Leptotyphlus

Leptotyphlus er en slekt av biller som hører til familien kortvinger (Staphylinidae). Denne gruppen er representert med tallrike arter i Mellom- og Sør-Europa, men finnes ikke nord for Østerrike.

Utseende 
Bittesmå (0,8-2,1 millimeter), slanke, blekt fargede (gjerne rødlige) kortvinger. Disse billene bærer preg av sitt underjordiske levesett. Øyne og flygevinger mangler helt. Kroppen er lang og tynn, jevbred, hodet forholdsvis stort med korte antenner. Pronotum og dekkvinger er ganske små, bakkroppen lang og smal. Beina er korte.

Levevis 
Disse billene lever dypt nede i jorda og kommer sjelden til overflaten. De fleste av de kjente artene lever i områder med middelhavsklima. Det har blitt spekulert i at årsaken til at gruppen mangler i Nord-Europa er at de ennå ikke har vandret inn etter siste istid, da disse billene trolig har ganske dårlig spredningsevne.

Utbredelse 
Slekten har mange arter i det sørlige Europa.

Systematisk inndeling 

 Ordenen biller, Coleoptera
 Underordenen Polyphaga
 Overfamilien åtsel- og rovbiller, Staphylinoidea
 Familien kortvinger (Staphylinidae) Latreille, 1802
 Underfamilien Leptotyphlinae Fauvel, 1874 – 435 arter i Europa
 Stamme Leptotyphlini Fauvel, 1874
 Slekten Leptotyphlus Fauvel, 1874 - ca. 200 arter

