J.F. GATES CLAR Neotropical

Microlepidoptera XIX:

Notes on and

; New Species of

Oecophoridae

Lepidoptera)

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY NUMBER 95
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SMITHSONIAN CONTRIBUTIONS TO

ZOOLOGY

NUMBER 95

J. F. Gates Clarke Neotropica l

Microlepidoptera XIX:

Notes on and

New Species of

Oecophoridae

(Lepidoptera)

SMITHSONIAN INSTITUTION PRESS

CITY OF WASHINGTON
ABSTRACT

Clarke, J. F. Gates. Neotropical Microlepidoptera XIX: Notes on and New Species

of Oecophoridae (Lepidoptera). Smithsonian Contributions to Zoology, number 95,

39 pages, 1971.—Costoma Busck ( = Phalarotarsa Meyrick) (new synonymy) ; Gonionota aethoptera, aethographa, cologramma, oxybela, charagma, hypoleuca, argopleura, amauroptera, gaiophanes, hemiglypta, eremia, insignata, menura, paravexillata, transversa, sphenogramma, selene, poecilia, phthiochroma and Hypercallia

heterochroma are described. Gonionota autocrena (Meyrick), Gonionota rosacea

(Forbes), new combination; G. dryodesma (-H. dryocrypta), H. miltopa Meyrick

{=Cryptolechia tunicata Busck), Phytomimia chlorophylla Walsingham (=Phytomimia silvicolor Meyrick, (new synonymy); Coptotelia bipunctalis (Warren), new

combination, ( = Coptotelia byrsocyma (Meyrick)) (new synonymy).

Official publication date is handstamped in a limited number of initial copies and is recorded

in the Institution's annual report, Smithsonian Year.

UNITED STATES GOVERNMENT PRINTING OFFICE

WASHINGTON : 1971

For sale by the Superintendent of Documents, U.S. Government Printing Office

Washington, D.C. 20402 - Price 50 cents (paper cover)
J. F. Gates Clarke Neotropical

Microlepidoptera XIX:

Notes on and

New Species of

Oecophoridae

(Lepidoptera)

Introduction

The purpose of this paper, the nineteenth in this

series, is to clarify points of synonymy, both generic

and specific, to illustrate previously unfigured species,

and to describe an accumulation of new taxa so that

their presence in the Neotropical fauna may be made

known.

The drawings for this paper were made by Mr.

Andre del Campo Pizzini, staff artist. The photographs were made by Mr. Victor Krantz, Smithsonian Photographic Laboratory.

Genus Costoma Busck

Costoma Busck, 1914:23 (Type species: Costoma basirosella Busck 1914:24 [by monotypy and original

designation]).

Phalarotarsa Meyrick, 1924:101 (Type species: Phalarotarsa cirrophaea Meyrick 1924:102 [by monotypy]) (New

synonymy).

I have compared the types of both genera, and although basirosella and cirrophaea are specifically distinct, there is no question about their generic identity.

/. F. Gates Clarke, Department of Entomology, National

Museum of Natural History, Smithsonian Institution,

Washington, D.C. 20560.

Costoma basirosella Busck

FIGURE 1

Costoma basirosella Busck, 1914:24.—Meyrick 1922:185.

TYPE.—United States National Museum.

TYPE LOCALITY.—Panama, Trinidad River.

DISTRIBUTION.—Panama."

The genitalia of basirosella are figured here, for

the first time, so that those of the male can be compared with those of cirrophaea (see Clarke, 1963:

373, pi. 183: figs. l-ld).

Costoma cirrophaea (Meyrick), new combination

FIGURE 2

Phalarotarsa cirrophaea Meyrick, 1924:102.—Clarke 1963:

373, pi. 183: figs. l-ld.

I have already figured this species (1963) where the

characters of the male are clearly illustrated. The

female genitalia are figured here from a specimen

from Bolivia, Prov. del Sara, Dep. Sta. Cruz, 450 m,

slide JFGC 12043, and can be compared with those of

basirosella.

TYPE.—British Museum (Natural History).

TYPE LOCALITY.—Bolivia, Prov. del Sara, Dep. Sta.

Cruz.

DISTRIBUTION.—Bolivia.

1
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Genus Gonionota Zeller

Gonionota aethoptera, new species

FIGURE 3; PLATE la

Alar expanse 24-25 mm.

Labial palpus light cinnamon brown; second segment with gray apical annulus; third segment suffused gray, apex buff. Antenna light cinnamon brown

with faint grayish annulations; scape buff ventrally.

Head light cinnamon brown mixed with gray in

lateral scale tufts. Thorax light cinnamon brown,

scales of crest tipped with dark brown. Forewing

ground color light cinnamon brown; scattered gray

scales along fold to tornus and irregularly in apical

half; termen narrowly blackish from vein 4 to tornus;

at apical third of costa a small white spot; at basal

third, in cell, two black spots; at end of cell a conFIGURE 1.—Costoma basirosella Busck, a, ventral view of

male genitalia with left harpe removed and aedeagus in

situ; b, ventral view of female genitalia.
NUMBER 95

spicuous white spot; dorsum with conspicuous dark

brown streak; cilia light cinnamon brown. Hindwing

light ocherous tawny overlaid with gray scales; cilia

pale grayish. Foreleg ocherous white; femur, tibia,

and first tarsal segment overlaid light cinnamon

tawny; midleg ocherous white with dark gray shade

on tibia; tarsal segments yellowish; hindleg ocherous

white with grayish suffusion and spots on tibia; tarsal

segments yellowish. Abdomen ocherous white, suffused grayish dorsally.

Male genitalia slide JFGC 12113. Harpe broadest

basally, gently narrowed toward the rounded cucullus; sacculus broadly sclerotized to slightly beyond

middle and terminating in a large setose protuberance. Gnathos a narrow, transverse, oval spined knob.

Uncus very narrow basally terminating in widely divergent arms. Vinculum a narrow band. Tegumen

rather broad, convex laterally, shorter than harpe.

Anellus a very strongly sclerotized, subrectangular

plate with a narrow dentate projection posteriorly at

each corner. Aedeagus very stout, curved, distally

shovel shaped; vesica armed with two sclerotized

plates each bearing a cluster of stout cornuti.

Female genitalia slide JFGC 12060. Ostium Vshaped, laterally strongly sclerotized. Antrum a very

narrow sclerotized ring. Inception of ductus seminalis at dorsoanterior edge of antrum. Ductus bursae

membranous with very finely granular surface, the

granular surface continuing as a narrow band to

anterior third of the membranous bursa copulatrix.

Signum an elongate, sclerotized, dentate plate. Lamella antevaginalis narrow, sclerotized. Lamella postvaginalis membranous with granular surface.

HOLOTYPE.—USNM 71083.

TYPE LOCALITY.—Venezuela, Aragua, Rancho

Grande, 1100 m.

DISTRIBUTION.—Venezuela.

Described from the $ holotype, one c? and 3 $ ?

paratypes all from the same locality as holotype and

with following dates, etc., o* (16-23. X. 66); ?

(24-31. X. 66), $ (1-5. XI. 66) all collected by

S. S. and W. D. Duckworth; $ (19. VI. 67), ?

(22-31. VII. 67), both collected by R. W. Poole.

Under cologramma I have pointed out that

aethoptera and several other species are related to it,

but aethoptera is a considerably larger moth than

cologramma, is generally darker in color, with a much

darker hindwing.

FIGURE 2.—Costoma cirrophaea (Meyrick). Ventral view

of female genitalia.
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Gonionota aethographa, new species

FIGURE 4; PLATE \b

Alar expanse 18 mm.

Antenna hazel, dorsally overlaid red. Head cinnamon buff, suffused dark gray on crown and with

roseate tinge. Thorax cinnamon buff with roseate

tinge; posterior tuft suffused dark gray. Forewing

ground color sayal brown; basal half of wing, marked

by an oblique line from costal two fifths to tornus,

cinnamon buff, shading to pinkish buff before dorsum; dorsum marked by a dark brown streak, broadest at middle, extending from basal fourth to tornus;

costa narrowly roseate and with slender pink streak at

apical third; in cell, at basal third, two small fuscous

spots and a similar single one at end of cell; on terFIGURE 3.—Gonionota aethoptera, new species, a, ventral

view of male genitalia with left harpe and aedeagus removed; b, aedeagus; c, ventral view of female genitalia.

: • 1 .

3c
NUMBER 95

FIGURE 4.—Gonionota aethographa, new species, a, ventral view of male genitalia with left

harpe and aedeagus removed; b, aedeagus.

men, between veins, five ill-defined, fuscous dashes;

cilia sayal brown. Hindwing sayal brown; cilia

slightly paler. Foreleg shining white, outer side of

femur overlaid brown; tibia brown on outer side

mixed carmine; first tarsal segment carmine on outer

side, remainder cinnamon buff; midleg white; tibia

suffused cinnamon and with black spot on outer side;

hindleg creamy white. Abdomen clay color, shading

to brown dorsoposteriorly; ventrally buff with sparse

clay-color irroration.

Male genitalia slide JFGC 12112. Harpe broadest

at base, gently tapered toward cucullus; from base of

costa to near middle of harpe, a strong sclerotized

bar; sacculus rather narrowly sclerotized, terminating

in a small setose protuberance at outer end of sclerotized base. Gnathos a broad, transverse, finely spinose

knob. Uncus (missing). Vinculum a narrow band

with median anterior convexity. Tegumen slightly

longer than aedeagus. Anellus rectangular, slightly

broadened proximally, and with deep concavity distally. Aedeagus stout, curved, pointed; vesica armed

with two clusters of comuti, each arising from a

small sclerotized plate, and four long, straight

cornuti.

HOLOTYPE.—USNM 71084.

TYPE LOCALITY.—Costa Rica, Las Cruces, nr. San

Vito.

DISTRIBUTION.—Costa Rica.

Described from the unique cf holotype (19-20.

III. 65, S. S. and W. D. Duckworth).

Nearest aethoptera, this species belongs in the hydrogramma-cologramma group. It differs from

aethoptera in the absence of the conspicuous white

spot at end of cell and by the presence of the four

strong cornuti in addition to the two clusters.

Gonionota cologramma, new species

FIGURE 5; PLATE \C

Alar expanse 20 mm.

Labial palpus tawny; second segment with a faint

rosy hue and with a few blackish scales apically;

third segment with blackish scales toward apex; apex

ocherous white. Antenna reddish basally, shading to

light brown toward apex and faintly annulated gray;

scape reddish dorsally, ocherous white ventrally.

