Leptusa

Leptusa er en slekt av biller som hører til den store underfamilien småkortvinger (Aleocharinae) i familien kortvinger (Staphylinidae). Det er beskrevet mer enn 400 arter, fire av dem er funnet i Norge.

Utseende  
Forholdsvis små (gjerne 2-4 millimeter lange), middels slanke kortvinger, kroppen nokså jevnbred. På farge er de ofte rødlige eller brunlige. Hodet er ganske bredt med nokså små fasettøyne. Pronotum er bredere enn langt, med rundede sider og hjørner. Dekkvingene er forholdsvis korte og har bakkanten tydelig innbuktet innenfor bakhjørnene. Føttene er fire-leddete.

Levevis 
Leptusa-artene er vanlige på skogbunnen, og kan også finnes under løs bark på døde trær.

Systematisk inndeling / norske arter 
Denne artsrike slekten blir delt opp i ca. 80 underslekter.

 Ordenen biller, Coleoptera
 Underordenen Polyphaga
 Overfamilien åtsel- og rovbiller, Staphylinoidea
 Familien kortvinger (Staphylinidae) Latreille, 1802
 Underfamilien småkortvinger (Aleocharinae) Fleming, 1821
 Stammen Homalotini Heer, 1839
 Slekten Leptusa Kraatz, 1856
 Leptusa pulchella (Mannerheim, 1831)
 Leptusa fumida (Erichson, 1839)
 Leptusa norvegica A. Strand, 1941
 Leptusa ruficollis (Erichson, 1839)

Liste over artene 

 Leptusa abchasica Bernhauer, 1936
 Leptusa abdominalis (Motschulsky, 1851)
 Leptusa abeillei Bernhauer, 1900
 Leptusa acuta Assing, 2002
 Leptusa aisenicola Pace, 2001
 Leptusa albergianensis Pace, 1989
 Leptusa allobrogica Pace, 2000
 Leptusa alpicola Brancsik, 1874
 Leptusa amisensis Pace, 1982
 Leptusa andina (Fauvel, 1866)
 Leptusa angustiarumberninae Scheerpeltz, 1972
 Leptusa anmashanensis Pace, 1996
 Leptusa annapurnensis Pace, 1982
 Leptusa antarctica (Kiesenwetter, 1877)
 Leptusa apennina Holdhaus, 1924
 Leptusa apfelbeckiana Likovský, 1984
 Leptusa araucana Pace, 2001
 Leptusa areraensis Scheerpeltz, 1973
 Leptusa argentierensis Pace, 1999
 Leptusa armatissima Assing, 2008
 Leptusa armeniaca Pace, 1989
 Leptusa artviniensis Pace, 1982
 Leptusa asiatica Bernhauer, 1909
 Leptusa asperata Eppelsheim, 1888
 Leptusa assingi Pace, 1996
 Leptusa asturica Pace, 1983
 Leptusa asturiensis Eppelsheim, 1880
 Leptusa athosensis Zerche, 2002
 Leptusa atriceps (Waterhouse, 1875)
 Leptusa atrocephala Bernhauer, 1905
 Leptusa austriaca Scheerpeltz, 1935
 Leptusa baldensis Ganglbauer, 1895
 Leptusa baldomontis Scheerpeltz, 1972
 Leptusa bellinorum Assing, 2004
 Leptusa benacensis Pace, 1980
 Leptusa bengalensis Pace, 1982
 Leptusa bergamasca Scheerpeltz, 1972
 Leptusa bernhaueri Scheerpeltz, 1926
 Leptusa binaghii Pace, 1979
 Leptusa biumbonata Scheerpeltz, 1973
 Leptusa blesioi Pace, 1981
 Leptusa bonvouloirii Brisout de Barneville, 1867
 Leptusa bozdaghensis Pace, 1983
