Fusulinid Biostratigraphy and

Correlations Between the

Appalachian and Eastern

Interir Basins

U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 1451

Prepared in cooperation ,with the 

Kentucky Geological Survey
Fusulinid Biostratigraphy and 

Correlations Between the 

Appalachian and Eastern 

Interior Basins

By RAYMOND C. DOUGLASS

U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 1451

Prepared in cooperation with the 

Kentucky Geological Survey

Fusulinids from western Kentucky and 

adjacent parts of Illinois and Indiana 

are compared and correlated with those 

from eastern Kentucky, Ohio, and 

western Pennsylvania

UNITED STATES GOVERNMENT PRINTING OFFICE, WASHINGTON: 1987
DEPARTMENT OF THE INTERIOR 

DONALD PAUL HODEL, Secretary

U.S. GEOLOGICAL SURVEY 

Dallas L. Peck, Director

Library of Congress Cataloging in Publication Data

Douglass, Raymond C.

Fusulinid biostratigraphy and correlations between the Appalachian and

eastern interior basins.

(U.S. Geological Survey professional paper; 1451)

Bibliography: p.

Supt. of Docs, no.: I 19.16:1451

1. Fusulinidae. 2. Paleontolougy Appalachian Region. 3.

Paleontology Middle West. 4. Stratigraphic correlation Appalachian

Region. 5. Stratigraphic correlation Middle West. I. Kentucky

Geological Society. II. Title. III. Series: Geological Survey professional

paper; 1451. 

QE75.P9 no. 1451 557.3 s [560M'728'0973] 86-600148 [QE772]

For sale by the Books and Open-File Reports Section, U.S. Geological Survey, 

Federal Center, Box 25425, Denver, CO 80225
CONTENTS

Abstract ____________________. 

Introduction __________________.

Acknowledgments ___________.

Previous studies ____________.

Current study ______________.

Geologic setting ____________.

Appalachian Basin ___________.

Eastern Interior (Illinois) Basin ____. 

Faunal succession ______________.

Zone of Millerella __________________

Zone of Profusulinella ______________.

Zone of Fusulinella ________________

Zone ofBeedeina __________________.

Subzone of Beedeina leei _________

Subzone of Wedekindellina _______

Subzone of Beedeina novamexicana 

Subzone of Beedeina girtyi ______

Subzone of Beedeina acme ________

Zone of Triticites __________________

Subzone of Triticites ohioensis ____

Page

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9

9

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II 

11 

11 

11

Faunal succession-Continued 

Zone of Triticites-Continued

Subzone of Triticites skinneri __________________

Subzone of Triticites callosus __________________

Subzone of Triticites beardi __________________

Correlations between the Eastern Interior and Appalachian 

Basins _____________________________________

Middle Pennsylvanian ___________________________

Beds of early Atokan age ______________________

Beds of late Atokan age _______________________

Beds of early Desmoinesian age _______________

Beds of middle Desmoinesian age _______________

Beds of late Desmoinesian age _________________

Upper Pennsylvanian ____________________________

Beds of Missourian age ______________________

Beds of Virgilian age _________________________

Lower Permian _________________________________

Register of localities and collections ___________________

Systematic descriptions _____________________________

References cited ___________________________________

Page

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12

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94

ILLUSTRATIONS

[Plates follow references cited]

PLATE 1. Profusulinella.

2. Profusulinella, Fusulinella.

3. Fusulinella.

4. Fusulinella

5. Fusulinella.

6. Fusulinella. 

1. Wedekindellina.

8. Beedeina.

9. Beedeina.

10. Beedeina.

PLATE 11. Beedeina.

12. Beedeina

13. Beedeina.

14. Beedeina.

15. Beedeina, Fusulinella.

16. Triticites, Beedeina, Wedekindellina.

17. Triticites, Beedeina, Wedekindellina

18. Triticites.

19. Triticites.

20. Triticites.

Ill
IV CONTENTS

Page

FIGURE 1. Index map showing localities for fusulinid collections 4

2. Representative fusulinids of eastern Kentucky and Ohio 6

3. Representative fusulinids of western Kentucky 7

4. Representative fusulinids of Illinois 8

5. Correlation chart 13 

6-31. Numerical-data graphs for

6. Profusulinella kentuckyensis Thompson and Riggs, 1959 28

7. Profusulinella ohioensis Douglass n. sp. 29

8. Fusulinella imprima Douglass n. sp. 31

9. Fusulinella primotina Douglass n. sp. 33

10. Fusulinella pria Douglass n. sp. 36

11. Fusulinella stouti Thompson, 1936 40

12. Fusulinella iownensis Thompson, 1934 44

13. Fusulinella sp. 45

14. Wedekindellina euthy'septa (Henbest), 1928 51

15. Beedeina sp. aff. B. novamexicana (Needham), 1937 54

16. Beedeina leei (Skinner), 1931 59

17. Beedeina sp. aff. B. spissiplicata (Dunbar and Henbest), 1942 61

18. Beedeina henbesti Douglass n. sp. 62

19. Beedeina ashlandensis Douglass n. sp. 65

20. Beedeina girtyi (Dunbar and Condra), 1927 67

21. Beedeina sp. aff. B. haworthi (Beede), 1916 69

22. Beedeina acme (Dunbar and Henbest), 1942 71

23. Beedeina carmani (Thompson), 1936 72

24. Triticites skinneri Thompson, 1936 75

25. Triticites ricei Douglass n. sp. 77

26. Triticites ohioensis Thompson, 1936 84

27. Triticites smythi Douglass n. sp. 85

28. Triticites kehni Douglass n. sp. 87

29. Triticites callosus Dunbar and Henbest, 1942 89

30. Triticites mediocris Dunbar and Henbest, 1942 91

31. Triticites beardi Douglass n. sp. 93

TABLES

Page

TABLES 1-28. Measurements for

1. Profusulinella kentuckyensis Thompson and Riggs, 1959 27

2. Profusulinella ohioensis Douglass n. sp. 28

3. Fusulinella imprima Douglass n. sp. 30

4. Fusulinella primotina Douglass n. sp. 32

5. Fusulinella pria Douglass n. sp. 34

6. Fusulinella stouti Thompson, 1936 37

7. Fusulinella iowensis Thompson, 1934 41

8. Fusulinella sp 44

9. Wedekindellina euthysepta (Henbest), 1928 46

10. Wedekindellina sp 51

11. Beedeina sp. aff. B. novamexicana (Needham), 1937 52

12. Beedeina leei (Skinner), 1931, and B. sp. aff. B. leei (Skinner), 1931 55

13. Beedeina sp. aff. B. spissiplicata (Dunbar and Henbest), 1942 60

14. Beedeina henbesti Douglass n. sp. 61

15. Beedeina ashlandensis Douglass n. sp. 63

16. Beedeina girtyi (Dunbar and Condra), 1927 66

17. Beedeina sp. aff. B. haworthi (Beede), 1916 68

18. Beedeina acme (Dunbar and Henbest), 1942, and B. sp. aff. B. acme 70

19. Beedeina carmani (Thompson), 1936 72

20. Beedeina sp. 73

21. Triticites skinneri Thompson, 1936 74
CONTENTS 

V

Page

22. Triticites ricei Douglass n. sp. 76

23. Triticites ohioensis Thompson, 1936 78

24. Triticites smythi Douglass n. sp. 85

25. Triticites kehni Douglass n. sp. 86

26. Triticites callosus Dunbar and Henbest, 1942 88

27. Triticites mediocris Dunbar and Henbest, 1942 90

28. Triticites beardi Douglass n. sp. 92

FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE 

APPALACHIAN AND EASTERN INTERIOR BASINS

By RAYMOND C. DOUGLASS

ABSTRACT

Fusulinids are common in marine rocks of the late Paleozoic succession in the Eastern Interior and Appalachian Basins. A detailed and 

systematic study of the fusulinids permits an improved correlation of 

the stratigraphy within and between these basins. Twenty-five species 

of fusulinids are recognized, of which 15 are forms previously named 

and 10 are new. Three additional forms are assigned only to genera. 

All the species are not represented in both basins, but they can be 

assigned to five zones and nine subzones that are recognized and correlated between the basins.

INTRODUCTION

In 1975, I was sent some limestone samples from 

southeastern Kentucky near the Virginia-Tennessee 

border, samples that contained many small fusulinids. I 

was asked whether these samples correlated with the 

Curlew Limestone Member of the Tradewater Formation of western Kentucky. Being unfamiliar with the 

Curlew, I asked whether comparative samples could be 

obtained from that unit. A suite of nine samples of the 

Curlew and mapped equivalents from five counties of 

western Kentucky was provided. On the basis of the contained fusulinids, these samples appeared to represent 

five different stratigraphic levels in three general 

groups-lower Atokan, upper Atokan, and Desmoinesian. The sample from the type locality of the Curlew in 

Union County, Ky., belonged about in the middle of this 

suite.

To get better control from the Middle Pennsylvanian 

part of the section, a set of core samples was studied 

from seven wells drilled at sites in six western Kentucky 

counties. It was gratifying to recognize in the core 

samples all the fossil horizons that had been studied in 

the surface samples. This study revealed that the 

Curlew Limestone Member was not represented in the 

samples submitted from one of the wells, but, because 

adjacent horizons were recognized, I was able to request 

samples from the probable intermediate interval and to 

find the missing Curlew fusulinids in them.

The corroboration of the surface-sample interpretation by the well data led to a flurry of activity to determine the relative position of many of the limestone beds 

in the surface exposures throughout western Kentucky. 

Additional well-core samples were investigated and correlated with the surface samples.

One of the obvious discrepancies noted early in the 

study was the difference between the type Curlew in 

Kentucky and the reported Curlew of southern Illinois. 

This difference pointed up the problem of correlation 

with the States adjacent to Kentucky, so samples from 

fusulinid-bearing units in Indiana and Illinois were collected and studied. These samples and the published 

record have made the correlation through the Illinois 

Basin possible.

Fusulinids from the Appalachian Basin had been 

described previously from Ohio, so samples were collected from various Middle and Upper Pennsylvanian 

horizons from which fusulinids had been reported. To 

round out the picture, samples were also collected from 

localities in western Pennsylvania, samples containing 

the easternmost fusulinids known in the United States.

ACKNOWLEDGMENTS

Most of the samples and stratigraphic data for this 

study were provided by C.L. Rice and T.M. Kehn of the 

U.S. Geological Survey (USGS), who both worked in 

cooperation with the Kentucky Geological Survey, Lexington, and by J.G. Beard of the Kentucky Geological 

Survey. Some core material was provided by the Institute of Mining and Mineral Research, University of 

Kentucky, Lexington. Additional faunas studied from 

collections in the U.S. National Museum of Natural 

History, Smithsonian Institution, Washington, B.C., 

were supplemented by collections that I made in Kentucky, Ohio, and Pennsylvania, with the aid of C.L. Rice 

and Richard Margerum, USGS; C.H. Cheong, Department of Geology, Seoul National University, Seoul, 

South Korea; and Rucha Ingavat, Geological Survey 

Division, Department of Mineral Resources, Bangkok, 

Thailand. Thin sections for the study were prepared 

with great skill and dexterity by Richard Margerum.