Head mixed gray, tawny, and reddish. Thorax tawny

with gray dorsal spot posteriorly; tegula tipped ocherous white. Fore wing ground color tawny, outer half

crossed by numerous, irregular, fine pale lines; costa
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

narrowly gray mixed with some pink scales; at apical

fifth of costa a gray spot; at basal third, in cell, two

blackish discal spots, one larger than the other; at

end of cell a white spot; around termen a series of

short blackish dashes; on fold, about middle a dark

brown spot; dorsum with a conspicuous brown stripe;

costal area, and from fold to the brown dorsal streak,

suffused rosy-pink; cilia tawny. Hindwing buff, somewhat darker toward anal angle. Foreleg ocherous

with scattered tawny and pink scales; midleg similar;

hindleg ocherous white. Abdomen ocherous white,

suffused tawny dorsally.

Male genitalia slide JFGC 12058. Harpe of about

equal width to beyond middle, then gently narrowed

to a truncate cucullus; sacculus broadly sclerotized,

recurved beyond middle. Gnathos a long, narrow,

transverse roll. Uncus broad basally, constricted at

middle, terminating in two divergent arms. Socii two

setose papillae. Vinculum a narrow band. Tegumen

about as broad as long, constricted at middle. Anellus a subrectangular plate with a slender pointed

process posteriorly at each corner. Aedeagus stout,

curved, apex linguiform; vesica armed with a cluster

of short, stout cornuti.

HOLOTYPE.—USNM 71085.

TYPE LOCALITY.—Venezuela, Aragua, Rancho

Grande, 1100 m.

DISTRIBUTION.—Venezuela.

Described from the unique c? holotype (10-21. II.

69, W. D. Duckworth and R. Dietz).

Superficially cologramma is much like G. hydrogramma (Meyrick), exhibiting the same type of pale

transverse strigulae found in that species, but the pale

hindwing of cologramma immediately distinguishes

it from the latter. In male genitalia the two are

strikingly similar, the chief difference being the

length of the basal bar of the clasper which extends

beyond middle of harpe in hydrogramma and terminates at or before middle of harpe in cologramma.

The cornuti are more numerous in cologramma than

in hydrogramma but must be compared (see Figure

5 and Clarke 1963: pi. 114: fig. 4a).

FIGURE 5.—Gonionota cologramma, new species, a, ventral view of male genitalia with left

harpe and aedeagus removed; b, aedeagus.
NUMBER 95

G. hydro gramma, anisodes, cologramma, aethographa and aethoptera constitute a group of similarly

colored species all having in common a dark dorsal

stripe on the forewing.

Gonionota autocrena (Meyrick), new combination

FIGURE 6; PLATE \d

Hypercallia autocrena Meyrick, 1930: (3)575.

Male genitalia slides JFGC 10812, 12071. Harpe of

nearly equal width throughout; cucullus bluntly

pointed; clasper a curved bar terminating in a cluster

of spines and hairlike setae; sacculus rather broadly

sclerotized. Gnathos a spined, transverse oval knob.

Uncus moderately broad basally, terminating in two

divergent points. Socii rather small setaceous

swellings at base of uncus. Vinculum U-shaped. Tegumen about as long as harpe. Anellus a sclerotized

plate, broad basally, narrowed, and with a long spine

at each corner posteriorly. Aedeagus moderately

slender, curved, pointed distally.

FIGURE 6.—Gonionota autocrena (Meyrick). a, ventral

view of male genitalia with left harpe and aedeagus removed; b, aedeagus; c, ventral view of female genitalia.
8 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Female genitalia slide JFGC 10813. Ostium a deep

cup. Genital plate subrectangular, sclerotized. Antrum sclerotized. Inception of ductus seminalis at

junction of antrum and membranous portion of

ductus bursae. Ductus bursae membranous and with

faint and sparse granulations. Bursa copulatrix membranous. Signum absent. Lamella postvaginalis membranous.

TYPE.—Museum National d'Histoire Naturelle,

Paris.

TYPE LOCALITY.—Brazil, Nova Friburgo, Organ

Mountain.

DISTRIBUTION.—Brazil.

In addition to the type, I have before me 2 cf cf,

$ from Brazil, Nova Teutonia (August 1948) Fritz

Plaumann.

The species is clearly referable to Gonionota and

has not been figured previously.

The following group of three new species, charagma, hypoleuca, and oxybela, belong with and are

related to incisa, phocodes and lichenista. It is virtually impossible to separate the six species on superficial characters, especially when the specimens are

worn, but the genitalia are distinct (except those of

lichenista are unknown) and preclude misidentification.

Gonionota oxybela, new species

FIGURE 7; PLATE le

Alar expanse 18—20 mm.

Labial palpus a mixture of tawny olive and sayal

brown with sparse, black irroration; apex of second

segment snuff brown; third segment snuff brown except base and apex. Antenna clay color, spotted and

suffused fuscous. Head a mixture of tawny olive and

sayal brown. Thorax bister brown with median

tawny olive line and sparse ochraceous-tawny irroration. Forewing ground color bister brown with

tawny-olive irroration; in outer half of wing considerable ochraceous-tawny scaling; costa ochraceous

FIGURE 7.—Gonionota oxybela, new species, a, ventral view of male genitalia with left harpe

and aedeagus removed; b, aedeagus.
NUMBER 95

tawny mixed with tawny olive; from basal third of

costa an ill-defined tawny-olive line extends to fold

where it continues outwardly to tornus; cilia bistre,

those in apical half mixed with ochraceous tawny.

Hindwing sepia above; beneath mostly tawny olive

with considerable ochraceous tawny along costa;

cilia sepia, ochraceous tawny-tipped around termen.

Foreleg white; tibia sepia on outer side; tarsal segments shading from sepia to clay color; midleg similar but tarsal segments paler; hindleg femur white

with sepia on outer side; tibia ocherous white with

sepia on outer side; spurs white and tibia with white

transverse line at middle and apex. Abdomen dorsally sepia with longitudinal, lateral, ochraceoustawny line; ventrally white except last three segments

mixed fuscous, ochraceous tawny, and clay color.

Male genitalia slide JFGC 4010. Harpe oval; cucullus produced as a long, slender process; sacculus

with large bulbous, spined process at outer end;

from base a dentate keel extends to one third. Gnathos

an oval spined knob. Uncus slender, curved, digitate.

Vinculum a narrow band with median anterior expansion. Tegumen as long as harpe. Anellus an elongate, rectangular plate with basal protuberance on

each side. Aedeagus long, curved; distally linguiform.

HOLOTYPE.—USNM 71086.

TYPE LOCALITY.—Peru, Cuzco Province.

DISTRIBUTION.—Peru.

Described from the <S holotype (no date or collector) and 4 o* o* paratypes, same data as holotype.

Of the several species to which oxybela is related,

it is nearest phocodes and differs from it mainly by

the long, pointed cucullus.

Gonionota charagma, new species

FIGURE 8; PLATE 1/

Alar expanse 16—20 mm.

Labial palpus olive buff irrorate with reddish

brown and blackish fuscous; apex of second segment

fuscous; third segment blackish fuscous in tuft, apex

buff. Antenna light reddish brown spotted grayish

fuscous. Head olive buff mixed grayish fuscous. Thorax olive buff strongly suffused fuscous; tegula with

scattered reddish brown scales. Forewing ground

color fuscous, overlaid and suffused reddish brown;

from basal third of costa an all-defined olive-buff

fascia extends to fold then contines to tornus; from

apex to vein 2, an ill-defined olive-buff transverse

fascia; cilia fuscous and reddish brown mixed. Hindwing sepia on upper side, underside light tawny olive

with olive buff, reddish brown, and grayish-fuscous

irrorations; cilia clay color with broad fuscous basal

band. Foreleg white, tibia fuscous on outer side; tarsal segments overlaid red brown; midleg white; tibia

fuscous on outer side except apically edged white;

hindleg similar except tibia with white median annulus and white median spurs. Abdomen fuscous dorsally with reddish-brown scales posterolaterally; ventrally shining white and reddish-brown scaling laterally on terminal segment.

Male genitalia slides JFGC 4015, 10929. Harpe

as long as vinculum and tegumen combined, of almost

uniform width throughout and only slightly broader

at middle; sacculus broadly sclerotized, confluent at

outer end with sclerotized rod arising from base of

clasper; termination of rod and sacculus armed with

a strongly setose bulbous process. Gnathos an oval

spined knob. Uncus broad basally, terminating in a

knob with shallow distal concavity. Vinculum a narrow band with median anterior process. Tegumen

broad basally, narrowed posteriorly. Anellus subrectangular, expanded basally on each side; posterior

edge concave. Aedeagus curved, stout, distal end

elongate, linguiform; cornuti absent.

Female genitalia slide JFGC 4016. Ostium narrowly

V-shaped. Genital plate strongly sclerotized; lamella

postvaginalis granular. Antrum narrowly sclerotized.

Inception of ductus seminalis dorsal and slightly

anterior to antrum. Ductus bursae membranous.

Bursa copulatrix membranous. Signum an elongate,

diamond-shaped, sclerotized, finely dentate plate.

HOLOTYPE.—USNM 71087.

TYPE LOCALITY.—Brazil, Nova Teutonia.

DISTRIBUTION.—Brazil.

Described from the o* holotype (IV. 1948, Fritz

Plaumann), 3 o* o* and one $ paratypes from Brazil

(Sta. Catharina, Oct. 1-4, 1934, Fritz Hoffmann).

In coloring charagma is very similar to incisa from

Bolivia, but incisa is darker, is practically devoid of

red-brown scaling and the underside of hindwing

is darker and is suffused. In the male genitalia the

cucullus of incisa is rounded (Clarke 1963, pi. 115:

figs. 2—2b) with a short point at outer end of costa.

In incisa there is also a triangular clasper about the

middle of harpe which is absent in charagma. The

harpe of charagma is twice as long as broad but that

of incisa is less than IJ/2 times as long as broad.
10 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Gonionota hypolenca, new species

FIGURE 9; PLATE \g

Alar expanse 16—22 mm.

Labial palpus first segment white; second segment

finely mottled white, chestnut brown and fuscous;

apex blackish fuscous; third segment almost wholly

blackish fuscous except basal annulus white and apex

ocherous white. Antenna buffy brown basally, grayish

toward distal end with blackish annulations. Head

buffy brown and grayish mixed. Thorax sepia; tegula

snuff brown, edged ocherous white. Fore wing ground

color snuff brown, many of the scales tipped whitish;

from basal third of costa a very faint, ill-defined white

line, interrupted in cell by a fuscous spot, extends to

fold; tornus with an ill-defined fuscous blotch; cilia

sepia. Hindwing sepia above, tawny olive, overlaid

and suffused grayish fuscous beneath; cilia grayish

fuscous with darker basal line. Foreleg white; tibia

sepia on outer side; tarsal segments clay color; midleg

similar; tibia blackish fuscous on outer side except

white basally and distally and a fine white line across

middle; spurs white; hindleg white; femur blackish

fuscous on outer side; tibia sepia on outer side, except spurs, a median line and apex white; tarsal

segments shading from sepia to clay color. Abdomen

sepia dorsally; laterally a longitudinal, cinnamon

brown line; ventrally white except terminal segment

sepia.

