Question: Which actor in The 39 Steps also starred in the series The Strain?

Context paragraphs:
Sean Mahon: Sean Mahon is an Irish stage and screen actor.  He is known for playing Nicky Giblin in the Broadway production of "The Seafarer", Richard Hannay in the Broadway production of "The 39 Steps" and Michael Hess (the lost son) in the award winning feature film, "Philomena".  In 2014 he was nominated for a best lead actor award for his portrayal of maverick cop Brian McGonigle in the Irish Drama Red Rock.  After much success in Ireland, Red Rock now plays on the BBC in the UK and on Amazon Prime in the US.

The 39 Steps (1959 film): The 39 Steps is a 1959 British thriller film produced by Betty Box, directed by Ralph Thomas and starring Kenneth More and Taina Elg.  It is a remake of the 1935 Alfred Hitchcock film, loosely based on the novel "The Thirty-Nine Steps" by John Buchan.

Rupert Penry-Jones: Rupert William Penry-Jones (born 22 September 1970) is an English actor, best known for his roles as Adam Carter in the BBC One spy drama series "Spooks", Clive Reader QC in the BBC One legal drama "Silk", policeman DI Joseph Chandler in the ITV murder mystery series "Whitechapel", and Mr. Quinlan in the FX horror series "The Strain".

Stephen Critchlow: Stephen Critchlow is an English actor, notable for his work in the theatre and appearances on radio series such as "Truly, Madly, Bletchley", "The Way We Live Right Now" and "Spats", along with radio episodes of "Torchwood" ("Lost Souls") and "Doctor Who" ("The Nowhere Place").  He has also appeared in "" as Kenneth Horne and is currently appearing in the West End version of "The 39 Steps".

Robert Donat: Friedrich Robert Donath (18 March 19059 June 1958) was an English film and stage actor.  He is best remembered for his roles in Alfred Hitchcock's "The 39 Steps" (1935) and "Goodbye, Mr. Chips" (1939), winning for the latter the Academy Award for Best Actor.

The 39 Steps (1935 film): The 39 Steps is a 1935 British thriller film directed by Alfred Hitchcock, starring Robert Donat and Madeleine Carroll.  Very loosely based on the 1915 adventure novel "The Thirty-Nine Steps" by John Buchan, the film is about an everyman civilian in London, Richard Hannay, who becomes caught up in preventing an organization of spies called the 39 Steps from stealing British military secrets.  After being mistakenly accused of the murder of a counter-espionage agent, Hannay goes on the run to Scotland with an attractive woman in the hopes of stopping the spy ring and clearing his name.

John Behlmann: John Behlmann is an American actor known for such plays, films and television series as "The 39 Steps", "Significant Other", "The Wolf of Wall Street", "Revolutionary Road", "Guiding Light" and "All My Children".

The 39 Steps (2008 film): The 39 Steps is a 2008 British television adventure thriller feature-length adaptation of the John Buchan novel "The Thirty-Nine Steps" produced by the BBC.  It was written by Lizzie Mickery, directed by James Hawes, and filmed on location in Scotland, starring Rupert Penry-Jones, Lydia Leonard, David Haig, Eddie Marsan, and Patrick Malahide.  Following three screen versions of the novel and the 1952 and 1977 television adaptations of "The Three Hostages", Penry-Jones became the sixth actor to portray Hannay on screen.  This adaptation is set on the eve of the First World War and sees mining engineer Richard Hannay caught up in an espionage conspiracy following the death of a British spy in his flat.

Biosynthesis of doxorubicin: Doxorubicin (DXR) is a 14-hydroxylated version of daunorubicin, the immediate precursor of DXR in its biosynthetic pathway.  Daunorubicin is more abundantly found as a natural product because it is produced by a number of different wild type strains of "streptomyces".  In contrast, only one known non-wild type species, "streptomyces peucetius" subspecies "cesius" ATCC 27952, was initially found to be capable of producing the more widely used doxorubicin.  This strain was created by Arcamone et al. in 1969 by mutating a strain producing daunorubicin, but not DXR, at least in detectable quantities.  Subsequently, Hutchinson's group showed that under special environmental conditions, or by the introduction of genetic modifications, other strains of "streptomyces" can produce doxorubicin.  His group has also cloned many of the genes required for DXR production, although not all of them have been fully characterized.  In 1996, Strohl's group discovered, isolated and characterized dox A, the gene encoding the enzyme that converts daunorubicin into DXR.  By 1999, they produced recombinant Dox A, a Cytochrome P450 oxidase, and found that it catalyzes multiple steps in DXR biosynthesis, including steps leading to daunorubicin.  This was significant because it became clear that all daunorubicin producing strains have the necessary genes to produce DXR, the much more therapeutically important of the two.  Hutchinson's group went on to develop methods to improve the yield of DXR, from the fermentation process used in its commercial production, not only by introducing Dox A encoding plasmids, but also by introducing mutations to deactivate enzymes that shunt DXR precursors to less useful products, for example baumycin-like glycosides.  Some triple mutants, that also over-expressed Dox A, were able to double the yield of DXR.  This is of more than academic interest because at that time DXR cost about $1.37 million per kg and current production in 1999 was 225 kg per annum.  More efficient production techniques have brought the price down to $1.1 million per kg for the non-liposomal formulation.  Although DXR can be produced semi-synthetically from daunorubicin, the process involves electrophilic bromination and multiple steps and the yield is poor.  Since daunorubicin is produced by fermentation, it would be ideal if the bacteria could complete DXR synthesis more effectively.

Parboiled rice: Parboiled rice (also called converted rice) is rice that has been partially boiled in the husk.  The three basic steps of parboiling are soaking, steaming and drying.  These steps also make rice easier to process by hand, boost its nutritional profile and change its texture.  About 50% of the world’s paddy production is parboiled.  The treatment is practiced in many parts of the world such as India, Bangladesh, Pakistan, Myanmar, Malaysia, Nepal, Sri Lanka, Guinea, South Africa, Italy, Spain, Nigeria, Thailand, Switzerland, USA and France.

Supporting paragraphs: The 39 Steps (2008 film) Rupert Penry-Jones
Answer: Rupert Penry-Jones