**EXHIBIT 21**

**SUBSIDIARIES OF APPLERA CORPORATION**

| **Name**                                            | **State or Jurisdiction of****Incorporation or Organization****** |
|----------------------------------------------------------|-------------------------------------------------------------------|
| Applera Charitable Foundation                            | Delaware, USA                                                     |
| Applera Overseas Corporation                             | New York, USA                                                     |
| Applied Biosystems Pty Ltd.                              | Australia                                                         |
| Applied Biosystems (Canada) Limited                      | Canada                                                            |
| Applied Biosystems/MDS SCIEX Instruments (1)             | Canada                                                            |
| Applied Biosystems (Thailand) Limited                    | Thailand                                                          |
| PE AG                                                    | Switzerland                                                       |
| Applera France S.A.                                      | France                                                            |
| PE (Sweden) AB                                           | Sweden                                                            |
| PE Stockholm AB                                          | Sweden                                                            |
| Applied Biosystems Finland OY                            | Finland                                                           |
| Applera Holding BV                                       | The Netherlands                                                   |
| Applera Finance BV                                       | The Netherlands                                                   |
| Applera Europe BV                                        | The Netherlands                                                   |
| Applied Biosystems Holdings Limited                      | UK                                                                |
| Applied Biosystems Ltd                                   | UK                                                                |
| PE (GB) Ltd.                                             | UK                                                                |
| Applera Polska Sp.zo.o.                                  | Poland                                                            |
| Applera Magyarorszag Kft (2)                             | Hungary                                                           |
| Applera Ceska Republica s.r.o.                           | Czech Republic                                                    |
| Spartan Ltd. (3)                                         | Channel Isles                                                     |
| Listronagh Company (4)                                   | Ireland                                                           |
| Applied Biosystems Asia Pte. Ltd.                        | Singapore                                                         |
| Applied Biosystems Malaysia Sdn. Bhd.                    | Malaysia                                                          |
| Applera Holding GmbH                                     | Germany                                                           |
| Applera South Africa (Pty.) Limited                      | South Africa                                                      |
| PE Manufacturing GmbH (5)                                | Germany                                                           |
| BSW Wohnstatten GmbH                                     | Germany                                                           |
| Applied Biosystems Manufacturing GmbH                    | Germany                                                           |
| Applera Austria Handels GmbH                             | Austria                                                           |
| Applied Biosystems Hong Kong, Limited                    | Hong Kong                                                         |
| Applied      Biosystems do Brasil Ltda.  (6)             | Brazil                                                            |
| ZAO      PE Biosystems (7)                               | Russia                                                            |
| Applied      Biosystems Korea LLC (8)                    | Korea                                                             |
| Applied Biosystems Taiwan Corporation                    | Delaware, USA                                                     |
| Applied      Biosystems de Mexico S. de R.L. de C.V. (9) | Mexico                                                            |
| Applied Biosystems Trading (Shanghai) Company Ltd        | China                                                             |
| Applera Insurance Company Limited                        | Bermuda                                                           |
| PE FSC, Inc.                                             | U.S. Virgin Islands                                               |
| Applera Hispania SA                                      | Spain                                                             |
| Applera International, Inc.                              | Delaware, USA                                                |
| PE Korea Corporation                                     | Delaware, USA                                                     |
| Applied Biosystems China, Inc.                           | Delaware, USA                                                     |
| GenScope, Inc.                                           | Delaware, USA                                                     |
| PE AgGen, Inc.                                           | Utah, USA                                                         |
| PerSeptive Biosystems, Inc.                              | Delaware, USA                                                     |
| Applera Deutschland GmbH                                 | Germany                                                           |
| Applied Biosystems Japan, Ltd.                           | Japan                                                             |
| PerSeptive Biosystems (Canada) Ltd.                      | Canada                                                            |

 

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| PNA Diagnostics ApS                      | Denmark         |
|------------------------------------------|-----------------|
| Boston Probes, Inc. (10)                 | Delaware, USA   |
| Paracel, Inc.                            | California, USA |
| Paracel Government Systems, Inc.         | Delaware, USA   |
| Axys Pharmaceuticals, Inc.               | Delaware, USA   |
| Axys 468 Littlefield LLC                 | California, USA |
| Foster City Holdings, LLC                | Delaware, USA   |
| Rockville Holdings, LLC                  | Delaware, USA   |
| Celera Diagnostics, LLC                  | Delaware, USA   |
| Celera Corporation                       | Delaware, USA   |
| Ambion, Inc.                             | Delaware, USA   |
| Ambion Europe, Limited                   | UK              |
| Ambion (UK) Limited                      | UK              |
| Ambion K.K.                              | Japan           |
| AB Advanced Genetic Analysis Corporation | Delaware, USA   |

**_______________________________________**

Note: Entities directly owned by subsidiaries of Applera Corporation are indented and listed below their immediate parent.  Ownership is 100% unless otherwise indicated.

| (1)  | 50.0% ownership.                                                                                                                                                                                             |
|------|--------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------|
| (2)  | 90.0% owned by Applera Holding BV and 10.0%        by Applera Finance BV (indirectly, in the aggregate, wholly owned by    Applera Corporation).                                                             |
| (3)  | 51.0% owned by Applera Overseas Corporation        and 49.0% by Applera Europe BV (indirectly, in the aggregate, wholly    owned by Applera Corporation).                                                    |
| (4)  | 3.9% owned by PE (Sweden) AB, 23.5% by Applera        Holding BV, 71.9% by Spartan Limited, and 0.7% by Applera Overseas Corporation    (indirectly, in the aggregate, wholly owned by Applera Corporation). |
| (5)  | 98.8% owned by Applera Holding GmbH, 0.5%        by Applera Overseas Corporation, and 0.7% by Listronagh Company (indirectly,    in the aggregate, wholly owned by Applera Corporation).                     |
| (6)  | .01% owned by Charles J. Heinzer and 99.99%        by Applera Overseas Corporation (directly and indirectly, in the aggregate,        wholly owned by Applera Corporation).                                  |
| (7)  | 0.1% owned by Applera Corporation and 99.9%        by Applera Overseas Corporation (directly and indirectly, in the aggregate,    wholly owned by Applera Corporation).                                      |
| (8)  | 20.0% owned by Applera Corporation and 80.0%        by Applera Overseas Corporation (directly and indirectly, in the aggregate,    wholly owned by Applera Corporation).                                     |
| (9)  | .01% owned by Applera Corporation and 99.99%        by Applera Overseas Corporation (directly and indirectly, in the aggregate,    wholly owned by Applera Corporation).                                     |
| (10) | 84.9% owned by Applera Corporation, and 15.1%        by PNA Diagnostics ApS (directly and indirectly, in the aggregate, wholly    owned by Applera Corporation).                                             |

Applera Corporation conducts business    through its Applied Biosystems group and Celera Genomics group. Also, we    previously operated a Celera Diagnostics business as a joint venture between    the Applied Biosystems group and the Celera Genomics group. During our 2006    fiscal year, Celera Diagnostics became part of the Celera Genomics group    pursuant to a reorganization described in our 2006 Annual Report on Form    10-K. Applera Corporation and its direct and indirect subsidiaries conduct    business under the names of these businesses and variants thereof. The Applied    Biosystems/MDS SCIEX Instruments partnership conducts business under its    partnership name and variants thereof.

 

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