Male genitalia slides JFGC 11539, 12047. Harpe

very broad, short; cucullus bluntly pointed; sacculus

\

FIGURE 8.—Gonionota charagma, new species, a, ventral view of male genitalia with left harpe

and aedeagus removed; b, aedeagus; c, ventral view of female genitalia.
NUMBE

R 95

11

broadly sclerotized, rugose; at end of sacculus a

round, strongly spined protuberance. Gnathos a long,

transverse spined knob. Uncus short, straight, with

apical group of setae; socii elongate setose lobes.

Vinculum a narrow band. Tegumen about as long

as harpe. Anellus an elongate plate with laterobasal

projections, constricted slightly before posterior end;

posterior margin concave. Aedeagus long, curved, ter-

minating in a long pointed process.

Female genitalia slides JFGC 11540, 12048. Ostium

oval, small. Genital plate strongly sclerotized except

central portion membranous, finely granular. Antrum

narrowly cylindrical, sclerotized. Inception of ductus

seminalis slightly anterior to antrum. Ductus bursae

membranous. Bursa copulatrix membranous. Signum

an elongate, diamond-shaped, dentate plate. Lamella

antevaginalis rugose. Lamella postvaginalis somewhat

granular.

HOLOTYPE.—USNM 71088.

FIGURE 9.—Gonionota hypoleuca, new species, a, ventral

view of male genitalia with left harpe and aedeagus re-

moved; b, aedeagus; c, ventral view of female genitalia.
12 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

TYPE LOCALITY.—Venezuela, Aragua, Rancho

Grande, 1100 m.

Described from the o* holotype (11-15. Jan. '66,

S. S. and W. D. Duckworth) 11 o* d" and 4 $ ?

paratypes, same locality and collectors (11-25. Jan.

'66; 24-31. X. '66; 1-5. XI. '66).

As pointed out above, all members of this group

are similar but hypoleuca appears to be the nearest

to the Peruvian phocodes from which it differs by

the abundance of white in the labial palpus. In genitalia the harpe of phocodes is very broad and shorter

in proportion to its length than that of hypoleuca,

and from the base of sacculus there is a long spatulate process in phocodes which is absent in hypoleuca.

I do not know the female of phocodes and can draw

no comparison with it.

Gonionota argopleura, new species

FIGURE 10; PLATE \h

Alar expanse 22 mm.

Labial palpus sordid whitish; second segment

speckled with tawny olive and clay color and shading

to snuff brown at apex; third segment snuff brown

except posterior edge and apex. Antenna tawny olive,

suffused grayish. Head and thorax tawny olive, the

scales tipped whitish. Forewing ground color fawn;

basal half of costa white, shading to gray, then merging with ground color; apical third of costa white

with a dark brown spot near inner end; on termen

a large ochraceous buff blotch between veins 4 and 7;

in cell, at basal two fifths, an ill-defined dark gray

spot; on middorsum a dark brown shade; cilia tawny

olive. Hindwing buff basally, shading to olive buff

toward margins; cilia buff. Foreleg light cinnamon

buff; midleg similar; tibia with gray spot at base of

spurs; hindleg ocherous white overlaid cinnamon

buff on outer side. Abdomen light grayish brown with

white scales mixed.

Female genitalia slide JFGC 10859. Ostium crescentic. Genital plate strongly sclerotized, rugose laterally and with two ridges converging anterior to

ostium. Antrum sclerotized. Inception of ductus seminalis at anterior edge of antrum. Ductus bursae

sclerotized posteriorly, ribbed. Bursa copulatrix membranous. Signum a narrow, oval, sclerotized transverse

plate.

FIGURE 10.—Gonionota argopleura, new species. Ventral

view of female genitalia.
NUMBER 95 13

HOLOTYPE.—USNM 71089.

TYPE LOCALITY.—Brazil, Santa Catharina.

DISTRIBUTION.—Brazil.

Described from the unique $ holotype (Sept. 26.

'34; Fritz Hoffmann).

In some respects argopleura is similar to the Colombian eurydryas but the sinuation of the costa is

beyond middle in argopleura and the latter lacks any

indication of the pale transverse striae of eurydryas;

also, eurydryas lacks the large ochraceous-buff blotch

before termen, found in argopleura.

Gonionota atnauroptera, new species

FIGURE 11; PLATE 2b

Alar expanse 16-20 mm.

Labial palpus ocherous white on inner side, irrorate

with fuscous and hazel; second segment hazel on

outer side with white-tipped fuscous scales in posterior tuft; third segment similarly marked, with

whitish apex. Antenna gray, somewhat paler apically;

scape whitish ventrally. Head hazel, scales white

tipped and appearing gray. Thorax hazel; dorsally

marked with yellow. Forewing ground color hazel;

costa narrowly reddish, extreme edge whitish; at

basal fifth an indistinct, transverse, fuscous fascia and

a similar outwardly oblique one at two fifths; at

apical third of costa a wedge-shaped spot varying

from white to reddish; in cell at basal third a pair of

minute black discal spots; at end of cell a black-edged

white spot, obsolete in one specimen; base of dorsum

yellow becoming orange rufous toward tornus; on

termen an ill-defined row of fuscous spots; cilia grayish fuscous.

Hindwing ocherous white basally, shading to pale

brownish apically; underside with sharply defined

row of blackish dashes around termen between veins;

cilia ocherous white to pale brownish at apex. Foreleg

ocherous white, shaded with hazel and reddish on

outer side; midleg similar; hindleg ocherous white

irrorate with fuscous on outer side. Abdomen pale

buff; dorsally an ill-defined, longitudinal, median

brownish suffusion; ventrally and laterally a longitudinal row of small brownish to black spots.

Male genitalia slides JFGC 10890, 12034, 12035.

Harpe broadest near base, gently tapered to the

pointed cucullus; clasper a cluster of curved spines

terminating before middle of harpe. Gnathos a rather

narrow, subrectangular, tranverse spined knob. Uncus

a broadly based slender stalk terminating in widely

divergent arms. Socii elongate setaceous ridges. Vinculum narrowly U-shaped. Tegumen shorter than

harpe, dilated laterally. Anellus a narrow sclerotized

plate, expanded basally and with a curved spinous

process from each corner posteriorly. Aedeagus rather

slender, strongly bent, apex linguiform; vesica with

small granular patches.

Female genitalia slide JFGC 12033. Ostium transverse, broad, ventral lip setaceous. Genital plate

sclerotized, with median granulations; lamella postvaginalis finely granular. Antrum sclerotized. Inception of ductus seminalis dorsal, at junction of antrum and membranous portion of ductus bursae.

Bursa copulatrix membranous. Signum a large, longitudinal, diamond-shaped dentate plate in posterior

half of bursa copulatrix.

HOLOTYPE.—USNM 71090.

TYPE LOCALITY.—Argentina, Tucuman, Ciudad

Universitaria, Cerro San Javier, 800 m.

DISTRIBUTION.—Argentina.

Described from the cf holotype (17. II. 1959, J. F.

G. Clarke) 3 cf cf and one 9 paratypes from the

type locality (17-20. II. 1959, Clarke).

In wing shape, with costa and dorsal margin

nearly parallel, amauroptera is similar to notodontella,

the type of this genus. In genitalia, however, it is as

near lecithitis as anything else but the vesica lacks

the strong cornutus of that species and signum is

strongly developed in amauroptera.

Among the five specimens before me no two are

alike. One specimen is indeed similar in coloring to

notodontella, the holotype shows a dark tornal blotch,

another is spotted, and two show considerable yellow

coloring.

Gonionota gaiophanes, new species

FIGURE 12; PLATE 2C

Alar expanse 21 mm.

Labial palpus buff mixed with gray; second segment brownish posteriorly. Antenna brown; scape

buff with some brown scaling anteriorly. Head buff

suffused clay color. Thorax clay color, scales buff

tipped. Forewing ground color clay color, strongly

overlaid sayal brown and weakly mottled with fuscuous; in tornal area an ill-defined fuscous blotch;

on termen from apex to tornus, six fuscous trans-
14 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

verse dashes; from basal third to apical third costa

narrowly yellowish; cilia concolorous with wing; underside of costal area strongly buff scaled, irrorate

with fuscuous and with two fuscous spots at two

fifths and middle. Hindwing pale clay color with

conspicuous fuscous shade between 2nd and 3rd

anals. Foreleg buff shaded clay color on outer side;

midleg similar to foreleg but more lightly shaded;

hindleg similar to midleg. Abdomen buff suffused

clay color ventrally.

Female genitalia slide JFGC 12082. Ostium transverse, crescentic. Genital plate subrectangular; lamella antevaginalis very finely scobinate. Antrum

sclerotized. Inception of ductus seminalis slightly anterior to antrum. Ductus bursae membranous. Bursa

copulatrix membranous. Signum small, cruciform,

each lateral element with 4 or 5 teeth.

HOLOTYPE.—USNM 71091.

TYPE LOCALITY.—Brazil, New Bremen.

DISTRIBUTION.—Brazil.

Described from the unique $ holotype (7. II.

1936, Fritz Plaumann).

In superficial appearance gaiophanes is similar to

aethoptera but lacks the dark dorsal band of the

FIGURE 11.—Gonionota amauroptera, new species, a, ventral view of male genitalia with left harpe and aedeagus

removed; b, aedeagus; c, ventral view of female genitalia.
NUMBER 95 15

latter species. The small but conspicuous white spot

at the end of cell in aethoptera is absent in gaiophanes.

Gonionota hemiglypta, new species

FIGURE 13; PLATE 2d

Alar expanse 17-21 mm.

Labial palpus ferruginous; second segment fuscous

dorsally and apically; third segment fuscous at middle with carmine scales at base and preapically; apex

yellowish. Antennal ferruginous basally, shading to

brown toward apex; segments each with tiny fuscous

spot. Head yellow with ferruginous, carmine-tipped

spreading scales laterally. Thorax yellow mixed with

carmine; tegula ferruginous, tipped with yellow and

carmine. Forewing ground color yellow with dorsal

half strongly overlaid carmine; costal half ferruginous

at base, this color becoming attenuated toward apex;

extreme costal edge narrowly whitish; at basal third,

in cell, a small black dot followed at end of cell by

a similar dot; termen and cilia, from vein 7 to tornus,

ferruginous. Hindwing ochraceous tawny; cilia yellowish white with ochraceous tawny suffusion. Foreleg ocherous white; femur overlaid ferruginous on

outer side; tibia carmine shaded; midleg ocherous

white; hindleg ocherous white with small ill-defined

fuscous spot at middle and another near distal end of

tibia. Abdomen grayish dorsoanteriorly, ferruginous

posterodorsally; ocherous white ventrally, sparsely

irrorate with black laterally.

