Spider Mites from Northwestern

and North Central Mexico

(Acarina: Tetranychidae)

D. M. TUTTLE, E. W. BAKER,

and M. ABBATIELLO

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY • NUMBER 171
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SMITHSONIA N CONTRIBUTION S T O ZOOLOG Y • NUMBE R 17 1

Spider Mites from Northwestern

and North Central Mexico

(Acarina: Tetranychidae)

D. M. Tuttle, E. W Baker,

and M. Abbatielto

SMITHSONIAN INSTITUTION PRESS

City of Washington

1974
ABSTRACT

Tuttle, D. M., E. W. Baker, and M. Abbatiello. Spider Mites from Northwestern and North Central Mexico (Acarina: Tetranychidae). Smithsonian

Contributions to Zoology, number 171, 18 pages, 28 figures, 1974.—Eighteen

genera of spider mites were collected by the authors during July and August

in 1970. Two of these are proposed and described as new, Crotonella and

Atetranychus. A total of sixty-nine species are reported, including thirteen new

species. Descriptions and figures of new species are as follows: Mcgregorella

lantanae, Paraplonobia (Langella) mexicana, Petrobia (Petrobia) waltheria,

Crotonella mazatlana, Mononycheilus wainsteini, M. waltheria, M. tephrosiae,

M. hyptis, Eoteranychus abutilon, E. brickiella, Atetranychus estebaneae, OHgonycnus, (Homonychus) constegia, and Tetranychus (Tetranychus) flechtmanni.

OFFICIAL PUBLICATION DATE is handstamped in a limited number of initial copies and is recorded

in the Institution's annual report, Smithsonian Year. SI PRESS NUMBER 5042. SERIES COVER

DESIGN: The coral Montastrea cavernosa (Linnaeus).

Library of Congress Cataloging in Publication Data

Tuttle, Donald Monroe, 1917—

Spider mites from northwestern and north central Mexico (Acarina: Tetranychidae)

(Smithsonian contributions to zoology, no. 171)

Supt. of Docs, no.: SI 1.27: 171.

1. Tetranychidae. 2. Arachnida—Mexico. I. Baker, Edward William, 1914- joint author.

II. Abbatiello, M., joint author. III. Title. IV. Series: Smithsonian Institution. Smithsonian

contributions to zoology, no. 171.

QL1.S54 no. 171 [QL458.2.T4] 595'.42 73-19794

For sale by the Superintendent ot Documents, U.S. Government Printing Office,

Washington. D.C. 20402. Price 60 cents
Spider Mites from Northwestern

and North Central Mexico

(Acarina: Tetranychidae)

D. M. Tuttle, E. W. Baker,

and M. Abbatiello

Introduction

This survey was undertaken by the authors to

determine the distribution of the spider mites of

the southwestern United States into Mexico and

to check for potential pest species. Tuttle and

Baker (1964; 1968) and Baker and Tuttle (1972)

have published on the spider mites of Arizona and

the southwestern United States, and this survey

is a continuation of these studies. As expected,

most of the mites collected proved to be known

species, mostly from the Southwest. Two new

genera and thirteen new species were found.

The collections were made b"y D. M. Tuttle, M.

Abbatiello, and E. W. Baker unless otherwise

stated; collections were made in the states of Sonora, Sinaloa, Nayarit, Jalisco, Zacatecas, Coahuila,

and Chihuahua. Unless otherwise stated, all collections were made in 1970.

We gratefully acknowledge the plant determinations made for us by Dr. Charles T. Mason, Jr.,

and Mrs. Caryl L. Sagar, Department of Botany,

University of Arizona.

The systematic arrangements are those of Tuttle

and Baker (1968).

D. M. Tuttle, University of Arizona, Yuma, Arizona. E. W.

Baker, V.S. Department of Agriculture, Deltsville, Maryland.

M. Abbalicllo, State University of New York at Farmingdale,

New York.

Genus Psendobryobia McGregor

Pseudobryobia McGregor, 1950:366.—Baker and Tuttle,

1972:2.

Pseudobryobia canescens Baker and Tuttle

Pseudobryobia canescens Baker and Tuttle, 1972:2.

A female and nymph were collected on A triplex

canescens (Pursh) Nuttall, 80 miles south of Ciudad Juarez, 8 August.

Pseudobryobia curiosa (Summers)

Bryobia curiosa Summers, 1958:290.

A female and three nymphs were taken on Zaluztinia triloba (Orton) Persoon, Zacatecas, 2 August.

Pseudobryobia drummondi (Ewing)

Petrobia drummondi Ewing, 1926:143.

Bryobia drummondi.—Pritchard and Baker, 1955:18.

Pseudobryobia drummondi.—McGregor 1950:368.—Baker and

Tuttle, 1972:2.

Pseudobryobia drummondi was collected from

the following host plants: Larrea tridentata (De

Candolle) Coville, Hermosillo, 17 July, Torreon,

5 August, Rancho Grande, 5 August, and 10 miles

east of Torreon, 5 August; Prosopis juliflora

1
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

(Swam) De Candolle, 10 miles east of Torreon,

5 August; and Acacia greggii Gray, 10 miles east

of Torreon, 5 August.

This mite was collected on Brickellia veronicaefolia (Humboldt, Bonpland, and Kunth) Gray,

Zacatecas, 2 August.

Pseudobryobia ephedrae (Tuttle and Baker)

Bryobia ephedrae Tuttle and Baker, 1968:7.

This species was collected on Ephedra trifurca

Torrey, 10 miles south of Chihuahua, 7 August.

Pseudobryobia species

A single specimen was found, a molting nymph

on Baileya pleniradiata Harvey and Gray, 10 miles

south of Chihuahua, 7 August. The nymphal setae

are short, broadly spatulate, and strongly spinate.

The developing nymph possesses setae similar to

those of P. drummondi (Ewing).

Genus Hystrichonychus McGregor

Hystrichonychus McGregor, 1950:272.—Tuttle and Baker,

1968:20.

Hystrichonychus gracilipes (Banks)

Tetranychus gracilipes Banks, 1900:72.

Hystrichonychus gracilipes.—McGregor 1950:39.—Tuttle and

Baker, 1968:27.

Many specimens were collected from Sphaeralcea

orcuttii Rose, Hermosillo, 17 July; S. angustifolia

(Cavara) David Don, Zacatecas, 2 August and

Rancho Grande, 4 August; Helianthus annuus Linnaeus, 160 miles north of Torreon, 6 August; and

Parthenium incanum Humboldt, Bonpland, and

Kunth, 6 August.

