U.S. Department of the Interior

U.S. Geological Survey

Open-File Report 2018–1112

Prepared for Commander Navy Region Southwest

Monitoring Breeding and Migration of Neotropical 

Migratory Birds at Naval Base Coronado, Remote 

Training Site, Warner Springs, San Diego County, 

California, 5-Year Summary 2013–17
Cover:

Top row: Photographs showing black-throated gray warbler (Setophaga nigrescens) by 

A. Houston, Bullock’s oriole (Icterus bullockii) by K. Hall, white-breasted nuthatch (Sitta 

carolinensis) by S. Howell.

Bottom row: Photographs showing red-breasted sapsucker (Sphyrapicus ruber) by S. Howell,

fox sparrow (Passerella iliaca) by S. Howell, and western kingbird (Tyrannus verticalis) by 

K. Hall. 
Monitoring Breeding and Migration of Neotropical 

Migratory Birds at Naval Base Coronado, Remote 

Training Site, Warner Springs, San Diego County, 

California, 5-Year Summary, 2013–17 

By Suellen Lynn, Katie A. Hall, Melanie C. Madden, and Barbara E. Kus

Prepared for Commander Navy Region Southwest

Open-File Report 2018–1112

U.S. Department of the Interior

U.S. Geological Survey
U.S. Department of the Interior

RYAN K. ZINKE, Secretary

U.S. Geological Survey

James F. Reilly II, Director

U.S. Geological Survey, Reston, Virginia: 2018 

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Suggested citation:

Lynn, S., Hall, K.A., Madden, M.C., and Kus, B.E., 2018, Monitoring breeding and migration of neotropical 

migratory birds at Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 

5-year summary, 2013–17: U.S. Geological Survey Open-File Report 2018–1112, 98 p., 

https://doi.org/10.3133/ofr20181112.

ISSN 2331-1258 (online)
iii

Contents

Executive Summary ................................................................................................................................................. 1

Introduction .............................................................................................................................................................. 3

Methods ................................................................................................................................................................... 4

Bird Banding......................................................................................................................................................... 4

Banding Schedule ................................................................................................................................................ 8

Data Analysis........................................................................................................................................................ 8

Results...................................................................................................................................................................10

Monitoring Avian Productivity and Survivorship..................................................................................................10

Overview of Captures.....................................................................................................................................10

Capture Rates ................................................................................................................................................10

Species Richness...........................................................................................................................................10

Relative Species Abundance .........................................................................................................................11

Age and Sex Structure ...................................................................................................................................11

Productivity.....................................................................................................................................................11

Survivorship ...................................................................................................................................................11

Population Trends ..........................................................................................................................................12

Fall Migration......................................................................................................................................................12

Overview of Captures.....................................................................................................................................12

Capture Rates ................................................................................................................................................12

Species Richness...........................................................................................................................................12

Relative Species Abundance .........................................................................................................................13

Age and Sex Structure ...................................................................................................................................13

Arrival Dates...................................................................................................................................................13

Discussion..............................................................................................................................................................14

References Cited ...................................................................................................................................................17

Figures

Figure 1. Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, 

California (U.S. Navy, 2013) ....................................................................................................................................5

Figure 2. Location of bird banding station, Naval Base Coronado, Remote Training Site, Warner Springs, 

San Diego County, California...................................................................................................................................6

Figure 3. Number of individuals captured per species during Monitoring Avian Productivity and Survivorship, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2013....................63

Figure 4. Number of individuals captured per species during Monitoring Avian Productivity and Survivorship, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2014....................63

Figure 5. Number of individuals captured per species during Monitoring Avian Productivity and Survivorship,

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2015....................64

Figure 6. Number of individuals captured per species during Monitoring Avian Productivity and Survivorship, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2016....................64

Figure 7. Number of individuals captured per species during Monitoring Avian Productivity and Survivorship, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2017....................65

Figure 8. Annual productivity for the ten most commonly captured resident species during Monitoring 

Avian Productivity and Survivorship, Naval Base Coronado, Remote Training Site, Warner Springs, 

San Diego County, California, 2013–17.................................................................................................................66
iv

Figure 9. Adult survivorship calculated with Program MARK for four commonly captured resident 

species during Monitoring Avian Productivity and Survivorship, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California, 2013–17...................................................67

Figure 10. Adult annual capture rate (individuals captured per 100 net-hours) of the 11 most commonly 

captured resident species during Monitoring Avian Productivity and Survivorship, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California, 2013–17...................................................68

Figure 11. Number of individual migrants captured per species during fall migration, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California, 2013.........................................................95

Figure 12. Number of individual migrants captured per species during fall migration, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California, 2014 .........................................................95

Figure 13. Number of individual migrants captured per species during fall migration, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California, 2015.........................................................96

Figure 14. Number of individual migrants captured per species during fall migration, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California, 2016.........................................................96

Figure 15. Number of new migrant species arriving per banding day during fall migration, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2013....................97

Figure 16. Number of new migrant species arriving per banding day during fall migration, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2014....................97

Figure 17. Number of new migrant species arriving per banding day during fall migration, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2015....................98

Figure 18. Number of new migrant species arriving per banding day during fall migration, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2016....................98

Tables 

Table 1. Global Positioning System locations of mist-nets at the bird banding station, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California.....................................................................7

Table 2. Species captured at Naval Base Coronado, Remote Training Site, Warner Springs, 

San Diego County, California, 2013–17.................................................................................................................20

Table 3. Species observed but not captured at Naval Base Coronado, Remote Training Site, Warner Springs, 

San Diego County, California, 2013–17.................................................................................................................22

Table 4. Number of birds captured and banded, during Monitoring Avian Productivity and Survivorship, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, 2013–17...............................24

Table 5. Number of birds recaptured during Monitoring Avian Productivity and Survivorship, Warner Springs, 

San Diego County, California, 2013–17.................................................................................................................27

Table 6. Capture frequency of individuals during Monitoring Avian Productivity and Survivorship, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2013....................28

Table 7. Capture frequency of individuals during Monitoring Avian Productivity and Survivorship, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2014....................29

Table 8. Capture frequency of individuals during Monitoring Avian Productivity and Survivorship, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2015....................31

Table 9. Capture frequency of individuals during Monitoring Avian Productivity and Survivorship, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2016....................32

Table 10. Capture frequency of individuals during Monitoring Avian Productivity and Survivorship, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2017....................33

Table 11. Capture rate by net and date during Monitoring Avian Productivity and Survivorship, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2013....................34

Table 12. Capture rate by net and date during Monitoring Avian Productivity and Survivorship, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2014....................36
v 

Table 13. Capture rate by net and date during Monitoring Avian Productivity and Survivorship, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2015....................38

Table 14. Capture rate by net and date during Monitoring Avian Productivity and Survivorship, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2016....................40

Table 15. Capture rate by net and date during Monitoring Avian Productivity and Survivorship, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2017....................42

Table 16. Number of captures by date during Monitoring Avian Productivity and Survivorship, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2013....................44

Table 17. Number of captures by date during Monitoring Avian Productivity and Survivorship, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2014....................46

Table 18. Number of captures by date during Monitoring Avian Productivity and Survivorship, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2015....................48

Table 19. Number of captures by date during Monitoring Avian Productivity and Survivorship, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2016....................50

Table 20. Number of captures by date during Monitoring Avian Productivity and Survivorship, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2017....................51

Table 21. Sex and age of individuals (banded and unbanded) captured during Monitoring Avian 

Productivity and Survivorship, Naval Base Coronado, Remote Training Site, Warner Springs, 

San Diego County, California, 2013.......................................................................................................................52

Table 22. Sex and age of individuals (banded and unbanded) captured during Monitoring Avian 

Productivity and Survivorship, Naval Base Coronado, Remote Training Site, Warner Springs, 

San Diego County, California, 2014.......................................................................................................................54

Table 23. Sex and age of individuals (banded and unbanded) captured during Monitoring Avian 

Productivity and Survivorship, Naval Base Coronado, Remote Training Site, Warner Springs, 

San Diego County, California, 2015.......................................................................................................................56

Table 24. Sex and age of individuals (banded and unbanded) captured during Monitoring Avian 

Productivity and Survivorship, Naval Base Coronado, Remote Training Site, Warner Springs, 

San Diego County, California, 2016.......................................................................................................................58

Table 25. Sex and age of individuals (banded and unbanded) captured during Monitoring Avian 

Productivity and Survivorship, Warner Springs, San Diego County, California, 2017 ............................................60

Table 26. Logistic regression models for the effect of year on survivorship for four resident bird species 

captured during Monitoring Avian Productivity and Survivorship, Naval Base Coronado, Remote Training Site, 

Warner Springs, San Diego County, California 2013–17 .......................................................................................62

Table 27. Number of migrant birds captured and banded during fall migration, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California, 2013–16...................................................69

Table 28. Capture frequency of migrant individuals during fall migration, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California, 2013.........................................................71

Table 29. Capture frequency of migrant individuals during fall migration, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California, 2014.........................................................72

Table 30. Capture frequency of migrant individuals during fall migration, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California, 2015.........................................................73

Table 31. Capture frequency of individual migrants during fall migration, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California, 2016.........................................................74

Table 32. Capture rate by net and date during fall migration, Naval Base Coronado, Remote Training Site, 

Warner Springs, San Diego County, California, 2013 ............................................................................................75

Table 33. Capture rate by net and date during fall migration, Naval Base Coronado, Remote Training Site, 

Warner Springs, San Diego County, California, 2014 ............................................................................................77
vi 

Table 34. Capture rate of migrant species by net and date during fall migration, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California, 2015.........................................................79

Table 35. Capture rate of migrant species by net and date during fall migration, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California, 2016.........................................................81

Table 36. Number of migrant species captures by date during fall migration, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California, 2013.........................................................83

Table 37. Number of migrant captures by date during fall migration, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California, 2014.........................................................85

Table 38. Number of migrant species captures by date during fall migration, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California, 2015.........................................................87

Table 39. Number of migrant captures by date during fall migration, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California, 2016.........................................................89

Table 40. Sex and age of individuals (banded and unbanded) captured during fall migration, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2013....................90

Table 41. Sex and age of migrant individuals (banded and unbanded) captured during fall migration, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2014....................91

Table 42. Sex and age of migrant individuals (banded and unbanded) captured during fall migration, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2015....................92

Table 43. Sex and age of migrant individuals (banded and unbanded) captured during fall migration, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2016....................93

Table 44. First and median capture dates for each migrant species during fall migration, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2013–16..............94

Conversion Factors

International System of Units to Inch/Pound

Multiply By To obtain

Length

millimeter (mm) 0.03937 inch (in.)

meter (m) 3.281 foot (ft) 

kilometer (km) 0.6214 mile (mi)

kilometer (km) 0.5400 mile, nautical (nmi) 

meter (m) 1.094 yard (yd) 

Area

hectare (ha) 2.471 acre

Datum

Horizontal coordinate information is referenced to the World Geographic System of 1984 (WGS 84).
vii 

Abbreviations

MAPS Monitoring Avian Productivity and Survivorship

IBP Institute for Bird Populations

PDT Pacific Daylight Time

PST Pacific Standard Time

RTSWS Remote Training Site, Warner Springs
viii
1 

Monitoring Breeding and Migration of Neotropical 

Migratory Birds at Naval Base Coronado, Remote 

Training Site, Warner Springs, San Diego County, 

California, 5-Year Summary, 2013–17 

By Suellen Lynn, Katie A. Hall, Melanie Madden, and Barbara E. Kus

Executive Summary

We operated a bird banding station on the Naval Base Coronado, Remote Training Site, 

Warner Springs (RTSWS), in northeastern San Diego County, California, during the bird breeding 

season (spring/summer) from 2013 to 2017 and during migration (fall) from 2013 to 2016. The station 

was established in spring 2013 as part of the Monitoring Avian Productivity and Survivorship (MAPS) 

program and continued into the fall for the first 4 years as part of a long-term monitoring program for 

neotropical migratory birds. 

We captured 705 individuals of 58 species during the MAPS/breeding season from 2013 to 

2017 (12–13 days each year in April through August), 79 percent of which were newly banded during 

the MAPS season (555), 8 percent of which were recaptures banded in previous years (57), and 13 

percent of which we released unbanded (64 hummingbirds and 29 other birds that were released or 

escaped prior to banding). Sixty individuals were captured more than once within a year during 

MAPS. Bird capture rate averaged 19 ± 1 captures per 100 net-hours (range 17–20) across 5 years. 

Annual species richness ranged from 28 (2017) to 42 (2014). The average species richness per day was 

highest in 2014 (9 ± 3) and lowest in 2016 (6 ± 2). Bushtit (Psaltriparus minimus) was the most 

abundant breeding species captured, followed by Spotted Towhee (Pipilo maculatus), Oak Titmouse 

(Baeolophus inornatus), Anna’s Hummingbird (Calypte anna), House Wren (Troglodytes aedon), 

Ash-throated Flycatcher (Myiarchus cinerascens), California Scrub-jay (Aphelocoma californica), 

Bewick’s Wren (Thryomanes bewickii), Acorn Woodpecker (Melanerpes formicivorus), California 

Towhee (Melozone crissalis), and Western Bluebird (Sialia mexicana). Each of these 11 breeding 

species accounted for at least 5 percent of captures in any 1 year. Fifty-seven percent of known-sex 

captures were female and 43 percent were male. Thirty-three percent of known-age captures were 

juveniles. Peaks in number of birds captured were in the first and last weeks of April, and the greatest 

number of species was captured in early May. 
2 

We were able to calculate productivity for 10 species (of the 11 most abundant species) with 

sufficient captures and recaptures for valid parameter estimation. Of these 10 species, Bushtit had the 

highest overall productivity as measured by the ratio of juveniles to adults captured, followed by 

Bewick’s Wren and Anna’s Hummingbird. There were no clear patterns in annual productivity among 

species. Across all 10 species, average productivity was highest in 2017 and lowest in 2014. Spotted 

Towhee, California Scrub-jay, Acorn Woodpecker, and Ash-throated Flycatcher all had low 

productivity overall. We had sufficient recaptures to analyze survivorship for four species: Bushtit, 

Spotted Towhee, Oak Titmouse, and House Wren. Bushtits had the highest survivorship rates, 

followed by Spotted Towhees, Oak Titmice, and House Wrens. Captures of Acorn Woodpeckers, 

California Scrub-jays, Oak Titmice, Spotted Towhees, and California Towhees decreased significantly 

and Bushtit captures increased significantly over 5 years. 

During fall migration (16–19 days in August through October), we captured 595 individual 

migratory birds of 44 species, 94 percent of which were newly banded in fall (558), 4 percent of which 

were recaptures banded in previous years (22), and 3 percent of which we released unbanded (11 

hummingbirds and 4 other birds that were released or escaped prior to banding). Twenty-seven 

individuals were captured more than once within a year in fall. Migratory bird capture rate averaged 22 

± 8 per 100 net-hours (range 12–30) and was highest in 2013 and lowest in 2016. Annual species 

richness during fall migration ranged from 23 (2016) to 30 (2015). Species richness averaged 5 ± 3 

species per day and was highest in 2013 (6 ± 2) and lowest in 2016 (4 ± 2). Spotted Towhees were the 

most abundant fall migrant captured, followed by Wilson’s Warblers (Cardellina pusilla), Yellow 

Warblers (Setophaga petechia), Orange-crowned Warblers (Oreothlypis celata), White-crowned 

Sparrows (Zonotrichia leucophrys), and Pacific-slope Flycatchers (Empidonax difficilis). Forty-two 

percent of known-sex individuals were female and 58 percent were male. Forty-nine percent of 

known-age captures were juveniles. The peak number of new species arrived during the fourth week of 

fall banding, early to mid-September. The highest number of birds and the greatest number of species 

were captured in September. 

First capture dates for eight migratory species were earlier each year they were captured from 

2013 to 2016; two species (Costa’s Hummingbird, Calypte costae, and Ruby-crowned Kinglet, 

Regulus calendula) had correspondingly earlier median capture dates each year. First capture dates for 

five migratory species were later each year they were captured; two species (Phainopepla, Phainopepla 

nitens, and Fox Sparrow, Passerella iliaca) had correspondingly later median capture dates each year.
3 

Introduction

Monitoring Avian Productivity and Survivorship (MAPS; administered by the Institute for Bird 

Populations [IBP], Point Reyes Station, California) is an international monitoring program that uses 

bird capture and banding data to compile basic demographic parameters of resident and migratory 

species, many of which are imperiled regionally and even globally. Age- and sex-specific data on 

annual survival, reproduction, and recruitment can be gathered and compared across stations to 

identify population trends for species of interest and can be used to identify proximate factors 

responsible for trends, specifically negative trends. In turn, information obtained from long-term 

monitoring of bird populations can be used to guide management activities intended to maintain or 

reestablish viable populations throughout the species’ ranges. 

We established a MAPS and migration monitoring station on the Naval Base Coronado, 

Remote Training Site, Warner Springs (RTSWS), in northeastern San Diego County, California, in 

2013. This station was established as part of a long-term monitoring program for neotropical migratory 

birds for Naval Base Coronado and was operated in a manner consistent with other banding stations 

participating in an effort to monitor birds worldwide. 

Understanding the factors that affect migratory bird populations during all phases of their life 

cycle is critical to species and ecosystem conservation planning. Bird migration studies are useful 

because they apply to species that occupy sites across a broad geographic range, and potentially a 

broad range of habitat types, during the phases of their life cycles. Because migration provides a link 

between wintering and breeding phases of the annual cycle, migration studies can elucidate the 

responses of species to diverse pressures affecting any of the phases of their annual cycles, including 

habitat alteration and loss, diseases, environmental contaminants, and global climate change. 

Studying birds during migration can also inform land managers about local factors that affect 

migratory stopover habitat. High-quality stopover habitat provides abundant resources that allow 

individuals to reestablish energy reserves within the short timeframe available for completing

migration (Skagen and others, 2004). Identifying these stopover sites is the first step in conserving and 

potentially enhancing habitats that benefit bird populations. 

This report summarizes 5 years of station operation from 2013 to 2017. There are five 

objectives for this project: (1) to calculate capture rates, relative abundance, and trends of various 

neotropical migratory bird species populations; (2) to estimate survivorship parameters for neotropical 

migratory bird species; (3) to estimate annual productivity for these species; (4) to augment existing 

distributional information for “sensitive” avian species; and (5) to document the demographics of 

neotropical migratory birds passing through the site during their fall southern migration to wintering 

grounds. 

This study was funded by Commander Navy Region Southwest, San Diego, California. 
4 

Methods

Bird Banding

We established the Warner Springs bird banding station at Naval Base Coronado, RTSWS, in 

spring 2013. The base includes approximately 5,052 hectares (ha) on predominantly undeveloped land 

leased from Cleveland National Forest, Vista Irrigation District, and the Bureau of Land Management 

(fig. 1). Vegetation at the station is a mix of cottonwood- (Populus fremontii) dominated riparian 

forest, big sagebrush (Artemisia tridentata) scrub, and oak (Quercus sp.) woodland. Typical woody 

species include cottonwood, live oak (Q. agrifolia), mule fat (Baccharis salicifolia), willow (Salix sp.),

and big sagebrush. 

Bird banding at the Warner Springs banding station followed standardized MAPS protocol 

(DeSante and others, 1993) with modifications for migration (see below). Ten nets were established at 

the station in April 2013, but because bird capture rates were low, we added three more nets in June 

2013 and one additional net in August 2013. The 14 mist-nets were placed 32–550 meters (m) apart 

and were erected in fixed locations chosen for their potential to capture birds moving through 

vegetation (fig. 2, table 1). Mist-nets were made of 30-millimeter (mm) black nylon mesh and were 12

m long by 2.6 m high with four trammels (“pockets”) running the length of the net. Nets were 

suspended from vertical aluminum poles anchored by permanent rebar stakes and covered a vertical 

area ranging from approximately 0.25 to 2.5 m above the ground. During MAPS, nets were opened 

within 30 minutes of dawn and remained open for 5 hours, typically closing between 1100 and 1200 

Pacific Daylight Time (PDT). During migration, nets were opened within 60 minutes of dawn and 

remained open for 3 hours, typically closing between 0900 and 1000 Pacific Standard Time (PST). 

Nets were not operated during inclement weather such as strong wind, rain, extreme heat, or extreme 

cold. Dates of net operation are detailed in the following section. 
5 

Source: Bing Maps Hybrid, © 2010 Microsoft Corporation and its data suppliers, Map contains the most current data to date which may change, and is compiled from a variety of references 

(See GIS Reference List in original report: Arnold 2017)

Figure 1. Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California (U.S. Navy, 2013). Pink star indicates location of 

banding station.
6 

Figure 2. Location of bird banding station, Naval Base Coronado, Remote Training Site, Warner Springs, San 

Diego County, California.
7 

Table 1. Global Positioning System locations of mist-nets at the bird banding station, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California. 

[Coordinates are in World Geodetic System of 1984 (WGS 84)]

Net location No. 

(see fig. 2) Longitude Latitude

1 −116.687782 33.310240

2 −116.687919 33.310646

3 −116.687600 33.311080

4 −116.686703 33.311412

5 −116.686925 33.311980

6 −116.686586 33.312421

7 −116.686953 33.313137

8 −116.686934 33.313416

9 −116.686359 33.314371

10 −116.687197 33.314280

11 −116.686138 33.314665

12 −116.686209 33.315095

13 −116.687447 33.312290

14 −116.688311 33.311221

Nets were checked every 20–40 minutes by observers working circuits. Hummingbirds, upland 

game birds, and raptors were not banded but were identified to species, age, and sex, when possible, 

and released immediately at the capture site. Hummingbirds were captured frequently at this station, 

and in 2014, we started clipping the right fourth rectrix of all hummingbirds captured as a way to 

temporarily mark them and help determine how many individual hummingbirds were being captured 

per year. All other birds were removed from nets, held in cloth bags labeled with the net number, and 

taken to a central processing location where they were banded with federal numbered aluminum bands. 

Data recorded for each individual caught included age, sex, skull pneumatization, breeding condition, 

weight, wing chord, fat deposition, feather wear, and molt status, using Pyle (1997) as a reference.

Birds were held for 5–60 minutes, depending on the number of birds captured during one net run. 

Birds showing signs of stress were prioritized for processing and were released with a minimum of 

data collected (band number, species, and if possible, age and sex). After processing, juveniles, 

brooding females, and resident birds from the more distant nets were released near the net in which 

they had been captured, and all other birds were released at the processing station. All species captured 

and (or) observed at the site were recorded in a daily list, along with any observed breeding behavior, 

to help determine breeding status and to document species that were present at the station but not 

captured. Breeding status was determined based on the number and timing of detections and whether 

breeding behavior was observed (for example, singing, nest building, carrying food, feeding young). 
8 

Typically, three to four personnel operated the station. Fieldwork was conducted by the 

following U.S. Geological Survey employees: Lisa Allen, Thomas Dayton, Patience Falatek, Jonathan 

Gunther, Katherine Hall, Sarah Harris, Alex Houston, Scarlett Howell, Jennifer Jacobs, Angela 

Johnson, Michael Lester, Suellen Lynn, Melanie Madden, Brandon Miller, Austin Parker, Ryan 

Pottinger, Devin Taylor, Michelle Treadwell, Charlie Vettes, Anne Winters, and Benjamin Zyla. The 

following volunteers provided assistance: Armando Aispuro, Arlene Arnold (U.S. Navy), Tiffany 

Shepherd (U.S. Navy), Erica Harris, Kim Roth, Jessica Looney, and Cassie Swindle. 

Banding Schedule

Because we were interested in examining both breeding and migratory activity at the Warner 

Springs Banding Station, we operated mist-nets during two seasons: (1) MAPS (breeding, 2013–17) 

and (2) fall (migration, 2013–16). 

During MAPS, the banding station was operated 1 day during every 10-day period between 

April 1 and August 9, following standard MAPS protocol (13 days each year, except for 2016 when 

weather limited banding operation to 12 days). In fall, the station was typically operated 2–3 days per 

week from August 15 through October 15 (16–19 days per year). Military training usually occurred at 

the site from Friday afternoon through Monday morning, which restricted our access for fall banding 

to Tuesday through Friday mornings. There were a few weeks when additional military activity or 

inclement weather prevented or further restricted our access. In 2013, our fall banding schedule was 

interrupted by the federal government furlough, and no banding occurred between September 26 and 

October 23, our last banding day. Our fall banding schedule was dynamic, in an attempt to maximize 

the capture of neotropical migrants while keeping project costs to a minimum. We began our week of 

banding on Tuesdays. If the capture rate for migrants was high on Tuesday (at least eight migrants), 

we continued banding on Wednesday; if capture rate was low, we resumed banding on Thursday. We 

followed this pattern for every day of banding, which meant we banded for a minimum of 2 days 

(either Tuesday/Wednesday or Tuesday/Thursday) and a maximum of 3 days 

(Tuesday/Wednesday/Thursday or Tuesday/Thursday/Friday) each week, weather and military training 

schedules permitting. 

Data Analysis

All banding data were entered into MAPSPROG, the IBP data entry program, for verification 

and error checking, which included cross-checking against previous years’ data. Finalized 

MAPSPROG data were submitted to IBP and analyzed as described below. 

Bird captures were quantified by species, age, sex, and number of captures for each year. We 

used the total number of captures (including newly banded birds, birds that were captured but not 

banded, and previously banded birds, or “recaptures”) to create a list of captured species and a total 

count of captures per species for the site. We intentionally did not band hummingbirds because they 

are difficult to band and recapture and would require a concentrated effort beyond the scope of a 

MAPS station. However, as described above, we trimmed a small section of the right fourth rectrix of 

each captured hummingbird upon initial capture to ascertain if an individual was new or a recapture 

within the same year. The total number of individuals captured in each year included only newly 

banded birds, first time recaptures of birds banded at the sites in previous years, and birds that were not 

banded. 
9 

We calculated annual capture rates for each species by dividing the total number of individuals 

captured (described above) by the total number of hours nets were open at the banding station (nethours) and multiplying this number by 100. Captures per 100 net-hours is a standard unit of 

measurement used for bird banding analyses. Calculating capture rates in this manner allows us to 

account for the discrepancy in the number of banding hours each day and to make between-year 

comparisons. 

Additionally, we used the total number of individuals to calculate relative species abundance

(the proportion of all individuals represented by a particular species), sex and age ratios (to determine 

the structure of breeding and migrating populations), productivity (see below), adult survivorship (see 

below), and first and median arrival dates for each species during migration. First and median arrival 

dates were intended to track potential changes in the timing of migration associated with climate 

change. 

During the fall, we restricted our analyses to neotropical migratory birds (species that were not 

resident breeders at Warner Springs), with the exception of two species (see below). Neotropical 

migratory species were defined as those covered under the Migratory Bird Treaty Act of 1918 (U.S. 

