by

S. W. Johnson, A. D. Neff, 

J. F. Thedinga, M. R. Lindeberg, 

and J. M. Maselko

NOAA Technical Memorandum NMFS-AFSC-239

Atlas of Nearshore Fishes of Alaska: 

A Synthesis of Marine Surveys from 1998 to 2011

October 2012 U.S. Department of Commerce | National Oceanic and Atmospheric Administration | National Marine Fisheries Service
NOAA Technical Memorandum NMFS

The National Marine Fisheries Service's Alaska Fisheries Science Center

uses the NOAA Technical Memorandum series to issue informal scientific and

technical publications when complete formal review and editorial processing

are not appropriate or feasible. Documents within this series reflect sound

professional work and may be referenced in the formal scientific and technical

literature.

The NMFS-AFSC Technical Memorandum series of the Alaska Fisheries

Science Center continues the NMFS-F/NWC series established in 1970 by the

Northwest Fisheries Center. The NMFS-NWFSC series is currently used by

the Northwest Fisheries Science Center.

This document should be cited as follows:

Johnson, S. W., A. D. Neff, J. F. Thedinga, M. R. Lindeberg, and

J. M. Maselko. 2012. Atlas of nearshore fishes of Alaska: A synthesis

of marine surveys from 1998 to 2011. U.S. Dep. Commer., NOAA

Tech. Memo. NMFS-AFSC-239, 261 p.

Reference in this document to trade names does not imply endorsement by

the National Marine Fisheries Service, NOAA.
 October 2012

NOAA Technical Memorandum NMFS-AFSC-239 

by

 S. W. Johnson, A. D. Neff, J. F. Thedinga, 

M. R. Lindeberg, and J. M. Maselko 

Atlas of Nearshore Fishes of Alaska: 

A Synthesis of Marine Surveys 

 from 1998 to 2011

Alaska Fisheries Science Center

Auke Bay Laboratories

17109 Lena Point Loop Road

Juneau, AK 99801

www.afsc.noaa.gov

U.S. DEPARTMENT OF COMMERCE

Rebecca M. Blank, Acting Secretary

National Oceanic and Atmospheric Administration

 Jane Lubchenco, Under Secretary and Administrator

National Marine Fisheries Service

Eric C. Schwaab, Assistant Administrator for Fisheries 
This document is available to the public through: 

National Technical Information Service 

U.S. Department of Commerce 

5285 Port Royal Road 

Springfield, VA 22161 

www.ntis.gov 
Sampling fishes with a seine in nearshore waters of western Prince William Sound, 

Alaska, in April 2006. 

 
 
iii 

ABSTRACT 

Information on the distribution, abundance, species composition, habitat use, and lifestage of 121 fish species caught in nearshore marine waters of Alaska is synthesized in this 

atlas. Data were collected by scientists from the Alaska Fisheries Science Center, Auke Bay 

Laboratories over a 14-year period (1998-2011). Fish were captured with a beach seine 

mostly in summer in four different habitat types (bedrock outcrops, eelgrass meadows, 

understory kelp beds, and sand or gravel beaches) at 93 locations and 555 unique sites within 

nine regions of Alaska spanning from the Arctic to southeastern Alaska. Data on species 

assemblages are summarized among regions and habitat types. Distribution maps, graphs of 

abundance and length frequency, and photos are presented for each species. This atlas is 

unique because the same methods were used to sample fish and habitat over a large extent of 

coastal Alaska, and the core group of researchers doing the work remained the same 

throughout all years. 

This synthesis reveals a wealth of information about nearshore fishes of Alaska. Our 

major findings include: 

 1. Nearshore marine waters support an abundant and diverse array of fishes; we 

captured an estimated 718,345 fish representing at least 121 species from 29 

families. 

 2. Four commercially important species (walleye pollock, Theragra chalcogramma; 

Pacific herring, Clupea pallasii; pink salmon, Oncorhynchus gorbuscha; and chum 

salmon, O. keta) accounted for 55% of our total overall catch. 

 3. Species distribution patterns varied greatly among regions; only two species (pink 

salmon and threespine stickleback, Gasterosteus aculeatus) were caught in all nine 

regions. 

 4. Abundance and species richness varied by region; mean catch-per-unit-effort 

(CPUE) ranged from 124 fish in Southcentral Alaska to 1,202 fish in the Aleutian 

Islands, and species richness ranged from 17 species in Bristol Bay to 67 species in 

Southeast – Northern Inside. 

 5. Species assemblages differed among habitat types in most regions, and mean 

CPUE and species richness were usually greatest in eelgrass or kelp. 
iv 

 6. Species composition by habitat type was unique in some regions (e.g., sand, 

Bristol Bay), largely the result of the presence of a few species (e.g., rainbow 

smelt, Osmerus mordax) that were absent in other regions. 

 7. Most fishes captured were juveniles, highlighting the importance of the nearshore 

to critical early life-history stages of managed and ecologically important species. 

 8. Forage species (e.g., Pacific herring and Pacific sand lance, Ammodytes 

hexapterus) may be the most susceptible fishes to shoreline disturbance because of 

their use of nearshore habitats for feeding and shelter as juveniles and for 

spawning as adults. 

 9. The high abundance and diversity of fishes in eelgrass and kelp warrant special 

protection of these habitats in the event of an oil spill or other shoreline 

disturbance, especially if these habitats are known spawning areas (e.g., Pacific 

herring). 

10. Our nearshore dataset is extremely important to managers responsible for oil spill 

response, natural resource damage assessment, essential fish habitat identification, 

and long-term monitoring. 

Resource managers in Alaska need information on fish use of shallow-water, marine 

habitats to protect areas critical to fisheries and to assess anthropogenic actions that may 

adversely affect fish habitat. The spatially explicit information in this atlas shows managers, 

scientists, and the public the statewide importance of nearshore marine habitats to fishery 

resources, and will provide a valuable baseline in the event of a major human or natural 

disturbance. 
v 

CONTENTS

Abstract .................................................................................................................................... iii 

List of Tables .......................................................................................................................... vii 

List of Figures ........................................................................................................................ viii 

Introduction ................................................................................................................................1 

Background ................................................................................................................................2 

Geographic Coverage and Sampling Protocol ...........................................................................3 

 Fish Capture and Processing ..........................................................................................7 

 Temperature and Salinity ...............................................................................................8 

 Data Analysis .................................................................................................................8 

 Caveats .........................................................................................................................10 

Summaries................................................................................................................................10 

 Statewide ......................................................................................................................10 

 Arctic (AR) ..................................................................................................................14 

 Aleutian Islands (AI)....................................................................................................15 

 Bristol Bay (BB) ..........................................................................................................16 

 Southcentral (SC) .........................................................................................................17 

 Prince William Sound (PWS) ......................................................................................18 

 Southeast – Northern Outside (SENO) ........................................................................19 

 Southeast – Northern Inside (SENI) ............................................................................20 

 Southeast – Southern Outside (SESO) .........................................................................22 

 Southeast – Southern Inside (SESI) .............................................................................23 

Discussion ................................................................................................................................24 

 Distribution ..................................................................................................................24 

 Abundance and Diversity .............................................................................................25 

 Species Composition and Habitat Use .........................................................................27 

 Fish Life Stages ............................................................................................................28 

 Forage Fishes ...............................................................................................................29 

 Essential Fish Habitat ..................................................................................................30 

 Environmental Assessment ..........................................................................................31 
vi 

Conclusions ..............................................................................................................................31 

The Atlas ..................................................................................................................................32 

 Species List ..................................................................................................................35 

 Species Pages ...............................................................................................................39 

 Species Photos ...........................................................................................................159 

Acknowledgments..................................................................................................................176 

Citations .................................................................................................................................177 

Tables .....................................................................................................................................185 

Figures....................................................................................................................................229 

Appendix ................................................................................................................................244 

 
vii 

LIST OF TABLES 

Table 1. Number of locations, sites, seine hauls, and fish captured in each region ............185 

Table 2. Total catch and frequency of occurrence (FO) of fish taxa by region...................186 

Table 3. Mean CPUE and FO of fish taxa by region ...........................................................192 

Table 4. Mean CPUE and FO of fish taxa by habitat type (all regions combined) .............198 

Table 5. Total catch and length metrics of fish taxa (all regions combined) ......................203 

Table 6. Water temperature and salinity by region .............................................................207 

Table 7. Total catch, mean CPUE, and FO of fish taxa (AR) .............................................208 

Table 8. Total catch, mean CPUE, and FO of fish taxa by habitat type (AI) ......................210 

Table 9. Total catch, mean CPUE, and FO of fish taxa (BB) .............................................212 

Table 10. Total catch, mean CPUE, and FO of fish taxa by habitat type (SC) .....................213 

Table 11. Total catch, mean CPUE, and FO of fish taxa by habitat type (PWS) ..................215 

Table 12. Total catch, mean CPUE, and FO of fish taxa by habitat type (SENO) ...............218 

Table 13. Total catch, mean CPUE, and FO of fish taxa by habitat type (SENI) .................221 

Table 14. Total catch, mean CPUE, and FO of fish taxa by habitat type (SESO) ................224 

Table 15. Total catch, mean CPUE, and FO of fish taxa by habitat type (SESI) ..................227 

 
viii 

LIST OF FIGURES 

Figure 1. Map with regions and sites surveyed in Alaska ...................................................229 

Figure 2. Seine frequency by month (1998-2011) ..............................................................230 

Figure 3. Photos of habitat types sampled ...........................................................................231 

Figure 4. Map with locations and sites surveyed (AR) .......................................................232 

Figure 5. Map with locations and sites surveyed (AI) ........................................................233 

Figure 6. Map with location and sites surveyed (BB) .........................................................234 

Figure 7. Map with locations and sites surveyed (SC) ........................................................235 

Figure 8. Map with locations and sites surveyed (PWS) ....................................................236 

Figure 9. Map with locations and sites surveyed (SENO) ..................................................237 

Figure 10. Map with locations and sites surveyed (SENI) ....................................................238 

Figure 11. Map with locations and sites surveyed (SESO) ...................................................239 

Figure 12. Map with locations and sites surveyed (SESI) ....................................................240 

Figure 13. Photos of beach seine and setting net ..................................................................241 

Figure 14. Similarities in fish assemblages among regions grouped by habitat type ...........242 

Figure 15. Similarities in fish assemblages among habitat types within regions ..................243 
INTRODUCTION 

 The National Marine Fisheries Service (NMFS) is mandated to identify and protect 

essential fish habitat (EFH) for all life stages of species included in a fishery management 

plan (FMP) (Minello 1999). Identifying EFH requires basic information on fish distribution, 

abundance, and habitat use. For many FMP species in Alaska this information is limited or 

absent, especially for early life stages. It is generally known, however, that juvenile life 

stages of many commercially and ecologically important fishes use nearshore marine habitats 

(Thayer et al. 1978, Beck et al. 2003). Nonetheless, traditional fishery surveys by resource 

agencies largely ignore the nearshore environment. To address this information gap, 

scientists from Auke Bay Laboratories (ABL) have led the effort on nearshore research for 

the Alaska Fisheries Science Center since 1998. 

 Nearshore marine habitats (< 6 m deep, < 20 m offshore) are some of the most 

productive in Alaska (Robards et al. 1999a, Dean et al. 2000). The rugged and extensive 

coastline of Alaska (about 70,000 km; ADNR 2012) provides a variety of nearshore habitats 

(e.g., sheltered and exposed rocky shores, eelgrass (Zostera marina) meadows, kelp forests, 

and sand beaches) for over 100 fish species (NMFS 2012a). Many species that use the 

nearshore (includes estuaries), contribute directly to Alaska’s economy. From 2000 to 2004, 

about 15% of the total landed weight (25 billion pounds) and 32% of the total dollar value 

($4.7 billion) from commercial landings in Alaska were attributed to estuarine (i.e., 

nearshore) fish and shellfish (Lellis-Dibble et al. 2008). In addition, ecologically important 

species such as forage fishes (e.g., Pacific herring, Clupea pallasii, and Pacific sand lance, 

Ammodytes hexapterus) use the nearshore for spawning and nursery habitat (Cooney 2007, 

Johnson et al. 2008). Forage fishes are abundant, schooling species preyed upon by many 

other fishes, seabirds, and marine mammals (Springer and Speckman 1997). 

 Nearshore habitats are susceptible to human and natural disturbance (Schmitten 1999, 

Harris et al. 2008). Thus, resource managers need information on fish use of shallow, 

nearshore habitats to protect areas critical to fisheries and to assess anthropogenic actions 

that may adversely affect EFH. Providing managers with a comprehensive synthesis of our 

nearshore surveys of coastal Alaska was our focus in preparing this publication. Specific 

objectives of this paper were to 1) interpret and synthesize data from 14 years (1998 to 2011) 
2 

of nearshore surveys across the state on fish distribution, catch, habitat use, and size; 2) provide 

detailed comparisons on fish distribution and species composition among different regions and 

habitat types; and 3) present distribution maps and photos of all fish species captured. The 

information presented in this synthesis will help managers, scientists, and the public better 

understand the statewide importance of nearshore habitats to fishery resources and will 

provide a valuable baseline for routine monitoring (e.g., every 5 or 10 years) and natural 

resource damage assessment. 

BACKGROUND

 Nearshore surveys were initiated by ABL scientists in 1998 when NOAA’s EFH funds 

first became available. Now with 14 years (1998 to 2011) of standardized sampling of coastal 

habitats across Alaska, we have one of the most complete and valuable datasets available to 

managers. The dataset described herein encompasses a suite of studies, but the sampling 

methods and core group of researchers remained the same throughout all years. Some 

surveys were based on opportunistic sampling, whereas others were funded studies (e.g., 

EFH and North Pacific Research Board) that focused on specific habitat types (i.e., eelgrass) 

(Murphy et al. 2000, Johnson et al. 2003a, Johnson and Thedinga 2005, Harris et al. 2008), 

nearshore prey of Steller sea lions (Eumetopias jubatus) (Thedinga et al. 2006a), forage fish 

ecology (Thedinga et al. 2006b, Johnson et al. 2008), or specific areas of the state such as the 

Aleutian Islands (Thedinga et al. 2008) or Prince William Sound (Johnson et al. 2010a). 

 In 2005, we published an atlas on nearshore fishes of southeastern Alaska (Johnson et 

al. 2005); the atlas contains information on distribution and catch data from beach seine 

surveys in a variety of nearshore habitat types sampled from 1998 to 2004. Beginning in 

2005, however, we expanded our nearshore surveys into other areas of Alaska including the 

Arctic, Aleutian Islands, Bristol Bay, Southcentral, and Prince William Sound. As we 

continued to survey throughout the state, it became clear that an easily updatable, online atlas 

was warranted. The first edition of our dynamic, online fish atlas was launched in October 

2006, and was recently updated in 2012 (NMFS 2012a). This interactive, online database is 

the primary source of information for managers at the NMFS Alaska Regional Office on 
3 

issues related to EFH and shoreline development. As designed, the online atlas is data rich 

and shows fish catch by region, site, species, and habitat, but offers no interpretation or 

synthesis of the data. Consequently, there was a need for a comprehensive written synthesis 

of our entire 14-year dataset. The intention of this synthesis is to provide a comprehensive 

data analysis across all regions of Alaska. 

GEOGRAPHIC COVERAGE AND SAMPLING PROTOCOL 

 The nearshore fish and habitat information described in this atlas is from nine regions 

of Alaska: Arctic (AR), Aleutian Islands (AI), Bristol Bay (BB), Southcentral (SC), Prince 

William Sound (PWS), Southeast – Northern Outside (SENO), Southeast – Northern Inside 

(SENI), Southeast – Southern Outside (SESO), and Southeast – Southern Inside (SESI) 

(Fig. 1). Most regions are identified by the general geographic location of sampling areas 

within the state. We did, however, divide the complex archipelago of southeastern Alaska 

into four separate regions (SENO, SENI, SESO, and SESI); boundaries were loosely drawn 

along gradients of oceanographic conditions and species distributions (e.g., shiner perch, 

Cymatogaster aggregata, and black rockfish, Sebastes melanops; Johnson et al. 2005). 

Marine waters of southeastern Alaska are characterized by a north-south temperature 

gradient (Quast 1968) and an inshore-offshore salinity gradient (Murphy and Orsi 1999). For 

our southeast regions, northern-southern divisions were made at Frederick Sound, and 

outside-inside divisions were based on general proximity to the outer coast or major ocean 

entrances. Similar geographic divisions of southeastern Alaska waters are reported in the 

literature (Quast 1968; Johnson et al. 2003a, b). Overall, our region boundaries match 

reasonably well with the classification of marine ecoregions of Alaska described in Piatt and 

Springer (2007). 

 Within our nine defined regions, we sampled fish with a beach seine at 93 general 

locations (e.g., specific bay or island). At each location, between 1 and 25 sites were sampled 

with a seine, and at least one seine haul was made at each site. At some locations, sites were 

sampled more than once in the same year (e.g., Benjamin Island in SENI) or across several 

years (e.g., Chukchi Sea in AR) (Appendix). A total of 1,009 seine hauls were made at 555 
4 

unique sites (Table 1, Appendix). All sites were sampled during daylight hours and within 2 

hours of low tide (range: +1.0 m to 1.5 m below mean lower low water, MLLW). A 

geographic position was obtained in the middle of each seine site with a hand-held global 

positioning system (GPS). Inclusive of all regions and years, most sampling effort (75%) was 

in summer (Fig. 2). 

 Individual sampling sites within each location were selected based on habitat type and 

had to be approachable by skiff and free of seining obstructions (e.g., large boulders). Habitat 

type was determined by the visual observation of the dominant substrate present and the 

presence or absence of attached vegetation. Up to four habitat types were sampled at each 

location: bedrock outcrops, eelgrass meadows, understory kelp beds, and sand or gravel 

beaches with no attached vegetation (Fig. 3). Bedrock outcrops (hereafter referred to as 

bedrock) are steep (> 20% gradient) and usually located in semi-exposed to semi-protected 

areas near the entrance of bays. Vegetation on bedrock walls is often dominated by patchy to 

lush kelps (Laminariales) (e.g., Agarum clathratum, Alaria marginata, and Saccharina 

latissima; Lindeberg and Lindstrom 2010). Eelgrass meadows are typically located inside 

protected coves or bays with flat to moderate gradient (5-10%) and predominantly finegrained sediments. Relative to MLLW, eelgrass occupies areas of the lower intertidal and 

subtidal zones from +1 m to 6 m. Understory kelp beds sampled were located in semiexposed to semi-protected areas with moderate gradient (10-20%) and predominantly cobble 

substrates. Relative to MLLW, understory kelps occupy lower intertidal and subtidal areas 

from 0 m to 30 m. Understory kelps typically form low-lying mats that, unlike canopy 

kelps, do not reach the water’s surface at high tide. Understory kelps (hereafter referred to as 

kelp) are dominated by patchy to lush Laminariales (e.g., A. marginata, Cymathaere 

triplicata, and S. latissima). Sand or gravel beaches (hereafter referred to as sand) are 

generally “pocket” beaches found inside semi-protected to protected bays with flat to 

moderate gradient (5-10%) and no attached vegetation. Algal mats (e.g., Cladophora 

seriacea and Pylaiella spp.) are sometimes present on sand beaches. Of the 555 unique sites 

sampled, 127 were bedrock, 144 were eelgrass, 149 were kelp, and 135 were sand. Of the 

three habitat types with attached vegetation (i.e., bedrock, eelgrass, and kelp), cover was 

usually greatest in eelgrass and least in bedrock. The habitats we sampled do not represent all 
5 

nearshore habitats available to fish, but are common in Alaska and can be effectively 

sampled with a beach seine. 

 The number of sites, time of sampling, and habitat types sampled varied among our 

regions. A brief description of our defined regions and sampling efforts within each follows. 

Arctic (AR): Survey area extended from Skull Cliffs in the Chukchi Sea east to Cooper 

Island in the Beaufort Sea; 81 hauls were made among two locations and 31 sites, mostly in 

August from 2004 to 2009 (Table 1; Fig. 4; Appendix). Sites in the Chukchi Sea (n = 19) 

were exposed, with no barrier islands or lagoons. Sites in the Beaufort Sea (n = 12) were 

spread along the Plover Islands, a group of barrier islands that separates the western Beaufort 

Sea from Elson Lagoon; fish were sampled on the Beaufort Sea side and Elson Lagoon side 

of some islands (e.g., Cooper Island) within this group. All sample sites in our AR region 

were sand habitat; sand or gravel beaches are the predominant nearshore habitat type in the 

Arctic. Our AR region is within the Beaufort-Chukchi Sea, Barrier Island-Lagoon System 

ecoregion (Piatt and Springer 2007). More detailed descriptions of our sampling in the 

Chukchi and Beaufort seas are provided in Johnson et al. (2010b) and Thedinga et al. (in 

press). 

Aleutian Islands (AI): Survey area was limited to Unalaska, Akutan, and Akun islands; 70 

hauls were made among six locations and 70 sites in June 2005 (Table 1; Fig. 5; Appendix). 

Habitat types sampled were bedrock, kelp, and sand; we did not observe or sample any 

eelgrass. Our AI region is within the Eastern Aleutians ecoregion (Piatt and Springer 2007). 

A more detailed description of our sampling in the Aleutian Islands is provided in Thedinga 

et al. (2008). 

Bristol Bay (BB): Survey area was limited to Nushagak Bay, a large estuary that opens into 

Bristol Bay; eight hauls were made at one location and eight sites in July 2009 (Table 1; 

Fig. 6; Appendix). All sample sites in our BB region were sand habitat, as sand beaches and 

mud flats are the predominant nearshore habitats in Nushagak Bay. Our BB region is within 

the Eastern Bering Sea – Alaska Coastal ecoregion (Piatt and Springer 2007). 

Southcentral (SC): Survey area extended from upper Cook Inlet near Anchorage to 

Resurrection Bay near Seward; 23 hauls were made among five locations and 22 sites in July 

2008, August 2009, and July 2010 (Table 1: Fig. 7; Appendix). All habitat types were 
6 

sampled in SC; only one eelgrass site was sampled, however, and this was in Resurrection 

Bay. Our SC region spans three ecoregions (Piatt and Springer 2007): our Knik Arm and Fire 

Island sites (locations 10 and 11; Fig. 7) are within the Western Cook Inlet – Shelikof Strait 

ecoregion; our Inner Kachemak Bay and Outer Kachemak Bay sites (locations 12 and 13; 

Fig. 7) are within the Southeastern Cook Inlet – Kodiak Upwelling ecoregion; and our 

Resurrection Bay sites (location 14; Fig. 7) are within the Northern Gulf of Alaska Shelf 

ecoregion. 

Prince William Sound (PWS): Survey area extended from waters near Latouche Island east 

to Valdez; 133 hauls were made among 13 locations and 50 sites, mostly in summer in 1999, 

2006, 2007, 2009, and 2010 (Table 1; Fig. 8; Appendix). All habitat types were sampled in 

PWS. Our PWS region is within the Prince William Sound Inside Waters ecoregion (Piatt 

and Springer 2007). A more detailed description of our sampling in Prince William Sound is 

provided in Johnson et al. (2010a). 

Southeast - Northern Outside (SENO): Survey area extended from Cross Sound south to 

southern Sitka Sound, mostly on or near the outer coasts of Chichagof and Baranof islands; 

164 hauls were made among 17 locations and 127 sites, mostly in summer in 1998, 1999, 

2001-2003, 2006, 2008, and 2010 (Table 1; Fig. 9; Appendix). All habitat types were 

sampled in SENO. Our SENO region is almost entirely within the Chichagof Shelf and 

Inside Waters ecoregion (Piatt and Springer 2007). More detailed descriptions of our 

sampling in northern outside waters of southeastern Alaska are provided in Johnson et al. 

(2003a, 2005). 

Southeast - Northern Inside (SENI): Survey area extended from Haines south to Frederick 

Sound; 382 hauls were made among 27 locations and 153 sites, mostly in summer from 1998 

to 2006 and 2008 to 2011 (Table 1; Fig. 10; Appendix). All habitat types were sampled in 

SENI. Our SENI region is within the Chichagof Shelf and Inside Waters ecoregion (Piatt and 

Springer 2007). More detailed descriptions of our sampling in northern inside waters of 

southeastern Alaska are provided in Harris et al. (2005, 2008), Johnson et al. (2003a, 2005), 

Johnson and Thedinga (2005), and Thedinga et al. (2006a). 

Southeast - Southern Outside (SESO): Survey area extended from southern Chatham Strait 

south to Dixon Entrance; 114 hauls were made among 12 locations and 60 sites, mostly in 
7 

summer from 1998 to 2000 and in 2007 (Table 1; Fig. 11; Appendix). All habitat types were 

sampled in SESO. Our SESO region is within the Prince of Wales Shelf and Inside Waters 

ecoregion (Piatt and Springer 2007). More detailed descriptions of our sampling in southern 

outside waters of southeastern Alaska are provided in Johnson et al. (2003a, 2005) and 

Murphy et al. (2000). 

Southeast - Southern Inside (SESI): Survey area extended from Frederick Sound south to 

near Ketchikan; 34 hauls were made among 10 locations and 34 sites in summer 2000 and 

2007 (Table 1; Fig. 12; Appendix). Habitat types sampled were bedrock, eelgrass, and kelp; 

no sand was sampled. Our SESI region is almost entirely within the Prince of Wales Shelf 

and Inside Waters ecoregion (Piatt and Springer 2007). More detailed descriptions of our 

sampling in southern inside waters of southeastern Alaska are provided in Johnson et al. 

(2003a, 2005). 

Fish Capture and Processing 

At each site, fish were captured with a beach seine within 20 m of shore in waters 

usually < 6 m deep. The seine is 37-m long with variable mesh that tapers from 5-m wide at 

the center to 1-m wide at the ends (Fig. 13). Outer panels are each 10-m of 32-mm stretch 

mesh, intermediate panels are each 4-m of 6-mm square mesh, and the bunt is 9-m of 3.2-mm 

square mesh. The seine has a leadline and a floatline so that the bottom contacts the substrate 

and the top floats on the surface. We set the seine as a “round haul” by holding one end on 

the beach, backing around in a skiff with the other end to the beach about 18 m from the 

start, and pulling the seine onto shore (Fig. 13). 

After retrieval of the seine, the entire catch was sorted, identified to species, and 

counted. A subsample (up to 50 fish) of each species was measured either to the nearest 

millimeter fork length (FL), or, in species without a distinct fork in caudal fin, to total length 

(TL). Fish were briefly anesthetized in a mixture of 1 part carbonated water to 2 parts 

seawater for identification and measurement. Smaller individuals (< 50 mm length) of some 

families of fish (e.g., Cottidae, Gadidae, and Hexagrammidae) that could not be easily 

identified in the field were grouped and recorded as juvenile sculpin, juvenile cod, or juvenile 

greenling. Misidentification problems with some Myoxocephalus species (i.e., frog sculpin, 
8 

M. stelleri; great sculpin, M. polyacanthocephalus; shorthorn sculpin, M. scorpius) in areas 

where their distribution overlaps (Mecklenburg et al. 2002), prompted us to combine these 

three species and refer to them collectively as Myoxocephalus spp. in most tables and figures. 

Similarly, we combine the northern rock sole (Lepidopsetta polyxystra) and southern rock 

sole (L. bilineata) and refer to them collectively as rock sole (Lepidopsetta spp.). 

The number of fish in large catches was estimated gravimetrically. To achieve this, a 

random subsample of approximately 500 fish was removed from the total catch, and the 

remaining fish were collectively weighed to the nearest 0.1 kg. Fish in the sub-sample were 

counted and weighed to the nearest gram. A mean weight of fish determined from the 

subsample was used to estimate the number of fish in the total catch. The proportion of each 

species in the subsample was also used to determine the species composition of the total 

catch. During processing, fish were held in buckets with aerators, and when data collection 

was complete, fish were released at the site of capture. 

Temperature and Salinity 

Surface water temperature and salinity (practical salinity scale, PSS) were measured at 

most seine sites each site visit. Temperature and salinity were measured at about 10-cm depth 

with a thermometer and hand-held refractometer. 

Data Analysis 

From our seine catches we determined relative abundance, frequency of occurrence, 

and species richness. Abundance is expressed in absolute numbers (i.e., number of individual 

fish captured) and catch-per-unit-effort (CPUE; i.e., number of fish captured per seine haul). 

Frequency of occurrence (FO) was determined for each species; FO is calculated as the 

number of seine hauls in which a species was captured divided by the total number of seine 

hauls and multiplied by 100. Calculated FO is expressed as percentages (0-100%) and as 

categories: abundant ( 50%), common (25-49%), occasional (10-24%), uncommon (5-9%), 

rare (< 5%), and absent. Species richness refers to the total number of fish species captured. 

Individuals identified only to family (e.g., juvenile sculpin, Cottidae) were counted in the 

total catch, but were only considered as a separate species for species richness calculations if 
9 

no other species from the same family were captured. In all tables, the values for total catch, 

CPUE, FO, species richness, and fish length are calculated cumulatively across all sampling 

efforts. 

Species composition was compared among regions for each habitat type and among 

habitat types within each region. An analysis of similarity (ANOSIM) was conducted using a 

Bray-Curtis similarity index; a fourth-root transformation of the catch data was used to 

reduce the influence of very abundant species. The resulting similarity matrices are presented 

using non-metric, multi-dimensional scaling (nMDS) ordination plots (Clarke and Warwick 

2001). These plots show a configuration of the data; the more similar samples are to each 

other, the closer they are to each other on the plot (Clarke and Warwick 2001). Stress values 

associated with each plot represent the dimensionality of the data; stress values < 0.1 

correspond to good ordination (Clarke and Warwick 2001). We restricted the ANOSIM 

analyses to summer data only (June-September), which encompassed the majority of our 

sampling effort. Species contributing most to either the similarity or dissimilarity among and 

within regions by habitat type were identified using a similarity percentage analysis 

(SIMPER; Clarke and Warwick 2001). Species that contributed at least 5% to the 

cumulative, within-group dissimilarity were selected as good discriminators among regions 

(for each habitat type) or habitat types (within regions). Species composition could not be 

compared among all regions by habitat type or among habitat types within all regions 

because not all habitat types were sampled in each region: bedrock was not sampled in the 

AR or BB; eelgrass was not sampled in the AR, BB, or AI, and only one eelgrass site was 

sampled in SC (omitted in ANOSIM analyses); kelp was not sampled in the AR or BB; and 

sand was not sampled in SESI. 

From our length-frequency data collected for each species, we determined the 

approximate percentage of juvenile versus adult life stages. We compared our speciesspecific, length-frequency distributions to species-specific, length-at-maturity estimates 

reported online (life-history tool; FishBase 2012). Length frequency data for pink salmon 

(Oncorhynchus gorbuscha) and sockeye salmon (O. nerka) do not include adults; we did 

capture a few adults of these species, but they were released without measuring and were 

considered insignificant in our catches. 
10 

Caveats 

Our sampling represents only a “snapshot”, temporally and spatially, of fish 

distribution and habitat use. The patchy distribution of some fish species and differences in 

water temperature, salinity, proximity to spawning areas, life stage, sampling effort, gear 

type, and time of sampling can determine the presence or absence of any given species at any 

given time. For example, the fact that we did not capture rainbow smelt (Osmerus mordax) in 

the Arctic with a beach seine does not mean that they are not present in the Arctic; we did 

catch rainbow smelt with a small trawl farther offshore, but those data are not included in this 

paper. In addition, the absence of a particular habitat type from any given region (e.g., sand 

in SESI) represents a lack of sampling of that habitat type and not necessarily the absence of 

that habitat type. 

SUMMARIES 

Statewide 

Nearshore marine waters in Alaska support an abundant and diverse assemblage of 

fishes. An estimated 718,345 fish representing at least 121 species from 29 families were 

captured in 1,009 beach seine hauls from 1998 to 2011 (Table 2). Four commercially 

important species accounted for 55% of the total catch: walleye pollock (Theragra

chalcogramma), Pacific herring, pink salmon, and chum salmon (Oncorhynchus keta) 

(Table 2). Many species (33%) were captured in only small or incidental numbers (< 10 fish); 

14 species were represented by a total catch of one fish (Table 2). 

Species distribution patterns varied greatly among regions. Only two species were 

captured in all nine regions of Alaska: pink salmon and threespine stickleback (Gasterosteus 

aculeatus) (Table 2). Juvenile sculpins (Cottidae) were also present in all nine regions, but 

were likely comprised of several different species. Forty-one species were captured in only 

one region of Alaska; for example, pond smelt (Hypomesus olidus) was captured only in 

Bristol Bay (BB) (Table 2). Most species were present in multiple, but not all regions of 

Alaska; for example, saffron cod (Eleginus gracilis) was frequent in Southcentral (SC) 
11 

(FO = 61%), common in Prince William Sound (PWS) (FO = 49%), occasional in Arctic 

(AR) (FO = 11%), but absent in Aleutian Islands (AI), BB, and all Southeast (SE) regions 

(Table 2). The only species categorized as frequent across all regions of Alaska was the 

crescent gunnel (Pholis laeta, FO = 55%); this species was captured frequently in most 

regions (Table 2), but usually in small numbers (mean CPUE = 12 fish; all Alaska regions 

combined) (Table 3). Conversely, the three most abundant species based on total catch 

(walleye pollock, Pacific sand lance, Pacific herring) were categorized as only occasional 

(FO = 10-24%; Table 2); these species were captured infrequently, but often in large 

numbers (mean CPUE = ~116 fish for all three species; all Alaska regions combined) 

(Table 3). 

