Appendix D 

Species List 
Appendix D 

Alaska Peninsula and Becharof NWR Comprehensive Conservation Plan D-1 

Species List 

This appendix includes a list of species for the Alaska Peninsula and Becharof National Wildlife 

Refuges. The table uses a number of abbreviations, which are defined as follows: 

? = possible/may be probable 

BB = Bristol Bay 

BRD = Breeder 

WTR = Winter resident 

MAR BRD = Marginal breeder; for example, found in 

the boreal in King Salmon and Katmai. Known boreal 

area is on edge of Becharof National Wildlife Refuge 

(NWR) (Big Creek drainage), so species probably 

found there, but on edge of range. 

RES = Resident (year-round) 

MIG = Migrant (a regular user during the spring or 

fall migration different from TRAN or INC) 

WTR = Winter 

O = Occasional 

NSHR = Nearshore; i.e., found on offshore islands (for 

example, belonging to the Alaska Maritime National 

Wildlife Refuge) or in ocean waters near shore, but not 

known to use “above mean high tide” on the Refuges 

ACC = Accidental: A species where the breeding and 

wintering ranges do not overlap and may be distant 

from Refuge lands, but where individuals of the species 

have been documented on the Refuges. Note that many 

of the ACC are Asiatic species. 
Appendix D: Species List 

D-2 Alaska Peninsula and Becharof NWR Compreh

Common Name Scientific Name Class/Order/Family 

FISHES—RESIDENT Arctic lamprey Lampetra japonica Cephalaspidomorphi/Petromyzontiformes/ 

Petromyzontidae 

Fr

Longnose sucker Catostomus catostomus Actinopterygii/Cypriniformes/Castomidae FrLeast Cisco Coregonus sardinella Actinopterygii/Salmoniformes/Salmonidae FrLake Whitefish Coregonus clupeaformis Actinopterygii/Salmoniformes/Salmonidae FrPygmy Whitefish Prosopium coulteri Actinopterygii/Salmoniformes/Salmonidae FrRound Whitefish Prosopium cylindraceum Actinopterygii/Salmoniformes/Salmonidae FrLake Trout Salvelinus namaycush Actinopterygii/Salmoniformes/Salmonidae FrArctic Char Salvelinus alpinus Actinopterygii/Salmoniformes/Salmonidae FrDolly Varden Salvelinus malma Actinopterygii/Salmoniformes/Salmonidae FrRainbow Trout/Steelhead Oncorhynchus mykiss Actinopterygii/Salmoniformes/Salmonidae FrArctic Grayling Thymallus arcticus Actinopterygii/Salmoniformes/Salmonidae FrPond Smelt Hypomesus olidus Actinopterygii/Osmeriformes/Osmeridae FrSlimy Sculpin Cottus cognatus Actinopterygii/Scorpaeniformes/Cottidae FrCoastrange Sculpin Cottus aleuticus Actinopterygii/Scorpaeniformes/Cottidae FrAlaska Blackfish Dallia pectoralis Actinopterygii/Esociformes/Umbridae FrNorthern Pike Esox lucius Actinopterygii/Esociformes/Esocidae FrBurbot Lota lota Actinopterygii/Gadiformes/Lotidae FrThree-spined Stickleback Gasterosteus aculeatus Actinopterygii/Gasterosteiformes/Gasterosteidae Fr

/M

Nine-spined Stickleback Pungitius pungitius Actinopterygii/Gasterosteiformes/Gasterosteidae FrFISHES—ANADROMOUS Pink Salmon Oncorhynchus gorbuscha Actinopterygii/Salmoniformes/Salmonidae ASockeye Salmon Oncorhynchus nerka Actinopterygii/Salmoniformes/Salmonidae AChinook Salmon Oncorhynchus tschawytscha Actinopterygii/Salmoniformes/Salmonidae ACoho Salmon Oncorhynchus kisutch Actinopterygii/Salmoniformes/Salmonidae AChum Salmon Oncorhynchus keta Actinopterygii/Salmoniformes/Salmonidae AEulachon Thaleichthys pacificus Actinopterygii/Osmeriformes/Osmeridae ARainbow Smelt Osmerus mordax Actinopterygii/Osmeriformes/Osmeridae APacific Lamprey Entosphenus tridentatus Cephalaspidomorphi/Petromyzontiformes/ 

Petromyzontidae 

A
A

Alaska Peninsula and Becharof NWR Comprehensive Conservation Plan 

Common Name Scientific Name Class/Order/Family 

FISHES—MARINE Pacific Herring Clupea harengus pallasii Actinopterygii/Cluperiformes/Cluperidae MSurf Smelt Hypomesus pretiosus Actinopterygii/Osmeriformes/Osmeridae MSaffron Cod Eleginu gracilis Actinopterygii/Gadiformes/Gadidae M

r 

Pacific Staghorn Sculpin Leptocottus armatus Actinopterygii/Scorpaeniformes/Cottidae BSharpnose Sculpin Clinocotus acuticeps Actinopterygii/Scorpaeniformes/Cottidae M

r 

Starry Flounder Platichthys stellatus Actinopterygii/Pleuronectiformes/Pleuronectidae M

