The English Mastiff is a breed of large dog. The lineage of modern dogs can be traced back to the early 19th century, but the modern type was stabilised in the 1880s and refined since. On the basis of what is known (and remembering that, as dogs are diploid animals, each gene location (locus) appears twice in every animal, so questions of dominance also must be resolved), the gene possibilities allowed by the Mastiff standard are AyBDEmh(kbr_or_ky)mS. This describes a dog which is fawn with a dark nose, non-dilute, black-masked, non-harlequin, brindled or not brindled, non-merle, and non-spotted. To allow for the rare exceptions we must include 'b' (brown mask and possible brown brindling), 'd' (blue mask and possible blue brindling), 'sp' (pied spotting), and perhaps 'a' (recessive black).
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