Histrichonychus sidae Pritchard and Baker

Histrichonychus sidae Pritchard and Baker, 1955:40.—Tuttle

and Baker, 1964:8.

Collections of this species were taken from Dalea

species, Fresnillo, 3 August; Sida diffusa Humboldt,

Bonpland, and Kunth, Chihuahua, 8 August; and

Sida species, Guadalajara, 31 July.

Hystrichonychus spinosus Tuttle and Baker

Hystrichonychus spinosus Tuttle and Baker, 1968:21.—Baker

and Tuttle, 1972:3.

Genus Monoceronychus McGregor

Monoceronyehus McGregor, 1945:100 Tuttle and Baker,

1968:3.

Monoceronychus californicus McGregor

Monoceronychus californicus McGregor, 1945:100.—Tuttle

and Baker, 1968:41.

A collection of M. californicus was collected

from Monanthochloe littoralis Engelmann, Los

Mochis, 24 July.

Genus Mcgregorella Baker and Tuttle

Mcgregorella Baker and Tuttle, 1972:5.

Mcgregorella incana Baker and Tuttle

Mcgregorella incana Baker and Tuttle, 1972:6.

Records of this species from Mexico include:

Aloysia wrightii (Gray) Heller, Chihuahua, 8

August; Coldenia canescens De Candolle, Chihuahua, 8 August; and Sida diffusa Humboldt, Bonpland, and Kunth, Chihuahua, 8 August.

Mcgregorella lantanae, new species

FIGURES 1-4

This species differs from M. incana Baker and

Tuttle in having lightly spinate dorsal body setae

and a different pattern of leg setae. The dorsal

body setae are much longer than those of M. trifida

Baker and Tuttle.

FEMALE.—Rostrum short; stylophore somewhat

elongate and slightly edentate anteriorly; peritreme

anastomosing distally. Dorsal body surface with

light reticulate pattern; propodosoma and hysterosoma separated by transverse striae; eyes appear

to be single; anterior to second pair of DC setae

another series of transverse striae. All dorsal body

setae, except short anterior propodosomal setae,

long, strong, lightly serrate, and set on strong elon-
NUMBER 171

FIGURES 1-4.—Mcgregorella lantanae, new species: 1, dorsum of female; 2, dorsum of male; 3, leg

I of male; 4, leg I of female.

gate tubercles as figured. Most leg setae short;

tibial setae longer than others; dorsal femoral and

genual setae short, stout, and serrate; tarsus III

with distal set of duplex setae, short and subequal

in length, the solenidion distal; tarsus IV similar

but duplex setae longer. Setal formula as follows:

1. 2-1-6-4-7 + 1-3 posterior to duplex

2. 2-1-5-4-6-3 posterior to duplex

3. 1-1-2-3-5-4 posterior to duplex

4. 1-1-2-3-5-4 posterior to duplex

Length of body 383jx; including rostrum 478^.

MALE.—Similar to female but body setae rela-
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

tively shorter; anterior propodosomal setae relatively longer; body divided by transverse suture

anterior to first pair of DC setae and posterior to

third pair. Eyes double. Duplex setae on tarsi

I-IV. Length of body 319u; including rostrum

415u.

HOLOTYPE.—Female, USNM 3537, ex Lantana

velutina Martens and Galeottit, Mazatlan, 26 July.

PARATYPES Six females and one male with the

above data.

Genus Paraplonobia Wainstein

Aplonobia (Paraplonobia) Wainstein, 1960:140.

Paraplonobia.—Tuttle and Baker, 1968:48.

Subgenus Paraplonobia (Paraplonobia) Wainstein

Aplonobia (Paraplonobia) Wainstein, 1960:140.

Paraplonobia (Paraplonobia).—Tuttle and Baker, 1968:50.

Paraplonobia (Paraplonobia) hilaria Tuttle and

Baker

Paraplonobia (Paraplonobia) hilaria Tuttle and Baker, 1968:

53.

Specimens of this species were taken on Hilaria

mutica (Buckley) Bentham, Torreon, 5 August.

Paraplonobia (Paraplonobia) trident Tuttle and

Baker

Paraplonobia (Paraplonobia) tridens Tuttle and Baker, 1968:

50.

This species was collected from Hilaria mutica

(Buckley) Bentham, Torreon, 5 August; and Tridens puchellus (Humboldt, Bonpland, and Kunth)

Hitchcock, 8 miles south of Ciudad Juarez, 8

August.

Paraplonobia (Paraplonobia) species

A single larva was found on Prosopis juliflora

(Swartz) De Candolle, Hermosillo, 18 July. The

long leg setation is distinctive compared to other

species.

Subgenus Paraplonobia (LangeUa) Wainstein

Paraplonobia (LangeUa) Wainstein, 1961:607.—Tuttle and

Baker, 1968:54.

Paraplonobia (Langella) artemisia Baker and

Tuttle

Paraplonobia (Langella) artemisia Baker and Tuttle, 1972:10.

This mite was taken from Artemisia ludoviciana

Nuttall, Chihuahua, 8 August; and Flourensia

cernna De Candolle, Chihuahua, 8 August.

Paraplonobia (Langella) euphorbiae Tuttle and

Baker

Paraplonobia (Langella) euphorbiae Tuttle and Baker, 1964:

11; 1968:56.

Mites of this species were collected from Acacia

greggii Gray, 10 miles east of Torreon, 5 August;

Scleropogon brevifolius Philippi, Chihuahua, 5 August; and Tidestromia lanuginosa (Nuttall) Standley, Cuencame, 4 August, and Torreon, 5 August.

Paraplonobia (Langella) mexicana, new species

FIGURE 5

This species appears to be closely related to P.

(L.) boutelouae Tuttle and Baker, differing only

in the setal count of all legs, in having somewhat

larger dorsal body setae, and in having the propodosomal shield striae shorter and much more

strongly tuberculate.

FEMALE.—Rostrum short and broad; stylophore

broadly rounded anteriorly; peritremes anastomosing distally. Dorsal body setae, except for the first

pair of propodosomals, broadly rounded, small and

serrate; first pair of propodosomals more elongate.

Propodosomal shield with longitudinal design of

somewhat wrinkled, short striae with obvious tubercles or lobes. Striae of hysterosoma transverse,

widely separated, wrinkled, and without tubercles

or lobes. Legs typical for genus, leg I longer than

others. The leg setal count is as follows:

1. 2-1-6-5-11 + 1-12+1

2. 2-1-5-4-7-$+2

S. 1-1-2-4-7-9+1

4. 1-1-3-5-8-9+1

In contrast, the setal counts of the legs of P.

boutelouae Tuttle and Baker are:

1. 2-1-4-3-8+1-12+2

2. 2-1-4-3-7-6

3. 1-1-3-2-8-10

4. i-i-3
NUMBER 171

FIGURES 5-10—-Paraplonobia (LangeUa) mexicana, new species: 5, dorsum of female. Petrobia

(Petrobia) waltheria, new species: 6, dorsum of female; 7, hystersomal striae and lobes.