Lister over artene 

 Leptotyphlus aberrans Orousset, 2013
 Leptotyphlus admirandus Orousset, 2013
 Leptotyphlus agrestis Orousset, 1984
 Leptotyphlus aithaliensis Orousset, 1983
 Leptotyphlus albareensis Coiffait, 1957
 Leptotyphlus albingaunicus Pace, 1977
 Leptotyphlus anachoreta Pace, 1975
 Leptotyphlus anchorifer Coiffait, 1957
 Leptotyphlus angelicus Coiffait, 1955
 Leptotyphlus antonius Coiffait, 1959
 Leptotyphlus antrorum Coiffait, 1959
 Leptotyphlus apenninus Pace, 1991
 Leptotyphlus arcsensis Coiffait, 1959
 Leptotyphlus argensis Coiffait, 1957
 Leptotyphlus argodi Coiffait, 1955
 Leptotyphlus argus
 Leptotyphlus arizanus Coiffait, 1959
 Leptotyphlus armifer Orousset, 2013
 Leptotyphlus ascoensis Orousset, 2013
 Leptotyphlus asinus Coiffait, 1962
 Leptotyphlus aspericeps
 Leptotyphlus asprensis Coiffait, 1957
 Leptotyphlus asturicus
 Leptotyphlus attis Coiffait, 1955
 Leptotyphlus audrasi Jarrige, 1949
 Leptotyphlus aurensis Coiffait, 1957
 Leptotyphlus babas
 Leptotyphlus baillauriensis Coiffait, 1962
 Leptotyphlus balachowskyi Hervé, 1966
 Leptotyphlus bastiensis Coiffait, 1973
 Leptotyphlus beieri
 Leptotyphlus belenus Coiffait, 1955
 Leptotyphlus bergistanus Hernando, 2012
 Leptotyphlus bicorniger Orousset, 1985
 Leptotyphlus bifoveolatus
 Leptotyphlus bifurcus Orousset, 1984
 Leptotyphlus bigerricus Coiffait, 1959
 Leptotyphlus boiteli
 Leptotyphlus bonadonai Coiffait, 1955
 Leptotyphlus bordei Coiffait, 1957
 Leptotyphlus bormanus Coiffait, 1955
 Leptotyphlus bouilloni Coiffait, 1959
 Leptotyphlus boulouensis Coiffait, 1962
 Leptotyphlus brescius Thélot, 1980
 Leptotyphlus brevicornis Coiffait, 1955
 Leptotyphlus calabricus Coiffait, 1970
 Leptotyphlus calcatogius Coiffait, 1959
 Leptotyphlus calenzanus Coiffait, 1959
 Leptotyphlus campanus Pace, 1977
 Leptotyphlus camulus Coiffait, 1955
 Leptotyphlus capito
 Leptotyphlus caqus
 Leptotyphlus cargeseensis Orousset, 1984
 Leptotyphlus cariniventris Coiffait, 1959
 Leptotyphlus castillonensis Coiffait, 1957
 Leptotyphlus chappuisi Coiffait, 1959
 Leptotyphlus charon Coiffait, 1955
 Leptotyphlus cheironensis Thélot, 1985
 Leptotyphlus chiavariensis Orousset, 1984
 Leptotyphlus circaeus Pace, 1977
 Leptotyphlus cirtulus
 Leptotyphlus coiffaiti Hervé, 1964
 Leptotyphlus collensis Coiffait, 1959
 Leptotyphlus compitalis Hervé, 1965
 Leptotyphlus confrater Normand, 1947
 Leptotyphlus corsicus
 Leptotyphlus couseranus Coiffait, 1959
 Leptotyphlus crassiphallus Coiffait, 1955
 Leptotyphlus cribratus Fauvel, 1890
 Leptotyphlus cyclops
 Leptotyphlus cyrneus Orousset, 1984
 Leptotyphlus dapsilis
 Leptotyphlus demeter Coiffait, 1955
 Leptotyphlus dentatus Coiffait, 1957
 Leptotyphlus denticulatus Coiffait, 1957
 Leptotyphlus devillei Coiffait, 1957
 Leptotyphlus diecki Orousset, 2013
 Leptotyphlus dispar Hervé, 1965
 Leptotyphlus dispersus Hervé, 1967
 Leptotyphlus distinctus Coiffait, 1955
 Leptotyphlus doderoi
 Leptotyphlus domensis Coiffait, 1957
 Leptotyphlus dubius Coiffait, 1957
 Leptotyphlus dujardini Hervé, 1967
 Leptotyphlus eaqus Coiffait, 1957
 Leptotyphlus elbanus Pace, 1980
 Leptotyphlus estellensis Struyve, 2013
 Leptotyphlus esus Coiffait, 1955
 Leptotyphlus eurydice Coiffait, 1962
 Leptotyphlus exilis (Mulsant & Rey, 1875)
 Leptotyphlus eximius  Orousset, 1988
 Leptotyphlus exsculptus
 Leptotyphlus ezeensis Coiffait, 1957
