Question: The Flying Fruit Fly Circus is in the twin cities separated by what landform?

Context paragraphs:
Tephritidae: The Tephritidae are one of two fly families referred to as fruit flies, the other family being the Drosophilidae.  The Tephritidae family does not include the biological model organisms of the genus "Drosophila" (in the family Drosophilidae), which is often called the "common fruit fly".  Nearly 5,000 described species of tephritid fruit fly are categorized in almost 500 genera of the Tephritidae.  Description, recategorization, and genetic analyses are constantly changing the taxonomy of this family.  To distinguish them from the Drosophilidae, the Tephritidae are sometimes called peacock flies, in reference to their elaborate and colorful markings.  The name comes from the Greek τεφρος, "tephros", meaning "ash grey".  They are found in all the ecozones.

Albury–Wodonga: Albury–Wodonga is the broad settlement incorporating the twin Australian cities of Albury and Wodonga, which are separated geographically by the Murray River and politically by a state border: Albury on the north of the river is part of New South Wales while Wodonga on the south bank is in Victoria.

Anastrepha suspensa: Anastrepha suspensa, also called the Greater Antilliean fruit fly or guava fruit fly is closely related to the Mexican fruit fly ("Anastrepha ludens").

Emma George: Emma George (born 1 November 1974 in Beechworth, Victoria) is a former Australian pole vaulter.  She set twelve world records in a row in the late nineties, but she lost the record on 26 May 2000 to Stacy Dragila and was unable to recapture it.  She was previously a trapeze artist in The Flying Fruit Fly Circus.  She was coached by world-renowned coach Mark Stewart, who also led Steve Hooker to Olympic gold.

Anastrepha obliqua: Anastrepha obliqua is a species of fruit fly.  It is the most important fruit fly pest of mangoes in Neotropics and attacks a wide range of other spicy fruits.  "A. obliqua" is widespread in Mexico, Central and South America and the West Indies.  It is sometimes called the West Indian fruit fly in English.

Flying Fruit Fly Circus School: The Flying Fruit Fly Circus School in Wodonga, Victoria is the only primary and secondary-level circus school in Australia.  It was founded in 1987 as the educational arm of The Flying Fruit Fly Circus.

Rhagoletis batava: Rhagoletis batava (seabuckthorn fruit fly) is a species of tephritid or fruit flies in the genus "Rhagoletis" of the family Tephritidae.  "Rhagoletis batava" larvae feed inside fruit flesh, and can be important pest of seabuckthorn.  This species is very similar to "Rhagoletis cerasi" (European cherry fruit fly)

Drosophila: Drosophila ( ) is a genus of flies, belonging to the family Drosophilidae, whose members are often called "small fruit flies" or (less frequently) pomace flies, vinegar flies, or wine flies, a reference to the characteristic of many species to linger around overripe or rotting fruit.  They should not be confused with the Tephritidae, a related family, which are also called fruit flies (sometimes referred to as "true fruit flies"); tephritids feed primarily on unripe or ripe fruit, with many species being regarded as destructive agricultural pests, especially the Mediterranean fruit fly.  One species of "Drosophila" in particular, "D. melanogaster", has been heavily used in research in genetics and is a common model organism in developmental biology.  The terms "fruit fly" and ""Drosophila"" are often used synonymously with "D. melanogaster" in modern biological literature.  The entire genus, however, contains more than 1,500 species and is very diverse in appearance, behavior, and breeding habitat.

Allan C. Spradling: Allan C. Spradling is an American scientist and principal investigator at the Carnegie Institution for Science and the Howard Hughes Medical Institute who studies egg development in the model organism, "Drosophila melanogaster", a fruit fly.  He is considered a leading researcher in the developmental genetics of the fruit fly egg and has developed a number of techniques in his career that have led to greater understanding of fruit fly genetics including contributions to sequencing its genome.  He is also an adjunct professor at Johns Hopkins University and at the Johns Hopkins University School of Medicine.

The Flying Fruit Fly Circus: The Flying Fruit Fly Circus is Australia's national youth circus and the only full-time circus school in Australia.  It is on the Murray River in the twin cities of Albury-Wodonga on the Victoria/New South Wales border.  In partnership with the Victorian Department of Education, the students/young artists in years 3-9 attend the selective entry Flying Fruit Fly Circus School where they undertake educational studies to complement their circus training, which is uniquely incorporated into the curriculum.  The Flying Fruit Fly Circus regularly performs in Australia and internationally at major arts centres, regional theatres, corporate events and festivals.

Supporting paragraphs: The Flying Fruit Fly Circus Albury–Wodonga
Answer: Murray River