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Catalog of Type Specimens 

of Recent Mammals: 

Rodentia (Sciuromorpha 

and Castorimorpha) 

in the National Museum 

of Natural History, 

Smithsonian Institution

Robert D. Fisher and Craig A. Ludwig

smithsonian contributions to zoology • number 640
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smithsonian contributions to zoology • numb e r 6 4 0

Catalog of Type Specimens 

of Recent Mammals: 

Rodentia (Sciuromorpha 

and Castorimorpha) 

in the National Museum 

of Natural History, 

Smithsonian Institution

Robert D. Fisher and Craig A. Ludwig

washington d.c.

2012
ABSTRACT

Fisher, Robert D., and Craig A. Ludwig. Catalog of Type Specimens of Recent Mammals: Rodentia (Sciuromorpha and 

Castorimorpha) in the National Museum of Natural History, Smithsonian Institution. Smithsonian Contributions to Zoology, number 640, iv + 97 pages, 2012.— The type collection of Recent mammals in the Division of Mammals, National 

Museum of Natural History, Smithsonian Institution, contains 843 specimens bearing names of 820 species group taxa of 

Rodentia (Sciuromorpha and Castorimorpha) as of July 2011. This catalog presents a list of these holdings, which comprise 798 holotypes, 14 lectotypes, seven syntypes (30 specimens), and one neotype. In addition, we include three holotypes 

and 10 specimens that are part of syntype series that should be in the collection but cannot be found and three syntypes 

that were originally in this collection but are now known to be in other collections. One specimen that no longer has namebearing status is included for the record. Forty-one of the names are new since the last type catalog. One new lectotype is 

designated. Suborders and families are listed as in Wilson and Reeder. Within families, currently recognized genera are arranged alphabetically. Within each currently recognized genus, accounts are arranged alphabetically by original published 

name. Information in each account includes original name and abbreviated citation thereto, current name if other than 

original, citation for first use of current name combination for the taxon (or new name combination if used herein for the 

first time), type designation, U.S. National Museum catalog number(s), preparation, age and sex, type locality, date of collection and name of collector, collector’s original number, and comments or additional information as appropriate. Digital 

photographs of each specimen serve as a condition report and will be linked to each electronic specimen record.

Cover images: Holotype of Callospermophilus chrysodeirus mitratus A. H. Howell, 1931 (USNM 138125). Left to right: 

dorsal, lateral, and ventral views of cranium. Photographs by D. F. Schmidt.

Published by Smithsonian Institution Scholarly Press

P.O. Box 37012, MRC 957

Washington, D.C. 20013-7012

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Compilation copyright © 2012 Smithsonian Institution

The text of this publication is in the public domain. The rights to all other content of this publication, including cover and 

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Library of Congress Cataloging-in-Publication Data

National Museum of Natural History (U.S.)

Catalog of type specimens of recent Mammals:Rodentia (SciuromorpIa and Castorimorpha) in the National Museum 

of Natural History, Smithsonian Institution / Robert D. Fisher and Craig A. Ludwig.

 p. cm. — (Smithsonian contributions to zoology ; no. 640)

Includes bibliographical references and index.

1. Rodents—Type specimens—Catalogs and collections—Washington (D.C.) 2. National Museum of Natural History 

(U.S.)—Catalogs. I. Fisher, Robert D. II. Ludwig, Craig A. III. Title.

QL737.R6N227 2012

599.35074’753--dc23

2012014838

ISSN: 0081-0282 (print); 1943-6696 (online)

The paper used in this publication meets the minimum requirements of the American National Standard for 

Permanence of Paper for Printed Library Materials Z39.48–1992.
Contents

INTRODUCTION 1

Type Categories Included 1

Organization 2

Formant of Accounts 2

Approach 2

Acknowledgments 3

ACCOUNTS OF NAME-BEARING TYPE SPECIMENS 3

Suborder Sciuromorpha Brandt, 1855 3

 Family Aplodontiidae Brandt, 1855 3

 Genus Aplodontia Richardson, 1829 3

 Family Sciuridae G. Fischer, 1814 3

 Genus Aretes G. M. Allen, 1940 3

 Genus Ammospermophilus Merriam, 1892 4

 Genus Callosciurus Gray, 1867 4

 Genus Callospermophilus, Merriam, 1897 10

 Genus Cynomys Rafinesque, 1817 11

 Genus Dremomys Heude, 1898 11

 Genus Exilisciurus Moore, 1958 11

 Genus Glaucomys Thomas, 1908 11

 Genus Heliosciurus Trouessart, 1880 13

 Genus Hylopetes Thomas, 1908 13

 Genus Ictidomys J. A. Allen, 1877 13

 Genus Iomys Thomas, 1908 14

 Genus Lariscus Thomas and Wroughton, 1909 14

 Genus Marmota Blumenbach, 1779 14

 Genus Menetes Thomas, 1908 15

 Genus Microsciurus J. A. Allen, 1895 15

 Genus Nannosciurus Trouessart, 1880 16

 Genus Notocitellus A. H. Howell, 1938 16

 Genus Otospermophilus Brandt, 1844 16

 Genus Paraxerus Forsyth Major, 1893 17

 Genus Petaurista Link, 1795 17
i v • smithsonian contributions to zoology

 Genus Petinomys Thomas, 1908 17

 Genus Prosciurillus Ellerman, 1947 17

 Genus Protoxerus Forsyth Major, 1893 18

 Genus Pteromys G. Cuvier, 1800 18

 Genus Ratufa Gray, 1867 18

 Genus Sciurus Linnaeus, 1758 20

 Genus Sundasciurus Moore, 1958 25

 Genus Tamias Illiger, 1811 26

 Genus Tamiasciurus Trouessart, 1880 31

 Genus Tamiops J. A. Allen, 1906 32

 Genus Urocitellus Obolenskij, 1927 33

 Genus Xerospermophilus Merriam, 1892 36

 Family Gliridae Muirhead, 1819 37

 Genus Eliomys Wagner, 1840 37

 Genus Graphiurus Smuts, 1832 37

 Genus Myomimus Ognev, 1924 38

Suborder Castorimorpha Wood, 1955 38

 Family Castoridae Hemprich, 1820 38

 Genus Castor Linnaeus, 1758 38

 Family Heteromyidae Gray, 1868 38

 Genus Chaetodipus Merriam, 1889 38

 Genus Dipodomys Gray, 1841 42

 Genus Heteromys Desmarest, 1817 47

 Genus Microdipodops Merriam, 1891 50

 Genus Perognathus Wied-Neuwied, 1839 50

 Family Geomyidae Bonaparte, 1845 54

 Genus Cratogeomys Merriam, 1895 54

 Genus Geomys Rafinesque, 1817 57

 Genus Orthogeomys Merriam, 1895 58

 Genus Pappogeomys Merriam, 1895 59

 Genus Thomomys Wied-Neuwied, 1839 60

 Genus Zygogeomys Merriam, 1895 72

REFERENCES 73

INDEX OF TAXA 83
INTRODUCTION

The collection of primary type specimens in the Division of Mammals, National 

Museum of Natural History (NMNH), Smithsonian Institution, formerly the U.S. National Museum (USNM) is the second largest in the world, numbering 3,213 specimens. 

Type specimens are “the international standards of reference that provide objectivity in 

zoological nomenclature” and as such “every institution in which name-bearing types 

are deposited should publish lists of name-bearing types in its possession or custody” 

(International Commission on Zoological Nomenclature, 1999:79).

This is the third type catalog for USNM mammals following Lyon and Osgood 

(1909), 1,415 specimens reported, and Poole and Schantz (1942) 2,824 specimens reported. However, we deviate from the previous catalogs and follow the precedent set 

by the Division of Fishes (Vari and Howe, 1991) and the Division and Amphibians and 

Reptiles (Reynolds et al., 2007) in which the entire type collection is not addressed as one 

but rather divided into smaller, more manageable units. We anticipate the current work 

to be the first of at least four such publications.

Type Categories Included

Specimens included in this catalog are one of the following as defined by the International Commission on Zoological Nomenclature (1999).

Holotype. The single specimen upon which a new nominal species group taxon is 

based in the original publication (Article 73.1).

Syntype. One of multiple specimens on which a species group name is equally based 

when no holotype is specified in the original publication and no subsequent designation of a lectotype has been published (Article 73.2). Syntypes are referred to as cotypes 

in Lyon and Osgood (1909) and Poole and Schantz (1942) as well as in other early 

literature.

Lectotype. One of multiple specimens upon which a species group name is originally 

based, designated in a publication subsequent to the original description to become the 

unique name bearer (Article 74.1).

Neotype. A specimen chosen as the name bearer of a species group taxon if no holotype, lectotype, syntype, or prior neotype is believed to be extant (Article 75.1).

Catalog of Type Specimens of Recent 

Mammals: Rodentia (Sciuromorpha and 

Castorimorpha) in the National Museum of 

Natural History, Smithsonian Institution

Robert D. Fisher, U.S. Geological Survey, Patuxent Wildlife Research Center, National Museum 

of Natural History, Smithsonian Institution, 

Washington, D.C. 20013-7012, USA; Craig A. 

Ludwig, Division of Mammals, Department of 

Vertebrate Zoology, National Museum of Natural History, Smithsonian Institution, Washington, D.C. 20013-7012, USA; Correspondence: 

fisherr@si.edu; ludwigc@si.edu. Manuscript received 30 September 2011; accepted 3 April 2012.
2 • smithsonian contributions to zoology

Organization

Suborders and families are arranged following Wilson and 

Reeder (2005). Within families, currently recognized genera are arranged alphabetically. Within each currently recognized genus, the 

accounts are arranged alphabetically by original published name.

Format of Accounts

We follow the format set by Lyon and Osgood (1909) and 

Poole and Schantz (1942) with two notable exceptions. At the 

time of these earlier catalogs the collections of the Division of 

Mammals and those of the Biological Survey/U.S. Fish and Wildlife Service were housed separately, as were their respective type 

collections. The earlier catalogs thus stated in which collection a 

specimen was located. The two collections, including the types, 

were merged many years ago, and thus no such designation is 

given here. The earlier catalogs also stated in general terms or 

code the condition of each specimen. We use digital photography 

combined with additional comments in the photographic metadata as necessary to report specimen condition.

Each account reads as follows: (1) Original name in bold

followed by the authors’ name; (2) citation of original description including publication, volume, page, and date; (3) currently 

used name if different from the original followed by citation for 

the first use of current name combination for the taxon. Author 

names in parentheses indicate the generic name used in the original description was different than the current. Current usage 

follows the authors in Wilson and Reeder (2005) unless more 

recent authors are cited. (4) type designation and USNM catalog number (s); (5) preparation of specimen; (6) age and sex; 

(7) collecting locality; (8) date collected and collector(s) name(s); 

(9) original number, which is usually the field number; and (10) 

comments that seem appropriate.

Further explanatory comments are in order for some of the 

information in these accounts. Every effort has been made to make 

obvious what was included in the original description as opposed 

to that added later from various sources. Added information, 

such as amended or restricted localities, is given with citations. 

Brackets enclose additional data not in the original description 

and, when not from commonly used sources such as maps or online gazetteers, are attributed to collector’s Field Catalog (FC), 

Museum Catalog (MC), Merriam Collection Catalog (MCC), 

skin label (SL), X-Catalog (XC), or a citation. Brackets surround 

USNM numbers of type specimens described prior to becoming 

part of the National Collection. Parentheses are used around metric equivalents and current place names when this information 

was not included in the original publication. In the early years of 

the U.S. National Museum a double number catalog system was 

used in which skins and alcoholics were entered in one catalog 

and skeletal material in another. Many specimens therefore have 

two numbers associated with them and the complete number is 

indicated by a fraction such as 23651/31045. This numerator on 

the skin label indicates 23651 as the skin to which skull 31045 

belongs; on the skull label 31045/23651 the numerator indicates 

31045 as the skull to which skin 23651 belongs. This catalog 

system was abandoned at the end of 1892 when a single numbering system was implemented in which all parts to a specimen 

were assigned the same number. The new system was put in place 

in January 1893 with number 50001. The C. Hart Merriam private collection (MCC), donated to the U.S. National Museum 

and mentioned in many accounts, also used the double numbering system. Finally, the X-Catalog requires explanation. Beginning with the Biological Survey and continuing with the Fish and 

Wildlife Service, specimens acquired by these agencies through 

exchange or purchase or specimens received from or collected by 

persons other than permanent staff, were entered in this catalog 

prior to being transferred to the Division of Mammals for cataloging. Frequently, these entries have data that were never entered 

into the Museum Catalog or onto the skin labels. The last entry in 

the X-Catalog was on 5 October 1951.

Approach

In the early 1970s the accounts in a copy of Poole and 

Schantz (1942) were cut out and pasted onto 5 × 7 index cards 

to create a card file of types. Additional cards were created and 

added to the file as new types were installed in the collection. The 

information on each card for the taxa included in this catalog 

was checked against the original description, the original skin 

label (when present), and the field catalog entry (when available). Other catalogs such as the Museum Catalog, Merriam 

Collection Catalog, and X-Catalog were consulted as needed. 

Information on each index card, often annotated by hand, was 

used to create each of the accounts.

Authors in Wilson and Reeder (2005) are followed for currently accepted names unless an online search of Zoological Record, conducted through 2009 for Sciuromorpha and 2010 for 

Castorimorpha, uncovered more recent taxonomic conclusions. 

Each specimen record in the collection database was corrected as 

needed to reflect new information. The type publication citation 

was added to the electronic record if not already present. Each 

Museum Catalog record was checked and updated with publication data as required in the hand ledgers.

Digital images are being created for each specimen and include the following: dorsal, lateral, and ventral views of crania; 

dorsal and lateral views of mandibles; dorsal and ventral views 

of skins (lateral if possibly diagnostic); and both sides of all labels. Additional views are included when deemed appropriate. In 

the case of lateral images, the traditional view of the left aspect 

is presented except in instances where the right aspect is more 

complete owing to damage on the left side. Specimen damage not 

captured in these images is recorded in the associated image metadata when readily apparent. No concerted effort was made to minutely examine specimens for damage. All images feature a metric 

scale bar. Images will be linked to the specimen database as they 
number 640 • 3

become available and can be accessed via the Internet. Images 

are being created in the order in which accounts are presented. 

We anticipate it will take about two years for all images to be 

completed for specimens in this catalog. These images will serve 

not only as specimen condition indicators, but in many instances 

they also will allow researchers to examine type specimens without having to travel to the U.S. National Museum and handle the 

specimens, thereby providing a convenience to researchers and 

minimizing possible damage to irreplaceable specimens.

Acknowledgments

We thank our supervisors, Robert P. Reynolds, USGS Patuxent Wildlife Research Center, NMNH, and Kristofer L. Helgen, 

Division of Mammals, NMNH, respectively, for support, encouragement, and time to devote to this project. Funds to purchase 

the photographic setup, in order of significance, were provided 

by Carol E. Butler, Chief of Collections, NMNH; Don E. Wilson, 

Chair, Department of Vertebrate Zoology, NMNH; and Robert 

P. Reynolds, to whom we are thankful. We are greatly indebted 

to David F. Schmidt, Division of Mammals, NMNH, with support from Linda K. Gordon, Division of Mammals, NMNH, who 

spent a considerable amount of time researching photographic 

equipment and methods, ordered the equipment, set up the photographic lab, and created images of the first 100 or so specimens before and for some time following his retirement. Advice 

on photographic equipment, setup, and technique was graciously 

provided to David by Melvin J. Wachowiak and Elizabeth Webb, 

Museum Conservation Institute, Smithsonian Institution; Roy 

and Jason Larimer, Visionary Digital; and James L. Di Loreto, Office of Imaging and Photographic Services, NMNH. Joelle Mintz, 

USGS Patuxent Wildlife Research Center, NMNH, checked the 

museum catalog hand ledgers and added publication data for 

all type records as needed. Alfred L. Gardner, USGS Patuxent 

Wildlife Research Center, NMNH, provided valuable insight to 

numerous taxonomic issues and guidance in interpreting various 

provisions of the International Code of Zoological Nomenclature. Alfred L. Gardner, Robert P. Reynolds, and David F. Schmidt 

provided comments that greatly improved the manuscript.

ACCOUNTS OF NAME-BEARING 

TYPE SPECIMENS

Suborder Sciuromorpha Brandt, 1855

Family Aplodontiidae Brandt, 1855

Genus Aplodontia Richardson, 1829

Aplodontia major Merriam

Ann. New York Acad. Sci. 3: 316, May 1886a.

= Aplodontia rufa californica (Peters, 1864). See Taylor (1918:473).

Lectotype: [186473] Skin and skeleton. Adult male. [SL and 

MCC: Blue Canyon], Placer County, California. Collected 7 

Oct 1885 by C. A. Allen. Original number [MCC: 1]. Merriam collection number 1501/2101.

Type not designated by number in original description. Poole 

and Schantz (1942:253) list this as a Lectotype without 

comment, and we can find no earlier designation. The skull 

is the first figured and the first listed in specimens examined.

Aplodontia major rainieri Merriam

Proc. Biol. Soc. Washington 13: 21, 31 Jan 1899a.

= Aplodontia rufa rainieri Merriam, 1899. See Taylor (1918: 465).

Holotype: 90144. Skin and skull. Adult male. Paradise Creek 

[FC: Paradise Park], south side of Mount Rainier, altitude 

5,200 ft (1,585 m), [Pierce County], Washington. Collected 

6 Aug 1897 by V. Bailey. Original number 6122.

Aplodontia olympica Merriam

Proc. Biol. Soc. Washington 13: 20, 31 Jan 1899a.

= Aplodontia rufa rufa (Rafinesque, 1817). See Dalquest and 

Sheffer (1945:37).

Holotype: 89549. Skin and skull. Young adult male. Queniult (Quinault) Lake, Olympic Mountains, [Grays Harbor County], Washington. Collected 24 Jul 1897 by R. T. 

Young. Original number 309.

Aplodontia pacifica Merriam

Proc. Biol. Soc. Washington 13: 19, 31 Jan 1899a.

= Aplodontia rufa pacifica Merriam, 1899. See Taylor (1918:467).

Holotype: 77372. Skin and skull. Adult female. Newport, [SL: 3 

mi (4.8 km) north of town], mouth of Yaquina Bay, [Lincoln 

County], Oregon. Collected 20 [SL: 26, which is correct] 

Mar 1896 by B. J. Bretherton. Original number 2219.

Aplodontia phaea Merriam

Proc. Biol. Soc. Washington. 13: 20, 31 Jan 1899a.

= Aplodontia rufa phaea Merriam, 1899. See Taylor (1918:480).

Holotype: 186475. Skin and skull. Adult male. Point Reyes, Marin 

County, California. Collected 1 Aug 1886 by C. A. Allen. 

Original number 142. Merriam collection number 2645/3273.

Family Sciuridae G. Fischer, 1814

Genus Aeretes G. M. Allen, 1940

Petaurista sulcatus A. B. Howell

J. Washington Acad. Sci. 17: 82, 19 Feb 1927.

= Aeretes melanopterus melanopterus (Milne-Edwards, 1867). 

See G. M. Allen (1940:746).

Holotype: 219206. Skin and skull. Adult female. Hsin-lungshan, 65 mi (105 km) northeast of Peking (Beijing), altitude 

3,000 ft (914 m), Chih-li (Hebei), China. Collected 18 Aug 

1917 by A. de C. Sowerby. Original number 1017.
4 • smithsonian contributions to zoology

Genus Ammospermophilus

Merriam,1892

Ammospermophilus leucurus extimus Nelson and Goldman

J. Washington Acad. Sci. 19: 281, 19 Jul 1929a.

Holotype: 146587. Skin and skull. Adult female. Saccaton, 15 

mi (24 km) north of Cape San Lucas, [SL and FC: altitude 

1,000 ft (305 m)], Lower California (Baja California Sur), 

Mexico. Collected 29 Dec 1905 by E. W. Nelson and E. A. 

Goldman. Original number 18805.

Ammospermophilus leucurus insularis Nelson and Goldman

Proc. Biol. Soc. Washington 22: 24, 10 Mar 1909.

Holotype: 146783. Skin and skull. Adult female. Espiritu Santo 

Island, [Gulf of California], Lower California (Baja California Sur), Mexico. Collected 7 Feb 1906 by E. W. Nelson and 

E. A. Goldman. Original number 19072.

Ammospermophilus leucurus pennipes A. H. Howell

J. Mamm. 12: 162, 14 May 1931.

Holotype: 75683. Skin and skull. Adult female. Grand Junction, 

[Mesa County], Colorado. Collected 11 Nov 1895 by A. H. 

Howell. Original number 494.

Ammospermophilus leucurus tersus Goldman

J. Washington Acad. Sci. 19: 435, 19 Nov 1929.

Holotype: 202645. Skin and skull. Young adult male. Lower 

end of Prospect Valley, altitude 4,500 ft (1,372 m), Grand 

Canyon, Hualapai Indian Reservation, [Mohave County], 

Arizona. Collected 3 Oct 1913 by E. A. Goldman. Original 

number 22269.

Spermophilus harrisii saxicolus Mearns

Proc. U.S. Natl. Mus. 18: 2, 25 Mar 1896a. Preprint of Proc. 

U.S. Natl. Mus. 18: 444, 23 May 1896b.

= Ammospermophilus harrisii saxicolus (Mearns, 1896). See 

Mearns (1907:307).

Holotype: 59869. Skin, skull, [and body with six embryos in alcohol]. Adult female. Tinajas Altas, Gila Mountains, Yuma 

County, Arizona. Collected 17 Feb 1894 by E. A. Mearns 

and F. X. Holzner. Original number 2983.

Spermophilus nelsoni Merriam

Proc. Biol. Soc. Washington 8: 129, 28 Dec 1893a.

= Ammospermophilus nelsoni (Merriam, 1893). See Lyon and 

Osgood (1909:172).

Holotype: 54651. Skin and skull. Adult male. Tipton, San Joaquin Valley, [FC: Tulare County], California. Collected 24 

Jun 1893 by C. P. Streator. Original number 2968.

Tamias interpres Merriam

N. Amer. Fauna 4: 21, 8 Oct 1890a.

= Ammospermophilus interpres (Merriam, 1890). See V. Bailey 

(1905:81).

Holotype: 18162/25060. Skin and skull. Adult female. El Paso, 

[El Paso County], Texas. Collected 10 Dec 1889 by V. Bailey. Original number 762.

Tamias leucurus Merriam

N. Amer. Fauna 2: 20, 30 Oct 1889a.

= Ammospermophilus leucurus leucurus (Merriam, 1889). See 

Mearns (1907:299).

Holotype: [186466. Skin and skull]. Adult male. San Gorgonio 

Pass, [Riverside County], California. Collected 16 May 

1885 by F. Stephens. Original number [MCC: 68]. Merriam 

collection number 1108/1600.

Tamias leucurus cinnamomeus Merriam

N. Amer. Fauna 3: 51, 11 Sep 1890b.

= Ammospermophilus leucurus cinnamomeus (Merriam, 1890). 

See Mearns (1907:299).

Holotype: 17987/24898. Skin and skull. Adult female. Echo 

Cliffs, [SL and FC: Moa Ave.], Painted Desert, [Coconino 

County], Arizona. Collected 22 Sep 1889 by C. H. Merriam 

and V. Bailey. Original number [FC: VB] 510.

Genus Callosciurus Gray, 1867

Callosciurus finlaysonii boonsongi Moore and Tate

Fieldiana Zool. 48: 184, 28 May 1965.

Holotype: 307801. Skin and skull. Adult male [SL and FC: female]. Phu Phan, a mountain in Sakon Nakhon (Amphoe 

Muang Sakon Nakhon) District, Sakon Nakhon Province, 

eastern Thailand. Collected 14 Jun 1954 by R. E. Elbel and 

B. Lekagul. Original number [SL: RE 3700].

Sciurus abbottii Miller

Proc. Washington Acad. Sci. 2: 224, 20 Aug 1900a.

= Callosciurus notatus (Boddaert, 1785). Unassigned to subspecies by Thorington and Hoffmann (2005:778).

Holotype: 101662. Skin and skull. Adult female. Big Tambelan 

Island, [Tambelan Islands, Kepulauan Riau Province, Indonesia]. Collected 10 Aug 1899 by W. L. Abbott. No original 

number.

Sciurus adangensis Miller

Smithsonian Misc. Coll. 45: 17, 6 Nov 1903a.

= Callosciurus caniceps adangensis (Miller, 1903). See Robinson 

and Kloss (1918a:209).

Holotype: 104389. Skin and skull. Adult male. Pulo Adang, Butang Islands [Satun, Thailand]. Collected 14 Dec 1899 by 

W. L. Abbott. Original number 153.

Sciurus altinsularis Miller

Smithsonian Misc. Coll. 45: 21, 6 Nov 1903a.

= Callosciurus caniceps (Gray, 1842) Unassigned to subspecies 

by Thorington and Hoffmann (2005:776).
number 640 • 5

Holotype: 111975. Skin and skull. Adult female. High Island, 

Mergui Archipelago, [Tanintharyi, Myanmar]. Collected 31 

Dec 1900 by W. L. Abbott. Original number 810.

Sciurus anambensis Miller

Proc. Washington Acad. Sci. 2: 223, 20 Aug 1900a.

= Callosciurus notatus (Boddaert, 1785). Unassigned to subspecies by Thorington and Hoffmann (2005:778).

Holotype: 101686. Skin and skull. Adult female. Pulo Siantan, Anambas Islands, [Kepulauan Riau Province, Indonesia]. Collected 12 Sep 1899 by W. L. Abbott. No original 

number.

Sciurus aoris Miller

Smithsonian Misc. Coll. 45: 10, 6 Nov 1903a.

= Callosciurus notatus miniatus (Miller, 1900). See Corbet and 

Hill (1992:291).

Holotype: 112418. Skin and skull. Adult female. Pulo Aor, off 

the coast of Johore (Johor), [Malaysia]. Collected 5 Jun 

1901 by W. L. Abbott. Original number 1002.

Sciurus arendsis Lyon

Proc. U.S. Natl. Mus. 40: 87, 25 Apr 1911a.

= Callosciurus notatus (Boddaert, 1785). Unassigned to subspecies by Thorington and Hoffmann (2005:778).

Holotype: 154276. Skin and skull. Adult male. Arends Island 

(Pulau Keramian) Java Sea, [Kalimantan Selatan, Borneo, 

Indonesia]. Collected 24 Nov 1908 by W. L. Abbott. Original number 6251.

Sciurus armalis Lyon

Proc. U.S. Natl. Mus. 40: 82, 25 Apr 1911a.

= Callosciurus prevostii atricapillus (Schlegel, 1863). See Thorington and Hoffmann (2005:779).

Holotype: 145420. Skin and skull. Adult male. Pulo Panebangan, [Kalimantan Barat], off west coast of Borneo, [Indonesia]. Collected 19 May 1907 by W. L. Abbott. Original 

number 5257.

Sciurus atratus Miller

Smithsonian Misc. Coll. 45: 13, 6 Nov 1903a.

= Callosciurus melanogaster atratus (Miller, 1903). See Hayman 

and Holt (1940:374).

Holotype: 121524. Skin and skull. Adult female. North Pagi Island (Pulau Pagai Utara), [Sumatera Barat], Sumatra, [Indonesia]. Collected 22 Nov 1902 by W. L. Abbott. Original 

number 2087.

Sciurus atricapillus atrox Miller

Smithsonian Misc. Coll. 61 (21): 23, 29 Dec 1913.

= Callosciurus prevostii (Desmarest, 1822). Unassigned to subspecies by Thorington and Hoffmann (2005:779).

Holotype: 196669. Skin and skull. Adult female. Talisaian 

Mountain, [Kalimantan Timur], Dutch Southeast Borneo, 

[Indonesia]. Collected 6 Feb 1913 by H. C. Raven. Original 

number 457.

Sciurus atristriatus Miller

Smithsonian Misc. Coll. 61 (21): 22, 29 Dec 1913.

= Callosciurus notatus (Boddaert, 1785). Unassigned to subspecies by Thorington and Hoffmann (2005:778).

Holotype: 196670. Skin and skull. Half-grown female. Lo Bon Bon, 

[SL: Mahakkan River, North bank], [FC: above Samamila], 

[Kalimantan Timur], Dutch Southeast Borneo, [Indonesia]. 

Collected 4 Jun 1912 by H. C. Raven. Original number 13.

Sciurus atrodorsalis tachin Kloss

J. Nat. Hist. Soc. Siam. 2: 178, Dec 1916. See Kloss (1919) for 

full description.

= Callosciurus erythraeus siamensis (Gray, 1860). See Thorington and Hoffmann (2005:776).

Holotype: [221566]. Skin and skull. Adult female. Pak Bu, 

Tachin, [Samut Sakhon], Central Siam (Thailand). Collected 

23 Oct 1916 by C. B. Kloss. Original number 2213.

Sciurus bentincanus Miller

Smithsonian Misc. Coll. 45: 19, 6 Nov 1903a.

= Callosciurus caniceps (Gray, 1842). Unassigned to subspecies 

by Thorington and Hoffmann (2005:776), but see Moore 

and Tate (1965:195) and Corbet and Hill (1992:288) who 

assign it to C. c. domelicus (Miller, 1903).

Holotype: 104383. Skin and skull. Adult female. Bentinck Island, Mergui Archipelago, [Tanintharyi, Myanmar]. Collected 11 Mar 1900 by W. L. Abbott. Original number 349.

Sciurus bilimitatus Miller

Smithsonian Misc. Coll. 45: 8, 6 Nov 1903a.

= Callosciurus nigrovittatus bilimitatus (Miller, 1903). See Robinson and Kloss (1918a:222).

Holotype: 105072. Skin and skull. Adult female. Tanjong Laboha, 

Tringanu (Terengganu), [Malaysia]. Collected 29 Sep 1900 

by C. B. Kloss. Original number W. L. Abbott 671.

Sciurus billitonus Lyon

Proc. U.S. Natl. Mus. 31: 592, 18 Dec 1906a.

= Callosciurus notatus (Boddaert, 1785). Unassigned to subspecies by Thorington and Hoffmann (2005:778).

Holotype: 124997. Skin and skull. Adult female. Buding Bay, 

Billiton Island, between Sumatra and Borneo, [Kepulauan 

Bangka Belitung, Indonesia]. Collected 5 Aug 1904 by W. 

L. Abbott. Original number 3539.

Sciurus borneoensis palustris Lyon

Proc. U.S. Natl. Mus. 33: 553, 24 Dec 1907a.

= Callosciurus prevostii (Desmarest, 1822). Unassigned to subspecies by Thorington and Hoffmann (2005:778).

Holotype: 142330. Skin and skull. Adult male. North (SL: Right) 

bank of the Kapaus River, below Pulo Limbang, [Kaliman-
6 • smithsonian contributions to zoology

tan Barat], western Borneo, [Indonesia]. Collected 22 Sep 

1905 by W. L. Abbott. Original number 4467.

Sciurus caniceps canigenus A. B. Howell

J. Washington Acad. Sci. 17: 81, 19 Feb 1927.

= Callosciurus erythraeus styani (Thomas, 1894). See G. M. 

Allen (1940:633).

Holotype: 241509. Skin and skull. Adult male. Hayenhsien, 

Hangchow Bay, [SL: NE] Chekiang (Zhejiang), China. Collected 10 Dec 1925 by A. de C. Sowerby. Original number 

1515.

Sciurus caniceps helvus Shamel

J. Mamm. 11: 72, 11 Feb 1930.

= Callosciurus caniceps (Gray, 1842). Unassigned to subspecies 

by Thorington and Hoffmann (2005:776), but see Moore 

and Tate (1965:196), who assigned it to C. c. caniceps 

(Gray, 1842).

Holotype: 253433. Skin and skull. Adult male. Island of Koh 

Tau, [Surat Thani], off the east coast of the Malay Peninsula, [Thailand]. Collected 17 Sep 1928 by H. M. Smith. 

Original number [SL: 2362].

Sciurus carimatae Miller

Proc. U.S. Natl. Mus. 31: 57, 23 Jul 1906a.

= Callosciurus prevostii atricapillus (Schlegel, 1863). See Corbet 

and Hill (1992:290).

Holotype: 125076. Skin and skull. Adult male. Telok Pai, Karimata Island, [Kalimantan Barat], off west coast of Borneo, 

[Indonesia]. Collected 27 Aug 1904 by W. L. Abbott. Original number 3662.

Sciurus carimonensis Miller

Proc. U.S. Natl. Mus. 31: 261, 11 Sep 1906b.

= Callosciurus prevostii (Desmarest, 1822). Unassigned to subspecies by Thorington and Hoffmann (2005:779).

Holotype: 122800. Skin and skull. Adult female [FC: 11 mi (18 

km) SW Tanjon Balie], Great Karimon Island [FC: Straits 

of Malacca], Rhio-Linga (Riau) Archipelago, [Kepulauan 

Riau, Sumatra, Indonesia]. Collected 24 May 1903 by W. L. 

Abbott. Original number 2423.

Sciurus casensis Miller

Smithsonian Misc. Coll. 45: 20, 6 Nov 1903a.

= Callosciurus caniceps casensis (Miller, 1903). See Robinson 

and Kloss (1918a:208).

Holotype: 104370. Skin and skull. Adult female. Chance Island, 

Mergui Archipelago [Tanintharyi, Myanmar]. Collected 28 

Dec 1899 by W. L. Abbott. Original number 185.

Sciurus condurensis Miller

Proc. U.S. Natl. Mus. 31: 260, 11 Sep 1906b.

= Callosciurus prevostii (Desmarest, 1822). Unassigned to subspecies by Thorington and Hoffmann (2005:779).

Holotype: 122876. Skin and skull. Adult female. Pulo Kundur, 

Rhio-Linga (Riau) Archipelago, [FC: east coast of Sumatra], 

[Kepulauan Riau, Indonesia]. Collected 13 Jun 1903 by 

W. L. Abbott. Original number 2486.

Sciurus datus Lyon

Proc. U.S. Natl. Mus. 40: 86, 25 Apr 1911a.

= Callosciurus notatus (Boddaert, 1785). Unassigned to subspecies by Thorington and Hoffmann (2005:778).

Holotype: 145393. Skin and skull. Adult female. Pulo Datu, off 

west coast of Borneo, [Kalimantan Barat], [FC: South China 

Sea], [Indonesia]. Collected 2 May 1907 by W. L. Abbott. 

Original number 5153.

Sciurus director Lyon

Proc. U.S. Natl. Mus. 36: 509, 7 Jun 1909a.

= Callosciurus notatus (Boddaert, 1785). Unassigned to subspecies by Thorington and Hoffmann (2005: 778).

Holotype: 145392. Skin and skull. Immature male. Direction 

Island, South China Sea [Kalimantan Barat, Borneo, Indonesia]. Collected 1 May 1907 by W. L. Abbott. Original 

number 5152.

Sciurus domelicus Miller

Smithsonian Misc. Coll. 45: 18, 6 Nov 1903a.

= Callosciurus caniceps domelicus (Miller, 1903). See Robinson 

and Kloss (1918a:207).

Holotype: 104381. Skin and skull. Adult female. Domel Island, 

Mergui Archipelago, [Tanintharyi, Myanmar]. Collected 24 

Feb 1900 by W. L. Abbott. Original number 322.

Sciurus dulitensis dilutus Miller

Smithsonian Misc. Coll. 61 (21): 23, 29 Dec 1913.

= Callosciurus notatus (Boddaert, 1785). Unassigned to subspecies by Thorington and Hoffmann (2005:778).

Holotype: 196712. Skin and skull. Adult female. Tanjong Batu, 

[Kalimantan Timur], Dutch Southeast Borneo, [Indonesia]. 

Collected 24 Aug 1912 by H. C. Raven. Original number 

177.

Sciurus epomophorus inexpectatus Kloss

J. Nat. Hist. Soc. Siam, 2: 178, Dec 1916.

= Callosciurus caniceps (Gray, 1842). Unassigned to subspecies 

by Thorington and Hoffmann (2005:776) but see Moore 

and Tate (1965:196) who assign it to C. c caniceps (Gray, 

1842).

Holotype: 221557. Skin and skull. Adult female. Koh Lak, Pran, 

[Prachuap Khiri Khan], southwest Siam (Thailand). Collected 15 Nov 1916 by C. B. Kloss. Original number 2434.

Sciurus erebus Miller

Proc. U.S. Natl. Mus. 26: 456, 3 Feb 1903b.

= Callosciurus prevostii piceus (Peters, 1866). See Robinson and 

Kloss (1918a:214).
number 640 • 7

Holotype: 114537. Skin and skull. Adult female. Tapanuli Bay, 

[Sumatera Utara], northwestern Sumatra, [Indonesia]. Collected 17 Mar 1902 by W. L. Abbott. Original number 1653.

Sciurus erythraeus pranis Kloss

J. Nat. Hist. Soc. Siam, 2: 178, Dec 1916.

= Callosciurus erythraeus pranis (Kloss, 1916). See Thorington 

and Hoffmann (2005:776).

Holotype: 221568. Skin and skull. Adult male. Koh Lak, Pran, 

[Prachuap Khiri Khan], southwest Siam (Thailand). Collected 9 Nov 1916 by C. B. Kloss. Original number 2395.

Sciurus finlaysoni trotteri Kloss

J. Nat. Hist. Soc. Siam, 2: 178, Dec 1916.

= Callosciurus finlaysonii trotteri (Kloss, 1916). See Robinson 

and Kloss (1918a:203).

Holotype: 236601. Skin and skull. Adult male. Koh Lan [SL: Island], [Chon Buri], inner Gulf of Siam (Thailand). Collected 

29 Oct 1916 by C. B. Kloss. Original number 2266.

Sciurus ictericus Miller

Smithsonian Misc. Coll. 45: 12, 6 Nov 1903a.

= Callosciurus notatus vittatus (Raffles, 1821). See Thorington 

and Hoffmann (2005:778).

Holotype: 121727. Skin and skull. Adult female. Tana Bala 

(Pulau Tanabala), Batu Islands [Sumatera Utara, Sumatra, 

Indonesia]. Collected 4 Feb 1903 by W. L. Abbott. Original 

number 2223.

Sciurus klossii Miller

Proc. Washington Acad. Sci. 2: 225, 20 Aug 1900a.

= Callosciurus nigrovittatus klossii (Miller, 1900). See Robinson 

and Kloss (1918a: 223).

Holotype: 101678. Skin and skull. Adult male. Kaju Ara, or Saddle Island (Pulau Mendarik), [SL: near] Tambelan group, 

[Kepulauan Riau, Indonesia]. Collected 15 Aug 1899 by 

W. L. Abbott. No original number.

Sciurus lamucotanus Lyon

Proc. U.S. Natl. Mus. 40: 85, 25 Apr 1911a.

= Callosciurus notatus (Boddaert, 1785). Unassigned to subspecies by Thorington and Hoffmann (2005:778).

Holotype: 145405. Skin and skull. Adult female. Pulo Lamukotan, [Kalimantan Barat], [FC: about 10 mi (16 km) off 

west coast of Borneo, [Indonesia]. Collected 8 May 1907 by 

W. L. Abbott. Original number 5201.

Sciurus lancavensis Miller

Smithsonian Misc. Coll. 45: 16, 6 Nov 1903a.

= Callosciurus caniceps (Gray, 1842). Unassigned to subspecies by 

Thorington and Hoffmann (2005:776) but see Moore and 

Tate (1965:187) who assign it to C. c. concolor (Blyth, 1856).

Holotype: 104390. Skin and skull. Adult male. Pulo (Pulau) 

Lankawi, off west coast of Malay Peninsula, about 75 mi 

(121 km) north of Penang, [Kedah, Malaysia]. Collected 1 

Dec 1899 by W. L. Abbott. Original number 101.

Sciurus lautensis Miller

Proc. Washington Acad. Sci. 3: 128, 26 Mar 1901a.

= Callosciurus notatus (Boddaert, 1785). Unassigned to subspecies by Thorington and Hoffmann (2005:778).

Holotype: 104683. Skin and skull. Adult female. Pulo Laut, 

North Natuna Islands, South China Sea, between Malay 

Peninsula and Borneo, [Kepulauan Riau, Indonesia]. Collected 6 Aug 1900 by W. L. Abbott. Original number 612.

Sciurus lucas Miller

Smithsonian Misc. Coll. 45: 20, 6 Nov 1903a.

= Callosciurus caniceps (Gray, 1842). Unassigned to subspecies 

by Thorington and Hoffmann (2005:776) but see Moore 

and Tate (1965:190), who assign it to C. c. bimaculatus

(Temminck, 1853).

Holotype: 104385. Skin and skull. Adult female. St. Luke Island, 

Mergui Archipelago, [Tanintharyi, Myanmar]. Collected 20 

Jan 1900 by W. L. Abbott. Original number 256.

Sciurus lutescens Miller

Proc. Washington Acad. Sci. 3: 124, 26 Mar 1901a.

= Callosciurus notatus (Boddaert, 1785). Unassigned to subspecies by Thorington and Hoffmann (2005:778).

Holotype: 104668. Skin and skull. Adult male. Sirhassen Island, 

South Natuna Islands, [Kepulauan Riau, Indonesia]. Collected 3 Jun 1900 by W. L. Abbott. Original number 429.

Sciurus marinsularis Lyon

Proc. U.S. Natl. Mus. 40: 89, 25 Apr 1911a.

= Callosciurus notatus (Boddaert, 1785). Unassigned to subspecies by Thorington and Hoffmann (2005:778).

Holotype: 151777. Skin and skull. Adult male. Pulo Laut, [FC: 

Saratok], off southeastern Borneo, [Kalimantan Selatan, Indonesia]. Collected 17 Dec 1907 by W. L. Abbott. Original 

number 5619.

Sciurus matthaeus Miller

Smithsonian Misc. Coll. 45: 19, 6 Nov 1903a.

= Callosciurus caniceps (Gray, 1842). Unassigned to subspecies 

by Thorington and Hoffmann (2005:776) but see Moore 

and Tate (1965:190) who assign it to C. c. bimaculatus

(Temminck, 1853).

Holotype: 111920. Skin and skull. Adult female. St. Matthew 

Island, Mergui Archipelago, [Tanintharyi, Myanmar]. 

Collected 11 Dec 1900 by W. L. Abbott. Original number 

774.

Sciurus melanops Miller

Proc. Acad. Nat. Sci. Philadelphia 54: 151, 11 Jun 1902.

= Callosciurus prevostii melanops (Miller, 1902). See Robinson 

and Kloss (1918a: 211).
8 • smithsonian contributions to zoology

Holotype: 113152. Skin and skull. Adult female. Indragiri River, 

[FC: at Tjenako], [Riau], eastern Sumatra, [Indonesia]. Collected 15 Sep 1901 by W. L. Abbott. Original number 1307.

Sciurus melanops penialius Lyon

Proc. U.S. Natl. Mus. 34: 637, 14 Sep 1908.

= Callosciurus prevostii melanops (Miller, 1902). See Thorington 

and Hoffmann (2005:779).

Holotype: 144364. Skin and skull. Adult female. Pulo Penjalei, 

at the mouth of the Kampar River, [Riau], eastern Sumatra, 

[Indonesia]. Collected 5 Feb 1907 by W. L. Abbott. Original 

number 5040.

Sciurus mendanauus Lyon

Proc. U.S. Natl. Mus. 31: 589, 18 Dec 1906a.

= Callosciurus prevostii (Desmarest, 1822). Unassigned to subspecies by Thorington and Hoffmann (2005:779).

Holotype: 124916. Skin and skull. Adult male. Pulo Mendanau, 

[FC: Kampong Petaling], west of Billiton Island, Malay Archipelago, [Kepulauan Bangka Belitung, Sumatra, Indonesia]. 

Collected 14 Jul 1904 by W. L. Abbott. Original number 3475.

Sciurus mimellus Miller

Proc. Washington Acad. Sci. 2: 218, 20 Aug 1900a.

= Callosciurus prevostii (Desmarest, 1822). Unassigned to subspecies by Thorington and Hoffmann (2005:779).

Holotype: 101668. Skin and skull. Adult male. Pulo Wai (Pulau 

Uwi), Tambelan Islands, South China Sea, between Malay 

Peninsula and Borneo, [Kepulauan Riau, Indonesia]. Collected 13 Aug 1899 by W. L. Abbott. No original number.

Sciurus mimiculus Miller

Proc. Washington Acad. Sci. 2: 219, 20 Aug 1900a.

= Callosciurus prevostii (Desmarest, 1822). Unassigned to subspecies by Thorington and Hoffmann (2005:779).

Holotype: 101616. Skin and skull. Adult male. Ste. (Saint) Barbe 

Island (Pulau Penjantan), South China Sea, between Sumatra and Borneo, [Kepulauan Riau, Indonesia]. Collected 1 

Aug 1899 by W. L. Abbott. No original number.

Sciurus notatus miniatus Miller

Proc. Washington Acad. Sci. 2: 79, 25 Jul 1900b.

= Callosciurus notatus miniatus (Miller, 1900). See Corbet and 

Hill (1992:291).

Holotype: 88415. Skin and skull. Adult female. On or near 

Khow Nom Plu, Trong (Trang), [SL: about 1,000 ft (305 

m)], Lower Siam (Thailand). Collected 25 Feb 1897 by 

W. L. Abbott. No original number.

Sciurus nyx Lyon

Proc. U.S. Natl. Mus. 34: 638, 14 Sep 1908.

= Callosciurus prevostii melanops (Miller, 1902). See Thorington 

and Hoffmann (2005:779).

Holotype: 143392. Skin and skull. Adult female. Pulo Rupat, off 

east coast of Sumatra, [Riau, Indonesia]. Collected 10 Mar 

1906 by W. L. Abbott. Original number 4691.

Sciurus pannovianus Miller

Smithsonian Misc. Coll. 45: 11, 6 Nov 1903a.

= Callosciurus notatus (Boddaert, 1785). Unassigned to subspecies by Thorington and Hoffmann (2005:778).

Holotype: 112351. Skin and skull. Adult male. Pulo Pannow 

(Penau), Atas (Badas) Islands , South China Sea, [Kepulauan 

Riau, Indonesia]. Collected 28 May 1901 by W. L. Abbott. 

Original number 952.

Sciurus pelapius Lyon

Proc. U.S. Natl. Mus. 40: 82, 25 Apr 1911a.

= Callosciurus prevostii atricapillus (Schlegel, 1863). See Corbet 

and Hill (1992:290).

Holotype: 145417. Skin and skull. Adult female. Pulo Pelapis, 

South Island, off west coast of Borneo, [Kalimantan Barat, 

Indonesia]. Collected 30 May 1907 by W. L. Abbott. Original number 5335.

Sciurus pemangilensis Miller

Smithsonian Misc. Coll. 45: 9, 6 Nov 1903a.

= Callosciurus notatus miniatus (Miller, 1900). See Corbet and 

Hill (1992:291).

Holotype: 112460. Skin and skull. Adult female. Pulo Pemangil, 

off coast of Johore (Johor), [SL and FC: South China Sea], 

[Malaysia]. Collected 12 Jun 1901 by W. L. Abbott. Original number 1062.

Sciurus peninsularis Miller

Smithsonian Misc. Coll. 45: 10, 6 Nov 1903a.

= Callosciurus notatus miniatus (Miller, 1900). See Corbet and 

Hill (1992:291).

Holotype: 112511. Skin and skull. Adult male. North bank of 

Endau River, Pahang, [Malaysia]. Collected 21 Jun 1901 by 

W. L. Abbott. Original number 1078.

Sciurus poliopus Lyon

Proc. U.S. Natl. Mus. 40: 88, 25 Apr 1911a. Primary junior 

homonym of Sciurus variegatus poliopus Fitzinger, 1867. 

Replaced with Sciurus conipus by Lyon (1911b:98).

= Callosciurus notatus (Boddaert, 1785). Unassigned to subspecies by Thorington and Hoffmann (2005:778).

Holotype: 151789. Skin and skull. Adult male. Pamukang (Pamukan) Bay, [Kalimantan Selatan], southern Borneo, [Indonesia]. 

Collected 5 Apr 1908 by W. L. Abbott. Original number 5923.

Sciurus pretiosus Miller

Proc. U.S. Natl. Mus. 26: 454, 3 Feb 1903b.

= Callosciurus notatus vittatus (Raffles, 1821). See Corbet and 

Hill (1992:292).
number 640 • 9

Holotype: 114325. Skin and skull. Adult female. Pulo Bangkaru, 

Banjak Islands, [Aceh, Sumatra, Indonesia]. Collected 20 

Jan 1902 by W. L. Abbott. Original number 1442.

Sciurus proserpinae Lyon

Smithsonian Misc. Coll. 48: 275, 4 Feb 1907b.

= Callosciurus prevostii atricapillus (Schlegel, 1863). See Corbet 

and Hill (1992:290).

Holotype: 142285. Skin and skull. Adult female. Pulo Temaju, 

about 4 mi (6.5 km) off west coast of Borneo, [Kalimantan 

Barat, Indonesia]. Collected 9 Jun 1905 by W. L. Abbott. 

Original number 4180.

Sciurus rubeculus Miller

Smithsonian Misc. Coll. 45: 22, 6 Nov 1903a.

= Callosciurus erythraeus rubeculus (Miller, 1903). See Robinson and Kloss (1918a:199).

Holotype: 86777. Skin and skull. Adult male. Khow Sai Dow, 

altitude 1,000 ft (305 m), Trong (Trang), Lower Siam (Thailand). Collected 21 Feb 1899 by W. L. Abbott. No original 

number.

Sciurus rubidiventris Miller

Proc. Washington Acad. Sci. 3: 127, 26 Mar 1901a.

= Callosciurus notatus (Boddaert, 1785). Unassigned to subspecies by Thorington and Hoffmann (2005:778).

Holotype: 104671. Skin and skull. Adult female. Bunguran Island, North Natunas (Natuna Islands), [Kepulauan Riau, 

Indonesia]. Collected 22 Jun 1900 by W. L. Abbott. Original number 498.

Sciurus rutiliventris Miller

Proc. Washington Acad. Sci. 3: 126, 26 Mar 1901a.

= Callosciurus notatus (Boddaert, 1785). Unassigned to subspecies by Thorington and Hoffmann (2005:778).

Holotype: 104658. Skin and skull. Adult male. Pulo Midei 

(Pulau Midai), Low Island, South Natuna Islands, [Kepulauan Riau, Indonesia]. Collected 24 May 1900 by W. L. 

Abbott. Original number 405.

Sciurus sanggaus Lyon

Proc. U.S. Natl. Mus. 33: 554, 24 Dec 1907a.

= Callosciurus prevostii atricapillus (Schlegel, 1863). See Corbet 

and Hill (1992:290).

Holotype: 142296. Skin and skull. Adult female. Sanggau, south 

bank of Kapuas River, [Kalimantan Barat], western Borneo, 

[Indonesia]. Collected 21 Aug 1905 by W. L. Abbott. Original number 4357.

Sciurus saturatus Miller

Proc. U.S. Natl. Mus. 26: 453, 3 Feb 1903b.

= Callosciurus notatus vittatus (Raffles, 1821). See Corbet and 

Hill (1992:292).

Holotype: 114629. Skin and skull. Adult female. Pulo Mansalar 

(Pulau Musala), off Tapanuli Bay, [Sumatera Utara], Sumatra, [Indonesia]. Collected 9 Mar 1902 by W. L. Abbott. 

Original number 1633.

Sciurus seraiae Miller

Proc. Washington Acad. Sci. 3: 125, 26 Mar 1901a.

= Callosciurus notatus (Boddaert, 1785). Unassigned to subspecies by Thorington and Hoffmann (2005:778).

Holotype: 104660. Skin and skull. Adult male. Pulo Seraia, 

South Natuna Islands, [Kepulauan Riau, Indonesia]. Collected 29 May 1900 by W. L. Abbott. Original number 415.

Sciurus serutus Miller

Proc. U.S. Natl. Mus. 31: 58, 23 Jul 1906a.

= Callosciurus notatus (Boddaert, 1785). Unassigned to subspecies by Thorington and Hoffmann (2005:778).

Holotype: 125025. Skin and skull. Adult male. Pulo Serutu, 

Karimata Islands, [Kalimantan Barat, Borneo, Indonesia]. 

Collected 17 Aug 1904 by W. L. Abbott. Original number 

3584.

Sciurus siriensis Lyon

Proc. U.S. Natl. Mus. 40: 87, 25 Apr 1911a.

= Callosciurus notatus (Boddaert, 1785). Unassigned to subspecies by Thorington and Hoffmann (2005:778).

Holotype: 151768. Skin and skull. Adult male. Pulo Mata Siri, 

Java Sea, [Kalimantan Selatan, Borneo, Indonesia]. Collected 7 Dec 1907 by W. L. Abbott. Original number 5580.

Sciurus sullivanus Miller

Smithsonian Misc. Coll. 45: 17, 6 Nov 1903a.

= Callosciurus caniceps (Gray, 1842). Unassigned to subspecies 

by Thorington and Hoffmann (2005:776) but see Moore 

and Tate (1965:190) who assign it to C. c. bimaculatus

(Temminck, 1853).

Holotype: 104377. Skin and skull. Adult female. Sullivan Island, 

Mergui Archipelago, [Tanintharyi, Myanmar]. Collected 1 

Feb 1900 by W. L. Abbott. Original number 294.

Sciurus tedongus Lyon

Proc. U.S. Natl. Mus. 31: 591, 18 Dec 1906a.

= Callosciurus notatus vittatus (Raffles, 1821). See Corbet and 

Hill (1992:292).

Holotype: 124717. Skin and skull. Adult male. Tanjong Tedong, 

island of Banka (Bangka), [Kepulauan Bangka Belitung], 

east coast of Sumatra, [Indonesia]. Collected 1 Jun 1904 by 

W. L. Abbott. Original number 3285.

Sciurus tenuirostris Miller

Proc. Washington Acad. Sci. 2: 221, 20 Aug 1900a.

= Callosciurus notatus miniatus (Miller, 1900). See Corbet and 

Hill (1992:291).
1 0 • smithsonian contributions to zoology

Holotype: 101753. Skin and skull. Adult female. Tioman Island, 

[Pahang, Malaysia]. Collected 30 Sep 1899 by W. L. Abbott. 

No original number.

Sciurus ubericolor Miller

Proc. U.S. Natl. Mus. 26: 455, 3 Feb 1903b.

= Callosciurus notatus vittatus (Raffles, 1821). See Corbet and 

Hill (1992:292).

Holotype: 114373. Skin and skull. Adult female. [FC: Ujong 

Tambaga, east side] Pulo Tuangku, Banjak Islands, [Aceh, 

Sumatra, Indonesia]. Collected 5 Feb 1902 by W. L. Abbott. 

Original number 1517.

Sciurus vittatus rupatius Lyon

Proc. U.S. Natl. Mus. 34: 640, 14 Sep 1908.

= Callosciurus notatus vittatus (Raffles, 1821). See Corbet and 

Hill (1992:292).

Holotype: 143406. Skin and skull. Adult female. Pulo Rupat, 

[Riau], off east coast of Sumatra, [Indonesia]. Collected 17 

Mar 1906 by W. L. Abbott. Original number 4733.

Sciurus vittatus tapanulius Lyon

Smithsonian Misc. Coll. 48: 280, 4 Feb 1907c.

= Callosciurus notatus vittatus (Raffles, 1821). See Corbet and 

Hill (1992:292).

Holotype: 114519. Skin and skull. Adult male. Tapanuli Bay, 

[Sumatera Utara], west coast of Sumatra, [Indonesia]. Collected 21 Feb 1902 by W. L. Abbott. Original number 1560.

Sciurus vittatus tarussanus Lyon

Smithsonian Misc. Coll. 48: 279, 4 Feb 1907c.

= Callosciurus notatus vittatus (Raffles, 1821). See Corbet and 

Hill (1992:292).

Holotype: 141038. Skin and skull. Adult female. [FC: Njalo], 

Tarussan (Tarusan) Bay, [Sumatera Barat], west coast of Sumatra, [Indonesia]. Collected 28 Dec 1904 by W. L. Abbott. 

Original number 3857.

Genus Callospermophilus Merriam, 1897

Callospermophilus chrysodeirus connectens A. H. Howell

J. Mamm. 12: 161, 14 May 1931.

= Callospermophilus lateralis connectens A. H. Howell, 1931. 

See Helgen et al. (2009:285).

Holotype: 212461. Skin and skull. Adult male. Homestead, [SL 

and FC: altitude 1,800 ft (549 m)], [Baker County], Oregon. 

Collected 1 Jun 1916 by H. H. Sheldon. Original number 

535.

Callospermophilus chrysodeirus mitratus A. H. Howell

J. Mamm. 12: 161, 14 May 1931.

= Callospermophilus lateralis mitratus A. H. Howell, 1931. See 

Helgen et al. (2009:285).

Holotype: 138125. Skin and skull. Adult male. South Yolla Bolly 

Mountain, [Tehama County] California. Collected 30 Jul 

1905 by J. F. Ferry. Original number 13.

Callospermophilus chrysodeirus trinitatis Merriam

Proc. Biol. Soc. Washington 14: 126, 19 Jul 1901a.

= Callospermophilus lateralis trinitatis Merriam, 1901. See Helgen et al. (2009:285).

Holotype: 95531. Skin and skull. Adult female. Trinity Mountains east of Hoopa Valley, [FC: summit east of Hoopa at 

Little Lake], altitude 5,700 ft (1737 m), [Humboldt County] 

California. Collected 10 Sep 1898 by V. Bailey. Original 

number 6693.

Callospermophilus lateralis arizonensis V. Bailey

Proc. Biol. Soc. Washington 26: 130, 21 May 1913a.

Holotype: 17527/24463. Skin and skull. Adult male. San Francisco Mountain (Peaks), [Coconino County], Arizona. Collected 8 Aug 1889 by C. H. Merriam and V. Bailey. Original 

number [FC: VB] 308.

Callospermophilus lateralis caryi A. H. Howell

Proc. Biol. Soc. Washington 30: 105, 23 May 1917.

= Callospermophilus lateralis castanurus (Merriam, 1890). See 

Helgen et al. (2009:285).

Holotype: 176826. Skin and skull. Adult female. Seven mi (11 

km) south of Fremont Peak, Wind River Mountains (Range), 

altitude 10,400 ft (3,170 m), [Fremont County], Wyoming. 

Collected 19 Jul 1911 by M. Cary. Original number 2211.

Callospermophilus lateralis certus Goldman

J. Mamm. 2: 232, 29 Nov 1921.

Holotype: 208891. [Skin and skull]. Adult male. North base of 

Charleston Peak, [Clark County], Nevada. Collected 29 Jun 

1915 by L. J. Goldman. Original number 2270.

Callospermophilus lateralis tescorum Hollister

Smithsonian Misc. Coll. 56(26): 2, 5 Dec 1911a.

Holotype: 174165. Skin and skull. Adult male. Head of Moose 

Pass branch of Smoky River, near Moose Pass, British Columbia, altitude 7,000 ft (2,134 m), Alberta, [Canada]. Collected 2 Aug 1911 by N. Hollister. Original number 3863.

Callospermophilus madrensis Merriam

Proc. Washington Acad. Sci. 3: 563, 29 Nov 1901b.

Holotype: 95363. [Skin and skull]. Adult female. Sierra Madre, 

near Guadalupe y Calvo, altitude 7,000 ft (2,134 m), Chihuahua, Mexico. Collected 27 Aug 1898 by E. W. Nelson 

and E. A. Goldman. Original number 12923.

Spermophilus chrysodeirus brevicaudus Merriam

Proc. Biol. Soc. Washington 8: 134, 28 Dec 1893a. Primary junior homonym of Spermophilus brevicauda Brandt, 1844. 
number 640 • 1 1

Replaced with Spermophilus (Callospermophilus) bernardinus by Merriam (1898a:782).

= Callospermophilus lateralis bernardinus (Merriam, 1898). See 

Helgen et al. (2009:285).

Holotype: 56661. Skin and skull. Adult female. San Bernardino 

Peak, [FC: altitude 9,500 ft (2,896 m)], [San Bernardino 

County], California. Collected 9 Oct 1893 by J. E. McLellan. Original number 274.

Tamias castanurus Merriam

N. Amer. Fauna 4: 19, 8 Oct 1890a.

= Callospermophilus lateralis castanurus (Merriam, 1890). See 

Helgen et al. (2009:285).

Holotype: 22733/30197. Skin and skull. Adult male. Park City, 

Wasatch Mountains, altitude 7,000 ft (2,134 m), [Summit 

County], Utah. Collected 3 Jul 1890 by V. Bailey. Original 

number 1383.

Tamias chrysodeirus Merriam

N. Amer. Fauna 4: 19, 8 Oct 1890a.

= Callospermophilus lateralis chrysodeirus (Merriam, 1890). See 

Helgen et al. (2009:285).

Holotype: [186464]. Skin and skull. Adult male. Fort Klamath, [Klamath County], Oregon. Collected 31 Jul 1888 by 

S. Parker. Original number 143. Merriam collection number 

4966/5720.