 Leptotyphlus falcata Hernando, 2012
 Leptotyphlus fallax Orousset, 1985
 Leptotyphlus fancelloi Hernando, 2006
 Leptotyphlus farinolensis Orousset, 1985
 Leptotyphlus faunus
 Leptotyphlus finalensis Pace, 1977
 Leptotyphlus fissuralis Coiffait, 1959
 Leptotyphlus fonfrigidus Coiffait, 1962
 Leptotyphlus foroiuliensis Pace, 1976
 Leptotyphlus foveiceps Dodero, 1908
 Leptotyphlus franzi
 Leptotyphlus furcatus Coiffait, 1957
 Leptotyphlus galiberti Coiffait, 1955
 Leptotyphlus gallaecianus
 Leptotyphlus garbetensis Coiffait, 1957
 Leptotyphlus gardensis Coiffait, 1959
 Leptotyphlus garomettensis
 Leptotyphlus giglionicus Coiffait, 1955
 Leptotyphlus girbeni Hernando, 2012
 Leptotyphlus gracilis Coiffait, 1957
 Leptotyphlus grannus Coiffait, 1955
 Leptotyphlus grouvellei
 Leptotyphlus hades Coiffait, 1955
 Leptotyphlus harmatus Orousset, 2013
 Leptotyphlus henryi Coiffait, 1959
 Leptotyphlus hervei Jarrige, 1949
 Leptotyphlus hipponensis
 Leptotyphlus ilvensis Orousset, 1983
 Leptotyphlus impressus Coiffait, 1955
 Leptotyphlus inopinatus Pace, 1975
 Leptotyphlus inornatus Orousset, 2013
 Leptotyphlus irenae Coiffait, 1952
 Leptotyphlus janasensis Hervé, 1966
 Leptotyphlus jeanneli
 Leptotyphlus joannis Hervé, 1971
 Leptotyphlus judicariae
 Leptotyphlus kabylianus
 Leptotyphlus kahleni Pace, 1991
 Leptotyphlus kovaci Šustek, 2000
 Leptotyphlus kyrnosus
 Leptotyphlus laetificus
 Leptotyphlus lamberti Thélot, 1979
 Leptotyphlus laminatus Coiffait, 1959
 Leptotyphlus laneyriei Hervé, 1965
 Leptotyphlus langastouensis Hervé, 1969
 Leptotyphlus laticeps Dodero, 1908
 Leptotyphlus lavagni
 Leptotyphlus lavezzianus Orousset, 2007
 Leptotyphlus lepinensis Pace, 1977
 Leptotyphlus lessinicus Pace, 1973
 Leptotyphlus levasseuri Coiffait, 1955
 Leptotyphlus liguricus Coiffait, 1957
 Leptotyphlus lobatus Coiffait, 1959
 Leptotyphlus londensis Coiffait, 1959
 Leptotyphlus lucensis Coiffait, 1962
 Leptotyphlus luci Hervé, 1972
 Leptotyphlus magrinii Pace, 1995
 Leptotyphlus mandibularis Coiffait, 1957
 Leptotyphlus maritimus Coiffait, 1957
 Leptotyphlus martellii Pace, 1974
 Leptotyphlus massatensis Coiffait, 1959
 Leptotyphlus maureillensis Coiffait, 1962
 Leptotyphlus maurettensis Hervé, 1968
 Leptotyphlus mauryensis Coiffait, 1962
 Leptotyphlus mayeti
 Leptotyphlus mendicus
 Leptotyphlus meriensis Coiffait, 1959
 Leptotyphlus micros
 Leptotyphlus mimus Pace, 1977
 Leptotyphlus minator Fancello, Hernando & Leo, 2009
 Leptotyphlus minervius Coiffait, 1959
 Leptotyphlus minos Coiffait, 1955
 Leptotyphlus moczarskianus
 Leptotyphlus molisensis Pace, 1977
 Leptotyphlus monachus  Hervé, 1967
 Leptotyphlus muricatus Orousset, 1988
 Leptotyphlus nardii Fancello, Hernando & Leo, 2009
 Leptotyphlus necropolis Coiffait, 1962
 Leptotyphlus nerviensis Coiffait, 1962
 Leptotyphlus nistosensis Coiffait, 1965
 Leptotyphlus nonzius
 Leptotyphlus normandi Coiffait, 1955
 Leptotyphlus ochsi Coiffait, 1955
 Leptotyphlus oglasensis Poggi, 1976
 Leptotyphlus olotensis Coiffait, 1959
 Leptotyphlus omessae
 Leptotyphlus orbensis
 Leptotyphlus orcus Coiffait, 1955
 Leptotyphlus orion
 Leptotyphlus orpheus Coiffait, 1957
 Leptotyphlus osellai Pace, 1973
 Leptotyphlus pauxillus
 Leptotyphlus peillensis Coiffait, 1957
 Leptotyphlus perpensus
 Leptotyphlus perroti Coiffait, 1957
 Leptotyphlus pertusatus