r 

Fourhorn Sculpin Myoxocephalus quadricornis Actinopterygii/Scorpaeniformes/Cottidae MAMPHIBIAN Wood Frog Rana sylvatica Amphibia/Anura/Ranidae REBIRDS—BREEDING or RESIDENT 134 TOTAL Greater White-fronted Goose Anser albifrons Aves/Anseriformes/Anatidae BRCackling Goose Branta huchinsoni Aves/Anseriformes/Anatidae BRTundra Swan Cygnus columbianus Aves/Anseriformes/Anatidae BRGadwall Anas strepera Aves/Anseriformes/Anatidae BRAmerican Wigeon Anas americana Aves/Anseriformes/Anatidae BRMallard Anas platyrhynchos Aves/Anseriformes/Anatidae BRNorthern Shoveler Anas clypeata Aves/Anseriformes/Anatidae BRNorthern Pintail Anas acuta Aves/Anseriformes/Anatidae BRGreen-winged Teal Anas crecca Aves/Anseriformes/Anatidae BRGreater Scaup Aythya marila Aves/Anseriformes/Anatidae BRCommon Eider Somateria mollissima Aves/Anseriformes/Anatidae BRHarlequin Duck Histrionicus histrionicus Aves/Anseriformes/Anatidae BRBlack Scoter Melanitta nigra Aves/Anseriformes/Anatidae BRLong-tailed Duck Clangula hyemalis Aves/Anseriformes/Anatidae BR
Appendix D: Species List 

D-4 Alaska Peninsula and Becharof NWR Compre

Common Name Scientific Name Class/Order/Family 

Common Merganser Mergus merganser Aves/Anseriformes/Anatidae BRed-breasted Merganser Mergus serrator Aves/Anseriformes/Anatidae BSpruce Grouse Falcipennis canadensis Aves/Galliformes/Phasianidae RWillow Ptarmigan Lagopus lagopus Aves/Galliformes/Phasianidae RRock Ptarmigan Lagopus muta Aves/Galliformes/Phasianidae RWhite-tailed Ptarmigan Lagopus leucura Aves/Galliformes/Phasianidae RRed-throated Loon Gavia stellata Aves/Gaviiformes/Gaviidae BPacific Loon Gavia pacifica Aves/Gaviiformes/Gaviidae BCommon Loon Gavia immer Aves/Gaviiformes/Gaviidae BRed-necked Grebe Podiceps grisegena Aves/Podicipediformes/Podicipedidae BDouble-crested Cormorant Phalacrocorax auritus Aves/Pelecaniformes/Phalacrocoracidae BRed-faced Cormorant Phalacrocorax urile Aves/Pelecaniformes/Phalacrocoracidae BPelagic Cormorant Phalacrocorax pelagicus Aves/Pelecaniformes/Phalacrocoracidae BOsprey Pandion haliaetus Aves/Falconiformes/Accipitridae BBald Eagle Haliaeetus leucocephalus Aves/Falconiformes/Accipitridae RNorthern Harrier Circus cyaneus Aves/Falconiformes/Accipitridae BNorthern Goshawk Accipiter gentilis Aves/Falconiformes/Accipitridae RRough-legged Hawk Buteo lagopus Aves/Falconiformes/Accipitridae BGolden Eagle Aquila chrysaetos Aves/Falconiformes/Accipitridae RMerlin Falco columbarius Aves/Falconiformes/Falconidae BGyrfalcon Falco rusticolus Aves/Falconiformes/Falconidae RPeregrine Falcon Falco peregrinus Aves/Falconiformes/Falconidae BSandhill Crane Grus canadensis Aves/Gruiiformes/Gruidae BBlack-bellied Plover Pluvialis squatarola Aves/Charadriiformes/Charadriidae BAmerican Golden-Plover Pluvialis dominica Aves/Charadriiformes/Charadriidae BPacific Golden-Plover Pluvialis fulva Aves/Charadriiformes/Charadriidae BSemipalmated Plover Charadrius semipalmatus Aves/Charadriiformes/Charadriidae BBlack Oystercatcher Haematopus bachmani Aves/Charadriiformes/Haematopodidae RGreater Yellowlegs Tringa melanoleuca Aves/Charadriiformes/Scolopacidae BLesser Yellowlegs Tringa flavipes Aves/Charadriiformes/Scolopacidae BSolitary Sandpiper Tringa solitaria Aves/Charadriiformes/Scolopacidae B
A

Alaska Peninsula and Becharof NWR Comprehensive Conservation Plan 

Common Name Scientific Name Class/Order/Family 

Wandering Tattler Heteroscelus incanus Aves/Charadriiformes/Scolopacidae BSpotted Sandpiper Actitis macularius Aves/Charadriiformes/Scolopacidae BHudsonian Godwit Limosa haemastica Aves/Charadriiformes/Scolopacidae BMarbled Godwit Limosa fedoa Aves/Charadriiformes/Scolopacidae BBlack Turnstone Arenaria melanocephala Aves/Charadriiformes/Scolopacidae BSurfbird Aphriza virgata Aves/Charadriiformes/Scolopacidae BWestern Sandpiper Calidris mauri Aves/Charadriiformes/Scolopacidae BLeast Sandpiper Calidris minutilla Aves/Charadriiformes/Scolopacidae BRock Sandpiper Calidris ptilocnemis Aves/Charadriiformes/Scolopacidae BDunlin Calidris alpina Aves/Charadriiformes/Scolopacidae BShort-billed Dowitcher Limnodromus griseus Aves/Charadriiformes/Scolopacidae BWilson's Snipe Gallinago delicata Aves/Charadriiformes/Scolopacidae BRed-necked Phalarope Phalaropus lobatus Aves/Charadriiformes/Scolopacidae BParasitic Jaeger Stercorarius parasiticus Aves/Charadriiformes/Laridae BLong-tailed Jaeger Stercorarius longicaudus Aves/Charadriiformes/Laridae BBonaparte's Gull Larus philadelphia Aves/Charadriiformes/Laridae BMew Gull Larus canus Aves/Charadriiformes/Laridae BGlaucous-winged Gull Larus glaucescens Aves/Charadriiformes/Laridae BSabine's Gull Xema sabini Aves/Charadriiformes/Laridae BBlack-legged Kittiwake Rissa tridactyla Aves/Charadriiformes/Laridae BArctic Tern Sterna paradisaea Aves/Charadriiformes/Laridae BAleutian Tern Sterna aleutica Aves/Charadriiformes/Laridae BCommon Murre Uria aalge Aves/Charadriiformes/Alcidae BThick-billed Murre Uria lomvia Aves/Charadriiformes/Alcidae BPigeon Guillemot Cepphus columba Aves/Charadriiformes/Alcidae BMarbled Murrelet Brachyramphus marmoratus Aves/Charadriiformes/Alcidae BKittlitz's Murrelet Brachyramphus brevirostris Aves/Charadriiformes/Alcidae BAncient Murrelet Synthliboramphus antiquus Aves/Charadriiformes/Alcidae BHorned Puffin Fratercula corniculata Aves/Charadriiformes/Alcidae BTufted Puffin Fratercula cirrhata Aves/Charadriiformes/Alcidae BGreat Horned Owl Bubo virginianus Aves/Strigiformes/Strigidae R
Appendix D: Species List 

