CE T. JAME A Partial Revision o

the Oriental Isomyia

, of the Viridaurea Group

(Diptera: Calliphoridae)

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY • 1970 NUMBER 67
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SMITHSONIAN CONTRIBUTIONS TO

ZOOLOGY

NUMBER 67

Maurice T. james A Partial Revision of

the Oriental Isomyia

of the Viridaurea Group

(Diptera: Calliphoridae)

SMITHSONIAN INSTITUTION PRESS

CITY OF WASHINGTON

1970
ABSTRACT

Maurice T. James. A Partial Revision of the Oriental Isomyia of the Viridaurea

Group (Diptera: Calliphoridae). Smithsonian Contributions to Zoology, 67: 1-14,

1970.—The group is defined and, except for some species of the delectans group,

reviewed. A key to species is presented. The following new species are described:

Isomyia lugubris and /. facialis from Thailand; /. chalconotum from Malaysia;

/. marginata, I. iris, and /. discalis from Philippine Islands; and /. aurifacies and

/. perisi from India. New combinations are /. gomezmenori (Peris), /. ceballosi (Peris),

and /. borneensis (Peris), all from Thelychaeta, and /. pseudolucilia (Malloch) from

Strongyloneura. A lectotype is designated for Thelychaeta chalybea Brauer and

Bergenstamm, a synonym of Isomyia viridaurea (Wiedemann).

Scientific Paper 3366, College of Agriculture,

Washington State University; work conducted under Project9043.

Official publication date is handstamped in a limited number of initial copies and is recorded

in the Institution's annual report, Smithsonian Year.

UNITED STATES GOVERNMENT PRINTING OFFICE

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Washington, D.C. 20402 - Price 25 cents (paper cover)
Maurice T. James A Partial Revision of

the Oriental Isomyia

of the Viridaurea Group

(Diptera: Calliphoridae)

The Oriental species of Isomyia have been studied by

several workers, the most comprehensive treatment

being that of Peris (1952). Peris used the generic name

Thelychaeta, but acknowledged the fact that Isomyia

had priority. He divided the genus into several groups,

one of which, the viridaurea group, was characterized

principally by having the thoracic squama lobulate,

that is, produced inwardly so that its inner margin

comes into contact with the scutellum. Some members

of the dotata group share this character but they are

distinguished from viridaurea and its allies by having

the stem vein or R^ (Peris says the subcosta) setulose

below, whereas this vein is bare below in the viridaurea

group. Peris refers one species, I. chrysoides (Walker),

to this group, even though its thoracic squama is not

of the type described above, on the basis of its overall

similarity to other members of the group, and I concur

with his opinion in this respect.

Malloch (1928), in his study of what he called

Strongyloneura, had laid some foundation for Peris'

grouping by pointing out the significance of the

squamal structure and the ventral setation of vein Mi.

He called what is essentially the viridaurea group his

group B, with the statement that it was "probably

equivalent to Thelychaeta, B. & B." Senior-White,

Aubertin, and Smart (1940) followed Malloch in the

essential aspects of their key but did not give any

grouping of the species; they used the generic name

Strongyloneura, though they indicated in a footnote

that the proper name was Isomyia. They included /.

chrysoides (Walker), along with /. viridaurea (WiedeMaurice T. James, Professor of Entomology, Washington

State University, Pullman, Washington 99163.

mann), /. pseudolucilia (Malloch), /. oestracea

(Seguy), and /. delectans (Walker), under the section of the key defined by the inwardly projecting

thoracic squama. Seguy expanded this section, which

he called the genus Isomyia, by the inclusion of four

more of his species; he retained I. chrysoides here, without explanation.

Peris was unable to deal satisfactorily with a complex which includes /. delectans (Walker), and I have

been but little more successful. A considerable amount

of material in this species complex and a careful study

of available types will be necessary before this problem can be solved. The viridaurea group apparently

is rich in species, and the present paper, with the key

it contains, can be considered only a contribution

toward the taxonomy of the group. The necessity of

describing so many species from single specimens is

unfortunate, but in all cases these species are so well

marked that this course is justified.

The type-species of Thelychaeta Brauer and Bergenstamm, 1891, is T. chalybea Brauer and Bergenstamm, a synonym of Musca viridaurea Wiedemann.

That of Isomyia Walker, 1860, is Musca delectans

Walker. Both nominal genera, therefore, are based on

species in the viridaurea group. Strongyloneura Bigot,

1886, as restricted by Peris, is quite a different group,

based on S. prasina Bigot as its type-species, and certainly is not congeneric with the Isomyia species.

Acknowledgments

This study was based on material from several collections, particularly those of the National Museum of

Natural History; British Museum (Natural History),

1
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

from the Malay States Museum; Natural History

Museum of Vienna; Michigan State University; and

Washington State University. Most holotypes and allotypes are in the National Museum of Natural History

(USNM). Paratypes of all species other than those described from unique specimens are in the Washington

State University collection; the holotype and allotype

of /. perisi are being donated to the National Museum

of Natural History, and one paratype of that species is

being given to the British Museum (Natural History);

the holotype and allotype of /. chalconotum are in

the British Museum (Natural History).

Morphology

Most terms used here are more or less standard, but

some require explanation. The term "face" is applied

to the area between the antennal bases and the oral

margin, and between the facials or facial ridges; it

includes what Townsend calls the epistoma, but which

actually is not that structure. No description is included

for the palpi and proboscis; they do not vary significantly in color or structure. The palpi are always

spatulate, rather inflated, and orange to orange-yellow.

The antennae vary little in structure; the arista is set

with long rays, longer above than below. The antennae

are separated from each other at the base, and usually

there is a more or less well-developed facial carina.

Thoracic chaetotaxy seems to be subject to some variation. An accessory propleural and an accessory stigmatal, in every case shorter and weaker than the true

propleural or stigmatal, may be present. Some variation

of significance occurs in the pteropleural(s) ; this is

indicated in the descriptions. There are three or four

strong lateral scutellars, the number being of definite

taxonomic significance; this count does not include a

strong setula or weak bristle (always present) basad

of the basal lateral scutellar. The term setula is applied

to stout, bristle-like hairs; the softer vestiture is designated as composed of hairs. A group of setulae occurs

below the notopleural suture, on the anterior part of

the mesopleuron; this group of setulae usually is clearly

differentiable from the rest of the vestiture of the

mesopleuron and is characteristic of the genus. A row

of bristles occurs on the posterior margin of the

mesopleuron, but in addition to this, and rising slightly

behind it, is a row of long hairs, about as long as the

bristles and lying parallel to them, of texture more like

setae but usually with some softer hairs intermixed.

This row of hairs is designated as the posterior

mesopleural fringe.

The apparent, rather than the actual, segmentation

of the abdomen is used; the first tergum, therefore,

actually is the second morphological tergum. The first

two sterna are evident. The structure of the male

genitalia shows only moderate variation within this

group, and genital structures are illustrated whenever

dissections are made. Measurements, except overall

length, are in micrometer units (60 = 1 mm). Unless

otherwise indicated, measurements are of the holotype

or allotype.

