Ptomaphaginus

Ptomaphaginus er en slekt av mycelbiller som særlig finnes knyttet til sopp og åtsler.

Utseende 
Små  (2-4 millimeter), halvmatte, mørkbrunlige mycelbiller, kroppsformen nærmest dråpeformet med lange bein. Antennene er litt kølleformede og 11-leddete. De fem ytterste leddene danner en avlang klubbe, men det åttende leddet er mye mindre enn de andre og er vanligvis forkortet til en liten skive. Hodet er oftest kraftig innskåret bak fasettøynene, med en smal hals, men dette kan være vanskelig å se da billene gjerne trekker hodet inn under brystskjoldet. Hannen har utvidete føtter på forbeina.

Levevis 
De fleste av artene lever i huler eller nede i jorda, og er gjerne blinde.

Utbredelse 
De aller fleste av artene er utbredt i Sørøst-Asia.

Systematisk inndeling 

 Ordenen biller, Coleoptera
 Underordenen Polyphaga
 Overfamilien åtsel- og rovbiller, Staphylinoidea
 Familien mycelbiller, Leiodidae Fleming, 1821
 Underfamilien kadaverbiller, Cholevinae
 Stammen Ptomaphagini
 Slekten Ptomaphaginus Portevin, 1914

Liste over artene 

 Ptomaphaginus agostii Schilthuizen & Perreau, 2008
 Ptomaphaginus alabensis  Schilthuizen & Perreau, 2018
 Ptomaphaginus anas Schilthuizen & Perreau, 2008
 Ptomaphaginus angusticornis (Portevin, 1921)
 Ptomaphaginus apiculatus Szymczakowski, 1975
 Ptomaphaginus assimilis Szymczakowski, 1972
 Ptomaphaginus baliensis Perreau, 1995
 Ptomaphaginus bengalicola Perreau, 1991
 Ptomaphaginus bryanti Jeannel, 1936
 Ptomaphaginus bryantioides Schilthuizen & Perreau, 2008
 Ptomaphaginus bucculentus Szymczakowski, 1974
 Ptomaphaginus burckhardti Schilthuizen & Perreau, 2008
 Ptomaphaginus caroli Schilthuizen & Perreau, 2008
 Ptomaphaginus chapmani Peck, 1981 
 Ptomaphaginus cherrapunjeensis Perreau, 1991
 Ptomaphaginus cilipes (Portevin, 1907)
 Ptomaphaginus clibanarius Szymczakowski, 1970
 Ptomaphaginus coronatus Szymczakowski, 1974
 Ptomaphaginus dao Perkovsky, 2001
 Ptomaphaginus depequkri Perreau, 1986
 Ptomaphaginus flavicornis (Motschulsky, 1863)
 Ptomaphaginus fornicatus Szymczakowski, 1972
 Ptomaphaginus franki Perreau, 1993
 Ptomaphaginus geigenmuellerae Perreau, 1993
 Ptomaphaginus giachinoi Schilthuizen & Perreau, 2008
 Ptomaphaginus gibberosus Szymczakowski, 1972
 Ptomaphaginus giganteus Szymczakowski, 1980
 Ptomaphaginus gonbe Nishikawa & Hayashi, 2016
 Ptomaphaginus gracilis Schweiger, 1956
 Ptomaphaginus grandis Schilthuizen & Perreau, 2018
 Ptomaphaginus guangxiensis Wang & Zhou, 2015
 Ptomaphaginus gutianshanicus Wang & Zhou, 2015
 Ptomaphaginus hamatus Szymczakowski, 1975
 Ptomaphaginus hanskii Schilthuizen & Perreau, 2018
 Ptomaphaginus heterotrichus Szymczakowski, 1975
 Ptomaphaginus honestus Szymczakowski, 1964
 Ptomaphaginus isabellarosselini  Schilthuizen, Njunjić & Perreau, 2018
 Ptomaphaginus ishizuchiensis Perreau, 1996
 Ptomaphaginus jacobsoni Szymczakowski, 1964
 Ptomaphaginus kinabaluensis Schilthuizen & Perreau, 2008
 Ptomaphaginus kinabatanganensis   Njunjić, Schilthuizen & Perreau, 2018
 Ptomaphaginus kosiensis Perreau, 1988
 Ptomaphaginus kurbatovi Schilthuizen & Perreau, 2008