Fish and Wildlife Service, 1998). Migratory birds were then categorized as resident or migrant, 

according to behavior, breeding condition, and breeding status records for the site (not for the region as 

a whole). Based on this categorization, neotropical migratory species that were captured during 

migration and were not known to breed at the site, such as Orange-crowned Warbler (Oreothlypis 

celata), were considered migrants even though they may breed nearby and be year-round residents for 

the region. Only non-breeding, neotropical migratory species were included in our analyses of fall

migration, except for Lark Sparrow (Chondestes grammacus) and Spotted Towhee (Pipilo maculatus), 

two neotropical migratory species that were known to breed at this site but were captured in much 

higher numbers during fall banding than during MAPS banding. The rationale for including these 

breeding species was to evaluate the importance of this site for them in the fall, possibly for postbreeding dispersal or as a stopover location for migrating individuals, and to examine whether this site 

may be more important to these species in the fall than during the breeding season. 

We calculated indices of productivity and survivorship for each year for a subset of bird 

species with sufficient captures and recaptures for valid parameter estimation. For productivity, we 

analyzed data collected for Anna’s Hummingbird (Calypte anna), Acorn Woodpecker (Melanerpes 

formicivorus), Ash-throated Flycatcher (Myiarchus cinerascens), California Scrub-jay (Aphelocoma 

californica), Oak Titmouse, (Baeolophus inornatus), Bushtit (Psaltriparus minimus), Bewick’s Wren 

(Thryomanes bewickii), House Wren (Troglodytes aedon), Spotted Towhee, and California Towhee 

(Melozone crissalis). We only had sufficient recaptures to analyze survivorship for Bushtit, House 

Wren, Oak Titmouse, and Spotted Towhee. Productivity was calculated as the number of juvenile 

captures (hatching-year) divided by the number of adult captures (after-hatching-year) to control for 

fluctuations in adult population size. 

We calculated annual survivorship in Program MARK (White and Burnham, 1999). We 

estimated only adult survivorship because the number of first-year captures was not large enough for 

meaningful analysis. For MARK analyses, we created encounter histories for each of the selected 

species based on whether or not an individual was captured at the station within a particular year. 

Multiple captures of individuals within a year were counted as a single encounter. We assumed 

constant recapture probability because we expended a constant effort each year. 

We used an information-theoretic approach (Akaike’s information criteria, or AIC; Burnham 

and Anderson, 2002) to evaluate support for models that tested the effect of year on survivorship. We 

used logistic regression with a logit link to build models. First, we generated a constant survival model 

to serve as a reference. We then modeled year (annual variation) and evaluated support for the model 

in relation to the constant survival model. Criteria for better supported models included lower AIC 

than other models, ∆AIC < 2, and AIC weight > 0.05.
10 

Results

Monitoring Avian Productivity and Survivorship

Overview of Captures

We had a total of 783 captures of 58 species in 4,224:10 net-hours during the 2013–17 

breeding seasons (tables 2, 4, 11–15 at back of report). Of these, 555 were newly banded during the 

MAPS season, 57 were recaptures originally banded in earlier years (table 4), and 93 were released 

unbanded (64 hummingbirds and 29 additional birds that escaped prior to banding or were released 

without banding; tables 6–10 at back of report), for a total of 705 individuals captured. Ninety percent 

of banded individuals (552/612; 50 species) were captured only once within a year during the MAPS 

season, 8 percent (50/612; 16 species) were captured twice, 1 percent (6/612; 4 species) were captured 

three times, and <1 percent (4/612; 2 species) were captured four times (tables 6–10). One 

hummingbird with a clipped tail feather was recaptured once during the breeding season. The total 

number of banded individuals per capture incidence does not include three banded individuals that 

were recaptured and escaped prior to recording the band number, thus making it impossible to 

determine whether these individuals had been captured multiple times that year. However, it does 

include one banded Spotted Towhee that was recaptured in 2016 and escaped prior to recording the 

band number because it was the first of this species captured that year and therefore could not have 

been banded earlier in the year. Additional species were observed at the Warner Springs Banding 

Station, but not captured (table 3 at back of report).

Capture Rates 

Annual capture rate averaged 19 ± 1 per 100 net-hours and ranged from 17 (2016) to 20 (2014) 

captures per 100 net-hours for all nets combined (tables 11–15 at back of report). Annual capture rate 

by net ranged from 6 ± 5 captures per 100 net-hours (net 5) to 38 ± 8 captures per 100 net-hours (net 

14). Capture rates for these nets were relatively consistent across years. Capture rates for most other 

nets varied widely by year. In particular, nets 1 and 12 had low capture rates (2–11 captures per 100 

net-hours) for the first 4 years, then jumped to 28 and 38 captures per 100 net-hours, respectively, in 

2017. Net 4 had over 30 captures per 100 net-hours in 2013, 2014, and 2017, but only 3–5 captures per 

100 net-hours for 2015 and 2016. 

Average captures per day were highest at the beginning of the season, with peaks in the first 

and last weeks of April (15 captures per day) and lowest in the first week of June and first week of 

August (9 captures per day) (tables 16–20 at back of report). The peak in early April was mainly 

resident breeders that were naïve to the nets. The peak in late April corresponded with influxes of 

spring migrant warblers and Warbling Vireos (Vireo gilvus). A secondary peak in captures per day 

occurred during the last week of July, driven by captures of Bushtits (flock of seven) and Spotted 

Towhees. 

Species Richness

The number of species captured per day ranged from 1 to 14 (tables 16–20). Species richness 

among captures ranged from an average of 6 ± 2 per day (2016) to 9 ± 3 per day (2014). Species

richness among captures was highest in early May and lowest in early June. 
11 

Relative Species Abundance

Bushtit was the most abundant breeding MAPS species captured over 5 years with 108 

individuals captured (1.3–4.5 total captures per 100 net-hours) (tables 4, 6–10, 16–20; figs. 3–7 at back 

of report). The second most abundant breeding species was Spotted Towhee with 42 individuals 

captured (0.2–2.3 captures per 100 net-hours). Additional breeding species accounting for at least 5 

percent of all individuals captured in any 1 year include Oak Titmouse (40 individuals, 0. 7–1.7 

captures per 100 net-hours), Anna’s Hummingbird (37 individuals, 0.4–1.2 captures per 100 nethours), House Wren (36 individuals, 0.3–2.3 captures per 100 net-hours), Ash-throated Flycatcher (28 

individuals, 0.3–1.7 captures per 100 net-hours), California Scrub-jay (26 individuals, 0.2–1.1 captures 

per 100 net-hours), Bewick’s Wren (25 individuals, 0.4–1.6 captures per 100 net-hours), Acorn 

Woodpecker (17 individuals, 0–1.6 captures per 100 net-hours), California Towhee (14 individuals, 0–

0.9 captures per 100 net-hours), and Western Bluebird (Sialia mexicana; 11 individuals, 0–1.1 captures 

per 100 net-hours). Together, these 11 species account for 54 percent (384/706) of all individuals 

captured across 5 years. 

Age and Sex Structure 

Fifty-seven percent (225/394) of known-sex captures were female and 43 percent (169/394) 

were male (tables 21–25 at back of report). Juveniles comprised 33 percent (230/690) of the knownage population. The proportion of juveniles of known-age captured varied annually, with the lowest 

proportion of juveniles captured in 2014 (0.20) and the highest proportion of juveniles captured in 

2017 (0.51). Juveniles of 22 breeding species were captured, with Bushtit contributing 39 percent 

(82/210), Bewick’s Wren and Anna’s Hummingbird each contributing 9 percent (18/210), Oak 

Titmouse contributing 8 percent (17/210), and House Wren contributing 6 percent (13/210) of the 

juvenile captures. Together, these five species accounted for 64 percent (148/230) of all hatching-year 

individuals captured during MAPS. 

Productivity 

Annual productivity varied by species and by year. Productivity for seven species (Anna’s 

Hummingbird, Acorn Woodpecker, California Scrub-jay, Oak Titmouse, Bewick’s Wren, House 

Wren, and Spotted Towhee) increased from 2013 to 2014 but then productivity for three of these 

species (Acorn Woodpecker, California Scrub-jay, and Spotted Towhee) decreased, two (Anna’s 

Hummingbird and House Wren) increased, and two (Oak Titmouse and Bewick’s Wren) remained the 

same from 2014 to 2015 (fig. 8 at back of report). Average productivity across all species was lowest 

in 2014 (0.8 juveniles per adult captured) and highest in 2017 (1.7 juveniles per adult captured). 

Bushtits had the highest overall productivity (4.3 juveniles per adult), followed by Bewick’s Wrens

(2.8 juveniles per adult) and Anna’s Hummingbirds (1.5 juveniles per adult). This calculation excluded 

2015 for Bewick’s Wrens and 2016 for Acorn Woodpeckers and California Towhees because no adults 

were captured and therefore productivity could not be calculated for those years. Spotted Towhees, 

California Scrub-jays, Acorn Woodpeckers, and Ash-throated Flycatchers had the lowest productivity, 

with < 0.1–0.4 juveniles per adult captured throughout the 5 years of banding. 

Survivorship 

Models with constant survivorship across years were better supported than models with

variation in survivorship by year for all species (table 26 at back of report). Bushtits had the highest 

annual survivorship (67 percent), followed by Spotted Towhees (54 percent), Oak Titmice (43 

percent), and House Wrens (18 percent) (fig. 9 at back of report). 
12 

Population Trends

Population size of adult resident bird species, as measured by the number of adult individuals 

captured per 100 net-hours, significantly decreased for five of the most abundant species captured 

during MAPS activities (Acorn Woodpecker, California Scrub-jay, Oak Titmouse, Spotted Towhee, 

and California Towhee) and significantly increased for one species (Bushtit) over the 5 years of the 

banding project (tables 16–20; fig. 10 at back of report).

Fall Migration

Overview of Captures

We had a total of 627 migrant captures of 44 species in 2,979 net-hours during the fall from 

2013 to 2016 (tables 27, 32–35 at back of report). Of these, 558 were newly banded during the fall, 22 

were recaptures from previous years or from the MAPS season, and 14 were released unbanded (10 

hummingbirds and 4 other birds that escaped prior to banding or were released without banding; tables 

27–31 at back of report), for a total of 594 individuals captured. Ninety-five percent of banded 

individuals (553/580; 39 species) were captured only once during fall migration, 4 percent (23/580; 8 

species) were captured twice, 1 percent (3/580; 3 species) were captured three times, and <1 percent 

(1/580; 1 species) were captured four times (tables 28–31). One hummingbird with a clipped tail 

feather was recaptured during migration. Additional species were observed during migration netting, 

but were not captured (table 3).

Capture Rates

The overall fall capture rate averaged 22 ± 8 per 100 net-hours for all nets combined (range 12–

30 captures per 100 net-hours; tables 32–35). The average capture rate during fall migration was 

highest in 2013, followed by 2014, and lowest in 2016. Average capture rates by net ranged from 9 to 

48 captures per 100 net-hours with net 14 the most productive and net 9 the least productive. Capture 

rates for all years were low in mid-August, increased to a peak in mid- to late September, then trailed 

off in October, except for 2015 (tables 36–39 at back of report). In 2015, capture rates peaked for a 

second time in early October, corresponding to an influx of White-crowned Sparrows (Zonotrichia 

leucophrys).

Species Richness

The number of fall migrant species captured per day ranged from 0 to 11 (tables 36–39).

Species richness among captures varied by year but generally increased in early September and stayed 

moderate to high through late September or early October. Overall species richness for fall averaged 5 

± 3 species per day, with the highest species richness in 2013 (6 ± 2) and the lowest in 2016 (4 ± 2). 
13 

Relative Species Abundance

Spotted Towhee was the most abundant fall migrant species across all 4 years, accounting for 

22 percent of all migrant captures (138/627, 0.8–9.4 individual captures per 100 net-hours; tables 27, 

36–39; figs. 11–14 at back of report). The second most abundant species was Wilson’s Warbler

(Cardellina pusilla), comprising 15 percent of all migrants (95/627, 1.1–6.6 captures per 100 nethours). Yellow Warbler (Setophaga petechia) was the third most abundant species, comprising 9 

percent of all migrants (56/627, 1.0–3.6 captures per 100 net-hours). Three additional species, Orangecrowned Warbler (8 percent, 51/627, 0.6–2.3 captures per 100 net-hours), White-crowned Sparrow (8 

percent, 52/627, 0.5–4.2 captures per 100 net-hours), and Pacific-slope Flycatcher (Empidonax 

difficilis, 5 percent, 32/627, 0.3–2.8 captures per 100 net-hours), each accounted for at least 5 percent 

of all individuals captured. Together, these six species accounted for 68 percent (424/627) of all fall

migrant individuals captured. 

Age and Sex Structure

Forty-two percent (150/359) of known-sex captures during migration were female and 58 

percent (209/359) were male (tables 40–43 at back of report). Juveniles comprised 49 percent

(254/520) of the known-age population. Juveniles of 35 species were captured, with Yellow Warbler

and Wilson’s Warbler each contributing 15 percent (38/254 and 37/254, respectively), followed by 

Spotted Towhee and Pacific-slope Flycatcher (11 percent, 28/254 and 27/254, respectively) and Whitecrowned Sparrow and Orange-crowned Warbler (6 percent, 16/254 and 14/254, respectively).

Together, these six species accounted for 63 percent (160/254) of all hatching-year individuals 

captured during fall. 

Arrival Dates

On average, the peak number of new species arriving (captured) per day across all 4 years 

occurred during the fourth week of banding, in early to mid-September (figs. 15–18 at back of report). 

Exceptions occurred in 2013 when the peak number of new migrant species arrived during the second 

week of banding (late August) and 2015, which also had a peak during the second-to-last week of 

banding (early October). The earliest species to arrive were Least Bell’s Vireo (Vireo bellii pusillus; 

August 15, 2013), Orange-crowned Warbler (August 15, 2014), Wilson’s Warbler (August 15, 2014), 

and Costa’s Hummingbird (Calypte costae; August 18, 2016). The latest new species to arrive were 

Ruby-crowned Kinglet (Regulus calendula; October 23, 2013; October 10, 2014), Audubon’s Warbler 

(Setophaga coronata auduboni; October 10, 2014), Oregon Junco (Junco hyemalis oregonus; October 

10, 2014), Sharp-shinned Hawk (Accipiter striatus; October 15, 2015), and Red-breasted Sapsucker 

(Sphyrapicus ruber; October 13, 2016; table 44 at back of report). 

First capture dates for eight migratory species (Costa’s Hummingbird, Western Wood-pewee

[Contopus sordidulus], Ruby-crowned Kinglet, Nashville Warbler [Oreothlypis ruficapilla], 

Audubon’s Warbler, Brewer’s Sparrow [Spizella breweri], Black-throated Sparrow [Amphispiza 

bilineata], and Oregon Junco) were progressively earlier over 4 years (table 44). For two of these 

species, Costa’s Hummingbird and Ruby-crowned Kinglet, median arrival dates were also earlier each 

year from 2013 to 2016. The remaining six species were not captured enough times to calculate 

median arrival dates for multiple years. When we examined dates these eight species were first 

observed at the banding station (either seen or captured), three of the eight species (Nashville Warbler, 

Brewer’s Sparrow, and Black-throated Sparrow) were detected at the banding station earlier each year 

from 2013 to 2016. 
14 

First capture dates for five species (Red-breasted Sapsucker, Phainopepla [Phainopepla nitens], 

MacGillivray’s Warbler [Geothlypis tolmiei], Fox Sparrow [Passerella iliaca], and House Finch 

[Carpodacus mexicanus]) were progressively later each year (table 44). For two of these species, 

Phainopepla and Fox Sparrow, median arrival dates were later each year they were captured from 2013 

to 2016. 

Discussion

Our monitoring results from 2013 to 2017 indicate that the RTSWS banding station provides 

breeding habitat for 37 species (24 captured and 13 observed only) and migratory stopover or 

wintering habitat for 79 migratory species (49 captured and 30 observed only). In addition, RTSWS

provides important breeding habitat for species of conservation concern, including Nuttall’s 

Woodpecker (Picoides nuttallii), Oak Titmouse, California Thrasher (Toxostoma redivivum), and 

Lawrence’s Goldfinch (Spinus lawrencei), and migratory stopover habitat for species of conservation 

concern including Costa’s Hummingbird, Rufous Hummingbird (Selasphorus rufus), Allen’s 

Hummingbird (Selasphorus sasin), Loggerhead Shrike (Lanius ludovicianus), Yellow Warbler, 

Wilson’s Warbler, Yellow-breasted Chat (Icteria virens), Brewer’s Sparrow, and Bell’s Sparrow 

(Artemisiospiza belli) (U.S. Fish and Wildlife Service, 2008; Partners in Flight Science Committee, 

2012; California Department of Fish and Wildlife, 2017). 

Naval Base Coronado, Remote Training Site, Warner Springs also provides stopover or 

dispersal habitat for two State and Federally listed species, Least Bell’s Vireo and Willow Flycatcher 

(Empidonax traillii). Willow Flycatchers were captured and banded 4 of the 5 years, for a total of 12 

individuals. None of these were identified to subspecies, although the Southwestern Willow Flycatcher 

(E. t. extimus) subspecies breeds along the San Luis Rey River just downstream of Lake Henshaw, 

10.5 kilometers (km) southwest of the banding station (Howell and Kus, 2015). Two suspected 

migrant Willow Flycatchers were detected at the banding station during focused Least Bell’s Vireo 

surveys conducted by the U.S. Navy in May 2015 (Arnold, 2015), but the RTSWS banding station 

lacks the surface water preferred by breeding Willow Flycatchers. Two Least Bell’s Vireos were 

captured at RTSWS—one was captured twice in August 2013 and a second was captured in May 2016. 

Both were given unique color band combinations in addition to the standard metal federal band for 

visual identification on the breeding and wintering grounds. Singing male Least Bell’s Vireos were 

also detected in May and June 2014 but were not captured. Focused surveys for Least Bell’s Vireo and 

Southwestern Willow Flycatcher were conducted on a portion of RTSWS on Forest Service lands 

north of the MAPS station by USGS in 2014, but none were detected (Howell and Kus, 2014).

Focused surveys for Least Bell’s Vireo were conducted by the U.S. Navy throughout riparian habitat 

on RTSWS in 2015, 2016, and 2017 (Arnold, 2015, 2016, 2017). A suspected breeding pair of Least 

Bell’s Vireos was detected approximately 1 km downstream of the banding station in 2015, and a pair 

with three dependent juveniles was detected there in 2017. Least Bell’s Vireos could potentially breed 

at the RTSWS banding station, although the riparian habitat was drier and sparser at the banding 

station than where the breeding vireos were detected downstream, and therefore, habitat at the banding 

station may not be ideal for this species. 
15 

The average capture rate during the breeding season was relatively stable from year to year, 

with peaks corresponding to an influx of migrant species when banding days coincided with peak 

migration days in the spring. In general, however, bird capture rates during the breeding season at the 

Warner Springs banding station were low. The average capture rate at RTSWS (19 captures per 100 

net-hours) was well below capture rates at other banding stations in other habitats in San Diego 

County, which ranged from 42 to 91 captures per 100 net-hours from 2013 to 2016 (Madden and 

others, 2014; Allen and Kus, 2014, 2015; Lynn and others, 2015; Hall and Kus, 2016; Lynn and Kus, 

2017). Many of the bird species that we observed and did not capture, or captured only rarely, were 

species with behaviors that were not compatible with the standard mist-netting configuration. These

species included raptors, woodpeckers, nuthatches, crows and ravens, and flycatchers, which tended to 

forage high in the canopy or close to tall, woody trees rather than moving through the lower foliage 

where mist-nets were located. Additionally, the size of the mesh on our mist-nets is also designed to 

catch smaller birds, such as warblers and sparrows, which were poorly represented among breeding 

species at the banding station. Finally, the riparian habitat at RTSWS may have been too isolated, dry, 

or sparse to attract the abundance and diversity of bird species captured at more productive banding 

stations. 

Capture rates for migrant species during fall migration (22 per 100 net-hours) was higher than 

for all species during MAPS banding and varied by year. Many migrant species tend to forage on the 

ground or in low vegetation (sparrows, warblers) and thus were more likely to be caught in our mistnets. The annual variation in migrant capture rate may be a result of missing migratory peaks during 

days we were not banding, may be evidence of decreasing populations, or may be a response of 

migrants to changing conditions at the banding station. However, the number and diversity of both 

migratory and resident species captured during fall migration emphasize the importance of Warner 

Springs as both stopover and post-breeding dispersal habitat.

In 2013, we observed a higher capture rate for Lark Sparrows and Spotted Towhees during the 

fall than during MAPS banding. Both of these species breed at RTSWS, but the fall captures were >80 

percent adults, likely immigrants from outside populations. In fall 2014, we began identifying Spotted 

Towhee to subspecies to distinguish between the resident and migratory individuals and better 

understand their demography at RTSWS. We identified two subspecies, P. m. megalonyx (locally 

breeding) and P. m. montanus (migratory) at Warner Springs. Of the towhees, we were able to identify 

two subspecies—70 percent were P. m. megalonyx and 30 percent were P. m. montanus—thus 

verifying that Warner Springs is valuable to both residents and migrants in the fall. Only one 

subspecies of Lark Sparrow (C. g. strigatus) is present in the western United States, so we were not 

able to distinguish between breeding and migratory populations of this species. 
16 

Five years is the minimum number of consecutive years necessary to obtain reliable 

productivity indices and survivorship estimates according to the MAPS program (DeSante and others, 

2011). As a demonstration of the benefit of increasing the number of banding years, we doubled the 

number of species for which we could calculate survivorship with the addition of 1 year of data (two 

species with sufficient sample size after 4 years, four species with sufficient sample size after 5 years).

However, considering that 5 years is a minimum, the significance found in our analyses should be 

interpreted cautiously when drawing overall conclusions. Nevertheless, several of the species we 

examined showed consistent decreasing population trends using 4 years as well as 5 years of banding 

data. One of these, Spotted Towhee, also had low productivity throughout the 5 years of banding. 

Capture rates of immigrant Spotted Towhees declined each year during fall banding as well, 

decreasing 89 percent from 2013 to 2016 (7 captures per 100 net-hours to 1 capture per 100 net-hours), 

suggesting that a decrease in the Spotted Towhee population may not be confined to our banding 

station but also includes populations that disperse to RTSWS in the fall. Alternatively, migratory 

populations may have shifted their fall routes, using different stopover and wintering habitat and 

avoiding RTSWS. While the RTSWS appeared well-vegetated in the spring, we noted a progressive 

senescence in mature trees and a drier, more sparsely foliaged understory each fall through 2016, 

likely a result of drought conditions in San Diego County. San Diego County experienced a drought 

between 2012 and 2015, with precipitation for each bio-year (July 1–June 30) from July 2011 to June 

2015 totaling 38–61 percent of average precipitation between 2005 and 2011 (Weather Company, 

LLC, 2017). Low precipitation compromises primary productivity and arthropod abundance and 

consequently the wildlife (birds) that depends on them. Rainfall during the 2015–16 and 2016–17 bioyears was 18 and 15 percent above the 2005–11 average, respectively, and we detected an increase in 

adult capture rates for five species and an increase in breeding productivity for four species from 2016 

to 2017. However, the long-term effects of drought are unlikely to be eliminated immediately with the 

resumption of normal rainfall accumulation, so general bird breeding productivity and population size 

may take several years to recover.

Because of low capture rates during the MAPS season, calculations of productivity, 

survivorship, and population trends were somewhat inconclusive for other bird species that bred at 

RTSWS. Productivity was relatively high for Bushtits, Anna’s Hummingbirds, and Bewick’s Wrens, 

primarily driven by high capture rates of juveniles from 2015 through 2017, likely a response to 

unseasonably high rainfall in May and July 2015 and an increase in annual rainfall in 2016 and 2017. 

All three of these species had a subsequent increase in population in 2016, and Anna’s Hummingbird 

and Bushtit also had increases in 2017. Of these three species, we could only calculate adult 

survivorship for Bushtits, which had an average survivorship of 67 percent. As described above, 

capture rates for many species were low because the habits of the species were not necessarily 

compatible with mist-netting configuration. Bushtit are mid-canopy foliage gleaners, and they forage 

in flocks that move between clumps of vegetation at about mist-net height. Therefore, it is not 

surprising that we were able to capture a sufficient sample of Bushtits for productivity and 

survivorship analyses. However, other species (particularly warblers) also move between vegetation 

clumps gleaning from foliage, although generally not in flocks except during migration. If these 

species were as abundant as Bushtits, we would expect to see capture rates similar to Bushtits.

Although continued monitoring including more years of average rainfall may determine if 

breeding bird capture rates at Warner Springs are attributed to weather, climate change, or some other 

factor(s), the general low capture rate during the breeding season (MAPS season) suggests that efforts 

to understand long-term breeding bird population trends may best be directed toward a more 

productive banding station than the current location at RTSWS. An understanding of demographics of 

bird populations at RTSWS may be better achieved with alternate survey techniques, such as point 

counts or area searches that would have a better chance of accumulating sufficient observations for 

valid analytical conclusions. 
17 

One of the potential effects of climate change on migratory species is the timing of migration. 

As temperatures increase, the period of time during which more northerly zones are suitable for 

breeding should increase. Migratory species could potentially take advantage of this by migrating 

north earlier in the season and remaining on their breeding grounds longer before migrating south. 

Alternatively, migratory species may shift their breeding grounds northward or higher in elevation to 

remain within their preferred climate for breeding. In either case, we would expect migrants to arrive 

earlier in the spring and later in the fall in our region.

Although capture rates during fall migration were higher than during the MAPS season, we 

were still only able to draw limited conclusions regarding migrant trends at RTSWS. Of the 49 

migratory species captured during fall banding, only 13 had trends in first capture dates, and only 8 of 

these trends were supported by additional data, including median capture dates and first detection dates 

(captured or just observed). Five species (Costa’s Hummingbird, Ruby-crowned Kinglet, Nashville 

Warbler, Brewer’s Sparrow, and Black-throated Sparrow) arrived earlier each year from 2013 to 2016 

based on first capture dates and either median capture dates or first detection dates. Four of these five 

species were captured only 2 or 3 out of the 4 years, and only two to five individuals of each species 

were captured throughout the 4 years. Only Ruby-crowned Kinglet had earlier first capture and median 

capture rates for each of the 4 years of banding, and this species had progressively earlier detection 

dates from 2013 to 2015, but then was detected later again in 2016. 

Three species (Phainopepla, MacGillivray’s Warbler, and Fox Sparrow) arrived later each year 

from 2013 to 2016, based on first capture dates and either median capture dates or first detection dates. 

Two of these three species each were only captured in 2 out of the 4 years with only five captures of 

each species throughout the 4 years of banding. Phainopepla was detected much more frequently than 

it was captured (detected all 4 years but only captured in 2 years; five individuals). First detections of 

this species at the banding station were on August 19 and 20 for 3 out of the 4 years. 

Continued banding activities at RTSWS may help elucidate trends in migrant arrival dates by 

increasing the probability of capturing cryptic migrant species. However, observational methods such 

as point counts or area counts may prove to be more accurate means of collecting arrival and 

abundance data for most migrant and breeding species.