D-6 Alaska Peninsula and Becharof NWR Compre

Common Name Scientific Name Class/Order/Family 

Northern Hawk Owl Surnia ulula Aves/Strigiformes/Strigidae BShort-eared Owl Asio flammeus Aves/Strigiformes/Strigidae BBoreal Owl Aegolius funereus Aves/Strigiformes/Strigidae BNorthern Saw-whet Owl Aegolius acadicus Aves/Strigiformes/Strigidae BBelted Kingfisher Ceryle alcyon Aves/Coraciiformes/Alcedinidae RDowny Woodpecker Picoides pubescens Aves/Piciformes/Picidae RAmerican Three-toed Woodpecker Picoides dorsalis Aves/Piciformes/Picidae BNorthern Flicker Colaptes auratus Aves/Piciformes/Picidae BOlive-sided Flycatcher Contopus cooperi Aves/Passeriformes/Tyrannidae BAlder Flycatcher Empidonax alnorum Aves/Passeriformes/Tyrannidae BNorthern Shrike Lanius excubitor Aves/Passeriformes/Laniidae RGray Jay Perisoreus canadensis Aves/Passeriformes/Corvidae RBlack-billed Magpie Pica hudsonia Aves/Passeriformes/Corvidae RNorthwestern Crow Corvus caurinus Aves/Passeriformes/Corvidae BCommon Raven Corvus corax Aves/Passeriformes/Corvidae RHorned Lark Eremophila alpestris Aves/Passeriformes/Alaudidae BTree Swallow Tachycineta bicolor Aves/Passeriformes/Hirundinidae BViolet-green Swallow Tachycineta thalassina Aves/Passeriformes/Hirundinidae BBank Swallow Riparia riparia Aves/Passeriformes/Hirundinidae BCliff Swallow Petrochelidon pyrrhonota Aves/Passeriformes/Hirundinidae BBarn Swallow Hirundo rustica Aves/Passeriformes/Hirundinidae BBlack-capped Chickadee Poecile atricapillus Aves/Passeriformes/Paridae RBoreal Chickadee Poecile hudsonica Aves/Passeriformes/Paridae RRed-breasted Nuthatch Sitta canadensis Aves/Passeriformes/Sittidae BBrown Creeper Certhia americana Aves/Passeriformes/Certhiidae BWinter Wren Troglodytes troglodytes Aves/Passeriformes/Troglodytidae RAmerican Dipper Cinclus mexicanus Aves/Passeriformes/Cinclidae BRuby-crowned Kinglet Regulus calendula Aves/Passeriformes/Regulidae BArctic Warbler Phylloscopus borealis Aves/Passeriformes/Sylviidae BGray-cheeked Thrush Catharus minimus Aves/Passeriformes/Turdidae BSwainson's Thrush Catharus ustulatus Aves/Passeriformes/Turdidae B
A

Alaska Peninsula and Becharof NWR Comprehensive Conservation Plan 

Common Name Scientific Name Class/Order/Family 

Hermit Thrush Catharus guttatus Aves/Passeriformes/Turdidae BAmerican Robin Turdus migratorius Aves/Passeriformes/Turdidae BVaried Thrush Ixoreus naevius Aves/Passeriformes/Turdidae BEastern Yellow Wagtail Motacilla tschutschensis Aves/Passeriformes/Motacillidae BAmerican Pipit Anthus rubescens Aves/Passeriformes/Motacillidae BOrange-crowned Warbler Vermivora celata Aves/Passeriformes/Parulidae BYellow Warbler Dendroica petechia Aves/Passeriformes/Parulidae BYellow-rumped Warbler Dendroica coronata Aves/Passeriformes/Parulidae BBlackpoll Warbler Dendroica striata Aves/Passeriformes/Parulidae BNorthern Waterthrush Seiurus noveboracensis Aves/Passeriformes/Parulidae BWilson's Warbler Wilsonia pusilla Aves/Passeriformes/Parulidae BAmerican Tree Sparrow Spizella arborea Aves/Passeriformes/Emberizidae BSavannah Sparrow Passerculus sandwichensis Aves/Passeriformes/Emberizidae BFox Sparrow Passerella iliaca Aves/Passeriformes/Emberizidae BSong Sparrow Melospiza melodia Aves/Passeriformes/Emberizidae RLincoln's Sparrow Melospiza lincolnii Aves/Passeriformes/Emberizidae BWhite-crowned Sparrow Zonotrichia leucophrys Aves/Passeriformes/Emberizidae BGolden-crowned Sparrow Zonotrichia atricapilla Aves/Passeriformes/Emberizidae BDark-eyed Junco Junco hyemalis Aves/Passeriformes/Emberizidae BLapland Longspur Calcarius lapponicus Aves/Passeriformes/Emberizidae BSnow Bunting Plectrophenax nivalis Aves/Passeriformes/Emberizidae BRusty Blackbird Euphagus carolinus Aves/Passeriformes/Icteridae BGray-crowned Rosy-Finch Leucosticte tephrocotis Aves/Passeriformes/Fringillidae BPine Grosbeak Pinicola enucleator Aves/Passeriformes/Fringillidae RWhite-winged Crossbill Loxia leucoptera Aves/Passeriformes/Fringillidae BCommon Redpoll Carduelis flammea Aves/Passeriformes/Fringillidae RPine Siskin Carduelis pinus Aves/Passeriformes/Fringillidae BBIRDS—WINTER or MIGRATION 38 Species Emperor Goose Chen canagica Aves/Anseriformes/Anatidae MSnow Goose Chen caerulescens Aves/Anseriformes/Anatidae MBrant Branta bernicla Aves/Anseriformes/Anatidae M
Appendix D: Species List 

