NOAA Technical Memorandum NMFS-AFSC-140


Characteristics of Benthic Sediments 

From Areas Open and Closed to 

Bottom Trawling in the Gulf of Alaska 

by 

R. P. Stone and M. M. Masuda

U.S. DEPARTMENT OF COMMERCE

National Oceanic and Atmospheric Administration 

National Marine Fisheries Service 

Alaska Fisheries Science Center 

October 2003 
NOAA Technical Memorandum NMFS 

The National Marine Fisheries Service's Alaska Fisheries Science Center 

uses the NOAA Technical Memorandum series to issue informal scientific and 

technical publications when complete formal review and editorial processing 

are not appropriate or feasible. Documents within this series reflect sound 

professional work and may be referenced in the formal scientific and technical 

literature. 

The NMFS-AFSC Technical Memorandum series of the Alaska Fisheries 

Science Center continues the NMFS-F/NWC series established in 1970 by the 

Northwest Fisheries Center. The NMFS-NWFSC series is currently used by 

the Northwest Fisheries Science Center. 

This document should be cited as follows: 

Stone, R. P., and M. M. Masuda. 2003. Characteristics of benthic 

sediments from areas open and closed to bottom trawling in the Gulf of 

Alaska. U.S. Dep. Commer., NOAA Tech. Memo. NMFS-AFSC-140, 

40 p. 

The appendices should be cited as follows: 

Stone, R. P., and A. Baldwin. 2003. Appendices to Characteristics of 

benthic sediments from areas open and closed to bottom trawling in the 

Gulf of Alaska. U.S. Dep. Commer., NOAA Tech. Memo. NMFS-AFSC140, 111 p. 

Reference in this document to trade names does not imply endorsement by 

the National Marine Fisheries Service, NOAA. 
NOAA Technical Memorandum NMFS-AFSC-140


Characteristics of Benthic Sediments 

From Areas Open and Closed to 

Bottom Trawling in the Gulf of Alaska 

by 

R. P. Stone and M. M. Masuda

Appendices 

by R. P. Stone and A. Baldwin* 

Auke Bay Laboratory 

Alaska Fisheries Science Center 

11305 Glacier Highway 

Juneau AK 99801-8626 

www.afsc.noaa.gov 

*Juneau Center School of Fisheries and Ocean Science 

University of Alaska Fairbanks 

11120 Glacier Highway 

Juneau AK 99801 

U.S. DEPARTMENT OF COMMERCE

Donald L. Evans, Secretary 

National Oceanic and Atmospheric Administration 

Vice Admiral Conrad C. Lautenbacher, Jr., U.S. Navy (ret.), Under Secretary and Administrator 

National Marine Fisheries Service 

William T. Hogarth, Assistant Administrator for Fisheries 

October 2003 
This document is available to the public through: 

National Technical Information Service 

U.S. Department of Commerce

5285 Port Royal Road 

Springfield, VA 22161 

www.ntis.gov 
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copy format. 
-iiiABSTRACT

 This report documents the sedimentary, biogeochemical, and infaunal characteristics of 

sediments from areas that were open or closed to bottom trawling in the central Gulf of Alaska. 

Sediment samples were collected from three sites during two cruises in 1998 and 1999. Bottom 

trawling had been prohibited in the closed areas for 11-12 years. Trawling intensity in the open 

areas was estimated at 11%-29% of the seafloor per year for the 5 years preceding this study. We 

found evidence that the sedimentary and biogeochemical characteristics of the seafloor have been 

altered to some degree by ambient levels of bottom trawling. Total organic carbon was higher in 

open areas at Site 1-1998 and at Site 2. Median grain size differed between open and closed 

areas at Sites 2 and 3, as did sediment sorting at Site 3. Grain size analysis at Site 3 provided 

evidence of a natural habitat gradient that may have been present prior to the closure in 1987. No 

significant differences in infauna measures of community structure were found between open and 

closed areas, other than a higher Simpson’s index of dominance in the open area at Site 2. 

Differences in sediment characteristics were subtle and minor, and no clear, repeated pattern of 

effects occurred among the three sites. Our results indicate that the effect of mobile fishing gears 

(i.e., otter trawls and scallop dredges) on sediment properties and infaunal communities at the 

three sites examined and at present levels of fishing is minor and not cause for serious concern at 

the present time. These results, used in conjunction with those on the effects of trawling on the 

epifaunal component of these communities, will provide resource managers with some insight 

into how current trawling activities in the Gulf of Alaska are affecting this important habitat. 

Infauna data appendices are included. 
-iv­
-vCONTENTS 

ABSTRACT ................................................................. iii 


INTRODUCTION ............................................................. 1

STUDYAREAS ............................................................... 2

METHODS .................................................................. 4

Estimation of Trawl Intensity .................................................. 4

 Experimental Design ......................................................... 5

 Statistical Methods .......................................................... 7

RESULTS .................................................................. 10 


 Sediment Samples .......................................................... 10 


 Sediment Characteristics ..................................................... 11 


 Differences in Sediment Composition ........................................... 11 


 Infaunal Community Structure ................................................. 13 


 Multivariate Analyses ....................................................... 14 


DISCUSSION ............................................................... 15 


ACKNOWLEDGMENTS ...................................................... 20 


CITATIONS ................................................................. 21 


TABLES .................................................................... 26 


FIGURES ................................................................... 31 

INTRODUCTION

 The Magnuson-Stevens Fishery Conservation and Management Act, as amended by the 

Sustainable Fisheries Act of 1996, requires the regional Fishery Management Councils and the 

National Marine Fisheries Service (NMFS) to minimize, to the extent practicable, adverse effects 

from fishing on essential fish habitat (U.S. Department of Commerce 1996). In 1998 and 1999, 

the Alaska Fisheries Science Center’s Auke Bay Laboratory initiated a study to determine the 

effects of bottom trawling on soft-bottom, low-relief marine habitat on the inner continental shelf 

of the central Gulf of Alaska (GOA).

 Previous studies worldwide have determined that bottom trawling alters seafloor habitat and 

has direct and indirect effects on benthic communities (Jones 1992, Auster et al. 1996). In 

addition to removing target species, bottom trawling incidentally removes, displaces, or damages 

non-target species (Ball et al. 2000) and physically alters the sedimentary, biogeochemical, and 

biological features of the seafloor (Churchill 1989). The chronic effects of these disturbances are 

unclear, due to a lack of potential reference sites where bottom trawling has been prohibited for 

any significant period. The degree of alteration likely depends on many factors, some of which 

may be area-specific; therefore, generalizing the effects of trawling over wide geographical areas 

is not prudent. Potential factors include 1) gear type, 2) spatial and temporal intensity of 

trawling, 3) substrate characteristics, and 4) oceanographic features near the seafloor (Jones 

1992).

 In April 1987 the North Pacific Fishery Management Council closed two areas near Kodiak 

Island, Alaska, to bottom trawling (Type 1 Areas). Scallop dredges are also prohibited in Type 1 

Areas. These areas were designated as important rearing habitat and migratory corridors for 
2


juvenile and molting crabs. The closures are intended to assist in rebuilding severely depressed 

Tanner crab (Chionoecetes bairdi) and red king crab (Paralithodes camtschaticus) stocks. In 

addition to crab resources, the closed areas and areas immediately adjacent to them support rich 

stocks of groundfish including flathead sole (Hippoglossoides elassodon), butter sole 

(Pleuronectes isolepis), Dover sole (Microstomus pacificus), rex sole (Glyptocephalus zachirus), 

Pacific halibut (Hippoglossus stenolepis), arrowtooth flounder (Atheresthes stomias), Pacific cod 

(Gadus macrocephalus), walleye pollock (Theragra chalcogramma), and several species of 

rockfish (Sebastes spp.) (Martin and Clausen 1995). Consequently, bottom trawling occurs 

immediately adjacent to the closed areas year-round. These closures provided a rare opportunity 

to study the effects of an active bottom trawl fishery on a productive, deepwater (> 100 m), softbottom marine ecosystem. Direct comparisons were possible between areas that were 

consistently trawled each year and areas where bottom trawling had been prohibited for 11 to 12 

years. The proximity of the closed and open sites allowed for comparison of fine-scale infauna 

and epifauna diversity and abundance, and microhabitat and community structure. This report 

documents the sedimentary, biogeochemical, and infaunal characteristics of sediments from areas 

open and closed to bottom trawling at three locations in the central GOA. 

STUDY AREAS

 Three study sites were established within and adjacent to Type 1 Area closures (Fig. 1). Study 

sites were chosen using two criteria: 1) the seafloor consisted of a soft-bottom substrate (i.e., 

sand, silt, and clay) and was relatively uniform in depth, and 2) trawling had occurred 

immediately adjacent to the closed area during the preceding 5 years. The first criterion was 
3


considered necessary to reduce variation in habitat and community structure associated with 

change in depth.

 Site 1 was located in an area known as the Chiniak Gully near the northeast side of Kodiak 

Island (Fig. 1). We investigated this site during June 1998 and August 1999 and established 10 

seafloor transects for each year. Maximum trawl intensity at Site 1 was estimated to be 29.4% of 

the seafloor per year. This site is characterized by strong, bi-directional bottom currents that flow 

predominately from the northwest and southeast. Depth within the transect area ranged from 105 

to 151 m, and the maximum depth differential along any transect was 18 m. 

 Site 2 was located in an area known as the Two-headed Gully near the southeast corner of 

Kodiak Island (Fig. 1). Ten seafloor transects were established at this site in June 1998. 

Maximum trawl intensity at Site 2 was estimated to be 19.4% of the seafloor per year. This site 

is characterized by moderate to light bottom currents. Depth within the transect area ranged from 

125 to 157 m, and the maximum depth differential along any transect was 15 m.

 Site 3 was located approximately 22 km south of the Trinity Islands (Fig. 1). Eleven seafloor 

transects were established at this site in August 1999, and maximum trawl intensity was 

estimated to be 11.4% of the seafloor per year. This site is characterized by light bottom 

currents. Depth within the transect area ranged from 124 to 137 m, and the maximum depth 

differential along any transect was 11 m. 
4 

METHODS 

Estimation of Trawl Intensity

 Using retrospective analysis of past trawling activity by the domestic commercial fishing fleet, 

we identified three areas where moderate bottom trawling had occurred immediately adjacent to 

Type 1 Area closures (Fig. 1). Trawl frequency was documented at each site for the 5-year 

period prior to our study through the NMFS North Pacific Groundfish Observer Program 

(NORPAC database). All observed hauls that originated within approximately 3 km of any point 

in the transect area were recorded (Fig. 2). Actual distances may have deviated slightly because 

the latitude and longitude of the starting position for each haul are rounded to the nearest nautical 

mile (nmi). Number of hauls by the fishing fleet (vessels between 18.3 m and 38 m in length) 

within each study site was estimated by dividing the observed number of hauls by 30% 

(percentage of fleet with observers). The number of hauls made by fishing vessels greater than 

38 m (125 feet) in length were estimated based on 100% observer coverage. All hauls are 

accounted for in the NORPAC database. Vessels smaller than 18.3 m (60 feet) in length are not 

required to participate in the observer program and therefore our estimates do not account for 

trawling by these vessels.

 We calculated the area of the seafloor swept by the trawl footrope (wing spread) based on a 

width of 17 m. Our calculations do not take into account any portion of the seafloor contacted by 

the trawl doors (otter boards), sweeps, or bridles. The length of each observed haul was 

calculated by multiplying the haul duration by an average vessel speed of 7.4 km/h (4 nmi/h). 

The area of each observed haul was calculated as the product of trawl length and trawl width. 
5


The total area of seafloor swept by footropes within each study area was calculated as the product 

of the estimated number of hauls and the mean area trawled per haul. Mean area trawled per haul 

was calculated independently for vessels between 18.3 m and 38 m in length and for those 

greater than 38 m in length. Values for total area trawled per site were calculated with the 

assumption that all hauls were nonoverlapping and were made entirely within the study area. 

Experimental Design

 At each site, we established parallel transects 3,000 m long and approximately 500 m apart. 

Each transect was bisected by the boundary closure line into equal 1,500 m sections in both the 

open and closed areas (Fig. 2). At Site 1, transects were 500 m apart during each year, but 250 m 

apart when all transects (both years) were combined (Fig. 2). 

 Bottom sediment to a depth of 10 cm was collected with a 0.04 m2 Shipek sediment sampler 

(volume 0.003 m3) along transects. The volume of each sediment sample was estimated from the 

measured height of sediment in the sampler. The design for sediment sampling differed during 

the 2 years of this study for Site 1 as illustrated in Figure 2. In 1998, three samples were 

collected at 250 m, 1,500 m, and 2,750 m along each transect (Fig. 2). In 1999, sediment was 

collected at six stations per transect following a systematic design. Stations were 500 m 

equidistant from a randomly generated starting point within the first 500 m (open end) of each 

transect, yielding three samples from each side of the trawl closure boundary (Fig. 2). Starting 

points were independently generated for each transect.

 Approximately 100 g was collected from the top 3 cm of each sediment sample to determine 

grain size, and 1 g from the top 1 cm for total organic carbon content (TOC) analysis. Infauna 
6


were macroscopically removed from those two subsamples. The remaining sediment was gently 

washed through a 500-μm sieve, and retained material was composited with the subsamples and 

preserved in 4% seawater-buffered formaldehyde containing 0.2 g/L Rose Bengal. In the 

laboratory, samples were transferred to a 70% ethanol solution, and organisms were enumerated 

and identified to the lowest possible taxonomic level. Whole animals and identifiable fragments 

were assigned an abundance value of 1. Annelids were further classified as sedentary or errant 

since these two forms may differ in their response to disturbance by fishing activities (Brylinsky 

et al. 1994). Organisms captured in the grab that are considered primarily epibenthic were not 

included in infauna analyses. These included the dwarf wrymouth (Cryptacanthodes aleutensis), 

juvenile arrowtooth flounders (Atheresthes stomias), an amphipod (Maera danae), three species 

of lysianassid amphipods (Anonnyx lilljeborgi, Lepidepecreum guarjanovae, and Wecomedon 

sp.), a crangonid shrimp species, Tanner crabs (Chionoecetes bairdi), Pacific lyre crabs (Hyas 

lyratus), hermit crabs (Pagurus ochotensis and Pagurus tanneri), a brachiopod (Terebratulina 

unguilica), an urchin (Strongylocentrotus sp.), sea stars (Luidia foliolata and Ctenodiscus 

crispatus), the pale moon snail (Euspira pallidus), and sea whips (Protoptilum sp.).

 Sediments were analyzed for grain size composition by wet sieving and were graded using the 

Wentworth classification scale (Buchanan 1984). Sorting was calculated with the Inclusive 

Graphic Standard Deviation method (Buchanan 1984), with higher values indicating greater 

grain-size variability. The TOC was analyzed with a Dohrmann Carbon Analyzer using a twopoint calibration curve. Twenty percent of the carbon samples were analyzed in triplicate to 

estimate analytical precision. The coefficient of variation ranged between 0.21% and 5.40% for 

these samples. 
7 

Statistical Methods

 Open areas were compared with closed areas using both univariate and multivariate methods. 

Samples from the three sites were analyzed separately, as sites were selected in part due to site 

contrasts. First- and second-year samples from Site 1 were also analyzed separately because 

fewer samples were collected during the first year. Differences in TOC, median grain size, and 

sediment sorting were tested with analysis of variance (ANOVA) models. We assessed 

community structure by analyzing the two components of species diversity: species richness and 

relative abundance of species (Magurran 1988). We used Margalef=s index, a function of total 

numbers of species and individuals, to measure species richness and Simpson’s index of 

dominance to measure the relative abundance of species (Clarke and Warwick 1994). Margurran 

(1988) notes that species richness and dominance indices are informative in environmental 

studies and cites several studies that demonstrated reduced species richness and increased 

dominance in stressed communities. Differences between open and closed areas in terms of 

species diversity and animal abundance, including total number of sedentary annelids and total 

number of errant annelids, were also tested with individual ANOVA models. The two-factor 

models included fixed factor status (variable indicating open or closed area) and fixed factor 

transect (blocking variable). In 1998, factor status had three levels (open, closed, and boundary) 

because a sample was collected on the boundary of the open and closed areas. Transects were 
8


assumed independent, and the two-factor ANOVA model used for the data was

 (1) 

yij = μ + α + β + ε , .. i j ij 

where μ is the general mean, α is the effect at the i-th level of status, β is the effect of the j- .. i j

the transect, and ε ij are independent N(0,σ2). 

 In 1998, one sample was collected for each level of status crossed with transect; that is, no 

replicate samples were collected at both sites. In 1999, three samples were systematically 

collected on either side of the boundary at fixed distances along a transect at both sites. Because 

samples may be autocorrelated along a transect, observations for these pseudo-replicate samples 

were averaged for testing area closure differences. In this case, yij in the model (Eq. 1) 

represented means and the model was fit with weighted least squares, with weights equaling the 

number of samples averaged. Prior to computation of indices with the software PRIMER (Clarke 

and Warwick 1994), numbers of individuals per species were standardized to the volume of one 

full grab (0.003 m3

). Dependent variables were Box-Cox transformed (Box and Cox 1964; 

Venables and Ripley 1999), if necessary, with a different power parameter λ estimated in the 

range [-2, 2] for each variable. One-tailed t-tests were used to test for reduced species richness 

(Margalef’s index), increased dominance (Simpson’s index), and decreased numbers of sedentary 

annelids in the open area. Concurring with the arguments of McConnaughey et al. (2000), we 

chose an "-level of 0.10 over the traditional "-level of 0.05 to protect against a Type II error.

 Sediment grabs less than 50% full were excluded from infauna analyses (see RESULTS ­

Sediment Samples), resulting in missing data for 1998 (Sites 1 and 2) when grab samples were 

not replicated. We imputed missing data multiple times (Schafer 1997; Schimert et al. 2001) in 
9


the analyses of species diversity (variables richness and dominance) and grouped taxa (all 

individuals, and sedentary and errant annelids). We assumed a Gaussian model for the squareroot of the data. For the Site-2 data, we used a noninformative prior (Schafer 1997; Schimert 

et al. 2001) for parameters of the Gaussian model and a ridge prior for Site-1 data. A data 

augmentation algorithm was used to generate three imputations from three parallel chains started 

from bootstrapped initial values (Schafer 1997; Schimert et al. 2001). We used Rubin’s rule 

(Rubin 1987; Schafer 1997) to combine parameter estimates of the ANOVA models (Schafer 

1997) for statistical inference. 

 Standardized multispecies counts were analyzed with two multivariate methods: nonmetric 

multidimensional scaling (MDS) -- a graphical technique -- and multivariate analysis of variance 

(MANOVA). First, MDS was used to map the k-variate samples into two dimensions, based on 

pairwise similarity coefficients computed between samples. Bray-Curtis similarities (Clarke 

1993), computed on double-root (or 4th root) transformed species counts, were used as measures 

of distance between samples. Multidimensional scaling fits were assessed by “stress” (Clarke 

and Warwick 1994), a measure of how well between-sample relationships are preserved in lower 

dimensions. Second, differences in multispecies and aggregated multispecies counts were tested 

with MANOVA. Rare species, defined as those that never account for greater than or equal to 

p% (where p is some small arbitrary value, see Clarke and Warwick 1994) of the abundance in 

any sample, were excluded from the MANOVA (Field et al. 1982). Species were also 

aggregated at the higher taxonomic level of phylum. These data-reduction techniques ensured 

sufficient degrees of freedom to determine significance of tests, and also eliminated many zeroes 

in the data, improving distributional properties. Because many samples less than 50% full at Site 
10


1 - 1998 and at Site 3 were excluded from analyses, multivariate tests for differences in status 

were not performed (see RESULTS - Sediment Samples).

 The dependent variables, arrays of counts, were transformed if necessary, and a battery of 

normality tests (Aitchison 1986) was performed to verify the model assumption of multivariate 

normality. Independent variables in the MANOVA models were status and transect. For each 

MANOVA, we computed the Wilks’ lambda or the likelihood ratio test statistic (Johnson and 

Wichern 1998). The α-level was 0.10. 

RESULTS 

Sediment Samples

 Sediment samples were collected from 181 of 186 stations during this study. For unknown 

reasons, we were unable to collect samples from five stations at Site 1 during 1999. The 

sediment samples collected were not consistently full grab samples; that is, the volume taken by 

the sampler was at times less than 0.003 m3

. All samples, including partial grab samples, yielded 

valid TOC and grain-size composition data, but we questioned the validity of comparing infauna 

data from samples of unequal volumes. Using linear regression, a positive relationship was 

found between fullness of grab samples (% volume) and both number of species and number of 

individuals; that is, fuller grab samples appeared to have more species and individuals (e.g., Fig. 

3). To ensure that numbers of species and individuals were comparable among all grab samples, 

we standardized counts (individuals and species) to a unit volume of one full grab (0.003 m3

). 

After standardizing species and individual counts, regression analysis showed a negative 
11


relationship between grab fullness and counts (e.g., Fig. 4), indicating that individuals and 

species were not uniformly distributed to the depth sampled by a full grab. Partial grabs probably 

sample only the uppermost few centimeters of the sediment where the majority of the meiofuana 

and microfauna live (Gray 1981). In general, for grabs greater than or equal to 50% full, 

standardized numbers of species and individuals were not related to grab fullness. Grabs less 

than 50% full were therefore excluded from analyses (Table 1). All infauna samples collected at 

Site 1 in 1999 were greater than or equal to 50% full (Table 1). 

Sediment Characteristics

 The TOC, median grain size, and sediment sorting varied between the three sites. Mean TOC 

was highest at Site 2 (7,276 ppm) and lowest at Site 3 (2,718 ppm) (Table 2). Mean TOC was 

similar at Site 1 in 1998 and 1999 and ranged from 2,537 to 5,116 ppm and from 2,941 to 

5,819 ppm, respectively (Table 2). Median grain size ranged from 229 :m (fine sand) at Site 3 

to 86 :m (very fine sand) at Site 2 (Table 3). Sediment grain size was relatively uniform at Sites 

1 and 2, but spanned a wider range of sizes at Site 3 (Table 3). This variability in grain size is 

reflected in high sorting coefficients for Site 3 sediments (Table 4). Site 2 had the best-sorted 

sediments (sorting class = moderately sorted, Gray 1981), whereas Site 3 had the worst-sorted 

sediments (sorting class = poorly sorted, Gray 1981) (Table 4). 

Differences in Sediment Composition

 The TOC was significantly higher in the open area than in the closed area at Site 1 in 1998 

(Fig. 5A) but not in 1999 (Fig. 5B; Table 5). At Site 2, the open area had significantly higher 
12


TOC than the closed area (Fig. 6A; Table 5). Significant differences in TOC were not detected at 

Site 3 (Fig. 6B; Table 5).

 Significant differences in median grain size and sediment sorting were not detected at Site 1 

during either year (Table 5). At Site 2, the closed area had significantly larger median grain size 

than the open area (Fig. 7A; Table 5). No significant difference in sediment sorting was detected 

at Site 2 (Table 5). At Site 3, similar to Site 2, the closed area had significantly larger median 

grain size than the open area (Fig.7B; Table 5). Sediments in the closed area were also 

significantly more poorly sorted, as indicated by higher sorting values, than those from the open 

area (Fig. 8; Table 5).

 A prominent demarcation in sediment type was observed from the research submersible Delta 

near the boundary between the open and closed areas at Site 3. This observation was consistent 

with the differences in sediment grain size measured there. A closer examination of these data 

revealed an ordering in grain size from the south end (A) to the north end (F) of the transects 

(Fig. 9). An ordering of samples of ABC, in 6 of 11 transects, and also an ordering of DEF, in 6 

of 11 transects, indicate the existence of a gradient (Fig. 9). If the transects were independent 

and the ordering of samples were random on either side, then we would expect an ordering of 

ABC or DEF in 1 of 6 transects. A randomization test with 10,000 permutations (Edgington 

1987) indicated that the probability of observing 6 of 11 transects at random with the ordering 

ABC or DEF was very low (P = 0.0048), therefore providing strong evidence for the existence of 

a gradient. If the differences in median grain size were due to a natural habitat gradient, then the 

grain size values would be expected to gradually change along the transect from the open end 
13


(sample A) to the closed end (sample F) as observed. Conversely, if the differences were due to 

the trawling closure then one would expect no ordering of the samples on the open side. 

Infaunal Community Structure

 One hundred and sixty-four unique taxonomic groups of infauna, representing 11 phyla and 

102 families, were identified from the sediment samples collected during this study (see 

Appendices volume). We identified 57% of the taxa to the species level and 88% to the genus 

level (see Appendices volume). Collectively, the annelids, both errant and sedentary forms, were 

the most numerous and diverse phylum at all sites. Amphipods and cumaceans were the most 

common arthropods, while the Amphiuridae were the dominant family of echinoderms at all 

sites. Several species of bivalves, most notably Axinopsida serricata and several species of 

Macoma, were common and abundant at all sites. 

 Significant differences in infauna abundance and species diversity between open and closed 

areas were not detected at Site 1 in either year. Site 2 exhibited a significantly higher index of 

dominance in the open area than in the closed area and no significant difference in species 

richness or animal abundance. Site 3 was not examined for differences in infauna between 

closed and open areas because several grab samples less than 50% full were excluded from 

infauna analyses (see RESULTS - Sediment Samples). There were no significant differences in 

numbers of sedentary annelids between open and closed areas at Site 1 in either year or at Site 2 

(Table 5). There were also no significant differences in total number of errant annelids between 

open and closed areas at Site 1 in either year or at Site 2 (Table 5). 
14 

Multivariate Analyses

 The nonmetric MDS plots indicated no discernible groupings of individual infauna samples by 

status at any of the three sites. Infauna samples that were averaged for the open or closed area by 

transect, however, appeared to group by transect number (left to right) (e.g., Fig. 10). Stress was 

high in all cases (0.18 at Site 1-1998, 0.21 at Site 2, 0.14 at Site 1-1999, and 0.18 at Site 3), 

indicating that the preservation of between-sample relationships in two dimensions was generally 

poor.

 At Site 1-1999, the multivariate tests showed no significant differences between open and 

closed areas (9-species MANOVA, Wilks’ lambda = 0.11, approximate F = 0.88 ~ F9,1, P = 0.69; 

7-phlya MANOVA, Wilks’ lambda = 0.11, approximate F = 0.88 ~ F9,1, P = 0.69). The 88 

species at Site 1-1999 were reduced to 9 (Mediomastus californiensis, Lumbrineris luti, 

Leitoscoloplos pugettensis, Chaetozone sp., Nephtys caeca, Diastylopsis dawsoni, Amphiodia 

sp., Axinopsida serricata, and Nutricola lordi) by retaining only those species that accounted for 

greater than or equal to 9% of the abundance in any one sample. Abundances for the 88 species 

were pooled into 7 phyla: Annelida, Arthropoda, Chordata, Echinodermata, Echiura, Mollusca, 

and Nemertea. 

 At Site 2, the multivariate tests showed no significant differences between open and closed 

areas (11-species MANOVA, Wilks’ lambda = 0.11, approximate F = 0.56 ~ F22,6, P = 0.85; 4­

phyla MANOVA, Wilks’ lambda = 0.68, approximate F = 0.53 ~ F8,20, P = 0.82). The 49 species 

at Site 2 were reduced to 12 (Magelona berkeleyi, Leitoscoloplos pugettensis, Chaetozone sp., 

Levinsenia gracilis, Hemipodus borealis, Nephtys caeca, Prionospio steenstrupi, Terebellides 
15


stroemi, Byblis veleronis, Nuculana minuta, Macoma eliminata, and Axinopsida serricata) by 

retaining only those species that accounted for greater than or equal to 16% of the abundance in 

any one sample. Byblis veleronis was excluded from the analysis because this species was 

observed in only one infauna sample. Abundances for the 49 species were pooled into 6 phyla: 

Annelida, Arthropoda, Echinodermata, Echiura, Mollusca, and Nemertea. Phyla Echinodermata 

and Echiura were excluded from the analysis because only two animals in each phyla were 

observed in all of the infauna samples. 

DISCUSSION

 We investigated the chronic effects of ambient levels of year-round bottom trawling (otter 

trawls and to a lesser extent scallop dredges at the open area at Site 1) on the benthos at three 

deepwater (> 100 m depth) locations in the central GOA. The sites investigated were separated 

by approximately 200 km, and although universally categorized as soft-bottom habitat, differ 

with respect to the levels of ambient trawling intensity (11% to 29%), sediment grain-size 

characteristics, total organic carbon content, depth range, and current regime. So our findings 

apply to a range of soft-bottom habitats in the central GOA.

 We found evidence that the biogeochemical and sedimentary characteristics of the seafloor in 

areas open to bottom trawling and scallop dredging have been altered to some degree. Total 

organic carbon was higher in areas open to benthic trawling at Site 1-1998 and at Site 2. We also 

detected differences in median grain size at Site 2 and differences in grain size and sorting at Site 

3. Differences at Site 3 may be associated with a natural habitat gradient that may have been 

present when trawling was prohibited 12 years earlier. In general, however, differences in 
16


sediment characteristics were subtle and minor, and no clear, repeated pattern of effects occurred 

among the three sites. Because sediments at Sites 1 and 2 were well sorted, we did not expect to 

find broad-scale changes in grain-size characteristics. These findings support the concept that 

surficial sediments are physically mixed by gear in contact with the seafloor. This is particularly 

evident from our analyses of TOC, where carbon within the sediment is likely transported to the 

surface during mixing.

 Infauna serve many important ecological functions in marine systems, including sediment 

redistribution (Hall 1994), and are also an important food item for some species of flatfish in our 

study areas (R. Stone, unpublished data). We found little evidence that changes have occurred to 

the diversity and abundance of infauna at the three sites investigated. We found no differences in 

measures of infauna community structure between open and closed areas, except for significantly 

higher Simpson’s index of dominance in the open area at Site 2. These findings were unexpected 

given the number of studies showing that trawling in soft-sediment communities affects the 

biomass and community structure of infauna (reviewed in Johnson 2002). Our results may 

indicate that resident infauna are indeed unaffected by ambient levels of trawling or, 

alternatively, that our sample sizes were not adequate to detect changes that have occurred. 

Drabsch et al. (2001) suggest that high spatial variability in infaunal assemblages can exceed 

changes induced by trawling, even with the most rigorous sampling designs. Infaunal 

assemblages at our study sites were highly variable even across small (< 500 m) spatial scales. 

We did not have any measure of natural variability a priori, and were consequently unable to 

conduct power analyses prior to collecting samples. Our sampling regime was constrained by 

ship time and the high cost of sample collection but was quite rigorous compared to other 
17


studies. We suspect that these are common limitations of most infauna studies, especially those 

in deepwater habitats.

 Future studies designed to show stress-induced change to infaunal communities should focus 

on the analysis of indicator species such as polychaetes (Pearson et al. 1983). Polychaetes differ 

in their response to disturbance such as bottom trawling depending on their life mode; errant 

species are typically less sensitive to disturbance than sedentary species (Frid et al. 1999). Errant 

species with small, robust bodies and short life spans adapt well to disturbance (Ball et al. 2000), 

while tubicolous species are typically sensitive to physical disturbance. Despite the fact that both 

forms were common at our study sites (see Appendices volume), we detected no difference in 

their abundance between areas open and closed to bottom trawling.

 Although our study sites are some of the more heavily trawled areas in the central GOA (Rose 

and Jorgensen, in press), trawling intensity is modest compared to other areas worldwide. We 

estimated trawling intensity at our sites at between 11% and 29% of the seafloor per year for the 

5 years preceding this study. The mobile gear fisheries in our three study areas include otter 

trawls that harvest finfish and scallop dredges that harvest weathervane scallops (Patinopecten 

caurinus; Site 1 only, discussed in Masuda and Stone in prep.); together only a fraction of our 

study areas were disturbed by this gear annually. Estimates of bottom trawling intensity on softbottom marine habitat have been similarly estimated at 1,290% per year in the North Sea (Frid et 

al. 1999), 300-400% per year on Nantucket Shoals (Churchill 1989), and 150-300% per year on 

the upper continental margin of northern California (Friedlander et al. 1999).

