The Old World Stenomidae:

A Preliminary Survey of the Fauna,

Notes on Relationships,

and Revision of the Genus Eriogenes

(Lepidoptera: Gelechioidea)

W. DONALD DUCKWORTH

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY • NUMBER 147
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SMITHSONIA N CONTRIBUTION S T O ZOOLOG Y • NUMBE R 147

The Old World Stenomidae:

A Preliminary Survey of the Fauna,

Notes on Relationships,

and Revision of the Genus Eriogenes

(Lepidoptera: Gelechioidea)

W. Donald Duckworth

SMITHSONIAN INSTITUTION PRESS

City of Washington

1973
ABSTRACT

Duckworth, W. Donald. The Old World Stenomidae: A Preliminary Survey of

the Fauna, Notes on Relationships, and Revision of the Genus Eriogenes (Lepidoptera: Gelechioidea). Smithsonian Contributions to Zoology, number 147,

21 pages, 9 figures, 7 maps, 1973.—A preliminary review of the presently known

composition of the microlepidopterous family Stenomidae in the Old World is

provided, including observations on distribution of the family, relationships

with other families in the superfamily Gelechioidea, and a catalog of the Old

World genera and species. The genus Eriogenes is revised and transferred from

the Xyloryctidae to the Stenomidae, and one new species is described.

OFFICIAL PUBLICATION DATE is handstamped in a limited number of initial copies and is recorded

in the Institution's annual report, Smithsonian Year. SI PRESS NUMBER 4777. SERIES COVER

DESIGN: The coral Montastrea cavernosa (Linnaeus).

Library of Congress Cataloging in Publication Data

Duckworth, W. Donald, 1935-

The Old World Stenomidae.

(Smithsonian contributions to zoology, no. 147)

Bibliography: p.

1. Stenomidae . I. Title. II. Series: Smithsonian Institution. Smithsonian contributions to

zoology, no. 147.

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For sale by the Superintendent of Documents,

U.S. Government Printing Office, Washington, D.C. 20402

Price 40 cents domestic postpaid or 30 cents GPO Bookstore
The Old World Stenomidae:

A Preliminary Survey of the Fauna,

Notes on Relationships,

and Revision of the Genus Eriogenes

(Lepidoptera: Gelechioidea)

W. Donald Duckworth

Introduction

For many years the microlepidopterous family

Stenomidae has been viewed as a uniquely New

World family with little, if any, representation in

the Old World fauna. The development of this

concept is difficult to trace precisely; however, to

a large extent it stems from the confusion among

earlier workers over the relationship between the

Stenomidae and the closely related family Xyloryctidae coupled with a lack of detailed studies dealing with either family.

Although both families were originally proposed

by Edward Meyrick, his concepts concerning them

fluctuated through the years primarily due to his

steadfast refusal to utilize characters other than

wing venation for his higher classification. This

dependance on a single character source led to

numerous misconceptions, particularly with the

higher categories. As the venational characters he

originally used to separate the xyloryctids and

stenomids began to break down with the continuing discovery and description of new species, Meyrick (1915) merged the two families under the

older name Xyloryctidae. From this point he proW. Donald Duckworth, Department of Entomology, National

Museum of Natural History, Smithsonian Institution, Washington, D.C. 20560.

ceeded during the remainder of his lifetime to describe hundreds of species from both the Old and

New World tropics under the family Xyloryctidae.

Other workers, principally August Busck, continued to utilize the original two-family concept

placing the New World species and genera in

Stenomidae. This geographical distinction was

further emphasized by Busck (1934) when in his

Stenomidae portion of the Lepidopterorum Catalogus series he only included genera and species

from the New World. Thus, through omission,

the genera and species from areas other than North

and South America remained largely unknown and,

until the present, uncataloged. In addition, the

Xyloryctidae (sensu stricto) have never been cataloged and remain essentially unstudied.

In recent years a number of papers have reflected

the distinctiveness of the two families; however,

due to the nature of the studies (faunistic, type

catalog) they do not provide a sufficiently clear

picture of the two families to serve as a basis for

future studies. Clarke (1955a) provides a thorough

account of Meyrick's treatment of xyloryctids and

stenomids in his introductory material to the catalog of Meyrick types; however, his detailed treatment (1955b) of the type specimens does not

include species from Australia where the majority

of xyloryctids and Old World stenomids occur.
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Diakonoff (1954) provided a complete and accurate definition of the two families based on his

study of the Microlepidoptera of New Guinea. His

descriptions coupled with those of Common (1970)

based on the Australian fauna provide the most

comprehensive characterization of the two families

yet published. Both studies, however, are restricted

in scope and the latter provides only fragmentary

information below family level.

During the course of revisionary studies of the

Western Hemisphere Stenomidae conducted by the

present author in recent years, the lack of information concerning the Old World elements of the

family has presented increasing difficulties, particularly in relation to higher category relationships and evolutionary trends. The present paper

is intended to provide a partial remedy to this

situation and, more importantly, to serve as a starting point for more comprehensive studies. An attempt has been made to provide as complete a

catalog as possible of the previously described taxa

that have been assigned to the family or that have

been determined to be more appropriately placed

in the family. No finality is suggested for this catalog, rather it is anticipated that numerous modifications will become necessary as additional

information is obtained.

A detailed revision of the genus Eriogenes Meyrick is included and represents the first step toward

a more thorough understanding of the Old World

stenomid fauna and its relation to the remainder

of the family. Previously assigned to the Xyloryctidae, the genus is herein transferred to the Stenomidae and a new species is described.

The comments which follow concerning the relationships between the Xyloryctidae and Stenomidae, as well as the statements concerning the

distribution of the Old World fauna, are preliminary. A great deal more work remains to be accomplished, especially at the generic and specific

levels, before more precise evaluations can be made.

In the meantime, it seems appropriate to bring

together as much of the existing information as

possible in order to facilitate future studies.

The author wishes to acknowledge with gratitude

the aid and cooperation of the following individuals and institutions who, through their support

and encouragement, have materially aided the present study: Mr. Allan Watson, Mr. P. E. S. Whalley,

Dr. Klaus Sattler, British Museum (Natural History); Dr. J. F. Franclemont, Cornell University;

Dr. I. F. B. Common, Commonwealth Scientific and

Industrial Research Organization, Canberra, Australia, for lending types and other specimens in

their charge for study. Special thanks are extended

to Dr. Ian F. B. Common, C. S. I. R. O., Canberra,

Australia and Mr. John S. Dugdale, Department

of Scientific & Industrial Research, Nelson, New

Zealand, for their aid in documenting the stenomid

fauna of their respective areas.

The author also wishes to acknowledge the assistance of Mr. George Venable, departmental illustrator, for the artwork; Mrs. Vera Milbank, technician, for bibliographic assistance; and the

National Museum of Natural History Photographic

Laboratory for the photographs.