D-8 Alaska Peninsula and Becharof NWR Compre

Common Name Scientific Name Class/Order/Family 

Canada Goose Branta canadensis parvipes Aves/Anseriformes/Anatidae MEurasian Wigeon Anas penelope Aves/Anseriformes/Anatidae MCanvasback Aythya valisineria Aves/Anseriformes/Anatidae MRedhead Aythya americana Aves/Anseriformes/Anatidae MSteller's Eider Polysticta stelleri Aves/Anseriformes/Anatidae MSpectacled Eider Somateria fischeri Aves/Anseriformes/Anatidae WKing Eider Somateria spectabilis Aves/Anseriformes/Anatidae MBufflehead Bucephala albeola Aves/Anseriformes/Anatidae WCommon Goldeneye Bucephala clangula Aves/Anseriformes/Anatidae WBarrow's Goldeneye Bucephala islandica Aves/Anseriformes/Anatidae WYellow-billed Loon Gavia adamsii Aves/Gaviiformes/Gaviidae MHorned Grebe Podiceps auritus Aves/Podicipediformes/Podicipedidae MSharp-shinned Hawk Accipiter striatus Aves/Falconiformes/Accipitridae MWhimbrel Numenius phaeopus Aves/Charadriiformes/Scolopacidae MBristle-thighed Curlew Numenius tahitiensis Aves/Charadriiformes/Scolopacidae MBar-tailed Godwit Limosa lapponica Aves/Charadriiformes/Scolopacidae MRuddy Turnstone Arenaria interpres Aves/Charadriiformes/Scolopacidae MRed Knot Calidris canutus Aves/Charadriiformes/Scolopacidae MSanderling Calidris alba Aves/Charadriiformes/Scolopacidae MSemipalmated Sandpiper Calidris pusilla Aves/Charadriiformes/Scolopacidae MBaird's Sandpiper Calidris bairdii Aves/Charadriiformes/Scolopacidae MPectoral Sandpiper Calidris melanotos Aves/Charadriiformes/Scolopacidae MBuff-breasted Sandpiper Tryngites subruficollis Aves/Charadriiformes/Scolopacidae MLong-billed Dowitcher Limnodromus scolopaceus Aves/Charadriiformes/Scolopacidae MRed Phalarope Phalaropus fulicaria Aves/Charadriiformes/Scolopacidae MPomarine Jaeger Stercorarius pomarinus Aves/Charadriiformes/Laridae MHerring Gull Larus argentatus Aves/Charadriiformes/Laridae MGlaucous Gull Larus hyperboreus Aves/Charadriiformes/Laridae MSnowy Owl Bubo scandiaca Aves/Strigiformes/Strigidae WSay's Phoebe Sayornis saya Aves/Passeriformes/Tyrannidae MNorthern Rough-winged swallow Stelgidopteryx ruficollis Aves/Passeriformes/Hirundinidae M
A

Alaska Peninsula and Becharof NWR Comprehensive Conservation Plan 

Common Name Scientific Name Class/Order/Family 

Golden-crowned Kinglet Regulus satrapa Aves/Passeriformes/Regulidae MTownsend's Warbler Dendroica townsendi Aves/Passeriformes/Parulidae MMcKay's Bunting Plectrophenax hyperboreus Aves/Passeriformes/Emberizidae MHoary Redpoll Carduelis hornemanni Aves/Passeriformes/Fringillidae MBIRDS—NEARSHORE 13 Species Surf Scoter Melanitta perspicillata Aves/Anseriformes/Anatidae NWhite-winged Scoter Melanitta fusca Aves/Anseriformes/Anatidae NNorthern Fulmar Fulmarus glacialis Aves/Procellariformes/Procellariidae NSooty Shearwater Puffinus griseus Aves/Procellariformes/Procellariidae NShort-tailed Shearwater Puffinus tenuirostris Aves/Procellariformes/Procellariidae NFork-tailed Storm-Petrel Oceanodroma furcata Aves/Procellariformes/Hydrobatidae NLeach's Storm-Petrel Oceanodroma leucorhoa Aves/Procellariformes/Hydrobatidae NCassin's Auklet Ptychoramphus aleuticus Aves/Charadriiformes/Alcidae NParakeet Auklet Aethia psittacula Aves/Charadriiformes/Alcidae NLeast Auklet Aethia pusilla Aves/Charadriiformes/Alcidae NWhiskered Auklet Aethia pygmaea Aves/Charadriiformes/Alcidae NCrested Auklet Aethia cristatella Aves/Charadriiformes/Alcidae NRhinoceros Auklet Cerorhinca monocerata Aves/Charadriiformes/Alcidae NBIRDS—ACCIDENTAL 29 Species Trumpeter Swan Cygnus buccinator Aves/Anseriformes/Anatidae ABlue-winged Teal Anas discors Aves/Anseriformes/Anatidae AGarganey Anas querquedula Aves/Anseriformes/Anatidae ABaikal Teal Anas formosa Aves/Anseriformes/Anatidae ARing-necked Duck Aythya collaris Aves/Anseriformes/Anatidae ATufted Duck Aythya fuligula Aves/Anseriformes/Anatidae ALesser Scaup Aythya affinis Aves/Anseriformes/Anatidae AHooded Merganser Lophodytes cucullatus Aves/Anseriformes/Anatidae AGreat Egret Ardea alba Aves/Ciconiiformes/Ardeidae ARed-tailed Hawk Buteo jamaicensis Aves/Falconiformes/Accipitridae AAmerican Kestrel Falco sparverius Aves/Falconiformes/Falconidae ALesser Sand-plover Charadrius mongolus Aves/Charadriiformes/Charadriidae A
Appendix D: Species List 