References Cited

Allen, K.S., and Kus, B.E., 2014, Neotropical migratory bird monitoring study at Marine Corps Base 

Camp Pendleton, California; 2013 annual data summary: Prepared for Assistant Chief of Staff, 

Environmental Security, Marine Corps Base Camp Pendleton.

Allen, K.S., and Kus, B.E., 2015, Neotropical migratory bird monitoring study at Marine Corps Base 

Camp Pendleton, California; 2015 annual data summary: Prepared for Assistant Chief of Staff, 

Environmental Security, Marine Corps Base Camp Pendleton.

Arnold, A., 2015, Least Bell’s Vireo presence/absence surveys at Remote Training Site Warner 

Springs, California in 2015: Prepared for Recovery Permit Coordinator, Carlsbad Fish and Wildlife 

Office, Carlsbad, Calif. 

Arnold, A., 2016, Least Bell’s Vireo presence/absence surveys at Remote Training Site Warner 

Springs, California in 2016: Prepared for Recovery Permit Coordinator, Carlsbad Fish and Wildlife 

Office, Carlsbad, Calif. 

Arnold, A., 2017, Least Bell’s Vireo presence/absence surveys at Remote Training Site Warner 

Springs, California in 2017: Prepared for Recovery Permit Coordinator, Carlsbad Fish and Wildlife 

Office, Carlsbad, Calif. 

Burnham, K.P., and Anderson, D.R., 2002, Model selection and multimodel inference: A practical 

information-theoretic approach (2d ed.): New York, Springer-Verlag, 488 p. 
18 

California Department of Fish and Wildlife, 2017, Special animals list: Natural Diversity Database, 

Periodic publication, 65 p., accessed October 2017 at 

https://nrm.dfg.ca.gov/FileHandler.ashx?DocumentID=109406&inline. 

DeSante, D.F., Williams, O.E., and Burton, K.M., 1993, The Monitoring Avian Productivity and 

Survivorship (MAPS) program: overview and progress, in Finch, D.M., and Stangel, P.W., eds., 

Status and management of neotropical migratory birds: General Technical Report RM-229, Fort 

Collins, Colo., U.S. Dept. of Agriculture, Forest Service, Rocky Mountain Forest and Range 

Experiment Station, p. 208–222. 

DeSante, D.F., Burton, K.M., Velez, P., Froehlich, D., and Kaschube, D., 2011, MAPS manual: 

Instructions for the establishment and operation of constant-effort bird-banding stations as part of the 

Monitoring Avian Productivity and Survivorship (MAPS) Program: Unpublished manuscript 

available from Institute for Bird Populations, Box 1346, Point Reyes Station, CA 94956. 

Hall, K.A., and Kus, B.E., 2016, Neotropical migratory bird monitoring study at Marine Corps Base 

Camp Pendleton, California; 2016 Annual Data Summary: Prepared for Assistant Chief of Staff, 

Environmental Security, Marine Corps Base Camp Pendleton.

Howell, S.L., and Kus, B.E., 2014, Status of the Least Bell’s Vireo and Southwestern Willow 

Flycatcher at Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, 

California, in 2014: Prepared for Commander Navy Region Southwest, San Diego, Calif. 

Howell, S.L., and Kus, B.E., 2015, The status of the Southwestern Willow Flycatcher at select sites 

along the San Luis Rey River in 2015, California: Prepared for San Diego Association of 

Governments, San Diego, Calif. 

Lynn, S., Madden, M., Houston, A., and Kus, B.E., 2015, Monitoring avian productivity and 

survivorship (MAPS) 5-year summary, Naval Outlying Landing Field, Imperial Beach, southwestern 

San Diego County, California, 2009–13: U.S. Geological Survey Open-File Report 2015–1035, 58 

p., https://dx.doi.org/10.3133/ofr20151035. 

Lynn, S. and Kus, B.E., 2017, Monitoring Breeding and Migration of Neotropical Migratory Birds at 

Point Loma, San Diego County, California, 5-Year Summary 2011–15: U.S. Geological Survey 

Open-File Report 2017–1042, 119 p., https://dx.doi.org/10.3133/ofr20171042. 

Madden, M.C., Allen, K.S., and Kus, B.E., 2014, Neotropical migratory bird monitoring study at 

Marine Corps Base Camp Pendleton, California; 2014 Annual Data Summary: Prepared for 

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http://rmbo.org/pifassessment. 

Pyle, P., 1997, Identification guide to North American birds, Part 1: Columbidae to Ploceidae: Bolinas, 

Calif., Slate Creek Press, 732 p. 

Skagen, S.K., Melcher, C.P., and Hazelwood, R., 2004, Migration stopover ecology of Western avian 

populations: a Southwestern migration workshop: U.S. Geological Survey Open-File Report 2004–

1452, 28 p. 

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web page, accessed October 31, 2017, at https://www.wunderground.com/history/airport/KRNM. 

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marked animals: Bird Study, v. 46 [Supplement], p. 120–139. 

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Federal Resource Laws of Interest to the U.S. Fish and Wildlife Service, accessed at 

https://www.fws.gov/laws/lawsdigest/migtrea.html. 

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Service, Division of Migratory Bird Management, 85 p., 

https://www.fws.gov/migratorybirds/pdf/grants/BirdsofConservationConcern2008.pdf. 
19 

U.S. Navy, 2013, Final Integrated Natural Resources Management Plan, Naval Base Coronado, 

California: Commander, Navy Installations Command, 580 p., accessed at 

https://www.cnic.navy.mil/content/dam/cnic/cnrsw/NAVFACSW%20Environmental%20Core/NBC

_INRMP_signed.pdf. 
20 

Table 2. Species captured at Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, 

California, 2013–17. 

[Species in italics are non-breeding migrants or transients. Special status: CC, Partners in Flight Species of Conservation 

Concern (U.S. Fish and Wildlife Service, 2008; Partners in Flight Science Committee, 2012; California Department of Fish 

and Wildlife, 2017); FC, Federal Species of Conservation Concern; FE, Federally listed as Endangered; SC, California 

State Species of Concern; SE, California State listed as Endangered. Season detected: B, Monitoring Avian Productivity 

and Survivorship (breeding season), F, fall (migratory season). Data collected by USGS and available from 

http://www.pwrc.usgs.gov/bbl] 

Alpha code Common name Scientific name Special status Season detected

SSHA1 Sharp-shinned Hawk Accipiter striatus F

CAQU California Quail Callipepla californica B, F

MODO1 Mourning Dove Zenaida macroura B, F

BCHU1 Black-chinned Hummingbird Archilochus alexandri B, F

ANHU1 Anna's Hummingbird Calypte anna B, F

COHU1 Costa's Hummingbird Calypte costae FC B, F

CAHU1 Calliope Hummingbird Stellula calliope B

RUHU1 Rufous Hummingbird Selasphorus rufus CC, FC B

ALHU1 Allen's Hummingbird Selasphorus sasin CC, FC B, F

ACWO1 Acorn Woodpecker Melanerpes formicivorus B, F

RNSA1 Red-naped Sapsucker Sphyrapicus nuchalis F

RBSA1 Red-breasted Sapsucker Sphyrapicus ruber F

NUWO1 Nuttall's Woodpecker Picoides nuttallii FC B, F

DOWO1 Downy Woodpecker Picoides pubescens B, F

RSFL1 Red-shafted Flicker Colaptes auratus cafer B, F

WEWP1 Western Wood-Pewee Contopus sordidulus B, F

WIFL1 Willow Flycatcher Empidonax traillii FC, SE B, F

HAFL1 Hammond’s Flycatcher Empidonax hammondii B

GRFL1 Gray Flycatcher Empidonax wrightii B, F

DUFL1 Dusky Flycatcher Empidonax oberholseri F

PSFL1 Pacific-slope Flycatcher Empidonax difficilis B, F

ATFL1 Ash-throated Flycatcher Myiarchus cinerascens B, F

WEKI1 Western Kingbird Tyrannus verticalis B

LBVI1 Least Bell's Vireo Vireo bellii pusillus CC, FE, SE B, F

HUVI1 Hutton's Vireo Vireo huttoni B, F

WAVI1 Warbling Vireo Vireo gilvus B, F

CASJ1 California Scrub-Jay Aphelocoma californica B, F

OATI1 Oak Titmouse Baeolophus inornatus CC, FC B, F

BUSH1 Bushtit Psaltriparus minimus B, F

WBNU1 White-breasted Nuthatch Sitta carolinensis B, F

BEWR1 Bewick's Wren Thryomanes bewickii B, F

HOWR1 House Wren Troglodytes aedon B, F

RCKI1 Ruby-crowned Kinglet Regulus calendula B, F

WEBL1 Western Bluebird Sialia mexicana B, F

SWTH1 Swainson's Thrush Catharus ustulatus B, F

HETH1 Hermit Thrush Catharus guttatus B, F
21 

Alpha code Common name Scientific name Special status Season detected

WREN Wrentit Chamaea fasciata CC B

NOMO1 Northern Mockingbird Mimus polyglottos F

CATH1 California Thrasher Toxostoma redivivum CC B, F

EUST European Starling Sturnus vulgaris B, F

PHAI1 Phainopepla Phainopepla nitens F

OCWA1 Orange-crowned Warbler Oreothlypis celata B, F

NAWA1 Nashville Warbler Oreothlypis ruficapilla B, F

YEWA1 Yellow Warbler Setophaga petechia FC, SC B, F

AUWA1 Audubon's Warbler Setophaga coronata auduboni B, F

BTYW1 Black-throated Gray Warbler Setophaga nigrescens F

BLPW1 Blackpoll Warbler Setophaga striata F

MGWA1 MacGillivray's Warbler Geothlypis tolmiei B, F

COYE1 Common Yellowthroat Geothlypis trichas B, F

WIWA1 Wilson's Warbler Cardellina pusilla CC B, F

YBCH1 Yellow-breasted Chat Icteria virens SC F

WETA1 Western Tanager Piranga ludoviciana B, F

GTTO1 Green-tailed Towhee Pipilo chlorurus F

SPTO1 Spotted Towhee Pipilo maculatus B, F

CALT1 California Towhee Melozone crissalis B, F

CHSP1 Chipping Sparrow Spizella passerina F

BRSP1 Brewer's Sparrow Spizella breweri CC B, F

LASP1 Lark Sparrow Chondestes grammacus B, F

BTSP1 Black-throated Sparrow Amphispiza bilineata B, F

BESP1 Bell's Sparrow Artemisiospiza belli SC B

FOSP1 Fox Sparrow Passerella iliaca F

SOSP1 Song Sparrow Melospiza melodia B, F

LISP1 Lincoln's Sparrow Melospiza lincolnii B, F

WCSP1 White-crowned Sparrow Zonotrichia leucophrys B, F

GCSP1 Golden-crowned Sparrow Zonotrichia atricapilla B

ORJU1 Oregon Junco Junco hyemalis oregonus B, F

BHGR1 Black-headed Grosbeak Pheucticus melanocephalus B, F

BUOR1 Bullock's Oriole Icterus bullockii B, F

PUFI1 Purple Finch Carpodacus purpureus F

HOFI1 House Finch Carpodacus mexicanus B, F

LEGO1 Lesser Goldfinch Spinus psaltria B, F

LAZB1 Lazuli Bunting Passerina amoena F

1

Protected species under the Migratory Bird Treaty Act of 1918.
22 

Table 3. Species observed but not captured at Naval Base Coronado, Remote Training Site, Warner Springs, 

San Diego County, California, 2013–17.

[Species in italics are non-breeding migrants or transients. Special status: CC, Partners in Flight Species of Conservation 

Concern (U.S. Fish and Wildlife Service, 2008; Partners in Flight Science Committee, 2012; California Department of Fish 

and Wildlife, 2017); FC, Federal Species of Conservation Concern; FE, Federally listed as Endangered; SC, California 

State Species of Concern; SE, California State listed as Endangered. Season detected: B, Monitoring Avian Productivity 

and Survivorship (breeding season), F, fall (migratory season). Data collected by USGS and available from 

http://www.pwrc.usgs.gov/bbl] 

Alpha code Common name Scientific name Special status Season detected

TUVU1 Turkey Vulture Cathartes aura B

MALL1 Mallard Anas platyrhynchos B, F

COHA1 Cooper's Hawk Accipiter cooperii B, F

RSHA1 Red-shouldered Hawk Buteo lineatus B, F

RTHA1 Red-tailed Hawk Buteo jamaicensis B, F

GOEA1 Golden Eagle Aquila chrysaetos B

AMKE1 American Kestrel Falco sparverius B, F

INPE Indian Peafowl Pavo cristatus B, F

WITU1 Wild Turkey Meleagris gallopavo B, F

KILL1 Killdeer Charadrius vociferus F

BTPI1 Band-tailed Pigeon Patagioenas fasciata B, F

EUCD Eurasian Collared-Dove Streptopelia decaocto B, F

WWDO1 White-winged Dove Zenaida asiatica F

COGD1 Common Ground-Dove Columbina passerina F

GRRO1 Greater Roadrunner Geococcyx californianus B

BANO1 Barn Owl Tyto alba B

WESO1 Western Screech-Owl Megascops kennicottii B, F

GHOW1 Great Horned Owl Bubo virginianus B, F

LENI1 Lesser Nighthawk Chordeiles acutipennis B, F

HAWO1 Hairy Woodpecker Picoides villosus F

BLPH1 Black Phoebe Sayornis nigricans F

SAPH1 Say's Phoebe Sayornis saya B, F

LOSH1 Loggerhead Shrike Lanius ludovicianus CC, FC F

CAVI1 Cassin's Vireo Vireo cassinii B

STJA1 Steller's Jay Cyanocitta stelleri F

AMCR1 American Crow Corvus brachyrhynchos B, F

CORA1 Common Raven Corvus corax B, F

BARS1 Barn Swallow Hirundo rustica F

MOCH1 Mountain Chickadee Poecile gambeli F

BGGN1 Blue-gray Gnatcatcher Polioptila caerulea B, F

BTGN1 Black-tailed Gnatcatcher Polioptila melanura B

TOSO1 Townsend's Solitaire Myadestes townsendi F

CEDW1 Cedar Waxwing Bombycilla cedrorum F

TOWA1 Townsend's Warbler Setophaga townsendi F

HEWA1 Hermit Warbler Setophaga occidentalis F

CLSW1 Cliff Swallow Petrochelidon pyrrhonota B

SAVS1 Savannah Sparrow Passerculus sandwichensis F
23 

Alpha code Common name Scientific name Special status Season detected

BLGR1 Blue Grosbeak Passerina caerulea B, F

WEME1 Western Meadowlark Sturnella neglecta B, F

BHCO1 Brown-headed Cowbird Molothrus ater B, F

HOOR1 Hooded Oriole Icterus cucullatus B, F

LAGO1 Lawrence's Goldfinch Spinus lawrencei CC, FC B, F

AMGO1 American Goldfinch Spinus tristis B, F

AMRO1 American Robin Turdus migratorius B, F

1

Protected species under the Migratory Bird Treaty Act of 1918.
24 

Table 4. Number of birds captured and banded, during Monitoring Avian Productivity and Survivorship, Naval Base Coronado, Remote Training Site, 

Warner Springs, San Diego County, 2013–17.

[See tables 2 and 3 for bird species codes. Species in italics are non-breeding neotropical migrants. Includes multiple captures of some individuals. Data collected by 

USGS and available from http://www.pwrc.usgs.gov/bbl] 

Total captures Total number of individuals captured New individuals banded

Year Year Year

Species 2013 2014 2015 2016 2017 Total 2013 2014 2015 2016 2017 Total 2013 2014 2015 2016 2017 Total

CAQU 0 1 1 3 1 6 0 1 1 3 1 6 0 0 0 0 0 0

MODO 0 0 0 1 0 1 0 0 0 1 0 1 0 0 0 0 0 0

BCHU 1 1 0 4 1 7 1 1 0 4 1 7 0 0 0 0 0 0

ANHU 9 9 6 3 10 37 9 9 6 3 9 36 0 0 0 0 0 0

COHU 1 2 8 1 0 12 1 2 8 1 0 12 0 0 0 0 0 0

CAHU 0 0 1 0 0 1 0 0 1 0 0 1 0 0 0 0 0 0

RUHU 0 1 3 0 0 4 0 1 3 0 0 4 0 0 0 0 0 0

ALHU 1 1 1 0 1 4 1 1 1 0 1 4 0 0 0 0 0 0

ACWO 12 3 2 0 1 18 11 3 2 0 1 17 11 3 2 0 1 17

NUWO 5 2 4 3 3 17 3 2 3 3 3 14 3 2 2 2 2 11

DOWO 1 2 0 3 0 6 1 1 0 2 0 4 1 1 0 2 0 4

RSFL 2 0 1 0 3 6 1 0 1 0 3 5 1 0 1 0 3 5

WEWP 0 0 1 0 0 1 0 0 1 0 0 1 0 0 1 0 0 1

WIFL 0 1 1 0 0 2 0 1 1 0 0 2 0 1 1 0 0 2

HAFL 0 2 0 0 0 2 0 2 0 0 0 2 0 2 0 0 0 2

GRFL 1 0 0 0 0 1 1 0 0 0 0 1 1 0 0 0 0 1

PSFL 0 3 5 1 0 9 0 3 5 1 0 9 0 3 5 1 0 9

ATFL 8 3 6 11 4 32 8 3 6 9 2 28 8 2 4 8 2 24

WEKI 0 0 0 1 0 1 0 0 0 1 0 1 0 0 0 1 0 1

LBVI 0 0 0 1 0 1 0 0 0 1 0 1 0 0 0 1 0 1

HUVI 0 0 0 0 2 2 0 0 0 0 1 1 0 0 0 0 1 1

WAVI 1 4 13 2 0 20 1 4 13 2 0 20 1 4 13 2 0 20

CASJ 8 7 2 4 5 26 8 7 2 4 5 26 8 6 0 4 4 22

OATI 11 6 8 13 11 49 8 6 6 9 11 40 8 1 4 7 7 27

BUSH 12 12 32 23 40 119 12 9 30 21 36 108 11 8 24 18 32 93

WBNU 3 7 2 1 4 17 3 6 2 1 4 16 2 3 0 1 2 8
25 

Total captures Total number of individuals captured New individuals banded

Year Year Year

Species 2013 2014 2015 2016 2017 Total 2013 2014 2015 2016 2017 Total 2013 2014 2015 2016 2017 Total

BEWR 3 5 7 12 10 37 3 4 4 7 7 25 3 3 4 5 5 20

HOWR 12 12 3 4 20 51 9 8 2 4 13 36 7 5 1 4 12 29

RCKI 0 0 1 0 0 1 0 0 1 0 0 1 0 0 1 0 0 1

WEBL 0 3 0 0 10 13 0 3 0 0 8 11 0 3 0 0 8 11

SWTH 1 1 3 1 0 6 1 1 3 1 0 6 1 1 3 1 0 6

HETH 0 2 0 0 0 2 0 2 0 0 0 2 0 2 0 0 0 2

WREN 4 1 0 0 0 5 4 1 0 0 0 5 3 1 0 0 0 4

CATH 2 0 1 5 3 11 2 0 1 4 3 10 2 0 1 2 3 8

EUST 0 1 0 0 0 1 0 1 0 0 0 1 0 0 0 0 0 0

OCWA 3 18 1 1 0 23 3 18 1 1 0 23 3 18 1 1 0 23

NAWA 1 4 1 0 0 6 1 4 1 0 0 6 1 4 1 0 0 6

YEWA 2 3 5 2 1 13 2 3 5 2 1 13 2 3 5 2 1 13

AUWA 0 2 3 0 12 17 0 2 3 0 12 17 0 2 3 0 12 17

MGWA 1 1 1 0 0 3 1 1 1 0 0 3 1 1 1 0 0 3

COYE 1 0 0 0 2 3 1 0 0 0 2 3 1 0 0 0 2 3

WIWA 4 17 14 15 4 54 4 17 14 15 4 54 4 16 14 15 4 53

WETA 1 2 0 0 0 3 1 2 0 0 0 3 1 2 0 0 0 3

SPTO 17 19 10 3 2 51 13 17 7 3 2 42 12 13 4 0 1 30

CALT 8 4 2 0 2 16 6 4 2 0 2 14 6 2 2 0 2 12

BRSP 1 0 0 0 0 1 1 0 0 0 0 1 1 0 0 0 0 1

LASP 0 0 0 4 0 4 0 0 0 4 0 4 0 0 0 4 0 4

BTSP 0 1 0 0 0 1 0 1 0 0 0 1 0 1 0 0 0 1

BESP 0 1 0 0 1 2 0 1 0 0 1 2 0 1 0 0 1 2

SOSP 0 2 0 1 1 4 0 2 0 1 1 4 0 2 0 1 1 4

LISP 0 0 0 0 1 1 0 0 0 0 1 1 0 0 0 0 1 1

WCSP 2 3 7 3 4 19 2 3 7 3 4 19 2 3 7 3 4 19

GCSP 0 0 1 0 0 1 0 0 1 0 0 1 0 0 1 0 0 1

ORJU 0 2 0 0 0 2 0 2 0 0 0 2 0 2 0 0 0 2

BHGR 1 1 2 2 3 9 1 1 2 2 3 9 1 1 2 2 3 9

BUOR 5 7 4 3 0 19 5 5 3 3 0 16 5 4 3 3 0 15

HOFI 0 0 1 0 0 1 0 0 1 0 0 1 0 0 1 0 0 1
26 

Total captures Total number of individuals captured New individuals banded

Year Year Year

Species 2013 2014 2015 2016 2017 Total 2013 2014 2015 2016 2017 Total 2013 2014 2015 2016 2017 Total

LEGO 0 2 0 0 0 2 0 2 0 0 0 2 0 2 0 0 0 2

Total 145 181 164 131 162 783 129 167 151 116 142 705 111 128 112 90 114 555
27 

Table 5. Number of birds recaptured during Monitoring Avian Productivity and Survivorship, Warner Springs, San 

Diego County, California, 2013–17.

[See tables 2 and 3 for bird species codes. Species in italics are non-breeding neotropical migrants. Includes multiple 

captures of some individuals. Recaptures originally banded during previous banding activities at Warner Springs, all 

seasons. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl] 

Species 

code

Recaptured individuals 

2014

Recaptured individuals 

2015

Recaptured individuals, 

2016

Recaptured individuals,

2017

Year originally 

banded Year originally banded

Year originally

banded

Year originally 

banded 

2013 Total 2013 2014 Total 2013 2014 2015 Total 2014 2015 2016 Total

NUWO 0 0 0 1 1 1 0 0 1 0 0 1 1

ATFL 1 1 1 1 2 0 0 1 1 0 0 0 0

CASJ 1 1 1 1 2 0 0 0 0 1 0 0 1

OATI 5 5 1 1 2 0 0 2 2 0 3 1 4

BUSH 1 1 3 1 4 0 1 0 1 0 0 0 0

WBNU 3 3 1 1 2 0 0 0 0 0 0 2 2

BEWR 0 0 0 0 0 0 0 1 1 0 1 0 1

HOWR 3 3 1 0 1 0 0 0 0 0 0 0 0

CATH 0 0 0 0 0 1 0 0 1 0 0 0 0

SPTO 4 4 1 2 3 0 1 0 2 1 0 0 1

CALT 2 2 0 0 0 0 0 0 0 0 0 0 0

BUOR 1 1 0 0 0 0 0 0 0 0 0 0 0

Total 21 21 9 8 17 2 2 4 9 2 4 4 10
28 

Table 6. Capture frequency of individuals during Monitoring Avian Productivity and Survivorship, Naval Base 

Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2013.

[See tables 2 and 3 for bird species codes. Species in italics are non-breeding neotropical migrants. Data collected by USGS 

and available from http://www.pwrc.usgs.gov/bbl] 

Number of individuals per capture incidence

(banded birds only) Total number of individuals

Species 1 capture 2 captures 3 captures Banded birds Unbanded birds All birds

BCHU 0 0 0 0 1 1

ANHU 0 0 0 0 9 9

COHU 0 0 0 0 1 1

ALHU 0 0 0 0 1 1

ACWO 10 1 0 11 0 11

NUWO 1 2 0 3 0 3

DOWO 1 0 0 1 0 1

RSFL 0 1 0 1 0 1

GRFL 1 0 0 1 0 1

ATFL 8 0 0 8 0 8

WAVI 1 0 0 1 0 1

CASJ 0 0 0 8 0 8

OATI 6 1 1 8 0 8

BUSH 11 0 0 11 1 12

WBNU 2 0 0 2 1 3

BEWR 3 0 0 3 0 3

HOWR 5 1 1 7 2 9

SWTH 1 0 0 1 0 1

WREN 3 0 0 3 1 4

CATH 2 0 0 2 0 2

OCWA 3 0 0 3 0 3

NAWA 1 0 0 1 0 1

YEWA 2 0 0 2 0 2

MGWA 1 0 0 1 0 1

COYE 1 0 0 1 0 1

WIWA 4 0 0 4 0 4

WETA 1 0 0 1 0 1

SPTO 9 2 1 12 1 13

CALT 4 2 0 6 0 6

BRSP 1 0 0 1 0 1

WCSP 2 0 0 2 0 2

BHGR 1 0 0 1 0 1

BUOR 5 0 0 5 0 5

Total 98 10 3 111 18 129
29 

Table 7. Capture frequency of individuals during Monitoring Avian Productivity and Survivorship, Naval Base 

Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2014.

[See tables 2 and 3 for bird species codes. Species in italics are non-breeding neotropical migrants. Data collected by USGS 

and available from http://www.pwrc.usgs.gov/bbl] 

Number of individuals per capture incidence

(banded birds only) Total number of individuals

Species 1 capture 2 captures 3 captures Banded birds Unbanded birds All birds

CAQU 0 0 0 0 1 1

BCHU 0 0 0 0 1 1

ANHU 0 0 0 0 9 9

COHU 0 0 0 0 2 2

RUHU 0 0 0 0 1 1

ALHU 0 0 0 0 1 1

ACWO 3 0 0 3 0 3

NUWO 2 0 0 2 0 2

DOWO 0 1 0 1 0 1

WIFL 1 0 0 1 0 1

HAFL 2 0 0 2 0 2

PSFL 3 0 0 3 0 3

ATFL 3 0 0 3 0 3

WAVI 4 0 0 4 0 4

CASJ 7 0 0 7 0 7

OATI 6 0 0 6 0 6

BUSH 7 2 0 91 0 91 

WBNU 5 1 0 6 0 6

BEWR 2 1 0 3 1 4

HOWR 5 2 1 8 0 8

WEBL 3 0 0 3 0 3

SWTH 1 0 0 1 0 1

HETH 2 0 0 2 0 2

WREN 1 0 0 1 0 1

EUST 0 0 0 0 1 1

OCWA 18 0 0 18 0 18

NAWA 4 0 0 4 0 4

YEWA 3 0 0 3 0 3

AUWA 2 0 0 2 0 2

MGWA 1 0 0 1 0 1

WIWA 16 0 0 16 1 17

WETA 2 0 0 2 0 2

SPTO 15 2 0 17 0 17

CALT 4 0 0 4 0 4

BTSP 1 0 0 1 0 1

BESP 1 0 0 1 0 1

SOSP 2 0 0 2 0 2

WCSP 3 0 0 3 0 3
30 

Number of individuals per capture incidence

(banded birds only) Total number of individuals

Species 1 capture 2 captures 3 captures Banded birds Unbanded birds All birds

ORJU 2 0 0 2 0 2

BHGR 1 0 0 1 0 1

BUOR 3 2 0 5 0 5

LEGO 2 0 0 2 0 2

Total 137 11 1 1491 18 1671 

1

Does not include one banded BUSH that was recaptured and escaped before recording the band number.
31 

Table 8. Capture frequency of individuals during Monitoring Avian Productivity and Survivorship, Naval Base 

Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2015.