Classification and Relationships

The greatest difficulty encountered in attempting

to define the family Stenomidae is the lack of

knowledge concerning the two most closely related

families, Xyloryctidae and Oecophoridae. Fortunately, at least for purposes of the present study,

the stenomids are relatively distinct morphologically, and while the oecophorids and xyloryctids

are yet to be clearly separated from each other,

it is possible to distinguish the stenomids from

them to a satisfactory degree and to indicate certain relationships.

The Stenomidae are distinguished by the following combination of characters: labial palpus upcurved, generally robust, apical segment usually

shorter than second segment; antenna long-ciliated

in the male, scape without pecten; ocelli absent;

maxillary palpus 4-segmented, usually partially

encircling base of proboscis; forewing with veins 7

and 8 (R4 and R5) usually separate, vein 2 (CuA2)

usually arising at or near lower angle of discal cell;

hind wing with vein 8 (Sc -j- R-i) approaching 7

(Rs), or 7 (R,) curved upward near upper angle

of discal cell, veins 6 (M-,) and 7 (Rs) usually

stalked; abdomen without spines on terga; male

genitalia with harpe simple to complex, bearing

specialized setae which are frequently bifurcate

apically.

As pointed out by Hodges (1966) many of the

genera currently assigned to the Xyloryctidae ap-
NUMBER 147

pear to be improperly placed, and until the family

is carefully studied it is impossible to provide an

accurate analysis of family limits; however, in my

opinion, the following characters provide the most

satisfactory definition possible at this time: labial

palpus upcurved; antenna simple, ciliated or pectinated in male, scape without pecten; maxillary

palpus 3- or 4-segmented, usually partially encircling base of proboscis; forewing with veins 7

and 8 (R4 and R5) usually stalked, vein 2 (CuA2)

arising well before lower angle of discal cell; hind

wing with vein 8 (Sc -|- Kt) separate and diverging

from 7 (R,) well before upper angle of discal cell,

veins 6 (Mj) and 7 (Rg) approximated, connate

or stalked; abdomen with spines on posterior half

of terga or along posterior margins; male genitalia

with harpe usually simple, without specialized

setae.

Diakonoff (1954) presented virtually the only

discussion of the relationships between the Stenomidae and Xyloryctidae to be found in the modern

literature. His conclusions were based primarily

on the New Guinea fauna and thus do not necessarily define either family in relation to their total

range and composition. In fact, there is evidence

which indicates that he did not always adhere to

his family definitions for placement of the New

Guinea genera. For example, the genus Eriogenes,

revised and transferred to the Stenomidae in the

present study, was included in the Xyloryctidae by

Diakonoff (1954:127) with the following comment:

"Meyrick regarded this genus to be related with

Agriophara (Stenomidae) which is certainly erroneous. The insect is a true Xyloryctid, is in our

opinion, allied to Cryptophasa, and does not show

any connections with the Stenomidae." In fact,

thorough study of the genus reveals that Eriogenes

agrees in every respect with the characters Diakonoff lists as distinctive for the Stenomidae and,

as discussed elsewhere in the present paper, demonstrates a closer relationship with the Neotropical

stenomids than to any portion of the Old World

fauna recognized to date.

As discussed in the introduction, the confusion

concerning the families Xyloryctidae and Stenomidae has a long history and involves personalities

as well as evaluation of characters. Personalities no

longer intrude into the problem; however, the relaMAP 1.—Generalized distribution of Old World Stenomidae.
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

MAP 2.—Distribution of Agriophara species (type-localities only): I, A. asaphes Diak.; 2,

A. atratella (Wlk.); 3, A. axesta Meyr.; 4, A. biornata Diak.; 5, A. bradleyi Diak.; 6, A. capnodes

Meyr.; 7, A. Cinderella (Newman); 8, A. cinerosa Rosenstock; 9, A. confertella (Wlk.); 10,

A. cremnopis Lower; 11, A. curta Lucas; 12, A. diminuta Rosenstock; 13, A. discoloba Turner;

14, A. dyscapna Turner; 15, A. fascifera Meyr.; 16, A. gravis Meyr.; 17, A. heterchrotna Diak;

18, A. horridula Meyr.; 19, A. hylinota Lower; 20, A. leptosemela Lower; 21, A. leucanthes

Turner; 22, A. leucosta Turner; 23, A. levis Meyr.; 24, A. lysimacha Meyr.; 25, A. musicolor

Meyr.; 26, A. neoxantha Meyr.; 27, A. nephelopa Diak.; 28, A. nodigera Turner; 29, A. parallela

Diak.; 30, A. parilis Meyr.; 31, A. phasmatopa (Meyr.); 32, A. plagiosema Turner; 33, A. platyscia

Lower; 34, A. poliopepla Turner; 35, A. polistis (Lower); 36, A. salinaria Meyr.; 37, A. tephroptera Lower; 38, A. virescens Diak.; 39, A. coricopa (Meyr.).

tive uncertainty over supergeneric classification, not

only in these two families but throughout the

Gelechioidea, continues to be the major obstacle

to a better understanding of their interrelationships and distribution in the world fauna. Hodges

(1966) and Powell (1973) have discussed this

problem in relation to their studies of the gelechiids and ethmiids, respectively, and both

emphasize the need for increased elucidation of

suprageneric taxa within the Gelechioidea on a

world basis.

Geographical Distribution

A generalized diagram of the distribution of Old

World stenomids is provided in Map 1. The patterns shown are based only on type-localities to

avoid errors of misidentification in those groups

not studied in detail. Maps 2-6 indicate the precise species localities for the various genera again

based on type-localities only. Map 7 provides the

total known distribution of the genus Eriogenes

and is discussed in detail in the treatment of that

genus.
NUMBER 147

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MAP 3.—Distribution of Phylomictis, Aproopta, Proscedes, and Nothochalara species (typelocalities only): 1, Ph. arctans Lucas; 2, Ph. decretoria Lucas; 3, Ph. eclecta Turner; 4, Ph.

idiotricha Meyr.; 5, Ph. leucopelta Lower; 6, Ph. lintearia Meyr.; 7, Ph. maligna Meyr.; 8, Ph.

monochroma Lower; 9, Ph. obliquata Lucas; 10, Ph. palaeomorpha Turner; 11, Ph. sarcinopa

Meyr.; 12, A. metanchlaena Turner; 13, Pr. torquigera Diak.; 14, N. sordida Diak.

Perhaps the most significant aspect of the distribution shown in Map 1 is the obvious absence

of records from most of the Oriental Region. This

reflects, of course, the serious lack of collecting of

Microlepidoptera accomplished to date in this important area. The impact of our ignorance of the

Oriental Region, as it relates to Microlepidoptera,

is brought into focus when it is realized that there

is increasing evidence, convincingly summarized

and elaborated by Smith (1970), that the original

center of distribution of flowering plants (angiosperms) was, in all probability, southeastern Asia

and adjacent Malesia. Viewed in conjunction with

the probability that the Order Lepidoptera owes

its diversity and success to the development of the

flowering plants, information concerning the

Microlepidoptera from the Oriental Region becomes basic to an understanding of the origin and

dispersal of the group. Thus, it seems reasonable

to assume that until additional field sampling is

achieved, any comments concerning relationships

in the Old World Microlepidoptera are exceedingly tenuous.