Catch-per-unit-effort and species richness varied by region and habitat type (Tables 3 

and 4). Mean CPUE (all taxa) was lowest in SC (124 fish) and greatest in AI (1,202 fish), 

whereas species richness was lowest in BB (17 species) and greatest in Southeast - Northern 

Inside (SENI) (67 species) (Table 3). Dominant species by region based on CPUE were 

capelin in the AR (178 fish); Pacific sand lance in AI (506 fish); rainbow smelt in BB (640 

fish); Pacific herring in SC (34 fish), PWS (186 fish), Southeast - Northern Outside (SENO) 

(125 fish), and Southeast - Southern Inside (SESI) (148 fish); walleye pollock in SENI (323 

fish), and pink salmon in Southeast - Southern Outside (SESO) (112 fish) (Table 3). Mean 

CPUE by habitat type (across all regions) was lowest in sand (478 fish) and greatest in 

eelgrass (890 fish), whereas cumulative species richness was lowest in bedrock (57 species) 

and greatest in sand (87 species) (Table 4). Dominant species by habitat type based on mean 

CPUE were Pacific herring in bedrock (180 fish); chum salmon in eelgrass (190 fish); 

walleye pollock in kelp (313 fish); and Pacific sand lance in sand (175 fish) (Table 4). Across 

all regions and habitat types, mean CPUE was 712 fish and cumulative species richness was 

121 (Table 3). 

Species composition differed significantly (P < 0.001) among regions grouped by 

habitat type: (ANOSIM: bedrock, Global R = 0.268; eelgrass, R = 0.363; kelp, R = 0.347; 

and sand, R = 0.486). Most regional bedrock assemblages (sample size in parentheses; 

AI (10), SC (4), PWS (31), SENO (70), SENI (67), SESO (9), and SESI (4)) differed 

significantly (0.180 < R < 0.574, P < 0.001) except SC versus SENO (R = 0.066, P > 0.05), 
12 

SC versus SENI (R = 0.110, P > 0.05), SC versus SESI (R = 0.396, P > 0.05), SENO versus 

SESI (R = 0.132, P > 0.05), and SESI versus SESO (R = 0.077, P > 0.05) (Fig. 14). Species 

composition in bedrock was unique in AI and PWS; species in these regions were very 

different from each other and from all other regions (Fig. 14). Based on pairwise SIMPER 

analysis, taxa that were good discriminators among regional bedrock assemblages were chum 

salmon, coho salmon (O. kisutch), crescent gunnel, kelp perch (Brachyistius frenatus), 

Pacific cod (Gadus macrocephalus), Pacific herring, Pacific sandfish (Trichodon trichodon), 

Pacific sand lance, pink salmon, rock greenling (Hexagrammos lagocephalus), shiner perch, 

silverspotted sculpin (Blepsias cirrhosus), unidentified cod, unidentified fish larvae, 

unidentified greenling, and walleye pollock. All regional eelgrass assemblages (sample size 

in parentheses; PWS (50), SENO (57), SENI (120), SESO (54), and SESI (13)) differed 

significantly (0.187 < R < 0.743, P < 0.001), except SENI versus SESI (R = 0.068, P > 0.05) 

(Fig. 14). Species composition in PWS eelgrass was most unlike any other region, largely 

due to the presence of saffron cod in PWS and their absence in all SE regions (Fig. 14). In 

addition, species composition in eelgrass differed between outside (SENO, SESO) and inside 

(SENI and SESI) regions of SE (Fig. 14). Species that were good discriminators among 

regional eelgrass assemblages were bay pipefish (Syngnathus leptorhynchus), chum salmon, 

saffron cod, shiner perch, threespine stickleback, snake prickleback (Lumpenus sagitta), and 

tubesnout (Aulorhynchus flavidus). All regional kelp assemblages (sample size in 

parentheses; AI (27), SC (8), PWS (45), SENO (26), SENI (117), SESO (34), and SESI (17)) 

differed significantly (0.116 < R < 0.916, P < 0.001), except SC versus SENI (R = 0.11, 

P > 0.05) (Fig. 14). Species composition in AI kelp was most unlike any other region 

(Fig. 14), partially due to the large number of unidentified cod and unidentified sculpin in our 

catches. Taxa that were good discriminators among regional kelp assemblages were chum 

salmon, crescent gunnel, Pacific cod, Pacific herring, Pacific sandfish, pink salmon, saffron 

cod, shiner perch, unidentified cod, unidentified sculpin, and walleye pollock. Most regional 

sand assemblages (sample size in parentheses; AR (81), AI (33), BB (8), SC (10), PWS (7), 

SENO (11), SENI (78), and SESO (17)) differed significantly (0.232 < R < 0.999, P < 0.001) 

except SENI versus SESO (R = 0.055, P > 0.05) and SENI versus PWS (R = 0.093, P > 0.05) 

(Fig. 14). Species composition in sand was most unique in AI and BB, partially due to the 
13 

high abundance of Pacific sand lance in AI and rainbow smelt and pond smelt in BB 

(Fig. 14). Taxa that were good discriminators among regional sand assemblages were capelin 

(Mallotus villosus), crescent gunnel, Myoxocephalus spp., northern sculpin (Icelinus 

borealis), Pacific herring, Pacific sand lance, Pacific staghorn sculpin (Leptocottus armatus), 

pink salmon, pond smelt, rainbow smelt, rock sole, saffron cod, shiner perch, threespine 

stickleback, unidentified cod, unidentified greenling, and unidentified sculpin. 

Based on estimated size at maturity, juvenile life stages dominated our catches for most 

species, particularly those included in a FMP for Alaska (Table 5). For example, all chum 

salmon, coho salmon, Pacific cod, pink salmon, and walleye pollock were juveniles; mean 

size (FL or TL) ranged from 52 mm to 96 mm (Table 5). For forage fishes, all Pacific herring 

and Pacific sandfish and most capelin (97%) and Pacific sand lance (83%) were also 

juveniles; mean FL ranged from 56 mm to 91 mm (Table 5). We captured a higher 

percentage of adults than juveniles for only a few species, including bay goby (Lepidogobius 

lepidus; 67%), least cisco (Coregonus sardinella; 100%), and northern sculpin (80%) 

(Table 5). 

Water temperature and salinity varied by region and were influenced by location, time 

of sampling, seasonal rainfall, and proximity to freshwater streams. Across all regions, 

temperature at time of sampling ranged from 0o C to 20o C, and salinity ranged from 0 PSS to 

35 PSS (Table 6). The coldest water temperature that we encountered was 0o C in AR, and 

the warmest water temperature was 20o C in PWS (Table 6). Similarly, the lowest salinity 

that we encountered was 0 PSS in SESI and the highest salinity was 35 PSS in AR and SENI 

(Table 6). More detailed temperature and salinity information from our survey locations and 

individual sites are available online (NMFS 2012a). 

Of the 121 species captured, 14 are a target or potential target (e.g., Arctic cod, 

Boreogadus saida and saffron cod) species in either a groundfish or salmon FMP in Alaska 

(Table 2) (NPFMC 2012). We also captured multiple forage fish species from several 

families (e.g., Ammodytidae, Osmeridae, and Trichodontidae) that are included in a 

groundfish FMP for Alaska; these species are included in a FMP because of their ecological 

importance. Target species captured include walleye pollock, pink salmon, chum salmon, 

Pacific cod, and coho salmon (Table 2), and important forage species captured include 
14 

capelin, Pacific sandfish, and Pacific sand lance. Pacific herring are an important commercial 

and forage species that we captured routinely, but are managed by the state and are not a 

target FMP species. 

In 14 years of surveys, we identified seven range extensions (not all have been 

published) and a dramatic increase in abundance of one species. Range extensions include 

bay goby, blackeye goby (Rhinogobiops nicholsii), brightbelly sculpin (Microcottus sellaris), 

kelp perch, roughback sculpin (Chitonotus pugetensis), saddleback sculpin (Oligocottus 

rimensis), and shiner perch; most of these extensions were north of their reported range 

(Csepp and Wing 1999, Mecklenburg et al. 2002). Some of the observed range extensions 

were relatively short distances (saddleback sculpin, ~ 60 km north) and likely the result of 

sampling in areas not sampled before. The dramatic increase in abundance was for saffron 

cod in nearshore habitats of western PWS (Johnson et al. 2009). 

Arctic (AR) 

A total of 20,803 fish representing 23 species were captured in 81 seine hauls from 

2004 to 2009 (Table 7). Mean CPUE was 257 fish for our AR region. Three taxa accounted 

for 88% of the total catch: capelin, juvenile sculpin, and Pacific sand lance. Capelin were 

abundant and often captured in large numbers (mean CPUE = 178 fish; FO = 51%), whereas 

juvenile sculpin were also abundant but usually captured in small numbers (mean CPUE = 26 

fish; FO = 64%) (Table 7). Thirteen species were captured in only small or incidental 

numbers (< 10 fish); six species were represented by a total catch of one fish (Table 7). More 

detailed descriptions of our catches in the Arctic are provided in Johnson et al. (2010b) and 

Thedinga et al. (in press). 

Catch and species richness varied between the Chukchi and Beaufort seas. Mean 

CPUE and species richness was 401 fish and 21 species in the Chukchi Sea (n = 47 hauls) 

and 58 fish and 15 species in the Beaufort Sea (n = 34 hauls). Within the Beaufort Sea 

location, there were also differences in catch and species richness between the Beaufort Sea 

side and the Elson Lagoon side of the Plover Islands group (e.g., Cooper Island); mean 

CPUE and species richness was 82 fish and 14 species on the Beaufort Sea side and 14 fish 

and 9 species on the Elson Lagoon side. Most (99%) capelin were captured in open waters of 
15 

the Chukchi (FO = 62%) and Beaufort seas (FO = 50%), and most (90%) whitefishes 

(Salmonidae, Coregoninae) were captured in Elson Lagoon (FO = 75%). Capelin were 

uncommon (FO = 8%) in Elson Lagoon, and whitefishes were rare (FO < 3%) in open waters 

of the Chukchi and Beaufort seas. Eleven species were unique to our AR region: Arctic cod, 

Arctic cisco (Coregonus autumnalis), Arctic sculpin (Myoxocephalus scorpioides), Arctic 

staghorn sculpin (Gymnocanthus tricuspis), Atlantic poacher (Leptagonus decagonus), 

fourhorn sculpin (M. quadricornis), fourline snakeblenny (Eumesogrammus praecisus), least 

cisco, longhead dab (Limanda proboscidea), variegated snailfish (Liparis gibbus), and 

veteran poacher (Podothecus veternus) (Table 2). 

Because sand was the only habitat type sampled in AR, species composition could not 

be compared to other habitat types. From regional comparisons of species composition 

within sand, however, the sand assemblage of fish in AR had an overall within-group, BrayCurtis similarity of 23.3% and consisted mainly of unidentified sculpin and capelin, which 

together accounted for 53% of the cumulative, within-group similarity. 

Aleutian Islands (AI) 

An estimated 84,118 fish representing 27 species were captured in 70 seine hauls in 

2005 (Table 8). Mean CPUE was 1,202 fish for our AI region. Four taxa accounted for 95% 

of the total catch: Pacific sand lance, juvenile cod, pink salmon, and Pacific sandfish 

(Table 8). Juvenile cod (mostly larvae) were likely a combination of Pacific cod and walleye 

pollock. Ten species were captured in only small or incidental numbers (< 10 fish); three 

species were represented by a total catch of one fish (Table 8). A more detailed description of 

our catches in the Aleutian Islands is provided in Thedinga et al. (2008). 

Catch-per-unit effort and species richness varied among habitat types. Mean CPUE was 

98 fish in bedrock, 1,648 fish in kelp, and 1,171 fish in sand. The most abundant taxa by 

habitat type were pink salmon and juvenile cod (CPUE = 34 fish each) in bedrock, juvenile 

cod (CPUE = 755 fish) in kelp, and Pacific sand lance (CPUE = 1,056 fish) in sand; FO 

ranged from 30 to 94% for these taxa (Table 8). Cumulative species richness was 14 in 

bedrock, 22 in kelp, and 17 in sand. Five species were captured in all habitat types, and 10 
16 

species were captured in only one habitat type (Table 8). No species were unique to our AI 

region (Table 2). 

Species composition differed significantly among habitat types in our AI region 

(ANOSIM: Global R = 0.503, P < 0.001). All habitat types (sample size in parentheses; 

bedrock (10), kelp (27), and sand (33)) differed significantly (0.280 < R < 0.651, P < 0.001);

each habitat type was unique based on species composition (Fig. 15). The AI bedrock 

assemblage had an overall within-group, Bray-Curtis similarity of 25.4% and consisted 

mainly of unidentified cod and unidentified fish larvae, which together accounted for 64% of 

the cumulative, within-group similarity. The AI kelp assemblage had an overall within-group 

similarity of 37.0% and consisted mainly of unidentified cod, unidentified sculpin, and pink 

salmon, which together accounted for 61% of the cumulative, within-group similarity. The 

AI sand assemblage had an overall within-group similarity of 33.6% and consisted mainly of 

Pacific sand lance that accounted for 57% of the cumulative, within-group similarity. Taxa 

that were good discriminators among AI habitat types were Pacific sandfish, Pacific sand 

lance, pink salmon, rock greenling, rock sole, silverspotted sculpin, unidentified cod, 

unidentified fish larvae, and unidentified sculpin. 

Bristol Bay (BB) 

A total of 6,775 fish representing 17 species were captured in 8 seine hauls in 2009 

(Table 9). Mean CPUE was 847 fish for our BB region. Two species accounted for 95% of 

the total catch: rainbow smelt and pond smelt. Both of these species were abundant 

(FO = 100%) and often captured in large numbers (rainbow smelt, CPUE = 640 fish; pond 

smelt, CPUE =166 fish) (Table 9). Ten species were captured in only small or incidental 

numbers (< 10 fish); two species were represented by a total catch of one fish (Table 9). Six 

species were unique to our BB region: Arctic lamprey (Lampetra camtschatica), belligerent 

sculpin (Megalocottus platycephalus), Bering poacher (Occella dodecahedron), blackline 

prickleback (Acantholumpenus mackayi), pond smelt, and rainbow smelt (Table 2). 

Because sand was the only habitat type sampled in BB, species composition could not 

be compared to other habitat types. From regional comparisons of species composition 

within sand, however, the sand assemblage of fish in BB had an overall within-group, Bray-
17 

Curtis similarity of 56.3% and consisted mainly of rainbow smelt and pond smelt, which 

together accounted for 67% of the cumulative, within-group similarity. 

Southcentral (SC) 

A total of 2,858 fish representing 42 species were captured in 23 seine hauls from 2008 

to 2010 (Table 10). Mean CPUE was 124 fish for our SC region. Six taxa accounted for 81% 

of the total catch: Pacific herring, pink salmon, walleye pollock, juvenile sculpin, saffron 

cod, and Pacific sand lance (Table 10). Twenty-four species were captured in only small or 

incidental numbers (< 10 fish); 12 species were represented by a total catch of one fish 

(Table 10). 

Catch-per-unit effort and species richness varied among habitat types. Mean CPUE was 

69 fish in bedrock, 78 fish in eelgrass, 177 fish in kelp, and 108 fish in sand (Table 10). The 

most abundant species captured by habitat type were Pacific sand lance in bedrock (CPUE = 

31 fish), masked greenling (H. octogrammus) in eelgrass (CPUE = 18 fish), pink salmon in 

kelp (CPUE = 48 fish), and Pacific herring in sand (CPUE = 78 fish); FO ranged from 20 to 

100% for these species (Table 10). Cumulative species richness was 12 in bedrock, 17 in 

eelgrass, 21 in kelp, and 26 in sand (Table 10). Pacific cod and whitespotted greenling 

(H. stelleri) were the only species captured in all habitat types, and 21 species were captured 

in only one habitat type (Table 10). Three species were unique to our SC region: longfin 

smelt (Spirinchus thaleichthys), prowfish (Zaprora silenus), and spotted snailfish 

(Liparis callydon) (Table 2). 

Species composition differed significantly among habitat types in our SC region 

(ANOSIM: Global R = 0.331, P < 0.001). All habitat types (bedrock, kelp, and sand) differed 

significantly (0.309 < R < 0.571, P < 0.001); species composition differed most between 

bedrock and kelp (Fig. 15). The SC bedrock assemblage had an overall within-group, BrayCurtis similarity of 11.9% and consisted mainly of unidentified greenling that accounted for 

70% of the cumulative, within-group similarity. The SC kelp assemblage had an overall 

within-group similarity of 42.2% and consisted mainly of saffron cod, crescent gunnel, and 

Pacific cod, which together accounted for 60% of the cumulative, within-group similarity. 

The SC sand assemblage had an overall within-group similarity of 17.1% and consisted 
18 

mainly of saffron cod, pink salmon, and sockeye salmon, which together accounted for 55% 

of the cumulative, within-group similarity. Taxa that were good discriminators among SC 

habitat types were coho salmon, crescent gunnel, masked greenling, Pacific cod, Pacific sand 

lance, pink salmon, saffron cod, silverspotted sculpin, sockeye salmon, unidentified 

greenling, and whitespotted greenling. 

Prince William Sound (PWS) 

A total of 61,423 fish representing 54 species were captured in 133 seine hauls between 

1999 and 2010 (Table 11). Mean CPUE was 462 fish for our PWS region. Four species 

accounted for 90% of the total catch: Pacific herring, saffron cod, pink salmon, and capelin 

(Table 11). Twenty-five species were captured in only small or incidental numbers (< 10 

fish); seven species were represented by a total catch of one fish (Table 11). A more detailed 

description of our catches in Prince William Sound is provided in Johnson et al. (2010a). 

Catch-per-unit-effort and species richness varied among habitat types. Mean CPUE was 

110 fish in bedrock, 739 fish in eelgrass, 283 fish in kelp, and 1,190 fish in sand (Table 11). 

The most abundant species captured by habitat type were pink salmon in bedrock (CPUE = 

72 fish), saffron cod in eelgrass (CPUE = 329 fish), capelin in kelp (CPUE = 148 fish), and 

Pacific herring in sand (CPUE = 1,161 fish); FO ranged from 13 to 74% for these species 

(Table 11). Cumulative species richness was 25 in bedrock, 41 in eelgrass, 43 in kelp, and 21 

in sand (Table 11). Ten species were captured in all habitat types, and 18 species were 

captured in only one habitat type (Table 11). Three species were unique to our PWS region: 

brightbelly sculpin (Microcottus sellaris), searcher (Bathymaster signatus), and smallmouth 

ronquil (B. leurolepis) (Table 2). 

Species composition differed significantly among habitat types in our PWS region 

(ANOSIM: Global R = 0.408, P < 0.001). Most habitat types (sample size in parentheses; 

bedrock (31), eelgrass (50), kelp (45), and sand (7)) differed significantly (0.164 < R < 0.823, 

P < 0.001), except bedrock versus sand (R = 0.11, P > 0.05); species composition in eelgrass 

was most unlike all other habitat types (Fig. 15). The PWS bedrock assemblage had an 

overall within-group, Bray-Curtis similarity of 18.2% and consisted mainly of crescent 

gunnel and Pacific herring, which together accounted for 56% of the cumulative, within-
19 

group similarity. The PWS eelgrass assemblage had an overall within-group similarity of 

44.8% and consisted mainly of saffron cod, crescent gunnel, and tubesnout, which together 

accounted for 55% of the cumulative, within-group similarity. The PWS kelp assemblage had 

an overall within-group similarity of 27.4% and consisted mainly of crescent gunnel, Pacific 

herring, padded sculpin (Artedius fenestralis), and Arctic shanny (Stichaeus punctatus), 

which together accounted for 55% of the cumulative, within-group similarity. The PWS sand 

assemblage had an overall within-group similarity of 34.2% and consisted mainly of Pacific 

herring and crescent gunnel, which together accounted for 54% of the cumulative, withingroup similarity. Species that were good discriminators among PWS habitat types were bay 

pipefish, crescent gunnel, Pacific herring, padded sculpin, pink salmon, saffron cod, and 

tubesnout. 

Southeast - Northern Outside (SENO) 

A total of 89,687 fish representing 62 species were captured in 164 seine hauls between 

1998 and 2010 (Table 12). Mean CPUE was 547 fish for our SENO region. Four species 

accounted for 79% of the total catch: Pacific herring, shiner perch, pink salmon, and chum 

salmon (Table 12). Nineteen species were captured in only small or incidental numbers (< 10 

fish); eight species were represented by a total catch of one fish (Table 12). More detailed 

descriptions of our catches in northern outside waters of southeastern Alaska are provided in 

Johnson et al. (2003a, 2005). 

Catch-per-unit-effort and species richness varied among habitat types. Mean CPUE was 

536 fish in bedrock, 614 fish in eelgrass, 551 fish in kelp, and 256 fish in sand (Table 12). 

The most abundant species captured by habitat type were pink salmon in bedrock (CPUE = 

233 fish), shiner perch in eelgrass (CPUE = 308 fish), and Pacific herring in kelp and sand 

(CPUE = 416 fish and 76 fish); FO ranged from 27 to 75% for these species (Table 12). 

Cumulative species richness was 40 in bedrock, 51 in eelgrass, 39 in kelp, and 27 in sand 

(Table 12). Fifteen species were captured in all habitat types, and 20 species were captured in 

only one habitat type (Table 12). Six species were unique to our SENO region: China 

rockfish (Sebastes nebulosus), Dover sole (Microstomus pacificus), kelp clingfish (Rimicola 
20 

muscarum), pygmy poacher (Odontopyxis trispinosa), saddleback sculpin, and wolf-eel 

(Anarrhichthys ocellatus) (Table 2). 

 Species composition differed significantly among habitat types in our SENO region 

(ANOSIM: Global R = 0.326, P < 0.001). The eelgrass assemblage differed significantly 

from all other habitat types (sample size in parentheses; bedrock (70), eelgrass (57), kelp 

(26), sand (11)) (0.437 < R < 0.700, P < 0.001); all other habitat comparisons were not 

significant (R < 0.122, P > 0.05) (Fig. 15). The SENO bedrock assemblage had an overall 

within-group, Bray-Curtis similarity of 12.2% and consisted mainly of shiner perch and 

unidentified greenling, which together accounted for 51% of the cumulative, within-group 

similarity. The SENO eelgrass assemblage had an overall within-group similarity of 41.7% 

and consisted mainly of shiner perch, crescent gunnel, bay pipefish, and unidentified 

greenling, which together accounted for 58% of the cumulative, within-group similarity. The 

SENO kelp assemblage had an overall within-group similarity of 21.3% and consisted 

mainly of kelp greenling (H. decagrammus), unidentified greenling, crescent gunnel, copper 

rockfish (S. caurinus), and shiner perch, which together accounted for 58% of the 

cumulative, within-group similarity. The SENO sand assemblage had an overall within-group 

similarity of 23.7% and consisted mainly of unidentified greenling and crescent gunnel, 

which together accounted for 53% of the cumulative, within-group similarity. Taxa that were 

good discriminators among SENO habitat types were bay pipefish, crescent gunnel, kelp 

greenling, Pacific cod, Pacific herring, Pacific sand lance, shiner perch, and unidentified 

greenling. 

Southeast - Northern Inside (SENI) 

A total of 399,648 fish representing 67 species were captured in 382 seine hauls 

between 1998 and 2011 (Table 13). Mean CPUE was 1,046 fish for our SENI region. Five 

species accounted for 87% of the total catch: walleye pollock, Pacific sand lance, chum 

salmon, Pacific herring, and pink salmon (Table 13). Twenty-one species were captured in 

only small or incidental numbers (< 10 fish); six species were represented by a total catch of 

one fish (Table 13). More detailed descriptions of our catches in northern inside waters of 

southeastern Alaska are provided in Johnson et al. (2003a, 2005). 
21 

Catch-per-unit-effort and species richness varied among habitat types. Mean CPUE was 

1,363 fish in bedrock, 1,313 fish in eelgrass, 980 fish in kelp, and 463 fish in sand (Table 13). 

The most abundant species captured by habitat type were walleye pollock in bedrock 

(CPUE = 468 fish), Pacific sand lance in eelgrass (CPUE = 439 fish), walleye pollock in kelp 

(CPUE = 729 fish), and pink salmon in sand (CPUE = 227 fish); FO ranged from 16 to 28% 

for these species (Table 13). Cumulative species richness was 31 in bedrock, 55 in eelgrass, 

52 in kelp, and 50 in sand (Table 13). Twenty-six species were captured in all habitat types, 

and 15 species were captured in only one habitat type (Table 13). Six species were unique to 

our SENI region: antlered sculpin (Enophrys diceraus), butter sole (Isopsetta isolepis), 

eulachon (Thaleichthys pacificus), ribbon snailfish (L. cyclopus), showy snailfish 

(L. pulchellus), and soft sculpin (Psychrolutes sigalutes) (Table 2). 

Species composition differed significantly among habitat types in our SENI region 

(ANOSIM: Global R = 0.363, P < 0.001). All habitat types (sample size in parentheses; 

bedrock (67), eelgrass (120), kelp (117), and sand (78)) differed significantly (0.131 < R <

0.663, P < 0.001); species composition in eelgrass was most unlike all other habitat types 

(Fig. 15). The SENI bedrock assemblage had an overall within-group, Bray-Curtis similarity 

of 15.2% and consisted mainly of crescent gunnel, silverspotted sculpin, and northern 

sculpin, which together accounted for 51% of the cumulative, within-group similarity. The 

SENI eelgrass assemblage had an overall within-group similarity of 35.6% and consisted 

mainly of crescent gunnel, Pacific staghorn sculpin, Myoxocephalus spp., and threespine 

stickleback, which together accounted for 51% of the cumulative, within-group similarity. 

The SENI kelp assemblage had an overall within-group similarity of 21.7% and consisted 

mainly of crescent gunnel, walleye pollock, silverspotted sculpin, Pacific cod, and Pacific 

herring, which together accounted for 54% of the cumulative, within-group similarity. The 

SENI sand assemblage had an overall within-group similarity of 20.9% and consisted mainly 

of crescent gunnel, rock sole, Myoxocephalus spp., and unidentified sculpin, which together 

accounted for 52% of the cumulative, within-group similarity. Species that were good 

discriminators among SENI habitat types were chum salmon, crescent gunnel, Pacific cod, 

Pacific herring, Pacific sand lance, Pacific staghorn sculpin, threespine stickleback, and 

walleye pollock. 
22 

Southeast - Southern Outside (SESO) 

A total of 39,772 fish representing 62 species were captured in 114 seine hauls between 

1998 and 2011 (Table 14). Mean CPUE was 349 fish for our SESO region. Five species 

accounted for 77% of the total catch: pink salmon, Pacific sand lance, shiner perch, bay 

pipefish, and threespine stickleback (Table 14). Twenty-four species were captured in only 

small or incidental numbers (< 10 fish); seven species were represented by a total catch of 

one fish (Table 14). More detailed descriptions of our catches in southern outside waters of 

southeastern Alaska are provided in Johnson et al. (2003a, 2005) and Murphy et al. (2000). 

Catch-per-unit-effort and species richness varied among habitat types. Mean CPUE was 

649 fish in bedrock, 492 fish in eelgrass, 144 fish in kelp, and 147 fish in sand (Table 14). 

The most abundant species captured by habitat type were Pacific sand lance in bedrock 

(CPUE = 390 fish), pink salmon in eelgrass and kelp (CPUE = 177 fish and 45 fish), and 

Pacific herring in sand (CPUE = 34 fish); FO ranged from 24 to 38% for these species 

(Table 14). Cumulative species richness was 18 in bedrock, 53 in eelgrass, 51 in kelp, and 27 

in sand (Table 14). Twelve species were captured in all habitat types, and 18 species were 

captured in only one habitat type (Table 14). Five species were unique to our SESO region: 

bocaccio (S. paucispinis), brown rockfish (S. auriculatus), roughback sculpin, sablefish 

(Anoplopoma fimbria), and yellowtail rockfish (S. flavidus) (Table 2). 

Species composition differed significantly among habitat types in our SESO region 

(ANOSIM: Global R = 0.493, P < 0.001). All habitat types (sample size in parentheses; 

bedrock (9), eelgrass (54), kelp (34), and sand (17)) differed significantly (0.181 < R < 0.913, 

P < 0.001); species composition differed most between bedrock and eelgrass (Fig. 15). The 

SESO bedrock assemblage had an overall within-group, Bray-Curtis similarity of 26.0% and 

consisted mainly of pink salmon that accounted for 70% of the cumulative, within-group 

similarity. The SESO eelgrass assemblage had an overall within-group similarity of 52.3% 

and consisted mainly of bay pipefish, crescent gunnel, and shiner perch, which together 

accounted for 53% of the cumulative, within-group similarity. The SESO kelp assemblage 

had an overall within-group similarity of 36.9% and consisted mainly of shiner perch, 

crescent gunnel, and copper rockfish, which together accounted for 60% of the cumulative, 

within-group similarity. The SESO sand assemblage had an overall within-group similarity 
23 

of 45.5% and consisted mainly of crescent gunnel, shiner perch, and Pacific staghorn sculpin, 

which together accounted for 59% of the cumulative, within-group similarity. Taxa that were 

good discriminators among SESO habitat types were bay pipefish, chum salmon, copper 

rockfish, crescent gunnel, Myoxocephalus spp., northern sculpin, pink salmon, shiner perch, 

snake prickleback, Pacific staghorn sculpin, and threespine stickleback. 

Southeast - Southern Inside (SESI) 

A total of 13,261 fish representing 47 species were captured in 34 seine hauls between 

1998 and 2011 (Table 15). Mean CPUE was 390 fish for our SESI region. Three species 

accounted for 72% of the total catch: Pacific herring, shiner perch, and bay pipefish 

(Table 15). Twenty species were captured in only small or incidental numbers (< 10 fish); 

four species were represented by a total catch of one fish (Table 15). More detailed 

descriptions of our catches in southern inside waters of southeastern Alaska are provided in 

Johnson et al. (2003a, 2005). 

Catch-per-unit-effort and species richness varied among habitat types. Mean CPUE was 

19 fish in bedrock, 490 fish in eelgrass, and 401 fish in kelp (Table 15). The most abundant 

species captured by habitat type were shiner perch in bedrock and eelgrass (CPUE = 11 fish 

and 206 fish), and Pacific herring in kelp (CPUE = 296 fish); FO ranged from 12 to 85% for 

these species (Table 15). Cumulative species richness was 9 in bedrock, 39 in eelgrass, and 

36 in kelp (Table 15). Seven species were captured in all habitat types, and 17 species were 

captured in only one habitat type (Table 15). Cabezon (Scorpaenichthys marmoratus) was 

the only species unique to our SESI region. 

Species composition differed significantly among habitat types in our SESI region 

(ANOSIM: Global R = 0.347, P < 0.001). All habitat types (sample size in parentheses; 

bedrock (4), eelgrass (13), and kelp (17)) differed significantly (0.169 < R < 0.864, 

P < 0.001); species composition differed most between bedrock and eelgrass (Fig. 15). The 

SESI bedrock assemblage had an overall within-group, Bray-Curtis similarity of 32.1% and 

consisted mainly of pink salmon that accounted for 80% of the cumulative, within-group 

similarity. The SESI eelgrass assemblage had an overall within-group similarity of 46.2% 

and consisted mainly of crescent gunnel, shiner perch, bay pipefish, and Pacific staghorn 
24 

sculpin, which together accounted for 57% of the cumulative, within-group similarity. The 

SESI kelp assemblage had an overall within-group similarity of 28.3% and consisted mainly 

of crescent gunnel and shiner perch, which together accounted for 54% of the cumulative, 

within-group similarity. Species that were good discriminators among SESI habitat types 

were bay pipefish, chum salmon, crescent gunnel, northern sculpin, pink salmon, shiner 

perch, snake prickleback, Pacific staghorn sculpin, threespine stickleback, and tubesnout. 

DISCUSSION 

Nearshore marine waters of Alaska provide a mosaic of habitat types that support a 

diverse assemblage of fishes. It is clear that shallow, nearshore habitats are used extensively 

by fishes, given that we captured 121 species and our mean CPUE was 712 fish. All 

nearshore habitats are used, but some provide more complexity and quality habitat than 

others (vegetated vs. nonvegetated). In addition, the importance of a species cannot be 

limited to commercial value only; many species that we captured provide ecological value as 

forage for other species. For example, we did not capture crescent gunnel in large numbers 

and they are not commercially important, but they are prey for river otter (Lutra canadensis), 

pigeon guillemot (Cepphus columba), and several fishes (Golet et al. 2000, Jewett et al. 

2002) including the commercially important lingcod (Beaudreau and Essington 2007). 

Because our knowledge of the nearshore environment has increased dramatically over the 

last 14 years, we can now provide spatially explicit information on the distribution, relative 

abundance, species composition, habitat use, and size of fishes in shallow, marine waters of 

Alaska. 

Distribution 

A wide range of species distribution patterns were evident from our surveys. Some of 

the more ubiquitous species were Pacific sand lance, pink salmon, and threespine 

stickleback; these species were captured from the Arctic to southeastern Alaska and appear to 

be adapted to a broad range of marine environments. Conversely, about a third of the species 

that we captured were restricted to one region (e.g., pond smelt in Bristol Bay) and could be 
25 

good indicator species to track changes in fish assemblages that may occur with climate 

change (Johnson and Thedinga 2005). Our catch data did not always agree with the reported 

distribution of some species. For example, Pacific herring is reported in nearshore waters of 

the Arctic (Mecklenburg et al. 2002), but we never captured Pacific herring in the Arctic in 

81 seine hauls. Low abundance of Pacific herring in the Arctic is likely the reason we 

captured none. 

