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Page 1

INTRODUCTION

This report provides estimates of mean lengths at age for 18 species of Northwest Atlantic finfish and shellfish,
together with maximum ages observed for selected finfish species collected during Northeast Fisheries Center (NEFC)
bottom trawl surveys (Table 1). If growth rates are sufficiently different between sexes or geographical areas, separate
mean lengths are given for each sex or area for which data are available. The information used to prepare this report was
taken from published sources and unpublished data files archived at the Woods Hole Laboratory in Woods Hole,
Massachusetts (all included in the references to this paper). Since some of this information is based on incomplete data
sets or has not been updated for many years, estimated lengths at age are meant to be used only in a general sense and are
not intended for use in generating growth equations for analytical purposes. The intent is to provide a general guide for
use in estimating the average age of a finfish or shellfish specimen at a given length. It is rarely possible to assign exact
ages to specimens without examining actual age structures (scales, otoliths, chondrophores, etc.), but the tables and graphs
should provide an approximation of the age of any given specimen. For example, a 19.7 in. (50.0 cm) female American
plaice would be estimated as being approximately nine years old.

Each table and corresponding graph gives information on estimated lengths at age in units of both inches and
centimeters. By convention, a birthdate of January 1 is assumed. This means that although an individual is not one-year
old biologically until one year from the time it was hatched, normal age determination convention considers it to be one-
year old the following January 1. This is done to simplify assignment of individual fish to year classes for stock status
assessments and other research. The lengths at age given in the tables represent the estimated lengths on the assigned
birthdate. For example, six-year-old male American plaice would be on average 15.1 in. (38.4 cm) long in January.

Table 1. Maximum ages observed for various finfish species collected during Northeast Fisheries Center research vessel
survey cruises, 1963 - 1988

Species Years Maximum age
Alewife 1973-1985 14
American plaice 1976-1984 24
Atlantic cod 1970-1988 18
Atlantic herring 1963-1988 18
Atlantic mackerel 1973-1987 19
Blueback herring 1982-1985 12
Bluefish 1985-1987 10
Butterfish 1977-1988 6
Haddock 1963-1988. 14
Pollock 1970-1988 19
Redfish 1975-1980 58
Red hake 1970-1985 14
Scup 1977-1980 1
Silver hake 1973-1987 14
Summer flounder 1975-1988 12
White hake 1987 only 18
Winter flounder 1976-1988 14
Witch flounder 1980-1988 30

Yellowtail flounder 1963-1988 17



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PAIRED TABLES AND FIGURES:
ESTIMATED LENGTH (SHELL HEIGHT) AT AGE


Page 2

REFERENCES

Almeida, F.P. Pers. comm. (red hake length at age). Nat.
Mar. Fish. Serv., Woods Hole, MA 02543.

Almeida, F.P. Pers. comm. (silver hake length at age). Nat.
Mar. Fish. Serv., Woods Hole, MA 02543.

Anderson, E.D., and A.J. Paciorkowski. 1980. A review of
the Northwest Atlantic mackerel fishery. Rapp. P.-V.
Reun. Cons. Int. Explor. Mer. 177: 175-211.

Anthony, V.C. 1972. Population dynamics of the Atlantic
herring in the Gulf of Maine. Ph.D. thesis, University
of Washington, Seattle, WA. 246 p.

Burnett, J. 1987. The population biology of the witch
flounder, Glyptocephalus cynoglossus (L.), in the
Gulf of Maine - Georges Bank region. M.Sc. thesis,
University of Massachusetts, Amherst, MA. 116 p.

Burnett, J.M. Pers. comm. (bluefish length at age). Nat.
Mar. Fish. Serv., Woods Hole, MA 02543.

Campbell, MJ., J.A. Penttila, and F.E. Nichy. Unpubl. ms.
Growth of scup. Nat. Mar. Fish. Serv., Woods Hole,
MA 02543.

Clark, S.H., W.J. Overholtz, and R.C. Hennemuth. 1982.
Review and assessment of the Georges Bank and Gulf
of Maine haddock fishery. J. Northwest Atl. Fish. Sci.
3: 1-27.