 Ptomaphaginus lacertosus Szymczakowski, 1970
 Ptomaphaginus laticornis Jeannel, 1936
 Ptomaphaginus latimanus Schilthuizen & Perreau, 2008
 Ptomaphaginus latipes (Pic, 1929)
 Ptomaphaginus layangensis Schilthuizen & Perreau, 2018
 Ptomaphaginus leucodon Szymczakowski, 1975
 Ptomaphaginus lipsae Perreau & Lemaire, 2018
 Ptomaphaginus loeblianus Schilthuizen & Perreau, 2008
 Ptomaphaginus longitarsis Portevin, 1914
 Ptomaphaginus louis Schilthuizen & Perreau, 2018
 Ptomaphaginus luoi Wang & Zhou, 2015
 Ptomaphaginus major Szymczakowski, 1972
 Ptomaphaginus marshalli Schilthuizen & Perreau, 2018
 Ptomaphaginus megalayanus Perreau, 1991
 Ptomaphaginus microphallus Schilthuizen & Perreau, 2018
 Ptomaphaginus minimus Perreau, 1998
 Ptomaphaginus mirabilis Szymczakowski, 1972
 Ptomaphaginus miyataorum Nishikawa, 2017
 Ptomaphaginus muluensis Schilthuizen & Perreau, 2018
 Ptomaphaginus murphyi Szymczakowski, 1970
 Ptomaphaginus nanus Schilthuizen & Perreau, 2018
 Ptomaphaginus newtoni Wang & Zhou, 2015
 Ptomaphaginus nipponensis Perreau, 1996
 Ptomaphaginus nitens Jeannel, 1936
 Ptomaphaginus obtusus Szymczakowski, 1959
 Ptomaphaginus okinawaensis Hoshina & Sugaya, 2003
 Ptomaphaginus oribates Szymczakowski, 1965
 Ptomaphaginus pallidicornis (Portevin, 1907)
 Ptomaphaginus palpalis Szymczakowski, 1974
 Ptomaphaginus palpaloides Perreau, 1988
 Ptomaphaginus pecki Perkovsky, 1995
 Ptomaphaginus perreaui Wang & Zhou, 2015
 Ptomaphaginus pilipennis Perreau, 1991
 Ptomaphaginus pilipennoides Perreau, 1991
 Ptomaphaginus pingtungensis Perreau, 1996
 Ptomaphaginus piraster Szymczakowski, 1975
 Ptomaphaginus portevini Szymczakowski, 1964
 Ptomaphaginus quadricalcarus Wang & Zhou, 2015
 Ptomaphaginus riedeli Perreau, 1995
 Ptomaphaginus rubidus (Champion, 1927)
 Ptomaphaginus rufus Jeannel, 1936
 Ptomaphaginus rugosus Perreau, 1991
 Ptomaphaginus ruzickai Wang & Zhou, 2015
 Ptomaphaginus sabahensis Schilthuizen & Perreau, 2008
 Ptomaphaginus sarawacensis Schilthuizen & Perreau, 2018
 Ptomaphaginus sauteri (Portevin, 1914)
 Ptomaphaginus scaber Szymczakowski, 1964
 Ptomaphaginus scaphaner Szymczakowski, 1972
 Ptomaphaginus schawalleri Perreau, 1992
 Ptomaphaginus shennongensis Wang & Zhou, 2015
 Ptomaphaginus shibatai Hayashi, 1969
 Ptomaphaginus similipes Schilthuizen & Perreau, 2008
 Ptomaphaginus similis Schweiger, 1956
 Ptomaphaginus sinuatus Schilthuizen, 1984
 Ptomaphaginus smetanai Perreau, 1988
 Ptomaphaginus takaosanus Nakane, 1982
 Ptomaphaginus takashii Nishikawa, 2017
 Ptomaphaginus tarsalis Szymczakowski, 1964
 Ptomaphaginus testaceus Schilthuizen & Perreau, 2018
 Ptomaphaginus thieleni Perreau, 1993
 Ptomaphaginus tomellerii Zoia, 1997
 Ptomaphaginus trautneri Perreau, 1993
 Ptomaphaginus troglodytes  Perreau & Růžička, 2018
 Ptomaphaginus truncatus Perreau, 1988
 Ptomaphaginus turensis Perreau, 1991
 Ptomaphaginus wenboi Wang & Zhou, 2015
 Ptomaphaginus wuzhishanicus Wang & Zhou, 2015
 Ptomaphaginus yaeyamaensis Hoshina & Sugaya, 2003
 Ptomaphaginus yui Wang & Zhou, 2015

Referanser

Kilder 
Leiodidae i Tree of life

Eksterne lenker 

 

Biller formelt beskrevet i 1914
Mycelbiller