 Although otter trawls may generally be the most environmentally benign type of benthic 

mobile fishing gear currently used (Hall 1994), many studies have shown that otter trawling on 
18


soft-bottom habitat causes significant changes to the seafloor (reviewed in Johnson 2002). No 

studies have been conducted on the effects of otter trawling on soft-bottom marine habitat in the 

GOA, and few studies have been conducted worldwide in similar habitat (sand-silt bottom) and 

at similar depths (105-157 m); thus, little research exists with which to compare our findings.

 Examination of the few studies comparable to ours provided evidence that both corroborated 

and contradicted our findings. Some evidence exists that areas open to trawling in deepwater 

areas of the Mediterranean Sea had increased TOC (Smith et al. 2000), while other studies have 

noted that experimental trawling reduces the abundance of flocculated organic matter on the 

seafloor (Schwinghamer et al. 1998). Engel and Kvitek (1998) similarly found that lightly 

trawled areas off the California Coast had a higher percent coverage of flocculated material than 

heavily trawled areas. These findings indicate that bottom trawls may affect potential carbon 

sources through two mechanisms: 1) the direct removal or resuspension of accumulated surficial 

organic material and debris from the seafloor, and 2) vertical redistribution of carbon within the 

sediment. 

 We found differences in grain-size characteristics at two study sites where bottom currents 

were weak. Changes in grain-size characteristics are important because grain-size is an 

important correlate for total carbon and nitrogen (Mayer et al. 1988). Engel and Kvitek (1998) 

found no differences in sediment grain-size characteristics between lightly trawled and heavily 

trawled areas. Likewise, a study of the effects of experimental otter trawling on deep water softsediment habitat of the Grand Banks found no effects on grain-size characteristics but did, 

however, note changes to the physical structure of the sediment (Schwinghamer et al. 1998). 
19


 One shortcoming in studying ambient levels of trawling intensity is that estimates of intensity 

and also of the spatial resolution of areas trawled are generally poor. In this study, for example, 

we estimated trawl intensity but were unable to precisely map the location of each trawl path 

because only the start position of each haul is recorded by the fisheries observer. Due to recordkeeping limitations of the fisheries observer program, precise locations of hauls within the open 

areas are unknown (i.e., designated “open” samples may have been collected in areas that were 

not actually trawled). Additionally, two important assumptions of our study were that the closed 

and open areas were identical at the time of the fishery closures in 1987 and that natural 

disturbances have equally affected those areas. 

 Our results indicate that the effect of mobile fishing gears (i.e., otter trawls and scallop 

dredges) on sediment properties and infaunal communities at the three sites examined and at 

present levels of fishing is minor and not cause for serious concern at the present time. These 

results are specific to these study sites, however, and should be cautiously applied to other areas 

of the GOA even when detailed habitat inventories become available. In 2001, we initiated a 

Before-After-Control-Impact (BACI) study at Site 1 using two levels of trawling (1× and 6×) to 

examine the acute effects of experimental bottom trawling on soft-bottom marine habitat. This 

design should provide insights into the potential processes (i.e., resuspension and redistribution 

of sediments) through which mobile fishing gear alters this habitat, and to identify threshold 

levels of trawling intensity that may cause acute changes in sediment characteristics and infaunal 

communities. These results used in conjunction with data on the effects of trawling on the 

epifaunal component of these communities (Stone et al. in press) should provide resource 
20


managers with a comprehensive picture of the effects of current trawling activities on this 

important habitat in the Gulf of Alaska. 

ACKNOWLEDGMENTS

 We thank the following people who helped with this project: John Karinen, Jeff Regelin, 

Patrick Malecha, Ken Krieger, and Linc Freese assisted with field collections; Aaron Baldwin 

and Jeff Regelin provided invaluable laboratory assistance; Chris Lunsford helped with trawling 

intensity estimations; and Jerry Pella, Jon Heifetz, Phil Rigby, Dean Courtney, and Mark 

Zimmermann reviewed this manuscript and provided many helpful suggestions. We are also 

grateful to Captain Wade Loofbourrow and the crew of the Alaska Department of Fish and Game 

RV Medeia for their assistance and support. 
21 

CITATIONS 

Aitchison, J. 1986. The statistical analysis of compositional data. Chapman and Hall Ltd., New 

York. 416 p. 

Auster, P. J., R. J. Malatesta, R. W. Langton, L. Watling, P. C. Valentine, C. L. S. Donaldson, 

E. W. Langton, A. N. Shepard, and I. G. Babb 1996. The impacts of mobile fishing gear 

on seafloor habitats in the Gulf of Maine (Northwest Atlantic): implications for 

conservation of fish populations. Rev. Fish. Sci. 4:185-202. 

Ball, B., B. Munday, and I. Tuck. 2000. Effects of otter trawling on the benthos and 

environment in muddy sediments, p. 69-82. In M. J. Kaiser and S. J. de Groot (editors), 

Effects of fishing on non-target species and habitats: biological, conservation, and socioeconomic issues. Blackwell Scientific Publications, Oxford, England. 

Box, G. E. P., and D. R. Cox. 1964. An analysis of transformations. J. R. Stat. Soc., Series B 

26:211-243. 

Brylinsky, M., J. Gibson, and D. C. Gordon, Jr. 1994. Impacts of flounder trawls on the 

intertidal habitat and community of the Minas Basin, Bay of Fundy. Can. J. Fish. Aquat. 

Sci. 51:650-661. 

Buchanan, J. B. 1984. Sediment Analysis, p. 41-65. In N. A. Holme and A. D. McIntyre 

(editors), Methods for the study of marine benthos, second edition. Blackwell Scientific 

Publications, Oxford, England. 

Churchill, J. H. 	1989. The effect of commercial trawling on sediment resuspension and 

transport over the Middle Atlantic Bight continental shelf. Cont. Shelf Res. 9:841-865. 
22


Clarke, K. R. 1993. Non-parametric multivariate analyses of changes in community structure. 

Aust. J. Ecol. 18:117-143. 

Clarke, K. R., and R. M. Warwick. 1994. Change in marine communities: an approach to 

statistical analysis and interpretation. Natural Environment Research Council, UK. 

144 p. 

Drabsch, S. L., J. E. Tanner, and S. D. Connell. 2001. Limited infaunal response to 

experimental trawling in previously untrawled areas. ICES J. Mar. Sci. 58:1261-1271. 

Edgington, E. S. 1987. Randomization tests. Second Edition. Marcel Dekker, Inc., New York. 

341 p. 

Engel, J., and R. Kvitek. 1998. Effects of otter trawling on a benthic community in Monterey 

Bay National Marine Sanctuary. Conserv. Biol. 12:1204-1214. 

Field, J. G., K. R. Clarke, and R. M. Warwick. 1982. A practical strategy for analyzing 

multispecies distribution patterns. Mar. Ecol. Prog. Ser. 8:37-52. 

Frid, C. L. J., R. A. Clark, and J. A. Hall. 1999. Long-term changes in the benthos on a heavily 

fished ground off the NE coast of England. Mar. Ecol. Prog. Ser. 188:13-20. 

Friedlander, A. M., G. W. Boehlert, M. E. Field, J. E. Mason, J. V. Gardner, and P. Dartnell. 

1999. Sidescan-sonar mapping of benthic trawl marks on the shelf and slope off Eureka, 

California. Fish. Bull. 97:786-801. 

Gray, J. S. 1981. The ecology of marine sediments. Cambridge University Press, Cambridge, 

England. 185 p. 

Hall, S. J. 1994. Physical disturbance and marine benthic communities: life in unconsolidated 

sediments. Oceanogr. Mar. Biol. Annu. Rev. 32:179-239. 
23


Johnson, K. A. 2002. A review of national and international literature on the effects of fishing 

on benthic habitats. U. S. Dep. Commer., NOAA Tech. Memo. NMFS-F/SPO-57, 72 p. 

Johnson, R. A., and D. W. Wichern. 1998. Applied multivariate statistical analysis. PrenticeHall, Upper Saddle River, New Jersey. 816 p. 

Jones, J. B. 1992. Environmental impact of trawling on the seabed: a review. N. Z. J. Mar. 

Freshwater Res. 26:59-67. 

Magurran, A. E. 1988. Ecological diversity and its measurement. Princeton University Press, 

Princeton, New Jersey. 179 p. 

Martin, M. H., and D. M. Clausen. 1995. Data report: 1993 Gulf of Alaska bottom trawl survey. 

U.S. Dep. Commer., NOAA Tech Memo. NMFS-AFSC-59, 217 p. 

Masuda, M., and R. P. Stone. In prep. Biological and spatial characteristics of the weathervane 

scallop Patinopecten caurinus at Chiniak Gully in the central Gulf of Alaska. Available 

from Auke Bay Laboratory, 11305 Glacier Hwy., Juneau. AK. 99801-8626. 

Mayer, L. M., S. A. Macko, and L. Cammen. 1988. Provenance, concentrations and nature of 

sedimentary organic nitrogen in the Gulf of Maine. Mar. Chem. 25:291-304. 

McConnaughey, R. A., K. L. Mier, and C. B. Dew. 2000. An examination of chronic trawling 

effects on soft-bottom benthos of the eastern Bering Sea. ICES J. Mar. Sci. 57:1377-

1388. 

Pearson, T. H., J. S. Gray, and P. J. Johannessen. 	1983. Objective selection of sensitive species 

indicative of pollution-induced change in benthic communities. 2. Data analyses. Mar. 

Ecol. Prog. Ser. 12:237-255. 
24


Rose, C. S., and E. M. Jorgensen. In press. Spatial and temporal distributions of trawling 

intensity off of Alaska: connecting the small scale to issues on an ecosystem scale. 

American Fisheries Society, Proceeding of the Symposium on the Effects of Fishing 

Activities on Benthic Habitats: linking geology, biology, socioeconomics, and 

management. 

Rubin, D. B. 1987. Multiple imputation for nonresponse in surveys. John Wiley and Sons, New 

York. 258 p. 

Schafer, J. L. 1997. Analysis of incomplete multivariate data. Chapman and Hall, London. 

430 p. 

Schimert, J., J. L. Schafer, T. Hesterberg, C. Fraley, and D. B. Clarkson. 2001. Analyzing data 

with missing values in S-PLUS. Insightful Corporation, Seattle, WA. 

Schwinghamer, P., D. C. Gordon, Jr., T. W. Rowell, J. Prena, D. L. McKeown, G. Sonnichsen, 

and J. Y. Guigné. 1998. Effects of experimental trawling on surficial sediment 

properties of a sandy-bottom ecosystem on the Grand Banks of Newfoundland. Conserv. 

Biol. 12:1215-122. 

Smith, C. J., K. N. Papadopoulou, and S. Diliberto. 2000. Impact of otter trawling on an eastern 

Mediterranean commercial trawl fishing ground. ICES J. Mar. Sci. 57:1340-1351. 

Stone, R. P., M. M. Masuda, and P. W. Malecha. In press. Effects of bottom trawling on softsediment communities in the Gulf of Alaska. American Fisheries Society, Proceeding of 

the Symposium on the Effects of Fishing Activities on Benthic Habitats: linking geology, 

biology, socioeconomics, and management. 
25 

U.S. Department of Commerce. 	1996. Magnuson-Stevens Fishery Conservation and 

Management Act. U.S. Dep. Commer., NOAA Tech. Memo. NMFS-F/SPO-23, 121 p. 

Venables, W. N., and B. D. Ripley. 	1999. Modern applied statistics with S-PLUS. SpringerVerlag, New York. 501 p. 
26


Table 1. – Numbers of sediment samples that were greater than or equal to 50% full. 


“Boundary” samples were not collected at either site sampled in 1999.


Status 

Site Open Boundary Closed 

Site 1-1998 8 9 5 

Site 2-1998 9 9 7 

Site 1-1999 29 - 26 

Site 3-1999 19 - 12 
27 

Table 2. – Total organic carbon content (ppm) of sediments collected at three sites near Kodiak 

Island, Alaska. Boundary samples were not collected in 1999. 

Site - Year Status n Mean (SD) Range 

Site 1-1998 All 

Open 

Closed 

Boundary 

Site 1-1999 All 

Open 

Closed 

Site 2-1998 All 

Open 

Closed 

Boundary 

Site 3-1999 All 

Open 

Closed 

30 4092 (617) 2537 - 5116 

10 4331 (616) 3404 - 5116 

10 3827 (684) 2537 - 4586 

10 4118 (488) 3402 - 4952 

54 4251 (623) 2941 - 5819 

29 4255 (617) 2998 - 5819 

25 4264 (642) 2941 - 5256 

30 7276 (1167) 5138 - 9728 

10 8025 (1306) 5987 - 9728 

10 6693 (911) 5167 - 7563 

10 7112 (908) 5138 - 8525 

66 2718 (724) 2110 - 7262 

33 2639 (496) 2127 - 4330 

33 2796 (898) 2110 - 7262 
28


Table 3.–Median grain size (:m) of sediments collected at three sites near Kodiak Island, Alaska. 


Boundary samples were not collected in 1999. 


Site - Year Status n Mean (SD) Range 

Site 1-1998 All 30 143 (29) 101 - 241 

Open 10 141 (25) 101 - 197 

Closed 10 152 (38) 113 - 241 

Boundary 10 137 (24) 104 - 184 

Site 1-1999 All 55 148 (13) 122 - 173 

Open 29 148 (11) 129 - 173 

Closed 26 148 (14) 122 - 171 

Site 2-1998 All 30 86 (13) 57 - 120 

Open 10 77 (12) 57 - 96 

Closed 10 97 (9) 87 - 120 

Boundary 10 86 (8) 67 - 93 

Site 3-1999 All 66 229 (61) 160 - 493 

Open 33 199 (30) 160 - 306 

Closed 33 260 (69) 187 - 493 
29

 Table 4. – Sorting coefficient (N) of sediments collected at three sites near Kodiak Island, 

Alaska. Boundary samples were not collected in 1999. 

Site - Year Status n Mean (SD) Range 

Site 1-1998 All 30 1.03 (0.13) 0.81 - 1.26 

Open 10 1.00 (0.10) 0.86 - 1.21 

Closed 10 1.05 (0.16) 0.81 - 1.26 

Boundary 10 1.04 (0.08) 0.87 - 1.20 

Site 1-1999 All 55 0.97 (0.09) 0.77 - 1.10 

Open 29 0.96 (0.08) 0.78 - 1.07 

Closed 26 0.98 (0.10) 0.77 - 1.10 

Site 2-1998 All 30 0.82 (0.07) 0.75 - 1.08 

Open 10 0.81 (0.06) 0.75 - 0.92 

Closed 10 0.84 (0.09) 0.76 - 1.08 

Boundary 10 0.80 (0.05) 0.75 - 0.88 

Site 3-1999 All 66 1.23 (0.56) 0.48 - 2.69 

Open 33 0.84 (0.26) 0.48 - 1.76 

Closed 33 1.62 (0.49) 0.82 - 2.69 
30 

Table 5. – Summary statistics for testing differences in status (variable indicating open or closed area) for sedime

infauna abundances, and species diversity indices. Statistics include value of the F statistic or |t| statistic (in the c

tests), degrees of freedom (df), P-values, and multiple R2 (%). An arrow indicates the direction of the variable in 

Significance at " = 0.10 is indicated by an asterisk (*). Summary statistics for the 1998 boundary samples versus 

samples are not reported. 

Site 

1 2 

1998 1999 1998 

Variable F or |t| df P R2 Open F or |t| df P R2 Open F or |t| df P R2 Open F or |t| df 

Sediment TOC 1.73 18 0.10* 31 8 0.003 1, 9 0.96 36 - 6.18 18 <0.001* 89 8 1.41 1, 10 

Median grain size 0.89 18 0.39 42 - 0.09 1, 9 0.77 78 - 4.75 18 <0.001* 66 9 24.78 1, 10 

Sorting 1.14 18 0.27 68 - 0.56 1, 9 0.47 80 - 1.22 18 0.24 49 - 70.8 1, 10 Species diversity Richness 0.32 18 0.38 42 - 1.01 9 0.83 72 - 0.38 18 0.35 29 - - - Dominance 0.80 18 0.22 63 - 1.69 9 0.94 83 - 1.51 18 0.07* 42 8 - - 

Grouped taxa All individuals 1.52 18 0.15 81 - 1.45 1, 9 0.26 85 - 1.20 18 0.24 33 - - - Sedentary 0.53 18 0.30 55 - 1.35 9 0.90 85 - 0.62 18 0.73 15 - - - Errant 0.91 18 0.37 35 - 0.52 1, 9 0.49 63 - 1.40 18 0.18 41 - - - 
31


Figure 1.--Map of Kodiak Island, Alaska, showing the three study sites and Type 1 closed areas 

(solid lines) that prohibit non-pelagic trawling year-round. Study Site 1 is located at the Chiniak 

Gully. Study Site 2 is located at the Two-headed Gully. Study Site 3 is located south of the 

Trinity Islands. 
32


Figure 2.--Sampling designs in 1998 and 1999 for Study Site 1 at the Chiniak Gully, near Kodiak 

Island, Alaska. Twenty transects bisected the boundary between the open and closed areas for 

non-pelagic trawling (solid line). Trawl intensity at this site was estimated within an 

approximate 3-km area of the transects (dashed line). 
33


Figure 3.--Relationship between grab fullness (% volume) and (A) number of species and (B) 

number of individuals for infauna samples collected at Site 2 in 1998. Both relationships were 

significant. Samples were collected in areas open to trawling (F), closed to trawling (M), and 

near the boundary (+). 
34


Figure 4.--Relationship between grab fullness (% volume) and (A) number of species and (B) 

number of individuals for infauna samples collected at Site 2 in 1998. Samples were 

standardized to the volume of one full grab. Both relationships were insignificant when grab 

samples less than 50% full (left of dotted line) were excluded. Samples were collected in areas 

open to trawling (F), closed to trawling (M), and near the boundary (+). 
35


Figure 5.--Total organic carbon (TOC, ppm) was significantly higher in sediments from the open 

area (F) compared to the closed area (M) at Site 1 in (A) 1998 (P = 0.10) but not in (B) 1999 (P 

= 0.96). Samples were also collected near the boundary during 1998 (+). 
36


Figure 6.--(A) Total organic carbon (TOC, ppm) was significantly higher (P = 0.001) in 

sediments from the open area (F) compared to the closed area (M) at Site 2 during 1998, but not 

(B) at Site 3 in 1999 (P = 0.26). Samples from open areas are denoted by F, samples from 

closed areas are denoted by M, and samples from near the boundary line during 1998, by +. 
37


Figure 7.--Median sediment grain size in the closed area was significantly larger compared to the 

open area at (A) Site 2 in 1998 (P < 0.001), and (B) Site 3 in 1999 (P < 0.001). Samples from 

open areas are denoted by F, samples from closed areas by M, and samples from near the 

boundary line during 1998, by +. 
38


Figure 8.--Sediments from the closed area (M) were significantly more poorly sorted (P < 0.001) 

than from the open area (F) at Site 3 in 1999. 
39


Figure 9.--Median grain size of sediments collected at 6 equidistant (500 m apart) stations along 

each of 11 transects at Site 3 in 1999. Station A was closest to the south end (farthest into the 

open area) and station F was closest to the north end (farthest into the closed area) of each 

transect. 
40


Figure 10.--Bivariate MDS plot produced after calculation of the Bray-Curtis similarity matrix on 

(A) individual benthic infauna samples (stress = 20%) and (B) transect-averaged benthic infauna 

samples (stress = 14%) from Site 1 during 1999. Samples from each transect were averaged by 

status (i.e., open or closed). Samples from open areas are denoted by F, samples from closed 

areas are denoted by M, and numbers indicate transect number. 

APPENDICES TO CHARACTERISTICS OF BENTHIC SEDIMENTS FROM AREAS 

OPEN AND CLOSED TO BOTTOM TRAWLING IN THE GULF OF ALASKA 

by 

Robert P. Stone 

Auke Bay Laboratory 

Alaska Fisheries Science Center 

National Marine Fisheries Service 

National Oceanic and Atmospheric Administration 

11305 Glacier Highway 

Juneau, AK 99801-8626 

and 

Aaron Baldwin 

Juneau Center School of Fisheries and Ocean Science 

University of Alaska, Fairbanks 

11120 Glacier Highway 

Juneau, Alaska 99801 

October 2003 
iii 

CONTENTS 

INTRODUCTION ............................................................. 1

TABLES ..................................................................... 2

REFERENCES ............................................................. 103
INTRODUCTION

 The information appended in this volume is provided for those interested in the detailed 

composition of infauna collected in sediment grab samples from three areas near Kodiak Island, 

Alaska. Samples were collected to examine potential chronic effects of bottom trawling (otter 

trawls, and to a lesser extent, scallop dredges) on the sedimentary, biogeochemical, and 

biological features of the seafloor in soft-bottom, low-relief marine habitat of the inner 

continental shelf in the central Gulf of Alaska.

 Numerous studies have been published worldwide that summarize analyses of infauna data, 

but few provide raw data sets from which comparative analyses can be performed (Pearson et al. 

1983). Infauna collections have been made at a number of locations in the Gulf of Alaska and 

Bering Sea, but no common database exists of species present in those areas. Consequently, the 

zoogeography of many species is poorly known and taxonomic efforts presently underway may 

be largely duplicated. These appendices are provided to assist in future taxonomic revisions of 

North Pacific species and for comparative analyses. Also included is a list of references used in 

the taxonomic identifications of infauna. 
2 

Appendix Table 1.--Total numbers of errant annelids collected in sediment grab samples at Site 1 

and Site 2. 

Order Family Species Site 1-1998 Site 1-1999 Site 2 

Cossurida Cossuridae Cossura soyeri 1 3 17 

Eunicida Lumbrineridae Lumbrineris sp. 7 

Eunicida Lumbrineridae Lumbrineris bicirrata 4 6 1 

Eunicida Lumbrineridae Lumbrineris inflata 1 1 

Eunicida Lumbrineridae Lumbrineris luti 142 410 14 

Magelonida Magelonidae Magelona berkeleyi 15 43 43 

Opheliida Opheliida Ophelia assimilis 1 

Opheliida Opheliida Travisia forbesii 5 6 1 

Opheliida Scalibregmidae Scalibregma inflatum 1 1 

Orbiniida Orbiinidae Leitoscoloplos pugettensis 90 199 36 

Orbiniida Paraonidae Aricidea lopezi 1 49 8 

Orbiniida Paraonidae Chaetozone sp. 34 52 76 

Orbiniida Paraonidae Levinsenia gracilis 1 6 34 

Phyllodocida Glyceridae Glycera sp. 3 

Phyllodocida Glyceridae Hemipodus borealis 40 42 15 

Phyllodocida Goniadidae Goniada brunnea 29 42 4 

Phyllodocida Nephtyidae Nephtys caeca 110 183 88 

Phyllodocida Nephtyidae Nephtys cornuta 22 68 23 

Phyllodocida Pholodidae Pholoe glabra 22 26 9 

Phyllodocida Phyllodocidae Eteone longa 2 4 

Phyllodocida Phyllodocidae Phyllodocida sp. 1 

Phyllodocida Polynoidae Hesperonoe sp. 2 

Phyllodocida Sphaerodoropsidae Sphaerodoropsis sp. 2 

Phyllodocida Syllidae Syllis sp. 15 29 
3 

Appendix Table 2.--Total numbers of sedentary annelids collected in sediment grab samples at 

Site 1 and Site 2. 

Order Family Species Site 1-1998 Site 1-1999 Site 2 

Capitellida Capitellidae Mediomastus californiensis 104 285 28 

Capitellida Capitellidae Notomastus giganteus 7 17 2 

Capitellida Maldanidae undetermined 11 14 

Capitellida Maldanidae Axiothella rubrocincta 13 1 

Capitellida Maldanidae Clymenella sp. 6 

Capitellida Maldanidae Euclymene zonalis 9 114 

Capitellida Maldanidae Praxillella gracilis 3 1 1 

Chaetopterida Chaetopteridae Spiochaetopterus costarum 5 1 

Eunicida Onuphidae Onuphis iridescens 11 24 4 

Oweniida Oweniidae Owenia cf. fusiformis 16 28 

Sabellida Sabellidae undetermined 3 6 2 

Sabellida Sabellidae Chone sp. 1 3 

Spionida Spionidae Dipolydora socialis 3 6 

Spionida Spionidae Prionospio lighti 3 4 

Spionida Spionidae Prionospio steenstrupi 45 90 111 

Spionida Spionidae Pygospio elegans 1 1 3 

Spionida Spionidae Spio cirrifera 1 7 2 

Terebellida Amphaeretidae Amphicteis scaphobranchiata 11 78 3 

Terebellida Pectinaridae Pectinaria moorei 3 

Terebellida Terebellidae Amphitrite cirrata 3 

Terebellida Terebellidae Scionella sp. 3 

Terebellida Terebellidae Artacama conifera 5 

Terebellida Terebellidae Terebellides stroemi 5 1 17 
4 

Appendix Table 3.--Number of individuals of each infauna taxon collected per grab sample at 

Transects 1 and 2 at Chiniak Gully during June 1998. 

Phylum Order Family Species 1A 1B 1C 2A 2B 2C 

Annelida Capitallida Capitellidae Mediomastus californiensis 1 5 3 

Annelida Capitallida Capitellidae Notomastus giganteus 

Annelida Capitallida Maldanidae 

Annelida Capitallida Maldanidae Asychis similis 

Annelida Capitallida Maldanidae Axiothella rubrocincta 

Annelida Capitallida Maldanidae Euclymene zonalis 

Annelida Capitallida Maldanidae Praxillella gracilis 

Annelida Cossurida Cossuridae Cossura soyeri 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 1 1 

Annelida Eunicida Lumbrineridae Lumbrineris inflata 

Annelida Eunicida Lumbrineridae Lumbrineris luti 1 3 11 

Annelida Eunicida Onuphidae 

Annelida Magelonida Magelonidae Magelona berkeleyi 2 

Annelida Opheliida Opheliidae Ophelina sp. 

Annelida Opheliida Opheliidae Travisia forbesii 

Annelida Opheliida Scalibregmidae Hyboscolex pacificus 

Annelida Opheliida Scalibregmidae Scalibregma inflatum 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 5 4 2 2 6 2 

Annelida Orbiniida Paraonidae Aricidea lopezi 

Annelida Orbiniida Paraonidae Chaetozone sp. 2 

Annelida Orbiniida Paraonidae Levinsenia gracilis 

Annelida Oweniida Oweniidae Owenia cf. fusiformis 

Annelida Phyllodocida Glyceridae Hemipodus borealis 2 1 1 1 

Annelida Phyllodocida Goniadidae Goniada brunnea 4 1 1 1 

Annelida Phyllodocida Hesionidae Micropodarke cf. dubiens 

Annelida Phyllodocida Nephtidae Nephtys caeca 6 9 1 9 3 

Annelida Phyllodocida Nephtidae Nephtys cornuta 5 

Annelida Phyllodocida Pholodidae Pholoe glabra 1 5 1 

Annelida Phyllodocida Phyllodocidae Eteone longa 1 
5


Annelida Phyllodocida Syllidae Syllis sp. 1 1 

Annelida Sabellida Sabellidae 

Annelida Sabellida Sabellidae Chone sp. 

Annelida Spionida Spionidae Dipolydora socialis 

Annelida Spionida Spionidae Laonice sp. 

Annelida Spionida Spionidae Prionospio sp. 

Annelida Spionida Spionidae Prionospio steenstrupi 2 1 

Annelida Spionida Spionidae Pygospio elegans 

Annelida Spionida Spionidae Spio cirrifera 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 4 1 

Annelida Terebellida Trichobranchidae Terebellides stroemi 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 1 2 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 

Arthropoda Amphipoda Corophiidae Corophium cf. insidiosum 

Arthropoda Amphipoda Phoxocephalidae Heterophoxus sp. 3 

Arthropoda Cumacea 1 

Arthropoda Decapoda Pinnixidae Pinnixa sp. 1 5 2 

Arthropoda Decapoda Xanthidae Lophopanopeus bellus 1 

Arthropoda Ostracoda Species I 

Echinodermata Dendrochiridotida 1 1 

Echinodermata Dendrochiridotida Phyllophoridae Pentamera sp. 

Echinodermata Ophiurida Amphiuridae Amphipholus sp. 3 8 6 24 21 38 

Mollusca 2 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 1 2 

Mollusca Mesogastropoda Eulimidae Eulima sp. 

Mollusca Mesogastropoda Naticidae Cryptonatica sp. 

Mollusca Mytiloida Mytilidae Musculus sp. 1 

Mollusca Neogastropoda Turridae Oenopota rosea 1 1 1 

Mollusca Nuculoida Nuculanidae Nuculana pernula 

Mollusca Nuculoida Nuculidae Ennucula tenuis 3 5 3 5 

Mollusca Nuculoida Yoldiidae Yoldia scissurata 2 3 1 

Mollusca Pholadomyoida Pandoridae Pandora bilirata 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 
6


Mollusca Veneroida Cultellidae Siliqua patula 1 10 2 6 

Mollusca Veneroida Tellinidae Macoma eliminata 1 1 3 2 

Mollusca Veneroida Tellinidae Macoma moesta 7 

Mollusca Veneroida Thyasiridae Axinospida serricata 14 23 6 20 32 25 

Mollusca Veneroida Veneridae Nutricola lordi 

Nemertea Heteronemertea Lineidae Linea sp. 1 

Total No. of Species 12 14 5 22 22 11 

Total No. of Individuals 41 69 16 97 114 80 
7 

Appendix Table 3.--Number of individuals of each infauna taxon collected per grab sample at 

Transects 3 and 4 at Chiniak Gully during June 1998. 

Phylum Order Family Species 3A 3B 3C 4A 4B 4C 

Annelida Capitallida Capitellidae Mediomastus californiensis 1 8 9 

Annelida Capitallida Capitellidae Notomastus giganteus 

Annelida Capitallida Maldanidae 

Annelida Capitallida Maldanidae Asychis similis 

Annelida Capitallida Maldanidae Axiothella rubrocincta 5 

Annelida Capitallida Maldanidae Euclymene zonalis 

Annelida Capitallida Maldanidae Praxillella gracilis 3 

Annelida Cossurida Cossuridae Cossura soyeri 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 2 

Annelida Eunicida Lumbrineridae Lumbrineris inflata 1 

Annelida Eunicida Lumbrineridae Lumbrineris luti 2 5 9 21 

Annelida Eunicida Onuphidae 1 1 

Annelida Magelonida Magelonidae Magelona berkeleyi 

Annelida Opheliida Opheliidae Ophelina sp. 

Annelida Opheliida Opheliidae Travisia forbesii 

Annelida Opheliida Scalibregmidae Hyboscolex pacificus 

Annelida Opheliida Scalibregmidae Scalibregma inflatum 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 5 11 1 1 1 

Annelida Orbiniida Paraonidae Aricidea lopezi 

Annelida Orbiniida Paraonidae Chaetozone sp. 

Annelida Orbiniida Paraonidae Levinsenia gracilis 

Annelida Oweniida Oweniidae Owenia cf. fusiformis 3 

Annelida Phyllodocida Glyceridae Hemipodus borealis 1 2 5 

Annelida Phyllodocida Goniadidae Goniada brunnea 1 1 1 4 

Annelida Phyllodocida Hesionidae Micropodarke cf. dubiens 

Annelida Phyllodocida Nephtidae Nephtys caeca 5 1 4 2 3 

Annelida Phyllodocida Nephtidae Nephtys cornuta 2 

Annelida Phyllodocida Pholodidae Pholoe glabra 2 4 2 4 1 

Annelida Phyllodocida Phyllodocidae Eteone longa 
8


Annelida Phyllodocida Syllidae Syllis sp. 1 2 1 

Annelida Sabellida Sabellidae 1 

Annelida Sabellida Sabellidae Chone sp. 

Annelida Spionida Spionidae Dipolydora socialis 2 

Annelida Spionida Spionidae Laonice sp. 

Annelida Spionida Spionidae Prionospio sp. 

