Duvalius

Duvalius  er en slekt av billene i delgruppen (underfamilien) skyggeløpere (Trechinae) i familiegruppen løpebiller.

Utseende 
Små til middelsstore (ca. 5 millimeter), spinkle, blanke, rødbrunlige løpebiller med lange antenner. De har reduserte øyne og er blinde.

Levevis 
Artene lever i huler, der de trolig ernærer seg av andre smådyr.

Utbredelse 
Slekten er utbredt i hulesystemer i Frankrike, Italia, på Balkan, i Lilleasia og Kaukasus.

Systematisk inndeling 
Systematikken følger Carabidae of the World. 

 ordenen biller, Coleoptera Linnaeus, 1758
 underordenen Adephaga Schellenberg, 1806
 overfamilien løpebiller og vannkalver, Caraboidea Latreille, 1802
 familien   løpebiller, Carabidae Latreille, 1802
 underfamilie skyggeløpere Trechinae Bonelli, 1810
 stammen Trechini Bonelli, 1810 
 delgruppe Trechina Bonelli, 1810
 Slekten Duvalius Delarouzee, 1859 
 Underslekten Biharotrechus Bokor, 1922
 Duvalius abnormis Knirsch, 1913
 Duvalius bedelensis Janak & P. Moravac, 1989
 Duvalius bekchievi Gueorgiev & Kostova, 2011
 Duvalius beshkovi Coiffait, 1970
 Duvalius biokovensis Holdhaus, 1911
 Duvalius breitianus Knirsch, 1912
 Duvalius brevipilosus Knirsch, 1927
 Duvalius budae Kenderessy, 1879
 Duvalius cicioarae Jeannel, 1930
 Duvalius cognatus J. Frivaldszky, 1879
 Duvalius coiffaiti Decou, 1967
 Duvalius comes Scheibel, 1934
 Duvalius droveniki Magrini, 1998
 Duvalius durmitorensis Apfelbeck, 1904
 Duvalius edithae Janak & P. Moravec, 1989
 Duvalius golesensis Winkler, 1926
 Duvalius hegeduesii J. Frivaldszky, 1880
 Duvalius herculis J. Frivaldszky, 1889
 Duvalius hickeri Knirsch, 1913
 Duvalius hurkai Janak & P. Moravec, 1989
 Duvalius kanabei Csiki, 1940
 Duvalius knirschi Janak & P. Monavec, 1989
 Duvalius laevigatus Bokor, 1913
 Duvalius maglicensis Winkler, 1933
 Duvalius mallaszii Csiki, 1901
 Duvalius mandibularis Jeannel, 1930
 Duvalius mariannae Knirsch, 1924
 Duvalius meschniggi Meixner, 1928
 Duvalius milleri J. Frivaldszky, 1862
 Duvalius nannus Jeannel, 1931
 Duvalius oltenicus Jeannel, 1928
 Duvalius paroecus J. Frivaldszky, 1865
 Duvalius peristericus J. Muller, 1914
 Duvalius pilifer Ganglbauer, 1891
 Duvalius redtenbacheri I. Frivaldszky von Frivald & J. Frivaldszky, 1857 
 Duvalius roseni Jeannel, 1929
 Duvalius scarisoarae Knirsch, 1913
 Duvalius semecensis Winkler, 1926
 Duvalius speiseri Ganglbauer, 1892
 Duvalius spiessi Jeannel & Mallasz, 1929
 Duvalius spinifer Jeannel, 1928
 Duvalius stilleri Reitter, 1913
 Duvalius sziladyi Csiki, 1904
 Duvalius transcarpathicus Shilenkov & Rizun, 1989
 Duvalius trescavicensis Ganglbauer, 1891
 Duvalius voitestii Jeannel, 1930
 Duvalius wercharatskii Rizun & Yanitskiy, 1994
 Duvalius winneguthi Apfelbeck, 1907
 Underslekten Duvalius (i snever forstand)
 Duvalius adelphus Magrini, Petrioli, Degiovanni & Bastianini, 2008
 Duvalius aliciae Magrini, Petrioli, Degiovanni & Bastianini, 2008
 Duvalius andreinii (Gestro, 1907)
 Duvalius andreuccii Magrini & Vanni, 1984
 Duvalius annamariae Vanni & Magrini, 1989
 Duvalius antonellae Casale, Giachino, Vailati & Vigna Taglianti, 1996
 Duvalius apuanus Dodero, 1917
