A Checklist of the North and

Middle American Crayfishes

(Decapoda: Astacidae and

Cambaridae)

HORTON H. HOBBS, JR.

m

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SMITHSONIA N CONTRIBUTION S T O ZOOLOG Y • NUMBE R 166

A Checklist of the North and

Middle American Crayfishes

(Decapoda: Astacidae and Cambaridae)

Horton H. Hobbs, Jr.

SMITHSONIAN INSTITUTION PRESS

City of Washington

1974
ABSTRACT

Hobbs, Horton H., Jr. A Checklist of the North and Middle American Crayfishes (Decapoda: Astacidae and Cambaridae). Smithsonian Contributions to

Zoology, number 166, 161 pages, 294 figures, 1974.—Included in this alphabetical list are all of the North and Middle American crayfishes, including two

fossil species, that were described prior to 1 January 1973. Within the treatment

of each specific or subspecific taxon is a synonymy in which are cited alterations

in generic and subgeneric assignments, erroneous spellings, and synonyms.

References also are given to recent descriptions, discussions, and keys. The composition and locations of type-series are cited, as are the type-locality, range,

and habitat of each species and subspecies. Nomina nuda that have been neither

validated nor synonymized with other crayfish taxa, together with one name

suppressed by the International Commission on Zoological Nomenclature, are

listed separately. Illustrations, which are arranged in related species groups,

are provided for each of the 294 recognized extant species and subspecies.

OFFICIAL PUBLICATION DATE is handstamped in a limited number of initial copies and is recorded

in the Institution's annual report, Smithsonian Year. SI PRESS NUMBER 4984. SERIES COVER DESIGN:

The coral Montastrea cavernosa (Linnaeus).

Library of Congress Cataloging in Publication Data.

Hobbs, Horton Holcombe, Jr., 1914—

A checklist of the North and Middle American crayfishes (Decapoda: Astacidae and Cambaridae).

(Smithsonian contributions to zoology no. 166)

1. Astacidae. 2. Cambaridae. 3. Crustacea—North America. 4. Crustacea—Central America.

I. Title. II. Series: Smithsonian Institution. Smithsonian contributions to zoology, no. 166.

QL1.S54 no. 166 [QL444.M33] 591'.08s [595'.3841] 73-16027

For tale by the Superintendent of Documents, U.S. Government Printing Office

Washington, D.C. 20402 • Price $2.50 (paper cover)
Contents

Page

Introduction 1

Nomina Nuda neither Validated nor Synonymized with Other

Crayfish Taxa 4

Suppressed Name 4

Species Inquirenda 4

The Checklist 5

Family Astacidae Latreille, 1802-1803 5

Family Cambaridae Hobbs, 1942 7

Subfamily Cambarellinae Laguarda, 1961 7

Subfamily Cambarinae Hobbs, 1942 9

Literature Cited 68

Explanation of Illustrations 80

Index 154

iii

A Checklist of the North and

Middle American Crayfishes

(Decapoda: Astacidae and Cambaridae)

Horton H. Hobbs, Jr.

Introduction

Preparation of this checklist of the crayfishes

of North and Middle America began late in the

1940's but its presentation for publication has been

repeatedly postponed because of the almost continuous appearance of descriptions of new taxa.

The most recent checklist appeared more than half

a century ago (Faxon, 1914); since then the number of described species and subspecies has nearly

tripled. As a result of this and because most descriptions subsequent to Faxon's checklist have

been published in short papers scattered among

many journals, a revised compilation is needed.

This list is being published despite the fact that

there remain many undescribed species; indeed,

descriptions of at least a dozen species are now

in preparation or in press.

To the best of my knowledge, all of the North

and Middle American crayfishes (including two

fossil species) that were described prior to 1 January 1973 are accounted for in this summary (most

also are incorporated in a recent key to the crayfish

fauna of the same area, Hobbs, 1972b). While those

few species that have been described since that date

are not treated, combinations involving older

names have been updated to June 1973.

Preceding the alphabetical list of species is a

list of nomina nuda that have not been validated,

Horton H. Hobbs, Jr., Department of Invertebrate Zoology,

National Museum of Natural History, Smithsonian Institution,

Washington, D.C. 20560.

together with one name that has been suppressed

by the International Commission on Zoological

Nomenclature. Included in the treatment of each

specific or subspecific taxon is a synonymy in

which are cited alterations in generic and subgeneric assignments, erroneous spellings, and synonyms; in only rare instances are references to

misidentifications listed. References also are given

to recent descriptions, discussions, and keys. The

composition and locations of the type-series are

cited and, where possible, the sexes of the syntypes

and primary-types are indicated. In instances in

which the type designations included a holotypic

male, form I, and a "paratypic male, form II"

(the latter clearly separated from the remainder

of the paratypic-series), the paratypic male has

been referred to here as "morphotype," the name

enclosed in quotation marks. In citing the types of

those taxa for which a morphotype was designated

by the describer, the quotation marks are omitted.

The type-locality, range, and habitats are given

for each taxon. The designations "primary, secondary, and tertiary burrowers" follow Hobbs

(1942b:20): primary burrowers are those largely

restricted to burrows; secondary burrowers are

those that generally occupy burrows but often

wander into open water during times when the

mouths of the burrows are flooded; and tertiary

burrowers are those that burrow "only in periods

of drought or occasionally, but not necessarily,

during the breeding season."

Insofar as possible the illustrations, which are

1
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

arranged in related species groups, are based upon

types of the species; and in the comparatively few

instances in which types were not available,

specimens were chosen from, or as close to, the

type-locality as possible (see Explanation of Illustrations).

LOCATIONS OF TYPES.—The following abbreviations are used for museums or collections in which

specimens of one or more type-series have been

deposited:

Alabama Mus. Nat. Hist.: Alabama Museum of Natural

History, Tuscaloosa, Alabama.

AMNH: American Museum of Natural History, New York,

New York.

AMS: Australian Museum, Sydney, Australia.

ANSP: Academy of Natural Sciences of Philadelphia, Pennsylvania.

BMNH: British Museum, Natural History, London, England.

BSNH: Boston Society of Natural History, Boston, Massachusetts.

Chas. M.: Charleston Museum, South Carolina.

CM: Carnegie Museum, Pittsburgh, Pennsylvania.

GM: Museum d'Histoire Naturelle, Geneva, Switzerland.

IBM: Instituto de Biologia de la Universidad Nacional

Autdnoma de Mexico, Mexico City, Mexico.

111. St. Lab. Nat. Hist.: Illinois State Laboratory of Natural

History, Urbana, Illinois.

La. Polytechnic Inst.: Louisiana Polytechnic Institute, Ruston, Louisiana.

MCZ: Museum of Comparative Zoology, Harvard University,

Cambridge, Massachusetts.

MHNP: Museum National d'Histoire Naturelle, Paris,

France.

Mus. Washburn Coll.: Museum of Washburn College, now

Department of Biology, Washburn University, Topeka,

Kansas.

Mus. Zool. Torino: Instituto e Museo di Zoologia, Delia

Universita di Torino, Italy.

NMV: National Museum of Victoria, Melbourne, Australia.

OSM: Ohio State Museum, Columbus, Ohio.

PM: Peabody Museum of Natural History, Yale University,

New Haven, Connecticut.

RR: Collection of Rendell Rhoades, Ashland College, Ashland, Ohio.

TU: Department of Zoology, Tulane University, New

Orleans, Louisiana.

UF: Department of Zoology, University of Florida, Gainesville, Florida.

USNM: National Museum of Natural History (formerly

under United States National Museum), Smithsonian Institution, Washington, D.C.

UZM: Universitetets Zoologiske Museum, Krystalgade, K0benhavn K., Denmark.

Wiirzburg Mus.: Wiirzburg Museum, Wiirzburg, Germany.

ZBM: Zoologisches Museum der Humboldt-Universitat,

Berlin, East Germany.

ZIAS: Zoological Institute, Academy of Sciences, Leningrad,

U.S.S.R.

In addition, type-specimens of a few species are

in the collections of the following: Troy C. Dorris,

Oklahoma State University, Stillwater, Oklahoma;

Max R. Matteson, University of Illinois, Urbana,

Illinois; Leslie Hubricht, Meridian, Mississippi;

Hugo L. James, University of Bridgeport, Bridgeport, Connecticut; Rollin D. Reimer, Texas A &

M, College Station, Texas; Joe B. Black, McNeese

State University, Lake Charles, Louisiana; Joseph

F. Fitzpatrick, Jr., Randolph-Macon Woman's

College, Lynchburg, Virginia; Jerry G. Walls,

Hightstown, New Jersey; Raymond W. Bouchard,

University of Tennessee, Knoxville; Rudolph Prins,

Western Kentucky University, Bowling Green; H.

H. Hobbs III, Christopher Newport College, Newport News, Virginia; and Coe College, Cedar

Rapids, Iowa.

Three large and important crayfish collections—

those of the University of Michigan Museum of

Zoology, the University of Kansas Museum of

Natural History, and the personal collection of

the author—have been transferred to the National

Museum of Natural History, Smithsonian Institution, and the type-specimens cited by many

previous authors as having been deposited in those

collections are now in the Smithsonian crayfish

collection.

A BRIEF HISTORY OF THE NOMENCLATURE OF

NORTH AMERICAN CRAYFISHES.—Prior to the publication of Girard's (1852) revision of the North

American Astaci, which treated 20 species, all of

the North American crayfishes were assigned to

the genus Astacus. Girard was the first to employ

Erichson's (1846) subgeneric designation Cambarus as a generic name, and with its adoption by

Hagen (1870) in his Monograph of the North

American Astacidae and the clear delineation

drawn between Cambarus, in the Atlantic watershed, and Astacus, largely restricted to the Pacific

slope, a generic stability was established that remained unchanged until 1933. The first checklist

of holarctic crayfishes was that of Faxon (1884),

in which 60 North American species were recognized. Faxon revised the Astacidae in 1885 but

made few changes in the "species groups" originally

proposed by Hagen. With some reshuffling of

species, Ortmann (1905c, 1905d, 1906a) proposed

the recognition of six subgenera of Cambarus:
NUMBER 166

Cambarus, Bartonius, Faxonius, Cambarellus, Procambarus, and Paracambarus. Fowler (1912), in

reviewing the Crustacea of New Jersey, pointed

out that Ortmann (1905c) overlooked Faxon's

(1898) selection of Astacus Bartonii Fabricius as

the type-species of Cambarus. Ortmann had chosen

that crayfish as the type-species of the subgenus

Bartonius, and Astacus Blandingii Harlan as that

of his subgenus Cambarus. Thus the name Bartonius was shown to be a junior objective synonym

of Erichson's Cambarus, and Ortmannicus was

proposed to replace Ortmann's subgenus Cambarus.

In his checklist of the crayfishes Faxon (1914)

included 110 species and subspecies from North

America but recognized no subgenera. An alteration in the generic assignment of the North American crayfishes had its inception with Creaser

(1933a) who elevated Ortmann's subgenus Faxonius to generic rank. Not until 1942, however, was

there a major departure from the generic structure

that had been established by Girard (1852) and

Hagen (1870).

Creaser (1933b) erected the monotypic subgenus

Faxonella to receive Cambarus clypeatus Hay. This

subgenus was questionably accorded generic status

by Creaser (1962) and was clearly so treated by

Fitzpatrick (1963).

Lyle (1938) designated the subgenus Girardiella

to encompass Faxon's (1914) Group II.

Hobbs (1942a) proposed the elevation of Ortmann's subgeneric groupings of Cambarus to generic rank, recognizing the subfamilies Astacinae

and Cambarinae, the latter comprising the following genera: Cambarus, Procambarus (including

Fowler's Ortmannicus and Lyle's Girardiella), Orconectes (= Ortmann's Faxonius), Cambarellus, Paracambarus, and Troglocambarus, the latter described

as new. Orconectes Cope (1872) was considered to

be a subjective senior synonym of Ortmann's

Faxonius.

Villalobos (1955) recognized the Cambaroidinae

as a subfamily of the Astacidae, and Laguarda

(1961) designated a fourth subfamily, the Cambarellinae. Fitzpatrick and Payne (1968) added the

genus Hobbseus, and Hobbs (1969b), in revising

the genus Cambarus, proposed the genus Fallicambarus. In the latter revision, 10 subgenera (see

below) of Cambarus were recognized. One of them,

Barbicambarus, was elevated to generic rank by

Bouchard (1972).

In 1972, Hobbs (1972a) recognized 16 subgenera

of Procambarus, resurrecting the subgenera Ortmannicus (Fowler, 1912) and Girardiella (Lyle,

1938) and returning Paracambarus to subgeneric

status in Procambarus. Later in the same year,

Hobbs' (1972b) key to most of the crayfishes, included herein, was made available.

Finally, in a synopsis of the families and genera

of crayfishes, Hobbs (1974) proposed a restructuring of the Astacuran infraorder Astacidea as follows:

INFRAORDFR ASTACIDEA

Superfamily Nephropoidea Dana, 1852 (lobsters and marine

allies)

Superfamily Parastacoidea Huxley, 1879 (crayfishes of the

Southern Hemisphere)

Superfamily Astacoidea Latreille, 1802-1803

Family Astacidae Latreille, 1802-1803

Genus Astacus Fabricius, 1775 (Europe and western

Asia)

Subgenus Astacus Fabricius, 1775

Subgenus Pontastacus Bott, 1950

Genus Austropotamobius Skorikov, 1908 (Europe)

Subgenus Austropotamobius Skorikov, 1908

Subgenus Atlantoastacus Bott, 1950

Genus Pacifastacus Bott, 1950

Family Cambaridae Hobbs, 1942a

Subfamily Cambarinae Hobbs, 1942a

Genus Barbicambarus Hobbs, 1969b

Genus Cambarus Erichson, 1846

Subgenus Cambarus Erichson, 1846

Subgenus Aviticambarus Hobbs, 1969b

Subgenus Depressicambarus Hobbs, 1969b

Subgenus Erebicambarus Hobbs, 1969b

Subgenus Hiaticambarus Hobbs, 1969b

Subgenus Jugicambarus Hobbs, 1969b

Subgenus Lacunicambarus Hobbs, 1969b

Subgenus Puncticambarus Hobbs, 1969b

Subgenus Veticambarus Hobbs, 1969b

Genus Fallicambarus Hobbs, 1969b

Genus Faxonella Creaser, 1933b

Genus Hobbseus Fitzpatrick and Payne, 1968

Genus Orconectes Cope, 1872

Genus Procambarus Ortmann, 1905d

Subgenus Procambarus Ortmann, I905d

Subgenus Acucauda Hobbs, 1972a

Subgenus Austrocambarus Hobbs, 1972a

Subgenus Capillicambarus Hobbs, 1972a

Subgenus Girardiella Lyle, 1938

Subgenus Hagen ides Hobbs, 1972a

Subgenus Leconticambarus Hobbs, 1972a

Subgenus Lonnbergius Hobbs, 1972a

Subgenus Mexicambarus Hobbs, 1972a

Subgenus Ortmannicus Fowler, 1912
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Subgenus Paracatnbarus Ortmann, 1906a

Subgenus Pennides Hobbs, 1972a

Subgenus Remoticambarus Hobbs, 1972a

Subgenus Scapulicambarus Hobbs, 1972a

Subgenus Tenuicambarus Hobbs, 1972a

Subgenus Villalobosus Hobbs, 1972a

Genus Troglocambarus Hobbs, 1942a

Subfamily Cambarellinae Laguarda, 1961

Genus Cambarellus Ortmann, 1905c

Subfamily Cambaroidinae Villalobos, 1955 (eastern Asia

and Japan)

Genus Cambaroides Faxon, 1884

ACKNOWLEDGMENTS.—Among the many persons

to whom I am indebted for assistance in compiling

this checklist are Dr. J. Forest, Museum National

d'Histoire Naturelle, Paris; the late Dr. H. Gisin,

Museum d'Histoire Naturelle, Geneva; Dr. H.-E.

Gruner, Zoologisches Museum, Berlin; Dr. H. W.

Levi, Museum of Comparative Zoology, Cambridge, Massachusetts; Dr. Paul W. Parmalee,

Illinois State Museum, Springfield; Dr. J. J.

Parodiz, Carnegie Museum, Pittsburgh, Pennsylvania; Dr. N. E. Pearson, Butler University, Indianapolis, Indiana; Dr. Philip W. Smith, Illinois

State Natural History Survey, Urbana; Dr. David

H. Stansbery, Ohio State Museum, Columbus; Dr.

Y. Starobogatov, Zoological Institute, Academy of

Sciences, U.S.S.R., Leningrad; Dr. J. Taubucci,

Instituto e Museo di Zoologia, Torino, Italy; Dr.

S. L. Tuxen, Universitetets Zoologiske Museum,

Copenhagen; Dr. Alejandro Villalobos, Universidad Nacional Autonoma de Mexico, Mexico City;

and Dr. J. C. Yaldwyn, formerly of the Australian

Museum, Sydney. I am also grateful to the following persons who have read or assisted in checking

certain parts of the manuscript: Thomas E. Bowman, Fenner A. Chace, Jr., Martha R. Cooper,

Margaret A. Daniel, Joseph F. Fitzpatrick, Jr.,

Georgia B. Hobbs, Richard L. Hoffman, and Mary

H. Moulton.

Nomina Nuda neither Validated nor Synonymized

with Other Crayfish Taxa

Catnbarus (Cambarus) lobdelli Lyle, 1938:76.

Cambarus (Faxonius) creaseri Lyle, 1938:76.

Cambarus (Girardiella) hagenianus carri Lyle, 1938:76.

Cambarus (Girardiella) hagenianus evansi Lyle, 1938:76.

Cambarus (Girardiella) hagenianus forestae Lyle, 1938:76.

Cambarus (Ortmannicus) cookae Lyle, 1938:76.

Cambarus (Ortmannicus) evictus Lyle, 1938:76.

Cambarus schmittii Penn, 1941:8 [questionably synonymized

with Cambarellus puer Hobbs].

Cambarus lucifugus Packard, 1888:82 [lapsus for Crangonyx

lucifugus Hay].

Suppressed Name

Astacus oreganus Randall, 1840:138. [Suppressed by Opinion

855 of the International Commission on Zoological Nomenclature, 1968:84.]

Species Inquirenda

Cambarus primaevus Packard

Cambarus primaevus Packard, 1880:222; 1881a:832; 1881b:391.

Cambarus primoevus Villalobos, 1955:14 [erroneous spelling]-

TYPES.—Not known to be extant.

TYPE-LOCALITY.—Lower Tertiary beds in the

Bear River Valley, "75 to 100 miles a little west

of north of Evanston, Wyo., and near the Utah

line" (Packard, 1881b:391).

RANGE.—Known only from the type-locality.

REMARKS.—It is highly doubtful that this crayfish is a member of the genus Cambarus as presently defined; and, on the basis of available

information, it cannot be assigned to any of the

subgenera. Possibly, it is not even a member of the

Cambaridae! The original combination of Packard is retained because, until the type specimens

or additional material of the species become available, no assessment of its relationships to other

crayfishes can be made.

Procambarus wiegmanni (Erichson)

Astacus (Cambarus) Wiegmanni Erichson, 1846:99.

Cambarus Wiegmanni.—Hagen, 1870:54.

Cambarus wiegmanni.—Faxon, 1890:621.

Procambarus wiegmanni.—Hobbs, 1942a:342 [by implication].

Procambarus weigmanni.—Villalobos, 1949:338 [erroneous

spelling].

Procambarus Weigmanni.—Villalobos, 1950:381-383 [erroneous spelling].

TYPES.—Type lost from BMNH.

TYPE-LOCALITY.—Mexico.

RANGE.—Unknown.

HABITAT.—Unknown.

REMARKS.—The identity of this crayfish has not

been recognized with certainty since it was described. Villalobos (1950) reviewed its history and

designated Erichson's name a nomen nudum. In-
NUMBER 166

asmuch as Erichson diagnosed and described the

species and stated that it occurs in Mexico, it cannot be considered a nomen nudum.

The species was referred to the genus Procambarus on the basis of Erichson's (1846:99) statement, "Des zweite Glied am dritten und am vierten

Beinpaar beim Mannchen mit einem hakenformigen Fortsatz." Only two crayfish genera are

known to occur in Mexico, and, of the two, this

statement could apply only to a member of the

genus Procambarus.

The Checklist

Family ASTACIDAE Latreffle, 1802-1803

Genus Pacifastacus Bott

Astacus.—Dana, 1852:522.

Cambarus.—Girard, 1852:90.

Cambrus Girard, 1852:90 [erroneous spelling].

Potamobius.—Ortmann, 1902:286.

Pacifastacus Bott, 1950:24. [Type-species by original designation, Astacus klamathensis Stimpson, 1857a:87. Gender:

masculine.]

Pasifastacus Sanders, 1967:33 [erroneous spelling].

Pacifastaculus Kamita, 1973:123 [erroneous spelling].

Pacifastacus chenoderma (Cope),

new combination

Astacus chenoderma Cope, 1871:606.—Rathbun, 1926:130, pi.

29: fig. 7; pis. 31-34.

?Astacus subgrundialis Cope, 1871:605. [Types not extant.

Type-locality, "fresh water deposit in the Territory of

Idaho, near Hot Springs Mountain."]

?Astacus breviforceps Cope, 1871:606. [Types not extant.

Type-locality, same as that for Astacus chenoderma, see

below.]

TYPES.—Syntype, USNM 353343; other syntypes

probably lost.

TYPE-LOCALITY.—"Catherine's creek, Idaho." According to an annotation in a reprint, this is "a

local name for the upper part of Castle Creek,

Owyhee County." A Pliocene deposit.

RANGE.—Miocene and Pliocene deposits in Idaho

and Oregon.

REMARKS.—Rathbun (1926) redescribed and figured Cope's A. chenoderma, suggesting that both A.

breviforceps and A. subgrundialis are probable

synonyms. Since the types are lost and accounts

of specimens assignable to either have not appeared

since Miss Rathbun's review of the species, I have

followed her in including these species as probable

synonyms of Pacifastacus chenoderma. The species

is transferred to the genus Pacifastacus because the

rostrum with as many as seven pairs of marginal

spines is a feature foreign to members of the Eurasian Astacidae but strongly resembles that of four

members of the American astacid genus Pacifastacus: P. connectens, P. fortis, P. gambelii, and P.

nigrescens.

Pacifastacus connectens (Faxon)

FIGURE 1

Astacus gambelii connectens Faxon, 1914:360, pi. 7: figs. 6, 10;

pi. 10.

Pacifastacus gambelii connectens.—Bott, 1950:24 [by implication].

Pacifastacus gambeli connectens.—Riegel, 1959:43.

Pacifastacus gambelii connectans.—Holt, 1960:61 [erroneous

spelling].

Pacifastacus connectens.—Hobbs, 1972b:22, figs. 12d, 14a.

TYPES.—Syntypes, USNM 23096 (3 $ , 9 ), MCZ

73285 ($).

TYPE-LOCALITY.—Snake River, Upper Salmon

Falls, Logan County, Idaho.

RANGE.—Idaho and Oregon.

HABITAT.—Streams.

Pacifastacus fortis (Faxon)

FIGURE 3

Astacus nigrescens fortis Faxon, 1914:360, pi. 7: figs. 5, 9; pi.

9: fig. 2. ,

Pacifastacus nigrescens fortis.—Bott, 1950:24 [by implication].

Pacifastacus fortis.—Hobbs, 1972b:23, figs. 13c, 14c.

TYPES.—Syntypes, USNM 44404 ($ , 2 9), MCZ

7383 (3, 9).

TYPE-LOCALITY.—Fall River, Fall City Mills,

Shasta County, California.

RANGE.—Known only from Shasta County, California.

HABITAT Streams.

Pacifastacus gambelii (Girard)

FIGURE 2

Cambrus Gambelii Girard, 1852:90 [lapsus for Cambarus].

Cambarus Gambeli.—Hagen, 1870:31.
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Astacus Gambeli Hagen, 1870:28.

Astacus Gambelii.—Hagen, 1870:90, pi. 1: figs. 97, 98; pi. 3:

fig. 170; pi. 11.

Astacus (Astacus) Gambelii.—Faxon, 1884:152 [by implication].

Astacus (Astacus) gambelii.—Faxon, 1898:666 by [implication].

Potamobius (Potamobius) gambeli.—Ortmann, 1902:286 [by

implication].

Astacus gambelii gambelii.—Faxon, 1914:360 [by implication].

Astacus gambelii Rathbun, 1926:133.

Pacifastacus gambelii gambelii Bott, 1950:24 [by implication].

Astacus gambeli.—Pennak, 1953:461.

Pacifastacus gambelii.—Kozloff, 1955:156 Hobbs, 1972b:22,

figs. 13a, 14b.

Pacifastacus gambeli.—Riegel, 1959:29, 43-44, figs. 3c, 8.

TYPES.—Syntypes, ANSP 1531 (10 + , mostly

fragments), 1532 (5 specimens).

TYPE-LOCALITY.—"California."

RANGE.—Pacific slope and in the Missouri River

drainage in California (?), Idaho, Montana, Nevada, Oregon, Utah, Washington, and Wyoming.

HABITAT.—Streams and lakes.

Potamobius (Potamobius) leniusculus.—Ortmann, 1902:286

[by implication].

Potamobius leniusculus.—Roberts, 1944:363.

Pacifastacus leniusculus.—Bott, 1950:24 [by implication]

Riegel, 1959:39, figs. 1D-H, 3A, 6, 10.

"Astacus" (Pacifastacus) leniusculus Brinck, 1968:85.

Pacifastacus leniusculus leniusculus Hobbs, 1972b:21, figs.

12a, 14e.

Pacifastaculus leniuscus.—Kamita, 1973:123 [erroneous spellings].

TYPES.—Syntypes, USNM 2019 {$), 2161 ($),

44408 (S).

TYPE-LOCALITY.—Columbia River and Puget

Sound, Washington.

RANGE.—British Columbia, California, Idaho,

Nevada, Oregon, and Washington. Introduced into

Sweden (Svardson, 1965:92) and Japan (Kamita,

1970:140).

HABITAT.—Streams and lakes.

REMARKS.—Astacus leniusculus was placed on the

Official List of Specific Names in Zoology, name

number 2244, in Opinion 855 of the International

Commission on Zoological Nomenclature (1968:84).

Pacifastacus leniusculus klamathensis (Stimpson)

FIGURE 6

Astacus Klamathensis Stimpson, 1857a:87.—Hagen, 1870:93,

pi. 3: fig. 169.—Faxon, 1885a:131, pi. 6: figs. 1, 2.

Astacus (Astacus) Klamathensis.—Faxon, 1884:151 [by implication].

Astacus klamathensis Faxon, 1890:634.

Potamobius (Potamobius) klamathensis.—Ortmann, 1902:286.

Pacifastacus klamathensis Bott, 1950:24 [by implication].—

Riegel, 1959:41, figs. 3B, 7, 10.

TYPES.—Not extant; "probably destroyed in the

Chicago fire in 1871" (Faxon, 1914:409).

TYPE-LOCALITY.—Lake Klamath, Klamath County, Oregon.

RANGE.—Washington south to central California.

HABITAT.—Lakes and cold, swift streams.

Pacifastacus leniusculus leniusculus (Dana)

FIGURE 5

Astacus leniusculus Dana, 1852:522, 524; 1855, pi. 33: fig.

la,b,—Hagen, 1870:94.

Astacus laeviusculus.—Lockington, 1878:304 [erroneous spelling]-

Astacus (Astacus) leniusculus.—Faxon, 1884:151 [by implication]; 1885a: 132, pi. 6: fig. 4.

Pacifastacus leniusculus trowbridgii (Stimpson)

FIGURE 7

Astacus Trowbridgii Stimpson, 1857a:87; 1857b:493 Hagen,

1870:93, pi. 3: fig. 171; pi. 10.

Astacus (Astacus) Trowbridgii Faxon, 1884:152 [by implication].

Astacus (Astacus) trowbridgii Faxon, 1898:666 [by implication].

Astacus trowbridgii.—Hay, 1899b:958.

Potamobius (Potamobius) trowbridgei.—Ortmann, 1902:286

[by implication].

Pacifastacus trowbridgii Bott, 1950:24 [by implication].—

Riegel, 1959:30, fig. lc-E.—Mason, 1970a:37, figs. 1-16;

1970b:969, figs. 1-7.

Astacus trowbridgi.—Pennak, 1953:461.

Astacus trowbridgei.—Pilgrim, 1960:249 [erroneous spelling].

Pacifastacus trowbridgeii Sanders, 1967:33 [erroneous spelling]-

Pasifastacus trowbridgeii.—Sanders, 1967:33 [erroneous spelling]-

Pacifastacus leniusculus trowbridgii.—Hobbs, 1972b:22, figs.

5a, 12b.

Pacifastacus trowbridgi Goldman, 1973:106 [erroneous spelling] •

Pacifastaculus trowbridgii Kamita, 1973:123 [erroneous

spelling].

TYPES.—Syntypes, USN M 2080 (24,2°) , MCZ

3510 (£ ), BSNH, PM.
NUMBER 166

TYPE-LOCALITY.—Columbia River above Astoria,

Oregon.

RANGE.—British Columbia, California, Idaho,

Oregon and Washington. Introduced into Japan

(Kamita, 1970:148).

HABITAT.—Streams and lakes.

TYPE-LOCALITY.—Potosf, Nuevo Le6n, 23 km

northwest of Galeana, Mexico.

RANGE.—Known only from the type-locality.

HABITAT.—"Colecta realizada en un deposito de

agua de una cuenca cerrada, situada en la parte

None de Valle Salado" (Villalobos, 1952:531).

Padfastacus nigrescens (Stimpson)

FIGURE 4

Astacus nigrescens Stimpson, 1857a:87; 1857b:492.—Hagen,

1870:92, pi. 3: fig. 168.

Astacus (Astacus) nigrescens.—Faxon, 1884:152 [by implication].

Astacus nigrescens nigrescens.—Faxon, 1914:360 [by implication].

Potamobius (Potamobius) nigrescens.—Ortmann, 1902:286

[by implication].

Padfastacus nigrescens nigrescens. Bott, 1950:24 [by implication].

Padfastacus nigrescens.—Riegel, 1959:44, figs. 3D, 9, 10.—

Hobbs, 1972b:23, figs. 13b, 14d.

Padfastacus nigresens.—Goldman, 1973:107 [erroneous spelling]-

TYPES.—Not extant; "probably destroyed in Chicago fire in 1871" (Faxon, 1914:408).

TYPE-LOCALITY.—Vicinity of San Francisco, California.

RANGE.—Vicinity of San Francisco, California.

HABITAT.—Streams.

REMARKS.—This species is almost, if not already,

extinct.

Family CAMBARTOAE Hobbs, 1942

Subfamily CAMBARELLINAE Laguarda, 1961

Genus Cambarellus Ortmann

Cambarellus Ortmann, 1905c: 106. [Type-species by original

designation, Cambarus Montezumae Saussure, 1857a: 102.

Gender: masculine.]

Cambarella Carlisle and Knowles, 1959:50 [erroneous spelling].

Cambarellus alvarezi Villalobos

FIGURE 14

Cambarellus alvarezi Villalobos, 1952:525, pi. 1: figs. 1-7;

pi. 2: figs. 1-5—Hobbs, 1972b:27, fig. 15f.

TYPES.—Holotype and morphotype, IBM 1-1952

(i I, $ II); paratypes, USNM.

Cambarellus areolatus (Faxon)

FIGURE 16

Cambarus Montezumae var. areolata Faxon, 1885a: 123.

Cambarus montezumae areolatus.—Faxon, 1898:661, pi. 66:

fig. 2.

Cambarus (Cambarellus) montezumae areolatus.—Ortmann,

1906a:23.

Cambarellus montezumae areolatus.—Hobbs, 1942a:350 [by

implication].—Villalobos, 1955:276.

Cambarellus areolatus.—Hobbs, 1972b:27, figs. 16b, 17b.

TYPES.—Syntypes, MCZ 3650 (<J I, 3 9 , ill) .

TYPE-LOCALITY.—Near Parras, Coahuila, Mexico.

RANGE.—Known only from the type-locality.

HABITAT.—Unknown.

Cambarellus chapalanus (Faxon)

FIGURE 15

Cambarus chapalanus Faxon, 1898:661, pi. 67: figs. 1, 2.

Cambarus (Cambarellus) chapalanus.—Ortmann, 1905c: 106.

Cambarus montezumae chapalanus.—Faxon, 1914:371.

Cambarus chapalamus.—Van Straelen, 1942:5 [erroneous

spelling].

Cambarellus montezumae chapalanus.—Hobbs, 1942a:350 [by

implication].

Cambarellus chapalanus.—Villalobos, 1955:277.—Hobbs,

1972b:28, figs. 16c, 17c.

TYPES.—Holotype, USNM 17698 (<$ I); paratypes, USNM, MCZ.

TYPE-LOCALITY.—Lake Chapala, State of Jalisco,

Mexico.

RANGE.—Jalisco and Michoacan, Mexico.

HABITAT.—Lentic situations.

Cambarellus diminutus Hobbs

FIGURE 12

Cambarellus diminutus Hobbs, 1945b:467, figs. 1, 2, 5, 6, 10,

12, 13, 17, 18, 20, 25, 26; 1972b:26, fig. 15b.

TYPES.—Holotype, allotype, and "morphotype,"

USNM 81554 (<j I, 9, <$ II); paratypes, USNM,

MCZ, CM, TU.
8 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

TYPE-LOCALITY.—Sand-bottomed stream, 3.5

miles south of Irvington, Mobile County, Alabama.

RANGE.—Mobile County, Alabama, and George

and Jackson counties, Mississippi.

HABITAT.—Pools, sluggish portions of streams

and ditches. Burrows when water disappears from

habitat.

Cambarellus montezumae (Saussure)

FIGURE 17

Cambarus Montezumae Saussure, 1857a: 102; 1858:459, fig. 22.

Cambarus Montezumae var. tridens von Martens, 1872:130.

[Syntypes, ZBM. Type-locality, Puebla, Mexico.]

Cambarus Montezumae Montezumae.—von Martens, 1872:130

[by implication].

Cambarus montezumae.—Faxon, 1890:633.

Cambarus montezumae montezumae.—Faxon, 1898:660 [by

implication].

Cambarus montezumae dugesii Faxon, 1898:660, pi. 66: fig. 1.

[Syntypes, USNM 16087 (4 311, 12$), MCZ. Type-locality,

State of Guanajuato, Mexico.]

Cambarus {Cambarellus) montezumae Ortmann, 1905c: 106.

Cambarus (Cambarellus) montezumae dugesi.—Ortmann,

1906a:20.

Cambarellus montezumae.—Hobbs, 1942a:340; 1972b:28, figs.

16f, 18b,d.

Cambarellus montezumae montezumae.—Hobbs, 1942a:350 [by

implication] Villalobos, 1943:588, pi. 1; 1955:256.

Cambarellus montezumae dugesii.—Hobbs, 1942a:350 [by

implication].

Cambarellus montezumae forma lermensis Villalobos, 1943:

603, pi. 2: figs. 3, 5, 8, 11, 14, 18, 20. [Holotype, allotype,

and "morphotype," IBM 50043 (009) ($1, $ , ^II); paratypes, IBM. Type-locality, Lerma, Estado de Mexico, Mexico.]

Cambarellus Montezumae.—Villalobos, 1943:587 [error in

title].

Cambarellus montezumae lermensis.—Villalobos, 1953:345,

359; 1955:262, pi. 59: figs. 3, 5, 8, 11, 14, 18, 20.

Cambarellus montezumae var. tridens Villalobos, 1953:345.

Cambarellus montezumae dugesi.—Villalobos, 1953:345, 358.

TYPES.—Syntypes, USNM 20583 ( $ I), GM (uncataloged, dry, 13 specimens), BMNH 3587.

TYPE-LOCALITY.—Swamps of the Valley of

Mexico; ponds in Chapultepec Park, Mexico.

RANGE.—Pacific slope and Valley of Mexico.

HABITAT.—Sluggish streams and lentic situations.

Cambarellus ninae Hobbs

FIGURE 11

Cambarellus ninae Hobbs, 1950a:89, figs. 1-17; 1972b:26, fig.

15d.

TYPES.—Holotype, allotype, and morphotype,

USNM 89768, 89770, 89769 ($ I, $, 3 II); paratypes, MCZ, ANSP, USNM, TU.

TYPE-LOCALITY.—Borrow ditches in the Aransas

Refuge (along East Shore Road), Aransas County,

Texas.

RANGE.—From Aransas to Matagorda and Wharton counties, Texas.

HABITAT.—Lentic habitats.

Cambarellus occidentalis (Faxon)

FIGURE 13

Cambarus montezumae occidentalis Faxon, 1898:661, pi. 66:

figs. 3, 4.

Cambarus (Cambarellus) montezumae occidentatis.—Ortmann, 1905c: 106 [by implication].

Cambarellus montezumae occidentalis Hobbs, 1942a:350 [by

implication].—Villalobos, 1953:358; 1955:276.

Cambarellus occidentalis Hobbs, 1972b:27, figs. 16a, 17a.

TYPES.—Syntypes, MCZ 3652 (2 a I, 2 3 II, 9

dry).

TYPE-LOCALITY.—Mazatlan, State of Sinaloa,

Mexico.

RANGE.—Michoacan and Sinaloa, Mexico.

HABITAT.—Lentic habitats.

Cambarellus patzcuarensis Villalobos

FIGURE 18

Cambarellus montezumae patzcuarensis Villalobos, 1943:607,

pi. 2: figs. 1, 6, 9, 10, 12, 15, 19, 21, 23; 1953:345, 359;

1955:265, pi. 59: figs. 1, 6, 9, 10, 12, 15, 19, 21, 23.

Cambarellus patzcuarensis.—Villalobos, 1955:280.—Hobbs,

1972b:28, figs. 16d, 17d.

TYPES.—Holotype, allotype, and "morphotype,"

IBM 90040(005) ($ I, $, all); paratypes, IBM.

TYPE-LOCALITY.—Lago de Patzcuaro, Michoadin,

Mexico.

RANGE.—Known only from the type-locality.

HABITAT.—Lake.

Cambarellus puer Hobbs

FIGURE 10

ICambarus schmittii Penn, 1941:8 [nomen nudum].

Cambarellus puer Hobbs, 1945b:469, figs. 7-9, 11, 14-16, 19,

21-24; 1972b:27, figs. 5c, 15e.—Penn, 195Ob:423.
NUMBER 166

TYPES.—Holotype, allotype, and "morphotype,"

USNM 81556 ($1, 9, $11); paratypes, USNM,

MCZ, ANSP, TU, USNM.

TYPE-LOCALITY.—Roadside ditch, 7 miles west

of Dayton, Liberty County, Texas, on U.S. Highway 90.

RANGE.—From Brazos and Matagorda counties,

Texas, to Green and Howard counties, Arkansas,

and eastward to Ascension Parish, Louisiana,

Holmes County, Mississippi, and western Tennessee.

HABITAT.—Swamps, ditches, sloughs, ponds, and

sluggish streams.

CambareUus schmitti Hobbs

FIGURE 9

CambareUus schmitti Hobbs, 1942b: 149, figs. 176-180, 286-

295; 1972b:26, fig. 15c.

TYPES.—Holotype, allotype, and "morphotype,"

USNM 81291 ($1, $, $11); paratypes, USNM,

MCZ, Chas. M., TU, UF (now in USNM).

TYPE-LOCALITY.—Small spring flowing into the

Suwannee River at Brandford, Suwannee County,

Florida.

RANGE.—From Mobile County, Alabama, eastward to the Suwannee River drainage system in

Florida.

HABITAT.—Clear streams and springs.

CambareUus shufeldtii (Faxon)

FIGURE 8

Cambarus Shufeldtii Faxon, 1884:134; 1885a:124, pi. 7: fig. 1;

pi. 10: figs. 8, 8', 8a, 8a'.

Cambarus shufeldtii.—Underwood, 1886:372.

Cambarus (CambareUus) shufeldti.—Ortmann, 1905c: 106.

CambareUus shufeldtii.—Hobbs, 1942a:350 [by implication];

1972b:25, fig. 15a—Penn, 1942:644; 1959:14, figs. 24, 45, 63,

77, 86^—Hobbs and Marchand, 1943:17, figs. 1, 11, 15, 21.

CambareUus shufeldti.—Pcnnak, 1953:462.

Cambarella shufeldti.—Carlisle and Knowles, 1959:50 [erroneous spelling].

TYPES.—Syntypes, USNM 4860 (3 $ I, 7 9 ), MCZ

3684 ($1, $11, 2? ).

TYPE-LOCALITY.—Near New Orleans, Louisiana.

RANGE.—Mississippi River drainage system in

southern Illinois, Tennessee, Arkansas, and in

Louisiana east of the Mississippi and Atchafalaya

rivers; Red River system in Louisiana and Texas;

and Pascagoula and Pearl river systems in Mississippi. Introduced into Rapides, St. Bernard, and

St. Tammany parishes, Louisiana (Penn, 1950b:

422), and into Harris County, Georgia (Penn, personal communication).

HABITAT.—Swamps, ditches, sloughs, lakes,

ponds, and sluggish streams. Burrows when water

disappears from habitat.

CambareUus zempoalensis Villalobos

FIGURE 19

CambareUus montezumae forma zempoalensis Villalobos,

1943:601, pi. 2: figs. 2, 4, 7, 13. 16, 17, 22.

CambareUus montezumae zempoalensis.—Villalobos, 1953:345,

359; 1955:257, pi. 59: figs. 2, 4, 7, 13, 16, 17, 22.

CambareUus zempoalensis Hobbs, 1972b:28, figs. 16e, 18a,c.

TYPES.—Holotype, allotype, and "morphotype,"

IBM 100042 (00FA) ($ I, $, $11); paratypes,

IBM.

TYPE-LOCALITY.—Lagunas de Zempoala, Morelos, Mexico.

RANGE.—Known only from the type-locality.

HABITAT.—Crater lakes.

Subfamily CAMBARINAE Hobbs, 1942

Genus Barbicambarus Hobbs

Barbicambarus Hobbs, 1969b:98. [Type-species by original

designation, Cambarus cornutus Faxon, 1884:120. Gender:

masculine.]

Barbicambarus cornutus (Faxon)

FIGURE 20

Cambarus cornutus Faxon, 1884:120; 1885a:80, pi. 5: figs. 1, 2;

pi. 9: figs. 3, 3'.—Cooper and Cooper, 1968:19-20, illustrated.

Cambarus (Bartonius) cornutus Ortmann, 1905c: 120.

Cambarus (Cambarus) cornutus.—Fowler, 1912:341 [by implication].

2d, 5, 13k, 14j, 17b.

Cambarus (Barbicambarus) cornutus Hobbs, 1969b:98, figs.

Barbicambarus cornutus.—Bouchard, 1972:56.

TYPES.—MCZ 3566 ($1).

TYPE-LOCALITY.—Green River, near Mammoth

Cave, Edmonson County, Kentucky.
10 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

RANGE.—Barren and upper Green river systems

in Kentucky and Tennessee.

HABITAT.—Streams.

Genus Cambarm Erichson

Astacus.—Fabricius, 1798:407.

Cambarus Erichson, 1846:97. [Type-species by subsequent designation (Faxon, 1898:644), Asiacus Bartonii Fabricius,

1798:407. Proposed as subgenus of Astacus; elevated to

generic rank by Girard, 1852:88. Gender: masculine.]

Camburus Faxon, 1885c:358 [erroneous spelling].

Camparus Williamson, 1899:47 [erroneous spelling].

S[ambarus] Steele, 1902:11 [erroneous spelling].

Bartonius Ortmann, 1905c:97. [Type-species by original designation, "C. bartoni" Fabricius, 1798:407. Proposed as subgenus of Cambarus, treated unintentionally as generic name

by Williamson, 1907:749, and declared a synonym of Cambarus by Fowler, 1912:341.]

Canbarus Thompson, 1967:47 [erroneous spelling].

cambarus Padgett, 1970:19 [lapsus calami].

Cambaras Bouchard, 1973:106 [erroneous spelling].

Subgenus Aviticamharus Hobbs

Aviticambarus Hobbs, 1969b:99. [Type-species by original

designation, Orconectes hamulatus Cope, 1881:881. Gender:

masculine.]

Cambarus (Aviticambarus) hamulatus (Cope)

FIGURE 21

Orconectes hamulatus Cope, 1881:881, pi. 7: fig. la.b.

Cambarus hamulatus.—Faxon, 1884:145.—Hobbs and Barr,

1960:17, figs. 1-10.

Cambarus (Bartonius) hamulatus.—Ortmann, 1905c: 120.

Cambarus (Cambarus) hamulatus Fowler, 1912:341 [by

implication].—Ortmann, 1931:96.

Cambarus hamueatus.—Fleming, 1938:303 [erroneous spelling]-

Cambarus (Aviticambarus) hamulatus.—Hobbs, 1969b:99,

figs. 5, 13f, 14f, 17j.

TYPES.—Syntypes, MCZ 3678 ( £ II, 9 ).

TYPE-LOCALITY.—Nickajack Cave, Marion County, Tennessee.

RANGE.—From the upper Sequatchie Valley,

Tennessee, southward to Blount County, Alabama.

HABITAT.—Subterranean streams.

Cambarus (Aviticambarus) jonesi Hobbs and Barr

FIGURE 22

Cambarus jonesi Hobbs and Barr, 1960:19, figs. 11-20.

Cambarus (Aviticambarus) jonesi.—Hobbs, 1969b:99, figs. 5,

17k.

TYPES.—Holotype, allotype, and morphotype,

USNM 104407, 104408, 104799 ($ I, $, all);

paratypes, TU, USNM.

TYPE-LOCALITY.—Cave Spring Cave, 12.1 miles

northwest of Valhermosa, Morgan County, Alabama.

RANGE.—Tennessee River basin between Florence and environs of Guntersville, Alabama.

HABITAT.—Subterranean waters.

Subgenus Cambarus Erichson

Cambarus Erichson, 1846:88. [Type-species by subsequent designation (Faxon, 1898:644), Astacus Bartonii Fabricius,

1798:407. Gender: masculine.]

Cambarus (Cambarus) bartonii bartonii

(Fabricius)

FIGURE 23

Astacus Bartonii Fabricius, 1798:407.

Astacus ciliaris Rafinesque, 1817:42. [Types not extant. Typelocality, brooks near Fishkill, Dutchess County, and Newburgh, Orange County, New York.]

Astacus pusillus Rafinesque, 1817:42. [Types not extant.

Type-locality, brooks in New York, near "Saratoga, Lake

George, Lake Champlain, Utica, Oswego, &c."]

Astacus bartonii De Kay, 1844:22.

Astacus Bartoni Tellkampf, 1845:85.

Astacus (Cambarus) Bartonii.—Erichson, 1846:97.

Cambarus Bartonii Girard, 1852:88.

Cambarus montanus Girard, 1852:88. [Types destroyed in

the Chicago fire in 1871; para type (?) ANSP 322 (3II),

from James River, Virginia. Type-locality, restricted by

Ortmann, 1931:114, "Tributary of James River, Rockbridge County, Virginia."]

Cambarus Bartoni.—Hagen, 1870:30.

Cambarus bartonii Packard, 1880:222.

Cambarus Bartonii Bartonii.—Faxon, 1885a:61 [by implication].

Cambarus bartoni Shufeldt, 1898:227.

Cambarus bartonii bartonii.—Hay, 1899b:959 [by implication].

Camparus bartonii.—Williamson, 1899:47 [erroneous spelling].

Cambarus bartoni bartoni.—Hay, 1902b:435.—Crocker, 1957:

42, pi. 1: figs. 2, 3, 6; pi. 2: figs. 5-7.

Cambarus (Bartonius) bartoni Ortmann, 1905c: 117, 120

[in part].

Cambarus bartoni typicus Ortmann, 1906b:450.

Asiacus bartoni.—Ortmann, 1906b:348.

Cambarus bartonius bartoni.—Chidester, 1908:710.
NUMBER 166 11

Cambarus (Cambarus) bartonii.—Fowler, 1912:341.

Cambarus bartonii montanus.—Faxon, 1914:385.

Cambarus montanus montanus Ortmann, 1931:106

Cambarus (Cambarus) bartoni bartoni Ortmann, 1931:130.

Cambarus bartoni montanus.—Ortmann, 1931:114.

Cambarus (Cambarus) bartonii bartonii.—Fleming, 1938:

303.—Hobbs, 1969b: 109, figs. 5, 13, 14, 19/.

Cambarus bartonius.—Roberts, 1944:370 [erroneous spelling].

Canbarus bartoni.—Thompson, 1967:47 [erroneous spelling].

Cambarus bartonni Coleman, 1972:21 [erroneous spelling].

Cambaras bartoni Bouchard, 1973:106 [erroneous spelling].

TYPES.—Type, UZM (all).

TYPE-LOCALITY.—North America; "probably

neighborhood of Philadelphia, Pa." (Faxon,

1914:423).

RANGE.—New Brunswick, Canada, to northern

Georgia, Ohio, and Tennessee to the Atlantic

Ocean; restricted to the mountains and foothills

in South Carolina and Georgia.

HABITAT.—Mostly lotic habitats, but occasionally

frequenting ponds and often burrowing along

streams and seepage areas.

Cambarus (Cambarus) bartonii carinirostris Hay

FICURE 24

Cambarus bartonii carinirostris Hay, 1914:384.

Cambarus (Cambarus) bartoni carinirostris Ortmann, 1931:

107.

Cambarus (Cambarus) bartonii carinirostris.—Hobbs, 1969b:

109, fig. 19m.

TYPES.—"Type" and paratypes, USNM 23962

(Sh 7$U, 15$); paratypes, MCZ 7399 ($ 1,

ill , $ ).

TYPE-LOCALITY.—Gandy Creek at Osceola, Randolph County, West Virginia.

RANGE.—Tributaries of the Cheat, Greenbrier,

and Tygart rivers in West Virginia. Exact range

not known.

HABITAT.—Streams of small to moderate size.

REMARKS.—Both Ortmann (1931) and Hobbs

(1972b) have remarked on the advisability of recognizing this species, and the latter indicated that

"it is questionable that this subspecies should be

recognized."

Cambarus (Cambarus) bartonii cavatus Hay

FIGURE 25

Cambarus bartoni cavatus Hay, 1902b:435.

Cambarus (Bartonius) bartoni.—Ortmann, 1905c: 120 [in

part].

Cambarus (Cambarus) bartonii cavatus.—Fowler, 1912:341

[by implication].—Hobbs, 1969b: 109, figs. 5, 19n.

Cambarus bartonii cavatus.—Faxon, 1914:425.

Cambarus (Cambarus) bartoni cavatus.—Ortmann, 1931:104.

Cambarus (Cambarus) species L Hobbs, 1969b: 109, fig. 5.

Cambarus species K.—Hobbs, 1969b:145 [not p. 109 and

fig. 5].

TYPES.—Syntypes, USNM 25017 (all, $, 2$

juv.).

TYPE-LOCALITY.—Powell River, Tazewell, Claiborne County, Tennessee.

RANGE.—Tributaries of the Tennessee River in

eastern Tennessee, northwestern Georgia, and

southwestern Virginia.

HABITAT.—Streams.

Cambarus (Cambarus) howardi Hobbs and Hall

FICURE 26

Cambarus (Cambarus) species K Hobbs, 1969b: 109, fig. 5.

Cambarus (Cambarus) howardi Hobbs and Hall, 1969:281,

figs. 1-12.

TYPES.—Holotype, allotype, and morphotype,

USNM 129866, 129867, 129868 (a I, 9, all);

paratypes, USNM.

TYPE-LOCALITY.—Sope Creek, tributary to the

Chattahoochee River at Paper Mill Road, 1.5 miles

above mouth, Cobb County, Georgia.

RANGE.—Chattahoochee drainage system from

Hall to Douglas counties, Georgia.

HABITAT.—Riffle areas of streams.

Cambarus (Cambarus) ortmanni Williamson

FIGURE 28

Bartonius ortmanni Williamson, 1907:749 [Bartonius intended

as a subgeneric name].

Cambarus (Bartonius) ortmanni Williamson, 1907:754, pi. 35:

figs. 1-3.

Cambarus (Cambarus) ortmanni.—Fowler, 1912:341 [by implication] Hobbs, 1969b: 109, figs. 5, 19o.

Cambarus ortmanni.—Creaser, 1932:336.