Annelida Spionida Spionidae Prionospio steenstrupi 1 1 

Annelida Spionida Spionidae Pygospio elegans 

Annelida Spionida Spionidae Spio cirrifera 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 

Annelida Terebellida Trichobranchidae Terebellides stroemi 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 2 1 2 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 

Arthropoda Amphipoda Corophiidae Corophium cf. insidiosum 1 

Arthropoda Amphipoda Phoxocephalidae Heterophoxus sp. 1 2 

Arthropoda Cumacea 

Arthropoda Decapoda Pinnixidae Pinnixa sp. 4 

Arthropoda Decapoda Xanthidae Lophopanopeus bellus 

Arthropoda Ostracoda Species I 

Echinodermata Dendrochiridotida 1 

Echinodermata Dendrochiridotida Phyllophoridae Pentamera sp. 

Echinodermata Ophiurida Amphiuridae Amphipholus sp. 18 28 14 64 10 37 

Mollusca 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 

Mollusca Mesogastropoda Eulimidae Eulima sp. 

Mollusca Mesogastropoda Naticidae Cryptonatica sp. 1 

Mollusca Mytiloida Mytilidae Musculus sp. 

Mollusca Neogastropoda Turridae Oenopota rosea 1 2 1 

Mollusca Nuculoida Nuculanidae Nuculana pernula 

Mollusca Nuculoida Nuculidae Ennucula tenuis 5 

Mollusca Nuculoida Yoldiidae Yoldia scissurata 4 

Mollusca Pholadomyoida Pandoridae Pandora bilirata 1 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 
9


Mollusca Veneroida Cultellidae Siliqua patula 2 3 2 6 1 

Mollusca Veneroida Tellinidae Macoma eliminata 2 3 3 

Mollusca Veneroida Tellinidae Macoma moesta 1 2 

Mollusca Veneroida Thyasiridae Axinospida serricata 11 11 22 27 36 

Mollusca Veneroida Veneridae Nutricola lordi 1 

Nemertea Heteronemertea Lineidae Linea sp. 

Total No. of Species 15 10 8 21 8 19 

Total No. of Individuals 55 67 44 148 23 140 
10 

Appendix Table 3.--Number of individuals of each infauna taxon collected per grab sample at 

Transects 5 and 6 at Chiniak Gully during June 1998. 

Phylum Order Family Species 5A 5B 5C 6A 6B 6C 

Annelida Capitallida Capitellidae Mediomastus californiensis 7 3 8 2 4 

Annelida Capitallida Capitellidae Notomastus giganteus 

Annelida Capitallida Maldanidae 4 6 

Annelida Capitallida Maldanidae Asychis similis 

Annelida Capitallida Maldanidae Axiothella rubrocincta 3 5 

Annelida Capitallida Maldanidae Euclymene zonalis 

Annelida Capitallida Maldanidae Praxillella gracilis 

Annelida Cossurida Cossuridae Cossura soyeri 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 

Annelida Eunicida Lumbrineridae Lumbrineris inflata 

Annelida Eunicida Lumbrineridae Lumbrineris luti 7 3 9 2 4 10 

Annelida Eunicida Onuphidae 1 1 

Annelida Magelonida Magelonidae Magelona berkeleyi 1 2 1 

Annelida Opheliida Opheliidae Ophelina sp. 

Annelida Opheliida Opheliidae Travisia forbesii 3 

Annelida Opheliida Scalibregmidae Hyboscolex pacificus 1 

Annelida Opheliida Scalibregmidae Scalibregma inflatum 1 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 2 3 1 3 

Annelida Orbiniida Paraonidae Aricidea lopezi 

Annelida Orbiniida Paraonidae Chaetozone sp. 1 

Annelida Orbiniida Paraonidae Levinsenia gracilis 

Annelida Oweniida Oweniidae Owenia cf. fusiformis 5 1 1 

Annelida Phyllodocida Glyceridae Hemipodus borealis 1 1 1 2 

Annelida Phyllodocida Goniadidae Goniada brunnea 2 1 4 2 

Annelida Phyllodocida Hesionidae Micropodarke cf. dubiens 

Annelida Phyllodocida Nephtidae Nephtys caeca 7 5 5 6 5 5 

Annelida Phyllodocida Nephtidae Nephtys cornuta 3 

Annelida Phyllodocida Pholodidae Pholoe glabra 1 

Annelida Phyllodocida Phyllodocidae Eteone longa 
11


Annelida Phyllodocida Syllidae Syllis sp. 1 3 

Annelida Sabellida Sabellidae 

Annelida Sabellida Sabellidae Chone sp. 

Annelida Spionida Spionidae Dipolydora socialis 

Annelida Spionida Spionidae Laonice sp. 

Annelida Spionida Spionidae Prionospio sp. 

Annelida Spionida Spionidae Prionospio steenstrupi 1 1 1 1 2 1 

Annelida Spionida Spionidae Pygospio elegans 

Annelida Spionida Spionidae Spio cirrifera 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 2 1 

Annelida Terebellida Trichobranchidae Terebellides stroemi 2 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 1 3 1 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 1 3 1 2 

Arthropoda Amphipoda Corophiidae Corophium cf. insidiosum 2 

Arthropoda Amphipoda Phoxocephalidae Heterophoxus sp. 2 4 7 3 2 3 

Arthropoda Cumacea 

Arthropoda Decapoda Pinnixidae Pinnixa sp. 5 9 3 

Arthropoda Decapoda Xanthidae Lophopanopeus bellus 

Arthropoda Ostracoda Species I 

Echinodermata Dendrochiridotida 3 1 1 

Echinodermata Dendrochiridotida Phyllophoridae Pentamera sp. 

Echinodermata Ophiurida Amphiuridae Amphipholus sp. 5 8 2 1 

Mollusca 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 

Mollusca Mesogastropoda Eulimidae Eulima sp. 1 

Mollusca Mesogastropoda Naticidae Cryptonatica sp. 

Mollusca Mytiloida Mytilidae Musculus sp. 

Mollusca Neogastropoda Turridae Oenopota rosea 1 

Mollusca Nuculoida Nuculanidae Nuculana pernula 

Mollusca Nuculoida Nuculidae Ennucula tenuis 

Mollusca Nuculoida Yoldiidae Yoldia scissurata 3 2 

Mollusca Pholadomyoida Pandoridae Pandora bilirata 1 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 1 
12


Mollusca Veneroida Cultellidae Siliqua patula 1 1 1 

Mollusca Veneroida Tellinidae Macoma eliminata 3 1 1 3 1 

Mollusca Veneroida Tellinidae Macoma moesta 2 2 1 1 

Mollusca Veneroida Thyasiridae Axinospida serricata 22 4 12 2 2 2 

Mollusca Veneroida Veneridae Nutricola lordi 

Nemertea Heteronemertea Lineidae Linea sp. 2 2 1 1 

Total No. of Species 20 16 27 18 14 15 

Total No. of Individuals 75 49 94 31 29 44 
13 

Appendix Table 3.--Number of individuals of each infauna taxon collected per grab sample at 

Transects 7 and 8 at Chiniak Gully during June 1998. 

Phylum Order Family Species 7A 7B 7C 8A 8B 8C 

Annelida Capitallida Capitellidae Mediomastus californiensis 9 2 2 6 16 

Annelida Capitallida Capitellidae Notomastus giganteus 6 

Annelida Capitallida Maldanidae 1 

Annelida Capitallida Maldanidae Asychis similis 1 

Annelida Capitallida Maldanidae Axiothella rubrocincta 

Annelida Capitallida Maldanidae Euclymene zonalis 9 

Annelida Capitallida Maldanidae Praxillella gracilis 

Annelida Cossurida Cossuridae Cossura soyeri 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 

Annelida Eunicida Lumbrineridae Lumbrineris inflata 

Annelida Eunicida Lumbrineridae Lumbrineris luti 4 5 2 3 5 3 

Annelida Eunicida Onuphidae 1 1 1 1 

Annelida Magelonida Magelonidae Magelona berkeleyi 3 2 1 

Annelida Opheliida Opheliidae Ophelina sp. 

Annelida Opheliida Opheliidae Travisia forbesii 

Annelida Opheliida Scalibregmidae Hyboscolex pacificus 

Annelida Opheliida Scalibregmidae Scalibregma inflatum 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 4 3 3 4 8 

Annelida Orbiniida Paraonidae Aricidea lopezi 1 

Annelida Orbiniida Paraonidae Chaetozone sp. 1 2 2 1 

Annelida Orbiniida Paraonidae Levinsenia gracilis 

Annelida Oweniida Oweniidae Owenia cf. fusiformis 3 

Annelida Phyllodocida Glyceridae Hemipodus borealis 2 1 5 4 

Annelida Phyllodocida Goniadidae Goniada brunnea 1 1 2 

Annelida Phyllodocida Hesionidae Micropodarke cf. dubiens 1 

Annelida Phyllodocida Nephtidae Nephtys caeca 5 3 3 3 6 

Annelida Phyllodocida Nephtidae Nephtys cornuta 1 1 

Annelida Phyllodocida Pholodidae Pholoe glabra 

Annelida Phyllodocida Phyllodocidae Eteone longa 1 
14


Annelida Phyllodocida Syllidae Syllis sp. 2 2 1 

Annelida Sabellida Sabellidae 

Annelida Sabellida Sabellidae Chone sp. 

Annelida Spionida Spionidae Dipolydora socialis 

Annelida Spionida Spionidae Laonice sp. 

Annelida Spionida Spionidae Prionospio sp. 

Annelida Spionida Spionidae Prionospio steenstrupi 4 2 4 4 

Annelida Spionida Spionidae Pygospio elegans 1 

Annelida Spionida Spionidae Spio cirrifera 1 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 

Annelida Terebellida Trichobranchidae Terebellides stroemi 1 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 1 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 1 1 6 2 

Arthropoda Amphipoda Corophiidae Corophium cf. insidiosum 

Arthropoda Amphipoda Phoxocephalidae Heterophoxus sp. 2 2 2 2 5 

Arthropoda Cumacea 1 

Arthropoda Decapoda Pinnixidae Pinnixa sp. 

Arthropoda Decapoda Xanthidae Lophopanopeus bellus 

Arthropoda Ostracoda Species I 1 

Echinodermata Dendrochiridotida 

Echinodermata Dendrochiridotida Phyllophoridae Pentamera sp. 1 

Echinodermata Ophiurida Amphiuridae Amphipholus sp. 

Mollusca 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 1 

Mollusca Mesogastropoda Eulimidae Eulima sp. 1 2 

Mollusca Mesogastropoda Naticidae Cryptonatica sp. 

Mollusca Mytiloida Mytilidae Musculus sp. 

Mollusca Neogastropoda Turridae Oenopota rosea 1 1 1 

Mollusca Nuculoida Nuculanidae Nuculana pernula 

Mollusca Nuculoida Nuculidae Ennucula tenuis 

Mollusca Nuculoida Yoldiidae Yoldia scissurata 

Mollusca Pholadomyoida Pandoridae Pandora bilirata 1 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 1 1 2 
15


Mollusca Veneroida Cultellidae Siliqua patula 

Mollusca Veneroida Tellinidae Macoma eliminata 1 

Mollusca Veneroida Tellinidae Macoma moesta 4 2 2 1 3 

Mollusca Veneroida Thyasiridae Axinospida serricata 5 2 2 3 14 

Mollusca Veneroida Veneridae Nutricola lordi 1 1 

Nemertea Heteronemertea Lineidae Linea sp. 

Total No. of Species 15 14 9 15 20 22 

Total No. of Individuals 43 29 15 37 48 86 
16 

Appendix Table 3.--Number of individuals of each infauna taxon collected per grab sample at 

Transects 9 and 10 at Chiniak Gully during June 1998. 

Phylum Order Family Species 9A 9B 9C 10A 10B 10C 

Annelida Capitallida Capitellidae Mediomastus californiensis 5 5 8 

Annelida Capitallida Capitellidae Notomastus giganteus 1 

Annelida Capitallida Maldanidae 

Annelida Capitallida Maldanidae Asychis similis 

Annelida Capitallida Maldanidae Axiothella rubrocincta 

Annelida Capitallida Maldanidae Euclymene zonalis 

Annelida Capitallida Maldanidae Praxillella gracilis 

Annelida Cossurida Cossuridae Cossura soyeri 1 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 

Annelida Eunicida Lumbrineridae Lumbrineris inflata 

Annelida Eunicida Lumbrineridae Lumbrineris luti 7 9 7 8 2 

Annelida Eunicida Onuphidae 1 2 

Annelida Magelonida Magelonidae Magelona berkeleyi 1 2 

Annelida Opheliida Opheliidae Ophelina sp. 1 

Annelida Opheliida Opheliidae Travisia forbesii 1 1 

Annelida Opheliida Scalibregmidae Hyboscolex pacificus 

Annelida Opheliida Scalibregmidae Scalibregma inflatum 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 2 7 1 5 2 2 

Annelida Orbiniida Paraonidae Aricidea lopezi 

Annelida Orbiniida Paraonidae Chaetozone sp. 10 5 3 4 1 2 

Annelida Orbiniida Paraonidae Levinsenia gracilis 1 

Annelida Oweniida Oweniidae Owenia cf. fusiformis 2 1 

Annelida Phyllodocida Glyceridae Hemipodus borealis 2 3 3 1 1 

Annelida Phyllodocida Goniadidae Goniada brunnea 2 

Annelida Phyllodocida Hesionidae Micropodarke cf. dubiens 

Annelida Phyllodocida Nephtidae Nephtys caeca 4 2 3 4 1 

Annelida Phyllodocida Nephtidae Nephtys cornuta 5 5 

Annelida Phyllodocida Pholodidae Pholoe glabra 1 

Annelida Phyllodocida Phyllodocidae Eteone longa 
17


Annelida Phyllodocida Syllidae Syllis sp. 

Annelida Sabellida Sabellidae 2 

Annelida Sabellida Sabellidae Chone sp. 1 

Annelida Spionida Spionidae Dipolydora socialis 1 

Annelida Spionida Spionidae Laonice sp. 1 

Annelida Spionida Spionidae Prionospio sp. 1 

Annelida Spionida Spionidae Prionospio steenstrupi 6 3 2 2 3 3 

Annelida Spionida Spionidae Pygospio elegans 

Annelida Spionida Spionidae Spio cirrifera 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 2 1 

Annelida Terebellida Trichobranchidae Terebellides stroemi 1 1 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 2 3 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 1 4 1 3 

Arthropoda Amphipoda Corophiidae Corophium cf. insidiosum 

Arthropoda Amphipoda Phoxocephalidae Heterophoxus sp. 4 3 1 1 

Arthropoda Cumacea 1 1 

Arthropoda Decapoda Pinnixidae Pinnixa sp. 1 1 

Arthropoda Decapoda Xanthidae Lophopanopeus bellus 

Arthropoda Ostracoda Species I 

Echinodermata Dendrochiridotida 1 

Echinodermata Dendrochiridotida Phyllophoridae Pentamera sp. 

Echinodermata Ophiurida Amphiuridae Amphipholus sp. 2 

Mollusca 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 2 

Mollusca Mesogastropoda Eulimidae Eulima sp. 

Mollusca Mesogastropoda Naticidae Cryptonatica sp. 

Mollusca Mytiloida Mytilidae Musculus sp. 

Mollusca Neogastropoda Turridae Oenopota rosea 

Mollusca Nuculoida Nuculanidae Nuculana pernula 3 

Mollusca Nuculoida Nuculidae Ennucula tenuis 1 

Mollusca Nuculoida Yoldiidae Yoldia scissurata 1 

Mollusca Pholadomyoida Pandoridae Pandora bilirata 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 
18


Mollusca Veneroida Cultellidae Siliqua patula 

Mollusca Veneroida Tellinidae Macoma eliminata 2 4 1 2 

Mollusca Veneroida Tellinidae Macoma moesta 2 1 

Mollusca Veneroida Thyasiridae Axinospida serricata 2 1 3 14 5 4 

Mollusca Veneroida Veneridae Nutricola lordi 1 6 

Nemertea Heteronemertea Lineidae Linea sp. 

Total No. of Species 19 15 17 12 16 16 

Total No. of Individuals 57 48 38 38 41 40 
19 

Appendix Table 4.--Number of individuals of each infauna taxon collected per grab sample at 

Transects 1 and 2 at Two-headed Gully during June 1998. 

Phylum Order Family Species 1A 1B 1C 2A 2B 2C 

Annelida Capitallida Capitellidae Mediomastus californiensis 2 3 

Annelida Capitallida Capitellidae Notomastus giganteus 

Annelida Capitallida Maldanidae Praxillella gracilis 

Annelida Chaetopterida Chaetopteridae Spiochaetopterus costarum 

Annelida Cossurida Cossuridae Cossura soyeri 3 1 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 

Annelida Eunicida Lumbrineridae Lumbrineris luti 1 1 

Annelida Eunicida Onuphidae 

Annelida Magelonida Magelonidae Magelona berkeleyi 2 5 

Annelida Opheliida Opheliidae Travisia forbesii 

Annelida Opheliida Scalibregmidae Hyboscolex pacificus 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 1 4 2 

Annelida Orbiniida Paraonidae Aricidea lopezi 1 1 

Annelida Orbiniida Paraonidae Chaetozone sp. 2 9 3 5 1 

Annelida Orbiniida Paraonidae Levinsenia gracilis 7 2 

Annelida Phyllodocida Glyceridae Hemipodus borealis 1 

Annelida Phyllodocida Goniadidae Goniada brunnea 1 

Annelida Phyllodocida Hesionidae 

Annelida Phyllodocida Nephtidae Nephtys caeca 9 4 5 3 

Annelida Phyllodocida Nephtidae Nephtys cornuta 1 1 

Annelida Phyllodocida Nereididae Cyclonereis clyclurus 

Annelida Phyllodocida Pholodidae Pholoe glabra 2 

Annelida Sabellida Sabellidae 

Annelida Spionida Spionidae Prionospio lighti 

Annelida Spionida Spionidae Prionospio steenstrupi 3 7 8 3 1 2 

Annelida Spionida Spionidae Pygospio elegans 

Annelida Spionida Spionidae Spio cirrifera 

Annelida Sternapsida Sternapsidae Sternapsis scutata 1 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 
20


Annelida Terebellida Terebellidae Artacama conifera 

Annelida Terebellida Trichobranchidae Terebellides stroemi 2 2 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 5 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 8 

Arthropoda Amphipoda Phoxocephalidae Heterophoxus sp. 

Arthropoda Calanoida 

Arthropoda Cumacea 1 1 

Arthropoda Ostracoda Species I 

Echinodermata Dendrochiridotida Phyllophoridae Pentamera sp. 

Echiura Echiuroinea Echiuridae Echiurus echiurus alaskensis 

Mollusca Cephalaspidea Atyidae Haminoea virescens 2 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 2 

Mollusca Mesogastropoda Naticidae Euspira pallida 

Mollusca Neogastropoda Neptuneidae Colus halli 

Mollusca Nuculoida Nuculanidae Nuculana fossa 1 

Mollusca Nuculoida Nuculanidae Nucula minuta 1 1 

Mollusca Nuculoida Nuculidae Ennucula tenuis 1 1 2 

Mollusca Nuculoida Yoldiidae Yoldia scissurata 2 

Mollusca Pholadomyoida Pandoridae Pandora bilirata 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 1 2 

Mollusca Veneroida Tellinidae Macoma eliminata 1 1 1 

Mollusca Veneroida Thyasiridae Axinospida serricata 4 10 4 2 3 

Nematoda 

Nemertea Heteronemertea Lineidae Cerebratulus sp. 

Nemertea Heteronemertea Lineidae Linea sp. 2 

Total No. of Species 9 8 11 11 9 14 

Total No. of Individuals 16 34 45 34 15 27 
21 

Appendix Table 4.--Number of individuals of each infauna taxon collected per grab sample at 

Transects 3 and 4 at Two-headed Gully during June 1998. 

Phylum Order Family Species 3A 3B 3C 4A 4B 4C 

Annelida Capitallida Capitellidae Mediomastus californiensis 3 4 2 

Annelida Capitallida Capitellidae Notomastus giganteus 

Annelida Capitallida Maldanidae Praxillella gracilis 

Annelida Chaetopterida Chaetopteridae Spiochaetopterus costarum 1 

Annelida Cossurida Cossuridae Cossura soyeri 4 3 2 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 1 

Annelida Eunicida Lumbrineridae Lumbrineris luti 1 

Annelida Eunicida Onuphidae 

Annelida Magelonida Magelonidae Magelona berkeleyi 1 1 4 

Annelida Opheliida Opheliidae Travisia forbesii 1 

Annelida Opheliida Scalibregmidae Hyboscolex pacificus 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 3 3 3 2 

Annelida Orbiniida Paraonidae Aricidea lopezi 2 

Annelida Orbiniida Paraonidae Chaetozone sp. 6 1 2 5 2 

Annelida Orbiniida Paraonidae Levinsenia gracilis 5 2 2 1 

Annelida Phyllodocida Glyceridae Hemipodus borealis 1 1 5 1 1 

Annelida Phyllodocida Goniadidae Goniada brunnea 

Annelida Phyllodocida Hesionidae 

Annelida Phyllodocida Nephtidae Nephtys caeca 1 4 5 

Annelida Phyllodocida Nephtidae Nephtys cornuta 3 2 6 2 

Annelida Phyllodocida Nereididae Cyclonereis clyclurus 

Annelida Phyllodocida Pholodidae Pholoe glabra 1 2 

Annelida Sabellida Sabellidae 1 

Annelida Spionida Spionidae Prionospio lighti 3 

Annelida Spionida Spionidae Prionospio steenstrupi 6 4 1 6 

Annelida Spionida Spionidae Pygospio elegans 

Annelida Spionida Spionidae Spio cirrifera 

Annelida Sternapsida Sternapsidae Sternapsis scutata 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 1 
22


Annelida Terebellida Terebellidae Artacama conifera 1 

Annelida Terebellida Trichobranchidae Terebellides stroemi 2 2 1 5 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 

Arthropoda Amphipoda Phoxocephalidae Heterophoxus sp. 

Arthropoda Calanoida 

Arthropoda Cumacea 

Arthropoda Ostracoda Species I 1 

Echiura Dendrochiridotida Phyllophoridae Pentamera sp. 1 

Echinodermata Echiuroinea Echiuridae Echiurus echiurus alaskensis 1 

Mollusca Cephalaspidea Atyidae Haminoea virescens 1 1 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 1 1 

Mollusca Mesogastropoda Naticidae Euspira pallida 

Mollusca Neogastropoda Neptuneidae Colus halli 

Mollusca Nuculoida Nuculanidae Nuculana fossa 

Mollusca Nuculoida Nuculanidae Nuculana minuta 1 8 

Mollusca Nuculoida Nuculidae Ennucula tenuis 1 1 

Mollusca Nuculoida Yoldiidae Yoldia scissurata 1 

Mollusca Pholadomyoida Pandoridae Pandora bilirata 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 

Mollusca Veneroida Tellinidae Macoma eliminata 1 7 4 2 

Mollusca Veneroida Thyasiridae Axinospida serricata 7 20 2 5 6 6 

Nematoda 

Nemertea Heteronemertea Lineidae Cerebratulus sp. 

Nemertea Heteronemertea Lineidae Linea sp. 

Total No. of Species 12 12 12 11 18 10 

Total No. of Individuals 40 41 28 31 47 27 
23 

Appendix Table 4.--Number of individuals of each infauna taxon collected per grab sample at 

Transects 5 and 6 at Two-headed Gully during June 1998. 

Phylum Order Family Species 5A 5B 5C 6A 6B 6C 

Annelida Capitallida Capitellidae Mediomastus californiensis 1 5 1 3 

Annelida Capitallida Capitellidae Notomastus giganteus 1 

Annelida Capitallida Maldanidae Praxillella gracilis 1 

Annelida Chaetopterida Chaetopteridae Spiochaetopterus costarum 

Annelida Cossurida Cossuridae Cossura soyeri 2 1 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 

Annelida Eunicida Lumbrineridae Lumbrineris luti 1 

Annelida Eunicida Onuphidae 

Annelida Magelonida Magelonidae Magelona berkeleyi 4 4 1 7 1 

Annelida Opheliida Opheliidae Travisia forbesii 

Annelida Opheliida Scalibregmidae Hyboscolex pacificus 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 3 3 3 2 

Annelida Orbiniida Paraonidae Aricidea lopezi 1 1 1 

Annelida Orbiniida Paraonidae Chaetozone sp. 2 1 3 9 

Annelida Orbiniida Paraonidae Levinsenia gracilis 5 5 1 

Annelida Phyllodocida Glyceridae Hemipodus borealis 2 2 

Annelida Phyllodocida Goniadidae Goniada brunnea 1 2 

Annelida Phyllodocida Hesionidae 

Annelida Phyllodocida Nephtidae Nephtys caeca 1 6 1 6 5 6 

Annelida Phyllodocida Nephtidae Nephtys cornuta 2 

Annelida Phyllodocida Nereididae Cyclonereis clyclurus 

Annelida Phyllodocida Pholodidae Pholoe glabra 1 2 

Annelida Sabellida Sabellidae 

Annelida Spionida Spionidae Prionospio lighti 

Annelida Spionida Spionidae Prionospio steenstrupi 4 5 3 1 2 9 

Annelida Spionida Spionidae Pygospio elegans 1 

Annelida Spionida Spionidae Spio cirrifera 1 

Annelida Sternapsida Sternapsidae Sternapsis scutata 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 
24


Annelida Terebellida Terebellidae Artacama conifera 

Annelida Terebellida Trichobranchidae Terebellides stroemi 3 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 1 2 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 

Arthropoda Amphipoda Phoxocephalidae Heterophoxus sp. 1 

Arthropoda Calanoida 1 

Arthropoda Cumacea 1 

Arthropoda Ostracoda Species I 

Echinodermata Dendrochiridotida Phyllophoridae Pentamera sp. 1 

Echiura Echiuroinea Echiuridae Echiurus echiurus alaskensis 

Mollusca Cephalaspidea Atyidae Haminoea virescens 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 1 

Mollusca Mesogastropoda Naticidae Euspira pallida 

Mollusca Neogastropoda Neptuneidae Colus halli 1 

Mollusca Nuculoida Nuculanidae Nuculana fossa 

Mollusca Nuculoida Nuculanidae Nuculana minuta 1 

Mollusca Nuculoida Nuculidae Ennucula tenuis 1 

Mollusca Nuculoida Yoldiidae Yoldia scissurata 1 

Mollusca Pholadomyoida Pandoridae Pandora bilirata 1 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 1 2 1 

Mollusca Veneroida Tellinidae Macoma eliminata 3 3 1 1 2 

Mollusca Veneroida Thyasiridae Axinospida serricata 19 3 8 10 10 4 

Nematoda 

Nemertea Heteronemertea Lineidae Cerebratulus sp. 

Nemertea Heteronemertea Lineidae Linea sp. 1 

Total No. of Species 18 16 10 7 12 14 

Total No. of Individuals 51 44 23 21 38 44 
25 

Appendix Table 4.--Number of individuals of each infauna taxon collected per grab sample at 

Transects 7 and 8 at Two-headed Gully during June 1998. 

Phylum Order Family Species 7A 7B 7C 8A 8B 8C 

Annelida Capitallida Capitellidae Mediomastus californiensis 2 1 

Annelida Capitallida Capitellidae Notomastus giganteus 

Annelida Capitallida Maldanidae Praxillella gracilis 

Annelida Chaetopterida Chaetopteridae Spiochaetopterus costarum 

Annelida Cossurida Cossuridae Cossura soyeri 1 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 

Annelida Eunicida Lumbrineridae Lumbrineris luti 1 2 

Annelida Eunicida Onuphidae 1 1 1 

Annelida Magelonida Magelonidae Magelona berkeleyi 2 1 4 1 3 

Annelida Opheliida Opheliidae Travisia forbesii 

Annelida Opheliida Scalibregmidae Hyboscolex pacificus 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 1 5 

Annelida Orbiniida Paraonidae Aricidea lopezi 1 

Annelida Orbiniida Paraonidae Chaetozone sp. 2 2 2 3 6 

Annelida Orbiniida Paraonidae Levinsenia gracilis 1 1 

Annelida Phyllodocida Glyceridae Hemipodus borealis 

Annelida Phyllodocida Goniadidae Goniada brunnea 

Annelida Phyllodocida Hesionidae 

Annelida Phyllodocida Nephtidae Nephtys caeca 2 2 1 5 6 

Annelida Phyllodocida Nephtidae Nephtys cornuta 3 2 

Annelida Phyllodocida Nereididae Cyclonereis clyclurus 

Annelida Phyllodocida Pholodidae Pholoe glabra 1 

Annelida Sabellida Sabellidae 1 

Annelida Spionida Spionidae Prionospio lighti 

Annelida Spionida Spionidae Prionospio steenstrupi 6 8 1 3 5 

Annelida Spionida Spionidae Pygospio elegans 1 

Annelida Spionida Spionidae Spio cirrifera 1 

Annelida Sternapsida Sternapsidae Sternapsis scutata 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 1 
26


Annelida Terebellida Terebellidae Artacama conifera 2 1 

Annelida Terebellida Trichobranchidae Terebellides stroemi 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 1 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 

Arthropoda Amphipoda Phoxocephalidae Heterophoxus sp. 1 

Arthropoda Calanoida 

Arthropoda Cumacea 1 

Arthropoda Ostracoda Species I 

Echinodermata Dendrochiridotida Phyllophoridae Pentamera sp. 

Echiura Echiuroinea Echiuridae Echiurus echiurus alaskensis 

Mollusca Cephalaspidea Atyidae Haminoea virescens 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 

Mollusca Mesogastropoda Naticidae Euspira pallida 1 

Mollusca Neogastropoda Neptuneidae Colus halli 

Mollusca Nuculoida Nuculanidae Nuculana fossa 

Mollusca Nuculoida Nuculanidae Nuculana minuta 1 

Mollusca Nuculoida Nuculidae Ennucula tenuis 1 1 

Mollusca Nuculoida Yoldiidae Yoldia scissurata 

Mollusca Pholadomyoida Pandoridae Pandora bilirata 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 2 1 

Mollusca Veneroida Tellinidae Macoma eliminata 3 3 3 2 

Mollusca Veneroida Thyasiridae Axinospida serricata 8 2 10 10 10 1 

Nematoda 

Nemertea Heteronemertea Lineidae Cerebratulus sp. 

Nemertea Heteronemertea Lineidae Linea sp. 3 1 1 2 

Total No. of Species 14 6 10 12 13 10 

Total No. of Individuals 36 16 26 29 34 25 
27 

Appendix Table 4.--Number of individuals of each infauna taxa collected per grab sample at 

Transects 9 and 10 at Two-headed Gully during June 1998. 

Phylum Order Family Species 9A 9B 9C 10A 10B 10C 

Annelida Capitallida Capitellidae Mediomastus californiensis 1 

Annelida Capitallida Capitellidae Notomastus giganteus 1 

Annelida Capitallida Maldanidae Praxillella gracilis 

Annelida Chaetopterida Chaetopteridae Spiochaetopterus costarum 

Annelida Cossurida Cossuridae Cossura soyeri 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 

Annelida Eunicida Lumbrineridae Lumbrineris luti 2 5 

Annelida Eunicida Onuphidae 1 

Annelida Magelonida Magelonidae Magelona berkeleyi 1 1 

Annelida Opheliida Opheliidae Travisia forbesii 

Annelida Opheliida Scalibregmidae Hyboscolex pacificus 1 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 1 

Annelida Orbiniida Paraonidae Aricidea lopezi 

Annelida Orbiniida Paraonidae Chaetozone sp. 5 5 

Annelida Orbiniida Paraonidae Levinsenia gracilis 2 

Annelida Phyllodocida Glyceridae Hemipodus borealis 1 

Annelida Phyllodocida Goniadidae Goniada brunnea 

Annelida Phyllodocida Hesionidae 1 

Annelida Phyllodocida Nephtidae Nephtys caeca 1 4 3 6 2 

Annelida Phyllodocida Nephtidae Nephtys cornuta 1 

Annelida Phyllodocida Nereididae Cyclonereis clyclurus 1 

Annelida Phyllodocida Pholodidae Pholoe glabra 

Annelida Sabellida Sabellidae 

Annelida Spionida Spionidae Prionospio lighti 1 

Annelida Spionida Spionidae Prionospio steenstrupi 3 9 3 7 1 

Annelida Spionida Spionidae Pygospio elegans 1 

Annelida Spionida Spionidae Spio cirrifera 

Annelida Sternapsida Sternapsidae Sternapsis scutata 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 1 
28


Annelida Terebellida Terebellidae Artacama conifera 1 

Annelida Terebellida Trichobranchidae Terebellides stroemi 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 1 2 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 1 1 

Arthropoda Amphipoda Phoxocephalidae Heterophoxus sp. 