 Leptusa brachati Pace, 1980
 Leptusa brevicollis Casey, 1893
 Leptusa brigantii Pace, 1979
 Leptusa brixiensis Pace, 1979
 Leptusa brucki W. Scriba, 1867
 Leptusa cabrerensis Pace, 1981
 Leptusa cadiensis Pace, 1989
 Leptusa calancensis Szallies, 2014
 Leptusa californiana Pace, 1989
 Leptusa calliceroides Assing, 2004
 Leptusa camunnensis Pace, 1981
 Leptusa canonica Casey, 1906
 Leptusa cansigliensis Pace, 1975
 Leptusa cantabrica Paśnik, 1998
 Leptusa carinata Bernhauer, 1909
 Leptusa carniolica Pace, 1983
 Leptusa carolinensis Pace, 1989
 Leptusa carpathica J. Weise, 1877
 Leptusa casalei Pace, 1979
 Leptusa cassagnaui Pace, 1984
 Leptusa caucasica Eppelsheim, 1878
 Leptusa cautinensis Pace, 2001
 Leptusa cavallensis Pace, 1980
 Leptusa cavallicola Pace, 1980
 Leptusa cavata Assing, 2003
 Leptusa centralis Pace, 1992
 Leptusa cephalotes Bernhauer, 1909
 Leptusa ceresoleana Pace, 1979
 Leptusa cerrutii Coiffait, 1970
 Leptusa chengduensis Pace, 2001
 Leptusa chilensis (Fauvel, 1866)
 Leptusa chinensis Pace, 1997
 Leptusa cimmeria Pace, 1996
 Leptusa circassica Bernhauer, 1935
 Leptusa coiffaiti Pace, 1989
 Leptusa confinis Pace, 1982
 Leptusa cordicollis Portevin, 1929
 Leptusa cornigera Assing, 2002
 Leptusa corsica Pace, 1989
 Leptusa crassiventris Scheerpeltz, 1973
 Leptusa cremata Assing, 2003
 Leptusa crenulata Bernhauer, 1899
 Leptusa cribrata Pace, 1996
 Leptusa cribratula (Casey, 1906)
 Leptusa cribripennis Kraatz, 1856
 Leptusa crinita Assing, 2007
 Leptusa cultellata Assing, 2008
 Leptusa cuneensis Pace, 1979
 Leptusa curvata Assing, 2006
 Leptusa cypria Pace, 1996
 Leptusa delphiniformis Assing, 2009
 Leptusa deplanata (K. Sawada, 1970)
 Leptusa deprehendens Pace, 1981
 Leptusa devoluyiensis Pace, 1989
 Leptusa diecki Pace, 1983
 Leptusa difficilis Pace, 1996
 Leptusa difformis (Mulsant & Rey, 1853)
 Leptusa discolor Assing, 2006
 Leptusa dissimilans Assing, 2004
 Leptusa dubia
 Leptusa durmitorensis Pace, 1984
 Leptusa elegans
 Leptusa emplenotoides Assing, 2006
 Leptusa epirensis Pace, 1989
 Leptusa erlangensis Pace, 1999
 Leptusa estrelensis Pace, 1981
 Leptusa etrusca Pace, 1979
 Leptusa europaea Pace, 1996
 Leptusa everestensis Pace, 1989
 Leptusa excaecata Assing, 2002
 Leptusa eximia
 Leptusa fauciumberninae Scheerpeltz, 1973
 Leptusa fauciumredortae Scheerpeltz, 1972
 Leptusa fauveli Eppelsheim, 1889
 Leptusa ferroi
 Leptusa fibula Assing, 2003
 Leptusa filiformis Scheerpeltz, 1972
 Leptusa fischeri Bernhauer, 1910
 Leptusa flagellata Assing, 2002
 Leptusa flagrifera Assing, 2009
 Leptusa flavicornis Brancsik, 1874
 Leptusa florae Scheerpeltz, 1926
 Leptusa formidabilis Pace, 1996
 Leptusa foroiuliensis Pace, 1980
 Leptusa franzi Pace, 1981