1
Types for comparison were kindly lent by the Peabody 

Museum of Natural History at Yale University, New 

Haven, Conn., the Illinois State Geological Survey at 

Urbana, and the State University of Iowa, Iowa City. 

Susanne Kaiser, USGS, assisted in compilation of data 

for this report, and Linda Jacobsen, USGS, assisted on 

the text figures. The typescript was read by C.L. Rice, 

T.M. Kehn, T.W. Henry, and D.A. Meyers, all USGS; 

their helpful comments are greatly appreciated.

PREVIOUS STUDIES

Several published studies are especially pertinent to 

this report. Dunbar and Henbest (1942) documented an 

extensive study of the fusulinids and stratigraphy of Illinois. Thompson (1936) described some of the fusulinids 

from Ohio; Thompson and Riggs (in Thompson, Shaver, 

and Riggs, 1959) studied early fusulinids from western 

Kentucky; and Thompson and Shaver (1964) evaluated 

early fusulinds from Illinois, Indiana, and Kentucky. 

Smyth (1957, 1974) documented fusulinids from Ohio 

and adjacent parts of Kentucky and Pennsylvania, and 

St. Jean (1957) studied some fusulinids from Indiana. 

Wanless (1975) presented a regional analyses of the 

Pennsylvanian System in the Appalachian region and 

the Illinois Basin region. Rice, Kehn, and Douglass 

(1979) published on Pennsylvanian correlations between 

the Eastern Interior and Appalachian Basins. Douglass 

(1979) illustrated the distribution of fusulinids and their 

correlation between the Illinois Basin and the Appalachian Basin.

CURRENT STUDY

This paper documents the fusulinid faunas and correlations between the Eastern Interior Basin and the 

Appalachian Basin, providing a biostratigraphic framework and a description of the fusulinids. The study 

started when a few samples were sent in for identification but grew rapidly as more samples were acquired. 

Surface samples were studied from all the exposed 

Pennsylvanian limestone units in western Kentucky. 

Figure 1 shows localities from which the samples were 

taken. Comparative material was also assembled from 

Illinois and Indiana (fig. 1). Supplemental samples were 

taken from many drill hole cores. The calcareous intervals represented in the cores were sampled for 

fusulinids. A large number of surface samples was also 

assembled from the Appalachian Basin, from Bell County in southeastern Kentucky along a broad line extending north-northeast through Ohio and east into western 

Pennsylvania. Two drill holes in eastern Kentucky were 

also sampled for the calcareous intervals containing

fusulinids. Figure 1 also shows the localities in the Appalachian Basin. The localities and the included collections are described elsewhere in this report. Every effort was made to obtain as complete a representation of 

the fusulinid faunas from the two basins as possible. 

Rumors of possible fusulinids in the proposed Pennsylvanian stratotype section in central West Virginia 

were pursued in the field and in laboratory studies, but 

no fusulinids were found. Collections from the Kentucky 

and Ohio areas near the West Virginia border commonly 

yield good Pennsylvanian fusulinids.

Preparation of useful thin sections of the fusulinids 

was a difficult task, especially in samples of nearly black 

calcareous shale and argillaceous limestone. Fortunately, Richard Margerum was able to use his knowledge 

and skill to obtain oriented sections even under the most 

difficult conditions.

Throughout the study, T.M. Kehn and C.L. Rice provided consultation on field relations and physical correlation to supplement the published stratigraphic data 

and were able to collect samples to fill out the 

stratigraphic sections in Kentucky and in the adjoining 

States.

The present study is based on these collections and on 

many samples available in the National Museum of 

Natural History (USNM) and in the collections at Yale 

University (YPM), the State University of Iowa (SUI), 

and the Illinois State Geological Survey (ISGS).

GEOLOGIC SETTING

The two depositions! areas of rocks of late Paleozoic 

age are separated by the structural high of the Cincinnati Arch. To the east is the Appalachian Basin, and to 

the west is the Eastern Interior or Illinois Basin. This 

study concerns those parts of the Appalachian Basin 

represented in western Pennsylvania, eastern Ohio, and 

eastern Kentucky and those parts of the Eastern Interior Basin represented in southern Illinois, southern 

Indiana, and western Kentucky (fig. 1.)

APPALACHIAN BASIN

Pennsylvanian marine deposits are common in an 

elongate basin extending from Pennsylvania to Tennessee, but fusulinid foraminifers are not represented 

throughout the entire basin. Fusulinids required open 

marine waters and are, therefore, found only in the 

parts of the basin where the seas encroached on the 

estuarine and riverine deposits sufficiently to dominate 

the environment. Many parts of the basin have deposits 

containing mollusks and even brachiopods that could 

tolerate a more brackish-water environment. The oldest
FAUNAL SUCCESSION

horizon containing fusiform fusulinids is in the Stoney 

Fork Member of the Breathitt Formation (formerly the 

Lost Creek Limestone of the Breathitt Formation of 

Morse, 1931), recognized in surface samples and drill 

cores in southeastern Kentucky (localities 45-53). 

Fusulinids of the same approximate age are found in the 

Boggs Limestone Member of the Pottsville Formation in 

eastern Ohio (localities 67, 74, 81). (Also, see fig. 2.) 

These two parts of the basin are separated by more than 

100 miles in which correlative beds have not been identified. The younger section in Ohio contains additional 

fusulinid-bearing marine horizons represented by the 

upper and lower Mercer Limestone Members (as used by 

Smyth, 1957) of the Pottsville Formation.

The section in eastern Kentucky lacks fusulinidbearing beds through most of the rest of the rocks of 

Middle Pennsylvanian age. Small indeterminate 

fusulinids were found at locality 55 in Carter County in 

rocks assigned to the Vanport Limestone Member of the 

Allegheny Formation. Abundant fusulinids occur on the 

Ohio border at locality 61 in Greenup County, Ky. The 

limestone at this locality was assigned to the Vanport 

Limestone by Phalen (1912) but is now considered 

equivalent to the Columbiana Limestone Member of the 

Allegheny Formation in Ohio. Fusulinid-bearing 

limestones are present at several levels through the 

Allegheny Formation in Ohio. The most persistent 

marine horizons are the Putnam Hill Limestone, the 

Vanport Limestone, and the Columbiana Limestone, all 

members of the Allegheny Formation. The principal 

area of outcrop for these units is in east-central Ohio, 

but the outcrops extend in a limited way into western 

Pennsylvania (localities 86, 87).

Marine zones of Late Pennsylvanian age are geographically more limited in the basin. The section in 

eastern Kentucky includes fusulinid-bearing beds of the 

Brush Creek and Cambridge Limestone Members of the 

Conemaugh Formation. These strata were found in outcrops and drill cores only in Boyd, Carter, and Lawrence 

Counties in northeastern Kentucky (localities 54-61). 

No occurrences have been confirmed from the adjoining 

areas of West Virginia. The Ohio part of the basin contains additional fusulinid horizons in the Conemaugh 

Formation in the areas of outcrop from Gallia County to 

Guernsey County. The principal units, as in Kentucky, 

are the Brush Creek and Cambridge Limestone 

Members that represent the Missourian Series. The 

Ames and Gaysport Limestone Members of the Conemaugh Formation also yield fusulinids at some 

localities in the same area. These horizons represent 

Virgilian age. One locality (88) in Pennsylvania at 

Brilliant Cut contains fusulinids that suggest Virgilian 

age. No fusulinids younger than the Conemaugh are 

known from the Appalachian Basin.

EASTERN INTERIOR (ILLINOIS) BASIN

The western part of Kentucky includes the southeastern part of the late Palezoic Eastern Interior Basin 

that covered much of Illinois and extended into adjoining States. The section in western Kentucky includes 

beds ranging in age from Early Pennsylvanian through 

the Late Pennsylvanian and locally contains a remnant 

of the Early Permian section (fig. 3).

Fusulinid-bearing marine beds, which are irregularly 

interspersed with sandstone, shale, and coal beds, are 

more persistent laterally than they are in the Appalachian Basin. A relatively complete fusulinid succession through the Middle and Upper Pennsylvanian is 

available from surface exposures and drill-core samples. 

The oldest fusulinid horizon is a little older than any 

represented in the Appalachian Basin and is found in the 

Lead Creek Limestone Member of the Tradewater Formation. This horizon is present in surface samples in 

southern Indiana and in surface and drill-core samples in 

seven counties in western Kentucky. It has not been 

recognized in Illinois (fig. 4). The youngest fusulinid 

horizon is of Early Permian age and is known only from 

an unnamed limestone in the Mauzy Formation in a 

graben of the Rough Creek Fault system (locality 19) in 

Union County, Ky. Rocks containing fusulinids of latest 

Pennsylvanian age are found at locality 1 in Cumberland 

County, 111.

FAUNAL SUCCESSION

Fusulinid zonations based on genera are commonly 

used to recognize divisions of the Pennsylvanian and 

Permian succession (Douglass, 1977). The rocks of the 

Eastern Interior Basin and the Appalachian Basin can 

be assigned to the fusulinid zones as commonly used, 

but, because of the large amount of data assembled for 

this study, it is appropriate to restudy the use of the 

zones as they apply specifically to these basins. The 

following discussion indicates the uses and the limitations of fusulinid zonation for correlation within and between the basins.

ZONE OF MILLERELLA

The zone of Millerella is commonly recognized as including beds of Early Pennsylvanian age below the first 

occurrence of Profusulinella. The genus Millerella is not 

well defined and ranges in age from at least Early Pennsylvanian to possible Permian. The upper limit of the 

zone is indefinite in the absence of Profusulinella. 

Millerella is present in rocks starting below the first 

Profusulinella and continuing into younger beds in 

several parts of the basins studied, but no attempt was
41

40°

39'

>,/''-'^H ^j.«;5r*i) ^ f4-;r v.t;n;:

Figure 1.-Localities of fusulinid collections. The localities marked with triangles are surface samples; those depicted with circles are drill cores.
FAUNAL SUCCESSION

84° 83

41°

40° -,cv.. 77^, - , ^80 

~ ,* \ I " A ^ 74 i ^ "terjiLJjMays^ 75\ A73

r I ""**" 76 A^-,, 72

"" ' >- , ) JPIKE '''( -i >s? , -, - -^ . ;,-;,

,^ 5,0'. _ |B«»OW-. I | l - -_.

