Recent Foraminifera 

of the 

Marshall Islands

Bikini and Nearby Atolls, Part 2, Oceanography (Biologic)

GEOLOGICAL SURVEY PROFESSIONAL PAPER 260-H
ERRATA SLIP FOR PROFESSIONAL PAPER 260-H

In the distribution tables (1-5) italic numbers indicate abundance 

of the species in percentages of the Foraminifera present in the 

sample. Arabic numbers indicate actual count of specimens.

Recent Foraminifera 

of the 

Marshall Islands

By JOSEPH A. CUSHMAN, RUTH TODD, and RITA J. POST

Bikini and Nearby Atolls, Part 2, Oceanography (Biologic)

GEOLOGICAL SURVEY PROFESSIONAL PAPER 260-H

UNITED STATES GOVERNMENT PRINTING OFFICE, WASHINGTON : 1954
UNITED STATES DEPARTMENT OF THE INTERIOR 

Douglas McKay, Secretary

GEOLOGICAL SURVEY 

W. E. Wrather, Director

For sale by the Superintendent of Documents, U. S. Government Printing Office 

Washington 25, D. C. - Price $1.75 (paper cover)
CONTENTS

Page

______.____-_-__-____-___-.-..______.____ 319

Introduction- ______---_-____--__-__-_-____-____--__ 319

General features_____----------_--_-_-__-________^__ 319

Fauna of reef flats.__.--________-_-___._________ 319

Fauna of lagoons_______________________________ 320

Fauna of outer slopes. __.-_.____-_-_. ___________ 321

Fauna of deep water. ___________________________ 321

Sylvania Guyot core.______________^___-____ 322

Species restricted to guyot-__.___-_-.-_______ 322

Nondiagnostic species._________----_-__^________ 323

Notes on distribution by families. __-.__._._._____ 325

Localities and depths of samples...__.-___^_-_________ 326

Rongerik Lagoon ________._._________^________._ 326

Rongelap Atoll__----_--__-___-_-___---.--______ 326

Bikini Atoll____---_-___-_-_-____-_-__-___-_-_-_ 326

Eniwetok Lagoon____________-__________________ 327

Outer slopes of Eniwetok Atoll___________________ 328

Systematic descriptions-.-__-_--__-____-_-_-____-__-_ 328

Saccamminidae- _---_-_--_____-___-_-_--_____-_- 328

Hyperamminidae. __--___--_________-_-_________ 328

Ammodiscidae. _ _--------_-___-_---___-_________ 328

Textulariidae__-_______--_________.___-____._. 328

Verneuilinidae- _________________________________ 330

Valvulinidae._-------_-_--__-__-_-___-_-_._____ 331

Miliolidae_____--_---_-__--__________.________ 332

Ophthalmidiidae-- _ _____________________________ 341

	Page 

Systematic descriptions—Continued

Trochamminidae. _______________________________ 342

Lagenidae _____________________________________ 342

Polymorphinidae_ ________-_-_-_----_--_-_-_-_-_ 344

Nonionidae_ __-__________________---_-----_---_ 345

Camerinidae ___________________________________ 346

Peneroplidae___________________________________ 348

Alveolinellidae________________________________ 348

Heterohelicidae- ________________________________ 349

Buliminidae_- _ _________-_______-_-__--_--___--_ 349

Ellipsoidinidae____________-_____--_____-_----__ 356

Rotaliidae. ___-----__----_---_--------------__- 356

Pegidiidae._--__-____-------------------___-___ 361

Amphisteginidae- _______________________________ 362

Calcarinidae.__________________________________ 363

Cymbaloporidae. _____-___-__--_-______-__---___ 364

Cassidulinidae_ _ ___-----_--_-_---_----_______. 365

Chilostomellidae- _______-______-_.-___---_______ 367

Globigerinidae________-_-_-_-_-___-__--_______ 368

Globorotaliidae_ _ _______________________________ 370

Anomalinidae. __________--______--___-__-__--__ 370

Planorbulinidae. _ __------___--__-_-___________ 372

Rupertiidae---------------------------_-----_-_ 373

Homotremidae -__________-___-__---_-__-_______ 373

Bibliography _-_______-________-__-_--_---________._ 374

Index..-_------_-_--_-_---_-_----_-_--__--_.______ 379

ILLUSTRATIONS

[Plates 82-93 follow page 384

PLATE 82. Miscellaneous large specimens of Foraminifera from families Verneuilinidae to Homotremidae.

83. Saccamminidae, Textulariidae, Verneuilinidae, Valvulinidae, Miliolidae.

84. Miliolidae.

85. Miliolidae, Ophthalmidiidae, Trochamminidae.

86. Lagenidae, Polymorphinidae, Nonionidae.

87. Camerinidae, Peneroplidae, Alveolinellidae, Heterohelicidae, Buliminidae.

88. Buliminidae, Ellipsoidinidae, Rotaliidae.

89. Rotaliidae, Pegidiidae.

90. Amphisteginidae, Calcarinidae, Cymbaloporidae, Cassidulinidae, Chilostomellidae.

91. Globigerinidae, Globorotaliidae, Anomalinidae, Planorbulinidae, Rupertiidae.

92. Galcarinidae and Homotremidae.

93. Homotremidae.

FIGURE 116. Composition of typical beach sample. _ ...______________-l__._._.___ ____________________

117. Composition of typical lagoon sample_----_-__-_-_______-_-__---_-----__-___-_--_--____-

118. Composition of typical deep-water sample.__--_________-_____________--_-___-_---__-___-

Page 

325

325

326

nx
IV CONTENTS

TABLES

Page

TABLE 1. Distribution of Saccarnminidae, Hyperamminidae, Ammodiscidae, Textulariidae, Verneuilinidae, Valvulinida^, and

Miliolidae in the Marshall Islands_-__________________________________________________________-__ Facing 338

2. Distribution of Miliolidae, Ophthalmidiidae, Trochamminidae, Lagenidae, Poty morphinidae, and Nonionidae in

the Marshall Islands.__________________________________________________________________________ Facing 346

3. Distribution of Camerinidae, Peneroplidae, Alveolinellidae, Heterohelicidae, Buliminidae, and Ellipsoidiniiae in

the Marshall Islands. __________________________________________________________________________ Facing 354

4. Distribution of Rotaliidae, Pegidiidae, Amphisteginidae, Calcarinidae, Cymbaloporidae, Gassidulinidae, and

Chilostomellidae in the Marshall Islarids_--_______-___--__________--____-__-__-___-___-____----__- Facing 362

5. Distribution of Globigerinidae, Globorotaliidae, Anomalinidae, Planorbulinidae, Rupertiidae, and Homotremidae iti

the Marshall Islands___________________________________________________________________ Facing 370
BIKINI AND NEARBY ATOLLS, MARSHALL ISLANDS

RECENT FORAMINIFERA OF THE MARSHALL ISLANDS

By Joseph A. Cushman, 1 Ruth Todd, and Rita J. Post

ABSTRACT

This paper records 331 species and varieties, most of which are 

formally described and illustrated, from 195 samples from the 

lagoons and outer slopes of the Marshall Islands: Rongerik, 

Rongelap, Bikini and Eniwetok. Seventeen species and two 

varieties are new and 33 are undeterminable specifically. The 

samples range in depth from the beach to 835 fathoms on 

Sylvania Guyot adjoining Bikini Atoll. Tables showing the 

distribution and abundance (as percentage of the total) of all 

the species in each sample are included. Species are listed 

according to their ecologic significance or lack of it. Depth, 

and access to the open ocean, appear to be the chief ecologic 

factors affecting distribution and abundance.

INTRODUCTION

A series of bottom foraminiferal samples from four 

coral atolls at the northwestern end of the Marshall 

group in the south Pacific has been studied both as to 

the total fauna present and as to the relative abundance 

of the species at various depths and positions within the 

lagoon and on the outer slopes of the atoll.

The samples available give a rather complete picture 

of the foraminiferal faunas present in the waters of and 

surrounding the atolls. The faunas may be divided as 

follows: reef flat and lagoon beach fauna, lagoon and 

shallow outer slope fauna, and deep- water fauna (chiefly 

a planktonic fauna with the addition of rare benthonic 

species). In each case the faunas may intergrade, or 

dominant species from either of the first two may work 

down the slope into the second or third and obscure 

its normal appearance.

Generally speaking, the faunas here studied are 

characteristic of the tropical Pacific. About 84 percent 

of the species are identified with known species from the 

tropical Pacific. About 6 percent of the species and varieties found seem to be new to science, and 10 percent 

are undeterminable specifically and may also be new. 

The faunas of the four atolls do not differ significantly

1 Dr. Cushman died during the progress of work on this fauna. It was therefore 

brought to completion by the two junior authors. The samples from Rongerik 

Atoll had been studied and the species identified and described by Dr. Cushman, 

and the work on Eniwetok Atoll begun, before his death. The work on the other 

two atolls was done largely by the other authors. The tables showing distribution 

and abundance were prepared by the junior authors. Responsibility for the descriptions of the various new species and varieties is indicated in each case.

among themselves, but there are notable differences 

between these atolls and certain other Pacific islands; 

for example, the absence of Baculogypsina and vr.rious 

medium to large species of Rotalia, such as E. calcar 

(D'Orbigny), and the great variety of the miliolids.

The percentages recorded hi the distribution table 

were estimated by means of a picking tray divided into 

segments to facilitate the counting of all specimen? in a 

unit area of a thinly scattered sample. Depending 

upon the variety of forms present, fifty to several 

hundred specimens were counted in each sample. In. 

the original picking of the samples the species were 

picked quantitatively up to 25 specimens. Beyond 

that point, percentages were estimated, but where the 

percentages are low the estimates are very rough.

In most cases the samples were treated with carbon 

tetrachloride to separate the smaller species from the 

larger and heavier tests of Cakarina, Amphistegina, 

Hezerostegina, Marginopora, etc., and from the fragments of coral, shell, and Halimeda, but both fractions 

were studied.

The most abundant single species in nearly all the 

lagoon samples (exclusive of beach samples) is Amphistegina madagawarientiis D'Orbigny, but its varying 

percentages are not so much significant of changes in 

its abundance as a reflection of the richness or barrenness of the accompanying fauna.

We appreciate the many helpful suggestions received 

during discussions of our work with Miss Frances L. 

Parker of Scripps Institution of Oceanography, Dr. W. 

Storrs Cole of Cornell University, Drs. Harry S. Ladd 

and Joshua I. Tracey of the U. S. Geological Survey 

and Dr. Kenneth O. Emery of the University of Southern California.

GENERAL FEATURES 

FAUNA OF BEEF FLATS

The fauna of the reef flat, represented by seven 

samples from Bikini, is characterized by large percentages of Cakarina spengleri (Gmelin), Margiropora 

vertebralis Blainville, Homotrema rubrum (Lamarck),

319
320 BIKINI AND NEARBY ATOLLS, MARSHALL ISLANDS

Miniacina miniacea (Pallas), and Carpenteria proteiformis Goes together with much smaller percentages of 

Amphistegina madagascariensis D'Orbigny. In addition 

there are some thirty other species which comprise a 

very small percentage of the total fauna and these are 

found mainly in pools on the reef flat. Local variations 

such as channels or pools where the water would be 

swift-moving or quiet affect the concentration of the 

species on the reef flat. A few specimens of planktonic 

forms brought in by the surf are found on the reef flat. 

The fauna of the beaches is almost entirely a transported one and consists mainly of much-worn specimens of Calcarina spengleri (Gmelin). The percentage 

of C. spengleri (Gmelin) on the beaches varies from 

place to place and in some places there is a considerable 

percentage of worn specimens of Amphistegina madagascariensis D'Orbigny and Marginopora vertebralis 

Blainville.

FAUNA OF LAGOONS

The fauna of the lagoons was studied in 13 samples 

from Rongerik ranging in depth from 74 to 150 feet, 

12 samples from Rongelap ranging in depth from 60 

to 192 feet, 58 samples from Bikini ranging in depth 

from 11 to 188 feet, and 45 samples from Eniwetok 

ranging in depth from 54 to 204 feet.

In the lagoon fauna as a whole, the dominant species 

is Amphistegina, madagascariensis D'Orbigny. Next in 

abundance in the lagoons is Heterostegina suborbicularis 

D'Orbigny, which in a few places is more abundant 

than Amphistegina. The dominance of Heterostegina 

suborbicularis D'Orbigny over Amphistegina madagascariensis D'Orbigny is probably related to bottom 

conditions. (See Meyers, 1943, p. 29.) Furthermore, 

the distribution of these two common species may be 

affected by additional unknown factors.

Leaving out of consideration the dominant one or 

several species, the remaining lagoon fauna may be rich 

in one or another of the Eeussella-Bolivina, Textularia, 

milioid, peneroplid, or globigerinid groups. The reasons 

for these variations are also obscure but probably 

bottom conditions have a very important part in 

determining what foraminiferal species are to be found. 

Many factors other than depth affect these faunas, 

which are not entirely life associations.

Other species found in considerable abundance in the 

lagoons are Calcarina hispida H. B. Brady, Marginopora vertebralis Blainville, Homotrema rubrum 

(Lamarck), and Miniacina miniacea (Pallas). Certain 

samples, the shallower ones and those close to the reef, 

have rather large percentages of worn specimens of 

Calcarina spengleri (Gmelin) which appear to have 

been transported down into the lagoon from the reef 

flat. In general, the percentage of C. spengleri (Gmelin)

decreases with depth and that of (7. hispida H. B. 

Brady increases.

The seven species mentioned above are a^ follows:

Heterostegina suborbicularis D'Orbigny 

Marginopora vertebralis Blainville 

Amphistegina madagascariensis D'Orbigny 

Calcarina hispida H. B. Brady

spengleri (Gmelin) 

Homotrema rubrum (Lamarck) 

Miniacina miniacea (Pallas)

These species may comprise from 50 to 75 percent 

of the total lagoon fauna. The remaining percentage is 

made up of about 185 other species, many of which are 

common and about 60 of which are very rare.

The following species are restricted to the lagoons (or 

with very rare occurrences outside):

Family Miliolidae

Quinqueloculina anguina var. arenata Said

Spiroloculina clara var. lirata Cushman, n. var. 

foveolata Egger

Hauerina serrata Cushman, n. sp.

Triloculina sp. A 

Family Ophthalmidiidae

Nubecularia lacitnensis Chapman 

Family Peneroplidae

Monalysidium politum Chapman 

Family Buliminidae

Bulirninella milletti Cushman

Bulimina fijiensis Cushman

Reus&ella sp. A 

Family Rotaliidae

Spirillina decorata H. B. Brady

vivipara var. revertens Rhumbler

Patellinella inconspicua (H. B. Brady)

Discorbis subbertheloti Cushman

Rotalia cf. R. beccarii var. tepida Cushmrn

Epistomaroides polystomelloides (Parker and Jones)

Siphonina tubulosa Cushman 

Family Pegidiidae?

Rugidia? spinosa Cushman, n. sp. 

Family Globigerinidae?

Globigerina? sp. 

Family Planorbulinidae

Gypsina plana (Carter)

One core, Bikini 218, taken inside Bikini lagoon about 

3 miles southwest of Bikini Island, and 2^ miles from 

the reef at its nearest place, was available for study. 

The sample showed a distinct difference in the composition of the fauna from top to bottom of the 16-inch core. 

The abundance of Globigerinoides conglo^ata (H. B. 

Brady) at the bottom of the core indicates that conditions had been different, resulting in a greater concentration of planktonic species and those characteristic 

of the outer slope. This might have been due to the 

proximity of a gap in the reef at the time of deposition 

of that part of the core, or to a change in the currents 

within the lagoon. At present, the pass nearest to the
RECENT FOBAMINIFEBA OF THE MARSHALL ISLANDS 321

location of the. core is about 6 miles to the south. The 

presence in considerable numbers at the bottom of the 

core of Globigerinoides conglobata (H. B. Brady) is significant in view of the fact that the species is not found, 

except very rarely, in any other of the lagoon samples, 

and occurs in abundance only in the deeper samples of 

the outer slopes and guyot. The presence of a considerable percentage of Calcarina spengleri (Gmelin) at 

the bottom of the core is consistent with other lagoon 

samples, however.

The two commonest pelagic species, Globigerinoides 

rvbra (D'Orbigny) and G. sacculifera, (H. B. Brady), 

are found almost throughout all the lagoons but are less 

abundant and less well developed there, as if those in 

the lagoons were all young specimens. The reasons for 

this are obscure. They may become more apparent as 

more is learned of the life history and the food supply 

of planktonic Foraminifera.

FAUNA OF OUTER SLOPES

The samples from the outer slopes of the atolls do not 

differ greatly in percentage composition from those inside the lagoons. They are, however, characterized by 

the appearance, as rare specimens, of various deepwater species that occur in greater abundance at greater 

depths. Cycloclypeus carpenteri H. B. Brady is characteristic of the outer slopes of Bikini Atoll from 58 to 

240 fathoms and was not found, except very rarely, at 

greater depths, such as on the guyot.

The following species occur only on the outer slopes 

of the atolls:

Siphotextularia crispata (H. B. Brady)

Gaudryina (Siphogaudryina) siphonifera (H. B. Brady)

Cycloclypeus carpenteri H. B. Brady

Discorbis tuberocapitata (Chapman)

Cassidulina pacifica Cushman

FAUNA OF DEEP WATER

The deep-water fauna is represented by 13 bottom 

samples on the outer slopes of Bikini Atoll from 55 to 

710 fathoms, 21 samples from Sylvania Guyot adjoining Bikini Atoll from 410 to 835 fathoms, and five samples from 24 to 500 fathoms on the outer slope of Eniwetok Atoll. The percentage composition of this fauna 

is entirely different from that of the reef flat or the lagoons. In the samples from the deeper locations, various species of the planktonic families Globigerinidae 

and Globorotaliidae together comprise 50 percent to 

98 percent of the total foraminiferal fauna. In the 

samples from the shallower locations the next most 

abundant species are Amphistegina, madagascariensis 

D'Orbigny and Amphistegina, radiata (Fichtel and Moll) 

and its varieties. In addition to these there are about

215 other species which are mostly rare and together 

form a relatively small proportion of the deep water 

fauna. Kepresentatives of the families Lagenidae, 

Polymorphinidae, Ellipsoidinidae, Cassidulinidae and 

Chilostomellidae are found almost exclusively in the 

deeper-water fauna.

The following species are restricted to the obiter 

slopes of the atolls and to the guyot, with rare occurrences inside the lagoons in a few casss:

Family Textulariidae

Textularia kerimbaensis Said

milletti Cushman

semialata Cushman 

Bigenerina sp. 

Family Verneuilinidae

Gaudryina triangularis var. angulata Cushman

trullissata Todd, n. sp.

(Siphogaudryina) transversaria (H. B. Brady) 

Family Valvulinidae

Karreriella bradyi (Cushman) 

Family Miliolidae

Quinqueloculina bradyana Cushman 

Spiroloculina circularis Chapman 

Pyrgo lucernula (Schwager)

murrhina (Schwager) 

Family Lagenidae

Robulus calcar (Linne")

gibbus (D'Orbigny)

limbosus (Reuss)

nigriseptus (Koch)

sp. A

sp. B

sp. C

sp. D 

Astacolus bradyi (Cushman)

planulata (Galloway and Wissler) 

Dentalina cf. D. filiformis (D'Orbigny) 

Nodosaria pauciloculata Cushman

sp. B

sp. C

Frondicularia sp. A of Boomgaart 

Lagena acuticosta Reuss

gracilis Williamson

Family Polymorphinidae

Dirnorphina tuberosa D'Orbigny 

Polymorphinella pacifica Cushman and Hanzawa 

Family Nonionidae

Nonion pacificum (Cushman) 

Astrononion tumidum Cushman and Edwards 

Family Buliminidae 

Fissurina sp. B

Bolivina abbreviata Heron-Alien and Earland 

nitida H. B. Brady 

robiista H. B. Brady 

Bitubulogenerina sp. 

Uvigerina ampullacea H. B. Brady 

proboscidea Schwager 

proboscidea var. vadescens Cushnian 

Siphogenerina dimorpha var. pacifica Cushman 

Angulogerina albatrossi var. ornata Cushman 

Trifarina bradyi Cushman
322 BIKINI AND NEARBY ATOLLS, MARSHALL ISLANDS

Family Rotaliidae

Gyroidina soldanii D'Orbigny 

Eponides tenera (H. B. Brady) 

Hdglundina elegans (D'Orbigny) 

Missii>sippina concentrica (Parker and Jones) 

Baggina philippinensis (Cushman)

philippinensis var. pilulifera Cushman and Todd 

Family Amphisteginidae

Amphistegina radiata (Fichtel and Moll)

radiata var. papillosa Said

radiata var. venosa (Fichtel and Moll)

radiata (Fichtel and Moll) var. 

Family Cassidulinidae

Cassidulina angulosa Cushman

costatula Cushman

delicata Cushman

elegans Sidebottom

marshallana Todd, n. sp.

patula Cushman

cf. C. spinifera Cushman and Jarvis

subglobosa H. B. Brady

subglobosa H. B. Brady var. 

Ehrenbergina pacifica Cushman 

Family Chilostomellidae

Pullenia salisburyi R. E. and K. C. Stewart 

Sphaeroidina bulloides D'Orbigny

compacta Cushman and Todd 

Family Globigerinidae

Globigerina cf. G. conglomerate/, Schwager

subcretacea Lomnicki

Globigerinella aequilateralis (H. B. Brady) 

Hastigerinella digitata Rhumbler 

Orbulina universa D'OrbignjPulleniatina obliquiloculata (Parker and Jones) 

Sphaeroidinella dehiscens (Parker and Jones) 

Candeina nitida D'Orbigny 

Family Globorotaliidae

Globorotalia crassa (D'Orbigny)

cf. G. Ursula (D'Orbigny)

menardii (D'Orbigny)

truncatulinoides (D'Orbigny)

tumida (H. B. Brady) 

Family Anomalinidae

Anomalina polymorpha Costa 

Planulina ariminensis D'Orbigny 

Cibicides cicatricosus (Schwager) 

Cibicidina sp.

SYI/VANIA GUYOT CORE

One of the cores from Sylvania Guyot, Bikini 1176, 

7 inches long, taken at a depth of 720 fathoms, contains 

a few species which suggest Tertiary age. These are

Gumbelinal marshallana Todd, n. sp. 

Pavonina triformis Parr 

Parafissurina sp.

Pleurostomella bolivinoides Schubert 

Osangularia bengalensis (Schwager)

The evidence is not conclusive and needs further support 

from additional samples. 

The position of the core in which these fossil species

were found is at the inner part of the neck connecting 

Sylvania Guyot to the base of the atoll ^ope. (See 

figure in chapter A of the present report.) Such a position, about 4300 feet below the top of the atoll would be 

favorable for the accumulation of any Tertiary or older 

material that might be exposed on the atoll slope above. 

Evidence from the well drilled to a depth of about 2500 

feet on Bikini and which penetrated beds thought to be 

Oligocene, indicates the probable existence of lower 

Tertiary beds as much as 1800 feet higher than the 

elevation of the core, Bikini 1176.. Thu? there is a 

possibility that the origin of these Tertiary species is an 

exposure somewhere on the atoll slope above, which 

sheds debris down onto the more gradual slope below. 

Only one of the species listed above, Favonina triformis Parr, was found in the material from the Bikini 

well at depths of 925-935)^ feet and 1167-1177# feet.

SPECIES RESTRICTED TO GTJYOT1

The following species are restricted to the guyot:

Family Textulariidae

Siphotextularia cf. S. concava (Karrer) 

Family Verneuilinidae

Gaudryinaflintii Cushman 

Family Valvulinidae

Schenckiella primaeva (Cushman) 

Family Miliolidae

Triloculina sp. B 

Family Lagenidae

Nodosaria sp. A 

Family Nonionidae

Nonion pompilioides (Fichtel and Moll) 

Family Heterohelicidae

Giimbelinal marshallana Todd, n. sp.

Bolivinita quadrilalera (Schwager) 

Family Buliminidae

Robertina subcylindrica (H. B. Brady) 

translucens Cushman and Parker

Bulimina aculeata D'Orbigny 

affi,nis D'Orbigny 

costata D'Orbigny 

rostrata'R. B. Brady

Fissurina sp. C

Loxostomum karrerianum (H. B. Brady)

Rectobolivina bifrons (H. B. Brady)

Uvigerina cf. U. peregrina Cushman 

porrecta var. fimbriata Sidebottom

Siphonodosaria abyssorum (H. B. Brady)

lepidula (Schwager) 

Family Rotaliidae

Osangularia bengalensis (Schwager) 

Family Cassidulinidae

Cassidulina gemma Todd, n. sp.

Ehrenbergina reticulata Cushman 

Family Chilostomellidae

Pullenia bulloides (D'Orbigny) 

Family Globorotaliidae

Globorotalia menardii \ax.fimbriata (H. B. Brady)
RECENT FORAMINIFERA OF THE MARSHALL ISLANDS 323

NONDIAGNOSTIC SPECIES

The following species are not very diagnostic because 

of their widespread distribution. However, those preceded by (*) are very rare or absent on the guyot, and 

those preceded by (**) are less abundant inside the 

lagoon.

Family Hyperamminidae

*Sageninafrondescens (H. B. Brady) 

Family Textulariidae

Textularia agglutinans D'Orbigny

candeiana D'Orbigny

cornea D'Orbigny

dupla Todd, n. sp.

*foliacea Heron-Alien and Earland

foliacea var. oceanica Cushman 

Family Verneuilinidae

Gaudryina (Siphogaudryina) rugulosa Cushman 

Family Valvulinidae

Valvulina davidiana Chapman 

Family Miliolidae

Quinqueloculina bidentata D'Orbigny

crassa var. subcuneata Cushman

distorqueata Cushman, n. sp.

cf. Q.ferussacii D'Orbigny

neostriatula Thalmann

parkeri (H. B. Brady)

cf. Q. seminulum (Linn6)

sulcata D'Orbigny 

Miliolinella australis (Parr) 

Triloculinella labiosa (D'Orbigny) 

Spiroloculina angulata Cushman

clara Cushman

communis Cushman and Todd

corrugata Cushman and Todd

marshallana Todd, n. sp. 

Articulina pacifica Cushman 

Hauerina bradyi Cushman

diversa Cushman

involuta Cushman

milletti Cushman, n. sp.

Schlumbergerina alveoliniformis (H. B. Brady) 

Triloculina cf. T. bassensis Parr

cf. T. bicarinata D'Orbigny

bikiniensis Todd, n. sp.

earlandi Cushman, n. sp.

involuta Todd, n. sp.

irregulari&> (D'Orbigay)

kerimbatica (Heron-Alien and Earland)

marshallana Todd, n. sp.

cf. T. oblonga (Montagu)

spinata Cushman

subplanciana Cushman, n. sp.

terquerniana (H. B. Brady)

tricarinata D'Orbigny

trigonula (Lamarck) 

Pyrgo denticulata (H. B. Brady)

denticulata var. striolata (H. B. Brady)

sp. 

Family Ophthalmidiidae

Cornu&pira planorbis Schultze 

Parrina bradyi (Millett) 

291053—54———2

Family Nonionidae

Nonionella translucens Cushman

sp. B 

Elphidium advenum (Cushman)

advenum var. dispar Cushman, n. var. 

jenseni (Cushman) 

simplex Cushman 

Family Camerinidae

Operculina ammonoides (Gronovius)

*Heterostegina suborbicularis D'Orbigny 

Family Peneroplidae

Spirolina acicularis (Batsch)

arietina (Batsch) 

Sorites marginalis (Lamarck)

*Marginopora vertebralis Blainville 

Family Heterohelicidae

Bolivinella folia (Parker and Jones)

folia var. ornata Gushman 

Family Buliminidae

Fissurina circularis Todd, n. sp.

formosa (Schwager)

lacunata (Burrows and Holland)

milletti Todd, n. sp. 

Bolivina compacta Sidebottom

pseudopygmaea Cushman

rhomboidalis (Millett)

striatula Cushman

subexcavata Cushman and Wickenden

tortuosa H. B. Brady 

Loxostomum convallarium (Millefct)

Hmbatum (H. B. Brady) 

Reussella simplex (Cushman) 

Mimosina pacifica Cushman 

Uvigerina porrecta H. B. Brady 

Siphogenerina raphana (Parker and Jones) 

Family Rotaliidae

*Discorbis concinna (H. B. Brady) 

crustata Cushman

*rnicens Cushman 

opima Cushman

*patelliformis (H. B. Brady)

*rugosa (D'Orbigny)^

*Poroeponides cribrorepandus Asano and TJchio

*Siphoninoides echinata (H. B. Brady) 

Family Pegidiidae

Rugidia corticata (Heron-Alien and Earland) 

Family Amphisteginidae

Amphistegina madagascariensis D'Orbigny 

Family Calcarinidae

Calceerinet hd»pida,tS .* B. Brady 

Family Cymbaloporidae

Cymbaloporetta bradyi (Cushman) 

squammosa (D'Orbigny)

Tretomphahis planus Cushman 

Family Cassidulinidae

Epistominella tubulifera (Heron-Alien and Earland)

sp.

Cassidulina minuta Cushman 

Family Globigerinidae

Globigerina bulloides D'Orbigny

Globigerinoides conglobata (H. B. Brady)

**rubra (D'Orbigny)

**sacculifera (H. B. Brady)
324 BIKINI AND NEAKBY ATOLLS, MAKSHALL ISLANDS

Family Anomalinidae

Anomalina cf. A. glabrata Cushman

*Anomalinella rostrata (H. B. Brady) 

Cibicides lobatulus (Walker and Jacob) 

mayori (Cushman) 

cf. C. pseudoungerianus (Cushman) 

Cibicidella variabilis (D'OrbignyJ 

Family Planorbulinidae

*Planorbulina acervalis H. B. Brady

*Acervulina inhaerens Schultze

*Gypsina globula (Reuss)

*vesicularis (Parker and Jones) 

Family Homotremidae

*Homotrema rubrum (Lamarck)

*Miniacina miniacea (Pallas)

The following species are too rare to be of value as 

indicators of ecologic conditions:

Family Saccamminidae

Proteonina sp. 

Family Ammodiscidae

Ammodiscus incertus (D'Orbigny) 

Family Verneuilinidae

Gaudryina cf. G. pauperata Earland 

Family Valvulinidae

Clavulina angularis D'Orbigny

pacifica Cushman 

Family Miliolidae

Quinqueloculina parkeri (H. B. Brady) var.

cf. Q. samoaensis Cushman 

Miliolinella oceanica (Cushman)

oceanica var. flintiana (Cushman) 

Massilina planata Cushman 

Spiroloculina acescata Cushman 

aequa Cushman 

mayori Cushman 

sp. A 

sp. B

Sigmoilina edwardsi (Schlumberger) 

Articulina elongata Cushman 

Tubinella funalis (H. B. Brady) 

Pyrgo millettii (Cushman) 

Family Ophthalmidiidae *

Vertebralina striata D'Orbigny 

Planispirina exigua (H. B. Brady) 

WiesnereUa auriculata (Egger) 

Family Trochamminidae

Trochammina cf. T. rotaliformis J. Wright 

Rotaliammina mayori Cushman 

Carterina spiculotesta (Carter)

Nouria polymorphinoides Heron-Alien and Earland 

Family Lagenidae (except as indicators of outer slopes and

deep water)

Lagena globosa (Montagu) 

spiralis H. B. Brady 

striato-punctata Parker and Jones 

williamsoni (Alcock) 

Family Polymorphinidae

Sigmomorphina semitecta var. terquemiana (Fornasini) 

Family Nonionidae 

Nonionella sp. A 

Elphidium milletti (Heron-Alien and Earland)

Family Peneroplidae

Peneroplis sp. 

Family Alveolinellidae

Neoalveolina pulchra (D'Orbigny) 

Family Buliminidae

Buliminella elegantissima (D'Orbigny) 

Buliminoides williamsoniana (H. B. Brady) 

Pseudobulimina sp. 

Oolina squamosa (Montagu) 

Fissurina auriculata var. costata (H. B. P^ady) 

radiato-marginata (Parker and Jones) 

sp. A

Virgulina earlandi Cushman 

Bolivina globulosa Cushman 

oceanica Cushman 

subreticulata Parr 

subtenuis Cushman 

Loxostomum mayori (Cushman) 

Bifarinella mackinnonii (Millett) 

Reussella sp. B 

Pavonina triformis Parr 

Chrysalidinella fijiensis Cushman 

Family Ellipsoidinidae

Pleurostomella bolivinoides Schubert 

Parafissurina sp. 

Family Rotaliidae

Spirillina decorata var. of Sidebottom 

inaequalis H. B. Brady 

tuberculato-limbata Chapman 

vivipara var. densepunctata Cushman 

Conicospirillina semi-involuta Cushman

sp. 

Patellina advena Cushman

advena var. altiformis Cushman 

Patellinella fijiensis Cushman

jugosa (H. B. Brady) 

Discorbis frustata Cushman 

Lamarckina scabra (H. B. Brady) 

Heronallenia sp. 

Paumotua terebra (Cushman) 

Family Pegidiidae

Pegidia dubia (D'Orbigny) 

Sphaeridia papillata Heron-Alien and Er.rland 

Family Cymbaloporidae

Pyropilus rotundatus Cushman 

Family Cassidulinidae

Ceratobulimina pacifica Cushman and Harris 

Epistominella cf. E. obtusa (Burrows anc1 Holland)

pulchra (Cushman) 

Family Chilostomellidae

Chilostomella oolina Schwager 

Family Globigerinidae

Globigerinoides elongata (D'Orbigny)

sacculifera var. fistulosa (Schubert) 

Family Anomalinidae

Laticarinina pauperata (Parker and Jones) 

Family Planorbulinidae

Planorbulinella larvata (Parker a 

Family Rupertiidae

Rupertia stabilis Wallich 

Carpenteria sp.
KECENT FORAMINIFERA OF THE MAKSHALL ISLANDS 325

NOTES ON DISTRIBUTION BY FAMILIES

Textulariidae and other arenaceous groups are 

present at all the atolls, less abundantly in deeper 

water and with certain species being very abundant in 

the lagoons. Miliolidae and Ophthalmidiidae are 

present and abundant at all the atolls, but are rare in 

deeper water. Lagenidae and Polymorphinidae are not 

present in Roiigerik lagoon. These groups are not 

found abundantly anywhere in the Marshall Islands, 

but their frequency is greater at the deeper locations. 

Nonionidae are present at all the atolls. Cameriniclae 

are present at all the atolls, comprising a considerable 

percentage of the lagoon fauna, but are rare in deeper 

water. Peneroplidae are present at all the atolls, rare 

in deeper water. One species, Marginopora vertebralis 

Blainville, comprises a considerable percentage of the 

reef fauna. Heterohelicidae and Ellipsoidinidae are 

found only very rarely. Buliminidae occur rarely at all 

the atolls, with fewer numbers but a greater variety of 

genera and species in deeper water. Rotaliidae are 

present at all the atolls but are rare. Amphisteginidae 

are present in large numbers at all the atolls, decreasing 

in numbers but increasing in variety of forms with 

depth en the outer slopes of the atolls. Calcarinidae 

are present at all the atolls. The two species found in 

this family are apparently restricted in their habitats, 

although the one reef-dwelling species, Calcarina 

spengleri (Gmelin),is found washed out great distances, 

both into the lagoons and down the outer slopes of the 

atolls. The other species, C. hispida H. B. Brady, 

flourishes within the lagoon at about 10 fathoms and 

deeper as well as in deep water on the outer slopes of the 

atolls. Cymbaloporidae are found equally in the 

lagoons and in the deeper water surrounding the atolls. 

Cassidulinidae and Chilostomellidae are found almost 

exclusively in the deep water on the outer slopes 

surrounding the atolls. There is only one notable 

exception, Cassidulina minuta Cushman, which is 

found as frequently inside the lagoons as out. Globigerinidae and Globorotaliidae are found most abundantly in the oozes from the deepest samples. These 

planktonic forms are also found inside the lagoons but 

in much smaller numbers. Anomalinidae and Planorbulinidae occur at all the atolls, but more rarely in 

deeper water. Homotremidae are found most abundantly in the shallower parts of the lagoons and very 

rarely in deep water.

Three generalized diagrams show the differences in 

percentage composition of the three types of environment. Figure 116 represents the composition of a

typical beach sample, figure 117 that of a typical 

lagoon sample, and figure 118 that of a typical deepwater sample.

Amphisfegma 

madagascariensis

FIGURE 116.—Composition of typical beach sample

FIGURE 117.—Composition of typical lagoon sample
326 BIKINI AND NEARBY ATOLLS, MARSHALL ISLANDS

Sample

Inside lagoon:

B-74

6

483

111

348

484 '

71

480

76

183

133

272

30

171

RONGELAP

Depth

ATOLL

(fathoms) Latitude (N.)

Beach sandstone,

Tufa Island.

+ %

-10

-13

-14

* -16

-18

-20

-22

-24

-26

-28

-30

-32

Longitude (E.)

lagoon beach, west end of

11°13'43"

14'19"

25W

10'12"

14'18"

20'25"

14'08"

22'24"

11'30"

23'30"

20'13"

16'34"

14'32"

166°53'50"

38'17"

167°01'48"

166°50'20"

38'30"

55'00"

39'00"

57'02"

48'48"

56'21"

40'46"

51'48"

48'46"

FIGURE 118.—Composition of typical deep-water sample

LOCALITIES AND DEPTHS OF SAMPLES

Outside lagoon:

468

463

456

462

-21

-32

-40

-49

11°27'20" 

27'37" 

22'57" 

26'42"

166°43'43"

54'26"

167°01'18"

166°58'23"

BIKINI ATOLL

Beach samples:

B-l Beach rock, east center, Bikini Island. 

B-20 Sand from lagoon beach, north end, Bikini Island. 

B-58 Polished beach sandstone, lagoon side of sand bar, at

west end of Uorikku Island, Bikini. 

B-105 Sand from "old reef zone" near micMle of Bikini

Island. 

734 Latitude 11°36'45" N., Longitude 165°32'59" E.

The samples examined in the course of this study

were obtained from the localities and depths indicated

l"»plr»w IL/OJ.V/ W •

Sample

99

102

8ft

56

54 

91

3

31

51

12

21

70

57

18

79

RONGERIK

Depth

(feet)

+ 3

+ 3

-74

-76

-85 

-95

-101

— 102

-111

-120

-127

-130

-138

-150

-150

LAGOON

Latitude (N.)

11°23'10"

23'56"

23'02"

23'30"

23'35" 

21'20"

22'34"

21'36"

22'32"

21'02"

21'55"

19'08"

22'57"

21'00"

22'36"

Longitude (E.)

167°21'54"

29'00"

26'53"

27'40"

28'35" 

23'06"

22'40"

25'44"

28'48"

24'05"

24'52"

26'36"

27'39"

24'48"

26'55"

Ree f flat samples:

B-23

B-52

B-72

B-107

B-132

B-135

B-305

Sample

From inside of I

37

33

34

Sand from reef flat, northwest end of Bikini Island

(Traverse 28).

Channel bottom, 6 feet, algal nodules from pools,

Oruk Island, Bikini.

Edge of pothole in reef, 100 yards from edge of reef,

Yi mile northwest of Beacon San, Bikini Island

(Traverse 26).

From a piece of living Lithothamnion, from landward edge of algal flat, 1 mile north of south end 

of Bikini Island.

Sand scraped from surface of reef flat, from rock

flat zone (Traverse 5) , about % mile north of Enyu

Island, Bikini.

Sand from inside edge of reef flat in shallow tidal 

pool, half way between Bikini and Aomoen Islands 

(Traverse 26).

Picked from small cavities in surface reef rock, sea

reef northeast of Bikini Island, 1,400 feet out from

shore in 1 foot of water at low tide.

Depth

(feet) Latitude (N.) Longitude (E.)

igoon:

-11 11°36'45" 165°32'55"

-22 36'13" 32'42"

-35 36'12" 32'40"
RECENT FORAMINIPERA OF THE MARSHALL ISLANDS

Sample

From inside of lagoon:

B-307

473

96

149

559

190

B-45

B-46

76

401

366 

384 

567

938

402

356

177

569

1052

1055

1012

165

1022

576

933

460

413

444

1116

1056

546

1117

580. 1

423

431

573

582

218(0"-4")

91SM9" 1 fi"^ ilo\,i* — J.O )

1010 

1028

1143

326

377

482

478

462

292

443

947

245

591

593

311

445

Depth

(feet)

-35

-38

-46

-57

-64

-69

-72

-72

-85

-90

-97 

-101 

-104

-104

-105

-118

-119

-123

-125

-128

-130

-132

-134

-140

-145

-147

-148

-148

-149

-154

-155

-157

-158

-159

-160

-160

-160

-161

-163 

-164

-165

-165

-171

-172

-176

-178

-179

-179

-179

-180

-180?

-180?

-184

-188

Latitude (N.) Longitude (E.)

From coral knoll in lagoon, top

of knoll 35 feet under water,

1 mile north of Eninman

Island, Bikini.

11°40'03" 165°21'27"

36'59" 32'05"

35'47" 32'13"

33'58" 13' 10"

36'19" 31'40"

Off Bikini Island about 1%

miles.

Off Bikini Island about \%

miles.

11°36'16" 165°31'28"

33'08" 16'14"

34'00" 14'03" 

40'18" 15'22" 

34'58" 13'55"

31'16" 27'27"

32'40" 16'14"

37'15" 13'34"

36'37" 30'34"

35'13" 14'12"

30'33" 29'18"

35'09" 24'32"

31'33" 19'33"

36'29" 29'56"

31'30" 25'10"

35'40" 15'25"

30'52" 31'02"

33'47" 21'27"

35'05" 17'42"

35'25" 22'57"

39'56" 23'50"

34'55" 19'56"

35'08" 29'57"

39' 18" 23'14"

35'37" 16'40"

• 36'54" 17'42"

40'24" 17'41"

35'35" 14'44"

35'30" 16'58"

35'23" 30'44"

32'24" 18'08" 

34'38" 28'47"

35'25" 17'28"

33'27" 24'48"

37'14" 14'52"

35' 17" 20'20"

37'52" 20'04"

35'00" 21'22"

33'45" 27'26"

36'27" 22'39"

33'47" 28' 18"

35'08" 27'50"

35'32" 18'02"

35'32" 18'26"

35'19" 26'05"

34'57" 22'48"-

Sample

From outer slopes:

Dredge samples:

1478

1493

1461

1458

1460

Bottom samples:

815

812

817

814

816

1173(0"-!"}

1 1 7^C1 rl—V\ 1 1 / o v 1 O /

1173(3"-5")

1173(5"-7"; 

1173(7"-9") 

819

818

830

Ouyot samples:

1174(l"-2")

1174(4"-6")

1174(6"-8")

1174(8"-10")

1174(10"-14">

1175

1175(0"-!")

1175(l"-2")

1175(2"-3")

1175(3"-4")

1175(4"-5")

1176(0"-!")

1176(l"-2")

1176(2"-3")

1176(5"-7")

1169

1170(0"-!")

1170(l"-2")

1170(2"-4")

H70(4"-6")

1172

Sample

10

11

120

231

336

332

286

287

288

289

290

293

Depth

(fathoms)

-13-8

-21-25

-44-25

-48-33

-91-58

-55

— 80

-114

-136

-219

-240

-315

-500

-710

— 410

-592

-592

-720

-710

-680

— 835

ENIWETOK

Depth

(fathoms)

+ 0.5

+ .5

+ .5

-9

-10

-12

-16

-18

-20

-22

-22

-23

Latitude (N.)

11°31'52"

29'26"

38' 26"

29'06"

38'27"

11°34'44"

29'31"

35'06"

29'18"

35'53"

40'40"

35' 53"

35'42"

50'30"

11°41'00"

42'33"

42'33"

44'47"

48'00"

48' 20"

47*35"

LAGOON

Latitude (N.)

11°21'15"

32'53"

33'30"

20'33"

30' 57"

31'42"

22'13"

22'20"

22'29"

22'38"

22'49"

23'24"

327

Longitude (E.)

165°32'53"

32'00"

31'43"

20'52"

31'46"

165°33'40"

32'19"

33'34"

30'59"

33'30"

14'05"

33'30"

33'52"

164°56'00"

165°14'01"

12'45"

12'45"

09'56"

01'52"

C1'27"

164°£9'20"

Longitude (E.)

162°J2'30"

Tl'35"

Tl'35"

15'50"

22'00"

SI'17"

12'28"

12'33"

12'41"

12'50"

12'59"

13'30"
328 BIKINI AND NEARBY ATOLLS, MARSHALL ISLANDS

Sample

329

218

291

292

296

145

326

328

295

298

321

323

299

305

325

310

317

318

320

324

300

301

302

307

312

313

314

316

322

151

303

304

306

311

183

49

Depth

(fathoms)

-23

-24

-24

-24 

, -24

-24

-24

-24

-26

-26

-26

-26

-27

-27

-27

-28

-28

-28

-28

-28

-29

-29

-30

-30

-30

-30

-30

-30

-30

-30

-31

-32

-32

-32

-34

-34

Latitude (N.) 

31'40"

22'59" 

23'11" 

24'16" 

27'19" 

31'15" 

31'29" 

24'02" 

24'41" 

30'16" 

30'36" 

24'54" 

26'37" 

31'03" 

27'55" 

29'08" 

29'28" 

29'57" 

30'51" 

25'07" 

25'29" 

25'41" 

27'07" 

28'18" 

28'30" 

28'38" 

28'59" 

30'23" 

27'55" 

25'55" 

26'08" 

26'49" 

28'05" 

33'26" 

32'00"

Longitude (E.) 

20'53"

13'08" 

13'19" 

14' 10" 

16'25" 

20'27" 

20'41" 

14'00" 

14'28" 

19'35" 

19'54" 

14'39" 

15'46" 

20'16" 

17'13" 

18'33" 

18'53" 

19'18" 

20'05" 

14'50" 

15'08"

17'37" 

17'49" 

18'GO" 

18'23" 

19'45" 

15'58" 

15'20" 

15'28" 

15'55" 

17'23" 

15'30" 

17'47"

OUTER SLOPES OF ENIWETOK ATOLL

Sample

118

117

116

115

119

Depth 

(fathoms)

-24

-88

-173

-214

-500

Latitude (N.) 

11°20'06" 

19'43" 

19'35" 

19'30" 

18'52"

Longitude (E.) 

162°16'19" 

16'24" 

16'37" 

16'40" 

15'24"

SYSTEMATIC DESCRIPTIONS

Family SACCAMMINIDAE 

Genus PROTEONINA Wffliamson, 1858

Proteonina sp. 

Plate 83, figure 1

A few specimens occur singly or very rarely in several 

samples at Bikini, Eniwetok, and Rongelap. The test 

is large, elongate, compressed, and smooth on the surface, being composed of large, flat fragments, sometimes 

tests of other Foraminifera, and considerable cement. 

There is a distinct, slender neck. The specimens from

Rongelap are composed of somewhat smaller fragments 

than those from Eiiiwetok. The one fro^Ta Bikini is 

rough on the surface and may be different.

The species does not seem to have been previously 

described, but present material is inadequate for 

description. The species shows a definite selective 

power in choosing large, flat fragments for the construction of its test. In this it may be compared to P. 

micacea Cushman (1918, p. 49, pi. 19, figs. 6, 7) but the 

present species has- a much smoother surface and regular 

shape.

Family HYPERAMMINIDAE 

Genus SAGENINA Chapman, 1900 

Sagenina frondescens (H. B. Brady)

Sagenella frondescens Brady, Quart. Jour. Micros. Sci., v. 19, p.

41, pi. 5, fig. 1, 1879; Challenger Rept., Zoology, v. 9, p. 278,

pi. 28, figs. 14, 15, 1884. 

Sagenina frondescens Chapman, Linnean Soc. Jour., Zoology,

v. 28, p. 4, pi. 1, figs. 1, 2; pi. 2, figs. 1, 2, 18P9. 

Cushman, U..S. Natl. Mus. Bull. 100, v. 4, p. €0, pi. 9, fig. 1;

pi. 14, fig. 1, 1921.

The species is found in abundance attached to 

Halimeda fronds at Bikini 1493 dredged on the outer 

slopes of Bikini at 125 to 150 feet. It also occurs 

attached to large specimens of Cycloclypeus carpenteri 

H. B. Brady obtained by dredging on the outer slopes 

of Bikini from 800 to 580 feet.

Family AMMODISCIDAE 

Genus AMMODISCUS Beuss, 1861 

Ammodiscus incertus (D'Orbigny)

Operculina incerta D'Orbigny, in De la Sagra, His^oire physique,

politique et naturelle de File de Cuba, Foraminiferes, p. 49,

pi. 6, figs. 16, 17, 1839. 

Ammodiscus incertus H. B. Brady, Challenger Repf,., Zoology, v.

9, p. 330, pi. 38, figs. 1-3, 1884. 

Cushman, U. S. Natl. Mus. Bull. 104, pt. 1, p. 9% pi. 39, 1918;

Bull. 100, v. 4, p. 62, pi. 5, figs. 1, 2, 1921.

A single megalospheric specimen was found at Bik 

1176 (5-7").

Family TEXTUIARHDAE 

Genus TEXTULARIA Defrance, 1824 

Textularia agglutinans D'Orbigny 

Plate 83, figure 2

Textularia aggluiinans D'Orbigny, in De la S'agra, Histoire 

physique, politique et naturelle de 1'Ile de Cuba, Foraminiferes, p. 144, pi. 1, figs. 17, 18, 32-34, 1839.

This species was described from shore sa rids of Cuba 

and is a common species in the warmer, shallow waters 

of both the Atlantic and Pacific. The specimens in the 

present material have more chambers and a more rapid 

increase in thickness than the typical form.
RECENT FORAMINIFERA OF THE MARSHALL ISLANDS 329

Textularia candeiana D'Orbigny 

Plate 83, figure 3

Textularia candeiana D'Orbigny, inDe la Sagra, Histoirephysique, 

politique et naturelle de 1'Ile de Cuba, Foraminiferes, p. 143, 

pi. 1, figs. 25-27, 1839.

The types of this species are from the Atlantic in the 

West Indian region but it has been recorded from 

shallow, warm waters of the Pacific and in some 

localities in deeper water.

Comparison between T. candeiana and T. agglutinans shows the following points of distinction: T. 

candeiana is more inflated and globular toward the later 

chambers and has a more tapering form, with very 

indistinct early chambers. T. agglutinans is of more 

even width and thickness nearly throughout and the 

chambers are higher and more distinct in the early 

part.

Textularia conica D'Orbigny

Plate 83, figure 4

Textularia conica D'Orbigny, in De la Sagra, Histoire physique, 

politique et naturelle de 1'Ile de Cuba, Foraminiferes, p. 143, 

pi. 1, figs. 19, 20, 1839.

This is evidently a species of shallow, warm waters. 

It is distinguished by its concave and nearly circular 

apertural face.

Textularia dupla Todd, n. sp. 

Plate 83, figure 6

Test compressed, about as broad as or broader than 

long, triangular in side view, apertural faces flattened 

or slightly concave, except convex in the young stages, 

periphery subacute, irregularly serrate; chambers indistinct, low, broad, much overlapping, later ones 

becoming slightly inflated; sutures indistinct in the 

early part, later becoming depressed, nearly horizontal; 

wall thin, finely granular, somewhat rough on the surface; aperture a low, arched slit at the base of the lastformed chamber bordered by an inconspicuous rim. 

Length 0.42 to 0.60 mm, breadth 0.40 to 0.80 mm, 

thickness 0.25 to 0.45 mm.

Holotype (USNM 548609) from Bikini lagoon, 179 

feet, 11°33'45" N., 165°27'26" E. (Bik 292).

This species resembles T. semialata Cushman but 

differs in its much broader form, its rougher surface, 

and the apertural end depressed instead of inflated. 

It also somewhat resembles T. abbreviata D'Orbigny 

(1846, p. 249, pi. 15, figs. 7-12) but differs in its.more 

numerous, lower, and broader chambers, in the test 

increasing steadily in breadth as growth proceeds, and 

in the very fine texture of its wall.

Textularia foliacea Heron-Alien and Earland 

Plate 83, figure 7

Textularia foliacea Heron-Alien and Earland, Zool. Soc. London 

Trans., v. 20, pt. 2, p. 628, pi. 47, figs. 17-20, 1915.

This is a common species in shallow water of the 

Indo-Pacific area and has been widely recorded in 

coral-reef regions.

Textularia foliacea var. oceanica Cushman 

Plate 83, figure 8

Textularia foliacea Heron-Alien and Earland, var. oceanica Cushman, U. S. Natl. Mus. Bull. 161, pt. 1, p. 8, pi. 1, fi^s. 11, 

12, 1932.

This variety differs from the typical form in being 

thicker and narrower, with higher chambers, and in 

having a rougher surface. It is considerably more 

abundant in the Marshall Islands than the typical form.

Textularia kerimbaensis Said

Plate 83, figure 11

Textularia conica D'Orbigny var. corrugata Heron-Alien and Earland (not T. corrugata Costa), Zool. Soc. London Trans., 

v. 20, p. 629, pi. 47, figs. 24-27, 1915.

Textularia corrugata Cushman, TJ. S. Natl. Mus. Bull. 161, pt. 1, 

p. 12, pi. 3, figs. 2, 4, 1932.

Textularia kerimbaensis Said, Cushman Lab. Foram. Research 

Special Pub. 26, p. 6, pi. 1, fig. 8, 1949.

Rare specimens were found in three Bikini samples. 

The species is distinguished by its broad, low chambers 

and deeply incised sutures.

Textularia milletti Cushman

Textularia milletti Cusbman, TJ. S. Natl. Mus. Bull. 71, pt. 2,

p. 13, text figs. 18, 19, 1911.

Spiroplectammina milletti Cushman, TJ. S. Natl. Mus. Bul\ 161, 

pt. 1, p. 7, pi. 1, fig. 5, 1932.

Test compressed, wedge-shaped, periphery of early 

portion acute in edge view, broadly rounded in side 

view; later portion very slightly tapering and gradually 

increasing in thickness to the truncate apertural end; 

chambers indistinct, low, broad, slightly curved; 

sutures distinct except in some specimens where they 

are obscured by the coarse texture of the wall surface, 

depressed and slightly curved; wall arenaceous, in 

most specimens very roughly finished; aperture a low, 

short opening in a reentrant at the base of the apertural 

face. Length about 1.2 mm, breadth about 0.75 mm, 

thickness about 0.50 mm.

The above description applies to specimens from 

Bikini and Eniwetok that have been compared and 

seem to be identical with types and other specimens 

of this species from deep water (323 to 891 fathoms) 

from several localities in the Pacific.
330 BIKINI AND NEARBY ATOLLS, MARSHALL ISLANDS

In the Marshall Islands, this species is found commonly at four Bikini stations between 592 and 720 

fathoms and at Eni 119 in 500 fathoms.

In the distribution chart this species was not distinguished from Textularia foliacea Heron-Alien and 

Earland, and the two were therefore plotted as one 

species. Actually, the deepest occurrence of T. foliacea 

is 136 fathoms at Bik 814 on the outer slope of the 

atoll.

The two species are, however, distinguishable both 

on the basis of shape of the test and surface of the wall, 

as follows: T. foliacea has a thicker, less wedge-shaped 

test than T. milletti, tapering gradually from the 

initial end, with fewer and higher chambers showing 

indentation along the periphery. T. milletti has a 

much rougher finish than T. foliacea.

The restricted occurrence of T. milletti to deep 

water, as well as that of T. foliacea to shallow water, 

makes these two species of value as ecologic indicators.

Textularia semialata Cushman 

Plate 83, figure 5

Textularia semialata Cushman, II. S. Natl. Mus. Proc., v. 44, 

p. 634, pi. 80, figs. 6, 7, 1913; Bull. 100, v. 4, p. 116, pi. 24, 

figs. 2, 3, 1921; Bull. 161, pt. 1, p. 9, pi. 2, figs. 1-3, 1932.

Test compressed throughout, rapidly broadening, 

triangular in side view, periphery slightly indented, 

siibacute on the earlier chambers, later becoming 

rounded; chambers much overlapping thus appearing 

very narrow on the surface, later chambers lightly inflated; but distinctly granular, very smoothly finished; 

aperture an arched opening at the base of the last 

formed chamber, with a distinct overhanging lip.

This species, originally described from the Philippines 

and known from the Tropical Pacific occurs in several 

samples at Bikini and Rongelap. It is distinctive in 

its overhanging apertural lip and its very thin, smooth 

wall.

Genus SIPHOTEXTULARIA Finlay, 1939

Siphotextularia cf. S. concava (Karrer) 

Plate 83, figure 9

Plecanium concavum Karrer, Akad. Wiss. Wien, Math.-naturwiss. 

Kl., Sitzungsber., Band 58, pt. 1, p. 129, pi. 1, fig. 3, 1868.

A very few specimens from two stations at Bikini 

seem very close to this species described from the 

Miocene of Hungary and widely recorded in the Recent 

seas.

Siphotextularia crispata (H. B. Brady) 

Plate 83, figure 10

Textularia crispata H. B. Brady, Challenger Rept., Zoology, v. 

9, p. 359, pi. 113, fig. 2, 1884.

Test elongate, quadrangular in section with the sides 

slightly, concave, compressed, plane of the test twisted

through about 90°, initial end acute, apertural end 

rounded, periphery truncate with acute and limbate 

angles, periphery serrate in side view, especially along 

the later chambers; chambers rather indistinct, not 

inflated but on the contrary appearing somewhat collapsed; sutures rather distinct, curved, limbate and 

slightly raised; wall very thin, translucent in the middle 

of the chambers, very finely granular, smoothly finished; 

aperture terminal, a narrow slit placed at an angle to 

the suture which separates the apertural faces of the 

two last-formed chambers, surrounded ly a slight, 

raised rim. Length 0.52 to 0.72 mm; breadth 0.30 to 

0.40 mm; thickness 0.17 to 0.25 mm.

Topotypes of Brady's species from 155 fathoms, off 

Raine Island, Torres Strait, indicate this species belongs 

in Siphotextularia.

It differs from S. concava (Karrer)'in its twisted test, 

its less distinctly concave faces, and its very delicate, 

translucent wall.

Genus BIGENERINA D'Orbigny, 182*

Bigenerina sp. 

Plate 83, figure 12

A few specimens of this genus were found in one 

Eniwetok and two Bikini samples, all from deep water. 

They seem to be different from any described species 

but are too rare to warrant description. The textularian portion is relatively broad and rather strongly 

compressed, and the chambers of the uniserial portion 

are indistinct and tend to decrease in diameter with 

growth. The aperture is protuding and surrounded by 

a raised rim.

Family VERNEUIUNIDAE

Genus GAUDRYINA D'Orbigny, 1839

Gaudryina flintii Cushman

Gaudryina subrotundata Flint (not G. subrotundata Sclrwager), 

U. S. Natl. Mus. Rept. for 1897, p. 287, pi. T3, fig. 1, 1899.

Gaudryina rugosa Goes (not G. rugosa D'Orbigny), Mus. Comp. 

Zool. Bull., v. 29, p. 39, 1896.

Gaudryina flintii Cushman, II. S. Natl. Mus. Bull. 71, pt. 2, p. 

63, text figs. 102a-c, 1911; Cushman Lab. Fcram. Research 

Special Pub. 7, p. 62, pi. 10, figs. 18-20, 1937.

This large, deep-water species occurs rarely in the 

guyot samples.

Gaudryina cf. G. pauperata Earland 

Plate 83, figure 13

Gaudryina pauperata Earland, Discovery Repts., v. 10, p. 121, 

pi. 5, figs. 47-49, 1934.

A few specimens occurring rarely in several samples 

from Eniwetok are somewhat similar to this minute 

species described from the Antarctic. They are slightly
RECENT FORAMINIFERA OF THE MARSHALL ISLANDS 331

larger, 0.28 to 0.32 mm in length, are composed of very 

coarse fragments for the size of the test, the axis of the 

test is usually slightly curved, and the last two chambers 

are distinctly inflated.

Gaudryina triangularis var. angulata Cushman 

Plate 83, figure 14

Gaudryina triangularis Cushman var. angulata Cushman, 

Carnegie Inst. Washington Pub. 342, p. 22, 1924; Bernice 

P. Bishop Mus., Bull. 27, p. 125, 1925 (1926); U. S. Natl. 

Mus. Bull. 161, pt. 1, p. 14, pi. 3, figs. 7a, b, 1932; Cushman 

Lab. Foram. Research Special Pub. 7, p. 66, pi. 10, figs. 

14a, b, 1937.

Although specimens exhibit considerable variation in 

shape, the variety is distinguished by its square end 

view and its rough and coarse surface. It occurs at 

Eniwetok, Bikini, and Rongelap.

Gaudryina trullissata Todd, n. sp. 

Plate 83, figure 15

Test small for the genus, tapering, and wedge-shaped 

initial end acute, triserial portion almost indistinguishable in some specimens, apertural end truncate, periphery broadly rounded in end view, irregularly serrate in 

side view; chambers indistinct in the early portion, low 

and broad in the later portion, projecting laterally 

outward; sutures indistinct except toward the periphery 

and in the later portion where they are deeply depressed; 

wall formed of very coarse, ill:sorted fragments, very 

roughly finished; aperture a narrow, low, arched opening at the base of the apertural face of the last-formed 

chamber, with a slight rim. Length 0.50 to 0.60 mm, 

breadth 0.25 to 0.35 mm; thickness 0.20 to 0.25 mm.

Holotype (USNM 548622) from Bikini atoll, 55 

fathoms, ll°34/44 // N., 165°33'40" E. (Bik 815).

This species differs from 0. quadrangularis Bagg 

(1908, p. 133, pi. 5, fig. 1) in its very much smaller 

size, greater compression and less distinct triserial 

portion.

Subgenus SIPHOGAUDRYINA Cushman, 1935

Gaudryina (Siphogaudryina) rugulosa Gushman

Plate 82, figure 1

Gaudryina rugulosa Cushman, U. S. Natl. Mus. Bull. 161, pt. 1,

p. 15, pi. 4, figs, la, b, 1932. 

Gaudryina (Siphogaudryina) rugulosa Cushman, Cushman Lab.

Foram. Research Special Pub. 7, p. 84, pi. 12, figs. 11, 12,

1937. 

Textularia rugosa H. B. Brady (not Reuss), Challenger Rept.,

Zoology, v. 9, p. 363, pi. 42, figs. 23, 24, 1884.

In our specimens the periphery is acute throughout 

and the lobes are very prominent. This species 

seems closely related to Q. (S.) rhodiensis Cushman

(1936, p. 12, pi. 2, figs. 9a, b) from the Pliocene of the 

of the Isle of Rhodes but that species is less broadly 

tapering.

Gaudryina (Siphogaudryina) siphonifera (H. B. Brady) 

Plate 83, figure 17

Textularia siphonifera H. B. Brady, Quart. Jour. Micros. Sci.,

v. 21, p. 53, 1881; Challenger Rept., Zoology, v. 9, p 362,

pi. 42, figs. 25-29, 1884. 

Gaudryina siphonifera Cushman, Cushman Lab. Foram. Research

Contr., v. 4, p. 109, pi. 16, figs. 1-5, 1928. 