Taxonomy

Key to the Oriental species of the Isomyia viridaurea group

1. Lunulc with several short black setulae 2

Lunule bare of setulae; fourth abdominal tergum, if densely pollinose, not contrasting with

preceding terga either in pollen or in coloration; color predominantly green or blue.... 3

2. Color overall black with purplish reflections; pollen of mesonotum and of dorsal aspect of

abdomen cinereous, rather uniform in density except on posterior margins of abdominal

terga, where it is inconspicuous, nowhere obscuring ground color.. I. lugubris, new species

Fourth abdominal tergum green dorsally, with dense, ashy pollen which, at least from

posterior view, largely conceals background, preceding terga dark blue to purple and

almost devoid of conspicuous pollen; mesonotum and, to a lesser extent, pleura deep

coppery (female) or green with coppery reflections (male), in striking contrast to the

dark blue to purple first three abdominal terga /. chalconotum, new species

3. Pile of mesopleuron, and of other pleural areas as well, soft and yellow to golden, except for

usual black setulae below notopleural suture; covering of anterior spiracle yellow 4

Pile of pleura more extensively black than indicated above, with some soft black hairs on

mesopleuron, sometimes remote from notopleural suture, and on sternopleuron 8

4. Basioosta bright yellow 5

Basicosta brown to black; abdomen violet, intermediate terga with broad black margins,

thorax blue with violet reflections on mesonotum and scutellum../. marginata, new species
NUMBER 6 7

5. Thoracic squama not tabulate; facets of eye of male strikingly larger above than below; pollen

of abdomen ordinary, usually not visible except from posterior view.

/. chrysoides (Walker)

Thoracic squama lobulate; facets of eye of male larger above than below but not

strikingly so 6

6. Pleura and abdomen densely pollinose in male, less so in female, but dorsal parts of fourth

tergum, when viewed laterally at an angle, with a tessellated pattern of pollen which,

in the proper light, obscures, or virtually obscures, the background; black bristles of

lateral slopes of first tergum surrounded, at least on three sides, by pale hairs.

/. viridaurea (Wiedemann)

Pleura and abdomen less densely pollinose and without a tessellated pattern; black bristles of

lateral slopes of first tergum with some black hairs in front and behind as well as above... 7

7. First abdominal tergum with black hairs at base; eyes of male separated by much less than

width of ocellar triangle, the frontale interrupted at narrowest part of frons.

/. gotnezmenori (Peris)

First abdominal tergum with yellow hairs at base; eyes of male separated by about width of

ocellar triangle, frontale evident /. pichoni (S6guy)

8. Four strong lateral scutellars, the last three spaced at closer intervals than the basal one.... 9

Three strong lateral scutellars, spaced at approximately equal intervals 12

9. Both squamae pale yellow; posterior mesopleural fringe yellow; some pile on notopleuron,

lower part of mesopleuron, and part of stemopleuron yellow to yellowish.

/. pseudolucilia (Malloch)

At least thoracic squama mostly brown; pile of notopleuron and of pleura wholly black

or dark brown 10

10. Gena, parafrontal, and parafacial black, lower part of parafacial and part of gena devoid of

pollen and shining; squamae whitish on basal part /. ceballosi (Peris)

Gena golden to brownish yellow, it and parafacial wholly pollinose; squamae entirely

brown to blackish brown 11

11. Lunule entirely pollinose; lunule, parafrontal, parafacial, and gena uniformly golden

brown pollinose. /. aurifacies, new species

Lunule shining anteriorly; pollen of parafrontal and parafacial not uniformly colored,

cinereous to brownish /. perisi, new species

12. Disc of mesonotum largely black, contrasting to lateral slopes and prescutellar area of

mesonotum, which, along with the scutellum, are bluish green or purple 13

Disc of mesonotum not conspicuously bicolored 14

13. Pleural pile wholly black to dark reddish brown, dense and long; squamae wholly brown;

a robust, purplish species /. iris, new species

Pile of pleura posterior to and including posterior mesopleural fringe yellow to golden, pile

much less dense than in I. iris; lower part of thoracic squama white, squamae otherwise

brown; a more slender, blue green species /. discalis, new species

14. Pleura with extensive yellow hair, at least around stigmatal and propleural bristles and on

part of stemopleuron; posterior mesopleural fringe golden 15

Pleural hairs entirely black or virtually so; posterior mesopleural fringe brown to black.... 17

15. Hairs covering anterior spiracle bright yellow to golden 16

Hairs covering anterior spiracle dark brown to black (some other species of the delectans

complex may run here) /. delectans (Walker)

16. Tibiae yellow; parafrontal in female subequal to frontale in width at middle of frons; parafacial in male almost twice as wide as antennal flagellum /. electa (Villeneuve)

Tibiae brown; parafrontal in female about half as wide as frontale; parafacial in male about

1.5 times as wide as antennal flagellum /• borneensis (Peris)

17. Upper and lower squamae wholly dark brown to black 18

Upper and lower squamae white, at least at base 19

18. Face dark brown to black, especially above vibrissae /. pictifacies palawanensis James

Face bright yellow to orange yellow part of the delectans complex

19. Upper squama, in both sexes, white on basal half or more; lower squama white at base;

larger species, 10.0-11.0 mm in length '• pictifacies (Bigot)

Upper and lower squamae wholly white in female, upper one white only anteroventrally in

male; smaller species, 7.0-9.0 mm in length /• facialis, new species
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Isomyia lugubris, new species

FEMALE.—Parafacials, parafrontals, occiput, ocellar

triangle, and postgena mostly black, covered, except

for glabrous area of occiput, with cinereous pollen,

densest on occipital orbits, on parafacials and parafrontals with a slightly yellowish tinge. An indefinite

area on each parafrontal, side of ocellar triangle, and

upper part of vertex orange yellow; pollen as on parafrontals. Frontale reddish brown, partly with a dull

purplish sheen. Cheek grooves and gena reddish brown,

gena becoming yellow next to black area; pollen of

gena golden, a very small, poorly defined, glabrous spot

at contact with parafacial, below eye. Lunule orangeyellow, largely glabrous but with some pollen visible

from behind; facials and face yellow, uniformly dulled

with white to yellow pollen. Frontale twice or more

width of parafrontal except below, on lower half almost

parallel-sided, bulging slightly at middle, then narrowing to vertex. Bristles, setulae, and hairs of head black

except on occiput, postgena, and gena along oral margin. About 11 frontals; frontal setulae prominent, some

approaching bristles in size; lunule with distinct black

setulae; parafacials with setulae arranged in about

three irregular rows. Antenna orange-yellow, flagellum

becoming clearer yellow at base and brownish yellow

toward apex; flagellum about twice length of pedicel ;

arista brown on basal fifth, yellow on subbasal fifth,

brown on apical three-fifths; rays brownish black.

Measurements in micrometer units: head width, 230;

width of vertex, 58, of frons at base of antenna, 97;

width of frontale at maximum, 38 (comparative width

of parafrontal at same place, 20), at antennal base, 35,

at anterior ocellus, 25; distance between vibrissae, 45;

vibrissa to oral margin, 20; oral margin to lower margin

of lunule, 80; width of parafacial at cheek groove, 20;

width of gena, 38; eye height, 145; antennae separated

by 7; length of pedicel, 17, of flagellum, 32.