TYPES.—Syntypes, CM 74.826 ($ I, $), MCZ

7587 (9 ).

TYPE-LOCALITY.—Six-Mile Creek and Craven

Ditch, tributary to Wabash River, above Bluffton,

Wells County, Indiana.
12 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

RANGE.—Southern Indiana and Ohio, and adjacent northern Kentucky.

HABITAT.—Burrows (secondary burrower).

Cambarus (Cambarus) sciotensis Rhoades

FIGURE 27

Cambarus bartoni sciotensis Rhoades, 1944b:96.

Cambarus sciotensis.—Holt, 1954:170 Hobbs, Holt, and

Walton, 1967:27, fig. 5.

Cambarus bartonii sciotensis Hobbs, 1955b:330.

Cambarus (Cambarus) sciotensis Hobbs, 1969b: 109, figs. 5,

19p.

TYPES.—Holotype, allotype, and "morphotype,"

USNM 81533 ($1, 9, $ II); paratypes, MCZ, RR,

OSM.

TYPE-LOCALITY. — Limestone Cliffs below

O'Shaughnessy Dam, Scioto River, Sec. 3, Liberty

Township, Delaware County, Ohio.

RANGE.—Scioto River drainage, Ohio, and Kanawha drainage in southwestern Virginia and West

Virginia.

HABITAT.—Riffle areas in small to large streams.

Subgenus Depressicambarus Hobbs

Depressicambarus Hobbs, 1969b: 102. [Type-species by original

designation, Astacus latimanus LeConte, 1856:402. Gender:

masculine.]

Cambarus (Depressicambarus) catagius

Hobbs and Perkins

FIGURE 36

Cambarus catagius Hobbs and Perkins, 1967:141, figs. 1-11.

Cambarus (Depressicambarus) catagius Hobbs, 1969b: 104,

figs. 8, 18c

TYPES.—Holotype, allotype, and morphotype,

USNM 117779, 117780, 117781 ($1, $, $ II);

paratypes, USNM, MCZ.

TYPE-LOCALITY.—Lawn at East Whittington

Street in southeastern section of Greensboro, Guilford County, North Carolina.

RANGE.—Known only from the type-locality.

HABITAT.—Burrows (primary burrower).

Cambarus (Depressicambarus) cymatilis Hobbs

FIGURE 37

Cambarus (Depressicambarus) cymatilis Hobbs, 197Ob:251,

figs. 3, 4.

TYPES.—Holotype, allotype, and morphotype,

USNM 129860, 129861, 129862 ($1, 9, $U);

paratypes, USNM.

TYPE-LOCALITY.—Western city limits of Chatsworth, Murray County, Georgia, in lawn and rose

garden of Mr. Charles S. Dunn, off Chestnut

Street.

RANGE.—Known from only three localities in the

immediate vicinity of Chatsworth, Murray County,

Georgia.

HABITAT.—Burrows (primary burrower).

Cambarus (Depressicambarus) englishi

Hobbs and Hall

FIGURE 30

Cambarus (Depressicambarus) englishi Hobbs and Hall, 1972:

151, fig. la-n.

TYPES.—Holotype, allotype, and morphotype,

USNM 131700, 131701, 132519 ($1, 9, all);

paratypes, USNM.

TYPE-LOCALITY.—Tallapoosa River, in riffle area

1 mile north of Tallapoosa, Haralson County,

Georgia, a few hundred yards east of bridge on

State Route 100.

RANGE.—Known only from the Tallapoosa River

in Haralson County, Georgia, and Clay County,

Alabama.

HABITAT.—Stream.

Cambarus (Depressicambarus) ftoridanus Hobbs

FIGURE 34

Cambarus (Cambarus) floridanus Hobbs, 1941b: 114, figs. 1, 4,

5, 8, 9, 16, 19, 22, 25, 31, 32.

Cambarus floridanus.—Hobbs, 1942a:356; 1942b:161, pi. 11:

figs. 191-195.

Cambarus (Depressicambarus) floridanus.—Hobbs, 1969b: 104,

figs. 8, 18d.

TYPES.—Holotype and allotype, USNM 79341

(<$I, 9), and "morphotype," USNM 79344

( $ II); paratypes, MCZ, USNM.

TYPE-LOCALITY.—12 miles west of Tallahassee

on State Route 19, Leon County, Florida.

RANGE.—Gadsden, Jackson, and Leon counties,

Florida.

HABITAT.—Burrows (primary burrower).
NUMBER 166 13

Cambarus (Depressicambarus) halli Hobbs

FIGURE 29

Cambarus halli Hobbs, 1968a:269, figs. 12-22.

Cambarus (Depressicambarus) halli Hobbs, 1969b: 104, figs.

8, 18e.

TYPES.—Holotype, allotype, and morphotype,

USNM 129288, 129289, 129290 ($ I, 9, $11);

paratypes, USNM.

TYPE-LOCALITY.—Tributary of the Tallapoosa

River, 1.3 miles south of the river on U.S. Highway 27, Haralson County, Georgia.

RANGE.—Tallapoosa drainage system in Alabama

and Georgia.

HABITAT.—Streams.

Cambarus (Depressicambarus) jordani Faxon

FIGURE 32

Cambarus Jordani Faxon, 1884:119; 1885a:83, pi. 3: fig. 3.

Cambarus jordani.—Hay, 1899b:959.

Cambarus (Bartonius) jordani.—Ortmann, 1905c: 120.

Cambarus (Cambarus) jordani.—Fowler, 1912:341 [by implication].

Cambarus (Depressicambarus) jordani.—Hobbs, 1969b: 104

[by implication]; 1972b: 114, figs. 97d, 99d.

TYPES.—"Holotype," MCZ 3561

TYPE-LOCALITY.—Etowah River, near Rome,

Floyd County, Georgia.

RANGE.—Coosa River system in Alabama and

Georgia.

HABITAT.—Streams.

Cambarus {Depressicambarus) latimanus

(LeConte)

FIGURE 33

Astacus latimanus LeConte, 1856:402.

Cambarus latimanus.—Hagen, 1870:83, pi. 1: figs. 43-46; pi.

3: fig. 162.—Faxon, 1885a:69, pi. 2: fig. 3.—Hobbs, 1942b:

158, figs. 186-190.

Cambarus obesus var. latimanus.—Packard, 1880:222.

Cambarus (Bartonius) latimanus.—Ortmann, 1905c: 120.

Cambarus (Cambarus) latimanus.—Fowler, 1912:341 [by implication].

Cambarus (Depressicambarus) latimanus.—Hobbs, 1969b: 104,

figs. If, 8, 13g, 14g, 18f.

TYPES.—Syntypes, MCZ 3378 ($ I dry), MCZ,

ANSP 329 ($).

TYPE-LOCALITY.—Athens, Clarke County, Georgia.

RANGE.—North Carolina to Alabama, chiefly in

the piedmont but extending into western Florida

along the Apalachicola River.

HABITAT.—Small streams and burrows (secondary burrower).

Cambarus (Depressicambarus) obstipus Hall

FIGURE 31

Cambarus obstipus Hall, 1959:221, figs. 1-9.

Cambarus (Depressicambarus) obstipus.—Hobbs, 1969b: 104,

figs. 8, 18g.

TYPES.—Holotype and allotype, USNM 102123,

102124 ($1, $); paratypes, USNM.

TYPE-LOCALITY.—Black Warrior River at Underwood's Ferry, 8 miles downstream from Cordova,

Walker County, Alabama.

RANGE.—Known only from the Black Warrior

River system in Alabama.

HABITAT.—Streams.

Cambarus (Depressicambarus) reduncus Hobbs

FIGURE 39

Cambarus reduncus Hobbs, 1956b:61, figs. 1-11.

Cambarus (Depressicambarus) reduncus.—Hobbs, 1969b: 104,

figs. 8, 18h.

TYPES.—Holotype, allotype, and morphotype,

USNM 99183, 99184, 99185 (a I, 9, $11); paratypes, TU, USNM.

TYPE-LOCALITY.—Flood plain pools of Little

River, 10.3 miles west of Winnsboro, Fairfield

County, South Carolina.

RANGE.—Piedmont province from Orange County, North Carolina, to Richland County, South

Carolina.

HABITAT Streams, ponds, and burrows (primary or secondary burrower).

Cambarus (Depressicambarus) sphenoides Hobbs

FIGURE 35

Cambarus sphenoides Hobbs, 1968a:262, figs. 1-11.

Cambarus (Depressicambarus) sphenoides Hobbs, 1969b:

104, figs. 8, 18i.
14 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

TYPES.—Holotype, allotype, and morphotype,

USNM 129325, 129326, 129327 ($ I, 9, all);

para types, USNM.

TYPE-LOCALITY.—Tributary to Clear Creek

(Emory River drainage), 11.2 miles north of Crossville, Cumberland County, Tennessee, on U.S.

Highway 127.

RANGE.—Stream tributaries of the Cumberland,

Emory, Sequatchie, and Tennessee river systems

on the Cumberland Plateau in Kentucky and

Tennessee.

HABITAT.—Streams.

Cambarus (Depressicambarus) striatus Hay

FIGURE 38

Camburus sp. (Ashland City, Tennessee) Faxon, 1885c:358

[lapsus for Cambarus].

Cambarus latimanus striatus Hay, 1902b:437.

Cambarus graysoni Faxon, 1914:393. [Types, MCZ 3593 (3II,

3 9) . Type-locality, Bear Creek, a tributary of Green

River, Grayson Springs, Grayson County, Kentucky.]

Cambarus (Cambarus) bartoni striatus.—Ortmann, 1931:140.

Cambarus (Cambarus) bartonii striatus.—Fleming, 1938:303.

Cambarus latimanus.—Fleming, 1939:311 [erroneous spelling].

Cambarus latimus.—Fleming, 1939:319 [erroneous spelling].

Cambarus bartonii striatus.—Hobbs, 1942a:354 [by implication].

Cambarus bartoni striatus.—Rhoades, 1944a: 114.

Cambarus striatus.—Hobbs, 1956b:61.

Cambarus (Depressicambarus) striatus.—Hobbs, 1969b: 104,

figs. 8, 18j.

TYPES.—Syntypes, USNM 25019 ($1, 3 5II,

4 9 ), MCZ 7348 ($ I, a II, 3 9).

TYPE-LOCALITY.—Nashville, Davidson County,

Tennessee.

RANGE.—Southern Kentucky and central Tennessee.

HABITAT.—Small streams, springs, and burrows

(secondary burrower).

Subgenus Erebicambarus Hobbs

Erebicambarus Hobbs, 1969b:99. [Type-species by original

designation, Cambarus bartoni tenebrosus Hay, 1902a:232.

Gender: masculine.]

Cambarus (Erebicambarus) cahni Rhoades

FIGURE 44

Cambarus (Cambarus) cahni Rhoades, 1941b: 146, fig. 36A-D.

Cambarus cahni.—Hobbs, 1942a:354 Hobbs and Barr, 1960:

25, figs. 31-37.

Cambarus (Erebicambarus) cahni.—Hobbs, 1969b: 100, figs.

6, 17c.

TYPES.—Holotype and allotype, USNM 80031,

80032 ($ I, 9 ); paratypes, MCZ, Alabama Mus.

Nat. Hist., ANSP, Leslie Hubricht, RR.

TYPE-LOCALITY.—Belgreen Cave, NW i/4 SW 14

sec. 12, T. 7 S., R. 13 W., Franklin County, Alabama.

RANGE.—Known only from the type-locality.

HABITAT.—Subterranean streams.

REMARKS.—It is probable that this species is

either identical to C. (£.) tenebrosus Hay or should

be accorded subspecific rank.

Cambarus (Erebicambarus) hubbsi Creaser

FlCURE 4 0

Cambarus (Bartonius) hubbsi Creaser, 1931a:4, figs. 7-12.

Cambarus hubbsi.—Hobbs, 1942a:356.—Williams, 1954:904,

figs. 233, 235-245.

Cambarus (Erebicambarus) hubbsi.—Hobbs, 1969b: 100, fig.

17d.

TYPES.—Holotype and allotype, USNM 98332,

98333 (si, 9); paratypes, USNM.

TYPE-LOCALITY.—Little Creek, tributary to St.

Francis River, 1 mile northeast of Chloride, Iron

County, Missouri.

RANGE.—Southeastern Missouri and northeastern

Arkansas.

HABITAT.—Rocky streams.

Cambarus (Erebicambarus) hubrichti Hobbs

FIGURE 42

Cambarus hubrichti Hobbs, 1952c:689, figs. 1-8.—Hobbs and

Barr, 1960:23, figs. 21-30.

Cambarus hubricht.—Hobbs, 1952c:693 [erroneous spelling].

Cambarus Hubrichti.—Cooper and Poulson, 1968:130, fig. 8.

Cambarus (Erebicambarus) hubrichti.—Hobbs, 1969b: 100,

fig. 17e.

TYPES.—Holotype and allotype, USNM 92295,

92296 («$ II, 9); paratypes, USNM.

TYPE-LOCALITY.—Stream in Lewis Cave, 15 miles

northwest of Doniphan, Ripley County, Missouri.

RANGE.—White River basin in Carter, Ripley,

and Oregon counties, Missouri.

HABITAT.—Subterranean waters.
NUMBER 166 15

Cambarus (Erebicambarus) laevis Faxon

FIGURE 46

Cambarus bartonii laevis Faxon, 1914:391.

Cambarus (Cambarus) bartoni laevis.—Ortmann, 1931:143.

Cambarus bartoni laevis.—Fleming, 1939:311.

Cambarus (Erebicambarus) laevis.—Hobbs, 1969b: 100, figs.

6, 17f.

Cambarus laevis.—Hobbs III, 1971:140.

TYPES.—Syntypes, MCZ 3812 ($11, 9). The

male should be considered the holotype and the

female a para type (see Faxon, 1914:392).

TYPE-LOCALITY.—Bloomington, Monroe County,

Indiana.

RANGE.—Southern Illinois and Indiana, southwestern Ohio, and northern Kentucky.

HABITAT.—Streams, springs, and caves.

Cambarus (Erebicambarus) ornatus Rhoades

FIGURE 45

Cambarus bartoni ornatus Rhoades, 1944a: 144, fig. 10A-F.

Cambarus bartonii ornatus.—Hobbs and Shoup, 1947:145.

Cambarus ornatus.—Hobbs and Barr, 1960:15.

Cambarus (Erebicambarus) ornatus.—Hobbs, 1969b: 100, figs.

6, 17g.

TYPES.—Holotype, allotype, and "morphotype,"

USNM 81330, 81331, 81332 ($ I, 9, $11); paratypes, MCZ, USNM, RR.

TYPE-LOCALITY.—Ellis Branch, 4 miles west of

Carrollton, Carroll County, Kentucky.

RANGE.—Ohio Basin from the lower Kentucky

River system to the Salt River, Kentucky.

HABITAT.—Intermittent streams.

Cambarus (Erebicambarus) rusticiformis Rhoades

FIGURE 41

Cambarus rusticiformis Rhoades, 1944a: 133, fig. 8A-F.

Cambarus (Erebicambarus) rusticiformis.—Hobbs, 1969b: 100,

figs. 2a, 6, 13h, 17h.

TYPES.—Holotype, allotype, and "morphotype,"

USNM 81324, 81325, 81326 ($ I, 9, 3 II); paratypes, MCZ, RR.

TYPE-LOCALITY.—Little River, 0.5 mile west of

Cadiz, Trigg County, Kentucky.

RANGE.—Cumberland drainage system in Kentucky and Tennessee.

HABITAT.—Swift streams.

Cambarus (Erebicambarus) tenebrosus Hay

FIGURE 43

Cambarus bartoni tenebrosus Hay, 19O2a:232.

Cambarus (Barionius) bartoni.—Ortmann, 1905c: 120 [in

part].

Cambarus (Cambarus) bartoni tenebrosus.—Fowler, 1912:341

[by implication].

Cambarus bartonii tenebrosus.—Faxon, 1914:424.

Cambarus tenebrosus.—Garman, 1920:42.—Hobbs, 1959:898.

Cambarus Bartoni tenebrosus.—Chappuis, 1927:92.

Cambarus (Bartonius) Bartoni tenebrosus.—Fage, 1931:373.

Cambarus (Cambarus) bartoni tenebrosus.—Ortmann, 1931:

145.

Cambarus (Bartonius) bartoni tenebrosus.—Hart, 1962:121.

Cambarus (Erebicambarus) tenebrosus.—Hobbs, 1969b: 100,

figs. 6, 14h, 17i.

TYPES.—Syntypes, USNM 22346 ($ II, 2 9).

TYPE-LOCALITY.—Echo River, Mammoth Cave,

Edmonson County, Kentucky.

RANGE.—Mammoth Cave southward to northern

Alabama.

HABITAT.—Streams and subterranean waters

(troglophilic).

Subgenus Hiaticambarus Hobbs

Hiaticambarus Hobbs, 1969b: 105. [Type-species by original

designation, Cambarus longulus Girard, 1852:90. Gender:

masculine.]

Cambarus (Hiaticambarus) chasmodactylus James

FlCURE 50

Cambarus longulus chasmodactylus James, 1966:14, fig. 1,

pi. lb,d,e,q,u-z,zz.

Cambarus chasmodactylus.—Hobbs, 1968b:K-17.

Cambarus (Hiaticambarus) chasmodactylus.—Hobbs, 1969b:

106, figs. 10, 18k.

TYPES.—Holotype, allotype, and morphotype,

USNM 115513, 115514, 115516 (<$I, 9, 3II);

paratypes, MCZ, TU, IBM, USNM, Hugo L.

James.

TYPE-LOCALITY.—East Fork of the Greenbrier

River, 9.7 miles west of Virginia State line on U.S.

Highway 250, Pocahontas County, West Virginia.

RANGE.—New River drainage system in northwestern North Carolina, Virginia, and eastern West

Virginia.

HABITAT.—Rocky streams.
16 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Cambarus (Hiaticambarus) girardianus Faxon

FIGURE 47

Cambarus Girardianus Faxon, 1884:117; 1885a:78, pi. 4: fig. 1;

pi. 9: figs. 2a, 2a'.

Cambarus girardianus.—Faxon, 1898:650.

Cambarus extraneus girardianus—Faxon, 1898:650.

Cambarus (Bartonius) extraneus girardianus—Ortmann,

1905c: 120 [by implication].

Cambarus (Cambarus) extraneus girardianus.—Fowler, 1912:

341 [by implication].

Cambarus extraneus Girardianus.—Fleming, 1938:299.

Cambarus (Hiaticambarus) girardianus.—Hobbs, 1969b: 106,

figs. 10, 18/.

TYPES Syntypes, MCZ 3560 ( $ II, 9 ), USNM

4882 (all, 2 9 )•

TYPE-LOCALITY.—Cypress Creek, Lauderdale

County, Alabama.

RANGE.—Tennessee River drainage system in

Alabama, Georgia, Mississippi, and Tennessee.

HABITAT.—Streams.

Cambarus (Hiaticambarus) longirostris Faxon

FIGURE 48

Cambarus Bartonii var. longirostris Faxon, 1885a:64.

Cambarus bartonii longirostris.—Faxon, 1890:623.

Cambarus bartonii spinirostris Faxon, 1890:623 [lapsus for

C. b. longirostris].

Cambarus longulus longirostris Hay, 1899b: 959.—James,

1966:9, fig. 2a,b, pi. lc,f,g,n,o,r,s.

Cambarus (Bartonius) bartoni longirostris Ortmann, 1905c:

135 [by implication].

Cambarus (Cambarus) bartonii longirostris.—Fowler, 1912:

341 [by implication].

Cambarus (Cambarus) longulus longirostris.—Ortmann, 1931:

121.

Cambarus longerosilis.—Brimley, 1938:503 [erroneous spelling].

Cambarus (Hiaticambarus) longirostris.—Hobbs, 1969b: 106,

figs. 10, 18m.

TYPE.—Holotype, MCZ 3629 ($ II).

TYPE-LOCALITY.—"Eastern Tennessee and West

Virginia" (Faxon, 1885a). Doe River, Elizabethton, Carter County, Tennessee, designated by Ortmann, 1931.

RANGE.—Tennessee River drainage in eastern

Tennessee, southwestern Virginia, and western

North Carolina, in the Coosa drainage in northeastern Alabama and Georgia.

HABITAT.—Rocky streams.

Cambarus (Hiaticambarus) longulus Girard

FIGURE 49

Cambarus longulus Girard, 1852:90.

Cambarus Bartonii longulus.—Faxon, 1885a:66 [by implication].

Cambarus longulus longulus—Hay, 1899b:959 [by implication].'—James, 1966:4, fig. 2c, pi. la,h-m,p,t,mm.

Cambarus (Bartonius) longulus.—Ortmann, 1905c: 120.

Cambarus (Cambarus) longutus longulus.—Fowler, 1912:341

[by implication] Ortmann, 1931:106.

Cambarus bartonii longulus.—Faxon, 1914:387.

Cambarus bartoni longulus.—Ortmann, 1931:118.

[Cambarus longulus] longulous James, 1966:22 [erroneous

spelling].

Cambaroides longulus.—Unestam, 1969:204 [lapsus for

Cambarus].

Cambarus (Hiaticambarus) longulus Hobbs, 1969b: 106, figs.

la, 10, 13c, 14c, 18n.

TYPES.—Destroyed in the Chicago fire, 1871.

TYPE-LOCALITY.—"Middle States" (Girard, 1852).

South River, Waynesboro, Augusta County, Virginia, designated by Ortmann, 1931. Emended by

Hobbs (1967a: 129) to Rockfish River, south of

Afton, Nelson County, Virginia.

RANGE.—Atlantic watershed from the James

drainage in Virginia south to the Yadkin drainage

in North Carolina, in mountains and piedmont.

HABITAT.—Rocky streams.

Subgenus Jugicambarus Hobbs

Jugicambarus Hobbs, 1969b: 106. [Type-species by original

designation, Cambarus bartonii asperimanus Faxon, 1914:

391. Gender: masculine.]

Cambarus (Jugicambarus) asperimanus Faxon

FIGURE 57

Cambarus bartonii asperimanus Faxon, 1914:391.

Cambarus (Cambarus) bartoni asperimanus.—Ortmann, 1931:

137.

Cambarus asperimanus.—Brimley, 1938:503.

Cambarus (Jugicambarus) asperimanus.—Hobbs, 1969b: 107,

figs. Id, 9, 13d, 14d, 18o.

TYPES.—Syntypes, USNM 47375 (2 3 I).

TYPE-LOCALITY.—Flat Creek, Montreat, Buncombe County, North Carolina.

RANGE.—Headwaters of the French Broad, Little

Tennessee, Catawba, Broad, Savannah, and Saluda

rivers in Georgia, North Carolina, and South Carolina.

HABITAT.—Small mountain streams.
NUMBER 166

Cambarus (Jugicambarus) bouchardi Hobbs

FIGURE 51

Cambarus (Veticambarus) bouchardi Hobbs, 1970b:245, figs.

1.2.

Cambarus (Jugicambarus) bouchardi.—Bouchard, 1973:105.

TYPES.—Holotype, allotype, and morphotype,

USNM 130295, 130296, 130297 (<$I, 9, a II);

paratypes, USNM, Raymond W. Bouchard.

TYPE-LOCALITY.—Perkins Creek, 6.9 miles north

of Oneida, Scott County, Tennessee, on U.S. Highway 27.

RANGE.—Known from only three localities in

the Big South Fork of the Cumberland River in

Scott County, Tennessee.

HABITAT.—Streams.

Cambarus (Jugicambarus) brachydactylus Hobbs

FIGURE 58

Cambarus brachydactylus Hobbs, 1953c:20, pi. 1: figs. 1-10.

Cambarus (Jugicambarus) brachydactylus Hobbs, 1969b:

107, 6gs. 9, 18p.

TYPES.—Holotype and allotype, USNM 93155

(Sh $ ); morphotype, USNM 93156 (<jll): paratypes, MCZ, USNM, TU.

TYPE-LOCALITY.—Louise Creek, 13.9 miles south

of Clarksville, Montgomery County, Tennessee.

RANGE.—Tributaries of the Cumberland River

on the western Highland Rim in Montgomery

and Dickson counties, Tennessee.

HABITAT.—Riffle areas of streams.

Cambarus (Jugicambarus) carolinus (Erichson)

FIGURE 62

Astacus Carolinus Erichson, 1846:87.

Astacus (Cambarus) Carolinus Erichson, 1846:96.

Astacus (Cambarus) carolinus.—Dana, 1852:522 [by implication].

Cambarus carolinus.—Girard, 1852:88.

Cambarus Carolinus Hagen, 1870:8.

Cambarus (Bartonius) carolinus.—Ortmann, 1905c: 120.

Cambarus carolinensis Adams, 1907:899 [erroneous spelling].

Cambarus (Cambarus) carolinus—Fowler, 1912:341 [by implication].

Cambarus carolinus carolinus.—Faxon, 1914:399.

Cambarus (Jugicambarus) carolinus.—Hobbs, 1969b: 107 [in

part], figs. 9, 19a.—Hobbs and Bouchard, 1973:51, figs. 3, 4.

17

TYPES.—Not extant. Topotypes, USNM 144164

(2 ^ I, 9, 9 with young), R. W. Bouchard.

TYPE-LOCALITY.—Greenville, Greenville County,

South Carolina. Restricted by Hobbs and Bouchard (1973:55) to "Along Wildcat Creek, 8.8 miles

north of the junction of U.S. Highway 29 and State

Route 101, on latter. . . ."

RANGE.—Known with certainty only in the

upper Enoree and Saluda drainage basins in South

Carolina.

HABITAT.—Burrows (primary burrower).

Cambarus (Jugicambarus) causeyi Reimer

FIGURE 65

Cambarus causeyi Reimer, 1966:9, figs. 1-8.

Cambarus (Jugicambarus) causeyi.—Hobbs, 1969b: 107, fig.

19b.

TYPES.—Holotype, allotype, USNM 116678,

116679 (a I, 9); paratype, USNM.

TYPE-LOCALITY.—Spring and natural pond, 4

miles west of Sandgap, Pope County, Arkansas, on

State Route 124.

RANGE.—Known only from the vicinity of the

type-locality.

HABITAT.—Complex burrows on hillside and

near spring and pond.

Cambarus (Jugicambarus) conasaugaensis

Hobbs and Hobbs

FIGURE 56

Cambarus conasaugaensis Hobbs and Hobbs, 1962:41, figs.

1-10.

Cambarus (Jugicambarus) conasaugaensis.—Hobbs, 1969b:

107, figs. 9, 19c.

TYPES.—Holotype, allotype, and morphotype,

USNM 107156, 107157, 107158 (& I, 9, $ II);

paratypes, USNM.

TYPE-LOCALITY Small tributary of the Conasauga River 2 miles east of Chatsworth, Murray

County, Georgia, on U.S. Highway 76.

RANGE.—Tributaries of the Coosawattee and

Conasauga rivers in Gilmer and Murray counties,

Georgia.

HABITAT.—Streams.
18 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Catnbarus (Jugicambarus) cryptodytes Hobbs

FIGURE 64

Cambarus cryptodytes Hobbs, 1941b:110, figs. 2, 3, 7, 11, 13,

15, 18, 21, 24, 28, 29; 1942b: 162, pi. 11: figs. 196-200—

Hobbs and Barr, 1960:31, figs. 48-57.

Cambarus (Jugicambarus) cryptodytes.—Hobbs, 1969b: 107,

figs. 9, 19d.

cambarus cryptodytes.—Padgett, 1970:19 [lapsus calami].

TYPES.—Holotype, allotype, and "morphotype,"

USNM 79339, 79340, 79343 (a I, 9, ill) ; paratypes, MCZ, USNM.

TYPE-LOCALITY.—R. W. Williams farm, 2 miles

south of Graceville, Jackson County, Florida.

RANGE.—Known from several caves in Jackson

County, Florida, and Climax Cave, 3 miles north

of Climax, Decatur County, Georgia.

HABITAT.—Subterranean waters.

Catnbarus (Jugicambarus) distorts Rhoades

FIGURE 53

Cambarus distorts Rhoades, 1944a: 136, fig. 9a-f.

Cambarus (Jugicambarus) distans.—Hobbs, 1969b: 107, figs.

9, 19e.

TYPES.—Holotype, allotype, and "morphotype,"

USNM 81327, 81329, 81328 (<$ I, 9, all); paratypes, CM, USNM, RR.

TYPE-LOCALITY.—"Cumberland River and small

tributary, just above Cumberland Falls, McCreary

County, Kentucky" (Rhoades, 1944a: 139).

RANGE.—Cumberland drainage in southeastern

Kentucky and tributaries on Cumberland Plateau

in Tennessee.

HABITAT.—Streams.

Cambarus {Jugicambarus) dubius Faxon

FIGURE 60

Cambarus dubius Faxon, 1884:114; 1885a:7O, pi. 4: fig. 3; pi.

8: figs. 7, V.

Cambarus carolinus dubius.—Faxon, 1914:396, 425.

Cambarus (Jugicambarus) dubius.—Hobbs and Bouchard,

1973:62.

TYPES.—Holotype, MCZ 3631 ($1).

TYPE-LOCALITY.—Terra Aha (Cranberry Summit), Preston County, West Virginia.

RANGE.—Poorly known, but extending southward through the Allegheny Mountains from

southwestern Pennsylvania into Virginia. The

same, or a closely allied, species also occurs in

southeastern Kentucky, eastern Tennessee, and

northwestern North Carolina.

HABITAT.—Burrows (primary burrower).

Cambarus (Jugicambarus) friaufi Hobbs

FIGURE 59

Cambarus friaufi Hobbs, 1953c:24, figs. 11-18.

Cambarus (Jugicambarus) friaufi Hobbs, 1969b: 107, figs. 9,

19f.

TYPES.—Holotype, and allotype, USNM 93157

( a I, 9 ); paratypes, MCZ, USNM.

TYPE-LOCALITY.—Small stream tributary of Cumberland River at Elmwood, Smith County, Tennessee.

RANGE.—Eastern Highland Rim in Smith and

De Kalb counties, Tennessee.

HABITAT.—Swift reaches of small streams.

Cambarus (Jugicambarus) gentryi Hobbs

FICURE 63

Cambarus (Jugicambarus) gentryi Hobbs, 1970a: 163, fig. 1.

TYPES.—Holotype, allotype, and morphotype,

USNM 130283, 130284, 130285 (a I, 9, all);

paratypes, USNM.

TYPE-LOCALITY.—Boggy area below seepage

along small tributary to Turnbull Creek (HarpethCumberland drainage system), 1 mile west of

Kingston Springs, Cheatham County, Tennessee.

RANGE.—Portions of the Cumberland (Stones

River, Harpeth River, and Yellow Creek) and Duck

rivers in middle Tennessee.

HABITAT.—Burrows (primary burrower).

Cambarus (Jugicambarus) monongalensis

Ortmann

FIGURE 61

Cambarus monongalensis Ortmann, 1905a:395.

Cambarus (Bartonius) monongalensis.—Ortmann, 1905c: 120;

1906b:398, pi. B: fig. 4; pi. 39: figs. 4, 10; pi. 40: fig. 5; pi.

41: figs. 2-4.

Cambarus (Cambarus) monongalensis.—Fowler, 1912:341 [by

implication].
NUMBER 166 19

Cambarus carolinus monongalensis.—Faxon, 1914:396, 399,

425.

Cambarus carolinus monongalenses.—Hand, 1954:437 [erroneous spelling].

Cambarus (Jugicambarus) monongalensis.—Hobbs, 1969b: 109,

figs. 9, 19g.

TYPES.—Syntypes, CM 74.316, USNM 30613

($1, 2 9), MCZ 6953 (<$ I, 9), ZIAS 1/44968 (4

specimens), ANSP 1601 ($1, 3$).

TYPE-LOCALITY.—Edgewood Park, Allegheny

County, Pennsylvania.

RANGE.—Western Pennsylvania and adjacent

northern West Virginia.

HABITAT.—Burrows (primary burrower).

Cambarus (Jugicambarus) obeyensis

Hobbs and Shoup

FIGURE 52

Cambarus obeyensis Hobbs and Shoup, 1947:138, figs. 1-11.

Cambarus (Jugicambarus) obeyensis Hobbs, 1969b: 107, figs.

9, 19h—Bouchard, 1973:105.

Cambarus (Veticambarus) obeyensis.—Hobbs, 1970b:243.

TYPES.—Holotype, allotype, and "morphotype,"

USNM 82260 ($ I, $, ^ II); paratypes, MCZ,

USNM, TU.

TYPE-LOCALITY.—Big Hurricane Creek on the

Monterey-Clarkrange road, Putnam-Cumberland

county line, Tennessee.

RANGE.—Headwaters of East Fork of Obey River

in Cumberland, Fentress, Putnam, and Overton

counties, Tennessee.

HABITAT.—Rocky streams.

Cambarus (Jugicambarus) parvoculus

Hobbs and Shoup

FIGURE 55

Cambarus parvoculus Hobbs and Shoup, 1947:142, figs. 12-22.

Cambarus (Jugicambarus) parvoculus.—Hobbs, 1969b: 107,

figs. 9, 19i.

TYPES.—Holotype, allotype, and "morphotype,"

USNM 82259 (<$L 9, all); paratypes, USNM,

MCZ, TU.

TYPE-LOCALITY.—Tributary to Big Hurricane

Creek, Fentress County (southwestern part), Tennessee.

RANGE.—Cumberland River drainage in Overton, Fentress, and Putnam counties, Tennessee,

and Bell County, Kentucky, and Tennessee River

drainage in Lee County, Virginia.

HABITAT.—Rocky streams.

Cambarus (Jugicambarus) setosus Faxon

FIGURE 66

Cambarus setosus Faxon, 1889:237, pi. 1: figs. 1-3, 7; pi. 2:

fig. 1_Hobbs and Barr, 1960:26, figs. 38-47.

Cambarus ayersii Steele, 1902:18, pi. 5: fig. A; pi. 6: fig. 14.

[Types, USNM 62316 (3II). Type-locality, Fisher's Cave

(Sequiota Cave in Galloway, T. 28 N., R. 21 W., sec. 9),

near Springfield, Green County, Missouri.]

Cambarus (Bartonius) setosus.—Ortmann, 1905c: 120.

Cambarus (Cambarus) setosus.—Fowler, 1912:341 [by implication].

Cambarus (Jugicambarus) setosus.—Hobbs, 1969b: 107. fig.

19j.

TYPES.—Syntypes, MCZ 4200 (4 $ II, 9), MCZ

4201 (3<$I, $ II, 4$ , 4ajuv., 2 9juv.), MCZ

4202 (2 $ II, 3 9, 3 $ juv., 4 9juv.), USNM 25828

(6 II, 9).

TYPE-LOCALITY.—Wilson's Cave [Whisner Cave,

2 miles northwest of Sarcoxie, T. 27 N., R. 29 W.,

sec. 6], near Springfield, Jasper County, Missouri.

RANGE.—Southwestern Missouri and northeastern Oklahoma (?).

HABITAT.—Subterranean streams.

Cambarus (Jugicambarus) tartarus

Hobbs and Cooper

FIGURE 68

Cambarus (Jugicambarus) tariarus Hobbs and Cooper, 1972:

51, figs. 1-12.

TYPES.—Holotype, allotype, and morphotype,

USNM 131951, 131411, 132754 (3 I, 9, ill).

TYPE-LOCALITY.—Stansberry-January Cave System, 4 miles north of Colcord (T. 21 N., R. 22 E.,

sec. II), Delaware County, Oklahoma.

RANGE.—Known only from the type-locality.

HABITAT Subterranean stream.

Cambarus (Jugicambarus) unestami

Hobbs and Hall

FIGURE 54

Cambarus (Depressicambarus) unestami Hobbs and Hall,

1969:287, figs. 13-24.

Cambarus (Jugicambarus) unestami Bouchard, 1973:105.
20 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

TYPES.—Holotype, allotype, and morphotype,

USNM 129863, 129864, 129865 (<jl, 9, $ II);

paratypes, USNM.

TYPE-LOCALITY.—Daniel Creek, 2.5 miles west

of the Walker County line on State Route 143,

Dade County, Georgia.

RANGE.—Tributaries of the Tennessee River in

Dade County, Georgia.

HABITAT.—Swift portions of small streams.

Cambarus (Jugicambarus) zophonastes

Hobbs and Bedinger

FIGURE 67

Cambarus zophonastes Hobbs and Bedinger, 1964:11, figs. 1,

2, 4-11.

Cambarus (Jugicambarus) zophonastes.—Hobbs, 1969b: 107,

fig. 19k.

TYPES.—Holotype and allotype, USNM 108356,

108357 (&1, 9); paratypes, USNM.

TYPE-LOCALITY.—Hell Creek Cave, Stone County, Arkansas (NE14NE14, sec. 30, T. 15 N., R.

10 W.).

RANGE.—Known only from the type-locality.

HABITAT.—Subterranean stream.

Subgenus Lacunicambarus Hobbs

Lacunicambarus Hobbs, 1969b: 110. [Type-species by original

designation, Cambarus diogenes Girard, 1852:88. Gender:

masculine.]

Cambarus (Lacunicambarus) diogenes diogenes

Girard

FIGURE 69

Astacus fossor Rafinesque, 1817:42. [Name suppressed by

International Commission, Opinion 522, 1958.]

Cambarus diogenes Girard, 1852:88.

Cambarus nebrascensis Girard, 1852:91. [Location of types

(if extant) unknown. Type-locality, Fort Pierre, Nebraska

(now in Stanley County, South Dakota).]

Cambarus Diogenes.—Hagen, 1870:6.

Cambarus obesus Hagen, 1870:81, pi. 1: figs. 39-42; pi. 3: fig.

163; pi. 9. [Syntypes: MCZ 195 (231, $) , 165 ($), 3361

(£1). 229 (£I), 3363 ($); MHNP (2 dry); St. Petersburg Mus., U.S.S.R. (2), lost. Type-locality, Lawn Ridge,

Illinois.]

Cambarus Diogenenes.—Abbott, 1873:83 [erroneous spelling].

Cambarus Diogenes Diogenes.—Faxon, 1884:144 [by implication].

Cambarus diogenes diogenes Hay, 1899b:959 [by implication].—Marlow, 1960:229.

S[ambarus] diogenes Steele, 1902:11 [erroneous spelling].

Cambarus (Bartonius) diogenes Ortmann, 1905c: 120; 1906b:

402, pi. A: fig. 3; pi. 39: fig. 11; pi. 40: figs. 6, 7.

Bartonius diogenes.—Williamson, 1907:749 [Bartonius intended as a subgeneric name].

Cambarus (Cambarus) diogenes.—Fowler, 1912:341, 348.

Cambarus (Lacunicambarus) diogenes diogenes—Hobbs,

1969b: 110, figs. 2c, 11, 13i, 14i, 20a.

TYPES.—Not known to be extant except for a

questionable paratype ($ ), ANSP.

TYPE-LOCALITY.—Vicinity of Washington, D. C.

RANGE.—Very widespread east of the Rockies

and south of the Great Lakes, except peninsular

Florida and the Alleghenies; not reported northeast of New Jersey in the East and east of western

Pennsylvania in the Mississippi drainage system.

HABITAT.—Burrows (primary burrower); occasionally found in ponds or streams.

REMARKS.—This is a species complex and needs

considerable attention.

Cambarus (Lacunicambarus) diogenes

ludovicianus Faxon

FIGURE 70

Cambarus Diogenes var. Ludoviciana Faxon, 1885:144.

Cambarus diogenes ludovicianus.—Hay, 1899b:959.—Marlow,

1960:229.

Cambarus diogenes, ludoviciana.—Steele, 1902:21.

Cambarus diogenes ludoviciana.—Penn, 1941:8.

Cambarus ludovicianus.—Roberts, 1944:363.

Cambarus (Lacunicambarus) diogenes ludovicianus.—Hobbs,

1969b: 110, fig. 20b.

TYPES.—Syntypes, USNM 5504 ( $ I, 3 II, 4?),

5625 (4II, 9), MCZ 3617 ($ I).

TYPE-LOCALITY.—New Orleans, Orleans Parish,

Louisiana.

RANGE.—Lower Mississippi drainage in Louisiana.

HABITAT.—Primary burrower.

Subgenus Puncticambarus Hobbs

Puncticambarus Hobbs, 1969b: 101. [Type-species by original

designation, Cambarus extraneus Hagen, 1870:73. Gender:

masculine.]

Cambarus (Puncticambarus) acuminatus Faxon

FIGURE 74

Cambarus acuminatus Faxon, 1884:113; 1885a:67, pi. 3: fig.
NUMBER 166 21

5; pi. 8: figs. 6a, 6a' Hobbs and Walton, 1966:136.

Cambarus (Barionius) acuminatus.—Ortmann, 1905c: 120.

Cambarus (Cambarus) acuminatus.—Fowler, 1912:341 [by

implication].

Cambarus bartonii acuminatus.—Faxon, 1914:424.

Cambarus montanus acuminatus.—Ortmann, 1931:106.

Cambarus bartoni acuminatus.—Ortmann, 1931:108.

Cambarus (Cambarus) montanus acuminatus.—Ortmann,

1931:108.

Cambarus (Puncticambarus) acuminatus.—Hobbs, 1969b: 101,

figs. 7, 14a, 17/.

TYPES.—Syntype, MCZ 3624 ($). "Cotypes"

formerly said to have been at Butler University

(Faxon, 1914:424) are no longer there.

TYPE-LOCALITY.—Saluda River, at Farr's Mills,

west of Greenville, Greenville County, South Carolina.

RANGE.—Atlantic watershed from Maryland to

Georgia.

HABITAT.—Streams.

Cambarus (Puncticambarus) chaugaensis

Prins and Hobbs

FIGURE 75

Cambarus (Puncticambarus) sp. E Hobbs, 1969b: 102, 134, 135.

Cambarus (Puncticambarus) chaugaensis Prins and Hobbs,

1972:413, fig. 1.

TYPES.—Holotype, allotype, and morphotype,

USNM 131926, 131927, 131928 (41, $, 3 II);

paratypes, USNM.

TYPE-LOCALITY.—Chauga River at Cassidy

Bridge (off County Road 290), Oconee County,

South Carolina.

RANGE.—Tributaries of the Savannah River in

Oconee County, South Carolina.

HABITAT.—Streams.

Cambarus (Puncticambarus) extraneus Hagen

FIGURE 71

Cambarus extraneus Hagen, 1870:73, pi. 1: figs. 88, 89; pi. 3:

fig. 156.

Cambarus extranus.—Steele, 1902:7 [erroneous spelling].

Cambarus (Bartonius) extraneus.—Ortmann, 1905c: 120.

Cambarus (Cambarus) extraneus.—Fowler, 1912:341 [by implication].

Cambarus extranius.—Unestam, 1969:203 [erroneous spelling].

Cambarus (Puncticambarus) extraneus.—Hobbs, 1969b: 101,

figs, lb, 7, 13b, 14b, 17m.

TYPES—MCZ 175 (3 <$ H, 9), USNM (9).

TYPE-LOCALITY.—"Tennessee River, Georgia"

(Hagen). The emendation by Ortmann (1931) to

Etowah River, Rome, Floyd County, Georgia, must

be rejected because this crayfish does not occur in

the Coosa (Etowah) basin.

RANGE.—Known only from the Chickamauga

Creek basin in northwestern Georgia.

HABITAT.—Streams.

Cambarus (Puncticambarus) nerterius Hobbs

FIGURE 73

Cambarus nerterius Hobbs, 1964:189, figs. 1-10.

Cambarus (Puncticambarus) nerterius.—Hobbs, 1969b: 101,

figs. 7. 17n.

TYPES.—Holotype, allotype, and morphotype,

USNM 111295, 111296, 111297 ($1, $, all);

paratypes, USNM.

TYPE-LOCALITY.—Matt's Black Cave, 2 miles

south of Renick, Greenbrier County, West Virginia.

RANGE.—Greenbrier River drainage system, West

Virginia.

HABITAT.—Subterranean waters.

Cambarus (Puncticambarus) reburrus Prins

FIGURE 76

Cambarus reburrus Prins, 1968:458, figs. 1-11.

Cambarus (Puncticambarus) reburrus.—Prins and Hobbs,

1972:412.

TYPES.—Holotype, allotype, and morphotype,

USNM 129501, 129502, 129503 (a I, 9, $ II);

paratypes, USNM, MCZ, Rudolph Prins.

TYPE-LOCALITY.—Small tributary to Horsepasture River from Sapphire Lake off U.S. Highway

64, 5.5 miles east of Cashiers, Jackson County,

North Carolina (Savannah Drainage).

RANGE.—Type-locality and tributaries of the

French Broad River in Buncombe, Madison, and

Transylvania counties, North Carolina.

HABITAT.—Streams.

Cambarus (Puncticambarus) robustus Cirard

FICURE 77

Cambarus robustus Girard, 1852:90.—Hagen, 1870:80, pi. 3:
22 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

6g. 167.—Crocker and Barr, 1968:118, figs. 30, 39, 48, 55,

65, 67, 81.

Cambarus Bartonii robustus.—Faxon, 1885a:9.

Cambarus Bartonii var. robusta.—Faxon, 1885a:61.

Cambarus bartonii robustus.—Faxon, 1890:622.

Cambarus (Bartonius) bartoni robustus.—Ortmann, 1905c:

122 [by implication].

Cambarus bartoni robustus.—Ortmann, 1905c: 135.

Cambarus (Cambarus) bartonii robustus.—Fowler, 1912:341

[by implication].

Cambarus (Cambarus) bartoni robustus.—Ortmann, 1931:126.

Cambarus (Bartonius) robustus.—Creaser, 1931b:260.

Cambarus (Puncticambarus) robustus Hobbs, 1969b: 101,

figs. 1c, 13a, 17o.

TYPES.—Syntype, ANSP 328 ($ II); others probably destroyed in the Chicago fire, 1871.

TYPE-LOCALITY.—Humber River, near Toronto,

Canada.

RANGE.—Ontario and New York, to Illinois, and

southward to Tennessee, North Carolina, and Virginia.

HABITAT.—Streams.

Cambarus (Puncticambarus) spicatus Hobbs

FIGURE 72

Cambarus spicatus Hobbs, 1956c: 116, figs. 1-11.

Cambarus (Puncticambarus) spicatus.—Hobbs, 1969b: 102,

figs. 7, 18a.

TYPES.—Holotype, allotype, and morphotype,

USNM 99323, 99324, 99325 (a I, $, SU); paratypes, TU, USNM.

TYPE-LOCALITY.—Little River, 10.3 miles west of

Winnsboro, Fairfield County, South Carolina.

RANGE.—Little River in Fairfield and Richland

counties, South Carolina.

HABITAT Streams.

Cambarus (Puncticambarus) veteranus Faxon

FIGURE 78

Cambarus bartonii veteranus Faxon, 1914:389, pi. 13: fig. 2.

Cambarus bartoni veteranus.—Ortmann, 1931:112.

Cambarus (Cambarus) montanus veteranus.—Ortmann, 1931:

112 [in part].

Cambarus veteranus.—Hobbs, 1955b:330.

Cambarus (Puncticambarus) veteranus.—Hobbs, 1969b: 102,

figs. 7, 18b.

TYPES.—Holotype, USNM 44712 (£ I); paratypes, USNM, MCZ.

TYPE-LOCALITY.—Indian Creek, Baileysville,

Wyoming County, West Virginia.

RANGE.—Guyandot drainage in southern West

Virginia. Limits of range very indefinite and poorly

known.

HABITAT.—Streams.

Subgenus Veticambarus Hobbs

Veticambarus Hobbs, 1969b:96. [Type-species by original designation, Cambarus pristinus Hobbs, 1965:268. Gender:

masculine.]

Cambarus (Veticambarus) pristinus Hobbs

FIGURE 79

Cambarus pristinus Hobbs, 1965:268, figs. 1-12.

Cambarus (Veticambarus) pristinus.—Hobbs, 1969b:98, figs.

\e, 5, 13/, 14ft, 17a.

TYPES.—Holotype, allotype, morphotype, USNM

115528, 115529, 115530 ($ I, $, all); paratypes,

USNM, MCZ.

TYPE-LOCALITY.—White Oak Creek, a tributary

to the Caney Fork of the Cumberland River, 3.9

miles east of the White-Cumberland county line

and 0.1 mile south of U.S. Highway 70S, Cumberland County, Tennessee.

RANGE.—Three localities in the Caney Fork

drainage in Cumberland County, Tennessee.

HABITAT.—Lotic habitats, under stones.

Genus Fallicambarus Hobbs

Fallicambarus Hobbs, 1969b:111. [Type-species by original

designation, Cambarus straumi Reimer, 1966:11. Gender:

masculine.]

Fallicambarus byersi (Hobbs)

FIGURE 87

Cambarus byersi Hobbs, 1941:118, figs. 6, 10, 14, 17, 20, 23,

26, 27, 30; 1942b: 167, pi. 12: figs. 206-210.

Procambarus byersi Hobbs, 1942b: 14 [erroneous combination].

Fallicambarus byersi.—Hobbs, 1969b:lll, fig. 20c; 1972b:101,

figs. 81e, 83e.

TYPES.—Holotype, allotype, USNM 79342 (a I,

9 ), and "morphotype," USNM 87969 (ill);

paratypes, USNM, MCZ.

TYPE-LOCALITY.—Phifer Creek, 5.5 miles north-
NUMBER 166 23

west of Pensacola, near U.S. Highway 90, Escambia County, Florida.

RANGE.—Okaloosa County, Florida, west to Bay

St. Louis, Hancock County, Mississippi.

HABITAT.—Primary burrower.

Fallicambarus dissitus (Penn)

FIGURE 88

Cambarus dissitus Penn, 1955:73, figs. 1-13.

Fallicambarus dissitus.—Hobbs, 1969b:111, fig. 20d; 1972b:99,

figs. 81c, 82d.

TYPES.—Holotype, allotype, and morphotype,

USNM 98125, 98126, 98127 (a I, $, $ II); paratypes, ANSP, AMNH, CM, TU, USNM.

TYPE-LOCALITY.—Three miles east of Choudrant,

Lincoln Parish, Louisiana.

RANGE.—Caldwell and Lincoln parishes, Louisiana.

HABITAT.—Burrows (primary burrower).

Fallicambarus fodiens (Cottle)

FIGURE 82

Astacus fodiens Cottle, 1863:217.

Cambarus argillicola Faxon, 1884:115. [Types, MCZ 3459

($ I* 4 $, 3 juv.) . Type-locality, Detroit, Michigan.]

Cambarus (Bartonius) argillicola Ortmann, 1905c: 120.

Bartonius argillicola.—Williamson, 1907:749 [Bartonius intended as subgeneric name].

Cambarus fodiens.—Huntsman, 1915:158, figs. 8f, 9d, 10e, lla,

12e—Crocker and Barr, 1968:129-135, figs. 28, 37, 46, 57,

62, 85.

Cambarus (Bartonius) fodiens.—Creaser, 1931b:260, fig. 37.

Fallicambarus fodiens,—Hobbs, 1969b:lll, fig. 20e; 1972b:102,

figs. 83c, 84b, 85b.

TYPES.—If extant, location unknown.

TYPE-LOCALITY.—"Upper Canada." Probably

Ontario.

RANGE.—Lower Ontario, Michigan, Ohio, Indiana, and Illinois southward to Arkansas and

southwestern Georgia.

HABITAT.—Lentic and lotic habitats and burrows (primary or secondary burrower).

Fallicambarus hedgpethi (Hobbs)

FIGURE 83

Cambarus hedgpethi Hobbs, 1948d:224, fig. Ha-fJt-jJ-—

Penn, 1959:14, figs. 9, 27, 46, 64, 86,—Reimer, 1969:53, figs.

2, 39.

Fallicambarus hedgpethi,—Hobbs, 1969b:111, fig. 20f; 1972b:

102, figs. 82c, 83d.

TYPES.—Holotype, morphotype, USNM 85146

($ I, $ II), and allotype, USNM 85147 ( ?); paratypes, USNM.

TYPE-LOCALITY.—Lower middle part of the

Aransas National Wildlife Refuge, Aransas County,

Texas.

RANGE.—Texas and Oklahoma eastward to

Louisiana and western Tennessee.

HABITAT.—Temporary bodies of water and burrows (primary burrower).

Fallicambarus hortoni Hobbs and Fitzpatrick

FIGURE 85

Fallicambarus hortoni Hobbs and Fitzpatrick, 1970:829, figs.

1-12—Hobbs, 1972b: 100, fig. 81d.