 Duvalius aroaniae Casale, Giachino, Vailati & Vigna Taglianti, 1996
 Duvalius auberti Grenier, 1864
 Duvalius auruncus Vigna Taglianti & Magrini, 2008
 Duvalius ausonicus Vigna Taglianti & Magrini, 2008
 Duvalius avoni Vigna Taglianti & Magrini, 2008
 Duvalius balazuci Bruneau de Mire, 1948
 Duvalius baldensis Putzeys, 1870
 Duvalius balearicus Henrot, 1964
 Duvalius bastianini Magrini, 1998
 Duvalius battonii Straneo, 1959
 Duvalius bensae Castro, 1892
 Duvalius bertagnii Magrini, 1998
 Duvalius berthae (Jeannel, 1910)
 Duvalius bianchii Jeannel, 1928
 Duvalius bischoffi Maschnigg, 1936
 Duvalius boldorii Jeannel, 1926
 Duvalius bonadonius Giordan & Raffaldi, 1982
 Duvalius bonzanoi Casale & Vigna Taglianti, 1990
 Duvalius boschi Jeannel, 1929
 Duvalius brandisi Ganglbauer, 1896
 Duvalius brandmayri Vigna Taglianti & Magrini, 2008
 Duvalius breiti Ganglbauer, 1900
 Duvalius brucki (Piccioli, 1870)
 Duvalius brujasi Sainte-Claire Deville, 1901
 Duvalius cadurcus Jeannel, 1955
 Duvalius cailloli Sainte-Claire Deville, 1902
 Duvalius calandrii Casale & Vigna Taglianti, 1990
 Duvalius canevae Gestro, 1885
 Duvalius caoduroi Pace, 1986
 Duvalius carantii Sella, 1874
 Duvalius carchinii Vigna Taglianti, Magrini & Vanni, 1993
 Duvalius casalei Sciaky, 1992
 Duvalius casellii Gestro, 1898
 Duvalius cerrutii Sbordoni & Di Domenico, 1967
 Duvalius cirocchii Magrini & Vanni, 1986
 Duvalius clairi Abeille de Perrin, 1881
 Duvalius comottii Casale, Giachino, Vailati & Vigna Taglianti, 1996
 Duvalius consortii Magrini & Degiovanni, 2006
 Duvalius convexicollis Peyerimhoff, 1904
 Duvalius cornilloni Giordan & Raffaldi, 1998
 Duvalius curtii Giordan & Raffaldi, 1983
 Duvalius dattii Vigna Taglianti & Magrini, 2008
 Duvalius degiovannii Magrini & Vanni, 1985
 Duvalius delphinensis Abeille de Perrin, 1869
 Duvalius diaphanus Rottenberg, 1874
 Duvalius didonnai Magrini, Vanni & Inguscio, 1996
 Duvalius diniensis Peyerimhoff, 1904
 Duvalius doderoi Gestro, 1885
 Duvalius dolops Jeannel, 1937
 Duvalius doriae Fairmaire, 1859
 Duvalius dragacevensis S.Curcic, Pavicevic & B.Curcic, 2001
 Duvalius exaratus Schaum, 1860
 Duvalius fodori Scheibel, 1937
 Duvalius franchettii Luigioni, 1926
 Duvalius franziniorum Vigna Taglianti & Magrini, 2008
 Duvalius fuchsi Scheibel, 1937
 Duvalius fulvii Casale, Giachino, Vailati & Vigna Taglianti, 1996
 Duvalius gabriellavernae Casale, Giachino, Vailati & Vigna Taglianti, 1999
 Duvalius genesti Casale & Vigna Taglianti, 1984
 Duvalius gentilei Gestro, 1885
 Duvalius georgii J. Muller, 1922
 Duvalius gestroi Dodero, 1900
 Duvalius ghidinii Gestro & Dodero, 1909
 Duvalius giachinoi Casale & Vigna Taglianti, 1990
 Duvalius graecus Casale, Giachino, Vailati & Vigna Taglianti, 1996
 Duvalius guareschii Moscandini, 1950
 Duvalius hartigi Magrini, Bavieri & Vigna Taglianti, 2006
 Duvalius innamoratorum Magrini & Cirocchi, 2002
 Duvalius iolandae Magrini & Vanni, 1986
 Duvalius iuliae Casale, Giachino, Vailati & Vigna Taglianli, 1996
 Duvalius iulianae Vigna Taglianti & Casale, 1973
 Duvalius jureceki Dodeno, 1917
 Duvalius krueperi Schaum, 1862
 Duvalius kyllenicus Scheibel, 1937