Thorax and abdomen black, with dull purplish reflections dorsally and on abdomen ventrally, purplish

to greenish on thoracic pleura; pollen cinereous, of

moderate density, less dense on apical part of scutellum,

mesopleuron and pteropleuron anterior to wing base

and squamae, base of abdomen, and apical transverse

bands on abdominal terga 2 to 4; pollen best visible

from posterior view, but distinctly dulling background

except in sparser areas indicated above; that of abdomen only indistinctly tessellated. Mesonotum, viewed

from behind, with traces of three vittae (a pair of

dorsocentrals, one between acrostichal rows) ; a lateral

vitta below postsutural supra-alars and extending onto

postalar callus, subshining. Setulae of mesonotum,

dorsal surface of scutellum, humerus except below,

mesopleuron, and abdomen except first and second

sterna black; postalar declivity with a few scattered,

yellowish to brownish, as well as blackish, hairs ventrally; posterior mesopleural fringe mixed black and

yellow; hairs of humerus below level of top of anterior

spiracle, sternopleuron, hypopleuron, prosternum, and

first and second abdominal sterna whitish to yellowish,

those of first abdominal sternum long. Coverings of

anterior and posterior spiracles black. Acrostichals,

1, 3; postsuturals located on posterior half of postscutum and continued forward by a row of about three

setulae that are stronger than the others; dorsocentrals,

2, 6; humerals, 3; accessory propleural and accessory

stigmatal present; pteropleurals, about 10, of equal

length but not of equal strength, forming a tuft; scutellars, 3 lateral, 1 discal; abdominal bristles weak, a

transverse row of median discals on fourth tergum;

1 pair apicals on second sternum.

Femora black with a weak greenish reflection; tibiae

reddish brown; basitarsi mostly reddish brown, remainder of tarsi blackish brown. Middle tibia with

strong, long ventral at about two-thirds length. Wing

yellowish hyaline, broadly brown apically, including

area beyond basal third of apical cell, and basally,

basad of furcation of Rs; basicosta black; bend of Mi

rounded, part beyond bend sinuate. Halteres yellow.

Squamae yellow, deeper so along margins; fringes pale

yellow to white. Length, 10 mm.

MALE.—Unknown.

HOLOTYPE.— $ , Thailand (Siam), Kuong Pen.,

Kao Chong, 1,000 feet, July 1933, Hugh M. Smith;

USNM 70751.

REMARKS.—This species belongs in the viridaurea

group as defined by Peris (1952), and, with the present

state of our knowledge, there is no reason to place

it elsewhere. Superficially, it resembles /. tristis (Bigot)

and its relatives. The black coloration of the thorax

and abdomen distinguish it from all other Oriental

members of the genus known to me; the tuft of

pteropleurals and (except for the next species) the

setulae on the lunule are distinctive.

Isomyia chalconotum, new species

FEMALE.—Head mostly black; gena mostly brownish

yellow to brown, becoming clearly yellow posteriorly

and along oral margin; cheek grooves pale reddish
NUMBER 6 7

brown; postgena black; frontale dark reddish brown;

lunule orange-yellow; face yellow; facials grading

from yellow to color of cheek groove and becoming

somewhat darker next to parafacials. Ocellar triangle,

parafrontals, parafacials, genae, cheek grooves, and

occiput, except glabrous area, densely golden pollinose,

color deepest on genae, pollen densest on occipital

orbits; a small subshining spot on gena below eye;

pollen of frontale concolorous with background; lunule

appearing glabrous but, when viewed posteriorly, with

golden pollen except on anterior margin; face more

lightly dusted with whitish pollen but its entire surface

dulled. Frontale about 1.3 to 1.4 times as wide as

parafrontal, gradually widening from anterior ocellus

to antennal base. Bristles, setulae, and hairs of head

mostly black; a few posterior bristles of oral row, also

hairs and setulae on posterior part of gena and lower

part of occiput, yellow to golden. Frontals, 9 to 13;

parafrontal and parafacial setulae short, latter in one

or two irregular rows. Several black setulae on lunule.

Antennae orange, flagellum somewhat brownish above;

arista brown, a subbasal area yellow. Measurements

in micrometer units: head width, 280; width of vertex,

60, of frons at base of antenna, 100; width of frontale

and parafrontal at base of antenna, 40 and 30, at

anterior ocellus, 27 and 17; distance between vibrissae,

60; vibrissa to oral margin, 35; oral margin to lower

margin of lunule, 115; parafacial at cheek groove, 20;

width of gena, 42; eye height, 175; antennae separated

at base by 9; length of pedicel, 22, of flagellum, 32.

Thorax green, but over most of its area so strongly

coppery in reflections as to obscure background except

under special lighting; green color shows best on

humeral slopes, postalar callus, margin of scutellum,

notopleuron and hypopleuron; pollen cinereous, that

of dorsal surface scant, more conspicuous anteriorly

on mesonotum, rather dense on lower part of humerus,

propleuron, lower anterior margin of mesopleuron,

most of sternopleuron, and posterior part of pteropleuron. Setulae of mesonotum, upper part of humerus,

dorsum and sides of scutellum, and upper anterior

margin of mesopleuron black, mesopleural black

setulae about 10 in number, in about two rows next

to notopleural suture. Pleural hairs, including those of

lower half of humerus, golden to yellow; posterior

mesopleural fringe golden. Some yellowish hairs on

ventral side of scutellum and on postalar declivity.

Covering of anterior spiracle yellow, that of posterior

spiracle brown. Thoracic bristles black; acrostichals,

2, 4; dorsocentral, 2, 4; humerals, 3-4; pteropleural,

1; an accessory propleural but no accessory stigmatal;

scutellars, 3 lateral, 1 discal.

Femora black, with green and bronze reflections.

Tibiae and tarsi reddish yellow, tarsi becoming darker

beyond apex of basitarsus. Coxae and femora with

cinereous pollen. Bristles and setulae of legs black.

Middle tibia with strong ventral at about two-thirds

length. Wing wholly brown; basicosta bright yellow;

Mi strongly bent but not angulate. Halteres, squamae,

and their hairs deep golden.

Abdomen blue-green (first tergum) or purple

(second and third terga) dorsally, without apparent

pale pollen, though some can be seen at a very oblique

angle; fourth tergum, in strong contrast, densely

cinereous pollinose, pollen obscuring background over

most of tergum regardless of light incidence; background quite evidently green at base of fourth tergum,

apparently purplish elsewhere. Apices of second and

third terga black banded. Ventral slopes of terga concolorous laterally with dorsal surfaces but second to

fourth largely coppery; on second and third terga

ventral surfaces merge from purple to green to copper.

Ventral faces of abdominal terga with conspicuous

cinereous pollen over much of surface, but not as dense

on fourth as dorsally. Hairs and setulae mostly black;

those on first and second sterna golden; considerable

golden hair on first tergum ventrally. Median discals of

fourth tergum in a very irregular row near base of

tergum; second sternum with a pair of strong black

apicals. Length, 12 mm.

MALE.—Eyes separated at narrowest by about 0.035

head width, or by little more than diameter of anterior ocellus, yet frontale narrowly visible throughout.

