NOAA Technical Report NMFS 46 December 1986

Distribution and Relative Abundance

of Pelagic Nonsalmonid Nekton

Off Oregon and Washington,

1979-84

Richard D. Brodeur

William G. Pearcy

u.s. DEPARTMENT OF COMMERCE

National Oceanic and Atmospheric Administration

National Marine Fisheries Service
NOAA TECHNICAL REPORT NMFS

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1. Synopsis of biulogical data on the Blue Crab, Callinecte.< sapidus Rlthbun, by

Mark R. Millikin and Austin B. Williams. March 1984, 39 p.

2. Development of hexagrammids IPisces: ScorpaeniformesJ in the Northea,tern

!"dcific Ocean, by Arthur W. Kendall, Jr., and Beverly Viuter. March 1984, 44 p.

3. Configurations and relative efficiencies ofshrimp trawls employed in southeastern

United States waters, by John W. Watson, Jr.. Ian K. Workman, Charles W. Taylor,

and Anthony F. Serra. March 1984, 12 p.

4. Management of northern fur seals on the Pribilof Islands, Alaska, 17~()-1981, by

Alton Y Roppel. April 1984, 26 p.

5. Net phytoplankton and zooplankton in the New York Bight, January 1976 tu

February 1978, with comments on the effects of wind, Gulf Stream eddies, and slope

water intrusions, by Daniel E. Smith and Jack W. Jossi. May 1984. 41 [J.

6. Ichthyoplankton survey of the estuarine and inshore waters of the Florida Everglades, May 1971 to February 1972, by L. Alan Collins, and John H. Finucane. July

1984. 75 p.

7. The feeding eeulugy ofsome zooplankters that are important prey iterr s of larval

fish, by Jefferson T Thrner. July \984, 28 p.

8. Proceedings of the International Workshop on Age Determination cf Oceanic

Pelagic Fishes: Tunas, Billfishes, and Sharks, by Eric D. Prince (convener and editor),

and Lynn M. Pulos (editor). December 1983, 211 p.

9. Sampling statistics in the Atlantic menhaden fishery. by Alexander J. Chester.

August \984, 16 p.

10. Proceedings of the Seventh U.S.-Japan Meeting on Aquaculture, Mari:le Finfish

Culture. Tokyo, Japan, October 3-4, 1978, by Carll Sindermann (editor). August 1984,

31 p.

11. Taxonomy of North American fish Eimeriidae, by Steve 1 Upton, David W.

Reduker, William L. Current, and Donald W. Duszynski. August 1984, 18 p.

12. Soviet-American Cooperative Research on Marine Mammals. Volume I-Pinnipeds, by Francis H. Fay, and Gennadii A. Fedoseev (editors). September 1994, 104 p.

13. Guidelines tor reducing porpoise mortality in tuna purse seining, by James M.

Coe, David B. Holts, and Richard W. Butler. September 1984, 16 p.

14. Synopsis of biological data on shortnose sturgeon, Ai:ipenser brevirostrwn LeSueur

1818, by Michael J. Dadswell, Bruce D. Taubert, Thomas S. Squiers, Dunald Marchette, and Jack Buckley. October 1984. 45 p.

15. Chaetognatha of the Caribbean sea and adjacent areas, by Harding B. Michel.

October 1984. 33 p.

16. Proceedings ofthe Ninth and Tenth U.S.-Japan Meetings on Aquaculture, by Carl

1 Sindermann (editor). November 1984, 92 p.

17 Identification and estimation of size from the beaks of 18 specIes of cephalopods

from the P:lcitic Ocean, by Gary A. Wolff. November 1984, 50 p.

18. A temporal and spatial study of invertebrate communities associated with hardhottom habitats in the South Atlantic Bight. by E. L. Wenner, P. Hinde, D. M. Knott,

and R. F. Van Dulah. November 1984, 104 p.

19. Synopsis of biological data on spottail pinfish, Diplodus holbrooki (Pisces:

Sparidael, by George H. Darcy. January 1985, 11 p.

20. Ichthyoplankton of the Continental Shelf near Kodiak Island, Alaska, by Arthur

W. Kendall, Jr., and Jean R. Dunn. January 1985, 89 p.

21. Annotated bibliography on hypoxia and its effects on marine life, with emphasis

un the Gulf of Mexico, by Mamice L. Renaud. February \985, 9 p.

22. Cungrid eels of the eastern Pacific and key to their Leptocephali, by Solomon

N. Raju. February 1985, 19 p.

23. Syn,'psis of biological data on the pinfish, Lagodon rhomboide, (Pisces:Sparidae),

by George H. Darcy. February 1985, 32 p.

24. Temperature cunditlOns in the colJ pool 1977-81: A comparison between southern

New England and i'iew York transects, by Sleven K. Cook. February \985, 22 p.

25. I"drasitul0l!y and pathology of marine organisms of the world ocean, by William

J Hdtgis, Jr. (editor). March 1985, n5 p.

26. Synopsis of biological Jata un the sand perch, Diplectrum Jormosum (Pisces:

Serrdnidae), by George H. Darcy. March 1985, 21 p.

27. Proceedings of the Eleventh lrS.-Japan Meeting on Aquaculture, Salmon Enhancement. Tokyu, Japan, October 19..20. 1982. by Carl J Sindermann (editor). March 1985,

IU2 P

28. Review of geographical stocks of tropical dolphins (Stenella spp. and Delphinus

delphis) in the eastern Pacific, by William F. Perrin, Michael D. Scott, G. Jay Walker,

and Virginia L. Ca". March 1985, 28 p.

29. Prevalence, mtensity, longevity, and persistence of Aflisakis sp. larvae and Laeistorhynchus tenuis metacestodes in San Francisco striped bass, by Mike Moser, Judy

A. Sakanari, Carol A. Reilly, and Jeannette Whipple. April 1985, 4 p.

30. Synopsis of biological data on the pink shrimp, Pandalus borealis Kr6yer, 1838.

by Sandra E. Shumway, Herbert C. Perkins, Daniel F. Schick, and Alden P. Stickney.

May \985. 57 p.

31. Shark catches from selected fisheries off the U.S. east coast, by Emory D. Anderson, John G. Casey, John J Hoey, and W. N. WitzelI. July 1985, 22 p.

32. Nutrient Distributions lor Georges Bank and adjacent waters in 1979, by A. F.

1 DraxJer, A. Matte, R. Waldhauer, and J E. O'Reilly. July [985, 34 p.

33. Manne flora and fauna of the i'iortheastern United States. Echinodermata:

E('hinoidea, by D. Keith Serafy and F. Julian Fell. September [985, 27 p.

34. Additions to a revision of the shark genus Curcharhinus: Synonymy of AprionoJOIl and Hypoprion, and description of a new species of CurdUlrhinus (Carcharhinidae).

by J A. E. Garrick. November 1985, 26 p.

35. Synoptic review of the literature on the Southern oyster drill Thai' haemastoma

floridana, by Philip A. Butler. November 1985, Y p.
NOAA Technical Report NMFS 46

Distribution and Relative Abundance

of Pelagic Nonsalmonid Nekton

Off Oregon and Washington,

1979-84

Richard D. Brodeur

William G. Pearcy

December 1986

u.s. DEPARTMENT OF COMMERCE

Malcolm Baldrige, Secretary

National Oceanic and Atmospheric Administration

Anthony J. Calio, Administrator

National Marine Fisheries Service

William E. Evans, Assistant Administrator for Fisheries
The National Marine Fisheries Service (NMFS) does not approve, recommend

or endorse any proprietary product or proprietary material mentioned in this

publication. No reference shall be made to NMFS, or to this publication furnished by NMFS, in any advertising or sales promotion which would indicate

or imply that NMFS approves. recommends or endorses any proprietary

product or proprietary material mentioned herein, or which has as its purpose

an intent to cause directly or indirectly the advertised product to be used or

purchased because of this NMFS publication.

II
CONTENTS

Introduction 1

Materials and Methods

Results

Physical and biotic environment 2

Summary of nekton collected 3

Dominant species

Loligo opalescens 3

Squalus acanrhias 4

Clupea harengus pallasi 4

Engraulis mordax 5

Merluccius productus 5

Cololabis saira 5

Anoplopoma fimbria 5

Trachurus symmetricus ='

Scomber japonicus 5

Pelagic assemblage analysis 5

Discussion 6

Acknowledgments 7

Citations 7

Figures 1 - Sampling area 9

2-16 - Hydrographic data 10

17-24 - Loligo opalescens 26

25-29 - Squalus acanthias 34

30-36 - Clupea harengus pallasi 39

37-42 - Engraulis mordax 46

43-46 - Merluccius productus 52

47-49 - Cololabis saira 56

50-53 - Anoplopoma fimbria 59

54-56 - Trachurus symmetricus 63

57-59 - Scomber japanicus 66

60 - Species associations 69

Appendix Tables 1-6 - Environmental data 71

7 - Species accounts 80

iii

Distribution and Relative

Abundance of Pelagic

Nonsalmonid Nekton off

Oregon and Washington,

1979-84

RICHARD D. BRODEUR'

College of Oceanography

Oregon State University

Marine Science Center

Newport, Oregon 97365

WILLIAM G. PEARCY

College of Oceanography

Oregon State University

Corvallis, Oregon 97331

ABSTRACT

Fifteen fine-mesh (32-mm mesh) pelagic purse seine surveys were conducted

between 1979 and 1984 off the Oregon and Washington coasts. Environmental

conditions varied greatly among the years sampled, and even within years, due

to variability in upwelling conditions and productivity and the effects of a strong

EI Nino from late 1982 to the middle of 1984. In the 843 sets made, a total of

115,891 specimens from 69 taxa was collected. Most individuals collected belonged

to nine dominant taxa. Seasonal and interannual variations in the abundance and

distribution patterns of these dominant taxa are presented in detail. A recurrent

group analysis delineated four major groupings of nekton.

'Present address: Fisheries Research Institute, WH-IO, University of Washington,

Seattle, Washington 98195.

INTRODUCTION _

The compilation of quantitative information on the abundance

patterns of pelagic nekton, and the spatiotemporal variations in these

patterns, lags far behind that available for demersal nekton in

temperate shelf waters off the west coast of North America.

Hydroacoustic and aerial surveys have been used with some success to determine location and approximate abundance of concentrations of pelagic species (Smith 1970; Mais 1974; Squire 1983),

but these do not provide detailed information on species composition or biological characteristics ofthe nekton. Non-acoustic pelagic

resource surveys are generally limited in geographic extent and duration of sampling. The pelagic fisheries that exist presently, such

as troll fisheries for salmon and albacore tuna, are usually very selective as to species caught and thus do not provide much information

on the pelagic ecosystem as a whole. Large purse seines and lampara nets are less selective and have been used to describe pelagic

species compositions (Cailliet et al. 1979; Allen and DeMartini

1983), but their use has generally been restricted to nearshore environments. Large-scale assessment of pelagic resources is needed

to determine latent fisheries resources and examine potential interactions, such as predation and competition, with presently utilized

fishery stocks.

From 1979 to 1984, researchers from the College of Oceanography at Oregon State University conducted 15 cruises in coastal

waters off Oregon and Washington. The primary purpose of these

cruises was to assess the abundance and determine distribution patterns and migration routes ofjuvenile and adult salmonids. Smallmesh purse seines were used to quantitatively sample large volumes

(up to one million m3) of water. In addition to the salmonid catch,

which has been summarized in various technical reports (Pearcy

1979,1980, Wakefieldetal. 1981; Fisheretal. 1983, 1984, Fisher

and Pearcy 1985), many associated species offishes and cephalopods

were caught and biological data gathered on these species. This

report summarizes the kinds, numbers, and size distribution of the

nonsalmonid nekton catches and describes their distribution in relation to some hydrographic features of the sampling area. A species

assemblage analysis is also given based on recurring groups of

species caught in the seine sets.

MATERIALS AND METHODS _

The purse seine surveys encompass the area from approximately

Cape Flattery off northern Washington (48°20') to Cape Blanco

off southern Oregon (43 °00'). Only those stations within 56 km

(30 nautical miles) of the coast were considered in this study. The

cruises conducted in 1979 and 1980 were exploratory, with stations selected to maximize the salmonid catch, and were not

systematically arranged. From 1981 to 1984, stations were occupied

along predetermined, parallel transects positioned along latitudinal

lines and spaccd approximately 37 km apart (Fig. 1). Purse seine

sets were generally made beginning at the 37 m (20 fm) isobath

and continuing at stations 6.9, 18,28, 37,46, and 56 km from

the coast. Additional sets were occasionally made in areas of interest

such as oceanographic fronts or sites of substantial bird activity.

The nets were set at various compass bearings, but the direction

to which the net was opened was generally perpendicular to the

coast.

The survey area was sampled at least once each month between

May and September during the six years of sampling, with some

part of June sampled every year (Table I). These months were
Table I-Summary of sampling by cruise, 1979-1984.

Numbers of sets include only quantitative. round hauls

taken within S6 km of the coast. -----------

Latitudinal

Cruise range sampled Nu. of

No. Year Date (North laL) sets

I 1979 June 18-29 46°20' -43° 18' 49

2 1980 June 20-28 46'20' -44°30' 33

3 1981 May 16-25 46°35' -44 0 .10' 63

4 1981 June 9-19 46°35'-43'11' 67

5 1981 July 9-19 46"35' -44'25 71

6 1981 August 8-19 41\'35' -43' I:.' 66

7 1982 May 19-June 2 48'20' -44 '()()' 62

8 1982 June 7-22 47'20' -44'20' 56

9 1982 Sept. 4-14 47°20' -44°20' 40

10 1983 May 16-27 48'20' -44 '20' 57

11 1983 June 9-27 48'20' -43'00' 58

12 1983 Sept. 15-24 48'20' -43 '2B' 52

13 1984 June 6-20 48 '20' -43 '28' 66

14 1984 July 19-30 48'00' -44'(){l' 40

15 1984 Sept. 1-15 48"20' -44 '()()' 63 _0.-

Tot31 843

specifically selected to coincide with peak periods of juvenile

salmonid migration through coastal waters off Oregon and Washington. Only successful, quantitative round-haul sets were included

in the analysis, although fishes were often caught in nonquaatitative

sets. Most sets were made in full daylight or twilight because of

the difficulties of setting the seine at night, although several complete diel series were made, mostly in 1981.

Four different purse seines were used in the sampling. The deployment circumferences of all the seine nets, including the main body

and bunt, were similar and varied between 494 and 530 m. Fishing

depth, measured by a depth meter attached to the lead line of the

net, varied between 15 m (cruises 5 and 6) and 67 m (cruises 7-9).

All seines were made of 32-mm (stretched) or finer mesh in the

main body of the net. Additional details on net design and sampling are given in Pearcy (1984).

At the conclusion of each set, the catch was either reeled onto

the stern or brailed over the side of the vessel when large volumes

of gelatinous zooplankton made normal net retrieval difficult. All

nonsalmonid nekton were sorted from the catch and retained for

processing. Fork length (FL) measurements were taken on most

species from the tip of the snout to the end of the middle caudal

ray with the exception of somE species where total length (TL; e.g.,

elasmobranchs and some teleosts) or dorsal mantle length (DML;

e.g., cephalopods) were measured. Specimens ~IOOO mm in length

were generally measured to the nearest centimeter, and those <1000

mm measured to the nearest millimeter.

Abundances and length-frequency distributions of large :atches

of a single species were estimated by subsampling the total catch

and counting and measuring a randomly chosen subsample. An expansion factor (estimated number in whole catch/number in subsample) was applied to the subsampled length-frequency data to

estimate the length-frequency distribution of the entire catch of these

dominant species when a suitable subsample was obtained. Most

fish were identified and measured at sea, although small or difficult

to identify species were preserved and brought back to the laboratory

for further identification. The abundances of some of the smaller

individuals collected were probably underestimated due to escapement through the meshes.

Hydrographic data, collected at most stations, included surface

temperature, salinity, and chlorophyll a (except 1979 and 1980)

measurements. Water depth, ambient light intensity, and water

2

transparency were measured at the beginning of each set. Surfaceto-bottom CTD casts were made from 1982 through 1984 to obtain

vertical profiles of temperature and salinity. A complete description of the sampling and results obtained for all cruises is given

in Pearcy (1979,1980), Wakefield et al. (1981), Fisher et al. (1983,

1984) and Fisher and Pearcy (1985), The station and pertinent

hydrographic data are given in Appendices 1 through 6 for each

year of sampling.

RESULTS _

Physical and biotic environment

The general features of the summer coastal hydrography and productivity off Oregon and Washington have been described by Anderson (1964), Smith (1968), Bourke et al. (1971), Barnes et al. (1972),

Huyer (1977), Huyer and Smith (1978), Peterson et al. (1979), and

Small and Menzies (1981). The following brief description of the

physical and biotic events occurring during the summer has been

condensed from these sources.

Two processes dominate the physical oceanography of the area:

upwelling and freshwater discharge, mainly from the Columbia

River. Upwelling occurs when nearshore surface waters are displaced offshore by wind-induced Ekman transport. Mass balance

is maintained by upwelling of colder, more saline and nutrient-rich

deeper water. This upwelling activity is concentrated within bands

usually oriented parallel to the coast 9 to 18 km offshore. Intermittent upwelling events may occur from April through September

and vary in intensity and duration.

Alongshore current flow is generally much stronger than zonal

flow. During the summer months, northwesterly winds set up driving a narrow coastal "jet" which flows equatorward at speeds close

to 35 cm/s at the surface and a more diffuse undercurrent which

flows northward at much slower speeds along the bottom. Inshoreoffshore flow is highly variable and subject to wind and tidal variations. The zonal surface flow is generally offshore during the upwelling season with a compensatory onshore flow at intermediate depths.

The Columbia River plume may be distinguish:1ble as a thin surface lens of warm, low-salinity water. It may extend up to 400 km

to the southwest of the mouth of the Columbia River during the

summer in response to the prevailing northwesterly winds and

offshore Ekman transport. The inner edge of the plume forms a

distinctive front where it contacts newly upwelled water. The outside boundaries of the plume are generally delineated by the

32.5°/00 isohaline.

Primary productivity is generally high in the spring and fall

months, but is often substantially lower in the summer due to the

depletion of nutrients in the euphotic zone. The highest productivity occurs following moderate-to-weak upwelling events when

sufficient nutrients and water column stability allow sustained

phytoplankton growth. The Columbia River plume waters generally

support lower production than neighboring upwelling areas, yet

higher levels than adjacent oceanic waters.

Oceanographic conditions during 1979-84 were highly variable

during the study period, both within years and between years for

the same months. The monthly mean upwelling index for June was

much higher during 1979 than 1980 (Table 2) and sea-surface

temperatures were lower in 1979 both within and outside the Columbia River plume (Figs. 2 and 3). During 1979, newly upwelled water

was apparent inshore in the southern part ofthe sampling area (Fig.

2). Some very low salinities «10°/00) were recorded north of the
Table 2-Comparison of mean monthly upwelling indices during

May-September 1979-841 and the long-term (1948-67) mean

upwelling index for 45°N, 125°W (Bakun 1973). Monthly values

during which cruises occurred are underlined. All values are expressed as m3

/second/l00 m of coastline.

Long-term

Month 1979 1980 1981 1982 1983 1984 mean

May 34 52 12 79 35 -2 34

June 86 32 -.! 59 19 37 48

July 30 103 107 51 18 121 74

Aug. 31 96 40 38 35 37 50

Sept. 0 9 -I 12 25 3 16

IA. Bakun, Pacific Environ. Group, Natl. Mar. Fish. Serv.,

NOAA, Monterey, CA, unpubl.

Columbia River in June of 1980, indicating a possible northward

flow of plume water, although salinities were low throughout the

study area (Fig. 3).

There were few indications of strong upwelling occurring during May and June of 1981. The mean upwelling index for both

months was substantially below normal (Table 2), and sea-surface

temperatures throughout the study area were above those normally

associated with upwelling (Figs. 4 and 5). Chlorophyll a concentrations were low during these months, except for the area north

of Cape Blanco in June (Fig. 5). Strong upwelling conditions (Table

2) were evident during July at a few inshore stations off central

Oregon, and higher chlorophyll a concentrations than the previous

months were observed inshore (Fig. 6). The surface temperatures

at most stations remained very high in July. Lack of runoff to the

north and moderate upwelling to the south resulted in high surface

salinities throughout much of the study area during August (Fig.

7). Surface chlorophyll a concentrations were very low south of

Yaquina Bay, and clear oceanic water prevailed.

Unusually strong upwelling in May of 1982 (Table 2) led to cool

sea-surface temperatures throughout the study area, with the coolest

waters found inshore off southern Oregon (Fig. 8). Surface salinities

were high except near the core of the Columbia River plume. High

surface chlorophyll a concentrations were found offshore (10-15

mile stations) off Washington and were generally found inshore

along the coast of Oregon. Chlorophyll a concentrations were

average to low during the June cruise except along the southernmost transect (Fig. 9). There were sharp salinity gradients outlining

the well-defined Columbia River plume which flowed to the southwest (Fig. 9). Sea-surface temperatures were much higher in

September with low surface temperatures found only at the southernmost stations (Fig. 10). The highest chlorophyll a concentrations

were associated with the Columbia River plume.

The strong El Nino of 1983 dominated ocean conditions off

Oregon and Washington especially during the May and June cruises.

Oceanic waters were advected onshore from the west and south and

greatly depressed the thermocline. Upwelling conditions were apparent only at the southernmost stations of the May survey (Fig.

11). June sea-surface temperatures averaged 3-4°C above those of

1982 (Fig. 12). The Columbia River plume was evident over a much

broader area and apparently contributed to much of the productivity occurring during these cruises. Greater than average upwelling occurred in September (Table 2) and sea-surface temperatures

were slightly lower than in 1982 (Fig. 13). Chlorophyll a concentrations in surface waters were higher during September than May

and June of 1983 and were generally higher than September of 1982

(Chung 1985).