Male genitalia slide JFGC 10820. Harpe narrowest

at base, dilated at middle, terminating in a broadly

rounded cucullus; from inside middle of costa, a conspicuous cluster of fine setae; clasper short, terminating in ventrally directed spines; from basal third a

stout curved spine; sacculus with small flap at middle. Gnathos an oval spined knob. Uncus broad

basally with widely separated curved lateral arms.

Vinculum a narrow band. Tegumen about as long as

broad. Anellus an elongate plate with deep excavation on each side and terminating in a pair of widely

divergent slender spines. Aedeagus moderately

slender, curved, pointed.

Female genitalia slide JFGC 10821. Ostium broad,

transverse, oval, with crenulate ventral edge. Genital

plate a narrow sclerotized band. Antrum sclerotized.

Inception of ductus seminalis at anterior edge of

antrum. Ductus bursae membranous. Bursa copulam

12

FIGURE 12.—Gonionota gaiophanes, new species. Ventral

view of female genitalia.
16 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

trix membranous. Signum a small, quadrate, dentate

plate.

HOLOTYPE.—USNM 71092.

TYPE LOCALITY.—Brazil, Nova Teutonia.

DISTRIBUTION.—Brazil.

Described from the cf holotype and one $ paratype .(IV. '48, Fritz Plaumann).

This species is similar to G. vivida (Meyrick) from

Bolivia but can be distinguished easily by the widely

separated arms of the uncus. In vivida the uncus

arises as a broadly based stalk, divided apically as in

many other members of Gonionota.

Gonionota eretnia, new species

FIGURE 14; PLATE 2e

Alar expanse 16-19 mm.

Labial palpus auburn with dark suffusion at base

of second segment. Antenna light auburn; scape auburn. Head auburn, scales tipped pale cinereous.

Thorax auburn, tegula tipped grayish. Forewing

ground color auburn; at apical third of costa a small

whitish streak; at basal third, in cell, a minute fuscous spot; at end of cell a pale, buff spot; between

veins 11 and 12 opposite base of 11, a minute fuscous

FIGURE 13.—Gonionota hemiglypta, new species, a, ventral

view of male genitalia with left harpe and aedeagus removed; b, aedeagus; c, ventral view of female genitalia.

spot; dorsum narrowly fuscous; cilia grayish auburn.

Hindwing fuscous, the scales loosely attached; cilia

grayish fuscous with pale basal line. Foreleg white;

femur and tibia auburn on outer side; tarsal segments suffused fuscous; mid- and hindlegs white suf-
NUMBE

R 95

17

fused fuscous; abdomen grayish fuscous above, white

ventrally.

Male genitalia slide JFGC 5091. Harpe less than

twice as long as broad, deeply hollowed to accom-

modate a stout, elongate, spined clasper; sacculus

broadly sclerotized; cuculus broad, blunt. Gnathos a

transverse, spined oval knob. Uncus very broad, deeply

cleft on posteromedian line; socii small setose papillae.

Vinculum a narrow band. Tegumen broad, rectan-

gular, not sharply differentiated from uncus. Anellus

a broad plate basally, constricted posteriorly, and

terminating in two slender lateral processes. Aedeagus

moderately short, slightly curved, simple.

Female genitalia slide JFGC 5092. Ostium cup-

shape, ventral side sclerotized. Genital plate cordi-

form, sclerotized. Antrum broadly sclerotized. Incep-

tion of ductus seminalis at anterior edge of antrum.

Ductus bursae membranous. Bursa copulatrix mem-

branous. Signum an oval dentate plate.

HOLOTYPE.—USNM 71093.

TYPE LOCALITY.—French Guiana, Cayenne.

DISTRIBUTION.—French Guiana, British Guiana.

Described from the cf holotype (no date; Wm.

Schaus) and one $ paratype, British Guiana, Bar-

FIGURE 14.—Gonionota eremia, new species, a, ventral view

of male genitalia with left harpe and aedeagus removed;

b, aedeagus; c, ventral view of female genitalia.
18 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

tica District, Kalacoon (20. X. 1920; no collector

indicated).

In genitalia eremia is near G. teganitis Meyrick but

differs widely in the nature of the clasper which is

short in teganitis and nearly as long as the harpe in

eremia. The presence of the dark dorsal edge places

eremia near aethoptera and aethographa, both described herein.

Gonionota dryodesma (Meyrick)

Hypercallia dryodesma Meyrick, 1916: (1)552.

Gonionota dryodesma (Meyrick), Clarke 1963: (4)230, pi.

112: figs. 4, 4a.

Hypercallia dryocrypta Meyrick, 1931: (4) 121. (New synonymy).

In Meyrick's description of the forewing of dryodesma he states "8 and 9 out of 7"; but in the case

of dryocrypta he wrote "9 out of 7." The types of

both have identical venation with 9 out of the stalk

of 7 and 8. A male from Venezuela matches the type

male of dryodesma and a female, collected in Venezuela at the same time as the male, matches the type

of dryocrypta.

In addition to the types, I have before me a o*

and $ , Venezuela, Aragua, Rancho Grande, 1100 m

(21-25. Jan. '66, S. S. and W. D. Duckworth); ? ,

Costa Rica, Turrialba (VII. 15-19.65, P. J. Spangler).

TYPES.—British Museum (Natural History) (dryodesma) ; U. S. National Museum (dryocrypta).

TYPE LOCALITIES.—French Guiana, St. Jean, Rio

Maroni (dryodesma) ; Guatemala, Cayuga (dryocrypta) .

DISTRIBUTION.—French Guiana, Venezuela, Guatemala, Costa Rica.

Gonionota insignata, new species

FIGURE 15; PLATE 2/

Alar expanse 16-18 mm.

Labial palpus ocherous white on inner side with

slight reddish suffusion and fuscous irrorations; remainder suffused reddish and profusely irrorate with

fuscous and gray; posterior tuft of second segment

mixed with much gray. Antenna gray with minute

fuscous spots at ends of segments; scape suffused reddish. Head grayish ocherous. Thorax dull ochraceous

orange, paler posteriorly. Forewing ground color

ochraceous orange irregularly blotched with grayish

fuscous, especially on dorsum; at basal third, in cell,

a fuscous spot, largely obscured by the irregular grayish fuscous blotching; at end of cell a white discal

spot; subterminally, from inside apex to tornus, a

series of 7 or 8 small fuscous spots; costa with some

scattered white scales and at apical third a pale,

wedge-shaped mark preceded by an ill-defined fuscous spot; cilia grayish fuscous mixed with some

white-tipped scales.

Hindwing ocherous white, slightly darker apically;

cilia ocherous white becoming darker toward apex.

Foreleg ocherous white, strongly marked reddish and

fuscous on outer side; midleg ocherous white with a

few grayish spots on outer side; hindleg ocherous

white with sparse grayish fuscous irroration. Abdomen sordid white infuscated dorsally; ventrally white

but tawny posteriorly.

Male genitalia slides JFGC 5093, 5094. Harpe

short, broadest at middle by a pronounced expansion

of sacculus; from base of sacculus to near middle a

low, free ridge; at junction of sacculus and a bar from

base of costa, a small knob armed with a cluster of

stout recurved setae. Gnathos an oval spined knob.

Uncus broad basally, somewhat narrowed at middle,

terminating in a flat knob. Vinculum a narrow ring.

Tegumen about as long as harpe, widest at base.

Anellus a sclerotized plate, widest above base, posterior edge deeply concave with a long, slender spine

at each corner. Aedeagus curved, moderately stout;

distal end spatulate.

HOLOTYPE.—USNM 71094.

TYPE LOCALITY.—Ecuador, Environs de Loja.

DISTRIBUTION.—Ecuador.

Described from the <$ holotype and two cf cf

paratypes from the same locality ("10-86, Dognin

Collection").

In general aspect insignata is similar to ioleuca but

the ground color of the latter is much grayer than

that of the former.

This is a variable species with the blotched effect

producing an irregular, broad, median area in the

holotype, absent in the paratypes, but one of them

with the blotched dorsal half of forewing.

Gonionota menura, new species

FIGURE 16; PLATE 2g

Alar expanse 24 mm.

Labial palpus ocherous white, heavily overlaid
PLATES
20

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

PLATE 1.—a, Gonionota aethoptera, new species; b, G. aethographa, new species; c, G. cologramma, new species; d, G. autocrena (Meyrick) ; e, G. oxybela, new species; /, G. charagma,

new species; g, G. hypoleuca, new species; h, G. argopleura, new species.
NUMBER 95

21

PLATE 2.—a, Gonionota phthiochroma, new species; b, G. amauroptera, new species; c, G.

gaiophanes, new species; d, G. hemiglypta, new species; e, G. eremia, new species; /, G. insignata, new species; g, G. menura, new species; h, G. parauexillata, new species.
22

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

J

PLATE 3.—a, Gonionota transuersa, new species; b, G. sphenogramma, new species; c, G. selene, new species; d, G. leucoporpa (Meyrick) ; e, G. lecithitis (Meyrick) ; /, G. poecilia, new

species; g, G. rosacea (Forbes) ; h, Hypercallia heterochroma, new species.
NUMBER 95 23

with amber brown and carmine scales; third segment

with apex broadly ocherous white. Antenna light

amber brown with darker annulations. Head ocherous white obscured by amber brown and carmine.

Thorax amber brown, spotted faintly with carmine.

Forewing ground color amber brown; basal third

faintly blotched with carmine; slightly before middle

of costa a small, ocherous white, quadrate spot edged

with carmine; at outer third costa excavated and

bordered with a white, carmine-edged, lunate mark;

remainder of costa very narrowly edged fuscous; in

cell at basal third a fuscous spot surrounded by a few

carmine scales; discal spot at end of cell obsolete;

between end of cell and termen an ill-defined, transverse row of fuscous spots; cilia amber brown.

Hindwing ocherous white basally, shading to

brown at apex; cilia matching. Foreleg ocherous

white overlaid with amber brown and carmine on

outer side; midleg similar; hindleg ocherous white

with scattered amber brown and carmine irrorations;

spurs brown tipped. Abdomen ocherous white, ventrally irrorate with tawny, fuscous, and brown.