Tamias cinerascens Merriam

N. Amer. Fauna 4: 20, 8 Oct 1890a.

= Callospermophilus lateralis cinerascens (Merriam, 1890). See 

Helgen et al. (2009:285).

Holotype: [186465]. Skin and skull. Adult female. Helena, altitude 4,500 ft (1,372 m), [Lewis and Clark County], Montana. Collected 13 Aug 1888 by C. H. Merriam. Original 

number 4. Merriam Collection number 4525/5177.

Genus Cynomys Rafinesque, 1817

Cynomys gunnisoni zuniensis Hollister

N. Amer. Fauna 40: 32, 20 Jun 1916.

Holotype: 137555. Skin and skull. Adult male. Wingate, McKinley County, New Mexico. Collected 26 Jun 1905 by 

N. Hollister. Original number 2374.

Cynomys leucurus Merriam

N. Amer. Fauna 3: 59, 11 Sep 1890b. See Merriam (1890c:33) 

for complete description.

Holotype: [186472]. Skin and skull. Adult female. Fort Bridger, 

[Uinta County], Wyoming. Collected 15 Sep 1888 by V. Bailey. 

Original number 224. Merriam collection number 4668/5319.

Cynomys mexicanus Merriam

Proc. Biol. Soc. Washington 7: 157, 27 Jul 1892a.

Holotype: 26423/33836. Skin and skull. Adult male. La Ventura, 

[FC: about 5,600 ft (1,707 m)], Coahuila, Mexico. Collected 24 Mar 1891 by C. P. Streator. Original number 625.

Spermophilus gunnisoni Baird

Proc. Acad. Nat. Sci. Philadelphia 7: 334, reported favorably for 

publication 24 Apr 1855a, reciept acknowledged 16 Jun 

1855 by the American Philosophical Society.

= Cynomys gunnisoni gunnisoni (Baird, 1855). See Hollister 

(1916:29).

Holotype: [501/1636. Skin and skull. Young adult]. Cochetopa 

Pass, [Saguache County, Colorado]. Collected [Lyon and 

Osgood (1909:161): first week of September 1853] by 

F. Kreutzfeldt. Original number [MC: Lt. E. G. Beckwith 22].

Type specimen not designated in the original description, but it is 

specified by number in Baird (1857:337).

Genus Dremomys Heude, 1898

Dremomys rufigenis lentus A. B. Howell

J. Washington Acad. Sci. 17: 80, 19 Feb 1927.

= Dremomys pernyi pernyi (Milne-Edwards, 1867). See G. M. 

Allen (1940:647).

Holotype: 240384. Skin and skull. Adult male. Near Wenchuanshein, altitude 6,000 ft (1,829 m), Szechwan (Sichuan), 

China. Collected 14 [SL and FC: 12] Aug 1924 by D. C. 

Graham. Original number 14.

Genus Exilisciurus Moore, 1958

Nannosciurus surrutilus Hollister

Proc. U.S. Natl. Mus. 46: 313, 31 Dec 1913.

= Exilisciurus concinnus (Thomas, 1888). See Heaney (1985:7).

Holotype: 144641. Skin and skull. Adult female. Summit [SL: of] 

Mount Bliss, altitude 5,750 ft (1,753 m), Mindanao, [SL: 

Philippine Islands] (Philippines). Collected 28 May 1906 by 

E. A. Mearns. Original number 6170.

Genus Glaucomys Thomas, 1908

Glaucomys bullatus A. H. Howell

Proc. Biol. Soc. Washington 28: 113, 27 May 1915a.

= Glaucomys sabrinus bangsi (Rhoads, 1897). See Davis (1939:231).

Holotype: 24271/31675. Skin and skull. Adult female. Sawtooth 

(Alturas) Lake, [FC: altitude 7,200 ft (2,195 m)], [Poole 

and Schantz (1942:534): east base of Sawtooth Mountains, 

Blaine County], Idaho. Collected 28 Sep 1890 by V. Bailey 

and B. H. Dutcher. Original number [FC: VB] 1883.

Glaucomys sabrinus coloratus Handley

Proc. Biol. Soc. Washington 66: 191, 2 Dec 1953.

Holotype: 292278. Skin and skull. Young adult female. Bald Knob, 

3.5 mi (5.6 km) south of the summit of Mount Mitchell, [SL: 
1 2 • smithsonian contributions to zoology

200 yds (183 m) below Bald Knob Cabin], altitude 5,000 ft 

(1,524 m), Yancey County, North Carolina. Collected 1 Mar 

1951 by M. G. Edwards. Original number [SL: 2].

Glaucomys sabrinus columbiensis A. H. Howell

Proc. Biol Soc. Washington 28: 111, 27 May 1915a.

Holotype: 94310. Skin and skull. Subadult male. Okanagan, 

British Columbia, [Canada]. Collected 9 May 1898 by A. C. 

Brooks. Original number 1214, [SL: 1762-X].

Glaucomys sabrinus fuscus Miller

Proc. Biol. Soc. Washington 49: 143, 22 Aug 1936.

Holotype: 260420. Skin and skull. Adult female. Cranberry 

Glades, altitude 3,300 ft (1,006 m), Pocahontas County, 

West Virginia. Collected 13 Jun 1936 by W. Perrygo and 

C. Lingebach. Original number 109.

Glaucomys sabrinus griseifrons A. H. Howell

J. Mamm. 15: 64, 15 Feb 1934

Holotype: 256993. Skin and skull. Adult female. Lake Bay, [SL: 

south of Wrangell], Prince of Wales Island, Alaska. Collected 18 Apr 1927 by F. R. Bates. Original number 10.

Glaucomys sabrinus latipes A. H. Howell

Proc. Biol. Soc. Washington 28: 112, 27 May 1915a.

Holotype: 68753. [Skin and skull]. Adult female. Glacier [FC: 

House, 44 mi (71 km) east of Revelstoke], British Columbia, [Canada]. Collected 13 Aug 1894 by J. A. Loring. Original number 2111.

Glaucomys volans guerreroensis Diersing

Southwestern Nat. 25: 162, 15 May 1980.

Holotype: 329704. Skin and skull. Adult male. One mile (1.6 

km) SW Omilteme, altitude 7,300 ft (2,225 m) Guerrero, 

Mexico. Collected 16 Apr 1963 by A. D. Stock. Original 

number 1149.

Glaucomys volans herreranus Goldman

J. Washington Acad. Sci. 26: 463, 15 Nov 1936a.

Holotype: 261695. Skin and skull. Young adult female. Mountains of Veracruz, Mexico. Collected 8 Nov 1924. Prepared 

by L. G. Rubio. No original number. 27589-X.

Glaucomys volans madrensis Goldman

J. Washington Acad. Sci. 26: 463, 15 Nov 1936a.

Holotype: 261694. Skin and skull. Adult, sex undetermined. Sierra Madre, Chihuahua, Mexico. Received Mar 1926 from 

A. L. Herrera. Original number [SL: 2]. 27588-X.

Glaucomys volans saturatus A. H. Howell

Proc. Biol. Soc. Washington 28: 110, 27 May 1915a.

Holotype: 178366. [Skin and skull]. Adult female. Dothan, 

[Houston County], Alabama. Collected 13 Mar 1912 by 

A. H. Howell. Original number 1960.

Glaucomys volans texensis A. H. Howell

Proc. Biol. Soc. Washington 28: 110, 27 May 1915a.

Holotype: 136400. [Skin and skull]. Adult male. [FC: Big Thicket], 

7 mi (11 km) northeast of Sour Lake, [Hardin County], Texas. 

Collected 15 Mar 1905 by J. H. Gaut. Original number 3480.

Sciuropterus alpinus klamathensis Merriam

Proc. Biol. Soc. Washington 11: 225, 15 Jul 1897a.

= Glaucomys sabrinus klamathensis (Merriam, 1897). See A. H. 

Howell (1918:52).

Holotype: 87310. [Skin and skull]. Adult female. Fort Klamath, 

altitude 4,200 ft (1,280 m), [Klamath County], Oregon. 

Collected 11 [XC and MCC: 17, which is correct] Jan 1897 

by B. L. Cunningham. Original number [XC: 25]. 355-X.

Sciuropterus alpinus zaphaeus Osgood

Proc. Biol. Soc. Washington 18: 133, 18 Apr 1905.

= Glaucomys sabrinus zaphaeus (Osgood, 1905). See A. H. 

Howell (1918: 43).

Holotype: 136137. [Skin and skull]. Adult female. Helm Bay, 

Cleveland Peninsula, southeastern Alaska. Collected 21 Jan 

1905 by C. Catt. No original number. [SL: 5196-X].

Sciuropterus oregonensis stephensi Merriam

Proc. Biol. Soc. Washington 13: 151, 13 Jun 1900a.

= Glaucomys sabrinus stephensi (Merriam, 1900). See A. H. 

Howell (1918: 57).

Holotype: 99830. [Skin and skull]. Young adult female. Sherwood [SL and XC: s], altitude 2,500 ft (762 m), Mendocino 

County, California. Collected 10 May 1894 by F. Stephens. 

Original number 2307 [SL: 2624-X].

Sciuropterus sabrinus macrotis Mearns

Proc. U.S. Natl. Mus. 21: 353, 4 Nov 1898a.

= Glaucomys sabrinus macrotis (Mearns, 1898). See A. H. Howell (1915a:111).

Holotype: 83152. [Skin and skull]. Adult female. Hunter Mountain, altitude 3,300 ft (1,006 m), Catskills (Catskill Mountains), Greene County, New York. Collected 31 Aug 1896 

by E. A. Mearns. Original number 4036.

Sciuropterus volans goldmani Nelson

Proc. Biol. Soc. Washington 17: 148, 6 Oct 1904.

= Glaucomys volans goldmani (Nelson, 1904). See A. H. Howell 

(1918:28).

Holotype: 132833. Skin and skull. Adult male. Twenty miles (32 

km) southeast of Teopisca, Chiapas, Mexico. Collected 8 

Apr 1904 by E. A. Goldman. Original number [FC: Nelson 

and Goldman] 16667.

Sciuropterus yukonensis Osgood

N. Amer. Fauna 19: 25, 6 Oct 1900a.

= Glaucomys sabrinus yukonensis (Osgood, 1900). See A. H. 

Howell (1918:41).
number 640 • 1 3

Holotype: 19909/35320. Skin and skull. Adult female. Camp 

Davidson, Yukon River, near Alaska-Canada boundary, 

Yukon, Canada. Collected 8 Dec 1890 by R. E. Carson. No 

original number.

Genus Heliosciurus Trouessart, 1880

Heliosciurus multicolor madogae Heller

Smithsonian Misc. Coll. 56 (17): 1, 28 Feb 1911.

= Heliosciurus gambianus madogae Heller, 1911. See Hayman 

and Holt (1940:402).

Holotype: 164828. [Skin and skull]. Adult male. Uma, 50 mi (80 

km) north of Nimule, Uganda (South Sudan). Collected 11 

Feb 1910 by K. Roosevelt. Original number J. A. Loring 8991.

Heliosciurus rufobrachiatus shindi Heller

Smithsonian Misc. Coll. 63 (7): 7, 24 Jun 1914.

= Heliosciurus undulatus (True, 1892). See Grubb (1982:201).

Holotype: 182768. [Skin and skull]. Adult male. Summit of 

Mount Umengo, altitude 6,000 ft (1,829 m), Taiti (Taita) 

Hills, [Coast Province], British East Africa (Kenya). Collected 11 Nov 1911 by E. Heller. Original number 4731.

Sciurus (Heliosciurus) rufobrachiatus libericus Miller

Proc. Washington Acad. Sci. 2: 633, 28 Dec 1900c.

= Heliosciurus rufobrachium maculatus (Temminck, 1853). See 

Thomas (1923:524).

Holotype: 83834. Skin and skull. Adult male. Mount Coffee, 

[Montserrado], Liberia, Africa. Collected 22 Mar 1897 by 

R. P. Currie. Original number 16.

Sciurus undulatus True

Proc. U.S. Natl. Mus. 15: 465, 26 Oct 1892.

= Heliosciurus undulatus (True, 1892). See Hollister (1919:11).

Lectotype: 19005/34731. Skin and skull. Male. Mount KilimaNjaro (Kilimanjaro), altitude 6,000 ft (1,829 m) [Kilimanjaro, Tanzania]. Collected 12 Jun 1888 by W. L. Abbott. 

[MC: Original number 7]. Lectotype selected by Lyon and 

Osgood (1909:196).

Genus Hylopetes Thomas, 1908

Sciuropterus amoenus Miller

Proc. U.S. Natl. Mus. 31: 264, 11 Sep 1906b.

= Hylopetes spadiceus spadiceus (Blyth, 1847). See Corbet and 

Hill (1992:315).

Holotype: 122883. Skin and skull. Adult male. Pulo Kundur, 

Rhio-Linga (Riau) Archipelago, [Kepulauan Riau], [FC: 

east coast Sumatra], [Indonesia]. Collected 12 Jun 1903 by 

W. L. Abbott. Original number [SL: 2483].

Genus Ictidomys J. A. Allen, 1877

Citellus tridecemlineatus arenicola A. H. Howell

Proc. Biol. Soc. Washington 41: 213, 18 Dec 1928.

= Ictidomys tridecemlineatus arenicola (A. H. Howell, 1928). 

See Helgen et al. (2009:291).

Holotype: 87686. [Skin and skull]. Adult male. Pendennis, [FC: 

Lane County], Kansas. Collected 22 Apr 1897 by J. A. Loring. Original number 3988.

Citellus tridecemlineatus hollisteri V. Bailey

Proc. Biol. Soc. Washington 26: 131, 21 May 1913a. 

= Ictidomys tridecemlineatus hollisteri (V. Bailey, 1913). See Helgen et al. (2009:291).

Holotype: 119025. Skin and skull. Adult female. Elk Valley, 

Mescalero Indian Reservation, Sacramento Mountains, altitude 8,000 ft (2,438 m), [Otero County], New Mexico. 

Collected 11 Sep 1902 by V. Bailey. Original number 7963.

Citellus tridecemlineatus monticola A. H. Howell

Proc. Biol. Soc. Washington 41: 214, 18 Dec 1928.

= Ictidomys tridecemlineatus monticola (A. H. Howell, 1928). 

See Helgen et al. (2009:291).

Holotype: 209255. Skin and skull. Adult male. Marsh Lake (also 

called Big Lake), White Mountains, altitude 9,000 ft (2,743 

m), [Apache County], Arizona. Collected 15 Jun 1915 by 

E. A. Goldman. Original number 22626.

Spermophilus mexicanus parvidens Mearns

Proc. U.S. Natl. Mus. 18: 1, 1 Mar 1896a. Preprint of Proc. U.S. 

Natl. Mus. 18: 443, 23 May 1896b.

= Ictidomys parvidens (Mearns, 1896). See Helgen et al. 

(2009:290).

Holotype: 63073. Skin and skull. Adult male. Fort Clark (Brackettville), Kinney County, Texas. Collected 21 Mar 1893 by 

E. A. Mearns. Original number 2312.

Spermophilus tridecemlineatus alleni Merriam

Proc. Biol. Soc. Washington 12: 71, 24 Mar 1898b.

= Ictidomys tridecemlineatus alleni (Merriam, 1898). See Helgen 

et al. (2009:291).

Holotype: 56050. Skin and skull. Male. [SL: near summit] [FC: 

west slope] Bighorn Mountains, [SL and FC: altitude 8,000 

ft (2,439 m)], [Washakie County], Wyoming. Collected 18 

Sep 1893 by V. Bailey. Original number 4383.

Spermophilus tridecemlineatus pallidus J. A. Allen

Monogr. N. Amer. Rodentia 11: 872, Aug 1877.

= Ictidomys tridecemlineatus pallidus (J. A. Allen, 1877). See 

Helgen et al. (2009:293).

Lectotype: 16237. Skin only. Mouth of Yellowstone River, Montana. Collected 18 Aug 1857 by F. V. Hayden. No original 

number. Lectotype selected by A. H. Howell (1938:112).

Spermophilus tridecemlineatus texensis Merriam

Proc. Biol. Soc. Washington 12: 71, 24 Mar 1898b.

= Ictidomys tridecemlineatus texensis (Merriam, 1898). See Helgen et al. (2009:293).
1 4 • smithsonian contributions to zoology

Holotype: [186471]. Skin and skull. Adult male. Gainesville, Cooke 

County, Texas. Collected 15 Apr 1886 by G. H. Ragsdale. Original number [MCC: 25]. Merriam collection number 2117/2647.

Genus Iomys Thomas, 1908

Iomys lepidus Lyon

Proc. U.S. Natl. Mus. 40: 78, 25 Apr 1911a.

= Iomys horsfieldii thomsoni (Thomas, 1900). See Corbet and 

Hill (1992:315).

Holotype: 153684. Skin, skull, [and skeleton]. Adult male. Batu Jurong, [Kalimantan Barat], southwestern Borneo, [Indonesia]. 

Collected 27 Jun 1908 by W. L. Abbott. Original number 6005.

Genus Lariscus Thomas and Wroughton, 1909

Funambulus castaneus Miller

Proc. Washington Acad. Sci. 2: 217, 20 Aug 1900a.

= Lariscus insignis (F. Cuvier, 1821). Unassigned to subspecies by 

Thorington and Hoffmann (2005:783).

Holotype: 101696. Skin and skull. Immature male. Pulo Siantan, 

Anambas Islands, [Kepulauan Riau, Indonesia]. Collected 

10 Sep 1899 by W. L. Abbott. No original number.

Funambulus obscurus Miller

Smithsonian Misc. Coll. 45: 23, 6 Nov 1903a.

= Lariscus obscurus obscurus (Miller, 1903). See Corbet and Hill 

(1992:301).

Holotype: 121640. Skin and skull. Adult female. South Pagi Island (Pulau Pagai Selatan), [Sumatera Barat], [FC: west], 

Sumatra, [Indonesia]. Collected 22 Nov 1902 by W. L. Abbott. Original number 2086.

Funambulus peninsulae Miller

Smithsonian Misc. Coll. 45: 25, 6 Nov 1903a.

= Lariscus insignis peninsulae (Miller, 1903). See Thorington 

and Hoffmann (2005:783).

Holotype: 86776. Skin and skull. Adult male. Khow Sai Dow, 

[SL: altitude 1,000 ft (305 m)], Trong (Trang), Lower Siam 

(Thailand). Collected 18 Feb 1899 by W. L. Abbott. No 

original number.

Funambulus rostratus Miller

Smithsonian Misc. Coll. 45: 24, 6 Nov 1903a.

= Lariscus insignis rostratus (Miller, 1903). See Robinson and 

Kloss (1918a:234).

Holotype: 121801. Skin and skull. Adult female. Tana Bala 

(Pulau Tanahbala), Batu Islands, [Sumatera Utara, Sumatra, 

Indonesia]. Collected 12 Feb 1903 by W. L. Abbott. Original number 2281.

Genus Marmota Blumenbach, 1779

Arctomys dacota Merriam

N. Amer. Fauna 2: 8, 30 Oct 1889b.

= Marmota flaviventris dacota (Merriam, 1889). See A. H. 

Howell (1914:15).

Holotype: [186474]. Skin and skull. Adult male. Custer, [Custer 

County], Black Hills [South] Dakota. Collected 21 Jul 1888 

by V. Bailey. Original number [SL: 113]. Merriam collection 

number 4478/5113.

Arctomys olympus Merriam

Proc. Acad. Nat. Sci. Philadelphia 50: 352, 4 Oct 1898c.

= Marmota olympus (Merriam, 1898). See Trouessart (1904:344).

Holotype: 90518. [Skin and skull]. Adult male. [SL: near] timberline at [FC: near] head of Soleduck River, Olympic Mountains, [Clallam County], Washington. Collected 27 Aug 1897 

by C. H. Merriam and V. Bailey. Original number VB 6210.

Marmota caligata cascadensis A. H. Howell

Proc. Biol. Soc. Washington 27: 17, 2 Feb 1914.

Holotype: 90134. [Skin and skull]. Adult female. Mount Rainier, [SL and FC: Indian Henrys’ Hunting Grounds], altitude 

6,000 ft (1,829 m), [Pierce County], Washington. Collected 

11 Aug 1897 by W. K. Fisher. Original number 422.

Marmota caligata nivaria A. H. Howell

Proc. Biol. Soc. Washington 27: 17, 2 Feb 1914.

= Marmota caligata okanagana (King, 1836). See Thorington 

and Hoffmann (2005:800).

Holotype: 72235. [Skin and skull]. Adult female. Mountains near 

Upper Saint Mary’s Lake, altitude about 6,100 ft (1,859 m), 

[Glacier County], Montana. [Note: FC: Saint Mary’s Lake, 

Montana, 1,600 ft (488 m) above lake]. Collected 27 May 

1895 by A. H. Howell. Original number 23.

Marmota caligata sheldoni A. H. Howell

Proc. Biol. Soc. Washington, 27: 18, 2 Feb 1914.

= Marmota caligata caligata (Eschscholtz, 1829). See Thorington 

and Hoffmann (2005:800).

Holotype: 137319. [Skin and skull]. Adult male. Montague Island, [SL: Zaikof Bay], [FC: Prince William Sound], Alaska. 

Collected 24 May 1905 by C. Sheldon. Original number 12. 

5461-X.

Marmota flaviventer luteola A. H. Howell

Proc. Biol. Soc. Washington 27: 15, 2 Feb 1914.

= Marmota flaviventris luteola A. H. Howell, 1914. See A. H. 

Howell (1915b:50).

Holotype: 186520. [Skin and skull]. Adult male. Woods Post 

Office, Medicine Bow Mountains, altitude about 7,500 ft 

(2,286 m), [Albany County], Wyoming. Collected 13 Aug 

1888 by V. Bailey. Original number 148. Merriam collection 

number 4655/5306.

Marmota flaviventer nosophora A. H. Howell

Proc. Biol. Soc. Washington 27: 15, 2 Feb 1914.

= Marmota flaviventris nosophora A. H. Howell, 1914. See 

A. H. Howell (1915b:46).
number 640 • 1 5

Holotype: 168494. [Skin and skull]. Adult female. Willow Creek, 

altitude 4,000 ft (1,219 m), 7 mi (11 km) east of Corvallis, 

[Ravalli County], Montana. Collected 8 Apr 1910 by A. H. 

Howell. Original number 1723.

Marmota flaviventer obscura A. H. Howell

Proc. Biol. Soc. Washington 27: 16, 2 Feb 1914.

= Marmota flaviventris obscura A. H. Howell, 1914. See A. H. 

Howell (1915b:53).

Holotype: 133505. [Skin and skull]. Adult female. Wheeler Peak, 

5 mi (8 km) south of Twining, altitude 11,300 ft (3,442 m), 

[Taos County], New Mexico. Collected 24 Jul 1904 by 

V. Bailey. Original number 8181.

Marmota flaviventer parvula A. H. Howell

Proc. Biol. Soc. Washington 27: 14, 2 Feb 1914.

= Marmota flaviventris avara (Bangs, 1899). See Thorington and 

Hoffmann (2005:801).

Holotype: 93690. [Skin and skull]. Adult female. [SL: Jefferson], 

Toquima Range, about 10 mi (16 km) north of Belmont, 

Nye County, Nevada. Collected 3 Jun 1898 by V. Bailey. 

Original number 6495.

Marmota flaviventer warreni A. H. Howell

Proc. Biol. Soc. Washington 27: 16, 2 Feb 1914.

= Marmota flaviventris luteola A. H. Howell, 1914. See Warren 

(1936:392).

Holotype: 202937. [Skin and skull]. Adult female. Crested Butte, 

Colorado. Restricted by Warren (1936:393) to “Smith 

trail,” 2 mi (3 km) west of Crested Butte, Gunnison County, 

Colorado, about 10,000 ft (3,049 m). Collected 11 Jul 1902 

by E. R. Warren. Original number 35. 10639-X.

Marmota flaviventris sierra A. H. Howell

N. Amer. Fauna 37: 43, 7 Apr 1915b.

= Marmota flaviventris flaviventris (Audubon and Bachman, 

1841). See Thorington and Hoffmann (2005:801).

Holotype: 30984/42859. Skin and skull. Adult female. Head of 

Kern River, Mount Whitney, altitude 9,300 ft (2,835 m), 

[Tulare County], California. Collected 3 Sep 1891 by V. Bailey. Original number 3242.

Marmota monax pretensis A. H. Howell

N. Amer. Fauna 37: 33, 7 Apr 1915b.

= Marmota monax canadensis (Erxleben, 1777). See Thorington 

and Hoffmann (2005:802).

Holotype: 203532. [Skin and skull]. Adult male. Revelstoke, 

British Columbia, [Canada]. Collected 12 May 1890 by 

W. Spreadborough. Original number 170 [SL: 10712-X]. 

Two additional numbers, 227 and 240, are on skin labels.

Marmota monax preblorum A. H. Howell

Proc. Biol. Soc. Washington 27: 14, 2 Feb 1914.

= Marmota monax rufescens A. H. Howell, 1914. See Thorington and Hoffmann (2005:802).

Holotype: 78360. [Skin and skull]. Adult male. Wilmington, 

[Middlesex County], Massachusetts. Collected 19 Apr 1896 

by E. A. Preble. Original number 127.

Marmota monax rufescens A. H. Howell

Proc. Biol. Soc. Washington 27: 13, 2 Feb 1914.

Holotype: 186521. [Skin and skull]. Adult male. Elk River, 

[Sherburne County], Minnesota. Collected 9 Apr 1886 by 

V. Bailey. Original number 237. Merriam collection number 

2563/3173.

Marmota sibila Hollister

Smithsonian Misc. Coll. 56 (35): 1, 7 Feb 1912a. Secondary 

junior homonym of Arctomys sibila Wolf, 1808. Replaced 

with Marmota oxytona by Hollister (1914:251).

= Marmota caligata okanagana (King, 1836). See Thorington 

and Hoffmann (2005:800).

Holotype: 174503. Skin and skull. Adult female. Head of Moose 

Pass Branch of the Smoky River, altitude 7,200 ft (2,195 m), 

[Poole and Schantz (1942:541): near Moose Pass, British 

Columbia], Alberta, Canada. Collected 3 Aug 1911 by N. 

Hollister and C. D. Walcott, Jr. Original number [FC: NH] 

3871.

Genus Menetes Thomas, 1908

Lariscus berdmorei amotus Miller.

Smithsonian Misc. Coll. 61 (21): 24, 29 Dec 1913

= Menetes berdmorei berdmorei (Blyth, 1849). See Moore and 

Tate (1965:297).

Holotype: 124152. Skin and skull. Adult male. Domel Island, 

Mergui Archipelago, [Tanintharyi, Myanmar]. Collected 30 

Jan 1904 by W. L. Abbott. Original number 3081.

Genus Microsciurus J. A. Allen, 1895

Microsciurus alfari venustulus Goldman

Smithsonian Misc. Coll. 56 (36): 4, 19 Feb 1912a.

Holotype: 171030. Skin and skull. Adult female. Gatun, Canal 

Zone (Colon), Panama. Collected 1 Mar 1911 by E. A. Goldman. Original number [FC: Nelson and Goldman] 20995.

Microsciurus isthmius vivatus Goldman

Smithsonian Misc. Coll. 60: 4, 20 Sep 1912b.

= Microsciurus mimulus isthmius (Nelson, 1899). See Handley 

(1966:778).

Holotype: 179565. [Skin and skull]. Adult female. Near Cana, 

altitude 3,500 ft (1,067 m), in the Pirri range of mountains, 

[Darien], eastern Panama. Collected 5 Jun 1912 by E. A. 

Goldman. Original number 21771.

Sciurus (Microsciurus) isthmius Nelson

Bull. Amer. Mus. Nat. Hist. 12: 77, 14 Apr 1899a.

= Microsciurus mimulus isthmius (Nelson, 1899). See Handley 

(1966:778).
1 6 • smithsonian contributions to zoology

Holotype: 3399/37344. Skin and skull. Truando River, [Choco], 

Colombia. A footnote to the original description says 

“The Truando River is a tributary on the west side of the 

Rio Atrato at the extreme southeast end of the Isthmus of 

Darien.” Collected by A. Schott, date unknown. Original 

number [SL: 232]. [Received from Lt. Michler. Skin cataloged 1 Mar 1859; skull, 25 Mar 1898].

Genus Nannosciurus Trouessart, 1880

Nannosciurus bancanus Lyon

Proc. Biol. Soc. Washington 19: 55, 1 May 1906b.

= Nannosciurus melanotis bancanus Lyon, 1906. See Robinson 

and Kloss (1918a:249).

Holotype: 124880. Skin and skull. Adult female. Klabat Bay, Island of Banka (Bangka), [Kepulauan Bangka Belitung], east 

of Sumatra, [Indonesia]. Collected 24 Jun 1904 by W. L. 

Abbott. Original number 3430.

Nannosciurus borneanus Lyon

Proc. Biol. Soc. Washington 19: 54, 1 May 1906b.

= Nannosciurus melanotis borneanus Lyon, 1906. See Robinson 

and Kloss (1918a:249).

Holotype: 142271. Skin and skull. Adult female. Sanggau, [FC: 

Kapuas River], [Kalimantan Barat], western Borneo, [Indonesia]. Collected 23 Aug 1905 by W. L. Abbott. Original 

number 4368.

Nannosciurus pulcher Miller

Proc. Acad. Nat. Sci. Philadelphia 54: 153, 11 Jun 1902.

= Nannosciurus melanotis pulcher Miller, 1902. See Robinson 

and Kloss (1918a:249).

Holotype: 113131. Skin and skull. Adult female. Sinkep (Singkep) Island, [FC: Sakana Bay], South China Sea, [Lingga Archipelago, Kepulauan Riau, Sumatra, Indonesia]. Collected 

4 Sep 1901 by W. L. Abbott. Original number 1274.

Nannosciurus sumatranus Lyon

Proc. Biol. Soc. Washington 19: 53, 1 May 1906b.

= Nannosciurus melanotis pulcher Miller, 1902. See Thorington 

and Hoffmann (2005:784).

Holotype: 141058. Skin and skull. Adult male. [FC: Nyalo], 

Tarussan (Tarusan) Bay, [Sumatera Barat], Sumatra, [Indonesia]. Collected 16 Jan 1905 by W. L. Abbott. Original 

number 3946.

Genus Notocitellus A. H. Howell, 1938

Citellus adocetus Merriam

Proc. Biol. Soc. Washington 16: 79, 29 May 1903a.

= Notocitellus adocetus adocetus (Merriam, 1903). See Helgen 

et al. (2009:277)

Holotype: 126129. [Skin and skull]. Adult female. La Salada, 

40 mi (64 km) south of Uruapan, Michoacan, Mexico. Collected 17 Mar 1903 by E. W. Nelson and E. A. Goldman. 

Original number 16183.

Spermophilus annulatus goldmani Merriam

Proc. Biol. Soc. Washington 15: 69, 22 Mar 1902a.

= Notocitellus annulatus goldmani (Merriam, 1902). See Helgen 

et al. (2009:278).

Holotype: 91259. Skin and skull. Adult female. Santiago, [FC: 

altitude 200 ft (61 m)], Tepic (Nayarit), Mexico. Collected 

18 Jun 1897 by E. W. Nelson and E. A. Goldman. Original 

number 11223.

Genus Otospermophilus Brandt, 1844

Citellus beecheyi parvulus A. H. Howell

J. Mamm. 12: 160, 14 May 1931.

= Otospermophilus beecheyi parvulus (A. H. Howell, 1931). See 

Helgen et al. (2009:284).

Holotype: 28068/40167. Skin and skull. Subadult female. Shepherd Canyon, Argus Mountains, [Inyo County], California. 

Collected 30 Apr 1891 by A. K. Fisher. Original number 

569.

Citellus beecheyi sierra A. H. Howell

N. Amer. Fauna 56: 153, 18 May 1938.

= Otospermophilus beecheyi sierra (A. H. Howell, 1938). See 

Helgen et al. (2009:284).

Holotype: 88421. Skin and skull. Adult female. Emerald Bay, 

Lake Tahoe, [Eldorado County], California. Collected 23 

May 1897 by J. A. Loring. Original number 4166.

Citellus grammurus utah Merriam

Proc. Biol. Soc. Washington 16: 77, 29 May 1903b.

= Otospermophilus variegatus utah (Merriam, 1903). See Helgen et al. (2009:284).

Holotype: [186468]. Skin and skull. Adult female. Foot of 

Wasatch Mountains, near Ogden, [Weber County], Utah. 

Collected 10 Oct 1888 by V. Bailey. Original number 291. 

Merriam collection number 4767/5427.

Citellus variegatus juglans V. Bailey

Proc. Biol. Soc. Washington 26: 131, 21 May 1913a.

= Otospermophilus variegatus grammurus (Say, 1823). See Helgen et al. (2009:284).

Holotype: 148289. [Skin and skull]. Adult male. Glenwood, 

altitude 5,000 ft (1,524 m), [FC: mouth of White Water 

Creek] on the Rio San Francisco at the southwest base of 

the Mogollon Mountains, [Catron County], New Mexico. 

Collected 2 Nov 1906 by V. Bailey. Original number 8587.

Spermophilus beecheyi fisheri Merriam

Proc. Biol. Soc. Washington 8: 133, 28 Dec 1893a.

= Otospermophilus beecheyi fisheri (Merriam, 1893). See Helgen et al. (2009:284).
number 640 • 1 7

Holotype: 29318/41385. Skin and skull. Adult male. Kern Valley, [FC: Kern River (South Branch)], 25 mi (40 km) above 

Kernville, Kern County, California. Restricted to South Fork 

of Kern River 3 mi (5 km) above Onyx, Kern County, California by A. B. Howell (1923:266). Collected 6 Jul 1891 by 

A. K. Fisher. Original number 741.

Spermophilus couchii Baird

Proc. Acad. Nat. Sci. Philadelphia 7: 332, reported favorably for 

publication 24 Apr 1855b.

= Otospermophilus variegatus couchii (Baird, 1855). See Helgen 

et al. (2009:284).

Lectotype: [338/1255]. Skin and skull. Santa Catarina, [Baird 

(1857:713): a few miles west of Monterrey], [Baird 

(1857:312): Nuevo Leon], Mexico. Collected Apr 1853 by 

D. N. Couch. Original number [Baird (1857:312): 190].

No type designated in the original description. Measurements 

given in the original description match those of this specimen 

as given by Baird (1857:312) and was therefore considered the 

type by Lyon and Osgood (1909:165). However, since Baird 

had two specimens before him when he described the taxon, 

the specimen selected by Lyon and Osgood is the Lectotype.

Genus Paraxerus Forsyth Major, 1893

Paraxerus kahari Heller

Smithsonian Misc. Coll. 56 (17): 2, 28 Feb 1911.

= Paraxerus ochraceus kahari Heller, 1911. See Hollister (1919:20).

Holotype: 164203. [Skin and skull]. Adult female. Meru [SL and 

FC: Mweru] Boma, northeast of Mount Kenia (Kenya), British East Africa (Kenya). Collected 24 Sep 1909 by E. Heller. 

Original number 1174.

Genus Petaurista Link, 1795

Petaurista batuana Miller

Smithsonian Misc. Coll. 45: 27, 6 Nov 1903a.

= Petaurista petaurista batuana Miller, 1903. See Robinson and 

Kloss (1918a:173).

Holotype: 121742. Skin and skull. Adult male. Tana Bala (Pulau 

Tanahbala), Batu Islands, [Sumatera Utara], Sumatra, [Indonesia]. Collected 5 Feb 1903 by W. L. Abbott. Original 

number 2233.

Petaurista mimicus Miller

Smithsonian Misc. Coll. 61(21): 27, 29 Dec 1913.

= Petaurista petaurista marchio Thomas, 1908. See Corbet and 

Hill (1992:309).

Holotype: 143341. Skin and skull. Adult female. Pulo Rupat, 

[Riau], eastern Sumatra, [Indonesia]. Collected 10 Mar 

1906 by W. L. Abbott. Original number 4695.

Petaurista petaurista nigrescens Medway

Malaysian Br. Roy. Asiat. Soc. 36: 116, Dec 1965.

Holotype: 19177/34899. Skin and skull. Male. Near Sandakan 

[Sabah, Borneo, Malaysia]. Collected 21 Oct 1887 by C. F. 

Adams. Original number [SL: AXX-100].

Petaurista rubicundus A. B. Howell

J. Washington Acad. Sci. 17: 82, 19 Feb 1927.

= Petaurista philippensis yunanensis (Anderson, 1875). See Corbet and Hill (1992:311).

Holotype: 240857. Skin only. Adult female. Mapientung [SL: 

Mapienting near Mt. Omei], Szechwan (Sichuan), China. 

Collected fall [FC: 24 Jul] 1924 by [FC: native hunters and 

secured by] D. C. Graham. No original number.

Petaurista terutaus Lyon

Proc. Biol. Soc. Washington 20: 17, 23 Feb 1907d.

= Petaurista petaurista terutaus Lyon, 1907. See Robinson and 

Kloss (1918b:224).

Holotype: 123934. Skin and skull. Adult male. Pulo Terutau (Ko 

Taruto), [Satun, Thailand]. Collected 9 Apr 1904 by W. L. 

Abbott. Original number 3219.

Genus Petinomys Thomas, 1908

Sciuropterus crinitus Hollister

Proc. Biol. Soc. Washington 24: 185, 23 Jun 1911b.

= Petinomys crinitus (Hollister, 1911). See Corbet and Hill 

(1992:318).

Holotype: 239217. Skin and skull. Adult male. Basilan Island, 

[Basilan Province], Philippines. Collected 12 Jan 1907 by 

R. C. McGregor and A. Celestino. No original number. Collection of the Philippine Bureau of Sciences number 101.

Sciuropterus maerens Miller

Smithsonian Misc. Coll. 45: 26, 6 Nov 1903a.

= Petinomys lugens (Thomas, 1895). See Corbet and Hill (1992: 

317).

Holotype: 121531. Skin and skull. Adult female. North Pagi Island (Pulau Pagai Utara), [Sumatera Barat], Sumatra, [Indonesia].Collected 14 Jan 1902 [FC: 1903, which is correct] 

by W. L. Abbott. Original number 2206.

Genus Prosciurillus Ellerman, 1947

Sciurus evidens Miller and Hollister

Proc. Biol. Soc. Washington 34: 99, 30 Jun 1921.

= Prosciurillus murinus murinus (Müller and Schlegel, 1844). 

See Corbet and Hill (1992:304).

Holotype: 217814. Skin and skull. Adult female. Pulo Lembeh, 

[Sulawesi Utara], off the shore of northeastern Celebes, [Indonesia]. Collected 16 [SL and FC: Jun, not Jan, as in original 

description] 1916 by H. C. Raven. Original number 2525.

Sciurus murinus necopinus Miller and Hollister

Proc. Biol. Soc. Washington 34: 98, 30 Jun 1921.
1 8 • smithsonian contributions to zoology

= Prosciurillus murinus necopinus (Miller and Hollister, 1921). 

See Corbet and Hill (1992:304).

Holotype: 218712. Skin and skull. Adult female. Goenoeng 

Lehio (southwest from Lake Lindoe), [Sulawesi Tengah], 

Middle Celebes, [Indonesia]. Collected 20 Jan 1917 by 

H. C. Raven. Original number 3107.

Genus Protoxerus Forsyth Major, 1893

Protoxerus stangeri bea Heller

Smithsonian Misc. Coll. 59(16): 2, 5 Jul 1912.

Holotype: 181786. [Skin and skull]. Adult male. Lukosa River, 

Kakumega (Kakamega) Forest, [Western Province], British East Africa (Kenya). Collected 8 Feb 1912 by E. Heller. 

Original number 5614.

Genus Pteromys G. Cuvier, 1800

Pteromys volans incanus Miller

Proc. Biol. Soc. Washington 31: 3, 21 Feb 1918.

= Pteromys volans volans (Linnaeus, 1758). See Corbet 

(1978:86).

Holotype: 200613. Skin and skull. Adult female. [FC: sex was 

marked incorrectly and should be male]. Verkhne Kolymsk, 

[Sakha (Yakutiya)], East Siberia (Russia). Collected 24 Apr 

1915 by C. Amory, Jr. Original number 374.

Genus Ratufa Gray, 1867

Ratufa anambae Miller

Proc. Washington Acad. Sci. 2: 215, 20 Aug 1900a.

= Ratufa bicolor melanopepla Miller, 1900. See Thorington and 

Hoffmann (2005:756).

Holotype: 101725. Skin and skull. Adult female, not male as 

in original description. Pulo Jimaja (Djemadja Island), 

Anamba Islands, [Kepulauan Riau, Indonesia]. Collected 25 

Sep 1899 by W. L. Abbott. No original number.

Ratufa angusticeps Miller

Proc. Washington Acad. Sci. 3: 130, 26 Mar 1901a.

= Ratufa bicolor melanopepla Miller, 1900. See Thorington and 

Hoffmann (2005:756).

Holotype: 104646. Skin and skull. Adult male. Pulo Lingung, off 

south coast of Bunguran [SL: Natuna Islands], [Kepulauan 

Riau, Indonesia]. Collected 17 Jun 1900 by W. L. Abbott. 

Original number 481.

Ratufa arusinus Lyon

Proc. U.S. Natl. Mus. 32: 442, 23 May 1907e.

= Ratufa affinis hypoleucos (Horsfield, 1823). See Thorington 

and Hoffmann (2005:756).

Holotype: 143351. Skin and skull. Adult male. Aru Bay, [Sumatera Utara], northeast coast of Sumatra, [Indonesia]. Collected 16 Jan 1906 by W. L. Abbott. Original number 4635.

Ratufa balae Miller

Smithsonian Misc. Coll. 45: 6, 6 Nov 1903a.

= Ratufa affinis hypoleucos (Horsfield, 1823). See Thorington 

and Hoffmann (2005:756).

Holotype: 121715. Skin and skull. Adult male. Tana Bala (Pulau 

Tanahbala), Batu Islands, [Sumatera Utara], Sumatra, [Indonesia]. Collected 5 Feb 1903 by W. L. Abbott. Original 

number 2224.

Ratufa bicolor major Miller

Proc. Biol. Soc. Washington 24: 28, 24 Feb 1911.

= Ratufa bicolor bicolor (Sparrman, 1778). See Thorington and 

Hoffmann (2005:756).

Holotype: 155666. Skin, skull, [and skeleton]. Adult female. 

Tjibodas, Mount Gede, altitude 4,500 ft (1,372 m), [Jawa 

Barat], Java, [Indonesia]. Collected 15 Aug 1909 by 

W. Palmer on the Bryant Expedition. Original number 505.

Ratufa bulana Lyon

Proc. U.S. Natl. Mus. 36: 482, 1 Jun 1909b.

= Ratufa affinis insignis Miller, 1903. See Thorington and Hoffmann (2005:756).

Holotype: 144412. Skin and skull. Adult female. Pulo Bulan, 

Rhio-Linga (Riau) Archipelago, [Kepulauan Riau, Sumatra, 

Indonesia]. Collected 23 Mar 1907 by W. L. Abbott. Original number 5130.

Ratufa carimonensis Miller

Proc. U.S. Natl. Mus. 31: 257, 11 Sep 1906b.

= Ratufa affinis insignis Miller, 1903. See Thorington and Hoffmann (2005:756).

Holotype: 122813. Skin and skull. Adult female. Great Karimon 

Island, Rhio-Linga (Riau) Archipelago [FC: Straits of Malacca], [Kepulauan Riau, Sumatra, Indonesia]. Collected 2 

Jun 1903 by W. L. Abbott. Original number 2465.

Ratufa catemana Lyon

Proc. U.S. Natl. Mus. 32: 443, 23 May 1907e.

= Ratufa affinis hypoleucos (Horsfield, 1823). See Thorington 

and Hoffmann (2005:756).

Holotype: 123124. Skin and skull. Adult male. Kateman River, 

[Riau], southeastern Sumatra, [Indonesia]. Collected 27 

Aug 1903 by W. L. Abbott. Original number 2759.

Ratufa celaenopepla Miller

Smithsonian Misc. Coll. 61 (21): 26, 29 Dec 1913.

= Ratufa bicolor phaeopepla Miller, 1913. See Moore and Tate 

(1965:52).

Holotype: 124149. Skin and Skull. Adult male. Domel Island, 

Mergui Archipelago, [Tanintharyi, Myanmar]. Collected 26 

Jan 1904 by W. L. Abbott. Original number 3073.

Ratufa condurensis Miller

Proc. U.S. Natl. Mus. 31: 258, 11 Sep 1906b.
number 640 • 1 9

= Ratufa affinis insignis Miller, 1903. See Thorington and Hoffmann (2005:756).

Holotype: 122879. Skin and skull. Adult male. Pulo Kundur, 

Rhio-Linga (Riau) Archipelago, [Kepulauan Riau, Sumatra, 

Indonesia]. Collected 25 Jun 1903 by W. L. Abbott. Original number 2552.

Ratufa confinis Miller

Proc. U.S. Natl. Mus. 31: 259, 11 Sep 1906b.

= Ratufa affinis bunguranensis (Thomas and Hartert, 1894). See 

Thorington and Hoffmann (2005:756).

Holotype: 113134. Skin and skull. Adult female. [FC: Sakana 

Bay], Sinkep (Singkep) Island, Rhio-Linga (Lingga) Archipelago, [Kepulauan Riau, Sumatra, Indonesia]. Collected 3 

Sep 1901 by W. L. Abbott. Original number 1265.

Ratufa conspicua Miller

Smithsonian Misc. Coll. 45: 5, 6 Nov 1903a.

= Ratufa affinis insignis Miller, 1903. See Thorington and Hoffmann (2005:756).

Holotype: 115528. Skin and skull. Adult male. Pulo Bintang, 

Rhio (Riau) Archipelago, [Kepulauan Riau, Sumatra, Indonesia]. Collected 19 Aug 1902 by W. L. Abbott. Original 

number 1900.

Ratufa cothurnata Lyon

Proc. U.S. Natl. Mus. 40: 93, 25 Apr 1911a.

= Ratufa affinis cothurnata Lyon, 1911. See Chasen and Kloss 

(1931:22).

Holotype: 145378. Skin and skull. Adult female. Foot of Mount 

Palung, near Sukadana, [Kalimantan Barat], western Borneo, [Indonesia]. Collected 2 Sep, not 9 Feb as in original 

description, 1907 by W. L. Abbott. Original number 5537.

Ratufa femoralis Miller

Proc. U.S. Natl. Mus. 26: 447, 3 Feb 1903b.

= Ratufa affinis hypoleucos (Horsfield, 1823). See Thorington 

and Hoffmann (2005:756)

Holotype: 114361. Skin and skull. Adult female. Pulo Tuangku, 

Banjak Islands, [Aceh, Sumatra, Indonesia]. Collected 27 

Jan 1902 by W. L. Abbott. Original number 1479.

Ratufa griseicollis Lyon

Proc. U.S. Natl. Mus. 40: 94, 25 Apr 1911a.

= Ratufa affinis cothurnata Lyon, 1911. See Thorington and 

Hoffmann (2005:756).

Holotype: 145372. Skin and skull. Adult male. Panebangan Island, [Kalimantan Barat], west coast of Borneo, [Indonesia]. 

Collected 24 May 1907 by W. L. Abbott. Original number 

5315.

Ratufa insignis Miller

Smithsonian Misc. Coll. 45: 4, 6 Nov 1903a.

= Ratufa affinis insignis Miller, 1903. See Dammerman (1926:317).

Holotype: 115531. Skin and skull. Adult male. Pulo Sugi, Rhio (Riau) 

Archipelago, [Kepulauan Riau, Sumatra, Indonesia]. Collected 

26 Aug 1902 by W. L. Abbott. Original number 1960.

Ratufa laenata Miller

Proc. U.S. Natl. Mus. 26: 449, 3 Feb 1903b.

= Ratufa bicolor palliata Miller, 1902. See Thorington and Hoffmann (2005:756).

Holotype: 114350. Skin and skull. Adult male. Pulo Tuangku, 

Banjak Islands, [Aceh, Sumatra, Indonesia]. Collected 27 

Jan 1902 by W. L. Abbott. Original number 1478.

Ratufa masae Miller

Smithsonian Misc. Coll. 45: 7, 6 Nov 1903a.

= Ratufa affinis hypoleucos (Horsfield, 1823). See Thorington 

and Hoffmann (2005:756).

Holotype: 121818. Skin and skull. Adult male. Tana Masa 

(Pulau Tanahmasa), Batu Islands, [Sumatera Utara, Sumatra, Indonesia]. Collected 21 Feb 1903 by W. L. Abbott. 

Original number 2330.

Ratufa melanopepla Miller

Proc. Washington Acad. Sci. 2: 71, 25 Jul 1900d.

= Ratufa bicolor melanopepla Miller, 1900. See Thorington and 

Hoffmann (2005:756).

Holotype: 83230. Skin and skull. Adult male. [SL: Telibon Island], Trong (Trang), Lower Siam (Thailand). Collected 27 

Feb 1896 By W. L. Abbott. No original number.

Ratufa melanopepla peninsulae Miller

Smithsonian Misc. Coll. 61 (21): 25, 29 Dec 1913.

= Ratufa bicolor melanopepla Miller, 1900. See Thorington and 

Hoffmann (2005:756).

Holotype: 83478/[49466]. Skin and skeleton. Adult male. Lay 

Song Hong, Trong (Trang), Lower Siam (Thailand). Collected 23 Sep 1896 by W. L. Abbott. No original number.

Ratufa nigrescens Miller

Proc. U.S. Natl. Mus. 26: 448, 3 Feb 1903b.

= Ratufa affinis hypoleucos (Horsfield, 1823). See Thorington 

and Hoffmann (2005:756).

Holotype: 114556. Skin and skull. Adult female. Pulo Mansalar 

(Pulau Musala), off Tapanuli Bay, [Sumatera Utara], [FC: 

west coast] Sumatra, [Indonesia]. Collected 11 Mar 1902 by 

W. L. Abbott. Original number 1641.

Ratufa notabilis Miller

Proc. Acad. Nat. Sci. Philadelphia 54: 150, 11 Jun 1902.

= Ratufa affinis bunguranensis (Thomas and Hartert, 1894). See 

Thorington and Hoffmann (2005:756).

Holotype: 113064. Skin and skull. Adult male. [FC: Mentuda 

Bay], west coast of Linga (Lingga) Island, [Lingga Archipelago, Kepulauan Riau, Sumatra, Indonesia]. Collected 24 

Aug 1901 by W. L. Abbott. Original number 1210.
2 0 • smithsonian contributions to zoology

Ratufa palliata Miller

Proc. Acad. Nat. Sci. Philadelphia 54: 147, 11 Jun 1902.

= Ratufa bicolor palliata Miller, 1902. See Robinson and Kloss 

(1918a:188).

Holotype: 113162. Skin and skull. Adult male. Indragiri River, 

[Riau], eastern Sumatra, [Indonesia]. Collected 23 Sep 1901 

by W. L. Abbott. Original number 1327.

Ratufa palliata batuana Lyon

Proc. U.S. Natl. Mus. 52: 445, 30 Dec 1916.

= Ratufa bicolor palliata Miller, 1902. See Thorington and Hoffmann (2005:756).

Holotype: 121707. Skin and skull. Adult male. Tana Bala (Pulau 

Tanahbala), off the Batu Islands, [Sumatera Utara, Sumatra, 

Indonesia]. Collected 11 Feb 1902 by W. L. Abbott. Original number 2263.

Ratufa phaeopepla Miller

Smithsonian Misc. Coll. 61 (21): 25, 29 Dec 1913.

= Ratufa bicolor phaeopepla Miller, 1913. See Chasen and Kloss 

(1930:69).

Holotype: 124235. Skin and skull. Adult male. Sungei Balik, 

Tenasserim (Tanintharyi), [Myanmar]. Collected 25 Feb 

1904 by W. L. Abbott. Original number 3141.

Ratufa piniensis Miller

Smithsonian Misc. Coll. 45: 8, 6 Nov 1903a.

= Ratufa affinis hypoleucos (Horsfield, 1823). See Thorington 

and Hoffmann (2005:756).

Holotype: 121840. Skin and skull. Adult male. Pulo Pinie (Pulau 

Pini), Batu Islands, [Sumatera Utara, Sumatra, Indonesia]. 

Collected 1 Mar 1903 by W. L. Abbott. Original number 

2343.

Ratufa polia Lyon

Proc. U.S. Natl. Mus. 31: 585, 18 Dec 1906a.

= Ratufa affinis polia Lyon, 1906. See Thorington and Hoffmann (2005:756).

Holotype: 125004. Skin and skull. Adult female. Bukit Menguru, 

Billiton Island, between Sumatra and Borneo, [Kepulauan 

Bangka Belitung, Sumatra, Indonesia]. Collected 9 Aug 

1904 by W. L. Abbott. Original number 3551.

Ratufa polia bancana Lyon

Proc. U.S. Natl. Mus. 31: 587, 18 Dec 1906a.

= Ratufa affinis bancana Lyon, 1906. See Thorington and Hoffmann (2005:756).

Holotype: 124680. Skin and skull. Adult male. Tanjong 

Rengsam, island of Banka (Bangka), [Kepulauan Bangka 

Belitung], east of Sumatra, [Indonesia]. Collected 27 May 

1904 by W. L. Abbott. Original number 3277.

Ratufa pyrsonota Miller

Proc. Washington Acad. Sci. 2: 75, 25 Jul 1900d.

= Ratufa affinis affinis (Raffles, 1821). See Thorington and Hoffmann (2005:756).

Holotype: 83483. Skin and skull. Adult male. [Poole and Schantz 

(1942:550): Lay Song Hong], Trong (Trang), Lower Siam 

(Thailand). Collected 29 Sep 1896 by W. L. Abbott. No 

original number.

Ratufa tiomanensis Miller

Proc. Washington Acad. Sci. 2: 216, 20 Aug 1900a.

= Ratufa bicolor melanopepla Miller, 1900. See Thorington and 

Hoffmann (2005:756).

Holotype: 101751. Skin and skull. Adult male. Tioman Island, 

[Pahang, Malaysia]. Collected 4 Oct 1899 by W. L. Abbott. 

No original number.

Ratufa vittata Lyon

Proc. U.S. Natl. Mus. 40: 94, 25 Apr 1911a.

= Ratufa affinis ephippium (Müller, 1838). See Thorington and 

Hoffmann (2005:756).

Holotype: 151758. Skin and skull. Adult female. Saratokon [FC: 

Saratok], Pulo Laut, [Kalimantan Selatan], off southeast 

coast of Borneo, [Indonesia]. Collected 19 Dec 1907 by 

W. L. Abbott. Original number 5632.

Ratufa vittatula Lyon

Proc. U.S. Natl. Mus. 40: 95, 25 Apr 1911a.

= Ratufa affinis ephippium (Müller, 1838). See Thorington and 

Hoffmann (2005:756).

Holotype: 151762. Skin and skull. Adult male. Pulo Sebuku, [Kalimantan Selatan], southeastern Borneo, [Indonesia]. Collected 2 Jan 1908 by W. L. Abbott. Original number 5720.

Genus Sciurus Linnaeus, 1758

Sciurus aberti chuscensis Goldman

Proc. Biol. Soc. Washington 44: 133, 17 Oct 1931a.

Holotype: 158553. [Skin and skull]. Adult female. Chusca 

Mountains, altitude 9,000 ft (2,743 m), [McKinley County], 

northwestern New Mexico. Collected 5 Oct 1908 by V. Bailey. Original number 8997.

Sciurus aberti concolor True

Proc. U.S. Natl. Mus. 17: 1, 24 Apr 1894a. Preprint of Proc. 

U.S. Natl. Mus. 17: 241, 15 Nov 1894b. Primary junior 

homonym of Sciurus concolor Blyth, 1855. Replaced with 

Sciurus aberti ferreus True (1900:183).

Holotype: 21423/36281. Skin and skull. Female. Loveland, Larimer County, Colorado. Collected by W. G. Smith. No original number or date collected [MC: cataloged 30 Oct 1893].

Sciurus aberti mimus Merriam

Proc. Biol. Soc. Washington 17: 130, 9 Jun 1904a.

= Sciurus aberti ferreus True, 1900. See Hoffmeister and Diersing 

(1978:412).
number 640 • 2 1

Holotype: 70908. [Skin and skull]. Adult female. Hall Peak, at 

south end of Cimarron Mountains, [Mora County], northeastern New Mexico. Collected 16 Jan 1895 by C. M. Barber. Original number 61.

Sciurus albipes colimensis Nelson

Proc. Biol. Soc. Washington 12: 152, 3 Jun 1898a.

= Sciurus aureogaster nigrescens Bennett, 1833. See Musser 

(1970:15).

Holotype: 33197/45202. Skin and skull. Adult male. Hacienda 

Magdalena, Colima, Mexico. Collected 19 Mar 1892 by 

E. W. Nelson. Original number 2239.

Sciurus albipes effugius Nelson

Proc. Biol. Soc. Washington 12: 152, 3 Jun 1898a.

= Sciurus aureogaster nigrescens Bennett, 1833. See Musser 

(1970:15).

Holotype: 70288. [Skin and skull]. Adult female. Mountains 

near Chilpancingo, [FC: altitude 9,800 ft (2,987 m)], Guerrero, Mexico. Collected 24 Dec 1894 by E. W. Nelson and 

E. A. Goldman. Original number 7271.

Sciurus albipes nemoralis Nelson

Proc. Biol. Soc. Washington 12: 151, 3 Jun 1898a.

= Sciurus aureogaster nigrescens Bennett, 1833. See Musser 

(1970:15).

Holotype: 35358/47623. Skin and skull. Adult male. Patzcuaro, [FC: altitude about 7,000 ft (2,137 m)], Michoacan, 

Mexico. Collected 23 Jul 1892 by E. W. Nelson. Original 

number 2905.

Sciurus albipes quercinus Nelson

Proc. Biol. Soc. Washington 12: 150, 3 Jun 1898a. Primary junior 

homonym of Sciurus quercinus Erxleben, 1777. Replaced 

with Sciurus albipes hernandezi by Nelson (1898b:783).

= Sciurus aureogaster nigrescens Bennett, 1833. See Musser 

(1970:15).

Holotype: 68202. [Skin and skull]. Adult female. Mountains 

on west side of valley of Oaxaca, [SL and FC: 15 mi (24 

km)W Oaxaca], [FC: altitude 9,500 ft (2,896 m)], Oaxaca, 

Mexico. Collected 15 Sep 1894 by E. W. Nelson and E. A. 

Goldman. Original number 6768.

Sciurus alleni Nelson

Proc. Biol. Soc. Washington 12: 147, 3 Jun 1898a.

Holotype: 25731/33131. Skin and skull. Adult male. Monterrey, Nuevo Leon, Mexico. Collected 22 Feb 1891 by C. P. 

Streator. Original number 563.

Sciurus arizonensis Coues

Amer. Nat. 1: 357, Sep 1867.

Holotype: [8475/37607]. Skin and skull. Female. Fort Whipple, 

[Yavapai County], Arizona. Collected 20 Dec 1865 by 

E. Coues. No original number.

There are several discrepancies in the data for this specimen and 

the original specimen label is missing. Coues (1867:357) did 

not designate a type by number but indicated that he took 

only one specimen. J. A. Allen (1877:741) lists 8475 as the 

type of S. arizonensis. Both give the date of collection as 

20 Dec 1865, but both the USNM skin and bone catalogs 

give the date as 6 Dec 1864. Coues (1867:357) stated the 

specimen was female but no sex is entered in either catalog. Coues (1867:357) stated he shot the specimen but an 

entry in the skin catalog under “Remarks” that is dittoed 

for several specimens from Fort Whipple states “Taken from 

Indians in a fight.” An unattributed comment in “Remarks” 

of the bone catalog, attributed to M. W. Lyon by Poole and 

Schantz (1942:553) states “Skull taken from mounted type, 

not naturally attached to skin, and it is very doubtful if it is 

the type skull. 31 Oct 98.” The skin was cataloged 12 Apr 

1866 and the skull 1 Nov 1898.

Sciurus aureogaster frumentor Nelson

Proc. Biol. Soc. Washington 12: 154, 3 Jun 1898a.

= Sciurus aureogaster nigrescens Bennett, 1833. See Musser 

(1970:15).

Holotype: 54259. [Skin and skull]. Adult male. Las Vigas, [FC: 

altitude 7,500 ft (2,286 m)], Veracruz, Mexico. Collected 18 

Jun 1893 by E. W. Nelson. Original number 5073.

Sciurus boothiae belti Nelson

Proc. Washington Acad. Sci. 1: 78, 9 May 1899b.

= Sciurus variegatoides belti Nelson, 1899. See Harris (1937:13).

