Brachinus

se artsliste

Brachinus  er en slekt av billene i delgruppen (underfamilien) bombardérbiller (Brachininae) i familiegruppen løpebiller. Ingen arter er registret fra Norge.

Utseende 
Små til middelsstore (gjerne 5-10 millimeter), spinkle løpebiller, dekkvingene forholdsvis brede. På farge er dekkvingene ofest mørke, ofte med metallglans, hode, pronotum og bein ofte rødlige. Hodet er omtrent femkantet, fasettøynene forholdsvis små, antennene lange og tynne. Pronotum er omtrent så bredt som hodet, bredere foran enn bak, med innsvungne kanter. Dekkvingene er noe flate, oftest med fine punktstriper.

Levevis 
Artene lever på varme, tørre, solåpne steder. De voksne billene er rovdyr og kan noen ganger finnes tallrikt under steiner. Larvene er ektoparasitter, i alle fall noen av artene angriper pupper av andre biller.
Billene i denne slekten forsvarer seg ved å sprute kokvarme kjemikalier mot angripere, dette kan de gjøre med stor presisjon.

Utbredelse 
Slekten har en holarktisk utbredelse, inkludert det meste av Europa, men er ikke funnet i Norge. Det er tallrike arter i Middelhavsområdet.

Systematisk inndeling 

 Ordenen biller, Coleoptera Linnaeus, 1758
 Underordenen Adephaga Schellenberg, 1806
 Overfamilien løpebiller og vannkalver, Caraboidea Latreille, 1802
 Familien løpebiller, Carabidae Latreille, 1802
 Underfamilien bombardérbiller, Brachininae Bonelli, 1810
 Slekten Brachinus Weber, 1801  – tallrike arter i det sørlige Europa