Figure 1-Continued.
li

EASTERN KENTUCKY OHIO

<«&* *&» » e

"*lp* «dfr ^»,

^ pm ^^%? ^n^' %

« {^^

Figure 2. - Representative fusulinids of eastern Kentucky and Ohio. The correlation of the stratigraphic units shown and their relationship to the fusulinid zones are shown on figure 5.
FAUNAL SUCCESSION

"'""'

 <as& -m

Figure 3. - Representative fusulinids of western Kentucky. The correlation of the stratigraphic units shown and their relationship to the

fusulinid zones are shown on figure 5.
8 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS

-T-V-T. -\

s*#-*">

-^> ** *S2? - .

rt ^SI^'^M ^ps ''IF ^^2^^^. '^

Figure 4.-Representative fusulinids of Illinois. The correlation of the stratigraphic units shown and their relationship to the fusulinid

zones are shown on figure 5.
FAUNAL SUCCESSION

made to define the extent or limits of the zone for this 

study.

ZONE OF PROFUSULINELLA

The lower part of the Atokan series of Middle Pennsylvanian age is included in the zone of Profusulinella. 

The zone includes the beds between the first occurrence 

of Profusulinella and the first occurrence of Fusulinella 

(Douglass, 1977, p. 478). Restudy of the type Atokan 

(Sutherland, 1982) and of its fusulinids (Douglass 1982; 

Douglass and Nestell, 1984) indicates that the Kentucky 

and Oklahoma sequences have many similarities. Both 

sequences have an indefinite lower boundary because 

early forms of Profusulinella are unknown in the section, and both have beds containing more advanced profusulinellas that are followed by younger beds containing fusulinids assigned to Fusulinella. A clear-cut 

distinction between Profusulinella and Fusulinella 

would make a separation of the zones easy, but this is a 

gradational evolutionary sequence (Douglass, 1981). 

Profusulinella grades into Fusulinella by the addition of 

an inner tectorium in the outer volutions of a form, the 

juvenile form of which is profusulinellid.

In western Kentucky, Profusulinella is present in surface and drill-core samples. The surface exposures are 

from the Lead Creek Limestone Member (Lead Creek 

Limestone of Crider, 1913) of the Tradewater Formation and are found through about 15 ft of section, 

primarily in two limestone beds a little more than 6 ft 

apart. The section available in drill cores shows a similar 

set of beds at the top and an occurrence of Profusulinella 30-45 ft deeper. This lower set of beds has 

not been recognized in outcrop.

Profusulinella kentuckyensis Thompson and Riggs, 

1959, characterizes the zone in western Kentucky. This 

species was described from the surface samples in 

Butler County (locality 36). The forms present in the 

drill cores show great variability (pi. 1) but are within 

the range of variation seen in the surface samples and 

have not been given a different name. None of the 

specimens from Kentucky resemble the early Profusulinellas described from Texas, New Mexico, and 

other Western States, and I assume that some of the 

forms from those areas represent an earlier stage of 

evolution. The species from Kentucky is a late form that 

has a tendency toward elongation and the incipient 

development of an inner tectorium. Surface samples 

have not yielded the next immediate stage in the evolutionary sequence, but the core hole at locality 21 in 

Webster County contains a fauna intermediate between 

P. kentuckyensis and Fusulinella iowensis, consisting of 

a primitive Fusulinella described as F. primotina n. sp.

The overlying horizon containing Fusulinella iowensis 

is 2.1-2.5 ft above F. primotina, and the underlying

Profusulinella kentuckyensis is about 30 ft. below F. 

primotina. For practical purposes, the top of the Profusulinella zone and the base of the Fusulinella zone 

should be taken as the base of the Curlew Limestone 

Member of the Tradewater Formation. The base of the 

Curlew is not a sharp boundary because the limestone 

grades down into nodular limestone and calcareous 

shale that could be considered part of the Curlew.

Shaver and Smith (1974) summarized the studies of 

Thompson and Riggs (in Thompson, Shaver, and Riggs, 

1959) and Thompson and Shaver (1964) for fusulinids of 

the Middle Pennsylvanian in Indiana and adjacent parts 

of Kentucky. Shaver and Smith reported P. kentuckyensis from the Fulda and Ferdinand Beds of the Lead 

Creek Limestone Member of the Mansfield Formation 

and P. burrensis from the Brazil Formation that they 

consider possibly younger than the Lead Creek 

Limestone. The first more advanced fusulinid that they 

reported is a Fusulinella sp. also from the Brazil Formation at a different locality. They proposed that the range 

of Profusulinella and Fusulinella may overlap in Indiana (Shaver and Smith, 1974, p. 17). F. iowensis occurs in the overlying Staunton Formation. Verification 

of the undescribed form of Fusulinella would suggest 

the Profusulinella persisted longer in Indiana than it 

seems to have persisted in Kentucky.

The zone of Profusulinella has not been identified in 

the Appalachian Basin. The oldest fusulinids present are 

in the Stoney Fork Member of the Breathitt Formation 

and the Boggs Limestone Member of the Pottsville Formation at locality 67 in Hocking County, Ohio. At this 

locality, the argillaceous nodular limestone has P. 

ohioensis n. sp. associated with F. imprima n. sp. The 

fusulinellid is a primitive form representing a stage of 

development similar to that of F. primotina n. sp. The 

form described as F. pria n. sp. from the Stoney Fork 

Member of the Breathitt Formation is also a primitive 

fusulinellid representing approximately the same early 

stage of development. The top of the zone of Profusulinella must be very close to the base of the Stoney 

Fork and to that of the Boggs. Again, as in Indiana, Profusulinella persisted in Ohio into the lower part of the 

zone of Fusulinella.

Other foraminifers commonly found in association 

with Profusulinella in the zone include Climacammina 

sp., Tetrataxis sp., Millerella sp., and, less commonly, 

Pseudostajfella sp..

ZONE OF FUSULINELLA

The upper part of the Atokan Series of Middle Pennsylvanian age is included in the zone of Fusulinella. The 

zone extends from the first occurence of Fusulinella to 

the first occurrence of Beedeina (Fusulina of previous 

use). The fact that Fusulinella evolved from Pro-
10 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS

fusulinella through transition and evolved into Beedeina 

through transition makes the zone limits dependent on 

taxonomic and nomenclatural decisions. The evolutionary changes took place rapidly enough so that, in 

practice, there is no great problem in assigning 

stratigraphic units. The section in the type area of the 

Atokan Series in southern Oklahoma shows a sequence 

of beds containing several species of Fusulinella above 

the Profusulinella zone (Douglass and Nestell, 1984). 

The species present in Oklahoma are not represented in 

the Kentucky section, but Fusulinella sp. aff. F. leyi, 

described from near the top of the section, is of about 

the same stage of development as forms from the type 

Curlew Limestone Member of the Tradewater Formation in Kentucky.

The zone of Fusulinella in the Eastern Interior Basin 

is relatively restricted in the stratigraphic section. The 

Kentucky section includes several beds of the type 

Curlew Limestone Member at locality 17 in Union County that contain F. iowensis and F. stouti. This section 

also includes the slightly older horizon of the Curlew 

Limestone Member found at locality 21 that contains F. 

primotina n. sp.

The equivalent interval to the Curlew Limestone 

Member is the Seville Limestone Member of the Spoon 

Formation, which contains similar species. The zone of 

Fusulinella in Indiana includes beds in the upper part of 

the Brazil Formation and the Perth Limestone Member 

of the Staunton Formation. The form Fusulinellal is 

mentioned by Thompson and Shaver (1964, p. 17) from 

strata tentatively assigned to the Brazil Formation or 

possibly to the Perth Limestone Member of the Staunton Formation. The Staunton also contains F. iowensis.

The Fusulinella zone in the Appalachian Basin is 

represented through the upper part of the Pottsville 

Formation by the Boggs Limestone Member and the upper and lower Mercer Limestone Members in Ohio and 

by the Stoney Fork Member of the Breathitt Formation 

in eastern Kentucky. The fauna includes P. ohioensis 

and F. imprima in the Boggs Limestone and F. pria in 

the Stoney Fork Member.

Fusulinella and Beedeina occur together in several 

areas, but the zone of Fusulinella is restricted to the 

beds below the first occurrence of Beedeina.

ZONE OF BEEDEINA

The zone of Beedeina includes all the beds between the 

zone of Fusulinella and the zone ofTriticites. This interval includes all the Desmoinesian Series of Middle Pennsylvanian age. The genus Beedeina is found throughout 

the zone as it evolves from a form differing only slightly 

from Fusulinella to the large, tightly fluted species of 

the latest Desmoinesian. The changes take place 

gradually through the zone, but some developmental

stages are striking enough to allow for division of the 

zone into recognizable subzones.

SUBZONE OF BEEDEINA LEEI

Beedeina leei is common in the "Curlew" Limestone 

Member of the Spoon Formation in southern Illinois and 

is the only fusulinid reported from this bed. The 

"Curlew" is the first marine limestone above the Seville 

Limestone Member of the Fusulinella zone. The name 

"Curlew" is based on miscorrelation with the Curlew 

Limestone Member of Kentucky, which is equivalent in 

age to the Seville Limestone Member. B. leei is the 

species characteristic of the Putnam Hill Limestone 

Member of the Allegheny Formation in Ohio. Here 

again, this species is the only fusulinid reported from the 

bed, and this unit is the first marine limestone above the 

upper Mercer Limestone Member of the Fusulinella 

zone. Thompson (1936, p. 677, 678) assigned the Putnam 

Hill specimens to his new species F. serotina but 

recognized the more advanced septal fluting that has led 

to the assignment of these specimens to Beedeina. A 

similar set of specimens was collected from the Mahoning Limestone of Rogers (1858) at locality 87 in Armstrong County, Pa. B. leei occurs with Beedeina sp. at 

this locality. There is no direct correlation with the Putnam Hill Limestone Member, but the fusulinids suggest 

this lowest subzone of the Desmoinesian.

The Kentucky sections include B. leei at several places 

both in outcrop (locality 32) and in drill cores (localities 

18, 35, and 40). The specimens come from an outcropping limestone that was thought to be "Curlew," 

from about 15 ft above the No. 4 coal at locality 32 in 

Christian County. The limestone below the coal here correlates with the Curlew Limestone Member and contains the Fusulinella fauna. At this locality, B. leei 

occurs with Wedekindellina euthysepta and may represent a younger horizon than the "Curlew" Limestone 

Member of Illinois and the Putnam Hill Limestone 

Member of Ohio.

The core samples from western Kentucky contain Beedeina leei and B. sp. aff. B. leei in several wells. At locality 18 in Union County, B. leei occurs as expected above 

the F. iowensis of the Fusulinella zone and below more 

advanced Beedeina with W. euthysepta in the Yeargins 

limestone member (informal usage) of the Tradewater 

Formation. The other drill-core samples show B. leei 

associated with a Wedekindellina sp. that cannot be further identified.