 Leptusa franziana Pace, 1981
 Leptusa frontalis
 Leptusa fuliginosa
 Leptusa fumida
 Leptusa gadesensis Pace, 1982
 Leptusa gaisbergeri
 Leptusa gansuensis Pace, 1997
 Leptusa ganzica Assing, 2002
 Leptusa gatineauensis
 Leptusa gerlitzensis Pace, 1983
 Leptusa ghoropanensis Pace, 1989
 Leptusa gibbera Assing, 2011
 Leptusa gimmeli
 Leptusa glabriceps Bernhauer, 1923
 Leptusa globulicollis (Mulsant & Rey, 1853)
 Leptusa gonggamontis Pace, 1997
 Leptusa gracilenta Pace, 1983
 Leptusa gracilipes
 Leptusa gracilis
 Leptusa gracillima Pace, 1983
 Leptusa granulicauda Eppelsheim, 1890
 Leptusa granulipennis Eppelsheim, 1880
 Leptusa grignaensis Scheerpeltz, 1973
 Leptusa hamulata Assing, 2010
 Leptusa hastata Assing, 2002
 Leptusa helambuica Assing, 2009
 Leptusa hercegovinensis Bernhauer, 1910
 Leptusa herminia Pace, 1982
 Leptusa hiarens Assing, 2011
 Leptusa himalayiana Pace, 1989
 Leptusa hlavaci Assing, 2006
 Leptusa hlisnikovskyi Pace, 1983
 Leptusa hoelzeli Scheerpeltz, 1948
 Leptusa honshuica Assing, 2002
 Leptusa hopffgarteni Eppelsheim, 1881
 Leptusa hummleri Bernhauer, 1910
 Leptusa hummleriana Bernhauer, 1935
 Leptusa ilamensis Pace, 1987
 Leptusa ilsae Bernhauer, 1935
 Leptusa improvisa Assing, 2009
 Leptusa indica
 Leptusa inexpectata
 Leptusa ionopolitana Pace, 1982
 Leptusa ishizuchiensis Pace, 1982
 Leptusa italica Pace, 1977
 Leptusa janbellini Assing, 2001
 Leptusa janczyki Pace, 1983
 Leptusa janetscheki Scheerpeltz, 1956
 Leptusa japonica
 Leptusa jeanneli Scheerpeltz, 1935
 Leptusa jelineki Pace, 1983
 Leptusa jinfomontis Pace, 2001
 Leptusa jiudingensis Pace, 1999
 Leptusa jucunda
 Leptusa juliana Pace, 1980
 Leptusa jurassica
 Leptusa kahleni Pace, 1980
 Leptusa kangdingensis Pace, 2010
 Leptusa kaohsiungensis Pace, 2007
 Leptusa kappenbergeri
 Leptusa karawankarum Pace, 1983
 Leptusa kaszabi Pace, 1981
 Leptusa kaufmanni
 Leptusa khnzoriani Pace, 1982
 Leptusa kitazawai (K. Sawada, 1970)
 Leptusa knabli Bernhauer, 1909
 Leptusa knappeorum Scheerpeltz, 1964
 Leptusa kocae Bernhauer, 1905
 Leptusa kochiana Bernhauer, 1936
 Leptusa komareki Pace, 1983
 Leptusa koralpicola Pace, 1983
 Leptusa korgei Scheerpeltz, 1970
 Leptusa koronensis
 Leptusa kosmajensis Pace, 1987
 Leptusa lackneri Assing, 2011
 Leptusa laevicauda (Scheerpeltz, 1958)
 Leptusa laeviuscula
 Leptusa lapidicola
 Leptusa laticeps Scheerpeltz, 1973
 Leptusa lativentris
 Leptusa lederi Eppelsheim, 1883
 Leptusa leonhardi
 Leptusa leonica Pace, 1981
 Leptusa leptotyphloides
 Leptusa lessiniensis Pace, 1978
 Leptusa limata Assing, 2002
 Leptusa linderi Scheerpeltz, 1966
 Leptusa linderiana Scheerpeltz, 1966
 Leptusa loebli Pace, 1984
 Leptusa lohsei Pace, 1989
 Leptusa lombarda Bernhauer, 1905