Gaudryina (Siphogaudryina) siphonifera Cushman, Cushman

Lab. Foram. Research Special Pub. 7, p. 83, pi. 12, fi-js. 9,

10, 1937.

This species is common in the Rongelap samples, 

but no specimens were found at any of the other isla nds. 

Some specimens are well enough preserved to show the 

closed ends of the tubes.

Gaudryina (Siphogaudryina) transversaria (H. B. Brady) 

Plate 83, figure 16

Textularia transversaria H. B. Brady, Challenger Rept., ZocTogy,

v. 9, p. 359, pi. 113, figs. 3-5, 1884. 

Gaudryina (Siphogaudryina) transversaria Said, Cushman Lab.

Foram. Research Special Pub. 26, p. 8, pi. 1, fig. 12, 1949.

Test small for the genus, strongly compressed, triserial portion very small and inconspicuous, rapidly 

increasing in width in the early part, later part of enual 

width or slightly decreasing, periphery irregularly 

serrate in side view, narrowly truncate in end view due 

to the broken-off peripheral extensions of the chambers; 

chambers numerous, indistinct in the early part, low, 

broad, not inflated; sutures indistinct in the early part, 

later ones horizontal, straight, slightly depressed; wall 

very thin usually translucent in the central part of the 

chambers, very finely granular, very smoothly finished; 

aperture a narrow, low, arched opening at the base of 

the apertural face of the last-formed chamber. Length 

0.35 to 0.55 mm, breadth 0.20 to 0,27 mm, thickness 

0.07 to 0.10 mm.

The species occurs rarely at Bikini and more commonly at Rongelap.

Family VALVULINIDAE

Genus VALVULINA D'Orbigny, 1826

Valvulina davidiana Chapman

Plate 82, figure 2

Valvulina davidiana Chapman, Linnean Soc. Jour., Zoology, v. 

28, p. 9, pi. 1, fig. 4; p. 207 (list), 1900; idem, p. 382, 386 

(lists), 1902.

Cushman, Cushman Lab. Foram. Research Special Pub. 8, 

p. 11, pi. 2, figs. 6, 7, 1937.

Test triangular, chambers triserially arranged and forming a 

more or less equilateral cone; aperture situated in a subcrescentic
332 BIKINI AND NEARBY ATOLLS, MARSHALL ISLANDS

depression, with a tooth-like valve projecting across, and sometimes a secondary one nearly meeting from the opposite side. 

Texture somewhat coarsely arenaceous, composed of calcareous 

particles; colour white to pale cream. Length ^5 inch (1 mm.).— 

Chapman, 1900.

The few records for this species are from shallow 

warm water of the Pacific including the lagoon of 

Rongelap Atoll, Marshall Ids.

Genus CLAVULINA D'Orbigny, 1826

Clavulina angularis D'Orbigny

Plate 83, figure 18

Clavulina angularis D'Orbigny, Annales sci. nat., v. 7, p. 268,

pi. 12, fig. 7, 1826.

Cushman, Cushman Lab. Foram. Research Special Pub. 8, 

p. 19, pi. 2, figs. 29-33, 1937.

Test elongate, sides nearly parallel in the adult, 

earlier portion triserial, later uniserial, triangular in 

transverse section, except for last-formed portion, 

which is rounded; chambers distinct, not inflated except 

the final one or two; sutures distinct, slightly depressed, 

especially toward the apertural end; wall arenaceous, 

rather roughly finished; aperture terminal, rounded, 

with a distinct, valvular tooth.

The species occurs rarely at Eniwetok and Rongelap.

Clavulina paciflca Cushman 

Plate 83, figure 19

Clavulina pacifica Cushman, Carnegie Inst. Washington Pub.

342, p. 22, pi. 6, figs. 7-11, 1924; U. S. Natl. Mus. Bull. 161,

pt. 1, p. 16, pi. 4, figs. 4, 7, 9, 1932. 

Clavulina angularis H. B. Brady (part, not D'Orbigny), Challenger

Rept., Zoology, v. 9, p. 396, pi. 48, figs. 22-24, 1884.

This Indo-Pacific species has a rather wide distribution. One specimen from Rongerik and a single fragment from Bikini seem to belong to this species. The 

texture of the wall is much smoother and the sutures 

not so deeply excavated as in C. angularis D'Orbigny.

Genus KARRERIELLA Cushman, 1933

Karreriella bradyi (Cushman)

Plate 83, figure 20

Oaudryina pupoides H. B. Brady (not D'Orbigny), Challenger 

Rept., Zoology, v. 9, p. 378, pi. 46, figs. 1-4, 1884.

(jfaudryina bradyi Cushman, U. S. Natl. Mus. Bull. 71, pt. 2, p. 

67, figs. 107a-c (in text), 1911.

Karreriella bradyi Cushman, Cushman Lab. Foram. Research 

Special Pub. 8, p. 135, pi. 16, figs. 6-11, 1937.

A few specimens from Bikini seem to belong to this 

-species, which is widely known in Recent seas and as a 

Jossil in ,the late Tertiary.

Genus SCHENCKIELLA Thalmann, 1942 

Schenckiella primaeva (Cushman")

Clavulina primaeva Cushman, U. S. Natl. Mu^. Proc., v. 44,

p. 635, pi. 80, figs. 4, 5, 1913; idem/Bull. 100, v. 4, p. 158,

pi. 30, figs. 2, 3, 5-7, 1921. 

Listerella primaeva Cushman, Cushman Lab. Foram. Research

Contr., v. 9, p. 36, pi. 4, figs. 5a, b, 193?; idem, Special

Pub. 8, p. 153, pi. 17, figs. 24-28, 1937. 

Schenckiella primaeva Cushman, Cushman Lab. Foram. Research

Special Pub. 8A, p. 53, 1947.

This species described from deep water off Borneo 

occurs fairly commonly in the guyot samples.

Family MILIOLIDAE

Genus QUINQUELOCUIINA D'Orbigny, 1826

Quinqueloculina anguina var. arenata Said

Plate 83, figure 21

Miliolina anguina (Terquem) var. agglutinans Wiesner (not 

Quinqueloculina;agglutinans D'Orbigny), in Heron-Alien and 

Earland, Zool. Soc. London Trans., v. 20, p. 575, 1915.

Quinqueloculina anguina Terquem var. agglutinans Cushman, 

Carnegie Inst. Washington Pub. 342, p. 60, pi. 22, figs. 5, 6, 

1924; U. S. Natl. Mus. Bull. 161, pt. 1, p. 18 pi. 5, figs, la-c, 

1932.

Quinqueloculina anguina Terquem var. arenata Said, Cushman 

Lab. Foram. Research Special Pub. 26, p. 9, pi. 1, fig. 25.

From the records this is evidently a common fpecies 

in warm, shallow waters of the Indo-Pacific area. 

Some of our specimens are not very noticeably arenageous, probably because only fine sediment was available for test building.

Quinqueloculina bidentata D'Orbigny 

Plate 83, figure 31

Quinqueloculina bidentata D'Orbigny, in De la Sagra, Histoire 

physique, politique et naturelle de 1'Ile de Cuba, Foraminiferes, p. 197, pi. 12, figs. 18-20, 1839.

This species was described from shore s^nds of Cuba. 

The Pacific specimens seem to be enough l ; ke the Atlantic ones to be included under this name. The apertural 

tooth is not bidentate, as shown in the.type figure, but 

platelike. In our series of specimens there is great 

variation in size and also in the shanoness of the 

peripheral angles. In some specimens th°! periphery is 

almost smoothly rounded.

Some specimens with a short-necked, compressed, 

and flaring aperture are included here but may be 

distinct. They seem to be smaller, thinner-walled, 

more compressed, and have a pebbly surface.

Quinqueloculina bradyana Cushman 

Plate 83, figure 26

Miliolina ^lndosa H. B. Brady (not Quinqueloculina undosa 

Karrer), Challenger Rept., Zoology, v. 9, p. 176, pi. 6, 

figs. 6-8, 1884.

Quinqueloculina bradyana Cushman, U. S. Natl. Mus. Bull. 

71, pt. 6, p. 52, pi. 18, figs. 2a-c, 1917.
RECENT FORAMINIFERA OF THE MARSHALL ISLANDS 333

A very few specimens similar to that figured were 

found at Bikini and Rongelap.

Quinqueloculina crassa var. subcuneata Cushman 

Plate 83, figure 22

Miliolina crassa Heron-Alien and Earland (part, not D'Orbigny), 

ZooL Soc. London Trans., v. 20, pt. 2, p. 572, pi. 42, fig. 41 

(not figs. 37-40), 1915.

Quinqueloculina crassa D'Orbigny var. subcuneata Cushman, 

U. S. Natl. Mus. Bull. 100, v. 4, p. 423, pi. 89, figs. 4a-c, 

1921; idem, Bull. 161, pt. 1, p. 21, pi. 5, figs. 8a-c, 1932.

Numerous specimens in our material seem identical 

with this variety which is widely distributed, particularly in the Indo-Pacific region. There is considerable 

variation in strength of the ornamentation. The wall 

is more highly polished than in Q. neostriatula Thalmann 

and the costae are raised. In Q. neostriatula the wall is 

finely roughened, although shining, and the ornamentation consists of fine, incised striae.

Quinqueloculina distorqueata Cushman, n. sp. 

Plate 83, figure 27

Test nearly as broad as long, initial end bluntly 

pointed, apertural end with a short, slender neck, 

periphery acute or slightly keeled, irregularly curved; 

chambers much compressed, distinct, enlarging rather 

rapidly as added, distorted in shape; sutures indistinct 

except for the keel of the preceding chamber; wall 

slightly roughened, with the sinuous keel of previous 

chambers making an irregular raised pattern; aperture 

small, rounded, at the end of a short and very slender 

neck, with a slight lip, usually without a definite 

apertural tooth. Length 0.65 to 0.80 mm, breadth 

0.45 to 0.65 mm.

Holotype, USNM 548545, from Rongerik lagoon, 

150 feet, 11°22'36" N., 167°26'55" E. (Rik 79).

This species is very common in certain samples from 

all the islands. They somewhat resemble forms referred 

to Quinqueloculina berthelotiana D'Orbigny (Cushman, 

1932, p. 19, pi. 5, figs. 2-4), but have a much more 

irregular form of the chambers which are sub-acute or 

keeled. This species differs from Q. samoaensis Cushman in its higher, sharper keels which make the test 

appear more twisted.

Quinqueloculina cf. Q. ferussacii D'Orbigny 

Plate 83, figure 30

A few specimens of moderate size and closely approaching the triloculine stage are questionably assigned 

to this species. They are elongate, with a moderately 

prominent neck, but without a lip, compressed, and are 

rather irregularly and variably ornamented with two 

or more costae on the periphery. The surface, of the 

wall is matte rather than polished.

This form is found only at Bikini and Eniwetok.

Quinqueloculina neostriatula Thalmann 

Plate 83, figure 28

Quinqueloculina striatula Cushman (not Deshayes, 1831) U. S.

Natl. Mus. Bull. 161, pt. 1, p. 27, pi. 7, figs. 3, 4, 193?. 

Quinqueloculina neostriatula Thalmann, Cushman Found. Foram.

Research Contr., v. 1, p. 45, 1950.

The types of this species are from Mokaujar Anchorage, Fiji Islands, and it seems to be widely distributed 

in shallow, warm waters of the South Pacific.

The ornamentation consists of fine incised lines, not 

costae, and the angles of the chambers are sharp. 

The aperture is a hood-like opening, not terminal and 

circular with a neck, although some specimens do have 

a neck.

Quinqueloculina parkeri (H. B. Brady) 

Plate 83, figure 23

"Qmnaueloculina with oblique ridges" Parker, Trans. Micros.

Soc. London, v. 6, p. 57, pi. 5, fig. 10, 1858. 

Miliolina parkeri H. B. Brady, Quart. Jour. Micros. Sci., v. 21,

p. 46, 1881; Challenger Kept., Zoology, v. 9, p. 177, pi. 7,

fig. 14, 1884. 

QuniQueloculina parkeri Cushman, TJ. S. Natl. Mus. Bul1 . 71,

pt. 6, p. 50, pi. 15, fig. 3, 1917; idem, Bull. 161, pt. 1, p. 25,

pi. 6, figs. 3, 4, 1932.

This is a species of warm, shallow waters of the IndoPacific area. It is found rarely at all the atolls.

Quinqueloculina parkeri (H. B. Brady) var. 

Plate 83, figure 24

A few specimens from all the islands differ from the 

typical form in the ornamentation being less distinct 

and consisting of fewer, coarser ridges, usually present 

only on the central part of the test.

Quinqueloculina cf. Q. samoaensis Cushman 

Plate 83, figure 25

Quinaueloculina samoaensis Cushman, Carnegie Inst. Washington Pub. 342, p. 59, pi. 21, figs. 4-7, 1924; U. S. Natl. Mus. 

Bull. 161, pt. 1, p. 26, pi. 7, figs, la-c, 1932.

A few specimens from Eniwetok seem close to this 

species described from 17 fathoms at Samoa. They 

are slightly smaller and most of them do not have as 

elongate an apertural neck.

Quinqueloculina cf. Q. seminulum (Linne) 

Plate 83, figure 29

A very few small specimens from all the islands 

similar to that figured, are referred to this widely recorded species. The wall is smooth and polished. 

They seem to be too large to be the nepionic or quinqueloculine stage of Triloculina cf. oblonga, (Montagu), 

although otherwise they resemble that species.
334 BIKINI AND NEAEBY ATOLLS, MAESHALL ISLANDS

Quinqueloculina sulcata D'Orbigny 

Plate 84, figures 1, 2

Quinqueloculina sulcata D'Orbigny, Annales sci. nat., v. 7, p. 301,

no. 17, 1826.

Cushman, U. S. Natl. Mus. Bull. 161, pt. 1, p. 28, pi. 7, figs. 

5-8, 1932.

The types of this species are from the Red Sea and it 

is recorded from a number of localities in warm, shallow 

waters about certain Pacific Islands.

The texture of the wall and the height and straightness of the ridges vary considerably. A group of 

specimens from Eniwetok are much more flattened than 

the more typical forms, but for the present are included 

in this species.

Genus MILIOLINELLA Wiesner, 1931

Miliolinella australis (Parr)

Plate 84, figures 3, 4

Quinqueloculina australis Parr, Royal Soc. Victoria Proc., v. 44, 

p. 7, pi. 1, fig. 8, 1932.

Test small for the genus, nearly circular, compressed, 

periphery subacute; chambers strongly curved, not inflated; sutures indistinct, curved, slightly depressed; 

wall thin, translucent in the middle portions of the 

chambers, surface very finely marked with faint, 

incised, longitudinal lines; aperture a high, arched 

opening with a slight lip, nearly closed by a projecting 

plate in front of the opening. Length and breadth both 

about 0.30 mm; thickness 0.15 to 0.20 mm.

This species was described from 7 miles east of Cape 

Pillar, Tasmania, in 100 fathoms and has been recorded 

from other localities off Australia.

A few specimens are considerably larger than the 

average but these appear to be the same, as they have 

the typical wall texture which is a good distinguishing 

feature of this species. The projecting plate in front 

of the aperture is frequently broken away but when 

present, the test shows in side view a rather deep slot 

between the aperture and the plate.

Miliolinella oceanica (Cushman)

Triloculina oceanica Cushman, U. S. Natl. Mus. Bull. 161, pt. 1> 

p. 54, pi. 12, figs. 3a-c, 1932.

A few typical specimens of this species which apparently belongs in MUiolinetta were found in two Bikini 

samples.

Miliolinella oceanica (Cushman), var. flintiana (Cushman)

Triloculina oceanica Cushman, var. flinliana Cushman, U. S. 

Natl. Mus. Bull. 161, pt. 1, p. 55, pi. 12, figs. 4a-c, 1932.

A single specimen was found in Rongerik sample 54.

Genus TRILOCTJLINELLA Riccio, 1950

Triloculinella labiosa (D'Orbigny)

Plate 84, figures 5, 6

Triloculina labiosa D'Orbigny, in De la Sagra Hhtoire physique1

politique et naturelle de 1'Ile de Cuba, Foraminiferes, p. 178»

pi. 10, figs. 12-14, 1839. 

Cushman, U. S. Natl. Mus. Bull. 161, pt. 1, p. 53, pi. 11, figs.

12a-c, 1932. 

Miliolina labiosa H. B. Brady, Challenger Rept., Zoology, v. 9,

p. 170, pi. 6, figs. 3-5, 1884.

Numerous specimens are found in most of the samples. This genus bears the same relationship to 

MiliolineUa as Triloculina, does to Quinqueloculina,.

Genus MASSILINA Schlumberger, 1893

Massilina planata Cushman

Plate 84, figure 16

Massilina planata Cushman, U. S. Natl. Mus. Bull. 161, pt. 1, 

p. 31, pi. 8, figs. 8a, b, 1932.

A few typical specimens were found at Eniwetok, 

Bikini, and Rongelap.

Genus SPIROLOCULINA D'Orbigny, 1826 

Spiroloculina acescata Cushman

Spiroloculina grateloupi D'Orbignj, var. acescata Cushman^

U. S. Natl. Mus. Bull. 161, pt. 1, p. 35, pi. 9, fig. 2, 1932. 

Spiroloculina acescata Cushman and Todd, Cushman Lab.

Foram. Research Special Pub. 11, p. 58, pi- 8, figs. 11, 12,.

194i. 

Spiroloculina planissima Heron-Alien and Earland (not

Lamarck), Zool. Soc. London Trans., v. 20, p. 556^

pi. 41, figs. 1-5, 1915.

A single specimen of this species known from the Fiji 

Islands was found at Eniwetok 310.

Spiroloculina aequa Cushman

Spiroloculina antillarum D'Orbigny, var. aequa Cushman, U. S.

Natl. Mus. Bull. 161, pt. 1, p. 38, pi. 9, fig. 13, 1932. 

Spiroloculina aequa Cushman and Todd, Cushmin Lab. Foranu

Research Special Pub. 11, p. 59, pi. 8, figs. 13-15, 1944.

A few typical specimens of this species described from 

inside the lagoon, Pinaki Atoll, Paumotu Ids., were 

found at Rongelap 111.

Spiroloculina angulata Cushman 

Plate 84, figures 11, 12

Spiioloculina grata Terquem, var. angulata Cushman, U. S. NatL

Mus. Bull. 71, pt. 6, p. 36, pi. 7, fig. 5, 1917. 

Spiroloculina antillarum D'Orbigny, var. angulata Cushmaa

(part), U. S. Natl. Mus. Bull. 100, v. 4, p. 408, pi. 81, fig. 5^

1921; idem, Bull. 161, pt. 1, p. 37, pi. 9, figs 6-9, 1932. 

Spiroloculina angulata Cushman and Todd, Cushman Lab.

Foram. Research Special Pub. 11, p. 50, pi. 7, figs. 18-22,,

1944. 

Spiroloculina grata H. B. Brady (not Terquem), Challenger Rept.,.

Zoology, v. 9, p. 155, pi. 10, figs. 16, 17, 22, 23, 1884.

The types of this species are from off Cebu, Philippine-
RECENT FORAMINIFERA OF THE MARSHALL ISLANDS 335

Islands. It is widely distributed in shallow, warm 

water of the Indo-Pacific.

The larger forms normally have a prominent ridge 

along the center of the periphery while the smaller ones 

have a flat and truncate periphery, and usually less distinct ornamentation. Both forms are figured. We believe the larger forms are microspheric and the much 

more abundant smaller ones are megalospheric.

Spiroloculina circularis Chapman 

Plate 84, figure 7

Spiroloculina dorsata Reuss, var. circularis Chapman, Biol.

Results Endeavour, v. 3, pt. 1, p. 7, pi. 1, fig. 1, 191E. 

Spiroloculina circularis Cushman and Todd, Cushman Lab.

Foram. Research Special Pub. 11, p. 49, pi. 7, figs. 15, 16,

1944.

A very few specimens of this species were found in 

three of the deep Bikini samples. It is a very small 

form, nearly circular in outline except for the basal and 

apertural ends of the last chamber which are protruding. 

The test is composed of very few chambers, rapidly increasing in size and thickness at the periphery which is 

truncate and bluntly keeled. The apertural neck is 

much contracted and has a slight phialine lip.

Spiroloculina clara Cushman 

Plate 84, figure 9

Spiroloculina clara Cushman, U. S. Natl. Mus. Bull. 161, pt. 1, 

p. 40, pi. 10, figs. 4, 5, 1932.

The types of this species were from Rongelap Atoll, 

and it has been found at other localities in the Pacific. 

It is the most abundant and widespread species of this 

genus in the Marshall Islands.

Spiroloculina clara var. lirata Cushman, n. var. 

Plate 84, figure 10

Variety differing from the typical form in the adult 

having an expanded periphery with raised ridges and 

the entire surface of the chambers more or less costate 

longitudinally.

Holotype, USNM 548477, from Rongerik lagoon, 102 

feet, 11°21'36" N., 167°25'44" E. (Rik 31).

Spiroloculina com munis Cushman and Todd 

Plate 84, figure 13

Spiroloculina communis Cushman and Todd, Cushman Lab.

Foram. Research Special Pub. 11, p. 63, pi. 9, figs. 4, 5, 7,

8, 1944. 

Spiroloculina excavata H. B. Brady (not D'Orbigny), Challenger

Rept., Zoology, v. 9, p. 151, pi. 9, figs. 5, 6, 1884. 

Spiroloculina impressa H. B. Brady (not Terquem), idem, p. 151,

pi. 10, figs. 3, 4, 1884. 

Spiroloculina grateloupi Cushman (not D'Orbigny), U. S. Natl.

Mus. Bull. 161, pt. 1, p. 34, pi. 8, figs. 10, 11 (not fig. 9), 1932.

A few specimens occur at Eniwetok, Bikini, and

Rongelap. They are rather rough-surfaced and not 

as large as the types.

Spiroloculina corrugata Cushman and Todd

Plate 84, figures 17, 18

Spiroloculina corrugatu Cushman and Todd, Cushman Lab.

Foram. Research Special Pub. 11, p. 61, pi. 8, figs. 22-25,

1944. 

Spiroloculina antillarum Cushman (not D'Orbigny) (part), IT. S.

Natl. Mus. Bull. 161, pt. 1, p. 36 (not pi. 9, figs. 3-5), lt)32.

The types of this species are from the Philippine 

region, 50 fathoms, off San Andreas Island, between 

Marinduque and Luzon. It is a variable species and 

widely distributed in the Indo-Pacific area.

In our specimens there is a variation from forms with 

cylindrical chambers and a very narrow test, to those 

with a broader test having flattened chambers, fainter 

costae, and a translucent wall. Both forms are figured.

Spiroloculina foveolata Egger 

Plate 84, figure 14

Spiroloculina foveolata Egger, K. bayer Akad. Wiss., Meth.- 

naturh. Abt., Abh., Kl. 2, Band 18, p. 224, pi. 1, figs. 

33, 34, 1893.

Cushman and Todd, Cushman Lab. Foram. Research Special 

Pub. 11, p. 48, pi. 7, figs. 7-12, 1944.

The types of this species are from off Mauritius and 

it seems to have a wide range in the Indo-Pacific 

region. Rare specimens were found at Rongerik and 

Rongelap.

Spiroloculina marshallana Todd, n. sp. 

Plate 84, figure 8

Test compressed throughout, slightly longer than 

broad, only slightly depressed in the middle, periphery 

squarely truncate or slightly convex or concave with 

thickened edges; chambers numerous, distinct, quadrangular in section, of uniform width, increasing slowly 

in size in the early stages, later more rapidly,,, basal and 

apertural ends about equally protruding; sutures distinct, slightly depressed; wall smooth, polished, somewhat translucent along the middle of the chambers; 

aperture small, quadrangular, nearly filled by a tooth. 

Length 0.75 to 0.92 mm; breadth 0.50 to 0.72 mm; 

thickness 0.10 to 0.13 mm.

Holotype, USNM 548615, from Bikini atoll, 136 

fathoms, 11°29'18" N., 165°30'59" E. (Bik 814).

This species differs from S. clara Cushman in its 

larger size and more nearly circular shape and its less 

distinctly translucent wall. It differs from S. reguUiris 

Cushman and Todd (1944, p. 51, pi. 7, figs. 26, 27) in 

its flatter form and protruding apical and apertural 

ends.

Specimens were found at all four islands but they 

were fairly common only at Bikini.
336 BIKINI AND NEARBY ATOLLS, MARSHALL ISLANDS

Spiroloculina mayori Cushman 

Plate 84, figure 15

Spiroloculina mayori Cushman, Carnegie Inst. Washington Pub.

342, p. 56, pi. 20, figs. 5, 6, 1924.

Cushman and Todd, Cushman Lab. Foram. Research Special 

Pub. 11, p. 56, pi. 8, figs. 1-6, 1944.

The types of this species are from 25 fathoms, off 

Utilei Reef, Pago Pago harbor, Samoa. The species is 

apparently limited to shallow, warm waters of the 

Pacific. A few young specimens were found at Eniwetok and Bikini.

Spiroloculina sp. A 

Plate 84, figures 21, 22

A very few specimens of a distinctive but apparently 

undescribed form occur in several Rongelap samples 

and single specimens were found at Rongerik 51 and 

Eniwetok 119. They are characterized by their broad, 

keel-shaped chambers which are very strongly embracing leaving only a relatively small depressed area in 

the middle of the test, and by their elongate and much 

contracted apertural neck. The wall is thick, procellaneous and polished; in some specimens, apparently 

immature forms, the periphery is truncate rather than 

acute, and several specimens, one of which is figured, 

show the one type of periphery on one side and the 

other type on the other side. In having this broad 

truncate periphery in the young form the species resembles S. mayori Cushman. Adult forms are somewhat similar to S. cushmani Hada (1931, p. 83, text 

fig. 36) but are more strongly compressed, and have an 

elongate apertural neck while S. cushmani has none.

Spiroloculina sp. B

Plate 84, figure 19

The single specimen figured seems to represent an 

undescribed species. It is unique in its slender and 

tubular apertural neck.

Genus SIGMOIIINA Schlumberger, 1887

Sigmoilina edwardsi (Schlumberger)

Plate 84, figure 20

Planispirina (Sigmoilina) edwardsi Schluiuberger, Soc. zool.

France Bull., v. 12, p. 483 (113), pi. 7, figs. 15-18; text fig. 8,

1887. 

Sigmoilina edwardsi Heron-Alien and Earland, Zool. Soc. London

Trans., v. 20, p. 584, pi. 45, figs. 19-21, 1915.

A very few rather small specimens were found in 

two Rongelap samples.

Genus ARTICULINA D'Orbigny, 18?«J 

Articulina elongata Cushman

Articulina elongata Cushman, Cushman Lab. Foram. Research 

Special Pub. 10, p. 16, pi. 4, figs. 7-9, 1944.

A fragment of a specimen showing a series of six 

uniserial chambers with the later chambers characteristically overhanging the earlier ones, was found at 

Rongelap 468. A few fragments were found at 

Eniwetok.

Articulina pacifica Cushman

Plate 84, figure 25

Articulina pacifica Cushman, Cushman Lab. Foram. Research 

Special Pub. 10, p. 17, pi. 4, figs. 14-18, 1944.

The types of this species are from Mokaujar Anchor 

age, Fiji. It is evidently a species of the warmer waters 

of the Indo-Pacific and occurs fairly commonly at all 

the atolls. In the second, or possibly third, uniserial 

chamber, the chamber is broad and flattened and the 

lip much recurved, suggesting A. mucronata (D'Orbigny) 

(1839a, p. 52, pi. 7, figs. 16-19).

Genus TUBINELLA Rhumbler, 190? 

• Tubinella funalis (H. B. Brady) 

Plate 84, figure 24

Articulina funalis H. B. Brady, Challenger Rept., Zoology, v. 9,

p. 185, pi. 13, figs. 6-11, 1884. 

Tubinella funalis Rhumbler, Jahrb. Zool., Ab... Syst., Band 24, p.

26, pi. 2, fig. 3, 1906; Foram. Plankton-Exped., pt. 1, pi. 2,

figs. 22, 23, 1911; pt. 2, p. 384, 1913.

The records indicate that this is a very widely distiibuted species. Only a single specimen was found at 

Rongerik 51.

Genus HAUEBINA D'Orbigny, 183?

Hauerina bradyi Cushman

Plate 84, figure 32

Hauerina bradyi Cushman, U. S. Natl. Mus. Pull. 71, pt. 6, 

p. 62 (not pi. 23, fig. 2), 1917; Cushmar Lab. Foram. 

Research Contr., v. 22, p. 11, pi. 2, figs. 14, 20, 21, 1946.

Hauerina cotnpressa H. B. Brady (not D'Orbigny), Challenger 

Rept., Zoology, v. 9, p. 190, pi. 11, figs. 12, 13, 1884.

The species is characterized by having a definite, 

elongated quinqueloculine stage in the middle, then a 

spiroloculine stage for several whorls, and in the adult 

having about 2% chambers per whorl. The wall, 

especially of the later chambers, is usually translucent 

and bluish. The test is almost completely evolute.

This species apparently has a fairly wide range in the 

Indo-Pacific region. It is much less common in the 

Marshall Islands than H. divfirsa Cushman.
KECENT FORAMINIFERA OF THE MARSHALL ISLANDS 337

Hauerina diversa Cushman 

Plate 84, figures 30, 31

Hauerina diversa Cushman, Cushman Lab. Foram. Research 

Contr., v. 22, pt. 1, p. 11, pi. 2, figs. 16-19, 1946.

Hauerina bradyi Cushman (not Cushman, 1917), U. S. Natl. 

Mus. Bull. 161, pt. 1, p. 44, pi. 10, figs. 12-15, 1932.

The types of this species are from shallow water off 

the beach at Hereheretue, Paumotu Islands, and it is 

also recorded from 12 fathoms, Levuka, Fiji, and in 18 

fathoms, Vavau Anchorage, Tonga Islands. It occurs 

commonly at all the atolls of the Marshall Islands.

This species differs from H. bradyi Cushman in being 

thicker and in having 3 to 5 chambers in the adult 

whorl; an inconspicuous and not raised, early quinqueloculine stage; and the test strongly involute with 

the last chambers consequently broad. The wall is 

usually not translucent, but may be. The earlier 

apertures are apparent as markings on the wall like 

retral processes along the sutures. The test tends to 

become uncoiled (see fig. 31) thus suggesting the 

Eocene genus Raadshocvenia van denBold (1946, p. 123) 

bu t these do not have labyrinthic chambers.

Hauerina involuta Cushman 

Plate 84, figures 28, 29

Hauerina involuta Cushman, Cushman Lab. Foram. Research

Contr., v. 22, p. 13, pi. 2, figs. 25-28, 1946. 

Hauerina ornatissima H. B. Brady (part) (not Karrer), Challenger

Rept., Zoology, v. 9, p. 192, pi. 7, figs. 15-17 (not figs. 18-22),

1884. 

Cushman (part), U. S. Natl. Mus. Bull. 161, pt. 1, p. 43 (not

pi. 10, figs. 16, 17), 1932.

The types of this species are from Rongelap Atoll, 

and it is evidently a common species in warm, shalloAv 

waters of the Pacific.

A very few small specimens from Eniwetok and 

Rongelap included here are much compressed and not 

involute, and somewhat resemble H. speciosa (Karrer) 

(1868, p. 135, pi. 1, fig. 8) from the Miocene of Europe. 

They are probably different but are inadequate for 

further identification.

Hauerina milletti Cushman, n. sp. 

Plate 84, figure 23

Miliolina circularis Bornemann var. sublineata Millett (not- H. 

B. Brady) Royal Micros. Soc. Jour., p. 501, pi. 11, figs. 4a, 

b, 1898.

Test in the adult stage rounded in front view and 

broadly elliptical in edge view, periphery broadly 

rounded; chambers distinct, strongly inflated, in the 

earlier stages with several making up the surface but hi 

the adult becoming involute and usually but three 

making up the entire surface; sutures distinct, depressed; wall thin, translucent, with very fine, numerous longitudinal costae; aperture with a finely perforate,

cribrate plate. Length 0.45 to 0.50 mm, diameter 0.40 

to 0.45 mm, thickness 0.25 to 0.28 mm.

Holotype, USNM 548501, from Rongerik lagoon, 

111 feet, 11°22'32" N., 167°28'48" E. (Rik 51).

This is the same as the form recorded and figured by 

Millett in the above reference from the Malay Archipelago. It differs from Brady's variety in the more 

irregular shape, and the cribrate aperture places it in 

the genus Hauerina.

Hauerina serrata Cushman, n. sp. 

Plate 84, figures 26, 27

Test much compressed, early quinqueloculine ,stage 

with strongly raised chambers, in the adult with the 

periphery very prominently wavy and serrate; chambers distinct, compressed in the adult, two forming the 

entire periphery, basal end projecting into a triangular 

lobe; sutures distinct, depressed; wall slightly granular; 

aperture slightly projecting, cribrate. Length 0.60 to 

0.70 mm, breadth 0.45 to 0.55 mm. ''

Holotype, USNM 548764, from Eniwetok lagcon, 

26 fathoms, 11°30'16" N., 162°19'35" E. (Eni 321),

This species differs from H. pacifica Cushman in the 

more regular form with two chambers making up the 

adult whorl and in the coarsely serrate periphery in 

the adult. Specimens occur at a number of stations 

in the vicinity of Eniwetok Atoll and also, but less 

commonly, at the other three islands.

Genus SCHLUMBERGERINA Munier-Chalmas, 1882

Schlumbergerina alveoliniformis (H. B. Brady)

Plate 85, figure 1

Miliolina alveoliniformis H. B. Brady, Quart. Jour. Micros, 

Sci., vol. 19, p. 54, 1879; Challenger Rept., Zoology, v. 9, 

p. 181, pi. 8, figs. 15-20, 1884.

Schlumbergerina alveoliniformis Cushman, U. S. Natl. Mus. 

Bull. 161, pt. 1, p. 29, pi. 8, fig. 1, 1932.

A few specimens from a number of Rongelap (samples 

seem to belong to this species. They are somewhat 

shorter and broader than Brady's type figures and 

the young forms show considerable variation in shr,pe. 

Most of the specimens appear worn and the cribrate 

apertural plate is absent in many of them. Rare 

specimens also occur at the other atolls.

Genus TRILOCULINA D'Orbigny, 1826

Triloculina cf. T. bassensis Parr

Plate 85, figure 14

Triloculina bassensis Parr, Royal Soc. Victoria Proc., v. 56 

(n. ser.), pt. 2, p. 198, pi. 8, figs. 7a-c, 1945.

A few specimens from all the atolls seem close ;to this 

species described from shore sands at Barwon Hep.ds, 

Victoria, Australia. They were compared with a tcpotype received from Mr. Parr and some differ in having
338 BIKINI AND NEARBY ATOLLS, MARSHALL ISLANDS

a slightly rougher surface but otherwise seem very 

close. They are characterized by the elongate arched 

aperture which is slightly grooved at the outer end, 

and by the wall covered by a pattern of very fine, 

incised longitudinal lines. In a few cases the wall is 

slightly crinkled transversely.

Triloculina cf. T. bicarinata D'Orbigny 

Plate 85, figure 2

Triloculina bicarinata D'Orbigny, 77? De la Sagra, Histoire 

physique, politique et naturelle de 1'Ile de Cuba, Foraminiferes, p. 180, pi. 10, figs. 18-20, 1839.

Cushman, U. S. Natl. Mus. Bull. 161, pt. 1, p. 60, pi. 13, figs. 

6a-c, 1932.

The figured specimen is similar to that figured from 

the Pacific, Mokaujar Anchorage, Fiji, but differs in 

not having the sharply angled periphery. It is probably distinct from the Atlantic species. The ornamentation is of a coarser texture in the Pacific specimens and tends to include elongate and irregular ridges.

Triloculina bikiniensis Todd, n. sp. 

Plate 85, figure 19

Test only slightly longer than broad, apical end 

rounded, apertural end not projecting, periphery acute; 

chambers curved, not inflated; sutures indistinct, not 

depressed; wall smooth, highly polished, somewhat 

translucent; aperture large, without a neck or lip, 

with an elongate tooth, slightly broadened or bifid at 

the tip. Length 0.40 to 0.48 mm, breadth 0.30 to 

0.37 mm.

Holotype, USNM 548598, from Bikini lagoon, 119 

feet, 11°36'37" N., 165°30'34" E. (Bik 177).

This species seems to be distinct from any known 

species but is very close to Quinoueloculina auberiana 

D'Orbigny (1839, p. 193, pi. 12, figs. 1-3) described 

from the West Indies, differing from that species (which 

generically is very close to Triloculina) in its less 

sharply acute periphery and its different apertural tooth.

It occurs at Bikini, Rongelap and Rongerik.

Triloculina earlandi Cushman, n. sp. 

Plate 85, figure 3

Miliolina terouemiana Heron-Alien and Earland (part) (not H. B. 

Brady), Zool. Soc. London Trans., v. 20, p. 563, pi. 41, 

figs. 26-28 (not figs. 29-31), 1915.

Test elongate, greatest breadth usually below the 

middle, triangular in end view, initial end rounded, 

apertural end tapering, periphery of the last-formed 

chamber broad, slightly convex and with distinct angles; chambers distinct, three visible in the adult, the 

last formed one making up a large proportion of the 

surface; sutures distinct, slightly depressed, the lastformed one usually slightly sinuous; wall ornamented 

by very fine longitudinal costae, often very slightly

oblique; aperture circular, at the end of r, slight neck, 

with a small, often slightly bifid tooth. Length 0.45 

to 0.50 mm, breadth 0.20 to 0.25 mm.

Holotype, USNM 548502, from Rongerik lagoon, 111 

feet, 11°22'32" N., 167°28'48" E. (Rik 51).

This abundant species in this material is apparently 

identical with the form noted above from the Kerimba 

Archipelago. It differs from Triloculino, terquemiana 

(H. B. Brady) in the more elongate form, slightly 

twisted chambers, distinct neck and rounded aperture 

with a bifid tooth.

Triloculina involuta Todd, n. sp. 

Plate 85, figure 4

Test of medium size for the genus, about twice as 

long as broad, compressed, nearly involute so that the 

last two chambers comprise most of the surface, basal 

end projecting and rounded, apertural end projecting in 

a short, thick, cylindrical neck, periphery broadly 

rounded; chambers curved, slightly inflated and strongly 

embracing the previous ones, very rapidly increasing in 

size as added; sutures indistinct, slightly depressed, 

either straight or irregular due to irregular overlapping 

of last chamber; wall dull-surfaced, ornamented by 

numerous, closely spaced costae, variable in strength, 

intercalating and tending to be slightly tangential, high 

and sharp when well preserved, best developed on the 

peripheral part of each chamber, extending onto the 

neck and curving around the basal end; aperture a 

large, circular, slightly flaring opening surrounded by a 

slightly thickened and polished rim, with a short bifid 

tooth. Length 1.10 to 1.50 mm, breadth 0.60 to 0.90 

mm, thickness 0.37 to 0.62 mm.

Holotype, USNM 548742, from Eniwetok atoU, 24 

fathoms, 11°20'06" N., 162°16'19" E. (Eni 118).

This species differs from T. linneiani D'Orbigny 

(1839a, p. 172, pi. 9, figs. 11-13) in its ornamentation, 

the costae being more closely spaced and tass regular in 

pattern, and in its more strongly embracing chambers 

resulting in a nearly involute form. It aTso* resembles 

T. planciana D'Orbigny (1839a, p. 173, pi. 9, figs. 17-19) 

but is much more strongly ornamented.

The generic position of adult forms of this species 

seems to be transitional between Triloculina and Flintia 

but as the young clearly show the triloculine, not 

biloculine, stage, the species is placed in Triloculina.

Triloculina irregularis (D'Orbigny) 

Plate 85, figure 12

Quinqueloculina irregularis D'Orbigny, Annales sci. nat., tome

7, p. 302, no. 25, 1826. 

Fornasini, Accad. sci. 1st. Bologna Mem., ser. 6, v. 2, p. 67,

pi. 3, fig. 14, 1905. 

Triloculina irregularis Cushman, U. S. Natl. Mus. Bull. 161,

pt. 1, p. 54, pi. 12, figs. 2a-c, 1932.
TABLE I.—Distribution of Saccamminidae, Hyperamminidae, Ammodiscidae, Textulariidae, Verneuilinidae, Valvulinidae, and Miliolidae in the Marshall Islands.

Family Saccamminidae

Proteonina BO.

Family Hyperamminidae

Sagenina frondescens

Family Ammodiscidae

Ammodiscus incertus

Family Textulariidae

Textularia aeglutinans

candeiana

conica

dupla n. sp.

foliacea

foliacea var. oceanica

kerimbaensis

semialata

Siphotextularia cf. S. concava

crispata

Bigenerina sp.

Family Verneuilinidae

Gaudryina flintii

cf. G. jjauperata

triangularis var. ansulata

trullissata n. sp.

Gaudryina (Siphogaudryina) rugulosa

transversaria

siphonifera

Family Valvulinidae

Valvulina davidiana

Clavulina angularis

pacifica

Karreriella bradyi

Schenckiella primaeva

Family Miliolidae

Quinqueloculina anguina var. arenata

bidentata

bradyana

crassa var. subcuneata

distorqueata n. sp.

cf. Q. ferussacii

neostriatula

parkeri

parkeri var.

cf. Q. samoaensis

cf. Q. seminula

sulcata

Miliolinella oceanica

oceanica var. flintiana

australis

Triloculinella labiosa

Massilina planata

Spiroloculina acescata

aequa

angulata

circularis

clara

clara var. lirata n. var.

communis

corrugata

foveolata

marshallana n. sp.

mayor!

sp. A

sp. B

Sigmoilina edwardsi

Articulina elongata

pacifica

Tubinella funalis

Hauerina bradyi

diversa

involuta

milletti n. sp.

serrata n. sp.

Schlumbergerina alveoliniformis

RONGERIK LAGOON

0) CM

0

0 COin

•* S

« S S S CM o 

rr- 00 05r1

1

1

1

1

2

15

5

1

4

1

13

4

6

7

3

13

3

10

14

2

1

4

3

6

14

4

8

13

14

3

12

1

2

7

1

7

1

5

4

6

5

1

16

15

8

4

6

2

7

1

11

2

5

7

3

8

3

1

4

2

1

2

1

6

1

21

1

1

4

1

2

1

4

1

1

1

19

1

1

5

1

16

1

3

10

1

3

2

2

6

1

1

2

4

4

2

4

1

2

1

18

15

6

1

4

1

1

15

5

8

11

*

22

5

g

\

4

1

4

1

6

1

7

6

1

5

1

6

16

1

15

5

10

12

8

14

1

10

1

11

2

12

6

1

3

17

9

12

2

4

2

1

12

10

1

2

1

7

1

2

23

17

8

4

1

1

4

8

3

4

11

2

7

7

6

12

3

1

4

1

1

24

1

11

1

10

3

1

6

21

2

6

7

10

3

2

1

5

1

1

6

3

2

2

7

3

12

lg

15

2

17

7

8

7

3

5

11

5

5

1

1

6

7

6

4

3

2

10

17

1

2

4

16

21

16

22

J

2

5

3

1

17

RONGELAP ATOLL

Inside the lagoon

* CO ^

a

00•* g

00•*

CO

•* 

•* r- o

CO•*

CO

r- oc

CO CM I- o

CO

Hill I2|l|

S

6

4

3

12

7

5

10

8

7

J

20

4

6

12

1

3

1

1

7

1

3

1

2

5

19

t

5

5

t

4

6

7

5

18

4

6

2

6

20

17

9

9

S

5

2

2

3

0

6

2

2

1

1

1

1

7

1

1

15

1

18

1

6

1

1

20

9

20

5

5

20

2

1

4

5

14

2

g

3

3

2

2

11

4

5

1

1

11

8

5

18

3

21

3

6

5

1

3

2

4

1

3

7

8

1

3

1

9

14

6

3

14

1

3

2

1

12

2

9

3

1

8

8

2

1

6

5

1

1

23

S

6

5

4

2

21

8

7

5

9

1

1

1

14

1

1

2

1

9

2

8

7

2

5

5

9

7

3

3

5

1

3

15

4

13

4

2

1

1

1

1

2

3

7

0

7

3

2

12

22

14

13

g

9

1

3

2

6

4

6

7

6

2

2

1

1

7

g

4

3

1

2

2

2

2

1

1

1

5

9

1

5

7

5

2

15

1

10

22

6

13

8

2

2

17

1

6

1

1

4

1

1

5

5

1

8

1

5

1

15

3

15

4

1

1

1

10

4

4

1

2

Outside 

the 

lagoon

00 CO

ss

CO

•*

CM

CO•*

111

8 10

2

1 4

1 1

3 2

3 3

2

2

1

1

1

1

1

6

1 1

1 3

713

3

1

1 13

1

1 1

523

1

2 5

1 5

1

I

1

I

) 1

17

1

i 1

)

i

3

7

12

1

8

2

2

1

2

1

2

2

1

2

1

1

4

1

6

6

5

2

2

1

1

1

2

1

1

1

1

BIKINI ATOLL

beach

-* o

CM

B n

in

ffl

o 

ffl

•*

CO

I

2

7

I

T

i

2

2

1

2

1

reef flat

0 CM M in

n «

rffl

ic

CO

CM

CO

ffl

CO

n

in

P: 

ffl

1

1

)

1

1

6

3

3

1

1

1

Inside the lagoon

r- ix

•*

c 

P

r-•* 0)

0)

S

«n

IT

o •*

m

TP 

n

S 0

•* CO 

CO

•*

00 

CO

rCO

CO CM

O•*

CO 

ul B cc

CMin

o

in

a

52

c

CO

CM 

CM

a

CD

r1

o

•* 5<

•* •* •* S

in

S

^f in 3

o

CO •* •*

en i- CM

"* 

o

I

CM

3

CM

0 

0

CM

0

•* CO 

CM

r- CM 

00

TP

00

•*

CM

CO •*

CM 

CM

•* •* •* in•* S

cc 

in CO •* •*

HI 111 1 1 Hlil 111 |3|

4

8 5

1

5

5

0

£.

1

8

1

2

8

11

0

1

8

5

4

3

8

1

2

4

1

1

3

1

1

1

3

9

8

1

15

5

5

3

5

5

20

1

5

2

7

2

8

4

7

3

3

1

1

3

17

2

1

1

5

5

1

2

8

6

2

5

6

4

16

8

2

1

4

1

4

1

1

7

1

3

2

2

15

2

20

2

3

1

1

4

1

6

6

1

1

2

1

1

1

4

1

1

/

1

1

1

2

7

1

1

1

7

1

5

1

1

2

9

1

9

1

1

1

1

/

2

2

5

4

1

2

9

1

1

16

13

1

1

11

4

1

2

8

2

1

4

3

3

1

8

5

11

3

1

9

1

7

2

2

4

1

2

1

1

1

2

1

5

15

1

19

2

2

2

1

1

1

2

1

6

1

2

1

2

1

2

2

2

2

1

3

1

2

1

g

2

1

2

1

9

2

5

14

3

1

1

6

5

12

1

23

2

2

1

8

20

1

1

5

3

20

4

1

2

1

1

1

2

2

2

2

I

3

1

1

2

3

1

1

2

2

1

6

2

5

2

3

3

6

1

2

5

4

6

2

3

9

1

1

2

7

J

9

1

2

4

3

6

1

5

14

19

7

6

3

4

1

4

14

13

5

6

10

6

18

11

2

11

2

8

9

4

1

6

5

7

1

11

2

5

3

4

2

1

2

1

1

2

1

S

5

9

5

6

7

3

5

5

4

1

15

7

8

4

3

1

1

,1

6

S

7

1

21

19

20

3

2

18

0

1

21

3

1

2

2

9

3

2

4

4

6

1

1

3

2

18

4

1

4

4

8

16

7

1

J

7

16

3

9

1

17

12

5

1

J

1

1

2

7

6

14

5

3

13

3

1

13

4

1

4

2

3

2

12

2

2

1

2

1

3

2

1

1

2

1

1

1

1

72

17

2

4

.t

2

9

24

1

6

1

1

2

3

6

6

20

6

3

5

1

22

8

2

1

1

1

2

1

3

8

7

1

7

1

i

1

g

3

2

1

1

21

4

12

1

1

13

12

3

5

2

6

1

12

8

10

1

4

5

3

3

1

2

3

2

3

6

1

5

1

4

9

4

10

6

2

L

5

2

5

5

1

2

3

4

6

4

3

2

?^

18

17

3

4

6

1

2

4

1

1

1

1

1

2

3

5

2

1

2

4

2

2

14

12

1

1

23

16

3

20

1

t

I

4

6

20

6

/

10

9

7

1

2

3

1

2

2

2

13

3

11

9

18

3

16

3

4

7

3

4

4

1

3

7

19

7

6

5

1

,1

2

1

1

2

2

1

7

24

5

14

1

9

7

22

2

2

9

t

1

7

7

2

19

6

3

3

8

2

20

1.

1

1

2

1

2

1

4

.1

S

1

6

3

7

11

11

3

1

1

5

1

4

3

6

9

2

3

1

1

2

4

1

2

6

6

10

8

2

15

20

4

2

5

4

2

1

2

?

3

1

2

1

12

3

1

4

1

8

1

1

1

1

3

1

1

2

1

5

t

g

8

2

1

1

9

6

2

21

3

2

3

8

6

3

4

4

9

7

1

6

7

1

5

6

6

3

4

4

4

3

3

1

1

2

1

4

2

1

.1

S

9

8

1

16

8

7

1

1

12

1

2

7

5

2

2

2

1

r

9

2

1

1

1

4

7

1

1

2

1

1

2

1

1

11

2

3

3

3

1

1

23

23

12

11

2

7

4

1

3

1

7

3

3

1

3

2

1

10

3

4

1

3

8

10

3

3

1

3

4

4

3

2

1

2

1

9

1

Outer slopes

Dredge 

samples

00I2

en •* 21

00

21

o

CO

2

15 5

3

S

5

3

2

1

1

12

13

5

1

8

2

1

1

10

3

4

1

4

1

1

1

3

3

4

1

1

2

9

1

1

1

2

2

2

1

1

2

1

1

4

5

3

1

5

2

1

1

2

1

5

R

1

1

1

2

1

Bottom samples

M

CM rco

21

CO CO

0

3

CO

CO

i-

"tric- 00

S

CO

o

03

III 1

1 1|

1 1 1 1 1 1

7

5

2

6

21

1

5

2

1

4

6

1

2

1

3

3

1

1

2

1

1

3

1

1

2

2

1

3

4

2

1

1

1

1

1

1

5

1

14

2

23

2

2

2

2

1

6

13

2

1

3

4

6

1

1

9

8

4

1

11

2

15

6

3

3

2

1

1

3

1

3

3

2

8

5

1

|

3

2

1

1

1

1

2

1

4

1

1

1

1

3

1

1

11

1

3

3

2

2

1

3

2

2

2

3

4

6

1

1

2

1

Guyot

"CM

•*

CO

21 •*

'bo

•* 

i3

"o

•* i2!

o

•* 

r- a

o

?

~n 

f-

"ip

•*

in 

io

CO

I

CM

CO 

IICO

c- to

o

o 

rCM

0

S

<"

CM

O 

>

1170(4-6")

CM

1 1

HI

T

6

1

3

1

1

1

4

1

1

1

2

4

3

2

2

1

2

6

1

1

2

2

7

1

3

1

3

3

1

3

1

1

2

1

1

2

2

3

1

4

2

1

i

i

3

1

3

7

1

1

2

1

1

1

7

1

4

5

1

1

9

2

1

2

1

1

3

1

12

3

1

1

1

2

1

1

3

7

2

2

2

1

1

1

1

5

3

J.L

i

3

1

1

1

1

1

1

ENIWETOK ATOLL,

Inside the lagoon

S a

o

CM n

CM 1

CM CO 

00

f00 

CM

00

oc

CM

CD 

00 

CM

o

CM

0) 

CM

CM S en

CM 

CM

CO

en

in

•*

CO 

CM 

«

CO 

CM

10

H

CO

en

CM 3

03 

CM

O>

en o NCO

o

CO

r- 00 0

•* o

= =

CM

3 o

CM

CO "

<*„

i 1 1 1 III 1 1 1 T2T 11 111 HI

I 1 1

1

4

3

1

2

1

4

4

3

19

4

1

11

1

2

1

18

4

50

2

12

1

1

1

5

5

1

1

2

1

4

3

2

1

1

8

0

1

2

1

1

1

10

7

15

2

1

6

5

7

4

1

4

2

7

1

3

1

1

1

5

15

2

7

4

4

3

6

8

10

4

3

4

5

5

1

S

2

S5

1

1

1

10

1

8

3

1

1

3

4

1

4

1

7

3

10

5

40

1

1

2

4

3

2

1

4

1

1

1

3

1

JO

5

6

1

2

1

1

1

1

2

1

t

5

S5

11

7

1

1

1

4

7

2

7

5

3

9

1

1

1

2

13

1

2

22

19

20

8

15

1

2

1

4

6

8

18

2

1

3

3

1

7

3

85

6

55

2

2

1

2

1

8

1

6

2

6

3

6

1

3

8

6

6

14

1

1

S

5

25

5

1

1

3

3

4

1

2

1

1

1

1

11

2

12

5

1

14

3

2

1

1

IS

9

60

1

2

1

9

1

4

18

1

11

4

1

3

2

10

8

3

11

2

9

i

S

1

4

1

I

8

1

1

5

1

10

4

4

7

6

6

21

2

10

1

S

9

1

5

1

15

1

1

1

S

22

14

5

13

3

t

6

6

9

3

3

5

1

20

8

1

5

18

4

2

1

2

1

3

9

0

10

2

S

1

8

2

1

3

2

3

3

2

1

5

1

4

1

15

6

15

17

0

1

8

9

3

5

1

3

4

2

3

4

4

6

5

1

4

3

7

1 1

8

4

7

1

1

1

1

10

3

3

1

2

2

2

9

12

8

3

4

13

1

1

2

1

16

2

2

18

8

1

4

2

1

6

^

0

*

3

21

1

7

1

3

2

1

1

17

16

14

1

8

9

17

13

1

1

5

3

0

4

2

8

13

1

0

3

7

4

4

1

1

3

3

5

3

1

2

1

S

6

5

5

2

5

6

20

5

3

12

3

2

2

4

1

1

1

1

2

4

4

3

1

2

SO

9

SO

7

8

15

1

1

1

11

1

2

3

11

2

4

1

1

7

5

1

7

2

7

9

5

4

2

1

1

1

15

1

g

6

16

4

11

18

1

11

20

1

10

1

1

12

6

7

5

1

0

20

16

5

1

3

7

1

3

3

IS

9

£0

3

4

1

4

14

18

15

3

2

6

7

5

10

1

7

1

1

1

13

3

3

10

2

3

15

6

20

20

1

5

11

g

14

1

5

1

1

7

1

2

7

7

2

5

2

3

8

1

3

1

14

21

10

1

1

2

S

14

5

3

15

2

3

1

2

1

21

/

3

1

1

6

3

8

2

7

3

9

6

3

7

2

1

7

1

4

14

1

13

2

5

9

9

b

2

5

9

0

7

0

1

1

1

3

1

17

1

3

1

1

2

1

t

5

S

0

5

1

12

12

4

1

1

7

2

4

5

7

2

4

11

9

13

13

8

2

0

6

9

8

3

1

1

S

7

t

1

6

1

1

2

1

2

S

7

2

1

1

1

3

13

1

7

1

3

3

2

2

2

2

5

4

1

3

2

1

1

3

1

3

1

6

2

2

4

8

6

3

1

1

2

1

4

1

1

2

2

1

1

1

1

•!

3

0

3

1

1

5 1

4 4

701

1

[

1

516 a

11

'HO 1

5

3 2

5

1

1

1

5

20

5

3

11

20

10

17

5

4

1

1

£

4 1

i e

i .

4 .

6 f

7

2

2

2

3

721

1

2

3

i 13 1

> 1

!

? 2

2

3 5

6

1

3 3

2 4

5 1

7

2

Outer 

slopes

B °?

1

* *

? 3

8 U

1

1

3

5

4

4 2

3 1

1

9 :

2 '<.

1

15 £

231

1

3

9

4

7

^•41

t

9

1

6

•> 7

1

1

??

2

2

9

1

1

1

1 1

1

1

i 4

0 3

3

1

5 1

? 6

1 2

2

0 2

1

1

1

2

1

1

1

1 2

2

1

1

3 5

1

291053 O - 54 (Face p. 338)
RECENT PORAMINIFERA OF THE MARSHALL ISLANDS 239

This species occurs fairly commonly at a few stations. 

The periphery is broadly truncate and keeled and the 

aperture is compressed and flaring and does not extend 

out beyond the outline of the test.

Triloculina kerimbatica (Heron-Alien and Earland) 

Plate 85, figures 10, 11

Miliolina kerimbatica Heron-Alien and Earland, Zool. Soc" 

London Trans., v. 20, pt. 2, p. 574, pi. 43, figs. 13-23, 1915' 

Quinqueloculina kerimbatica Cushman, Carnegie Inst. Washington Pub. 342, p. 60, pi. 22, fig. 9, 1924. 

Miliolina parkeri Millett (not H. B. Brady), Royal Micros. Soc.

Jour., 1898, p. 507, pi. 12, figs. 4a, b.

Test free, quinqueloculine. The walls of the chambers thick, 

irregularly furrowed in all directions with broad, deeply gougedout channels, running obliquely and irregularly across the face of 

each chamber and generally connecting with a deeper straight 

furrow excavated down the peripheral edge. This straight 

peripheral furrow, when exposed on an earlier chamber in the 

center of the test by the quinqueloculine arrangement of the shell, 

affords a very striking appearance by contrast with the transverse 

furrows on the surface of the surrounding chambers. Aperture 

large and furnished with a prominent tooth. The oral end is 

usually but slightly produced, but in some individuals the aperture is situated on a produced neck. The ridges between the 

furrows are flat on the top, «. e., the furrows are cleanly gouged 

out of the shell substance.—Heron-Alien and Earland, 1915.

The above is the original description of this species 

the types of which are from the Kerimba Archipelago. 

It has been recorded from a number of localities in the 

Indo-Pacific. Our specimens, although they vary 

somewhat, are much more uniformly ornamented than 

the series of figures given by Heron-Alien and Earland. 

The species appears to be closely related to that 

referred to T. cf. bicarinata D'Orbigny but the ornamentation consists of more definite wavy ridges than 

of a series of pits. The nepionic stages are quinqueloculine but the adults would indicate that the species 

should be placed in Triloculina.

Triloculina marshallana Todd, n. sp. 

Plate 85, figure 13

Test elongate-triangular, tricarinate, irregular in 

shape, apertural end more acute than apical end, periphery bluntly angled; chambers distinct, moderately 

inflated, each chamber as added projecting out with an 

angular face beyond the previous ones; sutures distinct, 

depressed; wall with a fine matte surface, glistening; 

aperture large, projecting, at the end of a short, broad 

neck, with an elongate tooth, bifid at the tip, surrounded 

by a slight lip above which the tooth is usually visible 

hi side view. Length 0.50 to 0.65 mm, breadth 0.30 

to 0.37 mm.

Holotype, USNM 548594, from Rongelap lagoon, 

10 fathoms, 11°14'19" N., 166°38'17" E. (Rap 483).

This species is characterized by its irregular and

angular appearance due to the chambers projecting out 

beyond the earlier formed ones. It differs from T. 

trigonula (Lamarck) in its more elongate form r.nd 

irregular shape and its projecting apertural neck.

Triloeulina cf. T. oblonga (Montagu) 

Plate 85, figures 5-7

Vermiculum oblongum Montagu, Testacea Brittanica, p. 522,

pi. 14, fig. 9, 1803. 

Triloculina oblonga D'Orbigny, Annales sci. nat., tome 7, p. 300,

no. 16, Modeles, no. 95, 1826. 

Parker, Jones, and H. B. Brady, Annals and Mag. Fat.

History, 3d ser., v. 16, p. 34, pi. 1, fig. 9, 1865. 

Miliolina oblonga H. B. Brady, Challenger Kept. Zoology, v. 9,

p. 160, pi. 5, figs. 4a, b, 1884.

Numerous specimens from all the atolls are tertatively referred to this species. They vary considerably 

in proportionate breadth, most of them being slenderer 

than the typical form which was described from off the 

British Isles. Except for their triloculine form, seme 

are close to the specimens here referred to Quinqueloculina, cf. seminulum (Linne") and it may be that tl ese 

two groups belong together, the ones referred to 

Quinqueloculina being the microspheric forms, and the 

ones referred to Triloculina the megalospheric forms. 

The slenderer forms are close to Quinqueloculina 

bosciana D'Orbigny (1839a, p. 191, pi. 11, figs. 22-24) 

described from the West Indies but are larger and 

triloculine rather than quinqueloculine.

Triloculina spinata Cushman 

Plate 85, figure 8

Triloculina spinata Cushman, U. S. Natl. Mus. Bull. 161, p*;. 1, 

p. 56, pi. 12, fig. 5, 1932.

This small, slender species is distinguished by the 

spine on the apical end and its elongate apertural neck. 

It was described from Rongelap Atoll and also occurs 

rarely in the present material from Eniwetok and 

Rongerik.

Triloculina subplanciana Cushman, n. sp. 

Plate 85, figure 17

Test elongate, about 2% tunes as long as bread, 

periphery subacute, initial end rounded, apertural end 

with a short neck; chambers distinct, slightly inflated, 

increasing rapidly in size as added; sutures distinct, 

very slightly depressed; wall thin, translucent, smooth 

or very finely costate; aperture terminal, with a si ort 

neck and phialine lip and a short tooth sometimes 

slightly expanded at the top. Length 0.45 to 0.53 mm, 

diameter 0.20 to 0.23 mm.

Holotype, USNM 548540, from Rongerik lagoon, 85 

feet, 11°23'35" N., 167°28'35" E. (Rik 54).

This species differs from Triloculina, plancwna
340 BIKINI AND NEAKBY ATOLLS, MARSHALL ISLANDS

D'Orbigny (1839a, p. 173, pi. 9, figs. 17-19) in the 

more compressed test, subacute periphery, and definitely 

projecting neck with a rounded lip and a short tooth. 

The wall is usually thin, showing the earlier chambers.

Triloculina terquemiana (H. B. Brady) 

Plate 85, figure 9

Miliolina terquemiana Brady, Challenger Rept., Zoology, v. 9,

p. 166, pi. 114, figs, la, b, 1884. 

Heron-Alien and Earland, Zool. Soc. London Trans., v. 20,

p. 563, pi. 41, figs. 29-31 (not figs. 26-28), 1915. 

Triloculina terquemiana Cushman, U. S. Natl. Mus. Bull. 71,

pt. 6, p. 72, pi. 27, figs. 2a, b, 1917.

This species, described from shallow sediments of 

Ceylon and Madagascar, occurs rarely at a few stations.

In its surface it resembles Quinqueloculina neostriatula Thahnann but differs in its trilocuiine shape, 

larger size, and its aperture, which is entirely terminal 

whereas that of Quinqueloculina, neostriatula is hoodlike.

Triloculina tricarinata D'Orbigny 

Plate 85, figures 15, 16

Triloculina tricarinata D'Orbigny, Annales sci. nat., v. 7, p. 299,

no. 7; Modeles, no. 94, 1826. 