[See tables 2 and 3 for bird species codes. Species in italics are non-breeding neotropical migrants. Data collected by USGS 

and available from http://www.pwrc.usgs.gov/bbl] 

Number of individuals per capture incidence

(banded birds only) Total number of individuals

Species 1 capture 2 captures 4 captures Banded 

bi d

Unbanded birds All birds

CAQU 0 0 0 0 1 1

ANHU 0 0 0 0 6 6

COHU 0 0 0 0 8 8

CAHU 0 0 0 0 1 1

RUHU 0 0 0 0 3 3

ALHU 0 0 0 0 1 1

ACWO 2 0 0 2 0 2

NUWO 2 1 0 3 0 3

RSFL 1 0 0 1 0 1

WEWP 1 0 0 1 0 1

WIFL 1 0 0 1 0 1

PSFL 5 0 0 5 0 5

ATFL 6 0 0 6 0 6

WAVI 13 0 0 13 0 13

CASJ 2 0 0 2 0 2

OATI 5 1 0 61 0 61 

BUSH 26 2 0 28 2 30

WBNU 2 0 0 2 0 2

BEWR 3 0 1 4 0 4

HOWR 1 1 0 2 0 2

RCKI 1 0 0 1 0 1

SWTH 3 0 0 3 0 3

CATH 1 0 0 1 0 1

OCWA 1 0 0 1 0 1

NAWA 1 0 0 1 0 1

YEWA 5 0 0 5 0 5

AUWA 3 0 0 3 0 3

MGWA 1 0 0 1 0 1

WIWA 14 0 0 14 0 14

SPTO 4 3 0 7 0 7

CALT 2 0 0 2 0 2

WCSP 7 0 0 7 0 7

GCSP 1 0 0 1 0 1

BHGR 2 0 0 2 0 2

BUOR 2 1 0 3 0 3

HOFI 1 0 0 1 0 1

Total 119 9 1 1291 22 1511

1

Does not include one banded OATI that was recaptured and escaped before recording the band number.
32 

Table 9. Capture frequency of individuals during Monitoring Avian Productivity and Survivorship, Naval Base 

Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2016.

[See tables 2 and 3 for bird species codes. Species in italics are non-breeding neotropical migrants. Data collected by USGS 

and available from http://www.pwrc.usgs.gov/bbl] 

Number of individuals per capture incidence

(Banded birds only) Total number of individuals

Species 1 capture 2 captures 3 captures 4 captures Banded birds Unbanded birds All birds

CAQU 0 0 0 0 0 3 3

MODO 0 0 0 0 0 1 1

BCHU 0 0 0 0 0 4 4

ANHU 0 0 0 0 0 3 3

COHU 0 0 0 0 0 1 1

NUWO 3 0 0 0 3 0 3

DOWO 1 1 0 0 2 0 2

PSFL 1 0 0 0 1 0 1

ATFL 7 2 0 0 9 0 9

WEKI 1 0 0 0 1 0 1

LBVI 1 0 0 0 1 0 1

WAVI 2 0 0 0 2 0 2

CASJ 4 0 0 0 4 0 4

OATI 5 4 0 0 9 0 9

BUSH 17 2 0 0 19 2 21

WBNU 1 0 0 0 1 0 1

BEWR 4 0 1 1 6 1 7

HOWR 4 0 0 0 4 0 4

SWTH 1 0 0 0 1 0 1

CATH 2 1 0 0 3 1 4

OCWA 1 0 0 0 1 0 1

YEWA 2 0 0 0 2 0 2

WIWA 15 0 0 0 15 0 15

SPTO 21 0 0 0 21 1 31 

LASP 4 0 0 0 4 0 4

SOSP 1 0 0 0 1 0 1

WCSP 3 0 0 0 3 0 3

BHGR 2 0 0 0 2 0 2

BUOR 3 0 0 0 3 0 3

Total 871 10 1 1 991 17 1161 

1

Includes one banded SPTO that was recaptured and escaped before recording the band number because it was the first 

capture of this species this year.
33 

Table 10. Capture frequency of individuals during Monitoring Avian Productivity and Survivorship, Naval Base 

Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2017. 

[See tables 2 and 3 for bird species codes. Species in italics are non-breeding neotropical migrants. Data collected by USGS 

and available from http://www.pwrc.usgs.gov/bbl] 

Number of individuals per capture incidence

(Banded birds only) Total number of individuals

Species 1 capture 2 captures 3 captures 4 captures Banded birds Unbanded birds All birds

CAQU 0 0 0 0 0 1 1

BCHU 0 0 0 0 0 1 1

ANHU 0 0 0 0 0 9 9

ALHU 0 0 0 0 0 1 1

ACWO 1 0 0 0 1 0 1

NUWO 3 0 0 0 3 0 3

RSFL 3 0 0 0 3 0 3

ATFL 0 2 0 0 2 0 2

HUVI 0 1 0 0 1 0 1

CASJ 5 0 0 0 5 0 5

OATI 11 0 0 0 11 0 11

BUSH 29 3 0 0 321 4 36

WBNU 4 0 0 0 4 0 4

BEWR 4 1 1 0 6 1 7

HOWR 9 1 0 2 12 1 13

WEBL 6 2 0 0 8 0 8

CATH 3 0 0 0 3 0 3

YEWA 1 0 0 0 1 0 1

AUWA 12 0 0 0 12 0 12

COYE 2 0 0 0 2 0 2

WIWA 4 0 0 0 4 0 4

SPTO 2 0 0 0 2 0 2

CALT 2 0 0 0 2 0 2

SAGS 1 0 0 0 1 0 1

SOSP 1 0 0 0 1 0 1

LISP 1 0 0 0 1 0 1

WCSP 4 0 0 0 4 0 4

BHGR 3 0 0 0 3 0 3

Total 111 10 1 2 124 18 142

1

Does not include one banded BUSH that was recaptured and escaped before recording the band number.
34 

Table 11. Capture rate by net and date during Monitoring Avian Productivity and Survivorship, Naval Base Coronado, Remote Training Site, Warner 

Springs, San Diego County, California, 2013.

[MAPS periods were established by the Institute for Bird Populations for program-wide consistency. In San Diego, banding began 3 periods early to encompass the 

earlier breeding season that occurs at southern latitudes. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl] 

MAPS 

period Date

Net

Totals by 

1 2 3 4 5 6 7 8 9 10 11 12 13 date

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 0:00 0:00 0:00 50:00

Captures 0 0 1 1 0 2 2 0 0 3 0 0 0 9

-3 4/5/2013 Captures per net-hour 0.00 0.00 0.20 0.20 0.00 0.40 0.40 0.00 0.00 0.60 -- -- -- 0.18

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 0:00 0:00 0:00 50:00

Captures 0 1 0 1 0 1 5 3 1 1 0 0 0 13

-2 4/12/2013 Captures per net-hour 0.00 0.20 0.00 0.20 0.00 0.20 1.00 0.60 0.20 0.20 -- -- -- 0.26

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 0:00 0:00 0:00 50:00

Captures 0 2 1 4 1 0 0 2 4 0 0 0 0 14

-1 4/26/2013 Captures per net-hour 0.00 0.40 0.20 0.80 0.20 0.00 0.00 0.40 0.80 0.00 -- -- -- 0.28

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 0:00 0:00 0:00 50:00

Captures 2 0 0 6 1 2 0 4 1 1 0 0 0 17

1 5/3/2013 Captures per net-hour 0.40 0.00 0.00 1.20 0.20 0.40 0.00 0.80 0.20 0.20 -- -- -- 0.34

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 0:00 0:00 0:00 50:00

Captures 0 0 0 0 0 2 2 3 0 0 0 0 0 7

2 5/17/2013 Captures per net-hour 0.00 0.00 0.00 0.00 0.00 0.40 0.40 0.60 0.00 0.00 -- -- -- 0.14

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 0:00 0:00 0:00 50:00

Captures 0 0 2 1 2 4 3 1 2 0 0 0 0 15

3 5/24/2013 Captures per net-hour 0.00 0.00 0.40 0.20 0.40 0.80 0.60 0.20 0.40 0.00 -- -- -- 0.30

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 65:00

Captures 0 1 5 0 1 0 2 1 1 0 1 1 2 15

4 6/7/2013 Captures per net-hour 0.00 0.20 1.00 0.00 0.20 0.00 0.40 0.20 0.20 0.00 0.20 0.20 0.40 0.23

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 65:00

Captures 0 2 1 1 0 0 0 0 3 0 0 0 0 7

5 6/14/2013 Captures per net-hour 0.00 0.40 0.20 0.20 0.00 0.00 0.00 0.00 0.60 0.00 0.00 0.00 0.00 0.11
35 

MAPS 

period Date

Net

Totals by 

1 2 3 4 5 6 7 8 9 10 11 12 13 date

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 65:00

Captures 0 0 0 4 0 0 2 1 3 1 0 1 0 12

6 6/21/2013 Captures per net-hour 0.00 0.00 0.00 0.80 0.00 0.00 0.40 0.20 0.60 0.20 0.00 0.20 0.00 0.18

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 65:00

Captures 0 1 0 3 0 1 0 8 0 0 0 0 1 14

7 7/4/2013 Captures per net-hour 0.00 0.20 0.00 0.60 0.00 0.20 0.00 1.60 0.00 0.00 0.00 0.00 0.20 0.22

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 65:00

Captures 2 1 2 0 0 2 0 0 1 0 0 0 1 9

8 7/12/2013 Captures per net-hour 0.40 0.20 0.40 0.00 0.00 0.40 0.00 0.00 0.20 0.00 0.00 0.00 0.20 0.14

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 65:00

Captures 1 1 0 0 0 1 0 0 0 1 0 0 1 5

9 7/26/2013 Captures per net-hour 0.20 0.20 0.00 0.00 0.00 0.20 0.00 0.00 0.00 0.20 0.00 0.00 0.20 0.08

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 65:00

Captures 1 3 0 0 0 0 0 2 1 0 0 0 1 8

10 8/2/2013 Captures per net-hour 0.20 0.60 0.00 0.00 0.00 0.00 0.00 0.40 0.20 0.00 0.00 0.00 0.20 0.12

Totals by net

Net-hours 65:0 65:0 65:0 65:0 65:0 65:0 65:0 65:0 65:00 65:0 35:0 35:0 35:0 755:00

Captures 6 12 12 21 5 15 16 25 17 7 1 2 6 145

Captures per net-hour 0.09 0.18 0.18 0.32 0.08 0.23 0.25 0.38 0.26 0.11 0.03 0.06 0.17 0.19
36 

Table 12. Capture rate by net and date during Monitoring Avian Productivity and Survivorship, Naval Base Coronado, Remote Training Site, Warner 

Springs, San Diego County, California, 2014.

[MAPS periods were established by the Institute for Bird Populations for program-wide consistency. In San Diego, banding began 3 periods early to encompass the 

earlier breeding season that occurs at southern latitudes. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl] 

MAPS 

period

Net Totals 

by

Date 1 2 3 4 5 6 7 8 9 10 11 12 13 14 date

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 50:00

Captures 0 1 4 2 3 0 2 2 1 2 1 0 5 2 25

-3 4/4/2014 Captures per net-hour 0.00 0.20 0.80 0.40 0.60 0.00 0.40 0.40 0.20 0.40 0.20 0.00 1.00 0.40 0.50

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 50:00

Captures 0 4 1 2 0 3 1 1 1 2 2 0 5 6 28

-2 4/11/2014 Captures per net-hour 0.00 0.80 0.20 0.40 0.00 0.60 0.20 0.20 0.20 0.40 0.40 0.00 1.00 1.20 0.56

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 50:00

Captures 0 1 0 3 0 1 1 1 2 0 0 1 2 1 13

-1 4/25/2014 Captures per net-hour 0.00 0.20 0.00 0.60 0.00 0.20 0.20 0.20 0.40 0.00 0.00 0.20 0.40 0.20 0.26

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 50:00

Captures 1 0 2 1 1 0 1 0 1 0 0 0 3 0 10

1 5/2/2014 Captures per net-hour 0.20 0.00 0.40 0.20 0.20 0.00 0.20 0.00 0.20 0.00 0.00 0.00 0.60 0.00 0.20

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 50:00

Captures 0 1 0 1 2 2 1 1 9 1 1 1 2 3 25

2 5/16/2014 Captures per net-hour 0.00 0.20 0.00 0.20 0.40 0.40 0.20 0.20 1.80 0.20 0.20 0.20 0.40 0.60 1.06

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 50:00

Captures 0 0 0 3 0 1 2 0 1 0 0 0 1 2 10

3 5/23/2014 Captures per net-hour 0.00 0.00 0.00 0.60 0.00 0.20 0.40 0.00 0.20 0.00 0.00 0.00 0.20 0.40 0.20

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 50:00

Captures 0 1 0 0 0 0 1 0 3 1 1 1 0 0 8

4 6/6/2014 Captures per net-hour 0.00 0.20 0.00 0.00 0.00 0.00 0.20 0.00 0.60 0.20 0.20 0.20 0.00 0.00 0.16

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 50:00

Captures 0 0 1 0 0 1 2 1 1 0 0 1 0 1 8

5 6/13/2014 Captures per net-hour 0.00 0.00 0.20 0.00 0.00 0.20 0.40 0.20 0.20 0.00 0.00 0.20 0.00 0.20 0.16
37 

MAPS 

period

Net Totals 

by

Date 1 2 3 4 5 6 7 8 9 10 11 12 13 14 date

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 50:00

Captures 1 0 0 3 0 1 0 0 1 0 0 0 0 1 7

6 6/20/2014 Captures per net-hour 0.20 0.00 0.00 0.60 0.00 0.20 0.00 0.00 0.20 0.00 0.00 0.00 0.00 0.20 0.66

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 50:00

Captures 0 0 0 1 1 0 0 1 0 0 0 1 2 0 6

7 7/1/2014 Captures per net-hour 0.00 0.00 0.00 0.20 0.20 0.00 0.00 0.20 0.00 0.00 0.00 0.20 0.40 0.00 0.12

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 50:00

Captures 0 0 0 0 0 0 0 0 1 3 2 0 1 1 8

8 7/11/2014 Captures per net-hour 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.20 0.60 0.40 0.00 0.20 0.20 0.16

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 3:40 5:00 4:30 50:00

Captures 1 0 2 7 1 1 4 1 1 0 1 0 0 5 24

9 7/25/2014 Captures per net-hour 0.20 0.00 0.40 1.40 0.20 0.20 0.80 0.20 0.20 0.00 0.20 0.00 0.00 1.11 0.48

Net-hours 4:30 4:30 4:20 4:30 4:30 4:30 4:30 4:30 4:30 4:30 4:30 4:30 4:30 4:30 44:50

Captures 0 1 2 0 1 0 1 0 0 0 0 0 1 3 9

10 8/1/2014 Captures per net-hour 0.00 0.22 0.46 0.00 0.22 0.00 0.22 0.00 0.00 0.00 0.00 0.00 0.22 0.67 0.20

Totals by net

Net-hours 64:30 64:30 64:20 64:30 64:30 64:30 64:30 64:30 64:30 64:30 64:30 63:10 64:30 64:00 901:00

Captures 3 9 12 23 9 10 16 8 22 9 8 5 22 25 181

Captures per net-hour 0.05 0.14 0.19 0.36 0.14 0.16 0.25 0.12 0.34 0.14 0.12 0.08 0.34 0.39 0.20
38 

Table 13. Capture rate by net and date during Monitoring Avian Productivity and Survivorship, Naval Base Coronado, Remote Training Site, Warner 

Springs, San Diego County, California, 2015.

[MAPS periods were established by the Institute for Bird Populations for program-wide consistency. In San Diego, banding began 3 periods early to encompass the 

earlier breeding season that occurs at southern latitudes. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl] 

MAPS 

period 

Net

Totals by 

Date 1 2 3 4 5 6 7 8 9 10 11 12 13 14 date

Net-hours 4:30 4:30 4:30 4:30 4:30 4:30 4:30 4:30 4:30 4:30 4:30 4:30 4:30 4:30 63:00

Captures 0 1 1 0 1 1 1 1 0 0 0 0 2 1 9

-3 4/3/2015 Captures per net-hour 0.00 0.22 0.22 0.00 0.22 0.22 0.22 0.22 0.00 0.00 1.00 2.00 3.00 4.00 0.14

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 70:00

Captures 0 0 1 0 0 1 7 1 0 0 1 0 1 3 15

-2 4/17/2015 Captures per net-hour 0.00 0.00 0.20 0.00 0.00 0.20 1.40 0.20 0.00 0.00 0.20 0.00 0.20 0.60 0.21

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 70:00

Captures 0 5 3 0 0 2 0 1 0 0 0 0 5 12 28

-1 4/24/2015 Captures per net-hour 0.00 1.00 0.60 0.00 0.00 0.40 0.00 0.20 0.00 0.00 0.00 0.00 1.00 2.40 0.40

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 70:00

Captures 0 1 3 1 0 2 0 1 1 3 0 0 2 3 17

1 5/1/2015 Captures per net-hour 0.00 0.20 0.60 0.20 0.00 0.40 0.00 0.20 0.20 0.60 0.00 0.00 0.40 0.60 0.24

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 70:00

Captures 0 0 2 1 0 1 1 0 1 0 1 2 0 4 13

2 5/19/2015 Captures per net-hour 0.00 0.00 0.40 0.20 0.00 0.20 0.20 0.00 0.20 0.00 0.20 0.40 0.00 0.80 0.19

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 70:00

Captures 0 1 2 0 0 0 0 0 1 0 1 1 0 1 7

3 5/29/2015 Captures per net-hour 0.00 0.20 0.40 0.00 0.00 0.00 0.00 0.00 0.20 0.00 0.20 0.20 0.00 0.20 0.10

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 70:00

Captures 0 0 0 0 0 0 0 0 0 12 0 0 0 1 13

4 6/5/2015 Captures per net-hour 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 2.40 0.00 0.00 0.00 0.20 0.19

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 70:00

Captures 0 1 0 0 0 0 6 0 0 0 0 0 0 0 7

5 6/12/2015 Captures per net-hour 0.00 0.20 0.00 0.00 0.00 0.00 1.20 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.10
39 

MAPS 

period 

Net

Totals by 

Date 1 2 3 4 5 6 7 8 9 10 11 12 13 14 date

Net-hours 5:30 5:30 5:30 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:30 72:00

Captures 0 0 4 0 0 4 1 3 0 0 0 1 4 1 18

6 6/26/2015 Captures per net-hour 0.00 0.00 0.73 0.00 0.00 0.80 0.20 0.60 0.00 0.00 0.00 0.20 0.80 0.18 0.25

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 70:00

Captures 1 0 0 0 0 1 1 0 0 3 0 1 1 1 9

7 7/2/2015 Captures per net-hour 0.20 0.00 0.00 0.00 0.00 0.20 0.20 0.00 0.00 0.60 0.00 0.20 0.20 0.20 0.13

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 70:00

Captures 0 0 0 0 0 0 2 1 1 6 0 0 1 0 11

8 7/17/2015 Captures per net-hour 0.00 0.00 0.00 0.00 0.00 0.00 0.40 0.20 0.20 1.20 0.00 0.00 0.20 0.00 0.16

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 70:00

Captures 0 1 1 0 1 0 0 1 1 1 0 1 1 3 11

9 7/24/2015 Captures per net-hour 0.00 0.20 0.20 0.00 0.20 0.00 0.00 0.20 0.20 0.20 0.00 0.20 0.20 0.60 0.16

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 70:00

Captures 0 0 1 0 0 0 1 0 0 2 1 0 1 0 6

10 7/31/2015 Captures per net-hour 0.00 0.00 0.20 0.00 0.00 0.00 0.20 0.00 0.00 0.40 0.20 0.00 0.20 0.00 0.09

Totals by net

Net-hours 65:00 65:00 65:00 64:30 64:30 64:30 64:30 64:30 64:30 64:30 64:30 64:30 64:30 65:00 905:00

Captures 1 10 18 2 2 12 20 9 5 27 4 6 18 30 164

Captures per net-hour 0.02 0.15 0.28 0.03 0.03 0.19 0.31 0.14 0.08 0.42 0.06 0.09 0.28 0.46 0.18
40 

Table 14. Capture rate by net and date during Monitoring Avian Productivity and Survivorship, Naval Base Coronado, Remote Training Site, Warner 

Springs, San Diego County, California, 2016.

[MAPS periods were established by the Institute for Bird Populations for program-wide consistency. In San Diego, banding began 3 periods early to encompass the 

earlier breeding season that occurs at southern latitudes. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl] 

MAPS 

period 

Net

Totals by 

Date 1 2 3 4 5 6 7 8 9 10 11 12 13 14 date

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 70:00

Captures 0 0 0 0 0 0 0 0 0 1 0 0 0 0 1

-2 4/15/2016 Captures per net-hour 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.20 0.00 0.00 0.00 0.00 0.01

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 70:00

Captures 0 0 0 0 0 0 3 0 0 0 0 2 3 2 10

-1 4/22/2016 Captures per net-hour 0.00 0.00 0.00 0.00 0.00 0.00 0.60 0.00 0.00 0.00 0.00 0.40 0.60 0.40 0.14

Net-hours 2:30 2:30 2:30 2:30 2:30 2:30 2:30 2:30 2:30 2:30 2:30 2:30 2:30 2:30 35:00

Captures 0 0 1 0 0 3 2 2 0 1 0 1 2 5 201 

1 5/6/2016 Captures per net-hour 0.00 0.00 0.40 0.00 0.00 1.20 0.80 0.80 0.00 0.40 0.00 0.40 0.80 2.00 0.57

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 70:00

Captures 0 0 0 0 0 0 2 1 1 0 1 0 1 2 8

2 5/12/2016 Captures per net-hour 0.00 0.00 0.00 0.00 0.00 0.00 0.40 0.20 0.20 0.00 0.20 0.00 0.20 0.40 0.11

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 70:00

Captures 1 0 2 1 0 4 2 3 0 1 1 0 2 0 17

3 5/27/2016 Captures per net-hour 0.20 0.00 0.40 0.20 0.00 0.80 0.40 0.60 0.00 0.20 0.20 0.00 0.40 0.00 0.24

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 70:00

Captures 0 1 0 0 0 0 0 1 0 1 0 1 1 2 7

4 6/3/2016 Captures per net-hour 0.00 0.20 0.00 0.00 0.00 0.00 0.00 0.20 0.00 0.20 0.00 0.20 0.20 0.40 0.10

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 70:00

Captures 0 0 0 0 0 1 0 0 6 0 3 0 1 3 14

5 6/17/2016 Captures per net-hour 0.00 0.00 0.00 0.00 0.00 0.20 0.00 0.00 1.20 0.00 0.60 0.00 0.20 0.60 0.20

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 4:40 5:00 5:00 69:40

Captures 0 2 1 0 0 2 0 0 2 2 0 0 2 2 13

6 6/23/2016 Captures per net-hour 0.00 0.40 0.20 0.00 0.00 0.40 0.00 0.00 0.40 0.40 0.00 0.00 0.40 0.40 0.19
41 

MAPS 

period 

Net

Totals by 

Date 1 2 3 4 5 6 7 8 9 10 11 12 13 14 date

Net-hours 3:30 3:30 3:30 3:30 3:30 3:30 3:30 3:30 3:30 3:30 3:30 3:30 3:30 3:30 49:00

Captures 1 0 1 2 0 0 2 1 0 0 1 0 1 1 10

7 7/1/2016 Captures per net-hour 0.29 0.00 0.29 0.57 0.00 0.00 0.57 0.29 0.00 0.00 0.29 0.00 0.29 0.29 0.20

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 70:00

Captures 0 0 0 0 0 1 2 0 0 0 0 1 1 0 5

8 7/14/2016 Captures per net-hour 0.00 0.00 0.00 0.00 0.00 0.20 0.40 0.00 0.00 0.00 0.00 0.20 0.20 0.00 0.07

Net-hours 4:10 4:10 4:10 4:10 4:10 4:10 4:10 4:10 4:10 4:10 4:10 4:10 4:10 4:10 58:20

Captures 0 0 0 0 0 0 4 0 1 1 4 1 0 2 13

9 7/22/2016 Captures per net-hour 0.00 0.00 0.00 0.00 0.00 0.00 0.96 0.00 0.24 0.24 0.96 0.24 0.00 0.48 0.22

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 70:00

Captures 0 0 0 0 0 6 0 0 0 0 0 0 3 4 13

10 8/5/2016 Captures per net-hour 0.00 0.00 0.00 0.00 0.00 1.20 0.00 0.00 0.00 0.00 0.00 0.00 0.60 0.80 0.19

Totals by net

Net-Hours 55:10 55:10 55:10 55:10 55:10 55:10 55:10 55:10 55:10 55:10 55:10 54:50 55:10 55:10 772:00

Captures 2 3 5 3 0 17 17 8 10 7 10 6 17 23 1311 

Captures per net-hour 0.04 0.05 0.09 0.05 0.00 0.31 0.31 0.15 0.18 0.13 0.18 0.11 0.31 0.42 0.17

1

Three birds captured but no net recorded.
42 

Table 15. Capture rate by net and date during Monitoring Avian Productivity and Survivorship, Naval Base Coronado, Remote Training Site, Warner 

Springs, San Diego County, California, 2017.