Our survey data provides a finer-scale of distribution information than can be found 

most anywhere else. For example, copper rockfish is reported throughout coastal waters of 

northern southeastern Alaska (Mecklenburg et al. 2002). In our nearshore surveys, however, 

copper rockfish was widely distributed and common (FO = 27%) in outside waters (SENO), 

but were absent in inside waters (SENI). Similarly, least cisco is reported in nearshore waters 

of the Arctic (Mecklenburg et al. 2002), yet we captured least cisco only inside Elson Lagoon 

and not on the Beaufort Sea side of the lagoon. 

Northward range extensions of short distance were observed for a few species. Major 

poleward shifts in marine species are predicted with a warming climate (Lubchenco et al. 

1993); northern shifts are already documented for several marine fish species in the North 

Sea (Perry et al. 2005). The appearance of new species in marine ecosystems, including those 

of Alaska, could have deleterious effects on existing fish stocks and food webs. Baseline 

species distributions, like those presented in this synthesis, can help scientists, resource 

managers, and the public to recognize, identify, and monitor changes in Alaska’s nearshore 

fish assemblages. 

Abundance and Diversity

Abundance (CPUE) varied greatly among regions, whereas species richness generally 

increased along a north to south gradient. Variability in catch among regions was likely due 

to the unique physical and biological characteristics of each region (e.g., SC), sampling only 

a single habitat type (sand) in some regions (AR and BB), and the sometimes large catches of 

schooling species such as Pacific sand lance in AI and walleye pollock in SENI. Poor habitat 

quality (e.g., fast current, turbid water, and no cover) in parts of our SC region (Cook Inlet) 

likely accounted for this region having the lowest mean CPUE. On a smaller geographic 
26 

scale, differing biological and physical conditions among individual bays within a region 

(e.g., PWS) can cause spatial variation in the relative densities of juvenile Pacific herring 

(Norcross et al. 2001). Similarly, within Cook Inlet, gradients in physical oceanography 

influence the distribution and abundance of some zooplankton and fish species (Speckman et 

al. 2005). Diversity gradients of increasing species richness from high latitudes towards the 

equator have been shown for many taxonomic groups (Krebs 1972, Cardillo 2002, Parmesan 

2006). In our surveys, species richness increased from the Arctic (23) to southeastern Alaska 

(67, SENI). Bristol Bay had the lowest species richness (17) of any region, probably due to 

our limited sampling (n = 8 hauls) in only one habitat type (sand). 

Abundance and diversity of fishes can vary not only by region, but also by season and 

time of day. Two limited studies that we did were a seasonal (summer/winter) study in SENI 

and a diel study in PWS. At two locations in our SENI region, catch and species richness 

were much lower in winter than in summer (Thedinga et al. 2006a). Higher abundance and 

diversity of fishes in summer versus winter has also been reported in other nearshore studies 

(Allen and Horn 1975, Orth and Heck 1980, Methven et al. 2001). In our PWS region, 

abundance of fish during day and night was similar in eelgrass and kelp, but species 

composition and mean size of some fish changed; for example, mean size of saffron cod was 

similar between day and night in eelgrass, but was greatest during the day in kelp (Thedinga 

et al. 2011). More fish were captured in vegetated and nonvegetated habitats at night than 

during the day in Chesapeake Bay (Orth and Heck 1980). In a diel study in Morro Bay, 

California, larger numbers of individuals and greater biomass of fishes were collected at 

night, but nearly equal numbers of species were captured during the day and night (Horn 

1980). 

Three or fewer species comprised 55% to 76% of the total fish catch in all regions of 

Alaska. Taxa that dominated the total catch in our regions collectively included capelin, 

chum salmon, Pacific herring, Pacific sand lance, pink salmon, rainbow smelt, saffron cod, 

shiner perch, unidentified cod, and walleye pollock. With the exception of Pacific herring 

and shiner perch, all of these taxa are included in a FMP for Alaska, highlighting the 

extensive spatial use of nearshore habitats by managed and forage fish species. Dominance of 

a few species in fish catches has been reported in other nearshore studies in California, 
27 

Canada, and Maine (Allen and Horn 1975, Allen 1982, Methven et al. 2001, Lazzari and 

Tupper 2002). 

Based on our 14 years of sampling throughout coastal Alaska, we consider some areas 

to be noteworthy. Due to factors such as high species abundance, diversity, and susceptibility 

to disturbance, two regions and one location deserve special consideration. First, the Arctic is 

an ecologically fragile area experiencing rapid changes in climate (warming) and loss of sea 

ice (Moline et al. 2008). Loss of sea ice from climate change threatens marine life and habitat 

(e.g., beach erosion) and has the potential to open up formerly inaccessible areas to oil and 

gas development, vessel traffic, and commercial fishing. Disturbance of the nearshore Arctic 

environment may have consequences for many species, particularly capelin that use this area 

for spawning and rearing (George et al. 2007). Capelin is an important forage species in the 

Arctic and was the most abundant species in our catches. Second, the Aleutian Islands are 

noteworthy because of the extremely high abundance (CPUE = 1,056 fish) and frequency 

(FO = 94%) of Pacific sand lance in sand habitats. Nowhere else in Alaska did we witness 

such high and consistent catches of such an ecologically important species. Pacific sand lance 

is a major prey species for 40 species of birds, 12 species of marine mammals, and 45 species 

of fish (Field 1988, Willson et al. 1999). Third, The Brothers Islands in SENI (Appendix) is 

unique because of the abundance and diversity of commercially important and forage fish 

species. The six most abundant species in summer, based on total catch at The Brothers 

Islands, were walleye pollock, Pacific herring, Pacific sand lance, Pacific cod, chum salmon, 

and Pacific sandfish (Thedinga et al. 2006a); all of these species are commercially important 

or forage fish species. Our catches of walleye pollock at The Brothers Islands were greater 

than anywhere else in Alaska. In addition, The Brothers Islands is an important year-round, 

haul-out for about 1,500 Steller sea lions (Thedinga et al. 2006a). 

Species Composition and Habitat Use

Species composition by habitat type was similar among some regions and unique in 

others. Regions with similar species composition were usually close spatially and restricted 

to the Gulf of Alaska, whereas regions with unique species assemblages were more spatially 

isolated (e.g., AI, AR, and BB). Species composition was similar among several regions in 
28 

bedrock, but only between a few regions in eelgrass, kelp, and sand. Small sample sizes may 

have influenced some of our results, especially in bedrock (n = 4 for SC and SESI). Regions 

with unique species assemblages by habitat type (e.g., BB sand) were largely the result of the 

presence of a few species (e.g., rainbow smelt) in large numbers that were absent in other 

regions. Our AI region had a species composition that was consistently unique across all 

habitat types sampled (bedrock, kelp, and sand); small boundaries based on bottom 

topography and current flow in the Eastern Aleutians ecoregion may determine the 

distribution of some taxa (Piatt and Springer 2007). Species composition was notably 

different in eelgrass and kelp among most SE regions. Inside waters of southeastern Alaska 

are more estuarine, more protected from wave action, and have more extreme seasonal 

temperature and salinity fluctuations than outside waters (Johnson et al. 2008), which likely 

affects fish distribution. 

In most regions that we sampled with more than one habitat type, species generally 

segregated by habitat. Eelgrass and kelp usually supported higher numbers of fish and more 

species than bedrock or sand; mean CPUE and species richness were greatest in eelgrass or 

kelp in most regions. Higher fish abundance and species richness in vegetated versus sparsely 

vegetated habitats has been reported elsewhere (Orth and Heck 1980, Bloomfield and 

Gillanders 2005, Franco et al. 2006). Submerged vegetation provides increased structure and 

complexity, prey availability, and protection from predators compared to bare or sparsely 

vegetated habitats (Heck and Orth 1980, Rozas and Odum 1988, Lazzari and Tupper 2002, 

Hamilton and Konar 2007). In addition, some species were largely restricted to one 

vegetation type; for example, 98% of the total catch of saffron cod in PWS was in eelgrass. 

Similarly at Kodiak Island, Alaska, saffron cod were almost exclusively restricted to eelgrass 

(Laurel et al. 2007). 

Fish Life Stages

Nearshore ecosystems are considered nurseries for juvenile life stages of many marine 

species (Thayer et al. 1978, Beck et al. 2003, Lellis-Dibble 2008). Most commercially 

important and forage fish species that we captured (e.g., Pacific herring and walleye pollock) 

were early juveniles (e.g., larvae and young-of-the-year), and were likely using the nearshore 
29 

for foraging and refuge habitat (Johnson et al. 2008). We also know that the shallow 

nearshore is used for spawning, particularly by forage fishes: capelin and Pacific sand lance 

spawn on fine gravel or sandy beaches (Pahlke 1985, Robards et al. 1999b), Pacific herring 

spawn on intertidal and shallow, subtidal vegetation (Cooney 2007), and Pacific sandfish 

spawn on rocky intertidal areas (Marliave 1981). Because many commercially important and 

forage fish species depend on the nearshore environment for shelter, food, and spawning 

habitat, disturbance of the nearshore from shoreline development, oil spills, or natural 

catastrophes (e.g., earthquake) are of great concern. The Exxon Valdez oil spill in Alaska in 

1989 is a clear example of the susceptibility of coastal habitats and biota to disturbance. The 

spill occurred in March just prior to the time that Pacific herring spawn inshore and juvenile 

salmon and other fish species enter the nearshore marine environment (Spies et al. 1996). 

One of the more difficult tasks of nearshore sampling is the correct identification of 

fishes. The capture of sometimes thousands of fish of different species and life stages can be 

challenging even to the trained eye. Identifying juveniles in the field can be especially 

difficult for some species, including pink and chum salmon; Pacific cod, Pacific tomcod, and 

saffron cod; black and yellowtail rockfish; copper and quillback rockfish; frog, great, and 

shorthorn sculpin; and juvenile greenlings. Field guides that we commonly used to help 

identify fish include Kramer et al. (1995), Mecklenburg et al. (2002), and Lamb and Edgell 

(2010). All three of these guides cover mostly adult life stages, however, and we are now in 

the process of compiling a pictorial field guide that includes photos, including early lifehistory stages, of all species that we have captured. 

Forage Fishes

Forage fishes, as a group, probably best characterize the shallow nearshore in Alaska in 

terms of overall distribution, abundance, and habitat use. Eight forage fish species (capelin, 

longfin smelt, Pacific herring, Pacific sandfish, Pacific sand lance, pond smelt, rainbow 

smelt, and surf smelt (Hypomesus pretiosus)) accounted for 39% of our total overall catch 

and were collectively captured in 43% of our seine hauls. In seven of the nine regions that 

we sampled, capelin, Pacific herring, Pacific sand lance, or rainbow smelt was the most 

abundant species. Of all the forage fishes, Pacific sand lance was the most abundant and 
30 

ubiquitous (CPUE = 120 fish, FO = 23%; all Alaska). From an ecosystem perspective, it is 

well established that forage fishes are of particular importance as prey to other fishes, 

seabirds, and marine mammals (Springer and Speckman 1997, Mundy and Hollowed 2005). 

Our consistent catches of forage fishes throughout much of coastal Alaska, and the reliance 

of most species on the nearshore for spawning, clearly identifies the nearshore as important 

habitat for forage fish populations. 

Essential Fish Habitat

Identification of nearshore EFH is difficult because some species and life stages use 

certain habitats for only a short time (e.g., spawning) and then leave. Bedrock may be an 

example of a transitional habitat; fish may use this habitat type only short-term as they 

migrate from sheltered bays toward the open ocean (Wertheimer and Celewycz 1996), 

whereas eelgrass and kelp may be used as foraging areas for several weeks (Johnson et al. 

2008). Eelgrass and kelp should be considered high-value fish habitat based on the high 

abundance and diversity of fishes in our surveys. Eelgrass has long been recognized 

worldwide as valuable fish habitat (Green and Short 2003), and our knowledge of understory 

kelps as important fish habitat is increasing (Dean et al. 2000, Hamilton and Konar 2007, 

Johnson et al. 2005). The mere presence of large numbers of fish or species is not enough, 

however, to label certain coastal habitats as essential. More information is needed on the 

function (e.g., spawning sites) of these habitats to individual species to warrant special 

protection. Pacific herring is one of the few nearshore species in Alaska that we have 

sufficient information to identify EFH; spawning beaches are well documented from years of 

aerial surveys (Carls et al. 2008). 

The shallow nearshore should be included in any ecosystem-based marine study. 

Different species and life stages can use the nearshore at any time and can be vastly different 

from offshore fish assemblages. For example, some of the species that we routinely captured 

in the nearshore Beaufort Sea (e.g., capelin, least cisco, and saffron cod) were sparse or 

absent in bottom trawl surveys in deeper waters (>30 m) of the Beaufort Sea (Rand and 

Logerwell 2010). 
31 

Environmental Assessment

This synthesis is the most comprehensive summary available on the nearshore 

environment of Alaska, and provides a baseline for long-term monitoring. Sites can be 

revisited over time to track changes in fish and habitat that may result from global climate 

change or shoreline development. In addition, Alaska’s coastline is being imaged and 

mapped with ShoreZone, an aerial-based, mapping protocol that classifies coastal 

environments into different along-shore, biotic and coastal features (e.g., continuous eelgrass 

and very protected shoreline), and quantifies the linear extents of habitats (e.g., ~6,000 km of 

eelgrass in southeastern Alaska). Since 2003, over 69% (51,745 km) of Alaska’s coastline 

has been imaged with ShoreZone (NMFS 2012b), including 85% of our fish survey sites. In 

the future, our nearshore fish survey dataset and ShoreZone imagery may be the most 

important tools available to managers for oil spill response, natural resource damage 

assessment, EFH identification, shoreline development evaluation, and long-term biological 

monitoring. Both of these datasets are available online (NMFS 2012a, 2012b) and will 

continue to be updated as more areas of the state are surveyed and imaged. 

CONCLUSIONS 

A summary of our major findings from 14 years (1998 to 2011) of nearshore fish 

surveys throughout Alaska (nine regions) are as follows: 

 1. Nearshore, marine waters support an abundant and diverse array of fishes; we 

captured an estimated 718,345 fish representing at least 121 species from 29 

families. 

 2. Four commercially important species (walleye pollock, Pacific herring, pink 

salmon, and chum salmon) accounted for 55% of our total overall catch. 

 3. Species distribution patterns varied greatly among regions; only two species (pink 

salmon and threespine stickleback) were caught in all nine regions. 

 4. Abundance and species richness varied by region; mean CPUE ranged from 124 

fish in Southcentral Alaska to 1,202 fish in the Aleutian Islands, and species 
32 

richness ranged from 17 species in Bristol Bay to 67 species in Southeast – 

Northern Inside. 

 5. Species assemblages differed among habitat types in most regions, and mean 

CPUE and species richness were usually greatest in eelgrass or kelp. 

 6. Species composition by habitat type was unique in some regions (e.g., sand, 

Bristol Bay), largely the result of the presence of a few species (e.g., rainbow 

smelt) that were absent in other regions. 

 7. Most fishes captured were juveniles, highlighting the importance of the nearshore 

to critical, early life-history stages of managed and ecologically important species. 

 8. Forage species (e.g., Pacific herring and Pacific sand lance) may be the most 

susceptible fishes to shoreline disturbance because of their use of nearshore 

habitats for feeding and shelter as juveniles and for spawning as adults. 

 9. The high abundance and diversity of fishes in eelgrass and kelp warrant special 

protection of these habitats in the event of an oil spill or other shoreline 

disturbance, especially if these habitats are known spawning areas (e.g., Pacific 

herring). 

 10. Our nearshore dataset is extremely important to managers responsible for oil spill 

response, natural resource damage assessment, EFH identification, and long-term 

monitoring. 

THE ATLAS 

Individual species pages and photos are presented for 121 fish species that we captured 

with a beach seine in nearshore waters of Alaska from 1998 to 2011. Species page and photo 

sections are arranged alphabetically by family first, and then by scientific name within 

family. Species pages are designed to quickly show distribution, abundance, habitat use, and 

length frequency information for each given species. The photo section provides a high 

resolution photo of every species captured; most were of actual fish that we caught. Length 

(millimeters) and life stage (juvenile, adult) information are provided for each photo; FL is 

fork length and TL is total length. 
33 

Each species page is standardized with the same layout: name banner, photo, graphs, 

and distribution maps. The name banner lists the species family name, common name, and 

scientific name from left to right. The fish photo shows the species overall appearance; a 

larger image is available on the page listed. Directly below the photo are graphs showing 

abundance and length frequency data. The large panel on the right side of each species page 

displays maps of the geographic distribution of the species by region. 

Abundance and length frequency graphs represent data from all nearshore surveys 

(1998 to 2011) encompassing a total of 1,009 seine hauls. Two abundance indices – mean 

catch-per-unit-effort (CPUE), where unit of effort = seine haul and percent frequency of 

occurrence (FO) – are shown by region (top graph) and habitat type (middle graph). First, 

mean CPUE by region is the species total catch by region, across all habitat types, divided by 

the total number of seine hauls by region (Table 3). Mean CPUE by habitat type is the 

species total catch by habitat type, across all regions, divided by the total number of seine 

hauls by habitat type (Table 4). Mean CPUE ranges from 0.1 fish to 1,000 fish (Y-axis, 

logarithmic scale). Second, percent FO by region is the total number of seine hauls in which 

the species was captured by region, across all habitat types, divided by the total number of 

seine hauls by region and multiplied by 100 (Table 3). Percent FO by habitat type is the total 

number of seine hauls in which the species was captured by habitat type, across all regions, 

divided by the total number of seine hauls by habitat type and multiplied by 100 (Table 4). 

Percent FO ranges from 0 to 100% (Y2-axis). The “n” represents the total catch of each 

species across all regions, habitat types, and sampling periods. Region acronyms (top graph, 

X-axis) are labeled and defined (e.g., AR, Arctic) in the distribution panel. Habitat types 

(middle graph, X-axis) are bedrock, eelgrass, kelp, and sand. The bottom graph displays 

length frequency data across all regions and habitat types. Percent frequency (0-100%, 

Y-axis) represents the number of fish measured within the different length classes (X-axis), 

ranging from < 50 mm to  400 mm, divided by the total number of fish measured. Length in 

millimeters is either fork length (FL) or total length (TL), depending on species (Table 5). 

The “n” represents the total number of fish measured across all regions, habitat types, and 

sampling periods. 
34 

Distribution maps show each of the nine regions that we sampled and are labeled 

accordingly (e.g., Arctic, AR and Aleutian Islands, AI). Within each regional map, sampling 

locations are shown as solid circles; large blue circles indicate species presence, and small 

black circles indicate species absence. The “n” represents the total number of seine hauls in 

each region. Maps of each region with key landmarks and individual sampling sites are 

shown in Figures 4-12. 

 
35 

Species List 

 
36 

Family Taxon Common name Page

*

Agonidae Agonopsis vulsa Northern spearnose poacher 40, 160

Leptagonus decagonus Atlantic poacher 41, 160

Occella dodecaedron Bering poacher 42, 160

Odontopyxis trispinosa Pygmy poacher 43, 160

Pallasina barbata Tubenose poacher 44, 160

Podothecus accipenserinus Sturgeon poacher 45, 160

Podothecus veternus Veteran poacher 46, 160

Ammodytidae Ammodytes hexapterus Pacific sand lance 47, 160

Anarhichadidae Anarrhichthys ocellatus Wolf-eel 48, 161

Anoplopomatidae Anoplopoma fimbria Sablefish 49, 161

Aulorhynchidae Aulorhynchus flavidus Tubesnout 50, 161

Bathymasteridae Bathymaster caeruleofasciatus Alaskan ronquil 51, 161

Bathymaster leurolepis Smallmouth ronquil 52, 161

Bathymaster signatus Searcher 53, 161

Ronquilus jordani Northern ronquil 54, 161

Clupeidae Clupea pallasii Pacific herring 55, 161

Cottidae Artedius fenestralis Padded sculpin 56, 162

Artedius harringtoni Scalyhead sculpin 57, 162

Artedius lateralis Smoothhead sculpin 58, 162

Chitonotus pugetensis Roughback sculpin 59, 162

Enophrys bison Buffalo sculpin 60, 162

Enophrys diceraus Antlered sculpin 61, 162

Enophrys lucasi Leister sculpin 62, 162

Gymnocanthus galeatus Armorhead sculpin 63, 162

Gymnocanthus tricuspis Arctic staghorn sculpin 64, 163

Hemilepidotus hemilepidotus Red Irish lord 65, 163

Hemilepidotus spinosus Brown Irish lord 66, 163

Icelinus borealis Northern sculpin 67, 163

Leptocottus armatus Pacific staghorn sculpin 68, 163

Megalocottus platycephalus Belligerent sculpin 69, 163

Microcottus sellaris Brightbelly sculpin 70, 163

Myoxocephalus jaok Plain sculpin 71, 163

Myoxocephalus polyacanthocephalus Great sculpin 74, 164

Myoxocephalus quadricornis Fourhorn sculpin 72, 164

Myoxocephalus scorpioides Arctic sculpin 73, 164

Myoxocephalus scorpius Shorthorn sculpin 74, 164

Myoxocephalus stelleri Frog sculpin 74, 164

Oligocottus maculosus Tidepool sculpin 75, 164

Oligocottus rimensis Saddleback sculpin 76, 164

Oligocottus snyderi Fluffy sculpin 77, 164

Scorpaenichthys marmoratus Cabezon 78, 165

Synchirus gilli Manacled sculpin 79, 165 *

species page, species photo
37 

Family Taxon Common name Page

*

Cyclopteridae Eumicrotremus orbis Pacific spiny lumpsucker 80, 165

Embiotocidae Brachyistius frenatus Kelp perch 81, 165

Cymatogaster aggregata Shiner perch 82, 165

Gadidae Boreogadus saida Arctic cod 83, 165

Eleginus gracilis Saffron cod 84, 165

Gadus macrocephalus Pacific cod 85, 165

Microgadus proximus Pacific tomcod 86, 166

Theragra chalcogramma Walleye pollock 87, 166

Gasterosteidae Gasterosteus aculeatus Threespine stickleback 88, 166

Pungitius pungitius Ninespine stickleback 89, 166

Gobiesocidae Rimicola muscarum Kelp clingfish 90, 166

Gobiidae Lepidogobius lepidus Bay goby 91, 166

Rhinogobiops nicholsii Blackeye goby 92, 166

Hemitripteridae Blepsias bilobus Crested sculpin 93, 166

Blepsias cirrhosus Silverspotted sculpin 94, 167

Nautichthys oculofasciatus Sailfin sculpin 95, 167

Hexagrammidae Hexagrammos decagrammus Kelp greenling 96, 167

Hexagrammos lagocephalus Rock greenling 97, 167

Hexagrammos octogrammus Masked greenling 98, 167

Hexagrammos stelleri Whitespotted greenling 99, 167

Ophiodon elongatus Lingcod 100, 167

Oxylebius pictus Painted greenling 101, 167

Liparidae Liparis callyodon Spotted snailfish 102, 168

Liparis cyclopus Ribbon snailfish 103, 168

Liparis florae Tidepool snailfish 104, 168

Liparis gibbus Variegated snailfish 105, 168

Liparis pulchellus Showy snailfish 106, 168

Osmeridae Hypomesus olidus Pond smelt 107, 168

Hypomesus pretiosus Surf smelt 108, 168

Mallotus villosus Capelin 109, 168

Osmerus mordax Rainbow smelt 110, 169

Spirinchus thaleichthys Longfin smelt 111, 169

Thaleichthys pacificus Eulachon 112, 169

Paralichthyidae Citharichthys sordidus Pacific sanddab 113, 169

Citharichthys stigmaeus Speckled sanddab 114, 169

Petromyzontidae Lampetra camtschatica Arctic lamprey 115, 169

Pholidae Apodichthys flavidus Penpoint gunnel 116, 169

Pholis laeta Crescent gunnel 117, 169

Pleuronectidae Isopsetta isolepis Butter sole 118, 170

Lepidopsetta spp. Rock sole 119, 170

Limanda aspera Yellowfin sole 120, 170

Limanda proboscidea Longhead dab 121, 170 *

species page, species photo
38 

Family Taxon Common name Page

*

Microstomus pacificus Dover sole 122, 170

Parophrys vetulus English sole 123, 170

Platichthys stellatus Starry flounder 124, 170

Pleuronectes glacialis Arctic flounder 125, 170

Pleuronectes quadrituberculatus Alaska plaice 126, 171

Pleuronichthys coenosus C-O sole 127, 171

Psettichthys melanostictus Sand sole 128, 171

Psychrolutidae Psychrolutes paradoxus Tadpole sculpin 129, 171

Psychrolutes sigalutes Soft sculpin 130, 171

Salmonidae Coregonus autumnalis Arctic cisco 131, 171

Coregonus sardinella Least cisco 132, 171

Oncorhynchus clarkii Cutthroat trout 133, 172

Oncorhynchus gorbuscha Pink salmon 134, 172

Oncorhynchus keta Chum salmon 135, 172

Oncorhynchus kisutch Coho salmon 136, 172

Oncorhynchus mykiss Steelhead trout 137, 172

Oncorhynchus nerka Sockeye salmon 138, 172

Oncorhynchus tshawytscha Chinook salmon 139, 172

Salvelinus malma Dolly Varden 140, 172

Scorpaenidae Sebastes auriculatus Brown rockfish 141, 173

Sebastes caurinus Copper rockfish 142, 173

Sebastes ciliatus Dark rockfish1 143, 173

Sebastes flavidus Yellowtail rockfish 144, 173

Sebastes maliger Quillback rockfish 145, 173

Sebastes melanops Black rockfish 146, 173

Sebastes nebulosus China rockfish 147, 173

Sebastes paucispinis Bocaccio 148, 173

Stichaeidae Acantholumpenus mackayi Blackline prickleback 149, 174

Anoplarchus insignis Slender cockscomb 150, 174

Eumesogrammus praecisus Fourline snakeblenny 151, 174

Lumpenus fabricii Slender eelblenny 152, 174

Lumpenus sagitta Snake prickleback 153, 174

Stichaeus punctatus Arctic shanny 154, 174

Xiphister atropurpureus Black prickleback 155, 174

Syngnathidae Syngnathus leptorhynchus Bay pipefish 156, 174

Trichodontidae Trichodon trichodon Pacific sandfish 157, 175

Zaproridae Zaprora silenus Prowfish 158, 175 *

species page, species photo 1

formerly the dark form of the dusky rockfish (Orr and Blackburn 2004) 

 
39 

Species Pages 
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Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Northern spearnose poacher

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

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Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 160

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

!

Northern spearnose poacher

Capture locations

Agonopsis vulsa Agonidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 4n = 3

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

40
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Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Atlantic poacher

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

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Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 160

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

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!!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Atlantic poacher Leptagonus decagonus Agonidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 1n = 1

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

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Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Bering poacher

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

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Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 160

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

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Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Bering poacher Occella dodecaedron Agonidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 1n = 1

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

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Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Pygmy poacher

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

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Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 160

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

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Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Pygmy poacher Odontopyxis trispinosa Agonidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 1n = 1

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

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Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Tubenose poacher

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

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% Frequency

Length (millimeters)

100

50

0

p. 160

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

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Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Tubenose poacher Pallasina barbata Agonidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 2,644n = 287

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

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Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Sturgeon poacher

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

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% Frequency

Length (millimeters)

100

50

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p. 160

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

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0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Sturgeon poacher Podothecus accipenserinus Agonidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 228n = 20

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

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Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Veteran poacher

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

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% Frequency

Length (millimeters)

100

50

0

p. 160

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

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0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Veteran poacher Podothecus veternus Agonidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 2n = 2

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

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Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Pacific sand lance

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

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% Frequency

Length (millimeters)

100

50

0

p. 160

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

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0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Pacific sand lance Ammodytes hexapterus Ammodytidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 121,089n = 3,324

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

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Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Wolf-eel

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

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Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 161

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

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0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Wolf-eel Anarrhichthys ocellatus Anarhichadidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 1n = 0

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

48
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Sablefish

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 161

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Sablefish Anoplopoma fimbria Anoplopomatidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 1n = 1

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

49
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!!!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Tubesnout

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

!

!

!!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 161

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Tubesnout Aulorhynchus flavidus Aulorhynchidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 7,614n = 1,151

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

50
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Alaskan ronquil

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 161

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Alaskan ronquil Bathymaster caeruleofasciatus Bathymasteridae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 4n = 0

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

51
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Smallmouth ronquil

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 161

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Smallmouth ronquil Bathymaster leurolepis Bathymasteridae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 1n = 1

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

52
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Searcher

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 161

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Searcher Bathymaster signatus Bathymasteridae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 2n = 2

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

53
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Northern ronquil

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 161

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Northern ronquil Ronquilus jordani Bathymasteridae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 6n = 6

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

54
!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Pacific herring

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

!

!

!!

!

!

!

!

! Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 161

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Pacific herring Clupea pallasii Clupeidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 106,451n = 3,126

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

55
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Padded sculpin

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

!

!

!!

!

!

!

! Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 162

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Padded sculpin Artedius fenestralis Cottidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 466n = 429

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

56
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Scalyhead sculpin

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

!!

!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 162

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Scalyhead sculpin Artedius harringtoni Cottidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 15n = 15

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

57
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Smoothhead sculpin

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 162

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Smoothhead sculpin Artedius lateralis Cottidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 17n = 14

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

58
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Roughback sculpin

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 162

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Roughback sculpin Chitonotus pugetensis Cottidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 13n = 13

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

59
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Buffalo sculpin

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

!

! Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 162

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Buffalo sculpin Enophrys bison Cottidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 354n = 78

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

60
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Antlered sculpin

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 162

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Antlered sculpin Enophrys diceraus Cottidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 7n = 7

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

61
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Leister sculpin

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 162

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Leister sculpin Enophrys lucasi Cottidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 40n = 21

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

62
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Armorhead sculpin

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 162

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Armorhead sculpin Gymnocanthus galeatus Cottidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 202n = 47

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

63
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Arctic staghorn sculpin

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 163

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Arctic staghorn sculpin Gymnocanthus tricuspis Cottidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 5n = 5

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

64
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!!

!

!

!!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!!

!

!!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Red Irish lord

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 163

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Red Irish lord Hemilepidotus hemilepidotus Cottidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 215n = 46

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

65
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Brown Irish lord

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 163

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Brown Irish lord Hemilepidotus spinosus Cottidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 19n = 19

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

66
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!!

!!

!

!!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!!!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Northern sculpin

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 163

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Northern sculpin Icelinus borealis Cottidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 1,265n = 59

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

67
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Pacific staghorn sculpin

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!!

! Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 163

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Pacific staghorn sculpin Leptocottus armatus Cottidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 1,157n = 253

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

68
!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Belligerent sculpin

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 163

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Belligerent sculpin Megalocottus platycephalus Cottidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 6n = 6

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

69
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Brightbelly sculpin

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

!

% Frequency

Length (millimeters)

100

50

0

p. 163

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Brightbelly sculpin Microcottus sellaris Cottidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 2n = 0

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

70
!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Plain sculpin

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

!

% Frequency

Length (millimeters)

100

50

0

p. 163

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Plain sculpin Myoxocephalus jaok Cottidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 5n = 5

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

71
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Fourhorn sculpin

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 164

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Fourhorn sculpin Myoxocephalus quadricornis Cottidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 6n = 6

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

72
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Arctic sculpin

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 164

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Arctic sculpin Myoxocephalus scorpioides Cottidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 40n = 40

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

73
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Frog, great, & shorthorn sculpin

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!!

!

!

!

!

!!

! Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 164

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Frog, great, & shorthorn sculpin Myoxocephalus spp. Cottidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 1,390n = 519

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

Frog Great

Shorthorn

74
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Tidepool sculpin

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

!

!

!!

!

!

!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 164

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Tidepool sculpin Oligocottus maculosus Cottidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 172n = 128

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

75
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Saddleback sculpin

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 164

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Saddleback sculpin Oligocottus rimensis Cottidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 1n = 1

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

76
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Fluffy sculpin

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 164

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Fluffy sculpin Oligocottus snyderi Cottidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 5n = 0

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

77
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Cabezon

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 165

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Cabezon Scorpaenichthys marmoratus Cottidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 8n = 0

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

78
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Manacled sculpin

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 165

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Manacled sculpin Synchirus gilli Cottidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 167n = 67

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

79
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Pacific spiny lumpsucker

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 165

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Pacific spiny lumpsucker Eumicrotremus orbis Cyclopteridae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 13n = 6

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

80
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Kelp perch

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 165

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Kelp perch Brachyistius frenatus Embiotocidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 125n = 78

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

81
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Shiner perch

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 165

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Shiner perch Cymatogaster aggregata Embiotocidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 29,000n = 2,662

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

82
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Arctic cod

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 165

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Arctic cod Boreogadus saida Gadidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 439n = 139

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

83
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Saffron cod

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

!

!

!!

!

!

!

! Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 165

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Saffron cod Eleginus gracilis Gadidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 17,492n = 2,316

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

84
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Pacific cod

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!!

! Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 165

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Pacific cod Gadus macrocephalus Gadidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 8,522n = 2,089

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

85
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Pacific tomcod

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

! Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 166

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Pacific tomcod Microgadus proximus Gadidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 16n = 16

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

86
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Walleye pollock

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

!

!!

!

!

!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 166

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Walleye pollock Theragra chalcogramma Gadidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 124,524n = 2,517

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

87
!

!

!

!

!

!

!

!

!!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Threespine stickleback

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

!

!

!!

!

!

!!

! Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 166

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Threespine stickleback Gasterosteus aculeatus Gasterosteidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 8,179n = 478

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

88
!

!

!

!

!

!