With the previous statements in mind, it seems

to me appropriate to examine the currently known

distribution of the Old World stenomid fauna,

albeit imperfect, and to consider possible explanations for the patterns observed.

When the various localities are examined in detail a definite tropical character to the distribution

of Old World stenomids emerges. Also, most of the

localities are in areas of tropical rain forest or

tropical montane forest as defined by Eyre (1968)

with the exception of southeastern Australia. This
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

4.—Distribution of Synchalara and Neospastis species (type-localities only).

• 5. argoplaca (Meyr.) 3 S. rhombota (Meyr.)

• S. byrsina (Meyr.) Q N. calipidias Meyr.

+ S. malacobryas Meyr. D N. encryphias (Meyr.)

• S. minax (Meyr.) it N. ichnaea (Meyr.)

pattern coincides with observations on the Neotropical stenomid fauna (Duckworth, 1969, 1971)

and, in my opinion, it is in this type of habitat

that stenomids have flourished and achieved their

initial diversification in both the Old and New

World. Secondary penetration of dryer habitats has

occurred but appears to be less important to the

overall family dispersal. The best example of this

pattern can be seen in the genera from Madagascar

(Maps 5 and 6) where only one species is described

from the western savanna area, the remainder occurring in the tropical forests which extend along

the eastern portion of the island.

A great deal more study at the generic level must

be accomplished before additional interpretation

of Old World stenomid distributions can be made.

The one genus which has been examined in detail

during the present study, Eriogenes, does indicate

a distinct relationship between the Old and New

World elements of the family and, furthermore,

appears to possess characteristics reflective of an

ancestral condition. If this is found to be the case

for other Old World genera, especially in the

Southeast Asia-Malesia area, it would seem likely

that the generalized model of angiosperm distribution proposed by Smith (1970) would hold true

for stenomids as well. In the case of stenomid entry into the New World, data at this point would

appear to favor Smith's southern route along eastern Australia and the New Caledonian-New Zealand insular chain into the West Antarctic

Archipelago and, utilizing island stepping stones,

into South America. There is no evidence, at this

point, which suggests a route crossing Beringia to

the north; the Nearctic stenomid fauna appears to

have originated in the Neotropical Region and is

only marginally successful in temperate habitats

(Duckworth, 1971).
NUMBER 147

MAP 5.—Distribution of Herbulotiana species (type-localities

only): 1, H. abceda Viette; 2, H. benoistella Viette; 3,

H. bernardiella Viette; 4, H. bicolorata Viette; 5, H. catalaella Viette; 6, H. collectella Viette; 7, H. halarcta (Meyr.);

8, H. longifascia Viette; 9, H. paulianella Viette; 10, H. rungsella Viette; 11, H. vadonella Viette; 12, H. violacea Viette;

13, H. atypicella Viette; 14, H. robustella Viette; 15, H. altitudinella Viette; 16, H. septella Viette.

MAP 6.—Distribution of Amontes, Mocquerysiella, and

Mnarolitia species (type-localities only).

• A. princeps Viette • Afn. paulianellum Viette

D Mo. albicosta Viette * Afn. sylvestrella Viette

O Mo. bourginella Viette

Revision of the Genus Eriogenes Meyrick

The genus Eriogenes was established by Meyrick

(1925) with a single included species, E. mesogypsa, with the indication that it was apparently

closely related to Agriophara. Diakonoff (1954)

included Eriogenes in the family Xyloryctidae in

his study of the New Guinea Microlepidoptera,

noting that Meyrick's placement of the genus as a

relative of Agriophara (Stenomidae) was in error.

Contrary to that opinion, in the present study it

has been determined that Eriogenes is, in fact, more

appropriately placed in the Stenomidae and shows

no relationship to the xyloryctids in any character

other than gross superficial appearance. In addition, detailed study of the morphological characters within the genus, particularly the genitalia,

suggest a relationship to certain genera in the Neotropical portion of the family.
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

MORPHOLOGY

Superficially, the members of this genus are large,

robust, densely scaled moths which resemble certain noctuids, or some of the larger xyloryctids such

as Cryptophasa, in general appearance. Although

comprised of only two species, the genus exhibits a

number of morphological characters worthy of special emphasis.

HEAD.—In both species the eye index is 1.0 or

greater, ranging from 1.0 to 1.3, suggesting that

these moths are nocturnal in adult activity patterns. For additional discussion of the eye index

and its significance refer to Duckworth (1971) and

Powell (1973). The maxillary palpi are 4-

segmented. The labial palpi are 3—segmented; the

apical segment is reduced and variable in size and

shape between the two species. The proboscis is

reduced and hidden between the basal segments of

the labial palpi when in the normal coiled position.

THORAX.—The thorax is very robust and clothed

laterally and ventrally with long hair scales. The

wing venation is homogeneous and no characters

of specific significance were observed. The occurrence of a cross vein between the upper edge of the

cell and vein 8 (Sc -f- R^) in the hind wing, mentioned by Diakonoff (1954), is variable and not a

dependable character at either the generic or specific level.

ABDOMEN.—No structures of generic significance

were observed on abdominal segments 1-8. Perhaps the most notable aspect of this area of the

body is the absence of secondary sexual structures,

with the exception of a small eversible ventral

pouch covered with a small mass of erectile hair

scales in the intersegmental membrane between

segments 7 and 8 in E. meyricki.

MALE GENITALIA.—By far, the most significant

and intriguing structural characters in this genus

are found in the male genitalia. The harpe consists of a simple lobe rather uniformly clothed over

the apical half with specialized setae. These specialized setae are broadened and multilobed apically, the number of lobes varying but apparently

always more than two. The uncus is large, broadly

articulated with the tegumen basally, and somewhat recurved near the midpoint. The gnathos is

pendulous and produced into a heavily sclerotized

plate apically. The anellus consists of a simple

plate with two lateral lobes. Both the shape of

the gnathal plate and the size and shape of the

lateral lobes of the anellus vary between the two

species and serve to distinguish them specifically.

The aedeagus is produced apically, and the vesica

is armed with cornuti in E. meyricki and unarmed

in E. eriogenes.

FEMALE GENITALIA.—No female specimens are

known for the species in this genus.

GEOGRAPHICAL DISTRIBUTION

This genus is presently known to occur only in

New Guinea and adjacent islands, including Ceram

in the Moluccas (Map 7). The lack of records for

the two species of Eriogenes seriously restricts any

effort to analyze distribution patterns or to project

ranges in any definitive fashion. The absence of

biological data, especially host plant information,

further complicates matters and renders all observations made at this point tentative.