[MAPS periods were established by the Institute for Bird Populations for program-wide consistency. In San Diego, banding began 3 periods early to encompass the 

earlier breeding season that occurs at southern latitudes. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl] 

MAPS 

period 

Net

Totals by 

Date 1 2 3 4 5 6 7 8 9 10 11 12 13 14 date

Net-hours 5:00 5:00 5:00 5:10 5:10 5:10 5:10 5:10 5:10 5:10 5:10 5:10 5:10 5:00 71:40

Captures 3 5 0 3 0 0 0 0 0 1 3 1 0 2 18

-3 4/5/2017 Captures per net-hour 0.60 1.00 0.00 0.58 0.00 0.00 0.00 0.00 0.00 0.19 0.58 0.19 0.00 0.40 0.25

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 70:00

Captures 3 0 0 0 0 0 1 2 0 0 0 5 0 2 13

-2 4/12/2017 Captures per net-hour 0.60 0.00 0.00 0.00 0.00 0.00 0.20 0.40 0.00 0.00 0.00 1.00 0.00 0.40 0.19

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 70:00

Captures 0 1 1 0 0 0 0 0 0 0 0 8 0 0 10

-1 4/28/2017 Captures per net-hour 0.00 0.20 0.20 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 1.60 0.00 0.00 0.14

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 70:00

Captures 0 2 0 0 0 0 0 0 0 0 1 0 1 0 4

1 5/3/2017 Captures per net-hour 0.00 0.40 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.20 0.00 0.20 0.00 0.06

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 70:00

Captures 0 1 0 0 1 0 0 1 0 1 0 0 0 1 5

2 5/17/2017 Captures per net-hour 0.00 0.20 0.00 0.00 0.20 0.00 0.00 0.20 0.00 0.20 0.00 0.00 0.00 0.20 0.76

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 70:00

Captures 0 4 0 3 0 2 1 1 1 3 0 3 0 1 19

3 5/30/2017 Captures per net-hour 0.00 0.80 0.00 0.60 0.00 0.40 0.20 0.20 0.20 0.60 0.00 0.60 0.00 0.20 0.27

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 70:00

Captures 2 0 0 0 0 0 0 0 1 0 0 0 0 1 4

4 6/7/2017 Captures per net-hour 0.40 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.20 0.00 0.00 0.00 0.00 0.20 0.06

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 70:00

Captures 1 2 0 1 1 3 2 0 0 3 0 1 0 0 14

5 6/16/2017 Captures per net-hour 0.20 0.40 0.00 0.20 0.20 0.60 0.40 0.00 0.00 0.60 0.00 0.20 0.00 0.00 0.20
43 

MAPS 

period 

Net

Totals by 

Date 1 2 3 4 5 6 7 8 9 10 11 12 13 14 date

Net-hours 3:50 3:50 3:50 4:30 4:30 4:30 4:30 4:30 4:30 4:30 4:30 4:30 4:30 3:30 60:00

Captures 5 1 0 1 0 0 2 1 0 0 0 3 0 1 14

6 6/21/2017 Captures per net-hour 1.30 0.26 0.00 0.22 0.00 0.00 0.44 0.22 0.00 0.00 0.00 0.67 0.00 0.29 0.55

Net-hours 5:00 5:00 4:20 5:00 5:00 4:20 4:20 4:20 4:20 4:20 4:20 4:20 5:00 5:00 64:40

Captures 1 2 1 3 0 0 0 1 0 1 0 0 0 7 16

7 7/6/2017 Captures per net-hour 0.20 0.40 0.23 0.60 0.00 0.00 0.00 0.23 0.00 0.23 0.00 0.00 0.00 1.40 0.25

Net-hours 5:00 5:00 3:50 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 68:50

Captures 2 1 0 7 0 3 1 4 1 1 0 1 0 1 22

8 7/12/2017 Captures per net-hour 0.40 0.20 0.00 1.40 0.00 0.60 0.20 0.80 0.20 0.20 0.00 0.20 0.00 0.20 0.32

Net-hours 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 4:10 4:20 5:00 4:30 5:00 5:00 68:00

Captures 0 0 0 4 0 6 0 0 0 3 0 1 0 0 14

9 7/26/2017 Captures per net-hour 0.00 0.00 0.00 0.80 0.00 1.20 0.00 0.00 0.00 0.69 0.00 0.22 0.00 0.00 0.21

Net-hours 5:00 5:00 4:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 5:00 4:30 4:30 68:00

Captures 1 0 0 2 0 0 0 0 1 2 1 1 0 1 9

10 8/9/2017 Captures per net-hour 0.20 0.00 0.00 0.40 0.00 0.00 0.00 0.00 0.20 0.40 0.20 0.20 0.00 0.22 0.13

Net-hours 63:50 63:50 61:00 64:40 64:40 64:00 64:00 64:00 63:10 63:20 64:00 63:30 64:10 63:00 891:10

Captures 18 19 2 24 2 14 7 10 4 15 5 24 1 17 162

Totals by net Captures per net-hour 0.28 0.30 0.03 0.37 0.03 0.22 0.11 0.16 0.06 0.24 0.08 0.38 0.02 0.27 0.18
44 

Table 16. Number of captures by date during Monitoring Avian Productivity and Survivorship, Naval Base 

Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2013.

[See tables 2 and 3 for bird species codes. Species in italics are non-breeding neotropical migrants. Includes multiple 

captures for some individuals. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl] 

MAPS Period

Captures 

per 100 

net-hours 

(755:00 

total nethours)

-3 -2 -1 1 2 3 4 5 6 7 8 9 10

Date

Species

4-05-2013

4-12-2013

4-26-2013

5-03-2013

5-17-2013

5-24-2013

6-07-2013

6-14-2013

6-21-2013

7-04-2013

7-12-2013

7-26-2013

8-02-2013

Total

BCHU 0 0 0 0 0 0 0 0 1 0 0 0 0 1 0.13

ANHU 0 0 1 1 1 2 4 0 0 0 0 0 0 9 1.19

COHU 0 0 0 0 0 0 0 0 0 0 0 0 1 1 0.13

ALHU 1 0 0 0 0 0 0 0 0 0 0 0 0 1 0.13

ACWO 1 0 0 1 1 3 0 0 4 1 1 0 0 12 1.59

NUWO 0 0 0 0 0 0 0 0 0 0 0 2 3 5 0.66

DOWO 0 0 0 0 0 0 0 0 0 0 1 0 0 1 0.13

RSFL 0 1 0 1 0 0 0 0 0 0 0 0 0 2 0.26

GRFL 0 0 1 0 0 0 0 0 0 0 0 0 0 1 0.13

ATFL 0 2 1 1 1 0 1 1 0 0 1 0 0 8 1.06

WAVI 0 0 0 0 0 0 0 0 0 0 0 1 0 1 0.13

CASJ 3 1 0 1 0 0 0 0 0 1 2 0 0 8 1.06

OATI 0 2 2 3 0 0 0 1 1 0 1 0 1 11 1.46

BUSH 0 0 0 0 0 2 1 0 0 8 0 0 1 12 1.59

WBNU 0 0 0 0 0 0 1 0 0 0 1 0 1 3 0.40

BEWR 0 0 0 0 0 0 2 0 0 0 1 0 0 3 0.40

HOWR 2 3 0 1 0 1 1 3 1 0 0 0 0 12 1.59

SWTH 0 0 0 0 1 0 0 0 0 0 0 0 0 1 0.13

WREN 0 0 0 0 0 0 0 0 3 1 0 0 0 4 0.53

CATH 0 0 0 0 0 2 0 0 0 0 0 0 0 2 0.26

OCWA 0 0 1 1 0 0 0 0 0 0 0 0 1 3 0.40

NAWA 0 1 0 0 0 0 0 0 0 0 0 0 0 1 0.13

YEWA 0 0 1 0 0 1 0 0 0 0 0 0 0 2 0.26

MGWA 0 0 1 0 0 0 0 0 0 0 0 0 0 1 0.13

COYE 0 1 0 0 0 0 0 0 0 0 0 0 0 1 0.13

WIWA 0 0 1 1 2 0 0 0 0 0 0 0 0 4 0.53

WETA 0 0 0 0 1 0 0 0 0 0 0 0 0 1 0.13

SPTO 1 1 4 1 0 3 3 0 2 1 0 1 0 17 2.25

CALT 0 0 0 1 0 1 1 1 0 2 1 1 0 8 1.06
45 

MAPS Period

Captures 

per 100 

net-hours 

(755:00 

total nethours)

-3 -2 -1 1 2 3 4 5 6 7 8 9 10

Date

Species

4-05-2013

4-12-2013

4-26-2013

5-03-2013

5-17-2013

5-24-2013

6-07-2013

6-14-2013

6-21-2013

7-04-2013

7-12-2013

7-26-2013

8-02-2013

Total

BRSP 0 0 1 0 0 0 0 0 0 0 0 0 0 1 0.13

WCSP 1 1 0 0 0 0 0 0 0 0 0 0 0 2 0.26

BHGR 0 0 0 0 0 0 1 0 0 0 0 0 0 1 0.13

BUOR 0 0 0 4 0 0 0 1 0 0 0 0 0 5 0.66

Captures per day 9 13 1

4

17 7 15 15 7 12 14 9 5 8 145 19.21

Total species 6 9 1

0 12 6 8 9 5 6 6 8 4 6 33 4.37
46 

Table 17. Number of captures by date during Monitoring Avian Productivity and Survivorship, Naval Base 

Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2014.

[See tables 2 and 3 for bird species codes. Species in italics are non-breeding neotropical migrants. Includes multiple 

captures for some individuals. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl] 

MAPS period

Captures 

per 100 nethours (total 

901:00 nethours)

-3 -2 -1 1 2 3 4 5 6 7 8 9 10

Date

Species

4-04-2014

4-11-2014

4-25-2014

5-02-2014

5-16-2014

5-23-2014

6-06-2014

6-13-2014

6-20-2014

7-01-2014

7-11-2014

7-25-2014

8-01-2014

Total

CAQU 0 0 0 0 1 0 0 0 0 0 0 0 0 1 0.11

BCHU 0 0 0 0 0 0 0 0 0 0 0 1 0 1 0.11

ANHU 1 0 0 0 3 0 1 1 0 0 1 1 1 9 1.00

COHU 0 0 0 0 1 0 0 0 1 0 0 0 0 2 0.22

RUHU 0 0 0 0 0 0 0 0 0 0 0 1 0 1 0.11

ALHU 0 0 0 0 0 0 0 1 0 0 0 0 0 1 0.11

ACWO 0 0 0 0 1 0 0 0 0 0 0 1 1 3 0.33

NUWO 0 0 0 1 0 0 0 0 0 0 0 0 1 2 0.22

DOWO 0 0 0 0 0 0 0 1 0 0 1 0 0 2 0.22

WIFL 0 0 0 0 0 0 0 1 0 0 0 0 0 1 0.11

HAFL 2 0 0 0 0 0 0 0 0 0 0 0 0 2 0.22

PSFL 0 0 0 0 1 1 0 0 0 0 0 1 0 3 0.33

ATFL 0 0 0 0 0 1 0 0 1 0 0 1 0 3 0.33

WAVI 0 1 0 0 0 1 0 0 0 0 0 1 1 4 0.44

CASJ 1 1 0 1 0 0 0 1 1 2 0 0 0 7 0.78

OATI 0 1 0 0 1 1 0 0 0 1 1 1 0 6 0.67

BUSH 1 0 0 0 0 0 2 1 0 0 1 7 0 12 1.33

WBNU 0 0 2 0 1 1 1 1 0 0 0 1 0 7 0.78

BEWR 0 0 0 0 0 0 0 0 0 0 1 2 2 5 0.55

HOWR 1 5 0 1 1 1 1 0 1 0 0 1 0 12 1.33

WEBL 0 0 0 0 2 0 0 0 1 0 0 0 0 3 0.33

SWTH 0 0 0 0 0 1 0 0 0 0 0 0 0 1 0.11

HETH 1 1 0 0 0 0 0 0 0 0 0 0 0 2 0.22

WREN 0 1 0 0 0 0 0 0 0 0 0 0 0 1 0.11

EUST 0 0 0 0 1 0 0 0 0 0 0 0 0 1 0.11

OCWA 6 11 1 0 0 0 0 0 0 0 0 0 0 18 2.00

NAWA 1 3 0 0 0 0 0 0 0 0 0 0 0 4 0.44

YEWA 0 0 0 0 1 2 0 0 0 0 0 0 0 3 0.33

AUWA 2 0 0 0 0 0 0 0 0 0 0 0 0 2 0.22

MGWA 0 0 1 0 0 0 0 0 0 0 0 0 0 1 0.11

WIWA 1 0 5 4 7 0 0 0 0 0 0 0 0 17 1.89

WETA 0 0 1 0 0 0 0 0 0 0 0 1 0 2 0.22

SPTO 5 0 1 0 1 1 2 1 1 2 1 4 0 19 2.11

CALT 0 1 0 1 0 0 0 0 0 0 0 0 2 4 0.44

BTSP 0 0 0 0 0 0 0 0 1 0 0 0 0 1 0.11

BESP 0 0 0 0 0 0 0 0 0 0 1 0 0 1 0.11

SOSP 0 0 0 0 0 0 0 0 0 1 1 0 0 2 0.22

WCSP 0 1 0 2 0 0 0 0 0 0 0 0 0 3 0.33

ORJU 1 1 0 0 0 0 0 0 0 0 0 0 0 2 0.22
47 

MAPS period

Captures 

per 100 nethours (total 

901:00 nethours)

-3 -2 -1 1 2 3 4 5 6 7 8 9 10

Date

Species

4-04-2014

4-11-2014

4-25-2014

5-02-2014

5-16-2014

5-23-2014

6-06-2014

6-13-2014

6-20-2014

7-01-2014

7-11-2014

7-25-2014

8-01-2014

Total

BHGR 0 0 0 0 0 0 0 0 0 0 0 0 1 1 0.11

BUOR 0 1 2 0 3 0 1 0 0 0 0 0 0 7 0.78

LEGO 2 0 0 0 0 0 0 0 0 0 0 0 0 2 0.22

Captures per day 25 28 13 10 25 10 8 8 7 6 8 24 9 181 20.09

Total species 13 12 7 6 14 9 6 8 7 4 8 14 7 42 4.66
48 

Table 18. Number of captures by date during Monitoring Avian Productivity and Survivorship, Naval Base 

Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2015.

[See tables 2 and 3 for bird species codes. Species in italics are non-breeding neotropical migrants. Includes multiple 

captures of some individuals. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl] 

MAPS period

Captures 

per 100 

net-hours 

(total 

905:00 nethours)

-3 -2 -1 1 2 3 4 5 6 7 8 9 10

Date

Species

4-03-2015

4-17-2015

4-24-2015

5-01-2015

5-19-2015

5-29-2015

6-05-2015

6-12-2015

6-26-2015

7-02-2015

7-17-2015

7-24-2015

7-31-2015

Total

CAQU 0 0 0 0 0 0 0 0 0 1 0 0 0 1 0.11

ANHU 0 1 0 0 0 3 1 0 0 0 0 1 0 6 0.66

COHU 0 0 0 1 0 0 0 0 1 3 1 1 1 8 0.88

CAHU 0 0 0 1 0 0 0 0 0 0 0 0 0 1 0.11

RUHU 1 1 0 1 0 0 0 0 0 0 0 0 0 3 0.33

ALHU 0 0 0 1 0 0 0 0 0 0 0 0 0 1 0.11

ACWO 0 0 0 1 0 0 0 0 1 0 0 0 0 2 0.22

NUWO 0 0 0 0 0 0 0 0 1 0 2 1 0 4 0.44

RSFL 0 0 1 0 0 0 0 0 0 0 0 0 0 1 0.11

WEWP 0 0 1 0 0 0 0 0 0 0 0 0 0 1 0.11

WIFL 0 0 0 0 1 0 0 0 0 0 0 0 0 1 0.11

PSFL 0 0 1 1 1 0 0 0 0 0 1 1 0 5 0.55

ATFL 0 2 0 0 0 0 0 0 1 1 0 2 0 6 0.66

WAVI 0 0 12 1 0 0 0 0 0 0 0 0 0 13 1.44

CASJ 0 0 0 0 1 0 0 0 0 1 0 0 0 2 0.22

OATI 0 1 0 0 0 1 1 0 2 2 0 0 1 8 0.88

BUSH 1 0 0 0 0 0 10 6 8 0 6 0 1 32 3.54

WBNU 0 0 0 0 1 0 0 0 0 0 0 1 0 2 0.22

BEWR 0 0 0 0 0 0 0 0 3 0 1 1 2 7 0.77

HOWR 0 0 0 0 0 0 1 0 1 0 0 1 0 3 0.33

RCKI 1 0 0 0 0 0 0 0 0 0 0 0 0 1 0.11

SWTH 0 0 0 0 3 0 0 0 0 0 0 0 0 3 0.33

CATH 0 0 0 0 0 1 0 0 0 0 0 0 0 1 0.11

OCWA 0 0 1 0 0 0 0 0 0 0 0 0 0 1 0.11

NAWA 0 0 1 0 0 0 0 0 0 0 0 0 0 1 0.11

YEWA 0 0 2 0 3 0 0 0 0 0 0 0 0 5 0.55

AUWA 0 2 0 1 0 0 0 0 0 0 0 0 0 3 0.33

MGWA 0 0 1 0 0 0 0 0 0 0 0 0 0 1 0.11

WIWA 0 0 7 6 1 0 0 0 0 0 0 0 0 14 1.55

SPTO 1 2 1 1 1 1 0 1 0 0 0 1 1 10 1.10

CALT 0 0 0 1 0 0 0 0 0 1 0 0 0 2 0.22

WCSP 4 2 0 1 0 0 0 0 0 0 0 0 0 7 0.77

GCSP 0 1 0 0 0 0 0 0 0 0 0 0 0 1 0.11

BHGR 0 2 0 0 0 0 0 0 0 0 0 0 0 2 0.22
49 

MAPS period

Captures 

per 100 

net-hours 

(total 

905:00 nethours)

-3 -2 -1 1 2 3 4 5 6 7 8 9 10

Date

Species

4-03-2015

4-17-2015

4-24-2015

5-01-2015

5-19-2015

5-29-2015

6-05-2015

6-12-2015

6-26-2015

7-02-2015

7-17-2015

7-24-2015

7-31-2015

Total

BUOR 1 1 0 0 1 1 0 0 0 0 0 0 0 4 0.44

HOFI 0 0 0 0 0 0 0 0 0 0 0 1 0 1 0.11

Captures per day 9 15 28 17 13 7 13 7 18 9 11 11 6 164 18.12

Total species 6 10 10 12 9 5 4 2 8 6 5 10 5 36 3.98
50 

Table 19. Number of captures by date during Monitoring Avian Productivity and Survivorship, Naval Base 

Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2016.

[See tables 2 and 3 for bird species codes. Species in italics are non-breeding neotropical migrants. Includes multiple 

captures of some individuals. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl] 

MAPS period

Captures per 

100 net-hours 

(772:00 nethours)

-2 -1 1 2 3 4 5 6 7 8 9 10

Date

Species

4-15-2016

4-22-2016

5-06-2016

5-12-2016

5-27-2016

6-03-2016

6-17-2016

6-23-2016

7-01-2016

7-14-2016

7-22-2016

8-05-2016

Total

CAQU 0 0 0 0 3 0 0 0 0 0 0 0 3 0.39

MODO 0 0 0 0 0 0 1 0 0 0 0 0 1 0.13

BCHU 0 0 0 3 0 0 0 0 0 1 0 0 4 0.52

ANHU 0 0 0 0 0 0 0 3 0 0 0 0 3 0.39

COHU 0 0 0 0 0 0 1 0 0 0 0 0 1 0.13

NUWO 0 0 0 0 0 0 0 2 0 0 0 1 2 0.26

DOWO 0 0 0 0 0 0 2 1 0 0 0 0 3 0.39

PSFL 0 0 1 0 0 0 0 0 0 0 0 0 1 0.13

ATFL 1 0 1 0 0 1 1 0 6 1 0 0 11 1.42

WEKI 0 0 0 0 0 0 0 0 1 0 0 0 1 0.13

LBVI 0 0 0 1 0 0 0 0 0 0 0 0 1 0.13

WAVI 0 0 2 0 0 0 0 0 0 0 0 0 2 0.26

CASJ 0 0 0 0 2 0 1 1 0 0 0 0 4 0.52

OATI 0 0 0 1 0 1 4 1 1 1 4 0 9 1.17

BUSH 0 2 0 0 6 0 0 0 0 0 7 8 8 1.04

WBNU 0 0 0 0 1 0 0 0 0 0 0 0 1 0.13

BEWR 0 2 1 1 0 2 1 2 1 1 0 1 11 1.42

HOWR 0 0 0 0 1 0 1 2 0 0 0 0 4 0.52

SWTH 0 0 1 0 0 0 0 0 0 0 0 0 1 0.13

CATH 0 0 0 0 1 2 1 0 0 1 0 0 5 0.65

OCWA 0 1 0 0 0 0 0 0 0 0 0 0 1 0.13

YEWA 0 0 2 0 0 0 0 0 0 0 0 0 2 0.26

WIWA 0 3 11 1 0 0 0 0 0 0 0 0 15 1.94

SPTO 0 0 1 0 1 0 0 1 0 0 0 0 3 0.39

LASP 0 0 0 0 0 0 0 0 1 0 0 3 1 0.13

SOSP 0 0 0 0 0 1 0 0 0 0 0 0 1 0.13

WCSP 0 2 0 1 0 0 0 0 0 0 0 0 3 0.39

BHGR 0 0 0 0 0 0 0 0 0 0 2 0 0 0.00

BUOR 0 0 0 0 2 0 1 0 0 0 0 0 3 0.39

Captures per day 1 10 20 8 17 7 14 13 10 5 13 13 131 16.97

Total species 1 5 8 6 8 5 10 8 5 5 3 4 28 3.63
51 

Table 20. Number of captures by date during Monitoring Avian Productivity and Survivorship, Naval Base 

Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2017.

[See tables 2 and 3 for bird species codes. Species in italics are non-breeding neotropical migrants. Includes multiple 

captures of some individuals. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl] 

MAPS period

Captures 

per 100 nethours 

(891:10 nethours)

-3 -2 -1 1 2 3 4 5 6 7 8 9 10

Date

Species

4/05/20174/12/20174/28/20175/03/20175/17/20175/30/20176/07/20176/16/20176/21/20177/06/20177/12/20177/26/20178/09/2017

Total

CAQU 0 0 0 0 1 0 0 0 0 0 0 0 0 1 0.11

BCHU 0 0 0 0 0 0 0 0 0 0 0 0 1 1 0.11

ANHU 0 1 0 0 0 1 3 0 2 1 0 2 0 10 1.12

ALHU 0 0 0 0 0 0 0 0 0 0 0 0 1 1 0.11

ACWO 0 0 0 0 0 0 0 0 0 0 1 0 0 1 0.11

NUWO 0 0 0 0 0 0 0 2 0 0 1 0 0 3 0.34

RSFL 2 0 0 0 0 0 0 0 1 0 0 0 0 3 0.34

ATFL 0 0 0 0 2 0 0 1 0 0 1 0 0 4 0.45

HUVI 0 0 0 0 0 0 0 0 1 1 0 0 0 2 0.22

CASJ 0 0 0 1 1 2 0 1 0 0 0 0 0 5 0.56

OATI 0 1 0 0 0 3 0 0 1 4 1 1 0 11 1.23

BUSH 0 0 5 1 0 5 0 0 5 1 13 7 3 40 4.49

WBNU 0 0 0 0 0 2 0 0 0 0 0 2 0 4 0.45

BEWR 0 0 0 0 0 0 0 1 1 4 2 0 2 10 1.12

HOWR 2 0 0 0 1 5 0 4 1 3 1 2 1 20 2.24

WEBL 0 1 2 0 0 0 0 2 2 2 1 0 0 10 1.12

CATH 0 0 0 1 0 0 0 1 0 0 1 0 0 3 0.34

YEWA 0 0 1 0 0 0 0 0 0 0 0 0 0 1 0.11

AUWA 8 4 0 0 0 0 0 0 0 0 0 0 0 12 1.35

COYE 0 2 0 0 0 0 0 0 0 0 0 0 0 2 0.22

WIWA 0 2 2 0 0 0 0 0 0 0 0 0 0 4 0.45

SPTO 1 0 0 0 0 0 1 0 0 0 0 0 0 2 0.22

CALT 1 0 0 0 0 0 0 0 0 0 0 0 1 2 0.22

BESP 0 0 0 0 0 1 0 0 0 0 0 0 0 1 0.11

SOSP 1 0 0 0 0 0 0 0 0 0 0 0 0 1 0.11

LISP 1 0 0 0 0 0 0 0 0 0 0 0 0 1 0.11

WCSP 2 2 0 0 0 0 0 0 0 0 0 0 0 4 0.45

BHGR 0 0 0 1 0 0 0 2 0 0 0 0 0 3 0.34

Captures per day 18 13 10 4 5 19 4 14 14 16 22 14 9 117 13.13

Total species 8 7 4 4 4 7 2 8 8 7 9 5 6 28 3.14
52 

Table 21. Sex and age of individuals (banded and unbanded) captured during Monitoring Avian Productivity and Survivorship, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California, 2013.

[See tables 2 and 3 for bird species codes. Species in italics are non-breeding neotropical migrants. Age: HY=hatching-year, AHY=after-hatching-year, SY=secondyear, ASY=after-second-year, TY=third year, ATY=after-third-year, I=indeterminable age. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl] 

Female Male Unknown sex

Age Female Age Male Age Unknown Species

Species HY AHY SY ASY TY ATY total HY AHY SY ASY TY ATY total HY AHY SY ASY I total total

BCHU 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 1

ANHU 1 3 0 0 0 0 4 3 2 0 0 0 0 5 0 0 0 0 0 0 9

COHU 0 1 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1

ALHU 0 1 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1

ACWO 0 1 1 0 1 1 4 0 1 0 0 1 4 6 1 0 0 0 0 1 11

NUWO 0 0 0 0 0 0 0 2 0 0 1 0 0 3 0 0 0 0 0 0 3

DOWO 0 0 0 0 0 0 0 1 0 0 0 0 0 1 0 0 0 0 0 0 1

RSFL 0 0 1 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1

GRFL 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 1 1

ATFL 0 2 0 0 0 0 2 0 0 0 0 0 0 0 0 1 1 4 0 6 8

WAVI 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 1 1

CASJ 0 0 1 0 0 0 1 0 0 0 0 0 0 0 0 4 0 3 0 7 8

OATI 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 6 1 0 0 8 8

BUSH 6 0 0 0 0 0 6 0 2 0 0 0 0 2 3 0 0 0 1 4 12

WBNU 0 2 0 0 0 0 2 0 0 0 0 0 0 0 0 0 0 0 1 1 3

BEWR 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 1 0 0 0 3 3

HOWR 0 2 0 0 0 0 2 0 1 0 0 0 0 1 1 5 0 0 0 6 9

SWTH 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 1 1

WREN 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 1 0 0 1 4 4

CATH 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 0 0 0 2 2

OCWA 0 2 0 1 0 0 3 0 0 0 0 0 0 0 0 0 0 0 0 0 3

NAWA 0 0 0 0 0 0 0 0 0 0 1 0 0 1 0 0 0 0 0 0 1

YEWA 0 0 1 0 0 0 1 0 0 0 1 0 0 1 0 0 0 0 0 0 2

MGWA 0 0 0 0 0 0 0 0 0 0 1 0 0 1 0 0 0 0 0 0 1

COYE 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1

WIWA 0 1 0 1 0 0 2 0 0 0 2 0 0 2 0 0 0 0 0 0 4

WETA 0 0 0 0 0 0 0 0 0 0 1 0 0 1 0 0 0 0 0 0 1
53 

Female Male Unknown sex

Age Female Age Male Age Unknown Species

Species HY AHY SY ASY TY ATY total HY AHY SY ASY TY ATY total HY AHY SY ASY I total total

SPTO 0 1 2 2 0 0 5 0 3 1 3 0 0 7 1 0 0 0 0 1 13

CALT 0 0 0 0 0 0 0 0 1 0 0 0 0 1 4 1 0 0 0 5 6

BRSP 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 1 1

WCSP 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 0 0 2 2

BHGR 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1

BUOR 0 0 1 2 0 0 3 0 0 2 0 0 0 2 0 0 0 0 0 0 5

Total 7 16 7 8 1 1 40 6 10 3 10 1 4 34 16 21 5 9 4 55 129
54 

Table 22. Sex and age of individuals (banded and unbanded) captured during Monitoring Avian Productivity and Survivorship, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California, 2014.