!!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Ninespine stickleback

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 166

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Ninespine stickleback Pungitius pungitius Gasterosteidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 36n = 35

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

89
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Kelp clingfish

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 166

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Kelp clingfish Rimicola muscarum Gobiescocidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 1n = 0

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

John Yasaki

90
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Bay goby

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 166

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Bay goby Lepidogobius lepidus Gobiidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 16n = 6

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

91
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Blackeye goby

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 166

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Blackeye goby Rhinogobiops nicholsii Gobiidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 108n = 22

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

92
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Crested sculpin

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 166

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Crested sculpin Blepsias bilobus Hemitripteridae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 29n = 11

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

93
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Silverspotted sculpin

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!!

! Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 167

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Silverspotted sculpin Blepsias cirrhosus Hemitripteridae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 1,867n = 226

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

94
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Sailfin sculpin

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 167

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Sailfin sculpin Nautichthys oculofasciatus Hemitripteridae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 18n = 4

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

95
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Kelp greenling

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 167

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Kelp greenling Hexagrammos decagrammus Hexagrammidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 303n = 170

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

96
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!!

!!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Rock greenling

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 167

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Rock greenling Hexagrammos lagocephalus Hexagrammidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 164n = 135

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

97
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Masked greenling

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

!

!

!!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 167

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Masked greenling Hexagrammos octogrammus Hexagrammidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 360n = 231

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

98
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Whitespotted greenling

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

!

!

!!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 167

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Whitespotted greenling Hexagrammos stelleri Hexagrammidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 719n = 591

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

99
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Lingcod

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

!

!!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 167

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Lingcod Ophiodon elongatus Hexagrammidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 336n = 196

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

100
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Painted greenling

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 167

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Painted greenling Oxylebius pictus Hexagrammidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 49n = 24

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

101
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Spotted snailfish

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 168

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Spotted snailfish Liparis callyodon Liparidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 3n = 3

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

102
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Ribbon snailfish

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 168

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Ribbon snailfish Liparis cyclopus Liparidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 1n = 1

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

103
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Tidepool snailfish

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 168

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Tidepool snailfish Liparis florae Liparidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 13n = 2

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

104
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Variegated snailfish

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 168

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Variegated snailfish Liparis gibbus Liparidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 2n = 2

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

105
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Showy snailfish

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 168

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Showy snailfish Liparis pulchellus Liparidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 7n = 0

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

Keith Clements

106
!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Pond smelt

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 168

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Pond smelt Hypomesus olidus Osmeridae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 1,331n = 202

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

107
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Surf smelt

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 168

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Surf smelt Hypomesus pretiosus Osmeridae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 932n = 326

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

108
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Capelin

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

!!!

!

!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 168

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Capelin Mallotus villosus Osmeridae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 21,770n = 1,305

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

109
!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Rainbow smelt

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 169

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Rainbow smelt Osmerus mordax Osmeridae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 5,117n = 319

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

110
!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Longfin smelt

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 169

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Longfin smelt Spirinchus thaleichthys Osmeridae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 27n = 27

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

111
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Eulachon

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 169

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Eulachon Thaleichthys pacificus Osmeridae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 6n = 6

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

112
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Pacific sanddab

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 169

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Pacific sanddab Citharichthys sordidus Paralichthyidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 83n = 48

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

113
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Speckled sanddab

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!!

!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 169

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Speckled sanddab Citharichthys stigmaeus Paralichthyidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 82n = 76

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

114
!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Arctic lamprey

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 169

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Arctic lamprey Lampetra camtschatica Petromyzontidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 2n = 2

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

115
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Penpoint gunnel

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 169

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Penpoint gunnel Apodichthys flavidus Pholidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 65n = 26

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

116
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Crescent gunnel

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

!

!

!!

!

!

!

!

!!

! Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 169

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Crescent gunnel Pholis laeta Pholidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 12,222n = 275

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

117
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Butter sole

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 170

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Butter sole Isopsetta isolepis Pleuronectidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 2n = 0

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

118
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Rock sole

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

!

% Frequency

Length (millimeters)

100

50

0

p. 170

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Rock sole Lepidopsetta spp. Pleuronectidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 801n = 539

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

119
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Yellowfin sole

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 170

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Yellowfin sole Limanda aspera Pleuronectidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 108n = 76

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

120
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Longhead dab

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 170

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Longhead dab Limanda proboscidea Pleuronectidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 12n = 12

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

121
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Dover sole

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 170

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Dover sole Microstomus pacificus Pleuronectidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 1n = 1

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

122
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

English sole

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 170

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

English sole Parophrys vetulus Pleuronectidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 186n = 80

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

123
!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Starry flounder

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

! Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 170

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Starry flounder Platichthys stellatus Pleuronectidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 261n = 188

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

124
!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Arctic flounder

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 170

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Arctic flounder Pleuronectes glacialis Pleuronectidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 11n = 11

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

125
!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Alaska plaice

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 171

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Alaska plaice Pleuronectes quadrituberculatus Pleuronectidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 6n = 6

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

126
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

C-O sole

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 171

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

C-O sole Pleuronichthys coenosus Pleuronectidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 5n = 4

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

127
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Sand sole

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 171

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Sand sole Psettichthys melanostictus Pleuronectidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 10n = 2

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

128
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Tadpole sculpin

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 171

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Tadpole sculpin Psychrolutes paradoxus Psychrolutidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 19n = 0

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

129
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Soft sculpin

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 171

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Soft sculpin Psychrolutes sigalutes Psychrolutidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 2n = 2

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

130
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Arctic cisco

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 171

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Arctic cisco Coregonus autumnalis Salmonidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 3n = 3

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

131
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Least cisco

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 171

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Least cisco Coregonus sardinella Salmonidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 39n = 39

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

132
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Cutthroat trout

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 172

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Cutthroat trout Oncorhynchus clarkii Salmonidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 28n = 25

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

133
!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Pink salmon

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

!

!

!!

!

!

!

! Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 172

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Pink salmon Oncorhynchus gorbuscha Salmonidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 89,556n = 4,472

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

134
!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Chum salmon

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

!

!

!!

!

!

!

! Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 172

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Chum salmon Oncorhynchus keta Salmonidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 76,385n = 3,856

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

135
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Coho salmon

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

!!

!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 172

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Coho salmon Oncorhynchus kisutch Salmonidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 3,100n = 1,349

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

136
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Steelhead trout

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 172

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Steelhead trout Oncorhynchus mykiss Salmonidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 10n = 10

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

137
!

!

!

!

!

!

!

!!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Sockeye salmon

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 172

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Sockeye salmon Oncorhynchus nerka Salmonidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 44n = 32

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

138
!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Chinook salmon

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 172

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Chinook salmon Oncorhynchus tshawytscha Salmonidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 9n = 7

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

139
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!!

!

!!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Dolly Varden

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

! Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 172

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Dolly Varden Salvelinus malma Salmonidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 529n = 420

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

140
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Brown rockfish

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 173

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Brown rockfish Sebastes auriculatus Scorpaenidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 54n = 53

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

141
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!!

!!

!

!

!!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Copper rockfish

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

!

!

!!

!

!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 173

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Copper rockfish Sebastes caurinus Scorpaenidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 886n = 710

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

142
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Dark rockfish

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 173

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Dark rockfish Sebastes ciliatus Scorpaenidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 62n = 60

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

143
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Yellowtail rockfish

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 173

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Yellowtail rockfish Sebastes flavidus Scorpaenidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 3n = 2

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

144
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Quillback rockfish

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 173

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Quillback rockfish Sebastes maliger Scorpaenidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 148n = 118

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

145
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!!

!!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Black rockfish

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 173

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Black rockfish Sebastes melanops Scorpaenidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 191n = 183

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

146
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

China rockfish

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 173

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

China rockfish Sebastes nebulosus Scorpaenidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 1n = 1

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

147
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Bocaccio

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 173

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Bocaccio Sebastes paucispinis Scorpaenidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 1n = 1

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

148
!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Blackline prickleback

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 174

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Blackline prickleback Acantholumpenus mackayi Stichaeidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 33n = 32

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

149
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Slender cockscomb

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

! Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 174

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Slender cockscomb Anoplarchus insignis Stichaeidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 10n = 6

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

150
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Fourline snakeblenny

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 174

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Fourline snakeblenny Eumesogrammus praecisus Stichaeidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 1n = 1

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

151
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Slender eelblenny

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 174

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Slender eelblenny Lumpenus fabricii Stichaeidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 55n = 51

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

152
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!!!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Snake prickleback

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

!

!!

!

!

!!

! Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 174

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Snake prickleback Lumpenus sagitta Stichaeidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 5,885n = 522

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

153
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Arctic shanny

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

!

!

!!

!

!

!

!

!!

! Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 174

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Arctic shanny Stichaeus punctatus Stichaeidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 287n = 161

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

154
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Black prickleback

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 174

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Black prickleback Xiphister atropurpureus Stichaeidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 4n = 1

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

155
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

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!

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!

!

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!

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!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!!!

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!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Bay pipefish

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

!

!

!

!

!!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 174

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Bay pipefish Syngnathus leptorhynchus Syngnathidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 8,054n = 592

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

156
!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

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!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Pacific sandfish

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 175

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Pacific sandfish Trichodon trichodon Trichodontidae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 21,943n = 815

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

157
!

!

!

!

!

!

!

!

!

!

!

!

!

!

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!

!

!

!

!

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!

!

!

!

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!

!

!

!

!

!

!!!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!!

!

!

!

!

!

!

!

!

!

!

!

!

Aleutian Islands (AI)

Bristol Bay (BB)

Southcentral (SC)

Southeast

Northern Outside

(SENO)

Southeast

Northern Inside (SENI)

Southeast

Southern Outside (SESO)

Southeast

Southern Inside (SESI)

Prowfish

Mean catch-per-unit-effort ( )

% Frequency of occurrence ( )

Length Frequency for All Regions and Habitats

0 25 50 km

0 25 50 km

0 25 50 km 0 25 50 km

0 25 50 km

0 25 50 km0 25 50 km

!

!

!

!

!!

!

!

!

!

!

!!

Prince William Sound (PWS)

% Frequency

Length (millimeters)

100

50

0

p. 175

0 25 50 km

AR

Alaska

BB

AI

SC

PWS

SENI

SENO

SESO

SESI

!

!

Arctic (AR)

0 25 50 km

Geographic Distribution

! Sample locations

! Capture locations

Prowfish Zaprora silenus Zaproridae

100500

Sand Bedrock Eelgrass Kelp

1,000

100

10

1

0.1

1,000

100

10

1

0.1

AR AI BB SC PWS SENO SENI SESO SESI

100500

<50

50-99

100-149

150-199

200-249

250-299

300-399

400 _

>

Abundance Indices by Region and Habitat

n = 1n = 1

n = 81

n = 70

n = 8

n = 23

n = 133

n = 164

n = 382

n = 114

n = 34

158
Species Photos 

159
Veteran poacher – Juvenile, 43 mm TL

Agonidae: Podothecus veternus

Tubenose poacher – Adult, 138 mm TL

Agonidae: Pallasina barbata

Sturgeon poacher – Adult, 196 mm TL

Agonidae: Podothecus accipenserinus

Pygmy poacher – Adult, 72 mm TL

Agonidae: Odontopyxis trispinosa

	
	
		
			

Ammodytidae: Ammodytes hexapterus

Bering poacher – Juvenile, 110 mm TL

Agonidae: Occella dodecaedron

Atlantic poacher – Juvenile, 28 mm TL

Agonidae: Leptagonus decagonus

Northern spearnose poacher – Juvenile, 115 mm TL

Agonidae: Agonopsis vulsa

160
Wolf-eel – Juvenile

Anarhichadidae: Anarrhichthys ocellatus

Tubesnout – Adult, 165 mm TL

Aulorhynchidae: 	



Smallmouth ronquil – Juvenile, 123 mm TL

Bathymasteridae: Bathymaster leurolepis



		
			

Anoplopomatidae: 	

Northern ronquil – Juvenile, 108 mm TL

Bathymasteridae: Ronquilus jordani

	!!"		
	#&		

Clupeidae: Clupea pallasii

Alaskan ronquil – Adult, 256 mm TL 

Bathymasteridae: Bathymaster caeruleofasciatus

Searcher – Adult

Bathymasteridae: Bathymaster signatus

161
Smoothhead sculpin – Juvenile, 87 mm TL

Cottidae: Artedius lateralis

Roughback sculpin – Juvenile, ~70 mm TL

Cottidae: Chitonotus pugetensis

Leister sculpin – Adult, ~150 mm TL

Cottidae: Enophrys lucasi

Buffalo sculpin – Juvenile, 100 mm TL

Cottidae: Enophrys bison

Armorhead sculpin – Adult, 250 mm TL

Cottidae: Gymnocanthus galeatus

Antlered sculpin – Juvenile, ~100 mm TL

Cottidae: Enophrys diceraus

Padded sculpin – Juvenile, 77 mm TL

Cottidae: Artedius fenestralis

Scalyhead sculpin – Adult, 75 mm TL

Cottidae: Artedius harringtoni

162
	
"'!	

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	@[		\

Cottidae: Leptocottus armatus

Red Irish lord – Adult, 378 mm TL

Cottidae: Hemilepidotus hemilepidotus

Plain sculpin – Juvenile, 296 mm TL

Cottidae: Myoxocephalus jaok

Northern sculpin – Juvenile, 57 mm TL

Cottidae: Icelinus borealis

Brown Irish lord – Juvenile, 81 mm TL

Cottidae: Hemilepidotus spinosus

Belligerent sculpin – Juvenile, 235 mm TL

Cottidae: Megalocottus platycephalus

Brightbelly sculpin – Adult, 110 mm TL

Cottidae: Microcottus sellaris

Arctic staghorn sculpin – Juvenile, 110 mm TL

Cottidae: Gymnocanthus tricuspis

163
Tidepool sculpin – Juvenile, 41 mm TL

Cottidae: Oligocottus maculosus

Shorthorn sculpin – Juvenile, 135 mm TL

Cottidae: Myoxocephalus scorpius

Saddleback sculpin – Juvenile, 47 mm TL

Cottidae: Oligocottus rimensis

'!'!	

?		
	#		\

Cottidae: Myoxocephalus quadricornis

Great sculpin – Juvenile, 136 mm TL

Cottidae: Myoxocephalus polyacanthocephalus

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	]		\

Cottidae: Myoxocephalus stelleri

Arctic sculpin – Juvenile, 72 mm TL

Cottidae: Myoxocephalus scorpioides


^^_	

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Cottidae: Oligocottus snyderi

164
Cabezon – Juvenile, 112 mm TL

Cottidae: Scorpaenichthys marmoratus

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	@[		

Embiotocidae: Brachyistius frenatus

!	'		
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Gadidae: Boreogadus saida

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Gadidae: Gadus macrocephalus

	
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Cyclopteridae: Eumicrotremus orbis

!	?!		
	&&		

Embiotocidae: Cymatogaster aggregata

Saffron cod – Juvenile, 265 mm TL

Gadidae: Eleginus gracilis

Manacled sculpin – Adult, 55 mm TL

Cottidae: Synchirus gilli

165
Walleye pollock – Juvenile, ~200 mm TL

Gadidae: Theragra chalcogramma

Threespine stickleback – Adult, 59 mm TL

Gasterosteidae: Gasterosteus aculeatus

	''		
			\

Gadidae: Microgadus proximus

Crested sculpin – Juvenile, 106 mm TL

Hemitripteridae: Blepsias bilobus

Blackeye goby – Juvenile, 79 mm TL

Gobiidae: Rhinogobiops nicholsii

Ninespine stickleback – Juvenile, 46 mm TL

Gasterosteidae: Pungitius pungitius

John Yasaki

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	@		\

Gobiescocidae: Rimicola muscarum

Bay goby – Adult, 87 mm TL

Gobiidae: Lepidogobius lepidus

166
Lingcod – Juvenile, 250 mm TL

Hexagrammidae: Ophiodon elongatus


	

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	@&		\

Hemitripteridae: Nautichthys oculofasciatus

Kelp greenling – Juvenile, 122 mm TL

Hexagrammidae: Hexagrammos decagrammus

Masked greenling – Juvenile, 207 mm TL

Hexagrammidae: Hexagrammos octogrammus

Painted greenling – Juvenile, 118 mm TL

Hexagrammidae: Oxylebius pictus

Rock greenling – Juvenile, 200 mm TL

Hexagrammidae: Hexagrammos lagocephalus

Silverspotted sculpin – Adult, 135 mm TL

Hemitripteridae: Blepsias cirrhosus

Whitespotted greenling – Juvenile, 163 mm TL

Hexagrammidae: Hexagrammos stelleri

167
'|_	


		
	

Liparidae: Liparis pulchellus

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	]		\

Liparidae: Liparis gibbus

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	&		

Osmeridae: Mallotus villosus

'	


		
			\

Liparidae: Liparis cyclopus

'	

		
			

Osmeridae: Hypomesus olidus

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			\

Liparidae: 


!^	

		
			

Osmeridae: Hypomesus pretiosus

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	[#		\

Liparidae: Liparis callyodon

Keith Clements

168
Arctic lamprey – Juvenile, 161 mm TL

Petromyzontidae: Lampetra camtschatica

'"	

		
			

Osmeridae: Spirinchus thaleichthys

Speckled sanddab – Adult, 123 mm TL

Paralichthyidae: Citharichthys stigmaeus

Crescent gunnel – Adult, 160 mm TL

Pholidae: Pholis laeta


'		
			

Osmeridae: 




	
		
	[		\

Paralichthyidae: Citharichthys sordidus

'|	

		
			

Osmeridae: Osmerus mordax

Penpoint gunnel – Adult, 293 mm TL

Pholidae: 



169
Longhead dab – Juvenile, 88 mm TL

Pleuronectidae: Limanda proboscidea



'|	
'
		
			\

Pleuronectidae: Limanda aspera

Dover sole – Juvenile, 107 mm TL

Pleuronectidae: 




!!_	'!		
			\

Pleuronectidae: Platichthys stellatus

English sole – Juvenile, ~75 mm TL

Pleuronectidae: Parophrys vetulus

Rock sole – Juvenile, 137 mm TL

Pleuronectidae: Lepidopsetta spp.

!	'!		
			\

Pleuronectidae: Pleuronectes glacialis

Butter sole – Juvenile, 180 mm TL

Pleuronectidae: Isopsetta isolepis

170
Sand sole – Juvenile, 38 mm TL

Pleuronectidae: Psettichthys melanostictus

Alaska plaice – Juvenile, 120 mm TL

Pleuronectidae: Pleuronectes quadrituberculatus

C-O sole – Juvenile, 190 mm TL

Pleuronectidae: Pleuronichthys coenosus


	
'		
	]&		

Salmonidae: Coregonus sardinella

!	
'		
	&		

Salmonidae: Coregonus autumnalis

Soft sculpin – Juvenile, 51 mm TL

Psychrolutidae: Psychrolutes sigalutes

Tadpole sculpin – Juvenile, 51 mm TL

Psychrolutidae: Psychrolutes paradoxus

171
'

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	&		

Salmonidae: Salvelinus malma


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	@&		

Salmonidae: Oncorhynchus mykiss

'{_	

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	]&		

Salmonidae: Oncorhynchus nerka

{	

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	&		

Salmonidae: Oncorhynchus gorbuscha

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	[&		

Salmonidae: Oncorhynchus clarkii

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Salmonidae: Oncorhynchus kisutch

	

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	&&		

Salmonidae: Oncorhynchus keta

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Salmonidae: Oncorhynchus tshawytscha

172


'|
	!'{
		
	[#		\

Scorpaenidae: 




{	!'{
		
	@&		\

Scorpaenidae: Sebastes maliger

!{	!'{
		
	[		\

Scorpaenidae: Sebastes ciliatus

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	@]		\

Scorpaenidae: Sebastes caurinus

	!'{
		
	&		\

Scorpaenidae: Sebastes nebulosus

!'|	!'{
		
	]		\

Scorpaenidae: Sebastes auriculatus

Bocaccio – Juvenile, ~165 mm TL

Scorpaenidae: Sebastes paucispinis


{	!'{
		
			\

Scorpaenidae: Sebastes melanops

173
Snake prickleback – Juvenile, 73 mm TL

Stichaeidae: Lumpenus sagitta

Slender eelblenny – Adult, 232 mm TL

Stichaeidae: Lumpenus fabricii

Slender cockscomb – Adult, 100 mm TL

Stichaeidae: Anoplarchus insignis

Black prickleback – Adult, 170 mm TL

Stichaeidae: Xiphister atropurpureus

Blackline prickleback – Juvenile, 170 mm TL

Stichaeidae: Acantholumpenus mackayi

_	??
		
	@#		\

Syngnathidae: Syngnathus leptorhynchus

Arctic shanny – Adult, 97 mm TL

Stichaeidae: Stichaeus punctatus

'!
	
{
_		
	&		\

Stichaeidae: Eumesogrammus praecisus

174
!'|
		
	]		\

Zaproridae: Zaprora silenus

	

		
	#		

Trichodontidae: Trichodon trichodon

175
ACKNOWLEDGMENTS 

Over the years, many individuals working in the field and in the laboratory have 

helped make this atlas possible. There are too many to thank individually. We are grateful to 

them all, whether they helped pull a seine, operate a boat, process fish, preserve specimens, 

or clean up gear. 

We would especially like to recognize Jim Noel for his programming expertise and 

assistance in automating the production of the species pages. Special thanks go to the 

following people for photos: Keith Clements of Emerald Diving for the showy snailfish, 

Robert Stone for the searcher, Frank Thrower for the cutthroat trout, and John Yasaki for the 

kelp clingfish. Andres Lopez, Adam Moles, and Stan Rice made valuable suggestions on 

earlier drafts of the manuscript. Funding for much of our field work and the production of 

this synthesis was provided by NOAA’s Essential Fish Habitat Program. Other funding or inkind services were provided by the NMFS’s Alaska Regional Office, the City of Craig 

(Prince of Wales Island), North Slope Borough (Barrow), North Pacific Research Board, and 

the U.S. Army Corps of Engineers (Anchorage). 

176
CITATIONS

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program. Retrieved August 2012 from 

http://alaskacoast.state.ak.us/Explore/Tourintro.html. 

Allen, L. G. 1982. Seasonal abundance, composition, and productivity of the littoral fish 

assemblage in upper Newport Bay, California. Fish. Bull., U.S. 80:769-789. 

Allen, L. G., and M. H. Horn. 1975. Abundance, diversity and seasonality of fishes in 

Colorado Lagoon, Alamitos Bay, California. Estuar. Coast. Mar. Sci. 3:371-380. 

Beaudreau, A. H., and T. E. Essington. 2007. Spatial, temporal, and ontogenetic patterns of 

predation on rockfishes by lingcod. Trans. Am. Fish. Soc. 136:1438-1452. 

Beck, M. W., K. L. Heck Jr., K. W. Able, D. L. Childers, D. B. Eggleston, B. M. Gillanders, 

B. S. Halpern, C. G. Hays, K. Hoshino, T. J. Minello, R. J. Orth, P. F. Sheridan, and 

M. P. Weinstein. 2003. The role of nearshore ecosystems in fish and shellfish 

nurseries. Issues Ecol. 11:1-12. 

Bloomfield, A. L., and B. M. Gillanders. 2005. Fish and invertebrate assemblages in 

seagrass, mangrove, saltmarsh, and nonvegetated habitats. Estuaries 28:63-77. 

Cardillo, M. 2002. Body size and latitudinal gradients in regional diversity of new world 

birds. Global Ecol. Biogeogr. 11:59-65. 

Carls, M. G., S. W. Johnson, M. R. Lindeberg, A. D. Neff, P. M. Harris, and R. Waples. 

2008. Status review of Lynn Canal herring (Clupea pallasii). Retrieved August 2012 

from http://alaskafisheries.noaa.gov/protectedresources/herring/statusrev0308.pdf. 

Clarke, K. R., and R. M. Warick. 2001. Changes in marine communities: an approach to 

statistical analysis and interpretation, 2nd edition. PRIMER-E: Plymouth, UK. 

Cooney, T. 2007. Pacific herring, p. 81-85. In R. B. Spies (editor), Long-term ecological 

change in the northern Gulf of Alaska. Elsevier Publications. 

Csepp, D. J., and B. L. Wing. 1999. Northern range extension and habitat observations for 

blackeye goby Rhinogobiops nicholsii and kelp perch Brachyistius frenatus in 

southeastern Alaska. Alaska Fish. Res. Bull. 6:78-84. 

177
Dean, T. A., L. Haldorson, D. R. Laur, S. C. Jewett, and A. Blanchard. 2000. The distribution 

of nearshore fishes in kelp and eelgrass communities in Prince William Sound, 

Alaska: associations with vegetation and physical habitat characteristics. Environ. 

Biol. Fish. 57:271-287. 

Field, L. J. 1988. Pacific sand lance, Ammodytes hexapterus, with notes on related 

Ammodytes species, p. 15–33. In N. J. Wilimovsky, L. S. Incze, S. J. Westrheim 

(editors), Species synopses, life histories of selected fish and shellfish of the 

Northeast Pacific and Bering Sea. Washington Sea Grant Program, Seattle, WA. 

FishBase. 2012. A global information system on fishes. Retrieved August 2012 from 

http://www.fishbase.org/search.php. 

Franco, A., P. Franzoi, S. Malavasi, F. Riccato, and P. Torricelli. 2006. Fish assemblages in 

different shallow water habitats of the Venice Lagoon. Hydrobiologia 555:159-174. 

George, C., L. Moulton, and M. Johnson. 2007. A field guide to the common fishes of the 

North Slope of Alaska. North Slope Borough, Dept. Wildlife Management, Barrow, 

AK. 

Golet, G. H., K. J. Kuletz, D. D. Roby, and D. B. Irons. 2000. Adult prey choice affects chick 

growth and reproductive success in pigeon guillemots. Auk 117:82-91. 

Green, E. P., and F. T. Short. 2003. World atlas of seagrasses. Univ. California Press, 

Berkeley, CA. 

Hamilton, J., and B. Konar. 2007. Implications of substrate complexity and kelp variability 

for south-central Alaskan nearshore fish communities. Fish. Bull., U.S. 105:189-196. 

Harris, P. M., S. W. Johnson, L. G. Holland, A. D. Neff, J. F. Thedinga, and S. D. Rice. 

2005. Hydrocarbons and fisheries habitat in Berners Bay, Alaska: Baseline 

monitoring associated with the Kensington Gold Mine. AFSC Proc. Rep. 2005-06, 44 

p. Alaska Fish. Sci. Cent., Natl. Mar. Fish. Serv., NOAA, Auke Bay Laboratories, 

17109 Pt. Lena Loop Rd., Juneau, AK, 99801-8626. 

Harris, P. M., A. D. Neff, S.W. Johnson, and J. F. Thedinga. 2008. Eelgrass habitat and 

faunal assemblages in the City and Borough of Juneau, Alaska. U.S. Dep. Commer., 

NOAA Tech. Memo. NMFS-AFSC-182, 46 p. 

178
Heck, K. L. Jr., and R. J. Orth. 1980. Seagrass habitats: the roles of habitat complexity, 

competition and predation in structuring associated fish and motile macroinvertebrate 

assemblages. p. 449-464. In V. S. Kennedy (editor), Estuarine perspectives. 

Academic Press. 

Horn, M. H. 1980. Diel and seasonal variation in abundance and diversity of shallow-water 

fish populations in Morro Bay, California. Fish. Bull., U.S. 78:759-770. 

Jewett, S. C., T. A. Dean, B. R. Woodin, M. K., Hoberg, and J. J. Stegeman. 2002. Exposure 

to hydrocarbons 10 years after the Exxon Valdez oil spill: evidence from cytochrome 

P4501A expression and biliary FACs in nearshore demersal fishes. Mar. Environ. 

Res. 54:21-48. 

Johnson, S. W., M. L. Murphy, D. J. Csepp, P. M. Harris, and J. F. Thedinga. 2003a. A 

survey of fish assemblages in eelgrass and kelp habitats of southeastern Alaska. U.S. 

Dep. Commer., NOAA Tech. Memo. NMFS-AFSC-139, 39 p. 

Johnson, S. W., M. L. Murphy, and D. J. Csepp. 2003b. Distribution, habitat, and behavior of 

rockfishes, Sebastes spp., in nearshore waters of southeastern Alaska: observations 

from a remotely operated vehicle. Environ. Biol. Fish. 66:259-270. 

Johnson, S. W., A. D. Neff, and J. F. Thedinga. 2005. An atlas on the distribution and habitat 

of common fishes in shallow nearshore waters of southeastern Alaska. U.S. Dep. 

Commer., NOAA Tech. Memo. NMFS-AFSC-157, 89 p. 

Johnson, S. W., and J. F. Thedinga. 2005. Fish use and size of eelgrass meadows in 

southeastern Alaska: a baseline for long-term assessment of biotic change. Northwest 

Sci. 79:141-155. 

Johnson, S. W., J. F. Thedinga, and K. M. Munk. 2008. Distribution and use of shallowwater habitats by Pacific sand lances in southeastern Alaska. Trans. Am. Fish. Soc. 

137:1455-1463. 

Johnson, S. W., J. F. Thedinga, and A. D. Neff. 2009. Invasion by saffron cod Eleginus 

gracilis into nearshore habitats of Prince William Sound, Alaska, USA. Mar. Ecol. 

Prog. Ser. 389:203-212. 

Johnson, S. W., J. F. Thedinga, A. D. Neff, P. M. Harris, M. R. Lindeberg, J. M. Maselko, 

and S. D. Rice. 2010a. Fish assemblages in nearshore habitats of Prince William 

Sound, Alaska. Northwest Sci. 84:266-280. 

179
Johnson, S. W., J. F. Thedinga, A. D. Neff, and C. A. Hoffman. 2010b. Fish fauna in 

nearshore waters of a barrier island in the western Beaufort Sea. U.S. Dep. Commer., 

NOAA Tech. Memo. NMFS-AFSC-210, 28 p. 

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flatfishes: families Bothidae, Cynoglossidae, and Pleuronectidae. Alaska Sea Grant 

College Program, University of Alaska Fairbanks, Mar. Advisory Bull. 47. 

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184
Table 1. -- Number of locations, sample sites, beach seine hauls, and total catch of nearshore, 

marine fishes by region in Alaska from 1998 to 2011. See Figure 1 for regions 

and sample sites. 

Region Locations Sites Hauls Catch 

Arctic 2 31 81 20,803 

Aleutian Islands 6 70 70 84,118 

Bristol Bay 1 8 8 6,775 

Southcentral 5 22 23 2,858 

Prince William Sound 13 50 133 61,423 

Southeast - Northern Outside 17 127 164 89,687 

Southeast - Northern Inside 27 153 382 399,648 

Southeast - Southern Outside 12 60 114 39,772 

Southeast - Southern Inside 10 34 34 13,261 

Total 93 555 1,009 718,345 

185
Table 2. -- Total catch and percent frequency of occurrence (FO) category by fish taxa for all regions of Alaska combined; FO also 

shown for each individual region. Fish were captured with a beach seine from 1998 to 2011. Categorical FO: A = abundant ( 50% of seine hauls), C = common (25-49%), O = occasional (10-24%), U = uncommon (5-9%), R = rare (< 5%). Blanks indicate a species was not captured. Regions: AR = Arctic, BB = Bristol Bay, SC = Southcentral, PWS = Prince William Sound, SENO and SENI = Southeast northern outside and inside, SESO and SESI = Southeast southern outside and inside. *Major or potential (Arctic) target species in a fisheries management plan. aMyoxocephalus spp. = 

frog, great, or shorthorn sculpin. bRock sole = northern or southern rock sole. See Figure 1 for regions. The Appendix 

lists spatial and temporal information on fish catch by region and habitat type. 

