Dyschirius

Dyschirius er en slekt av billene i delgruppen (underfamilien) graveløpere (Scaritinae) i familiegruppen løpebiller.

Utseende 
Små (2,5-7,5 millimeter), avlange, blanke, mørke løpebiller med kraftige bein. Pronotum og dekkvingene er rundet hver for seg, overflatisk kan de minne vagt om maur. Antennene er perlekjede-formede og forholdsvis korte. Hodet er forholdsvis lite med middelsstore fasettøyne og kjever. Pronotum er rundet og hvelvet, dekkvingene rundede. Beina er utviklet til gravebein.

Levevis 
Disse billene graver i bakken og lever for det meste underjordisk. Flere av artene lever på havstrender, der de gjerne graver seg ned i sand eller mudder ved flo sjø. De er rovdyr som jakter på små leddyr.

Utbredelse 
Slekten forekommer i alle verdensdeler bortsett fra Australia (og Antarktis). De fleste av artene lever Palearktis.

Systematisk inndeling 
Oversikten har med de europeiske artene.   Systematikken følger Carabidae of the World. 

 ordenen biller, Coleoptera Linnaeus, 1758
 underordenen Adephaga Schellenberg, 1806
 overfamilien løpebiller og vannkalver, Caraboidea Latreille, 1802
 familien   løpebiller, Carabidae Latreille, 1802
 underfamilie graveløpere Scaritinae Bonelli, 1810
 stammen Dyschiriini W. Kolbe, 1880 
 slekten Dyschirius Bonelli, 1810 - arter som forekommer i Norge:
 Dyschirius angustatus (Ahrens, 1830) 
 Dyschirius armatus Wollaston 1864
 Dyschirius caspius Putzeys 1866
 Dyschirius digitatus (Dejean 1825)
 Dyschirius humeratus Chaudoir 1850
 Dyschirius latipennis Seidlitz 1867
 Dyschirius numidicus Putzeys 1846
 Dyschirius obscurus (Gyllenhal, 1827)
 Dyschirius thoracicus (P. Rossi, 1790)
 Dyschirius zimini Znojko 1928

