EXHIBIT 21

SUBSIDIARIES OF APPLERA CORPORATION



|                                                    |  |                 |  |                                                       
|--------------------------------------------------- |  | --------------- |  | ------------------------------------------------------
|Name                                                |  |                 |  | State or Jurisdiction of Incorporation or Organization
|Applera Charitable Foundation                       |  | Delaware, USA  
|Applera Overseas Corporation                        |  | New York, USA  
|Applied Biosystems Pty Ltd.                         |  | Australia      
|Applied Biosystems (Canada) Limited                 |  | Canada         
|Applied Biosystems/MDS SCIEX Instruments (1)        |  | Canada         
|Applied Biosystems (Thailand) Limited               |  | Thailand       
|PE AG                                               |  | Switzerland    
|Applera France S.A.                                 |  | France         
|PE (Sweden) AB                                      |  | Sweden         
|PE Stockholm AB                                     |  | Sweden         
|Applied Biosystems Finland OY                       |  | Finland        
|Applera Holding BV                                  |  | The Netherlands
|Applera Finance BV                                  |  | The Netherlands
|Applera Europe BV                                   |  | The Netherlands
|Applied Biosystems Holdings Limited                 |  | UK             
|Applied Biosystems Ltd                              |  | UK             
|PE (GB) Ltd.                                        |  | UK             
|Applera Polska Sp.zo.o.                             |  | Poland         
|Applera Magyarorszag Kft (2)                        |  | Hungary        
|Applera Ceska Republica s.r.o.                      |  | Czech Republic 
|Spartan Ltd. (3)                                    |  | Channel Isles  
|Listronagh Company (4)                              |  | Ireland        
|Applied Biosystems Asia Pte. Ltd.                   |  | Singapore      
|Applied Biosystems Malaysia Sdn. Bhd.               |  | Malaysia       
|Applera Deutschland GmbH                            |  | Germany        
|Applera South Africa (Pty.) Limited                 |  | South Africa   
|PE Manufacturing GmbH (5)                           |  | Germany        
|BSW Wohnstatten GmbH                                |  | Germany        
|Applied Biosystems Manufacturing GmbH               |  | Germany        
|Applera Austria Handels GmbH                        |  | Austria        
|Applied Biosystems Hong Kong, Limited               |  | Hong Kong      
|Applied Biosystems do Brasil Ltda. (6)              |  | Brazil         
|ZAO PE Biosystems (7)                               |  | Russia         
|Applied Biosystems Korea LLC (8)                    |  | Korea          
|Applied Biosystems Taiwan Corporation               |  | Delaware, USA  
|Applied Biosystems de Mexico S. de R.L. de C.V. (9) |  | Mexico         
|Applied Biosystems Trading (Shanghai) Company Ltd.  |  | China          
|Applera Insurance Company Limited                   |  | Bermuda        
|Applera Hispania SA                                 |  | Spain          
|Applera International, Inc.                         |  | Delaware, USA  
|PE Korea Corporation                                |  | Delaware, USA  
|Applied Biosystems China, Inc.                      |  | Delaware, USA  
|GenScope, Inc.                                      |  | Delaware, USA  
|PE AgGen, Inc.                                      |  | Utah, USA      
|PerSeptive Biosystems, Inc.                         |  | Delaware, USA  
|Applied Biosystems Japan, Ltd.                      |  | Japan          
|PerSeptive Biosystems (Canada) Ltd.                 |  | Canada         
|PNA Diagnostics ApS                                 |  | Denmark        





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|                                         |  |                
|---------------------------------------- |  | ---------------
|Boston Probes, Inc. (10)                 |  | Delaware, USA  
|Paracel, Inc.                            |  | California, USA
|Paracel Government Systems, Inc.         |  | Delaware, USA  
|Axys Pharmaceuticals, Inc.               |  | Delaware, USA  
|Axys 468 Littlefield LLC                 |  | California, USA
|Foster City Holdings, LLC                |  | Delaware, USA  
|Celera Diagnostics, LLC                  |  | Delaware, USA  
|Rockville Holdings, LLC                  |  | Delaware, USA  
|Celera Corporation                       |  | Delaware, USA  
|Ambion, Inc.                             |  | Delaware, USA  
|Ambion Europe, Limited                   |  | UK             
|Ambion (UK) Limited                      |  | UK             
|AB Advanced Genetic Analysis Corporation |  | Delaware, USA  

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Note: Entities directly owned by subsidiaries of
Applera Corporation are indented and listed below their immediate parent. Ownership is 100% unless otherwise indicated.



|(1) | 50.0% ownership.
|--- | ----------------

|(2) | 90.0% owned by Applera Holding BV and 10.0% by Applera Finance BV (indirectly, in the aggregate, wholly owned by Applera Corporation).
|--- | --------------------------------------------------------------------------------------------------------------------------------------

|(3) | 51.0% owned by Applera Overseas Corporation and 49.0% by Applera Europe BV (indirectly, in the aggregate, wholly owned by Applera Corporation).
|--- | -----------------------------------------------------------------------------------------------------------------------------------------------

|(4) | 3.9% owned by PE (Sweden) AB, 23.5% by Applera Holding BV, 71.9% by Spartan Limited, and 0.7% by Applera Overseas Corporation (indirectly, in the aggregate, wholly owned by Applera
Corporation).
|--- | --------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------

|(5) | 98.8% owned by Applera Deutschland GmbH, 0.5% by Applera Overseas Corporation, and 0.7% by Listronagh Company (indirectly, in the aggregate, wholly owned by Applera Corporation).
|--- | ----------------------------------------------------------------------------------------------------------------------------------------------------------------------------------

|(6) | .01% owned by Charles J. Heinzer, an Applera employee, and 99.99% by Applera Overseas Corporation (directly and indirectly, in the aggregate, wholly owned by Applera Corporation).
|--- | -----------------------------------------------------------------------------------------------------------------------------------------------------------------------------------

|(7) | 0.1% owned by Applera Corporation and 99.9% by Applera Overseas Corporation (directly and indirectly, in the aggregate, wholly owned by Applera Corporation).
|--- | -------------------------------------------------------------------------------------------------------------------------------------------------------------

|(8) | 20.0% owned by Applera Corporation and 80.0% by Applera Overseas Corporation (directly and indirectly, in the aggregate, wholly owned by Applera Corporation).
|--- | --------------------------------------------------------------------------------------------------------------------------------------------------------------

|(9) | .01% owned by Applera Corporation and 99.99% by Applera Overseas Corporation (directly and indirectly, in the aggregate, wholly owned by Applera Corporation).
|--- | --------------------------------------------------------------------------------------------------------------------------------------------------------------

|(10) | 84.9% owned by Applera Corporation and 15.1% by PNA Diagnostics ApS (directly and indirectly, in the aggregate, wholly owned by Applera Corporation).
|---- | -----------------------------------------------------------------------------------------------------------------------------------------------------

Applera Corporation conducts business through its Applied Biosystems group and Celera group. Also, we previously operated a Celera Diagnostics business as a joint
venture between the Applied Biosystems group and the Celera group. During our 2006 fiscal year, Celera Diagnostics became part of the Celera group pursuant to a restructuring of the Celera Diagnostics business. Applera Corporation and its direct and
indirect subsidiaries conduct business under the names of these businesses, its corporate name, and variants thereof. The Applied Biosystems/MDS SCIEX Instruments partnership conducts business under its partnership name and variants thereof.