Question: Melissa blue has a subspecies of which endangered butterfly?
Context: Erikssonia edgei, commonly known as the Waterberg copper, tilodi copper or Edge's acraea copper, is an obligate myrmecophylous lycaenid butterfly, which is native to Limpopo, South Africa.  The critically endangered butterfly occurs in high-altitude grasslands on sandy substrates, and has only been obtained from the type and one subsequent locality.  The population at the type locality, a farm in the Waterberg, went extinct about 12 years after its 1980 discovery.  It was afforded species status in 2010, when no extant populations were known.  The status of two populations, discovered in 2013 at a private nature reserve to the southeast, remains indeterminate. Lemurs are strepsirrhine primates, all species of which are endemic to Madagascar.  They include the smallest primate in the world, Madame Berthe's mouse lemur, which weighs 30 g , and range up to the size of the indri, which can weigh as much as 9.5 kg .  However, recently extinct species grew much larger.  As of 2010, five families, 15 genera, and 101 species and subspecies of lemur were formally recognized.  Of the 101 species and subspecies, the International Union for Conservation of Nature (IUCN) classified eight as Critically Endangered, 18 as Endangered, 15 as Vulnerable, four as Near Threatened, eight as Least Concern and 41 as Data Deficient; seven were yet to be evaluated.  From 2000 through 2008, 39 new species were described and nine other taxa resurrected.  By 2014, the number of species and subspecies recognized had increased to 105; of these, the IUCN classified 24 as Critically Endangered, 49 as Endangered, 20 as Vulnerable, three as Near Threatened, three as Least Concern and four as Data Deficient; two were yet to be evaluated. The Mission blue ("Aricia icarioides missionensis") is a blue or lycaenid butterfly subspecies native to the San Francisco Bay Area of the United States.  The butterfly has been declared as endangered by the US federal government.  It is a subspecies of Boisduval's blue ("Aricia icarioides"). The Melissa blue ("Plebejus melissa") is a butterfly of the family Lycaenidae.  It is found in western North America, from Canada to Mexico.  The Karner blue ("Plebejus melissa samuelis") is a subspecies of the Melissa blue, and was described by the novelist/lepidopterist Vladimir Nabokov. The Palos Verdes blue ("Glaucopsyche lygdamus palosverdesensis") is a small endangered butterfly native to the Palos Verdes Peninsula in southwest Los Angeles County, California, United States.  As its distribution has been proven to be limited to one single site it has one of the best claims to being the world's rarest butterfly. The northern Rocky Mountain wolf ("Canis lupus irremotus") is a subspecies of gray wolf native to the northern Rocky Mountains.  It is a light-colored, medium to large-sized subspecies with a narrow, flattened frontal bone.  The subspecies was initially listed as Endangered on March 9, 1978.  On August 6, 2010, the northern Rocky Mountain wolf was ordered to be returned under Endangered Species Act protections by U.S. District Judge Donald Molloy in a decision overturning a previous ruling by the U.S. Fish and Wildlife Service based on the case Defenders of Wildlife versus Salazar .  Wolves were later removed from federal protection in Idaho and Montana on May 6, 2011 under unprecedented legislation passed by the U.S. Congress removing them from the endangered species list. The Saint Francis' satyr ("Neonympha mitchellii francisci") is an endangered butterfly found only in the US state of North Carolina.  First discovered in 1983, it was officially described in 1989 and listed as a federally endangered species by the US Fish and Wildlife Service in 1994.  It is a subspecies of "N. mitchellii" and is only known from a single metapopulation on Fort Bragg military base in Hoke and Cumberland counties.  The other subspecies, Mitchell's satyr ("Neonympha mitchellii mitchellii"), is also federally endangered. The Karner blue ("Plebejus melissa samuelis") is an endangered subspecies of small blue butterfly which was once found in significant numbers in the Miller section of Gary, Indiana community of the Indiana Dunes National Park.  The butterfly is now mainly found in other parts of the Great Lakes states, such as Wisconsin and Michigan, in small areas of New Jersey, and also in southern New Hampshire, and the Capital District region of New York.  The butterfly, whose life cycle depends on the wild blue lupine flower ("Lupinus perennis"), was classified as an endangered species in 1992.  In May 2000, the Canadian Species at Risk Act listed the Karner blue as being locally extinct in Canada.  This subspecies of "Plebejus melissa" was described by novelist Vladimir Nabokov. The salt marsh harvest mouse ("Reithrodontomys raviventris"), also known as the red-bellied harvest mouse and sometimes called the saltmarsh harvest mouse, is an endangered rodent endemic to the San Francisco Bay Area salt marshes in California.  There are two distinct subspecies, both endangered and listed together on federal and state endangered species lists.  The northern subspecies ("Reithrodontomys raviventris halicoetes") is lighter in color and inhabits the northern marshes of the bay, and the southern subspecies ("Reithrodontomys raviventris raviventris") lives in the East and South Bay marshes.  They are both quite similar in appearance to their congener species, the [Western harvest mouse, R. megalotis], to which they are not closely related.  Genetic studies of the northern subspecies have revealed that the salt marsh harvest mouse is most closely related to the plains harvest mouse, R. montanus, (), which occurs now in the Midwest].  Its endangered designation is due to its limited range, historic decline in population and continuing threat of habitat loss due to development encroachment at the perimeter of San Francisco Bay. Fender's blue butterfly ("Icaricia icarioides fenderi" syn.  "Plebejus icarioides fenderi") is an endangered subspecies of butterfly endemic to the Willamette Valley of northwestern Oregon, United States.  The species was first documented in the 1920s and was described to science in 1931 by biologist Ralph Macy.  He named it for his friend, Kenneth Fender, an entomologist and mail carrier.  The species was not seen after the 1930s and was presumed extinct.  Small populations were rediscovered in 1989.  Its eponym, Fender, had died nine years earlier.  In January 2000, Fender's blue butterfly was added to the Endangered Species List by the U.S. Fish and Wildlife Service.  The largest known populations now exist in the Baskett Slough National Wildlife Refuge.  A 2014 study reintroduced this species to William L. Finley National Wildlife Refuge.
Answer: Karner blue