NOAA Technical Memorandum NMFS-AFSC-16

U.S. DEPARTMENT OF COMMERCE

National Oceanic and Atmospheric Administration

National Marine Fisheries Service

Alaska Fisheries Science Center

June 1993

by

Clifford H. Fiscus

Catalogue of Cephalopods at the

National Marine Mammal Laboratory
NOAA Technical Memorandum NMFS

The National Marine Fisheries Service's Alaska Fisheries Science Center

uses the NOAA Technical Memorandum series to issue informal scientific and

technical publications when complete formal review and editorial processing

are not appropriate or feasible. Documents within this series reflect sound

professional work and may be referenced in the formal scientific and technical

literature.

The NMFS-AFSC Technical Memorandum series of the Alaska Fisheries

Science Center continues the NMFS-F/NWC series established in 1970 by the

Northwest Fisheries Center. The new NMFS-NWFSC series will be used by

the Northwest Fisheries Science Center.

This document should be cited as follows:

Fiscus, C. H. 1993. Catalogue of cephalopods at the National Marine

Mammal Laboratory. U.S. Dep. Commer., NOAA Tech. Memo. NMFSAFSC-16, 183 p.

Reference in this document to trade names does not imply endorsement by

the National Marine Fisheries Service, NOAA.
June 1993

NOAA Technical Memorandum NMFS-AFSC-16

by

Clifford H. Fiscus

 Catalogue of Cephalopods at the

National Marine Mammal Laboratory

Alaska Fisheries Science Center

7600 Sand Point Way N.E.

Seattle, WA 98115-0070

U.S. DEPARTMENT OF COMMERCE

Ronald H. Brown, Secretary

National Oceanic and Atmospheric Administration

Diana Josephson, Deputy Under Secretary
This document is available to the public through:

National Technical Information Service

U.S. Department of Commerce

5285 Port Royal Road

Springfield, VA 22161
Notice to Users of this Document 

This document is being made available in .PDF format for the convenience of users; however, the 

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iii

CONTENTS

Page No.

... 1 . . . . . . 43 .

..... 1 . .... 1 . •••••• 2 ••.•••••

PREFACE ..

CATALOGUE ..

Order Sepiida .

Family Sepiidae .

Sepia esculenta

Sepia kobiensis .....

Sepia andreana.

Order Sepiolida ...

Family Sepiolidae .

Rossia pacifica ..

Order Teuthida .

Suborder Myopsida

Family Loliginidae .

Loligo iaponica .

Loligo opalescens .

Loliqo pealei? .

Doryteuthis bleekeri .

Lolliguncula brevis .

Number Specimens

Each Category

. . . 1 ..

· 2 .

. 1. · 1 .

. . 6 ...

vii

1

1

1

1

1

1

2

2

2

4

4

4

4

15

15

15

Suborder Oegopsida

Family Enoploteuthidae .

Abraliopsis affinis 1 ..

Abraliopsis sp 14 ..

Watasenia scintillans 3 .

Pteryqioteuthis gemmata (giardi) 1 .

17

17

17

19

20

Family Octopoteuthidae .

Octopoteuthis deletron .

Taningia danae .

7 •••

. . 3 ..

21

21

22

Family Onychoteuthidae .

Onychoteuthis banksii 3 .

Onychoteuthis borealiiaponica 58 ....

Moroteuthis robusta 22 ..

23

23

23

38

Family Gonatidae .

Gonatus berryi .

Gonatus madokai ..

Gonatus middendorffi ..

Gonatus onyx .

Gonatus pyros .....

• • • 4 •.

. . . .11 .

· 8

.19

• . . . 8

44

44

45

47

50

54
iv

Number Specimens

Each Category

... 9

. 13 .

.3 . .63 ..

.. 2 ..... · . 1

..... 10 . •••••• 24 •••

Family Gonatidae (Cont.) ...

Gonatus sp. (fabricii type)

Gonatus sp .

Eogonatus tinro .

Gonatopsis borealis .

Gonatopsis makko .

Gonatopsis Type A Kubodera.

Berryteuthis anonychus.

Berryteuthis magister ..

Family Architeuthidae .

Architeuthis japonica .

Family Histioteuthidae . Histioteuthis dofleini .

Histioteuthis heteropsis ..

· . 12 .

8 ..

· . 2 .

55

56

57

61

62

78

78

78

82

92

92

95

95

96

Family Omrnastrephidae 98

Illex illecebrosus 1 98

Illex argentinus 1 98

Todarodes pacificus 4 99

Nototodarus gouldi 2 100

Nototodarus sloani 2 100

Ommastrephes bartrami 17 102

Dosidicus gigas 20 107

Sthenoteuthis (SYmplectoteuthis) oualaniensis 2 112

Eucleoteuthis (SYmplectoteuthis) luminosa .. 1 112

Family Thysanoteuthidae 113

Thysanoteuthis rombus 1 113

Family Chiroteuthidae .

Chiroteuthis calyx .

Chiroteuthis sp .

Family Mastigoteuthidae .

Mastigoteuthis sp ..

. 4 •

8

. 1 .

· 114

· 114

. . . 114

117

· 117

Family Cranchiidae 118

Cranchia scabra 3 118

Taonius pavo 4 119

Meqalocranchia sp 6 120

Galiteuthis sp 16 122

Galiteuthis phyllura 2 123

Mesonychoteuthis sp 1 125
v

Number Specimens

Each Category

. . . . . •22 •. ........ 1. ....... 3 ..

. . . . . . . 13 .

...... 3.

Order Vampyromorpha .

Family Vampyroteuthidae ..

Vam~yroteuthis infernalis .

Order Octopoda . Suborder Cirrata

Family Opisthoteuthidae ..

Opisthoteuthis californiana

Suborder Incirrata

Family Bolitaenidae .

Japetella heathi ..

Family Octopodidae .....

Octopus dofleini .

Octopus bimaculatus .

Octopus rubescens ..

Octopus sp .....

Benthoctopus sp. A ..

7 .

3 ..

6 .

128

· 128

· .. 128

. . 130

· 130

· .. 130

· 131

· .. 131

. . . . 133

. . . . . . 133

· 141

141

· 142

· 145

Family Tremoctopodidae .

Tremoctopus sp 3 ..

· 146

146

Family Ocythoidae ..

Ocythoe tuberculata . 7 ..

· 148

· 148

Family Argonautidae .

Argonauta pacifica 1 .

. . 150

. . . . 150

Family Allopsidae

Alloposus moll is ..

ACKNOWLEDGMENTS .

CITATIONS ....

2 .. . . 151

152

· 153

APPENDIX A - Sequential listing of catalogue numbers. 161

APPENDIX B - Marine mammal predators noted in catalogue

and cephalopod prey 169

APPENDIX C - Reexamination of cephalopod beaks from

previously published papers 175

APPENDIX D - Examination of cephalopod beaks from the

stomachs of marine mammals (To the best of

our knowledge, these data not published

elsewhere) 179

vii

PREFACE

The collection of fishes and cephalopods at the National

Marine Mammal Laboratory (NMML) of the Alaska Fisheries Science

Center (AFSC) was started in the late 1940s by Victor B.

Scheffer, Ford Wilke, and Karl W. Kenyon. The cephalopod

collection is used primarily as an aid in the identification of

cephalopod remains found in the stomachs, scats, and spewings of

marine mammals. As such, the parts of cephalopods which persist

after the soft parts are gone are prominent in the collection

(i.e., beaks, gladii, and statoliths). See Fiscus (1991) for

additional information on collection, preparation, and storage of

reference materials. The size of the cephalopod collection

increased dramatically in the late 1950s and 1960s with the

advent of large-scale NMML studies of the pelagic life of

northern fur seals, Callorhinus ursinus. Field parties continue

to collect and bring material back for the laboratory reference

collection.

Specimens are classified to the lowest possible taxon. The

collection presently contains cephalopods representing 22

families, 41 genera, and 58 species. Most of the material is

from the eastern North Pacific Ocean and the Bering Sea; however,

10 species from the western North Pacific Ocean, 9 from the

eastern tropical Pacific Ocean, 3 from the Atlantic Ocean, 1 from

the sub-Antarctic, and 2 from the southwest Pacific Ocean are

also represented in the collection.
viii

Specimens in the collection have been assigned NMML numbers

as listed in this catalogue. They are assigned catalogue numbers

when acquired; consequently, consecutive numbering does not hold

for each family. A listing by numerical order appears in

Appendix A.

The following information is listed for each specimen when

possible: date and location of collection, collector, name of

marine mammal from which sample was taken (and specimen number of

mammal) or method of collection, original specimen number,

description of specimen, parts saved, and the identifier of

specimen.

Abbreviations used in the catalogue include the following:

DML - dorsal mantle length; VML - ventral mantle length; MW -

mantle width; TL - total length; HCL - hectocotylus length; HAL -

hectocotylized arm length; LRL - lower rostal length; HL - hood

length; and CL - crest length.

Identification by scientists other than the author is

indicated in each specimen description; where no credit is given,

the identification was made by the author. Specimens that have

been listed in published reports are footnoted and the

publication is available from the NMML. Classification of the

collection follows that of Clarke and Truman (1988). Sweeney

et al. (1992) should be consulted for recent nomenclature changes

and geographical distribution.

As a working collection, it is not available for public

viewing. Specialists desiring permission to work with the
ix

collection must obtain approval from the NMML's Division

Director.

The catalogue was first published in 1984 (Tech. Memo. NMFS

F/NWC-65). This edition of the catalogue incorporates recent

nomenclatural changes in names, identifies some specimens

previously identified only to the family or genus level, to

species and includes additional measurements of specimens already

in the collection. An additional 110 new specimens have been

added to the collection since 1984. Included in the appendices

are revisions of identifications of cephalopods from earlier

published papers and recent identifications of cephalopopd

species from marine mammal stomachs not published elsewhere.

CATALOGUE

Order Sepiida

Family Sepiidae

NMML 30.

NMML 31.

NMML 32.

NMML 33.

NMML 34.

Sepiidae

Collected 18 November 1966, St. Paul Island, Alaska,

by R. J. Hajny. Washed ashore. Parts saved: 1

large cuttlebone.

Sepia esculenta

Collected 9 March 1950, 20 miles offshore Fukushima

Prefecture, Honshu, Japan, by F. Wilke. Parts

saved: 2 cuttlebones.

Sepia kobiensis?

Collected 24 April 1965, Kesennuma, Miyagi

Prefecture, Honshu, Japan, by H. Kajimura. Parts

saved: 1 entire specimen.

Sepia andreana

Collected 24 April 1965, Kesennuma, Miyagi

Prefecture, Honshu, Japan, by H. Kajimura. Parts

saved: 1 entire specimen.

Sepia esculenta?

Collected 24 April 1965, Kesennuma, Miyagi

Prefecture, Honshu, Japan, by H. Kajimura. Parts

saved: 1 entire specimen.
2

Order Sepiolida

Family Sepiolidae

NMML 26.

NMML 27.

NMML 28.

NMML 29.

NMML 238.

Rossia pacifica

Collected (no date), puget Sound, Washington.

Parts saved: 2 entire specimens.

Rossia pacifica

Collected (no date), Puget Sound, Washington.

Parts saved: 1 entire specimen.

Rossia pacifica

Collected 26 June 1973, lat. 55°00'N,

long. 167°31'W, Bering Sea, by C. H. Fiscus. Parts

saved: 1 entire specimen.

Rossia pacifica

Collected 24 April 1965, Kesennuma, Miyagi

Prefecture, Honshu, Japan, by H. Kajimura. Parts

saved: 5 entire specimens.

Rossia pacifica

Collected 3 November 1981, lat. 55°40'N,

long. 168°14'W. RV Miller Freeman, cruise 81-6, haul

38, by trawl. Depth 73 fathoms. Surface water

temperature 6.5°C, trawl temperature 5.6°C. Frozen

fresh. Male, DML 45 mm, weight 29 g (in

laboratory). Parts saved: 1 entire specimen. Beaks

and radula extracted and in same jar as specimen.
3

Family Sepiolidae (Cont.).

NMML 471. Rossia pacifica

Collected 29 March 1981, lat. 59°46'N,

long. 170018'E. South of Cape Olyutorskiy, Bering

Sea. By G. A. Antonelis. From the stomach of

Erignathus barbatus. Specimen ZV-8-81. Parts

saved: 1 pair beaks. Det. C. H. Fiscus.
4

Order Teuthida

Suborder Myopsida

Family Loliginidae

NMML 36.

NMML 37.

NMML 38.

NMML 39.

NMML 40.

NMML 41.

Loliqo japonica

Collected 24 April 1965, Kesennuma, Miyagi

Prefecture, Honshu, Japan, by H. Kajimura. Parts

saved: 3 entire specimens.

Loligo opalescens

Collected November 1959, Puget Sound, Washington, by

T. O'Brien. By trawl. Parts saved: 12+ entire

specimens - see 40 and 41 below (various sizes, not

necessarily all from same tow).

Loliqo opalescens

Collected 19 May 1964, lat. 40026'N, long. 124°42'W.

From stomach of Callorhinus ursinus, specimen US 64-

253. Parts saved: 12+ entire specimens.

Loligo opalescens

Collected 13 April 1958, Port Orford, Oregon, by C.

H. Fiscus. Surface test gill net. Parts saved: 2

entire specimens.

Loligo opalescens

Collected November 1959, puget Sound, Washington, by

T. O'Brien. Parts saved: 3 entire females; see NMML

37.

Loligo opalescens

Collected November 1959, puget Sound, Washington, by

T. O'Brien. Parts saved: 2 entire males; see NMML 37.
5

NMML 45.

NMML 44.

NMML 46.

NMML 43.

Family Loliginidae (Cont.).

NMML 42. Loliqo opalescens

Collected 29 January 1961, Bonella Grounds, Hecate

Strait, British Columbia, by K. Niggol and J.

Maddox. From trawler Sogn. Parts saved: 3 entire

specimens.

Loligo opalescens

Collected 17 March 1966, lat. 34°26'N,

long. 120024'W, east of Pt. Conception, California,

by R. K. Stroud. Water temperature 14°C. Parts

saved: 17+ small entire specimens.

Loligo opalescens

Collected 13 February 1968, off Cape Lookout,

Oregon, by R. K. Stroud. From trawler Ivanhoe.

Parts saved: 6 small entire squid.

Loligo opalescens

Collected 23 January 1961, lat. 49°00'N,

long. 125°49'W. From stomach of Q. ursinus,

specimen US61-1181. Parts saved: 6+ relatively

entire specimens plus beaks.

Loligo opalescens

Collected 27 January 1961, lat. 52°33'N,

long. 128°51'W. From stomach of Q. ursinus,

specimen US61-1194. Parts saved: beaks only.
NMML 48.

NMML 49.

6

Family Loliginidae (Cont.).

NMML 47. Loliqo opalescens

Collected 27 March 1970, lat. 4so46'N,

long. 124°2S'W. From stomach of Q. ursinus,

specimen US70-125. Parts saved: beaks only.

Loliqo opalescens

Collected 17 May 1970, lat. 46°27'N, long. 128°3S'W.

From stomach of Q. ursinus, specimen US70-271.

Parts saved: 2 pair beaks.

Loligo opalescens

NMML 50.

NMML 51.

NMML 52.

NMML 53.

Collected 19 May 1970, 46°46'N, 128 °35'W. long. lat.

specimen US70-293. ursinus,

saved:

1 pair beaks.

LoligQ QQalescens Collected 20 May 1970,

46°43'N, 127°37'W, long. lat.

sinus,

specimen US70-303.

Parts saved: 1 pair beaks.

Loligo opalescens

Collected (no date), puget Sound, Washington. Parts

saved: 1 small entire specimen.

Loligo opalescens

Collected (no date), off California? Parts saved:

2 pair beaks only - working collection.

Loliqo opalescens

Collected (no date), off California? Parts saved:

beaks only.
7

Family Loliginidae (Cont.).

NMML 54. Loligo opalescens

Collected (no date). Parts saved: pens only.

NMML 214. Loligo opalescens

Collected 26 April 1964, lat. 38°01'N,

long. 123°21'W. From stomach of ~. ursinus,

specimen US64-105. Parts saved: 1 fragment + pens

and buccal masses.

NMML 228. Loligo opalescens

Collected 29 January 1982, Elliott Bay, puget Sound,

Washington, by R. W. Mercer. By light attraction at

night in Lampera-type net. Female, nidamental gland

length 26 rom, DML 112 rom, weight 39 g. Reproductive

development intermediate. Parts saved: 2

statoliths.

NMML 229. Loligo opalescens

Collected 29 January 1982, Elliott Bay, puget Sound,

Washington, by R. W. Mercer. By light attraction at

night in Lampera-type net. Male, DML 128 rom, weight

51 g. Reproductive development immature. Parts

saved: beaks and radula.

NMML 229a. 2 statoliths of 229.
8

Family Loliginidae (Cont.)

NMML 230. Loligo opalescens

Collected 29 January 1982, Elliott Bay, puget Sound,

Washington, by R. W. Mercer. By light attraction at

night in Lampera-type net. Male, DML 127 rom, weight

50 g. Reproductive development immature. Parts

saved: 2 statoliths.

Loligo opalescens

Collected 29 January 1982, Elliott Bay, puget Sound,

Washington, by R. W. Mercer. By light attraction at

night in Lampera-type net. Female, nidamental gland

length 47 rom, DML 155 rom, weight 78 g. Reproductive

development mature. Parts saved: beaks and radula.

2 statoliths of 231.

NMML 231.

NMML 231a.

NMML 232.

NMML 232a.

NMML 233.

Loliqo opalescens

Collected 29 January 1982, Elliott Bay, puget Sound,

Washington, by R. W. Mercer. By light attraction at

night in Lampera-type net. Male, DML 115 rom, weight

45 g. Reproductive development immature. Parts

saved: beaks and radula.

2 statoliths of 232.

Loligo opalescens

Collected 29 January 1982, Elliott Bay, puget Sound,

Washington, by R. W. Mercer. By light attraction at

night in Lampera-type net. Female, nidamental gland

length 42 rom, DML 132 rom, weight 64 g. Reproductive

development mature. Parts saved: 2 statoliths.
9

Family Loliginidae (Cont.).

NMML 234. Loligo opalescens

Collected 29 January 1982, Elliott Bay, Puget Sound,

Washington, by R. W. Mercer, by light attraction at

night in Lampera-type net. Female, nidamental gland

length 12 mm, DML 89 mm, weight 18 g. Reproductive

development intermediate. Parts saved: beaks and

radula.

2 statoliths of 234.

Loligo opalescens

Collected 10 December 1983 off Redondo, Puget Sound,

Washington, by R. Lauzon and R. W. Mercer, by light

attraction at night in Lampera-type net. Male, DML

206 mm, weight 142 g. Reproductive development

mature - spermatophores present. Parts saved: 1

entire specimen.

Beaks and radula of 414.

2 statoliths of 414.

NMML 234a.

NMML 414.

NMML 414a.

NMML 414b.

NMML 415.

NMML 415a.

Loligo opalescens

Collected 10 December 1983 off Redondo, puget Sound,

Washington, by R. Lauzon and R. W. Mercer, by light

attraction at night in Lampera-type net. Male, DML

192 mm, weight 136 g. Reproductive development

mature - spermatophores present. Parts saved: 1

entire specimen.

Beaks and radula of 415.
NMML 416a.

NMML 416b.

NMML 417.

10

Family Loliginidae (Cont.).

NMML 415b. 2 statoliths of 415.

NMML 416. Loligo opalescens

Collected 10 December 1983 off Redondo, puget Sound,

Washington, by R. Lauzon and R. W. Mercer, by light

attraction at night in Lampera-type net. Male, DML

182 mrn, weight 112 g. Reproductive development

immature. Parts saved: 1 entire specimen.

Beaks and radula of 416.

1 statolith of 416.

Loliqo opalescens

Collected 10 December 1983 off Redondo, Puget Sound,

Washington, by R. Lauzon and R. W. Mercer, by light

attraction at night in Lampera-type net. Male, DML

180 mrn, weight 111 g. Reproductive development

immature. Parts saved: 1 entire specimen.

NMML 417a.

NMML 417b.

NMML 418.

NMML 418a.

NMML 418b.

Beaks and radula of 417.

1 statolith of 417.

Loligo opalescens

Collected 10 December 1983 off Redondo, puget Sound,

Washington, by R. Lauzon and R. W. Mercer, by light

attraction at night in Lampera-type net. Male, DML

180 mrn, weight 117 g. Reproductive development

immature. Parts saved: 1 entire specimen.

Beaks and radula of 418.

2 statoliths of 418.
11

Family Loliginidae (Cont.).

NMML 419. Loligo opalescens

Collected 10 December 1983 off Redondo, Puget Sound,

Washington, by R. Lauzon and R. W. Mercer, by light

attraction at night in Lampera-type net. Male, DML

183 rom, weight 103 g. Reproductive development

mature - spermatophores present. Parts saved: 1

entire specimen.

Beaks and radula of 419.

2 statoliths of 419.

NMML 419a.

NMML 419b.

NMML 420.

NMML 420a.

NMML 4210b.

NMML 421.

Loligo opalescens

Collected 10 December 1983 off Redondo, puget Sound,

Washington, by R. Lauzon and R. W. Mercer, by light

attraction at night in Lampera-type net. Male, DML

189 rom, weight 129 g. Reproductive development

mature - spermatophores present. Parts saved: 1

entire specimen.

Beaks and radula of 420.

1 statolith of 420.

Loligo opalescens

Collected 10 December 1983 off Redondo, puget Sound,

Washington, by R. Lauzon and R. W. Mercer, by light

attraction at night in Lampera-type net. Male,

DML 180 rom, weight 125 g. Reproductive development

mature - spermatophores present. Parts saved:

1 entire specimen.
12

Family Loliginidae (Cont.).

NMML 421a.

NMML 421b.

NMML 422

NMML 422a.

NMML 422b.

NMML 423.

NMML 423a.

NMML 424.

Beaks and radula of 421.

2 statoliths of 421.

Loligo opalescens

Collected 10 December 1983 off Redondo, puget Sound,

Washington, by R. Lauzon and R. W. Mercer, by light

attraction at night in Lampera-type net. Male

DML 192 rom, weight 137 g. Reproductive development

mature - spermatophores present. Parts saved:

1 entire specimen.

Beaks and radula of 422

2 statoliths of 422.

Loligo opalescens

Collected 11 January 1984 (1945-2100 hrs.) at

Edmonds, Puget Sound, Washington, by R. W. Mercer,

by jigging off fishing pier. Female, DML 170 rom.

Reproductive development mature - eggs. Parts

saved: Buccal mass. Det. by R. W. Mercer and S.

Maupin.

2 statoliths of 423.

Loligo opalescens

Collected 11 January 1984 (1945-2100 hrs.) at

Edmonds, puget Sound, Washington, by R. W. Mercer,

by jigging off fishing pier. Male, DML 175 rom.

Reproductive development mature. Parts saved:

Buccal mass. Det. by R. W. Mercer and S. Maupin.
13

Family Loliginidae (Cont.).

NMML 424a. 2 statoliths of 424.

NMML 425. Loliqo opalescens

Collected 11 January 1984 (1945-2100 hrs.) at

Edmonds, puget Sound, Washington, by R. W. Mercer,

by jigging off fishing pier. Male, DML 170 rom.

Reproductive development mature. Parts saved:

Buccal mass. Det. by R. W. Mercer and S. Maupin.

NMML 425a. 2 statoliths of 425.

NMML 426.

NMML 426a.

NMML 454.

Loligo opalescens

Collected 11 January 1984 (1945-2100 hrs.) at

Edmonds, puget Sound, Washington, by R. W. Mercer,

by jigging off fishing pier. Male, DML 160 rom.

Reproductive development mature. Parts saved:

Buccal mass. Det. by R. W. Mercer and S. Maupin.

2 statoliths of 426.

Loligo opalescens

Collected 4 May 1984, lat. 57°37.5'N,

long. 136°23.0'W, Salisbury Sound, Alaska by R. W.

Mercer. NOAA ship Chapman Cruise CH-84-03 by bottom

trawl. Depth 69 fathoms. Water temp. on bottom

6.9°C. Time: 1224 hrs. Reproductive development -

(Maturity): females 95%, males 60%. Parts saved:

58 entire specimens. Det. R. W. Mercer. (See Wing

and Mercer 1990 - west of Myriad Islands specimens.)
14

Family Lo1iginidae (Cont.).

NMML 457.

NMML 519.

NMML 519a.

NMML 519b.

Lo1igo o~a1escens

Collected 18 December 1987, Elliott Bay, puget

Sound, Washington, by H. Kajimura, by jigging under

lights at night. Sex, length, and weight of 15

specimens.

No. Length SexWeight

(mm)

(g)

M 190145

M 178130

M 177130

M 3543176120F17085M 178133

M 181135M 196155M181120

F16880

M202165

M177120197155F15390

15 pairs beaks.

Loligo opa1escens

Collected 4 August 1990, lat. 44°38'N,

long. 124°31'W, by jig 60 m., V37H5-1 U.S.-Japan

Cooperative Survey. Frozen upon capture, thawed in

laboratory. Lab measurements. Male, DML 127 mm,

weight 60 g. Parts saved: entire specimen.

Beaks and radu1a of 519. LRL 1.5 mm.

2 statoliths of 519.
15

NMML 35.

NMMl 55.

NMML 456.

NMML 520a.

NMML 520b.

NMML 58.

Family Loliginidae (Cont.).

NMML 520. Loliqo opalescens

Collected 4 August 1990, lat. 44°38'N,

long. 124°31'W, by jig 60 m., V37H5-2 U.S.-Japan

Cooperative Survey. Frozen upon capture, thawed in

laboratory. Lab measurements. Male, DML 108 rom,

weight 50 g. Parts saved: entire specimen.

Beaks and radula of 520. LRL 1.5 rom.

2 statoliths of 520.

Loligo pealei?

Collected (no date), Atlantic Ocean. Specimen

504315. Parts saved: beaks only.

Loligo pealei

Collected lat. 29°57.0'N, long. 86°45.0'W, by R. W.

Mercer. Depth 75 fathoms. Male, DML 297 rom. Parts

saved: statoliths. Det. R. W. Mercer.

Doryteuthis bleekeri

Collected 24 April 1965, Kesennuma, Miyagi

Prefecture, Honshu, Japan, by H. Kajimura. Parts

saved: 4 entire specimens.

Lolliguncula brevis (Fitch and Brownell 1971)

Collected 7 February 1969, Uruguay, South America.