TYPES.—Holotype, allotype, and morphotype,

USNM 129895, 129897, 129896 («$ I, $, $ II);

para types, USNM, J. F. Fitzpatrick, Jr.

TYPE-LOCALITY.—Low area along roadside ditch

leading to tributary of Cypress Creek, 7.5 miles

east of the Hardeman County line on State Route

57 (Hatchie River drainage), McNairy County,

Tennessee.

RANGE.—Known only from the type-locality.

HABITAT.—Burrows (primary burrower).

Fallicambarus macneesei (Black)

FIGURE 80

Cambarus macneesei Black, 1967:173, figs. 1-12.

Fallicambarus macneesei.—Hobbs, 1969b:111, fig. 20g; 1972b:

99, figs. 4c, 81a.

TYPES.—Holotype, allotype, and morphotype,

USNM 117681, 117683, 117682 (a I, $, all);

paratypes, USNM, TU, MCZ, Joe B. Black, J. F.

Fitzpatrick, Jr.

TYPE-LOCALITY.—Roadside ditches along East

McNeese Road, 1.8 miles west of intersection with

State Route 14, Lake Charles, Calcasieu Parish,

Louisiana.

RANGE.—Calcasieu River system in Calcasieu

Parish, Louisiana.
24 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

HABITAT.—Pools, roadside ditches, and burrows

(Psecondary burrower).

Fallicambarus oryktes (Penn and Marlow)

FIGURE 86

Cambarus oryktes Penn and Marlow, 1959:197, figs. 1-14.

Fallicambarus oryktes^-Hobbs, 1969b:111, fig. 20h; 1972b: 101,

figs. 81f, 83a.

TYPES.—Holotype, allotype, and morphotype,

USNM 101735, 101736, 101737 ($ I, 9, ill);

paratypes, TU, USNM.

TYPE-LOCALITY.—Roadside ditch at Alton, St.

Tammany Parish, Louisiana.

RANGE.—St. Tammany Parish, Louisiana, and

southern Mississippi.

HABITAT.—Burrows (primary burrower).

Fallicambarus strawni (Reimer)

FIGURE 81

Cambarus strawni Reimer, 1966:11, figs. 9-18.

Fallicambarus strawni Hobbs, 1969b:111, figs. 2f, 13j, 20i;

1972b:99, figs. 4a, 81b.

TYPES.—Holotype, allotype, and morphotype,

USNM 116675, 116676, 116677 (a I, 5, $ II); paratypes, R. D. Reimer.

TYPE-LOCALITY Small marshy area in the

Saline River drainage, 2.7 miles north of Dierks,

Howard County, Arkansas.

RANGE.—Sevier, Howard, and Pike counties,

Arkansas.

HABITAT.—Burrows, both simple and complex.

Fallicambarus uhleri (Faxon)

FIGURE 84

Cambarus Uhleri Faxon, 1884:116; 1885a:77, pi. 8: figs. 8, 8',

8a, 8a'.

Cambarus uhleri,—Hay, 1899b:959.—Meredith and Schwartz,

1960:28, figs. 11, 16.

Cambarus (Bartonius) uhleri.—Ortmann, 1905c: 120.

Cambarus (Cambarus) uhleri.—Fowler, 1912:341 [by implication].

Fallicambarus uhleri.—Hobbs, 1969b:111, fig. 20j; 1972b: 102,

figs. 82a, 83b, 84a, 85a.

TYPES.—Holotype, MCZ 3634 (a I); paratypes,

MCZ.

TYPE-LOCALITY.—"Swamp on Eastern Road near

Felsbury, Somerset County, Maryland" (restricted

by Faxon, 1914:426).

RANGE.—Atlantic coastal plain, from Maryland

to South Carolina.

HABITAT.—Lotic and lentic habitats and burrows (secondary burrower).

Genus Faxonella Creaser

Faxonella Creaser, 1933b:21. [Type-species by monotypy,

Cambarus clypeatus Hay, 1899a: 122. Gender: feminine.]

Faxonella beyeri (Penn)

FlCURE 91

Orconectes (Faxonella) beyeri Penn, 1950a: 166, figs. 1-9.

Faxonella beyeri.—Creaser, 1962:3 [by implication] Fitzpatrick, 1963:61 Hobbs, 1972b:29, fig. 19b.

TYPES.—Holotype, allotype, and morphotype,

USNM 90361, 90362, 90363 ($ I, $, all); paratypes, TU, USNM.

TYPE-LOCALITY.—Roadside ditch on U.S. Highway 84, 2 miles northeast of Naborton, De Soto

Parish, Louisiana.

RANGE.—Known only from De Soto and Natchitoches parishes, Louisiana.

HABITAT.—Roadside ditches.

Faxonella clypeata (Hay)

FIGURE 89

Cambarus clypeatus Hay, 1899a: 122, fig. 2.

Faxonius (Faxonella) clypeatus.—Creaser, 1933b: 19, pi. 1:

figs. 7, 8; pi. 2: figs. 1, 2.

Orconectes clypeatus.—Hobbs, 1942a:352 [by implication].

Orconectes (Faxonella) clypeata.—Hobbs, 1942b: 154, pi. 11:

figs. 181-185.

Orconectes (Faxonella) clypeatus.—Hobbs, 1959:890.

Faxonella clypeata.—Creaser, 1962:3 [by implication].—Fittpatrick, 1963:61.—Hobbs, 1972b:29, fig. 19c.

TYPES.—Holotype, USNM 17277 (9).

TYPE-LOCALITY.—Bay St. Louis, Hancock County, Mississippi.

RANGE.—Le Flore County, Oklahoma, and Marion County, Texas, east to Gadsden County,

Florida, and Richland County, South Carolina.

HABITAT.—Sluggish streams and lentic situations

and burrows (tertiary burrower).
NUMBER 166 25

Faxonella creaseri Walls

FIGURE 90

Faxonella creaseri Walls, 1968:413, figs. 1-9.—Hobbs, 1972b:

29, figs. 5w, 19a.

TYPES.—Holotype, allotype, and morphotype,

USNM 129215, 129216, 129214 ($ I, $, $ II);

paratypes, TU, MCZ, IBM, AMS, Joe B. Black,

J. F. Fitzpatrick, Jr., and Jerry G. Walls.

TYPE-LOCALITY.—Roadside ditch 3.0 miles north

of Grayson on U.S. Highway 165, Caldwell Parish,

Louisiana.

RANGE.—Known only from several localities in

Caldwell Parish, Louisiana.

HABITAT.—Temporary pools in roadside ditches.

Genus Hobbseus Fitzpatrick and Payne

Hobbseus Fitzpatrick and Payne, 1968:15. [Type-species by

original designation, Cambarus cristatus Hobbs, 1955a:95.

Gender: masculine.]

Hobbseus attenuatus Black

FIGURE 95

Hobbseus attenuatus Black, 1969:193, figs. 1-12.—Hobbs,

1972b: 75, fig. 58c.

TYPES.—Holotype, allotype, and morphotype,

USNM 129535, 129536, 129537 ($1, 9, ill);

paratypes, USNM, MCZ, TU, Joe B. Black, J. F.

Fitzpatrick, Jr.

TYPE-LOCALITY.—"Roadside ditch adjacent to

Noxapater Creek, on gravel road two miles north

of State Route 395, six miles west of Noxapater,

Winston County, Mississippi (R. 11 E, T. 14 N)."

RANGE.—Pearl River drainage in Winston and

Neshoba counties, Mississippi.

HABITAT.—Lentic situations.

Hobbseus cristatus (Hobbs)

FIGURE 93

Cambarus cristatus Hobbs, 1955a:95, figs. 1-11.

Hobbseus cristatus.—Fitzpatrick and Payne, 1968:15 Hobbs,

1972b: 75, figs. 5o, 58b.

TYPES.—Holotype, allotype, and morphotype,

USNM 96985, 96986, 96987 ($1, 9, 3 II); paratypes, MCZ, TU, USNM.

TYPE-LOCALITY.—Roadside ditch, 11.3 miles

south of Macon on U.S. Highway 45, Noxubee

County, Mississippi.

RANGE.—Kemper, Lauderdale, Lowndes, and

Noxubee counties, Mississippi.

HABITAT.—Lentic and lotic habitats and burrows (probably secondary burrower).

Hobbseus orconectoides Fitzpatrick and Payne

FIGURE 92

Hobbseus orconectoides Fitzpatrick and Payne, 1968:17, figs.

1-12.—Hobbs, 1972b:75, fig. 58a.

TYPES.—Holotype, allotype, and morphotype,

USNM 129191, 129193, 129194 ($ I, $, ^ II);

paratypes, MCZ, TU, OSM, USNM, J. F. Fitzpatrick, Jr.

TYPE-LOCALITY.—Woodland pond near Rock

Hill community, about 4.25 miles north of Starkville, Oktibbeha County, Mississippi (T. 19 N.,

R. 14 E., sec. 1).

RANGE.—Known only from the vicinity of the

type-locality.

HABITAT.—Lentic situations and burrows (probably secondary burrower).

Hobbseus prominens (Hobbs)

FIGURE 94

Cambarus prominens Hobbs, 1966a: 110, figs. 1-10.

Hobbseus prominens.—Fitzpatrick and Payne, 1968:15.—

Hobbs, 1972b:76, figs. 58e, 59b.

TYPES.—Holotype, allotype, and morphotype,

USNM 115603, 115604, 115605 (<$ I. 9, ill);

paratypes, USNM, MCZ.

TYPE-LOCALITY.—Roadside ditch, 3 miles west

of Demopolis, on U.S. Highway 80, Sumter County,

Alabama.

RANGE.—Mobile River drainage in Choctaw,

Clarke, Dallas, Hale, Marengo, Perry, and Sumter

counties, Alabama.

HABITAT.—Lentic and lotic habitats and burrows (probably secondary burrower).

Hobbseus vaUeculus (Fitzpatrick)

FIGURE %

Cambarus vaUeculus Fitzpatrick, 1967b: 163-167, figs. 1-12.
26 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Hobbseus valleculus.—Fitzpatrick and Payne, 1968:15—

Hobbs, 1972b: 76, figs. 58d, 59a.

TYPES.—Holotype, allotype, and morphotype,

USNM 117963, 117964, 117965 ($ I, S>, $ II);

paratypes, MCZ, TU, J. F. Fitzpatrick, Jr.

TYPE-LOCALITY.—Small creek, 6.8 miles south of

junction of State Routes 15 and 12 on State Route

15, Choctaw County, Mississippi.

RANGE.—Known only from the vicinity of the

type-locality, tributaries to Pearl River.

HABITAT.—Streams.

Genus Orconectes Cope

Astacus.—Rafinesque, 1817:42.

Orconectes Cope, 1872:419. [Type-species by subsequent designation (Fowler, 1912:339), Orconectes inermis Cope, 1872:

419. Gender: masculine.]

Camtarus Packard, 1888:156 [erroneous spelling].

Camborus Williamson, 1899:47 [erroneous spelling].

Cambrus Price, 1900:155 [erroneous spelling].

Orconectis Harris, 1903:113 [erroneous spelling].

Faxonius Ortmann, 1905c:97. [Type-species by original designation, Astacus limosus Rafinesque, 1817:42. Gender: masculine. Proposed as subgenus of Cambarus, elevated to

generic rank by Creaser, 1933a: 1.]

G[ambarus] Ortmann, 1905c: 112 [erroneous spelling].

Cambarrus Fleming, 1939:305 [erroneous spelTing].

Orconecctes Williams, 1952:330 [erroneous spelling].

Gambarus Croizat, 1958:908 [erroneous spelling].

Orconnectes Threinen, 1958:1 [erroneous spelling].

Camburus Bacescu, 1967:218 [erroneous spelling].

Orchonectes Dimond et al., 1968:760 [erroneous spelling].

Orconectus Ray and Stevens, 1970:58 [erroneous spelling].

Orconetes Avault, 1973:242 [erroneous spelling].

REMARKS.—Hobbs (1942a: 350) erroneously indicated that Orconectes inermis Cope was the typespecies by monotypy. Cope (1872:419) stated that

"The present genus embraces two species, O.

inermis of the Wyandotte and the O. pellucidus

of the Mammoth Cave."

Orconectes acares Fitzpatrick

FIGURE 132

Orconectes leptogonopodus acares Fitzpatrick, 1965:87, figs.

A-J.

Orconectes acares.—Hobbs, 1972b:94, fig. 74f.

TYPES.—Holotype, allotype, and morphotype,

USNM 115517, 115518, 115519 ($ I, $, 4II);

paratypes, USNM, MCZ, TU.

TYPE-LOCALITY.—Stream tributary to Ouachita

River, 6 miles northwest of Mount Ida, Montgomery County, Arkansas.

RANGE.—Tributaries of the Ouachita River in

Garland, Hot Springs, Montgomery, Perry, Pike,

Polk, and Saline counties, Arkansas.

HABITAT.—Streams.

Orconectes alabamensis (Faxon)

FIGURE 166

Cambarus Alabamensis Faxon, 1884:125; 1885a: 104, pi. 4: fig.

4; pi. 10: figs. 3, 3', 3a, 3a'.

Cambarus alabamensis.—Hay, 1899b:960.

Cambarus (Faxonius) alabamensis Ortmann, 1905c: 112.

Faxonius alabamensis Creaser, 1933a:3 [by implication];

1962:2 [by implication].

Faxonius (Faxonius) alabamensis.—Creaser, 1933b: 16 [by implication].

Orconectes alabamensis.—Hobbs, 1942a:352 [by implication].—Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) alabamensis.—Hobbs, 1942b: 154 [by

implication].

TYPES.—Syntypes, USNM 4876 (14 31. 16 9,

4 $ II), MCZ 3565 (2 <$ I, 2 $ , 2$ II).

TYPE-LOCALITY.—Second Creek, Waterloo, Lauderdale County, Alabama.

RANGE.—Known only from tributaries of the

Tennessee River along the Alabama-TennesseeMississippi border.

HABITAT.—Streams.

Orconectes australis australis (Rhoades)

FIGURE 97

Cambarus (Faxonius) pellucidus australis Rhoades, 1941b:

142, fig. 35A-F.

Orconectes pellucidus australis.—Hobbs, 1942a:353; 1948a:20,

figs. 6, 13.—Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) pellucidus australis.—Hobbs, 1942b:

154 [by implication].

Orconectes PeZZuridus—Cooper and Poulson, 1968:130, fig. 8.

Orconectes australis australis.—Hobbs and Barr, 1972:3, 11,

figs. 2, 3, 5o-v, 6, 7, 8i-cc.

TYPES.—Holotype, allotype, and "morphotype,"

USNM 79363, 79364, 79365 (<$ I, $, $U); paratypes, RR, ANSP, Alabama Mus. Nat. Hist., L.

Hubricht.

TYPE-LOCALITY.—Shelta Cavern (SE14NE14

sec. 27, T. 3 S., R. 1 W.), north of Huntsville, Madison County, Alabama.
NUMBER 166 27

RANGE.—Madison and Jackson counties, Alabama, north to middle Tennessee where it intergrades with O. australis packardi.

HABITAT.—Subterranean streams and pools.

Orconectes australis packardi Rhoades

FIGURE 98

Orconectes pellucidus packardi Rhoades, 1944a: 121, fig.

3a-f.—Hobbs, 1948a:20, figs. 8, 11 Fitzpatrick, 1963:61

[by implication].

Orconectes (Orconectes) pellucidus packardi.—Hobbs, 1959:

890 [by implication].

Orconectes pellucidus packardii.—Nicholas, 1960:133.

Orconectes australis packardi.—Hobbs and Barr, 1972:4, 22,

figs. 2, 4, 5a-n, 8a-h.

TYPES.—Holotype, allotype, and "morphotype,"

USNM 81310, 81312, 81311 ($ I, $, $ II); paratypes, MCZ, USNM, RR.

TYPE-LOCALITY.—Cumberland Crystal Cave

(Sloans Valley Cave), Alpine, Pulaski County, Kentucky.

RANGE.—Pulaski and McCreary counties, Kentucky, and south to Tennessee where it intergrades

with O. australis australis.

HABITAT.—Subterranean streams.

Orconectes barrenensis Rhoades

FIGURE 139

Orconectes rusticus barrenensis Rhoades, 1944a: 125, fig.

6A-F.—Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) rus-ticus barrenensis.—Hobbs, 1959:

893 [by implication].

Faxonius rusticus barrenensis.—Creaser, 1962:2 [by implication].

Orconectes rusticus barrensis Fitzpatrick, 1967a: 143 (erroneous spelling].

Orconectes barrenensis.—Hobbs, 1972b:97, figs. 74h, 76b.

TYPES.—Holotype, allotype, and "morphotype"

(designated as allotype by Rhoades), USNM 81319,

81321,81320 ($ I, $, ^ II); paratypes, CM, MCZ,

USNM, RR.

TYPE-LOCALITY.—Barren River, Beech Bend, 2

miles north of Bowling Green, Warren County,

Kentucky.

RANGE.—Barren River drainage in Kentucky and

Tennessee, and in the Green River system in Kentucky.

HABITAT.—Swift streams.

Orconectes bisectus Rhoades

FIGURE 125

Orconectes bisectus Rhoades, 1944a: 129, fig. 7A-D Fitzpatrick, 1963:61 [by implication].—Prins and Fitzpatrick,

1965:141, figs. 1-12.

Orconectes (Orconectes) bisectus.—Hobbs, 1959:890 [by implication].

Faxonius bisectus.—Creaser, 1962:2 [by implication].

TYPES.—Holotype and allotype, USNM 81322,

81323 (<$H, 9); paratypes, MCZ, RR.

TYPE-LOCALITY.—Brushy Fork, 1 mile west of

Repton, Crittenden County, Kentucky.

RANGE.—Crooked Creek drainage in Crittenden

County, Kentucky.

HABITAT.—Medium-sized streams.

Orconectes causeyi Jester

FIGURE 163

Orconectes causeyi Jester, 1967:518, figs. 1-12 Reimer, 1969:

52, figs. 4, 10, 24, 37.

TYPES.—Holotype, allotype, and morphotype,

USNM 115698, 115699, 115700 ($ I, $, ill) ;

paratypes, USNM, TU.

TYPE-LOCALITY.—Conchas Reservoir, 32 miles

northwest of Tucumcari on State Route 104, San

Miguel County, New Mexico.

RANGE.—"New Mexico, Texas, Arkansas, Oklahoma, Missouri, and Kansas" (Reimer, 1969).

HABITAT.—Streams and impounded lakes.

REMARKS.—This crayfish, insofar as I am able

to determine, is indistinguishable from Orconectes

virilis (Hagen). Until the relationships of these

two species and O. nais (Faxon) become more adequately understood, the conservative recognition

of the three seems advisable.

Orconectes compressus (Faxon)

FIGURE 169

Cambarus compressus Faxon, 1884:127; 1885a: 105, pi. 5: fig.

6; pi. 10: figs 2, 2', 2a, 2a'.

Cambarus (Faxonius) compressus Ortmann, 1905c: 112.

Faxonius compressus.—Creaser, 1933a:3 [by implication];

1962:2 [by implication].

Faxonius (Faxonius) compressus.—Creaser, 1933b: 16 [by implication].

Orconectes compressus.—Hobbs, 1942a:352 [by implication];

1949a: 17.—Fittpatrick, 1963:61 [by implication].
28 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Orconectes (Orconectes) compressus.-

implication].

-Hobbs, 1942b: 154 [by

TYPES.—Syntypes, USNM 4878 (11*1, 13$),

USNM 4879 (5 <$ I, 3 9 , $11), MCZ 3583 (3 $ I,

2 9 , $ II).

TYPE-LOCALITY.—Second Creek, Waterloo, Lauderdale County, Alabama. Restricted by Faxon

(1914:419).

RANGE.—Tributaries of the Tennessee River in

Alabama, Mississippi, and Tennessee; tributaries

of the Cumberland River in Tennessee and Kentucky; and tributaries of the Barren River in Tennessee and Kentucky.

HABITAT.—Clear, rock- or gravel-bedded streams.

Orconectes difficilis blacki Walls

FIGURE 107

Orconectes difficilis blacki Walls, 1972:454, figs. 3A-E, 4A.

TYPES.—Holotype, USNM 131255 ($1); paratypes, USNM, MCZ, TU, J. B. Black, J. F. Fitzpatrick, Jr., and J. G. Walls.

TYPE-LOCALITY.—Bearhead Creek, 4 miles southwest of Juanita on State Route 109, Beauregard

Parish, Louisiana.

RANGE.—Known only from six localities in Beauregard and Calcasieu parishes, Louisiana, and intergrading with O. d. hathawayi "in the Calcasieu

River drainage of Allen, Beauregard, Calcasieu,

and Vernon Parishes, Louisiana" (Walls, 1972:

455).

HABITAT.—Streams.

Orconectes difficilis difficilis (Faxon)

FIGURE 106

Cambarus difficilis Faxon, 1898:656, pi. 65: figs. 1-4.

Cambarus (Faxonius) difficilis.—Ortmann, 1905c: 113.

Faxonius difficilis.—Creaser, 1933a:3 [by implication]; 1962:2

[by implication].

Faxonius (Faxonius) difficilis.—Creaser, 1933b: 16 [by implication].

Cambarus difficulis.—Fleming, 1938:300 [erroneous spelling].

Orconectes difficilis.—Hobbs, 1942a:352 [by implication].

Williams, 1954:898, figs. 217-224, 233.—Fitzpatrick, 1963:61

[by implication].

Orconectes (Orconectes) difficilis Hobbs, 1942b: 154 [by implication]; 1959:892.

Orconectes difficilis difficilis—Walk, 1972:456.

TYPES.—Syntypes, USNM 19687 (9^1, 10 9,

5 $ II), MCZ 4359 ($1, 9 , $ II), Mus. Zool. Torino.

TYPE-LOCALITY.—McAlester, Pittsburg County,

Oklahoma.

RANGE.—Known from Pittsburg and Latimer

counties, Oklahoma, Upshur County, Texas, to

Washington County, Arkansas, southeastward to

Louisiana (see Walls, 1972).

HABITAT.—Rocky streams.

Orconectes difficilis hathawayi Perm

FIGURE 109

Orconectes (Orconectes) hathawayi Penn, 1952b: 1, figs. 1-14.

Faxonius hathawayi Creaser, 1962:2 [by implication].

Orconectes hathawayi.—Fitzpatrick, 1963:61 [by implication].

Orconectes difficilis hathawayi.—Walls, 1972:453.

TYPES.—Holotype, allotype, and morphotype,

USNM 92998, 92999, 93000, ( $ I, 9 , 3 II); paratypes, MCZ, CM, ANSP, USNM, AMNH, TU.

TYPE-LOCALITY.—Spring Creek, 5.1 miles east of

Hineston on State Route 85, Rapides Parish,

Louisiana.

RANGE.—Jackson and Rapides parishes south to

Vermilion Parish, Louisiana; intergrading with

O. d. blacki in the Calcasieu River system in Allen,

Beauregard, Calcasieu, and Vernon parishes.

HABITAT.—Streams.

Orconectes difficilis maletae Walls

FIGURE 108

Orconectes difficilis maletae Walls, 1972:456, figs. 2A-E, 4B.

TYPES.—Holotype, USNM 131254 ($1); paratypes, USNM, MCZ, TU, J. B. Black, J. F. Fitzpatrick, Jr., and J. G. Walls.

TYPE-LOCALITY.—Bayou Santabarb " (third

branch north of Kisatchie)" on State Route 117,

Natchitoches Parish, Louisiana.

RANGE.—Known from only seven localities in

Natchitoches and Sabine parishes, Louisiana, and

Upshur County, Texas. Specimens from Coal County, Oklahoma, are interpreted as intergrades between this and the nominate subspecies.

HABITAT.—Streams (?).
NUMBER 166 29

Orconectes erichsonianus (Faxon)

FIGURE 123

Cambarus erichsonianus Faxon, 1898:659, pi. 64: figs. 7-12.

Cambarus erichsoni.—Ortmann, 1905c: 109 [erroneous spelling]-

Cambarus (Faxonius) erichsonianus.—Ortmann, 1905c: 112.

Cambarus spinosus gulielmi Faxon, 1914:375. [Syntypes,

USNM 26379 (6 $11, 6$ , 4$juv.), MCZ 7448 ($11, $) .

Type-locality, stream from John Ross Spring, near Rossville, Walker County, Georgia.]

Faxonius erichsonianus.—Creaser, 1933a:3 [by implication];

1962:2 [by implication].

Faxonius (Faxonius) erichsonianus.—Creaser, 1933b:7 [by

implication].

Cambarus erichsonionus.—Fleming, 1938:299 [erroneous spelling]-

Orconectes erichsonianus.—Hobbs, 1942a:352 [by implication].—Fitzpatrick, 1963:61 [by implication]; 1967a: 131,

147, figs. 2-18, 20.

Orconectes (Orconectes) erichsonianus.—Hobbs, 1942b: 154

[by implication].

TYPES.—Syntypes, USNM 20787 (3 41, 4$),

MCZ 4347 ($1, 9, SlI).

TYPE-LOCALITY.—Rip Roaring Fork, 5 miles

northwest of Greeneville, Greene County, Tennessee.

RANGE.—Tennessee, Elk, and Coosa river systems in eastern Tennessee, northwestern Georgia,

and northern Alabama.

HABITAT.—Streams.

Orconectes eupunctus Williams

FIGURE 116

Orconecctes eupunctus Williams, 1952:330 [erroneous spelling

of generic name].

Orconectes eupunctus Williams, 1952:334, 335, pi. 1: figs. 1-8;

1954:840, figs. 41-49.—Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) eupunctus.—Hobbs, 1959:894.

Faxonius eupunctus.—Creaser, 1962:2 [by implication].

TYPES.—Holotype, allotype, and "morphotype,"

USNM 129200 ($1, 9, $11); paratypes, USNM,

MCZ.

TYPE-LOCALITY.—Eleven Point River at Riverton, Oregon County, Missouri.

RANGE Spring River and Eleven Point drainage systems in Arkansas and Missouri.

HABITAT.—Clear, cold, rapid streams.

Orconectes forceps (Faxon)

FIGURE 137

Cambarus forceps Faxon, 1884:133; 1885a: 119, pi. 5: fig. 4; pi.

9: figs. 5, 5', 5a, 5a'.

Cambarus (Faxonius) forceps.—Ortmann, 1905c: 112.

Cambarus (Faxonius) rusticus forceps.—Ortmann, 1931:72.

Faxonius forceps.—Creaser, 1933a:3 [by implication]; 1962:2

[by implication].

Faxonius (Faxonius) forceps.—Creaser, 1933b: 10 [by implication].

Orconectes rusticus forceps.—Hobbs, 1942a: 352 [by implication].—Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) rusticus forceps.—Hobbs, 1942b: 154

[by implication].

Orconectes forceps.—Fitzpatrick, 1967a: 143.—Hobbs, 1972b:

98, figs. 77i, 80d.

TYPES.—Syntypes, USNM 4880 (3 $ I, 4 9 ), MCZ

3582 (si, 9).

TYPE-LOCALITY.—Cypress Creek, Lauderdale

County, Alabama.

RANGE.—Tennessee River drainage, chiefly above

Walden Gorge, from northern Alabama to southwestern Virginia.

HABITAT.—Streams.

Orconectes harrisonii (Faxon)

FIGURE 111

Cambarus Harrisonii Faxon, 1884:130; 1885a:94, pi. 3: fig. 1;

pi. 9: figs. 9, 9'.

Cambarus harrisonii.—Hay, 1899b:960.

Cambarus harrisoni.—Ortmann, 1902:278.

Cambarus (Faxonius) harrisoni.—Ortmann, 1905c:111.

Faxonius harrisoni.—Creaser, 1933a:3 [by implication]; 1962:

2 [by implication].

Faxonius (Faxonius) harrisoni.—Creaser, 1933b:21 [by implication].

Faxonius harrisonii.—Creaser, 1934a: 1.

Orconectes harrisoni.—Hobbs, 1942a:352 [by implication].

Orconectes (Orconectes) harrisoni.—Hobbs, 1942b: 154 [by

implication].

Orconectes harrisonii.—Williams, 1954:839, figs. 35—41.—

Fitzpatrick, 1963:61 [by implication].

TYPES.—Syntypes 3586 I, 5 9), , MCZ

USNM 25826 ( $ I, $).

TYPE-LOCALITY.—Irondale, Washington County,

Missouri.

RANGE.—St. Genevieve and Washington counties, Missouri.

HABITAT.—Streams.

Orconectes hobbsi Penn

FIGURE 160

Orconectes hobbsi Penn, 1950c:381. figs. 1-10; 1959:18, figs. 32,

52, 68, 82.—Fitzpatrick, 1963:61 [by implication].
30 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Orconectes (Orconectes) hobbsi.—Hobbs, 1959:895.

Faxonius hobbsi.—Creaser, 1962:2 [by implication].

TYPES.—Holotype, allotype, and morphotype,

USNM 90952, 90953, 90954 ($1, 9, $ II); paratypes, MCZ, ANSP, USNM, TU, AMNH.

TYPE-LOCALITY.—Headwater creek of Bayou Lacombe at St. Tammany, 6 miles north of Lacombe

on State Route 187, St. Tammany Parish, Louisiana.

RANGE.—Lake Pontchartrain watershed in Louisiana and Mississippi.

HABITAT.—Rapidly flowing streams.

Orconectes hylas (Faxon)

FIGURE 134

Cambarus hylas Faxon, 1890:632.

Cambarus (Faxonius) hylas.—Ortmann, 1905c: 112.

Faxonius hylas Creaser, 1933a:3 [by implication]; 1962:2 [by

implication].

Faxonius (Faxonius) hylas.—Creaser, 1933b:21 [by implication].

Orconectes hylas.—Hobbs, 1942a:352 [by implication] Williams, 1954:847, figs. 74-82.—Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) hylas.—Hobbs, 1942b: 154 [by implication].

TYPES.—Syntypes, MCZ 3858 (3 $ II, $ ), USNM

25827 ($11, $> ) .

TYPE-LOCALITY.—West Fork of Black River,

Reynolds County, Missouri.

RANGE.—Tributaries of the Big and Black rivers

in southeastern Missouri.

HABITAT.—Streams.

Orconectes illinoiensis Brown

FIGURE 128

Orconectes illinoiensis Brown, 1956:163, figs. 1-9.

Faxonius illinoiensis Creaser, 1962:2 [by implication].

Orconectes illinoisensis.—Hobbs and Fitzpatrick, 1962:207

[erroneous spelling].—Fitzpatrick, 1967a: 131, 149, figs. 2-18,

21.

TYPES.—Holotype, allotype, and morphotype,

USNM 97997, 97998, 97999 (<$ I, $ , $ II); paratypes, Max R. Matteson, Troy C. Dorris, USNM.

TYPE-LOCALITY.—Cypress Creek, 3.25 miles south

of Mount Pleasant, Union County, Illinois.

RANGE.—Alexander, Gallatin, Hardin, Pope, and

Union counties, Illinois.

HABITAT—Streams.

Orconectes immunis (Hagen)

FIGURE 164

Cambarus immunis Hagen, 1870:71, pi. 1: figs. 101, 102; pi.

3: fig. 160; pi. 8: fig. b.

Cambarus signifer Herrick, 1882:253, fig. 7. [Types, MCZ

3515 (2«$I, $) . Type-locality, Grass Lake, Richfield,

Hennepin County, Minnesota.]

Cambarus immunis spinirostris Faxon, 1884:146. [Types,

USNM 4655 (6<$II, 8$), MCZ 3562 (2 £11, 2$). Typelocality, Reelfoot Lake, Obion County, Tennessee.]

Cambarus immunis immunis Faxon, 1884:146 [by implication].

Cambarus immunis var. spinosiris Harris, 1901b: 115 [erroneous spelling].

Cambarus rinmunis Steele, 1902:3 [erroneous spelling].

Cambarus (Faxonius) immunis.—Ortmann, 1905c:113.

Faxonius immunis.—Williamson, 1907:749 [Faxonius intended

as subgeneric name].—Creaser, 1933a:4; 1962:2.

Faxonius (Faxonius) immunis immunis Creaser, 1933b:21

[by implication].

Faxonius (Faxonius) immunis pedianus Creaser, 1933b: 14.

[Subgenus by implication. Holotype and allotype, USNM

98334, 98335 ($1, $) ; paratypes, USNM. Type-locality,

reservoir at Englewood, Denver, Colorado.]

Cambarus immunis Hagenianus.—Fleming, 1938:303 [lapsus

for C. immunis Hagen].

Orconectes immunis immunis.—Hobbs, 1942a:352 [by implication]—Williams, 1954:890, figs. 200-208 Fitzpatrick,

1963:61 [by implication].

Orconectes immunis pedianus.—Hobbs, 1942a:352.

Orconectes (Orconectes) immunis immunis Hobbs, 1942b:

154 [by implication].

Orconectes immunis.—Penn, 195Od:645.

Orconnectes immunis Threinen, 1958:2 [erroneous spelling].

Orconectes (Orconectes) immunis.—Hobbs, 1959:894.

Orchonectes immunis.—Dimond et al., 1968:760 [erroneous

spelling of generic name].

TYPES.—Syntypes, MCZ 188 (2 $ I, 2 9 ). The

"paratypes" of Faxon (1914:421), MCZ 3355, are

dubious "types" because this number was not listed

by Hagen (1870:73). The "paratype" said to be

in MHNP could not be located.

TYPE-LOCALITY.—Lawn Ridge, Marshall County,

Illinois.

RANGE.—Massachusetts to Wyoming and Alabama to Ontario.

HABITAT.—Ponds, roadside ditches, and sluggish

streams.

Orconectes incomptus Hobbs and Barr

FIGURE 99

Orconectes incomptus Hobbs and Barr, 1972:32, fig. 9.
NUMBER 166 31

TYPES.—Holotype, allotype, and morphotype,

USNM 130299, 130300, 130301 ($ I, 9, $ II);

paratypes, USNM, H. H. Hobbs III.

TYPE-LOCALITY.—Cherry Cave, lat. 36°28'09" N.,

long. 85°36'28" W., Jackson County, Tennessee.

RANGE.—Known only from three localities in

Jackson County, Tennessee.

HABITAT.—Subterranean waters.

Orconectes indianensis (Hay)

FIGURE 105

Cambarus indianensis Hay, 1896:494, fig. 9.

Cambarus (Faxonius) indianensis.—Ortmann, 1905c: 112.

Faxonius indianensis.—Creaser, 1933a:3 [by implication];

1962:2 [by implication].

Faxonius (Faxonius) indianensis.—Creaser, 1933b:21 [by implication].

Orconectes indianensis.—Hobbs, 1942a:352 [by implication].—Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) indianensis.—Hobbs, 1942b: 154 [by

implication].

TYPES.—Syntypes, USNM 14624 (2*1, 2 9),

MCZ 3859 (2 3 I, 2 9 )•

TYPE-LOCALITY.—Patoka River, Patoka, Gibson

County, Indiana.

RANGE.—Southern Illinois and Indiana.

HABITAT.—Streams.

westward to Crawford and Harrison counties, Indiana, where it intergrades with O. inermis testii.

HABITAT.—Subterranean waters.

Orconectes inermis testii (Hay)

FIGURE 101

Cambarus peltucidus var. testii Hay, 1891:148.

Cambarus pellucidus testii.—Hay, 1893:285.

Cambarus pellucidus testi.—Hay, 1902a:233.

Cambarus (Faxonius) pellucidus testi.—Ortmann, 1931:64 [by

implication].

Orconectes pellucidus testii.—Hobbs, 1942a: 352 [by implication].

Orconectes (Orconectes) pellucidus testii.—Hobbs, 1942b: 154

[by implication].

Orconectes pellucidus inermis.—Rhoades, 1959:401 [in part].

Orconectes inermis testii.—Hobbs and Barr, 1972:4, 49, figs.

1, 11, 12a,b.

TYPES.—Syntypes, USNM 17702 (2 ^ II, 9),

MCZ 7431 (3II, 9).

TYPE-LOCALITY.—Mayfield's Cave, SW14NE14

SW14 sec. 26, T. 9 N., R. 2 W., Monroe County,

Indiana.

RANGE.—Monroe County, Indiana, intergrading

immediately to the south with the typical subspecies.

HABITAT.—Subterranean waters.

Orconectes inermis inermis Cope

FIGURE 100

Orconectes inermis Cope, 1872:419 Hobbs, 1948a:20, figs.

10, 14.

Cambarus pellucidus inermis.—Faxon, 1885a:83 Packard,

1888:41.

Cambarus pellucidus variety inermis.—Hay, 1893:284.

Cambarus inermis.—Faxon, 1898:647.

Orconectis inermis.—Harris, 1903:113 [erroneous spelling].

Orconectes (Orconectes) inermis.—Hobbs, 1942b: 154 [by implication].

Orconectes pellucidus inermis.—Rhoades, 1959:401 [in part].

Orconectes inermis inermis.—Hobbs and Barr, 1972:4, 36, figs.

1, 10, 12c-z, 13.

TYPES.—Not extant. Topotypes, USNM 131405

(3 I, 9, $ II).

TYPE-LOCALITY.—Wyandotte Cave, Indiana. Actually Sibert's Well Cave, just below Wyandotte

Cave, Crawford County, Indiana, according to

Hobbs (1942a: 335).

RANGE.—From Hart County, Kentucky, northOrconectes iowaensis Fitzpatrick

FIGURE 127

Orconectes species A Fitzpatrick, 1967a:150-151, 167, 168,

169, figs. 2-18, 27.

Orconectes iowaensis Fitzpatrick, 1968:507, figs. 1-10.

TYPES.—Holotype, allotype, and morphotype,

USNM 117963, 117964, 117965 ($ I, 9, <S II);

paratypes, MCZ, Coe College, USNM.

TYPE-LOCALITY.—Turkey River, 2.5 miles south

of Cresco, Howard County, Iowa.

RANGE.—Mississippi drainage system in eastern

Iowa.

HABITAT.—Streams.

Orconectes jeffersoni Rhoades

FIGURE 119

Orconectes propinquus jeffersoni Rhoades, 1944a: 123, fig.

5A-F.—Fitzpatrick, 1963:61 [by implication].
32 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Orconectes (Orconectes) propinquus jeffersoni.—Hobbs, 1959:

894 [by implication].

Faxonius propinquus jeffersoni Creaser, 1962:2 [by implication].

Orconectes jeffersoni.—Fitzpatrick, 1967a:131, 152, figs. 2-18,

22.

TYPES.—Holotype, allotype, and "morphotype"

(designated by Rhoades as allotype), USNM 81316,

81318,81317 (3 1, 9, $ II); paratypes, MCZ, CM,

USNM, RR.

TYPE-LOCALITY.—Tributary of Muddy Fork of

Beargrass Creek, 2 miles east of Louisville corporation line on U.S. Highway 42, Jefferson County,

Kentucky.

RANGE.—Tributaries to Beargrass Creek, Ohio

River, Jefferson County, Kentucky.

HABITAT.—Streams.

Orconectes juvenilis (Hagen)

FIGURE 147

Cambarus juvenilis Hagen, 1870:66, pi. 1: figs. 29-33; pi. 3:

fig. 157.

Cambarus juvenalis.—Harris, 1900:271 [erroneous spelling].

Cambarus (Faxonius) juvenilis.—Ortmann, 1931:84 [in part].

Faxonius juvenilis.—Creaser, 1933a:3 [by implication]; 1962:

2 [by implication].

Faxonius (Faxonius) juvenilis.—Creaser, 1933b:21 [by implication].

Orconectes juvenilis.—Hobbs, 1942a:352 [by implication].—

Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) juvenilis Hobbs, 1942b: 154 [by implication].

Orconectes Juvenils.—Hobbs and Walton, 1966:136 [erroneous spelling in title].

Orconectes juvenalis—Merkle, 1969:228 [erroneous spelling].

TYPES.—Syntypes, MCZ 3347 (26,} II, $,8 3

juv.), USNM 4967 ( $ I), USNM 25830 ( $ I, 9),

MHNP (2<J, 2 9). Other types cited by Faxon

(1914:418) apparently no longer extant.

TYPE-LOCALITY.—Kentucky River, Little Hickman, Jessamine County, Kentucky.

RANGE.—Streams in the Ohio drainage system

in northern Kentucky and in Indiana (?). The limits of the ranges of O. juvenilis and the closely

related O. putnami and O. spinosus have not been

determined.

HABITAT.—Streams.

REMARKS.—Following Ortmann (1931), O. putnami and O. spinosus were considered synonyms

of O. juvenilis. Ortmann considered O. spinosus

Bundy (not Faxon) to be a "doubtful form" but

indicated that "it may be different from juvenilis."

Orconectes kentuckiensis Rhoades

FlCURE 113

Orconectes kentuckiensis Rhoades, 1944a: 122, fig. 4A-F.—

Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) kentuckiensis.—Hobbs, 1959:894.

Faxonius kentuckiensis.—Creaser, 1962:2 [by implication].

TYPES.—Holotype, allotype, and "morphotype"

(designated as allotype by Rhoades), USNM 81313,

81315, 81314 ($ I, 9, <$H); paratypes, MCZ,

ANSP, RR.

TYPE-LOCALITY.—Piney Creek, 3 miles west of

Shady Grove, Crittenden County, Kentucky.

RANGE.—Crittenden and Union counties, Kentucky.

HABITAT.—Mud-bottomed streams.

Orconectes lancifer (Hagen)

FlCURE 114

Cambarus lancifer Hagen, 1870:59, pi. 1: figs. 86, 87; pi. 3:

fig. 159.

Cambarus faxonii Meek, 1894:1042, figs. 1-4. [Syntypes, MCZ

4220 (43II, 3 9) , USNM 19331 (63,$ 9) , USNM 22425

(<$II, 9) . Type-locality, St. Francis River at Greenway

and Big Bay, Clay County, Arkansas.]

Cambarus (Faxonius) lancifer.—Ortmann, 1905c: 113.

Faxonius lancifer.—Penn, 1939:215.

Orconectes lancifer Hobbs, 1942a:352 [by implication].—

Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) lancifer.—Hobbs, 1942b: 154 [by implication].

TYPE.—Holotype, MCZ 306 (3 I).

TYPE-LOCALITY.—"Root Pond, Miss." Rocky

Ford near Etta, Union County, Mississippi, fide

Penn (1939:215).

RANGE.—Known from Texas, Louisiana, Arkansas, Mississippi, Tennessee, and extreme southern

Illinois.

HABITAT.—Sluggish streams and lentic situations.

Orconectes leptogonopodus Hobbs

FlCURE 131

Orconectes leptogonopodus Hobbs, 1948b: 146, figs. 24-32;

1972b:87, figs. 68a,e, 69b—Williams, 1954:854, figs. 18, 99-

106.—Fitzpatrick, 1963:61 [by implication].
NUMBER 166

Orconectes (Orconectes) leptogonopodus.—Hobbs, 1959:892.

Faxonius leptogonopodus leptogonopodus.—Creaser, 1962:2

[by implication].

Orconectes leptogonopodus leptogonopodus.—Fitzpatrick,

1965:87.

TYPES.—Holotype, allotype, and "morphotype,"

USNM 82262 (3 I, 9, $ II); paratypes, USNM,

MCZ, TU.

TYPE-LOCALITY.—McKinney's Creek, 4.7 miles

northeast of Hatfield, Polk County, Arkansas, on

U.S. Highway 71.

RANGE.—Red River drainage in Arkansas and

eastern Oklahoma.

HABITAT.—Small, clear, rocky streams.

Orconectes limosus. (Rafinesque)

FIGURE 103

Astacus limosus Rafinesque, 1817:42.

Astacus affinis Say, 1817:168. [Type unknown. Type-locality,

Delaware River.]

Astacus (Cambarus) affinis Erichson, 1846:96.

Cambarus Pealei Girard, 1852:87. [Syntypes, USNM 2081 (2 $ I,

2 $) . Type-locality, Potomac River at Washington, D.C.]

Cambarus affinis.—Girard, 1852:87.

Cambarus (Faxonius) limosus.—Ortmann, 1905c: 107.—Fowler, 1912:353, pis. 104, 105.

Cambarus limosus.—Ortmann, 1905c: 107.

Faxonius limosus.—Creaser, 1933a:3 [by implication]; 1962:2

[by implication].

Faxonius (Faxonius) limosus.—Creaser, 1933b:21 [by implication].

Orconectes limosus.—Hobbs, 1942a:352 [by implication];

1948a:20, fig. 22.—Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) limosus.—Hobbs, 1942b: 154 [by implication].

Faxonius affinis.—Hobbs, 1942c:55.

Orconectes limosa.—Mobberly, 1963:80 [erroneous spelling].

Camburus affinis.—Bacescu, 1967:218 [erroneous spelling].

Orconetes limosus.—Avault, 1973:242 [erroneous spelling].

TYPES.—Not known to exist.

TYPE-LOCALITY.—Delaware River near Philadelphia, Pennsylvania.

RANGE.—The Atlantic watershed from Maine to

the lower James River in Virginia. Introduced into

western Europe.

HABITAT.—Streams.

Orconectes longidigitus (Faxon)

FIGURE 142

Cambarus longidigitus Faxon, 1898:653, pi. 62: figs. 6-9.

Cambarus whitmani Steele, 1902:24, pi. 3: figs, cl, c2; pi. 5:

fig. B. [Location of types unknown. Type-locality, James

River, Missouri.]

Cambarus (Faxonius) longidigitus.—Ortmann, 1905c: 113.

Faxonius longidigitus.—Creaser, 1933a:3 [by implication];

1962:2 [by implication].

Faxonius (Faxonius) longidigitus.—Creaser, 1933b:21 [by implication].

Orconectes longidigitus.—Hobbs, 1942a:352 [by implication].—Williams, 1954:884, figs. 149, 183-190.

Orconectes (Orconectes) longidigitus.—Hobbs, 1942b: 154 [by

implication].

TYPES.—Syntypes, MCZ 4364 (2 <$ II, $ , 2 9 juv.),

USNM 19683 ( $ II, 3 $ , <J juv., 2 $ juv.).

TYPE-LOCALITY.—Oxford Bend, White River

(Izard County?), Arkansas.

RANGE.—Tributaries of the White and Little

Red rivers in Missouri and Arkansas.

HABITAT.—Streams.

Orconectes luteus (Creaser)

FIGURE 150

Faxonius (Faxonius) luteus Creaser, 1933b:7, pi. 1: figs. 3, 4

[subgenus by implication].

Faxonius luteus.—Creaser, 1934a: 1; 1962:2 [by implication].

Orconectes luteus.—Hobbs, 1942a:353^—Williams, 1954:872,

figs. 149-157.—Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) luteus.—Hobbs, 1942b: 154 [by implication].

Orconectes luetus.—Fitzpatrick, 1967a: 143 [erroneous spelling].

TYPES.—Holotype and allotype, USNM 98344,

98345 ($ I, $ ); paratypes, USNM.

TYPE-LOCALITY.—Niangua River at mouth of

Greasy Creek, 5 miles southeast of Buffalo, Dallas

County, Missouri.

RANGE.—From the Marais des Cygnes River,

Kansas, eastward through southern Missouri.

HABITAT.—Clear, swiftly flowing streams.

Orconectes macrus Williams

FIGURE 145

Orconectes nana macrus Williams, 1952:337, figs. 17-24; 1954:

851, figs. 82, 91-98.—Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) nana macrus.—Hobbs, 1959:892 [by

implication].

Faxonius nana macrus.—Creaser, 1962:2 [by implication].

Orconectes macrus.—Hobbs, 1972b:93, fig. 74e.

TYPES.—Holotype, allotype, and "morphotype,"
34

USNM 118847 (<jl, 9, all); paratypes, USNM,

MCZ.

TYPE-LOCALITY.—Spring River, 2 miles southwest of Mount Vernon, Lawrence County, Missouri.

RANGE.—Upper Arkansas River system in southwestern Missouri and northwestern Arkansas.

HABITAT.—Streams and in shallow burrows along

them.

Orconectes marchandi Hobbs

FIGURE 115

Orconectes marchandi Hobbs, 1948b: 140, figs. 1-4, 10-14,

17_Williams, 1954:843, figs. 1, 50-57.—Fitzpatrick, 1963:

61 [by implication].

Orconectes (Orconectes) marchandi.—Hobbs, 1959:895.

Faxonius marchandi.—Creaser, 1962:2 [by implication].

TYPES.—Holotype, allotype and "morphotype,"

USNM 82072, 82073, 82074 ($ I, 9 , $ II); paratypes, MCZ, USNM, TU.

TYPE-LOCALITY.—Stream 3.2 miles southeast of

Hardy, Sharp County, Arkansas, on U.S. Highway

63.

RANGE.—Spring River drainage system in Lawrence and Sharp counties, Arkansas, and Oregon

County, Missouri.

HABITAT.—Clear, rocky streams.

Orconectes medius (Faxon)

FIGURE 149

Cambarus medius Faxon, 1884:121; 1885a:107, pi. 3: fig. 4;

pi. 9: figs. 4, 4'.

Cambarus (Faxonius) medius.—Ortmann, 1905c: 112.

Faxonius medius.—Creaser, 1933a:3 [by implication]; 1962:2

[by implication].

Faxonius (Faxonius) medius.—Creaser, 1933b:21 [by implication].

Orconectes medius.—Hobbs, 1942a:352 [by implication].—

Williams, 1954:876, figs. 158-166._Fitzpatrick, 1963:61 [by

implication].

Orconectes (Orconectes) medius Hobbs, 1942b: 154 [by implication].

TYPES—Syntypes, MCZ 3585 (<$ I, 9).

TYPE-LOCALITY.—Irondale, Washington County,

Missouri.

RANGE.—Headwaters of the Big and Meramec

rivers in eastern Missouri.

HABITAT.—Streams.

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Orconectes meeki brevis Williams

FIGURE 156

Orconectes meeki brevis Williams, 1952:348, figs. 41-48; 1954:

881, figs. 166, 175-182.—Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) meeki brevis Hobbs, 1959:895 [by

implication].

Faxonius meeki brevis.—Creaser, 1962:2 [by implication].

TYPES.—Holotype, allotype, and "morphotype,"

USNM 118848 ($ I, $, $ II); paratypes, USNM,

MCZ.

TYPE-LOCALITY.—Stream, approximately 5 miles

north of Stilwell on U.S. Highway 59, Adair

County, Oklahoma.

RANGE.—Tributaries of the Arkansas River in

eastern Oklahoma and extreme northwestern

Arkansas.

HABITAT.—Rocky streams.

Orconectes meeki meeki (Faxon)

FIGURE 155

Cambarus meeki Faxon, 1898:657, pi. 65: figs. 5-9.

Cambarus meekii.—Hay, 1899b:960 [erroneous spelling].

Cambarus (Faxonius) meeki Ortmann, 1905c: 112.

G[ambarus] (Faxonius) meeki.—Ortmann, 1905c: 112 [erroneous spelling].

Faxonius meeki.—Creaser, 1933a:3 [by implication]

Faxonius (Faxonius) meeki Creaser, 1933b:21 [by implication].

Orconectes meeki.—Hobbs, 1942a:352 [by implication].

Orconectes (Orconectes) meeki meeki.—Hobbs, 1942b: 154 [by

implication].

Orconectes meeki meeki Williams, 1952:350; 1954:878, figs.

166-174.—Fitzpatrick, 1963:61 [by implication].

Faxonius meeki meeki.—Creaser, 1962:2 [by implication].

TYPES.—Syntypes, MCZ 4363 (2 3II, 3 9),

USNM 19680 (10 $ II,* 13 9), Mus. Zool. Torino

433.

TYPE-LOCALITY.—Walnut Fork of Big Piney

Creek, Swain, Newton County, Arkansas.

RANGE.—Arkansas, Red, and White river systems in Arkansas.

HABITAT.—Streams.

Orconectes menae (Creaser)

FlCURE 151

Faxonius (Faxonius) menae Creaser, 1933b:5, pi. 1: figs. 9, 10

[subgenus by implication].
NUMBER 166 35

Orconectes tnenae Hobbs, 1942a:353.—Williams, 1954:863,

figs. 123-131 Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) menae.—Hobbs, 1942b: 154 [by implication],

Faxonius menae.—Creaser, 1962:2 [by implication].

TYPES.—Holotype and allotype, USNM 98330,

98331 ($1, 9 ); paratypes, USNM, MCZ.

TYPE-LOCALITY.—Stream tributary to Irons Fork

of the Ouachita River, Mena, Polk County, Arkansas.