Liste over arter som ikke forekommer i Norge 
 Dyschirius abbreviatus Putzeys, 1846 i c g b
 Dyschirius abditus Fedorenko, 1993 c g
 Dyschirius addisabeba (Bulirsch, 2006) c g
 Dyschirius aenea Boheman, 1849
 Dyschirius aeneo Boheman, 1849 c
 Dyschirius aeneolus Leconte, 1850
 Dyschirius aeneus (Dejean, 1825) c g
 Dyschirius affinis Fall, 1901 i c g
 Dyschirius afghanus Jedlicka, 1967 c g
 Dyschirius agnatus Motschulsky, 1844 c g
 Dyschirius aida Schatzmayr, 1936 c g
 Dyschirius alajensis Znojko, 1930 c g
 Dyschirius alticola Lindroth, 1961 i c g
 Dyschirius amazonicus Fedorenko, 1991 c g
 Dyschirius ambiguus Fedorenko, 1994 c g
 Dyschirius amphibolus J. Muller, 1922
 Dyschirius amurensis Fedorenko, 1991 c g
 Dyschirius analis Leconte, 1852
 Dyschirius angolensis (Fedorenko, 2000) c g
 Dyschirius angusticollis Putzeys, 1867 c g
 Dyschirius anichtchenkoi (Bulirsch, 2015) c g
 Dyschirius apicalis Putzeys, 1846 c g
 Dyschirius apicius Gistel, 1857 c
 Dyschirius aratus Leconte, 1852
 Dyschirius arcanus Bruneau de Miré, 1952 c g
 Dyschirius arcifer Znojko, 1928 c g
 Dyschirius arnoldii Gryuntal, 1984 c g
 Dyschirius asper Andrewes, 1929 c g
 Dyschirius assegaaicus (Fedorenko, 2000) c g
 Dyschirius auriculatus Wollaston, 1867 c g
 Dyschirius bacillus Schaum, 1857 c g
 Dyschirius baehri (Bulirsch, 2006) c g
 Dyschirius baenningeri (Fedorenko, 2004) c g
 Dyschirius baicalensis Motschulsky, 1844 c g
 Dyschirius bechynei Kult, 1949 c g
 Dyschirius becvari (Bulirsch, 2006) c g
 Dyschirius beludscha Tschitscherine, 1904
 Dyschirius beludsha Tschitscherine, 1904 c g
 Dyschirius benedikti Bulirsch, 1995 c g
 Dyschirius bengalensis Andrewes, 1929 c g
 Dyschirius beydagensis Jeanne, 1996 c g
 Dyschirius bifrons Andrewes, 1929 c g
 Dyschirius binodosus Putzeys, 1878 c g
 Dyschirius boliviensis (Bulirsch, 2015) c g
 Dyschirius bonellii Putzeys, 1846 c g
 Dyschirius bousqueti (Bulirsch, 2006) c g
 Dyschirius braziliensis Fedorenko, 1999
 Dyschirius breviphthalmus Balkenohl & Lompe, 2003 c g
 Dyschirius brevispinus Leconte, 1878
 Dyschirius bruchi (Kult, 1950) c g
 Dyschirius bryanti (Kult, 1950) c g
 Dyschirius buglanensis Bulirsch, 1996 c g
 Dyschirius campicola Lindroth, 1961 i c g b
 Dyschirius cariniceps Baudi di Selve, 1864 c g
 Dyschirius carri Bousquet, 1996 c g
 Dyschirius carrorum Bousquet, 1997 i
 Dyschirius cerberus Larson, 1968 i c g
 Dyschirius chalceus Erichson, 1837 c g
 Dyschirius chalybeus Putzeys, 1846 c g
 Dyschirius championi Andrewes, 1929 c g
 Dyschirius changlingensis Li, 1992 c g
 Dyschirius cheloscelis Bates, 1873 c g
 Dyschirius chiricahuae Dajoz, 2004
 Dyschirius clorinda Bulirsch, 2009
 Dyschirius clypeatus Putzeys, 1867 c g
 Dyschirius comatus Bousquet, 1988 i c g
 Dyschirius compactus Lindroth, 1961 i c g
 Dyschirius consobrinus Leconte, 1852
 Dyschirius constrictus Andrewes, 1929 c g
 Dyschirius contortus (Fedorenko, 1997) c g
 Dyschirius crenulatus Putzeys, 1867 c g
 Dyschirius criddlei Fall, 1925 i c g
 Dyschirius crinifer Balkenohl, 1993 c g