Frontals, 14 or 15, uppermost reclinate, ascending a

little more than half way to ocellar triangle. Thorax

bright green, coppery reflections present in places but

noticeable only in certain lights. Pollen of mesonotum

and scutellum more noticeable than in female. Pteropleurals, 2. Pile of mesopleuron, except lowermost

part, and a few hairs on sternopleuron black. Middle

tibia without ventrals except apically. First three

abdominal terga as in female but purple areas more

extensive; fourth tergum showing same contrast with

preceding ones as in female, but background, which

varies from blue-green to purple, shows through more

clearly; hypopygium in contrast shining, pollen visible

only obliquely. Median discals of fourth tergum in a

much-better-ordered row and longer and stronger than

in female. Genitalia as in Figure la; inner forceps

(cerci) unusually narrow when viewed dorsally.
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

\J O

FIGURE 1.—Male genitalia, lateral view in all figures, also dorsal view of inner forceps in Isomyia

chalconotum: a, I. chalconotum, new species, from paratype; b, I. viridaurea (Wiedemann), from

Bangkok, Thailand; c, I. gomezmenori (Peris), topotypical; d, I. perisi, new species, from holotype; e, I. discalis, new species, from holotype; /, /. facialis, new species, from holotype.
NUMBER 6 7

HOLOTYPE.— $ , Malaysia (Federated Malay

States), Pahang, Fraser's Hill, 4,200 feet, 19 July 1936,

H. M. Pendlebury; British Museum (Natural History).

ALLOTYPE.— cf, Malaysia, Perak, Larut Hills, 3,700

to 4,500 feet, 11 February, H. M. Pendlebury.

PARATYPES.—Id1, 1? , same data as allotype but

collected 11 February and 15 February, respectively.

REMARKS.—This is a well-marked species, separable

from other members of this group except the preceding species by the setulose lunule and from all known

members of the group by the distinctly tricolored dorsal

surface of the thorax and abdomen, the densely pollinose fourth tergum strongly contracting with both

the thorax and the rest of the abdomen.

Isomyia marginata, new species

FEMALE.—Parafrontals, parafacials, and occipital

orbits apparently black but so densely covered with

golden pollen that color of background is virtually not

discernible; glabrous area of occiput mostly yellow,

paracranialia and broad borders of occiput brownish

black, latter becoming black on lower occiput and

postgena, these areas but moderately pollinose; gena

and oral margin yellow with dense golden pollen

which, however, becomes sparse on a subshining brown

patch on gena below eye; facialia, cheek grooves, and

face yellow, latter brownish yellow on upper half; face

with light yellow pollen which, however, dulls its entire

surface; frontale reddish yellow, dulled by concolorous

pollen; lunule yellow, glabrous. Head bristles black;

frontals, 9 or 10. Setulae of occipital orbit, parafrontals,

parafacials, and most of gena short, black; those of

parafrontals and parafacials numerous, latter finer in

texture and in three or four irregular rows. Hairs of

occiput and lower part of gena yellow, latter long.

Frontale apparently tapering gradually from ocellar

triangle to lunule, in type somewhat withdrawn below

so that its exact width is difficult to determine. Antennae orange-yellow, flagellum largely brown except at

base below; flagellum about twice length of pedicel;

arista reddish brown, yellow on a subbasal band. Measurements in micrometer units: head width, 265;

width of vertex, 60, of frons at base of antenna estimated at 90, of frontale at base of antenna estimated

at 27, of frontale at anterior ocellus estimated at 35;

distance between vibrissae, 55; vibrissa to oral margin,

30; oral margin to lower margin of lunule, 110; width

of parafacial at antennal base, 35, at cheek groove, 22;

width of gena, 55; eye height, 180; antennae separated

by 10; length of pedicel, 20, of flagellum, 40.

Mesonotum deep green-purple; a narrow longitudinal green vitta just inside each dorsocentral row,

bare of setulae, extending from anterior margin of

mesonotum to middle of postscutum; poorly defined

green areas along suture and on disc of mesonotum,

both presuturally and postsuturally, but mesonotum

predominantly purple and appearing to naked eye as

almost uniformly so; scutellum purple dorsally and

on sides; humerus purple above, green below; pleura

predominantly green but with indefinite purple areas

on sternopleuron and on anterior part of mesopleuron

and pteropleuron. Mesonotum and scutellum with light

gray tomentum, visible posteriorly, thickest on anterior

margin of mesonotum; scutellum ventrally densely

whitish pollinose on about median half, dull velvety

black laterally; pollen of pleura mostly cinereous, dense

on lower part of humerus, propleura, around anterior

spiracle, less so on extreme lower margin of mesopleuron except anterior slope, posterior part of

pteropleuron, and around posterior spiracle; visible

elsewhere only in proper light. Some long hairs basally

and short pubescence elsewhere on postalar declivity;

scutellum with rather abundant but short, fine, inconspicuous yellow hairs ventrally. Covering of anterior

spiracle bright yellow, of posterior one brown. Bristles

of thorax and setulae of mesonotum, scutellum, upper

parts of humerus, and upper anterior margin of mesopleuron black; hairs of pleura yellow; posterior mesopleural fringe light golden. Acrostichals, apparently

2, 3 (or 2, 4; postsutural acrostichals all on posterior

half of postscutum and an additional one may be

obliterated by the pin); dorsocentrals, 2, 4; humerals,

3; pteropleurals, 2; scutellars, 3 lateral, 1 discal; an

accessory propleural, no accessory stigmatals.

Femora partly black, partly strongly metallic; front

femur strongly metallic in part, ranging from purple

dorsally to purplish blue ventrally and bluish green on

basal half anteriorly; middle and hind femora in part

purplish posteriorly and blue-green on part of anterior

surface; tibiae yellowish brown; tarsi brown; bristles

and hairs black. Middle tibia with strong ventral at

two-third length. Wing hyaline; bend of Mj angular,

with a slight spur; basicosta brown broadly along margin, otherwise yellowish brown. Halteres yellow, stalk

somewhat reddish yellow; squamae and hairs yellow,

thoracic squama with tinge of brown posteriorly except along posterior rim.
8 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Abdomen purple dorsally; first tergum basally, broad

apices of second and third terga and a narrow median

vitta black; overall appearance to naked eye purple;

some green on fourth tergum, at apex, medially, and

on sides of second to fourth terga, at base of dorsal

surface; ventral aspect of terga merging into green,

considerably so on first, some bronzing on third; pollen

cinereous, invisible except when viewed from behind

on first three terga dorsally, most conspicuous there

on extreme anterior margins, more conspicuous on

fourth tergum, where pattern is tessellated; ventrally

conspicuous on first and median basal parts of second

to fourth, also on first sternum and, a little less so, on

second, visible elsewhere, as on dorsum, only in the

proper lighting. Bristles and setulae black; hairs on

ventral aspect of first tergum, anterior to lateral

bristles, and on sterna fine, pale, yellow to whitish. An

indefinite row of median discals on fourth tergum;

second sternum with one pair of strong, black apicals.

Length, 12 mm.

MALE.—Unknown.

HOLOTYPE.—• ? , Philippine Islands, Mindanao, Kolambugan; Baker; USNM 70752.

Isomyia chrysoides (Walker)

I have seen 6d <S and 7 $ $ from the H.M. Pendlebury Collection, now in the British Museum (Natural

History), from the following localities in Malaysia:

Pahang (Fraser's Hill, 4,000 ft., 25 January 1929;

Kuala Taku, about 550 ft., 5 December 1921) ; Perak

(Batang Padang, Jor Camp, 1,800-2,000 ft., 29 May

1923, 4 June 1923, and 20 August 1922; Larut Hills,

3,700-4,600 ft., 15 February 1932) ; Kuala Lumpur,

19 July 1933; and Selangor (Bukit Kutu, 3,450-3,500

ft, 18 and 19 April 1926, September 1915, 2 September

1929, and 22 September 1932).