 Duvalius laneyriei J. Ochs, 1948
 Duvalius lantosquensis Abeille da Perrin, 1881
 Duvalius latellai Vigna Taglianti & Magrini, 2008
 Duvalius laurentii Magrini & Bastianini, 2003
 Duvalius lemairei Giordan & Raffaldi, 1982
 Duvalius leonhardi Reitter, 1901
 Duvalius lepinensis Cenuti, 1950
 Duvalius lespesii Fairmaire, 1867
 Duvalius longhii Comolli, 1837
 Duvalius lucarellii Casale & Vigna Taglianti, 1990
 Duvalius lydiae Vigna Taglianti & Magrini, 2008
 Duvalius magdelainei Jeannel, 1914
 Duvalius magistrettianus Schatzmayr, 1940
 Duvalius maglianoi Giordan & Raffaldi, 1983
 Duvalius marii Vanni, Magrini & Pennisi, 1991
 Duvalius matocqi Giordan, 1987
 Duvalius meixneri Kreissl, 1993
 Duvalius merisioi Casale, Giachino, Vailati & Vigna Taglianti, 1996
 Duvalius milenae Casale, 1983
 Duvalius minozzii Dodero, 1917
 Duvalius mirei Deuve, 2001
 Duvalius mixanigi Daffner, 1993
 Duvalius moczarskyi J. Muller, 1917
 Duvalius montisageli Jeannel, 1947
 Duvalius montisoetae Casale, 1987
 Duvalius morisii Vigna Taglianti & Casale, 1973
 Duvalius muelleri Winkler, 1933
 Duvalius muriauxi Jeannel, 1957
 Duvalius nambinensis Boldori, 1935
 Duvalius nardii Vigna Taglianti & Magrini, 2008
 Duvalius occitanus Casale & Vigna Taglianti, 1993
 Duvalius ochsi Dodero, 1921
 Duvalius oertzeni L. Miller, 1884
 Duvalius olympiadicus Scheibel, 1937
 Duvalius oscus A. Franzini & C. Franzini, 1984
 Duvalius parnassianus Deuve, 2001
 Duvalius paulinae (Fagniez, 1922)
 Duvalius pecoudi Jeannel, 1937
 Duvalius pennisii Magrini & Vanni, 1984
 Duvalius perrinae Giordan, 1989
 Duvalius pominii Schatzmayr, 1943
 Duvalius raffaldii Curti, 1981
 Duvalius ramorinii (Gestro, 1887)
 Duvalius raymondi Delarouzee, 1859
 Duvalius regiszogui Maschnigg, 1936
 Duvalius roberti Abeille de Perrin, 1903
 Duvalius rossii Magrini & Vanni, 1991
 Duvalius ruffoi Magistretti, 1956
 Duvalius sardous (Dodero, 1917)
 Duvalius sbordonii Vigna Taglianti, Genest & Sciaky, 1980
 Duvalius sclanoi Magrini & Vanni, 1996
 Duvalius sicardi Fagniez, 1923
 Duvalius siculus (Baudi di Selva, 1882)
 Duvalius silvestrii (Gestro, 1896)
 Duvalius simoni (Abeille de Perrin, 1881)
 Duvalius smolikanus Casale, Giachino, Vailati & Vigna Taglianti, 1996
 Duvalius spaethi (Ganglbauer, 1904)
 Duvalius springeri (J. Muller, 1921)
 Duvalius starivlahi B.Gueorguiev, S.Curcic & B.Curcic, 2000
 Duvalius straneoi Jeannel, 1931
 Duvalius strupii Scheibel, 1937
 Duvalius sturanyi (Apfelbeck, 1904)
 Duvalius taygetanus Casale, 1979
 Duvalius turcati Giordan & Raffaldi, 1983
 Duvalius vallombrosus (C. & F. Rasetti, 1920)
 Duvalius vannii Magrini & Sclano, 1998
 Duvalius vermionensis (Casale, 1983)
 Duvalius vignai Casale, 1983
 Duvalius villiersi Giordan & Raffaldi, 1983
 Duvalius virginiae Magnini, Vanni & Cirocchi, 1996
 Duvalius volscus A. Franzini & G. Franzini, 1984
 Duvalius voraginis Jeannel & J. Ochs, 1938
 Duvalius waillyi Giordan & Raffaldi, 1982
 Duvalius weiratheri Scheibel, 1937
 Duvalius wichmanni Jeannel, 1929
 Duvalius wingelmuelleri (Ganglbauer, 1904)
 Duvalius winklerianus Jeannel, 1926
 Duvalius zaimisi Jeannel, 1929
 Underslekten Euduvalius Jeannel, 1928