 Dyschirius curvispinus Putzeys, 1846 i c g
 Dyschirius cylindricus (Dejean, 1825) c g
 Dyschirius darlingtoni Kult, 1950 c g
 Dyschirius darwini (Kult, 1950) c g
 Dyschirius dejeanii Putzeys, 1846 i c g b
 Dyschirius devroeyianus Burgeon, 1935 c g
 Dyschirius dimidiatus Chaudoir, 1846 c g
 Dyschirius disjunctus Andrewes, 1929 c g
 Dyschirius dispar Péringuey, 1896 c g
 Dyschirius distinctus Motschulsky, 1850 c
 Dyschirius dostali (Bulirsch & Fedorenko, 2007) c g
 Dyschirius ecuadorensis (Bulirsch, 2006) c g
 Dyschirius edentulus Putzeys, 1846 i c g
 Dyschirius eduardinus Burgeon, 1935 c g
 Dyschirius erwini Bulirsch, 2009
 Dyschirius erythrocerus Leconte, 1857 i c g b
 Dyschirius euphraticus Putzeys, 1846 c g
 Dyschirius euxinus Znojko, 1927 c g
 Dyschirius exaratus Putzeys, 1867 c g
 Dyschirius exochus Whitehead, 1970 i c g
 Dyschirius extensus Putzeys, 1846 c g
 Dyschirius fabbrii (Bulirsch & Magrini, 2006) c g
 Dyschirius facchinii Bulirsch, 2009
 Dyschirius fassatii Kult, 1949 c g
 Dyschirius fedorenkoi (Bulirsch, 2006) c g
 Dyschirius ferganensis Znojko, 1930 c g
 Dyschirius ferrugineus Bousquet, 1988 i c g
 Dyschirius fianarensis Bulirsch, 2006 c g
 Dyschirius filiformis Leconte, 1857
 Dyschirius flavicornis Péringuey, 1896 c g
 Dyschirius fleischeri Sainte-Claire Deville, 1904 c g
 Dyschirius fossifrons Putzeys, 1867 c g
 Dyschirius franzi Kult, 1954 c g
 Dyschirius freyi Jedlicka, 1958 c g
 Dyschirius fulgidus Motschulsky, 1850 c g
 Dyschirius fulvipes (Dejean, 1825) c g
 Dyschirius fulvus (Fedorenko, 2000) c g
 Dyschirius fusus Putzeys, 1878 c g
 Dyschirius genieri Bulirsch, 2009
 Dyschirius gerardi Burgeon, 1935 c g
 Dyschirius gibbipennis Leconte, 1857
 Dyschirius girardi Kult, 1949 c g
 Dyschirius globosus Herbst, 1784
 Dyschirius globulosus (Say, 1823) i c g b
 Dyschirius gracilis (Heer, 1837) c g
 Dyschirius guatemalenus Bates, 1881 c g
 Dyschirius haemorrhoidalis (Dejean, 1831) i c g b
 Dyschirius hajeki Bulirsch, 2009
 Dyschirius hassei Kult, 1954
 Dyschirius hessei Kult, 1954 c g
 Dyschirius heydeni A.Fleischer, 1899 c g
 Dyschirius hiemalis Bousquet, 1987 i g b
 Dyschirius himalaicus (Fedorenko, 1997) c g
 Dyschirius hingstoni Andrewes, 1929 c g
 Dyschirius hiogoensis Bates, 1873 c g
 Dyschirius hipponensis Pic, 1894 c g
 Dyschirius hoberlandti Kult, 1954 c g
 Dyschirius humiolcus Chaudoir, 1850 c g
 Dyschirius importunoides Jeanne, 1996 c g
 Dyschirius importunus Schaum, 1857 c g
 Dyschirius impressifrons Fedorenko, 1993 c g
 Dyschirius impressus Putzeys, 1846 c g
 Dyschirius impunctipennis Dawson, 1854 c g
 Dyschirius insularis (Bulirsch & Fedorenko, 2013) c g
 Dyschirius integer Leconte, 1852
 Dyschirius interior Fall, 1922 i c g b
 Dyschirius intermedius Putzeys, 1846 c g
 Dyschirius intricatus (Fedorenko, 2000) c g
 Dyschirius jedlickai Kult, 1940 c g
 Dyschirius jelineki Bulirsch, 2009
 Dyschirius jindrai Fedorenko, 2004
 Dyschirius jordanicus Fedorenko, 1996
 Dyschirius kabakovi (Fedorenko, 1996) c g
 Dyschirius kadleci Bulirsch, 2009
