Ptomaphagus

Ptomaphagus er en slekt av mycelbiller som særlig finnes knyttet til sopp og åtsler.

Utseende 
Små  (2-4 millimeter), halvmatte, mørkbrunlige mycelbiller, kroppsformen nærmest dråpeformet med lange bein. Antennene er litt kølleformede og 11-leddete. De fem ytterste leddene danner en avlang klubbe, men det åttende leddet er mye mindre enn de andre og er vanligvis forkortet til en liten skive. Hodet er oftest kraftig innskåret bak fasettøynene, med en smal hals, men dette kan være vanskelig å se da billene gjerne trekker hodet inn under brystskjoldet. Hannen har utvidete føtter på forbeina.

Levevis 
Artene kan finnes på små åtsler, mellom råtnende plantemateriale og av og til i musereir.

Systematisk inndeling 

 Ordenen biller, Coleoptera
 Underordenen Polyphaga
 Overfamilien åtsel- og rovbiller, Staphylinoidea
 Familien mycelbiller, Leiodidae Fleming, 1821
 Underfamilien kadaverbiller, Cholevinae
 Stammen Ptomaphagini
 Slekten Ptomaphagus Hellwig, 1795  – tre arter i Norge
 Ptomaphagus acuminatus Salgado, 2011
 Ptomaphagus acutus Peck and Gnaspini, 1997
 Ptomaphagus agilis (P.H.Lucas, 1847)
 Ptomaphagus alleghenyensis (Peck, 1979)
 Ptomaphagus altus Peck, 1973
 Ptomaphagus appalachianus (Peck, 1979)
 Ptomaphagus argenticornis Peck, 1977
 Ptomaphagus aritzensis Jeannel, 1934
 Ptomaphagus barbarae Peck, 1973
 Ptomaphagus barri Peck, 1973
 Ptomaphagus bedfordensis (Peck, 1979)
 Ptomaphagus biharicus Jeannel, 1934
 Ptomaphagus boliviensis Salgado, 2014
 Ptomaphagus bordoni (Jeannel, 1965)
 Ptomaphagus brevior Jeannel, 1949
 Ptomaphagus californicus (LeConte, 1854)
 Ptomaphagus caucasicus Jeannel, 1934
 Ptomaphagus cavernicola Schwarz, 1898
 Ptomaphagus championi Jeannel, 1936
 Ptomaphagus chendae J.Müller, 1921
 Ptomaphagus chenggongi Wang, Cheng-Bin, Nishikawa, Perreau, Růžička & Y.Hayashi, 2016
 Ptomaphagus chihuahuensis (Peck, 1976)
 Ptomaphagus chromolithus Peck, 1984
 Ptomaphagus circassicus (Reitter, 1888)
 Ptomaphagus clavalis (Reitter, 1884)
 Ptomaphagus cocytus Peck, 1973
 Ptomaphagus colima Peck, 1977
 Ptomaphagus conejera Peck, 1977
 Ptomaphagus consobrinus (LeConte, 1853)
 Ptomaphagus costanama Peck, 2003
 Ptomaphagus costarica Peck, 2003
 Ptomaphagus cubensis Salgado, 2012
 Ptomaphagus cumberlandus (Peck, 1979)
 Ptomaphagus dacicus Jeannel, 1934
 Ptomaphagus davisii Stephens, 1830
 Ptomaphagus delsur Peck, 1977
 Ptomaphagus densus Hatch, 1957
 Ptomaphagus distinctus Peck, 1977
 Ptomaphagus divaricatus Jeannel, 1934
 Ptomaphagus elabra Peck, 1971
 Ptomaphagus episcopus Peck, 1973
 Ptomaphagus fecundus (Barr, 1963)
 Ptomaphagus fiskei Peck, 1973
 Ptomaphagus fisus Horn, 1885
 Ptomaphagus flabellatus Szymczakowski, 1961
 Ptomaphagus forticornis A.Matthews, 1888
 Ptomaphagus fumosus (Peck, 1979)
 Ptomaphagus funiu Wang, Cheng-Bin, Perreau, Růžička & Nishikawa, 2017
 Ptomaphagus geomysi Peck and Skelley, 2002
 Ptomaphagus germari Schlechtendahl, 1888
 Ptomaphagus giaquintoi Jeannel, 1936
 Ptomaphagus gypsum Peck, 1973
 Ptomaphagus haba Wang, Cheng-Bin, Perreau, Růžička & Nishikawa, 2017
 Ptomaphagus hastatellus Szymczakowski, 1976
 Ptomaphagus hatchi Jeannel, 1933
 Ptomaphagus hazelae Peck, 1973
 Ptomaphagus hayashii Wang, 2016
 Ptomaphagus hirtus (Tellkampf, 1844)
 Ptomaphagus hispaniolensis Peck & Cook, 2014
 Ptomaphagus hubrichti Barr, 1958
 Ptomaphagus inbio Peck, 2003
 Ptomaphagus inyoensis Peck and Gnaspini, 1997
 Ptomaphagus izabal Perreau, 2013
 Ptomaphagus jalisco Peck, 1977