From the stomach of Pontophoria blainvillei,

specimen R.L.B. 465. Parts saved: beaks only.

Identification verified by G. L. Voss (letter 19

November 1969) .
16

Family Loliginidae (Cont.).

NMML 473. Lolliguncula brevis

Collected 20 January 1964, Pens Bay, Newfoundland,

Canada, by Frederick Aldrich. Parts saved: 2 entire

specimens. DML 44 and 41 rom.
17

Order Teuthida

Suborder Oegopsida

Family Enoploteuthidae

NMML 67.

NMML 68.

NMML 69.

NMML 70.

NMML 70a.

Abraliopsis affinis

Collected 9 April 1968, lat. 12°51'N, long. 93°18'W,

by W. Perrin. From the stomach of Stenella

graffmani, specimen CV285. Parts saved: 1 entire

specimen. Det. T. Okutani.

Abraliopsis sp.

Collected 9 April 1968, lat. 12°51'N, long. 93°18'W.

From the stomach of ~. graffmani, specimen CV285.

Parts saved: body fragments and beaks.

AbralioDsis Sp.l

Collected 22 January 1961, lat. 34°23'N,

long. 121°08'W. From the stomach of b. ursinus,

specimen US61-242. Parts saved: 2 entire specimens

+ 3 body fragments.

Abraliopsis Sp.l

Collected 19 March 1961, lat. 37°02'N,

long. 123°16'W. From the stomach of b. ursinus,

specimen US61-773. Parts saved: 24+ relatively

entire specimens.

Abraliopsis Sp.l

5 pens of 70.

IMost likely A. felis.
18

Family Enoploteuthidae (Cont.).

NMML 71. Abraliopsis Sp.l

Collected 19 March 1961, lat. 37°02'N,

long. 123°17'W. From the stomach of ~. ursinus,

specimen US61-774. Parts saved: 10 relatively

entire specimens plus vial of beaks.

NMML 71a. Abraliopsis Sp.l

1 entire specimen of 71.

NMML 71b. Abraliopsis sp.t

1 entire specimen of 71.

NMML 71c. Abraliopsis sp.t

1 entire specimen of 71.

NMML 72. Abraliopsis Sp.l

Collected 21 May 1965, lat. 37°49'N, long. 123°27'W.

From the stomach of ~. ursinus, specimen US6S-110.

Parts saved: beaks only.

NMML 73. Abraliopsis sp. (see MacAskie 1971)

Collected 2 May 1970, lat. 46°38'N, long. 126°37'W.

From the stomach of ~. ursinus, specimen US70-209.

Parts saved: 1 pair beaks.

NMML 74. Abraliopsis sp. (MacAskie 1971)

Collected 17 May 1970, lat. 46°27'N, long. 128°35'W.

From the stomach of ~. ursinus, specimen US 70-271.

Parts saved: 1 pair beaks.
NMML 64.

19

Family Enoploteuthidae (Cont.).

NMML 75. Abraliopsis sp. (MacAskie 1971)

Collected 19 May 1970, lat. 46°42'N, long. 129°06'W.

From the stomach of ~. ursinus, specimen US 70-283.

Parts saved: 1 upper beak.

NMML 76. Abraliopsis sp. (MacAskie 1971)

Collected 20 May 1970, lat. 46°41'N, long. 127°27'W.

From the stomach of ~. ursinus, specimen US70-308.

Parts saved: 1 pair beaks.

NMML 77. Abraliopsis sp.

Collected (date and location unknown). Parts saved:

1 relatively entire specimen.

Watasenia scintillans

Collected April 1950, Miyagi Prefecture, Honshu,

Japan, by F. Wilke. Parts saved: beaks. Det. F.

Wilke.

NMML 65.

NMML 66.

Watasenia scintillans

Collected 15 May 1950, Hachinohe, Aomori Prefecture,

Honshu, Japan, by F. Wilke. From the stomach of

Phocoenoides dalli. Parts saved: pens. Det. F.

Wilke.

Watasenia scintillans

Collected 11 March 1965, lat. 36°50'N,

long. 141°17'E, by H. Kajimura. From the stomach of

~. ursinus. Surface water temperature 10°C. Parts

saved: 10 entire specimens. Det. H. Kajimura.
20

Order Teuthida

Suborder Oegopsida

Family Enoploteuthidae

Subfamily pyroteuthinae

NMML 436. ?Pterygioteuthis qiardi (gemmata - Jefferts 1983)

Collected 18 August 1984, Guadalupe Island, Mexico,

by R. L. DeLong, from the scat of Arctocephalus

townsendi, specimen At84-13a. Parts saved: 1 upper

beak.
21

Order Teuthida

Suborder Oegopsida

Family Octopoteuthidae

NMML 144. Octopoteuthis sp. (deletron?) (see Young 1972)

NMML 145.

NMML 437.

NMML 474.

NMML 475.

Collected 8 February 1964 off Oregon, by W. E.

Pearcy. MT473 0-1000 m. Specimen NH50. Parts

saved: 1 pair beaks. Det. W. E. Pearcy.

Octopoteuthis deletron

Collected 20 April 1967, Mukkaw Bay, Washington by

R. K. Stroud. From the stomach of stranded Grampus

griseus, specimen 3769. Parts saved: 1 pair beaks,

crown fragments.

Octopoteuthis deletron

Collected 29 April 1984, San Miguel Island,

California, by G. A. Antonelis, by lavage from

stomach of Mirounga angustirostris, specimen A-27.

Parts saved: 1 lower beak.

Octopoteuthis deletron

Collected 11 June 1964, off central California.

From stomach of male ~. macrocephalus, specimen Rice

1964-1054. Parts saved: 1 upper beak, 1 lower beak.

LRL 9.0 mm.

Octopoteuthis deletron

Collected 3 May 1967, off central California. From

stomach of female ~. macrocephalus, specimen Rice

1967-114. Parts saved: 1 lower beak. LRL 7.5 mm.
NMML 491.

NMML 478.

22

Family Octopoteuthidae (Cont.).

NMML 476. Octopoteuthis deletron

Collected 23 April 1968, off central California.

From stomach of female R. macrocephalus, specimen

Rice 1968-108. Parts saved: 2 upper beaks, 2 lower

beaks. LRL 7.0 and 7.0 rom.

NMML 477. Octopoteuthis deletron

Collected 29 April 1968, off central California.

From stomach of female R. macrocephalus, specimen

Rice 1968-111. Parts saved: 1 upper beak, 2 lower

beaks. LRL 7.0 and 7.2 rom.

Taningia danae

Collected 25 November 1966, off central California.

From stomach of male R. macrocephalus, specimen Rice

1966-165. Parts saved: lower beak. LRL 12.2 rom.

Taningia danae

Collected 19 July 1990, West Cove, San Miguel

Island, California. From stomach of male Mirounga

angustirostris by lavage. Specimen Y1991/1992.

Parts saved: 2 lower beaks. LRL 15.0 and 8.0 rom,

NMML 492.

and 3 upper beaks.

Taningia danae

Collected 7 July 1989, San Miguel Island,

California. From stomach of male Mirounqa

angustirostris by lavage. Specimen TX750. Parts

saved: 3 pairs? beaks. LRLs 16.0, 12.5, and

11.2 rom.
23

Order Teuthida

Suborder Oegopsida

Family Onychoteuthidae

NMML 331.

NMML 332.

NMML 432.

NMML 142.

Onychoteuthis banksii

Collected 9 April 1968, lat. 12°51'N, long. 93°18'W.

From the stomach of Stenella sp., specimen CV285.

Parts saved: fragments of 3 specimens, buccal masses

of 4, including 4 pair beaks, and 1 pair beaks.

Onychoteuthis banksii

Collected 9 April 1968, lat. 12°51'N, long. 93°18'W.

From the stomach of Stenella sp., specimen CV285.

Loaned to C. F. E. Roper, U. S. National Museum,

18 October 1972.

Onychoteuthis banksii

Collected 18 August 1984, Guadalupe Island, Mexico,

by R. L. DeLong, from the scat of Arctocephalus

townsendi, specimen At-84-34a. Parts saved:

12 upper and 12 lower beaks.

Onychoteuthis borealijaponica

Collected 24 July 1957, south of Aleutian Islands,

by R. A. Hajny. Parts saved: 1 entire specimen,

DML 343 rom.
24

NMML 213.

NMML 212.

NMML 216.

NMML 216a.

NMML 216b.

Family Onychoteuthidae (Cont.).

NMML 211. Onychoteuthis borealiiaponica

Collected 2 July 1981, over Grays Canyon,

approximately 40 miles west of Westport, Washington,

by M. Bucy. FV Tres Cher, by dipnet-nightlight.

Female, DML 120 rom. Identified from whole specimen.

Parts saved: beaks only.

Onychoteuthis borealiiaponica

Collected 2 July 1981, over Grays Canyon,

approximately 40 miles west of Westport, Washington,

by M. Bucy. FV Tres Cher, by dipnet-nightlight.

Male, DML 162 rom. Identified from whole specimen.

Parts saved: beaks and radula.

Onychoteuthis borealijaponica

Collected 2 July 1981, over Grays Canyon,

approximately 40 miles west of Westport, Washington,

by M. Bucy. FV Tres Cher, by dipnet-nightlight.

Female, DML 225 rom. Identified from whole specimen.

Parts saved: beaks and radula.

Onychoteuthis borealijaponica

Collected 3 August 1981, lat. 47°21.2'N,

long. 124°54.5'W, by R. W. Mercer (Mercer and Bucy

1983). By jigging. Surface water temperature

16.5°C. Immature female, DML 186 rom, weight 176 g.

Parts saved: crown and pen.

Beaks of 216.

1 statolith of 216.
25

Family Onychoteuthidae (Cont.).

NMML 217.

NMML 217a.

NMML 218.

NMML 218a.

NMML 218b.

NMML 219.

NMML 219a.

NMML 219b.

NMML 220.

Onychoteuthis borealijaoonica

Collected 3 August 1981, lat. 47°21.2'N,

long. 124°54.5'W, by R. W. Mercer. By jigging.

Surface water temperature 16.5°C. Immature male,

DML 173 rom, weight 144 g. Parts saved: beaks.

2 statoliths of 217.

Onychoteuthis borealiiaponica

Collected 3 August 1981, lat. 47°21.2'N,

long. 124°54.5'W, by R. W. Mercer. By jigging.

Surface water temperature 16.5°C. Immature female,

DML 192 rom, weight 188 g. Parts saved: crown and

pen.

Beaks and radula of 218.

2 statoliths of 218.

Onychoteuthis borealijaponica

Collected 3 August 1981, lat. 47°21.2'N,

long. 124°54.5'W, by R. W. Mercer. By jigging.

Surface water temperature 16.5°C. Immature female,

DML 192 rom, weight 198 g. Parts saved: crown and

pen.

Beaks and radula of 219.

1 statolith of 219.

Onychoteuthis borealijaponica

Collected 3 August 1981, lat. 47°21.2'N,

long. 124°54.5'W, by R. W. Mercer. By jigging.

Surface water temperature 16.5°C. Immature male,

DML 182 rom, weight 141 g. Parts saved: beaks.
NMML 222.

NMML 222a

NMML 223.

NMML 223a.

NMML 224.

26

Family Onychoteuthidae (Cont.).

NMML 220a. 2 statoliths of 220.

NMML 221. Onvchoteuthis borealiiaponica

Collected 11 May 1981, lat. 47°26'N,

long. 125°09.7'W, by R. W. Mercer. By jigging.

Female, DML 251 rom. (field measurement). Parts

saved: 1 statolith.

Onychoteuthis borealiiaponica

Collected 27 August 1981, lat. 47°22.3'N,

long. 124°56.5'W, by R. W. Mercer. By jigging.

Surface water temperature 15.7°C (6.6°C at 33m).

Female, nidamental gland length 60 rom, DML 255 rom,

weight 448 g. Parts saved: beaks, radula.

2 statoliths of 222.

Onychoteuthis borealiiaponica

Collected 27 August 1981, lat. 47°22.3'N,

long. 124°56.5'W, By R. W. Mercer. By jigging.

Surface water temperature 15.7°C (6.6°at 33m).

Female, nidamental gland length 60 rom, DML 240 rom,

weight 337 g. Parts saved: beaks, radula.

2 statoliths of 223.

Onychoteuthis borealijaponica

Collected 27 August 1981, lat. 47°22.3'N,

long. 124°56.5'W, by R. W. Mercer. By jigging.

Surface water temperature 15.7°C. Male, mature,

many sperm packets, DML 253 rom, weight 345 g. Parts

saved: beaks, radula.
27

Family Onychoteuthidae (Cont.).

NMML 224a. 2 statoliths of 224.

NMML 224b. Male reproductive organs of 224.

NMML 225. Onychoteuthis borealijaponica

Collected 27 August 1981, lat. 47°23.3'N,

long. 125°56.5'W, by R. W. Mercer. By jigging.

Surface water temperature 15.7°C (6.6°C at 33 m).

Male, a few sperm packets present, DML 211 mm,

weight 243 g. Parts saved: beaks, radula.

NMML 225a. 2 statoliths of 225.

NMML 226.

NMML 226a.

NMML 227.

NMML 227a.

Onychoteuthis borealijaponica

Collected 27 August 1981, lat. 47°33.3'N,

long. 125°56.6'W, by R. W. Mercer. By jigging.

Surface water temperature 15.7°C (6.6°C at 33m).

Female, nidamental gland length 55 mm, DML 256 mm,

weight 411 g. Parts saved: beaks, radula.

2 statoliths of 226.

Onychoteuthis borealijaponica

Collected 13 September 1981, lat. 47°19.4'N,

long. 125°QQ'W, by R. W. Mercer. By jigging.

Surface water temperature 14.3°C (8.2°C at 33 mm) .

Frozen and thawed. Female, nidamental gland length

left 93 mm, right 93 mm, DML 288 mm, weight 558 g.

orange in color. Parts saved: beaks, radula.

2 statoliths of 227.
28

Family Onychoteuthidae (Cont.).

NMML 315. Onychoteuthis borealijaponica

Collected 16 August 1966, lat. 50015'N,

long. 176°22'W, by R. D. Carlson. MV Paragon cruise

3S, set 42. Water temperature 10.3°C. Parts saved:

6 pair beaks and radulas.

NMML 316. Onychoteuthis borealijaponica

Collected (date and position not given, but probably

24 July 1967, lat. 49°45'N, long. 176°22'W, as 10

squids examined from this set). RV George Kelez,

cruise 41, set probably lS. Water temperature

10.2°C. Parts saved: 7 pair beaks and radulas.

NMML 317. Onychoteuthis borealiiaponica

Collected 10 September 1969, lat. 52°30'N,

long. 160000'W. RV George Kelez, cruise 47, set 43.

Water temperature 11.7°C. Parts saved: 3 pair beaks

and radulas.

NMML 31S. Onychoteuthis borealijaponica

Collected 21 February 1961, lat. 35°1S'N,

long. 121°34'W. From the stomach of ~. ursinus,

specimen US61-696. Parts saved: 3 upper beaks,

3 lower beaks, 1 buccal mass.

NMML 319. Onychoteuthis borealiiaponica

Collected 19 May 1964, lat. 4000S'N, long. 124°39'W.

From the stomach of ~. ursinus, specimen US64-25S.

Parts saved: 2 lower beaks.
29

Family Onychoteuthidae (Cont.).

NMML 320. Onychoteuthis borealiiaponica

Collected 22 January 1966, lat. 36°53'N,

long. 122°35'W. From the stomach of Q. ursinus,

specimen US66-15. Parts saved: 1 crown, including

radula and beaks.

NMML 321. Onychoteuthis borealiiaponica

Collected 27 March 1970, lat. 46°27'N,

long. 124°25'W. From the stomach of Q. ursinus,

specimen us 70-125. Parts saved: 2 pair beaks.

NMML 322. Onychoteuthis borealiiaponica

Collected 30 March 1970, lat. 47°17'N,

long. 124°58'W. From the stomach of Q. ursinus,

specimen US70-135. Parts saved: 7 upper and 8 lower

beaks.

NMML 323. Onychoteuthis borealijaponica

Collected 21 April 1970, lat. 47°46'N,

long. 126°01'W. From the stomach of Q. ursinus,

specimen US70-194. Parts saved: 14 pair beaks.

NMML 324. Onychoteuthis borealijaponica

Collected 2 May 1970, lat. 46°38'N, long. 126°37'W.

From the stomach of Q. ursinus, specimen US70-209.

Parts saved: 10 upper beaks, 11 lower beaks.

NMML 325. Onychoteuthis borealijaponica

Collected 17 May 1970, lat. 46°29'N, long. 128°46'W.

From the stomach of Q. ursinus, specimen US70-273.

Parts saved: 11 upper beaks, 13 lower beaks.
30

Family Onychoteuthidae (Cont.).

NMML 326. Onychoteuthis borealiiaponica

Collected 20 May 1970, lat. 46°43'N, long. 127°38'W.

From the stomach of ~. ursinus, specimen US 70-302.

Parts saved: 1 pair beaks with buccal mass.

NMML 327. Onvchoteuthis borealijaponica

Collected 20 May 1970, lat. 46°45'N, long. 126°58'W.

From the stomach of ~. ursinus, specimen US70-318.

Parts saved: 19 upper beaks, 20 lower beaks.

NMML 328. Onychoteuthis borealijaponica

Collected (no date), San Miguel Island, California,

by E. C. Jameyson. Specimen ECJ-063. Parts saved:

1 pair beaks.

NMML 342. Onychoteuthis borealijaponica

Collected 1957, North Pacific Ocean, High Sea Salmon

Inventory. Parts saved: crown, gladius, 1 pair

beaks.

NMML 343. Onychoteuthis borealijaponica

Collected 16 August 1966, lat. 50015'N,

long. 176°22'W, by R. D. Carlson. MV Paragon,

cruise 38, set 42. Parts saved: 6 gladii from fresh

whole specimens.

NMML 344. Onychoteuthis borealijaponica

Collected 7 October 1965, lat. 48°22'N,

long. 173°00'E. RV George ~. Kelez, cruise 36,

set 5. Parts saved: gladii from 6 fresh whole specimens.
NMML 346.

NMML 347.

NMML 348.

31

Family Onychoteuthidae (Cont.).

NMML 345. Onychoteuthis borealijaponica

Collected 12 December 1960, lat. 38°41'N,

long. 124°25'W, by C. H. Fiscus. Night Light

Station 4-61. Water temperature 12°C. Parts saved:

5 entire specimens, DMLs 74, 73, 70, 67, and 52 rom,

measured 14 February 1983.

Onychoteuthis borealiiaponica

Collected 9 January 1961, lat. 38°42'N,

long. 123°51'W, by C.H. Fiscus. Night Light Station

6-61. Water temperature 11°C. Parts saved:

7 entire specimens, DMLs of 6 - 80, 71, 70, 65, 58,

and 54 rom, measured 14 February 1983.

Onychoteuthis borealijaponica

Collected 30 January 1961, lat. 33°32'N,

long. 120048'W, by C. H. Fiscus. Night Light

Station 12-61. Water temperature 14°C. Parts

saved: 2 entire specimens, DMLs 87 and 85 rom,

measured 14 February 1983.

Onychoteuthis borealiiaponica

Collected 25 January 1961, lat. 34°25'N,

long. 121°08'W. From the stomach of Q. ursinus,

specimen US61-375. Parts saved: 3 lower and 1 upper

beaks. Examined by E. Hochberg, 1980.
NMML 355.

32

Family Onychoteuthidae (Cont.).

NMML 349. Onychoteuthis borealijaponica

Collected 16 May 1970, lat. 46°24'N, long. 136°21'W.

From the stomach of Q. ursinus, specimen US70-252.

Parts saved: 2 relatively entire specimens, DMLs

approximately 160 rom, measured 14 February 1983.

NMML 350. Onychoteuthis borealijaponica

Collected 24 May 1970, lat. 47°38'N, long. 128°19'W.

From the stomach of C. ursinus, specimen US70-325.

Parts saved: 1 relatively entire specimen,

DML 170 rom, beaks in same container.

Onychoteuthis borealiiaponica

Collected 21 January 1961, lat. 34°35'N,

long. 121°05'W, by C. H. Fiscus. Night Light

Station 10-61. Parts saved: 7 entire specimens,

DMLs 121 112, 112, 107, 104, 72, and 67 rom, measured

NMML 356.

16 February 1983.

Onychoteuthis borealijaponica

Collected 5 March 1962, lat. 42°00'N,

long. 177°52'W, by R. D. Carlson. MV Bertha Ann,

cruise 22, set. 9, measured 29 April 1992. Female,

DML 370 rom, nidamental gland length 84 rom. Parts

saved: 1 entire specimen.
33

Family Onychoteuthidae (Cont.).

NMML 357. Onychoteuthis borealijaponica

Collected 5 March 1962, lat. 42°00'N,

long. 177°52'W, by R. D. Carlson. MV Bertha Ann,

cruise 22, set 9, measured 29 April 1992. Male?

DML 250 rom. Parts saved: 1 entire specimen.

NMML 358. Onychoteuthis borealijaponica

Collected 30 March 1962, lat. 46°54'N,

long. 154°53'W, by R. D. Carlson. MV Bertha Ann,

cruise 22, set. 19. Parts saved: 1 entire specimen.

NMML 359. Onychoteuthis borealiiaponica

Collected 1963, North Pacific, by R. Hitz. RV John

N. Cobb. Parts saved: 2 entire specimens.

NMML 360. Onychoteuthis borealijaponica

Collected 29 June 1967, lat. 48°00'N,

long. 176°21'W. RV George ~. Kelez, cruise 41,

set 6. Parts saved: 1 entire specimen.

NMML 361. Onychoteuthis borealijaponica

Collected September 1969, lat. 51°30' - 52°30'N,

long. 160000'W, by W. Ames. RV George ~. Kelez,

cruise 47, set 42 or 43. Parts saved: 3 entire

specimens - 2 males, 1 female, all immature.

NMML 362. Onychoteuthis borealijaponica

Collected 7 September 1969, lat. 51°00'N,

long. 160000'W, by W. Ames. RV George ~. Kelez,

cruise 47, set 41. Parts saved: 4 entire specimens

- females.
34

Family Onychoteuthidae (Cont.).

NMML 363. Onychoteuthis borealiiaponica

Collected 14 June 1981, lat. 41°N, long. 180°, by C.

Bouchet. RV Oshoro Maru research cruise, set 3, by

surface gill net. Frozen upon capture - thawed in

laboratory. Measured 14 July 1983. Male,

DML 227 rom, weight 231 g. Parts saved: 1 statolith.

Onychoteuthis borealijaponica

Collected 14 June 1981, lat. 41°N, long. 180°, by C.

Bouchet. RV Oshoro Maru research cruise, set 3, by

surface gill net. Measured 14 July 83. Male,

DML 186 rom, weight 109 g. Parts saved: beaks and

radula.

2 statoliths of 364.

NMML 364.

NMML 364a.

NMML 371.

NMML 371a.

Onychoteuthis borealijaDonica

Collected 20 July 1982, lat. 46°58', long. 177°31'W,

by P. J. Gearin. By surface gill net. Frozen upon

capture, thawed and measured in laboratory 20 July

1983. Female, nidamental gland length 59 rom,

DML 294 rom, weight 671 g. Parts saved: beaks and

radula.

2 statoliths of 371.
35

Family Onychoteuthidae (Cont.).

NMML 502. Onychoteuthis borealijaponica

Collected 14 August 1990, lat. 42°52'N,

long. 125°44'W, by jig 90 m, V37H15-1, U.S.-Japan

Cooperative Survey. Frozen upon capture, thawed in

laboratory. Lab measurements, female, DML 307 rom,

weight 600 g. Nidamental gland length 38 rom. Parts

saved: entire specimen.

Beaks and radula of 502. LRL 5.8 rom.

1 statolith of 502.

NMML 502a.

NMML 502b.

NMML 503.

NMML 503a.

NMML 503b.

NMML 504.

NMML 504a.

NMML 504b.

Onychoteuthis borealijaponica

Collected 19 August 1990, lat. 44°13'N,

long. 125°21'W, by jig 130 m, V68H19-3, U.S.-Japan

Cooperative Survey. Frozen upon capture, thawed in

laboratory. Lab measurements, female, DML 322 rom,

weight 880 g. Nidamental gland length 79 rom. Parts

saved: entire specimen.

Beaks and radula of 503. LRL 6.0 rom.

2 statoliths of 503.

Onychoteuthis borealijaponica

Collected 19 August 1990, lat. 44°13'N,

long. 125°21'W, by jig 130 m, V68H19-4, U.S.-Japan

Cooperative Survey. Frozen upon capture, thawed in

laboratory. Lab measurements, female, DML 308 rom,

weight 730 g. Nidamental gland length 54 rom. Parts

saved: entire specimen.

Beaks and radula of 504. LRL 5.6 rom.

2 statoliths of 504.
36

Family Onychoteuthidae (Cont.).

NMML 505. Onvchoteuthis borealijaponica

Collected 19 August 1990, lat. 44°13'N,

long. 125°21'W, by jig 130 m, V68H19-5, U.S.-Japan

Cooperative Survey. Frozen upon capture, thawed in

laboratory. Lab measurements female, DML 297 rom,

weight 570 g. Nidamental gland length 40 rom. Parts

saved: entire specimen.

Beaks and radula of 505. LRL 6.0 rom.

2 statoliths of 505.

Onychoteuthis borealiiaponica

Collected 8 August 1990, lat. 46°29'N,

long. 125°51'W, by jig 60 m, V37H9-3, U.S.-Japan

Cooperative Survey. Frozen upon capture, thawed in

laboratory. Lab measurements male, DML 165 rom,

weight 130 g. Parts saved: entire specimen.

Beaks and radula of 506. LRL not measureable.

1 statolith of 506.

NMML 505a.

NMML 505b.

NMML 506.

NMML 506a.

NMML 506b.

NMML 507.

NMML 507a.

NMML 507b.

Onychoteuthis borealijaponica

Collected 9 August 1990, lat. 45°49'N,

long. 124°36'W, by jig 60 m, V37H10-1, U.S.-Japan

Cooperative Survey. Frozen upon capture, thawed in

laboratory. Lab measurements male, DML 276 rom,

weight 260 g. Parts saved: entire specimen.

Beaks and radula of 507. LRL 5.0 rom.

2 statoliths of 507.
NMML508a.

NMML508b.

NMML 509.