D-10 Alaska Peninsula and Becharof NWR Compre

Common Name Scientific Name Class/Order/Family 

Black-headed Gull Larus ridibundus Aves/Charadriiformes/Laridae ABlack-tailed Gull Larus crassirostris Aves/Charadriiformes/Laridae ARing-billed Gull Larus delawarensis Aves/Charadriiformes/Laridae ASlaty-backed Gull Larus schistisagus Aves/Charadriiformes/Laridae AWestern Gull Larus occidentalis Aves/Charadriiformes/Laridae ARed-legged Kittiwake Rissa brevirostris Aves/Charadriiformes/Laridae ACaspian Tern Sterna caspia Aves/Charadriiformes/Laridae ARufous Hummingbird Selasphorus rufus Aves/Apodiformes/Trochilidae AHairy Woodpecker Picoides villosus Aves/Piciformes/Picidae AWestern Wood-Pewee Contopus sordidulus Aves/Passeriformes/Tyrannidae ASky Lark Alauda arvensis Aves/Passeriformes/Alaudidae ATownsend's Solitaire Myadestes townsendi Aves/Passeriformes/Turdidae AOlive-backed Pipit Anthus hodgsoni Aves/Passeriformes/Motacillidae ABohemian Waxwing Bombycilla garrulus Aves/Passeriformes/Bombacillidae ATennessee Warbler Vermivora peregrina Aves/Passeriformes/Parulidae AChipping Sparrow Spizella passerina Aves/Passeriformes/Emberizidae ABrown-headed Cowbird Molothrus ater Aves/Passeriformes/Icteridae ABrambling Fringilla montifringilla Aves/Passeriformes/Fringillidae AEXOTIC Two Species Rock Pigeon Columba livia Aves/Columbiiformes/Columbidae EEuropean Starling Sturnus vulgaris Aves/Passeriformes/Sturnidae EMAMMALS—TERRESTRIAL Montane (Dusky) Shrew Sorex monticolus Mammalia/Insectivora/Scoricidae RCinereus (Masked) Shrew Sorex cinereus Mammalia/Insectivora/Scoricidae RTundra Shrew Sorex tundrensis Mammalia/Insectivora/Scoricidae RLittle Brown Bat Myotis lucifugas Mammalia/Chiroptera/Vespertilionidae RCoyote Canis latrans Mammalia/Carnivora/Canidae RGray Wolf Canis lupus Mammalia/Carnivora/Canidae RRed Fox Vulpes vulpes Mammalia/Carnivora/Canidae RBrown Bear Ursus arctos Mammalia/Carnivora/Ursidae RShort-tailed Weasel Mustela erminea Mammalia/Carnivora/Mustelidae R
A

Alaska Peninsula and Becharof NWR Comprehensive Conservation Plan 

Common Name Scientific Name Class/Order/Family 

Least Weasel Mustela nivalis Mammalia/Carnivora/Mustelidae RMink Mustela vison Mammalia/Carnivora/Mustelidae RWolverine Gulo gulo Mammalia/Carnivora/Mustelidae RRiver Otter Lontra canadensis Mammalia/Carnivora/Mustelidae RLynx Lynx Canadensis Mammalia/Carnivora/Felidae RMoose Alces alces Mammalia/Artiodactyla/Cervidae RCaribou Rangifer tarandus Mammalia/Artiodactyla/Cervidae RHoary Marmot Marmota caligata Mammalia/Rodentia/Sciuridae RArctic Ground Squirrel Spermophilus parryii (undulatus) Mammalia/Rodentia/Sciuridae RRed Squirrel Tamiasciurus Hudsonicus Mammalia/Rodentia/Sciuridae RBeaver Castor canadesis Mammalia/Rodentia/Castoridae RRed-backed Vole Cletherionomys rutilus Mammalia/Rodentia/Muridae RTundra Vole Microtus oeconomus Mammalia/Rodentia/Muridae RMeadow Vole? Microtus pennsylvanicus Mammalia/Rodentia/Muridae RMuskrat Ondatra zibethicus Mammalia/Rodentia/Muridae RBrown Lemming Lemmus sibiricus Mammalia/Rodentia/Muridae RNorthern (Collared) Lemming Dicrostonyx groeniandicus Mammalia/Rodentia/Muridae RMeadow Jumping Mouse Zapus hudsonius Mammalia/Rodentia/Dipodidae RPorcupine Erethizon dorsatum Mammalia/Rodentia/Erethizontidae RSnowshoe (Varying) Hare Lepus americanus Mammalia/Lagomorpha/Leporidae RAlaskan (Tundra) Hare Lepus othus Mammalia/Lagomorpha/Leporidae RMAMMALS—MARINE Sea Otter Enhydra lutris Mammalia/Carnivora/Mustelidae RSteller Sea-lion Eumetopias jubatus Mammalia/Carnivora/Otariidae RHarbor Seal Phoca vitulina Mammalia/Carnivora/Phocidae RACC or NEARSHOREMARINE MAMMALS1Walrus Odobenus resmorus Mammalia/Carnivora/Odobenidae N