Isomyia viridaurea (Wiedemann)

Musca viridaurea Wiedemann, 1830, p. 397.

Thelychaeta chalybea Brauer and Bergenstamm, 1891, p. 391.

This is a common species that is distributed over a

large part of the eastern Oriental Region. A more

complete synonymy is given by Peris (1952). I have

seen specimens from Taiwan (as Formosa), Viet Nam

(as Indochina), Thailand (as Siam), Malaysia (Pahang, Kedah, Kuala Lumpur), Java, Sumatra, Borneo,

Bali, and the Philippine Islands; and Peris has recorded

it, in addition, from Assam and Burma.

The name viridaurea is aptly descriptive; the fly is

usually golden-green in overall appearance, though

some specimens will take on a bluish cast. The dense

pollen, especially of the abdomen and stemopleuron,

the yellow hairs of the thoracic pleura and ventral aspect of the abdomen, the yellow basicosta, and the

yellow covering of the anterior spiracle are good recognition marks. The black setulae below the notopleural

suture are usually arranged in two longitudinal rows

and contrast strongly with the pile. The second sternum is clothed with yellow hairs, some of which are

long and bristle-like; there is one pair of apical, black

bristles. Male genitalia as in Figure \b.

Musca viridaurea was described from an unstated

number of females from Java (no other data) "In

Westermann's und meiner Sammlung." According to

Zimsen (1954), there is one specimen in the Zoological Museum in Copenhagen. There is one female in

the Natural History Museum in Vienna with the

labels "Java," "Wd. viridaurea Coll. Winthem," and

"Musca viridaurea Wied." (Italics here refer to handwritten labels, the others being printed.) The last label

is in a different and apparently more recent hand, and

this may be the female stated to be in Westermann's

collection since it has a red type label; however, no

mention of the Winthem collection is made by Wiedemann. If a lectotype is to be designated, it should be

from the Copenhagen Museum, since that is the collection from which the specimens originally came.

Thelychaeta chalybea, from the description, was

based on both sexes; the only locality given is "Borneo";

no date, collector, or other information is given. Three

specimens in the Vienna collection have red type labels.

There are two females with the labels "F. Baczas/

1886/ Borneo" and "chalybea/ det. B. B." One of

these has an additional label, written in a different and

apparently more recent hand: "Thelychaeta chalybea

Br. Bgst." I am designating this specimen the lectotype, the other a paralectotype. The only male has the

labels "Sumatra" and "chalybea/ det. B. B." Probably

this male was the one referred to this species by Brauer

and Bergenstamm, but this is simply a conjecture,

since no Sumatra locality was given.

Isomyia gomezmenori (Peris), new combination

Thelychaeta gomez-menori Peris, 1951, p. 246.

The male genitalia (Figure \c) differ considerably

from those of other members of this group that I have
NUMBER 6 7

studied. A topotypical series of 3 c? S and 4 $ $ from

India, Uttar Pradesh, Mussoorie, 5,000-6,400 feet, 15-

18 May 1962, and 1 ? , Chakrata, 6,700 feet, all collected by J. R. Donahue and in the collection of

Michigan State University.

Isomyia pichoni Seguy

I have seen only the male, referred to this species by

Peris (1956).

Isomyia pseudolucilia (Malloch), new combination

Strongyloneura pseudolucilia Malloch, 1928, p. 483.

One female—North Viet Nam (Tonkin), Montes

Mauson, 2,000-3,000 feet, April-May, H. Fruhstorfer,

Natural History Museum of Vienna—was determined

as this species by Malloch and seems to belong here.

Dr. Raymond Gagne studied the type and kindly furnished me with information necessary for placing it in

my key (number and arrangement of scutellar bristles,

color of pleural pile).

Isomyia ceballosi (Peris), new combination

Thelychaeta ceballosi Peris, 1951, p. 246.

Described by Peris from a series from Borneo and

Burma. I am referring here 1 $ , Borneo, LebangHara, 1-7 January 1925, H. Winkler; Hamburg

Museum.

Isomyia aurifacies, new species

FEMALE.—Parafrontals, ocellar triangle, and occiput, black; frontale brownish black; parafacials and

cheek groove reddish brown, a black area occupying a considerable part of lower parafacial; gena reddish yellow, lunule and face more clearly yellow. Occipital orbits, parafrontals, parafacials, facialia, genae,

lunule, and upper facial carina with golden yellow

pollen, almost uniform but somewhat deeper in color

on parafacials; pollen of face and occiput other than

orbits yellow to pale yellow; lunule entirely pollinose. Bristles and hairs of head black except

those of occiput, which are pale yellow above to

mixed yellow and golden below. Setulae of parafacials

in about three irregular rows. Frons gradually widening

from vertex to antennal base; frontale widest at twofifths distance from vertex to antennal bases. Antennae

orange, flagellum somewhat brownish externally above;

setulae black; arista orange, becoming brown at apex,

rays mostly brown. Measurements in micrometer units:

head width, 310; width of vertex, 90, of frons at base

of antennae, 120, of frontale at maximum, 50, of parafrontal at same place, 26, of frontale and parafrontal

at antennal base, 37 and 42; distance between vibrissae, 67; vibrissa to oral margin, 30; oral margin to

lower margin of lunule, 105; parafacial at cheek groove,

29; width of gena, 48; eye height, 195; antennae separated by 9; length of pedicel, 18, of flagellum, 40.

Thorax green, scutellum more blue-green, with blue

to purplish reflections varying with light incidence;

mesonotum lightly but evenly cinereous pollinose, pollen visible best as viewed from behind, most noticeable

along anterior margin of mesonotum; pollen of pleura

scant except on lower part of sternopleuron. Posterior

fringe of mesopleuron and softer hairs of pteropleuron

brownish black, otherwise bristles, setulae, and hairs of

thorax black. Distinct narrow vittae between acrostichal and dorsocentral rows from anterior margin of

mesonotum to about two-fifths way between suture and

scutellum, also postsutural vittae between dorsocentral

and intra-alar rows and a triangular area behind posthumerals, devoid of setulae. Mesopleural pile only

moderately long and dense. Posterior declivity with

some black setulae. Coverings of anterior and posterior

spiracles blackish brown. Acrostichals, 1, 2; dorsocentrals, 2,4; humerals, 4; pteropleural, 1 plus 1 accessory;

scutellars, 4 lateral, 2 discal; 1 accessory propleural and

1 accessory stigmatal.

Coxae and femora largely black; front coxa anteriorly, middle coxa in large part, front femur posteriorly

on basal two-thirds, middle femur posteroventrally on

basal half, and a ventral area at base of hind femur

metallic green. Tibiae and basitarsi reddish yellow,

front basitarsus above and other tarsomeres becoming

brownish black. Middle tibia with strong ventral at

three-fifths length. Wing yellowish, veins golden, more

brownish toward base on lower surface; basicosta black;

Mx broadly rounded at bend but bent at almost a right

angle, strongly concave beyond bend; squamae uniformly brown.

Abdomen blue-green; first tergum largely with purplish reflections, especially ventrally; second to fourth

terga with black posterior margins, broader ventrally

than dorsally, and with a rather indistinct median

vitta. Pale pollen cinereous, scant dorsally, visible at
10 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

an angle at sides of second and third terga, much more

prominent ventrally, a dense spot on first under squamae, otherwise densest toward midventral margins of

first to fourth terga; second sternum with dense cinereous pollen. Hairs, bristles, and setulae mostly black;

second sternum basally with brownish yellow hairs,

otherwise with black setulae and about five pairs of

black bristles; an irregular discal row at about onethird of fourth tergum dorsally. Length, 12 mm.