The most interesting aspect of the distribution

patterns for the two species of Eriogenes is the divergence from mainland New Guinea into different

adjacent island groups. E. meyricki is recorded

from Ceram in the southern Moluccas, the Central

Range on mainland New Guinea, and throughout

the Bismark Archipelago. On the other hand, E.

mesogypsa extends from the Idenburg River in

West Irian, southeastward into the D'Entrecasteau

and Louisiade island groups. Similar patterns have

been noted for butterflies (Zeuner, 1943; Carpenter, 1953) and they appear to correlate with the

geological history of New Guinea (Toxopeus,

1950; Holloway and Jardine, 1968).

In the Miocene the northern ranges of New

Guinea were a part of a "Melanesian arc" of islands

which included the North Moluccas and Talaud

Islands to the west and the Bismarck and Solomon

Islands to the east. Farther south, land which now

constitutes the central range of New Guinea appeared in late Miocene, well isolated from the

Melanesian arc. During the Pliocene-Pleistocene

further uplifting occurred in the central range and

gradually the central and northern ranges were

joined. Other portions of New Guinea (e.g., Vogelkop) undoubtably existed as separate islands during the Pleistocene. Thus, the butterfly fauna, as
NUMBER 147

MAP 7.—Distribution of Eriogenes species.

E. mesogypsa Meyr • E. meyricki Duckworth

analyzed by Holloway and Jardine (1968) reflects

three faunal elements; one-third endemic, one-third

with Sunda-based affinities, and one-third Melanesian. They further conclude that this heterogeneity results from the Melanesian and endemic

elements having risen separately through dispersal

from Asia into the Melanesian arc and into the

formerly separate southern part of New Guinea

since the Miocene. Although insufficient data are

available to confirm or reject a similar pattern in

Eriogenes, it is interesting to note that E. meyricki

displays a "Melanesian" pattern and E. mesogypsa

appears to be more "endemic" in distribution.

The currently known distribution of the genus

also suggests the possibility that species of Eriogenes may occur farther west into Indonesia. This

area is virtually unknown insofar as Microlepidoptera are concerned (Diakonoff, 1954) and undoubtably served as a major point of dispersal to the

New Guinea region.

Available data suggest the possibility of a significant altitudinal difference for the two species

of Eriogenes. E. mesogypsa is much the better

documented species in this regard and appears to

occur at lower elevations up to approximately 2500

feet. One record, from Goodenough Island, indicates a range in elevation from 2500 feet to 4000

feet; however, the consistency of other records

would suggest that this specimen was probably captured at the lower limits of this range. Also, there

is a specimen from the Jimmi River, in the Western

Highlands, with an altitude record of 4700 feet.

This appears to be an error in that the Jimmi

River is situated in a low valley at an elevation

generally less than 2500 feet. At one point, according to Gressitt (1956), the river elevation is

460 meters (1508 ft). E. meyricki, on the other

hand, is recorded from Ceram at an elevation of

4600 feet, Telefomin in central New Guinea at

4500 to 5500 feet, and the other specimens from

Rook, St. Matthias and New Britain Islands are

without elevation data. Rook Island has elevations above 2500 feet and thus the potential for

correlation; however, St. Matthias is a much lower
10 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

island and would not appear to fit the concept.

Yet Hartert (1924), in a paper on the birds of St.

Matthias, indicates the island consists of a relatively

high plateau (2130 ft) partly covered with thick

forest which might be suitable habitat for species

that occur at greater altitudes on the higher islands.

CLASSIFICATION AND RELATIONSHIPS

Although a great deal of the Old World stenomid fauna remains to be studied in detail, and

undoubtably is yet to be discovered, the genus

Eriogenes is of considerable significance in that it

appears to be more closely related morphologically

to the Neotropical genus Chlamydastis and its allies

than to any Old World genus. As pointed out in

a previous paper (Duckworth, 1971), Chlamydastis

and several related genera seem to constitute a

major division within the family, characterized, in

part, by their possession of multilobed, palmatetype specialized setae on the harpe in the male

genitalia. Eriogenes has a less specialized type of

multilobed setae on the harpe, which suggests the

possibility that it represents a more primitive stage

in the development of this character. In addition,

virtually every structure of the male genitalia demonstrates both a relationship with Chlamydastis

and a more generalized character state than is

found in that genus. For example, in Chlamydastis

the specialized setae are restricted to the outer costal margin of the harpe, there is a tendency for

reduction in the length of the uncus, the harpe

shows a tendency to divide into a costal lobe and

saccular lobe, and the gnathos may be long and

pendulous or entirely absent. In Eriogenes the

specialized setae are uniformly distributed over the

apical half of the harpe, the uncus is very long,

the harpe is a large simple lobe, and the gnathos

is long and pendulous. Thus, in each of these

structures, a clear relationship exists between the

genera in terms of similarities of structure, with

Eriogenes possessing less reduction and modification than Chlamydastis. After consideration of

these and other characters (e.g., maculation and

venation), Eriogenes, in my opinion, represents a

distinct link between the Old and New World

stenomid faunas and possibly displays characteristics reflective of an ance'stral condition.

Genus Eriogenes Meyrick

Eriogenes Meyrick, 1925:159. [Type-species: Eriogenes

mesogypsa Meyrick by monotypy.]

Head densely scaled, lateral tufts raised. Male

antenna heavily ciliated ventrally, slightly fasciculate. Labial palpus recurved, not reaching vertex,

thickened with heavy scaling. Proboscis reduced,

concealed between labial palpi; maxillary palpus

4—segmented. Thorax and abdomen densely covered with long hair scales. Legs with all tibia and

tarsi expanded with hair scales. Forewing with

costa slightly arched, apex rounded, termen

oblique, tornus rounded; with 12 veins, all separate, discal cell extending beyond midpoint, 2

(CuA2) from end of cell, 7 (R5) to termen. Hind

wing broader than forewing; with 8 veins, 3 and 4

(CuAj and M3) connate, 6 and 7 (Mx and Rs)

connate, upper edge of cell curved upward toward

vein 8 (Sc -f- Rx) at apical four-fifths.

MALE GENITALIA.—Uncus simple, curved slightly

ventrad from middle; gnathos pendulous, fused

apically forming gnathal plate; harpe simple, apical

half clothed with apically multilobed specialized

setae; anellus with lateral lobes; vinculum fused;

Key to the Species of Eriogenes

BASED ON EXTERNAL FEATURES

Apical segment of labial palpus one-third to one-half the length of second segment; forewing

with discal spot on outer margin of median white fascia E. mesogypsa Meyrick

Apical segment of labial palpus less than one-third the length of second segment; discal spot

within median white fascia E. meyricki, new species

BASED ON THE MALE GENITALIA

Anellus with lateral lobes long, falciform; vesica of aedeagus without cornuti

E. mesogypsa Meyrick

Anellus with lateral lobes short, truncate; vesica of aedeagus with cornuti

E. meyricki, new species
NUMBER 147 11

FIGURES 1-4.—Head, legs, genitalia morphology: 1, caudolateral view of head of E. mesogypsa;

2, legs of E. mesogypsa; 3, male genitalia of E. mesogypsa (left harpe removed) ; 4, male

genitalia of E. meyricki (left harpe removed) .
12 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

aedeagus produced apically, vesica of aedeagus with

or without cornuti.

