tokens: Five imprinted , paternally expressed genes map to the PWS region , MKRN3 ( ref . 3 ) , NDN ( ref . 4 ) , NDNL1 ( ref . 5 ) , SNRPN ( refs 6 - 8 ) and IPW ( ref . 9 ) , as well as two poorly characterized framents designated PAR - 1 and PAR - 5 ( ref . 10 ) .