Region 

Common name Scientific name Catch Alaska AR AI BB SC PWS SENO SENI SESO SESIWalleye pollock* Theragra chalcogramma 124,524 O O O U O R U Pacific sand lance Ammodytes hexapterus 121,089 O C F O U O O O O 

Pacific herring Clupea pallasii 106,451 O F O C O O O U 

Pink salmon* Oncorhynchus gorbuscha 89,556 C R F C C C C C C O Chum salmon* Oncorhynchus keta 76,385 C R O O O C C C C Shiner perch Cymatogaster aggregata 29,000 O C R F F 

Juvenile cod Gadidae 22,438 O C C O R U U U Pacific sandfish Trichodon trichodon 21,943 U C R R U R Capelin Mallotus villosus 21,770 U F U R Saffron cod* Eleginus gracilis 17,492 U O F C Crescent gunnel Pholis laeta 12,222 F R C F F F F F Pacific cod* Gadus macrocephalus 8,522 O R F O O O O O Threespine stickleback Gasterosteus aculeatus 8,179 O R R F O O O O F C Bay pipefish Syngnathus leptorhynchus 8,054 C F C O F C Tubesnout Aulorhynchus flavidus 7,614 O R C O O O C Snake prickleback Lumpenus sagitta 5,885 O U O O O O C C Rainbow smelt Osmerus mordax 5,117 R F Juvenile sculpin Cottidae 4,705 C F F O O O O C O O Coho salmon* Oncorhynchus kisutch 3,100 O U C U O C C C 

186
Table 2. -- (Cont.). 

Region 

Common name Scientific name Catch Alaska AR AI BB SC PWS SENO SENI SESO SESITubenose poacher Pallasina barbata 2,644 O C R O O R O Silverspotted sculpin Blepsias cirrhosus 1,867 C C O U C C O O Fish larvae Division Teleostei 1,692 U O O U R R R Juvenile greenling Hexagrammidae 1,573 C O C O F O C O Myoxocephalus spp.a Myoxocephalus spp. 1,390 C R C C O O C C C 

Pond smelt Hypomesus olidus 1,331 R F 

Northern sculpin Icelinus borealis 1,265 C U C C F F Pacific staghorn sculpin Leptocottus armatus 1,157 O R U O C C C Surf smelt Hypomesus pretiosus 932 R R R R R O 

Copper rockfish Sebastes caurinus 886 O O C F O 

Rock sole*b Lepidopsetta spp. 801 O C C R R C U O 

Whitespotted greenling Hexagrammos stelleri 719 O C C O O O O 

Juvenile rockfish Scorpaenidae 667 U O U O R O Dolly Varden Salvelinus malma 529 O O O R U O O O 

Juvenile flatfish Pleuronectidae 502 U R C U R R U R Padded sculpin Artedius fenestralis 466 O R C U U U O 

Arctic cod* Boreogadus saida 439 R O 

Masked greenling Hexagrammos octogrammus 360 U O C C O R R 

Buffalo sculpin Enophrys bison 354 O R R U O O C 

Lingcod Ophiodon elongatus 336 U C U R R 

Kelp greenling Hexagrammos decagrammus 303 U R C R O U 

Arctic shanny Stichaeus punctatus 287 O R C O U 

Starry flounder Platichthys stellatus 261 U O F R R R O R U 

Sturgeon poacher Podothecus accipenserinus 228 R R R U R 

Red Irish lord Hemilepidotus hemilepidotus 215 U U R O R C U 

187
Table 2. -- (Cont.). 

Region 

Common name Scientific name Catch Alaska AR AI BB SC PWS SENO SENI SESO SESIArmorhead sculpin Gymnocanthus galeatus 202 R R U Black rockfish Sebastes melanops 191 R U O R 

English sole Parophrys vetulus 186 R R R U R O R 

Juvenile prickleback Stichaeidae 174 R C R 

Tidepool sculpin Oligocottus maculosus 172 U C R U R O 

Manacled sculpin Synchirus gilli 167 R R O U R R 

Rock greenling Hexagrammos lagocephalus 164 U C O O R R 

Quillback rockfish Sebastes maliger 148 R R O R R 

Kelp perch Brachyistius frenatus 125 R R C U 

Blackeye goby Rhinogobiops nicholsii 108 R U O U 

Yellowfin sole* Limanda aspera 108 R U R R U R 

Pacific sanddab Citharichthys sordidus 83 R U R U O 

Speckled sanddab Citharichthys stigmaeus 82 R R R R R O R 

Juvenile smelt Osmeridae 79 R R R R R Juvenile snailfish Liparidae 67 R O U U R R Penpoint gunnel Apodichthys flavidus 65 R R U R R Dark rockfish Sebastes ciliatus 62 R R U R R 

Slender eelblenny Lumpenus fabricii 55 R O R Brown rockfish Sebastes auriculatus 54 R O 

Juvenile poacher Agonidae 49 R O U O R 

Painted greenling Oxylebius pictus 49 R R R O U 

Sockeye salmon* Oncorhynchus nerka 44 R R O O R R R R 

Arctic sculpin Myoxocephalus scorpioides 40 R O Leister sculpin Enophrys lucasi 40 R R U Least cisco Coregonus sardinella 39 R O Ninespine stickleback Pungitius pungitius 36 R U F U 

188
Table 2. -- (Cont.). 

Region 

Common name Scientific name Catch Alaska AR AI BB SC PWS SENO SENI SESO SESIBlackline prickleback Acantholumpenus mackayi 33 R F Crested sculpin Blepsias bilobus 29 R R R R Cutthroat trout Oncorhynchus clarkii 28 R R R R R 

Longfin smelt Spirinchus thaleichthys 27 R U Brown Irish lord Hemilepidotus spinosus 19 R R R R U Juvenile Irish lord Hemilepidotus spp. 19 R R R R R 

Tadpole sculpin Psychrolutes paradoxus 19 R R U Sailfin sculpin Nautichthys oculofasciatus 18 R R R R R U 

Smoothhead sculpin Artedius lateralis 17 R R R R U Bay goby Lepidogobius lepidus 16 R R U Pacific tomcod Microgadus proximus 16 R O R Scalyhead sculpin Artedius harringtoni 15 R U R R R 

Pacific spiny lumpsucker Eumicrotremus orbis 13 R R R 

Roughback sculpin Chitonotus pugetensis 13 R R Tidepool snailfish Liparis florae 13 R R R R Longhead dab Limanda proboscidea 12 R U Arctic flounder Pleuronectes glacialis 11 R R C Sand sole Psettichthys melanostictus 10 R R R Slender cockscomb Anoplarchus insignis 10 R R R R 

Steelhead trout Oncorhynchus mykiss 10 R R R R Chinook salmon* Oncorhynchus tshawytscha 9 R R R R R 

Cabezon Scorpaenichthys marmoratus 8 R R 

Antlered sculpin Enophrys diceraus 7 R R Showy snailfish Liparis pulchellus 7 R R 

Alaska plaice* Pleuronectes quadrituberculatus 6 R R R O R Belligerent sculpin Megalocottus platycephalus 6 R C 

189
Table 2. -- (Cont.). 

Region 

Common name Scientific name Catch Alaska AR AI BB SC PWS SENO SENI SESO SESIEulachon Thaleichthys pacificus 6 R R Fourhorn sculpin Myoxocephalus quadricornis 6 R U 

Juvenile gunnel Pholidae 6 R R R R R 

Juvenile lumpsucker Cyclopteridae 6 R R R 

Northern ronquil Ronquilus jordani 6 R R R 

Unidentified cisco Coregonus spp. 6 R U Arctic staghorn sculpin Gymnocanthus tricuspis 5 R U 

C-O sole Pleuronichthys coenosus 5 R R R 

Fluffy sculpin Oligocottus snyderi 5 R R R 

Plain sculpin Myoxocephalus jaok 5 R R C R 

Alaskan ronquil Bathymaster caeruleofasciatus 4 R R U 

Black prickleback Xiphister atropurpureus 4 R R R 

Juvenile ronquil Bathymasteridae 4 R R Juvenile salmon* Oncorhynchus spp. 4 R O R R 

Northern spearnose poacher Agonopsis vulsa 4 R R R 

Arctic cisco Coregonus autumnalis 3 R R 

Spotted snailfish Liparis callyodon 3 R U 

Yellowtail rockfish Sebastes flavidus 3 R R 

Arctic lamprey Lampetra camtschatica 2 R C 

Brightbelly sculpin Microcottus sellaris 2 R R 

Butter sole Isopsetta isolepis 2 R R 

Searcher Bathymaster signatus 2 R R 

Soft sculpin Psychrolutes sigalutes 2 R R 

Variegated snailfish Liparis gibbus 2 R R 

Veteran poacher Podothecus veternus 2 R R 

Atlantic poacher Leptagonus decagonus 1 R R 

190
Table 2. -- (Cont.). 

Region 

Common name Scientific name Catch Alaska AR AI BB SC PWS SENO SENI SESO SESIBering poacher Occella dodecaedron 1 R O Bocaccio Sebastes paucispinis 1 R R 

China rockfish Sebastes nebulosus 1 R R Dover sole* Microstomus pacificus 1 R R Fourline snakeblenny Eumesogrammus praecisus 1 R R 

Juvenile clingfish Gobiescocidae 1 R R Juvenile trout Oncorhynchus spp. 1 R R 

Kelp clingfish Rimicola muscarum 1 R R 

Prowfish Zaprora silenus 1 R R Pygmy poacher Odontopyxis trispinosa 1 R R Ribbon snailfish Liparis cyclopus 1 R R Sablefish* Anoplopoma fimbria 1 R R 

Saddleback sculpin Oligocottus rimensis 1 R R Smallmouth ronquil Bathymaster leurolepis 1 R R 

Wolf-eel Anarrhichthys ocellatus 1 R R 

Total 718,435 

191
Table 3. -- Mean catch-per-unit-effort (CPUE, unit = seine haul), and percent frequency of occurrence (FO) by fish taxa for all of 

Alaska (AK) and by region. Fish were captured with a beach seine from 1998 to 2011. Subscript = number of seine hauls. Blanks indicate a species was not captured. Regions: AR = Arctic, BB = Bristol Bay, SC = Southcentral, PWS = Prince William Sound, SENO and SENI = Southeast northern outside and inside, SESO and SESI = Southeast southern outside and inside. *Major or potential (Arctic) target species in a fisheries management plan. aMyoxocephalus spp. = frog, great, 

or shorthorn sculpin. bRock sole = northern or southern rock sole. See Figure 1 for regions. The Appendix lists spatial and 

temporal information on fish catch by region and habitat type. 

Region 

AK1,009 AR81 AI70 BB8 SC23 PWS133 SENO164 SENI382 SESO114 SESI34

Common name CPUE FO CPUE FO CPUE FO CPUE FO CPUE FO CPUE FO CPUE FO CPUE FO CPUE FO CPUE FOWalleye pollock* 123.4 11 13.4 13 3.2 12 1.3 5 323.0 21 0.5 3 4.1 9Pacific sand lance 120.0 23 23.5 48 506.4 60 8.8 13 1.2 8 42.0 20 186.4 18 45.4 23 2.8 18Pacific herring 105.5 19 21.9 88 34.2 17 186.3 35 125.0 23 142.2 19 7.7 19 147.9 9Pink salmon* 88.8 31 <0.1 1 182.4 51 0.5 25 21.2 39 53.3 32 107.1 33 101.5 30 111.8 40 3.6 24Chum salmon* 75.7 27 1.4 3 1.1 13 0.2 17 1.3 20 78.8 26 160.5 34 11.6 48 16.2 26Shiner perch 28.7 18 121.0 45 2.1 1 44.9 68 94.4 71Juvenile cod 22.2 11 12.1 40 299.1 47 0.2 17 <0.1 <1 0.2 6 1.2 5 0.2 9

Pacific sandfish 21.7 6 149.7 34 <0.1 4 0.3 1 29.9 9 <0.1 3

Capelin 21.6 6 177.6 51 50.2 7 1.9 3

Saffron cod* 17.3 9 6.1 11 9.8 61 126.1 49Crescent gunnel 12.1 55 0.1 4 2.2 39 7.3 77 7.2 51 16.9 58 24.9 93 22.0 82Pacific cod* 8.4 17 0.3 1 2.1 52 7.8 21 4.8 18 17.1 24 0.7 10 0.2 12

Threespine stickleback 8.1 22 0.1 2 <0.1 3 0.5 50 2.6 13 1.2 24 0.5 12 9.3 22 30.8 54 23.9 41Bay pipefish 8.0 27 5.5 52 8.4 41 1.3 10 36.2 69 38.6 41

Tubesnout 7.5 21 0.1 4 7.4 44 4.2 21 14.8 22 1.5 18 4.1 41Snake prickleback 5.8 20 33.9 6 0.7 13 1.4 20 1.0 13 5.0 22 6.5 46 14.4 41Rainbow smelt 5.1 <1 639.6 100Juvenile sculpin 4.7 27 25.6 64 10.9 50 0.1 13 13.3 13 0.6 24 0.7 20 3.4 26 0.7 14 0.2 12Coho salmon* 3.1 18 0.9 6 1.3 26 0.1 8 0.7 17 6.7 25 2.3 26 1.6 26

192
Table 3. -- (Cont.). 

Region 

AK1,009 AR81 AI70 BB8 SC23 PWS133 SENO164 SENI382 SESO114 SESI34

Common name CPUE FO CPUE FO CPUE FO CPUE FO CPUE FO CPUE FO CPUE FO CPUE FO CPUE FO CPUE FOTubenose poacher 2.6 12 0.8 38 <0.1 4 7.5 10 3.5 23 0.5 4 0.5 15Silverspotted sculpin 1.9 25 2.7 33 0.4 17 0.1 5 1.9 35 3.4 38 0.3 10 0.5 18Fish larvae 1.7 5 1.0 23 3.0 17 1.4 9 <0.1 2 6.1 2 0.9 3

Juvenile greenling 1.6 25 0.3 13 2.7 39 1.3 17 4.8 50 1.0 22 1.4 36 0.3 18

Myoxocephalus spp.a 1.4 31 0.1 4 0.8 29 0.9 39 0.3 19 0.4 18 2.4 42 1.8 43 2.2 47

Pond smelt 1.3 <1 166.4 100

Northern sculpin 1.3 27 0.2 8 1.8 29 1.6 35 2.2 54 2.9 53Pacific staghorn sculpin 1.1 21 <0.1 4 0.1 6 0.5 20 2.0 28 1.9 44 2.6 44Surf smelt 0.9 2 <0.1 4 <0.1 <1 4.1 3 0.7 2 0.4 18

Copper rockfish 0.9 12 0.2 11 2.3 27 4.1 52 0.4 15Rock sole*b 0.8 15 2.6 31 0.7 26 <0.1 <1 0.1 2 1.4 27 0.2 8 0.7 12

Whitespotted greenling 0.7 17 1.4 39 0.8 30 1.1 20 0.8 15 0.6 23 0.5 18Juvenile rockfish 0.7 8 0.6 17 0.1 5 2.7 20 0.1 2 1.3 20Dolly Varden 0.5 10 1.4 20 0.2 13 0.1 4 0.3 6 0.8 15 0.3 10 0.4 15

Juvenile flatfish 0.5 6 <0.1 1 2.4 29 0.4 9 0.1 2 0.1 2 0.5 8 1.0 3

Padded sculpin 0.5 11 <0.1 4 0.8 37 0.7 9 0.5 9 0.2 7 0. 6 15Arctic cod* 0.4 1 5.4 17Masked greenling 0.4 9 0.2 10 1.4 35 1.0 27 0.8 16 0.1 2 0.1 3Buffalo sculpin 0.4 13 0.1 4 <0.1 3 0.1 7 0.7 20 0.5 22 0.7 32Lingcod 0.3 7 0.8 31 1.1 9 0.1 4 <0.1 4Kelp greenling 0.3 9 <0.1 3 1.3 32 0.1 3 0.6 22 0.1 6Arctic shanny 0.3 10 0.1 4 1.3 39 0.3 11 0.2 6Starry flounder 0.3 7 0.3 14 5.4 63 <0.1 4 0.1 3 <0.1 1 0.5 12 <0.1 2 0.1 9

Sturgeon poacher 0.2 3 0.7 4 0.8 4 0.1 5 0.1 <1Red Irish lord 0.2 8 0.1 7 0.1 2 0.3 16 0.1 4 1.1 30 0.1 6

193
Table 3. -- (Cont.). 

Region 

AK1,009 AR81 AI70 BB8 SC23 PWS133 SENO164 SENI382 SESO114 SESI34

Common name CPUE FO CPUE FO CPUE FO CPUE FO CPUE FO CPUE FO CPUE FO CPUE FO CPUE FO CPUE FOArmorhead sculpin 0.2 3 <0.1 1 0.5 7Black rockfish 0.2 4 0.1 6 1.1 20 0.1 2English sole 0.2 3 <0.1 1 0.1 4 0.1 5 <0.1 2 1.2 10 0.6 3

Juvenile prickleback 0.2 2 2.1 27 <0.1 <1Tidepool sculpin 0.2 7 0.4 26 0.1 4 0.2 6 0.2 4 0.4 12Manacled sculpin 0.2 3 0.3 3 0.7 12 0.3 6 <0.1 <1 <0.1 3

Rock greenling 0.2 5 1.2 33 0.2 13 0.5 13 <0.1 <1 <0.1 <1

Quillback rockfish 0.1 2 <0.1 2 0.8 10 <0.1 <1 <0.1 2Kelp perch 0.1 3 <0.1 <1 1.0 28 0.2 6Blackeye goby 0.1 2 0.3 7 0.5 10 0.1 6Yellowfin sole* 0.1 3 0.2 9 <0.1 4 <0.1 <1 0.2 7 <0.1 3Pacific sanddab 0.1 3 0.3 7 <0.1 <1 0.2 9 0.2 18Speckled sanddab 0.1 3 <0.1 4 <0.1 2 0.1 4 0.1 1 0.3 11 0.2 3Juvenile smelt 0.1 1 0.1 4 0.1 2 0.4 4 <0.1 2Juvenile snailfish 0.1 3 0.3 15 0.3 9 0.5 9 <0.1 1 <0.1 <1Penpoint gunnel 0.1 3 0.1 2 0.2 8 <0.1 2 0.1 4Dark rockfish 0.1 2 <0.1 1 0.2 6 0.1 2 <0.1 <1Slender eelblenny 0.1 <1 0.7 11 <0.1 <1Brown rockfish 0.1 1 0.5 10

Juvenile poacher <0.1 2 0.5 15 0.1 6 0.1 13 <0.1 4Painted greenling <0.1 2 <0.1 2 <0.1 2 0.3 12 0.1 6Sockeye salmon* <0.1 2 <0.1 1 0.1 13 0.2 13 0.1 2 0.1 4 <0.1 1 <0.1 2Arctic sculpin <0.1 1 0.5 16Leister sculpin <0.1 2 <0.1 2 0.1 5Least cisco <0.1 <1 0.5 10

194
Table 3. -- (Cont.). 

Region 

AK1,009 AR81 AI70 BB8 SC23 PWS133 SENO164 SENI382 SESO114 SESI34

Common name CPUE FO CPUE FO CPUE FO CPUE FO CPUE FO CPUE FO CPUE FO CPUE FO CPUE FO CPUE FONinespine stickleback <0.1 1 0.1 9 2.9 75 0.1 9Blackline prickleback <0.1 <1 4.1 50Crested sculpin <0.1 2 <0.1 4 0.1 2 0.1 4Cutthroat trout <0.1 1 <0.1 <1 <0.1 3 0.1 <1 0.1 3

Longfin smelt <0.1 <1 1.2 9Brown Irish lord <0.1 <1 <0.1 2 <0.1 1 0.1 3 0.1 6Juvenile Irish lord <0.1 <1 <0.1 <1 0.1 4 <0.1 <1 <0.1 <1Tadpole sculpin <0.1 <1 <0.1 <1 0.5 6

Sailfin sculpin <0.1 2 <0.1 4 <0.1 2 <0.1 <1 <0.1 1 0.1 7Smoothhead sculpin <0.1 <1 <0.1 <1 <0.1 <1 0.1 4 0.2 6Bay goby <0.1 <1 <0.1 <1 0.1 6Pacific tomcod <0.1 <1 0.6 13 <0.1 <1

Scalyhead sculpin <0.1 1 0.1 5 <0.1 2 <0.1 <1 <0.1 <1

Pacific spiny lumpsucker <0.1 <1 <0.1 <1 <0.1 2Roughback sculpin <0.1 <1 0.1 <1

Tidepool snailfish <0.1 1 <0.1 2 <0.1 2 <0.1 3Longhead dab <0.1 <1 0.2 7Arctic flounder <0.1 <1 <0.1 1 1.3 25Sand sole <0.1 <1 0.1 4 <0.1 <1

Slender cockscomb <0.1 <1 0.1 3 <0.1 <1 <0.1 <1Steelhead trout <0.1 <1 <0.1 <1 <0.1 2 <0.1 <1Chinook salmon* <0.1 <1 <0.1 4 <0.1 <1 <0.1 <1 <0.1 2Cabezon <0.1 <1 0.2 3

Antlered sculpin <0.1 <1 <0.1 <1Showy snailfish <0.1 <1 <0.1 <1

195
Table 3. -- (Cont.). 

Region 

AK1,009 AR81 AI70 BB8 SC23 PWS133 SENO164 SENI382 SESO114 SESI34

Common name CPUE FO CPUE FO CPUE FO CPUE FO CPUE FO CPUE FO CPUE FO CPUE FO CPUE FO CPUE FOAlaska plaice* <0.1 <1 <0.1 1 <0.1 1 0.4 13 <0.1 <1Belligerent sculpin <0.1 <1 0.8 25Eulachon <0.1 <1 <0.1 <1Fourhorn sculpin <0.1 <1 0.1 6Juvenile gunnel <0.1 <1 <0.1 <1 <0.1 1 <0.1 <1 <0.1 3Juvenile lumpsucker <0.1 <1 <0.1 2 <0.1 <1Northern ronquil <0.1 <1 <0.1 2 <0.1 <1

Unidentified cisco <0.1 <1 0.1 7Arctic staghorn sculpin <0.1 <1 0.1 5C-O sole <0.1 <1 <0.1 1 <0.1 3Fluffy sculpin <0.1 <1 <0.1 1 <0.1 3Plain sculpin <0.1 <1 <0.1 1 0.4 25 <0.1 <1Alaskan ronquil <0.1 <1 <0.1 <1 0.1 6

Black prickleback <0.1 <1 <0.1 2 <0.1 <1

Juvenile ronquil <0.1 <1 <0.1 <1

Juvenile salmon* <0.1 <1 0.3 13 <0.1 4 <0.1 <1Northern spearnose poacher <0.1 <1 <0.1 <1 0.1 3

Arctic cisco <0.1 <1 <0.1 1Spotted snailfish <0.1 <1 0.1 9Yellowtail rockfish <0.1 <1 <0.1 <1Arctic lamprey <0.1 <1 0.3 25Brightbelly sculpin <0.1 <1 <0.1 <1Butter sole <0.1 <1 <0.1 <1Searcher <0.1 <1 <0.1 2Soft sculpin <0.1 <1 <0.1 <1

196
Table 3. -- (Cont.). 

Region 

AK1,009 AR81 AI70 BB8 SC23 PWS133 SENO164 SENI382 SESO114 SESI34

Common name CPUE FO CPUE FO CPUE FO CPUE FO CPUE FO CPUE FO CPUE FO CPUE FO CPUE FO CPUE FOVariegated snailfish <0.1 <1 <0.1 2Veteran poacher <0.1 <1 <0.1 1Atlantic poacher <0.1 <1 <0.1 1Bering poacher <0.1 <1 0.1 13Bocaccio <0.1 <1 <0.1 <1China rockfish <0.1 <1 <0.1 <1

Dover sole* <0.1 <1 <0.1 <1Fourline snakeblenny <0.1 <1 <0.1 1Juvenile clingfish <0.1 <1 <0.1 <1

Juvenile trout <0.1 <1 <0.1 <1

Kelp clingfish <0.1 <1 <0.1 <1Prowfish <0.1 <1 <0.1 4Pygmy poacher <0.1 <1 <0.1 <1Ribbon snailfish <0.1 <1 <0.1 <1Sablefish* <0.1 <1 <0.1 <1

Saddleback sculpin <0.1 <1 <0.1 <1Smallmouth ronquil <0.1 <1 <0.1 <1Wolf-eel <0.1 <1 <0.1 <1

Catch 718,345 20,803 84,118 6,775 2,858 61,423 89,687 399,648 39,772 13,261

Number of species 121 23 27 17 42 54 62 67 62 47

Mean CPUE 712 257 1,202 847 124 462 547 1,046 349 390

197
Table 4. -- Mean catch-per-unit-effort (CPUE, unit = seine haul), and percent frequency of 

occurrence (FO) by fish taxa and habitat type for all of Alaska. Fish were 

captured with a beach seine from 1998 to 2011. Subscript = number of seine 

hauls. Blanks indicate a species was not captured. *Major or potential (Arctic) 

target species in a fisheries management plan. aMyoxocephalus spp. = frog, great, 

or shorthorn sculpin. b

Rock sole = northern or southern rock sole. The Appendix 

lists spatial and temporal information on fish catch by region in Alaska and 

habitat type. 

Habitat type 

Bedrock195 Eelgrass295 Kelp274 Sand245

Common name CPUE FO CPUE FO CPUE FO CPUE FO 

Walleye pollock* 160.9 9 7.4 15 313.3 16 20.9 2 

Pacific sand lance 75.3 13 186.9 16 30.2 21 175.4 40 

Pacific herring 180.2 19 113.4 19 93.8 23 49.6 15 

Pink salmon* 145.8 30 66.6 34 75.6 34 84.8 24 

Chum salmon* 66.4 17 190.2 40 21.0 29 6.4 16 

Shiner perch 4.8 10 84.4 34 8.8 17 3.1 4 

Juvenile cod 2.1 6 1.3 9 74.5 10 5.1 18 

Pacific sandfish 58.4 9 <0.1 1 37.1 9 1.6 6 

Capelin 1.2 3 <0.1 <1 24.6 4 60.4 18 

Saffron cod* 0.3 3 55.7 13 1.6 10 2.3 7 

Crescent gunnel 0.7 26 34.3 94 5.0 59 2.4 26 

Pacific cod* 9.0 13 10.3 24 7.2 24 7.2 5 

Threespine stickleback 1.1 4 24.5 53 0.3 13 2.6 9 

Bay pipefish 0.2 6 25.6 66 1.0 18 0.7 6 

Tubesnout 0.5 10 23.7 53 1.7 12 0.3 3 

Snake prickleback <0.1 <1 9.9 53 9.1 9 1.9 9 

Rainbow smelt 20.9 3 

Juvenile sculpin 0.2 7 3.7 29 3.4 30 10.8 38 

Coho salmon* 0.5 8 7.4 30 2.1 19 1.0 11 

Tubenose poacher 0.1 3 8.7 29 0.2 6 0.1 4 

Silverspotted sculpin 0.8 21 3.7 41 1.9 30 0.4 5 

Fish larvae 2.3 5 <0.1 1 4.1 4 0.5 11 

Juvenile greenling 0.3 12 3.5 47 1.2 24 0.6 11 

Myoxocephalus spp.a

 <0.1 4 3.2 54 0.7 27 1.0 31 

Pond smelt 5.4 3 

Northern sculpin 0.2 10 3.0 48 0.8 28 0.5 14 

Pacific staghorn sculpin <0.1 2 3.5 54 0.1 6 0.3 13 

Surf smelt 2.4 4 0.8 3 <0.1 <1 

198
Table 4. -- (Cont.). 

Habitat type 

Bedrock195 Eelgrass295 Kelp274 Sand245

Common name CPUE FO CPUE FO CPUE FO CPUE FO 

Copper rockfish 0.4 7 2.0 21 0.7 17 <0.1 1 

Rock sole*b <0.1 <1 0.7 17 0.4 9 2.0 30 

Whitespotted greenling 0.1 7 1.7 31 0.5 21 0.2 4 

Juvenile rockfish 1.4 10 1.0 13 0.4 6 <0.1 1 

Dolly Varden 0.1 2 0.5 10 0.4 14 1.1 13 

Juvenile flatfish <0.1 1 0.6 6 0.2 3 1.1 13 

Padded sculpin 0.1 5 1.0 20 0.5 13 0.1 4 

Arctic cod* 1.8 6 

Masked greenling <0.1 3 0.9 15 0.3 12 <0.1 2 

Buffalo sculpin <0.1 2 0.6 16 0.3 15 0.3 14 

Lingcod <0.1 2 0.9 13 0.2 9 <0.1 2 

Kelp greenling 0.1 6 0.7 14 0.3 13 0.1 2 

Arctic shanny <0.1 3 0.6 12 0.3 17 0.1 4 

Starry flounder 0.3 12 <0.1 1 1.0 13 

Sturgeon poacher 0.5 5 <0.1 1 0.3 4 

Red Irish lord <0.1 1 0.5 17 0.2 9 <0.1 2 

Armorhead sculpin <0.1 1 <0.1 2 0.7 7 

Black rockfish 0.2 7 0.5 7 <0.1 2 

English sole 0.6 7 <0.1 <1 <0.1 3 

Juvenile prickleback <0.1 <1 0.7 9 

Tidepool sculpin 0.1 5 0.3 14 0.2 5 <0.1 2 

Manacled sculpin 0.1 4 0.2 3 0.2 5 <0.1 <1 

Rock greenling <0.1 4 0.2 4 0.3 10 <0.1 2 

Quillback rockfish 0.3 2 0.1 3 0.2 3 

Kelp perch 0.1 3 0.1 3 0.3 7 <0.1 <1 

Blackeye goby <0.1 3 <0.1 1 0.2 5 0.1 <1 

Yellowfin sole* 0.3 7 <0.1 1 0.1 4 

Pacific sanddab <0.1 1 0.1 6 <0.1 <1 0.1 2 

Speckled sanddab 0.2 7 <0.1 1 0.1 2 

Juvenile smelt 0.3 2 <0.1 1 <0.1 <1 0.1 2 

Juvenile snailfish <0.1 1 <0.1 1 0.1 3 0.1 5 

Penpoint gunnel <0.1 <1 0.2 7 <0.1 2 <0.1 <1 

Dark rockfish 0.1 3 0.1 2 0.1 3 <0.1 <1 

Slender eelblenny <0.1 <1 0.2 4 

Brown rockfish 0.2 2 <0.1 1 

Juvenile poacher 0.2 7 

199
Table 4. -- (Cont.). 

Habitat type 

Bedrock195 Eelgrass295 Kelp274 Sand245

Common name CPUE FO CPUE FO CPUE FO CPUE FO 

Painted greenling <0.1 2 <0.1 1 0.1 5 

Sockeye salmon* <0.1 2 0.1 2 <0.1 1 0.1 3 

Arctic sculpin 0.2 5 

Leister sculpin <0.1 1 <0.1 <1 0.1 4 <0.1 2 

Least cisco 0.2 3 

Ninespine stickleback 0.1 6 

Blackline prickleback 0.1 2 

Crested sculpin <0.1 2 <0.1 2 <0.1 3 <0.1 <1 

Cutthroat trout <0.1 <1 0.1 3 <0.1 1 <0.1 <1 

Longfin smelt 0.1 <1 

Brown Irish lord <0.1 1 <0.1 2 

Juvenile Irish lord 0.1 3 <0.1 <1 

Tadpole sculpin 0.1 <1 <0.1 <1 

Sailfin sculpin <0.1 <1 <0.1 1 <0.1 4 

Smoothhead sculpin <0.1 2 <0.1 <1 <0.1 1 

Bay goby <0.1 <1 <0.1 2 <0.1 <1 

Pacific tomcod <0.1 <1 <0.1 <1 <0.1 <1 

Pacific spiny lumpsucker <0.1 <1 <0.1 3 

Roughback sculpin <0.1 <1 

Scalyhead sculpin <0.1 1 <0.1 2 <0.1 1 

Tidepool snailfish <0.1 2 <0.1 1 <0.1 <1 

Longhead dab <0.1 2 

Arctic flounder <0.1 1 

Sand sole <0.1 1 

Slender cockscomb <0.1 <1 <0.1 1 <0.1 <1 

Steelhead trout <0.1 1 <0.1 1 <0.1 <1 

Cabezon <0.1 <1 

Chinook salmon* <0.1 <1 <0.1 <1 <0.1 <1 <0.1 <1 

Antlered sculpin <0.1 <1 <0.1 <1 

Showy snailfish <0.1 <1 

Alaska plaice* <0.1 <1 <0.1 1 

Belligerent sculpin <0.1 <1 

Eulachon <0.1 <1 

Fourhorn sculpin <0.1 2 

Juvenile gunnel <0.1 1 <0.1 <1 <0.1 <1 <0.1 <1 

Juvenile lumpsucker <0.1 <1 <0.1 <1 <0.1 <1 

200
Table 4. -- (Cont.). 

Habitat type 

Bedrock195 Eelgrass295 Kelp274 Sand245

Common name CPUE FO CPUE FO CPUE FO CPUE FO 

Northern ronquil <0.1 1 

Unidentified cisco <0.1 2 

Arctic staghorn sculpin <0.1 2 

C-O sole <0.1 1 <0.1 <1 

Fluffy sculpin <0.1 1 <0.1 <1 <0.1 <1 

Plain sculpin <0.1 2 

Alaskan ronquil <0.1 <1 <0.1 <1 

Black prickleback <0.1 <1 <0.1 <1 

Juvenile ronquil <0.1 <1 

Juvenile salmon* <0.1 <1 <0.1 <1 <0.1 <1 

Northern spearnose poacher <0.1 <1 <0.1 <1 

Arctic cisco <0.1 <1 

Spotted snailfish <0.1 <1 <0.1 <1 

Yellowtail rockfish <0.1 <1 

Arctic lamprey <0.1 <1 

Brightbelly sculpin <0.1 <1 

Butter sole <0.1 <1 

Searcher <0.1 <1 

Soft sculpin <0.1 <1 

Variegated snailfish <0.1 <1 

Veteran poacher <0.1 <1 

Atlantic poacher <0.1 <1 

Bering poacher <0.1 <1 

Bocaccio <0.1 <1 

China rockfish <0.1 <1 

Dover sole* <0.1 <1 

Fourline snakeblenny <0.1 <1 

Juvenile clingfish <0.1 <1

Juvenile trout <0.1 <1 

Kelp clingfish <0.1 <1

Prowfish <0.1 <1 

Pygmy poacher <0.1 <1 

Ribbon snailfish <0.1 <1 

Sablefish* <0.1 <1 

Saddleback sculpin <0.1 <1

201
Table 4. -- (Cont.). 

Habitat type 

Bedrock195 Eelgrass295 Kelp274 Sand245

Common name CPUE FO CPUE FO CPUE FO CPUE FO 

Smallmouth ronquil <0.1 <1 

Wolf-eel <0.1 <1

Catch 139,453 262,521 199,312 117,059 

Number of species 57 83 79 87 

Mean CPUE 715 890 727 478 

202
Table 5. -- Total catch by species, number measured (n), mean length in millimeters (fork 

length (FL) or total length (TL) depending on species), standard deviation (SD), 

range, and the proportion (%) of each species estimated to be mature. Fish were 

captured with a beach seine in Alaska from 1998 to 2011. Percent mature is based 

on estimated size at first maturity from FishBase (www.fishbase.org). *Major or 

potential (Arctic) target species in a fisheries management plan. a

Myoxocephalus

spp. = frog, great, or shorthorn sculpin. b

Rock sole = northern or southern rock 

sole. 