Holotype: 36477/48847. Skin and skull. Adult female. Escondido 

River, 50 mi (80 km) from Bluefields, [Atlantico Sur], Nicaragua. Collected 12 Oct, not 22 Nov as in original description, 

1892 by C. W. Richmond. Original number [SL and FC: 124].

Sciurus boothiae managuensis Nelson

Proc. Biol. Soc. Washington 12: 150, 3 Jun 1898a.

= Sciurus variegatoides managuensis Nelson, 1898. See Harris 

(1937:17).

Holotype: 62476. [Skin and skull]. Adult male. Managua River, 

[Izabal], Guatemala. Collected 12 Feb 1895 by C. McElroy. 

Original number [SL: 4].

Sciurus boothiae underwoodi Goldman

J. Washington Acad. Sci. 22: 275, 19 May 1932a.

= Sciurus variegatoides underwoodi Goldman, 1932. See Harris 

(1937:9).

Holotype: 250219. [Skin and skull]. Adult male. Monte Redondo, altitude 5,100 ft (1,554 m), about 30 mi (48 km) 

northwest of Tegucigalpa, [Francisco Morazan], Honduras. 

Collected 8 Dec 1931 by C. F. Underwood. Original number 

644 [SL: 26780-X].

Sciurus carolinensis hypophaeus Merriam

Science 7: 351, 16 Apr 1886b.
2 2 • smithsonian contributions to zoology

Lectotype: [193864. Skin and skull]. [SL: Adult female. Elk 

River], Sherbourne (Sherburne) County, Minnesota. [SL: 

Collected 20 Jan 1886 by V. Bailey. Original number 29]. 

Merriam Collection number 1949/2483. Lectotype selected 

by V. Schantz in 1938 but apparently not published until 

Poole and Schantz (1942:554).

Sciurus carolinensis minutus H. H. Bailey

Bull. Bailey Mus. Lib. Nat. Hist. 12: 4, 5 Jan 1937a. Primary 

junior homonym of Sciurus minutus duChaillu, 1861. Replaced with Sciurus carolinensis matecumbei by H. H. Bailey (1937b:516).

= Sciurus carolinensis extimus Bangs, 1896. See Hubbard and 

Banks (1970:329).

Holotype: 347444. Skin and skull. Adult female. Lower Key 

Largo, Monroe County, Florida. Collected 13 June 1923 

by H. H. Bailey. No original number. Bailey Collection of 

Natural History number 178.

Sciurus carolinensis yucatanensis J. A. Allen

Monogr. N. Amer. Rodentia. 11: 705, Aug 1877.

= Sciurus yucatanensis yucatanensis J. A. Allen, 1877. See Miller 

(1912:330).

Syntypes: 8503/37496. Skin and skull. Female. Merida, Yucatan, 

[Mexico]. Collected [SL: 3] Mar 1865 by A. Schott, [SL: 

Comision cientifica de Yucatan, Jose Salagar Llarregui.] 

Original number 229.

8504/37493. Skin and skull. No sex. Locality as above. Collected Mar 1865 by A. Schott. Original number 332. Considered a Paratype by previous authors, Lyon and Osgood 

(1909:197), Poole and Schantz (1942:554), and Helgen and 

McFadden (2001:106); J. A. Allen (1877:716) listed this 

specimen by number along with numbers 8502 and 8505, 

and all three are Syntypes. Number 8502 is in the collection 

of the Museum of Comparative Zoology, Harvard (MCZ 

5398) (Helgen and McFadden, 2001:106). Number 8505 

cannot be found.

Sciurus castanonotus Baird

Proc. Acad. Nat. Sci. Philadelphia 7: 332, reported favorably for 

publication 24 Apr 1855b, receipt acknowledged 15 Jun 

1855 by the American Philosophical Society. Original spelling of castanotus corrected by Baird (1857:266).

= Sciurus aberti aberti Woodhouse, 1853. See J. A. Allen 

(1877:735).

Lectotype: 121/1107. [Skin and skull]. [Lyon and Osgood 

(1909:183): Adult female]. On the Mimbres [SL: Coppermines, Gila]. In designating the Lectotype, Lyon and 

Osgood (1909:182) give the type locality as Coppermines, 

near the present site of Georgetown, Grant County, New 

Mexico. Baird (1857:707) in his list of localities says of 

Coppermines, “A former station of the United StatesMexican Boundary Survey, subsequently called Fort Webster. Not indicated clearly whether on a tributary of the Gila 

or Mimbres. About Latitude 33°, Longitude 108°.” Baird 

(1857:709) says of Fort Webster “See also Coppermines. 

Altitude 6,350 ft (1,935 m). Latitude 32°47’, Longitude 

108°04’.” J. P. Hubbard, who has researched old New Mexico collecting localities for decades, indicates the proper locality designation should read “On the Mimbres [River=the 

copper mines=former Santa Rita del Cobre in present 

Grant County, New Mexico (approximately 32°47.560′N, 

108°4.710′W)]” J. P. Hubbard (in litt., 26 October 2010 

to A. L. Gardner). Collected [SL: 1851] by J. H. Clark. No 

original number.

Sciurus chiricahuae Goldman

Proc. Biol. Soc. Washington 46: 71, 27 Apr 1933a.

= Sciurus nayaritensis chiricahuae Goldman, 1933. See Lee and 

Hoffmeister (1963:189).

Holotype: 244124. Skin and skull. Adult female. Cave Creek, 

altitude 5,200 ft (1,585 m), Chiricahua Mountains, Cochise 

County, Arizona. Collected 16 Aug 1923 by E. A. Goldman. 

Original number 23527.

Sciurus colliaei nuchalis Nelson

Proc. Washington Acad. Sci. 1: 59, 9 May 1899b.

Holotype: 32657/44580. Skin and skull. Adult male. Manzanillo, [FC: altitude 50 ft (15 m)], Colima, Mexico. Collected 

2 Feb 1892 by E. W. Nelson. Original number [SL: 1828].

Sciurus deppei vivax Nelson

Proc. Biol. Soc. Washington 14: 131, 9 Aug 1901.

Holotype: 107932. Skin and skull. Adult female. Apazote, [SL 

and FC: near Yohaltun], Campeche, Mexico. Collected 8 

Jan 1901 by E. W. Nelson and E. A. Goldman. Original 

number [SL: 14429].

Sciurus dorsalis Woodhouse

Proc. Acad. Nat. Sci. Philadelphia 6: 110, reported favorably 

for publication 29 Jun 1852, receipt acknowledged 20 Aug 

1852 by the Smithsonian Institution. Primary junior homonym of Sciurus dorsalis Gray, 1848.Renamed Sciurus aberti by Woodhouse (1853a:220).

= Sciurus aberti aberti Woodhouse, 1853. See Miller (1912:333).

Holotype: 2430. Skin only. Sex unknown. San Francisco Mountain (Peaks), [Coconino County], New Mexico (Arizona), 

[Lyon and Osgood (1909:184): on the routes of Captains 

Sitgraves and Whipple, about latitude 35° and longitude 

111°30′]. Collected Oct 1851 by S. W. Woodhouse. No 

original number.

Sciurus fossor Peale

U.S. Explor. Exped. 8: 55, 1848.

= Sciurus griseus griseus Ord, 1818. See Miller (1912:332).

Holotype: No type specimen designated in original description. 

Southern Oregon is thought to be the type locality. Cassin 

(1858:50) states that “Specimen in Mus. Acad. Philadelphia 
number 640 • 2 3

and Nat. Mus. Washington.” There are currently no specimens in the collection that could possibly be a type of this 

taxon.

Sciurus fossor anthonyi Mearns

Proc. U.S. Natl. Mus. 20: 1, 5 Mar 1897a. Preprint of Proc. U.S. 

Natl. Mus. 20: 501, 19 Jan 1898b.

= Sciurus griseus anthonyi Mearns, 1897. See Elliot (1901:57).

Holotype: 60928. Skin and skull. Adult female. Campbell’s 

ranch at Laguna, San Diego County, California. Collected 

10 Jun 1894 by E. A. Mearns. Original number 3642.

Sciurus goldmani Nelson

Proc. Biol. Soc. Washington 12: 149, 3 Jun 1898a.

= Sciurus variegatoides goldmani Nelson, 1898. See Dickey 

(1928:8).

Holotype: 77903. Skin and skull. Adult male. Huehuetan, Chiapas, Mexico. Collected 28 Feb 1896 by E. W. Nelson and 

E. A. Goldman. Original number 9435.

Sciurus granatensis agricolae Hershkovitz

Proc. U.S. Natl. Mus. 97: 16, 25 Aug 1947.

Holotype: 279775. Skin and skull. Adult male. Colonia Agricola 

de Caracolicito, Rio Ariguani, on the southern slopes of the 

Sierra Nevada de Santa Marta, altitude 335 m [SL: 400 m], 

Department of Magdalena, Colombia. Collected 1 Mar 

1942 by P. Hershkovitz. Original number 208.

Sciurus granatensis norosiensis Hershkovitz

Proc. U.S. Natl. Mus. 97: 21, 25 Aug 1947.

Holotype: 279949. Skin and skull. Adult male. Norosi, Mompos, 

altitude 120 m, Department of Bolivar, Colombia. Collected 

26 Jun 1943 by P. Hershkovitz. Original number 2150.

Sciurus granatensis perijae Hershkovitz

Proc. U.S. Natl. Mus. 97: 23, 25 Aug 1947.

Holotype: 279892. Skin and skull. Adult male. Sierra Negra, 

above Villanueva, on the western slope of the Sierra de 

Perija, altitude 1,265 m, Valledupar, Department of Magdalena (Cesar), Colombia. Collected 25 Dec 1942 by P. Hershkovitz. Original number 1230.

Sciurus granatensis tarrae Hershkovitz

Proc. U.S. Natl. Mus. 97: 26, 25 Aug 1947.

Holotype: 279964. Skin and skull. Adult female. Rio Tarra, a 

small tributary of the Upper Catatumbo, San Calixto, altitude about 200 m, Department of Norte de Santander, Colombia. Collected 25 Jul 1943 by P. Hershkovitz. Original 

number 2260.

Sciurus griseoflavus chiapensis Nelson

Proc. Washington Acad. Sci. 1: 69, 9 May 1899b.

= Sciurus aureogaster nigrescens Bennett, 1833. See Musser 

(1970:15).

Holotype: 75957. Skin and skull. Adult male. San Cristobal, 

[FC: altitude 8,400 ft (2,560 m)], Chiapas, Mexico. Collected 22 Sep 1895 by E. W. Nelson and E. A. Goldman. 

Original number [SL: 8447].

Sciurus infuscatus Cabrera

Bol. Soc. Esp. Hist. Nat. 5: 227, Apr 1905.

= Sciurus vulgaris infuscatus Cabrera, 1905. See Hayman and 

Holt (1940:330). See Grill et al. (2009:2687) for current 

usage.

Holotype: [152266. Skin and skull]. Adult male. Navas del 

Marques, altitude 2,410 m, province of Avila, [Castilla y 

Leon], Spain. Collected 26 Oct 1904 by M. M. de la Escalera. No original number. The USNM number was not given 

in the original description as the specimen was in the collection of A. Cabrera at the time. On the back of the original 

label is written “Type of the species” (name given on front 

of label), and the measurements on the label match those in 

the original description.

Sciurus kaibabensis Merriam

Proc. Biol. Soc. Washington 17: 129, 9 Jun 1904a.

= Sciurus aberti kaibabensis Merriam, 1904. See Hoffmeister 

and Diersing (1978:410).

Holotype: 130982. [Skin and skull]. Adult male. Head of Bright 

Angel Creek, top of Kaibab Plateau, north side of Grand Canyon of Colorado, [Coconino County], Arizona. Collected 1 

Dec 1903 by J. T. Stewart. No original number [SL: 4470-X].

Sciurus limitis Baird

Proc. Acad. Nat. Sci. Philadelphia 7: 331, reported favorably for 

publication 24 Apr 1855b, receipt acknowledged 15 Jun 

1855 by the American Philisophical Society.

= Sciurus niger limitis Baird, 1855. See Osgood (1907a:45).

Holotype: [351/1265]. Skin and skull. Sex undetermined. Devils 

River, [Val Verde County], Texas. Collected by J. J. Clark, 

United States and Mexican Boundary Survey, under Major 

W. H. Emory. Cataloged 12 Dec 1854. No original number. 

No specimen number was given in the original description. 

Lyon and Osgood (1909:187) determined this specimen was 

the type.

Sciurus negligens Nelson

Proc. Biol. Soc. Washington 12: 147, 3 Jun 1898a.

= Sciurus deppei negligens Nelson, 1898. See Hooper (1953:4).

Holotype: 93028. [Skin and skull]. Adult female. Alta Mira, [FC: 

altitude about 75 ft (23 m)], Tamaulipas, Mexico. Collected 

18 Apr 1898 by E. A. Goldman. Original number [FC: Nelson and Goldman] 12319.

Sciurus nelsoni Merriam

Proc. Biol. Soc. Washington 8: 144, 29 Dec 1893b.

= Sciurus aureogaster nigrescens Bennett, 1833. See Musser 

(1970:15).
2 4 • smithsonian contributions to zoology

Holotype: 51157. [Skin and skull]. Adult female. Huitzilac, [FC: 

altitude 8,500 ft (2,591 m)], Morelos, Mexico. Collected 

1 Jan 1893 by E. W. Nelson. Original number 4144.

Sciurus nelsoni hirtus Nelson

Proc. Biol. Soc. Washington 12: 153, 3 Jun 1898a.

= Sciurus aureogaster nigrescens Bennett, 1833. See Musser 

(1970:15).

Holotype: 55325. [Skin and skull]. Adult male. Tochimilco, [FC: 

altitude 7,000 ft (2,134 m)], Puebla, Mexico. Collected 

7 Aug 1893 by E. W. Nelson and E. A. Goldman [SL: only 

Nelson]. Original number [FC: EWN] 5295.

Sciurus niger avicennia A. H. Howell

J. Mamm. 1: 37, 28 Nov 1919.

Holotype: 231498. Skin and skull. Adult male. Everglades, Lee 

(Collier) County, Florida. Collected 14 Mar 1919 by A. H. 

Howell. Original number 2325.

Sciurus niger bryanti H. H. Bailey

Bull. Bailey Mus. and Lib. Nat. Hist. 1: 1, 1 Aug 1920.

= Sciurus niger cinereus Linnaeus, 1758. See Barkalow (1954:23).

Holotype: 347443. Skin and skull. Adult female. Dorchester 

County, Maryland. Collected 21 Dec 1917 by a hunter for 

R. W. Jackson. Original number RWJ 2. Previously Bailey 

collection numbers 602 and 123.

Sciurus oculatus tolucae Nelson

Proc. Biol. Soc. Washington 12: 148, 3 Jun 1898a.

Holotype: 55927. [Skin and skull]. Adult male. North slope Volcan 

Toluca, [FC: altitude 11,000 ft (3,353 m)], Mexico, Mexico. 

Collected 8 Sep 1893 by E. W. Nelson and E. A. Goldman 

[SL: only Nelson]. Original number [FC: EWN] 5419.

Sciurus poliopus peregrinator Nelson

Proc. Biol. Soc. Washington 17: 149, 6 Oct 1904.

= Sciurus aureogaster nigrescens Bennett, 1833. See Musser 

(1970:15).

Holotype: 70279. [Skin and skull]. Adult female. Piaxtla, [FC: 

altitude 3,900 ft (1,189 m)], Puebla, Mexico. Collected 25 

Nov 1894 by E. W. Nelson and E. A. Goldman. Original 

number 7104.

Sciurus poliopus senex Nelson

Proc. Biol. Soc. Washington 17: 148, 6 Oct 1904.

= Sciurus aureogaster nigrescens Bennett, 1833. See Musser 

(1970:15).

Holotype: 126208. [Skin and skull]. Adult female. La Salada, 

40 mi (64 km) south of Uruapan, southern Michoacan, 

Mexico. Collected 14 Mar 1903 by E. W. Nelson and E. A. 

Goldman. Original number 16127.

Sciurus richmondi Nelson

Proc. Biol. Soc. Washington 12: 146, 3 Jun 1898a.

Holotype: 36481/48851. Skin and skull. Adult female. Escondido River, [SL: 50 mi (80 km) from] (emended to read 

above by Nelson, 1899b:100) Bluefields, [Atlantico Sur], 

Nicaragua. Collected 4 Oct 1892 by C. W. Richmond. Original number 118.

Sciurus socialis cocos Nelson

Proc. Biol. Soc. Washington 12: 155, 3 Jun 1898a.

= Sciurus aureogaster nigrescens Bennett, 1833. See Musser 

(1970:15).

Holotype: 70644. [Skin and skull]. Adult male. Acapulco, [FC: 

near sea level], Guerrero, Mexico. Collected 11 Jan 1895 by 

E. W. Nelson and E. A. Goldman. Original number 7360.

Sciurus socialis littoralis Nelson

Proc. Biol. Soc. Washington 20: 87, 11 Dec 1907.

= Sciurus aureogaster nigrescens Bennett, 1833. See Musser 

(1970:15).

Holotype: 71322. [Skin and skull]. Adult female. Puerto Angel, 

[FC: 100–300 ft (30–91 m)], Oaxaca, Mexico. Collected 11 

Mar 1895 by E. W. Nelson and E. A. Goldman. Original 

number [SL: 7627].

Sciurus thomasi Nelson

Proc. Washington Acad. Sci. 1: 71, 9 May 1899b.

= Sciurus variegatoides thomasi Nelson, 1899. See Harris (1937:24).

Holotype: 12044/23367. Skin and skull. Sex undetermined. Talamanca, Costa Rica. Collected [MC: 1874] by W. M. Gabb. 

Original number [MC: 45].

Sciurus truei Nelson

Proc. Washington Acad. Sci. 1: 61, 9 May 1899b.

= Sciurus colliaei truei Nelson, 1899. See Anderson (1962:12).

Holotype: 96229. [Skin and skull]. Adult male. Camoa, Rio Mayo, 

Sonora, Mexico. Collected 20 Jan 1899 by E. A. Goldman. 

Original number [FC: Nelson and Goldman] 13405.

Sciurus variabilis choco Goldman

Smithsonian Misc. Coll. 60(22): 4, 28 Feb 1913.

= Sciurus granatensis morulus Bangs, 1900. See Cabrera 

(1961:366).

Holotype: 179561. [Skin and skull]. Adult male. Cana, altitude 

3,500 ft (1,067 m), [Poole and Schantz (1942:559): Pirri 

Mountains], [Darien], eastern Panama. Collected 28 May 

1912 by E. A. Goldman. Original number 21752.

Sciurus variegatoides helveolus Goldman

Smithsonian Misc. Coll. 56(36): 3, 19 Feb 1912a.

Holotype: 171540. [Skin and skull]. Adult male. Corozal, Canal 

Zone (Panama), Panama. Collected 15 Jun 1911 by E. A. Goldman. Original number [FC: Nelson and Goldman] 21166.

Sciurus vulgaris chiliensis Sowerby

Ann. Mag. Nat. Hist. (ser. 9) 7: 253, Mar 1921.
number 640 • 2 5

Holotype: 219185. Skin and skull. Adult female. Wu-ling-shan, 

Tung Ling area, altitude 3,500 ft (1,067 m), 75 mi (121 km) 

northeast of Peking (Beijing), Chihli (Hebei), China. Collected 25 Aug 1917 by A. de C. Sowerby. Original number 

1033.

Sciurus yucatanensis baliolus Nelson

Proc. Biol. Soc. Washington 14: 131, 9 Aug 1901.

Holotype: 107939. [Skin and skull]. Adult male. Apazote, [FC: 

near Yohaltun], Campeche, Mexico. Collected 8 Jan 1901 

by E. A. Goldman. Original number [FC: Nelson and 

Goldman] 14428.

Genus Sundasciurus Moore, 1958

Sciurus balae Miller

Smithsonian Misc. Coll. 45: 14, 6 Nov 1903a.

= Sundasciurus lowii balae (Miller, 1903). See Thorington and 

Hoffmann (2005:787).

Holotype: 121799. Skin and skull. Adult male. Tana Bala (Pulau 

Tanahbala), Batu Islands, [Sumatera Utara, Sumatra, Indonesia]. Collected 12 Feb 1903 by W. L. Abbott. Original 

number 2282.

Sciurus bancarus Miller

Proc. U.S. Natl. Mus. 26: 451, 3 Feb 1903b.

= Sundasciurus tenuis bancarus (Miller, 1903). See Thorington 

and Hoffmann (2005:788).

Holotype: 114311. Skin and skull. Adult male. Pulo Bangkaru, 

Banjak Island, [Aceh, Sumatra, Indonesia]. Collected 17 Jan 

1902 by W. L. Abbott. Original number 1422.

Sciurus hippurellus Lyon

Smithsonian Misc. Coll. 50: 27, 8 Apr 1907f.

= Sundasciurus hippurus borneensis (Gray, 1867). See Thorington and Hoffmann (2005:786).

Holotype: 142274. Skin and skull. Adult female. Batu Ampar, 

on the Landak River, [Kalimantan Barat], western Borneo, 

[Indonesia]. Collected 11 Jul 1905 by W. L. Abbott. Original number 4260.

Sciurus hippurosus Lyon

Smithsonian Misc. Coll. 50: 26, 8 Apr 1907f.

= Sundasciurus hippurus hippurosus (Lyon, 1907). See Thorington and Hoffmann (2005: 786).

Holotype: 141031. Skin and skull. Adult female. Tarussan (Tarusan) Bay, [Sumatera Barat], west coast of Sumatra, [Indonesia]. Collected 18 Dec 1904 by W. L. Abbott. Original 

number 3826.

Sciurus humilis Miller

Smithsonian Misc. Coll. 61 (21): 24, 29 Dec 1913.

= Sundasciurus lowii humilis (Miller, 1913). See Thorington and 

Hoffmann (2005:787).

Holotype: 123116. [Skin and skull]. Immature female, M3 in 

place, permanent premolars just appearing. Kateman River, 

[Riau], East Sumatra, [Indonesia]. Collected 20 Aug 1903 

by W. L. Abbott. Original number 2751.

Sciurus lingungensis Miller

Proc. Washington Acad. Sci. 3: 123, 26 Mar 1901a.

= Sundasciurus lowii natuensis (Thomas, 1895). See Corbet and 

Hill (1992:296).

Holotype: 104693. Skin and skull. Adult male. Pulo Lingung, 

off southern extremity of Bungurun, North Natuna Islands, 

[Kepulauan Riau, Indonesia]. Collected 19 Jun 1900 by 

W. L. Abbott. Original number 494.

Sciurus mansalaris Miller

Proc. U.S. Natl. Mus. 26: 451, 3 Feb 1903b.

= Sundasciurus tenuis (Horsfield, 1824). Unassigned to subspecies by Thorington and Hoffmann (2005:788).

Holotype: 114633. Skin and skull. Adult male. Pulo Mansalar 

(Pulau Musala), off Tapanuli Bay, [Sumatera Utara], [FC: 

west coast of] Sumatra, [Indonesia]. Collected 2 Mar 1902 

by W. L. Abbott. Original number 1583.

Sciurus mansalaris batus Lyon

Proc. U.S. Natl. Mus. 52: 443, 30 Dec 1916.

= Sundasciurus tenuis (Horsfield, 1824). Unassigned to subspecies by Thorington and Hoffmann (2005:788).

Holotype: 121732. Skin and skull. Adult male. Tana Bala (Pulau Tanahbala), Batu Island[s], [Sumatera Utara, Sumatra, Indonesia]. 

Collected 4 Feb 1903 by W. L. Abbott. Original number 2217.

Sciurus parvus Miller

Proc. Biol. Soc. Washington 14: 33, 5 Apr 1901b.

= Sundasciurus tenuis parvus (Miller, 1901). See Thorington and 

Hoffmann (2005:788).

Holotype: 84509. Skin and skull. Adult male. Nulu, altitude 

1,000 ft (305 m), Sarawak, Borneo, [Malaysia]. Collected 

Oct 1894 by C. Hose. No original number.

Sciurus piniensis Miller

Smithsonian Misc. Coll. 45: 14, 6 Nov 1903a.

= Sundasciurus lowii balae (Miller, 1903). See Thorington and 

Hoffmann (2005:787).

Holotype: 121800. Skin and skull. Adult female. Pulo Pinie 

(Pulau Pini), Batu Islands, [Sumatera Utara], Sumatra, [Indonesia]. Collected 1 Mar 1903 by W. L. Abbott. Original 

number 2344.

Sciurus procerus Miller

Proc. Washington Acad. Sci. 3: 122, 26 Mar 1901a.

= Sundasciurus tenuis procerus (Miller, 1901). See Thorington 

and Hoffmann (2005:788).

Holotype: 104698. Skin and skull. Adult male. Bunguran Island, Natunas (North Natuna Islands), [Kepulauan Riau, 
2 6 • smithsonian contributions to zoology

Indonesia]. Collected 18 Jul 1900 by W. L. Abbott. Original 

number 574.

Sciurus pumilus Miller

Smithsonian Misc. Coll. 45: 15, 6 Nov 1903a.

= Sundasciurus fraterculus (Thomas, 1895). See Thorington and 

Hoffmann (2005:786).

Holotype: 121627. Skin and skull. Adult male, not female as 

in original description. South Pagi Island (Pulau Pagai Selatan), [Sumatera Barat], Sumatra, [Indonesia]. Collected 27 

Nov 1902 by W. L. Abbott. Original number 2098.

Sciurus tenuis surdus Miller

Proc. Washington Acad. Sci. 2: 80, 25 Jul 1900b.

= Sundasciurus tenuis tenuis (Horsfield, 1824). See Thorington 

and Hoffmann (2005:788).

Holotype: 84412. Skin and skull. Adult male. Trong (Trang), 

Lower Siam (Thailand). Collected 3 Feb 1897 by [SL: W. L. 

Abbott]. No original number.

Sundasciurus rabori Heaney

Proc. Biol. Soc. Washington 92: 281, 16 Aug 1979.

Holotype: 477989. Skin and skull. Adult male. Magtaguimbong, 

altitude 3,600–4,350 ft (1,097–1,326 m), approximately 

8°48′N, 117°40′E, Mt. Mantalingajan, [Palawan Province], Palawan Island, Republic of the Philippines. Collected 

16 Apr 1962 by D. S. Rabor. Original number [SL: 1444].

Genus Tamias Illiger, 1811

Eutamias amoenus albiventris Booth

The Murrelet 28: 7, 30 Apr 1947.

= Tamias amoenus albiventris (Booth, 1947). See Sutton (1992:2).

Holotype: 275175. Skin and skull. Adult female. Wickiup Spring, 

23 mi (37 km) west of Anatone, on the Asotin-Garfield 

County boundary, Washington. Collected 23 Jul 1943 by 

E. S. Booth. Original number 910 [SL: 32541-X].

Eutamias amoenus ochraceus A. H. Howell

J. Mamm. 6: 54, 9 Feb 1925.

= Tamias amoenus ochraceus (A. H. Howell, 1925). See Hayman 

and Holt (1940:432).

Holotype: 161049. Skin and skull. Adult male. [FC: Head of] 

Studhorse Canyon, [FC: altitude 6,500 ft (1,981 m)], Siskiyou Mountains, [Siskiyou County], California. Collected 

27 Sep 1909 by N. Hollister. Original number 3511.

Eutamias amoenus operarius Merriam

Proc. Biol. Soc. Washington 18: 164, 29 Jun 1905.

= Tamias minimus operarius (Merriam, 1905). See Hayman and 

Holt (1940:431).

Holotype: 129808. [Skin and skull]. Young adult female. Gold 

Hill, [FC: in canyon near Rowena], 7,400 ft (2,256 m), 

[Boulder County], Colorado. Collected 8 Oct 1903 by 

V. Bailey. Original number 8160.

Eutamias amoenus vallicola A. H. Howell

J. Mamm. 3: 179, 4 Aug 1922.

= Tamias amoenus vallicola (A. H. Howell, 1922). See Hayman 

and Holt (1940:432).

Holotype: 168027. Skin and skull. Adult female. Bass Creek, 

near [SL: NW] Stevensville, altitude 3,725 ft (1,135 m), [Ravalli County], Montana. Collected 23 Mar 1910 by C. Birdseye. Original number 1052.

Eutamias asiaticus altaicus Hollister

Proc. Biol. Soc. Washington 25: 183, 24 Dec 1912b.

= Tamias sibiricus sibiricus (Laxmann, 1769). See Ognev 

(1966:421).

Holotype: 175501. Skin and skull. Adult male. Tapucha [SL: 

Dapucha], altitude 6,900 ft (2,103 m), Altai Mountains, 

[Altayaskiy], Siberia (Russia). Collected 8 Aug 1912 by 

N. Hollister. Original number 4474.

Eutamias asiaticus umbrosus A. B. Howell

J. Washington Acad. Sci. 17: 80, 19 Feb 1927.

= Tamias sibiricus umbrosus (A. B. Howell, 1927). See Hayman 

and Holt (1940:436).

Holotype: 240744. Skin and skeleton. Adult male. [FC: Archuan], 140 mi (225 km) south of Lanchowfu (Lanzhou), 

Kansu (Gansu), China. Collected 21 Aug 1923 by F. R. Wulsin. Original number 1118.

Eutamias atristriatus V. Bailey

Proc. Biol. Soc. Washington 26: 129, 21 May 1913a.

= Tamias minimus atristriatus (V. Bailey, 1913). See Hayman and 

Holt (1940:431).

Holotype: 119028. [Skin and skull]. Adult female. On Penasco 

Creek, altitude 7,400 ft (2,256 m), 12 mi (19 km) east of 

Cloudcroft, Sacramento Mountains, New Mexico. Restricted by Frey et al. (2009:269) to “Sacramento Mountains, [Otaro County], New Mexico, on Penasco Creek 

[=James Canyon], 12 mi (19 km) east of Cloudcroft [by 

road], altitude 7,400 ft (2,256 m).” Collected 6 Sep 1902 by 

V. Bailey. Original number 7953.

Eutamias bulleri solivagus A. H. Howell

J. Mamm. 3: 179, 4 Aug 1922.

= Tamias durangae (J. A. Allen, 1903). See Levensen et al. 

(1985:242).

Holotype: 116882. Skin and skull. Adult female. Sierra Guadalupe, [FC: altitude about 9,500 ft (2,896 m)], Coahuila, 

Mexico. Collected 1 May 1902 by E. W. Nelson and E. A. 

Goldman. Original number 15169.

Eutamias canicaudus Merriam

Proc. Biol. Soc. Washington 16: 77, 29 May 1903b.

= Tamias amoenus canicaudus (Merriam, 1903). See Hayman 

and Holt (1940:432).

Holotype: 27007/34428. Skin and skull. Adult female. Spokane [SL and FC: Falls], [FC: altitude 1,700 ft (518 m)], 
number 640 • 2 7

[Spokane County], Washington. Collected 11 Apr 1891 by 

C. P. Streator. Original number 639.

Eutamias caniceps Osgood

N. Amer. Fauna 19: 28, 6 Oct 1900a.

= Tamias minimus caniceps (Osgood, 1900). See Hayman and 

Holt (1940:431).

Holotype: 99200. [Skin and skull]. Adult female. Lake Labarge, 

[SL: Yukon River], Northwest Territory (Yukon), Canada. 

Collected 13 Jul 1899 by W. H. Osgood. Original number 603.

Eutamias caurinus Merriam

Proc. Acad. Nat. Sci. Philadelphia 50: 352, 4 Oct 1898c.

= Tamias amoenus caurinus (Merriam, 1898). See Hayman and 

Holt (1940:432).

Holotype: 90636. Skin and skull. Adult male. Timberline near 

head of Soleduck River, Olympic Mountains, [Clallam 

County], Washington. Collected 27 Aug 1897 by C. H. 

Merriam. Original number [FC: V. Bailey] 6211.

Eutamias cinereicollis canipes V. Bailey

Proc. Biol. Soc. Washington 15: 117, 2 Jun 1902.

= Tamias canipes canipes (V. Bailey, 1902). See Levenson et al. 

(1985:222).

Holotype: 109229. [Skin and skull]. Adult female. [FC: Head 

of Dog Canyon], altitude 7,000 ft (2,134 m), Guadalupe 

Mountains, [Culberson County], Texas. Collected 24 Aug 

1901 by V. Bailey. Original number 7827.

Eutamias cinereicollis cinereus V. Bailey

Proc. Biol. Soc. Washington 26: 130, 21 May 1913a.

= Tamias cinereicollis cinereus (V. Bailey, 1913). See Hayman 

and Holt (1940:434).

Holotype: 167029. [Skin and skull]. Adult male. Copper Canyon, Magdalena Mountains, altitude 8,200 ft (2,499 m), 

[Socorro County], New Mexico. Collected 1 Sep 1909 by 

E. A. Goldman. Original number [FC: Nelson and Goldman] 20435.

Eutamias consobrinus clarus V. Bailey

Proc. Biol. Soc. Washington 31: 31, 16 May 1918.

= Tamias minimus consobrinus J. A. Allen, 1890. See Hayman 

and Holt (1940:431).

Holotype: 227313. [Skin and skull]. Adult male. Swan Lake Valley, Yellowstone National Park, [SL: Wyoming]. Collected 

13 Sep 1917 by V. Bailey. Original number 9945.

Eutamias dorsalis carminis Goldman

Proc. Biol. Soc. Washington 51: 56, 18 Mar 1938a.

= Tamias dorsalis carminis (Goldman, 1938). See Levenson et 

al. (1985:222).

Holotype: 263378. Skin and skull. Young male. Carmen Mountains, altitude 7,400 ft (2,256 m), Coahuila, Mexico. Collected 20 Sep 1937 by R. S. Sturgis [XC: shot by W. H. 

Hazlette]. Original number 73. [SL: 28066-X].

Eutamias dorsalis sonoriensis Callahan and Davis

Southwestern Nat. 22: 71, 1 Mar 1977.

= Tamias dorsalis sonoriensis (Callahan and Davis, 1977). See 

Hart (1992:1).

Holotype: [398643]. Skin, skull, and stained baculum. Adult 

male. Twenty-four kilometers NE Guaymas [SL: 1 mi 

(1.6 km) SW Maytorena], altitude 100 m [SL: about 300 ft 

(91 m)], Sonora, Mexico. Collected 13 Nov 1973 by J. Callahan. Original number 123. University of Arizona number 

22788.

Eutamias dorsalis utahensis Merriam

Proc. Biol. Soc. Washington 11: 210, 1 Jul 1897b.

= Tamias dorsalis utahensis (Merriam, 1897). See Elliot 

(1901:69).

Holotype: 186457. Skin and skull. Adult male. Ogden [MCC: 

foothills of Wasatch Mountains], [Weber County], Utah. 

Collected 9 Oct 1888 by V. Bailey. Original number 289. 

Merriam collection number 4788/5448.

Eutamias hopiensis Merriam

Proc. Biol. Soc. Washington 18: 165, 29 Jun 1905.

= Tamias sp. Illiger, 1811. See Thorington and Hoffmann 

(2005:816) and Patterson (1984:452).

Holotype: 67768. [Skin and skull]. Adult female. Keam Canyon [FC: 80 mi (129 km) N of Holbrook], Painted Desert, 

[Navajo County], Arizona. Collected 27 Jul 1894 by A. K. 

Fisher. Original number 1688.

Eutamias ludibundus Hollister

Smithsonian Misc. Coll. 56 (26): 1, 5 Dec 1911a.

= Tamias amoenus ludibundus (Hollister, 1911). See Hayman 

and Holt (1940:432).

Holotype: 174225. Skin and skull. Adult female. Yellowhead 

Lake, altitude 3,700 ft (1,128 m) [SL and FC: 3,614 ft 

(1,102 m)], British Columbia, Canada. Footnote states 

“This is the Cowdung Lake of many maps. It lies about six 

miles (9.7 km) west of the Alberta-British Columbian line, 

in Yellowhead Pass.” Collected 29 Aug 1911 by N. Hollister. Original number 3987.

Eutamias merriami meridionalis Nelson and Goldman

Proc. Biol. Soc. Washington 22: 23, 10 Mar 1909.

= Tamias obscurus meridionalis (Nelson and Goldman, 1909). 

See Best and Granai (1994:1).

Holotype: 139597. [Skin and skull]. Adult female. Aguaje de San 

Estaban, about 25 mi (40 km) northwest of San Ignacio, altitude about 1,200 ft (366 m), Lower California (Baja California Sur), Mexico. Collected 5 Oct 1905 by E. W. Nelson 

and E. A. Goldman. Original number 18268.

Eutamias minimus arizonensis A. H. Howell

J. Mamm. 3: 178, 4 Aug 1922.

= Tamias minimus operarius (Merriam, 1905). See Thorington 

and Hoffmann (2005:815) and Conley (1970:700).
2 8 • smithsonian contributions to zoology

Holotype: 205869. Skin and skull. Adult male. Prieto Plateau 

at south end of Blue Range, [SL and FC: altitude 9,000 ft 

(2,743 m)], Greenlee County, Arizona. Collected 7 Sep 1914 

by E. G. Holt. Original number 384.

Eutamias minimus caryi Merriam

Proc. Biol. Soc. Washington 21: 143, 9 Jun 1908a.

= Tamias minimus caryi (Merriam, 1908). See Hayman and Holt 

(1940: 431).

Holotype: 150740. [Skin and skull]. Young adult male. Medano 

Ranch, [SL: 15 mi (24 km) NE of Mosca], [FC: altitude 

7,600 ft (2,316 m)], San Luis Valley, [Costilla County], Colorado. Collected 24 Oct 1907 by M. Cary. Original number 

1176.

Eutamias minimus confinis A. H. Howell

J. Mamm. 6: 52, 9 Feb 1925.

= Tamias minimus confinis (A. H. Howell, 1925). See Hayman 

and Holt (1940:431).

Holotype: 168957. Skin and skull. Adult female. Head of Trapper’s Creek, altitude 8,500 ft (2,591 m), west slope of Bighorn Mountains, [Big Horn County], Wyoming. Collected 

7 Jun 1910 by M. Cary. Original number 1956.

Eutamias minimus grisescens A. H. Howell

J. Mamm. 6: 52, 9 Feb 1925.

= Tamias minimus grisescens (A. H. Howell, 1925). See Hayman 

and Holt (1940:431).

Holotype: 89701. Skin and skull. Sub-adult male. Farmer, Douglas County, Washington. Collected 31 Jul 1897 by J. A. Loring. Original number 4539.

Eutamias minimus jacksoni A. H. Howell

J. Mamm. 6: 53, 9 Feb 1925.

= Tamias minimus neglectus J. A. Allen, 1890. See Levenson et 

al. (1985:222).

Holotype: 227423. Skin and skull. Adult male. Crescent Lake, 

Oneida County, Wisconsin. Collected 7 Sep 1917 by H. H. T. 

Jackson. Original number 927.

Eutamias obscurus davisi Callahan

J. Mamm. 58: 193, 31 May 1977.

= Tamias obscurus davisi (Callahan, 1977). See Best and Granai 

(1994:1).

Holotype: 398642. Skin, skull, and stained baubellum. Adult 

female [SL: lactating]. Barker’s Reservoir, altitude 1,300 

m, 16 km SW Twentynine Palms, San Bernardino County, 

California. Collected 5 Aug 1974 by J. Callahan. Original 

number 143 [SL: University of Arizona number 22761].

Eutamias oreocetes Merriam

Proc. Biol. Soc. Washington 11: 207, 1 Jul 1897b.

= Tamias minimus oreocetes (Merriam, 1897). See Hayman and 

Holt (1940:431).

Holotype: 72468. [Skin and skull]. Nursing adult female. Timberline near Summit, [FC: altitude 7,500 ft (2,286 m)], 

Teton Mountains [Poole and Schantz (1942:529): on Great 

Northern Railroad, Flathead County], Montana. Collected 

14 Jun 1895 by V. Bailey. Original number 5024.

Eutamias pallidus cacodemus Cary

Proc. Biol. Soc. Washington 19: 89, 4 Jun 1906.

= Tamias minimus cacodemus (Cary, 1906). See Hayman and 

Holt (1940:431).

Holotype: 138737, not 138137 as in original description. [Skin 

and skull]. Adult female. [FC: Head of Corral Draw, 40 mi 

(64 km) east of Hermosa], Sheep Mountain, Big Badlands, 

[Pennington County], South Dakota. Collected 2 Sep 1905 

by M. Cary. Original number 682.

Eutamias palmeri Merriam

Proc. Biol. Soc. Washington 11: 208, 1 Jul 1897b.

= Tamias palmeri (Merriam, 1897). See Elliot (1901:69).

Holotype: 26075/33481. Skin and skull. Adult male. Charleston 

Peak, altitude about 2,450 m or 8,000 ft, [Clark County], 

Nevada. Collected 13 Feb 1891 by T. S. Palmer and E. W. 

Nelson. Original number [FC: EWN] 432.

Eutamias quadrivittatus animosus Warren

Proc. Biol. Soc. Washington 22: 105, 25 Jun 1909.

= Tamias quadrivittatus (Say, 1823). See Bachman (1839:74).

Holotype: [154163. Skin and skull]. Adult female. Irwin Ranch, 

T29S, R52W., altitude 5,000 ft (1,524 m), Las Animas 

County, Colorado. Collected 29 Apr 1909 by E. R. Warren. 

Original number 3428.

Eutamias quadrivittatus australis Patterson

J. Mamm. 61: 460, 20 Aug 1980.

= Tamias quadrivittatus (Say, 1823). See Bachman (1839:74).

Holotype: 530290. Skin, skull, and baculum. Adult male. Three 

and one-half miles (5.6 km) south, 1.5 mi (2.4 km) east 

Organ, 6,800 ft (2,073 m), North Fork Pine Tree Trail, Aguirre Springs, T22S R4E Sec. 30, Dona Ana County, New 

Mexico. Collected 12 Sep 1978 by B. D. Patterson. Original 

number 169 [SL: New Mexico State University Department 

of Biology number 7893].

Eutamias ruficaudus A. H. Howell

Proc. Biol. Soc. Washington 33: 91, 30 Dec 1920.

= Tamias ruficaudus ruficaudus (A. H. Howell, 1920). See Hayman and Holt (1940:434).

Holotype: 72294. Skin and skull. Adult male [SL and FC: 

female]. Upper St. Mary’s Lake, [Glacier County], Montana. Collected 30 May 1895 by A. H. Howell. Original 

number 27.

Eutamias ruficaudus simulans A. H. Howell

J. Mamm. 3: 179, 4 Aug 1922.
number 640 • 2 9

= Tamias ruficaudus simulans (A. H. Howell, 1922). See Hayman and Holt (1940:434).

Holotype: 28487/40591. Skin and skull. Adult female. Coeur 

d’Alene, [Kootenai County], Idaho. Collected 1 Jun 1891 

by C. P. Streator. Original number 881.

Eutamias senescens Miller

Proc. Acad. Nat. Sci. Philadelphia 50: 349, 4 Oct 1898.

= Tamias sibiricus senescens (Miller, 1898). See Hayman and 

Holt (1940:436).

Holotype: 83395. Skin and skull. Adult female. Low barren hills, 

15 mi (24 km) west of Peking (Beijing), [SL: Western Hills], 

[Beijing], China. Collected 21 Aug 1896 by G. D. Wilder. 

No original number.

Eutamias sibiricus barberi Johnson and Jones

Proc. Biol. Soc. Washington 68: 175, 31 Dec 1955.

= Tamias sibiricus barberi (Johnson and Jones, 1955). See Koh 

et al. (2009:1).

Holotype: 298042. Skin and skull. Adult female. Central National Forest, near Puo’ yong-ni, 37°44′N, 127°12′E, 

[South] Korea. Collected 26 Aug 1952 by W. H. Lawrence. 

Original number B12068.

Eutamias speciosus inyoensis Merriam

Proc. Biol. Soc. Washington 11: 208, 1 Jul 1897b.

= Tamias umbrinus inyoensis (Merriam, 1897). See Levenson et 

al. (1985:223).

Holotype: 29387/41462. Skin and skull. Young adult male. [FC: 

4,600 ft (1,402 m) above Black Canyon = 8,238 ft (2,511 

m), west slope] White Mountains, Inyo County, California. 

Collected 7 Jul 1891 by E. W. Nelson. Original number 

1069.

Eutamias speciosus sequoiensis A. H. Howell

J. Mamm. 3: 180, 4 Aug 1922.

= Tamias speciosus sequoiensis (A. H. Howell, 1922). See Levenson et al. (1985:222).

Holotype: 30899/42799. Skin and skull. Adult female. Mineral 

King, east fork of Kaweah River, altitude 7,300 ft (2,225 

m), [Tulare County], California. Collected 12 Sep 1891 by 

V. Bailey. Original number 3259.

Eutamias townsendii alleni A. H. Howell

J. Mamm. 3: 181, 4 Aug 1922.

= Tamias sonomae alleni (A. H. Howell, 1922). See Best (1993:1).

Holotype: 135177. Skin and skull. Adult male. Inverness, Marin 

County, California. Collected 16 Nov 1904 by N. Hollister. 

Original number 1378.

Eutamias townsendii ochrogenys Merriam

Proc. Biol. Soc. Washington 11: 195, 1 Jul 1897b.

= Tamias ochrogenys (Merriam, 1897). See Gannon et al. 

(1993:1).

Holotype: 67182. [Skin and skull]. Adult male. Mendocino, 

[Mendocino County], California. Collected 17 Jul 1894 by 

J. E. McLellan. Original number 1015.

Eutamias townsendii siskiyou A. H. Howell

J. Mamm. 3: 180, 4 Aug 1922.

= Tamias siskiyou siskiyou (A. H. Howell, 1922). See Gannon 

et al. (1993:1).

Holotype: 161033. Skin and skull. Adult female. Near summit 

of White Mountain, Siskiyou Mountains, altitude 6,000 ft 

(1,829 m), [FC: Siskiyou County], California. Collected 16 

Sep 1909 by N. Hollister. Original number 3432.

Eutamias umbrinus sedulus White

Univ. Kansas Publ., Mus. Nat. Hist. 5: 573, 1 Dec 1953.

= Tamias umbrinus sedulus (White, 1953). See Bergstrom (1999:393).

Holotype: 158181. Skin, skull, and baculum. Adult male. Mount 

Ellen, Henry Mountains, Garfield County, Utah. Collected 

13 Oct 1908 by W. H. Osgood. Original number 3667.

Tamias alpinus Merriam

Proc. Biol. Soc. Washington 8: 137, 28 Dec 1893a.

Holotype: 30507/42491. Skin and skull. Young adult female. 

[FC: Head] Big Cottonwood Meadows, High Sierra, altitude 3,050 m or 10,000 ft, just south of Mount Whitney, 

[Inyo County], California. Collected 12 Aug 1891 by B. H. 

Dutcher. Original number 191.

Tamias amoenus J. A. Allen

Bull. Amer. Mus. Nat. Hist. 3: 90, Jun 1890.

= Tamias amoenus amoenus J. A. Allen, 1890. See Hayman and 

Holt (1940:432).

Holotype: 186460. Skin and skull. Adult female. Fort Klamath, 

[Klamath County], Oregon. Collected 16 May 1887 by J. C. 

Merrill. [MCC: Original number 5]. Merriam collection 

number 3469/[MCC: 4096].

Tamias asiaticus borealis J. A. Allen

Monog. N. Amer. Rodentia 793, Aug 1877.

= Tamias minimus borealis J. A. Allen, 1877. See Hayman and 

Holt (1940:431).

Lectotype: 6506. [Skin, broken skull inside]. Fort Laird, British 

America (Mackenzie District, Northwest Territories, Canada). Collected [Poole and Schantz (1942:524): about 1860] 

by W. S. Hardisty. Cataloged 1863. Original number [SL: 

1086]. No type designated in original description. Lectotype 

selected by J. A. Allen (1890:109).

Tamias callipeplus Merriam

Proc. Biol. Soc. Washington 8: 136, 28 Dec 1893a.

= Tamias speciosus callipeplus Merriam, 1893. See Best et al. 

(1994:1).

Holotype: 31299/43164. Skin and skull. Young adult male. Summit of Mount Pinos [SL and FC: Mt. McGill], altitude [Lyon 
3 0 • smithsonian contributions to zoology

and Osgood (1909:174): 9,000 ft (2,743 m)], or [Miller and 

Kellogg (1955:231): 8,800 ft (2,682 m)], Ventura County, 

California. Collected 20 October 1891 by E. W. Nelson. 

Original number 1344.

Tamias cinereicollis J. A. Allen

Bull. Amer. Mus. Nat. Hist. 3: 94, Jun 1890.

= Tamias cinereicollis cinereicollis J. A. Allen, 1890. See Hayman 

and Holt (1940:434).

Holotype: 17597/[24533]. Skin and skull. Adult female. [FC: 

Little Spring], San Francisco Mountain (Peaks), [Coconino 

County], Arizona. Collected 2 Aug 1889 by C. H. Merriam 

and V. Bailey. Original number [FC: VB], [SL: 260].

Tamias cooperi Baird

Proc. Acad. Nat. Sci. Philadelphia 7: 334, reported favorably for 

publication 24 Apr 1855a.

= Tamias townsendii cooperi Baird, 1855. See Baird (1857:300).

Syntypes: 212/1183. Skin and skull. On eastern side of the Cascade Mountains, Washington Territory. Emended by Cooper 

(1868:531) to read Clickitat (Klickitat) Pass, altitude 4,500 ft 

(1,372 m), Cascade Mountains, [Skamania County], Washington. Collected Jul 1853 by J. G. Cooper. No original number.

211/1182. Skin only. Other data as given above. This specimen 

is now number 4754, Museum of Comparative Zoology, 

Harvard University.

No type is designated in the original description, but Baird 

(1857:301) states that Tamias cooperi was based on the two 

specimens listed above.

Tamias dorsalis Baird

Proc. Acad. Nat. Sci. Philadelphia 7: 332, reported favorably for 

publication 24 Apr 1855a, receipt acknowledged 15 Jun 

1855 by the American Philisophical Society.

= Tamias dorsalis dorsalis Baird, 1855. See Hayman and Holt 

(1940:435).

Syntypes: 120. Mounted specimen with skull inside. On the 

Mimbres [MC: Coppermines, Gila]. Baird (1857:707) in 

his list of localities says of Coppermines, “A former station of the United States-Mexican Boundary Survey, subsequently called Fort Webster. Not indicated clearly whether 

on a tributary of the Gila or Mimbres. About Latitude 33°, 

Longitude 108°.” Baird (1857:709) says of Fort Webster 

“See also Coppermines. Altitude 6,350 ft (1,935 m). Latitude 32°47′, Longitude 108°04′.” J. P. Hubbard, who has 

researched old New Mexico collecting localities for decades, 

indicates the proper locality designation should read “On 

the Mimbres [River = the copper mines = former Santa Rita 

del Cobre in present Grant County, New Mexico (approximately 32°47.560′N, 108°4.710′W)]” J. P. Hubbard (in litt., 

26 October 2010 to A. L. Gardner). Collected 1851 by J. H. 

Clark. No original number.

119/3151. Skin and partial mandible. Other data as given above. 

This specimen is now number 4759, Museum of Comparative Zoology, Harvard University.

No type is designated in the original description, but Baird 

(1857:300) listed these two specimens as those on which 

Tamias dorsalis was based.

Tamias macrorhabdotes Merriam

Proc. Biol. Soc. Washington 3: 25, 27 Jan 1886c.

= Tamias quadrimaculatus Gray, 1867. See Hayman and Holt 

(1940:435).

Lectotype: [186458. Skin and skull.] Sierra Nevada Mountains 

of central California [SL: Blue Canon (Canyon)], [Placer 

County] California. Collected [SL: 6 Oct 1885] by C. A. 

Allen. Original number [MCC: 1] [SL: Merriam collection 

number 1488/2109].

No type is designated in the original description. Merriam 

(1897b:203) states that the type locality for his macrorhabdotes is Blue Canon (Canyon) but again does not indicate 

a type and we have not found a published specimen number associated with this name prior to Poole and Shantz 

(1942:527). There is compelling reason to believe that Merriam selected this specimen because it was cataloged in a 

separate block in the USNM catalog along with the other 

type specimens from his personal collection upon donation 

in 1911. It is also delineated as a type in his private collection catalog. In order to remove ambiguity to the application of this name, we designate this specimen as the 

Lectotype.

Tamias minimus consobrinus J. A. Allen

Bull. Amer. Mus. Nat. Hist. 3: 112, Jun 1890.

Holotype: [186456]. Skin and skull. Adult male. [Poole and 

Schantz (1942:528): Parley’s Canyon], Wasatch foothills, 18 

mi (29 km) east of Salt Lake City, [Salt Lake County], Utah. 

Collected 31 Oct 1888 by V. Bailey. Original number [SL: 

361]. Merriam collection number 4883/[5630].

Tamias minimus melanurus Merriam

N. Amer. Fauna 4: 22, 8 Oct 1890a.

= Tamias minimus pictus J. A. Allen, 1890. See Hayman and 

Holt (1940:431).

Holotype: 23048/30494. Skin and skull. Adult male [FC: immature male]. West side of Snake River, near Blackfoot, [Bingham County], Idaho. Collected 17 Jul 1890 by V. Bailey and 

B. H. Dutcher. Original number [FC: VB] 1451.

Tamias minimus pictus J. A. Allen

Bull. Amer. Mus. Nat. Hist. 3: 115, Jun 1890.

Holotype: [186459]. Skin and skill. Adult male. Kelton, [Box 

Elder County], Utah. Collected 25 Oct 1888 by V. Bailey. 

Original number [SL: 342]. Merriam collection number 

4882/[SL: 5629].

Tamias obscurus J. A. Alllen

Bull. Amer. Mus. Nat. Hist. 3: 70, Jun 1890.

= Tamias obscurus obscurus J. A. Allen, 1890. See Best and Granai (1994:1).
number 640 • 3 1

Holotype: 18050/[24954]. Skin and skull. Adult female. San 

Pedro [Martir] Mountains, [Poole and Schantz (1942:529): 

near Vallecitos], Lower California (Baja California), Mexico. Collected 1 May 1889 by C. H. Townsend. Original 

number [SL: 7].

Tamias panamintinus Merriam

Proc. Biol. Soc. Washington, 8: 134, 28 Dec 1893a.

= Tamias panamintinus panamintinus Merriam, 1893. See Hayman and Holt (1940:432).

Holotype: 27603/39702. Skin and skull. Adult male. Johnson 

Canyon, [FC: altitude about 6,000 ft (1,829 m)], between 

Death Valley and Panamint Valley, Panamint Mountains, [Inyo County], California. Restricted by Grinnell 

(1933:128) to vicinity of Hungry Bill’s Ranch, about 5,000 

ft (1,534 m). Collected 3 Apr 1891 by E. W. Nelson. Original number 723.

Tamias quadrivittatus gracilis J. A. Allen

Bull. Amer. Mus. Nat. Hist. 3: 99, Jun 1890.

= Tamias quadrivittatus (Say, 1823). See Bachman (1839:74).

Holotype: 17144/[24077]. Skin and skull. [SL: Hardly adult; FC: 

young of year] female. San Pedro, [Socorro County], New 

Mexico. Collected 4 Jul 1889 by V. Bailey. Original number 

[SL: 166].

Tamias quadrivittatus luteiventris J. A. Allen

Bull. Amer. Mus. Nat. Hist. 3: 101, Jun 1890.

= Tamias amoenus luteiventris J. A. Allen, 1890. See Elliot 

(1901:76).

Holotype: 11991/[37996]. Skin and skull. Adult male. Chief 

Mountain (Waterton) Lake, [Poole and Schantz (1942:531): 

3.5 mi (5.6 km) north of the United States–Canada boundary, Alberta, Canada]. Collected 24 Aug 1874 by E. Coues. 

Original number [SL: 4596].

Tamias quadrivittatus pallidus J. A. Allen

Proc. Boston Soc. Nat. Hist. 16: 289, 1874.

= Tamias minimus pallidus J. A. Allen, 1874. See Hayman and 

Holt (1940:431).

Lectotype: 11656/[38311]. Skin and skull. [Lyon and Osgood 

(1909:177): Nearly adult]. Camp Thorne, near present 

town of Glendive, Yellowstone River, [Dawson County], 

Montana. Collected 18 Jul 1873 by J. A. Allen. Original 

number [SL: 200]. No type designated in original description. Lectotype designated by Cary (1906:88).

Tamias senex J. A. Allen

Bull. Amer. Mus. Nat. Hist. 3: 83, Jun 1890.

= Tamias senex senex J. A. Allen, 1890. See Sutton and Patterson 

(2000:307).

Holotype: [186461]. Skin and skull. Adult, sex undetermined. 

Summit of Donner Pass, Placer County, California. Collected 1 Jul 1885 by L. Belding. Original number [MCC: 3]. 

Merriam collection number 1133/[5882].

Tamias speciosus Merriam

In J. A. Allen, Bull. Amer. Mus. Nat. Hist. 3: 86, Jun 1890.

= Tamias speciosus speciosus Merriam, 1890. See Best et al. (1994:1).

Holotype: [186462]. Skin and skull. Adult [FC and MCC: immature] male. [Poole and Schantz (1942:532): Head of Whitewater Creek], altitude 7,500 ft (2,286 m), San Bernardino 

Mountains, [San Bernardino County], California. Collected 

22 Jun 1885 by F. Stephens. Original number [MCC: 108]. 

Merriam collection number 1148/1804.

Tamias striatus doorsiensis Long

Proc. Biol. Soc. Washington 84: 201, 4 Nov 1971.

Holotype: [565665]. Skin and skull. Adult male. Peninsula State 

Park, Door County, Wisconsin. Collected 28 Sep 1969 by 

J. Small. Original number 15. Wisconsin State University 

Museum of Natural History number 1633.

Tamias striatus fisheri A. H. Howell

J. Mamm. 6: 51, 9 Feb 1925.

Holotype: 193370. Skin and skull. Sub-adult female. Merritts’ 

Corners, 4 mi (6.4 km) west [SL: east] of Sing Sing (Ossining), 

[Westchester County], New York. Type locality restricted by 

Paradiso (1963:580) to Merritts Corners, approximately 3.75 

mi (6 km) east and 1.75 mi (3 km) north of Ossining, Westchester County, New York, 41°11′27′′N latitude, 73°47′51′′W 

longitude Collected 23 Aug 1884 by A. K. Fisher. No original 

number. Merriam collection number 801/1338.

Tamias umbrinus J. A. Allen

Bull. Amer. Mus. Nat. Hist. 3: 96, Jun 1890.

= Tamias umbrinus umbrinus J. A. Allen, 1890. See Levenson et 

al. (1985:223).

Holotype: [186463]. Skin and skull. Male. Unita Mountains, 

south of Fort Bridger [SL and MCC: originally said Fort 

Bridger, Wyoming; Uinta Mountains, Utah, 30 mi (48 km) 

south of Fort Bridger was added later]. Type locality restricted by A. H. Howell (1929a:94) to Blacks Fork, altitude 

about 9,500 ft (2,896 m), [Summit County], Uinta Mountains, Utah. Collected 20 [SL and MCC: 19] Sep 1888 by 

V. Bailey. Original number [SL and MCC: 228]. Merriam 

collection number 4690/[5341].

Genus Tamiasciurus Trouessart, 1880

Sciurus douglasii cascadensis J. A. Allen

Bull. Amer. Mus. Nat. Hist. 10: 277, 22 Jul 1898.

= Tamiasciurus douglasii mollipilosus (Audubon and Bachman, 

1841). See Anderson (1947:121).

Holotype: 80229. Skin and skull. Adult male. Mount Hood, 

[FC: W. base below timberline, altitude 5,000 ft (1,524 m)], 

Oregon. Collected 9 Sep 1896 by V. Bailey. Original number 

[SL: 5874].

Sciurus fremonti neomexicanus J. A. Allen

Bull. Amer. Mus. Nat. Hist. 10: 291, 22 Jul 1898.
3 2 • smithsonian contributions to zoology

= Tamiasciurus hudsonicus mogollonensis (Mearns, 1890). See 

Kelson (1951a:17).

Holotype: 71690. [Skin and skull]. Adult male. Rayado Canyon, Colfax County, New Mexico. Collected 1 Apr 1895 by 

C. M. Barber. Original number [SL: 89].

Sciurus hudsonicus abieticola A. H. Howell

J. Mamm. 10: 75, 11 Feb 1929b.

= Tamiasciurus hudsonicus abieticola (A. H. Howell, 1929). See 

Kellogg (1937:459).

Holotype: 193638. Skin and skull. Adult female. Highlands, 

[Macon County], North Carolina. Collected 9 Mar 1886 

by C. L. Boynton. Original number 30. Merriam collection 

number 2296/2806.

Sciurus hudsonicus baileyi J. A. Allen

Bull. Amer. Mus. Nat. Hist. 10: 261, 22 Jul 1898.

= Tamiasciurus hudsonicus baileyi (J. A. Allen, 1898). See Hayman and Holt (1940:346).

