Ataenius

Se systematisk inndeling

Ataenius er en artsrik slekt av biller som hører til familien skarabider (Scarabaeidae).

Utseende 
Små (1,5 -7 millimeter), avlange, blanke, mørke gjødselbiller. Hodet er stort og bredt, noe hvelvet, gjerne innbuktet foran.

Levevis 
Disse billene har larver som utvikler seg i jorda, der de etter på gressrøtter. De kan gjøre en del skade på golfbaner og lignende

Utbredelse 
Slekten er utbredt i alle verdensdeler bortsett fra Antarktis.

Systematisk inndeling 

 Ordenen biller, Coleoptera
 Underordenen Polyphaga
 Overfamilien skarabider, Scarabaeoidea
 Familien skarabider, (Scarabaeidae) Latreille, 1806
 Underfamilien Gjødselbiller, Aphodiinae Leach, 1815
 Stammen Eupariini Schmidt, 1910 
 Slekten Ataenius Harold, 1867

Liste over artene 
 Ataenius abancay Stebnicka, 2005
 Ataenius abditus (Haldeman, 1848)
 Ataenius abdominalis Petrovitz, 1973
 Ataenius aciculus Hinton, 1937
 Ataenius acutulus Schmidt, 1913
 Ataenius aequalis Harold, 1880
 Ataenius aequatorialis Petrovitz, 1961
 Ataenius africanus Endrödi, 1967
 Ataenius alternatus (Melsheimer, 1845)
 Ataenius ambaritae Stebnicka, 1988
 Ataenius angulatus Petrovitz, 1964
 Ataenius annanus Stebnicka & Howden, 1997
 Ataenius aphodiiformis Bordat & Théry, 2012
 Ataenius apicalis Hinton, 1937
 Ataenius arenosus Harold, 1868
 Ataenius argillaceus Schmidt, 1916
 Ataenius arrowi Hinton, 1936
 Ataenius asper Petrovitz, 1975
 Ataenius atramentarius (Erichson, 1847)
 Ataenius attenuator Harold, 1874
 Ataenius australasiae Boheman, 1858
 Ataenius australis Harold, 1875
 Ataenius balthasari Petrovitz, 1973
 Ataenius barberi Cartwright, 1974
 Ataenius basiceps Lea, 1923
 Ataenius beattyi Chapin, 1940
 Ataenius benarabyensis Stebnicka & Howden, 1997
 Ataenius bicolor Petrovitz, 1963
 Ataenius biroi Endrödi, 1951
 Ataenius bispinulosus Schmidt, 1911
 Ataenius blapoides Balthasar, 1947
 Ataenius bolivarensis Stebnicka, 2007
 Ataenius bolivari Stebnicka, 2001
 Ataenius borgmeieri Hinton, 1936
 Ataenius borjae Stebnicka, 2005
 Ataenius brevicollis (Wollaston, 1854)
 Ataenius brevis Fall, 1930
 Ataenius brevitarsis Petrovitz, 1964
 Ataenius brouni Sharp, 1876
 Ataenius brunneus Schmidt, 1922
 Ataenius buenavistae Stebnicka, 2001
 Ataenius caesoides Fairmaire, 1899
 Ataenius caicarae Stebnicka, 2005
 Ataenius californicus Horn, 1887
 Ataenius canoasus Stebnicka, 2007
 Ataenius carinatipennis Petrovitz, 1973
 Ataenius carinator Harold, 1874
 Ataenius cartago Stebnicka, 2007
 Ataenius cartwrighti Chalumeau & Gruner, 1974
 Ataenius castaniellus Bates, 1887
 Ataenius catarinaensis Stebnicka, 2007
 Ataenius catenulatus (Erichson, 1847)
 Ataenius cavagnaroi Cartwright, 1970
 Ataenius ceylonensis Schmidt, 1912
 Ataenius chapini Hinton, 1937
 Ataenius chilensis (Solier, 1851)
 Ataenius chinacotae Stebnicka, 2007
 Ataenius circulusbrumalis Petrovitz, 1964
 Ataenius clavatus Schmidt, 1916
 Ataenius clitellarius Petrovitz, 1973
 Ataenius cochabambae Stebnicka, 2005