Length 

Common name Catch n Mean SD Range % 

Walleye pollock* 124,524 2,517 58.9 TL 24.9 25-242 0 

Pacific sand lance 121,089 3,324 90.9 FL 29.5 24-190 17 

Pacific herring 106,451 3,126 56.8 FL 29.9 24-212 0 

Pink salmon* 89,556 4,472 52.2 FL 16.0 23-123 0 

Chum salmon* 76,385 3,856 64.5 FL 16.6 31-125 0 

Shiner perch 29,000 2,662 90.6 FL 26.4 40-170 8 

Pacific sandfish 21,943 815 56.2 FL 28.4 16-150 0 

Capelin 21,770 1,305 57.6 FL 17.4 27-160 3 

Saffron cod* 17,492 2,316 118.3 TL 62.1 22-365 <1 

Crescent gunnel 12,222 275 124.3 TL 34.8 22-240 17 

Pacific cod* 8,522 2,089 71.3 TL 29.0 21-270 0 

Threespine stickleback 8,179 478 42.5 TL 21.6 11-93 37 

Bay pipefish 8,054 592 171.4 TL 50.1 53-315 29 

Tubesnout 7,614 1,151 97.5 TL 43.4 13-261 35 

Snake prickleback 5,885 522 152.8 TL 85.9 37-420 7 

Rainbow smelt 5,117 319 53.6 FL 26.8 20-270 <1 

Coho salmon* 3,100 1,349 96.3 FL 22.0 32-191 0 

Tubenose poacher 2,644 287 61.1 TL 19.3 25-155 5 

Silverspotted sculpin 1,867 226 57.8 TL 31.2 19-165 7 

Myoxocephalus spp.a

 1,390 519 121.4 TL 113.2 17-485 13 

Pond smelt 1,331 202 77.4 FL 13.1 32-111 0 

Northern sculpin 1,265 59 87.8 TL 28.9 29-179 80 

Pacific staghorn sculpin 1,157 253 193.3 TL 60.2 64-349 13 

Surf smelt 932 326 94.6 FL 32.6 39-183 0 

Copper rockfish 886 710 88.4 TL 37.3 19-235 0 

Rock sole*b

 801 539 124.7 TL 99.2 30-522 8 

Whitespotted greenling 719 591 110.3 TL 71.6 48-360 3 

Dolly Varden 529 420 224.5 FL 79.4 100-535 3 

Padded sculpin 466 429 73.3 TL 19.7 18-133 13 

 

 

203
Table 5. -- (Cont.). 

 

Length 

Common name Catch n Mean SD Range % 

Arctic cod* 439 139 40.1 FL 11.5 25-83 0 

Masked greenling 360 231 140.6 TL 78.5 51-480 6 

Buffalo sculpin 354 78 78.7 TL 49.6 13-220 0 

Lingcod 336 196 115.2 TL 54.5 27-405 0 

Kelp greenling 303 170 142.3 TL 86.8 49-500 3 

Arctic shanny 287 161 88.2 TL 22.5 25-134 48 

Starry flounder 261 188 207.8 TL 122.1 17-560 2 

Sturgeon poacher 228 20 67.0 TL 46.6 25-196 5 

Red Irish lord 215 46 120.4 TL 85.6 31-378 7 

Armorhead sculpin 202 47 82.1 TL 15.9 48-151 0 

Black rockfish 191 183 123.3 TL 35.8 42-329 0 

English sole 186 80 89.0 TL 40.4 36-212 0 

Tidepool sculpin 172 128 52.3 TL 11.7 24-96 11 

Manacled sculpin 167 67 31.4 TL 10.6 7-55 4 

Rock greenling 164 135 214.1 TL 102.5 44-495 16 

Quillback rockfish 148 118 63.5 TL 35.5 17-195 0 

Kelp perch 125 78 93.8 FL 34.2 43-181 13 

Blackeye goby 108 22 79.5 TL 15.3 45-106 9 

Yellowfin sole* 108 76 108.0 TL 37.1 60-285 1 

Pacific sanddab 83 48 80.0 TL 31.0 35-160 0 

Speckled sanddab 82 76 93.1 TL 28.4 35-163 30 

Penpoint gunnel 65 26 119.1 TL 88.5 32-300 12 

Dark rockfish 62 60 94.2 TL 33.9 58-280 0 

Slender eelblenny 55 51 49.3 TL 26.8 35-232 2 

Brown rockfish 54 53 111.4 TL 35.8 55-200 0 

Painted greenling 49 24 85.1 TL 38.9 27-211 4 

Sockeye salmon* 44 32 79.7 FL 78.9 29-498 3 

Arctic sculpin 40 40 97.7 TL 41.9 18-221 8 

Leister sculpin 40 21 131.6 TL 53.9 56-235 43 

Least cisco 39 39 288.8 FL 28.8 229-341 100 

Ninespine stickleback 36 35 46.9 TL 14.0 26-78 11 

Blackline prickleback 33 32 61.6 TL 34.5 34-165 0 

Crested sculpin 29 11 140.3 TL 27.3 88-190 27 

Cutthroat trout 28 25 207.4 FL 46.0 139-306 0 

Longfin smelt 27 27 107.0 FL 26.1 63-155 37 

 

 

204
Table 5. -- (Cont.). 

 

Length 

Common name Catch n Mean SD Range % 

Brown Irish lord 19 19 96.4 TL 45.3 35-199 5 

Tadpole sculpin 19 0 

Sailfin sculpin 18 4 96.8 TL 50.8 28-138 50 

Smoothhead sculpin 17 14 94.2 TL 19.8 72-133 50 

Bay goby 16 6 74.3 TL 19.7 52-100 67 

Pacific tomcod 16 16 109.3 TL 53.0 72-235 13 

Scalyhead sculpin 15 15 71.0 TL 22.7 42-132 27 

Pacific spiny lumpsucker 13 6 26.3 TL 13.6 10-47 0 

Roughback sculpin 13 13 85.3 TL 21.2 60-123 0 

Tidepool snailfish 13 2 32.0 TL 0.0 32 0 

Longhead dab 12 12 66.6 TL 18.9 44-91 0 

Arctic flounder 11 11 62.5 TL 16.7 46-110 0 

Sand sole 10 2 38.0 TL 5.7 34-42 0 

Slender cockscomb 10 6 94.5 TL 19.1 70-117 67 

Steelhead trout 10 10 188.3 FL 31.9 149-234 0 

Chinook salmon* 9 7 131.9 FL 33.7 77-170 0 

Cabezon 8 0 

Antlered sculpin 7 7 112.6 TL 36.9 88-169 0 

Showy snailfish 7 0 

Alaska plaice* 6 6 135.2 TL 77.8 69-269 0 

Belligerent sculpin 6 6 209.8 TL 46.9 118-247 0 

Eulachon 6 6 40.5 FL 2.1 38-44 0 

Fourhorn sculpin 6 6 122.2 TL 56.2 60-183 0 

Northern ronquil 6 6 116.2 TL 37.0 55-170 0 

Arctic staghorn sculpin 5 5 41.6 TL 12.2 30-56 0 

C-O sole 5 4 166.5 TL 76.4 98-265 25 

Fluffy sculpin 5 0 

Plain sculpin 5 5 225.0 TL 89.5 76-296 0 

Alaskan ronquil 4 0 

Black prickleback 4 1 160.0 TL 0.0 160 0 

Northern spearnose poacher 4 3 81.0 TL 31.1 54-115 0 

Arctic cisco 3 3 83.0 FL 3.6 79-86 0 

Spotted snailfish 3 3 77.0 TL 13.9 61-86 33 

Yellowtail rockfish 3 2 116.5 TL 2.1 115-118 0 

Arctic lamprey 2 2 163.0 TL 2.8 161-165 0 

 

 

205
Table 5. -- (Cont.). 

 

Length 

Common name Catch n Mean SD Range % 

Brightbelly sculpin 2 0 

Butter sole 2 0 

Searcher 2 2 92.0 TL 1.4 91-93 0 

Soft sculpin 2 2 38.5 TL 17.7 26-51 0 

Variegated snailfish 2 2 50.5 TL 27.6 31-70 0 

Veteran poacher 2 2 28.0 TL 4.2 25-31 0 

Atlantic poacher 1 1 25.0 TL 0.0 25 0 

Bering poacher 1 1 104.0 TL 0.0 104 0 

Bocaccio 1 1 245.0 TL 0.0 245 0 

China rockfish 1 1 80.0 TL 0.0 80 0 

Dover sole* 1 1 107.0 TL 0.0 107 0 

Fourline snakeblenny 1 1 29.0 TL 0.0 29 0 

Kelp clingfish 1 0 

Prowfish 1 1 106.0 TL 0.0 106 0 

Pygmy poacher 1 1 71.0 TL 0.0 71 100 

Ribbon snailfish 1 1 103.0 TL 0.0 103 100 

Sablefish* 1 1 215.0 FL 0.0 215 0 

Saddleback sculpin 1 1 47.0 TL 0.0 47 100 

Smallmouth ronquil 1 1 123.0 TL 0.0 123 0 

Wolf-eel 1 0 0 

Total 686,351 38,720 

206
Table 6. -- Range in surface water temperature and salinity (practical salinity scale, PSS) by 

region in Alaska where nearshore fishes were captured with a beach seine from 

1998 to 2011. Sampling spanned from January to September; most sampling, 

however, was from June to August. Blank indicates missing data. See Figure 1 for 

regions. 

Region Temperature (o C) Salinity (PSS)

Arctic 0.0-11.0 10-35 

Aleutian Islands 5.0-10.0 22-33 

Bristol Bay 11.0-14.5 

Southcentral 6.0-14.5 5-29 

Prince William Sound 1.0-20.0 2-33 

Southeast - Northern Outside 6.5-18.0 3-33 

Southeast - Northern Inside 2.5-16.0 4-35 

Southeast - Southern Outside 6.0-15.5 12-33 

Southeast - Southern Inside 8.0-16.0 0-33 

207
Table 7. -- Total catch, mean catch-per-unit-effort (CPUE, unit = seine haul), and percent 

frequency of occurrence (FO) by fish taxa and habitat type in the Arctic (AR) 

region of Alaska. Subscript = number of hauls. Page = species page. Fish were 

captured with a beach seine from 2004 to 2009. *Potential target species in a 

fisheries management plan. aMyoxocephalus spp. = shorthorn sculpin. See Figure 

4 for the AR region map. The Appendix lists spatial and temporal information on 

fish catch by habitat type. 

Habitat type 

Sand81

Common name Catch CPUE FO Page 

Capelin 14,383 177.6 51 109 

Juvenile sculpin 2,072 25.6 64 

Pacific sand lance 1,906 23.5 48 47 

Juvenile cod* 983 12.1 40 

Saffron cod* 492 6.1 11 84 

Arctic cod* 439 5.4 17 83 

Juvenile prickleback 173 2.1 27 

Fish larvae 84 1.0 23 

Slender eelblenny 54 0.7 11 152 

Juvenile poacher 41 0.5 15 

Arctic sculpin 40 0.5 16 73 

Least cisco 39 0.5 10 132 

Juvenile snailfish 20 0.3 15 

Yellowfin sole 14 0.2 9 120 

Longhead dab 12 0.2 7 121 

Ninespine stickleback 11 0.1 9 89 

Fourhorn sculpin 6 0.1 6 72 

Unidentified cisco 6 0.1 7 

Arctic staghorn sculpin 5 0.1 5 64 

Threespine stickleback 5 0.1 2 88 

Myoxocephalus spp.a

 4 0.1 4 74 

Arctic cisco 3 <0.1 1 131 

Variegated snailfish 2 <0.1 2 105 

Veteran poacher 2 <0.1 1 46 

Alaska plaice 1 <0.1 1 126 

Arctic flounder 1 <0.1 1 125 

Atlantic poacher 1 <0.1 1 41 

Fourline snakeblenny 1 <0.1 1 151 

Juvenile flatfish 1 <0.1 1 

208
Table 7. -- (Cont.). 

Habitat type 

Sand81

Common name Catch CPUE FO Page 

Pink salmon 1 <0.1 1 134 

Plain sculpin 1 <0.1 1 71 

Catch 20,803 20,803 

Number of species 23 23 

Mean CPUE 257 257 

209
Table 8. -- Total catch, mean catch-per-unit-effort (CPUE, unit = seine haul), and percent 

frequency of occurrence (FO) by fish taxa and habitat type in the Aleutian Islands 

(AI) region of Alaska. Subscript = number of hauls. Page = species page. Fish 

were captured with a beach seine in 2005. Blanks indicate a species was not 

captured. *Major target species in a fisheries management plan. aRock sole = 

northern or southern rock sole. b

Myoxocephalus spp. = frog, great, or shorthorn 

sculpin. See Figure 5 for the AI region map. The Appendix lists spatial and 

temporal information on fish catch by habitat type. 

Habitat type 

Bedrock10 Kelp27 Sand33

Common name Catch CPUE FO CPUE FO CPUE FO Page

Pacific sand lance 35,451 0.2 20 21.8 33 1,056.4 94 47 

Juvenile cod* 20,934 34.0 70 755.1 74 6.3 18 

Pink salmon* 12,770 34.1 30 379.2 63 66.4 48 134 

Pacific sandfish 10,478 3.4 30 373.4 48 11.0 24 157 

Snake prickleback 2,374 87.0 11 0.8 3 153 

Juvenile sculpin 761 1.4 30 12.4 81 12.5 30 

Fish larvae 208 19.7 50 0.3 15 0.1 9 

Silverspotted sculpin 186 2.5 50 6.0 67 94 

Rock solea

 179 1.0 15 4.6 55 119 

Juvenile flatfish 168 0.1 10 1.1 22 4.2 39 

Chum salmon* 100 3.2 4 0.4 3 135 

Dolly Varden 99 0.3 15 2.8 30 140 

Rock greenling 81 0.4 30 2.8 67 0.1 6 97 

Coho salmon* 64 0.3 4 1.7 9 136 

Myoxocephalus spp.b

 55 0.1 10 0.4 19 1.3 42 74 

Sturgeon poacher 51 1.6 9 45 

Juvenile greenling 24 0.9 30 <0.1 3 

Pacific cod* 23 0.9 4 85 

Starry flounder 23 <0.1 4 0.7 27 124 

Juvenile snailfish 21 0.1 10 0.7 19

Manacled sculpin 19 1.9 20 79 

Masked greenling 14 0.1 10 0.3 15 0.2 6 98 

Red Irish lord 7 0.1 10 0.2 15 65 

Black rockfish 6 0.2 15 146 

Juvenile poacher 6 0.2 12 

Crescent gunnel 4 0.2 11 117 

Sockeye salmon* 3 0.1 3 138 

Armorhead sculpin 2 0.1 4 63 

Kelp greenling 2 0.1 10 <0.1 4 96 

210
Table 8. -- (Cont.). 

Habitat type 

Bedrock10 Kelp27 Sand33

Common name Catch CPUE FO CPUE FO CPUE FO Page

Threespine stickleback 2 0.1 10 <0.1 4 88 

Alaska plaice* 1 <0.1 3 126 

Dark rockfish 1 <0.1 4 143 

English sole 1 <0.1 3 123 

Catch 84,118 983 44,485 38,650 

Number of species 27 14 22 17 

Mean CPUE 1,202 98 1,648 1,171 

211
Table 9. -- Total catch, mean catch-per-unit-effort (CPUE, unit = seine haul), and percent 

frequency of occurrence (FO) by fish taxa and habitat type in the Bristol Bay (BB) 

region of Alaska. Subscript = number of hauls. Page = species page. Fish were 

captured with a beach seine in 2009. *Major target species in a fisheries 

management plan. See Figure 6 for the BB region map. The Appendix lists spatial 

and temporal information on fish catch by habitat type. 

Habitat type

Sand8

Common name Catch CPUE FO Page 

Rainbow smelt 5,117 639.6 100 110 

Pond smelt 1,331 166.4 100 107 

Pacific herring 175 21.9 88 55 

Starry flounder 43 5.4 63 124 

Blackline prickleback 33 4.1 50 149 

Ninespine stickleback 23 2.9 75 89 

Arctic flounder 10 1.3 25 125 

Chum salmon* 9 1.1 13 135 

Belligerent sculpin 6 0.8 25 69 

Tubenose poacher 6 0.8 38 44 

Pink salmon* 4 0.5 25 134 

Threespine stickleback 4 0.5 50 88 

Alaska plaice* 3 0.4 13 126 

Plain sculpin 3 0.4 25 71 

Arctic lamprey 2 0.3 25 115 

Juvenile salmon* 2 0.3 13

Bering poacher 1 0.1 13 42 

Juvenile poacher 1 0.1 13

Juvenile sculpin 1 0.1 13

Sockeye salmon* 1 0.1 13 138 

Catch 6,775 6,775

Number of species 17 17

Mean CPUE 847 847

212
Table 10. -- Total catch, mean catch-per-unit-effort (CPUE, unit = seine haul), and percent 

frequency of occurrence (FO) by fish taxa and habitat type in the Southcentral 

(SC) region of Alaska. Subscript = number of hauls. Page = species page. Fish 

were captured with a beach seine from 2008 to 2010. Blanks indicate a species 

was not captured. *Major target species in a fisheries management plan. 

a

Myoxocephalus spp. = frog, great, or shorthorn sculpin. bRock sole = northern or 

southern rock sole. See Figure 7 for the SC region map. The Appendix lists 

spatial and temporal information on fish catch by habitat type. 

Habitat type 

Bedrock4 Eelgrass1 Kelp8 Sand10

Common name Catch CPUE FO CPUE FO CPUE FO CPUE FO Page

Pacific herring 786 0.3 25 3.0 100 78.2 20 55

Pink salmon* 488 22.8 25 47.6 38 1.6 50 134

Walleye pollock* 309 38.6 38 87

Juvenile sculpin 306 37.5 13 0.6 20

Saffron cod 225 10.0 100 25.1 100 1.4 50 84

Pacific sand lance 202 31.3 25 7.7 20 47

Juvenile greenling 63 4.8 75 5.4 63 0.1 10

Threespine stickleback 60 6.0 30 88

Crescent gunnel 51 0.5 25 3.0 100 5.8 88 117

Pacific cod* 48 1.3 50 3.0 100 4.1 75 0.7 30 85

Masked greenling 33 18.0 100 1.8 75 0.1 10 98

Whitespotted greenling 32 1.0 25 4.0 100 2.6 63 0.3 20 99

Fish larvae 31 0.8 25 2.8 10

Coho salmon* 30 5.3 25 0.3 13 0.7 40 136

Longfin smelt 27 2.7 20 111

Myoxocephalus spp.a

 20 1.0 100 0.6 50 1.4 40 74

Rock sole*b

 17 1.8 50 0.3 20 119

Snake prickleback 17 13.0 100 0.5 25 153

Pacific tomcod 14 12.0 100 0.1 13 0.1 10 86

Juvenile rockfish 13 0.3 25 1.5 38 

Juvenile snailfish 11 0.3 25 1.3 13 

Juvenile flatfish 9 0.9 20

Silverspotted sculpin 9 0.5 25 0.9 38 94

Chum salmon* 5 0.3 25 0.1 13 0.3 20 135

Dolly Varden 5 0.4 25 0.2 10 140

Juvenile cod 5 0.5 38 0.1 10

Rock greenling 4 0.4 25 0.1 10 97

Sockeye salmon* 4 0.4 30 138

Buffalo sculpin 3 0.3 10 60

Juvenile smelt 3 0.3 10

213
Table 10. -- (Cont.). 

Habitat type 

Bedrock4 Eelgrass1 Kelp8 Sand10

Common name Catch CPUE FO CPUE FO CPUE FO CPUE FO Page

Spotted snailfish 3 1.0 100 0.2 10 102

Tubesnout 3 3.0 100 50

Arctic shanny 2 0.3 13 154

English sole 2 2.0 100 123

Ninespine stickleback 2 0.2 20 89

Sand sole 2 0.2 10 128

Chinook salmon* 1 0.1 10 139

Crested sculpin 1 0.1 13 93

Juvenile poacher 1 0.1 10

Juvenile salmon* 1 0.1 13 

Pacific sandfish 1 0.3 25 157

Pacific staghorn sculpin 1 1.0 100 68

Padded sculpin 1 1.00 100 56

Prowfish 1 0.1 10 158

Sailfin sculpin 1 0.1 13 95

Speckled sanddab 1 0.1 10 114

Starry flounder 1 0.1 10 124

Surf smelt 1 1.0 100 108

Tubenose poacher 1 1.0 100 44

Yellowfin sole* 1 1.0 100 120

Catch 2,858 277 78 1,419 1,084 

Number of species 42 12 17 21 26 

Mean CPUE 124 69 78 177 108 

214
Table 11. -- Total catch, mean catch-per-unit-effort (CPUE, unit = seine haul), and percent 

frequency of occurrence (FO) by fish taxa and habitat type in the Prince William 

Sound (PWS) region of Alaska. Fish were captured with a beach seine in 1999, 

2006, 2007, 2009, and 2010. Subscript = number of hauls. Page = species page. 

Blanks indicate a species was not captured. *Major target species in a fisheries 

management plan. a

Myoxocephalus spp. = frog, great, or shorthorn sculpin. 

b

Rock sole = northern or southern rock sole. See Figure 8 for the PWS region 

map. The Appendix lists spatial and temporal information on fish catch by 

habitat type. 

Habitat type 

Bedrock31 Eelgrass50 Kelp45 Sand7

Common name Catch CPUE FO CPUE FO CPUE FO CPUE FO Page

Pacific herring 24,780 30.8 29 295.2 22 20.9 47 1160.6 71 55

Saffron cod 16,775 2.0 19 328.5 74 5.1 44 8.1 29 84

Pink salmon* 7,092 72.1 32 21.0 24 83.3 42 8.1 14 134

Capelin 6,670 0.3 10 148.0 13 109

Pacific cod* 1,041 0.4 13 19.1 32 1.6 18 85

Tubesnout 983 0.8 10 14.8 84 4.9 31 50

Crescent gunnel 967 1.0 48 15.3 94 3.6 82 1.3 57 117

Bay pipefish 731 0.3 13 12.9 88 1.8 47 156

Walleye pollock* 423 0.3 6 7.5 24 0.9 4 87

Snake prickleback 188 <0.1 3 3.6 44 0.1 2 0.4 29 153

Chum salmon* 176 0.3 19 0.8 22 2.9 20 135

Arctic shanny 169 0.2 13 2.3 44 0.9 51 1.1 43 154

Juvenile greenling 167 0.1 6 2.7 30 0.6 13 

Pacific sand lance 162 0.1 6 1.9 8 1.5 11 47

Threespine stickleback 156 <0.1 3 2.8 46 0.3 18 88

Masked greenling 133 0.1 6 1.5 40 1.2 31 98

Lingcod 111 0.1 6 1.8 46 0.4 33 0.1 14 100

Padded sculpin 101 0.2 16 0.8 40 1.0 42 1.3 71 56

Whitespotted greenling 101 0.1 10 1.3 40 0.7 36 0.1 14 99

Manacled sculpin 89 0.1 10 1.1 6 0.4 20 1.4 14 79

Juvenile sculpin 75 0.2 10 0.3 22 1.2 31 0.7 57

Tidepool sculpin 51 0.4 26 0.6 40 0.2 13 75

Myoxocephalus spp.a

 43 0.6 30 0.2 16 0.6 43 74

Northern sculpin 25 0.4 16 0.1 7 67

Copper rockfish 22 0.2 13 0.1 6 0.3 18 142

Silverspotted sculpin 18 0.3 12 <0.1 2 94

Coho salmon* 16 <0.1 3 0.2 10 0.1 9 136

215
Table 11. -- (Cont.). 

 

Habitat type 

Bedrock31 Eelgrass50 Kelp45 Sand7

Common name Catch CPUE FO CPUE FO CPUE FO CPUE FO Page

Dolly Varden 13 <0.1 4 <0.1 2 1.4 29 140

Red Irish lord 12 0.2 4 <0.1 2 65

Pacific staghorn sculpin 11 0.1 8 <0.1 2 0.6 43 68

Starry flounder 10 <0.1 2 0.2 4 0.1 14 124

Juvenile flatfish 9 1.3 29

Juvenile rockfish 9 0.1 10 <0.1 4 

Scalyhead sculpin 7 0.1 6 0.1 4 <0.1 4 57

Crested sculpin 6 0.1 6 93

Juvenile smelt 6 0.9 29

Penpoint gunnel 6 0.1 4 <0.1 2 116

Slender cockscomb 6 <0.1 2 <0.1 4 0.3 14 150

Sockeye salmon* 6 0.1 4 138

Juvenile lumpsucker 5 0.1 3 0.1 2 

Northern ronquil 5 0.1 4 54

Quillback rockfish 5 <0.1 2 0.1 4 145

Buffalo sculpin 4 0.1 7 0.1 14 60

Black prickleback 3 0.1 4 155

Brown Irish lord 3 <0.1 2 <0.1 4 66

Fish larvae 3 0.4 29

Leister sculpin 3 <0.1 2 <0.1 2 62

Painted greenling 3 0.1 4 101

Speckled sanddab 3 <0.1 4 <0.1 2 114

Brightbelly sculpin 2 0.3 14 70

Pacific tomcod 2 0.3 14 86

Sailfin sculpin 2 <0.1 4 95

Searcher 2 <0.1 4 53

Surf smelt 2 <0.1 2 108

Juvenile Irish lord 1 <0.1 2

Juvenile cod 1 <0.1 3

Juvenile gunnel 1 <0.1 3

Pacific spiny lumpsucker 1 <0.1 2 80

Plain sculpin 1 0.1 14 71

Rock sole*b

 1 0.1 14 119

Slender eelblenny 1 <0.1 2 152

Smallmouth ronquil 1 <0.1 2 52

216
Table 11. -- (Cont.). 

 

Habitat type 

Bedrock31 Eelgrass50 Kelp45 Sand7

Common name Catch CPUE FO CPUE FO CPUE FO CPUE FO Page

Smoothhead sculpin 1 <0.1 3 58

Yellowfin sole* 1 <0.1 2 120

Catch 61,423 3,410 36,936 12,747 8,330

Number of species 54 25 41 43 21

Mean CPUE 462 110 739 283 1,190

217
Table 12. -- Total catch, mean catch-per-unit-effort (CPUE, unit = seine haul), and percent 

frequency of occurrence (FO) by fish taxa and habitat type in the Southeast 

Northern Outside (SENO) region of Alaska. Subscript = number of hauls. Page = 

species page. Fish were captured with a beach seine in 1998, 1999, 2001 to 2003, 

2006, 2008, and 2010. Blanks indicate a species was not captured. *Major target 

species in a fisheries management plan. a

Myoxocephalus spp. = frog, great, or 

shorthorn sculpin. b

Rock sole = northern or southern rock sole. See Figure 9 for 

the SENO region map. The Appendix lists spatial and temporal information on 

fish catch by habitat type. 

Habitat type 

Bedrock70 Eelgrass57 Kelp26 Sand11

Common name Catch CPUE FO CPUE FO CPUE FO CPUE FO Page

Pacific herring 20,497 87.8 20 47.1 23 416.5 31 76.0 27 55

Shiner perch 19,851 9.9 24 307.9 75 38.9 38 54.2 27 82

Pink salmon* 17,563 232.7 30 6.7 44 11.3 19 54.5 27 134

Chum salmon* 12,918 120.9 17 73.1 42 7.2 15 9.6 18 135

Pacific sand lance 6,887 73.6 11 22.3 23 1.0 35 39.8 27 47

Bay pipefish 1,380 0.5 10 23.2 89 0.4 27 0.8 27 156

Tubenose poacher 1,230 21.3 19 0.6 15 0.1 9 44

Crescent gunnel 1,172 0.3 13 18.9 96 2.0 46 1.7 64 117

Fish larvae 1,006 <0.1 1 <0.1 2 38.6 8

Pacific cod* 789 0.4 3 8.1 32 11.5 35 85

Juvenile greenling 785 0.5 23 9.9 79 4.6 50 5.6 73

Tubesnout 681 0.8 14 10.0 35 2.0 19 50

Surf smelt 669 11.7 9 108

Juvenile rockfish 443 3.5 20 3.0 28 0.9 8 0.3 9

Copper rockfish 379 0.8 7 4.8 47 1.6 42 0.5 18 142

Silverspotted sculpin 308 0.2 13 4.6 63 1.0 38 0.6 18 94

Northern sculpin 297 0.1 7 4.8 60 0.4 27 0.4 18 67

Walleye pollock* 216 0.1 3 3.4 7 0.8 8 87

Kelp greenling 212 0.3 14 2.3 40 1.9 62 1.0 36 96

Lingcod 183 <0.1 1 3.0 18 0.3 8 0.1 9 100

Whitespotted greenling 180 <0.1 4 2.9 44 0.4 15 99

Black rockfish 176 0.6 19 2.3 32 0.2 8 146

Snake prickleback 161 2.8 35 0.4 18 153

Quillback rockfish 136 0.9 6 0.6 14 1.4 15 145

Masked greenling 127 2.0 35 0.5 27 98

Sturgeon poacher 125 2.2 12 45

Padded sculpin 121 <0.1 1 2.1 21 <0.1 4 56

Coho salmon* 118 0.4 6 1.4 35 0.4 15 136

Juvenile sculpin 109 0.1 7 1.4 30 0.4 23 1.0 36

218
Table 12. -- (Cont.). 

Habitat type 

Bedrock70 Eelgrass57 Kelp26 Sand11

Common name Catch CPUE FO CPUE FO CPUE FO CPUE FO Page

Threespine stickleback 89 <0.1 1 1.5 32 <0.1 4 88

Pacific staghorn sculpin 80 <0.1 1 1.2 47 0.1 8 0.6 18 68

Rock greenling 77 0.1 6 1.0 21 0.6 23 97

Myoxocephalus spp.a

 70 <0.1 1 1.0 40 0.1 4 0.6 45 74

Juvenile smelt 62 0.8 3 <0.1 2 0.2 8 0.3 9

Manacled sculpin 55 <0.1 3 0.1 5 1.8 19 79

Blackeye goby 54 0.1 7 <0.1 4 0.7 12 2.6 18 92

Dolly Varden 50 0.3 4 0.5 7 0.1 8 0.1 9 140

Pacific sanddab 48 <0.1 1 0.4 12 2.0 27 113

Red Irish lord 47 0.7 35 0.2 12 0.5 27 65

Pacific sandfish 41 1.6 8 157

Juvenile cod* 39 0.7 18

Dark rockfish 37 0.1 3 0.5 9 0.2 12 143

Penpoint gunnel 34 <0.1 1 0.6 21 116

Buffalo sculpin 21 0.3 18 0.2 18 60

Sockeye salmon* 20 0.1 4 0.2 4 0.2 4 138

Speckled sanddab 20 0.4 11 114

Juvenile Irish lord 16 0.3 9 0.1 9

English sole 16 0.1 5 0.7 45 123

Rock sole*b

 16 0.3 5 0.1 9 119

Juvenile flatfish 15 <0.1 1 0.1 4 1.0 9

Tidepool sculpin 12 0.2 9 0.1 4 0.1 9 75

Starry flounder 7 0.1 2 0.4 9 124

Bay goby 6 0.6 9 91

Painted greenling 4 <0.1 3 <0.1 2 <0.1 4 101

Scalyhead sculpin 4 0.1 5 57

Brown Irish lord 3 <0.1 2 <0.1 4 66

Chinook salmon* 3 <0.1 1 139

Cutthroat trout 3 <0.1 1 133

C-O sole 2 <0.1 4 127

Fluffy sculpin 2 <0.1 3 77

Juvenile gunnel 2 <0.1 1 <0.1 2

Kelp perch 2 0.1 4 81

Smoothhead sculpin 2 <0.1 1 58

China rockfish 1 <0.1 4 147

Dover sole* 1 <0.1 2 122

219
Table 12. -- (Cont.). 

Habitat type 

Bedrock70 Eelgrass57 Kelp26 Sand11

Common name Catch CPUE FO CPUE FO CPUE FO CPUE FO Page

Juvenile salmon* 1 <0.1 2

Kelp clingfish 1 <0.1 1 90

Pygmy poacher 1 <0.1 2 43

Saddleback sculpin 1 <0.1 1 76

Sailfin sculpin 1 <0.1 2 95

Steelhead trout 1 <0.1 2 137

Wolf-eel 1 <0.1 1 48

Catch 89,687 37,551 35,003 14,318 2,815

Number of species 62 40 51 39 27

Mean CPUE 547 536 614 551 256

 

 

220
Table 13. -- Total catch, mean catch-per-unit-effort (CPUE, unit = seine haul), and percent 

frequency of occurrence (FO) by fish taxa and habitat type in the Southeast 

Northern Inside (SENI) region of Alaska. Fish were captured with a beach seine 

from 1998 to 2006 and 2008 to 2011. Subscript = number of hauls. Page = 

species page. Blanks indicate a species was not captured. *Major target species 

in a fisheries management plan. a

Myoxocephalus spp. = frog, great, or shorthorn 

sculpin. b

Rock sole = northern or southern rock sole. See Figure 10 for the SENI 

region map. The Appendix lists spatial and temporal information on fish catch by 

habitat type. 