MALE.—Unknown.

HOLOTYPE.— $ , India, Assam, 10 miles north of

Tinsukia, 11 April 1944, D. E. Hardy; USNM 70753.

Isomyia perisi, new species

MALE.—Head largely black. Gena, cheek groove,

lower outer part of faciale, and lunule reddish brown,

face and rest of faciale reddish yellow; middle of gena

rather indefinitely brownish; pollen of parafrontals,

parafacials, and occiput cinereous, that of faciale, parafacial, cheek groove, and gena concolorous with background; lunule with clearly evident yellowish pollen,

visible from any angle, on more than its posterior half.

Occiput with long yellow hairs medially, most conspicuous below glabrous area of upper occiput but also

visible back of proboscis, other hairs, setulae, and

bristles of head black. Frontals about 12, upper ones

becoming size of larger setulae; setulae abundant on

parafrontals and parafacials; lunule without setulae.

Width of frons at narrowest little more than diameter

of anterior ocellus, frontale completely obliterated at

narrowest. Antennal scape and pedicel reddish brown,

flagellum reddish yellow, brown above except at base.

Arista reddish brown to brown, subbasal third yellow;

rays brown. Measurements in micrometer units: head

width, 310; width of frons at minimum, 8, at vertex,

30; distance between vibrissae, 62; vibrissa to oral

margin, 45; oral margin to lower margin of lunule,

130; parafacial at antennal base, 20, at cheek groove,

32; width of gena, 70; eye height, 210; antennae separated by 10; length of pedicel, 20, of flagellum, 40.

Thorax blue-green with purplish reflections, especially laterally; pollen brownish gray, clearly evident

from posterior view over most of mesonotum, scant on

scutellum, lacking or scant on most pleural areas, most

evident there on part of the sternopleuron; darker

pollen resulting in a pair of narrow presutural vittae

between acrostichal and dorsocentral rows; hairs,

setulae, and bristles mostly black, some around anterior spiracle and posterior mesopleural fringe dark

brown. Coverings of anterior spiracle brownish black,

that of posterior black. Acrostichals, 2, 3, anterior presutural and postsutural much weaker than others;

dorsocentrals, 2, 4; scutellars, 4 lateral, 1 or 2 discal;

pteropleural, 1, surrounded by setulae as long as the

bristle, some of them almost bristle-like; 1 accessory

propleural, no accessory stigmatals.

Coxae and femora black, anterior pair with strong

blue-green reflections, coxa anteriorly and femur

posteriorly virtually blue-green, others with traces of

such; tibiae and basitarsi reddish brown to brown,

anterior pair palest; trochanters and tarsi beyond

basitarsi black. Middle tibia without median ventral.

Wing pale brown, almost hyaline; bend of Mj pronounced, forming approximately a 100° angle, but

rounded at bend. Basicosta black; halteres reddish yellow; squamae wholly dark brown.

Abdomen bluish green. Pollen scant except on second sternum and inner half of ventral areas of second

and third terga, where it is dense and cinereous. First

abdominal tergum and posterior margins of second and

third, also a broad median vitta dorsally on second and

a narrow one on third, black, with purple reflections;

ventral sides of first three terga with strong violet

reflections, fourth less so; black areas of posterior

margins of second and third much more broadly developed ventrally than dorsally. Fourth tergum well set

(except on basal third) with discals, not arranged in

rows. Second sternum with about eight pairs of bristles,

but with no stronger apical pair, and with long yellowish hairs; bristles, setulae, and hairs of abdomen black,

unless otherwise noted. Genitalia as in Figure Id.

Length, 13 mm.

FEMALE.—Frontale brownish black. Pollen of parafrontals and parafacials variable in color, usually

brownish to yellowish brown, darker on parafrontals

than on parafacials, sometimes mostly cinereous as in

the male. Pale hairs of upper occiput not as long or as

deep yellow as in male. Measurements in micrometer

units: head width, 310; width of vertex, 90, of frons

at base of antennae, 125, of frontale and parafrontal

at anterior ocellus, 35 and 25, of same at antennal base

42 and 42, of parafacial at cheek groove, 30. Frons

gradually broadening from anterior ocellus to base of

antennae. Middle tibia with a strong ventral a little

beyond middle. Discals of fourth tergum arranged

mostly in a row just anterior to midline of tergum.

Length, 13 mm (allotype: range, 12-13 mm).
NUMBER 6 7 11

HOLOTYPE.— <S, South India, Madras, Anamalai

Hills, Kadamparai, 3,500 feet, May 1963; P. S.

Nathan; USNM 70754.

ALLOTYPE.— 9 same data as holotype.

PARATYPES.—5 $ $ , same data; 5 ? <j>, same data

but Anamalai Hills, Cinchona, May 1964; 1 9 , same

data but Cinchona, September 1964.

REMARKS.—This new species may be the species

discussed by Peris (1952, p. 165) as "Th. sp.?" and

referred by him to the delectans complex, which he

thought consisted of more than one species but which

he was unable to resolve. Like I. ceballosi (Peris), /.

pseudolucilia (Malloch), and / aurifacies, new species,

this new species has four marginal scutellars which are,

however, not equidistant from one another, the distance between the basal one and the next in line being

the greatest. The species of this complex are all large

and robust, and the second sternum, which is broader

than usual for the viridaurea group, is set with a number of black bristles rather than with only the usual

strong apical pair.

I take pleasure in dedicating this species to Dr.

S. V. Peris, who has done so much toward elucidating

our knowledge of the Rhiniinae.

Isomyia iris, new species

MALE.—Occiput, ocellar triangle, frons, parafacials,

and a large area on anterior part of gena black; face

yellow, facials, cheek groove, rest of gena, and lunule

orange to orange-yellow; pollen cinereous, densest on

parafrontals, parafacials, and occipital orbits, becoming golden on cheek groove, much of gena, and face,

only moderately dense on gena, the entire ground color

showing through; f rontalia on upper half golden pollinose; lunule apparently shining, yet when viewed from

behind upper part distinctly yellow-pollinose. Bristles,

hair and setulae black to blackish brown, those of occiput yellowish, becoming yellowish brown to brown

below; setulate of parafacials moderately long, in

two fairly well defined rows. Frontals, 8 or 9,

row continued with a few setulae above to a point

about half way from antennal base to ocelli;

lunule without setulae. Width of frons at narrowest area less than diameter of anterior ocellus,

frontale completely obliterated at narrowest. Antenna

orange, flagellum largely yellow, upper surface brown;

arista orange at base, then yellow, becoming brown

on apical half; rays brown. Measurements in micrometer units: head width 275; width of frons at minimum, 4, at vertex, 30; distance between vibrissae, 55;

vibrissae to oral margin 35; oral margin to lower margin

of lunule, 105; parafacial at antennal base, 15, at

cheek groove, 17; width of gena 45; eye height 195;

antennae separated by 8; length of pedicel 20, of flagellum, 43.