3

The surface temperatures were slightly elevated during June 1984

due to lower than normal upwelling that month (Table 2) and the

lingering effects of the EI Nino. Newly upwelled water was evident only off central Oregon (Fig. 14). Few stations showed high

chlorophyll a concentrations. Upwelling-favorable winds occurred

though much of early July and were manifested in the low

temperature and high salinity and chlorophyll values seen inshore

along the central Oregon coast (Fig. 15). Sea-surface temperatures

along the Washington Coast were only slightly higher than those

registered in June. Sea-surface warming continued into September

with values exceeding 14°C found throughout much of the study

area south of the Quinault River (Fig. 16). Upwelling during

September was well below that of the longterm mean (Table 2).

Chlorophyll a concentrations were fairly high in the northern area

but dropped off substantially to the south.

Summary of nekton collected

Three invertebrate and 66 nonsalmonid vertebrate taxa were collected in the 843 sets included in the study (Table 3). A total of

115,891 specimens was collected, of which 25.7% were cephalopods. The overwhelming majority (>99.9%) of the cephalopods

were of one species, Loligo opalescens, which was the most frequently occurring and numerous species collected during the study

(Table 3). The 86,059 fishes caught represented 3 classes, 12 orders,

and 31 families.

The ten most abundant families made up only 39.7 % of the total

number ofspecies but included 98.7 % of the individuals collected

during the study (Table 4). The greatest diversity was found in the

family Scorpaenidae, represented by 15 species, all in the genus

Sebastes. The majority of the species comprising these dominant

families were schooling species, with the exception of the Scorpaenidae which were mostly juvenile stages. The numbers caught

of these species often varied substantially between sets, and even

between repeat sets, within the same area.

A systematic listing of all the nektonic taxa collected in our sampling that were identified to species is presented in Appendix Table

7. The nine most abundant species, representing 95.9% of the total

specimens caught, all occurred in at least 5 % of the collections.

The distribution and relative abundance patterns and size distributions of these dominant species will be discussed in detail for each

cruise in which they were an important part of the catch.

Dominant species

Loligo opalescens-We collected 29,793 specimens ofL. opalescens

at 211 stations. This species was found in substantial numbers during each of our 15 cruises. During 1979 and 1980, mostly smaller

«100 mm DML) individuals, probably less than I yr old (Hixon

1983), were collected in greatest numbers in the southern part of

the study area (Fig. 17). Loligo had a much broader size range during 1981 and again were collected in the highest numbers off central and southern Oregon (Figs. 18 and 19). Adult L. opalescens

occurred throughout much of the study area in May of 1982 (Fig.

20), but by September of that year this species was represented only

by juveniles and was distributed south of the Columbia River (Fig.

21). The distribution ofthis species was much more limited during

the warm year of 1983 and appeared to be found farther to the north

(Figs. 21 and 22). Catches were much lower than in previous years,

especially offsouthern Oregon. These trends continued during 1984
Table 3-Phylogenetic listing of nonsalmonid pelagic nekton captured in purse seines 1979-84, including percent frequency of occurrence and total number caught.

Common and scientific names of fishes follow Robins et al. (1980), and cephalopods follow Roper et al. (1984).

% Frequency Total no. % Frequency Total no.

Common Name Scientific Name of occurrence caught Common Name Scientific Name of occurrence caught---

Market squid Loligo opalescens 25.2 29,793 Bocaccio S. paucispinis 0.7 12

Pacific c1ubhook squid Onychoteuthis Canary rockfish S. pillniger 0.8 9

borealijaponicus 04 10 Redstripe rockfish S. pmriger 0.2 14

Giant octopus Octopus dojleini 0.1 I Stripetail rockfish S. saxicola 0.6 17

Pacific lamprey Lompetra trideruata 1.8 16 Sablefish Anoplopoma fimbria 7. I 12,967

Spiny dogfish Squalus aean/hias 184 5,738 Kelp greenling Hexagrammos

Thresher shark Alopias vulpinus 0.6 5 decllgramnzus 2.9 90

Soupfin shark Galeorhinus zyopterus 1.8 19 Lingcod Ophiodon elongatus 1.3 88

Blue shark Prionace glauca 3.6 38 Brown Irish lord Hemilepidotus spinosus 0.8 15

Big skate Raja binoculata 0.6 5 Pac. staghorn sculpin Leptocottus armotus 04 7

Longnose skate R. rhina 0.2 2 Cabezon Scorpaenichthys

Pacific electric ray Torpedo californica 0.4 3 marmora/us 0.8 8

American shad Alosa sapidissima 3.8 1,185 N. spearnose poacher Agonopsis vulsa 0.6 5

Pacific herring Clupea harengus pallasi 14.9 7,803 Showy snailfish Liparis pulchellus 0.\ I

Pacific sardine Sardinops sagax 0.2 2 Jack mackerel Truchurus symmelricus 6.9 5,700

Northern anchovy Engraulis mordax 7.8 15.592 Halfmoon MeJialuna californiensis O. I 12

Whitebait smelt Allosmerus elongatus 0.8 33 Pacific sandfish Trichodon trichodon 0.1 I

Surf smelt Hypomesus pretioS/iS 1.7 491 Wolf-eel Anarrhichthys ocellatus 7.7 142

Night smelt Spirinchus starksi 0.2 6 Prowfish Zllproro sifenus 0.1 I

Eulachon 711Qleichthys pacificus 0.2 2 Pacific sand lance Ammodytes hexapterus 0.1 1

Northern clingfish Gobiesox maeandricus 0.\ I Pacific bonito Sardo chiliensis 02 2

Pacific tomcod Microgadus proximus 3.0 89 Chub mackerel Scomber japonicus 7.0 29,466

Pacific hake Merluccius productus 8.5 2,680 Pacific pompano Peprilus simillimus 1.2 31

Pacifie saury Cololabis sairo 6.9 1,436 Medusafish lcichthys lockingtoni 74 136

King-of-the-salmon Trachipterus altivelis 2.5 42 Ragfish [costeus aenigmQlicus 06 5

Darkblotched rockfish Sebastes crameri 0.1 I Pacific sanddab Citharichthys sordidus 1.6 114

Spl ilnose rockfish S. diploproa 0.5 13 SpeckJed sanddab C. stigmaeus 0.2 3

Pugel Sound rockfish S. emphaeus 0.1 I Arrowtooth flounder Atheresthes stvmias 0.1 2

Widow rockfish S. entomelas 2.7 276 Rex sole Glyptocephalus zachirus 0.1 I

Yellowtail rockfish S. j1avidus 4.0 555 Butter sale lsopsetta isolepis 0.1 I

Chilipepper S. goodei 0.2 2 Slender sole Lyopsetta exilis 0.6 7

Shortbelly rockfish S. jordani 29 82 Duver sole Microstomus pacificus O. I 1

Quillback rockfish S. maliger O. I I English sale Parophrys vetulus 0.5 28

Black rockfish S. melanops 9.3 538 Starry flounder Plotichthys stellatus 1.7 33

Blue rockfish S. mystinus 3.8 429 Sand sale Psettichthys melanostictus 0.1 7

Tiger rockfish S. nigrocinclUs 0.2 18 Ocean sunfish Mola mola 24 28

Table 4-Contribution to the total catch by the

ten most abundant families caught in purse

seines, 1979-84, in order of abundance.

No. of No. of % of total

Family species specimens specimens

Loliginidae I 29,793 25.7

Scombridae 2 29,468 254

Engraulidae I 15,592 13.5

Anoplomomatidae I 12,967 11.2

Clupeidae 3 8,990 7.8

Squalidae 1 5,738 5.0

Carangidae 1 5,700 4.9

Gadidae 2 2,769 24

Scorpaenidae 15 1,982 1.7

Scomberesocidae I 1,436 1.2

Total 98.8

4

with few large individuals caught in our sampling (Figs. 23 and

24). Overall, the highest catches of L. opalescens occurred in waters

with surface temperatures between 10° and 13 °C.

Squalus acanthias-We collected 5,738 specimens of S. acanthias

at 154 stations in our sampling. They were collected during every

cruise but were generally not abundant during the August and

September cruises. A broad size range of S. acanthias was found

over much of the sampling area in June of 1979, but abundances

were low at all stations (Fig. 25). Low abundances of mostly smaller

individuals were found during June and July of 1981 (Figs. 25 and

26). In contrast, high numbers of S. acanthias were found in 1982,

especially in the northern transects (Figs. 26 and 27). Several distinct

size classes were collected during 1983 but the numbers were lower

and the distributions compressed shoreward relative to other years

(Figs_ 27 and 28). The number of S. acanthias caught during June

of 1984 was close to that of previous cruises, but numbers again

declined by July (Fig. 29). The highest concentrations of S. acanthias occurred in waters with surface temperatures between 10°

and 14°C.

Clupea harengus pallasi-A total of 7.803 specimens of Clupea

was collected at 125 stations during this study. They occurred during every cruise, although the number of occurrences during June

of 1981 was limited (n = 2) and therefore not included as a figure.
Most C. harengus pallasi were collected off Yaquina Bay in central Oregon and off the Columbia River in June of 1979 and 1980,

respectively (Fig. 30). During 1981, the highest numbers were found

close to shore off the Columbia River but were farther offshore

off central Oregon (Figs. 31 and 32). Abundances were lower and

occurrences closer to shore in the three cruises made during 1982

(Figs. 32 and 33). The distribution was limited during 1983 to the

few areas that had relatively cold «14°C) surface temperatures

(Figs. 34 and 35). Abundances were low in 1984, and again the

catches were restricted to colder waters, especially in September

(Figs. 35 and 36). Overall, the highest catches of Clupea were found

in waters of relatively cold (10°-14°C) surface temperatures and

low «30%0) surface salinities.

Engraulis mordax-We collected 15,592 specimens of E. mordax

at 65 stations during our cruises. This species was collected during

each of our cruises, although the occurrences during June of 1979,

August of 1981, June of 1982, and September of 1983 were few

and these were not plotted. In addition, an insufficient number of

length measurements was taken on two cruises to present lengthfrequency diagrams. The distribution of E. mordax was similar for

most cruises and was centered within the Columbia River plume

(Figs. 37 to 42), where the northern subpopulation of northern anchovy is known to spawn (Richardson 1980). During June of 1981,

however, this species occurred in high numbers off southern Oregon

(Fig. 38), but many of these were juveniles and were possibly

advected to the south at this time. The distribution of this species

appears to be confined for the most part to waters of surface salinity <300f0o.

Merluccius productus-Throughout our survey, we collected 2,680

specimens of M. productus at 71 stations. Our catches of this species

were highly variable between years, with some years showing high

apparent abundances (1982 and 1984) and other years having no

catches (1980) or very low catches (1979, 1981, and 1983). During 1982 the catches in May were distributed mostly south of the

Columbia River, whereas in June they were found mostly off

Washington (Figs. 43 and 44). By September, the catches were

centered off the Columbia River (Fig. 44). A similar migration was

apparent during the summer of 1984, but the catches were still

distributed far to the north in September (Figs. 45 and 46). The

highest catches of M. productus occurred where surface temperatures fell between 10° and l3°C during most cruises.

Cololabis saira-A total of 1,436 specimens of C. saira was collected at 58 stations in this study. This species was not collected

in any of the three cruises made during May and was infrequently

caught during June and only at the offshore stations where higher

water temperatures prevailed. Cololabis saira was more abundant

during the warmer months of August and September, mostly at the

offshore stations (Fig. 47). During the warm years of 1983 and

1984, it was found fairly close to shore (Figs. 48 and 49). A wide

size range of C. saira was caught and, in some cases, several size

classes were evident (Figs. 47 and 49). This species was generally

associated with offshore waters with surface temperatures> 13dc.

Anoplopoma fimbria-Juveniles of this species were found in very

high abundances when they did occur in our samples. We collected

12,967 specimens of A. fimbria at 59 stations, mostly from June

to September. During June through August of 1981, A. fimbria

juveniles were found almost exclusively at the outermost one or

two stations along each transect (Figs. 50 and 51). There was a

5

steady increase in modal size through these months, which represented the progression of a single (age 0) year class (Boehlert and

Yoklavich 1985). Their distribution was closer to shore during

September of 1982 and during June and September of 1983 (Figs.

51 and 52). They were again distributed farther offshore during

September of 1984 (Fig. 53). Collections of A. fimbria were made

at stations with surface temperatures 14°C.

Trachurus symmetricus-During our study we collected 5,700 T.

symmetricus at 58 stations. There were, however, only four occurrences prior to September of 1982, when it occurred in low

numbers mostly south of the Columbia River (Fig 54). This species

was very abundant thereafter with the exception of the September

cruise of 1984. During May of 1983, T. symmetricus was found

only at the southernmost stations, but by June it was found in large

numbers south of the Columbia River (Figs. 54 and 55). By

September, this species had reached the northernmost transect, but

the bulk of the catches was still south of the Columbia River (Fig.

55). This species was widely dispersed throughout much of the study

area during June of 1984 (Fig. 56) and was caught at most stations

with surface temperatures exceeding 14°C. Two widely separated

length groups were evident during some months (Figs. 55 and 56),

representing both juveniles (2-3 yr old) and adults (> 7 yr old;

Mallicoate and Parrish 1981). The majority of T. symmetricus caught

throughout the study were caught in temperatures exceeding l3°C.

Scomberjaponicus-This species occurred in very dense concentrations during the warmer years of the study. Altogether, we collected 29,466 specimens of S. japonicus at 59 stations. Only two

of these specimens, both collected at a single station in June of 1982,

were taken prior to 1983. The largest collections of this species

in May of 1983 were along the southernmost transects (Fig. 57).

By June of 1983, they were distributed throughout much of the study

area with very large concentrations located south of the Columbia

River. Very high numbers of S. japonicus were taken along almost

every transect in June of 1984, with the notable exception of a large

upwelling area off the central Oregon coast (Fig. 58). Concentrations of this species were less evident during July, and its distribution was restricted to a relatively small area to the south by

September (Figs. 58 and 59). Based on our length-frequency data,

the majority of the individuals captured were approximately 2-3

years old (Mallicoate and Parrish 1981). Surface temperature appears to be the dominant factor governing the distibution of this

species. The majority of S. japonicus collected during this survey

were taken at stations with water temperatures exceeding 13dc.

Pelagic assemblage analysis

To determine which species tend to be associated together in a

similar environment, we used recurrent group analysis (Fager 1957,

1963) to group species according to their co-occurrences. In this

analysis, an index of affinity (I) is calculated for all possible species

pairs as follows:

where Jub is the number of joint occurrences, Naand Nb are the

total occurrences of species a and b, and Nb Na . This index

ranges from 0.0 (the species pair was never caught in the same sample) to 1.0 (the species pair always co-occurred). Only species which
occurred in more than 1.0% of the total number of collections were

included in the analysis. All species ofjuvenile Sebastes were combined in our analysis.

Although only occurrence rather than abundance data are used

in this analysis, it was considered appropriate for this data set due

to slightly different sampling gear used on some cruises and the

high variances associated with the numbers of many of the pelagic

species sampled. Recurrent group analysis has previously been used

with apparent success for pelagic (Cailliet et al. 1979), me~opelagic

(McKelvie 1985), and demersal fishes (Fager and Longhurst 1968;

Allen 1982).

We initially grouped our species at several different affinity levels

and decided that the 0.3 level gave the most biologically interpretable

groupings. This level is somewhat lower than that used in most of

the previously mentioned studies. The generally low affinity values

we found were probably a result of patchiness in the distributions

of many of the schooling species and the low number of species

per collection. Once the recurrent groups were formed containing

all the possible species pairs showing positive affinities, intergroup

connections were calculated based on the number of positive affinities divided by the total number of possible pairings between

members of the groups.

The results of our analysis for all our collections (Fig. 60) showed

several main groups and species that were affiliated with these

groups (i.e., showing positive affinities with only some members

of the group). A small group ofinshore pelagic juvenile fishes was

formed containing rockfishes (Sebastes spp.) and wolfeels (A.

ocellatus) and associated kelp greenling (H. decagrammus). This

inshore juvenile fish group was weakly connected to a large midshelf group which contained many of the dominant species collected

in this study. Strongly associated with this main group were American shad (A. sapidissima) and black rockfish adults (S. melanops).

Surf smelt (H. pretiosus) and two other main groupings were found

to be weakly associated. The first ofthese includes two main species

(T. symmetricus and S. japonicus) and one associated species (P.

simillimus) which have centers of distribution south of the study

area and tended to occur in our samples only during warm years.

The second main grouping consisted of three species (I. lockingtoni,

A. fimbria, and Mola mola) which were generally found along the

outer shelf region. Associated with this grouping were Pacific saury

(c. saira) and the king-of-the-salmon (T. altivelis) which have

centers of distribution offshore of the study area.

DISCUSSION _

This study represents the first large-scale survey of the pelagic

nekton off Oregon and Washington and presents new information

on the abundance and distribution of many epipelagic species of

nekton. Many of these species have not been adequately sampled

by previous midwater and bottom trawl surveys (e.g., Gabriel 1983).

By systematically sampling at predetermined stations during most

cruises and by knowing the area sampled, we could make crude

estimates of the abundances of many species which are quantitatively

sampled by the purse seines. Unfortunately, we do not have precise

estimates of the vulnerability of most of the species to our gear.

For some midwater and demersal species (Le., Merluccius productus

and Squalus acanthias). we may be sampling only the upper layers

of the population. Diel vertically migrating species. such as

Engraulis mordax, may be substantially underestimated during daytime collections compared with bottom trawl collections (Laroche

and Richardson 1980). Moreover, highly attenuated species (Colo6

labis saira) and juveniles of most species were observed to escape

through the meshes during pursing and retrieval of the net, and captures of these species may be considered only as incidental.

The usefulness of this study lies not in estimating precise absolute

abundances, but rather in examining seasonal, interannual, and

geographic changes in the relative abundance of the various species,

particularly those which dominate the catches. Many species of

pelagic and demersal fishes are known to show substantial variation in year-class strength (Hollowed et al. in press) and independent (nonfishery) estimates of the stock size are important in

characterizing the variability in recruitment patterns.

Along with the substantial environmental changes that occurred

within one sampling year were major changes in the relative rank

order of abundance (ROA) for the different cruise months. The ROA

of the dominant species for May, June, July-August, and September

for all years combined are given in Table 5. Although some species

were among the ten most abundant for each of the four time periods

examined, several trends were apparent in the relative rankings.

Several species with cold-water affinities (Loligo opalescens,

Squalus acanthias, Hypomesus pretiosus, and Sebastes spp.

juveniles) generally decreased in relative ROA through the summer. Other species with more warm-water affinities (Anoplopoma

fimbria juveniles and Cololabis .mira) generally increased in relative

importance through the sampling season. The remainder of the

dominant species either showed no change or were inconsistent in

their relative ROA patterns. Seasonal variations in the abundance

of Sebastes spp. and A. fimbria may also reflect ontogenetic changes

as the juveniles settle out to the adult demersal habitat.

Although some differences in sampling effort occurred among

years, the June cruises provided us with a time series of observations with which to make interannual comparisons (Table 6). The

rankings during 1978, 1980, and 1981 were similar despite some

differences in environmental conditions among the three years. During June of 1982, juvenile rockfishes (Sebastes spp.) and northern

anchovy (E. mordax) decreased in relative importance while two

higWy migratory species (M. productus and T. symmetricus) became

more important. Major interannual changes in the ROA were

associated with the occurrence of the strong EI :-.Iino event during

June of 1983 and 1984 (Table 6). The dominant taxa (Loligo

opalescens and Sebastes spp.) from 1979-82 had much lower ROA's

during the final two years. They were replaced in relative rankings by two species (Scomberjaponicus and Trachurus symmetricus)

that usually have centers of distribution off California. Other species

with warm-water southern or offshore affinities (Peprilus simillimus,

lcichthys lockingtoni, and Mola mola) showed increased abundances

during the latter two years (Pearcy et al. 1985).

Spatial variations in abundance may be difficult to interpret

because of the many different migration patterns exhibited by these

pelagic species. The marine distribution of several anadromous

osmerids and estuarine-spawning C. harengus pallasi may change

seasonally depending on the spawning cycle. Adult LoNgo opalescens are known to move southward and shoreward in late spring

and early summer to spawn off Oregon (Starr 1985), whereas adult

E. mordax move offshore to spawn, apparently within the Columbia River plume, during summer (Laroche and Richardson 1980).

Other species undergo north-south seasonal migrations associated

with feeding. For instance, M. productus spawn off southern

California in the winter, and the juveniles move northward along

the shelf in the spring and occupy feeding grounds within our study

area during the summer (Bailey et al. 1982). During August, they

begin to move offshore and occupy greater depths before migrating

south in the fall. During the warm year of 1983, there was a well-
Table 5-Rankings of relative abundance of dominant

nonsalmonid nekton by cruise month. Only those species

ranked among the most abundant at least two months are

listed. Rankings below the top ten are indicated by a

dashed line. Numbers of cruises in each sampling period

are given in parentheses.

Rankings

JulyMay June August September

Species (3) (6) (3) (3)

Scomber japonicus 3 1 6 I

Loligo opalescens 2 2 4 6

Clupea harengus pallasi 5 6 2 4

Engraulis mordax 1 8 3 7

Anoplopoma fimbria 9 I 3

Squalus acan/hias 4 5 5

Merluccius productus 7 7 8 5

Trachurus symmetricus 8 3 9 8

Cololabis saira 10 2

Sebastes sp.' 10 4

Hypornesus pretiosus 6 10

'Includes juveniles of several species.