37

Family Onychoteuthidae (Cont.).

NMML 508. Onychoteuthis borealijaponica

Collected 24 August 1990, lat. 46°23'N,

long. 126°30'W, by jig 90 m, V37H25-2, U.S.-Japan

Cooperative Survey. Frozen upon capture, thawed in

laboratory. Lab measurements male, DML 233 rom,

weight 280 g. Parts saved: entire specimen.

Beaks and radula of 508. LRL 5.2 rom.

2 statoliths of 508.

Onychoteuthis borealijaponica

U.S.-Japan Cooperative Survey 1990. Spermiduct and

related organs - various stages of development,

NMML 143.

spermatophores.

a. V58H19-2 - Collected 19 August 1990,

lat. 44°12'N, long. 124°54'W. Male, DML 239 rom,

weight 320 g. Gonad weight 19.62 g.

b. V63H4-1 - Collected 4 August 1990,

lat. 44°56'N, long. 124°52'W. Male, DML 207 rom,

weight 200 g. Gonad weight 7.1 g.

c. V37Hl-4 - Collected 1 August 1990,

lat. 42°57'N, long. 124°57'W. Male, DML 228 rom,

weight 270 g. Gonad weight 8.24 g.

Onychoteuthidae (Chaunoteuthis?)

Collected 20 May 1972, lat. 15°51'N, long. 105°45'W,

by C. W. Oliver. MV Lois Seaver I, by dip net.

Det. by C. H. Fiscus and D. Holts. Shipped to C. F.

E. Roper, 18 October 1972, to verify.
NMML 241.

38

Family Onychoteuthidae (Cont.).

NMML 208. Moroteuthis robusta

Collected 25 September 1960, off central California.

From the stomach of Physeter macrocephalus, specimen

Rice 287. Parts saved: 26 upper beaks, 22 lower

beaks. LRLs 12.0, 11.5, 11.5, 11.0, 11.0, 11.0,

10.5, 10.5, 10.5, 10.5, 10.0, 9.5, 9.0, and 8.0 rom.

Note: Three upper and 8 lower beaks (not paired)

deposited in the collection of the Marine Biological

Association of the United Kingdom, Plymouth,

England, June 1981. F12 Plymouth 81.

Moroteuthis robusta

NMML 329.

NMML 330.

Collected 20 July 1981, lat. 50019.9'N,

long. 174°21.7'E, by P. J. Gearin. In surface gill

net. DML 130 cm, TL 369 cm (field measurements) .

Parts saved: 1 pair beaks, stomach contents.

Moroteuthis robusta

Collected 8 September 1962, off central California.

From stomach of male £. macrocephalus, specimen

Rice-723. Parts saved: 1 pair beaks and radula.

LRL 7.5 rom.

Moroteuthis robusta

Collected 20 May 1970, lat. 46°45'N, long. 126°58'W.

From stomach of Q. ursinus, specimen US70-318.

Parts saved: 1 pair beaks.
NMML 337.

39

Family Onychoteuthidae (Cont.).

NMML 336. Moroteuthis robusta

Collected (date unknown) off west coast Vancouver

Island, British Columbia, by F. Wilke. From stomach

of male g. macrocephalus. Parts saved: 3 upper and

6 lower beaks.

Moroteuthis robusta

NMML 338.

NMML 339.

NMML 340.

Collected 21 May 1965, lat. 37°49'N, long. 123°27'W.

From stomach of ~. ursinus, specimen US65-110.

Parts saved: cartilagenous end cone of gladius and

flesh.

Moroteuthis robusta

Collected 23 August 1959, off central California.

From the stomach of a male g. macrocephalus,

specimen Rice 128. Parts saved: cartilagenous end

cone of gladius.

Moroteuthis robusta

Collected 25 October 1965, off central California.

From stomach of male g. macrocephalus, specimen Rice

1966-155. Parts saved: 1 pair beaks, radula, and

cartilagenous end cone and gladius fragment.

LRL 9.0 rom.

Moroteuthis robusta

Collected 24 July 1969, off central California.

From stomach of male g. macrocephalus, specimen Rice

1969-158. Parts saved: 3 lower beaks. LRLs 7.5,

11.0, and 12.5 rom.
NMML 352.

40

Family Onychoteuthidae (Cont.).

NMML 341. Moroteuthis robusta

Collected 12 May 1970, off central California. From

stomach of female R. macrocephalus, specimen Rice

1970-22. Parts saved: 2 pair beaks (larger pair

LRL 12.5 rom) the smaller pair a good example of

color development and for comparison with similar

size O. borealijaponica.

NMML 351. Moroteuthis robusta

Collected 5 September 1963, off central California.

From stomach of male P. macrocephalus, specimen Rice

932. Parts saved: 1 gladius fragment and

cartilagenous end cone.

Moroteuthis robusta

NMML 353.

NMML 354.

Collected 8 March 1966, lat. 35°18'N,

long. 121°44'W, by R. Stroud. Picked up dead on

surface. Parts saved: gladius and cartilagenous end

cone (end cone 257 rom).

Moroteuthis robusta

Collected 1963, off Oregon-Washington. RV John N.

Cobb. Parts saved: 1 entire specimen, DML 346 rom.

Moroteuthis robusta (Pattie 1968)

Collected January 1965, off Strait of Juan de Fuca,

by B. H. Pattie. By otter trawl. Parts saved: 2

arms, gladius, radula, DML 860 rom. Cartilagenous

end cone and beaks loaned to I. B. MacAskie, Fish.

Res. Board of Canada, Nanaimo, B. C., 13 November 1970.
NMML 513.

41

Family Onychoteuthidae (Cont.).

NMML 388. Moroteuthis robusta

Collected 1 November 1967, off central California.

From the stomach of a female g. macrocephalus,

specimen Rice 1967-234. Parts saved: 1 upper beak.

NMML 389. Moroteuthis robusta

Collected 24 July 1969, off central California.

From the stomach of a male g. macrocephalus,

specimen Rice 1969-157. Parts saved: 1 pair beaks

and radula (nice example of relatively small beaks) .

LRL 6.0 rom.

Moroteuthis robusta

NMML 513a.

NMML 513b.

NMML 514.

Collected 24 August 1990, lat. 46°23'N,

long. 126°30'W, by jig, 90 M, V37H25-1, U.S.-Japan

Cooperative Survey. Frozen upon capture, thawed in

laboratory. Lab measurements female DML 478 rom,

weight 1,460 g. Nidamental gland length 63.0 rom.

Parts saved: entire specimen.

Beaks and radula of 513. LRL 6.8 rom.

2 statoliths of 513.

Moroteuthis robusta

Collected 31 August 1990, lat. 44°05'N,

long. 126°56'W, by jig, 90 m, V37H32-1, U.S.-Japan

Cooperative Survey. Frozen upon capture, thawed in

laboratory. Lab measurements male, DML 694 rom,

weight 3,780 g. Parts saved: entire specimen.
42

Family Onychoteuthidae (Cont.).

NMML 514a. Beaks and radula of 514. LRL 8.0 rom.

NMML 514b. 1 statolith of 514.

NMML 515. Moroteuthis robusta

Collected 17 August 1990, lat. 43°08'N,

long. 125°38'W, by jig, 100 m, V58-H17-1, U.S.-Japan

Cooperative Survey. Frozen upon capture, thawed in

laboratory. Lab measurements female, DML 490 rom,

weight 720 g. Nidamental gland length 59.0 rom.

Parts saved: crown (head, arms, tentacles).

Beaks and radula of 515. LRL 6.5 rom.

1 statolith of 515.

NMML 515a.

NMML 515b.

NMML 516.

NMML 516a.

NMML 516b.

Moroteuthis robusta

Collected 12 August 1990, lat. 42°15'N,

long. 126°32'W, by jig, 55 m, V68H12-1, U.S.-Japan

Cooperative Survey. Frozen upon capture, thawed in

laboratory. Lab measurements female, DML 440 rom,

weight 1,300 g. Nidamental gland length 46.0 rom.

Gonad weight 1.76 g. Parts saved: 1 tentacle,

1 arm, gladius.

Beaks and radula of 516. LRL 5.2 rom.

2 statoliths of 516.
43

Family Onychoteuthidae (Cont.).

NMML 517. Moroteuthis robusta

Collected 24 August 1990, lat. 46°23'N,

long. 126°30'W, by jig, 90 m. V37H25, U.S.-Japan

Cooperative Survey. Frozen upon capture, thawed in

laboratory. Lab measurements female, DML 810 rom,

weight 6,080 g. Nidamental gland length 120 rom.

Parts saved: entire specimen.

Beaks and radula of 517. LRL 9.0 rom.

2 statoliths of 517.

NMML 517a.

NMML 517b.

NMML 518.

NMML 518a.

Moroteuthis robusta

Collected 24 August 1990, lat. 46°23'N,

long. 126°30'W, by jig, 90 m. V37H25, U.S.-Japan

Cooperative Survey. Frozen upon capture, thawed in

laboratory. Lab measurements female, DML 1,080 rom,

weight 13,140 g. Nidamental gland length 222 rom.

Parts saved: crown (head, arms, tentacles).

Beaks and radula of 518. LRL 11.5 rom.
44

Order Teuthida

Suborder Oegopsida

Family Gonatidae

Listing of genera and species follows that of Okutani et al.,

1988

NMML 305.

NMML 439

NMML 440.

NMML 441.

Gonatus cf. berryi

Collected 30 May 1970, lat. 46°13'N, long. 125°20'W.

From the stomach of ~. ursinus, specimen US70-351.

Parts saved: 3 upper, 2 lower beaks.

Gonatus cf. berryi

Collected 28 April 1984, San Miguel Island,

California, by G. A. Antonelis, by lavage from

stomach of Mirounga angustirostris, specimen A-21.

Parts saved: 1 lower beak. Det. by F. G. Hochberg

and C. H. Fiscus.

Gonatus cf. berryi

Collected 30 April 1984, San Miguel Island,

California, by G. A. Antonelis, by lavage from

stomach of M. angustirostris, specimen A-42. Parts

saved: 2 lower beaks, 1 upper beak. Det. by F. G.

Hochberg and C. H. Fiscus.

Gonatus cf. berryi

Collected 30 April 1984, San Miguel Island,

California, by G. A. Antonelis, by lavage from

stomach of M. anqustirostris, specimen A-43. Parts

saved: 2 lower beaks.
NMML 290.

45

Family Gonatidae (Cont.).

NMML 289. Gonatus madokai 2

Collected 2 May 1970, lat. 46°38'N, long. 126°37'W.

From the stomach of ~. ursinus, specimen US70-209.

Parts saved: 1 pair beaks.

Gonatus madokai

NMML 291.

NMML 292.

NMML 293.

Collected 17 May 1970, lat. 46°28'N, long. 128°28'W.

From the stomach of ~. ursinus, specimen US70-266.

Parts saved: 1 head, arms, tentacles and buccal mass

with 5 rowed radula and 1 pair beaks.

Gonatus madokai

Collected 19 May 1970, lat. 46°46'N, long. 128°35'W.

From the stomach of ~. ursinus, specimen US70-293.

Parts saved: 1 pair beaks.

Gonatus madokai

Collected 20 May 1970, lat. 46°41'N, long. 127°27'W.

From the stomach of ~. ursinus, specimen US70-309.

Parts saved: 1 pair beaks.

Gonatus madokai

Collected 24 May 1970, lat. 47°29'N, long. 128°39'W.

From the stomach of ~. ursinus, specimen US70-328.

Parts saved: 1 pair beaks.

2 Note No. 289-295 identified 1 August 1987 C. H. Fiscus
NMML 295.

46

Family Gonatidae (Cont.).

NMML 294. Gonatus madokai

Collected 24 May 1970, lat. 47°24'N, long. 128°40'W.

From the stomach of ~. ursinus, specimen US70-336.

Parts saved: 1 pair beaks. (Note - good match with

NMML 379a.)

Gonatus madokai

NMML 296.

NMML 379.

NMML 379a.

Collected 24 May 1970, lat. 47°24'N, long. 128°38'W.

From the stomach of ~. ursinus, specimen US70-337.

Parts saved: 1 pair beaks.

Gonatus madokai

Collected 29 July 1973, lat. 56°08'N,

long. 171°01'W. From the stomach of ~. ursinus,

specimen US73-110. Parts saved: 1 pair beaks.

Gonatus madokai

Collected 27 July 1982, lat. 52°56.8'N,

long. 176°01.2'W, by R. W. Mercer. MV Dominator,

cruise 82-1, haul 70, by midwater trawl. Depth

16 fathoms, bottom 2,000 fathoms. Time: 1235 hrs.

DML 210 rom, weight 45 g. Parts saved: 1 entire

specimen. Det. K. Jefferts.

Beaks and radula of 379. LRL 6.0 rom, HL 6.0 rom,

CL 13.0+ rom.
NMML 179.

NMML 427a.

NMML 180.

47

Family Gonatidae (Cont.).

NMML 427. Gonatus madokai

Collected 15 July 1981, lat. 51°19'N,

long. 173°27'E, by P. J. Gearin. From the stomach

of Fratercula cirrhata. Gearin 82TP, by surface

gillnet. DML not measurable - small 30-40 rom.

Parts saved: 10 entire small specimens. Det. by

K. Jefferts.

Beaks, radula, and fragment of 427.

Gonatus madokai?

Collected (no date), North Pacific, probably from

Pacific Salmon Investigations. Parts saved: 4

entire specimens, DMLs 97, 175, 150, and 180 rom.

Gonatus middendorffi

Collected 8 June 1970, lat. 51°03'N, long. 175°01'W.

RV George ~. Kelez, cruise 49, set 30, by surface

gill net. Male, DML 295 rom. Parts saved: 1 entire

NMML 179a.

NMML 183.

NMML 183a.

specimen.

Beaks of 179.

Gonatus middendorffi

Collected 15 July 1966, lat. 50037'N,

long. 176°22'W, by R. D. Carlson. MV Paraqon,

cruise 38, set 16. Parts saved: entire specimen,

DML 257 rom. (tentacle club - 2 large hooks followed

by a row of small hooks). Det. T. Kubodera and K.

Jefferts, 21 March 1983.

Buccal mass and pen of 183.
NMML 188.

NMML 187.

NMML 205.

48

Family Gonatidae (Cont.).

NMML 185. Gonatus middendorffi

Collected 7 August 1968, lat. 51°34'N,

long. 176°33'W. RV Miller Freeman, cruise 44, set

32. Parts saved: 1 entire specimen, DML 82 rom,

tentacle lengths approximately 80 rom (right tentacle

club; 1 large, 1 small hook). Beaks in same jar.

Det. T. Kubodera and K. Jefferts, 21 March 1983.

Gonatus middendorffi

Collected (no date or location). Parts saved: 2

entire specimens, DMLs 62 and 53 rom (one small hook

on tentacle club visible). Det. T. Kubodera and K.

Jefferts, 21 March 1983.

Gonatus middendorffi

Collected (no date or location). Parts saved:

1 entire specimen, DML 99 rom (one tentacle club,

1 large hook, 1 smaller hook distal, 1 or more

proximal hooks). Beaks in same jar. Det. T.

Kubodera and K. Jefferts, 21 March 1983.

Gonatus middendorffi

Collected 31 July 1973, lat. 55°58'N,

long. 170012'W. From stomach of ~. ursinus,

specimen US73-185. Parts saved: 2 entire specimens,

DMLs 51 and 40 rom (one hook visible on tentacle

clubs). Det. T. Kubodera and K. Jefferts, 21 March

1983.
NMML 141

NMML 410a.

NMML 263a.

NMML 410.

49

Family Gonatidae (Cont.).

NMML 263. Gonatus middendorffi

Collected August 1971, between lat. 51° and 52°N,

long. 1700Wand 1700E. RV George ~. Kelez,

cruise 53, by surface gill net. Female, DML 322 rom,

preserved weight 316 g. Parts saved: 1 entire

specimen. (Note: from second bag examined 16

September 1971 by C. H. Fiscus). Det. T. Kubodera

and K. Jefferts, 21 March 1983.

Beaks and radula of 263.

Gonatus middendorffi

Collected 25 July 1982, lat. 58°4.3'N,

long. 179°33.9'W, by R. W. Mercer. MV Dominator,

cruise 82-1, haul 64, by midwater trawl. Depth

350 fathoms, bottom 1,970 fathoms. Time: 0045 hrs.

DML 220 rom. Parts saved: 1 entire specimen.

Det. K. Jefferts and T. Kubodera.

Beaks and radula of 410. LRL 4.5 rom, HL 5.0 rom,

CL 9.0 + rom.

Gonatus middendorffi or ~. madokai

Collected (date and location unknown (Alaskan

waters?) From stomach of male Eumetopias jubatus,

specimen SUV-5-76. Either~. middendorffi or

~. madokai. Parts saved: 1 lower beak.
50

NMML 191a.

NMML 191c.

NMML 191b.

NMML 181a.

NMML 191.

NMML 191d.

NMML 196.

Family Gonatidae (Cont.).

NMML 181. Gonatus onyx

Collected 9 January 1961, lat. 38°42'N,

long. 123°51'W, by C. H. Fiscus. Night Light

Station 6-61. Water temperature 11°C. Parts saved:

6 entire specimens DMLs 63, 55, 42, 45, --, and

36 rom.

Lower beak of largest specimen of 181.

Gonatus ~

Collected 10 January 1961, lat. 38°08'N,

long. 123°35'W, by C. H. Fiscus. Night Light

Station 7-61. Parts saved: 14 entire specimens.

DML size range 38-55 rom.

Gonatus onyx

Arms and tentacles of 2 specimens of 191.

Gonatus onyx

Gladii of 2 specimens of 191.

Gonatus onyx

Beaks of specimens of 191. Radula 5-rowed. DMLs of

specimens 191 a-b-c range from 45 to 46 rom.

Beaks of 55 rom DML specimen of 191.

Gonatus ~

Collected 6 January 1961, lat. 37°41'N,

long. 123°29'W, by C. H. Fiscus. Night Light

Station 5-61. Surface water temperature 11°C.

Parts saved: 2 entire specimens, DMLs 58 rom and 60

rom. Very good condition, not sexed.
NMML 405.

51

Family Gonatidae (Cont.).

NMML 197. Gonatus onyx

Collected 13 March 1961, lat. 37°31'N,

long. 123°06'W, by C. H. Fiscus. Night Light

Station 20-61. Parts saved: 1 entire specimen, DML

22 mm.

Gonatus ~?

Collected 13 March 1961, lat. 37°31'N,

long. 123°06'W, by C. H. Fiscus. Night Light

Station 20-61. Surface water temperature 13°C.

Parts saved: 1 entire specimen. Det. K. Jefferts.

Note: Jeffert's comment - larval, no arm hooks,

alternating 3 and 2 rows of suckers on tentacular

stalk.

NMML 275.

NMML 276.

Gonatus ~

Collected 27 March 1970, lat. 46°27'N,

long. 124°25'W. From the stomach of ~. ursinus,

specimen US70-125. Parts saved: 2 upper, 1 lower

beaks. (identified 9 Aug. 1987 - C. H. Fiscus)

Gonatus ~

Collected 30 March 1970, lat. 47°17'N,

long. 124°58'W. From the stomach of ~. ursinus,

specimen US70-135. Parts saved: 2 upper, 1 lower

beaks. (identified 2 Aug. 1987 - C. H. Fiscus)
52

Family Gonatidae (Cont.).

NMML 277. Gonatus onvx

Collected 2 May 1970, lat. 46°38'N, long. 126°37'W.

From the stomach of ~. ursinus, specimen US70-209.

Parts saved: 4 upper, 5 lower beaks. (identified

9 Aug. 1987 - C. H. Fiscus)

NMML 438. Gonatus onyx

Collected 1 May 1984, San Miguel Island, California,

by G. A. Antonelis, by lavage from stomach

M. angustirostris, specimen A-48. Parts saved:

1 lower, 1 upper beak.

NMML 462. Gonatus onyx

Collected 16 April 1985, lat. 48°00'N,

long. 125°20'W. By E. Hacker, NOAA Platforms of

Opportunity Program Survey, Haul 62. By

"NorEastern" bottom trawl (non-closing). Depth

249 fathoms. DMLs 75, 70, 70, 65, and 64 rom. Parts

saved: 5 pair beaks. Det. E. Hacker.

NMML 463. Gonatus ~

Collected 16 April 1985, lat. 47°58'N,

long. 125°18'W. By E. Hacker, NOAA Platforms of

Opportunity Program Survey, Haul 63. By

"NorEastern" bottom trawl (non-closing). Depth

227 fathoms. DML 65 rom. Parts saved: beaks. Det.

E. Hacker.
NMML 466.

53

Family Gonatidae (Cont.).

NMML 464. Gonatus onyx

Collected 30 April 1985, lat. 47°20'N,

long. 124°28'W. By E. Hacker, NOAA Platforms of

Opportunity Program Survey, Haul 100. By

"NorEastern" bottom trawl (non-closing). Depth

221 fathoms. DML 80 rom. Parts saved: beaks. Det.

E. Hacker.

NMML 465. Gonatus ~

Collected 2 May 1985, lat. 47°49'N, long. 125°12'W.

By E. Hacker, NOAA Platforms of Opportunity Program

Survey, Haul 110. By "NorEastern" bottom trawl

(non-closing). Depth 245 fathoms. DMLs 63, 54, 53,

and 52 rom. Parts saved: 3 pair beaks. Det. E.

Hacker.

Gonatus ~

Collected 2 May 1985, lat. 47°48'N, long. 125°10'W.

By E. Hacker, NOAA Platforms of Opportunity Program

Survey, Haul 111. By "NorEastern" bottom trawl

(non-closing). Depth 243 fathoms. DML 60 rom.

Parts saved: beaks. Det. E. Hacker.

NMML 467. Gonatus onyx

Collected 21 May 1985, lat. 45°16'N, long. 124°24'W.

By E. Hacker, NOAA Platforms of Opportunity Program

Survey, Haul 201. By "NorEastern" bottom trawl

(non-closing). Depth 209 fathoms. DMLs 75 and

40 rom. Parts saved: 2 pair beaks. Det. E. Hacker.
NMML 281.

NMML 380.

NMML 380a.

NMML 407.

54

Family Gonatidae (Cont.).

NMML 468. Gonatus onyx

Collected 21 May 1985, lat. 45°14'N, long. 124°21'W.

By E. Hacker, NOAA Platforms of Opportunity Program

Survey, Haul 202. By "NorEastern" bottom trawl

(non-closing). Depth 202 fathoms. DMLs 75 and

50 mm. Parts saved: 2 pair beaks. Det. E. Hacker.

Gonatus pyros

Collected 1 August 1982, lat. 53°13.6'N,

long. 163°50.3'W, by R. W. Mercer. MV Dominator,

cruise 82-1, haul 75, by midwater trawl. Depth

372 fathoms, bottom 2,300 fathoms. Time: 0031 hrs.

DML 92 mm, weight 31 g. Parts saved: 1 entire

specimen. Det. K. Jefferts.

Beaks and radula of 380.

Gonatus pyros

Collected 28 May 1964, lat. 45°35'N,

long. 124°53.3W. RV Commando, by shrimp trawl.

Depth 750 fathoms. Parts saved: 1 entire specimen.

Det. K. Jefferts. Note: (Jeffert's comment)

DML <35 mm, has remnants of optic photophore.

Gonatus pyros

Collected 20 May 1970, lat. 46°41'N, long. 127°28'W.

From the stomach of Q. ursinus, specimen US70-307.

Parts saved: 2 pair beaks. (identified 2 Aug. 1987

- C. H. Fiscus)
NMML 285.

NMML 287.

NMML 286.

55

Family Gonatidae (Cont.).

NMML 282. Gonatus sp. (type "A") (~. pyros? 2 Aug. 1987 -

C. H. Fiscus). Collected 20 May 1970, lat. 46°41'N,

long. 127°21'W. From the stomach of h. ursinus,

specimen US70-310. Parts saved: 1 pair beaks in

buccal mass.

NMML 284. Gonatus sp. (type "A") (~. pyros? 2 Aug. 1987 -

C. H. Fiscus). Collected 20 May 1970, lat. 46°41'N,

long. 127°27'W. From the stomach of h. ursinus,

specimen US70-308. Parts saved: 1 upper, 2 lower

beaks.

Gonatus pyros

Collected 24 May 1970, lat. 47°38'N, long. 128°27'W.

From the stomach of h. ursinus, specimen US70-326.

Parts saved: 4 pair beaks. (identified 2 Aug. 1987

- C. H. Fiscus)

Gonatus pyros

Collected 30 May 1970, lat. 46°13'N, long. 125°20'W.

From the stomach of h. ursinus, specimen US70-351.

Parts saved: 2 upper, 1 lower beak. (identified

2 Aug. 1987 - C. H. Fiscus)

Gonatus pyros?

Collected 30 May 1970, lat. 46°13'N, long. 125°22'W.

From the stomach of h. ursinus, specimen US70-352.

Parts saved: 2 pair beaks. (identified 2 Aug. 1987

- C. H. Fiscus)
56

Family Gonatidae (Cont.).

NMML 125.

NMML 126.

NMML 128.

NMML 129a.

NMML 134.

Gonatus sp. (fabricii type)3

Collected 26 May 1964, off central California. From

stomach of female P. macrocephalus, specimen Rice

1047. Parts saved: 2 lower beaks, 1 upper beak.

LRL 8.0 and 8.0 rom.

Gonatus sp. (fabricii type)

Collected 20 April 1965, off central California.

From stomach of male ~. macrocephalus, specimen Rice

1965-5. Parts saved: 1 lower beak. LRL 7.5 rom.

Gonatus sp. (fabricii type)

Collected 1 December 1966, off central California. From

stomach of female ~. macrocephalus, specimen Rice 1966-

180. Parts saved: 1 pair beaks. LRL 8.0 rom.

Gonatus sp. (fabricii type)

Collected 9 December 1966, off central California. From

stomach of male ~. macrocephalus, specimen Rice 1966-181.

Parts saved: 1 lower beak. LRL 8.5 rom.

Gonatus sp. (fabricii type)

Collected 17 October 1967, off central California. From

stomach of male ~. macrocephalus, specimen Rice 1967-231.

Parts saved: 1 lower beak (type "2"). LRL 8.5 rom.

3~. fabricii has not been identified from the North Pacific

or its adjacent seas. Beaks similar to this species have been

reported from marine mammal stomachs from the California Current

System. See Fiscus et al. 1989, p. 5 for discussion of Gonatus

sp. (cf. ~. fabricii)
NMML 184.