Holotype: 56040. [Skin and skull]. Adult male. [FC: West slope] 

Bighorn Mountains, altitude 8,440 ft (2,560 m), [Washakie 

County], Wyoming. Collected 19 Sep 1893 by C. H. Merriam, not V. Bailey as in original description. Original number [SL: V. Bailey 4390].

Sciurus hudsonicus mearnsi Townsend

Proc. Biol. Soc. Washington 11: 146, 9 Jun 1897.

= Tamiasciurus mearnsi (Townsend, 1897). See Lindsay (1981:680).

Holotype: 8266/25170. Skin and skull. Age and sex undetermined. 

San Pedro Martir Mountains, at about 7,000 ft (2,134 m), 

Lower California (Baja California), [Mexico]. Collected May 

1889 by C. H. Townsend. No original number.

Sciurus hudsonicus petulans Osgood

N. Amer. Fauna 19: 27, 6 Oct 1900a.

= Tamiasciurus hudsonicus petulans (Osgood, 1900). See A. H. 

Howell (1936:136).

Holotype: 97457. Skin and skull. Adult female. White Pass [FC: 

glacier or summit], altitude 1,870 ft (570 m), southern Alaska. 

Collected 4 Jun 1899 by W. H. Osgood. Original number 370.

Sciurus hudsonicus ventorum J. A. Allen

Bull. Amer. Mus. Nat. Hist. 10: 263, 22 Jul 1898.

= Tamiasciurus hudsonicus ventorum (J. A. Allen, 1898). See 

Davis (1939:229).

Holotype: 56030. Skin and skull. Adult [SL and FC: male], not 

female as given in original description. South Pass City, [FC: 

6 mi (10 km) north, altitude 8,000 ft (2,438 m)], [Wind 

River Mountains, Fremont County], Wyoming. Collected 6 

Sep 1893 by V. Bailey. Original number [SL: 4305].

Sciurus suckleyi Baird

Proc. Acad. Nat. Sci. Philadelphia 7: 333, reported favorably for 

publication 24 Apr 1855a, receipt acknowledged 15 Jun 

1855 by the American Philisophical Society.

= Tamiasciurus douglasii douglasii (Bachman, 1839). See Hayman and Holt (1940:347).

Holotype: 272/[37606]. Skin and skull. Adult male. [SL and 

MC: Fort] Steilacoom, Puget Sound, [Pierce County], Washington. Collected [MC: 15 Jan 1854] by G. Suckley. Original number 1.

No type designated in the original description, but three measurements of a specimen were given. In a later publication, 

Baird (1857:276) gives further details about the taxon and 

in the table on page 278 only one specimen, number 272, 

has the same measurements as given in the original description that led Lyon and Osgood (1909:194) and subsequent 

authors to consider it the type.

Tamiasciurus hudsonicus kenaiensis A. H. Howell

Proc. Biol. Soc. Washington 49: 136, 22 Aug 1936.

Holotype: 107603. Skin and skull. Adult male. Hope [FC: City], 

Cook Inlet, Alaska. Collected 6 Sep 1900 by W. H. Osgood. 

Original number 1360.

Tamiasciurus hudsonicus murii A. H. Howell

Proc. Biol. Soc. Washington 56: 67, 16 Jun 1943.

= Tamiasciurus hudsonicus minnesota (J. A. Allen, 1899). See 

Anderson (1947:119).

Holotype: 234039. Skin and skull. Adult male. Moorhead, [Clay 

County], Minnesota. Collected 11 Feb 1918 by O. J. Murie. 

Original number 1174. 21595-X.

Tamiasciurus hudsonicus pallescens A. H. Howell

Proc. Biol. Soc. Washington 55: 13, 12 May 1942.

Holotype: 261625. Skin and skull. Adult female. Eight miles (13 km) 

east of Upham, McHenry County, North Dakota. Collected 8 

Jan 1936 by C. J. Henry. Original number H-135. 27532-X.

Tamiasciurus hudsonicus preblei A. H. Howell

Proc. Biol. Soc. Washington 49: 133, 22 Aug 1936.

Holotype: 133862. Skin and skull. Adult [SL and FC: male], 

not female as given in original description. Fort Simpson, 

Mackenzie [District, Northwest Territories, Canada]. Collected 29 Feb 1904 by E. A. Preble. Original number 5141.

Tamiasciurus hudsonicus wasatchensis Hardy

Proc. Biol. Soc. Washington 63: 13, 26 Apr 1950.

= Tamiasciurus hudsonicus fremonti (Audubon and Bachman, 

1853). See Durrant and Hansen (1954:91).

Holotype: 276398. Skin and skull. Adult male. About 10,000 

ft (3,048 m) in spruce fir area along Skyline Drive east of 

Mount Nebo, Juab County, near Juab–Utah County line, 

Utah. Collected 19 Aug 1942 by R. Hardy. Original number 

2431. 33443-X.

Genus Tamiops J. A. Allen, 1906

Sciurus novemlineatus Miller

Proc. Biol. Soc. Washington 16: 147, 12 Nov 1903c.
number 640 • 3 3

= Tamiops mcclellandi leucotis (Temminck, 1853). See Moore 

and Tate (1965:242).

Holotype: 84403. Skin and skull. Adult male. In heavy forest 

among the hills of Trong (Trang), altitude 1,500 ft (457 m), 

Lower Siam (Thailand). Collected 19 Feb 1897 by W. L. Abbott. No original number.

Tamiops mcclellandi liantis Kloss

J. Nat. Hist. Soc. Siam 3: 370, 31 Dec 1919.

= Tamiops rodolphii rodolphii (Milne-Edwards, 1867). See 

Moore and Tate (1965:244).

Holotype: [221542]. Skin and skull. Adult female. Satahip (Sattahip), near Cape Liant, [Poole and Schantz (1942:562): 

Shelter Bay], [Chon Buri], southeast Siam (Thailand). Collected 2 Nov 1916 by C. B. Kloss. Original number 2337.

Tamiops rodolphii elbeli Moore

Amer. Mus. Novitates 1879: 4, 28 Feb 1958.

Holotype: 294876. Skin and skull. Female. Ban Lad, Pookeio 

(Amphoe Phu Khieo) District, Chaiyaphum Province, Thailand, about latitude 16°40′N, longitude 101°45′E. Collected 

18 Jan 1952 by R. E. Elbel. Original number RE 369.

Tamiops vestitus Miller

Proc. Biol. Soc. Washington 28: 115, 27 May 1915.

= Tamiops swinhoei vestitus Miller, 1915. See Moore and Tate 

(1965:250).

Holotype: 199561. Skin and skull. Adult male. Hsin-lung-shan, 

[SL: 3,000 ft (914 m)], south of Jehol and 65 mi (105 km) 

northeast of Peking [Beijing], Chi li (Hebei), China. Collected 

15 Feb 1915 by A. de C. Sowerby. Original number 754.

Genus Urocitellus Obolenskij, 1927

Citellus columbianus ruficaudus A. H. Howell

Proc. Biol. Soc. Washington 41: 212, 18 Dec 1928.

= Urocitellus columbianus ruficaudus (A. H. Howell, 1928). See 

Helgen et al. (2009:297).

Holotype: 231942. Skin and skull. Adult female. [FC: head of] 

Wallowa Lake, [SL: altitude 4,000 ft (1,219 m)], Wallowa 

County, Oregon. Collected 13 Apr 1919 by G. G. Cantwell. 

Original number 1093.

Citellus canus vigilis Merriam

Proc. Biol. Soc. Washington 26: 137, 21 May 1913.

= Urocitellus canus (Merriam, 1898). See Helgen et al. (2009: 

297).

Holotype: 168361. [Skin and skull]. Adult female. Vale, [Malheur County], eastern Oregon. Collected 29 Apr 1910 by 

S. G. Jewett. Original number 30.

Citellus elegans nevadensis A. H. Howell

Proc. Biol. Soc. Washington 41: 211, 18 Dec 1928.

= Urocitellus elegans nevadensis (A. H. Howell, 1928). See Helgen et al. (2009:297).

Holotype: 156788. Skin and skull. Adult female. Paradise, Humboldt County, Nevada. Collected 3 Mar 1908 by S. E. Piper. 

Original number 112.

Citellus idahoensis Merriam

Proc. Biol. Soc. Washington 26: 135, 21 May 1913.

= Urocitellus mollis idahoensis (Merriam, 1913). See Helgen et 

al. (2009:297).

Holotype: 168290. [Skin and skull]. Adult female. Payette, at 

junction of Payette and Snake River, [Payette County], 

Idaho. Collected 23 Apr 1910 by S. G. Jewett. Original 

number 17.

Citellus leurodon Merriam

Proc. Biol. Soc. Washington 26: 136, 21 May 1913.

= Urocitellus mollis mollis (Kennicott, 1863). See Helgen et al. 

(2009:297).

Holotype: 169031. [Skin and skull]. Young adult male. Murphy, 

[Owyhee County], in hills of southwestern Idaho west of 

Snake River. Collected 30 May 1910 by S. G. Jewett. Original number 112.

Citellus mollis artemesiae Merriam

Proc. Biol. Soc. Washington 26: 137, 21 May 1913.

= Urocitellus mollis artemesiae (Merriam, 1913). See Helgen et 

al. (2009:297).

Holotype: 23489/30907. Skin and skull. Adult male. Birch 

Creek, [A. H. Howell (1938:65): about 10 mi (16 km) south 

of Nicholia], [Clark County], Idaho. Collected 9 Aug 1890 

by V. Bailey. Original number 1573.

Citellus mollus [sic] pessimus Merriam

Proc. Biol. Soc. Washington 26: 138, 21 May 1913.

= Urocitellus mollis artemesiae (Merriam, 1913). See Helgen et 

al. (2009:297).

Holotype: 23925/31330. Skin and skull. Adult male. Lower part 

of Big Lost River, [Butte County], east central Idaho. Collected 23 Jul 1890 by C. P. Streator. Original number 53.

Citellus mollis washoensis Merriam

Proc. Biol. Soc. Washington 26: 138, 21 May 1913.

= Urocitellus mollis mollis (Kennicott, 1863). See Helgen et al. 

(2009:277).

Holotype: 16946/23861. Skin and skull. Adult [FC: male]. Carson Valley [FC: Carson City], [Douglas County], western 

Nevada. Collected 23 May 1889 by C. A. Keeler. Original 

number 34.

Citellus nebulicola Osgood

Proc. Biol. Soc. Washington 16: 26, 19 Mar 1903.

= Urocitellus parryii nebulicola (Osgood, 1903). See Helgen et 

al. (2009:297).

Holotype: 59145. [Skin and skull]. Adult female. Nagai Island, 

Shumagin Islands, Alaska. Collected 24 Jun 1893 by C. H. 

Townsend. No original number.
3 4 • smithsonian contributions to zoology

Citellus plesius ablusus Osgood

Proc. Biol. Soc. Washington 16: 25, 19 Mar 1903.

= Urocitellus parryii ablusus (Osgood, 1903). See Helgen et al. 

(2009:297).

Holotype: 119815. [Skin and skull]. Adult male. Nushagak, 

Alaska, Collected 16 Sep 1902 by W. H. Osgood and A. G. 

Maddren. Original number [FC: WHO] 2043.

Citellus stejnegeri J. A. Allen

Bull. Amer. Mus. Nat. Hist. 19: 142, 31 Mar 1903.

= Urocitellus parryii stejnegeri (J. A. Allen, 1903). See Helgen et 

al. (2009:297).

Holotype: 63226. [Skin and skull]. Juvenile male? Near Petropaulski, southeastern Kamchatka, [Russia]. Collected [MC: 

1895] by L. Stejneger. No original number.

Citellus townsendii brunneus A. H. Howell

Proc. Biol. Soc. Washington 41: 211, 18 Dec 1928.

= Urocitellus brunneus brunneus (A. H. Howell, 1928). See Helgen et al. (2009:297).

Holotype: 201963. Skin and skull. Adult female. New Meadows, Adams County, Idaho. Collected 11 Jul 1913 by L. E. 

Wyman. Original number 178.

Citellus washingtoni washingtoni A. H. Howell

N. Amer. Fauna 56: 69, 18 May 1938.

= Urocitellus washingtoni (A. H. Howell, 1938). See Helgen et 

al: (2009:298).

Holotype: 27948/40068. Skin and skull. Adult male. Touchet 

[FC: sage plain at mouth of Touchet River], Walla Walla 

County, Washington. Collected 18 May 1891 by C. P. 

Streator. Original number 817.

Citellus washingtoni loringi A. H. Howell

N. Amer. Fauna 56: 71, 18 May 1938.

= Urocitellus washingtoni (A. H. Howell, 1938). See Helgen et 

al. (2009:298).

Holotype: 89805. Skin and skull. Adult male. Douglas, [Douglas 

County], Washington. Collected 1 Aug 1897 by J. A. Loring. Original number 4547.

Spermophilus armatus Kennicott

Proc. Acad. Nat. Sci. Philadelphia 15: 158, ordered published 

30 Jun 1863, receipt acknowledged 28 Nov 1863 by the 

Smithsonian Institution.

= Urocitellus armatus (Kennicott, 1863). See Helgen et al. (2009: 

297).

Syntypes: No numbers are mentioned in the original description. Kennicott briefly describes four new spermophiles in 

the collection of the Smithsonian Institution collected by 

C. Drexler at Fort Bridger, [Uinta County], Utah (Wyoming). Subsequently, Lyon and Osgood (1909:163) considered all specimens from Fort Bridger taken by Drexler 

and in the National Collection before 1863 to be Cotypes 

(Syntypes) of the species. These specimens, listed below, all 

have armatus written on their labels, possibly by Kennicott, 

and they are entered in the museum catalog as Spermophilus 

armatus. The skins were all collected in April through June 

1858 and the specimens in alcohol were probably taken at 

the same time.

3463/4890. Skin and skull. Female. Collected 26 May 1858. 

Original number 454. Skin cannot be found.

3464. Skin only. Female. Collected 25 Jun 1858. Original number 728. [SL: Camp Scott].

3466/4808. Skin and skull. Male. Collected 10 Jun 1858. Original number 610. Skin cannot be found.

3467. Skin only. Male. Collected 2 May 1858. Original number 

261. Cannot be found.

3470. Skin only. Female. Collected 26 May 1858. Original number 455. Cannot be found.

3472/4799. Skin and skull. Male. Collected 11 Apr 1858. Original number 167. Cannot be found.

3474. Skin only. Male. Collected 14 Apr 1858. Original number 

215. Cannot be found.

3475. Skin only. Female. Collected 14 Apr 1858. Original number 197. Cannot be found.

3476. Skin only with partial skull inside. Female. Collected 15 

Apr 1858. Original number 229. [SL: Camp Scott].

3478. Skin only. Male. Collected 2 Apr 1858. Original number 

140. Cannot be found.

3481. Skin only. Male. Collected 19 May 1858. Original number 

375. Cannot be found.

4221/4794. Skin and skull. No information associated with 

specimen. Remarks in the catalog state “Died at S. I. after 

two years’ confinement.” It was probably collected at the 

time the others were taken, Lyon and Osgood (1909:164). 

Skin cannot be found.

5958. Alcoholic. Cannot be found.

5959. Alcoholic.

5960. Alcoholic.

5961. Skin only preserved in alcohol. Remainder cannot be found.

No additional data is given for the alcohol-preserved specimens. 

Lyon and Osgood (1909:164) indicate they were cataloged 

5 Feb 1863, but according to the catalog it is possible that 

they were cataloged in 1862.

Spermophilus barrowensis Merriam

Proc. Washington Acad. Sci. 2: 19, 14 Mar 1900b.

= Urocitellus parryii kennicottii (Ross, 1861). See Helgen et al. 

(2009:297).

Holotype: 14061/37824. Skin and skull. Adult male. Point Barrow, Alaska. [SL: Collected 30 May 1883 by P. L. Ray. Original number 1428].

Spermophilus beldingi Merriam

Ann. New York Acad. Sci. 4: 317, 28 Dec 1888.

= Urocitellus beldingi beldingi (Merriam, 1888). See Helgen et 

al. (2009:297).
number 640 • 3 5

Holotype: [186467]. Skin and skull. Adult female. Donner, 

Placer County, California. Collected [SL: 22 Jun 1886 by 

C. A. Allen. Original number 103]. Merriam collection 

number 2712/3342.

Spermophilus beringensis Merriam

Proc. Washington Acad. Sci. 2: 20, 14 Mar 1900b.

= Urocitellus parryii kennicottii (Ross, 1861). See Helgen et al. 

(2009:297).

Holotype: 15253. Skin only. Adult male. Cape Lisbourne, Coal 

Veins, Alaska. Collected May 1885 by H. D. Woolfe. No 

original number.

Spermophilus brunneus endemicus Yensen

J. Mamm. 72: 596, 16 Aug 1991.

= Urocitellus brunneus endemicus (Yensen, 1991). See Helgen et 

al. (2009:297).

Holotype: 565927. Skin and skull. Female. Sand Hollow, 1 km 

east, up canyon from OX Ranch headquarters, T9N, R4W, 

Sec 7, NE 1⁄4; 5.6 km north, 5 km east Payette; elevation 

750 m, Payette County, Idaho. Collected 30 Mar 1989 by 

E. Yensen. Original number 932.

Spermophilus elegans Kennicott

Proc. Acad. Nat. Sci. Philadelphia 15: 158, ordered published 

30 Jun 1863, receipt acknowledged 28 Nov 1863 by the 

Smithsonian Institution.

= Urocitellus elegans elegans (Kennicott, 1863). See Helgen et 

al. (2009:297).

Syntypes: No numbers are mentioned in the original description. Kennicott briefly describes four new spermophiles in 

the collection of the Smithsonian Institution collected by 

C. Drexler at Fort Bridger, [Uinta County], Utah (Wyoming). Subsequently, Lyon and Osgood (1909:165) considered all specimens from Fort Bridger taken by Drexler and 

in the National Collection before 1863 to be cotypes (syntypes) of the species. These specimens, listed below, all have 

elegans written on their labels, possibly by Kennicott, and 

they are entered in the museum catalog as Spermophilus elegans. The skins were collected in April and May 1858, and 

the specimens in alcohol were probably taken at the same 

time. Specimens without a date of collection were cataloged 

in late 1862 or early 1863.

3468/4796. Skin and skull. Female. Collected 11 Apr 1858. 

Original number 168. The skin and mandible cannot be 

found.

3469. Skin only. Female. Collected 11 Apr 1858. Original number 169. [SL: Camp Scott].

3473. Skin only. Male. Collected 11 Apr 1858. Original number 

166. Cannot be found.

3479/4810. Skin and skull. Female. Collected 17 Apr 1858. 

Original number 233. [SL: Camp Scott].

3480. Skin with partial skull inside. Male. Collected 14 Apr 

1858. Original number 216.

4003. Skin only with fragmentary skull inside, formerly mounted. 

Female. Cataloged Jun 1860. Original number 3.

5950/4814. Skin and skull. Skin, without body, in alcohol.

5951. Alcoholic, abdominal viscera removed.

5952. Alcoholic, abdominal viscera removed.

5953. Alcoholic, abdominal viscera removed.

5954. Alcoholic, abdominal viscera removed. Young.

5955.Mounted skin with body in alcohol.

5956. Alcoholic, abdominal viscera removed.

5957. Body without feet or head in alcohol. Skin cannot be found.

Spermophilus empetra plesius Osgood

N. Amer. Fauna 19: 29, 6 Oct 1900a.

= Urocitellus parryii plesius (Osgood, 1900). See Helgen et al. 

(2009:297).

Holotype: 98931. Skin and skull. Adult female. Bennett City, 

head of Lake Bennett, British Columbia, [Canada]. Collected 19 Jun 1899 by W. H. Osgood. Original number 465.

Spermophilus mollis Kennicott

Proc. Acad. Nat. Sci. Philadelphia 15: 157, ordered published 

30 Jun 1863, receipt acknowledged 28 Nov 1863 by the 

Smithsonian Institution.

= Urocitellus mollis mollis (Kennicott, 1863). See Helgen et al. 

(2009:297).

Lectotype: 3777/4798. Skin and skull. Age and sex undetermined. Camp Floyd, near Fairfield, Wasatch (Utah) County, 

Utah. Collected 18 Mar 1859 by C. S. McCarthy. Original 

number 164.

No type designated in the original description. Specimens upon 

which the description was based came from Camp Floyd and 

Rocky Mountains and were collected by C. Drexler and C. S. 

McCarthy. Miller and Rehn (1901:53) restricted the type locality to Camp Floyd. Subsequently, A. H. Howell (1938:63) 

selected the specimen given above as the Lectotype.

Spermophilus mollis canus Merriam

Proc. Biol. Soc. Washington 12: 70, 24 Mar 1898b.

= Urocitellus canus (Merriam, 1898). See Helgen et al. (2009:297).

Holotype: 78681. [Skin and skull]. Adult female. [FC: 7 mi (11 

km) east] Antelope, Wasco County, Oregon. Collected 21 

Jun 1896 by V. Bailey. Original number 5561.

Spermophilus mollis stephensi Merriam

Proc. Biol. Soc. Washington 12: 69, 24 Mar 1898b.

= Urocitellus mollis mollis (Kennicott, 1863). See Helgen et al. 

(2009:297).

Holotype: 29492/41567. Skin and skull. Adult male. Queen Station, near head of Owens Valley, altitude about 6,000 ft 

(1,829 m), [Esmeralda County], Nevada. Collected 12 Jul 

1891 by F. Stephens. Original number 718.

Spermophilus mollis yakimensis Merriam

Proc. Biol. Soc. Washington 12: 70, 24 Mar 1898b.
3 6 • smithsonian contributions to zoology

= Urocitellus townsendii townsendii (Bachman, 1839). See Helgen et al. (2009:298).

Holotype: 89331. [Skin and skull]. Adult male. Mabton, Yakima 

County, Washington. Collected 16 Jul 1897 by W. K. Fisher. 

Original number 323.

Spermophilus oregonus Merriam

Proc. Biol. Soc. Washington 12: 69, 24 Mar 1898b.

= Urocitellus beldingi oregonus (Merriam, 1898). See Helgen et 

al. (2009:297).

Holotype: 89177. [Skin and skull]. Adult female. Swan Lake Valley, Klamath Basin, [Klamath County], Oregon. Collected 

12 Jun 1897 by V. Bailey. Original number 6005.

Spermophilus osgoodi Merriam

Proc. Washington Acad. Sci. 2: 18, 14 Mar 1900b.

= Urocitellus parryii osgoodi (Merriam, 1900). See Helgen et al. 

(2009:297).

Holotype: 12789/37822. Skin and skull. Adult male. Fort 

Yukon, Alaska. Collected 29 Apr 1877 by L. M. Turner. 

Original number 1635.

Spermophilus parryii kodiacensis J. A. Allen

Proc. Boston Soc. Nat. Hist. 16: 292, 1874.

= Urocitellus parryii kodiacensis (J. A. Allen, 1874). See Helgen 

et al. (2009:297).

Lectotype: 9242/38543. Skin and skull. Adult female. Kodiak Island, Alaska. Collected Jun 1868 by F. Bischoff. No original 

number. No type designated in original description. Lectotype designated by A. H. Howell (1938:103).

Genus Xerospermophilus Merriam, 1892

Citellus spilosoma pallescens A. H. Howell

Proc. Biol. Soc. Washington 41: 212, 18 Dec 1928.

= Xerospermophilus spilosoma pallescens (A. H. Howell, 1928). 

See Helgen et al. (2009:295).

Holotype: 79535. Skin and skull. Adult male. La Ventura, [FC: 

altitude 5,650 ft (1,722 m)], Coahuila, Mexico. Collected 

10 Aug 1896 by E. W. Nelson and E. A. Goldman. Original 

number 10016.

Spermophilus canescens Merriam

N. Amer. Fauna 4: 38, 8 Oct 1890d.

= Xerospermophilus spilosoma canescens (Merriam, 1890). See 

Helgen et al. (2009:295).

Holotype: 17873/24810. Skin and skull. Immature male. Willcox, [FC: on alkalie flats], Cochise County, Arizona. Collected 16 Nov 1889 by V. Bailey. Original number 676.

Spermophilus cryptospilotus Merriam

N. Amer. Fauna 3: 57, 11 Sep 1890b.

= Xerospermophilus spilosoma cryptospilotus (Merriam, 1890). 

See Helgen et al. (2009:295).

Holotype: 17676/24612. Skin and skull. Adult male. [FC: Near] 

Tenebito (Dinnebito) Wash, [Merriam (1890b:57): about 40 

km (25 mi) east of the Little Colorado], Painted Desert, [Coconino County], Arizona. Collected 17 Aug 1889 by C. H. 

Merriam. Original number [FC: V. Bailey] 374.

Spermophilus mohavensis Merriam

N. Amer. Fauna 2: 15, 30 Oct 1889c.

= Xerospermophilus mohavensis (Merriam, 1889). See Helgen 

et al. (2009:294).

Holotype: [186469]. Skin and skull. Adult male. Mohave River, 

California. Type locality restricted by Grinnell and Dickson 

(1919:667) to Near Rabbit Springs, about 15 mi (24 km) east 

of Mohave River at Hesperia, San Bernardino County, California. Collected 29 Jun 1886 by F. Stephens. Original number [MCC: 315]. Merriam Collection number 2594/3223.

Spermophilus neglectus Merriam

N. Amer. Fauna 2: 17, 30 Oct 1889d.

= Xerospermophilus tereticaudus neglectus (Merriam, 1889). 

See Helgen et al. (2009:295).

Holotype: [186470]. Skin and skull. [SL: Adult] male. The valley 

1 mi (1.6 km) west of Dolan’s Spring, altitude about 3,000 

ft (914 m), Arizona. A. H. Howell (1938:187) interpreted 

this to be 12 mi (19 km) northwest of Chloride, Mohave 

County. Collected 9 Feb 1889 by V. Bailey. Original number 

[SL: 566]. Merriam collection number 5262/5952.

Spermophilus obsoletus Kennicott

Proc. Acad. Nat. Sci. Philadelphia 15: 157, ordered published 

30 Jun 1863, receipt acknowledged 28 Nov 1863 by the 

Smithsonian Institution.

= Xerospermophilus spilosoma obsoletus (Kennicott, 1863). See 

Helgen et al. (2009:295).

Lectotype: 3222/37998. Skin and skull. Female. Fifty miles (80 

km) west of Fort Kearney, Nebraska. Collected 9 Aug 1857 

by J. G. Cooper. Original number 44. Lectotype designated 

by A. H. Howell (1938:131).

Spermophilus perotensis Merriam

Proc. Biol. Soc. Washington 8: 131, 28 Dec 1893a.

= Xerospermophilus perotensis (Merriam, 1893). See Helgen et 

al. (2009:294).

Holotype: 54274. [Skin and skull]. Adult female. Perote, [FC: 

7,800 ft (2,377 m)], Veracruz, Mexico. Collected 8 Jun 

1893 by E. W. Nelson. Original number 4976.

Spermophilus spilosoma annectens Merriam

Proc. Biol. Soc. Washington 8: 132, 28 Dec 1893a.

= Xerospermophilus spilosoma annectens (Merriam, 1893). See 

Helgen et al. (2009:294).

Holotype: 30410/42396. Skin and skull. Young adult male. [FC 

and amended by A. H. Howell (1938:128): The Tanks, 

12 mi (19 km) from Point (Port) Isabel], Padre Island, [Cam-
number 640 • 3 7

eron County], Texas. Collected 24 Aug 1891 by W. Lloyd. 

Original number 694.

Spermophilus spilosoma arens V. Bailey

Proc. Biol. Soc. Washington 15: 118, 2 Jun 1902.

= Xerospermophilus spilosoma canescens (Merriam, 1890). See 

Helgen et al. (2009:295).

Holotype: 64977. Skin and skull. Adult male. El Paso, [El Paso 

County], Texas. Collected 10 May 1894 by A. K. Fisher. 

Original number 1446.

Spermophilus spilosoma major Merriam

N. Amer. Fauna 4: 39, 8 Oct 1890d.

= Xerospermophilus spilosoma major (Merriam, 1890). New 

name combination

Holotype: 17116/24049. Skin and skull. Adult female. Albuquerque, [Bernalillo County], New Mexico. Collected 22 

Jul 1889 by V. Bailey. Original number 225.

Spermophilus spilosoma macrospilotus Merriam

N. Amer. Fauna 4: 38, 8 Oct 1890d.

= Xerospermophilus spilosoma canescens (Merriam, 1890). See 

Helgen et al. (2009:295).

Holotype: 16750/23663. Skin and skull. Adult female. Oracle, 

Pinal County, Arizona. Collected 11 Jun 1889 by V. Bailey. 

Original number 129.

Spermophilus spilosoma marginatus V. Bailey

Proc. Biol. Soc. Washington 15: 118, 2 Jun 1902.

= Xerospermophilus spilosoma major (Merriam, 1890). New 

name combination.

Holotype: 108927. [Skin and skull]. Adult male. [FC: 4 mi 

(6 km) E] Alpine, [Brewster County], Texas. Collected 5 Jul 

1901 by V. Bailey. Original number 7702.

Spermophilus spilosoma obsidianus Merriam

N. Amer. Fauna 3: 56, 11 Sep 1890b.

= Xerospermophilus spilosoma pratensis (Merriam, 1890). See 

Helgen et al. (2009:295).

Holotype: 17674/24610. Skin and skull. Adult male. Cedar belt 

northeast of San Francisco Mountain (Peaks), [FC: below 

Partridge Spring, altitude 7,250 ft (2,210 m)], [Coconino 

County, Arizona]. Collected 1 Oct 1889 by V. Bailey. Original number 557.

Spermophilus spilosoma pratensis Merriam

N. Amer. Fauna 3: 55, 11 Sep 1890b.

= Xerospermophilus spilosoma pratensis (Merriam, 1890). See 

Helgen et al. (2009:295).

Holotype: 17659/24595. Skin and skull. Adult female. [FC: 

Little Spring], Pine plateau at north foot of San Francisco 

Mountain (Peaks), [Coconino County], Arizona. Collected 

5 Aug 1889 by C. H. Merriam and V. Bailey. Original number [FC:] VB 285.

Spermophilus tereticaudus Baird

Mammals in Rep. Expl. Surv. 8(1): 315, 1857.

= Xerospermophilus tereticaudus tereticaudus (Baird, 1857). See 

Helgen et al. (2009:295).

Syntypes: No type designated in the original description, which 

is apparently based equally upon these specimens, which are 

listed by number.

1584/2419. Skin and skull. Not fully adult male. Original number [SL: 236].

1585. Skin of the head and neck. The body is supposed to be in 

alcohol but cannot be found. [SL: Female].

2490. Alcoholic. Young female.

All specimens were collected at [Old] Fort Yuma, [Imperial 

County], California by G. H. Thomas. Skins cataloged 24 

Jun 1856, the alcoholic 9 Apr 1857. Poole and Schantz 

(1942:519) add to the locality “on right bank of the Colorado River, opposite present town of Yuma, Arizona.” 

Baird (1857:709) in his list of localities says of Fort Yuma: 

“at mouth of Gila River. Altitude 355 ft (108 m). Latitude 

32°32′, Longitude 114°36′.”

Family Gliridae Muirhead, 1819

Genus Eliomys Wagner, 1840

Eliomys quercinus denticulatus Ranck

Bull. U.S. Natl. Mus. 275: 216, 2 Oct 1968.

= Eliomys munbyanus (Pomel, 1856). See Holden (2005:834).

Holotype: 322757. Skin, skull, [partial skeleton]. Adult female. 

El Gatrun, Fezzan Province, [Murzuq], Libya. Collected 11 

Jan 1962 by G. L. Ranck. Original number 1289.

Eliomys cincticauda Miller

Proc. Biol. Soc. Washington 14: 39, 25 Apr 1901c.

= Eliomys quercinus (Linnaeus, 1766). See Holden (2005:835).

Holotype: 103030. Skin and skull. Adult male. Sorrento, near 

Naples, [Napoli Province, Campania], Italy. Collected 31 

May 1900 by D. Coolidge. Original number 1118.

Genus Graphiurus Smuts, 1832

Eliomys parvus True

Proc. U.S. Natl. Mus. 16: 601, 25 Oct 1893.

= Graphiurus kelleni (Reuvens, 1890). See Holden (2005:824).

Holotype: 21005/[36056]. Skin and skull. Female. Along the 

Tana River, between the coast and Hameye, [Coast Province], East Africa (Kenya). Collected [MC: Nov 1892] by 

W. A. Chanler and L. von Hohnel. No original number.

Graphiurus murinus isolatus Heller

Smithsonian Misc. Coll. 59: 3, 5 Jul 1912.

= Graphiurus murinus (Desmarest, 1822). See Holden (2005:826).

Holotype: 181788. Skin, skull cannot be found. Adult female. 

[SL: summit] Mount Umengo, altitude 5,000 ft (1,524 m), 
3 8 • smithsonian contributions to zoology

Taita Mountains, [Coast Province], British East Africa 

(Kenya). Collected [SL: 11] Nov 1911 by E. Heller. Original 

number 4719.

Graphiurus murinus johnstoni Heller

Smithsonian Misc. Coll. 59 (16): 2, 5 Jul 1912.

= Graphiurus murinus (Desmarest, 1822). See Holden (2005:826).

Holotype: 181787. Skin and skull. Adult male. [SL: summit] 

Mount Gargues (Uaragess), Mathews Range, altitude 7,000 

ft (2,134 m) [SL: 7,100 ft (2,164 m)], [Rift Valley], British 

East Africa (Kenya). Collected 26 Aug 1911 by E. Heller. 

Original number 4114.

Graphiurus personatus Heller

Smithsonian Misc. Coll. 56 (17): 2, 28 Feb 1911.

= Graphiurus kelleni (Reuvens, 1890). See Holden (2005:824).

Holotype: 164827. [Skin and skull]. Adult female. Rhino Camp, 

Lado Enclave, [Uganda]. Collected 11 Jan 1910 by J. A. 

Loring. Original number 8644.

Genus Myomimus Ognev, 1924

Myomimus setzeri Rossolimo

Vestn. Zool. 4: 51, 1976.

Holotype: 369888. Skin and skull. Adult male. Four kilometers 

West Branch, Kordistan Province, Iran. Collected 30 May 

1965, not 9 Jun 1964 as in original description, by R. G. 

Tuck. Original number 1600.

Suborder Castorimorpha Wood, 1955

Family Castoridae Hemprich, 1820

Genus Castor Linnaeus, 1758

Subspecies of Castor canadensis are not recognized by Helgen 

(2005:842), but those listed below are recognized by Hall 

(1981:601–605) and Baker and Hill (2003:288).

Castor canadensis acadicus V. Bailey and Doutt

J. Mamm. 23: 87, 14 Feb 1942.

Holotype: 174525. [Skin and skull]. Adult male. Nepisiquit 

River, New Brunswick, Canada. Collected 25 Sep 1911 by 

K. Roosevelt. No original number.

Castor canadensis baileyi Nelson

Proc. Biol. Soc. Washington 40: 125, 26 Sep 1927.

Holotype: 228006. [Skin and skull]. Adult male. Humboldt 

River, 4 mi (6.4 km) above Winnemucca, [Humboldt 

County], Nevada. Collected 13 Oct 1917 by J. R. Bunch. 

Original number [SL: 5]. [SL: 19885-X].

Castor canadensis frontador Mearns

Proc. U.S. Natl. Mus. 20: 2, 5 Mar 1897a; Preprint of Proc. U.S. 

Natl. Mus. 20: 502, 19 Jan 1898b.

Holotype: 20750/35883. Skin and skull. Adult male. Near monument 98 of the Mexican Boundary line, San Pedro River, 

Sonora, Mexico. Collected 24 Oct 1892 by E. A. Mearns 

and F. X Holzner. Original number 2151.

Castor canadensis mexicanus V. Bailey

Proc. Biol. Soc. Washington 26: 191, 23 Oct 1913b.

Holotype: 96522. [Skin and skull]. Adult female. [SL: Blackwell’s 

Ranch], Ruidoso Creek, 6 mi (9.6 km) below Ruidoso, [Lincoln County], New Mexico. Collected 29 Sep 1898, not 

1899 as in original description, by C. Barber. Original number 83. 1991-X.

Castor canadensis michiganensis V. Bailey

Proc. Biol. Soc. Washington 26: 192, 23 Oct 1913b.

Holotype: 170561. [Skin and skull]. Adult female. Tahquamenaw 

(Tahquamenon) River, 5 mi (8 km) above [Tahquamenon] 

falls, Luce County, Michigan. Collected 20 Sep [SL: 1910] 

by C. Birdseye. Original number 1270.

Castor canadensis missouriensis V. Bailey

J. Mamm. 1: 32, 28 Nov 1919.

Holotype: 205763. Skin and skull. Immature male. Apple Creek, 

7 mi (11.3 km) east of Bismarck, [Burleigh County], North 

Dakota. Collected 1 Sep 1914 by V. Bailey. Original number 

9703.

Castor canadensis repentinus Goldman

J. Mamm. 13: 266, 9 Aug 1932b.

Holotype: 161613. [Skin and skull]. Adult female. Bright Angel 

Creek, altitude 4,000 ft (1,219 m), Grand Canyon of the 

Colorado River, [Coconino County], Arizona. Collected 14 

Sep 1909 by C. Birdseye. Original number 723.

Castor canadensis texensis V. Bailey

N. Amer. Fauna 25: 122, 24 Oct 1905.

Holotype: 135744. [Skin and skull]. Old, sex unknown. Cummings Creek, [XC: 9 mi (14 km) from New Ulm], Colorado 

County, Texas. Collected 25 Dec 1900 by F. Brune. Kept in 

captivity until 10 Jan 1901. No original number. 5139-X.

Castor subauratus shastensis Taylor

Univ. California Publ. Zool. 12: 433, 20 Mar 1916.

= Castor canadensis shastensis Taylor, 1916. See Grinnell 

(1933:166).

Holotype: 50978. Skull only. Adult male. Cassel (Hat Creek), Pit 

River, Shasta County, California. Collected 3 Jan 1893 by 

H. E. Williams. No original number.

Family Heteromyidae Gray, 1868

Genus Chaetodipus Merriam, 1889

Chaetodipus intermedius beardi Weckerly, Gennaro, and Best

Southwestern Nat. 33: 100, 30 Mar 1988.
number 640 • 3 9

Holotype: [564498]. Skin and cranium (skull). Adult female. 

Twenty-six miles (42 km) north, 15.5 mi (25 km) east Engle, 

Socorro County, New Mexico. Collected 23 Aug 1982 by 

F. W. Weckerly. Original number 236. Eastern New Mexico University (ENMU) Natural History Museum number 

8918.

Perognathus anthonyi Osgood

N. Amer. Fauna 18: 56, 20 Sep 1900b.

= Chaetodipus fallax anthonyi (Osgood, 1900). See Williams et 

al. (1993:137).

Holotype: 81058. [Skin and skull]. Adult female. South Bay, 

Cerros (Cedros) Island, Lower California (Baja California), 

Mexico. Collected 29 Jul 1896 by A. W. Anthony. Original 

number 71.

Perognathus arenarius ambiguus Nelson and Goldman

Proc. Biol. Soc. Washington 42: 108, 25 Mar 1929b.

= Chaetodipus arenarius ambiguus (Nelson and Goldman, 

1929). See Lackey (1991a:1).

Holotype: 140011. [Skin and skull]. Young adult male, molars 

slightly worn. Yubay, altitude 2,000 ft (610 m), 30 mi (48 

km) southeast of Calamahue, Lower California (Baja California), Mexico. Collected 18 Sep 1905 by E. W. Nelson and 

E. A. Goldman. Original number 18141.

Perognathus arenarius sublucidus Nelson and Goldman

Proc. Biol. Soc. Washington 42: 109, 25 Mar 1929b.

= Chaetodipus arenarius sublucidus (Nelson and Goldman, 

1929). See Lackey (1991a:1).

Holotype: 146896. [Skin and skull]. Adult male. La Paz, Lower California (Baja California Sur), Mexico. Collected 16 Feb 1906 

by E. W. Nelson and E. A. Goldman. Original number 19146.

Perognathus armatus Merriam

N. Amer. Fauna 1: 27, 25 Oct 1889e.

= Chaetodipus californicus californicus (Merriam, 1889) See 

Williams et al. (1993:135).

Holotype: [186505]. Skin and skull. Adult female. Mount Diablo, [Contra Costa County], California. Collected 28 Mar 

1882 by W. E. Bryant. No original number. [Merriam collection number] 672/6321.

Perognathus artus Osgood

N. Amer. Fauna 18: 55, 20 Sep 1900b.

= Chaetodipus artus (Osgood, 1900). See Hafner and Hafner 

(1983:25).

Holotype: 96298. [Skin and skull]. Adult female. Batopilas, [FC: 

about 2,500 ft (762 m)], Chihuahua, Mexico. Collected 6 

Oct 1898 by E. A. Goldman. Original number [FC: Nelson 

and Goldman] 13090.

Perognathus baileyi Merriam

Proc. Acad. Nat. Sci. Philadelphia 46: 262, 27 Sep 1894a. Preprint 

of Proc. Acad. Nat. Sci. Philadelphia 46: 262, 23 Oct 1894a.

= Chaetodipus baileyi baileyi (Merriam, 1894). See Paulson 

(1988a:1).

Holotype: 17838/24775. Skin and skull. Adult female. Magdalena, Sonora, Mexico. Collected 3 Nov 1889 by V. Bailey. 

Original number 633.

Perognathus baileyi domensis Goldman

Proc. Biol. Soc. Washington 41: 204, 18 Dec 1928a.

= Chaetodipus baileyi baileyi (Merriam, 1894). See Paulson 

(1988a:1).

Holotype: 248002. Skin and skull. Adult female. Castle Dome, 

at base of Castle Dome Peak, altitude 1,400 ft (427 m), [FC: 

Yuma County], Arizona. Collected 13 Apr 1927 by G. G. 

Cantwell. Original number 2882.

Perognathus baileyi extimus Nelson and Goldman

J. Washington Acad. Sci. 20: 223, 19 Jun 1930a.

= Chaetodipus rudinoris extimus (Nelson and Goldman, 1930). 

See Riddle et al. (2000:167).

Holotype: 146672. Skin and skull. Adult female. Tres Pachitas, 

altitude 700 ft (213 m), [FC about] 36 mi (58 km) south 

of La Paz, Lower California (Baja California Sur), Mexico. 

Collected 25 Dec 1905 by [E. W.] Nelson and [E. A.] Goldman. Original number 18785.

Perognathus baileyi insularis Townsend

Bull. Amer. Mus. Nat. Hist. 31: 122, 14 Jun 1912.

= Chaetodipus baileyi insularis (Townsend, 1912). See Paulson 

(1988a:1).

Holotype: [198410. Skin and skull]. Male. Tiburon Island, [Gulf of 

California, Sonora, Mexico]. Collected 13 Apr 1911 by [SL: 

H. E. Anthony. Original number 153]. Specimen described as 

American Museum of Natural History number 31846.

Perognathus californicus Merriam

N. Amer. Fauna 1: 26, 25 Oct 1889e.

= Chaetodipus californicus californicus (Merriam, 1889). See 

Williams et al. (1993:135).

Holotype: [186506]. Skin and skull. [MCC: Adult] female. 

Berkeley, [Alameda County], California. Collected 8 Nov 

1888 by T. S. Palmer and C. A. Keeler. No original number. 

[Merriam collection number] 5132/5827.

Perognathus californicus dispar Osgood

N. Amer. Fauna 18: 58, 20 Sep 1900b.

= Chaetodipus californicus dispar (Osgood, 1900). See Williams 

et al. (1993:135).

Holotype: 32116/43928. Skin and skull. Adult male. Carpenteria, 

[FC: 250 ft (76 m)], Santa Barbara County, California. Collected 19 Dec 1891 by E. W. Nelson. Original number 1655.

Perognathus californicus ochrus Osgood

Proc. Biol. Soc. Washington 17: 128, 9 Jun 1904.

= Chaetodipus californicus ochrus (Osgood, 1904). See Williams 

et al. (1993:136).
4 0 • smithsonian contributions to zoology

Holotype: 130348. [Skin and skull]. Young adult female. Santiago Springs, 16 mi (26 km) southwest of McKittrick, Kern 

County, California. Collected 30 Jul 1903 by L. J. Goldman. 

Original number [SL: 728].

Perognathus dalquesti Roth

J. Mamm. 57: 562, 27 Aug 1976.

= Chaetodipus dalquesti (Roth, 1976). See Hafner and Hafner 

(1983:25).

Holotype: 510851. Skin, skull, [and baculum]. Adult male. Four 

miles (6.4 km) southeast Migrino (32 km south southeast 

Todos Santos), elevation near sea level, Baja California Sur, 

Mexico, 23°10′N, 110°07′W. Collected 7 Jan 1973 by E. L. 

Roth. Original number 1974. University of Arizona number 

UA 20108.

Perognathus (Chaetodipus) eremicus Mearns

Bull. Amer. Mus. Nat. Hist. 10: 300, 31 Aug 1898c.

= Chaetodipus eremicus eremicus (Mearns, 1898). See Patton 

(2005a:854).

Holotype: 21052/36094. Skin and skull. Adult female. Fort 

Hancock, El Paso (Hudspeth) County, Texas. Collected 27 

Jun 1893 by E. A. Mearns. Original number 2380.

Perognathus fallax Merriam

N. Amer. Fauna 1: 19, 25 Oct 1889e.

= Chaetodipus fallax fallax (Merriam, 1889). See Williams et al. 

(1993:137).

Holotype: 15889/22684. Skin and skull. Adult male. San Bernardino, California. Type locality restricted to “Reche 

Canyon, 3 mi (4.8 km) southeast of Colton, San Bernardino County, California” by Osgood (1900b:55). Grinnell (1933:153) gives 1,250 ft (381 m) as the altitude. 

Collected 21 Apr 1887 by F. Stephens. Original number 

[SL: 424].

Perognathus fallax inopinus Nelson and Goldman

Proc. Biol. Soc. Washington 42: 110, 25 Mar 1929b.

= Chaetodipus fallax inopinus (Nelson and Goldman, 1929). See 

Williams et al. (1993:138).

Holotype: 81059. [Skin and skull]. Adult male. Turtle (also 

known as Bartolome) Bay (Bahia Tortugas), Lower California (Baja California Sur), Mexico. Collected 1 Aug 1896 by 

A. W. Anthony. Original number 82.

Perognathus fallax pallidus Mearns

Proc. Biol. Soc. Washington 14: 135, 9 Aug 1901.

= Chaetodipus fallax pallidus (Mearns, 1901). See Williams et 

al. (1993:138).

Holotype: 61007. Skin and skull. Adult female. Mountain Spring, 

halfway up the east slope of the Coast Range Mountains on 

the Mexican Boundary Line, San Diego (Imperial) County, 

California. Collected 16 May 1894 by E. A. Mearns. Original number 3520.

Perognathus formosus Merriam

N. Amer. Fauna 1: 17, 25 Oct 1889e.

= Chaetodipus formosus formosus (Merriam, 1889). See Williams et al. (1993:140).

Holotype: [186508]. Skin and skull. Adult male. [FC: 1 mi (1.6 

km) west] St. George, [FC: 3,000 ft (914 m)], [Washington 

County], Utah. Collected 2 Jan 1889 by V. Bailey. Original 

number [SL: 469]. [Merriam collection number] 5213/5908.

Perognathus formosus domisaxensis Cockrum

J. Washington Acad. Sci. 46: 131, 7 May 1956.

= Chaetodipus formosus formosus (Merriam, 1889). See Williams et al. (1993:140).

Holotype: 249006. Skin and skull. Adult female. Houserock Valley, 15 mi (24 km) west of the Navajo Bridge, Coconino 

County, Arizona. Collected 6 Aug 1929 by V. Bailey. Original number 10758.

Perognathus formosus melanocaudus Cockrum

J. Washington Acad. Sci. 46: 132, 7 May 1956.

= Chaetodipus formosus formosus (Merriam, 1889). See Williams et al, (1993:140).

Holotype: 262918. Skin and skull. Adult female. [FC: Interior 

of lava beds], Lower end of Toroweap Valley, rim of Grand 

Canyon, Mojave County, Arizona. Collected 30 Jul 1937 by 

L. C. Goldman. Original number 341.

Perognathus goldmani Osgood

N. Amer. Fauna 18: 54, 20 Sep 1900b.

= Chaetodipus goldmani (Osgood, 1900). See Hafner and Hafner (1983:25).

Holotype: 96673. [Skin and skull]. Adult female. Sinaloa, 

Sinaloa, Mexico. Collected 15 Feb 1899 by E. A. Goldman. 

Original number [FC: Nelson and Goldman] 13428.

Perognathus hispidus Baird

Mammals in Rep. Expl. Surv. 8(1): 421, 1857.

= Chaetodipus hispidus hispidus (Baird, 1857). See Paulson 

(1988b:1).

Lectotype: 577/1696. Skin and skull. Adult female. Charco Escondido, [Tamaulipas], Mexico. Collected in 1853 by D. N. 

Couch. No original number. No type designated in original 

description. Lectotype designated by Merriam (1889e:23–

24). Osgood (1900b:43) states that the skull of this specimen is composite.

Perognathus hispidus zacatecae Osgood

N. Amer. Fauna 18: 45, 20 Sep 1900b.

= Chaetodipus hispidus zacatecae (Osgood, 1900). See Paulson 

(1988b:1).

Holotype: 91877. [Skin and skull]. Young adult female. Valparaiso, [FC: 6,200 ft (1890 m)], Zacatecas, Mexico. Collected 

16 Dec 1897 by E. A. Goldman. Original number [FC: Nelson and Goldman] 11968.
number 640 • 4 1

Perognathus intermedius Merriam

N. Amer. Fauna 1: 18, 25 Oct 1889e.

= Chaetodipus intermedius intermedius (Merriam, 1889). See 

Williams et al. (1993:144).

Holotype: [186509]. Skin and skull. Adult male. Mud Spring, 

[Mohave County], Arizona. Collected 26 Feb 1889 by 

V. Bailey. Original number [SL: 616]. [Merriam collection 

number] 5309/6000.

Perognathus (Chaetodipus) intermedius canescens Merriam

Proc. Acad. Nat. Sci. Philadelphia 46: 267, 27 Sep 1894a. Preprint 

of Proc. Acad. Nat. Sci. Philadelphia 46: 267, 23 Oct 1894a.

= Chaetodipus nelsoni canescens (Merriam, 1894). See Williams 

et al. (1993:147).

Holotype: 51016. [Skin and skull]. Young adult male. Jaral, [FC: 

4,000 ft (1,219 m)], Coahuila, Mexico. Collected 14 Jan 

1893 by C. P. Streator. Original number 2557.

Perognathus intermedius phasma Goldman

Proc. Biol. Soc. Washington 31: 22, 16 May 1918.

= Chaetodipus intermedius phasma (Goldman, 1918). See Williams et al. (1993:145).

Holotype: 203003. [Skin and skull]. Adult female. Tinajas Altas, 

[Hoffmeister (1986:286): Gila Mountains], altitude about 

1,400 ft (427 m), Yuma County, Arizona. Collected 23 Nov 

1913 by E. A. Goldman. Original number 22309.

Perognathus (Chaetodipus) nelsoni Merriam

Proc. Acad. Nat. Sci. Philadelphia 46: 266, 27 Sep 1894a. Preprint of Proc. Acad. Nat. Sci. Philadelphia 46: 266, 23 Oct 

1894a.

= Chaetodipus nelsoni nelsoni (Merriam, 1894). See Williams et 

al. (1993:148).

Holotype: 50214. [Skin and skull]. Old female. Hacienda La 

Parada, [FC: Altitude about 6,000 ft (1,829 m)], [Miller 

(1924:284): about 25 mi (40 km) northwest of the city of 

San Luis Potosi], San Luis Potosi, Mexico. Collected 19 Aug 

1892 by E. W. Nelson. Original number 3207.

Perognathus obscurus Merriam

N. Amer. Fauna 1: 20, 25 Oct 1889e.

= Chaetodipus intermedius intermedius (Merriam, 1889). See 

Williams et al. (1993:144).

Holotype: [186510]. Skin and skull. Adult female. Camp 

Apache, Grant (Hidalgo) County, New Mexico. Collected 

30 Apr 1886 by A. W. Anthony. Original number [MCC: 

62]. [Merriam collection number] 2340/2848.

Perognathus paradoxus Merriam

N. Amer. Fauna 1: 24, 25 Oct 1889e.

= Chaetodipus hispidus paradoxus (Merriam, 1889). See Paulson (1988b:1).

Holotype: [186513]. Skin and skull. [MCC: Adult] female. [MC: 

Banner], Trego County, Kansas. Collected [SL: 14], not 17 

as in original description, Apr 1884 by A. B. Baker. No original number. [Merriam collection number] 940/1544.

Perognathus paradoxus spilotus Merriam

N. Amer Fauna 1: 25, 25 Oct 1889e.

= Chaetodipus hispidus spilotus (Merriam, 1889). See Paulson 

(1988b:1).

Holotype: [186514]. Skin only. Adult female. Gainesville, Cook 

(Cooke) County, Texas. Collected 8 Oct 1886 by G. H. Ragsdale. No original number. [Merriam collection number] 5293. 

Merriam (1889e:25) based his description of the skull on 

USNM 23096, a young adult female from the type locality.

Perognathus penecillatus [sic] Woodhouse

Proc. Acad. Nat. Sci. Philadelphia 6: 200, presented at meeting 

of 28 Dec 1852b. Receipt acknowledged 7 Feb 1853 by the 

Smithsonian Institution.

= Chaetodipus penicillatus penicillatus (Woodhouse, 1852). See 

Williams et al. (1993:150).

Holotype: [2676/37437]. Skin, skull, [and baculum]. [Lyon and 

Osgood (1909:54): Adult] male. San Francisco Mountain 

(Peaks), New Mexico (Arizona). Type locality restricted by 

Hoffmeister and Lee (1967: 368) to 1 mi (1.6 km) southwest of Parker, Yuma County, Arizona. Collected [Poole and 

Schantz (1942:435): October] [Baird (1857:419): 1851 by 

S. W. Woodhouse]. Type not designated by number in original description but was based on a single specimen from the 

type locality. Reference to Baird (1857:419) indicates that 

2676 is the specimen on which the description is based.

Perognathus penicillatus albulus Nelson and Goldman

Proc. Biol. Soc. Washington 36: 159, 1 May 1923.

= Chaetodipus arenarius albulus (Nelson and Goldman, 1923). 

See Williams et al. (1993:127).

Holotype: 146864. [Skin and skull]. Adult male. Magdalena Island, [FC: 20 ft (6 m)], Lower California (Baja California 

Sur), Mexico. Collected 3 Dec 1905, not 1906 as in original 

publication, by E. W. Nelson and E. A. Goldman. Original 

number 18733.

Perognathus penicillatus ammophilus Osgood

Proc. Biol. Soc. Washington 20: 20, 23 Feb 1907b.

= Chaetodipus arenarius ammophilus (Osgood, 1907). See Williams et al. (1993:128).

Holotype: 146859. [Skin and skull]. Adult male. Margarita 

(Santa Margarita) Island, [FC: 25 ft (7.6 m)], Lower California (Baja California Sur), Mexico. Collected 29 Nov 

1905 by E. W. Nelson and E. A. Goldman. Original number 

[SL: 18655].

Perognathus penicillatus angustirostris Osgood

N. Amer. Fauna 18: 47, 20 Sep 1900b.

= Chaetodipus penicillatus angustirostris (Osgood, 1900). See 

Williams et al. (1993:149).
4 2 • smithsonian contributions to zoology

Holotype: 73881. [Skin and skull]. Adult male. Carrizo Creek, 

Colorado Desert, [Imperial County], California. Collected 

31 Mar 1895 by A. W. Anthony. Original number 22.

Perognathus penicillatus goldmani Townsend

Bull. Amer. Mus. Nat. Hist. 31: 122, 14 Jun 1912. Primary junior 

homonym of Perognathus goldmani Osgood (1900b:54). 

Replaced with Perognathus penicillatus seri by Nelson 

(1912:116).

= Chaetodipus penicillatus seri (Nelson, 1912). See Williams et 

al. (1993:151).

Holotype: [198411. Skin and skull]. Male. Tiburon Island, [Gulf 

of California, Sonora, Mexico]. Collected 13 Apr 1911 by 

[SL: H. E.. Anthony. Original number 155]. Specimen described as American Museum of Natural History number 

31845.

Perognathus penicillatus serosus Goldman

Proc. Biol. Soc. Washington 52: 34, 11 Mar 1939a. Primary 

junior homonym of Perognathus spinatus serosus Burt 

(1932:167). Replaced with Perognathus penicillatus sobrinus by Goldman (1939b:257).

= Chaetodipus penicillatus sobrinus (Goldman, 1939). See Williams et al. (1993:151).

Holotype: 27598/39697. Skin and skull. Adult male. Sand flat 

along Virgin River, 7 mi (11 km) above Bunkerville, Clark 

County, Nevada. Note: This locality is on the ArizonaNevada border. Hardy (1949:435) placed it in Arizona; 

Hoffmeister and Nader (1963:92) concluded it was in Nevada. Collected 9 May 1891 by V. Bailey. Original number 

2743.

Perognathus penicillatus siccus Osgood

Proc. Biol. Soc. Washington 20: 20, 23 Feb 1907b.

= Chaetodipus arenarius siccus (Osgood, 1907). See Williams et 

al. (1993:130).

Holotype: 146890. [Skin and skull]. Adult male. Ceralbo (Cerralvo/Jacque Cousteau) Island, Lower California (Baja California Sur), Mexico. Collected 13 Feb 1906 by E. W. Nelson 

and E. A. Goldman. Original number [SL: 19131].

Perognathus pernix rostratus Osgood

N. Amer. Fauna 18: 51, 20 Sep 1900b.

= Chaetodipus pernix rostratus (Osgood, 1900). See Williams et 

al. (1993:152).

Holotype: 95818. [Skin and skull]. Young adult male. Camoa, 

[FC: about 800 ft (244 m)], Rio Mayo, Sonora, Mexico. 

Collected 28 Oct 1898 by E. A. Goldman. Original number 

[FC: Nelson and Goldman] 13167.

Perognathus spinatus Merriam

N. Amer. Fauna 1: 21, 25 Oct 1889e.

= Chaetodipus spinatus spinatus (Merriam, 1889). See Lackey 

(1991b:1).

Holotype: [186516]. Skin, skull, [and baculum]. Adult male. 

Twenty-five miles (40 km) below [SL: south of] The Needles, 

Lower Colorado River, [San Bernardino County], California. Collected 23 Mar 1889 by V. Bailey. Original number 

[SL: 683]. [Merriam collection number] 5449/6137.

Perognathus spinatus magdalenae Osgood

Proc. Biol. Soc. Washington 20: 21, 23 Feb 1907b.

= Chaetodipus spinatus magdalenae (Osgood, 1907). See Lackey 

(1991b:1).

Holotype: 146102. [Skin and skull]. Adult female. Magdalena 

Island, [FC: 20–50 ft (6–15 m)], Lower California (Baja 

California Sur), Mexico. Collected 25 Nov 1905 by E. W. 

Nelson and E. A. Goldman. Original number [SL: 18633].

Perognathus spinatus nelsoni Townsend

Bull. Amer. Mus. Nat. Hist. 31: 122, 14 Jun 1912. Primary junior 

homonym of Perognathus nelsoni Merriam (1894a:266). 

Replaced with Perognathus spinatus occultus by Nelson 

(1912:116)

= Chaetodipus spinatus occultus (Nelson, 1912). See Lackey 

(1991b:1).

Holotype: [198409. Skin and skull]. Male. Carmen Island, [Gulf 

of California, Baja California Sur, Mexico]. Collected 3 Apr 

1911 by [SL: H. E. Anthony. Original number 99]. Specimen described as American Museum of Natural History 

number 31855.

Perognathus (Chaetodipus) stephensi Merriam

Proc. Acad. Nat. Sci. Philadelphia 46: 267, 27 Sep 1894a. Preprint of Proc. Acad. Nat. Sci. Philadelphia 46: 267, 23 Oct 

1894a.

= Chaetodipus penicillatus stephensi (Merriam, 1894). See Williams et al. (1993:151).

Holotype: 27774/39873. Skin and skull. Adult male. [Grinnell 

(1933:153): Edge of a little mesa near Triangle spring], Mesquite Valley, northwest arm of Death Valley, [Inyo County], 

California. Collected 6 Apr 1891 by F. Stephens. Original 

number 258.

Genus Dipodomys Gray, 1841

Dipodomys ambiguus Merriam

N. Amer. Fauna 4: 42, 8 Oct 1890e.

= Dipodomys merriami ambiguus Merriam, 1890. See Merriam 

(1898d:31).

Holotype: 18147/25045. Skin and skull. Adult male. El Paso, [El 

Paso County], Texas. Collected 13 Dec 1889 by V. Bailey. 

Original number 782.

