In shape they resemble the old southern hound in miniature, but with more neatness and beauty; and they also resemble that hound in style of hunting. Alongside the Bloodhound and Basset Hound, the beagle has one of the best developed senses of smell of any dog. In the 1950s, John Paul Scott and John Fuller began a 13-year study of canine behavior. Beagles are better at ground-scenting (following a trail on the ground) than they are at air-scenting, and for this reason they have been excluded from most mountain rescue teams in favor of collies, which use sight in addition to air-scenting and are more biddable. The long ears and large lips of the beagle probably assist in trapping the scents close to the nose.