 Ataenius cognatus (LeConte, 1858)
 Ataenius columbicus Harold, 1880
 Ataenius communis Hinton, 1936
 Ataenius complicatus Harold, 1869
 Ataenius confertus Fall, 1909
 Ataenius convexus Robinson, 1940
 Ataenius coriarius Schmidt, 1912
 Ataenius corporaali Boucomont, 1924
 Ataenius corrosus Chapin, 1940
 Ataenius costulifer Balthasar, 1951
 Ataenius crenaticollis Petrovitz, 1973
 Ataenius crenatipennis MacLeay, 1871
 Ataenius crenator Harold, 1876
 Ataenius crenatostriatus (Blanchard, 1851)
 Ataenius crenulatus Schmidt, 1910
 Ataenius cribrithorax Bates, 1887
 Ataenius cristobalensis Cook & Peck, 2000
 Ataenius cucutae Stebnicka, 2007
 Ataenius cylindrus Horn, 1871
 Ataenius dentatus Koshantshikov, 1916
 Ataenius depilis Petrovitz, 1976
 Ataenius deserti Blackburn, 1894
 Ataenius desertus Horn, 1871
 Ataenius ecruensis Stebnicka & Lago, 2005
 Ataenius edungalbae Stebnicka & Howden, 1997
 Ataenius elegans Harold, 1868
 Ataenius elisaensis Stebnicka, 2002
 Ataenius elongatulus MacLeay, 1888
 Ataenius elongatus (Palisot de Beauvois, 1811)
 Ataenius endroedyyoungai Endrödi, 1973
 Ataenius erinaceus Petrovitz, 1961
 Ataenius eringundae Stebnicka & Howden, 1997
 Ataenius erratus Fall, 1930
 Ataenius eurynotus Lea, 1923
 Ataenius excisicollis Petrovitz, 1961
 Ataenius exiguus Brown, 1932
 Ataenius fattigi Cartwright, 1948
 Ataenius floreanae Cook & Peck, 2000
 Ataenius forgiei Bordat & Théry, 2012
 Ataenius forsteri Balthasar, 1960
 Ataenius freyi Petrovitz, 1961
 Ataenius gammonensis Stebnicka & Howden, 1997
 Ataenius garamas Peyerimhoff, 1929
 Ataenius gascoyneensis Stebnicka & Howden, 1997
 Ataenius gibbus Blackburn, 1904
 Ataenius gilesi Stebnicka & Howden, 1997
 Ataenius glabriventris Schmidt, 1911
 Ataenius glaseri Cartwright, 1974
 Ataenius gracilis (Melsheimer, 1845)
 Ataenius granchacoensis Balthasar, 1939
 Ataenius granocostatus Schmidt, 1912
 Ataenius griffini Cartwright, 1974
 Ataenius gruneri Chalumeau, 1979
 Ataenius guanacastae Stebnicka, 2005
 Ataenius guayasi Stebnicka, 2001
 Ataenius gungareei Stebnicka & Howden, 1997
 Ataenius guriensis Stebnicka, 2005
 Ataenius heinekeni (Wollaston, 1854)
 Ataenius hesperius Cartwright, 1974
 Ataenius hirsutus Horn, 1871
 Ataenius hispaniolae Chalumeau, 1982
 Ataenius hispidus Harold, 1867
 Ataenius holopubescens Hinton, 1938
 Ataenius horticola Harold, 1869
 Ataenius howdeni Chalumeau, 1978
 Ataenius hrubanti Balthasar, 1967
 Ataenius huanus Stebnicka, 2007
 Ataenius humptydooensis Stebnicka & Howden, 1997
 Ataenius hygrophilus Paulian, 1947
 Ataenius icanus Balthasar, 1941
 Ataenius illaetabilis Lea, 1923
 Ataenius imbricatoides Schmidt, 1909
 Ataenius imbricatus (Melsheimer, 1845)
 Ataenius imparilis Blackburn, 1904
 Ataenius impiger Schmidt, 1916
 Ataenius impressus (Petrovitz, 1963)
 Ataenius incanus Balthasar, 1951
 Ataenius indutus Petrovitz, 1961
 Ataenius inquisitus Horn, 1887
 Ataenius insculptus Horn, 1887
 Ataenius insolitus Schmidt, 1909
 Ataenius insulae Chalumeau & Gruner, 1974