Mesonotum black centrally, with dull bronze reflections when seen through pollen; broad sides and prescutellar area deep metallic blue-green, more of a deep

green in supra-alar area and in front of scutellum, deep

blue to purple elsewhere; central part of prescutellar

green area forming a triangle from postalar callus almost to suture, where it is continued by a vitta between

acrostichal rows to anterior presutural acrostichal;

dorsocentrals set on a narrow purple vitta with violet

reflections; black areas strongly cinereous pollinose, as

viewed from behind, leaving a narrow black vitta between each acrostichal and dorsocentral row and a

large triangular area between dorsocentral row and

posthumerals without pale pollen; areas of contact between black and metallic areas merging in proper light

from green to deep blue to purple to violet (this suggests the name iris, "rainbow"). Scutellum and pleura

deep blue with purplish reflections; under side of scutellum cinereous pollinose on median third; pollen elsewhere cinereous to brown, generally not conspicuous

except on sternopleuron. All thoracic bristles, setulae,

and most hairs black to brownish black; hairs of

mesopleuron especially long and dense, especially on

upper part, where they tend to obscure the usual black

setulae; these hairs appear brownish under certain

lights; posterior mesopleural fringe brown; some black

erect hairs on posterior declivity and under sides of

scutellum. Covering of anterior and posterior spiracles

deep brown. Acrostichals, 2, 3, anterior postsutural

weak; dorsocentrals, 2, 4; scutellars, 3 lateral 1 discal;

humerals 3; pteropleural, 1; no accessory propleural or

stigmatal.

Legs black, with black setulae and bristles; front

femur with strong purplish reflections, almost appearing purple, posteriorly and on basal part anteriorly,

other femora with limited purplish reflections posteriorly; tibiae more brownish black, front tibia in

places yellowish brown. Middle tibia without a ventral.

Wing almost hyaline, very slightly brownish; bend of

Mx abrupt but not angular. Basicosta brown to dark

brown, darker anteriorly. Halteres yellow. Squamae

wholly and uniformly brown except rim yellowish.
12 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Abdomen deep blue with strong purple reflections;

basal part of first tergum dorsally and broad apices

of second and third, also a median vitta on first three,

black; ventral aspects of second and third becoming

bronze medially. Pollen of dorsal surface inconspicuous,

visible as thin cinereous pollen when viewed posteriorly;

that of ventral surface much more apparent, dense on

first tergum and toward inner margins of second and

third, where it forms tessellated patterns. First sternum

apparently black, second purple, densely cinereous

pollinose. Bristles and setulae mostly black; hairs of

first and second sterna white, but second also, in addition to apical bristles, with numerous strong black

setulae that appear as fine bristles. Genitalia not

dissected. Length, 11 mm.

FEMALE.—Unknown.

HOLOTYPE.— cf, Philippine Islands, Mindanao,

Zamboanga; Baker; USNM 70767.

Isomyia discalis, new species

MALE.—Occiput, ocellar triangle, frons, parafacials,

and postgena black; lunule orange; face yellow; facials,

cheek groove, and gena yellow to yellowish brown, gena

with a large indefinite blackish spot anteriorly. Ocellar

triangle and occiput with thick whitish pollen; parafacials and genae with thick yellowish pollen, obscuring

ground color except on part of the black spot of the

gena, which is subshining, and on a small spot on lower

part of parafacial; face golden pollinose; frontale dark

brown; lunule apparently shining, yet when viewed

from behind upper part distinctly yellow pollinose.

Hairs of lower occiput, postgena, and posterior part

of gena yellow, bristles and setulae of head all black;

hairs of parafacials moderately long and abundant, in

two or three very irregular rows; eight or nine frentals,

row continued with some long setulae to about half

way from base of antennae to ocelli; lunule without

setulae. Width of frons at narrowest less than diameter

of anterior ocellus, frontale there completely obliterated. Antenna orange, flagellum more yellowish

below, brownish above; arista reddish brown at base,

then yellow, becoming brown on apical half. Measurements in micrometer units: head width, 240; width of

frons at minimum, 3, at vertex, 25; distance between

vibrissae, 45; vibrissa to oral margin, 30; oral margin

to lower margin of lunule, 100; eye height, 175;

antennae separated by 7; length of pedicel, 30, of

flagellum, 62.

Mesonotum black centrally, opaque to subshining,

broadly metallic green laterally and posteriorly, the

green presutural area extending forward to third postsutural dorsocentral; a narrow median green vitta

between acrostichal rows, interrupting black area but

not reaching anterior margin; some deep purplish reflections where black and green areas meet and along

suture; scutellum and pleura green, in places with

bluish reflections, a median area of hypopleuron purple.

Pollen cinereous, thin on mesonotum, scutellum, and

most pleural areas, where it is visible only posteriorly,

more evident on sternopleuron. All bristles and most

setulae black; abundant black hairs on upper part of

mesopleuron in addition to usual black setulae; lowermost hairs of humerus yellow, others black; most hairs

of mesopleuron black, a few below yellow, posterior

mesopleural fringe golden; sternopleuron with hairs on

anterior half black, otherwise yellow; other pleural

hairs mostly yellow. Posterior declivity with some short

brownish hairs. Underside of scutellum densely whitish

pollinose medially, opaque black on sides; a few scattered yellowish hairs scattered over ventral surface.

Covering of anterior spiracle brown, of posterior dark

brown. Acrostichals, apparently 2, 5, posterior presutural and posterior two postsuturals much the

strongest; four postsuturals visible in type, apparently

a fifth (the third of the series) obliterated by the pin;

dorsocentrals, 2, 4; scutellars, 3 lateral, 1 discal;

humerals, 3; pteropleural, 1; an accessory propleural

but no accessory stigmatals.

Legs black, with black hairs and bristles; basitarsi

with somewhat of a reddish tinge; femora with a

greenish reflection on part of anterior surface, front

femur also with similar reflection posteriorly. Middle

tibia with small but distinct ventral at two-thirds

length. Wing uniformly light brown; bend of Mi

broadly rounded, not angulate, part beyond bend

lightly sinuate. Basicosta yellow, somewhat brownish

inwardly. Halteres orange. Squamae brown, rim and

fringe yellow; outer half of thoracic squama white.

Abdomen basically green, fourth tergum with

golden-green reflection; actually much of dorsal surface

is black, including all of first tergum, second tergum

except laterobasal angles, and posterior margins and

median vitta on third and fourth, both broader on

third than on fourth; terga ventrally with black

posterior markings on second to fourth segments, about

half length of segment on second and third, narrow
NUMBER 6 7 13

on fourth; first sternum brownish, second green; green

areas of abdomen mostly with dense whitish pollen

which, in proper light, completely obscures background; pattern on fourth tergum clearly tessellated.

Bristles, setulae, and hairs mostly black; first and second

sterna, and most of ventral aspect of first tergum, with

whitish to yellowish hairs, those of second sternum

long; second sternum with one pair of strong apical

bristles. Genitalia as in Figure \e. Length, 10.5 mm.

FEMALE.—Unknown.

HOLOTYPE.— <$, Philippine Islands, Biliran; Baker;

USNM 70768.

REMARKS.—This is a well-marked species, separable

from all others in the group known to me (except the

preceding species) by the thoracic coloration. It is

unfortunate that females of both these species are unavailable. Malloch had obviously recognized this species

as distinct and undescribed and had placed the label

"discalis" on the type specimen but had not described it.

Isomyia delectans (Walker)

Walker's type was from Celebes. The only other

specimens that I can refer to this species with reasonable certainty also come from that island: 2 $ $ ,

South Celebes, Samanga, November 1895, H. Fruhstorfer, National History Museum of Vienna; 2 c? c?