SUBZONE OF WEDEKINDELLINA

The genus Wedekindellina is found through the lower 

beds of the Desmoinesian series in both basins but is 

more common in the Eastern Interior Basin than in the
FAUNAL SUCCESSION 11

Appalachian Basin. The first occurrences are within the 

zone of Beedeina leei, as previously noted. The most 

common occurrences are in the Stonefort Limestone 

Member of the Spoon Formation and its equivalents in 

Illinois, the Yeargins limestone member of the 

Tradewater Formation in Kentucky, and the Vanport 

Limestone Member of the Allegheny Formation in Ohio 

and Pennsylvania. Reports of Wedekindellina in 

younger beds (Dunbar and Henbest, 1942, p. 30) are not 

included, because the specimens are now assigned to 

Eowaeringella Skinner and Wilde, 1967.

SUBZONE OF BEEDEINA NOVAMEXICANA

Beedeina novamexicana in known from Illinois in the 

Stonefort Limestone Member of the Spoon Formation. 

The Kentucky specimens related to this species are from 

the Yeargins limestone member (informal usage) of the 

Tradewater Formation. Wedekindellina is a common 

element in this subzone, as noted above. This subzone 

may be represented in the Appalacian Basin by the Vanport and Columbiana Limestone Members of the 

Allegheny Formation. The Vanport has B. carmani 

(Thompson) and W. euthysepta, and the Columbiana has 

B. ashlandensis n. sp. Both limestones correlate with the 

Yeargins limestone member of western Kentucky, but 

the Wedekindellina is the only fusulinid common to all 

these limestones.

SUBZONE OF BEEDEINA GIRTYI

Beedeina girtyi is common in the Brereton and Herrin 

Limestone Members of the Carbondale Formation in Illinois. The same subzone is represented in Kentucky in 

the Providence Limestone Member of the Sturgis Formation. No marine beds of equivalent age are known 

from the Appalachian Basin. The fusulinids of the B. girtyi subzone include B. girtyi, B. haworthi, and the 

specimens described by Dunbar and Henbest (1942) as 

Fusulina illinoisensis that are probably closely related 

to B. girtyi. Additional specimens that are approaching 

the stage of B. acme are found in some samples assigned 

to the Providence Limestone Member, but they have not 

developed the characteristic subcylindrical shape of that 

species.

SUBZONE OF BEEDEINA ACME

Beedeina acme is found in the marine limestone beds 

of the Modesto Formation in Illinois and in the Madisonville Limestone Member of the Sturgis Formation in 

Kentucky. The names Piasa, Cutler, and Lonsdale 

Limestone were used for these beds by Dunbar and 

Henbest (1942, p. 25, 26), leaving some question as to 

their relative stratigraphic position but placing them all

in the latest part of the Desmoinesian sequence with the 

last representatives of Beedeina (their Fusulina). Dunbar and Henbest (1942) recognized five species of Beedeina (their Fusulina) in this unit, of which only B. acme 

is recognized in the Kentucky section. No marine 

horizons equivalent to these beds were found in the Appalachian Basin.

ZONE OF TRITICITES

The zone of Triticites includes all the rocks of Late 

Pennsylvanian (Missourian and Virgilian) age. It includes the rocks containing the first Triticites, but it 

does not include the upper limits of the range of the 

genus; Triticites also occurs in the overlying zones of 

Schwagerina and Pseudoschwagerina. Subzones of 

Triticites can be distinguished within the Eastern Interior and Appalachian Basins.

SUBZONE OF TRITICITES OHIOENSIS

Triticites ohioensis was described from the Brush 

Creek and Cambridge Limestone Members of the Conemaugh Formation in Ohio. The subzone includes the 

lowest beds of the Brush Creek in southern Ohio and 

eastern Kentucky, the Cambridge in Ohio, and probably 

the Portersville Member of the Conemaugh Formation 

in Ohio, from which Smyth (1957, p. 262) reported probable T. ohioensis. The equivalent interval in the Eastern 

Interior Basin includes the Shoal Creek and Livingston 

Limestone Members of the Bond Formation and the 

Omega Limestone Member of the Mattoon Formation in 

Illinois and the Carthage Limestone Member of the 

Sturgis Formation in Kentucky. Other fusulinids found 

in this subzone include T. venustus in Illinois, T. kehni n. 

sp. in Kentucky, and T. smythi n. sp. in Ohio.

SUBZONE OF TRITICITES SKINNERI

Triticites skinneri was described from the Ames 

Limestone Member of the Conemaugh Formation in 

Ohio. It is common in that unit and has been reported 

from the Gaysport Member (Smyth, 1957, p. 262), but 

that occurrence has not been verified. T. skinneri is 

associated with T. ricei n. sp. in the Ames Limestone 

Member. T. cullomensis was also reported from the 

Ames Limestone Member (Smyth, 1957, p. 273, 274) in 

Ohio, but the specimens illustrated (pi. 3, figs. 23, 24) 

are more properly assigned to T. skinneri. The 

specimens of T. cullomensis from Brilliant Cut in Pittsburgh, Pa., are referred to the Ames Limestone Member 

but may represent a slightly younger horizon. The 

Shumway Limestone Member of the Mattoon Formation in Illinois carries a faunna of T. turgidus and T. 

pauper that closely resemble T. skinneri in stage of 

development and probably represent the subzone.
12 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS

SUBZONE OF TRITICITES CALLOSUS

The Greenup Limestone Member of the Mattoon Formation has a fusulinid fauna including Triticites callosus 

and T. mediocris that suggest a late developmental 

stage of the genus, approaching the T. ventricosus 

known from rocks of Early Permian age in the Midcontinent. This subzone is the youngest fusulinid unit known 

from the Illinois part of the Eastern Interior Basin, and 

it is only known from Cumberland County, 111. (locality 

1). No equivalent marine beds are known from the 

Apalachian Basin. Outcrops in western Kentucky do not 

include a marine horizon representing this subzone. The 

one drill core that might have penetrated the subzone 

(locality 19) did not have fusulinids to verify it.

SUBZONE OF TRITICITES BEARDI

A unit that in the Midcontinent could be called the 

zone of Triticites ventricosus is represented in the 

graben in the Rough Creek fault system. The unit is only 

known from one drill core and contains a form described 

as T. beardi n. sp. This form represents a developmental 

stage similar to forms described from rocks of Early 

Permian (Wolfcampian) age. No fusulinids of this age 

are known from other parts of either the Eastern Interior or Appalachian Basins.

CORRELATIONS BETWEEN THE 

EASTERN INTERIOR AND APPALACHIAN BASINS

An introduction to the correlations of units between 

the Eastern Interior and Appalachian Basins was 

presented by Rice, Kehn, and Douglass in 1979, on the 

basis of the studies completed at that time. Precise correlation of beds between the basins is not possible 

because sedimentation was not uniform in the separate 

basins and because marine beds are discontinuous even 

within each basin. Correlations are, nevertheless, possible on the basis of the faunal succession described. 

Figure 5 summarizes the correlations, showing the approximate equivalence of many of the named beds in five 

generalized sections. Correlation between beds that 

have no fusulinid faunas is not suggested, and the intervening beds are placed only approximately.

The succession will be treated from bottom to top, 

starting with the oldest beds that have fusiform 

fusulinids and moving upsection. The correlations possible for each fusulinid-bearing unit are indicated.

MIDDLE PENNSYLVANIAN

BEDS OF EARLY ATOKAN AGE

The oldest unit bearing fusiform fusulinids is the Lead 

Creek Limestone Member of the Tradewater Formation. This member is exposed in western Kentucky and 

in southern Indiana (not shown in fig. 5) and is known 

from several drill cores in western Kentucky. The 

member consists of several beds, two of which were 

called the Fulda and Ferdinand Limestone Members 

(Mansfield Formation) in Indiana by Thompson and 

Shaver (1964). These beds were later called the Ferdinand and Fulda Beds of the Lead Creek Limestone 

Member of the Mansfield Formation by Shaver and 

Smith (1974). Other older beds that might be considered 

to belong to this unit are found in some of the drill cores, 

but no names have been applied to them. No fusiform 

fusulinids of this age are known for Illinois or from the 

Appalachian Basin. The member probably correlates 

with strata of the Abbott Formation in Illinois and, 

perhaps, in part, with the Magoffin Member of the 

Breathitt Formation in Kentucky. Both the Lead Creek 

and the Magoffin are very close in age but are older than 

the Boggs Limestone Member of the Pottsville 

Limestone in Ohio.

BEDS OF LATE ATOKAN AGE

A fairly close correlation can be made between the 

lowest beds of the Curlew Limestone Member of the 

Tradewater Formation at locality 21, the Stoney Fork 

Member of the Breathitt Formation in eastern Kentucky, and the Boggs Limestone Member of the Pottsville Formation in Ohio. Each of these units has a 

primitive Fusulinella intermediate between Profusulinella and typical fusulinellas. The species are 

distinct but closely related. No representatives of this 

level are known from surface exposures in the Eastern 

Interior Basin.

This level is commonly overlain in drill cores and in 

surfaces deposits of the Eastern Interior Basin by the 

Curlew Limestone Member of the Tradewater Formation as it is defined in Union County, Ky. The Curlew 

Limestone Member correlates with the Seville 

Limestone Member of the Spoon Formation in Illinois, 

the Perth Limestone Member of the Staunton Formation in Indiana, and the Mercer Limestone Member of 

the Pottsville Formation in Ohio. No fusulinid-bearing 

equivalents are known from eastern Kentucky.

BEDS OF EARLY DESMOINESIAN AGE

The beds called Curlew Limestone by Dunbar and 

Henbest (1942, p. 20), which are younger than the type 

Curlew of Kentucky, are probably correlative with the 

Creal Springs Limestone Member of the Spoon Formation (Wanless, 1975, p. 79). They correlate with the 

lower part of the Yeargins limestone member of the 

Tradewater Formation in surface and drill-core samples
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14 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS

and with the Putnam Hill Limestone Member of the 

Allegheny Formation in Ohio.

BEDS OF MIDDLE DESMOINESIAN AGE

The Stonefort Limestone Member of the Spoon Formation in Illinois correlates with the Yeargins limestone 

member of the Tradewater Formation in western Kentucky. The beds containing fusulinids of this general age 

in the Appalachian Basin have few species in common, 

and a precise correlation cannot be made. The Vanport 

and Columbiana Limestone Members of the Allegheny 

Formation in Ohio and Kentucky and the Vanport 

Limestone Member in Pennsylvania are all assigned a 

middle Desmoinesian age.

BEDS OF LATE DESMOINESIAN AGE

The Brereton and Herrin Limestone Members of the 

Carbondale Formation in Illinois correlate with the 

Providence Limestone Member of the Sturgis Formation in western Kentucky.

The Piasa, Cutler (now included in the Piasa), and 

Lonsdale Members of the Modesto Formation in Illinois 

correlate with the Madisonville Limestone Member of 

the Sturgis Formation in Kentucky.