Dipodomys californicus Merriam

N. Amer. Fauna 4: 29, 8 Oct 1890e.

= Dipodomys californicus californicus Merriam, 1890. See Kellogg (1916:366).
number 640 • 4 3

Holotype: 16618/23544. Skin and skull. Adult male. Ukiah, restricted to the west edge of the main road running south from 

Ukiah and about a mile (1.6 km) south of the then center of 

town by Grinnell (1933:156), Mendocino County, California. 

Collected 4 May 1889 by T. S. Palmer. Original number 46.

Dipodomys compactus True

Proc. U.S. Natl. Mus. 11: 160, 5 Jan 1889.

= Dipodomys compactus compactus True, 1889. See Schmidly 

and Hendricks (1976:235).

Holotype: 19665/35227. Skin only, skull lost. Adult female. 

Padre Island, Cameron County, Texas. Collected 3 Apr 

1888. Collector unknown. No original number. Received 

from C. K. Worthen. No type designated in original description. The description was based on a single specimen and 

the data, including three measurements, match that of the 

specimen listed above.

Dipodomys deserti Stephens

Amer. Nat. 21: 42, Jan 1887.

= Dipodomys deserti deserti Stephens, 1887. See Goldman 

(1923:139).

Holotype: [15629/22522. Skin and skull]. [Lyon and Osgood 

(1909:57): Adult] female. Mohave River, restricted to Mohave River bottom at upper crossing on old road from 

Cajon Pass to Rabbit Springs, 3 mi (4.8 km) or 4 mi (6.4 

km) from, and opposite Hesperia, by Grinnell (1933:165), 

[San Bernardino County], California. Collected 29 Jun 

1886 by [SL: F. Stephens]. Original number, under which it 

was published, 314.

Dipodomys deserti sonoriensis Goldman

Proc. Biol. Soc. Washington 36: 139, 1 May 1923.

Holotype: 242306. [Skin and skull]. Adult male, La Libertad 

Ranch, 30 mi (48 km) east of Sierra Seri, Sonora, Mexico. 

Collected 3 Jan 1922 by C. Sheldon. No original number. 

[SL: 22611-X].

Dipodomys elator Merriam

Proc. Biol. Soc. Washington 9: 109, 21 Jun 1894b.

Holotype: 64802. [Skin and skull]. Adult male. Henrietta, Clay 

County, Texas. Collected 13 Apr 1894 by J. A. Loring. Original number 1804.

Dipodomys heermanni gabrielsoni Goldman

Proc. Biol. Soc. Washington 38: 33, 12 Mar 1925.

= Dipodomys californicus californicus Merriam, 1890. See Patton et al. (1976:162).

Holotype: 244514. [Skin and skull]. Adult male. Brownsboro, 

Jackson County, Oregon. Collected 21 Jun 1924 by I. N. 

Gabrielson. Original number [SL: M 8. 23625-X].

Dipodomys insularis Merriam

Proc. Biol. Soc. Washington 20: 77, 22 Jul 1907.

= Dipodomys merriami insularis Merriam, 1907. See Best and 

Janecek (1992:5).

Holotype: 79053. [Skin and skull]. Adult female. San Jose Island, 

Gulf of California, Lower California (Baja California Sur), 

Mexico. Collected 6 Aug 1895 by J. E. McLellan. Original 

Number 1457.

Dipodomys margaritae Merriam

Proc. Biol. Soc. Washington 20: 76, 22 Jul 1907.

= Dipodomys merriami margaritae Merriam, 1907. See Lidicker 

(1960:204).

Holotype: 146058. [Skin and skull]. Young adult male. Margarita Island, [FC: 20 ft (6 m)], Lower California (Baja California Sur), [SL: Mexico]. Collected 1 Dec 1905 by [E. W.] 

Nelson and [E. A.] Goldman. Original number 18711.

Dipodomys merriami atronasus Merriam

Proc. Biol. Soc. Washington 9: 113, 21 Jun 1894b.

Holotype: 50276. [Skin and skull]. Adult male. Hacienda La 

Parada, [FC: about 6,000 ft (1,829 m)], [Miller (1924:291): 

about 25 mi (40 km) northwest of the city of San Luis Potosi], San Luis Potosi, Mexico. Collected 20 Aug 1892 by 

E. W. Nelson. Original number 3229.

Dipodomys merriami exilis Merriam

Proc. Biol. Soc. Washington 9: 113, 21 Jun 1894b.

= Dipodomys nitratoides exilis Merriam, 1894. See Grinnell 

(1921:96).

Holotype: 34843/43823. Skin and skull. Young adult male. 

Fresno, [FC: 328 ft (100 m) on sand hills 3 mi (5 km) west 

of town], San Joaquin Valley, [Fresno County], California. 

Grinnell (1933:161) restricted the locality to 4 mi (6.4 km) 

north of railroad station, along Southern Pacific right of 

way. Collected 23 Sep 1891 by V. Bailey. Original number 

3277.

Dipodomys merriami kernensis Merriam

Proc. Biol. Soc. Washington 20: 77, 22 Jul 1907.

= Dipodomys merriami merriami Mearns, 1890. See Grinnell 

(1921:97).

Holotype: 108884. [Skin and skull]. Adult male. Onyx [FC: 

PO, south fork Kern River], west end of Walker Pass, Kern 

County, California. Collected 25 Jul 1901 by W. K. Fisher. 

Original number 2106.

Dipodomys merriami mayensis Goldman

Proc. Biol. Soc. Washington 41: 141, 15 Oct 1928b.

Holotype: 96437. [Skin and skull]. Adult male. Alamos, [FC: 

about 1,200 ft (366 m)], Sonora, Mexico. Collected 19 Dec 

1898 by E. A. Goldman. Original number [FC: Nelson and 

Goldman] 13302.

Dipodomys merriami nevadensis Merriam

Proc. Biol. Soc. Washington 9: 111, 21 Jun 1894b.
4 4 • smithsonian contributions to zoology

= Dipodomys merriami merriami Mearns, 1890. See Grinnell 

(1921:97).

Holotype: 54552. [Skin and skull]. Adult female. Pyramid Lake, 

[FC: at Agency], [Washoe County], Nevada. Collected 26 

Jun 1893 by V. Bailey. Original number 3990.

Dipodomys merriami nitratoides Merriam

Proc. Biol. Soc. Washington 9: 112, 21 Jun 1894b.

= Dipodomys nitratoides nitratoides Merriam, 1894. See Grinnell (1921:96).

Holotype: 54674. [Skin and skull]. Adult male. Tipton, San Joaquin Valley, [Tulare County], California. Collected 25 Jun 

1893 by C. P. Streator. Original number 2978.

Dipodomys merriami nitratus Merriam

Proc. Biol. Soc. Washington 9: 112, 21 Jun 1894b.

= Dipodomys merriami merriami Mearns, 1890. See Grinnell 

(1921:97).

Holotype: 25359/32772. Skin and skull. Adult male. Keeler, east 

side of Owens Lake, [Inyo County], California. Collected 

29 Dec 1890 by E. W. Nelson. Original number 160.

Dipodomys merriami regillus Goldman

Proc. Biol. Soc. Washington 50: 75, 22 Jun 1937a.

= Dipodomys merriami merriami Mearns, 1890. See Lidicker 

(1960:165).

Holotype: 203017. Skin and skull. Young adult male. Tule Well, 

Tule Desert between Cabeza Prieta Mountains and Tule 

Mountains, [Hoffmeister (1986:312): 35 mi (56 km) south 

of Mohawk], altitude 1,000 ft (305 m), Yuma County, Arizona. Collected 4 Dec 1913 by E. A. Goldman. Original 

number 22345.

Dipodomys microps alfredi Goldman

Proc. Biol. Soc. Washington 50: 221, 28 Dec 1937b.

Holotype: 262846. [Skin and skull]. Adult female. Gunnison Island, altitude about 4,300 ft (1,311 m), Great Salt Lake, 

[Box Elder County], Utah. Collected 1 Jun 1937 by A. M. 

Bailey and J. Niedrach. No original number. [SL: Colorado 

Museum of Natural History number 2994. 28020-X].

Dipodomys microps bonnevillei Goldman

Proc. Biol. Soc. Washington 50: 222, 28 Dec 1937b.

Holotype: 31894/43755. Skin and skull. Adult female. Kelton, altitude about 4,300 ft (1,311 m), Box Elder County, 

Utah. Collected 7 Nov 1891 by V. Bailey. Original number 

3490.

Dipodomys microps celsus Goldman

J. Washington Acad. Sci. 14: 372, 19 Sep 1924.

Holotype: 243101. [Skin and skull]. Adult male. Six miles (9.6 

km) north of Wolf Hole, altitude 3,500 ft (1,067 m), [Mohave County], Arizona. Collected 16 Oct 1922 by E. A. 

Goldman. Original number 23411.

Dipodomys microps leucotis Goldman

Proc. Biol. Soc. Washington 44: 135, 17 Oct 1931a.

Holotype: 250036. [Skin and skull]. Adult male. Six miles (9.6 km) 

west of Colorado River (Navajo) Bridge, altitude 3,700 ft 

(1,128 m), Houserock Valley, north side of Marble Canyon 

of Colorado River, [Coconino County], Arizona. Collected 

8 Jun 1931 by E. A. Goldman. Original number 23570.

Dipodomys microps russeolus Goldman

J. Mamm. 20: 353, 14 Aug 1939c.

Holotype: 263895. Skin and skull. Adult male. Dolphin Island, 

altitude 4,250 ft (1,295 m), Great Salt Lake, [Box Elder 

County], Utah. Collected 5 Jun 1938 by W. H. Marshall. 

Original number 65.

Dipodomys microps subtenuis Goldman

J. Mamm. 20: 354, 14 Aug 1939c.

Holotype: 263917. Skin and skull. Adult female. Carrington Island, altitude 4,250 ft (1,295 m), Great Salt Lake, [Toole 

County], Utah. Collected 30 Jun 1938 by W. H. Marshall. 

Original number 182.

Dipodomys mitchelli Mearns

Proc. U.S. Natl. Mus. 19: 719, 30 Jul 1897b.

= Dipodomys merriami mitchelli Mearns, 1897. See Burt 

(1938:48).

Holotype: 63188. Skin and skull. Adult female. Tiburon Island, 

Gulf of California, Sonora, Mexico. Collected 23 Dec 1895 

by J. W. Mitchell. Original number 3.

Dipodomys montanus Baird

Proc. Acad. Nat. Sci. Philadelphia 7: 334, reported favorably for 

publication 24 Apr 1855a.

= Dipodomys ordii montanus Baird, 1855. See Grinnell (1921:96).

Holotype: 490/1631. Skin and skull. Near Fort Massachusetts (Fort Garland), [SL: N. Rio Grande Valley], [Costilla 

County, Colorado]. Collected [Baird (1857:412): 1853] by 

F. Kreutzfeldt. No original number. Type not designated by 

number in original description, but a footnote to the list of 

specimens in Baird (1857:412) states that 490 is the type.

Dipodomys nelsoni Merriam

Proc. Biol. Soc. Washington 20: 75, 22 Jul 1907.

Holotype: 79439. [Skin and skull]. Adult male. La Ventura, [FC: 

5,650 ft (1,722 m)], Coahuila, Mexico. Collected 10 Aug 

1896 by [E. W.] Nelson and [E. A.] Goldman. Original number 9998.

Dipodomys oklahomae Trowbridge and Whitaker

J. Mamm. 21: 343, 13 Aug 1940.

= Dipodomys ordii oklahomae Trowbridge and Whitaker, 1940. 

See Davis (1942:332).

Holotype: 265454. Skin and skull. Young adult female. North 

bank of the South Canadian River, 2.25 mi (3.6 km) south 
number 640 • 4 5

of Norman, Cleveland County, Oklahoma. Collected 16 Mar 

1934 by H. L. Whitaker. No original number. University of 

Oklahoma Museum of Zoology number 14517. 29312-X.

Dipodomys ordii Woodhouse

Proc. Acad. Nat. Sci. Philadelphia 6: 235, 13 Apr 1853b.

= Dipodomys ordii ordii Woodhouse, 1853. See Grinnell 

(1921:96).

No name-bearing status: 18135/25033. Skin and skull. Adult 

female. El Paso, [El Paso County], Texas. Collected 11 

Dec 1889 by V. Bailey. Original number 769. Williams et 

al. (1993:78) designated this specimen as Neotype of this 

taxon following Merriam (1890e:45). However, Williams 

et al. (1993) overlooked Koopman (1976:14), who lists a 

specimen thought to be from the type series as a Cotype 

(Syntype). The Syntype has precedence and the specimen 

given here currently has no standing as a name bearer.

Dipodomys ordii cineraceus Goldman

J. Mamm. 20: 352, 14 Aug 1939c.

Holotype: 263890. Skin and skull. Adult male. Dolphin Island, 

altitude 4,250 ft (1,295 m), Great Salt Lake, [Box Elder 

County], Utah. Collected 4 Jun 1938 by W. H. Marshall. 

Original number 57.

Dipodomys ordii cleomophila Goldman

J. Washington Acad. Sci. 23: 469, 15 Oct 1933b.

= Dipodomys ordii longipes (Merriam, 1890). See Setzer 

(1949:556).

Holotype: 226348. Skin and skull. Adult female. Five miles (8 

km) northeast of Winona, altitude 6,200 ft (1,890 m), Coconino County, Arizona. Collected 16 Jul 1917 by E. A. 

Goldman. Original number 23101.

Dipodomys ordii cupidineus Goldman

J. Washington Acad. Sci. 14: 372, 19 Sep 1924.

Holotype: 243093. [Skin and skull]. Adult male. Kanab Wash, at 

[SL: near] southern boundary of Kaibab Indian Reservation, 

[Mohave County], Arizona. Collected 12 Oct 1922 by E. A. 

Goldman. Original number 23384.

Dipodomys ordii evexus Goldman

J. Washington Acad. Sci. 23: 468, 15 Oct 1933b.

Holotype: 150990. Skin and skull. Adult male. Salida, altitude 

7,000 ft (2,134 m), Chaffee County, Colorado. Collected 10 

Nov 1907 by M. Cary. Original number 1245.

Dipodomys ordii fuscus Setzer

Univ. Kansas. Publ., Mus. Nat. Hist. 1: 555, 27 Dec 1949. Primary junior homonym of Dipodomys agilis fuscus Boulware (1943:393).

= Dipodomys ordii durranti Setzer. J. Washington Acad. Sci. 42: 

391, 17 Dec 1952. Replacement name for Dipodomys ordii 

fuscus Setzer.

Holotype: 93886. Skin and skull. Adult male. Juamave (Jaumave), [FC: 2,000 ft (610 m)], Tamaulipas, Mexico. Collected 3 Jun 1898 by E. W. Nelson and E. A. Goldman. 

Original number 12437.

Dipodomys ordii marshalli Goldman

Proc. Biol. Soc. Washington 50: 223, 28 Dec 1937b.

Holotype: 262655. [Skin and skull]. Adult female. Bird (Hat) 

Island, altitude 4,300 ft (1,310 m), Great Salt Lake, [Box 

Elder County], Utah. Collected 22 Jun 1937 by W. H. Marshall. Original number [SL: 39]. 27969-X.

Dipodomys ordii medius Setzer

Univ. Kansas. Publ., Mus. Nat. Hist. 1: 519, 27 Dec 1949.

Holotype: 118526. [Skin and skull]. Male. Santa Rosa, Guadalupe County, New Mexico. Collected 5 Oct 1902 by J. H. 

Gaut. Original number 565.

Dipodomys ordii nexilis Goldman

J. Washington Acad. Sci. 23: 470, 15 Oct 1933b.

Holotype: 149938. Skin and skull. Adult male. [FC: Dry creek] 

5 mi (8 km) west of Naturita, Montrose County, Colorado. 

Collected 20 Jul 1907 by M. Cary. Original number 1068.

Dipodomys ornatus Merriam

Proc. Biol. Soc. Washington 9: 110, 21 Jun 1894b.

= Dipodomys phillipsii ornatus Merriam, 1894. See Genoways 

and Jones (1971:281).

Holotype: 57990. [Skin and skull]. Adult female. Berriozabal, 

[FC: 6,800 ft (2,073 m)], Zacatecas, Mexico. Collected 29 

Dec 1893 by E. A. Goldman. Original number [FC: Nelson 

and Goldman] 5613.

Dipodomys perotensis Merriam

Proc. Biol. Soc. Washington 9: 111, 21 Jun 1894b.

= Dipodomys phillipsii perotensis Merriam, 1894. See Davis 

(1944:391).

Holotype: 54285. [Skin and skull]. Adult female. Perote, [FC: 

8,000 ft (2,438 m)], Veracruz, Mexico. Collected 21 May 

1893 by E. W. Nelson. Original number 4840.

Dipodomys platycephalus Merriam

Proc. Biol. Soc. Washington 20: 76, 22 Jul 1907.

= Dipodomys merriami platycephalus Merriam, 1907. See Huey 

(1927:66).

Holotype: 139882. [Skin and skull]. Adult male. Calmalli, [FC: 

1,200 ft (366 m)], Lower California (Baja California), Mexico. Collected 1 Oct 1905 by E. W. Nelson and E. A. Goldman. Original number 18248.

Dipodomys spectabilis Merriam

N. Amer. Fauna 4: 46, 8 Oct 1890e.

= Dipodomys spectabilis spectabilis Merriam, 1890. See Grinnell 

(1921:95).
4 6 • smithsonian contributions to zoology

Holotype: 17886/24823. Skin and skull. Adult male. Dos Cabezos (Cabezas), [FC: 5,300 ft (1,615 m)], Cochise County, 

Arizona. Collected 22 Nov 1889 by V. Bailey. Original number 695.

Dipodomys spectabilis baileyi Goldman

Proc. Biol. Soc. Washington 36: 140, 1 May 1923.

Holotype: 97185. [Skin and skull]. Adult male. Forty miles (64 

km) west of Roswell [FC: at Cedar Point ranch], [Lincoln 

County], New Mexico. Collected 13 Jun 1899 by V. Bailey. 

Original number 6961.

Dipodomys spectabilis clarencei Goldman

J. Washington Acad. Sci. 23: 467, 15 Oct 1933b.

= Dipodomys spectabilis baileyi Goldman, 1923. See Nader 

(1978:41).

Holotype: 158824. Skin and skull. Adult male. Blanco, San Juan 

County, New Mexico. Collected 19 Nov 1908 by C. Birdseye. Original number 443.

Dipodomys spectabilis cratodon Merriam

Proc. Biol. Soc. Washington 20: 75, 22 Jul 1907.

Holotype: 78953. [Skin and skull]. Adult male. Chicalote, [FC: 

6,180 ft (1,884 m)], Aguascalientes, Mexico. Collected 2 Jul 

1896 by E. W. Nelson and E. A. Goldman. Original number 

9734.

Dipodomys spectabilis perblandus Goldman

J. Washington Acad. Sci. 23: 466, 15 Oct 1933b.

Holotype: 17748/24689. Skin and skull. Adult female. Calabasas, altitude about 3,500 ft (1,067 m), Santa Cruz County, 

Arizona. Collected 27 Oct 1889 by V. Bailey. Original number 611.

Dipodomys spectabilis zygomaticus Goldman

Proc. Biol. Soc. Washington 36: 140, 1 May 1923.

Holotype: 96432. [Skin and skull]. Adult male. Parral, [FC: 

6,000 ft (1,829 m)], southern Chihuahua, Mexico. Collected 17 Sep 1898 by E. A. Goldman. Original number 

[FC: Nelson and Goldman] 13030.

Dipodops longipes Merriam

N. Amer. Fauna 3: 72, 11 Sep 1890b.

= Dipodomys ordii longipes (Merriam, 1890). See Grinnell 

(1921:96).

Holotype: 17703/24639. Skin and skull. Young adult male. Foot 

of Echo Cliffs, Painted Desert, [SL and FC: Moa Ave], [FC: 

Coconino County], Arizona. Collected 22 Sep 1889 by 

C. H. Merriam. Original number [FC: V. Bailey] 512.

Perodipus agilis tularensis Merriam

Proc. Biol. Soc. Washington 17: 143, 14 Jul 1904b.

= Dipodomys heermanni tularensis (Merriam, 1904). See Grinnell (1921:95).

Holotype: 127158. [Skin and skull]. Adult female. Alila (Earlimart), Tulare County, California. Collected 23 Jun 1903 

by L. J. Goldman. Original number 563.

Perodipus cabezonae Merriam

Proc. Biol. Soc. Washington 17: 144, 14 Jul 1904b.

= Dipodomys simulans simulans (Merriam, 1904). See Williams 

et al. (1993:86).

Holotype: 54055. [Skin and skull]. Adult female. Cabezon, Colorado Desert, [Grinnell (1922:96): San Gorgonio Pass, Riverside County], California. Collected 31 May 1893 by C. P. 

Streator. Original number 2859.

Perodipus goldmani Merriam

Proc. Biol. Soc. Washington 17: 143, 14 Jul 1904b.

= Dipodomys heermanni goldmani (Merriam, 1904). See Grinnell (1921:95).

Holotype: 118924. [Skin and skull]. Adult male. Salinas, mouth 

of Salinas Valley, Monterey County, California. Collected 

4 Sep 1902 by L. J. Goldman. Original number 431.

Perodipus ingens Merriam

Proc. Biol. Soc. Washington 17:141, 14 Jul 1904b.

= Dipodomys ingens (Merriam, 1904). See Grinnell (1921:95).

Holotype: 128805. [Skin and skull]. Adult male. Painted Rock, 

20 mi (32 km) [Grinnell (1922:69): 12 mi (19 km)] southeast of Simmler, Carrizo Plain, San Luis Obispo County, 

California. Collected 6 Aug 1903 by L. J. Goldman. Original number 777.

Perodipus microps Merriam

Proc. Biol. Soc. Washington 17: 145, 14 Jul 1904b.

= Dipodomys microps microps (Merriam, 1904). See Grinnell 

(1921:96).

Holotype: 25288/32701. Skin and skull. Adult male. Lone Pine, 

Owens Valley, [FC: lower, or eastern base of Alabama 

Mountains], Inyo County, California. Collected 22 Dec 

1890 by E. W. Nelson. Original number 138.

Perodipus microps levipes Merriam

Proc. Biol. Soc. Washington 17: 145, 14 Jul 1904b.

= Dipodomys microps levipes (Merriam, 1904). See Hall (1931:5).

Holotype: 27176/34575. Skin and skull. Adult male. Perognathus Flat, altitude 5,200 ft (1,585 m), [FC: at head of] Emigrant Gap, Panamint Mountains, [Inyo County], California. 

Restricted to near Bench Mark 4899, 2 mi (3 km) northwest 

of Harrisburg, on U.S Geological Survey. Ballarat Quadrangle, edition of Mar 1913 by Grinnell (1922:105). Collected 

16 Apr 1891 by V. Bailey. Original number 2668.

Perodipus microps preblei Goldman

J. Mamm. 2: 233, 29 Nov 1921.

= Dipodomys microps preblei (Goldman, 1921). See Miller 

(1924:297).
number 640 • 4 7

Holotype: 79340. [Skin and skull]. Adult female. Narrows, Malheur Lake, Harney County, Oregon. Collected 23 Jul 1896 

by E. A. Preble. Original number 1201.

Perodipus montanus utahensis Merriam

Proc. Biol. Soc. Washington 17: 143, 14 Jul 1904b.

= Dipodomys ordii utahensis (Merriam, 1904). See Grinnell 

(1921:96).

Holotype: 55115. [Skin and skull]. Adult male. Odgen, [Weber 

County], Utah. Collected 15 Jul 1893 by V. Bailey. Original 

number 4085.

Perodipus morroensis Merriam

Proc. Biol. Soc. Washington 20: 78, 22 Jul 1907.

= Dipodomys heermanni morroensis (Merriam, 1907). See Boulware (1943:393).

Holotype: 31626/43499. Skin and skull. Adult female. [Grinnell 

(1922:59): About 4 mi (6.4 km) south of] Morro, San Luis 

Obispo County, California. Collected 11 Nov 1891 by E. W. 

Nelson. Original number 1464.

Perodipus ordii columbianus Merriam

Proc. Biol. Soc. Washington 9: 115, 21 Jun 1894b.

= Dipodomys ordii columbianus (Merriam, 1894). See Grinnell 

(1921:96).

Holotype: 24181/31594. Skin and skull. Adult female. Umatilla, 

[Miller (1924:298): at mouth of Umatilla River], Plains of 

Columbia, [Umatilla County], Oregon. Collected 18 Oct 

1890 by C. P. Streator. Original number 386.

Perodipus ordii luteolus Goldman

Proc. Biol. Soc. Washington 30: 112, 23 May 1917a.

= Dipodomys ordii luteolus (Goldman, 1917). See Grinnell 

(1921:96).

Holotype: 160408. [Skin and skull]. Adult male. Casper, [FC: 

north side of Platte River, 4,700 ft (1,433 m)], [Natrona 

County], Wyoming. Collected 2 Sep 1909 by M. Cary. Original number 1690.

Perodipus panamintinus Merriam

Proc. Biol. Soc. Washington 9: 114, 21 Jun 1894b.

= Dipodomys panamintinus panamintinus (Merriam, 1894). See 

Hall (1946:409).

Holotype: 28566/40670. Skin and skull. Adult male. On head of 

Willow Creek, [FC: 5,040 ft (1,536 m)], Panamint Mountains, [Inyo County], California. Collected 12 May 1891 by 

E. W. Nelson. Original number 853.

Perodipus perplexus Merriam

Proc. Biol. Soc. Washington 20: 79, 22 Jul 1907.

= Dipodomys agilis perplexus (Merriam, 1907). See Grinnell 

(1921:96).

Holotype: 29261/41328. Skin and skull. Adult male. Walker 

Basin, [Grinnell (1933:163): 3,400 ft (1,036 m)], Kern 

County, California. Collected 15 Jul 1891 by V. Bailey. 

Original number 3053.

Perodipus simulans peninsularis Merriam

Proc. Biol. Soc. Washington 20: 79, 22 Jul 1907.

= Dipodomys simulans peninsularis (Merriam, 1907). See Williams et al. (1993:85).

Holotype: 139872. [Skin and skull]. Young adult male. Santo 

Domingo [Huey (1951:246): (Santo Domingo Landing), 

latitude 28°51′N], [FC: 150 ft (46 m)], Lower California 

(Baja California), Mexico. Collected 27 Sep 1905 by [E. W.] 

Nelson and [E. A.] Goldman. Original number 18215.

Perodipus stephensi Merriam

Proc. Biol. Soc. Washington 20: 78, 22 Jul 1907.

= Dipodomys stephensi (Merriam, 1907). See Grinnell (1921:95).

Holotype: [186503]. Skin and skull. Adult male. San Jacinto Valley, [Grinnell (1922:67): a little west of the present town of 

Winchester, toward Menifee], Riverside County, California. 

Collected 27 Nov 1885 by F. Stephens. Original number 

[MCC: 206]. Merriam collection number 1808/2340.

Perodipus streatori Merriam

Proc. Biol. Soc. Washington 9: 113, 21 Jun 1894b.

= Dipodomys heermanni heermanni (LeConte, 1853). See Grinnell (1921:97).

Holotype: 64310. [Skin and skull]. Adult female. Carbondale, at 

west foot of Sierra Nevada, Amador County, California. Collected 3 Apr 1894 by C. P. Streator. Original number 3673.

Perodipus streatori simulans Merriam

Proc. Biol. Soc. Washington 17: 144, 14 Jul 1904b.

= Dipodomys simulans simulans (Merriam, 1904). See Williams 

et al. (1993:86).

Holotype: 33105/45103. Skin and skull. Adult female. Dulzura, 

San Diego County, California. Collected 24 Nov 1891 by 

C. H. Marsh. Original number 255.

Perodipus venustus Merriam

Proc. Biol. Soc. Washington 17: 142, 14 Jul 1904b.

= Dipodomys venustus venustus (Merriam, 1904). See Grinnell 

(1921:96).

Holotype: 51852. [Skin and skull]. Adult male. Santa Cruz, 

Santa Cruz County, California. Collected 12 Mar 1893 by 

G. B. Badger. Original number 46.

Genus Heteromys Desmarest, 1817

Heteromys annectens Merriam

Proc. Biol. Soc. Washington 15: 43, 5 Mar 1902b.

= Heteromys pictus annectens Merriam, 1902. New name combination.

Holotype: 71510. [Skin and skull]. Adult male. Pluma, [FC: 

3,000 ft (914 m)], Oaxaca, Mexico. Collected 18 Mar 
4 8 • smithsonian contributions to zoology

1895 by E. W. Nelson and E. A. Goldman. Original number 7674.

Heteromys australis conscius Goldman

Smithsonian Misc. Coll. 60 (22): 8, 28 Feb 1913.

Holotype: 178699. [Skin and skull]. Adult male. Cana, altitude 

2,000 ft (610 m), [Darien], [Miller (1924:266): mountains 

of] eastern Panama. Collected 8 Mar 1912 by E. A. Goldman. Original number 21468.

Heteromys catopterius Anderson and Gutierrez

Bull. Amer. Mus. Nat. Hist. 331: 40, 15 Dec 2009.

Holotype: 517536. Skin and skull. Adult male. Rancho Grande, 

3,576 ft (1,090 m), near Biological Station, 13 km northwest Maracay, Aragua, Venezuela, 10°21′N, 67°40′W. Collected 20 Mar 1960 by C. O. Handley, Jr. and D. I. Rhymer. 

Original number COH, Jr. 7778.

Heteromys crassirostris Goldman

Smithsonian Misc. Coll. 60 (2): 10, 20 Sep 1912b.

= Heteromys desmarestianus crassirostris Goldman, 1912. See 

Goldman (1920:117).

Holotype: 179016. [Skin and skull]. Adult male. Near head 

of Rio Limon, altitude 5,000 ft (1,524 m), Mount Pirri, 

[Darien], eastern Panama. Collected 26 Apr 1912 by E. A. 

Goldman. Original number 21649.

Heteromys desmarestianus planifrons Goldman

J. Washington Acad. Sci. 27: 418, 15 Oct 1937c.

Holotype: 250348. Skin and skull. Adult female. San Geronimo, 

Pirris, western Costa Rica. Collected 10 Apr, not 12 Apr as 

in original description, 1931 by C. F. Underwood. Original 

number [SL: 259]. 26913-X, not 26914-X as in original description.

Heteromys desmarestianus subaffinis Goldman

J. Washington Acad. Sci. 27: 420, 15 Oct 1937c.

Holotype: 12904/38591. Skin and skull. Adult male. Angostura, 

altitude about 1,980 ft (604 m), southern side of Rio Reventazon, opposite Turrialba, Costa Rica. Collected May 1876 

by J. C. Zeledon. Original number 98.

Heteromys goldmani Merriam

Proc. Biol. Soc. Washington 15: 41, 5 Mar 1902b.

See Rogers and Gonzales (2010:914) for recognition of goldmani at the specific level.

Holotype: 77576. [Skin and skull]. Adult male. Chicharras, [FC: 

3,200 ft (975 m)], Chiapas, Mexico. Collected 7 Feb 1896 

by E. W. Nelson and E. A. Goldman. Original number 9244.

Heteromys goldmani lepturus Merriam

Proc. Biol. Soc. Washington 15: 42, 5 Mar 1902b.

= Heteromys desmarestianus desmarestianus Gray, 1868. See 

Rogers and Schmidly (1982:375).

Holotype: 73382. [Skin and skull]. Adult male. Mountains 

near Santo Domingo, [FC: 1,600 ft (488 m)], [Goldman 

(1911:25): a few miles west of Guichicovi], Oaxaca, Mexico. Collected 20 Jun 1895 by E. W. Nelson and E. A. Goldman. Original number 8120.

Heteromys griseus Merriam

Proc. Biol. Soc. Washington 15: 42, 5 Mar 1902b.

= Heteromys desmarestianus desmarestianus Gray, 1868. See 

Rogers and Schmidly (1982:383).

Holotype: 76062. [Skin and skull]. Adult male. Mountains near 

Tonala, [FC: 2,800 ft (853 m)], Chiapas, Mexico. Collected 

15 Aug 1895 by E. W. Nelson and E. A. Goldman. Original 

number 8339.

Heteromys (Xylomys) nelsoni Merriam

Proc. Biol. Soc. Washington 15: 43, 5 Mar 1902b.

= Heteromys nelsoni Merriam, 1902. See Goldman (1911:31).

Holotype: 77920. [Skin and skull]. Adult male. Pinabete, [FC: 

8,200 ft (2,499 m)], Chiapas, Mexico. Collected 11 Feb 

1896 by E. W. Nelson and E. A. Goldman. Original number 

9281.

Heteromys oasicus Anderson

Amer. Mus. Nov. 3396: 9, 27 Mar 2003.

Holotype: 456325. Skin and skull. Adult male. Cerro Santa 

Ana, 550 m, 49 km north, 32 km west of Coro [Handley 

(1976:47): Peninsula de Paraguana, 15 km SSW Pueblo 

Nuevo], Falcon, Venezuela. Collected 25 Jul 1968 by N. E. 

Peterson. Original number SVP (Smithsonian Venezuelan 

Project) 24036.

Heteromys panamensis Goldman

Smithsonian Misc. Coll. 56 (36): 9, 19 Feb 1912a.

= Heteromys desmarestianus panamensis Goldman, 1912. See 

Goldman (1920:117).

Holotype: 171107. [Skin and skull]. Adult male. Cerro Azul, 

altitude 2,800 ft (853 m), near headwaters of the Chagres 

River, [Panama], Panama. Collected 23 Mar 1911 by E. A. 

Goldman. Original number [FC: Nelson and Goldman] 

20999.

Heteromys temporalis Goldman

N. Amer. Fauna 34: 26, 7 Sep 1911.

= Heteromys desmarestianus temporalis Goldman, 1911. See 

Rogers and Schmidly (1982:385).

Holotype: 63719. [Skin and skull]. Adult female. Motzorongo, 

Veracruz, Mexico. Collected 3 Mar 1894 by [E. W.] Nelson 

and [E. A.] Goldman. Original number 5915.

Heteromys zonalis Goldman

Smithsonian Misc. Coll. 56 (36): 9, 19 Feb 1912a.

= Heteromys desmarestianus zonalis Goldman, 1912. See Goldman (1920:116).
number 640 • 4 9

Holotype: 170976. [Skin and skull]. Adult female. Rio Indio, 

near Gatun, Canal Zone (Colon), Panama. Collected 15 Feb 

1911 by E. A. Goldman. Original number [FC: Nelson and 

Goldman] 20885.

Liomys canus Merriam

Proc. Biol. Soc. Washington 15: 44, 5 Mar 1902b.

= Heteromys irroratus alleni Coues, 1881. New name combination. See Hafner et al. (2007:1140) for use of Heteromys.

Holotype: 96259. [Skin and skull], Adult male. [SL: Near] Parral, [FC: 6,200 ft (1,890 m)], Chihuahua, Mexico. Collected 

21 Sep 1898 by E. W. Nelson and E. A. Goldman. Original 

number 13036.

Liomys crispus Merriam

Proc. Biol. Soc. Washington 15: 49, 5 Mar 1902b.

= Heteromys salvini crispus (Merriam, 1902). New combination. See Hafner et al. (2007:1140) for use of Heteromys.

Holotype: 75105. [Skin and skull]. Adult male, Tonala, [FC: 

500 ft (152 m)], Chiapas, Mexico. Collected 7 Aug 1895 by 

E. W. Nelson and E. A. Goldman. Original number 8283.

Liomys crispus setosus Merriam

Proc. Biol. Soc. Washington 15: 49, 5 Mar 1902b.

= Heteromys salvini crispus (Merriam, 1902). New name combination. See Hafner et al. (2007:1140) for use of Heteromys.

Holotype: 77588. [Skin and skull]. Adult female. Huehuetan, 

[FC: 500 ft (152 m)], Chiapas, Mexico. Collected 22 Feb 

1896 by E. W. Nelson and E. A. Goldman. Original number 

9364.

Liomys guerrerensis Goldman

N. Amer. Fauna 34: 62, 7 Sep 1911.

= Heteromys irroratus guerrerensis (Goldman, 1911). New 

name combination. See Hafner et al. (2007:1140) for use 

of Heteromys.

Holotype: 127523. [Skin and skull]. Young female. Omilteme, 

Guerrero, Mexico. Collected 17 May 1903 by [E. W.] Nelson and [E.A.] Goldman. Original number 16435.

Liomys heterothrix Merriam

Proc. Biol. Soc. Washington 15: 50, 5 Mar 1902b.

= Heteromys salvini salvini Thomas, 1893. New name combination. See Hafner et al. (2007:1140) for use of Heteromys.

Holotype: 90161. [Skin and skull]. Adult male. San Pedro Sula, 

[Cortes], Honduras. Collected 16 Jul 1897 by J. C. Ingersoll. Original number 20. 884-X.

Liomys irroratus pretiosus Goldman

N. Amer. Fauna. 34: 58, 7 Sep 1911.

= Heteromys irroratus texensis (Merriam, 1902). New name combination. See Hafner et al. (2007:1140) for use of Heteromys.

Holotype: 93097. [Skin and skull]. Adult male. Metlaltoyuca, 

[FC: about 800 ft (244 m)], Puebla, Mexico. Collected 25 

Jan 1898 by E. A. Goldman. Original number [FC: Nelson 

and Goldman] 12081.

Liomys obscurus Merriam

Proc. Biol. Soc. Washington 15: 48, 5 Mar 1902b.

= Heteromys pictus pictus Thomas, 1893. New name combination. See Hafner et al. (2007:1140) for use of Heteromys.

Holotype: 108563. [Skin and skull]. Adult female. Carrizal, 

Veracruz, Mexico. Collected 12 May 1901 by E. W. Nelson 

and E. A. Goldman. Original number 14714.

Liomys orbitalis Merriam

Proc. Biol. Soc. Washington 15: 48, 5 Mar 1902b.

= Heteromys pictus pictus Thomas, 1893. New name combination. See Hafner et al. (2007: 1140) for use of Heteromys.

Holotype: 65452. [Skin and skull]. Adult female. Catemaco, [FC: 

1,000 ft (305 m)], Veracruz, Mexico. Collected 29 Apr 1894 

by E. W. Nelson and E. A. Goldman. Original number 6129.

Liomys parviceps Goldman

Proc. Biol. Soc. Washington 17: 82, 21 Mar 1904.

= Heteromys pictus plantinarensis (Merriam, 1902). New name 

combination. See Hafner et al. (2007:1140) for use of Heteromys.

Holotype: 126477. [Skin and skull]. Adult female. La Salada, 

40 mi (64 km) south of Uruapan, Michoacan, Mexico. Collected 19 Mar 1903 by E. W. Nelson and E. A. Goldman. 

Original number 16194.

Liomys phaeura Merriam

Proc. Biol. Soc. Washington 15: 48, 5 Mar 1902b.

= Heteromys pictus pictus Thomas, 1893. New name combination. See Hafner et al. (2007:1140) for use of Heteromys.

Holotype: 71500. [Skin and skull]. Adult female, Pinotepa, [FC: 

700 ft (213 m)], Oaxaca, Mexico. Collected 21 Feb 1895 by 

E. W. Nelson and E. A. Goldman. Original number 7553.

Liomys pictus isthmius Merriam

Proc. Biol. Soc. Washington 15: 46, 5 Mar 1902b.

= Heteromys pictus pictus Thomas, 1893. New name combination. See Hafner et al. (2007:1140) for use of Heteromys.

Holotype: 73367. [Skin and skull]. [Lyon and Osgood (1909:45): 

Adult] male. Tehuantepec, [FC: 150 ft (46 m)], [Oaxaca], 

Mexico. Collected 28 Apr 1895 by E. W. Nelson and E. A. 

Goldman. Original number 7796.

Liomys pictus rostratus Merriam

Proc. Biol. Soc. Washington 15: 46, 5 Mar 1902b.

= Heteromys pictus pictus Thomas, 1893. New name combination. See Hafner et al. (2007:1140) for use of Heteromys.

Holotype: 71488. [Skin and skull]. Adult male. Near Ometepec, 

[FC: 200 ft (61 m)], Guerrero, Mexico. Collected 14 Feb 

1895 by E. W. Nelson and E. A. Goldman. Original number 

7447.
5 0 • smithsonian contributions to zoology

Liomys plantinarensis Merriam

Proc. Biol. Soc. Washington 15: 46, 5 Mar 1902b.

= Heteromys pictus plantinarensis (Merriam, 1902). New name 

combination. See Hafner et al. (2007:1140) for use of 

Heteromys.

Holotype: 33595/45630. Skin and skull. Adult female. Plantinar, 

[FC: about 4,000 ft (1,219 m)], Jalisco, Mexico. Collected 4 

Apr 1892 by E. W. Nelson, not E. A. Goldman as in original 

description. Original number 2383.

Liomys sonorana Merriam

Proc. Biol. Soc. Washington 15: 47, 5 Mar 1902b.

= Heteromys pictus hispidus (J. A. Allen, 1897). New name combination. See Hafner et al. (2007:1140) for use of Heteromys.

Holotype: 96252. [Skin and skull]. Adult male. Alamos, [FC: 

about 1,200 ft (366 m)], Sonora, Mexico. Collected 19 Dec 

1898 by E. A. Goldman. Original number [FC: Nelson and 

Goldman] 13299.

Liomys texensis Merriam

Proc. Biol. Soc. Washington 15: 44, 5 Mar 1902b.

= Heteromys irroratus texensis (Merriam, 1902). New name 

combination. See Hafner et al. (2007:1140) for use of 

Heteromys.

Holotype: 58670. [Skin and skull]. Adult female. Brownsville, 

[Cameron County], Texas. Collected 19 Feb 1894 by J. A. 

Loring. Original number 1672.

Liomys torridus Merriam

Proc. Biol. Soc. Washington 15: 45, 5 Mar 1902b.

= Heteromys irroratus torridus (Merriam, 1902). New name combination. See Hafner et al. (2007:1140) for use of Heteromys.

Holotype: 69645. [Skin and skull]. Adult female. Cuicatlan, [FC: 

1,800 ft (549 m)], Oaxaca, Mexico. Collected 14 Oct 1894 

by E. W. Nelson and E. A. Goldman. Original number 6904.

Liomys torridus minor Merriam

Proc. Biol. Soc. Washington 15: 45, 5 Mar 1902b.

= Heteromys irroratus torridus (Merriam, 1902). New name 

combination. See Hafner et al. (2007:1140) for use of 

Heteromys.

Holotype: 70301. [Skin and skull]. Adult female. Huajuapam, 

[FC: 5,500 ft (1,676 m)], Oaxaca, Mexico. Collected 18 

Nov 1894 by E. W. Nelson and E. A. Goldman. Original 

number 7061.

Liomys veraecrucis Merriam

Proc. Biol. Soc. Washington 15: 47, 5 Mar 1902b.

= Heteromys pictus pictus Thomas, 1893. New name combination. See Hafner et al. (2007:1140) for use of Heteromys.

Holotype: 65457. [Skin and skull]. Adult female. San Andres 

Tuxtla, [FC: 1,000 ft (305 m)], Veracruz, Mexico. Collected 

7 May 1894 by E. W. Nelson and E. A. Goldman. Original 

number 6174.

Genus Microdipodops Merriam, 1891

Microdipodops californicus Merriam

Proc. Biol. Soc. Washington 14: 128, 19 Jul 1901c.

= Microdipodops megacephalus californicus Merriam, 1901. See 

Hall (1941:250).

Holotype: 101227. [Skin and skull]. Young adult male. Sierra 

Valley, near Vinton, [FC: pass, 5,100 ft (1,544 m)], Plumas 

County, California. Collected 7 Aug 1900 by W. K. Fisher. 

Original number 1596.

Microdipodops megacephalus Merriam

N. Amer. Fauna 5: 116, 30 Jul 1891a.

= Microdipodops megacephalus megacephalus Merriam, 1891. 

See Miller (1912:280).

Holotype: 24417/31823. Skin and skull. Adult male. Halleck, 

[Miller and Rhen (1901:155): East Humboldt Valley, Elko 

County], Nevada. Collected 23 Oct 1890 by V. Bailey. Original number 2005.

Microdipodops megacephalus lucidus Goldman

Proc. Biol. Soc. Washington 39: 127, 27 Dec 1926.

= Microdipodops pallidus pallidus Merriam, 1901. See Hall 

(1941:269).

Holotype: 210397. Skin and skull. Adult male. Clayton Valley, 8 

mi (13 km) southeast of Blair, altitude about 4,500 ft (1,372 

m) [SL and FC: 4,000 ft (1,219 m)], [Esmeralda County], 

Nevada. Collected 19 Oct 1915 by L. J. Goldman. Original 

number 2424.

Microdipodops megacephalus oregonus Merriam

Proc. Biol. Soc. Washington 14: 127, 19 Jul 1901c.

Holotype: 80128. [Skin and skull]. Young adult male. Lake Alvord, Alvord Desert, [Harney County], eastern Oregon. 

Restricted to Wild Horse Creek, 4 mi (6 km) northwest of 

Alvord Lake by Hall (1941:248). Collected 18 Aug 1896 by 

C. P. Streator. Original number 5430.

Microdipodops pallidus Merriam

Proc. Biol. Soc. Washington 14: 127, 19 Jul 1901c.

= Microdipodops pallidus pallidus Merriam, 1901. See Hall 

(1941:269).

Holotype: 93520. [Skin and skull]. Adult female. [SL and 

FC: Mountains] 10 mi (16 km) east of Stillwater [Hall 

(1941:269): Mountain Well], near Sink of the Humboldt 

and Carson, Churchill County, Nevada. Collected 11 May 

1898 by H. C. Oberholser. Original number 101.

Genus Perognathus Wied-Neuwied, 1839

Cricetodipus parvus Peale

U.S. Explor. Exped. 8: 53, 1848.

= Perognathus parvus parvus (Peale, 1848). See Miller 

(1912:264).
number 640 • 5 1

Holotype: No number given in original description. [Cassin (1858:48): alcoholic]. Young female. Oregon. Osgood 

(1900b:34) assumed the locality to be The Dalles, Wasco 

County, Oregon. Date of collection and collector unknown. 

The Type specimen was part of the U.S. Exploring Expedition collection that was sent to Washington, D.C., and later 

became part of the USNM. No specimen currently in the 

collection meets any criteria of the type.

Otognosis longimembris Coues

Proc. Acad. Nat. Sci. Philadelphia 27: 305, 31 Aug 1875a.

= Perognathus longimembris longimembris (Coues, 1875). See 

Miller (1924:276).

Holotype: 9858/37356. Alcoholic [Skull removed]. Female. Fort 

Tejon, [Osgood (1900b:33): Canada de las Uvas (Grapevine 

Canyon), Kern County], California. Collected [Poole and 

Schantz (1942:431): 1857–1858] by J. Xantus. No original 

number.

Perognathus amplus Osgood

N. Amer. Fauna 18: 32, 20 Sep 1900b.

= Perognathus amplus amplus Osgood, 1900. See Miller and 

Kellogg (1955:363).

Holotype: 34626/46711. Skin and skull. Adult male. Fort [SL: 

Camp] Verde, [Yavapai County], Arizona. Collected 26 Jun 

1892 by J. A. Loring. Original number 272.

Perognathus amplus jacksoni Goldman

J. Washington Acad. Sci. 23: 465, 15 Oct 1933b.

= Perognathus amplus amplus Osgood, 1900. See Hoffmeister 

(1986: 267).

Holotype: 212780. Skin and skull. Adult male. Congress Junction, 

altitude 3,000 ft (914 m), Yavapai County, Arizona. Collected 

21 Jun 1916 by H. H. T. Jackson. Original number 381.

Perognathus amplus pergracilis Goldman

J. Washington Acad. Sci. 22: 387, 19 Jul 1932c.

= Perognathus amplus amplus Osgood, 1900. See McKnight 

(2005:830).

Holotype: 227528. [Skin and skull]. Young adult male. Hackberry, altitude 3,500 ft (1,067 m), Mohave County, Arizona. 

Collected 14 Sep 1917 by E. A. Goldman. Original number 

23304.

Perognathus amplus rotundus Goldman

J. Washington Acad. Sci. 22: 387, 19 Jul 1932c.

= Perognathus amplus amplus Osgood, 1900. See Hoffmeister 

(1986:267).

Holotype: 250470. [Skin and skull]. Adult male. Welton, Yuma 

County, Arizona. Collected 9 Nov 1931 by B. Bailey. Original number A4353. 27029-X.

Perognathus amplus taylori Goldman

J. Washington Acad. Sci. 22: 488, 19 Oct 1932d.

Holotype: 250533. [Skin and skull]. Adult female. Santa Rita Range 

Reserve, altitude 4,000 ft (1,219 m), near Northeast Station, 

35 mi (56 km) south of Tucson, Pima County, Arizona. Collected 3 Aug 1930 by W. P. Taylor. Original number 1899.

Perognathus apache Merriam

N. Amer. Fauna 1: 14, 25 Oct 1889e.

= Perognathus flavescens apache Merriam, 1889. See Williams 

(1978:45).

Holotype: [186504]. Skin and skull. Adult male. [SL: Near 

Keam’s Canyon], Navajo County, Arizona. Collected 22 

May 1888 by [MCC: J. Sullivan]. Original number [MCC: 

11]. Merriam collection number 4253/4984.

Perognathus apache caryi Goldman

Proc. Biol. Soc. Washington 31: 24, 16 May 1918.

= Perognathus flavescens caryi Goldman, 1918. See Williams 

(1978:51).

Holotype: 148206. [Skin and skull]. Adult male. Eight miles (13 

km) west of Rifle, Garfield County, Colorado. Collected 4 

Oct 1906 by M. Cary. Original number 937.

Perognathus apache cleomophila Goldman

Proc. Biol. Soc. Washington 31: 23, 16 May 1918.

= Perognathus flavescens apache Merriam, 1889. See Williams 

(1978:45).

Holotype: 226344. Skin and skull. Adult male. [SL: 3 mi (5 km) 

northwest] Winona, altitude 6,400 ft (1,951 m), Coconino 

County, Arizona. Collected 19 Jul 1917 by E. A. Goldman. 

Original number 23127.

Perognathus apache melanotis Osgood

N. Amer. Fauna 18: 27, 20 Sep 1900b.

= Perognathus flavescens melanotis Osgood, 1900. See Williams 

(1978:54).

Holotype: 97416. [Skin and skull]. Adult female. Casas Grandes, 

[FC: 4,300 ft (1,311 m)], Chihuahua, Mexico. Collected 21 

May 1899 by E. A. Goldman. Original number [FC: Nelson 

and Goldman] 13750.

Perognathus apache relictus Goldman

J. Mamm. 19: 495, 14 Nov 1938b.

= Perognathus flavescens relictus Goldman, 1938. See Williams 

(1978:54).

Holotype: 150768. [Skin and skull]. Young adult male. Medano 

Springs Ranch, altitude 7,600 ft (2,316 m), 15 mi (24 km) 

northeast of Mosca, San Luis Valley, [Alamosa County], 

southern Colorado. Collected 2 Nov 1907 by M. Cary. 

Original number 1222.

Perognathus bimaculatus Merriam

N. Amer. Fauna 1: 12, 25 Oct 1889e.

= Perognathus flavus bimaculatus Merriam, 1889. See Osgood 

(1900b:24).
5 2 • smithsonian contributions to zoology

Holotype: 8455/23789. Skin and skull. Adult male. Fort Whipple, [Yavapai County], Arizona. Collected 21 May 1865 by 

E. Coues. Original number [SL: 1499].

Perognathus bombycinus Osgood

Proc. Biol. Soc. Washington 20: 19, 23 Feb 1907b.

= Perognathus longimembris bombycinus Osgood, 1907. See 

Nelson and Goldman (1929b:104).

Holotype: 136123. [Skin and skull]. Adult male. Yuma, [Yuma 

County], Arizona. Collected 18 Mar 1905 by E. A. Goldman. Original number [FC: Nelson and Goldman 16844].

Perognathus callistus Osgood

N. Amer. Fauna 18: 28, 20 Sep 1900b.

= Perognathus fasciatus callistus Osgood, 1900. See Jones 

(1953:524).

Holotype: 88245. Skin and skull. Young adult male. Kinney 

Ranch, Green River Basin, near [Williams and Genoways 

(1979:99): about 22 mi (35 km) south] Bitter Creek, Sweetwater County, Wyoming. Collected 14 May 1897 by J. A. 

Loring. Original number 4122.

Perognathus columbianus Merriam

Proc. Acad. Nat. Sci. Philadelphia 46: 263, 27 Sep 1894a. Preprint of Proc. Acad. Nat. Sci. Philadelphia 46: 263, 23 Oct 

1894a.

= Perognathus parvus columbianus Merriam, 1894. See Dalquest (1948:299).

Holotype: 27351/39450. Skin and skull. Young adult male. 

Pasco, Plains of Columbia on east side of Columbia River, 

near mouth of Snake River, [Franklin County], Washington. 

Collected 9 May 1891 by C. P. Streator. Original number 

768.

Perognathus fasciatus Wied-Neuwied

Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. 19: 369, 1839.

= Perognathus fasciatus fasciatus Wied-Neuwied, 1839. See 

Miller (1924:272).

Neotype: 168599. Skin and skull. Adult male. Buford, Williams 

County, North Dakota. Collected 6 May 1910 by H. E. Anthony. Original number [SL: 9].

Neotype designated by Williams and Genoways (1979:95).

Perognathus fasciatus flavescens Merriam

N. Amer. Fauna 1: 11, 25 Oct 1889e.

= Perognathus flavescens flavescens Merriam, 1889. See Osgood 

(1900b:20).

Holotype: [186507]. Skin and skull. Adult male. Kennedy, 

[Cherry County], Nebraska. Collected 11 Jun 1888 by 

V. Bailey. Original number [SL: 23]. Merriam collection 

number 4303/5027.

Perognathus fasciatus litus Cary

Proc. Biol. Soc. Washington 24: 61, 22 Mar 1911.

= Perognathus fasciatus fasciatus Wied-Neuwied, 1839. See Williams and Genoways (1979:95).

Holotype: 160600. [Skin and skull]. Adult female. Sun, [FC: sand 

flat along Sweetwater River], Sweetwater Valley, [Natrona 

County], Wyoming. Collected 18 Sep 1909 by M. Cary. 

Original number 1778.

Perognathus flavescens perniger Osgood

Proc. Biol. Soc. Washington 17: 127, 9 Jun 1904.

Holotype: 57725. [Skin and skull]. Young adult female. Vermillion [Clay County], South Dakota. Collected 22 Aug 1889 

by G. S. Agersborg. No original number.

Perognathus flavus Baird

Proc. Acad. Nat. Sci. Philadelphia 7: 332, reported favorably 

for publication 24 Apr 1855, receipt acknowledged 15 Jun 

1855 by the American Philisophical Society.

= Perognathus flavus flavus Baird, 1855. See Miller (1912:261).

Holotype: [148/1130]. Fragment of skull, skin lost. Age and sex 

unknown. El Paso, [El Paso County], Texas. Collected [MC: 

1851] by J. H. Clark. No original number.

Type not designated by number, but reference to Baird (1857:425) 

reveals this specimen to be the only one that fits Bairds’ 

description.

Perognathus flavus hopiensis Goldman

Proc. Biol. Soc. Washington 45: 89, 21 Jun 1932e.

Holotype: 248014. [Skin and skull]. Adult female. Oraibi, altitude 6,000 ft (1,829 m), Hopi Indian Reservation, Navajo 

County, Arizona. Collected 5 Jun 1927 by G. G. Cantwell. 

Original number 3037, not 3027 as in original description.

Perognathus flavus mexicanus Merriam

Proc. Acad. Nat. Sci. Philadelphia 46: 265, 27 Sep 1894a. Preprint of Proc. Acad. Nat. Sci. Philadelphia 46: 265, 23 Oct 

1894a.

Holotype: 50714. [Skin and skull]. Young adult [SL: male]. Tlalpam, [FC: 7,600 ft (2,316 m)], Valley of Mexico, Federal 

District (Distrito Federal), [Mexico]. Collected 4 Dec 1892 

by E. W. Nelson. Original number 3978.

Perognathus flavus piperi Goldman

Proc. Biol. Soc. Washington 30: 148, 27 Jul 1917b.

Holotype: 168650. [Skin and skull]. Adult male. Twenty-three 

miles (37 km) southwest of Newcastle, [Weston County], 

Wyoming. Collected 25 May 1910 by S. E. Piper. Original 

number 283.

Perognathus flavus sonoriensis Nelson and Goldman

J. Washington Acad. Sci. 24: 267, 15 Jun 1934a.

Holotype: 250885. Skin and skull. Adult female. Costa Rica 

Ranch, lower Sonora River, Sonora, Mexico. Collected 13 

Dec 1932 by V. Bailey and F. Winthrop. Original number 

VB 11282.
number 640 • 5 3

Perognathus fuliginosus Merriam

N. Amer. Fauna 3: 74, 11 Sep 1890b.

= Perognathus flavus fuliginosus Merriam, 1890. See Osgood 

(1900b:25).

Holotype: 17708/24644. Skin and skull. Immature male. Cedar 

belt, northeast of [SL: main peak] San Francisco Mountain 

(Peaks), altitude 7,000 ft (2,134 m), [FC: below Partridge 

Spring, 7,250 ft (2,210 m)], Coconino County, Arizona. 

Collected 4 Oct 1889 by V. Bailey. Original number 559.

Perognathus inornatus Merriam

N. Amer. Fauna 1: 15, 25 Oct 1889e.

= Perognathus inornatus inornatus Merriam, 1889. See Miller 

(1924:277).

Holotype: 13394/23790. Alcoholic, skull removed. Young adult 

male. Fresno [Fresno County], California. Collected, date 

unknown, by [MC: G. Eisen]. Alcoholic cataloged 6 Feb 

1882, skull 3 Jul 1889. No original number.

Perognathus longimembris arizonensis Goldman

Proc. Biol. Soc. Washington 44: 134, 17 Oct 1931a.

Holotype: 250032. [Skin and skull]. Adult female. Ten miles 

(16 km) south of Jacobs Pools, altitude 4,000 ft (1,219 m), 

Houserock Valley, north side of Marble Canyon of Colorado River, [Coconino County], Arizona. Collected 17 Jun 

1931 by E. A. Goldman. Original number 23589.

Perognathus longimembris panamintinus Merriam

Proc. Acad. Nat. Sci. Philadelphia 46: 265, 27 Sep 1894a. Preprint of Proc. Acad. Nat. Sci. Philadelphia 46: 265, 23 Oct 

1894a.

Holotype: 27767/39866. Skin and skull. Young adult male. 

Perognathus Flat [(Grinnell (1922:105): near Bench Mark 

4899, 2 mi (3.2 km) northeast of Harrisburg, on U.S. Geological Survey Ballarat Quadrangle, edition of Mar 1913)], 

[SL: altitude 5,200 ft (1,585 m)], [FC: Emigrant Wash], 

Panamint Mountains, [Inyo County], California. Collected 

16 Apr 1891 by V. Bailey. Original number 2675.

Perognathus merriami gilvus Osgood

N. Amer. Fauna 18: 22, 20 Sep 1900b.

Holotype: 35939/48273. Skin and skull. Adult male. Eddy 

(Carlsbad), [FC: Eddy County], New Mexico. Collected 18 

Sep 1892 by B. H. Dutcher. Original number 329.

Perognathus mollipilosus Coues

Proc. Acad. Nat. Sci. Philadelphia 27: 296, 31 Aug 1875a.

= Perognathus parvus mollipilosus Coues, 1875. See Osgood 

(1900b:36).

Holotype: 7251. Alcoholic. Female, age unknown. Fort Crook, 

[Shasta County], California. Collected by J. Feilner , date 

unknown. No original number. The museum number is incorrect; in the Skin Catalog it is a Scalopus aquaticus, in 

the Bone Catalog a Podiceps. The catalog entry for USNM 

7339 matches the published data for this type, but this specimen cannot be found.

Perognathus monticola Baird

Mammals in Rep. Expl. Surv. 8(1): 422, 1857.

= Perognathus parvus parvus (Peale, 1848). See Miller (1924: 

278).

Holotype: 451/1585. Skin and skull, formerly mounted. [Lyon 

and Osgood (1909:53): Adult] female. West of the Rocky 

Mountains, St Mary’s ? In his list of localities, Baird 

(1857:713) expands the locality to “St. Mary’s Mission, 

Rocky Mountains. A catholic missionary station among the 

Flatheads in Washington Territory, west of Rocky Mountains, on the St. Mary’s Fork of the Bitterroot. Near 46°30′

Latitude 112°00′ Longitude.” Osgood (1900b:36) considered the specimen more likely came from The Dalles, 

[Wasco County], Oregon. Collected in 1853 by G. Suckley. 

No original number.