Liste over artene 
 Brachinus aabaaba Erwin, 1970
 Brachinus abbreviatus (Castelnau, 1835)
 Brachinus abyssinicus Chaudoir, 1876
 Brachinus adustipennis Erwin, 1969
 Brachinus aeger Chaudoir, 1876
 Brachinus aeneicostis Bates, 1883
 Brachinus afghanus Jedlicka, 1967
 Brachinus agraphus (Alluaud, 1899)
 Brachinus albarracinus Wagner, 1926
 Brachinus alexandri F. Battoni, 1984
 Brachinus alexiguus Erwin, 1970
 Brachinus algoensis Peringuey, 1896
 Brachinus alternans Dejean, 1825
 Brachinus americanus (Leconte, 1844)
 Brachinus andalusiacus Rambur, 1837
 Brachinus andreaei Basilewsky, 1964
 Brachinus angustatus Dejean, 1831
 Brachinus ankarensis Jedlicka, 1962
 Brachinus annulicornis Chaudoir, 1842
 Brachinus apicalis Erichson, 1843
 Brachinus arboreus Chevrolat, 1834
 Brachinus aristokrates Liebke, 1934
 Brachinus armiger Dejean, 1831
 Brachinus atramentarius Mannerheim, 1837
 Brachinus atripennis Ballion, 1871
 Brachinus azureipennis Chaudoir, 1876
 Brachinus baeticus Rambur, 1837
 Brachinus bagdatensis Pic, 1902
 Brachinus barbarus Lucas, 1846
 Brachinus bayardi Dejean, 1831
 Brachinus bellicosus L. Dufour, 1820
 Brachinus berytensis Reiche & Saulcy, 1855
 Brachinus bigutticeps Chaudoir, 1876
 Brachinus bilineatus Castelnau, 1835
 Brachinus bipustulatus Quensel, 1806
 Brachinus bodemeyeri Apfelbeck, 1904
 Brachinus brevicollis Motschulsky, 1844
 Brachinus brittoni Ali, 1967
 Brachinus bruchi Liebke, 1939
 Brachinus brunneus Castelnau, 1834
 Brachinus brunniventris Motschulsky, 1864
 Brachinus caffer Boheman, 1848
 Brachinus canaliculatus Fairmaire, 1897
 Brachinus capnicus Erwin, 1970
 Brachinus catalonicus Jeanne, 1988
 Brachinus chalchihuitlicue Erwin, 1970
 Brachinus charis Andrewes, 1923
 Brachinus chaudoirianus Jakobson, 1908
 Brachinus chinensis Chaudoir, 1850
 Brachinus chirriador Erwin, 1970
 Brachinus chuji Jedlicka, 1956
 Brachinus cibolensis Erwin, 1970
 Brachinus ciliatus Liebke, 1933
 Brachinus cinctellus Chaudoir, 1876
 Brachinus cinctipennis Chevrolat, 1835
 Brachinus circumtinctus Bates, 1892
 Brachinus collarti Basilewsky, 1948
 Brachinus conformis Dejean, 1831
 Brachinus congicus (Basilewsky, 1958)
 Brachinus connectoides (Jeannel, 1949)
 Brachinus connectus Dejean, 1831
 Brachinus consanguineus Chaudoir, 1876
 Brachinus constrictus Reitter, 1919
 Brachinus cordicollis Dejean, 1826
 Brachinus costatulus Quensel, 1806
 Brachinus costiger Chaudoir, 1876
 Brachinus costipennis Motschulsky, 1859
 Brachinus costulipennis Liebke, 1928
 Brachinus crepitans Linne, 1758  – kjent fra Sverige, Finland og Danmark
 Brachinus cruciatus Quensel, 1806
 Brachinus cyanipennis Say, 1823
 Brachinus cyanochroaticus Erwin, 1969
 Brachinus daliensis Kirschenhofer, 2011
 Brachinus dawnaensis Kirschenhofer, 2003
 Brachinus demoulini Basilewsky, 1962
 Brachinus dieganus (Jeannel, 1949)
 Brachinus diffusus Chaudoir, 1876
 Brachinus dilapsus Antoine, 1941
 Brachinus dilatatus Klug, 1832
 Brachinus dilatipennis Reitter, 1919
 Brachinus dilottii Maran, 1933
 Brachinus dissimilis (Basilewsky, 1943)
 Brachinus distans Lorenz, 1998