Crotonella mazottana, new species: 8, dorsum of female; 9, peritreme; 10, tubercle and base of

seta DQ.
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

The outer coxal setae of legs I and II of P.

boutelouae are much stouter than those of the

new species. Length of body 472^i; including rostrum 542u.

MALE.—Not known.

HOLOTYPE.—Female, USNM 3538, ex Prosopis

pubescens Bentham, Los Mochis, 24 July.

PARATYPE.—Female with the same data.

Paraplonobia (Langella) prosopsis (Tuttle and

Baker)

Aplonobia prosopis Tuttle and Baker, 1964:13.

Paraplonobia (Langella) prosopis.—Tuttle and Baker, 1968:

56.

Specimens were taken from the following plant

hosts in Mexico: Cercidium microphylum (Torrey) Rose and Johnson, Alamos, 21 July; and

Prosopis juliflora (Swartz) De Candolle, Ciudad

Obregon, 22 July, 2 miles south of Zacatecas, 2

August, 10 miles south of Chihuahua, 7 August,

and 80 miles south of Ciudad Juarez, 8 August.

Paraplonobia (Langella) species

A single nymph was collected on Acacia vernicosa Standley, Fresnillo, 3 August. The dorsal striation pattern is distinctive in forming starlike

crenulated lobes.

Genus Georgiobia Wainstein

Georgiobia Wainstein. 1960:138—Tuttle and Baker, 1968:

58.—Baker and Tuttle, 1972:12.

Georgiobia ambrosiae Tuttle and Baker

Georgiobia ambrosiae Tuttle and Baker, 1968:59.

This species was collected from Ambrosia confertiflora (De Candolle) Rydberg, Hermosillo, 17

July.

Georgiobia haplopappi Tuttle and Baker

Georgiobia haplopappi Tuttle and Baker. 1968:62.—Baker

and Tuttle, 1972:12.

Georgiobia haplopappi was collected from the

following host plants in Mexico: Ambrosia confer ti flora (De Candolle) Rydberg, Hermosillo, 17

July; Parthenium hysterophorus Linnaeus, Los

Mochis, 24 July; Ruellia nudiflora (Engelmann

and Gray) Urban, Torreon, 5 August; and Gaillardia pulchella Fougeroux, Ciudad Juarez, 9

August.

Genus Petrobia Murray

Petrobia Murray, 1877:118.—Wainstein, 1960:133.

Subgenus Petrobia (Petrobia) Murray

Petrobia Murray, 1877:118.—Wainstein, 1960:133.—Tuttle

and Baker, 1968:71.

Petrobia (Petrobia) waltheria, new species

FIGURES 6, 7

This species is distinctive in having short, broad

serrate dorsal body setae. This mite would key

out to P. latens (Miiller) in Pritchard and Baker

(1955), but differing in the type of setae.

FEMALE.—Rostrum short and broad; stylophore

narrowing anteriorly and rounded; peritremes

slightly anastomosing distally. Propodosomal shield

pebbled; hysterosoma with irregular transverse

striae bearing rounded lobes. Dorsal body setae

short, distinctly broadening distally, serrate and

about equal in length except for the Pj and DC3

setae; humeral setae smaller. Leg I as long as body;

II and III shorter; IV longer than II and III but

not as long as I. Leg setae short, slender, proximal

setae stronger and distinctly serrate. Length of

body $00\i; including rostrum 351ji.

MALE.—Not known.

HOLOTYPE.—Female, USNM 3539, ex Waltheria

americana Linnaeus, Mazatlan, 26 July.

PARATYPE.—Female with the above data.

Specimens were also collected in Mazatlan, 26

July on Cynodon dactylon (Linnaeus) Persoon and

Sida species.

Genus Neotrichobia Tuttle and Baker

Neotrichobia Tuttle and Baker, 1968:73.—Baker and Tuttle,

1972:14.

Neotrichobia arizonensis Tuttle and Baker

Neotrichobia arizonensis Tuttle and Baker, 1968:74.—Baker

and Tuttle, 1972:14.
NUMBER 171

FIGURES 11—16.—Mononychellus wainsteini, new species: 11, dorsum of female; 12, aedeagus;

13, dorsal seta. Mononychellus waltheria, new species: 14, dorsum of female; 15, dorsal seta.

Mononychellus tephrosiae, new species: 16, dorsum of female.
8

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

This species was collected in Mexico on Allionia

incarnata Linnaeus, Hermosillo, 18 July, 10 miles

south of Chihuahua, 7 August; Boerhaavia species,

Torreon, 4 August; Palafoxia linearis (Cavara)

Lagasca, 8 miles east of Torreon, 5 August; and

Scleropogon brevifolius Philippi, Chihuahua, 8

August.

Genus Crotonella, new genus

The dorsal striae form spicules; the dorsal setae

are marginal except for DCX_3; the empodia are

split distally; and the duplex setae are distal, and

approximate on tarsus I.

TYPE-SPECIES.—Crotonella mazatlana, new

species.

Crotonella mazatlana, new species

FIGURES 8-10

FEMALE.—Rostrum short and broad; stylophore

slightly edentate anteriorly; peritremes slightly

hooked distally. Dorsum of body covered with

spicules formed by the striae; dorsum divided into

three units by transverse striae between propodosoma and hysterosoma, and posterior to DC! and

humeral setae. Venter of body with normal striae;

ventral body setae long, slender, and simple. Dorsal body setae, other than DC1-3, marginal and on

prominent tubercles which basally possess fingerlike projections except for propodosomal setal

bases which have blunt processes. Setae broad at

base and tapering distally, with strong blunt serrations basally.

Leg I setal pattern as follows: tarsus with 2 setae

proximal to duplex setae; tibia with 3 slender setae;

genu with 3 strong and 1 slender setae; femur

with 2 strong and 2 slender setae; trochanter with

1 slender seta; and coxa with 2 slender setae.

Leg II pattern as follows: tarsus with 1 slender

seta proximal to duplex setae; tibia with 2 strong

and 2 slender setae; genu with 2 strong and 2

slender setae; femur with 1 strong and 2 slender

setae; trochanter with 1 long slender seta; and

coxa with 2 long slender setae.

Length of body 287u; including rostrum 238u.

HOLOTYPE.—Female, USNM 3540, ex Croton

species, Mazatlan, 26 July.