[See tables 2 and 3 for bird species codes. Species in italics are non-breeding neotropical migrants. Age: HY=hatching-year, AHY=after-hatching-year, SY=secondyear, ASY=after-second-year, TY=third year, ATY=after-third-year, I=indeterminable age. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl] 

Female Male Unknown sex

Age Female Age Male Age Unknown Species

Species HY AHY SY ASY ATY total HY AHY SY ASY total HY AHY SY ASY I total total

CAQU 0 0 0 0 0 0 0 1 0 0 1 0 0 0 0 0 0 1

BCHU 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 1 1

ANHU 2 2 0 0 0 4 3 1 1 0 5 0 0 0 0 0 0 9

COHU 0 1 0 0 0 1 1 0 0 0 1 0 0 0 0 0 0 2

RUHU 0 0 0 0 0 0 1 0 0 0 1 0 0 0 0 0 0 1

ALHU 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 1

ACWO 0 0 0 1 0 1 0 1 0 0 1 1 0 0 0 0 1 3

NUWO 0 0 0 0 1 1 1 0 0 0 1 0 0 0 0 0 0 2

DOWO 0 0 0 0 0 0 0 1 0 0 1 0 0 0 0 0 0 1

WIFL 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 1 1

HAFL 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 0 2 2

PSFL 0 0 0 0 0 0 0 0 0 0 0 1 1 1 0 0 3 3

ATFL 0 0 0 0 0 0 0 0 0 0 0 0 1 1 1 0 3 3

WAVI 0 0 0 0 0 0 0 0 0 0 0 0 3 0 1 0 4 4

CASJ 0 1 0 1 0 2 0 0 0 0 0 2 1 1 1 0 5 7

OATI 0 0 0 0 0 0 0 0 0 0 0 1 0 2 3 0 6 6

BUSH 2 2 0 0 0 4 0 2 0 0 2 3 0 0 0 0 3 9

WBNU 0 0 0 3 0 3 0 0 0 3 3 0 0 0 0 0 0 6

BEWR 0 1 0 0 0 1 0 0 0 0 0 3 0 0 0 0 3 4

HOWR 0 1 0 0 0 1 0 1 0 1 2 1 2 0 2 0 5 8

WEBL 0 0 0 1 0 1 0 0 0 1 1 1 0 0 0 0 1 3

SWTH 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 1 1

HETH 0 0 0 0 0 0 0 0 0 0 0 0 1 0 1 0 2 2

WREN 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 1 1

EUST 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 1 1

OCWA 0 0 1 6 0 7 0 0 3 5 8 0 0 0 3 0 3 18

NAWA 0 0 0 0 0 0 0 0 0 4 4 0 0 0 0 0 0 4

YEWA 0 0 3 0 0 3 0 0 0 0 0 0 0 0 0 0 0 3
55 

Female Male Unknown sex

Age Female Age Male Age Unknown Species

Species HY AHY SY ASY ATY total HY AHY SY ASY total HY AHY SY ASY I total total

AUWA 0 0 0 2 0 2 0 0 0 0 0 0 0 0 0 0 0 2

MGWA 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 1

WIWA 0 3 5 2 0 10 0 0 4 1 5 0 0 0 1 1 2 17

WETA 0 0 0 1 0 1 0 1 0 0 1 0 0 0 0 0 0 2

SPTO 1 3 0 3 0 7 0 1 1 4 6 4 0 0 0 0 4 17

CALT 0 0 0 0 0 0 0 0 0 0 0 1 2 0 1 0 4 4

BTSP 0 0 0 0 0 0 0 0 0 1 1 0 0 0 0 0 0 1

BESP 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 1 1

SOSP 0 0 0 0 0 0 0 0 0 0 0 2 0 0 0 0 2 2

WCSP 0 0 0 0 0 0 0 0 0 0 0 0 2 1 0 0 3 3

ORJU 0 0 2 0 0 2 0 0 0 0 0 0 0 0 0 0 0 2

BHGR 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 1

BUOR 0 0 0 3 0 3 0 0 0 2 2 0 0 0 0 0 0 5

LEGO 0 0 0 1 0 1 0 0 0 1 1 0 0 0 0 0 0 2

Total 5 14 12 25 1 57 6 9 9 23 47 22 16 6 17 2 63 167
56 

Table 23. Sex and age of individuals (banded and unbanded) captured during Monitoring Avian Productivity and Survivorship, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California, 2015.

[See tables 2 and 3 for bird species codes. Species in italics are non-breeding neotropical migrants. Age: HY=hatching-year, AHY=after-hatching-year, SY=secondyear, ASY=after-second-year, ATY=after-third-year, I=indeterminable age. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl] 

Female Male Unknown sex

Age Female Age Male Age Unknown Species

Species HY AHY SY ASY ATY I total HY AHY SY ASY total HY AHY SY ASY total total

CAQU 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 1 1

ANHU 1 1 0 0 0 1 3 3 0 0 0 3 0 0 0 0 0 6

COHU 3 1 0 0 0 0 4 3 1 0 0 4 0 0 0 0 0 8

CAHU 0 1 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 1

RUHU 0 1 0 0 0 0 1 0 2 0 0 2 0 0 0 0 0 3

ALHU 0 1 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 1

ACWO 0 1 0 0 1 0 2 0 0 0 0 0 0 0 0 0 0 2

NUWO 0 0 1 1 0 0 2 1 0 0 0 1 0 0 0 0 0 3

RSFL 0 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 1

WEWP 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 1 1

WIFL 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 1 1

PSFL 0 1 0 0 0 0 1 0 0 0 0 0 0 2 2 0 4 5

ATFL 0 0 0 1 0 0 1 0 1 0 0 1 0 1 1 2 4 6

WAVI 0 0 0 0 0 0 0 0 0 0 0 0 0 2 2 9 13 13

CASJ 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 2 2

OATI 0 0 0 0 0 0 0 0 0 0 0 0 1 3 1 1 6 6

BUSH 17 0 0 3 0 1 21 0 1 0 0 1 8 0 0 0 8 30

WBNU 0 0 0 0 0 0 0 0 0 0 2 2 0 0 0 0 0 2

BEWR 0 0 0 0 0 0 0 0 0 0 0 0 4 0 0 0 4 4

HOWR 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 1 2 2

RCKI 0 1 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 1

SWTH 0 0 0 0 0 0 0 0 0 0 0 0 0 3 0 0 3 3

CATH 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 1 1

OCWA 0 1 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 1

NAWA 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 1 1

YEWA 0 0 2 1 0 0 3 0 0 0 2 2 0 0 0 0 0 5

AUWA 0 1 0 1 0 0 2 0 0 1 0 1 0 0 0 0 0 3

MGWA 0 0 0 0 0 0 0 0 0 1 0 1 0 0 0 0 0 1
57 

Female Male Unknown sex

Age Female Age Male Age Unknown Species

Species HY AHY SY ASY ATY I total HY AHY SY ASY total HY AHY SY ASY total total

WIWA 0 2 1 1 0 0 4 0 0 3 7 10 0 0 0 0 0 14

SPTO 1 0 0 3 0 0 4 0 0 0 3 3 0 0 0 0 0 7

CALT 0 0 0 0 0 0 0 0 1 0 0 1 1 0 0 0 1 2

WCSP 0 0 0 0 0 0 0 0 0 0 0 0 0 3 4 0 7 7

GCSP 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 1 1

BHGR 0 0 0 1 0 0 1 0 0 0 1 1 0 0 0 0 0 2

BUOR 0 0 0 1 0 0 1 0 0 0 2 2 0 0 0 0 0 3

HOFI 0 1 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 1

Total 22 13 4 13 2 2 56 7 6 5 17 35 17 17 12 14 60 151
58 

Table 24. Sex and age of individuals (banded and unbanded) captured during Monitoring Avian Productivity and Survivorship, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California, 2016.

[See tables 2 and 3 for bird species codes. Species in italics are non-breeding neotropical migrants. Age: HY=hatching-year, AHY=after-hatching-year, SY=secondyear, ASY=after-second-year, ATY=after-third-year, I=indeterminable age. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl] 

Female Male Unknown sex

Age Female Age Male Age Unknown Species

Species HY AHY SY ASY total HY AHY SY ASY ATY total HY AHY SY ASY I total total

CAQU 0 0 0 0 0 0 0 0 0 0 0 3 0 0 0 0 3 3

MODO 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 1 1

BCHU 0 1 0 0 1 0 0 0 0 0 0 3 0 0 0 0 3 4

ANHU 0 2 0 0 2 1 0 0 0 0 1 0 0 0 0 0 0 3

COHU 0 0 0 0 0 1 0 0 0 0 1 0 0 0 0 0 0 1

NUWO 0 0 0 0 0 2 0 0 0 1 3 0 0 0 0 0 0 3

DOWO 0 0 0 0 0 0 0 0 0 0 0 2 0 0 0 0 2 2

PSFL 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 1 1

ATFL 0 1 2 1 4 0 0 0 1 0 1 1 2 0 1 0 4 9

WEKI 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 1

LBVI 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 1 1

WAVI 0 0 0 0 0 0 0 0 0 0 0 0 2 0 0 0 2 2

CASJ 0 0 0 0 0 0 0 0 0 0 0 2 1 0 1 0 4 4

OATI 0 0 0 0 0 0 0 0 0 0 0 7 0 2 0 0 9 9

BUSH 8 2 0 1 11 0 1 0 0 0 1 7 0 0 0 2 9 21

WBNU 0 0 0 0 0 0 0 0 1 0 1 0 0 0 0 0 0 1

BEWR 0 2 0 0 2 0 0 0 0 0 0 4 0 1 0 0 5 7

HOWR 0 1 0 0 1 0 0 0 0 0 0 2 1 0 0 0 3 4

SWTH 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 1 1

CATH 0 0 0 0 0 0 0 0 0 0 0 2 1 0 1 0 4 4

OCWA 0 0 0 0 0 0 1 0 0 0 1 0 0 0 0 0 0 1

YEWA 0 0 0 0 0 0 2 0 0 0 2 0 0 0 0 0 0 2

WIWA 0 6 0 0 6 0 3 0 3 0 6 0 3 0 0 0 3 15

SPTO 0 1 0 0 1 0 1 0 1 0 2 0 0 0 0 0 0 3

LASP 0 0 0 0 0 0 0 0 0 0 0 4 0 0 0 0 4 4

SOSP 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 1 1

WCSP 0 0 0 0 0 0 0 0 0 0 0 0 3 0 0 0 3 3

BHGR 0 0 0 1 1 0 0 0 1 0 1 0 0 0 0 0 0 2
59 

Female Male Unknown sex

Age Female Age Male Age Unknown Species

Species HY AHY SY ASY total HY AHY SY ASY ATY total HY AHY SY ASY I total total

BUOR 0 1 0 0 1 1 0 1 0 0 2 0 0 0 0 0 0 3

Total 8 17 3 3 31 5 8 1 7 1 22 38 17 3 3 2 63 116
60 

Table 25. Sex and age of individuals (banded and unbanded) captured during Monitoring Avian Productivity and Survivorship, Warner Springs, San Diego 

County, California, 2017.

[See tables 2 and 3 for bird species codes. Species in italics are non-breeding neotropical migrants. Age: HY=hatching-year, AHY=after-hatching-year, SY=secondyear, ASY=after-second-year, ATY=after-third-year, I=indeterminable age. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl] 

Female Male Unknown sex

Age Female Age Male Age Unknown Species

Species HY AHY SY ASY ATY Total HY AHY SY ASY ATY Total HY AHY SY ASY I total total

CAQU 0 1 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 1

BCHU 0 0 0 0 0 0 0 1 0 0 0 1 0 0 0 0 0 0 1

ANHU 3 4 0 0 0 7 1 1 0 0 0 2 0 0 0 0 0 0 9

ALHU 0 0 0 0 0 0 1 0 0 0 0 1 0 0 0 0 0 0 1

ACWO 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 1

NUWO 0 0 0 0 0 0 2 0 1 0 0 3 0 0 0 0 0 0 3

RSFL 1 0 0 0 1 2 0 0 0 0 1 1 0 0 0 0 0 0 3

ATFL 0 0 0 2 0 2 0 0 0 0 0 0 0 0 0 0 0 0 2

HUVI 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 1 1

CASJ 0 0 0 2 0 2 0 0 0 0 0 0 2 0 0 1 0 3 5

OATI 0 0 0 1 0 1 0 0 0 0 0 0 7 0 1 2 0 10 11

BUSH 9 3 0 0 0 12 0 2 0 0 0 2 19 0 0 0 3 22 36

WBNU 0 0 0 0 0 0 2 0 0 2 0 4 0 0 0 0 0 0 4

BEWR 0 0 0 0 0 0 0 0 0 0 0 0 5 0 0 1 1 7 7

HOWR 0 1 0 0 0 1 0 0 0 0 0 0 8 3 0 0 1 12 13

WEBL 0 1 0 0 0 1 0 0 1 1 0 2 5 0 0 0 0 5 8

CATH 0 0 0 0 0 0 0 0 0 0 0 0 2 1 0 0 0 3 3

YEWA 0 0 0 0 0 0 0 0 0 1 0 1 0 0 0 0 0 0 1

AUWA 0 2 1 2 0 5 0 4 1 2 0 7 0 0 0 0 0 0 12

COYE 0 0 1 1 0 2 0 0 0 0 0 0 0 0 0 0 0 0 2

WIWA 0 0 0 2 0 2 0 0 1 1 0 2 0 0 0 0 0 0 4

SPTO 0 0 0 1 0 1 0 0 0 1 0 1 0 0 0 0 0 0 2

CALT 0 0 0 0 0 0 0 1 0 0 0 1 1 0 0 0 0 1 2

SAGS 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 1 1

SOSP 0 0 0 0 0 0 0 1 0 0 0 1 0 0 0 0 0 0 1

LISP 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 1 1

WCSP 0 0 0 0 0 0 0 0 0 0 0 0 0 1 3 0 0 4 4

BHGR 1 0 0 0 0 1 0 0 1 1 0 2 0 0 0 0 0 0 3
61 

Female Male Unknown sex

Age Female Age Male Age Unknown Species

Species HY AHY SY ASY ATY Total HY AHY SY ASY ATY Total HY AHY SY ASY I total total

Total 14 12 3 11 1 41 6 10 5 9 1 31 51 5 5 4 5 70 142
62 

Table 26. Logistic regression models for the effect of year on survivorship for four resident bird species captured 

during Monitoring Avian Productivity and Survivorship, Naval Base Coronado, Remote Training Site, Warner 

Springs, San Diego County, California 2013–17.

[Models are ranked from best to worst based on Akaike’s information criteria for small samples (AICC), ΔAICC, and 

Akaike weights (w). AICC is based on −2 x loge likelihood (L) and the number of parameters (K) in the model. Parameter 

estimates (β), standard error (SE), odds ratios and 95 percent confidence intervals (CI) for the best supported model 

explaining adult survivorship. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl] 

a. Oak Titmouse

Model Deviance # Parameters AICC ΔAICC AICC Weight

Constant 

survivorship

9.4 3 42.8 0 0.94

Year effect 6.0 6 48.2 5.5 0.06

b. Bushtit

Model Deviance # Parameters AICC ΔAICC AICC Weight

Constant 

survivorship

9.2 3 26.7 0 0.75

Year effect 4.4 6 28.6 2.2 0.25

c. House Wren

Model Deviance # Parameters AICC ΔAICC AICC Weight

Constant 

survivorship

4.7 3 23.3 0 0.99

Year effect 3.0 6 32.1 8.9 0.01

d. Spotted Towhee

Model Deviance # Parameters AICC ΔAICC AICC Weight

Constant 

survivorship

8.5 3 50.7 0 0.98

Year effect 8.1 6 58.4 7.7 0.02
63 

Figure 3. Number of individuals captured per species during Monitoring Avian Productivity and Survivorship, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2013. Colored bars 

represent the 11 most commonly captured resident species from 2013 to 2017. See tables 2 and 3 for bird 

species codes. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl. 

Figure 4. Number of individuals captured per species during Monitoring Avian Productivity and Survivorship,

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2014. Colored bars 

represent the 11 most commonly captured resident species from 2013 to 2017. See tables 2 and 3 for bird 

species codes. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl. 
64 

Figure 5. Number of individuals captured per species during Monitoring Avian Productivity and Survivorship, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2015. Colored bars 

represent the 11 most commonly captured resident species from 2013 to 2017. See tables 2 and 3 for bird 

species codes. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl. 

Figure 6. Number of individuals captured per species during Monitoring Avian Productivity and Survivorship, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2016. Colored bars 

represent the 11 most commonly captured resident species from 2013 to 2017. See tables 2 and 3 for bird 

species codes. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl. 
65 

Figure 7. Number of individuals captured per species during Monitoring Avian Productivity and Survivorship, 

Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2017. Colored bars 

represent the 11 most commonly captured resident species from 2013 to 2017. See tables 2 and 3 for bird 

species codes. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl. 
66 

Figure 8. Annual productivity for the ten most commonly captured resident species during Monitoring Avian 

Productivity and Survivorship, Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, 

California, 2013–17. Dotted lines connect missing years when productivity was not calculated because no adults 

were captured of this species this year. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl. 
67 

Figure 9. Adult survivorship calculated with Program MARK for four commonly captured resident species during 

Monitoring Avian Productivity and Survivorship, Naval Base Coronado, Remote Training Site, Warner Springs, 

San Diego County, California, 2013–17. Data collected by USGS and available from 

http://www.pwrc.usgs.gov/bbl. 
68 

Figure 10. Adult annual capture rate (individuals captured per 100 net-hours) of the 11 most commonly captured 

resident species during Monitoring Avian Productivity and Survivorship, Naval Base Coronado, Remote Training 

Site, Warner Springs, San Diego County, California, 2013–17. Data collected by USGS and available from 

http://www.pwrc.usgs.gov/bbl. 
69 

Table 27. Number of migrant birds captured and banded during fall migration, Naval Base Coronado, Remote Training Site, Warner Springs, San Diego 

County, California, 2013–16.

[See tables 2 and 3 for bird species codes. Includes multiple captures of some individuals. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl] 

Total captures Total number of individuals 

captured New individuals banded Recapture year / Year originally banded

Year Year Year 2014 2015 2016

Species 2013 2014 2015 2016 Total 2013 2014 2015 2016 Total 2013 2014 2015 2016 Total 2013 2013 2014 2013 2015 Total

SSHA 0 0 1 0 1 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0

BCHU 2 1 2 1 6 2 1 2 1 6 0 0 0 0 0 0 0 0 0 0 0

COHU 0 0 2 2 4 0 0 1 2 3 0 0 0 0 0 0 0 0 0 0 0

RUHU 1 0 0 0 1 1 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0

RNSA 0 3 0 0 3 0 3 0 0 3 0 3 0 0 3 0 0 0 0 0 0

RBSA 0 1 0 1 2 0 1 0 1 2 0 1 0 1 2 0 0 0 0 0 0

WEWP 0 1 1 0 2 0 1 1 0 2 0 1 1 0 2 0 0 0 0 0 0

WIFL 3 4 1 2 10 3 4 1 2 10 3 4 1 2 10 0 0 0 0 0 0

GRFL 0 0 0 1 1 0 0 0 1 1 0 0 0 1 1 0 0 0 0 0 0

DUFL 1 0 0 0 1 1 0 0 0 1 1 0 0 0 1 0 0 0 0 0 0

PSFL 22 5 2 3 32 21 5 2 3 31 21 5 2 3 31 0 0 0 0 0 0

LBVI 2 0 0 0 2 1 0 0 0 1 1 0 0 0 1 0 0 0 0 0 0

WAVI 3 1 2 3 9 2 1 2 3 8 2 1 2 3 8 0 0 0 0 0 0

RCKI 4 2 4 2 12 4 2 4 2 12 4 2 4 2 12 0 0 0 0 0 0

SWTH 1 1 8 0 10 1 1 8 0 10 1 1 8 0 10 0 0 0 0 0 0

HETH 2 0 0 0 2 2 0 0 0 2 2 0 0 0 2 0 0 0 0 0 0

WREN 1 0 0 0 1 1 0 0 0 1 1 0 0 0 1 0 0 0 0 0 0

NOMO 0 0 0 1 1 0 0 0 1 1 0 0 0 1 1 0 0 0 0 0 0

PHAI 3 0 2 0 5 3 0 2 0 5 3 0 2 0 5 0 0 0 0 0 0

OCWA 18 17 5 11 51 17 16 5 11 49 17 15 5 11 48 1 0 0 0 0 1

NAWA 1 3 1 0 5 1 3 1 0 5 1 3 1 0 5 0 0 0 0 0 0

YEWA 28 7 11 10 56 28 7 11 10 56 28 7 11 10 56 0 0 0 0 0 0

AUWA 0 1 8 1 10 0 1 8 1 10 0 1 8 1 10 0 0 0 0 0 0

BTYW 1 1 2 1 5 1 1 2 1 5 1 1 2 1 5 0 0 0 0 0 0

BLPW 0 1 0 0 1 0 1 0 0 1 0 1 0 0 1 0 0 0 0 0 0

MGWA 4 1 0 0 5 4 1 0 0 5 4 1 0 0 5 0 0 0 0 0 0

COYE 3 5 6 5 19 3 5 6 5 19 3 5 6 5 19 0 0 0 0 0 0

WIWA 51 22 15 7 95 49 22 14 7 92 47 22 14 7 90 0 0 0 0 0 0
70 

Total captures Total number of individuals 

captured New individuals banded Recapture year / Year originally banded

Year Year Year 2014 2015 2016

Species 2013 2014 2015 2016 Total 2013 2014 2015 2016 Total 2013 2014 2015 2016 Total 2013 2013 2014 2013 2015 Total

YBCH 9 1 4 1 15 5 1 2 1 9 5 1 2 1 9 0 0 0 0 0 0

WETA 4 6 3 3 16 4 6 3 3 16 4 6 3 3 16 0 0 0 0 0 0

GTTO 1 0 1 0 2 1 0 1 0 2 1 0 1 0 2 0 0 0 0 0 0

SPTO 52 69 12 5 138 44 64 11 5 124 40 54 9 4 107 8 1 0 1 0 171

CHSP 0 0 1 0 1 0 0 1 0 1 0 0 1 0 1 0 0 0 0 0 0

BRSP 0 1 1 0 2 0 1 1 0 2 0 1 1 0 2 0 0 0 0 0 0

LASP 9 4 1 4 18 9 4 1 4 18 8 4 1 3 16 0 0 0 0 0 12 

BTSP 0 1 1 0 2 0 1 1 0 2 0 1 1 0 2 0 0 0 0 0 0

FOSP 0 3 0 2 5 0 3 0 2 5 0 3 0 2 5 0 0 0 0 0 0

SOSP 0 0 2 0 2 0 0 2 0 2 0 0 2 0 2 0 0 0 0 0 0

LISP 3 3 1 5 12 3 3 1 5 12 3 3 1 5 12 0 0 0 0 0 0

WCSP 4 5 34 9 52 4 4 31 9 48 4 4 29 8 45 0 1 1 0 1 3

ORJU 0 1 3 0 4 0 1 3 0 4 0 1 3 0 4 0 0 0 0 0 0

LAZB 0 0 0 1 1 0 0 0 1 1 0 0 0 1 1 0 0 0 0 0 0

PUFI 0 1 0 0 1 0 1 0 0 1 0 1 0 0 1 0 0 0 0 0 0

HOFI 1 0 3 0 4 1 0 3 0 4 1 0 3 0 4 0 0 0 0 0 0

Total 234 172 140 81 627 216 165 132 81 594 206 153 124 75 558 9 2 1 1 1 221,2

1

Seven SPTO were originally banded during MAP, then recaptured the same year in the fall.

2

One LASP was originally banded during MAPS, then recaptured in the fall.
71 

Table 28. Capture frequency of migrant individuals during fall migration, Naval Base Coronado, Remote Training 

Site, Warner Springs, San Diego County, California, 2013.

[See tables 2 and 3 for bird species codes. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl] 

Number of individuals per capture incidence

(banded birds only) Total number of individuals

Species 1 capture 2 captures 3 captures 4 captures Banded birds Unbanded birds All birds

BCHU 0 0 0 0 0 2 2

RUHU 0 0 0 0 0 1 1

WIFL 3 0 0 0 3 0 3

DUFL 1 0 0 0 1 0 1

PSFL 20 1 0 0 21 0 21

LBVI 0 1 0 0 1 0 1

WAVI 1 1 0 0 2 0 2

RCKI 4 0 0 0 4 0 4

SWTH 1 0 0 0 1 0 1

HETH 2 0 0 0 2 0 2

WREN 1 0 0 0 1 0 1

PHAI 3 0 0 0 3 0 3

OCWA 16 1 0 0 17 0 17

NAWA 1 0 0 0 1 0 1

YEWA 28 0 0 0 28 0 28

BTYW 1 0 0 0 1 0 1

MGWA 4 0 0 0 4 0 4

COYE 3 0 0 0 3 0 3

WIWA 46 0 1 0 47 2 49

YBCH 3 1 0 1 5 0 5

WETA 4 0 0 0 4 0 4

GTTO 1 0 0 0 1 0 1

SPTO 37 6 1 0 44 0 44

LASP 8 0 0 0 8 1 9

LISP 3 0 0 0 3 0 3

WCSP 4 0 0 0 4 0 4

HOFI 1 0 0 0 1 0 1

Total 196 11 2 1 210 6 216
72 

Table 29. Capture frequency of migrant individuals during fall migration, Naval Base Coronado, Remote Training 

Site, Warner Springs, San Diego County, California, 2014.

[See tables 2 and 3 for bird species codes. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl] 

Number of Individuals per capture 

incidence (banded birds only) Total number of individuals

Species 1 capture 2 captures Banded birds Unbanded birds All birds

BCHU 0 0 0 1 1

RNSA 3 0 3 0 3

RBSA 1 0 1 0 1

WEWP 1 0 1 0 1

WIFL 4 0 4 0 4

PSFL 5 0 5 0 5

WAVI 1 0 1 0 1

RCKI 2 0 2 0 2

SWTH 1 0 1 0 1

OCWA 15 1 16 0 16

NAWA 3 0 3 0 3

YEWA 7 0 7 0 7

AUWA 1 0 1 0 1

BTYW 1 0 1 0 1

BLPW 1 0 1 0 1

MGWA 1 0 1 0 1

COYE 5 0 5 0 5

WIWA 22 0 22 0 22

YBCH 1 0 1 0 1

WETA 6 0 6 0 6

SPTO 59 5 64 0 64

BRSP 1 0 1 0 1

LASP 4 0 4 0 4

BTSP 1 0 1 0 1

FOSP 3 0 3 0 3

LISP 3 0 3 0 3

WCSP 3 1 4 0 4

ORJU 1 0 1 0 1

PUFI 1 0 1 0 1

Total 157 7 164 1 165
73 

Table 30. Capture frequency of migrant individuals during fall migration, Naval Base Coronado, Remote Training 

Site, Warner Springs, San Diego County, California, 2015.