 Ptomaphagus jamesi Peck, 1973
 Ptomaphagus joanna (Peck, 1979)
 Ptomaphagus jonesi (Peck, 1979)
 Ptomaphagus julius Peck, 1973
 Ptomaphagus kuntzeni Sokolowski, 1957
 Ptomaphagus laselva Peck, 1977
 Ptomaphagus laticornis Jeannel, 1949
 Ptomaphagus leo Peck, 1973
 Ptomaphagus lincolnensis Peck, 1978
 Ptomaphagus liquidambar Peck, 1977
 Ptomaphagus loedingi Hatch, 1933
 Ptomaphagus longicornis Jeannel, 1949
 Ptomaphagus manzano Peck, 1978
 Ptomaphagus masumotoi Nishikawa, 2011
 Ptomaphagus mckenziei Peck, 1977
 Ptomaphagus medius (Rey, 1889)  - finnes i Norge
 Ptomaphagus merritti Tishechkin, 2007
 Ptomaphagus meximontanus Peck, 1973
 Ptomaphagus microps Peck, 1978
 Ptomaphagus mitchellensis (Hatch, 1933)
 Ptomaphagus nashvillensis (Peck, 1979)
 Ptomaphagus neojonesi Peck & Newton, 2017
 Ptomaphagus nepalensis Perreau, 1988
 Ptomaphagus nevadicus Horn, 1880
 Ptomaphagus newtoni Peck, 1973
 Ptomaphagus nubloso Peck, 2003
 Ptomaphagus oaxaca Peck, 1973
 Ptomaphagus orichalcum (Peck, 1979)
 Ptomaphagus oriental Peck, 1977
 Ptomaphagus parashant Peck and Wynne, 2013
 Ptomaphagus pecki Gnaspini, 1996
 Ptomaphagus piccoloi Wang, Cheng-Bin, Růžička, Nishikawa, Perreau & Y.Hayashi, 2016
 Ptomaphagus piperi Hatch, 1933
 Ptomaphagus pisgahensis (Peck, 1979)
 Ptomaphagus pius (Seidlitz, 1887)
 Ptomaphagus pruina Peck, 1977
 Ptomaphagus pruinosus (Reitter, 1881)
 Ptomaphagus puertoricensis Peck, 1972
 Ptomaphagus pyrenaeus Jeannel, 1934
 Ptomaphagus quercus Peck, 1977
 Ptomaphagus reddelli Peck, 1973
 Ptomaphagus rhagianicus Sbordoni, 1967
 Ptomaphagus richlandensis (Peck, 1979)
 Ptomaphagus ruficollis Portevin, 1903
 Ptomaphagus sardus (Seidlitz, 1887)
 Ptomaphagus schwarzi Hatch, 1933
 Ptomaphagus sciakyi (Zoia, 1993)
 Ptomaphagus scottsboroensis (Peck, 1979)
 Ptomaphagus sericatus (Chaudoir, 1845) - finnes i Norge
 Ptomaphagus setiger (Horn, 1885)
 Ptomaphagus shapardi Sanderson, 1939
 Ptomaphagus sibiricus Jeannel, 1934
 Ptomaphagus solanum Peck, 1973
 Ptomaphagus sonorensis (Peck, 1976)
 Ptomaphagus spelaeus (Bilimek, 1867)
 Ptomaphagus steevesi (Peck, 1979)
 Ptomaphagus subtruncatus Mäklin, 1881
 Ptomaphagus subvillosus (Goeze, 1777) - finnes i Norge
 Ptomaphagus suteri (Peck, 1979)
 Ptomaphagus szymczakowskii Peck, 1977
 Ptomaphagus tabascensis Sbordoni, 1974
 Ptomaphagus talamanca Peck, 1973
 Ptomaphagus tauricus Jeannel, 1934
 Ptomaphagus tenuicornis (Rosenhauer, 1856)
 Ptomaphagus texanus Melander, 1902
 Ptomaphagus thebeatles Schilthuizen et al., 2020
 Ptomaphagus thomomysi Hatch, 1957
 Ptomaphagus tierrabaja Peck, 1977
 Ptomaphagus tingtingae Wang, Cheng-Bin, Nishikawa, Perreau, Růžička & Y.Hayashi, 2016
 Ptomaphagus torodei Peck, 1984
 Ptomaphagus trezzii Perreau, 2009
 Ptomaphagus troglodytes Blas & Vives, 1983
 Ptomaphagus troglomexicanus Peck, 1968
 Ptomaphagus tuza Peck, 1977
 Ptomaphagus ulkei Horn, 1885
 Ptomaphagus valentinei Jeannel, 1933
 Ptomaphagus validus (Kraatz, 1852)
 Ptomaphagus vallombrosae (Seidlitz, 1887)
 Ptomaphagus varicornis (rosenhauer, 1847) - finnes i Norge
 Ptomaphagus volcanica Peck, 1977
 Ptomaphagus walteri Peck, 1973
 Ptomaphagus whiteselli Barr, 1963
 Ptomaphagus yasutoshii Nishikawa, 1993
 Ptomaphagus yuvila Peck, 1977

Referanser

Kilder 
 Leiodidae i Tree of life

 Silfverberg, H. 1992. Enumeratio Coleopterorum Fennoscandiae, Daniae et Baltiae (liste over Nordens biller). Helsinki.

Eksterne lenker 
 Artsdatabanken - Ptomaphagus 
 Ptomapahgus på GBIF 
 

Biller formelt beskrevet i 1795
Mycelbiller