Table 6-Rankings of relative abundance of dominant

nonsalmonid nekton for June cruises, 1979-84. Only those

species ranked among the most abundant at least two

years are listed. Rankings below the top ten are indicated

by a dashed line.

Rankings

Species 197919801981 1982 1983 1984

Loligo opalescens I I I I 5 4

Squalus acan/hias 5 6 4 3 3 3

Clupea harengus pallasi 3 3 9 4 6 5

Sebasres sp.' 2 2 2 7 10

Trachurus symmetricus 5 2 2

Engraulis mordax 4 4 8

Scomber japonicus I I

Merluccius productus 2 6

Anoplopoma fimbria 8

Sebasres melanops 6 9 9

Cololabis saira 5 10

Alosa sapidissima 8 10

'Includes juveniles of several species.

defined northward movement of S. japonicus and T. symmetricus

schools through the study area. Some evidence exists that these

schools remained within the study area during the winter of 1983-84

(Pearcy et al. 1985); both species were widely distributed throughout

the study area in 1984 and S. japonicus was found in large numbers

off the Canadian coast (Ashton et aJ. 1985).

Oceanographic measurements taken concurrently with our collections allow us to examine the environmental preferences of these

species and may be used to predict which species are likely to occur in a particular water type. Our results suggest that seemingly

small changes in surface temperature may be associated with drastic

changes in species composition and abundance. For instance, a

1-4°C increase in mean surface temperature between May and June

of 1982 and 1983 (Fisher et al. 1984) was associated with a complete shift from an assemblage dominated by northern temperate

species (Loligo opalescens and Squalus acanthias) to an assemblage

dominated by more southern fauna (Scomber japonicus and

Trachurus symmetricus). Surface temperature alone, however, may

7

not be an accurate indicator of environmental conditions. A major

northward shift in the location of the subarctic boundary caused

by an intensification of the northward-flowing Davidson Current

during the EI Nino resulted in numerous biological anomalies at

all trophic levels (Wooster and Fluharty 1985; Mysak 1986). In

essence, our study illustrates the indirect effects of a major environmental perturbation, such as the 1982-84 EI Nino, on nektonic

organisms.

ACKNOWLEDGMENTS __

We thank Craig Banner, Alton Chung, Joe Fisher, Dean Gushee,

Jon Shenker, and Waldo Wakefield for their generous assistance

in the field. We are indebted to Wayne Laroche for identifying and

measuring the juvenile rockfishes. Alton Chung, Joe Fisher, Harriet Lorz, and Eric Rexstad assisted with the data compilation, computer analysis, and figure preparation. This study is the result of

research funded by the Northwest and Alaska Fisheries Center of

the National Marine Fisheries Service. NOAA.

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Oregon in 1985. Oregon Dep. Fish. Wild!. Inf. Rep. 85-9,41 p.

WAKEFIELD, W. W., J. P. FISHER, and W. G. PEARCY.

1981. Studies ofjuvenile salmonids off the Oregon and Washington coast, 1981.

Oregon State Univ. Lab. Ref. 81-13, 51 p.

WOOSTER, W. S., and D. L. FLUHARTY (editors).

1985. EI Nino North: Nino effects in the Eastern Subarctic Pacific Ocean. Wash.

Sea Grant Prog. Pub!. 85-3, Seattle, Wash., 312 p.
~-....-nl.~~, YAQUINA

rr'" BAY

I ,'iJ

Figure 1

Location of sampling area and transect lines used in the study.

9
Surface temperature (OC) Salinity (%0)

Figure 2

Station locations and isopleths of surface temperature and salinity, June 1979.

10
QUINAULT R.

47° 47°

25 20

o. , O'O~~

46°

.:.. 14

Surface temperature (0C) Salinity (°/00)

Figure 3

Station locations and isopleths of surface temperature and salinity, June 1980.

11
Figures 4 through 16

Station locations and isopleths of surface temperature, salinity, and chlorophyll a, summer 1981-84.

MAY 1981

I I I I

Surface temperature (OC)

I I I I

QUINAULT R.

28

.

J~~"~~, .....

Salinity (°/00)

Figure 4

12

I I I I

CAPE FLATTERY

QUINAULT R.

Chlorophyll a (Jlg/L)
/7···15 C)

·f 14 • •• ·"14

1\...(

Surface temperature (OC)

JUNE 1981

QUINAULT R.

Salinity (°/00)

Figure 5

13

1 I I I

QUINAULT R.

Chlorophyll a (J,lg/L)
JULY 1981

I I 1 I

CAPE FLATTERY CAPE FLATTERY

QUINAULT R.

.....

ft3.!: I)

• 14

r-;

26 COLUMBIA R.

. {.

28

Surface temperature (0C) Salinity (°/00) Chlorophyll a (J.tg/L)

Figure 6

14
I I I I

CAPE FLATTERY

Surface temperature (OC)

AUGUST 1981

I I I I

Salinity (%0)

Figure 7

15

I I I I

QUINAULT R.

Chlorophyll a (j.lg/L)
MAY 1982

Surface temperature (DC) Salinity (°/00)

Figure 8

16

Chlorophyll a <J.tg/L)
JUNE 1982

I I I I I I I I I I I I

II 32

47° .~...

32~2

.." .. .. ....

46°

Surface temperature (0C) Salinity (%0) Chlorophyll a ~g/L)

Figure 9

17
SEPfEMBER 1982

YAQUINA BAY 1.0

..;: }... 33

Surface temperature (0C) Salinity (°/00) Chlorophyll a {J.tg/L)

Figure 10

18
MAY 1983

1250 1240 1230 1240

I I I I I II I I I I I r-;---,-....,--'...---:;---,---.-.,..:-:;=---7-,....'"~

[ 00

--( O",~cr'

I I I I

0

o 0 0

00 0 45 0

0

,~ YAOUINA BAY

440 8.0

440

Surface temperature (OC) Salinity (Ofoo) Chlorophyll a ~g/L)

Figure 11

19
JUNE 1983

...

124°

I I I I

:Ol4~.~ lCAPE FLATTERY

6.0

.~

5.0 ) ..

1.0

125°

I I I I

...

...

I I I I

13 CAPE FLATTERY

,~

16~

I I I I

<1.0

: ...

45° .... 45°

1.0

.f YAQUINA BAY

44° 44°

Surface temperature (0C) Salinity (°/00) Chlorophyll a ~g/L)

Figure 12

20
SEPTEMBER 1983

I I I I

CAPE FLATTERY

16

'°1 47° 47° '.'t'

46°

Surface temperature (0C) Salinity (°/00) Chlorophyll a (j.lg/L)

Figure 13

21
JUNE 1984

Surface temperature (ec) Salinity (°/00)

Figure 14

22

Chlorophyll a {Jig/L)
JULY 1984

I I I I

32 33

I I I I

48° 48°

47° l 47°

I

E

46°

Surface temperature CC) Salinity (°/00) Chlorophyll a (J.tg/L)

Figure 15

23
SEPTEMBER 1984

I I I I I

}...

32

15

..

Surface temperature (OC) Salinity (°/00) Chlorophyll a (j.tg/L)

Figure 16

24
Figures 17 through 59

Distribution, relative abundance and length-frequency distributions of dominant species in summer catches, 1979-84.

25
Loligo opalescens

100,------------------,

48° 48°

QUINAULT R.

47° 47°

45° 45°

o~ YAQUINA BAY OPCbo+

+ = 0 0

44° CP+ o = 1-5 I-S

o = 6-10 44° 0 6-10

o =11-25 o =11-2S

o =26-se =26-S0

o =SI-le0 o =SI-100

0= > 100 o· > 100

43° 43°

?5

50

?5

... '"

'" 50 I:

25

0

35 60 85 110 135 160

DORSl=lL Mf=lHTL£ LENGTH (MM) DORSAL MANTLE LEHGTH <MM>

June 1979 June 1980

Figure 17

26
Loligo opalescens

QUINAULT R.

o

\-5

o 6-\0

o "\\-25

o "26-50

0"5\-\00

0" > \00

100 ~-----------------,

75

50

+ 0 +<C9

+ - 0

() 0 - \-5

o - 6-\0

o -\\-25

-26-50

0-5 \-\00

0- > \00

100~====~=====:::::===~ 75

50

1S0

DORSAL MANTLE LENGTH <Mn)

May 1981

DORSAL MANTLE LENGTH (Mn)

June 1981

Figure 18

27
Loligo opalescens

CP++~

" + - " 44° 1-5 + o - 1-5 44° o - 6-1" 6-1"

o -11-25

0-26-5"

0-51 - 1""

O· > I""

43° 43°

100

75 75

50 50

25 25

0

35 60 85 118 135 160 60 85 110 135 160

DORSAL MANTLE LENGTH CM") DORSAL MANTl( L[NGTH eM">

July 1981 August 1981

Figure 19

28
Loligo opalescens

125°

48° 48° ++0+

0++

0+0 +0+

47° 470. 0+ + + +

+ 0 Q. r- <0..

+ 0+ e

++ .. 46° +0 0

oQoo+ ....

+ s 0 0

44° o - 1-5 1-5 44° o - S-10 S-10

o -11-25 o -11-25

o -2S-50 o -ZS-50

0-51-100 0-51 -\00

0- > 1011 0-> 100

43° 43°

,.. ::==============~ 75

5. 5.

25 25

6. 85 II. 135 168

• .,....,.~...... ........,..e;...,.,....,....,.,.....9'l'....-.......~..,....,.J

35

DDRSAL MNTLE LENGTH eM) DORSAL MANTl£ l[HOTH eM")

May 1982 June 1982

Figure 20

29
+ - 0

o • 1-5

o - 6-111

0-11-25

o -26-50

0-:11-11111

0- > 11111

+ + ++

o

1-:1

6-111

Loligo opalescens

1250

+ +

o

+ +++

+ +

+ +++

+ + +

75

59 59

25 25

69 95 1t9 135 169

9L,-...~.......,..,..,~..y.........J:;lT'-o'''''''''''~''''''''''''''~''''''''

35

DORSAL MANTLE LENGTH enM) DORSAL MANTLE LENGTHCn">

September 1982 May 1983

Figure 21

30
Loligo opalescens

++ + [CAPE FLA_:r_T_E_R~Y H

\ .. + 0

0

+ + + ++

+ 3 II + - 0

44° 0++ o • 1-5 o - 1-5

44° + ++ o - 6-10 o - 6-10

o -11-25 0-11-25

o -26-50 o -26-50

r o -51-100 0-51-100

0-> 100 0-> to" f43° +++ 43°

100 100

75 75

50 50

25 25

0 0

35 .0 95 119 135 1.0 35 .0 95 110 135 1.0

DORSAL MANTLE LENGTH enn> DORSAL MANTLE LENGTH <nn>

June 1983 September 1983

Figure 22

31
100~==============::,

o

1-5

6-10

+ ..

+ +0 + +

o

+ + 0 +

Loligo opalescens

1250

+++*

+ + +0

100,------------------,

75 75

50 50

25

DORSAL MANTLE LENGTH eMM) DORSAL MANTLE LENGTH <M">

June 1984 July 1984

Figure 23

32
Loligo opalescens

+.IJ

45° + + ++ 45°

+++-.+

YAQUINA BAY +

++ .... + _ 0

44° ++ ++

o _ 1-5

o _ 6-10 44°

0-11-25

c26-50

0-51-100

0- > 100

43° 43°

100

75

50

25

0

35 .0 OS 110 135 ,.0

DORSAL MANTLE LENGTH eM")

September 1984

Figure 24

33
Squalus acanthias

1250

QUINAULT R. QUINAULT R.

30

+ a 0

o 1-5

o 6-10

0=11-25

o =26-50

o ~51-I,a0

o a ) 100

...

+ + ++-tC

20

.0

50~====~=======;

o

1-5

6-10

=11-25

~26-50

o ~51-100

0= ) 100

+0

20

760 920 10B0

TOTAL LENGTH <MN) TOTAL LENGTH (MM)

June 1979 June 1981

Figure 25

34
Squalus acanthias

125°

000+

YAQUINA BAY

+ + + +

+ ++

s.~~==========:::

+ - 0

l + - 0

44° o • 1-5 o - 1-5 44° o - 6-10 o - 6-10

o -11-25 o -11-25

o -26-50 0-26-50

o -51-100 o -51-100

0- > 100 0- > 100

•• ••

J. J.

2. 2. I.

TOTAL LENGTH CMM) TOTAL LENGTH (M",

July 1981 May 1982

Figure 26

35
Squalus acanthias

929 19a0

•

1-5

o - 6-t.

o -t 1-25

-26-5.

0-51 - 1••

0- ) 10.

7.9

++

+ ++

TOTAL LENGTH en">

May 1983

+ + 0>

29

39

S9 r-----------------,

.9

Figure 27

36

o

1-5

o - 6-te

o -11-25

0-26-50

0-51-100

0- > 180

June 1982

+00

TOTAL LlNGTH (MM)

29

39

.9

S9 r-----------------,
8

1-5

CJ - 6-18

o -11-2:1

o -26-58

o -:l1-18e

0- > lell

+

+€1

+

Squalus acanthias

1250

8

1-:1

6-18

o -11-2:1

-26-:18

0-:11-188

0- > te8

+ 00

•• ••

3. 3.

2. 2.

I.

TOTAL LENGTH CM"> TOTAL LENGTH enn>

June 1983 September 1983

Figure 28

37
440 600 760 920 1080

01,-..;"...,...-.'ir.Q.,J.l.l.l.Y,l,l..,l,l.."'"T""~"'<;>''''''''~''''''~r.-.'

280

+ + + +

QUINAULT R.

+

+ + ++

10

20

40

30

440 600 760 920 t989

)CAPE FLATTERY

0 00+<

0::',\

t

UINAULT R.

0++

DODD?

(,

"00

'0

S

1-5

o .. S-le

o ~11-25

-26-5S

o -51-ISS

O· ) ISS I

J

43

0

'--'--'---'---'--L--'---'---'--L-L-J........!.-l.J.

TOTAL LENGTH eM"> TOTAL LENGTH (MM)

June 1984 July 1984

Figure 29

38
Clupea harengus pallasi

125°

QUINAULT R.

..

0l,......~'T""".,.<;l;W.J.I.II;>..,~~T"""""'~'T"""~ .....,..,..,J

105 135 165 195 225 255 285

.... e

<> .. 1-5

¢ .. 6-le

<> -11-25

<> "26-se

(> =51-lee

0- > lee

June 1980

FORK LENGTH (MM)

20

40

.. .. e

44° .... o .. 1-5 .. ¢ .. 6-le

<> "11-25

<> -26-5e

(> "51-lee

O· > lee

43°

60

40

20

0

10S 13S 16S 19S 22S 2SS 20S

FORK LENGTH (MM)

June 1979

Figure 30

39
Clupea harengus pallasi

QUINAULT R.

e + - e

44° I-S <> = I-S

6-10 44° <> • 6-le

<> =11-2S <> -1I-2S

o =26-se o -26-se

o -SI-Iee o -sl-lee

0- > lee 0- > lee

29

9L,....,..,I,l,.l;I;J..........,..,I,I;.;i,J,.1..l;I.J.......~~......~"'T".,....J

U:5 IJS 165 195 225 25:5 28:5

FORK LENGTH cr1")

May 1981

29

9 l,.................,J.,l.J.W,J.I.l,.l;I,J,.1..W.l.ITL,...,.,........,..........<.>r""'" us t JS 165 19S 225 255 285

FORK LeNGTH eMn>

July 1981

Figure 31

40
o

I-~

o - 6-10

o -11-25

<> -26-50

0-51-100

<> - > 100

o.

+ + 0

+ ++

•• • o • I-~

<> • 5-1'

() -11-2~

Clupea harengus pallasi

1240

.....

S0~============~

40 40

20 20

o 'r.-~""T~"""M""".y,J""""""'y;>'''''''''''''''''~''''''''''''' 18:5 135 165 195 225 255 295

0'r-.....l,.1f...........r.....,.y-.J.....,""Y......J.,l,lT""""'~'T'-

18S 135 165 195 225 255 285

F"ORK LENGTH 01") fORK LENGTH <MM)

August 1981 May 1982

Figure 32

41
Clupea harengus pallasi

+

+-100

45°

+ ++

YAQUINA BAY

+ +

+ = 0

() = 1-5 44° () s 6-10

<:> =11-25

o =26-50

<> =51-100

<> = > 100

43°

sa

.a

2a

•las 135 ISS 195 225 255 285

fORK LENGTH (MM)

September 1982

fORK LENGTH (MM)

aly-.-,~..,...~.o,J.,l,J.,..,..,..,,...Q.,I;l.,....~..,...~ .......,........J

10S 135 165 195 225 255 285

June 1982

2a

.a

Figure 33

42
Clupea harengus pallasi

+<> + +

<) + ++

QUINAULT R.

+ + ++ 43°

S•

••

2.

•'.5 135 1S5 195 225 255 2.5

fORI< LENGTH (MM>

June 1983

fORK LENGTH (MM)

May 1983

• l,....<;<;I.J.,J.,Jl,.l.l,l;I,J;;Q,<~:;O".......,~~,.........~.,.......J

105 IJS 165 195 225 255 285

2 •

••

+<>~<>-K> +

+ = 0 + = 0

44° <) = 1-5 <) = 1-5 +++ 44° <> = 6-10 6-10

<> -11-25 =11-25

=26-50 =26-50

o =51-100 0=51-100

0= > 100 0- > 100

Figure 34

43
+ - II

o - t-s

(> - 6-tll

<> -11-2S

<> -26-S9

o -SI-199

(> - ) 199

++ + ++

+ + + +++

+ - II

o • 1-5

(> - 6-tll

<> -11-25

<> -26-SII

o -St-tllll

(> - ) tllll

Clupea harengus pallasi

1250

+

++

+

+ + +

•• ••

2. 20

t3~ 165 19S 225 255 295

rORK LENGTH (M") fORK LENGTH (MM)

September 1983 June 1984

Figure 35

44
Clupea harengus pallasi

+ + ++

+ ++

+ - 0

() • 1-~

6-10

-11-2:5

+ + + +to

o

1-:5

o - 6-10

<> -11-2:5

-26-:50

0-:51-100

<> - > 100

+

++ +

+

48° 48° + + + + +++0

-<00

+ + ++

47° + ++ +

.9 '9

29 29

9'f-,-,~..,.,~.......,,.......~"T""~J..Y.,1,.L,.l,J.lf.l.~ .........,J

185 tJS 165 195 225 255 285

9l,-.-~.,..,J.,I.,J,.L,J.J,M'1,.l.l;;y;J.,.t;I;l;l;>~-.-.',.....~ .........,J

185 135 165 195 225 255 285

fORK LENGTH eM") fORK LENGTH en")

July 1984 September 1984

Figure 36

45
Engraulis mordax

1250

o

1-5

o • 6-10

0-11-25

-26-50

0-51 - 10'"

0- ) 100

QUINAULT R.

YAQUINA BAY

+ .. ++......

+ - 0

o • 1-5

o - 6-10

o -11-25

o -26-50

o ~51-100

o ) 100

<5

30 '"

OJ

30

115 130 160 175

o '-r-~~~~......,=>...,.-'--,--'-r'--.-'--.-'--,'-.,..-L......,~

10"

15

115 130 160 175

0'-r---.--'=;=i--,...L,--'-T-L..-.L,.-'-r-~~-r-~--r--.-J

100

15

fORK LENGTH (MM) fORK LENGTH (MM)

June 1980 May 1981

Figure 37

46
Engraulis mordax

+ + .......... YAQUINA BAY

+ + ++++ + + .......

+ 0

o 1-5

o • 6-10

o =11-25

o ~26-50

o ~51-100

O' > 100

+ - 0

o • 1-5

o - 6-10

o -11-25

o a26-50

o a51-100

0= > 100

45 45

Je Je

15

115 IJe 145 Ise 175

e L,-.-JL..J--,l--.-t~......;=L,....J.,,-L,...L,-'-r.L.-.r:::;::l=y=....,..J

lJe 145 Ise 175 lee eL,L..J=..-~~.--~..--,rG=L,.-'-r~~-.-....-J

lee

rORK LENGTH (nM) rORK LENGTH (Mn)

June 1981 July 1981

Figure 38

47
September 1982

roRK LEHSTH CM)

15

No data avilable

3.

Engraulis mordax

125°

+ ++

+ ++ +

+ - e

44° + o - 1-:5 + o - 6-10

o -11-25

-26-5e

o -51-lee

0- ) lee

43°

••

• 5

3.

15

•1•• 115 13. ,.5 1•• I?>

fORK LENGTH (M")

May 1982

Figure 39

48
o

I-~

o • &-10

o ·11-2~

·2&-~0

o ·~1-100

O. ) 100

+

+++

+ + +

+ + +

o

I-~

&-10

Engraulis mordax

1250

+

+++

+ ++

++++

++ + +

45 45

39 39

15 15

115 139 145 169 P5

9'-r-~~-,-~~r-,....a...,.-J-,-<-'...L..,.-'-r'>=;="-r-9"""'"

115 139 145 169 17S 199

9L..--~~,-~..-r:?-,..L,,..J..,...L,.-'-r-'=f"-r-~-r-.-J

199

rORK L[NGTH ("") rORK LENGTH en">

May 1983 June 1983

Figure 40

49
Engraulis mordax

o

1-:5

6-10

+

+

0+ +

+ 0 + ... tCOLUMBIA R.

+ + + +

.. :====='========~

48° 48° + + + + + + + +

+++*

+ + ++

47° 47° + ++ +

<S .5

3. 3.