 Leptusa longalata Assing, 2017
 Leptusa longilobata Assing, 2007
 Leptusa lophophororum Pace, 1987
 Leptusa luigionii Bernhauer, 1935
 Leptusa luremontis Pace, 1999
 Leptusa lurensis Pace, 1989
 Leptusa magellanica Pace, 1989
 Leptusa majevicensis Scheerpeltz, 1935
 Leptusa major Bernhauer, 1900
 Leptusa maleschevskaensis
 Leptusa manasluensis Assing, 2002
 Leptusa mancinii Pace, 1980
 Leptusa mandli Scheerpeltz, 1953
 Leptusa manfredi Pace, 1980
 Leptusa marasica Assing, 2006
 Leptusa marianii Pace, 1979
 Leptusa marieluiseae
 Leptusa marmotae Assing, 2002
 Leptusa martensi Pace, 1987
 Leptusa media Scheerpeltz, 1973
 Leptusa merditana
 Leptusa merkli Bernhauer, 1900
 Leptusa meschniggi Bernhauer, 1935
 Leptusa meybohmi Assing, 2014
 Leptusa michai Assing, 2002
 Leptusa microphthalma
 Leptusa microvolans Pace, 1997
 Leptusa migrituber Assing, 2017
 Leptusa minutissima Pace, 1983
 Leptusa moczarskii Bernhauer, 1915
 Leptusa monacha (Fauvel, 1898)
 Leptusa monachorum Bernhauer, 1912
 Leptusa monguzzii Pace, 1989
 Leptusa montana (Fauvel, 1866)
 Leptusa montiscathedrae Pace, 1978
 Leptusa montisgrappae Pace, 1975
 Leptusa montisgrignae Scheerpeltz, 1973
 Leptusa montisparii Scheerpeltz, 1953
 Leptusa montispasubii Pace, 1975
 Leptusa montiumcarnorum (Scheerpeltz, 1958)
 Leptusa mucronemontis Scheerpeltz, 1972
 Leptusa munelensis
 Leptusa muscicola
 Leptusa myrmidon
 Leptusa nahuelbutana
 Leptusa nanula
 Leptusa nemoricultrix Pace, 1995
 Leptusa nepalica Scheerpeltz, 1976
 Leptusa nepalorum Pace, 1982
 Leptusa netolitzkyi Bernhauer, 1914
 Leptusa newar Pace, 1989
 Leptusa nigerrima Pace, 1983
 Leptusa nitidiventris
 Leptusa nonveilleri Pace, 1984
 Leptusa norica Pace, 1979
 Leptusa norvegica
 Leptusa notabilis Scheerpeltz, 1972
 Leptusa nubigena
 Leptusa numidica Pace, 1983
 Leptusa nurdaghensis Assing, 2003
 Leptusa obirensis Scheerpeltz, 1935
 Leptusa obscura
 Leptusa occulta Pace, 1975
 Leptusa ochsi Pace, 1983
 Leptusa oertzeni Eppelsheim, 1888
 Leptusa olympica Pace, 1994
 Leptusa opaca
 Leptusa opacina Pace, 2001
 Leptusa oreophila
 Leptusa oropaensis Pace, 1979
 Leptusa orumboviorum Pace, 1982
 Leptusa osellai Pace, 1977
 Leptusa pangeoensis Assing & Wunderle, 1999
 Leptusa paradoxa Pace, 1980
 Leptusa parallela
 Leptusa parvibulbata Assing, 2008
 Leptusa pascuorum Pace, 1975
 Leptusa pasubiana Scheerpeltz, 1953
 Leptusa pedemontana Bernhauer, 1935
 Leptusa peinantamontis Pace, 2007
 Leptusa pelionensis Bernhauer, 1916
 Leptusa peninsulae Pace, 1989
 Leptusa peregrina Pace, 1995
 Leptusa peristerica Pace, 1989
 Leptusa persica Assing, 2009
 Leptusa petzeniensis Pace, 1989
 Leptusa peyerimhoffi Bernhauer, 1935
 Leptusa piceata (Mulsant & Rey, 1853)
