Question: What was the former parent company of the pharmacy chain that discovered and developed Flosequinan?

Context paragraphs:
Navarro Discount Pharmacies: Navarro Discount Pharmacies, Inc., frequently referred to as Navarro is a pharmacy chain, photo service, and pharmacy benefit manager in the United States.  The company was acquired by CVS Pharmacy in September, 2014 and is operated as a separate brand and business of CVS, headquartered in Miami, Florida.  The company mainly operates in Miami-Dade and Broward Counties and currently has 28 stores.  Navarro Discount Pharmacies has a 17% market share in South Florida, ahead of Walgreens and CVS/pharmacy.

USA Drug: USA Drug is a convenience store and pharmacy chain in the United States that operates more than 160 stores in Arkansas, Oklahoma, Mississippi, Missouri, and Tennessee.  It was founded in Pine Bluff, Arkansas in 1986.  Its headquarters are still located in Pine Bluff with additional offices in Little Rock, Arkansas, Tulsa, Oklahoma, and Memphis, Tennessee.  Its parent company, Walgreens, operates Super D Drug Stores and Ikes Deep Discount Drug, both acquired in October 1997, and May's Drug, Med-X Drug, Drug Warehouse, and Drug Mart, all acquired in July 2004, as "Part of the USA Drug Family".  Wal-Mart and CVS/Pharmacy are two of its primary competitors.  The company's slogan is "America's Low Price Drug Store".

Charlie the Tuna: Charlie the Tuna is the cartoon mascot and spokes-tuna for the StarKist brand.  He was created in 1961 by Tom Rogers of the Leo Burnett Agency after StarKist hired Leo Burnett in 1958.  StarKist Tuna is the name of a brand of tuna currently owned by Dongwon Industries, a South Korea-based conglomerate.  StarKist itself is based in Pittsburgh, the home of its former parent company, H. J. Heinz Company, sharing its headquarters on the site of Three Rivers Stadium with another former parent company, Del Monte Foods' Pittsburgh headquarters.

Boots UK: Boots UK (formerly Boots the Chemists Ltd.), trading as Boots, is a pharmacy chain in the United Kingdom, Ireland, Norway, Thailand and other territories.  The company's former parent, The Boots Company Plc, merged with Alliance UniChem in 2006 to form Alliance Boots.  In 2007, Alliance Boots was bought by Kohlberg Kravis Roberts and Stefano Pessina, taking the company private, and moving its headquarters to Switzerland, the first ever FTSE 100 company bought by a private equity firm.  In 2012, Walgreens bought a 45% stake in Alliance Boots, with the option to buy the rest within three years.  It exercised this option in 2014, and as a result Boots became a subsidiary of the new company, Walgreens Boots Alliance, on 31 December 2014.

Osco Drug and Sav-on Drugs: Osco Drug and Sav-on Drugs were the names of a pair of chain pharmacies that operated in the United States.  Osco Drug was founded by the Skaggs family.  Alpha Beta grocery store was purchased by American Stores in 1961.  Skaggs Drug Centers bought American Stores in 1979 and assumed the American Stores name.  Sav-on Drugs was a California-based pharmacy chain that was acquired by Osco's parent company in 1980.  Both Osco and Sav-on stores eventually came under the ownership of American Stores, then Albertsons, and finally SuperValu before the stores were sold off.

Hartig Drug: Hartig Drug Stores is a chain of pharmacy/retail stores based out of Dubuque, Iowa.  The company provides pharmacy services not only to the general public, but to area institutions as well.  Along with this pharmacy focus, Hartig Drug stores offer convenience retail items.  The company has operated for over 100 years; currently, it is the second-oldest family-run pharmacy chain in the United States.

Well Pharmacy: Well, formerly The Co-operative Pharmacy, is the largest independent pharmacy business and the third largest overall pharmacy chain after Boots the Chemists and Lloyds Pharmacy (except in Wales, where Well is the biggest chain overall) in the United Kingdom.

Apotek 1 Gruppen: Apotek 1 Gruppen AS is a Norwegian pharmacy wholesaler ("Apokjeden Distribusjon AS") and retailer (Apotek 1) owned by German Phoenix Pharmahandel.  It owns the largest pharmacy chain in Norway, Apotek 1.  The company started a cooperation with a number of pharmacists who wanted a wholesaler independent of the state owned Norsk Medisinaldepot in 1995, when most pharmacies in the country joined the company.  1997 saw the first attempts to brand the pharmacies, as Plussapotek.  The first branded pharmacy as Apotek 1 was launched in 1999 in Manglerud Senter.  In 2001, the pharmacy market was deregulated, and Tamro was at first chosen as a wholesaler, before eventually buying the entire company in 2004.

Flosequinan: Flosequinan is a quinolone vasodilator that was discovered and developed by Boots UK and was sold for about a year under the trade name Manoplax.  It had been approved in 1992 in the US and UK to treat people with heart failure who could not tolerate ACE inhibitors or digitalis.

Uniprix: Uniprix is a Quebec pharmacy chain founded in 1977 and based in Saint-Leonard, Montreal.  It operates under four brands: Uniprix, Unipharm, Clinique Santé and Uniclinique (inside clinics).  Combining its four brands, Uniprix is the second-largest pharmacy chain in Quebec (after Jean Coutu) and is mutually exclusive with the Pharmasave chain in English Canada.

Supporting paragraphs: Flosequinan Boots UK
Answer: The Boots Company Plc