Parker, Jones, and H. B. Brady, Annals and Mag. Nat.

History, 3d ser., v. 16, p. 34, pi. 1, fig. 8, 1865. 

Cushman, U. S. Natl. Mus. Bull. 161, pt. 1, p. 59, pi. 13, figs.

3a, b, 1932.

Miliolina tricarinata H. B. Brady, Challenger Rept., Zoology. 

v. 9, p. 165, pi. 3, figs. 17a, b, 1884.

There are many records for this species but apparently not all referring to the same form, and no attempt 

is made to include them. Triangular forms with sharp 

angles are here included under this name.

Triloculina trigonula (Lamarck) 

Plate 85. figure 18

Miliola trigonula Lamarck, Annales Museum, tome 5, p. 351,

no. 3, 1804; v. 9, pi. 17, fig. 4, 1807. 

Triloculina trigonula D'Orbigny, Annales sci nat., tome 7, p. 299,

no. 1, pi. 16, figs. 5-9; Modeles, no. 93, 1826. 

Cushman, U. S. Natl. Mus. Bull. 161, pt. 1, p. 56, pi. 13, figs.

la, b, 1932. 

^Hliolina trigonula Williamson, Recent Foraminifera of Great

Britain, p. 84, pi. 7, figs. 180-182, 1858. 

H. B. Brady, Challenger Rept., Zoology, v. 9, p. 164, pi. 3,

figs. 14-16, 1884.

A few specimens of a triangular form with rounded 

gles are placed in this species.

Triloculina sp. A 

Plate 85, figure 20

Test elongate, compressed, 2J4 to 3 times as long as 

broad, periphery acute; chambers not inflated, of about 

equal breadth throughout, curved at basal end; sutures 

indistinct; wall having a rather coarse matte surface 

occasionally with slight keels on the periphery ; aperture

an

elongate, compressed, surrounded by a narrow lip, filled 

by an elongate tooth. Length 1.00 to 1.25 mm, breadth 

0.25 to 0.45 mm.

This species occurs in some numbers at Eniwetok. 

It is distinguished by its elongate and compressed form.

Triloculina sp. B 

Plate 85, figure 21

Test elongate, triangular in section, 2 to 1% times as 

long as broad, peripheral angles acute but rounded, 

apical end bluntly pointed, apertural e»id obliquely 

truncate so aperture is visible in side view; chambers 

not inflated; sutures indistinct, slightly depressed; wall 

smooth, polished; aperture circular, without any 

thickened rim or lip, partially filled by delicate bifid 

U-shaped tooth which extends across about one-third 

the diameter of the opening. Length 0.5£ to 0.78 mm, 

breadth 0.22 to 0.38 mm.

The species was found only in the deeper waters 

outside Bikini Atoll.

Genus PYR60 Defrance, 1824

Pyrgo denticulata (H. B. Brady)

Plate 85, figure 22

Bilocultna ringens Lamarck var. denticulata H. B. Brady, 

Challenger Rept., Zoology, v. 9, p. 143, pi. 3, figs. 4, 5, 1884.

Biloculina denticulata Cushman, U. S. Natl. Mus. Bull. 71, pt. 6, 

p. 80, pi. 33, fig. 1, 1917.

Pyrgo denticulata Cushman, U. S. Natl. Mus. Bull. 104, pt. 6, p. 

69, pi. 18, figs. 3, 4, 1929.

This is a species typical of coral reef regions, especially 

in the Pacific. There is considerable variation in the 

development of the basal spines. It is common in the 

Marshall Islands material, occurring in numerous 

samples.

Pyrgo denticulata var. striolata (H. B. B"ady) 

Plate 85, figure 23

Biloculina ringens Lamarck var. striolata H. B. Brady, Challenger

Rept., Zoology, v. 9, p. 143, pi. 3, figs. 7, 8, 1884. 

Biloculina denticulata (H. B. Brady) var. 'striolata Cushman, U. S.

Natl. Mus. Bull. 71, pt. 6, p. 80, pi. 33, figs. 2, 3, 1917. 

Pyrgo denticulata (H. B. Brady) var. striolata Cnshman, U. S.

Natl. Mus. Bull. 104, pt. 6, p. 69, pi. 18, fi-js. 5a-c, 1929;

idem, Bull. 161, pt. 1, p. 63, pi. 14, figs. 10, 11, 1932.

Specimens occur in a number of samples but are not 

abundant. The specimens show the typical striae in 

varying degree.

Pyrgo lucernula (Schwager) 

Plate 85, figure 24

Biloculina lucernula Schwager, Aforara-Exped., CJeol. Theil,

Band. 2, p. 202, pi. 4, figs. 14, 17, 1866. 

Pyrgo lucernula Thalmann, Eclogae geol. Helvetiae, v. 25, pp.

295, 296, 1932. 

Biloculina bulloides H. B. Brady (not D'Orbigny), Challenger

Rept., Zoology, v. 9, p. 142, pi. 2, figs. 5, 6, 1884.
RECENT FORAMINIFERA OF THE MARSHALL ISLANDS 341

Flint, U. S. Natl. Mus. Rept. for 1897, p. 293, pi. 38, fig. 5, 

1899.

This species, described from the Pliocene of Kar 

Nicobar, occurs rarely in a number of samples at 

Bikini and in one sample from Eniwetok. Specimens 

vary considerably in shape but are characterized by a 

protruding apertural neck.

Pyrgo millettii (Cushman) 

Plate 85, figure 25

Biloculina millettii Cushman, U. S. Natl. Mus. Bull. 71, pt. 6, p.

81, pi. 34, figs. 4, 5, 1917. 

Pyrgo millettii Cushman, U. S. Natl. Mus. Hull. 161, pt. 1, p. 66,

pi. 15, figs. 4, 5, 1932. 

Miliolina durrandii Millett (part), Royal Micros. Soc. Jour ,

1898, p. 268, pi. 6, figs. 8-10 (not fig. 7). 

Heron-Alien and Earland, Zool. Soc. London Trans., v. 20, p.

565, pi. 42, figs. 11-16, 1915.

A very few specimens, all immature and in the 

triloculine stage, occur at Eniwetok and Bikini. This is 

a unique species in that its compression is at right 

angles to the usual plane of compression in this genus.

Pyrgo murrhina (Schwager)

Biloculina murrhina Schwager, Novara-Exped., Geol. Theil,

Band 2, p. 203, pi. 4, fig. 15, 1866. 

Pyrgo murrhina Cushman, U. S. Natl. Mus. Bull. 161, pt. 1, p.

64, pi. 15, figs. 1-3, 1932.

A few specimens from Bikini, circular, and keeled 

around the periphery, seem to belong to this species 

described from the Pliocene of Kar Nicobar and widely 

recorded in Kecent seas.

Pyrgo sp. 

Plate 85, figure 26

A few specimens similar to that figured have a 

roughly quadrangular test, with sutures not depressed 

and chambers not inflated nor angular along the 

periphery. They are somewhat similar to P. elongata, 

(D'Orbigny) (Parker, Jones, and Brady, 1871, p. 247, 

pi. 8, fig. 6) but have an elongate, slitlike aperture.

Family OPHTHALMIDHDAE

Genus CORNUSPIRA Schultze, 1854

Cornuspira planorbis Schultze

Plate 85, figure 27

Cornuspira planorbis Schultze, Organismus Pclythal., p. 40, pi. 2,

fig. 21, 1854. 

Cornuspira involvens Cushman (not Reuss), U. S. Natl. Mus.

Bull. 161, pt. 1, p. 67, pi. 16, figs. 2a, b, 1932.

The species occurs rarely in the Marshall Islands.

Genus VERTEBRALINA D'Orbigny, 1826 

Vertebralina striata D'Orbigny

Vertebralina striata D'Orbigny, Annales sci. nat., tome 7, p. "S3,

no. 1; Modeles, no. 81, ive livraison, 1826. 

Cushman, U. S. Natl. Mus. Bull. 161, pt. 1, p. 73, pi. 16, figs.

8-10, 1932.

Cushman and Todd, Cushman Lab. Foram. Research Cortr., 

v. 20, pt. 3, p. 74, pi. 12, figs. 7-11. 1944.

A few nepionic specimens, some fragmentary, were 

found at Bikini and Kongelap.

Genus PIANISPIRINA Seguenza, 1880 

Planispirina exigua (H. B. Brady)

Plate 85, figure 28

Hauerina exigua H. B. Brady, Quart. Jour. Micros. Sci., v. 19,

p. 53, 1879. 

Planispirina exigua H. B. Brady, Challenger Rept., Zoology, v.

9, p. 196, pi. 12, figs. 1-4; woodcut, fig. 5b, 1884. 

Cushman, U. S. Natl. Mus. Bull. 161, pt. 1, p. 71, pi. 16, figs.

7a, b, 1932.

Kare specimens were found at Eniwetok 329 and a 

single specimen at Rongelap 480.

Genus WIESNERELLA Cushman, 1933

Wiesnerella auriculata (Egger)

Plate 85, figure 30

Planispirina auriculata Egger, K. bayer. Akad. Wiss., Math.-

naturh. Abt., Abh., Kl. 2, Band 18, p. 245, pi. 3, figs.13-15,

1893. 

Cushman, U. S. Natl. Mus. Bull. 161, pt. 1, p. 72, pi. 16, figs

6a-c, 1932. 

Wiesnerella auriculata Cushman, Cushman Lab. Foram. Research

Contr., v. 9, p. 33, pi. 3, figs. 7-9, 1933.

This species has a wide distribution in the Atlantic 

and Pacific. Typical specimens occur rarely in several 

samples from Eniwetok, Bikini, and Rongelap.

Genus NUBECULARIA Defrance, 1825

Nubecularia lacunensis Chapman

Plate 85, figure 29

Nubecularia lacunensis Chapman. Linnean Soc. Jour., Zoology, 

v. 28, p. 169, pi. 19, fig. 2; p. 206 (list), 1900; v. 28, p. 381, 

386, 1902.

Test porcellaneous, opaque or translucent, apparently 

attached by flat or concave surface, periphery acute 

and irregular; first few chambers coiled, later ones 

irregularly spreading.

A few specimens were found at all the islands.
342 BIKINI AND NEARBY ATOLLS, MARSHALL ISLANDS

Genus PARKIN A Cushman, 1931 

Parrina bradyi (Millett) 

Plate 85, figures 31-34

Nubecularia bradyi Millett, Royal Micros. Soc. Jour., 1898'

p. 261, pi. 5, figs. 6a, b. 

Silvestria bradyi Schubert, Palaeont. Zeitschr., Band 3, p. 166,

1920. 

Parrina bradyi Cushman, Cushman Lab. Foram. Research

Contr., v. 7, p. 20, 1931; U. S. Natl. Mus. Bull. 161, pt. 1,

p. 74, pi. 17, figs. 1-4, 1932. 

Nubecularia inflata H. B. Brady (not Terquem), Challenger Rept.,

Zoology, v. 9, p. 135, pi. 1, figs. 5-8, 1884.

It is highly variable in shape, and some immature 

specimens resemble Miliolinella labiosa (D'Orbigny). 

This species is widely distributed in warm, shallow 

waters of the Mediterranean and about the islands of 

the Indo-Pacific.

Family TROCHAMMINIDAE

Genus TROCHAMMINA Parker and Jones, 1859

Trochammina cf. T. rotaliformis J. Wright

Plate 85, figure 35 

Trochammina rotaliformis J. Wright Heron-Alien and Earland,

Royal Irish Acad. Proc., v. 31, pt. 64, p. 52, pi. 3, 

13, 1913.

11-

A few specimens from Eniwetok, Bikini, and Rongelap are similar to this species. They are small and 

rounded, with indistinct chambers and sutures, and 

3 or 4 chambers in the last whorl.

Genus ROTALIAMMINA Cushman, 1924

Rotaliammina mayori Cushman

Plate 85, figure 36

Rotaliammina mayori Cushman, Carnegie Inst. Washington 

Pub. 342, p. 11, pi. 1, figs. 4, 5, 1924.

Typical specimens of this species, described from 

shallow water of Samoa, occurs at Rongelap 468. This 

is an attached form and from the dorsal side appears 

to be a Trochammina surrounded by a thin flange of 

lighter colored material.

From the ventral side it is apparent that no ventral 

wall to the chambers exists, but the septal partitions 

indicate the chamber divisions.

Genus CARTERINA H. B. Brady, 1884 

Carterina spiculotesta (Carter)

Plate 85, figure 37

Rotalia spiculotesta Carter, Annals and Mag. Nat. History, 4th

ser., v. 20, p. 470, pi. 16, figs. 1-3, 1877. 

Carterina spiculotesta H. B. Brady, Challenger Rept., Zoology,

v. 9, p. 346, pi. 41, figs. 7-10, 1884.

A single specimen was found in Bikini sample 814. 

The thin wall is very distinctive in being formed of a

single layer of elongate sponge spicule? very neatly 

cemented together.

Genus NOURIA Heron-Alien and Earlar-i, 1914 

Nouria polymorphinoides Heron-Alien aid Earland

Nouria polymorphinoides Herou-Allen and Earland, Zool. Soc_ 

London Trans., v. 20, pt. 12, p. 376, pi. 37, figs. 1-14, 1914, 

Cushman, U. S. Natl. Mus. Bull. 161, pt. 1, p. 78, pi. 17, fig. 9, 

1932.

Test longer than broad, somewhat compressed, periphery rounded; chambers few, tending to be biserial 

in adult, irregular; sutures slightly depressed; wall 

arenaceous, composed of sand grains of various sizes 

rather neatly cemented; aperture simple, terminal. 

Maximum length 1.5 mm.

The types of this species are from the Kerimba 

Archipelago. It has been widely recorded in the 

Pacific. Very rare specimens were found in the 

Marshall Islands.

Family LAGENIDAE

The occurrence of species of this family in the Marshall Islands is of particular interest because, unlike 

the other families, this family is very restricted in its 

distribution. None were found at Ronge^ik, and only a 

few at Eniwetok and Rongelap. The family is fairly well 

represented at Bikini by a large number of species, but 

not in abundance. The presence of species of the Lagenidae seems to be indicative of moderate to great depth 

and open-sea conditions.

Genus ROBULUS Montfort, 180S

Robulus calcar (Linne)

Plate 86, figure 4

Robulus calcar (Linnaeus) Cushman, U. S. ! T atl. Mus. Bull. 

161, pt. 2, p. 7, pi. 2, figs. 3a, b, 1933.

A few small, apparently young, specimens were found 

at several Bikini stations.

Robulus gibbus (D'Orbigny) 

Plate 86, figure 2

Cristellaria gibba D'Orbigny, Annales sci. nat., v. 7, p. 292, no. 

17, 1826; D'Orbigny, in De la Sagra, Histoire physique, 

politique et naturelle de 1'Ile de Cuba, Foraminiferes, p. 

40, pi. 7, figs. 20, 21, 1839.

Parker, Jones, and H. B. Brady, Annals and Mag. Nat. History, 4th ser., v. 8, v. 253, 1871.

H. B. Brady, Challenger Rept., Zoology, v. 9, p. 546, pi. 69, 

figs. 8, 9, 1884.

Robulus gibbus Cushman, U. S. Natl. Mus. Bull. 161, pt. 2, p. 

6, pi. 2, figs. 2, 6, 7, 1933.

A few specimens, similar to that figured, occur at 

several Bikini stations. They are compact, relatively 

thick forms, with a blunt periphery, anc1 the chambers 

and sutures are rather indistinct.
RECENT FORAMINIFERA OP THE MARSHALL ISLANDS 343

Eobulus limbosus (Reuss)

Plate 86, figure 1

Robulina limbosa Reuss, Akad. Wiss. Wien, Math.-naturwiss.

Kl., Sitzungsber., Band 48, pt. 1, p. 55, pi. 6, fig. 69, 1863. 

Robulus limbosus Cushman, U. S. Natl. Mus. Bull. 161, pt. 2,

p. 3, pi. 1, figs. 4-6; pi. 2, figs. 5a, b, 1933. 

Cushman and McCulloch, Allan Hancock Pacific Exped., v.

6, no. 6, p. 297, pi. 38, fig. 8; pi. 39, figs. 1-4, 1950.

Specimens with few chambers (5 to 7), a keeled periphery, prominent umbones, and tangential slightly 

curved and limbate sutures are placed in this species. 

They occur rarely at a few Bikini stations.

Robulus nigriseptus (Koch) 

Plate 86, figure 3

'Cristellaria nigrisepta Koch, Schweizer. palaeont. Gesell. Ber., 

Band 19, no. 3, p. 740, fig. 14 (in text), 1926.

Robulus nigriseptus Cushman, U. S. Natl. Mus. Bull. 161, pt. 2, 

p. 5, pi. 1, figs. 9a, b, 1933.

A very few specimens from two Bikini stations and 

one Rongelap station seem to be the same as those 

recorded from the Tropical Pacific. They differ, however, from the type of B. nigriseptus in having a blunt, 

not keeled, periphery. They are also like B. clericii 

(Fornasini) (1895, text fig.) described from the Pliocene of Italy but the characteristic curvature at the 

inner ends of the sutures is not as strongly developed 

as in that species.

Robulus sp. A 

Plate 86, figure 6

Specimens with numerous chambers (8 to 10), 

strongly curved sutures, acute or slightly keeled periphery, and clear umbones occur rarely in several Bikini 

samples. They seem to be identical with the species 

referred to B. expansus (Cushman) var. planulatus 

(Cushman) (1933, p. 8, pi. 2, figs. 4a, b) but comparison 

Tvith the holotype of that variety shows they are distinct 

in being much smaller, biconvex and not compressed, 

.and lacking any surface ornamentation.

This species resembles specimens referred to R. vortex 

(Fichtel and Moll) (Cushman, 1933, p. 5, pi. 2, figs, 

la, b) and B. orbicularis (D'Orbigny) (Cushman and 

McCulloch, 1950, p. 296, pi. 38, figs. 4-7), but differs 

in the clear umbones and somewhat less strongly curved 

; sutures.

Robulus sp. B 

Plate 86, figure 5

This compressed species occurs rather commonly at 

Bikini 1174 (1-2") and more rarely at Bikini 1174. 

The umbones are clear, giving the effect of a slightly 

evolute test. Seven or eight chambers comprise the 

last whorl and the sutures are strongly curved. The 

periphery is acute .and slightly keeled.

Robulus sp. C 

Plate 86, figure 8

A few specimens from Bikini and Rongelap, similar 

to that figured, are inadequate for description. They 

are distinctly keeled and the curved sutures are limbr.te 

with the middle portion of each suture raised as a ridge 

of clear shell material.

Robulus sp. D 

Plate 82, figure 3

This large species is represented by only three specimens from three Bikini stations, but is figured for future 

reference. Six chambers comprise the last whorl, and 

around the periphery the large radiate apertures, 

belonging to each previous chamber, are visible.

Genus ASTACOLUS Montfort, 1808

Astacolus bradyi (Cushman)

Plate 86, figure 9

Cristellaria bradyi Cushman, U. S. Natl. Mus. Proc., v. 51, p.

659, 1917; idem, Bull. 100, v. 4, p. 243, pi. 48, figs, la, b,

1921. 

Cristellaria costata H. B. Brady (in part?), Challenger Re^t.,

Zoology, v. 9, p. 555, pi. 71, fig. 8, 1884.

Two nepionic specimens were found at Bikini.

Astacolus planulata Galloway and Wissler 

Plate 86, figure 7

Astacolus planulatus Galloway and Wissler, Jour. Paleontology,

v. 1, p. 46, pi. 8, fig. 5, 1927. 

Planularia planulata Cushman and Gray, Cushman Lab. Foram.

Research Special Pub. 19, p. 12, pi. 2, fig. 18, 1946. 

Cushman and McCulloch, Allan Hancock Pacific Exped., v.

6, no. 6, p. 303, pi. 40, figs. 1-5, 1950.

Specimens were rare at a few stations at Bikini. The 

one figured shows more of a coiled stage than most of 

the others.

Genus DENTALINA D'Orbigny, 1826

Dentalina cf. D. filiformis (D'Orbigny)

Plate 86, figure 10

Nodosaria filiformis D'Orbigny, H. B. Brady, Challenger Re^t., 

Zoology, v. 9, p. 500, pi. 63, figs. 3-5.

Very rare specimens from several Bikini stations are 

referred here with some question. They are slender, 

have an initial spine, and oblique sutures.

Genus NODOSARIA Lamarck, 1812

Nodosaria pauciloculata Cushman

Plate 86, figure 14

Nodosaria pauciloculata Cushman, TJ. S. Natl. Mus. Proc., v. 51, 

p. 655, 1917.

A few two-chambered specimens from Eniwetok 

Bikini seem to be the young of this species.
344 BIKINI AND NEARBY ATOLLS, MARSHALL ISLANDS

Nodosaria sp. A 

Plate 86, figure 11

Rare specimens, mostly fragmentary, represent a 

slender species with slightly inflated chambers, transverse sutures, and granular wall.

Nodosaria sp. B 

Plate 86, figure 12

Rare specimens, similar to that figured, are inadequate for description.

Nodosaria sp. C 

Plate 86, figure 13

This form may be an anomalous kind of Lagena and 

is recorded here for future reference. The ornamentation resembles that of N. prava Cushman (1933, p. 14, 

pi. 4, figs. 1-4) but the much greater constriction 

between the chambers and the transverse rings distinguish it from that species.

Genus FRONDICULARIA Defrance, 1824

Frondicularia sp. A of Boomgaart

Plate 86, figure 15

Frondicularia sp. A Boomgaart, Thesis Univ. Utrecht, p. 84, 

pi. 11, fig. 1, 1949.

Three single specimens, from Bikini 814, Bikini 817, 

and Eniwetok 117, seem very close to a specimen 

described and figured from late Tertiary sediments 

from a well at Bodjonegoro, Java. The present 

specimens are larger in size and number of chambers, 

but otherwise seem to be the same.

Genus LAGENA Walker and Jacob, 1798

Lagena acuticosta Reuss

Plate 86, figure 16

Lagena acuticosta Reuss, Cushman, U. S. Natl. Mus. Bull. 161, 

pt. 2, p. 34, pi. 8, figs. 9, 10, 12, 1933.

This species occurs rarely at Bikini. 

Lagena globosa (Montagu) 

Plate 86, figure 17

Lagena globosa (Montagu), Cushman, U. S. Natl. Mus. Bull. 104, 

pt. 4, p. 20, pi. 4, figs. 1, 2, 1923.

Single specimens were found at a few stations at, 

Bikini, Eniwetok, and Rongelap.

Lagena gracilis Williamson 

Plate 86, figure 18

Lagena gracilis Williamson, Cushman, U. S. Natl. Mus. Bull. 161, 

pt. 2, p. 33, pi. 8, figs. 5-7, 1933.

This species occurs rarely at several Bikini stations.

Lagena spiralis H. B. Brady 

Plate 86, figure 19

Lagena spiralis H. B. Brady, Cushman, U. S. Natl. Mus. Bull. 

161, pt. 2, p. 30, pi. 7, figs. 15, 16, 1933.

This is known as a species of the Indo-Pacific, 

occurring in shoal waiter. It occurs in typical form in 

the Eniwetok samples but is rare.

Lagena striato-punctata Parker and Jones

Lagena striato-punctata Parker and Jones, Cushman, U. S. Natl. 

Mus. Bull. 104, pt. 4, p. 55, pi. 10, fig. 10, 1923.

Very rare and very small specimens from Bikini and 

Eniwetok have the ornamented costae characteristic 

of this species.

Lagena williamsoni (Alcock) 

Plate 86, figure 20

Lagena williamsoni (Alcock), Cushman, U. S. ] Tatl. Mus. Bull. 

161, pt. 2, p. 34, pi. 8, fig. 8, 1933.

Single specimens were found at Bikini s nd Eniwetok.

Family POLYMORPHINIDAE

Genus DIMORPHINA D'Orbigny, 1826

Dimorphina tuberosa D'Orbigny

Plate 86, figures 21, 22

Dimorphina tuberosa D'Orbigny, Annales sci. nat., tome 7, p.

264, no. 1; Modeles, no. 60, iii e livraison, IP'26. 

Parker, Jones, and H. B. Brady, Annals and Mag. Nat. History, 3d ser., v. 16, p. 28, pi. 2, fig. 53, 1865.

Common specimens from several Bikini samples 

seem to belong in this species described from the 

Mediterranean. Only one specimen shows the polyserial arrangement of the early chambers. In all the 

other specimens the test is uniserial throughout.

Genus SIGMOMORPHINA Cushman and Ozawa, 1928 

Sigmomorphina semitecta (Reuss), var. terquemiana (Fornasini)

Sigmomorphina semitecta (Reuss), var. terquemiana (Fornasini), 

Cushman and Ozawa, U. S. Natl. Mus. Froc., v. 77, art. 

6, p. 129, pi. 33, figs. 4, 5; pi. 34, figs. 2, 3; pi. 35, fig. 1, 1930.

Specimens occur rarely at Bikini. 

Genus POLYMORPHINELLA Cushman and Hinzawa, 1936 

Polymorphinella paciflca Cushman and F«.nzawa 

Plate 86, figures 23, 24

Polymorphinella pacifica Cushman and Hanzawa, Cushman Lab.

Foram. Research Contr., v. 12, pt. 2, p. 47, 1936. 

Cristellaria schloenbachi Reuss, anomalous specimen, H. B.

Brady, Challenger Rept., Zoology, v. 9, pi. 67, fig. 8, 1884.

A few specimens from several Bikini stations apparently belong in this species which was named from
RECENT FORAMINIFERA OF THE MARSHALL ISLANDS 345

a specimen figured by Brady in the Challenger Report 

from Torres Strait. Two specimens are figured to 

show the two extremes of form. The species is only 

slightly biserial in the initial part and so may appear 

to be an anomalous form of Astacolus.

Family NONIONIDAE

Genus NONION Montfort, 1808

Nonion pacificum (Cushman)

Plate 86, figure 25

Nonionina umbilicatula (Montagu) var. pacifica Cushman, Carnegie Inst. Washington Pub. 342, p. 48, pi. 16, fig. 3, 1924.

Nonion umbilicatulum (Montagu) var. pacificum Cushman, 

Scripps Inst. Oceanography Bull., Tech. ser., v. 1, no. 10, 

p. 149, pi. 2, fig. 5, 1927.

Nonion pacificum Cushman, U. S. Natl. Mus. Bull. 161, pt. 2, 

p. 44, pi. 10, figs. 9a, b, 1933; B. P. Bishop Mus. Bull. 119, 

p. 120, pi. 14, figs. 7a, b, 1934- U. S. Geol. Survey Prof. 

Paper 191, p. 25, pi. 6, fig. 25, 1939.

Cushman and Todd, Cushman Lab. Foram. Research Special 

Pub. 15, p. 36, pi. 5, fig. 26, 1945.

Typical specimens occur at Bikini. They are very 

coarsely perforate, similar to N. pompilioides, but differ 

in being more compressed and not having llmbate 

sutures.

Nonion pompilioides (Fichtel and Moll) 

Plate 86, figure 26

''Nautilus melo" Soldani, Testaceographica, p. 38, pi. 8, figs, zz,

A, B, C, 1798. 

Nautilus pompilioides Fichtel and Moll, Testacea microscopica,

p. 31, pi. 2, figs, a-c, 1798. 

Nonionina pompilioides D'Orbigny, Annales sci. nat., tome 7,

p. 294, no. 15, 1826. 

H. B. Brady, Challenger Kept., Zoology, v. 9, p. 727, pi. 109,

figs. 10, 11, 1884. 

Nonion pompilioides Cushman, U. S. Natl. Mus. Bull. 104, pt. 7,

p. 4, pi. 1, figs. 7-11: pi. 2, figs. 1, 2, 1929; U. S. Geol. Survey

Prof. Paper 191, p. 19, pi. 5, figs. 9-12, 1939.

This species is common in several Bikini samples. 

It is known from the later Tertiary of southern Europe 

and is widely distributed in Recent seas.

Genus ASTROM ONION Cushman and Edwards, 1937 

Astrononion tumidum Cushman and Edwards

Plate 86, figure 27

Astrononion tumidum Cushman and Edwards, Cushman Lab.

Foram. Research Contr., v. 13, p. 33, pi. 3, fig. 17, 1937. 

Cushman, U. S. Geol. Survey Prof. Paper 191, p. 37, pi. 10, 

fig. 11, 1939.

Nonionina stelligera (part) H. B. Brady (not D'Orbigny), Challenger Kept., Zoology, v. 9, p. 728, pi. 109, fig. 5, 1884.

This species, named from the South Atlantic, occurs 

rarely at Bikini and Eniwetok.

Genus NONIONELLA Cushman, 1926

Nonionella translucens Cushman

Plate 86, figure 29

Nonionella translucens Cushman, U. S. Natl. Mus. Bull. 161, 

pt. 2, p. 45, pi. 11, figs. 2a-c, 1933; U. S. Geol. Surrey 

Prof. Paper 191, p. 34, pi. 9, figs. 13a-c, 1939.

The types of this species are from a depth of 12 

fathoms off Levuka, Fiji. It is apparently a species of 

the Pacific but not confined to shallow waters.

Nonionella sp. A

Nonionella sp. Cushman, U. S. Natl. Mus. Bull. 161, pt. 2, p. 

46, pi. 11, fig. 3, 1933.

A very few specimens from Bikini and Eniwetok 

seem to be the same as the specimen recorded in the 

above reference from 860 fathoms, in the tropical 

Pacific.

Nonionella sp. B §

Plate 86, figure 28

Test small, elongate, strongly compressed, periphery 

rounded; chambers about 10 in the last whorl, rapidly 

increasing in height as added, not inflated; sutures 

distinct, not depressed, slightly curved at the outer 

ends especially of the later ones; wall thin, transparent; 

aperture indistinct. Length 0.25 to 0.35 mm, breacfth 

0.17 to 0.20 mm, thickness 0.05 mm.

This species seems to be distinct but is inadequate 

for complete description. It resembles N. japonica 

(Asano), var. mexicana Cushman and McCulloch (1940, 

p. 160, pi. 17, figs. lOa-c) in its strongly compressed 

form but has fewer chambers and is somewhat larger. 

It also somewhat resembles N. limbato-striata Cushman 

(1934, p. 121, pi. 14, figs. lOa-c) from the Neogene of 

Fiji but is a narrower and more elongate form.

Genus ELPHIDIUM Montfort, 1808

Elphidium advenum (Cushman)

Plate 86, figure 30

Polystomella advena Cushman, Carnegie Inst. Washington Pub.

311, p. 56, pi. 9, figs. 11, 12, 1922. 

Elphidium advenum Cushman, U. S. Natl. Mus. Bull. 104, pt.

7, p. 25, pi. 10, figs. 1, 2, 1930; idem, Bull. 161, pt. 2, p. 50,

pi. 12, figs. 1-3, 1933; U. S. Geol. Survey Prof. Paper 191,

p. 60, pi. 16, figs. 31-35, 1939.

This widely distributed species in the Atlantic and 

Pacific occurs at all four atolls, buc less commorJy 

than the variety dispar. The occurrence of these t^vo 

forms may be of significance, as the typical form is 

common at Bikini and very rare at Eniwetok while the 

variety is abundant at Eniwetok and was not found at 

all at Bikini.
346 BIKINI AND NEARBY ATOLLS, MARSHALL ISLANDS

Elphidium advenum var. dispar Cushman, n. var. 

Plate 86, figure 31

Variety differing from the typical form in having the 

umbonal region slightly depressed, with the boss very 

much reduced or wanting, and in having the retral 

processes comprising a larger proportion of the surface 

of the test with the intervening chamber ridges slightly 

raised. Diameter 0.40 to 0.45 mm, thickness 0.20 to 

0.25 mm.

Holotype of variety (USNM 548508) from Kongerik 

lagoon, 111 feet, ll°22/32 // N., 167°28'48" E. (Kik 51).

This is the most common form of the genus in this 

material. It seems related to E. advenum (Cushman) 

in number and curvature of chambers, but may be a 

distinct species. It differs from E. fimbriatulum 

(Cushman) (1918, p. 20, pi. 8, fig. 5) in its more 

numerous and more strongly curved chambers, and its 

depressed urmboiial region.

Elphidium jenseni (Cushman) 

Plate 86, figure 32

Polystomella jenseni Cushman, Carnegie Inst. Washington Pub.

342, p. 49, pi. 16, figs. 4 (?), 6, 1924. 

Elphidium jenseni Cushman, U. S. Natl. Mus. Bull. 161, pt. 2,

p. 48, pi. 11, figs. 6, 7, 1933; U. S. Geol. Survey Prof. Paper

191, p. 62, pi. 17, figs. 14, 15, 1939. 

Polystomella macella (Fichtel and Moll), var. Jensen, Linnean

Soc. New South Wales Proc., v. 29, p. 817, pi. 23, fig. 4,

1904 (1905).

From the records this is a species of rather shallow 

warm water of the South Pacific. It is found mostly 

in water of less than 25 fathoms but may be carried 

into deeper water by currents. It may be confused 

with E. advenum var. dispar but is much more strongly 

compressed throughout.

Elphidium milletti (Heron-Alien and Earland)

Polystomella verriculata Millett (not H. B. Brady), Royal

Micros. Soc. Jour., 1904, p. 604, pi. 11, fig. 3. 

Polystomella milletti Heron-Alien and Earland, Zool. Soc.

London Trans., v. 20, p. 735, pi. 53, figs. 38-42, 1915. 

^Elphidium milletti Cushman, U. S. Natl. Mus. Bull. 161, pt. 2,

p. 49, pi. 11, figs. 8a, b, 1933; U. S. Geol. Survey Prof.

Paper 191, p. 58, pi. 16, figs. 20-22, 1939.

A single typical specimen was found at 3 to 5 inches 

in the core at Bikini 1173.

Elphidium simplex Cushman 

Plate 86, figure 33

Elphidium simplex Cushman, U. S. Natl. Mus. Bull. 161, pt. 2, 

p. 52, pi. 12, figs. 8, 9, 1933; U. S. Geol. Survey Prof. Paper 

191, p. 62, pi. 17, fig. 10, 1939.

The types of this species are from Vavau Anchorage' 

Tonga Islands. It is recorded rather widely in the 

Pacific.

Family CAMERINIDAE

Genus OPERCULINA D'Orhigny, 1826

Operculina ammonoides (Gronovius)

Plate 87, figure 1

Nautilus ammonoides Gronovius, Zoophylacium Gronovianum

etc., p. 282, pi. 19, figs. 5, 6, 1781. 

Linn6, Systema naturae Linnaei, Ed. 13 (Gmelin's), p. 3370,

1788. 

Operculina ammonoides Parker and Jones, Introd. Foram.,

Appendix, p. 310, 1862. 

H. B. Brady, Challenger Rept., Zoology, v. 9, p. 745, pi. 112,

figs. 1, 2, 1884; Royal Micros. Soc. Jour., 1887, p. 926. 

Operculina gaimardi D'Orbigny, Annales sci. nat., tome 7, p.

281, no. 5, 1826. 

Fornasini, Soc. geol italiana Boll., v. 22, fa^c. 3, p. 2 (396),

pi. 14, fig. 4, 1903.

Cushman, U. S. Natl. Mus. Bull. 161, pt. 2, p. 55, pi. 13, 1933.

Operculina granulosa (Leymerie) (?), Cushman, U. S. Natl.

Mus. Bull. 161, pt. 2, p. 56, pis. 14, 15; pi. 16, figs. 1-3, 1933.

Numerous specimens, mostly worn, occur at Eniwetok, Bikini, and Rongerik. Some eoecimens are 

ornamented by bosses along the early suture lines and 

the umbonal areas are somewhat raised.

Genus HETEROSTEGINA D'Orhigny, 1826 

Heterostegina suborbicularis D'Orhigny 

Plate 82, figures 7, 8; plate 87, figure 2

Heterostegina suborbicularis D'Orbigny, Annales sci. nat., tome 7,

p. 305, no 1, 1826. 

Fornasini, Soc. geol. italiana Boll., v. 22, fa«c. 3, p. 2 (396),

pi. 14, figs. 5-7, 1903. 

Hofker, Sz&ogra-Exped., Mon. t, p. 70, pi. 35; pi. 36, figs. 3,

6-12, 1927; Resultats Sci. Voyage Indes Orientales Neerlandaises, v. 2, fasc. 1, p. 6, pi. 1, figs. 11, 12, 1930; Vidensk.

Medd. fra Dansk naturh. Foren., v. 93, 1932, p. 153, 1933. 

Cushman, U. S. Natl. Mus. Bull. 161, pt. 2, p. 58, pi. 17,

figs. 6a, b, 1933. 

Heterostegina curva Moebius, Foraminiferen von Mauritius,

p. 105, pi. 13, figs. 1-6, 1880. 

Egger, K. bayer. Akad. Wiss., Math.-naturh. Abt., Abh., Kl. 2,

Band 18, p. 434, pi. 20, figs. 26-33, 1893. 

Cushman, U. S. Natl. Mus. Bull. 161, pt. 2, p. 59, pi. 17,

figs. 1-5, 1933.

Test large, lenticular in edge view, rapidly flaring in 

side view, central portion much thickened, peripheral 

portion thin, periphery rounded; chambers distinct in 

some specimens, indistinct in others, tecoming more 

indistinct toward the middle portion on account of the 

thickening of the test, divided into chamberlets progressively from the peripheral end of each chamber; 

sutures very strongly curved and becoming sigmoid at 

their inner ends, slightly limbate, beconing indistinct 

toward the central part of the test; wall smooth except 

for slightly raised and limbate sutures.

The largest specimen observed in the present material 

is 14 mm in diameter. Several specimens show
TABLE 2.—Distribution ofMiliolidae, Ophthalmidiidae, Trochamminidae, Lagenidae, Polymorphinidae, and Nonionidae, in the Marshall Islands.

Family Millolidae (continued)

Triloculina cf. T. bassensis

cf. T. bicarinata

earlandi n. sp.

involuta n. sp.

irregular Is

kerimbatica

marshallana n. sp.

cf. T. oblonga

spinata

subplanciana n. sp.

terquemiana

tricar inata

trigonula

bikiniensJs n. sp.

sp. A

sp. B

Pyrgo denticulata

denticulata var. striolata

lucernula

millettii

murrhina

sp.

Family Ophthalmidiidae

Cornuspira planorbis

Vertebralina striata

Planispirina exigua

Wiesnerella auriculata

Nubecularia lacunensis

Parrina bradyi

Family Trochamminidae

Trochammina cf. T. rotaliformis

Rotaliammina mayori

Carterina spiculotesta

Nouria polymorphinoides

Family Lagenidae

Robulus calcar

fiibbus

limbosus

nigriseptus

sp. A

sp. B

sp. C

sp. D

Astacolus bradyi

planulata

Dentalina cf. D. filiformis

Nodosaria pauciloculata

sp. A

sp. B

sp. (J

Frondicularia sp. A of Boomgaart

Lagena acuticosta

fllobbsa

gracilis

spiralis

striato-punctata

williamsoni

Family Polymorphinidae

Dimorphina tuberosa

Sigmomorphina semitecta var. terquemiana

Polymorphinella pacifica

Family Nonionidae

Nonion pacificum

potnpilioides

Astronomon tutnidutn

Nonionella translucens

sp. A

sp. B

Elphidium advenutn

advenum var. dispar n. var.

jenseni

milletti

simplex

RONGERIK LAGOON

OJ 

OS

5

1

3

n

o

0

1

2

16

2

3

13

3

4

to

1

1

13

1

3

10

4

7

2

2

1

4

4

3

4

17

9

Tf

1

2

11

2

3

5

1

2

14

2

S

4

14

en

1

1

2

3

4

«

1

21

21

b

1

2

1

t

2

4

5

co

12

4

1

1

1

3

12

13

1

£

1

1

13

8

7

1

2

5

3

4

4

2

10

1

6

7

J

4

I

2

1

21

13

1

8

2

4

7

3

3

3

S

9

19

1

1

1

1

4

4

o

4

1

14

9

5

1

fa

2

1

1

7

I

8

in

2

1

9

2

1

2

8

3

3

3

9

t

2

2

3

2

16

1

1?

ent15

3

1

fa

fa

2

1

1

1

1

10

7

1

6

1

RONGELAP ATOLL

Inside the lagoon

Tf 

tn

CO co

CO

Tf

10

4

15

3

23

9

,'0

1

5

2

4

4

7

3

16

1

6

3

3

1

5

3

18

5

3

21

2

2

5

17

9

2

S

1

/

CO

Tf

3

9

2

4

1

2

1

3

4

1

Tf

CO*

/

1

3

13

10

8

4

3

1

1

1

7

1

3

7

4

c8

12

4

2

1

1

2

4

1

?

0 

COTf

1

1

1

8

5

1

1

3

7

4

6

6

CO

21

2

2

1

10

4

a

i

20

4

10

2

1

5

6

6

S

1

4

3

22

2

1

8

2

1

10

3

t

6

14

£

i

i

3

10

6

5

1

8

5

5

3

7

5

1

1

11

o

1

3

£

Outside 

the 

lagoon

CO

Tf

1-

19

10

1

24

7

5

1

3

3

8

1

4

CO 

CO•*

1

2

1

5

1

2

7

4

1

1

2

Tf

3

2

2

2

1

1

1

1

2

4

1

1

1

1

1

4

1

n

CO•*

1

2

1

1

BIKINI ATOLL

Beach

K

3

2

o

n

in

CQ

o

n

i

i

•*

CO

Reef flat

co 

n

n

i

2

i(J 

CO

2

3

1

2

1

n

o

n

.CO

w

i

i

CO

n

o

CO

n

Inside the lagoon

5

7

2

5

11

4

23

1

4

4

4

4

1

S

2

3

S

2

6

1

2

6

1

3

1

1

1

Tf

10

4

3

2

8

3

1

2

5

1

2

2

o

n

2

2

1

4

2

1

8

2

1

1

1

COt-•*

9

2

2

1

2

3

1

1

2

1

2

CO

o

4

1

I

1

3

2

1

1

1

en•*

2

1

3

1

m 

«

1

2

1

2

4

2

2

o 

en

1

1

9

3

1

1

Tf

n

i

i

i

i

2

2

2

7

2

Tf

n

12

3

4

1

5

2

1

1

1

4

2

10

4

CO

2

1

1

12

2

1

2

3

2

3

1

6 •*

3

1

9

1

1

4

CO

3

4

2

2

3

1

2

1

•<f

1

1

2

1

1

3

3

2

2

cCO

4

5

14

1

7

4

1

3

2

1

CO 

OJ

8

4

1

1

1

1

1

4

n

o

•*

4

1

4

1

2

2

2

1

2

4

CO

1

5

2

5

4

3

5

c- 

[>

4

3

1

2

8

3

2

3

1

5

2

7

2

2

1

1

1

4

6

CD

2

15

10

1

8

3

1

1

1

1

10

10

in 

o

5

3

3

2

1

2

1

3

1

1

3

2

o

1

1

2

?

1

2

o

11

1

5

1

1

7

10

11

m

S

I

n

o

1

8

20

1

14

8

3

6

8

2

4

1

10

5

tm

4

1

3

2

2

10

6

1

2

1

3

6

?2

11

CO

o

20

1

12

1

6

1

5

1

1

2

1

9

1

t

o

Tf

3

1

1

CO

Tf

1

2

Tf 

Tf 

Tf

CO

2

1

7

4

1

3

1

to

10

in 

o

1

5

Tf in

3

6

1

1

1

12

tfl

o

CO

16

4

19

3

5

6

2

5

14

4

CO 

CC•*

3

3

1

4

2

1

?

Tf

6

7

3

1

1

2

1

4

2

2

•i

6

to

3

4

1

1

3

1

CO

6

4

4

18

16

1

1

8

5

3

16

4

%f 

o

2

1

1

6

1

3

3

1

5

7

1

1

2

t— 1

2

5

9

14

2

2

3

6

1

1

2

5

2

1

1

3

o

S

5

1

6

8

1

1

9

1

6

14

7

CO

o

2

1

3

2

1

1

1

1

3

•* »

7

1

2

1

1

tc1

1

1

1

1

1

2

1

n

Tf

3

2

2

1

5

1

•*

1

1

1

en

7

5

13

2

2

2

3

4

1

13

3

CO•* •*

1

1

2

1

5

en

3

6

8

3

5

7

4

5

1

I

?

2

in

Tf

OJ

1

53in

1

1

2

2

3

2

2

2

2

1

1

CO 

enm

1

S

1

1

2

1

1

2

*

Tf

3

1

1

1

Outer slopes

Dredge 

samples

CO C- •*

9

1

3

2

1

2

1

1

6

1

1

2

1

1

CO

Tf

1

1

1

3

1

21

2

4

2

1

1

o

Tf

2

3

1

1

2

1

2

Bottom samples

,

2

2

2

4

1

2

1

5

2

n

1

1

CO

4

2

2

1

1

1

1

1

3

3

3

5

1

1

1

1

1

1

1

2

•*

4

2

8

2

3

5

2

6

1

1

2

1

1

1

1

1

1

3

1

4

1

2

1

2

CO

3

2

2

1

1

4

1

o

CO

1

2

1

2

2

1

CO

1

2

1

1

2

J

CO

1

2

1

1

t> ~ai

COt^

en 

CO

2

3

2

1

4

1

2

1

1

1

1

1

1

1

1

1

2

2

3

CO

1

2

1

1

2

2

3

1

1

2

1

3

2

1

1

1

2

6

1

o

CO

Guyot

^ 

c2

2

6

4

1

3

3

6

11

2

2

1

5

2

1

1

1

2

4

3

7

4

3

3

Tf 

Tf

C1

1

1

13

4

2

2

10

1

2

Tf

1

4

4

1

9

3

1

1

1

1

9

"b

Tf

3

1

1

1

3

2

1

3

1

1

1174(10-14")

3

3

2

1

3

1

1

1

2

2

2

1

2

3

1

2

1

6

1

t>

2

2

1

1

1

1

2

1

1

1

1

2

1

5

1

2

o

5

i

i

2

1

2

2

3

2

1

1

1

2

2

3

3

m 

c^

1

2

2

2

1

4

1

4

1

1

2

1

2

4

1

2

n

in 

c3

1

6

1

2

2

2

1

1

1

1

3

2

2

%f

1

2

2

1

4

1

1

1

1

1

3

2

1

£• 

c3

4

2

3

2

1

2

1

o

CO 

C1

1

3

1

1

2

1

1

1

eg

1

5

1

2

t1

1

1

m

1

1

2

1

1

1

4

1

1

1

1

1

a.

CO

1

o 

o

C1

5

1

1

1

'«

o

C1

Vf

CO

o

C2

1

CO

Tf

o

3

1

eg

cENIWETOK ATOLL

Inside the lagoon

o

1

1

2

=!

1

2

o

S

1

4

1

5

1

1

CO

n

1

1

3

2

CO 

CO

n

CO

4

1

5

CO 

33

2

2

1

1

3

1

CO

1

1

2

1

3

1

2

1

3

11

1

2

1

9

2

7

6

n

19

5

2

1

1

6

8

7

00

1

3

4

1

3

1

1

2

3

o

OJ 

M

4

6

2

1

5

O} 

<M

1

12

3

1

2

1

1

3

5

5

en

11

2

4

18

1

3

2

2

7

2

3

3

5

1

4

18

fa

1

S

10

2

5

4

2

5

3

2

10

1

7

16

6

M

3J

2

1

1

2

2

CO

en *

4

4

2

3

1

5

«

23

1

4

5

2

2

3

4

2

4

6

2

J

2

1

«

8

11

2

4

2

?

2

5

4

1

1

1

16

en

CO

9

4

1

1

5

en 

eg

8

2

4

2

2

4

2

fa

2

n

5

5

fa

10

3

1

7

5

7

3

2

13

4

10

ig

CO

OJ

1

3

1

1

2

s

1

7

J

12

1

8

21

5

2

1

10

1

15

5

8

S

10

2

7

23

1

3

7

10

6

1

3

1

o

5

5

3

fa

3

5

1

1

1

1

2

15

1

8

2

CO

1

2

8

9

1

3

1

2

3

2

7

3

2

a

8

i

12

1

4

1

6

4

1

2

18

o 

eg

CO

3

1

11

14

3

1

17

3

2

1

3

4

1

8

1

•«f

N

2

2

3

fa

fa

o 

o

CO

2

17

13

4

3

7

5

1

1

fa

I

fa

13

10

6

CO

?

P

2

1

2

2

1

18

4

11

eg

o

1

2

1

1

o

?

1

2C

9

2

1

4

3

1

7

/

2

S

CO

1

10

4

10

4

6

1

1

fa

1

6

1

4

13

a

i

5

*

fa

1

•*

10

8

8

1

fa

3

1

2

1

3

Ifa

CO

CO

2

2

2

3

3

2

1

CC

5

4

fa

5

fa

1

1

1

3

13

b

11

in

1

1

2

2

1

1

o

1

Tf

O

1

16

15

1

8

2

5

3

1

5

1

1

9

1

4

s

1

6

6

1

4

1

1

2

55

6

12

2

2

1

1

1

1

8

1

4

CO 

CO

1

4

5

4

7

7

5

3

2

1

11

/

TJOuter 

slopes

S

6

4

18

16

10

1

1

6

1

1

2

fa

5

^

9

2

15

1

2

2

4

2

1

2

2

1

3

2

S

1

1

1

1

4

1

3

1

S3

1

4

2

1

1

1

2

1

1 2

1

1

1

1

1

2

1

12

10

Z91053 O - 54 (Face p. 346)
RECENT FORAMINIFERA OF THE MARSHALL ISLANDS 347

evidence of breaking and then regrowth with larger 

chambers.

It is difficult to separate the two forms called H. 

suborbicularis and H. curva as their distinction seems 

to be an arbitrary rather than a natural one. It is, 

therefore, believed that only one species of Heterostegina 

is represented in the present material.

The types of H. suborbicularis were from the Hawaiian 

Islands. The very thick umbonate forms with a more 

opaque wall have been included under this specific 

name.

The types of H. curva were from off Mauritius. The 

flatter forms with the chambers visible te the center of 

the test have been assigned to this species.

It is possible that some forms we have included here 

are the same as forms referred to Heterostegina depressa 

D'Orbigny (Cushman, 1933, p. 57, pi. 16, figs. 5-9 

(not fig. 4) but these do not appear to be typical of 

H. depressa according to the type figure.

Genus CYCLOCLYPEUS W. B. Carpenter, 1856

Cycloclypeus carpenter! H. B. Brady

Plate 82, figure 9

Cycloclypeus W. B. Carpenter, Philosophical Trans., v. 146,

p. 555, pi. 30, figs. 1, 3, 1856; Introd. Foraminifera, p. 292,

pi. 19,-figs. 2-7, 1862. 

Tan, Wetenschappelijke Mededeelingen no. 19, Dienst

Mijnbouw Nederlandsch-Indie, p. 81, 82, 1932. 

Cycloclypeus carpenteri H. B. Brady, Quart. Jour. Micros. Sci.,

v. 21, p. 67, 1881; Challenger Kept., Zoology, v. 9, p. 751,

1884. 

Chapman, Linnean Soc. Jour., Zoology, v. 28, p. 22, pi. 2,

figs. 6, 7; pi. 3, figs. 1-5, 1900. 

Hofker, Siboga-Exped., Mon. 4, p. 71, pi. 24, fig. 1; pi. 37;

pi. 38, figs. 1-9, 12, 13, 1927. 

Cycloclypeus guembelianus H. B. Brady, Quart. Jour. Micros.

Sci., v. 21, p. 66, 1881; Challenger Kept., Zoology, v. 9,

p. 751, pi. Ill, figs. 8a, b (young form), 1884. 

Hanzawa, Short Papers Instit. Geol. Pal., Tohoku Univ.,

Sendai, no. 3, p. 1, pis. 1, 2, text figs. 1-10, 1951.

This species, described from off the coast of Borneo 

and off Fiji, occurs rarely in several deep-water samples 

at Bikini and Rongelap, and very abundantly in a 

dredging from off Bikini. The microspheric and megalospheric forms of the species are quite distinct in size and 

appearance; the megalospheric ones having been 

described as C. guembelianus. The megalospheric specimens are 15 to 18 mm. in diameter, papillate along the 

sutures, and rather translucent except in'the central 

thickened portion. The microspheric specimeas are 

much larger, up to 48 mm in diameter, smooth, thickwalled and not translucent, with the chambers less

distinctly visible, and the periphery often slightly 

ruffled.

In a sample dredged from 800 to 580 feet, off Bikini, 

11°38'33" N., 165°31'18" E., abundant specimens of 

both forms were obtamed. Some of this material preserved in neutralized formalin was tested for protoplasm by the rose bengal test and gave a positive rer<> 

tion. The megalospheric forms showed the protoplarrn 

present in the central, thickened portion only. The 

test was not conclusive for the microspheric forms inasmuch as the surface of the shell substance, even in obviously dead specimens, appeared to take the stain in 

varying degrees.

Examination by Dr. Margaret Jepps (1953, p. 1114), 

including decalcification of additional preserved material from the same dredging sample, disclosed the 

internal structure and caaal system of the shells as well 

as proved the existence of multinuclear protoplarm 

within a microspheric specimen.

All the large, microspheric specimens obtained were 

found to bear various types of attached organisms: 

bryozoans, worm tubes, and Forarninifera; and a few 

specimens showed pin-prick holes, the results of the 

activity of the boring sponge Cliona (Chapman, 1900, 

p. 24).

Among the Foraminifera found attached to Cycloclypeus is an unidentified form that leaves shallow p; ts 

from 0.5 to 2 mm in diameter thickly scattered over the 

surface. A few pits were worn deeply enough into the 

shell substance as to give access to the interior of the 

chambers of the Cycloclypeus, Eight specimens of 

these pitting organisms were found attached to the 

surface of one specimen, but, although the results of 

then* activities could be observed on several of the larger 

microspheric individuals, no more specimens we^e 

seen. The test is irregular in outline, flat with a ttin 

wall on the attached side, convex but flattened on the 

exposed side. The exposed surface is smooth, granular, 

and ornamented with a few glassy papillae apparently 

without any definite pattern. The wall is calcareous 

and appears to be finely perforate. Internally the test 

consists of a large central chamber surrounded by tv^o 

or more crescent-shaped chambers. No aperture was 

observed. Sutures between the chambers are rot 

obvious from the exposed surface of the test but m*».y 

be seen when the test is moistened. The affinities of 

this form are not revealed by the available material. 

Further material may show its family and generic 

relationships and whether it feeds upon other Foraminifera.

291053—54
348 BIKINI AND NEARBY ATOLLS, MARSHALL ISLANDS

Family PENEROPLIDAE 

Genus PENEROPLIS Montfort, 1808

Peneroplis sp. 

Plate 87, figure 3

A few specimens from Rongelap appear to represent 

an undescribed species. The test is strongly compressed, slightly umbilicate; the chambers 9 to 11 in the 

last whorl; the wall translucent and finely pitted; the 

apertures multiple on the face of the last-formed 

chamber.

Genus SPIROLINA Lamarck, 1804

Spirolina acicularis (Batsch) 

Plate 87, figure 6

Naidilus (Lituus) acicularis Batsch, Conch. Seesandes, p. 4, pi. 6,

fig. 16, 1791. 

Spirolina acicularis Cushman, U. S. Natl. Mus. Bull. 104, pt. 7,

p. 42, pi. 15, figs. 1-3, 1930; idem, Bull. 161, pt. 2, p. 63,

pi. 19, figs. 6, 7, 1933.

Very rare, small specimens, may be referred to this 

species which has been recorded from various localities 

in shallow water of the Pacific Islands and the Tropical 

Atlantic. The types are from the Mediterranean. 

One fragmentary specimen shows as many as 20 uniserial chambers.

Spirolina arietina (Batsch)

Plate 87, figures 4, 5

Nautilus (Lituus) arietinus, Batsch (in part), Conch. Seesandes,

p. 4, pi. 6, fig. 15c, 1791. 

Peneroplis arietinus Parker, Jones, and H. B. Brady, Annals and

Mag. Nat. History, 3rd ser. v. 16, p. 26, pi. 1, fig. 18, 1865. 

H. B. Brady, Challenger Kept., Zoology, v. 9, p. 204, pi. 13,

figs. 18, 19, 22, 1884. 

Spirolina arietina Cushman, U. S. Natl. Mus. Bull. 104, pt. 7,

p. 43, pi. 15, figs. 4, 5, 1930; idem, Bull. 161, pt. 2; p. 62,

pi. 19, figs. 4, 5, 1933.

This is a widely distributed species in the warm shallow waters of tropical seas. It is common in this 

material.

Genus MONALYSIDIUM Chapman, 1900

Monalysidium politum Chapman

Plate 87, figure 7

Peneroplis (Monalysidium) pohta Chapman, Linnean Soc. Jour.,

Zoology, v. 28, p. 4, pi. 1, fig. 5 (?), 1900. 

Monalysidium politum Heron-Alien and Earland, Zool. Soc.

London Trans., v. 20, p. 603, fig. 43G (in text), 1915. 

Cushman, U. S. Natl. Mus. Bull. 161, pt. 2, p. 63, pi. 19,

figs. 8, 9, 1933.

Very rare specimens in this material are referred to 

this widely recorded species. They are all fragmentary.

Some show faint striae as the fragments of Spirolina 

acicularis do, but in this species the chambers are more 

distinctly separated by incised sutures.

Genus SORITES Ehrenberg, 1840

Sorites marginalis (Lamarck)

Plate 82, figure 4

Orbulites marginalis Lamarck, Syst. Anim. sans Vert., tome 2, p.

196, no. 1, 1816. 

Orbitolites marginalis W. B. Carpenter, Philosophical Trans.,

v. 174, p. 560, fig. 1 (in text), 1883; Challenger Kept., Zoology,

v. 7, "Orbitolites," p. 20, pi. 3, figs. 1-7; pi. 4, figs. 1-5, 1883. 

H. B. Brady,^Challenger Rept., Zoology, v. 9, p. 214, pi. 15,

figs. 1-5, 1884. 

Sorites marginalis Cushman, U. S. Natl. Mus. Bull. 104, pt. 7,

p. 49, pi. 18, figs. 1-4, 1930.

This species is not common in this material and is 

mostly represented by immature specimens.

Genus MARGINOPORA Blainville, J830

Marginopora vertebralis Blainville

Plate 82, figures 5, 6

Marginopora vertebralis Blainville, Manuel d'Actinologie ou de

Zoophytologie, p. 412, pi. 69, fig. 6, 1834. 

Van der Vlerk, Wetenschappelijke Meded?elingen, no. 1,

p. 11, pi. 4, figs. 14, 15, 1924. 

Hofker, Siboga-Exped., Mon. 4a, p. 160, pi. 57, figs. 1, 2; pi.

61, figs. 4, 5, 11; pi. 62 figs. 1-9, 11, 12, 19°0. 

Cushman, U. S. Natl. Mus. Bull. 161, pt. 2, p. 67, pi. 19, figs.

11, 12, 1933. 

Amphisorus hemprichii Ehrenberg, Akad. Wiss. Berlin Abhandl.,

1838, p. 134, pi. 3, fig. 3, 1840.

We are following W. Storrs Cole in placing together 

Amphisorus and Marginopora, considering the former as 

immature specimens of the latter. The thicker, more 

robust specimens with depressed centers are very abundant in very shallow water and on the reef flats. They 

are evidently strong enough to resist wave action to 

some extent. The thinner, two-chambers-thick specimens with more translucent walls, those formerly 

referred to Amphisorus, are found with the thicker ones 

and, in addition, occur in deeper water.

Family ALVEOLINELLIDAE

Genus NEOALVEOLINA Silvestri, 1928

Neoalveolina pulchra (D'Orbigny)

Plate 87, figure 8

Alveolina pulchra D'Orbigny, in De la Sagra, Histoire physique, 

politique et naturelle de File de Cuba, Foraminiferes, p. 70, 

pi. 8, figs. 19, 20, 1839.

Neoalveolina pulchra Reichel, Soc. Pal. Suisse Mem., v. 58, p. 95, 

1937.

Rare specimens occur at all the atolls.
RECENT FOKAMINIPEKA OF THE MARSHALL ISLANDS 349

Family HETEROHELICIDAE

Genus GUMBELINA Egger, 1899

Gumbelina? marshallana Todd, n. sp.

Plate 87, figure 9

Test small, slender, somewhat compressed, periphery 

rounded, very slightly indented; chambers few, distinct, 

6 to 8 pairs comprising the test, rapidly increasing in 

size as added; sutures distinct, depressed; wall finely 

perforate; aperture a high, narrow, slit extending from 

the suture into the face of the last-formed chamber. 

Length 0.23 to 0.30 mm, breadth 0.13 to 0.15 mm, 

thickness 0.07 to 0.10 mm.

Holotype (USNM 548727) from core on Sylvania 

Guyot, bottom of the core at 5 to 7 inches, 720 fathoms, 

11°44'47" N., 165°09'56" E. (Bik 1176(5"-7")).

This species closely resembles G. cubensis Palmer 

(1934, p. 74, text figs. 1-6) from the Oligocene of Cuba 

and adjacent areas but differs in its slenderer test, 

slightly less inflated chambers, and its high aperture, 

elongated in the plane of the test.

This genus has not previously been known from 

sediments younger than Oligocene. This form is 

questionably placed in this genus because of the type 

of aperture which resembles that characteristic of 

Bolivina, although it is not unknown in the genus 

Gumbelina.

This species may have its origin in fossil, rather than 

Recent, sediments in a manner suggested in the discussion of the unique faunal composition of the core, 

Bikini 1176, whereby a submarine outcrop of Tertiary 

sediments higher on the atoll slope might have shed 

debris down the slope of the atoll.

Genus BOLIVINITA Cushman, 1927 

Bolivinita quadrilatera (Schwager)

Textularia quadrilatera Schwager, Mwara-Exped., Geol. Theil,

Band 2, p. 253, pi. 7, fig. 103, 1866. 

H. B. Brady, Challenger Kept., Zoology, v. 9, p. 358, pi. 42,

figs. 8-12, 1884. 

Bolivinita quadrilatera Cushman, B. P. Bishop Mus. Bull. 119,

p. 121, pi. 14, figs. 12a, b, 1934; U. S. Natl. Mus. Bull. 161,

pt. 3, p. 2, pi. 1, 1942.

A few specimens, some as long as 1.15 mm, occur in 

three deep-water samples off Bikini.

Genus BOLIYINELLA Cushman, 1927

Bolivinella folia (Parker and Jones)

Plate 87, figures 10, 11

Textularia folia Parker and Jon$3, Philosophical Trans., v. 155,

p. 370, 420, pi. 18, fig. 19, 1865. 

H. B. Brady (in part), Challenger Kept., Zoology, v. 9, p. 357,

pi. 42, figs. 3-5 (not 1, 2), 1884. 

Bolivinella folia Cushman Lab. Foram. Research Contr., v. 2,

pt. 4, p. 79, 1927; U. S. Natl. Mus. Bull. 161, pt. 3, p. 3,

pi. 2, figs. 1-4, 6, 1942.

This is a characteristic species of warm shallow waters 

of the Pacific.

