tokens: Comparison of the linkage map of mouse chromosome 9 ( MMU9 ) and that of human chromosome 11 ( HSA11 ) indicates that there is a chromosomal rearrangement due to an inversion between Ets1 and Atm - Npat - Acat1 and that the inversion of MMU9 originated from the chromosomal breakage at the boundary between Gria4 and Atm - Npat - Acat1 on HSA11 .