[See tables 2 and 3 for bird species codes. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl] 

Number of individuals per capture 

incidence (banded birds only) Total number of individuals

Species 1 capture 2 captures 3 captures Banded birds Unbanded birds All birds

SSHA 0 0 0 0 1 1

BCHU 0 0 0 0 2 2

COHU 0 0 0 0 1 1

WEWP 1 0 0 1 0 1

WIFL 1 0 0 1 0 1

PSFL 2 0 0 2 0 2

WAVI 2 0 0 2 0 2

RCKI 4 0 0 4 0 4

SWTH 8 0 0 8 0 8

PHAI 2 0 0 2 0 2

OCWA 5 0 0 5 0 5

NAWA 1 0 0 1 0 1

YEWA 11 0 0 11 0 11

AUWA 8 0 0 8 0 8

BTYW 2 0 0 2 0 2

COYE 6 0 0 6 0 6

WIWA 13 1 0 14 0 14

YBCH 1 0 1 2 0 2

WETA 3 0 0 3 0 3

GTTO 1 0 0 1 0 1

SPTO 10 1 0 11 0 11

CHSP 1 0 0 1 0 1

BRSP 1 0 0 1 0 1

LASP 1 0 0 1 0 1

BTSP 1 0 0 1 0 1

SOSP 2 0 0 2 0 2

LISP 1 0 0 1 0 1

WCSP 28 3 0 31 0 31

ORJU 3 0 0 3 0 3

HOFI 3 0 0 3 0 3

Total 122 5 1 128 4 132
74 

Table 31. Capture frequency of individual migrants during fall migration, Naval Base Coronado, Remote Training 

Site, Warner Springs, San Diego County, California, 2016.

[See tables 2 and 3 for bird species codes. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl] 

Number of individuals per capture 

incidence (banded birds only) Total number of individuals

Species 1 capture Banded birds Unbanded birds All birds

BCHU 0 0 1 1

COHU 0 0 2 2

RBSA 1 1 0 1

WIFL 2 2 0 2

GRFL 1 1 0 1

PSFL 3 3 0 3

WAVI 3 3 0 3

RCKI 2 2 0 2

NOMO 1 1 0 1

OCWA 11 11 0 11

YEWA 10 10 0 10

AUWA 1 1 0 1

BTYW 1 1 0 1

COYE 5 5 0 5

WIWA 7 7 0 7

YBCH 1 1 0 1

WETA 3 3 0 3

SPTO 5 5 0 5

LASP 4 4 0 4

FOSP 2 2 0 2

LISP 5 5 0 5

WCSP 9 9 0 9

LAZB 1 1 0 1

Total 78 78 3 81
75 

Table 32. Capture rate by net and date during fall migration, Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 

2013.

[Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl]

Banding 

Day

Net Totals 

Date 1 2 3 4 5 6 7 8 9 10 11 12 13 14 by date

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 1:10 40:10

Captures 0 0 0 0 0 0 0 0 0 1 0 1 0 1 3

1 8/15/13 Captures per net-hour 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.33 0.00 0.33 0.00 0.86 0.07

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 1 0 0 1 0 0 0 1 0 0 0 1 1 1 6

2 8/20/13 Captures per net-hour 0.33 0.00 0.00 0.33 0.00 0.00 0.00 0.33 0.00 0.00 0.00 0.33 0.33 0.33 0.14

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 0 0 0 0 0 0 0 0 0 0 0 1 1 2 4

3 8/22/13 Captures per net-hour 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.33 0.33 0.67 0.10

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 1 1 0 0 1 0 2 1 1 0 0 0 2 1 10

4 8/27/13 Captures per net-hour 0.33 0.33 0.00 0.00 0.33 0.00 0.67 0.33 0.33 0.00 0.00 0.00 0.67 0.33 0.24

Net-hours 3:30 3:30 3:30 3:30 3:30 3:30 3:30 3:30 3:30 3:30 3:30 3:30 3:30 3:30 49:00

Captures 1 0 2 2 0 0 1 2 1 0 3 0 2 3 17

5 8/28/13 Captures per net-hour 0.29 0.00 0.57 0.57 0.00 0.00 0.29 0.57 0.29 0.00 0.86 0.00 0.57 0.86 0.35

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 0 0 0 0 0 1 2 2 0 0 0 1 2 0 8

6 8/29/13 Captures per net-hour 0.00 0.00 0.00 0.00 0.00 0.33 0.67 0.67 0.00 0.00 0.00 0.33 0.67 0.00 0.19

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 1 3 1 0 0 0 1 0 0 1 1 0 1 2 11

7 9/3/13 Captures per net-hour 0.33 1.00 0.33 0.00 0.00 0.00 0.33 0.00 0.00 0.33 0.33 0.00 0.33 0.67 0.26

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 0 0 0 2 0 2 0 0 0 0 1 1 1 0 7

8 9/5/13 Captures per net-hour 0.00 0.00 0.00 0.67 0.00 0.67 0.00 0.00 0.00 0.00 0.33 0.33 0.33 0.00 0.17

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 0 0 4 1 0 0 0 2 1 1 0 1 0 1 11

9 9/10/13 Captures per net-hour 0.00 0.00 1.33 0.33 0.00 0.00 0.00 0.67 0.33 0.33 0.00 0.33 0.00 0.33 0.26
76 

Banding 

Day

Net Totals 

Date 1 2 3 4 5 6 7 8 9 10 11 12 13 14 by date

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 1 0 0 1 0 1 5 1 0 1 1 0 1 2 14

10 9/12/13 Captures per net-hour 0.33 0.00 0.00 0.33 0.00 0.33 1.67 0.33 0.00 0.33 0.33 0.00 0.33 0.67 0.33

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 0 1 1 5 0 0 0 0 1 0 0 1 5 2 16

11 9/17/13 Captures per net-hour 0.00 0.33 0.33 1.67 0.00 0.00 0.00 0.00 0.33 0.00 0.00 0.33 1.67 0.67 0.38

Net-hours 6:20 6:20 6:20 6:20 5:50 5:50 4:20 4:50 4:20 4:50 4:50 4:20 4:20 4:20 73:10

Captures 2 3 4 7 3 9 3 5 1 0 1 1 4 0 43

12 9/18/13 Captures per net-hour 0.32 0.47 0.63 1.11 0.51 1.54 0.69 1.03 0.23 0.00 0.21 0.23 0.92 0.00 0.59

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 0 1 0 1 0 2 0 0 0 1 1 1 3 1 11

13 9/19/13 Captures per net-hour 0.00 0.33 0.00 0.33 0.00 0.67 0.00 0.00 0.00 0.33 0.33 0.33 1.00 0.33 0.26

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 1 1 2 3 0 0 0 0 0 2 2 0 4 3 18

14 9/24/13 Captures per net-hour 0.33 0.33 0.67 1.00 0.00 0.00 0.00 0.00 0.00 0.67 0.67 0.00 1.33 1.00 0.43

Net-hours 6:00 6:00 6:00 6:00 6:00 4:30 3:10 3:10 3:10 3:10 3:10 3:10 6:00 6:00 65:30

Captures 1 3 4 4 0 4 2 1 0 0 1 2 2 6 30

15 9/26/13 Captures per net-hour 0.17 0.50 0.67 0.67 0.00 0.89 0.63 0.32 0.00 0.00 0.32 0.63 0.33 1.00 0.46

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 2 1 0 0 0 0 1 0 0 1 1 0 0 1 7

16 9/27/13 Captures per net-hour 0.67 0.33 0.00 0.00 0.00 0.00 0.33 0.00 0.00 0.33 0.33 0.00 0.00 0.33 0.17

Net-hours 3:10 3:10 3:10 3:10 3:10 3:10 3:10 3:10 3:10 3:10 3:10 3:10 3:10 3:10 44:20

Captures 1 1 1 0 1 2 3 1 0 1 1 1 5 0 18

17 10/23/13 Captures per net-hour 0.32 0.32 0.32 0.00 0.32 0.63 0.95 0.32 0.00 0.32 0.32 0.32 1.58 0.00 0.41

Totals by net

Net-hours 58:00 58:00 58:00 58:00 57:30 56:00 53:10 53:40 53:10 53:40 53:40 53:10 56:00 54:10 776:10

Captures 12 15 19 27 5 21 20 16 5 9 13 12 34 26 234

Captures per net-hour 0.21 0.26 0.33 0.47 0.09 0.38 0.38 0.30 0.09 0.17 0.24 0.23 0.61 0.48 0.30
77 

Table 33. Capture rate by net and date during fall migration, Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, California, 

2014.

[Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl]

Banding Net

Totals 

by

day Date 1 2 3 4 5 6 7 8 9 10 11 12 13 14 date

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 30:00

Captures 0 0 1 1 0 0 1 1 1 0 0 0 0 0 5

1 8/15/14 Captures per net-hour 0.00 0.00 0.33 0.33 0.00 0.00 0.33 0.33 0.33 0.00 0.00 0.00 0.00 0.00 0.17

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 30:00

Captures 0 0 0 0 2 0 0 0 0 0 0 0 0 0 2

2 8/19/14 Captures per net-hour 0.00 0.00 0.00 0.00 0.67 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.07

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 30:00

Captures 1 0 0 1 1 0 0 0 0 0 1 0 0 0 4

3 8/21/14 Captures per net-hour 0.33 0.00 0.00 0.33 0.33 0.00 0.00 0.00 0.00 0.00 0.33 0.00 0.00 0.00 0.13

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 30:00

Captures 1 0 0 0 0 2 0 0 1 0 0 1 0 2 7

4 8/26/14 Captures per net-hour 0.33 0.00 0.00 0.00 0.00 0.67 0.00 0.00 0.33 0.00 0.00 0.33 0.00 0.67 0.23

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 30:00

Captures 0 0 1 0 0 1 1 0 0 0 0 0 0 0 3

5 8/28/14 Captures per net-hour 0.00 0.00 0.33 0.00 0.00 0.33 0.33 0.00 0.00 0.00 0.00 0.00 0.00 0.00 1.77

Net-hours 4:30 4:30 4:30 4:30 4:30 4:30 4:30 4:30 4:30 4:30 4:30 4:30 4:30 4:30 45:00

Captures 1 2 2 0 3 2 1 3 2 0 2 0 1 4 23

6 9/10/14 Captures per net-hour 0.22 0.44 0.44 0.00 0.67 0.44 0.22 0.67 0.44 0.00 0.44 0.00 0.22 0.89 0.51

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 30:00

Captures 1 2 0 1 0 0 0 0 1 1 1 1 3 0 11

7 9/16/14 Captures per net-hour 0.33 0.67 0.00 0.33 0.00 0.00 0.00 0.00 0.33 0.33 0.33 0.33 1.00 0.00 0.37

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 30:00

Captures 0 1 3 0 0 0 0 1 0 1 0 1 1 0 8

8 9/18/14 Captures per net-hour 0.00 0.33 1.00 0.00 0.00 0.00 0.00 0.33 0.00 0.33 0.00 0.33 0.33 0.00 0.27

Net-hours 3:50 3:50 3:50 3:50 3:50 3:50 3:50 3:50 3:50 3:50 3:50 3:50 3:20 3:50 38:20

Captures 0 1 6 1 1 4 0 1 2 1 1 0 4 4 26

9 9/23/14 Captures per net-hour 0.00 0.26 1.57 0.26 0.26 1.04 0.00 0.26 0.52 0.26 0.26 0.00 1.20 1.04 0.86

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 30:00

Captures 1 0 1 1 0 2 1 0 0 1 0 0 2 1 10

10 9/24/14 Captures per net-hour 0.33 0.00 0.33 0.33 0.00 0.67 0.33 0.00 0.00 0.33 0.00 0.00 0.67 0.33 0.33
78 

Banding Net

Totals 

by

day Date 1 2 3 4 5 6 7 8 9 10 11 12 13 14 date

Net-hours 3:50 3:50 3:50 3:50 3:50 3:50 3:20 3:50 3:50 3:50 3:50 3:20 3:50 3:50 37:50

Captures 0 0 2 2 2 4 1 1 1 1 0 1 0 6 21

11 9/30/14 Captures per net-hour 0.00 0.00 0.52 0.52 0.52 1.04 0.30 0.26 0.26 0.26 0.00 0.30 0.00 1.57 0.56

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 30:00

Captures 0 0 0 0 1 1 0 0 1 0 0 0 1 3 7

12 10/1/14 Captures per net-hour 0.00 0.00 0.00 0.00 0.33 0.33 0.00 0.00 0.33 0.00 0.00 0.00 0.33 1.00 0.23

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 30:00

Captures 0 1 0 0 0 2 2 2 1 0 0 2 1 2 13

13 10/7/14 Captures per net-hour 0.00 0.33 0.00 0.00 0.00 0.67 0.67 0.67 0.33 0.00 0.00 0.67 0.33 0.67 0.43

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 30:00

Captures 2 0 3 0 2 0 1 0 1 2 1 0 0 2 14

14 10/9/14 Captures per net-hour 0.67 0.00 1.00 0.00 0.67 0.00 0.33 0.00 0.33 0.67 0.33 0.00 0.00 0.67 0.47

Net-hours 6:00 6:00 6:00 6:00 6:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 6:00 6:00 45:00

Captures 0 0 0 0 1 2 0 0 0 1 1 1 1 2 9

15 10/10/14 Captures per net-hour 0.00 0.00 0.00 0.00 0.17 0.67 0.00 0.00 0.00 0.33 0.33 0.33 0.17 0.33 0.20

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 30:00

Captures 1 0 0 0 1 0 0 0 0 1 1 2 3 0 9

16 10/15/14 Captures per net-hour 0.33 0.00 0.00 0.00 0.33 0.00 0.00 0.00 0.00 0.33 0.33 0.67 1.00 0.00 0.30

Totals by net

Net-hours 54:10 54:10 54:10 54:10 54:10 51:10 50:40 51:10 51:10 51:10 51:10 50:40 53:40 54:10 735:50

Captures 8 7 19 7 14 20 8 9 11 9 8 9 17 26 172

Captures per net-hour 0.15 0.13 0.35 0.13 0.26 0.39 0.16 0.18 0.21 0.18 0.16 0.18 0.32 0.48 0.23
79 

Table 34. Capture rate of migrant species by net and date during fall migration, Naval Base Coronado, Remote Training Site, Warner Springs, San Diego 

County, California, 2015.

[Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl]

Banding 

day Date

Net Totals 

by 

1 2 3 4 5 6 7 8 9 10 11 12 13 14 date

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

1 8/19/15 Captures per net-hour 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0

2 8/21/15 Captures per net-hour 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1

3 8/25/15 Captures per net-hour 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.33 0.02

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 2 0 1 0 0 0 0 0 0 0 0 0 0 0 3

4 8/27/15 Captures per net-hour 0.67 0.00 0.33 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.07

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 0 3 0 0 2 0 0 0 0 0 0 0 1 0 6

5 9/1/15 Captures per net-hour 0.00 1.00 0.00 0.00 0.67 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.33 0.00 0.14

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 0 0 1 0 0 0 0 1 0 1 1 0 1 1 6

6 9/3/15 Captures per net-hour 0.00 0.00 0.33 0.00 0.00 0.00 0.00 0.33 0.00 0.33 0.33 0.00 0.33 0.33 0.14

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 0 0 0 0 0 0 0 0 0 0 0 0 2 2 4

7 9/8/15 Captures per net-hour 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.67 0.67 0.10

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 0 0 1 0 0 0 0 0 1 0 1 0 3 4 10

8 9/10/15 Captures per net-hour 0.00 0.00 0.33 0.00 0.00 0.00 0.00 0.00 0.33 0.00 0.33 0.00 1.00 1.33 0.24

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 0 0 0 1 0 0 2 2 1 0 0 0 2 1 9

9 9/16/15 Captures per net-hour 0.00 0.00 0.00 0.33 0.00 0.00 0.67 0.67 0.33 0.00 0.00 0.00 0.67 0.33 0.21

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 0 2 0 0 0 0 0 0 0 1 0 0 0 1 4

10 9/18/15 Captures per net-hour 0.00 0.67 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.33 0.00 0.00 0.00 0.33 0.10
80 

Banding 

day Date

Net Totals 

by 

1 2 3 4 5 6 7 8 9 10 11 12 13 14 date

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 0 0 1 0 0 1 2 1 0 0 0 0 1 3 9

11 9/23/15 Captures per net-hour 0.00 0.00 0.33 0.00 0.00 0.33 0.67 0.33 0.00 0.00 0.00 0.00 0.33 1.00 0.21

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 0 0 0 1 0 1 1 3 0 1 0 0 1 3 11

12 9/25/15 Captures per net-hour 0.00 0.00 0.00 0.33 0.00 0.33 0.33 1.00 0.00 0.33 0.00 0.00 0.33 1.00 0.26

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 0 2 4 0 0 2 2 0 1 1 0 0 0 3 15

13 9/29/15 Captures per net-hour 0.00 0.67 1.33 0.00 0.00 0.67 0.67 0.00 0.33 0.33 0.00 0.00 0.00 1.00 0.36

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 1 2 0 0 0 1 0 0 0 2 0 0 2 1 9

14 9/30/15 Captures per net-hour 0.33 0.67 0.00 0.00 0.00 0.33 0.00 0.00 0.00 0.67 0.00 0.00 0.67 0.33 0.21

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 0 0 0 0 0 0 0 0 0 0 0 0 1 2 3

15 10/6/15 Captures per net-hour 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.33 0.67 0.07

Net-hours 3:50 3:50 3:50 3:50 3:50 3:50 3:50 3:50 3:50 3:50 3:50 3:50 3:50 3:50 53:40

Captures 1 2 0 1 0 1 2 5 1 3 2 0 1 3 22

16 10/8/15 Captures per net-hour 0.26 0.52 0.00 0.26 0.00 0.26 0.52 1.30 0.26 0.78 0.52 0.00 0.26 0.78 0.41

Net-hours 2:00 3:00 3:00 2:00 3:00 3:00 3:00 3:00 3:00 2:30 2:30 2:30 3:00 2:30 38:00

Captures 0 0 0 0 0 1 0 1 0 0 0 0 6 1 9

17 10/9/15 Captures per net-hour 0.00 0.00 0.00 0.00 0.00 0.33 0.00 0.33 0.00 0.00 0.00 0.00 2.00 0.40 0.24

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 1 1 0 4 0 1 5 1 0 0 0 0 0 1 14

18 10/14/15 Captures per net-hour 0.33 0.33 0.00 1.33 0.00 0.33 1.67 0.33 0.00 0.00 0.00 0.00 0.00 0.33 0.33

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 0 0 1 1 0 1 0 1 0 0 0 0 1 0 5

19 10/15/15 Captures per net-hour 0.00 0.00 0.33 0.33 0.00 0.33 0.00 0.33 0.00 0.00 0.00 0.00 0.33 0.00 0.12

Totals by net

Net-hours 56:50 57:50 57:50 56:50 57:50 57:50 57:50 57:50 57:50 57:20 57:20 57:20 57:50 57:20 805:40

Captures 5 12 9 8 2 9 14 15 4 9 4 0 22 27 140

Captures per net-hour 0.09 0.21 0.16 0.14 0.03 0.16 0.24 0.26 0.07 0.16 0.07 0.00 0.38 0.47 0.17
81 

Table 35. Capture rate of migrant species by net and date during fall migration, Naval Base Coronado, Remote Training Site, Warner Springs, San Diego 

County, California, 2016.

[Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl]

Banding 

day

Net

Totals 

by 

Date 1 2 3 4 5 6 7 8 9 10 11 12 13 14 date

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 0:22 0:22 0:22 0:22 31:30

Captures 0 0 0 0 0 0 0 0 0 1 0 0 0 0 1

1 8/18/16 Captures per net-hour 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.33 0.00 0.00 0.00 0.00 0.03

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 0 0 0 0 0 0 0 0 0 0 0 1 1 0 2

2 8/19/16 Captures per net-hour 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.33 0.33 0.00 0.05

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1

3 8/23/16 Captures per net-hour 0.33 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.02

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1

4 9/1/16 Captures per net-hour 0.33 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.33 0.00 0.00 0.00 0.00 0.00 0.02

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 0 0 0 0 0 1 0 0 0 0 0 0 0 1 2

5 9/2/16 Captures per net-hour 0.00 0.00 0.00 0.00 0.00 0.33 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.33 0.05

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 2 0 0 0 0 1 0 0 0 0 0 0 1 2 6

6 9/6/16 Captures per net-hour 0.67 0.00 0.00 0.00 0.00 0.33 0.00 0.00 0.00 0.00 0.00 0.00 0.33 0.67 0.14

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 0 1 0 0 0 0 0 0 0 0 0 1 0 3 5

7 9/8/16 Captures per net-hour 0.00 0.33 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.33 0.00 1.00 0.12

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 2 1 0 1 0 3 1 0 0 3 0 0 1 3 15

8 9/15/16 Captures per net-hour 0.67 0.33 0.00 0.33 0.00 1.00 0.33 0.00 0.00 1.00 0.00 0.00 0.33 1.00 0.36

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 0 2 0 0 0 1 0 0 0 3 0 0 0 3 9

9 9/16/16 Captures per net-hour 0.00 0.67 0.00 0.00 0.00 0.33 0.00 0.00 0.00 1.00 0.00 0.00 0.00 1.00 0.21

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 0 0 2 0 0 0 0 2 0 0 1 0 1 1 7

10 9/22/16 Captures per net-hour 0.00 0.00 0.67 0.00 0.00 0.00 0.00 0.67 0.00 0.00 0.33 0.00 0.33 0.33 0.17
82 

Banding 

day

Net

Totals 

by 

Date 1 2 3 4 5 6 7 8 9 10 11 12 13 14 date

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 0 0 0 1 2 0 0 0 0 1 0 0 3 0 7

11 9/29/16 Captures per net-hour 0.00 0.00 0.00 0.33 0.67 0.00 0.00 0.00 0.00 0.33 0.00 0.00 1.00 0.00 0.17

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 0 0 0 0 1 0 1 2 0 0 0 0 2 3 9

12 9/30/16 Captures per net-hour 0.00 0.00 0.00 0.00 0.33 0.00 0.33 0.67 0.00 0.00 0.00 0.00 0.67 1.00 0.21

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 0 0 0 0 0 0 0 0 0 0 0 0 0 2 2

13 10/4/16 Captures per net-hour 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.67 0.05

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 0 0 0 0 0 0 1 0 0 0 0 0 0 1 2

14 10/6/16 Captures per net-hour 0.00 0.00 0.00 0.00 0.00 0.00 0.33 0.00 0.00 0.00 0.00 0.00 0.00 0.33 0.05

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 2 0 0 0 0 0 2 1 0 0 0 0 1 0 6

15 10/11/16 Captures per net-hour 0.67 0.00 0.00 0.00 0.00 0.00 0.67 0.33 0.00 0.00 0.00 0.00 0.33 0.00 0.14

Net-hours 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 3:00 42:00

Captures 0 1 0 1 0 1 0 0 0 0 0 0 0 3 6

16 10/13/16 Captures per net-hour 0.00 0.33 0.00 0.33 0.00 0.33 0.00 0.00 0.00 0.00 0.00 0.00 0.00 1.00 0.14

Totals by net

Net-hours 48:00 48:00 48:00 48:00 48:00 48:00 48:00 48:00 48:00 48:00 45:22 45:22 45:22 45:22 661:30

Captures 8 5 2 3 3 7 5 5 0 8 1 2 10 22 81

Captures per net-hour 0.17 0.10 0.04 0.06 0.06 0.15 0.10 0.10 0.00 0.17 0.02 0.04 0.22 0.48 0.12
83 

Table 36. Number of migrant species captures by date during fall migration, Naval Base Coronado, Remote Training Site, Warner Springs, San Diego 

County, California, 2013.

[See tables 2 and 3 for bird species codes. Includes multiple captures for some individuals. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl]

Banding day

Captures per 100 nethours (776:10 total nethours)

1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17

Date

Species

8-15-2013

8-20-2013

8-22-2013

8-27-2013

8-28-2013

8-29-2013

9-03-2013

9-05-2013

9-10-2013

9-12-2013

9-17-2013

9-18-2013

9-19-2013

9-24-2013

9-26-2013

9-27-2013

10-23-2013

Total

BCHU 0 0 0 0 0 0 2 0 0 0 0 0 0 0 0 0 0 2 0.26

RUHU 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 1 0.13

WIFL 0 0 0 0 1 1 0 0 0 0 0 0 0 0 0 1 0 3 0.39

DUFL 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0.13

PSFL 0 1 0 1 1 1 1 1 0 2 1 0 1 0 10 2 0 22 2.84

LBVI 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 0.26

WAVI 0 0 0 0 0 0 0 0 0 0 0 1 0 1 1 0 0 3 0.39

RCKI 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 4 4 0.52

SWTH 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 1 0.13

HETH 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 1 2 0.26

WREN 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 1 0.13

PHAI 0 0 0 0 0 0 0 0 0 0 3 0 0 0 0 0 0 3 0.39

OCWA 0 0 0 2 2 0 1 0 1 0 0 7 1 1 2 1 0 18 2.32

NAWA 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 1 0.13

YEWA 0 0 1 0 2 0 0 0 0 1 3 16 1 0 4 0 0 28 3.61

BTYW 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 1 0.13

MGWA 0 0 0 0 0 1 0 0 2 0 0 1 0 0 0 0 0 4 0.52

COYE 0 0 0 0 0 0 0 0 0 0 0 1 1 1 0 0 0 3 0.39

WIWA 0 2 0 2 3 1 2 3 4 3 5 12 1 3 8 0 2 51 6.57

YBCH 0 0 0 2 1 1 0 1 0 1 0 0 1 1 1 0 0 9 1.16

WETA 0 0 0 1 2 0 0 0 1 0 0 0 0 0 0 0 0 4 0.52

GTTO 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 1 0.13

SPTO 2 2 2 1 4 2 4 2 2 5 3 3 3 6 3 3 5 52 6.70

LASP 0 1 0 0 0 0 1 0 0 0 0 2 0 1 1 0 3 9 1.16
84 

Banding day

Captures per 100 nethours (776:10 total nethours)

1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17

Date

Species

8-15-2013

8-20-2013

8-22-2013

8-27-2013

8-28-2013

8-29-2013

9-03-2013

9-05-2013

9-10-2013

9-12-2013

9-17-2013

9-18-2013

9-19-2013

9-24-2013

9-26-2013

9-27-2013

10-23-2013

Total

LISP 0 0 0 0 0 0 0 0 0 0 0 0 1 2 0 0 0 3 0.39

WCSP 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 3 4 0.52

HOFI 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 1 0.13

Captures per day 3 6 4 10 17 8 11 7 11 14 16 43 11 18 30 7 18 234 30.15

Total species 2 4 3 7 9 7 6 4 6 7 6 8 9 10 8 4 6 27 3.48
85 

Table 37. Number of migrant captures by date during fall migration, Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, 

California, 2014.