Bolivinella folia var. ornata Cushman 

Plate 87, figures 12, 13

Bolivinella folia (Parker and Jones) var. ornata Cushman, 

Cushman Lab. Foram. Research Contr., v. 5, p. 32, pi. 5, 

figs. 3, 4, 1929; U. S. Natl. Mus. Bull. 161, pt. 3, p. 5, pi. 2, 

figs. 5, 8, 1942.

The types of this variety were from shore sand of 

Hardwicke Bay, Australia, and the only other recorded 

occurrence is from 12 fathoms, off Levuka, Fiji. It is 

of interest that it has been found in a number of samples 

in this material.

In addition to the peripheral spines this variety 

differs in being thicker in the middle, while the typical 

form is equally compressed throughout.

Family BTJLIMINIDAE 

Genus BULIMINELLA Cushman, 1911 

Buliminella elegantissima (D'Orbigny)

Bulimina elegantissima D'Orbigny, Voyage dans TAme'rique 

me'ridionale, tome 5, pt. 5, Foraminiferes, p. 51, pi. 7, 

figs. 13, 14, 1839.

Buliminella elegantissima Cushman and Parker, U. S. Geol. 

Survey Prof. Paper 210-D, p. 67, pi. 17, figs. 10-12, 1947.

A single small but typical specimen of this cosm®- 

politan species was found at Eniwetok 329.

Buliminella milletti Cushman 

Plate 87, figures 11, 15

Buliminella milletti Cushman, Cushman Lab. Foram. Research 

Contr., v. 9, p. 78, pi. 8, figs. 5, 6, 1933; U. S. Natl. Mus. 

Bull. 161, pt. 3, p. 7, pi. 3, figs. 1-4, 1942.

The types of this species are from Mokaujar AncHorage, Fiji, and it is recorded from numeious statiois in 

shallow warm waters of the Pacific and also in the 

Mediterranean and the West Indian region of the 

Atlantic. A few of the specimens have a pointed initial 

end resembling B. madagascariensis (D'Orbigny), var. 

spicata Cushman and Parker (1947, p. 64, pi. 16, 

fig. 20) but not the distinct basal spine characteristic 

of that variety. With these are very rare specimens that 

tend toward B. parallela Cushman and Parker (1947, 

p. 69, pi. 17, fig. 22) but not enough to separate them 

definitely.

Genus BULIMINOIDES Cushman, 1911

Buliminoides williamsoniana (H. B. Brady)

Plate 87, figures 16, 17

Bulimina williamsoniana H. B. Brady, Quart. Jour. Micros.

Sci., v. 21, p. 56, 1881; Challenger Rept., Zoology, v. 9,

p. 408, pi. 51, figs. 16, 17, 1884. 

Buliminoides williamsoniana Cushman, U. S. Natl. Mus. Bull.

71, pt. 2, p. 90, fig; 144 (in text), 1911; idem, Bull. 161, pt. 3,.

p. 8, pi. 3, figs. 7-9, 1942.

This is a widely distributed species in the Indo-
350 BIKINI AND NEARBY ATOLLS, MARSHALL ISLANDS

Pacific region and occurs also in the West Indian region 

of the Atlantic.

Genus ROBERTINA D'Orbigny, 1846

Robertina subcylindrica (H, B. Brady)

Plate 87, figure 18

Bulimina subcylindrica H. B. Brady, Quart. Jour. Micros. Sci.,

v. 21, p. 56, 1881; Chalknger Rept., Zoology, v. 9, p. 404,

pi. 50, figs. 16a, b, 1884. 

Robertina subcylindrica Cushman and Parker, Cushman Lab.

Foram. Research Contr., v. 12, p. 95, pi. 16, figs. lOa, b,

1936. 

Cushman, U. S. Natl. Mus. Bull. 161, pt. 3, p. 9, pi. 3, fig. 14,

1942.

This species occurs rarely in the core at Bikini 1176. 

Robertina translucens Cushman and Parker 

Plate 87, figure 19

Robertina translucens Cushman and Parker, Cushman Lab. 

Foram. Research Contr., v. 12, p. 99, pi. 16, figs. 8a, b, 1936, 

U. S. Geol. Survey Prof. Paper 210-D, p. 75, pi. 18, figs. 

15a, b, 1947.

Two typical specimens were found at Bikini 1174.

Genus PSEUDOBULIMINA Earland, 1934 

Pseudobulimina sp. 

Plate 87, figure 20

A few specimens from Bikini and Rongelap, insufficient for complete description, seem to represent 

an undescribed species. The specimens are quite 

strongly compressed and broader in proportion to 

length than those referred to Pseudobulimina convoluta 

(Williamson) (Said, 1949, p. 26, pi. 4, fig. 4), although 

the available specimens show a considerable degree of 

variation in shape.

Genus BULIMINA D'Orbigny, 1826

Bulimina aculeata D'Orbigny

Plate 87, figure 21

Bulimina aculeata D'Orbigny, Annales sci. nat., tome 7, p. 269,

no. 7, 1826.

Cushman and Parker, U. S. Geol. Survey Prof. Paper 210-D, 

p. 120, pi. 28, figs. 8-11, 1947.

This widely distributed species occurs at several of 

the deep-water Bikini stations.

Bulimina affinis D'Orbigny 

Plate 87, figure 22

Bulimina affinis D'Orbigny, in De la Sagra, Histoire physique, 

politique et naturelle de 1'Ile de Cuba, Foraminiferes, 

tome 6, p. 105, pi. 2, figs. 25, 26, 1839.

Cushman and Parker, U. S. Geol. Survey Prof. Paper 210-D, 

p. 122, pi. 28, figs. 23-25, 1947.

A few specimens were found in the core at Bikini 1176.

Bulimina costata D'Orbigny 

Plate 87, figure 23

Bulimina costata D'Orbigny, Annales sci. nat., tome 7, p. 269,

no. 1, 1826.

Cushman and Parker, U. S. Geol. Survey Prof. Paper 210-D, 

p. 115, pi. 27, figs. 2, 3, 1947.

This thin-walled, coarsely punctate species occurs in 

several deep-water samples at Bikini. The present 

specimens have been compared with specimens from 

the Pliocene of Italy and resemble them cT osely.

Bulimina fijiensis Cushman 

Plate 87, figure 25

Bulimina fijiensis Cushman Lab. Foram. Research Contr., v. 9, 

p. 79, pi. 8, figs. 7a-c, 1933; U. S. Natl. Mus. Bull. 161, 

pt. 3, p. 11, pi. 3, figs. 10, 11, 1942.

The types are from 12 fathoms, Nairai, Fiji, and it 

is also recorded from 12 fathoms, Levuka, Fiji, and 

Tom Zanzibar. It occurs commonly at all four atolls.

Bulimina rostrata H. B. Brady 

Plate 87, figure 24

Bulimina rostrata H. B. Brady, Challenger, Rept., Zoology, v. 9, 

p. 408, pi. 51, figs. 14, 15, 1884.

This species, described from 428 father's off the Ki 

Islands, occurs at several deep-water stations at 

Bikini.

Genus OOLINA D'Orbigny, 1839

Oolina squamosa (Montagu) 

Plate 87, figure 26

Vermiculum squamosum Montagu, Testacea Britannica, p. 526 

pi. 14, fig. 2, 1803.

Entosolenia squamosa Williamson, Annals and Mag. Nat. 

History, 2d ser., v. 1, p. 18, pi. 2, fig. 19, 1848; Recent 

Foraminifera of Great Britain, p. 12, pi. 1, fT. 29, 1858.

Rare but characteristic specimens of this widely 

known species were found at Bikini ari Rongelap. 

With these was found a single specimen of 0. catenulata 

(Williamson) (1848, p. 19, pi. 2, fig. 20) which differs 

in that the cells of the reticulate ornamentation are 

arranged in longitudinal columns whereas in 0. squamosa 

no alignment of the cells is noticeable.

Genus FISSTJRINA Reuss, 1850 

Fissurina auriculata (H. B. Brady) var. costata (H. B. Brady)

Lagena auriculata var. costata H. B. Brady, QueH. Jour. Micr. 

Sci., v. 21, p. 61, 1881; Challenger Rept., Zoology, v. 9r 

p. 487, pi. 60, fig. 38, 1884.

Cushman, U. S. Natl. Mus. Bull. 161, pt. 2, p. 23, pi. 6, fig. 2, 

1933.

A single specimen from Bikini 1175 at 5^2 fathoms is 

the same as specimens from deep water in the Tropical 

Pacific.
RECENT FOBAMINIFEBA OF THE MARSHALL ISLANDS 351

Fissurina circularis Todd, n. sp. 

Plate 87, figure 27

Lagena marginata Cushman (part) (not Montagu), U. S. Natl. 

Mus. Bull. 161, pt. 2, p. 17, pi. 4, figs. 11, 14; pi. 5, figs. 4, 

6, 8, 9 (not pi. 4, figs. 9, 12, 15, 16; pi. 5, fig. 2), 1933.

Test small for the genus, compressed, very slightly 

longer than broad, apertural end not extending out from 

the circular outline of the test, periphery acute, sometimes slightly keeled; wall thin, transparent in the 

central portion, with an opaque band around the 

periphery giving the appearance of a broad keel, sometimes both sides of the test are ornamented by faint 

ridges of shell material just inside and parallel to the 

periphery; aperture an elongate slit, without a lip, 

connecting with the interior by an elongate tube extending half or more of the length of the test. Diameter 

0.25 to 0.30 mm, thickness 0.12 mm.

Holotype (USNM 548767) from Eniwetok lagoon, 24 

fathoms, 11°31'15" N., 162°20'27" E. (Eni 326).

This species differs from F. marginata (Walker and 

Boys) (1784, p. 2, pi. 1, fig. 7) in its circular form and 

its aperture not projecting out from the outline of the 

test. It is fairly widespread in the Marshall Islands 

material.

Fissurina formosa (Schwager)

Plate 87, figure 31

Lagena formosa Schwager (in part), Afoflara-Exped., Geol. Theil,

v. 2, p. 206, pi. 4, figs. 19a, 19d (not 19b, 19c), 1866. 

H. B. Brady, Challenger Rept., Zoology, v. 9, p. 480, pi. 60, 

figs. 10, 18-20, 1884.

Entosolenia formosa Cushman, Foraminifera, 4th Ed., Key, pi. 

21, fig. 22, 1948.

A few specimens showing considerable variation in 

shape and size, occur rather rarely at Rongerik, Bikini, 

and Eniwetok.

Fissurina lacunata (Burrows and Holland) 

Plate 87, figure 28

Lagena lacunata Burrows and Holland, (in Jones), Paleont. Soc.,

1895, p. 205, pi. 7, fig. 12. 

Lagena orbignyana var. lacunata Sidebottom, Manchester Lit.

Philos. Soc. Mem. and Proc., v. 54, no. 16, p. 19, pi. 2, fig.

14, 1910. 

Cushman, U. S. Natl. Mus. Bull. 161, pt. 2, p. 27, pi. 7, figs.

1-5, 8, 1933. 

Fissurina lacunata Parr, Royal Soc. Victoria Proc., v. 56 (n.

ser.), pt. 2, p. 203, 1945. 

Lagena castrensis H. B. Brady (not Schwager), Challenger Rept.,

Zoology, v. 9, p. 485, pi. 60, figs. 1, 2, 1884. 

Lagena orbignyana var. castrensis Millett, Royal Micros. Soc.

Jour., 1901, p. 626, pi. 14, fig. 20.

This small species, triple-keeled on the periphery and 

with the body of the test having a reticulate ornamentation, occurs rarely at all four atolls. It has a more 

definite neck than F. orbignyana (Seguenza) (1862, p.

66, pi. 2, figs. 25, 26) and so seems to be more than 

varietally distinct from that species.

Fissurina milletti Todd, n. sp. 

Plate 87, figure 30

Lagena marginato-perforata Millett (not Seguenza), Royal Micros.

Soc. Jour., 1901, p. 621, pi. 14, fig. 4.

Cushman (part), U. S. Natl. Mus. Bull. 161, pt. 2, p. 19, pi. 5, 

figs, la, b; pi. 6, figs, la, b (not pi. 4, figs. 13a, b), 1933.

Test small for the genus, compressed, about twic^ as 

long as broad, basal end rounded, apertural end extended into a shoit neck with everted lip, periphery rith 

a thickened, blunt keel; wall thin, transparent, ornamented thickly and evenly by small pits which appear 

opaque on the clear wall; aperture compressed, surrounded by a thickened rim, connecting with the interior 

by means of a tube extending about one quarter the 

length of the test. Length 0.25 mm, breadth 0.13 mm, 

thickness 0.10 mm.

Holotype (USNM 548743) from Eniwetok atoll, 500 

fathoms, 11°18'52" N., 162°15'24" E. (Eni 119).

This species differs from F. marginato-perforata (Seguenza) (1880, p. 332, pi. 17, fig. 34) in the finer texture of 

its ornamentation, its narrow, blunt peripheral keel and 

its short neck with backwardly curved lip. It appears 

to be the same as the specimens referred to hi the above 

synonymy; Millett's record being from the Mr,lay 

Archipelago and the other from various localities in the 

Tropical Pacific. In the present material it occurs at 

numerous stations but is nowhere common. It shows 

remarkably little variation.

It somewhat resembles F. perforata (Moebius, 1880, 

p. 90, pi. 8, fig. 6) from off Mauritius but that species 

lacks the definite neck that is characteristic of F. milletti, n. sp.

Fissurina radiato-marginata (Parker and Jones) 

Plate 87, figure 29

Lagena radiato-marginata Parker and Jones, Philosophical

Trans., v. 155, p. 355, pi. 18, fig. 3, 1865. 

H. B. Brady, Challenger Rept., Zoology, v. 9, p. 481, pi, 61,

figs. 8, 9, 1884. 

Cushman, U. S. Natl. Mus. Bull. 161, pt. 2, p. 31, pi. 7, figs.

17, 18, 1933.

Fissurina radiato-marginata Parr, Royal Soc. Victoria Proc., v. 58 

(n. ser.), pts. 1-2, p. 122, pi. 6, fig. 11, 1945 (1947).

This species, described from Australian coral-i n,efs 

and recorded from various localities in the Pacific, is 

represented by two specimens from Bikini 218 core 

(12-16").

Fissurina sp. A

Plate 87, figure 32

Test very small, inflated, globular with a projecting 

apertural neck, periphery with 3 very slight keels, cen-
352 BIKINI AND NEAEBY ATOLLS, MARSHALL ISLANDS

tral body of test translucent so that the internal tube is 

visible. Length 0.18 mm, breadth 0.15 mm, thickness 

0.10 mm.

This small species seems to be undescribed. It occurs 

fairly commonly at Eniwetok and is found rarely at 

Rongelap and Rongerik.

Fissurina sp. B 

Plate 87, figure 33

Lagena orbignyana Cushman (not Seguenza), TJ. S. Natl. MusBull. 161, pt. 2, p. 26. pi. 6, figs. 7, 8, 11, 1933.

A species with a broad peripheral keel and secondary 

keels on either side, and a projecting apertural neck, is 

represented by a few specimens at Bikini and Eniwetok. 

It seems to be different from typical F. orbignyana 

(Seguenza) (1862, p. 66, pi. 2, figs. 25, 26) in its very 

broad keels and projecting neck.

Fissurina sp. C 

Plate 87, figure 34

A species with thickened base showing numerous keels 

and a compressed and pointed apertural end, similar to 

the specimen figured, occurs rarely at Bikini and 

Eniwetok.

Fissurina spp.

In addition to those here recorded, there are several 

other species of Fissurina, represented by too few or 

poor specimens to describe.

Genus VIRGULINA D'Orbigny, 1826

Virgulina earlandi Cushman

Plate 87, figure 35

Virgulina earlandi Cushman, Cushman Lab. Foram. Research

Special Pub. 6, p. 49, pi. 7, figs. 8a-c, 1936; idem, Special

Pub. 9, p. 30, pi. 4, figs. 27, 28, 1937. 

Virgulina schreibersiana Millett (not Czjzek), Royal Micros. Soc.

Jour., 1900, p. 280, pi. 11, figs. 13a, b. 

Heron-Alien and Earland, Zool. Soc. London Trans, v. 20, p.

642, pi. 49, figs. 1-12, 1915.

This species, described from the Kerimba Archipelago 

occurs rarely. Most of the specimens are young and are 

slenderer than paratypes with which they have been 

compared.

Genus BOLIVINA D'Orbigny, 1839

Bolivina abbreviata Heron-Alien and Earland 

Plate 87, figure 36

Bolivina limbata H. B. Brady var. abbreviata Heron-Alien and

Earland, Linnean Soc. Jour., Zoology, v. 35, p. 622, pi. 36,

figs. 25-27, 1924. 

Bolivina abbreviata Cushman, Cushman Lab. Foram. Research

Special Pub. 9, p. 143, pi. 18, figs. 34, 35, 1937; U. S. Natl.

Mus. Bull. 161, pt. 3, p. 33, pi. 9, fig. 5, 1942.

A few specimens were found at Bikini.

Bolivina compacta Sidebottom 

Plate 87, figure 37

Bolivina robusta H. B. Brady var. compacta Sid?bottom, Manchester Lit. Philos. Soc. Mem. and Proc., v. 49, no. 5, p. 15, 

pi. 3, fig. 7,1905.

Bolivina compacta Cushman, U. S. Natl. Mus. Bull. 71, pt. 2, p. 

36, fig. 58, 1911; U. S. Natl. Mus. Bull. 161, pt. 3, p. 20, pi. 7, 

figs 2, 3, 1942.

This is apparently a widely distributed species and 

has been recorded from many localities in the warm, 

shallow waters of the Pacific.

Bolivina globulosa Cushman

Bolivina globulosa Cushman, Cushman Lab. Foram. Research 

Contr., v. 9, p. 80, pi. 8, figs. 9a, b, 1933.

A single specimen of this species described from 2883 

fathoms and recorded from other deep water samples 

from the tropical Pacific, was found at Bikini 1175

(0-1"). 

The measurements as originally given are in error.

The holotype measures 0.30 mm in length and 0.13 mm 

in breadth. The present specimen is slightly larger; 

0.48 mm long and 0.21 mm wide, but shows the same 

highly polished wall surface.

Bolivina nitida H. B. Brady 

Plate 87, figure 38

Bolivina nitida H. B. Brady, Challenger Rept., Zoology, v. 9, p.

420, pi. 52, fig. 30a, b, 1884.

Cushman, Cushman Lab. Foram. Research Special Pub. 9, p. 

136, pi. 17, figs. 25a, b, 1937; U. S. Natl. Mus. Bull. 161, pt. 

3, p. 25, pi. 7, figs. 5a, b, 1942.

This species known from medium to deep-water 

samples in the Pacific occurs rarely but in typical form 

at Bikini and Rongelap.

Bolivina oceanica Cushman

Bolivina oceanica Cushman, Cushman Lab. Foram. Research 

Contr., v. 9, p. 81, pi. 8, figs. lOa, b, 1933; idem, Special Pub. 

9, p. 147, pi. 19, figs. 31a, b, 1937; U. S. Natl. Mus. Bull. 161, 

pt. 3, p. 24, pi. 7, figs. 4a, b, 1942.

Single specimens from two samples, Rongerik 13 and 

54, resemble this species but may not belor? here. The 

species is recorded from numerous Pacific stations but 

all in comparatively deep water.

Bolivina pseudopygmaea Cushmar 

Plate 87, figure 39

Bolivina pseudopygmaea Cushman, Cushman Lab. Foram. 

Research Contr., v. 9, p. 79, pi. 8, figs. 8a, b, 1933; idem, 

Special Pub. 9, p. 147, pi. 19, figs. 32a, b, 1037; U. S. Natl. 

Mus. Bull. 161, pt. 3. p. 29, pi. 8, figs. 6a, b, 1942.

The types and all other records for this species are 

from the vicinity of the Marshall Islands. Therefore
RECENT FORAMINIFERA OP THE MARSHALL ISLANDS 353

it is not surprising to find typical specimens in abundance in this material.

Bolivina rhomboidalis (Millett) 

Plate 87, figures 41, 42

Textularia rhomboidalis Millett, Royal Micros. Soc. Jour., 1899,

p. 559, pi. 7, figs. 4a, b. 

Bolivina rhomboidalis Cushman, U. S. Natl. Mus. Bull. 161, pt.

3, p. 19, pi. 6, figs. 7, 8, 1942.

The types of this species are from the Malay Archipelago. It is widely distributed in rather warm, 

shallow waters.

Bolivina robusta H. B. Brady

Plate 87, figure 40

Bolivina robusta H. B. Brady, Quart. Jour. Micros. Sci., v. 21, p. 

57, 1881; Challenger Rept., Zoology, v. 9, p. 421, pi. 53, figs. 

7-9, 1884.

Cushman, U. S. Natl. Mus. Bull. 161, pt. 3, p. 17, pi. 6, figs. 

2a, b, 1942.

This species described from the Ki Islands and Fiji 

occurs at Bikini and Eniwetok in typical form except 

lacking the basal spine. Specimens are distinctive 

in their crenulated sutures.

Bolivina striatula Cushman 

Plate 87, figure 43

Bolivina striatula Cushman, Carnegie Inst. Washington Pub. 311, 

p. 27, pi. 3, fig. 10, 1922; U. S. Natl. Mus. Bull. 161, pt. 3, 

p. 30, pi. 9, fig. 1, 1942.

This species has a very wide distribution. Specimens referred to it are quite abundant in the present 

material.

Bolivina subexcavata Cushman and Wickenden 

Plate 87, figure 44

Bolivina subexcavata Cushman and Wickenden, U. S. Natl. 

Mus. Proc., v. 75, art. 9, p. 9, pi. 4, figs. 4a, b, 1929.

The types of this species were from off Juan Fernandez. Specimens from several samples have been compared with the types and seem identical.

Bolivina subreticulata Parr

Bolivina subreticulata Parr, Royal Soc. Victoria Proc., v. 44, p. 12,

pi. 1, figs. 2la, b, 1932.

Cushman, U. S. Natl. Mus. Bull. 161, pt. 3, p. 31, pi. 9, fig. 2, 

1942.

Two specimens from Bikini 1175 core at 1 to 2 inches 

have been compared with topotypes of this species from 

155 fathoms, off Raine Island, and seem to be the same. 

In ornamentation they are similar to B. robusta H. B. 

Brady but are shorter and broader, and have more 

strongly raised costae.

Bolivina subtenuis Cushman

Bolivina subtenuis Cushman, Cushman Lab. Foram. Research

Special Pub. 6, p. 57, pi. 8, fig. 10, 1936; idem, Special P ib.

9, p. 148, pi. 19, figs. 33, 34, 1937; U. S. Natl. Mus. Bull. 161,

pt. 3, p. 29, pi. 8, fig. 5, 1942, 

Bolivina tennis H. B. Brady (not Marsson), Challenger Re^>t.,

Zoology, v. 9, p. 419, pi. 52, fig. 29, 1884.

A single specimen of this rare species, known from 

Fiji and Samoa, was found at Kongelap 183, at 24 fms.

Bolivina tortuosa H. B. Brady

Plate 87, figures 45-47

Bolivina tortuosa H. B. Brady (part), Quart. Jour. Micros. S°i., 

v. 21, p.-27, 1881; Challenger Rept., Zoology, v. 9, p. 420, pi. 

52, figs. 31, 32 (not 33, 34), 1884.

Cushman, U. S. Natl. Mus. Bull. 161, pt. 3, p. 20, pi. 7, figs, la, 

b, 1942.

The types of this species are from 17 fathoms off the 

Admiralty Islands. It is very widely recorded both 

living and fossil. It is especially abundant in rather 

warm, shallow waters, and is abundant in our mater:'al. 

The species is characterized by rather coarse perforations which appear white against the translucent wall, 

and some specimens showing these perforations are 

placed here even though not strongly twisted.

Genus LOXOSTOMUM Ehrenberg, 1854 

toxostomum convallarium (Millett) 

Plate 88, figures 4, 5

Bolivina convallaria Millett, Royal Micros. Soc. Jour., 1900 p 

544, pi. 4, fig. 6.

Loxostoma convallarium Cushman, Cushman Lab. Foram. Research Special Pub. 9, p. 191, pi. 22, figs. 11-13, 1937; U. S. 

Natl. Mus. Bull. 161, pt. 3, p. 37, pi. 10, fig. 6, 1942.

This is evidently a Pacific species. It is rather common at Eniwetok.^

Loxostomum karrerianum (H. B. Brady) 

Plate 88, figure 6

Bolivina karreriana H. B. Brady, Quart. Jour. Micros. Sci., v. 21, 

p. 28, 1881; Challenger Rept., Zoology, v. 9, p. 424, pi. 53, 

figs. 19-21, 1884.

Loxostoma karrerianum Cushman, Cushman Lab. Foram. Research Special Pub. 9, p. 184, pi. 21, figs. 17a, b, 1937.

A few typical specimens were found in two deep-water 

samples, Bikini 1174 and 1175.

Loxostomum limbatum (H. B. Brady) 

Plate 88, figure 7

Bolivina limbata H. B. Brady, Quart. Jour. Micros. Sci., v. 21,

p. 27, 1881; Challenger Rept., Zoology, v. 9, p. 419, pi. 52;

figs. 26-28, 1884. 

Loxostoma limbatum Cushman, Cushman Lab. Foram. Research

Special Pub. 9, p. 186, pi. 21, figs. 26-29, 1937; U. S. Natl.

Mus. Bull. 161, pt. 3, p. 35, pi. 10, figs, la, b, 1942.

The types of this species are from off the Hawaiian 

Islands. It has a very wide distribution particularly 

in warm, shallow waters.
354 BIKINI AND NEARBY ATOLLS, MARSHALL ISLANDS

loxostomum mayori (Cushman) 

Plate 88, figure 8

Bolivina mayori Cushman, Carnegie Inst. Washington Pub. 311,

p. 27, pi. 3, figs. 5, 6, 1922. 

Loxostomum mayori Cushman, TJ. S. Natl. Mus. Bull. 161, pt. 3,

p. 38, pi. 11, figs. 1, 2, 1942.

A few specimens of this widely distributed species 

were found at Eniwetok and Rongerik. In the coarsely 

perforated test they are similar to L. limbatum but 

differ in the sutures not being limbate and the whole 

test being much slenderer.

Genus BIFARINEIIA Cushman and Hanzawa, 1936

Bifarinella mackinnonii (Millett)

Plate 88, figure 9

Bifarina mackinnonii Millett, Royal Micros. Soc. Jour., 1900'

p. 281, pi. 2, figs. 15a, b.

Cushman, Cushman Lab. Foram. Research Special Pub. 9, 

p. 200, pi. 23, figs. 6, 7, 1937.

The genus Bifarinetta, known from a single species 

described from the late Tertiary of the Ryukyu Islands 

and found in deep water of the Red Sea, is represented 

by a different species in the Marshall Islands. Only 

three specimens were found but they seem to be referable to the form described from the Malay Archipelago 

at 14 fms. They differ from B. ryukyuensis Cushman 

and Hanzawa (1936, p. 46, pi. 8, figs. 7, 8) in the much 

slenderer test lacking the coarse papillate and spinose 

ornamentation. The available specimens indicate that 

the present species holds its stage of oblique sutures 

much longer and the test axis continues to be twisted 

throughout growth.

Genus RECTOBOLIVINA Cushman, 1927 

Rectobolivina bifrons (H. B. Brady)

Sagrina bifrons H. B Brady, Quart. Jour. Micros. Sci., v. 21,

p. 64, 1881; Challenger Rept., Zoology, v. 9, p. 582, pi. 75,

figs. 18-20, 1884. 

Siphogenerina (Sagrina) bifrons Egger, K. bayer. Akad. Wiss.,

Math.-naturh. Abt., Abh., Kl. 2, Band *18, p. 317, pi. 9,

figs. 25, 26, 29, 1893. 

Siphogenerina bifrons Cushman, U. S. Natl. Mus. Bull. 71, pt. 3,

p. 105, pi. 45, figs. 1, 2, 5-7, 1913. 

Rectobolivina bifrons Cushman, Cushman Lab. Foram. Research

Contr., v. 3, p. 68, 1927; idem, Special Pub. 9, p. 204, pi. 23,

figs. 13, 14, 1937.

A few specimens were found in Bikini coie 1175 at 

1 to 2 inches and 2 to 3 inches.

Genus BITUBULOGENERINA Howe, 1934

Bitubulogenerina sp.

Plate 88, figure 11

Test small, compressed, attenuated, periphery 

strongly indented; chambers few, not inflated, basal

portion flaring outward from the test and strongly 

undercut with the result that the test resembles a 

stalk surrounded by a spiral fringe; wall thin, translucent; aperture large, terminal, rounded, with a slight 

lip. Length 0.25 to 0.32 mm, breadth 0.12 to 0.15 mm. 

This seems to be the first record of thi-* genus in the 

Recent although certain species having terminal apertures but described as Bolivina may belong here. The 

present specimens may possibly be young forms of a 

Bifarina, but in the material available, no indication 

of uniserial development can be noticed. This species 

differs from early stages of Bijarina jimbriata (Millett) 

(Cushman, 1937, p. 200, pi. 23, fig. 4) in the chambers 

being much more attenuated:

Genus REUSSELLA Galloway, 19f 3

Reussella simplex (Cushman)

Plate 88, figures 1, 2

Trimosina simplex Cushman, Washington Acsd. Sci. Jour., v.

19, p. 158, figs. 2a, b (in text), 1929; U. S. Natl. Mus. Bull.

161, pt. 3, p. 44, pi. 12, figs. 7a, b, 1942. 

Reussella simplex Cushman, Cushman Lab. Foram. Research

Contr., v. 21, p. 40, pi. 7, figs, 5a, b, 1945.

This species is known only from the Indo-Pacific area. 

It is very abundant in this material.

Reussella sp. A 

Plate 88, figure 3

Rare specimens from a number of Eniwetok stations 

and one Rongerik station differ from R. simplex in their 

smaller, slenderer, and less regular tests and the wall 

which is rather granular on the surface instead of 

smooth, translucent, and ornamented by coarse perforations as in R. simplex. They closely resemble 

young specimens of R. spinulosa, var. atlantica Cushman (1947, p. 91, pi. 20, figs. 6, 7) described from off 

Florida.

Reussella sp. B

Rare specimens from Bikini core 1175 at 3 to 4 inches 

differ from R. simplex (Cushman) in having a few 

heavy, blunt spines projecting outward, nearly horizontally, from the three peripheral angles.

Genus MIMOSINA Millett, 1900 

Mimosina pacifica Cushman

Mimosina pacifica Cushman, Cushman Lab. Foram. Research 

Contr., v. 9, p. 77, pi. 8, figs. 3a, b, 1933; U. S. Natl. Mus. 

Bull. 161, pt. 3, p. 45, pi. 12, figs. 8a, b, 1942.

This species was described from material collected in 

12 fathoms off Levuka, Fiji. The otl °sr records are 

from Mokaujar Anchorage, Fiji, and Pongelap Atoll, 

Marshall Islands. It is fairly common, occurring at 

all four atolls.
TABLE 3.—Distribution of Camerinidae, Peneroplidae, Alveolinellidae, Heterohelicidae, Buliminidae, and Ellipsoidinidae in the Marshall Islands.

Family Camerinidae

Operculina ammorioides

Heterostegina suborbicularis

Cycloclypeus carpenter!

Family Peneroplidae

Peneroplis sp.

Spirolina acicularis

arietina

Monalysidium politum

Sorites marginalis

Marginopora vertebralis

Family Alveolinellidae

Neoalveolina pulchra

Family Heterohelicidae

Giimbelina? marshallana n. sp.

Bolivinita quadrilatera

Bolivinella folia

folia var. ornata

Family Buliminidae

Buliminella elegantissima

milletti

Buliminoides williamsoniana

Robertina subcylindrica

translucens

Pseudobulimina sp.

Bulimina aculeata

affinis

costata

fi.iiensis

rostrata

Oolina squamosa

Fissurina auriculata var. costata

circularly n. sp.

formosa

lacunata

milletti n. sp.

radiato -mar ginata

sp. A

sp. B

sp. C

Virgulina earlandi

Bolivina abbreviata

compacta

globulosa

nitida

oceanica

pseudopygmaea

rhomboidalis

robusta

striatula

subexcavata

subreticulata

subtenuis

tortuosa

Loxostomum convallarium

karrerianum

limbatum

mayor!

Bifarinella mackinnonii

Rectobolivina bifrons

Bitubuloeenerina sp.

Reussella simplex

sp. A

sp. B

Mimosina pacif ica

Pavonina triformis

1 Chrysalidinella fijiensis

Uvigerina ampullacea

cf. U. peregrina

porrecta

porrecta var. f imbriata

proboscidea

proboscidea var. vadescens

Siphogenerina dimorpha var. pacifica

raphana

Siphonodosaria abyssorum

lepidula

Angulogerina albatrossi var. ornata

Trifarina bradyi

Family Ellipsoidinidae

Pleurostomella bolivinoides

Parafissurina sp.

RONGERIK LAGOON

d

1

£

<N

C

7

o

40

6

1

1

1

if.

8

2

12

4

8

•* 

m

S

4

2

1

t

Ol

S

9

U

«

3

9

2

S5

«

go

ts

m

1

3

8

2

8

15

2

1

5

t

N

4

1

3

£

o i>

5

7

6

8

i>

to

1

4

2

CO

4

3

1

2

t>

90

7

1

2

1

1

1

8

1

2

6

11

1

3

15

3

1

1

4

1

1

2

4

9

2

21

1

1

1

1

7

1

6

2

1

4

10

2

1

1

1

1

1

11

8

2

2

1

1

2

7

16

£

8

6

6

2

1

1

b

1

2

10

1

22

2

1

19

17

21

5

1

1

1

1

1

2

10

12

2

1

6

1

1

/

8

2

12

14

2

1

3

1

1

1

3

4

1

1

1

2

2

1

3

4

3

1

4

4

6

1

1

5

2

1

1

2

3

2

3

8

2

1

4

2

3

6

6

3

8

1

4

2

1

19

10

4

2

S

5

SO

3

RONGELAP ATOLL

Inside the lagoon

•* 

i>

m

*

.

4

CO•*

1

4

1

19

8

i

S

5

5

3

5

21

3

10

8-

CO•* 

co

5

1

2

^c

CO^c

1

15

17

1

4

SO

t4

7

1

1

17

7

12

6

5

6

3

1

18

5

2

3

2

3

6

7

7

9

2

2

2

2

2

11

1

7

3

10

S

9

1

1

2

13

5

4

1

5

2

3

16

1

S

2

1

4

1

2

2

1

10

£

2

1

o

CO•*

2

1

1

1

10

c3

2

1

9

14

S

11

10

2

3

3

4

3

m

1

2

7

S

5

1

2

(N 

t8

£

1

1

7

7

o

3

1

i>

1

1

3

1

4

2

2

23

S

2

1

2

4

1

3

3

2

1

i

2

3

1

1

7

8

1

1

22

5

4

1

9

3

2

3

2

2

1

16

10

7

1

i

6

3

*

7

7

6

2

2

3

1

1

2

1

1

4

1

6

6

1

2

5

2

7

4

1

2

Outside 

the 

lagoon

CO

•*

4

i

1

3

3

i

^c

10

5

f

10

2

18

(N

^

2

1

5

1

2

2

1

3

5

12

8

9

1

12

6

4

1

6

1

2

1

1

1

11

1

4

3

1

1

1

5

3

1

16

3

6

1

1

BIKINI ATOLL

Beach

m

65

o

(N

ra

5

CO

ra

in 

o

«

£0

•* 

tB

Reef flat

ra

25

n

i

20

(N 

I>

ra

9

i>

o

a

<N

M

n

so

m

a

go

in

o 

m

m

15

1

2

7

6

Inside the lagoon

^

1

30

£

15

•*

/O

i>

o

CO

cq

4*

1

10

i>^c

1

1

3

10

S

2

1

2

•*

1

1

5

en

1

10

4

o 

2

6

1

16

in•*

CQ

8

ts

•*

n

5

5

c10

Z0I9

o

1

2

1

£

S

2

14

10

•*

/O

1

1

5

in

3

6

1

i5

CO 

en

1

2

20

(N

O •*

1

3

3

3

t

S

2

2

1

i> i>

1

1

6

2

o>

CD

1

2

6

5

5

<Nin

o

1

1

in 

c

1

CM

O

2

3

4

in

2

2

c

4

3

2

8

t>

5

3

1

1

8

2

01

2

2

6

1

5

4

o 

•*

1

5

1

•*

1

1

•* 

^ •* S

1

1

2

4

2

in 

o

1

2

•*

4

8

4

i>

75

o 

in

2

£

10

3

3

CO

•*

2

2

4

1

2

•»

1

5

5

6

in

11

4

2

7

CM

CO

£

1

7

4

6

^* 

o

i

£

1

2

£

i

7

1

11

o

o

5

1

2

13

2

8

c

/

17

^

2

(N

3

X

2

ti>

SO

5

2

•*

1

4

5

1

1

CO

•*

CM

^

85

7

(N

2

20

2

2

1

CO••J" •*

2

IS

1

1

i>•*

3

85

6

1

•* en in

20

1 1 1 1

1

3

3

10

£

*

3

2

2

5

6

4

5

16

2

3

2

1

1

3

2

6

1

3

1

2

3

3

8

1

1

1

1

6

1

1

10

1

1

3

1

1

1

2

1

1

1

8

2

1

1

1

1

5

1

3

1

2

1

5

1

3

3

6

2

1

3

1

1

2

4

1

1

1

1

4

1

3

1

4

10

S

2

2

1

2

2

1

4

1

1

1

1

1

1

2

1

1

2

1

2

2

2

1

1

13

4

5

9

1

1

1

2

2

1

1

8

2

1

1

1

1

1

4

1

1

3

7

£

4

3

1

1

4

5

1

10

5

1

£

5

1

5

3

3

2

1

6

2

7

1

S

2

2

1

2

2

1

1

11

6

4

23

1

3

1

1

4

7

£

1

1

2

1

4

2

2

5

IS

£5

1

3

1

1

4

3

*

1

1

1

1

4

6

1

1

1

1

1

1

1

2

6

12

1

3

11

1

9

1

7

2

2

1

11

£

1

1

1

2

14

1

3

22

1

1

2

1

4

1

13

2

4

4

1

2

4

1

3

1

1

6

1

22

1

1

1

1

4

14

5

3

2

2

6

13 15

3

1

5

5

4

11

2

1

1

1

12

1

1

20

1

1

3

1

1

1

4

5

10

1

S

1

1

4

1

1

1

1

1

12

1

6

1

m

10

m

b

17

2

in •* 

^c

1

2

1

1

2

1

1

6

1

1

1

Outer slopes

Dredge 

samples

COI>

S

10

15

10

5

•*

i

i

10

CO

v

3

2

3

9

17

7

2 1

1

1

2

16

3

1

1

1

2

1

2

4 1

o

•*

5

15

1

3

1

4

Bottom samples

53

1

5

12

1

6

CO

3

1

i>

1

2

2

1

1

1

1

1

1

1

12

1

1

17

€

5

1

1

1

12

1

10

1

5

2

4

2

1

•*

CO

1

7

5

1

1

1

6

i?

o

COi>

5

4

2

'to

i>

2

6

6

1

In

COi>

1

4

3

3

C

"

1

en

i>

2

4

en

1

3

3

3

o

1 j 1

2

1

1

2

6

2

2

1

2

9

1

12

1

3

1

2

1

11

1

8

1

1

1

1

1

2

2

1

1

3

1

1

2

1

1

1

5

7

1

1

15

5

5

9

4

1

1

3

1

2

7

3

10

6

12

1

2

1

4

24

5

18

8

2

7

Guyot

C

^f t^

1

€

TII 

i>

10

1

1

C

Tl1 

t^

1

2

•*

-

3 

o

^c in

-

1

1

1

1

1

2

1

3

4

7

9

1

1

2

S

1

1

£

1

2

10

12

/

1

16

4

5

1

1

3

2

2

8

/

12

7

20

1

12

4

1

4

1

2

2

1

1

1

1

2

2

2

9

1

16

13

5

8

12

12

2

1

3

1

6

3

4

12

2

2

3

10

1

1

12

10

3

3

1

2

9

3

1

4

2

1

1

4

3

1

1

2

1

1

4

19

1

2

2

*

1

19

12

1

2

5

2

6

1

1

3

1

1

1

2

1

12

2

10

1

1

2

17

11

9

1

1

1

2

2

C

in

"•

'CM

in

-1

"to

m

-1

^c

m

-1

in

t>

C

S

C

I>

CO

t^

"tJ2

2

3

1

1

7

1

1

2

1

5

1

5

3

13

2

2

5

2

2

6

1

20

4

2

1

2

3

5

2

1

1

1

8

1

2

1

4

1

4

2

2

2

2

1

2

1

1

7

4

8

1

4

3

4

1

1

9

2

4

16

2

5

6

14

1

3

5

5

4

18

2

7

1

3

3

1

6

3

1

9

1

3

9

/

1

4

2

6

2

10

14

1

5

2

2

2

2

1

8

4

2

10

1

/

1

5

3

2

5

3

3

2

1

3

2

1

1

2

1

7

2

2

3

1

3

2

11

1

1

1

4

1

3

1

4

2

1

5

1

8

4

7

1

1

2

5

7

2

2

4

7

2

1

2

1

2

1

1

1

1

1

2

2

1

2

3

6

11

i

1

2

6

2

2

2

3

1

1

2

3

1

3

2

11

1

1

5

1

5

5

1

11

12

3

9

6

1

1

1

1

CD 

CO

1

C

O

I>

1

=CM

o 

I>

1

2

1

4

2

3

V

o 

t%

C

o 

t^ S

1

3

1

1

1

ENIWETOK ATOLL

Inside the lagoon

o

1

8

1

S

1

1

o

S

10

a

7

1

5

2

CO 

CO

3

1

CM

6

8

1

5

1

1

1

1

7

1

12

21

16

1

2

5

S

10

2

i>

85

1

1

CO 

(N

2

9

CO

1

JO

3

o

(N

6

1

2

S

2

4

2

11

en

(N

CO

10

35

1

1

1

5

M

i

1

3

2

2

1

<N

1

0) 

(N

3

6

1

2

en

CM

2

1

3

1

1

1

1

1

3

1

5

15

1

4

1

2

5

15

1

2

1

2

5

1

1

10

1

2

1

4

1

9

1

24

7

1

1

1

1

10

15

1

2

1

1

2

3

1

1

18

10

2

1

1

1

1

1

8

6

14

12

7

1

4

1

8

2

1

1

1

1

4

1

11

7

3

5

1

1

3

3

2

1

2

£

2

1

6

18

2

3

8

to 

CO

9

13

1

1

3

6

5

6

1

2

4

13

6

20

3

1

10

2

1

4

1

2

CO 

CM

S

85

1

4

8

2

1

1

2

1

1

1

11

5

2

1

1

2

in 

en

3

1

4

CO

6

£0

4

CO

£

/O

2

7

CO

7

en

17

1

3

o

2

10

5

4

2

S

I/

15

1

19

1

1

1

1

3

5

2

1

5

4

1

10

5

1

6

1

£

1

2

2

1

2

17

1

3

5

1

3

1

10

1

1

2

1

6

1

2

1

8

1

80

3

3

1

2

2

*

9

15

7

1

S

2

2

1

4

2

2

4

1

1

4

8

20

4

1

1

£

3

o

3

8

f^

CO

5

50

2

1

3

1

1

20

1

o

(N

1

1

5

4

4

•* o 

o o

5

10

1

4

2

19

6

5

22

2

2

ro

1

4

1

13

4

£

1

1

1

1

2

5

10

2

1

16

4

10

5

3

1

4

4

£

11

16

2

1

1

5

5

1

15

14

1

1

1

1

2

£

1

2

5

3

1

1

3

2

2

4

1

1

1

8

3

5

12

2

1

5

5

3

18

2

3

21

3

1

S

4

(N

O 

CO

i>

o

12

1

1

2

3

1

1

6

1

1

1

£

4

4

17

5

5

1

CO ,

3

85

1

9

2

2

£

1

•*

CO

5

80

2

2

11

1

6

10

2

3

CM 

(N S

15

30

2

5

£0

2

5

2

2

1

1

£

1

5

2

1

5

3

2

1

2

13

2

£

5

2

1

1

/

1

1

20

3

8

2

£

4

2

3

2

9

1

1

1

5

1

1

1

2

3

1

6

2

17

£

1

1

5

5

CO

o

i" 

o

CO

to 

o

CO

3

8

1

£

4

13

2

1

1

4

3

11

2

2

1

/O

2

2

8

4

1

10

1

JO

1

1

CO 3

4

7

3

14

S

9

6

2

2

6

2

2

10

1

2

10

6

16

1

4

2

1

1

1

1

1

5

2

1

9

4

3

J5

1

5

•*

2

1

1

Outer 

slopes

S

3

6

7

TO

S

4

6

2

12

S

1

3

5

1

2

1

2

1

17

1

6

1

4

10

2

11

1

22

2

4

1

11

4

10

10

2

1

1

in 01 

S S

1

2

1

1

2

1

7

£

3

3

1

1

1 4

1

2

4

9

6

5

6

1

10

1

1

Z91053 O - 54 ( Face p. 354)
RECENT FORAMINIFERA OF THE MARSHALL ISLANDS 355

Genus PAVONINA D'Orbigny, 1826 

Pavonina triformis Parr 

Plate 88, figure 10

Pavonina triformis Parr, Royal Soc. Victoria Proc., v. 45, p. 29,

pi. 7, figs. 1-3, 1933. 

Pavonina flabelliformis Howchin (not D'Orbigny), Royal Soc.

South Australia Trans. and Proc., v. 12, p. 7, 1889. 

Heron-Alien and Earland, Royal Micros. Soc. Jour., 1924, p.

141, pi. 8, fig. 22.

A single specimen from 5 to 7 inches in Bikini core 

1176 seems referable to this species described and known 

from the Miocene and Oligocene of Australia. It has 

a more prominently projecting initial part than the 

Recent P. flabelliformis D'Orbigny (1826, p. 260, pi. 

10, figs. 10-12), including at the beginning a triserial, 

then a biserial stage. Preservation of the surface of 

the test is not good but the wall is finely beaded except 

over the initial part where it is irregularly costate.

Genus CHRTSALlDlNELLA Schubert, 1907 

Chrysalidinella fljiensis Cushman

Plate 88, figure 12

Chrysalidinella fijiensis Cushman, Cushman Lab. Foram. 

Research Contr., v. 21, p. 53, pi. 8, figs. 24a, b, 1945.

Chrysalidinella dimorpha Cushman (not H. B. Brady), Washington Acad. Sci. Jour., v. 19, p. 159, fig. 3 (in text), 1929; 

U. S. Natl. Mus. Bull. 161, pt. 3, p. 46, pi. 13, fig. 1, 

1942.

A single specimen from Rongerik 31 is typical but 

no other specimens were found. It was described from 

40 to 50 fathoms, off Fiji.

Genus UVIGERINA D'Orbigny, 1826

Uvigerina ampullacea H. B. Brady

Plate 88, figure 19

Uvigerina asperula var. ampullacea H. B. Brady, Challenger

Rept., Zoology, v. 9, p. 579, pi. 75, figs. 10, 11, 1884. 

Uvigerina ampullacea Egger, K. bayer. Akad. Wiss., Math.-

naturh. Abt., Abh., Kl. 2, Band 18, p. 313, pi. 9, fig. 37,

1893. 

Cushman, U. S. Natl. Mus. Bull. 161, pt. 3, p. 46, pi. 13, figs.

2-6, 1942.

This species occurs in a few deep-water samples at 

Bikini. It is characterized by a compact, rounded 

initial portion and a slender, attenuated apertural end. 

The species was described from Ascension Island and 

has been widely recorded in both the Atlantic and 

Pacific, and in the later Tertiary as well.

Uvigerina cf. U. peregrina Cushman 

Plate 88, figure 13

Uvigerina peregrina Cushman, U. S. Natl. Mus. Bull. 104, pt. 4, 

p. 166, pi. 42, figs. 7-10, 1923.

Abundant specimens from Bikini 1174 seem closer to 

this western Atlantic species than to any other. They 

are not well preserved, as the ornamentation is ob291053—54———4

scured by fine sediment, but otherwise they seem very

close.

Uvigerina porrecta H. B. Brady

Plate 88, figures 14, 15

Uvigerina porrecta H. B. Brady, Quart. Jour. Micros. Sci., v. 19, 

p. 60, pi. 8, figs. 15, 16, 1879; Challenger Rept., Zoology, v. 9, 

p. 577, pi. 74, figs. 21-23, 1884.

Cushman, U. S. Natl. Mus. Bull. 161, pt. 3, p. 48, pi. 13, figs. 

7, 8, 1942.

Typical specimens occur at all four atolls. T1 is 

species was described from Torres Strait and has b*-en 

widely recorded in the Pacific.

Uvigerina porrecta var. flmbriata Sidebottom 

Plate 88, figure 16

Uvigerina porrecta H. B. Brady var. fimbriata Sidebottom, Royal

Micros. Soc. Jour>, 1918, p. 147, pi. 5, fig. 23. 

Cushman, U. S. Natl. Mus. Bull. 161, pt. 3, p. 49, pi. 13, figs. 

9, 10, 1942.

A single specimen of this form was found at Rongelap 

463. The wall is covered by rather coarse spines 

throughout. The variety is unique in its extreme 

attenuation.

Uvigerina proboscidea Schwager 

Plate 88, figure 18

Uvigerina proboscidea Schwager, Novam-Exped., Geol. Theil,

Band 2, p. 250, pi. 7, fig. 96, 1866.

Cushman, U. S. Natl. Mus. Bull. 161, pt. 3, p. 49, pi 14, figs. 

1-4, 1942.

This species described from the Pliocene of F!ar 

Nicobar occurs commonly in the deeper-water samples 

of Bikini and rarely at Eniwetok. In ornamentation 

it is similar to U. ampullacea H. B. Brady, but differs 

in shape of test, being more robust and compact, r.nd 

not attenuated toward the apertural end.

Uvigerina proboscidea var. vadescens Cushman 

Plate 88, figure 17

Uvigerina proboscidea Schwager var. vadescens Cushman, Cushman Lab. Foram. Research Contr., v. 9, p. 85, pi. 8, figs. 14, 

15, 1933; U. S. Natl. Mus. Bull. 161, pt. 3, p. 50, pi. 14, figs. 

5-9, 1942.

This variety occurs in a number of deep-water samples 

at Bikini and Eniwetok, but is abundant at only a few 

stations. It was described from 21 fathoms, Guam 

Anchorage, Ladrone Islands.

Genus SIPHOGENERINA Schlumberger, 1883

Siphogenerina dimorpha var. paciflca Cushmau

Plate 88, figure 22

Siphogenerina dimorpha (Parker and Jones) var. pacifica Cashman, U. S. Natl. Mus. Proc., v. 67, art. 25, p. 13, pi. 2, fife. 9; 

pi. 3, figs. 6a, b, 1926; idem, Bull. 161, pt. 3, p. 53, pi. 15, 

fig. 4, 1942.

Sagrina dimorpha H. B. Brady (part), Challenger Rept., Zoology, 

v. 9, p. 582, pi. 76, figs. 1-3, 1884.

Single typical specimens were found at Bikini 1175
356 BIKINI AND NEABBY ATOLLS, MARSHALL ISLANDS

and Eniwetok 119. This small species is characterized 

by its cancellated wall and the depressions along the 

base of the chambers at the sutures.

Siphogenerina raphana (Parker and Jones) 

Plate 88, figures 23, 24

Uvigerina (Sagrind) raphanus Parker and Jones, Philosophical 

Trans., v. 155, p. 364, pi. 18, figs. 16, 17, 1865.

Sagrina raphanus H. B. Brady, Challenger Kept., Zoology, v. 9, 

p. 585, pi. 75, figs. 21-24, 1884.

Siphogenerina (Sagrina) raphanus Egger, K. bayer. Akad. Wiss., 

Math.-naturh. Abt., Abh., Kl. 2, Band 18, p. 317, pi. 9, 

fig. 36, 1893.

Siphogenerina raphanus Cushman, U. S. Natl. Mus. Proc., v. 

67, art. 25, p. 4, pi. 1, figs. 1-4; pi. 2, figs. 1-3, 10; pi. 5, figs. 

1, 2, 1926; idem, Bull. 161, pt. 3, p. 55, pi. 15, figs. 6-9, 1942.

Siphogenerina raphanus (Parker and Jones) var. costulata Cushman, idem, Proc., v. 51, p. 662, 1917; idem, Bull. 100, v. 

4, p. 281, pi. 56, fig. 6, 1921.

Siphogenerina costata Schlumberger, Feuille Jeun. Nat., ann. 13, 

p. 118, fig. B, 1883.

This species is recorded from numerous localities in 

fairly shallow warm water. The figured specimens 

illustrate the difference between microspheric and 

megalospheric forms.

Genus SIPHONODOSARIA A. Silvestri, 1924

Siphonodosaria abyssorum (H. B. Brady)

Plate 88, figure 26

Nodosaria abyssorum H. B. Brady, Quart. Jour. Micros. Sci., v.

21, p. 63, 1881. 

Nodosaria (?) abyssorum H. B. Brady, Challenger Kept., Zoology,

v. 9, p. 504, pi. 63, figs. 8, 9, 1884.

This species was described from 1825 fathoms in the 

South Pacific, southwest of Juan Fernandez. A few 

typical specimens occur in the core at Bikini 1176.

Siphonodosaria lepidula (Schwager) 

Plate 88, figures 27, 28

Nodosaria lepidula Schwager, Novara-Exped., Geol. Theil, v. 2,

p. 210, pi. 5, figs. 27, 28, 1866. 

Siphonodosaria lepidula van der Sluis and de Vletter, Ned. Akad.

Wetenschappen Proc., v. 45, p. 1014 (list), 1942.

This species described from the Pliocene of Kar Nicobar also occurs in the core at Bikini 1176, and is fairly 

common. Two specimens are figured to show the variation in ornamentation.

Genus ANGULOGERINA Cushman, 1927

Angulogerina albatrossi var. ornata Cushman

Plate 88, figure 20

Angulogerina albatrossi Cushman var. ornata Cushman, Cushman 

Lab. Foram. Research Contr., v. 8, p. 45, pi. 6, figs. 13, 14, 

1932; U. S. Natl. Mus. Bull. 161, pt. 3, p. 57, pi. 15, figs. 

10, 11, 1942.

Rare specimens from Bikini and Eniwetok are referred 

to this variety, although differing somewhat in having

an elongated apertural neck and rather prominent 

trilateral keels extending up onto the necK

Genus TRIFARINA Cushman, 192,°

Trifarina bradyi Cushman

Plate 88, figure 21

Trifarina bradyi Cushman, U. S. Natl. Mus. Bull. 104, pt. 4, p.

99, pi. 22, figs. 3-9, 1923; idem, Bull. 161, pt. 3, p. 59, pi. 15,

figs. 13a, b, 1942. 

Rhabdogonium tricarinatum H. B. Brady (not Vaginulina iricarinata D'Orbigny), Challenger Kept., Zoolcgy, v. 9, p. 525,

pi. 67, figs. 1-3, 1884. 

Triplasia tricarinata Cushman (not Vaginulina tricarinata

D'Orbigny), U. S. Natl. Mus. Bull. 71, pt. 3, p. 62, pi. 39,

fig. 2, 1913.

This cosmopolitan species occurs rarely at Bikini and 

Eniwetok. Specimens are shorter and proportionately 

broader than the types.

Family ELLIPSOIDINIDAE

Genus PLEUROSTOMELLA Keuss, 1860

Pleurostomella bolivinoides Schubert

Plate 88, figure 29

Pleurostomella bolivinoides Schubert, Geol. Reichsanstalt Abh.,

v. 20, pt. 4, p. 58, figs. 4a, b (in text), 1911. 

Thalmann, Eclogae geol. Helvetiae, v. 30, p. 337, pi. 21, figs.

la, b, 1937. 

Pleurostomella sp. nov. Schubert, Geol. Reichsanstalt Abh.,

Band 20, pt. 4, p. 57, figs. 4a, b (in text), 1911. 

Pleurostomella, schuberti Cushman and Harris, Cushman Lab.

Foram. Research Contr., v. 3, p. 133, pi. 25, f gs. 29a, b, 1927.

A single typical specimen was found at 5-7" in the 

core at Bikini 1176. The species was described from 

late Tertiary sediments in the Bismarck Archipelago 

and recorded from 375 fathoms in the Philippines.

Genus PARAFISSUEINA Parr, 1947 

Parafissurina sp. 

Plate 88, figure 25

A single specimen belonging in this genus was found 

at the bottom of the core at Bikini 1176. It is somewhat similar to figures of P. ventricosa (Silvestri) (1903, 

p. 10, text figs. 6a-e) from the Miocene of Italy.

Family ROTALHDAE

Genus SPIRILLINA Ehrenberg, 1843

Spirillina decorata H. B. Brady

Plate 88, figures 32, 33

Spirillina decorata H, B. Brady, Challenger Rept., Zoology, v. 9, 

p. 633, pi. 85, figs. 22-25, 1884.

Test with the dorsal side flattened or slightly concave, ventral side convex, periphery ac\ite; chamber 

with the coils increasing very gradually in width as 

added, slightly involute on the ventral side; suture 

distinct, depressed on the ventral side; wall oa the 

dorsal side with fine, generally radiate markings, more
RECENT FORAMINIFERA OF THE MARSHALL ISLANDS 357

definite and coarser on the ventral side, and somewhat 

depressed; aperture at the end of the tubular chamber. 

Diameter of present specimens up to 0.35 mm.

A few specimens may be referred to this decidedly 

variable species.

Spirillina decorata H. B. Brady var. of Sidebottom 

Plate 88, figure 35

Spirillina decorata H. B. Brady var. Sidebottom, Manchester 

Lit. Philos. Soc. Mem. and Proc., v. 52, no. 13, p. 8, pi. 2, 

figs. 6a, b, 1908.

A very few specimens from Rongelap and Bikini resemble the form recorded by Sidebottom from the 

Island of Delos. The wall is radially ornamented, like 

S. decorata, but the peripheral spines vary from small 

and numerous, as in the specimen figured by Sidebottom, to fewer and more prominent, as in the specimen figured here.

Spirillina inaequalis H. B. Brady 

Plate 88, figures 36, 37

Spirillina inaequalis H. B. Brady, Quart Jour. Micros. Sci., v. 19, 

p. 278, pi. 8, figs. 25a, b, 1879; Challenger Kept., Zoology, 

v. 9, p. 631, pi. 85, figs. 8-11, 1884.

Specimens of this widely distributed Indo-Pacific 

species are very rare in these collections.

Spirillina tuberculato-limbata Chapman 

Plate 88, figure 34

Spirillina tuberculato-limbata Chapman, Linnean Soc. Jour., 

Zoology, v. 28; p. 11, pi. 1, figs. 8a-c, 1899.

Two specimens, from Bikini 819 and Eniwetok 298, 

were found of this species described from the Funafuti 

borings. On one side the suture is obscured by the 

papillate ornamention and on the other side the suture 

is strongly limbate.

Spirillina vivipara var. densepunctata Cushman 

Plate 88, figure 31

Spirillina vivipara Ehrenberg var. densepunctata Cushman, U. S. 

Natl. Mus. Bull. 104, pt. 8, p.4, pi. 1, figs. 5a, b, 1931.

This variety differs from the typical form in the 

broader coils that are ornamented by very numerous, 

fine, and evenly spaced perforations. Diameter 0.28 

to 0.35 mm.

The types of this variety are from 7 feet, Largo 

Shoal, San Juan Harbor, Puerto Rico, and it is also 

recorded from Rio de Janeiro Harbor, Brazil. Rare 

specimens from Rongerik Atoll seem identical.

Spirillina vivipara var. revertens Rhumbler 

Plate 88, figure 30

Spinilina vivipara Ehrenberg var. revertens Rhumbler, Jahrb. 

Zool., Abt. Syst., Band 24, p. 32, pi. 2, figs. 8-10, 1906; 

Foram. Plankton-Exped., pt. 1, pi. 5, fig. 8; pi. 6, figs. 7-10, 

1911; idem, pt. 2, p. 430, 1913.

Variety differing from the typical in a thicker test 

and in having the last portion of the final whorl s%btly 

bent under on the ventral side.

The types of this variety are from off Laj'san, 

Chatham Islands. It is recorded from a number of 

localities in the Pacific but is very rare in our material.

Genus CONICOSPIRILLINA Cushman, 1927 

Conicospirillina semi-involuta Cushman

Plate 88, figure 38

Conicospirillina semi-involuta, C ashman, Cushman Lab. Foram. 

Research Contr., v. 9, p. 86, pi. 8, figs. 19a-c, 1933.

A single specimen of this species described from 40 to 

50 fathoms off Fiji was found at Sta. 51, 111 feet, in the 

lagoon at Rongerik.

Conicospirillina sp. 

Plate 88, figure 39

Very rare specimens represent a small, low, scahlike 

species of Conicospirillina. The wall is thin and transparent and the spiral suture indistinct. The periphery 

is acute and the ventral side depressed in the middle.

Genus FATEIUNA Williamson, 1858 

Patellina advena Cushman

Patellina advena Cushman, U. S. Geol. Survey Prof. Paper 129-F, 

p. 135, pi. 31, fig. 9, 1922.

Test plano-convex, variable in relative height, the 

earliest portion of one or more undivided tubular w^rls 

of greatest extent in the microspheric form, later portion 

composed of chambers spirally arranged, the length of 

the earliest chambers often more than a half whor1 but 

in the adult usually reduced to a half whorl in length, 

the latest portion in gerontic specimens sometimes 

showing a tendency to become undivided and tub'Uar; 

chambers partially divided by numerous internal septa, 

visible from the exterior and forming a radiating pattern; ventral side with numerous radiating lines; wall 

calcareous, fairly thin, finely perforate; aperture on the 

ventral side near the middle, a rounded opening into the 

ventral face. Diameter of present specimens up to 

0.23 mm.

This species occurs in both the Atlantic and Pacific. 

It is very rare in the present material from Bikini and 

Rongerik.

Patellina advena var. altiformis Cushman 

Plate 89, figure 1

Patellina advena Cushman var. altiformis Cushman, Cushman 

Lab. Foram. Research, Contr., v. 9, p. 87, pi. 9, figs. 8a, b, 

1933.

Variety differing from the typical form in the very 

high, pointed conical test.
358 BIKINI AND NEABBY ATOLLS, MABSHALL ISLANDS

The types of the variety are from 40 to 50 fathoms off 

Fiji, the only previous record. It is interesting to find 

a single specimen at Kongelap 463.

Genus PATEIIINEIIA Cushman, 1928

Patellinella fijiana Cushman

Plate 89, figure 2

Patellinella fijiana Cushman, Cushman Lab. Foram. Research 

Contr., v. 9, p. 87, pi. 9, figs. 7a, b, 1933.

The types of this species are from off Nairai, Fiji. 

Single specimens were found at Bikini 818 and Rongelap 

462. The species is characterized by its spinose wall, 

except over the ventral surface of the last two chambers, 

where it is smooth and polished.

Patellinella inconspicua (H. B. Brady) 

Plate 89, figure 4

Textularia inconspicua H. B. Brady, Challenger Rept., Zoology,

v. 9, p. 357, pi. 42, figs. 6a-c, 1884. 