No data availai.lle

15 15

115 13. 145 I •• 175

.L,.-~~-,-~~.-~~,-.~---.~~--r-.-J

I ••

fORK LENGTH (MM) fORK LENGTH CM")

June 1984 July 1984

Figure 41

50
Engraulis mordax

{

APE FLATTERY../

+ ++ + "'-~-~

--I + + + +

+++\

+-+0 +

+ +++

++ ++

+

o

1-:5

6-10

6°1 .5

30

IS

f"ORK LENGTH (MM)

September 1984

Figure 42

51
S0,-----------------,

Merluccius productus

1250

S0~=========::::=::;

00+ 0

+ - 0 + - 0-

44° 0 - 1-5 + 0 - 1-5 + 44° <) - 6-10 <) - 6-10

0 -11-25 o -11-25

o -26-50 o -26-50

<> -51-lil0 <> -51-100

0- > 10il 0- > 100

40 40

30 30

20 20

3S0 410 470 S30 SS0

o.......~.....,r'"""'"~TT"" ......J.,.J..JW.......J..,J.,JW......y.,.t.-,..,.,., 2S0

10

fORK LeNGTH ("") fORK LENGTH <MM)

July 1981 May 1982

Figure 43

52
+. 0

o iii t-~

o - 6-\0

<> -\ 1-25

<> -26-50

<> -51-100

<> - ) \011

+ +

+ +++

+

+ +

+ +++

+ + +

i-

+ - 0

o - \-S

o - 6-\0

<> -\\-2S

<> -26-S0

<> -5\-\00

o - ) \00

+

Merluccius productus

1250

+ +.

++ -<>

++0+

+0<)

+

4. 4.

3. 3.

2.

59.

2.

35. 410 47. 53. 59.

• L,-.-~"""'"T~~,.o,.J.,l,J..Y..J..l..,l,J.,Jy,J-.Q,.J.,J,-~,I,J.,J

290

,.

fORK LENGTH (MM) rORK LrhGTH en">

June 1982 September 1982

Figure 44

53
+

Merluccius productus

124°

+ + ++

48° 48° + + ++ + + + +

+++*

+ +0+

+ 0+

47° 47°

+++

45° + + + ++ 45°

00 + +

YAQUINA BAY

+ + + +++

+ = 0 + • 0

44° ++ + ++ o = I-~ 0000. o • I-~

¢ • 6-10 ¢ • 6-10 44°

0 ·11-2~ o ·11-2~

=26-~0 ·26-~0

<> ·~1-100 <> ·~1-100

O· > 100 O· > 100

43° 43°

50 ~-----------------, 50 ~-----------------,

40 40

30 30

20 20

10 10

350 410 470 530 590

0l,-...-~.......,..,.l,Jl,J,.l..y..L,J..,:l,J,.l.,J.l,,J..,1,.r;y.,J.,J:;J..J..,....~"'"

350 410 470 530 590 290

0l,-...-~.......,......,l,J,.l..y..o,<;i.l,.l.-, ....l,.ly-.."T<?T-."T"""~"'"

290

rORK LEHGHI (MM) rORK LEHGTH (MM)

June 1984 July 1984

Figure 45

54
Merluccius productus

125°

)"CAPE FLATTERY

+ .+ +;

48° ++.,,\

\

-to+ \

\

+

++.

+ - 0

44° ++ ++ <> - 1-:5

<> - 6-10

<> -11-2:5

-26-:50

<> -:51-100

<>. > 100

43°

50

40

30

20

10

0

2S0 350 410 470 530 SS0

rORK LENGTH (MrO

September 1984

Figure 46

55
Cololabis saira

o

1-:5

(> • 6-19

<> -1\-2:5

o -26-50

<> -:51-100

<> - > 100

39:===============~

QUINAULT R.

29 29

No data available

19

17. 2 •• 23. 2 •• 2 ••

9lr-..~~~.,.Q...r;:rM,l......--.--.~~....,....J;1,.I;ItL,.I.,J

178 208 23. 2.8 2.8 ,.9 8lr-..~~~~~r..-~......,~~""'"T~~"".,...J

148

fORK LENGTH en"> rORl( LrHiHH eM">

June 1980 At:gust 1981 L- . -----'

Figure 47

56
Cololabis saira

QUINAULT R.

+ ++

+ + +

+ + +

+

+ 0 ++<>

+ III + - III

44° \-5

<» -\-5 44° +++

o - 6-\111 o - 6-\111

<> -1\-25 <> -\\-25

-26-5111 o -26-5111

0-5 \-\111111

0- > \111111

20 20

10 10

170 200 230 260 290

o L,....~~.y4J-.J..J......,....,..............,'-,J.,........c.y...............,."..........

170 200 230 260 290 140

0l,..~~rr-~.......,,.........~....~l.Qc;J,-.,.L;l,....,J:~.....,J

1<0

rORI< L[NGTH <nn> fORK LeNGTH enn>

June 1983 September 1983

Figure 48

57
Cololabis saira

+

20

10

++ ++

+ - 0

<> = \-5

<> - 6-\0

o -\\-25

-26-50

fORK L[NBTW 0110

September 1984

Figure 49

58
Anoplopoma fimbria

125°

+ = 0

44° .... o = 1-5 44° 0 6-10

o =11-25

o =26-50

o =51-100

0=> 100

49 49

39 39

29 29

199 239 279 319

rORK LENGTH CM"> FORK L£NGTH enn>

June 1981 July 1981

Figure SO

59
Anoplopoma fimbria

1250

59~============'

QUINAULT R.

+ + "'"

+ . 1I + - 1I

44° + 0 . 1-~ o • 1-5 44°

C - 6-111 o • 6-11l

o "11-2~ o -11-2~

o "26-~1l o -26-50

o -~1-llllI 0-51-100

0- > IlIll 0- > 100

49 49

39 39

29 29

279 319

FORK LENGTH eM"- rORK LENGTH enM)

August 1981 September 1982

Figure 51

60
Anoplopoma fimbria

190 230 270 310

+++0+++

50

40

30

20

10

0

110 150

10

40

20

30

50,--------------------,

+ + ++

" + - " 44° 1-:1 o - I-S + ++ 44° 6-1" o - 6-\"

0-\1-2:1 o -1\-2S

-26-:1" o -26-S"

0-5 \-\"" o -SI-tllll

0" ) 100 0" > \0"

raRK LENGTH CMM) rORk LENGTH eM")

June 1983 September 1983

Figure 52

61
Anoplopoma fimbria

c+ ++

L

+O'El

+ . " 44° ++ ++ 0 . 1-5

6-1"

,.:============~

••

3.

2 •

19. 23. 27. 31. I'. • 1,-.-.~.........~~~~........-..4;1.J.1;#~....-~,..J 110

rORK LeNGTH <M")

September 1984

Figure 53

62
Trachurus symmetricus

o

1-5

6-10

+

+++

+ +l!)O ++

+ - 0

o • 1-5

o - 6-111

o -1l-25

0-26- 511

0-51-11111

0-> 11111

+0

48° 48° ++ + +

+ + ++

+ + ++

47° + ++

~470 ++++

•• ••

•• ••

2. 2 •

• \-mI~'?-"""""",,,,,,,,,,,,,,,,,,,,,~,,,,,,,,~,,,,,,~~

2SB 319 368 418 468 518 568 6t8

fORK LENGTH <nM) fORk LENGTH U1t"

September 1982 May 1983

Figure 54

63
Trachurus symmetricus

125°

48° 48°

+ + +

QUINAULT R. + + ++

47° + ++ 47° ++

:~

45° +0 45°

+ +

0

+ - 0

44° o - 1-5 44° o - 6-10

0-11-25

0-26- 50

0-51-100

O· > 100

43° 43°

60 60

40 40

20

o

2Se 310 360 410· 460 510 sse Gte

rORIC LENGTH <MM)

20

o

260F~""""""J'"""'~~""""~""""- 310 360 410 460 510 560 610

rORK L[HGTH (MM)

June 1983 September 1983

Figure 55

64
+ + + +

Track-urus symmetricus

125°

0+0+

0

44° I-~ 44° 6-111

o ·11-2~

·26-~1I

o ·~1-1011

o· > 1110

43° 43°

•• ••

•• ••

2. 2.

•

26B JIB J69 41B 469 :519 :569 6tB

fORK L£HGTH ("'0

.~"",,¥IJWJtl¥........"T"'"~"""""":':l'~'!."f}~~ 269 319 369 04U 460 519 :569 61B

rORI< LENGTH (MM)

June 1984 July 1984

Figure 56

65
QUINAULT R.

0++

Scomber japonicus

125°

I I

+ + + +

+ + + +

125;::~~========'=====::::::~

0 0

44° 0 . 1-:1 1-:1

a - 6-10 6-10

o -11-2:1

-26-:10

o -:ll-le0

D- ) 100

100 100

75 75

50 50

260 290 320 350 380

o ...........,J..,J,.l.l.JL,J.,J..y.,,L.J.,.L,J.,J..,L.J.,,L.J.,.J:;y:;>'O'~ ..............,J

230

June 1983

FORK LENGTH (n")

25

Figure 57

66

300

May 1983

rOR.c LENGTH enM>
+ + ++

+ +0+

125 ;::'='='='=~========~

[):J + +

Scomber japonicus

1230 1250

...-.---.,....:...:;:..~.....-r--r---.,.....:..r-~.,...-r-,,-...;..:;

12'~===':;:~=========~

188 188

75 75

.8 5B

25 25

298 328 3:58 38B

rORK LeNGTH <nro

2SB

8l,-..-~.......-.I,.J.,.J..l,W..t.l,,I.,.J.,I~.o;J.,.l,.-~ .....~,.......I

298 328 "B 388 23B

fORK ltHGTH enn>

2SB

8 ["""~,I.,.J.,IW,.L..l,.J.,J..l,.J..l,W:.J..-,,...I;:~,......<;1.r'r~,.......I

238

June 1984 July 1984

Figure S8

67
Scomber japonicus

+ + + +

12S;:::=============~ ,..

7S

S.

2S

29.26. 32. 35. 38.

• ~~'""'""T~,......l,.W.J...l.J.,J,.J.,J.,.I,,l;l.Jly-.J:;J,..c;""'i'~~

23.

fORK LeNGTH eM">

September 1984

Figure 59

68
Recurrent Group Analysis

Inshore Midshelf Midshelf Outer shelf

juveniles temperate southern temperate

Sebastes spp. £Oligo - - - - - - - - - - - - - - - - - - Icichthys

0.30 opalescens lockingtoni

Anarrhichthys

ocellatus Clupea Trachurus Anoplopoma

I

harengus symmetricus fimbria

pallasi -

0.50 Scomber Mola mola

I Engraulis japonicus

o.L mordax

Hexagrammos I

decagrammus Merluccius 0.50 I productus [

Trachipterus 0.33 -

Squalus Peprilus altivelis

acanthias - 1 simillimus

I Cololabis

0.40 saira

0.40 I

Alosa

sapidissima

'-- Sebastes

melanops

Hypomesus _ J

pretiosus

Figure 60

Species associations of marine nekton off Washington and Oregon, 1979-84, according to Fager's recurrent group analysis (affinity level

= 0.30; sample size = 843). Species within squares show positive affinities to all others within the square. Intergroup affinities are shown

with connecting lines between squares. Intergroup affinities above 0.25 are given, and those below 0.25 are shown as dashed lines.

69

Appendix Table I-Station and environmental data, June 1979. Appendix Table 2-Station and environmental data, June 1980.

Off- Time Salin- Off- Time SalinDate Lat. Long. shore Depth at Temp. ity Date Lat. Long. shore Depth at Temp. ity

Set (June) (N) (W) (Ian) (m) stan (0C) r:/oo) Set (June) (N) (W) (Ian) (m) stan (0C) r:/oo)

I 18 46°13' 124°10' 10.2 40 0630 13.5 28.2 I 20 46°16' 124°35' 37.0 463 1410 14.9 30.2

2 18 46°13' 124°16' 16.8 77 0855 13.5 9 21 46°30' 124°44' 50.0 732 1230 15.1 30.5

3 18 46°13' 124°21' 22.4 102 1014 13.5 22.9 10 21 46°20' 124°39' 42.6 183 1446 15.2 29.8

4 18 46°13' 124°27' 29.8 117 1125 14.0 24.4 II 21 46°18' 124°27' 31.5 183 1539 15.4 29.7

7 19 46°20' 124°10' 6.7 26 1310 15.0 28.8 12 21 46°19' 124°21' 23.1 110 1636 15.2 30.4

8 19 46°20' 124°13' 9.6 38 1417 15.0 28.5 13 21 46°18' 124°21' 21.3 110 1713 14.6 30.7

9 19 46°20' 124°17' 15.2 73 1605 15.0 22.3 14 21 46°18' 124°20' 20.3 110 1753 15.2 9.9

10 19 46°18' 124°15' 11.5 66 1730 15.0 16.2 15 21 46°19' 124°18' 17.6 82 1857 15.1 8.2

11 19 46°13' 124°13' 14.8 68 1915 16.0 18.8 16 21 46°19' 124°12' 10.1 37 2002 14.9 10.9

12 20 46°12' 124°15' 16.4 77 0752 14.5 24.8 17 22 46°11' 124°07' 9.8 37 0803 13.3 24.4

16 21 46°10' 124°04' 6.4 22 0724 13.3 27.8 18 22 46°1 I' 124°13' 18.5 73 0911 14.0 16.2

17 21 46'10' 124'08 10.4 40 0843 13.5 27.1 19 22 46° 10' 124°20' 27.8 110 1027 14.4 18.8

18 21 46°10' 124°12' 15.2 73 0957 13.8 24.5 20 22 46°09' 124°28' 37.0 132 1140 15.2 29.9

19 21 46'10' 124°14' 17.1 75 1107 12.9 26.5 24 22 46° 10' 124°14' 21.2 73 1620

20 21 46'10' 124'19' 22.2 99 1219 13.9 23.2 25 23 46°19' 124°12' 9.3 38 0755 14.0 17.7

21 21 46° 10' 124°21' 24.3 112 1325 15.1 22.3 26 23 46° 11' 124°08' 10.2 37 0912 14.1 28.1

22 21 46°10' 124°25' 28.8 130 1444 15.2 21.5 27 25 46°10' 124°13' 18.5 88 1032 13.4 29.0

23 22 45°28' 124'02' 1.6 26 1300 13.4 31.7 28 26 45°36' 124°00' 3.1 37 1332 15.3 27.7

24 22 45°28' 124°06' 6.4 75 1417 14.5 31.3 29 26 45°36' 124'06' 12.0 82 1420 15.2 28.7

25 23 44°38' 124'07' 2.7 27 0704 10.2 33.5 30 26 45'28' 124°00' 2.4 33 1551 15.3 26.7

26 23 44°38' 124°11' 6.2 46 0800 10.4 33.3 31 27 44°39' 124°07' 5.5 38 0655 13.8 31.1

27 23 44°38' 124° 13' 8.6 55 0924 11.2 33.3 32 27 44'41' 124°11' 9.3 55 0746 14.6 29.6

28 23 44'38' 124° 18' 14.1 77 1130 11.2 32.7 33 27 44°40' 124°18' 18.5 77 0857 14.9 28.0

29 23 44'38' 124°26' 23.4 73 1300 12.7 32.3 34 27 44'39' 124°26' 27.8 81 1005 14.8 28.7

30 26 44°34' 124°31' 30.6 135 1000 11.8 32.1 36 27 44°39' 124°22' 23.1 88 1200

31 26 44°34' 124°36' 37.4 196 1111 12.6 32.0 37 27 44°39' 124°15' 17.5 73 1310 15.4 28.9

32 26 44°33' 124°32' 31.8 135 1232 12.3 32.1 38 27 44°39' 124°1 I' 9.3 53 1425 15.1 29.9

33 26 44°31' 124°23' 21.1 53 1411 11.2 32.6 39 27 44°40' 124°08' 6.5 68 1514 14.7 31.0

34 26 44'30' 124°18' 14.4 80 1539 11.3 32.2 40 27 44°42' 124°09' 6.5 54 1605 14.3 31.2

35 26 44°30' 124°1 I' 6.2 55 1656 11.2 33.6 41 28 44°33' 124°08' 3.7 37 0730 12.3 32.2

36 26 44°30' 124°07' 2.2 31 1825 11.2 33.6 42 28 44'30' 124°11' 9.3 55 0837 12.7 32.2

37 27 44'37' 124°08' 3.0 35 0602 9.4 33.7 43 28 44°32' 124°18' 16.7 79 0943 13.1 31.9

38 27 44°37' 124°09' 3.2 37 0807 9.4 44 28 44°33' 124°25' 25.9 44 1058 14.0 30.8

39 27 44°37' 124°12' 8.2 55 0921 10.7 33.6

42 27 44°37' 124°20' 16.0 79 1432 10.3 33.0

43 28 44°37' 124'12' 8.1 57 0410 10.0 33.6

44 28 44°00' 124°14' 5.8 55 0943 9.2 33.6

45 28 44°03' 124°22' 14.2 92 1103 10.5 33.4

46 28 44'03' 124°27' 20.5 112 1310 10.5 33.4

47 29 43°25' 124°22' 4.8 27 0607 10.1 33.5

48 29 43°25' 124°24' 6.4 73 0717 11.0 33.2

49 29 43°25' 124'27' 10.4 102 0820 11.1 33.2

50 29 43°26' 124°34' 19.2 141 0905 12.6 32.9

51 29 43°26' 124°38' 24.0 249 1101 12.5 32.8

52 29 43°27' 124'46' 32.6 520 1220 13.2 32.4

53 29 43°17' 124'38' 16.0 128 1420 12.0 32.7

54 29 43° 17' 124°31' 8.1 92 1538 10.5 33.6

55 29 43°17' 124°27' 3.2 53 1647 10.6 33.5

56 29 43°17' 124°26' 1.9 29 1739 10.5 33.5

71
Appendix Table 3-Station and environmental data, summer 1981.

Off- Time Salin- Chloro- Off- Time Salin- ChloroLal. Long. shore Depth at Temp. ity phyll a Lal. Long. shore Depth at Temp. ity phyll a

Set Date (N) (W) (kIn) (m) start ('C) ~/oo) (J;g/L) Set Date (N) (W) (kIn) (m) start ('C) ~/oo) (J;g/L)

May June

I 16 44'38' 124'07' 3.9 40 0640 10.7 32.7 3.6 74 9 46'35' 124'34' 38.3 201 2102 13.6 31.4 0.2

2 16 44'40' 124'09' 7.0 48 1020 11.1 32.5 - 75 10 46'20' 124'33' 37.0 134 0640 13.4 31.1 0.3

3 16 44'38' 124'13' 12.8 67 1229 12.1 29.1 3.9 76 10 46'20' 124'25' 28.1 126 0821 14.2 24.9 2.0

4 16 44'38' 124'18' 28.3 79 1425 12.6 26.9 1.1 77 10 46'20' 124'18' 18.5 44 1102 14.8 20.1 3.2

5 16 44'38' 124'25' 28.3 67 1703 12.8 27.2 0.6 78 10 46'20' 124'14' 13.3 51 1329 15.1 12.0 3.7

6 16 44'38' 124'18' 18.7 77 1854 12.4 27.5 2.3 79 10 46'20' 124'12' 10.6 37 1452 15.2 14.7 4.1

7 16 44'38' 124'18' 18.9 77 2053 12.9 26.3 - 80 10 46' 10' 124'07' 9.3 37 1718 15.1 23.2 1.4

8 16 44'37' 124'18' 18.5 79 2342 12.6 26.1 - 81 10 46' 10' 124'09' 13.3 49 1842 15.2 24.9 1.4

9 17 44'39' 124'18' 19.1 77 0425 12.2 27.3 - 82 10 46' 10' 124'13' 18.5 73 2052 15.0 26.7 0.6

10 17 44'38' 124' 18' 18.9 75 0702 12.3 - 0.6 83 II 46' 10' 124'20' 28.1 110 0638 14.8 25.2 0.3

II 17 44'38' 124'18' 18.5 75 0904 12.4 27.3 0.7 84 II 46'10' 124'27' 37.0 128 0917 14.8 24.6 0.5

12 18 44'25' 124'08' 4.1 40 1157 12.6 - 2.1 85 II 46' 10' 124'34' 46.3 152 1112 14.1 31.0 0.4

13 18 44'25' 124'1 I' 8.0 49 1307 13.0 27.6 1.0 86 II 46'10' 124'34' 46.3 154 1159 14.3 29.6 - 15 18 44'25' 124'15' 13.1 62 1718 12.9 27.9 - 87 II 46'10' 124'42' 56.2 366 1402 14.4 31.6 0.2

16 18 44'25' 124'19' 18.5 68 1834 12.1 29.7 5.9 88 II 46'10' 124'19' 26.8 104 1710 15.3 25.7 0.3

17 18 44'25' 124'26' 28.0 79 2051 12.7 - 0.7 89 II 46'10' 124'20' 15.0 27 1929 15.1 26.1 0.3

18 18 44'25' 124'32' 35.7 97 2305 12.3 29.8 - 90 II 46' 10' 124'21' 8.8 61 2030 15.0 26.0 - 19 19 44'51' 124'05' 3.1 37 0745 12.3 28.2 3.3 91 II 46' II' 124'20' 27.4 60 2322 15.1 25.5 - 20 19 44'51' 124'08' 7.0 60 0904 12.7 27.4 0.8 92 12 45'55' 124'34' 46.3 170 0846 13.8 31.3 0.7

21 19 44'52' 124'13' 13.0 99 1055 13.2 28.8 0.3 93 12 45'55' 124'27' 37.0 146 1014 14.2 28.1 0.8

22 19 44'52' 124'17' 18.3 124 1258 13.6 27.7 0.2 94 12 45'55' 124'20' 28.5 135 1205 15.0 24.9 0.7

23 19 44'52' 124'24' 27.8 152 1430 12.9 27.9 0.2 95 12 45'55' 124' 13' 18.9 103 1400 15.2 25.5 0.5