Arthropoda Calanoida 

Arthropoda Cumacea 3 1 1 1 

Arthropoda Ostracoda Species I 

Echinodermata Dendrochiridotida Phyllophoridae Pentamera sp. 

Echiura Echiuroinea Echiuridae Echiurus echiurus alaskensis 1 

Mollusca Cephalaspidea Atyidae Haminoea virescens 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 1 

Mollusca Mesogastropoda Naticidae Euspira pallida 

Mollusca Neogastropoda Neptuneidae Colus halli 

Mollusca Nuculoida Nuculanidae Nuculana fossa 

Mollusca Nuculoida Nuculanidae Nuculana minuta 

Mollusca Nuculoida Nuculidae Ennucula tenuis 1 

Mollusca Nuculoida Yoldiidae Yoldia scissurata 

Mollusca Pholadomyoida Pandoridae Pandora bilirata 1 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 

Mollusca Veneroida Tellinidae Macoma eliminata 1 1 

Mollusca Veneroida Thyasiridae Axinospida serricata 3 5 2 6 1 

Nematoda 1 

Nemertea Heteronemertea Lineidae Cerebratulus sp. 1 

Nemertea Heteronemertea Lineidae Linea sp. 

Total No. of Species 7 10 7 5 12 10 

Total No. of Individuals 13 23 17 18 25 13 
29 

Appendix Table 5.--Number of individuals of each infauna taxon collected per grab sample at 

Transect 1 in Chiniak Gully during August 1999. 

Phylum Order Family Species 1A 1B 1C 1D 1E 

Annelida Capitallida Capitellidae Mediomastus californiensis 2 12 

Annelida Capitallida Capitellidae Notomastus giganteus 

Annelida Capitallida Maldanidae 

Annelida Capitallida Maldanidae Axiothella rubrocincta 

Annelida Capitallida Maldanidae Clymenella sp. 

Annelida Capitallida Maldanidae Euclymene zonalis 3 

Annelida Capitallida Maldanidae Praxillella gracilis 

Annelida Chaetopterida Chaetopteridae Spiochaetopterus costarum 

Annelida Cossurida Cossuridae Cossura soyeri 

Annelida Eunicida Lumbrineridae Lumbrineris sp. 3 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 

Annelida Eunicida Lumbrineridae Lumbrineris inflata 

Annelida Eunicida Lumbrineridae Lumbrineris luti 3 2 12 16 

Annelida Eunicida Onuphidae Onuphis iridescens 1 1 

Annelida Magelonida Magelonidae Magelona berkeleyi 

Annelida Opheliida Opheliidae Ophelina sp. 

Annelida Opheliida Opheliidae Travisia forbesii 

Annelida Opheliida Scalibregmidae Scalibregma inflatum 1 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 1 2 4 5 

Annelida Orbiniida Paraonidae Aricidea lopezi 2 

Annelida Orbiniida Paraonidae Chaetozone sp. 2 

Annelida Orbiniida Paraonidae Levinsenia gracilis 

Annelida Oweniida Oweniidae Owenia cf. fusiformis 

Annelida Phyllodocida Glyceridae Glycera sp. 1 

Annelida Phyllodocida Glyceridae Hemipodus borealis 1 1 1 1 

Annelida Phyllodocida Goniadidae Goniada brunnea 3 2 

Annelida Phyllodocida Nephtidae Nephtys caeca 1 3 2 2 

Annelida Phyllodocida Nephtidae Nephtys cornuta 7 1 

Annelida Phyllodocida Pholodidae Pholoe glabra 4 2 2 
30


Annelida Phyllodocida Phyllodocidae Eteone longa 

Annelida Phyllodocida Phyllodocidae Phyllodocida sp. 

Annelida Phyllodocida Polynoidae Hesperonoe sp. 

Annelida Phyllodocida Spaerodoropsidae Spaerodoropsis sp. 1 

Annelida Phyllodocida Syllidae Syllis sp. 

Annelida Sabellida Sabellidae 1 

Annelida Sabellida Sabellidae Chone sp. 

Annelida Spionida Spionidae Dipolydora socialis 1 

Annelida Spionida Spionidae Prionospio lighti 1 

Annelida Spionida Spionidae Prionospio steenstrupi 3 

Annelida Spionida Spionidae Pygospio elegans 

Annelida Spionida Spionidae Spio cirrifera 1 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 3 

Annelida Terebellida Pectinaridae Pectinaria moorei 1 

Annelida Terebellida Terebellidae Amphitrite cirrata 

Annelida Terebellida Terebellidae Scionella sp. 

Annelida Terebellida Trichobranchidae Terebellides stroemi 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 2 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 1 

Arthropoda Amphipoda Ischyroceridae 1 6 

Arthropoda Amphipoda Phoxocephalidae Heterophoxus sp. 5 

Arthropoda Cumacea Diastylopsidae Diastylopsis dawsoni 16 375 75 296 

Arthropoda Cumacea Leuconiidae Leptocuma sp. 

Arthropoda Cumacea Leuconiidae Leucon magnadentata 1 

Arthropoda Decapoda Pinnixidae Pinnixa sp. 4 4 3 

Arthropoda Pantopoda Ammotheidae Achelia sp. 1 

Arthropoda Pantopoda Nymphonidae Nymphon sp. 

Chordata Stolidobranchia Mogulidae 

Echinodermata Apodida Chiridotidae 

Echinodermata Clypeasteroida Dendrasteridae Echinarchinus parma 1 

Echinodermata Dendrochiridotida Phyllophoridae Pentamera sp. 1 

Echinodermata Dendrochiridotida Sclerodactylidae Eupentacta sp. 

Echinodermata Molpadiida Molpadiidae Molpadia intermedia 
31


Echinodermata Ophiurida Amphiuridae Amphiodia sp. 21 27 2 24 24 

Echiura Echiuroinea Echiuridae Echiurus echiurus alaskensis 1 

Mollusca Cephalaspidea Atyidae Haminoea virescens 

Mollusca Cephalaspidea Cylichnidae Acteocina harpa 1 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 1 

Mollusca Mesogastropoda Eulimidae Eulima sp. 

Mollusca Mesogastropoda Naticidae Euspira pallida 

Mollusca Myoida Hiatellidae Hiatella arctica 

Mollusca Neogastropoda Amphissidae Alia gausapata 

Mollusca Neogastropoda Turridae Oenopota rosea 2 

Mollusca Nuculoida Nuculidae Ennucula tenuis 2 

Mollusca Nuculoida Yoldiidae Yoldia hyperborea 1 2 1 4 1 

Mollusca Nuculoida Yoldiidae Yoldia seminuda 

Mollusca Pholadomyoida Lyonsiidae Lyonsia californica 

Mollusca Pholadomyoida Pandoridae Pandora filosa 1 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 

Mollusca Veneroida Cultellidae Siliqua alta 2 10 4 

Mollusca Veneroida Mactridae Spisula polynema 

Mollusca Veneroida Tellinidae Macoma eliminata 1 2 2 1 

Mollusca Veneroida Tellinidae Macoma lipara 32 

Mollusca Veneroida Tellinidae Macoma moesta 1 1 2 1 

Mollusca Veneroida Thyasiridae Axinospida serricata 11 8 39 29 

Mollusca Veneroida Veneridae Compsomyax subdiaphana 

Mollusca Veneroida Veneridae Nutricola lordi 9 

Nemertea Heteronemertea Lineidae Cerebratulus sp. 

Nemertea Heteronemertea Lineidae Linea sp. 2 1 2 

Total No. of Species 10 17 33 17 16 

Total No. of Individuals 60 458 98 189 379 
32 

Appendix Table 5.--Number of individuals of each infauna taxon collected per grab sample at 

Transect 2 in Chiniak Gully during August 1999. 

Phylum Order Family Species 2A 2B 2C 2D 2E 2F 

Annelida Capitallida Capitellidae Mediomastus californiensis 1 1 

Annelida Capitallida Capitellidae Notomastus giganteus 

Annelida Capitallida Maldanidae 2 

Annelida Capitallida Maldanidae Axiothella rubrocincta 

Annelida Capitallida Maldanidae Clymenella sp. 

Annelida Capitallida Maldanidae Euclymene zonalis 

Annelida Capitallida Maldanidae Praxillella gracilis 

Annelida Chaetopterida Chaetopteridae Spiochaetopterus costarum 1 

Annelida Cossurida Cossuridae Cossura soyeri 

Annelida Eunicida Lumbrineridae Lumbrineris sp. 1 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 

Annelida Eunicida Lumbrineridae Lumbrineris inflata 

Annelida Eunicida Lumbrineridae Lumbrineris luti 6 2 11 

Annelida Eunicida Onuphidae Onuphis iridescens 

Annelida Magelonida Magelonidae Magelona berkeleyi 1 2 

Annelida Opheliida Opheliidae Ophelina sp. 

Annelida Opheliida Opheliidae Travisia forbesii 

Annelida Opheliida Scalibregmidae Scalibregma inflatum 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 4 2 7 9 5 7 

Annelida Orbiniida Paraonidae Aricidea lopezi 

Annelida Orbiniida Paraonidae Chaetozone sp. 

Annelida Orbiniida Paraonidae Levinsenia gracilis 

Annelida Oweniida Oweniidae Owenia cf. fusiformis 

Annelida Phyllodocida Glyceridae Glycera sp. 

Annelida Phyllodocida Glyceridae Hemipodus borealis 

Annelida Phyllodocida Goniadidae Goniada brunnea 

Annelida Phyllodocida Nephtidae Nephtys caeca 2 4 2 1 1 4 

Annelida Phyllodocida Nephtidae Nephtys cornuta 3 

Annelida Phyllodocida Pholodidae Pholoe glabra 4 1 1 
33


Annelida Phyllodocida Phyllodocidae Eteone longa 1 

Annelida Phyllodocida Phyllodocidae Phyllodocida sp. 

Annelida Phyllodocida Polynoidae Hesperonoe sp. 1 1 

Annelida Phyllodocida Spaerodoropsidae Spaerodoropsis sp. 

Annelida Phyllodocida Syllidae Syllis sp. 1 

Annelida Sabellida Sabellidae 1 

Annelida Sabellida Sabellidae Chone sp. 

Annelida Spionida Spionidae Dipolydora socialis 

Annelida Spionida Spionidae Prionospio lighti 

Annelida Spionida Spionidae Prionospio steenstrupi 1 

Annelida Spionida Spionidae Pygospio elegans 

Annelida Spionida Spionidae Spio cirrifera 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 1 1 

Annelida Terebellida Pectinaridae Pectinaria moorei 

Annelida Terebellida Terebellidae Amphitrite cirrata 

Annelida Terebellida Terebellidae Scionella sp. 

Annelida Terebellida Trichobranchidae Terebellides stroemi 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 4 2 2 3 1 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 

Arthropoda Amphipoda Ischyroceridae 2 2 1 7 

Arthropoda Amphipoda Phoxocephalidae Heterophoxus sp. 1 

Arthropoda Cumacea Diastylopsidae Diastylopsis dawsoni 1 3 1 30 

Arthropoda Cumacea Leuconiidae Leptocuma sp. 

Arthropoda Cumacea Leuconiidae Leucon magnadentata 

Arthropoda Decapoda Pinnixidae Pinnixa sp. 3 9 4 5 3 

Arthropoda Pantopoda Ammotheidae Achelia sp. 

Arthropoda Pantopoda Nymphonidae Nymphon sp. 1 

Chordata Stolidobranchia Mogulidae 1 

Echinodermata Apodida Chiridotidae 1 

Echinodermata Clypeasteroida Dendrasteridae Echinarchinus parma 

Echinodermata Dendrochiridotida Phyllophoridae Pentamera sp. 1 

Echinodermata Dendrochiridotida Sclerodactylidae Eupentacta sp. 

Echinodermata Molpadiida Molpadiidae Molpadia intermedia 
34


Echinodermata Ophiurida Amphiuridae Amphiodia sp. 44 24 17 17 15 28 

Echiura Echiuroinea Echiuridae Echiurus echiurus alaskensis 

Mollusca Cephalaspidea Atyidae Haminoea virescens 

Mollusca Cephalaspidea Cylichnidae Acteocina harpa 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 

Mollusca Mesogastropoda Eulimidae Eulima sp. 

Mollusca Mesogastropoda Naticidae Euspira pallida 

Mollusca Myoida Hiatellidae Hiatella arctica 

Mollusca Neogastropoda Amphissidae Alia gausapata 

Mollusca Neogastropoda Turridae Oenopota rosea 1 2 1 

Mollusca Nuculoida Nuculidae Ennucula tenuis 2 3 

Mollusca Nuculoida Yoldiidae Yoldia hyperborea 1 2 1 2 

Mollusca Nuculoida Yoldiidae Yoldia seminuda 1 

Mollusca Pholadomyoida Lyonsiidae Lyonsia californica 

Mollusca Pholadomyoida Pandoridae Pandora filosa 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 

Mollusca Veneroida Cultellidae Siliqua alta 5 3 8 4 8 

Mollusca Veneroida Mactridae Spisula polynema 1 1 

Mollusca Veneroida Tellinidae Macoma eliminata 1 2 3 

Mollusca Veneroida Tellinidae Macoma lipara 1 

Mollusca Veneroida Tellinidae Macoma moesta 2 3 1 

Mollusca Veneroida Thyasiridae Axinospida serricata 31 24 34 6 6 41 

Mollusca Veneroida Veneridae Compsomyax subdiaphana 

Mollusca Veneroida Veneridae Nutricola lordi 

Nemertea Heteronemertea Lineidae Cerebratulus sp. 

Nemertea Heteronemertea Lineidae Linea sp. 3 1 

Total No. of Species 16 11 21 15 10 22 

Total No. of Individuals 108 72 100 56 38 157 
35 

Appendix Table 5.--Number of individuals of each infauna taxon collected per grab sample at 

Transect 3 in Chiniak Gully during August 1999. 

Phylum Order Family Species 3A 3B 3C 3D 3F 

Annelida Capitallida Capitellidae Mediomastus californiensis 3 1 3 2 

Annelida Capitallida Capitellidae Notomastus giganteus 

Annelida Capitallida Maldanidae 1 

Annelida Capitallida Maldanidae Axiothella rubrocincta 

Annelida Capitallida Maldanidae Clymenella sp. 2 

Annelida Capitallida Maldanidae Euclymene zonalis 2 3 5 3 

Annelida Capitallida Maldanidae Praxillella gracilis 

Annelida Chaetopterida Chaetopteridae Spiochaetopterus costarum 

Annelida Cossurida Cossuridae Cossura soyeri 

Annelida Eunicida Lumbrineridae Lumbrineris sp. 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 1 

Annelida Eunicida Lumbrineridae Lumbrineris inflata 

Annelida Eunicida Lumbrineridae Lumbrineris luti 7 7 1 8 1 

Annelida Eunicida Onuphidae Onuphis iridescens 2 1 

Annelida Magelonida Magelonidae Magelona berkeleyi 

Annelida Opheliida Opheliidae Ophelina sp. 

Annelida Opheliida Opheliidae Travisia forbesii 

Annelida Opheliida Scalibregmidae Scalibregma inflatum 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 6 5 6 1 1 

Annelida Orbiniida Paraonidae Aricidea lopezi 

Annelida Orbiniida Paraonidae Chaetozone sp. 

Annelida Orbiniida Paraonidae Levinsenia gracilis 

Annelida Oweniida Oweniidae Owenia cf. fusiformis 1 

Annelida Phyllodocida Glyceridae Glycera sp. 

Annelida Phyllodocida Glyceridae Hemipodus borealis 

Annelida Phyllodocida Goniadidae Goniada brunnea 2 2 2 3 2 

Annelida Phyllodocida Nephtidae Nephtys caeca 6 4 

Annelida Phyllodocida Nephtidae Nephtys cornuta 2 1 

Annelida Phyllodocida Pholodidae Pholoe glabra 1 1 2 
36


Annelida Phyllodocida Phyllodocidae Eteone longa 

Annelida Phyllodocida Phyllodocidae Phyllodocida sp. 

Annelida Phyllodocida Polynoidae Hesperonoe sp. 

Annelida Phyllodocida Spaerodoropsidae Spaerodoropsis sp. 

Annelida Phyllodocida Syllidae Syllis sp. 1 3 

Annelida Sabellida Sabellidae 

Annelida Sabellida Sabellidae Chone sp. 

Annelida Spionida Spionidae Dipolydora socialis 

Annelida Spionida Spionidae Prionospio lighti 

Annelida Spionida Spionidae Prionospio steenstrupi 2 1 

Annelida Spionida Spionidae Pygospio elegans 

Annelida Spionida Spionidae Spio cirrifera 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 2 2 

Annelida Terebellida Pectinaridae Pectinaria moorei 1 

Annelida Terebellida Terebellidae Amphitrite cirrata 

Annelida Terebellida Terebellidae Scionella sp. 

Annelida Terebellida Trichobranchidae Terebellides stroemi 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 2 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 

Arthropoda Amphipoda Ischyroceridae 1 

Arthropoda Amphipoda Phoxocephalidae Heterophoxus sp. 3 

Arthropoda Cumacea Diastylopsidae Diastylopsis dawsoni 3 1 1 213 

Arthropoda Cumacea Leuconiidae Leptocuma sp. 

Arthropoda Cumacea Leuconiidae Leucon magnadentata 1 

Arthropoda Decapoda Pinnixidae Pinnixa sp. 7 3 1 3 

Arthropoda Pantopoda Ammotheidae Achelia sp. 

Arthropoda Pantopoda Nymphonidae Nymphon sp. 

Chordata Stolidobranchia Mogulidae 1 1 1 5 

Echinodermata Apodida Chiridotidae 1 

Echinodermata Clypeasteroida Dendrasteridae Echinarchinus parma 1 1 

Echinodermata Dendrochiridotida Phyllophoridae Pentamera sp. 1 2 

Echinodermata Dendrochiridotida Sclerodactylidae Eupentacta sp. 1 

Echinodermata Molpadiida Molpadiidae Molpadia intermedia 
37


Echinodermata Ophiurida Amphiuridae Amphiodia sp. 22 53 19 36 45 

Echiura Echiuroinea Echiuridae Echiurus echiurus alaskensis 

Mollusca Cephalaspidea Atyidae Haminoea virescens 

Mollusca Cephalaspidea Cylichnidae Acteocina harpa 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 1 

Mollusca Mesogastropoda Eulimidae Eulima sp. 

Mollusca Mesogastropoda Naticidae Euspira pallida 1 

Mollusca Myoida Hiatellidae Hiatella arctica 

Mollusca Neogastropoda Amphissidae Alia gausapata 

Mollusca Neogastropoda Turridae Oenopota rosea 2 

Mollusca Nuculoida Nuculidae Ennucula tenuis 

Mollusca Nuculoida Yoldiidae Yoldia hyperborea 1 1 1 4 6 

Mollusca Nuculoida Yoldiidae Yoldia seminuda 

Mollusca Pholadomyoida Lyonsiidae Lyonsia californica 

Mollusca Pholadomyoida Pandoridae Pandora filosa 1 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 1 

Mollusca Veneroida Cultellidae Siliqua alta 4 6 4 12 7 

Mollusca Veneroida Mactridae Spisula polynema 

Mollusca Veneroida Tellinidae Macoma eliminata 1 2 

Mollusca Veneroida Tellinidae Macoma lipara 

Mollusca Veneroida Tellinidae Macoma moesta 1 1 1 1 

Mollusca Veneroida Thyasiridae Axinospida serricata 17 18 6 32 24 

Mollusca Veneroida Veneridae Compsomyax subdiaphana 1 

Mollusca Veneroida Veneridae Nutricola lordi 2 

Nemertea Heteronemertea Lineidae Cerebratulus sp. 

Nemertea Heteronemertea Lineidae Linea sp. 1 1 1 1 

Total No. of Species 21 22 20 23 16 

Total No. of Individuals 85 120 62 127 311 
38 

Appendix Table 5.--Number of individuals of each infauna taxon collected per grab sample at 

Transect 4 in Chiniak Gully during August 1999. 

Phylum Order Family Species 4A 4B 4C 4D 4F 

Annelida Capitallida Capitellidae Mediomastus californiensis 3 2 3 

Annelida Capitallida Capitellidae Notomastus giganteus 

Annelida Capitallida Maldanidae 1 1 

Annelida Capitallida Maldanidae Axiothella rubrocincta 

Annelida Capitallida Maldanidae Clymenella sp. 

Annelida Capitallida Maldanidae Euclymene zonalis 1 1 

Annelida Capitallida Maldanidae Praxillella gracilis 

Annelida Chaetopterida Chaetopteridae Spiochaetopterus costarum 

Annelida Cossurida Cossuridae Cossura soyeri 

Annelida Eunicida Lumbrineridae Lumbrineris sp. 3 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 1 

Annelida Eunicida Lumbrineridae Lumbrineris inflata 

Annelida Eunicida Lumbrineridae Lumbrineris luti 7 18 14 4 5 

Annelida Eunicida Onuphidae Onuphis iridescens 1 1 

Annelida Magelonida Magelonidae Magelona berkeleyi 1 

Annelida Opheliida Opheliidae Ophelina sp. 

Annelida Opheliida Opheliidae Travisia forbesii 

Annelida Opheliida Scalibregmidae Scalibregma inflatum 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 4 1 7 11 6 

Annelida Orbiniida Paraonidae Aricidea lopezi 

Annelida Orbiniida Paraonidae Chaetozone sp. 

Annelida Orbiniida Paraonidae Levinsenia gracilis 

Annelida Oweniida Oweniidae Owenia cf. fusiformis 1 

Annelida Phyllodocida Glyceridae Glycera sp. 

Annelida Phyllodocida Glyceridae Hemipodus borealis 2 1 1 1 

Annelida Phyllodocida Goniadidae Goniada brunnea 1 1 3 3 

Annelida Phyllodocida Nephtidae Nephtys caeca 2 1 7 6 

Annelida Phyllodocida Nephtidae Nephtys cornuta 3 2 

Annelida Phyllodocida Pholodidae Pholoe glabra 1 2 1 
39


Annelida Phyllodocida Phyllodocidae Eteone longa 

Annelida Phyllodocida Phyllodocidae Phyllodocida sp. 

Annelida Phyllodocida Polynoidae Hesperonoe sp. 

Annelida Phyllodocida Spaerodoropsidae Spaerodoropsis sp. 

Annelida Phyllodocida Syllidae Syllis sp. 2 1 1 

Annelida Sabellida Sabellidae 1 

Annelida Sabellida Sabellidae Chone sp. 

Annelida Spionida Spionidae Dipolydora socialis 

Annelida Spionida Spionidae Prionospio lighti 2 

Annelida Spionida Spionidae Prionospio steenstrupi 3 1 

Annelida Spionida Spionidae Pygospio elegans 

Annelida Spionida Spionidae Spio cirrifera 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 4 1 

Annelida Terebellida Pectinaridae Pectinaria moorei 

Annelida Terebellida Terebellidae Amphitrite cirrata 

Annelida Terebellida Terebellidae Scionella sp. 

Annelida Terebellida Trichobranchidae Terebellides stroemi 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 1 2 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 

Arthropoda Amphipoda Ischyroceridae 2 2 1 

Arthropoda Amphipoda Phoxocephalidae Heterophoxus sp. 1 1 

Arthropoda Cumacea Diastylopsidae Diastylopsis dawsoni 5 148 1 

Arthropoda Cumacea Leuconiidae Leptocuma sp. 

Arthropoda Cumacea Leuconiidae Leucon magnadentata 1 

Arthropoda Decapoda Pinnixidae Pinnixa sp. 2 1 4 

Arthropoda Pantopoda Ammotheidae Achelia sp. 

Arthropoda Pantopoda Nymphonidae Nymphon sp. 

Chordata Stolidobranchia Mogulidae 1 5 

Echinodermata Apodida Chiridotidae 

Echinodermata Clypeasteroida Dendrasteridae Echinarchinus parma 

Echinodermata Dendrochiridotida Phyllophoridae Pentamera sp. 

Echinodermata Dendrochiridotida Sclerodactylidae Eupentacta sp. 

Echinodermata Molpadiida Molpadiidae Molpadia intermedia 
40


Echinodermata Ophiurida Amphiuridae Amphiodia sp. 24 33 23 28 14 

Echiura Echiuroinea Echiuridae Echiurus echiurus alaskensis 

Mollusca Cephalaspidea Atyidae Haminoea virescens 

Mollusca Cephalaspidea Cylichnidae Acteocina harpa 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 2 

Mollusca Mesogastropoda Eulimidae Eulima sp. 1 

Mollusca Mesogastropoda Naticidae Euspira pallida 1 

Mollusca Myoida Hiatellidae Hiatella arctica 

Mollusca Neogastropoda Amphissidae Alia gausapata 

Mollusca Neogastropoda Turridae Oenopota rosea 

Mollusca Nuculoida Nuculidae Ennucula tenuis 1 1 

Mollusca Nuculoida Yoldiidae Yoldia hyperborea 4 3 1 

Mollusca Nuculoida Yoldiidae Yoldia seminuda 

Mollusca Pholadomyoida Lyonsiidae Lyonsia californica 

Mollusca Pholadomyoida Pandoridae Pandora filosa 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 1 

Mollusca Veneroida Cultellidae Siliqua alta 4 3 1 3 

Mollusca Veneroida Mactridae Spisula polynema 

Mollusca Veneroida Tellinidae Macoma eliminata 

Mollusca Veneroida Tellinidae Macoma lipara 

Mollusca Veneroida Tellinidae Macoma moesta 2 1 

Mollusca Veneroida Thyasiridae Axinospida serricata 6 12 7 15 

Mollusca Veneroida Veneridae Compsomyax subdiaphana 

Mollusca Veneroida Veneridae Nutricola lordi 

Nemertea Heteronemertea Lineidae Cerebratulus sp. 1 

Nemertea Heteronemertea Lineidae Linea sp. 1 1 

Total No. of Species 21 21 22 13 15 

Total No. of Individuals 78 93 88 210 54 
41 

Appendix Table 5.--Number of individuals of each infauna taxon collected per grab sample at 

Transect 5 in Chiniak Gully during August 1999. 

Phylum Order Family Species 5A 5B 5C 5D 5F 

Annelida Capitallida Capitellidae Mediomastus californiensis 1 10 10 

Annelida Capitallida Capitellidae Notomastus giganteus 1 1 

Annelida Capitallida Maldanidae 

Annelida Capitallida Maldanidae Axiothella rubrocincta 

Annelida Capitallida Maldanidae Clymenella sp. 

Annelida Capitallida Maldanidae Euclymene zonalis 5 5 4 2 5 

Annelida Capitallida Maldanidae Praxillella gracilis 1 

Annelida Chaetopterida Chaetopteridae Spiochaetopterus costarum 1 

Annelida Cossurida Cossuridae Cossura soyeri 

Annelida Eunicida Lumbrineridae Lumbrineris sp. 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 

Annelida Eunicida Lumbrineridae Lumbrineris inflata 

Annelida Eunicida Lumbrineridae Lumbrineris luti 1 6 11 13 13 

Annelida Eunicida Onuphidae Onuphis iridescens 2 

Annelida Magelonida Magelonidae Magelona berkeleyi 2 

Annelida Opheliida Opheliidae Ophelina sp. 1 

Annelida Opheliida Opheliidae Travisia forbesii 1 1 

Annelida Opheliida Scalibregmidae Scalibregma inflatum 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 2 3 2 6 

Annelida Orbiniida Paraonidae Aricidea lopezi 1 

Annelida Orbiniida Paraonidae Chaetozone sp. 

Annelida Orbiniida Paraonidae Levinsenia gracilis 

Annelida Oweniida Oweniidae Owenia cf. fusiformis 1 

Annelida Phyllodocida Glyceridae Glycera sp. 

Annelida Phyllodocida Glyceridae Hemipodus borealis 1 1 

Annelida Phyllodocida Goniadidae Goniada brunnea 2 3 2 1 

Annelida Phyllodocida Nephtidae Nephtys caeca 4 5 8 7 6 

Annelida Phyllodocida Nephtidae Nephtys cornuta 2 1 1 1 

Annelida Phyllodocida Pholodidae Pholoe glabra 1 
42


Annelida Phyllodocida Phyllodocidae Eteone longa 

Annelida Phyllodocida Phyllodocidae Phyllodocida sp. 

Annelida Phyllodocida Polynoidae Hesperonoe sp. 

Annelida Phyllodocida Spaerodoropsidae Spaerodoropsis sp. 

Annelida Phyllodocida Syllidae Syllis sp. 2 

Annelida Sabellida Sabellidae 

Annelida Sabellida Sabellidae Chone sp. 

Annelida Spionida Spionidae Dipolydora socialis 

Annelida Spionida Spionidae Prionospio lighti 

Annelida Spionida Spionidae Prionospio steenstrupi 1 1 2 

Annelida Spionida Spionidae Pygospio elegans 

Annelida Spionida Spionidae Spio cirrifera 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 1 2 2 1 1 

Annelida Terebellida Pectinaridae Pectinaria moorei 

Annelida Terebellida Terebellidae Amphitrite cirrata 

Annelida Terebellida Terebellidae Scionella sp. 

Annelida Terebellida Trichobranchidae Terebellides stroemi 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 4 1 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 1 1 

Arthropoda Amphipoda Ischyroceridae 1 2 

Arthropoda Amphipoda Phoxocephalidae Heterophoxus sp. 1 2 2 1 1 

Arthropoda Cumacea Diastylopsidae Diastylopsis dawsoni 1 3 

Arthropoda Cumacea Leuconiidae Leptocuma sp. 

Arthropoda Cumacea Leuconiidae Leucon magnadentata 1 

Arthropoda Decapoda Pinnixidae Pinnixa sp. 4 1 1 

Arthropoda Pantopoda Ammotheidae Achelia sp. 

Arthropoda Pantopoda Nymphonidae Nymphon sp. 

Chordata Stolidobranchia Mogulidae 4 3 1 1 

Echinodermata Apodida Chiridotidae 1 

Echinodermata Clypeasteroida Dendrasteridae Echinarchinus parma 

Echinodermata Dendrochiridotida Phyllophoridae Pentamera sp. 

Echinodermata Dendrochiridotida Sclerodactylidae Eupentacta sp. 

Echinodermata Molpadiida Molpadiidae Molpadia intermedia 
43


Echinodermata Ophiurida Amphiuridae Amphiodia sp. 3 16 16 7 

Echiura Echiuroinea Echiuridae Echiurus echiurus alaskensis 

Mollusca Cephalaspidea Atyidae Haminoea virescens 

Mollusca Cephalaspidea Cylichnidae Acteocina harpa 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 

Mollusca Mesogastropoda Eulimidae Eulima sp. 

Mollusca Mesogastropoda Naticidae Euspira pallida 1 

Mollusca Myoida Hiatellidae Hiatella arctica 1 1 

Mollusca Neogastropoda Amphissidae Alia gausapata 

Mollusca Neogastropoda Turridae Oenopota rosea 1 

Mollusca Nuculoida Nuculidae Ennucula tenuis 

Mollusca Nuculoida Yoldiidae Yoldia hyperborea 1 2 2 1 

Mollusca Nuculoida Yoldiidae Yoldia seminuda 

Mollusca Pholadomyoida Lyonsiidae Lyonsia californica 1 

Mollusca Pholadomyoida Pandoridae Pandora filosa 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 1 1 

Mollusca Veneroida Cultellidae Siliqua alta 1 1 1 

Mollusca Veneroida Mactridae Spisula polynema 

Mollusca Veneroida Tellinidae Macoma eliminata 2 

Mollusca Veneroida Tellinidae Macoma lipara 

Mollusca Veneroida Tellinidae Macoma moesta 4 4 2 4 2 

Mollusca Veneroida Thyasiridae Axinospida serricata 4 3 4 17 

Mollusca Veneroida Veneridae Compsomyax subdiaphana 

Mollusca Veneroida Veneridae Nutricola lordi 

Nemertea Heteronemertea Lineidae Cerebratulus sp. 