Paiellinella inconspicua Cushman, Cushman Lab. Foram.

Research Contr., v. 4, p. 5, pi. 1, figs. 8a-c, 1928.

Test small, conical but somewhat compressed laterally, earliest stage with a short, spiral chamber hi the 

microspheric form, remainder of the test with two 

chambers making up each whorl, periphery rounded, 

ventral face concave; chambers distinct but not inflated, 

increasing rather rapidly but evenly in size as added; 

sutures distinct, not depressed; wall smooth; aperture 

a semi-elliptical depression at the inner border of the 

ventral face of the last-formed chamber. Height up to 

0.23 mm, breadth 0.20 mm, thickness 0.15 mm.

The records for this species are from the Pacific and 

Australian regions. It is rare in the material from 

Rongerik, Eniwetok and Kongelap but occurs in several 

samples.

Patellinella jugosa (H. B. Brady) 

Plate 89, figure 3

Textularia jugosa H. B. Brady, Challenger Rept., Zoology, v. 9,

p. 358, pi. 42, figs. 7a,b, 1884. 

Texlularia inconspicua H. B. Brady var. jugosa Millett, Royal

Micros. Soc. Jour., 1898, p. 558, pi. 7, figs. 2a,b.

A single specimen from Kongerik 12 in 120 feet is the 

only one found. It is not entirely typical but is like 

others found in the Fiji and Caroline Islands. With 

more material it may prove to be a new species.

Genus DISCORBIS Lamarck, 1804

Discorbis concinna (H. B. Brady)

Plate 89, figures 12, 13

Discorbina concinna H. B. Brady, Challenger Rept., Zoology, v.

9, p. 646, pi. 90, figs. 7, 8, 1884. 

Discorbis concinna Cushman, U. S. Natl. Mus. Bull. 71, pt. 5,

p. 16, pi. 5, fig. 3, 1915; idem, Bull. 104. pt. 8, p. 21, 1931.

Test small, plano-convex, dorsal side rather evenly

convex, ventral side flattened or somewhat concave, 

periphery subacute but not keeled, two or more whorls 

in the adult; chambers few, about four in the adult 

whorl, slightly inflated, the final one makirg up nearly 

one-half of the periphery; sutures distinct, slightly depressed, very strongly curved; wall smooth, distinctly 

perforate; aperture ventral, beneath a distinct lip-like 

projection of the ventral border of the last-formed 

chamber. Diameter 0.25 to 0.35 mm.

This is apparently an Indo-Pacific spec/es from the 

records. Specimens are fairly common in some of the 

samples. They are characterized by a circular test 

with narrow arcuate chambers, and the wall sometimes 

polished.

Discorbis crustata Cushman 

Plate 89, figure 6

Discorbis crustata Cushman, Cushman Lab. Foram. Research 

Contr., v. 9, p. 88, pi. 9, figs. 4a-c, 1933.

This species described from 24 fathoms, Nairai, Fiji, 

occurs in typical form, but rarely, at all four atolls. 

It is distinctive in its polished wall.

Discorbis frustata Cushman 

Plate 89, figure 5

Discorbis frustata Cushman, Cushman Lab. Foram. Research 

Contr., v. 9, p. 88, pi. 9, figs. 2a-c, 1933.

The types are from off Fiji in 40 to 50 fathoms, the 

only previous record. It is extremely rar^, occurring 

only at Rongerik 51 and 80.

Discorbis micens Cushman 

Plate 89, figures 8, 9

Discorbis micens Cushman, Cushman Lab. Foram. Research 

Contr., v. 9, p. 89, pi. 9, figs. 5a-c, 1933.

The types are from off the Paumotu Islands in 675 

fathoms and it has not been recorded from shallow 

water. A number of specimens with a rounded periphery seem to be very close to this specie^. As compared with D. concinna (H. B. Brady) thir species has 

an elongate test, with only four chamberr in the last 

whorl, and the last chamber is easily discernible.

Discorbis opima Cushman 

Plate 89, figures 10, 11

Discorbis opima Cushman, Cushman Lab. Foram. Research 

Contr., v. 9, p. 88, pi. 9, figs. 3a-c. 1933.

The types are from 24 fathoms, off Nairai, Fiji. It 

is a common species in the Pacific and occurs in a number of samples from all the atolls. It is characterized 

by a rounded periphery, very coarse perforations, and 

limbate dorsal sutures.
BECENT FOBAMINIFEEA OF THE MABSHALL ISLANDS 359

Discorbis pateliiformis (H. B. Brady) 

Plate 89, figure 7

Discorbina pateliiformis H. B. Brady, Challenger Rept., Zoology, 

v. 9, p. 647, pi. 88, figs. 3a-c; pi. 89, figs, la-c, 1884.

Discorbis pateliiformis Cushman, U. S. Natl. Mus. Bull. 71, pt. 

5, p. 17, pi. 5, figs. 5a-c, 1915.

Fairly common specimens of this high-spired species 

occur at all four atolls. They vary considerably in 

height and acuteness of the spire but exhibit the radial 

ornamentation of the ventral side typical of this species.

Discorbis rugosa (D'Orbigny) 

Plate 89, figure 14

Rosalina rugosa D'Orbigny, Voyage dans I'Ame'rique me'ridionale, 

tome 5, pt. 5, Foraminiferes, p. 42, pi. 2, figs, 12-14, 1839.

Discorbina rugosa H. B. Brady, Challenger Rept., Zoology, v. 9, 

p. 652, pi. 87, figs. 3a-c; pi. 91, figs. 4a-c, 1884.

A few specimens, mostly immature, of this widely 

recorded species were found at all four atolls. They 

are large and are distinguished by their very coarsely 

perforate wall and chambers which are flat on the dorsal 

surface and inflated ventrally.

Discorbis subbertheloti Cushman 

Plate 89, figure 15

Discorbis bertheloli H. B. Brady (part) (not D'Orbigny), Challenger Rept., Zoology, v. 9, p. 650, pi. 89, figs. lOa, b, 1884.

Discorbis subbertheloti Cushman, Carnegie Inst. Washington 

Pub. 342, p. 33, pi. 10, fig. 1, 1924.

Test attached, plano-convex, early chambers in a 

low spire, composed of about two and a half whorls, 

periphery subacute to rounded; chambers distinct, 

four to five in the adult whorl, slightly if at all inflated, 

increasing rapidly in size as added, slightly keeled on 

the ventral side; sutures distinct, very slightly depressed 

strongly curved both on the dorsal and ventral sides; 

wall thin, very finely but distinctly perforate; aperture 

a curved slit on the ventral margin of the last-formed 

chamber. Diameter about 0.45 mm.

The types of this species are from rather shallow 

water off Samoa. It is evidently a species of comparatively warm, shallow waters. Specimens referable 

to it occurred in a number of the samples. It is 

characterized by very fine perforations, and a flattened 

dorsal surface.

Discorbis tuberocapitata (Chapman) 

Plate 89, figure 16

Discorbina tuberocapitata Chapman, Linnean Soc. Jour., Zoology,

v. 28, p. 11, pi. 1, figs. 9a-c, 1899. 

Discorbis tuberocapitata Chapman, Annals and Mag. Nat.

History, llth ser., v. 11, p. 100 (list), 1944.

This species occurs fairly commonly at Rongelap. 

It was described from Funafuti Atoll and recorded as

better developed on the outer part of the reef tl <m in 

the lagoon. It was also found in the Funafuti fcoring.

Genus IAMARCKINA Berthelin, 1881

lamarckina scabra (H. B. Brady)

Plate 89, figure 17

Pulvinulina oblonga var. sca&m H. B. Brady, Challenger Rept., 

Zoology, v. 9, p. 689, pi. 106, figs. 8a-c, 1884.

Lamarckina scabra Cushman, Cushman Lab. Foram. Research 

Contr., v. 2, pt. 1, p. 13, 1926.

A single specimen was found at Bikini 1175 st 592 

fathoms. The species was described from 155 fathoms 

in Torres Strait.

Genus HERONALLENIA Chapman and Parr, 1931 

Heronallenia sp. ^ 

Plate 89, figure 18

An apparently undescribed species of Heronattenia is 

represented by a very few specimens from EiuVetok, 

Bikini and Rongelap. It is small, 0.25 to 0.30 mm, 

compressed, and slightly arched with the dorsal side 

convex and the ventral side concave; the periphery is 

subacute. The chambers are few and rapidly increase 

in size as added. The sutures are distinct and curved 

on the dorsal side. The wall is thin, translucent, finely 

punctate on the dorsal side, and ornamented by fine 

lines radiating out from the umbilicus on the vontral 

side. It is distinctive in its few chambers very rrmdly 

increasing in size.

Genus GYROIDINA D'Orbigny, 1826

Gyroidina soldanii D'Orbigny

Plate 89, figure 19

Gyroidina soldanii D'Orbigny, Annales sci. nat., tome 7, p. 278,

no. 5; Modeles, no. 36, ii« livraison, 1826. 

Parker, Jones, and H. B. Brady, Annals and Mag. Nat. 

History, 3d ser., v. 16, p. 25, pi. 3, fig. 86, 1865; idem, 4th 

ser., v. 8, p. 176, pi. 12, fig. 151, 1871. 

Cushman, U. S. Natl. Mus. Bull. 104, pt. 8, p. 38, pi. 8, figs.

3-8, 1931.

Rotalia soldanii H. B. Brady, Challenger Rept., Zoology, v. 9, 

p. 706, pi. 107, figs. 6, 7, 1884.

Specimens from Bikini and Eniwetok have been compared with topotypes from Rimini, Italy, and seem 

identical.

Genus EPONIDES Montfort, 1808 

Eponides tenera (H. B. Brady) 

Plate 89, figure 20

Truncatulina tenera H. B. Brady, Challenger Rept., Zoology, v. 9,

p. 665, pi. 95, figs, lla-c, 1884. 

Eponides tenera Cushman, Scripps Inst. Oceanography Bull,,

Tech. ser., v. 1, no. 10, p. 163, pi. 5, figs. 6, 7, 1927.

Typical specimens were found in several Bikini
360 BIKINI AND NEARBY ATOLLS, MARSHALL ISLANDS

samples and at Eniwetok 119. The species was 

described from the west coast of Patagonia.

Genus OSANGULARIA Brotzen, 1940

Osangularia bengalensis (Schwager)

Plate 89, figure 21

Anomalina bengalensis Schwager, Novara-lSixped., Geol. Theil,

Band 2, p. 259, pi. 7, fig. Ill, 1866. 

Parrella bengalensis Finlay, Royal Soc. New Zealand Trans.,

v. 68, p. 523, 1939.

Single typical specimens of this species described 

from the Pliocene of Kar Nicobar were found in Bikini 

core 1176, at a depth of 1 to 2 inches in the core, and at 

Bikini 1172.

Genus*OTALIA Lamarck, 1804 

Rotalia cf. R. beccarii var. tepida Cushman 

Plate 89, figure 22

Rotalia beccarii (Linn6) var. tepida Cushman, Carnegie Inst. 

Washington Pub. 344, p. 79, pi. 1, 1926.

Fairly common specimens from Eniwetok, Rongelap, 

and Rongerik seem very close to, if not identical with 

this variety known from warm, shallow waters of the 

West Indian region. Most of the Pacific specimens 

have 6 rather than 7 chambers in the last whorl.

Genus HOGLUNDINA Brotzen, 1948 

Hoglundina elegans (D'Orbigny) 

Plate 89, figure 23

Epistomina elegans (D'Orbigny) Cushman, U. S. Natl. Mus. 

Bull. 104, pt. 8, p. 65, pi. 13, figs. 6a-c, 1931.

A few well-developed and typical specimens occur at 

several deep-water stations at Bikini and at Eniwetok 

119, at 500 fathoms.

Genus POROEPONIDES Cushman, 1944

Poroeponides cribrorepandus Asano and Uchio

Plate 89, figures 24, 25

Poroeponides cribrorepandus Asano and Uchio, in Asano, Illustrated Catalogue of Japanese Tertiary Smaller Foraminifera, 

pt. 14, Rotaliidae, p. 18, figs. 134, 135, 1951.

Eponides repanda Cushman and Kellett (not Fichtel and Moll), 

U. S. Natl. Mus. Proc., v. 75, art. 25, p. 11, pi. 4, figs. 7a-c, 

1929.

Test large, biconvex, ventral side more strongly 

so than the dorsal, periphery slightly lobulate, limbate, 

blunt; chambers distinct, 6 to 9 in the last-formed 

whorl, gradually increasing in size as added, later ones 

very slightly inflated; sutures distinct, limbate, oblique 

on the dorsal side, radial on the ventral side, last one 

or two ventral sutures depressed instead of limbate; 

wall very finely perforate, usually roughened in front

of the aperture; apertures of two kinds, a low, arched 

opening under the edge of the last-formed chamber, 

extending from the periphery to the umbilicus, and a 

series of evenly spaced pores over the apertural face. 

Diameter 1.00 to 1.50 mm, thickness 0.55 to 0.85 mm.

This species differs from P. lateralis C^erquem) in 

its more regular form with chambers not becoming 

long and arcuate, and in being more strongly convex 

on the ventral than on the dorsal side. In its general 

shape and limbate sutures it rather close1 y resembles 

specimens in the Cushman Collection referred to 

Eponides repandus (Fichtel and Moll) from the Bay 

of Naples, Italy (Cushman Coll. 12844). One of these 

shows a single supplementary pore on the apertural 

face. It is our opinion that the genus Poroeponides, 

being based upon a character that may be found in 

varying degrees even within a single spec; es, is therefore inadequate as a genus. However, until more is 

learned of the cause of development of supplementary 

apertural pores, we are continuing to use Poroeponides 

for those forms that differ from Eponides in the presence 

of supplementary pores on the apertural face.

There seem to be two forms of this species, both of 

which are figured. The larger form is irore regular 

in outline with a small apertural face and consequently 

fewer pores. The smaller form tends to be irregular in 

shape with a large flaring apertural fac^, and more 

supplementary pores. The two forms seem to. grade 

into one another.

Genus EPISTOMAEOIDES Uchio, 195°

Epistomaroides polystomelloides (Parker and Jones)

Plate 89, figure 26

Discorbina polystomelloides Parker and Jones, Philos. Trans.,

1865, p. 421, pi. 19, fig. 8. 

H. B. Brady, Challenger Kept., Zoology, v. 9, p. 652, pi. 91,

fig. 1, 1884. 

Epistomaroides polystomelloides Uchio, Japanese Jour. Geol.

Geogr., v. 22, p. 158, pi. 7, figs. 1-3, 1952.

A few specimens from Rongelap, Bikini, and Rongerik, and more abundant ones from Eniwetok seem 

to be the same as this Indo-Pacific species.

Genus MISSISSIPPINA Howe, 1930

Mississippina concentrica (Parker and Jones)

Plate 89, figure 27

Pulvinulina concentrica Parker and Jones, Ms., in H. B. Brady, 

Linnean Soc. London Trans., v. 24, p. 470, pi. 48, figs. 

14a, b, 1864. 

H. B. Brady, Challenger Kept., Zoology, v. 9, p. 686, pi. 105,

figs, la-c, 1884. 

Eponides concentrica Cushman, U. S. Natl. Mus. Bull. 104,

pt. 8, p. 43, pi. 9, figs. 4, 5, 1931.

Colom, Inst. Espaflol Oceanografia, Notas y Resiimenes, 

ser. 2, no. 108, p. 39, pi. 7, figs. 136, 137, 194^.,
KECENT FORAMINIFEKA OP THE MAKSHALL ISLANDS 361

Mississippina concentrica Chapman, Royal Soc. South Australia

Trans., v. 65, p. 174, 1941.

Said, Cushman Lab. Foram. Research Special Pub. 26, p. 37, 

pi. 4, fig. 1, 1949.

Rare specimens were found in the following deepwater samples: Bikini 814, 817, 818, and Rongelap 456. 

The species was recorded by H. B. Brady from various, 

mostly deep-water, samples in the Atlantic, Mediterranean and South Pacific.

Genus PAUMOTUA Loeblich, 1952

Paumotua terebra (Cushman)

Plate 89, figure 28

Eponides terebra Cushman, Cushman Lab. Foram Research

Contr., v. 9, p. 89, pi. 10, figs, la-c, 1933. 

Paumotua terebra Loeblich, Washington Acad. Sci. Jour., v. 42,

No. 6, p. 193, fig. 1, 1952.

Three specimens were found at Bikini 817 in 114 

fathoms. The present specimens show an opaque 

band close inside the periphery on the ventral side, 

and the ventral openings, although not as numerous 

or regular as in the holotype, are distributed along 

this band. The Bikini specimens are more strongly 

compressed than the types from Anu Anuraro Atoll, 

Paumotu Islands, 405 fathoms.

Genus SIPHONINA Reuss, 1849

Siphonina tubulosa Cushman

Plate 89, figures 29, 30

Siphonina tubulosa Cushman, Carnegie Inst. Washington Pub. 

342, p. 40, pi. 13, figs. 1, 2, 1924; U. S. Natl. Mus. Proc., v. 

72, art. 20, p. 10, pi. 1, figs. 3, 5, 1927.

Truncatulina reticulata H. B. Brady (part) (not Czjzek), Challenger Rept., Zoology, v. 9, p. 669, pi. 96, figs. 5-7 (not fig. 

8), 1884.

Test compressed, keeled, the last-formed whorl consisting of three or four chambers; sutures slightly 

depressed; wall with numerous, short, tubular projections and the periphery with a border of longer projections of varying length; aperture somewhat compressed with a short, tubular neck and widely flaring lip. 

Maximum diameter 0.50 mm, usually less.

The types are from off Samoa. Rare specimens occur 

in our material.

Genus SIPHONINOIDES Cushman, 1927 

Siphoninoides echinata (H. B. Brady)

Plate 89, figures 31, 32

• 

Planorbulina echinata H. B. Brady, Quart. Jour. Micros. Sci., v.

19, p. 283, pi. 8, figs. 31a-c, 1879. 

Truncatulina echinata H. B. Brady, Challenger Rept., Zoology, v.

9, p. 670, pi. 96, figs. 9-14, 1884.

Siphonina echinata Cushman, U. S. Natl. Mus. Bull. 71, pt. 5, 

p. 42, pi. 18, figs. 1-4, 1915.

Siphoninoides echinata Cushman, Cushman Lab. Forarc. Re" 

search Contr., v. 3, p. 77, pi. 16, figs. 12a-c, 1927; U. S. Natl. 

Mus. Proc., v. 72, art. 20, p. 13, pi. 4, figs. 7, 8, 1927.

Test nearly spherical, composed of about two 

irregular whorls, the last-formed one consistirg of 

about four chambers; wall coarsely punctate, thick, 

smooth except for a few blunt, spinose projections; 

aperture a short tube with a circular opening and slight 

lip. Diameter up to 0.40 mm.

This is a widely distributed species in the IndoPacific region and occurs also in the Atlantic, It 

occurred in a number of the samples but only rare^.

Genus BAGGINA Cushman, 1926 

Baggina philippinensis (Cushman)

Pulvinulina hauerii H. B. Brady (part) (not D'Orbigny), Challenger Rept., Zoology, v. 9, p. 690, pi. 106, fig. 7 (not f -r. 6),

1884. 

Pulvinulina philippinensis Cushman, II. S. Natl. Mus. Bull. 100,

v. 4, p. 331, pi. 58, fig. 2, 1921. 

Baggina philippinensis LeRoy, Colorado School of Mines Q iart.,

v. 36, no. 1, p. 84, pi. 6, figs. 36, 37, 1941. 

Cushman and Todd, Cushman Lab. Foram. Research Contr.,

v. 20, p. 104, pi. 17, figs. 1-3, 1944. 

Discorbis allomorphinoides Cushman (not Reuss), U. S. Natl.

Mus. Bull. 100, v. 4, p. 306, pi. 58, fig. 1, 1921.

A single typical specimen was found at Bikini 1174

Baggina philippinensis var. pilulifera Cushman and Todd 

Plate 89, figure 33

Baggina philippinensis (Cushman) var. pilulifera Cushman and 

Todd, Cushman Lab. Foram. Research Contr., v. 20, p. 

105, pi. 17, figs. 4a-c, 1944.

A specimen from Bikini 815 in 55 fathoms sterns 

identical with this variety in having a series of knobs of 

clear shell material in front of the aperture. The only 

other record for this form was from Recent dredgings, 

12% miles east of Cape Byron, Australia.

Family PEGIDIIDAE

Genus PEGIDIA Heron- Alien and Earland, 1928 

Pegidia dubia (D'Orbigny)

Rotalia dubia D'Orbigny, Annales sci. nat., v. 7, p. 274, no. 34,

1826. 

Fornasini, Accad. sci. 1st. Bologna Mem., ser. 6, v. 5, p. 46,

pi. 1, figs. 14, 14a, b, 1908. 

Pegidia dubia Heron-Alien and Earland, Royal Micros. Soc.

Jour., 1928, p. 290, pi. 1, figs. 8-15.

A single typical specimen was found at Bikini 1461, 

dredged at 44 to 25 fathoms on the outer slopg of 

Bikini Atoll. This species was described from Maur this 

and recorded as rare from the Kerimba Archipe^go, 

the Java Sea, and the Philippines.
362 BIKINI AND NEARBY ATOLLS, MARSHALL ISLANDS

Genus SPHAERIDIA Heron-Alien and Earland, 1928 

Sphaeridia papillata Heron-Alien and Earland

Sphaeridia papillata Heron-Alien and Earland, Royal Micros. 

Soc. Jour., 1928, p. 294, pis. 2, 3, figs. 27-37.

A single specimen was found in Bikini lagoon at 

105 feet. It is known from the Kerimba Archipelago 

and Mauritius.

Genus RUGIDIA Heron-Alien and Earland, 1928

Rugidia corticata (Heron-Alien and Earland)

Plate 89, figure 34

Sphaeroidina corticata Heron-Alien and Earland, Zool. Soc.

London Trans., v. 20, pt. 17, p. 681, pi. 51, figs. 14-18, 1915. 

Rugidia corticata Heron-Alien and Earland, Royal Micros.

Soc. Jour., 1928, p. 289, pi. 1, figs. 5-7.

Test irregular in shape, one side flattened, the other 

strongly convex, composed in the adult of three or 

four visible, subglobular chambers; wall thick, coarsely 

punctate, covered with irregular cortications giving a 

rough appearance; apertures along the sutures on the 

convex side. Diameter up to 0.25 mm.

This species is known only from the Indo-Pacific 

region. Specimens from several samples seem like the 

figures of this species except that almost all have but 

three chambers.

Rugidia? spinosa Cushman, n. sp. 

Plate 89, figures 35-37

Test small, irregular in shape, one side formed of the 

last two chambers, strongly convex, the other side 

usually somewhat angled; chambers in the adult 

globular, increasing very rapidly in size as added; 

sutures of the later portion depressed; wall of the 

pointed portion with irregular raised areas generally 

elongate toward the point of the test, in the globular 

chambers irregular in shape; aperture a very small, 

rounded opening near the inner margin of the chamber. 

Diameter up to 0.30 mm.

Holotype (USNM 548542) from Rongerik lagoon, 

85 feet, 11°23'35" N., 167°28'35" E. (Rik 54).

This species is a puzzling one and is referred to this 

genus with considerable question. It differs from R. 

corticata (Heron-Alien and Earland) in the pointed 

early chambers, different surface ornamentation and 

the very small rounded apertures.

Family AMPHISTEGINIDAE 

Genus AMPHISTEGINA D'Orbigny, 1826

Amphistegina madagascariensis D'Orbigny 

Plate 90, figures 1, 2

Amphistegina madagascariensis D'Orbigny, Annales sci. nat.,

tome 7, p. 304, no. 5, 1826.

Fornasini, Accad. sci. 1st. Bologna Rend., v. 7, p. 3, pi. 2, 

figs. 5a-c, 1903.

Cushman, Carnegie Inst. Washington Pub. 34" p. 49, pi. 17,

fig. 3, 1924.

Amphistegina lessonii D'Orbigny var. madagascariensis Cushman, U. S. Natl. Mus. Bull. 100, v. 4, p. 372, 1921.

Test large, slightly carinate, strongly but unequally 

biconvex, umbonal region of the ventral side very 

convex, of clear shell material; chambers fairly distinct, 

few in number, not inflated; sutures strc^igly curved, 

often slightly angular in the middle, usually flush 

with the surface; wall smooth except or the ventral 

side near the aperture where it is spinose; aperture 

on the ventral side of the last-formed chamber. Diameter up to 1.65 mm.

The types of this species were from off Madagascar. 

It is very abundant in the present collections. It has 

probably been referred to many times rs A. lessonii 

D'Orbigny (1826, p. 304, pi. 17, figs. 1-4) which, however, is a thinner form with more numerous chambers 

per whorl. It is a species of the shallow, warm waters 

of the Indo-Pacific. In the Marshall Islands it occurs 

abundantly in the lagoons and down to moderate depths 

on the outer slopes of the reefs, comprising in some 

samples as much as 80 percent of the foraminifera 

present. In the deep-water samples the species is 

present but much less abundantly and the specimens 

are very much smaller.

Amphistegina radiata (Fichtel and Moll) 

Plate 90, figure 3

Nautilus radiatus Fichtel and Moll, Testacea microscopica, p.

58, pi. 8, figs, a-d, 1798. 

Amphistegina radiata Said, Cushman Lab. Foram. Research

Special Pub. 26, p. 38, pi. 4, fig. 10, 1949.

Typical specimens of this species described from the 

Red Sea were found in deep waters off Bikini and 

Eniwetok.

Amphistegina radiata var. papillosa Said 

Plate 90, figures 5, 6

Amphistegina radiata (Fichtel and Moll) var. papillosa Said,

Cushman Lab. Foram. Research Special Pub. 26, p. 39,

pi. 4, fig. 12, 1949. 

Amphistegina radiata Cushman (part) (not Fichtel and Moll),

Carnegie Inst. Washington Pub. 342, p. 49, pi. 17, fig. 2

(not fig. 1), 1924. 

Hofker, Resultats Sci. Voyage Indes Orientales Neerlandaises,

tome 2, fasc. 1, p. 6, pi. 1, figs. 8-10, 1930. 

Amphistegina lessonii Dakin (not D'Orbigny), Ceylon Pearl

Oyster Fish. Rept., v. 5, p. 240, pi. 1, fig. 13, 1906.

Variety differing from the typical form in its ornamentation, which consists of 2 or more rows of papillae 

between adjacent sutures on both sides of the test. 

Some papillae are joined together into short raised 

ridges. The present specimens include both planoconvex and biconvex forms. Diameter as much as 

1.00 mm, thickness as much as 0.50 mm.
TABLE 4. — Distribution of Rotaliidae, Pegidiidae, Amphisteginidae, Calcarinidae, Cymbaloporidae, Cassidulinidae, and Chilostomellidae in the Marshall Islands.

Family Rotaliidae 

Spirillina decorata 

decor ata var. of Sidebottom 

inaequalis 

tuberculato-limbata 

vivipara var. densepunctata 

vivipara var. revertens 

Conicospirillina semi-involuta 

sp. 

Patellina advena 

advena var. altiformis 

Patellinella fijiana 

inconspicua 

jugosa 

Discorbis concinna 

crustata 

frustata 

micene 

opima 

patelliformis 

rugosa 

subbertheloti 

tuberocapitata 

Lamarckina scabra 

Heronallenia sp. 

Gyroidina eoldanii 

Eponides tenera 

Osangularia bengaleneis 

Rotalia cf. R. beccarii var. tepida 

Hoglundina elegans 

Poroeponides cribrorepandus 

Epistomaroides polystomelloidee 

Mississippina concentrica 

Paumotua terebra 

Siphonina tubulosa 

Siphoninoides echinata 

Baggina philippinensis 

philippinensis var. pilulifera 

Family Pegidiidae 

Pegidia dubia 

Sphaeridia papillata 

Rugidia corticata

Family Amphisteginidae 

Amphistegina madagascariensis 

radiata 

radiata var. papillosa 

radiata var. venosa 

radiata var. 

Family Calcarinidae 

Calcarina hispida 

spengleri 

Family Cymbaloporidae 

Cymbaloporetta bradyi 

squammosa 

Pyropilus rotundatus 

Tretomphalus planus 

Family Cassidulinidae 

Ceratobulimina pacifica 

Epistominella cf . E. obtusa 

pulchra 

tubulifera 

sp. 

Cassidulina anguloea 

costatula 

delicata 

elegans 

gemma n. sp. 

marshallana n. sp. 

minuta 

pacifica 

patula 

cf. C. spinifera 

subglobosa 

subelobosa var. 

Cassidulinoides sp. 

Ehrenbergina pacifica 

reticulata 

Family Chilostomellidae 

Chilostomella oolinsi 

Pullenia bulloides 

Salisbury! 

Sphaeroidina bulloides 

compacta

RONGERIK LAGOON

01

en

1

95

2

CM 

O

6 

98

2 

15 

1

3 

1

3 

2

15

15 

15

1 

6

1

1

9 

3 

1

1

1

1 

1

15

8

1

3

4

m 

J

1

1

7

1

17 

5 

3 

2

4

SO

18 

10

s

7

1

2

1

70

-

3

3

4 

2

5

1

3 

8

40

t

14

1

2 

4

13

8

4 

2 

8

2

6

,

ir>

1

3

3 

4

1

1

2 

4

1 

14 

17 

1 

1 

9

4 

3 

7

5

10

"

t

2

5 

3

1 

9

2 

12

70

1

22 

3

1 

2

3 

6

3

1 

3

12

50

13

2

1

6

t3

10 

5

1 

6

1 

2

70

*

14 

1

6

1

2 

2

5 

8

4

2 

1 

2

SB

a

4 

1 

3

1

1 

2

3 

1

4 

2

55

1 

3

7

D1

1 

6

1 

6

2

1

to

S

1 

1

1

RONGELAP ATOLL

Inside the lagoon

<H

H

5 

BO

*

*

CO

•a1 

1

1 

10

2 

15 

7 

2

1 

1

6

45

1

18 

6

13 

2

3

2

5 

12

6 

17 

5 

12 

2

18

2 

5

1

50

11

I 

16 

1 

10

1

CO•&

6 

5

2

50

-

3 

5

*

•* 

•*r

11

1 

3 

2 

2

2 

3

55

S

6

c1 

7

3 

2

2

60

1

6 

2 

1

19

o

•*

4

2 

2 

2

50

3

6 

3

11

CDD9

1

1 

3

1

1 

2

60

6

1

3

2 

1

1

1

10

1 

1

3

2 

1

60

4 

11

1 

5

1

1

1

17

1 

4 

6 

5

7

1

40

17 

7

13 

1

:-

4

1

1 

1 

1 

3 

1

2

4 

3

60

t

12

1

6 

1

1

3

2 

3

5

IT10

Outside 

the 

lagoon

00

CD•a1

1

1

9 

13

2 

1 

2

11

50

go

1

5 

5

19 

2

2

3 

2

7 

2

CO•*

1

2

1

2 

2

8 

1 

2

3

7 

3

£5 

5

1

f

7

9 

3

3

CO

•* 

1

2

1

10

17 

1

45 

2C 

3

35

16

1 

1

12 

2

3

1

CM

•*

1

3 

1

.

15

50

M

1

BIKINI ATOLL

Beach

IT

9

B

6 

11

0

tt

I 

90

m

PC

15

»

o

ffi

1

5

7*

1 

2

•* 

D8

85

1

Reef flat

M

m 

i

5

55

1

m

tt

2

1

5 

1

*0

B

9

7

ITCC

1, 

S

to

s

-

s

1

to

w

1 

2

tt

t 

75

z> 

tt

J5 

50

Inside the lagoon

2

4

1

1

8 

2 

1

2

JO

40

5

18

2

5

2 

1

10

,1

18 

3

•a1

3

8

1

7

70

t

9

to

K

3

1 

2

14

SO

I

11

D- •*

1 

2

10 

2 

2

40

8 

«0

9 

4

1 

1

3

3

1

F

i

4

•a1

1

1

55

1 

60

m

2 

4

1

60

2 

4

o

2 

3

1

45

11 

S5

1

v

pa

2 

1

1

55

5

7

to •*

m

i

2

1 

1

40

19 

SO

2 

5

4

1 

2

1 

2

80

14 

6

5 

4

1

o

•a1

1 

2

50

i

1 

1

CO

2

80

2

3

1

•*

00

1

4

2 

3 

2

60

2

8 

13

1

D1

1 

2

50

1 

5

3

03

CO 

05

3 

3

65

1 

7

2

o

•*

6 

2

3 

1

80

1

1

1

2

1

1 

4

1

75

i

2

2 

6

1

c- t19 

6

1 

4

70

18 

6

16

CO

1

3

1

1 

1

1 

2

1

80

12 

1

5

CM

O

1

2

1 

2

70

2 

4

4

m0

3 

6

5 

2

3

7 

1

0

1

2

1

60

5

1

±

*

CM 

O

1

1

7

7 

4

2

1

50

24 

10

8 

3

3 

12

COD- m

12 

4

3

1

3 

2

70

1

*

4

1

1

16 

2

1

1 

1

1

80

12

7 

i

4 

1

3

o

•a1

1

-

5 

6

•*

1 

1

2

2

1

1

•* •* •*

1

1

6 

1 

2

1

3

13

10

3

1 

2

CO

o

1

3 

1

3 

3

•a1

5

1 

1

50

i

4 

6

t3

7

1

1

0

1

5

1

1 

3

1 

1

70

*

18

2

1 

1

1 

1

75

7 

9

2

CO

1 

1 

1 

2

1

70

8

13 

6

1

COi~

4 

2

1

90

i

3 

4

11 

4

60

«

i

V 

o

16 

2

75

il

20 

4

fc

1 

3

2

1

i

4 

3

2 

1

45

23 

W

23

8

1 

1

o

6 

4

1

3

1

65

14

1 

13

9 

1

3

70

3 

1

•* CO

3

65

7 

2

1

Dc1 

1 

1

40

4 

3

CM

CO

1

75

10

1

00i~ 

4

3

1 

2

CO

1

1

2

65

2

1 

1

7

5 

1

2

1

1

65

7 

7

2

•a1

1

65

5 

1

2

c-•*

1 

4

1 

1

45

5

1 

1

1

1

•a1 

1

2

8

Q

1 

1

01

5 

11 

3

1

SO

18

1 

8

2

*

1 

1

2

«

19

7

1 

2

•* 

1 

1

«

5 

4

Outer slopes

Dredge 

samples

00I~

1

1 

1

65

7

1

1 

1

2 

2

1 

1

8

50

13 

3

S 

5

3 

3

1 

1

2 

2 

2

3 

4

1

»

6

25 

85

1 

2

CO

2

9

70

1 

1

i

o

4 

1

50 

5 

3

15 

5

1

Bottom samples

1

4 

2

2

3 

1

.

10 

1

11

1

4

15 

2

JO

1 

2

90

2

1

1

1 

1

c~

00

1 

1

2

1

1 

6

2

3

60

17 

1

7 

2

6 

1

4

1

15 

7

10

1 

14

•*

5 

1

2

3 

2

3

40

8 

1

3

13 

1

8 

3

2 

JO

14

CO

1 

2

40

11 

6

6 

1

13 

2

1 

16

1

c4

^

14 

8 

5

10 

28

1 

1

CO

c9

1

Is

14 

9

13 

J5

2

1

2

1

To

CO

c11

SO

1 

15

3

6 

15

1 

2

*[•- 

c1

3 

1

CD 

C2

10 

3

1 

1

CO

1

2 

2

55

CO

1

6

1 

2 

4

9

1 

1 

1

2

9 

4 

1 

7 

11 

1

5

22 

9

5 

2

o

2 

16

2

3

5

Guyot

'CM

•*

4

9 

10

6 

1

SO

6 

1 

5

1 

24

1 

19

24 

16 

2

19 

23

1

*

1

22

6 

/ 

2

•*

1

3

7

4

J5 

15

4

10 

8

1

1

4 

2

5 

6

10 

11

3

00

•*

2

6 

7

6

12 

2 

6 

2

1

2

13 

5

3

3 

6

11 

5

3 

2

4 

1

_r

o

•a1

1

1

1

7

7

7

13 

12

3

3

3

1

8

1 

5

2 

1

o

•*

2

11 

5

7 

1

6

2

23

2 

1 

2

23 

2 

6

1

4

10

1

4 

2

3

7

2 

1

1

1

7 

1 

4 

8 

13 

1 

1

1

1

11 

3

6 

1

5 

2

3

18 

4

1 

6 

1 

1

2 

2 

1

2 

11

1 

2

'M

5 

5

5

1 

1

5

1 

1 

2

3

8

1

1

6 

2

5 

1 

12

1 

1

1 

3

To

2 

4

7

6 

1 

1

1

3

1 

12 

3

1 

4

2

2

1 

1 

6 

1

3 

1

^ 

m

7 

4

3 

2

JO 

5

2

2

4

1

5 

4

8

1 

7

2 

5

3 

3

U5 

1

3 

2

4

9 

5

9 

2

2

5 

4

9 

6

1

CO

1

2

1 

2

2

3 

3

2

CO

4 

5

1

1 

2

3

1 

16

14

8 

1

1

1

CO

3

8 

2

1

1

1

1

c5 

1 

1

10

1

1

2 

7

1

1 

1

5

2

7 

2

1

1

1

o

1 

8

3

2

1 

12

1

0

1

1

^ 

o

1

CD 

O

5

1 

1 

2

1 

1

CM

1

1

1

7 

16 

8

1 

1

1

ENIWETOK ATOLL

Inside the lagoon

o 

2

S

£5

1

1

1

m

4

o

2

2

1

1 

1

1

5

4 

4

2

1

8

1 

3

4 

2

15

1

2 

15

2

1

4

CD

2 

96

CM

1

15

80

1

1 

1

1

6 

3

1 

2

4

«0

1

7 

9

c5 

1

12

1 

6 

3 

2

10

1 

1

3

5

10 

10

3 

1

1

CO

1

3

3 

2

1

2 

3

£5

i

2

15

1 

2

01

7 

2

4 

1

1

10

2 

5

5 

1

0

1 

4

2 

3

1

«0

1 

17

1

1

1 

16

1 

2 

2 

6

1 

1

2 

4

1

55

4 

*

1

1

1

1

10

4 

11

1

4

2

2 

5

1 

1

45

5

17 

2

9 

2

00

2

10

6

3 

1

8 

4

5

1 

1

1 

2

10

1 

1

1 

1

2

50

1 

3 

1

3

1

CO

75

2

17 

2

1

1

t

8

1 

2

4 

1

1

1

13 

1

2

10

1 

3

1

75

2

1 

7

19

4 

3

3

2 

1

17

1

4 

5

5 

2 

4

50

3 

1 

15

1

13 

4

1 

1

50

2

5 

1

1 

12

1

1 

1

50

2

10

8

2 

8 

2

1

2 

5

3 

1

2

40

6 

4

5

1

5

3

01

1

8

1 

2

1

SO

1

9

1

2

1

J

1 

2

1 

3

50

g

2

J

13

mCM

3 

2

2 

1

1

1 

7 

5

9 

5

55

3

1 

11

J 

7

o

CO

23

7 

2 

1

2 

1

40

6

2 

2

11 

2

1

t1 

1

2 

4

2

1

2 

4

SO

1

7 

6

CO

1 

1

4

2

8

1

21

3

15

1

1

1

o 

4

1 

2

6 

1

50

2d

19 

7

2 

1

•*

3 

2

2

1

60

9 

2

1

1

2

0

1

t

1 

8

1

6

1 

5

2

4

50

1

6 

15

5 

1

CO

9 

1

2

50

5 

3

14

6

N

1 

1

17

c1

t

1 

1

2 

2

1

50

7

13 

1

10

CM

2

1 

1

2 

2

3

5 

1

SO

1

5 

2

20

2 

1

4

3 

1

25

1 

1

1

4

•* 

2

2 

1

13

7

2 

2

7

1 

4

3

1

55

3

2 

J

1 

6

CD

5 

4

1 

3

1

1

40

3

1 

7

1

14 

4

50

6

6 

4

17 

3

4

55

3

4

CO

1 

1

1

1

7 

2

•a1 o

CO

2

8

2 

1

1 

1

25

18 

5 

5

2

4

CO

o

1

12 

1

1

«5

3

7 

2

5 

2

2 

1

1 

1

50

1

1

19

2

1 

19

1

6

1

5

J

2

3

19 

2

•a1 

1

4

8 

13

1 

1

Outer 

slopes

CO

7 

3

6

2 

4

1

40

5 

4

5

2

1

2

1 

1

1 

3

1

1

50

2

2

1 

7

7 

1

4 

t

1

CD

5 

1

1 

1

10

^

5

1

3

8 

1

2

4 

5

1

1

11 8 

1 

1

1

3 

2

5

14

1 

1 

5 

4

2 

S

2

J 

1

3

1 

1

291053 O - 54 (Face p. 36E)
RECENT FORAMINIFEKA OF THE MARSHALL ISLANDS 363

With a large series of specimens it is possible to find 

almost every gradation between the typical form and 

this papillose variety. Nevertheless, the ornamental 

form seems worthy of distinction and so arbitrary 

limits have been set. In the Marshall Islands the 

variety is much more abundant than the typical form.

Amphistegina radiata var. venosa (Fichtel and Moll) 

Plate 90, figure 7

Nautilus venosus Fichtel and Moll, Testacea microscopica, p. 59, 

pi. 8, figs, e-h, 1798.

Amphistegina radiala (Fichtel and Moll) var. venosa Said, Gushman Lab. Foram. Research Special Pub. 26, p. 39, pi. 4, 

fig. 11, 1949.

This smooth, unornamented variety, having fewer 

chambers than the typical form, is found rarely at 

Bikini.

Amphistegina radiata (Fichtel and Moll) var. 

Plate 90, figure 4

Rare specimens from Eniwetok, Bikini and Rongelap 

are distinct from A. radiata in having a large strongly 

raised knob of clear shell material on the ventral side.

Family CAICARINIDAE

Genus CAICARINA D'Orbigny, 1826

Calcarina hispida H. B. Brady

Plate 90, figures 9-12

Calcarina hispida H. B. Brady, Quart. Jour. Micros. Sci., v. 16,

p. 406, 1876; Challenger Rept., Zoology, v. 9, p. 713, pi.

108, figs. 8, 9, 1884. 

Calcarina quoyi D'Orbigny, Annales sci. nat., tome 7, p. 276,

no. 6, 1826, nomen nudem. 

Fornasini, Accad. sci. 1st. Bologna Mem., ser. 6, v. 5, p. 45, pi.

3, figs. 8, 9, 1908. 

Calcarina hispid variety, Carpenter, Parker and Jones, Introd.

Foram., p. 218, pi. 14, figs. 6, 7, 1862.

Test small for the genus, compressed, dorsal surface 

slightly convex, ventral umbilicus slightly depressed, 

periphery rounded, slightly lobulated; chambers indistinct except the last few, narrow, inflated; sutures 

obscured except on the ventral side where they are 

radial and depressed; wall finely perforate, thickly 

covered by short coarse blunt spines throughout, except 

over the last one or two chambers, and by polished 

tubercules of shell material more thickly distributed 

at the middle of both dorsal and ventral sides, ornamented along the periphery by numerous (6 to 15 or 

more) large slender tapering spines which are themselves channeled and spinose, the large spines usually 

not much diverging from the plane of the test; aperture 

not observed. Diameter, exclusive of spines, 0.50 to 

1.00 mm, thickness 0.35 to 0.45 mm.

This species is very well developed in the Marshall

Islands samples and specimens do not appear rrorn. 

The species occurs fairly abundantly in the lagoor and 

at moderate depths on the outer slopes of the reefs. 

Its ecologic range appears to be entirely different from 

that of C. spengleri (Gmelin). These two species 

should prove useful in paleoecologic studies.

Calcarina hispida was found in material from the 

top of a coral knoll in Bikini Lagoon (B307) at a depth 

of only 35 feet. It would thus appear that depth is 

of less importance as a factor in determining whether 

the species will live in a certain location than is relation 

to the reef and to accumulation of detritus from above.

Calcarina spengleri (Gmelin) 

[[Plate 82, figures 10, 11; plate 92, figures 1-7

"Ammonshorn" Spengler, Dansk. Skr., Band 1, p. 373, pi. 

2, figs. 9a-c, 1781.

Nautilus spengleri Gmelin, Systema naturae Linnaei, Ed. 13,

p. 3371, 1788. 

Fichtel and Moll, Testacea microscopica, p. 84, pi. 14, figs.

g-i; pi. 15, figs, a-h, 1798. 

Calcarina spengleri D'Orbigny, Annales sci. nat., tome 7, p. 276,

no. 4, 1826. 

Carpenter, Parker and Jones, Introd. Foram., p. 216, pi. 14,

figs. 1-5, 8-10; figs. 33-35 (in text), 1862. 

H. B. Brady, Challenger Rept., Zoology, v. 9, p. 712, pi, 108,

figs. 5, 7, 1884. 

Hofker, Siboga-Exped., Mon. 4, p. 45, pi. 21, figs. 3-6, 8-10;

pi. 22, 1927; Resultats Sci. Voyage Indes Orientales Neerlandaises, tome 2, fasc. 1, p. 8, pi. 3, figs. 1-8, 1930. 

Calcarina defrancii H. B. Brady (not D'Orbigny), Cha'Unger

Rept., Zoology, v. 9, p. 714, pi. 108, figs. 6a-c, 1884.

Test large, lenticular, biconvex; chambers numerous, narrow, inflated, all except the last few on the 

ventral side obscured by the supplementary skeleton; 

sutures radial, depressed, observable only between the 

last few chambers on the ventral side; wall of the 

chambers coarsely perforate, thin, fragile, quickly 

covered during development of the test by a l^avy 

supplementary skeleton, the surface of which is tl ickly 

covered by highly polished tubercules, the test ornamented by long, coarse spines varying in number from 

6 to 30 or more, usually most abundant along the 

periphery, sometimes bifurcating at their ends, the 

spines themselves smooth but longitudinally channeled; aperture not observed and it is believed that 

the coarsely perforate wall of the last several chambers 

serves as the aperture. Diameter,. exclusive of spines, 

1.00 to 2.00 mm, thickness 0.60 to 1.25 mm.

In the Marshall Islands this species is found at two 

different locations and in two correspondingly different states of preservation: 1, on the reef flat in a fresh 

unworn condition with the shell wall white and the 

spines more or less complete; and 2, on the lagoon 

beaches where the specimens are spineless, being worn 

almost beyond recognition, and the shell wall is li
364 BIKINI AND NEARBY ATOLLS, MARSHALL ISLANDS

orange-colored. This species apparently lives only on 

the reef flat, a fact pointed out by Dr. Joshua I. Tracey, 

Jr., who found specimens in abundance, together with 

Marginopora vertebralis Blainville, both species rather 

firmly settled into but not attached to the small cavities on the surface and underneath the beach rocks on 

the reef flats around the outer sides of the islands and 

reefs of Bikini Atoll. The species occurs even more 

abundantly in a layer of velvety algae on the bare reef 

flat, particularly on the outer edge of the flat.

The spines of C. spengleri are well adapted for enabling the test to cling in the cavities and also for holding 

the body of the test away from actual contact with 

rock or other specimens and thus allowing free access 

of the ocean water to all parts of the animal. The 

degree of development of spines is probably related to 

the conditions under which the animal lives, especially 

turbidity of the water, and is not a specific character, 

as had previously been believed (see Cushman, p. 367, 

1919).

Two specimens from Bikini and one from Eniwetok 

similar to that figured, pi. 82, fig. 11, represent an 

extreme development of arborescent spines. A somewhat similar form was described as "Calcarina hispida 

Brady var. pulchella Chapman" (1900a, p. 15, pi. 1, 

fig. 10) from Funafuti Atoll, but the present specimens 

seem, in the surface texture of the wall, closer to Calcarina spengleri and the spines are even more numerous 

and project from the dorsal and ventral as well as the 

peripheral areas.

This species is one of the most abundant in the 

Marshall Islands, comprising as much as 95 to 98 

percent of the Foraminifera in some of the lagoon 

beach sands. It is washed out into the lagoon in small 

numbers, and is carried out onto the outer slopes of the 

atolls. It occurs rarely in the moderately deep samples 

on the outer slopes of the reefs, but is never found in the 

deep bottom samples of the adjacent guyot.

Family CYMBALOPORIDAE

Genus CYMBALOPORETTA Cushman, 1928

Cymbaloporetta bradyi (Cushman)

Plate 90, figures 13, 14

Cymbalopora poeyi (D'Orbigny) var. H. B. Brady, Challenger

Rept., Zoology, v. 9, p. 637, pi. 102, figs. 14a-d, 1884. 

Cymbalopora poeyi Rhumbler (not D'Orbigny), Jahrb. ZooL,

Abt. Syst., Band 24, p. 71, pi. 15, figs. 59a, b, 1906. 

Cymbalopora poeyi (D'Orbigny) var. bradyi Cushman, U. S.

Natl. Mus. Bull. 71, pt. 5, p. 25, pi. 10, figs. 2a-c; pi. 14,

figs. 2a-c, 1915. 

Cymbalopora bradyi Cushman, Carnegie Inst. Washington Pub.

342, p. 34, pi. 10, figs. 2-4, 1924. 

Cymbaloporetta bradyi Cushman, U. S. Natl. Mus. Bull. 104, pt.

8, p. 85, 1931.

Test with the chambers in a very flattened, irregular

spire, numerous, with as many as ten or more in the 

last-formed whorl visible on the ventral side; sutures 

distinct and slightly depressed on the dorsal side, 

ventrally deeply depressed and nearly radial; wall 

rather coarsely perforate. Diameter up to 0.90 mm.

This species is common in the Indo-Pacifio region but 

less so in the Atlantic. It is somewhat var able in the 

amount of flattening of the spire and in the number of 

chambers. It is common in the Marshall Islards.

Cymbaloporetta squammosa (D'Orbign^) 

Plate 90, figures 15, 16

Rotalia squammosa D'Orbigny, Annales sci. nat., tome 7, p. 272,

no. 8, 1826. 

Rosalina saitammosa D'Orbigny (in De la SagTa), Histoire

physique, politique et naturelle de File de Cuba, Foraminiferes, p. 91, pi. 3, figs. 12-14, 1839. 

Cymbalopora sauammosa Cushman, Carnegie Inst. Washington

Pub. 311, p. 41, pi. 6, figs. 4-6, 1922. 

Cymbaloporetta squammosa Cushman, Cushman Lab. Foram.

Research Contr., vol. 4, p. 7, 1928; U. S. Natl. Mus. Bull.

104, pt. 8, p. 83, pi. 16, fig. 4, 1931. 

Rosalina poeyi D'Orbigny (in De la Sagra), Histoire physique,

politique et naturelle de File de Cuba, Foraminiferes, p. 92,

pi. 3, figs. 18-20, 1839. 

Cymbalopora poeyi Carpenter, Parker, and Jones, Introd. Foram.,

p. 215, pi. 13, figs. 10-12, 1862. 

Discorbina poeyi Goes, K. svenska vetensk. akad. Handl., Band

19, no. 4, p. 107, pi. 8, figs. 264, 265, 1882.

This species is much higher spired than C. bradyi and 

has fewer chambers visible on the ventral side.

Genus PYROPILUS Cushman, 1934

Pyropilus rotundatus Cushman

Plate 90, figure 8

Pyropilus rotundatus Cushman, Cushman Lab. Foram. Research 

Contr., v. 10, p. 100, pi. 13, figs. 1, 2, 1934.

The types are from off Rangiroa in the Pacific and it 

has not been recorded elsewhere. It is interesting to 

find rare but typical specimens in the material from 

Rongerik, Eniwetok ard Rongelap in the shallow water 

of the lagoons.

Genus TRETOMPHALUS Moebius, 1880

Tretomphalus planus Cushman

Plate 90, figures 17, 18

Tretomphalus bulloides (D'Orbigny) var. plant Cushman,

Carnegie Inst. Washington Pub. 342, p. 36, pi. 10, fig. 8,

1924. 

Tretomphalus planus Cushman, Cushman Lab. Fon.m. Research

Contr., v. 10, p. 94, pi. 11, figs, lla-c; pi. 12, figs. 18-22,

1934.

Martin W. Johnson (1949, pp. 238, 241, 243) found 

T. planus occurring abundantly as plankton inside 

Bikini lagoon but not outside to the east and north.

In the bottom sediments, specimens were found at 

Rongelap, Bikini, and Eniwetok, most commonly at the
RECENT FORAMINIFERA OF THE MARSHALL ISLANDS 365

latter atoll. Relatively few specimens showing the 

spherical float-chamber were found.

Family CASSIDULINIDAE

Genus CERATOBUIIMINA Toula, 1915

Ceratobulimina pacifica Cushman and Harris

Ceratobulimina pacifica Cushman and Harris, Cushman Lab. 

Foram. Research Contr., v. 3, p. 176, pi. 29, figs. 9a-c, 1927.

A single specimen, probably immature, was found in 

Bikini core 1176 at 2 to 3 inches. The species was 

described from the Philippines in 494 fathoms.

Genus EPISTOMINELLA Husezima and Maruhasi, 1944

Epistominella cf. E. obtusa (Burrows and Holland)

Plate 90, figure 19

Pulvinulina exigua H. B. Brady var. obtusa Burrows and Holland, Geol. Assoc. Proc., v. 15, p. 49, pi. 2, fig. 25, 1897.

A single specimen from Bikini 1174 (1-2") seems 

close to this Eocene species and is figured for future 

reference. It also somewhat resembles one of the 

specimens included as Pulvinulina exigua H. B. Bracly 

(1884, p. 696, pi. 103, fig. 14 (not fig. 13)), but is not 

the same as that species, which is convex on the dorsal 

side and nearly flat on the ventral, whereas the opposite 

is true of the present specimen.

Epistominella pulchra (Cushman)

Pulvinulinella pulchra Cushman, Cushman Lab. Foram. Research Contr., v. 9, p. 92, pi. 9, figs. lOa-c, 1933.

A single specimen apparently identical with this 

species described from near Nairai, Fiji, was found at 

Eniwetok 321.

Epistominella tubulifera (Heron-Alien and Earland) 

Plate 90, figure 20

Truncatulina tubulifera Heron-Alien and Earland, Zool. Soc.

London Trans., v. 20, p. 710, pi. 52, figs. 37-40, 1915. 

Cushman, Carnegie Inst. Washington Pub. 342, p. 38, pi. 11, 

fig. 8, 1924.

Test small, biconvex, the ventral side much more 

convex than the dorsal, periphery acute, serrate; chambers fairly distinct on the dorsal side, much more so on 

the ventral side, six to eight in the adult whorl, increasing gradually in size as added, the peripheral edge 

fringed and serrate; sutures on the dorsal side indistinct 

except last two or three which are depressed, very 

oblique and curved, on the ventral side distinct, slightly 

depressed, strongly curved; wall on the ventral side 

smooth, on the dorsal side with a thickened fringe along 

the outer margin and in the adult with spinose projections both near the periphery and in the middle of each 

chamber, some of the projections open at their ends; 

aperture elongate, at the basal margin of the lastformed chamber on the ventral side and extending into 

the ventral face more or less parallel l-o the peripl ^ry. 

Diameter 0.38 to 0.47 mm, thickness 0.20 to 0.25 mm. 

This species was described from the general IndoPacific region and is present in the Marshall Isk.nds. 

It occurs at all the atolls, rarely in the lagoons and 

more abundantly in the deeper waters on the outer 

slopes of the atolls.

Epistominella sp. 

Plate 90, figure 21

Rare specimens from Eniwetok, Bikini, and Rong^lap 

seem to belong in this genus but are insufficient for 

complete description. They are small (about 0.35 

mm) and smoothly rounded in outline as well as on 

the periphery. The chambers are not inflated and the 

sutures are indistinct. The wall is smooth and polished 

but coarsely and conspicuously perforate. The aperture under the ventral edge of the last-formed chamber, 

appears to extend from the umbilicus to the periphery 

with a looplike extension parallel to and just ventral to 

the periphery.

In its coarsely perforate wall this species suggests 

E. pulchra (Cushman) but is not the same, as in that 

species the perforations are in definite rows, outl; ning 

chamber edges and sutures, and the periphery is acute.

Genus CASSIDUUNA D'Orbigny, 1826 

Cassidulina angulosa Cushman

Plate 90, figure 22

Cassidulina angulosa Cushman, Cushman Lab. Foram. Research 

Contr., v. 9, p. 93, pi. 10, figs. 6a, b, 1933.

Rare specimens from deep-water samples at Fikini 

and Eniwetok are identical with this species described 

from 468 fathoms in the Marshall Islands.

Cassidulina costatula Cushman 

Plate 90, figure 24

Cassidulina costatula Cushman, Cushman Lab. Foram. Research 

Contr., v. 9, p. 94, pi. 10, figs. 7a, b, 1933.

This species described from 594 fathoms in the 

Paumotu Islands, occurs fairly commonly at Bikini and 

rarely at Eniwetok. It is similar to C. decorata Sidebottom (1910b, p. 107, pi. 4, fig. 2) described from the 

southwest Pacific but is not as smoothly globular as 

that species.

Cassidulina delicata Cushman 

Plate 90, figure 25

Cassidulina delicata Cushman, Scripps Inst. Oceanography Bull., 

Tech. Ser., v. 1, no. 10, p. 168, pi. 6, fig. 5, 1927.

Fairly common specimens from Bikini and from 

Eniwetok 119 are referred to this species described from 

the west coast of America. Although larger, they seem
366 BIKINI AND NEAKBY ATOLLS, MARSHALL ISLANDS

to be the same in having a very elongate, curved aperture and the alternating chambers occupying only a 

very small space between the main series on each side.

Cassidulina elegans Sidebottom 

Plate 90, figure 23

Cassidulina elegans Sidebottom, Quekett Micros. Club Jour., 

ser. 2, v. 11, p. 106, pi. 4, figs, la-c, 1910.

Very rare specimens from four deep-water samples 

off Bikini seem to be the same as Sidebottom's species 

described from 1050 fathoms in the southwest Pacific. 

The angulation of the chambers in the present specimens is even stronger than shown in Sidebottom's 

figure. This species is also related to C. alternans 

Yabe and Hanzawa (1926, p. 53, text figs, a, b) described from south of the Riukiu Islands, but comparison with a specimen of that species shows they are 

not the same.

Cassidulina gemma Todd, n. sp. 

Plate 90, figures 26, 27

Test of medium size, outline lobulated; chambers 

distinct, inflated, about six pairs forming the last 

whorl; sutures distinct, limbate, depressed, especially 

the later ones; wall thin, translucent, conspicuously 

perforate; aperture an elongate loop-shaped opening 

extending into the concave face of the last-formed 

chamber, surrounded by a slight, raised lip, in the 

early stages nearly circular instead of elongate. Length 

0.35 to 0.45 mm, breadth 0.25 to 0.35 mm, thickness 

0.25 to 0.35 mm.

Holotype (USNM 548668) from Sylvania guyot, 835 

fathoms, 11°47'35" N., 164°59/20// E. (Bik 1172).

This species differs from <7. subglobosa, H. B. Brady 

in its inflated chambers and consequently lobulated 

test and in its translucent and distinctly perforate wall.

It occurs commonly at Bikini 1172 and a single 

specimen was found at Bikini 1175.

Cassidulina marshallana Todd, n. sp. 

Plate 90, figures 28, 29

Test nearly spherical, very slightly compressed; 

chambers not inflated, 4 or 5 pairs forming the last 

whorl, very slowly increasing in size as added; sutures 

indistinct, flush; wall thick, opaque, smooth, polished, 

very finely perforate; aperture a long, much curved 

slit hi the ventral face of the last-formed chamber, wall 

below aperture concave with sharp, outward flaring 

edge, and wall above smoothly rounded and bent 

inward. Diameter 0.55 to 1.10 mm.

Holotype (USNM 548685) from core on Sylvania 

guyot, at 1 to 2 inches hi the core, 410 fathoms, 

11°41'00" N., 165°14/01 // E. (Bik 1174 (l"-2")).

This species differs from C. subglobosa H. B. Brady 

in its more smoothly rounded test and the very long 

aperture which is curved downward almost into a 

semicircle. In its aperture this specie,-' somewhat 

suggests C. caudriae Cushman and Stainforth (1945, 

p. 64, pi. 12, figs. 2, 3). In some specimens the aperture 

gives the impression of being a large arcHed opening 

nearly filled by a large platelike tooth, but with proper 

lighting it can be seen that only one end of the long 

arched opening actually joins the suture, although the 

opposite end nearly does.

This species was found at Bikini 1174 (1-2) and 

Eniwetok 119 only.

Cassidulina minuta Cushman 

Plate 90, figure 35

Cassidulina minuta Cushman, Cushman Lab. Fo-am. Research 

Contr., v. 9, p. 92, pi. 10, figs. 3a-c, 1933.

The types of this species are from off t^e Paumotu 

Islands at 844 fathoms. This is by far the most 

abundant species of the genus in the present material, 

and unlike the other species of the genur is not restricted to the deep-water samples.

Cassidulina pacifica Cushman

Cassidulina calabra H. B. Brady Xn°t Seguenzi), Challenger 

Kept., Zoology, v. 9, p. 431, pi. 113, figs. 8a-c 1884.

Cassidulina pacifica Cushman, Cushman Lab. Foram. Research 

Contr., v. 1, pt. 3, p. 53, pi. 9, figs. 14-16, 1975.

Very rare specimens from two Rongelap samples are 

referred to this small globular species, the types of 

which are from Torres Strait.

Cassidulina patula Cushman

Cassidulina patula Cushman, Cushman Lab. Foram. Research 

Contr., v. 9, p. 92, pi. 10, figs. 2a-c, 1933.

Very rare specimens from several deep-wr.ter sampler 

at Bikini and Eniwetok belong to this species described 

from 486 fathoms in the Paumotu Islands, They aredistinctive in their few chambers and distinctly perforate wall.

Cassidulina cf. C. spinifera Cushman and Jarvis 

Plate 90, figure 34

Cassidulina spinifera Cushman and Jarvis, Cushman Lab. 

Foram. Research Contr., v. 5, p. 17, pi. 3, figs, la, b, 1929.

A single specimen from Bikini 1170 is interesting in 

its close relationship to this species known from the 

Oligocene of Trinidad. 11 is figured for future reference..
RECENT FORAMINIFERA OF THE MARSHALL ISLANDS 367

Cassidulina subglobosa H. B. Brady 

Plate 90, figures 30-32

Cassidulina subglobosa H. B. Brady, Quart. Jour. Micros. Sci., 

v. 21, p. 60, 1881; Callenger Kept., Zoology, v. 9, p. 430, pi. 

54, figs. 17a-c, 1884.

Specimens referable to Brady's species occur rather 

commonly in a number of Bikini samples and at 

Eniwetok 119 and 116, all deep-water samples. Specimens vary considerably in size and inflation of the 

chambers and several are figured to illustrate this.

Cassidulina subglobosa H. B. Brady var. 

Plate 90, figure 33

A few specimens, similar to that figured, from Bikini 

1174 (1-2) and 1175 and Eniwetok 119 show a pattern 

of straight grooves radiating out in all directions from 

the aperture.