RANGE.—Tributaries of the Ouachita River in

Polk and Montgomery counties, Arkansas.

HABITAT.—Rocky creeks.

Orconectes minis (Ortmann)

FIGURE 143

Cambarus (Faxonius) rusticus mirus Ortmann, 1931:81.

Faxonius (Faxonius) mirus.—Creaser, 1933b: 10 [by implication].

Orconectes rusticus mirus.—Hobbs, 1942a:352 [by implication] Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) rusticus mirus.—Hobbs, 1942b: 154

[by implication].

Faxonius mirus.—Creaser, 1962:2 [by implication].

Orconectes mirus.—Hobbs, 1972b:97, figs. 74i, 76c,e.

TYPES.—Syntypes, CM 74.1378 ( $ I, 3 $ II, 9).

TYPE-LOCALITY.—Hurricane Creek, Cumberland

Springs, Moore County, Tennessee.

RANGE.—Elk and Duck river drainages in Alabama and Tennessee.

HABITAT.—Small creeks.

Orconectes mississippiensis (Faxon)

FIGURE 161

Cambarus mississippiensis Faxon, 1884:123; 1885a: 101, pi. 3:

fig. 2; pi. 10: figs. 4, 4', 4a, 4a'.

Cambarus (Faxonius) mississippiensis Ortmann, 1905c: 113.

Faxonius mississippiensis Creaser, 1933a:3 [by implication];

1962:2 [by implication].

Faxonius (Faxonius) mississippiensis.—Creaser, 1933b: 19 [by

implication].

Orconectes mississippiensis.—Hobbs, 1942a:352 [by implication].—Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) mississippiensis.—Hobbs, 1942b: 154

[by implication].

TYPES.—Syntypes, USNM 19760 ($ I, 9),

USNM 19759 ($ II, 9), MCZ 3563 (9 )•

TYPE-LOCALITY.—Eastern Mississippi. Restricted

to Macon, Noxubee County, Mississippi, by Faxon

(1914:422).

RANGE.—Eastern Mississippi.

HABITAT.—Streams and roadside ditches.

Orconectes nais (Faxon)

FIGURE 165

Cambarus Nais Faxon, 1885b: 140.

Cambarus nais.—Underwood, 1886:371.

Cambarus nais.—Faxon, 1890:631.

Cambarus pilosus Hay, 1899a: 121, fig. 1 [Types, USNM 19761

(6 3II), MCZ 7389 ($ II). Type-locality, Beloit, Mitchell

County, Kansas.]

Cambarus pelosus.—Harris, 1901:115 [erroneous spelling].

Cambarus (Faxonius) nais.—Ortmann, 1905c: 113.—Creaser

and Ortenberger, 1933:37, figs. 10, 14, 24.

Faxonius nais.—Creaser, 1933a:3; 1962:2 [by implication].

Faxonius (Faxonius) nais.—Creaser, 1933b:5.

Orconectes nais.—Hobbs, 1942a:352 [by implication].—Williams, 1954:886, figs. 191-199.—Fitzpatrick, 1963:61 [by

implication].

Orconectes (Orconectes) nais.—Hobbs, 1942b: 154 [by implication].

Orconectus nais.—Ray and Stevens, 1970:58 [erroneous spelling]-

TYPES.—Syntypes, MCZ 3755 (2*1, $11, 9),

USNM 129213 (3 <$ I, 3 & II); Mus. Washburn Coll.

(no longer there).

TYPE-LOCALITY.—Labette County, Kansas.

RANGE.—Great Plains and Ozark region.

HABITAT.—Streams.

REMARKS.—The syntypes in the United States

National Museum are specimens collected by W.

S. Newlon from the type-locality and were received

from the Kansas Museum of Natural History; presumably they represent a part of the lot that was

said by Faxon (1914) to have been at Washburn

College. See remarks under O. causeyi.

Orconectes nana Williams

FIGURE 144

Orconectes nana nana Williams, 1952:333, figs. 9-16; 1954:849,

figs. 82-90 Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) nana nana.—Hobbs, 1959:892 [by

implication],

Faxonius nana nana Creaser, 1962:2 [by implication].

Orconectes nana.—Hobbs, 1972b:96, figs. 74g, 76a.

TYPES.—Holotype, allotype, and "morphotype,"

USNM 129824, 129825, 129826 ($ I, 9, 6II);

paratypes, USNM, MCZ.

TYPE-LOCALITY.—Flint Creek, 5 miles east of

Kansas, Delaware County, Oklahoma.
36 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

RANGE.—Spring River drainage in Arkansas and

eastern Oklahoma.

HABITAT.—Swift, cold streams.

Kansas River in Colorado, Nebraska, and eastcentral Kansas.

HABITAT.—Rocky streams.

Orconectes neglectus chaenodactylus Williams

FIGURE 140

Orconectes neglectus chaenodactylus Williams, 1952:344, figs.

33-40; 1954:869, figs. 140-148 Fitzpatrick, 1963:61 [by

implication].

Orconectes (Orconectes) neglectus chaenodactylus—Hobbs,

1959:893 [by implication].

Faxonius neglectus chaenodactylus.—Creaser, 1962:2 [by implication].

TYPES.—Holotype, allotype, and "morphotype,"

USNM 118840 (<$L 9, ill); paratypes, USNM,

MCZ.

TYPE-LOCALITY.—Whites Creek, 8.6 miles southeast of Ava, Douglas County, Missouri.

RANGE.—North Fork of the White River in Arkansas and Missouri, intergrading in the headwaters with the nominate subspecies.

HABITAT.—Streams.

Orconectes neglectus neglectus (Faxon)

FIGURE 141

Cambarus neglectus Faxon, 1885b: 142.

Cambarus (Faxonius) neglectus.—Ortmann, 1905c 112.—

Creaser and Ortenburger, 1933:17, figs. 9, 15, 23.

Faxonius neglectus.—Creaser, 1933a:3 [by implication].

Faxonius (Faxonius) neglectus.—Creaser, 1933b:9 [by implication].

Cambarus neglecta.—Parks, Smith, and Garrett, 1939:2 [erroneous spelling].

Orconectes neglectus.—Hobbs, 1942a:352 [by implication].

Orconectes (Orconectes) neglectus Hobbs, 1942b: 154 by implication].

Orconectes neglectus neglectus.—Williams, 1952:344 [by implication]; 1954:866, figs. 132-139, 148—Fitzpatrick, 1963:

61 [by implication].

Faxonius neglectus neglectus.—Creaser, 1962:2 [by implication].

TYPES—Syntypes, MCZ 3757 ($ I, 9); MCZ

3590 (<JH) [questionable type]; Mus. Washburn

Coll. [no longer there].

TYPE-LOCALITY.—Mill Creek, Wabaunsee County, Kansas.

RANGE—Streams in the White (except North

Fork) and Arkansas river systems in Missouri, Oklahoma, and Arkansas, and in tributaries of the

Orconectes obscurus (Hagen)

FIGURE 117

Cambarus obscurus Hagen, 1870:69, pi. 1: figs. 72-75; pi. 3:

fig. 154.

Astacus obscurus Hagen, 1870:5 [lapsus].

Cambarus propinquus var. obscura.—Faxon, 1885a:92.

Cambarus propinquus obscurus.—Hay, 1899b:960.

Cambarus (Faxonius) obscurus.—Ortmann, 1905c:112; 1906b:

369, pi. A: figs. 1, 2; pi. 39: fig. 7; pi. 40: fig. 1; pi. 42:

figs. 2, 3.

Faxonius obscurus.—Creaser, 1933a:5; 1962:2 [by implication].

Faxonius (Faxonius) obscurus.—Creaser, 1933a:5 [by implication].

Orconectes obscurus Hobbs, 1942a:352 [by implication]

Crocker, 1957:36, 53, 75, pi. 3: figs. 5, 6; pi. 5: fig. 2 Fitzpatrick, 1963:61 [by implication]; 1967a: 131, 160, figs. 3,

11-15, 25.

Orconectes (Orconectes) obscurus.—Hobbs, 1942b: 154 [by implication].

Cambarus obscura.—Fitzpatrick, 1967a: 161 [erroneous spelling]-

TYPES.—Lectotype, MCZ 181 ($ I); paralectotypes, MCZ 181 (9 $ I, 6 2 ), MCZ 3353 (7*1, 9 ),

MCZ 3354 (3 ^ II), USNM 4971 (<51), MHNP

( S , 9 ), Wiirzburg Mus.; "cotypes" previously reported to be in the Australian Museum are not

there. The lectotype (g I) was selected by Fitzpatrick (1967a: 162) from MCZ 181, and all other

"types" were designated paralectotypes.

TYPE-LOCALITY.—Genesee River, Rochester,

Monroe County, New York.

RANGE.—Southeastern Ontario, New York, Pennsylvania, eastern Ohio, West Virginia, northern

Virginia, and western Maryland.

HABITAT.—Rocky streams.

Orconectes ozarkae Williams

FIGURE 153

Orconectes ozarkae Williams, 1952:339, figs. 25-32; 1954:860,

figs. 115-123 Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) ozarkae.—Hobbs, 1959:892.

Faxonius ozarkae.—Creaser, 1962:2 [by implication].

TYPES.—Holotype, allotype, USNM 118846 (a I,

9 ) and "morphotype," USNM 118844 (jH);

paratypes, USNM, MCZ.
NUMBER 166 37

TYPE-LOCALITY.—Hackney Creek, 4.6 miles westsouthwest of Stuart, Sharp County, Arkansas.

RANGE.—White River system in southern Missouri and northern Arkansas.

HABITAT.—Streams.

Orconectes palmeri creolanus (Creaser)

FIGURE 159

Faxonius (Faxonius) creolanus Creaser, 1933b:16, pi. 1: figs. 1,

2 [subgenus by implication].

Cambarus creolanus.—Penn, 1941:8.

Orconectes creolanus.—Hobbs, 1942a:352.

Orconectes (Orconectes) creolanus Hobbs, 1942b: 154 [by

implication].

Orconectes (Orconectes) palmeri creolanus.—Penn, 1952a:744.

Orconectes palmeri creolanus.—Penn, 1953e:6; 1957:231, figs.

24, 27, 30.—Fitzpatrick, 1963:61 [by implication].

Faxonius palmeri creolanus.—Creaser, 1962:2 [by implication].

TYPES.—Holotype and allotype, USNM 101972

and 98341 (4 I, $); paratypes, MCZ, USNM.

TYPE-LOCALITY.—Tributary to Amite River, 0.5

mile north of Ethel, East Feliciana Parish, Louisiana.

RANGE.—Mississippi and Louisiana in "the Lake

Pontchartrain drainage area plus the Pearl and

Pascagoula rivers" (Penn, 1957:258).

HABITAT.—Rapid and sluggish streams.

Orconectes palmeri longimanus (Faxon)

FIGURE 158

Cambarus palmeri longimanus Faxon, 1898:655, pi. 64: figs.

1-6.

Cambarus longimanus Creaser and Ortenburger, 1933:16,

38, figs. 4, 16, 26.

Faxonius longimanus Creaser, 1933a:3 [by implication].

Faxonius (Faxonius) longimanus.—Creaser, 1933b: 19 [by implication].

Orconectes palmeri longimanus.—Hobbs, 1942a:352 [by implication].—Williams, 1954:894, figs. 200, 209-216.—Penn,

1957:231, figs. 25, 28.—Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) palmeri longimanus.—Hobbs, 1942b:

154 [by implication].

Faxonius palmeri longimanus Creaser, 1962:2 [by implication].

TYPES.—Holotype, MCZ 7390 ( $ I); paratypes,

MCZ, USNM, Mus. Zool. Torino.

TYPE-LOCALITY.—Red River, Arthur City, Lamar County, Texas.

RANGE.—Texas, Oklahoma, Kansas, Arkansas,

and Louisiana, "all western tributaries of the Mississippi River from the Arkansas River to the

Gulf, and Gulf drainage streams from the Mississippi River to the Guadalupe River in Texas"

(Penn, 1957:258).

HABITAT Streams.

Orconectes palmeri palmeri (Faxon)

FIGURE 157

Cambarus palmeri Faxon, 1884:124; 1885a: 103, pi. 3: fig. 6;

pi. 10: figs. 5a, 5a'.

Cambarus palmeri palmeri.—Faxon, 1898:655 [by implication].

Cambarus (Faxonius) palmeri.—Ortmann, 1905c: 113.

Faxonius palmeri.—Creaser, 1933a:3 [by implication].

Faxonius (Faxonius) palmeri Creaser, 1933b: 19 [by implication].

Orconectes palmeri palmeri.—Hobbs, 1942a:352 [by implication].—Hobbs and Marchand, 1943:28, figs. 7, 13, 19, 26.—

Penn, 1957:231, figs. 26, 29—Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) palmeri.—Hobbs, 1942b: 154 [by implication].

Faxonius palmeri palmeri.—Creaser, 1962:2 [by implication].

TYPES.—Syntypes, USNM 4872 (8 3 II, 7$) ,

MCZ 3564 (2 <$ II, 2 9 ).

TYPE-LOCALITY.—Creek at eastern side of Reelfoot Lake, Obion County, Tennessee.

RANGE.—Lower Mississippi Valley, occurring in

western Tennessee and Mississippi and in eastern

Missouri and Arkansas.

HABITAT.—Streams.

Orconectes pellucidus (Tellkampf)

FIGURE 102

Astacus pellucidus Tellkampf, 1844a:684; 1844b:383; 1845:85.

Astacus (Cambarus) pellucidus—Erichson, 1846:95.

Cambarus pellucidus Girard, 1852:87.—Hagen, 1870:55, pi.

1: figs. 68-71; pi. 3: fig. 148; pi. 6.

Cambarus pelulcidus Hagen, 1870:106 [erroneous spelling].

Orconectes pellucidus.—Cope, 1872:409.—Hobbs and Barr,

1972:58, figs. 1, 14-16.

Cambarus typhlobius Joseph, 1880:202 [see Holthuis, 1964].

Cambarus coecus Joseph, 1881:237 [nomen midiim; see

Holthuis, 1964].

Cambarus stygius Joseph, 1881:241 [nomen nudum]; 1882:12

[validation of nomen nudum of 1881, becoming homonym

of Cambarus stygius 'Rxmdy—Procambarus acutus acutus

(Girard); see Holthuis, 1964].
38 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Cambarus (Orconectes) pellucidus Hovey, 1882:222 [in

part].

Cambarus Stygius Faxon, 1884:139.

Cambarus pellucidus pellucidus.—Packard, 1888:41 [by implication].

Camtarus (Orconectes) pellucidus, form inermis.—Packard,

1888:156 [erroneous spelling].

Astacus Cambarus Stigius.—Rabe, 1890:9 [erroneous spelling].

Cambrus pellucidus.—Price, 1900:155 [erroneous spelling].

Cambarus (Faxonius) pellucidus.—Ortmann, 1905c: 107.

Cambarus caecus Bouvier, 1940:68 [erroneous spelling].

Cambarus pelludicus.—Dearolf, 1942:50 [erroneous spelling].

Cambarus Pellucidus.—Jackson, 1942:4.

Orconectes pellucidus pellucidus.—Hobbs, 1942a:352 [by implication].—Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) pellucidus pellucidus Hobbs,

1942b: 154 [by implication].

Cambarus (Cambarus) pellucidus.—Balss, 1944:402.

Orconectes pellucidus pelluicdus.—Hobbs, 1948a:16 [erroneous spelling].

Cambarus Coecus.—Bott, 1950:25.

Gambarus typhlobius.—Croizat, 1958:908 [erroneous spelling].

TYPES.—Holotype, ZBM 1562 ($ I).

TYPE-LOCALITY.—Mammoth Cave, Edmonson

County, Kentucky.

RANGE.—Area extending "southwestward from

Hart County to Trigg County, Kentucky, and

Montgomery County, Tennessee" (Hobbs and Barr,

1972).

HABITAT.—Subterranean waters and springs.

Orconectes perfectus Walls

FIGURE 110

Orconectes perfectus Walls, 1972:451, figs. 1A-E, 4C.

TYPES.—Holotype, USNM 131256 (31); paratypes, USNM.

TYPE-LOCALITY.—Satilpa Creek, 8.7 miles northwest (not "east") of Grove Hill on Chilton Road,

Clarke County, Alabama.

RANGE.—Known from five localities in Clarke

and Choctaw counties, Alabama, and Clay and

Lowndes counties, Mississippi.

HABITAT.—Streams.

Orconectes peruncus (Creaser)

FIGURE 135

Cambarus (Faxonius) peruncus Creaser, 1931a:7, figs. 13-17.

Faxonius (Faxonius) peruncus.—Creaser, 1933b: 12 [by implication].

Faxonius peruncus Creaser, 1934a: 1; 1962:2 [by implication].

Orconectes peruncus Hobbs, 1942a:353.—Williams, 1954:

845, figs. 18, 66-73.—Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) peruncus.—Hobbs, 1942b: 154 [by

implication].

TYPES.—Holotype and allotype, USNM 98339,

98340 ($ I, $); paratypes, USNM.

TYPE-LOCALITY.—Little Creek, tributary to St.

Francis River, 1 mile northeast of Chloride, Iron

County, Missouri.

RANGE.—Headwaters of the St. Francis River in

central Missouri.

HABITAT.—Rocky streams.

Orconectes placidus (Hagen)

FIGURE 138

Cambarus placidus Hagen, 1870:65, pi. 1: figs. 76-79; pi. 3:

fig. 158.

Cambarus rusticus, var. placida Faxon, 1885a:65.

Cambarus rusticus, var. placidus.—Faxon, 1885b: 142.

Cambarus rusticus placidus.—Faxon, 1890:630.

Cambarus (Faxonius) rusticus placidus.—Ortmann, 1931:76.

Faxonius placidus.—Creaser, 1933a:3 [by implication]; 1962:2

[by implication].

Faxonius (Faxonius) placidus Creaser, 1933b: 10 [by implication].

Orconectes rusticus placidus.—Hobbs, 1942a:352 [by implication] Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) rusticus placidus Hobbs, 1942b: 154

[by implication].

Orconectes placidus.—Hobbs, 1966b:69; 1967a: 125.

TYPES.—Syntypes, MCZ 289 (4 ^11,3$ ), USNM

4966 (4 II).

TYPE-LOCALITY.—Lebanon, Wilson County, Tennessee.

RANGE.—Cumberland, Duck, and Tennessee

drainage systems in southern Kentucky, Tennessee,

and northern Alabama.

HABITAT.—Streams.

Orconectes propinquus (Girard)

FIGURE 118

Cambarus propinquus Girard, 1852:88.

Astacus propinquus.—Hagen, 1870:61.

Cambarus propinquus propinquus.—Faxon, 1885a:91 [by implication].

Camborus propinquus.—Williamson, 1899:47 [erroneous spelling]-

Cambarus propinquus typicus.—Ortmann, 1905c: 133.
NUMBER 166 39

Cambarus (Faxonius) propinquus.—Ortmann, 1905c: 112;

19O6b:358, pi. 39: fig. 6a,b.

Faxonius propinquus.—Williamson, 1907:754.—Creaser,

1933a: 1.

Faxonius (Faxonius) propinquus propinquus Creaser, 1933a:

3 [by implication].

Cambarus propinguus Fleming, 1939:304 [erroneous spelling]-

Cambarus propinkuus Fleming, 1939:305 [in part; erroneous

spelling].

Cambarrus propinquus Fleming, 1939:305 [erroneous spelling]-

Orconectes propinquus propinquus.—Hobbs, 1942a:352 [by

implication] Crocker, 1957:35, 48, 75; frontispiece; pi. 3:

figs. 1, 2; pi. 5: fig. 1 Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) propinquus propinquus.—Hobbs,

1942b: 154 [by implication].

Orconectes propinquus Hoff, 1943:281.—Fitzpatrick, 1967a:

143, figs. 2-19.

Orconnectes propinquus Threinen, 1958:1 [erroneous spelling]-

Faxonius propinquus propinquus.—Creaser, 1962:2 [by implication].

TYPES.—Destroyed in the Chicago fire in 1871.

TYPE-LOCALITY.—Three localities were originally given by Girard. From these, Faxon designated Garrison Creek, Sackett's Harbor, Jefferson

County, New York, as the type-locality. Ortmann

cited Oswego, Oswego County, New York. Crocker

(1957:35) and Fitzpatrick (1967a: 146) accepted

Ortmann's designation.

RANGE.—Ontario and New York west to Illinois

and Wisconsin. #

HABITAT.—Streams.

Orconectes punctimanus (Creaser)

FIGURE 133

Faxonius (Faxonius) punctimanus Creaser, 1933b: 1, pi. 1: figs.

5, 6 [subgenus by implication].

Faxonius punctimanus.—Creaser, 1934a: 1; 1962:2 [by implication].

Orconectes punctimanus Hobbs, 1942a:353 Williams,

1954:856, figs. 107-114, 200.—Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) punctimanus.—Hobbs, 1942b: 154 [by

implication].

TYPES.—Holotype and allotype, USNM 98342,

98343 (a I, 9 ); paratypes, USNM, MCZ.

TYPE-LOCALITY.—Rubidoux Creek at Waynesville, Pulaski County, Missouri.

RANGE.—Between the White, St. Francis, and

Missouri rivers in Arkansas and Missouri.

HABITAT.—Streams.

Orconectes putnami (Faxon)

FICURE 148

Cambarus Putnami Faxon, 1884:131; 1885a: 118, pi. 5: fig. 5;

pi. 9: figs. 6, 6', 6a, 6a'.

Cambarus putnami.—Underwood, 1886:372.

Cambarus (Faxonius) putnami.—Ortmann, 1905c: 112.

Cambarus (Faxonius) juvenilis Ortmann, 1931:84 [in part].

Orconectes putnami.—Rhoades, 1944a: 129.—Hobbs, 1972b:86,

figs. 68d, 69a.

TYPES.—Syntypes, MCZ 3568 (4 $ I, 7 9 , 8 $ juv.,

5$juv.), USNM 10130 (,jl).

TYPE-LOCALITY.—Bear Creek, Grayson Springs,

Grayson County, Kentucky.

RANGE.—Ohio drainage system in western Kentucky, southern Indiana (?), and Tennessee (?). The

limits of the range of O. putnami and the closely

related O. juvenilis and O. spinosus have not been

determined.

HABITAT.—Streams.

REMARKS.—Following Ortmann (1931) this

species has been largely relegated to synonymy with

O. juvenilis (Hagen). Even Rhoades, in using the

combination Orconectes putnami for the first time,

considered it a synonym of O. juvenilis. Not until

1972 was it proposed that it again be accorded

specific status.

Orconectes quadruncus (Creaser)

FIGURE 130

Faxonius (Faxonius) quadruncus Creaser, 1933b: 10, pi. 1: figs.

11, 12 [subgenus by implication].

Faxonius quadruncus—Creaser, 1934a: 1; 1962:2 [by implication].

Orconectes quadruncus Hobbs, 1942a:353.—Williams, 1954:

844, figs. 1, 58-65 Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) quadruncus.—Hobbs, 1942b: 154 [by

implication].

TYPES.—Holotype and allotype, USNM 98346,

98347 (£l, 9); paratypes, USNM, MCZ.

TYPE-LOCALITY.—Stout's Creek, tributary to St.

Francis River, between Ironton and Arcadia, Iron

County, Missouri.

RANGE.—Restricted to the headwaters of the St.

Francis River system in Iron, St. Genevieve, and

Madison counties, Missouri.

HABITAT.—Rapidly flowing, rocky streams.
40 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Orconectes rafinesquei Rhoades

FIGURE 124

Orconectes rafinesquei Rhoades, 1944a: 116, fig. 1A-F.—Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) rafinesquei.—Hobbs, 1959:893.

Faxonius rafinesquei.—Creaser, 1962:2 [by implication].

TYPES.—Holotype, allotype, and morphotype,

USNM 81304, 81306, 81305 ($1, $, «J II); paratypes, ANSP, MCZ, RR.

TYPE-LOCALITY.—Rough River, at Falls-ofRough, Grayson-Breckenridge counties, Kentucky.

RANGE.—Breckenridge, Grayson, and Ohio counties in central Kentucky.

HABITAT.—Streams.

Orconectes rusticus.—Spoor, 1955:78 Hobbs, 1972b:92, figs.

74c, 75b,d.

Orconnectes rusticus.—Threinen, 1958:3 [erroneous spelling].

Faxonius rusticus rusticus.—Creaser, 1962:2 [by implication].

TYPES.—"Types probably destroyed in the Chicago fire in 1871" (Faxon, 1914:418). A questionable type (in very poor condition) is USNM 4968

($ II); also questionable is a dry female, ANSP

195.

TYPE-LOCALITY.—Ohio River at Cincinnati,

Hamilton County, Ohio.

RANGE.—Michigan, Ohio, Indiana, Kentucky,

Tennessee, and southern Ontario; introduced in

Massachusetts.

HABITAT.—Streams, ponds, and lakes.

Orconectes rhoadesi Hobbs

FIGURE 167

Cambarus validus Faxonianus.—Fleming, 1938:303 [error for

"C. validus Faxon"].

Orconectes rhoadesi Hobbs, 1949a: 19, pi. 3: figs. 1-10.—Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) rhoadesi.—Hobbs, 1959:895.

Faxonius rhoadesi.—Creaser, 1962:2 [by implication].

TYPES.—Holotype, morphotype, USNM 87953

(Si, & II), and allotype, USNM 87954 ($ ); paratypes, USNM.

TYPE-LOCALITY.—Otter Creek, between Granny

White Pike and Hillsboro Pike, about 7 miles

south of Nashville, Davidson County, Tennessee.

RANGE.—Cumberland, Duck, and Tennessee

basins in central Tennessee.

HABITAT.—Streams.

Orconectes sanbornii erismophorous

Hobbs and Fitzpatrick

FICURE 122

Orconectes propinquus erismophorous Hobbs and Fitzpatrick,

1962:208, figs. 1-15.

Orconectes sanborni erismophorus.—Fitzpatrick, 1967a:131,

figs. 2-18, 24 [erroneous spelling].

Orconectes sanborni erismophorous.—Fitzpatrick, 1967a: 160.

Orconectes sanbornii erismophorous [original genitive of

species].

TYPES.—Holotype, allotype, and morphotype,

USNM 107597, 107598, 107599 («$ I, $, all);

rjferatypes, TU, USNM.

TYPE-LOCALITY.—Crane Nest Creek at Pee Wee,

Wirt County, West Virginia.

RANGE.—Little Kanawha River system, West

Virginia.

HABITAT.—Streams.

Orconectes rusticus (Cirard)

FIGURE 136

Cambarus rusticus Girard, 1852:88.—Faxon, 1885a: 108, pi. 9:

figs. 8, 8', 8a, 8a'.

Cambarus (Faxonius) rusticus.—Ortmann, 1905c: 112.

Faxonius rusticus.—Williamson, 1907:753 [Faxonius intended

as a subgeneric name].—Creaser, 1933a:5.

Cambarus (Faxonius) rusticus rusticus.—Ortmann, 1931:82.

Faxonius (Faxonius) ruslicus rusticus.—Creaser, 1933b:21 [by

implication].

Orconectes rusticus rusticus.—Hobbs, 1942a:352 [by implication].—Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) rusticus rusticus.—Hobbs, 1942b: 154

[by implication].

Orconectes sanbornii sanbornii (Faxon)

FIGURE 121

Cambarus Sanbornii Faxon, 1884:128.

Cambarus propinquus Sanbornii.—Faxon, 1885a:91, pi. 5: fig.

3; pi. 9: figs. 10, KK, 10a, 10a'.

Cambarus propinquus var. sanborni.—Underwood, 1886:372.

Cambarus propinquus var. sanbornii Osborn and Williamson, 1898:21.

Cambarus propinquus sanbornii Faxon, 1898:660.

Cambarus propinquus sanborni Ortmann, 1905c: 128.

Cambarus obscurus sanborni Ortmann, 1906b:437.

Faxonius sanborni—Creaser, 1933a:3 [by implication].

Faxonius (Faxonius) sanborni.—Creaser, 1933b:21 [by implication].
NUMBER 166 41

Orconectes propinquus sanborni.—Hobbs, 1942a:352 [by implication] Fitzpatrick, 1963:61 [by implication].

Orconectes {Orconectes) propinquus sanborni.—Hobbs,

1942b: 154 [by implication].

Faxonius sanborni sanborni Creaser, 1962:2 [by implication].

Orconectes sanborni sanborni.—Fitzpatrick, 1967a: 131, 157,

figs. 2-18, 23.

Orconectes sanborni Stevenson, 1967:208.

Orconectes sanbornii sanbornii [original genitive].

TYPES.—Lectotype, MCZ 3692 ( $ I); paralectotypes, MCZ 3587 (lost), MCZ 3692 (ill , 9) [see

Fitzpatrick (1967a: 157)].

TYPE-LOCALITY.—Oberlin, Lorain County, Ohio.

Restricted by Faxon (1914:418).

RANGE.—Southern Ohio and adjacent West Virginia and Kentucky.

HABITAT.—Streams.

Orconectes shoupi Hobbs

FICURE 129

Cambarus propinkuus.—Fleming, 1939:305 [in part; erroneous spelling].

Cambarus propinquus sanborni.—Fleming [not Faxon], 1939:

305 [in part], pi. 14.

Orconectes shoupi Hobbs, 1948a: 14, figs. 1-5, 9, 15, 16, 19,

20 Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) shoupi.—Hobbs, 1959:893.

Faxonius shoupi.—Creaser, 1962:2 [by implication].

TYPES.—Holotype, allotype, and morphotype,

USNM 84072 (<jl, 9, ill); paratypes, MCZ,

USNM.

TYPE-LOCALITY.—Mill Creek, tributary of Cumberland River, east of Oglesby near Antioch Pike,

10 miles south of Nashville, Davidson County,

Tennessee.

RANGE.—Known only from the vicinity of Nashville, Tennessee.

HABITAT.—Streams.

Orconectes sloanii (Bundy)

FIGURE 112

Cambarus sloanii Bundy, 1876:24.

Cambarus Sloanii.—Faxon, 1885a:89, pi. 4: fig. 5; pi. 10: figs.

1, V, la, la'.

Cambarus sloanei.—Ortmann, 1902:278 [erroneous spelling].

Cambarus (Faxonius) sloanei.—Ortmann, 1905c: 107 [erroneous spelling].

Faxonius sloanii.—Creaser, 1933a:3 [by implication]; 1962:2

[by implication].

Faxonius (Faxonius) sloanii.—Creaser, 1933b:21 [by implication].

Cambarus sloani.—Rhoades, 1941a:93.

Orconectes sloani.—Hobbs, 1942a:352 [by implication]; 1948a:

20.—Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) sloani.—Hobbs, 1942b: 154 [by implication]; 1959:894.

Orconectes sloanii.—Eberly, 1955:281.

Orconectes sloanei.—Eberly, 1958:2 [erroneous spelling].

TYPES.—Syntypes, MCZ 3806 («$ I, $).

TYPE-LOCALITY.—New Albany, Floyd County,

Indiana.

RANGE.—Southern Indiana and southwestern

Ohio.

HABITAT.—Streams.

Orconectes spinosus (Bundy)

FIGURE 146

Cambarus spinosus Bundy, 1877:173 Faxon, 1885a: 115, pi.

9: figs. 7, 7', 7a, 7a'.

Cambarus (Faxonius) spinosus.—Ortmann, 1905c: 112.

Cambarus (Faxonius) juvenilis.—Ortmann, 1931:84 [in part].

Orconectes spinosus.—Hobbs, 1944:340; 1972b:87, figs. 68b,g,

69d.

TYPES.—Syntypes, MCZ 3540 (2 $ II, 2 $ ), MCZ

3541 (<jll, 9), USNM 19779 (3 «j II, 2?).

TYPE-LOCALITY.—Etowah, Oostanaula, and

Coosa rivers, in the vicinity of Rome, Floyd County, Georgia.

RANGE.—Stream tributaries of the Coosa, Kanawha (?), and Tennessee rivers in Alabama, Georgia,

eastern Tennessee, Virginia (?), and West Virginia (?). The limits of the ranges of O. spinosus

and the closely related O. juixnilis and O. putnami have not been determined.

HABITAT.—Streams.

REMARKS.—Following Ortmann (1931) this

species has been largely relegated to synonomy

with O. juvenilis (Hagen). Not until 1972 was it

proposed that it again be accorded specific status.

Orconectes transfuga Fitzpatrick

FIGURE 152

Orconectes transfuga Fitzpatrick, 1966b: 178, figs. A-T.

Orconectes transfugus.—Fitzpatrick, 1967a: 143 [erroneous

spelling].

TYPES.—Holotype, allotype, USNM 114763 (<j I,
42 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

9 ), and morphotype, USNM 114762 ($ II); paratypes, MCZ, USNM, J. F. Fitzpatrick, Jr.

TYPE-LOCALITY.—Rogue River, 6 miles upstream

from Grants Pass, Jackson County, Oregon.

RANGE.—Only two localities known, both in

Rogue River drainage in Jackson County, Oregon.

HABITAT.—Streams.

Orconectes tricuspis Rhoades

FIGURE 126

Orconectes tricuspis Rhoades, 1944a:117, fig. 2A-F.—Hobbs,

1948a:20.—Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) tricuspis.—Hobbs, 1959:894.

Faxonius tricuspis.—Creaser, 1962:2 [by implication].

TYPES.—Holotype, allotype, and "morphotype"

(latter designated as allotype by Rhoades), USNM

81307, 81309, 81308 ($ I, 5, $\l); paratypes,

ANSP, MCZ, RR.

TYPE-LOCALITY.—Pete Lights' spring, 3 miles

east of Canton, Trigg County, Kentucky.

RANGE.—Cumberland drainage in Lyon, Trigg,

and Christian counties, Kentucky.

HABITAT.—Small streams.

Orconectes validus (Faxon)

FIGURE 168

Cambarus validus Faxon, 1914:382, pi. 7: figs. 3, 4, 8; pi. 13:

fig. 1.

Faxonius validus Creaser, 1933a:3 [by implication]; 1962:2

[by implication].

Faxonius (Faxonius) validus.—Creaser, 1933b:21 [by implication].

Cambarus validus Faxonianus.—Fleming, 1938:303 [not validus Faxon; see O. rhoadesi].

Orconectes validus Hobbs, 1942a:352 [by implication].—

Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) validus.—Hobbs, 1942b: 154 [by implication].

TYPE.—Holotype, MCZ 301 («jl).

TYPE-LOCALITY.—Huntsville, Madison County,

Alabama.

RANGE.—Tennessee drainage below Walden

Gorge in Alabama and southern Tennessee, and

upper Black Warrior and Tombigbee drainages in

Alabama.

HABITAT.—Streams.

Orconectes virginiensis Hobbs

FIGURE 120

Orconectes virginiensis Hobbs, 1951a: 122, figs. 1—10.—Fitzpatrick, 1963:61 [by implication]; 1967a: 131, 164, figs. 2-18,

26.

Orconectes (Orconectes) virginiensis.—Hobbs, 1959:893.

Faxonius virginiensis.—Creaser, 1962:2 [by implication].

Orconectes virginianus.—Fitzpatrick, 1967a: 133 [erroneous

spelling].

TYPES.—Holotype, allotype, and morphotype,

USNM 91659, 91660, 91661 ($ I, $, a II); paratypes, USNM.

TYPE-LOCALITY.—Rowanty Creek, tributary of

the Nottoway River, 3.3 miles south of Reams

Station on U.S. Highway 301, Dinwiddie County,

Virginia.

RANGE.—Chowan drainage system in North

Carolina and Virginia.

HABITAT.—Streams.

Orconectes virilis (Hagen)

FIGURE 162

Cambarus virilis Hagen, 1870:63, pi. 1: figs. 23-28; pi. 2: figs.

128-132; pi. 3: fig. 155; pi. 8.

Cambarus debllis Bundy, 1876:24. [Type, MCZ 3449 ($11).

Type-locality, Barraboo River, Ironton, Sauk County, Wisconsin.]

Cambarus wisconsinensis Bundy, 1876:4. [Type, MCZ 3448

($ II). Type-locality, Racine, Racine County, Wisconsin.]

Cambarus couesi Streets, 1877:803. [Syntypes, USNM 3154

(7 31, 3$juv., 5$juv.), MCZ 3545 ($1, $ , $ II). Typelocality, Red River of the North, near Pembina, Pembina

County, North Dakota.]

Cambarus viriles.—Bundy, 1882:180 [erroneous spelling].

Cambarus viridis Moenkhaus, 1904:17 [erroneous spelling].

Cambarus (Faxonius) virilis.—Ortmann, 1905c:113.—Creaser,

1932:326, figs. 1, 2, 8.

Faxonius virilis.—Creaser, 1933a:3; 1962:2 [by implication].

Faxonius (Faxonius) virilis.—Creaser, 1933b:21 [by implication].

Orconectes virilis.—Hobbs, 1942a:352 [by implication]; 1972b:

91, figs. 72h, 73e.—Fitzpatrick, 1963:61 [by implication].

Orconectes (Orconectes) virilis.—Hobbs, 1942b: 154 [by implication].

Orconnectes virilis.—Threinen, 1958:1 [erroneous spelling].

Cambarus virilus Hama, 1961:329 [erroneous spelling].

Camburus viridis.—Baccscu, 1967:218 [erroneous spelling].

Orconectes (Cambarus) virilis.—Mason, 197Ob:975.

TYPES.—Syntypes, MCZ 194 ( $ I, $11, 4 9 ,

2 9 ovig.), MCZ 196 (7 $ I, 3 $ II, 3 9, <5 juv.,

9juv.), MCZ 203 ($11, 39) , MCZ 1151 ($1),
NUMBER 166

MCZ 3342 (£ I, 5 ^ II, 9, 2<?juv., 9 juv.), MCZ

3343 (4 3 I, 9 ), MCZ 3344 ( $ II, 2 9 , $ juv.,

3 9 juv.), MHNP, Wiirzburg Mus., AMS.

TYPE-LOCALITY.—"Lake Superior," restricted by

Faxon (1914:420).

RANGE.—From Saskatchewan to Ontario, Canada, and from Montana and Wyoming to New

York and southwestern Maine. Introduced into

California (Riegel, 1959:29, 37), Maryland (Meredith and Schwartz, 1960:28), parts of New England, and Tennessee (R. W. Bouchard, personal

communication).

HABITAT.—Lakes and streams.

Orconectes ivilliamsi Fitzpatrick

FIGURE 154

Orconectes williamsi Fitzpatrick, 1966a: 145, figs. 1-10.

TYPES.—Holotype, allotype, and morphotype,

USNM 115520, 115521, 115522 ($1, 9, all);

paratypes, MCZ, OSM, J. F. Fitzpatrick, Jr.

TYPE-LOCALITY.—White River, 2.8 miles east of

Pettigrew, Madison County, Arkansas.

RANGE.—Three localities in headwaters of the

White River in Madison County, Arkansas.

HABITAT.—Pools in small streams.

Orconectes wrighti Hobbs

FIGURE 104

Orconectes wrighti Hobbs, 1948c:85, pi. 3: figs. 1-9.—Fitzpatrick, 1963:61 [by implication].

Orconectes {Orconectes) wrighti.—Hobbs, 1959:893.

Faxonius wrighti.—Creaser, 1962:2 [by implication].

TYPES.—Holotype, USNM 85144 ( $ I); allotype

and morphotype, USNM 85145 (9 , ill); paratypes, USNM.

TYPE-LOCALITY.—Robinson Creek on State

Route 57, Hardin County, Tennessee.

RANGE.—Known only from Hardin County, Tennessee.

HABITAT.—Streams.

Genus Procambarus Ortmann

Astacus Harlan, 1830:464.

Camborus Williamson, 1899:47 [erroneous spelling].

Cambarus.—Ortmann, 1905c:96. [Proposed as typical subgenus of Cambarus for which Astacus Blandingii Harlan,

1830:464, was designated type-species.]

Procambarus Ortmann, 1905d:437. [Type-species by subsequent designation (Fowler, 1912:340), Cambarus Digueti

Bouvier, 1897:225. Gender: masculine. Proposed as subgenus of Cambarus and elevated to generic rank by Hobbs,

1942a:341.]

Paracambarus Ortmann, 1906a: 1. [Type-species by monotypy,

Cambarus (Paracambarus) paradoxus Ortmann, 1906a:3.

Gender: masculine. Proposed as subgenus of Cambarus,

elevated to generic rank by Hobbs, 1942a:344, and reduced

to subgenus of Procambarus by Hobbs, 1972a: 10.]

Ortmannicus Fowler, 1912:341. [Replacement name for Ortmann's subgenus Cambarus. Utilized as generic name by

Rhoades, 1944a: 114.]

Cambarellus.—Creaser, 1933b:21 [lapsus calami].

procambarus.—Padgett, 1970:19 [lapsus calami].

Subgenus Acucauda Hobbs

Acucauda Hobbs, 1972a:5. [Type-species by original designation, Procambarus fitzpatricki Hobbs, 1971a:461. Gender:

feminine.]

Procambarus (Acucauda) fitzpatricki Hobbs

FIGURE 170

Cambarus (Paracambarus) harnedi Lyle, 1938:76 [nomen

nudum].

Procambarus fitzpatricki Hobbs, 1971a:46I, fig. 1.

Procambarus (Acucauda) fitzpatricki Hobbs, 1972a:5, figs.

1/, 2a, 4a-g.

TYPES.—Holotype, allotype, and morphotype,

USNM 131205, 131206, 131207 ($ I, 9, ill);

paratypes, USNM.

TYPE-LOCALITY.—Roadside ditch at junction of

State Route 67 and Woolmarket Road, just north

of D'lberville, Harrison County, Mississippi.

RANGE.—Southern Mississippi between the Wolf

and Pascagoula rivers.

HABITAT.—Burrows (primary or secondary burrower ).

Subgenus Austrocambarus Hobbs

Austrocambarus Hobbs, 1972a:5. [Type-species by original

designation, Procambarus vazquezae Villalobos, 1954b:328.

Gender: masculine.]

Procambarus (Austrocambarus) acanthophorus

Villalobos

FK.URK 180

Procambarus acanthophorus Villalobos, 1948:175, pi. 1: figs.

1-10; pi. 2: figs. 1-9.
44 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Procambarus (Austrocambarus) acanthophorus.—Hobbs,

1972a:6.

TYPES.—Holotype, allotype, and morphotype,

IBM 90847(035) ($1, 9, *II); paratypes, IBM,

USNM.

TYPE-LOCALITY.—Four km east of Tuxtepec,

Oaxaca, Mexico.

RANGE.—Atlantic drainage in southern Veracruz

and Oaxaca, Mexico.

HABITAT.—Streams and canals.

Procambarus (Austrocambarus) atkinsoni

(Ortmann)

FIGURE 183

Cambarus (Procambarus) atkinsoni Ortmann, 1913:414.

Procambarus atkinsoni.—Hobbs, 1942a:342 [by implication]—

Hobbs and Villalobos. 1964:346, figs. 40-50.

Procambarus Atkinsoni.—Creaser, 1962:7 [erroneous spelling].

Procambarus (Austrocambarus) atkinsoni.—Hobbs, 1972a:6.

TYPES.—Lectoholotype, lectoallotype, and lectomorphotype, CM 74.924 ( $ I, $ , <J II); paralectotypes, CM 74.924, 74.925. Designated by Hobbs

and Villalobos, 1964:347, 349, 350.

TYPE-LOCALITY.—Tributaries of Rio de los

Indios, Isla de Pinos, Cuba.

RANGE.—Isla de Pinos, Cuba.

HABITAT.—Streams (?).

Procambarus (Austrocambarus) aztecus (Saussure)

FIGURE 172

Cambarus aztecus Saussure, 1857b:503; 1858:460, figs. 23, 23a,

23b.

Cambarus mexicanus.—[not Erichson.] Most authors from

Faxon, 1884, to present [in part].

Procambarus aztecus.—Hobbs, 1942a:342 [by implication].

Procambarus (Austrocambarus) aztecus.—Hobbs, 1972a:6.

TYPES.—Syntypes, USNM 20682 ( $ I), GM (uncataloged, dry).

TYPE-LOCALITY.—"Pris a Tomatlan, Dans les

Terres Chaudes." (Veracruz, Mexico.)

RANGE.—Vicinity of Jalapa, Veracruz, Mexico.

HABITAT.—Streams.

REMARKS.—Specimens described as Procambarus

aztecusby Villalobos (1954b:314) are not members

of this species and must receive a new name.

Procambarus (Austrocambarus) cubensis cubensis

(Erichson)

FIGURE 185

Astacus (Cambarus) Cubensis Erichson, 1846:100.

Cambarus Cubensis.—Girard, 1852:87 [by implication].—

Faxon, 1885a:51, pi. 2: fig. 1; pi. 8: figs. 5, 5', 5a, 5a'.

Cambarus consobrinus Saussure, 1857a: 101. [Syntypes, GM

(uncataloged, dry, 2$), specimens formerly in MHNP no

longer there, BMNH 1163 (2$), USNM 20684 (,$11).

Type-locality, ponds in the central part of Cuba.]

Cambarus (Cambarus) cubensis.—Ortmann, 1905c: 101.

Cambarus (Procambarus) cubensis.—Ortmann, 1905d:438.

Cambarus cubensis cubensis.—Faxon, 1912:458 [by implication].

Cambarus cubensis consobrinus.—Faxon, 1912:458.

Cambarellus cubensis.—Creaser, 1933b:21 [lapsus calami].

Procambarus cubensis cubensis Hobbs, 1942a:342 [by implication] Hobbs and Villalobos, 1964:319, figs. 1-13.

Procambarus cubensis consobrinus Hobbs, 1942a:342 [by

implication],

Procambarus consobrinus.—Villalobos, 1954b:303; 1955:160.

Procambarus (Austrocambarus) cubensis cubensis Hobbs,

1972a:6.

TYPES.—BMNH 4140.

TYPE-LOCALITY.—Cuba.

RANGE.—Scattered localities throughout Cuba,

except in the southwestern mountains.

HABITAT.—Lentic and lotic situations.

Procambarus (Austrocambarus) cubensis rivalis

(Faxon)

FIGURE 184

Cambarus cubensis rivalis Faxon, 1912:459.

Procambarus cubensis rivalis.—Hobbs, 1942a:342 [by implication].—Hobbs and Villalobos, 1964:335, figs. 14-24.

Procambarus (Austrocambarus) cubensis rivalis Hobbs,

1972a:6.

TYPES.—Syntypes, MCZ 7406 (2 $11, 3 5 ).

TYPE-LOCALITY.—San Diego de los Banos, Province of Pinar del Rio, Cuba.

RANGE.—Known only from a few localities in

the province of Pinar del Rio, Cuba.

HABITAT.—Mountain streams.

Procambarus (Austrocambarus) llamasi Villalobos

FIGURE 181

Procambarus llamasi Villalobos, 1954b:364, pis. 16, 17.

Procambarus (Austrocambarus) llamasi.—Hobbs, 1972a:6.
NUMBER 166 45

TYPES.—Holotype, allotype, and morphotype,

IBM (a I, 9, 8II); paratypes, IBM, USNM.

TYPE-LOCALITY.—Santa Rita, 47 km east of

Escarcega, Campeche, Mexico.

RANGE.—Veracruz to Yucatan, Mexico, and

northern Guatemala.

HABITAT.—Small streams and drainage ditches.

Procambarus (Austrocambarus) mexicanus

(Erichson)

FIGURE 173

Astacus (Cambarus) mexicanus Erichson, 1846:99.

Astacus mexicanus.—Dana, 1852:522.

Cambarus mexicanus.—Saussure, 1858:461.

Cambarus (Cambarus) mexicanus.—Ortmann, 1905c: 101.

Cambarus (Procambarus) mexicanus.—Ortmann, 1905d:438.

Procambarus mexicanus.—Hobbs, 1942a:342 [by implication].—Villalobos, 1954b:307, pis. 1, 2.

Procambarus Mexicanus.—Creaser, 1962:7 [erroneous spelling].

Procambarus (Austrocambarus) mexicanus.—Hobbs, 1972a:6.

TYPES.—Lost, formerly in ZBM. Neoholotype,

ANSP 421 (<J I); neoallotype and neoparatypes,

IBM (Villalobos, 1954b:312).

TYPE-LOCALITY.—Mexico. Restricted by Villalobos (1954b:312) to "El Mirador de Zacuapan," 8

km northeast of Huatusco, Veracruz.

RANGE.—Known from a single locality.

HABITAT.—Not reported.

Procambarus (Austrocambarus) mirandai

Villalobos

FIGURE 176

Procambarus mirandai Villalobos, 1954b:355, pis. 13-15.

Procambarus mirandae.—Hobbs and Villalobos, 1964:313

[erroneous spelling].

Procambarus (Austrocambarus) mirandai Hobbs, 1972a:6.

TYPES.—Holotype, allotype, and morphotype,

IBM (a I, 9, $11); paratypes, IBM, USNM.

TYPE-LOCALITY.—Cerro Hueco, 4 km southeast

of Tuxtla Gutierrez, Chiapas, Mexico.

RANGE.—Known only from the type-locality.

HABITAT.—"Los ejemplares fueron colectados

dentro de una gruta, que es la salida de un rio

subterrdneo."

Procambarus (Austrocambarus) niveus

Hobbs and Villalobos

FIGURE 186

Procambarus niveus Hobbs and Villalobos, 1964:342, figs.

26-39.

Procambarus (Austrocambarus) niveus.—Hobbs, 1972a:6.

TYPES.—Holotype and allotype, USNM 109076,

109077 (al, 9 ).

TYPE-LOCALITY.—"Cuevas de Santo Tomas,

Sierra de los Organos, cerca de Ponce, Pinar del

Rio, Cuba."

RANGE.—Known only from the type-locality.

HABITAT.—Subterranean waters.

Procambarus (Austrocambarus) pilosimanus

(Ortmann)

FIGURE 182

Cambarus (Procambarus) pilosimanus Ortmann, 1906a:6,

fig. 2.

Procambarus pilosimanus.—Hobbs, 1942a:342 [by implication] Villalobos, 1954b:371, pis. 18, 19.

Procambarus pelosimanus.—Creaser, 1962:7 [erroneous spelling].

Procambarus (Austrocambarus) pilosimanus.—Hobbs, 1972a:6.

TYPES.—Syntypes, MHNP (8 <J , 6 9 ), CM 74.678.

TYPE-LOCALITY.—"Coche, pres de la rivere de

Coban, Guatemala." Probably Rio Cahab6n at

Coban, Province of Alta Verapaz, Guatemala.

RANGE.—Guatemala, British Honduras, and

Chiapas and Quintana Roo, Mexico.

HABITAT.—Lentic habitats.

Procambarus (Austrocambarus) rodriguezi Hobbs

FIGURE 174

Procambarus rodriguezi Hobbs, 1943b:203, pi. 2.

Procambarus (Austrocambarus) rodriguezi.—Hobbs, 1972a:6.

TYPES.—Holotype, allotype, and "morphotype,"

USNM 81302 (a I, 9, all); paratypes, USNM,

IBM, MCZ.

TYPE-LOCALITY.—Cueva de Ojo de Agua, westnorthwest of Hacienda Potrero Viejo, Paraje

Nuevo, Cordoba, Veracruz, Mexico.

RANGE.—Known only from the type-locality.

HABITAT.—Subterranean stream.
46 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Procambarus (Austrocambarus) ruthveni ruthveni

(Pearse)

FIGURE 177

Cambarus ruthveni Pearse, 1911:110, fig. 3.

Procambarus ruthveni.—Hobbs, 1942a:342 [by implication].

Procambarus ruthveni ruthveni.—Villalobos, 1954b:336, pis.

9, 10.

Procambarus (Austrocambarus) ruthveni ruthveni.—Hobbs,

1972a: 6.

TYPES.—Holotype and allotype, USNM 98348,

9834 9 (si, 9) .

TYPE-LOCALITY.—Cuatotolapan, Cant6n de Acayucan, Veracruz, Mexico.

RANGE.—Known from only one additional locality: "San Juan Diaz Covarrubias, 10 Km. SO.

de Cuatotolapan, Ver." (Villalobos, 1954b: 344).

HABITAT.—Temporary ponds, pools, and burrows (secondary burrower).

RANGE.—Known only from the type-locality.

HABITAT.—Under rocks along the shore of Laguna de Catemaco.

Procambarus (Austrocambarus) veracruzanus

Villalobos

FIGURE 175

Procambarus veracruzanus Villalobos, 1954b:323, pis. 5, 6.