Nemertea Heteronemertea Lineidae Linea sp. 1 1 3 

Total No. of Species 18 20 18 23 27 

Total No. of Individuals 36 63 61 70 88 
44 

Appendix Table 5.--Number of individuals of each infauna taxon collected per grab sample at 

Transect 6 in Chiniak Gully during August 1999. 

Phylum Order Family Species 6A 6B 6C 6D 6E 6F 

Annelida Capitallida Capitellidae Mediomastus californiensis 9 6 6 3 8 

Annelida Capitallida Capitellidae Notomastus giganteus 2 2 

Annelida Capitallida Maldanidae 1 1 1 

Annelida Capitallida Maldanidae Axiothella rubrocincta 

Annelida Capitallida Maldanidae Clymenella sp. 

Annelida Capitallida Maldanidae Euclymene zonalis 3 7 2 3 5 

Annelida Capitallida Maldanidae Praxillella gracilis 

Annelida Chaetopterida Chaetopteridae Spiochaetopterus costarum 1 1 

Annelida Cossurida Cossuridae Cossura soyeri 

Annelida Eunicida Lumbrineridae Lumbrineris sp. 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 1 

Annelida Eunicida Lumbrineridae Lumbrineris inflata 

Annelida Eunicida Lumbrineridae Lumbrineris luti 11 11 12 8 8 12 

Annelida Eunicida Onuphidae Onuphis iridescens 1 2 

Annelida Magelonida Magelonidae Magelona berkeleyi 2 2 4 

Annelida Opheliida Opheliidae Ophelina sp. 

Annelida Opheliida Opheliidae Travisia forbesii 1 

Annelida Opheliida Scalibregmidae Scalibregma inflatum 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 4 3 2 4 

Annelida Orbiniida Paraonidae Aricidea lopezi 3 1 

Annelida Orbiniida Paraonidae Chaetozone sp. 1 

Annelida Orbiniida Paraonidae Levinsenia gracilis 

Annelida Oweniida Oweniidae Owenia cf. fusiformis 5 

Annelida Phyllodocida Glyceridae Glycera sp. 

Annelida Phyllodocida Glyceridae Hemipodus borealis 2 1 1 1 1 

Annelida Phyllodocida Goniadidae Goniada brunnea 1 2 1 1 

Annelida Phyllodocida Nephtidae Nephtys caeca 3 7 6 7 3 6 

Annelida Phyllodocida Nephtidae Nephtys cornuta 1 3 2 2 1 3 

Annelida Phyllodocida Pholodidae Pholoe glabra 1 
45


Annelida Phyllodocida Phyllodocidae Eteone longa 

Annelida Phyllodocida Phyllodocidae Phyllodocida sp. 1 

Annelida Phyllodocida Polynoidae Hesperonoe sp. 

Annelida Phyllodocida Spaerodoropsidae Spaerodoropsis sp. 

Annelida Phyllodocida Syllidae Syllis sp. 1 

Annelida Sabellida Sabellidae 1 

Annelida Sabellida Sabellidae Chone sp. 

Annelida Spionida Spionidae Dipolydora socialis 1 

Annelida Spionida Spionidae Prionospio lighti 

Annelida Spionida Spionidae Prionospio steenstrupi 5 7 3 3 2 1 

Annelida Spionida Spionidae Pygospio elegans 

Annelida Spionida Spionidae Spio cirrifera 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 1 2 1 1 4 

Annelida Terebellida Pectinaridae Pectinaria moorei 1 

Annelida Terebellida Terebellidae Amphitrite cirrata 1 

Annelida Terebellida Terebellidae Scionella sp. 1 

Annelida Terebellida Trichobranchidae Terebellides stroemi 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 3 3 1 6 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 2 2 2 1 

Arthropoda Amphipoda Ischyroceridae 4 6 5 

Arthropoda Amphipoda Phoxocephalidae Heterophoxus sp. 2 2 7 

Arthropoda Cumacea Diastylopsidae Diastylopsis dawsoni 1 

Arthropoda Cumacea Leuconiidae Leptocuma sp. 

Arthropoda Cumacea Leuconiidae Leucon magnadentata 

Arthropoda Decapoda Pinnixidae Pinnixa sp. 1 2 

Arthropoda Pantopoda Ammotheidae Achelia sp. 

Arthropoda Pantopoda Nymphonidae Nymphon sp. 

Chordata Stolidobranchia Mogulidae 

Echinodermata Apodida Chiridotidae 

Echinodermata Clypeasteroida Dendrasteridae Echinarchinus parma 

Echinodermata Dendrochiridotida Phyllophoridae Pentamera sp. 

Echinodermata Dendrochiridotida Sclerodactylidae Eupentacta sp. 

Echinodermata Molpadiida Molpadiidae Molpadia intermedia 
46


Echinodermata Ophiurida Amphiuridae Amphiodia sp. 2 1 1 

Echiura Echiuroinea Echiuridae Echiurus echiurus alaskensis 1 1 

Mollusca Cephalaspidea Atyidae Haminoea virescens 

Mollusca Cephalaspidea Cylichnidae Acteocina harpa 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 2 

Mollusca Mesogastropoda Eulimidae Eulima sp. 

Mollusca Mesogastropoda Naticidae Euspira pallida 

Mollusca Myoida Hiatellidae Hiatella arctica 1 1 

Mollusca Neogastropoda Amphissidae Alia gausapata 

Mollusca Neogastropoda Turridae Oenopota rosea 3 

Mollusca Nuculoida Nuculidae Ennucula tenuis 

Mollusca Nuculoida Yoldiidae Yoldia hyperborea 3 

Mollusca Nuculoida Yoldiidae Yoldia seminuda 

Mollusca Pholadomyoida Lyonsiidae Lyonsia californica 

Mollusca Pholadomyoida Pandoridae Pandora filosa 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 

Mollusca Veneroida Cultellidae Siliqua alta 1 1 

Mollusca Veneroida Mactridae Spisula polynema 

Mollusca Veneroida Tellinidae Macoma eliminata 6 1 6 1 

Mollusca Veneroida Tellinidae Macoma lipara 

Mollusca Veneroida Tellinidae Macoma moesta 7 3 2 4 

Mollusca Veneroida Thyasiridae Axinospida serricata 8 2 3 8 3 3 

Mollusca Veneroida Veneridae Compsomyax subdiaphana 

Mollusca Veneroida Veneridae Nutricola lordi 1 

Nemertea Heteronemertea Lineidae Cerebratulus sp. 1 1 

Nemertea Heteronemertea Lineidae Linea sp. 1 4 2 

Total No. of Species 25 22 21 20 23 16 

Total No. of Individuals 79 77 63 60 56 52 
47 

Appendix Table 5.--Number of individuals of each infauna taxon collected per grab sample at 

Transect 7 in Chiniak Gully during August 1999. 

Phylum Order Family Species 7A 7B 7C 7D 7E 7F 

Annelida Capitallida Capitellidae Mediomastus californiensis 1 3 11 22 2 13 

Annelida Capitallida Capitellidae Notomastus giganteus 1 

Annelida Capitallida Maldanidae 

Annelida Capitallida Maldanidae Axiothella rubrocincta 1 

Annelida Capitallida Maldanidae Clymenella sp. 1 

Annelida Capitallida Maldanidae Euclymene zonalis 3 1 1 2 

Annelida Capitallida Maldanidae Praxillella gracilis 

Annelida Chaetopterida Chaetopteridae Spiochaetopterus costarum 

Annelida Cossurida Cossuridae Cossura soyeri 2 

Annelida Eunicida Lumbrineridae Lumbrineris sp. 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 

Annelida Eunicida Lumbrineridae Lumbrineris inflata 

Annelida Eunicida Lumbrineridae Lumbrineris luti 3 8 14 18 2 9 

Annelida Eunicida Onuphidae Onuphis iridescens 1 1 1 2 

Annelida Magelonida Magelonidae Magelona berkeleyi 1 2 

Annelida Opheliida Opheliidae Ophelina sp. 

Annelida Opheliida Opheliidae Travisia forbesii 1 1 

Annelida Opheliida Scalibregmidae Scalibregma inflatum 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 2 1 3 3 7 

Annelida Orbiniida Paraonidae Aricidea lopezi 2 1 4 2 

Annelida Orbiniida Paraonidae Chaetozone sp. 3 1 

Annelida Orbiniida Paraonidae Levinsenia gracilis 

Annelida Oweniida Oweniidae Owenia cf. fusiformis 2 3 

Annelida Phyllodocida Glyceridae Glycera sp. 

Annelida Phyllodocida Glyceridae Hemipodus borealis 1 2 

Annelida Phyllodocida Goniadidae Goniada brunnea 

Annelida Phyllodocida Nephtidae Nephtys caeca 5 7 2 6 1 

Annelida Phyllodocida Nephtidae Nephtys cornuta 2 1 3 2 

Annelida Phyllodocida Pholodidae Pholoe glabra 
48


Annelida Phyllodocida Phyllodocidae Eteone longa 

Annelida Phyllodocida Phyllodocidae Phyllodocida sp. 

Annelida Phyllodocida Polynoidae Hesperonoe sp. 

Annelida Phyllodocida Spaerodoropsidae Spaerodoropsis sp. 

Annelida Phyllodocida Syllidae Syllis sp. 1 3 1 1 

Annelida Sabellida Sabellidae 

Annelida Sabellida Sabellidae Chone sp. 1 

Annelida Spionida Spionidae Dipolydora socialis 

Annelida Spionida Spionidae Prionospio lighti 

Annelida Spionida Spionidae Prionospio steenstrupi 2 3 3 5 2 

Annelida Spionida Spionidae Pygospio elegans 

Annelida Spionida Spionidae Spio cirrifera 1 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 2 2 4 1 2 3 

Annelida Terebellida Pectinaridae Pectinaria moorei 

Annelida Terebellida Terebellidae Amphitrite cirrata 

Annelida Terebellida Terebellidae Scionella sp. 1 

Annelida Terebellida Trichobranchidae Terebellides stroemi 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 7 3 2 3 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 1 

Arthropoda Amphipoda Ischyroceridae 8 1 2 3 

Arthropoda Amphipoda Phoxocephalidae Heterophoxus sp. 1 1 6 1 1 1 

Arthropoda Cumacea Diastylopsidae Diastylopsis dawsoni 

Arthropoda Cumacea Leuconiidae Leptocuma sp. 

Arthropoda Cumacea Leuconiidae Leucon magnadentata 1 

Arthropoda Decapoda Pinnixidae Pinnixa sp. 1 

Arthropoda Pantopoda Ammotheidae Achelia sp. 

Arthropoda Pantopoda Nymphonidae Nymphon sp. 

Chordata Stolidobranchia Mogulidae 

Echinodermata Apodida Chiridotidae 

Echinodermata Clypeasteroida Dendrasteridae Echinarchinus parma 

Echinodermata Dendrochiridotida Phyllophoridae Pentamera sp. 

Echinodermata Dendrochiridotida Sclerodactylidae Eupentacta sp. 

Echinodermata Molpadiida Molpadiidae Molpadia intermedia 
49


Echinodermata Ophiurida Amphiuridae Amphiodia sp. 3 

Echiura Echiuroinea Echiuridae Echiurus echiurus alaskensis 

Mollusca Cephalaspidea Atyidae Haminoea virescens 

Mollusca Cephalaspidea Cylichnidae Acteocina harpa 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 

Mollusca Mesogastropoda Eulimidae Eulima sp. 

Mollusca Mesogastropoda Naticidae Euspira pallida 

Mollusca Myoida Hiatellidae Hiatella arctica 

Mollusca Neogastropoda Amphissidae Alia gausapata 

Mollusca Neogastropoda Turridae Oenopota rosea 

Mollusca Nuculoida Nuculidae Ennucula tenuis 1 1 

Mollusca Nuculoida Yoldiidae Yoldia hyperborea 3 1 1 1 

Mollusca Nuculoida Yoldiidae Yoldia seminuda 

Mollusca Pholadomyoida Lyonsiidae Lyonsia californica 

Mollusca Pholadomyoida Pandoridae Pandora filosa 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 

Mollusca Veneroida Cultellidae Siliqua alta 

Mollusca Veneroida Mactridae Spisula polynema 

Mollusca Veneroida Tellinidae Macoma eliminata 4 2 3 1 

Mollusca Veneroida Tellinidae Macoma lipara 

Mollusca Veneroida Tellinidae Macoma moesta 1 6 3 1 1 6 

Mollusca Veneroida Thyasiridae Axinospida serricata 11 6 12 5 7 13 

Mollusca Veneroida Veneridae Compsomyax subdiaphana 

Mollusca Veneroida Veneridae Nutricola lordi 1 

Nemertea Heteronemertea Lineidae Cerebratulus sp. 

Nemertea Heteronemertea Lineidae Linea sp. 1 1 

Total No. of Species 22 14 26 18 14 19 

Total No. of Individuals 65 48 79 76 30 73 
50 

Appendix Table 5.--Number of individuals of each infauna taxon collected per grab sample at 

Transect 8 in Chiniak Gully during August 1999. 

Phylum Order Family Species 8A 8B 8C 8D 8E 8F 

Annelida Capitallida Capitellidae Mediomastus californiensis 18 16 1 4 6 20 

Annelida Capitallida Capitellidae Notomastus giganteus 1 

Annelida Capitallida Maldanidae 1 

Annelida Capitallida Maldanidae Axiothella rubrocincta 

Annelida Capitallida Maldanidae Clymenella sp. 2 

Annelida Capitallida Maldanidae Euclymene zonalis 4 2 1 3 

Annelida Capitallida Maldanidae Praxillella gracilis 

Annelida Chaetopterida Chaetopteridae Spiochaetopterus costarum 1 

Annelida Cossurida Cossuridae Cossura soyeri 1 

Annelida Eunicida Lumbrineridae Lumbrineris sp. 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 1 

Annelida Eunicida Lumbrineridae Lumbrineris inflata 

Annelida Eunicida Lumbrineridae Lumbrineris luti 18 12 8 4 8 12 

Annelida Eunicida Onuphidae Onuphis iridescens 1 

Annelida Magelonida Magelonidae Magelona berkeleyi 1 3 1 1 5 

Annelida Opheliida Opheliidae Ophelina sp. 

Annelida Opheliida Opheliidae Travisia forbesii 

Annelida Opheliida Scalibregmidae Scalibregma inflatum 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 5 7 7 2 16 

Annelida Orbiniida Paraonidae Aricidea lopezi 2 1 1 2 4 

Annelida Orbiniida Paraonidae Chaetozone sp. 3 2 

Annelida Orbiniida Paraonidae Levinsenia gracilis 2 2 

Annelida Oweniida Oweniidae Owenia cf. fusiformis 3 1 

Annelida Phyllodocida Glyceridae Glycera sp. 1 

Annelida Phyllodocida Glyceridae Hemipodus borealis 1 2 2 2 

Annelida Phyllodocida Goniadidae Goniada brunnea 1 1 

Annelida Phyllodocida Nephtidae Nephtys caeca 8 9 1 1 9 1 

Annelida Phyllodocida Nephtidae Nephtys cornuta 1 2 1 3 

Annelida Phyllodocida Pholodidae Pholoe glabra 
51


Annelida Phyllodocida Phyllodocidae Eteone longa 1 1 

Annelida Phyllodocida Phyllodocidae Phyllodocida sp. 

Annelida Phyllodocida Polynoidae Hesperonoe sp. 

Annelida Phyllodocida Spaerodoropsidae Spaerodoropsis sp. 1 

Annelida Phyllodocida Syllidae Syllis sp. 1 2 

Annelida Sabellida Sabellidae 

Annelida Sabellida Sabellidae Chone sp. 

Annelida Spionida Spionidae Dipolydora socialis 2 

Annelida Spionida Spionidae Prionospio lighti 

Annelida Spionida Spionidae Prionospio steenstrupi 1 4 1 2 

Annelida Spionida Spionidae Pygospio elegans 1 

Annelida Spionida Spionidae Spio cirrifera 1 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 3 2 1 3 2 3 

Annelida Terebellida Pectinaridae Pectinaria moorei 

Annelida Terebellida Terebellidae Amphitrite cirrata 

Annelida Terebellida Terebellidae Scionella sp. 

Annelida Terebellida Trichobranchidae Terebellides stroemi 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 4 2 1 6 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 1 

Arthropoda Amphipoda Ischyroceridae 3 9 4 3 3 1 

Arthropoda Amphipoda Phoxocephalidae Heterophoxus sp. 2 2 2 

Arthropoda Cumacea Diastylopsidae Diastylopsis dawsoni 

Arthropoda Cumacea Leuconiidae Leptocuma sp. 

Arthropoda Cumacea Leuconiidae Leucon magnadentata 

Arthropoda Decapoda Pinnixidae Pinnixa sp. 

Arthropoda Pantopoda Ammotheidae Achelia sp. 

Arthropoda Pantopoda Nymphonidae Nymphon sp. 

Chordata Stolidobranchia Mogulidae 

Echinodermata Apodida Chiridotidae 1 

Echinodermata Clypeasteroida Dendrasteridae Echinarchinus parma 

Echinodermata Dendrochiridotida Phyllophoridae Pentamera sp. 1 1 

Echinodermata Dendrochiridotida Sclerodactylidae Eupentacta sp. 

Echinodermata Molpadiida Molpadiidae Molpadia intermedia 
52


Echinodermata Ophiurida Amphiuridae Amphiodia sp. 2 

Echiura Echiuroinea Echiuridae Echiurus echiurus alaskensis 

Mollusca Cephalaspidea Atyidae Haminoea virescens 1 1 

Mollusca Cephalaspidea Cylichnidae Acteocina harpa 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 1 

Mollusca Mesogastropoda Eulimidae Eulima sp. 

Mollusca Mesogastropoda Naticidae Euspira pallida 

Mollusca Myoida Hiatellidae Hiatella arctica 

Mollusca Neogastropoda Amphissidae Alia gausapata 

Mollusca Neogastropoda Turridae Oenopota rosea 

Mollusca Nuculoida Nuculidae Ennucula tenuis 

Mollusca Nuculoida Yoldiidae Yoldia hyperborea 2 2 

Mollusca Nuculoida Yoldiidae Yoldia seminuda 

Mollusca Pholadomyoida Lyonsiidae Lyonsia californica 

Mollusca Pholadomyoida Pandoridae Pandora filosa 1 1 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 

Mollusca Veneroida Cultellidae Siliqua alta 

Mollusca Veneroida Mactridae Spisula polynema 

Mollusca Veneroida Tellinidae Macoma eliminata 1 1 2 6 1 

Mollusca Veneroida Tellinidae Macoma lipara 

Mollusca Veneroida Tellinidae Macoma moesta 2 1 1 2 2 4 

Mollusca Veneroida Thyasiridae Axinospida serricata 9 7 6 7 14 5 

Mollusca Veneroida Veneridae Compsomyax subdiaphana 

Mollusca Veneroida Veneridae Nutricola lordi 2 7 2 

Nemertea Heteronemertea Lineidae Cerebratulus sp. 

Nemertea Heteronemertea Lineidae Linea sp. 1 

Total No. of Species 20 27 13 18 26 21 

Total No. of Individuals 87 96 36 39 96 80 
53 

Appendix Table 5.--Number of individuals of each infauna taxon collected per grab sample at 

Transect 9 in Chiniak Gully during August 1999. 

Phylum Order Family Species 9A 9B 9D 9E 9F 

Annelida Capitallida Capitellidae Mediomastus californiensis 2 7 2 8 

Annelida Capitallida Capitellidae Notomastus giganteus 2 

Annelida Capitallida Maldanidae 3 

Annelida Capitallida Maldanidae Axiothella rubrocincta 

Annelida Capitallida Maldanidae Clymenella sp. 1 

Annelida Capitallida Maldanidae Euclymene zonalis 4 1 1 3 

Annelida Capitallida Maldanidae Praxillella gracilis 

Annelida Chaetopterida Chaetopteridae Spiochaetopterus costarum 

Annelida Cossurida Cossuridae Cossura soyeri 

Annelida Eunicida Lumbrineridae Lumbrineris sp. 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 

Annelida Eunicida Lumbrineridae Lumbrineris inflata 

Annelida Eunicida Lumbrineridae Lumbrineris luti 4 13 4 14 

Annelida Eunicida Onuphidae Onuphis iridescens 1 3 

Annelida Magelonida Magelonidae Magelona berkeleyi 2 2 6 

Annelida Opheliida Opheliidae Ophelina sp. 

Annelida Opheliida Opheliidae Travisia forbesii 1 

Annelida Opheliida Scalibregmidae Scalibregma inflatum 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 13 2 3 1 

Annelida Orbiniida Paraonidae Aricidea lopezi 2 8 

Annelida Orbiniida Paraonidae Chaetozone sp. 2 1 

Annelida Orbiniida Paraonidae Levinsenia gracilis 1 

Annelida Oweniida Oweniidae Owenia cf. fusiformis 1 2 

Annelida Phyllodocida Glyceridae Glycera sp. 1 

Annelida Phyllodocida Glyceridae Hemipodus borealis 1 2 3 

Annelida Phyllodocida Goniadidae Goniada brunnea 1 

Annelida Phyllodocida Nephtidae Nephtys caeca 6 2 3 

Annelida Phyllodocida Nephtidae Nephtys cornuta 1 3 1 

Annelida Phyllodocida Pholodidae Pholoe glabra 1 
54


Annelida Phyllodocida Phyllodocidae Eteone longa 

Annelida Phyllodocida Phyllodocidae Phyllodocida sp. 

Annelida Phyllodocida Polynoidae Hesperonoe sp. 

Annelida Phyllodocida Spaerodoropsidae Spaerodoropsis sp. 

Annelida Phyllodocida Syllidae Syllis sp. 1 1 1 1 

Annelida Sabellida Sabellidae 

Annelida Sabellida Sabellidae Chone sp. 1 

Annelida Spionida Spionidae Dipolydora socialis 

Annelida Spionida Spionidae Prionospio lighti 

Annelida Spionida Spionidae Prionospio steenstrupi 6 1 1 

Annelida Spionida Spionidae Pygospio elegans 

Annelida Spionida Spionidae Spio cirrifera 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 3 1 1 2 

Annelida Terebellida Pectinaridae Pectinaria moorei 

Annelida Terebellida Terebellidae Amphitrite cirrata 1 1 

Annelida Terebellida Terebellidae Scionella sp. 

Annelida Terebellida Trichobranchidae Terebellides stroemi 1 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 3 2 1 1 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 1 1 

Arthropoda Amphipoda Ischyroceridae 1 1 1 

Arthropoda Amphipoda Phoxocephalidae Heterophoxus sp. 3 5 2 

Arthropoda Cumacea Diastylopsidae Diastylopsis dawsoni 155 1 2 

Arthropoda Cumacea Leuconiidae Leptocuma sp. 

Arthropoda Cumacea Leuconiidae Leucon magnadentata 

Arthropoda Decapoda Pinnixidae Pinnixa sp. 5 5 

Arthropoda Pantopoda Ammotheidae Achelia sp. 

Arthropoda Pantopoda Nymphonidae Nymphon sp. 

Chordata Stolidobranchia Mogulidae 3 

Echinodermata Apodida Chiridotidae 

Echinodermata Clypeasteroida Dendrasteridae Echinarchinus parma 

Echinodermata Dendrochiridotida Phyllophoridae Pentamera sp. 

Echinodermata Dendrochiridotida Sclerodactylidae Eupentacta sp. 

Echinodermata Molpadiida Molpadiidae Molpadia intermedia 1 
55


Echinodermata Ophiurida Amphiuridae Amphiodia sp. 37 

Echiura Echiuroinea Echiuridae Echiurus echiurus alaskensis 

Mollusca Cephalaspidea Atyidae Haminoea virescens 1 

Mollusca Cephalaspidea Cylichnidae Acteocina harpa 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 

Mollusca Mesogastropoda Eulimidae Eulima sp. 

Mollusca Mesogastropoda Naticidae Euspira pallida 

Mollusca Myoida Hiatellidae Hiatella arctica 

Mollusca Neogastropoda Amphissidae Alia gausapata 

Mollusca Neogastropoda Turridae Oenopota rosea 1 

Mollusca Nuculoida Nuculidae Ennucula tenuis 3 

Mollusca Nuculoida Yoldiidae Yoldia hyperborea 3 2 1 

Mollusca Nuculoida Yoldiidae Yoldia seminuda 

Mollusca Pholadomyoida Lyonsiidae Lyonsia californica 

Mollusca Pholadomyoida Pandoridae Pandora filosa 1 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 1 

Mollusca Veneroida Cultellidae Siliqua alta 

Mollusca Veneroida Mactridae Spisula polynema 

Mollusca Veneroida Tellinidae Macoma eliminata 1 2 

Mollusca Veneroida Tellinidae Macoma lipara 

Mollusca Veneroida Tellinidae Macoma moesta 1 1 1 8 

Mollusca Veneroida Thyasiridae Axinospida serricata 7 33 25 7 12 

Mollusca Veneroida Veneridae Compsomyax subdiaphana 

Mollusca Veneroida Veneridae Nutricola lordi 2 1 2 

Nemertea Heteronemertea Lineidae Cerebratulus sp. 

Nemertea Heteronemertea Lineidae Linea sp. 1 1 

Total No. of Species 19 17 18 25 24 

Total No. of Individuals 58 91 245 49 74 
56 

Appendix Table 5.--Number of individuals of each infauna taxon collected per grab sample at 

Transect 10 in Chiniak Gully during August 1999. 

Phylum Order Family Species 10A 10B 10C 10D 10E 10F 

Annelida Capitallida Capitellidae Mediomastus californiensis 7 16 18 12 3 6 

Annelida Capitallida Capitellidae Notomastus giganteus 1 3 2 1 

Annelida Capitallida Maldanidae 1 1 

Annelida Capitallida Maldanidae Axiothella rubrocincta 

Annelida Capitallida Maldanidae Clymenella sp. 

Annelida Capitallida Maldanidae Euclymene zonalis 2 7 9 4 3 4 

Annelida Capitallida Maldanidae Praxillella gracilis 

Annelida Chaetopterida Chaetopteridae Spiochaetopterus costarum 

Annelida Cossurida Cossuridae Cossura soyeri 

Annelida Eunicida Lumbrineridae Lumbrineris sp. 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 1 1 

Annelida Eunicida Lumbrineridae Lumbrineris inflata 1 

Annelida Eunicida Lumbrineridae Lumbrineris luti 3 8 6 4 6 2 

Annelida Eunicida Onuphidae Onuphis iridescens 1 1 

Annelida Magelonida Magelonidae Magelona berkeleyi 1 2 2 

Annelida Opheliida Opheliidae Ophelina sp. 

Annelida Opheliida Opheliidae Travisia forbesii 

Annelida Opheliida Scalibregmidae Scalibregma inflatum 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 1 3 3 

Annelida Orbiniida Paraonidae Aricidea lopezi 7 2 4 

Annelida Orbiniida Paraonidae Chaetozone sp. 1 3 4 2 3 24 

Annelida Orbiniida Paraonidae Levinsenia gracilis 1 

Annelida Oweniida Oweniidae Owenia cf. fusiformis 1 7 

Annelida Phyllodocida Glyceridae Glycera sp. 

Annelida Phyllodocida Glyceridae Hemipodus borealis 1 3 1 3 1 

Annelida Phyllodocida Goniadidae Goniada brunnea 2 

Annelida Phyllodocida Nephtidae Nephtys caeca 2 3 4 3 

Annelida Phyllodocida Nephtidae Nephtys cornuta 3 2 3 4 

Annelida Phyllodocida Pholodidae Pholoe glabra 1 
57


Annelida Phyllodocida Phyllodocidae Eteone longa 1 

Annelida Phyllodocida Phyllodocidae Phyllodocida sp. 

Annelida Phyllodocida Polynoidae Hesperonoe sp. 

Annelida Phyllodocida Spaerodoropsidae Spaerodoropsis sp. 

Annelida Phyllodocida Syllidae Syllis sp. 2 2 

Annelida Sabellida Sabellidae 1 1 

Annelida Sabellida Sabellidae Chone sp. 1 

Annelida Spionida Spionidae Dipolydora socialis 1 1 

Annelida Spionida Spionidae Prionospio lighti 

Annelida Spionida Spionidae Prionospio steenstrupi 2 8 7 2 4 

Annelida Spionida Spionidae Pygospio elegans 

Annelida Spionida Spionidae Spio cirrifera 2 2 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 1 4 3 1 3 1 

Annelida Terebellida Pectinaridae Pectinaria moorei 

Annelida Terebellida Terebellidae Amphitrite cirrata 

Annelida Terebellida Terebellidae Scionella sp. 1 

Annelida Terebellida Trichobranchidae Terebellides stroemi 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 1 6 1 1 2 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 1 1 2 1 

Arthropoda Amphipoda Ischyroceridae 5 2 

Arthropoda Amphipoda Phoxocephalidae Heterophoxus sp. 1 3 2 

Arthropoda Cumacea Diastylopsidae Diastylopsis dawsoni 1 

Arthropoda Cumacea Leuconiidae Leptocuma sp. 2 

Arthropoda Cumacea Leuconiidae Leucon magnadentata 1 2 

Arthropoda Decapoda Pinnixidae Pinnixa sp. 1 

Arthropoda Pantopoda Ammotheidae Achelia sp. 

Arthropoda Pantopoda Nymphonidae Nymphon sp. 

Chordata Stolidobranchia Mogulidae 1 

Echinodermata Apodida Chiridotidae 

Echinodermata Clypeasteroida Dendrasteridae Echinarchinus parma 

Echinodermata Dendrochiridotida Phyllophoridae Pentamera sp. 1 1 

Echinodermata Dendrochiridotida Sclerodactylidae Eupentacta sp. 

Echinodermata Molpadiida Molpadiidae Molpadia intermedia 
58


Echinodermata Ophiurida Amphiuridae Amphiodia sp. 1 

Echiura Echiuroinea Echiuridae Echiurus echiurus alaskensis 

Mollusca Cephalaspidea Atyidae Haminoea virescens 

Mollusca Cephalaspidea Cylichnidae Acteocina harpa 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 1 1 

Mollusca Mesogastropoda Eulimidae Eulima sp. 

Mollusca Mesogastropoda Naticidae Euspira pallida 1 1 

Mollusca Myoida Hiatellidae Hiatella arctica 

Mollusca Neogastropoda Amphissidae Alia gausapata 1 

Mollusca Neogastropoda Turridae Oenopota rosea 

Mollusca Nuculoida Nuculidae Ennucula tenuis 

Mollusca Nuculoida Yoldiidae Yoldia hyperborea 3 1 

Mollusca Nuculoida Yoldiidae Yoldia seminuda 

Mollusca Pholadomyoida Lyonsiidae Lyonsia californica 

Mollusca Pholadomyoida Pandoridae Pandora filosa 1 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 2 

Mollusca Veneroida Mactridae Spisula polynema 

Mollusca Veneroida Cultellidae Siliqua alta 8 1 

Mollusca Veneroida Tellinidae Macoma eliminata 1 2 1 4 8 

Mollusca Veneroida Tellinidae Macoma lipara 

Mollusca Veneroida Tellinidae Macoma moesta 3 2 6 1 2 2 

Mollusca Veneroida Thyasiridae Axinospida serricata 25 14 21 12 16 11 

Mollusca Veneroida Veneridae Compsomyax subdiaphana 

Mollusca Veneroida Veneridae Nutricola lordi 1 7 11 13 22 

Nemertea Heteronemertea Lineidae Cerebratulus sp. 