No fusulinids of this age are known from the Appalachian Basin, but the upper beds of the Breathitt Formation in Kentucky and of the Allegheny Formation in 

Ohio should be equivalent in age.

UPPER PENNSYLVANIAN

BEDS OF MISSOURIAN AGE

The Shoal Creek and Livingston Limestone Members 

of the Bond Formation and the Omega Limestone 

Member of the Mattoon Formation in Illinois correlate 

approximately with the Carthage Limestone Member of 

the Sturgis Formation in western Kentucky. The Carthage Limestone Member probably represents the upper 

part of these beds. The Appalachian Basin equivalents 

are the several beds assigned to the Brush Creek and 

Cambridge Limestone Members of the Conemaugh Formation.

BEDS OF VIRGILIAN AGE

The Shumway Limestone Member of the Mattoon Formation in Illinois correlates with the Ames Limestone 

Member of the Conemaugh Formation in Ohio. No 

equivalents are known in eastern Kentucky. The Ames 

Limestone Member of Pennsylvania is probably a little 

younger than the Ames in Ohio.

The Greenup Limestone Member of the Mattoon Formation is younger than the Ames of Pennsylvania and is 

the youngest fusulinid horizon known from the Pennsylvanian of either basin.

LOWER PERMIAN

Strata of Early Permian age are known from only one 

locality in the Eastern Interior Basin (Kehn, Beard, and 

Williamson, 1982). Fusulinids of Early Permian age are 

from a drill core at locality 19 in Union County, Ky.

REGISTER OF LOCALITIES AND COLLECTIONS

[Localities are keyed to index map, fig 1]

Locality Collection

ILLINOIS

1 ___£13995 __Cumberland County. Pennsylvanian-Permian. 

1,100 ft from north line and 1,900 ft from 

east line sec. 3, T. 9 N., R. 9 E., Toledo 

15-min Quadrangle. Approx. 1 mi northwest 

of Greenup, about 300 ft north of Illinois 

Highway 121. T.M. Kehn colln. KB-42. 

Triticites callosus, T. mediocris.

f!4012 __Cumberland County. Upper Pennsylvanian.

Approx. 1 mi airline northwest of Greenup. 

About 2,150 ft from east line and 1,900 ft 

from north line sec. 3, T. 9 N., R. 9 E., Toldeo 

15-min Quadrangle. Greenup Limestone 

Member of Mattoon Formation. H. Schwalb 

and J. Beard colln. KB-44. Triticites 

callosus, T. mediocris.

f 14013 __Cumberland County. Upper Pennsylvanian.

Re-collection of KB-42 (f!3995) in the 

Greenup Limestone Member. T.M. Kehn 

colln. KB-41, re-collection at same locality 

but from bed about 4 ft lower in section. 

Triticites callosus, T. mediocris.

2 ___fl3875 ___Williamson County. Middle Pennsylvanian.

NEV4,SEV4, sec. 25, T. 10 S., R. 4 E., Harrisburg 15-min Quadrangle. Type locality for 

Stonefort Limestone Member of Spoon Formation. T.M. Kehn, collector, 1977. 

Wedekindellina euthysepta, Beedeina sp. aff. 

B. novamexicana.

f!3988 __Williamson County. Middle Pennsylvanian.

Stonefort Limestone Member type locality. 

Sec. 25, T. 10 S., R. 4 E., Harrisburg 15-min 

Quadrangle at Dunbar and Henbest Station 

371. Limestone above 3-4 ft of black platy 

shale with light-gray plastic clay shale at 

base and overlain by sandstone. C.L. Rice 

colln. KG-78-6. Wedekindellina euthysepta, 

Beedeina sp. aff. B. novamexicana.

3 ___f!3996 __Saline County. Middle Pennsylvanian.

Bankston Fork Limestone Member of Carbondale Formation near Henbest Station 233 

in 1- to 3-ft-thick limestone in old strip pit.
REGISTER OF LOCALITIES AND COLLECTIONS 15

REGISTER OF LOCALITIES AND COLLECTIONSContinued

Locality Collection

ILLINOIS-Continued

Sec. 3, T. 9 S., R. 6 E., 50 ft from south line 

and 600-1,000 ft from west line of Harrisburg Quadrangle. J. Beard, collector, 

1978. Beedeina sp.

4 ___f!3989 __Saline County. Middle Pennsylvanian.

Type "Curlew" of Illinois. West side of 

SWV4NWV4NWV4 sec. 27, T. 10 S., R. 6 E., 

Harrisburg 15-min Quadrangle. Dunbar and 

Henbest (1942) Station 234. C.L. Rice colln. 

KG-78-6. Beedeina leei.

5 ___f!3987 ___Saline County. Middle Pennsylvanian.

Creal Springs Limestone Member of Spoon 

Formation type locality. Old sandstone 

quarry in SWV4SWV4 sec. 30, T. 10 S., R. 3 

E., Harrisburg 15-min Quadrangle. C.L. Rice 

colln. KG-78-6. Wedekindellina euthysepta, 

Beedeina sp.

6 ___fl3999 ___Saline County. Middle Pennsylvanian.

Herrin Limestone Member at Henbest Station 377. Sec. 31, T. 9 S., R. 6 E., 1,900 ft 

from north line and 1,300 ft from west line of 

Harrisburg Quadrangle. 2Vz to 3 ft of 

limestone above water in old strip pit. J. 

Beard, collector, 1978. Beedeina sp. aff. B. 

haworthi.

7 ___f!4011 __Gallatin County. Upper Pennsylvanian.

In town limits of New Haven, along south 

bank of Wabash River. 1,200 ft from east line 

and 100 ft from south line of sec. 17, T. 7 S., 

R. 10 E., Emma 7V2-min Quadrangle. Shoal 

Creek Limestone Member of Bond Formation. J. Beard colln. KB-39. Triticites sp. aff. 

T. venustus.

________________INDIANA________________

8 f!3948 __Dubois County. Middle Pennsylvanian.

North side of road, 700 ft west and 6,500 ft 

south of northeast corner of Dale 7V2-min 

Quadrangle, just east of pipeline. C.L. Rice 

colln. RF-d from float blocks of Lead Creek 

Limestone Member of Mansfield Formation 

from excavation of pipeline trench? 

Millerella sp., Profusulinella kentuckyensis.

9 ___f!3949 __Spencer County. Middle Pennsylvanian.

Indiana Highway 545 roadcut in 

SWV4NEV4SWV4 sec. 11, T. 5 S., R. 4 W., 

Fulda 7V2-min Quadrangle, IVfe mi south of 

Fulda on northeast side of road intersection 

at 540-ft altitude. C.L. Rice colln. RF-1 

through 2V2 ft of Lead Creek Limestone 

Member. Profusulinella kentuckyensis. 

10 f!3947 Spencer County. Middle Pennsylvanian.

Limestone on west side of road at about 465 

ft altitude in NEV4 sec. 20, T. 6 S., R. 4 W., 

Lewisport 7V2-min Quadrangle. Probably 

same lociality as Shaver and Smith (1974)

REGISTER OF LOCALITIES AND COLLECTIONSContinued

Locality___Collection____________________________________

_____________INDIANA - Continued_____________

locality 28. Fulda Bed of Lead Creek 

Limestone Member. C.L. Rice colln. RF-2. 

Millerella sp., Profusulinella kentuckyensis. 

f 13980 __Spencer County. Middle Pennsylvanian.

1,850 ft east and 650 ft south of northwest 

corner sec. 20, T. 6 S., R. 4 W., Huff 

Township, in northeast corner of Lewisport 

7V2-min Quadrangle. Float block of Lead 

Creek Limestone Member at locality 28 of 

Shaver and Smith (1974, p. 34). C.L. Rice, 

collector, 197'8. Millerella sp., Profusulinella 

kentuckyensis.

_______________KENTUCKY_______________

11 ___f 13907 __Hancock County. Middle Pennsylvania.

Core hole Gil-1, Tell City Quadrangle, 

Kentucky-Indiana, approx. 6 mi southwest of 

Tell City, Ind. Approx. 4,100 ft from west 

boundary and 12,900 ft from south boundary 

of quadrangle. T.M. Kehn colln. KG-77-7. 

Core sample at depth of 119.4 ft. Millerella 

sp., Profusulinella sp.

f!3908 __Hancock County. Middle Pennsylvanian.

Same locality as f!3907, but core sample at 

depth of 120 ft. T.M. Kehn colln. KG-77-7. 

Millerella sp., Profusulinella kentuckyensis. 

 Hancock County. Middle Pennsylvanian.

Same locality as f!3907, but core sample at 

depth of 122.6 ft. T.M. Kehn colln. KG-77-7. 

Millerella sp., Profusulinella kentuckyensis.

f!3910 __Hancock County. Middle Pennsylvanian.

Same locality as f!3907, but core sample at 

depth of 145.2 ft. T.M. Kehn colln. KG-77-7. 

Millerella sp., Profusulinella kentuckyensis.

f!3911 __Hancock County. Middle Pennsylvanian.

Same locality as f!3907, but core sample at 

depth of 148.5 ft. T.M. Kehn colln. KG-77-7. 

Millerella sp., Profusulinella kentuckyensis.

12 ___f!4080 __Hancock County. Middle Pennsylvanian.

About 6,000 ft west of Petri, 700 ft south of 

U.S. Highway 60, 18,550 ft from south 

boundary and 14,200 ft from west boundary 

of Tell City 7V2 -min Quadrangle (GQ-356). 

Well started at 490-ft altitude. Sampled at 39 

ft. T.M. Kehn, collector, 1979. Millerella sp., 

Profusulinella kentuckyensis.

f!4081 __Hancock County. Middle Pennsylvanian.

Same locality as f!4080, but sectioned at 37 

ft. T.M. Kehn, collector, 1979. Millerella sp., 

Profusulinella kentuckyensis.

f!4082 __Hancock County. Middle Pennsylvanian.

Same locality as f!4080, but sectioned at 35 

ft. T.M. Kehn, collector, 1979. Millerella sp., 

Profusulinella kentuckyensis.

13 ___f!3922 __Daviess County. Middle Pennsylvanian.

Carter coordinates, sec. 7, Q-31, 1,250 ft 

from north line and 1,225 ft from east line.

f!3909 __.
16 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS

REGISTER OF LOCALITIES AND COLLECTIONSContinued

Locality Collection

KENTUCKY - Continued

About 4.5 mi southwest of Lewisport. 

Rockport and Lewisport Quadrangles. Core 

hold Gil-28, at 43.1 ft. T.M. Kehn colln. 

KG-77-7. Millerella sp., Profusulinella kentuckyensis.

f!3923 __Daviess County. Middle Pennsylvanian.

Same locality as f!3922, but depth of 46.3 ft. 

T.M. Kehn, colln. KG-77-7. Millerella sp., 

Profusulinella kentuckyensis.

f!3924 __Daviess County. Middle Pennsylvanian.