 Brachinus dorsalis Dejean, 1831
 Brachinus dryas Andrewes, 1936
 Brachinus dryas Andrewes, 1936
 Brachinus efflans Dejean, 1830
 Brachinus ejaculans Fischer von Waldheim, 1828
 Brachinus elegans Chaudoir, 1842
 Brachinus elegantulus Erichson, 1842
 Brachinus elongatulus Chaudoir, 1876
 Brachinus enonensis Peringuey, 1899
 Brachinus eucosmus Andrewes, 1937
 Brachinus evanescens Bates, 1892
 Brachinus exhalans P. Rossi, 1792
 Brachinus explodens Duftschmid, 1812
 Brachinus explosus Erwin, 1970
 Brachinus exquisitus Bates, 1892
 Brachinus fageli Basilewsky, 1962
 Brachinus fasciatocollis (Fairmaire, 1901)
 Brachinus favicollis Erwin, 1965
 Brachinus femoralis Gistel, 1857
 Brachinus flavicapillus Bates, 1892
 Brachinus flaviventris Chaudoir, 1876
 Brachinus flavus Jedlicka, 1955
 Brachinus foochowi Kirschenhofer, 1986
 Brachinus formosanus Jedlicka, 1939
 Brachinus forticostis Liebke, 1934
 Brachinus frontalis Chaudoir In Oberthur, 1883
 Brachinus fulminatus Erwin, 1969
 Brachinus fulvipennis Chaudoir, 1876
 Brachinus fumans (Fabricius, 1781)
 Brachinus fumator Gistel, 1857
 Brachinus fuscicornis Dejean, 1826
 Brachinus fuscipennis Dejean, 1825
 Brachinus galactoderus Erwin, 1970
 Brachinus gebhardis Erwin, 1965
 Brachinus genicularis Mannerheim, 1837
 Brachinus geniculatus Dejean, 1831
 Brachinus gentilis Erichson, 1843
 Brachinus ghindanus Liebke, 1934
 Brachinus grandis Brulle, 1838
 Brachinus hamatus Fischer von Waldheim, 1828
 Brachinus hazardi Andrewes, 1930
 Brachinus hexagrammus Chaudoir, 1876
 Brachinus hirsutus Bates, 1884
 Brachinus hoffmanni Liebke, 1927
 Brachinus humeralis Ahrens, 1812
 Brachinus hylaenus Reichardt, 1967
 Brachinus ichabodopsis Erwin, 1970
 Brachinus illotus Chaudoir, 1876
 Brachinus immaculicornis Dejean, 1826
 Brachinus immarginatus Brulle, 1838
 Brachinus imperialensis Erwin, 1965
 Brachinus imporcitis Erwin, 1970
 Brachinus incomptus Bates, 1873
 Brachinus inconditus Peringuey, 1896
 Brachinus inops Andrewes, 1932
 Brachinus intactus Bates, 1892
 Brachinus intermedius Brulle, 1838
 Brachinus irakus Liebke, 1933
 Brachinus italicus Dejean, 1831
 Brachinus janthinipennis (Dejean, 1831)
 Brachinus javalinopsis Erwin, 1970
 Brachinus jucundus Dejean, 1831
 Brachinus kalalovae Roubal, 1932
 Brachinus kansanus Leconte, 1862
 Brachinus kavanaughi Erwin, 1969
 Brachinus knirschi Jedlicka, 1931
 Brachinus kollari Liebke, 1934
 Brachinus krynickii Jedlick
 Brachinus kryzhanovskii Belousov & Kabak, 1992
 Brachinus laetus Dejean, 1831
 Brachinus laevicostis Liebke, 1934
 Brachinus langenhani Liebke, 1934
 Brachinus lapidarius Basilewsky, 1962
 Brachinus lateralis Dejean, 1831
 Brachinus latipennis Peyerimhoff, 1907
 Brachinus latus Lorenz, 1998
 Brachinus lavaudeni (Jeannel, 1949)
 Brachinus leprieuri Gory, 1833
 Brachinus lesnei Andrewes, 1923
 Brachinus lethierryi Reiche, 1868
 Brachinus lewecki Liebke, 1928
 Brachinus lewisii Bates, 1873
 Brachinus limbellus Chaudoir, 1876
 Brachinus limbicollis Chaudoir, 1876
 Brachinus limbiger Chaudoir, 1876
 Brachinus lombokensis Kirschenhofer, 2010 ID