Female genitalia slide JFGG 12126. Ostium wide,

broadly funnel shaped. Lamella postvaginalis granular. Antrum not differentiated. Inception of ductus

seminalis lateral, from a small membranous area at

posterior two thirds of ductus bursae. Ductus bursae

strongly sclerotized for most of its length with an

expanded area anteriorly on left side. Bursa copulatrix membranous. Signum an elongate oval, sclerotized, dentate plate.

HOLOTYPE.—USNM 71095.

TYPE LOCALITY.—Panama, Cerro Campana, nr.

Chica.

DISTRIBUTION.—Panama.

Described from the $ holotype (2-5. IV. 65, S. S.

and W. D. Duckworth).

In coloring and pattern menura is similar to isastra and insulana but isastra has a much grayer

hindwing; insulana is a smaller insect than menura

and has a proportionately narrower forewing. The

heavily sclerotized ductus bursae of menura further

distinguishes it from insulana.

Gonionota paravexillata, new species

FIGURE 17; PLATE 2h

Alar expanse 17 mm.

Labial palpus ocherous white; second segment

heavily overlaid red brown and with carmine suffusion on inner side; second segment similar with few

black scales in tuft, and apex buff. Antenna fuscous,

somewhat paler and suffused reddish basally; scape

ocherous white ventrally, reddish brown dorsally.

FIGURE 15.—Gonionota insignata, new species, a, ventral

view of male genitalia with left harpe and aedeagus removed; b, aedeagus.
24 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Head a mixture of reddish and yellowish whitetipped scales. Thorax a mixture of snuff brown,

orange, and scarlet; base of tegula snuff brown. Forewing ground color snuff brown; from basal third

of costa a broad scarlet fascia curves inwardly and

broadens to base of wing; the scarlet band is mixed

with orange and a few sepia scales across fold; the

fascia is bordered outwardly by an ill-defined sepia

line; on costa, just beyond apical third, a sharply defined, small white triangle from which arises an illdefined, outwardly curved, narrow, sepia fascia which

extends to dorsum at tornus; at end of cell a small

white dot preceded and followed by a few fuscous

scales; dorsum narrowly edged fuscous; cilia sepia.

Hindwing snuff brown, slightly darker toward apex,

buff between cell and costa; cilia sepia with grayish

tips. Foreleg ocherous white; femur grayish fuscous

on outer side; tibia snuff brown with reddish hue on

outer side; first tarsal segment with scarlet dash; midleg ocherous white; tibia grayish fuscous on outer

side; hindleg ocherous white with faint grayish suffusion on tibia and tarsal segments. Abdomen sepia

dorsally, light buff ventrally.

Male genitalia slide JFGC 12140. Harpe broadest

about middle; cucullus rounded; clasper with distal,

free, spined arm extending to edge of cucullus; sacculus recurved ventrally slightly beyond middle.

Gnathos a transverse, spined oval knob. Uncus a

long, slender stalk with widely divergent terminal

arms. Vinculum U-shaped. Tegumen as long as

harpe, moderately narrow, slightly dilated laterally.

Anellus a rectangular plate with long spinous process

from each corner posteriorly. Aedeagus stout, curved,

distally an upturned point; vesica armed with a cluster of stout cornuti.

HOLOTYPE.—USNM 71096.

TYPE LOCALITY.—Venezuela, Aragua, Rancho

Grande, 1100 m.

DISTRIBUTION.—Venezuela.

Described from the unique o* holotype (21-25

Jan. 1966. S. S. and W. D. Duckworth).

As the name suggests, this species is very similar

to vexillata but may be distinguished readily by the

genitalia. The terminal spines of the clasper of vexillata are directed inward and toward sacculus but in

paravexillata they are directed toward cucullus. The

cornuti of vexillata consist of two groups in linear arrangement but in paravexillata there is one large

cluster.

16

FIGURE 16.—Gonionota menura, new species. Ventral view

of female genitalia.
NUMBER 95 25

Gonionota transversa, new species

FIGURE 18; PLATE 3a

Alar expanse 17-19 mm.

Labial palpus buff; second segment irrorate with

tawny and fuscous; posterior tuft fuscous; third segment with a broad fuscous median annulus. Antenna

dresden brown with paler annulations; scape dresden

brown dorsally, buff ventrally. Head buff; frons and

laterally dresden brown; dorsally mixed with scattered ochraceous-orange scales. Thorax buff mixed

with ochraceous-orange and dresden brown. Forewing ground color dresden brown; basal third of

wing, except rectangular patch of ground color on

costa, buff, mixed with ochraceous orange and fuscous; outer third of wing buff but this color nearly

obscured by darker blotches and irrorations; on

costa, at apical third, a buff spot; in cell, at one third,

a pair of small fuscous discal spots; at end of cell a

similar, but smaller, spot; cilia dresden brown. Hindwing ocherous white, slightly darker toward margins;

cilia ocherous white at anal angle, becoming darker

toward apex. Foreleg ocherous white; femur and

tibia fuscous on outer side; tarsal segments suffused

ochraceous orange; midleg ocherous white with small

fuscous spot on tibia; hindleg ocherous white with a

few scattered grayish scales. Abdomen grayish dorsally; ventrally buff.

Male genitalia slides JFGC 4631, 12131. Harpe of

nearly equal width throughout; cucullus broadly

rounded; clasper terminating in a small cluster of

stout spines (one example with one large and two

small ones). Gnathos a pear-shaped spined knob.

Uncus a broadly based curved stalk, distally bilobed.

Vinculum U-shaped. Tegumen arched, shorter than

harpe. Anellus a subrectangular, sclerotized plate; at

each corner posteriorly a flattened projection. Aedeagus stout, curved, pointed distally; vesica armed with

one large cluster of stout cornuti and two groups of

granulations.

HOLOTYPE.—USNM 71097.

TYPE LOCALITY.—Brazil, Castro Parana.

DISTRIBUTION.—Brazil.

Described from the o* holotype and 3 cf o* paratypes with identical data (no date; Wm. Schaus

Coll.).

Although transversa is similar to ioleuca, and is

from the same locality, the former can be distinguished immediately from the latter by the genitalia.

In ioleuca the spined part of the clasper extends to

the end of cucullus but in transversa the clasper is

weak and has only a few curved spines about middle

of harpe. Moreover the vesica of transversa is armed

with a large cluster of stout cornuti, a feature absent

in ioleuca.

Gonionota sphenogramma, new species

FIGURE 19; PLATE 36

Alar expanse 17-18 mm.

Labial palpus brown; second segment grayish

posteriorly; third segment apex yellowish. Antenna

FIGURE 17.—Gonionota paravexillata, new species, a, ventral view of male genitalia with left harpe and aedeagus

removed; b, aedeagus.
26 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

18c

FIGURE 18.—Gonionota transversa, new species, a, ventral view of male genitalia with left harpe

and aedeagus removed; b, aedeagus; c, variation in clasper.

brown, faintly and narrowly annulated fuscous. Head

light brown, scales tipped grayish. Thorax brown and

gray mixed. Forewing ground color brown with pronounced gray dusting, particularly in basal half; at

basal two fifths of costa a small white spot preceded

by dark gray; basal third of costa yellowish; costa

at middle, dark gray, recurved, and followed outwardly by a shallow excavation, the latter white,

broadened outwardly into a wedge-shaped mark,

edged yellowish; on costa, preceding apex, a slender

white streak; discal spot at basal third dark gray, at

end of cell white; tornus dusted dark gray; cilia concolorous except dark gray from vein 5 to tornus.

Hindwing light gray suffused brownish toward margins; cilia pale brownish buff basally, sordid white

apically, and with a gray subbasal line. Foreleg

ocherous white; femur brown on outer side; tibia

brown on outer side with carmine apically; 1st tarsal

segment carmine on outer side; midleg similar to

foreleg but with two brownish spots on tibia; hindleg ocherous with ill-defined, small brownish markings. Abdomen ocherous white, infused brownish

dorsally; irrorate with scattered brown scales laterally.

Male genitalia slide JFGC 12083. Harpe broadest

before middle, then tapered to the bluntly pointed

cucullus; from base of costa a slender strongly sclerotized bar to middle of harpe continued as a long

free arm nearly to cucullus, bulbous at base, and

armed with stout setae; sacculus swollen and with

a prominent ridge in distal half. Gnathos a transverse, oval spined knob. Uncus broad, divided, and

setose distally. Socii low, setose prominences. Vinculum a narrow band. Tegumen moderately broad,

narrowed posteriorly. Anellus a pear-shaped plate

with posterolateral hooks. Aedeagus stout, curved,

sharply pointed.

Female genitalia slide JFGC 12084. Ostium transverse, slitlike. Genital plate subcordiform, rugose,

strongly setose. Antrum broadly sclerotized. Inception of ductus seminalis slightly anterior to antrum.

Ductus bursae membranous. Bursa copulatrix membranous. Signum a longitudinal, oval, dentate plate.

HOLOTYPE.—USNM 71098.

TYPE LOCALITY.—Venezuela, Aragua, Rancho

Grande, 1100 m.

DISTRIBUTION.—Venezuela.
NUMBE

R 95

27

Described from the d

1

 holotype (16-19. I. 66, S. S.

and W. D. Duckworth), 4 d d (11-25. I. 1966),

and one $ paratypes (31. X. 1966) all collected by

S. S. and W. D. Duckworth.

This species is very near G. anelicta (Meyrick)

from Bolivia, but has a much paler hindwing. Un-

fortunately, the type of anelicta has no abdomen

(Clarke 1963:229, pi. Ill: fig. 3), so no comparison

of the genitalia can be made.

Gonionota selene, new species

FIGURE 20; PLATE 3C

Alar expanse 20 mm.

Labial palpus gray brown; second segment with

some ocherous white on inner side; third segment

apex ocherous white. Antenna gray but browner

toward base and ocherous white ventrally; scape

brownish dorsally, ocherous white ventrally. Head

gray brown, scales tipped whitish. Thorax light

chrome yellow with median longitudinal brown line;

tegula brown anteriorly. Forewing ground color rus-

set; at base of dorsum a light chrome yellow blotch

extending across fold and bordered on outer edge

FIGURE 19.—Gonionota sphenogramma, new species, a, ven-

tral view of male genitalia with left harpe and aedeagus re-

moved; b, aedeagus; c, ventral view of female genitalia.
28 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

with a short fuscous dash; costa gray brown except

at two thirds a white, wedge-shaped mark; at two

fifths of costa a short chrome yellow transverse fascia

not reaching middle of wing; at end of cell a small

fuscous dot; dorsum, beyond the chrome yellow basal

blotch, broadly fuscous to termen at vein 3; dark

dorsal area overlaid avellaneous in central part; from

apex to vein 5 an ill-defined, quadrate chrome yellow

area; cilia russet. Hindwing whitish ocherous, basally

shading to pale brown at apex and grayish fuscous in

anal area; cilia sordid white mixed grayish brown.