 Dyschirius kaliki Kult, 1949 c g
 Dyschirius katanganus Burgeon, 1935 c g
 Dyschirius kirghizicus Fedorenko, 1994 c g
 Dyschirius kirschenhoferi (Bulirsch, 2012) c g
 Dyschirius kryzhanovskii Gryuntal, 1984 c g
 Dyschirius lacustris Andrewes, 1929 c g
 Dyschirius laevifasciatus G.Horn, 1878
 Dyschirius laeviusculus Putzeys, 1846 c g
 Dyschirius lambertoni Vuillet, 1910 c g
 Dyschirius larochellei Bousquet, 1988 i c g b
 Dyschirius lgockii A.Fleischer, 1912 c g
 Dyschirius limpopo (Fedorenko, 2000) c g
 Dyschirius longicollis Motschulsky, 1844 c g
 Dyschirius longipennis Putzeys, 1867 c g
 Dyschirius longulus Leconte, 1850 i c g b
 Dyschirius luticola Chaudoir, 1850 c g
 Dyschirius luzonicus Kult, 1949 c g
 Dyschirius macroderus Chaudoir, 1850 c g
 Dyschirius macrophthalmus (Fedorenko, 1999) c g
 Dyschirius mafuga (Bulirsch, 2006) c g
 Dyschirius magrinii (Bulirsch, 2012) c g
 Dyschirius mahratta Andrewes, 1929 c g
 Dyschirius malawicus (Bulirsch, 2006) c g
 Dyschirius marani Kult, 1949 c g
 Dyschirius matisi Lafer, 1989 c g
 Dyschirius melancholicus Putzeys, 1867 i c g
 Dyschirius mesopotamicus J. Muller, 1922
 Dyschirius microthorax Motschulsky, 1844 c g
 Dyschirius milloti Jeannel, 1949 c g
 Dyschirius minarum Putzeys, 1867 c g
 Dyschirius minutus (Dejean, 1825) c g
 Dyschirius montanus Leconte, 1879
 Dyschirius moraveci (Bulirsch, 2006) c g
 Dyschirius morio Putzeys, 1867 c g
 Dyschirius mortchaensis Bruneau de Miré, 1952 c g
 Dyschirius natalensis (Fedorenko, 1997) c g
 Dyschirius neoteutonus Fedorenko, 1991 c g
 Dyschirius neresheimeri Wagner, 1915
 Dyschirius nianus Fedorenko, 1993 c g
 Dyschirius nigricornis Motschulsky, 1844 i c g
 Dyschirius nitens Putzeys, 1878 c g
 Dyschirius nitidus (Dejean, 1825) c g
 Dyschirius ogloblini (Kult, 1950) c g
 Dyschirius opistholius Alluaud, 1936 c g
 Dyschirius ordinatus Bates, 1873 c g
 Dyschirius orientalis Putzeys, 1867 c g
 Dyschirius owen Dajoz, 2004
 Dyschirius pacificus Lindroth, 1961 i c g
 Dyschirius paleki Kult, 1949 c g
 Dyschirius pallipennis (Say, 1823) i c g b
 Dyschirius pampicola Putzeys, 1867 c g
 Dyschirius parallelus Motschulsky, 1844 c g
 Dyschirius parvulus Péringuey, 1896 c g
 Dyschirius patruelis Leconte, 1852
 Dyschirius paucipunctus Andrewes, 1929 c g
 Dyschirius pauxillus Wollaston, 1864 c g
 Dyschirius peringueyi Kult, 1954 c g
 Dyschirius persicus Fedorenko, 1994 c g
 Dyschirius peruanus Fedorenko, 1991 c g
 Dyschirius perversus Fall, 1922 i c g
 Dyschirius peyrierasi Basilewsky, 1976 c g
 Dyschirius pfefferi Kult, 1949 c g
 Dyschirius pilosus Leconte, 1857 i c g b
 Dyschirius planatus Lindroth, 1961 i c g b
 Dyschirius planiusculus Putzeys, 1867 c g
 Dyschirius politus (Dejean, 1825) i c g b
 Dyschirius pumilus (Dejean, 1825) i c g b
 Dyschirius punctatus (Dejean, 1825) c g
 Dyschirius pusillus (Dejean, 1825) c g
 Dyschirius quadrimaculatus Lindroth, 1961 i c g b
 Dyschirius recurvus Putzeys, 1867 c g
 Dyschirius reitteri Kult, 1949 c g
 Dyschirius remotepunctatus Putzeys, 1867 c
 Dyschirius roubali Maran, 1938 c g