25 19 44'51' 124'20' 22.8 135 1707 13.4 28.5 0.2 96 12 45'55' 124'08' 12.6 81 1605 14.8 27.3 0.7

26 19 44'52' 124' 15' 15.5 112 1928 13.5 27.6 - 97 12 45'55' 124'04' 7.2 66 1719 15.2 27.4 0.5

27 20 44'55' 124'01' 3.5 37 0621 13.3 27.5 1.3 98 12 45'55' 124'00' 3.2 33 2010 14.4 28.7 0.8

28 20 45'55' 124'04' 7.4 66 0742 13.5 25.6 1.9 99 13 44'51' 124'06' 3.7 46 1213 14.9 28.3 1.3

29 20 45'55' 124'08' 13.3 81 0919 13.3 17.0 1.8 100 13 44'51' 124'08' 7.0 62 1426 14.8 29.5 0.2

30 20 45'55' 124'13' 18.5 104 1105 13.2 26.2 2.1 101 13 44'51' 124'13' 13.0 99 1632 14.7 29.7 0.4

31 20 45'55' 124'20' 27.8 135 1505 13.5 26.5 1.9 102 13 44'52' 124'17' 18.5 124 1835 14.8 29.5 0.4

32 20 45'55' 124'27' 37.2 146 1718 13.4 27.6 1.4 103 13 44'52' 124'24' 27.8 150 2108 14.7 29.7 0.3

33 20 45'55' 124'34' 46.1 165 1933 12.9 - 0.2 104 14 44'40' 124'32' 37.6 135 0648 14.4 29.2 - 35 21 46'10' 124'28' 38.3 130 0550 12.1 29.7 1.2 105 14 44'40' 124'39' 45.5 278 0821 14.7 29.3 0.1

36 21 46' 10' 124'34' 46.6 152 08.\5 12.4 29.2 0.1 106 14 44'40' 124'24' 27.8 90 1021 14.9 29.2 0.2

37 21 46' 10' 124'34' 46.6 152 0854 12.4 29.2 0.1 107 14 44'38' 124'18' 17.9 75 1215 15.7 28.4 0.4

38 21 46' 10' 124'41' 54.0 194 1105 12.8 30.7 0.2 108 14 44'38' 124'13' 13.0 64 1352 15.4 28.7 0.3

39 21 46'10' 124'29' 38.7 130 1250 12.4 - 2.1 109 14 44'38' 124'09' 6.9 48 1530 15.3 - 1.1

40 21 46'10' 124'20' 28.1 110 1431 13.4 22.9 3.0 110 14 44'39' 124'06' 3.7 40 1722 15.3 29.2 1.4

41 21 46'10' 124'13' 18.3 73 1630 12.9 22.9 1.7 III 15 44'25' 124'08' 3.9 38 0738 13.9 30.0 3.9

42 21 46' 10' 124'08' 12.8 48 1747 13.2 24.3 1.6 112 15 44'25' 124'10' 8.1 48 0912 14.6 29.9 1.0

43 21 46'10' 124'07' 11.3 38 1850 13.4 24.3 1.3 113 15 44'25' 124'15' 13.0 64 1110 15.7 29.4 0.3

44 22 46'20' 124'12' 10.4 37 1119 13.7 - 2.4 114 15 44'25' 124'19' 18.5 68 1226 15.9 29.2 - 45 22 46'20' 124'14' 13.0 51 1242 13.8 15.2 2.9 115 15 44'25' 124'26' 27.8 77 1431 15.9 29.4 1.1

47 22 46'20' 124'25' 27.8 124 1814 12.9 29.3 1.0 116 15 44'25' 124'33' 37.2 101 1622 15.6 29.5 0.3

48 22 46'20' 124'32' 37.0 134 2018 13.0 15.2 - 117 15 44'24' 124'33' 37.2 99 1707 15.6 29.5 0.2

49 22 46'20' 124'33' 37.0 134 2103 13.0 30.8 0.1 118 16 44'1 I' 124'51' 36.2 293 0550 15.4 30.9 - 50 22 46'20' 124'40' 47.7 915 2315 12.9 31.5 0.1 119 16 43'1 I' 124'45' 27.8 247 0740 15.4 30.0 - 51 23 46'20' 124'40' 46.6 915 0019 12.8 31.4 - 120 16 43' II' 124'38' 18.5 123 1010 14.8 30.2 0.2

52 23 46'35' 124'41' 46.6 640 0620 12.8 30.8 - 121 16 43' II' 124'33' 13.0 84 1301 14.8 30.1 -

53 23 46'35' 124'33' 37.0 320 0755 12.7 30.9 0.2 122 16 43' II' 124'30' 6.9 60 1441 12.8 31.4 6.0

54 23 46'35' 124'26' 28.0 95 0932 12.9 29.5 0.9 123 16 43'1 I' 124'27' 3.7 40 1640 12.8 31.7 4.2

55 23 46'35' 124'18' 18.1 119 1131 12.8 24.9 3.4 124 16 43'22' 124'24' 3.7 40 1954 13.6 30.7 10.8

56 23 46'35' 124'14' 12.6 46 1326 13.9 22.0 3.3 125 16 43'22' 124'28' 10.7 93 2128 14.1 30.4 1.5

57 23 46'35' 124'11' 9.6 37 1508 14.2 22.6 - 126 17 43'31' 124'16' 2.6 38 0635 14.4 30.0 1.2

58 24 45'55' 124'13' 18.7 104 1016 12.8 29.2 0.5 127 17 43'32' 124'16' 1.8 22 0745 13.2 30.4 3.1

59 24 45'55' 124'08' 12.9 81 1140 12.8 29.1 0.7 128 17 43'3 I' 124'16' 1.8 24 0835 13.2 30.4 3.1

61 24 45'55' 124'04' 7.0 64 1504 13.4 29.1 0.8 129 17 43'32' 124'16' 1.8 22 1034 13.3 30.7 3.8

62 24 45'55' 124'01' 3.5 38 1619 13.6 27.5 1.0 130 17 43'32' 124'25' 13.3 110 1252 13.4 31.0 1.0

63 24 45'55' 124'20' 28.0 135 1865 12.7 30.7 0.2 131 17 44'01' 124'1 I' 3.5 46 1840 14.2 29.8 4.0

64 24 46'05' 124'10' 18.9 79 2232 13.2 27.8 1.4 132 17 44'01' 124'14' 7.4 62 2001 14.9 30.3 1.1

65 25 46'09' 124'20' 28.0 110 0047 13.7 23.4 1.5 133 18 44'38' 124'07' 4.3 40 0501 14.6 29.8 3.0

66 25 46'10' 124'12' 18.5 70 0325 13.3 25.3 2.4 134 18 44'38' 124'10' 7.4 48 0646 14.8 30.1 3.3

67 25 46'10' 124'07' 11.1 38 0542 13.3 25.5 - 135 18 44'38' 124'12' 10.4 57 0917 14.6 29.2 1.3

68 25 46'10' 124'08' 12.6 0 0643 12.9 27.4 - July

June 140 9 46'35' 124'08' 9.6 37 1115 13.9 - - 69 9 46'35' 124'11' 9.6 37 1052 13.8 26.9 5.3 141 9 46'35' 124'12' 15.9 56 1253 13.8 31.5 2.2

70 9 46'35' 124'09' 7.2 26 1255 13.7 28.0 4.9 142 9 46'35' 124'20' 19.4 68 1412 14.7 31.4 0.7

71 9 46'35' 124'14' 12.6 46 1417 13.4 29.4 3.0 143 9 46'35' 124'26' 27.8 101 1655 14.7 31.6 2.8

72 9 46'35' 124'18' 18.3 66 1550 13.2 - 1.0 144 9 46'35' 124'34' 38.5 223 1826 14.7 31.3 0.3

73 9 46'35' 124'25' 27.8 95 1804 13.7 1.0 0.4 145 9 46'35' 124'25' 27.6 95 2029 14.2 31.4 1.4

72
Appendix Table 3-Continued.

Off- Time Salin- Chloro- Off- Time Salin- ChloroLal. Long. shore Depth at Temp. ity phyll a Lal. Long. shore Depth at Temp. ity phyll a

Set Date (N) (W) (Ian) (m) start (oC) ~/oo) <!,g/L) Set Date (N) (W) (Ian) (m) start (0C) ~/oo) <!,g/L)

July Aug.

146 9 46°35' 124°20' 20.9 71 2149 13.7 31.8 - 213 10 46°20' 124°19' 18.9 79 1148 15.9 18.8 10.0

147 10 46°20' 124°12' 10.6 37 0750 15.3 11.6 2.0 214 10 46°20' 124°26' 28.5 128 1314 15.1 32.4 2.4

148 10 46°20' 124°20' 12.4 48 0927 14.8 13.8 3.7 215 10 46°20' 124°32' 36.8 135 1430 15.1 32.4 1.1

149 II 46°10' 124°10' 11.1 40 0906 13.7 27.1 13.7 216 11 46°35' 124°40' 46.1 640 0957 141 32.0 12.8

150 11 46° 10' 124°10' 14.4 51 0958 13.4 26.3 12.8 217 11 46°35' 124°32' 36.5 183 1109 13.7 32.3 7.3

151 11 46° 10' 124°09' 11.7 53 1111 14.2 26.0 8.7 218 11 46°35' 124°26' 27.9 97 1214 15.0 32.7 1.4

152 11 46°11' 124°18' 25.0 103 1257 17.8 18.1 2.7 219 11 46°35' 124°19' 18.9 64 1435 14.1 32.7 7.7

153 11 46° 10' 124°13' 18.3 73 1435 16.9 22.2 5.5 220 11 46°36' 124°14' 13.0 48 1614 13.8 32.8 62

154 11 46° 10' 124°12' 18.3 73 1510 14.6 26.3 - 221 11 46°35' 124°11' 9.4 37 1721 13.4 32.7 6.5

156 11 46°12' 124°24' 31.8 130 1908 15.8 20.8 5.4 222 12 46° 10' 124°34' 45.9 150 0709 15.3 32.0 05

157 11 46° 10' 124°35' 47.0 476 2057 14.8 31.1 0.2 223 12 46°11' 124°26' 35.5 130 0856 15.0 32.1 0.7

158 12 46°20' 124°39' 45.7 824 0621 14.7 31.2 0.4 224 12 46°10' 124°19' 27.4 108 1011 15.7 20.7 9.9

159 12 46°20' 124°33' 37.5 135 0758 14.7 16.4 3.3 225 12 46° 10' 124°19' 27.4 108 1048 15.6 22.2 -

160 12 46°21' 124°26' 28.3 117 0919 15.4 13.3 4.0 226 12 46°10' 124°18' 25.5 106 1426 16.3 23.0 5.0

161 12 46°20' 124°18' 18.5 75 1145 16.2 14.6 3.0 227 12 46°10' 124°13' 18.1 73 1536 16.5 22.2 13.1

162 12 46°20' 124°18' 17.8 75 1233 16.0 15.0 3.8 228 12 46° 10' 124°06' 10.7 37 1813 14.7 24.8 7.4

163 12 46°20' 124°18' 17.6 75 1317 16.4 14.9 2.9 229 13 45°57' 124°35' 46.3 146 0700 15.6 32.4 0.7

164 12 46°20' 124°14' 13.3 55 1459 16.4 16.5 3.6 230 13 45°55' 124°27' 37.0 145 0826 153 28.9 4.9

166 12 46°19' 124°14' 13.1 53 1601 16.7 16.4 - 231 13 45°55' 124°20' 27.2 135 1019 15.4 25.1 8.0

167 12 46°20' 124°12' 10.6 37 1709 17.3 15.2 2.7 232 13 45°55' 124°20' 27.4 137 1217 15.6 25.7 6.0

168 12 46°20' 124°18' 18.5 81 2134 16.4 14.0 - 233 13 45°55' 124°13' 18.7 104 1329 14.5 31.0 2.7

169 12 46°20' 124° 18' 18.5 81 2346 15.0 17.1 - 234 13 45°56' 124°08' 12.0 82 1543 14.2 31.8 2.4

171 13 46°20' 124°19' 18.7 81 0329 15.1 17.6 - 235 13 45°56' 124°08' 12.9 82 1644 14.2 31.9 3.1

172 13 46°20' 124°18' 18.7 77 0525 15.0 18.8 5.1 236 13 45°55' 124°04' 7.8 70 1753 14.6 32.1 1.3

173 13 46°20' 124°19' 18.7 81 0612 15.0 17.4 - 237 13 45°55' 124°02' 5.2 55 2000 14.4 32.1 0.9

174 13 46°20' 124°18' 18.5 79 0659 15.0 18.4 - 238 14 45°20' 124°20' 28.3 183 0723 15.5 28.5 3.7

175 13 45°55' 124°01' 2.6 37 1525 14.5 28.6 2.7 239 14 45°20' 124° 13' 18.9 152 0850 14.5 31.7 0.7

176 13 45°55' 124°04' 7.8 70 1633 15.2 28.8 4.4 240 14 45°20' 124°08' 12.8 108 1026 14.5 31.8 1.1

177 13 45°55' 124°08' 13.0 82 1725 16.3 24.5 3.2 241 14 45°20' 124°04' 7.2 73 1118 14.3 31.9 2.6

178 13 45°55' 124°08' 11.7 82 1819 15.4 22.9 3.1 242 14 45°19' 124°01' 4.1 44 1206 13.5 32.3 11.5

179 13 45°55' 124°08' 13.0 84 1924 16.3 22.7 3.6 243 15 44°38' 124°06' 3.3 37 1117 13.4 33.0 1.3

180 13 45°55' 124°08' 13.0 84 2132 16.4 19.9 - 244 15 44°38' 124°07' 4.6 46 1216 13.3 33.1 1.0

181 14 45°55' 124°08' 13.0 82 0517 15.1 27.0 2.9 245 15 44°38' 124°14' 13.3 66 1406 13.0 33.1 6.7

182 14 45°55' 124°13' 18.1 104 0625 14.9 23.3 5.0 246 15 44°38' 124° 18' 19.2 81 1506 12.5 33.0 10.1

183 14 45°55' 124°20' 28.5 137 0909 15.8 17.4 2.9 247 15 44°38' 124°25' 27.6 66 1641 13.9 32.3 3.3

184 14 45°55' 124°25' 34.8 146 1026 16.0 19.9 2.7 248 15 44°38' 124°32' 36.5 132 1843 15.0 31.8 1.0

185 15 45°55' 124°34' 46.4 168 1146 15.4 31.4 0.3 249 15 44°38' 124°39' 46.6 238 1952 15.0 31.6 0.7

186 17 44°38' 124°09' 8.9 46 0718 9.1 33.4 1.3 250 15 44°38' 124°14' 13.1 68 2252 12.7 34.9 -

187 17 44°38' 124°14' 12.8 64 0812 11.3 32.5 9.5 251 16 44°38' 124°14' 12.8 66 0143 12.1 32.9 -

188 17 44°38' 124°18' 18.9 77 0912 11.6 32.3 7.5 252 16 44°39' 124°13' 12.8 66 0430 12.2 32.8 -

189 17 44°38' 124°18' 18.9 77 0939 11.9 32.3 - 253 16 44°38' 124°14' 13.3 64 0803 12.1 32.8 6.6

190 17 44°39' 124°20' 20.9 77 1052 12.5 32.0 4.0 254 16 44°52' 124°04' 2.4 37 1429 13.2 33.1 4.5

191 17 44°38' 124°20' 21.6 73 1136 12.2 31.9 3.5 255 16 44°52' 124°09' 8.0 64 1540 12.9 33.0 1.9

192 17 44°38' 124°25' 28.1 73 1513 14.0 31.7 3.2 257 16 44°51' 124° 13' 13.3 103 1716 12.5 33.0 3.4

193 17 44°38' 124°25' 28.3 71 1551 14.2 31.8 - 258 17 44°52' 124° 17' 18.5 114 0621 12.6 32.9 5.7

194 17 44°38' 124°32' 37.6 135 1726 14.2 31.1 3.7 259 17 44°52' 124°24' 29.1 148 0732 14.5 31.4 0.6

195 17 44°38' 124°38' 45.5 258 1840 14.3 29.1 1.1 260 17 44°25' 124°33' 36.8 103 1108 14.6 32.4 0.5

196 18 44°52' 124°31' 37.4 209 0800 15.3 28.5 - 261 17 44°25' 124°26' 28.1 81 1217 14.0 32.4 0.6

197 18 44°52' 124°24' 27.8 154 0906 15.0 30.0 0.6 262 17 44°25' 124°20' 19.1 70 1353 13.2 32.6 2.8

198 18 44°51' 124°16' 18.0 126 1016 15.1 31.0 0.5 263 17 44°25' 124°15' 13.0 66 1530 13.3 33.4 0.7

199 18 44°51' 124°13' 13.3 104 1210 14.4 30.9 1.0 264 17 44°25' 124°10' 7.2 49 1659 13.1 33.4 2.1

200 18 44°51' 124°13' 13.9 103 1245 14.6 33.2 0.9 265 17 44°25' 124°09' 4.6 37 1814 12.5 33.4 5.3

201 18 44°51' 124°06' 3.7 37 1357 12.4 32.1 2.5 266 18 43°11' 124°45' 28.1 311 0926 15.0 32.4 0.6

202 18 44°52' 124°09' 9.7 66 1459 13.9 32.1 5.7 267 18 43°11' 124°38' 19.2 128 1034 14.1 32.7 0.5

203 18 44°38' 124°07' 3.7 37 1702 11.7 33.3 3.0 268 18 43°12' 124°33' 12.4 81 1133 13.3 32.8 1.8

204 18 44°25' 124°08' 4.3 42 2003 12.9 32.8 5.6 169 18 43°12' 124°28' 7.2 60 1318 12.6 33.0 1.9

205 18 44°24' 124°11' 7.1 49 2052 10.6 33.1 6.4 270 18 43°12' 124°27' 4.1 46 1359 11.7 33.4 0.4

206 18 44°25' 124°15' 13.5 64 2203 12.9 32.3 - 271 18 43°21' 124°29' 10.9 95 1557 14.3 32.7 2.5

207 18 44°25' 124°19' 18.5 71 2300 12.6 32.2 - 272 18 43°26' 124°19' 2.8 44 1736 12.7 32.9 1.3

208 19 44°25' 124°26' 27.2 77 0025 13.2 31.9 - 273 19 44°01' 124°34' 34.4 143 0704 14.5 32.3 0.4

209 19 44°25' 124°32' 35.9 97 0140 13.6 31.7 - 274 19 44°01' 124°18' 12.6 86 0915 13.9 32.7 0.7

Aug. 275 19 44°01' 124°13' 6.7 58 1005 13.5 32.6 1.1

211 10 46°20' 124°12' 9.9 37 0851 15.0 19.1 - 276 19 44°01' 124°13' 7.0 58 1037 13.5 32.6 1.0

212 10 46°20' 124°14' 13.3 53 1006 15.7 17.6 - 277 19 44°01' 124°10' 2.6 39 1118 13.1 32.8 1.0

73
Appendix Table 4-Station and environmental data, summer 1982.

Off- Time Salin- Chloro- Off- Time Salin- ChlaroLat. Long. shore Depth at Temp. ity phyll a Lat. Long. shore Depth at Temp. ity phyll a

Set Date (N) (W) (kIn) (m) start (0C) ('/00) v.g/L) Set Date (N) (W) (kIn) (m) start (0C) ('/00) v.g/L)

May June

I 19 48°21" 124°50' 9.8 62 1123 10.8 32.2 5.5 78 7 47°00' 124°25' 18.5 57 2002 12.3 31.3 - 2 19 48°20' 124°55' 18.3 318 1256 10.4 31.7 3.9 79 8 46°41' 124°18' 17.4 62 0646 12.1 28.2 -

3 19 48°20' 125°02' 27.2 218 1426 11.5 31.4 14.6 80 8 46°41" 124°21" 21.1 71 0851 12.4 27.0 4.8

4 19 48°20' 125°10' 37.2 143 1600 12.2 30.9 0.8 81 8 46°41" 124°29' 31.8 97 1045 11.6 30.9 6.5

5 19 48°20' 125°17' 45.7 137 1734 11.8 30.8 0.8 82 10 46°40' 124°19' 19.8 70 0723 11.9 31.4 2.3

6 19 48°20' 124°50' 10.5 59 2025 9.5 32.7 2.8 83 10 46°40' 124°26' 27.6 88 0902 11.6 32.2 3.0

7 20 47°59' 124°49' 11.5 55 0837 11.0 31.0 0.9 84 10 46°41' 124°32' 35.3 119 1041 11.7 32.1 1.5

8 20 48°00' 124°56' 19.2 93 1320 11.7 30.7 0.4 85 10 46°30' 124°32' 37.2 560 1243 11.7 32.1 0.2

9 20 48°00' 125°03' 36.8 121 1505 12.0 30.8 0.3 86 10 46°30' 124°25' 27.6 99 1430 12.3 31.6 1.5

10 20 48°00' 125° II" 37.4 154 1839 11.9 30.9 0.3 87 10 46°30' 124°18' 18.5 68 1616 11.8 31.7 8.1

13 21 47°40' 124°41' 20.7 59 0921 10.0 31.6 9.7 88 10 46°30' 124°18' 18.1 66 1705 11.7 - - 14 21 47°40' 124°46' 27.9 80 1113 11.2 31.2 4.3 89 II 46°23' 124°16' 15.4 60 0730 11.6 31.4 5.8

15 21 47°40' 124°53' 37.6 115 1335 12.1 30.9 0.3 90 II 46°23' 124°18' 19.1 71 0940 12.5 31.6 3.6

16 21 47°20' 124°47' 36.8 329 1700 11.7 30.9 0.6 91 11 46°23' 124°27' 30.0 110 1128 12.4 32.0 l.l