Genus CASSIDUIINOIDES Cushman, 1927 

Cassidulinoides sp.

A small, compact and compressed form, with a 

translucent wall appears to belong hi this genus but is 

different from C. bradyi (Norman) (H. B. Brady, 1884, 

p. 431, pi. 54, figs. 6-10) hi being less elongate and 

having fewer uncoiled chambers. It is similar to the 

form referred to C. bradyi (Norman) from fairly deep 

water (90 to 300 meters) of the Atlantic continental 

shelf, off the northeastern United States (Parker, 1948, 

p. 224, 237, pi. 3, fig. 12).

Genus EHRENBERGINA Reuss, 1850

Ehrenbergina paciflca Cushman

Plate 90, figure 36

Ehrenbergina serrata H. B. Brady (part) (not Reuss), Challenger 

Kept., Zoology, v. 9, pi. 55, figs. 6, 7, 4 (?) (not figs. 2, 3, 

5 (?)), 1884.

Ehrenbergina pacifica Cushman, U. S. Natl. Mus. Proc., v. 70, 

art. 16, p. 5, pi. 2, figs. 2a-c, 1927.

This species, the types of which are from off the Ki 

Islands, in 129 fathoms, occurs fairly commonly at 

Bikini and in two Eniwetok samples. The wall is 

smooth and polished and the peripheral angles and 

spines project outward nearly horizontally.

Ehrenbergina reticulata Cushman 

Plate 90, figure 37

Ehrenbergina reticulata Cushman, Cushman Lab. Foram. Research Contr., v. 9, p. 94, pi. 10, figs. 9a, b, 1933.

Rare specimens from Bikini 1175 and 1176 (5"-7") 

are identical with this species described from 1937 

fathoms, off the Cook Islands. In the figured specimen 

the last-formed chamber is without surface ornamentation but is densely perforate.

Family CHIIOSTOMELLIDAE

Genus CHILOSTOMELIA Reuss, 1850

Chilostomella oolina Schwager

Chilostomella oolina Schwager, R. com. geol. Italia Boll. v. 9, p.

527, pi. 1, fig. 16, 1878.

Cushman and Todd, Cushman Lab. Foram. Research Contr., 

v. 25, p. 91, pi. 15, figs. 23, 24, 1949.

A single specimen was found hi Bikini 1175 (0-1).

Genus PUIIENIA Parker and Jones, 1862 

Pullenia bulloides (D'Orbigny)

Nonionina bulloides D'Orbigny, Annales sci. nat., tome 7, p 293, 

1826; Foraminiferes fossiles du bassin tertiaire de Vienne, 

p. 107, pi. 5, figs. 9. 10, 1846.

Pullenia bulloides Cushman and Todd, Cushman Lab. Foram. 

Research Contr., v. 19, p. 13, pi. 2, figs. 15-18, 1943.

Very rare specimens from Bikini may be referred to 

this widely recorded species.

Pullenia Salisbury! R. E. and K. C. Stewart 

Plate 90, figure 38

Pullenia salisburyi R. E. and K. C. Stewart, Jour. Paleontology, 

v. 4, p. 72, pi. 8, figs. 2a, b, 1930.

This species described from the Pliocene of California 

and known from the Miocene to the Recent is more 

common in the Marshall Islands than is P. bulloides 

(D'Orbigny). Specimens vary considerably in relative 

thickness.

Genus SPHAEROIDINA D'Orbigny, 1826

Sphaeroidina bulloides D'Orbigny

Plate 90, figure 39

Sphaeroidina bulloides D'Orbigny, Annales sci. nat., tome 7,

p. 267, no. 1; Modeles, no. 65, 1826.

Cushman and Todd, Cushman Lab. Foram. Research Contr., 

v. 25, p. 13, pi. 3, figs. 8-11, 1949.

A few specimens from several of the deep Bikini and 

Eniwetok samples are referred to this widely reccrded 

species. They are very thin walled and translucent 

and smaller than topotypes of S. bulloides from the 

Adriatic, but otherwise seem very similar.

Sphaeroidina compacta Cushman and Todd

Sphaeroidina compacta Cushman and Todd, Cushman Lab.

Foram. Research Contr., v. 25, p. 19, pi. 4, figs. 14a, b, 1949. 

Sphaeroidina bulloides Cushman (not D'Orbigny), U. S. NatL

Mus. Bull. 71, pt. 4, p. 18, pi. 12, figs, la, b, 1914.

Two rather typical specimens of this species described from the Yucatan Channel between the Gulf of 

Mexico and the Caribbean Sea and also known from 

the Pacific near Guam, were found in the sample from 

Bikini 1174 (l"-2").
368 BIKINI AND NEARBY ATOLLS, MARSHALL ISLANDS

Family GLOBIGERINIDAE

Genus GLOBIGEEINA D'Orbigny, 1826

Globigerina bulloides D'Orbigny

Plate 91, figure 2

Globigerina bulloides D'Orbigny, Annales sci. nat., tome 7, p. 277, 

no. 1; Modules, jeune, n° 17, i te livr., et adulte, ive n° 76, 

1826; (in Barker-Webb and Berthelot), Histoire naturelle 

des lies Canaries, tome 2, pt. 2, Foraminifferes, p. 132, 

pi. 2, figs. 1-3, 28, 1839.

Williarnson, Recent Foraminifera of Great Britain, p. 56, 

pi. 5, figs. 116-118, 1858.

Parker, Jones, and H. B. Brady, Annals and Mag. Nat. 

History, 3d ser., v. 16, pp. 21, 31, pi. 2, figs. 55, 56, 1865.

Cushman, Cushman Lab. Foram. Research Contr., v. 17, p. 

38, pi. 10, figs. 1-13, 1941.

A few specimens, mostly smaller than topotypes from 

Rimini on the Adriatic, are very similar to this widely 

distributed species. They are distinguished by a rather 

finely spinose wall and by having four chambers, 

rapidly increasing in size, in the last whorl.

Globigerina cf. G. conglomerata Schwager 

Plate 91, figure 4

Globigerina conglomerata Schwager, A/wara-Exped., Geol. Theil, 

v. 2, p. 255, pi. 7, fig. 113, 1866.

In the deeper samples at Bikini a rather small, compact form, showing 4 chambers in the last whorl, and 

having a smooth, polished, but conspicuously punctate 

wall is somewhat similar to this species described from 

the Pliocene of Kar Nicobar and widely recorded in 

Recent seas.

Globigerina subcretacea Lomnicki 

Plate 91, figure 1

Globigerina subcretacea Lomnicki, Akad. Umiej. Krakowie, Czesc 

II, p. 57, 1901.

Globigerina subcretacea Chapman, Linnean Soc. Jour., Zoology, 

v. 28, p. 404 (list); p. 410, pi. 36, figs. 16a, b, 1902.

Globigerina cretacea H. B. Brady (part) (not D'Orbigny), Challenger Rept., Zoology, v. 9, p. 596, pi. 82, figs. lOa-c (not 

figs, lla-c), 1884.

Test large, compact, somewhat compressed, spire low 

or depressed, periphery indented; chambers inflated, 

six or seven comprising the adult whorl, not much increasing in size as added, the last chamber in some 

instances slightly smaller than previous ones; sutures 

distinct, incised; wall rather thick for the genus, smoothsurfaced and covered with closely spaced fine pits with 

a pore at the bottom of each pit, the pits showing no 

particular arrangement; aperture small, semicircular, 

slightly rimmed, under the ventral edge of the lastformed chamber about half way between the periphery 

and the umbilicus. Diameter up to 0.70 mm.

The species is based upon Brady's figure 10 on plate 

82 for which the same new name, subcretacea, was proposed independently and nearly simultaneously by two 

authors: Lomnicki (1901) and Chapman (1902). 

Brady's figure 10, mentioned above, is of a specimen 

from 580 fathoms off the Ki Islands. Tv e specimen 

figured by Chapman is from deep dredgings around 

Funafuti.

This species occurs abundantly in the deep-water 

Bikini samples and was found rarely at Kiiwetok and 

Rongelap.

Globigerina? sp.

Plate 91, figure 3

A few specimens of a very minute forrr, about 0.22 

mm in greatest diameter, with 5 or 6 chambers in the 

last whorl, and a nearly smooth but distinctly perforate 

wall, occur in some numbers. A specimen is here 

figured for future reference.

As it occurs fairly commonly inside the lagoon at 

Rongerik, it does not seem likely that it is an oceanic 

form and its reference to Globigerina is doubtful.

Genus GLOBIGERINOIDES Cushman, 1927

Globigerinoides 2 conglobata (H. B. Brady)

Plate 91, figure 12

Globigerina conglobata H. B. Brady, Quart. Jour. Micros. Sci., 

v. 19, p. 72, 1879; Challenger Rept., Zoology, v. 9, p. 603, 

pi. 80, figs. 1-5; pi. 82, fig. 5, 1884.

Globigerinoides conglobata Cushman, Am. Jour. Sci., v. 239, pi. 2, 

fig. 2, 1941.'

This species occurs abundantly at the deep-water 

stations at Bikini, Eniwetok, and Rongelap.

At Bikini, it is not known inside the lagoon except 

in the core at 218. It occurs commonly at the bottom 

of the core, 12 to 16 inches, but only a sir^le specimen 

was found at the top of the core. This would suggest 

that 218 was open to oceanic waters when the sediments 

at 12 to 16 inches below the surface of the core were 

deposited.

Globigerinoides elongata (D'Orbigny) 

Plate 91, figure 5

Globigerina elongata D'Orbigny, Annales sci. nat., tome 7, p. 277,

no. 4, 1826.

Parker, Jones, and H. B. Brady, Annals and Mag. Nat. History, 4th ser., v. 8, p. 175, pi. 11, fig. 129, 1871. 

Fornasini, Accad. sci. 1st. Bologna Mem., ser. £. v. 7, p. 11, pi.

3, figs. 8-10, 1899.

Globigerinoides elongata Cushman, Cushman Lab. Foram. Research Contr., v. 17, p. 40, pi. 10, figs. 20-23; pi. 11, figs. 

3a-c, 1941.

A single specimen, here figured, is very similar to this 

species described from the Adriatic at Rimini and the

2 Regarding gender of Qlobigerinoides, I prefer to follow the recommendation of 

Roland W. Brown that a generic name ending in -oides, an adjectival suffix meaning 

like, takes its gender from the gender of the generic name on whnh it is based. Thus 

Qloblgerinoides, like Globigerina, is feminine; Ammodiscoides, like Ammodiscus, is 

masculine; and Haplophragmoides, like Haplophragmiutn, is nerter. R. Todd.
RECENT FORAMINIFERA OF THE MARSHALL ISLANDS 369

Pliocene of Castel Arquato, Italy. It is unique in its 

very high spire and belongs in Globigerinoides as it has 

supplementary apertures.

Globigerinoides rubra (D'Orbigny) 

Plate 91, figure 6

Globigerina rubra D'Orbigny (in De la Sagra), Histoire physique,

politique et naturelle de Pile de Cuba, Foraminiferes, p. 82,

pi. 4, figs. 12-14, 1839. 

Globigerinoides rubra Cushman, Am. Jour. Sci., v. 239, pi. 2, fig. 3,

1941; Cushman Lab. Foram. Research Contr., v. 21, p. 75,

pi. 12, figs. 6-9, 1945.

This species was found at all four atolls. Only a few 

of the specimens, generally the smallest, show the red 

color.

Globigerinoides sacculifera (H. B. Brady)

Plate 91, figure 7

Globigerina sacculifera H. B. Brady, Quart. Jour. Micros. Sci., v.

19, p. 73, 1879; Challenger Kept., Zoology, v. 9, p. 604, pi. 80,

figs. 11-17; pi. 82, fig. 4, 1884. 

Globigerinoides sacculifera Cushman, Am. Jour. Sci., v. 239, pi. 3,

fig. 1, 1941.

This species and G. rubra (D'Orbigny) include the 

bulk of the specimens of the Globigerinidae in the Marshall Islands. Both species occur more or less abundantly at all four atolls, even inside the lagoons. However, specimens from inside the lagoons are notably 

smaller than normal size.

Globigerinoides sacculifera var. fistulosa (Schubert) 

Plate 91, figure 13

Globigerina fistulosa Schubert, K. k. geol. Reichsanstalt Verb., no.

14, p. 324 (7), fig. 2 (in text), 1910; K. k. geol. Reichsanstalt

Abh., Band 20, pt. 4, p. 100, figs. 13a-c (in text), 1911. 

Globigerinoides sacculifera (H. B. Brady) var. fistulosa Cushman,

Cushman Lab. Foram. Research Special Pub. 5, pi. 34, figs.

6a-c, 1933.

A few specimens were found in the core at Bikini 1176. 

The variety was described from Globigerina, marls of 

New Guinea, and recorded from the Pliocene of Java. 

The figured specimen represents an extreme development of the fistulose processes on the later chambers.

Genus Globigerinella Cushman, 1927

Globigerinella aequilateralis (H. B. Brady)

Plate 91, figure 8

Globigerina aequilateralis H. B. Brady, Quart. Jour. Micros. Sci., 

v. 19, p. 71, 1879; Challenger Rept., Zoology, v. 9, p. 605, pi. 

80, figs. 18-21, 1884.

Globigerinella aequilateralis Cushman, Am. Jour. Sci., v. 239, pi. 2, 

fig 1, 1941.

This species occurs commonly in several of the deepwater samples at Bikini and Eniwetok.

Genus HASTIGERINELLA Cushman, 1927

Hastigerinella digitata (Rhumbler)

Plate 91, figures 9, 10

Hastigerina digitata Rhumbler, Foram. Plankton-Exped., v. 3,

pt. 1, pi. 37, figs. 9a, b, 1911. 

Hastigerinella digitata Wiesner, Deutsche Siid-Polar Exped.,

Band 20, Zoology, p. 135, pi. 22, figs. 264-266, 1931.

A few specimens, mostly small and immature, were 

found at Eniwetok and Bikini. Young specimens differ 

from Globigerinella aequilateralis (H. B. Brady) in the 

chambers being more rapidly enlarging and rather 

attenuated and the aperture not extending over onto 

the dorsal side as in Globigerinella. The texture c* the 

wall surface in both species is very similar.

Genus ORBUIINA D'Orbigny, 1839 

Orbulina universa D'Orbigny

Orbulina universa D'Orbigny (in Barker-Webb and Berthelot), 

Histoire naturelle des lies Canaries, tome 2, pt. 2, Foraminiferes, p. 123, pi. 1, fig. 1, 1839; (in De la Sagra), Hfstoire 

physique, politique et naturelle de Pile de Cuba, Foraminiferes, p. 3, pi. 1, fig. 1, 1839. 

Cushman, Am. Jour. Sci., v. 239, pi. 4, fig. 2, 1941.

Orbulina universa occurs abundantly in most of the 

deep-water Bikini samples and rarely at Eniwetol- and 

Hongelap, but no specimens were found from inside 

Hongerik lagoon. The occurence of this and other 

pelagic species seems to depend more on exposed position 

than depth of the sample.

Genus PUIIENIATINA Cushman, 1927

Pulleniatina obliquiloculata (Parker and Jones)

Plate 91, figure 11

Pullenia obliquiloculata Parker and Jones,^Philos. Trans., v. 155,

p. 368, pi. 19, figs. 4a, b, 1865. 

H. B. Brady, Challenger Rept., Zoology, v. 9, p. 618, pi. 84,

figs. 16-20, 1884. 

Pulleniatina obliquiloculata Cushman, Am. Jour. Sci., v. 239,

pi. 4, fig. 3, 1941. .

This planktonic species has a similar distribution hi 

the deeper-water samples of Bikini and Eniwetok.

Genus SPHAEROIDINELLA Cushman, 1927

Sphaeroidinella dehiscens (Parker and Jones)

Plate 91, figure 14

Spaeroidina dehiscens Parker and Jones, Philos. Trans., v. 155,

p. 369, pi. 19, figs. 5a-c, 1865. 

H. B. Brady, Challenger Rept., Zoology, v. 9, p. 621, pi. 84,

figs. 8-11, 1884. 

Sphaeroidinella dehiscens Cushman, Am. Jour. Sci., v. 239, pi. 5,

fig. 1, 1941.

This species is well developed and abundant in most 

of the deep-water samples from Bikini and Eniwetok.
370 BIKINI AND NEARBY ATOLLS, MARSHALL ISLANDS

Genus CANDEINA D'Orbigny, 1839

Candeina nitida D'Orbigny

Plate 91, figure 15

Candeina nitida D'Orbigny (in De la Sagra), Histoire physique, 

politique et naturelle de 1'Ile de Cuba, Foraminiferes, p. 108, 

pi. 2, figs. 27, 28, 1839.

H. B. Brady, Challenger Rept., Zoology, v. 9, p. 622, pi. 82, 

figs. 13-20, 1884.

Cushman, Am. Jour. Sci. v. 239, pi. 1, fig. 1, 1941.

This cosmopolitan species occurs in the deep-water 

Bikini and Eniwetok samples.

Family GLOBOROTAIHDAE

Genus GLOBOEOTAIIA Cushman, 1927

Globorotalia crassa (D'Orbigny)

Plate 91, figure 16

Pulvinulina crassa D'Orbigny, sp. H. B. Brady, Challenger Rept., 

Zoology, v. 9, p. 694, pi. 103, figs. 11, 12, 1884.

Globorotalia crassa Heron-Alien and Earland, Discovery Repts., 

v. 4, p. 428, 1932.

This form, which is midway between G. truncatulinoides (D'Orbigny) and G. cf. G. hirsute, (D'Orbigny), 

occurs rarely at Bikini and Eniwetok. It is less thick 

and has blunter angles at the periphery and along the 

face of the last-formed chamber than G. truncatulinoides, 

and is a thicker, more compact form than G. cf. G. 

hirsuta.

Globorotalia cf. G. hirsuta (D'Orbigny) 

Plate 91, figure 17

Rotalina hirsuta D'Orbigny (in Barker-Webb and Berthelot), 

Histoire naturelle des lies Canaries, tome 2, pt. 2, Foratniniferes, p. 131, pi. 1, figs. 37-39, 1839.

Globorotalia hirsuta Cushman, U. S. Natl. Mus. Bull. 104, pt. 8, 

p. 99, pi. 17, figs. 6a-c, 1931.

Specimens resembling this species described from 

the Canary Islands, but differing in being considerably 

more inflated ventrally, occur rarely in several Bikini 

samples and one Eniwetok sample. The ventral side, 

particularly the earlier chambers, shows a coarsely 

hispid surface.

Globorotalia menardii (D'Orbigny) 

Plate 91, figure 19

Rotalia menardii D'Orbigny, Annales sci. nat., tome 7, p. 273,

no. 26; Modeles, no. 10, ire livraison, 1826. 

Parker, Jones, and H. B. Brady, Annals and Mag. Nat.

History, 3d ser., v. 16, p. 20, pi. 3, fig. 81, 1865. 

Pulvinulina menardii H. B. Brady, Challenger Rept., Zoology,

v. 9, p. 690, pi. 103, figs. 1, 2, 1884. 

Flint, U. S. Natl. Mus. Rept. for 1897, p. 329, pi. 73, fig. 3,

1899. 

Globorotalia menardii Cushman, U. S. Natl. Mus. Bull. 104,

pt. 8, p. 91, pi. 17, figs, la-c, 1931; Am. Jour. Sci., v. 239,

pi. 3, fig. 2, 1941.

This species occurs at most stations with G. tumida 

(H. B. Brady) from which it differs in its less inflated

test and clear or translucent wall without the heavy 

beading characteristic of the latter species.

Globorotalia menardii var. fimbriata (H. B. Brady) 

Plate 91, figure 20

Pulvinulina menardii var. fimbriata H. B. Brady, Challenger 

Rept., Zoology, v. 9, p. 691, pi. 103, figs. 3a, b, 1884.

Globorotalia menardii (D'Orbigny) var. fimbriata C 'shman, U. S. 

Natl. Mus. Bull. 104, pt. 8, p. 94, pi. 17, figs. 2a, b, 1931.

Only three specimens from Bikini 1175 are referable 

to this variety which has a fringed and spinose periphery.

Globorotalia truncatulinoides (D'Orbigrv) 

Plate 91, figure 18

Rotalina truncatulinoides D'Orbigny (in Barker-Webb and 

Berthelot), Histoire naturelle des lies Canaries, tome 2, 

pt. 2, Foraminiferes, p. 132, pi. 2, figs. 25-27, 1839.

Globorotalia truncatulinoides Cushman and Wiekenden, U. S. 

Natl. Mus. Proc., v. 75, art. 9, p. 14, pi. 6, figs. 3a-c, 1929. 

Cushman, Am. Jour. Sci., v. 239, pi. 4, fig. 1, 1941.

This species occurs commonly in a few deep-water 

samples from Bikini and Eniwetok.

Globorotalia tumida (H. B. Brady) 

Plate 91, figure 21

Pulvinulina tumida H. B. Brady, Challenger Rept., Zoology, v. 9,

p. 692, pi. 103, figs. 4-6, 1884. 

Globorotalia tumida Cushman, U. S. Natl. Mus. B ill. 104, pt. 8,

p. 95, pi. 17, figs. 3a-c,1931; Am. Jour. Sci., v. 239, pi. 3,

fig. 3, 1941.

This widely distributed species occurs abundantly 

in a few of the deeper-water samples from Bikini, and 

more rarely at Eniwetok and Rongelap.

Family ANOMALINIDAE

Genus ANOMALINA D'Orbigny, 182C

Anomalina cf. A. glabrata Cushman

Plate 91, figure 23

Anomalina glabrata Cushman, Carnegie Inst. Washington Pub. 

342, p. 39, pi. 12, figs. 5-7, 1924.

A few specimens from numerous samples from all the 

atolls seem close to this species known from the South 

Pacific but these specimens are somewhat larger than 

the types and more coarsely punctate.

Anomalina polymorpha Costa 

Plate 91, figure 22

Anomalina polymorpha Costa, Accad. pontaniana Atti, v. 7, pt. 

2, p. 252, pi. 21, figs. 7-9, 1856.

A few specimens of this distinctive speeds described 

from the Pliocene of Italy and known from Kecent 

seas, were found in Bikini core 1174, more abundant at 

the base than at the top, and a single specimen from 

Bikini 1175.
TABLE 5.—Distribution of Globigerinidae, Globorotaliidae, Anomalinidae, Planorbulinidae, Rupertiidae, and Homotremidae in the Marshall Islands.

Family Globigerinidae

Globigerina bulloides

ci. G. conglomerata

Globigerina? sp.

Globigerinoides conglobata

elongata 

rubra

sacculifera 

sacculifera var. fistulosa

Globigerinella aequilateralis

Hastigerinella digitata

Orbulina universa

Pulleniatina obliquiloculata

Candeina nitida

Family Globorotaliidae

Globorotalia crassa

cf. G. hirsuta

menardii

menardii var. fimbriata

truncatulinoides

Family Anomalinidae

Anomalina cf . A. glabrata

polymorpha

Planulina ariminensis

Laticarinina pauperata

Cibicides cicatricosus

lobatulus

mayor!

cf. C. pseudoungerianus

Cibicidina sp.

Family Planorbulinidae

Planorbulinella larvata 

Acervulina inhaerens

Gypsina globula 

vesicular is

Family Rupertiidae

Rupertia stabilis

Carpenteria proteiformis ?

sp.

Family Homotremidae

Homotrema rubrum

Miniacina miniacea

Approximate grand total

RONGERIK LAGOON

0

9

99

O

4

43

O

1

4

3

1

1

3

415

COm

1

1

2

2

'I

1

1

679

•*

1 

1

It

618

OS

1

6

188

CO

2

3 

1

6

1

485

CO

1

2

9

«

3

488

a

2

1

1

12

6

t

978

2

4

1

3

394

3

3

4

1

£

429

O

3

6

2

11

587

in

6

3

J

10

335

3

4

1

377

c1

3

2

6

2

3

683

RONGELAP ATOLL,

Inside the lagoon

•*

m

CO

24

78

CO •*

1

1

750

1

2

B5£

>*

1

1

1

2

316

•* 

f

1

4

1

492

c1

5

1

6

3

2

470

O 

COf

1

427

CO

c2

1

4

4

430

05

2

i

i

i

513

s

2

1

1

5

3

1 

1

306

C*

1

5

3

821

O

1

1

2

48

£;

6

1

22

Outside 

the 

lagoon

CO

f

1

2

1

2

1

1

12

10 

1

5

1

748

i

i

2

4

11

1

4

18

512

•*

3

1

1

2

12

5

1

9

17

444.

«

CO 

*f

1

1

9

137

BIKINI ATOLL

beach

m

i

8

1

1

122

O

m

75

m

m

50

mo

m

94

•* 

c81

reef flat

m m

i

1

10

106

4

1

19

176

C*

m

7

69

co

m

X

m

i

i

5

90

m

i

100

m 

o

m

10

100

Ins

S

i

2

4

17

502

S

12

217

f

1

10

1

302

O

m

i

4

4

1

2

2

271

•*

6

244

CO

3

3

218

S

01

1

6

3

139

o

OT

m

1

5

134

1

2

4

3

217

i

4

218

«* 

B

3

2

359

c~

1

a

2

1

14

290

O 

f

T

33

i

2-

1

175

1

2

3

172

f

1

214

S

1

1

0 

f

CO

2

1

301

1

3

1

257

11

282

I

2

271

c-

,

1

2

17

1 

1

1

444

O

1

m 

o

5

2

2

399

1

1

2

2

149

5

2

9

1

3

73

0

10

1

1

391

3

1

1

9

1

33

0

5

1

2

3

1

592

i d e

t-

<*

2

6

4

2

623

0)

3

20

1

7

4

1

581

the lagoo

0 

CO•*

b

1

189

3

15

f 

f

•* 3

3

7

4

1

1

375

0

3

4

8

1

38

f 

in

4

4

3

1

13

3

286

n

3

5

1

51

O 

CO

4

10

1

3 

3

479

•*

to 

f

4

m

2

1 

1

214

1

1

2

2

4

3 

5

311

3

1

238

1

•I

1

2

8

1

585

O 

I

1

3

1

2

4

4

3

368

M

M

1

1

13

15

4

4

486

O 

0

1

9

CO 

O

2

3

9

1

534 212

•*

2

i

i

i

217

t>

2

1 

3.

178

•*

2

184

c-

•*

3

»

48

COM<

1

1

101

CM

1

19

1

3

423

*T 

*T

1

2

169

c1

8

1

1

2

1

3

344

f

2

2

32

m

4

4

3

2

8

1

264

m

1

22

1

59

S

2

2

1

258

f

•*

4

1

2

152

Outer slopes

Dredge 

samples

c~

i<

l

i

2

1

2

295

*T

9

li8-

3

2

1

2

10

287

CD 

f

1

1

1

1

4

2

1

JO

176

CO

f

O 

3

1

1

1

1

1

3

1

10

168

3

186

Bottom samples

CO

4

I

6

2

1

1

1

1

1

1

4

5

4

4

433

s

i

6

1

1

1 

1

3

63

CO

12

7

3

2

4

1

2

1

2

21

3

1

1

1 

2

3

384

•* 

CO

1

1

10

5

3

2

3

1

t

3

1

2

531

CO

CO

1

10

19

1

1

2

11

7

1

235

0

5

3

1

1

2

124

1

1

10

5

4

3

1

1

2

1

5

1

1

2

1

237

I

4

8

1

2

2 

1

3

176

c1

1

8

'01

2

28

CO

tO

3

2

10

1

3

2

1

2

225

CO

1

10

CO

12

6

0

1

35

8

10

1

1

4

3

7

1

2

21

3

1

1

1

470

1

10

2

1

193

Guyot

^

1

14

S

ti

10

11

2

H

1

2

8

11

4

11

2

782

CO 

c-^

15

1

15

1

2

6

1

5

12

1

4

1

481

"co

CO

30

S

15

1

1

8

4

2

7

2

1

431

O 

f

S

10

10

w

1

2

4

4

3

1

16

1

391

^r

0

f

6

7

15

9

8

b

14

7

B

3

426

S

2

18

to

£6

1

3

3

1

7

11

9

495

O

£j

16

S

M

2

1

1

3

1

385

m 

c~

i

ts

to

4

1

3

7

2

2

1

554

i

10

ts

S3

1

1

1

6

4

8

1

469

"•«•

m 

c~

10

I

i« 

1

1

b

1

1

12

553

>a« 

c~

2

to

ts

30

1

1

3

8

9

1

389

0

c~

2

6

23

15

S7 

2

1

CO

i

ts

S3

9

2l

1

5

347

3

10

477

CO

to

It

13

2

2

1

6

2

1

331

r-

£;

0 

S

1

2

4

<

li 

1

3

2

3

2

2

1

9

2

515

1

6

«0

to

10

1

1

3

2

1

242

O 

0

1

st

1 

s

10

1

3

7

1

336

O 

S

18

5

M

3

12

1

305

^J1

0 

C3

2

15

10

U

1

4

2

295

S 

0

t3

it

ts

ss

i

9

1

1

2

321

r1

14

ts

SS

8

13

5

4

244

ENIWETOK ATOLL

o

i

1

119

S

1

66

O 

«

to

18.

S

1

3

5

1

1

1

5

272

g

2

63

s

3

6

130

Inside the lagoon

CO

CO

3

283

cco

2

3

2 

3

514

CO

CO

3

4

1

12

423

s

3

6

254

0 

01

2

5

210

«

3

7

1

3 

2

11

451

to

2

2

2

8

1

644

CO

3

1

455

S

1

1

19

1

20

217

CO

1

93

3

i

2

4

4

361

1

2

2

2

69

1

1

1

3

en

CM

4

1

626

10

292

N

1

1

5

309

CM

2

1

i CM

5

1

53C 10 192

0

1

1

6

I

1

IN

2

1

2

2

3

679 826

O

m

1

3

1

12

13

482

^

1

1 

1

3

1

365

CO

1

2

1

1

1

593

O

3

2

1

4

8

5

793

•«< 

CM

1

30C

0 

0 

CO

1

5 

2

13

13

578

O 

CO

1

1

2

1

426

N

O

2

78

co

4

1

2

473

CO

2

4

1

3

585

53

1

1 

3

3

255

•*

1

3

5

19

659

S

1

3

6

13

10

325

a

2

1

1

461

3

i

7

4

171

0

70

•* 

0

6

5

4

5

624

O

2

2

1

2

2

28C

S

1

1

1

1

1 

1

2

376

S

1

3

7

3

626

01

•*

4

54

Outer 

slopes

S

3

3

1

4

1

2

B

2

493

3

i

6

1

1

1

1

1

1

2

4

5

10

384

CD

2

9

11

1

8

183

3

i

3

1

1 

1

01

2

1

i

15

St

15

3

2

3

1

1

1

23

3

1

4

14

12

2

2

38 534

291053 O - 54 (Face p. 370)
RECENT FORAMINIPERA OP THE MARSHALL ISLANDS 371

Genus PLANULINA D'Orbigny, 1826 

Planulina ariminensis D'Orbigny

Planulina ariminensis D'Orbigny, Annales sci. nat., tome 7, p.

280, no. ], pi. 5 (14), figs. 1-3 bis., Modeles, no. 49, iie livr.,

1826. 

Anomalina ariminensis H. B. Brady, Challenger Rept., Zoology,

v. 9, p. 674, pi. 93, figs. 10, 11, 1884.

A few well-preserved specimens occur in the guyot 

samples. The species was described from the Adriatic 

Sea and is known from the late Tertiary of the Mediterranean. It is widely recorded in Recent seas.

Genus LATICARININA Galloway and Wissler, 1927 

Laticarinina pauperata (Parker and Jones)

Pulvinulina repanda var. menardii, subvar. pauperata Parker and

Jones, Philosophical Trans., v. 155, p. 395, pi. 16, figs. 50, 51,

1865. 

Pulvinulina pauperata H. B. Brady, Challenger Rept., Zoology,

v. 9, p. 696, pi. 104, figs. 3-11, 1884. 

Laticarinina pauperata Cushman and Todd, Cushman Lab.

Foram. Research Contr., v. 18, p. 15, pi. 4, figs. 1-6, 1942.

A single nepionic individual was found at Bikini 1170.

Genus ANOMALINELLA Cushman, 1927

Anomalinella rostrata (H. B. Brady)

Plate 91, figure 24

Truncatulina rostrata H. B. Brady, Quart. Jour. Micros. Sci.,

v. 21, p. 65V 1881; Challenger Rept., Zoology, v. 9, p. 668, pi.

94, figs. 6a-c, 1884. 

Anomalinella rostrata Cushman, Cushman Lab. Foram. Research

Contr., v. 3, p. 93, 1927; idem, Special Pub. 1, p. 322, pi. 50,

fig. 1, 1928.

Test bilaterally symmetrical, involute, last-formed 

whorl consisting of about ten chambers, periphery 

keeled; chambers of uniform shape, increasing very 

slowly and evenly in size as added; sutures distinct, 

limbate, slightly curved; wall very coarsely perforate; 

aperture on ventral margin of last-formed chamber 

between periphery and umbilical area with a supplementary aperture just below peripheral margin, elongate, and parallel to axis of coiling. Length up to 1 

mm.

This is a rather common species in the Indo-Pacific 

region and occurs in a number of the samples.

Genus CIBICIDES Montfort, 1808

Cibicides cicatricosus (Schwager)

Plate 91, figures 25, 26

Anomalina cicatricosa Schwager, AToworo-Exped., Geol. Theil,

Band 2, p. 260, pi. 7, figs. 108, 4, 1866. 

Cushman, Geol. Soc. Japan Jour., v. 46, no. 546, p. 153 (43),

pi. 10 (6), figs. 19 a-c, 1939. 

Cibicides cicatricosus Thalmann, Eclogae geol. Helvetiae, tome

25, p. 309, 1932. 

Truncatulina akneriana H. B. Brady (not D'Orbigny), Challenger

Rept., Zoology, v. 9, p. 663, pi. 94, figs. 8a-c, 1884.

In a few deep-water samples from Eniwetok and 

Bikini, this species, described from the Pliocene of Kar 

Nicobar, occurs fairly commonly. It is a distinctly 

perforate species. About 10 chambers comprise the 

last-formed whorl, the dorsal side is only slightly 

evolute and the earlier whorls largely obscured by shell 

material, the sutures are limbate and distinct, and the 

aperture extends slightly over onto the dorsal side. 

In adult tests the wall is about equally perforate on 

both dorsal and ventral sides, but in young specimens 

the ventral side, or the early chambers on the ventral 

side, are almost free of the coarse perforations.

Cibicides lobatulus (Walker and Jacob) 

Plate 91, figures 27, 28

Nautilus lobatulus Walker and Jacob, Adams Essays, Kanmacher's ed., p. 642, pi. 14, fig. 36, 1798.

Truncatulina lobatula D'Orbigny, Foraminiferes fosstbs du 

bassin tertiaire de Vienne, p. 168, pi. 9, figs. 18-23, 1846.

Cibicides lobatulus Cushman, U. S. Natl. Mus. Bull. 104, pt. 8, 

p. 118, pi. 21, figs. 3a-c, 1931.

Truncatulina lobata D'Orbigny (in Barker-Webb and Berthelot), 

Histoire naturelle des lies Canaries, tome 2, pt. 2, Foraminiferes, p. 134, pi. 2, figs. 22-24, 1839.

This cosmopolitan species is found at Bikini, E'liwetok, and Rongelap, more abundantly at the deeper 

stations. It shows much variation and occurs in two 

forms, examples of each of which are figured. The 

difference in these two forms seems to be due to the 

attachment of the one (fig. 27) resulting in a flat or 

concave dorsal surface and consequently an acute 

periphery, while the other (fig. 28) shows no evidence 

of having been attached and the chambers on the dorsal 

side are slightly inflated, and the periphery smcothly 

rounded. This second, rounded form seems to be 

limited to the deeper-water samples and to be found 

only among the larger specimens while those showing 

evidence of attachment are found in either de?,p or 

shallow-water samples. Young specimens almost all 

show evidence of attachment. Both kinds shovr the 

closely spaced, coarse perforations on both dorsal and 

ventral surfaces, typical of the species.

Cibicides mayori (Cushman) 

Plate 91, figures 29, 30

Truncatulina mayori Cushman, Carnegie Inst. Washington Pub. 

342, p. 39, pi. 12, figs. 3, 4, 1924.

Test attached, flattened, planoconvex or with dorsal 

side slightly concave, not completely involute ventrally 

thus leaving a small knob of shell material in the center 

of the convex ventral side, periphery very irregular, 

with a very thin, fringe-like, sometimes serrate keel; 

chambers indistinct, numerous, 5 to 10 in the last 

whorl, later ones irregular in size and shape, very 

slightly inflated; sutures indistinct, limbate and curved.
372 BIKINI AND NEARBY ATOLLS, MARSHALL ISLANDS

slightly depressed on the ventral side; wall thin, 

smooth, perforations very coarse, less closely spaced on 

the ventral side than on the dorsal; aperture an elongate, very low and inconspicuous slit extending from 

near the umbo on the ventral side to the periphery and 

slightly over onto the dorsal side. Diameter 0.50 to 

0.75 mm, thickness 0.15 to 0.20 mm.

Specimens from Bikini, Eniwetok and Rongelap have 

been compared with the types of this species from off 

Samoa, in 17, 18, and 25 fathoms, and seem to be the 

same, although they exhibit considerable variation. 

The Marshall Islands specimens have more chambers 

to the last whorl, as many as 10, and only a few specimens have the perforations on the dorsal surface 

which, as shown by reexamination of the holotype, do 

not extend entirely through the test, but only in to the 

previous chambers. These perforations seem to be the 

unclosed dorsal portions of earlier apertures.

Cibicides cf. C. pseudoungerianus (Cushman) 

Plate 91, figure 31

Truncatulina pseudoungeriana Cushman, U. S. Geol. Survey 

Prof. Paper 129-E, p. 97, pi. 20, fig. 9, 1922.

Cibicides pseudoungeriana Cushman, U. S. Natl. Mus. Bull. 104, 

pt. 8, p. 123, pi. 22, figs. 3-7, 1931; Am. Jour. Sci., v. 239, 

p. 138, pi. 5, figs. 16-19, 1941.

A few specimens seem closely related to this species 

described from the Oligocene of Mississippi and widely 

recorded, fossil and Recent. They are smaller than 

the types and have 13 or 14 chambers in the last whorl. 

They have a more prominent umbilical boss on the 

ventral side than do the types, and the difference in 

coarseness of perforations between the ventral and 

dorsal sides is very striking.

Genus CIBICIDINA Bandy, 1949

Cibicidina sp. 

Plate 91, figure 32

A very few specimens from four Bikini deep-water 

samples, similar to that figured, seem to be undescribed. 

The species resembles C. mississippiensis (Cushman) 

(1935, p. 54, pi. 22, figs. 3a-c) in general structure. 

The chambers are very strongly inflated ventrally and 

the dorsal side is slightly concave and only very 

slightly evolute. The wall is distinctly perforate.

Genus CIBICIDELIA Cushman, 1927

Cibicidella variabilis (D'Orbigny)

Plate 82, figure 13

Truncatulina variabilis D'Orbigny, Annales sci. nat., vol. 7, 

p. 279, No. 8, 1826; (in Barker-Webb and Berthelot), Histoire naturelle des lies Canaries, tome 2, pt. 2, Foraminiferes, p. 135, pi. 2, fig. 29, 1839.

Parker, Jones, and H. B. Brady, Annals and Mag. Nat. History, 4th ser., v. 8, p. 177, pi. 12, fig. 138, 1871.

Cibicidella variabilis Cushman, Cushman Lab. Foram. Research 

Contr., v. 3, p. 93, 1927; idem, Special Pub. 1, p. 323, pi. 

50, figs. 5-7; pi. 51, fig. 5, 1928; U. S. Natl. Mus. Bull. 

104, pt. 8, p. 127, pi. 24, fig. 3, 1931.

This is a highly variable species, the eTly stages 

like Cibicides but the later chambers taking many 

different shapes, probably partly due to the surface 

to which it is attached. It is very widely recorded 

and more than one species may be included.

Family PLANORBULINIDAE 

Genus PLANORBUHNA D'Orbigny, 182^

Planorbulina acervalis H. B. Brady 

Plate 82, figure 14; plate 91, figures 34-36

Planorbulina acervalis H. B. Brady, Challenger Reit., Zoology, 

v. 9, p. 657, pi. 92, fig. 4, 1884.

Numerous specimens from all four atolls r.re referred 

to this species. It is evidently very variable and has 

been confused with Acervulina inhaerens Schultze. 

The two are very difficult to separate in some instances, 

and it is difficult to place the references with any 

degree of consistency. Usually it is possible to distinguish the initial few chambers in a regular coil on the 

dorsal side before the chambers assume the;r irregular 

manner of growth.

Genus PLANORBULINELLA Cushman, 1?97 

Planorbulinella larvata (Parker and Jones)

Planorbulina larvata Parker and Jones, Philos. Trans., v. 155,

p. 380, pi. 19, fig. 3, 1865.

H. B. Brady, Challenger Rept., Zoology, v. 9, p. 658, pi. 92, 

figs. 5, 6, 1884.

Planorbulinella larvata Cushman, Cushman Lab. Foram. Research Special Pub. 4, pi. 29, figs. 5a, b, 1933; idem, Special 

Pub. 5, pi. 37, figs. 3a, b, 1933.

This usually common species is noteworthy because 

of its rarity in the Marshall Islands. Only a single 

specimen, from Bikini core 1176 at 2 to 3 inches, was 

found.

Genus ACERVULINA Schultze, 1854

Acervulina inhaerens Schultze

Plate 91, figures 37, 38

Acervulina inhaerens Schultze, Organismus Polythal., 1854, p.

68, pi. 6, fig. 12. 

Bronn, Klassen und Ordnungen Thier-Reichs, Band 1, p.

72, pi. 6, fig. 17, 1859. 

Gypsina inhaerens H. B. Brady, Challenger Rept., Zoology, v. 9,

p. 718, pi. 102, figs. 1-6, 1884.

From an examination of the records of this species 

it seems that more than one species is recorded under 

this name. Specimens referred to it here are common in 

many of the samples and it is evidently a species of 

rather warm, shallow water.
RECENT FORAMINIFERA OP THE MARSHALL' ISLANDS 373

The separation between this species and Planorbulina 

acervalis is hard to make. In this species the later 

chambers are more piled up on top of the earlier chambers and in some cases the species is very distinctive 

in having a radial pattern of sutures on the ventral side. 

It has very small chambers, not inflated, and not as 

coarsely perforate as in Planorbulina acervalis.

Genus GYPSINA Carter, 1877

Gypsina globula (Reuss)

Plate 91, figure 39

Ceriopora globulus Reuss, Haidinger's Naturwiss. Abh., Band 2,

p. 33, pi. 5, fig. 7, 1847. 

Gypsina globulus H. B. Brady, Challenger Rept., Zoology, v. 9,

p. 717, pi. 101, fig. 8, 1884.

A few specimens may be placed under this name. 

When well preserved they are generally spherical in 

shape, although a number of immature ones have a 

flat surface on one side. The individual chambers are 

usually smaller than in Q. vesicularis, resulting in a 

finer textured surface.

Gypsina plana (Carter) 

Plate 93, figure 1

Polytrema planum Carter, Annals and Mag. Nat. History, 4th

ser., v. 17, p. 211, pi. 13, figs. 18, 19, 1876. 

Chapman, Linneari Soc. Jour., Zoology, v. 28, p. 209 (list);

p. 201, pi. 20, figs. 6, 7, 1900; Annals and Mag. Nat. History,

7th ser., v. 7, p. 82, 1901; Linnean Soc. Jour., Zoology, v. 28,

p. 396, pi. 35, figs. 2, 4; p. 407 (list), 1902. 

Gypsina inhaerens Schultze var. plana Yabe and Hanzawa,

Tohoku Imp. Univ. Sci. Repts., 2nd ser., Geol., v. 11, p. 179,

pi. 2, fig. 6, 1929; v. 14, p. 37, pi. 1, fig. 6; pi. 5, fig. 7; pi. 9,

fig. 12; pi. 10, fig. 8, 1930. 

Acervulina inhaerens Schultze var. plana Hanzawa, idem, v. 12,

p. 156, pi. 3, fig. 4, 1931. 

Acervulina inhaerens Schultze var. plana Carter var. Chapman,

Annals and Mag. Nat. History, llth ser., v. 11, p. 101 (list),

1944.

Rather large masses of this encrusting form are found 

attached to coral at a relatively shallow depth, 14 

fathoms, in Bikini Lagoon, and as a cementing coating 

on Halimeda segments at Bikini 788 also in the lagoon. 

It differs from Miniacina miniacea in consisting of a 

smooth, very fine textured coating over a large area 

rather than a more local development with branches 

extending upward from the surface. In cut section 

(pi. 93, fig. Ib) the structure is identical with that 

observed in cut sections of free specimens of Gypsina.

This species has been reported in Tertiary material. 

Chapman (1944) reported it as "one of the chief agents 

in binding together the loose reef material" in the 

Funafuti boring.

Gypsina vesicularis (Parker and Jones) 

Plate 82, figure 12

Orbitolina vesicularis Parker and Jones, Annals and Mag. Nat.

History, 3d ser., v. 6, p. 31, no. 5, 1860.

Gypsina vesicularis Carter, idem, 4th ser., v. 20, p. 173, 1877. 

H. B. Brady, Challenger Rept., Zoology, v. 9, p. 718, pi. 101,

figs. 9-12, 1884.

This species was found at all four atolls. Specimens 

are much larger and coarser textured than Q. globula 

and are disc-shaped, usually thickened in the center.

Family RUPERTIIDAE

Genus RUPERTIA Wallich, 1877

Rupertia stabilis Wallich

Plate 91, figure 33

Rupertia stabilis Wallich, Annals and Mag. Nat. History, 4th 

ser., v. 19, pp. 501-504, pi. 20, 1877.

A single young specimen, here figured, was found at 

Rongelap 456.

Genus CARPENTERIA Gray, 1858 

Carpenteria proteiformis Goes?

Carpenleria balaniformis Gray var. proteiformis Goes, K. svenska 

vetensk. akad. Hand!., Band 19, no. 4, p. 94, pi. 6, figs. 

208-214; pi. 7, figs. 215-219, 1882.

Carpenteria proteiformis Brady, Challenger Rept., Zoology, v. 9, 

p. 679, pi. 97, figs. 8-14, 1884.

Specimens consisting of an irregularly growing series 

of thick-walled, coarsely perforate chambers are fcTind 

fairly abundantly in a few samples. They are mostly 

light to dark red or orange.

Carpenteria sp. 

Plate 82, figure 15

The specimen figured was the only ones of this sort 

found. It may belong in C. monticularis Carter (1877, 

p. 211, pi. 13, figs. 9-12) which is known from the 

Pacific.

Family HOMOTREMIDAE 

Genus HOMOTREMA Hickson, 1911

Homotrema rubrum (Lamarck) 

Plate 82, figure 17; plate 92, figure 8; plate 93, figure 2

Millepora rubra Lamarck, Histoire naturelle des Animaux sans

Vertebres, tome 2, p. 202, 1816. 

Polytrema rubra Dujardin, Histoire naturelle des Zoophytes,

1841, p. 259. 

Carpenter, Parker, and Jones, Introd. Foram., p. 235, p1 . 13,

figs. 18-20, 1862.
374 BIKINI AND NEAEBY ATOLLS, MARSHALL ISLANDS

Homotrema rubrum Hickson, Linnean Soc. London Trans.,

Zoology, ser. 2, v. 14, pp. 445, 454, pi. 30, fig. 2; pi. 31,

fig. 9; pi. 32, figs. 19, 22, 28, 1911. 

Cushman, Carnegie Inst. Washington Pub. 311, p. 53, pi. 14,

figs. 6-8, 1922. 

Hofker, Siboga-Exped., Mon. 4, p. 31, pi. 13, figs. 8, 9; pi. 14,

figs. 12-20; pi. 15, 1927. 

Chapman and Parr, Royal Soc. Victoria Proc., v. 43, pt. 2,

p. 238, pi. 9, figs. 9, 10, 1931. 

Emiliani, Cushman Found. Foram. Research Contr., v. 2,

pp. 143-146, pis. 15, 16, 1951.

This is a species of shallow, warm waters, attached 

to shell or Halimeda or coral fragments. In our 

samples it is mostly represented by fragments evidently 

taken into deeper water by wave and current action. 

A few well-preserved specimens were found attached 

inside cavities. In these specimens (see plate 92, 

fig. 8, and plate 93, fig. 2) sponge spicules may be 

observed protruding from the high conical apertures.

Genus MINIACINA Galloway, 1933 

Miniacina miniacea (Pallas)

Plate 82, figure 16

Millepora miniacea Pallas, Elenchus Zoophytorum, p. 251, The

Hague, 1766. 

Esper, Die Pflanzenthiere, Theil 1, p. 225, pi. 17, figs. 1-4,

1791. 

Polytrema miniacea Carter, Annals and Mag. Nat. History, 4th

ser., v. 17, p. 185, pi. 13, figs. 1-6, 1876. 

Moebius, Foraminiferen von Mauritius, p. 85, pi. 7, figs. 1-17,

1880. 

H. B. Brady, Challenger Kept., Zoology, v. 9, p. 721, pi. 100,

figs. 5-9; pi. 101, fig. 1, 1884. 

Chapman, Linnean Soc. Jour., Zoology, v. 28, p. 16, pi. 4,

fig. 7, 1899. 

Hickson, Linnean Soc. London Trans., Zoology, ser. 2, v. 14,

p. 453, pi. 30, fig. 1; pi. 31; fig. 8, 1911. 

Heron-Alien and Earland, Soc. Sci. Hist. Nat. Corse Bull.,

p. 139, pi. 2, figs. 40-49, 1922. 

Hofker, Sibo'ga-Exped., Mon. 4, p. 27, pi. 12; pi. 13, figs. 1-7;

pi. 14, figs. 1-11, 1927. 

Miniacina miniacea Cushman, Cushman Lab. Foram. Research

Special Pub. 5, pi. 37, figs. 33-36, 1933. 

Colom, Inst. Espanol Oceanografia, Notas y Resumenes, ser. 2,

no. 108, p. 45, pi. 11, figs.'218, 220-222, 1942.

This species, red in color, occurs at all four atolls 

and is distinguished from Homotrema rubrum (Lamarck) 

by having fine perforations throughout, whereas in 

Homotrema the walls between the foramina are solid 

and thus appear glossy. Also, M. miniacea is a 

slenderer and more branching form. Numerous fragments show the enlarged disclike area by which the 

specimens are attached.

Some specimens lack the red color and may belong 

to the variety alba (Carter) (1877, p. 213, pi. 13, fig. 14).

BIBLIOGRAPHY

Asano, Kiyoshi, 1950-51, Illustrated catalogue of Japanese 

Tertiary smaller Foraminifera—Part 1, Nonionidae; part 2, 

Buliminidae; part 3, Textulariidae; part 4, Valvulinidae; 

part 5, Verneuilinidae; part 6, Miliolidae; part 7, Cassidulinidae; part 8, Polymorphinidae; part 9, Ophthalmidiidae; part 10, Lituolidae; part 11, Trochamminidae; part 12, 

Chilostomellidae; part 13, Anomalinidae; part 14, Rotaliidae; part 15, Lagenidae: Compiled and edited by Leo W. 

Stach. Tokyo, Japan.

Bagg, R. M., 1908, Foraminifera collected near the Hawaiian 

Islands by the steamer Albatross in 1902: U. S. Natl. 

Mus. Proc., v. 34, p. 113-172, pi. 5.

van den Bold, W. A., 1946, Contribution to the study of 

Ostracoda with special reference to the Tertiary and 

Cretaceous microfauna of the Caribbean region: Univ. 

Utrecht Thesis, p. 1-167, pis. 1-18, maps.

Brady, H. B., 1884, Report on the Foraminifera dredged by 

H. M. S. Challenger, during the years 187?-76: Reports 

of the Scientific Results of the Voyage of H. M. S. Challenger, 

v. 9 (Zoology), London, 814 p., 115 pis.

Carter H. J., 1877, On the locality of Carpenteria balaniformis, 

with description of a new species, and other Foraminifera 

found in and about Tubipora musica: Annals and Mag. Nat. 

History, 4th ser., v. 19, p. 209-219, pi. 13.

Chapman, Frederick, 1895, On some Foraminifen. obtained by 

the Royal Indian Marine Survey's S. S. Investigator, from 

the Arabian Sea near the Laccadive Islands: Zool. Soc. 

London Proc., p. 3-55, pi. 1.

———— 1900a, On some new and interesting Forr minifera from 

the Funafuti Atoll, Ellice Islands: Linnean Soc. Jour., 

Zoology, v. 28, p. 1-27, pis. 1-4.

———— 1900b, Foraminifera from the lagoon at Funafuti: 

tinnean Soc. Jour., Zoology, v. 28, p. 161-210, pis. 19, 20.

——— 1902, On the Foraminifera collected round the Funafuti 

Atoll from shallow and moderately deep water: Linnean 

Soc. Jour., Zoology, v. 28, p. 379-433, pis. 35, 36.

———— 1906, On some Foraminifera and Ostracoda obtained 

off Great Barrier Island, New Zealand: New Zealand Inst. 

Trans., v. 38, 1905, p. 77-112, pi. 3.

1907, Recent Foraminifera of Victoria; some littoral

gatherings: Quekett Micros. Club Jour., v. 10, p. 117-146, 

pis. 9, 10.

———— 1910, On the Foraminifera and Ostracoda f~om soundings 

(chiefly deep-water) collected round Funafuti by H. M. S. 

Penguin: Linnean Soc. Jour., Zoology, v. 30, p. 388-444, 

pis. 54-57.

———— 1912, Report on a sample of Globigerina ooze from 1122 

fathoms, east of Tasmania: Zool. Results Endeavour, 

1909-10, pt. 3, p. 309-311.

———— 1915, Report on the Foraminifera and Ostrr.coda obtained 

by the F. I. S. Endeavour from the east coast of Tasmania, 

and off Cape Wiles, South Australia: Biol. Results Endeavour, 

1909-14, p. 1-51, pis. 1-3.

———— 1922, Sea-floor deposits from soundings: Australasian 

Antarctic Exped. 1911-14, Sci. Repts., ser. .A. v. 2, Oceanography, pt. I, p. 1-60, pis. 1-3.

———— 1941, Report on foraminiferal soundings and dredgings 

of the F. I. S. Endeavour along the continental shelf of 

the south-east coast of Australia: Royal Soc. South Australia 

Trans., v. 65, pt. 2, p. 145-211, pis. 7-9.
RECENT FORAMINIFERA OF THE MARSHALL ISLANDS 375

Cushman, J. A., 1910, A monograph of the Foraminifera of the 

North Pacific Ocean. Part 1. Astrorhizidae and Lituolidae: 

U. S. Natl. Mus. Bull. 71, pt. 1, p. i-xiv, 1-134, text figs. 

1-203.

———— 1911, A monograph of the Foraminifera of the North 

Pacific Ocean. Part 2. Textulariidae: U. S. Natl. Mus. 

Bull. 71, pt. 2, p. i-xiii, 1-108, text figs. 1-156.

———— 1913, A monograph of the Foraminifera of the North 

Pacific Ocean. Part 3. Lagenidae: U. S. Natl. Mus. Bull. 

71, pt. 3, p. i-ix, 1-125, pis. 1-47.

———— 1914, A monograph of the Foraminifera of the North 

Pacific Ocean. Part 4. Chilostomellidae, Globigerinidae, 

Nummulitidae: U. S. Natl. Mus. Bull. 71, pt. 4, p. i-vi, 

1-46, pis. 1-19.

———— 1915, A monograph of the Foraminifera of the North 

Pacific Ocean. Part 5. Rotaliidae: U. S. Natl. Mus. Bull. 71, 

pt. 5, p. i-vii, 1-87, pis. 1-31, text figs. 1-62.

———— 1917, A monograph of the Foraminifera of the North 

Pacific Ocean. Part 6. Miliolidae: U. S. Natl. Mus. Bull. 71, 

pt. 6, p. i-vii, 1-108, pis. 1-39, text figs. 1-52.

———— 1918, The Foraminifera of the Atlantic Ocean. Part 1. 

Astrorhizidae: U. S. Natl. Mus. Bull. 104, pt. 1, p. i-vii, 

1-111, pis. 1-39.

———— 1919, Some Pliocene and Miocene Foraminifera of the 

coastal plain of the United States: U. S. Geol. Survey Bull. 

676, 1918, p. 1-100, pis. 1-31.

———— 1921, Foraminifera of the Philippine and adjacent seas: 

U. S. Natl. Mus. Bull. 100, v. 4, p. 1-608, pis. 1-100, text 

figs. 1-52.

———— 1922, Shallow-water Foraminifera of the Tortugas 

region: Carnegie Inst. Washington Pub. 311, V. 17, p. 1-85, 

pis. 1-14.

———— 1926, Foraminifera of Tropical Central Pacific: Bernice P. 

Bishop Mus., Bull. 27, Tanager Exped. Pub. 1, 1925, p. 

121-144.

———— 1927, Recent Foraminifera from off the west coast of 

America: Scripps Inst. Oceanography Bull., Tech. Ser., 

v. 1, no. 10, p. 119-188, pis. 1-6.

———— 1932, The Foraminifera of the Tropical Pacific collections 

of the Albatross, 1899-1900. Part I. Astrorhizidae to 

Trochamminidae: U. S. Natl. Mus. Bull. 161, pt. 1, p. i-vi', 

1-88, pis. 1-17.

———— 1933, The Foraminifera of the Tropical Pacific collections 

of the Albatross, 1899-1900. Part II. Lagenidae to 

Alveolinellidae: U. S. Natl. Mus. Bull. 161, pt. 2, p. i-vi, 

1-79, pis. 1-19.

———— 1934, Smaller Foraminifera from Vitilevu, Fiji: Bernice P. 

Bishop Mus. Bull. 119, p. 102-142, pis. 10-18.

———— 1935, Upper Eocene Foraminifera of the southeastern 

United States: U. S. Geol. Survey Prof. Paper 181, 88 p., 

23 pis.

———— 1936, New genera and species of the families Verneuilinidae and Valvulinidae and of the subfamily Virgulininae: 

Cushman Lab. Foram. Research Special Pub. 6, p. 1-71, 

pis. 1-8.

———— 1937, A monograph of the subfamily Virgulininae of the 

foraminiferal family Buliminidae: Cushman Lab. Foram. 

Research Special Pub. 9, 228 p., 24 pis.

———— 1942, The Foraminifera of the Tropical Pacific collections 

of the Albatross, 1899-1900. Part III. Heterohelicidae 

and Buliminidae: U. S. Natl. Mus. Bull. 161, pt. 3, p. i-v, 

1-67, pis. 1-15.

Cushman, J. A., 1947, New species and varieties of Foraminifera 

from off the southeastern coast of the United States: Cushman Lab. Foram. Research Contr., v. 23, p. 86-92, pis. 18-20.

———— 1948, Foraminifera, their classification and economi-5 use.

4th edition, revised and enlarged, with an illustrated key to

the genera: Harvard Univ. Press, Cambridge, Mas*?., p.

i-viii, 1-605, 86 pis., 9 text figs. 

Cushman, J. A., and Hanzawa, Shoshiro, 1936, New gener 1* and

species of Foraminifera of the late Tertiary of the Pacific:

Cushman Lab. Foram. Research Contr., v. 12, p. 45-48,

pi. 8. 

Cushman, J. A., and McCulloch, Irene, 1939, A report on some

some arenaceous Foraminifera: Allan Hancock IHcific

Exped., v. 6, no. 1, p. 1-113, pis. 1-12.

———— 1940, Some Nonionidae in the collections of the Allan 

Hancock Foundation: Allan Hancock Pacific Exped., v. 6, 

no. 3, p. 145-178, pis. 17-20.

———— 1942, Some Virgulininae in the collections of the Allan 

Hancock Foundation: Allan Hancock Pacific Exped., v. 6, 

no. 4, p. 179-230, pis. 21-28.

———— 1948, The species of Bulimina and related genera in the 

collections of the Allan Hancock Foundation: Allan Hancock 

Pacific Exped., v. 6, no. 5, p. 231-294, pis. 29-36.

1950, Some Lagenidae in the collections of the Allan

Hancock Foundation: Allan Hancock Pacific Exped., v. 6,

no. 6, p. 295-364, pis. 37-48. 

Cushman, J. A., and Parker, F. L., 1947, Bulimina and related

foraminiferal genera: U. S. Geol. Survey Prof. Paper 210-D,

p. 55-176, pis. 15-30. 

Cushman, J. A., and Stainforth, R. M., 1945, The Foraminifera

of the Cipero marl formation of Trinidad, British West

Indies: Cushman Lab. Foram. Research Special Pub. 14,

p. 1-75, pis. 1-16. 

Cushman, J. A., and Todd, Ruth, 1944, The genus SpirolotuUna

and its species: Cushman Lab. Foram. Research Special

Pub. 11, p. 1-82, pis. 1-9. 

Dakin, W. J., 1906, Report on the Foraminifera collected by

Prof. Herdman at Ceylon in 1902: Ceylon Pearl Oyster

Fish. Rept., v. 5, p. 225-242, 1 pi. 

D'Orbigny, A. D., 1826, Tableau methodique de la classe des

Ce"phalopodes: Annals des Sci. Naturelles, tome 7, p, 245-

314, pis. 10-17.

———— 1839a, Foraminiferes, in Ramon de la Sagra, Histoire 

physique, politique et naturelle de 1'Ile de Cuba: Paris, 

p. xlviii, 1-224, 12 pis.

———— 1839b, Voyage dans 1'Ame'rique Me"ridionale, Foraminiferes: Paris and Strasbourg, tome 5, pt. 5, p. 1-86, 9 pis.

——— 1839c, Foraminiferes, in Barker-Webb and Berfthelot 

Histoire naturelle des lies Canaries: Paris, tome 2, pt. 2, 

p. 119-146. 3 pis.

———— 1846, Foraminiferes fossiles du Bassin tertiaire de Vienne: 

Paris, 312 p., 21 pis.

1849, 1850, 1852, Prodrome de pale"ontologie stratigraphique universelle des animaux mollusques et rayonnfe:

Paris, 3 volumes: v. 1, 1849, p. 161, 242, 293, 324; v. 2,

1850, p. 41, 95, 110, 143, 185, 210, 279, 334, 407, 427; v. 3,

1852, p. 151, 190. 

Egger, J. G., 1893, Foraminiferen aus meeresgrundp'oben,

gelothet von 1874 bis 1876 von S. M. Sch. Gazelle: K. bayer.

Akad. Wiss., Math.-naturh. Abt., Abh., Kl. 2, Bard 18,

p. 195-458, pis. 1-21. 

Fornasini, Carlo, 1895, Cristellaria cleridi n. sp.: Riv. Italiana

Paleontologia, Bologna.
376 BIKINI AND NEARBY ATOLLS, MARSHALL ISLANDS

Goddard, E. J., 1907, The results of deep-sea investigation in 

the Tasman sea. L. The expedition of H. M. C. S. Miner.: 

Records Australian Mus., v. 6, 1905-1908, p. 305-311, text 

figs. 44-48.