 Leptotyphlus petretensis
 Leptotyphlus phami Orousset, 1987
 Leptotyphlus phoenix Coiffait, 1962
 Leptotyphlus picardi Lavagne, 1917
 Leptotyphlus picenus Pace, 1977
 Leptotyphlus pluto Coiffait, 1955
 Leptotyphlus poirioni Hervé, 1971
 Leptotyphlus pontelecciensis Orousset, 1987
 Leptotyphlus priapus Coiffait, 1955
 Leptotyphlus proprianensis
 Leptotyphlus pugnator Orousset, 2013
 Leptotyphlus pyrenaeus Coiffait, 1955
 Leptotyphlus queinneci Orousset, 1984
 Leptotyphlus remensis Pace, 1977
 Leptotyphlus reticulatus Coiffait, 1959
 Leptotyphlus revelierianus Orousset, 1988
 Leptotyphlus revelierii
 Leptotyphlus rhadamanthus Coiffait, 1957
 Leptotyphlus ribasensis Coiffait, 1957
 Leptotyphlus rivularis Coiffait, 1962
 Leptotyphlus romanus Pace, 1975
 Leptotyphlus ruffoi Pace, 1975
 Leptotyphlus saevus Orousset, 2013
 Leptotyphlus samniticus Pace, 1977
 Leptotyphlus sardiniensis Herman, 2001
 Leptotyphlus sardous Coiffait, 1955
 Leptotyphlus sassii Hervé, 1966
 Leptotyphlus saturnius
 Leptotyphlus saulcyi Croissandeau, 1891
 Leptotyphlus savonensis Pace, 1977
 Leptotyphlus sbordonii Pace, 1991
 Leptotyphlus schuleri Hervé, 1967
 Leptotyphlus senensis
 Leptotyphlus senyensis Coiffait, 1957
 Leptotyphlus serianensis Coiffait, 1957
 Leptotyphlus sibyllinicus Pace, 2002
 Leptotyphlus siccanus
 Leptotyphlus simplarius
 Leptotyphlus siscoi Hernando, 2015
 Leptotyphlus solarii Coiffait, 1957
 Leptotyphlus sospelensis Coiffait, 1957
 Leptotyphlus sourniensis Coiffait, 1962
 Leptotyphlus strobinoi Hervé, 1967
 Leptotyphlus subhervei
 Leptotyphlus sublaevis Fauvel, 1874
 Leptotyphlus sublaneyriei Hervé, 1965
 Leptotyphlus subloevis
 Leptotyphlus subpyrenaeus Coiffait, 1959
 Leptotyphlus sucellus Coiffait, 1955
 Leptotyphlus sylvanus Coiffait, 1955
 Leptotyphlus taggiensis Coiffait, 1962
 Leptotyphlus tellus
 Leptotyphlus thaumasios Orousset, 1987
 Leptotyphlus toscanicus
 Leptotyphlus transensis Coiffait, 1959
 Leptotyphlus tressensi Coiffait, 1965
 Leptotyphlus trifurcatus Coiffait, 1959
 Leptotyphlus troglophilus Coiffait, 1957
 Leptotyphlus tronqueti Struyve, 2013
 Leptotyphlus tuniseus Normand, 1910
 Leptotyphlus truggiensis Orousset, 1984
 Leptotyphlus tyrrhenius Dodero, 1908
 Leptotyphlus uccellinensis Pace, 1979
 Leptotyphlus vandeli Coiffait, 1959
 Leptotyphlus varagensis Thélot, 1980
 Leptotyphlus velatus Coiffait, 1955
 Leptotyphlus venator Orousset, 2013
 Leptotyphlus veneriatus
 Leptotyphlus ventralis Coiffait, 1962
 Leptotyphlus venus Coiffait, 1955
 Leptotyphlus veronensis Pace, 1978
 Leptotyphlus vicus Coiffait, 1959
 Leptotyphlus villacidrinus Fancello, Hernando & Leo, 2009
 Leptotyphlus villiersi
 Leptotyphlus viti Pace, 1978
 Leptotyphlus vivesi Coiffait, 1976
 Leptotyphlus vulcanus
 Leptotyphlus weulerssae Orousset, 2013
 Leptotyphlus zariquieyi Coiffait, 1955

Referanser

Kilder 
 Fauna Europaea Web Service Version 2.4 (2011) Utbredelsesdatabase for europeiske dyr: 
 Grebennikov, V.V. og A.F. Newton, 2008: Minute larvae of Leptotyphlinae (Coleoptera: Staphylinidae): description of three genera with discussion on the monophyly and phylogenetic position of the subfamily as inferred from larval morphology. Zootaxa, 1817: 49-58. 
 Gusarov, V.: Prosjektbeskrivelse. Med bilder

Eksterne lenker 
 
 Bilde av to arter fra Russland: 

Leptotyphlinae
Biller formelt beskrevet i 1874