 Dyschirius rufimanus A.Fleischer, 1898 c g
 Dyschirius rufipes (Dejean, 1825) c g
 Dyschirius rugifer Putzeys, 1878 c g
 Dyschirius ruthmuellerae Bulirsch, 2009
 Dyschirius sabahensis Bulirsch, 2009
 Dyschirius safraneki (Bulirsch & Fedorenko, 2013) c g
 Dyschirius salinus Schaum, 1843 c g
 Dyschirius salivagans Leconte, 1875
 Dyschirius saudiarabicus Balkenohl, 1994 c g
 Dyschirius schaumii Putzeys, 1867 c g
 Dyschirius scriptifrons A.Fleischer, 1898 c g
 Dyschirius sculptus Bousquet, 1988 i c g
 Dyschirius sellatus Leconte, 1857
 Dyschirius selvas (Fedorenko, 1999) c g
 Dyschirius semistriatus (Dejean, 1825) c g
 Dyschirius senegalensis Bruneau de Miré, 1952 c g
 Dyschirius setosus Leconte, 1857
 Dyschirius sevanensis Iablokoff-Khnzorian, 1962 c g
 Dyschirius sextoni Bousquet, 1987 i c g
 Dyschirius shaumii Putzeys, 1866
 Dyschirius similatus Betta, 1847 c
 Dyschirius sjoestedti G.Muller, 1935
 Dyschirius sjostedti G.Müller, 1935 c g
 Dyschirius smetanai (Bulirsch, 2011) c g
 Dyschirius smyrnensis (Fedorenko, 1996) c g
 Dyschirius snizeki (Bulirsch, 2011) c g
 Dyschirius soda Dajoz, 2004
 Dyschirius sonamargensis Balkenohl, 1994 c g
 Dyschirius speculifer Andrewes, 1929 c g
 Dyschirius sphaericollis (Say, 1823) i c g b
 Dyschirius sphaerulifer Bates, 1873 c g
 Dyschirius stellula Andrewes, 1936 c g
 Dyschirius steno Bates, 1873 c g
 Dyschirius stenoderus Putzeys, 1873 c g
 Dyschirius strumosus Erichson, 1837 c g
 Dyschirius subarcticus Lindroth, 1961 i c g
 Dyschirius subcylindricus Motschulsky, 1849
 Dyschirius sublaevis Putzeys, 1846 i c g b
 Dyschirius substriatus (Duftschmid, 1812) c g
 Dyschirius syriacus Putzeys, 1867 c g
 Dyschirius szeli Bulirsch, 2006 c g
 Dyschirius tamil Andrewes, 1929 c g
 Dyschirius tardipes Gistel, 1857 c
 Dyschirius tenuescens Andrewes, 1929 c g
 Dyschirius tenuispinus Lindroth, 1961 i c g b
 Dyschirius terminatus Leconte, 1848
 Dyschirius thailandicus (Fedorenko, 1996) c g
 Dyschirius timidus Lindroth, 1961 i c g
 Dyschirius tonkinensis (Bulirsch & Fedorenko, 2013) c g
 Dyschirius tricuspis Andrewes, 1929 c g
 Dyschirius tridentatus Leconte, 1852
 Dyschirius tristis Stephens, 1827 c g
 Dyschirius truncatus Leconte, 1857
 Dyschirius unipunctatus Fall, 1901 i c g
 Dyschirius ussuriensis Fedorenko, 1991 c g
 Dyschirius vadoni Jeannel, 1946 c g
 Dyschirius vanhillei Basilewsky, 1962
 Dyschirius varidens Fall, 1910 i c g b
 Dyschirius verticalis Putzeys, 1878 c g
 Dyschirius vietnamicus (Fedorenko, 2000) c g
 Dyschirius wayah Dajoz, 2005
 Dyschirius weyrauchi Kult, 1950 c g
 Dyschirius yezoensis Bates, 1883 c g
 Dyschirius zambesiensis (Fedorenko, 2000) c g
 Dyschirius zanzibaricus Chaudoir, 1878 c g

Referanser

Litteratur 
 Bertram G. Rye. 1908. Biller I. Løpebiller. Danmarks Fauna Bind 3. G.E.C. GAD, København.

Eksterne lenker 
 
  Dyschirius – Database Carabidae of the World – engelskspråklig
  Dyschirius - Fauna Europaea - engelskspråklig
 Norsk Entomologisk forening 

graveløpere
Biller formelt beskrevet i 1810