———— and Jensen, H. I., 1907, Contributions to a knowledge of 

Australian Foraminifera. Part 2: Linnean Soc. New 

South Wales Proc., v. 32, p. 291-318, pi. 6.

Hada, Yoshine, 1931, Report of the biological survey of Mutsu 

Bay. 19. Notes on the Recent Foraminifera from Mutsu 

Bay: Tohoku Imp. Univ. Sci. Repts., 4th ser., Biology, 

v. 6, no. 1, p. 45-148, text figs. 1-95.

Hanzawa, Shoshiro, 1948> A comparative study of the fossil 

foraminiferal fauna of the Ryukyu Islands: Report of the 

committee on a treatise on marine ecology and paleoecology, 

1946-1947, no. 7, p. 77-88.

———— 1951, Recent and fossil Cydoclypeus from the Ryukyu 

Islands and their adjacent sea: Short Papers from the Inst. 

Geology Paleontology, Tohoku Univ., Sendai, no. 3, p. 1-12, 

pis. 1, 2, text figs. 1-10.

Heron-Alien, Edward, and Earland, Arthur, 1914-1915, The 

Foraminifera of the Kerimba archipelago (Portuguese East 

Africa): Zool. Soc. London Trans., v. 20, pt. 12, 1914, p. 

363-390, pis. 35-37; pt. 17, 1915, p. 543-794, pis. 40-53.

———— 1924, The Foraminifera of Lord Howe Island, South 

Pacific: Linnean Soc. Jour., Zoology, v. 35, p. 599-647, 

pis. 35-37.

1928, On the Pegididae, a new family of Foraminifera:

Royal Micros. Soc. Jour., v. 48, p. 283-299, pis. 1-3.

Hickson, S. J., 1911, On Polylrema and some allied genera: 

Linnean Soc. London Trans., v. 14, pt. 3, p. 443-462, 

pis. 30-32, 1 text fig.

Hofker, J., 1927, The Foraminifera of the Siboga Expedition. 

Part I, Families Tinoporidae, Rotaliidae, Nummultidae, 

Amphisteginidae: Siboga-Exped., Mon. 4, p. 1-78, pis. 1-38.

Jensen, H. L, 1905, Contributions to a knowledge of Australian 

Foraminifera: Linnean Soc. New South Wales Proc., 1904, 

pt. 4, p. 810-831, pi. 23.

Jepps, M. W., 1953, Nuclei of Cydoclypeus carpenteri Brady: 

Nature, v. 171, p. 1114, 3 text figs.

Johnson, M. W., 1949, Zooplankton as an index of water exchange between Bikini Lagoon and the open sea: Am. 

Geophys. Union Trans., v. 30, no. 2, p. 238-244, text 

figs. 1-15.

Karrer, Felix, 1868, Die Miocene Foraminiferen-fauna von 

Kostej ini Banat: K. Akad. Wiss. Wien Sitzungsber., 

Band 58, Heft 1, p. 111-193, pis. 1-5.

Lalicker, C. G., and McCulloch, Irene, 1940, Some Textulariidae 

of the Pacific Ocean: Allan Hancock Pacific Exped., v. 6, 

no. 2, p. 115-143, pis. 13-16.

Lister, J. J., 1895, Contributions to the life-history of the Foraminifera: Philos. Trans., v. 186, p. 401-453, pis. 6-9.

Marie, Pierre, 1941, Sur la faune de Foraminiferes des depots 

littoraux actuels de la mer Rouge et de Djiboute: Soc. Linn. 

Normandie M6m., n. ser., tome 1, pt. 1, p. 53-71.

Martinotti, Anna, 1920, Foraminiferi della spiaggia di Tripoli: 

Soc. Italiana Sci. Nat. Atti, v. 59, p. 249-334, pis. 10-13.

Millett, F. W., 1898-1904, Report on the Recent Foraminifera 

of the Malay archipelago contained in anchor-mud collected 

by Mr. A. Durrand, F. R. M. S.: Royal Micros. Soc. Jour., 

1898, p. 258-269, pis. 5, 6; p. 499-513, pis. 11, 12; p. 607-614, 

pi. 13; 1899, p. 249-255, pi. 4; p. 357-365, pi. 5; p. 557-564, 

pi. 7; 1900, p. 6-13, pi. 1; p. 273-281, pi. 2; p. 539-549, 

pi. 4; 1901, p. 1-11, pi. 1; p. 485-497, pi. 8; p. 619-628, 

pi. 14; 1902, p. 509-528, pi. 11; 1903, p. 253-275, pi. 5; 

p. 685-704, pi. 7; 1904, p. 489-506, pi. 10; p. 597-609, pi. 11.

Moebius, Karl, 1880, Foraminiferen von Mauritius, in Moebiusr 

F. Richfcer, and E. von Mertens, Beitrage zur Meeresfauna 

der Insel Mauritius und der Seychellen: Berlin, p. 63-110r 

pis. 1-14.

Morishima, Masao, 1948a, The accumulation o* foraminiferal 

tests in inlets of Wakasa Bay on the Island Sea of Japan: 

Report of the committee on a treatise on marine ecology 

and paleoecology, 1946-1947, no. 7, p. 89-91.

————, 1948b, Foraminiferal thanatocoenoses of Ago Bay, Kii 

Peninsula, Japan: Report of the committee on a treatise on 

marine ecology and paleoecology, 1947-H48, no. 8, p. 

111-117, 2 charts.

Myers, E. H., 1942, Ecologic relationships of some Recent and1 

fossil Foraminifera: Report of the committee on marine 

ecology as related to paleontology 1941-1942, no. 2, p. 31-36.

———— 1943, Ecologic relationships of larger Foraminifera: 

Report of the committee on marine ecology as related to 

paleontology, 1942-1943, no. 3, p. 26-30.

Oinomikado, Tuneteru, and Stach, L. W., 1948, Recent shallow 

water foraminiferal assemblages from Niigrta Prefecture, 

Japan: Report of the committee on a treatise on marine 

ecology and paleoecology 1947-1948, no. 8, p. 104-110.

Palmer, D. K., 1934, The foraminiferal genus Gwmbelina in the 

Tertiary of Cuba: Soc. Cubana Hist. Nat. Mem., v. 8r 

no. 2, p. 73-76, text figs. 1-8.

Parker, F. L., 1948, Foraminifera of the continental shelf from 

the Gulf of Maine to Maryland: Mus. Comp. Zoology 

Bull., v. 100, no. 2, p. 213-253, pis. 1-7, text figs. 1-4, 

tables 1-10.

Parker, W. K., and Jones, T. R., 1863, On the nomenclature of 

the Foraminifera 10—The species enumerated by D'Orbigny 

in the Annales des Sciences Naturelles, tome vii, 1826: 

Annals Mag. Nat. History, ser. 3, v. 12, p. 429-441.

Parker, F. L., Jones, T. R., and Brady, H. B., 1865a, On the 

nomenclature of the Foraminifera, 11—The species enumerated by Batsch in 1791: Annals Mag. Nat. History, ser. 3r 

v. 15, p. 225-232.

———— 1865b, On the nomenclature of the Foraminifera, 12— 

The species enumerated by D'Orbigny in tl •? Annales des 

Sciences Naturelles, tome vii, 1826—(3) Th-? species illustrated by models: Annals Mag. Nat. History, ser. 3, v. 16, 

p. 15-41, pis. 1-3.

1871, On the nomenclature of the Foraminifera, 14—

The species enumerated by D'Orbigny in tl 3 Annales desSciences Naturelles, tome vii, 1826—(4) The species founded 

upon the figures in Soldani's Testaceographir, ac Zoophytographia: Annals Mag. Nat. History, ser. 4, v. 8, p. 145-179, 

238-266, pis. 8-12.

Sidebottom, Henry, 1904-1909, Report on the Recent Foraminifera from the coast of the island of Delos (Grecian archipelago). Parts I-VI: Manchester Lit. Philos. Soc. Mem. 

and Proc., v. 48, pt. 2, 1904, p. 1-26, pis. 2-5; v. 49, pt. 2, 

1905, p. 1-22, pis. 1-3; v. 50, pt. 2, 1906, p. 1-18, pis. 1, 2; 

v. 51, pt. 3, 1907, p. 1-28, pis. 1-4; v. 52, pt. 3 1908, p. 1-28, 

pis. 1-5; v. 53, pt. 3, 1909, p. 1-32, pis. 1-5.

———— 1910a, Report on the Recent Foraminifera from the Bay 

of Palermo, Sicily, 14-20 fathoms. (OS the harbour): 

Manchester Lit. Philos. Soc. Mem. and Proc., v. 54, pt. 3, 

p. 1-36, pis. 1-3.

———— 1910b, Two new species of Cassidulina: Quekett Micros^ 

Club Jour., v. 11, p. 105-108, pi. 4.

———— 1912-1913, Lagenae of the south-west Pacific Ocean. 

From soundings taken by H. M. S. Wa'erwitch, 1895:: 

Quekett Micros. Club Jour., v. 11, 1912, p. 375-434, pis.. 

14-21; v. 12, 1913, p. 161-210, pis. 15-18.
RECENT FOEAMINIFEEA OF THE MARSHALL ISLANDS 377

Sidebottom, Henry, 1918, Report on the Recent Foraininifera

dredged off the east coast of Australia. H. M. S. Dart, station

19 (14 May 1895), Lat. 29°22' S., Long. 153°51' E., 465

fathoms—Pteropod ooze: Royal Micros. Soc. Jour., p. 1-25,

121-153, 249-264, pis. 1-6. 

Silvestri, Alfredo, 1903, Forme nuove o poco conosciute di

Protozoi Miocenici Pieinontesi: Accad. Sci. Torino Atti,

v. 39, p. 1-14, text figs. 1-7. 

Soldani, Ambrogio, 1780, Saggio orittografico ovvero osservazioni

sopra le terre nautilitiche ed ammonitiche della Toscana:

Sienna, 146 p., 25 pis. 

Parr, W. J., 1945, Recent Foraminifera from Barwon Heads,

Victoria: Royal Soc. Victoria Proc., v. 56 (new ser.), pt. 2,

p. 189-227, pis. 8-12, figs. 1, 2 (maps). 

Reuss, A. E., 1864, Beitrage zur Kenntniss der tertiaren foraminiferenfauna—3, Die Foraminiferen des Septarienthones

von Offenbach; 4, Die Foraminiferen des Septarienthones

von Kreuznach: K. Akad. Wiss. Wien Sitzungsber., Band

48, pt. 1, 1863, p. 36-69, pis. 1-8. 

Rhumbler, Ludwig, 1902, Die Doppelschalen von Orbitolites und

anderer Foraminiferen, vom entwicklungsmechanischen

Standpunkt aus betrachtet: Archiv. fiir Protistenkunde,

Band 1, p. 193-296, pis. 7, 8. 

Said, Rushdi, 1949, Foraminifera of the northern Red Sea:

Cushman Lab. Foram. Research Special Pub. 26, 44 p., 4 pis. 

———— 1950a, Additional Foraminifera from the northern Red

Sea: Cushman Found. Foram. Research Contr., v. 1, pts.

1, 2, p. 4-9, pi. 1.

Said, Rushdi, 1950b, The distribution of Foraminifera in the 

northern Red Sea: Cushman Found. Foram. Research 

Contr., v. 1, pts. 1, 2, p. 9-29, distribution table.

Schultze, M. S., 1854, tJber den organismus der PolythaJamien 

(Foraminiferen), nebst bemerkungen iiber die Rhizcnoden 

im allgemeinen: Leipzig, 68 p., 7 pis.

Seguenza, Giuseppe, 1862, Descrizione del Foraminiferi monotalamici delle marne Mioceniche del distretto di Messina: 

Messina, 84 p., 2 pis.

———— 1880, Le formazioni Terziarie nella Provincia di Reggio 

(Calabria): R. Accad. Lincei Atti, ser. 3, v. 6, p. 1-446, 

pis. 1-17.

Walker, George, and Boys, William, 1784, Testacea rainuta 

rariora, nuperrime detecta in arena littoris Sandvicensis: a 

Gul. Boys, arm. S. A. S. Multa addidit, et omnium figuras 

ope microscopii ampliatas accurate delineavit Geo. Vralker: 

London.

Williamson, W. C., 1848, On the Recent British species of the 

genus Lagena: Annals Mag. Nat. History, ser. 2, v. 1, p. 

1-20, pis. 1, 2.

———— 1858, On the Recent Foraminifera of Great F-itain:

Ray Soc., London, 107 p., 7 pis. 

Yabe, Hisakatsu and Hanzawa, Shoshiro, 1926, Globigerina

ooze from the sea lying south of Okinawa-Jima (The Riukiu

Islands): Japanese Jour. Geology and Geography, v. 4,

1925, p. 47-54, 1 text fig.

INDEX

<tt~breviata, Bolivina.. — ........... 321, 352, tab. 3, pi. 87

Bolivina limbata-.---.-..--...-......--------- 352

Textularia.-.........—..................... 329

abyssorum, Nodosaria.-. —————— --------- — — 356

Siphonodosaria................ 322, 366, tab. 3, pi. 88

acer.a.is, Planorbulina..... 324, 37%, 373, tab. 5, pis. 82,91

Acereulina inhaeiens.--- _____ 324, 372, tab. 5, pi. 91

inhaetensplana....... .............————.— 373

acescata, Spiroloculina.....---. ———_ 324, 334, tab. 1

Spirolocuhna grateloupi----......... ——— -__ 334

acicularis, Nautilus (iiiMMs) --------------------- 348

Spirolina---................. 323, 848, tab. 3, pi. 87

aculeata, Bulimina.-.—-------- 322, 350, tab. 3, pi. 87

acuticosta, Lagena,..—.-.—...... _ 321, 344, tab. 2, pi. 86

advena, Patellina.............. .—.__ 324, 357, tab. 4

PolystomeUa-.........- .....———. —— —— -_ 345

attiformis, Patellina.......... 324, 357, tab. 4, pi. 89

advenum, Elphidium...— — - 323, 345, 346, tab. 2, pi. 86 

advenum dispar, Elphidium... 323, 345, 346, tab. 2, pi. 86 

aequa, Spiroloculina_.——. —— __ 324, 334, tab. 1

Spiroloculina antittarum...... ......—— -.__- 334

aeguilateralis, Globigerina................... ...--__ 369

Globigerinella.....--—... 322, 326, 369, tab. 5, pi. 91

affinis, Bulimina.............. .__ 322, 350, tab. 3, pi. 87

agglutinans, Miliolina anguina..................... 332

Quinqueloculina-------...-------- — _ —..__ 332

anguina-.....--. ...... ——..- —— — _____ 332

Textularia..-............. 323, 328, 329, tab. 1, pi. 83

akneriana, Truncatulina... — — -------_--.._ 371

alba, Miniacina....--..---......-................. 374

albatrossi ornata, Angulogerina...- 321, 356, tab. 3, pi. 88 

allomorphinoides, Discorbis....................... 361

alternans, Cassidulina.............. —— „.. ——.- 366

attiformis, Patellina advena,....-- 324, 357, tab. 4, pi. 89

Alveolinc pulchra............—...... ——— - — — 348

Alveolinellidae_______------_______.. 348

alveoliniformis, Miliolina.--.-- ——...—————.. 337

Schlumbergerina........ .__ 323, 337, tab. 1, pi. 85

Ammodiscidae—..._-------——.--_____.. 328

Ammodiscoides.............__..______.. 368

Ammodiscus—— —— —————— — ———— __ 368

incertus........__....——....... 324, 328, tab. 1

ammonoides, Nautilus............................ 346

Operculina—...— —........ 323, 346, tab. 3, pi. 87

Amphisorus.............................___— 348

hemprichii.............................___.. 348

Amphistegina——....———————_————— 319, 330

lessonii.....-............ ...— ——————— — 362

madagascariensis....—......... .....,-..- — — 319,

320, 321, 323, 325, 326, 368, tab. 4, pi. 90

radiata..-— 321, 322, 326, 368, 363, tab. 4, pi. 90

radiata papittosa............ 322, 362, tab. 4, pi. 90

venosa............__ 322, 363, tab. 4, pi. 90

var—.....................____ 363, tab. 4, pi 90

Amphisteginidae...... ———————_—.——-- 362

amputtacea, Uvigerina............ 321, 355, tab. 3, pi. 88

Uvigerina asperula.......................... .. 355

anguina agglutinans, Miliolina.................... 332

agglutinans, Quinqueloculina-................. 332

arenata, Quinqueloculina.-.—. 320, 332, tab. 1, pi. 83 

angularis, Clavulina...........— 324, 332, tab. 1, pi. 83

angulata, Gaudryina triangularis.- 321, SS1, tab. 1, pi. 83

Spiroloculina............._. 323, 334, tab. 1, pi. 84

antittarum................................ 334

grata.................................—... 334

Angulogerina albatrossi ornata.... 321, 356, tab. 3, pi. 88

angulosa, Cassidulina............. 322, 365, tab. 4, pi. 90

Anomalina ariminensis........................... 371

cicatricosa..................................... 371

glabrata....................... 324, 370, tab. 5, pi. 91

polymorpha..................- 322, 370, tab. 5, pi. 91

Anomalinetta rostrata............ 324, 371, tab. 5, pi. 91

Anomallnidae——-...._____..________ 370

291053—54——5

[Italic numbers indicate descriptions]

antiUarum, Spiroloculina.-—........ ....—..... __ 335

aequa, Spi}oloculina.... ...——— ——— —— —— 334

angulata, Spiroloculina..-....... —————— -. 334

arenata, Quinqueloculina anguina- 320, SSi, tab. 1, pi. 83 

arietina, Spirolina................ 323, 348, tab. 3, pi. 87

arietinus, Nautilus (Lituus)....................-. 348

Peneroplis.......................-...........- 348

ariminensis, Anomalina--.-......-..---..-------- 371

Planulina......................— 322, 37^, tab. 5

Articulina elongata.................... 324, 336, tab. 1

funalis........................................ 336

mucronata.....------------------------------ 336

pacifica,—-.................... 323,336, tab. 1, pi. 84

asperulaampullacea, Uvigerina......--------....-- 355

Astacolus...---........-----------.----------....- 345

bradyL..-----................. 321, 343, tab. 2, pi.

planulata.................... 321, 343, tab. 2, pi. 86

planitlatus.--—.................--_.--—----.- 343

Astrononion tumidum.---......... 321, 345, tab. 2, pi.

atlantica, Reussella spinulosa. ...... —- — — ——— 354

auberiatia, Quingueloculina...._————————— 338

auriculata, Planispirina... ..__. ————— —— —— 341

Wiesnerella.................. 324, 341, tab. 2, pi. 85

costata, Fissurina................. 324, 350, tab. 3

australis, Miliolinella............- 323, 334, tab. 1, pi. 84

B 

Baculogypsina.---.............................— 319

BagginaphUippinensis,.__._.____- 322, 36^, tab. 4 

philippinensis pilulifera....... 322, 36^, tab. 4, pi. 89

balaniformisproteiformis, Carpenteria --------- — 373

bassensis, Triloculina-..-......... 323, 337, tab. 2, pi. 85

Beach fauna-...__------_____...——... 320,325

beccarii tepida, Rotalia..—........ 320, 360, tab. 4, pi. 89

bengalensis, Anomitina...................—...... 360

Osangularia...... ._______ 322, 360, tab. 4, pi. 89

Ponella..---.......—........——.————. 360

bertheloti, Discorbis—.—..............-........-. 359

berthelotiana, Quinqueloculina..——————————— 333 

bicarinata, Triloculina.._--... 323,335, 339, tab. 2, pi. 85 

bidentata, Quinqueloculina—...... 323, 338, tab. 1, pi. 83

Bifarina.. ________________________ 354 

fimbriata.-— __„.__--—--———-_------ 354

mpckinnonii----.............................. 354

Bifarinella.--.........................__—__ 354

mackinnonii.. ................ 324, 354, tab. 3, pi. 88

ryukyuensis..... ____________——--_ 354

bifrons, Rectobolivina..................... 322, 354, tab. 3

Sagrina.......—————..———————————— 364

Siphogenerina................................. 364

(Sagrina)....-............................... 354

Bigenerina sp_ ___________ 321, S30, tab. 1, pi. 83

Bikini Atoll, localities and depths of samples—— 326-327

bikiniensis, Triloculina.___..___ 323, SS£, tab. 2, pi. 85

Biloculina bulloides.--.---... _...———————— 340

denticulata__________________•___ 340

striolata...........—............—........ 340

lucernula.-- ______________——__ 340 

millettii....................................... 341

murrhina..................................... 341

ringens denticulata............................ 340

striolato-_____________-----__ 340

Bttubulogcnerina, sp............... 321, 354, tab. 3, pi. 88

Bdlimna........................................ 349,354

abbreviata..................... 321, S52, tab. 3, pi. 87

compacta..................... 323, 352, tab. 3, pi. 87

convallaria.................................... 353

ff.o6a.osa-.-----_________..— 324, 350, tab. 3

karreriana.................................... 353

limbata....................................... 353

abbreviata................................. 352

mayori........................................ 354

nitida......................... 321, 350, tab. 3, pi. 87

oceanica.......................___ 324, 852, tab. 3

pseudopygmaea............... 323, 858, tab. 3, pi. 87

Bolivina—Continued

rhomboidalis.................. 323, 353, tab. 3 pi. 87

robusta........................ 321, 353, tab. 3 pi. 87

compacta.............—————.—————. 352

striatula..---................ 323, 353, tab. 3 pi. 87

subexcavata.............__.. 323, 353, tab. 3 pi. 87

subreticulata......................... 324, 353. tab. 3

subtenuis............................ 324, 353, tab. 3

tennis-.......................—............. 353

tortuosa..----................ 323, 353, tab. 3 pi. 87

BMmneUa folia-...—........... 323, 349, tab. 3 pi. 87

folia ornata——.........._- 323, 349, tab. 3 pi. 87

Bolivinitaguadrilatera.—............... 322,349, tab. 3

bolivinoides, Pleurostomella... 322, 324, 356, tab. 3, pi. 88 

bosciana, Quinquflocutina......................... 339

bradyana, Quinqueloculina........ 321,332, tab. lr pi. 83

bradyi, Astacolus.................. 321, 343, tab. 2, pi. 86

Cassidulinoides... ____..__.______ 367 

Cristellaria..——.............................. 343

Cymbalopora_____—————————————— 364 

poeyi.-.-................................. 364

Cymbaloporetta............... 323, 364, tab. 4r pi. 90

Gaudryina—.................—.............. 332

Hauerina—-............ 323, 336, 337, tab. 1, pi. 84

Karrerietta.-——....-.-.. 321, S3., tab. 1, pi. 83

Nubecularia............ _____—____—. 342

Parrina....—................ 323, 348, tab. 2, pi. 85

SUvestria-.—.....................—..—... 342

Trifarina--------...........-. 321, 356, tab. 3, pi. 88-

Bulimina aculeata- ............... 322, 350, tab. 3, pi. 87

affinis........................ 322, 350, tab. 3, pi. 87

costata........................ 322, 350, tab. 3, pi. 87

eleganttssima... —— ———————— ——————— 349 

fijiensis.-_.---_.___.-. 320, 350, tab. 3, pi. 87 

rostrata.-.____________ 322, 350, tab. 3, pi. 87 

subcylindrica________..——______ 350 

wittiamsmiana.________.„______ 349 

Buliminella elegantissima.............. — 324, 349, tab. 3

madagascariensis spicata___._———————— 349 

milktti—........—......... 320, 349, tab. 3, pi. 87

parallela—.................................... 349

Buliminidae—————__-—————.... 349

Buliminoides williamsoniana...... 324, 349, tab. 3, pi. 87

bulloides, Biloculina..................______. 340

Glotigerina———............ 323, S6S, tab. 5, pi. 91

Nonionina.................................... 367

PuUenia—.......................... 322, 367, tab. 4

Sphaeroidina——-............ 322, 367, tab. 4, pi. 90

Tretomphalus................................. 364

O 

calabra, Cassidulina............................... 366

calcar, Robulus...... „——————— 321, 342, tab. 2, pi. 86

Rotalia....................................... 319

Calcarina_____________._______-- 319,325 

defrancii___________________._ 363 

hispida.-............... 320,323,325,363, tab.4, pi. 90

pulchella..— .............—.............-- 364

quoyi.......................................-. 363

spsnglerL... 319,320,321,325,363,364, tab. 4, pis 82,92 

Calcarinidae—...__...._____...-_-__-- 363

Camerinidae.------________________. 346

candeiana, Textularia.............— 323,359, tab. 1, pi.83

Candeina nitida——................ 322,370, tab. S, pi. 91

carpenteri, Cydodypeus—--........ 321,347, tab.3,pi.82

Carpenteria balamformis proteifor mis--............ 373

proteiformis....................--.... 320,373 tab.5

sp—————————.—— 324,373,tab.S pl.82

Carterina spiculotesta.............— 324,34,,, tab. 2, pi. 85

Cassiditlinaatternans...........................-. 366

angulosa........................ 322,365, tab. 4. pi. 90

calabra..............-.......................... 366

caudriae.-..-................................. 366

costatula........................ 322,365, tab. 4. pi. 90

decorata...-.............-..................... 365

379
380 INDEX

Cassidulina alternans—Continued

delicata......................... 322, 365, tab.4,pi. 90

elegans......................... 322,366,tab.4,pi.90

gemma......................... 322, 366, tab.4,pi.90

marshallana.................... 322,366,tab.4,pi.90

minuta..................... 323,325,366,tab.4,pi.90

pacifica............................... 321,366,tab.4

patula................................ 322,356,tab.4

spinifera........................ 322,366,tab.4, pi.90

subglobosa.................. 322,366,367,tab.4,pi.90

vax............................. 367, tab.4,pi.90

Cassidulinidae.———_—_______________ 355 

Cossidulinoides bradyL............................ 367

Sf>.................................... 322,367,tab.4

castrensis, Lagena................................. 351

Lagena orbiffnyana............................ 351

catenulata, Oolina................................. 350

caudriae, Cassidulina.............................. 366

CeratobuUminapadfica......... ____-- 324,365,tab.4

Ceriopora globulus................. .._____... 373

Chilostomella oolina...................... 324,367, tab.4

ChUostomellidae-.-..._.____.________ 367 

Chrysalidinella dimorpha.......................... 355

fijiensis—...................... 324,355,tab.3,pi.88

Cibicidella variabilis................. 324,370, tab. 5, pi. 82

Cibiddes.......................................... 372

cicatricosus...........____-- 322,377, tab. 5, pi. 91

lobatulus....................... 324, S71, tab. 5, pi. 91

mayori......................... 324,37;,tab.5,pi.91

pseudoungerianus.-............. 324,370, tab. 5, pi. 91

Cibiddina mississippiensis........................ 372

sp___——————————— 322,370, tab. 5, pi. 91

cicatricosa, Anomalina................. _____— 371

cicatricosus, Cibiddes.............— 322, 371, tab. 5, pi. 91

drcularis, Fissurina................ 323,357, tab. 3, pi. 87

Spiroloculina................... 321,335, tab. l,pl.84

dorsata—................................. 335

sublineata, MUiolina........................... 337

cZoro, Spiroloculina................. 323,335, tab. 1, pi.84

lirata, Spiroloculina............. 320,335, tab. 1, pi.84

Clavulina angularis................. 324,332, tab. 1, pi. 83

padfica............................... 324,330,pi.83

primaeva----................................. 332

deridi, Robulus................................... 343

Cliona............................................ 347

communis, Spiroloculina............ 323,335, tab. 1,pi.84

compacta, Bolivina.................. 323,352, tab. 3, pi. 87

Bolivina robusta............................... 352

Sphaeroidina......................... 322,367,tab.4

compressa, Hauerina... .....-____...____. 336

concava, Siphotextularia............. 322, SSO, tab. 1, pi. 83

concavum, Plecanium............................. 330

concentrica, Epinoides............................. 360

Mississippina............... 322, SCO, 361, tab. 4, pi. 89

Pulvinulina....... ~........................-- 360

condnna, Discorbina.............................. 358

Discorbis.....................— 323,358, tab. 4, pi. 89

Conditions affecting variations of species, lagoon

fauna________________ 320-321

reef-flat fauna___________________ 320

conglobata, Globigerina................•............

Globigerinoides-—- 320,321,323,326,368, tab. 5, pi. 91

•conglomerata, Globigerina. -......... 322,368, tab. 5, pi. 91

tonica, Textularia................... 323,305, tab. 1, pi.83

corrugata, Textularia........................-- 329

•Conicospirillina-............—.................... 357

semi-involuta................... 324,357, tab. 4, pi. 88

sp————_——...————.... 324,357, tab. 4, pi.

tonvallaria, Bolivina............................... 353

convattarium, Loxostoma.......................... 353

Loxostomum.................-- 323,353,tab.3,pi.88

.convoluta, Pseudobulimina........................ 350

Cores, Bikini lagoon——.—...__.____-- 320-321 

Sylvania Guyot......._..__._______ 322

•Cornuspirainvolvens.............................. 341

planorbis.....................-. 323, S41, tab. 2,pi. 85

corrugata, Spiroloculina...........— 323,335, tab. 1, pi. 84

Textulana..............--.................... 329

conica.................................... 329

^orticata, Rugidia................... 323,360, tab. 4, pi. 89

Sphaeroidina.................................. 362

costata, Bulimina.................. 322, SSO, tab.3, pi. 87

Cristettaria.................................... 343

Fissurina auriculata................ 324, 350, tab. 3

Siphogenerina................................. 356

costatula, Cassidulina............. 322, S65, tab. 4, pi. 90

Siphogenerina raphanus....................... 356

rassa, Globorotalia................ 322,370, tab. 5, pi. 91

Miliolina—................................... 333

Pulvinulina................................... 370

subcuneata, Quinqueloculina... 323,333, tab. 1, pi. 83 

cribrorepandus, Poroeponides...... 323, SCO, tab. 4, pi. 89

crispata, Siphotextularia........... 321,330, tab. 1, pi. 83

Textularia.................................... 330

Cristettaria brad^L................................ 343

costata........................................ 343

gibba.......................................... 342

nigrisepta..................................... 343

schloenbachi................................... 344

crustata, Discorbis................. 323,358, tab. 4, pi. 89

cubensis, Gnmbelina.............................. 349

curva, Heterostegina............................. 346, 347

cushmani, Spiroloculina........................... 336

Cydodypeus...................................... 34'

carpenteri..................... 321, 347, tab. 3, pi. 82

guembelianus................................. 347

Cymbalopora bradyi............................... 364

poeyi......................................... 364

bradyi.................................... 364

squammosa................................... 364

Cymbaloporetta bradyi........... 323, 364, tab. 4, pi. 90

squammosa................... 323, 364, tab. 4, pi. 90

Cymbaloporidae_--_—..__.__.__.....____—...-. 364

D

davidiana, Vahulina.............. 323, SSI, tab. 1, pi. 82

decorata, Cassidulina-............................. 365

Spirfflina........... 320, 324, 356, 357, tab. 4, pi. 88

var.- —— —————_——-—— 357, tab. 4, pi.88 

Deep-water fauna.._..................... 321-322,326

defrancii, Calcarina............................... 363

dehiscens, Sphaeroidina............................ 369

Sphaeroidinetta........... 322, 326, 369, tab. 5, pi. 91

delicata, Cassidulina.............. 322,365, tab. 4, pi. 90

densepunctata, Spirillina vivipara-- 324,357, tab. 4, pi. 88 

Dentalinafiliformis-..-.......... 321, 343, tab. 2, pi. 86

denticulata, Biloculina............................ 340

Biloculina ringens............................. 340

Pyrgo......................... 323, 340, tab. 2, pi. 85

striolata, Biloculina........................... 340

Pyrgo—.................. 323, S40, tab. 2, pi. 85

depressa, Heterostegina............................ 347

digitata, Hastigerina............................... 369

HastigerineUa................. 322, S69, tab. 5, pi. 91

, Chrysalidinella......................... 355

Sagrina....................................... 355

padfica, Siphogenerina........ 321, 355, tab. 3, pi. 88

Dimorphinatuberosa.............. 321, 844, tab. 2, pi. 86

Discorbina condnna—............................ 358

patettiformis.................................. 359

poeyi.......................................... 364

polystomdloides............................... 360

rugosa........................................ 359

tuberocapitata................................. 359

Discorbis aUomorphinoides........................ 361

bertheloti-.................................... 359

condnna...................... 323, SSS, tab. 4, pi.

crustata....................... 323, 358, tab. 4, pi.

frustata....................... 324, 358, tab. 4, pi.

micens—..................... 323,358, tab. 4, pi.

opi'ma........................ 323,358, tab. 4, pi.

patelliformis.-................ 323,355, tab. 4, pi.

rugosa........................ 323, 359, tab. 4, pi.

subbertheloti................... 320,350, tab. 4, pi.

tuberocapitata................. 321,355, tab. 4, pi.

dispar, Elphidium advenum— 323, 345,346, tab. 2, pi. 86 

distorqueata, Ouinqueloculina...... 323, 333, tab. 1, pi. 83

diversa, Hauerina............ 323, 336, 337, tab. 1, pi.

dorsata drcularis, Spiroloculina.................... 335

dubia, Pegidia........................... 324, 367, tab. 4

Rotalia....................................... 361

dupla, Textularia................. 323,325, tab. 1, pi. 83

durrandii, Miliolina.......................____ 341

E

earlandi, TrUoculina.............. 323,338, tab. 2, pi. 85

Virgulina..................... 324,350, tab. 3, pi. 87

echinata, Planorbulma...........'................. 361

Siphonina.................................... 361

Siphoninoides................. 323,361, tab. 4, pi. 8»