1 Of the species listed in this section, only gray whales and rrcas have been documented adjacent to Refuges boundaries. While the othe

documented within Refuges boundaries, they are present in nearshore areas. 
Appendix D: Species List 

D-12 Alaska Peninsula and Becharof NWR Compre

Common Name Scientific Name Class/Order/Family 

Northern Fur Seal Callorhinus ursinus Mammalia/Carnivora/Otariidae TN. Right Whale Eubalaena glacialis Mammalia/Cetacea/Balaenidae NMinke Whale Balaenoptera acutorostrata Mammalia/Cetacea/Balaenopteridae NHumpback Whale Megaptera novaeangliae Mammalia/Cetacea/Balaenopteridae NGray Whale Eschrichtius robustus Mammalia/Cetacea/Eschrichtidae NPacific White-sided Dolphin Lagenorhynchus obliquidens Mammalia/Cetacea/Delphinidae NPilot Whale Globicephala sp Mammalia/Cetacea/Delphinidae NKiller Whale Orcinus Orca Mammalia/Cetacea/Delphinidae NBeluga Delphinapterus leucas Mammalia/Cetacea/Monodentidae NHarbor Porpoise Phocoena phocoena Mammalia/Cetacea/Phocoenidae NDall Porpoise Phocoenoides dalli Mammalia/Cetacea/Phocoenidae N
Appendix D: Species List 

Alaska Peninsula and Becharof NWR Comprehensive Conservation Plan D-13 

Common Name Scientific Name (family, species) (DOI 1974)

Club Moss Family LYCOPODIACEAE 

Fir Club Moss Lycopodium selago

Stiff Club Moss Lycopodium annotinum 

Common Club Moss Lycopodium clavatum 

Alpine Club Moss Lycopodium alpinum 

 

Quiltwort Family ISOETACFAE

Isoetes muricata

 

Horsetail Family EQUISETACEAE

Equisetum fluviatile

Equisetum silvaticum 

Equisetum arvense 

 

Adder’s Tongue Family OPHIOGLOSSACEAE

Moonwort Botrychium lunaria

Botrychium lanceolatum 

 

Mountain Parsley Family CRYPTOGRAMMACEAE

Parsley Fern Cryptogramma crispa

 

Marsh Fern Family THEYPTERIDACEAE

Thelypteris phegopteris

 

Lady Fern Family ATHYRIACEAE

Lady Fern Athyrium filix-femina

Fragile Fern Cystopteris fragilis 

 

Shield Fern Family ASPIDIACEAE

Dryopteris dilatata

Gymnocarpium dryopteris 

 

Licorice Fern Family POLYPODIACEAE

Polypodium vulgare

 

Bur Reed Family SPARGANIACEAE

Sparganium hyperboreum

 

Pondweed Family POTAMOGETONACEAE

Eelgrass Zostera marina

Potamogeton alpinus 

Potamogeton perfoliatus 

Potamogeton filiformis 

Ditch Grass Ruppia spiralis 

 

Arrow Grass Family JUNCAGINACEAE

Trigolochin maritimum

Trigolochin palustris 
Appendix D: Species List 

D-14 Alaska Peninsula and Becharof NWR Comprehensive Conservation Plan 

Common Name Scientific Name (family, species) (DOI 1974)

Grass Family GRAMINEAE

Holy Grass Hierochloe alpina

Vanilla grass Hierochloe odorata 

Mountain Timothy Phleum commutatum 

Timothy Phleum pratense 

Foxtail Alopecurus aequalis 

Polar Grass Arctagr stis latifolia 

Wood Reed Grass Cinna latifolia 

Bent Grass Agrostis alaskana 

Agrostis exarata 

Agrostis seabra 

Agrostis geminata 

Bluejoint Calamogrostis canadensis 

Deschampsia caespitosa 

Deschampsia beringensis 

Vahlodea atropurpurea 

Trisetum spicatum 

Poa arctica 

Poa eminens 

Poa palustris 

Poa nemoralis 

Poa hispidula 

Poa stenantha 

Poa annua 

Poa brachyanthera 

Alkali Grass Puccinellia nutkaensis 

Fuscue Grass Festuca altaica 

Festuca rubra 

Brome Grass Bromus ciliatus 

Hordeum brach 

Yantherum 

Lyme Grass Elymus arenarius 

 

Sedge Family CYPERACEAE

Cotton Grass Eriophorum angustifolium

Eriophorum russeolum 

Trichophorum caespitosum 

Sedge Carex anthoxanthea 

Carex circinnata 

Carex nigricans 

Carex macrocephala 

Carex macloviana 

Carex canescens 

Carex Kelloggii 

Carex aquatlilis 

Carex Lyngbyaei 

Carex stylosa 

Carex Gmelini 

Carex machrochaeta 

Carex spectabilis 

Carex nesophila 
Appendix D: Species List 

Alaska Peninsula and Becharof NWR Comprehensive Conservation Plan D-15 

Common Name Scientific Name (family, species) (DOI 1974)