57

Family Gonatidae (Cont.).

NMML 136. Gonatus sp. (fabricii type)

Collected 23 April 1968, off central California.

From stomach of female E. macrocephalus, specimen

Rice 1968-108. Parts saved: 1 pair beaks

(type "2"). LRL 7.5 rom. F15 PlYmouth 1981. Det.

M. R. Clarke.

NMML 138. Gonatus sp. (fabricii type)

Collected 13 June 1969, off central California.

From stomach of female E. macrocephalus, specimen

Rice 1969-137. Parts saved: 1 lower beak

(type "2"). LRL 8.8 rom. F17 PlYmouth 1981. Det.

M. R. Clarke.

NMML 139. Gonatus sp. (fabricii type)

Collected 25 February 1966, lat. 37°46'N,

long. 123°26'W. From stomach of Lagenorhynchus

obliquidens, specimen 3764. Parts saved: 1 pair

beaks. LRL 9.0 rom.

NMML 447. Gonatus sp. (fabricii type)

Collected 30 April 1984, San Miguel Island,

California, by G. A. Antonelis, by lavage from

stomach of M. angustirostris, specimen A-43. Parts

saved: 1 lower beak. LRL 6 rom.

Gonatus sp.

Collected (no date or location). RV John N. Cobb.

Specimen 1963. Parts saved: 1 gladius only.

DML 180 rom, gladius length 170 rom.
NMML 242.

58

Family Gonatitae (Cont.).

NMML 186. Gonatus sp.

Collected 27 November 1966, off central California.

From the stomach of female £. macrocephalus,

specimen Rice 1966-169. Parts saved: 1 entire

specimen. Badly damaged, beaks dissected out (type

"A") . (Note: On permanent loan to Oregon State

University. )

NMML 190. Gonatus .§n.

Collected 1963 (no location). RV John Q. Cobb.

DML 180 mrn. Parts saved: beaks and radula only

(type "B").

Gonatus .§n.

Collected 3 November 1981, lat. 55°40'N,

long. 168°14'W. RV Miller Freeman, cruise 81-6,

haul 38, by trawl. Depth 73 fathoms. Surface

temperature 6.5°C, trawl temperature 5.6°C. Frozen

fresh. DML 30 mrn approximately. Parts saved:

1 entire specimen (in poor shape). Det. K.

Jefferts.

NMML 297. Gonatus sp.

Collected 25 August 1973, lat. 56°01'N,

long. 170007'W. From the stomach of Q. ursinus,

specimen US73-360. Parts saved: arms/tentacles and

buccal mass/radula and 1 pair beaks. Resembles

undescribed specimen of T. Kubodera. Examined 21

March 1983 by T. Kubodera and K. Jefferts.
NMML 279.

59

Family Gonatidae (Cont.)

NMML 406. Gonatus sp.

Collected 13 April 1964, lat. 35°52'N,

long. 121°52'W, by C. H. Fiscus. Parts saved:

1 entire specimen. Det. K. Jefferts. Note:

(Jeffert's comment) larval (about 6 mm), has 5

alternating rows of suckers on stalk. Probably~.

onyx or~. californiensis.

NMML 408. Gonatus .§D.

Collected 28 May 1964, lat. 45°35'N,

long. 124°53.3'W. RV Commando, shrimp trawl, depth

750 fathoms. Parts saved: 1 entire specimen. Det.

K. Jefferts. Note: (Jeffert's comment) not ~.

pyros, no trace of optic photophore, clubs missing.

NMML 278. Gonatus sp. (type "A") (~. onyx? ~. californiensis.)

Collected 4 May 1970, lat. 46°08'N, long. 125°22'W.

From the stomach of ~. ursinus, specimen US70-225.

Parts saved: 3 upper, 3 lower beaks. (2 Aug. 1987 -

C. H. Fiscus)

Gonatus sp. (type "A") (~. califoriensis?)

Collected 14 May 1970, lat. 46°38'N, long. 124°51'W.

From the stomach of ~. ursinus, specimen US70-235.

Parts saved: 1 pair beaks. (2 Aug. 1987 - C. H.

Fiscus)
NMML 521a.

60

Family Gonatidae (Cont.)

NMML 280. Gonatus sp. (type "A") (~. onyx?)

Collected 15 May 1970, lat. 46°21'N, long. 124°53'W,

from the stomach of b. ursinus, specimen US70-237.

Parts saved: 4 pair beaks. (2 Aug. 1987 - C. H.

Fiscus)

NMML 283. Gonatus sp. (type "A") (1 beak and buccal mass ~.

pyros?, 1 buccal mass ~. ~)

Collected 24 May 1970, lat. 47°38'N, long. 128°19'W.

From the stomach of b. ursinus, specimen US70-325.

Parts saved: 3 pair beaks, 2 buccal masses, and

crown. (2 Aug. 1987 - C. H. Fiscus)

NMML 288. Gonatus sp. (type "A") ~. ~ plus one other

species

Collected 31 July 1973, lat. 56°01'N, long. 70012'W.

From the stomach of C. ursinus, specimen US73-179.

Parts saved: 9 pair beaks. (9 Aug. 1987 - C. H.

Fiscus)

NMML 521. Gonatus sp.

Collected 6 August 1990, lat. 47°19'N,

long. 125°23'W, by jig 60 m, V37H7-1. U.S.-Japan

Cooperative Survey. Frozen upon capture, thawed in

laboratory. Lab measurements. Male, DML 123 rom,

weight 33.0 g. Radula 5-row. Parts saved: entire

specimen.

Beaks and radula of 521. LRL 2.8 rom.
61

NMML 412.

NMML 411a.

NMML 413a.

NMML 412a.

NMML 413.

Family Gonatidae (Cont.).

NMML 411. Eogonatus tinro (see Okutani et al. 1988)

Collected 25 July 1984, lat. 58°4.3'N,

long. 179°33.9'W, by R. W. Mercer. MV Dominator,

cruise 82-1, haul 64, by midwater trawl. Depth

350 fathoms, bottom 1,970 fathoms. Time: 0045 hrs.

Female, DML 184 rom, LRL 5 rom. Parts saved: 1 entire

specimen. Det. K. Jefferts and T. Kubodera.

Beaks of 411.

Eogonatus tinro

Collected 25 July 1984, lat. 58°4.3'N,

long. 179°33.9'W, by R. W. Mercer. MV Dominator,

cruise 82-1, haul 64, by midwater trawl. Depth

350 fathoms, bottom 1,970 fathoms. Time: 0045 hrs.

Male, DML 147 rom, LRL 4.2 rom. Parts saved: 1 entire

specimen. Det. K. Jefferts and T. Kubodera.

Beaks and radula of 412.

Eogonatus tinro

Collected 1 August 1982, lat. 53°13.6'N,

long. 163°50.3'W, by R. W. Mercer. MV Dominator,

cruise 82-1, haul 75, by midwater trawl. Depth

372 fathoms, bottom 2,300 fathoms. Time: 0031 hrs.

Parts saved: 1 entire specimen. Det. K. Jefferts

and T. Kubodera.

Beaks and radula of 413.
62

Family Gonatidae (Cont.)

NMML 413b. Spermatophores of 413. 1 spermatophore length

13 mm, 1 - 11 mm, 1 - incomplete.

NMML 157. Gonatopsis borealis

Collected 7 February 1970, lat. 49°40'N,

long. 155°00'W. RV George ~. Kelez, cruise 48,

set 5, mesh size 2 inches. Parts saved: 3 entire

specimens, DMLs 182 mm, male; 186 mm, male; and

160 mm, male?

NMML 158. Gonatopsis borealis

Collected 28 January 1971, lat. 50010'N,

long. 160000'W. RV George ~. Kelez, cruise 51,

set 2, mesh size 2 inches. Parts saved: 4 entire

specimens, male, DMLs 145, 162, 140, and 142 mm.

NMML 159. Gonatopsis borealis

Collected 29 January 1971, lat. 49°30'N,

long. 160000'W. RV George ~. Kelez, cruise 51,

set 3, mesh size 2 inches. Parts saved: 2 entire

specimens.

NMML 160. Gonatopsis borealis

Collected 22 February 1970, lat. 48°25'N,

long. 163°25'W. RV Georqe ~. Kelez, cruise 48,

set 10. Parts saved: 2 entire specimens, DMLs 190

and 190 mm, both adult males.
63

Family Gonatidae (Cont.)

NMML 161. Gonatopsis borealis

Collected 23 February 1970, lat. 49°16'N,

long. 161°44'W. RV George ~. Kelez, cruise 48,

set 11, mesh size 2 1/2 inches. Parts saved:

1 entire specimen, DML 195 rom.

NMML 162. Gonatopsis borealis

Collected 5 February 1970, lat. 51°00'N,

long. 155°00'W. RV George ~. Kelez, cruise 48,

set 3, mesh size 2 inches. Parts saved: 1 entire

specimen, female.

NMML 163. Gonatopsis borealis

Collected 1 March 1970, lat. 53°30'N,

long. 160000'W. RV Georqe ~. Kelez, cruise 48,

set 16, mesh size 2 inches. Parts saved: 1 entire

specimen, DML approximately 140 rom.

NMML 164. Gonatopsis borealis

Collected 1 February 1971, lat. 50010'N,

long. 165°00'W. RV George ~. Kelez, cruise 51,

set 5. Female, DML 150 romapproximately. Parts

saved: 1 entire specimen.

NMML 165. GonatoDsis borealis

Collected 3 August 1972, lat. 51°00'N,

long. 177°20'E. RV George ~. Kelez, cruise 54,

set 15. Female, DML 145 romapproximately. Parts

saved: 1 entire specimen.
NMML 167.

NMML 168.

NMML 168a.

64

Family Gonatidae (Cont.).

NMML 166. Gonatopsis borealis

Collected 11 September 1971, lat. 51°31'N,

long. 179°21.5'E, by T. Newby. Depth 83 fathoms.

Time: 0455 hrs. Parts saved: 1 entire specimen,

DML 85 rom.

Gonatopsis borealis

Collected (no date or location). Parts saved:

1 entire specimen, DML 142 rom.

Gonatopsis borealis

Collected (no date), along 176°W long., probably

Pacific Salmon Investigations material. Female, DML

approximately 150 rom; male, DML approximately

130 rom. Parts saved: 2 entire specimens.

Gonatopsis borealis

Beaks of 168.

NMML 169.

NMML 169a.

Gonatopsis borealis

Collected 28 March 1962, lat. 49°03'N,

long. 154°54'W, by R. D. Carlson. MV Bertha Ann,

set 17. Water temperature 5.6°C. Parts saved: 1

entire specimen, internal organs missing, DML 200

rom.

Gonatopsis borealis

Beaks, radula, and gladius of 169.
65

Family Gonatidae (Cont.).

NMML 170. Gonatopsis borealis

Collected (no date), North Pacific. Parts saved:

1 head and arms, DML 157 rom.

NMML 170a. Gonatopsis borealis

Beaks of 170.

NMML 171.

NMML 172.

NMML 173.

NMML 173a.

NMML 174.

Gonatopsis borealis

Collected 4 February 1970, lat. 51°30'N,

long. 155°00'W. RV Georqe ~. Kelez, cruise 48,

set 2, mesh size 2 inches. Male, DML 145 rom. Parts

saved: 1 entire specimen.

Gonatopsis borealis

Collected 23 May 1969, lat. 50045'N, long. 165°00'W.

RV George ~. Kelez, cruise 46, set 18. Female,

DML 248 rom. Parts saved: 1 entire specimen. Beaks

in separate vial.

Gonatopsis borealis

Collected 8 May 1969, lat. 50000'N, long. 165°00'W.

RV George ~. Kelez, cruise 46, set 10. Parts saved:

2 entire specimens, females, DMLs 213 and 215 rom.

Gonatopsis borealis

Beaks of specimens 173.

Gonatopsis borealis

Collected 1 July 1966, lat. 50015'N, long. 176°22'W.

MV Paragon, cruise 38, set 6. Parts saved: 1 pair

beaks, gladius, and radula.
66

Family Gonatidae (Cont.).

NMML 175. Gonatopsis borealis

Collected (no date), North Pacific, probably from

Pacific Salmon Investigations. Specimen

photographed, DML 170 rom. Parts saved: 1 entire

specimen.

NMML 176. Gonatopsis borealis

Collected 11 July 1966, lat. 50030'N,

long. 176°22'W. MV Paraqon, cruise 38, set 12.

Parts saved: 1 entire specimen.

NMML 177. Gonatopsis borealis

Collected (no date) North Pacific. Parts saved:

4 gladii.

NMML 178. Gonatopsis borealis

Collected (no date) North Pacific. Parts saved:

1 pair beaks, DML 166 rom.

NMML 200. Gonatopsis borealis

Collected 1963, North Pacific. RV John N. Cobb

(probably off Columbia River, by R. Hitz. Parts

saved: 3 entire specimens, DMLs 115, 75, and 58 rom.

NMML 201. Gonatopsis borealis

Collected 1 July 1966, lat. 50015'N, long. 176°22'W,

by R. D. Carlson. MV Paragon, cruise 38, set 6.

Parts saved: 1 entire specimen, DML 115 rom.
NMML 206a.

67

Family Gonatidae (Cont.).

NMML 202. Gonatopsis borealis

Collected 11 July 1966, lat. 50030'N,

long. 176°22'W, by R. D. Carlson. MV Paragon,

cruise 38, set 12. Parts saved: beaks, radula, and

gladius only.

NMML 203. Gonatopsis borealis

Collected 26 October 1966, off central California.

From stomach of male £. macrocephalus, specimen Rice

1966-155. Parts saved: arms, buccal mass - beaks

dissected out. LRL 9.0 rom.

NMML 206. Gonatopsis borealis

Collected 1 December 1966, off central California.

From stomach of male £. macrocephalus, specimen Rice

1966-173. Parts saved: head, arms, buccal mass

only. Beaks and radula dissected out 8 May 1981.

F11 PlYmouth 1981.

Gonatopsis borealis

NMML 209.

NMML 209a.

Beaks and radula of 206. LRL 8.0 rom.

Gonatopsis borealis

Collected 5 September 1963, off central California.

From stomach of male £. macrocephalus, specimen Rice

932. Parts saved: 1 entire specimen (the 26.5 cm

gladius from the whole specimen). Lower beak

separated and 2 gladii: 27.5 cm, 26.5 cm. F2

PlYmouth 1981.

Gonatopsis borealis

Lower beak of 209. LRL 7.5 rom.
68

Family Gonatidae (Cont.).

NMML 249.

NMML 249a.

NMML 249b.

NMML 250.

NMML 250a.

NMML 250b.

Gonatopsis borealis4

Collected 21 October 1981, lat. 54°40'N,

long. 166°21'W. RV Miller Freeman, cruise 81-6,

haul 18, by trawl. Depth of trawl 100 fathoms,

bottom 180 fathoms. Trawl temperature 5.00C.

Frozen upon capture - thawed in laboratory. Male,

DML 124 rom, weight 82 g. Parts saved: 1 entire

specimen.

Beaks and radula of 249.

2 statoliths of 249.

Gonatopsis borealis

Collected 21 October 1981, lat. 54°40'N,

long. 166°21'W. RV Miller Freeman, cruise 81-6,

haul 18, by trawl. Depth of trawl 100 fathoms,

bottom 180 fathoms. Trawl temperature 5.00C.

Frozen upon capture - thawed in laboratory. Female,

DML 107 rom, weight 55 g. Parts saved: 1 entire

specimen.

Beaks and radula of 250.

2 statoliths of 250.

4Cruise 81-6, haul 18 contained 13 Berrvteuthis (one-mantle

only) and 7 Gonatopsis (one-mantle only). All Berryteuthis in

much worse shape than Gonatopsis, most chromatophores gone, color

white, tissue soft, many tentacles missing. Gonatopsis retained

skin and chromatophores, tissue hard in comparison to

Berryteuthis. Berryteuthis beaks and radulas extracted easily

during examination. Gonatopsis beaks were tight to surrounding

tissues and were extracted after a day of soaking.
NMML 300.

NMML 298a.

NMML 299.

69

Family Gonatidae (Cont.).

NMML 298. Gonatopsis borealis

Collected 26 July 1959, lat. 51°31'N,

long. 176°42'W. By surface trawl. Parts saved:

15 entire specimens. DMLs 29, 30, 37, 38, 44, 45,

45, 51, 52, 54, 57, 58, 60, 67, and 76 rom (measured

27 January 1983, some wear from length of time in

storage) .

1 gladius (60 rom) and 1 upper beak of 298.

Gonatopsis borealis

Collected 10 January 1961, lat. 38°08'N,

long. 123°35'W, by C. H. Fiscus. Night Light

Station 7-61. Water temperature 11°C. Parts saved:

1 entire specimen, DML approximately 30 rom (measured

25 January 1983) .

Gonatopsis borealis

Collected 17 January 1961, lat. 35°18'N,

long. 122°16'W, by C. H. Fiscus. Night Light

Station 8-61. Water temperature 13°C. Parts saved:

1 entire specimen, DML 40 rom (measured 25 January

1983) .
NMML 303.

NMML 301a.

NMML 301b.

NMML 301c.

NMML 302.

70

Family Gonatidae (Cont.).

NMML 301. Gonatopsis borealis

Collected 20 January 1961, lat. 34°50'N,

long. 121°40'W, by C. H. Fiscus. Night Light

Station 9-61. Water temperature 13.5°C. Parts

saved: 15 entire specimens, DMLs 31, 32, 35, 37, 38,

39, 39, 42, 42, 45, 45, 46, 50, 53, and 63 rom

(measured 27 January 1983, some wear from length of

time in storage) .

2 gladius of 301.

2 pair beaks, 7 rowed radulas from additional

specimens DMLs 41 and 71 rom of 301.

Tentacles of specimens listed in 301a and 301b.

Gonatopsis borealis

Collected 31 January 1961, lat. 33°32'N,

long. 120048'W, by C. H. Fiscus. Night Light

Station 12-61. Parts saved: 1 entire specimen, DML

approximately 60 rom (measured 23 January 1983) .

Gonatopsis borealis

Collected 20 August 1964, lat. 55°57'N,

long. 169°36'W, by C. H. Fiscus. Night Light

Station 7-64. Parts saved: 1 entire specimen, DML

approximately 38 rom (measured 25 January 1983) .

Det. T. Okutani.
71

NMML 306.

NMML 308.

NMML 307.

NMML 309.

Family Gonatidae (Cont.).

NMML 304. Gonatopsis borealis

Collected 30 May 1970, lat. 46°13'N, long. 125°22'W.

From the stomach of Q. ursinsus, specimen US70-352.

Parts saved: buccal masses from 3.

Gonato~sis borealis

Collected (no date), North Pacific. Parts saved:

gladius, DML 157 rom.

Gonatopsis borealis

Collected (no date), North Pacific. Parts saved:

gladius, DML 166 rom.

Gonatopsis borealis

Collected 12 December 1960, lat. 38°41'N,

long. 124°25'W, by C. H. Fiscus. Night Light

Station 4-61. Water temperature 12°C. Parts saved:

11 entire specimens, DMLs 55, 51, 50, 47, 47, 42,

41, 37, 37, 36, and 34 rom (measured 27 January 1983,

some wear due to length of time in storage) .

Gonatopsis borealis

Collected 6 January 1961, lat. 37°41'N,

long. 123°29'W, by C. H. Fiscus. Night Light

Station 5-61. Water temperature 11°C. Parts saved:

3 entire specimens, DMLs 50, 48, and 42 rom (measured

27 January 1983, some wear due to length of time in

storage) .
72

Family Gonatidae (Cont.).

NMML 310. Gonatopsis borealis

Collected 9 January 1961, lat. 38°42'N,

long. 123°51'W, by C. H. Fiscus. Night Light

Station 6-61. Water temperature 11°C. Parts saved:

15 entire specimens, DMLs 51, 47, 43, 42, 38, 38,

37, 35, 35, 34, 32, 31, 27, and 25 rom; 1

unmeasurable (measured 27 January 1983, some wear

due to length of time in storage) .

NMML 311. Gonatopsis borealis

Collected 22 January 1961, lat. 33°51'N,

long. 120051'W, by C. H. Fiscus. Night Light

Station 11-61. Water temperature 13.5°C. Parts

saved: 2 entire specimens, DMLs 42 and 45 rom (some

wear due to length of time in storage) .

NMML 312. Gonatopsis borealis

Collected 8 February 1961, lat. 35°19'N,

long. 121°43'W, by C. H. Fiscus. Night Light

Station 14-61. Water temperature 14°C. Parts

saved: 1 entire specimen, DML 57 (some wear due to

length of time in storage) .

NMML 313. Gonatopsis borealis

Collected 6 June 1965, lat. 37°38'N, long. 123°14'W.

From the stomach of ~. ursinus, specimen US65-248.

Parts saved: 1 gladius, arms, buccal mass, radula,

beaks.
NMML 333.

NMML 334.

73

Family Gonatidae (Cont.).

NMML 314. Gonatopsis borealis

Collected 20 May 1970, lat. 46°43'N, long. 127°38'W.

From the stomach of Q. ursinus, specimen US70-302.

Parts saved: 1 head, arms, buccal mass, radula,

beaks.

Gonatopsis borealis

Collected 12 May 1970, off central California. From

the stomach of female £. macrocephalus, specimen

Rice 1970-22. Parts saved: 1 relatively entire

specimen, DML 202 mm, LRL 8.0 mm, 3 lower beaks, LRL

8.8, 8.5, and 8.2 mm, 1 gladius fragment.

Gonatopsis borealis

Collected 24 May 1970, lat. 47°38'N, long. 128°19'W.

From the stomach of Q. ursinus, specimen US 70-325.

Parts saved: 1 entire specimen, plus 2 arms and

buccal masses - includes beaks. DMLs 138, 135, and

128 mm.

NMML 335. Gonatopsis borealis

Collected 10 June 1968, lat. 53°15'N,

long. 164°50'W. From the stomach of Q. ursinus,

specimen US68-820. Parts saved: 23 heads, arms,

buccal masses.
74

Family Gonatidae (Cont.).

NMML 372. Gonatopsis borealis

Collected 24 June 1982, lat. 47°55'N,

long. 177°39'E, by P. J. Gearin. By surface gill

net. Frozen upon capture, thawed and measured in

laboratory 20 July 1983. Female, DML 262 mm,

nidamental gland length - left 109 mm, right

114 mm - weight 678 g. Mature, eggs ripe. Parts

saved: beaks and radula.

NMML 372a. 1 statolith of 372.

NMML 391.

NMML 392.

Gonatopsis borealis

Collected 27 July 1982, lat. 53°19'N,

long. 176°43.4'W, by R. W. Mercer. MV Dominator,

cruise 82-1, haul 69, by midwater trawl. Depth

10 fathoms, bottom 2,000 fathoms. Time: 0230 hrs.

DML 39 mm, weight 3.1 g. Parts saved: 1 entire

specimen, beaks separate in same vial. Det. K.

Jefferts.

Gonatopsis borealis

Collected 27 June 1982, lat. 53°19'N,

long. 176° 43.4'W. by R. W. Mercer. MV Dominator,

cruise 82-1, haul 69, by midwater trawl. Depth

10 fathoms, bottom 2,000 fathoms. Time: 0230 hrs.

DML 41 mm, weight 3.9 g. Parts saved: 1 entire

specimen, beaks separate in same vial. Det. K.

Jefferts.
NMML 394.

75

Family Gonatidae (Cont.).

NMML 393. Gonatopsis borealis

Collected 27 July 1982, lat. 53°19'N,

long. 176°43.4'W, by R. W. Mercer. MV Dominator,

cruise 82-1, haul 69, by midwater trawl. Depth

10 fathoms, bottom 2,000 fathoms. Time: 0230 hrs.

DML 44 ern., weight 4.2 g. Parts saved: 1 entire

specimen, beaks separate in same vial. Det. K.

Jefferts.

Gonatopsis borealis

Collected 27 July 1982, lat. 53°19'N,

long. 176°43.4'W, by R. W. Mercer. MV Dominator,

cruise 82-1, haul 69 by midwater trawl. Depth

10 fathoms, bottom 2,000 fathoms. Time: 0230 hrs.

DML 45 em., weight 4.6 g. Parts saved: 1 entire

specimen, beaks separate in same vial. Det. K.

Jefferts.

NMML 395. Gonatopsis borealis

Collected 27 July 1982, lat. 53°19'N,

long. 176°43.4'W, by R. W. Mercer. MV Dominator,

cruise 82-1, haul 69, by midwater trawl. Depth

10 fathoms, bottom 2,000 fathoms. Time: 0230 hrs.

DML 50 rom, weight 7.7 g. Parts saved: 1 entire

specimen, beaks separate in same vial. Det. K.

Jefferts.
76

Family Gonatidae (Cont.).

NMML 396. Gonatopsis borealis

Collected 27 July 1982, lat. 53°19'N,

long. 176°43.4'W, by R. W. Mercer. MV Dominator,

cruise 82-1, haul 69, by midwater trawl. Depth

10 fathoms, bottom 2,000 fathoms. Time: 0230 hrs.

DML 51 rom, weight 7.5 g. Parts saved: 1 entire

specimen, beaks separate in same vial. Det. K.

Jefferts.

Gonatopsis borealis

Collected 9 June 1980, lat. 49°09.6'N,

long. 172°01.3'E, by W. A. Walker. Hoyo Maru 81,

set 4, by 55 rom mesh surface gill net. Frozen upon

capture, then placed in isopropyl alcohol. Female,

DML 170 rom. Parts saved: 1 entire specimen.

Measurements and dissection by C. H. Fiscus. Det.

W. A. Walker.

Beaks and radula of 397.

Statoliths of 397.

NMML 397.

NMML 397a.

NMML 397b.

NMML 398. Gonatopsis borealis

Collected 9 June 1980, lat. 49°09.6'N,

long. 172°01.3'E, by W. A. Walker. Hoyo Maru 81,

set 4, by 55 rom mesh surface gill net. Frozen upon

capture, then placed in isopropyl alcohol. Female,

DML 160 rom. Parts saved: beaks and radula.

Measurements and dissection by C. H. Fiscus, 1

January 1984. Det. W. A. Walker.
77

Family Gonatidae (Cont.).

NMML 510. Gonatopsis borealis

Collected 26 August 1990, lat. 45°59'N,

long. 126°17'W, by jig 90 m, V37 H27-1 U.S.-Japan

Cooperative Survey. Frozen upon capture, thawed in

laboratory. Lab measurements, male, DML 318 rom,

weight 860 g. Parts saved: entire specimen.