PARATYPE.—Female, broken, with the above

data.

Genus Allonychus Pritchard and Baker

Allonychus Pritchard and Baker, 1955:137.

Allonychus littoralis (McGregor)

Allonychus littoralis,—Baker and Pritchard, 1963:313.

This species was taken from Quercus reticulatus

Humbold and Bonpland, Zapotlanejo, 30 July. A

male and a female were found. The ventral hairs

of empodium I of the male form a clawlike structure; the ventral hairs of empodia 11-IV of the male

and I-1V of the female are free. The aedeagus appears to be slender and sigtnoid.

Genus Mononychellus Wainstein

Mononychus Wainstein, 1960:198 [preoccupied].

Mononychellus Wainstein, 1971:589.

Mononychellus wainsteini, new species

FIGURES 11-13

This species is distinctive in that the dorsal

body setae reach to or about to the bases of the

setae of the next row; the seta are subequal in

length. The head of the aedeagus is rounded and

equally hooked.

FEMALE.—Rostrum short and strong; stylophore

rounded anteriorly; peritreme ending in simple

bulb. Dorsal body setae reaching about to or to

the bases of setae of next row; setae subequal in

length, strong, serrate, parallel sided, and tapering

to point distally. Striae with broad lobes, longitudinal between DC3 and V-like posterior to DC4.

Ventral body setae normal and slender. Length

of body 363u, including rostrum 447^.

MALE.—Dorsal body setae similar to those of

female, but shorter. Aedeagus typical for genus

but head with equal angulations dorsally and ventrally. Specimen broken and not measurable.

HOLOTYPE.—Female, USNM 3541, ex Eysenhardlia species, Zapotlenejo, 30 July.

PARATYPES.—One male and seven females with

the above data.
NUMBER 171

18

22

FIGURES 17—22.—Mononychellus hyptis, new species: 17, dorsum of female; 18, two lateral views

of acdeagus. Eotetranychus abutilon, new species: 19, peritreme; 20, aedeagus. Eotetranychus

brickellia, new species: 21, acdeagus; 22, peritreme of male.

This species is similar to Mononychellus siccus

(Pritchard and Baker), which was collected from

a leguminous shrub at Phoenix, Arizona. It differs

in that the setae are shorter and much stouter,

and that the head of the aedeagus of Af. siccus

appears to be rounded or knoblike.

Mononychellus tcaltheria, new species

FIGURES 14, 15

This mite was first believed to be Mononychellus

planki (McGregor) (McGregor, 1950:300; Pritchard and Baker, 1955:148). However, the dorsal

body setae of the female broaden distally, whereas

in Af. planki these setae narrow distally, and the

mite is well out of the known range of Af. planki.

It is here described as new.

FEMALE.—In general, very closely allied to Af.

planki in having the stylophore rounded anteriorly,

the peritreme knobbed distally, and in having a

similar reticulate dorsal striation pattern. The dorsal body setae of Af. planki are slender and nar-
10 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

rowing distally; those of the new species are

obviously broadened distally, although about equal

length to those of M. planki. Length of body 319^;

including rostrum 408u.

HOLOTYPE.—Female, USNM 3542, ex Waltheria

americana Linnaeus, Mazatlan, 26 July.

PARATYPES.—Ten females with the above data.

One female was also taken on bean leaf, La

Calera, Nicaragua, 12 January 1960 by F. A.

Estrada.

Mononychellus tephrosiae, new species

FIGURES 16

This species is close to M. tanajoa (Bondar)

(Bondar, 1938:441; Flechtmann and Baker, 1970:

160), differing in that the dorsocentral hysterosomal

setae are at least more than one-half as long as

the distance to the bases of the setae of the next

row.

FEMALE.—Rostrum short and broad; stylophore

broadly rounded anteriorly; peritreme ending in

a simple bulb. Dorsal striation pattern as figured,

being longitudinal between DC3 setae. All dorsal

body setae long, expanded distally, and reaching

half the distance or more to the bases of the setae

of the next row; all setae of about equal length.

Length of body 319u; including rostrum 402^.

HOLOTYPE.—Female, USNM 3543, ex Tephrosia

talpa Sereno Watson, Tequila, 26 July.

PARATYPES.—Nine females with the above data.

Other specimens studied are from Olneya tesota

Gray, Hermosillo, 17 July, and from Solarium

rostratum Dunal, Guadalajara, 31 July.

Mononychellus hyptis, new species

FICURES 17, 18

This species has the short setae of M. caribbeanae

(McGregor), but these dorsal body setae are much

stronger, and the males are present. Climatically,

the mite is found on the dry west coast of Mexico.

FEMALE.—Stylophore rounded anteriorly; peritremes with simple knob distally. Propodosomal

setae 1 and 3 longer than 2, all three pairs obviously widened distally. Propodosomal striae

longitudinal, broadly lobed, and in an irregular

pattern. Hysterosomal setae Dx-D4 small, broadening distally; marginals longer and becoming progressively longer posteriorly and also broadening

distally. Striae of hysterosoma irregular, longitudinal between setae D3, and with broad lobes. Venter

with striae and setae typical for genus. Length of

body 415^; including rostrum 478^.

MALE.—Setae typical for males of the genus,

being long and more slender than in females, but

with same length pattern. Aedeagus typical for

males of genus, being hooked almost equally distally and with slight indentation if seen properly.

Length of body 338u; including rostrum 415(i.

HOLOTYPE.—Female, USNM 3544, ex Hyptis

species, Zapotlanejo, 30 July.

PARATYPES.—Two males and six females with

the above data.

Genus Eotetranychus Oudemans

Eotetranychus Oudemans, 1931:224 Pritchard and Baker,

1955:138.—Tuttle and Baker, 1968:85.

Eotetranychus abutilon, new species

FICURES 19, 20

This mite keys out to Eotetranychus perplexus

(McGregor) in the key to the females in Pritchard

and Baker (1955). The aedeagus of the males differ

considerably.

MALE.—Sensillum of palpus small, about as long

as broad. Peritremes slightly hooked distally.

Tarsus I with clawlike empodium; other empodia

with free hairs. Setal count of legs as follows:

1. 2-1-8-5-8 + 3-4+4 (proximal to duplexes)

2. 2-1-7-5-8-1 + 1

3. 1-1-5-4-5-1 + 1

4. 1-1-4-4-6-5+1

Aedeagus sharp sigmoid, curving ventrally. Length

of body 223u; including rostrum 287u.

FEMALE.—With characters of Eotetranychus perplexus (McGregor). Tibia II with eight setae;

tarsus I with four setae proximal to duplex setae.