Same locality as f!3922, but depth of 

70.6-76.5 ft. T.M. Kehn, colln. KG-77-7. 

Millerella sp., Profusulinella kentuckyensis.

14 __f!3632 __Hancock County. Middle Pennsylvanian.

Lead Creek Limestone Member, 7,300 ft 

from south line and 15,400 ft from west line 

of Tell City Quadrangle. Approx. 3.75 mi 

west of Hawesville in roadcut of paved county road. Limestone 0-20 ft above Lead Creek 

coal bed (not exposed). T.M. Kehn colln. 

KB-7-1 from lower 6 in. of 1-ft limestone. 

Same locality as Thompson and Shaver 

(1964) locality 58. Millerella sp., Profusulinella kentuckyensis. 

f!3981 __Hancock County. Middle Pennsylvanian.

Carter coordinates sec. 25, R-33, 900 ft from 

north line and 1,800 ft from west line, Tell 

City 7V2-min Quadrangle. Ferdinand Bed of 

the Lead Creek Limestone Member, Tradewater Formation. C.L. Rice, collector, 1978. 

Millerella sp., Profusulinella kentuckyensis.

15 f!3921 ___Daviess County. Middle Pennsylvanian.

Carter coordinates sec. 1, N-27, 1,400 ft 

from north line and 550 ft from east line. 

About 13,700 ft from south and 550 ft from 

east boundary of Curdsville Quadrangle. 

Core hole Gil-27.

a) 203.6-204.0-ft depth.

b) 244.7-248.8-ft depth.

T.M. Kehn colln. KG-77-7. Beedeina sp.

16 f!3982 Union County. Upper Pennsylvanian.

8,000 ft from south line and 4,350 ft from 

west line of Uniontown Quadrangle boundary on north bank of Sibley Creek. Approx. 

6,000 ft southeast of east point of Wabash 

Island. Carter coordinates sec. 16, P-19, 

2,750 ft from south line, 16,000 ft from east 

line. Carthage Limestone Member of Sturgis 

Formation. T.M. Kehn colln. B-l. Triticites 

kehni.

17 __f 13629 __Union County. Middle Pennsylvanian.

Type locality for Curlew Limestone Member 

on south side of southeast end of Indian Hill, 

altitude 435 ft, approx. 1 mi southwest of 

Dekoven, 23,300 ft from north line and 

12,200 ft from west line of Dekoven 

Quadrangle boundary. Approx. 20 ft above 

No. 4 coal bed (not exposed) and 40 ft below

REGISTER OF LOCALITIES AND COLLECTIONS

Continued

Locality Collection

KENTUCKY - Continued

sandstone capping hill. Limestone about 3 ft 

thick. Sample from basal 1 ft. T.M. Kehn colln. KB-1. Fusulinella stouti, F. iowensis.

f!3896 __Union County. Middle Pennsylvanian.

South side of southwest end of Indian Hill, 

altitude 370 ft. Approx. 1 mi southwest of 

Dekoven. Curlew Limestone Member of 

Tradewater Formation, approx. 8 ft below 

sandstone capping hill. T.M. Kehn colln. 

KB-26. Fusulinella iowensis.

f!3897 __Union County. Middle Pennsylvanian.

East side of Indian Hill, 23,300 ft from north 

line and 12,200 ft from west line of Dekoven 

Quadrangle boundary. Lowest limestone exposed. About 6-8 in. of flaggy limestone 

below 3-ft limestone bed 30-35 ft below 

sandstone capping hill. T.M. Kehn colln. 

KB-l-a. Fusulinella stouti, F. iowensis.

f!3898 __Union County. Middle Pennsylvanian.

Same locality as above, but collected from 

3-ft limestone. T.M. Kehn colln. KB-l-b. 

Fusulinella stouti, F. iowensis.

f!3899 __Union County. Middle Pennsylvanian.

Same locality as f!3897, but collected from 

small patch of limestone about 2-ft square in 

saddle of ridge and about 30 ft above the 

other limestone. T.M. Kehn colln. KB-l-c. 

Fusulinella stouti, F. iowensis.

18 f!3743 __Union County. Middle Pennsylvanian.

2,100 ft from west line and 1,950 ft from

south line of Dekoven 7V2-min Quadrangle.

Altitude 414 ft. Drill hole Gil-15, depth

374-375 ft. C.L. Rice, collector, 1976.

Fusulinella iowensis. 

f!3744 __Union County. Middle Pennsylvanian.

Same locality as f!3743, but depth of 267 ft.

C.L. Rice, collector, 1976. Wedekindellina

euthysepta, Beedeina leei. 

f!3745 __Union County. Middle Pennsylvanian.

Same locality as f!3743, but depth of 65 ft.

C.L. Rice, collector, 1976. Wedekindellina

euthysepta, Beedeina sp. aff. B. novamexicana.

19 ___f!3876 __Union County. Upper Pennsylvanian.

Carter coordinates sec. 13, N-20, 300 ft from 

south line and 1,700 ft from east line in 

northwest corner of Bordley 7V2-min Quadrangle. Altitude 455 ft, total depth drilled 

1,841.05 ft, depth 194.4 ft. Limestone. T.M. 

Kehn, collector, 1977. Triticites beardi.

f!3877 __Union County. Upper Pennsylvanian.

Same locality as f!3876, but depth of 194.7 

ft. T.M. Kehn, collector, 1977. Triticites sp.

f!3878 __Union County. Upper Pennsylvanian.

Same locality as f!3876, but depth of 932.6 

ft. T.M. Kehn, collector, 1977. Triticites sp.

f!3879 __Union County. Upper Pennsylvanian.

Same locality as f!3876, but depth of 933.2 

ft. T.M. Kehn, collector, 1977. Triticites sp.
REGISTER OF LOCALITIES AND COLLECTIONS 17

REGISTER OF LOCALITIES AND COLLECTIONSContinued

Locality Collection

KENTUCKY - Continued

20 ___fL3633 __Crittenden County. Middle Pennsylvania.

Curlew Limestone Member of Tradewater 

Formation. Approx. 5.3 mi west of Providence, north bank of Kentucky Highway 

120, south side of Bald Hill, approx. 10,300 ft 

from south line and 4,600 ft from west line of 

Providence Quadrangle boundary. Same 

locality as No. 61 of Thompson and Shaver 

(1964). T.M. Kehn colln. KB-2-1 from 6 in. 

of fossililferous limestone at base of lower 

3-ft limestone. Fiisulinella stouti.

21 ___f!3925 __Webster County. Middle Pennsylvanian.

Carter coordinates sec. 3, K-21, 550 ft from 

east line and 2,400 ft from south line. About 

11,400 ft from south and 10,200 ft from east 

boundary of Providence Quadrangle. About 

7,500 ft northwest of Providence. Core hole 

Gil-31. Depth 398.1-398.4 ft. T.M. Kehn, 

collector, 1977. Wedekindellina euthysepta, 

Beedeina sp. aff. B. leei.

f!3926 __Webster County. Middle Pennsylvanian.

Same locality as f!3925, but depth of 

469.0-469.4 ft. T.M. Kehn, collector, 1977. 

Fusulinella iowensis.

f!3927 __Webster County. Middle Pennsylvanian.

Same locality as f 13925, but depth of 

471.5-471.7 ft. T.M. Kehn, collector, 1977. 

Fusulinella primotina.

f 13928 __Webster County. Middle Pennsylvanian.

Same locality as f!3925, but depth of 506.0 

ft. T.M. Kehn, collector, 1977. Millerella sp., 

Profusulinella kentuckyensis.

22 ___f!3977 __Webster County. Upper Pennsylvanian.

22,000 ft from north line and 16,000 ft from 

west line of Nebo Quadrangle boundary 

(GQ-777). Approx. 4 mi northeast of Providence. Limestone reported as 2-4 ft thick, 

partly exposed in sinkhole at altitude 410 ft. 

T.M. Kehn colln. 0-16-176. Triticites kehni. 

f!3984 __Webster County. Middle Pennsylvanian.

22,000 ft from north line and 15,900 ft from 

west line of Nebo Quadrangle boundary. 

Carter coordinates sec. 19, L-22, 2,200 ft 

from south line, 1,400 ft from west line. Approx. 4.75 mi northwest of Nebo. Carthage 

Limestone Member of Sturgis Formation 

about 6 ft thick, exposed in sinkhole. T.M. 

Kehn colln. F-178. Triticites kehni.

23 ___f!3915 __Hopkins County. Middle Pennsylvanian.

Carter coordinates sec. 22, J-21, 50 ft from 

north line and 2,100 ft from west line. About 

6,000 ft from south and 7,600 ft from east 

boundary of Dalton Quadrangle. Core hole 

Gil-17, sample from 230.2-ft depth. T.M. 

Kehn, collector, 1977. Wedekindellina sp.

REGISTER OF LOCALITIES AND COLLECTIONS

Continued

Locality Collection

KENTUCKY - Continued

24 ___f!3946 __Caldwell County. Middle Pennsylvanian.

Roadcut on north side of Route 293 across 

from Felker Cemetery. 17,600 ft from west 

line and 17,300 ft from south line of Dalton 

Quadrangle (GQ-490). C.L. Rice colln. JD 

from limestone in fault block about 100 ft 

above a coal bed. 1977. Millerella sp., Profusulinella sp.

25 ___f!3916 __Hopkins County. Middle Pennsylvanian.

Carter coordinates sec. 23, J-23, 2,025 ft 

from south line and 2,300 ft from west line. 

About 2,000 ft from south boundary and 200 

ft from east boundary of Coiltown 

Quadrangle. Core hole Gil-18, sample from 

283.9 ft depth. T.M. Kehn, collector, 1977. 

Wedekindellina sp., Beedeina sp.

26 ___f 13872 __Hopkins County. Middle Pennsylvanian.

Approx. 18,250 ft from north line and 5,600 

ft from west line of Madisonville East 

7V2-min Quadrangle. East side of Madisonville, 800 ft south of Center St. and adjacent 

to Louisville and Nashville Railroad tracks, 

west side of Madison Square shopping 

center. Madisonville Limestone Member of 

Sturgis Formation. Four samples in descending order through about 14 ft of basal part of 

member. T.M. Kehn colln. KB-16-4, 1-2-ft 

limestone at base of member below 2-ft shale 

break. Orange-weathering gray silty 

limestone. Beedeina acme. 

f!3873 __Hopkins County. Middle Pennsylvanian.

Approx. 15,800 ft from north line and 15,800 

ft from east line of Madisonville West 

7V2-min Quadrangle. In strip-mine highwall 

No. 14 coal bed, 2 mi west of Madisonville. 

Madisonville Limestone Member of Sturgis 

Formation lower beds. T.M. Kehn colln. 

KB-19-3. Lower 6-in. limestone below shale 

break in lower part of Madisonville 

Limestone Member. Beedeina sp.