Nemertea Heteronemertea Lineidae Linea sp. 2 

Total No. of Species 21 24 29 21 25 21 

Total No. of Individuals 71 95 121 67 86 100 
59 

Appendix Table 6.--Number of individuals of each infauna taxon collected per grab sample at 

Transect 1 in Trinity Islands during August 1999. 

Phylum Order Family Species 1A 1B 1C 1D 1E 1F 

Annelida 1 

Annelida Capitallida Capitellidae Mediomastus californiensis 3 1 

Annelida Capitallida Maldanidae 1 

Annelida Capitallida Maldanidae Axiothella rubrocincta 1 

Annelida Capitallida Maldanidae Clymenella sp. 

Annelida Capitallida Maldanidae Euclymene zonalis 

Annelida Capitallida Maldanidae Praxillella gracilis 

Annelida Capitallida Maldanidae Praxilura sp. 

Annelida Chaetopterida Chaetopteridae Spiochaetopterus costarum 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 

Annelida Eunicida Lumbrineridae Lumbrineris luti 1 

Annelida Eunicida Onuphidae Onuphis iridescens 

Annelida Magelonida Magelonidae Magelona berkeleyi 2 4 3 1 1 

Annelida Opheliida Opheliidae Travisia forbesii 

Annelida Opheliida Opheliidae Travisia japonica 

Annelida Opheliida Scalibregmidae Hyboscolex pacificus 

Annelida Opheliida Scalibregmidae Scalibregma inflatum 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 1 

Annelida Orbiniida Paraonidae Aricidea antennata 

Annelida Orbiniida Paraonidae Aricidea lopezi 1 3 

Annelida Orbiniida Paraonidae Chaetozone sp. 1 1 2 

Annelida Orbiniida Paraonidae Cirratulus sp. 

Annelida Orbiniida Paraonidae Levinsenia gracilis 1 

Annelida Orbiniida Paraonidae Tharyx sp. I 2 1 

Annelida Orbiniida Paraonidae Tharyx sp. II 1 2 

Annelida Phyllodocida Aphrodotidae Aphrodites japonica 

Annelida Phyllodocida Glyceridae Hemipodus borealis 1 1 1 

Annelida Phyllodocida Goniadidae Goniada brunnea 

Annelida Phyllodocida Hesionidae 1 
60


Annelida Phyllodocida Nephtidae Nephtys sp. 

Annelida Phyllodocida Nephtidae Nephtys caeca 1 1 2 

Annelida Phyllodocida Nephtidae Nephtys cornuta 1 1 

Annelida Phyllodocida Nereididae Nereis sp. 

Annelida Phyllodocida Pholodidae Pholoe glabra 1 

Annelida Phyllodocida Pholodidae Sigalona sp. 

Annelida Phyllodocida Phyllodocidae Eteone longa 

Annelida Phyllodocida Polynoidae 2 1 

Annelida Phyllodocida Polynoidae Hesperonoe sp. 

Annelida Phyllodocida Polynoidae Hololepida magna 

Annelida Phyllodocida Syllidae Syllis sp. 

Annelida Sabellida Sabellidae 

Annelida Sabellida Sabellidae Potamilla sp. 

Annelida Sabellida Serpulidae 

Annelida Spionida Spionidae Dipolydora socialis 

Annelida Spionida Spionidae Laonice sp. 

Annelida Spionida Spionidae Prionospio lighti 

Annelida Spionida Spionidae Prionospio steenstrupi 1 1 2 3 

Annelida Spionida Spionidae Pygospio elegans 

Annelida Spionida Spionidae Spio cirrifera 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 

Annelida Terebellida Pectinaridae Pectinaria moorei 

Annelida Terebellida Sabellaridae Sabellaria cementarium 

Annelida Terebellida Terebellidae 

Annelida Terebellida Terebellidae Scionella sp. 1 

Annelida Terebellida Terebellidae Thelepas sp. 

Annelida Terebellida Trichobranchidae Terebellides stroemi 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 2 5 6 6 3 2 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 

Arthropoda Amphipoda Dexaminidae 1 

Arthropoda Amphipoda Ischyroceridae 

Arthropoda Amphipoda Oedicerotidae Monoculoides sp. 

Arthropoda Amphipoda Synopiidae Tiron biocellata 
61


Arthropoda Calanoida 

Arthropoda Cumacea Diastylidae Diastylis sp. 

Arthropoda Cumacea Diastylopsidae Diastylopsis dawsoni 

Arthropoda Cumacea Leuconiidae Eudorella sp. 1 

Arthropoda Cumacea Leuconiidae Lamprops fuscata 

Arthropoda Cumacea Leuconiidae Leucon fulvus 

Arthropoda Cumacea Leuconiidae Leucon magnadentata 

Arthropoda Decapoda Pinnixidae Pinnixa sp. 1 

Arthropoda Isopoda Gnathidae Gnathia sp. 1 1 

Arthropoda Ostracoda Species I 

Arthropoda Ostracoda Species II 1 

Arthropoda Ostracoda Species III 

Arthropoda Tanaidacea Paratanaidea Leptognathia sp. 

Echinodermata Apodida Chiridotidae 

Echinodermata Dendrochiridotida 

Echinodermata Ophiurida Amphiuridae Amphiodia sp. 2 1 

Echiura Echiuroinea Echiuridae Echiurus echiurus alaskensis 1 

Mollusca Archaeogastropoda Trochidae Solariella vancouverense 

Mollusca Cephalaspidea Atyidae Haminoea virescens 1 

Mollusca Dentaliida Dentaliidae Antalis pretiosum 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 

Mollusca Mesogastropoda Naticidae Euspira pallida 

Mollusca Myoida Hiatellidae Hiatella arctica 

Mollusca Mytiloida Mytilidae Crenella sp. 1 

Mollusca Neogastropoda Buccinidae Buccinum sp. 

Mollusca Neogastropoda Cancellaridae Admete sp. 1 

Mollusca Neogastropoda Turridae Oenopota rosea 

Mollusca Neogastropoda Turridae Ophiodermella sp. 

Mollusca Neoloricata Ischnochitonidae Lepidozona sp. 

Mollusca Nuculoida Nuculidae Ennucula tenuis 1 

Mollusca Osteroida Propeomussidae Parvamussium alaskensis 

Mollusca Pholadomyoida Cuspidariidae Cardiomya pectinata 

Mollusca Pholadomyoida Pandoridae Pandora filosa 
62


Mollusca Pyramidelliacea Pyramidellidae Odostomia sp. 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 2 1 1 

Mollusca Veneroida Astartiidae Astarte esquimalti 3 

Mollusca Veneroida Carditidae Cyclocardia sp. 

Mollusca Veneroida Cultellidae Siliqua alta 

Mollusca Veneroida Malletiidae Tindaria brunea 

Mollusca Veneroida Tellinidae Macoma eliminata 5 4 1 1 1 2 

Mollusca Veneroida Tellinidae Macoma moesta 2 1 

Mollusca Veneroida Thyasiridae Axinospida serricata 2 1 1 1 

Mollusca Veneroida Thyasiridae Thyasira flexuosa 

Nemertea Heteronemertea Lineidae Cerebratulus sp. 

Nemertea Heteronemertea Lineidae Linea sp. 

Priapulida Priapulamorpha Priapulidae Priapulus caudata 1 

Sipunucula Sipunuculida Golfingiidae 

Aschelminths 

Total No. of Species 13 17 15 14 9 7 

Total No. of Individuals 22 30 26 23 12 12 
63 

Appendix Table 6.--Numbers of individuals of each infauna taxon collected per grab sample at 

Transect 2 in Trinity Islands during August 1999. 

Phylum Order Family Species 2A 2B 2C 2D 2E 2F 

Annelida 1 

Annelida Capitallida Capitellidae Mediomastus californiensis 1 3 1 

Annelida Capitallida Maldanidae 

Annelida Capitallida Maldanidae Axiothella rubrocincta 1 1 

Annelida Capitallida Maldanidae Clymenella sp. 

Annelida Capitallida Maldanidae Euclymene zonalis 

Annelida Capitallida Maldanidae Praxillella gracilis 

Annelida Capitallida Maldanidae Praxilura sp. 

Annelida Chaetopterida Chaetopteridae Spiochaetopterus costarum 1 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 

Annelida Eunicida Lumbrineridae Lumbrineris luti 

Annelida Eunicida Onuphidae Onuphis iridescens 

Annelida Magelonida Magelonidae Magelona berkeleyi 1 2 4 3 

Annelida Opheliida Opheliidae Travisia forbesii 1 1 

Annelida Opheliida Opheliidae Travisia japonica 

Annelida Opheliida Scalibregmidae Hyboscolex pacificus 

Annelida Opheliida Scalibregmidae Scalibregma inflatum 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 

Annelida Orbiniida Paraonidae Aricidea antennata 

Annelida Orbiniida Paraonidae Aricidea lopezi 1 1 1 

Annelida Orbiniida Paraonidae Chaetozone sp. 4 4 1 

Annelida Orbiniida Paraonidae Cirratulus sp. 

Annelida Orbiniida Paraonidae Levinsenia gracilis 

Annelida Orbiniida Paraonidae Tharyx sp. I 1 1 1 

Annelida Orbiniida Paraonidae Tharyx sp. II 3 

Annelida Phyllodocida Aphrodotidae Aphrodites japonica 

Annelida Phyllodocida Glyceridae Hemipodus borealis 

Annelida Phyllodocida Goniadidae Goniada brunnea 1 1 

Annelida Phyllodocida Hesionidae 
64


Annelida Phyllodocida Nephtidae Nephtys sp. 

Annelida Phyllodocida Nephtidae Nephtys caeca 

Annelida Phyllodocida Nephtidae Nephtys cornuta 

Annelida Phyllodocida Nereididae Nereis sp. 

Annelida Phyllodocida Pholodidae Pholoe glabra 1 1 

Annelida Phyllodocida Pholodidae Sigalona sp. 3 

Annelida Phyllodocida Phyllodocidae Eteone longa 

Annelida Phyllodocida Polynoidae 1 1 1 

Annelida Phyllodocida Polynoidae Hesperonoe sp. 

Annelida Phyllodocida Polynoidae Hololepida magna 

Annelida Phyllodocida Syllidae Syllis sp. 

Annelida Sabellida Sabellidae 1 

Annelida Sabellida Sabellidae Potamilla sp. 

Annelida Sabellida Serpulidae 

Annelida Spionida Spionidae Dipolydora socialis 

Annelida Spionida Spionidae Laonice sp. 

Annelida Spionida Spionidae Prionospio lighti 

Annelida Spionida Spionidae Prionospio steenstrupi 2 1 2 1 

Annelida Spionida Spionidae Pygospio elegans 

Annelida Spionida Spionidae Spio cirrifera 1 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 

Annelida Terebellida Pectinaridae Pectinaria moorei 

Annelida Terebellida Sabellaridae Sabellaria cementarium 

Annelida Terebellida Terebellidae 1 

Annelida Terebellida Terebellidae Scionella sp. 1 1 

Annelida Terebellida Terebellidae Thelepas sp. 

Annelida Terebellida Trichobranchidae Terebellides stroemi 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 2 3 8 13 2 2 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 2 

Arthropoda Amphipoda Dexaminidae 2 

Arthropoda Amphipoda Ischyroceridae 1 

Arthropoda Amphipoda Oedicerotidae Monoculoides sp. 

Arthropoda Amphipoda Synopiidae Tiron biocellata 
65


Arthropoda Calanoida 

Arthropoda Cumacea Diastylidae Diastylis sp. 1 

Arthropoda Cumacea Diastylopsidae Diastylopsis dawsoni 

Arthropoda Cumacea Leuconiidae Eudorella sp. 1 

Arthropoda Cumacea Leuconiidae Lamprops fuscata 

Arthropoda Cumacea Leuconiidae Leucon fulvus 1 

Arthropoda Cumacea Leuconiidae Leucon magnadentata 

Arthropoda Decapoda Pinnixidae Pinnixa sp. 

Arthropoda Isopoda Gnathidae Gnathia sp. 2 2 

Arthropoda Ostracoda Species I 

Arthropoda Ostracoda Species II 

Arthropoda Ostracoda Species III 

Arthropoda Tanaidacea Paratanaidea Leptognathia sp. 

Echinodermata Apodida Chiridotidae 

Echinodermata Dendrochiridotida 

Echinodermata Ophiurida Amphiuridae Amphiodia sp. 1 1 

Echiura Echiuroinea Echiuridae Echiurus echiurus alaskensis 

Mollusca Archaeogastropoda Trochidae Solariella vancouverense 

Mollusca Cephalaspidea Atyidae Haminoea virescens 

Mollusca Dentaliida Dentaliidae Antalis pretiosum 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 

Mollusca Mesogastropoda Naticidae Euspira pallida 

Mollusca Myoida Hiatellidae Hiatella arctica 

Mollusca Mytiloida Mytilidae Crenella sp. 

Mollusca Neogastropoda Buccinidae Buccinum sp. 

Mollusca Neogastropoda Cancellaridae Admete sp. 

Mollusca Neogastropoda Turridae Oenopota rosea 

Mollusca Neogastropoda Turridae Ophiodermella sp. 

Mollusca Neoloricata Ischnochitonidae Lepidozona sp. 

Mollusca Nuculoida Nuculidae Ennucula tenuis 

Mollusca Osteroida Propeomussidae Parvamussium alaskensis 

Mollusca Pholadomyoida Cuspidariidae Cardiomya pectinata 1 

Mollusca Pholadomyoida Pandoridae Pandora filosa 
66


Mollusca Pyramidelliacea Pyramidellidae Odostomia sp. 1 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 

Mollusca Veneroida Astartiidae Astarte esquimalti 1 

Mollusca Veneroida Carditidae Cyclocardia sp. 1 

Mollusca Veneroida Cultellidae Siliqua alta 

Mollusca Veneroida Malletiidae Tindaria brunea 

Mollusca Veneroida Tellinidae Macoma eliminata 6 1 3 4 1 

Mollusca Veneroida Tellinidae Macoma moesta 3 3 2 

Mollusca Veneroida Thyasiridae Axinospida serricata 2 2 

Mollusca Veneroida Thyasiridae Thyasira flexuosa 

Nemertea Heteronemertea Lineidae Cerebratulus sp. 

Nemertea Heteronemertea Lineidae Linea sp. 1 

Priapulida Priapulamorpha Priapulidae Priapulus caudata 1 

Sipunucula Sipunuculida Golfingiidae 

Aschelminths 

Total No. of Species 11 11 12 10 16 12 

Total No. of Individuals 12 22 24 30 31 18 
67 

Appendix Table 6.--Number of individuals of each infauna taxon collected per grab sample at 

Transect 3 in Trinity Islands during August 1999. 

Phylum Order Family Species 3A 3B 3C 3D 3E 3F 

Annelida 

Annelida Capitallida Capitellidae Mediomastus californiensis 7 4 11 1 2 

Annelida Capitallida Maldanidae 

Annelida Capitallida Maldanidae Axiothella rubrocincta 

Annelida Capitallida Maldanidae Clymenella sp. 

Annelida Capitallida Maldanidae Euclymene zonalis 

Annelida Capitallida Maldanidae Praxillella gracilis 

Annelida Capitallida Maldanidae Praxilura sp. 

Annelida Chaetopterida Chaetopteridae Spiochaetopterus costarum 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 1 

Annelida Eunicida Lumbrineridae Lumbrineris luti 

Annelida Eunicida Onuphidae Onuphis iridescens 1 

Annelida Magelonida Magelonidae Magelona berkeleyi 3 3 2 3 

Annelida Opheliida Opheliidae Travisia forbesii 1 1 

Annelida Opheliida Opheliidae Travisia japonica 

Annelida Opheliida Scalibregmidae Hyboscolex pacificus 

Annelida Opheliida Scalibregmidae Scalibregma inflatum 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 

Annelida Orbiniida Paraonidae Aricidea antennata 

Annelida Orbiniida Paraonidae Aricidea lopezi 3 1 

Annelida Orbiniida Paraonidae Chaetozone sp. 1 4 

Annelida Orbiniida Paraonidae Cirratulus sp. 

Annelida Orbiniida Paraonidae Levinsenia gracilis 1 

Annelida Orbiniida Paraonidae Tharyx sp. I 

Annelida Orbiniida Paraonidae Tharyx sp. II 1 1 

Annelida Phyllodocida Aphrodotidae Aphrodites japonica 

Annelida Phyllodocida Glyceridae Hemipodus borealis 1 1 

Annelida Phyllodocida Goniadidae Goniada brunnea 

Annelida Phyllodocida Hesionidae 
68


Annelida Phyllodocida Nephtidae Nephtys sp. 

Annelida Phyllodocida Nephtidae Nephtys caeca 1 

Annelida Phyllodocida Nephtidae Nephtys cornuta 1 1 

Annelida Phyllodocida Nereididae Nereis sp. 

Annelida Phyllodocida Pholodidae Pholoe glabra 

Annelida Phyllodocida Pholodidae Sigalona sp. 

Annelida Phyllodocida Phyllodocidae Eteone longa 

Annelida Phyllodocida Polynoidae 1 1 

Annelida Phyllodocida Polynoidae Hesperonoe sp. 1 

Annelida Phyllodocida Polynoidae Hololepida magna 

Annelida Phyllodocida Syllidae Syllis sp. 

Annelida Sabellida Sabellidae 3 1 1 

Annelida Sabellida Sabellidae Potamilla sp. 

Annelida Sabellida Serpulidae 1 

Annelida Spionida Spionidae Dipolydora socialis 

Annelida Spionida Spionidae Laonice sp. 

Annelida Spionida Spionidae Prionospio lighti 1 

Annelida Spionida Spionidae Prionospio steenstrupi 1 2 1 1 

Annelida Spionida Spionidae Pygospio elegans 

Annelida Spionida Spionidae Spio cirrifera 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 1 

Annelida Terebellida Pectinaridae Pectinaria moorei 

Annelida Terebellida Sabellaridae Sabellaria cementarium 

Annelida Terebellida Terebellidae 1 

Annelida Terebellida Terebellidae Scionella sp. 1 

Annelida Terebellida Terebellidae Thelepas sp. 1 

Annelida Terebellida Trichobranchidae Terebellides stroemi 1 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 5 4 13 12 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 

Arthropoda Amphipoda Dexaminidae 1 1 

Arthropoda Amphipoda Ischyroceridae 

Arthropoda Amphipoda Oedicerotidae Monoculoides sp. 

Arthropoda Amphipoda Synopiidae Tiron biocellata 
69


Arthropoda Calanoida 1 

Arthropoda Cumacea Diastylidae Diastylis sp. 

Arthropoda Cumacea Diastylopsidae Diastylopsis dawsoni 

Arthropoda Cumacea Leuconiidae Eudorella sp. 

Arthropoda Cumacea Leuconiidae Lamprops fuscata 

Arthropoda Cumacea Leuconiidae Leucon fulvus 

Arthropoda Cumacea Leuconiidae Leucon magnadentata 

Arthropoda Decapoda Pinnixidae Pinnixa sp. 

Arthropoda Isopoda Gnathidae Gnathia sp. 2 

Arthropoda Ostracoda Species I 1 

Arthropoda Ostracoda Species II 

Arthropoda Ostracoda Species III 1 

Arthropoda Tanaidacea Paratanaidea Leptognathia sp. 1 

Echinodermata Apodida Chiridotidae 

Echinodermata Dendrochiridotida 3 

Echinodermata Ophiurida Amphiuridae Amphiodia sp. 4 1 

Echiura Echiuroinea Echiuridae Echiurus echiurus alaskensis 

Mollusca Archaeogastropoda Trochidae Solariella vancouverense 

Mollusca Cephalaspidea Atyidae Haminoea virescens 

Mollusca Dentaliida Dentaliidae Antalis pretiosum 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 

Mollusca Mesogastropoda Naticidae Euspira pallida 

Mollusca Myoida Hiatellidae Hiatella arctica 1 

Mollusca Mytiloida Mytilidae Crenella sp. 

Mollusca Neogastropoda Buccinidae Buccinum sp. 

Mollusca Neogastropoda Cancellaridae Admete sp. 

Mollusca Neogastropoda Turridae Oenopota rosea 

Mollusca Neogastropoda Turridae Ophiodermella sp. 

Mollusca Neoloricata Ischnochitonidae Lepidozona sp. 1 

Mollusca Nuculoida Nuculidae Ennucula tenuis 

Mollusca Osteroida Propeomussidae Parvamussium alaskensis 1 

Mollusca Pholadomyoida Cuspidariidae Cardiomya pectinata 

Mollusca Pholadomyoida Pandoridae Pandora filosa 
70


Mollusca Pyramidelliacea Pyramidellidae Odostomia sp. 1 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 

Mollusca Veneroida Astartiidae Astarte esquimalti 

Mollusca Veneroida Carditidae Cyclocardia sp. 

Mollusca Veneroida Cultellidae Siliqua alta 

Mollusca Veneroida Malletiidae Tindaria brunea 1 

Mollusca Veneroida Tellinidae Macoma eliminata 15 2 3 

Mollusca Veneroida Tellinidae Macoma moesta 1 1 2 

Mollusca Veneroida Thyasiridae Axinospida serricata 4 1 1 2 

Mollusca Veneroida Thyasiridae Thyasira flexuosa 

Nemertea Heteronemertea Lineidae Cerebratulus sp. 

Nemertea Heteronemertea Lineidae Linea sp. 1 

Priapulida Priapulamorpha Priapulidae Priapulus caudata 

Sipunucula Sipunuculida Golfingiidae 

Aschelminths 

Total No. of Species 19 12 13 5 6 17 

Total No. of Individuals 51 26 37 6 11 34 
71 

Appendix Table 6.--Number of individuals of each infauna taxon collected per grab sample at 

Transect 4 in Trinity Islands during August 1999. 

Phylum Order Family Species 4A 4B 4C 4D 4E 4F 

Annelida 

Annelida Capitallida Capitellidae Mediomastus californiensis 3 5 3 1 6 2 

Annelida Capitallida Maldanidae 

Annelida Capitallida Maldanidae Axiothella rubrocincta 

Annelida Capitallida Maldanidae Clymenella sp. 

Annelida Capitallida Maldanidae Euclymene zonalis 

Annelida Capitallida Maldanidae Praxillella gracilis 

Annelida Capitallida Maldanidae Praxilura sp. 

Annelida Chaetopterida Chaetopteridae Spiochaetopterus costarum 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 

Annelida Eunicida Lumbrineridae Lumbrineris luti 1 

Annelida Eunicida Onuphidae Onuphis iridescens 1 

Annelida Magelonida Magelonidae Magelona berkeleyi 3 2 4 1 5 

Annelida Opheliida Opheliidae Travisia forbesii 

Annelida Opheliida Opheliidae Travisia japonica 

Annelida Opheliida Scalibregmidae Hyboscolex pacificus 

Annelida Opheliida Scalibregmidae Scalibregma inflatum 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 2 

Annelida Orbiniida Paraonidae Aricidea antennata 

Annelida Orbiniida Paraonidae Aricidea lopezi 1 1 

Annelida Orbiniida Paraonidae Chaetozone sp. 1 5 2 1 3 3 

Annelida Orbiniida Paraonidae Cirratulus sp. 

Annelida Orbiniida Paraonidae Levinsenia gracilis 1 

Annelida Orbiniida Paraonidae Tharyx sp. I 1 

Annelida Orbiniida Paraonidae Tharyx sp. II 2 

Annelida Phyllodocida Aphrodotidae Aphrodites japonica 

Annelida Phyllodocida Glyceridae Hemipodus borealis 1 1 

Annelida Phyllodocida Goniadidae Goniada brunnea 2 

Annelida Phyllodocida Hesionidae 
72


Annelida Phyllodocida Nephtidae Nephtys sp. 

Annelida Phyllodocida Nephtidae Nephtys caeca 1 1 

Annelida Phyllodocida Nephtidae Nephtys cornuta 1 4 

Annelida Phyllodocida Nereididae Nereis sp. 

Annelida Phyllodocida Pholodidae Pholoe glabra 1 

Annelida Phyllodocida Pholodidae Sigalona sp. 

Annelida Phyllodocida Phyllodocidae Eteone longa 

Annelida Phyllodocida Polynoidae 1 

Annelida Phyllodocida Polynoidae Hesperonoe sp. 

Annelida Phyllodocida Polynoidae Hololepida magna 1 

Annelida Phyllodocida Syllidae Syllis sp. 

Annelida Sabellida Sabellidae 

Annelida Sabellida Sabellidae Potamilla sp. 

Annelida Sabellida Serpulidae 

Annelida Spionida Spionidae Dipolydora socialis 

Annelida Spionida Spionidae Laonice sp. 

Annelida Spionida Spionidae Prionospio lighti 1 

Annelida Spionida Spionidae Prionospio steenstrupi 2 3 2 4 3 

Annelida Spionida Spionidae Pygospio elegans 

Annelida Spionida Spionidae Spio cirrifera 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 1 

Annelida Terebellida Pectinaridae Pectinaria moorei 

Annelida Terebellida Sabellaridae Sabellaria cementarium 1 

Annelida Terebellida Terebellidae 1 

Annelida Terebellida Terebellidae Scionella sp. 1 1 

Annelida Terebellida Terebellidae Thelepas sp. 

Annelida Terebellida Trichobranchidae Terebellides stroemi 1 1 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 7 4 1 1 1 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 

Arthropoda Amphipoda Dexaminidae 1 

Arthropoda Amphipoda Ischyroceridae 1 1 

Arthropoda Amphipoda Oedicerotidae Monoculoides sp. 

Arthropoda Amphipoda Synopiidae Tiron biocellata 
73


Arthropoda Calanoida 

Arthropoda Cumacea Diastylidae Diastylis sp. 

Arthropoda Cumacea Diastylopsidae Diastylopsis dawsoni 

Arthropoda Cumacea Leuconiidae Eudorella sp. 

Arthropoda Cumacea Leuconiidae Lamprops fuscata 1 

Arthropoda Cumacea Leuconiidae Leucon fulvus 

Arthropoda Cumacea Leuconiidae Leucon magnadentata 1 

Arthropoda Decapoda Pinnixidae Pinnixa sp. 

Arthropoda Isopoda Gnathidae Gnathia sp. 1 

Arthropoda Ostracoda Species I 

Arthropoda Ostracoda Species II 

Arthropoda Ostracoda Species III 

Arthropoda Tanaidacea Paratanaidea Leptognathia sp. 

Echinodermata Apodida Chiridotidae 

Echinodermata Dendrochiridotida 

Echinodermata Ophiurida Amphiuridae Amphiodia sp. 3 4 1 

Echiura Echiuroinea Echiuridae Echiurus echiurus alaskensis 

Mollusca Archaeogastropoda Trochidae Solariella vancouverense 1 

Mollusca Cephalaspidea Atyidae Haminoea virescens 

Mollusca Dentaliida Dentaliidae Antalis pretiosum 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 

Mollusca Mesogastropoda Naticidae Euspira pallida 

Mollusca Myoida Hiatellidae Hiatella arctica 

Mollusca Mytiloida Mytilidae Crenella sp. 1 

Mollusca Neogastropoda Buccinidae Buccinum sp. 

Mollusca Neogastropoda Cancellaridae Admete sp. 

Mollusca Neogastropoda Turridae Oenopota rosea 

Mollusca Neogastropoda Turridae Ophiodermella sp. 1 

Mollusca Neoloricata Ischnochitonidae Lepidozona sp. 

Mollusca Nuculoida Nuculidae Ennucula tenuis 

Mollusca Osteroida Propeomussidae Parvamussium alaskensis 

Mollusca Pholadomyoida Cuspidariidae Cardiomya pectinata 

Mollusca Pholadomyoida Pandoridae Pandora filosa 
74


Mollusca Pyramidelliacea Pyramidellidae Odostomia sp. 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 

Mollusca Veneroida Astartiidae Astarte esquimalti 1 

Mollusca Veneroida Carditidae Cyclocardia sp. 

Mollusca Veneroida Cultellidae Siliqua alta 1 

Mollusca Veneroida Malletiidae Tindaria brunea 

Mollusca Veneroida Tellinidae Macoma eliminata 3 7 3 8 

Mollusca Veneroida Tellinidae Macoma moesta 4 2 1 

Mollusca Veneroida Thyasiridae Axinospida serricata 1 1 1 

Mollusca Veneroida Thyasiridae Thyasira flexuosa 

Nemertea Heteronemertea Lineidae Cerebratulus sp. 

Nemertea Heteronemertea Lineidae Linea sp. 1 1 1 

Priapulida Priapulamorpha Priapulidae Priapulus caudata 1 

Sipunucula Sipunuculida Golfingiidae 1 

Aschelminths 

Total No. of Species 12 15 14 8 17 16 

Total No. of Individuals 28 41 21 11 35 29 
75 

Appendix Table 6.--Number of individuals of each infauna taxon collected per grab sample at 

Transect 5 in Trinity Islands during August 1999. 

Phylum Order Family Species 5A 5B 5C 5D 5E 5F 

Annelida 

Annelida Capitallida Capitellidae Mediomastus californiensis 1 10 1 10 1 5 

Annelida Capitallida Maldanidae 1 1 

Annelida Capitallida Maldanidae Axiothella rubrocincta 

Annelida Capitallida Maldanidae Clymenella sp. 

Annelida Capitallida Maldanidae Euclymene zonalis 

Annelida Capitallida Maldanidae Praxillella gracilis 

Annelida Capitallida Maldanidae Praxilura sp. 

Annelida Chaetopterida Chaetopteridae Spiochaetopterus costarum 1 1 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 2 1 

Annelida Eunicida Lumbrineridae Lumbrineris luti 

Annelida Eunicida Onuphidae Onuphis iridescens 1 

Annelida Magelonida Magelonidae Magelona berkeleyi 2 5 1 5 3 

Annelida Opheliida Opheliidae Travisia forbesii 1 1 

Annelida Opheliida Opheliidae Travisia japonica 

Annelida Opheliida Scalibregmidae Hyboscolex pacificus 

Annelida Opheliida Scalibregmidae Scalibregma inflatum 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 1 

Annelida Orbiniida Paraonidae Aricidea antennata 

Annelida Orbiniida Paraonidae Aricidea lopezi 1 2 

Annelida Orbiniida Paraonidae Chaetozone sp. 1 1 4 8 4 

Annelida Orbiniida Paraonidae Cirratulus sp. 1 

Annelida Orbiniida Paraonidae Levinsenia gracilis 2 

Annelida Orbiniida Paraonidae Tharyx sp. I 1 1 1 

Annelida Orbiniida Paraonidae Tharyx sp. II 1 1 

Annelida Phyllodocida Aphrodotidae Aphrodites japonica 

Annelida Phyllodocida Glyceridae Hemipodus borealis 1 4 

Annelida Phyllodocida Goniadidae Goniada brunnea 1 

Annelida Phyllodocida Hesionidae 
76


Annelida Phyllodocida Nephtidae Nephtys sp. 

Annelida Phyllodocida Nephtidae Nephtys caeca 2 2 

Annelida Phyllodocida Nephtidae Nephtys cornuta 1 

Annelida Phyllodocida Nereididae Nereis sp. 

Annelida Phyllodocida Pholodidae Pholoe glabra 1 

Annelida Phyllodocida Pholodidae Sigalona sp. 

Annelida Phyllodocida Phyllodocidae Eteone longa 1 

Annelida Phyllodocida Polynoidae 

Annelida Phyllodocida Polynoidae Hesperonoe sp. 

Annelida Phyllodocida Polynoidae Hololepida magna 1 

Annelida Phyllodocida Syllidae Syllis sp. 1 

Annelida Sabellida Sabellidae 

Annelida Sabellida Sabellidae Potamilla sp. 1 1 

Annelida Sabellida Serpulidae 

Annelida Spionida Spionidae Dipolydora socialis 

Annelida Spionida Spionidae Laonice sp. 