Carex pluriflora 

Carex saxatilis 

 

Rush Family JUNCACEAE

Juncus arcticus

Juncus mertensianus 

Juncus castaneus 

Wood Rush Luzula wahlenbergii 

Luzula parviflora 

Luzula arcuata 

Luzula tundricola 

Luzula multiflora 

Luzula spicata 

 

Lily Family LILIACEAE

False Asphodel Tofieldia coccinea

Kamchatka Fritillary, Sarana Fritillaria camschatcensis 

Alp Lily Lloydia serotina 

Twisted-stalk Streptopus amplexifolius 

 

Iris Family IRIDACEAE

Wild Flag Iris setosa

 

Orchid Family ORCHIDACEAE

Lady’s Slipper Cypripedium guttatum

Key Flower Dactylorhiza aristata 

Frog Orchid Coeloglossum viride 

Bog Orchid Platanthera convallariaefoia 

Ladies’ Tresses Platanthera dilatata 

Spiranthes romanzoffiana 

Listera cordata 

 

Willow Family SALICACEAE

Balsam Poplar, Cottonwood Populus balsamifera

Netted Willow Salix reticulata 

Salix routundifolia 

Salix arctica 

Salix fuscescens 

Salix cyclophylla 

Salix glauca 

Salix barclayi 

Salix commutata 

Alaska Willow Salix alaxensis 

Salix rulchra 

 

Birch Family BETULCEAE

Sitka Alder Alnus crispa

Alnus sinuata 

Betula kenaica 
Appendix D: Species List 

D-16 Alaska Peninsula and Becharof NWR Comprehensive Conservation Plan 

Common Name Scientific Name (family, species) (DOI 1974)

Nettle Family URTICACEAE

Urtica gracilis

 

Buckwheat Family POLYGONACEAE

Koenigia islandica

Sheep Sorrel Rumex acetosella 

Rumex graminifolius 

Rumex fenestratus 

Mountain Sorrel Oxyria digyna 

Polygonum viviparum 

Mountain Sorrel Polygonum caurianum 

 

Goosefoot Family CHENOPODIACEAE

Atiplex alaskensis

 

Pursalane Family PORTULACEAE

Spring Beauty Claytonia sibrica

Claytonia sarmentosa 

Claytonia chamissoi 

Water Blinks Montia fontana 

 

Pink Family CARYOPHYLLACEAE

Chickweed Stellaria media

Stellaria crispa 

Stellaria humifusa 

Stellaria calycantha 

Stellaria sitchana 

Stellaria monantha 

Mouse-ear Chickweed Cerastium berringianum 

Cerastium fischerianum 

Cerastium fontanum 

Pearlwort Sagina intermedia 

Sagina crassicaulis 

Minuartia macrocarpa 

Minuartia arctica 

Minuartia rubella 

Honckenya peploides 

Grove Sandwort Moehringia lateriflora 

Moss Campion Silene acaulis 

 

Water Lily Family NYMPHAEACEAE

Yellow pond Lily Nuphar polysepalum

 

Crowfoot Family RANUNCULACEAE

Caltha palustris

Actaea rubra 

Monkshood Aconitum delphinifolium 

Aconitum maximum 

Coptis trifolia 

Anemone richardsonii 

Anemone parviflors 
Appendix D: Species List 

Alaska Peninsula and Becharof NWR Comprehensive Conservation Plan D-17 

Common Name Scientific Name (family, species) (DOI 1974)

Anemone narcissiflora 

Buttercup, Crowfoot Ranunculus sp. 

White Water Crowfoot Ranunculus trichophyllus 

Creeping Spearwort Ranunculus reptans 

Ranunculus eschscholtzii 

Ranunculus Bongardi 

Meadow Rue Thalictrum sparsiflorum 

 

Poppy Family PAPAVERACEAE

Papaver alaskanum

 

Earth Smoke Family FUMARIACEA

Corydalis pauciflora

 

Mustard Family CRUCIFERAE

Winter Cress Barbarea orthoceras

Yellow Cress Rorippa islandica 

Bitter Cress Cardamine bellidifolia 

Cuckoo Flower Cardamine pratensis 

Capsella rubella 

Draba nivalis 

Draba borealis 

Rock Cress Arabis lyrata 

Arabis hirsuta 

 

Sundew Family DROSERACEA

Drosera rotundifolia

 

Stonecrop Family CRASSUIACEAE

Roseroot Sedum rosea

 

Saxifrage Family SAXIFRAGACEAE

Leatherleaved Saxifrage Leptarrhena pyrolifolia

Purple Mountain Saxifrage Saxifraga oppositifolia 

Saxifraga serpyllifolia 

Box Saxifrage Saxifraga hirculus 

Spotted Saxifrage Saxifraga bronchialis 

Saxifraga punctata 

Saxifraga bracteata 

Saxifraga rivularis 

Saxifraga unalaschcensis 

Snow Saxifrage Saxifraga nivalis 

Coast Saxifrage Saxifraga ferruginea 

Tufted Saxifrage Saxifraga caespitosa 

Alpine Heuchera Heuchera glabra 

Fringe Cups Tellima grandiflora 

Northern Water Carpet Chrysosplenium tetrandrum 

Chrysosplenium wrightii 

Parnassia kotzebuei 

Parnassia palustris 
Appendix D: Species List 

D-18 Alaska Peninsula and Becharof NWR Comprehensive Conservation Plan 

Common Name Scientific Name (family, species) (DOI 1974)

Rose Family ROSACEAE

Alaskan Spiraea Spiraea beauverdiana

Luetkea pectinata 

Goatsbeard Aruncus sylvester 

Rubus arcticus 

Cloudberry Rubus chamaemorus 

Salmonberry Rubus spectabilis 

Marsh Fivefinger Potentilla palustris 

Potentilla villosa 

Pacific Silverweed Potentilla Egedii 

Sibbaldia procumbens 

Avens Geum macrophyllum 

Geum Rossii 

Burnet Sanguisorba stipulata 

Fragaria chiloensis 

Dryas sp. 