Perognathus nevadensis Merriam

Proc. Acad. Nat. Sci. Philadelphia 46: 264, 27 Sep 1894a. Preprint of Proc. Acad. Nat. Sci. Philadelphia 46: 264, 23 Oct 

1894a.

= Perognathus longimembris nevadensis Merriam, 1894. See 

Grinnell (1933:147).

Holotype: 54828. [Skin and skull]. Adult male. Halleck, East 

Humboldt Valley, [Elko County], Nevada. Collected 4 Jul 

1893 by V. Bailey. Original number 4070.

Perognathus olivaceus Merriam

N. Amer. Fauna 1: 15, 25 Oct 1889e.

= Perognathus parvus olivaceus Merriam, 1889. See Osgood 

(1900b:37).

Holotype: [186511]. Skin and skull. [SL: Adult] male. Kelton, [FC: foothills 8 mi (13 km) north], [Miller and Rehn 

(1901:159): near north end of Great Salt Lake], [Box Elder 

County], Utah. Collected 24 Oct 1888 by V. Bailey. Original 

number [SL: 338]. Merriam collection number 4876/5623.

Perognathus olivaceus amoenus Merriam

N. Amer. Fauna 1: 16, 25 Oct 1889e.

= Perognathus parvus olivaceus Merriam, 1889. See Osgood 

(1900b:37).

Holotype: [186512]. Skin, skull [and baculum]. [SL: Adult] 

male. Nephi, [FC: 5,200 ft (1,585 m)], [Juab County], Utah. 

Collected 23 Nov 1888, according to original description 

and SL even though FC and MCC say 24 Nov, by V. Bailey. Original number [SL: 404]. Merriam collection number 

5103/5795.

Perognathus pacificus Mearns

Bull. Amer. Mus. Nat. Hist. 10: 299, 31 Aug 1898c.

= Perognathus longimembris pacificus Mearns, 1898. See von 

Bloeker (1932:128).
5 4 • smithsonian contributions to zoology

Holotype: 61022. Skin and skull. Adult female. Edge of Pacific 

Ocean, at the last Mexican Boundary Monument, No. 258, 

[San Diego County, California]. Collected 12 Jul 1894 by 

E. A. Mearns. Original number [FC: Mearns and Holzner] 

3787.

Perognathus panamintinus arenicola Stephens

Proc. Biol. Soc. Washington 13: 153, 13 Jun 1900.

= Perognathus longimembris bangsi Mearns, 1898. See Grinnell 

(1913:329).

Holotype: 99828. [Skin and skull]. [Lyon and Osgood (1909:48): 

Adult] male. San Felipe Narrows, San Diego County, California. Collected 11 Apr 1892 by F. Stephens. Original number 2056. [SL: 2622-X].

Perognathus panamintinus brevinasus Osgood

N. Amer. Fauna 18: 30, 20 Sep 1900b.

= Perognathus longimembris brevinasus Osgood, 1900. See 

Miller (1924:276).

Holotype: [186515]. Skin and skull. Adult female. San Bernardino, [San Bernardino County], California. Collected 

2 May 1885 by F. Stephens. Original number [SL: 49]. Merriam collection number 1109/1661.

Perognathus parvus clarus Goldman

Proc. Biol. Soc. Washington 30: 147, 27 Jul 1917b.

Holotype: 178939. [Skin and skull]. Adult male. Cumberland, 

[Lincoln County], Wyoming. Collected 18 May 1912 by 

S. G. Jewett. Original number 976.

Perognathus parvus idahoensis Goldman

Proc. Biol. Soc. Washington 35: 105, 17 Oct 1922.

Holotype: 236394. [Skin and skull]. Adult male. Echo Crater, 

20 mi (32 km) southwest of Arco, [SL: Snake River Desert], 

Blaine (Butte) County, Idaho. Collected 14 Jun 1921 by L. J. 

Goldman. Original number 2752.

Perognathus parvus magruderensis Osgood

N. Amer. Fauna 18: 38, 20 Sep 1900b.

= Perognathus parvus olivaceus Merriam, 1889. See Hall 

(1946:367).

Holotype: 28427/40531. Skin and skull. Adult male. Mount 

Magruder, altitude 8,000 ft (2,438 m) [FC: 7,900–8,000 ft 

(2,408–2,438 m)], [Esmeralda County], Nevada. Collected 

6 Jun 1891 by V. Bailey. Original number 2899.

Perognathus parvus plerus Goldman

J. Mamm. 20: 352, 14 Aug 1939c.

= Perognathus parvus olivaceus Merriam, 1889. See Durrant 

(1952:241).

Holotype: 263971. [Skin and skull]. Old adult male. North end 

of Stansbury Island, altitude 4,250 ft (1,295 m), Great Salt 

Lake, [Tooele County], Utah. Collected 23 Jun 1938 by 

W. H. Marshall. Original number [SL: 117].

Family Geomyidae Bonaparte, 1845

Genus Cratogeomys Merriam, 1895

Cratogeomys castanops angusticeps Nelson and Goldman

Proc. Biol. Soc. Washington 47: 139, 13 Jun 1934b.

= Cratogeomys castanops castanops (Baird, 1852). See Hafner 

et al. (2008:203).

Holotype: 24503/31908. Skin and skull. Adult male. Eagle Pass, 

[Miller and Kellogg (1955:342): 3,000 ft (914 m)], [Maverick County],Texas. Collected 11 Nov 1890 by C. P. Streator. 

Original number 434.

Cratogeomys castanops consitus Nelson and Goldman

Proc. Biol. Soc. Washington 47: 140, 13 Jun 1934b.

Holotype: 50924. Skin and skull. Young adult male. Gallego, 

altitude 5,500 ft (1,676 m), Chihuahua, Mexico. Collected 

16 Dec 1892 by C. P. Streator. Original number 2416.

Cratogeomys castanops convexus Nelson and Goldman

Proc. Biol. Soc. Washington 47: 142, 13 Jun 1934b.

= Cratogeomys castanops castanops (Baird, 1852). See Hafner 

et al. (2008:203).

Holotype: 127356. Skin and skull. Adult female. Seven miles 

(11 km) east of Las Vacas, Rio Grande Valley, Coahuila, 

Mexico, opposite Del Rio, Texas. Collected 9 Jun 1903 by 

J. H. Gaut. Original number 1469.

Cratogeomys castanops excelsus Nelson and Goldman

Proc. Biol. Soc. Washington 47: 143, 13 Jun 1934b.

= Cratogeomys castanops consitus Nelson and Goldman, 1934. 

See Hafner et al. (2008:203).

Holotype: 246533. Skin and skull. Adult male. San Pedro, 10 mi (16 

km) west of Laguna de Mayran, Coahuila, Mexico. Collected 

28 Mar 1926 by E. A. Goldman. Original number 23561.

Cratogeomys castanops goldmani Merriam

N. Amer. Fauna 8: 160, 31 Jan 1895.

= Cratogeomys goldmani goldmani Merriam, 1895. See Hafner 

et al. (2008: 204).

Holotype: 57965. [Skin and skull]. Young adult female. Canitas, 

Zacatecas, Mexico. Collected 24 Dec 1893 by E. A. Goldman. Original number 286.

Cratogeomys castanops hirtus Nelson and Goldman. 

Proc. Biol. Soc. Washington 47: 138, 13 Jun 1934b.

Holotype: 58325. Skin and skull. Adult male. Albuquerque, altitude 5,000 ft (1,524 m), [Bernalillo County], New Mexico. 

Collected 17 Jan 1894 by J. A. Loring. Original number 1558.

Cratogeomys castanops lacrimalis Nelson and Goldman

Proc. Biol. Soc. Washington 47: 137, 13 Jun 1934b.

= Cratogeomys castanops castanops (Baird, 1852). See Hafner 

et al. (2008:203).
number 640 • 5 5

Holotype: 119071. Skin and skull. Adult male. Roswell, altitude 

3,500 ft (1,069 m) [FC: altitude about 50 ft (15 m) above 

town altitude], Chaves County, New Mexico. Collected 13 

Sep 1902 by J. H. Gaut. Original number 476.

Cratogeomys castanops peridoneus Nelson and Goldman

Proc. Biol. Soc. Washington 47: 148, 13 Jun 1934b.

= Cratogeomys goldmani subnubilus Nelson and Goldman, 

1934. See Hafner et al. (2008:204).

Holotype: 82049. Skin and skull. Adult male. Rio Verde, altitude 

[FC: about] 3,000 ft (914 m), San Luis Potosi, Mexico. Collected 10 Jan 1897 by [E. W.] Nelson and [E. A.] Goldman. 

Original number 10423.

Cratogeomys castanops perplanus Nelson and Goldman

Proc. Biol. Soc. Washington 47: 136, 13 Jun 1934b.

= Cratogeomys castanops castanops (Baird, 1852). See Hafner 

et al. (2008:203).

Holotype: 97171. Skin and skull. Adult male. Tascosa, altitude 

3,000 ft (914 m), Oldham County, Texas. Collected 5 Jun 

1899 by V. Bailey. Original number 6941.

Cratogeomys castanops planifrons Nelson and Goldman

Proc. Biol. Soc. Washington 47: 146, 13 Jun 1934b.

= Cratogeomys goldmani subnubilus Nelson and Goldman, 

1934. See Hafner et al. (2008:204).

Holotype: 93942. Skin and skull. Adult male. Miquihuana, altitude 5,000 ft (1,524 m), southern Tamaulipas, not Nuevo 

Leon as in original description, Mexico. Collected 9 Jun 

1898 by [E. W.] Nelson and [E. A.] Goldman. Original number 12527.

Cratogeomys castanops subnubilus Nelson and Goldman

Proc. Biol. Soc. Washington 47: 145, 13 Jun 1934b.

= Cratogeomys goldmani subnubilus Nelson and Goldman, 

1934. See Hafner et al. (2008:204).

Holotype: 79482. Skin and skull. Adult male. Carneros, altitude 

6,800 ft (2,073 m), Coahuila, Mexico. Collected 12 Aug 

1896 by [E. W.] Nelson and [E. A.] Goldman. Original number 10018.

Cratogeomys castanops subsimus Nelson and Goldman

Proc. Biol. Soc. Washington 47: 144, 13 Jun 1934b.

= Cratogeomys castanops consitus Nelson and Goldman, 1934. 

See Hafner et al. (2008:203).

Holotype: 51048. Skin and skull. Adult male. Jaral (San Antonio 

de Jaral), southeastern Coahuila, Mexico. Collected 14 Jan 

1893 by C. P. Streator. Original number 2555.

Cratogeomys castanops tamaulipensis Nelson and Goldman

Proc. Biol. Soc. Washington 47: 141, 13 Jun 1934b.

Holotype: 116535. Skin and skull. Adult female. Matamoras, 

Tamaulipas, Mexico. Collected 8 Feb 1902 by [E. W.] Nelson and [E. A.] Goldman. Original number 14885.

Cratogeomys estor Merriam

N. Amer. Fauna 8: 155, 31 Jan 1895.

= Cratogeomys perotensis Merriam, 1895. See Hafner et al. 

(2005:1103).

Holotype: 54308. [Skin and skull]. Adult male. Las Vigas, altitude 8,000 ft (2,438 m), Veracruz, [Mexico]. Collected 12 

Jun 1893 by E. W. Nelson. Original number 5005.

Cratogeomys fulvescens Merriam

N. Amer. Fauna 8: 161, 31 Jan 1895. See Hafner et al. 

(2005:1103) for current usage.

Holotype: 58168. [Skin and skull]. Adult male. Chalchicomula, 

[FC: 8,200 ft (2,499 m)], Puebla, Mexico. Collected 15 Jan 

1894 by E. W. Nelson and E. A. Goldman. Original number 

5651.

Cratogeomys fulvescens subluteus Nelson and Goldman

Proc. Biol. Soc. Washington 47: 152, 13 Jun 1934b.

= Cratogeomys fulvescens Merriam, 1895. See Hafner et al. 

(2005:1103).

Holotype: 54300. Skin and skull. Adult female, not male as in 

original description. Perote, altitude 7,800 ft (2,377 m), Veracruz, Mexico. Collected 3 Jun 1893 by [E. W.] Nelson and 

[E. A.] Goldman. Original number [FC: EWN] 4929.

Cratogeomys merriami irolonis Nelson and Goldman

Proc. Biol. Soc. Washington 47: 150, 13 Jun 1934b.

= Cratogeomys perotensis Merriam, 1895. See Hafner et al. 

(2005:1103).

Holotype: 53494. Skin and skull. Adult female. Irolo, altitude 

7,600 ft (2,316 m), Hidalgo, Mexico. Collected 30 Mar 

1893 by [E. W.] Nelson and [E. A.] Goldman. Original number [FC: EWN] 4520.

Cratogeomys merriami saccharalis Nelson and Goldman

Proc. Biol. Soc. Washington 47: 149, 13 Jun 1934b.

= Cratogeomys merriami (Thomas, 1893). See Hafner et al. 

(2005:1103).

Holotype: 55347. Skin and skull. Adult male. Atlixco, altitude 

5,400 ft (1,646 m), Puebla, Mexico. Collected 29 Jul 1893 by 

[E. W.] Nelson and [E. A.] Goldman. Original number 5279.

Cratogeomys oreocetes Merriam

N. Amer. Fauna 8: 156, 31 Jan 1895.

= Cratogeomys merriami (Thomas, 1893). See Hafner et al. 

(2005:1103).

Holotype: 57963. [Skin and skull]. Young adult female. Mount 

Popocatepetl, altitude 11,000 ft (3,353 m), [Mexico], 

Mexico. Collected 7 Jan 1894 by E. W. Nelson and E. A. 

Goldman. Original number 5647, not 47 as in original description.

Cratogeomys peregrinus Merriam

N. Amer. Fauna 8: 158, 31 Jan 1895.
5 6 • smithsonian contributions to zoology

= Cratogeomys merriami (Thomas, 1893). See Hafner et al. 

(2005:1103).

Holotype: 57964. [Skin and skull]. Old female. Mount Iztaccihuatl, altitude 11,500 ft (3,505 m), [Mexico], Mexico. 

Collected 9 Jan 1894 by E. W. Nelson and E. A. Goldman. 

Original number 5650, not 50 as in original description.

Cratogeomys perotensis Merriam

N. Amer. Fauna 8: 154, 31 Jan 1895. See Hafner et al. (2005: 

1103) for current usage.

Holotype: 54299. [Skin and skull]. Adult female. Cofre de Perote, 

altitude 9,500 ft (2,896 m), Veracruz, Mexico. Collected 28 

May 1893 by E. W. Nelson. Original number 4889.

Geomys clarkii Baird

Proc. Acad. Nat. Sci. Philadelphia 7: 332, reported favorably for 

publication 24 Apr 1855b, receipt acknowledged 15 Jun 

1855 by the American Philosophical Society.

= Cratogeomys castanops castanops (Baird, 1852). See Hafner 

et al. (2008:203).

Holotype: [6/1624. Skin, skull, and partial skeleton]. Presidio 

del Norte, on the Rio Grande, [Chihuahua, Mexico]. Nelson and Goldman (1934b:140) add “at or near the present 

town of Ojinaga.” Collected by J. H. Clark; date unknown. 

Skin cataloged 12 Feb 1852; skull 15 Feb 1855. Type not 

designated by number in original description, but only one 

specimen, a skin and skull, is mentioned. Reference to Baird 

(1857:384) shows that he had two specimens from Presidio 

de Norte collected by Clark; a skull only and the one given 

above whose measurements match those in the original 

description.

Geomys fumosus Merriam

Proc. Biol. Soc. Washington 7: 165, 29 Sep 1892b.

= Cratogeomys fumosus fumosus (Merriam, 1892). See Hooper 

(1948:302).

Holotype: 33202/45207. Skin and skull. Adult male. Colima 

City, [Colima], Mexico. Restricted to 3 mi (4.8 km) west of 

Colima City, altitude 1,700 ft (518 m), Colima, Mexico by 

Miller and Kellogg (1955:346). Collected 27 Mar 1892 by 

E. W. Nelson. Original number 2338.

Geomys gymnurus Merriam

Proc. Biol. Soc. Washington 7: 166, 29 Sep 1892b.

= Cratogeomys fumosus fumosus (Merriam, 1892). See Hafner 

et al. (2004:1177).

Holotype: 33579/45614. Skin and skull. Adult female. Zapotlan, [FC: about 400 ft (122 m)], Jalisco, Mexico. Collected 

16 Apr 1892 by E. W. Nelson. Original number 2460.

Platygeomys gymnurus inclarus Goldman

J. Mamm. 20: 88, 14 Feb 1939d.

= Cratogeomys fumosus fumosus (Merriam, 1892). See Hafner 

et al. (2004:1177).

Holotype: 33582/45617. Skin and skull. Young female. North 

slope Sierra Nevada de Colima, altitude 10,000 ft (3,048 

m), Jalisco, Mexico. Collected 22 Apr 1892 by [E. W.] 

Nelson and [E. A.] Goldman. Original number [FC: EWN] 

2532.

Platygeomys gymnurus imparilis Goldman

J. Mamm. 20: 89, 14 Feb 1939d.

= Cratogeomys fumosus imparilis (Goldman, 1939). See Hafner 

et al. (2004:1178).

Holotype: 34918/47183. Skin and skull. Adult male. Patzcuaro, altitude [FC: about] 7,000 ft (2,134 m), Michoacan, 

Mexico. Collected 23 Jul 1892 by [E. W.] Nelson and [E. A.] 

Goldman. Original number [FC: EWN] 2918.

Platygeomys neglectus Merriam

Proc. Biol. Soc. Washington 15: 68, 22 Mar 1902a.

= Cratogeomys fumosus tylorhinus (Merriam, 1895). See Hafner 

et al. (2004:1178).

Holotype: 81218. [Skin and skull]. Adult male. Cerro de la 

Calentura, altitude 9,000 ft (2,743 m) [FC: about 9,500 ft 

(2,896 m)], about 8 mi (13 km) northwest of Pinal de Amoles, Queretaro, Mexico. Collected 14, not 4 as in original 

description, Sep 1896 by E. A. Goldman. Original number 

[FC: Nelson and Goldman] 10142.

Platygeomys planiceps Merriam

N. Amer. Fauna 8: 168, 31 Jan 1895.

= Cratogeomys planiceps (Merriam, 1895). See Hafner et al. 

(2004:1178).

Holotype: 55906. Skin and skull. Adult male. North slope Volcan Toluca, [FC: 9,000 ft (2,743 m)], [Mexico, Mexico]. 

Collected 12 Sep 1893 by E. W. Nelson. Original number 

5466.

Platygeomys tylorhinus Merriam

N. Amer. Fauna 8: 167, 31 Jan 1895.

= Cratogeomys fumosus tylorhinus (Merriam, 1895). See Hafner 

et al. (2004:1178).

Holotype: 51883. [Skin and skull]. Adult male. Tula, [FC: 6,800 

ft (2,073 m)], Hidalgo, Mexico. Collected 13 Mar 1893 by 

E. W. Nelson. Original number 4442.

Platygeomys tylorhinus angustirostris Merriam

Proc. Biol. Soc. Washington 16: 81, 29 May 1903a.

= Cratogeomys fumosus angustirostris (Merriam, 1903). See 

Hafner et al. (2004:1177).

Holotype: 125688. [Skin and skull]. Adult female. Patamban, 

[FC: 10,000 ft (3,048 m)], Michoacan, Mexico. Collected 

2 Feb 1903 by [E. W.] Nelson and [E. A.] Goldman. Original number 15850.

Platygeomys varius Goldman

J. Mamm. 20: 90, 14 Feb 1939d.
number 640 • 5 7

= Cratogeomys fumosus angustirostris (Merriam, 1903). See 

Hafner et al. (2004:1177).

Holotype: 126144. Skin and skull. Adult male. Uruapan, altitude about 6,000 ft (1,829 m), Michoacan, Mexico. Collected 8 Mar 1903 by [E. W.] Nelson and [E. A.] Goldman. 

Original number 16105.

Platygeomys zinseri Goldman

J. Mamm. 20: 91, 14 Feb 1939d.

= Cratogeomys fumosus angustirostris (Merriam, 1903). See 

Hafner et al. (2004:1177).

Holotype: 78971. Skin and skull. Adult male. Lagos, altitude 

6,150 ft (1,875 m), Jalisco, Mexico. Collected 29 Jun 1896 

by [E. W.] Nelson and [E. A.] Goldman. Original number 

9718.

Pseudostoma castanops Baird

In Stansbury, Expl. Surv. Valley Great Salt Lake, Utah, App. C., 

Jun 1852: p. 313.

= Cratogeomys castanops castanops (Baird, 1852). See Miller 

(1912:247).

Holotype: 4007/3861. Skin and skull. Unknown age and sex. 

Along the prairie road to Bent’s Fort. [Merriam (1895:159): 

on the Arkansas River near the present town of Las Animas, 

Bent County, Colorado]. Collected by [Baird (1857:385): 

J. W.] Abert, date unknown. [Cataloged 18 Jun 1860].

Type not designated by number, but Baird (1857:385) states there 

was only one specimen collected by Abert at this locality.

Genus Geomys Rafinesque, 1817

Geomys arenarius Merriam

N. Amer. Fauna 8: 139, 31 Jan 1895.

= Geomys arenarius arenarius Merriam, 1895. See Davis (1940:22).

Holotype: 18117/25015. Skin and skull. Adult male. El Paso, 

[El Paso County], Texas. Collected 14, not 13 as in original 

description, Dec 1899 by V. Bailey. Original number 798.

Geomys breviceps Baird

Proc. Acad. Nat. Sci. Philadelphia 7: 335, reported favorably for 

publication 24 Apr 1855a, receipt acknowledged 15 Jun 

1855 by the American Philisophical Society.

= Geomys breviceps breviceps Baird, 1855. See Davis (1938:488).

Holotype: 156/1138. Skin and skull. Adult female. [SL: Prairie 

Mer Rouge], Morehouse Parish, Louisiana. Collected [SL: 

1852] by J. Fairie. No original number.

Type not designated by number in original description, but measurements therein can only be associated with those given 

for 156 in tables on pages 379 and 380 in Baird, 1857.

Geomys breviceps ammophilus Davis

Bull. Texas Agr. Exp. Stat. 590: 16, 23 Oct 1940.

= Geomys attwateri ammophilus Davis, 1940. See Patton 

(2005b:862).

Holotype: 97010. Skin and skull. Adult female. Cuero, De Witt 

County, Texas. Collected 26 Apr 1899 by V. Bailey. Original 

number 6841.

Geomys breviceps attwateri Merriam

N. Amer. Fauna 8: 135, 31 Jan 1895.

= Geomys attwateri attwateri Merriam, 1895. See Patton 

(2005b:862).

Holotype: 51382. [Skin and skull]. Adult male. Rockport, Aransas County, Texas. Collected 18 Nov 1892 by H. H. Keays. 

Original number 36.

Geomys breviceps dutcheri Davis

Bull. Texas Agr. Exp. Stat. 590: 12, 23 Oct 1940.

= Geomys breviceps sagittalis Merriam, 1895. See Heaney and 

Timm (1983:18).

Holotype: 64591. Skin and skull. Adult female. Fort Gibson, 

Muskogee County, Oklahoma. Collected 27 Apr 1894 by 

B. H. Dutcher. Original number 584.

Geomys breviceps llanensis V. Bailey

N. Amer. Fauna 25: 129, 24 Oct 1905.

= Geomys texensis llanensis V. Bailey, 1905. See Jolley et al. 

(2000:1026).

Holotype: 97086. [Skin and skull]. Adult male. Llano, [Llano 

County], Texas. Collected 15 May 1899 by V. Bailey. Original number 6912.

Geomys breviceps sagittalis Merriam

N. Amer. Fauna 8: 134, 31 Jan 1895.

Holotype: 32936/44957. Skin and skull. Adult male. Clear 

Creek, Galveston Bay, [SL: Galveston County], Texas. Collected 28 Mar 1892 by W. Lloyd. Original number 1181.

Geomys bursarius lutescens Merriam

N. Amer. Fauna 4: 51, 8 Oct 1890f.

= Geomys lutescens Merriam, 1890. For specific status of G. lutescens see Sudman et al. (2006:668) and Chambers et al. 

(2009:537).

Holotype: 16677/23595. Skin and skull. Adult male, not female 

as in original description. Sandhills on Birdwood Creek, 

Lincoln County, Nebraska. Collected 27 May 1889 by A. B. 

Baker. Original number [SL: 11].

Geomys bursarius wisconsinensis Jackson

Proc. Biol. Soc. Washington 70: 33, 28 Jun 1957.

Holotype: 243005. Skin and skull. Adult female. Lone Rock, 

Richland County, Wisconsin. Collected 13 Sep 1922 by 

H. H. T. Jackson. Original number 1604.

Geomys personatus True

Proc. U.S. Natl. Mus. 11: 159, 5 Jan 1889.

= Geomys personatus personatus True, 1889. See Davis 

(1940:26).
5 8 • smithsonian contributions to zoology

Syntypes: 19667/37999. Skin and skull. [Lyon and Osgood 

(1909: 71: Adult] male.

19668/38000. Skin and skull. [Lyon and Osgood (1909:71: 

Adult] female.

Padre Island, [Cameron County], Texas. Collected 11 Apr 1888, 

collector unknown. Purchased from C. K. Worthen. No 

type designated by number in original description. Measurements and sex are given for two specimens, and these are the 

only specimens in the collection that are from 11 Apr 1888. 

Their measurements match those in the original description 

as well.

Geomys personatus fallax Merriam

N. Amer. Fauna 8: 144, 31 Jan 1895.

Holotype: 32031/43845. Skin and skull. Adult male. South side 

of Nueces Bay, [Nueces County], Texas. Collected 30 Nov 

1891 by W. Lloyd. Original number 949.

Geomys personatus tropicalis Goldman

Proc. Biol. Soc. Washington 28: 134, 29 Jun 1915.

= Geomys tropicalis Goldman, 1915. See Alvarez (1963:426).

Holotype: 92946. [Skin and skull]. Adult male. Alta Mira, [FC: 

about 75 ft (23 m)], Tamaulipas, Mexico. Collected 18 Apr 

1898 by E. A. Goldman. Original number [FC: Nelson and 

Goldman] 12320.

Geomys texensis Merriam

N. Amer. Fauna 8: 137, 31 Jan 1895.

= Geomys texensis texensis Merriam, 1895. See Smolen et al. 

(1993:10).

Holotype: [186502]. Skin and skull. Adult female. Mason, 

Mason County, Texas. Collected 17 Dec 1885 by I. B. 

Henry. Original number [MCC: 27]. Merriam collection 

number 1690/2259.

Geomys tuza mobilensis Merriam

N. Amer. Fauna 8: 119, 31 Jan 1895.

= Geomys pinetis mobilensis Merriam, 1895. See Harper 

(1952:37). See Sudman et al. (2006:673) and Chambers et 

al. (2009:543) for use of G. p. mobilensis.

Holotype: 33988/46023. Skin and skull. Adult male. [FC: Point 

Clear], Mobile Bay, [Baldwin County], Alabama. Collected 

26 Apr 1892 by R. J. Thompson. Original number 50.

Genus Orthogeomys Merriam, 1895

Heterogeomys hispidus chiapensis Nelson and Goldman

Proc. Biol. Soc. Washington 42: 151, 30 Mar 1929c.

= Orthogeomys hispidus chiapensis (Nelson and Goldman, 

1929). See Russell (1968:531).

Holotype: 76737. [Skin and skull]. Adult female. Tenejapa, altitude 7,800 ft (2,377 m), about 16 mi (26 km) northeast of 

San Cristobal, Chiapas, Mexico. Collected 28 Nov 1895 by 

E. W. Nelson and E. A. Goldman. Original number 8682.

Heterogeomys hispidus concavus Nelson and Goldman

Proc. Biol. Soc. Washington 42: 148, 30 Mar 1929c.

= Orthogeomys hispidus concavus (Nelson and Goldman, 1929). 

See Russell (1968:531).

Holotype: 81227. [Skin and skull]. Adult male. Pinal de Amoles, 

altitude [SL: about] 5,500 ft (1,676 m), Queretaro, Mexico. 

Collected 9 Sep 1896 by E. W. Nelson and E. A. Goldman. 

Original number 10130.

Heterogeomys hispidus isthmicus Nelson and Goldman

Proc. Biol. Soc. Washington 42: 149, 30 Mar 1929c.

= Orthogeomys hispidus isthmicus (Nelson and Goldman, 

1929). See Hall (1981:511).

Holotype: 78062. [Skin and skull]. Adult male. Jaltipan, altitude 

100 ft (30 m), Veracruz, Mexico. Collected 2 May 1896 by 

E. W. Nelson and E. A. Goldman. Original number 9574.

Heterogeomys hispidus teapensis Goldman

J. Washington Acad. Sci. 29: 176, 15 Apr 1939e.

= Orthogeomys hispidus teapensis (Goldman, 1939). See Hall 

(1981:511).

Holotype: 100035. Skin and skull. Adult female. Teapa, Tabasco, Mexico. Collected 23 Mar 1900 by [E. W.] Nelson 

and [E. A.] Goldman. Original number 14056.

Heterogeomys hispidus tehuantepecus Goldman

J. Washington Acad. Sci. 29: 175, 15 Apr 1939e.

= Orthogeomys hispidus tehuantepecus (Goldman, 1939). See 

Russell (1968:531).

Holotype: 73434. Skin and skull. Adult male. Mountains 12 mi 

(19 km) northwest of Santo Domingo and about 60 mi (97 

km) north of Tehuantepec City, altitude 1,600 ft (488 m), 

Oaxaca, Mexico. Collected 18 Jun 1895 by [E. W.] Nelson 

and [E. A.] Goldman. Original number 8092.

Heterogeomys hispidus yucatanensis Nelson and Goldman

Proc. Biol. Soc. Washington 42: 150, 30 Mar 1929c.

= Orthogeomys hispidus yucatanensis (Nelson and Goldman, 

1929). See Russell (1968:531).

Holotype: 100344. [Skin and skull]. Adult male. Campeche, altitude 50 ft (15 m), Campeche, Mexico. Collected 7 Jun 1900 

by E. W. Nelson and E. A. Goldman. Original number 14233.

Heterogeomys torridus Merriam

N. Amer. Fauna 8: 183, 31 Jan 1895.

= Orthogeomys hispidus torridus (Merriam, 1895). See Russell 

(1968:531).

Holotype: 63629. [Skin and skull]. Adult female. Chichicaxtle, 

Veracruz, Mexico. Collected 15 Feb 1894 by E. W. Nelson 

[SL: and E. A. Goldman]. Original number [FC: Nelson and 

Goldman] 5850.

Macrogeomys costaricensis Merriam

N. Amer. Fauna 8: 192, 31 Jan 1895.
number 640 • 5 9

= Orthogeomys cherriei costaricensis (Merriam, 1895). See Russell (1968:532).

Holotype: 12911/22551. Skin and skull. Unsexed juvenile. Pacuare, [Limon], Costa Rica. Collected 1876 by J. Cooper. 

Original number 96.

Macrogeomys dariensis Goldman

Smithsonian Misc. Coll. 60 (2): 8, 20 Sep 1912b.

= Orthogeomys dariensis (Goldman, 1912). See Russell (1968:532).

Holotype: 179587. [Skin and skull]. Adult male. Cana, altitude 

2,000 ft (610 m), [Darien], in the mountains of eastern Panama. Collected 31 May 1912 by E. A. Goldman. Original 

number 21760.

Macrogeomys dolichocephalus Merriam

N. Amer. Fauna 8: 189, 31 Jan 1895.

= Orthogeomys heterodus dolichocephalus (Merriam, 1895). 

See Russell (1968:532).

Holotype: 8627/36295. Skin and skull. Adult male. San Jose, 

[San Jose], Costa Rica. Collected Jan 1866 by J. C. Zeledon. No original number. Goodwin (1946:377) says the 

type probably came from Zarcero or Palmira.

Orthogeomys grandis alleni Nelson and Goldman

J. Mamm. 11: 156, 9 May 1930b.

Holotype: 70586. [Skin and skull]. Adult male. Near Acapulco, 

altitude 2,000 ft (610 m), Guerrero, Mexico. Collected 21 

Jan 1895 by [E. W.] Nelson and [E. A.] Goldman. Original 

number 7409.

Orthogeomys grandis annexus Nelson and Goldman

Proc. Biol. Soc. Washington 46: 195, 26 Oct 1933.

Holotype: 75949. Skin and skull. Adult female. Tuxtla Gutierrez, altitude 2,600 ft (792 m), Chiapas, Mexico. Collected 

11 Sep 1895 by [E. W.] Nelson and [E. A.] Goldman. Original number 8418.

Orthogeomys grandis felipensis Nelson and Goldman

J. Mamm. 11: 157, 9 May 1930b.

Holotype: 67030. [Skin and skull]. Adult male. Cerro San Felipe, 

altitude 10,000 ft (3,048 m), 10 mi (16 km) north of Oaxaca, Oaxaca, Mexico. Collected 28 Aug 1894 by [E. W.] 

Nelson and [E. A.] Goldman. Original number 6669.

Orthogeomys grandis guerrerensis Nelson and Goldman

J. Mamm. 11: 158, 9 May 1930b.

Holotype: 126536. [Skin and skull]. Adult female. El Limon, in 

the valley of the Rio de las Balsas, about 20 mi (32 km) 

northwest of La Union, Guerrero, [Mexico]. Collected 

2 Apr 1903 by [E. W.] Nelson and [E. A.] Goldman. Original number 16286.

Orthogeomys grandis vulcani Nelson and Goldman

Proc. Biol. Soc. Washington 44: 105, 17 Oct 1931.

Holotype: 76745. [Skin and skull]. Adult male. Volcan Santa 

Maria, altitude 9,000 ft (2,743 m), Quezaltenango, Guatemala. Collected 28 Jan 1896 by [E. W.] Nelson and [E. A.] 

Goldman. Original number 9243.

Orthogeomys latifrons Merriam

N. Amer. Fauna 8: 178, 31 Jan 1895.

= Orthogeomys grandis latifrons Merriam, 1895. See Nelson 

and Goldman (1930b:156).

Holotype: [61239. Skin and skull. Adult, sex unknown]. Guatemala, 

exact locality unknown. [Nelson and Goldman (1930b:156): 

probably from the lowlands near the Pacific coast]. [Date collected and collector unknown; cataloged 19 Nov 1894]. Original number 2, World’s Fair exhibit of Guatemala.

Orthogeomys nelsoni Merriam

N. Amer. Fauna 8: 176, 31 Jan 1895.

= Orthogeomys grandis nelsoni Merriam, 1895. See Nelson and 

Goldman (1930b:156).

Holotype: 66751. [Skin and skull]. Adult male. Mount Zempoaltepec, altitude 8,000 ft (2,438 m), Oaxaca, Mexico. 

Collected 8 Jul 1894 by E. W. Nelson and E. A. Goldman. 

Original number 6376.

Genus Pappogeomys Merriam, 1895

Geomys nelsoni Merriam

Proc. Biol. Soc. Washington 7: 164, 29 Sep 1892b.

= Pappogeomys bulleri bulleri (Thomas, 1892). See Hall (1981:516).

Holotype: 33587/45622. Skin and skull. Old male. North slope 

Sierra Nevada de Colima, altitude 1,908 m or 6,500 ft, 

Jalisco, Mexico. Collected 11 Apr 1892 by E. W. Nelson. 

Original number 2436.

Pappogeomys albinasus Merriam

N. Amer. Fauna 8: 149, 31 Jan 1895.

= Pappogeomys bulleri albinasus Merriam, 1895. See Goldman 

(1939f:94).

Holotype: 34138/46215. Skin and skull. Adult male [originally 

processed as female]. Altemajac, [FC: about 4,500 ft (1,372 

m)], a suburb of Guadalajara, Jalisco, Mexico. Collected 21 

May 1892 by E. W. Nelson. Original number 2654.

Pappogeomys bulleri amecensis Goldman

J. Mamm. 20: 97, 14 Feb 1939f.

= Pappogeomys bulleri bulleri (Thomas, 1892). See Hafner et al. 

(2009:54).

Holotype: 82189. Skin and skull. Adult male. Mountains near 

Ameca, altitude 6,500 ft (1,981 m), Jalisco, Mexico. Collected 20 Feb 1897 by [E. W.] Nelson and [E. A.] Goldman. 

Original number 10554.

Pappogeomys bulleri flammeus Goldman

J. Mamm. 20: 95, 14 Feb 1939f.
6 0 • smithsonian contributions to zoology

= Pappogeomys bulleri bulleri (Thomas, 1892). See Hall (1981: 

516).

Holotype: 88117. Skin and skull. Adult male. Milpillas, 5 mi 

(8 km) southwest of San Sebastian, altitude 3,800 ft (1,158 

m), northwestern Jalisco, Mexico. Collected 15 Mar 1897 

by [E. W.] Nelson and [E. A.] Goldman. Original number 

10716.

Pappogeomys bulleri lagunensis Goldman

J. Mamm. 20: 96, 14 Feb 1939f.

= Pappogeomys bulleri bulleri (Thomas, 1892). See Hall (1981: 

516).

Holotype: 88121. Skin and skull. Adult male. La Laguna, altitude 6,500 ft (1,981 m), Sierra de Juanacatlan, northwestern 

Jalisco, Mexico. Collected 24 Mar 1897 by [E. W.] Nelson 

and [E. A.] Goldman. Original number 10802.

Pappogeomys bulleri nayaritensis Goldman

J. Mamm. 20: 94, 14 Feb 1939f.

Holotype: 88124. Skin and skull. Adult male. Jalisco, about 

10 mi (16 km) south of Tepic, altitude 5,000 ft (1,524 m), 

Nayarit, Mexico. Collected 10 Apr 1897 by [E. W.] Nelson 

and [E. A.] Goldman. Original number 10886.

Genus Thomomys Wied-Neuwied, 1839

Geomys fulvus Woodhouse

Proc. Acad. Nat. Sci. Philadelphia 6: 201, presented at meeting 

of 28 Dec 1852c, receipt acknowledged 7 Feb 1853 by the 

Smithsonian Institution.

= Thomomys bottae fulvus (Woodhouse, 1852). See Goldman 

(1935a:156).

Holotype: [2674. Mounted skin, no skull. Sex and age unknown]. 

San Francisco Mountain (Peaks), [Coconino County], New 

Mexico (Arizona). Collected [Poole and Schantz (1942:306): 

October] [Baird (1857:403) 1851] by S. W. Woodhouse. 

No original number. Type not designated by number in 

original description. Woodhouse based his description on a 

single specimen, and comparing his data with that of Baird 

(1857:403) shows the above specimen to be the one.

Thomomys alexandrae Goldman

J. Washington Acad. Sci. 23: 464, 15 Oct 1933b.

= Thomomys bottae alexandrae Goldman, 1933. See Benson 

(1935:449).

Holotype: 250969. Skin and skull. Adult male. Plain 5 mi (8 km) 

southeast of Rainbow Lodge, altitude 6,200 ft (1,890 m), 

near Navajo Mountain, Coconino County, Arizona. Collected 16 Jun 1933 by E. A. Goldman. Original number 

23613.

Thomomys alpinus Merriam

Proc. Biol. Soc. Washington 11: 216, 15 Jul 1897c.

= Thomomys bottae alpinus Merriam, 1897. See Goldman 

(1935a:154).

Holotype: 30528/42512. Skin and skull. Adult male. [FC: Head 

of] Big Cottonwood Meadows, 8 mi (13 km) southeast of 

Mount Whitney Peak, altitude 10,000 ft (3,048 m), [Inyo 

County], California. Collected 6 Aug 1891 by B. H. Dutcher. 

Original number 167.

Thomomys alpinus awahnee Merriam

Proc. Biol. Soc. Washington 21: 146, 9 Jun 1908b.

= Thomomys bottae awahnee Merriam, 1908. See Goldman 

(1935a:154).

Holotype: 133076. [Skin and skull]. Adult female. Yosemite Valley, [Miller and Kellogg (1955:278): 4,000 ft (1,219 m) near 

Old Sentinel Hotel], [Mariposa County], California. Collected 14 Jun 1904 by N. Hollister. Original number 870.

Thomomys angularis Merriam

Proc. Biol. Soc. Washington 11: 214, 15 Jul 1897c.

= Thomomys bottae bottae (Eydoux and Gervais, 1836). See 

Patton and Smith (1990:112).

Holotype: 58123. [Skin and skull]. Adult male. Los Banos, 

Merced County, California. Collected 1 Jan 1894 by J. E. 

McLellan. Original number 418.

Thomomys angularis pascalis Merriam

Proc. Biol. Soc. Washington 14: 111, 19 Jul 1901d.

= Thomomys bottae pascalis Merriam, 1901. See V. Bailey (1915:54).

Holotype: 32779/44702. Skin and skull. Adult male. Fresno, 

[FC: plain east of county hospital], San Joaquin Valley, 

[Fresno County], California. Collected 4 Mar, not May 

as in original description, 1892 by C. P. Streator. Original 

number 1634.

Thomomys apache V. Bailey

Proc. Biol. Soc. Washington 23: 79, 4 May 1910.

= Thomomys bottae aureus J. A. Allen, 1893. See Youngman 

(1958:370).

Holotype: 135366. [Skin and skull]. Adult male. Lake La Jara, 

altitude 7,500 ft (2,286 m), on the Jicarilla Apache Indian 

Reservation, [Rio Arriba County], New Mexico. Collected 

19 Sep 1904 by J. H. Gaut. Original number 3289.

Thomomys aureus lachuguilla V. Bailey

Proc. Biol. Soc. Washington 15: 120, 2 Jun 1902.

= Thomomys bottae lachuguilla V. Bailey, 1902. See Goldman 

(1938a:55).

Holotype: 110336. [Skin, skull, and baculum]. Adult male. Arid 

foothills near El Paso, [FC: Franklin Mountains], [Miller 

and Kellogg (1955:305): dry wash, 1 mi (1.6 km) northeast] 

El Paso, [El Paso County], Texas. Collected 24 Sep 1901 by 

V. Bailey. Original number 7858.

Thomomys aureus perpes Merriam

Proc. Biol. Soc. Washington 14: 111, 19 Jul 1901d.

= Thomomys bottae bottae (Eydoux and Gervais, 1836). See 

Patton and Smith (1990:112).
number 640 • 6 1

Holotype: 25091/32503. Skin and skull. Adult male. Lone Pine, 

Owens Valley, [Inyo County], California. Collected 23 Dec 

1890 by E. W. Nelson. Original number 145.

Thomomys aureus pervagus Merriam

Proc. Biol. Soc. Washington 14: 110, 19 Jul 1901d.

= Thomomys bottae pervagus Merriam, 1901. See Goldman 

(1935a:157).

Holotype: 58293. [Skin and skull]. Adult male. Espanola, [Rio 

Arriba County], New Mexico. Collected 4 Jan 1894 by J. A. 

Loring. Original number 1548.

Thomomys baileyi Merriam

Proc. Biol. Soc. Washington 14: 109, 19 Jul 1901d.

= Thomomys bottae baileyi Merriam, 1901. See Anderson 

(1966:195).

Holotype: 18256/25159. Skin and skull. Adult female. [FC: On 

plain east of] Sierra Blanca, [Miller and Kellogg (1955:309): 

railway station at junction of Texas Pacific and Southern 

Pacific Railroads], [Hudspeth County], Texas. Collected 28 

Dec 1889 by V. Bailey. Original number 870.

Thomomys baileyi spatiosus Goldman

Proc. Biol. Soc. Washington 51: 58, 18 Mar 1938a.

= Thomomys bottae spatiosus Goldman, 1938. See Anderson 

(1966:195). 

Holotype: 100427. Skin and skull. Adult male. Alpine, altitude 

4,500 ft (1,372 m), Brewster County, Texas. Collected 26 

May 1900 by V. Bailey. Original number 7368.

Thomomys bottae actuosus Kelson

Univ. Kansas Publ., Mus. Nat. Hist. 5: 67, 1 Oct 1951b.

Holotype: 119053. Skin and skull. Adult male. Corona, [FC: 

6,666 ft (2,032 m)], Lincoln County, New Mexico. Collected 13 Oct 1902 by J. H. Gaut. Original number 591.

Thomomys bottae alienus Goldman

J. Washington Acad. Sci. 28: 338, 15 Jul 1938c.

= Thomomys bottae mearnsi V. Bailey, 1914. See Hoffmeister 

(1986:233).

Holotype: 261926. Skin and skull. Adult male. Mammoth, altitude 2,400 ft (732 m), [SL: east side] San Pedro River, Pinal 

County, Arizona. Collected 4 Nov 1936 by E. A. Goldman. 

Original number 23746.

Thomomys bottae birdseyei Goldman

Proc. Biol. Soc. Washington 50: 134, 10 Sep 1937d.

Holotype: 161654. Skin and skull. Adult male. Pine Valley 

Mountains, altitude 8,300 ft (2,530 m), 5 mi (8 km) [FC: 4 

mi (6 km)] east of Pine Valley, Washington County, Utah. 

Collected 10 Oct 1909 by C. Birdseye. Original number 

861.

Thomomys bottae convergens Nelson and Goldman

J. Mamm. 15: 123, 15 May 1934c.

Holotype: 250894. Skin and skull. Adult male. Costa Rica Ranch, 

delta of Sonora River, [Miller and Kellogg (1955:308): 

about 40 mi (64 km) west] southwest of Hermosillo, Sonora, Mexico. Collected 13 Dec 1932 by V. Bailey and F. 

Winthrop. Original number [FC: VB] 11280.

Thomomys bottae cultellus Kelson

Univ. Kansas Publ., Mus. Nat. Hist. 5: 64, 1 Oct 1951b.

Holotype: 70919. Skin and skull. Adult male. Halls Peak, Mora 

County, New Mexico. Obtained 13 Jan 1895 by G. Barber. 

Original number [SL: 56].

Thomomys bottae desitus Goldman

J. Washington Acad. Sci. 26: 113, 15 Mar 1936b.

= Thomomys bottae desertorum Merriam, 1901. See Hoffmeister (1986:221).

Holotype: 227802. Skin and skull. Adult male. Big Sandy River, 

near Owen (Owens), [Miller and Kellogg (1955:294): about 

4 mi (6 km) southeast of Wickieup], altitude 2,000 ft (610 

m), Mohave County, Arizona. Collected 21 Sep 1917 by 

E. A. Goldman. Original number 23332.

Thomomys bottae divergens Nelson and Goldman

J. Mamm. 15: 122, 15 May 1934c.

Holotype: 250892. Skin and skull. Adult male. Four miles (6 km) 

west of Huachinera, altitude 4,000 ft (1,219 m), Bavispe 

River, northeastern Sonora, Mexico. Collected 8 Nov 1932 

by V. Bailey and F. Winthrop. Original number [FC: VB] 

11098.

Thomomys bottae extenuatus Goldman

Proc. Biol. Soc. Washington 48: 149, 31 Oct 1935b.

= Thomomys bottae mearnsi V. Bailey, 1914. See Hoffmeister 

(1986:233).

Holotype: 17872/24809. Skin and skull. Adult male. Willcox, 

altitude 4,000 ft (1,219 m), Cochise County, Arizona. Collected 27 Nov 1889 by V. Bailey. Original number 712.

Thomomys bottae guadalupensis Goldman

J. Washington Acad. Sci. 26: 117, 15 Mar 1936b.

Holotype: 109225. Skin and skull. Adult male. [FC: Head of] 

McKittrick Canyon, Guadalupe Mountains, altitude 7,800 

ft (2,377 m), [Culberson County], Texas. Collected 22 Aug 

1901 by V. Bailey. Original number 7821.

Thomomys bottae howelli Goldman

J. Washington Acad. Sci. 26: 116, 15 Mar 1936b.

Holotype: 75684. Skin and skull. Adult female. Grand Junction, 

altitude 4,600 ft (1,402 m), Mesa County, Colorado. Collected 7 Nov 1895 by A. H. Howell. Original number 493.

Thomomys bottae hualpaiensis Goldman

J. Washington Acad. Sci. 26: 114, 15 Mar 1936b.

= Thomomys bottae desertorum Merriam, 1901. See Hoffmeister (1986:229).
6 2 • smithsonian contributions to zoology

Holotype: 227796. Skin and skull. Adult male. Hualpai Peak, 

Hualpai Mountains, altitude 7,000 ft (2,134 m), Mohave 

County, Arizona. Collected 6 Oct 1917 by E. A. Goldman. 

Original number 23351.

Thomomys bottae imitabilis Goldman

Proc. Biol. Soc. Washington 52: 30, 11 Mar 1939g.

Holotype: 146839. Skin and skull. Adult male. La Paz, southern 

Lower California (Baja California Sur), Mexico. Collected 

16 Feb 1906 by [E. W.] Nelson and [E. A.] Goldman. Original number 19163.

Thomomys bottae incomptus Goldman

Proc. Biol. Soc. Washington 52: 29, 11 Mar 1939g.

Holotype: 140671. Skin and skull. Adult male. San Jorge, near 

Pacific coast west of Pozo Grande and about 25 mi (40 km) 

southwest of Comondu, altitude 50 ft (15 m), southern 

Lower California (Baja California Sur), Mexico. Collected 

12 Nov 1905 by [E. W.] Nelson and [E. A.] Goldman. Original number 18546.

Thomomys bottae internatus Goldman

J. Washington Acad. Sci. 26: 115, 15 Mar 1936b.

Holotype: 150997. Skin and skull. Adult male. Salida, [FC: 

east Arkansas Valley], altitude 7,000 ft (2,134 m), Chaffee 

County, Colorado. Collected 10 Nov 1907 by M. Cary. 

Original number 1247.

Thomomys bottae levidensis Goldman

Proc. Biol. Soc. Washington 55: 76, 26 Jun 1942.

Holotype: 191962. Skin and skull. Adult male. Manti, [FC: in 

low valley near river], altitude about 5,500 ft (1,676 m), 

Sanpete County, Utah. Collected 6 Dec 1888 by V. Bailey. 

Original number 427. [SL: Merriam collection number 

5137/5832].

Thomomys bottae minimus Durrant

Proc. Biol. Soc. Washington 52: 161, 11 Oct 1939.

Holotype: 263942. Skin and skull. Adult male. [FC: north end] 

Stansbury Island, Great Salt Lake, Tooele County, Utah. Collected 25 Jun 1938 by W. H. Marshall. Original number 141.

Thomomys bottae minor V. Bailey

Proc. Biol. Soc. Washington 27: 116, 10 Jul 1914.

= Thomomys bottae laticeps Baird, 1855. See Patton and Smith 

(1990:115).

Holotype: 146463. [Skin and skull]. Adult male. Fort Bragg, 

Mendocino County, California. Collected 30 Nov 1905 by 

J. H. Gaut. Original number 4187.

Thomomys bottae nicholi Goldman

J. Washington Acad. Sci. 28: 337, 15 Jul 1938c.

= Thomomys bottae planirostris Burt, 1931. See Hoffmeister 

(1986:237).

Holotype: 262864. Skin, skull [and baculum]. Adult male. 

Twenty miles (32 km) south of Wolf Hole [FC: on] road to 

Parashonts, altitude 5,000 ft (1,524 m), Shivwits Plateau, 

Mohave County, Arizona. Collected 6 Aug 1937 by L. C. 

Goldman. Original number 363.

Thomomys bottae operosus Hatfield

Bull. Chicago Acad. Sci. 6: 151, 12 Jan 1942.

= Thomomys bottae fulvus (Woodhouse, 1852). See Hoffmeister 

(1986:231).

Holotype: [287886]. Skin and skull. Adult female. Six miles (10 

km) north of Yarnell, [Miller and Kellogg (1955:294): altitude 4,400 ft (1,341 m)], Peeples Valley, Yavapai County, 

Arizona. Collected 30 May 1937 by R. Komarek. No original number. Chicago Academy of Science number 4061.

Thomomys bottae optabilis Goldman

J. Washington Acad. Sci. 26: 116, 15 Mar 1936b.

Holotype: 149962. Skin and skull. Adult male. Coventry, Naturita Creek Valley, altitude 6,500 ft (1,981 m), Montrose 

County, Colorado. Collected 31 Jul 1907 by M. Cary. Original number 1105.

Thomomys bottae opulentus Goldman

Proc. Biol. Soc. Washington 48: 150, 31 Oct 1935b.

Holotype: 167318. Skin and skull. Adult female. Las Palomas, 

on the Rio Grande, [FC: 4,200 ft (1,280 m)], Sierra County, 

New Mexico. Collected 24 Oct 1909 by E. A. Goldman. 

Original number [FC: Nelson and Goldman] 20675.

Thomomys bottae parvulus Goldman

J. Washington Acad. Sci. 28: 339, 15 Jul 1938c.

= Thomomys bottae catalinae Goldman, 1931. See Hoffmeister 

(1986:222).

Holotype: 262813. Skin and skull. Adult male. [SL: Along highway], [FC: Top of] pass between Santa Catalina and Rincon 

Mountains [FC: in lower edge of oaks], [Miller and Kellogg 

(1955:292): 25 mi (40 km) east-northeast of Tucson], altitude 4,500 ft (1,372 m), Pima County, Arizona. Collected 

5 Jun 1937 by L. C. Goldman. Original number 213.

Thomomys bottae patulus Goldman

J. Washington Acad. Sci. 28: 341, 15 Jul 1938c.

= Thomomys bottae albatus Grinnell, 1912. See Hoffmeister 

(1986:226).

Holotype: 262899. Skin and skull. Adult male. Bottomland along 

Hassayampa River, altitude 2,000 ft (610 m), 2 mi (3.2 km) 

below Wickenburg, Maricopa County, Arizona. Collected 

16 Sep 1937 by L. C. Goldman. Original number 460.

Thomomys bottae pervarius Goldman

Proc. Biol. Soc. Washington 51: 57, 18 Mar 1938a.

Holotype: 18201/25105. Skin and skull. Young adult male. 

Lloyd Ranch, 35 mi (56 km) south of Marfa, altitude 4,200 
number 640 • 6 3

ft (1,280 m), Presidio County, Texas. Collected 20 Jan 1890 

by V. Bailey. Original number 900.

Thomomys bottae pinalensis Goldman

J. Washington Acad. Sci. 28: 342, 15 Jul 1938c.

Holotype: 245709. Skin and skull. Adult female. Oak Flat, 

[Miller and Kellogg (1955:293): altitude 3,900 ft (1,189 

m)], 5 mi (8 km) east of Superior, Pinal Mountains, [Pinal 

County], Arizona. Collected 22 May 1924 by W. P. Taylor. 

Original number 1667.

Thomomys bottae russeolus Nelson and Goldman

Proc. Biol. Soc. Washington 22: 25, 10 Mar 1909.

Holotype: 139920. [Skin and skull]. Young adult male. San 

Angel, [FC: 100 ft (30 m)], 30 mi (48 km) west of [Miller 

and Kellogg (1955:289): 16 mi (26 km) west southwest of] 

San Ignacio, Lower California (Baja California Sur), Mexico. Collected 15 Oct 1905 by E. W. Nelson and E. A. Goldman. Original number 18355.

Thomomys bottae virgineus Goldman

Proc. Biol. Soc. Washington 50: 133, 10 Sep 1937d.

= Thomomys bottae planirostris Burt, 1931. See Hoffmeister 

(1986:237).

Holotype: 262016. Skin and skull. Adult male. Beaverdam 

Creek, [FC: near mouth just above Littlefield], near confluence with Virgin River, at Littlefield, altitude 1,500 ft 

(457 m), [Mohave County], northwestern Arizona. Collected 16 Oct 1936 by L. C. Goldman. Original number 67.

Thomomys bottae winthropi Nelson and Goldman

J. Mamm. 15: 122, 15 May 1934c.

Holotype: 251028. Skin and skull. Adult male. Hermosillo, Sonora, Mexico. Collected 21 Dec 1932 by F. Winthrop and 

B. Bailey. Original number [SL: 32-202]. 27203-X.

Thomomys bridgeri Merriam

Proc. Biol. Soc. Washington 14: 113, 19 Jul 1901d.

= Thomomys talpoides bridgeri Merriam, 1901. See Goldman 

(1939h:234).

Holotype: 18858/25592. Skin, skull [and baculum]. Adult male. 

Fort Bridger, Wyoming. Restricted to Mountainview, Smiths 

Fork, 4 mi (6 km), airline, southeast of Fort Bridger, Uinta 

County, Wyoming by Hooper (1943:503). Collected 27 

May 1890 by V. Bailey. Original number 1207.

Thomomys cabezonae Merriam

Proc. Biol. Soc. Washington 14: 110, 19 Jul 1901d.

= Thomomys bottae nigricans Rhoads, 1895. See Patton and 

Smith (1990:117).

Holotype: 53987. [Skin and skull]. Adult male. Cabezon, [Miller 

and Kellogg (1955:284): 1,791 ft (546 m)], San Gorgonio 

Pass, [Riverside County], California. Collected 3 Jun 1893 

by C. P. Streator. Original number 2906.

Thomomys canus V. Bailey

Proc. Biol. Soc. Washington 23: 79, 4 May 1910.

= Thomomys bottae canus V. Bailey, 1910. See Goldman (1935a: 

155).

Holotype: 78365. [Skin and skull]. Adult male. Deep Hole, at 

north end of Smoke Creek Desert, [Washoe County], Nevada. Collected 14 May 1896 by C. P. Streator. Original 

number 5169.

Thomomys clusius Coues

Proc. Acad. Nat. Sci. Philadelphia 27: 138, 15 Jun 1875b.

Holotype: 3051/[36805]. Skin and Skull. Adult female. Bridger’s 

Pass, Rocky Mountains. Restricted by V. Bailey (1915: 100) 

to Bridger Pass, 18 mi (29 km) southwest of Rawlins, Carbon County, Wyoming. Collected 28, not 21 as in original 

description, Jul 1857 by W. A. Hammond. Original number 

[SL: 88].

Thomomys clusius fuscus Merriam

N. Amer. Fauna 5: 69, 30 Jul 1891b.

= Thomomys talpoides fuscus Merriam, 1891. See Davis (1939: 

253).

Holotype: 24267/31671. Skin and skull. Adult female. Mountains [FC: Canyon] at head of Big Lost River, [Custer 

County], Idaho. Collected 23 Sep 1890 by [SL: V. Bailey 

and] B. H. Dutcher. Original number [FC: VB] 1847.

Thomomys clusius ocius Merriam

Proc. Biol. Soc. Washington 14: 114, 19 Jul 1901d.

= Thomomys talpoides ocius Merriam, 1901. See Durrant 

(1946:17).

Holotype: 18852/25586. Skin and skull. Adult male. Fort 

Bridger, Wyoming. Restricted to Mountainview, Smiths 

Fork, 4 mi (6 km), airline, southeast of Fort Bridger, Uinta 

County, Wyoming by Hooper (1943:503). Collected 24 

May 1890 by V. Bailey. Original number 1194.

Thomomys desertorum Merriam

Proc. Biol. Soc. Washington 14: 114, 19 Jul 1901d.

= Thomomys bottae desertorum Merriam, 1901. See Goldman 

(1935a:156).

Holotype: [186523]. Skin and skull. Adult male. Mud Spring, 

[FC: lowest part of] Detrital Valley, [Mohave County], Arizona. Collected 21 Feb 1889 by V. Bailey. Original number 

598. Merriam collection number 5387/6082.

Thomomys douglasii oregonus Merriam

Proc. Biol. Soc. Washington 14: 115, 19 Jul 1901d.

= Thomomys mazama oregonus Merriam, 1901. See Hall (1981: 

466).

Holotype: 56939. [Skin and skull]. Adult male. [V. Bailey 

(1915:117): Ely’s orchard, Ely, near] Oregon City, Willamette Valley, [Clackamas County], Oregon. Collected 24 

Oct 1893 by C. P. Streator. Original number 3340.
6 4 • smithsonian contributions to zoology

Thomomys douglasii shawi Taylor

Proc. Biol. Soc. Washington 34: 121, 30 Jun 1921.

= Thomomys talpoides shawi Taylor, 1921. See Hall and Dalquest (1939:4).

Holotype: 232807. Skin and skull. Adult male. Owyhigh Lakes, 

altitude 5,100 ft (1,554 m), Mount Rainier, [Pierce County], 

Washington. Collected 9 Aug 1919 by G. G. Cantwell. 

Original number 1464.

Thomomys douglasii tacomensis Taylor

Proc. Biol. Soc. Washington 32: 169, 30 Sep 1919.

= Thomomys mazama tacomensis Taylor, 1919. See Johnson and 

Benson (1960:21).

Holotype: 231096. [Skin and skull]. Young adult female. [FC: 

Meadow Park Golf Links], 6 mi (10 km) south of Tacoma, 

Pierce County, Washington. Collected 24 Dec 1918 by G. G. 

Cantwell. Original number 857.

Thomomys douglasii yelmensis Merriam

Proc. Biol. Soc. Washington 13: 21, 31 Jan 1899a.

= Thomomys mazama yelmensis Merriam, 1899. See Johnson 

and Benson (1960:21).