17 21 47°20' 124°39' 27.0 93 1849 11.4 31.3 0.5 92 II 46°23' 124°33' 37.4 139 1259 12.0 31.8 0.4

18 21 47°20' 124°32' 18.3 57 2024 10.1 31.9 21.3 93 II 46° 10' 124°10' 14.4 57 1609 10.8 29.0 2.4

19 22 47"00' 124°25' 18.7 57 1305 11.6 32.1 5.8 95 12 46°11" 124°09' 18.0 84 0840 12.4 26.5 4.2

20 22 47°00' 124°32' 27.4 79 1646 11.7 32.2 2.9 96 12 46°00' 124° II' 19.2 88 1000 12.4 26.3 -

21 22 47°00' 124°25' 18.3 55 2016 11.4 32.1 - 97 12 46°01" 124°14' 23.7 95 1113 12.6 26.5 3.2

22 23 46°40' 124°18' 18.0 62 1023 10.5 31.6 1.5 98 13 45°51' 124°04' 7.4 68 0959 12.6 26.1 2.5

23 23 46°40' 124°18' 18.0 62 1137 10.5 31.6 5.9 100 13 45°50' 124°21' 30.3 137 1521 13.3 30.0 0.9

25 23 46°40' 124°26' 27.9 90 1406 11.4 32.3 12.1 101 13 45°50' 124°15' 22.4 124 1647 13.0 26.6 1.8

26 23 46°40' 124°32' 37.0 124 1715 11.5 31.5 10.1 102 14 45°40' 124°02' 6.9 59 1945 11.5 30.1 11.0

27 23 46°40' 124°16' 15.4 51 2000 11.2 31.8 2.0 103 14 45°40' 124°03' 9.1 68 0736 10.9 30.9 8.7

28 24 46°30' 124°18' 17.8 62 0702 10.8 31.5 7.4 104 14 45°41" 124°10' 18.3 108 1040 12.1 28.9 2.0

29 24 46°21" 124°14' 13.0 49 0903 11.4 31.1 2.9 106 16 45°20' 124°05' 9.3 146 0621 8.8 33.0 0.6

31 24 46°20' 124°18' 18.1 150 1408 11.8 31.0 4.6 107 16 45°20' 124°12' 16.1 148 0818 11.1 31.8 l.l

33 27 46°00' 124°04' 10.6 57 1235 10.4 31.2 - 108 16 45°20' 124°13' 16.8 15 1037 11.2 - - 34 27 46°00' 124°10' 18.7 86 1413 10.4 30.0 5.4 109 17 45°00' 124°05' 6.5 62 1455 9.2 33.0 0.6

35 27 46°00' 124°10' 18.5 88 1555 11.0 31.2 5.4 110 17 45°00' 124°15' 19.1 152 1723 10.6 32.1 1.1

36 27 45°60' 124°17' 27.2 119 1816 11.4 32.2 8.2 III 18 44°41" 124°25' 26.5 93 0949 9.7 32.5 0.5

37 27 46°00' 124°25' 37.9 143 2002 10.6 32.2 - 112 18 44°40' 124°32' 37.0 134 1130 11.8 32.1 1.0

38 27 46°00' 124°31' 46.3 152 2131 10.3 31.9 8.9 113 18 44°40' 124°39' 46.6 285 1316 13.4 31.9 1.7

39 28 46°09' 124°19' 27.4 106 1212 12.3 17.6 3.4 114 18 44°41" 124°46' 54.2 95 1512 13.8 31.5 1.2

41 28 46°08' 124°29' 41.3 134 1736 10.8 32.0 - 115 19 44°39' 124°18' 18.7 77 1856 9.2 33.0 0.4

42 28 46°10' 124°34' 45.5 155 1920 11.2 31.8 4.9 116 19 44°40' 124°11" 9.4 60 2033 8.9 33.8 0.4

44 30 45°41" 124°04' 10.0 71 1005 9.3 33.1 8.5 117 19 44°20' 124°13' 8.7 60 1427 11.3 33.5 17.1

45 30 45°41" 124°10' 18.5 108 1121 11.4 31.5 6.5 118 19 44°20' 124°20' 18.1 77 1600 11.4 33.5 10.7

46 30 45°40' 124°18' 28.1 141 1305 12.0 29.2 6.7 119 19 44°20' 124°28' 28.7 93 1750 9.8 33.2 3.4

47 30 45°40' 124°25' 36.8 166 1506 12.2 27.4 5.0 120 19 44°20' 124°34' 36.8 95 1920 9.6 32.8 1.4

48 30 45°20' 124°06' 9.6 88 1921 10.6 32.0 16.9 121 20 45°00' 124°06' 6.7 64 0945 10.3 32.8 1.1

49 30 45°20' 124°05' 8.5 88 2021 10.4 32.0 - 122 20 45°00' 124°15' 19.2 148 1310 12.9 30.2 1.7

50 31 45°21" 124°12' 17.2 148 0742 9.8 32.3 7.1 123 20 45°00' 124°22' 28.1 205 1454 13.1 30.6 1.1

51 31 45°20' 124°19' 27.4 192 0952 10.8 31.3 3.1 124 20 45°00' 124°22' 28.1 203 1652 13.1 - - 52 31 45°21" 124°26' 35.5 366 1140 11.3 31.0 2.3 125 21 45°21" 124°26' 35.2 22 0&48 12.7 30.4 1.3

53 31 45°20' 124°34' 46.4 454 1400 11.7 30.8 2.3 126 21 45°21' 124°34' 44.4 59 1133 13.3 30.4 0.9

54 31 45°00' 124°28' 36.5 346 1659 11.8 31.8 0.8 127 21 45°20' 124°26' 36.6 26 1405 13.2 30.0 - 55 31 45°01" 124°22' 27.8 201 1841 11.2 31.6 1.1 128 21 45°20' 124°00' 2.4 11 1850 9.0 33.6 - 56 31 45°00' 124°15' 19.1 146 2109 10.2 31.6 1.4 129 22 45°20' 124°12' 15.2 141 0633 11.4 31.6 3.3

June 130 22 45°21' 124°20' 26.8 190 0849 12.1 31.1 3.1

57 1 45°01" 124°05' 5.9 59 0748 9.7 32.7 21.3 Sept.

58 I 45°00' 124°05' 6.1 60 0852 - - - 131 4 47°20' 124°32' 18.3 57 0810 15.2 30.9 4.3

59 I 45°01' 124°08' 9.4 88 1124 10.6 32.4 15.0 132 4 47°20' 124°39' 27.2 93 1204 15.4 31.5 1.5

60 I 44°40' 124°11" 9.6 60 1425 10.8 32.3 10.1 133 4 47°20' 124°47' 37.4 571 1452 15.1 - 0.5

61 I 44°41" 124°18' 18.0 82 1545 10.8 33.1 7.5 136 5 47°00' 124°25' 18.1 55 0703 14.8 31.5 0.6

62 I 44°41" 124°24' 26.8 90 1711 11.3 31.8 1.9 137 5 47°00' 124°29' 23.5 70 0925 15.0 31.8 0.5

63 1 44°41" 124°31' 36.1 137 1850 11.5 32.5 0.4 138 5 47°00' 124°32' 28.1 77 1041 15.5 32.1 l.l

64 2 44°20' 124°34' 36.6 93 0534 10.4 32.2 4.2 139 5 47°00' 124°32' 27.9 79 1157 15.6 31.9 - 65 2 44°20' 124°26' 26.8 90 0715 9.9 33.2 17.4 140 5 47°00' 124°40' 37.6 106 1452 17.0 31.4 0.3

66 2 44°20' 124°21' 19.2 79 0846 9.3 32.9 12.3 141 5 46°40' 124°17' 16.1 55 1843 16.2 30.0 1.0

67 2 44°20' 124°12' 8.3 59 1041 9.4 33.2 16.4 142 6 46°40' 124°16' 15.4 55 0638 15.8 29.0 5.4

68 2 44°00' 124°15' 8.5 73 1400 9.9 32.9 8.9 143 6 46°40' 124°22' 22.9 77 0835 16.1 - 2.7

69 2 44°00' 124°25' 18.3 110 1550 10.2 33.0 13.3 144 6 46°40' 124°22' 22.2 77 0945 16.1 - - 71 7 47°20' 124°32' 18.5 57 0626 11.3 32.8 0.4 145 6 46°40' 124°26' 27.6 88 1105 16.1 31.1 1.0

72 7 47"20' 124°39' 28.0 95 0815 10.8 32.4 7.9 146 6 46°40' 124°33' 37.4 126 1309 16.5 31.4 0.4

73 7 47°21" 124°46' 35.5 159 1028 11.7 31.8 0.7 147 6 46°20' 124°32' 35.9 137 1730 15.4 25.8 7.9

75 7 47°00' 124°40' 37.2 104 1608 11.5 31.8 0.8 148 6 46°20' 124°25' 27.6 126 2007 15.5 25.4 5.9

76 7 47°00' 124°32' 27.9 79 1736 10.8 37..3 4.6 149 7 46°20' 124° 14' 13.0 53 0643 15.6 24.9 7.7

77 7 47°00' 124°25' 18.5 57 1907 12.3 31.3 - 150 7 46°20' 124°18' 18.5 79 0948 15.8 25.2 9.2

74
Appendix Table 4-Continued.

Off- Time Sa1in- ChloroLat. Long. shore Depth at Temp. ity phyll a

Set Date (N) (W) Oem) (m) start (0C) ~/oo) (j.Ig/L)

Sept.

151 7 46°20' 124°18' 17.0 77 1050

153 7 45°59' 124°24' 35.7 137 1634 17.4 3.0

154 8 45°56' 124°03' 4.4 57 0744 14.1 31.8 4.1

155 8 45°56' 124°07' 9.4 79 1002 15.0 31.5 1.1

156 9 45°51' 124°11' 17.2 104 1046 16.5 1.0 1.6

157 9 45°50' 124°19' 27.9 144 1345 16.6 29.1 1.2

158 9 45°40' 124°35' 37.2 172 1640 16.7 31.2 0.5

159 9 45°39' 124°17' 27.4 143 1810 15.9 31.4 0.9

160 9 45°40' 124°01' 5.7 55 2050 15.2 31.5 1.0

161 11 45°21' 124°02' 3.7 55 1707 13.9 32.0

162 11 45°21' 124°05' 9.1 88 1903 14.4 31.8

163 12 45°21' 124°05' 7.8 84 0717 14.8 31,7 4,1

164 12 45°20' 124°12' 18.1 143 0912 15.8 31.5 0.8

165 12 45°21' 124°08' 10.4 104 1104 14.5 31.9 3.0

166 13 44°40' 124°13' 12.4 68 1705 14.0 32.3 2,0

167 14 44°40' 124°18' 19,1 80 0706 13.4 32.9 2.3

168 14 44°40' 124°25' 27.6 93 0900 13,7 32.1 1.6

169 14 44°20' 124°27' 27.9 93 1217 13.9 32.8 1.0

170 14 44°20' 124°21' 19.4 79 1402 13.5 2,0 2.5

171 14 44°20' 124°13' 8.7 55 1534 13.0 32.9 1.1

172 14 44°19' 124° 11' 5.7 51 1629 13.0 33.1 2.7

173 14 44°19' 124°10' 5,2 51 1735 12.4 2.0 2.8

75
Appendix Table S-Slation and environmental data, summer 1983.

Off- Time Salin- Chlaro- Off- Time Salin- ChloroLa\. Long. shore Depth al Temp. ity phyll a La\. Long. shore Depth at Temp. ity phyll a

Set Date (N) (W) (kIn) (m) start (0C) ~/oo) (j;g/L) Set Date (N) (W) (kIn) (m) start (0C) ~/oo) (j;g/L)

May June

I 16 48°20' 124°44' 5.9 42 1245 10.1 32.2 2.0 73 14 48°20' 125°09' 36.3 119 1242 13.2 3 \.4 1.3

2 16 48°20' 124°46' 9.1 41 1445 10.6 3\.9 2.6 74 14 48°20' 124°54' 18.0 302 1500 12.7 3 \.4 3.2

3 16 48°20' 124°55' 18.7 302 1643 10.9 3\.3 2.9 75 15 48°00' 124°48' 9.3 38 0951 12.9 3\.1 6.1

4 16 48°20' 125°02' 27.6 181 1830 11.1 30.7 8.2 76 15 48°00' 124°56' 18.9 90 1132 13.5 31.3 1.3

5 16 48°20' 125° 10' 36.0 191 2005 11.7 30.2 3.8 77 15 48°00' 125°03' 28.0 123 1300 14.0 3\.1 1.7

6 16 48°21' 125°18' 45.8 112 2218 12.2 30.9 2.1 78 15 47°40' 124°51' 34.5 106 1708 15.4 3\.1 5.2

7 17 48°00' 124°48' 8.9 35 0920 11.2 31.2 I\.I 79 15 47°40' 124°44' 25.0 71 1815 16.0 3\.5 \.4

8 17 48°00' 124°56' 18.5 88 1100 I\.9 30.4 7.0 80 15 47°40' 124°38' 15.9 42 1940 15.5 3\.5 -

9 17 48°00' 125°04' 28.7 123 1236 13.0 31.3 \.0 81 16 47"20' 124°47' 37.8 549 1102 14.2 3\.5 0.2

10 17 48°00' 125°10' 35.8 146 1414 12.5 3 \.4 0.2 82 16 47°20' 124°39' 27.8 92 1215 14.5 3\.6 0.9

11 17 47°40' 124°53' 36.7 115 1744 12.7 3 \.0 0.2 83 16 47°20' 124°32' 18.5 59 1310 14.2 3 \.6 0.3

12 17 47°41' 124°45' 27,1 77 1910 12.4 3 \.0 0.8 84 16 47°20' 124°27' I\.7 33 1410 14.7 28.2 1.7

13 17 47"41' 124°38' 15.8 46 2033 1\.8 3\.0 8.8 85 22 46°21 ' 124°12' 10.7 38 0610 15,9 18.1 \.6

i4 18 47°40' 124°35' 13.5 35 0637 11.9 30.7 2.7 87 23 46°00' 124°01' 7.6 37 0717 15.4 26.5 0.4

15 18 47°20' 124°27' 1\.9 38 1026 12.9 30.2 1.3 88 23 46°01' 124°10' 18.7 84 0917 15.7 25.2 0.3

16 18 47°20' i24°32' 18.0 54 1317 12.9 30.5 \.6 89 23 46°00' 124°17' 27.4 112 1045 15.2 25.3 0.8

17 18 47°20' 124°39' 27.4 91 1510 13.0 3\.0 0.6 90 23 46°00' 124°06' 13.0 68 1240 16.3 25.2 0.3

18 18 47°20' 124°47' 36.9 549 1659 13.1 30.7 0.3 91 23 45°39' 124°25' 37.4 165 1616 16.3 29.7 -

19 18 47°00' 124°19' 11.3 37 2043 12.6 30.9 0.3 92 23 45°40' 124°17' 27.4 141 1727 16.4 29.8 0.1

20 19 47°00' 124°25' 18.7 53 1423 13.3 30.6 0.2 93 23 45°40' 124°10' 18.0 104 1831 16.2 27.2 0.2

21 19 47°00' 124°26' 19.6 55 1728 i3.3 - - 94 23 45°41' 124°03' 8.9 68 1942 16.1 25.3 0.3

22 19 47°00' 124°33' 28.4 79 1900 13.1 30.6 0.2 95 23 45°40' 124°00' 4.3 38 2039 16.0 25.1 0.3

23 20 46°40' 124° 13' 11.3 40 0829 12.8 30.7 0.6 96 24 45°20' 124°19' 27.4 192 1036 15.9 29.9 0.1

24 20 46°40' 124°19' 18.9 66 1021 13.0 30.9 0.3 97 24 45°20' 124°13' 18.9 145 1200 15.5 30.3 0.1

25 20 46°40' 124°26' 27.4 84 1153 12.9 29.7 0.1 98 24 45°21' 124°08' 13.2 110 1327 15,5 29.0 0.3

26 20 46°40' 124°33' 36.7 119 i342 13.4 30.0 0.3 99 24 45°20' 124°06' 9.5 84 1437 15.7 28.8 0.2

28 21 46°21' 124°11' 10.0 35 0902 12.3 30.1 \.6 100 24 45°17' 124°21' 4.4 44 1557 15.7 28.1 0.4

29 21 46°21' 124° 18' 18.5 81 1344 13.2 3\.2 0.3 101 24 45°01' 124°01' 27.2 198 1910 16.0 30.5 0.1

30 2i 46°20' 124°26' 28.5 128 1545 13.7 30.4 0.1 102 24 45°00' 124°15' 18.2 143 2029 16.0 29.7 0.1

31 21 46°21' 124°33' 37.1 137 1718 13.9 29.9 0.1 103 24 45°01' 124°07' 9.1 84 2145 15.8 29.7 0.1

32 21 46°20' 124°25' 27.6 128 1850 13.4 30.6 0.1 104 24 45°01' 124°04' 4.4 42 2245 15.3 29.4 0.2

33 22 46°00' 124°01' 6.5 37 0900 12.4 31.1 0.5 105 25 44°40' 124°07' 5.0 45 1046 14.9 3\.1 \.2

34 22 46°00' 124°06' 13.5 73 1036 13.6 29.1 0.5 106 25 44°40' 124°18' 18.7 79 1221 15.4 30.8 0.3

35 22 46°00' 124°10' 18.7 86 1154 13.9 25.6 \.1 107 26 43°28' 124°19' 3.3 41 0615 12.6 32.2 0.7

36 22 46°00' 124°17' 27.6 117 1317 13.8 24.6 2.0 108 26 43°28' 124°22' 8.9 90 0719 14.5 3\.2 0.5

37 23 45°41' 123°59' 3.9 38 0907 1\.9 3\.8 0.5 109 26 43°28' 124°30' 18.5 117 0841 15.6 3\.2 0.2

38 23 45°41' 124°03' 8.9 68 1024 12.9 31.3 0.6 110 26 43°28' 124°30' 18.5 119 1016 15.9 - -

39 23 45°41' 124°11' 18,5 106 1154 14.1 29.4 0.1 III 26 43°28' 124°36' 27.1 187 1131 16.3 3\.0 0.1

40 23 45°41' 124°18' 28.5 i43 1325 15.3 27.6 0.2 112 26 43°28' 124°44' 37.4 516 1254 16.4 30.1 0.1

41 23 45°24' 124°23' 32.1 421 1606 15.0 26.6 0.7 113 26 43°00' 124°47' 27.1 150 1637 16,2 30.5 0.2

42 23 45°21' 124°19' 26.9 183 1834 14.5 26.9 0.2 114 26 43°00' 124°41' 18.2 135 1748 16.4 30.7 0.3

43 23 45°20' 124°13' 18.7 146 1949 14.6 26.9 0.2 115 26 43°00' 124°34' 8.5 90 1902 14.5 32.0 \.2

45 24 45°19' 124°06' 7.8 78 1012 14.0 28.5 0.1 116 26 43°00' 124°29' 2.6 42 2016 14.1 32.9 1.7

46 24 45°00' 124°04' 4.1 37 1312 13.5 3\.6 \.2 117 27 44°00' 124°11' 4.1 42 0859 14.8 31.7 0.5

47 24 45°00' 124°17' 2\.5 141 1449 13.8 29.3 0.2 118 27 44°00' 124°14' 8.9 71 1009 15.6 3\.6 0.3

48 24 45°00' 124°22' 27.8 201 1619 13.9 29.6 0.1 119 27 44°00' 124°22' 18.3 110 1153 15.8 31.3 0.1

49 24 44°56' 124°09' 9.1 82 1815 13.9 28.3 0.2 120 27 44°20' 124°08' 2.8 42 1531 15.4 3\.9 0.4

50 25 44°40' 124°08' 5.6 42 1028 I \.8 32.6 8.5 Sep\.