Procambarus (Austrocambarus) veracruzanus Hobbs, 1972a:

6.

TYPES.—Holotype, allotype, and morphotype,

IBM (si, 9 , S II); paratypes, IBM.

TYPE-LOCALITY.—Presidio, 30 km southeast of

Cordoba, Veracruz, Mexico. "Cuenca del rio Papaloapan."

RANGE.—Known only from the type-locality.

HABITAT.—Stream.

Procambarus (Austrocambarus) ruthveni

zapoapensis Villalobos

FIGURE 178

Procambarus ruthveni zapoapensis Villalobos, 1954b:347, pis.

11, 12.

Procambarus (Austrocambarus) ruthveni zapoapensis.—

Hobbs, 1972a:6.

TYPES.—Holotype, allotype, and morphotype,

IBM (si, $, 3II); paratypes, IBM, USNM.

TYPE-LOCALITY.—Zapoapan de Cabana, 11 km

southeast of Catemaco, Veracruz, Mexico.

RANGE.—Known only from the vicinity of the

type-locality.

HABITAT.—Small, rocky streams.

Procambarus (Austrocambarus) vazquezae

Villalobos

FIGURE 171

Procambarus vazquezae Villalobos, 1954b:328, pis. 7, 8.

Procambarus vasquezae.—Hobbs, 1966b:71 [erroneous spelling]-

Procambarus (Austrocambarus) vazquezae Hobbs, 1972a:6,

figs. 2b, 5a-g.

TYPES.—Holotype, allotype, and morphotype,

IBM (si, ?, ^11); paratypes, IBM, USNM.

TYPE-LOCALITY.—"Playa Norte de la Laguna de

Catemaco," Veracruz, Mexico.

Procambarus (Austrocambarus) williamsoni

(Ortmann)

FIGURE 179

Cambarus (Procambarus) williamsoni Ortmann, 1905d:439,

figs. 1-3.

Procambarus williamsoni.—Hobbs, 1942a:342 [by implication].

Procambarus (Austrocambarus) williamsoni.—Hobbs, 1972a:6.

TYPES.—Syntypes, CM 74.560 ($1, 9, $ II),

CM 74.561 (si, 9, ill).

TYPE-LOCALITY.—Los Amates, Province of Izabal,

four to five miles south of Rio Motagua, Guatemala. (Not "Malagua" as recorded by Ortmann.)

RANGE.—Known only from the vicinity of the

type-locality and from Lago Yojoa, Honduras.

HABITAT.—Unknown.

Subgenus Capillicambarus Hobbs

Capillicambarus Hobbs, 1972a:6. [Type-species by original

designation, Cambarus (Cambarus) hinei Ortmann, 1905b:

401. Gender: masculine].

Procambarus (Capillicambarus) hinei (Ortmann)

FIGURE 187

Cambarus (Cambarus) hinei Ortmann, 1905b:401, fig. 1.

Procambarus (Ortmannicus) hinei.—Fowler, 1912:341 [by implication].—Creaser, 1934b:4 [by implication].
NUMBER 166 47

Procambarus hinei.—Hobbs, 1942a:342 [by implication].—

Penn, 1953b:63, figs. 1-15.

Procambarus (Capillicambarus) hinei Hobbs, 1972a:6, figs.

In, 2c, 6a-g.

TYPES.—Syntypes, OSM 443.1 ( $ I), 443.2 (9 ),

444.1 (9), and 444.2 (9), USNM 98355 ( $ I, 9).

TYPE-LOCALITY.—One-fourth mile from Gulf

Beach, near Cameron, Cameron Parish, Louisiana.

RANGE.—Southern Louisiana and southeastern

Texas.

HABITAT.—Pools and roadside ditches.

Procambarus (Capillicambarus) incilis Penn

FIGURE 188

Procambarus incilis Penn, 1962:222, figs. 1-11.

Procambarus (Capillicambarus) incilis.—Hobbs, 1972a:6.

TYPES.—Holotype, allotype, and morphotype,

USNM 106794, 106795, 106796 ($ I, 9, $ II);

paratypes, USNM, TU.

TYPE-LOCALITY.—7.5 miles east of Edna, Jackson County, Texas.

RANGE.—Southeastern Texas in Jackson, Matagorda, and Wharton counties.

HABITAT.—Roadside ditches.

Subgenus Girardiella Lyle

Girardiella Lyle, 1938:76. [Type-species by monotypy, Cambarus Hagenianus Faxon, 1884:141. Gender: feminine.]

Procambarus (Girardiella) gracilis (Bundy)

FIGURE 193

Cambarus gracilis Bundy, 1876:5.

Cambarus gracillis Bundy, 1882:182 [erroneous spelling].

Cambarus (Cambarus) gracilis.—Ortmann, 1905c: 101.

Cambarus (Ortmannicus) gracilis Fowler, 1912:341 [by implication].

Cambarus (Girardiella) gracilis.—Lyle, 1938:76 [by implication].

Procambarus gracilis.—Hobbs, 1942a:342 [by implication].—

Williams, 1954:826, figs. 10-18.

Procambarus (Girardiella) gracilis Hobbs, 1972a:7, fig. Ik.

TYPES.—Syntypes, 111. St. Lab. Nat. Hist, (lost),

MCZ 3794 ( $ I, 9 ), MCZ 3454 (<$ I).

TYPE-LOCALITY.—Normal, McLean County, Illinois. Here restricted.

RANGE.—Texas, Oklahoma, Kansas, Arkansas,

Missouri, Iowa, Illinois, and Wisconsin.

HABITAT.—Burrows (primary burrower).

Procambarus (Girardiella) hagenianus (Faxon)

FIGURE 192

Cambarus advena Hagen, 1870:86-87 [in part], figs. 90-92,

164. [The description is of hagenianus. The Georgia specimens mentioned are P. pygmaeus and P. advena.]

Cambarus Carolinus Hagen [not Erichson], 1870:31.

Cambarus Hagenianus Faxon, 1884:141.

Cambarus hagenianus.—Hay, 1902c:38.—Faxon, 1914:366, pi.

1; pi. 7: figs, la-c, 7.

Cambarus (Cambarus) hagenianus Ortmann, 1905c:101.

Cambarus (Ortmannicus) hagenianus.—Fowler, 1912:341 [by

implication],

Cambarus (Girardiella) hagenianus hagenianus.—Lyle, 1938:

76.

Procambarus hagenianus Hobbs, 1942a:342 [by implication].

Procambarus (Girardiella) hagenianus.—Hobbs, 1972a:7, figs.

2d, 7a-g.

TYPES.—Holotype, MCZ 232 ($ I).

TYPE-LOCALITY.—Given as Charleston, South

Carolina, but definitely based on mislabeled specimens.

RANGE.—Central and eastern Mississippi and

western Alabama.

HABITAT.—Burrows (primary burrower).

Procambarus (Girardiella) simulans

regiomontanus Villalobos

FlCURE 191

Procambarus simulans regiomontanus Villalobos, 1954a:289,

pis. 1, 2.

Procambarus (Girardiella) simulans regiomontanus.—Hobbs,

1972a: 7.

TYPES.—Holotype, allotype, and morphotype,

IBM (i I, $ , $ II); paratypes, IBM, USNM.

TYPE-LOCALITY.—Five km north of Monterrey,

Nuevo Leon, Mexico.

RANGE.—Known only from the type-locality.

HABITAT.—Not recorded.

Procambarus (Girardiella) simulans simulans

(Faxon)

FIGURE 190

Cambarus simulans Faxon, 1884:112.

Cambarus similans.—Ortmann, 1892:12 [erroneous spelling].

Cambarus gallinus Cockerell and Porter, 1900:434, fig. 1.

[Syntypes, MCZ 7342 (3£11), USNM 23915 ($11. 2 9) .

23916 (31, 9) . 129197 (2&U, 29) , ANSP 1456 (,$1),
48 SMITHSON'IAN CONTRIBUTIONS TO ZOOLOGY

1455 (2 9 )• Type-locality, Gallinas River at Las Vegas, New

Mexico.]

Cambarus baumgarlneri Harris, 1901b: 115 [nomen nudum].

Cambarus (Cambarus) simulans.—Ortmann, 1905c: 101.

Cambarus (Ortmannicus) simulans.—Fowler, 1912:341 [by

implication].

Procambarus simulans.—Hobbs, 1942a:342 [by implication].

Procambarus simulans simulans.—Villalobos, 1954a:296.

Procambarus (Girardiella) simulans simulans.—Hobbs,

1972a:7.

TYPES.—Syntypes, MCZ 3646 ($ I, 4$), MCZ

3647 (23 $ II, 30 9 , 4 3 juv., 5 9 juv.), USNM 4150

(a I, 3«jII, 5 a juv.), ZIAS 1/44975 (2 S , 9).

TYPE-LOCALITY.—Dallas, Texas.

RANGE.—New Mexico, Texas, Louisiana, Oklahoma, Kansas, Arkansas, and Colorado.

HABITAT.—Lentic and lotic situations, and burrows (secondary burrower).

REMARKS.—The 3 <j II syntypes, MCZ 7342,

could not be located.

Procambarus (Girardiella) tulanei Penn

FIGURE 189

Procambarus tulanei Penn, 1953a: 163, figs. 1-12.

Procambarus (Girardiella) tulanei Hobbs, 1972a:7.

TYPES.—Holotype, allotype, and morphotype,

USNM 93655, 93656, 79928 ($ I, 9, 3 II); paratypes, USNM, TU, La. Polytechnic Inst.

TYPE-LOCALITY.—Tributary of Bayou D'Arbonne, 4 miles west of Dubach on State Route 288,

Lincoln Parish, Louisiana.

RANGE.—Ouachita and Red drainage systems in

Louisiana and Arkansas.

HABITAT.—Lentic and lotic situations, and burrows (secondary burrower).

Subgenus Hagenides Hobbs

Hagenides Hobbs, 1972a:7. [Type-species by original designation, Astacus advena LeConte, 1856:402. Gender: masculine.]

Procambarus (Hagenides) advena (LeConte)

FIGURE 194

Astacus advena LeConte, 1856:402.

Cambarus advena.—Hagen, 1870:87, pi. 3: fig. 164; pi. 7.

Faxon, 1884:140.

Cambarus Carolinus.—Hagen, 1870:87-88 [latter in part], pi.

I: figs. 51-54; pi. 3: fig. 165.

Cambarus (Cambarus) advena.—Ortmann, 1905c: 101.

Cambarus (Ortmannicus) advena.—Fowler, 1912:341 [by implication].

Procambarus advena.—Hobbs, 1942a:34O.

Procambarus (Hagenides) advena.—Hobbs, 1972a:7.

TYPES.—Syntypes, MCZ 3379 (9), ANSP 321

TYPE-LOCALITY.—"Georgia inferiore." Here restricted to 2.5 miles west of Riceboro, Liberty

County, Georgia.

RANGE.—Lower coastal plain of Georgia, between Savannah and Altamaha rivers.

HABITAT.—Burrows (primary burrower).

REMARKS.—As pointed out by Faxon (1884:140),

the descriptions of P. advena and C. carolinus were

transposed in Hagen's monograph, and figures 51-

54 and 165 are actually of P. advena.

Procambarus (Hagenides) geodytes Hobbs

FIGURE 195

Cambarus advena geodytes Hobbs, 1937:154 [nomen nudum].

Procambarus geodytes Hobbs, 1942b:80, figs. 61-65, 276-285.

Procambarus (Hagenides) geodytes Hobbs, 1972a:7.

TYPES.—Holotype and allotype, USNM 81279

(31, 9); "morphotype," USNM 81280 (3II);

paratypes, MCZ, USNM, UF (now in USNM).

TYPE-LOCALITY.—Orange Springs in the northeastern part of Marion County, Florida.

RANGE.—Along the St. Johns River and its tributaries from Putnam County south to Orange

County, Florida.

HABITAT.—Burrows in flatwoods and seepage

areas (primary burrower).

Procambarus (Hagenides) pygmaeus Hobbs

FIGURE 196

Procambarus pygmaeus Hobbs, 1942b:83, figs. 66-70, 296-304.

Procambarus (Hagenides) pygmaeus.—Hobbs, 1972a:7.

TYPES.—Holotype, allotype, and "morphotype,"

USNM 81285 (3 I, 9, all); paratypes, MCZ,

USNM.

TYPE-LOCALITY.—About 16 miles north of Fargo

on State Route 89, Clinch County, Georgia.

RANGE.—Georgia south of the Savannah River

and in Gulf, Liberty, Leon, and Wakulla counties,

Florida.
NUMBER 166 49

HABITAT.—Lentic and lotic situations and burrows (secondary burrower).

Procambarus (Hagenides) rogersi campestris

Hobbs

FIGURE 198

Procambarus rogersi campestris Hobbs, 1942b:90, figs. 81-85;

1945a:255, figs. 3, 4, 7, 16, 18, 21, 24.

Procambarus (Hagenides) rogersi campestris.—Hobbs, 1972a:7.

TYPES.—Holotype, allotype, and "morphotype,"

USNM 79384 ($ I, $, $ II); paratypes, MCZ,

USNM.

TYPE-LOCALITY.—About 12 miles south of Tallahassee, Leon County, Florida (Sec. 16-17, T. 2 S.,

R. 2 W.).

RANGE.—Leon and Wakulla counties, Florida.

HABITAT.—Burrows in flatwoods and seepage

areas (primary burrower).

Procambarus (Hagenides) rogersi expletus

Hobbs and Hart

FIGURE 199

Procambarus rogersi expletus Hobbs and Hart, 1959:179, figs.

27-39.

Procambarus (Hagenides) rogersi expletus.—Hobbs, 1972a:7.

TYPES.—Holotype, allotype, and morphotype,

USNM 102125, 102127, 102126 (£ I, $, ,5II);

paratypes, USNM.

TYPE-LOCALITY.—Seepage slope, 0.9 mile north

of Clarksville, Calhoun County, Florida, on State

Route 73.

RANGE.—Known only from the type-locality.

HABITAT.—Burrows in flatwoods and seepage

areas (primary burrower).

Procambarus (Hagenides) rogersi ochlocknensis

Hobbs

FIGURE 197

Procambarus rogersi ochlocknensis Hobbs, 1942b:89, figs. 76-

80; 1945a:252, figs. 1, 10, 11, 13, 15, 19, 20, 26.

Procambarus (Hagenides) rogersi ochlocknensis.—Hobbs,

1972a:7.

TYPES.—Holotype, allotype, and "morphotype,"

USNM 79383 ($ I, $, $ II); paratypes, MCZ,

USNM.

TYPE-LOCALITY.—7.3 miles west of Quincy, on

U.S. Highway 90, Gadsden County, Florida.

RANGE.—Gadsden and Liberty counties, Florida.

HABITAT.—Burrows in seepage areas (primary

burrower).

Procambarus (Hagenides) rogersi rogersi

(Hobbs)

FICURE 200

Cambarus rogersi Hobbs, 1938a:62, figs. 1-11.

Procambarus rogersi.—Hobbs, 1942a:344.

Procambarus rogersi rogersi.—Hobbs, 1942b:89, figs. 71-75.

Procambarus (Hagenides) rogersi rogersi.—Hobbs, 1972a:7.

TYPES.—Holotype, allotype, and "morphotype,"

USNM 75120, 75121, 75589 («jl, $, 4 II); paratypes, MCZ, USNM.

TYPE-LOCALITY.—Flatwoods, 4 miles north of

Blountstown on State Route 71, Calhoun County,

Florida.

RANGE.—Eastern Calhoun County, Florida.

HABITAT.—Burrows in flatwoods and seepage

areas (primary burrower).

Procambarus (Hagenides) truculentus Hobbs

FIGURE 201

Procambarus truculentus Hobbs, 1954:111, figs. 1-13.

Procambarus Truculentus.—Hobbs, 1954:111.

Procambarus (Hagenides) truculentus.—Hobbs, 1972a:7.

TYPES.—Holotype, allotype, and morphotype,

USNM 95670, 95671, 95672 ($ I, $, $ II); paratypes, MCZ, TU, USNM.

TYPE-LOCALITY.—Seepage area 11 miles north of

Lyons, Emanuel County, Georgia.

RANGE.—Between the Altamaha and Savannah

rivers in Georgia.

HABITAT.—Burrows in flatwoods and seepage

areas (primary burrower).

Subgenus Leconticambarus Hobbs

Leconticambarus Hobbs, 1972a:7. [Type-species by original

designation, Cambarus barbatus Faxon, 1890:621. Gender:

masculine.]

Procambarus (Leconticambarus) alleni (Faxon)

FIGURE 202

Cambarus Alleni Faxon, 1884:110; 1885a:35, pi. 1: fig. 1; pi.

8: figs. 2, 2'.
50 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Cambarus alleni.—Faxon, 1890:619.

Cambarus (Cambarus) alleni.—Ortmann, 1905c: 102.

Cambarus (Ortmannicus) alleni.—Fowler, 1912:341 [by implication].—Creaser, 1934b:4 [by implication].

Procambarus alleni.—Hobbs, 1942a:342 [by implication];

1942b:69, figs. 51-55.

Procambarus (Leconticambarus) alleni.—Hobbs, 1972a:8.

TYPE.—Holotype, MCZ 3531 (a I).

TYPE-LOCALITY.—St. Johns River, Hawkinsville

(1 mile upstream from bridge at Crow's Bluff, between Eustis and Deland), Lake County, Florida.

RANGE.—East of St. Johns River and all of

peninsular Florida south of Levy and Marion

counties.

HABITAT.—Lentic and lotic situations and burrows.

Procambarus (Leconticambarus) apalachicolae

Hobbs

FIGURE 206

Procambarus apalachicolae Hobbs, 1942b:55, figs. 26-30, 256-

265.

Procambarus (Leconticambarus) apalachicolae Hobbs,

1972a:8.

TYPES.—Holotype and allotype, USNM 81272

(a I, 9); "morphotype," USNM 81273 (all);

paratypes, MCZ, USNM.

TYPE-LOCALITY.—Roadside ditch, 11.1 miles west

of Beacon Hill on U.S. Highway 98, Bay County,

Florida.

RANGE.—Coastal flatwoods in Bay and Gulf

counties, Florida.

HABITAT.—Burrows (secondary burrower).

Procambarus (Leconticambarus) barbatus

(Faxon)

FIGURE 204

Astacus penicillatus LcConte (not Astacus penicillatus Olivier,

1791), 1856:401.

Cambarus penicillatus.—Hagen, 1870:16, 53, figs. 93, 94 (not

95 and 96, which are P. ancylus), 149.

Cambarus barbatus Faxon, 1890:621 [in part].

Cambarus (Cambarus) barbatus.—Ortmann, 1905c: 102.

Cambarus (Ortmannicus) barbatus,—Fowler, 1912:341 [by

implication].

Procambarus barbatus—Hobbs, 1942a:342 [by implication];

1942b:39, figs. 1-5.

Procambarus (Leconticambarus) barbatus Hobbs, 1972a:8,

figs. 1m, 2f. 9a-g.

TYPE.—Neotype of A. penicillatus LeConte,

MCZ 279 (s I), here designated. Specimens from

the Escambia River at Flomaton cited by Faxon

(1890) and listed as paratypes by him (1914:414)

are paratypes of Procambarus escambiensis Hobbs.

TYPE-LOCALITY.—"Habitat in fossis Georgiae

inferioris" (LeConte, 1856:401). Here restricted to

2.5 miles west of Riceboro, Liberty County,

Georgia, in pine flatwoods.

RANGE.—In the coastal plain, from north of the

Altamaha River in Georgia to the Edisto River

in South Carolina.

HABITAT.—Burrows (secondary burrower).

REMARKS.—The type of LeConte's Astacus penicillatus is no longer extant. Inasmuch as Hagen

(1870) and Faxon (1884, 1885a, 1890, and 1914)

confused other species with it, and, in the latter

work, Faxon included members of two species in

his type-series of C. barbatus ( — A. penicillatus),

his "Type" is here designated the neotype of LeConte's A. penicillatus to establish the identity of

the latter's species.

Procambarus (Leconticambarus) capillatus

Hobbs

FIGURE 211

Procambarus capillatus Hobbs, 1971b:83, fig. 1.

Procambarus (Leconticambarus) capillatus.—Hobbs, 1972a:8.

TYPES.—Holotype, allotype, and morphotype,

USNM 131454, 131455, 131456 (a I, $, all);

paratypes, USNM.

TYPE-LOCALITY.—Drainage ditch adjacent to

Burnt Corn Creek (Escambia River drainage) on

State Route 41 northwest of Brewton, Escambia

County, Alabama.

RANGE.—Escambia River basin in Conecuh and

Escambia counties, Alabama, and perhaps Escambia County, Florida.

HABITAT.—Temporary ponds, pools, and burrows

(secondary burrower).

Procambarus (Leconticambarus) econfinae

Hobbs

FIGURE 207

Procambarus econfinae Hobbs, 1942b:49, figs. 16-20, 236-245.

Procambarus (Leconticambarus) econfinae Hobbs, 1972a:8.
NUMBER 166 51

TYPES.—Holotype, allotype, and "morphotype,"

USNM 81276 ($ I, $, $ II); paratypes, MCZ,

USNM.

TYPE-LOCALITY.—Flatwoods in the northern part

of Panama City, Bay County, Florida.

RANGE.—Environs of Panama City, Bay County,

Florida.

HABITAT.—Flatwoods, in ditches and temporary

ponds (secondary burrower).

Procambarus (Leconticambarus) escambiensis

Hobbs

FIGURE 208

Cambarus barbatus Faxon, 1890:621 [in part] Faxon, 1914:

414 [in part].

Procambarus escambiensis Hobbs, 1942b:46, figs. 11-15, 226-

235.

Procambarus (Leconticambarus) escambiensis.—Hobbs, 1972a:

8.

TYPES.—Holotype, allotype, USNM 81278 (<$ I,

9 ), and "morphotype," USNM 81277 ( $ II); paratypes, MCZ, USNM.

TYPE-LOCALITY.—About 100 yards east of Perdido River on U.S. Highway 90, Escambia County,

Florida.

RANGE.—Known from the type-locality and from

the Escambia River floodplain at Flomaton, Escambia County, Alabama; reported from the latter

locality as Cambarus barbatus by Faxon (1890)

and listed as paratypes of C. barbatus ^Procambarus barbatus] by him (1914).

HABITAT.—Temporary bodies of water in flatwoods and floodplains (secondary burrower).

Procambarus (Leconticambarus) hubbelli (Hobbs)

FIGURE 213

Cambarus hubbelli Hobbs, 1938b:90 [nomen nudum; validated by Hobbs, 1940a:406, fig. 19.]

Procambarus hubbelli.—Hobbs, 1942a:342.

Procambarus (Leconticambarus) hubbelli.—Hobbs, 1972a:8.

TYPES.—Holotype and allotype, USNM 76593

(6 I, $); "morphotype," USNM 79351 (3II);

paratypes, MCZ, USNM, Chas. M.

TYPE-LOCALITY.—One mile east of Bonifay on

U.S. Highway 90, Holmes County, Florida.

RANGE.—Choctawhatchee River drainage in Alabama and Florida.

HABITAT.—Temporary ponds, pools, and burrows (secondary burrower).

Procambarus (Leconticambarus) kilbyi (Hobbs)

FIGURE 212

Cambarus kilbyi Hobbs, 1940a:410, fig. 20.

Procambarus kilbyi.—Hobbs, 1942a:343.

Procambarus (Leconticambarus) kilbyi.—Hobbs, 1972a:8.

TYPES.—Holotype, allotype, and "morphotype,"

USNM 76594 ($ I, $, SU); paratypes, MCZ,

USNM.

TYPE-LOCALITY.—Seven miles northwest of

Blountstown on State Route 71, Calhoun County,

Florida.

RANGE.—Coastal flatwoods from Calhoun and

Gulf counties east and south to Levy County,

Florida.

HABITAT.—Temporary ponds, pools, and burrows (secondary burrower).

Procambarus (Leconticambarus) latipleurum

Hobbs

FIGURE 205

Procambarus latipleurum Hobbs, 1942b:52, figs. 21-25, 246-

255.

Procambarus (Leconticambarus) latipleurum.—Hobbs, 1972a:8.

TYPES.—Holotype, allotype, and "morphotype,"

USNM 81281 («jl, 9, a II); paratypes, MCZ,

USNM, UF (now in USNM).

TYPE-LOCALITY.—5.8 miles west of Wewahitchka

on State Route 22, Gulf County, Florida.

RANGE.—Gulf County, Florida, north of the

Wetappo Canal.

HABITAT.—Temporary ponds, pools, and burrows (secondary burrower).

Procambarus (Leconticambarus) milleri Hobbs

FICURE 203

Procambarus milleri Hobbs, 1971c: 115, figs. 1-11, 16.

Procambarus (Leconticambarus) milleri.—Hobbs, 1972a:8.

TYPES.—Holotype and morphotype, USNM

131257, 131258 (<$ I, ^ II); paratypes, USNM.

TYPE-LOCALITY.—Well at Little Bird Nursery

and Garden Store at 8427 Bird Road, Miami, Dade

County, Florida (Sec. 15, T. 54 S., R. 40 E.).
52

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

RANGE.—Known only from the type-locality.

HABITAT.—Subterranean waters.

Procambarus (Leconticambarus) pubischelae

Hobbs

FIGURE 209

Procambarus pubischelae Hobbs, 1942b:41, figs. 6-10, 216-225.

Procambarus pubischaelae.—Villalobos, 1955:38 [erroneous

spelling].

Procambarus (Leconticambarus) pubischelae.—Hobbs, 1972a:8.

TYPES.—Holotype, allotype, and "morphotype,"

USNM 81284 (a I, $, 3II); paratypes, MCZ,

ANSP, USNM.

TYPE-LOCALITY.—Cypress ponds and roadside

ditch 9.4 miles north of Lake City on U.S. Highway 441, Columbia County, Florida.

RANGE.—From the Altamaha River in Georgia

south to Alachua County, Florida.

HABITAT.—Burrows (secondary burrower).

Procambarus (Leconticambarus) rathbunae

(Hobbs)

FIGURE 210

Cambarus rathbunae Hobbs, 1940a:414, fig. 21.

Procambarus rathbunae.—Hobbs, 1942a:344.

Procambarus {Leconticambarus) rathbunae.—Hobbs, 1972a:8.

TYPES.—Holotype, allotype, and "morphotype,"

USNM 76595 ($ I, $, <$ II); paratypes, MCZ,

USNM.

TYPE-LOCALITY.—Near the Yellow River at Milligan, Okaloosa County, Florida, at the intersection

of U.S. Highway 90 and State Route 4.

RANGE.—Okaloosa and Holmes counties, Florida.

HABITAT.—Temporary bodies of water and burrows (secondary burrower).

Procambarus (Leconticambarus) shermani

Hobbs

FIGURE 214

Procambarus shermani Hobbs, 1942b:61, figs. 36-40, 266-275.

Procambarus (Leconticambarus) shermani Hobbs, 1972a:8.

TYPES.—Holotype and "morphotype," USNM

81287 (a I, ,$11); allotype, USNM 81288 (?);

paratypes, MCZ, USNM.

TYPE-LOCALITY.—About 12 miles southwest of

Jay, Santa Rosa County, Florida, in the Escambia

River swamp along McCaskill's Mill Creek.

RANGE.—Escambia River drainage in Florida,

Pearl River County, Mississippi, and St. Tammany

Parish, Louisiana.

HABITAT.—Streams and sloughs.

Subgenus Lonnbergius Hobbs

Lonnbergius Hobbs, 1972a:8. [Type-species by original designation, Cambarus acherontis Lonnberg, 1895:6. Gender:

masculine.]

Procambarus (Lonnbergius) acherontis

(Lonnberg)

FICURE 215

Cambarus acherontis Lonnberg, 1895:6, figs. l-5b Hobbs,

1940a: 390, fig. 15a-k.

Cambarus (Cambarus) acherontis Ortmann, 1905c: 102.

Cambarus (Ortmannicus) acherontis.—Fowler, 1912:341 [by

implication].

Procambarus acherontis.—Hobbs, 1942a:342 [by implication].

Procambarus (Lonnbergius) acherontis Hobbs, 1972a:8, figs.

2g, lOa-g.

TYPE.—Holotype, ZIAS 1/4412 (1 specimen).

TYPE-LOCALITY.—Subterranean rivulet about 42

feet from surface, Lake Brantley, Seminole County,

Florida.

RANGE.—Only one additional locality known:

Palm Springs, near Sanlando Springs, Seminole

County, Florida.

HABITAT.—Springs and underground streams.

REMARKS.—Hobbs (1940a:393), endeavoring to

clarify the confusion engendered by Faxon as to

the identity of the species (see P. lucifugus lucifugus) and believing that both of Lonnberg's types

were lost, designated neotypes from the Palm

Springs locality. Dr. Y. Starobogatov kindly has

informed me that one of these specimens is still

extant (see above); thus, the neotype designation

should be rejected.

Subgenus Mexicambarus Hobbs

Mexicambarus Hobbs, 1972a:8. [Type-species by original designation, Cambarus (Cambarus) bouvieri Ortmann, 1909:

159. Gender: masculine.]
NUMBER 166

Procambarus (Mexicambarus) bouvieri (Ortmann)

FIGURE 216

Cambarus (Cambarus) bouvieri Ortmann, 1909:159, figs. 1, 2.

Cambarus (Ortmannicus) bouvieri Fowler, 1912:341 [by

implication].

Procambarus bouvieri.—Hobbs, 1942a:342 [by implication]—

Villalobos, 1946:224, pi. 2: figs. 1-12.

Procambarus (Mexicambarus) bouvieri.—Hobbs, 1972a:9, figs.

2h, lla-g.

TYPES.—Syntypes, MHNP (2 <$ I, $ ), CM 74.864

TYPE-LOCALITY.—Uruapan, Michoacan, Mexico.

RANGE.—Two localities in the vicinity of Uruapan, Michoacan, Mexico.

HABITAT.—Streams.

Subgenus Ortmannicus Fowler

Ortmannicus Fowler, 1912:341. [Type-species by original designation, Astacus Blandingii Harlan, 1830:464. Gender:

masculine. Ortmannicus is a replacement name for Ortmann's subgenus Cambarus, 1905c:97.]

Procambarus (Ortmannicus) acutissimus (Girard)

FIGURE 246

Cambarus acutissimus Girard, 1852:91.

Cambarus acutus [in part].—Many authors prior to 1942.

Cambarus blandingii acutus [in part] Most authors prior to

1942.

Procambarus blandingii acutus [in part] Most authors from

1942 to 1955.

Procambarus acutissimus.—Hobbs, 1942a:342 [by implication]; 1955a:98; 1962a:286, fig. 51.

Procambarus (Ortmannicus) acutissimus Hobbs, 1972a:9.

TYPES.—ANSP 309 (2 $ II).

TYPE-LOCALITY.—Affluent of Mobile River,

Kemper County, Mississippi.

RANGE.—Tombigbee, Alabama, and Choctawhatchee drainages in east-central Mississippi and westcentral Alabama.

HABITAT.—Sluggish streams, ponds, and roadside ditches.

Procambarus (Ortmannicus) acutus acutus

(Girard)

FIGURE 240

Cambarus blandingii [in part].—Most authors from 1852 to

53

Cambarus acutus Girard, 1852:91.

1962.

Cambarus blandingii blandingii [in part].—Most authors

from 1852 to 1942.

Cambarus acutus var. A.—Hagen, 1870:36.

Cambarus acutus var. B.—Hagen, 1870:36.

Cambarus stygius Bundy, 1876:3. [Type not extant. Typelocality, Lake Michigan at Racine, Racine County, Wisconsin.]

Cambarus Blandingii var. acuta.—Faxon, 1884:136; 1885a:20,

pi. 7: figs. 2, 2', 2", 2a, 2a'.

Cambarus blandingii acutus.—Faxon, 1890:619.

Camborus blandingii acutus.—Williamson, 1899:47 [erroneous

spelling].

Cambarus blandingi acutus.—Ortmann, 1905c: 105.

Cambarus (Cambarus) blandingi acutus.—Ortmann, 1905c: 126

[by implication].

Cambarus (Ortmannicus) blandingii acutus.—Fowler, 1912:

341 [by implication].

Cambarus (Cambarus) blandingii acutus.—Fleming, 1938:301.

Procambarus blandingii acutus.—Hobbs, 1942a:343 [by implication].

Procambarus blandingii blandingii [in part].—Most authors

from 1942 to 1962.

Ortmannicus blandingii acutus.—Rhoades, 1944a: 114.

Cambarus blandingi acutis.—Roberts, 1944:364 [erroneous

spelling].

Procambarus acutus acutus.—Hobbs, 1967a: 130.

Procambarus (Ortmannicus) acutus acutus Hobbs, 1972a:9.

TYPES.—Lost in Chicago fire, 1871 (Faxon,

1914:413).

TYPE-LOCALITY.—From an affluent of Mobile

River in Kemper County, Mississippi.

RANGE.—Coastal plain and piedmont from

Maine to Georgia, from Florida panhandle to

Texas, and from Minnesota to Ohio; intergrades

with cuevachicae in southwestern Texas and northern Mexico.

HABITAT.—Sluggish to moderately flowing

streams and most lentic situations.

Procambarus (Ortmannicus) acutus cuevachicae

(Hobbs)

FIGURE 241

Cambarus blandingii cuevachicae Hobbs, 1941 a: 1, fig. 1.

Procambarus blandingii cuevachicae.—Hobbs, 1942a:S42.—

Villalobos, 1955:70, pi. 13: figs. 1-7; pi. 14: figs. 1-6.

Procambarus acutus cuevachicae.—Hobbs, 1966b:70.

Procambarus (Ortmannicus) acutus cuevachicae.—Hobbs,

1972a:9.

TYPES.—Holotype, allotype, and morphotype,

USNM 80030 (3 I, $, ill) ; paratypes, USNM,

PM.
54 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

TYPE-LOCALITY.—La Cueva Chica, a limestone

cave about 1 mile northeast of Pujal, San Luis

Potosf, Mexico.

RANGE.—Four localities in the states of San Luis

Potosi and Puebla, Mexico.

HABITAT Subterranean and lentic situations.

Procambarus (Ortmannicus) ancylus Hobbs

FIGURE 233

Cambarus penicillatus.—Hagen, 1870:54 [in part], figs. 95, 96.

Procambarus ancylus Hobbs, 1958a: 164, figs. 13-24.

Procambarus {Ortmannicus) ancylus.—Hobbs, I972a:9.

TYPES.—Holotype, allotype, and morphotype,

USNM 115050 ($1, $, $ II); paratypes, USNM.

TYPE-LOCALITY.—Summerville, Dorchester County, South Carolina.

RANGE.—Bladen, Brunswick, and Columbus

counties, North Carolina, south to Lexington,

Richland, and Colleton counties, South Carolina.

HABITAT.—Lentic and lotic situations and burrows.

Procambarus (Ortmannicus) angustatus

(LeConte)

FIGURE 226

Astacus angustatus LeConte, 1856:401.

Cambarus angustatus.—Hagen, 1870:50, figs. 65-67, 146.

Cambarus (Cambarus) angustatus.—Ortmann, 1905c: 102.

Cambarus (Ortmannicus) angustatus Fowler, 1912:341 [by

implication.]

Procambarus angustatus.—Hobbs, 1942a:342 [by implication].

Procambarus (Ortmannicus) angustatus.—Hobbs, 1972a:9.

TYPE.—Holotype, ANSP 444 («$ I).

TYPE-LOCALITY.—"Georgia inferiore, in aquae

purae rivulos qui inter colliculos arenosos (Sandhills) currunt."

RANGE.—Known from the type-specimens only.

Repeated searches for this species in Georgia have

been unsuccessful.

HABITAT.—Streams.

Procambarus (Ortmannicus) bivittatus Hobbs

FIGURE 255

Procambarus bivittatus Hobbs, 1942b:96, figs. 96-100, 305-314.

Procambarus (Ortmannicus) bivittatus.—Hobbs, 1972a: 10.

TYPES.—Holotype, USNM 81274 ( $ I); allotype

and "morphotype," USNM 81275 ( 9 , 6 II); paratypes, MCZ, USNM.

TYPE-LOCALITY.—Sloughs along Escambia River

on State Route 62, Escambia County, Florida.

RANGE.—Escambia River drainage in Escambia

and Santa Rosa counties, Florida, to the Pearl

River drainage in St. Tammany and Washington

parishes, Louisiana.

HABITAT.—Streams and sloughs.

Procambarus (Ortmannicus) blandingii (Harlan)

FIGURE 239

Astacus Blandingii Harlan, 1830:464.

Astacus (Cambarus) Blandingii.—Erichson, 1846:98.

Cambarus Blandingii Girard, 1852:91.

Cambarus Blandingii var. Blandingii—Faxon, 1884:136 [by

implication].

Cambarus blandingii blandingii.—Faxon, 1890:619 [by implication].

Cambarus (Cambarus) blandingi.—Ortmann, 1905c: 102.

Cambarus (Cambarus) blandingi blandingi Ortmann, 1905c:

126 [by implication].

Cambarus blandingi-typicus.—Ortmann, 1905c: 105.

Cambarus (Ortmannicus) blandingii.—Fowler, 1912:341.

Procambarus blandingii blandingii.—Hobbs, 1942a:342 [by

implication].

Procambarus blandingii.—Hobbs, 1967a: 130.

Procambarus (Ortmannicus) blandingii.—Hobbs, 1972a:9, figs.

2i, 12a-g.

TYPE.—Holotype, ANSP 4177 ( <$ I).

TYPE-LOCALITY.—Camden, Kershaw County,

South Carolina.

RANGE.—From the Santee River to the Pee Dee

River system in South Carolina and southern

North Carolina.

HABITAT.—Lentic and lotic situations and burrows.

Procambarus (Ortmannicus) caballeroi

Villalobos

FIGURE 257

Procambarus caballeroi Villalobos, 1944b: 175, pis. 1, 2.

Procambarus (Ortmannicus) caballeroi.—Hobbs, 1972a:9.

TYPES.—Holotype, allotype, and morphotype,

IBM 50040 (004) ( $ I, $, $ II); paratypes, IBM,

USNM.
NUMBER 166 55

TYPE-LOCALITY.—"Riachuelos del Sur de Villa

Juarez," Puebla, Mexico.

RANGE.—Known only from the type-locality.

HABITAT.—Streams.

Procambarus (Ortmannictts) chacei Hobbs

FIGURE 219

Procambarus chacei Hobbs, 1958b:5, pi. 2: figs. 14-26.

Procambarus sp. A.—Hobbs, 1958c:72, 76, 77, 79, 83-S6, 90.

Procambarus (Ortmannicus) chacei.—Hobbs, 1972a:9.

TYPES.—Holotype, allotype, and morphotype,

USNM 101289, 101290, 101291 (a I, 9, $ II);

paratypes, MCZ, ANSP, IBM, TU, USNM.

TYPE-LOCALITY.—Cedar Creek, 3 miles east of

Lykes, Richland County, South Carolina, on U.S.

Highway 76.

RANGE.—From the Wateree River in South Carolina south to the Canoochee River in Georgia.

HABITAT.—Streams.

Procambarus (Ortmannicus) enoplosternum

Hobbs

FIGURE 218

Procambarus enoplosternum Hobbs, 1947a:5, figs. 2, 5, 10, 12,

14, 18, 20, 23-25, 32.

Procambarus (Ortmannicus) enoplosternum.—Hobbs, 1972a:9.

TYPES.—Holotype, allotype, and "morphotype,"

USNM 82263 ($ I, $, $11); paratypes, MCZ,

USNM, TU.

TYPE-LOCALITY.—Rocky Creek, 6 miles south of

Lyons on U.S. Highway 1, Toombs County,

Georgia.

RANGE.—Lower Oconee and Ohoopee drainages

in Georgia.

HABITAT.—Streams.

REMARKS.—Hobbs (1947a: 14) erred in his explanation of the illustrations of this species: figure

14 (not 13) represents the basipodites and ischiopodites of the fourth and third pereiopods of

P. enoplosternum.

Procambarus (Ortmannicus) epicyrtus Hobbs

FIGURE 220

Procambarus epicyrtus Hobbs, 1958b: 1, pi. 1: figs. 1—IS.

Procambarus (Ortmannicus) epicyrtus.—Hobbs, 1972a:9.

TYPES.—Holotype, allotype, and morphotype,

USNM 101286, 101287, 101288 ($1, 5, <$ II);

paratypes, MCZ, ANSP, IBM, TU, USNM.

TYPE-LOCALITY.—South Ogeechee Creek, 6.8

miles south of Sylvania on U.S. Highway 306,

Screven County, Georgia.

RANGE.—Tributaries of the Ogeechee and

Oconee rivers in Georgia.

HABITAT.—Streams.

Procambarus (Ortmannicus) evermanni (Faxon)

FIGURE 237

Cambarus evermanni Faxon, 1890:620.

Cambarus (Cambarus) evermanni.—Ortmann, 1905c: 102.

Cambarus (Ortmannicus) evermanni.—Fowler, 1912:341 [by

implication].

Procambarus evermanni.—Hobbs, 1942a:342 [by implication];

1942b: 107, figs. 111-115.

Procambarus (Ortmannicus) evermanni.—Hobbs, 1972a: 10.

TYPE.—Holotype, MCZ 3834 ($ I).

TYPE-LOCALITY.—Escambia River at Flomaton,

Escambia County, Alabama.

RANGE.—In addition to the type-locality, known

from Santa Rosa and Escambia counties, Florida,

to Jackson County, Mississippi.

HABITAT.—Sluggish streams. Specimen from Mississippi dug from simple burrows in drying creek

bed.

Procambarus (Ortmannicus) fallax (Hagen)

FIGURE 235

Cambarus fallax Hagen, 1870:45, figs. 103-105.

Cambarus (Cambarus) fallax.—Ortmann, 1905c: 102.

Cambarus (Ortmannicus) fallax.—Fowler, 1912:341 [by implication].—Creaser, 1934b:4 [by implication].

Procambarus fallax.—Hobbs, 1942a:342 [by implication];

1942b:111, figs. 116-120.

Procambarus (Ortmannicus) fallax.—Hobbs, 1972a:9.

TYPES Syntypes, MCZ 3526 ($ I, «$ II), BSNH

(& II, 9 ), USNM 63294 («j II, 9 )•

TYPE-LOCALITY.—Florida. Here restricted to St.

Johns River at Welaka, Putnam County, Florida.

RANGE.—Tributaries of the Satilla River,

Georgia, southward through peninsular Florida.

HABITAT.—Lentic and lotic situations and burrows.
56 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Procambarus (Ortmannicus) gonopodocristatus

Villalobos

FIGURE 260

Procambarus gonopodocristatus Villalobos, 1958:279, pis. 1-3.

Procambarus (Ortmannicus) gonopodocristatus.—Hobbs,

1972a: 10.

TYPES.—Holotype, allotype, and morphotype,

IBM (<$I, 9, $ II); paratypes, USNM, AMNH.

TYPE-LOCALITY.—Paso Largo, 16 km northeast

of Martinez de la Torre, Veracruz, Mexico.

RANGE.—Only one other locality known—

"Maria de la Torre, km. 406 de la carretera Martinez de la Torre, Nautla," Veracruz, Mexico.

HABITAT.—Lentic situations.

Procambarus (Ortmannicus) hayi (Faxon)

FIGURE 244

Cambarus hayi Faxon, 1884:108.

Cambarus (Cambarus) hayi.—Ortmann, 1905c: 102.

Cambarus (Ortmannicus) hayi.—Fowler, 1912:341 [by implication].—Creaser, 1934b:4 [by implication].

Procambarus hayi.—Hobbs, 1942a:342 [by implication].

Procambarus (Ortmannicus) hayi.—Hobbs, 1972a:9.

TYPES.—Syntypes, MCZ 3533 ( s I, 2 $ II, 9,

cjjuv.), USNM 19751 ($11), 19752 (all, 9),

21850 (2 a I, 9).

TYPE-LOCALITY.—Macon, Noxubee County, Mississippi.

RANGE.—Tributaries of the Tombigbee and Tallahatchie rivers in Alabama and Mississippi, and

Hatchie drainage in Tennessee.

HABITAT.—Sluggish streams and lentic situations.

drainage systems in South Carolina.

HABITAT.—Streams.

Procambarus (Ortmannicus) horsti

Hobbs and Means

FIGURE 230

Procambarus horsti Hobbs and Means, 1972:301, fig. 2.

TYPES.—Holotype, allotype, and morphotype,

USNM 132043, 132044, 132045 ($1, 9, 6II);

paratypes, USNM.

TYPE-LOCALITY.—Big Blue Springs (tributary to

the Wacissa River), 2.2 miles south of the crossroads in the town of Wacissa, Jefferson County,

Florida (SE14NW14 sec. 12, T. 2 S., R. 3 E.).

RANGE.—Known only from the type-locality.

HABITAT.—Subterranean waters.

Procambarus (Ortmannicus) hybus

Hobbs and Walton

FIGURE 249

Procambarus hybus Hobbs and Walton, 1957:39, figs. 1-12.

Procambarus (Ortmannicus) hybus.—Hobbs, 1972a: 10.

TYPES.—Holotype, allotype, and morphotype,

USNM 99581, 99582, 99583 ($ I, 9, Sll); paratypes, TU, USNM.

TYPE-LOCALITY.—Roadside ditch 1.7 miles north

of Boligee, Greene County, Alabama.

RANGE.—Tombigbee drainage in Alabama and

Mississippi.

HABITAT.—Temporary ponds, pools, and burrows (secondary burrower).

Procambarus (Ortmannicus) hirsutus Hobbs

FIGURE 224

Procambarus hirsutus Hobbs, 1958a: 160, figs. 1-12.

Procambarus sp. B.—Hobbs, 1958c:72, 76, 77, 79, 84, 86, 90.

Procambarus (Ortmannicus) hirsutus Hobbs, 1972a:9.

TYPES.—Holotype, allotype, and morphotype,

USNM 101148, 101150, 101149 ($1, 9, $ II);

paratypes, TU, Chas. M., USNM.

TYPE-LOCALITY.—Salkehatchie River, 1.9 miles

south of Barnwell, Barnwell County, South Carolina, on State Route 3.

RANGE.—Edisto, Salkehatchie, and Savannah

Procambarus (Ortmannicus) jaculus

Hobbs and Walton

FIGURE 248

Procambarus jaculus Hobbs and Walton, 1957:48, figs. 24-

34.—Walls and Black, 1967:60.

Procambarus (Ortmannicus) jaculus.—Hobbs, 1972a: 10.

TYPES.—Holotype, allotype, and morphotype,

USNM 99584, 99585, 99586 («$ I, 9, $11); paratype, USNM.

TYPE-LOCALITY.—Roadside ditch 1 mile west of

Scott-Rankin county line on U.S. Highway 80,

Rankin County, Mississippi.
NUMBER 166

RANGE.—Hinds, Rankin, and Scott counties,

Mississippi, and Avoyelles Parish, Louisiana.

HABITAT.—Temporary lentic situations and burrows (secondary burrower).

Procambarus (Ortmannicus) lecontei (Hagen)

FIGURE 243

Cambarus LeContei Hagen, 1870:47, figs. 15-18, 145.

Cambarus Lecontei Faxon, 1884:110.

Cambarus lecontei.—Hay, 1899b:959.

Cambarus (Cambarus) lecontei Ortmann, 1905c: 102.

Cambarus (Ortmannicus) lecontei Fowler, 1912:341 [by implication].—Creaser, 1934b:4 [by implication].

Procambarus lecontei Hobbs, 1942a:342 [by implication];

1952a:210, fig. 81.

Procambarus (Ortmannicus) lecontei Hobbs, 1972a:9.

TYPES.—Syntypes, MCZ 217 ( i I, 7 9 , 2 ^ II),

USNM 4958 («$ I), AMS (lost), MHNP ($ I, 9),

Wiirzburg Mus.

TYPE-LOCALITY.—Mobile County, Alabama.

RANGE.—Mobile County, Alabama, and Stone

County, Mississippi.

HABITAT.—Streams.

Procambarus (Ortmannicus) leonensis Hobbs

FIGURE 236

Procambarus leonensis Hobbs, 1942b: 114, figs. 121-125;

1943a:49, figs. 1, 6, 7, 10, 12-14, 16, 17, 22, 26, 29, 31.

Procambarus (Ortmannicus) leonensis.—Hobbs, 1972a:9.

TYPES.—Holotype, allotype, and "morphotype,"

USNM 81091 ($ I, $, «$II); paratypes, MCZ,

USNM, ANSP.

TYPE-LOCALITY.—Sinkhole pond, 3 miles southwest of Tallahassee on State Route 20, Leon

County, Florida. (Now beneath runway at airport.)

RANGE.—Between the Apalachicola and Suwannee rivers, Florida.

HABITAT.—Lentic and lotic situations, and burrows.

Procambarus (Ortmannicus) lepidodactylus

Hobbs

FIGURE 222

Procambarus lepidodactylus Hobbs, 1947b:25, figs. 1-12.

Procambarus (Ortmannicus) lepidodactylus.—Hobbs, 1972a:9.

57

TYPES.—Holotype and allotype, USNM 84198

(<$I, 9); morphotype, USNM 84199 ($ II); paratypes, USNM.

TYPE-LOCALITY.—Juniper Creek, 1 mile southwest of Patrick on U.S. Highway 1, Chesterfield

County, South Carolina.

RANGE.—Santee and Pee Dee drainage systems

in eastern South Carolina and Columbus County,

North Carolina.

HABITAT.—Streams.

Procambarus (Ortmannicus) lewisi

Hobbs and Walton

FIGURE 254

Procambarus lewisi Hobbs and Walton, 1959:39, figs. 1-12.

Procambarus (Ortmannicus) lewisi.—Hobbs, 1972a: 10.

TYPES.—Holotype, allotype, and morphotype,

USNM 102467, 102468, 102469 ($ I, $, <$ II);

paratypes, TU, USNM.

TYPE-LOCALITY.—Roadside ditch 18.8 miles east

of Montgomery on U.S. Highway 80, Macon

County, Alabama.

RANGE.—Barbour, Lowndes, Macon, and Montgomery counties, Alabama.

HABITAT.—Sluggish streams and roadside ditches.

Procambarus (Ortmannicus) litosternum Hobbs

FIGURE 221

Procambarus litosternum Hobbs, 1947a:9, figs. 3, 4, 9, 11, 13,

16, 19, 21, 26, 29, 30.

Procambarus (Ortmannicus) litosternum.—Hobbs, 1972a:9.

TYPES.—Holotype, allotype, and "morphotype,"

USNM 82261 («jl, $, <$ II); paratypes, USNM.

TYPE-LOCALITY.—Stream 5 miles northeast of

Swainsboro on U.S. Highway 25, Emanuel County,

Georgia.

RANGE.—Tributaries of the Canoochee, Ogeechee, and Newport rivers in Bulloch, Jenkins,

Emanuel, and Liberty counties, Georgia.

HABITAT.—Streams.

REMARKS.—Hobbs (1947a: 14) erred in his explanation of the illustrations of this species: figure

13 (not 15) represents the lateral view of the first

pleopod of male, form I, of P. litosternum.
58 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Procambarus (Ortmannicus) lophotus

Hobbs and Walton

FIGURE 245

Procambarus lophotus Hobbs and Walton, 1960:123, figs. 1-11.

Procambarus (Ortmannicus) lophotus.—Hobbs, 1972a:9.

TYPES.—Holotype, allotype, and morphotype,

USNM 104404, 104405, 104406, ($ I, $, $11);

paratypes, TU, USNM.

TYPE-LOCALITY.—Roadside ditch 3.4 miles northeast of Haynesville on State Route 111, Lowndes

County, Alabama.

RANGE.—Alabama drainage system from Clarke

County, Alabama, to Gordon and Catoosa counties,

Georgia, and in the Tennessee drainage in Polk

County, Tennessee, and northwestern Georgia.

HABITAT.—Lentic and lotic situations and burrows (tertiary burrower).

Procambarus (Ortmannicus) lucifugus alachua

(Hobbs)

FIGURE 227

Cambarus lucifugus alachua Hobbs, 1940a:402, fig. 18a-/.

Procambarus lucifugus alachua.—Hobbs, 1942a:343.

Procambarus (Ortmannicus) lucifugus alachua Hobbs,

1972a: 10.

TYPES.—Holotype, allotype, and "morphotype,"

USNM 76592 ($ I, $, ^ II); paratypes, USNM.