Annelida Spionida Spionidae Prionospio lighti 

Annelida Spionida Spionidae Prionospio steenstrupi 1 8 2 6 2 2 

Annelida Spionida Spionidae Pygospio elegans 

Annelida Spionida Spionidae Spio cirrifera 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 1 

Annelida Terebellida Pectinaridae Pectinaria moorei 

Annelida Terebellida Sabellaridae Sabellaria cementarium 

Annelida Terebellida Terebellidae 1 

Annelida Terebellida Terebellidae Scionella sp. 1 1 

Annelida Terebellida Terebellidae Thelepas sp. 1 

Annelida Terebellida Trichobranchidae Terebellides stroemi 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 10 1 8 2 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 1 

Arthropoda Amphipoda Dexaminidae 1 

Arthropoda Amphipoda Ischyroceridae 1 

Arthropoda Amphipoda Oedicerotidae Monoculoides sp. 

Arthropoda Amphipoda Synopiidae Tiron biocellata 
77


Arthropoda Calanoida 

Arthropoda Cumacea Diastylidae Diastylis sp. 

Arthropoda Cumacea Diastylopsidae Diastylopsis dawsoni 

Arthropoda Cumacea Leuconiidae Eudorella sp. 

Arthropoda Cumacea Leuconiidae Lamprops fuscata 

Arthropoda Cumacea Leuconiidae Leucon fulvus 

Arthropoda Cumacea Leuconiidae Leucon magnadentata 

Arthropoda Decapoda Pinnixidae Pinnixa sp. 

Arthropoda Isopoda Gnathidae Gnathia sp. 1 

Arthropoda Ostracoda Species I 

Arthropoda Ostracoda Species II 

Arthropoda Ostracoda Species III 

Arthropoda Tanaidacea Paratanaidea Leptognathia sp. 

Echinodermata Apodida Chiridotidae 1 

Echinodermata Dendrochiridotida 

Echinodermata Ophiurida Amphiuridae Amphiodia sp. 4 

Echiura Echiuroinea Echiuridae Echiurus echiurus alaskensis 1 1 

Mollusca Archaeogastropoda Trochidae Solariella vancouverense 

Mollusca Cephalaspidea Atyidae Haminoea virescens 

Mollusca Dentaliida Dentaliidae Antalis pretiosum 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 

Mollusca Mesogastropoda Naticidae Euspira pallida 

Mollusca Myoida Hiatellidae Hiatella arctica 

Mollusca Mytiloida Mytilidae Crenella sp. 

Mollusca Neogastropoda Buccinidae Buccinum sp. 1 

Mollusca Neogastropoda Cancellaridae Admete sp. 

Mollusca Neogastropoda Turridae Oenopota rosea 1 

Mollusca Neogastropoda Turridae Ophiodermella sp. 

Mollusca Neoloricata Ischnochitonidae Lepidozona sp. 

Mollusca Nuculoida Nuculidae Ennucula tenuis 

Mollusca Osteroida Propeomussidae Parvamussium alaskensis 

Mollusca Pholadomyoida Cuspidariidae Cardiomya pectinata 

Mollusca Pholadomyoida Pandoridae Pandora filosa 
78


Mollusca Pyramidelliacea Pyramidellidae Odostomia sp. 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 2 

Mollusca Veneroida Astartiidae Astarte esquimalti 2 1 

Mollusca Veneroida Carditidae Cyclocardia sp. 

Mollusca Veneroida Cultellidae Siliqua alta 

Mollusca Veneroida Malletiidae Tindaria brunea 

Mollusca Veneroida Tellinidae Macoma eliminata 2 6 9 4 1 

Mollusca Veneroida Tellinidae Macoma moesta 2 2 

Mollusca Veneroida Thyasiridae Axinospida serricata 2 2 1 5 

Mollusca Veneroida Thyasiridae Thyasira flexuosa 

Nemertea Heteronemertea Lineidae Cerebratulus sp. 

Nemertea Heteronemertea Lineidae Linea sp. 1 1 1 

Priapulida Priapulamorpha Priapulidae Priapulus caudata 

Sipunucula Sipunuculida Golfingiidae 2 

Aschelminths 

Total No. of Species 17 20 11 18 9 15 

Total No. of Individuals 24 58 16 64 14 31 
79 

Appendix Table 6.--Number of individuals of each infauna taxon collected per grab sample at 

Transect 6 in Trinity Islands during August 1999. 

Phylum Order Family Species 6A 6B 6C 6D 6E 6F 

Annelida 

Annelida Capitallida Capitellidae Mediomastus californiensis 8 2 17 4 1 7 

Annelida Capitallida Maldanidae 

Annelida Capitallida Maldanidae Axiothella rubrocincta 1 

Annelida Capitallida Maldanidae Clymenella sp. 1 

Annelida Capitallida Maldanidae Euclymene zonalis 

Annelida Capitallida Maldanidae Praxillella gracilis 

Annelida Capitallida Maldanidae Praxilura sp. 

Annelida Chaetopterida Chaetopteridae Spiochaetopterus costarum 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 

Annelida Eunicida Lumbrineridae Lumbrineris luti 2 1 

Annelida Eunicida Onuphidae Onuphis iridescens 

Annelida Magelonida Magelonidae Magelona berkeleyi 6 2 2 3 2 

Annelida Opheliida Opheliidae Travisia forbesii 1 

Annelida Opheliida Opheliidae Travisia japonica 

Annelida Opheliida Scalibregmidae Hyboscolex pacificus 1 

Annelida Opheliida Scalibregmidae Scalibregma inflatum 1 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 2 

Annelida Orbiniida Paraonidae Aricidea antennata 3 

Annelida Orbiniida Paraonidae Aricidea lopezi 1 1 3 

Annelida Orbiniida Paraonidae Chaetozone sp. 5 1 1 1 

Annelida Orbiniida Paraonidae Cirratulus sp. 

Annelida Orbiniida Paraonidae Levinsenia gracilis 1 

Annelida Orbiniida Paraonidae Tharyx sp. I 1 

Annelida Orbiniida Paraonidae Tharyx sp. II 

Annelida Phyllodocida Aphrodotidae Aphrodites japonica 

Annelida Phyllodocida Glyceridae Hemipodus borealis 1 1 

Annelida Phyllodocida Goniadidae Goniada brunnea 1 2 1 1 

Annelida Phyllodocida Hesionidae 
80


Annelida Phyllodocida Nephtidae Nephtys sp. 

Annelida Phyllodocida Nephtidae Nephtys caeca 1 2 

Annelida Phyllodocida Nephtidae Nephtys cornuta 1 

Annelida Phyllodocida Nereididae Nereis sp. 

Annelida Phyllodocida Pholodidae Pholoe glabra 

Annelida Phyllodocida Pholodidae Sigalona sp. 

Annelida Phyllodocida Phyllodocidae Eteone longa 1 1 1 

Annelida Phyllodocida Polynoidae 1 

Annelida Phyllodocida Polynoidae Hesperonoe sp. 1 1 

Annelida Phyllodocida Polynoidae Hololepida magna 1 

Annelida Phyllodocida Syllidae Syllis sp. 

Annelida Sabellida Sabellidae 1 

Annelida Sabellida Sabellidae Potamilla sp. 1 

Annelida Sabellida Serpulidae 

Annelida Spionida Spionidae Dipolydora socialis 

Annelida Spionida Spionidae Laonice sp. 

Annelida Spionida Spionidae Prionospio lighti 

Annelida Spionida Spionidae Prionospio steenstrupi 15 1 4 1 1 4 

Annelida Spionida Spionidae Pygospio elegans 

Annelida Spionida Spionidae Spio cirrifera 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 

Annelida Terebellida Pectinaridae Pectinaria moorei 1 

Annelida Terebellida Sabellaridae Sabellaria cementarium 

Annelida Terebellida Terebellidae 

Annelida Terebellida Terebellidae Scionella sp. 

Annelida Terebellida Terebellidae Thelepas sp. 1 

Annelida Terebellida Trichobranchidae Terebellides stroemi 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 8 1 5 6 37 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 1 

Arthropoda Amphipoda Dexaminidae 

Arthropoda Amphipoda Ischyroceridae 1 

Arthropoda Amphipoda Oedicerotidae Monoculoides sp. 

Arthropoda Amphipoda Synopiidae Tiron biocellata 
81


Arthropoda Calanoida 

Arthropoda Cumacea Diastylidae Diastylis sp. 

Arthropoda Cumacea Diastylopsidae Diastylopsis dawsoni 

Arthropoda Cumacea Leuconiidae Eudorella sp. 1 1 

Arthropoda Cumacea Leuconiidae Lamprops fuscata 

Arthropoda Cumacea Leuconiidae Leucon fulvus 

Arthropoda Cumacea Leuconiidae Leucon magnadentata 1 

Arthropoda Decapoda Pinnixidae Pinnixa sp. 

Arthropoda Isopoda Gnathidae Gnathia sp. 1 

Arthropoda Ostracoda Species I 

Arthropoda Ostracoda Species II 

Arthropoda Ostracoda Species III 

Arthropoda Tanaidacea Paratanaidea Leptognathia sp. 

Echinodermata Apodida Chiridotidae 

Echinodermata Dendrochiridotida 

Echinodermata Ophiurida Amphiuridae Amphiodia sp. 3 2 2 1 1 

Echiura Echiuroinea Echiuridae Echiurus echiurus alaskensis 

Mollusca Archaeogastropoda Trochidae Solariella vancouverense 1 

Mollusca Cephalaspidea Atyidae Haminoea virescens 1 

Mollusca Dentaliida Dentaliidae Antalis pretiosum 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 

Mollusca Mesogastropoda Naticidae Euspira pallida 

Mollusca Myoida Hiatellidae Hiatella arctica 

Mollusca Mytiloida Mytilidae Crenella sp. 

Mollusca Neogastropoda Buccinidae Buccinum sp. 

Mollusca Neogastropoda Cancellaridae Admete sp. 

Mollusca Neogastropoda Turridae Oenopota rosea 

Mollusca Neogastropoda Turridae Ophiodermella sp. 

Mollusca Neoloricata Ischnochitonidae Lepidozona sp. 

Mollusca Nuculoida Nuculidae Ennucula tenuis 

Mollusca Osteroida Propeomussidae Parvamussium alaskensis 

Mollusca Pholadomyoida Cuspidariidae Cardiomya pectinata 

Mollusca Pholadomyoida Pandoridae Pandora filosa 
82


Mollusca Pyramidelliacea Pyramidellidae Odostomia sp. 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 1 

Mollusca Veneroida Astartiidae Astarte esquimalti 

Mollusca Veneroida Carditidae Cyclocardia sp. 

Mollusca Veneroida Cultellidae Siliqua alta 

Mollusca Veneroida Malletiidae Tindaria brunea 

Mollusca Veneroida Tellinidae Macoma eliminata 7 1 3 4 1 24 

Mollusca Veneroida Tellinidae Macoma moesta 2 1 1 

Mollusca Veneroida Thyasiridae Axinospida serricata 2 1 1 1 1 13 

Mollusca Veneroida Thyasiridae Thyasira flexuosa 

Nemertea Heteronemertea Lineidae Cerebratulus sp. 

Nemertea Heteronemertea Lineidae Linea sp. 2 

Priapulida Priapulamorpha Priapulidae Priapulus caudata 1 

Sipunucula Sipunuculida Golfingiidae 

Aschelminths 

Total No. of Species 17 8 17 14 17 18 

Total No. of Individuals 66 10 40 26 25 104 
83 

Appendix Table 6.--Number of individuals of each infauna taxon collected per grab sample at 

Transect 7 in Trinity Islands during August 1999. 

Phylum Order Family Species 7A 7B 7C 7D 7E 7F 

Annelida 

Annelida Capitallida Capitellidae Mediomastus californiensis 7 11 5 1 4 1 

Annelida Capitallida Maldanidae 

Annelida Capitallida Maldanidae Axiothella rubrocincta 2 1 

Annelida Capitallida Maldanidae Clymenella sp. 1 

Annelida Capitallida Maldanidae Euclymene zonalis 

Annelida Capitallida Maldanidae Praxillella gracilis 1 

Annelida Capitallida Maldanidae Praxilura sp. 1 

Annelida Chaetopterida Chaetopteridae Spiochaetopterus costarum 1 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 2 1 1 

Annelida Eunicida Lumbrineridae Lumbrineris luti 1 

Annelida Eunicida Onuphidae Onuphis iridescens 1 1 1 1 

Annelida Magelonida Magelonidae Magelona berkeleyi 3 6 2 3 2 

Annelida Opheliida Opheliidae Travisia forbesii 

Annelida Opheliida Opheliidae Travisia japonica 1 

Annelida Opheliida Scalibregmidae Hyboscolex pacificus 1 

Annelida Opheliida Scalibregmidae Scalibregma inflatum 1 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 1 

Annelida Orbiniida Paraonidae Aricidea antennata 

Annelida Orbiniida Paraonidae Aricidea lopezi 5 1 1 1 

Annelida Orbiniida Paraonidae Chaetozone sp. 1 3 2 2 6 4 

Annelida Orbiniida Paraonidae Cirratulus sp. 

Annelida Orbiniida Paraonidae Levinsenia gracilis 

Annelida Orbiniida Paraonidae Tharyx sp. I 1 1 1 

Annelida Orbiniida Paraonidae Tharyx sp. II 2 1 

Annelida Phyllodocida Aphrodotidae Aphrodites japonica 

Annelida Phyllodocida Glyceridae Hemipodus borealis 1 2 

Annelida Phyllodocida Goniadidae Goniada brunnea 1 

Annelida Phyllodocida Hesionidae 1 
84


Annelida Phyllodocida Nephtidae Nephtys sp. 1 

Annelida Phyllodocida Nephtidae Nephtys caeca 2 1 1 2 

Annelida Phyllodocida Nephtidae Nephtys cornuta 1 3 2 

Annelida Phyllodocida Nereididae Nereis sp. 

Annelida Phyllodocida Pholodidae Pholoe glabra 1 1 

Annelida Phyllodocida Pholodidae Sigalona sp. 

Annelida Phyllodocida Phyllodocidae Eteone longa 1 1 

Annelida Phyllodocida Polynoidae 

Annelida Phyllodocida Polynoidae Hesperonoe sp. 

Annelida Phyllodocida Polynoidae Hololepida magna 

Annelida Phyllodocida Syllidae Syllis sp. 

Annelida Sabellida Sabellidae 1 

Annelida Sabellida Sabellidae Potamilla sp. 

Annelida Sabellida Serpulidae 

Annelida Spionida Spionidae Dipolydora socialis 1 

Annelida Spionida Spionidae Laonice sp. 

Annelida Spionida Spionidae Prionospio lighti 1 

Annelida Spionida Spionidae Prionospio steenstrupi 7 2 4 3 3 4 

Annelida Spionida Spionidae Pygospio elegans 

Annelida Spionida Spionidae Spio cirrifera 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 1 

Annelida Terebellida Pectinaridae Pectinaria moorei 

Annelida Terebellida Sabellaridae Sabellaria cementarium 

Annelida Terebellida Terebellidae 1 

Annelida Terebellida Terebellidae Scionella sp. 1 1 2 

Annelida Terebellida Terebellidae Thelepas sp. 1 

Annelida Terebellida Trichobranchidae Terebellides stroemi 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 2 9 3 2 3 3 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 1 

Arthropoda Amphipoda Dexaminidae 

Arthropoda Amphipoda Ischyroceridae 1 1 

Arthropoda Amphipoda Oedicerotidae Monoculoides sp. 

Arthropoda Amphipoda Synopiidae Tiron biocellata 
85


Arthropoda Calanoida 

Arthropoda Cumacea Diastylidae Diastylis sp. 2 

Arthropoda Cumacea Diastylopsidae Diastylopsis dawsoni 

Arthropoda Cumacea Leuconiidae Eudorella sp. 

Arthropoda Cumacea Leuconiidae Lamprops fuscata 

Arthropoda Cumacea Leuconiidae Leucon fulvus 

Arthropoda Cumacea Leuconiidae Leucon magnadentata 

Arthropoda Decapoda Pinnixidae Pinnixa sp. 

Arthropoda Isopoda Gnathidae Gnathia sp. 6 

Arthropoda Ostracoda Species I 

Arthropoda Ostracoda Species II 

Arthropoda Ostracoda Species III 

Arthropoda Tanaidacea Paratanaidea Leptognathia sp. 

Echinodermata Apodida Chiridotidae 

Echinodermata Dendrochiridotida 

Echinodermata Ophiurida Amphiuridae Amphiodia sp. 4 2 1 1 

Echiura Echiuroinea Echiuridae Echiurus echiurus alaskensis 

Mollusca Archaeogastropoda Trochidae Solariella vancouverense 3 

Mollusca Cephalaspidea Atyidae Haminoea virescens 

Mollusca Dentaliida Dentaliidae Antalis pretiosum 1 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 

Mollusca Mesogastropoda Naticidae Euspira pallida 

Mollusca Myoida Hiatellidae Hiatella arctica 

Mollusca Mytiloida Mytilidae Crenella sp. 

Mollusca Neogastropoda Buccinidae Buccinum sp. 

Mollusca Neogastropoda Cancellaridae Admete sp. 

Mollusca Neogastropoda Turridae Oenopota rosea 

Mollusca Neogastropoda Turridae Ophiodermella sp. 

Mollusca Neoloricata Ischnochitonidae Lepidozona sp. 

Mollusca Nuculoida Nuculidae Ennucula tenuis 

Mollusca Osteroida Propeomussidae Parvamussium alaskensis 

Mollusca Pholadomyoida Cuspidariidae Cardiomya pectinata 

Mollusca Pholadomyoida Pandoridae Pandora filosa 
86


Mollusca Pyramidelliacea Pyramidellidae Odostomia sp. 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 1 1 1 

Mollusca Veneroida Astartiidae Astarte esquimalti 1 

Mollusca Veneroida Carditidae Cyclocardia sp. 

Mollusca Veneroida Cultellidae Siliqua alta 

Mollusca Veneroida Malletiidae Tindaria brunea 

Mollusca Veneroida Tellinidae Macoma eliminata 3 7 1 2 5 

Mollusca Veneroida Tellinidae Macoma moesta 2 3 2 3 

Mollusca Veneroida Thyasiridae Axinospida serricata 4 4 2 5 1 

Mollusca Veneroida Thyasiridae Thyasira flexuosa 

Nemertea Heteronemertea Lineidae Cerebratulus sp. 

Nemertea Heteronemertea Lineidae Linea sp. 

Priapulida Priapulamorpha Priapulidae Priapulus caudata 1 

Sipunucula Sipunuculida Golfingiidae 

Aschelminths 1 

Total No. of Species 18 16 23 18 15 22 

Total No. of Individuals 50 55 38 30 38 36 
87 

Appendix Table 6.--Number of individuals of each infauna taxon collected per grab sample at 

Transect 8 in Trinity Islands during August 1999. 

Phylum Order Family Species 8A 8B 8C 8D 8E 8F 

Annelida 

Annelida Capitallida Capitellidae Mediomastus californiensis 6 3 6 3 1 2 

Annelida Capitallida Maldanidae 

Annelida Capitallida Maldanidae Axiothella rubrocincta 

Annelida Capitallida Maldanidae Clymenella sp. 2 1 

Annelida Capitallida Maldanidae Euclymene zonalis 

Annelida Capitallida Maldanidae Praxillella gracilis 

Annelida Capitallida Maldanidae Praxilura sp. 

Annelida Chaetopterida Chaetopteridae Spiochaetopterus costarum 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 

Annelida Eunicida Lumbrineridae Lumbrineris luti 2 1 1 2 

Annelida Eunicida Onuphidae Onuphis iridescens 1 1 1 1 

Annelida Magelonida Magelonidae Magelona berkeleyi 4 2 2 2 1 1 

Annelida Opheliida Opheliidae Travisia forbesii 

Annelida Opheliida Opheliidae Travisia japonica 

Annelida Opheliida Scalibregmidae Hyboscolex pacificus 1 

Annelida Opheliida Scalibregmidae Scalibregma inflatum 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 2 

Annelida Orbiniida Paraonidae Aricidea antennata 

Annelida Orbiniida Paraonidae Aricidea lopezi 1 1 1 2 

Annelida Orbiniida Paraonidae Chaetozone sp. 2 3 7 3 3 2 

Annelida Orbiniida Paraonidae Cirratulus sp. 1 

Annelida Orbiniida Paraonidae Levinsenia gracilis 1 

Annelida Orbiniida Paraonidae Tharyx sp. I 

Annelida Orbiniida Paraonidae Tharyx sp. II 

Annelida Phyllodocida Aphrodotidae Aphrodites japonica 

Annelida Phyllodocida Glyceridae Hemipodus borealis 1 1 1 1 

Annelida Phyllodocida Goniadidae Goniada brunnea 

Annelida Phyllodocida Hesionidae 
88


Annelida Phyllodocida Nephtidae Nephtys sp. 

Annelida Phyllodocida Nephtidae Nephtys caeca 2 5 1 2 1 1 

Annelida Phyllodocida Nephtidae Nephtys cornuta 

Annelida Phyllodocida Nereididae Nereis sp. 

Annelida Phyllodocida Pholodidae Pholoe glabra 

Annelida Phyllodocida Pholodidae Sigalona sp. 

Annelida Phyllodocida Phyllodocidae Eteone longa 1 1 

Annelida Phyllodocida Polynoidae 

Annelida Phyllodocida Polynoidae Hesperonoe sp. 2 

Annelida Phyllodocida Polynoidae Hololepida magna 

Annelida Phyllodocida Syllidae Syllis sp. 

Annelida Sabellida Sabellidae 

Annelida Sabellida Sabellidae Potamilla sp. 1 

Annelida Sabellida Serpulidae 

Annelida Spionida Spionidae Dipolydora socialis 

Annelida Spionida Spionidae Laonice sp. 

Annelida Spionida Spionidae Prionospio lighti 

Annelida Spionida Spionidae Prionospio steenstrupi 19 6 6 8 2 3 

Annelida Spionida Spionidae Pygospio elegans 1 

Annelida Spionida Spionidae Spio cirrifera 1 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 

Annelida Terebellida Pectinaridae Pectinaria moorei 

Annelida Terebellida Sabellaridae Sabellaria cementarium 

Annelida Terebellida Terebellidae 1 1 

Annelida Terebellida Terebellidae Scionella sp. 1 

Annelida Terebellida Terebellidae Thelepas sp. 1 

Annelida Terebellida Trichobranchidae Terebellides stroemi 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 7 11 1 4 1 3 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 

Arthropoda Amphipoda Dexaminidae 

Arthropoda Amphipoda Ischyroceridae 1 

Arthropoda Amphipoda Oedicerotidae Monoculoides sp. 

Arthropoda Amphipoda Synopiidae Tiron biocellata 
89


Arthropoda Calanoida 1 1 

Arthropoda Cumacea Diastylidae Diastylis sp. 

Arthropoda Cumacea Diastylopsidae Diastylopsis dawsoni 

Arthropoda Cumacea Leuconiidae Eudorella sp. 

Arthropoda Cumacea Leuconiidae Lamprops fuscata 

Arthropoda Cumacea Leuconiidae Leucon fulvus 

Arthropoda Cumacea Leuconiidae Leucon magnadentata 

Arthropoda Decapoda Pinnixidae Pinnixa sp. 

Arthropoda Isopoda Gnathidae Gnathia sp. 

Arthropoda Ostracoda Species I 

Arthropoda Ostracoda Species II 

Arthropoda Ostracoda Species III 

Arthropoda Tanaidacea Paratanaidea Leptognathia sp. 

Echinodermata Apodida Chiridotidae 

Echinodermata Dendrochiridotida 

Echinodermata Ophiurida Amphiuridae Amphiodia sp. 1 2 1 

Echiura Echiuroinea Echiuridae Echiurus echiurus alaskensis 

Mollusca Archaeogastropoda Trochidae Solariella vancouverense 1 

Mollusca Cephalaspidea Atyidae Haminoea virescens 

Mollusca Dentaliida Dentaliidae Antalis pretiosum 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 

Mollusca Mesogastropoda Naticidae Euspira pallida 

Mollusca Myoida Hiatellidae Hiatella arctica 

Mollusca Mytiloida Mytilidae Crenella sp. 

Mollusca Neogastropoda Buccinidae Buccinum sp. 

Mollusca Neogastropoda Cancellaridae Admete sp. 

Mollusca Neogastropoda Turridae Oenopota rosea 

Mollusca Neogastropoda Turridae Ophiodermella sp. 

Mollusca Neoloricata Ischnochitonidae Lepidozona sp. 

Mollusca Nuculoida Nuculidae Ennucula tenuis 

Mollusca Osteroida Propeomussidae Parvamussium alaskensis 

Mollusca Pholadomyoida Cuspidariidae Cardiomya pectinata 

Mollusca Pholadomyoida Pandoridae Pandora filosa 
90


Mollusca Pyramidelliacea Pyramidellidae Odostomia sp. 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 1 

Mollusca Veneroida Astartiidae Astarte esquimalti 

Mollusca Veneroida Carditidae Cyclocardia sp. 1 

Mollusca Veneroida Cultellidae Siliqua alta 

Mollusca Veneroida Malletiidae Tindaria brunea 

Mollusca Veneroida Tellinidae Macoma eliminata 3 6 3 2 3 6 

Mollusca Veneroida Tellinidae Macoma moesta 1 1 1 2 

Mollusca Veneroida Thyasiridae Axinospida serricata 1 5 2 1 

Mollusca Veneroida Thyasiridae Thyasira flexuosa 1 

Nemertea Heteronemertea Lineidae Cerebratulus sp. 1 

Nemertea Heteronemertea Lineidae Linea sp. 1 

Priapulida Priapulamorpha Priapulidae Priapulus caudata 

Sipunucula Sipunuculida Golfingiidae 

Aschelminths 

Total No. of Species 16 13 13 13 23 16 

Total No. of Individuals 53 42 35 34 31 29 
91 

Appendix Table 6.--Number of individuals of each infauna taxon collected per grab sample at 

Transect 9 in Trinity Islands during August 1999. 

Phylum Order Family Species 9A 9B 9C 9D 9E 9F 

Annelida 

Annelida Capitallida Capitellidae Mediomastus californiensis 10 3 1 3 1 

Annelida Capitallida Maldanidae 

Annelida Capitallida Maldanidae Axiothella rubrocincta 

Annelida Capitallida Maldanidae Clymenella sp. 

Annelida Capitallida Maldanidae Euclymene zonalis 

Annelida Capitallida Maldanidae Praxillella gracilis 

Annelida Capitallida Maldanidae Praxilura sp. 1 

Annelida Chaetopterida Chaetopteridae Spiochaetopterus costarum 1 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 

Annelida Eunicida Lumbrineridae Lumbrineris luti 1 

Annelida Eunicida Onuphidae Onuphis iridescens 1 

Annelida Magelonida Magelonidae Magelona berkeleyi 7 9 10 5 4 3 

Annelida Opheliida Opheliidae Travisia forbesii 1 

Annelida Opheliida Opheliidae Travisia japonica 

Annelida Opheliida Scalibregmidae Hyboscolex pacificus 

Annelida Opheliida Scalibregmidae Scalibregma inflatum 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 1 1 

Annelida Orbiniida Paraonidae Aricidea antennata 

Annelida Orbiniida Paraonidae Aricidea lopezi 1 3 2 1 1 

Annelida Orbiniida Paraonidae Chaetozone sp. 5 6 4 2 3 

Annelida Orbiniida Paraonidae Cirratulus sp. 

Annelida Orbiniida Paraonidae Levinsenia gracilis 3 

Annelida Orbiniida Paraonidae Tharyx sp. I 

Annelida Orbiniida Paraonidae Tharyx sp. II 

Annelida Phyllodocida Aphrodotidae Aphrodites japonica 

Annelida Phyllodocida Glyceridae Hemipodus borealis 1 

Annelida Phyllodocida Goniadidae Goniada brunnea 2 1 1 2 

Annelida Phyllodocida Hesionidae 
92


Annelida Phyllodocida Nephtidae Nephtys sp. 

Annelida Phyllodocida Nephtidae Nephtys caeca 1 2 1 1 2 

Annelida Phyllodocida Nephtidae Nephtys cornuta 1 1 1 1 

Annelida Phyllodocida Nereididae Nereis sp. 1 

Annelida Phyllodocida Pholodidae Pholoe glabra 

Annelida Phyllodocida Pholodidae Sigalona sp. 

Annelida Phyllodocida Phyllodocidae Eteone longa 

Annelida Phyllodocida Polynoidae 1 

Annelida Phyllodocida Polynoidae Hesperonoe sp. 

Annelida Phyllodocida Polynoidae Hololepida magna 

Annelida Phyllodocida Syllidae Syllis sp. 

Annelida Sabellida Sabellidae 3 

Annelida Sabellida Sabellidae Potamilla sp. 

Annelida Sabellida Serpulidae 

Annelida Spionida Spionidae Dipolydora socialis 1 

Annelida Spionida Spionidae Laonice sp. 

Annelida Spionida Spionidae Prionospio lighti 

Annelida Spionida Spionidae Prionospio steenstrupi 17 11 4 2 1 1 

Annelida Spionida Spionidae Pygospio elegans 

Annelida Spionida Spionidae Spio cirrifera 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 2 

Annelida Terebellida Pectinaridae Pectinaria moorei 

Annelida Terebellida Sabellaridae Sabellaria cementarium 

Annelida Terebellida Terebellidae 1 1 

Annelida Terebellida Terebellidae Scionella sp. 1 1 

Annelida Terebellida Terebellidae Thelepas sp. 

Annelida Terebellida Trichobranchidae Terebellides stroemi 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 4 8 11 3 2 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 1 

Arthropoda Amphipoda Dexaminidae 1 

Arthropoda Amphipoda Ischyroceridae 

Arthropoda Amphipoda Oedicerotidae Monoculoides sp. 

Arthropoda Amphipoda Synopiidae Tiron biocellata 
93


Arthropoda Calanoida 

Arthropoda Cumacea Diastylidae Diastylis sp. 

Arthropoda Cumacea Diastylopsidae Diastylopsis dawsoni 

Arthropoda Cumacea Leuconiidae Eudorella sp. 2 

Arthropoda Cumacea Leuconiidae Lamprops fuscata 

Arthropoda Cumacea Leuconiidae Leucon fulvus 

Arthropoda Cumacea Leuconiidae Leucon magnadentata 

Arthropoda Decapoda Pinnixidae Pinnixa sp. 

Arthropoda Isopoda Gnathidae Gnathia sp. 1 2 

Arthropoda Ostracoda Species I 

Arthropoda Ostracoda Species II 

Arthropoda Ostracoda Species III 

Arthropoda Tanaidacea Paratanaidea Leptognathia sp. 

Echinodermata Apodida Chiridotidae 

Echinodermata Dendrochiridotida 

Echinodermata Ophiurida Amphiuridae Amphiodia sp. 6 1 

Echiura Echiuroinea Echiuridae Echiurus echiurus alaskensis 

Mollusca Archaeogastropoda Trochidae Solariella vancouverense 

Mollusca Cephalaspidea Atyidae Haminoea virescens 

Mollusca Dentaliida Dentaliidae Antalis pretiosum 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 

Mollusca Mesogastropoda Naticidae Euspira pallida 

Mollusca Myoida Hiatellidae Hiatella arctica 

Mollusca Mytiloida Mytilidae Crenella sp. 

Mollusca Neogastropoda Buccinidae Buccinum sp. 

Mollusca Neogastropoda Cancellaridae Admete sp. 

Mollusca Neogastropoda Turridae Oenopota rosea 

Mollusca Neogastropoda Turridae Ophiodermella sp. 

Mollusca Neoloricata Ischnochitonidae Lepidozona sp. 

Mollusca Nuculoida Nuculidae Ennucula tenuis 

Mollusca Osteroida Propeomussidae Parvamussium alaskensis 

Mollusca Pholadomyoida Cuspidariidae Cardiomya pectinata 

Mollusca Pholadomyoida Pandoridae Pandora filosa 
94


Mollusca Pyramidelliacea Pyramidellidae Odostomia sp. 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 

Mollusca Veneroida Astartiidae Astarte esquimalti 

Mollusca Veneroida Carditidae Cyclocardia sp. 1 

Mollusca Veneroida Cultellidae Siliqua alta 

Mollusca Veneroida Malletiidae Tindaria brunea 

Mollusca Veneroida Tellinidae Macoma eliminata 3 3 2 3 

Mollusca Veneroida Tellinidae Macoma moesta 1 1 

Mollusca Veneroida Thyasiridae Axinospida serricata 6 1 3 2 

Mollusca Veneroida Thyasiridae Thyasira flexuosa 

Nemertea Heteronemertea Lineidae Cerebratulus sp. 