Truncatulina................................. 361

dwardsi, Planispirina (Sigmoilind)................ 336

Sigmottina.................... 324,336, tab. 1, pi. 84

Ehrenbergina padfica............. 322,367, tab. 4, pi. 00

reticulata..................... 322, 367, tab. 4, pi. 90

serrata........................................ 367

elegans, Cassidulina... ____.._ 322, 366, tab. 4, pi. 90 

Epistomina............................t___ 360

Hoglundina......................... 322, SCO, pi. 89

•legantissima, Bulimina........................... 349

Buliminetta......................... 324, S49, tab. 3

EUipsoidinidae—__—._.__....__.___.. 356

elongata, Articulina .................... 324, 336, tab. 1

Globigerina-.................................. 368

Globigerinoides-...-.----..---. 324,368, tab. 5, pi. 91

Pyrgo........................................ 341

Elphidium advenum.......... 323,345,346, tab. 2, pi. 86

advenum dispar........... 323,345,346, tab. 2, pi. 86

fimbriatulum...... ______.______._. 346

jenseni-...................... 323,346, tab. 2, pi. 86

milletti-—-......................... 324,346, tab. 2

simplex....................... 323. 346, tab. 2, pi. 86

Eniwetok Atoll, localities and depth-* of samples. 327-328

Entosoleniaformosa............................... 351

squamosa..................................... 350

Epinoides concentrica............................. 360

Epistomaroidespolystomelloides... 320,360, tab. 4, pi. 89

Epistomina elegans............................... 360

Epistominella obtusa.............. 324,365, tab. 4, pi. 90

pulchra—........................... 324,365, tab. 4

tubulifera.- —............... 323,366, tab. 4, pi. 90

sp-_—----------__----- 323,365, tab. 4, pi. 90

Eponides....—.................................. 360

repanda...................................... 360

tenera..-..................... 322,359, tab. 4, pi. 89

terebra....................... ..t ______.-- 361

excavata, Spiroloculina............................ 335

exigua, Hauerina................................. 341

Planispirina...................... 341, tab. 2, pi. 85

Pulvinulina.................................. 365

obtusa, Pulvinulina...........____.._.... 365

expansusplanulatus, Robulus..................... 343

F 

ferussadi, Ouingueloculina........ 323,333, tab. 1, pi. 83

fijiana, Patellinella..................~ 358, tab. 4, pi. 89

fijiensis, Bulimina..............— 320,350, tab. 3, pi. 87

Chrysalidinella................ 324,355, tab. 3, pi. 88

Patettinetta................................... 324

filiformis, Dentalina............... 321,343, tab. 2, pi. 86

Nodosaria.................................... 343

fimbriata, Bifarina................................ 354

Globorotalia menardii.......... 322,370, tab. 5, pi. 91

Pulvinulina menardii......................... 370

Uvigerina porrecta..______ 322,355, tab. 3, pi. 88 

fimbriatulum, Elphidium-.-.-.-.................. 346

Fissurina auriculata costata.............. 324,350, tab. 3

drcularis...................... 323,S51, tab. 3, pi. 87

formosa....................... 323, SSI, tab. 3, pi. 87

lacunata...................... 323,351, tab. 3, pi. 87

marginata.................................... 351

marginato-perforata........................... 351

mittetti—..................... 323,357, tab. 3, pi. 87

orbignyana...........................___.. 351

perforata...................................... 351

radiato-margmata............. 324,357, tab. 3, pi. 87

sp. A.____......._..-__ 324,357, tab. 3, pi. 87

sp. B—————— —....... — 321, 350, tab. 3, pi. 87

sp. C_———... ....——.... 322,350, tab. 3, pi. 87

spp____________________________ 352 

fistulosa, Globigerina.............................. 366

Globigerinoides sacculifera...... 324,365, tab. 5, pi. 91
INDEX 381

flabettiformis, Pavonina....... ...-._--______. 355

Flintia........................................... 338

flintiana, Miliolinetta oceanica.. .....__. 324,334, tab. 1

flintii, Gaudryina..... ._-_.__.__ 322,330, tab. 1

folia, Bolimnella.—............... 323, 349, tab. 3, pi. 87

TextiOaria.................................... 349

ornata, Bolivinella............. 323, 349, tab. 3, pi. 87

foliacea, Textularia.-........ 323,329,330, tab. 1, pi. 83

oceanica, Textularia........... 323,3#fl, tab. 1, pi. 83

formosa, Entosolenia.............................. 351

Fissurina_______._..... 323,35?, tab. 3, pi. 87

Lagena-...................................... 351

foveolata, Spiroloculina. __--. 320,335, tab. 1, pi. 84 

frondescens, Sagenina.............__. 323,328, tab. 1

Frondicularia sp. A _. _ _ _ __. __ _ 321, 344, tab. 2, pi. 86

frustata, Discorbis _______.___ 324,358, tab. 4, pi. 89

funalis, Articvtina................................ 336

Tubinella..................... 324,336, tab. 1, pi. 84

G 

gaimardt, Operculina..—---. ......__.._.__. 346

Gaudryinabradyi.---_.....................:...... 332

flintii.............---...........— 322,330, tab. 1

pauperata—-................ 324,330, tab. 1, pi. 83

pupoides—-_..-_--._---_---_-_._-.-_--...... 332

quadrangularis...—.. — -.—.----. — ........... 331

rugosa........................................ 330

rugulosa-..........—... ........._______ 331

siphonifera............... .........______ 331

subrotundata.................................. 330

triangularis angulata........ __ 321,33?, tab. 1, pi. 83

truttissata—.................. 321,831, tab. 1, pi. 83

(Siphogaudryina) rhodiensis......_____-. 331

rugulosa.................. 323,33?, tab. 1, pi. 82

siphfnifera.................—... 321,331, pi. 83

_rans»ersorio___-...______ 321,331, pi, 83

gemma, Cassidulina............... 322,366, tab. 4, pi. 90

gibba, CristeUaria.-.. ——......................... 342

gibbus, Robulus................... 321,342, tab. 2, pi. 86

glabrata, Anomalina............— 324,370, tab. 5, pi. 91

Globigerina..................................... 368,369

aequilateralis................................. 369

bulloides...................... 323,368, tab. 5, pi. 91

conglobata..... .........__..__._____. 368

conglomerata......... _...-..__ 322, 368, tab. 5, pi. 91

elongata—..................... __-______._-. 368

fistulosa. .........._.....____.__.___-- 369

rubra..................................____ 369

sacculifera...............__....._......... 369

subcretacea.................... 322, 368, tab. 5, pi. 91

sp—— .—_-.__---- — .__- 320, 368, tab. 5, pi. 91

Globigerinella.... — .............________.. 369

aequilateralis.....--..-. 322, 326, 869, tab. 5, pi. 91

Globigerinidae. ____-...--__._________-- 368

Globigerinoides............................- 326,368,369

conglobata...-... 320, 321, 323, 326, 368, tab. 5, pi. 91 

elongata,............. ____-- 324, 368, tab. 5, pi. 91

rubra... — — ......... 321, 323, 326, 369, tab. 5, pi. 91

sacculifera-.......... 321, 323, 326, 369, tab. 5, pi. 91

fistulosa................ 324,369, tab. 5, pi. 91

Globorotalia crassa................ 322, 370, tab. 5, pi. 91

hirsuta—--------.....—.—... 322, S70, tab. 5, pi. 91

menardii......_____-. 322, 326, 370, tab. 5, pi. 91

fimbriata—.............. 322, 370, tab. 5, pi. 91

tmncatulinoides.. — -......... 322, 370, tab. 5, pi. 91

tumida................... 322, 326, 370, tab. 5, pi. 91

Globorotaliidae................................... 370

globosa, Lagena............. ...... 324, 344, tab. 2, pi. 86

ghbula, Gypsina.................. 324, 373, tab. 5, pi. 91

globulosa, Bolivina...—.-..............— 324, 352, tab. 3

globulus, Ceriopora.... .--.-.-...._____._._ 373

gracilis, Lagena....... _._____-- 321, 344, tab. 2, pi. 86

granulosa, Operculma............................. 346

grata, Spiroloculina......................_..._.__. 334

angulata, Spiroloculina........................ 334

grateloupi, Spiroloculina.......................... 335

acescata, Spiroloculina..............._...... 334

guembeUanus, Cycloclyptus...................... 347

Gumbelm a........................................ 349

cubensis. ........_.----....-.--............. 349

marshallana....—............ 322,349, tab. 3, pi. 87

Guyot, species.-_......_.__......_._____ 322

Gypsina...............——..—..............—.— 373

globula—..................... 324, 373, tab. 5, pi. 91

inhaerens- ................................... 372

plana.—.................................. 373

yiana................................ 320, 373, pi. 93

vesicularis-................... 324, 373, tab. 5, pi. 82

Gyroidina soldanii.............. ... 322, 359, tab. 4, pi. 89

H 

Halimeda........................... 328,373,374, pi. 93

Haplophragmium...............................-. 368

Haplophragmoides....._.-------._------...-.--.-- 368

Hastigerina digitata............................... 369

Hastigerinella digitata............. 322, 369, tab. 5, pi. 91

hauerii, Pulvinulina.....--_-----....--------.--.-- 361

Hauerina ..........._.......__.___...... 337

bradyi———..-..—.--. 323, 336, 337, tab. 1, pi. 84 

compressa.................................... 336

diversa——... —— —— —— . 323, 336, 337, tab. 1, pi. 84 

exigua............................... ......... 341

involuta.—-........................ 323, 337, pi. 84

miUetti—.-.-- — ......----.. 323, 337, tab. 1, pi. 84

ornatissima................................... 337

pacifica-... .................____...__— 337

serrata............ ............ 320, 337, tab. 1, pi. 84

tpeciosa....................................... 337

hemprichii, Amphisorus........................... 348

Heronallenia...................................... 359

sp—— — —— —— — — -- 324, 359, tab. 4, pi. 89

Heterohelicidae ._____.....____..__.____ 349

Heterostegina............... .__.___..... — - 319,347

curva....................................... 346,347

depressa.........—...................... — .— 347

suborbicularis 320, 323, 325, 346, 347, tab. 3, pis. 82, 87 

hirsuta, Globorotalia..—. .......... 322, 370, tab. 5, pi. 91

Rotalina—................................... 370

ft.s-p.da, Calcarina........ 320, 323, 325, 363, tab. 4, pi. 90

pulchella, Calcarina........................... 364

Hoglundina elegans.........—.... 322, 360, tab. 4, pi. 89

Homotrema...............__________..— 325,374

rubrum.. 319, 320, 323, 373, 374, tab. 5, pis. 82, 92, 93 

Homotremidae — _..........._._._-_..__.. 373

Hyperamminidae_._.______-------____ 328

impressa, Spiroloculina.................... __.,_-- 335

inaequalis, Spirillina.............. 324, 357, tab. 4, pi. 88

incertus, Ammodiscus.. — -.............. 324, SS8, tab. 1

inconspicua, Patellinella.......... 320, 358, tab. 4, pi. 89

Textularia........... .____________.. 358

inflata, Nubecularia............................... 342

inhaerens, Acervulina............. 324, 378, tab. 5, pi. 91

Gypsina...................................... 372

plana, Acervulina.._______.'.._____-. 373

Gypsina.._________._________ 373

involwta, Hauerina______. __ 323, 337, tab. 1, pi. 84

Triloculina--.......——..........— 323, 338, pi. 85

involvens, Cornusptra............................. 341

^rregulans, Qumgueloculina....................... 338

Triloculina................... 323, 338, tab. 2, pi. 85

japonica meyicana, Nonionella..................... 345

jenseni, Elphidium................ 323, 346, tab. 2, pi. 86

PolystomeHa.................................. 346

jugosa, Patettinetta.——............ 324,358, tab. 4, pi. 89

Textularia— _____—__________.__ 358

K 

karreriana, Bolivina............................... 353

karrerianum, Loxostoma.......................... 353

Loxostomum.——............. 322,353, tab. 3, pi. 88

Kantrietta bradyi.................. 321,330, tab. 1, pi. 83

kerimbaensis, Textulaiia........... 321,3X9, tab. 1, pi. 83

ktrimbatica, Miliolina.............. _______.. 339

Quinqueloculina................_______.. 339

Triloculina.................... 323,339, tab. 2, pi. 85

L 

labiosa, Miliolina.................................. 334

Miliolinetta................................... 342

labiosa, Miliolina— Continued

TtttociaineUa...—..———. 323,334, tab. 1, pi. 82 

lacunata, Fissurina................ 323,351, tab. 3, pi. 87

Lagena............................ —..—..-.. 351

orbignyana................................ 351

lacunenais, Nubecidaria............ 320, 341, tab. 2, pi. 85

Lagena-------...—............................... 344

acutkosta..——......——.... 321,344, tab. 2, pi. 86

castrensis..............................—.— 351

formosa.............................—........ 351

globosa..............————.. 324, 344, tab. 2, pi. 88

gracilis........................ 321,344, tab. 2, pi. 86

lacunata............ __———. ———— —.. — -. 351

marginata.................................... 351

margtnato-perforata... —————.————— ----- 351

orbignyana.................................... 352

castrensis................................. 351

lacunata.................................. 351

radiato-marginata....—---............... — ... 351

spiralis...........—...—...... 324, 344, tab. 2, pi. 86

striato-punetata... ................... 324,344, tab. 2

wiUiamsoni.................... 324, 344, tab. 2, pi. 86

Lagenidae....--.-----------------.-----—----- 342

Lagoonfauna—— —— ——————— 320-311,325

Lamarckina scabra................. 324,359, tab. 4, pi. 89

larvata, Planorbulina.-... ————..———————— 372 

PlanorbidineUa......—— — —— .—— 324,572,tab. 5

lateralis, Poroeponides...........—....-.--.-- — --. 360

Laticarinina pauperata--—-.............- 324,37?, tab. 5

lepidula, Nodosaria.. ................——.—..... 356

Siphonodosaria................ 322,356, tab. 3, pi. 88

lessonii, Amphistegina... ........._—————— 362

limbata, Bolivina—.—............................. 353

abbreviata, Bolivina——....................... 352

limbato-striata, Nonionella.......--....-........... 346

limbatum, Loxostomum........ 323,353,354, tab. 3, pi. 88

limbosa, Robulina.....................—......... 343

limbosus, Robulus..............-- 321,343, tab. 2, pi. 86

linneiana, Triloculina-............................ 338

Urata, Spiroloculina clara.......... 320,335, tab. 1, pi. 84

Listeretta primaeva................................ 332

Lithothamnion........__-- ——... ————— —— 326

(IMuus) acicularis, Nautilus...............——— 348

arze.tB.ts, Nautilus.....--..................... 348

lobatula, Truncatulina............................ 371

lobatulus, Cibkides—.............. 324, 371, tab. 5, pi. 91

Nautilus...................................... 371

Loxostoma convaUarium........................... 353

karrerianum.................................. 353

Loxostomum convallarium......... 323,353, tab. 3, pi. 88

karrerianum.-. ____....—_ 322,353,tab. 3, pi.88

limbatum.................. 323,353,354, tab. 3, pi. 88

mayori...........—......-.... 324, 354, tab. 3, pi. 88

lucernula, Biloculina.......................—..... 340

Pyrgo—.-.—————__ 321,340, tab. 2, pi. 85

M 

macella, Polystomella--........................... 346

mackinnonii, Bifarina...........——.......—.—. 354

Bifarinetta..................... 324,354, tab. 3, pi. 88

madagascariensis, Amphistegina...——.. — ——— 319,

320,321,323,325,326,362, tab. 4, pi. 90

spicata, BulimineUa........................... 349

marginalis, Orbulites.............................. 348

Sorites......................... 323,_?4S-tab. 3, pi. 82

marginata. Fissurina...........................-.. 351

marginoto-perforata, Fissurina-................. ... 351

Lagena.......... ———————— —— —. ——. 351

Marginopora.................................... 319,348

vertebralis——— 319,320,323,325,348,364, tab. 3, pi. 82

marshallana, Cassidulina.......... 322,366, tab. 4, pi. 90

GHmbelina.................... 322,349, tab. 3, pi. 87

Spiroloculina.................. 323,335, tab. 1, pi. 84

Triloculina.................... 323,339, tab. 2, pi. 85

Massilina planata................... 324,334, tab. 1, pi. 84

mayori, Bolivina.................................. 354

Cibicides—................... 324,377, tab. 5, pi. 91

Loxostomum..-.. — ... — —.... 324,354,tab. 3, pl.88

Potaliammina______.__ 324,342, tab. 2, pi. 85 

Spiroloculina.................. 324,336, tab. 1. pi. 84

Truncatulina.............. ————————— — 371
382 INDEX

melo, Nautilus— ———————.. —————— - — ---_ 345 

menardii, QloborotaUa____—_ 322,326, 370, tab. 5, pi. 91

PuMnulina....___>___________———-- 370

Ratalia---.................................... 370

PuMnulina...............————————— 370

fimtriata, Oloborotalia.......... 322,370, tab. 5, pi. 91

pauperata, PuMmdina itpanda..........-.-__ 371

mexicana, Noniondlajaponka............ ___--.-__ 345

micacea, Proieonina......................_..-_---_ 328

micens, Discorbis............. 323, 3S8, tab. 4, pi. 89

Miliolidae... ————————— ———————„. 332

Miliolatrigonula.-....— _.-.__-_____---_-.__ 340

Miliolina alveoliniformis .......................... 337

anguina agglutinans...-_..-------._------_---. 332

circularis sublineata........................... 337

crassa......... ___— —._-_____— ——. 333

<-Wr<m<-?.----------------------------— ------ 341

kerimbatica................................... 339

labiosa ------...-----__.._____—— —— __ 334

otlonga...... _..——..—....———— _... 339

parkeri----.-.......-__.___.-. — .. — _ 333,339

terquemiana .............-......-..-...--.- 338,340

tricarinata .................................... 340

trigonula ... _———..————-—-————... 340 

undosa.---..... ._----------_-----__... 332

MJ7iaHn««a_._————— — — ———————_ 334

ausfralis....................... 323, 334, tab. 1, pi. 84

labiosa-... _.___--.-.-----.---.-----_._-_-_._ 342

oceanica-...... -..__-——_......... 324, SS4, tab. 1

flintiana.-....... — ._..„.._..__.... 324, S34

MiUepora miniacea.-.-........................._ 374

re&ro-----------------------_-----—---.-.. 373

miHetti, Bulimhiella..------ 320, S49, tab. 3, pi. 87

Elphidium.----..................... 324, 848, tab. 2

Fissurina...................... 323,361, tab. 3, pi.87

Hauerina...................... 323, 337, tab. 1, pi.84

Polystomella-............ __...__...___ 346

Spiroplectammina.........._.....--.-...... 329

Textularia.--. __-___....___.... 321,300,330

millettii, Biloculina................................ 341

Pyrgo.......................... 324,34.7, tab. 2, pi. 85

Mimosinapacifica .....---.-......... 323, 354, tab. 3

miniacea, MWepora..............________ 374

Miniancina............ 320, 324,373, 374, tab. 5, pi. 82

Polytrema.................................... 374

Miniacina.-..........--_...__--..__.._.. 325

alba.—...-. - — -- — --.-..-...--............. 374

miniacea...-......--- 320, 324, 373, 374, tab. 5, pi. 82

minuta, Cassidulina.. ______ 323, 325, S66, tab. 4, pi. 90

mississippiensis, Cibidda.......................... 372

Mississippina concentrica.... 322, S60, 361, tab. 4, pi. 89

Mmalysidium polUum........ 320, 348, tab. 3, pi. 87

(Monalysidium) polita, Peneroplis... ________ 348 

mucronata, Articulina............................ 336

murrhina, Biloculina........................._.. 341

Pyrgo............................ 321, 341, tab. 2

N 

Nautilus ammonoides -----------.--—....---.... 346

lobatulus...................................... 371

melo.............______._________ 345

pompilioides...-------.-.....--------... ...... 345

raaiatus.---------------- ...-__-_.--.... 362

spengleri...................................... 363

venoms.......----------------------___... 363

(Lituus) adcularts...------....- ______.. 348

arietinus.-.--.......-.....-............... 348

Neoalveolinapulchra............ 324, 348, tab. 3, pi. 87

neostriatula, Quinqueloculina....... 323, 333, tab. 1, pi. 83

nigrisepta, Cristellaria..................____... 343

Robulus...................... 321, 34S, tab. 2, pi. 86

nitida, Bolivina........._____ 321, 35S, tab. 3, pi. 87

Candeina---................. 322, 370, tab. 5, pi. 91

Nodosaria abyssorum.—........................... 356

filiformis.....—............................... 343

lepidula __._____..._______-__-__ 356 

pauciloculota................ 321, 848, tab. 2, pi. 86

prava......................................... 344

sp. A-------..-.-.--.---.—— 322, S44, tab. 2, pi. 86

sp. B——............—.-.-. 321, 344, tab. 2, pi. 86

sp. C—...-...--....-...__ 321, 344, tab. 2, pi. 86

Nondiagnosticspecies,..- ——.... ———.----__ 323-324

Nonion pacificum................ 321, 348, tab. 2, pi. 86

pompilioides................ 322, 345, tab. 2, pi. 86

umbilicatulum pacificum........ ..--.._-...... 345

Nonionella....._ ______________.--.—. 345

japonica mexicana..... ..—————- — _ ———. 345

limbato-striata___.. ———— ------- — ——.. ——. 345

translucent................. . 323, 34$, tab. 2, pi. 86

sp. A_...........__............ 324, S45, tab. 2

sp. B—--——.-..——— 323, 345, tab. 2, pi. 86 

Nonionidae-.- ———————-——— — -- 325,345

Nonionina bulloides... ————— —....-------_.— 367

pompilioides-......- _...____-___--.—— 345

sMHffero.------------------------------------ 345

umbilicatula pacifica..........--..-----.-..... 345

Nouriapolymoiphinoides------.------- 324, 34%, tab. 2

Nubecularia bradyi..------- — —.-.--___._ —.— 342

inflata........... „.———— ——-_———— 342

lacunensis.................. 320, 341, tab. 2, pi. 85

O

obliquiloculata, PuUenia...—————.————-——— 369 

Pulleniatina—-........ 3$2, 326, 369, tab. 5, pi. 91

oblonga, Miliolina................................. 339

Triloculina................ 323, 333, 339, tab. 2, pi. 85

scabra, Pulvinulina........................... 359

oblongum, Vermiculum.......—....——————.- 339

obtusa, Epistominella............ 324, 365, tab. 4, pi. 90

Pulvinulina exigua............ ._._-.__..._ 365

oceanica, Bohvina........................ 324, 352, tab. 3

MiUollnella...-.-—--.............. 324, 334, tab. 1

Teitulariafoliacea.................... 323, 329, pi. 83

Triloculina................................... 334

flintiana, Miliolhiella................ 324, 334, tab. 1

oolina, CMlostomella..................... 324, 367, tab. 4

Oolin a catenulata. ___... —— --_.-----_-__-——.. 350

squamosa. ... ._________ 324, 350, tab. 3, pi. 87

Operculina ammonoides........... 323, 346, tab. 3, pi. 87

gaimardi..........— .......................... 346

granulosa...............----------—--------- 346

Ophthalmidiidae.-.---. —— -.——- —— --_----- 341

opima, Discorbis.................. 323, S58, tab. 4, pi. 89

orbicularis, Robvlus............................... 343

orbignyana, Fissurina............................. 351

Lagena...--............--.................... 352

castrensis, Lagena........._-..--_-.-.-----.- 351

lacunata, Lagena.............................. 351

Orbitulina vesicularis...-..-....................... 373

Orbulina universa....---...-...-.... 322, 326, 369, tab. 5

Orbulties marginalis........................------- 348

ornata, Angulogerina albatrossi..-- 321, 356, tab. 3, pi. 88 

Boil vinett a folia............... 323, 349, tab. 3, pi. 87

ornatissima, Hauerina-.--_____..-__..._ 337 

Osangularia bengalensis.--- ____ 322, 360, tab. 4, pi. 89 

Outer-slope fauna___-..-.—---------- —— ——.- 321

pacifica, Articulina............... 323, S36, tab. 1, pi. 84

Cassidulina--.-.--.....-..---....-- 321, 366, tab. 4

Ceratobulimina....................-- 324, 356, tab. 4

Clavulina. _______________-- 324,33;.', pi. 83 

Ehrenbergina................ . 322,357, tab. 4, pi. 90

Hauerina........................... ._____ 337

Mimosina- ......__.__.__.... 323,354, tab. 3

Polymorphinella____.___ 321, 344, tab. 2, pi. 86

Siphogenerina dimorpha... __ 321, 355, tab. 3, pi. 88

pacificum, Nonion-.... _____-- 321, 345, tab. 2, pi. 86

Nonion umbilicatulum__________._-.. 345 

papillata, Sphaeridia................ ...-- 324, S6S, tab. 4

papillosa, Amphistegina radiata... 322, 36S, tab. 4, pi. 90 

Parafissurina ventricosa........................... 356

sp——............———- 322, 324, 350, tab. 3, pi. 88

parallela, Ruliminella... ___.________..-- 349 

parkeri, Miliolina.-............................. 333,339

Ouinquelocuhna.......... 323, 324, 333, tab. 1, pi. 83

var.--______________ 333, tab. 1, pi. 83 

Parrella bengalensis__________-__.._-- 360 

Parrina bradyi.................... 323, S4S, tab. 2, pi. 85

patelliformis, Discorbina............--............ 359

Discorbis...-........ _____ 323,359, tab. 4, pi. 89

PattUina advena._._.__._.__.___._ 324,357, tab. 4 

advena altiformis....... .... — . 3:74, 357, tab. 4, pi. 89

Pat IHnettafijiana—..---..---...-- 368, tab. 4, pi. 89 

fijimsis ...___----..__-._______.-- 324

inconspicua............... .--- 3£0, 358, tab. 4., pi. 89

jugosa. -------________ 324, 358, tab. 4, pi. 89

patula, Cassidulina-.......... .„.__-- 322, 366, tab. 4

pauciloculata, Nodosaria... --.-.. 321, 343, tab. 2, pi. 86 

Paumotuaterebra......... ____ 324, 361, tab. 4, pi. 89

pauperata, Gaudryina._______ 324,330, tab. 1, pi. 83 

Laticarinina..-...................... 324, 371, tab. 5

Pulvinulina............ —.-..__————-- 371

repanda menardii---..-.................-- 371

Pavoninaflabelliformis--............. ____.—— 355

Mformis——.............. 322, 324, 555, tab. 3, pi. 88

Pegidiadubia...........-----........--. 324,36.7, tab. 4

Pegidiidae..__...„ — ..... — .--...—......— 361

Peneroplidae._____..-_._____-_____-- 348 

Peneroplis arietinus .— ..................-........ 348

(Monalysidvum) polita.—.. ......._______. 348

sp. ————— — ————— 324, 348, tab. 3, pi. 87

peregrina, Uvigerina........ .__-_ 322, 355, tab. 3, pi. 88

perforata, Fissurina.-................ _—.——-- 351

philippinensis, Baggina.... .......____ 322, 361, tab. 4

Pulvinulina.-..-... ______-...-___..-- 361 

piluhfera, Baggina....- — -.-. 39.2, 361, tab. 4, pi. 89

piluUfera, Baggina philippinensis. 322, 361, tab. 4, pi. 89 

plana, Acervulinainhaerens-.____---------- 373

Gypsina...........................— 320,373, pi. 93

inhaerens-... —————....——.————-- 373

plmata, Massilma.-.-............ 324, 334, tab. 1, pi. 84

planciana, Triloculina........................-- 338,339

Planispirina auriculata...-..--.... __—.—.-- 341

exigua-.........._._______-- 34.7, tab. 2, pi. 85

(Sig moilina') ei'wardsi......................... 336

pianissimo, Spiroloculina................... —... 334

planorbis, Cornuspira____.__ 373, 341, tab. 2, pi. 85

PlanorbuJina acervalis.--- 324, 373, 373, tab. 5, pis. 82, 91

echinata.....---------------------------------- 361

larvata-.-..--.-. — ........................— 372

Planorbulinella larvata-. ................ 324, 378, tab. 5

Planorbulinidae..._____--...______._._ 372 

Planulariaplanulata.--.-.-.----.-----.----------- 343

planulata, Astacolus.............. 3.1, 34S, tab. 2, pi. 86

Planularia__--_.-..--..---..-.---- —._..-. 343

planulatus, Astacolus........-....----.--....-..-. 343

Robulus expansus-.-.-.-----.--...------------ 343

Planulina ariminensis....... ____.... 322, 371, tab. 5

planum, Polytrema.... ..-.--_--...._----------- 373

planus, Tretomphalus.... ......... 3?3, 364, tab. 4, pi. 90

Plecaniumconcavum.--------------- — .---------- 330

Pleurostomella bolMnoides.... 322, 324, 356, tab. 3, pi. 88

sp_._.__--.....-..--------.„-.-----_— 356

poeyi, Cymbalopora.. _---..---.-._—....——— 364

Discorbina.__________-_____----- 364

Rosalina............................____... 364

bradyi, Cymbalopora..... ...___.__...— 364

polita, Peneroplis (Monalysidiu m)_................ 348

politum, Monalysidhim.---.-----. 3"«), 348, tab. 3, pi. 87

polymorpha, Anomalina.......... 3.2, 370, tab. 5, pi. 91

Polymorphinella pacifica.....__ 3,1, 344, tab. 2, pi. 86

Polymorphinidae-_—- —— —.-..- — -... —.- 344 

polymorphinoides, Nouria. _____..___ 324, 34^, tab. 2 

Polystometta advena............................... 345

jenseni—................-------------------... 346

macella....-.-......-------------------------- 346

milletti-............ —————————— 346

verriculata.-----..--------------.----------.— 346

polystomelloides, Discorbina................--.-.-- 360

Epistomairoides.............. 320, 360, tab. 4, pi. 89

Polytrema miniacea__._____________ 374 

planum..........______.________ 373

rubra______.________________ 373 

pompilioides, Nautilus.--....-.................... 345

Nonion....................... 3:72, 345, tab. 2, pi. 86

Nonionina....... ._.__-----------_...... 345

Poroeponides..................................... 360

cribrorepandus.... _____... 3:73, 360, tab. 4, pi. 89

lateralis....................................... 360

porrecta, Uvigerina................ 333, 355, tab. 3, pi. 88

fimbriata, Uvigerina........... 3:72, 355, tab. 3, pi. 88
INDEX 383

•prava, Nodosaria.... _______.---.-.---___ 344

primaeva, Clavuliwa................._-.-..__. 332

Listerella..................................... 332

Schenckiella......................... 322, 332, tab. 1

probosddea, Uvigerina.........-.-- 321, S55, tab. 3, pi. 88

vadescens, Uvigerina....... .— 321, 355, tab. 3, pi. 88

proteiformis, Carpenteria. -.............. 320,373, tab. 5

Carpenteria balaniformis................. — ... 373

Proteonina micacea..........................--_.. 328

sp—— — ————— — . 324, 328, tab. 1, pi. 83

Pseudobuliminaconvoluta......................... 350

sp. ——-.-_———————-- 324,350, tab. 3, pi. 87

pseudopygmaea, Bolivina.......... 323, 35%, tab. 3, pi. 87

pseudoungeriana, Truneatulina.................... 372

pseudoungerianus, Cibiddes....... 324, 872, tab. 5, pi. 91

pulchella, Calcarina hispida....................... 364

pulchra, Alveolina................................ 348

Epistominella....................... 324,365, tab. 4

NtoaJveolina.................. 324, 348, tab. 3, pi. 87

Pulvinulina.-------.......................... 365

PuUaiia bulloides....................... 322, 367, tab. 4

obliquiloculata.... _.--------------------------_

salisburyi..................... 322, 367, tab. 4, pi. 90

Putteniatina obliquiloculata... 322, 326, 369, tab. 5, pi. 91 

Pulvinulina concentrica........................... 360

crassa........................................ 370

exigua........................................ 365

obtusa-.---............................... 365

hauerii........................................ 361

menardii..................................... 370

fimbriata................................. 370

oblongascabra................................. 359

pauperata.................................... 371

philippinmsis—............—.......... ._- 361

repanda menardii pauperata................... 371

tumida....................................... 370

Pulvinulinella pulchra............................ 365

pupoides, Gaudryina.............................. 332

Pyrgo denticvlata................. 323,340, tab. 2, pi. 85

dentkulata striolata........... 323, 340, tab. 2, pi. 85

dongata...................................... 341

lucernula..................... 321, 340, tab. 2, pi. 85

mittettii....................... 324, 34^, tab. 2, pi. 85

murrhina........................... 321, 341, tab. 2

sp............................ 323, S41, tab. 2, pi. 85

Pyroptlus rotundatus--.......-... 324, 864, tab. 4, pi. 90

Q

quadrangularis, Oaudryina........................ 331

quadrilatera, Bolivinita............--.... 322, 349, tab. 3

Textularia.................................... 349

Quinquelocitlina................................ 333,334

agglutinans............................._... 332

anguina agglvtinans........................... 332

arenata................... 320, 322, tab. 1, pi. 83

auberiana................... ----------------- 338

berthelotiana.................................. 333

bidentata-.................... 323, 332, tab. 1, pi. 83

bosdana...................................... 339

bradyana..................... 321,332, tab. 1, pi. 83

crassasubcuneata............. 323,333, tab. 1, pi. 83

dlstorqueata................... 323, 333, tab. 1, pi. 83

ferussacii..................... 323,333, tab. 1, pi. 83

irregularis.................................... 338

kerimbatica................................... 339

neostriatula.............. 323, 333, 340, tab. 1, pi. 83

parkeri................... 323, 324, 333, tab. 1, pi. 83

vsa:........................... 333, tab. 1, pi. 83

samoaensis............—.-... 324, 333, tab. 1, pi. 83

seminulum............. 323, 333, 339, tab. 1, pi. 83

striatula.... ...................-------------- 333

suJcata....................--_ 323, 334, tab. 1, pi. 84

undosa....................................... 332

quoyi, Calcarina.................................. 363

R 

Raadshoovenia.................................... 337

radicta, Amphiitegina. 321,322,326,362,363, tab. 4, pi. 90 

papillosa, Amphistegina..... 322, 362, tab. 4, pi. 90

venosa, Amphistegina........ 322, 363, tab. 4, pJ. 90

var., Amphistegina..------------- 363, tab. 4, pi. 90

radiato-marginata, Fissurina....~ 324, 351, tab. 3, pi. 87

Lagena....................................... 351

radiatus, Nautilus................................ 362

raphana, Siphogenerina........... 323, 356, tab. 3, pi. 88

raphanus, Sagrina................................ 356

Siphogenerina................................. 356

(Sagrina)...................................... 356

Uvigerina (Sagrina")........................... 356

costulata, Siphogenerina....................... 356

Rectobolivina bifrons.................... 322, 354, tab. 3

Reef-flat fauna......__....._...........--- 319-320

regularis, Spiroloculina........................... 335

repanda, Eponides................................ 360

menardii pauperata, Pulvinulina ............. 371

reticulata, Ehrenbergina............ 322,367, tab. 4, pi. 90

Truneatulina................................. 361

Reussdla simplex............ 323, 325, 354, tab. 3, pi. 88

spinulosa atlantica............................ 354

sp. A---.___.--.-..--..---. 320, 354, tab. 3, pi. 88 

sp. B—-----„„—------......... 324, 354, tab. 3

Reussella-Bolivina..-..------..--—----------.---- 320

revertens, SpiriUma mvipara........ 320,357, tab. 4, pi. 88

Rhabdogoniumtricarinatum.....................-- 356

rhodiensis, Oaudryina (Siphogaudryina)........--. 331

rhomboidalis, Bolivina........ __ 323, 353, tab. 3, pi. 87

Textularia..................................-- 353

ringensdenticulata, Biloculina..................— 340

striolata, Eiloculina........................... 340

Robertinasubcylindrica........-.. 322, 350, tab. 3, pi. 87

translucens........................ 322, 350, pi. 87

Robulinalimbosa.--.............................. 343

Robulus calcar................... 321, 343, tab. 2, pi. 86

cleridi.....................................--. 343

expansusplanulatus-----.............-------- 343

gibbus........................ 321, 342, tab. 2, pi. 86

limbosus..................... 321, 343, tab. 2, pi. 86

nigriseptus................... 321, 343, tab. 2, pi. 86

orbicularis.................................... 343

sp. \........................ 321, 343, tab. 2, pi. 86

sp. ^......................... 321, S^S, tab. 2, pi. 86

sp. C-.- — - —............... 321, 343, tab. 2, pi. 86

sp. D......................... 321, S4S, tab. 2, pi. 82

robusta, PoUvina................-. 321, 353, tab. 3, pi. 87

compacta, Bolivina............................ 352

Rongelap Atoll, localities arid depths of samples-- 326 

Rongerik lagoon, localities and depths of samples. 326 

RoxaUna poeyi.......... ___.....__-----------— 364

rugosa------------.-------------------.------- 359

rostrata, Anomalmella............ 324, 37^, tab. 5, pi. 91

Biklimina..................... 322, 350, tab. 3, pi. 87

Truneatulina................................. 371

Rotalia beccarii tepida............. 320, 3fO, tab. 4, pi. 89

calcar.......-................................. 319

dubia......................................... 361

menardii...................................... 370

soldanii....................................... 359

spiculotesta ___________._...._.-__.__.. 342

squammosa...............................-.-- 364

Rotaliammina mayori............. 324, 342, tab. 2, pi. 85

rotaliformis, Trochammina........ 324, 84$, tab. 2, pi. 85

Rotaliidae........................................ 356

Rotalina hirsute-,..._...--._._..__-.____-- 370 

truncatultnotdes................----.----....-- 370

rotundatus, Pyropilus............. 324, 364, tab. 4, pi. 90

rubra, Globigerina................................. 369

Olobigerinoides......... 321,323,326,369, tab. 5, pi.91

Mtllepora...............................-..-.. 373

Polytrema....... — . — _..—.......— . — ...— 373

rubrum, Homotrema........................... 319, 320,

323, 373, 374, tab. 5, pis. 82, 92, 93 

Rugidia corticata.................. 323, 362, tab. 4, pi. 89

spinosa....................... 320, 362, tab. 4, pi. 89

rugosa, Discorbina................................ 359

Discorbis..................... 323, 3fi9, tab. 4, pi. 89

Oaudryina.................................... 330

Rosalina...................................... 359

Textularia.... ......__...______.__-- 331

\a, Gaudryina.............................. 331

Gaudryina (Siphogaudryina). 323, 33^, tab. 1, pi. 82 

Rupertia stabiJis................. 324, 373, tab. 5, pi. 91

Kupertiidae--------------....—...——....--. 373

i, Bijarinella........................... 354

Saccamminidae.-_______..___.__..... 328 

sacculijera, Globigerina.........................-.. 369

Globigerinoides........ 321, 323,326, S69, tab. B, pi. 91

fistulosa, Globigerinoides....-- 324, 369, tab. 5 pi. 91

Sagenina frondescens.................. 323, S28, tab. 1

Sagrina bifrons.................................... 354

dimorpha..................................... 355

raphanus...--................................ 356

(Sagrina) bifrons, Siphogenerina.--................ 354

raphanus, Siphogenerina..----................ 366

Uvigerina................................. 356

salisburyi, Pullenia. ___.--____ 322, 367, tab. 4. pi. 90 

samoaensis, Quinqueloculina---.......... 324, SSS pi.£3

Samples, means of estimating percentages__.... 319

treatment for separation--________... 319 

scabra, Lamarckina............... 324, S59, tab. 4, pi. 89

Pulvinulina oblonga........................... 359

Schenckiella primaeva.................... 322,332 tab. 1

schloenbachi, Cnstellaria.....-..................... 344

Schlumbergerina alveoliniformis.... 323, 337, tab. 1, pi. 85

schreibersiana, Virgulina.......................... 352

semialata, Textularia.-........ 321, 329, 350, tab. 1. pi. 83

semi-involUa, Conicospirillina...-- 324, 357, tab. 4, pi. 88 

seminulum, Ouinqueloculina......... 323, SSS, 33fl, pi. 83

semitectaterquemiana, Sigmomorphina.-- 324, 344- tab. 2 

serrata, Ehrenbergina.............................. 367

Hauerina. — .—.............. 320, 337, tab. 1. pi. 84

Sigmoilina edwardsi............... 324, 336, tab. 1, pi. 84

(Sigmoilina) edwardsi, Planispirina............... 336

Sigmomorphina semitecta terquemiana..-. 324, 344,- tab. 2 

simplex, Blphidium.-............. 323, 346, tab. 2, pi. 86

Reussella................. 323, 325, 354, tab. 3, pi. 88

Trimosina.................................... 354

Siphogaudryina---............... ............ —.. 331

(Siphogaudryina) rhodiensis, Gaudryina-....-..... 331

rugulosa, Gaudryina.......... 323, 33^, tab. 1. pi. 82

siphonifera, Gaudryina.--............ 321, 33^, pi. 83

transversaria, Gaudryina-- - ---.... 321, SS1, pi. 83

Siphogenerina btfrons.............................. 354

costata........................................ 356

dimorpha pacifica............. 321, 355, tab. 3, pi. 88

raphana....--..----...--..... 323, 356, tab. 3, pi. 88

raphanus.-----.-.-........................... 356

costulata--................................ 356

(Sagrina) bifrons........................ — ... 354

raphanus...--..-...................- — -.- 356

siphonifera, Gaudryina.--.................... 331, tab. 1

Gaudryina (Siphogaudryina)......... 321, 33^, pi. 83

Textularia.................................... 331

Si/phoninaechinata.------......................... 361

tubulosa ------_._______ 320, 36^, tab. 4. pi. 89

Siphoninoides echinata............. 323, 361, tab. 4, pi. 89

Siphonodosaria abyssorum... -..... 322, 356, tab. 3, pi. 88

lepidula..-—................ 322, 356, tab. 3. pi. 88

Siphotextvloaria—................................. 330

concava.--.. __________ 322, 330, tab. 1, pi. 83 

crispata....................... 321, 330, tab. 1. pi. 83

Silvestria bradyi.--.--............................. 342

soldanii, Gyroidina................ 322, 359, tab. 4, pi. 89

Rotalia....................................... 359

Sorites marginalis................. 323, 348, tab. 3, pi. 82

speciosa, Hauerina................................ 337

spengleri, Calcarina.----.......................... 319,

320, 321, 325, 363, 364, tab. 4, pis 82, 92 

Nautilus—................................... 363

Sphaeridiapapillata..................... 324,362, tab. 4

Sphaeroidina bulloides...-......... 322, 367, tab. 4, pi. 90

compacta............................ 322, 367, tab. 4

corticata-- ____-__-_--__--__--_____. 362

dehiscens.-..-................................. 369

Sphaeroidinella dehiscens...... 322, 326, 369, tab. E, pi. 91

spicata, Buliminella madagascariensis...^...... ... 349

spiculotesta, Carterina.-........... 324, 842, tab. 2, pi. 85

Rotalia.-.—................................. 342

spinata, Triloculina.-............. 323, 339, tab. 2, pi. 85

spinifera, Cassidulina............. 322, 366, tab. 4. pi. 90

dpinosa, Rugidia.-................. 320,362, tab. 4, pi. 89
384 INDEX

spinulosa atlantica, Reussella..-.....--............ 354

spirtdis, Lagena................... 324, 844, tab. 2, pi. 86

Spirillina decorata.---.... 320, 324, 356, 357, tab. 4, pi. 88

decorata var.--.._____-__ 357, tab. 4, pi, 88 

inaequalis................._. 324, 357, tab. 4, pi. 88

tuberculato-limbata............ 324, 357, tab. 4, pi. 88

vivipara densepunctata......... 324,557, tab. 4, pi, 88

resertens......................... 320, 357, pi. 88

Spirolina acicularis................ 323, 348, tab. 3, pi. 87

arietina--.................... 323, 348, tab. 3, pi, 87

Spiroloculina acescata.--................. 324, 334, tab. 1

aequa-— —— — __._____.__ 324, 334, tab. 1

angulata-..................... 323, 334, tab. 1, pi. 84

antittarum.................................... 335

aequa..................................... 334

angulata-----............................. 334

circularis...................... 321, 335, tab. 1, pi. 84

Clara.......................... 323, 335, tab. 1, pi. 84

lirata..................... 320, 335, tab. 1, pi. 84

communis.-.................. 323,535, tab. 1, pi. 84

corrugata...................... 323, 335, tab. 1, pi. 84

cushmani..................................... 336

dor sat a circularis.............................. 335

excavata.--..--................................ 335

foveolata...................... 320,555, tab. l,pl. 84

grata.---..--.-.-................—......__ 334

angulata--......--__..__-------__. 334

grateloupi.--..................... _ _____ 335

acescata--- ------.......................... 334

impressa..._.-._--............................ 335

marshattana.----... -____ 323,335, tab. 1, pi. 84

mayori....................... 324, 336, tab. 1, pi. 84

pianissimo-..—............................... 334

regularls...................................... 335

sp. A——.--————— 324, 336, tab. 1, pi. 84

Sp. B—._———.—..— 324, 836, tab. 1, pi. 84 

Spiroplectammina mittetti......................... 329

sguammosa, Cymbalopora......................... 364

Cymbaloporetta............... 323, 364, tab. 4, pi. 90

Rotalia....................................... 364

sguamosa, Entosolenia............................. 350

Oolina--.-.................... 324, 350, tab. 3, pi. 87

squamosum, Vermiculum......................... 350

stabilis, Rupertia.................. 324, 375, tab. 5, pi. 91

stelligera, Nonionina.............................. 345

striata, Vertebralina..................... 324, 341, tab. 2

striato-punctata, Lagena................. 324, 344, tab. 2

Striaiula, Bolivina—.............. 323, 353, tab. 3, pi. 87

Quingueloculina..........'....................

striolata, Biloculina denticulata............ ..... _.. 340

Biloculina ringens............................. 340

Pyrgodenticulata............. 323, 340, tab. 2, pi. 85

subbertheloti, Discorbis............ 320, 359, tab. 4, pi. 89

subcretacea, Globignrina............ 322, 368, tab. 5, pi. 91

subcuneata, Quinqueloculina crassa. 323,333, tab. 1, pi. 83 

subcylindrica, Bulimina.-......................... 350

Robertina..................... 322, 350, tab. 3, pi. 87

subexcavata, Bolivina-............. 323, 353, tab. 3, pi. 87

subglobosa, Cassidulina........ 322, 366, 367, tab. 4, pi.

var., Cassidulina.................. 367, tab. 4, pi. 90

sublineata, MUiolina circularis..................... 337

suborbicularis, Heterostegina............ __._-. 320,

323, 325, 346, 347, tab. 3, pis. 82, 87 

subplanciana, Triloculina......... 323, 339, tab. 2, pi. 85

subreticulata, Bolivina.................. 324, 353, tab. 3

subrotundata, Oaudryina.......................... 330

subtenuis, Bolivina--.................... 324, 353, tab. 3

sulcata, Quinqueloculina-..-...... 323, 334, tab. 1, pi. 84

tenera, Eponides.................. 322, 359, tab. 4, pi.

Truncatulina................................. 359

tennis, Bolivina..-..-............................. 353

tepida, Rotalia beccarii............. 320, 360, tab. 4, pi. 89

terebra, Eponides.. — ............................. 361

Paumotua-.................. 324, 361, tab. 4, pi. 89

terqutmiana, MUiolina.........______----- 338, 340

Sigmomorphinasemitecta---........ 324, 344, tab. 2

Triloculina............... 323, 338, 340, tab. 2, pi. 85

Textularia........................................ 320

abbreviata..................................... 329

agglutinans-.............. 323, 328, 329, tab. 1, pi. 83

candeiana..................... 323, 329, tab. 1, pi. 83

conica........................ 323, 329, tab. 1, pi. 83

corrugata...................... —+- ------ 329

corrugata..................................... 329

crispata....................................... 330

dupla—...................... 323, 329, tab. 1, pi. 83

folia......................................... 349

foliacea...-- — -........ 323, 529, 330, tab. 1, pi. 83

oceanica.----............. 323, 329, tab. 1, pi. 83

inconspicua................................... 358

jugosa.............................---------- 358

kerifnbtamis.. —.............. 321, 329, tab. 1, pi. 83

mittetti.......-....---....---------- 321, 329, 330

quadrilatera................................... 349

rhomboidalis-----......--------------- — ----- 353

rugosa......-~----.................-—-.--.. 331

semialata................. 321, 329,330, tab. 1, pi. 83

siphohifera............... —................... 331

transversaria.................................. 331

Textulariidae...------.--.-------.---------—— 328

tortuosa, Bolieina........—..-.... 323, 555, tab. 3, pi. 87

transluscens, Nonionella........... 323, 545, tab. 2, pi. 86

Robertina.................-— 322, 350, tab. 3, pi. 87

transversaria, Guadryina (Siphogaudryina). 321,331, pi. 83 

Textularia.—................................ 331

Tretomphalus bulloides—......................... 364

planus........................ 323, 364, tab. 4, pi. 90

triangularis angulata, Oaudryina.. 321, 33/, tab. 1, pi. 83 

tricarinata, MUiolina............................... 340

Triloculina................... 323, 340, tab. 2, pi. 85

Vaginulina................................... 356

tricarinatum, Rhabdogonium...................... 356

Trifarina bradyi................... 321, 366, tab. 3, pi. 88

triformis, Pavonina........... 322, 324, 555, tab. 3, pi. 88

trigonula, Mttiola..... ____--__________- 340

MUiolina...............----------—-..- — .. 340

Triloculina............... 323, 339, 340, tab. 2, pi. 85

Triloculina...——......................... 334, 338, 339

bassensis-..............——... 323, 337, tab. 2, pi. 85

bicarinata................. 323, 33S, 339, tab. 2, pi. 85

bikiniensis...... — ........... 323, 338, tab. 2, pi. 85

earlandi...... _________ 323, 338, tab. 2, pi. 85

involuta....................... 323, 33S, tab. 2, pi. 85

irregularis.................... 323, 33S, tab. 2, pi. 85

kerimbatica......................... 323, 539, pi. 85

linneiana..................................... 338

marshallana.................. 323, 539, tab. 2, pi. 85

oblonga.—.,............. 323, 333, 539, tab. 2, pi. 85

oceanica...................................... 334

planeiana...........................——.. 338, 339

spinata....................... 323, 339, tab. 2, pi. 85

subplanciana..-........—.... 323,339, tab. 2, p. 85

terquemiana............... 323,338,340, tab. 2, pi. 85

tricarinata.................... 323,340, tab. 2, pi. 85

trigonula.................. 323, 339,340, tab. 2, pi. 86

sp. A__--....-.....---.....-.. 320, 3^0, tab. 2, pi. 85

sp. B.......----.............. 322,540, tab. 2, pi. 85

Triloculinellalabiosa.............. 323,33-f, tab. 1, pi. 84

Trimosina simplex...........—................... 354

Trochammina.................................... 342

rotaliformis................... 324, 34,2, tab. 2, pi. 85

Troehamminidae_-------__--..--.---------.-- 342

trullissata, Gaudryina............. 32?.331, tab. 1, pi. 8$

Truncatulina akneriana.......... ________.- 371

echinata...................................... 361

lobatiila-—..........———................ 371

mayori....................................... 371

pseudoungeriana.............................. 372

reticulata..................................... 361

rostrata....—................................. 371

tenera........................................ 359

tubulifera............................._._ 365

variabilis—.................................. 372

trvncatulinoides, Globorotalia...... 322,570, tab. 5, pi. 91

Rotalina—................................... 370

tuberculato-limbata, Spirillina..... 324,357, tab. 4, pi. 88

tuberocapitata, Discorbina......................... 35fr

Discorbis..-................... 321,359, tab. 4, pi. 8»

tuberosa, Dimorphma.-........... 321,344, tab. 2, pi. 86

Tubinfllafumalis................. 324,536, tab. 1, pi. 84

tubulifera, Epistominella.......... 323,365, tab. 4, pi. 90

Truncatulina. —.-—.—................——. 365

tubulosa, Siphonina............... 320,361, tab. 4, pi. 89

tumida, Globorotalia........... 322,326,570, tab. 5, pi. 91

Pulvinulina.................... ._______ 370

tumidum, Astrononion............ 321,345, tab. 2, pi. 86

IT

umbilicatulapacifica, Nonionina.................. 345

umbilicatulumpacificum, Nonion................. 345

undosa, Miliolina................................. 332

Quinqueloculina......———... ——— -..-—— 332

universa, Orbulina....................322,326,369, tab. 5

Uvigerina ampullacea............. 331,355, tab. 3, pi. 88

asperula ampullacea.......................... 355

peregrina.....—.............. 312,366, tab. 3, pi. 88

porrerta...................._ 373,355, tab. 3, pi. 88

fimbriata.—.——.—— 332,355, tab. 3, pi. 88 

proboscidea—................. 331,355, tab. 3, pi. 88

vadescens.................. 331,355, tab. 3, pi. 88

(Sagrina") raphanus........................... 356

vadescens, Uvigerina probostidea... 321,355, tab. 3, pi. 88 

Vaginulina tricarinata---................-.....--- 356

Valvulina davidiana............... 323,35J, tab. 1, pi. 82

Valvulinidae--_................................... 331

»orio6i/is, Cibiddella............... 394,372, tab. 5, pi. 82

Truncatulina-................................ 372

venosa, Ampfiistegina radiata...... 332,365, tab. 4, pi. 90

venosus, Nautilus................................. 363

ventricosa, Parafissurina.......................... 356

Vermiculum oblongum............... ———. — -- 339

squamosum..... ._—————.—————--——— 350

Verneuilinidae——.——.-........................ 330

veniculata, Polystomella.................... — ...- 346

Vertebralina striata...................... 324,54*, tab. 2

vertebralis, Marginopora........................ 319,320,

323,325,54S, 364,, tab. 3, pi. 82 

vesicularis, Gypsina............... 524,373, tab. 5, pi. 82

Orbitulina.................................... 373

Virgulina earlandi—.-............ S24.352, tab. 3, pi. 87

schreibersiana................................. 352

vivipara densepunctata, Spirillina.. 324,357, tab. 4, pi. 88

revertens, Spirittina.................. 320,357, pi. 88

W

Wiesnerella auriculata............. c2±,341, tab. 2, pi. 85

wittiamsoni, Lagena.——.......... ?24,344, tab. 2, pi. 86

williamsoniana, Bulimina........................ 349

BuliminoidfS-................. 324,349, tab. 3, pi. 87

U. S. GOVERNMENT PRINTING OFFICE: 19S4
PLATES 82-93
PLATE 82

[All figures X 15]

FIGURE 1. Gaudryina (Siphogaudryina) rugulosa Cushtnan (p. 331). 

USNM 548610; Bikini 292.

2. Valvulina davidiana Chapman (p. 331).

USNM 548487; Rongerik 51; a, Side view; b, top view.

3. Robulus sp. D. (p. 343).

USNM 548619; Bikini 814.

4. Sorites marginalia (Lamarck) (p. 348).

USNM 548472; Rongerik 21. 

5, 6. Marginopora vertebralis Blainville. (p. 348).

5, USNM 548558; Rongerik 99. 6, USNM 548550; Rongerik 79. 

7, 8. Heterostegina suborbicidaris D'Orbigny (p. 346).

7, USNM 548746; 8, USNM 548747; Eniwetok 145. 

9. Cycloclypeus carpenteri H. B. Brady (p. 347).

USNM 548632; Bikini 817; Megalospheric specimen. 

10, 11. Calcarina spengleri (Gmelin) (p. 363).

10, USNM 548559; Rongerik 99. 11, USNM 548611; Bikini 304; Specimen showing development of arborescent 

spines.

12. Gypsina vesicularis (Parker and Jones) (p. 373). 

USNM 548752; Eniwetok 291.

13. Cibicidella variabilis (D'Orbigny) (p. 372). 

USNM 548536; Rongerik 51.

14. Planorbulina acervalis H. B. Brady (p. 372). 

USNM 548537; Rongerik 51.

15. Carpenteria sp. (p. 373).

USNM 548554; Rongerik 80; a, Top view; 6, side view.

16. Miniacina miniacea (Pallas) (p. 374).

USNM*548755; Eniwetok 306; a, Top view; b, side view.

17. Homotrema rubrum (Lamarck) (p. 373).

USNM 548737; Eniwetok 11; Eroded fragment of an attached specimen.
GEOLOGICAL cJ PROFESSIONAL PAPER 260 PLATE 82

MIS* rij \M.OI s i \IU;K M»F.UMF.N> OF FOR\MIMI-F.KA
GEOLOGICAL SURVEY PROFESSIONAL PAPER 260 PLATE 83

SACCAMMIMDAE, TEXTTLAKIIDAE, \ERYELILIMDAE, VALV CLIMDAE, MILIOLIDAE
PLATE 83

[All figures X 40, except as indicated]

FIGUBE 1. Proteonina sp. (p. 328).

USNM 548745; Eniwetok 145.

2. Textularia agglutinans D'Orbigny (p. 328). 

USNM 548483; Rongerik 51.

3. Textulana candeiana D'Orbigny (p. 329). 

USNM 548484; Rongerik 51.

4. Textularia conica D'Orbigny (p. 329). 

USNM 548555; Rongerik 91.

5. Textulana semialata Cushman (p. 330). 

USNM 548583; Rongelap 468.

6. Textularia dupla Todd, n. sp. (p. 329). 

Holotype, USNM 548609; Bikini 292.

7. Textularia foliacea Heron-Alien and Earland (p. 329). 

USNM 548485; Rongerik 51.

8. Textularia foliacea var. oceanica Cushman (p. 329). 

USNM 548486; Rongerik 51.

9. Siphotextularia cf. S. concava (Karrer) (p. 330). 

USNM 548672; Bikini 1174 (1-2").

10. Siphotextularia crispata (H. B. Brady) (p. 330). 

USNM 548568; Rongelap 456.

11. Textularia kerimbaensis Said (p. 329). 

USNM 548629; Bikini 817.

12. Bigenerina sp. (p. 330).

USNM 548671; Bikini 1174 (1-2").

13. Gaudryina cf. G. pauperata Earland (p. 330). 

USNM 548748,X80; Eniwetok 218.

14. Gaudryina triangularis var. angulata Cushman (p. 331). 

USNM 548649; Bikini 819.

15. Gaudryina trullissata Todd, n. sp. (p. 331).

Holotype, USNM 548622,X80; Bikini 815.

16. Gaudryina (Siphogaudryina) transversaria (H. B. Brady) (p. 331). 

USNM 548612,X80; Rongelap 463.

17. Gaudryina (Siphogaudryina) siphonifera (H. B. Brady) (p. 331). 

USNM 548576; Rongelap 463.

18. Clavulina angularis D'Orbigny (p. 332). 

USNM 548591; Rongelap 483.

19. Clavulina pacifica Cushman (p. 332). 

USNM 548488; Rongerik 51.

20. Karreriella bradyi (Cushman) (p. 332). 

USNM 548698; Bikini 1175.

21. QuinQueloculina anguina var. arenata Said (p. 332). 

USNM 548489; Rongerik 51.

22. Quinoueloculina crassa var. subcuneata Cushman (p. 333). 

USNM 548470; Rongerik 18.

23. Quinc/ueloculina parkeri (H. B. Brady) (p. 333). 

USNM 548491,X15; Rongerik 51.

24. Quino^eloculina parkeri (H. B. Brady) var. (p. 333) 

USNM 548492,X15; Rongerik 51."

25. QuinQueloculina cf. Q. samoaensis Cushman (p. 333). 

USNM 548757; Eniwetok 310.

26. Quinqueloculina bradyana Cushman (p. 332).

USNM 548614; Bikini 814; a, Side view; 6, apertural view

27. Quinqueloculina distorqueata Cushman, n. sp. (p. 333). 

Holotype, USNM 548545; Rongerik 79.

28. Quinqueloculina neostriatula Thalmann (p. 333). 

USNM 548464; Rongerik 3.

29. Quinqueloculina cf. Q. seminulum (Linne") (p. 333) 

USNM 548562; Rongelap 111.

30. Quinqueloculina cf. Q. ferussacii D'Orbigny (p. 333).

USNM 548599; Bikini 190; a, Side view; 6, apertural view

31. Quinqueloculina bidentata D'Orbigny (p. 332).

USNM 548490; Rongerik 51; a, Side view; 6, apertural view.
PLATE 84

[All figures X 40, except as indicated]

FIGURES 1, 2. Quinqueloculina sulcata D'Orbigny (p. 334).

TJSNM 548493; Rongerik 51. 

3, 4. Miliolinella australis (Parr) (p. 334).

3, TJSNM 548476, X 80; Rongerik 31. 4, TJSNM 548577, X 80; Rongelap 463. 

5, 6. Triloculinella labiosa (D'Orbigny) (p. 334).

5, TJSNM 548761, X 80; Eniwetok 317. 6, TJSNM 548494, X 80; Rongerik 51; Young specimen.

7. Spiroloculina circularis Chapman (p. 335) 

TJSNM 548650; X 80; Bikini 819.

8. Spiroloculina marshallana Todd, n. sp. (p. 335). 

Holotype, TJSNM 548615; Bikini 814.

9. Spiroloculina clara Cushman (p. 335). 

TJSNM 548496 ; Rongerik 51.

10. Spiroloculina clara var. lirata Cushman, n. var. (p. 335).

Holotype, TJSNM 548477; Rongerik 31. 

11, 12. Spiroloculina angulata Cushman (p. 334). 

TJSNM 548495; Rongerik 51.

13. Spiroloculina communis Cushman and Todd (p. 335). 

TJSNM 548738, Eniwetok 116.

14. Spiroloculina foveolata Egger (p. 335). 

TJSNM 548497; Rongerik 51.

15. Spiroloculina mayori Cushman (p. 336).

TJSNM 548613; Bikini 1028; a, Front view: b, end view.

16. Massilina pianola Cushman (p. 334).

TJSNM 548762; Eniwetok 317. 

17, 18. Spiroloculina corrugaia Cushman and Todd (p. 335).

17, TJSNM 548739; Eniwetok 116. 18, TJSNM 548758; Eniwetok 310.

19. Spiroloculina sp. B. (p. 336).

TJSNM 548478; Rongerik 31.

20. Sigmoilina edwardsi (Schlumberger) (p. 336). 

TJSNM 548602, X 80; Bikini 218 (12-16"). 

21, 22. Spiroloculina sp. A. (p. 336).

21, TJSNM 548584; Rongelap 468. 22, TJSNM 548592; Rongelap 483.

23. Hauerina milletti Cushman, n. sp. (p. 337). 

Holotype, TJSNM 548501; Rongerik 51.

24. Tubinella funalis (H. B. Brady) (p. 336). 

TJSNM 548498, X 80; Rongerik 51.

25. Articulina pacifica Cushman (p. 336)7

TJSNM 548546, X 80; Rongerik 79. 

26, 27. Hauerina serrata Cushman, n. sp. (p. 337).

26, Paratype, TJSNM 548765; 27 Holotype, TJSNM 548764; Eniwetok 321. 

28, 29. Hauerina involuta Cushman (p. 337).

TJSNM 548500; Rongerik 51. 

30, 31. Hauerina diversa Cushman (p. 337).

30, TJSNM 548499; Rongerik 51. 31, TJSNM 548593; Rongelap 483. 

32. Hauerina bradyi Cushman (p. 336).

TJSNM 548760; Eniwetok 314.
GEOLOGICAL PROFESSIONAL PAPER 260 PLATE 84

MILIOLIDAE
GEOLOGICAL SURVEY PROFESSIONAL PAPER 260 PLATE 85

MILIOLIDAE, OPHTHALMID IIDAE, TROCHAMMIiMDAE
PLATE 85

[All figures X 40, except as indicated]

FIGURE 1. Schlumbergerina alveoliniformis (H: B. Brady) (p. 337). 

USNM 548569, X 15; Rongelap 456.

2. Triloculina cf. T. bicarinata D'Orbigny (p. 338).

USNM 548556, X 15; Rongerik 99; a, Side view; &, apertural view.

3. Triloculina earlandi Cushman, n. sp. (p. 338). 

Holotype, USNM 548502; Rongerik 51.

4. Triloculina involuta Todd, n. sp. (p. 338).

Holotype, USNM 548742, X 15; Eniwetok 118. 

5-7. Triloculina cf. T. oblonga (Montagu) (p. 339). 

USNM 548504; Rongerik 51.

8. Triloculina spinata Cushman (p. 339). 

USNM 548595, X 80; Rongelap 483.

9. Triloculina terquemiana (H. B. Brady) (p. 340).

USNM 548557; Rongerik 99; a, Side view; &, apertural view. 

10, 11. Triloculina kerimbatica (Heron-Alien and Earland) (p. 339).

10, USNM 548585; Rongelap 468; a, Side view; b, apertural view.

11, USNM 548503; Rongerik 51. 

12. Triloculina irregularis (D'Orbigny) (p. 338).

USNM 548596; Bikini 37; a, Side view; 6, apertural view. 

; 13. Triloculina marshallana Todd, n. sp. (p. 339).

Holotype, USNM 548594; Rongelap 483; a, Side view; &, apertural view. 

14. Triloculina cf. T. bassensis Parr. (p. 337).

USNM 548597; Bikini 177; a, Side view; b, apertural view. 

15, 16. Triloculina tricarinata D'Orbigny (p. 340). 

USNM 548505; Rongerik 51.

17. Triloculina subplanciana Cushman, n. sp. (p. 339). 

Holotype, USNM 548540, X 80; Rongerik 54.

18. Triloculina trigonula (Lamarck) (p. 340). 

USNM 548628; Bikini 816.

19. Triloculina bikiniensis Todd, n. sp. (p. 338). 

Holotype, USNM 548598; Bikini 177.

20. Triloculina sp. A. (p. 340).

USNM 548756; Eniwetok 307.

21. Triloculina sp. B. (p. 340).

USNM 548635; Bikini 818; a, Side view; b, apertural view.

22. Pyrgo denticulata (H. B. Brady) (p. 340). 

USNM 548506; Rongerik 51.

23. Pyrgo denticulata var. striolata (H. B. Brady) (p. 340). 

USNM 548507, X 50; Rongerik 51.

24. Pyrgo lucernula (Schwager) (p. 340). 

USNM 548659; Bikini 1170.

25. Pyrgo millettii (Cushman), juv. (p. 341). 

USNM 548616; Bikini 814.

26. Pyrgo sp. (p. 341).

USNM 548617; Bikini 814.

27. Cornuspira planorbis Schultze (p. 341). 

USNM 548471, X 80; Rongerik 21.

28. Planispirina exigua (H. B. Brady) (p. 341). 

USNM 548769, X 80; Eniwetok 329.

29. Nubecularia lacunensis Chapman (p. 341). 

USNM 548544, X 80; Rongerik 56.

30. Wiesnerella auriculata (Egger) (p. 341). 

USNM 548754, X 80; Eniwetok 304. 

31-34. Parrina bradyi (Millett) (p. 342). 

USNM 548547; Rongerik 79.

35. Trochammina cf. T. rotaliformis J. Wright (p. 342). 

USNM 548749, X 80; Eniwetok 218.

36. Rotaliammina mayori Cushman (p. 342).

USNM 548586, X 80; Rongelap 468; Ventral view, showing chamber walls.

37. Carterina spiculotesta (Carter) (p. 342). 

USNM 548618, X 80; Bikini 814.
PLATE 86

[All figures X 40, except as indicated]

FIGURE 1. Robulus limbosus (Reuss) (p. 343). 

USNM 548699; Bikini 1175.

2. Robulus gilbus (D'Orbigny) (p. 342). 

USNM 548630; Bikini 817.

3. Robulus nigriseptus (Koch) (p. 343). 

USNM 548673; Bikini 1174 (1-2").

4. Robulus calcar (Linne) (p. 342).

USNM 548636, X 80; Bikini 818.

5. Robulus sp. B. (p. 343).

USNM 548674; Bikini 1174 (1-2").

6. Robulus sp. A. (p. 343).

USNM 548651; Bikini 819.

7. Astacolus planulata Galloway and Wissler (p. 343). 

USNM 548676; Bikini 1174 (1-2").

8. Robulus sp. C. (p. 343).

USNM 548675; Bikini 1174 (1-2").

9. Astacolus bradyi (Cushman), juv. (p. 343). 

USNM 548652; Bikini 819.

10. Dentaltna cf. D. filiformis (D'Orbigny) p. 343). 

USNM 548653; Bikini 819.

11. Nodosaria sp. A. (p. 344).

USNM 548717; Bikini 1176 (1-2").

12. Nodosaria sp. B. (p. 344).

USNM 548677; Bikini 1174 (1-2").

13. Nodosaria sp. C. (p. 344).

USNM 548678; Bikini 1174 (1-2").

14. Nodosaria pauciloculata Cushman, juv. (p.[343). 

USNM 548740; Eniwetok 117.

15. Frondicularia sp. A of Boomgaart (p. 344). 

USNM 548631; Bikini 817.

16. Lagena acuticosta Reuss (p. 344).

USNM 548718, X 80; Bikini 1176 (1-2").

17. Lagena globosa (Montagu) (p. 344).

USNM 548719, X 80; Bikini 1176 (1-2").

18. Lagena gracilis Williamson (p. 344). 

USNM 548637, X 80; Bikini 818.

19. Lagena spiralis H. B. Brady (p. 344). 

USNM 548759, X 80; Eniwetok 312.

20. Lagena williamsoni (Alcock) (p. 344).

USNM 548700, X 80; Bikini 1175. 

21, 22. Dimorphina tuberosa D'Orbigny (p. 344).

USNM 548679; Bikini 1174 (1-2"). 

23, 24. Polymorphinella pacifica Cushman and Hanzawa (p. 344).

USNM 548680; Bikini 1174 (1-2").

25. Nonion pacificum (Cushman) (p. 345). 

USNM 548638; Bikini 818.

26. Nonion pompilioides (Fichtel and Moll) (p. 345). 

USNM 548701; Bikini 1175.

27. Astrononion tumidum Cushman and Edwards (p. 345). 

USNM 548639, X 80; Bikini 818.

28. Nonionella sp. B. (p. 345).

USNM 548563, X 80; Rongelap 111.

29. Nonionella translucens Cushman (p. 345). 

USNM 548552, X 80; Rongerik 80.

30. Elphidium advenum (Cushman) (p. 345). 

USNM 548553, X 80; Rongerik 80.

31. Elphidium advenum var. dispar Cushman, n. var. (p. 346). 

Holotype, USNM 548508, X 80; Rongerik 51.

32. Elphidium jenseni (Cushman) (p. 346). 

USNM 548509, X 80; Rongerik 51.

33. Elphidium simplex Cushman (p. 346). 

USNM 548510, X 80; Rongerik 51.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 260 PLATE 86

LAGEIMDAE, POL\ MORPHLMDAE, ISON1OMDAE
GEOLOGICAL SUR\ EY PROFESSIONAL PAPER 260 PLATE 87

CAMERIMDAh, PEJNFROPLIDAE, ALVEOLINELLID4E, HETKROHELICIDVE, Bl LIMIMDAE
PLATE 87

[All figures X 80, except as indicated]

FIGURE 1. Operculina ammonoides (Gronovius). USNM 548770, X 15; Eniwetok 329. (p. 346).

2. Heterostegina suborbicularis D'Orbigny. USNM 548548, X 15; Rongerik 79. (p. 346).

3. Peneroplis sp. USNM 548564, X 40; Rongelap 111. (p. 348). 

4, 5. Spirolina arietina (Batsch). USNM 548511, X 40; Rongerik 51. (p. 348).

6. Spirolina acicularis (Batsch). USNM 548565; Rongelap 111. (p. 348).

7. Monalysidium politum Chapman. USNM 548549; Rongerik 79. (p. 348).

8. Neoalveolina pulchra (D'Orbigny). USNM 548570, X 40; Rongelap 456. (p. 348).

9. Gumbelina? marshallana Todd, n. sp. Holotype, USNM 548727; Bikini 1176- (5-7"). (p. 349). 

10, 11. Bolivindla folia (Parker and Jones). USNM 548512; Rongerik 51. (p. 349). 

12, 13. Bolivinella folia var. ornata Cushman. USNM 548513; Rongerik 51. (p. 349). 

14, 15. Buliminella milletti Cushman. USNM 548514; Rongerik 51. (p. 349). 

16, 17. Buliminoides williamsoniana (H. B. Brady). USNM 548515; Rongerik 51. (p. 349).

18. Robertina subcylindrica (H. B. Brady). USNM 548728, X 40; Bikini 1176 (5-7"). (p. 350).

19. Robertina translucens Cushman and Parker. USNM 548690, X40; Bikini 1174 (10-14"). (p. 350).

20. Pseudobulimina sp. USNM 548578, X 40; Rongelap 463. (p. 350).

21. Bulimina aculeata D'Orbigny. USNM 548702, X 40; Bikini 1175. (p. 350).

22. Bulimina affinis D'Orbigny. USNM 548729, X 15; Bikini 1176 (5-7"). (p. 350).

23. Bulimina costata D'Orbigny. USNM 548703; Bikini 1175. (p. 350).

24. Bulimina rostrata H. B. Brady. USNM 548704; Bikini 1175. (p. 350).

25. Bulimina fijiensis Cushrnan. USNM 548473; Rongerik 21. (p. 350).

26. Oolina squamosa (Montagu). USNM 548579; Rongelap 463. (p. 350).

27. Fissurina circularis Todd, n. sp. Holotype, USNM 548767; Eniwetok 326. (p. 351).

28. Fissurina lacunata (Burrows and Holland). USNM 548654; Bikini 819. (p. 351).

29. Fissurina radiato-marginata (Parker and Jones). USNM 548604; Bikini 218 (12-16"). (p. 351).

30. Fissurina milletti Todd, n. sp. Holotype, USNM 548743; Eniwetok 119. (p. 351).

31. Fissurina formosa (Schwager). USNM 548603; Bikini 218 (12-16"). (p. 351).

32. Fissurina sp. A. USNM 548763; Eniwetok 318. (p. 351).

33. Fissurina sp. B. USNM 548655; Bikini 819. (p. 352).

34. Fissurina sp. C. USNM 548705; Bikini 1175. (p. 352).

35. Virgulina earlandi Cushman. USNM 548605; Bikini 218 (12-16"). (p. 352).

36. Bolivina abbreviata Heron-Alien and Earland. USNM 548720; Bikini 1176 (1-2"). (p. 352).

37. Bolivina compacta Sidebottom. USNM 548706; Bikini 1175. (p. 352).

38. Bolivina nitida H. B. Brady. USNM 548640; Bikini 818. (p. 352).

39. Bolivina pseudopygmaea Cushrnan. USNM 548479; Rongerik 31. (p. 352).

40. Bolivina robusta H. B. Brady. USNM 548707; Bikini 1175. (p. 353). 

41, 42. Bolivina rhomboidalis (Millett). USNM 548516; Rongerik 51. (p. 353).

43. Bolivina striatula Cushman. USNM 548768; Eniwetok 326. (p. 353).

44. Bolivina subexcavata Cushrnan and Wickenden. USNM 548517; Rongerik 51. (p. 353). 

45-47. Bolivina tortuosa H. B. Brady. 45, 46, USNM 548518; 47, USNM 548519; Rongerik 51. (p. 353).
PLATE 88

[All figures X 80, except as indicated]

FIGURES 1, 2. Reus&ella simplex (Cushman) (p. 354).

USNM 548522; Rongerik 51. 

3. Reussella sp. A. (p. 354).

USNM 548753; Eniwetok 301. 

4, 5. Loxostomum convallarium (Millett) (p. 353).

4, USNM 548520; Rongerik 51. 5, USNM 548567; Rongelap 133.

6. Loxostomum karrerianum (H. B. Brady) (p. 353). 

USNM 548691; Bikini 1174 (10-14").

7. Loxostomum limbatum (H. B. Brady) (p. 353). 

USNM 548521; Rongerik 51.

8. Loxostomum mayori (Cushman) (p. 354). 

USNM 548766; Eniwetok 321.

9. Bifarinella mackinnonii (Millett) (p. 354).

USNM 548571, X 40; Rongelap 456. , 10. Pavonina triformis Parr (p. 355).

USNM 548730, X 40; Bikini 1176 (5-7").

11. Bitubulogenerina sp. (p. 354).

USNM 548587; Rongelap 468. ;

12. Chrysalidinella fijiensis Cushman (p. 355). 

USNM 548480, X 40; Rongerik 31.

13. Uvigerina cf. U. peregrina Cushman (p. 355).

USNM 548692, X 40; Bikini 1174 (10-14"). 

14, 15. Uvigerina porrecta H. B. Brady (p. 355). 

USNM 548641; Bikini 818.

16. Uvigerina porrecta var. fimbriata Sidebottom (p. 355). 

USNM 548580; Rongelap 463.

17. Uvigerina proboscidea var. vadescens Cushman (p. 355). 

USNM 548681; Bikini 1174 (1-2").

18. Uvigerina proboscidea Schwager (p. 355). 

USNM 548660, X 40; Bikini 1170.

19. Uvigerina ampullacea H. B. Brady (p. 355).

USNM 548731, X 40; Bikini 1176 (5-7").

20. Angulogerina albatrossi var. ornata Cushman (p. 356). 

USNM 548656, X 40; Bikini 819.

21. Trifarina bradyi Cushman (p. 356). 

USNM 548623; Bikini 815.

22. Siphogenerina dimorpha var. pacifica Cushman (p. 355).

USNM 548708; Bikini 1175. 

23, 24. Siphogenerina raphana (Parker and Jones) (p. 356).

USNM 548620, X 40; Bikini 814; 23, Megalospheric specimen; 24, microspheric specimen.

25. Parafissurina sp. (p. 356). ' 

USNM 548734; Bikini 1176 (5-7").

26. Siphonodosaria abyssorum (H. B. Brady) (p. 356).

USNM 548732, X 40; Bikini 1176 (5-7"). j 

27, 28. Siphonodosaria lepidula (Schwager) (p. 356). 

USNM 548721, X 40; Bikini 1176 (1-2").

29. Pleurostomella bolivinoides Schubert (p. 356). ; 

USNM 548733; Bikini 1176 (5-7"). I

30. Spirillina vivipara var. revertens Rhumbler (p. 357). 

USNM 548468; Rongerik 12.

31. Spirillina vivipara var. densepunctata Cushman (p. 357).

USNM 548551; Rongerik 79.

32, 33. Spirillina decorata H. B. Brady (p. 356).

USNM 548465; Rongerik 3.

34. Spirillina tuberculato-limbata Chapman (p. 357). \ 

USNM 548657; Bikini 819.

35. Spirillina decorata H. B. Brady var. of Sidebottom (p. 357).

USNM 548581; Rongelap 463.

36, 37. Spirillina inaequalis H. B. Brady (p. 357).

USNM 548523; Rongerik 51.

38. Conicospirillina semi-involuta Cushman (p. 357). 

USNM 548524; Rongerik 51.

39. Conicospirillina sp. (p. 357).

USNM 548606; Bikini 218 (12-16").
GEOLOGICAL SURVEY PROFESSIONAL PAPER 260 PLATE

BULIMINIDAE, ELLIPSOIDINIDAE, ROTALIIDAE
(JEOLOOICAL SURVEY PROFESSIONAL PAPER 260 PLATE 89

ROT \L1I DAE, PEG ID IIDAL
PLATE 89

[All figures X 80, except as indicated]

FIGURE 1. Patellina advena var. altifor'mis Cushman (p. 357).

USNM 548582; Rongelap 463; a, Dorsal view; 6, side view.

2. Patellinella fijiana Cushman (p. 358). 

USNM 548642; Bikini 818.

3. Patellinella jugosa (H. B. Brady) (p. 358). 

USNM 548469; Rongerik 12.

4. Patellinella inconspicua (H. B. Brady) (p. 358). 

USNM 548541; Rongerik 54.

5. Discorbis frustata Cushman (p. 358). 

USNM 548526; Rongerik 51.

6. Discorbis crustata Cushman (p. 358). 

USNM 548588; Rongelap 468.

7. Discorbis patelliformis (H. B. Brady) (p. 359).

USNM 548529; Rongerik 51; a, Ventral view; b, side view. 

8, 9. Discorbis micens Cushman (p. 358).

USNM 548527, X 40; Rongerik 51; 8, Dorsal view; 9, ventral view. 

10, 11. Discorbis opima Cushman (p. 358).

USNM 548528, X 40; Rongerik 51; 10, Dorsal view; 11, ventral view. 

12, 13. Discorbis concinna (H. B. Brady) (p. 358).

USNM 548525, X 40; Rongerik 51; 12, Ventral view; 13, dorsal view.

14. Discorbis rugosa (D'Orbigny) (p. 359).

USNM 548624, X 40; Bikini 815; a, Dorsal view; b, ventral view.

15. Discorbis subbertheloti Cushman (p. 359).

USNM 548643; Bikini 818; a, Dorsal view; b, ventral view.

16. Discorbis tuberocapitata (Chapman) (p. 359). .

USNM 548574, X 40; Rongelap 462; a, Dorsal view; b, side view.

17. Lamarckina scabra (H. B. Brady) (p. 359). 

USNM 548709, X 40; Bikini 1175.

18. Heronallenia sp. (p. 359).

USNM 548607; Bikini 218 (12-16").

19. Gyroidina soldanii D'Orbigny (p. 359).

USNM 548693, X 40; Bikini 1174 (10-14").

20. Eponides tenera (H. B. Brady) (p. 359).

USNM 548682, X 40; Bikini 1174 (1-2").

21. Osangularia bengalensis (Schwager) (p. 360). 

USNM 548722, X 40; Bikini 1176 (1-2").

22. Rotalia cf. R. beccarii var. tepida Cushman (p. 360). 

USNM 548600; Bikini 218.

23. Hoglundina elegans (D'Orbigny) (p. 360).

USNM 548694, X 15; Bikini 1174 (10-14"). 

24, 25. Poroeponides cribrorepandus Asano and Uchio (p. 360).

24, USNM 548590, X 15; Ventral view; 25, USNM 548589, X15; Dorsal view; Rongelap 468.

26. Epistomaroides polystomelloides (Parker and Jones) (p. 360). 

USNM 548566, X 15; Rongelap 111.

27. Mississippina concentrica (Parker and Jones) (p. 360).

USNM 548621, X 40; Bikini 814; a, Dorsal view; b, ventral view.

28. Paumotua terebra (Cushman) (p. 361).

USNM 548633, X 40; Bikini 817; a, Dorsal view, b, ventral view. 

29, 30. Siphonina tubulosa Cushman (p. 361).

USNM 548530; Rongerik 51; 29, Dorsal view; 30, ventral view. 

31, 32. Siphoninoides echinata (H. B. Brady) (p. 361).

USNM 548474, X 40; Rongerik 21.

33. Baggina philippinensis var. pilulifera Cushman and Todd (p. 361). 

USNM 548625, X 15; Bikini 815.

34. Rugidia corticata (Heron-Alien and Earland) (p. 362).

USNM 548475; Rongerik 21. 

35-37. Rugidiaf spinosa Cushman, n. sp. (p. 362).

35, 36, Paratypes, USNM 548543; 37, Holotype, USNM 548542; Rongerik 54.
PLATE 90

[All figures X 40, except as indicated

FIGURES 1, 2. Amphistegina madagascariensis D'Orbigny (p. 362).

USNM 548531; Rongerik 51; 1, Ventral view; 2, dorsal view.

3. Amphistegina radiata (Fichtel and Moll) (p. 362). 

USNM 548741; Eniwetok 117.

4. Amphistegina radiata (Fichtel and Moll) var. (p. 363).

USNM 548572, X 15; Rongelap 456; a, Dorsal view; 6. ventral view. 

5, 6. Amphistegina radiata var. papillosa Said (p. 362).

5, USNM 548634; Bikini 817. 6, USNM 548575, X 15; Rongelap 462; a, Dorsal view; 6, ventral view.

7. Amphistegina radiata var. venosa (Fichtel and Moll) (p. 363). 

USNM 548644; Bikini 818.

8. Pyropilus rotundatus Cushman (p. 364).

USNM 548560; Rongerik 99. 

9-12. Calcarina hispida H. B. Brady (p. 363).

USNM 548532; Rongerik 51; 9, 10, Ventral views; 11, 12, dorsal views. 10, 11, Young specimens. 

13, 14. Cymbaloporetta bradyi (Cushman) (p. 364).

USNM 548533; Rongerik 51; 13, Ventral view; 14, dorsal view. 

15, 16. Cymbaloporetta squammosa (D'Orbigny) (p. 364).

USNM 548534; Rongerik 51; 15, Ventral view; 16, dorsal view. 

17, 18. Tretomphalus planus Cushman (p. 364).

USNM 548751; Eniwetok 290; 17, Dorsal view; 18, side view.

19. Epistominella cf. E. obtusa (Burrows and Holland) (p. 365).

USNM 548683; Bikini 1174 (1-2"); a, Dorsal view; 6, ventral view.

20. Epistominella tubulifera (Heron-Alien and Earland) (p. 365).

USNM 548744; Eniwetok 119; a, Dorsal view; 6, ventral view.

21. Epistominella sp. (p. 365).

USNM 548645, X 80; Bikini 818; a, Dorsal view; 6, peripheral view.

22. Cassidulina angulosa Cushman (p. 365). 

USNM 548726; Bikini 1175.

23. Cassidulina elegans Sidebottom (p. 366). 

USNM 548646; Bikini 818.

24. Cassidulina costatula Cushman (p. 365).

USNM 548684; Bikini 1174 (1-2"); a, Side view; 6, peripheral view.

25. Cassidulina delicata Cushman (p. 365).

USNM 548710; Bikini 1175. 

26, 27. Cassidulina gemma Todd, n. sp. (p. 366).

26, Paratype, USNM 548669; 27, Holotype, USNM 548668; Bikini 1172. 

28, 29. Cassidulina marshallana Todd, n. sp. (p. 366).

28, Holotype, USNM 548685, X 15; 29, Paratype, USNM 548686, X15; Bikini 1174 (1-2"). 

30-32. Cassidulina subglobosa H. B. Brady (p. 367).

USNM 548687; Bikini 1174 (1-2").

33. Cassidulina subglobosa H. B. Brady var. (p. 367). 

USNM 548711; Bikini 1175.

34. Cassidulina cf. C. spinifera Cushman and Jarvis (p. 366). 

USNM 548661; Bikini 1170.

35. Cassidulina minuta Cushman (p. 366). 

USNM 548466, X80; Rongerik 3.

36. Ehrenbergina pacifica Cushman (p. 367). 

USNM 548712, X 80; Bikini 1175.

37. Ehrenbergina reticulata Cushman (p. 367). 

USNM 548735; Bikini 1176 (5-7").

38. Pullenia salisburyi R. E. and K. C. Stewart (p. 367). 

USNM 548713; Bikini 1175.

39. Sphaeroidina bulloides D'Orbigny (p. 367). 

USNM 548695; Bikini 1174 (10-14").
GEOLOGICAL SURVEY PROFESSIONAL PAPER 260 PLATE 90

AMPHISTEGIMDAE, CALCAKIMDAE, CYMBALOPORIDAE, CASSIDL'LIMDAL, CHILOSToM ELLIDAE
GEOLOGICAL SURVEY PROFESSIONAL PAPER 260 PLATE 91

GLOBIGERIMDAE, GLOBOROTAL1IDAE, ANOMALIMDAK, PLANORB L LINIDAE, RUPERT1IDAE
PLATE 91

[All figures X 40, except as indicated]

FIGURE 1. Globigerina subcretacea Lomnicki (p. 368). 

USNM 548662; Bikini 1170.

2. Globigerina bulloides D'Orbigny (p. 368). 

USNM 548723; Bikini 1176 (1-2").

3. Globigerina? sp. (p. 368).

USNM 548750, X 80; Eniwetok 288.

4. Globigerina ef. G. conglomerata Schwager (p. 368). 

USNM 548724; Bikini 1176 (1-2").

5. Globigerinoides elongata (D'Orbigny) (p. 368). 

USNM 548664; Bikini 1170.

6. Globigerinoides rubra (D'Orbigny) (p. 369). 

USNM 548658; Bikini 819.

7. Globigerinoides sacculifera (H. B. Brady) (p. 369). 

USNM 548626, X 15; Bikini 815.

8. Globigerinella aequilateralis (H. B. Brady) (p. 369).

USNM 548647; Bikini 818.

9, 10. Hastigerinella digitata (Rhumbler) (p. 369). 

USNM 548601; Bikini 218.

11. Pulleniatina obliquiloculata (Parker and Jones) (p. 369). 

USNM 548714; Bikini 1175.

12. Globigerinoides conglobata (H. B. Brady) (p. 368). 

USNM 548663, X 15; Bikini 1170.

13. Globigerinoides sacculifera var. fistulosa (Schubert) (p. 369). 

USNM 548736, X 15; Bikini 1176 (5-7").

14. Sphaeroidinella dehiscens (Parker and Jones) (p. 369). 

USNM 548648, X 15; Bikini 818.

15. Candeina nitida D'Orbigny (p. 370). 

USNM 548725; Bikini 1176 (1-2").

16. Globorotalia crassa (D'Orbigny) (p. 370). 

USNM 548670; Bikini 1172.

17. Globorotalia cf. G. Ursula (D'Orbigny) (p. 370). 

USNM 548665; Bikini 1170.

18. Globorotalia truncatulinoides (D'Orbigny) (p. 370).

USNM 548666; Bikini 1170; a, Ventral view; b, side view.

19. Globorotalia menardii (D'Orbigny) (p. 370). 

USNM 548715, X 15; Bikini 1175.

20. Globorotalia menardii var. fimbriata (H. B. Brady) (p. 370). 

USNM 548716, X 15: Bikini 1175.

21. Globorotalia, tumida (H. B. Brady) (p. 370). 

USNM 548667, X 15; Bikini 1170.

22. Anomalina polymorpha Costa (p. 370).

USNM 548696, X 15; Bikini 1174 (10-14").

23. Anomalina cf. A. glabrata Cushman (p. 370). 

USNM 548481; Rongerik 31.

24. Anomalinella rostrata (H. B. Brady) (p. 371).

USNM 548535; Rongerik 51. 

25, 26. Cibicides cicatricosus (Schwager) (p. 371).

USNM 548697; Bikini 1174 (10-14"). 

27, 28. Cibicides lobatulus (Walker and Jacob) (p. 371;.

USNM 548627; Bikini 815; 27, Dorsal view; 28, ventral view. 

29,30. Cibicides mayori (Cushman) (p. 371).

29, USNM 548561; Rongelip 71; Dorsal view. 30, USNM 548608; Bikini 218 (12-16"); Ventral view.

31. Cibicides cf. C. pseudoungerianus (Cushman) (p. 372). 

USNM 548688, Bikini 1174 (1-2").

32. Cibicidina sp. (p. 372).

USNM 548689; Bikini 1174 (1-2"); a, Dorsa, view; b, ventral view.

33. Rupertia stabilis Wallich (p. 373). 

USNM 548573; Rongelap 456. 

34-36. Planorbulina acervalis H. B. Brady (p. 372).

USNM 548538; Rongerik 51. 34, 35, Ventral views, X 40; 36, dorsal view, X 50. Note that specimens 34 and 36

were attached by the dorsal sides, while specimen 35 appears to have been attached by the ventral side. 

37, 38. Acervulina inhaerens Schultze (p. 372).

USNM 548482; Rongerik 31. 37, Ventral view, X 40; 38, dorsal view, X 50. 

39. Gypsina globula (Reuss) (p. 373). 

USNM 548539; Rongerik 51.
PLATE 92

[All figures X 20]

FIGURES 1-7. Calcarina spengleri (Gmelin) (p. 363).

Free specimens sectioned and photographed by W. Storrs Cole. 1, 3-6, Horizontal sections; 2, 7, vertical sections. 

1, 2, From reef off village of Odomari, Okinawa Shima, Ryukyu Islands. 3-7, From Arno Atoll, Marshall Islands. 

8. Homotrema rubrum (Lamarck) (p. 373).

USNM 548772; Bikini 314. Specimen encrusting a cavity made by a boring organism in algal material. Siliceous 

sponge spicules protrude from the apertures.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 260 PLATE 92

CALCARINIDAE AND HOMOTREMIDAE
PL ATI: 03

IKIMOTKI.M1DV1
PLATE 93

[Both figures X 14^]

FIGUKE 1. Gypsina plana (Carter) (p. 373).

TJSNM 548771; Bikini 788, Bikini Lagoon. Encrustation on Halimeda segments showing cellular structure in the 

cut section and the indistinct honeycomb pattern of areolae on the surface, a, Halimeda in cut section; b, cellular 

structure of Gypsina plana in cut section; c, surface of Gypsina plana. 

2. Homotrema rubrum (Lamarck) (p. 373).

TJSNM 548772; Bikini 314. Three specimens, at a, b, and c, encrusting a cavity in algal material. Specimen a is 

white, b is pink, and c is red. Siliceous sponge spicules protrude from the high, conical apertural areas. Surface 

areolae are best shown on specimen c between the apertural cones.

o