FEMALE GENITALIA.—Unknown.

DISCUSSION.—This genus is readily separated

from other Old World stenomid genera by the reduced proboscis, distinctive maculation of the forewings, robust thorax and abdomen covered with

long hair scales, and distinctive male genitalia. In

the latter, the long uncus, pendulous gnathos, and

simple harpe with multilobed specialized setae are

characteristic.

No female specimens of species in this genus are

known and, consequently, it is impossible to provide distinguishing characteristics based on structures of the female genitalia.

Eriogenes mesogypsa Meyrick

FIGURES 1-3, 5-8; MAP 7

Eriogenes mesogypsa Meyrick, 1925:159.

Alar expanse 30-35 mm.

Antenna with scape light brown suffused with

white, shaft light brown dorsally. Head dark brown

medially, whitish tufts posterior to antennae, face

brown suffused with white; labial palpus dark

brown, apical segment and apex of second segment

sprinkled with white, apical segment one-third to

one-half the length of second segment. Thorax and

tegulae brown suffused with white dorsally. Legs

white shaded with light brown; foreleg with coxa

overcast with pale brown; femur, tibia, tarsi overcast with dark brown; midleg with tarsi brown;

hind leg with tarsi white. Abdomen light brown to

white. Forewing brown, darker basally, a median,

triangulate, white fascia from costa to dorsum,

discal spot dark brown on outer margin of white

fascia, apical third brown along costa suffused

with white toward tornus, a faint, outwardly curved

brown line from costal three-fourths to tornus,

cilia with interrupted brown line basally, white

beyond. Hind wing light brown to white, cilia

light brown basally, white beyond.

MALE GENITALIA (WDD 4199, lectotype).—Uncus rounded apically; gnathal plate sharply narrowing before apex, apex narrowly recurved; harpe

broadly rounded apically; anellar plate with ventral margin straight, two upright, falciform lateral

lobes; aedeagus long, narrow, more than one-half

the length of harpe, apex produced into a broad,

knifelike flange; vesica without cornuti.

FEMALE GENITALIA.—Unknown.

TYPE.—Lectotype male, British Museum (Natural History).

TYPE-LOCALITY.—Kumusi River, Northeast British New Guinea at low elevation.

HOST PLANT.—Unknown.

DISTRIBUTION. — Presently known from New

Guinea and adjacent islands at low elevations.

ADULT RECORDS. — NEW GUINEA: Kumusi

River, low elevation; Bernhard Camp, Idenburg

River, 50 m (Jan.); Hydrographer Mountains,

2500 ft (Jan., Feb.); Western Highlands, Jimi

River, 4700 ft (July, Sept.). D'ENTRECASTEAU

ISLANDS: Goodenough Island, 2500-4000 ft

(April, May, June). MURUA (Woodlark Island);

Kulamadau (Jan., May). LOUISIADE ARCHIPELAGO: Sudest Island (Tagula), Mount Riu,

2000 ft (Jan., Feb., March, April); Rossel Island,

Mount Rossel, 2100 ft (Nov., Dec); Rossel Island,

Abaleti (Nov., Dec); Misima Island, Umana Camp,

500 ft (June, July).

DISCUSSION.—Meyrick originally described this

species from four specimens and noted that one

of the four, from Ceram, was somewhat atypical.

During the course of this study it was determined

that the specimen from Ceram represented a separate species which, although very similar superficially, may be readily distinguished by both genitalia characters and maculation. In addition to

the original three specimens, in excess of twenty

additional specimens of E. mesogypsa from various

localities have been studied.

For the most part, the material studied was collected many years ago and, consequently, it is

impossible to determine the effect of time on the

scale colors. Meyrick (1925) indicates a copper

sheen in his original description which is not discernible in the specimens now, and it should be

noted that the original series was collected at least

ten years before Meyrick's description. In addition,

most of the specimens, including the original series,

are mouldy and somewhat greased. Thus, the colors

are undoubtedly much duller than would be found

in fresh specimens. I suspect the brown wing colors

are more red-brown than they appear in the specimens now; however, additional information on

this subject must await procurement of fresh

material.
NUMBER 147 13

FIGURES 5-9.—Head morphology, wing venation, left wings: 5, head of E. mesogypsa, frontal

view, scales removed (labial palpus of E. Meyricki below); 6, maxilla of E. mesogypsa; 7, wing

venation of E. mesogypsa; 8, left wings of E. mesogypsa; 9, left wings of E. meyricki.
14 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Eriogenes meyricki, new species

FIGURES 4, 9; MAP 7

Alar expanse 27-35mm.

Antenna with scape and dorsum of shaft light

brown mixed with white. Head light brown medially, whitish tufts posterior to antennae, face light

brown; labial palpus dark brown basally to middle

of second segment, lighter brown mixed with white

to apex, apical segment less than one-third length

of second segment. Thorax and tegulae brown,

heavily suffused with white. Legs as in E. mesogypsa. Abdomen pale brown to white ventrally,

brown dorsally. Forewing brown, a median, irregularly shaped, white fascia from costa to dorsum,

discal spot dark brown within the outer margin of

white fascia, apical third with two relatively

distinct transverse brown lines, one bordering the

outer margin of white fascia, the other extending

from apical three-fourths of costa to dorsum before

tornus, cilia white with interrupted brown line

basally. Hind wing brown, cilia brown.

MALE GENITALIA (WDD 4200, holotype).—Uncus

truncate apically; gnathal plate gradually narrowing to broadly recurved apex; harpe acute apically;

anellar plate with ventral margin indented at midpoint, two short, truncate lateral lobes; aedeagus

short, broad, less than one-half the length of harpe,

apex produced into a narrow, apically acute projection; vesica with two large, heavily sclerotized

cornuti.

FEMALE GENITALIA.—Unknown.

TYPE.—Holotype male, British Museum (Natural History).

TYPE-LOCALITY.—Central Ceram, 4600 ft.

HOST PLANT.—Unknown.

DISTRIBUTION.—Presently known from the Moluccas, New Guinea, and the Bismarck Archipelago.

ADULT RECORDS.—MOLUCCAS: Ceram, 4600 ft

(Jan.). NEW GUINEA: Telefomin (Eliptamin),

4500-5500 ft (June, Sept.). BISMARCK ARCHIPELAGO: Rook Island (July); New Britain, Keravat (no date). St. MATTHIAS GROUP: St.

Matthias Island (July).

DISCUSSION.—Described from the male holotype:

"Central Ceram, 4600 ft., Jan. '20, C. F. & J. Pratt";

two male paratypes: "St. Matthias Id., July, 1923,

A. F. Eichhorn"; three male paratypes: "Rook Id.,

July, 1913, A. S. Meek, Joicey Bequest, Brit. Mus.,

1934-120"; and one male paratype: "Bismarck

Archipel., Rook Island, Meek, 6-7-13, M 768, Paravicini Coll., B. M. 1937-383." Two of the above

paratypes deposited in the National Museum of

Natural History, Smithsonian Institution, the

remainder in the British Museum (Natural History). Two male paratypes: "New Guinea, Telefomin (Eliptamin), 4500-5500 ft., 19 June-14 Sept.