Beaks and radula of 510. LRL not measurable.

2 statoliths of 510.

NMML 510a.

NMML SlOb.

NMML 511.

NMML 511a.

NMML 511b.

NMML 512.

NMML 512a.

NMML 512b.

Gonatopsis borealis

Collected 14 August 1990, lat. 43°26'N,

long. 125°26'W, by jig 50 m, V68 H14-1 U.S.-Japan

Cooperative Survey. Frozen upon capture, thawed in

laboratory. Measurements, female, DML 284 rom,

weight 660 g. Nidamental gland length 57.0 rom.

Parts saved: entire specimen.

Beaks and radula of 511. LRL not measurable.

2 statoliths of 511.

Gonatopsis borealis

Collected 14 August 1990, lat. 43°26'N,

long. 125°26'W, by jig 50 m, V68 H14-2 U.S.-Japan

Cooperative Survey. Frozen upon capture, thawed in

laboratory. Lab measurements, male, DML 261 rom,

weight 500 g. Parts saved: entire specimen.

Beaks and radula of 512. LRL not measurable.

2 statoliths of 512.
NMML 524.

78

Family Gonatidae (Cont.).

NMML 523. Gonatopsis makko

Collected 26 March 1959, from stomach of Ziphius

cavirostris. Amchitka Island, Alaska, by K. Kenyon.

Parts saved: 1 lower beak damaged and worn, LRL

6.5 mm.

GonatoDsis makko

Collected 26 March 1959, from stomach of Ziphius

cavirostris, Amchitka Island, Alaska, by K. Kenyon.

Parts saved: 3 lower beaks, LRLs 8.5, 7.0, and

7.0 mm.

NMML 409.

NMML 189.

Gonatopsis - (Type A Kubodera)

Collected 28 November 1965, lat. 53°30'N,

long. 166°38'W, Unalaska Bay, Unalaska I., by C. H.

Fiscus. RV~.~. Reed, by shrimp trawl. Depth

60 fathoms. Parts saved: 1 entire specimen. Det.

K. Jefferts. Note: (Jeffert's comment, DML

approximately 29 mm, tentacles reduced, strong

chromatophore pattern on aboral surface.

Berryteuthis anonychus

Collected 1961, southwest of Columbia River. RV

John R. Cobb, Station 24, Atomic Energy Commission

Project. Parts saved: 14 small entire specimens.

Det. K. Jefferts.
79

Family Gonatidae (Cont.).

NMML 381. Berryteuthis anonychus

Collected 25 July 1982, lat. 58°04.3'N,

long. 179°33.9'W, by R. W. Mercer. MV Dominator,

cruise 82-1, haul 64, by midwater trawl. Depth

350 fathoms, bottom 1,970 fathoms. Time: 0045 hrs.

Female, DML 81 mrn, weight 19 g. Parts saved:

1 entire specimen. Det. K. Jefferts.

Beaks and radula of 381.

Berryteuthis anonychus

Collected 25 July 1982, lat. 58°04.3'N,

long. 179°33.9'W, by R. W. Mercer. MV Dominator,

cruise 82-1, haul 64, by midwater trawl. Depth

350 fathoms, bottom 1,970 fathoms. Time: 0045 hrs.

Female, DML 87 mrn, weight 29 g. Parts saved:

1 entire specimen. Det. K. Jefferts.

Beaks and radula of 382.

NMML 381a.

NMML 382.

NMML 382a.

NMML 383.

NMML 383a.

NMML 383b.

Berryteuthis anonychus

Collected 25 July 1982, lat. 58°04.3'N,

long. 179°33.9'W, by R. W. Mercer. MV Dominator,

cruise 82-1, haul 64, by midwater trawl. Depth

350 fathoms, bottom 1,970 fathoms. Time: 0045 hrs.

Female, DML 80 mrn, weight 16 g. Parts saved:

1 entire specimen. Det. K. Jefferts.

Beaks and radula of 383.

Statoliths of 383.
NMML 401.

80

Family Gonatidae (Cont.).

NMML 399. Berryteuthis anonvchus

Collected 15 June 1980, lat. 51°34.1'N,

long. 173°35.5'E, by W. A. Walker. From stomach of

Oncorhynchus nerka. Hoyo Maru 81, set 9, by surface

gill net. Initially in 10% formalin, transferred to

isopropyl alcohol. Female, DML 93 rom, weight

27.7 g. Parts saved: 1 entire specimen.

Measurements in laboratory 1 January 1984 by C. H.

Fiscus. Det. W. A. Walker.

NMML 400. Berryteuthis anonychus

Collected 15 June 1980, lat. 54°34.1'N,

long. 173°35.5'E, by W. A. Walker. From stomach of

O. nerka. Hoyo Maru 81, set 9, by surface gill net.

Initially in 10% formalin, transferred to isopropyl

alcohol. Female, DML 183 rom, weight 25 g. Parts

saved: 1 entire specimen. Measurements in

laboratory 1 January 1984 by C. H. Fiscus. Det. W.

A. Walker.

Berryteuthis anonychus

Collected 15 June 1980, lat. 51°34.1'N,

long. 173°35.5'E, by W. A. Walker. From the stomach

of Q. nerka. Hoyo Maru 81, set 9, by surface gill

net. Frozen on capture, transferred to isopropyl

alcohol. Adult female, DML 92 rom, specimen greatly

flattened. Parts saved: 1 entire specimen.

Measurements and dissection by C. H. Fiscus, January

1984. Det. W. A. Walker.
81

Family Gonatidae (Cont.).

NMML 401a. Beaks and radula of 401.

NMML 401b. Statoliths of 401.

NMML 402. Berryteuthis anonychus

Collected 15 June 1980, lat. 51°34.1'N,

long. 173°35.5'E, by W. A. Walker. From the stomach

of Q. nerka. Hoyo Maru 81, set 9, by surface gill

net. Frozen on capture, transferred to isopropyl

alcohol. Adult male, spermatophores present. DML

75 mm+ (distorted), specimen greatly flattened.

Parts saved: 1 entire specimen. Measurements and

dissection by C. H. Fiscus, January 1984. Det. W.

A. Walker.

Beaks and radula of 402.

Statoliths of 402.

NMML 402a.

NMML 402b.

NMML 403.

NMML 403a.

NMML 403b.

Berryteuthis anonychus

Collected 15 June 1980, lat. 51°34.1'N,

long. 173°35.5'E, by W. A. Walker. From the stomach

of Q. nerka. Hoyo Maru 81, set 9, by surface gill

net. Frozen on capture, transferred to isopropyl

alcohol. Adult male, spermatophores present. DML

70 mm, specimen greatly flattened. Parts saved:

1 entire specimen. Measurements and dissection by

C. H. Fiscus, January 1984. Det. W. A. Walker.

Beaks and radula of 403.

Statoliths of 403.
82

Family Gonatidae (Cont.).

NMML 404. Berryteuthis anonychus

Collected 6 August 1963, lat. 54°19'N,

long. 166°29'W. From the stomach of Q. ursinus,

specimen US63-411. DML 15 mm. Parts saved:

1 entire specimen. Det. K. Jefferts. Note:

(Jeffert's comment) body shape as in ~. anonychus,

club tip with only minute sucker buds. No enlarged

suckers on club, no arm hooks.

Berryteuthis magister

Collected August 1971, North Pacific.

Kelez, cruise 53. Male, DML 152 mm.

entire specimen.

Berryteuthis magister

Collected 1963, probably North Pacific. RV John N.

Cobb. Parts saved: 4 entire specimens DMLs 108,

92, 132, and 140 mm, plus buccal mass of much larger

specimen.

Berryteuthis magister

Collected 10 April 1963, southwest of Columbia

River, by W. Pererya. RV John N. Cobb, in

325 fathoms. DML approximately 240 mm. Parts

saved: 1 entire specimen, beaks intact.

2 entire specimens, DMLs 220 and 200 mm, of 193

haul) .

1 buccal mass of 193.

NMML 182.

NMML 192.

NMML 193.

NMML 193a.

NMML 193b.

RV George ~.

Parts saved: 1
NMML 195.

83

Family Gonatidae (Cont.).

NMML 194. Berryteuthis maqister

Collected 26 June 1973, lat. 55°00'N,

long. 167°31'W, by C. H. Fiscus. MV Mark~, cruise

73-36, station A-1, haul 1. Depth 160 fathoms.

29 specimens, 5 of which were sent to PlYmouth, May

1981. DMLs 130, 153, 162, 154, 81, 170, 84, 150,

97, 80, 76, 79, 75, 90, 102, 95, 70, 67, 82, 66, 67,

92, 68, 72, 75, 68, 65, 64, and 55 rom. Not sexed.

Parts saved: 24 entire specimens.

Berryteuthis magister

Collected 26 June 1973, lat. 55°20'N,

long. 168°05'W, by C. H. Fiscus. MV Mark~, cruise

73-36, station B-18, haul 2. 13 specimens, listed

below. Parts saved: buccal masses of 195 a-g.

(rom) Parts saved* Weight(g)Sex DML

1,525Buccal mass F 363

- 153 110 Buccal mass

285F 213235 193

Buccal mass F20 85

Buccal mass 300F 199

- 65 15 Buccal mass

120F

180-300165-35-15-

the field after collection.
84

Family Gonatidae (Cont.).

NMML 198. Berrvteuthis magister

Collected (no date), North Pacific. Parts saved:

several gladii.

NMML199. Berryteuthis magister

Collected 14 February 1962, lat. 55°05.5'N,

long. 167°40.5'W, by H. Kajimura. DML 154 rom.

Parts saved: beaks, gladius and radula only.

NMML 204. Berryteuthis magister

Collected 29 August 1973, lat. 56°15'N,

long. 168°35'W. From stomach of Q. ursinus,

specimen US73-433. Parts saved: 1 entire specimen,

DML 96 rom. Buccal mass and beaks dissected out

(good beaks) .

NMML 243. Berryteuthis maqister

Collected 4 November 1981, lat. 53°47.5'N,

long. 164°15.2'W. RV Miller Freeman cruise 80-6,

haul 41, by trawl. Depth of trawl on bottom,

51 fathoms. Surface temperature 8°C, trawl

temperature 7.6°C. Frozen fresh. Female, DML 198

mm, weight 224 g. Parts saved: beaks and radula.

NMML 243a. Statoliths of 243.
85

Family Gonatidae (Cont.).

NMML 244. Berryteuthis magister

Collected 2 November 1981, lat. 55°20.7'N,

long. 167°47.2'W. RV Miller Freemam, cruise 81-6,

haul 37, by trawl. Depth of trawl on bottom,

96 fathoms. Surface temperature 7.3°C, trawl

temperature 5.5°C. Frozen fresh. Female, DML

290 rom, nidamental gland length 40 rom, weight 803 g,

ovary weight 145 g, egg diameter 3-4 rom. Parts

saved: beaks and radula.

NMML 244a. Statoliths of 244.

NMML 244b.

NMML 247.

NMML 247a.

Section of tissue, etc.

Berryteuthis magister

Collected 20 October 1981, lat. 54°14'N,

long. 166°02'W. RV Miller Freeman, cruise 81-6,

haul 17, by trawl. Depth 59 fathoms. Trawl

temperature 4.4°C. Frozen upon capture, thawed in

laboratory. Male, DML 185 rom, weight 216 g, right

tentacle 3/4 missing, left tentacle 265 rom. Parts

saved: 1 entire specimen.

Beaks and radula of 247.
86

Family Gonatidae (Cont.).

NMML 248. Berryteuthis magister

Collected 20 October 1981, lat. 54°22'N,

long. 165°45'W. RV Miller Freeman, cruise 81-6,

haul 15, by trawl. Depth 64 fathoms. Trawl

temperature 5.0oC. Frozen upon capture, thawed in

laboratory. Mature female, DML 273 rom, weight

643 g, longest tentacle 385 rom, nidamental gland

length - left, 123 rom, right, 126 rom, egg diameter

2.5 - 3 rom. Parts saved: 1 entire specimen.

NMML 248a. Beaks and radula of 248.

NMML 251.

NMML 251a.

NMML 251b.

Berryteuthis magister

Collected 20 October 1981, lat. 54°22'N,

long. 165°45'W. RV Miller Freeman, cruise 81-6,

haul 15, by trawl. Depth 64 fathoms. Trawl

temperature 5.0oC. Frozen upon capture, thawed in

laboratory. Female, DML 273 rom, weight 687 g,

longest tentacle 365 rom, approximately equal,

nidamental gland length 112 rom, approximately equal.

Egg diameter, 3 rom. Parts saved: 1 entire specimen.

Beaks and radula of 251.

Statoliths of 251.
NMML 253.

NMML 253a.

NMML 253b.

87

Family Gonatidae (Cont.).

NMML 252. Berryteuthis magister

Collected 20 October 1981, lat. 54°14'N,

long. 166°02'W. RV Miller Freeman, cruise 81-6,

haul 17, by trawl. Depth 59 fathoms. Trawl

temperature 4.4°C. Frozen upon capture, thawed in

laboratory. Female, DML 252 rnm, weight 569 g (fresh

weight 601 g), nidamental gland length, 82 rnm, egg

diameter, 1 - 1.5 rnm. Parts saved: 1 entire

specimen - beaks extracted and in same container.

Berryteuthis magister

Collected 20 October 1981, lat. 54°14'N,

long. 166°02'W. RV Miller Freeman, cruise 81-6,

haul 17, by trawl. Depth 59 fathoms. Trawl

temperature 4.4°C. Frozen upon capture, thawed in

laboratory. Female, DML 285 rnm, weight 830 g (fresh

weight 927 g), tentacles approximately same, 430 rnm,

nidamental gland length 62 rnm, approximately same

size. Eggs - none apparent. Ovary - very small

(spent?). Parts saved: 1 entire specimen.

Beaks and radula of 253.

Statoliths of 253.
NMML 254a.

NMML 255.

88

Family Gonatidae (Cont.).

NMML 254. Berryteuthis maqister

Collected 21 October 1981, lat. 54°40'N,

long. 166°21'W. RV Miller Freeman, cruise 81-6,

haul 18, by trawl. Depth of trawl 100 fathoms,

bottom 180 fathoms. Trawl temperature 5.0oC.

Frozen upon capture, thawed in laboratory. Male,

DML 95 mm, weight 33 g. Parts saved: beaks and

radula.

2 statoliths of 254.

Berryteuthis magister

Collected 21 October 1981, lat. 54°40'N,

long. 166°21'W. RV Miller Freeman, cruise 81-6,

haul 18, by trawl. Depth of trawl 100 fathoms,

bottom 180 fathoms. Trawl temperature 5.0oC.

Frozen upon capture, thawed in laboratory. Male,

DML 170 mm, weight 172 g. Parts saved: beaks and

NMML 255a.

NMML 256.

NMML 256a.

radula.

2 statoliths of 255.

Berryteuthis maqister

Collected 21 October 1981, lat. 54°40'N,

long. 166°21'W. RV Miller Freeman, cruise 81-6,

haul 18, by trawl. Depth of trawl 100 fathoms,

bottom 180 fathoms. Trawl temperature 5.0oC.

Frozen upon capture, thawed in laboratory. Male,

DML 63 mm, weight 15 g. Parts saved: beaks and

radula.

1 statolith from 256.
89

Family Gonatidae (Cont.).

NMML 257. Berryteuthis magister

Collected 21 October 1981, lat. 54°40'N,

long. 166°21'W. RV Miller Freeman, cruise 81-6,

haul 18, by trawl. Depth of trawl 100 fathoms,

bottom 180 fathoms. Trawl temperature 5.0oC.

Frozen upon capture, thawed in laboratory. Male,

DML 80 rom, weight 21 g. Parts saved: beaks and

radula.

NMML 257a. 2 statoliths from 257.

NMML 258.

NMML 258a.

NMML 455.

Berryteuthis maqister

Collected 21 October 1981, lat. 54°40'N,

long. 166°21'W. RV Miller Freeman, cruise 81-6,

haul 18, by trawl. Depth of trawl 100 fathoms,

bottom 180 fathoms. Trawl temperature 5.0oC.

Frozen upon capture, thawed in laboratory. Male,

DML 147 rom, weight 109 g. Parts saved: beaks and

radula.

1 statolith from 258.

Berryteuthis magister

Collected 15 July 1984, lat. 55°05'N,

long. 166°40'W. By special deep haul, depth

188 fathoms. By R. W. Mercer. Sex unknown, DML 250

rom, weight 439 g. Parts saved: statoliths. Det. R.

W. Mercer.
NMML 453.

NMML 472.

90

Family Gonatidae (Cont.).

NMML 452. Berryteuthis maqister

Collected 24 March 1986, lat. 54°34.3'N,

long. 167°32.3'W. By bottom trawl. Depth 350 to

400 fathoms. Time: 2100 hrs. Male, DML 263, weight

645.8 g. Reproductive development mature --

spermatophores present. Parts saved: statoliths.

Det. W. Walker and E. Hacker.

Berryteuthis maqister

Collected 24 March 1986, lat. 54°34.3'N,

long. 167°32.3'W. By bottom trawl. Depth 350 to

400 fathoms. Time: 2100 hrs. Male, DML 223, weight

378.8 g. Reproductive development mature --

spermatophores present. Parts saved: statoliths.

Det. W. Walker and E. Hacker.

Berryteuthis magister

Collected 7 February 1990, in Port Susan, puget

Sound, Washington, lat. 48°08.1'N, long. 122°23.9'W.

By R. L. DeLong. NOAA ship Miller Freeman. By

bottom trawl. Depth 64 fathoms (117 m). Time not

reported. Female, DML 238 mrn, weight 665 g.

Reproductive development mature -- egg diameter

3-4 mrn. Nidamental gland length 100 mrn.

approximately equal. Left tentacle length 230 mrn.

Right tentacle broken off near base. Fin length

120 mrn. Arm lengths, 1 - 100, 2 - 150, 3 - 115, and

4 - 120 mrn. Parts saved: entire specimen. Det. C.

H. Fiscus.
91

Family Gonatidae (Cont.).

NMML 472a. Beaks and radula of 472. LRL 6.7 rom.

NMML 472b. 2 statoliths
92

Order Teuthida

Suborder Oegopsida

Family Architeuthidae

NMML 110.

NMML 111.

NMML 112.

NMML 113.

NMML 114.

Architeuthis japonicas

Collected 16 September 1961, off central California.

From stomach of male £. macrocephalus, specimen Rice

535. Parts saved: 1 upper beak.

Architeuthis japonica

Collected 15 June 1963, off central California.

From stomach of male £. macrocephalus, specimen Rice

813. Parts saved: 1 upper beak.

Architeuthis japonica

Collected 25 November 1966, off central California.

From stomach of male £. macroceDhalus, specimen Rice

1966-165. Parts saved: 1 upper beak.

Architeuthis iaponica

Collected 1 December 1966, off central California.

From stomach of female g. macrocephalus, specimen

Rice 1966-177. Parts saved: 1 lower beak fragment.

Architeuthis japonica

Collected 9 December 1966, off central California.

From stomach of male g. macrocephalus, specimen Rice

1966-185. Parts saved: 1 upper beak.

SNote: (F. G. Hochberg, Santa Barbara Natural History

Museum, Santa Barbara, CA 93105) pers. commun. 24 Aug. 1987

Architeuthis from around the Pacific Rim - California to Japan

can be attributed to the species A. japonica.
93

Family Architeuthidae (Cont.).

NMML 115. Architeuthis japonica

Collected 12 May 1967, off central California. From

stomach of female R. macrocephalus, specimen Rice

1967-128. Photographed; see illustration in Fiscus

and Rice (1974). Parts saved: 1 pair beaks, lower

beak-patch on wing stage. LRL 11.0 rom.

NMML 116. Architeuthis japonica

Collected 23 April 1968, off central California.

From stomach of female R. macrocephalus, specimen

Rice 1968-108. Parts saved: 1 upper beak.

NMML 117. Architeuthis japonica

Collected 23 April 1968, off central California.

From stomach of male P. macrocephalus, specimen Rice

1968-109. Parts saved: 1 upper beak fragment.

NMML 118. Architeuthis japonica

Collected 13 June 1969, off central California.

From stomach of female R. macrocephalus, specimen

Rice 1969-143. Parts saved: 1 upper beak.

NMML 119. Architeuthis japonica

Collected 19 November 1969, off central California.

From stomach of male R. macrocephalus, specimen Rice

1969-201. Parts saved: 1 lower beak fragment.

LRL 12.0 rom.
94

Family Architeuthidae (Cont.).

NMML 120. Architeuthis japonica

Collected 24 February 1970, off Central California.

From stomach of male g. macrocephalus, specimen Rice

1970-001. Parts saved: 1 lower beak fragment.

(Note: next to 115, this is best lower beak in

collection). LRL 12.5 rom.

NMML 486. Architeuthis japonica

Collected 19 July 1990, West Cove, San Miguel

Island, California. From stomach of male Mirounga

angustirostris by lavage, specimen Y1991/1992.

Parts saved: 1 lower beak, patch on wing stage,

LRL 10.3 rom.
95

Order Teuthida

Suborder Oegopsida

Family Histioteuthidae

NMML 59.

NMML 59a.

NMML 60.

NMML 60a.

NMML 61.

NMML 61a.

NMML 62.

Histioteuthis dofleini

Collected 1 November 1967, off central California.

From stomach of female R. macrocephalus, specimen

Rice 1967-234. Parts saved: body, head, arms,

tentacles, buccal mass. Det. C. F. E. Roper, U.S.

National Museum. Additional information and photos

in file.

Radula and beaks of 59, LRL 5.0 rom.

Histioteuthis dofleini

Collected 23 April 1968, off central California.

From stomach of male R. macrocephalus, specimen Rice

1968-109. Parts saved: arms, tentacles.

Beaks of 60, LRL 6.0 rom. F6 PlYmouth 1981 (verified

by M. R. Clarks).

Histioteuthis dofleini

Collected 27 November 1966, off central California.

From stomach of female R. macrocephalus, specimen

Rice 1966-170. Parts saved: head, arms, tentacles.

Beaks of 61. LRL 5.0 rom.

Histioteuthis dofleini

Collected 27 November 1966, off central California.

From stomach of female R. macrocephalus, specimen

Rice 1966-169. Parts saved: 1 pair beaks. LRL

6.0 rom.
NMML 443.

NMML 479.

96

Family Histioteuthidae (Cont.).

NMML 63. Histioteuthis dofleini (identified 9 Aug. 1987 -

C. H. Fiscus) Collected 14 May 1969, off central

California. From stomach of male R. macrocephalus,

specimen Rice 1969-120. Parts saved: 3 buccal

masses. LRLs 5.0, 5.2, and 6.0 rom.

NMML 433. Histioteuthis dofleini

Collected 18 August 1984, Guadalupe Island, Mexico,

by R. L. DeLong, from the scat of A. townsendi,

specimen At-84-34a. Parts saved: 1 pair? beaks.

NMML 442. Histioteuthis dofleini

Collected 30 April 1984, San Miguel Island,

California, by G. A. Antonelis, by lavage from

stomach of M. angustirostris, specimen A-43. Parts

saved: 2 lower beaks. Det. F. G. Hochberg and C. H.

Fiscus.

Histioteuthis dofleini

Collected 14 June 1966, off central California.

From stomach of male R. macrocephalus, specimen Rice

1966-51. Parts saved: 1 lower beak. LRL 4.2 rom.

Histioteuthis heteropsis

Collected 30 April 1984, San Miguel Island,

California, by G. A. Antonelis, by lavage from

stomach of M. angustirostris, specimen A-43. Parts

saved: 5 lower beaks. Det. F. G. Hochberg and C. H.

Fiscus.
97

Family Histioteuthidae (Cont.).

NMML 480. Histioteuthis heteropsis

Collected 19 May 1968, off central California. From

stomach of male ~. macrocephalus, specimen Rice

1968-226. Parts saved: 1 lower beak. LRL 3.0 rom.
98

Order Teuthida

Suborder Oegopsida

Family Ornmastrephidae

Subfamily Illicinae

NMML 108.

NMML 522.

NMML 522a.

Illex illecebrosus

Collected spring 1970, off Newfoundland, Canada, by

M. C. Mercer. Parts saved: 16 pair beaks (in

separate vials, each containing 1 pair of beaks) .

DML and sex is listed. Det. M. C. Mercer.

Illex argentinus

Collected summer 1990, from Japanese squid fishery

vicinity of Falkland Islands, South Atlantic Ocean,

by jig, Wakashio Maru No. 68. Obtained by observer

B. Bradley. Frozen upon capture, thawed in

laboratory. Lab measurements: female, DML 350 rom,

weight 1,070 g. Nidamental gland length 96.5 rom.

Parts saved: entire specimen.

Beaks and radula of 522. LRL 5.5 rom.

NMML 522b. 2 statoliths of 522.

NMML 522 c. Club end of a tentacle - from same package as 522

but not from 522.
99

Family Ommastrephidae

Subfamily Todarodinae

NMML 7S.

NMML 79.

NMML 79a.

NMML SO.

NMML SOa.

NMML SOb.

Todarodes pacificus (Ommastrephes sloani pacificus)

Collected May 1950, Tokyo Fish Market, by F. Wilke.

Det. F. Wilke. Loaned to C. F. E. Roper, curator,

u. S. National Museum, 25 July 1967 (see notes and

reprint section).

Todarodes pacificus (Q. ~. pacificus)

Collected April 1950, off Onagawa, Japan, by F.

Wilke. From the stomach of ~. ursinus 247. Det. F.

Wilke. Loaned to C. F. E. Roper, U. S. National

Museum, 25 July 1967 (see notes and reprint

section). In Nat'l. Museum Collection

Todarodes pacificus (Q. ~. pacificus)

Collected April 1950, off Onagawa, Japan, by F.

Wilke. From the stomach of ~. ursinus 247. Parts

saved: buccal mass and beaks only. Det. F. Wilke.

Todarodes pacificus

Collected S November 1965, Seattle Public Market.

Probably from Japan. Parts saved: 2 entire

specimens, DMLs 222 and 232 mrn, gladii lengths

213 and 225 mrn.

2 pair beaks of SO.

Suckers of SO.
100

Family Ommastrephidae (Cont.).

Subfamily Todarodinae

NMML 428.

NMML 429.

NMML 430.