Foreleg ocherous white; femur gray brown on outer

side; tibia russet on outer side; midleg similar but

lighter and speckled; hindleg whitish ocherous; tibia

and tarsal segments speckled brown and gray. Abdomen ocherous white with a fewr

 rust-colored scales

mixed; ventrally finely irrorate with fuscous.

Female genitalia slide JFGC 10935. Ostium crescent shaped; posteroventral edge convex. Genital

plate rather narrow, U-shaped; lamella postvaginalis

membranous. Antrum subrectangular, sclerotized.

Inception of ductus seminalis slightly anterior to antrum. Ductus bursae finely granular, moderately

short. Bursa copulatrix membranous. Signum a large

diamond-shaped, dentate, sclerotized plate.

HOLOTYPE.—USNM 71099.

TYPE LOCALITY.—Brazil, Santa Catharina.

DISTRIBUTION.—Brazil.

Described from the unique $. holotype (21 Jan.

'36, Fritz Hoffmann).

Superficially selene resembles incalescens but where

the wedge-shaped white costal mark is situated there

is a deep excavation of costa in selene which is absent in incalescens. The outer discal spot is white

in incalescens but fuscous in selene and the prominent crimson coloring of incalescens is absent in

selene. Although the detailed differences seem great,

the moths look strikingly similar to the naked eye.

Gonionota leucoporpa (Meyrick), new combination

FIOURE 21; PLATE 3d

Hypercallia leucoporpa Meyrick, 1926:313.—Clarke 1963:

293, pi. 143: fig. 4.

Male genitalia slide JFGC 12052. Harpe broadest

slightly before middle, abruptly narrowed to cucullus;

clasper terminating about middle of harpe in a cluster of fine spines. Gnathos a transverse, spined oval

knob. Uncus a broadly based rather wide stalk terFIGURE 20.—Gonionota selene, new species. Ventral view of

female genitalia.
NUMBE

R 95

minating in two blunt divergent points. Socii indi-

cated by setaceous, raised, broad ridges. Vinculum

U-shaped. Tegumen about two thirds the length of

harpe. Anellus a sclerotized oval plate with a pair

of divergent pointed processes posteriorly. Aedeagus

moderately slender, curved, distal end upturned;

cornutus a small sclerotized plate.

Female genitalia slide JFGC 12053. Ostium slitlike,

transverse. Genital plate sclerotized, produced an-

teriorly, narrowed laterally. Antrum sclerotized, short.

Inception of ductus seminalis at junction of antrum

and membranous portions of ductus bursae. Ductus

bursae membranous with minute granulations. Bursa

copulatrix membranous. Signum a very small, irregu-

lar sclerotized plate.

TYPE.—British Museum (Natural History).

TYPE LOCALITY.—Colombia, Sosomoko.

DISTRIBUTION.—Colombia, Venezuela.

As pointed out by Clarke (1963, p. 293) the ab-

domen of the type is missing. The genitalia of leucoporpa, therefore, have not been figured previously

FIGURE 21.—Gonionota leucoporpa (Meyrick). a, ventral

view of male genitalia with left harpe and aedeagus re-

moved; b, aedeagus; c, ventral view of female genitalia. 29

21c

i
30 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

and are figured here from specimens from Venezuela,

Aragua, Rancho Grande, 1100 m, 11-19. Jan. 1966,

S. S. and W. D. Duckworth.

Gonianota lecithitis (Meyrick)

FIGURE 22; PLATE 3e

Coptotelia lecithitis Meyrick, 1912:703.—Clarke 1963:238,

pi. 116: figs. 4-4b.

Male genitalia slides JFGC 4007, 12213. Harpe

broadest about middle; cucullus rounded; clasper

terminating in a cluster of curved spines. Gnathos

an oval spined knob. Uncus a broadly based stalk

dilated posteriorly. Socii, small, narrow setaceous

ridges. Vinculum U-shaped. Tegumen two thirds the

length of harpe. Anellus a broad plate basally, narrowed posteriorly, terminating in two divergent

points. Aedeagus moderately stout, curved distally

ending in a small peak; vesica armed with a strongly

barbed cornutus.

Female genitalia slide JFGC 4008. Ostium round,

ventral edge irregular. Genital plate narrow. Antrum

somewhat thickened. Inception of ductus seminalis

FIGURE 22.—Gonionota lecithitis (Meyrick). a, ventral view

of male genitalia with left harpe and aedeagus removed;

b, aedeagus; c, ventral view of female genitalia.

at anterior edge of antrum. Ductus bursae membranous, short. Bursa copulatrix membranous. Signum faintly indicated.

TYPE.—British Museum (Natural History).

TYPE LOCALITY.—Argentina, Parana.

DISTRIBUTION.—Argentina.

The illustration in Clarke (1963) is inadequate.

The cucullus appears pointed in the photograph and

is the result of pressure on the genitalia. Moreover

the barbed cornutus and the small peak at the distal

end of the aedeagus do not show in Figure 4a. The

female has not been illustrated previously and is in-
NUMBER 95 31

eluded for the sake of completeness. It is figured

from a specimen from Tucuman, Argentina.

Gonionota poecilia, new species

FIGURE 23; PLATE 3/

Alar expanse 16-19 mm.

Labial palpus light ochraceous buff; second segment irrorate with ochraceous tawny; posterior tuft

grayish fuscous; third segment similar but irrorate

with reddish scales and apex ocherous white. Antenna

buff with grayish fuscous annulations; scape grayish

fuscous dorsally, buff ventrally. Head light avellaneous. Thorax sayal brown mixed with scarlet and

orange yellow. Ground color of forewing sayal

brown; dorsal two thirds mostly scarlet and orangeyellow blotches and spots set off by a fuscous reticulum; on basal third of dorsum a triangular spot of

ground color, its base on dorsal edge; on basal third

of costa an orange yellow triangle containing scarlet

scales; a similarly colored wedge-shaped mark on

outer third of costa; at basal third, in cell, a large

white spot followed at end of cell by a similar, but

smaller, spot; subterminally an outwardly curved

row of fuscous dashes from vein 9 to vein lc; terminal edge from apex around tornus to base of dorsum narrowly fuscous; cilia sayal brown.

Hindwing ocherous white, basally shading to clay

color toward margins; termen narrowly dark clay

color; cilia ocherous white at anal angle becoming

darker toward apex. Foreleg light ochraceous buff;

femur and tibia sayal brown on outer side, mid- and

hindlegs light ochraceous buff; very sparsely irrorate

with brownish. Abdomen deep olive buff dorsally,

light ochraceous buff ventrally, irrorate with brown

and fuscous scales.

Male genitalia slide JFGC 12015. Harpe broadest before middle, gradually tapered to a narrow,

bluntly pointed cucullus; clasper a sclerotized rod

terminating in a free knob clothed with one large

and numerous small spines. Gnathos an oval spined

knob. Uncus broad, flattened, dilated distally; socii

short, digitate setaceous processes. Vinculum narrowly rounded. Tegumen about two thirds the

length of harpe, broad basally. Anellus pear shape;

from each corner, posteriorly, a long spine. Aedeagus strongly bent, sharply pointed distally; vesica

unarmed.

Female genitalia slide JFGC 12085. Ostium slitlike. Genital plate very broad, rounded anteriorly;

lamella antevaginalis and lamella postvaginalis membranous, granular. Antrum strongly sclerotized for a

short distance inside ostium. Inception of ductus

seminalis lateral, at junction of antrum and membranous part of ductus bursae. Ductus bursae membranous. Bursa copulatrix membranous, very finely

granular posteriorly. Signum an elongate, sclerotized,

dentate plate.

HOLOTVPE.—USNM 71101.

TYPE LOCALITY.—Venezuela, Aragua, Rancho

Grande, 1100 m.

DISTRIBUTION.—Venezuela.

Described from the o* holotype, 12 d o* and 7

9 9 para types, all from the same locality (January,

October, and November dates, 1966, S. S. and W.

D. Duckworth) and one 9 (22-30. VI. 1967, R. W.

Poole).

Superficially, poecilia resembles festicola more

than it does anything else presently assigned to the

genus, but is easily distinguished by the genitalia. In

festicola there is a long, pointed process, parallel to

costa, from base of harpe and a strongly recurved

clasper from basal third of harpe (Clarke 1963: pi.

114: fig. lb), both absent in poecilia. The vesica of

festicola is armed, that of poecilia unarmed. The

uncus is broad and flattened in poecilia, stalked in

festicola. The female genitalia of festicola are unknown, so cannot be compared with poecilia.

Gonionota rosacea (Forbes), new combination

FIGURE 24; PLATE 3g

Hypercallia rosacea Forbes, 1931:4 (4):379, pi. 42: fig. 18.

Male genitalia slides JFGC 12050, 12051. Harpe

oval, terminating in a bluntly pointed cucullus;

clasper expanded dorsally into a large, dentate disc

with a flattened curved process from base nearly

reaching ventral edge of harpe. Gnathos a transverse, elongate spined knob. Uncus a broadly based,

short stalk with short divergent arms apically. Socii

indicated mainly by a few setae. Vinculum Ushaped. Tegumen about two thirds the length of

harpe, convex laterally. Anellus broad basally, narrowed, and with a long spine from each side posteriorly. Aedeagus slightly curved, apically digitate;

vesica armed with one beaklike curved cornutus

with a spined plate at base.
32 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

TYPE.—Cornell University, Ithaca, New York.

TYPE LOCALITY.—Haiti.

DISTRIBUTION.—Cuba, Haiti, Dominican Republic.

There are two specimens before me from the

Dominican Republic (Jarabacoa, 3-4 June 1969

and Convento, 12 km. S. of Contanza, 6-13 June

1969, both collected by Flint and Gomez) which

extends the range of this beautiful species.

So far the species is known only from males.

The genitalia of rosacea have not been figured

previously. The enormously dilated clasper distinguishes rosacea from any other species of Gonionota

known to me.

Gonionota phthiochroma, new species

FIGURE 25; PLATE 2a

Alar expanse 23 mm.

Antenna very pale grayish, basally shading to

buff apically; dorsally, near base a slight carmine

suffusion; scape grayish fuscous dorsally, white ventrally. Head grayish fuscous, anteriorly shading

through ochraceous salmon to buff posteriorly.