 Ataenius insularis Lea, 1923
 Ataenius integer Harold, 1868
 Ataenius integricollis Lea, 1923
 Ataenius intermedius Bates, 1887
 Ataenius iquitosae Stebnicka, 2007
 Ataenius isabelae Franz, 1985
 Ataenius jamaicensis Chapin, 1940
 Ataenius jardinensis Stebnicka, 2002
 Ataenius jelineki Chalumeau, 1982
 Ataenius kapalgaensis Stebnicka & Howden, 1997
 Ataenius klapperichi Howden, 1978
 Ataenius kochi Balthasar, 1941
 Ataenius koebelei Blackburn, 1904
 Ataenius koelleri Balthasar, 1963
 Ataenius koghianus Paulian, 1991
 Ataenius lamarensis Stebnicka, 2007
 Ataenius lamotteiroyi Bordat, 1992
 Ataenius lanei Petrovitz, 1973
 Ataenius languidus Schmidt, 1911
 Ataenius latus Petrovitz, 1963
 Ataenius lenkoi Petrovitz, 1973
 Ataenius liogaster Bates, 1887
 Ataenius lobatus Horn, 1871
 Ataenius londrinae Stebnicka, 2007
 Ataenius longiclavus Petrovitz, 1970
 Ataenius lucanus Horn, 1871
 Ataenius luctuosus (Burmeister, 1877)
 Ataenius luteomargo Chapin, 1940
 Ataenius madagassicus Bordat, 1990
 Ataenius maghribinicus Baraud, 1985
 Ataenius martinezi Petrovitz, 1973
 Ataenius miamii Cartwright, 1934
 Ataenius michelii Chalumeau, 1978
 Ataenius microtrichopterus Lea, 1923
 Ataenius millstreamae Stebnicka & Howden, 1997
 Ataenius montanus Schmidt, 1911
 Ataenius monteithi Paulian, 1991
 Ataenius monticola Paulian, 1940
 Ataenius montreuili Bordat & Théry, 2012
 Ataenius morator Harold, 1869
 Ataenius murchisoni Stebnicka & Howden, 1997
 Ataenius nakpandurii Endrödi, 1973
 Ataenius nanus (DeGeer, 1774)
 Ataenius napoensis Stebnicka, 2007
 Ataenius nigricans Paulian, 1933
 Ataenius noques Stebnicka, 2007
 Ataenius noronhai Stebnicka, 2007
 Ataenius nudus Blackburn, 1904
 Ataenius nugator Harold, 1880
 Ataenius oaxacaensis Stebnicka & Lago, 2005
 Ataenius ocumarensis Stebnicka, 2001
 Ataenius oklahomensis Brown, 1930
 Ataenius onkonensis Stebnicka, 2005
 Ataenius opacipennis Schmidt, 1910
 Ataenius opatrinus Harold, 1867
 Ataenius opatroides (Balthasar, 1947)
 Ataenius ovatulus Horn, 1871
 Ataenius pacificus (Sharp, 1879)
 Ataenius palmaritoensis Stebnicka, 2007
 Ataenius palmerstoni Blackburn, 1891
 Ataenius palustris (Montrouzier, 1860)
 Ataenius panamensis Hinton, 1936
 Ataenius parallelipennis Petrovitz, 1973
 Ataenius parvus Lea, 1923
 Ataenius pearlensis Stebnicka, 2006
 Ataenius peregianensis Stebnicka & Howden, 1997
 Ataenius peregrinator Harold, 1877
 Ataenius pereirai Petrovitz, 1970
 Ataenius perforatus Harold, 1867
 Ataenius pertuga Balthasar, 1961
 Ataenius petrovitzi Balthasar, 1960
 Ataenius picinus Harold, 1867
 Ataenius platensis (Blanchard, 1847)
 Ataenius plaumanni Petrovitz, 1973
 Ataenius polyglyptus Bates, 1887
 Ataenius pseudimparilis Stebnicka & Howden, 1997
 Ataenius pseudocarinator Balthasar, 1947
 Ataenius pseudoclavatus Stebnicka, 2005
 Ataenius pseudocommunis Stebnicka, 2001
 Ataenius pseudostercorator Stebnicka, 2003
 Ataenius pseudousingeri Galante, Stebnicka & Verdú, 2003
 Ataenius punctatohirsutus Schmidt, 1909