Holotype: 31916/43777. Skin and skull. Adult male. Tenino, 

Yelm Prairie, [Thurston County], Washington. Collected 24 

Oct 1891 by C. P. Streator. Original number 1385.

Thomomys fossor kaibabensis Goldman

J. Washington Acad. Sci. 28: 333, 15 Jul 1938c.

= Thomomys talpoides kaibabensis Goldman, 1938. See Goldman (1939h:234).

Holotype: 262891. Skin, skull [and baculum]. Adult male. De 

Motte Park, altitude 9,000 ft (2,743 m), Kaibab Plateau, 

Coconino County, Arizona. Collected 10 Sep 1937 by L. C. 

Goldman. Original number 443.

Thomomys fossor levis Goldman

J. Washington Acad. Sci. 28: 336, 15 Jul 1938c.

= Thomomys talpoides levis Goldman, 1938. See Goldman 

(1939h:234).

Holotype: 158979. Skin and skull. Adult female. Seven Mile 

Flat, altitude 10,000 ft (3,048 m), 5 mi (8 km) north of 

Fish Lake, Fish Lake Plateau, Sevier County, Utah. Collected 1 Oct 1908 by W. H. Osgood. Original number 

3616.

Thomomys fossor moorei Goldman

J. Washington Acad. Sci. 28: 335, 15 Jul 1938c.

= Thomomys talpoides moorei Goldman, 1938. See Goldman 

(1939h:234).

Holotype: 248222. Skin and skull. Adult male. One mile 

(1.6 km) south of Fairview, altitude 6,000 ft (1,829 m), Sanpete County, Utah. Collected 19 Feb 1928 by A. W. Moore. 

Original number [SL: 355]. 24799-X.

Thomomys fossor parowanensis Goldman

J. Washington Acad. Sci. 28: 334, 15 Jul 1938c.

= Thomomys talpoides parowanensis Goldman, 1938. See Goldman (1939h:234).

Holotype: 158072. Skin and skull. Adult male. Brian Head, 

altitude 11,000 ft (3,353 m), Parowan Mountains, Iron 

County, Utah. Collected 8 Sep 1908 by W. H. Osgood. 

Original number 3483.

Thomomys fulvus catalinae Goldman

J. Washington Acad. Sci. 21: 419, 19 Oct 1931b.

= Thomomys bottae catalinae Goldman, 1931. See Goldman 

(1935a:157).

Holotype: 244081. [Skin and skull]. Adult male. Summerhaven, 

altitude 7,500 ft (2,286 m), Santa Catalina Mountains, Pima 

County, Arizona. Collected 6 Aug 1923 by E. A. Goldman. 

Original number 23517.

Thomomys fulvus collinus Goldman

J. Washington Acad. Sci. 21: 421, 19 Oct 1931b.

= Thomomys bottae mearnsi V. Bailey, 1914. See Hoffmeister 

(1986:233).

Holotype: 66053. [Skin and skull]. Adult male. Fly Park, altitude 9,000 ft (2,743 m) [FC: 10,000 ft (3,048 m)], Chiricahua Mountains, Cochise County, Arizona. Collected 10 Jun 

1894 by A. K. Fisher. Original number 1527.

Thomomys fulvus emotus Goldman

Proc. Biol. Soc. Washington 46: 76, 27 Apr 1933a.

= Thomomys umbrinus emotus Goldman, 1933. See Nelson and 

Goldman (1934c:116).

Holotype: 157005. Skin and skull. Adult male. Animas Peak, 

altitude 8,000 ft (2,438 m), Animas Mountains, Hidalgo 

County, southwestern New Mexico. Collected 3 Aug 1908 

by E. A. Goldman. Original number [FC: Nelson and Goldman] 19929.

Thomomys fulvus flavidus Goldman

J. Washington Acad. Sci. 21: 417, 19 Oct 1931b.

= Thomomys bottae chrysonotus Grinnell, 1912. See Goldman 

(1935a:156).

Holotype: 181065. [Skin and skull]. Adult male. Parker, altitude 

350 ft (107 m), Yuma County, Arizona. Collected 1 Feb 

1913 by E. A. Goldman. Original number 21810.

Thomomys fulvus grahamensis Goldman

J. Washington Acad. Sci. 21: 420, 19 Oct 1931b.

= Thomomys bottae mearnsi V. Bailey, 1914. See Hoffmeister 

(1986:233).

Holotype: 204313. [Skin and skull]. Adult male. Graham (Pinaleno) Mountains, altitude 9,200 ft (2,804 m), Graham 

County, Arizona. Collected 7 Jun 1914 by E. G. Holt. Original number 269.
number 640 • 6 5

Thomomys fulvus intermedius Mearns

Proc. U.S. Natl. Mus. 19: 719, 30 Jul 1897b.

= Thomomys umbrinus intermedius Mearns, 1897. See Nelson 

and Goldman (1934c:117).

Holotype: 21473/36697. Skin and skull. Adult male. Aspen and 

spruce zone at the summit of the Huachuca Mountains, altitude 9,000 ft (2,743 m), [Cochise County], southern Arizona. 

Hoffmeister (1986:243) states that the locality should just be 

Huachuca Mountains, Arizona. Collected 6 Sep 1893 by F. X. 

Holzner. Original number [FC: Mearns and Holzner] 1013.

Thomomys fulvus modicus Goldman

J. Washington Acad. Sci. 21: 418, 19 Oct 1931b.

= Thomomys bottae modicus Goldman, 1931. See Nelson and 

Goldman (1934c:122).

Holotype: 59203. [Skin and skull]. Adult male. La Osa, near 

Mexican Boundary, southern end of Altar Valley, Pima 

County, Arizona. Collected 14 Dec 1893 by E. A. Mearns 

and F. X. Holzner. Original number 2681.

Thomomys fulvus mutabilis Goldman

Proc. Biol. Soc. Washington 46: 75, 27 Apr 1933a.

= Thomomys bottae fulvus (Woodhouse, 1852). See Hoffmeister 

(1986:231).

Holotype: 214611. Skin and skull. Adult male. Camp Verde, 

[FC: 3 mi (5 km) south at Bell Ranch], altitude 3,200 ft 

(975 m), [Yavapai County], Arizona. Collected 25 Jul 1916 

by W. P. Taylor. Original number 269.

Thomomys fulvus peramplus Goldman

J. Washington Acad. Sci. 21: 423, 19 Oct. 1931b.

= Thomomys bottae peramplus Goldman, 1931. See Goldman 

(1935a:157).

Holotype: 247632. Skin and skull. Adult male. Wheatfield Creek, 

[Miller and Kellogg (1955:297): about 27 mi (43 km) east of 

Chin Lee (Chinle)], west slope of Tunicha Mountains, altitude 

7,000 ft (2,134 m), Apache County, northeastern Arizona. 

Collected 23 Jun 1927 by P. Trapier. Original number 720.

Thomomys fulvus phasma Goldman

Proc. Biol. Soc. Washington 46: 72, 27 Apr 1933a.

= Thomomys bottae pusillus Goldman, 1931. See Hoffmeister 

(1986:238).

Holotype: 203026. Skin and skull. Adult female. Two miles (3.2 

km) south of Tule Tank, Tule Desert, near Mexican Boundary, [Miller and Kellogg (1955:290): altitude 1,200 ft (366 

m)], Yuma County, Arizona. Collected 8 Dec 1913 by E. A. 

Goldman. Original number 22355.

Thomomys fulvus pusillus Goldman

J. Washington Acad. Sci. 21: 422, 19 Oct 1931b.

= Thomomys bottae pusillus Goldman, 1931. See Goldman 

(1935a:157).

Holotype: 209290. [Skin and skull]. Adult female. Coyote Mountains, [Miller and Kellogg (1955:291): Quinlan Mountains, 

40 mi (64 km) west-southwest of Tucson], altitude 3,000 ft 

(914 m), Pima County, Arizona. Collected 4 Sep 1915 by E. 

A. Goldman. Original number 22722.

Thomomys fulvus suboles Goldman

Proc. Biol. Soc. Washington 41: 203, 18 Dec 1928a.

= Thomomys bottae desertorum Merriam, 1901. See Hoffmeister (1986:229).

Holotype: 244163. [Skin and skull]. Adult male. Alluvial bottom at Old Searchlight Ferry, Colorado River, altitude 

1,000 ft (305 m), [Miller and Kellogg (1955:308): about 

40 mi (64 km)] northwest of Kingman [Miller and Kellogg 

(1955:308): or 28 mi (45 km) west of Chloride, Mohave 

County], Arizona. Collected 26 Sep 1923 by L. C. Goldman. Original number 17. [SL: 23428-X].

Thomomys fulvus subsimilis Goldman

Proc. Biol. Soc. Washington 46: 74, 27 Apr 1933a.

= Thomomys bottae subsimilis Goldman, 1933. See Goldman 

(1935a:156).

Holotype: 227803. Skin and skull. Adult female. Harquahala Mountains, altitude 3,000 ft (914 m), [Yuma County], Arizona. Collected 14 Oct 1917 by E. A. Goldman. Original number 23356.

Thomomys fulvus texensis V. Bailey

Proc. Biol. Soc. Washington 15: 119, 2 Jun 1902.

= Thomomys bottae texensis V. Bailey, 1902. See Goldman 

(1935a:157).

Holotype: 22511/[31408], not 2251 as in original description. 

[Skin and skull]. Adult male. Head of Limpia Creek, altitude 5,500 ft (1,676 m), Davis Mountains, [Jeff Davis 

County], Texas. Collected 7 Jan 1890 by V. Bailey. Original 

number 876.

Thomomys fuscus columbianus V. Bailey

Proc. Biol. Soc. Washington 27: 117, 10 Jul 1914.

= Thomomys talpoides columbianus V. Bailey, 1914. See Goldman (1939h:234).

Holotype: 23722/31117. Skin and skull. Adult male. Touchet, 

[FC: about 700 ft (213 m) on edge of plain], Walla Walla 

County, Washington. Collected 10 Sep 1890 by C. P. 

Streator. Original number 234.

Thomomys fuscus fisheri Merriam

Proc. Biol. Soc. Washington 14: 111, 19 Jul 1901d.

= Thomomys talpoides fisheri Merriam, 1901. See Goldman 

(1939h:234).

Holotype: 101238. [Skin and skull]. [Lyon and Osgood 

(1909:64): Young] adult male. Beckwith, Sierra Valley, Plumas County, California. Collected 3 Aug 1900 by W. K. 

Fisher. Original number 1547.
6 6 • smithsonian contributions to zoology

Thomomys fuscus loringi V. Bailey

Proc. Biol. Soc. Washington 27: 118, 10 Jul 1914.

= Thomomys talpoides loringi V. Bailey, 1914. See Goldman 

(1939h:234).

Holotype: 68746. [Skin and skull]. Adult male. South Edmonton, [FC: 2 mi (3.2 km) south of Edmonton], Alberta, 

[Canada]. Collected 23 Sep 1894 by J. A. Loring. Original 

number 2437.

Thomomys fuscus saturatus V. Bailey

Proc. Biol. Soc. Washington 27: 117, 10 Jul 1914.

= Thomomys talpoides saturatus V. Bailey, 1914. See Davis 

(1939:253).

Holotype: 28781/40885. Skin and skull. Adult male. Silver, altitude 4,300 ft (1,311 m) [FC: 3,200 ft (975 m)], near Saltese, 

Coeur d’Alene Mountains, in the western corner of Missoula County, Montana. Collected 20 Jun 1891 by C. P. 

Streator. Original number 979.

Thomomys goldmani Merriam

Proc. Biol. Soc. Washington 14: 108, 19 Jul 1901d.

= Thomomys umbrinus goldmani Merriam, 1901. See Nelson 

and Goldman (1934c:115).

Holotype: 58075. [Skin and skull]. Adult male. Mapimi, [Nelson 

and Goldman (1934c:115): 3,800 ft (1,158 m)], Durango, 

Mexico. Collected 15 Dec 1893 by E. A. Goldman. Original 

number 240.

Thomomys hesperus Merriam

Proc. Biol. Soc. Washington 14: 116, 19 Jul 1901d.

= Thomomys mazama hesperus Merriam, 1901. See Hall (1981: 

466).

Holotype: 69825. [Skin and skull]. Adult female. Tillamook, 

[Tillamook County], Oregon. Collected 9 Nov 1894 by J. E. 

McLellan. Original number 1189.

Thomomys idahoensis Merriam

Proc. Biol. Soc. Washington 14: 114, 19 Jul 1901d.

= Thomomys idahoensis idahoensis Merriam, 1901. See Thaeler 

(1972:420).

Holotype: 23482/30900. Skin and skull. Adult male. Birch 

Creek, [Bailey (1915:108): 10 mi (16 km) south of Nicholia, 

at about 6,400 ft (1,951 m) altitude, Lemhi County], Idaho. 

Collected 8 Aug 1890 by C. P. Streator. Original number 

129.

Thomomys lachuguilla confinalis Goldman

J. Washington Acad. Sci. 26: 119, 15 Mar 1936b.

= Thomomys bottae confinalis Goldman, 1936. See Goldman 

(1938a:55).

Holotype: 117571. Skin and skull. Subadult male. Thirtyfive miles (56 km) east of Rock Springs, altitude 2,450 ft 

(747 m), [Miller and Kellogg (1955:306): north fork of 

Guadalupe River, 15 mi (24 km) west of Japonica, Kerr 

County], Texas. Collected 11 Jul 1902 by V. Bailey. Original number 7910.

Thomomys lachuguilla limitaris Goldman

J. Washington Acad. Sci. 26: 118, 15 Mar 1936b.

= Thomomys bottae limitaris Goldman, 1936. See Goldman 

(1938a:55).

Holotype: 110339. Skin and skull. Adult male. [FC: On mesa] 

4 mi (6.4 km) west of Boquillas, Brewster County, Texas. 

Collected 28 May 1901 by V. Bailey. Original number 7659.

Thomomys laticeps Baird

Proc. Acad. Nat. Sci. Philadelphia 7: 335, reported favorably for 

publication 24 Apr 1855a, receipt acknowledged 15 Jun 

1855 by the American Philosophical Society.

= Thomomys bottae laticeps Baird, 1855. See V. Bailey (1915:46).

Holotype: [513/1648. Skin and skull. Age and sex unknown]. 

Humboldt Bay, [Humboldt County, California]. Collected 

on unknown date by Lieut. [W. P.] Trowbridge. Cataloged 

21 Feb 1855. Type not designated by number in original description. Baird (1857:393) indicates he only had this single 

specimen.

Thomomys latirostris Merriam

Proc. Biol. Soc. Washington 14: 107, 19 Jul 1901d.

= Thomomys bottae aureus J. A. Allen, 1893. See Goldman 

(1935a:156).

Holotype: 18003/24914. Skin and skull. Adult male. [Hall and 

Kelson (1959:426): Tanner Crossing, about 3 mi (4.8 km) 

above Cameron; Hoffmeister (1986:227): says actually 1 mi 

(1.6 km) south, 1 2/3 mi (2.7 km) east of Cameron], [FC: 

4,000 ft (1,219 m)], Little Colorado River, Painted Desert, 

[Coconino County], Arizona. Collected 22 Sep 1899 by V. 

Bailey. Original number 504.

Thomomys leucodon Merriam

Proc. Biol. Soc. Washington 11: 215, 15 Jul 1897c.

= Thomomys bottae leucodon Merriam, 1897. See V. Bailey 

(1915:47).

Holotype: 32034/43848. Skin and skull. Adult male. Grant 

(Grants) Pass, Rogue River Valley, [Josephine County], 

Oregon. Collected 17 Dec 1891 by C. P. Streator. Original 

number 1394.

Thomomys leucodon navus Merriam

Proc. Biol. Soc. Washington 14: 112, 19 Jul 1901d.

= Thomomys bottae navus Merriam, 1901. See V. Bailey 

(1915:49).

Holotype: 57791. [Skin and skull]. Adult male. Red Bluff, [FC: 

Tehama County], California. Collected 26 Dec 1893 by 

C. P. Streator. Original number 3462.

Thomomys limosus Merriam

Proc. Biol. Soc. Washington 14: 116, 19 Jul 1901d.
number 640 • 6 7

= Thomomys talpoides limosus Merriam, 1901. See Goldman 

(1939h:235).

Holotype: 89724. [Skin and skull]. Adult male. White Salmon, 

gorge of the Columbia, [Klickitat County], Washington. Collected 26 Jun 1897 by J. A. Loring. Original number 4382.

Thomomys magdalenae Nelson and Goldman

Proc. Biol. Soc. Washington 22: 24, 10 Mar 1909.

= Thomomys bottae magdalenae Nelson and Goldman, 1909. 

See Goldman (1943:147).

Holotype: 146832. [Skin and skull]. Adult male. Magdalena Island, [FC: 20 ft (6 m)], Lower California (Baja California 

Sur), Mexico. Collected 3 Dec 1905 by E. W. Nelson and 

E. A. Goldman. Original number 18743.

Thomomys mazama Merriam

Proc. Biol. Soc. Washington 11: 214, 15 Jul 1897c.

= Thomomys mazama mazama Merriam, 1897. See Hall (1981: 

466).

Holotype: 80502. [Skin and skull]. Adult male. Head of Anna 

Creek, [V. Bailey (1915: 123): near] Crater Lake, Mount 

Mazama, altitude 6,000 ft (1,829 m), [Klamath County], 

Oregon. Collected 3 Sep 1896 by E. A. Preble. Original 

number 1485.

Thomomys mearnsi V. Bailey

Proc. Biol. Soc. Washington 27: 117, 10 Jul 1914.

= Thomomys bottae mearnsi V. Bailey, 1914. See Anderson 

(1966:195).

Holotype: 157008. [Skin and skull]. Adult male. Gray’s Ranch, 

[SL: altitude 5,000 ft (1,524 m)], Animas Valley [Hidalgo 

County], southwest corner of New Mexico. Collected 

10 Aug 1908 by E. A. Goldman. Original number [FC: Nelson and Goldman] 19939.

Thomomys melanops Merriam

Proc. Biol. Soc. Washington 13: 21, 31 Jan 1899a.

= Thomomys mazama melanops Merriam, 1899. See Johnson 

and Benson (1960:21).

Holotype: 90630. Skin and skull. Adult female. Timberline at head 

of Soleduck (Sole Duc) River, altitude 5,200 ft (1,585 m), 

Olympic Mountains, [Clallam County], Washington. Collected 28 Aug 1897 by V. Bailey. Original number 6219.

Thomomys mewa Merriam

Proc. Biol. Soc. Washington 21: 146, 9 Jun 1908b.

= Thomomys bottae mewa Merriam, 1908. See V. Bailey (1915:50).

Holotype: 133183. [Skin and skull]. Adult male. Raymond, 

Madera County, California. Collected 28 Jun 1904 by N. 

Hollister. Original number 908.

Thomomys monticola pinetorum Merriam

N. Amer. Fauna 16: 97, 28 Oct 1899b.

= Thomomys monticola J. A. Allen, 1893. See Goldman (1943:146).

Holotype: 95152. [Skin and skull]. Adult male. Sisson, Siskiyou 

County, California. Collected 4 Sep 1898 by R. T. Fisher. 

Original number 173.

Thomomys muralis Goldman

J. Washington Acad. Sci. 26: 112, 15 Mar 1936b.

= Thomomys bottae desertorum Merriam, 1901. See Hoffmeister (1986:229).

Holotype: 202580. Skin and skull. Adult male. Lower end of 

Prospect Valley, [Miller and Kellogg (1955:207): about 25 

mi (40 km) west-southwest of Supai], Grand Canyon, altitude 4,500 ft (1,372 m), Hualpai Indian Reservation, [Coconino County], Arizona. Collected 3 Oct 1913 by E. A. 

Goldman. Original number 22264.

Thomomys myops Merriam

Proc. Biol. Soc. Washington 14: 112, 19 Jul 1901d.

= Thomomys talpoides fuscus Merriam, 1891. See Dalquest and 

Scheffer (1944:438).

Holotype: 91066. Skin and skull. Adult female. Conconully, east 

base Cascade Range, [Okanogan County], Washington. 

Collected 11 Sep 1897 by J. A. Loring. Original number 

4650.

Thomomys nasicus Merriam

Proc. Biol. Soc. Washington 11: 216, 15 Jul 1897c.

= Thomomys mazama nasicus Merriam, 1897. See Hall (1981: 

466).

Holotype: 79815. [Skin and skull]. Adult male. Farewell Bend 

(Bend), Des Chutes River, [Deschutes County], Oregon. Collected 4 Aug 1896 by E. A. Preble. Original number 1274.

Thomomys neglectus V. Bailey

Proc. Biol. Soc. Washington 27: 117, 10 Jul 1914.

= Thomomys bottae bottae (Eydoux and Gervais, 1836). See 

Patton and Smith (1990:112).

Holotype: 137869. [Skin and skull]. Adult male. Bear Flat Meadows, altitude 6,400 ft (1,951 m), San Antonio (Baldy) Peak, 

San Gabriel Mountains, [Los Angeles County], California. 

Collected 23 Jul 1905 by J. H. Gaut. Original number 3719.

Thomomys nelsoni Merriam

Proc. Biol. Soc. Washington 14: 109, 19 Jul 1901d.

= Thomomys umbrinus nelsoni Merriam, 1901. See Anderson 

(1972:292).

Holotype: 96451. [Skin and skull]. Adult female. [FC: Near] Parral, [FC: 6,000 ft (1,829 m)], Chihuahua, Mexico. Collected 

18 Sep 1898 by E. W. Nelson and E. A. Goldman. Original 

number 13035.

Thomomys nevadensis Merriam

Proc. Biol. Soc. Washington 11: 213, 15 Jul 1897c.

= Thomomys townsendii nevadensis Merriam, 1897. See V. Bailey (1915:44).
6 8 • smithsonian contributions to zoology

Holotype: 25001/32413. Skin and skull. Young adult male. [FC: 

On flats in valley 5 mi (8 km) south of] Austin, [Miller and 

Kellogg (1955:324): 5 mi (8 km) west, Reese River bottomlands], [Lander County], Nevada. Collected 11 Nov 1890 

by V. Bailey. Original number 2097.

Thomomys nevadensis atrogriseus V. Bailey

Proc. Biol. Soc. Washington 27: 118, 10 Jul 1914.

= Thomomys townsendii townsendii (Bachman, 1839). See 

Miller (1924:236).

Holotype: 181196. [Skin and skull]. Adult male. Nampa, 

[Canyon County], Idaho. Collected 15 Mar 1913 by L. E. 

Wyman. Original number 35. [SL: 10181-X].

Thomomys niger Merriam

Proc. Biol. Soc. Washington 14: 117, 19 Jul 1901d.

= Thomomys mazama niger Merriam, 1901. See Hall (1981: 

466).

Holotype: 69407. [Skin and skull]. Adult male. Seaton, near 

mouth of Umpqua River, [Douglas County], Oregon. Collected 6 Oct 1894 by J. E. McLellan. Original number 1147.

Thomomys operarius Merriam

Proc. Biol. Soc. Washington 11: 215, 15 Jul 1897c.

= Thomomys bottae operarius Merriam, 1897. See Goldman 

(1935a:155).

Holotype: 25054/32466. Skin and skull. Adult male. Keeler, 

[Miller and Rehn (1901:147): east side of] Owens Lake, 

Inyo County, California. Collected 29 Nov 1890 by E. W. 

Nelson. Original number 1.

Thomomys orizabae Merriam

Proc. Biol. Soc. Washington 8: 145, 29 Dec 1893b.

= Thomomys umbrinus umbrinus (Richardson, 1829). See 

Castro-Campilo and Ramirez-Pulido (2000:32).

Holotype: 53616. [Skin and skull]. Adult female. Mount Orizaba, 9,500 ft (2,896 m), Puebla, Mexico. Collected 25 Apr 

1893 by E. W. Nelson. Original number 4744.

Thomomys pectoralis Goldman

J. Washington Acad. Sci. 26: 120, 15 Mar 1936b.

= Thomomys bottae pectoralis Goldman, 1936. See Patton 

(2005b:867).

Holotype: 244372. Skin and skull. Adult male. Vicinity of [FC: 

on ridge near] Carlsbad Cave, Carlsbad Cave National 

Monument, Eddy County, New Mexico. Collected 17 Mar 

1924 by V. Bailey. Original number 10222.

Thomomys perditus Merriam

Proc. Biol. Soc. Washington 14: 108, 19 Jul 1901d.

= Thomomys bottae perditus Merriam, 1901. See Patton 

(2005b:867).

Holotype: 25605/32998. Skin and skull. Adult male. [FC: On 

mountainside], Lampazos [Miller and Kellogg (1955:329): 

Lampazos de Narango], Nuevo Leon, Mexico. Collected 22 

Jan 1891 by C. P. Streator. Original number 512.

Thomomys peregrinus Merriam

Proc. Biol. Soc. Washington 8: 146, 29 Dec 1893b.

= Thomomys umbrinus umbrinus (Richardson, 1829). See 

Castro-Campillo and Ramirez-Pulido (2000:33).

Holotype: 50130. [Skin and skull]. Young adult female. Salazar, 

[FC: 10,300 ft (3,139 m)], Mexico, [Mexico]. Collected 24, 

not 23 as in original description, Oct 1892 by E. W. Nelson. 

Original number 3668.

Thomomys perpallidus absonus Goldman

J. Washington Acad. Sci. 21: 425, 19 Oct 1931b.

= Thomomys bottae planirostris Burt, 1931. See Hoffmeister 

(1986:237).

Holotype: 250016. [Skin and skull]. Adult male. Jacob’s Pools, 

altitude 4,000 ft (1,219 m), Houserock Valley, Coconino 

County, northern Arizona. Collected 7 Jun 1931 by E. A. 

Goldman. Original number 23569.

Thomomys perpallidus dissimilis Goldman

J. Washington Acad. Sci. 21: 425, 19 Oct 1931b.

= Thomomys bottae dissimilis Goldman, 1931. See Goldman 

(1935a:156).

Holotype: 158526. [Skin and skull]. Adult female. East slope of 

Mount Ellen, altitude 8,000 ft (2,438 m), Henry Mountains, 

Garfield County, Utah. Collected 15 Oct 1908 by W. H. Osgood. Original number 3677.

Thomomys perpallidus osgoodi Goldman

J. Washington Acad. Sci. 21: 242, 19 Oct 1931b.

= Thomomys bottae osgoodi Goldman, 1931. See Goldman 

(1935a:156).

Holotype: 158530. [Skin and skull]. Adult male. Hanksville, 

Wayne County, Utah. Collected 20 Oct 1908 by W. H. Osgood. Original number 3701.

Thomomys pryori V. Bailey

Proc. Biol. Soc. Washington 27: 116, 10 Jul 1914.

= Thomomys talpoides pryori V. Bailey, 1914. See V. Bailey 

(1915:104).

Holotype: 66469. [Skin and skull]. Adult male. Head of Sage 

Creek, altitude 6,000 ft (1,829 m), Pryor Mountains, [Carbon County], Montana. Collected 16 Jul 1894 by V. Bailey. 

Original number 4646.

Thomomys pygmaeus Merriam

Proc. Biol. Soc. Washington 14: 115, 19 Jul 1901d.

= Thomomys idahoensis pygmaeus Merriam, 1901. See Thaeler 

(1972:425).

Holotype: 55271. [Skin, skull, and baculum]. Adult male. Montpelier Creek, [Miller (1924:240): about 10 mi (16 km) 

northeast of Montpelier], altitude 6,700 ft (2,042 m), [Bear 
number 640 • 6 9

Lake County], Idaho. Collected 29 Jul 1893 by V. Bailey. 

Original number 4150.

Thomomys quadratus Merriam

Proc. Biol. Soc. Washington 11: 214, 15 Jul 1897c.

= Thomomys talpoides quadratus Merriam, 1897. See Goldman 

(1939h:234).

Holotype: 57134. [Skin and skull]. Adult male. The Dalles, 

[Wasco County], Oregon. Collected 2 Nov 1893 by C. P. 

Streator. Original number 3359.

Thomomys sheldoni V. Bailey

N. Amer. Fauna 39: 93, 15 Nov 1915.

= Thomomys umbrinus madrensis Nelson and Goldman, 1934. 

See Anderson (1972:291).

Holotype: 90819. [Skin and skull]. Adult male. Santa Teresa, 

altitude 6,800 ft (2,073 m), Tepec (Nayarit), Mexico. Collected 10 Aug 1897 by E. W. Nelson and E. A. Goldman. 

Original number 11443.

Thomomys simulus Nelson and Goldman

J. Mamm. 15: 120, 15 May 1934c.

= Thomomys bottae simulus Nelson and Goldman, 1934. See 

Dunnigan (1967:144).

Holotype: 96463. Skin and skull. Adult male. Alamos, altitude 

[FC: about] 1,200 ft (366 m), southern Sonora, Mexico. 

Collected 29 Dec 1898 by E. A. Goldman. Original number 

[FC: Nelson and Goldman] 13339.

Thomomys simulus parviceps Nelson and Goldman

J. Mamm. 15: 121, 15 May 1934c.

= Thomomys atrovarius parviceps Nelson and Goldman, 1934. 

See Hafner et al. (2011:1377).

Holotype: 96743. Skin and skull. Young adult female. Chacala, 

[FC: about] 3,000 ft (914 m), western Durango, Mexico. 

Collected 6 Mar 1899 by E. A. Goldman. Original number 

[FC: Nelson and Goldman] 13485.

Thomomys sinaloae Merriam

Proc. Biol. Soc. Washington 14: 108, 19 Jul 1901d.

= Thomomys bottae sinaloae Merriam, 1901. See Nelson and 

Goldman (1934c:124).

Holotype: 96745. [Skin and skull]. Adult male. Altata, [FC: 

about 175 ft (53 m)], Sinaloa, Mexico. Collected 28 Mar 

1899 by E. A. Goldman. Original number [FC: Nelson and 

Goldman] 13607.

Thomomys sturgisi Goldman

Proc. Biol. Soc. Washington 51: 56, 18 Mar 1938a.

= Thomomys bottae sturgisi Goldman, 1938. See Patton and 

Smith (1990:148).

Holotype: 263376. Skin and skull. Adult female. Carmen Mountains, 

altitude 6,000 ft (1,829 m), Coahuila, Mexico. Collected 21 Sep 

1937 by R. S. Sturgis. Original number 74. [SL: 28064-X].

Thomomys talpoides agrestis Merriam

Proc. Biol. Soc. Washington 21: 144, 9 Jun 1908a.

Holotype: 150725. [Skin and skull]. Adult female. Medano 

Ranch, [SL and FC: 15 mi (24 km) northeast of Mosca], 

[FC: 7,600 ft (2,316 m)], San Luis Valley, [Alamosa County], 

Colorado. Collected 29 Oct 1907 by M. Cary. Original 

number 1205.

Thomomys talpoides andersoni Goldman

J. Mamm. 20: 235, 14 May 1939h.

Holotype: 69001. Skin and skull. Adult male. Medicine Hat, on 

the South Saskatchewan River, Alberta, Canada. Collected 

14 Oct 1894 by J. A. Loring. Original number 2567.

Thomomys talpoides badius Goldman

J. Mamm. 20: 242, 14 May 1939h.

= Thomomys talpoides yakimensis Hall and Dalquest, 1939. See 

Dalquest and Scheffer (1944:440).

Holotype: 91073. Skin and skull. Adult male. Wenatchee, Chelan 

County, Washington. Collected 19 Sep 1897 by J. A. Loring. 

Original number 4695.

Thomomys talpoides bullatus V. Bailey

Proc. Biol. Soc. Washington 27: 115, 10 Jul 1914.

Holotype: 55159, not 55795 as in original description. [Skin and 

skull]. Adult male. Powderville, [Powder River County], 

Montana. Collected 21 Jul 1893 by A. K. Fisher. Original 

number 1402.

Thomomys talpoides caryi V. Bailey

Proc. Biol. Soc. Washington 27: 115, 10 Jul 1914.

Holotype: 168946. [Skin and skull]. Adult female. Head of 

Trapper Creek, [FC: below timberline at] altitude 9,500 ft 

(2,896 m), [FC: west slope] Bighorn Mountains, [Big Horn 

County], Wyoming. Collected 10 Jun 1910 by M. Cary. 

Original number 1993.

Thomomys talpoides couchi Goldman

J. Mamm. 20: 243, 14 May 1939h.

= Thomomys mazama couchi Goldman, 1939. See Johnson and 

Benson (1960:21).

Holotype: 243092. Skin and skull. Adult male. Four miles (6 

km) north of Shelton, Mason County, Washington. Collected 27 Jun 1922 by L. K. Couch. Original number [SL: 

15]. 22896-X.

Thomomys talpoides ericaeus Goldman

J. Mamm. 20: 243, 14 May 1939h.

= Thomomys talpoides devexus Hall and Dalquest, 1939. See 

Dalquest and Scheffer (1944:434).

Holotype: 242445. [Skin and skull]. Adult male. Badger Mountains, altitude 3,000 ft (914 m), 8 mi (13 km) southwest of 

Waterville, Douglas County, Washington. Collected 8 Nov 

1921 by G. G. Cantwell. Original number 2813.
7 0 • smithsonian contributions to zoology

Thomomys talpoides glacialis Dalquest and Scheffer

Proc. Biol. Soc. Washington 55: 97, 13 Aug 1942.

= Thomomys mazama glacialis Dalquest and Scheffer, 1942. See 

Johnson and Benson (1960:21).

Holotype: 272033. Skin and skull. Adult male. Two miles (3.2 

km) south of Washington, Pierce County, Washington. Collected 19 Dec 1941 by W. W. Dalquest. Original number 

2672. [SL: 30556-X].

Thomomys talpoides incensus Goldman

J. Mamm. 20: 240, 14 May 1939h.

Holotype: 67096. Skin and skull. Adult female. Shuswap, Yale 

District, British Columbia, [Canada]. Collected 1 Sep 1894 

by C. P. Streator. Original number 4228.

Thomomys talpoides kelloggi Goldman

J. Mamm. 20: 237, 14 May 1939h.

Holotype: 225907. Skin and skull. Adult male. West Boulder 

Creek, Absaroka Mountain, 18 mi (29 km) southeast of 

Livingston, Park County, Montana. Collected 29 Jun 1917 

by R. Kellogg. Original number 964.

Thomomys talpoides louiei Gardner

J. Mamm. 31: 92, 21 Feb 1950.

= Thomomys mazama louiei Gardner, 1950. See Johnson and 

Benson (1960:21).

Holotype: 276436. Skin and skull. Adult female. CrownZellerbach’s Cathlamet Tree Farm, altitude 2,500 ft 

(762 m), 12 mi (19 km) north northeast of Cathlamet, Wahkiakum County, Washington. Collected 27 May 1949 by W. 

A. Moore. Original number 760. [SL: 33482-X].

Thomomys talpoides medius Goldman

J. Mamm. 20: 241, 14 May 1939h.

Holotype: 66653. Skin and skull. Adult female. Silver King Mine, 

summit of Toad Mountain, 6 mi (9.7 km) south of Nelson, 

Kootenay District, British Columbia, [Canada]. Collected 

23 Jul 1894 by J. A. Loring. Original number 2042.

Thomomys talpoides nebulosus V. Bailey

Proc. Biol. Soc. Washington 27: 116, 10 Jul 1914.

Holotype: 202495. [Skin and skull]. Adult male. Jack Boyden’s 

Ranch, 5 mi (8 km) above mouth of Sand Creek Canyon, altitude 3,750 ft (1,143 m), [Miller (1924:248): 15 mi (24 km) 

northeast of Sundance], Black Hills, [Crook County], Wyoming. Collected 25 Aug 1913 by V. Bailey. Original number 

9508.

Thomomys talpoides perpallidus Merriam

Science 8: 588, 24 Dec 1886d.

= Thomomys bottae perpallidus Merriam, 1886. See Goldman 

(1935a:155).

Lectotype: [191935. Skin and skull. Adult male]. [V. Bailey 

(1915:68): Palm Springs (Agua Caliente), 6 mi (9.7 km) 

south of Palm Springs Station, at sea level on the edge of 

the] Colorado Desert, [Riverside County], California. Collected [SL: 22 Mar] 1886 by F. Stephens. Original number 

[MCC: 235]. [SL: Merriam collection number 2218/2734]. 

No type specimen designated in original description. Lectotype designated by V. Bailey (1915:68).

Thomomys talpoides pugetensis Dalquest and Scheffer

Proc. Biol. Soc. Washington 55: 96, 13 Aug 1942.

= Thomomys mazama pugetensis Dalquest and Scheffer, 1942. 

See Johnson and Benson (1960:21).

Holotype: 272032. Skin, skull [and baculum]. Adult male. Three 

miles (4.8 km), not 4 (6.4 km) as in original description, 

south of Olympia, Thurston County, Washington. Collected 

31 Dec 1940, not 1941 as in original description, by W. W. 

Dalquest. Original number 2022. [SL: 30555-X]. Changes 

from original description correspond to skin label data; also 

see Lidicker (1971:13).

Thomomys talpoides relicinus Goldman

J. Mamm. 20: 239, 14 May 1939h.

Holotype: 243549. Skin and skull. Adult female. Twin Springs, 

20 mi (32 km) north of Minidoka, Snake River Desert, 

[Minidoka County], Idaho. Collected 22 Apr 1923 by L. J. 

Goldman. Original number 2841.

Thomomys talpoides retrorsus Hall

Univ. Kansas Publ., Mus. Nat. Hist. 5: 83, 1 Oct 1951.

Holotype: 69840. Skin and skull. Subadult male. Flagler, Kit 

Carson County, Colorado. Collected 26 Nov 1894 by C. P. 

Streator. Original number 4460.

Thomomys talpoides tenellus Goldman

J. Mamm. 20: 238, 14 May 1939h.

Holotype: 169753. Skin and skull. Adult female. [SL and FC: 

Pahaska Tepee], Whirlwind Peak, Absaroka Range, altitude 

10,500 ft (3,200 m), Park County, Wyoming. Collected 

3 Aug 1910 by M. Cary. Original number 2114.

Thomomys talpoides trivialis Goldman

J. Mamm. 20: 236, 14 May 1939h.

Holotype: 225918. Skin and skull. Adult male. Near head of Big 

Timber Creek, [Miller and Kellogg (1955:312): about 15 mi 

(24 km) northwest Big Timber], altitude 5,200 ft (1,585 m), 

aneroid reading, Crazy Mountains, [Sweet Grass County], 

Montana. Collected 20 Jun 1917 by M. A. Hanna. Original 

number 349.

Thomomys talpoides tumuli Dalquest and Scheffer

Proc. Biol. Soc. Washington 55: 96, 13 Aug 1942.

= Thomomys mazama tumuli Dalquest and Scheffer, 1942. See 

Johnson and Benson (1960:21).

Holotype: 272034. Skin and skull. Adult male. Seven miles 

(11 km) [SL: 5 mi (8 km)] north of Tenino, Thurston 
number 640 • 7 1

County, Washington. Collected 2 Jan 1942 by W. W. Dalquest. Original number 2781. [SL: 30557-X].

Thomomys talpoides whitmani Drake and Booth

Walla Walla Coll. Publ. Dep. Biol. Sci. and Biol. Sta. 1: 52, 25 

Nov 1952.

Holotype: [286835]. Skin and skull, [SL and MC: 14 Jan 1957, 

skin destroyed by moths]. Adult male. Whitman National 

Monument, 6 mi (9.7 km) west of Walla Walla, altitude 750 

ft (229 m), Walla Walla County, Washington. Collected 8 

May 1952 by E. S. Booth. Original number 1694.

Thomomys townsendii lenis Goldman

Proc. Biol. Soc. Washington 55: 75, 25 Jun 1942.

= Thomomys bottae lenis Goldman, 1942. See Durrant 

(1946:53).

Holotype: 264805. Skin and skull. Adult male. Richfield, Sevier 

County, Utah. Collected 11 Mar 1928 by A. W. Moore. 

Original numbers [SL: 203 and 385]. 28835-X.

Thomomys uinta Merriam

Proc. Biol. Soc. Washington 14: 112, 19 Jul 1901d.

= Thomomys talpoides uinta Merriam, 1901. See Goldman 

(1939h:234).

Holotype: 22501/30051. Skin, skull [and baculum]. Adult male. 

[Miller (1924:250): Black’s Fork], [V. Bailey (1915:113): 

north base of Gilbert Peak], altitude 10,000 ft (3,048 m), 

Uinta Mountains, [Summit County], Utah. Collected 6 Jun 

1890 by V. Bailey. Original number 1262.

Thomomys umbrinus albigularis Nelson and Goldman

J. Mamm. 15: 106, 15 May 1934c.

= Thomomys umbrinus umbrinus (Richardson, 1829). See 

Castro-Campillo and Ramirez-Pulido (2000:33).

Holotype: 51888. Skin and skull. Adult female. El Chico, altitude 

9,800 ft (2,987 m), Sierra de Pachuca, Hildalgo, Mexico. 

Collected 27 Mar 1893, not 1896 as in original description, 

by [E. W.] Nelson and [E. A.] Goldman. Original number 

EWN 4505.

Thomomys umbrinus analogus Goldman

Proc. Biol. Soc. Washington 51: 59, 18 Mar 1938a.

= Thomomys bottae analogus Goldman, 1938. See Patton and 

Smith (1990:19).

Holotype: 116994. Skin and skull. Adult male. Sierra Guadalupe, 

[Hall and Kelson (1959:419): about 12 mi (19 km) south of 

General Cepeda], [FC: about 6,500 ft (1,981 m)], southeastern Coahuila, Mexico. Collected 25 Apr 1902 by [E. W.] Nelson and [E. A.] Goldman. Original number 15117.

Thomomys umbrinus caliginosus Nelson and Goldman

J. Mamm. 15: 116, 15 May 1934c.

= Thomomys umbrinus madrensis Nelson and Goldman, 1934. 

See Anderson (1972:291).

Holotype: 250887. [Skin and skull]. Adult male. Eight miles (13 

km) west of Altamirano, altitude 8,000 ft (2,438 m), Sierra 

Madre, northwestern Chihuahua, Mexico, near Sonora 

boundary west of Casas Grandes. Collected 3 Nov 1932 by 

V. Bailey and F. Winthrop. Original number [FC: VB] 11071.

Thomomys umbrinus chihuahuae Nelson and Goldman

J. Mamm. 15: 114, 15 May 1934c.

Holotype: 96452. Skin and skull. Adult male. Sierra Madre, altitude [FC: about] 7,000 ft (2,134 m), about 65 mi (105 

km) east of Batopilas, Chihuahua, Mexico. Collected 1 Oct 

1898 by E. A. Goldman. Original number [FC: Nelson and 

Goldman] 13076.

Thomomys umbrinus chiricahuae Nelson and Goldman

J. Mamm. 15: 117, 15 May 1934c.

= Thomomys bottae mearnsi V. Bailey, 1914. See Hoffmeister 

(1986:233).

Holotype: 247828. Skin and skull. Adult male. Pinery Canyon, 

altitude 7,500 ft (2,286 m), west slope of Chiricahua Mountains, [Cochise County], Arizona. Collected 28 Apr 1927 by 

G. G. Cantwell. Original number 2918.

Thomomys umbrinus crassidens Nelson and Goldman

J. Mamm. 15: 113, 15 May 1934c.

Holotype: 91986. Skin and skull. Adult male. Sierra de Valparaiso, altitude 8,700 ft (2,652 m), western Zacatecas, Mexico. 

Collected 30 Nov 1897 by [E. W.] Nelson and [E. A.] Goldman. Original number 11793.

Thomomys umbrinus durangi Nelson and Goldman

J. Mamm. 15: 114, 15 May 1934c.

Holotype: 94603. Skin and skull. Adult male. Durango, [FC: altitude 6,200 ft (1,890 m)], Durango, Mexico. Collected 28 

[FC: 29] Jun 1898 by [E. W.] Nelson and [E. A.] Goldman. 

Original number 12656.

Thomomys umbrinus enixus Nelson and Goldman

J. Mamm. 15: 112, 15 May 1934c.

Holotype: 90834. Skin and skull. Adult male. Sierra Moroni, 

altitude 8,500 ft (2,591 m), near Plateado, Zacatecas, Mexico. Collected 2 Sep 1897 by [E. W.] Nelson and [E. A.] 

Goldman. Original number 11606.

Thomomys umbrinus eximius Nelson and Goldman

J. Mamm. 15: 118, 15 May 1934c.

Holotype: 96458. Skin and skull. Adult male. Sierra de Choix, 

altitude [FC: about] 5,000 ft (1,524 m), about 20 mi (32 km) 

[FC and SL: 50 mi (80 km)] northeast of Choix, Sinaloa, 

Mexico. Collected 19 Oct 1898 by E. A. Goldman. Original 

number [FC: Nelson and Goldman] 13137.

Thomomys umbrinus extimus Nelson and Goldman

J. Mamm. 15: 119, 15 May 1934c.
7 2 • smithsonian contributions to zoology

= Thomomys atrovarius atrovarius J. A. Allen, 1898. See Hafner 

et al. (2011: 1377).

Holotype: 88131. Skin and skull. Adult male. Colomo, altitude 

600 ft (183 m), southern Nayarit, Mexico. Collected 5 Apr 

1897, not 1898 as in original description, by [E. W.] Nelson 

and [E. A.] Goldman. Original number 10878.

Thomomys umbrinus madrensis Nelson and Goldman

J. Mamm. 15: 115, 15 May 1934c.

Holotype: 98203. Skin and skull. Adult male. Pilares Canyon, 

altitude 6,400 ft (1,951 m), 10 mi (16 km) northeast of Colonia Garcia, and about 25 mi (40 km) southwest of Casas 

Grandes, Chihuahua, Mexico. Collected 23 Jun 1899 by [E. 

W.] Nelson and [E. A.] Goldman. Original number 13810.

Thomomys umbrinus martinensis Nelson and Goldman

J. Mamm. 15: 108, 15 May 1934c.

= Thomomys umbrinus umbrinus (Richardson, 1829). See 

Castro-Campillo and Ramirez-Pulido (2000:33).

Holotype: 55622. Skin and skull. Adult female. San Martin Texmelucan, altitude 7,400 ft (2,256 m), Puebla, Mexico. Collected 13 Aug 1893 by [E. W.] Nelson and [E. A.] Goldman. 

Original number 5305.

Thomomys umbrinus musculus Nelson and Goldman

J. Mamm. 15: 119, 15 May 1934c.

= Thomomys atrovarius atrovarius J. A. Allen, 1898. See Hafner 

et al. (2011:1377).

Holotype: 90818. Skin and skull. Adult male. Pedro Pablo, altitude 3,500 ft (1,067 m), about 22 mi (35 km) east of Acaponeta, Sierra Teponahuaxtla, Nayarit, Mexico. Collected 3 

Aug 1897 by [E. W.] Nelson and [E. A.] Goldman. Original 

number 11440.

Thomomys umbrinus potosinus Nelson and Goldman

J. Mamm. 15: 111, 15 May 1934c.

Holotype: 82057. Skin and skull. Adult male. La Tinaja, altitude 

[FC: about] 6,000 ft (1,829 m), about 20 mi (32 km) northeast of San Luis Potosi, [San Luis Potosi], Mexico. Collected 

22 Jan 1897 by [E. W.] Nelson and [E. A.] Goldman. Original number 10469.

Thomomys umbrinus sonoriensis Nelson and Goldman

J. Mamm. 15: 118, 15 May 1934c.

Holotype: 250893. Skin and skull. Old adult female, not male as 

in original description. Ten miles (16 km) east of Chinapa, 

altitude 3,000 ft (914 m), Sonora River Valley, northern 

Sonora, Mexico. Collected 21 Nov 1932 by V. Bailey and 

F. Winthrop. Original number [FC: VB] 11135.

Thomomys umbrinus supernus Nelson and Goldman

J. Mamm. 15: 110, 15 May 1934c.

Holotype: 81687. Skin and skull. Adult male. Santa Rosa, altitude between 8,500 ft (2,591 m), not 9,500 as in original 

description, and 10,000 ft (3,048 m), about 7 mi (11 km) 

northeast of Guanajuato, Guanajuato, Mexico. Collected 

15 Nov 1896 by [E. W.] Nelson and [E. A.] Goldman. Original number 10353.

Thomomys umbrinus tolucae Nelson and Goldman

J. Mamm. 15: 109, 15 May 1934c.

= Thomomys umbrinus umbrinus (Richardson, 1829). See 

Castro-Campillo and Ramirez-Pulido (2000:33).

Holotype: 55908. Skin and skull. Adult male. North slope Volcano of Toluca, altitude 9,500 ft (2,896 m), [Mexico], Mexico. Collected 5 Sep 1893 by [E. W.] Nelson and [E. A.] 

Goldman. Original number [FC: EWN] 5385.

Thomomys umbrinus vulcanius Nelson and Goldman

J. Mamm. 15: 109, 15 May 1934c.

= Thomomys umbrinus umbrinus (Richardson, 1829). See 

Castro-Campillo and Ramirez-Pulido (2000:33).

Holotype: 51885. Skin and skull. Adult female. Volcano of Popocatepetl, [FC: timberline], altitude 12,900 ft (3,932 m), 

Mexico, Mexico. Collected 27 Feb 1893 by [E. W.] Nelson 

and [E. A.] Goldman. Original number [FC: EWN] 4407.

Thomomys umbrinus zacatecae Nelson and Goldman

J. Mamm. 15: 112, 15 May 1934c.

Holotype: 57972. Skin and skull. Adult male. Berriozabal, altitude 6,600 ft (2,012 m) [FC: 6,800 ft (2,073 m)], Zacatecas, 

Mexico. Collected 29 Dec 1893 by [E. W.] Nelson and [E. 

A.] Goldman. Original number 5607.

Genus Zygogeomys Merriam, 1895

Zygogeomys trichopus Merriam

N. Amer. Fauna 8: 196, 31 Jan 1895.

= Zygogeomys trichopus trichopus Merriam, 1895. See Goldman (1938d:211).

Holotype: 50107. [Skin and skull]. Adult male. Nahuatzin, [FC: 

8,500 ft (2,591 m)], Michoacan, Mexico. Collected 11 Oct 

1892 by E. W. Nelson. Original number 3571.

Zygogeomys trichopus tarascensis Goldman

Proc. Biol. Soc. Washington 51: 211, 23 Dec 1938d.

Valid subspecies according to J. L. Patton (in litt.) 17 Jan 2011.

Holotype: 34922/47187. Skin and skull. Adult female. Mountains 6 mi (10 km) southeast of Patzcuaro, altitude [SL and 

FC: about] 8,000 ft (2,438 m), Michoacan, Mexico. Collected 24 Jul 1892 by [E. W.] Nelson and [E. A.] Goldman. 

Original number [FC: EWN] 2943.
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———. 1890. A Review of Some of the North American Ground Squirrels of the Genus Tamias. Bulletin of the 

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———. 1898. Revision of the Chickarees, or North American Red Squirrels (Subgenus Tamiasciurus). Bulletin of 

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———. 1903. Reports on the Mammals Collected in Northeastern Siberia by the Jesup North Pacific Expedition, 

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———. 1966. Taxonomy of Gophers, Especially Thomomys in Chihuahua, Mexico. Systematic Zoology, 15:189–

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———. 1972. Mammals of Chihuahua, Taxonomy and Distribution. Bulletin of the American Museum of Natural 

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———. 1937a. Two New North American Mammals. Bulletin of the Bailey Museum and Library of Natural 

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———. 1937b. Change of Name of Sciurus carolinensis minutus Bailey. Journal of Mammalogy, 18:516.

Bailey, V. 1902. Seven New Mammals from Western Texas. Proceedings of the Biological Society of Washington, 

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———. 1905. Biological Survey of Texas. North American Fauna, 25:1–222. http://dx.doi.org/10.3996/nafa

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———. 1910. Two New Pocket Gophers of the Genus Thomomys. Proceedings of the Biological Society of Washington, 23:79–80.

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———. 1932e. Two New Rodents from Arizona. Proceedings of the Biological 

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———. 1933a. Five New Rodents from Arizona and New Mexico. Proceedings of 

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———. 1935a. Pocket Gophers of the Thomomys bottae Group in the United 

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———. 1936a. Two New Flying Squirrels from Mexico. Journal of the Washington 

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———. 1937a. A New Kangaroo Rat from Southwestern Arizona. Proceedings of 

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———. 1937b. Four New Mammals from Utah. Proceedings of the Biological Society of Washington, 50:221–226.

———. 1937c. New Rodents from Middle America. Journal of the Washington 

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———. 1938a. Six New Rodents from Coahuila and Texas and Notes on the Status 

of Several Described Forms. Proceedings of the Biological Society of Washington, 51:55–61.

———. 1938b. A New Pocket Mouse from Colorado. Journal of Mammalogy, 

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———. 1938c. New Pocket Gophers of the Genus Thomomys, from Arizona and 

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———. 1939a. A New Pocket Mouse of the Genus Perognathus from Nevada. 