Thorax ochraceous salmon anteriorly, ochraceous

tawny posteriorly; tegula ochraceous tawny anteriorly, ochraceous salmon and buff posteriorly. Forewing ground color costal two fifths ochraceous buff,

dorsal three fifths ochraceous salmon; extreme edge

of costa ocherous white; basal fifth of wing, inside

costa ochraceous salmon and ochraceous tawny; in

FIGURE 23.—Gonionota poecilia, new species, a, ventral

view of male genitalia with left harpe and aedeagus removed; b, aedeagus; c, ventral view of female genitalia.
NUMBER 95 33

FIGURE 24.—Gonionota rosacea (Forbes), a, ventral view of

male genitalia with left harpe and aedeagus removed; b,

aedeagus.

FIGURE 25.—Gonionota phthiochroma, new species, a, ventral view of male genitalia with left

harpe and aedeagus removed; b, aedeagus.
34 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

cell, at basal third a minute black spot and a similar

one at end of cell; outer half of wing with a few

scattered black scales; cilia ochraceous tawny at

apex, shading to ochraceous salmon and buff around

termen to tornus; costal two fifths, underside,

heavily suffused fuscous. Hindwing ocherous white,

slightly darker and with a pink tinge apically; cilia

very pale brownish at apex, shading to ocherous

white around termen to base; underside sparsely irrorate fuscous and with a series of conspicuous fuscous spots around termen. Foreleg ocherous white;

outer side of femur and tibia ochraceous tawny;

mid- and hindlegs ocherous white with sparse fuscous irroration. Abdomen ocherous white, dorsoposteriorly shading to buff; a pair of brownish spots

ventroanteriorly and black spots posterolaterally.

Male genitalia slide JFGC 12212. Harpe twice

as long as broad; cucullus rounded; clasper stout,

terminating in three large and three small recurved

spines. Gnathos a transverse oval spined knob. Uncus

a broadly based stalk with short divergent arms

apically. Socii small, setaceous papillae. Vinculum

broadly U-shaped. Tegumen about two thirds the

length of harpe, dilated basally. Anellus an elongate

sclerotized plate, broadest basally, narrowed distally.

Aedeagus moderately slender, curved, distal end

linguiform; vesica unarmed except for a few

granulations.

HOLOTYPE.—USNM 71102.

TYPE LOCALITY.—Bolivia, Cochabamba, Incahaca, Tropical cloud area, 2100 m.

DISTRIBUTION.—Bolivia.

Described from the unique cf holotpe (27-VIII5. IX. 56, L. Pefia).

The nearest relative of phthiochroma is isophylla

Meyrick from Peru but phthiochroma lacks any

trace of the "reddish gray" hindwing described for

isophylla. Moreover the fuscous suffusion of the

costal two fifths of the forewing of phthiochroma is

absent in isophylla. The male genitalia of the two

are strikingly similar, showing the obvious close relationship, but the curved spines at the end of clasper of isophylla are longer than those of phthiochroma.

The following species, including those described

in the foregoing pages, are now assigned to the

Genus Gonionota:

acrocosma (Meyrick) 1912, Transactions of the Entomological Society of London, 1911:703—COLOMBIA

aethographa, new species—COSTA RICA

aethoptera, new species—VENEZUELA

amauroptera, new species—ARGENTINA

amphicrena (Meyrick) 1912, Transactions of the Entomological Society of London, 1911:703—COLOMBIA

anelicta (Meyrick) 1926, Exotic Microlepidoptera, 3:312—

BOLIVIA

anisodes (Meyrick) 1916, Exotic Microlepidoptera, 1:

552—FRENCH GUIANA

argopleura, new species—BRAZIL

autocrena (Meyrick) 1930, Exotic Microlepidoptera, 3:

575—BRAZIL

bourquini Clarke 1964, Proceedings of the United States

National Museum, 115(3480): 70, fig. 4, pi. 2: figs. 4,

5—ARGENTINA, BRAZIL

borquiniella (Kohler) 1940, An. Soc. Cient. Argentina, 128:

369, fig. 1, new combination—ARGENTINA

captans (Meyrick) 1931, Exotic Microlepidoptera, 4:121,

new combination—BRAZIL

charagma, new species—BRAZIL

citronota (Meyrick) 1932, Exotic Microlepidoptera, 4:279,

new combination—BRAZIL

cologramma, new species—VENEZUELA

comastis Meyrick 1909, Transactions of the Entomological

Society of London, 1909:25—PERU

constellata (Meyrick) 1912, Transactions of the Entomological Society of London, 1911:701—COLOMBIA

contrasta Clarke 1964, Proceedings of the United States

National Museum, 115(3480):64, figs. 2a-d; pi. 1:

fig. 4; 2: fig. 2—PERU

cristata Walsingham 1912, in Godman and Salvin, Biologia

Centrali-Americana, 42 (Lepidoptera-Heterocera, 4) :

129—PANAMA

determinata Clarke 1964, Proceedings of the United States

National Museum, 115(3480):68, fig. 3; pi. 1: fig. 5;

2: fig. 3—BRITISH GUIANA, VENEZUELA

dissita Clarke 1964, Proceedings of the United States National Museum, 115(3480) :67, fig. 2e; pi. 3: fig. 5—

TRINIDAD

dryodesma (Meyrick) 1916, Exotic Microlepidoptera, 1:

552—FRENCH GUIANA

[=dryocrypta (Meyrick)]—GUATEMALA

eremia, new species—FRENCH GUIANA

erotopis (Meyrick) 1926, Exotic Microlepidoptera, 3:

315—BOLIVIA

erythroleuca (Meyrick) 1928, Exotic Microlepidoptera, 3:

473—PERU

eurydryas (Meyrick) 1926, Exotic Microlepidoptera, 3:

314—COLOMBIA

euthyrsa (Meyrick) 1930, Exotic Microlepidoptera, 3:

577—ECUADOR

excavata Clarke 1964, Proceedings of the United States

National Museum, 115(3480): 79, fig. 6e; pi. 1: fig. 2;

3: fig. 3—MEXICO
NUMBER 95 35

extima Clarke 1964, Proceedings of the United States National Museum, 115(3480): 76, fig. 5b, pi. 1: fig. 1; 3:

fig. 2—COSTA RICA

festicola (Meyrick) 1924, Exotic Microlepidoptera, 3:104—

PERU

ftmbriata Clarke 1964, Proceedings of the United States

National Museum, 115(3480):82, figs. 7c-e; pi. 1: fig.

3; 3: fig. 1—PANAMA

gaiophanes, new species—BRAZIL

habristis (Meyrick) 1914, Exotic Microlepidoptera, 1:189—

BRITISH GUIANA

hemiglyptal new species—BRAZIL

hydrogramma (Meyrick) 1912, Transactions of the Entomological Society of London, 1911:702—COLOMBIA

hypoleuca, new species—VENEZUELA

hyptiotes Clarke 1964, Proceedings of the United States

National Museum, 115(3480) : 80, fig. 7f; pi. 3: fig. 6—

MEXICO

incalescens (Meyrick) 1914, Exotic Microlepidoptera, 1:

192—COLOMBIA

incisa Meyrick 1909, Transactions of the Entomological Society of London, 1909:27—BOLIVIA

incontigua Clarke 1964, Proceedings of the United States

National Museum, 115(3480) :74, fig. 5a; pi. 2: fig. 6—

VENEZUELA

insignata, new species—ECUADOR

insulana Clarke 1968, Proceedings of the United States National Museum, 125(3654) :5, fig. 3, pi. 2—DOMINICA

intonans (Meyrick) 1933, Exotic Microlepidoptera, 4:368—

ARGENTINA

ioleuca (Meyrick) 1912, Transactions of the Entomological

Society of London, 1911: 703—ARGENTINA

isastra (Meyrick) 1926, Exotic Microlepidoptera, 3:312—

COLOMBIA

isodryas (Meyrick) 1921, Exotic Microlepidoptera, 2:389—

BRAZIL

isophylla Meyrick 1909, Transactions of the Entomological

Society of London, 1909:25—PERU

lecithitis (Meyrick) 1912, Transactions of the Entomological Society of London, 1911:703—BRAZIL

leucoporpa (Meyrick) 1926, Exotic Microlepidoptera, 3:

313—COLOMBIA

lichenista (Meyrick) 1926, Exotic Microlepidoptera, 3:

312—COLOMBIA

melobaphes Walsingham 1912, in Godman and Salvin, Biologia Centralia-Americana, 42

(Lepidoptera-Heterocera, 4): 129, pi. 4: fig. 27—COSTA

RICA, PANAMA

menura, new species—PANAMA

militaris (Meyrick) 1914, Exotic Microlepidoptera, 1:191—

COLOMBIA

mitts (Meyrick) 1914, Exotic Microlepidoptera, 1:190—

PERU

notodontella Zeller 1877, Horae Societatis Entomologicae

Rossicae, 13:381, pi. 5: fig. 132—COLOMBIA

oligarcha (Meyrick) 1913, Transactions of the Entomological Society of London, 1913:178—PERU

oriphanta (Meyrick) 1928, Exotic Microlepidoptera, 3:

473—COLOMBIA

oxybela, new species—PERU

paravexillata, new species—VENEZUELA

periphereia Clarke 1964, Proceedings of the United States

National Museum, 115(3480) :77, figs. 6c, d; pi. 3: fig.

4—ECUADOR

persistis (Meyrick) 1914, Exotic Microlepidoptera, 1:191—

PERU

phocodes Meyrick 1909, Transactions of the Entomological

Society of London, 1909:27—PERU

phthiochroma, new species—BOLIVIA

pialea (Meyrick) 1921, Exotic Microlepidoptera, 2:389—

BRAZIL

poecilia, new species—VENEZUELA

praeclivis (Meyrick) 1921, Exotic Microlepidoptera, 2:

389—PERU

prolectans (Meyrick) 1926, Exotic Microlepidoptera, 3:

314—COLOMBIA

pyrocausta (Meyrick) 1931, Exotic Microlepidoptera, 4:

121—COLOMBIA

pyrrhotrota (Meyrick) 1932, Exotic Microlepidoptera 4:

280, new combination—BOLIVIA

rhacina Walsingham 1912, in Godman and Salvin, Biologia

Centrali-Americana, 42 (Lepidoptera-Heterocera 4):

130, pi. 4: fig. 29—GUATEMALA

rosacea (Forbes) 1931, Journal of the Department of Agriculture of Porto Rico, 4(4): 379, pi. 42: fig. 18—CUBA,

HAITI, DOMINICAN REPUBLIC

satrapis (Meyrick) 1914, Exotic Microlepidoptera, 1:191—

COLOMBIA

saulopis Meyrick 1909, Transactions of the Entomological

Society of London, 1909:28—PERU

selene, new species—BRAZIL

sphenogramma, new species—VENEZUELA

teganitis Meyrick 1909, Transactions of the Entomological

Society of London, 1909:26—PERU

transversa, new species—BRAZIL

uberrima (Meyrick) 1914, Exotic Microlepidoptera, 1:

190—PERU

ustimacula (Zeller) 1875, Verhandlungen der K.K. Zoologisch-botanischen Gesellschaft in WIEN, 24:440, pi. 12:

fig. 10 (=zelleri Butler)—CHILE

vexillata (Meyrick) 1913, Transactions of the Entomological

Society of London, 1913:179—PERU

vivida (Meyrick) 1924, Exotic Microlepidoptera, 3:103—

BOLIVIA

Genus Coptotelia Zeller

Coptotelia bipunctalis (Warren), new combination

Hyphypena bipunctalis Warren, 1889:232.