Habitat type 

Bedrock67 Eelgrass120 Kelp117 Sand78

Common name Catch CPUE FO CPUE FO CPUE FO CPUE FO Page

Walleye pollock* 123,381 468.0 21 13.6 23 728.8 28 65.7 8 87

Pacific sand lance 71,213 87.8 15 439.4 16 59.9 19 71.8 22 47

Chum salmon* 61,304 61.1 16 428.3 47 39.1 35 16.0 29 135

Pacific herring 54,306 418.5 18 131.3 18 75.8 21 21.2 19 55

Pink salmon* 38,773 119.2 19 72.1 37 37.8 29 227.1 28 134

Pacific sandfish 11,422 169.4 21 <0.1 3 0.4 9 0.3 8 157

Pacific cod* 6,532 25.5 25 13.2 24 12.6 29 22.5 13 85

Crescent gunnel 6,437 1.0 31 46.7 89 3.5 49 4.7 45 117

Tubesnout 5,635 0.3 9 45.2 56 1.3 5 0.5 8 50

Threespine stickleback 3,539 0.1 4 29.3 54 0.1 9 0.1 6 88

Coho salmon* 2,556 0.6 10 16.5 41 4.2 23 0.6 15 136

Snake prickleback 1,912 11.9 53 0.7 6 5.3 17 153

Tubenose poacher 1,336 0.2 9 10.7 54 0.2 9 0.2 9 44

Juvenile sculpin 1,301 0.1 4 7.9 39 1.9 26 1.7 23

Silverspotted sculpin 1,296 1.8 37 6.6 57 2.6 36 1.0 14 94

Myoxocephalus spp.a 919 <0.1 4 5.8 71 0.9 32 1.6 47 74

Shiner perch 817 6.8 3 82

Pacific staghorn sculpin 766 <0.1 1 5.9 69 <0.1 3 0.6 24 68

Capelin 717 3.4 4 <0.1 1 0.7 5 5.3 4 109

Northern sculpin 591 0.6 22 2.9 47 1.0 35 1.1 26 67

Rock sole*b

 541 <0.1 1 1.4 34 0.4 9 4.2 63 119

Bay pipefish 508 3.8 28 0.6 6 156

Juvenile cod* 450 0.9 6 2.8 10 0.1 2 0.6 3

Juvenile greenling 363 <0.1 3 1.8 39 0.6 19 0.9 17

Fish larvae 360 3.7 4 <0.1 3 0.9 4 <0.1 1

Dolly Varden 322 <0.1 1 1.0 18 0.6 19 1.8 17 140

Whitespotted greenling 316 0.2 7 1.8 24 0.5 16 0.4 6 99

Buffalo sculpin 249 0.1 4 1.2 18 0.4 22 0.7 31 60

Surf smelt 247 0.3 3 1.8 3 <0.1 3 108

221
Table 13. -- (Cont.). 

Habitat type 

Bedrock67 Eelgrass120 Kelp117 Sand78

Common name Catch CPUE FO CPUE FO CPUE FO CPUE FO Page

Padded sculpin 204 0.3 3 0.8 15 0.7 9 0.1 6 56

Armorhead sculpin 200 0.1 3 0.1 3 2.3 23 63

Juvenile flatfish 186 0.6 13 0.2 3 1.2 15

Starry flounder 172 0.7 23 <0.1 1 1.1 21 124

Arctic shanny 111 <0.1 1 0.5 12 0.3 17 0.2 8 154

Yellowfin sole* 91 0.7 16 <0.1 2 0.1 5 120

Tidepool sculpin 79 0.3 10 0.3 4 0.1 6 75

Juvenile rockfish 54 <0.1 1 0.2 4 0.2 2 <0.1 1

Masked greenling 45 <0.1 3 0.3 2 <0.1 2 <0.1 1 98

Sturgeon poacher 44 0.1 7 0.1 3 0.3 9 45

Leister sculpin 37 <0.1 3 <0.1 1 0.2 9 0.2 8 62

Lingcod 37 0.1 3 0.2 6 0.1 5 100

Red Irish lord 25 <0.1 1 0.1 3 0.1 7 0.1 3 65

Kelp greenling 24 0.1 3 0.1 7 <0.1 1 96

Dark rockfish 23 0.1 6 0.1 3 <0.1 1 143

Crested sculpin 22 0.1 6 <0.1 3 0.1 7 0.1 1 93

Speckled sanddab 19 0.1 3 0.1 3 114

Penpoint gunnel 16 0.1 4 <0.1 2 <0.1 3 116

Cutthroat trout 13 0.1 5 <0.1 3 <0.1 1 133

Juvenile snailfish 13 0.1 3 <0.1 2

Pacific spiny lumpsucker 12 <0.1 2 0.1 6 80

English sole 10 0.1 6 123

Tidepool snailfish 10 0.1 5 <0.1 1 <0.1 1 104

Juvenile smelt 8 <0.1 1 <0.1 3 <0.1 3

Sand sole 8 0.1 3 128

Sockeye salmon* 8 <0.1 1 <0.1 1 0.1 4 138

Steelhead trout 8 <0.1 2 <0.1 3 <0.1 1 137

Antlered sculpin 7 <0.1 2 0.1 1 61

Showy snailfish 7 0.1 2 106

Eulachon 6 0.1 1 112

Juvenile ronquil 4 <0.1 1

Sailfin sculpin 4 <0.1 3 95

Chinook salmon* 3 <0.1 1 <0.1 1 139

Manacled sculpin 3 <0.1 1 <0.1 2 79

Slender cockscomb 3 <0.1 1 <0.1 1 <0.1 1 150

Butter sole 2 <0.1 1 118

222
Table 13. -- (Cont.). 

Habitat type 

Bedrock67 Eelgrass120 Kelp117 Sand78

Common name Catch CPUE FO CPUE FO CPUE FO CPUE FO Page

Juvenile gunnel 2 <0.1 1 <0.1 1

Pacific sanddab 2 <0.1 1 113

Quillback rockfish 2 <0.1 1 <0.1 1 145

Scalyhead sculpin 2 <0.1 1 <0.1 1 57

Soft sculpin 2 <0.1 2 130

Alaska plaice* 1 <0.1 1 126

Black prickleback 1 <0.1 1 155

Juvenile Irish lord 1 <0.1 1

Juvenile clingfish 1 <0.1 1

Juvenile lumpsucker 1 <0.1 1

Juvenile prickleback 1 <0.1 1

Juvenile trout 1 <0.1 1

Northern ronquil 1 <0.1 1 54

Ribbon snailfish 1 <0.1 1 103

Rock greenling 1 <0.1 1 97

Tadpole sculpin 1 <0.1 1 129

Catch 399,648 91,318 157,590 114,633 36,107

Number of species 67 31 55 52 50

Mean CPUE 1,046 1,363 1,313 980 463

223
Table 14. -- Total catch, mean catch-per-unit-effort (CPUE, unit = seine haul), and percent 

frequency of occurrence (FO) by fish taxa and habitat type in the Southeast 

Southern Outside (SESO) region of Alaska. Subscript = number of hauls. Page = 

species page. Fish were captured with a beach seine from 1998 to 2000 and 

2007. Blanks indicate a species was not captured. *Major target species in a 

fisheries management plan. a

Myoxocephalus spp. = frog, great, or shorthorn 

sculpin. b

Rock sole = northern or southern rock sole. See Figure 11 for the SESO 

region map. The Appendix lists spatial and temporal information on fish catch by 

habitat type. 

Habitat type 

Bedrock9 Eelgrass54 Kelp34 Sand17

Common name Catch CPUE FO CPUE FO CPUE FO CPUE FO Page

Pink salmon* 12,744 164.1 78 176.5 31 45.0 38 11.9 53 134

Pacific sand lance 5,174 390.1 33 19.6 15 14.8 29 6.1 29 47

Shiner perch 5,121 22.2 11 71.1 78 26.9 76 9.8 47 82

Bay pipefish 4,122 70.8 100 5.6 56 6.4 35 156

Threespine stickleback 3,513 23.3 22 50.0 72 1.3 35 32.8 47 88

Crescent gunnel 2,842 0.7 44 39.6 100 14.6 91 12.1 100 117

Chum salmon* 1,321 42.2 33 8.3 46 8.7 53 11.5 53 135

Pacific herring 877 0.1 11 4.8 17 1.2 24 34.0 24 55

Snake prickleback 745 13.1 72 0.8 26 0.7 29 153

Copper rockfish 470 1.8 44 6.0 56 3.9 71 0.1 6 142

Coho salmon* 261 0.7 22 1.5 20 1.3 29 7.5 41 136

Northern sculpin 254 3.2 67 1.4 41 1.8 71 67

Pacific staghorn sculpin 211 0.2 11 3.3 63 0.2 18 1.5 53 68

Myoxocephalus spp.a

 203 0.2 22 2.0 48 0.8 35 3.7 53 74

Tubesnout 174 2.1 28 0.9 12 1.6 12 50

Juvenile greenling 161 2.0 52 1.4 29 0.3 18

Juvenile rockfish 148 1.7 33 1.7 20 1.2 24 0.1 6

English sole 137 2.5 15 <0.1 3 0.1 12 123

Red Irish lord 122 1.9 44 0.6 29 65

Kelp perch 115 0.6 33 0.7 19 2.0 53 0.2 6 81

Juvenile flatfish 114 2.1 4 0.1 6

Pacific cod* 84 0.3 13 1.9 12 85

Juvenile sculpin 74 0.8 15 0.7 18 0.5 12

Whitespotted greenling 72 0.1 11 0.8 26 0.7 26 0.2 12 99

Kelp greenling 63 0.9 28 0.4 26 0.1 6 96

Walleye pollock* 57 <0.1 2 1.7 6 87

Buffalo sculpin 55 0.3 20 0.7 21 0.9 41 60

Brown rockfish 54 0.9 13 0.2 12 141

Tubenose poacher 54 1.0 9 44

224
Table 14. -- (Cont.). 

Habitat type 

Bedrock9 Eelgrass54 Kelp34 Sand17

Common name Catch CPUE FO CPUE FO CPUE FO CPUE FO Page

Blackeye goby 52 0.1 11 0.1 4 1.4 24 92

Painted greenling 38 0.3 22 0.1 6 0.9 26 101

Silverspotted sculpin 34 0.5 15 0.3 9 94

Speckled sanddab 32 0.4 15 0.1 6 0.4 18 114

Dolly Varden 28 <0.1 4 0.2 12 1.1 29 140

Juvenile cod* 26 0.4 11 0.1 6 0.3 12

Pacific sanddab 25 0.2 13 0.7 18 113

Rock sole*b

 24 0.2 7 0.2 9 0.2 12 119

Padded sculpin 20 0.3 9 0.1 9 56

Tidepool sculpin 17 0.1 11 0.2 4 0.2 6 75

Roughback sculpin 13 0.2 2 59

Bay goby 10 <0.1 2 0.3 18 91

Brown Irish lord 10 0.2 2 0.1 6 66

Sailfin sculpin 10 0.1 11 0.1 6 0.2 12 95

Black rockfish 9 0.2 4 146

Cutthroat trout 9 0.2 2 133

Penpoint gunnel 9 0.1 4 0.1 6 116

Masked greenling 8 0.1 2 <0.1 3 0.1 6 98

Sturgeon poacher 8 0.2 2 45

Smoothhead sculpin 7 0.1 2 0.1 9 58

Lingcod 5 0.1 11 <0.1 4 0.1 6 100

Quillback rockfish 5 0.2 6 145

C-O sole 3 <0.1 4 <0.1 3 127

Fluffy sculpin 3 <0.1 2 0.1 6 77

Yellowtail rockfish 3 0.1 3 144

Alaskan ronquil 2 <0.1 2 51

Chinook salmon* 2 <0.1 3 0.1 6 139

Juvenile snailfish 2 0.1 3

Scalyhead sculpin 2 <0.1 2 57

Sockeye salmon* 2 <0.1 2 <0.1 3 138

Starry flounder 2 <0.1 4 124

Tidepool snailfish 2 0.1 6 104

Bocaccio 1 <0.1 2 148

Juvenile Irish lord 1 <0.1 2

Dark rockfish 1 <0.1 3 143

Northern spearnose poacher 1 <0.1 3 40

225
Table 14. -- (Cont.). 

Habitat type 

Bedrock9 Eelgrass54 Kelp34 Sand17

Common name Catch CPUE FO CPUE FO CPUE FO CPUE FO Page

Rock greenling 1 <0.1 3 97

Sablefish* 1 <0.1 2 49

Slender cockscomb 1 <0.1 3 150

Steelhead trout 1 <0.1 3 137

Catch 39,772 5,839 26,545 4,893 2,495

Number of species 62 18 53 51 27

Mean CPUE 349 649 492 144 147

 

226
Table 15. -- Total catch, mean catch-per-unit-effort (CPUE, unit = seine haul), and percent 

frequency of occurrence (FO) by fish taxa and habitat type in the Southeast 

Southern Inside (SESI) region of Alaska. Subscript = number of hauls. Page = 

species page. Fish were captured with a beach seine in 2000 and 2007. Blanks 

indicate a species was not captured. *Major target species in a fisheries 

management plan. a

Myoxocephalus spp. = frog, great, or shorthorn sculpin. 

b

Rock sole = northern or southern rock sole. See Figure 12 for the SESI region 

map. The Appendix lists spatial and temporal information on fish catch by 

habitat type. 

Habitat type 

Bedrock4 Eelgrass13 Kelp17 

Common name Catch CPUE FO CPUE FO CPUE FO Page

Pacific herring 5,030 0.1 8 295.8 12 55

Shiner perch 3,211 11.3 50 206.2 85 28.6 65 82

Bay pipefish 1,313 0.3 25 100.3 85 0.5 12 156

Threespine stickleback 811 61.2 77 0.9 24 88

Crescent gunnel 749 0.3 25 41.9 100 12.0 82 117

Chum salmon* 552 0.3 25 4.7 15 28.8 35 135

Snake prickleback 488 35.2 77 1.8 24 153

Tubesnout 138 0.8 25 9.9 77 0.4 18 50

Walleye pollock* 138 8.1 18 87

Pink salmon* 121 3.5 75 1.1 15 5.5 18 134

Northern sculpin 98 4.8 54 2.1 65 67

Pacific sand lance 94 0.5 23 5.2 18 47

Pacific staghorn sculpin 88 6.1 85 0.5 24 68

Myoxocephalus spp.a

 76 4.1 69 1.4 41 74

Coho salmon* 55 2.5 31 1.4 29 136

Rock sole*b 23 0.1 8 1.3 18 119

Buffalo sculpin 22 0.8 38 0.7 35 60

English sole 20 1.5 8 123

Padded sculpin 19 0.3 25 1.1 23 0.2 6 56

Tadpole sculpin 18 1.4 15 129

Whitespotted greenling 18 0.9 15 0.4 24 99

Tubenose poacher 17 0.5 23 0.7 12 44

Silverspotted sculpin 16 0.5 15 0.6 24 94

Copper rockfish 15 0.2 8 0.8 24 142

Surf smelt 13 0.7 15 0.2 24 108

Tidepool sculpin 13 0.5 23 0.4 6 75

Dolly Varden 12 0.1 8 0.7 24 140

Juvenile greenling 10 0.6 31 0.1 12

227
Table 15. -- (Cont.). 

Habitat type 

Bedrock4 Eelgrass13 Kelp17 

Common name Catch CPUE FO CPUE FO CPUE FO Page

Cabezon 8 0.5 6 78

Kelp perch 8 2.0 50 81

Pacific sanddab 8 0.4 31 0.2 12 113

Smoothhead sculpin 7 0.3 25 0.5 8 58

Speckled sanddab 7 0.5 8 114

Juvenile sculpin 6 0.2 15 0.2 12

Arctic shanny 5 0.3 12 154

Pacific cod* 5 0.1 8 0.2 18 85

Painted greenling 4 0.2 12 101

Brown Irish lord 3 0.1 8 0.1 6 66

Cutthroat trout 3 0.2 8 133

Northern spearnose poacher 3 0.2 8 40

Starry flounder 3 0.2 23 124

Alaskan ronquil 2 0.1 8 0.1 6 51

Blackeye goby 2 0.1 12 92

Kelp greenling 2 0.1 8 0.1 6 96

Red Irish lord 2 0.2 15 65

Juvenile gunnel 1 0.1 6

Manacled sculpin 1 0.1 8 79

Pacific sandfish 1 0.1 6 157

Tidepool snailfish 1 0.1 8 104

Yellowfin sole* 1 0.1 6 120

Catch subtotal 13,261 75 6,369 6,817

Number of species 47 9 39 36

Mean CPUE 390 19 490 401

 

228
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Figure 1. -- Regions and sites sampled for nearshore fishes in Alaska from 1998 to 2011. Fish were sampled with a beach seine

in nine regions (inset) encompassing 555 sites (solid circles in full-scale map). See the Appendix for spatial andtemporal site information on fish catch by habitat type (i.e., bedrock, eelgrass, kelp, and sand).

229
Percent frequency of beach seine hauls by month from 1998 to 2011 in nearshore, 

marine waters of Alaska. 

Figure 2. -- 

0 

10 

20 

30 

Jan Feb Mar Apr May Jun Jul Aug Sep 

Frequency (%) 

n = 1,009 hauls 

Month 

230
Figure 3. -- Examples of the four habitat types sampled with a beach seine for nearshore, 

marine fishes in Alaska from 1998 to 2011. Clockwise from upper left: bedrock 

outcrop, eelgrass meadow, understory kelp bed, and sand or gravel beach. 

231
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Canada

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Figure 4. -- Locations and sites sampled for nearshore fishes in the Arctic (AR) region of Alaska from 2004 to 2009. Fish were

sampled with a beach seine at two locations (numbered boxes) encompassing 31 sites (solid circles). Sites sampledwere all sand beaches. See the Appendix for spatial and temporal site information on fish catch by habitat type.

232
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Figure 5. -- Locations and sites sampled for nearshore fishes in the Aleutian Islands (AI) region of Alaska in 2005. Fish were

sampled with a beach seine at six locations (numbered boxes) encompassing 70 sites (solid circles). Sites sampled

were bedrock, kelp, and sand habitat types. See the Appendix for spatial and temporal site information on fish

catch by habitat.

233
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Figure 6. -- Location and sites sampled for nearshore fishes in the Bristol Bay (BB) region of Alaska in 2009. Fish were

sampled with a beach seine at one location (numbered box) encompassing eight sites (solid circles). Sites sampledwere all sand beaches. See the Appendix for spatial and temporal site information on fish catch by habitat type.

234
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Figure 7. -- Locations and sites sampled for nearshore fishes in the Southcentral (SC) region of Alaska from 2008 to 2010. Fish

were sampled with a beach seine at five locations (numbered boxes) encompassing 22 sites (solid circles). Sites

sampled were bedrock, eelgrass, kelp, and sand habitat types. See the Appendix for spatial and temporal siteinformation on fish catch by habitat.

235
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Figure 8. -- Locations and sites sampled for nearshore fishes in the Prince William Sound (PWS) region of Alaska in 1999,

2006, 2007, 2009, and 2010. Fish were sampled with a beach seine at 13 locations (numbered boxes) encompassing

50 sites (solid circles). Sites sampled were bedrock, eelgrass, kelp, and sand habitat types. See the Appendix forspatial and temporal site information on fish catch by habitat.

236
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Figure 9. -- Locations and sites sampled for nearshore fishes in the Southeast Northern Outside (SENO) region of Alaska in

1998, 1999, 2001 to 2003, 2006, 2008, and 2010. Fish were sampled with a beach seine at 17 locations (numberedboxes) encompassing 127 sites (solid circles). Sites sampled were bedrock, eelgrass, kelp, and sand habitat types.

See the Appendix for spatial and temporal site information on fish catch by habitat.

237
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Figure 10. -- Locations and sites sampled for nearshore fishes in the Southeast Northern Inside (SENI) region of Alaska from

1998 to 2006 and 2008 to 2011. Fish were sampled with a beach seine at 27 locations (numbered boxes)encompassing 153 sites (solid circles). Sites sampled were bedrock, eelgrass, kelp, and sand habitat types. See theAppendix for spatial and temporal site information on fish catch by habitat.

238
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82

81

80

79

77

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75

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Figure 11. -- Locations and sites sampled for nearshore fishes in the Southeast Southern Outside (SESO) region of Alaska from

1998 to 2000 and 2007. Fish were sampled with a beach seine at 12 locations (numbered boxes) encompassing 60sites (solid circles). Sites sampled were bedrock, eelgrass, kelp, and sand habitat types. See the Appendix forspatial and temporal site information on fish catch by habitat.

239
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0 25 50 km

Figure 12. -- Locations and sites sampled for nearshore fishes in the Southeast Southern Inside (SESI) region of Alaska in 2000

and 2007. Fish were sampled with a beach seine at 10 locations (numbered boxes) encompassing 34 sites (solidcircles). Sites sampled were bedrock, eelgrass, and kelp habitat types. See the Appendix for spatial and temporalsite information on fish catch by habitat.

240
Figure 13. -- The 37-m long variable mesh beach seine used to sample nearshore fishes in nine 

regions of Alaska (top), and setting the seine from a small boat in bedrock habitat 

(bottom). 

241
SESO 

AI 

SENO 

SENI 

SESI 

SC 

PWS 

AI 

SENO 

SENI 

SESO 

SESI 

SC 

PWS 

AI 

SENO 

SENI SESO 

BB 

SC 

PWS 

AR 

2D Stress: 0.05 

2D Stress: 0.07 2D Stress: 0.12 

2D Stress: 0.0 

Bedrock 

Kelp Sand 

Eelgrass 

SENO 

SENI 

SESO 

SESI 

PWS 

Relative similarities of nearshore fish assemblages among seven different 

regions of Alaska, grouped within four different habitat types. The 2nd stage 

nMDS plots show the centroid locations (ANOSIM R value) of fish 

assemblages by region within each habitat type (bedrock, eelgrass, kelp, and 

sand). The closer each region is to each other in the plot indicates more 

similarity among regional species assemblages; regions within circles are not 

significantly different, P > 0.05. A stress value < 0.1 corresponds to good 2-D 

representation of the nMDS ordination. Regions are Aleutian Islands (AI), 

Southcentral (SC), Prince William Sound (PWS), Southeast northern outside 

(SENO) and inside (SENI), and Southeast southern outside (SESO) and inside 

(SESI). See Figure 1 for region boundaries. 

Figure 14. -- 

242
Bedrock Eelgrass 

Sand 

Kelp 

Sand 

Eelgrass Kelp 

Bedrock 

SC Kelp 

Bedrock 

Sand 

Eelgrass 

Kelp 

Bedrock 

Sand 

Eelgrass 

Kelp 

Bedrock 

Sand 

Bedrock Kelp 

Sand 

SENI SESO 

SC 

PWS SENO 

AI 

SESI 

2D Stress: 0.0 2D Stress: 0.0 

2D Stress: 0.0 

2D Stress: 0.0 2D Stress: 0.0 

2D Stress: 0.0 

2D Stress: 0.08 Eelgrass 

Kelp 

Bedrock 

Relative similarities of nearshore fish assemblages by habitat type, grouped 

within seven different regions of Alaska. The 2nd stage nMDS plots show the 

centroid locations (ANOSIM R value) of fish assemblages by habitat type 

(bedrock, eelgrass, kelp, and sand) within each region; not all habitats were 

sampled in each region. The closer each habitat is to each other in the plot 

indicates more similarity among habitat-specific species assemblages; habitats 

within circles are not significantly different, P > 0.05. A stress value < 0.1 

corresponds to good 2-D representation of the nMDS ordination. Regions are 

Aleutian Islands (AI), Southcentral (SC), Prince William Sound (PWS), 

Southeast northern outside (SENO) and inside (SENI), and Southeast southern 

outside (SESO) and inside (SESI). See Figure 1 for region boundaries. 

Figure 15. -- 

243
Appendix. -- Spatial and temporal site information on fish catch (all species) by habitat type 

in nearshore waters of Alaska from 1998 to 2011. Regions are in bold, and 

locations are numbered from 1 to 93 in parentheses. Sites sampled were 

bedrock, eelgrass, kelp, and sand habitat types. Coordinates = N latitude and W 

longitude. Hauls are number of beach seine hauls. Regions are shown in Figure 

1, and locations and sites by region are shown in Figures 4-12. 

Region, Location, Site, Coordinates Dates sampled Hauls Catch

Arctic 

Chukchi Sea (1) 

sand 71.3883 , 156.4773 8/05, 8/06 2 1,592

sand 71.3869 , 156.4804 8/05 1 5

sand 71.3816 , 156.4802 8/05, 8/06, 8/07, 8/08, 8/09, 9/09 6 1,049

sand 71.3510 , 156.5706 8/07, 8/08, 8/09, 9/09 4 4,231

sand 71.3155 , 156.7221 8/05, 8/06 2 64

sand 71.3056 , 156.7550 8/07, 8/08, 8/09, 9/09 4 1,246

sand 71.3023 , 156.7646 8/05, 8/06 2 117

sand 71.2930 , 156.7907 8/05, 8/06 2 142

sand 71.2805 , 156.8216 8/05, 8/06 2 163

sand 71.2440 , 156.8947 8/05, 8/06 2 259

sand 71.2375 , 156.9101 8/07, 8/08, 8/09, 9/09 4 508

sand 71.2143 , 156.9565 8/05, 8/06 2 92

sand 71.1816 , 157.0179 8/05, 8/06 2 24

sand 71.1644 , 157.0481 8/07, 8/08, 8/09, 9/09 4 7,575

sand 71.0897 , 157.1791 8/07, 8/08, 8/09, 9/09 4 1,682

sand 70.8432 , 157.9452 8/05 1 30

sand 70.8399 , 157.9678 8/05 1 19

sand 70.8377 , 157.9835 8/05 1 20

sand 70.8355 , 158.0004 8/05 1 10

Beaufort Sea (2) 

sand 71.3685 , 156.4014 8/05, 8/06 2 283

sand 71.3681 , 156.4025 8/05, 8/06 2 81

sand 71.3484 , 156.2742 8/05 1 17

sand 71.3472 , 156.2710 8/05 1 10

sand 71.3381 , 156.1677 8/05 1 9

sand 71.2721 , 155.9245 8/05, 8/06 2 4

sand 71.2372 , 155.7178 8/04, 8/05, 8/06, 8/07, 8/09, 9/09 6 210

sand 71.2322 , 155.6858 8/04, 8/05, 8/06 3 11

sand 71.2322 , 155.6350 8/04, 8/05, 8/06, 8/07, 9/09 5 1,054

sand 71.2313 , 155.6347 8/04, 8/05, 8/06 3 39

sand 71.2272 , 155.5817 8/04, 8/05, 8/06, 8/07, 9/09 5 242

sand 71.2261 , 155.5880 8/04, 8/05, 8/06 3 15

244
Appendix -- (Cont.). 

Region, Location, Site, Coordinates Dates sampled Hauls Catch

Aleutian Islands

Reese Bay (3) 

bedrock 54.0079 , 166.6954 6/05 1 202

bedrock 54.0031 , 166.6931 6/05 1 1

bedrock 53.9928 , 166.8219 6/05 1 16

sand 53.9994 , 166.7126 6/05 1 6

sand 53.9991 , 166.7055 6/05 1 18

sand 53.9927 , 166.8258 6/05 1 3,993

sand 53.9793 , 166.8372 6/05 1 1,663

Unalaska Bay (4) 

kelp 53.9321 , 166.6410 6/05 1 2,051

kelp 53.9194 , 166.4380 6/05 1 197

sand 53.9498 , 166.6264 6/05 1 8,166

sand 53.9226 , 166.6480 6/05 1 168

sand 53.9146 , 166.6409 6/05 1 1,095

sand 53.9122 , 166.4387 6/05 1 1,454

sand 53.9017 , 166.4449 6/05 1 29

sand 53.8989 , 166.4647 6/05 1 2,037

sand 53.8857 , 166.6299 6/05 1 16,181

Beaver Inlet (5) 

bedrock 53.7808 , 166.5109 6/05 1 8

bedrock 53.7785 , 166.4858 6/05 1 263

bedrock 53.7774 , 166.5000 6/05 1 373

bedrock 53.7316 , 166.2934 6/05 1 9

bedrock 53.7104 , 166.5582 6/05 1 50

bedrock 53.7019 , 166.5399 6/05 1 21

kelp 53.8204 , 166.2117 6/05 1 12,215

kelp 53.7924 , 166.5065 6/05 1 253

kelp 53.7885 , 166.2176 6/05 1 4,994

kelp 53.7825 , 166.2737 6/05 1 458

kelp 53.7770 , 166.2363 6/05 1 326

kelp 53.7590 , 166.2991 6/05 1 173

kelp 53.7394 , 166.2909 6/05 1 18

kelp 53.7313 , 166.3202 6/05 1 873

kelp 53.7306 , 166.3269 6/05 1 926

kelp 53.7131 , 166.5377 6/05 1 154

kelp 53.7088 , 166.5541 6/05 1 98

kelp 53.7019 , 166.5413 6/05 1 354

245
Appendix -- (Cont.). 

Region, Location, Site, Coordinates Dates sampled Hauls Catch

sand 53.8092 , 166.2087 6/05 1 150

sand 53.7893 , 166.5201 6/05 1 17

Unalga Pass (6) 

kelp 53.9178 , 166.2621 6/05 1 203

kelp 53.9169 , 166.2675 6/05 1 944

kelp 53.9141 , 166.2577 6/05 1 65

kelp 53.9130 , 166.2657 6/05 1 288

sand 53.9465 , 166.3480 6/05 1 72

sand 53.9457 , 166.3405 6/05 1 9

sand 53.9457 , 166.3427 6/05 1 48

sand 53.9456 , 166.3388 6/05 1 8

sand 53.9129 , 166.2703 6/05 1 121

sand 53.9123 , 166.2711 6/05 1 113

Akutan Island (7) 

bedrock 54.1200 , 165.7696 6/05 1 40

kelp 54.1751 , 165.8075 6/05 1 15,149

kelp 54.1743 , 165.8080 6/05 1 3,143

kelp 54.1674 , 165.8041 6/05 1 34

kelp 54.1363 , 165.7577 6/05 1 626

kelp 54.1141 , 165.7463 6/05 1 329

sand 54.1703 , 165.8081 6/05 1 105

sand 54.1696 , 165.8074 6/05 1 23

sand 54.1681 , 165.8054 6/05 1 298

sand 54.1455 , 165.7354 6/05 1 61

sand 54.1450 , 165.7385 6/05 1 660

sand 54.1286 , 165.8230 6/05 1 217

sand 54.1180 , 165.7657 6/05 1 31

sand 54.1172 , 165.7622 6/05 1 1,419

Akun Island (8) 

kelp 54.2392 , 165.6227 6/05 1 18

kelp 54.2379 , 165.6246 6/05 1 35

kelp 54.2373 , 165.6248 6/05 1 483

kelp 54.2365 , 165.6248 6/05 1 78

sand 54.2445 , 165.6114 6/05 1 87

sand 54.2439 , 165.6088 6/05 1 326

sand 54.2430 , 165.6055 6/05 1 6

sand 54.2412 , 165.5976 6/05 1 17

sand 54.2388 , 165.5917 6/05 1 43

sand 54.2334 , 165.5852 6/05 1 9

246
Appendix -- (Cont.). 

Region, Location, Site, Coordinates Dates sampled Hauls Catch

Bristol Bay 

Nushagak Bay (9) 

sand 58.8565 , 158.4909 7/09 1 261

sand 58.8465 , 158.5545 7/09 1 2,443

sand 58.8460 , 158.5545 7/09 1 1,761

sand 58.8218 , 158.5503 7/09 1 89

sand 58.7848 , 158.6676 7/09 1 923

sand 58.7725 , 158.6598 7/09 1 203

sand 58.5895 , 158.7789 7/09 1 873

sand 58.5783 , 158.7734 7/09 1 222

Southcentral 

Knik Arm (10) 

sand 61.3451 , 149.7154 7/10 1 4

sand 61.3442 , 149.7229 7/10 1 1

Fire Island (11) 

sand 61.1713 , 150.2225 8/09, 7/10 2 123

sand 61.1569 , 150.2605 8/09 1 0

sand 61.1282 , 150.2847 8/09 1 4

Inner Kachemak Bay (12) 

bedrock 59.5945 , 151.2793 7/08 1 259

kelp 59.5782 , 151.2687 7/08 1 163

sand 59.5795 , 151.2974 7/08 1 20

sand 59.5670 , 151.3511 7/08 1 8

Outer Kachemak Bay (13) 

bedrock 59.5122 , 151.4627 7/08 1 9

bedrock 59.5065 , 151.5331 7/08 1 2

bedrock 59.4830 , 151.5001 7/08 1 7

kelp 59.5133 , 151.4581 7/08 1 29

kelp 59.5130 , 151.4231 7/08 1 671

kelp 59.5052 , 151.5308 7/08 1 33

kelp 59.5036 , 151.4380 7/08 1 8

kelp 59.4684 , 151.5319 7/08 1 354

kelp 59.4516 , 151.4207 7/08 1 121

kelp 59.4368 , 151.3989 7/08 1 40

sand 59.4860 , 151.5711 7/08 1 23

Resurrection Bay (14) 

eelgrass 60.1026 , 149.3644 8/09 1 78

sand 60.0783 , 149.3474 8/09 1 901

247
Appendix -- (Cont.). 

Region, Location, Site, Coordinates Dates sampled Hauls Catch

Prince William Sound

Flemming Island (15) 

bedrock 60.1824 , 148.0297 4/06, 7/06, 7/07 3 24

eelgrass 60.1684 , 148.0407 4/06, 7/06, 9/06, 7/07, 8/07 5 899

kelp 60.1671 , 148.0437 4/06, 7/06, 9/06, 7/07, 8/07 5 412

Whale Bay, East Arm (16) 

bedrock 60.1835 , 148.1755 4/06, 7/06, 9/06, 7/07 4 256

eelgrass 60.2091 , 148.1528 4/06, 7/06, 9/06, 7/07, 8/07 5 1,935

kelp 60.1868 , 148.1726 4/06, 7/06, 9/06, 7/07, 8/07 5 856

Whale Bay, West Arm (17) 

bedrock 60.2258 , 148.2537 4/06, 7/06, 9/06, 7/07 4 46

eelgrass 60.2287 , 148.2502 4/06, 7/06, 9/06, 7/07, 8/07 5 1,101

kelp 60.2358 , 148.2372 4/06, 7/06, 9/06, 7/07, 8/07 5 699

Jackpot Bay (18) 