TYPE-LOCALITY.—A small cave, Hog Sink, about

10 miles west of Gainesville, Alachua County,

Florida (Sec. 24, T. 10 S., R. 18 E.).

RANGE.—Southwestern Alachua and western Gilchrist counties, Florida.

HABITAT.—Subterranean waters.

Procambarus (Ortmannicus) lucifugus lucifugus

(Hobbs)

FIGURE 228

Cambarus acherontis Faxon [not Lonnberg], 1898:645, pi. 62:

figs. 1-5.

Cambarus lucifugus lucifugus Hobbs, 1940a:398, figs. 17,

18m,n.

Procambarus lucifugus lucifugus Hobbs, 1942a:343.

Procambarus (Ortmannicus) lucifugus lucifugus.—Hobbs,

1972a:9.

TYPES.—Holotype, allotype, and "morphotype,"

USNM 77916, 77917, 77918 (s I, 9, $11); paratypes, MCZ, USNM.

TYPE-LOCALITY.—Gum Cave, about 7 miles

southwest of Floral City, Citrus County, Florida.

RANGE.—Citrus and Hernando counties northward to Marion County, Florida, where it intergrades with P. lucifugus alachua.

HABITAT.—Subterranean waters.

Procambarus (Ortmannicus) lunzi (Hobbs)

FICURE 234

Cambarus lunzi Hobbs, 1940b:3, figs. 1-10.

Procambarus lunzi.—Hobbs, 1942a:343.

Procambarus (Ortmannicus) lunzi.—Hobbs, 1972a:9.

TYPES.—Holotype and allotype, USNM 79301

( i h 9).

TYPE-LOCALITY.—1.4 miles southwest of Early

Branch on State Route 28, Hampton County, South

Carolina.

RANGE.—Hampton and Beaufort counties, South

Carolina, southward to Mclntosh County, Georgia.

HABITAT.—Lentic situations and burrows (secondary or tertiary burrower).

Procambarus (Ortmannicus) mancus

Hobbs and Walton

FIGURE 251

Procambarus mancus Hobbs and Walton, 1957:44, figs. 13-23.

Procambarus (Ortmannicus) mancus.—Hobbs, 1972a: 10.

TYPES.—Holotype, allotype, and morphotype,

USNM 99578, 99579, 99580 ($ I, $, $11); paratypes, TU, USNM.

TYPE-LOCALITY.—Roadside ditch, 5 miles south

of Meridian, Lauderdale County, Mississippi.

RANGE.—Known only from Lauderdale and Newton counties, Mississippi.

HABITAT.—Burrows (secondary burrower).

Procambarus (Ortmannicus) orcinus

Hobbs and Means

FICURE 231

Procambarus orcinus Hobbs and Means, 1972:394, fig. I.

TYPES.—Holotype, allotype, and morphotype,
NUMBER 166 59

USNM 132031, 132032, 132033 («?I, 9, «H);

paratypes, USNM.

TYPE-LOCALITY.—Gopher Sink, 3.1 miles southwest of Florida Road 61 and 0.2 mile east of Florida

Road 369 (SWi/4NW^NEi^, sec. 16, T. 2 S., R.

1 W.), Leon County, Florida.

RANGE.—Known from four localities in Leon and

Wakulla counties, Florida.

HABITAT.—Subterranean waters.

Procambarus (Ortmannicus) pallidus (Hobbs)

FIGURE 229

Cambarus acherontis.—Hobbs [not Lonnberg], 1937:154.

Cambarus acherontis pallidus Hobbs, 1938b:9O [nomen

nudum].

Cambarus pallidus Hobbs, 1940a:394, fig. 16.

Procambarus pallidus Hobbs, 1942a:343.

procambarus pallidus.—Padgett, 1970:19 [lapsus calami].

Procambarus (Ortmannicus) pallidus.—Hobbs, 1972a: 10.

TYPES.—Holotype and allotype, USNM 76591

(£l, $ ); "morphotype," USNM 79350 (<$ II);

paratypes, MCZ, USNM.

TYPE-LOCALITY.—Warren's Cave, 11 miles northwest of Gainesville, Alachua County, Florida (sec.

13, T. 9 S., R. 18 E.).

RANGE.—Subterranean waters in the northwestern part of Alachua County to Suwannee County,

Florida.

HABITAT.—Subterranean waters.

Procambarus (Ortmannicus) pearsei pearsei

(Creaser)

FICURE 253

Cambarus (Ortmannicus) pearsei Creaser, 1934b: 1, figs. 1-3.

Procambarus pearsei.—Hobbs, 1942a:343.

Procambarus pearsei pearsei.—Hobbs and Walton, 1958:7.

Procambarus (Ortmannicus) pearsei pearsei.—Hobbs, 1972a:

10.

TYPES.—Holotype, USNM 98336 (£ I); allotype,

USNM 98337 (9); paratypes, USNM.

TYPE-LOCALITY.—Pond and ditch on State Route

22, south of Fayetteville, Cumberland County,

North Carolina.

RANGE.—Cumberland County, North Carolina,

south to Horry County, South Carolina, intergrading with P. pearsei plumimamis in Johnston

and Pitt counties, North Carolina.

HABITAT.—Lentic situations and burrows (secondary burrower).

Procambarus (Ortmannicus) pearsei plumimanus

Hobbs and Walton

FICURE 252

Procambarus pearsei plumimanus Hobbs and Walton, 1958:8,

figs. 1-13.

Procambarus (Ortmannicus) pearsei plumimanus.—Hobbs,

1972a: 10.

TYPES.—Holotype, allotype, and morphotype,

USNM 100940, 100941, 100942 («J I, 9, <J II);

paratypes, TU, USNM.

TYPE-LOCALITY.—Roadside ditch 2.2 miles southeast of Havelock on U.S. Highway 70, Craven

County, North Carolina.

RANGE.—Carteret, Craven, and Duplin counties,

North Carolina.

HABITAT.—Temporary ponds, pools, and burrows (secondary burrower).

Procambarus (Ortmannicus) pictus (Hobbs)

FIGURE 217

Cambarus pictus Hobbs, 1940a:419, fig. 22.

Procambarus pictus.—Hobbs, 1942a:344.

Procambarus (Ortmannicus) pictus.—Hobbs, 1972a:9.

TYPES.—Holotype, allotype, and "morphotype,"

USNM 76596 (£ I, 9, £ II); paratypes, MCZ,

USNM.

TYPE-LOCALITY.—A small, swift swamp stream,

about 2 miles southwest of Green Cove Springs,

Clay County, Florida.

RANGE.—Known from several small tributaries

of Black Creek in Clay County, Florida.

HABITAT.—Streams.

Procambarus (Ortmannicus) planirostris Penn

FIGURE 250

Procambarus planirostris Penn, 1953c:71, figs. 1-12.

Procambarus (Ortmannicus) planirostris.—Hobbs, 1972a: 10.

TYPES.—Holotype, allotype, and morphotype,

USNM 95674, 95675, 95676 (<$ I, 9 , <$H); paratypes, ANSP, TU, USNM.

TYPE-LOCALITY.—Flatwoods, 1 mile south of
60 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Walker on State Route 336, Livingston Parish,

Louisiana.

RANGE.—"Florida" parishes of southeastern Louisiana, and Perry, Pearl River, and Winston counties, Mississippi.

HABITAT.—Temporary ponds, pools, and burrows (secondary burrower).

Procambarus (Ortmannicus) pubescens (Faxon)

FIGURE 223

Cambarus pubescens Faxon, 1884:109.

Cambarus (Cambarus) pubescens.—Ortmann, 1905c:101.

Cambarus (Ortmannicus) pubescens Fowler, 1912:341 [by

implication].

Procambarus pubescens.—Hobbs, 1942a:342 [by implication];

1947a:l, figs. 1, 6-8, 15, 17, 22, 27. 28, 31.

Procambarus (Ortmannicus) pubescens.—Hobbs, 1972a:9.

TYPES.—Syntypes, USNM 3181 ( $ II, 9).

TYPE-LOCALITY.—McBean Creek, south of Augusta, Richmond County, Georgia.

RANGE.—In the Oconee, Ogeechee, and Savannah drainage systems in Georgia and South Carolina.

HABITAT.—Streams.

REMARKS.—Hobbs (1947a: 14) erred in his explanations of the illustrations of this species: figure 15 (not figure 14) represents the basipodites

and ischiopodites of the fourth and third pereiopods of P. pubescens.

Procambarus (Ortmannicus) pycnogonopodus

Hobbs

FICURE 238

Procambarus pycnogonopodus Hobbs, 1942b:117, figs. 126-

130; 1943a:53, figs. 3, 4, 8, 9, 11, 15, 19-21, 24, 25, 27, 28,

30,32.

Procambarus (Ortmannicus) pycnogonopodus. Hobbs,

1972a:9.

TYPES.—Holotype, allotype, and "morphotype,"

USNM 81092 (<jl, 9, ^ II); paratypes, MCZ,

ANSP, USNM.

TYPE-LOCALITY.—A roadside excavation and adjoining intermittent stream in flatwoods 5.8 miles

west of Wewahitchka on State Route 22, Gulf

County, Florida.

RANGE.—From the Apalachicola River westward

to the Choctawhatchee River, Florida.

HABITAT.—Lentic and lotic situations, and burrows.

Procambarus (Ortmannicus) seminolae Hobbs

FIGURE 232

Procambarus seminolae Hobbs, 1942b: 142, figs. 166-170, 335-

346.

Procambarus (Ortmannicus) seminolae.—Hobbs, 1972a:9.

TYPES.—Holotype, allotype, and "morphotype,"

USNM 81286 ($1, 9, $ II); paratypes, MCZ,

ANSP, USNM, UF (now in USNM).

TYPE-LOCALITY.—Roadside excavation about 9

miles northeast of Gainesville on State Route 24,

Alachua County, Florida.

RANGE.—From the Altamaha River drainage in

Georgia to Marion County, Florida.

HABITAT.—Lentic and lotic situations, and burrows (secondary or tertiary burrower).

Procambarus (Ortmannicus) texanus Hobbs

FIGURE 242

Procambarus texanus Hobbs, 1971b:87, fig. 2.

Procambarus (Ortmannicus) texanus Hobbs, 1972a:9.

TYPES.—Holotype, allotype, and morphotype,

USNM 131457, 131458, 131459 (£ I, 9, «H);

paratypes, USNM.

TYPE-LOCALITY.—Fish hatchery near Smithville,

Bastrop County, Texas.

RANGE.—Known only from the type-locality.

HABITAT.—Ponds.

Procambarus (Ortmannicus) toltecae Hobbs

FIGURE 259

Procambarus toltecae Hobbs, 1943b: 198, figs. 1-14.

Procambarus (Ortmannicus) toltecae.—Hobbs, 1972a: 10.

TYPES.—Holotype and allotype, USNM 81301

( $ I, 9 ); paratypes, USNM.

TYPE-LOCALITY.—Puente de Xilitla (not Palitla

as previously recorded), small stream 8 km north

of Tomazunchale, San Luis Potosi, Mexico.

RANGE.—Southern Tamaulipas and southeastern

San Luis Potosi.

HABITAT.—Streams.
NUMBER 166 61

Procambarus (Ortmannicus) verrucosus Hobbs

FIGURE 247

Procambarus verrucosus Hobbs, 1952a:212, fig. 82.

Procambarus (Ortmannicus) verrucosus.—Hobbs, 1972a:9.

TYPES.—Holotype, allotype, and morphotype,

USNM 90743 ($\, 9 , £ II); paratypes, USNM.

TYPE-LOCALITY.—Tributary of Calebea Creek,

3.9 miles south of Tuskegee, Macon County, Alabama, on U.S. Highway 29.

RANGE.—Tributaries of the Tallapoosa and

Chattahoochee rivers in southeastern Alabama.

HABITAT.—Streams.

Procambarus (Ortmannicus) viaeviridis (Faxon)

FIGURE 256

Cambarus viae-viridis Faxon, 1914:370, pi. 5.

Cambarus (Ortmannicus) viae viridis Creaser, 1934b:4 [by

implication].

Procambarus viae-viridis.—Hobbs, 1942a:342 [by implication].

Procambarus viaeviridis.—Penn, 1959:5.

Procambarus (Ortmannicus) viaeviridis Hobbs, 1972a:9.

TYPES.—Syntypes, MCZ 7336 (2 <$ L 2 <5 II, 6 $ ).

TYPE-LOCALITY.—St. Francis River, Greenway,

Clay County, Arkansas.

RANGE.—Green and Clay counties, Arkansas, to

Tuscaloosa County, Alabama.

HABITAT.—Sluggish streams and lentic situations.

Procambarus (Ortmannicus) villalobosi Hobbs

FIGURE 258

Procambarus villalobosi Hobbs, 1969a:41, pi. 1: figs. 1-11.

Procambarus (Ortmannicus) villalobosi.—Hobbs, 1972a: 10.

TYPES.—Hoiotype and allotype, USNM 118644

and 118645 ($ I, $ ).

TYPE-LOCALITY.—Cueva del Agua, 75 km east of

Valles, San Luis Potosf, Mexico.

RANGE.—Known only from the type-locality.

HABITAT.—Streams.

Procambarus (Ortmannicus) youngi Hobbs

FICURE 225

Procambarus youngi Hobbs, 1942b: 131, figs. 146-150, 325-334.

Procambarus (Ortmannicus) youngi.—Hobbs, 1972a:9.

TYPES.—Holotype and allotype, USNM 81289

(£l, 9); "morphotype," USNM 81290 (,$ II);

paratypes, MCZ, USNM.

TYPE-LOCALITY.—Guard House Branch, about 1

mile west of Wewahitchka on State Route 22, Gulf

County, Florida.

RANGE.—Known from the type-locality; Wetappo

Creek, 6.4 miles east of Bay County line on State

Route 22; and St. Marks River in Leon and

Wakulla counties, Florida.

HABITAT.—Streams.

Subgenus Paracambarus Ortmann

Paracambarus Ortmann, 1906a: 1. [Type-species by original

designation, Cambarus (Paracambarus) paradoxus Ortmann,

1906a:3. Gender: masculine.]

Procambarus (Paracambarus) ortmannii

(Villalobos)

FIGURE 261

Paracambarus ortmannii Villalobos, 1949:331, pi. 1: figs. 1-7;

pi. 2: figs. 1-7.

Paracambarus ortmanni.—Villalobos, 1953:346.

Procambarus (Paracambarus) ortmanni Hobbs, 1972a: 10.

Procambarus (Paracambarus) ortmannii [original genitive].

TYPES.—Holotype, allotype, and morphotype,

IBM 2-1149-la(OF6) (<j I, 9, 5II); paratype,

IBM.

TYPE-LOCALITY.—"Los Estajos, 6 km NE de

Zihuateutla, Pue. en un pequeno arroyito que

corre por el lado derecho del camino que va al

rancho El Mirador," Mexico.

RANGE.—Known only from the type-locality.

HABITAT.—Stream.

Procambarus (Paracambarus) paradoxus

(Ortmann)

FIGURE 262

Cambarus (Paracambarus) paradoxus Ortmann, 1906a:3, fig. 1.

Paracambarus paradoxus.—Hobbs, 1942a:344—Villalobos,

1947a:233, pi. 1; pi. 3: figs. 1-4.

Procambarus Paradoxus.—Creaser, 1962:7 [lapsus calami].

Procambarus (Paracambarus) paradoxus.—Hobbs, 1972a: 10,

figs. 2j, 13a-g.

TYPES.—Syntypes, ANSP 5358 (3 specimens),

MHNP (25+ specimens), CM 74.677, MCZ 6955

U I, 6U, 9 )•
62 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

TYPE-LOCALITY.—Tetela de Ocampo y La

Canada, 35 km northeast of Zacapoaxtla, Estado

de Puebla, Mexico.

RANGE.—Known only from the area of the typelocality.

HABITAT.—Streams.

Subgenus Pennides Hobbs

Pennides Hobbs, 1972a: 10. [Type-species by original designation, Procambarus natchitochae Penn, 1953d:5. Gender:

masculine.]

Procambarus (Pennides) ablusus Penn

FIGURE 265

Procambarus ablusus Penn, 1963:121, figs. 1-10.

Procambarus (Pennides) ablusus.—Hobbs, 1972a: 10.

TYPES.—Holotype, allotype, and morphotype,

USNM 108200, 108202, 108201 ($ I, $, $ II);

paratypes, TU.

TYPE-LOCALITY.—Hatchie River, 12.1 miles east

of Ripley on State Route 4, Tippah County, Mississippi.

RANGE.—Hatchie River system in Hardeman

County, Tennessee, and Alcorn and Tippah counties, Mississippi.

HABITAT.—Streams.

Procambarus (Pennides) dupratzi Penn

FICURE 270

Procambarus dupratzi Penn, 1953d: 1, figs. 1-7, 10-14.

Procambarus (Pennides) dupratzi Hobbs, 1972a: 10.

TYPES.—Holotype, allotype, and morphotype,

USNM 93652, 93653, 93654 ($ I, $, 5 II); paratypes, USNM, AMNH, ANSP, MCZ, CM, TU.

TYPE-LOCALITY.—Attoyac Bayou, 5.6 miles southwest of Timpson on U.S. Highway 59, Shelby

County, Texas.

RANGE.—Drainage systems of the Trinity, Red,

Neches, Sabine, and Calcasieu rivers in Texas,

Arkansas, and Louisiana.

HABITAT.—Streams.

Procambarus (Pennides) echinatus Hobbs

FICURE 269

Procambarus echinatus Hobbs, 1956a: 117, figs. 1-17.

Procambarus (Pennides) echinatus.—Hobbs, 1972a: 10.

TYPES.—Holotype, allotype, and morphotype,

USNM 99180, 99181, 99182 ($ I, 9, $ II); paratypes, TU, USNM.

TYPE-LOCALITY.—Salkehatchie River, 1.9 miles

south of Barnwell on State Route 3, Barnwell

County, South Carolina.

RANGE.—Salkehatchie and Edisto drainage systems in South Carolina.

HABITAT.—Streams.

Procambarus (Pennides) elegans Hobbs

FICURE 266

Procambarus elegans Hobbs, 1969c:329, figs. 1-11, 24, 25.

Procambarus (Pennides) elegans.—Hobbs, 1972a: 10.

TYPES.—Holotype, allotype, and morphotype,

USNM 129892, 129893, 115838 ($ I, 5, ill);

paratype, USNM.

TYPE-LOCALITY.—Spillway from Corney Lake, an

impounded tributary of the Ouachita River, Claiborne Parish, Louisiana.

RANGE.—Ouachita drainage system in Claiborne

and Union parishes, Louisiana.

HABITAT.—Streams.

Procambarus (Pennides) gibbus Hobbs

FICURE 275

Procambarus gibbus Hobbs, 1969c:337, figs. 12-23.

Procambarus (Pennides) gibbus.—Hobbs, 1972a: 10.

TYPES.—Holotype, allotype, and morphotype,

USNM 129804, 129803, 129805 ($ I, $, $ II);

paratypes, USNM.

TYPE-LOCALITY.—Tributary to Muckalee Creek

(Flint River drainage), 3.2 miles north of Americus

on U.S. Highway 19, Sumter County, Georgia.

RANGE.—Flint River drainage system in Crawford, Lee, Schley, and Sumter counties, Georgia.

HABITAT.—Streams.

Procambarus (Pennides) lagniappe Black

FIGURE 273

Procambarus lagniappe Black, 1968:5, figs. 1-12.

Procambarus (Pennides) lagniappe Hobbs, 1972a: 10.
NUMBER 166 63

TYPES.—Holotype, allotype, and morphotype,

USNM 119088, 119089, 119090 («j I, 9, 5 II);

paratypes, TU, J. B. Black.

TYPE-LOCALITY.—Pawticfaw Creek (tributary to

Sucarnoochee Creek, tributary to Tombigbee

River), 6.0 miles south of De Kalb, Kemper County,

Mississippi, on State Route 39.

RANGE.—Tributaries of the Tombigbee River in

Kemper County, Mississippi.

HABITAT.—Streams.

Procambarus (Pennides) lylei

Fitzpatrick and Hobbs

FIGURE 268

Procambarus lylei Fitzpatrick and Hobbs, 1971:95, figs. 1-14.

Procambarus (Pennides) lylei.—Hobbs, 1972a: 10.

TYPES.—Holotype, allotype, and morphotype,

USNM 131533, 131534, 131535 ($ I, 9, a II);

paratypes, J. F. Fitzpatrick, Jr.

TYPE-LOCALITY.—Shutispear Creek, 2.7 miles

south of State Route 8 on State Route 9, Calhoun

County, Mississippi.

RANGE.—Known from only two localities in the

Yalobusha River drainage in Calhoun County,

Mississippi.

HABITAT.—Streams.

Procambarus (Pennides) natchitochae Penn

FIGURE 264

Procambarus natchitochae Penn, 1953d:5, figs. 8, 9, 15-19.

Procambarus (Pennides) natchitochae.—Hobbs, 1972a: 10, figs.

2k, 14a-g.

TYPES.—Holotype, allotype, and morphotype,

USNM 93649, 93650, 93651, ($ I, 9, 3 II); paratypes, USNM, AMNH, ANSP, TU.

TYPE-LOCALITY.—Tributary of Spring Creek at

Melder, on State Route 85, Rapides Parish, Louisiana.

RANGE.—Tributaries of Bayou Teche, Red, and

Calcasieu rivers in Louisiana, Arkansas, and Texas.

HABITAT.—Streams.

Procambarus (Pennides) ouachitae Penn

FIGURE 276

Procambarus ouachitae Penn, 1956:109, figs. 1-17.

Procambarus quachitae.—Penn, 1956:113 [erroneous spelling].

Procambarus ouichitae.—Hobbs, 1968b:K26 [erroneous spelling]-

Procambarus (Pennides) ouachitae.—Hobbs, 1972a: 10.

TYPES.—Holotype, allotype, and morphotype,

USNM 99222, 99223, 99224 («$ I, 9 , $ 11); paratypes, USNM, ANSP, AMNH, CM.

TYPE-LOCALITY.—Tributary of the South Fork

of the Saline River, 2.4 miles north of U.S. Highway 70 on a local road 6.5 miles northeast of Hot

Spring (7.7 miles south-southwest of Owensville),

Garland County, Arkansas.

RANGE.—Ouachita and Arkansas river systems in

southwestern Arkansas.

HABITAT.—Streams.

Procambarus (Pennides) penni Hobbs

FIGURE 272

Procambarus penni Hobbs, 1951b:273, figs. 1-11.

Procambarus (Pennides) penni.—Hobbs, 1972a: 10.

TYPES.—Holotype, allotype, and morphotype,

USNM 91662, 91663, 91664 (4 I, 9, & II); paratypes, TU, USNM.

TYPE-LOCALITY.—Talisheek Creek, at Talisheek,

St. Tammany Parish, Louisiana.

RANGE.—Pearl and Pascagoula drainage systems

in eastern Louisiana and south-central Mississippi.

HABITAT.—Streams.

Procambarus (Pennides) raneyi Hobbs

FIGURE 277

Procambarus raneyi Hobbs, 1953b:412, figs. 1-13.

Procambarus (Pennides) raneyi.—Hobbs, 1972a: 10.

TYPES.—Holotype, allotype, and morphotype,

USNM 95124, 95125, 95126 (s I, 9, <J II); paratypes, MCZ, TU, USNM.

TYPE-LOCALITY South fork of the Broad River,

1 mile south of Carlton on the Oglethorpe-Madison

county line, Georgia.

RANGE Savannah River drainage in Georgia

and South Carolina and headwaters of the Ocmulgee River in De Kalb County, Georgia.

HABITAT.—Streams.
64 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Procambarus (Pennides) spiculifer (LeConte)

FIGURE 274

Astacus spiculifer LeConte, 1856:401.

Cambarus spiculifer.—Hagen, 1870:48, figs. 59-62, 147.

Cambarus (Cambarus) spiculifer.—Ortmann, 1905c: 101.

Cambarus (Ortmannicus) spiculifer.—Fowler, 1912:341 [by

implication].—Creaser, 1934b:4 [by implication].

Procambarus spiculifer.—Hobbs, 1942a:342 [by implication];

1942b: 119, figs. 131-135.

Procambarus (Pennides) spiculifer.—Hobbs, 1972a: 10.

TYPES.—Syntypes, ANSP 316 ($11), MCZ 172

(2«jl, 4? , 2<$juv., 3$juv.), MCZ 3376 ($ I),

USNM 4962 (specimens decayed), MHNP ($1,

TYPE-LOCALITY.—Athens, Clarke County, Georgia.

RANGE.—Western Alabama eastward to the Altamaha River, Georgia.

HABITAT.—Lotic situations.

Procambarus (Pennides) suttkusi Hobbs

FICURE 263

Procambarus suttkusi Hobbs, 1953a: 173, pi. 9.

Procambarus suitkuzi.—Hobbs, 1968b:Kll [erroneous spelling]-

Procambarus (Pennides) suttkusi.—Hobbs, 1972a: 10.

TYPES.—Holotype, allotype, and morphotype,

USNM 93730, 93731, 93732 («jl, $, $ II); paratypes, USNM, TU.

TYPE-LOCALITY.—Tributary of Claybank Creek,

2 miles west of Ozark, Dale County, Alabama.

RANGE.—Choctawhatchee drainage system in

Alabama and northern Florida.

HABITAT.—Streams.

Procambarus {Pennides) versutus (Hagen)

FIGURE 267

Cambarus versutus Hagen, 1870:51, figs. 55-58, 150.

Cambarus (Cambarus) versutus.—Ortmann, 1905c: 101.

Cambarus (Ortmannicus) versutus.—Fowler, 1912:341 [by implication].—Creaser, 1934b:4 [by implication].

Procambarus versutus—Hobbs, 1942a:342 [by implication];

1942b: 126, figs. 136-140.

Procambarus (Pennides) versutus.—Hobbs, 1972a: 10.

TYPES—Syntypes, MCZ 190 (3 <? II, 9 ), USNM

4963 (£ I), MHNP (2 specimens), AMS (lost).

TYPE-LOCALITY.—Spring Hill, Mobile County,

Alabama.

RANGE.—From western Alabama to the Apalachicola River in Florida.

HABITAT.—Streams.

Procambarus (Pennides) vioscai Penn

FIGURE 271

Procambarus vioscai Penn, 1946:27, fig. 1.

Procambarus vioscae.—Williams, 1954:819 [erroneous spelling] •

Procambarus (Pennides) vioscai.—Hobbs, 1972a: 10.

TYPES.—Holotype, USNM 79925 ( $ I); allotype,

USNM 79931 (9); paratypes, USNM, Alabama

Mus. Nat. Hist., TU.

TYPE-LOCALITY.—Big Creek at Fishville, about

3 miles east of Pollock, Grant Parish, Louisiana.

RANGE.—Tributaries of the Red River in Arkansas and Louisiana eastward to the Pascagoula River

in Mississippi.

HABITAT.—Streams.

Subgenus Procambarus Ortmann

Procambarus Ortmann, 1905d:437. [Type-species by subsequent designation (Fowler, 1912:340), Cambarus Digueti

Bouvier, 1897:225. Gender: masculine.]

REMARKS.—Hobbs (1942a:341; 1972a: 11) erred

in citing himself as having designated the typespecies of Procambarus. See above.

Procambarus (Procambarus) digueti (Bouvier)

FIGURE 278

Cambarus Digueti Bouvier, 1897:225.

Cambarus carinatus Faxon, 1898:648, pi. 63. [Type, USNM

17699 (£ I). Type-locality, Guadalajara, Mexico.]

Cambarus (Cambarus) digueti.—Ortmann, 1905c:98, 101.

Cambarus (Procambarus) digueti.—Ortmann, 1905d:438.

Cambarus digniti.—Mctcalf, 1929:11 [erroneous spelling].

Procambarus digueti.—Hobbs, 1942a:342 [by implication].—

Villalobos, 1946:220, pi. 1.

Procambarus (Procambarus) digueti.—Hobbs, 1972a: 11, figs.

11, 3a, 15a-g.

TYPES.—Syntypes, USN M 30579 (2 <J I, 9) ,

MHN P (8<5, 69) , C M 74.483.

TYPE-LOCALITY.—Affluents of Ri o Santiago, Jalisco, Mexico.
NUMBER 166 65

RANGE.—Pacific slope in states of Jalisco and

Michoacan, Mexico.

HABITAT.—Streams.

Subgenus Remoticambarus Hobbs

Remoticambarus Hobbs, 1972a: 11. [Type-species by original

designation, Procambarus pecki Hobbs, 1967b:2. Gender:

masculine.]

Procambarus (Remoticambarus) pecki Hobbs

FIGURE 279

Procambarus pecki Hobbs, 1967b:2, figs. 1-12.

Procambarus (Remoticambarus) pecki Hobbs, 1972a: 11, figs.

2m, 16a-g.

TYPES.—Holotype, allotype, and morphotype,

USNM 117684, 117685, 117686 ($ I, 9, $ II);

paratypes, MCZ, TU, USNM.

TYPE-LOCALITY.—McKinney Pit Cave, about 2.5

miles west of Tuscumbia, Colbert County, Alabama (Sec. 10, T. 4 S., R. 12 W.).

RANGE.—Tennessee drainage system in Colbert,

Lauderdale, and Morgan counties, Alabama.

HABITAT.—Subterranean streams and pools.

Subgenus Scapulicambarus Hobbs

Scapulicambarus Hobbs, 1972a: 11. [Type-species by original

designation, Cambarus clarkii paeninsulanus Faxon, 1914:

369. Gender: masculine.]

Procambarus (Scapulicambarus) clarkii (Cirard)

FIGURE 283

Cambarus Clarkii Girard, 1852:91 Hagen, 1870:39, figs. 7-10,

99, 100, 133, 134, 142, pi. 4.

Cambarus clarkii.—Faxon, 1898:644.

Cambarus (Cambarus) clarki.—Ortmann, 1905c: 102.

Cambarus (Ortmannicus) clarkii.—Fowler, 1912:341 [by implication].

Cambarus clarkii clarkii.—Faxon, 1914:369 [by implication].

Procambarus clarkii.—Hobbs, 1942a:342 [by implication];

1962b:273, figs. 1-9.

Procambarus clarki.—Penn and Hobbs, 1958:466, figs. 2, 21,

36, 49, 65.

Procambarus ctarcii.—Unestam, 1969:203 [erroneous spelling].

Procambarus (Scapulicambarus) clarkii.—Hobbs, 1972a: 12.

TYPES.—Destroyed in Chicago fire, 1871 (Faxon,

1914:414).

TYPE-LOCALITY.—Between San Antonio and El

Paso del Norte, Texas.

RANGE.—Northern Mexico to Escambia County,

Florida, and north to southern Illinois.

HABITAT.—Lentic and lotic situations and burrows.

Procambarus (Scapulicambarus) howellae Hobbs

FIGURE 281

Procambarus howellae Hobbs, 1952b: 167, figs. 1, 4, 5, 8-14.

Procambarus (Scapulicambarus) howellae.—Hobbs, 1972a: 12.

TYPES.—Holotype and allotype, USNM 93158

(<5 I, $); morphotype, USNM 93159 ($ II); paratypes, MCZ, USNM.

TYPE-LOCALITY.—Drainage ditch on campus of

Wesleyan College, Rivoli, Bibb County, Georgia.

RANGE.—Lower piedmont and upper coastal

plain of Georgia between the Ocmulgee and Ogeechee rivers.

HABITAT.—Lotic and lentic situations.

Procambarus (Scapulicambarus) okaloosae

Hobbs

FIGURE 282

Procambarus okaloosae Hobbs, 1942b: 100, figs. 101-105, 315—

324.

Procambarus (Scapulicambarus) okaloosae.—Hobbs, 1972a; 12.

TYPES—Holotype, USNM 81282 (<$ I); allotype

and "morphotype," USNM 81283 (5 , ill) ; paratypes, MCZ, USNM.

TYPE-LOCALITY.—Roadside ditch at intersection

of U.S. Highway 90 and State Route 4, Milligan,

Okaloosa County, Florida.

RANGE.—Perdido, Escambia, and Yellow drainage systems in Alabama and Florida.

HABITAT.—Lentic and lotic situations.

Procambarus (Scapulicambarus) paeninsulanus

(Faxon)

FIGURE 280

Cambarus clarkii paeninsulanus Faxon, 1914:369.

Cambarus clarkipaenensulanus.—Hobbs, 1937:154 [erroneous

spelling].

Procambarus clarkii paeninsulanus.—Hobbs, 1942a:342 [by

implication].
66 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Procambarus paeninsulanus.—Hobbs, 1942b: 104, figs. 106—110.

Cambarus paeninsulanus.—Hobbs, 1972a: 12 [erroneous combination].

Procambarus (Scapulicambarus) paeninsulanus.—Hobbs,

1972a: 12, figs. le,f, 2n, 3b, 17a-g.

TYPES.—Holotype, MCZ 3530 ( $ II); paratypes,

MCZ, USNM.

TYPE-LOCALITY.—Three miles below Horse

Landing, St. Johns River, Putnam County, Florida.

RANGE.—South Georgia, and Florida east of the

Choctawhatchee drainage south to Hillsborough

County.

HABITAT.—Lentic and lotic situations, and burrows (tertiary).

Procambarus (Scapulicambarus) troglodytes

(LeConte)

FIGURE 284

Asiacus troglodytes LeConte, 1856:400.

Astacus fossarum LeConte, 1856:401. [Syntypes, MCZ 3377

($I dry), ANSP 314 (9). Type-locality, ditches, lower

Georgia.]

Astacus maniculatus LeConte, 1856:401. [Types not extant.

Type-locality, ditches, lower Georgia.]

Cambarus troglodytes.—Hagen, 1870:41, figs. 11-14, 141.

Cambarus troglodytus.—Hagen, 1870:33 [erroneous spelling].

Cambarus (Cambarus) troglodytes.—Qrtmann, 1905c:102.

Cambarus (Ortmannicus) troglodytes.—Fowler, 1912:341 [by

implication].—Creaser, 1934b:4 [by implication].

Procambarus troglodytes.—Hobbs, 1942a:342 [by implication].

Procambarus (Scapulicambarus) troglodytes.—Hobbs, 1972a:

12.

TYPES.—Syntypes, MCZ 3375 ($ I dry), ANSP

4175 (fragments).

TYPE-LOCALITY.—Rice fields in Georgia.

RANGE.—Between the Altamaha and Pee Dee

rivers, in Georgia and South Carolina.

HABITAT.—Lentic and lotic situations, and burrows (tertiary burrower).

Subgenus Tenuicambarus Hobbs

Tenuicambarus Hobbs, 1972a: 12. [Type-species by original

designation, Procambarus tenuis Hobbs, 1950b: 194. Gender:

masculine.]

Procambarus (Tenuicambarus) tenuis Hobbs

FIGURE 285

Procambarus tenuis Hobbs, 1950b: 194, figs. 1-12.

Procambarus (Tenuicambarus) tenuis—Hobbs, 1972a: 12, figs.

2o, 3d, 18a-g.

TYPES.—Holotype, allotype, and morphotype,

USNM 90390 (s I, $, ^11); paratypes, USNM.

TYPE-LOCALITY.—Six miles east of Page and just

west of Oklahoma state line, Le Flore County,

Oklahoma.

RANGE.—Le Flore, Pushmataha, and Pittsburg

counties, Oklahoma, and Montgomery and Polk

counties, Arkansas.

HABITAT.—Springs and clear, cold streams.

Subgenus Villalobosus Hobbs

Villalobosus Hobbs, 1972a: 12. [Type-species by original designation, Paracambarus riojae [sic] Villalobos, 1944a: 161. Gender: masculine.]

Procambarus (Villalobosus) contrerasi (Creaser)

FIGURE 289

Cambarus (Cambarus) contrerasi Creaser, 1931a:l, figs. 1-6.

Cambarus contrerasi.—Creaser, 1931a: 1.

Procambarus contrerasi.—Hobbs, 1942a:342.—Villalobos,

1950:373, pis. 2, 3.

Procambarus (Villalobosus) contrerasi.—Hobbs, 1972a: 12.

TYPES.—Holotype, USNM 98353 (<5 I); paratype, USNM.

TYPE-LOCALITY.—Tributary of the Rio Cazones

near Agua Fria, 12 miles south of Miahuapan,

State of Puebla, Mexico.

RANGE.—Tributaries of the Rio Cazones and

the Rio Tecolutla in the vicinity of Zihuateutla

and Agua Fria, Puebla, Mexico.

HABITAT.—Streams.

Procambarus (Villalobosus) erichsoni Villalobos

FIGURE 288

Procambarus erichsoni Villalobos, 1950:384, pis. 5-7.

Procambarus (Villalobosus) erichsoni.—Hobbs, 1972a: 12.

TYPES.—Holotype, allotype, and morphotype,

IBM 52048(048) ($ I, $, $ II); paratypes, IBM,

USNM.

TYPE-LOCALITY.—"Arroyos Puendo, Mamay y

Bojoy, Tenango de Doria, Hidalgo," Mexico.

RANGE.—Tributaries of the Rio Pantepec (to
NUMBER 166 67

Rio Tuxpan) in the vicinity of Tenango de Doria,

Hidalgo, Mexico.

HABITAT.—Streams.

Procambarus (Villalobosus) hoffmanni

(Villalobos)

FIGURE 287

Paracambarus hoffmanni Villalobos, 1944a: 169, pi. 2.

Procambarus hoffmanni.—Villalobos, 1950:411.

Procambarus Hoffmanni Hobbs, 1962a:279 [lapsus calami].

Procambarus (Villalobosus) hoffmanni.—Hobbs, 1972a: 12.

TYPES.—Holotype, allotype, and "morphotype,"

IBM 100043(013) (3 I, 9, 3 II); paratypes, IBM,

USNM.

TYPE-LOCALITY.—"Vertedor de Demasia de la

Presa de Necaxa, Necaxa," Puebla, Mexico. (Villalobos, 1955:107.)

RANGE.—Tributaries of the Rio Tecolutla and

Rio Nautla in Puebla and Veracruz, Mexico.

HABITAT.—Streams.

Procambarus (Villalobosus) hortonhobbsi

Villalobos

FIGURE 290

Procambarus hortonhobbsi Villalobos, 1950:402, pis. 10, 11.

Procambarus (Villalobosus) hortonhobbsi Hobbs, 1972a: 12.

TYPES.—Holotype, allotype, and morphotype,

IBM 21249(085) ($ I, 9, £ II); paratypes, IBM,

USNM.

TYPE-LOCALITY.—El Coyular, 7 km northeast of

La Union, Municipio de Zihuateutla, Puebla,

Mexico.

RANGE.—Known only from the type-locality.

HABITAT.—Streams.

Procambarus (Villalobosus) riojai (Villalobos)

FIGURE 286

Paracambarus riojae Villalobos, 1944a: 161, pi. 1.

Procambarus riojae.—Villalobos, 1950:401.

Procambarus riojai.—Hobbs, 1971d:3.

Procambarus (Villalobosus) riojai.—Hobbs, 1972a: 12, figs. 2p,

3c, 19a-g.

TYPES.—Holotype, allotype, and "morphotype,"

IBM 50045(027) (£l, 9, ill); paratypes, IBM,

USNM.

TYPE-LOCALITY.—"Huauchinango, Estado de

Puebla, riachuelos tributarios del rio Necaxa,"

Mexico.

RANGE.—Headwater tributaries of the Rio Cazones and Rio Tecolutla in Hidalgo and Puebla,

Mexico.

HABITAT.—Streams.

Procambarus (Villalobosus) teziutlanensis

(Villalobos)

FIGURE 292

Paracambarus teziutlanensis Villalobos, 1947a:240, pi. 2; pi.

3: figs. 5-9.

Procambarus teziutlanensis.—Villalobos, 1950:372.

Procambarus tziutlanensis Hobbs, 1971d:12 [erroneous

spelling].

Procambarus (Villalobosus) teziutlanensis.—Hobbs, 1972a: 12.

TYPES.—Holotype, allotype, and morphotype,

IBM 121245(029) ($ I, 9, S II); paratypes, IBM,

USNM.

TYPE-LOCALITY.—"Chignautla, 5 km al Oriente

de Teziutlan, Estado de Puebla," Mexico.

RANGE.—Headwater tributaries of the Rfo

Nautla in Puebla and Veracruz, Mexico.

HABITAT.—Streams.

Procambarus (Villalobosus) tlapacoyanensis

(Villalobos)

FIGURE 293

Paracambarus tlapacoyanensis Villalobos, 1947b:537, pis. 1, 2.

Procambarus tlapacoyanensis.—Villalobos, 1950:372.

Procambarus (Villalobosus) tlapacoyanensis.—Hobbs, 1972a:

12.

TYPES.—Holotype, allotype, and morphotype,

IBM 51947(031) (£ I, 9, $ II); paratypes, IBM,

USNM.

TYPE-LOCALITY.—Canada de Tomata, Tlapacoyan, Veracruz, Mexico.

RANGE.—Tributaries of the Rio Nautla in the

vicinity of Tlapacoyan, Veracruz, Mexico.

HABITAT.—Streams.

Procambarus (Villalobosus) zihuateutlensis

Villalobos

FIGURE 291

Procambarus zihuateutlensis Villalobos, 1950:394, pis. 8, 9.

Procambarus (Villalobosus) zihuateutlensis.—Hobbs, 1972a:

12.
68 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

TYPES.—Holotype, allotype, and morphotype,

IBM 21149(080) ($ I, $, $ II); paratypes, IBM.

TYPE-LOCALITY.—Arroyo de Tlatentiloyan, Los

Estajos, Municipio de Zihuateutla, Puebla, Mexico.

RANGE.—Headwater tributaries of the Rio Tecolutla in the vicinity of Zihuateutla, Puebla,

Mexico.

HABITAT.—Streams.

Genus Troglocambarus Hobbs

Troglocambarus Hobbs, 1942a:345. [Type-species by original

designation, Troglocambarus tnaclanei Hobbs, 1942a:345.

Gender: masculine.]

Troglocambarus maclanei Hobbs

FIGURE 294

Troglocambarus maclanei Hobbs, 1942a:345, pis. 1, 2; 1971c:

122, figs. 17a-g, 19a.

TYPES.—Holotype, allotype, and "morphotype,"

USNM 79385, 79386, 79387 (<j I, $, $ II); paratypes, MCZ, USNM.

TYPE-LOCALITY.—Squirrel Chimney, a small cave

about 11 miles northwest of Gainesville, Alachua

County, Florida (Sec. 21, T. 9 S., R. 18 E.).

RANGE.—Citrus and Hernando to Alachua counties, Florida.

HABITAT.—Subterranean waters.

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Fitzpatrick, J. F., Jr., and Horton H. Hobbs, Jr.

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22(1187):121-123, 2 figures.

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1902a. Observations on the Crustacean Fauna of the Region about Mammoth Cave, Kentucky. Proceedings

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1902b. Observations on the Crustacean Fauna of Nickajack

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1902c. On the Proper Application of the Name Cambarus

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1937. Some Florida Crawfishes and Their Habitat Distribution. [Abstract.] Proceedings of the Florida

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1938a. A New Crayfish from Florida. Journal of the

Washington Academy of Sciences, 28(2):6I-65, 11

figures.

1938b. Two New Crawfishes from Florida, Cambarus hubbelli and Cambarus acherontis pallidus. [Abstract.]

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2:90-91.

1940a. Seven New Crayfishes of the Genus Cambarus from

Florida, with Notes on Other Species. Proceedings

of the United States National Museum, 89(3097):

387-423, figures 14-22.

1940b. A New Crayfish from South Carolina. Charleston

Museum Leaflet, 14:3-7, 10 figures.

1941a. A New Crayfish from San Luis Potosf, Mexico

(Decapoda, Astacidae). Zoologica, 26(1): 1-4, 1 figure.

1941b. Three New Florida Crayfishes of the Subgenus

Cambarus (Decapoda, Astacidae). American Midland Naturalist, 26(1): 110-121, 2 plates.

1942a. A Generic Revision of the Crayfishes of the Subfamily Cambarinae (Decapoda, Astacidae) with the

Description of a New Genus and Species. American

Midland Naturalist, 28(2): 334-357, 3 plates.

1942b. The Crayfishes of Florida. University of Florida

Publications, Biological Science Series, 3(2): v + 179

pages, 24 plates.

1942c. On the First Pleopod of the Male Cambari (Decapoda, Astacidae). Proceedings of the Florida Academy of Sciences, 1940, 5:55-61, 2 plates.

1943a. Two New Crayfishes from the Panhandle of Florida

(Decapoda, Astacidae). Proceedings of the Florida

Academy of Sciences, 6(l):49-58, 32 figures.

1943b. Two New Crayfishes of the Genus Procambarus

from Mexico (Decapoda, Astacidae). Lloydia, 6:

198-206, 2 plates.

1944. [Identification of crayfishes.] In C. Clayton Hoff,

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327-357, 31 figures.
72

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

1945a. The Subspecies and Intergrades of the Florida Burrowing Crayfish, Procambarus rogersi (Hobbs).

Journal of the Washington Academy of Sciences, 1952a.

35(8):247-26O, 34 figures.

1945b. Two New Species of Crayfishes of the Genus Cambarellus from the Gulf Coastal States, with a Key

to the Species of the Genus (Decapoda, Astacidae). 1952b.

American Midland Naturalist, 34 (2):466-474, 26

figures.

1947a. Two New Crayfishes of the Genus Procambarus

from Georgia, with Notes on Procambarus pubescens

(Faxon) (Decapoda, Astacidae). Quarterly Journal 1952c.

of the Florida Academy of Science, 9(1): 1-18, 32

figures.

1947b. A Key to the Crayfishes of the Pictus Subgroup of

the Genus Procambarus, with the Description of a 1953a.

New Species from South Carolina (Decapoda,

Astacidae). Florida Entomologist, 30(3):25-31, 12

figures.

1948a. On the Crayfishes of the Limosus Section of the 1953b.

Genus Orconectes (Decapoda, Astacidae). Journal

of the Washington Academy of Sciences, 38(1): 14-21,

28 figures.

1948b. Two New Crayfishes of the Genus Orconectes from

Arkansas, with a Key to the Species of the Hylas 1953c.

Group (Decapoda, Astacidae). American Midland

Naturalist, 39(1): 139-150, 30 figures.

1948c. A New Crayfish of the Genus Orconectes from

Southern Tennessee (Decapoda, Astacidae). Pro- 1954.

ceedings of the Biological Society of Washington,

61(15):85-91, 9 figures.

1948d. A New Crayfish of the Genus Cambarus from

Texas, with Notes on the Distribution of Cambarus 1955a.

fodiens (Cottle). Proceedings of the United States

National Museum, 98(3230):223-231, figure 17.

1949a. A New Crayfish of the Genus Orconectes from the 1955b.

Nashville Basin in Tennessee, with Notes on the

Range of Orconectes compressus (Faxon) (Decapoda,

Astacidae). Proceedings of the Biological Society of

Washington, 62 (7): 17-24, 10 figures. 1956a.

1949b. [Identifications of crayfishes.] In Perry C. Holt, A

Comparative Study of the Reproductive Systems of

Xironogiton instabilius instabilius Moore and Cambarincola philadelphica Leidy (Annelida, Oligo- 1956b.

chaeta, Branchiobdellidae). Journal of Morphology,

84(3):535-571, 27 figures.

1950a. A New Crayfish of the Genus Cambarellus from

Texas (Decapoda, Astacidae). Proceedings of the 1956c.

Biological Society of Washington, 63(18):89-96, 17

figures.

1950b. A New Crayfish of the Genus Procambarus from

Oklahoma and Arkansas (Decapoda, Astacidae).

Journal of the Washington Academy of Sciences, 1958a.

40(6): 194-198, 12 figures.

1951a. A New Crayfish of the Genus Orconectes from

Southeastern Virginia (Decapoda, Astacidae). Vir- 1958b.

ginia Journal of Science, 2 (2): 122-128, 10 figures.

1951b. A New Crayfish of the Genus Procambarus from

Louisiana with a Key to the Species of the Spiculifer 1958c.

Group. Journal of the Washington Academy of

Sciences, 41(8): 272-276, 11 figures.

A New Crayfish from Alabama, with Notes on

Procambarus lecontei (Hagen). Proceedings of the

United States National Museum, 102(3297):209-219,

figures 81, 82.

A New Crayfish of the Genus Procambarus from

Georgia with a Key to the Species of the Clarkii

Subgroup (Decapoda, Astacidae). Quarterly Journal

of the Florida Academy of Sciences, 15(3): 165-174,

14 figures.

A New Albinistic Crayfish of the Genus Cambarus

from Southern Missouri with a Key to the Albinistic

Species of the Genus (Decapoda, Astacidae). American Midland Naturalist, 48 (3):689-693, 8 figures.

A New Crayfish of the Genus Procambarus from

Alabama and Florida (Decapoda, Astacidae). Proceedings of the Biological Society of Washington,

66(22): 173-178, plate 9.

On the Ranges of Certain Crayfishes of the Spiculifer

Group of the Genus Procambarus, with the Description of a New Species (Decapoda, Astacidae).

Journal of the Washington Academy of Sciences,

43(12):412-417, 13 figures.

Two New Crayfishes from the Highland Rim in

Tennessee (Decapoda, Astacidae). Journal of the

Tennessee Academy of Science, 28(l):2O-27, 18

figures.

A New Crayfish from the Upper Coastal Plain of

Georgia (Decapoda, Astacidae). Quarterly Journal

of the Florida Academy of Sciences, 17(2): 110-118,

13 figures.

A New Crayfish of the Genus Cambarus from Mississippi. Proceedings of the Biological Society of

Washington, 68(15):95-IOO, 11 figures.

Ostracods of the Genus Entocythere from the New

River System in North Carolina, Virginia, and West

Virginia. Transactions of the American Microscopical Society, 74(4):325-333, 10 figures.

A New Crayfish of the Genus Procambarus from

South Carolina (Decapoda, Astacidae). Journal of

the Washington Academy of Sciences, 46(4): 117-121,

17 figures.

A New Crayfish of the Genus Cambarus from North

Carolina and South Carolina (Decapoda, Astacidae).

Journal of the Elisha Mitchell Scientific Society,

72(l):61-67, 11 figures.

A New Crayfish of the Extraneus Section of the

Genus Cambarus with a Key to the Species of the

Section (Decapoda, Astacidae). Proceedings of the

Biological Society of Washington, 69(18): 115-121, 11

figures.

Two New Crayfishes of the Genus Procambarus

from South Carolina. Journal of the Washington

Academy of Sciences, 48(5): 160-168, 24 figures.

Two New Crayfishes of the Genus Procambarus

from South Carolina and Georgia. Notulae Naturae,

307: 10 pages, 26 figures.

The Evolutionary History of the Pictus Group of
NUMBER 166 73

the Crayfish Genus Procambarus (Decapoda, Astacidae). Quarterly Journal of the Florida Academy

of Sciences, 21(1):71-91, 20 figures.

1959. Pages 883-901, figures 31.18-31.41, in W. T. Edmondson, Fresh-Water Biology. Second edition. New

York: John Wiley & Sons.

1962a. Notes on the Affinities of the Members of the

Blandingii Section of the Crayfish Genus Procambarus (Decapoda, Astacidae). Tulane Studies

in Zoology, 9(5):273-293, 72 figures.

1962b. La Presencia de Procambarus clarkii (Girard) en los

Estados de Chihuahua y Sonora, Mexico (Decapoda,

Astacidae). Anales del Instituto de Biologia, Universidad Nacional Autonoma de Mexico, 33(1, 2):

273-276, 9 figures.

1964. A New Cave-Dwelling Crayfish from the Greenbrier

Drainage System, West Virginia (Decapoda, Astacidae). Proceedings of the Biological Society of

Washington, 77(21): 189-194, 10 figures.

1965. A New Crayfish of the Genus Cambarus from Tennessee with an Emended Definition of the Genus

(Decapoda, Astacidae). Proceedings of the Biological Society of Washington, 78(33):265-273, 12 figures.

1966a. A New Crayfish from Alabama with Observations

on the Cristatus Section of the Genus Cambarus

(Decapoda, Astacidae). Proceedings of the Biological

Society of Washington, 79(15): 109-116, 10 figures.

1966b. An Illustrated Key to the Species of the Genus

Ankylocythere with a Description of a New Species

from Louisiana (Ostracoda, Entocytheridae). Proceedings of the Louisiana Academy of Sciences, 29:

67-75, 18 figures.