Nototodarus gouldi6

Collected 1 August 1983, lat. 40055.5'S,

long. 173°24'E, 1138 hrs, by R. H. Matlin, 1 Jan.

1983 (1.1.0 no. 18), by trawl, depth 45 m. Female,

DML 310 rom, weight 930 g. Parts saved: beaks. Det.

by R. H. Mattlin, New Zealand Fish. Res. Div. Note:

see footnote this page.

Nototodarus gouldi

Collected 1 August 1983, lat. 40055.5'S,

long. 173°24'E, 1138 hrs, by R. H. Mattlin, 1 Jan.

1983 (1.1.0 no. 32), by trawl, depth 45 m. Male,

DML 220 rom, weight 405 g. Parts saved: beaks. Det.

R. H. Mattlin, New Zealand Fish. Res. Div. Note:

see footnote this page.

Nototodarus sloani

Collected 1 November 1983, lat. 44°40'S,

long. 172°17.8'E, 1620 hrs, by R. H. Mattlin,

14 Jan. 1983 (4.5.0 no. 4), by trawl, depth 53 m.

Male, DML 303 rom, weight 720 g. Parts saved: beaks.

Det. by R. H. Mattlin, New Zealand Fish. Res. Div.

Note: see footnote this page.

~ote: Nototodarus gouldi is found north of subtropical

convergence. Nototodarus sloani is found south of subtropical

convergence. See Smith, et al. 1987 for additional information.
101

Family Ommastrephidae (Cont.).

Subfamily Todarodinae

NMML 431. Nototodarus sloani

Collected 1 December 1983, lat. 46°01.2'S,

long. 170029.5'E, 0713 hrs, by R. H. Mattlin,

15 Jan. 1983 (5.1.0 no. 4), by trawl, depth 50 m.

Male, DML 210 rom, weight 200 g. Parts saved: beaks.

Det. by R. H. Mattlin, New Zealand Fish. Res. Div.

Note: see footnote 6.
102

Family Ommastrephidae

Subfamily Ommastrephinae

NMML 81.

NMML 82.

NMML 84.

NMML 85.

NMML 86.

NMML 86a.

NNML 210.

Ommastrephes bartrami

Collected 1957, south of Aleutian Islands, by R. A.

Hajny. Can 3. Parts saved: 1 entire specimen.

Ommastrephes bartrami

Collected 1957, south of Aleutian Islands, by R. A.

Hajny. Parts saved: 2 entire specimens.

Ommastrephes bartrami

Collected 1957, south of Aleutian Islands, by R. A.

Hajny. Parts saved: 1 tentacle and 1 arm.

Ommastrephes bartrami

1 pair beaks, radula, pen of 84.

Ommastrephes bartrami

Collected 13 October 1965, lat. 44°40'N,

long. 178°02'E, by R. D. Carlson. RV George ~.

Kelez, cruise 36, set 10. Parts saved: 1 head,

arms, tentacles.

1 pair beaks and radula of 86.

Ommastrephes bartrami

Collected 2 July 1981, over Grays Canyon

approximately 40 miles west of Westport, Washington,

by M. Bucy. FV Tres Cher, by dip net-nightlight.

Female, DML 404 rom, weight 2.051 kg. Identified

from whole specimen. Parts saved: beaks and radula.

LRL 9.5 rom.
103

Family Ommastrephidae

Subfamily Ommastrephinae (Cont.).

NMML 365.

NMML 365a.

NMML 365b.

NMML 366.

NMML 366a.

NMML 366b.

Ommastrephes bartrami

Collected 14 June 1981, lat. 41°N, long. 180°, by C.

Bouchet. RV Oshoro Maru research cruise, set 3, by

surface gill net. Frozen upon capture, thawed in

laboratory. Laboratory measurements 14 July 1983,

Female, DML 368 rom, nidamental gland length 49 rom,

weight 1,905 g (1 fin missing). Parts saved:

1 entire specimen.

Beaks and radula of 365.

2 statoliths of 365.

Oromastrephes bartrami

Collected 14 June 1981, lat. 41°N, long. 180°, by

C. Bouchet. RV Oshoro Maru research cruise, set 3,

by surface gill net. Frozen upon capture, thawed in

laboratory. Laboratory measurements 14 July 1983.

Female, DML 392 rom, nidamental gland length 54 rom,

weight 1,976 g. Parts saved: 1 entire specimen.

Beaks and radula of 366. LRL 10.0 rom.

2 statoliths of 366.
104

Family Ommastrephidae

Subfamily Ommastrephinae (Cont.).

NMML 367.

NMML 367a.

NMML 494.

NMML 494a.

NMML 494b.

NMML 495.

NMML 495a.

NMML 495b.

Ommastrephes bartrami

Collected 14 June 1981, lat. 41°N, long. 180°. by

C. Bouchet. RV Oshoro Maru research cruise, set 3,

by surface gill net. Frozen upon capture, thawed in

laboratory. Laboratory measurements 14 July 1983.

Female, DML 382 rom, nidamental gland length 44 rom,

weight 1,674 g, 1 tentacle missing. Parts saved:

beaks and radula.

2 statoliths of 367.

Ommastrephes bartrami

Collected 27 August 1990, lat. 45°10'N,

long. 126°43'W, by jig 80 m, V37H28-2, U.S.-Japan

Cooperative Survey. Frozen upon capture, thawed in

laboratory. Lab measurements, male, DML 300 rom,

weight 810 g. Parts saved: entire specimen.

Beaks and radula of 494. LRL 6.8 rom.

2 statoliths of 494.

Ommastrephes bartrami

Collected 13 August 1990, lat. 42°12.5'N,

long. 124°54'W, by jig 100 m, V63H13-3, U.S.-Japan

Cooperative Survey. Frozen upon capture, thawed in

laboratory. Lab measurements, female, DML 345 rom,

weight 1,100 g. Parts saved: entire specimen.

Beaks and radula of 495. LRL 7.1 rom.

2 statoliths of 495.
105

Family Ommastrephidae

Subfamily Ommastrephinae (Cont.).

NMML 496.

NMML 496a.

NMML 496b.

NMML 497.

NMML 497a.

NMML 497b.

NMML 498.

NMML 498a.

NMML 498b.

Ommastrephes bartrami

Collected 26 August 1990, lat. 46°18'N,

long. 128°18'W, by jig 120 m, V58H26-3, U.S.-Japan

Cooperative Survey. Frozen upon capture, thawed in

laboratory. Lab measurements, female, DML 298 rom,

weight 740 g. Parts saved: entire specimen.

Beaks and radula of 496. LRL 7.0 rom.

2 statoliths of 496.

Ommastrephes bartrami

Collected 26 August 1990, lat. 46°35'N,

long. 128°29'W, by jig 100 m, V63H26-4, U.S.-Japan

Cooperative Survey. Frozen upon capture, thawed in

laboratory. Lab measurements, male, DML 348 rom,

weight 1,310 g. Parts saved: entire specimen.

Beaks and radula of 497. LRL not measurable.

2 statoliths of 497.

Oromastre~hes bartrami

Collected 22 August 1990, lat. 45°15'N,

long. 126°44'W, by jig 120 m, V58H22-1, U.S.-Japan

Cooperative Survey. Frozen upon capture, thawed in

laboratory. Lab measurements, female, DML 488 rom,

weight 3,360 g. Parts saved: entire specimen.

Beaks and radula of 498. LRL 11.0 rom.

2 statoliths of 498.
106

Family Ommastrephidae

Subfamily Ommastrephinae (Cont.).

NMML 499.

NMML 499a.

NMML 499b.

NMML 500.

NMML sOOa.

NMML sOOb.

NMML 501.

Ornmastreohes bartrami

Collected 25 August 1990, lat. 47°34'N,

long. 126°38'W, by jig 90 m, V37H26-3, U.S.-Japan

Cooperative Survey. Frozen upon capture, thawed in

laboratory. Lab measurements, female, DML 480 mm,

weight 3,030 g. Parts saved: entire specimen.

Beaks and radula of 499. LRL 9.5 mm.

2 statoliths of 499.

Ommastrephes bartrami

Collected 25 August 1990, lat. 47°34'N,

long. 126°38'W, by jig 90 m, V37H26-s, U.S.-Japan

Cooperative Survey. Frozen upon capture, thawed in

laboratory. Lab measurements, male, DML 342 mm,

weight 1,200 g. Parts saved: entire specimen.

Beaks and radula of 500. LRL 7.2 mm.

2 statoliths of 500.

Ornmastrephes bartrami

U.S-Japan Cooperative Survey, 1990. Spermiduct and

related organs - various stages of development,

spermatophores.

a. V37H23-3. Collected 22 August 1990,

lat. 44°2s'N, long. 126°44'W. Male, DML 373 mm,

weight 1,520 g, gonad weight 26 g, spermiduct weight

43.7 g.
107

Family Ommastrephidae

Subfamily Ommastrephinae (Cont.).

b. V37H29-3. Collected 28 August 1990,

lat. 44°45'N, long. 127°20'W. Male, DML 310 rom,

weight 780 g, gonad weight 11.94 g.

c. V63H30-3. Collected 30 August 1990,

lat. 44°29'N, long. 126°49'W. Male, DML 390 rom,

weight 1,800 g, gonad weight 33.31 g.

d. V63H32-1. Collected 1 September 1990,

lat. 44°22'N, long. 126°28'W. Male, DML 325 rom,

weight 1,260 g, gonad weight 20.31 g.

NMML 87.

NMML 88.

Dosidicus giQas7

Collected 21-22 January 1972, lat. 9°N,

long. 86°20'W, by G. M. Treinen. Specimen DH-1.

DML 205 rom, weight 250 g. Parts saved: buccal mass

only.

Dosidicus gigas

Collected 21-22 January 1972, lat. 9°N,

long. 86°20'W, by G. M. Treinen. Specimen DH-2.

Male, DML 188 rom, weight 168 g. Parts saved: buccal

mass only.

7Determination and measurements by C. H. Fiscus and D. R.

Holts. See NMML reference file, Table 1, for complete

information on catalogue numbers 87-95.
108

Family Ommastrephidae

Subfamily Ommastrephinae (Cont.).

NMML 89.

NMML 90.

NMML 91.

NMML 92.

NMML 93.

Dosidicus gigas

Collected 21-22 January 1972, lat. 9°N,

long. 86°20'W, by G. M. Treinen. Specimen DH-3.

Male, DML 171 rom, weight 153 g. Parts saved: buccal

mass only.

Dosidicus giqas

Collected 20-22 January 1972, lat. 9°N,

long. 86°20'W, by G. M. Treinen. Specimen DH-4.

Male, DML 167 rom, weight 127 g. Parts saved: buccal

mass only.

Dosidicus gigas

Collected 21-22 January 1972, lat. 9°N,

long. 86°20'W, by G. M. Treinen. Specimen DH-5.

Male, DML 165 rom, weight 130 g. Parts saved: buccal

mass only.

Dosidicus gigas

Collected 21-22 January 1972, lat. 9°N,

long. 86°20'W, by G. M. Treinen. Specimen DH-6.

Male, DML 167 rom, weight 120 g. Parts saved: buccal

mass only. LRL 3.5 rom.

Dosidicus giqas

Collected 14 April 1972, lat. 9°29'N, long. 100050'W,

by C. W. Oliver. Specimen DH-17. Female, DML 174 rom,

weight 135 g. Parts saved: buccal mass only.
109

Family Ommastrephidae

Subfamily Ommastrephinae (Cont.).

NMML 94.

NMML 95.

NMML 96.

NMML 97.

NMML 97a.

Dosidicus qigas

Collected 14 April 1972, lat. 9°29'N,

long. 100050'W, by C. W. Oliver. Specimen DH-18.

Female, DML 210 mm, weight 237 g. Parts saved:

buccal mass only.

Dosidicus giqas

Collected 16 January 1972, lat. 7°10'N,

long. 94°56'W, by D. B. Holts. Specimen DH-19.

Female, DML 185 mm, weight 230 g. Parts saved:

buccal mass only.

Dosidicus gigas

Collected 21-22 January 1972, lat. 9°N,

long. 86°20'W. MV Eastern Pacific. Parts saved:

8 entire specimens.

Dosidicus gigas

Collected 14 July 1976, Devereux Beach, Goleta,

California, by C. Smith, B. Britt, and K. Rindeaub,

of Santa Barbara Museum of Natural History, Santa

Barbara, California. Washed ashore. Parts saved:

1 entire specimen, DML 288 mm.

Buccal mass (radula, beaks) of 97.
110

Family Omrnastrephidae

Subfamily Omrnastrephinae (Cont.).

NMML 98.

NMML 98a.

NMML 99.

NMML 100.

NMML 101.

Dosidicus qiqas

Collected 10 April 1968, lat. 10031'N,

long. 92°52'W, by W. F. Perrin. Juvenile male?,

DML 378 rom, weight 495 g. Parts saved: 1 entire

specimen.

Beaks of 98 lower beak - um2.er beak

rum

6.5 rom 5.8 rom20.0 rom 6.0 rom24.5 rom10.0 rom4.5 rom10.0 rom

Collected 12 September 1976, Simonton Cove, San

Miguel Island, California, by G. A. Antonelis.

Washed ashore. Parts saved: buccal mass (beaks and

radula). LRL 8.0 rom.

Dosidicus gigas

Collected 12 September 1976, Simonton Cove, San

Miguel Island, California, by G. A. Antonelis.

Washed ashore. Parts saved: buccal mass.

LRL 9.5 rom.

Dosidicus gigas

Collected 12 September 1976, Simonton Cove, San

Miguel Island, California, by G. A. Antonelis.

Washed ashore. Parts saved: buccal mass.

LRL 8.5 rom.
111

Family Ommastrephidae

Subfamily Ommastrephinae (Cont.).

NMML 102.

NMML 102a.

NMML 103.

NMML 104.

NMML 105.

NMML 434.

Dosidicus gigas

Collected 21 July 1966, lat. 11°33'N, long. 91°20'W,

by W. F. Perrin. MV Conte Bianco, by dip net.

Time: 2200 hrs. Parts saved: 1 entire specimen.

Det. by C. F. E. Roper.

Buccal mass of beaks of 102.

Dosidicus giqas

Collected 17 April 1968, lat. 7°50'N, long. 92°32'W,

by W. F. Perrin. MV Carol Virqinia, by dip net at

night. Parts saved: 2 entire specimens. Det. by C.

F. E. Roper.

Dosidicus gigas

Collected 18 April 1968, lat. 8°10'N, long. 92°25'W,

by W. F. Perrin. MV Carol Virginia, by dip net at

night. Parts saved: 1 entire specimen. Det. C. F.

E. Roper.

Dosidicus giqas

Collected 17 April 1968, lat. 7°50'N, long. 92°32'W,

by W. F. Perrin. From the stomach of Thunnus

albacares. MV Carol Virginia. Parts saved: 3

specimens. Det. C. F. E. Roper.

Dosidicus gigas

Collected 18 August 1984, Guadalupe Island, Mexico

by R. L. DeLong, from the scat of A. townsendi,

specimen AT-84-6. Parts saved: 2 upper and 2 lower

beaks.
112

Family Ommastrephidae

Subfamily Ommastrephinae (Cont.).

NMML 106.

NMML 106a.

NMML 107.

NMML 107a.

NMML 435.

Sthenoteuthis (Svrnplectoteuthis) oualaniensis8

Collected 18 April 1968, lat. 8°10'N, long. 92°25'W,

by W. F. Perrin. MV Carol Virginia, by dip net.

Time: 2,000 hrs. Parts saved: 1 entire specimen.

Det. by C. F. E. Roper.

Buccal mass and beaks of 106.

Sthenoteuthis (Symplectoteuthis) oualaniensis

Collected 21 July 1966, lat. 11°33'N, long. 91°21'W,

by W. F. Perrin. By dip net. Time: 2200 hrs.

Parts saved: 1 entire specimen. Det. C. F. E.

Roper.

Buccal mass and beaks of 107.

Eucleoteuthis (Symplectoteuthis) luminosa

Collected 18 August 1984, Guadalupe Island, Mexico,

by R. L. DeLong, from the scat of A. townsendi,

specimen A-84-1. Parts saved: 4 upper and 5 lower

beaks.

8See Roeleveld, M. A. (1982), p. 253 footnote.
113

Order Teuthida

Suborder Oegopsida

Family Thysanoteuthidae

NMML 109. Thysanoteuthis rombus

Collected lS April 1972, lat. 8°20'N,

long. 102°40'W, by C. W. Oliver. MV Lois Seaver, by

dip net. Time: 2000 hrs. Parts saved: 2 entire

specimens. Note: About 6 were around the vessel,

including 1 approximately 2 ft long.
114

Order Teuthida

Suborder Oegopsida

Family Chiroteuthidae

NMML 146.

NMML 146a.

NMML 147.

NMML 147a.

NMML 148.

Chiroteuthis calyx (See Young 1972 and Jefferts

1983). Collected 27 March 1968, lat. 49°46'N,

long. 176°29'W, by R. D. Carlson. RV George ~.

Kelez, cruise 43, set 5, by gill net (75 ft depth).

Surface water temperature 3.6°C. Parts saved: head

and arms only, tentacles and body missing.

Beaks and radula of 146.

Chiroteuthis sp.

Collected March 1966, between latitudes 34°50'N and

36°50'N, off California. Remains caught on outside

of stomach of C. ursinus, specimen US66-297. (A

series number cannot be assigned with certainty.

All stomachs in barrel were taken in March 1966,

between latitudes 34°50'N and 36°50'N off

California). Parts saved: arms, tentacle

(incomplete), buccal mass.

Beaks of 147.

Chiroteuthis sp.

Collected 20 May 1970, lat. 46°43'N, long. 127°35'W.

From the stomach of ~. ursinus, specimen US70-302.

Parts saved: arms, tentacles (somewhat frayed),

buccal mass, beaks.
115

NMML 152.

NMML 154.

NMML 153.

NMML 155.

Family Chiroteuthidae (Cont.).

NMML 150. Chiroteuthis sp.

Collected 15 May 1970, lat. 46°22'N, long. 125°03'W.

From the stomach of Q. ursinus, specimen US70-239.

Parts saved: I pair beaks.

NMML 151. Chiroteuthis calyx (see Stroud 1968)

Collected 20 April 1967, Mukkaw Bay, Washington.

From the stomach of stranded Grampus griseus,

specimen 3769. Parts saved: Beaks and fragments of

7 specimens.

Chiroteuthis sp.

Collected 24 May 1970, lat. 47°38'N, long. 128°27'W.

From the stomach of Q. ursinus, specimen US70-326.

Parts saved: 1 pair beaks.

Chiroteuthis sp.

Collected 9 July 1976, Fort Stevens State Park,

Oregon, by R. K. Stroud. From stomach of stranded

Mirounga angustirostris, specimen MM76-13. Parts

saved: 1 lower beak.

Chiroteuthidae

Collected 9 April 1968, lat. 12°51'N, long. 93°18'W,

by W. F. Perrin. From the stomach of Stenella

graffmani, specimen CV285. Parts saved: small

fragments.

Chiroteuthis sp.

Collected 24 May 1970, lat. 47°38'N, long. 128°19'W.

From the stomach of ~. ursinus, specimen US70-325.

Parts saved: 1 pair large beaks.
116

Family Chiroteuthidae (Cont.).

NMML 156. Chiroteuthis sp.

Collected 3 May 1970, lat. 46°24'N, long. 126°12'W.

From the stomach of Q. ursinus. Specimen US70-218.

Parts saved: 1 pair beaks.

NMML 444. Chiroteuthis calyx

Collected 29 April 1985, San Miguel Island,

California, by G. A. Antonelis, by lavage from

stomach of M. angustirostris, specimen A-26. Parts

saved: 2 lower beaks.

NMML 445. Chiroteuthis calyx

Collected 29 April 1984, San Miguel Island,

California, by G. A. Antonelis, by lavage from

stomach of M. angustirostris, specimen A-34. Parts

saved: 3 lower beaks.
117

Order Teuthida

Suborder Oegopsida

Family Mastigoteuthidae

NMML 149. ?Mastigoteuthis sp.

Collected 25 November 1966, off central California.

From the stomach of E. macrocephalus, specimen Rice

1966-165. Male. Parts saved: 1 lower beak.

LRL 8.0 rom, HL 8.0 rom, CL 15.0 rom, wings dark. Det.

C. H. Fiscus and M. R. Clarke. F-4 Plymouth, June

1981.
118

Order Teuthida

Suborder Oegopsida

Family Cranchiidae

Subfamily Cranchiinae

NMML 124.

NMML 446.

NMML 487.

Cranchia scabra

Collected 6 February 1959, lat. 35°09'N,

long. 121°35'W, by C. H. Fiscus. From the stomach

of Delphinus delphis, specimen 3801, (Tac-2-59).

Parts saved: 2 relatively entire specimens.

Cranchia scabra

Collected 29 April 1984, San Miguel Island,

California, by G. A. Antonelis, by lavage from

stomach of M. angustirostris, specimen A-28. ~arts

saved: 2 lower beaks, 1 upper beak.

Cranchia scabra

Collected 8 May 1986, Adams Cove, San Miguel Island,

California. From stomach of female

M. angustirostris by lavage, specimen E912. Parts

saved: 1 lower beak, LRL 2.5 rom.
119

Order Teuthida

Suborder Oegopsida

Family Cranchiidae

Subfamily Taoniinae

NMML 373.

NMML 373a.

NMML 273b.

NMML 374.

NMML 375

Taoni us pavo9

Collected 25 July 1982, lat. 58°04.3'N,

long. 179°33.9'W, by R. W. Mercer. MV Dominator,

cruise 82-1, haul 64.4, by midwater trawl. Depth

350 fathoms, bottom 1,970 fathoms. Time: 0045 hrs.

DMLs 258 and 272 rom (crowns missing). Parts saved:

4 entire specimens. Det. K. Jefferts.

1 entire specimen, DML 227 rom.

Beaks and radula, DML 300 rom, LRL 6.0 rom.

Taonius pavo

Collected 27 July 1982, lat. 53°24.6'N,

long. 176°40.2'W, by R. W. Mercer. MV Dominator,

cruise 82-1, haul 68, by midwater trawl. Depth

350 fathoms, bottom 2,000 fathoms. Time: 0013 hrs.

DML 158 rom. Parts saved: 1 entire specimen. Det.

K. Jefferts.

Taonius pavo

Collected 1 August 1982, lat. 53°13.6'N,

long. 163°50,3'W, by R. W. Mercer. MV Dominator,

cruise 82-1, haul 75, by midwater trawl. Depth

372 fathoms, bottom 2,300 fathoms. Time: 0031 hrs.

DMLs 166, 195, and 245 rom. Parts saved: 3 entire

specimens (2 + 1 body + crown). Det. K. Jefferts.

~ote: NMML Nos. 373, 374, 375, 122, 122a, 123 loaned to N.

Voss 13 April 1987.
120

Family Cranchiidae

subfamily Taoniinae (Cont.).

NMML 274a.

NMML 122.

NMML 122a.

NMML 123.

NMML 274.

Taonius pavo

Collected 13 December 1966, off central California.

From the stomach of male, E. macrocephalus, specimen

Rice 1966-189. Parts saved: 1 lower beak,

LRL 8.5mm.

Taonius sp.

Collected 4 September 1964, lat. 45°55'N,

long. 124°55'W. MV Commando, depth 400 fathoms.

Time: 1600 hrs. Parts saved: 1 entire specimen.

Somewhat distorted, DML 185 mm.

Beaks and buccal mass of 122. LRL 5.5 mm.

Taoniinae

Collected 28 May 1964, lat. 45°35'N,

long. 124°53.3'W. MV Commando. Parts saved:

1 entire specimen.

Megalocranchia (=Phasmatopsis)

Collected 13 December 1966, off central California.

From the stomach of male E. macrocephalus, specimen

Rice 1966-189. Parts saved: 2 upper, 1 lower beaks,

LRL 15 mm, F5 Plymouth 1981. Det. M. R. Clarke.
121

Family Cranchiidae

Subfamily Taoniinae (Cont.).

NMML 390

NMML 481.

NMML 482.

NMML 483.

NMML 484.

Megalocranchia sp.

Collected 16 September 1961, off central California.

From the stomach of male R. macrocephalus, specimen

Rice 535. Parts saved: 1 lower beak, LRL 14.5 rom.

Megalocranchia sp.

Collected 14 June 1966, off central California.

From the stomach of male R. macrocephalus, specimen

Rice 1966-51. Parts saved: 1 lower beak,

LRL 7.5 rom.

Megalocranchia sp.

Collected 9 December 1966, off central California.

From the stomach of female R. macrocephalus,

specimen Rice 1966-184. Parts saved: 1 upper beak,

1 lower beak, LRL 6.5 rom.

Megalocranchia sp.

Collected 12 May 1967, off central California. From

the stomach of male R. macrocephalus, specimen Rice

1967-131. Parts saved: 1 upper beak, 1 lower beak,

LRL 13.5 rom.

Megalocranchia sp.

Collected 15 May 1968, off central California. From

stomach of male R. macrocephalus, specimen Rice

1968-120. Parts saved: 1 upper beak.
122

Family Cranchiidae

Subfamily Taoniinae (Cont.).

NMML 126a.

NMML 127.

NMML 127a.

NMML 127b.

NMML 127c.

NMML 132.

Galiteuthis sp.

Collected 20 April 1965, off central California.

From the stomach of male g. macrocephalus, specimen

Rice 1965-5. Parts saved: 1 lower beak.

Galiteuthhis sp.

Collected 29 October 1966, off central California.

From the stomach of male g. macrocephalus, specimen

Rice 1966-156. Parts saved: 1 pair beaks.

Galiteuthis sp.

1 buccal mass, 7-rowed radula, head and arms

fragment of 127. Number 3 of 4 jars. Beaks,

LRL 5.0 rom (Fiscus - type "1").

Galiteuthis sp. (pacifica?)

1 buccal mass, 7-rowed radula, head and arms

fragment of 127. Number 4 of 4 jars (Fiscus -

type "1"). Beaks, LRL 5.5 rom.

Galiteuthis sp. (pacifica?)

1 pair of beaks of 127b. F14 PlYmouth 1981. Det.

C. H. Fiscus and M. R. Clarke.

Galiteuthis sp.

Collected 9 May 1967, off central California. From

the stomach of male g. macrocephalus, specimen Rice

1967-119. Parts saved: 1 lower beak.
123

Family Cranchiidae

Subfamily Taoniinae (Cont.).

NMML 137.

NMML 215.

NMML 215a.

NMML 376.

