passages.id: split_8_test_856_passage
passages.type: progene_text
passages.text: Previous studies have shown that the production of extracellular enzymes ( pectate lyase [ Pel ] , polygalacturonase [ Peh ] , cellulase [ Cel ] , and protease [ Prt ] ) and harpin ( Ecc ) ( the elicitor of hypersensitive reaction ) in Erwinia carotovora subsp. carotovora is regulated by RsmA , an RNA - binding protein , and rsmB , a regulatory RNA ( Rsm stands for regulator of secondary metabolites ) ( Y. Liu et al. , Mol. Microbiol. 29 : 219 - 234 , 1998 ) .