Peritreme slightly hooked. Sensillum of palpal

tarsus about 2 times as long as broad. Striae of

genital plate and area anterior to plate transverse.

Setal count of legs as follows:
NUMBER 171 11

1. 2-1-10-5-9+1-4 (proximaU)

2. 2-1-7-5-8-1 + 1 (proximals)

S. 1-1-4-4-6-2+1 (dorsals)

4. 1-1-4-4-6-2+1 (dorsals)

Length of body 306u; including rostrum 382u.

HOLOTYPE.—Male, USNM 3545, ex Abutilon

species, Zapotlanejo, 13 July.

PARATYPES.—Three males and eight females with

the above data.

Eotetranychus brickellia, new species

FIGURE 21, 22

This species is similar to Eotetranychus perplexur (McGregor) except that the head of the

aedeagus is much smaller.

MALE.—Terminal sensillum of palpus minute;

peritreme with simple knob distally. Tibia I with

9 tactile setae; tibia II with 8 tactile setae. The

aedeagus is strong, bent ventrally distally, and

with a small distal head forming a dorsal ventral

angulation. Length of body 255u; including rostrum 319u.

FEMALE.—Terminal sensillum of palpus strong,

about twice as long as broad; peritreme ending in

simple knob. Tibia I with 9 tactile setae; tibia II

with 8 tactile setae; tarsus I with 4 tactile setae

proximal to duplex setae. Striae of genital area

transverse. Length of body 287u; including rostrum

383u.

HOLOTYPE.—Male, ex Brickellia veronicaefolia

(Humboldt, Bonpland, and Kunth) Gray, Fresnillo, 3 August.

PARATYPES.—Two males and two females with

the above data.

Two males and eight females were also in the

above series.

Eotetranychus species

These mites were collected from Polanisia species,

Mazatlan, 28 July and Sida species, Zapotlanejo,

30 July.

The females key out to E. perplexus (McGregor), based on the setal count of tibia II and

tarsus I. Males were not present, and should be

studied for specific determination.

Eotetranychus deflexus (McGregor)

Tetranychus deflexus McGregor

Eotetranychus deflexus.—Pritchard and Baker, 1955:206.

The peritremes of the females are straight distally. There are 8 tactile setae on tibia II. These

characters agree with those of E. deflexus, which

have been found on Symphoricarpos oreophilus

Gray and S. palmeri G. N. Jones in Arizona

(Tuttle and Baker, 1968:90). No males were found.

Specimens were collected in Mexico from Celtis

iguanaea (Jacquin), Sargent, Alamos, 26 July; and

Crescentia alata Humboldt, Bonpland, and Kunth,

Mazatlan, 28 July.

Eotetranychus ecclisis? Pritchard and Baker

Eotetranychus ecclisis Pritchard and Baker, 1955:210.

Only females were found. These key out to E.

ecclisis, which was described from Artemisia ludoviciana subspecies mexicana (Willdenow) Keck,

Cuernavaca. We have taken it on Quercus reticulata Humboldt and Bonpland, Zapotlanejo, 30

Eotetranychus fallugia Tuttle and Baker

Eotetranychus fallugia Tuttle and Baker, 1968:88.

This species was taken on Cassia wislizeni Gray,

Chihuahua, 7 August.

Eotetranychus fremonti Tuttle and Baker

Eotetranychus fremonti Tuttle and Baker, 1964:26.

Specimens of this species were collected from

Acacia constricta Bentham, Torreon, 6 August;

Mentzelia involucrata Watson, 10 miles southwest

of Chihuahua, 7 August; Populus fremontii Watson,

Los Mochis, 24 July; Populus tremuloides Michaux,

Ciudad Obregon, 22 July; Prosopis juliflora

(Swartz) De Candolle, 10 miles southwest of Chihuahua, 7 August; and Ruellia nudiflora (Engelmann and Gray) Urban, Torreon, 6 August.

Eotetranychus lewisi (McGregor)

Tetranychus lewisi McGregor, 1943:147.

Eotetranychus lewisi.—Tuttle and Baker 1968:91.

Identifications based on males and females rep-
12

resent material from Ceiba acuminata (Watson)

Rose, Mazatlan, 26 July; Ipomoea species, Guadalajara, 31 July; and Mimosa laxiflora Bentham,

Hermosillo, 19 July. The following are believed

to be E. leivisi because of the hosts, distribution,

and female morphology (no males were found):

Bebbia juncea (Bentham) Greene, Hermosillo, 19

July; Cardiospermum corindum Linnaeus, Hermosillo, 19 July; Brickellia californica (Torrey and

Gray) Gray, Los Mochis, 23 July; Croton sonorae

Torrey, Hermosillo, 19 July; and Mimosa biuncifero Bentham, Fresnillo, 3 August.

A series of females was collected in which the

peritreme is not strongly hooked or bent as in

the typical E. lewisi. These are tentatively placed

here. Host plant records include: Abutilon malacum Watson, Chihuahua, 8 August; Ambrosia confertifolia (De Candolle) Rydberg, Tepic, 26 July;

Croton ciliatoglandulosus Ortega, Guadalajara, 31

July; Croton species, Jimenez, 4 August; and

Heterotheca species, Guadalajara, 31 July.

Eotetranychus malvastris (McGregor)

Tetranychus malvastris McGregor, 1950:290.

Eotetranychus malvastris.—Tuttle and Baker, 1964:20.—Tuttle and Baker, 1968:91.

Males and females were in a series on Abutilon

species, Hermosillo, 18 July, Topolobampo, 24

July, and Ma/atlan, 26 July; Pluchea odorata (Linnaeus) Case, Mazatlan, 26 July; and Sida species,

Mazatlan, 26 July.

Eotetranychus prosopis Tuttle and Baker

Eotetranychus prosopis Tuttle and Baker, 1964:24; 1968:89.

Collections in Mexico included two additional

host records: Acacia vernicosa Standley, Fresnillo,

3 August; and Hibiscus palustris Linnaeus, Ciudad

Obregon, 21 July.

Eotetranychus vaughni Baker and Pritchard

Eotetranychus vaughni Baker and Pilchard, 1962:321.

Specimens were taken from Guazuma ultnifolia

Lamarck, Ciudad Obregon, 21 July.

The figure given by Baker and Pritchard for

this species shows the aedeagal head with sharp

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

angulations anteriorly and posteriorly. If seen from

a slightly different angle, the anterior portion of

the knob is rounded.

Eotetranychus yumensis (McGregor)

Tetranychus yumensis McGregor, 1934:256.

Eotetranychus yumensis.—Tuttle and Baker, 1968:97.