NMML 448.

Galiteuthis sp.

Collected 29 April 1968, off central California.

From the stomach of female £. macrocephalus,

specimen Rice 1968-111. Parts saved: 1 gladius, 1

beak (Fiscus - type "1" - patch on wing stage) .

Galiteuthis sp. (probably phvllura)

Collected 23 April 1968, off central California.

From the stomach of £. macrocephalus, specimen Rice

1968-108. Parts saved: 2 heads, arms, buccal mass

fragments, lower beak (Fiscus - type "1"). F16

PlYmouth 1981. Det. M. R. Clarke, PlYmouth,

June 1981.

Beaks of 215.

Galiteuthis phyllura

Collected 27 July 1982, lat. 53°24.6'N,

long. 176°40.2'W, by R. W. Mercer. MV Dominator,

cruise 82-1, haul 68, by midwater trawl. Depth

350 fathoms, bottom 2,000 fathoms. Time: 0013 hrs.

DMLs 104 and 105 rom. Parts saved: 2 entire

specimens. Det. K. Jefferts.

Galiteuthis sp.

Collected 30 April 1984, San Miguel Island,

California, by G. A. Antonelis, by lavage from

stomach of M. angustirostris, specimen A-38. Parts

saved: 2 lower beaks. Det. F. G. Hochberg and C. H.

Fiscus.
124

Family Cranchiidae

Subfamily Taoniinae (Cont.).

NMML 449.

NMML 450.

NMML 126a.

NMML 129.

Galiteuthis sp.

Collected 30 April 1984, San Miguel Island,

California, by G. A. Antonelis, by lavage from

stomach of M. anqustirostris, specimen A-42. Parts

saved: 1 lower beak, 1 upper beak. Det. F. G.

Hochberg and C. H. Fiscus.

Galiteuthis sp.

Collected 28 April 1984, San Miguel Island,

California, by G. A. Antonelis, by lavage from

stomach of M. angustirostris, specimen A-21. Parts

saved: 1 lower beak, LRL 5.5 rom. Det. F. G.

Hochberg and C. H. Fiscus.

Galiteuthis sp.

Collected 20 April 1965, off central California.

From the stomach of male g. macrocephalus, specimen

Rice 1965-5. Parts saved: 1 lower beak.

Galiteuthis sp.

Collected 9 December 1966, off central California.

From the stomach of male g. macrocephalus, specimen

Rice 1966-181. Parts saved: 1 lower beak,

LRL 6.0 rom.
125

Family Cranchiidae

Subfamily Taoniinae (Cont.).

NMML 135.

NMML 135a.

NMML 136a.

NMML 121.

Galiteuthis sp.

Collected 16 November 1967, off central Califiornia.

From the stomach of male £. macrocephalus, specimen

Rice 1967-241. Parts saved: 2 lower beaks (Fiscus -

type "1"), LRL 5.0 rom (patch on wing stage), LRL 6.0

rom (wing dark same as NMML 129) .

Galiteuthis sp.

Collected 16 November 1967, off central California.

From the stomach of male £. macrocephalus, specimen

Rice 1967-241. Parts saved: 1 lower beak (Fiscus -

type "2"), LRL 5.5 rom.

Galiteuthis sp.

Collected 23 April 1968, off central California.

From the stomach of female £. macrocephalus,

specimen Rice 1968-108. Parts saved: 1 lower beak

(Fiscus - type "1").

Mesonychoteuthis sp.

Collected 19 October 1970, off central California.

From the stomach of male £. macrocephalus, specimen

Rice 1970-43. Parts saved: 2 pair beak fragments,

LRLs 30 and 32 rom. Det. M. R. Clarke, July 1981.

F-1 PlYmouth 81.
126

Family Cranchiidae

Subfamily Taoniinae (Cont.).

NMML 125a

NMML 130.

NMML 131.

NMML 133.

Cranchiidae10 (resembles Galiteuthis)

Collected 26 May 1964, off central California. From

the stomach of female g. macrocephalus, specimen

Rice 1047. Parts saved: 1 lower beak.

Cranchiidae (resembles Galiteuthis)

Collected 5 April 1967, off central California.

From the stomach of male g. macrocephalus, specimen

Rice 1967-105. Parts saved: 1 lower beak (Fiscus -

type "1").

Cranchiidae

Collected 5 April 1967, off central California.

From the stomach of male g. macrocephalus, specimen

Rice 1967-105. Parts saved: 1 gladius (incomplete),

measured 305 nun.

Cranchiidae

Collected 3 June 1967, off central California. From

the stomach of female g. macrocephalus, specimen

Rice 1967-133. Parts saved: 2 gladii. 1 complete

gladius measured 450 nun; 1 incomplete gladius

measured 350 nun.

l~emainder of Cranchiidae listed only to family. See

Fiscus1 memorandum of 23 January 1983 "Review of cranchiids or

beaks thought to belong to the family Cranchiidae" (NMML 125-141)

in notes and reprint file.
127

Family Cranchiidae

Subfamily Taoniiinae (Cont.)

NMML 140.

NMML 141a.

Cranchiidae

Collected 10 March 1966, lat. 35 °08'N,

long. 121°25'W. From the stomach of ~. obliquidens,

specimen 3218. Parts saved: 1 pair beaks.

Cranchiidae?

Collected (date and location unknown) (Alaska?).

From the stomach of male Eumetopias jubatus,

specimen SUV-5-76. Parts saved: 1 beak.
128

Order Vampyromorpha

Family Vampyroteuthidae

NMML 269.

NMML 270.

NMML 384.

NMML 385.

Vampyroteuthis infernalis

Collected 25 November 1966, off central California.

From the stomach of male g. macrocephalus, specimen

Rice 1966-165. Parts saved: 1 lower beak, HL 13 mrn.

F-3 Plymouth 81. Det. C. H. Fiscus and M. R.

Clarke.

Vampyroteuthis infernalis

Collected 1 December 1969, off central California.

From the stomach of female g. macrocephalus,

specimen Rice 1969-178. Parts saved: 3 lower beaks,

HL 14, 13.5, 11 mrn. F-7 Plymouth 81. Det. C. H.

Fiscus and M. R. Clarke.

Vampyroteuthis infernalis

Collected 26 May 1964, off central California. From

the stomach of female g. macrocephalus, specimen

Rice 1047. Parts saved: 1 lower, HL 12.5 mrn, 2

upper beaks.

Vampyroteuthis infernalis

Collected 19 May 1968, off central California. From

the stomach of male g. macrocephalus, specimen Rice

1968-125. Parts saved: 1 lower, HL 11.0 mrn, 1 upper

beaks.
NMML 451

129

Family Vampyroteuthidae

NMML 386. Vampyroteuthis infernalis

Collected 12 May 1969, off central California. From

the stomach of female ~. macrocephalus, specimen

Rice 1969-115. Parts saved: 1 lower, HL 14.0 mm,

1 upper beaks.

NMML 387. Vampyroteuthis infernalis

Collected 10 May 1969, off central California. From

the stomach of female ~. macrocephalus, specimen

Rice 1969-107. Parts saved: 2 lower beaks,

HL 11.5 and 9.0 rom.

Vam?vroteuthis infernalis

Collected 28 April 1984, San Miguel Island,

California, by G. A. Antonelis, by lavage from the

stomach of M. angustirostris, specimen A-24. Parts

saved: 1 lower beak, HL 7.0 rom; 1 upper beak.
130

Order Octopoda

Suborder Cirrata

Family Opisthoteuthidae

NMML 1.

NMML 1a.

NMML 2.

NMML 2a.

NMML 3.

Opisthoteuthis californiana (Pereyra 1965)

Collected 11 December 1961, lat. 42°55.9'N,

long. 124°48.1'W, southwest of the mouth of the

Columbia River. MV Commando 3 S. Station 14A. In

375 fathoms. Parts saved: 1 entire specimen.

Beaks of 1

Opisthoteuthis californiana (Pereyra 1965)

Collected 17 September 1962, lat. 59°34'N,

long. 145°45'W, Gulf of Alaska. International

Pacific Halibut Commission, haul W208. Parts saved:

1 entire specimen.

Beaks of 2.

Opisthoteuthis californiana (Pereyra 1965)

Collected 17 September 1962, lat. 59°33'N,

long. 145°17'W, Gulf of Alaska. International

Pacific Halibut Commission, haul W207. Parts saved:

1 entire specimen.
131

Order Octopoda

Suborder Incirrata

Family Bolitaenidae

NMML 56.

NMML 57.

NMML 377.

Japetella heathi

Collected (no date), south of the mouth of the

Columbia River, Oregon. On loan from AFSC, RACE,

Columbia River outfall studies. Parts saved: 1

entire specimen. Det. by W. Pearcy.

Japetella heathi

Collected 23-24 August 1971, lat. 50050'N,

long. 177°19'E. RV George ~. Kelez, cruise 53,

Parts saved: 1 entire specimen.

Japetella heathi

Collected 27 July 1982, lat. 53°24.6'N,

long. 176°40.2'W, by R. W. Mercer. MV Dominator,

NMML 377a.

NMML 378.

cruise 82-1, haul 68, by midwater trawl. Depth

350 fathoms, bottom 2,000 fathoms. Time: 0013 hrs.

Parts saved: 3 entire specimens. Det. K. Jefferts.

Beaks of 1 specimen with DML approximately 60 rom.

Japetella heathi

Collected 1 August 1982, lat. 53°13.6'N,

long. 163°50.3'W, by R. W. Mercer. MV Dominator,

cruise 82-1, haul 75, by midwater trawl. Depth

372 fathoms, bottom 2,300 fathoms. Time: 0031 hrs.

Parts saved: 6 entire specimens. Det. K. Jefferts.
132

Family Bolitaenidae (Cont.).

NMML 488. Japetella sp.

Collected 24 April 1985, San Miguel Island,

California, from stomach of female

M. angustirostris, specimen A90. Parts saved:

1 pair beaks.

NMML 489. Japetella sp.

Collected 12 May 1986, San Miguel Island,

California, from stomach of male M. angustirostris

by lavage, specimen E-980. Parts saved: 2 pair

beaks.
133

Order Octopoda

Suborder Incirrata

Family Octopodidae

NMML4.

NMML4a.

NMML 5.

NMML 6.

NMML 7.

Octopus dofleini

Collected 18 September 1962, lat. 59°40'N,

long. 145°41'W, Gulf of Alaska. International

Pacific Halibut Commission, haul 210. Parts saved:

1 entire specimen. Det. F. G. Hochberg.

Beaks and radula of 4.

Octopus dofleini

Collected 18 September 1962, lat. 59°40'N,

long. 145°41'W, Gulf of Alaska. International

Pacific Halibut Commission, haul 210. Parts saved:

1 entire specimen, male. Det. F. G. Hochberg.

Octopus dofleini

Collected 22 June 1960, Caton Island, Alaska, by K.

W. Kenyon and T. O'Brien. Tide pool. Time:

0630 hrs. Parts saved: 1 entire specimen. Det. F.

G. Hochberg.

Octo~us dofleini

Collected 10 December 1965, Naden Harbor, Graham

Island, Queen Charlotte Islands, British Columbia,

Canada, by C. H. Fiscus. Parts saved: 1 entire

specimen. Det. F. G. Hochberg.
134

Family Octopodidae (Cont.).

NMML 207.

NMML 235.

NMML 235a.

NMML 236.

Octopus dof1eini

Collected 14 May 1964, off central California. From

stomach of male ~. macrocephalus, specimen Rice

1019. Parts saved: radulas. 3 pair beaks,

Crest Length Hood Lenqth

18.0 rom41.5 rom16.2 rom37.8 rom16.0 rom 37.8 rom

Octopus dofleini

Collected 29 October 1981, lat. 56°40'N,

long. 170003.6'W. RV Miller Freeman, cruise 81-6,

haul 29, by trawl. Depth of trawl and bottom,

32 fathoms. Surface water temperature 7.6°C, trawl

temperature 7.7°C. Frozen fresh. DML 56 rom, weight

30 g (in laboratory). Parts saved: 1 entire

specimen. Det. F. G. Hochberg. On permanent loan

to Santa Barbara Museum of Natural History.

Beaks and radula of 235.

Octopus dofleini

Collected 1 November 1981, lat. 55°04.6'N,

long. 166°58.6'W. RV Miller Freeman, cruise 81-6.

haul 32, by trawl. Depth 85 fathoms. Surface water

temperature 7.3°C, trawl temperature 5.7°C. Frozen

fresh. Female, DML 350 rom, weight 14.2 kg in

laboratory, weight of ovary 775 g. Parts saved:

beaks and radula.
NMML 237a.

NMML 239.

135

Family Octopodidae (Cont.).

NMML 236a. Statoliths of 236.

NMML 237. Octopus dofleini

Collected 27 October 1981, lat. 57°0'N,

long. 169°33'W. RV Miller Freeman, cruise 81-6,

haul 24, by trawl. Depth 32 fathoms. Surface water

temperature 7.6°C, trawl temperature 7.7°. Frozen

fresh. Male, DML 340 mm, weight 7 kg in laboratory.

Parts saved: 1 entire specimen.

Beaks and radula of 237.

Octopus dofleini

Collected 21 October 1981, lat. 54°50'N,

long. 166°32'W. RV Miller Freeman, cruise 81-6,

haul 19, by trawl. Depth 100 fathoms. Surface

water temperature 7.9°C, trawl temperature 5.00C.

Frozen fresh. Male, field weight 13.4 kg,

HL 245 mm. Parts saved: third right hectocotilized

NMML 239a.

NMML 239b.

arm .

Statoliths of 239.

Beaks and radula of 239.
NMML 245a.

NMML 240a.

NMML 245.

136

Family Octopodidae (Cont.).

NMML 240. Octopus dofleini

Collected 18 October 1981, lat. 55°41'N,

long. 165°49'W. RV Miller Freeman, cruise 81-6,

haul 14, by trawl. Depth 62 fathoms. Surface water

temperature 8.7°C, trawl temperature 5.3°C. Frozen

fresh. Female (based on no hectocotilization on

arms), DML 380 rom (organs removed in field,

measurement approximate), field weight 18.6 kg.

Parts saved: beaks and radula

Statoliths of 240.

Octopus dofleini

Collected October 1981, lat. 57°03'N,

long. 140041'W. RV Miller Freeman, cruise 81-6,

haul 28, by trawl. Depth 48 fathoms. Trawl

temperature 7.0oC. Frozen fresh. Male, DML 115 rom,

weight 382 g (in laboratory). Parts saved: 1 entire

specimen. Det. F. G. Hochberg. On permanent loan

to Santa Barbara Museum of Natural History.

Beaks and radula of 245.
NMML 246a.

NMML 259.

NMML 259a.

NMML 260.

137

Family Octopodidae (Cont.).

NMML 246. Octopus dofleini

Collected 21 October 1981, lat. 54°51'N,

long. 166°52'W. RV Miller Freeman, cruise 81-6,

haul 19, by trawl. Depth 107 fathoms. Surface

water temperature 7.9°C, trawl temperature 5.0oC.

Frozen fresh. Male, DML 120 rom, weight 386 g (in

laboratory) . Parts saved: 1 entire specimen. Det.

F. G. Hochberg. On permanent loan to Santa Barbara

Museum of Natural History.

Beaks and radula from 246.

Octopus dofleini

Collected 20 October 1981, lat. 54°14'N,

long. 166°02'W. RV Miller Freeman, cruise 81-6,

haul 17, by trawl. Depth 59 fathoms. Trawl

temperature 4.4°C. Frozen fresh. Female,

DML 43 rom, weight 18 g (fresh weight 25.2 g). Parts

saved: 1 entire specimen. Det. F. G. Hochberg. On

permanent loan to Santa Barbara Museum of Natural

History.

Beaks and radula of 259.

Octopus dofleini

Collected (no date), Bering Sea. RV Miller Freeman,

cruise 81-6, by trawl. Frozen fresh. Female,

DML 95 rom, weight 227 g. Parts saved: 1 entire

specimen. Det. F. G. Hochberg. On permanent loan

to Santa Barbara Museum of Natural History.
138

Family Octopodidae (Cont.).

NMML 260a. Beaks and radula of 260.

NMML 261. Octopus dofleini

Collected 4 November 1981, lat. 54°N,

long. 164°34'W. RV Miller Freeman, cruise 81-6,

haul 40, by trawl. Depth 51 fathoms. Trawl

temperature 7.6°C. Frozen fresh. Male, DML 240 rom,

weight 5.4 kg in laboratory. Ligula 85 rom, open.

Parts saved: 1 entire specimen.

NMML 261a. Beaks and radula of 261.

NMML 262.

NMML 262a.

NMML 262b.

Octopus dofleini

Collected 3 November 1981, lat. 55°42'N,

long. 168°39'W. RV Miller Freeman, cruise 81-6,

haul 39, by trawl. Depth 77 fathoms. Trawl

temperature 7.8°C. Frozen fresh. Female, DML 390

rom, field weight 21.7 kg. Weight of ovary 2,286 g,

eggs not ripe, small, 3-4 mm. Parts saved: arm R2

(length 1,540 mm) .

Beaks and radula (note: radula agrees with Sasaki

(1929) text, Fig. 35) of 262.

Statoliths of 262.
NMML 368a.

NMML 369.

NMML 369a.

NMML 369b.

139

Family Octopodidae (Cont.).

NMML 368. Octopus dofleini

Collected 2 October 1982, lat. 56°42'N,

long. 169°32'W, by T. R. Loughlin. RV Miller

Freeman, cruise 82-3, haul 34, by trawl. Bottom

depth 52 fathoms. Surface temperature 7.2°C, trawl

temperature 5.8°C. Frozen fresh, thawed in

laboratory. Laboratory measurements 16 July 1983.

Female, DML 140 rom, total length 490 rom, field

weight 454 g. Suckers at web edge somewhat enlarged

on arms 1 and 2. Parts saved: 1 entire specimen.

Beaks and radula of 368.

Octopus dofleini

Collected 28 September 1982, lat. 55°40'N,

long. 165°11'W, by T. R. Loughlin. RV Miller

Freeman, cruise 82-3, haul 16, by trawl. Bottom

depth 58 fathoms. Surface temperature 7.9°C. bottom

5.3°C. Frozen fresh, thawed in laboratory.

Laboratory measurements 19 July 1983. Male,

DML 390 rom, TL 1,565 rom, field weight 22.7 kg.

HAL 1170 rom, HL 240 rom, funnel length, 90 rom.

Largest sucker diameter 45 to 50 rom. Gill lamella

left 11 + 12, right 12 + 12. Gonad weight 446 g.

Parts saved: third right arm.

Beaks and radula of 369.

2 statoliths of 369.
NMML 469.

NMML 370a.

NMML 459.

140

Family Octopodidae (Cont.).

NMML 370. Octopus dofleini

Collected 3 October 1982, lat. 56°20'N,

long. 168°18'W, by T. R. Loughlin. RV Miller

Freeman, cruise 82-3, haul 38, by trawl. Bottom 86

fathoms. Surface temperature 7.5°C, bottom 3.9°C.

Frozen upon capture, thawed in laboratory.

Laboratory measurements 19 July 1983. Male,

DML 280 mm, TL 1,425 mm, field weight 15.4 kg,

HAL 825 mm, HL 235 mm. Largest sucker diameter

40 mm. Gill lamella left 12 + 12. right 12 + 12.

Parts saved: third right arm.

Beaks and radula of 370.

Octopus dofleini

Collected 26 June 1973, lat. 55°20'N,

long. 168°05'W, by C. H. Fiscus. MV Mark I, cruise

73-36, Sta. B 18, haul 2, by bottom trawl. Depth

142 fathoms (259 m). Water temperature, surface

7.2°C, bottom 4.0oC. Female, DML 70 mm. Parts

saved: entire specimen. Det. F. G. Hochberg.

Octopus dofleini

Collected 26 June 1973, lat. 55°00'N,

long. 167°31'W, by C. H. Fiscus. MV Mark~, cruise

73-36, Sta. A-6, haul 1, by bottom trawl. Depth not

recorded. Water temperature, surface not recorded,

bottom not recorded. Male, DML not measured. Parts

saved: entire specimen. Det. F. G. Hochberg.
NMML 265.

NMML 264.

recorded.

141

Family Octopodidae (Cont.).

NMML 470. Octopus dofleini

Collected 26 June 1973, lat. 55°00'N,

long. 167°31'W, by C. H,. Fiscus. MV Mark ~, cruise

73-36 Sta. A-I, haul 1, by bottom trawl. Depth not

Water temperature, surface not recorded, bottom not

recorded. Female, DML not measured. Parts saved:

entire specimen. Det. F. G. Hochberg.

Octopus bimaculatus

Collected 4 August 1978, northshore of west end of

San Miguel Island, California, by G. A. Antonelis.

From Zalophus californianus scat. Original number

26A, SMI collection. Parts saved: 1 upper beak

only. Det. F. G. Hochberg.

Octopus rubescens

Collected 31 July 1979, north shore of west end, San

Miguel Island, California, by G. A. Antonelis. From

~. californianus scat. Original specimen number

GAA-87, SMI collection. Parts saved: 13 lower, 18

upper beaks only. Det. F. G. Hochberg.
142

Family Octopodidae (Cont.).

NMML 266. Octopus rubescens

Collected 31 July 1979, north shore of west end, San

Miguel Island, California, by G. A. Antonelis. From

Z. californianus scat. Original specimen number

GAA-94, SMI collection. Parts saved: 16 lower,

16 upper beaks only. Det. F. G. Hochberg.

NMML 267. Octopus rubescens

Collected 16 May 1979, Adams Cove, San Miguel

Island, California, by G. A. Antonelis. From

Z. californianus scat. Original specimen number

GAA-127, SMI Collection. Parts saved: 3 lower,

2 upper beaks. Det. F. G. Hochberg.

NMML 8. Octopus sp.

Collected 9 July 1960, Amak Island, Alaska, by C. H.

Fiscus. From stomach of Phoca vitulina, specimen

3723. Parts saved: 1 pair beaks.

NMML 9. Octopus sp.

Collected 28 August 1966, Puget Sound, Washington,

by H. Kajimura. Parts saved: 1 pair beaks from

specimen with 8 inch tentacles.

NMML 10. Octopus sp.

Collected 10 August 1967, Puget Sound, Washington,

by H. Kajimura. Parts saved: 1 pair beaks.
143

Family Octopodidae (Cont.).

NMML 11. Octopus sp.

Collected 29 January 1962, Bering Sea, by H.

Kajimura. From stomach of Gadus macrocephalus.

Parts saved: 2 pair beaks.

NMML 12. Octopus sp. (Kenyon 1965)

Collected 6 March 1962, St. Makerius Point, Amchitka

Island, Alaska, by K. W. Kenyon. From stomach of

adult male g. vitulina, specimen KWK 62-217. Parts

saved: 1 pair beaks plus 1 large fragment.

NMML 13. Octopus sp. (Kenyon 1965)

Collected 6 March 1962, St. Makerius Point, Amchitka

Island, Alaska, by K. W. Kenyon. From stomach of

juvenile female g. vitulina, specimen KWK 62-218.

Parts saved: 2 pair beaks.

NMML 14. Octopus sp. (Kenyon 1965)

Collected 26 October 1962, St. Makerius Point,

Amchitka Island, Alaska, by K. W. Kenyon. From

stomach of Enhydra lutris, specimen KWK 62-307.

Parts saved: 1 pair beaks.

NMML 15. Octopus sp. (Johnson et ale 1966)

Collected 22 May 1961, Point Hope, Alaska. From

stomach of Erignathus barbatus, specimen M-1562.

Parts saved: 5 lower, 7 upper beaks. (4

Q. dofleini, 1 Benthoctopus? - 27 Jan. 1988.)
NMML 25.

NMML 18.

NMML 493.

144

Family Octopodidae (Cont.).

NMML 16. Octopus sp. (Johnson et ale 1966)

Collected 20 March 1961, Point Hope, Alaska. From

stomach of Phoca hispida, specimen M1223. Parts

saved: 1 pair beaks. (Q. dofleini - 27 Jan. 1988.)

NMML 17. Octopus sp. (Johnson et ale 1966)

Collected 4 June 1961, Point Hope, Alaska. From

stomach of P. hispida, specimen M1785. Parts saved:

2 lower, 3 upper beaks. (Q. dofleini - 27 Jan.

1988.)

Octopus sp.

Collected 30 March 1970, lat. 47°17'N,

long. 124°58'W. From stomach of ~. ursinus,

specimen US-70-135. Parts saved: 1 beak.

Octopus sp.

Collected 26 April 1961, Point Hope, Alaska. From

stomach of P. hispida, specimen 1438. Parts saved:

1 upper beak.

Octopus sp.

Collected 17 December 1987, Elliott Bay (Seattle,

WA), by H. Kajimura. By squid jig. DML 29 rom

(preserved), weight 10.8 g (preserved). Parts

saved: entire specimen.
145

Family Octopodidae (Cont.).

NMML 458. Benthoctopus sp. A.

Collected 26 June 1973, lat. 55°20'N,

long. 168°05'W, by C. H. Fiscus. MV Mark~, cruise

73-36, Sta. B18, haul 2, by bottom trawl. Depth

142 fathoms (259 m). Water temperature, surface

7.2°C, bottom 4.00C. Male, DML 35 rom. Parts saved:

entire specimen - beaks separate. Det. F. G.

Hochberg.

NMML 460. Benthoctopus sp. A.

Collected 26 June 1973, lat. 55°20'N,

long. 168°05'W, by C. H. Fiscus. MV Mark~, cruise

73-76, Sta. B18, haul 2, by bottom trawl. Depth

142 fathoms (259 m). Water temperature, surface

7.2°C, bottom 4.0oC. Female, DML56 rom. Parts

saved: entire specimen - beaks separate. Det. F. G.

Hochberg.

NMML 461. Benthoctopus sp. A.

Collected 26 June 1973, lat. 55°20'N,

long. 168°05'W, by C. H. Fiscus. MV Mark~, cruise

73-36, Sta. B18, haul 2, by bottom trawl. Depth

142 fathoms (259 m). Water temperature, surface

7.2°C, bottom 4.0oC. Female, DML 25 rom. Parts

saved: entire specimen- beaks separate. Det. F. G.

Hochberg.
146

Order Octopoda

Suborder Incirrata

Family Tremoctopodidae

NMML 271.

NMML 272.

NMML 272a.

NMML 273.

NMML 273a.

Tremoctopus sp.