51 25 44°40' 124°18' 18.7 81 1315 14.0 28.9 0.2 121 15 48°20' 124°47' 8.9 38 0808 14.1 3\.8 1.3

52 25 44°40' 124°24' 27.4 95 1546 14.1 30.1 0.1 122 15 48°20' 124°51' 13.2 71 0934 14.6 3\.5 0.8

53 26 44°20' 124°09' 3.7 37 0745 I\.I 32.8 8.8 123 15 48°20' 124°54' 18.3 307 1107 14.5 3\.8 \.7

54 26 44°20' 124°13' 9.3 60 0920 1\.9 32.7 5.8 124 15 48°20' 125°02' 27.8 225 1234 14.9 3\.6 1.3

56 27 44°20' 124°20' 18.5 77 0648 I\.4 32.8 0.2 125 15 48°20' 125°10' 37.1 121 1451 15.1 3\.6 -

57 27 44°20' 124°27' 28.5 90 0824 I\.6 32.8 7.0 126 15 48°20' 125°17' 46.3 178 1602 15.2 31.5 1.4

58 27 44°19' 124°27' 27.1 92 0958 11.7 - - 127 15 48°18' 125°25' 55.6 113 1735 14.5 3\.6 3.2

59 27 44°20' 124°33' 36.5 92 1105 I\.9 32.8 8.0 128 16 47°40' 124°36' 14.8 42 0732 15.9 30.8 0.6

60 27 44°20' 124°41' 46,0 82 1240 12.2 32.1 0.3 129 16 47°40' 124°41' 23.2 70 1004 15.8 3\.0 0.7

June 130 16 47°40' 124°50' 32.4 104 1155 15.8 3\.0 \.0

61 9 46°30' 124°22' 23.2 81 1046 12.6 3\.8 5.0 131 16 47"20' 124°32' 18.5 60 1635 16.2 30.6 2.8

62 12 46°29' 124°20' 22.2 73 1102 15.9 29.4 5.5 132 16 47°20' 124°28' 13.3 44 1748 15.6 30.7 \.9

63 12 46°41' 124°12' 9.1 37 1324 15.2 26.8 5.0 133 17 47°00' 124°28' 13.0 42 0821 14.7 30.9 2.1

65 13 46°40' 124°19' 18.5 62 0750 14.7 31.5 \.2 134 17 47°00' 124°29' 23.2 71 1037 15.8 30.8 1.7

66 13 46°40' 124°26' 28.4 82 0935 15.4 3\.4 0.4 135 17 47°00' 124°34' 29.7 88 1146 15.9 30.6 2.4

67 13 46°40' 124°33' 36.9 124 1055 15.9 3\.6 0.3 136 i7 46°40' 124°27' 29.0 90 1451 16.4 30.9 \.0

68 13 47°00' 124°32' 28.1 77 1348 15.5 31.3 0.2 137 17 46°40' 124°17' 16.7 59 1624 i5.8 30.8 3.6

69 13 47°00' 124°24' 17.4 53 1509 15.3 25.5 2.0 138 17 46°40' 124°14' I\.I 40 1834 15.6 3\.1 5.2

70 13 47°00' 124°20' 12.4 48 1945 14.9 26.7 3.1 139 19 46°21' 124°13' 11.6 42 0755 13.3 - 4.4

71 14 48°20' 124°47' 8.5 40 0605 12.9 3\.6 4.4 140 19 46°24' 124°19' 18.9 71 0943 14.4 3\.2 3.6

72 14 48°20' 125°04' 30.2 187 1055 12.9 31.7 2.1 142 19 46°28' 124°27' 29.7 101 1220 15.1 3\.1 2.5

76
Appendix Table S-Continued.

Off- Time Salin- ChloroLat. Long. shore Depth at Temp. ity phyll a

Set Date (N) (W) (km) (m) start (0C) ~/oo) (I-'g/L)

Sept.

143 20 45°59' 124°01' 8.0 44 0846 13.8 30.6 2.5

144 20 46°00' 124°06' 13.0 71 1000 13.8 30.6 7.4

145 20 46°01' 124°1 I' 18.5 88 1111 14.1 30.8 7.4

146 20 46°00' 124°18' 27.8 119 1251 14.0 31.3 6.6

147 20 45°40' 124°10' 18.5 107 1519 14.4 30.4 5.0

148 20 45°40' 124°03' 9.3 70 1710 14.6 30.4 5.2

149 20 45°40' 124°03' 8.9 70 1800 14.7

150 20 45°39' 123°59' 3.7 40 1903 14.0 30.8 6.1

151 21 45°20' 124°20' 28.7 192 1920 14.9 31.2 6.9

152 21 45°20' 124°12' 18.5 145 2034 14.2 31.5 8.8

153 21 45°20' 124°1 I' 13.9 110 2132 13.4 31.4 8.8

154 22 45°20' 124°05' 9.3 82 0648 12.9 31.6 9.4

155 22 45°20' 124°01' 3.7 45 0747 12.9 31.5 7.7

156 22 45°17' 124°01' 3.7 41 0850 12.1 31.9 88

157 22 45°10' 124°01' 3.2 43 1031 11.3 32.3 5.0

158 22 45°00' 124°03' 3.3 41 1227 12.2 32.6 3.3

159 22 45°00' 124°08' 9.3 88 1357 13.7 32.0 4.7

160 22 45°00' 124°15' 18.5 148 1505 14.3 31.8 2.6

161 22 44°49' 124°06' 2.3 41 1713 12.5 32.6 4.7

163 23 44°40' 124°07' 4.3 40 0923 13.0 32.5 5.7

164 23 44°40' 124° II' 9.6 60 1025 12.6 32.5 4.4

165 23 44°40' 124° 18' 18.7 82 1133 14.0 32.2 2.0

166 23 44°20' 124°20' 18.5 77 1426 15.4 32.0 0.4

167 23 44°20' 124°13' 9.3 62 1544 14.6 32.2 2.7

168 23 44°20' 124°34' 3.2 38 1647 13.2 32.6 7.3

169 23 44°20' 124°09' 3.7 38 1737 13.9

170 23 44°30' 124°08' 3.7 44 1922 14.1 32.2 1.9

171 24 44°00' 124°1 I' 3.9 44 0821 12.4 32.8 6.4

172 24 44°00' 124°15' 9.3 73 0912 12.3 33.0 5.8

173 24 44°00' 124°22' 18.5 112 1042 13.6 32.7 0.9

174 24 43°28' 124°19' 3.0 47 1719 13.3 32.8 0.7

77
Appendix Table 6-Station and environmental data, summer 1984.

Off- Time Salin- Chloro- Off- Time Salin- ChloroLat. Long. shore Depth at Temp. ity phyll a Lat. Long. shore Depth at Temp. ity phyll a

Set Date (N) (W) (Icrn) (m) start (0C) ~/oo) (jAg/L) Set Date (N) (W) (Icrn) (m) start (0C) ~/oo) (jAg/L)

June July

I 4 48°20' 124°49' 11.4 91 1755 11.8 32.3 1.2 82 23 44°40' 124°18' 18.1 76 0730 9.4 33.5 18.0

2 5 48°20' 124°55' 18.8 305 0555 9.8 32.3 0.9 84 23 44°40' 124°26' 29.0 100 1611 11.9 32.3 21.7

3 5 48°20' 125°03' 28.4 183 0850 10.8 31.0 1.8 85 23 44°40' 124°32' 37.0 131 1737 11.9 32.2 16.6

4 5 48°20' 125° 10' 37.0 179 1126 12.8 30.3 1.2 86 23 44°40' 124°39' 46.8 274 1908 12.4 31.9 6.1

5 5 48°00' 125°10' 36.6 153 1615 12.7 30.0 0.7 87 24 44°01' 124°36' 37.9 146 0051 12.0 32.8 2.2

6 5 48°00' 125°03' 27.1 117 1750 12.5 29.7 0.6 88 24 44°28' 124°28' 26.8 126 0330 12.0 32.7 4.2

7 5 48°00' 124°55' 17.7 89 1910 12.5 29.7 10.1 89 24 44°00' 124°21' 17.3 106 0458 12.2 32.9 3.1

8 5 48°00' 124°49' 10.1 43 2103 12.1 30.0 1.4 90 24 44°00' 124°15' 9.0 73 0610 11.6 33.2 10.4

9 6 47°40' 124°37' 16.6 43 0813 11.9 30.4 6.3 91 24 44°00' 124°11' 4.0 45 0709 9.9 33.9 12.5

10 6 47°40' 124°39' 19.2 51 1015 12.4 30.4 3.8 92 25 45°00' 124°04' 4.4 49 0748 10.0 33.4 29.4

11 6 47°40' 124°46' 28.1 80 1405 13.3 29.5 0.9 94 25 45°00' 124°08' 9.6 82 1846 13.7 31.2 1.8

12 6 47°40' 124°53' 37.5 113 1545 12.7 30.1 3.3 95 25 45°00' 124°05' 18.6 142 2008 14.1 30.9 1.2

13 6 47°40' 125°01' 47.3 170 1731 12.6 30.1 1.8 96 25 45°00' 124°23' 29.2 215 2138 14.4 31.3 0.6

15 8 47°20' 124°28' 13.1 42 0930 12.3 29.5 3.2 97 26 45°30' 124°02' 5.3 47 0852 13.2 31.4 2.2

16 8 47°20' 124°32' 18.5 58 1200 12.1 29.2 0.8 98 26 45°31' 124°06' 10.9 80 1002 13.7 31.1 1.4

17 8 47°20 124°39' 27.3 89 1300 12.4 30.3 1.9 99 26 45°31' 124°12' 19.0 128 1116 14.0 30.4 1.1

18 8 47°00' 124°40' 37.3 100 1550 12.0 30.2 2.6 100 26 45°30' 124°19' 28.6 164 1228 14.4 30.9 0.7

19 8 47°00' 124°32' 27.7 73 1735 12.1 30.4 2.2 101 26 46°00' 124°24' 36.4 142 1608 13.8 31.3 2.8

20 8 47°00' 124°25' 18.3 54 1920 12.0 30.0 2.0 102 26 46°00' 124°31' 45.8 155 1724 12.2 32.2 1.5

21 8 47°00' 124°19' 11.6 40 2050 12.2 28.1 1.5 103 26 46°00' 124°16' 27.1 III 1918 12.8 31.5 3.7

22 9 46°40' 124°13' 10.3 40 1220 13.3 25.9 2.9 104 26 46°00' 124°10' 18.6 87 2040 13.3 31.2 0.6

23 9 46°40' 124°20' 17.5 64 1345 13.0 27.0 0.8 105 26 46°01' 124°03' 9.8 47 2205 13.3 31.9 1.2

24 9 46°40' 124°26' 27.3 89 1509 13.0 30.5 1.5 106 27 46°30' 124°18' 18.5 65 1211 13.8 27.2 13.7

25 9 46°40' 124°34' 37.3 124 1654 12.8 30.3 2.7 109 27 46°30' 124°33' 37.9 219 1759 14.4 31.9 0.6

26 10 46°20' 124°13' 11.8 40 0915 12.9 14.2 1.6 110 27 46°30' 124°40' 46.8 457 1907 15.5 31.8 0.2

27 10 46°20' 124°34' 38.2 137 1710 13.9 30.1 2.1 III 28 47°00' 124°19' 13.6 47 1437 12.9 32.6 1.2

28 10 46°20' 124°25' 27.5 124 1850 12.9 29.1 1.4 112 28 47"00' 124°27' 21.0 64 1545 13.2 32.7 2.5

29 11 46°20' 124°19' 19.4 82 1350 14.3 17.5 1.6 114 29 47°00' 124°32' 27.7 78 0606 13.3 32.0 2.2

30 II 46°00' 124°01' 7.9 40 1750 13.7 26.9 2.2 115 29 46°59' 124°39' 36.6 102 0812 13.8 31.9 1.5

31 II 46°00' 124°04' 9.9 49 1855 13.4 26.8 1.8 116 29 47°00' 124°47' 46.2 142 1030 14.9 31.8 0.5

32 12 46°00' 124°24' 36.8 139 1030 13.2 19.6 2.3 117 29 47°30' 124°56' 44.4 197 1540 14.2 31.8 0.3

33 12 46°00' 124°17' 27.7 118 1223 13.1 22.9 2.0 118 29 47°30' 124°48' 34.7 126 1707 14.5 31.8 1.9

34 12 46°00' 124°10' 18.1 84 1447 13.3 25.1 2.6 119 29 47°30' 124°41' 25.9 82 1833 13.5 32.2 5.9

35 12 45°40' 124°10' 18.3 104 1740 13.2 27.8 1.2 120 29 47°30' 124°36' 18.8 49 2000 13.2 32.8 1.0

36 12 45°40' 124°03' 9.2 67 1918 11.8 30.5 1.4 121 30 48°00' 124°50' 11.1 47 0807 13.8 32.0 1.4

37 13 45°40' 124°18' 27.9 142 1600 14.1 24.8 1.8 122 30 48°00' 124°56' 18.8 87 0952 14.0 32.1 0.6

38 13 45°40' 124°25' 37.0 170 1722 13.9 22.9 3.4 123 30 48°00' 125°03' 27.9 120 1150 13.5 32.0 1.4

39 13 45°20' 124°06' 9.8 82 2027 12.0 30.6 1.8 124 30 48°00' 125°11' 37.5 155 1325 14.3 32.1 0.8

40 14 45°20' 124°13' 18.3 142 0906 12.9 27.2 1.3 Sept.

41 14 45°20' 124°20' 27.7 184 1026 13.2 27.9 1.0 140 1 48°20' 124°49' 10.7 54 0711 10.1 32.5 1.8

44 16 45°00' 124°04' 4.6 45 0829 9.4 33.1 0.6 141 I 48°20' 124°54' 17.5 290 0916 12.5 31.5 3.0

45 16 45°01' 124°08' 9.8 84 1007 10.4 32.0 1.1 142 I 48°21' 125°00' 22.2 252 1034 13.4 31.8 5.5

46 16 45°00' 124°15' 18.8 144 1131 11.9 30.7 1.0 143 1 48°21' 125°00' 22.2 241 1139 12.4 - -

47 16 45°00' 124°22' 28.4 203 1304 12.9 30.0 1.3 144 I 48°21' 125°10' 37.0 186 1316 14.9 31.8 7.4

48 16 45°00' 124°29' 36.4 345 1440 14.0 27.0 0.9 145 I 48°00' 125° II' 37.1 159 1624 15.2 31.2 2.5

49 18 44°40' 124°07' 4.9 43 0517 8.4 33.7 0.2 146 1 48°00' 125°03' 27.7 122 1753 14.8 31.5 4.7

51 19 44°40' 124°17' 17.9 76 0032 8.9 32.7 0.3 147 1 48°00' 124°56' 18.3 89 1921 14.1 31.2 7.1

52 19 44°40' 124°18' 19.2 87 0154 10.3 31.8 0.3 148 I 48°00' 124°49' 10.7 43 2054 13.2 31.7 7.1

53 19 44°40' 124°31' 36.4 135 0320 11.2 31.1 1.2 149 2 47"40' 124°37' 16.6 43 0823 12.4 32.5 3.3

54 19 44°40' 124°38' 44.7 252 0435 11.6 30.8 1.7 150 2 47°40' 124°43' 24.0 65 0942 12.4 32.3 9.3

55 19 44°20' 124°43' 46.9 100 0755 10.5 31.5 1.1 151 2 47°40' 124°46' 28.4 82 1054 14.5 31.9 3.3

56 19 44°20' 124°34' 35.8 89 1001 10.4 32.0 0.5 152 2 47°20' 124°43' 36.2 549 1403 15.4 31.9 0.8

57 19 44°20' 124°28' 29.0 89 1122 10.9 32.7 0.4 153 2 47°20' 124°39' 27.5 93 1521 15.3 32.1 - 58 19 44°20' 124°20' 17.9 73 1250 11.4 33.3 2.2 154 2 47°20' 124°36' 23.3 73 1630 15.0 32.4 2.7

59 19 44°20' 124°14' 9.0 54 1414 12.1 33.2 4.5 155 2 47"20' 124°29' 14.6 45 1737 15.5 32.1 1.3

60 19 44°20' 124°10' 4.8 43 1513 10.8 33.3 1.8 157 3 47°00' 124°21' 13.6 43 0849 14.0 32.4 - 61 19 44°00' 124°11' 3.7 42 1829 11.3 33.1 5.9 158 3 47°01' 124°27' 20.7 62 1027 14.6 32.4 3.9

62 20 44°00' 124°15' 8.6 73 0549 10.2 32.6 1.8 159 3 47°00' 124°27' 21.0 60 1124 14.6 32.6 -

63 20 44°00' 124°22' 17.9 111 0726 10.8 31.9 1.4 160 3 47°00' 124°32' 27.9 76 1326 15.4 32.2 2.5

64 20 44°00' 124°29' 27.7 128 0848 10.5 32.1 0.7 161 3 47°00' 124°39' 36.8 98 1457 16.3 32.2 0.9

65 20 44°00' 124°35' 36.0 150 1006 10.2 31.9 0.8 162 3 47°00' 124°47' 46.4 140 1614 15.0 31.9 1.3

66 20 43°27' 124°43' 35.1 512 1430 12.8 32.6 2.4 163 3 47°00' 124°32' 27.5 76 1806 16.0 32.2 1.3

67 20 43°28' 124°37' 26.8 208 1540 13.4 31.8 3.4 164 4 46°40' 124°14' 12.0 45 0907 14.2 32.0 2.5

68 20 43°28' 124°31' 19.0 122 1648 13.1 31.9 1.1 165 4 46°40' 124°19' 18.5 65 L043 14.4 32.4 2.3

69 20 43°27' 124°24' 9.8 95 1803 11.0 32.9 3.0 166 4 46°40' 124°18' 17.9 64 1142 14.8 32.2 0.4

70 20 43°28' 124°19' 2.9 43 1913 11.8 33.1 4.7 167 4 46°40' 124°26' 28.3 86 1326 15.0 32.4 1.6

July 168 4 46°40' 124°31' 37.1 122 1502 15.8 31.8 0.5

80 19 44°40' 124°08' 5.7 45 0608 8.3 33.6 7.0 169 4 46°20' 124°13' 11.4 45 1835 15.4 21.2 3.6

81 23 44°40' 124°10' 9.0 58 0557 9.5 33.5 20.9 170 5 46°20' 124°18' 18.1 73 1841 14.9 25.3 2.7

78
Appendix Table 6-Continued.

Off- Time Salin- ChloroLat. Long. shore Depth at Temp. ity phyll a

Set Date (N) (W) (krn) (m) start (0C) ~/oo) (Jlg/L)

Sept.

171 5 46°21' 124°24' 26.0 109 1956 16.5 30.0 0.8

172 6 46°20' 124°25' 27.7 122 0816 14.2 24.1 2.7

173 6 46°20' 124°33' 37.1 137 0931 16.1 30.1 0.7

174 6 46°20' 124°18' 19.6 76 1131 15.6 20.1 3.0

175 10 46°00' 124°03' 9.2 43 0907 14.8 30.8 1.8

176 10 46°00' 124°24' 37.0 144 1143 16.5 31.5 0.3

178 11 45°40' 124°00' 4.8 45 0644 15.5 31.0 0.5

179 11 45°40' 124°04' 9.2 69 0748 15.3 31.3 0.5

180 11 45°40' 124°11' 18.5 104 0927 15.6 30.7 1.1

181 11 45°40' 124°18' 28.4 140 1045 15.4 30.8 1.2

182 II 45°20' 124°13' 17.3 137 1430 16.1 31.8 0.5

183 11 45°20' 124°05' 9.2 80 1600 16.0 31.6 0.5

184 11 45°21' 124°02' 5.3 45 1833 15.9 31.6 0.5

186 12 45°00' 124°08' 9.2 84 1212 15.3 32.3 0.9

187 12 45°00' 124°15' 18.5 139 1343 15.7 31.9 0.5

188 12 45°00' 124°22' 27.7 195 1500 17.2 31.8 0.3

189 12 45°00' 124°04' 4.6 45 1714 15.1 32.9 l.l

190 13 44°20' 124°14' Il.l 43 1040 14.8 32.8 1.5

191 13 44°20' 124°12' 9.2 60 1132 14.8 32.8 1.8

192 13 44°20' 124°20' 18.5 76 1245 15.2 32.5 0.5

193 13 44°20' 124°28' 27.9 91 1352 15.5 32.7 0.5

194 13 44°20' 124°13' 9.4 58 1550 14.9 32.8 1.8

195 13 44°30' 124°10' 6.2 45 1819 14.3 33.0 4.1

196 14 44°40' 124°07' 4.9 43 0807 13.4 33.1 8.5

197 14 44°40' 124°10' 9.2 56 0917 14.4 32.9 2.7

198 14 44°40' 124°18' 18.6 80 1030 14.4 32.9 2.1

199 14 44°40' 124°25' 28.6 93 1141 14.8 32.2 l.l

200 14 44°40' 124°32' 38.1 139 1258 15.6 32.1 0.8

201 15 44°00' 124°28' 26.2 124 1305 15.0 31.9 0.8

202 15 44°00' 124°23' 19.2 111 1417 15.3 32.1 0.7

203 15 44°00' 124°15' 9.6 76 1524 15.3 32.3 0.7

204 15 44°00' 124°11' 4.0 43 1622 15.3 32.7 1.8

205 15 44°00' 124°11' 3.8 45 1655

79
Appendix Table 7-lndividual accounts for all taxa collected in purse seines off the coast of Oregon and Washington, 1979-84. The nine most abundant taxa are

disc'ussed in detail in the text. Capture data on the less common species are listed in the following abbreviated sequence: year and set number, followed by lengths

of specimens taken. For those sets which contained 10 or more individuals of a species, the number collected is followed by a hyphen and the mean length and

range in lengths (in parentheses). Specimens for which no lengths are available are designated by NLA. All lengths given are fork length in mm unless otherwise

specified. The systematic arrangement and common and scientific nomenclature used below follow Roper et aI. (1984) for cephalopods and Robins et aI. (1980) for fIShes.

CEPHALOPODA Prionace glauca (Linnaeus), blue shark

Rajiformes (total lengths)

Teuthoidea (dorsal mantle lengths)

Loliginidae

Loligo opalescens Berry, market squid

See text for catch and length-distribution data.

Onychoteuthidae

Onychoteuthis borealijaponicus Okada, Pacific clubhook squid

80-012; 72

81-050; 242

84-096; 155,164,186,196,198,211 ,NLA,NLA

Octopoda

Octopodidae

Octopus dojleini (Wulker), giant octopus

84-148; I - NLA

80-020; 1370,1570

81-141; 1190

81-195; 1170

81-196; 1170,1660

81-200; 1750

81-209; 1250

81-215; 1170

81-217; 1310

81-223; 1780

81-232; 1750

81-249; 920,1520

81-267; 1520

81-273; 950

82-055; 1740

82-154; 1520

82-165; 1370,1520

83-123; 1980

83-163; 1310

83-164; 1320

83-165; 940

83-166; 1490,1450

83-167; 1540,1930

83-172; 1350

84-100; 1115

84-101; 1040,1880

84-110; 2 - NLA

84-160; 1830

84-167; 1440

84-171; 1525

84-180; 995

AGNATHA

Petromyzontiformes (total lengths)

Petromyzontidae

Lampetra tridentOla (Oairdner), Pacific lamprey

79-045; 552

80-028; 1 - NLA

82-034; 186

82-042; 650

82-060; 605

82-079; 192

82-106; 600

82-117; 565

83-045; 2 - NLA

83-061; 590

83-118; I - NLA

84-039; 596

84-055; 570

84-089; 251

84-143; 215

CHONDRICHTHYES

Squaliformes (total lengths)

Squalidae

Squalus acanthias Linnaeus, spiny dogfish

See text for catch and length-distribution data.