bedrock 60.3442 , 148.2315 4/06, 7/06, 9/06, 7/07 4 24

eelgrass 60.3593 , 148.2377 4/06, 7/06, 9/06, 7/07, 8/07 5 13,901

kelp 60.3400 , 148.2222 4/06, 7/06, 9/06, 7/07, 8/07 5 1,743

sand 60.3605 , 148.2483 4/06 1 59

Chenega Island (19) 

bedrock 60.3584 , 148.0958 4/06, 7/06, 9/06, 7/07 4 38

eelgrass 60.3601 , 148.0894 4/06, 7/06, 9/06, 7/07, 8/07 5 14,865

kelp 60.3629 , 148.0902 4/06, 7/06, 9/06, 7/07, 8/07 5 379

Ewan Bay (20) 

bedrock 60.3763 , 148.1379 4/06, 7/06, 9/06, 7/07 4 1,985

eelgrass 60.3812 , 148.1538 4/06, 7/06, 9/06, 7/07, 8/07 5 390

kelp 60.3728 , 148.1377 4/06, 7/06, 9/06, 7/07, 8/07 5 225

Culross Passage (21) 

bedrock 60.6573 , 148.1863 5/06, 7/06, 9/06, 7/07 4 1,028

eelgrass 60.6244 , 148.2018 5/06, 7/06, 9/06, 7/07 4 574

kelp 60.6343 , 148.1719 5/06, 7/06, 9/06, 7/07 4 5,466

Culross Island (22) 

bedrock 60.7446 , 148.2206 5/06, 7/06, 9/06, 7/07 4 9

eelgrass 60.7447 , 148.2189 5/06, 7/06, 9/06, 7/07 4 807

kelp 60.7455 , 148.2143 5/06, 7/06, 9/06, 7/07 4 2,027

Shotgun Cove (23) 

eelgrass 60.7944 , 148.5332 7/06 1 127

eelgrass 60.7854 , 148.5623 7/06 1 29

eelgrass 60.7851 , 148.5623 7/06 1 63

kelp 60.7963 , 148.5337 7/06 1 122

248
Appendix -- (Cont.). 

Region, Location, Site, Coordinates Dates sampled Hauls Catch

Whittier (24) 

sand 60.7855 , 148.7176 8/09 1 444

sand 60.7831 , 148.7171 8/09 1 3,062

sand 60.7798 , 148.7153 8/09 1 67

sand 60.7778 , 148.7140 8/09 1 4,612

Valdez (25) 

eelgrass 61.1288 , 146.4705 7/10 1 305

kelp 61.1228 , 146.3402 7/10 1 142

kelp 61.1156 , 146.5863 7/10 1 367

sand 61.1259 , 146.6034 7/10 1 25

sand 61.1257 , 146.4099 7/10 1 61

Bligh Island (26) 

eelgrass 60.8689 , 146.7561 7/99 1 232

eelgrass 60.8678 , 146.7536 7/99 1 88

eelgrass 60.8636 , 146.8069 7/99 1 243

kelp 60.8650 , 146.8072 7/99 1 20

kelp 60.8650 , 146.8072 7/99 1 16

Tatitlek (27) 

eelgrass 60.8647 , 146.6844 7/99 1 285

eelgrass 60.8622 , 146.6794 7/99 1 173

eelgrass 60.8617 , 146.6669 7/99 1 313

eelgrass 60.8603 , 146.6742 7/99 1 175

eelgrass 60.8586 , 146.7083 7/99 1 431

kelp 60.8592 , 146.6711 7/99 1 46

kelp 60.8592 , 146.7092 7/99 1 227

Southeastern - Northern Outside 

Torch Bay (28) 

eelgrass 58.3306 , 136.7715 7/10 1 1,897

kelp 58.3403 , 136.8075 7/10 1 54

kelp 58.3227 , 136.8020 7/10 1 808

Dundas Point (29) 

eelgrass 58.3355 , 136.2052 7/10 1 419

kelp 58.3341 , 136.2046 7/10 1 9

sand 58.3266 , 136.2154 7/10 1 6

Port Althorp (30) 

eelgrass 58.1111 , 136.2789 8/99, 6/08 2 162

eelgrass 58.1094 , 136.2750 8/99 1 134

kelp 58.1553 , 136.3081 8/99 1 58

kelp 58.1297 , 136.3000 8/99, 6/08 2 317

249
Appendix -- (Cont.). 

Region, Location, Site, Coordinates Dates sampled Hauls Catch

Yakobi Island (31) 

kelp 58.0989 , 136.4950 8/99 1 10,049

kelp 58.0964 , 136.4947 8/99 1 11

kelp 58.0775 , 136.4553 8/99 1 18

kelp 58.0742 , 136.4511 8/99 1 33

Islas Bay (32) 

bedrock 57.8093 , 136.3622 7/10 1 4

eelgrass 57.8437 , 136.3763 7/10 1 215

eelgrass 57.8310 , 136.3945 7/10 1 136

kelp 57.8297 , 136.3916 7/10 1 61

kelp 57.8223 , 136.3610 7/10 1 1,119

kelp 57.8026 , 136.3774 7/10 1 69

Neva Strait (33) 

bedrock 57.3439 , 135.6750 4/98 1 351

bedrock 57.3403 , 135.7094 4/98 1 10

bedrock 57.3400 , 135.7061 4/98, 8/98 2 2,084

bedrock 57.3358 , 135.7111 8/98 1 1,049

bedrock 57.3244 , 135.6689 4/98 1 43

bedrock 57.3231 , 135.6706 4/98 1 16

bedrock 57.3225 , 135.6694 4/98, 8/98 2 17

bedrock 57.3193 , 135.6979 8/98 1 1,194

bedrock 57.3186 , 135.6719 4/98 1 11

St. John Baptist Bay (34) 

bedrock 57.2925 , 135.6050 8/98 1 510

bedrock 57.2922 , 135.6044 8/98 1 41

bedrock 57.2922 , 135.6036 4/98 1 14

bedrock 57.2903 , 135.5874 4/98, 8/98 2 13

bedrock 57.2900 , 135.5883 4/98, 8/98 2 30

eelgrass 57.2834 , 135.5501 8/98 1 141

eelgrass 57.2834 , 135.5502 4/98, 8/98 2 171

sand 57.2858 , 135.5576 4/98, 8/98 2 52

sand 57.2847 , 135.5630 8/98 1 23

Krestof Sound (35) 

bedrock 57.2211 , 135.5762 6/06 1 1,983

eelgrass 57.2417 , 135.6227 6/06 1 2,572

kelp 57.2299 , 135.5985 6/06 1 31

Nakwasina Sound (36) 

bedrock 57.2497 , 135.3686 8/98 1 10

bedrock 57.2422 , 135.3519 8/98 1 13

250
Appendix -- (Cont.). 

Region, Location, Site, Coordinates Dates sampled Hauls Catch

bedrock 57.2422 , 135.3519 5/98, 8/98 2 59

bedrock 57.2372 , 135.3647 5/98 1 136

bedrock 57.2372 , 135.3647 5/98 1 1

bedrock 57.2364 , 135.3550 5/98, 8/98 2 3,207

bedrock 57.2319 , 135.3708 5/98, 8/98 2 6,122

bedrock 57.2203 , 135.3964 5/98 1 4,300

bedrock 57.2197 , 135.3942 8/98 1 36

eelgrass 57.1975 , 135.3833 6/01, 6/02, 1/03, 6/03, 6/08 5 1,329

eelgrass 57.1975 , 135.3842 5/98, 8/98 2 559

eelgrass 57.1964 , 135.3847 6/01, 6/02, 1/03, 6/03, 6/08 5 4,921

eelgrass 57.1964 , 135.3853 8/98 1 486

sand 57.1939 , 135.3878 5/98, 8/98 2 471

sand 57.1938 , 135.3891 8/98 1 32

Katlian Bay (37) 

bedrock 57.1803 , 135.3378 8/98 1 64

bedrock 57.1803 , 135.3378 5/98 1 7

bedrock 57.1772 , 135.3078 8/98 1 393

bedrock 57.1768 , 135.2965 5/98 1 196

bedrock 57.1661 , 135.3586 5/98, 8/98 2 2,055

bedrock 57.1652 , 135.3627 5/98 1 3,000

bedrock 57.1639 , 135.3294 8/98 1 53

bedrock 57.1588 , 135.3781 5/98 1 2

bedrock 57.1564 , 135.3533 5/98, 8/98 2 18

bedrock 57.1473 , 135.3763 5/98 1 4,000

bedrock 57.1469 , 135.3902 5/98 1 0

bedrock 57.1442 , 135.3939 5/98 1 3,001

bedrock 57.1440 , 135.3937 5/98 1 128

bedrock 57.1424 , 135.3758 6/06 1 0

bedrock 57.1419 , 135.3773 5/98 1 1

eelgrass 57.1859 , 135.3478 5/98, 8/98 2 1,121

eelgrass 57.1645 , 135.3112 8/99 1 129

eelgrass 57.1644 , 135.3104 8/99 1 96

eelgrass 57.1640 , 135.3144 5/98, 8/98 2 1,311

eelgrass 57.1529 , 135.3610 6/06 1 1,597

kelp 57.1647 , 135.3222 8/99 1 19

kelp 57.1646 , 135.3231 8/99 1 21

kelp 57.1551 , 135.3535 6/06 1 302

sand 57.1606 , 135.3419 5/98, 8/98 2 1,750

sand 57.1602 , 135.3423 8/98 1 473

251
Appendix -- (Cont.). 

Region, Location, Site, Coordinates Dates sampled Hauls Catch

Middle Island (38) 

eelgrass 57.0931 , 135.4494 8/99 1 163

eelgrass 57.0923 , 135.4481 8/99 1 532

eelgrass 57.0889 , 135.4504 8/99 1 188

kelp 57.0916 , 135.4493 8/99 1 28

Sitka (39) 

bedrock 57.0375 , 135.2929 6/06 1 67

eelgrass 57.0436 , 135.3037 6/06 1 358

kelp 57.0429 , 135.3001 6/06 1 298

kelp 57.0389 , 135.3276 6/06 1 63

Silver Bay (40) 

bedrock 57.0094 , 135.1639 8/98 1 10

bedrock 57.0044 , 135.1569 8/98 1 1

bedrock 56.9972 , 135.1644 8/98 1 121

bedrock 56.9858 , 135.1333 8/98 1 7

Sandy Cove (41) 

eelgrass 56.9788 , 135.3172 8/99 1 2,561

eelgrass 56.9786 , 135.3108 6/01, 6/02, 1/03, 6/03, 6/08 5 5,256

eelgrass 56.9782 , 135.3117 8/99 1 1,993

eelgrass 56.9781 , 135.3119 6/01, 6/02, 1/03, 6/03, 6/08 5 3,134

kelp 56.9792 , 135.3205 8/99 1 318

kelp 56.9786 , 135.3108 8/99 1 409

Pirate Cove (42) 

eelgrass 56.9853 , 135.3753 6/02 1 1,870

eelgrass 56.9852 , 135.3753 8/99 1 168

eelgrass 56.9849 , 135.3713 6/01, 5/03 2 493

eelgrass 56.9844 , 135.3717 6/01, 6/02, 5/03 3 322

kelp 56.9856 , 135.3712 8/99 1 45

kelp 56.9850 , 135.3739 8/99 1 22

Redoubt Bay (43) 

bedrock 56.9258 , 135.3524 6/06 1 2

bedrock 56.8828 , 135.3469 6/06 1 3,061

eelgrass 56.9110 , 135.3260 6/06 1 120

eelgrass 56.9106 , 135.3246 6/06 1 85

eelgrass 56.8788 , 135.3411 6/06 1 364

kelp 56.9253 , 135.3450 6/06 1 141

kelp 56.8772 , 135.3383 6/06 1 15

sand 56.9090 , 135.4002 6/06 1 8

252
Appendix -- (Cont.). 

Region, Location, Site, Coordinates Dates sampled Hauls Catch

Biorka Channel (44) 

bedrock 56.8475 , 135.4964 8/98 1 3

bedrock 56.8472 , 135.4178 8/98 1 2

bedrock 56.8472 , 135.5089 8/98 1 0

bedrock 56.8456 , 135.4944 8/98 1 7

bedrock 56.8450 , 135.4933 8/98 1 0

bedrock 56.8433 , 135.3931 8/98 1 6

bedrock 56.8433 , 135.3931 8/98 1 54

bedrock 56.8419 , 135.5172 8/98 1 1

bedrock 56.8394 , 135.4239 8/98 1 21

bedrock 56.8378 , 135.5247 8/98 1 0

bedrock 56.8378 , 135.5322 8/98 1 3

bedrock 56.8364 , 135.4142 8/98 1 6

bedrock 56.8364 , 135.4175 8/98 1 5

bedrock 56.8303 , 135.4247 8/98 1 2

Southeastern - Northern Inside 

Haines (45) 

sand 59.2348 , 135.4384 8/10 1 38

sand 59.2307 , 135.4394 8/10 1 23

Berners Bay (46) 

eelgrass 58.6719 , 134.9148 6/04, 7/05, 8/06 3 663

eelgrass 58.6693 , 134.9127 6/04, 7/05, 8/06 3 434

eelgrass 58.6686 , 134.9121 6/04, 7/05, 8/06 3 596

kelp 58.7057 , 134.9464 6/04, 7/05, 8/06 4 3,093

kelp 58.6954 , 134.9356 6/04, 7/05, 8/06 4 395

kelp 58.6941 , 134.9342 6/04, 7/05, 8/06 4 122

sand 58.7852 , 135.0202 5/01 1 2

sand 58.7778 , 135.0078 5/01 1 192

sand 58.7738 , 134.9353 5/01 1 1,098

sand 58.7735 , 134.9862 5/01 1 46

St. James Bay (47) 

eelgrass 58.6100 , 135.2094 7/00 1 233

eelgrass 58.5731 , 135.1825 7/00 1 164

Bridget Cove (48) 

bedrock 58.6492 , 134.9689 7/05 1 3

eelgrass 58.6421 , 134.9556 6/04, 7/05, 8/06 3 3,439

eelgrass 58.6348 , 134.9480 6/04, 7/05, 8/06, 7/09 4 6,318

eelgrass 58.6303 , 134.9449 6/04, 7/05, 8/06 3 3,584

eelgrass 58.6285 , 134.9425 6/98, 8/98, 8/99 4 5,174

253
Appendix -- (Cont.). 

Region, Location, Site, Coordinates Dates sampled Hauls Catch

eelgrass 58.6274 , 134.9453 6/98, 8/98, 8/99 4 104

kelp 58.6419 , 134.9581 6/04, 7/05, 8/06 4 1,017

kelp 58.6405 , 134.9551 6/04, 7/05, 8/06 4 1,148

kelp 58.6386 , 134.9545 6/04, 7/05, 8/06 3 734

Yankee Cove (49) 

sand 58.6013 , 134.9187 4/01 1 4

sand 58.5902 , 134.9045 4/01 1 5

Benjamin Island (50) 

bedrock 58.5611 , 134.8996 7/01, 3/02, 7/02, 3/03, 7/03, 3/04 6 538

bedrock 58.5481 , 134.9256 8/01, 3/02, 7/02, 3/03, 7/03, 3/04 6 451

bedrock 58.5444 , 134.9206 8/01, 3/02, 3/03, 7/03, 3/04 5 218

kelp 58.5752 , 134.9263 4/01, 7/01, 3/02, 7/02, 3/03, 7/03, 3/04 7 120

kelp 58.5675 , 134.9150 4/01, 7/01, 3/02, 7/02, 3/03, 7/03, 3/04 7 4,707

kelp 58.5631 , 134.9017 4/01, 7/01, 3/02, 7/02, 3/03, 7/03, 3/04 7 934

sand 58.5717 , 134.9242 4/01 1 83

sand 58.5644 , 134.9102 7/01, 3/02, 7/02, 3/03, 7/03, 3/04 6 4,734

sand 58.5547 , 134.9061 4/01, 8/01, 3/02, 7/02, 3/03, 7/03, 3/04 7 271

sand 58.5547 , 134.8983 4/01, 8/01, 3/02, 7/02, 3/03, 7/03, 3/04 7 201

Favorite Channel (51) 

eelgrass 58.4309 , 134.7648 7/05 1 1,470

kelp 58.5031 , 134.8669 4/01, 8/01, 3/02, 7/02, 3/03, 6/03, 3/04 7 215

kelp 58.5019 , 134.8664 4/01, 8/01, 3/02, 7/02, 3/03, 6/03, 3/04 7 1,405

sand 58.5131 , 134.8419 4/01, 8/01, 3/02, 7/02, 3/03, 6/03, 3/04 7 934

sand 58.5119 , 134.8356 4/01, 8/01, 3/02, 7/02, 3/03, 6/03, 3/04 7 140

sand 58.5053 , 134.8106 4/01, 8/01 2 87

sand 58.5008 , 134.7994 4/01, 8/01 2 92

Auke Bay (52) 

eelgrass 58.3853 , 134.6517 6/05 1 328

eelgrass 58.3819 , 134.6603 6/05, 8/06 2 2,097

eelgrass 58.3811 , 134.6924 4/99, 5/99, 6/99, 7/99 4 4,739

eelgrass 58.3809 , 134.6918 6/05, 8/06 3 1,090

eelgrass 58.3801 , 134.6918 8/06 1 364

eelgrass 58.3756 , 134.7255 7/10 1 218

sand 58.3777 , 134.7200 7/10 1 29

Peterson Creek (53) 

eelgrass 58.3018 , 134.6813 6/05 2 54

Funter Bay (54) 

bedrock 58.2436 , 134.9178 8/98 1 1

bedrock 58.2336 , 134.9156 8/98 1 216

254
Appendix -- (Cont.). 

Region, Location, Site, Coordinates Dates sampled Hauls Catch

bedrock 58.2311 , 134.9203 8/98 1 3

bedrock 58.2283 , 134.9164 8/98 1 15

eelgrass 58.2569 , 134.9003 8/99, 6/01, 6/02, 2/03, 6/03, 6/08, 6/09, 6/10 8 1,482

eelgrass 58.2553 , 134.9106 8/98 1 2,088

eelgrass 58.2553 , 134.9097 8/99, 6/01, 6/02, 2/03, 6/03, 6/08, 6/10 7 1,105

eelgrass 58.2550 , 134.9058 8/98 1 65

kelp 58.2550 , 134.8881 8/99 1 42

kelp 58.2503 , 134.8953 8/99 1 19

sand 58.2495 , 134.9075 8/98 1 9

sand 58.2483 , 134.9081 8/98 1 40

Couverden (55) 

eelgrass 58.2103 , 135.1178 6/08 1 886

kelp 58.1992 , 135.1045 6/08 1 121

Neka Bay (56) 

eelgrass 58.0500 , 135.6725 8/99 1 89

eelgrass 58.0497 , 135.6706 8/99, 6/08 2 684

eelgrass 57.9894 , 135.5976 6/08 1 183

kelp 58.0522 , 135.6458 8/99 1 40

kelp 58.0328 , 135.6422 8/99, 6/08 2 51

Tenakee Inlet (57) 

bedrock 57.8734 , 135.5983 6/06 1 128

eelgrass 57.9445 , 135.7302 6/06 1 294

eelgrass 57.8936 , 135.6225 6/06 1 157

eelgrass 57.8714 , 135.6157 6/06 1 188

sand 57.9171 , 135.6881 6/06 1 1,155

sand 57.8716 , 135.5700 6/06 1 86

Tenakee Springs (58) 

bedrock 57.7789 , 135.1319 8/98 1 1

bedrock 57.7775 , 135.1272 8/98 1 2

bedrock 57.7766 , 135.1565 6/06 1 1

bedrock 57.7742 , 135.1981 8/98 1 0

bedrock 57.7742 , 135.1981 8/98 1 10

bedrock 57.7550 , 135.1129 6/06 1 809

eelgrass 57.7758 , 135.1911 8/98 1 119

eelgrass 57.7756 , 135.1914 8/98 1 81

eelgrass 57.7387 , 135.2709 6/06 1 815

eelgrass 57.7367 , 135.3836 6/01, 6/02, 1/03, 6/03, 6/08, 6/09, 6/10 7 4,207

eelgrass 57.7364 , 135.3875 6/01, 6/02, 1/03, 6/03, 6/08, 6/09, 6/10 7 1,367

255
Appendix -- (Cont.). 

Region, Location, Site, Coordinates Dates sampled Hauls Catch

kelp 57.7775 , 135.1560 6/06 1 1,110

kelp 57.7390 , 135.1819 6/06 1 52

sand 57.7798 , 135.1562 6/06 1 11,076

sand 57.7764 , 135.1817 8/98 1 47

sand 57.7758 , 135.1828 8/98 1 42

Hoonah Sound (59) 

eelgrass 57.7597 , 135.7931 8/02 1 438

eelgrass 57.7431 , 135.7931 8/02 1 562

eelgrass 57.6166 , 135.6247 6/08 1 727

kelp 57.6131 , 135.5884 6/08 1 138

Ushk Bay (60) 

bedrock 57.5675 , 135.6381 8/98 1 5

bedrock 57.5667 , 135.6256 8/98 1 10,124

bedrock 57.5667 , 135.6255 8/98 1 124

bedrock 57.5647 , 135.6106 8/98 1 3

eelgrass 57.5705 , 135.6400 8/02, 2/03, 5/03, 8/03 4 5,381

eelgrass 57.5656 , 135.6544 8/98 1 140

eelgrass 57.5628 , 135.6572 8/98 1 92

eelgrass 57.5618 , 135.6116 8/02, 2/03, 5/03, 8/03 4 48,520

sand 57.5617 , 135.6606 8/98 1 44

sand 57.5617 , 135.6617 8/98 1 33

Sitkoh Bay (61) 

bedrock 57.5108 , 134.9272 8/98 1 137

bedrock 57.5050 , 134.9269 8/98 1 656

bedrock 57.5031 , 134.9111 8/98 1 18

bedrock 57.4922 , 134.8919 8/98 1 5

eelgrass 57.5325 , 134.9761 8/98 1 97

eelgrass 57.5303 , 134.9767 8/98 1 742

sand 57.5264 , 134.9675 8/98 1 33

sand 57.5253 , 134.9669 8/98 1 1,052

Kelp Bay (62) 

kelp 57.2697 , 134.8683 6/00 1 2,026

kelp 57.2656 , 134.8858 6/00 1 433

Cosmos Cove (63) 

eelgrass 57.2428 , 134.8811 6/00 1 567

eelgrass 57.2425 , 134.8717 6/00 1 414

Chaik Bay (64) 

eelgrass 57.3142 , 134.4711 6/01, 6/02, 6/03 3 26,721

eelgrass 57.3133 , 134.4728 6/01, 6/02, 6/03 3 14,614

256
Appendix -- (Cont.). 

Region, Location, Site, Coordinates Dates sampled Hauls Catch

Pybus Bay (65) 

eelgrass 57.3872 , 134.1761 6/00 1 242

eelgrass 57.3764 , 134.1836 6/00 1 77

kelp 57.3672 , 134.1378 6/00 1 260

kelp 57.3378 , 134.1222 6/00 1 68

Brothers Islands (66) 

bedrock 57.3037 , 133.8433 7/01, 7/02, 3/03, 7/03, 3/04 5 2,049

bedrock 57.3033 , 133.8630 3/02, 7/02, 3/03, 7/03, 8/03, 3/04, 6/11 7 12,456

bedrock 57.3026 , 133.8045 7/01, 3/02, 7/02, 7/03, 3/04 5 32,855

bedrock 57.2987 , 133.7964 3/02, 7/02, 7/03, 8/03, 3/04 5 23,458

bedrock 57.2910 , 133.7950 3/03 1 2

bedrock 57.2829 , 133.8417 3/02, 7/02, 3/03, 7/03, 3/04 5 6,979

eelgrass 57.2928 , 133.8150 7/01, 3/02, 7/02, 3/03, 7/03, 8/03, 3/04, 6/11 8 12,679

kelp 57.3092 , 133.8161 7/01, 7/02, 3/03, 7/03, 8/03, 3/04, 6/11 7 35,528

kelp 57.3086 , 133.8314 7/01, 2/02, 7/02, 3/03, 7/03, 8/03, 3/04 7 9,510

kelp 57.3078 , 133.8497 7/01, 2/02, 7/02, 3/03, 7/03, 3/04 6 6,486

kelp 57.3036 , 133.8603 7/01, 2/02, 7/02, 3/03, 7/03, 3/04 6 13,812

kelp 57.2997 , 133.8250 7/01, 3/02, 7/02, 3/03, 7/03, 3/04 6 2,366

kelp 57.2894 , 133.8633 7/01, 3/02, 7/02, 3/03, 7/03, 3/04 6 25,595

sand 57.2989 , 133.8233 7/01, 3/02, 7/02, 3/03, 7/03, 8/03, 3/04 7 6,989

sand 57.2958 , 133.8656 7/01, 3/02, 7/02, 3/03, 7/03, 8/03, 3/04, 6/11 8 7,002

Gambier Bay (67) 

bedrock 57.4235 , 133.8961 6/11 1 18

kelp 57.4372 , 133.9068 6/11 1 1,345

kelp 57.4322 , 133.8924 6/11 1 799

Hobart Bay (68) 

kelp 57.4233 , 133.4525 5/00 1 14

kelp 57.4222 , 133.4536 5/00 1 19

Seymour Canal (69) 

bedrock 57.7468 , 134.0170 6/11 1 33

eelgrass 57.7299 , 133.9961 6/11 1 698

kelp 57.7190 , 133.9986 6/11 1 211

Taku Harbor (70) 

kelp 58.0714 , 134.0232 6/11 1 7

sand 58.0645 , 134.0142 6/11 1 22

sand 58.0631 , 134.0240 6/11 1 30

sand 58.0612 , 134.0265 6/11 1 22

257
Appendix -- (Cont.). 

Region, Location, Site, Coordinates Dates sampled Hauls Catch

Gastineau Channel (71) 

kelp 58.2047 , 134.2155 7/05 1 395

kelp 58.1926 , 134.2638 7/05 1 296

sand 58.2282 , 134.2659 7/05 1 446

Southeastern - Southern Outside 

Port Conclusion (72) 

kelp 56.2553 , 134.6633 7/00 1 119

kelp 56.2536 , 134.6653 7/00 1 35

Shakan Bay (73) 

eelgrass 56.1950 , 133.5167 7/00 1 298

eelgrass 56.1936 , 133.5169 7/00 1 301

kelp 56.1483 , 133.5378 7/00 1 39

kelp 56.1453 , 133.5408 7/00 1 38

Warm Chuck Inlet (74) 

eelgrass 55.7758 , 133.5319 7/99 1 265

eelgrass 55.7708 , 133.5361 7/99 1 629

eelgrass 55.7581 , 133.4708 7/99 1 370

eelgrass 55.7564 , 133.4711 7/99 1 433

Klawock Inlet (75) 

bedrock 55.5289 , 133.1900 5/98 1 3,564

bedrock 55.5258 , 133.1407 6/98 1 235

bedrock 55.5186 , 133.1489 5/98 1 255

eelgrass 55.5778 , 133.0941 4/98, 5/98, 6/98, 9/98 4 4,145

eelgrass 55.5772 , 133.0944 4/98, 5/98, 6/98, 9/98 4 1,030

eelgrass 55.5361 , 133.1056 4/98, 5/98, 6/98, 9/98, 7/99 5 1,856

eelgrass 55.5358 , 133.1053 4/98, 5/98, 6/98, 9/98, 7/99 5 1,323

eelgrass 55.5208 , 133.1617 5/98 1 116

eelgrass 55.5003 , 133.1647 4/98, 5/98, 6/98, 9/98, 7/99 5 2,475

eelgrass 55.5000 , 133.1644 4/98, 5/98, 6/98, 9/98, 7/99 5 3,002

eelgrass 55.4875 , 133.1414 4/98, 5/98, 6/98, 9/98 4 2,162

eelgrass 55.4872 , 133.1414 4/98, 5/98, 6/98, 9/98 4 3,443

eelgrass 55.4764 , 133.1881 5/98 1 21

kelp 55.5125 , 133.1297 5/98, 6/98, 9/98, 7/99 4 302

kelp 55.5072 , 133.1339 5/98, 6/98, 9/98, 7/99 4 267

kelp 55.5011 , 133.1675 4/98, 5/98, 6/98, 9/98, 7/99 5 1,791

kelp 55.5008 , 133.1672 4/98, 5/98, 6/98, 9/98, 7/99 5 470

sand 55.5622 , 133.0994 4/98, 5/98, 6/98, 9/98 4 342

sand 55.5619 , 133.0997 4/98, 5/98, 6/98, 9/98 4 1,503

258
Appendix -- (Cont.). 

Region, Location, Site, Coordinates Dates sampled Hauls Catch

sand 55.5125 , 133.1297 4/98, 5/98, 6/98, 9/98 4 307

sand 55.5072 , 133.1339 4/98, 5/98, 6/98, 9/98 4 258

sand 55.4819 , 133.1881 5/98 1 85

Port San Antonio (76) 

eelgrass 55.3658 , 133.5856 7/99 1 310

eelgrass 55.3575 , 133.5869 7/99 1 105

kelp 55.3597 , 133.5883 7/99 1 85

kelp 55.3597 , 133.5878 7/99 1 91

Nichols Bay (77) 

bedrock 54.7211 , 132.1257 6/07 1 5

eelgrass 54.7401 , 132.1536 6/07 1 1,781

kelp 54.7259 , 132.1241 6/07 1 74

kelp 54.7051 , 132.1005 6/07 1 146

Moira Sound (78) 

bedrock 54.9869 , 132.1050 6/07 1 1,028

eelgrass 54.9711 , 132.0924 6/07 1 254

kelp 54.9856 , 132.1098 6/07 1 121

Dall Bay (79) 

bedrock 55.1562 , 131.7453 6/07 1 217

eelgrass 55.1561 , 131.7498 6/07 1 686

kelp 55.1551 , 131.7421 6/07 1 64

Bostwick Inlet (80) 

eelgrass 55.2364 , 131.7500 8/00 1 509

eelgrass 55.2342 , 131.7300 8/00 1 130

kelp 55.2200 , 131.7361 8/00 1 592

kelp 55.2078 , 131.7278 8/00 1 107

Sylburn Harbor (81) 

bedrock 55.1872 , 131.5824 6/07 1 31

eelgrass 55.1782 , 131.5791 6/07 1 267

kelp 55.1760 , 131.5972 6/07 1 153

Reef Harbor (82) 

bedrock 54.9776 , 131.2560 6/07 1 451

eelgrass 54.9684 , 131.2492 6/07 1 550

kelp 54.9744 , 131.2538 6/07 1 27

Kah Shakes Cove (83) 

bedrock 55.0479 , 130.9951 6/07 1 53

eelgrass 55.0426 , 130.9995 6/07 1 84

kelp 55.0635 , 130.9917 6/07 1 182

kelp 55.0470 , 131.0105 6/07 1 190

259
Appendix -- (Cont.). 

Region, Location, Site, Coordinates Dates sampled Hauls Catch

Southeastern - Southern Inside

Farragut Bay (84) 

kelp 57.1244 , 133.2053 6/00 1 198

kelp 57.1144 , 133.2050 6/00 1 58

Port Camden (85) 

kelp 56.7160 , 133.9523 6/00 1 177

kelp 56.6525 , 133.9569 6/00 1 5,235

Kah Sheets Bay (86) 

eelgrass 56.5183 , 133.0969 8/00 1 157

eelgrass 56.5167 , 133.0958 8/00 1 203

kelp 56.5153 , 133.1208 8/00 1 68

kelp 56.5119 , 133.1269 8/00 1 43

Exchange Cove (87) 

eelgrass 56.2478 , 133.0781 7/00 1 51

eelgrass 56.2111 , 133.0683 7/00 1 281

kelp 56.2400 , 133.0719 7/00 1 24

kelp 56.2336 , 133.0733 7/00 1 42

Steamer Bay (88) 

eelgrass 56.1575 , 132.6978 7/00 1 96

eelgrass 56.1531 , 132.6911 7/00 1 190

kelp 56.1625 , 132.7100 7/00 1 54

kelp 56.1581 , 132.6967 7/00 1 51

Zimovia Strait (89) 

eelgrass 56.1917 , 132.3033 8/00 1 196

eelgrass 56.1886 , 132.3164 8/00 1 1,010

kelp 56.2269 , 132.3778 8/00 1 71

kelp 56.2236 , 132.3825 8/00 1 40

Twelve Mile Arm (90) 

eelgrass 55.4283 , 132.6594 8/00 1 1,517

eelgrass 55.4156 , 132.6881 8/00 1 515

kelp 55.4486 , 132.6539 8/00 1 73

kelp 55.4286 , 132.6594 8/00 1 106

Cholmondeley Sound (91) 

bedrock 55.2516 , 132.3874 6/07 1 53

eelgrass 55.2541 , 132.4418 6/07 1 240

kelp 55.2515 , 132.3815 6/07 1 160

Clover Passage (92) 

bedrock 55.5599 , 131.7041 6/07 1 13

eelgrass 55.5446 , 131.7032 6/07 1 1,013

kelp 55.5565 , 131.7058 6/07 1 312

260
Appendix -- (Cont.). 

Region, Location, Site, Coordinates Dates sampled Hauls Catch

Thorne Arm (93) 

bedrock 55.3572 , 131.2667 6/07 1 2

bedrock 55.3190 , 131.2580 6/07 1 7

eelgrass 55.3868 , 131.2886 6/07 1 900

kelp 55.3209 , 131.2566 6/07 1 105

261

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