[See tables 2 and 3 for bird species codes. Includes multiple captures for some individuals. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl] 

MAPS period

Captures per 100 

net-hours (total 

735:50 net-hours)

1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16

Date

Species

8-15-2014

8-19-2014

8-21-2014

8-26-2014

8-28-2014

9-10-2014

9-16-2014

9-18-2014

9-23-2014

9-24-2014

9-30-2014

10-01-2014

10-07-201410-09-201410-10-201410-15-2014

Total

BCHU 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0.14

RNSA 0 0 0 0 0 0 0 0 0 1 0 0 0 2 0 0 3 0.41

RBSA 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 1 0.14

WEWP 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 1 0.14

WIFL 0 0 1 0 1 2 0 0 0 0 0 0 0 0 0 0 4 0.54

PSFL 0 0 0 0 0 0 1 2 1 0 0 0 1 0 0 0 5 0.68

WAVI 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 1 0.14

RCKI 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 2 0.27

SWTH 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 1 0.14

OCWA 2 0 0 2 0 3 0 1 2 1 5 1 0 0 0 0 17 2.31

NAWA 0 0 0 0 0 1 0 0 0 2 0 0 0 0 0 0 3 0.41

YEWA 0 0 0 0 0 3 1 0 0 1 2 0 0 0 0 0 7 0.95

AUWA 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 1 0.14

BTYW 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 1 0.14

BLPW 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 1 0.14

MGWA 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 1 0.14

COYE 0 0 0 0 0 0 0 0 0 1 1 2 1 0 0 0 5 0.68

WIWA 2 2 1 1 0 3 2 1 7 0 2 0 1 0 0 0 22 2.99

YBCH 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 1 0.14

WETA 0 0 0 1 0 2 0 0 2 0 1 0 0 0 0 0 6 0.82

SPTO 1 0 1 2 2 5 4 4 10 3 4 3 9 10 5 6 69 9.38

BRSP 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 1 0.14

LASP 0 0 0 0 0 1 2 0 1 0 0 0 0 0 0 0 4 0.54

BTSP 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 1 0.14

FOSP 0 0 0 0 0 0 0 0 1 0 0 0 0 2 0 0 3 0.41
86 

MAPS period

Captures per 100 

net-hours (total 

735:50 net-hours)

1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16

Date

Species

8-15-2014

8-19-2014

8-21-2014

8-26-2014

8-28-2014

9-10-2014

9-16-2014

9-18-2014

9-23-2014

9-24-2014

9-30-2014

10-01-2014

10-07-201410-09-201410-10-201410-15-2014

Total

LISP 0 0 0 0 0 0 0 0 1 1 1 0 0 0 0 0 3 0.41

WCSP 0 0 0 0 0 0 0 0 0 0 2 0 0 0 1 2 5 0.68

ORJU 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 1 0.14

PUFI 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 1 0.14

Captures per day 5 2 4 7 3 23 11 8 26 10 21 7 13 14 9 9 172 23.37

Total species 3 1 4 5 2 11 6 4 9 7 11 4 5 3 5 3 29 3.94
87 

Table 38. Number of migrant species captures by date during fall migration, Naval Base Coronado, Remote Training Site, Warner Springs, San Diego 

County, California, 2015.

[See tables 2 and 3 for bird species codes. Includes multiple captures of some individuals. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl] 

Banding day

Captures per 100 

net-hours (total 

805:40 net-hours)

1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19

Date

Species

8-19-2015

8-21-2015

8-25-2015

8-27-2015

9-01-2015

9-03-2015

9-08-2015

9-10-2015

9-16-2015

9-18-2015

9-23-2015

9-25-2015

9-29-2015

9-30-2015

10-06-201510-08-201510-09-201510-14-201510-15-2015

Total

SSHA 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 0.12

BCHU 0 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 0.25

COHU 0 0 0 0 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 2 0.25

WEWP 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 1 0.12

WIFL 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 1 0.12

PSFL 0 0 0 0 0 2 0 0 0 0 0 0 0 0 0 0 0 0 0 2 0.25

WAVI 0 0 0 0 0 0 0 2 0 0 0 0 0 0 0 0 0 0 0 2 0.25

RCKI 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 0 2 0 4 0.50

SWTH 0 0 0 0 0 0 1 0 5 2 0 0 0 0 0 0 0 0 0 8 0.99

PHAI 0 0 0 0 0 0 0 0 0 0 0 1 1 0 0 0 0 0 0 2 0.25

OCWA 0 0 0 0 0 0 0 1 0 0 0 1 2 0 0 1 0 0 0 5 0.62

NAWA 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0.12

YEWA 0 0 0 1 0 0 0 1 0 0 1 2 1 3 1 0 1 0 0 11 1.36

AUWA 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 5 2 8 0.99

BTYW 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 1 0 0 0 2 0.25

COYE 0 0 0 0 1 0 0 0 0 0 2 0 2 1 0 0 0 0 0 6 0.74

WIWA 0 0 0 1 2 1 1 1 1 1 0 1 4 2 0 0 0 0 0 15 1.86

YBCH 0 0 0 0 0 1 1 1 0 0 0 1 0 0 0 0 0 0 0 4 0.50

WETA 0 0 0 0 0 0 0 0 0 0 2 0 1 0 0 0 0 0 0 3 0.37

GTTO 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 1 0.12

SPTO 0 0 1 0 0 1 0 0 0 1 2 2 1 1 0 2 0 1 0 12 1.49

CHSP 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 1 0.12

BRSP 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0.12

LASP 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 1 0.12

BTSP 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 1 0.12
88 

Banding day

Captures per 100 

net-hours (total 

805:40 net-hours)

1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19

Date

Species

8-19-2015

8-21-2015

8-25-2015

8-27-2015

9-01-2015

9-03-2015

9-08-2015

9-10-2015

9-16-2015

9-18-2015

9-23-2015

9-25-2015

9-29-2015

9-30-2015

10-06-201510-08-201510-09-201510-14-201510-15-2015

Total

SOSP 0 0 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 2 0.25

LISP 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 1 0.12

WCSP 0 0 0 0 0 0 0 0 0 0 2 2 3 2 2 8 7 6 2 34 4.22

ORJU 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 3 0 0 0 3 0.37

HOFI 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 3 0 0 0 3 0.37

Captures per day 0 0 1 3 6 6 4 10 9 4 9 11 15 9 3 22 9 14 5 140 17.37

Total species 0 0 1 3 5 5 4 9 5 3 5 8 8 5 2 9 3 4 3 30 3.72
89 

Table 39. Number of migrant captures by date during fall migration, Naval Base Coronado, Remote Training Site, Warner Springs, San Diego County, 

California, 2016.

[See tables 2 and 3 for bird species codes. Includes multiple captures of some individuals. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl] 

Banding Day

Captures per 

100 net-hours 

(661:00 nethours)

1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16

Date

Species

8-18-2016

8-19-2016

8-23-2016

9-01-2016

9-02-2016

9-06-2016

9-08-2016

9-15-2016

9-16-2016

9-22-2016

9-29-2016

9-30-2016

10-04-201610-06-201610-11-201610-13-2016

Total

BCHU 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 1 0.15

COHU 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 0.30

RBSA 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 0.15

WIFL 0 0 0 0 1 1 0 0 0 0 0 0 0 0 0 0 2 0.30

GRFL 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0.15

PSFL 0 0 0 0 0 0 1 0 1 1 0 0 0 0 0 0 3 0.45

WAVI 0 0 0 0 0 0 0 1 2 0 0 0 0 0 0 0 3 0.45

RCKI 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 1 2 0.30

NOMO 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 1 0.15

OCWA 0 0 0 0 0 0 1 3 3 0 2 0 0 0 2 0 11 1.66

YEWA 0 0 0 1 0 2 1 5 0 1 0 0 0 0 0 0 10 1.51

AUWA 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 1 0.15

BTYW 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 1 0.15

COYE 0 0 0 0 0 1 0 0 2 0 1 0 0 0 1 0 5 0.76

WIWA 0 0 0 0 0 1 2 1 0 2 0 1 0 0 0 0 7 1.06

YBCH 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 1 0.15

WETA 0 0 0 0 0 0 0 0 1 1 0 0 1 0 0 0 3 0.45

SPTO 0 1 0 0 0 0 0 3 0 0 0 1 0 0 0 0 5 0.76

LASP 0 0 0 0 0 0 0 1 0 0 1 0 0 0 0 2 4 0.61

FOSP 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 0 2 0.30

LISP 0 0 0 0 0 0 0 0 0 1 2 1 0 0 0 1 5 0.76

WCSP 0 0 0 0 0 0 0 1 0 0 1 4 1 0 1 1 9 1.36

LAZB 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 1 0.15

Captures per day 1 2 1 1 2 6 5 15 9 7 7 9 2 2 6 6 81 12.25

Total species 1 2 1 1 2 5 4 7 5 6 5 6 2 2 4 5 23 3.48
90 

Table 40. Sex and age of individuals (banded and unbanded) captured during fall migration, Naval Base 

Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2013.

[See tables 2 and 3 for bird species codes. Species in italics are non-breeding neotropical migrants. Age: HY=hatchingyear, AHY=after-hatching-year, SY=second-year, ASY=after-second-year, I=indeterminable age. Data collected by USGS 

and available from http://www.pwrc.usgs.gov/bbl] 

Female Male Unknown sex

Age Female Age Male Age Unknown Species

Species HY AHY SY I total HY AHY ASY I total HY AHY I total total

BCHU 1 0 0 1 2 0 0 0 0 0 0 0 0 0 2

RUHU 0 1 0 0 1 0 0 0 0 0 0 0 0 0 1

WIFL 0 0 0 0 0 0 0 0 0 0 2 1 0 3 3

DUFL 0 0 0 0 0 0 0 0 0 0 0 1 0 1 1

PSFL 0 0 0 0 0 0 0 0 0 0 20 1 0 21 21

LBVI 0 0 0 0 0 0 1 0 0 1 0 0 0 0 1

WAVI 0 0 0 0 0 0 0 0 0 0 2 0 0 2 2

RCKI 0 3 0 0 3 0 1 0 0 1 0 0 0 0 4

SWTH 0 0 0 0 0 0 0 0 0 0 1 0 0 1 1

HETH 0 0 0 0 0 0 0 0 0 0 2 0 0 2 2

WREN 0 0 0 0 0 0 0 0 0 0 0 0 1 1 1

PHAI 0 1 0 0 1 1 0 0 0 1 1 0 0 1 3

OCWA 1 1 0 0 2 4 6 0 1 11 0 0 4 4 17

NAWA 0 0 0 0 0 0 0 0 0 0 0 1 0 1 1

YEWA 3 3 0 0 6 3 4 0 0 7 12 1 2 15 28

BTYW 0 0 0 0 0 1 0 0 0 1 0 0 0 0 1

MGWA 1 3 0 0 4 0 0 0 0 0 0 0 0 0 4

COYE 0 1 0 0 1 0 2 0 0 2 0 0 0 0 3

WIWA 7 8 0 3 18 10 16 0 0 26 0 3 2 5 49

YBCH 1 1 0 0 2 0 0 0 0 0 3 0 0 3 5

WETA 2 0 0 0 2 2 0 0 0 2 0 0 0 0 4

GTTO 0 0 0 0 0 0 0 0 0 0 1 0 0 1 1

SPTO 4 11 1 2 18 2 17 2 5 26 0 0 0 0 44

LASP 0 0 0 0 0 0 0 0 0 0 1 7 1 9 9

LISP 0 0 0 0 0 0 0 0 0 0 2 1 0 3 3

WCSP 0 0 0 0 0 0 0 0 0 0 1 3 0 4 4

HOFI 0 0 0 0 0 1 0 0 0 1 0 0 0 0 1

Total 20 33 1 6 60 24 47 2 6 79 48 19 10 77 216
91 

Table 41. Sex and age of migrant individuals (banded and unbanded) captured during fall migration, Naval Base 

Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2014.

[See tables 2 and 3 for bird species codes. Species in italics are non-breeding neotropical migrants. Age: HY=hatchingyear, AHY=after-hatching-year, ASY=after-second-year, I=indeterminable age. Data collected by USGS and available 

from http://www.pwrc.usgs.gov/bbl] 

Female Male Unknown sex

Age Female Age Male Age Unknown Species

Species HY AHY ASY I total HY AHY ASY I total HY AHY I total total

BCHU 0 0 0 0 0 0 1 0 0 1 0 0 0 0 1

RNSA 0 1 0 0 1 0 2 0 0 2 0 0 0 0 3

RBSA 0 0 0 0 0 0 0 0 0 0 0 1 0 1 1

WEWP 0 0 0 0 0 0 0 0 0 0 1 0 0 1 1

WIFL 0 0 0 0 0 0 0 0 0 0 3 1 0 4 4

PSFL 0 0 0 0 0 0 0 0 0 0 5 0 0 5 5

WAVI 0 0 0 0 0 0 0 0 0 0 1 0 0 1 1

RCKI 0 0 0 0 0 0 1 0 0 1 0 0 1 1 2

SWTH 0 0 0 0 0 0 0 0 0 0 1 0 0 1 1

OCWA 2 0 1 3 6 1 0 0 3 4 1 0 5 6 16

NAWA 0 1 0 0 1 1 0 0 1 2 0 0 0 0 3

YEWA 0 0 0 0 0 2 3 0 0 5 2 0 0 2 7

AUWA 0 0 0 0 0 1 0 0 0 1 0 0 0 0 1

BTYW 1 0 0 0 1 0 0 0 0 0 0 0 0 0 1

BLPW 0 1 0 0 1 0 0 0 0 0 0 0 0 0 1

MGWA 0 0 0 0 0 0 1 0 0 1 0 0 0 0 1

COYE 0 0 0 1 1 0 1 0 3 4 0 0 0 0 5

WIWA 1 1 0 0 2 9 1 0 4 14 4 0 2 6 22

YBCH 0 0 0 0 0 0 1 0 0 1 0 0 0 0 1

WETA 0 2 0 0 2 1 1 0 0 2 2 0 0 2 6

SPTO 11 14 3 2 30 3 26 4 1 34 0 0 0 0 64

BRSP 0 0 0 0 0 0 0 0 0 0 0 1 0 1 1

LASP 0 0 0 0 0 0 0 0 0 0 1 3 0 4 4

BTSP 0 0 0 0 0 0 0 0 0 0 1 0 0 1 1

FOSP 0 0 0 0 0 0 0 0 0 0 2 1 0 3 3

LISP 0 0 0 0 0 0 0 0 0 0 3 0 0 3 3

WCSP 0 0 0 0 0 0 0 0 0 0 1 3 0 4 4

ORJU 0 1 0 0 1 0 0 0 0 0 0 0 0 0 1

PUFI 0 0 0 0 0 0 0 0 0 0 1 0 0 1 1

Total 15 21 4 6 46 18 38 4 12 72 29 10 8 47 165
92 

Table 42. Sex and age of migrant individuals (banded and unbanded) captured during fall migration, Naval Base 

Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2015.

[See tables 2 and 3 for bird species codes. Species in italics are non-breeding neotropical migrants. Age: HY=hatchingyear, AHY=after-hatching-year, ASY=after-second-year, I=indeterminable age. Data collected by USGS and available 

from http://www.pwrc.usgs.gov/bbl] 

Female Male Unknown sex

Age Female Age Male Age Unknown Species

Species HY AHY ASY I total HY AHY I total HY AHY ASY I total total

SSHA 0 0 0 0 0 1 0 0 1 0 0 0 0 0 1

BCHU 0 1 0 0 1 1 0 0 1 0 0 0 0 0 2

COHU 0 1 0 0 1 0 0 0 0 0 0 0 0 0 1

WEWP 0 0 0 0 0 0 0 0 0 1 0 0 0 1 1

WIFL 0 0 0 0 0 0 0 0 0 1 0 0 0 1 1

PSFL 0 0 0 0 0 0 0 0 0 0 2 0 0 2 2

WAVI 0 0 0 0 0 0 0 0 0 2 0 0 0 2 2

RCKI 0 0 0 0 0 0 0 0 0 1 0 0 3 4 4

SWTH 0 0 0 0 0 0 0 0 0 5 3 0 0 8 8

PHAI 0 0 0 2 2 0 0 0 0 0 0 0 0 0 2

OCWA 0 0 0 1 1 3 0 1 4 0 0 0 0 0 5

NAWA 0 0 0 0 0 1 0 0 1 0 0 0 0 0 1

YEWA 3 1 0 0 4 6 1 0 7 0 0 0 0 0 11

AUWA 0 3 0 0 3 3 0 0 3 2 0 0 0 2 8

BTYW 0 0 0 0 0 1 1 0 2 0 0 0 0 0 2

COYE 0 0 0 0 0 3 1 1 5 1 0 0 0 1 6

WIWA 1 8 0 0 9 3 2 0 5 0 0 0 0 0 14

YBCH 0 0 0 0 0 0 1 0 1 1 0 0 0 1 2

WETA 0 0 0 0 0 2 1 0 3 0 0 0 0 0 3

GTTO 0 0 0 0 0 0 0 0 0 1 0 0 0 1 1

SPTO 3 3 1 0 7 2 2 0 4 0 0 0 0 0 11

CHSP 0 0 0 0 0 0 0 0 0 1 0 0 0 1 1

BRSP 0 0 0 0 0 0 0 0 0 0 1 0 0 1 1

LASP 0 0 0 0 0 0 0 0 0 0 1 0 0 1 1

BTSP 0 0 0 0 0 0 0 0 0 1 0 0 0 1 1

SOSP 0 0 0 0 0 0 0 0 0 1 1 0 0 2 2

LISP 0 0 0 0 0 0 0 0 0 1 0 0 0 1 1

WCSP 0 0 0 0 0 0 0 0 0 12 17 2 0 31 31

ORJU 0 0 0 0 0 0 0 0 0 1 0 0 2 3 3

HOFI 0 0 0 0 0 0 0 2 2 0 0 0 1 1 3

Total 7 17 1 3 28 26 9 4 39 32 25 2 6 65 132
93 

Table 43. Sex and age of migrant individuals (banded and unbanded) captured during fall migration, Naval Base 

Coronado, Remote Training Site, Warner Springs, San Diego County, California, 2016.

[See tables 2 and 3 for bird species codes. Age: HY=hatching-year, AHY=after-hatching-year, SY=second-year, 

ASY=after-second-year, I=indeterminable age. Data collected by USGS and available from http://www.pwrc.usgs.gov/bbl] 

Female Male Unknown sex

Age Female Age Male Age Unknown Species

Species HY AHY I total HY AHY ASY I total HY AHY SY I total total

BCHU 0 1 0 1 0 0 0 0 0 0 0 0 0 0 1

COHU 0 1 1 2 0 0 0 0 0 0 0 0 0 0 2

RBSA 0 0 0 0 0 0 0 0 0 0 1 0 0 1 1

WIFL 0 0 0 0 0 0 0 0 0 1 1 0 0 2 2

GRFL 0 0 0 0 0 0 0 0 0 1 0 0 0 1 1

PSFL 0 0 0 0 0 0 0 0 0 2 1 0 0 3 3

WAVI 0 0 0 0 0 0 0 0 0 2 1 0 0 3 3

RCKI 0 0 0 0 0 1 0 0 1 0 0 0 1 1 2

NOMO 0 0 0 0 0 0 0 0 0 1 0 0 0 1 1

OCWA 1 0 1 2 1 2 0 0 3 0 0 0 6 6 11

YEWA 2 2 0 4 3 1 0 0 4 2 0 0 0 2 10

AUWA 0 0 0 0 1 0 0 0 1 0 0 0 0 0 1

BTYW 0 1 0 1 0 0 0 0 0 0 0 0 0 0 1

COYE 0 0 0 0 1 0 0 2 3 1 0 0 1 2 5

WIWA 1 2 0 3 1 2 0 0 3 0 0 0 1 1 7

YBCH 0 0 0 0 0 1 0 0 1 0 0 0 0 0 1

WETA 0 2 0 2 0 0 0 0 0 1 0 0 0 1 3

SPTO 1 0 0 1 1 1 1 0 3 1 0 0 0 1 5

LASP 0 0 0 0 0 0 0 0 0 2 2 0 0 4 4

FOSP 0 0 0 0 0 0 0 0 0 2 0 0 0 2 2

LISP 0 0 0 0 0 0 0 0 0 3 2 0 0 5 5

WCSP 0 0 0 0 0 0 0 0 0 2 6 1 0 9 9

LAZB 0 0 0 0 0 0 0 0 0 1 0 0 0 1 1

Total 5 9 2 16 8 8 1 2 19 22 14 1 9 46 81
94 

Table 44. First and median capture dates for each migrant species during fall migration, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California, 2013–16.

[See tables 2 and 3 for bird species codes. Median was calculated for species with at least two individuals captured. Data 

collected by USGS and available from http://www.pwrc.usgs.gov/bbl] 

First arrival date Median arrival date Number of individuals captured

Species 2013 2014 2015 2016 Average 2013 2014 2015 2016 2013 2014 2015 2016

SSHA 10/15 10/15 1

BCHU 9/3 8/21 8/27 9/30 9/5 9/3 8/29 2 1 2 1

COHU 9/8 8/18 8/29 9/9 8/18 2 2

RUHU 9/12 9/12 1

RNSA 9/24 9/24 10/9 3

RBSA 10/7 10/13 10/10 1 1

WEWP 9/16 9/10 9/13 1 1

WIFL 8/28 8/21 9/10 9/2 8/31 8/29 9/3 9/4 3 4 1 2

GRFL 8/23 8/23 1

DUFL 8/27 8/27 1

PSFL 8/20 9/16 9/3 9/8 9/4 9/26 9/18 9/3 9/16 21 5 2 3

LBVI 8/15 8/15 1

WAVI 9/18 9/10 9/10 9/15 9/14 9/21 9/10 9/16 2 1 2 3

RCKI 10/23 10/10 10/8 10/6 10/12 10/23 10/12 10/11 10/9 4 2 4 2

SWTH 9/10 8/26 9/8 9/4 9/16 1 1 8 0

HETH 9/24 9/24 10/8 2

WREN 9/19 9/19 1

NOMO 9/2 9/2 1

PHAI 9/17 9/25 9/21 9/17 9/27 3 2

OCWA 8/27 8/15 9/10 9/8 8/31 9/18 9/20 9/29 9/16 17 16 5 11

NAWA 9/17 9/10 9/1 9/9 9/24 1 3 1

YEWA 8/22 9/10 8/27 9/1 8/31 9/18 9/16 9/29 9/15 28 7 11 10

AUWA 10/10 10/9 9/30 10/7 10/14 1 8 1

BTYW 8/28 9/30 9/16 10/6 9/20 9/27 1 1 2 1

BLPW 9/23 9/23 1

MGWA 8/29 9/30 9/14 9/10 4 1

COYE 9/18 9/24 9/1 9/6 9/13 9/19 10/1 9/26 9/16 3 5 6 5

WIWA 8/20 8/15 8/27 9/6 8/25 9/17 9/17 9/17 9/15 49 22 14 7

YBCH 8/27 9/10 9/3 9/22 9/8 9/12 9/14 5 1 2 1

WETA 8/27 8/26 9/23 9/16 9/8 8/28 9/16 9/23 9/22 4 6 3 3

GTTO 9/12 9/16 9/14 1 1

SPTO 8/15 8/15 8/25 8/19 8/19 9/12 9/30 9/25 9/15 44 64 11 5

CHSP 10/8 10/8 1

BRSP 9/10 9/1 9/6 1 1

LASP 8/20 9/10 9/25 9/15 9/10 9/24 9/16 10/6 9 4 1 4

BTSP 9/30 9/16 9/23 1 1

FOSP 9/23 10/11 10/3 10/9 10/11 3 2

SOSP 9/3 9/3 9/6 2

LISP 9/19 9/23 10/8 9/22 9/26 9/24 9/24 9/29 3 3 1 5

WCSP 9/24 9/30 9/23 9/15 9/23 10/23 10/5 10/8 9/30 4 4 31 9

ORJU 10/10 10/8 10/9 10/8 1 3

LAZB 9/6 9/6 1

PUFI 10/1 10/1 1

HOFI 8/29 10/8 9/18 10/8 1 3
95 

Figure 11. Number of individual migrants captured per species during fall migration, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California, 2013. See tables 2 and 3 for four-letter bird 

species codes. Colored bars represent the six most commonly captured migrant species from 2013 to 2016. Data 

collected by USGS and available from http://www.pwrc.usgs.gov/bbl. 

Figure 12. Number of individual migrants captured per species during fall migration, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California, 2014. See tables 2 and 3 for four-letter bird 

species codes. Colored bars represent the six most commonly captured migrant species from 2013 to 2016. Data 

collected by USGS and available from http://www.pwrc.usgs.gov/bbl. 
96 

Figure 13. Number of individual migrants captured per species during fall migration, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California, 2015. See tables 2 and 3 for four-letter bird 

species codes. Colored bars represent the six most commonly captured migrant species from 2013 to 2016. Data 

collected by USGS and available from http://www.pwrc.usgs.gov/bbl. 

Figure 14. Number of individual migrants captured per species during fall migration, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California, 2016. See tables 2 and 3 for four-letter bird 

species codes. Colored bars represent the six most commonly captured migrant species from 2013 to 2016. Data 

collected by USGS and available from http://www.pwrc.usgs.gov/bbl. 
97 

Figure 15. Number of new migrant species arriving per banding day during fall migration, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California, 2013. Data collected by USGS and 

available from http://www.pwrc.usgs.gov/bbl. 

Figure 16. Number of new migrant species arriving per banding day during fall migration, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California, 2014. Data collected by USGS and 

available from http://www.pwrc.usgs.gov/bbl. 
98 

Figure 17. Number of new migrant species arriving per banding day during fall migration, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California, 2015. Data collected by USGS and 

available from http://www.pwrc.usgs.gov/bbl. 

Figure 18. Number of new migrant species arriving per banding day during fall migration, Naval Base Coronado, 

Remote Training Site, Warner Springs, San Diego County, California, 2016. Data collected by USGS and 

available from http://www.pwrc.usgs.gov/bbl. 
Publishing support provided by the U.S. Geological Survey

Science Publishing Network, Tacoma Publishing Service Center

For more information concerning the research in this report, contact the

Director, Western Ecological Research Center

U.S. Geological Survey

3020 State University Drive East

Sacramento, California 95819

http://werc.usgs.gov
Lynn and others—Monitoring Breeding and Migration of Neotropical Migratory Birds at Naval Base Coronado, Calif., 5-Year Summary 2013–17—Open-File Report 2018-1112

ISSN 2331-1258 (online)

https://doi.org/10.3133/ofr20181112