 Leptusa piciformis Assing, 2009
 Leptusa pilatensis Scheerpeltz, 1972
 Leptusa pingtungensis Pace, 1995
 Leptusa pinkeri
 Leptusa pirinensis
 Leptusa pizzocolensis Pace, 1981
 Leptusa plitvicensis Pace, 1980
 Leptusa poggii Pace, 1979
 Leptusa pollicita Assing, 2010
 Leptusa portusnaoniensis Pace, 1989
 Leptusa pratensis Pace, 1989
 Leptusa priesneri Pace, 1989
 Leptusa proiecta Assing, 2008
 Leptusa pseudoalpestris Scheerpeltz, 1935
 Leptusa pseudocaucasica Pace, 1983
 Leptusa pseudopaca
 Leptusa pseudosmokyiensis
 Leptusa puellaris
 Leptusa puetzi Assing, 2008
 Leptusa pulchella
 Leptusa punctithorax Bernhauer, 1935
 Leptusa punctulata (Mulsant & Rey, 1875)
 Leptusa pusio
 Leptusa puthzi Assing, 2002
 Leptusa pyrenaica Pace, 1982
 Leptusa qinlingensis Pace, 1999
 Leptusa quinqueimpressa Assing, 2008
 Leptusa rambouseki
 Leptusa recta Assing, 2006
 Leptusa reinosensis Pace, 1996
 Leptusa reitteri Eppelsheim, 1880
 Leptusa repentina Scheerpeltz, 1972
 Leptusa rhaetoromanica
 Leptusa rhilensis Pace, 1983
 Leptusa rizensis Pace, 1996
 Leptusa rorata Pace, 1995
 Leptusa rosai Pace, 1979
 Leptusa roscidavallensis Pace, 1981
 Leptusa rossica Bernhauer, 1915
 Leptusa rougemonti Pace, 1997
 Leptusa rousi Pace, 1983
 Leptusa ruffoi Pace, 1983
 Leptusa ruficollis
 Leptusa ruficornis
 Leptusa rugatipennis
 Leptusa sabauda Pace, 1989
 Leptusa salernitana Pace, 1979
 Leptusa salonichia Bernhauer, 1914
 Leptusa samia Assing, 2004
 Leptusa sampeyrensis Pace, 2009
 Leptusa sanfilippoi Pace, 1980
 Leptusa sarensis Pace, 1983
 Leptusa savonensis Pace, 1977
 Leptusa scabripennis (Mulsant & Rey, 1875)
 Leptusa schaschli
 Leptusa schawalleri Assing, 2011
 Leptusa schoenmanni Pace, 1983
 Leptusa schuelkei Pace, 1999
 Leptusa secreta Bernhauer, 1900
 Leptusa semiscabra Scheerpeltz, 1972
 Leptusa semivolans Pace, 1996
 Leptusa sengleti Pace, 1984
 Leptusa serbica Scheerpeltz, 1935
 Leptusa seriana Scheerpeltz, 1973
 Leptusa serullazi
 Leptusa shaanxiensis Pace, 1999
 Leptusa sibirica Pace, 1982
 Leptusa sibyllinica Pace, 1977
 Leptusa sica Assing, 2003
 Leptusa sichuanensis Pace, 1997
 Leptusa simoni Eppelsheim, 1878
 Leptusa sinorum Pace, 2001
 Leptusa smetanai Pace, 1989
 Leptusa smetanaiella Pace, 1989
 Leptusa smetanaorum Pace, 1989
 Leptusa smokyiensis Pace, 1989
 Leptusa soganlica Assing, 2007
 Leptusa solieri Pace, 1989
 Leptusa soriciformis Assing, 2009
 Leptusa sororella Pace, 1989
 Leptusa sparsa (Fauvel, 1898)
 Leptusa spirarum Pace, 1996
 Leptusa spoliata Assing, 2002
 Leptusa stimulans Assing, 2008
 Leptusa stocksi
 Leptusa stoeckleini Bernhauer, 1914
 Leptusa storkani Roubal, 1917
 Leptusa subalpina
 Leptusa subcarpathica Roubal, 1930
 Leptusa subcaucasica Pace, 1983