27 ___f!3785 __Hopkins County. Middle Pennsylvanian.

Providence Limestone Member of Sturgis 

Formation from approx. 300 ft north of main 

intersection in Earlington and within the city 

limits. 400 ft from north line, 600 ft from 

west line of Madisonville West 7V2 min 

Quadrangle. Carter coordinates sec. 20, 

J-24. From 40-in. fossiliferous limestone just 

below No. 12 coal bed and 6 in. above top of 

No. 11 coal bed. T.M. Kehn colln. K224-2. 

Beedeina girtyi. 

f!3786 __Hopkins County. Middle Pennsylvanian.

Same locality as f!3785, but from 12-in. 

fossiliferous limestone above No. 12 coal bed, 

which is 14 in. thick. T.M. Kehn colln. 

K224-1. Beedeina sp.

28 ___f!3874 __Hopkins County. Middle Pennsylvanian.

Approx. 20,450 ft from north line and 9,400
18 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS

REGISTER OF LOCALITIES AND COLLECTIONS -

Continued

Locality___Collection_______

____________KENTUCKY - Continued____________

ft from east line of Madisonville East TVa-min 

Quadrangle. Approx. 4 mi east of Madisonville in high wall of Cimeron strip mine on 

Nos. 11, 12, and 13 coal beds. Providence 

Limestone Member of Sturgis Formation. 

T.M. Kehn colln. KB-20-1 from 3-ft limestone at base of formation. Beedeina girtyi, 

B. sp. aff. B. acme.

29 ___£13782 __Muhlenberg County. Middle Pennsylvanian.

USGS loc. 23770-PC (field no. P-19-270). 

Millport 7V2-min Quadrangle. Vogue Mine 

(13-J-26). Lat 37° 17.69' N., long 87°22.42' 

W. 1,900 ft from north line and 1,850 ft from 

east line. Millport 7V2-min Quadrangle. Providence Limestone Member of Sturgis Formation. G.J. Franklin, collector, 9-11-69. Beedeina girtyi.

f!3783 __Muhlenberg County. Middle Pennsylvanian.

USGS loc. 23771-PC (field no. P-19-273). 

Vogue Mine (13-J-26). Lat 37° 17.75' N., 

long 87°22.49' W. 1,530 ft from north line 

and 2,350 ft from east line, Millport 7V2-min 

Quadrangle. Providence Limestone Member 

of Sturgis Formation. G.J. Franklin, collector, 9-11-69. Beedeina sp.

f 13784 __Muhlenberg County. Middle Pennsylvanian. 

USGS loc. 23774-PC (field no. P-19-290). 

Vogue Mine (19-J-26). Lat 37° 16.81' N., 

long 87°21.68' W. 1,250 ft from north line 

and 1,650 ft from east line, Millport 7V2-min 

Quadrangle. Providence Limestone Member 

of Sturgis Formation. G.J. Franklin, collector, 9-23-69. Beedeina sp. aff. B. haworthi.

30 ___f 13903 __Muhlenberg County. Middle Pennsylvanian.

Carter coordinates sec. 13,1-27, 800 ft. from 

north line, 1,500 ft from east line, Graham 

7V2-min Quadrangle. Approx. 1 mi south of 

Yeargins Chapel, in roadbank (Kentucky 

Highway 175) and approx. 500 ft north of Illinois Central Railroad crossing. Yeargins 

limestone member (informal usage) below 

No. 6. coal. T.M. Kehn colln. KB-22. Beedeina sp.

31 ___£13740 __Christian County. Middle Pennsylvanian.

2,500 ft from north line and 150 ft from east 

line, Dawson Springs SE 7V2-min Quadrangle. Altitude 554 ft. Depth of 181-184 ft. 

Core hole DDK, G16. C.L. Rice, collector, 

1976. Profusulinella kentuckyensis.

£13741 __Christian County. Middle Pennsylvanian.

Same locality as £13740, but depth of 36-38 

ft. Core hole DDK, G16. C.L. Rice, collector, 

1976. Wedekindellina euthysepta, Beedeina 

sp. aff. B. spissiplicata.

f!3741B _Christian County. Middle Pennsylvanian.

Same locality as f!3740, but depth of 74.5 ft. 

C.L. Rice, collector, 1976. Fusulinella 

iowensis.

REGISTER OF LOCALITIES AND COLLECTIONS

Continued

Locality Collection

KENTUCKY - Continued

f!3742 __Christian County. Middle Pennsylvanian.

Same locality as f 13740, but depth of 4.1-4.3 

ft. Core hole DDK, G16. C.L. Rice, collector, 

1976. Wedekindellina euthysepta, Beedeina 

sp. aff. B. leei.

32 ___£13635 __Christian County. Middle Pennsylvanian.

Approx. 2,500 ft northwest of Empire. 

12,200 ft from north line and 1,800 ft from 

east line of Dawson Springs SE Quadrangle. 

From top of limestone 0-40 ft below No. 4 

coal bed. Limestone 5-15 ft. T.M. Kehn colln. 

KB-3 from lower 5 ft of fossiliferous 

limestone. Fusulinella stouti, Fusulinella 

sp., F. iowensis. 

f!3636 __Christian County. Middle Pennsylvanian.

Northwest of Empire, approx. 400 ft northwest of f 13635 (KB-3). 11,800 ft from north 

line and 2,450 ft from east line of Dawson 

Springs SE Quadrangle. "Curlew Limestone" 4 ft thick about 15 ft above base of No. 

4 coal bed and associated overlying shale. 

T.M. Kehn colln. KB-4, pieces of limestone 

and some picked fusulinids. Wedekindellina 

euthysepta, Beedeina leei. 

f 13983 __Christian County. Middle Pennsylvanian.

12,200 ft from north boundary and 2,600 ft 

from east boundary of Dawson Springs SE 

Quadrangle (GQ-1365). T.M. Kehn colln. 

B-2. Beedeina sp. aff. B. leei.

33 ___f!3917 __Muhlenberg County. Middle Pennsylvanian.

Carter coordinates sec. 15, 1-31, 1,550 ft 

from north line and 2,000 ft from east line. 

About 13,500 ft from north boundary and 

2,700 ft from west boundary of Rochester 

Quadrangle. Core hole Gil-19, 282.8-ft 

depth. T.M. Kehn colln. KG-77-7. 

Wedekindellina sp., Beedeina sp.

f!3918 __Muhlenberg County. Middle Pennsylvanian.

Same locality as f!3917, but depth of 284 ft. 

T.M. Kehn, collector, 1977. Beedeina sp.

f!3749 __Muhlenberg County. Middle Pennsylvanian.

3,080 ft from west line and 1,640 ft from 

south line, Rochester 7V2-min Quadrangle. 

Altitude 483 ft. Core hole DDH, G10, at 

depth of 293-296 ft. C.L. Rice, collector, 

1976. Profusulinella kentuckyensis.

f!3750 __Muhlenberg County. Middle Pennsylvanian.

Same locality as f!3749, but depth of 150 ft. 

C.L. Rice, collector, 1976. Beedeina sp. aff. 

B. spissiplicata.

34 ___f!3913 __Butler County. Middle Pennsylvanian.

Lead Creek Limestone Member of Tradewater Formation from core hole Gil-12 about 

4,000 ft southeast of Rochester. Carter coordinates sec. 13, 1-32, 22,800 ft from south 

line and 1,025 ft from west line. About 

16,000 ft from north boundary and 1,400 ft 

from east boundary of Rochester Quadrangle
REGISTER OF LOCALITIES AND COLLECTIONS 19

REGISTER OF LOCALITIES AND COLLECTIONSContinued

Locality Collection _ ____

____________KENTUCKY-Continued____________

(GQ-1171). At 147.6-ft depth. T.M. Kehn, 

collector, 1977. Millerella sp., Profusulinella 

sp. 

f 13914 __Butler County. Middle Pennsylvanian.

Same locality as £13913, but depth of 148.5 

ft. T.M. Kehn, collector, 1977. Millerella sp., 

Profusulinetta kentuckyensis.

35 ___f!3919 __Butler County. Middle Pennsylvanian.

Carter coordinates sec. 25, J-33, 1,700 ft 

from north line and 1,050 ft from west line. 

About 4,400 ft from south boundary and 

13,200 ft from west boundary of Cromwell 

Quadrangle. Core hole Gil-23, at depth of 

177.6-177.8 ft. T.M. Kehn, collector, 1977. 

Beedeina leei, B. sp. aff. B. spissiplicata. 

f!3920 __Butler County. Middle Pennsylvanian.

Same locality as f!3919, but depth of 371.2 

ft. T.M. Kehn, collector, 1977. Millerella sp., 

Profusulinetta kentuckyensis.

36 ___f!3630 __Butler County. Middle Pennsylvanian.

"Curlew Limestone" approx. 20 ft above No. 

4 coal bed, 1,500 ft from north line and 

18,500 ft from east line of Morgantown 

Quadrangle. West side of U.S. Highway 231, 

approx. 600 ft north of north end of highway 

bridge over Green River. Same as locality 

Ky-2 and Ky-3 of Thompson and Shaver 

(1964). T.M. Kehn colln. KB-5-1. 1975. Profusulinella kentuckyensis.

f!3631 __Butler County. Middle Pennsylvanian.

Same locality as f!3630, but lower 6 in. of 

3-ft limestone. T.M. Kehn, colln. KB-5-2. 

1975. Profusulinella kentuckyensis.

f!3900 __Butler County. Middle Pennsylvanian.

West side of U.S. Highway 231, approx. 600 

ft north of north end of bridge over Green 

River. Approx. 1.4 mi north of Morgantown. 

New exposures at locality KB-5 (f 13630-1). 

KB-5A from 6 in. limestone at base of principal limestone and separated from it by 2-3 

in. of clay shale. T.M. Kehn, collector, 1977. 

Millerella sp., Profusulinella kentuckyensis.

f 13901 __Butler County. Middle Pennsylvanian.

Same locality as f 13900, but KB-5B from upper limestone approx. 20 in. thick. T.M. 

Kehn, collector, 1977. Profusulinella kentuckyensis.

f!3994 __Butler County. Middle Pennsylvanian.

18,600 ft from north line and 15,850 ft from 

west line of quadrangle border. Approx. 

11,000 ft southeast of Pleasant Rock Church. 

Carter coordinates sec. 19,1-34, 500 ft from 

north line and 1,400 ft from west line of 

Morgantown 7V2-min Quadrangle. T.M. Kehn 

colln. KB-40. Profusulinella kentuckyensis.

f!4034 __Butler County. Middle Pennsylvanian.

Carter coordinates sec. 19, 1-34, 1,300 ft 

from west line and 800 ft from north line,

REGISTER OF LOCALITIES AND COLLECTIONS

Continued

Locality Collection

KENTUCKY - Continued

Morgantown 7Vz-min Quadrangle. Shale between the two limestone beds of Morgantown limestone (informal usage) at or near 

KB-40 (f!3994). J. Beard, collector, 1979. 

