Exhibit 21



SUBSIDIARIES OF APPLIED BIOSYSTEMS INC.



|                                                               |  |                              
|-------------------------------------------------------------- |  | -----------------------------
|                                                               |  | State or Jurisdiction of     
|Name                                                           |  | Incorporation or Organization
|                                                               | 
|Applera Charitable Foundation                                  |  | Delaware, USA                
|Applied Biosystems Overseas Corporation                        |  | New York, USA                
|Applied Biosystems Pty Ltd.                                    |  | Australia                    
|Applied Biosystems (Canada) Limited                            |  | Canada                       
|Applied Biosystems/MDS Analytical Technologies Instruments (1) |  | Canada                       
|Applied Biosystems (Thailand) Limited                          |  | Thailand                     
|PE AG (9)                                                      |  | Switzerland                  
|Applera France S.A.                                            |  | France                       
|PE (Sweden) AB (9)                                             |  | Sweden                       
|PE Stockholm AB (9)                                            |  | Sweden                       
|Applied Biosystems Finland OY (9)                              |  | Finland                      
|Applied Biosystems B.V.                                        |  | The Netherlands              
|Applied Biosystems Finance BV                                  |  | The Netherlands              
|Applied Biosystems Europe BV                                   |  | The Netherlands              
|Applied Biosystems Holdings Limited (9)                        |  | UK                           
|Applied Biosystems Ltd                                         |  | UK                           
|PE (GB) Ltd. (9)                                               |  | UK                           
|Applera Polska Sp.zo.o.                                        |  | Poland                       
|Applera Magyarorszag Kft (2)                                   |  | Hungary                      
|Applera Ceska Republica s.r.o.                                 |  | Czech Republic               
|Applied Biosystems Asia Pte. Ltd.                              |  | Singapore                    
|Applied Biosystems Malaysia Sdn. Bhd.                          |  | Malaysia                     
|Applera Deutschland GmbH                                       |  | Germany                      
|Applera South Africa (Pty.) Limited                            |  | South Africa                 
|PE Manufacturing GmbH (3) (9)                                  |  | Germany                      
|BSW Wohnstatten GmbH (9)                                       |  | Germany                      
|Applied Biosystems Manufacturing GmbH (9)                      |  | Germany                      
|Applera Austria Handels GmbH                                   |  | Austria                      
|Applied Biosystems Hong Kong, Limited                          |  | Hong Kong                    
|Applied Biosystems do Brasil Ltda. (4)                         |  | Brazil                       
|ZAO PE Biosystems (5) (9)                                      |  | Russia                       
|Applied Biosystems Korea LLC (6)                               |  | Korea                        
|Applied Biosystems Taiwan Corporation                          |  | Delaware, USA                
|Applied Biosystems de Mexico S. de R.L. de C.V. (7)            |  | Mexico                       
|Applied Biosystems Trading (Shanghai) Company Ltd.             |  | China                        
|Applied Biosystems Insurance Company Limited                   |  | Bermuda                      
|Applera Hispania SA                                            |  | Spain                        
|Applied Biosystems International, Inc.                         |  | Delaware, USA                
|PE Korea Corporation                                           |  | Delaware, USA                
|Applied Biosystems China, Inc.                                 |  | Delaware, USA                
|PerSeptive Biosystems, Inc.                                    |  | Delaware, USA                
|Applied Biosystems Japan, Ltd.                                 |  | Japan                        
|PerSeptive Biosystems (Canada) Ltd.                            |  | Canada                       
|PNA Diagnostics ApS                                            |  | Denmark                      
|Boston Probes, Inc. (8)                                        |  | Delaware, USA                



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|                                         |  |              
|---------------------------------------- |  | -------------
|Ambion, Inc.                             |  | Delaware, USA
|Ambion Europe, Limited                   |  | UK           
|AB Advanced Genetic Analysis Corporation |  | Delaware, USA



Note:    Entities directly owned by subsidiaries of Applied Biosystems Inc. are indented and listed below their immediate parent. Ownership is 100% unless otherwise indicated.



|(1) | 50.0% ownership.
|--- | ----------------

|(2) | 90.0% owned by Applied Biosystems B.V. and 10.0% by Applied Biosystems Finance BV (indirectly, in the aggregate, wholly owned by Applied Biosystems Inc.).
|--- | ----------------------------------------------------------------------------------------------------------------------------------------------------------

|(3) | 98.8% owned by Applera Deutschland GmbH, 0.5% by Applied Biosystems Overseas Corporation, and 0.7% by Applied Biosystems Europe B.V. (indirectly, in the aggregate, wholly owned by
Applied Biosystems Inc.).
|--- | -------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------

|(4) | .01% owned by Charles J. Heinzer, an Applied Biosystems Inc. employee, and 99.99% by Applied Biosystems Overseas Corporation (directly and indirectly, in the aggregate, wholly owned
by Applied Biosystems Inc.).
|--- | ------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------

|(5) | 0.1% owned by Applied Biosystems Inc. and 99.9% by Applied Biosystems Overseas Corporation (directly and indirectly, in the aggregate, wholly owned by Applied Biosystems Inc.).
|--- | --------------------------------------------------------------------------------------------------------------------------------------------------------------------------------

|(6) | 20.0% owned by Applied Biosystems Inc. and 80.0% by Applied Biosystems Overseas Corporation (directly and indirectly, in the aggregate, wholly owned by Applied Biosystems Inc.).
|--- | ---------------------------------------------------------------------------------------------------------------------------------------------------------------------------------

|(7) | .01% owned by Applied Biosystems Inc. and 99.99% by Applied Biosystems Overseas Corporation (directly and indirectly, in the aggregate, wholly owned by Applied Biosystems Inc.).
|--- | ---------------------------------------------------------------------------------------------------------------------------------------------------------------------------------

|(8) | 84.9% owned by Applied Biosystems Inc., and 15.1% by PNA Diagnostics ApS (directly and indirectly, in the aggregate, wholly owned by Applied Biosystems Inc.).
|--- | --------------------------------------------------------------------------------------------------------------------------------------------------------------

|(9) | This entity is considered dormant and/or has no material operations/assets.
|--- | ---------------------------------------------------------------------------

Applied Biosystems Inc. and its direct and indirect subsidiaries conduct business under the Applied Biosystems name and variants thereof, and similarly subsidiaries may conduct business under their entity name or variants thereof. The
Applied Biosystems/MDS Analytical Technologies Instruments partnership conducts business under its partnership name and variants thereof.