Fogarty, M.J. 1981. Review and assessment of the summer
flounder (Paralichthys dentatus) fishery in the North-
west Atlantic. Nat. Mar. Fish. Serv., Northeast Fish.
Ctr., Woods Hole Lab. Ref. Doc. No. 81-25. 54 p.

Lux, F.E. 1969. Age and growth of the winter flounder,
Pseudopleuronectes americanus, on Georges Bank.
Fish. Bull., U.S. 71: 505-512.

Mayo, R.K. 1980. Exploitation of redfish, Sebastes
marinus (L.) in the Gulf of Maine - Georges Bank
region, with particular reference to the 1971 year class.
J. Northwest Atl. Fish. Sci, 1: 21-37.

Mayo, R.K. Pers. comm. (pollock length at age). Nat. Mar.
Fish. Serv., Woods Hole, MA 02543.

Moseley, S.E. 1986. Age structure, growth, and intraspeci-
fic growth variations of yellowtail flounder, Limanda
ferruginea (Storer), on four northeastern United States
fishing grounds. M.Sc. thesis, University of Massa-
chusetts, Amherst, MA. 110 p.

Penttila, J.A., and V.M. Gifford. 1976. Growth and
mortality rates for cod from the Georges Bank and
Gulf of Maine areas. Int. Comm. Northwest Atl. Fish.
Res, Bull. 12; 29-36.

Serchuk, F.M., and S.A. Murawski. 1980. Assessment and
status of surf clam Spisula solidissima (Dillwyn) popu-
lations in offshore Middle Atlantic waters of the United
States. Nat. Mar. Fish. Serv., Northeast Fish. Ctr.,
Woods Hole Lab. Ref. Doc. No. 80-33. 46 p.

Serchuk, F.M. Pers. comm. (sea scallop length at age).
Nat. Mar. Fish. Serv., Woods Hole, MA 02543.

Sullivan, L.F. 1982. American plaice, Hippoglossoides
platessoides, in the Gulf of Maine: I. The fishery II.
Age and growth III. Spawning and larval distribution.
MSc. thesis, University of Rhode Island, Kingston,
RI. 95 p.

Waring, G.T. Pers. comm. (butterfish length at age). Nat.
Mar. Fish. Serv., Woods Hole, MA 02543.


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Guidelines for Estimating Lengths at Age
for 18 Northwest Atlantic Finfish

and Shellfish Species

U.S. DEPARTMENT OF COMMERCE
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National Marine Fisheries Service
Northeast Fisheries Center
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May 1989


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Table of Contents

Introduction...
References

ne

Table 1. Maximum ages observed for various finfish species collected during Northeast Fisheries Center

research vessel survey cruises, 1963-1988 ... ~1
Paired Tables and Figures: Estimated length (shell height) at age
American plaice ...... 24
Atlantic cod... 6
Atlantic herring ...... . 8

Atlantic mackerel ....

Bluefish

Butterfish

Haddock

Pollock .....

18

Redfish

Red hake

Scup ....

Sea scallop .....

Silver hake ..

Summer flounder......

Surf clam

- 32

Winter flounder

Witch flounder

Yellowtail flounder ..

Page iii


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NOAA Technical Memorandum NMFS-F/NEC-66

This TM series is used for documentation and timely communication of preliminary results,
interim reports, or special purpose information, and has not received complete formal review,
editorial control, or detailed editing.

Guidelines for Estimating Lengths at Age
for 18 Northwest Atlantic Finfish
and Shellfish Species

Judith A. Penttila, Gary A. Nelson, and John M. Burnett, Ill

Woods Hole Lab., National Marine Fisheries Service, Woods Hole, MA 02543

U. S. DEPARTMENT OF COMMERCE
Robert A. Mosbacher, Secretary

National Oceanic and Atmospheric Administration
William E. Evans, Administrator

National Marine Fisheries Service
James W. Brennan, Assistant Administrator for Fisheries

Northeast Fisheries Center
Woods Hole, Massachusetts

May 1989


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Recent issues in this series

36. Growth and Survival of Larval Fishes in Relation to the Trophodynamics of Georges Bank Cod and
Haddock. By Geoffrey C. Laurence and R. Gregory Lough. January 1985. xvi + 150 p., 67 figs., 15 tables, 1 app. NTIS
Access. No. PB85-220093/AS.

