100/2112), Citipati sp. (MPC-D 100/42) and Rinchenia mongoliensis (MPC-D 100/32-1). The specimen is not well enough preserved to determine whether or not the nasal was pneumatic, as it clearly was in Rinchenia and other mature oviraptorid individuals. The presence of a crest is generally associated with oviraptorines rather than ingeniines, but may in fact be correlated with size and maturity; most ingeniines are in fact smaller than most oviraptorines [2]. The anteroventral margin of the premaxilla is almost perpendicular to the ventral margin of the jugal (Figure 5). The height of the premaxilla below the external naris is 44.8 mm. The premaxillary-maxillary suture is nicely preserved on the right side of the skull (Figure 5). If a line is drawn through the ventral margins of the quadratojugal and maxilla (Fig. 5A), then the tomial margin of the premaxilla clearly projects well below the maxilla. This may ultimately turn out to be a size correlated character of little taxonomic utility. The lacrimal is platelike and projects lateral to the orbit in both MPC-D 107/15 and the holotype. Although the skull roof cannot be seen as well as in the holotype, the nasal of MPC-D 107/15 appears to have been no longer than the frontal. It appears unlikely that the frontals and parietals were pneumatized like the nasals, because the small sections of preserved surfaces of these bones are flat in MPC-D 107/15. There is no evidence for a separate prefrontal in either this specimen or the holotype. The postorbital process of the jugal extends posterodorsally from the suborbital process of the jugal (Figure 5). The basal tubera cannot be seen in MPC-D 107/15, but appear to be well developed in the holotype [14]. As in other oviraptorids, the jaw is deep (the maximum height of the dentary is a third of the jaw length), and the external mandibular fenestra is large, making up more than a quarter of the jaw length (Figure 5). A prominent process of the surangular invades the back margin of the external mandibular fenestra as in all oviraptorids. Postcranially, the ends of most bones were destroyed, presumably by insect scavengers (Figures 3, 4). Most of the scapula (180 mm) is present, and the total length appears to have been about 185 mm. The minimum shaft width is 15 mm, and the distal end expands to 27 mm. Although the full length of the humerus is not preserved, the minimum shaft width (transverse) has a diameter of 19 mm. Amongst sixteen oviraptorosaurs (Table S4), there is a strong correlation (r2 = 0.95) between humerus length and transverse shaft width, which suggests that the humerus would have been 152 mm long in MPC-D 107/15. The ratio of the lengths of the scapula to the humerus can therefore be calculated as 1.2, which is typical of the majority of oviraptorids. The distal end of the deltoid process is preserved, but little can be said other than it was well developed and extended far down the shaft as in other oviraptorids. Only small sections of the shafts of the radius (the diameter of the shaft width is 12 mm) and ulna (SW = 14 mm) are preserved, but show that the diameter of the shaft of the radius was 0.86 that of the ulna. The length of the radius was estimated as 144 mm using 10 pairs of measurements of oviraptorid radii, plus the measurements of radii from seven individuals of the oviraptorosaurs Avimimus and Caudipteryx (b = 1.35, k = 0.75, r2 = 0.89). The estimates based on shaft widths suggest that the ratio of radius length to humerus length is 0.95, which is close to the same ratios in Citipati osmolskae (MPC-D 100/ 979), and Conchoraptor gracilis (MPC-D 110/21). In contrast, the ratio is less than 0.80 in Ingenia (MPC-D 100/32, 100/33, 110/03) and Rinchenia (MPC-D 100/32-1). The ratio is intermediate between the two extremes in all other oviraptorid taxa. The hands were also badly damaged, but it is clear that the phalanges (I-1, I-2) of the first digit were more massive than those of the second digit (II-2, II-3) (Fig. 6A), that the distal end of the second digit does not extend far beyond the distal end of the first digit, and New Specimens of Nemegtomaia from Mongolia Figure 6. Hands of Nemegtomaia barsboldi. A, preserved elements and reconstructed manus of MPC-D 107/15 in dorsal (a1) and lateral (a2) views: B, right manus of MPC-D 107/16 in dorsal (b1) and medial (b2) views; C, left manus of MPC-D 107/16 in medial (c1) and dorsal (c2) views, with the preserved distal portions of radius and ulna. lr, left radius; lu, left ulna. Scale bar 5 cm. Illustration by Marco Auditore. doi:10.1371/journal.pone.0031330.g006 that the third metatarsal is slender. These are all characters that are diagnostic for Ingeniinae. The pubic shaft is preserved in MPC-D 107/15, and was clearly curved anteroventrally (is anteriorly concave in lateral view) as in other oviraptorids. Most of the femur is preserved, although the distal end is in poor condition (Fig. 4B). Its length can be estimated from the transverse diameter of the shaft (31 mm) and is approximately 286 mm when compared with 26 oviraptorids and dromaeosaurids (b = 1.22, PLoS ONE | www.plosone.org 7 February 2012 | Volume 7 | Issue 2 | e31330