1967a. The Current Status of the Crayfishes Listed by

Girard (1852) in His "A Revision of the North

American Astaci. . . ." Crustaceana, (12)2:124-132.

1967b. A New Crayfish from Alabama Caves with Notes on

the Origin of the Genera Orconectes and Cambarus

(Decapoda: Astacidae). Proceedings of the United

States National Museum, 123(3621): 1-17, 21 figures.

1968a. Two New Crayfishes of the Genus Cambarus from

Georgia, Kentucky, and Tennessee (Decapoda, Astacidae). Proceedings of the Biological Society of

Washington, 81(31):261-274, 22 figures.

1968b. Crustacea: Malacostraca. Pages K1-K36, 32 figures,

in Fred K. Parrish, Keys to Water Quality Indicative Organisms (Southeastern United States). Federal Water Pollution Control Administration, Department of the Interior.

1969a. Procambarus villalobosi, un Nuevo Cambarino dc

San Luis Potosi, Mexico (Decapoda, Astacidae).

Anales del Instituto de Biologia, Universidad

Nacional Autonoma de Mexico, Serie Ciencias del

Mar y Limnologia, 38(l):41-46, 11 figures.

1969b. On the Distribution and Phylogcny of the Crayfish Genus Cambarus. In Perry C. Holt, Richard L.

Hoffman, and C. Willard Hart, Jr., The Distributional History of the Biota of the Southern Appalachians, Part I: Invertebrates. Virginia Polytechnic

Institute, Research Division Monograph, 1:93-178,

20 figures.

1969c. Two New Species of the Crayfish Genus Procambarus (Decapoda, Astacidae) with Keys to the Members of the Spiculifer Group. Proceedings of the

Biological Society of Washington, 83 (24):329-348,

38 figures.

1970a. A New Crayfish from the Nashville Basin, Tennessee. Proceedings of the Biological Society of

Washington, 83(14): 161-169, 1 figure.

1970b. New Crayfishes of the Genus Cambarus from Tennessee and Georgia (Decapoda, Astacidae). Proceedings of the Biological Society of Washington,

83(23): 241-259, 4 figures.

1971a. A New Crayfish of the Genus Procambarus from

Mississippi (Decapoda: Astacidae). Proceedings of

the Biological Society of Washington, 83(40): 459-

468, 1 figure.

1971b. New Crayfishes of the Genus Procambarus from

Alabama and Texas (Decapoda, Astacidae). Proceedings of the Biological Society of Washington,

84(ll):81-94, 2 figures.

1971c. A New Troglobitic Crayfish from Florida. Quarterly Journal of the Florida Academy of Sciences,

34(2): 114-124, 19 figures.

1971d. The Entocytherid Ostracods of Mexico and Cuba.

Smithsonian Contributions to Zoology, 81: 55 pages,

31 figures.

1972a. The Subgenera of the Crayfish Genus Procambarus

(Decapoda: Astacidae). Smithsonian Contributions

to Zoology, 117: 22 pages, 20 figures.

1972b. Crayfishes (Astacidae) of North and Middle America,

Identification Manual 9, x + 173 pages, 115 figures.

In Biota of Freshwater Ecosystems. United States

Environmental Protection Agency, Water Pollution

Research Control Series.

1974. Synopsis of the Families and Genera of Crayfishes

(Crustacea, Decapoda). Smithsonian Contributions

to Zoology, 164: 32 pages, 27 figures.

Hobbs, Horton H., Jr., and Thomas C. Barr, Jr.

1960. The Origins and Affinities of the Troglobitic Crayfishes of North America (Decapoda, Astacidae), I.

The Genus Cambarus. American Midland Naturalist, 64(1): 12-33, 57 figures.

1972. Origins and Affinities of the Troglobitic Crayfishes

of North America (Decapoda: Astacidae), II. Genus

Orconectes. Smithsonian Contributions to Zoology,

105: 84 pages, 16 figures.

Hobbs, Horton H., Jr., and M. S. Bedinger

1964. A New Troglobitic Crayfish of the Genus Cambarus

(Decapoda, Astacidae) from Arkansas with a Note

on the Range of Cambarus cryptodytes Hobbs.

Proceedings of the Biological Society of Washington,

77(3):9-15, 11 figures.

Hobbs, Horton H., Jr., and Raymond W. Bouchard

1973. A New Crayfish from the Cumberland River System with Notes on Cambarus carolinus (Erichson).

Proceedings of the Biological Society of Washington,

86(5):41-68, 4 figures.
74

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Hobbs, Horton H., Jr., and Martha R. Cooper

1972. A New Troglobitic Crayfish from Oklahoma (Decapoda: Astacidae). Proceedings of the Biological

Society of Washington, 85(3):49-56, 1 figure.

Hobbs, Horton H., Jr., and J. F. Fitzpatrick, Jr.

1962. A New Crayfish of the Propinquus Group of the

Genus Orconectes from the Ohio Drainage System

in West Virginia (Decapoda, Astacidae). Proceedings of the Biological Society of Washington, 75 (30):

207-214, 15 figures.

1970. A New Crayfish of the Genus Falticambarus from

Tennessee (Decapoda, Astacidae). Proceedings of

the Biological Society of Washington, 82(64):829-

836, 12 figures.

Hobbs, Horton H., Jr., and Edward T. Hall, Jr.

1969. New Crayfishes from Georgia (Decapoda, Astacidae).

Proceedings of the Biological Society of Washington,

82(21):281-294, 24 figures.

1972. A New Crayfish from the Tallapoosa River in

Georgia (Decapoda: Astacidae). Proceedings of the

Biological Society of Washington, 85(12): 151-161,

1 figure.

Hobbs, Horton H., Jr., and C. W. Hart, Jr.

1959. The Freshwater Decapod Crustaceans of the Apalachicola Drainage System in Florida, Southern

Alabama, and Georgia. Bulletin of the Florida

State Museum, 4(5): 145-191, 39 figures.

Hobbs, Horton H., Jr., and H. H. Hobbs III

1962. A New Crayfish of the Genus Cambarus from

Georgia (Decapoda, Astacidae). Proceedings of the

Biological Society of Washington, 75(5):41-45, 10

figures.

Hobbs, Horton H., Jr., Perry C. Holt, and Margaret Walton

1967. The Crayfishes and Their Epizootic Ostracod and

Branchiobdellid Associates of the Mountain Lake,

Virginia, Region. Proceedings of the United States

National Museum, 123(3602): 1-84, 22 figures.

Hobbs, Horton H., Jr., and Lewis J. Marchand

1943. A Contribution toward a Knowledge of the Crayfishes of the Reelfoot Lake Area. Journal of the

Tennessee Academy of Science, 18(l):6-35, 27 figures.

Hobbs, Horton H., Jr., and D. Bruce Means

1972. Two New Troglobitic Crayfishes (Decapoda, Astacidae) from Florida. Proceedings of the Biological Society of Washington, 84 (46): 393-409, 2 figures.

Hobbs, Horton H., Jr., and Frank O. Perkins

1967. A New Burrowing Crayfish from North Carolina

(Decapoda, Astacidae). Proceedings of the Biological Society of Washington, 80(21);141-146, 11 figures.

Hobbs, Horton H., Jr., and Charles S. Shoup

1947. Two New Crayfishes (Decapoda, Astacidae) from

the Obey River Drainage in Tennessee. Journal

of the Tennessee Academy of Science, 22(2): 138-145,

22 figures.

Hobbs, Horton H., Jr., and Alejandro Villalobos

1964. Los Cambarinos de Cuba. Anales del Instituto de

Biologia, Universidad Nacional Autdnoma de Mexico, 34(12): 307-366, 50 figures.

Hobbs, Horton H., Jr., and Margaret Walton

1957. Three New Crayfishes from Alabama and Mississippi (Decapoda: Astacidae). Tulane Studies in

Zoology, 5(3):39-52, 34 figures.

1958. Procambarus pearsei plumimanus, a New Crayfish

from North Carolina (Decapoda, Astacidae).

Journal of the Elisha Mitchell Scientific Society,

74(1):7-12, 13 figures.

1959. A New Crayfish of the Genus Procambarus from

Alabama (Decapoda, Astacidae). Proceedings of the

Biological Society of Washington, 72(10):39-44, 12

figures.

1960. A New Crayfish of the Genus Procambarus from

Southern Alabama (Decapoda, Astacidae). Proceedings of the Biological Society of Washington, 73(20):

123-129, 11 figures.

1966. Orconectes Juvenils (Hagen) in Mountain Lake,

Virginia: an Unplanned Experiment in Interspecific

Competition (Decapoda, Astacidae). Virginia Journal of Science, 17(3):136-14O.

Hobbs III, H. H.

1971. New Entocytherid Ostracods of the Genera Ankylocythere and Dactylocythere. Proceedings of the

Biological Society of Washington, 84(17): 137-146,

2 figures.

Hoff, C. Clayton

1943. Two New Ostracods of the Genus Entocythere and

Records of Previously Described Species. Journal

of the Washington Academy of Sciences, 33(9):276-

286, 2 figures.

Holt, Perry C.

1954. A New Branchiobdellid of the Genus Cambarincola

(Oligochaeta, Branchiobdellidae) from Virginia.

Virginia Journal of Science, 5(3): 168-172, 5 figures.

1960. The Genus Ceratodrilus Hall (Branchiobdellidae,

Oligochaeta), with the Description of a New Species.

Virginia Journal of Science, ll(2):53-77, 16 figures.

Holthuis, L. B.

1964. On the Status of the Two Allegedly European Crayfishes, Cambarus typhlobius Joseph, 1880, and Austropotamobius pallipes bispinosus Karaman, 1961

(Decapoda, Astacidae). Crustaceana, 7 (l):42-48,

plate 3.

Hovey, Horace C.

1882. Celebrated American Caverns, Especially Mammoth,

Wyandot, and Luray, together with Historical, Scientific and Descriptive Notices of Caves and Grottoes in other Lands; with Maps and Illustrations.

xii + 288 pages, illustrated. Cincinnati: Robert

Clarke and Company.

Huntsman, A. G.

1915. The Fresh-Water Malacostraca of Ontario. Contributions to Canadian Biology, 1911-1914:145-163,

13 figures.

Huxley, Thomas H.

1879. On the Classification and the Distribution of the

Crayfishes. Proceedings of the Zoological Society of

London, 1878:751-788, 7 figures.
NUMBER 166 75

Jackson, George F.

1942. Animal Life of Caverns. Bulletin of the National

Speleological Society, 4:3-7.

James, Hugo A.

1966. Range and Variations of Subspecies of Cambarus

longulus (Decapoda, Astacidae). Proceedings of the

United States National Museum, 119(3544): 1-24, 2

figures, 1 plate.

Jester, Douglas B.

1967. A New Crawfish of the Genus Orconectes from

New Mexico (Decapoda, Astacidae). American Midland Naturalist, 77(2):518-524, 12 figures.

Joseph, Gustav

1880. Demonstration eines neuen zur Familie der Astaciden gehorenden blinden Grottenkrebses aus

Krain. Schlesische gesellschaft fur vaterldndische

kultur, Breslau. Jahresbericht, 57:202.

1881. Erfahrungen im wissenschaftlichen Sammeln und

Beobachten der den Krainer Trofsteingrotten

eigenen Arthropoden. Berliner entomologische

zeitschrift, 25:233-282.

1882. Systematisches Verzcichniss der in der TropfsteinGrotten von Krain einheimischen Arthropoden

nebst Diagnosen der vom Verfasser entdeckten und

bisher noch nicht beschriebenen Arten. Schluss der

Abhandl.; Erfahrungen im wissenschaftlichen

Sammeln und Beobachten der den Krainer

Tropfsteingrottcn eigenen Arthropoden. Berliner

entomologische zeitschrift, 26:1-50.

Kamita, Tsuncichi

1970. Studies on the Fresh-Water Shrimps, Prawns and

Crawfishes of Japan. 213 pages. Matsue.

1973. Crayfishes Brought to Japan from Mexico. Zoological Magazine (Dobutsugaku Zasshi), 83(2): 123-126, 4

figures.

Kozloff, Eugene N.

1955. Two New Species of Entocy there (Ostracoda:

Cytheridae) Commensal on Pacifastacus Gambelii

(Girard). American Midland Naturalist, 53(1):

156-161, 24 figures.

Laguarda, Alfredo

1961. Contribucion al Estudio Com para tivo de la Formula

Branquial en la Familia Astacidae (Crustacea: Decapoda). 74 pages, 22 plates. Facultad de Ciencias,

Departmento dc Biologia, Univcrsidad Nacional

Autonoma de Mexico.

Latreille, P. A.

1802-1803. Histoire Naturelle, General el Particuliere des

Crustaces et des Insectes. 387 pages, plates 16-37.

Paris: F. Dufart.

LeConte, John

1856. Descriptions of New Species of Astactts from Georgia. Proceedings of the Academy of Natural Sciences of Philadelphia, 7:400-402.

Lockington, W. N.

1878. Remarks upon the Thalassinidea and Astacidae of

the Pacific Coast of North America, with Description of a New Species. Annals and Magazine of

Natural History, series 5, 2(10):299-304.

Lonnberg, Einar

1895. Cambarids from Florida. Bihang Till Koniglische

Svenska Vetenskaps-Akademiens Handlingar, 20(4):

3-14, figures l-5b.

Lyle, Clay

1938. The Crawfishes of Mississippi, with Special Reference to the Biology and Control of Destructive

Species. [Abstract.] Iowa State College Journal of

Science, 13:75-77.

Marlow, Guy

1960. The Subspecies of Cambarus diogenes. American

Midland Naturalist, 64(l):229-250, 8 figures.

Martens, Eduard von

1872. Ueber Cubanesche Crustacean—nach den Samtnlungen Dr. J. Grundlach's. Archiv fur Naturgeschichte, Berlin, 38:77-147, plates 4, 5.

Mason, John C.

1970a. Egg-Laying in the Western North American Crayfish, Pacifastacus trowbridgii (Stimpson) (Decapoda,

Astacidae). Crustaceana, 19 (1):37—44, 16 figures.

1970b. Copulatory Behavior of the Crayfish, Pacifastacus

trowbridgii (Stimpson). Canadian Journal of Zoology, 48(5):969-976, 7 figures.

Meek, S. E.

1894. A New Cambarus from Arkansas. American Naiuralist, 28(336): 1042-1043, 4 figures.

Meredith, W. G., and F. J. Schwartz

1960. Maryland Crayfishes. Maryland Department of

Research and Education, Educational Series, 46: 32

pages, 17 figures.

Merkle, E. Lynn

1969. Home Range of Crayfish Orconectes juiienalis.

American Midland Naturalist, 81(l):228-235, 5 figures.

Metcalf, Maynard M.

1929. Parasites and the Aid They Give in Problems of

Taxonomy, Geographical Distribution, and Paleogeography. Smithsonian Miscellaneous Collections,

81(8): 1-36, 3 figures.

Mobberly, William C, Jr.

1963. Hormonal and Environmental Regulation of the

Molting Cycle in the Crayfish Faxonella clypeata.

Tulane Studies in Zoology, ll(3):79-96, 9 figures.

Mocnkhatis, W. J.

1904. An Extra Pair of Appendages Modified for Copulatory Purposes in Cambarus I'iridis. Indiana Uniirrsity Science Bulletin, 1:17-18, 2 figures.

Nicholas, Brother G.

1960. Checklist of Macroscopic Troglobitic Organisms of

the United States. American Midland Naturalist,

64(1): 123-160.

Olivier, G. A.

1791. ftcrevisse. Astacus. Pages 327-349 in Histoire Naturelle des Insectes, Encyclopedic Methodique . . . ,

Zoology, volume 6. Paris and Liege.

Ortmann, A. E.

1892. Die Decapoden-Krebsc des Strassburger Museum. . . . Zoologischen Jahrbiichern, 6:1-58, 1 plate.

1902. The Geographical Distribution of Fresh-Water De-
76

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

capods and Its Bearing upon Ancient Geography.

Proceedings of the American Philosophical Society,

41 (171):267-400, 8 figures.

1905a. The Crawfishes of Western Pennsylvania. Annals

of the Carnegie Museum, 3(2):387-406.

1905b. A New Species of Cambarus from Louisiana. Ohio

Naturalist, 6(2):401-4O3, 1 figure.

1905c. The Mutual Affinities of the Species of the Genus

Cambarus, and Their Dispersal over the United

States. Proceedings of the American Philosophical

Society, 44(180):91-136, plate 3.

1905d. Procambarus, a New Subgenus of the Genus Cambarus. Annals of the Carnegie Museum, 3(3):435-

442, 3 figures.

1906a. Mexican, Central American and Cuban Cambari.

Proceedings of the Washington Academy of Sciences, 8:1-24, 4 figures.

1906b. The Crawfishes of the State of Pennsylvania.

Memoirs of the Carnegie Museum, 2(10): iv +343-

523, plates A, B, 39-43.

1909. Une £crevisse nouvelle du Mexique, Cambarus

(Cambarus) bouvieri nov. sp. Annales des Sciences

Naturelles, new series, 7:159-166, 4 figures.

1913. A New Species of the Genus Cambarus from the

Isle of Pines. Annals of the Carnegie Museum,

8(3-4):414-417.

1931. Crawfishes of the Southern Appalachians and the

Cumberland Plateau. Annals of the Carnegie Museum, 20 (2):61-160.

Osborn, R. C, and E. B. Williamson

1898. The Crayfish of Ohio. Sixth Annual Report of the

Ohio State Academy of Science, page 21.

Packard, A. S., Jr.

1880. Fossil Crawfish from the Tertiaries of Wyoming.

American Naturalist, 14(3):222-223.

1881a. A Fossil Tertiary Crayfish. American Naiuralist,

15(10):832-834, 2 figures.

1881b. On a Crayfish from the Lower Tertiary Beds of

Western Wyoming. Bulletin of the United States

Geological and Geographical Survey, 6(2):391-397,

2 figures.

1888. The Cave Fauna of North America, with Remarks

on the Anatomy of the Brain and Origin of the

Blind Species. Memoirs of the National Academy

of Sciences, 4:3-156, 21 figures, 27 plates.

Padgett, Allen

1970. Georgia Cave Fauna. Georgia Speleological Society

Bulletin, 1970:19-20.

Parks, Hal B., Clay Smith, and Jack Garrett

1939. The East Texas Crayfish and Lepidopterous Insects.

Technical Bulletin of Stephen F. Austin State

Teachers College, 1(5):2.

Pearse, A. S.

1911. Report on the Crustacea Collected by the University of Michigan-Walker Expedition in the State of

Vera Cruz, Mexico. 13th Report of the Michigan

Academy of Science, pages 108-113, 4 figures.

Penn, George Henry, Jr.

1939. The Type Locality of the Crawfish (Faxonius lancifer). American Midland Naturalist, 22 (1):215.

1941. Preliminary Report of a Survey of the Crawfishes

of Louisiana. Abstracts of Papers Presented at the

Eighty-Eighth Annual Meeting of the New Orleans

Academy of Sciences, page 8.

1942. Observations on the Biology of the Dwarf Crawfish,

Cambarellus shufeldtii (Faxon). American Midland

Naturalist, 28(3):644-647, 1 figure.

1946. A New Crawfish of the Genus Procambarus from

Louisiana (Decapoda: Astacidae). Journal of the

Washington Academy of Sciences, 36(l):27-29, 1

figure.

1950a. A New Crawfish of the Genus Orconectes from

Louisiana (Decapoda: Astacidae). Journal of the

Washington Academy of Sciences, 40(5): 166-169, 9

figures.

1950b. The Genus Cambarellus in Louisiana (Decapoda,

Astacidae). American Midland Naturalist, 44(2):

421-426, 1 figure.

1950c. A New Orconectes from the Pontchartrain Watershed in Louisiana and Mississippi (Decapoda: Astacidae). Journal of the Washington Academy of

Sciences, 40(11):381-384, 10 figures.

1950d. Utilization of Crawfishes by Cold-Blooded Vertebrates in the Eastern United States. American

Midland Naturalist, 44(3):643-658, 3 figures.

1952a. The Genus Orconectes in Louisiana (Decapoda,

Astacidae). American Midland Naiuralist, 47(3):

743-748.

1952b. A New Crawfish of the Viritis Section of the Genus

Orconectes (Decapoda, Astacidae). Natural History Miscellanea, 109: 7 pages, 14 figures.

1953a. A New Crawfish of the Genus Procambarus from

Louisiana and Arkansas (Decapoda: Astacidae).

Journal of the Washington Academy of Sciences,

43(5): 163-166, 12 figures.

1953b. A Redescription of the Crawfish Procambarus hinei

(Ortmann) (Decapoda, Astacidae). Tulane Studies

in Zoology, l(5):63-68, 15 figures.

1953c. A New Burrowing Crawfish of the Genus Procambarus from Louisiana and Mississippi (Decapoda,

Astacidae). Tulane S-tudies in Zoology, 1 (6):71-76,

12 figures.

1953d. Two New Crawfishes of the Genus Procambarus

from Texas, Louisiana, and Arkansas (Decapoda,

Astacidae). American Museum Novitates, 1636: 10

pages, 19 figures.

1953e. Classification. In Mel Washburn, Percy Viosca, Jr.,

and George Henry Penn, All About Crawfish.

Louisiana Conservationist, 5(6):2-7, illustrated.

1955. A New Cambarus of the Diogenes Section from

North Louisiana (Decapoda, Astacidae). Tulane

Studies in Zoology, 3(4):73-81, 13 figures.

1956. A New Crawfish of the Genus Procambarus from

Arkansas (Crustacea, Astacidae). Lloydia, 19(2):

109-119, 17 figures.
NUMBER 166 77

1957. Variation and Subspecies of the Crawfish Orconectes

palmen. Tulane Studies in Zoology, 5(10):231-262,

30 figures.

1959. An Illustrated Key to the Crawfishes of Louisiana,

with a Summary of Their Distribution within the

State (Decapoda, Astacidae). Tulane Studies in

Zoology, 7(l):3-20, 86 figures.

1962. A New Crawfish of the Hinei Section of the Genus

Procambarus (Decapoda, Astacidae). Crustaceana,

3(3):222-226, 11 figures.

1963. A New Crawfish from the Hatchie River in Mississippi and Tennessee (Decapoda, Astacidae). Proceedings of the Biological Society of Washington,

76(15): 121-125, 10 figures.

Penn, George Henry, Jr., and Horton H. Hobbs, Jr.

1958. A Contribution toward a Knowledge of the Crayfishes of Texas (Decapoda, Astacidae). Texas Journal of Science, 10(4):452-483, 68 figures.

Penn, George Henry, Jr., and Guy Marlow

1959. The Genus Cambarus in Louisiana. American Midland Naturalist, 61(1): 191-203, 14 figures.

Pennak, Robert W.

1953. Fresh-Water Invertebrates of the United Staies.

ix + 769 pages, 470 figures. New York: The Ronald

Press Company.

Pilgrim, R. L. C.

1960. Muscle Receptor Organs in Some Decapod Crustacea. Comparative Biochemical Physiology, 1:

248-257, 3 figures.

Price, F. M.

1900. The Eyes of Cambarus pellucidus [title, without

text, of a paper presented to the Academy], Proceedings of the Indiana Academy of Science, 1899:

155.

Prins, Rudolph

1968. A New Crayfish of the Genus Cambarus from North

Carolina (Decapoda, Astacidae). Journal of the

Elisha Mitchell Scientific Society, 84 (4):458-461, 11

figures.

Prins, Rudolph, and Joseph F. Fitzpatrick, Jr.

1965. The First-Form Male of Orconectes bisectus

Rhoades, a Poorly Known Kentucky Crawfish.

American Midland Naturalist, 74(1):141-147, 12

figures.

Prins, Rudolph, and Horton H. Hobbs, Jr.

1972. A New Crayfish of the Subgenus Punciicambarus

from the Savannah River Drainage with Notes on

Cambarus (P.) reburrus Prins (Decapoda, Astacidae).

Proceedings of the Biological Society of Washington,

84(47):411-420, 1 figure.

Rabe, Felix

1890. La question de Vecrevisse. 86 pages, 11 figures.

Sezanne: H. Patoux.

Rafinesque, C. S.

1817. Synopsis of Four New Genera and Ten New Species

of Crustacea, Found in the United States. American Monthly Magazine and Critical Revue, 2:40-43.

Randall, J. W.

1840. Catalogue of the Crustacea Brought by Thomas

Nuttall and J. K. Townsend, from the West Coast

of North America and the Sandwich Islands. . . .

Journal of the Academy of Natural Sciences, 3(1):

106-147, 5 plates.

Rathbun, Mary J.

1926. The Fossil Stalk-Eyed Crustacea of the Pacific Slope

of North America. Bulletin of the United States

National Museum, 138: vii+155 pages, 6 figures, 39

plates.

Ray, Johnny, and Verl Stevens

1970. Using Baytex to Control Crayfish in Ponds. Progressive Fish-Culturist, 33:58-69.

Reimer, Rollin DeWayne

1966. Two New Species of the Genus Cambarus from

Arkansas (Decapoda, Astacidae). Tulane Studies

in Zoology, 13(1):9-15, 18 figures.

1969. A Report on the Crawfishes (Decapoda, Astacidae)

of Oklahoma. Proceedings of the Oklahoma Academy of Sciences, 48:49-65, 46 figures.

Rhoades, Rendell

1941a. The Distribution of Cambarus sloani Bundy, in

Ohio. Ohio Journal of Science, 41(2):93-98, 6 figures.

1941b. Notes on Some Crayfishes from Alabama Caves, with

the Description of a New Species and a New Subspecies. Proceedings of the United States National

Museum, 91(3129): 141-148, figures 35, 36.

1944a. The Crayfishes of Kentucky, with Notes on Variation, Distribution and Descriptions of New Species

and Subspecies. American Midland Naturalist,

31(1): 111-149, 10 figures.

1944b. Further Studies on Distribution and Taxonomy of

Ohio Crayfishes and the Description of a New Subspecies. Ohio Journal of Science, 44 (2):95-99.

1959. A Review of the Status of Orconectes inermis Cope

and a Suggested Nomenclatural Change. American

Midland Naturalist, 61 (2):399-402.

Riegel, J. A.

1959. The Systematics and Distribution of Crayfishes in

California. California Fish and Game, 45(l):29-50,

10 figures.

Roberts, Tilden W.

1944. Light, Eyestalk Chemical, and Certain Other Factors as Regulators of Community Activity for the

Crayfish Cambarus virilis Hagen. Ecological Monographs, 14 (4): 350-392, 21 figures.

Sanders, Robert R.

1967. Alphabetical Synopsis of the Crayfish Pasifastacus

trowbridgeii (Stimpson). Carolina Tips, 30(9):3S,

1 figure.

Saussurc, Henri de

1857a. Note carcinologique sur la famille des Thalassides

et sur celle des Astacides. Revue et Magasin de

Zoologie Pure et Applique'e, series 2, 9:99-102.

1857b. Diagnoses de quelques crustaces nouveaux de

I'Amerique tropicale. Revue et Magasin de Zoologie Pure ei Appliquee, series 2, 9:501-505.

1858. Memoire sur divers crustaces nouveaux des Antilles
78

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

et du Mexique. Memoires de la Societe d'Histoire

Naturelle de Geneve, 14:417-496, 6 plates.

Say, Thomas

1817. An Account of the Crustacea of the United States.

Journal of the Academy of Natural Sciences of

Philadelphia, 1(6): 165-169.

Shufeldt, R. W.

1898. Some Observations on Crayfish. Shooting and Fishing (December 29), 25(11):227.

Skorikov, A. S.

1908. Contributions a la classification des Potamobiidae

d'Europe et d'Asie. Annuaire de Musee Zoologique

de I'Academie Imperiale des Sciences de St. Petersbourg, 12:115-118. [1907, in Russian.]

Spoor, W. A.

1955. Loss and Gain of Heat-Tolerance by the Crayfish.

Biological Bulletin, 108(l):77-87, 2 figures.

Steele, Mary

1902. The Crayfish of Missouri. University of Cincinnati

Bulletin, series 2, 2(10): 1-54, 6 plates.

Stevenson, J. Ross

1967. Lack of Correlation of Some Growth and Developmental Processes in the Crayfish. [Abstract.] American Zoologist, 7(2):208.

Stimpson, William

1857a. Notices of New Species of Crustacea of Western

North America; Being an Abstract from a Paper to

be Published in the Journal of the Society. Proceedings of the Boston Society of Natural History,

6 (6): 84-89.

1857b. On the Crustacea and Echinodermata of the Pacific

Shores of North America, Part I. Crustacea. Boston

Journal of Natural History, 6\4):444-532.

Streets, Thomas H.

1877. Description of Cambarus couesi, New Species of

Crawfish from Dakota. Bulletin of the United

States Geological Survey, 3:803-804.

Svardson, Gunnar

1965. The American Crayfish Pacifastacus leniusculus

(Dana) Introduced into Sweden. Institute of Freshwater Research, Drottningholm, report 46:90-94, 1

figure.

Tellkampf, Theodor A.

1844a. Ausfliig nach der Mammuthhohle in Kentucky.

Das Ausland, 168:671-672; 169:675-676; 170:679-680;

171:683-684; 172:687-688; 173:691-692; 174:695-696;

175:699-700.

1844b. Ueber den blinden Fisch der Mammuthhohle in

Kentucky, mit Bemerkungen iibcr cinige andere in

dieser Hohle lebende Thiere. Archiv fur Anatomie,

Physiologie und wissenschaftliche Medicin, Heft

IV, pages 381-394, plate 9.

1845. Memoirs on the Blind Fishes and Some Other Animals Living in the Mammoth Cave in Kentucky.

New York Journal of Medicine and Collateral

Science, July 1845:84-93, 1 plate.

Thompson, David H.

1967. A Comparison of Visual Acuity between Cave and

Epigeal Crayfish. The Wisconsin Speleologist,

6(3-4):46-54.

Threincn, C. W.

I9">8. Wisconsin Crayfish. Wisconsin Conservation Bulletin, 23(7): 1-3.

Underwood, Lucien M.

1886. List of the Described Species of Fresh Water Crustacea from America, North of Mexico. Bulletin of

the Illinois State Laboratory of Natural History,

2(5): 323-386.

Unestam, Torgny

1969. Resistance to the Crayfish Plague in Some American, Japanese and European Crayfishes. Institute

of Freshwater Research, Drottningholm, report 49:

202-209.

Van Straelen, Victor

1942. A propos de la distribution des ecrevisses, des

homards et des crabes d'eau douce. Bulletin du

Musee Royal d'Histoire des Sciences Naturelles de

Belgique, 18(56): 1-11.

Villalobos, Alejandro

1943. Estudios de los Cambarinos Mexicanos, I. Observaciones sobre Cambarellus Montezumne (Saussure) y

Algunas de sus Formas con Descripci6n de una

Subspecie Nueva. Anales del Institute) de Biologia,

Universidad Nacional Autonoma de Mexico, 14(2):

587-611, 2 plates.

1944a. Estudios de los Cambarinos Mexicanos, II. Dos

Especies Nucvas del Genero Paracambarus. Anales

del Instituto de Biologia, Universidad Nacional

Autonoma de Mexico, 15(1): 161-174, 2 plates.

1944b. Estudios de los Cambarinos Mexicanos, III. Una

Especie Nueva de Procambarus, Procambarus caballeroi n. sp. Anales del Instituto de Biologia,

Universidad Nacional Autonoma de Mexico, 15(1):

175-184, 2 plates.

1946. Estudios de los Cambarinos Mexicanos, IV. Consideraciones Acerca de la Posici6n Sistematica y

Redescripci6n de Procambarus digueti (Bouvier) y

Procambarus bouvieri (Ortmann). Anales del Instituto de Biologia, Universidad Nacional Autonoma

de Mexico, 17(l,2):215-230, 2 plates.

1947a. Estudios de los Cambarinos Mexicanos, V. Redescripci6n de Paracambarus paradoxus (Ort.) y Descripci6n de una Especie Nueva del Mismo Genero.

Anales del Instiiuto de Biologia, Universidad Nacional Autonoma de Mexico, 18(l):233-247, 3 plates.

1947b. Estudios de los Cambarinos Mexicanos, VI. Descripci6n de una Nueva Especie del Genero Paracambarus. Anales del Instituto de Biologia, Universidad Nacional Autonoma de Mexico, 18 (2):537-546,

2 plates.

1948. Fstudios de los Cambarinos Mexicanos, VII. Descripci6n de una Nueva Especie del Genero Procambarus, Procambarus acanthophorus n. sp. Anales

del Instituto de Biologia, Universidad Nacional

Autonoma de Mexico, 19(1): 175-182, 2 plates.

1949. Estudios de los Cambarinos Mexicanos, VIII. Descripci6n de una Nueva Especie del Genero Para-
NUMBER 166 79

cambarus. Anales del Instituto de Iiiologia, Universidad National Autonoma de Mexico, 20(1,2):

331-339, 2 plates.

1950. Contribution al Estudio dc los Cambarinos Mexicanos, IX. Estudio Taxonomico dc un Grupo de

Especies del Genero Procambarus. Anales del

Instituto de Biologia, Universidad National Autonoma de Mexico, 21 (2):367-413, 11 plates.

1952. Estudios de los Cambarinos Mcxicanos, X. Una

Nucva Especie del Genero Cambarellus del Estado

de Nuevo Lc6n. Anales del Instituto de Biologia,

Universidad National Autonoma de Mexico, 22(2):

525-532, 2 plates.

1953. Distribution Geografica y Notas Ecol6gicas dc los

Cambarinos Mcxicanos. Memoria del Congreso

Cientifico Mexicano, VII. Ciencias Biologicas, IV

Centenario de la Universidad de Mexico, 7:343-

374, 5 figures.

1954a. Estudios de los Cambarinos Mcxicanos, XI. Una

Nueva Subspecie de Procambarus simulans del

Estado de Nuevo Lc6n. Anales del Instiluto de

Biologia, Universidad National Autonoma de Mexico, 25(l,2):289-298, 2 plates.

1954b. Estudios de los Cambarinos Mexicanos, XII, Parte

1. Revisi6n de la Especie Afines a Procambarus

mexicanus (Erichson), con Descripci6n de Nuevas

Formas. Anales del Instituto de Biologia, Universidad National Autonoma de Mexico, 25(I,2):299-

379, 19 plates.

1955. Cambarinos de la Fauna Mexicana. (Crustacea Decapoda). xvi + 290 pages, 62 plates. Tesis, Facultad

de Ciencias. Univcrsidad Nacional Aut6noma de

Mexico.

1958. Estudios de los Cambarinos Mcxicanos, XIII. Dcscripci6n de una Nueva Especie de Cambarino del

Estado de Veracruz (Crustacea, Decapoda). Anales

del Instituto de Biologia, Unwersidad Nacional

Autonoma de Mexico, 28(l,2):279-288, 3 plates.

Walls, Jerry G.

1968. A New Faxonella from Northeast Louisiana (Decapoda, Astacidae). Proceedings of the Biological

Society of Washington, 81(4I):413-418, 9 figures.

1972. Three New Crawfishes Related to Orconectes difficilis (Faxon) (Decapoda: Astacidae). Proceedings

of the Biological Society of Washington, 84(53):449-

458, 4 figures.

Walls, Jerry G., and Joe B. Black

1967. New Variations and Records of the Crawfish Procambarus jaculus Hobbs (Crustacea, Decapoda, Astacidae). Proceedings of the Louisiana Academy

of Sciences, 30:60-62.

Williams, Austin B.

1952. Six New Crayfishes of the Genus Orconectes (Decapoda: Astacidae) from Arkansas, Missouri and

Oklahoma. Transactions of the Kansas Academy

of Science, 55(2):330-351, 48 figures.

1954. Spcciation and Distribution of the Crayfishes of the

Ozark Plateaus and Ouachita Provinces. University

of Kansas Science Bulletin, 36(12): 803-918, 253

figures.

Williamson, E. B.

1899. Notes on Ohio Astacidae. Pages 47-48 in Seventh

Annual Report of the Ohio State Academy of

Science.

1907. Notes on the Crayfish of Wells County, Indiana,

with Description of New Species. 31st Annual Report of the Department of Geology and Natural

Resources, Indiana, 1906:749-763, plate 35.
FIGURES 1-294

EXPLANATION OF ILLUSTRATIONS

All of the illustrations except Figure \4d, which was redrawn from Villalobos

(1952:527), were made from camera lucida drawings sketched during the preparation of this manuscript. The lettering utilized in Figure 1 also applies to Figures

2 through 7; that in Figure 8, to Figures 9 through 169; and that in Figure 170,

to Figures 171 through 294.

FIGURES 1-7.—a, dorsal view of carapace of male; b, mesial view of first pleopod of male;

c, lateral view of same; d, dorsal view of chela of male.

FIGURES 8-88 and 92-169 a, dorsal view of carapace of male, form I; b, mesial view of first

pleopod of male, form I; c, lateral view ;of same; d, lateral view of first pleopod of male, form

II; e, annulus ventralis; /, dorsal view of chela of male, form I.

FIGURES 89-91 Same as in Figure 8 except c and d are caudal views of first pleopods of firstand second-form males, respectively.

FIGURES 170-196 and 200-294 a, dorsal view of carapace of male, form I; b, lateral view of

first pleopod of male, form I; c, lateral view of tip of same; d, mesial view of tip of same;

e, lateral view of first pleopod of male, form II; /, annulus ventralis; g, dorsal view of chela of

male, form I.

FIGURFS 197-199.—Same as in Figure 170 except c is caudal view of tip of first pleopod of

male, form I.
NUMBER 166 81

2. Pacifastacus gambelii

4. Pacifastacus nigrescens

b V c V± d

1. Pacifastacus connectens

3. Pacifastacus fortis

5. Pacifastacus leniusculus leniusculus

r 1*

6. Pacifastacus leniusculus klamathensis 7. Pacifastacus leniusculus trowbridgii

FIGURES 1-7
82 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

8. Cambarellus shufeldtii

9. Cambarellus schmitti

10. Cambarellus puer

11. Cambarellus ninae

FIGURES 8-11
NUMBER 166

83

12. Cambarellus diminutus

13. Cambarellus occidentalis

14. Cambarellus alvarezi

15. Cambarellus chapalanus

FICURES 12-15
84

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

16. Cambarellus areolatus

17. Cambarellus montezumae

18. Cambarellus patzcuarensis

19. Cambarellus zempoalensis

FICURES 16-19
NUMBER 166 85

20. Barbicambarus cornutus

21. Cambarus (Aviticambarus) hamulatus

22. Cambarus (Aviticambarus) jonesi

23. Cambarus (Cambarus) bartonii bartonii

FIGURES 20-23
86

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

24. Cambarus (Cambarus) bartonii carinirostris

25. Cambarus (Cambarus) bartonii cavatus

26. Cambarus (Cambarus) howardi

/ s

27. Cambarus (Cambarus) sciotensis

FIGURES 24-27
NUMBER 166 87

28. Cambarus (Cambarus) ortmanni

29. Cambarus (Depressicambarus) halli

30. Cambarus (Depressicambarus) englishi

31. Cambarus (Depressicambarus) obstipus

FICURES 28-31
88 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

32. Cambarus (Depressicambarus) jordani

33. Cambarus (Depressicambarus) latimanus

34. Cambarus (Depressicambarus) floridanus

35. Cambarus (Depressicambarus) sphenoides

FIGURES 32-35
NUMBER 166 89

36. Cambarus (Depressicambarus) catagius

37. Cambarus (Depressicambarus) cymatilis

38. Cambarus (Depressicambarus) striatus

39. Cambarus (Depressicambarus) reduncus

FICURES 36-39
90 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

40. Cambarus (Erebicambarus) hubbsi

41. Cambarus (Erebicambarus) rusticiformis

42. Cambarus (Erebicambarus) hubrichti

43. Cambarus (Erebicambarus) tenebrosus

FICURES 40-43
NUMBER 166 91

44. Cambarus (Erebicambarus) cahni

45. Cambarus (Erebicambarus) ornatus

46. Cambarus (Erebicambarus) laevis

a ^

47. Cambarus (Hiaticambarus) girardianus

FIGURES 44-47
92 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

48. Cambarus (Hiaticambarus) longirostris

49. Cambarus (Hiaticambarus) longulus

50. Cambarus (Hiaticambarus) chasmodactylus

51. Cambarus (Jugicambarus) bouchardi

FICURES 48-51
NUMBER 166 93

52. Cambarus (Jugicambarus) obeyensis

53. Cambarus (Jugicambarus) distorts

54. Cambarus (Jugicambarus) unestami

55. Cambarus (Jugicambarus) parvoculus

FIGURES 52-55
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

56. Cambarus (Jugicambarus) conasaugaensis

57. Cambarus (Jugicambarus) asperimanus

58. Cambarus (Jugicambarus) brachydactylus

59. Cambarus (Jugicambarus) friaufi

FICURES 56-59
NUMBER 166 95

60. Cambarus (Jugicambarus) dubius

61. Cambarus (Jugicambarus) monongalensis

62. Cambarus (Jugicambarus) carolinus

63. Cambarus (Jugicambarus) gentryi

FICURES 60-63
96 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

64. Cambarus (Jugicambarus) crytodytes

65. Cambarus (Jugicambarus) causeyi

66. Cambarus (Jugicambarus) setosus

67. Cambarus (Jugicambarus) zophonastes

FICURES 64-67
NUMBER 166 97

68. Cambarus (Jugicambarus) tartarus

69. Cambarus (Lacunicambarus) diogenes diogenes

70. Cambarus (Lacunicambarus) diogenes ludovicianus

71. Cambarus (Puncticambarus) extraneus

FIGURES 68-71
98

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

72. Cambarus (Puncticambarus) spicatus

73. Cambarus (Puncticambarus) nerterius

74. Cambarus (Puncticambarus) acuminatus

75. Cambarus (Puncticambarus) chaugaensis

FIGURES 72-75
NUMBER 166

99

76. Cambarus (Puncticambarus) reburrus

77. Cambarus (Puncticambarus) robustus

78. Cambarus (Puncticambarus) veteranus

A J

".•>••:• -•-••/ •

79. Cambarus (Veticambarus) pristinus

FIGURES 76-79
100 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

80. Fallicambarus macneesei

81. Fallicambarus strawni

82. Fallicambarus fodiens

83. Fallicambarus hedgpethi

FIGURES 80-83
NUMBER 166 101

84. Fallicambarus uhleri

85. Fallicambarus hortoni

86. Fallicambarus oryktes

87. Fallicambarus byersi

FIGURES 84-87
102

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

88. Fallicambarus dissitus

89. Faxonella clypeata

90. Faxonella creaseri

91. Faxonella beveri

FIGURES 88-91
NUMBER 166

103

92. Hobbseus orconectoides

93. Hobbseus cristatus

94. Hobbseus prominens

95. Hobbseus attenuatus

FICURES 92-95
104

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

96. Hobbseus valleculus

97. Orconectes australis australis

[ \

98. Orconectes australis packardi

II

99. Orconectes incomptus

FICURES 96-99
NUMBER 166 105

r u

100. Orconectes inermis inermis

101. Orconectes inermis testii

102. Orconectes pellucidus

\ f

•t <•• :

*-

103. Orconectes limosus

FICURES 100-103
106 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

104. Orconectes wrighti

105. Orconectes indianensis

106. Orconectes difficilis difficilis

107. Orconectes difficilis blacki

FICURES 104-107
NUMBER 166

107

:

108. Orconectes difficilis maletae

109. Orconectes difficilis hathawayi

110. Orconectes perfectus

111. Orconectes harrisonii

FICURES 108-111
108

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

112. Orconectes sloanii

113. Orconectes kentuckiensis

114. Orconectes lancifer

115. Orconectes marchandi

FICURES 112-115
NUMBER 166 109

116. Orconectes eupunctus

117. Orconectes obscurus

118. Orconectes propinquus

119. Orconectes jeffersoni

FICURES 116-119
110

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

120. Orconectes virginiensis

121. Orconectes sanbornii sanbornii

122. Orconectes sanbornii erismophorous

123. Orconectes erichsonianus

FIGURES 120-123
NUMBER 166 111

124. Orconectes rafinesquei

125. Orconectes bisectus

126. Orconectes tricuspis

127. Orconectes iowaensis

FICURES 124-127
112

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

128. Orconectes illinoiensis

129. Orconectes shoupi

130. Orconectes quadruncus

131. Orconectes leptogonopodus

FICURES 128-131
NUMBER 166 113

132. Orconectes acares

133. Orconectes punctimanus

134. Orconectes hylas

135. Orconectes peruncus

FIGURES 132-135
114

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

136. Orconectes rusticus

137. Orconectes forceps

138. Orconectes placidus

139. Orconectes barrenensis

FICURES 136-139
NUMBER 166 115

140. Orconectes neglectus chaenodactylus

141. Orconectes neglectus neglectus

142. Orconectes longidigitus

143. Orconectes minis

FICURES 140-143
116

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

144. Orconectes nana

145. Orconectes macrus

146. Orconectes spinosus

147. Orconectes juvenilis

FICURES 144-147
NUMBER 166

117

148. Orconectes putnami

149. Orconectes medius

150. Orconectes luteus

151. Orconectes menae

FICURES 148-151
118

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

152. Orconectes transfuga

153. Orconectes ozarkae

154. Orconectes williamsi

155. Orconectes meeki meeki

FIGURES 152-155
NUMBER 166

119

156. Orconectes meeki brevis

157. Orconectes palmeri palmeri

158. Orconectes palmeri longimanus

159. Orconectes palmeri creolanus

FICURES 156-159
120

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

160. Orconectes hobbsi

161. Orconectes mississippiensis

. . J

'i

w*l

/ . • ( •

• • ;

162. Orconectes virilis

163. Orconectes causeyi

FIGURES 160-163
NUMBER 166 121

164. Orconectes immunis

165. Orconectes nais

166. Orconectes alabamensis

167. Orconectes rhoadesi

FICURES 164-167
122

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

168. Orconectes validus

169. Orconectes compressus

b V> d

170. Procambarus (Acucauda) fitzpatricki

171. Procambarus (Austrocambarus) vazquezae

FIGURES 168-171
NUMBER 166 123

172. Procambarus (Austrocambarus) aztecus

173. Procambarus (Austrocambarus) mexicanus

174. Procambarus (Austrocambarus) rodriguezi

175. Procambarus (Austrocambarus) veracruzanus

FICURES 172-175
124 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

r

176. Procambarus (Austrocambarus) mirandai

177. Procambarus (Austrocambarus) ruthveni ruthveni

178. Procambarus (Austrocambarus) ruthveni zapoapensis

11

179. Procambarus (Austrocambarus) williamsoni

FICURES 176-179
NUMBER 166 125

180. Procambarus (Austrocambarus) acanthophorus

181. Procambarus (Austrocambarus) llamasi

182. Procambarus (Austrocambarus) pilosimanus

Li

li

183. Procambarus (Austrocambarus) atkinsoni

FICURES 180-183
126

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

184. Procambarus (Austrocambarus) cubensis rivalis

185. Procambarus (Austrocambarus) cubensis cubensis

unknown

186. Procambarus (Austrocambarus) niveus

; j

m

187. Procambarus (Capillicambarus) hinei

FICURES 184-187
NUMBER 166

127

188. Procambarus (Capillicambarus) incilis

189. Procambarus (Girardiella) tulanei

190. Procambarus (Girardiella) simulans simulans

191. Procambarus (Girardiella) simulans regiomontanus

FICURES 188-191
128

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

192. Procambarus (Girardiella) hagenianus

193. Procambarus (Girardiella) gracilis

194. Procambarus (Hagenides) advena

195. Procambarus (Hagenides) geodytes

FICURES 192-195
NUMBER 166 129

196. Procambarus (Hagenides) pygmaeus

197. Procambarus (Hagenides) rogersi ocklocknensis

198. Procambarus (Hagenides) rogersi campestris

199. Procambarus (Hagenides) rogersi expletus

FIGURES 196-199
130

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

200. Procambarus (Hagenides) rogersi rogersi

201. Procambarus (Hagenides) truculentus

202. Procambarus (Leconticambarus) alleni

203. Procambarus (Leconticambarus) milleri

unknown

FIGURES 200-203
NUMBER 166

131

204. Procambarus (Leconticambarus) barbatus

205. Procambarus (Leconticambarus) latipleurum

206. Procambarus (Leconticambarus) apalachicolae

207. Procambarus (Leconticambarus) econfinae

FICI-RFS 204-207
132 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

208. Procambarus (Leconticambarus) escambiensis

209. Procambarus (Leconticambarus) pubischelae

210. Procambarus (Leconticambarus) rathbunae

211. Procambarus (Leconticambarus) capillatus

FIGURES 208-211
NUMBER 166 133

212. Procambarus (Leconticambarus) kilbyi

213. Procambarus (Leconticambarus) hubbelli

214. Procambarus (Leconticambarus) shermani

215. Procambarus (Lonnbergius) acherontis

FICURES 212-215
134 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

216. Procambarus (Mexicambarus) bouvieri

217. Procambarus (Ortmannicus) pictus

218. Procambarus (Ortmannicus) enoplosternum

219. Procambarus (Ortmannicus) chacei

FIGURES 216-219
NUMBER 166 135

220. Procambarus (Ortmannicus) epicyrtus

221. Procambarus (Ortmannicus) litosternum

222. Procambarus (Ortmannicus) lepidodactylus

223. Procambarus (Ortmannicus) pubescens

FIGURES 220-223
136 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

224. Procambarus (Ortmannicus) hirsutus

225. Procambarus (Ortmannicus) youngi

unknown unknown

226. Procambarus (Ortmannicus) angustatus

227. Procambarus (Ortmannicus) lucifugus alachua

FIGURES 224-227
NUMBER 166 137

228. Procambarus (Ortmannicus) lucifugus lucifugus

229. Procambarus (Ortmannicus) pallidus

230. Procambarus (Ortmannicus) horsti

231. Procambarus (Ortmannicus) orcinus

FICURES 22&-231
138

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

232. Procambarus (Ortmannicus) seminolae

233. Procambarus (Ortmannicus) ancylus

234. Procambarus (Ortmannicus) lunzi

235. Procambarus (Ortmannicus) fallax

FIGURES 232-235
NUMBER 166

139

236. Procambarus (Ortmannicus) leonensis

237. Procambarus (Ortmannicus) evermanni

238. Procambarus (Ortmannicus) pycnogonopodus

239. Procambarus (Ortmannicus) blandingii

FICURES 236-239
140 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

240. Procambarus (Ortmannicus) acutus acutus

241. Procambarus (Ortmannicus) acutus cuevachicae

242. Procambarus (Ortmannicus) texanus

243. Procambarus (Ortmannicus) lecontei

FICURES 240-243
NUMBER 166

141

244. Procambarus (Ortmannicus) hayi

245. Procambarus (Ortmannicus) lophotus

246. Procambarus (Ortmannicus) acutissimus

247. Procambarus (Ortmannicus) verrucosus

FICURES 244-247
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

248. Procambarus (Ortmannicus) jaculus

249. Procambarus (Ortmannicus) hybus

250. Procambarus (Ortmannicus) planirostris

251. Procambarus (Ortmannicus) mancus

FICURES 248-251
NUMBER 166 143

252. Procambarus (Ortmannicus) pearsei plumimanus

253. Procambarus (Ortmannicus) pearsei pearsei

254. Procambarus (Ortmannicus) lewisi

255. Procambarus (Ortmannicus) bivittatus

FIGURES 252-255
144

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

256. Procambarus (Ortmannicus) viaeviridis

257. Procambarus (Ortmannicus) caballeroi

unknown

258. Procambarus (Ortmannicus) villalobosi

259. Procambarus (Ortmannicus) toltecae

\ I

FIGURES 256-259
NUMBER 166

260. Procambarus (Ortmannicus) gonopodocristatus

261. Procambarus (Paracambarus) ortmannii

262. Procambarus (Paracambarus) paradoxus

unknown

263. Procambarus (Pennides) suttkusi

FIGURES 260-263
146 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

264. Procambarus (Pennides) natchitochae

265. Procambarus (Pennides) ablusus

266. Procambarus (Pennides) elegans

7\

r

267. Procambarus (Pennides) versutus

FICURES 264-267
NUMBER 166

268. Procambarus (Pennides) lylei

269. Procambarus (Pennides) echinatus

270. Procambarus (Pennides) dupratzi

271. Procambarus (Pennides) vioscai

FICURES 268-271
148 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

272. Procambarus (Pennides) penni

273. Procambarus (Pennides) lagniappe

274. Procambarus (Pennides) spiculifer

275. Procambarus (Pennides) gibbus

''

FIGURES 272-275
NUMBER 166 149

276. Procambarus (Pennides) ouachitae

277. Procambarus (Pennides) raneyi

278. Procambarus (Procambarus) digueti

a

n

279. Procambarus (Remoticambarus) pecki

FICURES 276-279
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

280. Procambarus (Scapulicambarus) paeninsulanus

281. Procambarus (Scapulicambarus) howellae

282. Procambarus (Scapulicambarus) okaloosae

283. Procambarus (Scapulicambarus) clarkii

FIGURES 280-283
NUMBER 166 151

284. Procambarus (Scapulicambarus) troglodytes

285. Procambarus (Tenuicambarus) tenuis

286. Procambarus (Villalobosus) riojai

287. Procambarus (Villalobosus) hoffmanni

FIGURES 284-287
152 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

288. Procambarus (Villalobosus) erichsoni

289. Procambarus (Villalobosus) contrerasi

290. Procambarus (Villalobosus) hortonhobbsi

291. Procambarus (Villalobosus) zihuateutlensis

FIGURES 288-291
NUMBER 166

292. Procambarus (Villalobosus) teziutlanensis

293. Procambarus (Villalobosus) tlapacoyanensis

294. Troglocambarus maclanei

FICURES 292-294
Index

(This index covers current valid names, nomina nuda, and synonyms; misspellings and combinations not presently in use are excluded)

ablusus, Procambarus (Pennides), 62, 146 (fig. 265)

acanthophorus, Procambarus (Austrocambarus), 43-44, 125

(fig. 180)

acares, Orconectes, 26, 113 (fig. 132)

acherontis, Procambarus, 52

Procambarus (Lonnbergius), 52, 133 (fig. 215)

Acucauda, subgenus, 3, 43

acuminatus, Cambarus (Puncticambarus), 20-21, 98 (fig. 74)

acutissimus, Procambarus (Ortmannicus), 53, 141 (fig. 246)

acutus acutus, Procambarus, 37

Procambarus (Ortmannicus), 53, 140 (fig. 240)

acutus cuevachicae, Procambarus (Ortmannicus), 53-54, 140

(fig. 241)

acutus, Procambarus acutus, 37

advena, Astacus, 48

Procambarus (Hagenides), 48, 128 (fig. 194)

affinis, Astacus, 33

alabamensis, Orconectes, 26, 121 (fig. 166)

alachua, Procambarus (Ortmannicus) lucifugus, 58, 136 (fig.