Collected 15 April 1973, lat. 6°17'N, long. 81°25'W,

by K. Sexton. MV Antonia Q., set 39. Female,

DML 75 rom, VML 80 rom, MW 55 rom, weight 159 g

(preserved). 4 aquiferous pores present. Parts

saved: 1 entire specimen.

Tremoctopus sp.

Collected 1973, lat. 6°00'N, long. 94°00'W, by

"JAY", set 12. Female, DML 140 rom, VML 100 rom,

MW 80-90 rom, weight 381 g (preserved). 4 aquiferous

pores present. Markings on dorsal surface near

pores (See Robson 1932). Best condition of

271-273.) Parts saved: 1 entire specimen.

Beaks and radula of 272.

Tremoctopus sp.

Collected 27 March 1973, lat. 9°38'N, long. 95°57'W,

by J. Ploeger. MV JFK~, set 15. Time: 1600 hrs.

Female, DML 120 rom, VML 97 rom, MW 65 rom, weight

351 g (preserved). 4 aquiferous pores present.

Parts saved: 1 entire specimen.

Beaks and radula of 273.
147

Family Tremoctopodidae (Cant.).

NOTE: All three (271-273) specimens badly crumpled. DML,

VML, and MW only approximate. Arms of most

specimens broken in varying degrees, as follows:

271. (1), (2) both broken off; (3) R - 100 rom,

272.

273.

L-80each 110 Im'Cl, (4) Im'Cl.

(1)

R - 200 Im'Cl, L - 175 Im'Cl (partial 2), (2)R - 165 Im'Cl, L - 215 L(3) Im'Cl,

R - 160 85 Im'Cl(4)broken. L -

(1)

broken off, R - 300(3)Im'Cl, L broken, (2)

broken off. (4)
148

Order Octopoda

Suborder Incirrata

Family Ocythoidae

NMML 19.

NMML 20.

NMML 21.

NMML 22.

Ocythoe tuberculata

Collected January and February 1961, California.

From the stomachs of b. ursinus, specimens US61.

Parts saved: 23 lower, 16 upper beaks (beaks from

several stomachs, see 1961 NMML files and Pelagic

Fur Seal report for 1961) .

Ocythoe tuberculata

Collected 18 January 1961, lat. 3so18'N,

long. 122°08'W, California. From the stomach of

b. ursinus, specimen US61-10S. Parts saved: 1 pair

beaks.

Ocythoe tuberculata

Collected 18 January 1961, lat. 3so2S'N,

long. 122°11'W, California. From the stomach of

b. ursinus, specimen US61-107. Parts saved: 1 pair

beaks. Det. F. G. Hochberg.

Ocythoe tuberculata

Collected 22 January 1961, lat. 33°S3'N,

long. 120oSS'W, California. From the stomach of

b. ursinus, specimen US61-26S. Parts saved: 1 pair

beaks. Det. F. G. Hochberg.
NMML 490.

NMML 23a.

NMML 24.

149

Family Ocythoidae (Cont.).

NMML 23. Ocythoe tuberculata

Collected 25 January 1961, lat. 34°25'N,

long. 121°08'W, California. From the stomach of

h. ursinus, specimen US61-375. Parts saved: 1 body

fragment and tentacles. Det. F. G. Hochberg.

1 pair beaks of 23.

Ocythoe tuberculata

Collected 31 January 1961, lat. 33°17'N,

long. 120018'W, California. From the stomach of

h. ursinus, specimen US61-425. Parts saved: 1 body

fragment, tentacles, beaks.

Ocythoe tuberculata

Collected 7 July 1989, San Miguel Island,

California. From stomach of male Mirounqa

angustirostris by lavage, specimen Tx 750. Parts

saved: 1 lower and 2 upper beaks.
150

Order Octopoda

Suborder Incirrata

Family Argonautidae

NMML 83. Arqonauta pacifica

Collected 17 March 1973, lat. 22°21'N,

long. 110045'W, by J. Ploeger. MV JFK ~, Southwest

Fisheries Center. Parts saved: 1 entire specimen.
151

Order Octopoda

Suborder Incirrata

Family Allopsidae

NMML 268.

NMML 485.

Alloposus moll is

Collected 25 November 1966, off central California.

From the stomach of male ~. macrocephalus, specimen

Rice 1966-165. Parts saved: 2 lower beaks.

HL 13 rom, CL 27 rom (F-3 PlYmouth 81). Det. C. H.

Fiscus and M. R. Clarke.

Alloposus mollis

Collected 16 September 1961, off central California.

From stomach of male ~. macrocephalus, specimen Rice

1961-535. Parts saved: 1 lower beak. HL 13.0 rom,

CL 25.0 rom damaged.
152

ACKNOWLEDGMENTS

The author gratefully acknowledges the help of the many

people who collected or identified specimens, and to others who

donated specimens collected during cruises of the Pacific Salmon

Investigations, and of the Resource Assessment and Conservation

Engineering Division, AFSC. These people are named in the

catalogue.

The catalogue could not have been assembled without the

able assistance of Kathy Cunningham, who cheerfully suffered

through the several revisions and additions to the catalogue.

Her typing and editorial assistance are most gratefully

acknowledged.
153

CITATIONS

CLARKE, M. R., and E. R. TRUMAN

1988. Introduction (to the Mollusca Vol. 12) Chap. I,

pp 1-10. Table 1. Classifications of the Genera of

Recent Cephalopods. Academic Press.

FISCUS, C. H.

1984. Catalogue of Cephalopods at the National Marine

Mammal Laboratory, NOAA Tech. Memo NMFS F/NWC 65.

FISCUS, C. H.

1991. Notes on North Pacific Gonatids: Identification of

body fragments and beaks from marine mammal stomachs.

West. Soc. Malacol. Ann. Rpt. 23:2-6.

FISCUS, C. H., and D. W. RICE.

1974. Giant squids, Architeuthis sp., from stomachs of

sperm whales captured off California. Calif. Fish Game

60(2) :91-93.

FISCUS, C. H., D. W. RICE, and A. A. WOLMAN.

1989. Cephalopods from the stomachs of sperm whales taken

off California. U.S. Dep. Commer., NOAA Tech. Rep.

NMFS 83, 12 p.

FITCH, J. E., and R. L. BROWNELL, Jr.

1971. Food habits of the franciscana pontophoria

blainvillei (Cetacea: Platanistidae from South America) .

Bull. Mar. Sci. 21:626-636.

JEFFERTS, K.

1983. Zoogeography and systematics of cephalopods of the

northeastern Pacific Ocean. Ph.D. Diss., Oregon State

University, Corvallis, Oregon, 291 p.
154

JOHNSON, M. L., C. H. FISCUS, B. T. OSTENSON, and M. L. BARBOUR.

1966. Marine mammals. In N. J. Wi1imovsky and J. N. Wolfe

(editors), Environment of the Cape Thompson region,

Alaska. Book 2, p. 877-924. U. S. Atomic Energy Comm.,

Energy Res. Dev. Admin., Tech. Inf. Cent., Oak Ridge,

Tenn. (Available from U. S. Dep. Commer., Natl. Tech.

Inf. Serv., Springfield, VA, as PNE-481.)

KENYON, K. W.

1965. Food of harbor seals at Amchitka Island, Alaska.

J. Mammal. 46:123-124.

MACASKIE, I. B.

1971. A range extension for the squid Abraliopsis. J.

Fish. Res. Board Can. 28:620-621.

MERCER, R. W., and M. BUCY.

1983. Experimental squid jigging off the Washington coast.

U. S. Nat. Mar. Fish. Serv., Mar. Fish. Rev. 45(7-8-

9) :56-62.

OKUTANI, T., T. KUBODERA, and K. JEFFERTS.

1988. Diversity, distribution and ecology of gonatid

squids in the subarctic Pacific: A review. Bull. Ocean

Res. Inst., University of Tokyo, no. 26 (Part 1),

159-192.

PATTIE, B. H.

1968. Notes on giant squid Moroteuthis robusta (Dall)

Verrill, trawled off the southwest coast of Vancouver

Island, Canada. Wash. Dept. Fish., Fish. Res. Pap.

3 (1) :47-50.
155

PEREYRA, W. T.

1965. New records and observations on the flapjack

devilfish, Opisthoteuthis californiana, Berry. Pac. Sci.

19 :427-441.

ROBSON, G. C.

1932. A monograph of the recent Cephalopoda. Part 2, the

Octopoda. Br. Museum, Natl. Hist., London, 359 p.

ROELEVELD, M. A.

1982. Interpretation of tentacular club structure in

Sthenoteuthis oualaniensis (Lesson 1830) and Ommastrephes

bartramii (Lesueur 1821) (Cephalopoda, Ommastrephidae).

Ann. S. Afr. Mus. 89(4)1982:249-264, 6 Figs.

SASAKI, M.

1929. A monograph of the dibranchiate cephalopods of the

Japanese and adjacent waters. J. Fac. Agri. Hokkaido

Univ., vol. 20 supplementary number, 357 p. plus 30

plates.

SMITH, P. J., R. H. MATTLIN, M. A. ROELEVELD, and T. OKUTANI

1987. Arrow squids of the genus Nototodorus in New Zealand

waters: systematics, biology and fisheries. N. Z. J.

Mar. Freshwater Res., no. 121:315-326.

STROUD, R. K.

1968. Risso dolphin in Washington state. J. Mammal.

49:347-348.
156

SWEENEY, M. J., C. F. E. ROPER, K. M. MANGOLD, M. R. CLARKE, and

S. V. BOLETZKY (editors).

1992. "Larval" and juvenile cephalopods: A manual for

their identification. Smithson. Contrib. Zoology, 513, 282 p.

WING, B., and R. W. MERCER

1990. Temporary northern range extension of the squid

Loliqo opalescens in southeast Alaska. Veliger

33 (3) :238-240.

YOUNG, R. E.

1972. The systematics and areal distribution of pelagic

cephalopods from the seas off southern California.

Smithson, Contrib. Zool. 97, 159 p.




161

APPENDIX A

Checklist of NMML numbers: Cephalopod collection (by family)

Specimens are assigned catalogue numbers as they are acquired by

NMML, consequently consecutive numbering does not hold for each

family. This checklist is a sequential listing of all numbers

used and the family to which the specimen is assigned.

No.

1

2

3

4

5

6

7

8

9

10

11

12

13

14

15

16

17

18

19

20

21

22

23

24

25

26

27

28

29

30

31

32

33

34

35

36

37

Family

Opisthoteuthidae "

"

Octopodidae

Ocythoidae"

"

"

"

"

Octopodidae

Sepiolidae"

"

"

Sepiidae"

"

"

"

Loliginidae "

"

No.

38

39

40

41

42

43

44

45

46

47

48

49

50

51

52

53

54

55

56

57

58

59

60

61

62

63

64

65

66

67

68

69

70

71

72

73

74

Family

Loliginidae "

"

"

"

"

"

"

"

"

"

Bolitaenidae "

Loliginidae

Histioteuthidae "

"

"

"

Enoploteuthidae
162

Cephalopod Collection (Cont. ) .

Family No.

Enoploteuthidae Cranchiidae 122

" 123 "

124

Ommastrephidae 125

" 126 "

" " 127" 128

" 129

Argonautidae 130Ommastrephidae 131

132133134135

136137

, 138

139140141 "

Onychoteuthidae 142

143 "

Octopouthidae 144

, 145 "

Chiroteuthidae 146147 "

" 148

Mastigoteuthidae 149

Chiroteuthidae 150151

152

153154155

Thysanoteuthidae 156

Gonatidae Architeuthidae 157

" 158 ", 159

160

161

162

163164165166167

Cranchildae 168
163

Cephalopod Collection (Cont. ) .

Family No.

Gonatidae Onychoteuthidae 216

" 217

218219220221

222

223

, 224 "

225226

227

Loliginidae 228

229 "230231232 "

233 "234 "

Octopodidae 235

, 236 "

237 "

Sepiolidae 238Octopodidae 239

240 "

Onychoteuthidae 241

Gonatidae 242243 "244 "

Octopodidae 245

246 "

Gonatidae 247248249250251252253

Octopod~dae 254Onychoteuthidae 255

Gonatidae 256

Ommastrephidae 257Onychoteuthidae 258

Octopodidae " 259

" 260 "

Loliginidae 261

Cranchiidae 262
164

Cephalopod Collection (Cont.) .Family No.

Gonatidae 310

Octopodidae 311 "

" 312

313314

Onychoteuthidae Alloposidae 315

Vampyroteuthidae 316 "

" 317 "

Tremoctopodidae 318

" 319" 320

Cranchiidae 321Gonatidae 322

" 323, 324

325326327328

329 "

330331332

Gonatidae 333334 "

335 "

Onychoteuthidae 336

337

" 338

339

" 340

" 341

" 342

343344345 "346347348 "349 "350 "351352353354 "355 "356
165

Cephalopod Collection (Con t .) .

Family Family No.

Gonatidae Onychoteuthidae 404

405

406407408

409

410411

Ommastrephidae 412

" 413

Loliginidae 414Octopodidae 415

" 416

" 417

Onychoteuthidae 418

Gonatidae 419Cranchiidae 420

" 421

422423 "

Bolitaenidae 424

" 425

Gonatidae 426 "Gonatidae" 427

Ommastrephidae " 428

" 429 "

430

Vampyroteuthidae 431Onychoteuthidae " 432

Histioteuthidae 433

Ommastrephidae 434Onychoteuthidae 435 "

" Enoploteuthidae 436

Octopoteuthidae Cranchiidae 437

Gonatidae 438

" 439

440 "

" 441" Histioteuthidae 442

443 "

Chiroteuthidae 444

" 445

Cranchiidae 446

" 447

448 "449 "

450
166

NMML Numbers: Cephalopod Collection (Cont.).

No. Family

451 Vampyroteuthidae

452

Gonatidae

453

"

54 Loliginidae

455

Gonatidae

4576

"

458 Octopodidae 60 459

"

61

"

462

Gonatidae

698456 467 463

Octopodidae

470

"

71 Sepiolidae

472

Gonatidae

473

Loliginidae

474

Octopoteuthidae

475

"

7678

"

79

Histioteuthidae

81 480

Cranchiidae

482

"

8345 Alloposidae

486

Architeuthidae

487

Cranchiidae

488

Bolitaenidae

489

"

90

Ocythoidae

491

Octopoteuthidae

492

"

93 Octopodidae

494

Ommastrephiade

495

"

967

No. Family

498 -

Ommastrephidae

499

"

5001

"

502 Onychoteuthidae

503

"

5045678 50910

Gonatidae

512 511

"

513 Onychoteuthidae

514

"

515678

"

519

Loliginidae

520

"

521

Gonatidae

522

Ommastrephidae

5234

"


169

APPENDIX B

Marine Mammal Predators Noted in Catalogue and Cephalopod Prey

MARINE MAMMAL

Zalophus californianus

Eumetopias jubatus

Callorhinus ursinus

Arctocephalus townsendi

Enhydra lutris

Phoca vitulina

Phoca hispida

Erignathus barbatus

CEPHALOPOD PREY

Octopus rubescens

Octopus bimaculatus

Gonatus sp.

Cranciidae

Loligo opalescens

Abraliopsis sp.

Watasenia scintillans

Onychoteuthis borealiiaponica

Moroteuthis robusta

Gonatus berryi

Gonatus madokai

Gonatus middendorffi

Gonatus onyx

Gonatus pyros

Gonatus sp.

Gonatopsis borealis

Berryteuthis anonychus

Berryteuthis magister

Todarodus pacificus

Chiroteuthis sp.

Octopus sp.

Ocythoe tuberculata

Onychoteuthis banksii

Histioteuthis dofleini

Dosidicus gigas

Eucleoteuthis luminosa

?Pterygioteuthis giardi

Octopus sp.

Octopus dofleini

Octopus sp.

Octopus sp.

Rossia pacifica
170

Marine Mammal Predators Notes in Catalogue and Cephalopod Prey

(Cont.) .

MARINE MAMMAL

Mirounga angustirostris

Pontoporia blainvillei

Stenella qraffmani

Stenella sp.

Delphinus delphis

Laqenorhynchus obliquidens

Grampus griseus

Phocoenoides dalli

CEPHALOPOD PREY

Octopoteuthis deletron

Taningia danae

Gonatus berryi

Gonatus onyx

Gonatus sp. (fabricii type)

Architeuthis japonica

Histioteuthis dofleini

Histioteuthis heteropsis

Chiroteuthis calyx

Cranica scabra

Taonius pavo

Galiteuthis sp.

Vampyroteuthis infernal is

Japetella sp.

Ocythoe tuberculata

Lolliquncula brevis

Abraliopsis affinis

Chiroteuthidae

Onychoteuthis banksii

Cranchia scabra

Gonatus sp. (fabricii type)

Cranchiidae

Octopoteuthis deletron

Chiroteuthis calyx

Watasenia scintillans
171

Marine Mammal Predators Noted in Catalogue and Cephalopod Prey

(Cont. ) .

MARINE MAMMAL

Physeter macrocephalus

CEPHALOPOD PREY

Octopoteuthis deletron

Moroteuthis robusta

Gonatus sp. (fabricii type)

Gonatus sp.

Gonatopsis borealis

Architeuthis japonica

Histioteuthis dofleini

Histioteuthis heteropsis

Mastigoteuthis sp.

Taonius pavo

Meqalocranchia

Galiteuthis sp.

Galiteuthis pacifica?

Mesonychoteuthis sp.

Cranchiidae

Vampyroteuthis internal is

Octopus dofleini

Alloposus mollis



175

APPENDIX C

Reexamination of cephalopod beaks from the stomach of Grampus qriseus MMBL No. 3769.

Published by R.K. Stroud - Risso dolphin in Washington State, J. Mammal. 49(2) 347-348.

Original examination of beaks by Fiscus in 1967 - reexamined 8-11 October 1987.

Table 2. Stroud stomach contents of Grampus qriseus (MMBL No. 3769)

Oriqinal I D 8-11 October 1987 ID

Number

1

1

Onvchoteuthis banksii

Octopodoteuthis sicula

Chiroteuthis veranvi

Gonatus fabricii

Gonatidae - form A 2

Gonatidae - form B 2

Unidentified beaks

16

4

7

13

2

Onvchoteuthis borealiiaponica

Octopoteuthis deletron

In NMML ColI. #145

Chiroteuthis ~

In NMML ColI. #151

Gonatus pvros

Taonius pavo

Taonius pavo

Chiroteuthis ~ (very small)

unidentified (very small)

Number

1 pair beaks

1 pair beaks

I 13 lower beaks

8 upper beaks

4 pair beaks

14 pair ? beaks

1 pair beaks

1 lower beak

Nomenclature change.

On reexamination, difference between A and B attributed to wear on lower beaks.



179

APPENDIX D

Examination of cephalopod beaks from the stomach of a Koqia sp.

Collected 30 December 1969, by R. L. DeLong

Location: stranded on beach at 16-17th Avenue, Santa Cruz, Santa Cruz County, California.

Specimens - no assigned numbers - 1 jar labelled Kogia ColI. R. L. DeLong

Species Number

Beaks

Moroteuthis robusta

Ommastrephes bartrami

Histioteuthis dofleini

Gonatus ~

Gonatus sp.

unidentified beaks

1

1

3

2

2

lower beak

pair beaks

lower beaks

pair ? beaks

lower beak fragments

LRL 13.5 rom.

LRL 13.0 rom.

upper beaks (Histioteuthis/Cranchiid) 4

1 fragment

Moroteuthis ? 1 fragment

Note. All beaks much darkened and worn from long stay in stomach.
180

Examination of cephalopod beaks from the stomach of a Koqia breviceps.

Collected 5 October 1977, by J. Rozdilsky

Location: stranded, freshly dead, on beach 13 km north of Oak Harbor, Whidbey Island,

puget Sound, Washington. Lat. 48°38'W. Long. 122°17'W. Skeletal remains in

Burke Museum, University of Washington, Seattle.

Cephalopod specimens CC-6: 11 vials, NMML voucher collection.

Original examination of beaks by Fiscus, 4 October 1978, reexamined 7 July 1992.

Original I D 7 Jul,y,1992

Number Number

Unidentified beaks

Gonatus sp.? lower beaks

upper beaks

Unident. beaks small & fragments

lower beaks

upper beaks

1st Chamber stomach

Octopoteuthis sp.

2nd Chamber stomach

Histioteuthis sp. lower beaks

upper beaks

Octopoteuthis sp. lower beaks

upper beaks

Gonatidae

most likely Gonatopsis lower bks

borealis upper bks

Onvchoteuthis borealiiaponicus

lower beaks

upper beaks

Octo£oteuthis deletron?

6 upper beaks

5 upper beaks

3 upper beaks

1 upper beak

1 upper beak

1 upper beak

1 upper beak

5 lower beaks

6 lower beaks

2 upper beaks

7 lower beaks

5 lower beaks

1 lower beak

1 lower beak

17 lower beaks

15 upper beaks

17 lower beaks

22 upper beaks

Octo£oteuthis deletron

Onvchoteuthis

boreali~onica 2

includes

Histioteuthis sp.

Taonius sp.

Galiteuthis sp.

Taonius pavo

Galiteuthis sp.

Gonatus sp. (fab. type) 3

Gonatidae

Unidentified beaks

Types 1

2

3

4

5

6

Gon/Berrv 1

Histioteuthis sp.

1

5

6

1

4

1

14

2

18

13

4

8

17

22

beaks pair

Cranchidae

upper beaks

lower beaks

upper beaks

3rd Chamber stomach

Octopoteuthis sp.

Histioteuthis sp.

unidentified

2

1

1

Octopoteuthis deletron

Histioteuthis dofleini

Gonatidae

2 upper beaks

1 lower beak

1 upper beak

I Gon/Berrv - Gonatopsis borealis or Berrvteuthis maqister.

2 Nomenclature change.

3 See Note ~ Catalogue of Cephalopods.
181

Examination of cephalopod beaks from the stomach of a Koqia.

Collected January 1966 by W. J. Houck

Location: stranded on beach in Big Lagoon, Arcata, California. Skeletal remains in

Museum, Humboldt State College, Arcata, California.

Cephalopod specimens CC-8:- 6 vials, NMML voucher collection.

Original examination by Fiscus, 8 August 1966, reexamined 7 July 1992.

Oriqinal I D 7 Julv 1992Onvchoteuthis boreali- 3 lower beaks 2 lower beaks

iaponica 1

1 1 upper beak

1 lower beak

Chiroteuthis ~ 1 1 lower beak 1

Octopoteuthis deletron5 lower beaks I 5

3 3 upper beaksGonatus madokai 2 ?lower beaks 2 lower beaks

1 1 upper beak

upper beak Gonatidae 1 upper beak
182

Cephalopod specimens CC-7 - 1 vial from stomach of Delphinus delphis.

Collected 1968 by W. J. Houck, Humboldt State University, Arcata, California.

Oriqinal I D 7 Julv 1992

lower beaks Octopoteuthis deletron 3 lower beaks 3

4 3 upper beaks

lower beak ? Onychoteuthis 1

1 upper beak

lower beaks Taonius pavo 1 lower beak 32

Gonatus berrvi

lower beaks3 ?

2 upper beaks unidentified beaks

Cephalopod specimens CC-I0 - 3 vials from stomach of Mirounqa anqustirostris

Collected 9 July 1976 by R. K. Stroud, Oregon State University, Corvallis, Oregon.

Location: stranded, Fort Stevens State Park, Oregon. Stroud MM 76-13.

Oriqinal I D 8 July 1992

1

7 pairs beaks Histioteuthis dofleini 7 pairs beaks

Onychoteuthis 1 pair beaks 1 pir beaks

borealiiaponica

Octopoteuthis deletron lower beaks 10 9 lower beaks

7 7 upper beaks

vial No. 4 not found 1 lower beak

vial No.4 not found 6 lower beaks

vial No. 4 not found 4

1 lower beak

vial No. lower beaks 13

vial No. 7 upper beaks

Cephalopod specimens CC-ll - 1 vial from stomach of Delphinus delphis male.

Collected 6 March 1976 by R. K. Stroud, Oregon State University, Corvallis, Oregon.

Location: stranded, 1/4 mile north of Netarts Bay, Oregon. Stroud MM 76-3.

Oriqinal I D 8 July 1992

1 upper beak

Abraliopsis sp.1 upper beak lower beak

4 pairs ? 1 lower beak4 lower beaks

3 upper beaks

1 upper beak
183

Cephalopod specimens CC-12 - 1 vial from stomach of Mirounqa anqustirostris.

Collected 28 February 1975 by R. K. Stroud, Oregon State University, Corvallis, Oregon.

Location: stranded, Depoe Bay, Oregon. Stroud MM 75-2.

8 July 1992

Gonatus ~ ?

Cephalopod specimens CC-13 - 1 vial from stomach of Delphinus delphis.

1st examination

1 lower beak

2 upper beaks

Collected January 1975 by R. K. Stroud, Oregon State University, Corvallis, Oregon.

Location: stranded, Neskowin, Oregon. Stroud MM 74-19.

8 Julv 1992 1st examiniation

LoligQ opalescens

1 lower beak

Onvchoteuthis 13 upper beaks

10 lower beaks

borealiiaponica

10 upper beaks

Gonatus ~

3 upper beaks

3 upper beaks ? Gon/Berrv

1 lower beak

Histioteuthis sp.

3 lower beaks

2 upper beaks unidentified beaks Taonius pavo

5 upper beaks

Cephalopod specimens CC-15 - 1 vial from stomach of Phvseter macrocephalus.

Collected 19 August 1959, collector unknown.

Location: taken off west coast of Vancouver Island, Canada.

8 Julv 1992 1st examination

Onvchoteuthis

6 lower beaks

borealiiaponica

3 upper beaks

Gonatus sp.

1 lower beak

Chiroteuthis ~

4 lower beaks

Taonius pavo

15 lower beaks

Galliteuthis sp.

1 lower beak

unidentified beaks

12 upper beaks

RECENT TECHNICAL MEMORANDUMS

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5 GUTTORMSEN, M., R. NARITA, J. GHARRETT, G. TROMBLE, and J. BERGER. 1992. Summary

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4 GUTTORMSEN, M., R. NARITA, and J. BERGER. 1992. Summary of U. S. observer sampling

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PB93-127546.

3 JOHNSON, P. A., S. D. RICE, and M. M. BABCOCK (compilers). 1992. Impacts of oil pollution

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2 KAJIMURA, H., and E. SINCLAIR. 1992. Fur seal investigations, 1990, 192 p. NTIS No. PB93-

109080.