37. Regional Action Plan. By Bruce E. Higgins, Ruth Rehfus, John B. Pearce, Robert J. Pawlowski, Robert L.
Lippson, Timothy Goodger, Susan Mello Roe, and Douglas W. Beach. April 1985. ix + 84 p., 4 figs., 6 tables, 9 app.
NTIS Access. No. PB85-219962/AS.

38. The Shelf/Slope Front South of Nantucket Shoals and Georges Bank as Delineated by Satellite Infrared
Imagery and Shipboard Hydrographic and Plankton Observations. By J. B. Colton, Jr., J. L. Anderson, J. E.
O’Reilly, C. A. Evans-Zetlin, and H. G. Marshall. May 1985. vi + 22 p., 14 figs. NTIS Access. No. PB85-221083/AS.

39. USA Historical Catch Data, 1904-82, for Major Georges Bank Fisheries. By Anne M. T. Lange and Joan E.
Palmer. May 1985. iii + 21 p., 12 figs., 2 tables. NTIS Access. No. PB85-233948/AS.

40. Indexing the Economic Health of the U.S. Fishing Industry’s Harvesting Sector. By Virgil J. Norton, Morton
M. Miller, and Elizabeth Kenney. May 1985. v + 42 p., 44 figs., 25 tables, 1 app. NTIS Access. No. PB85-217958/
AS.

41. Calculation of Standing Stocks and Energetic Requirements of the Cetaceans of the Northeast United States
Outer Continental Shelf. By Robert D. Kenney, Martin A. M. Hyman, and Howard E. Winn. May 1985. iv + 99 p.,
1 fig., 5 tables, 1 app. NTIS Access. No. PB85-239937/AS.

42. Status of the Fishery Resources Off the Northeastern United States for 1985. By Conservation & Utilization
Division, Northeast Fisheries Center. August 1985. iii + 137 p., 46 figs., 49 tables. NTIS Access. No. PB86-125473/
AS.

43. Status of the Fishery Resources Off the Northeastern United States for 1986. By Conservation & Utilization
Division, Northeast Fisheries Center, September 1986. iii + 130 p., 45 figs., 48 tables. NTIS Access. No. PB87-122115/
AS.

44. NOAA’s Northeast Monitoring Program (NEMP): A Report on Progress of the First Five Years (1979-84)
and a Plan for the Future. By Robert N. Reid, Merton C. Ingham, and John B. Pearce, eds., and Catherine E. Warsh
(water quality), Robert N. Reid (sediments & bottom organisms), Adriana Y. Cantillo (trace contaminants in tissues),
and Edith Gould (biological effects), topic coords. May 1987. xi + 138 p., 13 figs., 1 table, 9 app. NTIS Access. No.
PB87-210100.

45. Food and Distribution of Juveniles of Seventeen Northwest Atlantic Fish Species, 1973-1976. By Ray E.
Bowman, Thomas R. Azarovitz, Esther S. Howard, and Brian P. Hayden. May 1987. xi + 57 p., 10 figs., 19 tables. NTIS
Access. No. PB87-215851/AS.

46. Influence of Freshwater Inflows on Estuarine Productivity. By James G. Turek, Timothy E. Goodger, Thomas
E. Bigford, and John S. Nichols. May 1987. iii + 26 p. NTIS Access. No. PB87-213666/AS.

47. MARMAP Surveys of the Continental Shelf from Cape Hatteras, North Carolina, to Cape Sable, Nova Scotia
(1977-1984). Atlas No. 2. Annual Distribution Patterns of Fish Larvae. By Wallace W. Morse, Michael P. Fahay,
and Wallace G. Smith. May 1987. viii + 215 p., 27 figs., 2 tables. NTIS Access. No. PB87-232831/AS.

48. Indexed Bibliography of the Bay Scallop (Argopecten irradians). By Barbara D. Sabo (Gibson) and Edwin W.
Rhodes. May 1987. iii + 85 p. NTIS Access. No. PB87-231411/AS.

49. Northeast Fisheries Center Framework for Inshore Research. By Research Planning & Coordination Staff,
Northeast Fisheries Center. July 1987. vi + 44 p., 2 figs., 2 tables. NTIS Access. No. PB87-232286/AS.

(continued on inside back cover)


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