227)

alleni, Procambarus (Leconticambarus), 49-50, 130 (fig. 202)

alvarezi, Cambarellus, 7, 83 (fig. 14)

ancylus, Procambarus (Ortmannicus), 54, 138 (fig. 233)

angustatus, Procambarus (Ortmannicus), 54, 136 (fig. 226)

apalachicolae, Procambarus (Leconticambarus) , 50, 131 (fig.

206)

areolatus, Cambarellus, 7, 84 (fig. 16)

argillicola, Cambarus, 23

asperimanus, Cambarus bartonii, 16

Cambarus (Jugicambarus) , 16, 94 (fig. 57)

Astaci, 2

Astacidae, 2, 3, 5

Astacidea, 3

Astacoidea, 3

Astacus, 2, 3

advena, 48

affinis, 33

Bartonii, 3, 10

Blandingii, 3, 53, 54

breviforceps, 5

ciliaris, 10

fossarum, 66

fossor, 20

klamathensis, 5, 6

latimanus, 12, 13

limosus, 26, 33

maniculatus, 66

oreganus, 4

penicillatus, 50

pusillus, 10

subgrundialis, 5

Astacus, subgenus, 3

atkinsoni, Procambarus (Austrocambarus), 44, 125 (fig. 183)

Atlantoastacus, subgenus, 3

attenuatus, Hobbseus, 25, 103 (fig. 95)

australis australis, Orconectes, 26-27, 104 (fig. 97)

australis packardi, Orconectes, 27, 104 (fig. 98)

Austrocambarus, subgenus, 3, 43

Austropotamobius, 3

Austropotamobius, subgenus, 3

Aviticambarus, subgenus, 3, 10

ayersii, Cambarus, 19

aztecus, Procambarus (Austrocambarus) , 44, 123 (fig. 172)

barbatus, Cambarus, 49, 50

Procambarus (Leconticambarus), 50, 131 (fig. 204)

Barbicambarus, 3, 9

cornutus, 9, 85 (fig. 20)

Barbicambarus, subgenus, 3

barrenensis, Orconectes, 27, 114 (fig. 139)

bartoni tenebrosus, Cambarus, 14, 15

bartonii asperimanus, Cambarus, 16

Bartonii, Astacus, 3, 10

bartonii, Cambarus (Cambarus) bartonii, 10-11, 85 (fig. 23)

carinirostris, Cambarus (Cambarus), 11, 86 (fig. 24)

cavatus, Cambarus (Cambarus), 11, 86 (fig. 25)

spinirostris, Cambarus, 16

Bartonius, subgenus, 3

baumgartneri, Cambarus, 48

beyeri, Faxonella, 24, 102 (fig. 91)

bisectus, Orconectes, 27, 111 (fig. 125)

bivittatus, Procambarus (Ortmannicus) , 54, 143 (fig. 255)

blacki, Orconectes difficilis, 28, 106 (fig. 107)

Blandingii, Astacus, 3, 53, 54

blandingii, Procambarus (Ortmannicus) , 54, 139 (fig. 239)

bouchardi, Cambarus (Jugicambarus), 17, 92 (fig. 51)

bouvieri, Cambarus (Cambarus), 52, 53

Procambarus (Mexicambarus), 53, 134 (fig. 216)

brachydactylus, Cambarus (Jugicambarus), 17, 94 (fig. 58)

breviforceps, Astacus, 5

brevis, Orconectes meeki, 34, 119 (fig. 156)

byersi, Fallicambarus, 22-23, 101 (fig. 87)

caballeroi, Procambarus (Ortmannicus), 54-55, 144 (fig. 257)

cahni, Cambarus (Erebicambarus), 14, 91 (fig. 44)

Cambarellinae, 4, 7

Cambarellus, 3, 4, 7

alvarezi, 7, 83 (fig. 14)

154
NUMBER 166 155

areolatus, 7, 84 (fig. 16)

chapalanus, 7, 83 (fig. 15)

diminutus, 7-8, 83 (fig. 12)

montezumae, 8, 84 (fig. 17)

montezumae lermensis, 8

ninae, 8, 82 (fig. 11)

occidentalis, 8, 83 (fig. 13)

patzcuarensis, 8, 84 (fig. 18)

puer, 4, 8-9, 82 (fig. 10)

schmitti, 9, 82 (fig. 9)

shufeldtii, 9, 82 (fig. 8)

zempoalensis, 9, 84 (fig. 19)

Cambarellus, subgenus, 3

Cambaridae, 3, 4, 7

Cambarinae, 3, 9

Cambaroides, 4

Cambaroidinae, 4

Cambarus, 2, 3, 4, 10

argillicola, 23

(Aviticambarus) hamulatus, 10, 85 (fig. 21)

(Aviticambarus) jonesi, 10, 85 (fig. 22)

ayersii, 19

barbatus, 49, 50

bartoni tenebrosus, 14, 15

bartonii asperimanus, 16

bartonii spinirostris, 16

baumgartneri, 48

(Cambarus) bartonii bartonii, 10-11, 85 (fig. 23)

(Cambarus) bartonii carinirostris, 11, 86 (fig. 24)

(Cambarus) bartonii cavatus, 11, 86 (fig. 25)

(Cambarus) bouvieri, 52, 53

(Cambarus) hinei, 46

(Cambarus) howardi, 11, 86 (fig. 26)

(Cambarus) lobdelli, 4

(Cambarus) ortmanni, 11-12, 87 (fig. 28)

(Cambarus) sciotensis, 12, 86 (fig. 27)

carinatus, 64

clarkii paeninsulanus, 65

clypeatus, 3, 24

coecus, 37, 38

consobrinus, 44

couesi, 42

cristatus, 25

debilis, 42

(Depressicambarus) catagius, 12, 89 (fig. 36)

(Depressicambarus) cymatilis, 12, 89 (fig. 37)

(Depressicambarus) englishi, 12, 87 (fig. 30)

(Depressicambarus) floridanus, 12, 88 (fig. 34)

(Depressicambarus) halli, 13, 87 (fig. 29)

(Depressicambarus) jordani, 13, 88 (fig. 32)

(Depressicambarus) latimanus, 13, 88 (fig. 33)

(Depressicambarus) obstipus, 13. 87 (fig. 31)

(Depressicambarus) reduncus, 13, 89 (fig. 39)

(Depressicambarus) sphenoides, 13-14, 88 (fig. 35)

(Depressicambarus) striatus, 14, 89 (fig. 38)

digueti, 64

diogenes, 20

(Erebicambarus) cahni, 14, 91 (fig. 44)

(Erebicambarus) hubbsi, 14, 90 (fig. 40)

Cambarus—continued

(Erebicambarus) hubrichti, 14, 90 (fig. 42)

(Erebicambarus) laevis, 15, 91 (fig. 46)

(Erebicambarus) ornatus, 15, 91 (fig. 45)

(Erebicambarus) rusticiformis, 15, 90 (fig. 41)

(Erebicambarus) tenebrosus, 15, 90 (fig. 43)

erichsoni, 29

extraneus, 20, 21

faxonii, 32

(Faxonius) creaseri, 4

gallinus, 47

(Girardiella) hagenianus carri, 4

(Cirardiella) hagenianus evansi, 4

(Girardiella) hagenianus forestae, 4

gracilis, 47

graysoni, 14

(Hiaticambarus) chasmodactylus, 15, 92 (fig. 50)

(Hiaticambarus) girardianus, 16, 91 (fig. 47)

(Hiaticambarus) longirostris, 16, 92 (fig. 48)

(Hiaticambarus) longulus, 16, 92 (fig. 49)

immunis spinirostris, 30

(Jugicambarus) asperimanus, 16, 94 (fig. 57)

(Jugicambarus) bouchardi, 17, 92 (fig. 51)

(Jugicambarus) brachydactylus, 17, 94 (fig. 58)

(Jugicambarus) carolinus, 17, 95 (fig. 62)

(Jugicambarus) causeyi, 17, 96 (fig. 65)

(Jugicambarus) conasaugaensis, 17, 94 (fig. 56)

(Jugicambarus) cryptodytes, 18, 96 (fig. 64)

(Jugicambarus) distans, 18, 93 (fig. 53)

(Jugicambarus) dubius, 18, 95 (fig. 60)

(Jugicambarus) friaufi, 18, 94 (fig. 59)

(Jugicambarus) gentryi, 18, 95 (fig. 63)

(Jugicambarus) monongalensis, 18-19, 95 (fig. 61)

(Jugicambarus) obeyensis, 19, 93 (fig. 52)

(Jugicambarus) parvoculus, 19, 93 (fig. 55)

(Jugicambarus) setosus, 19, 96 (fig. 66)

(Jugicambarus) tartarus, 19, 97 (fig. 68)

(Jugicambarus) unestami, 19-20, 93 (fig. 54)

(Jugicambarus) zophonastes, 20, 96 (fig. 67)

(Lacunicambarus) diogenes diogenes, 20, 97 (fig. 69)

(Lacunicambarus) diogenes ludovicianus, 20, 97 (fig. 70)

longulus, 15, 16

lucifugus, 4

montanus, 10

Montezumae, 7, 8

montezumae dugesii, 8

Montezumae var. tridens, 8

nebrascensis, 20

obesus, 20

(Ortmannicus) cookae, 4

(Ortmannicus) evictus, 4

(Paracambarus) harnedi, 43

(Paracambarus) paradoxus, 61

Pealei, 33

penicillatus, 50, 54

pilosus, 35

primaevus, 4

pristinus, 22

(Puncticambarus) acuminatus, 20-21, 98 (fig. 74)
156

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Cambarus—continued

(Puncticambarus) chaugaensis, 21, 98 (fig. 75)

(Puncticambarus) extraneus, 21, 97 (fig. 71)

(Puncticambarus) nerterius, 21, 98 (fig. 73)

(Puncticambarus) reburrus, 21, 99 (fig. 76)

(Puncticambarus) robustus, 21-22, 99 (fig. 77)

(Puncticambarus) spicatus, 22, 98 (fig. 72)

(Puncticambarus) veteranus, 22, 99 (fig. 78)

schmittii, 4, 8

signifer, 30

spinosus gulielmi, 29

strawni, 22, 24

stygius, 37, 38, 53

typhlobius, 37

(Veticambarus) pristinus, 22, 99 (fig. 79)

whitmani, 33

wisconsinensis, 42

Cambarus, subgenus, 3, 10

campestris, Procambarus (Hagenides) rogersi, 49, 129 (fig.

198)

capillatus, Procambarus (Leconticambarus), 50, 132 (fig. 211)

Capillicambarus, subgenus, 3, 46

carinatus, Cambarus, 64

carinirostris, Cambarus (Cambarus) bartonii, 11, 86 (fig. 24)

carolinus, Cambarus (Jugicambarus), 17, 95 (fig. 62)

carri, Cambarus (Girardiella) hagenianus, 4

catagius, Cambarus (Depressicambarus) , 12, 89 (fig. 36)

causcyi, Cambarus (Jugicambarus), 17, 96 (fig. 65)

Orconectes, 27, 120 (fig. 163)

cavatus, Cambarus (Cambarus) bartonii, 11, 86 (fig. 25)

chacei, Procambarus (Ortmannicus), 55, 134 (fig. 219)

chaenodactylus, Orconectes neglectus, 36, 115 (fig. 140)

chapalanus, Cambarellus, 7, 83 (fig. 15)

chasmodactylus, Cambarus (Hiaticambarus), 15, 92 (fig. 50)

chaugaensis, Cambarus (Puncticambarus), 21, 98 (fig. 75)

chenoderma, Pacifastacus, 5

ciliaris, Astacus, 10

clarkii paeninsulanus, Cambarus, 65

clarkii, Procambarus (Scapulicambarus) , 65, 150 (fig. 283)

clypeata, Faxonella, 24, 102 (fig. 89)

clypeatus. Cambarus, 3, 24

coecus, Cambarus, 37, 38

compressus, Orconectes, 27-28, 122 (fig. 169)

conasaugaensis, Cambarus (Jugicambarus), 17, 94 (fig. 56)

connectens, Pacifastacus, 5, 81 (fig. 1)

consobrinus, Cambarus, 44

contrerasi, Procambarus (Villalobosus) , 66, 152 (fig. 289)

cookac, Cambarus (Ortmannicus) , 4

cornutus, Barbicambarus, 9, 85 (fig. 20)

couesi, Cambarus, 42

Crangonyx lucifugus, 4

crtascri, Cambarus (Faxonius), 4

Faxonella, 25, 102 (fig. 90)

crcolanus, Orconectes palmeri, 37, 119 (fig. 159)

cristatus, Cambarus, 25

Hobbseus, 25, 103 (fig. 93)

cryptodytcs, Cambarus (Jugicambarus) , 18, 96 (fig. 64)

cubensis cubensis, Procambarus (Austrocambarus), 44, 126

(fig. 185)

cubensis rivalis, Procambarus (Austrocambarus) ,44, 126 (fig.

184)

cuevachicae, Procambarus (Ortmannicus) acutus, 53-54, 140

(fig. 241)

cymatilis, Cambarus (Depressicambarus), 12, 89 (fig. 37)

debilis, Cambarus, 42

Depressicambarus, subgenus, 3, 12

difficilis blacki, Orconectes, 28, 106 (fig. 107)

difficilis difficilis, Orconectes, 28, 106 (fig. 106)

difficilis hathawayi, Orconectes, 28, 107 (fig. 109)

difficilis maletae, Orconectes, 28, 107 (fig. 108)

digueti, Cambarus, 64

Procambarus (Procambarus) , 64-65, 149 (fig. 278)

diminutus, Cambarellus, 7-8, 83 (fig. 12)

diogenes, Cambarus, 20

diogenes, Cambarus (Lacunicambarus), 20, 97 (fig. 69)

ludovicianus, Cambarus (Lacunicambarus), 20, 97 (fig. 70)

dissitus, Fallicambarus, 23, 102 (fig. 88)

distans, Cambarus (Jugicambarus), 18, 93 (fig. 53)

dubius, Cambarus (Jugicambarus), 18, 95 (fig. 60)

dugesii, Cambarus montezumae, 8

dupratzi, Procambarus (Pennides), 62, 147 (fig. 270)

echinatus, Procambarus (Pennides), 62, 147 (fig. 269)

econfinae, Procambarus (Leconticambarus), 50-51, 131 (fig.

207)

elegans, Procambarus (Pennides), 62, 146 (fig. 266)

englishi, Cambarus (Depressicambarus), 12, 87 (fig. 30)

enoplosternum, Procambarus (Ortmannicus) , 55, 134 (fig. 218)

epicyrtus, Procambarus (Ortmannicus), 55, 135 (fig. 220)

Erebicambarus, subgenus, 3, 14

erichsoni, Cambarus, 29

Procambarus (Villalobosus), 66-67, 152 (fig. 288)

erichsonianus, Orconectes, 29, 110 (fig. 123)

erismophorous, Orconectes sanbornii, 40, 110 (fig. 122)

escambiensis, Procambarus, 50

Procambarus (Leconticambarus), 51, 132 (fig. 208)

eupunctus, Orconectes, 29, 109 (fig. 116)

evansi, Cambarus (Girardiella) hagenianus, 4

evermanni, Procambarus (Ortmannicus) , 55, 139 (fig. 237)

evictus, Cambarus (Ortmannicus) , 4

expletus, Procambarus (Hagenides) rogersi, 49, 129 (fig. 199)

extraneus, Cambarus, 20, 21

Cambarus (Puncticambarus) , 21, 97 (fig. 71)

fallax, Procambarus (Ortmannicus), 55, 138 (fig. 235)

Fallicambarus, 3, 22

byersi, 22-23, 101 (fig. 87)

dissitus, 23, 102 (fig. 88)

fodiens, 23, 100 (fig. 82)

hedgpethi, 23, 100 (fig. 83)

hortoni, 23, 101 (fig. 85)

macneesei, 23-24, 100 (fig. 80)

oryktes, 24, 101 (fig. 86)

strawni, 24, 100 (fig. 81)

uhleri, 24, 101 (fig. 84)

Faxonella, 3, 24

beyeri, 24, 102 (fig. 91)
NUMBER 166 157

Faxonella—continued

clypeata, 24, 102 (fig. 89)

creased, 25, 102 (fig. 90)

Faxonella, subgenus, 3

faxonii, Cambarus, 32

Faxon i us, 3, 26

(Faxonius) immunis pedianus, 30

Faxonius, subgenus, 3, 26

fitzpatricki, Procambarus, 43

Procambarus (Acucauda), 43, 122 (fig. 170)

floridanus, Cambarus (Depressicambarus) , 12, 88 (fig. 34)

fodiens, Fallicambarus, 23, 100 (fig. 82)

forceps, Orconectes, 29, 114 (fig. 137)

forestae, Cambarus (Cirardiella) hagenianus, 4

fortis, Pacifastacus, 5, 81 (fig. 3)

fossarum, Astacus, 66

fossor, Astacus, 20

friaufi, Cambarus (Jugicambarus) , 18, 94 (fig. 59)

gallinus, Cambarus, 47

gambelii, Pacifastacus, 5-G, 81 (fig. 2)

gentryi, Cambarus (Jugicambarus) , 18, 95 (fig. 63)

geodytes, Procambarus (Hagenides), 48, 128 (fig. 195)

gibbus, Procambarus (Pennides), 62, 148 (fig. 275)

girardianus, Cambarus (Hiaticambarus) , 16, 91 (fig. 47)

Girardiella, subgenus, 3, 47

gonopodocristatus, Procambarus (Ortmannicus) , 56, 145 (fig.

260)

gracilis, Cambarus, 47

Procambarus (Girardiella) , 47, 128 (fig. 193)

graysoni, Cambarus, 14

gulielmi, Cambarus spinosus, 29

hagenianus carri, Cambarus (Girardiella), 4

hagenianus evansi, Cambarus (Girardiella), 4

hagenianus forestae, Cambarus (Girardiella) , 4

hagenianus, Procambarus (Girardiella), 47, 128 (fig. 192)

Hagenides, subgenus, 3, 48

halli, Cambarus (Depressicambarus), 13, 87 (fig. 29)

hamulatus, Cambarus (Aviticambarus), 10, 85 (fig. 21)

Orconectes, 10

harnedi, Cambarus (Paracambarus), 43

harrisonii, Orconectes, 29, 107 (fig. Ill)

hathawayi, Orconectes difficilis, 28, 107 (fig. 109)

hayi, Procambarus (Ortmannicus) , 56, 141 (fig. 244)

hcdgpethi, Fallicambarus, 23, 100 (fig. 83)

Hiaticambarus, subgenus, 3, 15

hinei, Cambarus (Cambarus), 46

Procambarus (Capillicambarus) , 46-47, 126 (fig. 187)

hirsutus, Procambarus (Ortmannicus), 56, 136 (fig. 224)

Hobbseus, 3, 25

attenuatus, 25, 103 (fig. 95)

cristatus, 25, 103 (fig. 93)

orconectoides, 25, 103 (fig. 92)

prominens, 25, 103 (fig. 94)

valleculus, 25-26, 104 (fig. 96)

hobbsi, Orconectes, 29-30, 120 (fig. 160)

hoffmanni, Procambarus (Villalobosus) , 67, 151 (fig. 287)

horsti, Procambarus (Ortmannicus) , 56, 137 (fig. 230)

hortonhobbsi, Procambarus (Villalobosus), 67, 152 (fig. 290)

hortoni, Fallicambarus, 23, 101 (fig. 85)

howardi, Cambarus (Cambarus), II, 86 (fig. 26)

howellae, Procambarus (Scapulicambarus), 65, 150 (fig. 281)

hubbelli, Procambarus (Leconticambarus), 51, 133 (fig. 213)

hubbsi, Cambarus (Erebicambarus), 14, 90 (fig. 40)

hubrichti, Cambarus (Erebicambarus), 14, 90 (fig. 42)

hybus, Procambarus (Ortmannicus), 56, 142 (fig. 249)

hylas, Orconectes, 30, 113 (fig. 134)

illinoiensis, Orconectes, 30, 112 (fig. 128)

immunis, Orconectes, 30, 121 (fig. 164)

pedianus, Faxonius (Faxonius), 30

spinirostris, Cambarus, 30

incilis, Procambarus (Capillicambarus), 47, 127 (fig. 188)

incomptus, Orconectes, 30-31, 104 (fig. 99)

indianensis, Orconectes, 31, 106 (fig. 105)

inermis inermis, Orconectes, 31, 105 (fig. 100)

inermis, Orconectes, 26, 31

testii, Orconectes, 31, 105 (fig. 101)

iowaensis, Orconectes, 31, 111 (fig. 127)

jaculus, Procambarus (Ortmannicus), 56-57, 142 (fig. 248)

jeffersoni, Orconectes, 31-32, 109 (fig. 119)

jonesi, Cambarus (Aviticambarus), 10, 85 (fig. 22)

jordani, Cambarus (Depressicambarus), 13, 88 (fig. 32)

Jugicambarus, subgenus, 3, 16

juvenilis, Orconectes, 32, 116 (fig. 147)

kentuckiensis, Orconectes, 32, 108 (fig. 113)

kilbyi, Procambarus (Leconticambarus), 51, 133 (fig. 212)

klamathensis, Astacus, 5, 6

Pacifastacus leniusculus, 6, 81 (fig. 6)

Lacunicambarus, subgenus, 3, 20

laevis, Cambarus (Erebicambarus), 15, 91 (fig. 46)

lagniappe, Procambarus (Pennides) , 62-63, 148 (fig. 273)

lancifer, Orconectes, 32, 108 (fig. 114)

latimanus, Astacus, 12, 13

Cambarus (Depressicambarus), 13, 88 (fig. 33)

latipleurum, Procambarus (Leconticambarus), 51, 131 (fig.

205)

lecontei, Procambarus (Ortmannicus) , 57, 140 (fig. 243)

Leconticambarus, subgenus, 3, 49

leniusculus klamathensis, Pacifastacus, 6, 81 (fig. 6)

leniusculus leniusculus, Pacifastacus, 6, 81 (fig. 5)

leniusculus trowbridgii, Pacifastacus, 6-7, 81 (fig. 7)

leonensis, Procambarus (Ortmannicus) , 57, 139 (fig. 236)

lepidodactylus, Procambarus (Ortmannicus), 57, 135 (fig. 222)

leptogonopodus, Orconectes, 32-33, 112 (fig. 131)

lermcnsis, Cainbarellus montezumae, 8

lewisi, Procambarus (Ortmannicus), 57, 143 (fig. 254)

limosus, Astacus, 26, 33

Orconectes, 33, 105 (fig. 103)

litosternum, Procambarus (Ortmannicus), 57, 135 (fig. 221)

llamasi, Procambarus (Austrocambarus), 44-45, 125 (fig. 181)

lobdelli, Cambarus (Cambarus), 4

longidigitus, Orconectes, 33, 115 (fig. 142)

longimanus, Orconectes palmeri, 37, 119 (fig. 158)

longirostris, Cambarus (Hiaticambarus), 16, 92 (fig. 48)
158 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

longulus, Cambarus, 15, 16

Cambarus (Hiaticambarus) , 16, 92 (fig. 49)

Lonnbergius, subgenus, 3, 52

lophotus, Procambarus (Ortmannicus) , 58, 141 (fig. 245)

lucifugus alachua, Procambarus (Ortmannicus), 58, 136 (fig.

227)

lucifugus, Cambarus, 4

Crangonyx, 4

lucifugus, Procambarus (Ortmannicus), 58, 137 (fig. 228)

ludovicianus, Cambarus (Lacunicambarus) diogenes, 20, 97

(fig. 70)

lunzi, Procambarus (Ortmannicus), 58, 138 (fig. 234)

luteus, Orconectes, 33, 117 (fig. 150)

lylei, Orconectes, 63, 147 (fig. 268)

madanei, Troglocambarus, 68, 153 (fig. 294)

macneesei, Fallicambarus, 23-24, 100 (fig. 80)

macrus, Orconectes, 33-34, 116 (fig. 145)

maletae, Orconectes difficilis, 28, 107 (fig. 108)

mancus, Procambarus (Ortmannicus), 58, 142 (fig. 251)

maniculatus, Astacus, 66

marchandi, Orconectes, 34, 108 (fig. 115)

medius, Orconectes, 34, 117 (fig. 149)

meeki brevis, Orconectes, 34, 119 (fig. 156)

meeki meeki, Orconectes, 34, 118 (fig. 155)

menae, Orconectes, 34-35, 117 (fig. 151)

Mexicambarus, subgenus, 3, 52

mexicanus, Procambarus (Austrocambarus), 45, 123 (fig. 173)

milled, Procambarus (Leconticambarus), 51-52, 130 (fig. 203)

mirandai, Procambarus (Austrocambarus), 45, 124 (fig. 176)

mirus, Orconectes, 35, 115 (fig. 143)

mississippiensis, Orconectes, 35, 120 (fig. 161)

monongalensis, Cambarus (Jugicambarus), 18-19, 95 (fig. 61)

montanus, Cambarus, 10

montezumae, Cambarellus, 8, 84 (fig. 17)

dugesii, Cambarus, 8

lermensis, Carbarellus, 8

var. tridens, Cambarus, 8

Montezumae, Cambarus, 7, 8

nais, Orconectes, 35, 121 (fig. 165)

nana, Orconectes, 35-36, 116 (fig. 144)

natchitochae, Procambarus, 62, 63

Procambarus (Pennides), 63, 146 (fig. 264)

nebrascensis, Cambarus, 20

neglectus chaenodactylus, Orconectes, 36, 115 (fig. 140)

neglectus neglectus, Orconectes, 36, 115 (fig. 141)

Nephropoidea, 3

nerterius, Cambarus (Puncticambarus) , 21, 98 (fig. 73)

nigrescens, Pacifastacus, 5, 7, 81 (fig. 4)

ninae, Cambarellus, 8, 82 (fig. 11)

niveus, Procambarus (Austrocambarus) , 45, 126 (fig. 186)

obesus, Cambarus, 20

obeyensis, Cambarus (Jugicambarus), 19, 93 (fig. 52)

obscurus, Orconectes, 36, 109 (fig. 117)

obstipus, Cambarus (Depressicambarus), 13, 87 (fig. 31)

occidentalis, Cambarellus, 8, 83 (fig. 13)

ochlocknensis, Procambarus (Hagenides) rogersi, 49, 129 (fig.

197)

okaloosae, Procambarus (Scapulicambarus), 65, 150 (fig. 282)

orcinus, Procambarus (Ortmannicus), 58-59, 137 (fig. 231)

Orconectes, 3, 26

acares, 26, 113 (fig. 132)

alabamensis, 26, 121 (fig. 166)

australis australis, 26-27, 104 (fig. 97)

australis packardi, 27, 104 (fig. 98)

barrenensis, 27, 114 (fig. 139)

bisectus, 27, 111 (fig. 125)

causeyi, 27, 120 (fig. 163)

compressus, 27-28, 122 (fig. 169)

difficilis blacki, 28, 106 (fig. 107)

difficilis difficilis, 28, 106 (fig. 106)

difficilis hathawayi, 28, 107 (fig. 109)

difficilis maletae, 28, 107 (fig. 108)

erichsonianus, 29, 110 (fig. 123)

eupunctus, 29, 109 (fig. 116)

forceps, 29, 114 (fig. 137)

hamulatus, 10

harrisonii, 29, 107 (fig. Ill)

hobbsi, 29-30, 120 (fig. 160)

hylas, 30, 113 (fig. 134)

illinoiensis, 30, 112 (fig. 128)

immunis, 30, 121 (fig. 164)

incomptus, 30-31, 104 (fig. 99)

indianensis, 31, 106 (fig. 105)

inermis, 26, 31

inermis inermis, 31, 105 (fig. 100)

inermis testii, 31, 105 (fig. 101)

iowaensis, 31, 111 (fig. 127)

jeffersoni, 31-32, 109 (fig. 119)

juvenilis, 32, 116 (fig. 147)

kentuckiensis, 32, 108 (fig. 113)

lancifer, 32, 108 (fig. 114)

leptogonopodus, 32-33, 112 (fig. 131)

limosus, 33, 105 (fig. 103)

longidigitus, 33, 115 (fig. 142)

luteus, 33, 117 (fig. 150)

macrus, 33-34, 116 (fig. 145)

marchandi, 34, 108 (fig. 115)

medius, 34, 117 (fig. 149)

meeki brevis, 34, 119 (fig. 156)

meeki meeki, 34, 118 (fig. 155)

menae, 34-35, 117 (fig. 151)

mirus, 35, 115 (fig. 143)

mississippiensis, 35, 120 (fig. 161)

nais, 35, 121 (fig. 165)

nana, 35-36, 116 (fig. 144)

neglectus chaenodactylus, 36, 115 (fig. 140)

neglectus neglectus, 36, 115 (fig. 141)

obscurus. 36, 109 (fig. 117)

ozarkae, 36-37, 118 (fig. 153)

palmeri creolanus, 37, 119 (fig. 159)

palmeri longimanus, 37, 119 (fig. 158)

palmeri palmeri. 37, 119 (fig. 157)

pellucidus, 26, 37-38, 105 (fig. 102)

perfectus, 38, 107 (fig. 110)

peruncus, 38, 113 (fig. 135)

placidus, 38, 114 (fig. 138)
NUMBER 166 159

Orconectes—continued

propinquus, 38, 39, 109 (fig. 118)

punctimanus, 39, 113 (fig. 133)

putnami, 32, 39, 117 (fig. 148)

quadruncus, 39, 112 (fig. 130)

rafinesquei, 40, 111 (fig. 124)

rhoadesi, 40, 121 (fig. 167)

rusticus, 40, 114 (fig. 136)

sanbornii erismophorous, 40, 110 (fig. 122)

sanbornii sanbornii, 40-41, 110 (fig. 121)

shoupi, 41, 112 (fig. 129)

sloanii, 41, 108 (fig. 112)

spinosus, 32, 41, 116 (fig. 146)

transfuga, 41-42, 118 (fig. 152)

tricuspis, 42, 111 (fig. 126)

validus, 42, 122 (fig. 168)

virginiensis, 42, 110 (fig. 120)

virilis, 42-13, 120 (fig. 162)

williamsi, 43, 118 (fig. 154)

wrighti, 43, 106 (fig. 104)

orconectoides, Hobbseus, 25, 103 (fig. 92)

oreganus, Astacus, 4

ornatus, Cambarus (Erebicambarus) , 15, 91 (fig. 45)

ortmanni, Cambarus (Cambarus), 11-12, 87 (fig. 28)

Ortmannicus, subgenus, 3, 43, 53

ortmannii, Procambarus (Paracambarus), 61, 145 (fig. 261)

oryktes, Fallicambarus, 24, 101 (fig. 86)

ouachitae, Procambarus (Pennides), 63, 149 (fig. 276)

ozarkae, Orconectes, 36-37, 118 (fig. 153)

Pacifastacus, 3, 5

chenoderma, 5

connectens, 5, 81 (fig. 1)

fortis, 5, 81 (fig. 3)

gambelii, 5-6, 81 (fig. 2)

leniusculus klamathensis, 6, 81 (fig. 6)

leniusculus leniusculus, 6, 81 (fig. 5)

leniusculus trowbridgii, 6-7, 81 (fig. 7)

nigrescens, 5, 7, 81 (fig. 4)

packardi, Orconectes australis, 27, 104 (fig. 98)

paeninsulanus, Cambarus clarkii, 65

Procambarus (Scapulicambarus) , 65-66, 150 (fig. 280)

pallidus, Procambarus (Ortmannicus), 59, 137 (fig. 229)

palmeri creolanus, Orconectes, 37, 119 (fig. 159)

palmed longimanus, Orconectes, 37, 119 (fig. 158)

palmeri palmeri, Orconectes, 37, 119 (fig. 157)

Paracambarus, 3, 43

riojae, 66, 67

Paracambarus, subgenus, 4, 43, 61

paradoxus, Cambarus (Paracambarus), 61

Procambarus (Paracambarus) , 61-62, 145 (fig. 262)

Parastacoidea, 3

parvoculus, Cambarus (Jugicambarus), 19, 93 (fig. 55)

patzcuarensis, Cambarellus, 8, 84 (fig. 18)

Pealei, Cambarus, 33

pearsei pearsei, Procambarus (Ortmannicus), 59, 143 (fig. 253)

pearsei plumimanus, Procambarus (Ortmannicus), 59, 143

(fig- 252)

pecki, Procambarus, 65

pecki—continued

Procambarus (Remoticambarus), 65, 149 (fig. 279)

pedianus, Faxonius (Faxonius) immunis, 30

pellucidus, Orconectes, 26, 37-38, 105 (fig. 102)

penicillatus, Astacus, 50

Cambarus, 50, 54

penni, Procambarus (Pennides), 63, 148 (fig. 272)

Pennides, subgenus, 4, 62

perfectus, Orconectes, 38, 107 (fig. 110)

peruncus, Orconectes, 38, 113 (fig. 135)

pictus, Procambarus (Ortmannicus), 59, 134 (fig. 217)

pilosimanus, Procambarus (Austrocambarus), 45, 125 (fig.

182)

pilosus, Cambarus, 35

placidus, Orconectes, 38, 114 (fig. 138)

planirostris, Procambarus (Ortmannicus), 59-60, 142 (fig. 250)

plumimanus, Procambarus (Ortmannicus) pearsei, 59, 143

(fig- 252)

Pontastacus, subgenus, 3

Potamobius, 5

primaevus, Cambarus, 4

pristinus, Cambarus, 22

Cambarus (Veticambarus), 22, 99 (fig. 79)

Procambarus, 3, 5, 43

acherontis, 52

(Acucauda) fitzpatricki, 43, 122 (fig. 170)

acutus acutus, 37

(Austrocambarus) acanthophorus, 43-44, 125 (fig. 180)

(Austrocambarus) atkinsoni, 44, 125 (fig. 183)

(Austrocambarus) aztecus, 44, 123 (fig. 172)

(Austrocambarus) cubensis cubensis, 44, 126 (fig. 185)

(Austrocambarus) cubensis rivalis, 44, 126 (fig. 184)

(Austrocambarus) llamasi, 44-45, 125 (fig. 181)

(Austrocambarus) mexicanus, 45, 123 (fig. 173)

(Austrocambarus) mirandai, 45, 124 (fig. 176)

(Austrocambarus) niveus, 45, 126 (fig. 186)

(Austrocambarus) pilosimanus, 45, 125 (fig. 182)

(Austrocambarus) rodriguezi, 45, 123 (fig. 174)

(Austrocambarus) ruthveni ruthveni, 46, 124 (fig. 177)

(Austrocambarus) ruthveni zapoapensis, 46, 124 (fig. 178)

(Austrocambarus) vazquezae, 46, 122 (fig. 171)

(Austrocambarus) veracruzanus, 46, 123 (fig. 175)

(Austrocambarus) williamsoni, 46, 124 (fig. 179)

(Capillicambarus) hinei, 46-47, 126 (fig. 187)

(Capillicambarus) incilis, 47, 127 (fig. 188)

escambiensis, 50

fitzpatricki, 43

(Girardiella) gracilis, 47, 128 (fig. 193)

(Girardiella) hagenianus, 47, 128 (fig. 192)

(Girardiella) simulans regiomontanus, 47, 127 (fig. 191)

(Girardiella) simulans simulans, 47-48, 127 (fig. 190)

(Girardiella) tulanei. 48, 127 (fig. 189)

(Hagenides) advena, 48, 128 (fig. 194)

(Hagenides) geodytes, 48, 128 (fig. 195)

(Hagenides) pygmaeus, 48-49, 129 (fig. 196)

(Hagenides) rogersi campestris, 49. 129 (fig. 198)

(Hagenides) rogersi expletus, 49. 129 (fig. 199)

(Hagenides) rogersi ochlocknensis, 49, 129 (fig. 197)

(Hagenides) rogersi rogersi, 49, 130 (fig. 200)
160

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Procambarus—continued

(Hagenides) truculentus, 49, 130 (fig. 201)

(Leconticambarus) alleni, 49-50, 130 (fig. 202)

(Leconticambarus) apalachicolae, 50, 131 (fig. 206)

(Leconticambarus) barbatus, 50, 131 (fig. 204)

(Leconticambarus) capillatus, 50, 132 (fig. 211)

(Leconticambarus) econfinae, 50-51, 131 (fig. 207)

(Leconticambarus) escambiensis, 51, 132 (fig. 208)

(Leconticambarus) hubbelli, 51, 133 (fig. 213)

(Leconticambarus) kilbyi, 51, 133 (fig. 212)

(Leconticambarus) latipleurum, 51, 131 (fig. 205)

(Leconticambarus) milleri, 51-52, 130 (fig. 203)

(Leconticambarus) pubischelae, 52, 132 (fig. 209)

(Leconticambarus) rathbunae, 52, 132 (fig. 210)

(Leconticambarus) shermani, 52, 133 (fig. 214)

(Lonnbergius) acherontis, 52, 133 (fig. 215)

(Mexicambarus) bouvieri, 53, 134 (fig. 216)

natchitochae, 62, 63

(Ortmannicus) acutissimus, 53, 141 (fig. 246)

(Ortmannicus) acutus acutus, 53, 140 (fig. 240)

(Ortmannicus) acutus cuevachicae, 53-54, 140 (fig. 241)

(Ortmannicus) ancylus, 54, 138 (fig. 233)

(Ortmannicus) angustatus, 54, 136 (fig. 226)

(Ortmannicus) bivittatus, 54, 143 (fig. 255)

(Ortmannicus) blandingii, 54, 139 (fig. 239)

(Ortmannicus) caballeroi, 54-55, 144 (fig. 257)

(Ortmannicus) chacei, 55, 134 (fig. 219)

(Ortmannicus) enoplosternum, 55, 134 (fig. 218)

(Ortmannicus) epicyrtus, 55, 135 (fig. 220)

(Ortmannicus) evermanni, 55, 139 (fig. 237)

(Ortmannicus) fallax, 55, 138 (fig. 235)

(Ortmannicus) gonopodocristatus, 56, 145 (fig. 260)

(Ortmannicus) hayi, 56, 141 (fig. 244)

(Ortmannicus) hirsutus, 56, 136 (fig. 224)

(Ortmannicus) horsti, 56, 137 (fig. 230)

(Ortmannicus) hybus, 56, 142 (fig. 249)

(Ortmannicus) jaculus, 56-57, 142 (fig. 248)

(Ortmannicus) lecontei, 57, 140 (fig. 243)

(Ortmanniciis) leonensis, 57, 139 (fig. 236)

(Ortmannicus) lepidodactylus, 57, 135 (fig. 222)

(Ortmannicus) lewisi, 57, 143 (fig. 254)

(Ortmannicus) litostcrnum, 57, 135 (fig. 221)

(Ortmannicus) lophotus, 58, 141 (fig. 245)

(Ortmannicus) lucifugus alachua, 58, 136 (fig. 227)

(Ortmannicus) lucifugus lucifugus, 58, 137 (fig. 228)

(Ortmannicus) lunzi, 58, 138 (fig. 234)

(Ortmannicus) mancus, 58, 142 (fig. 251)

(Ortmannicus) orcinus, 58-59, 137 (fig. 231)

(Ortmannicus) pallidus, 59, 137 (fig. 229)

(Ortmannicus) pearsci pearsei, 59, 143 (fig. 253)

(Ortmannicus) pearsei plumimanus, 59, 143 (fig. 252)

(Ortmannicus) pictus, 59, 134 (fig. 217)

(Ortmannicus) planirostris, 59-60, 142 (fig. 250)

(Ortmannicus) pubescens, 60, 135 (fig. 223)

(Ortmannicus) pyenogonopodus, 60, 139 (fig. 238)

(Ortmannicus) seminolae, 60, 138 (fig. 232)

(Ortmannicus) texanus, 60, 140 (fig. 242)

(Ortmannicus) toltccae, 60, 144 (fig. 259)

(Ortmannicus) verrucosus, 61, 141 (fig. 247)

Procambarus—continued

(Ortmannicus) viaeviridis, 61, 144 (fig. 256)

(Ortmannicus) villalobosi, 61, 144 (fig. 258)

(Ortmannicus) youngi, 61, 136 (fig. 225)

(Paracambarus) ortmannii, 61, 145 (fig. 261)

(Paracambarus) paradoxus, 61-62, 145 (fig. 262)

pecki, 65

(Pennides) ablusus, 62, 146 (fig. 265)

(Pennides) dupratzi, 62, 147 (fig. 270)

(Pennides) echinatus, 62, 147 (fig. 269)

(Pennides) elegans, 62, 146 (fig. 266)

(Pennides) gibbus, 62, 148 (fig. 275)

(Pennides) lagniappe, 62-63, 148 (fig. 273)

(Pennides) lylei, 63, 147 (fig. 268)

(Pennides) natchitochae, 63, 146 (fig. 264)

(Pennides) ouachitae, 63, 149 (fig. 276)

(Pennides) penni, 63, 148 (fig. 272)

(Pennides) raneyi, 63, 149 (fig. 277)

(Pennides) spiculifer, 64, 148 (fig. 274)

(Pennides) suttkusi, 64, 145 (fig. 263)

(Pennides) versutus, 64, 146 (fig. 267)

(Pennides) vioscai, 64, 147 (fig. 271)

(Procambarus) digueti, 64-65, 149 (fig. 278)

(Remoticambarus) pecki, 65, 149 (fig. 279)

(Scapulicambarus) clarkii, 65, 150 (fig. 283)

(Scapulicambarus) howellae, 65, 150 (fig. 281)

(Scapulicambarus) okaloosae, 65, 150 (fig. 282)

(Scapulicambarus) paeninsulanus, 65-66, 150 (fig. 280)

(Scapulicambarus) troglodytes, 66, 151 (fig. 284)

(Tenuicambarus) tenuis, 66, 151 (fig. 285)

tenuis, 66

vazquezae, 43, 46

(Villalobosus) contrerasi, 66, 152 (fig. 289)

(Villalobosus) erichsoni, 66-67, 152 (fig. 288)

(Villalobosus) hoffmanni, 67, 151 (fig. 287)

(Villalobosus) hortonhobbsi, 67, 152 (fig. 290)

(Villalobosus) riojai, 67, 151 (fig. 286)

(Villalobosus) teziutlanensis, 67, 153 (fig. 292)

(Villalobosus) tlapacoyanensis, 67, 153 (fig. 293)

(Villalobosus) zihuateutlensis, 67-68, 152 (fig. 291)

wiegmanni, 4

Procambarus, subgenus, 3, 64

prominens, Hobbseus, 25, 103 (fig. 94)

propinquus, Orconectes. 38-39, 109 (fig. 118)

pubescens, Procambarus (Ortmannicus) , 60, 135 (fig. 223)

pubischelae, Procambarus (Leconticambarus), 52, 132 (fig. 209)

puer, Cambarellus, 4, 8-9, 82 (fig. 10)

Puncticambarus, subgenus, 3, 20

punctimanus, Orconectes, 39, 113 (fig. 133)

pusillus, Astacus, 10

putnami, Orconectes, 32, 39, 117 (fig. 148)

pyenogonopodus, Procambarus (Ortmannicus), 60, 139 (fig.

238)

pygmaeus, Procambarus (Hagenides), 48-49, 129 (fig. 196)

quadruncus, Orconectes, 39, 112 (fig. 130)

rafinesquei, Orconectes, 40, 111 (fig. 124)

raneyi, Procambarus (Pennides), 63, 149 (fig. 277)
NUMBER 166 161

rathbunae, Procambarus (Leconticambarus), 52, 132 (fig. 210)

reburrus, Cambarus (Puncticambarus), 21, 99 (fig. 76)

reduncus, Cambarus (Depressicambarus), 13, 89 (fig. 39)

regiomontanus, Procambarus (Girardiella) simulans, 47, 127

(fig. 191)

Remoticambarus, subgenus, 4, 65

rhoadesi, Orconectes, 40, 121 (fig. 167)

riojae, Paracambarus, 66, 67

riojai, Procambarus (Villalobosus) , 67, 151 (fig. 286)

rivalis, Procambarus (Austrocambarus) cubensis, 44, 126 (fig.

184)

robustus, Cambarus (Puncticambarus), 21-22, 99 (fig. 77)

rodriguezi, Procambarus (Austrocambarus), 45, 123 (fig. 174)

rogersi campestris, Procambarus (Hagenides) , 49, 129 (fig198)

rogersi expletus, Procambarus (Hagenides), 49, 129 (fig. 199)

rogersi ochlocknensis, Procambarus (Hagenides) , 49, 129 (fig.

197)

rogersi rogersi, Procambarus (Hagenides), 49, 130 (fig. 200)

rusriciformis. Cambarus (Erebicambarus), 15, 90 (fig. 41)

rusticus, Orconectes, 40, 114 (fig. 136)

ruthveni ruthveni, Procambarus (Austrocambarus), 46, 124

(fig- 177)

ruthveni zapoapensis, Procambarus (Austrocambarus) , 46, 124

(fig. 178)

sanbornii erismophorous, Orconectes, 40, 110 (fig. 122)

sanbornii sanbornii, Orconectes, 40-41, 110 (fig. 121)

Scapulicambarus, subgenus, 4, 65

schmitti, Cambarellus, 9, 82 (fig. 9)

schmittii, Cambarus, 4, 8

sciotensis, Cambarus (Cambarus) , 12, 86 (fig. 27)

seminolae, Procambarus (Ortmannicus), 60, 138 (fig. 232)

setosus, Cambarus (Jugicambarus), 19, 96 (fig. 66)

shermani, Procambarus (Leconticambarus), 52, 133 (fig. 214)

shoupi, Orconectes, 41, 112 (fig. 129)

shufeldtii, Cambarellus, 9, 82 (fig. 8)

signifer, Cambarus, 30

simulans regiomontanus, Procambarus (Girardiella), 47, 127

(fig. 191)

simulans simulans, Procambarus (Girardiella), 47-48, 127

[fig. 190)

sloanii, Orconectes, 41, 108 (fig. 112)

sphenoides, Cambarus (Depressicambarus) , 13-14, 88 (fig. 35)

spicatus, Cambarus (Puncticambarus), 22, 98 (fig. 72)

spiculifer, Procambarus (Pennides), 64, 148 (fig. 274)

spinirostris, Cambarus bartonii, 16

Cambarus immunis, 30

spinosus gulielmi, Cambarus, 29

spinosus, Orconectes, 32, 41, 116 (fig. 146)

strawni, Cambarus, 22, 24

Fallicambarus, 24, 100 (fig. 81)

striatus, Cambarus (Depressicambarus) , 14, 89 (fig. 38)

stygius, Cambarus, 37, 38, 53

subgrundialis, Astacus, 5

suttkusi, Procambarus (Pennides) , 64, 145 (fig. 263)

tartarus, Cambarus (Jugicambarus) , 19, 97 (fig. 68)

tenebrosus, Cambarus bartoni, 14, 15

Cambarus (Erebicambarus), 15, 90 (fig. 43)

Tenuicambarus, subgenus, 4, 66

tenuis, Procambarus, 66

Procambarus (Tenuicambarus) ,66 , 151 (fig. 285)

testii, Orconectes inermis, 31, 105 (fig. 101)

texanus, Procambarus (Ortmannicus), 60, 140 (fig. 242)

teziutlanensis, Procambarus (Villalobosus), 67, 153 (fig. 292)

tlapacoyanensis, Procambarus (Villalobosus), 67-68, 152 (fig.

293)

toltecae, Procambarus (Ortmannicus), 60, 144 (fig. 259)

transfuga, Orconectes, 41-42, 118 (fig. 152)

tricuspis, Orconectes, 42, 111 (fig. 126)

tridens, Cambarus Montezumae, 8

Troglocambarus, 3, 4, 68

maclanei, 68, 153 (fig. 294)

troglodytes, Procambarus (Scapulicambarus) ,66, 151 (fig. 284)

trowbridgii, Pacifastacus leniusculus, 6-7, 81 (fig. 7)

truculentus, Procambarus (Hagenides), 49, 130 (fig. 201)

tulanei, Procambarus (Girardiella), 48, 127 (fig. 189)

typhlobius, Cambarus, 37

uhleri, Fallicambarus, 24, 101 (fig. 84)

unestami, Cambarus (Jugicambarus), 19-20, 93 (fig. 54)

validus, Orconectes, 42, 122 (fig. 168)

valleculus, Hobbseus, 25-26, 104 (fig. 96)

vazquezae, Procambarus, 43, 46

Procambarus (Austrocambarus), 46, 122 (fig. 171)

veracruzanus, Procambarus (Austrocambarus), 46, 123 (fig.

175)

verrucosus, Procambarus (Ortmannicus), 61, 141 (fig. 247)

versutus, Procambarus (Pennides), 64, 146 (fig. 267)

veteranus, Cambarus (Puncticambarus) , 22, 99 (fig. 78)

Veticambarus, subgenus, 3, 22

viaeviridis, Procambarus (Ortmannicus), 61, 144 (fig. 256)

villalobosi, Procambarus (Ortmannicus), 61, 144 (fig. 258)

Villalobosus, subgenus, 4, 66

vioscai, Procambarus (Pennides) ,64, 147 (fig. 271)

virginiensis, Orconectes, 42, 110 (fig. 120)

virilis, Orconectes, 42-43, 120 (fig. 162)

whitmani, Cambarus, 33

wiegmanni, Procambarus, 4

williamsi, Orconectes, 43, 118 (fig. 154)

williamsoni, Procambarus (Austrocambarus), 46, 124 (fig. 179)

wisconsinensis, Cambarus, 42

wrighti, Orconectes, 43, 106 (fig. 104)

youngi, Procambarus (Ortmannicus), 61, 136 (fig. 225)

zapoapensis, Procambarus (Austrocambarus) ruthveni, 46, 124

(fig. 178)

zempoalcnsis, Cambarellus, 9, 84 (fig. 19)

zihuateutlensis, Procambarus (Villalobosus), 67-68, 152 (fig.

291)

zophonastes, Cambarus (Jugicambarus), 20, % (fig. 67)

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