1959, W. W. Brandt, Genitalia Slide G617"; one

male paratype: "NEW BRITAIN, Keravat, collected by Wm. Brandt, Sir Edward Hallstrom,

Genitalia Slide G1616"; all deposited in the Australian National Insect Collection, C. S. I. R. O.,

Canberra, Australia..

This species seems very closely related to E.

mesogypsa and may be separated by the characters

indicated under the discussion pertaining to that

species. In general, it seems E. meyricki is on the

average smaller than E. mesogypsa, occurs at higher

elevations, and is darker in maculation.

Catalog of the Old World Stenomidae

Included in the following catalog are all those

genera and species which have been assigned to the

Stenomidae by previous workers or determined

more appropriately placed with the family during

the present study. The information provided concerning location of types has been gathered from

various sources and where questionable the data

is preceded by a question mark. The genera are

arranged alphabetically and the included species

are also listed alphabetically under the generic

headings. Type-locality data are essentially as found

in the original description, with an occasional

addition (state, district, province) for clarity.

As mentioned in the introduction, no finality is

implied for this catalog. Many of the species included have not been studied and the genera are,

in many cases, poorly defined. Thus, the catalog is

intended as a point of departure for future studies

and a summation of the family's current composition in the Old World.

Genus Agriophara Rosenstock

MAP 2

Agriophara Rosenstock, 1885:39. [Type-species: Agriophara

cinerosa Rosenstock by subsequent designation, Fletcher,

1929:9.]
NUMBER 147 15

Hypeuryntis Meyrick, 1897:389. [Type-species: Hypeuryntis

coricopa Meyrick by monotypy.]

1. A. asaphes Diakonoff [Agriophara], 1948:193.

Type-locality: Paniai Lake, New Guinea.

Type: holotype female, Rijksmuseum van Natuurlijke Historie, Leiden.

2. A. atratella (Walker) [Acrobasis], 1866:1712.

Type-locality: Moreton Bay, Queensland,

Australia.

Type: holotype male, British Museum (Natural History).

3. A. axesta Meyrick [Agriophara], 1890:79.

Type-locality: Wirrabara, South Australia,

Australia.

Type: syntypes of both sexes, British Museum

(Natural History).

4. A. biornata Diakonoff [Agriophara], 1954:147.

Type-locality: Iebele Camp, 2250 m, West

Irian, New Guinea.

Type: holotype male, Rijksmuseum van Natuurlijke Historie, Leiden.

5. A. bradleyi Diakonoff [Agriophara], 1954:146.

Type-locality: Mist Camp, 1800 m, West Irian,

New Guinea.

Type: holotype male, Rijksmuseum van Natuurlijke Historie, Leiden.

6. A. capnodes Meyrick [Agriophara], 1890:77.

Type-locality: Mount Lofty, South Australia,

Australia.

Type: syntypes of both sexes, British Museum

(Natural History).

7. A. Cinderella (Newman) [Chimabacche], 1855:

288.

Type-locality: Mt. Alexander Range, Victoria,

Australia.

Type: holotype PBritish Museum (Natural

History).

8. A. cinerosa Rosenstock [Agriophara], 1885:439.

Type-locality: South Australia.

Type: holotype male, British Museum (Natural History).

9. A confertella (Walker) [Cryptolechia], 1864:

758.

Type-locality: Moreton Bay, Queensland; Sydney, New South Wales, Australia.

Type: syntypes of both sexes, British Museum

(Natural History).

10. A. coricopa (Meyrick) [Hypeuryntis], 1897:389.

Type-locality: Wellington, New Zealand.

Type: holotype male, British Museum (Natural History).

11. A. cremnopis Lower [Agriophara], 1894:93.

Type-locality: Duaringa, Queensland, Australia.

Type: holotype male, PSouth Australia Museum, Adelaide.

12. A. curta Lucas [Agriophara], 1900:161.

Type-locality: Brisbane, Queensland, Australia.

Type: holotype male, South Australia Museum, Adelaide.

13. A. diminuta Rosenstock [Agriophara], 1885:

440.

Type-locality: South Australia.

Type: holotype male, British Museum (Natural History).

14. A. discobola Turner [Agriophara], 1897:32.

Type-locality: Gisborne, Victoria, Australia.

Type: syntypes of both sexes, ?Australian National Insect Collection, Canberra.

15. A. dyscapna Turner [Agriophara], 1917:99.

Type-locality: Gisborne, Victoria, Australia.

Type: holotype male, PAustralian National

Insect Collection, Canberra.

16. A. fascifera Meyrick [Agriophara], 1890:80.

Type-locality: Sydney and Bathurst, 2500 ft,

New South Wales, Australia.

Type: syntypes of both sexes, British Museum

(Natural History).

17. A. gravis Meyrick [Agriophara], 1890:77.

Type-locality: Sydney, New South Wales, Australia; Deloraine, Tasmania.

Type: syntypes of both sexes, British Museum

(Natural History).

18. A. heterchroma Diakonoff [Agriophara], 1954:

144.

Type-locality: Iebele Camp, 2250 m, West

Irian, New Guinea

Type: holotype female, Rijksmuseum van

Natuurlijke Historie, Leiden.

19. A. horridula Meyrick [Agriophara], 1890:77.

Type-locality: Sydney, New South Wales,

Australia.

Type: holotype male, British Museum (Natural History).

20. A. hyalinota Lower [Agriophara], 1899:104.

Type-locality: Parkside, South Australia; Duaringa, Queensland, Australia.
16 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Type: syntypes of both sexes, South Australia

Museum, Adelaide.

21. A. leptosemela Lower [Agriophara], 1893:174.

Type-locality: Parkside, South Australia,

Australia.

Type: holotype male (without abdomen),

South Australia Museum, Adelaide.

22. A. leucanthes Turner [Agriophara], 1897:31.

Type-locality: Brisbane, Queensland, Australia.

Type: syntype males, Australian National Insect Collection, Canberra.

23. A. leucosta Lower [Agriophara], 1893:173.

Type-locality: Parkside, South Australia,

Australia.

Type: syntypes of both sexes, PSouth Australia

Museum, Adelaide.

24. A. levis Meyrick [Agriophara], 1921:446.

Type-locality: Brisbane, Queensland, Australia.

Type: holotype, British Museum (Natural

History).

25. A. lysimacha Meyrick [Agriophara], 1915:410.

Type-locality: Woodlark Island, New Guinea.

Type: holotype male, British Museum (Natural History).

26. A. murinella (Walker) [Cryptolechia], 1860:

757.

Type-locality: Australia.

Type: holotype female (without abdomen),

British Museum (Natural History).

27. A. musicolor Meyrick [Agriophara], 1930:560.

Type-locality: Biagi, Mambare River, 5000 ft,

New Guinea.