 Duvalius erichsonii Schaufuss, 1864
 Duvalius lucidus J. Muller, 1903
 Duvalius maglajensis Apfelbeck, 1908
 Duvalius novaki J. Muller, 1911
 Duvalius petrochilosi Coiffait, 1965
 Duvalius reiseri Ganglbauer, 1891
 Duvalius ruffoanus Casale, Giachino, Vailati & Vigna Taglianti, 1996
 Duvalius zhalovi Gueorgiev, 2005
 Underslekten Hungarotrechus Bokor, 1922
 Duvalius babicola Knirsch, 1925
 Duvalius bokori Csiki, 1910
 Duvalius bucurensis Janak, 1993
 Duvalius centenarius Knirsch, 1925
 Duvalius corpulentus Weise, 1875
 Duvalius delamarei Decou, 1967
 Duvalius deubelianus Csiki, 1903
 Duvalius dieneri Csiki, 1910
 Duvalius gaali Mallasz, 1928
 Duvalius ganglbauerianus Knirsch, 1913
 Duvalius godeanus Janak, 1993
 Duvalius goemoeriensis Bokor, 1922
 Duvalius gracilis Petri, 1912
 Duvalius hungaricus Csiki, 1903
 Duvalius kimakowiczi Ganglbauer, 1891
 Duvalius merkli J. Frivaldszky, 1877
 Duvalius microphthalmus L. Miller, 1859
 Duvalius onaci Moldovan, 1993
 Duvalius poporogui Decou, 1973
 Duvalius proceroides Jeannel, 1926
 Duvalius procerus Putzeys, 1847
 Duvalius richardi Knirsch, 1925
 Duvalius roubali Jeannel, 1926
 Duvalius ruthenus Reitter, 1878
 Duvalius subterraneus (L. Miller, 1868)
 Duvalius szaboi Csiki, 1914
 Underslekten Neoduvalius G. Muller, 1913
 Duvalius bolei Pralmer, 1963
 Duvalius bradycephalus Jeannel, 1928
 Duvalius cuniculinus Knirsch, 1929
 Duvalius cvijici (Jeannel, 1924)
 Duvalius eurydice Schaufuss, 1881
 Duvalius humerosus Knirsch, 1926
 Duvalius klimeschi Winkler, 1914
 Duvalius kodrici Scheibel, 1938
 Duvalius langhofferi Csiki, 1913
 Duvalius neumanni J. Muller, 1911
 Duvalius opermanni Scheibel, 1933
 Duvalius petraeus Knirsch, 1927
 Duvalius reitteri Miller, 1881
 Duvalius schatzmayri J. Muller, 1912
 Duvalius styx Apfelbeck, 1904
 Duvalius vranensis Breit, 1904
 Underslekten Paraduvalius Knirsch, 1924
 Duvalius balcanicus (J. Frivaldszky, 1879)
 Duvalius beroni V.B. Gueorguiev, 1971
 Duvalius bulgaricus Knirsch, 1924
 Duvalius bureschi Jeannel, 1928
 Duvalius deltshevi V.B. Gueorguiev, 1965
 Duvalius garevi Casale & Genest, 1991
 Duvalius germanae Casale & Vigna Taglianti, 1990
 Duvalius glabellus G. Etonti & M. Etonti, 1984
 Duvalius hanae Hurka, 1990
 Duvalius joakimovi Gueorgiev, 2005
 Duvalius joannidisi Casale, Giachino & M. Elonti, 1990
 Duvalius karelhurkai Farkac, 1990
 Duvalius kotelensis Genest, 1978
 Duvalius legrandi Genest, 1983
 Duvalius marani Knirsch, 1930
 Duvalius papasoffi Mandl, 1942
 Duvalius petrovi Gueorgiev, 2005
 Duvalius philippensis Jeannel, 1937
 Duvalius pirinensis Gueorgiev, 2005
 Duvalius pretneri V.B. Gueorguiev, 1971
 Duvalius pruinosus Jeannel, 1937
 Duvalius rajtchevi Genest & Juberthie, 1983
 Duvalius regisborisi Buresch, 1926
 Duvalius stankovitchi Jeannel, 1924
 Duvalius sydowi Jeannel, 1930
 Duvalius winkleri Jeannel, 1924
 Duvalius zivkovi (Knirsch, 1925)
 Underslekten Platyduvalius Jeannel, 1928
 Duvalius gogalai Pretnen, 1963
 Duvalius macedonicus J. Muller, 1917
 Underslekten Trechopsis Peyerimhoff, 1908
 Duvalius ferreresi Lagar Mascare, 1976
 Underslekten Typhloduvalius Hurka & Pulpan, 1980
 Duvalius gebhardti Bokor, 1926