 Brachinus longipalpis Wiedemann, 1821
 Brachinus longulus Chaudoir, 1876
 Brachinus luzonicus Chaudoir, 1876
 Brachinus macrocerus Chaudoir, 1876
 Brachinus mactus Peringuey, 1904
 Brachinus madecassus (Jeannel, 1949)
 Brachinus magyari Jedlicka, 1960
 Brachinus marginellus Dejean, 1826
 Brachinus marginipennis Motschulsky, 1864
 Brachinus marginiventris Brulle, 1838
 Brachinus marinus Motschulsky, 1845
 Brachinus marleyi Barker, 1919
 Brachinus maublanci (Colas, 1942) F
 Brachinus mauretanicus Bedel, 1914
 Brachinus medius Harris, 1828
 Brachinus melanarthrus Chaudoir, 1876
 Brachinus melancholicus Schmidt-Goebel, 1846
 Brachinus merkli Kirschenhofer, 2003
 Brachinus mesopotamicus Ali, 1967
 Brachinus methneri Liebke, 1934
 Brachinus mexicanus Dejean, 1831
 Brachinus micheli (Jeannel, 1949)
 Brachinus microamericanus Erwin, 1969
 Brachinus microrrhabdus (Alluaud, 1899)
 Brachinus midoli Alluaud, 1917
 Brachinus minor Jedlicka, 1958
 Brachinus mobilis Erwin, 1970
 Brachinus modestus Schmidt-Goebel, 1846
 Brachinus motschulskyi Jedlick
 Brachinus muchei Jedlicka, 1967
 Brachinus natalicus Peringuey, 1896
 Brachinus neglectus Leconte, 1844
 Brachinus niger Chaudoir, 1876
 Brachinus nigricans Chaudoir, 1850
 Brachinus nigricornis Gebler, 1829
 Brachinus nigridorsis Nakane, 1962
 Brachinus nigripes G.R.Waterhouse, 1841
 Brachinus nigrovirescens Basilewsky, 1951
 Brachinus nobilis Dejean, 1831
 Brachinus nuristanus Jedlicka, 1967
 Brachinus oaxacensis Erwin, 1970
 Brachinus obliquetruncatus Peris, 1875
 Brachinus obliterus Peringuey, 1896
 Brachinus oblongus Dejean, 1825
 Brachinus obscuripennis Motschulsky, 1864
 Brachinus obscuritarsis Motschulsky, 1864
 Brachinus obtusus (Thunberg, 1784)
 Brachinus olgae Arribas, 1993
 Brachinus olidus Reiche, 1842
 Brachinus oneili Peringuey, 1899
 Brachinus opacipennis Motschulsky, 1864
 Brachinus orestes Kirschenhofer, 2003
 Brachinus orestes Kirschenhofer, 2003
 Brachinus orientalis Chaudoir, 1876
 Brachinus ornatus Fairmaire, 1901
 Brachinus otini Antoine, 1963
 Brachinus ovipennis LeConte, 1862
 Brachinus oxygonus Chaudoir, 1843
 Brachinus pachygaster Perty, 1830
 Brachinus pallidipes Reitter, 1919
 Brachinus pallidus Erwin, 1965
 Brachinus pallipes Dejean, 1826
 Brachinus papua Darlington, 1968
 Brachinus pateri Puel, 1938
 Brachinus patruelis Leconte, 1844
 Brachinus paviei Lesne, 1896
 Brachinus pecoudi Puel, 1925
 Brachinus pectoralis Dejean, 1825
 Brachinus peltastes Andrewes, 1931
 Brachinus perplexus Dejean, 1831
 Brachinus perrieri (Jeannel, 1949)
 Brachinus phaeocerus Chaudoir, 1868
 Brachinus philippinensis Tian & Deuve, 2007
 Brachinus piceus Chaudoir, 1876
 Brachinus pictus Hope, 1833
 Brachinus plagiatus Reiche, 1868
 Brachinus planiusculus Motschulsky, 1864
 Brachinus posticus Dejean, 1831
 Brachinus praestans Andrewes, 1931
 Brachinus promontorii Peringuey, 1888
 Brachinus proximus Fairmaire, 1887
 Brachinus pseudocruciatus Reitter, 1909
 Brachinus psophia Audinet-Serville, 1821
 Brachinus puberulus Chaudoir, 1868
 Brachinus punctaticollis Heller, 1923