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———. 1939b. A Substitute Name for Perognathus penicillatus serosus. Journal of 

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———. 1890c. Description of a New Prairie Dog from Wyoming. North American 

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Index of Taxa

abbottii

Sciurus, 4

aberti

Sciurus, 20, 22, 23

abieticola

Sciurus, 32

Tamiasciurus, 32

ablusus

Citellus, 34

Urocitellus, 34

absonus

Thomomys, 68

acadicus

Castor, 38

actuosus

Thomomys, 61

adangensis

Callosciurus, 4

Sciurus, 4

adocetus

Citellus, 16

Notocitellus, 16

Aeretes

melanopterus, 3

affinis

Ratufa, 18, 19, 20

agilis

Dipodomys, 45, 47

Perodipus, 46

agrestis

Thomomys, 69

agricolae

Sciurus, 23

albatus

Thomomys, 62

albigularis

Thomomys, 71

albinasus

Pappogeomys, 59

albipes

Sciurus, 21
8 4 • smithsonian contributions to zoology

albiventris

Eutamias, 26

Tamias, 26

albulus

Chaetodipus, 41

Perognathus, 41

alexandrae

Thomomys, 60

alfari

Microsciurus, 15

alfredi

Dipodomys, 44

alienus

Thomomys, 61

alleni

Eutamias, 29

Heteromys, 49

Ictidomys, 13

Orthogeomys, 59

Sciurus, 21

Spermophilus, 13

Tamias, 29

alpinus

Sciuropterus, 12

Tamias, 29

Thomomys, 60

altaicus

Eutamias, 26

altinsularis

Sciurus, 4

ambiguus

Chaetodipus, 39

Dipodomys, 42

Perognathus, 39

amecensis

Pappogeomys, 59

ammophilus

Chaetodipus, 41

Geomys, 57

Perognathus, 41

Ammospermophilus

cinnamomeus, 4

extimus, 4

harrisii, 4

insularis, 4

interpres, 4

leucurus, 4

nelsoni, 4

pennipes, 4

saxicolus, 4

tersus, 4

amoenus

Eutamias, 26

Perognathus, 53

Sciuropterus, 13

Tamias, 26, 27, 29, 31

amotus

Lariscus, 15

amplus

Perognathus, 51

analogus

Thomomys, 71

anambae

Ratufa, 18

anambensis

Sciurus, 5

andersoni

Thomomys, 69

angularis

Thomomys, 60

angusticeps

Cratogeomys, 54

Ratufa, 18

angustirostris

Chaetodipus, 41

Cratogeomys, 56, 57

Perognathus, 41

Platygeomys, 56

animosus

Eutamias, 28

annectens

Heteromys, 47

Spermophilus, 36

Xerospermophilus, 36

annexus

Orthogeomys, 59

annulatus

Notocitellus, 16

Spermophilus, 16

anthonyi

Chaetodipus, 39

Perognathus, 39

Sciurus, 23

aoris

Sciurus, 5

apache

Perognathus, 51

Thomomys, 60

Aplodontia

californica, 3

major, 3

olympica, 3

pacifica, 3

phaea, 3

rainieri, 3

rufa, 3

APLODONTIIDAE, 3

Arctomys

dacota, 14

olympus, 14

arenarius

Chaetodipus, 39, 41, 42

Geomys, 57

Perognathus, 39

arendsis

Sciurus, 5

arenicola

Citellus, 13

Ictidomys, 13

Perognathus, 54

arens

Spermophilus, 37

arizonensis

Callospermophilus, 10

Eutamias, 27

Perognathus, 53

Sciurus, 21

armalis

Sciurus, 5

armatus

Perognathus, 39

Spermophilus, 34

Urocitellus, 34

artemesiae

Citellus, 33

Urocitellus, 33

artus

Chaetodipus, 39

Perognathus, 39

arusinus

Ratufa, 18

asiaticus

Eutamias, 26

Tamias, 29

atratus

Callosciurus, 5

Sciurus, 5

atricapillus

Callosciurus, 5, 6, 8, 9

Sciurus, 5

atristriatus

Eutamias, 26

Sciurus, 5

Tamias, 26

atrodorsalis

Sciurus, 5

atrogriseus

Thomomys, 68

atronasus

Dipodomys, 43

atrovarius

Thomomys, 69, 72

atrox

Sciurus, 5

attwateri

Geomys, 57

aureogaster

Sciurus, 21, 23, 24

aureus

Thomomys, 60, 61, 66

australis

Eutamias, 28

Heteromys, 48

avara

Marmota, 15

avicennia

Sciurus, 24

awahnee

Thomomys, 60

badius

Thomomys, 69

baileyi

Castor, 38

Chaetodipus, 39

Dipodomys, 46

Perognathus, 39

Sciurus, 32

Tamiasciurus, 32

Thomomys, 61

balae

Ratufa, 18

Sciurus, 25

Sundasciurus, 25

baliolus

Sciurus, 25

bancana

Ratufa, 20

bancanus

Nannosciurus, 16

bancarus

Sciurus, 25

Sundasciurus, 25

bangsi

Glaucomys, 11

Perognathus, 54

barberi

Eutamias, 29

Tamias, 29

barrowensis

Spermophilus, 34

batuana

Petaurista, 17

Ratufa, 20

batus

Sciurus, 25

bea

Protoxerus, 18

beardi

Chaetodipus, 38

beecheyi

Citellus, 16

Otospermophilus, 16

Spermophilus, 16

beldingi

Spermophilus, 34

Urocitellus, 34, 36

belti

Sciurus, 21

bentincanus

Sciurus, 5

berdmorei

Lariscus, 15

Menetes, 15

beringensis

Spermophilus, 35

bernardinus

Callospermophilus, 11

Spermophilus, 11

bicolor

Ratufa, 18, 19, 20

bilimitatus

Callosciurus, 5, 7

Sciurus, 5

billitonus

Sciurus, 5

bimaculatus

Callosciurus, 7, 9

Perognathus, 51

birdseyei

Thomomys, 61

bombycinus

Perognathus, 52

bonnevillei

Dipodomys, 44

boonsongi
number 640 • 8 5

Callosciurus, 4

boothiae

Sciurus, 21

borealis

Tamias, 29

borneanus

Nannosciurus, 16

borneensis

Sundasciurus, 25

borneoensis

Sciurus, 5

bottae

Thomomys, 60, 61, 62, 63, 64, 

65, 66, 67, 68, 69, 70, 71

brevicauda

Spermophilus, 10

brevicaudus

Spermophilus, 10

breviceps

Geomys, 57

brevinasus

Perognathus, 54

bridgeri

Thomomys, 63

brunneus

Citellus, 34

Spermophilus, 35

Urocitellus, 34, 35

bryanti

Sciurus, 24

bulana

Ratufa, 18

bullatus

Glaucomys, 11

Thomomys, 69

bulleri

Eutamias, 26

Pappogeomys, 59, 60

bunguranensis

Ratufa, 19

bursarius

Geomys, 57

cabezonae

Perodipus, 46

Thomomys, 63

cacodemus

Eutamias, 28

Tamias, 28

californica

Aplodontia, 3

californicus

Chaetodipus, 39

Dipodomys, 42, 43

Microdipodops, 50

Perognathus, 39

caligata

Marmota, 14, 15

caliginosus

Thomomys, 71

callipeplus

Tamias, 29

callistus

Perognathus, 52

Callosciurus

adangensis, 4

atratus, 5

atricapillus, 5, 6, 8, 9

bilimitatus, 5, 7

bimaculatus, 7, 9

boonsongi, 4

caniceps, 4, 5, 6, 7, 9

casensis, 6

concolor, 7

domelicus, 5, 6

erythraeus, 5, 6, 7, 9

finlaysonii, 4, 7

klossii, 7

melanogaster, 5

melanops, 7, 8

miniatus, 5, 8, 9

nigrovittatus, 5, 7

notatus, 4, 5, 6, 7, 8, 9, 10

piceus, 6

pranis, 7

prevostii, 5, 6, 7, 8, 9

rubeculus, 9

siamensis, 5

styani, 6

trotteri, 7

vittatus, 7, 8, 9, 10

Callospermophilus

arizonensis, 10

bernardinus, 11

caryi, 10

castanurus, 10, 11

certus, 10

chrysodeirus, 10, 11

cinerascens, 11

connectens, 10

lateralis, 10, 11

madrensis, 10

mitratus, 10

tescorum, 10

trinitatis, 10

canadensis

Castor, 38

Marmota, 15

canescens

Chaetodipus, 41

Perognathus, 41

Spermophilus, 36

Xerospermophilus, 36, 37

canicaudus

Eutamias, 26

Tamias, 26

caniceps

Callosciurus, 4, 5, 6, 7, 9

Eutamias, 27

Sciurus, 6

Tamias, 27

canigenus

Sciurus, 6

canipes

Eutamias, 27

Tamias, 27

canus

Citellus, 33

Liomys, 49

Spermophilus, 35

Thomomys, 63

Urocitellus, 33, 35

carimatae

Sciurus, 6

carimonensis

Ratufa, 18

Sciurus, 6

carminis

Eutamias, 27

Tamias, 27

carolinensis

Sciurus, 21, 22

caryi

Callospermophilus, 10

Eutamias, 28

Perognathus, 51

Tamias, 28

Thomomys, 69

cascadensis

Marmota, 14

Sciurus, 31

casensis

Callosciurus, 6

Sciurus, 6

castaneus

Funambulus, 14

castanonotus

Sciurus, 22

castanops

Cratogeomys, 54, 55, 56, 57

Pseudostoma, 57

castanurus

Callospermophilus, 10, 11

Tamias, 11

Castor

acadicus, 38

baileyi, 38

canadensis, 38

frontador, 38

mexicanus, 38

michiganensis, 38

missouriensis, 38

repentinus, 38

shastensis, 38

subauratus, 38

texensis, 38

CASTORIDAE, 38

CASTORIMORPHA, 38

catalinae

Thomomys, 62, 64

catemana

Ratufa, 18

catopterius

Heteromys, 48

caurinus

Eutamias, 27

Tamias, 27

celaenopepla

Ratufa, 18

celsus

Dipodomys, 44

certus

Callospermophilus, 10

Chaetodipus

albulus, 41

ambiguus, 39

ammophilus, 41

angustirostris, 41

anthonyi, 39

arenarius, 39, 41, 42

artus, 39

baileyi, 39

beardi, 38

californicus, 39

canescens, 41

dalquesti, 40

dispar, 39

eremicus, 40

extimus, 39

fallax, 39, 40

formosus, 40

goldmani, 40

hispidus, 40, 41

inopinus, 40

insularis, 39

intermedius, 38, 41

magdalenae, 42

nelsoni, 41

occultus, 42

ochrus, 39

pallidus, 40

paradoxus, 41

penicillatus, 41, 42

pernix, 42

phasma, 41

rostratus, 42

rudinoris, 39

seri, 42

siccus, 42

sobrinus, 42

spilotus, 41

spinatus, 42

stephensi, 42

sublucidus, 39

zacatecae, 40

cherriei

Orthogeomys, 59

chiapensis

Heterogeomys, 58

Orthogeomys, 58

Sciurus, 23

chihuahuae

Thomomys, 71

chiliensis

Sciurus, 24

chiricahuae

Sciurus, 22

Thomomys, 71

choco

Sciurus, 24

chrysodeirus

Callospermophilus, 10, 11

Spermophilus, 10

Tamias, 11

chrysonotus

Thomomys, 64
8 6 • smithsonian contributions to zoology

chuscensis

Sciurus, 20

cincticauda

Eliomys, 37

cineraceus

Dipodomys, 45

cinerascens

Callospermophilus, 11

Tamias, 11

cinereicollis

Eutamias, 27

Tamias, 27, 30

cinereus

Eutamias, 27

Sciurus, 24

Tamias, 27

cinnamomeus

Ammospermophilus, 4

Tamias, 4

Citellus

ablusus, 34

adocetus, 16

arenicola, 13

artemesiae, 33

beecheyi, 16

brunneus, 34

canus, 33

columbianus, 33

elegans, 33

grammurus, 16

hollisteri, 13

idahoensis, 33

juglans, 16

leurodon, 33

loringi, 34

mollis, 33

monticola, 13

nebulicola, 33

nevadensis, 33

pallescens, 36

parvulus, 16

pessimus, 33

plesius, 34

ruficaudus, 33

sierra, 16

spilosoma, 36

stejnegeri, 34

townsendii, 34

tridecemlineatus, 13

utah, 16

variegatus, 16

vigilis, 33

washingtoni, 34

washoensis, 33

clarencei

Dipodomys, 46

clarkii

Geomys, 56

clarus

Eutamias, 27

Perognathus, 54

cleomophila

Dipodomys, 45

Perognathus, 51

clusius

Thomomys, 63

cocos

Sciurus, 24

colimensis

Sciurus, 21

colliaei

Sciurus, 22, 24

collinus

Thomomys, 64

coloratus

Glaucomys, 11

columbianus

Citellus, 33

Dipodomys, 47

Perodipus, 47

Perognathus, 52

Thomomys, 65

Urocitellus, 33

columbiensis

Glaucomys, 12

compactus

Dipodomys, 43

concavus

Heterogeomys, 58

Orthogeomys, 58

concinnus

Exilisciurus, 11

concolor

Callosciurus, 7

Sciurus, 20

condurensis

Ratufa, 18

Sciurus, 6

confinalis

Thomomys, 66

confinis

Eutamias, 28

Ratufa, 19

Tamias, 28

conipus

Sciurus, 8

connectens

Callospermophilus, 10

conscius

Heteromys, 48

consitus

Cratogeomys, 54, 55

consobrinus

Eutamias, 27

Tamias, 27, 30

conspicua

Ratufa, 19

convergens

Thomomys, 61

convexus

Cratogeomys, 54

cooperi

Tamias, 30

costaricensis

Macrogeomys, 58

Orthogeomys, 59

cothurnata

Ratufa, 19

couchi

Thomomys, 69

couchii

Otospermophilus, 17

Spermophilus, 17

crassidens

Thomomys, 71

crassirostris

Heteromys, 48

cratodon

Dipodomys, 46

Cratogeomys

angusticeps, 54

angustirostris, 56, 57

castanops, 54, 55, 56, 57

consitus, 54, 55

convexus, 54

estor, 55

excelsus, 54

fulvescens, 55

fumosus, 56, 57

goldmani, 54, 55

hirtus, 54

imparilis, 56

irolonis, 55

lacrimalis, 54

merriami, 55, 56

oreocetes, 55

peregrinus, 55

peridoneus, 55

perotensis, 55, 56

perplanus, 55

planiceps, 56

planifrons, 55

saccharalis, 55

subluteus, 55

subnubilus, 55

subsimus, 55

tamaulipensis, 55

tylorhinus, 56

Cricetodipus

parvus, 50

crinitus

Petinomys, 17

Sciuropterus, 17

crispus

Heteromys, 49

Liomys, 49

cryptospilotus

Spermophilus, 36

Xerospermophilus, 36

cultellus

Thomomys, 61

cupidineus

Dipodomys, 45

Cynomys

gunnisoni, 11

leucurus, 11

mexicanus, 11

zuniensis, 11

dacota

Arctomys, 14

Marmota, 14

dalquesti

Chaetodipus, 40

Perognathus, 40

dariensis

Macrogeomys, 59

Orthogeomys, 59

datus

Sciurus, 6

davisi

Eutamias, 28

Tamias, 28

denticulatus

Eliomys, 37

deppei

Sciurus, 22, 23

deserti

Dipodomys, 43

desertorum

Thomomys, 61, 63, 65, 67

desitus

Thomomys, 61

desmarestianus

Heteromys, 48

devexus

Thomomys, 69

dilutus

Sciurus, 6

Dipodomys

agilis, 45, 47

alfredi, 44

ambiguus, 42

atronasus, 43

baileyi, 46

bonnevillei, 44

californicus, 42, 43

celsus, 44

cineraceus, 45

clarencei, 46

cleomophila, 45

columbianus, 47

compactus, 43

cratodon, 46

cupidineus, 45

deserti, 43

durranti, 45

elator, 43

evexus, 45

exilis, 43

fuscus, 45

gabrielsoni, 43

goldmani, 46

heermanni, 43, 46, 47

ingens, 46

insularis, 43

kernensis, 43

leucotis, 44

levipes, 46

longipes, 45, 46

luteolus, 47

margaritae, 43

marshalli, 45

mayensis, 43

medius, 45

merriami, 42, 43, 44, 45
number 640

• 8 7

microps, 44, 46

mitchelli, 44

montanus, 44

morroensis, 47

nelsoni, 44

nevadensis, 43

nexilis, 45

nitratoides, 43, 44

nitratus, 44

oklahomae, 44

ordii, 44, 45, 46, 47

ornatus, 45

panamintinus, 47

peninsularis, 47

perblandus, 46

perotensis, 45

perplexus, 47

phillipsii, 45

platycephalus, 45

preblei, 46

regillus, 44

russeolus, 44

simulans, 46, 47

sonoriensis, 43

spectabilis, 45, 46

stephensi, 47

subtenuis, 44

tularensis, 46

utahensis, 47

venustus, 47

zygomaticus, 46

Dipodops

longipes, 46

director

Sciurus, 6

dispar

Chaetodipus, 39

Perognathus, 39

dissimilis

Thomomys, 68

divergens

Thomomys, 61

dolichocephalus

Macrogeomys, 59

Orthogeomys, 59

domelicus

Callosciurus, 5, 6

Sciurus, 6

domensis

Perognathus, 39

domisaxensis

Perognathus, 40

doorsiensis

Tamias, 31

dorsalis

Eutamias, 27

Sciurus, 22

Tamias, 27, 30

douglasii

Sciurus, 31

Tamiasciurus, 31, 32

Thomomys, 63, 64

Dremomys

lentus, 11

pernyi, 11

rufigenis, 11

dulitensis

Sciurus, 6

durangae

Tamias, 26

durangi

Thomomys, 71

durranti

Dipodomys, 45

dutcheri

Geomys, 57

effugius

Sciurus, 21

elator

Dipodomys, 43

elbeli

Tamiops, 33

elegans

Citellus, 33

Spermophilus, 35

Urocitellus, 33, 35

Eliomys

cincticauda, 37

denticulatus, 37

munbyanus, 37

parvus, 37

quercinus, 37

emotus

Thomomys, 64

empetra

Spermophilus, 35

endemicus

Spermophilus, 35

Urocitellus, 35

enixus

Thomomys, 71

ephippium

Ratufa, 20

epomophorus

Sciurus, 6

erebus

Sciurus, 6

eremicus

Chaetodipus, 40

Perognathus, 40

ericaeus

Thomomys, 69

erythraeus

Callosciurus, 5, 6, 7, 9

Sciurus, 7

estor

Cratogeomys, 55

Eutamias

albiventris, 26

alleni, 29

altaicus, 26

amoenus, 26

animosus, 28

arizonensis, 27

asiaticus, 26

atristriatus, 26

australis, 28

barberi, 29

bulleri, 26

cacodemus, 28

canicaudus, 26

caniceps, 27

canipes, 27

carminis, 27

caryi, 28

caurinus, 27

cinereicollis, 27

cinereus, 27

clarus, 27

confinis, 28

consobrinus, 27

davisi, 28

dorsalis, 27

grisescens, 28

hopiensis, 27

inyoensis, 29

jacksoni, 28

ludibundus, 27

meridionalis, 27

merriami, 27

minimus, 27, 28

obscurus, 28

ochraceus, 26

ochrogenys, 29

operarius, 26

oreocetes, 28

pallidus, 28

palmeri, 28

quadrivittatus, 28

ruficaudus, 28

sedulus, 29

senescens, 28

sequoiensis, 29

sibiricus, 29

simulans, 28

siskiyou, 29

solivagus, 26

sonoriensis, 27

speciosus, 29

townsendii, 29

umbrinus, 29

umbrosus, 26

utahensis, 27

vallicola, 26

evexus

Dipodomys, 45

evidens

Sciurus, 17

excelsus

Cratogeomys, 54

exilis

Dipodomys, 43

Exilisciurus

concinnus, 11

eximius

Thomomys, 71

extenuatus

Thomomys, 61

extimus

Ammospermophilus, 4

Chaetodipus, 39

Perognathus, 39

Sciurus, 22

Thomomys, 71

fallax

Chaetodipus, 39, 40

Geomys, 58

Perognathus, 40

fasciatus

Perognathus, 52

felipensis

Orthogeomys, 59

femoralis

Ratufa, 19

ferreus

Sciurus, 20

finlaysoni

Sciurus, 7

finlaysonii

Callosciurus, 4, 7

fisheri

Otospermophilus, 16

Spermophilus, 16

Tamias, 31

Thomomys, 65

flammeus

Pappogeomys, 59

flavescens

Perognathus, 51, 52

flavidus

Thomomys, 64

flaviventer

Marmota, 14, 15

flaviventris

Marmota, 14, 15

flavus

Perognathus, 51, 52, 53

formosus

Chaetodipus, 40

Perognathus, 40

fossor

Sciurus, 22, 23

Thomomys, 64

fraterculus

Sundasciurus, 26

fremonti

Sciurus, 31

Tamiasciurus, 32

frontador

Castor, 38

frumentor

Sciurus, 21

fuliginosus

Perognathus, 53

fulvescens

Cratogeomys, 55

fulvus

Geomys, 60

Thomomys, 60, 62, 64, 65

fumosus

Cratogeomys, 56, 57

Geomys, 56

Funambulus

castaneus, 14
8 8 • smithsonian contributions to zoology

Funambulus (continued)

obscurus, 14

peninsulae, 14

rostratus, 14

fuscus

Dipodomys, 45

Glaucomys, 12

Thomomys, 63, 65, 66, 67

gabrielsoni

Dipodomys, 43

gambianus

Heliosciurus, 13

GEOMYIDAE, 54

Geomys

ammophilus, 57

arenarius, 57

attwateri, 57

breviceps, 57

bursarius, 57

clarkii, 56

dutcheri, 57

fallax, 58

fulvus, 60

fumosus, 56

gymnurus, 56

llanensis, 57

lutescens, 57

mobilensis, 58

nelsoni, 59

personatus, 57, 58

pinetis, 58

sagittalis, 57

texensis, 57, 58

tropicalis, 58

tuza, 58

wisconsinensis, 57

gilvus

Perognathus, 53

glacialis

Thomomys, 70

Glaucomys

bangsi, 11

bullatus, 11

coloratus, 11

columbiensis, 12

fuscus, 12

goldmani, 12

griseifrons, 12

guerreroensis, 12

herreranus, 12

klamathensis, 12

latipes, 12

macrotis, 12

madrensis, 12

sabrinus, 11, 12

saturatus, 12

stephensi, 12

texensis, 12

volans, 12

yukonensis, 12

zaphaeus, 12

GLIRIDAE, 37

goldmani

Chaetodipus, 40

Cratogeomys, 54, 55

Dipodomys, 46

Glaucomys, 12

Heteromys, 48

Notocitellus, 16

Perodipus, 46

Perognathus, 40, 42

Sciuropterus, 12

Sciurus, 23

Spermophilus, 16

Thomomys, 66

gracilis

Tamias, 31

grahamensis

Thomomys, 64

grammurus

Citellus, 16

Otospermophilus, 16

granatensis

Sciurus, 23, 24

grandis

Orthogeomys, 59

Graphiurus

isolatus, 37

johnstoni, 38

kelleni, 37, 38

murinus, 37, 38

personatus, 38

griseicollis

Ratufa, 19

griseifrons

Glaucomys, 12

griseoflavus

Sciurus, 23

grisescens

Eutamias, 28

Tamias, 28

griseus

Heteromys, 48

Sciurus, 22, 23

guadalupensis

Thomomys, 61

guerrerensis

Heteromys, 49

Liomys, 49

Orthogeomys, 59

guerreroensis

Glaucomys, 12

gunnisoni

Cynomys, 11

Spermophilus, 11

gymnurus

Geomys, 56

Platygeomys, 56

harrisii

Ammospermophilus, 4

Spermophilus, 4

heermanni

Dipodomys, 43, 46, 47

Heliosciurus

gambianus, 13

maculatus, 13

madogae, 13

multicolor, 13

rufobrachiatus, 13

rufobrachium, 13

shindi, 13

undulatus, 13

helveolus

Sciurus, 24

helvus

Sciurus, 6

hernandezi

Sciurus, 21

herreranus

Glaucomys, 12

hesperus

Thomomys, 66

heterodus

Orthogeomys, 59

Heterogeomys

chiapensis, 58

concavus, 58

hispidus, 58

isthmicus, 58

teapensis, 58

tehuantepecus, 58

torridus, 58

yucatanensis, 58

HETEROMYIDAE, 38

Heteromys

alleni, 49

annectens, 47

australis, 48

catopterius, 48

conscius, 48

crassirostris, 48

crispus, 49

desmarestianus, 48

goldmani, 48

griseus, 48

guerrerensis, 49

hispidus, 50

irroratus, 49, 50

lepturus, 48

nelsoni, 48

oasicus, 48

panamensis, 48

pictus, 47, 49, 50

planifrons, 48

plantinarensis, 49, 50

salvini, 49

subaffinis, 48

temporalis, 48

texensis, 49, 50

torridus, 50

zonalis, 48

heterothrix

Liomys, 49

hippurellus

Sciurus, 25

hippurosus

Sciurus, 25

Sundasciurus, 25

hippurus

Sundasciurus, 25

hirtus

Cratogeomys, 54

Sciurus, 24

hispidus

Chaetodipus, 40, 41

Heterogeomys, 58

Heteromys, 50

Orthogeomys, 58

Perognathus, 40

hollisteri

Citellus, 13

Ictidomys, 13

hopiensis

Eutamias, 27

Perognathus, 52

horsfieldii

Iomys, 14

howelli

Thomomys, 61

hualpaiensis

Thomomys, 61

hudsonicus

Sciurus, 32

Tamiasciurus, 32

humilis

Sciurus, 25

Sundasciurus, 25

Hylopetes

spadiceus, 13

hypoleucos

Ratufa, 18, 19, 20

hypophaeus

Sciurus, 21

ictericus

Sciurus, 7

Ictidomys

alleni, 13

arenicola, 13

hollisteri, 13

monticola, 13

pallidus, 13

parvidens, 13

texensis, 13

tridecemlineatus, 13

idahoensis

Citellus, 33

Perognathus, 54

Thomomys, 66, 68

Urocitellus, 33

imitabilis

Thomomys, 62

imparilis

Cratogeomys, 56

Platygeomys, 56

incanus

Pteromys, 18

incensus

Thomomys, 70

inclarus

Platygeomys, 56

incomptus

Thomomys, 62

inexpectatus
number 640

• 8 9

Sciurus, 6

infuscatus

Sciurus, 23

ingens

Dipodomys, 46

Perodipus, 46

inopinus

Chaetodipus, 40

Perognathus, 40

inornatus

Perognathus, 53

insignis

Lariscus, 14

Ratufa, 18, 19

insularis

Ammospermophilus, 4

Chaetodipus, 39

Dipodomys, 43

Perognathus, 39

intermedius

Chaetodipus, 38, 41

Perognathus, 41

Thomomys, 65

internatus

Thomomys, 62

interpres

Ammospermophilus, 4

Tamias, 4

inyoensis

Eutamias, 29

Tamias, 29

Iomys

horsfieldii, 14

lepidus, 14

thomsoni, 14

irolonis

Cratogeomys, 55

irroratus

Heteromys, 49, 50

Liomys, 50

isolatus

Graphiurus, 37

isthmicus

Heterogeomys, 58

Orthogeomys, 58

isthmius

Liomys, 49

Microsciurus, 15

Sciurus, 15

jacksoni

Eutamias, 28

Perognathus, 51

johnstoni

Graphiurus, 38

juglans

Citellus, 16

kahari

Paraxerus, 17

kaibabensis

Sciurus, 23

Thomomys, 64

kelleni

Graphiurus, 37, 38

kelloggi

Thomomys, 70

kenaiensis

Tamiasciurus, 32

kennicottii

Urocitellus, 34, 35

kernensis

Dipodomys, 43

klamathensis

Glaucomys, 12

Sciuropterus, 12

klossii

Callosciurus, 7

Sciurus, 7

kodiacensis

Spermophilus, 36

Urocitellus, 36

lachuguilla

Thomomys, 60, 66

lacrimalis

Cratogeomys, 54

laenata

Ratufa, 19

lagunensis

Pappogeomys, 60

lamucotanus

Sciurus, 7

lancavensis

Sciurus, 7

Lariscus

amotus, 15

berdmorei, 15

insignis, 14

obscurus, 14

peninsulae, 14

rostratus, 14

lateralis

Callospermophilus, 10, 11

laticeps

Thomomys, 62, 66

latifrons

Orthogeomys, 59

latipes

Glaucomys, 12

latirostris

Thomomys, 66

lautensis

Sciurus, 7

lenis

Thomomys, 71

lentus

Dremomys, 11

lepidus

Iomys, 14

lepturus

Heteromys, 48

leucodon

Thomomys, 66

leucotis

Dipodomys, 44

Tamiops, 33

leucurus

Ammospermophilus, 4

Cynomys, 11

Tamias, 4

leurodon

Citellus, 33

levidensis

Thomomys, 62

levipes

Dipodomys, 46

Perodipus, 46

levis

Thomomys, 64

liantis

Tamiops, 33

libericus

Sciurus, 13

limitaris

Thomomys, 66

limitis

Sciurus, 23

limosus

Thomomys, 66, 67

lingungensis

Sciurus, 25

Liomys

canus, 49

crispus, 49

guerrerensis, 49

heterothrix, 49

irroratus, 49

isthmius, 49

minor, 50

obscurus, 49

orbitalis, 49

parviceps, 49

phaeura, 49

pictus, 49

plantinarensis, 50

pretiosus, 49

rostratus, 49

setosus, 49

sonorana, 50

texensis, 50

torridus, 50

veraecrucis, 50

littoralis

Sciurus, 24

litus

Perognathus, 52

llanensis

Geomys, 57

longimembris

Otognosis, 51

Perognathus, 51, 52, 53, 54

longipes

Dipodomys, 45, 46

Dipodops, 46

loringi

Citellus, 34

Thomomys, 66

louiei

Thomomys, 70

lowii

Sundasciurus, 25

lucas

Sciurus, 7

lucidus

Microdipodops, 50

ludibundus

Eutamias, 27

Tamias, 27

lugens

Petinomys, 17

luteiventris

Tamias, 31

luteola

Marmota, 14, 15

luteolus

Dipodomys, 47

Perodipus, 47

lutescens

Geomys, 57

Sciurus, 7

Macrogeomys

costaricensis, 58

dariensis, 59

dolichocephalus, 59

macrorhabdotes

Tamias, 30

macrospilotus

Spermophilus, 37

macrotis

Glaucomys, 12

Sciuropterus, 12

maculatus

Heliosciurus, 13

madogae

Heliosciurus, 13

madrensis

Callospermophilus, 10

Glaucomys, 12

Thomomys, 69, 71, 72

maerens

Sciuropterus, 17

magdalenae

Chaetodipus, 42

Perognathus, 42

Thomomys, 67

magruderensis

Perognathus, 54

major

Aplodontia, 3

Ratufa, 18

Spermophilus, 37

Xerospermophilus, 37

managuensis

Sciurus, 21

mansalaris

Sciurus, 25

marchio

Petaurista, 17

margaritae

Dipodomys, 43

marginatus

Spermophilus, 37

marinsularis

Sciurus, 7
9 0 • smithsonian contributions to zoology

Marmota

avara, 15

caligata, 14, 15

canadensis, 15

cascadensis, 14

dacota, 14

flaviventer, 14, 15

flaviventris, 14, 15

luteola, 14, 15

monax, 15

nivaria, 14

nosophora, 14

obscura, 15

okanagana, 14, 15

olympus, 14

parvula, 15

preblorum, 15

pretensis, 15

rufescens, 15

sheldoni, 14

sibila, 15

sierra, 15

warreni, 15

marshalli

Dipodomys, 45

martinensis

Thomomys, 72

masae

Ratufa, 19

matecumbei

Sciurus, 22

matthaeus

Sciurus, 7

mayensis

Dipodomys, 43

mazama

Thomomys, 63, 64, 66, 67, 

68, 69, 70

mcclellandi

Tamiops, 33

mearnsi

Sciurus, 32

Tamiasciurus, 32

Thomomys, 61, 64, 67, 71

medius

Dipodomys, 45

Thomomys, 70

megacephalus

Microdipodops, 50

melanocaudus

Perognathus, 40

melanogaster

Callosciurus, 5

melanopepla

Ratufa, 18, 19, 20

melanops

Callosciurus, 7, 8

Sciurus, 7, 8

Thomomys, 67

melanopterus

Aeretes, 3

melanotis

Nannosciurus, 16

Perognathus, 51

melanurus

Tamias, 30

mendanauus

Sciurus, 8

Menetes

berdmorei, 15

meridionalis

Eutamias, 27

Tamias, 27

merriami

Cratogeomys, 55, 56

Dipodomys, 42, 43, 44, 45

Eutamias, 27

Perognathus, 53

mewa

Thomomys, 67

mexicanus

Castor, 38

Cynomys, 11

Perognathus, 52

Spermophilus, 13

michiganensis

Castor, 38

Microdipodops

californicus, 50

lucidus, 50

megacephalus, 50

oregonus, 50

pallidus, 50

microps

Dipodomys, 44, 46

Perodipus, 46

Microsciurus

alfari, 15

isthmius, 15

mimulus, 15

venustulus, 15

vivatus, 15

mimellus

Sciurus, 8

mimiculus

Sciurus, 8

mimicus

Petaurista, 17

mimulus

Microsciurus, 15

mimus

Sciurus, 20

miniatus

Callosciurus, 5, 8, 9

Sciurus, 8

minimus

Eutamias, 27, 28

Tamias, 26, 27, 28, 29, 30, 

31

Thomomys, 62

minnesota

Tamiasciurus, 32

minor

Liomys, 50

Thomomys, 62

minutus

Sciurus, 22

missouriensis

Castor, 38

mitchelli

Dipodomys, 44

mitratus

Callospermophilus, 10

mobilensis

Geomys, 58

modicus

Thomomys, 65

mogollonensis

Tamiasciurus, 32

mohavensis

Spermophilus, 36

Xerospermophilus, 36

mollipilosus

Perognathus, 53

Tamiasciurus, 31

mollis

Citellus, 33

Spermophilus, 35

Urocitellus, 33, 35

monax

Marmota, 15

montanus

Dipodomys, 44

Perodipus, 47

monticola

Citellus, 13

Ictidomys, 13

Perognathus, 53

Thomomys, 67

moorei

Thomomys, 64

morroensis

Dipodomys, 47

Perodipus, 47

morulus

Sciurus, 24

multicolor

Heliosciurus, 13

munbyanus

Eliomys, 37

muralis

Thomomys, 67

murii

Tamiasciurus, 32

murinus

Graphiurus, 37, 38

Prosciurillus, 17, 18

Sciurus, 17

musculus

Thomomys, 72

mutabilis

Thomomys, 65

Myomimus

setzeri, 38

myops

Thomomys, 67

Nannosciurus

bancanus, 16

borneanus, 16

melanotis, 16

pulcher, 16

sumatranus, 16

surrutilus, 11

nasicus

Thomomys, 67

natuensis

Sundasciurus, 25

navus

Thomomys, 66

nayaritensis

Pappogeomys, 60

Sciurus, 22

nebulicola

Citellus, 33

Urocitellus, 33

nebulosus

Thomomys, 70

necopinus

Prosciurillus, 18

Sciurus, 17

neglectus

Platygeomys, 56

Spermophilus, 36

Tamias, 28

Thomomys, 67

Xerospermophilus, 36

negligens

Sciurus, 23

nelsoni

Ammospermophilus, 4

Chaetodipus, 41

Dipodomys, 44

Geomys, 59

Heteromys, 48

Orthogeomys, 59

Perognathus, 41, 42

Sciurus, 23, 24

Spermophilus, 4

Thomomys, 67

nemoralis

Sciurus, 21

neomexicanus

Sciurus, 31

nevadensis

Citellus, 33

Dipodomys, 43

Perognathus, 53

Thomomys, 67, 68

Urocitellus, 33

nexilis

Dipodomys, 45

nicholi

Thomomys, 62

niger

Sciurus, 23, 24

Thomomys, 68

nigrescens

Petaurista, 17

Ratufa, 19

Sciurus, 21, 23, 24

nigricans

Thomomys, 63

nigrovittatus

Callosciurus, 5, 7

nitratoides

Dipodomys, 43, 44

nitratus

Dipodomys, 44

nivaria
number 640 • 9 1

Marmota, 14

norosiensis

Sciurus, 23

nosophora

Marmota, 14

notabilis

Ratufa, 19

notatus

Callosciurus, 4, 5, 6, 7, 8, 9, 

10

Sciurus, 8

Notocitellus

adocetus, 16

annulatus, 16

goldmani, 16

novemlineatus

Sciurus, 32

nuchalis

Sciurus, 22

nyx

Sciurus, 8

oasicus

Heteromys, 48

obscura

Marmota, 15

obscurus

Eutamias, 28

Funambulus, 14

Lariscus, 14

Liomys, 49

Perognathus, 41

Tamias, 27, 28, 30

obsidianus

Spermophilus, 37

obsoletus

Spermophilus, 36

Xerospermophilus, 36

occultus

Chaetodipus, 42

Perognathus, 42

ochraceus

Eutamias, 26

Paraxerus, 17

Tamias, 26

ochrogenys

Eutamias, 29

Tamias, 29

ochrus

Chaetodipus, 39

Perognathus, 39

ocius

Thomomys, 63

oculatus

Sciurus, 24

okanagana

Marmota, 14, 15

oklahomae

Dipodomys, 44

olivaceus

Perognathus, 53, 54

olympica

Aplodontia, 3

olympus

Arctomys, 14

Marmota, 14

operarius

Eutamias, 26

Tamias, 26, 27

Thomomys, 68

operosus

Thomomys, 62

optabilis

Thomomys, 62

opulentus

Thomomys, 62

orbitalis

Liomys, 49

ordii

Dipodomys, 44, 45, 46, 47

Perodipus, 47

oregonensis

Sciuropterus, 12

oregonus

Microdipodops, 50

Spermophilus, 36

Thomomys, 63

Urocitellus, 36

oreocetes

Cratogeomys, 55

Eutamias, 28

Tamias, 28

orizabae

Thomomys, 68

ornatus

Dipodomys, 45

Orthogeomys

alleni, 59

annexus, 59

cherriei, 59

chiapensis, 58

concavus, 58

costaricensis, 59

dariensis, 59

dolichocephalus, 59

felipensis, 59

grandis, 59

guerrerensis, 59

heterodus, 59

hispidus, 58

isthmicus, 58

latifrons, 59

nelsoni, 59

teapensis, 58

tehuantepecus, 58

torridus, 58

vulcani, 59

yucatanensis, 58

osgoodi

Spermophilus, 36

Thomomys, 68

Urocitellus, 36

Otognosis

longimembris, 51

Otospermophilus

beecheyi, 16

couchii, 17

fisheri, 16

grammurus, 16

parvulus, 16

sierra, 16

utah, 16

variegatus, 16, 17

pacifica

Aplodontia, 3

pacificus

Perognathus, 53

pallescens

Citellus, 36

Tamiasciurus, 32

Xerospermophilus, 36

palliata

Ratufa, 19, 20

pallidus

Chaetodipus, 40

Eutamias, 28

Ictidomys, 13

Microdipodops, 50

Perognathus, 40

Spermophilus, 13

Tamias, 31

palmeri

Eutamias, 28

Tamias, 28

palustris

Sciurus, 5

panamensis

Heteromys, 48

panamintinus

Dipodomys, 47

Perodipus, 47

Perognathus, 53, 54

Tamias, 31

pannovianus

Sciurus, 8

Pappogeomys

albinasus, 59

amecensis, 59

bulleri, 59, 60

flammeus, 59

lagunensis, 60

nayaritensis, 60

paradoxus

Chaetodipus, 41

Perognathus, 41

Paraxerus

kahari, 17

ochraceus, 17

parowanensis

Thomomys, 64

parryii

Spermophilus, 36

Urocitellus, 33, 34, 35, 36

parviceps

Liomys, 49

Thomomys, 69

parvidens

Ictidomys, 13

Spermophilus, 13

parvula

Marmota, 15

parvulus

Citellus, 16

Otospermophilus, 16

Thomomys, 62

parvus

Cricetodipus, 50

Eliomys, 37

Perognathus, 50, 52, 53, 54

Sciurus, 25

Sundasciurus, 25

pascalis

Thomomys, 60

patulus

Thomomys, 62

pectoralis

Thomomys, 68

pelapius

Sciurus, 8

pemangilensis

Sciurus, 8

penialius

Sciurus, 8

penicillatus

Chaetodipus, 41, 42

Perognathus, 41, 42

peninsulae

Funambulus, 14

Lariscus, 14

Ratufa, 19

peninsularis

Dipodomys, 47

Perodipus, 47

Sciurus, 8

pennipes

Ammospermophilus, 4

peramplus

Thomomys, 65

perblandus

Dipodomys, 46

perditus

Thomomys, 68

peregrinator

Sciurus, 24

peregrinus

Cratogeomys, 55

Thomomys, 68

pergracilis

Perognathus, 51

peridoneus

Cratogeomys, 55

perijae

Sciurus, 23

perniger

Perognathus, 52

pernix

Chaetodipus, 42

Perognathus, 42

pernyi

Dremomys, 11

Perodipus

agilis, 46

cabezonae, 46

columbianus, 47

goldmani, 46

ingens, 46

levipes, 46

luteolus, 47

microps, 46
9 2 • smithsonian contributions to zoology

Perodipus (continued)

montanus, 47

morroensis, 47

ordii, 47

panamintinus, 47

peninsularis, 47

perplexus, 47

preblei, 46

simulans, 47

stephensi, 47

streatori, 47

tularensis, 46

utahensis, 47

venustus, 47

Perognathus

albulus, 41

ambiguus, 39

ammophilus, 41

amoenus, 53

amplus, 51

angustirostris, 41

anthonyi, 39

apache, 51

arenarius, 39

arenicola, 54

arizonensis, 53

armatus, 39

artus, 39

baileyi, 39

bangsi, 54

bimaculatus, 51

bombycinus, 52

brevinasus, 54

californicus, 39

callistus, 52

canescens, 41

caryi, 51

clarus, 54

cleomophila, 51

columbianus, 52

dalquesti, 40

dispar, 39

domensis, 39

domisaxensis, 40

eremicus, 40

extimus, 39

fallax, 40

fasciatus, 52

flavescens, 51, 52

flavus, 51, 52, 53

formosus, 40

fuliginosus, 53

gilvus, 53

goldmani, 40, 42

hispidus, 40

hopiensis, 52

idahoensis, 54

inopinus, 40

inornatus, 53

insularis, 39

intermedius, 41

jacksoni, 51

litus, 52

longimembris, 51, 52, 53, 54

magdalenae, 42

magruderensis, 54

melanocaudus, 40

melanotis, 51

merriami, 53

mexicanus, 52

mollipilosus, 53

monticola, 53

nelsoni, 41, 42

nevadensis, 53

obscurus, 41

occultus, 42

ochrus, 39

olivaceus, 53, 54

pacificus, 53

pallidus, 40

panamintinus, 53, 54

paradoxus, 41

parvus, 50, 52, 53, 54

penicillatus, 41, 42

pergracilis, 51

perniger, 52

pernix, 42

phasma, 41

piperi, 52

plerus, 54

relictus, 51

rostratus, 42

rotundus, 51

seri, 42

serosus, 42

siccus, 42

sobrinus, 42

sonoriensis, 52

spilotus, 41

spinatus, 42

stephensi, 42

sublucidus, 39

taylori, 51

zacatecae, 40

perotensis

Cratogeomys, 55, 56

Dipodomys, 45

Spermophilus, 36

Xerospermophilus, 36

perpallidus

Thomomys, 68, 70

perpes

Thomomys, 60

perplanus

Cratogeomys, 55

perplexus

Dipodomys, 47

Perodipus, 47

personatus

Geomys, 57, 58

Graphiurus, 38

pervagus

Thomomys, 61

pervarius

Thomomys, 62

pessimus

Citellus, 33

petaurista

Petaurista, 17

Petaurista

batuana, 17

marchio, 17

mimicus, 17

nigrescens, 17

petaurista, 17

philippensis, 17

rubicundus, 17

sulcatus, 3

terutaus, 17

yunanensis, 17

Petinomys

crinitus, 17

lugens, 17

petulans

Sciurus, 32

Tamiasciurus, 32

phaea

Aplodontia, 3

phaeopepla

Ratufa, 18, 20

phaeura

Liomys, 49

phasma

Chaetodipus, 41

Perognathus, 41

Thomomys, 65

philippensis

Petaurista, 17

phillipsii

Dipodomys, 45

piceus

Callosciurus, 6

pictus

Heteromys, 47, 49, 50

Liomys, 49

Tamias, 30

pinalensis

Thomomys, 63

pinetis

Geomys, 58

pinetorum

Thomomys, 67

piniensis

Ratufa, 20

Sciurus, 25

piperi

Perognathus, 52

planiceps

Cratogeomys, 56

Platygeomys, 56

planifrons

Cratogeomys, 55

Heteromys, 48

planirostris

Thomomys, 62, 63, 68

plantinarensis

Heteromys, 49, 50

Liomys, 50

platycephalus

Dipodomys, 45

Platygeomys

angustirostris, 56

gymnurus, 56

imparilis, 56

inclarus, 56

neglectus, 56

planiceps, 56

tylorhinus, 56

varius, 56

zinseri, 57

plerus

Perognathus, 54

plesius

Citellus, 34

Spermophilus, 35

Urocitellus, 35

polia

Ratufa, 20

poliopus

Sciurus, 8, 24

potosinus

Thomomys, 72

pranis

Callosciurus, 7

Sciurus, 7

pratensis

Spermophilus, 37

Xerospermophilus, 37

preblei

Dipodomys, 46

Perodipus, 46

Tamiasciurus, 32

preblorum

Marmota, 15

pretensis

Marmota, 15

pretiosus

Liomys, 49

Sciurus, 8

prevostii

Callosciurus, 5, 6, 7, 8, 9

procerus

Sciurus, 25

Sundasciurus, 25

Prosciurillus

murinus, 17, 18

necopinus, 18

proserpinae

Sciurus, 9

Protoxerus

bea, 18

stangeri, 18

pryori

Thomomys, 68

Pseudostoma

castanops, 57

Pteromys

incanus, 18

volans, 18

pugetensis

Thomomys, 70

pulcher

Nannosciurus, 16

pumilus

Sciurus, 26

pusillus
number 640

• 9 3

Thomomys, 65

pygmaeus

Thomomys, 68

pyrsonota

Ratufa, 20

quadratus

Thomomys, 69

quadrimaculatus

Tamias, 30

quadrivittatus

Eutamias, 28

Tamias, 28, 31

quercinus

Eliomys, 37

Sciurus, 21

rabori

Sundasciurus, 26

rainieri

Aplodontia, 3

Ratufa

affinis, 18, 19, 20

anambae, 18

angusticeps, 18

arusinus, 18

balae, 18

bancana, 20

batuana, 20

bicolor, 18, 19, 20

bulana, 18

bunguranensis, 19

carimonensis, 18

catemana, 18

celaenopepla, 18

condurensis, 18

confinis, 19

conspicua, 19

cothurnata, 19

ephippium, 20

femoralis, 19

griseicollis, 19

hypoleucos, 18, 19, 20

insignis, 18, 19

laenata, 19

major, 18

masae, 19

melanopepla, 18, 19, 20

nigrescens, 19

notabilis, 19

palliata, 19, 20

peninsulae, 19

phaeopepla, 18, 20

piniensis, 20

polia, 20

pyrsonota, 20

tiomanensis, 20

vittata, 20

vittatula, 20

regillus

Dipodomys, 44

relicinus

Thomomys, 70

relictus

Perognathus, 51

repentinus

Castor, 38

retrorsus

Thomomys, 70

richmondi

Sciurus, 24

rodolphii

Tamiops, 33

rostratus

Chaetodipus, 42

Funambulus, 14

Lariscus, 14

Liomys, 49

Perognathus, 42

rotundus

Perognathus, 51

rubeculus

Callosciurus, 9

Sciurus, 9

rubicundus

Petaurista, 17

rubidiventris

Sciurus, 9

rudinoris

Chaetodipus, 39

rufa

Aplodontia, 3

rufescens

Marmota, 15

ruficaudus

Citellus, 33

Eutamias, 28

Tamias, 28, 29

Urocitellus, 33

rufigenis

Dremomys, 11

rufobrachiatus

Heliosciurus, 13

Sciurus, 13

rufobrachium

Heliosciurus, 13

rupatius

Sciurus, 10

russeolus

Dipodomys, 44

Thomomys, 63

rutiliventris

Sciurus, 9

sabrinus

Glaucomys, 11, 12

Sciuropterus, 12

saccharalis

Cratogeomys, 55

sagittalis

Geomys, 57

salvini

Heteromys, 49

sanggaus

Sciurus, 9

saturatus

Glaucomys, 12

Sciurus, 9

Thomomys, 66

saxicolus

Ammospermophilus, 4

Spermophilus, 4

SCIURIDAE, 3

SCIUROMORPHA, 3

Sciuropterus

alpinus, 12

amoenus, 13

crinitus, 17

goldmani, 12

klamathensis, 12

macrotis, 12

maerens, 17

oregonensis, 12

sabrinus, 12

stephensi, 12

volans, 12

yukonensis, 12

zaphaeus, 12

Sciurus

abbottii, 4

aberti, 20, 22, 23

abieticola, 32

adangensis, 4

agricolae, 23

albipes, 21

alleni, 21

altinsularis, 4

anambensis, 5

anthonyi, 23

aoris, 5

arendsis, 5

arizonensis, 21

armalis, 5

atratus, 5

atricapillus, 5

atristriatus, 5

atrodorsalis, 5

atrox, 5

aureogaster, 21, 23, 24

avicennia, 24

baileyi, 32

balae, 25

baliolus, 25

bancarus, 25

batus, 25

belti, 21

bentincanus, 5

bilimitatus, 5

billitonus, 5

boothiae, 21

borneoensis, 5

bryanti, 24

caniceps, 6

canigenus, 6

carimatae, 6

carimonensis, 6

carolinensis, 21, 22

cascadensis, 31

casensis, 6

castanonotus, 22

chiapensis, 23

chiliensis, 24

chiricahuae, 22

choco, 24

chuscensis, 20

cinereus, 24

cocos, 24

colimensis, 21

colliaei, 22, 24

concolor, 20

condurensis, 6

conipus, 8

datus, 6

deppei, 22, 23

dilutus, 6

director, 6

domelicus, 6

dorsalis, 22

douglasii, 31

dulitensis, 6

effugius, 21

epomophorus, 6

erebus, 6

erythraeus, 7

evidens, 17

extimus, 22

ferreus, 20

finlaysoni, 7

fossor, 22, 23

fremonti, 31

frumentor, 21

goldmani, 23

granatensis, 23, 24

griseoflavus, 23

griseus, 22, 23

helveolus, 24

helvus, 6

hernandezi, 21

hippurellus, 25

hippurosus, 25

hirtus, 24

hudsonicus, 32

humilis, 25

hypophaeus, 21

ictericus, 7

inexpectatus, 6

infuscatus, 23

isthmius, 15

kaibabensis, 23

klossii, 7

lamucotanus, 7

lancavensis, 7

lautensis, 7

libericus, 13

limitis, 23

lingungensis, 25

littoralis, 24

lucas, 7

lutescens, 7

managuensis, 21

mansalaris, 25

marinsularis, 7

matecumbei, 22

matthaeus, 7

mearnsi, 32

melanops, 7, 8
9 4 • smithsonian contributions to zoology

Sciurus (continued)

mendanauus, 8

mimellus, 8

mimiculus, 8

mimus, 20

miniatus, 8

minutus, 22

morulus, 24

murinus, 17

nayaritensis, 22

necopinus, 17

negligens, 23

nelsoni, 23, 24

nemoralis, 21

neomexicanus, 31

niger, 23, 24

nigrescens, 21, 23, 24

norosiensis, 23

notatus, 8

novemlineatus, 32

nuchalis, 22

nyx, 8

oculatus, 24

palustris, 5

pannovianus, 8

parvus, 25

pelapius, 8

pemangilensis, 8

penialius, 8

peninsularis, 8

peregrinator, 24

perijae, 23

petulans, 32

piniensis, 25

poliopus, 8, 24

pranis, 7

pretiosus, 8

procerus, 25

proserpinae, 9

pumilus, 26

quercinus, 21

richmondi, 24

rubeculus, 9

rubidiventris, 9

rufobrachiatus, 13

rupatius, 10

rutiliventris, 9

sanggaus, 9

saturatus, 9

senex, 24

seraiae, 9

serutus, 9

siriensis, 9

socialis, 24

suckleyi, 32

sullivanus, 9

surdus, 26

tachin, 5

tapanulius, 10

tarrae, 23

tarussanus, 10

tedongus, 9

tenuirostris, 9

tenuis, 26

thomasi, 24

tolucae, 24

trotteri, 7

truei, 24

ubericolor, 10

underwoodi, 21

undulatus, 13

variabilis, 24

variegatoides, 21, 23, 24

variegatus, 8

ventorum, 32

vittatus, 10

vivax, 22

vulgaris, 23, 24

yucatanensis, 22, 25

sedulus

Eutamias, 29

Tamias, 29

senescens

Eutamias, 28

Tamias, 29

senex

Sciurus, 24

Tamias, 31

sequoiensis

Eutamias, 29

Tamias, 29

seraiae

Sciurus, 9

seri

Chaetodipus, 42

Perognathus, 42

serosus

Perognathus, 42

serutus

Sciurus, 9

setosus

Liomys, 49

setzeri

Myomimus, 38

shastensis

Castor, 38

shawi

Thomomys, 64

sheldoni

Marmota, 14

Thomomys, 69

shindi

Heliosciurus, 13

siamensis

Callosciurus, 5

sibila

Marmota, 15

sibiricus

Eutamias, 29

Tamias, 26, 29

siccus

Chaetodipus, 42

Perognathus, 42

sierra

Citellus, 16

Marmota, 15

Otospermophilus, 16

simulans

Dipodomys, 46, 47

Eutamias, 28

Perodipus, 47

Tamias, 29

simulus

Thomomys, 69

sinaloae

Thomomys, 69

siriensis

Sciurus, 9

siskiyou

Eutamias, 29

Tamias, 29

sobrinus

Chaetodipus, 42

Perognathus, 42

socialis

Sciurus, 24

solivagus

Eutamias, 26

sonomae

Tamias, 29

sonorana

Liomys, 50

sonoriensis

Dipodomys, 43

Eutamias, 27

Perognathus, 52

Tamias, 27

Thomomys, 72

sp.

Tamias, 27

spadiceus

Hylopetes, 13

spatiosus

Thomomys, 61

speciosus

Eutamias, 29

Tamias, 29, 31

spectabilis

Dipodomys, 45, 46

Spermophilus

alleni, 13

annectens, 36

annulatus, 16

arens, 37

armatus, 34

barrowensis, 34

beecheyi, 16

beldingi, 34

beringensis, 35

bernardinus, 11

brevicauda, 10

brevicaudus, 10

brunneus, 35

canescens, 36

canus, 35

chrysodeirus, 10

couchii, 17

cryptospilotus, 36

elegans, 35

empetra, 35

endemicus, 35

fisheri, 16

goldmani, 16

gunnisoni, 11

harrisii, 4

kodiacensis, 36

macrospilotus, 37

major, 37

marginatus, 37

mexicanus, 13

mohavensis, 36

mollis, 35

neglectus, 36

nelsoni, 4

obsidianus, 37

obsoletus, 36

oregonus, 36

osgoodi, 36

pallidus, 13

parryii, 36

parvidens, 13

perotensis, 36

plesius, 35

pratensis, 37

saxicolus, 4

spilosoma, 36, 37

stephensi, 35

tereticaudus, 37

texensis, 13

tridecemlineatus, 13

yakimensis, 35

spilosoma

Citellus, 36

Spermophilus, 36, 37

Xerospermophilus, 36, 37

spilotus

Chaetodipus, 41

Perognathus, 41

spinatus

Chaetodipus, 42

Perognathus, 42

stangeri

Protoxerus, 18

stejnegeri

Citellus, 34

Urocitellus, 34

stephensi

Chaetodipus, 42

Dipodomys, 47

Glaucomys, 12

Perodipus, 47

Perognathus, 42

Sciuropterus, 12

Spermophilus, 35

streatori

Perodipus, 47

striatus

Tamias, 31

sturgisi

Thomomys, 69

styani

Callosciurus, 6

subaffinis

Heteromys, 48

subauratus

Castor, 38
number 640 • 9 5

sublucidus

Chaetodipus, 39

Perognathus, 39

subluteus

Cratogeomys, 55

subnubilus

Cratogeomys, 55

suboles

Thomomys, 65

subsimilis

Thomomys, 65

subsimus

Cratogeomys, 55

subtenuis

Dipodomys, 44

suckleyi

Sciurus, 32

sulcatus

Petaurista, 3

sullivanus

Sciurus, 9

sumatranus

Nannosciurus, 16

Sundasciurus

balae, 25

bancarus, 25

borneensis, 25

fraterculus, 26

hippurosus, 25

hippurus, 25

humilis, 25

lowii, 25

natuensis, 25

parvus, 25

procerus, 25

rabori, 26

tenuis, 25, 26

supernus

Thomomys, 72

surdus

Sciurus, 26

surrutilus

Nannosciurus, 11

swinhoei

Tamiops, 33

tachin

Sciurus, 5

tacomensis

Thomomys, 64

talpoides

Thomomys, 63, 64, 65, 66, 

67, 68, 69, 70, 71

tamaulipensis

Cratogeomys, 55

Tamias

albiventris, 26

alleni, 29

alpinus, 29

amoenus, 26, 27, 29, 31

asiaticus, 29

atristriatus, 26

barberi, 29

borealis, 29

cacodemus, 28

callipeplus, 29

canicaudus, 26

caniceps, 27

canipes, 27

carminis, 27

caryi, 28

castanurus, 11

caurinus, 27

chrysodeirus, 11

cinerascens, 11

cinereicollis, 27, 30

cinereus, 27

cinnamomeus, 4

confinis, 28

consobrinus, 27, 30

cooperi, 30

davisi, 28

doorsiensis, 31

dorsalis, 27, 30

durangae, 26

fisheri, 31

gracilis, 31

grisescens, 28

interpres, 4

inyoensis, 29

leucurus, 4

ludibundus, 27

luteiventris, 31

macrorhabdotes, 30

melanurus, 30

meridionalis, 27

minimus, 26, 27, 28, 29, 30, 

31

neglectus, 28

obscurus, 27, 28, 30

ochraceus, 26

ochrogenys, 29

operarius, 26, 27

oreocetes, 28

pallidus, 31

palmeri, 28

panamintinus, 31

pictus, 30

quadrimaculatus, 30

quadrivittatus, 28, 31

ruficaudus, 28, 29

sedulus, 29

senescens, 29

senex, 31

sequoiensis, 29

sibiricus, 26, 29

simulans, 29

siskiyou, 29

sonomae, 29

sonoriensis, 27

sp., 27

speciosus, 29, 31

striatus, 31

townsendii, 30

umbrinus, 29, 31

umbrosus, 26

utahensis, 27

vallicola, 26

Tamiasciurus

abieticola, 32

baileyi, 32

douglasii, 31, 32

fremonti, 32

hudsonicus, 32

kenaiensis, 32

mearnsi, 32

minnesota, 32

mogollonensis, 32

mollipilosus, 31

murii, 32

pallescens, 32

petulans, 32

preblei, 32

ventorum, 32

wasatchensis, 32

Tamiops

elbeli, 33

leucotis, 33

liantis, 33

mcclellandi, 33

rodolphii, 33

swinhoei, 33

vestitus, 33

tapanulius

Sciurus, 10

tarascensis

Zygogeomys, 72

tarrae

Sciurus, 23

tarussanus

Sciurus, 10

taylori

Perognathus, 51

teapensis

Heterogeomys, 58

Orthogeomys, 58

tedongus

Sciurus, 9

tehuantepecus

Heterogeomys, 58

Orthogeomys, 58

temporalis

Heteromys, 48

tenellus

Thomomys, 70

tenuirostris

Sciurus, 9

tenuis

Sciurus, 26

Sundasciurus, 25, 26

tereticaudus

Spermophilus, 37

Xerospermophilus, 36, 37

tersus

Ammospermophilus, 4

terutaus

Petaurista, 17

tescorum

Callospermophilus, 10

texensis

Castor, 38

Geomys, 57, 58

Glaucomys, 12

Heteromys, 49, 50

Ictidomys, 13

Liomys, 50

Spermophilus, 13

Thomomys, 65

thomasi

Sciurus, 24

Thomomys

absonus, 68

actuosus, 61

agrestis, 69

albatus, 62

albigularis, 71

alexandrae, 60

alienus, 61

alpinus, 60

analogus, 71

andersoni, 69

angularis, 60

apache, 60

atrogriseus, 68

atrovarius, 69, 72

aureus, 60, 61, 66

awahnee, 60

badius, 69

baileyi, 61

birdseyei, 61

bottae, 60, 61, 62, 63, 64, 65, 

66, 67, 68, 69, 70, 71

bridgeri, 63

bullatus, 69

cabezonae, 63

caliginosus, 71

canus, 63

caryi, 69

catalinae, 62, 64

chihuahuae, 71

chiricahuae, 71

chrysonotus, 64

clusius, 63

collinus, 64

columbianus, 65

confinalis, 66

convergens, 61

couchi, 69

crassidens, 71

cultellus, 61

desertorum, 61, 63, 65, 67

desitus, 61

devexus, 69

dissimilis, 68

divergens, 61

douglasii, 63, 64

durangi, 71

emotus, 64

enixus, 71

ericaeus, 69

eximius, 71

extenuatus, 61

extimus, 71

fisheri, 65

flavidus, 64

fossor, 64
9 6 • smithsonian contributions to zoology

Thomomys (continued)

fulvus, 60, 62, 64, 65

fuscus, 63, 65, 66, 67

glacialis, 70

goldmani, 66

grahamensis, 64

guadalupensis, 61

hesperus, 66

howelli, 61

hualpaiensis, 61

idahoensis, 66, 68

imitabilis, 62

incensus, 70

incomptus, 62

intermedius, 65

internatus, 62

kaibabensis, 64

kelloggi, 70

lachuguilla, 60, 66

laticeps, 62, 66

latirostris, 66

lenis, 71

leucodon, 66

levidensis, 62

levis, 64

limitaris, 66

limosus, 66, 67

loringi, 66

louiei, 70

madrensis, 69, 71, 72

magdalenae, 67

martinensis, 72

mazama, 63, 64, 66, 67, 68, 

69, 70

mearnsi, 61, 64, 67, 71

medius, 70

melanops, 67

mewa, 67

minimus, 62

minor, 62

modicus, 65

monticola, 67

moorei, 64

muralis, 67

musculus, 72

mutabilis, 65

myops, 67

nasicus, 67

navus, 66

nebulosus, 70

neglectus, 67

nelsoni, 67

nevadensis, 67, 68

nicholi, 62

niger, 68

nigricans, 63

ocius, 63

operarius, 68

operosus, 62

optabilis, 62

opulentus, 62

oregonus, 63

orizabae, 68

osgoodi, 68

parowanensis, 64

parviceps, 69

parvulus, 62

pascalis, 60

patulus, 62

pectoralis, 68

peramplus, 65

perditus, 68

peregrinus, 68

perpallidus, 68, 70

perpes, 60

pervagus, 61

pervarius, 62

phasma, 65

pinalensis, 63

pinetorum, 67

planirostris, 62, 63, 68

potosinus, 72

pryori, 68

pugetensis, 70

pusillus, 65

pygmaeus, 68

quadratus, 69

relicinus, 70

retrorsus, 70

russeolus, 63

saturatus, 66

shawi, 64

sheldoni, 69

simulus, 69

sinaloae, 69

sonoriensis, 72

spatiosus, 61

sturgisi, 69

suboles, 65

subsimilis, 65

supernus, 72

tacomensis, 64

talpoides, 63, 64, 65, 66, 67, 

68, 69, 70, 71

tenellus, 70

texensis, 65

tolucae, 72

townsendii, 67, 68, 71

trivialis, 70

tumuli, 70

uinta, 71

umbrinus, 64, 65, 66, 67, 68, 

69, 71, 72

virgineus, 63

vulcanius, 72

whitmani, 71

winthropi, 63

yakimensis, 69

yelmensis, 64

zacatecae, 72

thomsoni

Iomys, 14

tiomanensis

Ratufa, 20

tolucae

Sciurus, 24

Thomomys, 72 

torridus

Heterogeomys, 58

Heteromys, 50

Liomys, 50

Orthogeomys, 58

townsendii

Citellus, 34

Eutamias, 29

Tamias, 30

Thomomys, 67, 68, 71

Urocitellus, 36

trichopus

Zygogeomys, 72

tridecemlineatus

Citellus, 13

Ictidomys, 13

Spermophilus, 13

trinitatis

Callospermophilus, 10

trivialis

Thomomys, 70

tropicalis

Geomys, 58

trotteri

Callosciurus, 7

Sciurus, 7

truei

Sciurus, 24

tularensis

Dipodomys, 46

Perodipus, 46

tumuli

Thomomys, 70

tuza

Geomys, 58

tylorhinus

Cratogeomys, 56

Platygeomys, 56

ubericolor

Sciurus, 10

uinta

Thomomys, 71

umbrinus

Eutamias, 29

Tamias, 29, 31

Thomomys, 64, 65, 66, 67, 

68, 69, 71, 72

umbrosus

Eutamias, 26

Tamias, 26

underwoodi

Sciurus, 21

undulatus

Heliosciurus, 13

Sciurus, 13

Urocitellus

ablusus, 34

armatus, 34

artemesiae, 33

beldingi, 34, 36

brunneus, 34, 35

canus, 33, 35

columbianus, 33

elegans, 33, 35

endemicus, 35

idahoensis, 33

kennicottii, 34, 35

kodiacensis, 36

mollis, 33, 35

nebulicola, 33

nevadensis, 33

oregonus, 36

osgoodi, 36

parryii, 33, 34, 35, 36

plesius, 35

ruficaudus, 33

stejnegeri, 34

townsendii, 36

washingtoni, 34

utah

Citellus, 16

Otospermophilus, 16

utahensis

Dipodomys, 47

Eutamias, 27

Perodipus, 47

Tamias, 27

vallicola

Eutamias, 26

Tamias, 26

variabilis

Sciurus, 24

variegatoides

Sciurus, 21, 23, 24

variegatus

Citellus, 16

Otospermophilus, 16, 17

Sciurus, 8

varius

Platygeomys, 56

ventorum

Sciurus, 32

Tamiasciurus, 32

venustulus

Microsciurus, 15

venustus

Dipodomys, 47

Perodipus, 47

veraecrucis

Liomys, 50

vestitus

Tamiops, 33

vigilis

Citellus, 33

virgineus

Thomomys, 63

vittata

Ratufa, 20

vittatula

Ratufa, 20

vittatus

Callosciurus, 7, 8, 9, 10

Sciurus, 10

vivatus

Microsciurus, 15

vivax

Sciurus, 22
number 640

• 9 7

volans

Glaucomys, 12

Pteromys, 18

Sciuropterus, 12

vulcani

Orthogeomys, 59

vulcanius

Thomomys, 72

vulgaris

Sciurus, 23, 24

warreni

Marmota, 15

wasatchensis

Tamiasciurus, 32

washingtoni

Citellus, 34

Urocitellus, 34

washoensis

Citellus, 33

whitmani

Thomomys, 71

winthropi

Thomomys, 63

wisconsinensis

Geomys, 57

Xerospermophilus

annectens, 36

canescens, 36, 37

cryptospilotus, 36

major, 37

mohavensis, 36

neglectus, 36

obsoletus, 36

pallescens, 36

perotensis, 36

pratensis, 37

spilosoma, 36, 37

tereticaudus, 36, 37

yakimensis

Spermophilus, 35

Thomomys, 69

yelmensis

Thomomys, 64

yucatanensis

Heterogeomys, 58

Orthogeomys, 58

Sciurus, 22, 25

yukonensis

Glaucomys, 12

Sciuropterus, 12

yunanensis

Petaurista, 17

zacatecae

Chaetodipus, 40

Perognathus, 40

Thomomys, 72

zaphaeus

Glaucomys, 12

Sciuropterus, 12

zinseri

Platygeomys, 57

zonalis

Heteromys, 48

zuniensis

Cynomys, 11

Zygogeomys

tarascensis, 72

trichopus, 72

zygomaticus

Dipodomys, 46
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requirements for smithsonian series publication