Hypercallia brysocyma Meyrick, 1921: (2)388; 1922:

180:163, no. 55.—Gaede, in Bryk 1939: 92:256. (New

synonymy).

Coptotelia bysocyma (Meyrick), Clarke 1951: 11:350, pi.

5: figs. 4-4b; 1963: 4:149, pi. 71: figs. 1-lb.

I have compared carefully the slide of the genitalia
36 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

of byrsocyma (JFGC 4622) with that of bipunctalis

(JFGC 4632) and am convinced of the above

synonymy.

TYPES.—British Museum (Natural History).

TYPE LOCALITIES.—Brazil, R. Jurua (bipunctalis);

Brazil, Obidos {byrsocyma).

DISTRIBUTION.—Brazil.

Genus Hypercallia Stephens

Hypercallia miltopa (Meyrick)

Coptotelia miltopa Meyrick, 1912:700.

Cryptolechia tunicata Busck, 1914:29. (New synonymy).

Hypercallia miltopa (Meyrick), 1922:163, pi. 6: fig. 101.—

Clarke 1963:294, pi. 144: figs. 3-3b.

Machimia tunicata (Busck), Meyrick 1922:82.

The specific synonymy here is clear but the generic

position of miltopa is uncertain. In all probability,

when the generic limits of the South American

species have been established, a new genus will be

required for this species and its congeners.

TYPES.—U.S. National Museum (tunicata);

British Museum (Natural History) (miltopa).

TYPE LOCALITIES.—Panama, Porto Bello (tunicata); Colombia, San Antonio (miltopa).

DISTRIBUTION.—Colombia, Panama.

Hypercallia heterochroma, new species

FIGURE 26; PLATE 3h.

Alar expanse 19-21 mm.

Labial palpus yellow; second segment edged with

brown anteriorly, in distal half, and apically; third

segment buff with sparse carmine scaling anteriorly.

Antenna pale yellow; basal seventh with blackish

fuscous, dorsoanterior longitudinal line; beyond this

faintly marked fuscous and remainder blackish

fuscous annulated; at basal two fifths a few carmine

spots. Head yellow with light carmine blotch posteriorly. Thorax brown with large black spot posteriorly; tegula edged outwardly with yellow and

carmine scales. Ground color of forewing brown

sparsely irrorate pale yellow; base of costa and two

triangular spots, one at basal third, the second at

two thirds of costa, yellow; apex from coastal four

fifths to vein 4, yellow; around apex to vein 5, a

slender, broken, brown line; costa brown mixed with

scarlet, where ground color intrudes between the

yellow costal markings; these markings edged scarlet;

from the yellow costal mark at basal third a zigzag

scarlet mark extends to middle of wing; on dorsum

an inwardly curved scarlet mark extends from basal

fifth to near middle; beyond end of cell a large oval

scarlet blotch, with a conspicuous, transverse, narrow series of black scales, extends to tornus; at basal

third, in cell, a conspicuous fuscous-edged white

spot; at end of cell a very small, ill-defined whitish

spot; cilia from apex to vein 4, yellow, remainder

brown. Hindwing yellowish white, basally shading

to grayish then pink toward termen; costa and apex

gray; cilia mixed pale yellow and gray. Foreleg

femur and tibia ocherous white, the tibia scarlet on

outer side; tarsal segments pale yellow; first tarsal

segment scarlet on outer side; midleg pale yellow to

ocherous white; hindleg ocherous white. Abdomen

ocherous white suffused pale grayish dorsally.

Male genitalia slide JFGC 12086. Harpe simple,

broadest before middle; sacculus sharply angulate;

cucullus, narrow, truncate. Gnathos a wide, transverse, oval spined knob. Uncus broad, stout, terminating in a small, sparsely setose protuberance.

Vinculum a moderately broad band, considerably

widened at middle. Tegumen wide, short, constricted

posteriorly. Anellus subrectangular, deeply excavate

posteriorly, and with a large, fleshy, setose lobe on

each side posteriorly. Aedeagus stout, nearly straight,

narrowed distally; vesica armed with one strong,

thornlike cornutus.

Female genitalia slide JFGC 12087. Ostium cup

shaped. Genital plate wide, moderately sclerotized.

Antrum greatly swollen, very lightly sclerotized. Inception of ductus seminalis lateral and slightly anterior to antrum. Ductus bursae membranous but

showing some slightly thickened longitudinal striations. Bursa copulatrix membranous. Signum a tiny

sclerotized thornlike plate.

HOLOTYPE.—USNM 71103.

TYPE LOCALITY.—Venezuela, Aragua, Rancho

Grande, 1100 m.

DISTRIBUTION.—Venezuela.

Described from the cT holotype (July 22-31,

1967) and one $ para type from the same locality

(June 22-30, 1967), both collected by R. W. Poole.

I have placed this species in Hypercallia temporarily so that it can be associated with its nearest

relative miltopa, the latter already assigned to that

genus. Eventually miltopa, heterochroma, and their

congeners will be placed in a new genus.
26b

FIGURE 26.—Hypercallia heterochroma, new species, a, ventral view of male genitalia with left

harpe and aedeagus removed; b, aedeagus; c, ventral view of female genitalia with bursa

copulatrix to left.
38 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Although very closely related to miltopa, heterochroma can be distinguished easily by its scarlet

blotches on the forewing and the absence of the

large, white discal dash found in miltopa.

Genus Phytomimia Walsinghani

Phytomimia chlorophylla Walsingham

Phytomomia chlorophylla Walsingham, 1912:134.

Phytomimia silvicolor Meyrick, 1932:279. (New synonymy).

I have examined Meyrick's type of silvicolor in the

Naturhistorisches Museum, in Vienna, and Walsingham's type of chlorophylla in the British Museum

(Natural History) The genitalia of the types, and

additional material before me from Mexico and

Costa Rica, all agree.

TYPES.—British Museum (Natural History)

(chlorophylla) ; Naturhistorisches Museum, Vienna

(silvicolor).

TYPE LOCALITIES.—Mexico, Guerrero, Venta de

Zopilote (chlorophylla); Panama, Lino, 2,650 feet

(815 m) (silvicolor).

DISTRIBUTION.—Mexico, Guatemala, Panama.

Literature Cited

Busck, August

1914. New Genera and Species of Microlepioptera from

Panama. Proceedings of the United States National Museum, 47:1-67.

Clarke, J. F. Gates

1951. The Moths of the Genus "Coptotelia" Zeller. De

Ada Zoologica Lilloana, 11:335-352, plates 1-5.

1963. Catalogue of the Type Specimens of Microlepidoptera in the British Museum (Natural History) described by Edward Meyrick, 4:1-521,

plates 1—252. London, England. British Museum

(Natural History).

Forbes, Wm. T. M.

1931. Supplementary Report on the Heterocera or Moths

of Porto Rico. The Journal of the Department of

Agriculture of Porto Rico, 4:339-394, plates

42-47.

Gaede, M.

1939. In Bryk, Lepidopterorum Catalogus. Oecophoridae.

92:209-476. 's-Gravenhage: W. Junk.

Meyrick, E.

1912. XXX. Descriptions of South American MicroLepidoptera. Transactions of the Entomological

Society of London, 1911:673-718.

1916. Exotic Microlepidoptera, 1:547-577. Taylor and

Francis. London.

1921. Exotic Microlepidoptera, 2:385-400. Taylor and

Francis. London.

1922. In Wytsman, Genera Insectorum. Lepidoptera

Heterocera. Oecophoridae, 180:1-200, plates 1-6.

1924. Exotic Microlepidoptera, 3:97-112. Taylor and

Francis. London.

1926. Exotic Microlepidoptera, 3:305-320. Taylor and

Francis. London.

1930. Exotic Microlepidoptera, 3:561-576. Taylor and

Francis. London.

1931. Exotic Microlepidoptera, 4:113-128. Taylor and

Francis. London.

1932. Exotic Microlepidoptera, 4:273-288. Taylor and

Francis. London.

Walsingham, Lord (Thomas de Grey)

1909- In Godman and Salvin, Biologia Centrali-Ameri1915. cana 42. {Lepidoptera-Heterocera, 4) :i-xii, 1-24

(1909); 25-40 (1910); 41-112 (1911; 113-168

(1912); 169-224 (1913); 225-392 (1914); 393-

482 (1915); figures 1-30, plates 1-10. London.

Warren, William

1889. VIII. On the Pyralidina Collected in 1874 and

1875 by Dr. J. W. H. Trail in the Basin of the

Amazons. Transactions of the Entomological Society of London, 1889:227-295.
NUMBER 95 39

Paper

I, II

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V

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VIII

IX

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Author

Clarke

Clarke

Duckworth

Obraztsov

Clarke

Obraztsov

Duckworth

Obraztsov

Duckworth

Obraztsov

Duckworth

Duckworth

Clarke

Duckworth

Clarke

Clarke

XVIII Duckworth

Neotropical Microlepidoptera Series

in Proceedings of the United States National Museum

(Papers are prepared with the aid of National Science Foundation Grants.)

Subject Year

Blastodacnidae and Aegeriidae

Gonionota melobaph.es and relatives

A new genus of Stenomidae

The tortricid genus Proeulia

The genera Orsotricha and Palinorsa

The tortricid genus Pseudomeritastis

The stenomid genus Falculina

The tortricid genus Pseudatteria

Taxa erroneously placed in Stenomidae

The tortricid genus Idolatteria

The stenomid genus Lethata

The stenomid genus Loxotoma

Chilean Microlepidoptera described by Blanchard

The stenomid genus Thioscelis

New species of Oecophoridae

New species of Phaloniidae

in Smithsonian Contributions to Zoology

Oecophorid genus Peleopoda 1970

Volume Number

1962

1964

1964

1964

1964

1966

1966

1966

1966

1966

1967

1967

1967

1967

1968

1968

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125

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