1 $ , North Celebes, Tondano, 25 May 1940, R. G.

Wind, Washington State University.

Isomyia borneensis (Peris), new combination

Thelychaeta borneensis Peris, 1951, p. 247.

This species was described (Peris, 1951) and redescribed in more detail (Peris, 1952) from a single

female, erroneously stated to be a male in the type

citation in the second work. I have seen six specimens

which conform with Peris' descriptions for the most

part; the femora, described as metallic green or violet,

are variable as to the extent of that color; and the

abdomen is distinctly pollinose dorsally, at least in

large part, if viewed at an angle. New records are:

2dV , 1? , Borneo, Kupat, 7-15, September 1927,

British Museum (Natural History); 2 $ $ , Borneo,

Bettotan, 18 August 1927, British Museum (Natural

History); 1 cT, Sumatra, Fort de Koch, 920 meters,

1925, E. Jacobson, Amsterdam Museum.

Isomyia spp.

A considerable residue of material, forming, along

with /. delectans, I. pictifacies, and /. electa, a part

of the delectans complex, must await further study

after accumulation of more material.

Isomyia facialis, new species

MALE.—Head black; lower part of face from

slightly above level of vibrissa, also oral margin,

yellow; cheek grooves and lunule reddish brown.

Pollen yellowish brown on inner margin of parafacial

and on cheek groove, becoming yellow on parafacials,

a paler yellow on gena and occiput, a darker brown

on parafrontals; yellowish to whitish on face. Lunule

clearly pollinose, with a glabrous area above each

antennal base. Hairs of occiput whitish above, yellowish below, blackish toward orbits; otherwise all bristles,

setulae, and hairs of head black. About eight frontals,

row continued by setulae above; setulae of parafrontals sparse, of parafacials sparse and fine but rather

long, mostly in one row. Lunule without setulae. Frons

at narrowest much less than diameter of anterior

ocellus, frontale obliterated there and parafrontals

almost so. Antennal scape and pedicel orange; flagellum (based on paratype, missing in holotype) brown

above and apically, yellow below. Measurements in

micrometer units: head width, 230; width of frons at

minimum, 3, at vertex, 25; distance between vibrissae,

44; vibrissa to oral margin, 36; oral margin to lower

margin of lunule, 103; parafacial at antennal base, 13,

at cheek groove, 17; width of gena, 35; eye height,

165; antennae separated by 5; length of pedicel (paratype) 16, of flagellum, 32.

Thorax green, more blue-green laterally, with

bronze reflections on mesonotum except laterally; all

hairs, setulae, and bristles dark brownish black to

black. Mesonotum and scutellum, except laterally,

with grayish pollen visible from posterior view; traces

of bronze to black presutural vittae between dorsocentral and acrostichal rows. Pollen of pleura scant.

Posterior declivity with hairs much more distinct and

abundant than usual in this group; scutellum with

black hairs ventrally. Covering of anterior and posterior

spiracles dark brown. Acrostichals, 2, 4, postsuturals

set close together on posterior half of postscutum;

dorosocentrals, 2, 4; scutellars, 3 lateral, 1 discal;

humerals, 3 or 4; pteropleural, 1; an accessory propleural but no accessory stigmatals.
14 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Coxae and femora black, coxae anteriorly and

femora anteroventrally partly metallic green, more extensively with metallic green reflections; tibiae and

basitarsi reddish yellow, tarsi presumably becoming

darker apically (missing beyond second tarsomere in

holotype and all male paratypes); middle tibia without ventrals. Wing hyaline, bend of Mi broadly

rounded, strongly sinuate beyond bend. Basicosta

black. Halteres yellow, knob a little darker. Squamae

brown; outer basal part of thoracic squama white.

Abdomen green; first tergum largely, second and

third terga on apical margins, broader ventrally than

dorsally, and on an indefinite median line, black;

second and third terga more blue-green ventrally;

second to fourth terga dorsally and fourth tergum

ventrally with bronze reflections, black areas of second

and third terga ventrally with purplish reflections.

Second sternum green. Pollen cinereous on dorsal surface, visible from posterior view, clearly evident ventrally on terga except toward sides and on second

sternum. Second sternum with yellow hairs and with

about three pairs of black bristles, also with a few black

setulae; otherwise all abdominal bristles, setulae, and

hairs black. An irregular row of dorsals on fourth

tergum. Genitalia as in Figure If. Length, 10 mm

(paratypes, 9.0 to 10.0 mm).

FEMALE.—Lower part of gena becoming yellowish

toward oral margin. Pollen of head more cinereous,

some yellowish on parafacials, more yellowish on gena.

Frontale reddish brown, pollen concolorous; widest at

about half distance from anterior ocellus to lunule,

narrowing gradually above and below. Measurements

in micrometer units: head width, 215; width of frons

at vertex, 50, at base of antennae, 80, of frontale at

broadest and parafrontal at same point, 30 and 17,

same at anterior ocellus, 25 and 15, same at base of

antenna, 25 and 27. Middle tibia with a strong ventral

bristle two-thirds distance from base. Halteres white.

Discals of fourth tergum mostly in a transverse row,

as in the male. Length, 10 mm.

HOLOTYPE.—o*, Thailand (Siam), Bangkok, H. M.

Smith; USNM 70769.

ALLOTYPE.— $ , 4 <$ <$, same data as holotype.

PARATYPES.—2 $ $ , 4 c? cf, same data as holotype.

REMARKS.—Unfortunately, no specimen in the

series is complete; the antennal flagellum and arista are

missing in all but one male and one female paratype

and the legs are broken in all specimens. The holotype,

in spite of its broken antenna, was chosen as the most

complete and representative specimen in the series.

Literature Cited

Brauer, F., and J. E. von Bergenstamm

1891. Die Zweifliigler des Kajserlichen Museums zu

Wien. V. Vorarbciten zu einer Monographic dcr

Muscaria Schizometopa (exclusive Anthomyidae).

Part 2. Denkschriften der Kais. Akademie der

Wissenschaften, Wien, 58:305-446.

Malloch, J. R.

1928. Exotic Muscaridae (Diptera). XXI. Annals and

Magazine of Natural History, (10)1:465-494.

Peris, S. V.

1951. Descripciones Preliminares de Nuevos Rhiniini

(Dipt. Calliphoridae). Eos, 27:237-247.

1952. La Subfamilia Rhiniinae (Dipt. Calliphoridae).

Anales de la Estacion Experimental de Aula Dei:

3:1-224.

1956. Nuevas Notas sobre Rhiniini con Descripcion de

Formas Nuevas. Eos, 32:231-254.

Seguy, E.

1949. Les Calliphorides Thelychaetiformes du Museum

de Paris. Revista Brasiliera de Entomologia, 9:115-

142.

Senior-White, R.; D. Aubertin; and J. Smart

1940. The Fauna of British India. Diptera. Volume 6,

Family Calliphoridae. xiii -f- 288 pages. London:

Taylor and Francis.

Wiedemann, C. R. W.

1830. Aussereuropaische Zweifliigelige Insekten. II. xii -f

684 pages.

Zimsen, E.

1954. The Insect Types of C. R. W. Wiedemann in the

Zoological Museum in Copenhagen. Spolia Zoologici Musei Hauiensis, 14:5-43.

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