 Ataenius puncticollis LeConte, 1868
 Ataenius punctipennis Harold, 1868
 Ataenius purator Harold, 1868
 Ataenius quintanaroo Stebnicka, 2006
 Ataenius raccurti Chalumeau, 1978
 Ataenius rakovici Bordat & Théry, 2012
 Ataenius raslani Bordat & Théry, 2012
 Ataenius raucus Schmidt, 1908
 Ataenius robustus Horn, 1871
 Ataenius rosinae Balthasar, 1967
 Ataenius rubrotessellatus (Blanchard, 1843)
 Ataenius sabinoi Cartwright, 1974
 Ataenius saltae Stebnicka, 2007
 Ataenius santarosae Stebnicka, 2007
 Ataenius saramari Cartwright, 1939
 Ataenius saulensis Stebnicka, 2006
 Ataenius scabrelloides Petrovitz, 1962
 Ataenius scabrellus Schmidt, 1909
 Ataenius scalptifrons Bates, 1887
 Ataenius schmidti Stebnicka, 2003
 Ataenius sculptilis Harold, 1868
 Ataenius sculptor Harold, 1868
 Ataenius scutellaris Harold, 1867
 Ataenius seaforthensis Stebnicka & Howden, 1997
 Ataenius semicoecus MacLeay, 1888
 Ataenius semicornutus MacLeay, 1871
 Ataenius semipilosus Van Dyke, 1928
 Ataenius setiger Bates, 1887
 Ataenius setosus Schmidt, 1909
 Ataenius seydeli Endrödi, 1964
 Ataenius siminasus Petrovitz, 1973
 Ataenius simplicipes Mulsant & Rey, 1870
 Ataenius skelleyi Stebnicka, 2007
 Ataenius sparsicollis Blackburn, 1904
 Ataenius speculator Blackburn, 1891
 Ataenius spinipennis Lea, 1923
 Ataenius spretulus (Haldeman, 1848)
 Ataenius stebnickae Bordat & Théry, 2012
 Ataenius steinheili Harold, 1874
 Ataenius stephani Cartwright, 1974
 Ataenius stercorator (Fabricius, 1775)
 Ataenius striatocrenatus Fairmaire, 1889
 Ataenius strigatus (Say, 1823)
 Ataenius strigicaudus Bates, 1887
 Ataenius strigifrons Schmidt, 1920
 Ataenius suineata Deloya, 2012
 Ataenius synnotensis Stebnicka & Howden, 1997
 Ataenius talpoides Stebnicka, 2001
 Ataenius tambopatae Stebnicka, 2001
 Ataenius tarumensis Stebnicka, 2007
 Ataenius temperei Chalumeau & Gruner, 1974
 Ataenius terminalis (Chevrolat, 1864)
 Ataenius texanus Harold, 1874
 Ataenius tindalensis Stebnicka & Howden, 1997
 Ataenius tomentosus Petrovitz, 1969
 Ataenius torridus Blackburn, 1892
 Ataenius tovarensis Stebnicka, 2001
 Ataenius tuberculatus Schmidt, 1911
 Ataenius uriarrae Stebnicka & Howden, 1997
 Ataenius usingeri Hinton, 1937
 Ataenius utahensis Cartwright, 1974
 Ataenius vandykei Cartwright, 1974
 Ataenius variopunctatus Schmidt, 1922
 Ataenius versicolor Schmidt, 1916
 Ataenius vethianus Schmidt, 1909
 Ataenius vinacoensis Stebnicka, 2006
 Ataenius walkeri Blackburn, 1904
 Ataenius waltherhorni Balthasar, 1938
 Ataenius warisensis Stebnicka, 1998
 Ataenius wenzelii Horn, 1887
 Ataenius windjanae Stebnicka & Howden, 1997
 Ataenius yungasus Stebnicka, 2001

Referanser

Kilder 
 Generic Guide to New World Scarab Beetles - Ataenius 
 Z. Stebnicka og P.K. Lago (1995) The New World species of Ataenius Harold, 1867. V. Revision of the A. strigatus group (Scarabaeidae: Aphodiinae: Eupariini). Insecta Mundi 19: 55-83 
 biolib.cz - Oversikt over stammer og slekter

Eksterne lenker 

 

Gjødselbiller
Dyr formelt beskrevet i 1867