 Leptusa subconvexa (Mulsant & Rey, 1875)
 Leptusa subnivalis (Roubal, 1911)
 Leptusa substricta Assing, 2017
 Leptusa sudetica Lokay, 1900
 Leptusa sulcicollis Bernhauer, 1932
 Leptusa svanetica Assing, 2017
 Leptusa taichii Kishimoto, 2008
 Leptusa taichungensis Pace, 1996
 Leptusa taiwanensis Pace, 1992
 Leptusa tarvisiana Pace, 1989
 Leptusa taurica Assing, 2004
 Leptusa taurinensis Pace, 1989
 Leptusa tectusoides Assing, 2002
 Leptusa temperei Pace, 1989
 Leptusa tenchiensis Pace, 1996
 Leptusa tendamontis Scheerpeltz, 1972
 Leptusa tendicola Pace, 1999
 Leptusa tenerrima Bernhauer, 1900
 Leptusa tenuicornis Assing, 2006
 Leptusa tenuissima Scheerpeltz, 1972
 Leptusa thailandensis Pace, 2004
 Leptusa tirolensis Bernhauer, 1900
 Leptusa titillans Assing, 2002
 Leptusa tomorensis Pace, 1989
 Leptusa toumayeffi Scheerpeltz, 1972
 Leptusa transversiceps Pace, 1983
 Leptusa trapezuntis Pace, 1989
 Leptusa triangulata Assing, 2017
 Leptusa tricolor W. Scriba, 1870
 Leptusa tridentina Pace, 1980
 Leptusa trumplinensis Pace, 1981
 Leptusa turgida Assing, 2006
 Leptusa tusculanensis Pace, 1981
 Leptusa ultracollensis Pace, 1982
 Leptusa umbhakica Assing, 2011
 Leptusa vailatii Pace, 1989
 Leptusa vallisvenyi Scheerpeltz, 1972
 Leptusa veniorum Pace, 1999
 Leptusa ventosensis Pace, 1996
 Leptusa venusta (Hochhuth, 1849)
 Leptusa vercorensis Pace, 1989
 Leptusa vermiensis Pace, 1981
 Leptusa veronensis Pace, 1983
 Leptusa vietnamensis Pace, 1992
 Leptusa wechseliensis Pace, 1980
 Leptusa weiratheri Scheerpeltz, 1972
 Leptusa winkleri Scheerpeltz, 1926
 Leptusa winkleriana Scheerpeltz, 1965
 Leptusa winneguthi Linke, 1938
 Leptusa winneguthiana Pace, 1983
 Leptusa woerndlei Scheerpeltz, 1935
 Leptusa wolongensis Assing, 2002
 Leptusa wunderlei Pace, 1996
 Leptusa wuyica Assing, 2002
 Leptusa xanthopyga Eppelsheim, 1881
 Leptusa xiahensis Pace, 1997
 Leptusa xianensis Pace, 1999
 Leptusa xuemontis Pace, 2001
 Leptusa yakorum Pace, 1987
 Leptusa yamato (K. Sawada, 1990)
 Leptusa yunnanensis Pace, 2001
 Leptusa zanettiorum Pace, 1980
 Leptusa zerchei Pace, 1989
 Leptusa zhemomontis Assing, 2010
 Leptusa zhongdianensis Pace, 2010
 Leptusa zoiai Pace, 1982

Kilder 
 Park, J.-S., C.E. Carlton og M.L. Ferro 2010. Diversity and taxonomic review of Leptusa Kraatz (Coleoptera: Staphylinidae: Aleocharinae) from Great Smoky Mountains National Park, U.S.A., with descriptions of four new species. Zootaxa 2662: 1-27. 
 Silfverberg, H. 1992. Enumeratio Coleopterorum Fennoscandiae, Daniae et Baltiae (Liste over Nordens biller). Helsinki.
 Fauna Europaea Web Service Version 2.4 (2011) Utbredelsesdatabase for europeiske dyr.

Eksterne lenker 
 

Småkortvinger
Biller formelt beskrevet i 1856