 

Pea Family LEGUMINOSAE

Lupine Lupinus nootkatensis

Milk Vetch Astragalus polaris 

Oxtropis Maydellians 

Oxtropis nigrescens 

Lathyrus maritimus 

Vetchling Lathyrus palustris 

 

Geranium Family GERANIACEAE

Geranium erianthum

 

Water Startwort Family CALLITRICHACEA

Callitriche verna

 

Violet Family VIOLACEAE

Viola biflora

Viola langsdorfii 

Viola epipsila 

 

Evening Primrose Family ONAGRACEAE

Willow Herb Epilobium angustifolium

River Beauty Epilobium latifolium 

Epilobium luteum 

Epilobium palustre 

Epilobium leptocarpum 

Epilobium glandulosum 

Epilobium sertulatum 

Epilobium anagallidifolium 

Epilobi Hornemannii 

Enchanter’s Nightshade Circaea alpina 

 

Water Milfoil Family HALORAGACEAE

Mare’s Tail Hippuris vulgaris

Hippuris tetraphylla 
Appendix D: Species List 

Alaska Peninsula and Becharof NWR Comprehensive Conservation Plan D-19 

Common Name Scientific Name (family, species) (DOI 1974)

Parsley Family UMBELLIFERAE

Water Hemlock Cicuta mackenzieana

Beach Lovage Ligusticum scoticum 

Hemlock Parsley Conioselinum chinese 

Wild Celery Angelica lucida 

Cow parsnip Heracleum lanatum 

Bupleurum triradiatum 

 

Dogwood Family CORNACEAE

Swedish Dwarf Cornel Cornus suecica

Bunchberry; Canadian DwarfCornel Cornus canadensis 

 

Wintergreen Family PYROLACEAE

Pyrola asarifolia

Pyrola secunda 

Pyrola minor 

 

Crowberry Family EMPETRACEAE

Empetrum nigrum. 

 

Heath Family ERICACEAE

Labrador Tea Ledum palustre

Kamchatka Rhododendron Rhododendron camtschaticum 

Alpine Azalea Loiseleuria procumbens 

Mountain Heather Phyllodoce aleutica 

Alaska Moss Heath Cassiope stelleriana 

Cassiope lycopodioides 

Andromeda polifolia 

Arctostaphylos alpina 

Arctostaphylos uva-ursi 

Lingonberry Vaccinium vitis-idaea 

Alpine Blueberry Vaccinium uliginosum 

Vaccinium ovalifolium 

Cranberry Oxycoccus microcarpus 

Diapensia Family DIAPENSIACEAE

Diapensia lapponica

Primrose Family PRIMULACEAE

Primula cuneifolia

Greenland Primrose Primula egaliksensis 

Androsace chamaejasme 

Androsace alaskana 

Starflower Trientalis europaea 

Leadwort Family PLUMBAGINACEAE

Thrift Armeria maritima

Genetian Family GENTIANACEAE

Gentiana algida

Gentiana amarella 
Appendix D: Species List 

D-20 Alaska Peninsula and Becharof NWR Comprehensive Conservation Plan 

Common Name Scientific Name (family, species) (DOI 1974)

Buckbean Menyanthes trifoliata 

Swertia perennis 

Polemoaium Family POLEMONIACEAE

Polenomium acutiflorum

Borage Family BORAGINACEAE

Myosotis alpestris

Mertensia maritima 

Figwort Family SCROPHULARIACEAE

Mimulus sp.

Veronica sp. 

Lagotis glauca 

Castilleja unalaschensis 

Rhinanthus minor 

Pedicularis kanei 

Pedicularis capitata 

Bladderwort Family LENTIBULARIACEAE

Pinguicula vulgaris

Madder Family RUBIACEAE

Galeum sp.

Honeysuckle Family CAPRIFOLIACEAE

Sambucus racemosa

Viburnum edula 

Valerian Family VALERIANACEAE

Valeriana capitata

Bluebell Family CAMPANULACEAE

Campanula lasiocarpa

 

Composite Family COMPOSITAE 

Goldenrod Solidago multiradiata

Solidago lepida 

Aster sibiricus 

Coastal Fleabane Erigeron peregrinus 

Pussytoe Antennaria monocephala 

Pearly Everlasting Anaphalis argaritacea 

Yarrow Achillea borealis 

Pineapple Weed Matricaria matricatiodes 

Arctic Daisy Chrysanthemum arcticum 

Wormwood Artemisia globularia 

Artemisia tilesii 

Artemisia arctica 

Petasites frigidus 

Petasites hyperboreusS 

Arnica Lessingii 

Arnica Chamissonis 
Appendix D: Species List 

Alaska Peninsula and Becharof NWR Comprehensive Conservation Plan D-21 

Common Name Scientific Name (family, species) (DOI 1974)

Groundsel, Ragwort Senecio resedifolius 

Senecio pseudo-Arnica 

Dandelion Taraxacum trignolobum 

Rattlesnake Root Prenanthes alata 

References Cited 

DOI. 1974. “Proposed Katmai National Park, Alaska.” 

Final environmental impact statement. Washington, 

D.C.: U.S. Department of the Interior.