 Brachinus puncticollis Schmidt-Goebel, 1846
 Brachinus pygmaeus Dejean, 1826
 Brachinus quadriguttatus Gebler, 1829
 Brachinus quadripennis Dejean, 1825
 Brachinus reyi Andrewes, 1924
 Brachinus rikatlae Peringuey, 1896
 Brachinus rugipennis Chaudoir, 1868
 Brachinus sallei Chaudoir, 1876
 Brachinus sanctus Motschulsky, 1864
 Brachinus schmidti Andrewes, 1927
 Brachinus scitulus Schmidt-Goebel, 1846
 Brachinus sclopeta Fabricius, 1792
 Brachinus scotomedes L. Redtenbacher, 1867
 Brachinus scotti Liebke, 1934
 Brachinus scriptus Chaudoir, 1878
 Brachinus scutellatus Chaudoir, 1876
 Brachinus sericeus Dejean, 1831
 Brachinus servillei Marc, 1839
 Brachinus sexmaculatus Dejean, 1825
 Brachinus sexnotatus Liebke, 1934
 Brachinus sexpustulatus (Fabricius, 1775)
 Brachinus seyrigi (Alluaud, 1935)
 Brachinus sibiricus Motschulsky, 1864
 Brachinus sicardi Jeannel, 1949
 Brachinus sichemita Reiche & Saulcy, 1855
 Brachinus simulans Peringuey, 1896
 Brachinus solidipalpis Tian & Deuve, 2007
 Brachinus solidus Peringuey, 1899
 Brachinus somereni Burgeon, 1947
 Brachinus sonorous Erwin, 1970
 Brachinus sordidus Andrewes, 1933
 Brachinus stappersi Liebke, 1934
 Brachinus stenoderus Bates, 1873
 Brachinus stevensi Andrewes, 1924
 Brachinus stevensianus Jedlicka, 1956
 Brachinus strictus Morvan, 1971
 Brachinus stygius Andrewes, 1933
 Brachinus subcostatus Dejean, 1825
 Brachinus suberbianus Alluaud, 1935
 Brachinus sublaevis Chaudoir, 1868
 Brachinus subnotatus Chaudoir, 1844
 Brachinus suensoni Kirschenhofer, 1986
 Brachinus sulcipennis (Jeannel, 1949)
 Brachinus suturalis (Jeannel, 1949)
 Brachinus suturatus Chaudoir, 1876
 Brachinus suturellus Chaudoir, 1876
 Brachinus szetschuanensis Jedlicka, 1963
 Brachinus talyschensis Motschulsky, 1850
 Brachinus tenuicollis LeConte, 1844
 Brachinus tenuis Lorenz, 1998
 Brachinus testaceus Rambur, 1837
 Brachinus tetracolon Chaudoir, 1876
 Brachinus tetragrammus Chaudoir, 1876
 Brachinus tetraspilotus Chaudoir, 1876
 Brachinus tetrastigma Fairmaire, 1897
 Brachinus texanus Chaudoir, 1868
 Brachinus thieleni Kirschenhofer, 1987
 Brachinus tianshanicus Mikhailov, 1976
 Brachinus tigridis Ali, 1967
 Brachinus truncatulus Fairmaire, 1901
 Brachinus tsara Alluaud, 1918
 Brachinus turkestanicus Liebke, 1928
 Brachinus vadoni Jeannel, 1949
 Brachinus vagus Peringuey, 1899
 Brachinus variegatus (Roth, 1851)
 Brachinus variventris Schaufuss, 1862
 Brachinus velutinus Erwin, 1965
 Brachinus vicinus Dejean, 1826
 Brachinus vigilans Chaudoir, 1876
 Brachinus viridipennis Dejean, 1831
 Brachinus vitticollis Chaudoir, 1876
 Brachinus vulcanoides Erwin, 1969
 Brachinus xanthophryus Chaudoir, 1876
 Brachinus xanthopleurus Chaudoir, 1876
 Brachinus yunnanus Jedlicka, 1963

Litteratur 
 Bertram G. Rye. 1908. Biller I. Løpebiller. Danmarks Fauna Bind 3. G.E.C. GAD, København.

Eksterne lenker 
 
 Tree of Life – Bombardérbiller (Brachininae, Carabidae) – engelskspråklig
 Brachininae - fotogalleri - engelskspråklig
 Norsk Entomologisk forening

Bombardérbiller
Biller formelt beskrevet i 1801