Nemertea Heteronemertea Lineidae Linea sp. 

Priapulida Priapulamorpha Priapulidae Priapulus caudata 

Sipunucula Sipunuculida Golfingiidae 1 

Aschelminths 

Total No. of Species 19 10 15 14 14 10 

Total No. of Individuals 74 45 43 26 22 19 
95 

Appendix Table 6.--Number of individuals of each infauna taxon collected per grab sample at 

Transect 10 in Trinity Islands during August 1999. 

Phylum Order Family Species 10A 10B 10C 10D 10E 10F 

Annelida 

Annelida Capitallida Capitellidae Mediomastus californiensis 5 2 4 2 1 2 

Annelida Capitallida Maldanidae 

Annelida Capitallida Maldanidae Axiothella rubrocincta 

Annelida Capitallida Maldanidae Clymenella sp. 

Annelida Capitallida Maldanidae Euclymene zonalis 

Annelida Capitallida Maldanidae Praxillella gracilis 

Annelida Capitallida Maldanidae Praxilura sp. 

Annelida Chaetopterida Chaetopteridae Spiochaetopterus costarum 1 2 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 1 

Annelida Eunicida Lumbrineridae Lumbrineris luti 1 3 

Annelida Eunicida Onuphidae Onuphis iridescens 1 

Annelida Magelonida Magelonidae Magelona berkeleyi 4 3 1 1 4 

Annelida Opheliida Opheliidae Travisia forbesii 1 1 

Annelida Opheliida Opheliidae Travisia japonica 

Annelida Opheliida Scalibregmidae Hyboscolex pacificus 

Annelida Opheliida Scalibregmidae Scalibregma inflatum 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 2 2 

Annelida Orbiniida Paraonidae Aricidea antennata 1 

Annelida Orbiniida Paraonidae Aricidea lopezi 5 5 1 2 

Annelida Orbiniida Paraonidae Chaetozone sp. 16 4 1 4 7 

Annelida Orbiniida Paraonidae Cirratulus sp. 

Annelida Orbiniida Paraonidae Levinsenia gracilis 1 1 

Annelida Orbiniida Paraonidae Tharyx sp. I 

Annelida Orbiniida Paraonidae Tharyx sp. II 1 

Annelida Phyllodocida Aphrodotidae Aphrodites japonica 

Annelida Phyllodocida Glyceridae Hemipodus borealis 2 2 1 

Annelida Phyllodocida Goniadidae Goniada brunnea 1 1 1 

Annelida Phyllodocida Hesionidae 
96


Annelida Phyllodocida Nephtidae Nephtys sp. 

Annelida Phyllodocida Nephtidae Nephtys caeca 2 4 3 2 2 

Annelida Phyllodocida Nephtidae Nephtys cornuta 1 1 1 1 

Annelida Phyllodocida Nereididae Nereis sp. 

Annelida Phyllodocida Pholodidae Pholoe glabra 

Annelida Phyllodocida Pholodidae Sigalona sp. 

Annelida Phyllodocida Phyllodocidae Eteone longa 

Annelida Phyllodocida Polynoidae 

Annelida Phyllodocida Polynoidae Hesperonoe sp. 1 

Annelida Phyllodocida Polynoidae Hololepida magna 

Annelida Phyllodocida Syllidae Syllis sp. 

Annelida Sabellida Sabellidae 1 1 

Annelida Sabellida Sabellidae Potamilla sp. 1 1 

Annelida Sabellida Serpulidae 

Annelida Spionida Spionidae Dipolydora socialis 1 

Annelida Spionida Spionidae Laonice sp. 

Annelida Spionida Spionidae Prionospio lighti 

Annelida Spionida Spionidae Prionospio steenstrupi 31 37 7 9 8 7 

Annelida Spionida Spionidae Pygospio elegans 

Annelida Spionida Spionidae Spio cirrifera 1 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 

Annelida Terebellida Pectinaridae Pectinaria moorei 

Annelida Terebellida Sabellaridae Sabellaria cementarium 1 

Annelida Terebellida Terebellidae 1 

Annelida Terebellida Terebellidae Scionella sp. 1 

Annelida Terebellida Terebellidae Thelepas sp. 

Annelida Terebellida Trichobranchidae Terebellides stroemi 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 25 8 21 5 5 1 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 1 

Arthropoda Amphipoda Dexaminidae 

Arthropoda Amphipoda Ischyroceridae 1 

Arthropoda Amphipoda Oedicerotidae Monoculoides sp. 1 

Arthropoda Amphipoda Synopiidae Tiron biocellata 1 
97


Arthropoda Calanoida 

Arthropoda Cumacea Diastylidae Diastylis sp. 1 

Arthropoda Cumacea Diastylopsidae Diastylopsis dawsoni 

Arthropoda Cumacea Leuconiidae Eudorella sp. 1 

Arthropoda Cumacea Leuconiidae Lamprops fuscata 

Arthropoda Cumacea Leuconiidae Leucon fulvus 

Arthropoda Cumacea Leuconiidae Leucon magnadentata 

Arthropoda Decapoda Pinnixidae Pinnixa sp. 

Arthropoda Isopoda Gnathidae Gnathia sp. 1 2 

Arthropoda Ostracoda Species I 

Arthropoda Ostracoda Species II 

Arthropoda Ostracoda Species III 

Arthropoda Tanaidacea Paratanaidea Leptognathia sp. 

Echinodermata Apodida Chiridotidae 

Echinodermata Dendrochiridotida 

Echinodermata Ophiurida Amphiuridae Amphiodia sp. 1 1 

Echiura Echiuroinea Echiuridae Echiurus echiurus alaskensis 1 

Mollusca Archaeogastropoda Trochidae Solariella vancouverense 

Mollusca Cephalaspidea Atyidae Haminoea virescens 1 

Mollusca Dentaliida Dentaliidae Antalis pretiosum 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 

Mollusca Mesogastropoda Naticidae Euspira pallida 1 

Mollusca Myoida Hiatellidae Hiatella arctica 

Mollusca Mytiloida Mytilidae Crenella sp. 

Mollusca Neogastropoda Buccinidae Buccinum sp. 

Mollusca Neogastropoda Cancellaridae Admete sp. 

Mollusca Neogastropoda Turridae Oenopota rosea 

Mollusca Neogastropoda Turridae Ophiodermella sp. 

Mollusca Neoloricata Ischnochitonidae Lepidozona sp. 

Mollusca Nuculoida Nuculidae Ennucula tenuis 

Mollusca Osteroida Propeomussidae Parvamussium alaskensis 

Mollusca Pholadomyoida Cuspidariidae Cardiomya pectinata 

Mollusca Pholadomyoida Pandoridae Pandora filosa 1 
98


Mollusca Pyramidelliacea Pyramidellidae Odostomia sp. 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 1 

Mollusca Veneroida Astartiidae Astarte esquimalti 

Mollusca Veneroida Carditidae Cyclocardia sp. 

Mollusca Veneroida Cultellidae Siliqua alta 

Mollusca Veneroida Malletiidae Tindaria brunea 1 

Mollusca Veneroida Tellinidae Macoma eliminata 1 3 1 6 5 

Mollusca Veneroida Tellinidae Macoma moesta 2 1 3 1 

Mollusca Veneroida Thyasiridae Axinospida serricata 2 4 2 1 1 

Mollusca Veneroida Thyasiridae Thyasira flexuosa 

Nemertea Heteronemertea Lineidae Cerebratulus sp. 

Nemertea Heteronemertea Lineidae Linea sp. 1 

Priapulida Priapulamorpha Priapulidae Priapulus caudata 

Sipunucula Sipunuculida Golfingiidae 

Aschelminths 

Total No. of Species 18 23 18 15 15 13 

Total No. of Individuals 103 84 53 36 39 28 
99 

Appendix Table 6. -- Number of individuals of each infauna taxon collected per grab sample at 

Transect 11 in Trinity Islands during August 1999. 

Phylum Order Family Species 11A 11B 11C 11D 11E 11F 

Annelida 

Annelida Capitallida Capitellidae Mediomastus californiensis 3 4 3 4 4 

Annelida Capitallida Maldanidae Axiothella rubrocincta 1 

Annelida Capitallida Maldanidae Clymenella sp. 

Annelida Capitallida Maldanidae 

Annelida Capitallida Maldanidae Euclymene zonalis 1 

Annelida Capitallida Maldanidae Praxillella gracilis 

Annelida Capitallida Maldanidae Praxilura sp. 

Annelida Chaetopterida Chaetopteridae Spiochaetopterus costarum 

Annelida Eunicida Lumbrineridae Lumbrineris bicirrata 

Annelida Eunicida Lumbrineridae Lumbrineris luti 1 1 

Annelida Eunicida Onuphidae Onuphis iridescens 1 1 

Annelida Magelonida Magelonidae Magelona berkeleyi 1 1 1 2 3 2 

Annelida Opheliida Opheliidae Travisia forbesii 1 

Annelida Opheliida Opheliidae Travisia japonica 

Annelida Opheliida Scalibregmidae Hyboscolex pacificus 

Annelida Opheliida Scalibregmidae Scalibregma inflatum 

Annelida Orbiniida Orbiniidae Leitoscoloplos pugettensis 2 1 1 1 

Annelida Orbiniida Paraonidae Aricidea antennata 

Annelida Orbiniida Paraonidae Aricidea lopezi 1 3 

Annelida Orbiniida Paraonidae Chaetozone sp. 8 4 2 9 4 

Annelida Orbiniida Paraonidae Cirratulus sp. 

Annelida Orbiniida Paraonidae Levinsenia gracilis 

Annelida Orbiniida Paraonidae Tharyx sp. I 1 

Annelida Orbiniida Paraonidae Tharyx sp. II 

Annelida Phyllodocida Aphrodotidae Aphrodites japonica 1 

Annelida Phyllodocida Glyceridae Hemipodus borealis 

Annelida Phyllodocida Goniadidae Goniada brunnea 2 1 1 1 2 

Annelida Phyllodocida Hesionidae 
100


Annelida Phyllodocida Nephtidae Nephtys sp. 

Annelida Phyllodocida Nephtidae Nephtys caeca 1 2 1 2 

Annelida Phyllodocida Nephtidae Nephtys cornuta 1 1 5 1 

Annelida Phyllodocida Nereididae Nereis sp. 

Annelida Phyllodocida Pholodidae Pholoe glabra 1 

Annelida Phyllodocida Pholodidae Sigalona sp. 

Annelida Phyllodocida Phyllodocidae Eteone longa 1 1 

Annelida Phyllodocida Polynoidae 

Annelida Phyllodocida Polynoidae Hesperonoe sp. 

Annelida Phyllodocida Polynoidae Hololepida magna 

Annelida Phyllodocida Syllidae Syllis sp. 

Annelida Sabellida Sabellidae 1 1 

Annelida Sabellida Sabellidae Potamilla sp. 

Annelida Sabellida Serpulidae 

Annelida Spionida Spionidae Dipolydora socialis 

Annelida Spionida Spionidae Laonice sp. 1 

Annelida Spionida Spionidae Prionospio lighti 1 

Annelida Spionida Spionidae Prionospio steenstrupi 12 4 4 13 11 9 

Annelida Spionida Spionidae Pygospio elegans 

Annelida Spionida Spionidae Spio cirrifera 

Annelida Terebellida Amphaeretidae Amphicteis scaphobranchiata 1 1 

Annelida Terebellida Pectinaridae Pectinaria moorei 

Annelida Terebellida Sabellaridae Sabellaria cementarium 

Annelida Terebellida Terebellidae 1 

Annelida Terebellida Terebellidae Scionella sp. 1 1 

Annelida Terebellida Terebellidae Thelepas sp. 1 

Annelida Terebellida Trichobranchidae Terebellides stroemi 

Arthropoda Amphipoda Ampeliscidae Ampelisca cf. unsocalae 2 1 4 7 

Arthropoda Amphipoda Ampeliscidae Byblis veleronis 2 

Arthropoda Amphipoda Dexaminidae 

Arthropoda Amphipoda Ischyroceridae 

Arthropoda Amphipoda Oedicerotidae Monoculoides sp. 1 1 

Arthropoda Amphipoda Synopiidae Tiron biocellata 
101


Arthropoda Calanoida 

Arthropoda Cumacea Diastylidae Diastylis sp. 1 

Arthropoda Cumacea Diastylopsidae Diastylopsis dawsoni 1 

Arthropoda Cumacea Leuconiidae Eudorella sp. 1 1 

Arthropoda Cumacea Leuconiidae Lamprops fuscata 

Arthropoda Cumacea Leuconiidae Leucon fulvus 

Arthropoda Cumacea Leuconiidae Leucon magnadentata 

Arthropoda Decapoda Pinnixidae Pinnixa sp. 

Arthropoda Isopoda Gnathidae Gnathia sp. 1 

Arthropoda Ostracoda Species I 

Arthropoda Ostracoda Species II 

Arthropoda Ostracoda Species III 

Arthropoda Tanaidacea Paratanaidea Leptognathia sp. 

Echinodermata Apodida Chiridotidae 

Echinodermata Dendrochiridotida 

Echinodermata Ophiurida Amphiuridae Amphiodia sp. 3 

Echiura Echiuroinea Echiuridae Echiurus echiurus alaskensis 1 2 

Mollusca Archaeogastropoda Trochidae Solariella vancouverense 

Mollusca Cephalaspidea Atyidae Haminoea virescens 1 

Mollusca Dentaliida Dentaliidae Antalis pretiosum 

Mollusca Dentaliida Dentaliidae Rhabdus rectius 1 

Mollusca Mesogastropoda Naticidae Euspira pallida 

Mollusca Myoida Hiatellidae Hiatella arctica 

Mollusca Mytiloida Mytilidae Crenella sp. 

Mollusca Neogastropoda Buccinidae Buccinum sp. 1 

Mollusca Neogastropoda Cancellaridae Admete sp. 

Mollusca Neogastropoda Turridae Oenopota rosea 

Mollusca Neogastropoda Turridae Ophiodermella sp. 

Mollusca Neoloricata Ischnochitonidae Lepidozona sp. 

Mollusca Nuculoida Nuculidae Ennucula tenuis 

Mollusca Osteroida Propeomussidae Parvamussium alaskensis 

Mollusca Pholadomyoida Cuspidariidae Cardiomya pectinata 

Mollusca Pholadomyoida Pandoridae Pandora filosa 
102


Mollusca Pyramidelliacea Pyramidellidae Odostomia sp. 

Mollusca Pyramidelliacea Pyramidellidae Turbonilla sp. 

Mollusca Veneroida Astartiidae Astarte esquimalti 

Mollusca Veneroida Carditidae Cyclocardia sp. 

Mollusca Veneroida Cultellidae Siliqua alta 

Mollusca Veneroida Malletiidae Tindaria brunea 

Mollusca Veneroida Tellinidae Macoma eliminata 4 5 1 1 11 

Mollusca Veneroida Tellinidae Macoma moesta 1 3 2 

Mollusca Veneroida Thyasiridae Axinospida serricata 1 1 2 2 2 

Mollusca Veneroida Thyasiridae Thyasira flexuosa 

Nemertea Heteronemertea Lineidae Cerebratulus sp. 

Nemertea Heteronemertea Lineidae Linea sp. 1 

Priapulida Priapulamorpha Priapulidae Priapulus caudata 

Sipunucula Sipunuculida Golfingiidae 

Aschelminths 

Total No. of Species 14 23 14 10 18 16 

Total No. of Individuals 35 35 30 26 50 53 
103 

CITATIONS 

Blake, J. A. 1995. Family Pholodidae, p. 175-188. In J. A. Blake, B. Hilbig, and P. V. Scott 

(editors), Taxonomic atlas of the benthic fauna of the Santa Maria Basin and western 

Santa Barbara Channel, volume 5--The Annelida, part 2--Polychaeta: Phyllodocida 

(Syllidae and scale-bearing families), Amphinomida, and Eunicida. Santa Barbara 

Museum of Natural History, Santa Barbara, California. 

Blake, J. A. 1996. Family Cirratulidae, p. 263-384. In J. A. Blake, B. Hilbig, and P. V. Scott 

(editors), Taxonomic atlas of the benthic fauna of the Santa Maria Basin and western 

Santa Barbara Channel, volume 6--The Annelida, part 3--Polychaeta: Orbiniidae to 

Cossuridae. Santa Barbara Museum of Natural History, Santa Barbara, California. 

Blake, J. A. 1996. Family Magelonidae, p. 253-262. In J. A. Blake, B. Hilbig, and P.V. Scott 

(editors), Taxonomic atlas of the benthic fauna of the Santa Maria Basin and western 

Santa Barbara Channel, volume 6--The Annelida, part 3--Polychaeta: Orbiniidae to 

Cossuridae. Santa Barbara Museum of Natural History, Santa Barbara, California. 

Blake, J. A. 1996. Family Orbiniidae, p. 1-26. In J. A. Blake, B. Hilbig, and P.V. Scott 

(editors), Taxonomic atlas of the benthic fauna of the Santa Maria Basin and western 

Santa Barbara Channel, volume 6--The Annelida, part 3--Polychaeta: Orbiniidae to 

Cossuridae. Santa Barbara Museum of Natural History, Santa Barbara, California. 

Blake, J. A. 1996. Family Paraonidae, p. 27-80. In J. A. Blake, B. Hilbig, and P.V. Scott 

(editors), Taxonomic atlas of the benthic fauna of the Santa Maria Basin and western 

Santa Barbara Channel, volume 6--The Annelida, part 3--Polychaeta: Orbiniidae to 

Cossuridae. Santa Barbara Museum of Natural History, Santa Barbara, California. 
104


Blake, J. A. 1996. Family Spionidae, p. 81-232. In J. A. Blake, B. Hilbig, and P.V. Scott 

(editors), Taxonomic atlas of the benthic fauna of the Santa Maria Basin and western 

Santa Barbara Channel, volume 6--The Annelida, part 3--Polychaeta: Orbiniidae to 

Cossuridae. Santa Barbara Museum of Natural History, Santa Barbara, California. 

Blake, J. A. 1997. Family Phyllodocidae, p. 109-178. In J. A. Blake, B. Hilbig, and P.V. 

Scott (editors), Taxonomic atlas of the benthic fauna of the Santa Maria Basin and 

western Santa Barbara Channel, volume 4--The Annelida, part 1--Oligochaeta and 

Polychaeta: Phyllodocida (Phyllodocidae to Paralacydoniiae). Santa Barbara Museum 

of Natural History, Santa Barbara, California. 

Blake, J. A. 2000. Family Capitellidae, p. 47-96. In J. A. Blake, B. Hilbig, and P.V. Scott 

(editors), Taxonomic atlas of the benthic fauna of the Santa Maria Basin and western 

Santa Barbara Channel, volume 7--The Annelida, part 4--Polychaeta: Flabelligeridae to 

Sternapsidae. Santa Barbara Museum of Natural History, Santa Barbara, California. 

Blake, J. A. 2000. Family Flabelligeridae, p. 1-24. In J. A. Blake, B. Hilbig, and P.V. Scott 

(editors), Taxonomic atlas of the benthic fauna of the Santa Maria Basin and western 

Santa Barbara Channel, volume 7--The Annelida, part 4--Polychaeta: Flabelligeridae to 

Sternapsidae. Santa Barbara Museum of Natural History, Santa Barbara, California. 

Blake, J. A. 2000. Family Opheliidae, p. 145-168. In J. A. Blake, B. Hilbig, and P.V. Scott 

(editors), Taxonomic atlas of the benthic fauna of the Santa Maria Basin and western 

Santa Barbara Channel, volume 7--The Annelida, part 4--Polychaeta: Flabelligeridae to 

Sternapsidae. Santa Barbara Museum of Natural History, Santa Barbara, California. 
105


Blake, J. A. 2000. Family Oweniidae, p. 97-128. In J. A. Blake, B. Hilbig, and P.V. Scott 

(editors), Taxonomic atlas of the benthic fauna of the Santa Maria Basin and western 

Santa Barbara Channel, volume 7--The Annelida, part 4--Polychaeta: Flabelligeridae to 

Sternapsidae. Santa Barbara Museum of Natural History, Santa Barbara, California. 

Blake, J. A. 2000. Family Scalibregmatidae, p. 129-144. In J. A. Blake, B. Hilbig, and P.V. 

Scott (editors), Taxonomic atlas of the benthic fauna of the Santa Maria Basin and 

western Santa Barbara Channel, volume 7--The Annelida, part 4--Polychaeta: 

Flabelligeridae to Sternapsidae. Santa Barbara Museum of Natural History, Santa 

Barbara, California. 

Bousfield, E. L., and J. A. Kendall. 1994. The amphipod superfamily Dexaminoidea on the 

North American Pacific coast; families Atylidae and Dexaminidae: systematics and 

distributional ecology. Amphipacifica 1:3-66. 

Butler, T. H. 1980. Shrimps of the Pacific Coast of Canada. Can. Bull. Fish. Aquat. Sci., No. 

202, 280 p. 

Conlan, K. E. 1997. The superfamily Corophioidea, p. 177-223. In J. A. Blake and P. V. 

Scott (editors), Taxonomic atlas of the benthic fauna of the Santa Maria Basin and 

western Santa Barbara Channel, volume 12--The Crustacea, part 3--The Amphipoda. 

Santa Barbara Museum of Natural History, Santa Barbara, California. 

Coyle, K. O. 1982. The amphipod genus Grandifoxus Barnard (Gammaridea: 

Phoxocephalidae) in Alaska. J. Crustacean Biol. 2:430-450. 

Derjavin, A. N. 1926. The Cumacea of the Kamtchatka Expedition. Russische 

Hydrobiologische Zeitschrift 9:174-181. 
106


Fauchald, K. 1977. The polychaete worms--definitions and keys to the orders, families, and 

genera. Natural History Museum of Los Angeles County Scientific Series No. 28, 

188 p. 

Fauchald, K. 1982. Revision of Onuphis, Nothria, and Paradiopatra (Polychaeta: Onuphidae) 

based upon type material. Smithson. Contrib. Zool. 356:1-109. 

Foster, N. R. 1991. Intertidal bivalves: A guide to the common marine bivalves of Alaska. 

University of Alaska Press, Fairbanks, Alaska, 152 p. 

Griffith, L. M. 1967. The intertidal univalves of British Columbia. British Columbia 

Provincial Museum, Handbook 26, Victoria, British Columbia, 101 p. 

Harbo, R. M. 1997. Shells and shellfish of the Pacific Northwest: A field guide. Harbour 

Publishing Madeira Park, British Columbia, Canada, 270 p. 

Hart, J. F. L. 1930. Some Cumacea of the Vancouver Island region. Contributions to 

Canadian Biology and Fisheries 6:25-40. 

Hart, J. F. L. 1982. Crabs and their relatives of British Columbia. British Columbia 

Provincial Museum, Handbook 40, Victoria, British Columbia, 266 p. 

Hilbig B. 1995. Family Onuphidae, p. 229-262. In J. A. Blake, B. Hilbig, and P.V. Scott 

(editors), Taxonomic atlas of the benthic fauna of the Santa Maria Basin and western 

Santa Barbara Channel, volume 4--The Annelida, part 2--Polychaeta: Phyllodocida 

(Syllidae and scale-bearing families), Amphinomida, and Eunicida. Santa Barbara 

Museum of Natural History, Santa Barbara, California. 
107


Hilbig B. 1995. Family Lumbrineridae, p. 279-314. In J. A. Blake, B. Hilbig, and P.V. Scott 

(editors), Taxonomic atlas of the benthic fauna of the Santa Maria Basin and western 

Santa Barbara Channel, volume 4--The Annelida, part 2--Polychaeta: Phyllodocida 

(Syllidae and scale-bearing families), Amphinomida, and Eunicida. Santa Barbara 

Museum of Natural History, Santa Barbara, California. 

Hilbig, B. 1997. Family Glyceridae, p. 187-206. In J. A. Blake, B. Hilbig, and P.V. Scott 

(editors), Taxonomic atlas of the benthic fauna of the Santa Maria Basin and western 

Santa Barbara Channel, volume 4--The Annelida, part 1--Oligochaeta and Polychaeta: 

Phyllodocida (Phyllodocidae to Paralacydoniiae). Santa Barbara Museum of Natural 

History, Santa Barbara, California. 

Hilbig, B. 1997. Family Goniadidae, p. 207-222. In J. A. Blake, B. Hilbig, and P.V. Scott 

(editors), Taxonomic atlas of the benthic fauna of the Santa Maria Basin and western 

Santa Barbara Channel, volume 4--The Annelida, part 1--Oligochaeta and Polychaeta: 

Phyllodocida (Phyllodocidae to Paralacydoniiae). Santa Barbara Museum of Natural 

History, Santa Barbara, California. 

Hilbig, B. 1997. Family Nephtyidae, p. 317-350. In J. A. Blake, B. Hilbig, and P.V. Scott 

(editors), Taxonomic atlas of the benthic fauna of the Santa Maria Basin and western 

Santa Barbara Channel, volume 4--The Annelida, part 1--Oligochaeta and Polychaeta: 

Phyllodocida (Phyllodocidae to Paralacydoniiae). Santa Barbara Museum of Natural 

History, Santa Barbara, California. 
108


Hilbig, B. 2000. Family Ampharetidae, p. 169-230. In J. A. Blake, B. Hilbig, and P.V. Scott 

(editors), Taxonomic atlas of the benthic fauna of the Santa Maria Basin and western 

Santa Barbara Channel, volume 7--The Annelida, part 4--Polychaeta: Flabelligeridae to 

Sternapsidae. Santa Barbara Museum of Natural History, Santa Barbara, California. 

Hilbig, B. 2000. Family Terebellidae, p. 231-230. In J. A. Blake, B. Hilbig, and P.V. Scott 

(editors), Taxonomic atlas of the benthic fauna of the Santa Maria Basin and western 

Santa Barbara Channel, volume 7--The Annelida, part 4--Polychaeta: Flabelligeridae to 

Sternapsidae. Santa Barbara Museum of Natural History, Santa Barbara, California. 

Hilbig, B. 2000. Family Trichobranchidae, p. 295-310. In J. A. Blake, B. Hilbig, and P.V. 

Scott (editors), Taxonomic atlas of the benthic fauna of the Santa Maria Basin and 

western Santa Barbara Channel, volume 7--The Annelida, part 4--Polychaeta: 

Flabelligeridae to Sternapsidae. Santa Barbara Museum of Natural History, Santa 

Barbara, California. 

Hobson, K. D., and K. Banse. 1981. Sedentariate and archiannelid polychaetes of British 

Columbia and Washington. Can. Bull. Fish. Aquat. Sci. No. 209, 144 p. 

Hong, J. S., and M. R. Park. 1999. The genus Eudorella (Crustacea: Cumacea) from the 

Yellow Sea, with description of a new species. J. Mar. Biol. Assoc. U. K. 79:445-457. 

Jarrett, N. N., and E. L. Bousfield. 1994a. The amphipod superfamily Phoxocephaloidea on 

the Pacific Coast of North America. Family Phoxocephalidae. Part I. Metharpiniidae, 

new subfamily. Amphipacifica 1:58-140. 
109


Jarrett, N. N., and E. L. Bousfield. 1994b. The amphipod superfamily Phoxocephaloidea on 

the Pacific Coast of North America. Family Phoxocephalidae. Part II. Subfamilies 

Pontharpiniinae, Parharpiniinae, Brolginae, Phoxocephalinae, and Harpiniinae. 

Systematics and ecology. Amphipacifica 1:71-150. 

Kessler, D. W. 1985. Alaska’s saltwater fishes and other sea life. Alaska Northwest 

Publishing Company, Anchorage, Alaska, 358 p. 

Kozloff, E. N. 1996. Marine invertebrates of the Pacific Northwest with additions and 

corrections. University of Washington Press, Seattle, Washington, 539 p. 

Kudenov, J. D., and L. H. Harris. 1995. Family Syllidae. p. 1-98. In J. A. Blake, B. Hilbig, 

and P.V. Scott (editors), Taxonomic atlas of the benthic fauna of the Santa Maria Basin 

and western Santa Barbara Channel, volume 4--The Annelida, part 2--Polychaeta: 

Phyllodocida (Syllidae and scale-bearing families), Amphinomida, and Eunicida. Santa 

Barbara Museum of Natural History, Santa Barbara, California. 

Martin, J. W., and D. L. Zmarzly. 1994. Pinnixa scamit, a new species of pinnotherid crab 

(Decapoda: Brachyura) from the continental slope off California. Proc. Biol. Soc. 

Wash. 107:354-359. 

Pearson, T. H., J. S. Gray, and P. J. Johannessen. 1983. Objective selection of sensitive 

species indicative of pollution-induced change in benthic communities. 2. Data 

analyses. Mar. Ecol. Prog. Ser. 12:237-255. 

Quayle, D. B. 1970. Intertidal bivalves of British Columbia. Second edition. British 

Columbia Provincial Museum, Handbook 17, Victoria, British Columbia, 104 p. 
110


Rice, T. C. 1971. Marine shells of the Pacific Northwest. Ellison Industries, Edmunds, 

Washington, 102 p. 

Roccatagliata D., and R. W. Heard. 1992. Diastylopsis goekei, a new species (Crustacea, 

Cumacea, Diastylidae) from Antarctic waters. Proc. Biol. Soc. Wash. 105:743-752. 

Thomas, J. D., and L. D. McCann. 1997. The families Argissidae, Dexaminidae, Eusiridae, 

Gammaridae, Leucothoidae, Melphidippidae, Oedicerotidae, Pardaliscidae, 

Phoxocephalidae, Podoceridae, Stegocephalidae, Stenothoidae, Stilipedidae, 

Synopiidae, and Urothoidae, p. 21-136. In J. A. Blake and P. V. Scott (editors), 

Taxonomic atlas of the benthic fauna of the Santa Maria Basin and western Santa 

Barbara Channel, volume 12--The Crustacea, part 3--The Amphipoda. Santa Barbara 

Museum of Natural History, Santa Barbara, California. 

Watling, L. 1991. Revision of the cumacean family Leuconidae. J. Crustacean Biol. 

11:569-582. 

Watling, L. 1997. The families Ampeliscidae, Amphilochidae, Liljeborgiidae, and Pleustidae, 

p. 137-176. In J. A. Blake and P. V. Scott (editors), Taxonomic atlas of the benthic 

fauna of the Santa Maria Basin and western Santa Barbara Channel, volume 12--The 

Crustacea, part 3--The Amphipoda. Santa Barbara Museum of Natural History, Santa 

Barbara, California. 
111 

Watling, L., and L. D. McCann. 1997. Descriptions of the species of the suborders 

Anthuridea, Epicaridea, Flabellifera, Gnathiidea, and Valvifera, p. 9-58. In J. A. Blake 

and P. V. Scott (editors), Taxonomic atlas of the benthic fauna of the Santa Maria Basin 

and western Santa Barbara Channel, volume 11--The Crustacea, part 2--The Isopoda, 

Cumacea, and Tanaidacea. Santa Barbara Museum of Natural History, Santa Barbara, 

California. 

Wetzer, R., and R. C. Brusca. 1997. Cumacea, p. 121-178. In J. A. Blake and P. V. Scott 

(editors), Taxonomic atlas of the benthic fauna of the Santa Maria Basin and western 

Santa Barbara Channel, volume 11--The Crustacea, part 2--The Isopoda, Cumacea, and 

Tanaidacea, Santa Barbara Museum of Natural History, Santa Barbara, California. 

Zmarzly, D. L. 1992. Taxonomic review of pea crabs in the genus Pinnixa (Decapoda: 

Brachyura: Pinnotheridae) occurring on the California shelf, with descriptions of two 

new species. J. Crustacean Biol. 12:677-713. 
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