Specimens were collected from the following

host plants: Cassia covesii Gray, Ciudad Obregon,

22 July; Jatropha cinerea (Ortega) Mueller, Jean

(of Aargau) Ciudad Obregon, 22 July; Jatroph

cardiophylla (Torrey) Mueller, Jean (of Aargau),

22 July and Hermosillo, 27 July; Olneya tesota

Gray, Hermosillo, 19 July; Pluchea odorata (Linnaeus) Cassini, Topolobampo, 24 July; Sarcostemma pannosum (Decaisne) Schlechter, Topolobampo, 21 July; and Trichloris crinita (Lagasca) Perodi, Torreon, 5 August.

Eotetranychus mastichi DeLeon

Eotetranychus mastichi DeLeon, 1957:111 Tuttle and Baker,

1968:90.

A series was taken on Berlandiera lyrata Bentham, Fresnillo, 3 August.

Eotetranychus species

Several females were collected from Bouteloua

curtipendula (Michaux) Torrey, Rancho Grande,

4 August. These are similar to E. potentillae Tuttle

and Baker, but differ in having long solenidia on

the tarsi. Males should be studied.

Genus N eotetranychus Tragardh

Neotetranychus TrMgardh, 1915:23 Tuttle and Baker, 1968:

104.

N eotetranychus gloriosus Estebanes and Baker

Neotetranychus gloriosus Estebanes and Baker, 1968:73.

This mite was collected by the authors from

Agave tequilana Weber, Mazatlan, 28 July.

The dorsal speculate pattern of the striae is not

typical, but the ventral body striae do form a

basket-weave pattern.
NUMBER 171 IS

Genus Schizotetranychus Tragardh

Schizotetranychus Tragardh, 1915:277.—Tuttle and Baker,

1968:92.

One collection of S. nugax was made from

Hilaria mutica (Buckley) Bentham, Torreon, 5

August.

Schizotetranychus boutelouae Tuttle and Baker

Schizotetranychus boutelouae Tuttle and Baker, 1968:96.

Collections from Mexico include the following

host plants: Commelina dianthifolia Delile, 8 miles

north of Chihuahua, 8 August; and Stipa eminent

Cavara, Rancho Grande, 4 August.

Schizotetranychus celtides Tuttle and Baker

Schizotetranychus celtides Tuttle and Baker, 1968:93.

Specimens were found on Sporobulus flexuosus

(Thurber) Rydberg, Torreon, 5 August; and Tridens pulchellus (Humboldt, Bonpland, and Kunth)

Hitchcock, Chihuahua, 7 August.

Schizotetranychus elymus McGregor

Schizotetranychus elymus McGregor, 1950:310—Tuttle and

Baker, 1968:104.

A series was taken from Panicum obtusum Humboldt, Bonpland, and Kunth, Rancho Grande, 4

August and Fresnillo, 13 August.

Schizotetranychus eremophilus McGregor

Schizotetranychus eremophilus McGregor, 1950:311.—Pritchard and Baker, 1955:251.—Tuttle and Baker, 1964:32.—

Tuttle and Baker, 1968:95.

This species was collected from Cynodon dactylon (Linnaeus) Persoon, Mazatlan, 26 July and

20 miles south of Torreon, 4 August.

Schizotetranychus montanae Tuttle and Baker

Schizotetranychus montanae Tuttle and Baker, 1968:99.

This species was found on Pappophorum mucronulatum Nees, Torreon, 5 August.

Schizotetranychus nugax Pritchard and Baker

Schizotetranychus nugax Pritchard and Baker, 1955:264.—

Tuttle and Baker, 1968:95.

Schizotetranychus species

A series of these mites was found on two grasses

in Mexico: Scleropogon brevifolius Philippi, Chihuahua, 8 August; and Setaria macrostachya Humboldt, Bonpland, and Kunth, Chihuahua, 8 August.

This mite is similar to S. eremophilus McGregor

in having the same general dorsal setal pattern.

However, the Mexican mites have much shorter

dorsal body setae, which do not or barely reach

the bases of the setae of the next row.

Genus Anatetranychus Womersley

Anatetranychus Womersley, 1940:261 Tuttle and Baker,

1968:107.

Anatetranychus daleae Tuttle and Baker

Anatetranychus daleae Tuttle and Baker, 1968:110.

Specimens were collected from Dalea emoryi

Gray, Hermosillo, 8 July.

Anatetranychus albiflora Tuttle and Baker

Anatetranychus albiflora Tuttle and Baker, 1968:113.

Collections of this species were taken from

Brickellia species, Fresnillo, 3 August; Dalea tuberculata Lagasca, Fresnillo, 3 August, and Rancho

Grande, 3 August; Dalea species, Fresnillo, 3 August; and Ipomoea species, Fresnillo, 3 August.

Genus Atetranychus, new genus

This genus is similar to Anatetranychus but

differs in having a single set of duplex setae on

tarsus I, and in that the dorsal hysterosomal striae

and the lateral striae of the propodosoma are very

widely separated and irregular. The empodia are

clawlike and without ventral hairs. The propodosomal shield is pebbled. The dorsal body setae are

long and set on prominent tubercles.

TYPE-SPECIES.—Atetranychus estebanesae, new

species.
14

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

FIGURES 23-28.—Atetranychus estebanesae, new species: 23, dorsum of female; 24, tarsus of leg

I of female; 25, aedeagus; 26, dorsal seta. Oligonychus (Homonychus) constegia, new species:

27, leg I of female. Tetranychus (Tetranychus) flechtmanni, new species: 28, aedeagus.

Atetranychus estebanesae, new species

FIGURES 23-26

This species is named for Srta. Maria Luisa

Estebanes of the Instituto de Biologia of the Universidad Nacional Autonoma de Mexico.

FEMALE.—Rostrum elongate; stylophore broadly

rounded anteriorly; peritreme ending in large

anastomosing ball. Dorsal body setae long, slender,

serrate, and expanding distally, set on prominent

tubercles. Propodosomal shield covered with lobes,

in general these lobes form a longitudinal pat-
NUMBER 171 15

tern; laterad of the shield striae widely separated

and angularly broken. Striae of hystersoma, in

general, transverse and similar to those of propodosoma, widely separated and angularly broken.

All dorsal setal tubercles surrounded by area of

small lobes, those between DC4 setae contiguous.

Ventral striae more or less similar to dorsal striae.

Leg setal pattern as follows:

1. 2-1-9-4-13+1-19+duplex

2. 2-1-5-4-9-15+duplex

S. 1-1-5-4-9-14

4. 1-1-5-4-11-14

Length of body 666u; including rostrum 826u.