Type: holotype male, PDeutsches Entomologisches Institut, Berlin.

28. A. neoxanta Meyrick [Agriophara], 1930:410.

Type-locality: Woodlark Island, New Guinea;

Cooktown, Queensland, Australia.

Type: syntypes of both sexes, British Museum

(Natural History).

29. A. nephelopa Diakonoff [Agriophara], 1954:

148.

Type-locality: Moss Forest Camp, 2600-2800

m, West Irian, New Guinea.

Type: male holotype, Rijksmuseum van Natuurlijke Historie, Leiden.

30. A. nodigera Turner [Agriophara], 1900:11.

Type-locality: Warwick, Queensland, Australia.

Type: holotype female, Australian National

Insect Collection, Canberra.

31. A. parallela Diakonoff [Agriophara], 1954:142.

Type-locality: Lower Mist Camp, 1400-1600

m, West Irian, New Guinea.

Type: holotype female, Rijksmuseum van Natuurlijke Historie, Leiden.

32. A. parilis Meyrick [Agriophara], 1918:197.

Type-locality: Setekwa River, 2000-3000 ft.,

New Guinea.

Type: holotype male, British Museum (Natural History).

33. A. phasmatopa (Meyrick) [Stenoma], 1910:460.

Type-locality: Gizo, New Georgia, Solomon

Islands.

Type: syntype males, British Museum (Natural History).

34. A. plagiosema Turner [Agriophara], 1897:32.

Type-locality: Brisbane, Queensland, Australia.

Type: holotype male, Australian National Insect Collection, Canberra.

35. A. platyscia Lower [Agriophara], 1908:117.

Type-locality: Tasmania.

Type: holotype female (without abdomen),

South Australia Museum, Adelaide.

36. A. poliopepla Turner [Agriophara], 1897:31.

Type-locality: Brisbane, Queensland, Australia.

Type: syntypes of both sexes, Australian National Insect Collection, Canberra.

37. A. polistis (Lower) [Agriophara], 1923:55.

Type-locality: Dorrigo, New South Wales,

Australia.

Type: syntype males, South Australia Museum,

Adelaide.

38. A. salinaria Meyrick [Agriophara], 1931:43.

Type-locality: Florida, Solomon Islands.

Type: holotype male, PBritish Museum (Natural History).

39. A. tephroptera Lower [Agriophara], 1903:230.

Type-locality: Broken Hill, New South Wales,

Australia.

Type: holotype male, South Australia Museum, Adelaide.

40. A virescens Diakonoff [Agriophara], 1954:143.

Type-locality: Cyclops Mountains, 400-900 m,

West Irian, New Guinea.

Type: holotype male, Rijksmuseum van Natuurlijke Historie, Leiden.
NUMBER 147 17

Genus Amontes Viette

MAP 6

Amontes Viette, 1958:116. [Type-species: Amontes princeps

Viette by original designation.]

1. A. princeps Viette [Amontes], 1958:116.

Type-locality: env. de Perinet, fore"t d'Analamazoatra, 910 m, Madagascar.

Type: holotype male, Museum National D'Histoire Naturelle, Paris.

Genus Aproopta Turner

MAP 3

Aproopta Turner, 1919:171. [Type-species: Aproopta melanchlaena Turner by monotypy.]

1. A. melanchlaena Turner [Aproopta], 1919:172.

Type-locality: Katoomba, New South Wales,

Australia.

Type: holotype male, Australian Museum,

Sydney.

Genus Eriogenes Meyrick

MAP 7

Eriogenes Meyrick, 1925:159. [Type-species: Eriogenes mesogypsa Meyrick by monotypy.]

1. E. mesogypsa Meyrick [Eriogenes], 1925:159.

Type-locality: Kumusi River, Northeast British New Guinea, at low elevation.

Type: lectotype male, British Museum (Natural History).

2. E. meyricki Duckworth [Eriogenes], 1973:14.

Type-locality: Central Ceram, 4600 ft,

Moluccas.

Type: holotype male, British Museum (Natural History).

Genus Herbulotiana Viette

MAP 5

Herbulotiana Viette, 1954:17. [Type-species: Herbulotiana

abceda Viette by original designation.]

1. H. abceda Viette [Herbulotiana], 1954:23.

Type-locality: baie d'Antongil, Madagascar.

Type: holotype male, Museum National D'Histoire Naturelle, Paris.

2. • H. altitudinella Viette [Herbulotiana], 1962:

24.

Type-locality: Plateau Soaindrana, 2070 m,

Massif de l'Andringitra, Central Madagascar.

Type: holotype male, Museum National D'Histoire Naturelle, Paris.

3. H. atypicella Viette [Herbulotiana], 1956a: 115.

Type-locality: route d'Anosibe, km 57, Madagascar Est.

Type: holotype male, Museum National D'Histoire Naturelle, Paris.

4. H. benoistella Viette [Herbulotiana], 1954:18.

Type-locality: baie d'Antongil, Madagascar.

Type: holotype male, Museum National D'Histoire Naturelle, Paris.

5. H. bernardiella Viette [Herbulotiana], 1954:26.

Type-locality: env. de Maroantsetra, for£t

d'Ambodivoangy, Madagascar.

Type: holotype male, Museum National D'Histoire Naturelle, Paris.

6. H. bicolorata Viette [Herbulotiana], 1954:20.

Type-locality: baie d'Antongil, Madagascar.

Type: holotype male, Museum National D'Histoire Naturelle, Paris.

7. H. catalaella Viette [Herbulotiana], 1954:22.

Type-locality: env. de Maroantsetra, for£t

d'Ambodivoangy, Madagascar.

Type: holotype male, Museum National D'Histoire Naturelle, Paris.

8. H. collectella Viette [Herbulotiana], 1956a:111.

Type-locality: env. de Maroantsetra, for£t

d'Ambodivoangy, Nord-Est Madagascar.

Type: holotype male, Museum National D'Histoire Naturelle, Paris.

9. H. halarcta (Meyrick) [Agriophara], 1917:59.

Type-locality: Antananarivo, Madagascar.

Type: holotype male, British Museum (Natural History).

10. H. longifascia Viette [Herbulotiana], 1954:26.

Type-locality: baie d'Antongil, Madagascar.

Type: holotype female, Museum National

D'Histoire Naturelle, Paris.

11. H. paulianella Viette [Herbulotiana], 1954:24.

Type-locality: env. de Maroantsetra, foret

d'Ambodivoangy, Madagascar.

Type: holotype male, Museum National D'Histoire Naturelle, Paris.

12. H. robustella Viette [Herbulotiana], 1956a: 113.

Type-locality: env. de Perinet, for£t d'Analamazoatra, 910 m, Madagascar Est.
18 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Type: holotype male, Museum National D'Histoire Naturelle, Paris.

13. H. rungsella Viette [Herbulotiana], 1954:19.

Type-locality: env. de Maroantsetra, foret

d'Ambodivoangy, Madagascar.