Alopiidae

Alopias vulpinus (Bonnaterre), thresher shark

79-046; 3650

82-139; 3910

82-157; 3350

84-024; 3750

84-027; 2015

Carcharhinidae

Galeorhinus zyopterus Jordan and Gilbert, soupfin shark

80-020; 1490 84-109; 1520

81-090; 1590 84-124; 1580

82-139; 1420 84-168; 1530

82-143; 1830 84-169; 1625

82-151; 1610

82-157; 1270,1620,1650

82-165; 1550,1570,1651

83-067; 1680

83-085; 1500

83-104; 1320

84-101; 1575

80

Rajidae

Raja binoculata (Girard), big skate

82-086; 1200

82-138; 1490

83-080; 610

84-003; 1400

84-106; 1270

Raja rhina Jordan and Gilbert, longnose skate

82-118; 898

84-114; 1290

Torpedinidae

Torpedo californica Ayres, Pacific electric ray

81-234; 527

83-153; 480

84-095; 635

OSTEICHTHYES

Clupeiformes

Clupeidae

Alosa sapidissima (Wilson), American shad

79-001; 360,364,369,389,440,489,492 83-152; 312,318,408

79-016; 400 83-153; 235,392,435

79-023; 419,462,470,480 84-030; 350,351,355,360,

79-043; 484 362,388,402,415,440

80-016; 2 - NLA 84-097; 425

80-018; 3 - NLA 84-114; 357,9 (303-446)

80-025; I - NLA

81-041; 282

81-065; 274,286

81-078; 271,294,353

81-079; 271,272,276,277,283,30 I

81-146; 264,270,270,271,275,282,286,290

81-148; 278,300

81-150; 312

81-151; 276,287,295,339

81-168; 26 - 323.0 (267-435)

81-169; 25 - 314.4 (269-472)

81-171; 39 - 372.7 (270-494)

81-172; 286,297,300,377,384,421

81-173; 285,290,397

81-180; 286,292,297,298,299,300,302,306

81-181; 361,418

81-211; 883 - 304.8 (263-409)

81-212; 76 - 305.1 (274-426)

82-131; 12 - 327.8 (285-384)

83-028; 320

83-062; 364
Appendix Table 7-continued.

Clupea harengus pallasi Valenciennes. Pacific herring

See text for catch and length-distribution data.

Sardinops sagax (Jenyns). Pacific sardine

84-048; 250

84-194; 243.248,250,265

Engraulidae

Engrau/is mordax Girard, northern anchovy

See text for catch and length-distribution data.

Salmoniformes

Osmeridae

Allosmerus e/ongatus (Ayres), whitebait smelt

79-043; 110,111,117,126,126

80-017; 20 - 3 lengths available, 102,102,104

82-023; 123

82-029; 89,96

82-128; 99

83-070; 3 - NLA

83-128; 104

Hypomesus pretiosus (Girard), surf smelt

79-037; 134

79-038; 161

80-018; 143

81-001; 128,130,137,140,141,151,154,159

81-147; 160

81-186; 158,164

82-039; 151

82-057; 47 - 147.0 (135-156)

83-014; 27 - 174.0 (159-190)

83-028; 36 - 66.3 (39-84)

83-033; 300 - 165.9 (151-178)

83-053; 20 - 165.7 (162-170)

83-075; 50 - 173.0 (165-187)

84-049; 140,141,145,149,154,154,160

84-060; 155

Spirinchus starksi (Fisk), night smelt

82-079; 140

83-006; 143,146,150,151,162

Tha/eichthys pacificus (Richardson), eulachon

84-148; 120

84-199; 164

Gobiesociformes

Gobiesocidae

Gobiesox maeandricus (Girard), northern c1ingfish

83-128; 32

Gadiformes

Gadidae

Merluccius productus (Ayres), Pacific hake

See text for catch and length-distribution data.

Microgadus proximus (Girard), Pacific tomcod

79-003; 154 82-138; 5 - NLA

79-019; 57 83-121; 91,93

79-020; 64 84-009; 53,55

79-021; 32,50,50,55 84-015; I - NLA

81-079; 46 84-020; 20 - 8 lengths avail., 38,42,44,45,47,49,50,53

81-083; I - NLA 84-147; 73

81-089; I - NLA 84-148; 87,90,103

81-152; 3 - NLA 84-190; 57,61

81-162; I - NLA 84-191; 64

81-163; I - NLA 84-196; 45

81-168; 5 - NLA 84-203; 44,50,51,52

81-263; 56

82-132; 25 - 3 lengths available, 54,63,71

81

Antheriniformes

Scomberesocidae

Co/a/obis saira (Brevoort), Pacific saury

See text for catch and length-distribution data.

Lampriformes

Trachipteridae (standard lengths)

Trachipterus a/tivelis Kner, king-of-the-salmon

81-038; I - NLA

81-043; I - NLA

81-051; 183

81-105; 92

81-118; 267

81-119; 93,142,154,192

81-120; 70,94

81-130; 91,113,114,122,160,161,170,176,210

81-271; 135

82-143; 385

82-145; 121

82-148; 110,120,200,220

82-157; I - NLA

82-168; 156

83-101; 188

83-102; 108

83-112; 193,221

84-169; I - NLA

84-180; 95,115

84-187; 180,198

84-193; 143,205,245

84-198; 224,282

Perciformes

Scorpaenidae (standard lengths)

Sebastes crameri (Jordan), darkblotched rockfish

81-107; 51

Sebastes diploproa (Gilbert), splitnose rockfish

83-124; 32,33,34,36,38,38,39,44,45

83-130; 28,33

84-165; 48

84-200; 35

Sebastes emphaeus (Starks), Puget Sound rockfish

81-048; 37

Sebastes entomelas (Jordan and Gilbert), widow rockfish

79-024; 52,55,59,60

79-025; 61,61,68,68

79-031; 56

79-032; 48 - 61.1 (54-68)

79-036; 65

79-044; 12 - 60.5 (49-68)

79-051; 16 - 69.5 (61-75)

79-053; 41 - 69.9 (56-78)

79-054; 69,71

80-001; 25 - 54.6 (51-58)

81-102; 10 - 51.9 (48-55)

81-105; 46

81-113; 52,52,54

81-115; 54

81-118; 40,43,44,45,45

81-119; 23 - 46.7 (40-51)

81-120; 49 - 51.5 (39-57)

81-121; 13 - 53.3 (48-58)

81-125; 10 - 51.8 (46-57)

82-053; 49

82-079; 50,50

82-083; 50,53

84-027; 52
84-111; 425,430,465,490

84-112; 425,474,488

84-114; 441

84-119;497

84-121; I - NLA

84-123; 356

84-149; 520

84-150; 365,455,470,480,495

84-157; 75 - 423.0 (375-485)

84-175; 502

84-186; 512

Appendix Table 7-continued.

OSTEICHTHYES (continued)

PerciCormes

Scorpaenidae (continued)

Sebastes flavidus (Ayres), yellowtail rockfish

Juveniles:

79-024; 53,54

79-024; 59 - 53.3 (47-58)

79-<l44; 48,51,55

79-052; 52,55

79-053; 54,54,55,55,55,56,57,59

80-001; 220 - 48.9 (45-53)

80-010; 45,47,47,48,50

80-014; 46,49,50,50,50

80-032; 48

80-033; 11 - 47.8 (41-54)

81-102; 40

81-104; 41,43,47,48

81-105; 46,46,46,47,48

81-113; 44,47

81-117; 37,45,45,48

81-118; 29 - 42.0 (35-47)

81-119; 35 - 41.1 (31-47)

81-120; 114 - 45.5 (30-50)

81-121; 12 - 47.0 (40-52)

81-125; 47,48

82-083; 44

83-071; 44

83-078; 46

83-107; 49

84-054; 47

Adults:

82-034; 442

82-037; 1 - NLA

82-072; 490

82-076; 290,485,586

82-113; 420

84-082; 465

84-084; 16 - 412.0 (356-472)

84-150; 1 - NLA

84·189; 387

Sebastes goodei (Eigenmann and Eigenmann), chilipepper

81-062; 36

81-098; 45

Sebastes jordani (Gilbert), shortbelly rockfish

79-025; 65

79-053; 69,70

80-032; 60

81-049; 44

81-075; 55,57,58

81-102; 10 - 57.1 (54-60)

81-107; 53,53,57,59

81-113; 23 - 56.4 (50-60)

81-115; 52,55,55,57,57,57,58,58

81-119; 45

81-121; 55,56,58,63

81-125; 55,59,59

81-201; 56

82-046; 49,54

82-052; 58

82-053; 47,51,52,53

82-054; 55

82-085; 53

82-091; 63

82-100; 61

83-081; 55,56,58,60,61,63

84-027; 52

84-055; 61

84-066; 58

82

Sebastes ma/iger (Jordan and Gilbert), quillback rockfish

79-051; 22

Sebastes me/anops Girard, black rockfish

Juveniles:

79-024; 22 - 51.3 (43-55)

79-025; 48,50,50,52,53,53,53,55,57

79-028; 43,51,52,52,53,54,56

79-032; 49 - 52.8 (48-56)

79-035; 54

79-036; 54

79-043; 53

79-044; 50,51,53,54

79-047; 54

79-050; 52

79-051; 10 - 54.7 (48-58)

79-053; 10 - 55.5 (51-60)

80-001; 103 - 47.2 (43-51)

80-009; 50

80-011; 44,45,47

80-014; 47

80-033; 23 - 48.0 (43-53)

81-031; 42

81-102; 44,44

81-104; 1 - NLA

81-105; 10 - 44.8, (40-49)

81-118; 28 - 42.9 (38-48)

81-119; 23 - 44.7 (40-50)

81-120; 28 - 45.2 (41-50)

81-121; 46,47,47,47,47,48,48,50

81-125; 44

81-126; 48

81-127; 45

82-068; 40

83-071; 46

Adults:

79-007; 482

79-025; 489

80-015; 292

80-016; 1 - NLA

80-031; 1 - NLA

80-041; 1 - NLA

80-044; I - NLA

81-151; 2 - 504,NLA

81-154; 395

81-221; 504

81-254; 491

82-008; 347

82-016; 380

82-018; 362

82-019; 470,484,515

82-027; 492

82-051; 171

82-056; 425

82-071; 450,481,510,508

82-072; 11 - 493.8 (440-530)

82-077; 535

82-088; 488,512,530

82-131; 476,477,510,512,517

82-161; 532

83-007; 294

83-018; 305

83-057; 400

83-070; 10 - 437.3 (392-490)

83-080; 411,437,438,490,511

83-116; 505

84-015; 520

84-020; 470,489,492,515,530,532,540,558

84-021; 476,490

84-080; 492

84-097; 470

84-105; 475
Appendix Table 7-conlinued.

--.:..:---------------------------------_._----

Sebastes mystinus (Jordan and Gilbert), blue rockfish

Juveniles:

79-023; 57

79-024; 12 - 57.1 (51-60)

79-025; 53,57,58,58,58,59

79-026; 54

79-027; 60

79-028; 51,53,56,56,58,59,59,59

79-032; 83 - 58.1, (51-63)

79-042; 51,52,66

79-043; 55

79-044; 53,54,58,59,59,60,60,64

79-047; 60

79-051; 13 - 61.5 (58-71)

79-052; 55

79-053; 58 - 60.6 (53-66)

79-054; 65

80-001; 123 - 51.3 (45-58)

80-011; 51

80-014; 51,53

80-033; II - 54.0 (50-57)

81-005; 47

81-102; 49,40,56

81-104; 58

81-113; 1 - 1\LA

81-115; 54

81-117; 47

81-118; 47,47,49

81-119; 21 - 465 (42-50)

81-120; 34 - 48.0 (45-51)

81-121; 23 - 50.7 (47-58)

81-201; 48,48

Adults:

82-160; 440

82-170; 386,407

Sebastes nigrocinctus Ayres, tiger rockfish

83-124; 17 - 44.3 (30-56)

84-157;42

Sebastes paucispinis Ayres, bocaccio

79-051; 80

79-053; 79,83,86,86,89,89

81-119; 53

81-120; 53,60

81-121; 41

81-125; 65

Sebastes pilllliger (Gill), canalY rockfish

79-026; 44

79-030; 46

79-031; 39

79-032; 46

79-044; 45,46,47

79-052; 45

82-052; 38

Sebastes proriger (Jordan and Gilbert) redstripe rockfish

84-192; 27

84-201; 13 - 27.7 (23-35)

Sebastes saxicnlu (Gilbert), stripetail rockfish

81-075; 35

81-098; 35,35,36

81-105; 35,35,35,35

81-119; 29

81-120; 30.33,34,35,36,36,37,38

Anoplopomatidae

Anoplopoma fimbria (Pallas), sablefish

See text for catch and length-distribution data.

83

Hexagrammidae

Hexagrammos decagrammus (Pallas), kelp greenling

79-031; 19 - 57.3 (53-60)

79-042; 63,64,65

79-044; 64

81-048; 61

81-049; 67

81-062; 63,65,66,68

81-071; 61,61,64,64,64,65,67,67,68

81-098; 60,67,67,71

81-121; 63,66,67

81-132; 27 - 65.1 (54-72)

82-060; 73

82-062; 72

82-064; 3 - NLA

82-069; 68

82-092; 58

82-102; 65

83-053; 48

84-016; 54

84-029; 65

84-038; 62,65

84-056; 65,69

84-061; 67

84-067; 65

84-068; 68

Ophiodon elongatus Girard, lingcod

Juveniles:

81-022; 67

81-036; 53,59,60,61,61,63,66

81-037; II - 53.1 (48-66)

81-038; 50 - 58.6 (48-66)

81-042; 71

81-048; 59,60,61,64,65,65,65,69

81-049; 65

81-093; 76

82-085; 51,54,59,60,62,63

82-125; 1 - NLA

Adults:

84-158; 635

Coltidae

Hemilepidotus spinosus (Ayres), brown Irish lord (standard lengths)

79-044; 25,26,29

79-053; 43

80-030; 1 - NLA

81-048; 27

81-049; 28,28

82-092; 23,24,25,25,25,26

83-043; 27

Leptocottus armatus Girard, Pacific staghorn sculpin

79-026; 175

82-160; 198,213,226,226,228

83-037; 195

Scorpaenichthys marmoratus (Ayres), cabezon (standard lengths)

80-001; 46

80-013; 41

80-014; 49

80-092; 46

83-045; 35

83-064; 41,43

83-112; 43
Appendix Table 7-continued.

OSTEICHTHYES (continued)

Perciformes

Agonidae

Agonopsis vul'a (Jordan and Gilben), nonhern spearnose poacher

82-052; 58

82-100; 68

82-120; 63

84-084; 73

84-116; 65

Cyclopteridae (total lengths)

Liparis pulchellus Ayres, showy snalifish

84-148; 99

Carangidae

Trachurus symmetricus (Ayres), jack mackerel

See text for catch and length-distribution data.

Kyphosidae

Medialuna californiensis (Sleindachner), halfmoon

83-135; 12 - 231.9 (210-247)

Trichodontidae

Trichodon trichodon (Tilesius), Pacific sandfish

79-047; 178

Zaproridae

Znprora silenus Jordan, prowfish (total length)

82-155; 200

Ammodytidae

Ammody/es hexap/erus Pallas, Pacific sand lance

82-128; 79

Scombridae

Sarda chiliensis (Cuvier), Pacific bonito

83-133; 640,653

Scomber japonicus Houttuyn, chub mackerel

See texl for catch and length-distribution data.

Stromateidae

Peprilus simillimus (Ayres), Pacific pompano

79-037; 173

82-039; 166

83-014; 176

83-020; 196

83-021; 179

83-050; 6 - NLA

83-056; 192

83-058; 155,163,169,170,175,17<),180

84-015; 168,170,180,183,185.190,191,195.195

84-155; 175,179,189

lcich/hys locking/oni Jordan and Gilbcn, medusafish Anarhichadidae

Anarrhich/hys oce/latus Ayres, wolf-eel

79-001; 457

79-002; 506

79-003; 477

79-012; 1 - NLA

79-016; 464,48:

79-020; 463

79-024; 505

79-025; 458

79-026; 458,460,472,508,518

79-027; 494,503,532,548,599

79-028; 520

79-029: 492

79-030; 478

79-033; 425,485,610

79-034; 487,496,577 ,587

79-035; 18 - 488.9 (432-546)

79-037; 510,620

79-038; 502,508,508,512,528

79-039; 458,463,473,487,489,492,494,533,548

79-043; 457,48 1,483,500,512,532,557,586

79-044; 437,475,480,488,493

79-045; 418,443,453,477,509,512

79-046; 436,475

79-047; 432,505

79-048; 434,482,560

79-049; 418,50l,NLA

79-050; 413,438,453,463,468,483,502,502,509

79-051; 277,328,357,387,397,405,452

79-053; 451,455,511

79-054; 458

79-055; 405,540

80-037; 591

80-043; 521

80-044; 537

81-099; I - NLA

81-110; I - NLA

81-188; 530

81-190; 438

81-214; 456

81-241; 472

82-028; I - LA

82-044; 1200

82-068; 516,613

82-079; 495

82-098; 1440

82-101; 399

82-103; 610

82-117; 490

82-1 \8; 460

82-121; 2 - NLA

82-151; 490

82-160; 640

83-025; 1252

83-042; 480

83-065; 1220

83-094; 1 - NLA

83-102; I - NLA

83-104; I - NLA

83- 108; 580

84-082; 525

84-092; 500

84-094; 570

84-097; 515,530

84-102; 515

84-150; 600

84-203; 452

84

79-038; 110

81-119; 58

81-125; 60

81-175; 101

81-220; 116

81-272; I - NLA

82-002; 175

82-004; 139

82-085; 155

82-132; 127

82-143; 170,183

82-144; 91

82-150; 151,153

82-153; 154

82-155; 189

82-156; 150

82-157; 67,86

82-160; 2 - NLA

82-162; 160,170,172

82-165; 185

82-167; 124

82-168; 110,155

82-170; 120

82-172; 117,142,161

83-033; 152

83-037; 125

83-107; 90,94,100

83-110; 90,95,114

83-1 I 1; 103

83-112; 65,110

83-114; 95

83-130; 95,151

83-131; 84,134,162

83-135; 47

83-137; 121

83-140; 138

83-142; 178

83-143; 98

83-147; 71,113.121

83-148; 119,125,134,194

83-149; 132

83-150; 86,89

83-153; 79,80,105,138

83-154; 128

83-155; 110,119,130,132

83-156; 82,130

83-158; 61

83-159; 95,96,98,102,104,136,147

83-160; 169

83-161; 83,100,117,163,187

83-163; 75,80,118

83-164; 195

83-165; 130,163,176

83-166; 86,104,120

83-167; 66,68,80.87,97,124,144

83-170; 103,105,111,113,132,150

83-171; 91

83-172; 95,114,169,174

83-173; 107,110

84-189; 83

84-191; 113

84-203; 80
Appendix Table 7-continued.

Icosteidae

Icosreus aenigmalicus Lockmgton, ragfish

79-012; 1 - NLA

81·112; 69

82·135; 134

82-138; 272

83-107; 270

Pleuronectiformes

Bothidae (total length)

Citharichthys sordidus (Girard), Pacific sanddab

79-017; 208

79-044; 28 - 258.8 (233-315)

80-001; 39

81-010; 39

81-109; 165

81-211; 220

81-250; 272

82-068; I - NLA

83-172; 268,282,299,316

84-061; 190,270

84-105; 265,300

84-169; 263

84-191; 70 - 236.2 (203-281)

Citharicluhys stiwnaeus Jordan and Gilbert, speckled sanddab

80-033; 37

82-128; 59,63

Pleuroneclidae (total lengths)

Atheresthes .<tomias (Jordan and Gilbert), arrowtooth flounder

83-006; 120,130

Glyptocephaills wchirus Loekington, rex wie

81-125; 80

!sopselta isulepis (Lockinglon), butter sale

79-049; 210

Lyopselta exilis (Jordan and Gilbert), slender sale

79-030; 136

82-038; 3 - NLA

82-114: 53

~4-089; 132

84-091; 165

Microstumus pacific'lIs (Lockington), Dover sale

82-114; 52

Paruphrys vetulus Girard, English sale

82-160; 232,NLA

83-056; 208

84-105; 240

84-148; 24 - 173.0 (120-191)

Platiclullys stellatus (Pallas), starry flounder

79-027; 415,418,420,431,445,460,463

79-047; 331,340,530

80-041; 389

81-042; 215

81-124; 311

81-204; 408,475

81-228; 316,410

82-060; 407,493,517,547

82-116; 477,490,517,523,540,542,589

82·117; 450

82-122; I - NLA

82·161,603

83-075; 509

84-148; 456,565

Pseltichthys me/anostictus Girard, sand sole

82-128; 125,130,152,185, 197.NLA,NLA

"~c. U.S. GOVERNMENT PRINTING OFFICE:1987-791-121/61003 85

Tetraodontiformes

Molidae (total lengths)

Mola fIlula (Linnaeus), ocean sunfish

81-103; 650

81·140; 066

81-139; 2235

82·138: 660

~2-143; 737,838,804

82-145; 610

83-114; 405

83-115; 470

83-117; 425

83-130; 450

83·136; 340,360,430

84-033; 1780

84-110; 1675

84-163; 350,380

84·165; 440

84-167; 400

84-l"I3; 440

84-184; 680

b4-188; 375,600,695,995

~4-201;415