MALE.—In general similar to female in setal

patterns. Entire dorsum with pattern of tubercles,

those on propodosoma forming longitudinal patterns, those on hysterosoma transverse pattern.

Legs I broken off; leg II with single set of duplex

setae. Aedeagus long, strong, simply curved, and

attenuate distally. Length of body 35lu; including

rostrum 382u.

HOLOTYPE.—Female, USNM 3547, ex Verbena

canescens Humboldt, Bonpland, and Kunth, Fresnillo, 3 August.

PARATYPES.—A female and a male with the above

data.

Genus Oligonychus Berlese

Oligonychus Berlese, 1886:24.—Tuttle and Baker, 1968:116.

Subgenus Oligonychus (Oligonychus) Berlese

Oligonychus (Oligonychus).—Wainstein, 1960:203.—Tuttle

and Baker. 1968:118.

Oligonychus (Oligonychus) ununguis (Jacobi)

Tetranychus ununguis Jacobi, 1905:239.

Oligonychus (Oligonychus) ununguis*

1968:118.

-Tuttle and Baker,

The spruce spider mite was collected from Thuja

occidentalis Linnaeus, Ciudad Obregon, 22 July.

Oligonychus (Oligonychus) species

Only females were collected from Setaria macrostachya Humboldt, Bonpland, and Kunth, Chihuahua, 8 August. Males are needed for specific

identification.

Subgenus Oligonychus (Homonychus) Wainstein

Oligonychus (Homonychus) Wainstein, 1960:216.—Tuttle and

Baker, 1968:119.

Oligonychus (Homonychus) conostegia, new

species

FIGURE 27

This species is similar to O. (H.) gambelii Tuttle

and Baker (1968:120), differing in that there are

four tactile setae and one solenidion proximal to

the duplex setae on tarsus I. O. gambelii has only

two tactiles and one solenidion proximal to the

duplex setae.

FEMALE.—Rostrum short and broad; stylophore

broadly rounded; peritreme ending in a simple

bulb. Dorsal body striae with strong, rounded

lobes; striae longitudinal between DC3 setae. Dorsal body setae long, strong, serrate, and on small

tubercles. All setae of about equal length except

for short DC5 setae. Tarsus I with four tactiles

and one solenidion proximal to duplex setae. Setal

pattern of legs as follows:

1. 2-1-8-5-7+1-4+1 (proximals)

2. 2-1-6-5-5-3+1

3. 1-1-2-2-5-5+1

4. 1-1-1-2-5-5+1

Length of body 255u; including rostrum 383u.

MALE.—Not known.

HOLOTYPE.—Female, USNM 3548, ex Conostegia

xalapensis (Bonpland) David Don, Tepic, 28 July.

Oligonychus (Homonychus) platani? (McGregor)

Oligonychus (Homonychus) platani (McGregor), 1950:349.—

Tuttle and Baker, 1968:120.

This mite is similar in many respects to O. (H.)

platani (McGregor), but the tactile setae of the

duplex setae are longer. Males were not found.

This species was collected on Boerhaavia species,

Torreon, 2 August.

Subgenus Oligonychus (Pritchardinychus)

Wainstein

Oligonychus (Pritchardinychus) Wainstein, 1960:217.
16 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Oligonychus (Pritchardinychus) species

A series of females was collected from the following plants in Mexico: Euphorbia albomarginata

Torrey and Gray, 120 miles north of Chihuahua,

8 August; Hyptis species, Guadalajara, 31 July;

Pithecolobium duke (Roxburgh) Bentham, Topolobampo, 24 July; Setaria macrostachya Humboldt,

Bonpland, and Kunth, Chihuahua, 8 August; and

Sida species, Chihuahua, 8 August.

Subgenus Oligonychus (Reckiella) Turtle and

Baker

Oligonychus (Reckiella) Tuttle and Baker, 1968:122.

Oligonychus (Reckiella) mexicanus (McGregor

and Ortega)

Paratetranychus mexicanus McGregor and Ortega, 1953:3.

Paratetranychus indicus Hirst.—Pritchard and Baker, 1955:

354 [misidentification].

Oligonychus mexicanus.—Estebanes and Baker, 1968:95.

Two collections were made by D. M. Tuttle on

Zea mays var. saccharata (Sturtevant) Bailey, Torreon, 26 July and Chapingo, 2 July 1969.

This species was placed in synonymy with O.

indicus by Pritchard and Baker in 1955. Examination of Indian material in the British museum

proved that the two species were different.

Oligonychus (Reckiella) species

Without males no specific determinations can

be made. A series of females was found on the

following plants: Artemisia bigelovii Gray, Torreon, 5 August; Chenopodium album Linnaeus,

Zapotlanejo, 30 July; Cynodon dactylon (Linnaeus)

Persoon, Mazatlan, 26 July; Gossypium hirsutum

Linnaeus, 8 miles south of Torreon, 5 August;

Helianthus annuus Linnaeus, 107 miles south of

Torreon, 6 August; Parthenium incanum Humboldt, Bonpland, and Kunth, Torreon, 6 August;

Setaria lutescens (Weigel) Hubbard, Zapotlanejo,

30 July; Solarium rostratum Dunal, Guadalajara,

31 July; Sporobolus flexuosus (Thurber) Rydberg,

Torreon, 5 August; Verbena Carolina Linnaeus,

Guadalajara, 31 July; and Zea mays var. saccharata

(Sturtevant) Bailey, Guadalajara, 31 July.

Genus Tetranychus Dufour

Tetranychus Dufour, 1832:276.

Subgenus Tetranychus (Tetranychus) Dufour

Tetranychus (Tetranychus) Dufour, 1636:276.—Wainstein

1960:149.—Tuttle and Baker, 1968:126.

Tetranychus (Tetranychus) dnnabarinus

(Boisduval)

Acarus dnnabarinus Boisduval, 1867:88.

Tetranychus (Tetranychus) dnnabarinus.—Tuttle and Baker,

1968:129.

Specimens ol this mite were taken on Gossypium

hirsutum Linnaeus, Santa Rosa, Valle del Fuerte,

4 August; Mangifera indica Linnaeus, Acoponeta,

28 July; Morus rubra Linnaeus, San Miguel, east

of Torreon, 5 August; and Rosa delecta Rehder,

Guadalajara, 31 July.

Tetranychus (Tetranychus) desertorum Banks

Tetranychus desertorum Banks, 1900:76.—Tuttle and Baker,

1968:126.

