Wildflowers such as lupine (Lupinus argenteus) and arnica (Arnica cordifolia) often grow under the forest canopy, but the most conspicuous wildflower displays occur in open meadows and sagebrush-steppe. The appearance of springbeauties (Claytonia lanceolata), glacier lilies (Erythronium grandiflorum), and steershead (Dicentra uniflora) announce spring in the park. Soon colors splash the slopes, especially on the northern range—yellow from arrowleaf balsamroot (Balsamorhiza sagittata), white from phlox (Phlox multiflora), reds and oranges from paintbrush (Castilleja), and blue from penstemon (Penstemon montanus) and lupine. Goldenrod (Solidago missouriensis) and purple asters indicate the coming of fall.