Liste over arter ikke plassert til underslekt 

 Duvalius abyssimus Reboleira & Ortuño, 2014
 Duvalius alexeevi Belousov, 1992
 Duvalius antoniae Reitter, 1892
 Duvalius armeniacus Casale, 1979
 Duvalius arnoldii Jeannel, 1962
 Duvalius baborensis Bruneau de Mire, 1955
 Duvalius bicikensis Perrault, 1971
 Duvalius bodoanus Reitter, 1913
 Duvalius bortesii Casale & Vigna Taglianti, 1984
 Duvalius bruschii Vigna Taglianti, 1999
 Duvalius dvoraki P. Moravec, 1986
 Duvalius gusevi Belousov, 1989
 Duvalius hetschkoi (Reitter, 1911)
 Duvalius huberi Casale, 2011
 Duvalius huetheri Jeannel, 1934
 Duvalius iblis Payerimhoff, 1910
 Duvalius iljukhini Dalzhanski & Ljovuschkin, 1985
 Duvalius jurjurae Peyerimhoff, 1909
 Duvalius koeni Muilwijk & Felix, 2008
 Duvalius kryshanovskii Jeannel, 1962
 Duvalius kurnakovi Jeannel, 1960
 Duvalius lapiei Peyerimhoff, 1908
 Duvalius lencinai Mateu & Ortuno, 2006
 Duvalius martensi Casale, 1983
 Duvalius martinae Jeanne, 1996
 Duvalius megrel Belousov, 1992
 Duvalius miroshnikovi Belousov & Zamotajlov, 1995
 Duvalius mohammadzadehi Muilwijk & Felix, 2008
 Duvalius ovtshinnikovi Belousov, 1992
 Duvalius panoecus J. Frivaldszky, 1865
 Duvalius phoenicius Vigna Taglianti, 1973
 Duvalius putshkovi Belousov & Kabak, 1999
 Duvalius saueri Farkac & P. Moravec, 1993
 Duvalius sokolovi Ljovuschkin, 1963
 Duvalius stepanavanensis Iablokoff-Khnzorian, 1963
 Duvalius stopicensis (Jeannel, 1924)
 Duvalius vartashensis Belousov, 1989
 Duvalius yatsenkokhmelevskyi Iablokoff-Khnzorian, 1960
 Duvalius zlatiborensis (Curcic, Brajkovic & Curcic, 2005)

Referanser

Eksterne lenker 
 Bilde av Duvalius delphinensis
 

skyggeløpere
Biller formelt beskrevet i 1859