Records of this species from material collected

in 1970 included the following host plants: Ambrosia confertiflora De Candolle, Ciudad Obregon,

22 July; Buddleia scordioides Humboldt, Bonpiand, and Kunth, Rancho Grande, 4 August; Cassia

crotolariodes Kunth, Fresnillo, 3 August; Encelia

farinosa Gray, Hermosillo, 19 July; Marrubium

vulgare Linnaeus, Zacatecas, 2 August; Mentzelia

pumila (Nuttall) Torrey and Gray, Hermosillo,

18 July; Parthenium hysterophorus Linnaeus, Los

Mochis, 24 July; Ruellia nudiflora (Engelmann

and Gray) Urban, Torreon, 5 August; and

Sphaeralcea angustifolia (Cavara) David Don,

Rancho Grande, 4 August.

Females of what may be this species, but differing in that the proximal duplex setal are not quite

on a line with the proximal duplex setae and that

tarsus I is not as slender, were collected on Sida

species, Mazatlan, 26 July.
NUMBER 171 17

Tetranychus (Tetranychus) fiechtmanni, new

species

FIGURE 28

This species is related to T. (T.) aimipenis

Flechtmann and Baker (1970) in having a posteriorly protruding spur on the aedeagus. In T.

armipenis this spur is pointed distally; in the

species here described it is either blunt or slightly

indented.

MALE.—Striae of both sexes not lobed, that of

T. (T.) armipenis $ lobed. Empodium I with

ventral hairs forming claw; other empodia with

ventral hairs free. Sensillum of palpus longer than

broad. Knob of aedeagus directed dorsal and with

the terminal angulation ending well beyond the

level of the bend, as in T. pacificus McGregor.

Posteriorly, below the knob, is a short projecting

blunt or indented spur as figured. Length of body

363^i; including rostrum 446[i.

FEMALE.—Typical for the subgenus Tetranychus.

Length of body 446JA; including rostrum 574u.

HOLOTYPE.—Male, USNM 3549, ex Haplopappus spinulosus (Pursh) De Candolle, Zacatecas,

5 August. „

PARATYPE.—Male with the above data.

This species has been named in honor of Dr.

Carlos Flechtmann of the University of Sao Paulo

at Piracicaba, Sao Paulo, Brazil.

Tetranychus (Tetranychus) gigas Fritchard

and Baker

Tetranychus gigas Pritchard and Baker, 1955:405.

Tetranychus (Tetranychus) gigas.—Tuttle and Baker, 1968:

127.

This species was found on Parthenium incanum

Humboldt, Bonpland, and Kunth, Torreon, 5

August.

Tetranychus (Tetranychus) hydrangeae Pritchard

and Baker

Tetranychus hydrangeae Pritchard and Baker, 1955:425.

Specimens were collected on Ambrosia ambrosioides Cavara, Ciudad Obregon, 22 July; Clethra

alnifolia Linnaeus, Hermosillo, 17 July; and

Cucumis melo Linnaeus, Antunez, 5 May 1968, by

Servando Lopez B.

Tetranychus (Tetranychus) merganser Boudreaux

Tetranychus merganser Boudreaux, 1954:181.

This species was taken from Solanum rostratum

Ounal, Tepic, 25 July.

Tetranychus (Tetranychus) polys Pritchard and

Baker

Tetranychus polys Pritchard and Baker, 1955:396.

Tetranychus (Tetranychus) polys.—Tuttle and Baker, 1968:

131.

A series was taken from Solatium elaeagnifolium

Cavara, Topolobampo, 24 July.

Tetranychus (Tetranychus) urticae Koch

Tetranychus urticae Koch, 1836:10.

Tetranychus (Tetranychus) urticae.—Tuttle and Baker, 1968:

129.

This species appears to be out of its northern

range, but the females collected appear to be

typical.

Specimens in Mexico were collected on Acacia

greggii Gray, 10 miles east of Torreon, 5 August;

and Rosa dilecta Rehder, Guadalajara, 31 July.
18

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Literature Cited

Baker, E. W., and A. E. Pritchard

1963. Aranas rojas de America Central. Revista Sociedad

Mexicana de Historia Natural, 23:309-340. [Bears

the date 1962.]

Baker, E. W., and D. M. Tuttle

1972. New Species and Further Notes on the Tetranychoidea Mostly from the Southwestern United

States (Acarina: Tetranychidae and Tenuipalpidae).

Smithsonian Contributions to Zoology, 116: 37

pages.

Banks, N.

1900. The Red Spiders of the United States (Tetranychus

and Stigmaeus). United States Department of Agriculture Division of Entomology Technical Series,

8:65-77.

Berlese, A.

1886. Acari dannosi alle piante coltivati. 31 pages. Padova.

Boisduval, A.

1867. Essai sur Ventomologie horticole. 648 pages.

Bondar, A.

1938. Xotas entomologicas da Bahia. II. Revista de Entomologia (Brazil), 9 (3-4):441-443.

Boudreaux, B.

1954. New Species of Tetranychid Mites (Acarina). PanPacific Entomology, 30(3): 181-186.

DeLeon, D.

1957. Two New Eotetranychus and a New Oligonychus

from Southern Florida (Acarina: Tetranychidae).

Florida Entomologist, 40(3): 111-113.

Dufour, L.

1832. Description et figure du Tetranychus lintearicus,

arachnide nouvelle de la tribu des acarides. Annales

des Sciences Naturelles Paris, 25: 276 pages.

Estebanes, M. L., and E. W. Baker

1968. Aranas rojas de Mexico. Anales Escuela Nacional

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Ewing, H. E.

1926. Two New Spider Mites (Tetranychidae) from

Death Valley, California (Acarina). Entomological

News, 37:142-143.

Fletchmann, C. H. W., and E. W. Baker

1970. A Preliminary Report on the Tetranychidae of

Brazil. Annals Entomological Society of America,

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Jacobi, A.

1905. Eine Spinnmilbe (Tetranychus ununguis n. sp.) als

Koniferenschadling. Naturwiss Zeitschrift Landwirtschaftltch forswesen, 3:239-247.

Koch, C. L.

1836. Deutschlands Crustaceen Myriapoden und Arachniden. 1:8.

McGregor, E. A.

1934. A New Spinning Mite on Citrus at Yuma, Arizona.

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1934. A New Spider Mite on Citrus in Southern California (Acarina: Tetranychidae). Proceedings Entomological Society Washington, 45(5): 127-129.

1950. Mites of the Family Tetranychidae. The American Midland Naturalist, 44(2):257-420.

McGregor, E. A., and A. Ortega

1953. Una nueva arana roja de Mexico Paratetranychus

mexicanus sp. nov. Folleto Technico, 10:1-17.

Murray, A.

1877. Economic Entomology, Aptera. 443 pages. London:

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-.•> U.S . GOVE 0/13
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