Type: holotype female, Museum National

D'Histoire Naturelle, Paris.

14. H. septella Viette [Herbulotiana], 1956a: 112.

Type-locality: env. de Ranomafana, district

d'Ifanadiana, 700 m, Madagascar Est.

Type: holotype male, Museum National D'Histoire Naturelle, Paris.

15. H. vadonella Viette [Herbulotiana], 1956b:466.

Type-locality: env. de Maroantsetra, foret

d'Ambodivoangy, Nord-Est Madagascar.

Type: holotype male, Museum National D'Histoire Naturelle, Paris.

16. H. violacea Viette [Herbulotiana], 1954:21.

Type-locality: baie d'Antongil, Madagascar/

Type: holotype male, Museum National D'Histoire Naturelle, Paris.

Genus Mnarolitia Viette

MAP 6

Mnarolitia Viette, 1954:16. [Type-species: Mnarolitia paulianellum Viette by original designation.]

1. M. paulianellum Viette [Mnarolitia], 1954:16.

Type-locality: env. de Morafenobe, foret de

Mahajeby, Ouest Madagascar.

Type: holotype male, Museum National D'Histoire Naturelle, Paris.

2. M. sylvestrella Viette [Mnarolitia], 1968:84.

Type locality: env. de Perinet, foret d'Analamazaotra, 910 m, Madagascar Est.

Type: holotype male, Museum National D'Histoire Naturelle, Paris.

Genus Mocquerysiella Viette

MAP 6

Mocquerysiella Viette, 1954:13. [Type-species: Mocquerysiella

alibcosta Viette by original designation.]

1. M. albicosta Viette [Mocquerysiella], 1954:14.

Type-locality: baie d'Antongil, Nord-Est Madagascar.

Type: holotype male, Museum National D'Histoire Naturelle, Paris.

M. bourginella Viette [Mocquerysiella] 1954:15.

Type-locality: env. de Maroantsetra, foret

d'Ambodivoangy, Nord-Est Madagascar.

Type: holotype male, Museum National D'Histoire Naturelle, Paris.

Genus Neospastis Meyrick

MAP 4

Neospastis Meyrick, 1917:59. [Type-species: Agriophara encryphias Meyrick by original designation.]

1. N. calpidias Meyrick [Neospastis], 1917:59.

Type-locality: Nilgiris, Pykara, 7000 ft, India.

Type: lectotype male, British Museum (Natural History).

2. N. encryphias (Meyrick) [Agriophara], 1907a:

743.

Type-locality: Khasi Hills, Assam, India.

Type: lectotype male, British Museum (Natural History).

3. N. ichnaea (Meyrick) [Stenoma], 1914:118.

Type-locality: Anshi, Kanara, India.

Type: lectotype male, British Museum (Natural History).

Genus Nothochalara Diakonoff

MAP 3

Nothochalara Diakonoff, 1954:135. [Type-species: Nothochalara sordida Diakonoff by original designation.]

1. N. sordida Diakonoff [Nothochalara], 1954:136.

Type-locality: Araucaria Camp, 800 m, West

Irian, New Guinea.

Type: holotype female, Rijksmuseum van Natuurlijke Historic Leiden.

Genus Phylomictis Meyrick

MAP 3

Phylomictis Meyrick, 1890:74. [Type-species: Phylomictis

maligna Meyrick by monotypy.]

Comoscotopa Lower, 1902:239. [Type-species: Comoscotopa

leucopelta Lower by monotypy.]

1. P. arctans Lucas [Phylomictis], 1900:159.
NUMBER 147 19

Type-locality: Brisbane, Queensland, Australia.

Type: syntypes of both sexes, PSouth Australia

Museum, Adelaide.

2. P. decretoria Lucas [Phylomictis], 1900:160.

Type-locality: Brisbane, Queensland, Australia.

Type: holotype female, PSouth Australia Museum, Adelaide.

3. P. eclecta Turner [Phylomictis], 1906:142.

Type-locality: Burpengary, near Brisbane,

Queensland, Australia.

Type: holotype male, Australian National Insect Collection, Canberra.

4. P. idiotricha Meyrick [Phylomictis], 1921:446.

Type-locality: Brisbane, Queensland, Australia.

Type: holotype male, British Museum (Natural History).

5. P. leucopelta (Lower) [Comoscotopa], 1902:

240.

Type-locality: Mount Gambier, South Australia, Australia.

Type: holotype male, PSouth Australia Museum, Adelaide.

6. P. lintearia Meyrick [Phylomictis], 1921:445.

Type-locality: Brisbane, Queensland, Australia.

Type: holotype male, British Museum (Natural History).

7. P. maligna Meyrick [Phylomictis], 1890:75.

Type-locality: Melbourne, Victoria, Australia.

Type: holotype male, PBritish Museum (Natural History).

8. P. monochroma Lower [Phylomictis], 1892:17.

Type-locality: Parkside, South Australia, Australia.

Type: holotype female (without abdomen),

South Australia Museum, Adelaide.

9. P. obliquata Lucas [Phylomictis], 1900:160.

Type-locality: May Orchard, Brisbane, Queensland, Australia.

Type: syntypes of both sexes, PSouth Australia

Museum, Adelaide.

10. P. palaeomorpha Turner [Phlomictis], 1897:30.

Type-locality: Brisbane, Queensland, Australia.

Type: syntype male, Australian National Insect Collection, Canberra.

11. P. sarcinopa Meyrick [Phylomictis], 1920:322.

Type-locality: Brisbane, Queensland, Australia.

Type: holotype female, British Museum (Natural History).

Genus Proscedes Diakonoff

MAP 3

Proscedes Diakonoff, 1954:137. [Type-species: Proscedes torquigera Diakonoff by original designation.]

1. P. torquigera Diakonoff [Proscedes], 1954:138.

Type-locality: Rattan Camp, 1200 m, West

Irian, New Guinea.

Type: holotype male, Rijksmuseum van Natuurlijke Historic Leiden.

Genus Synchalara Meyrick

MAP 4

Synchalara Meyrick, 1917:60. [Type-species: Agriophara rhombota Meyrick by original designation.]

1. S. argoplaca (Meyrick) [Agriophara], 1907a:743.

Type-locality: Maskeliya, Ceylon.

Type: lectotpe male, British Museum (Natural

History).

2. S. byrsina (Meyrick) [Agriophara], 1907b: 151.

Type-locality: Khasi Hills, Assam, India.

Type: lectotype male, British Museum (Natural History).

3. S. malacobryas Meyrick [Synchalara], 1938:518.

Type-locality: Mt. Tafa, 8500 ft, New Guinea.

Type: male holotype, British Museum (Natural

History).

4. S. minax (Meyrick) [Agriophara], 1907b: 152.

Type-locality: Khasi Hills, Assam, India.

Type: lectotype male, British Museum (Natural History).

5. S. rhombota (Meyrick) [Agriophara], 1907a:981.

Type-locality: Khasi Hills, Assam, India.

Type: lectotype male, British Museum (Natural

History).
20

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

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