Millerella sp., Profusulinella kentuckyensis.

37 ___f!3746 __Butler County. Middle Pennsylvanian.

1,730 ft from east line and 1,520 ft from 

south line, South Hill 7V2-min Quadrangle, 

altitude 535 ft. Depth 168 ft. C.L. Rice, collector, 1976. Millerella sp., Profusinella kentuckyensis.

38 ___f!3747 __Ohio County. Middle Pennsylvanian.

2,700 ft from south line and 500 ft from east 

line, Flener 7V2-min Quadrangle. Altitude 

570 ft. Depth of 199 ft. C.L. Rice, collector, 

1976. Millerella sp., Profusulinella kentuckyensis. 

f!3748 __Ohio County. Middle Pennsylvanian.

Same locality as f!3747, but depth of 143 ft. 

C.L. Rice, collector, 1976. Millerella sp., 

Projusulinetta kentuckyensis.

39 ___f!3904 __Butler County. Middle Pennsylvanian.

Approx. 2,200 ft south of Green River Parkway bridge over Green River and approx. 

4,400 ft from south boundary and 1,200 ft 

from west boundary of Flener Quadrangle 

(GQ-1049). T.M. Kehn, colln. KB-23. 

Beedeina sp. aff. B. novamexicana. 

f!3905 __Butler County. Middle Pennsylvanian.

East bank of Green River Parkway. Approx. 

1.4 mi south of bridge over Green River, 

1,200 ft south of Kentucky Highway 403 

bridge over parkway. Flener Quadrangle. 

Mapped as near No. 6 coal. T.M. Kehn colln. 

KB-25A. Wedekindellina euthysepta. 

f!3906 __Butler County. Middle Pennsylvanian.

Same locality as f!3905, but in 1-2-ft 

limestone 3V2 ft above KB-25A. T.M. Kehn, 

colln. KB-25B. Wedekindellina euthysepta, 

Beedeina sp. aff. B. leei.

40 ___f 13751 __Ohio County. Middle Pennsylvanian.

2,600 ft from south line and 2,250 ft from 

west line, Cromwell 7V2-min Quadrangle. 

Core Gil-9 at depth of 332 ft. C.L. Rice, collector, 1976. Beedeina sp. aff. B. 

spissiplicata. 

f!3752 __Ohio County. Middle Pennsylvanian.

Same locality as f!3751, but depth of 

313-316 ft. C.L. Rice, collector, 1976. 

Wedekindellina euthysepta, Beedeina sp. aff. 

B. novamexicana. 

f!3753 __Ohio County. Middle Pennsylvanian.

Same locality as f 13751, but depth of 185 ft. 

C.L. Rice, collector, 1976. Beedeina leei.

41 ___f!3902 __Ohio County. Middle Pennsylvanian.

Carter coordinates sec. 14, K-33, 500 ft from 

north line and 600 ft from west line. North 

bank of Western Kentucky Parkway. Ap-
20 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS

REGISTER OF LOCALITIES AND COLLECTIONSContinued

Locality___Collection_______ ____

____________KENTUCKY - Continued____________

prox. 4,000 ft east of parkway service plaza. 

Horton Quadrangle. Limestone below No. 6 

coal bed. T.M. Kehn colln. KB-21. 

Wedekindellina euthysepta, Beedeina sp. aff. 

B. novamexicana.

42 ___f!4098 __Ohio County. Middle Pennsylvanian.

West side of Green River Turnpike at 

altitude 480 ft. Carter coordinates sec. 16, 

J-34, 3,000 ft from north line and 1,100 ft 

from west line; at 9,100 ft from south line 

and 1,100 ft from west line of Cromwell 

Quadrangle. J. Beard colln. 79-11. Beedeina 

sp.

43 ___f!4097 __Ohio County. Middle Pennsylvanian.

On Green River Turnpike 1,700 ft northwest 

of intersection. West U.S. 62 on southwest 

side of turnpike. Carter coordinates sec. 21, 

L-32, 1,500 ft from west line and 1,650 ft 

from north line; 8,850 ft from west line and 

19,500 ft from south line of Horton 

Quadrangle. J. Beard colln. 79-12. 

Fusulinella iowensis.

44 ___f!3912 __Ohio County. Middle Pennsylvanian.

Core hole Gil-6. Carter coordinates sec. 24, 

N-32, 850 ft from north line and 600 ft from 

east line. About 9,900 ft from north boundary and 3,000 ft from east boundary of Pleasant Ridge Quadrangle (GQ-766). Sample 

from 45.4-ft depth in Lead Creek Limestone 

Member of Tradewater Formation. T.M. 

Kehn, collector, 1977. Projusulinella sp.

45 ___f!4099 __Bell County. Middle Pennsylvanian.

Core from drill hole 11,850 ft east and 19,350 

ft north of southwest corner of Beverly 

7V2-min Quadrangle. Sample from 56 ft 

above Hazard Coal bed. C.L. Rice colln. 

79-10. Fusulinella pria.

46 ___f!3574 __Bell County. Middle Pennsylvanian.

Kentucky coordinates 211,250 N., 2,640,550 

E., Beverly Quadrangle, on strip bench west 

of Kentucky Highway 1201, about 2Vi mi 

southwest of Beverly at head of Red Bird 

Creek. Russell Ping colln. LC3-2. Jan. 19, 

1975. Fusulinella pria.

f!3570 __Bell County. Middle Pennsylvanian.

Russell Ping colln. LC3-3 at same locality 

but stratigraphically higher in bed. 

Fusulinella pria.

f!3571 __Bell County. Middle Pennsylvanian.

Russell Ping colln. LC3-4 at same locality 

but stratigraphically higher within the 59-in. 

bed. Fusulinella pria.

f!3572 __Bell County. Middle Pennsylvanian.

Russell Ping collns. LC3-5, LC3-6 at same 

locality but stratigraphically higher in bed. 

Fusulinella pria.

f!3573 __Bell County. Middle Pennsylvanian.

Russell Ping colln. LC3-7 at same locality, 

but at top of limestone bed. Fusulinella pria.

REGISTER OF LOCALITIES AND COLLECTIONSContinued

Locality___Collection____________________________________

____________KENTUCKY - Continued____________

f!3788 __Bell County. Middle Pennsylvanian.

to Kentucky coordinates 211,250 N., 2,640,550 

f!3794 E., Beverly Quadrangle, on strip bench west 

of Kentucky Highway 1201, about 2V2 mi 

southwest of Beverly at head of Red Bird 

Creek. Samples through 30 ft in ascending 

order. C.L. Rice, collector, 1976. Fusulinella 

pria.

47 ___f!3855 __Clay County. Middle Pennsylvanian.

to Stoney Fork Member from drill core at Bell 

f!3866 Lookout Tower, 4,400 ft east and 900 ft 

south of northwest corner of Beverly 7V2-min 

Quadrangle. BFT-1 (f!3855) at bottom to 

BFT-11 (f 13866) at top. Base of 48.5 ft of 

limestone at 1,899-ft altitude C.L. Rice, collector, 1977. Fusulinella pria.

48 ___f!3585 __Leslie County. Middle Pennsylvanian.

Carter coordinates sec. 21, H-74, 1,100 ft 

from north line and 1,450 ft from west line, 

Cutshin 7V2-min Quadrangle. Strip bench at 

altitude of 1,715 ft above sea level on divide 

between John Creek and Camp Creek of Middle Fork of Kentucky River, 1,800 ft northwest of hill marked 1,725 ft on topographic 

map. Locality 4-3. Top is 5 in. from top of 

limestone. C.L. Rice, collector, 1975. 

Fusulinella pria.

f!3586 __Leslie County. Middle Pennsylvanian.

Same area as f!3585, but locality 4-4. Top is 

7 in. from top of limestone. C.L. Rice, collector, 1975. Mitterella sp.

f!3587 __Leslie County. Middle Pennsylvanian.

Same area as f 13585, but locality 4-5. Top is 

3 in. from top of limestone. C.L. Rice, collector, 1975. Fusulinella sp.

f!3588 __Leslie County. Middle Pennsylvanian.

Same area as f!3585, but locality 4-6. About 

2 in. from top of limestone. C.L. Rice, collector, 1975. Fusulinella sp.

f!3589 __Leslie County. Middle Pennsylvanian.

Same area as f!3585, but locality 5-3. Middle 

of limestone. C.L. Rice, collector, 1975. 

Fusulinella sp.

f!3590 __Leslie County. Middle Pennsylvanian.

Same area as f 13585, but locality 5-4. Top of 

limestone. C.L. Rice, collector, 1975. 

Fusulinella pria.

49 ___f!4028 __Clay County. Middle Pennsylvanian.

11,600 ft east and 9,650 ft north of northwest corner of Creekville 7V2-min 

Quadrangle. Altitude 1,675 ft. Unnamed 

limestone at depth of about 96 ft and about 

173 ft above Magoffin Member of Breathitt 

Formation. C.L. Rice, collector, 1978. 

Fusulinella pria. 

f 14336 __Clay County. Middle Pennsylvanian.

Lat 37°03'12" N., long 83°37'17" W., northwest of Mudlick Church at head of Mudlick
REGISTER OF LOCALITIES AND COLLECTIONS 21

REGISTER OF LOCALITIES AND COLLECTIONSContinued

Locality Collection

KENTUCKY - Continued

50 __J13591

to 

£13596

Fork, Creekville TVa-min Quadrangle. G.W. 

Cumbee, collector, 1980. Fusulinella pria. 

___Leslie County. Middle Pennsylvanian.

On strip bench at altitude of 1,620+ ft on 

divide between Big Branch of Bull Creek and 

Cucumber Branch of Hell For Certain Creek, 

both of Middle Fork of Kentucky River, 

2,400 ft east of hill marked 1,748 ft on 

Hayden West 7V2-min Quadrangle. C.L. Rice 

colln. LC 7-1 (f!3591) through LC 7-6 

(fl3596). June 1975. Fusulinella pria.

51 ___f!3828 __Perry County. Middle Pennsylvanian.

Limestone from upper 2 ft of Stoney Fork 

Member of Breathitt Formation on strip 

bench just east of BM 1230 on Kentucky 

Highway 28, 2,300 ft west and 17,100 ft 

south of northeast corner of Buckhorn 

7V2-min Quadrangle. C.L. Rice colln. BIB. 

KG-76-7. Fusulinella pria.

52 ___f!3597 __Breathitt County. Middle Pennsylvanian.

Carter coordinates sec. 11, L-74, 700 ft from 

north line and 50 ft from east line, Haddix 

7V2-min Quadrangle. Type locality of Stoney 

Fork Member of Breathitt Formation on 

divide between Lost Creek and Big Branch 

of North Fork of Kentucky River. Just north 

of gap through which road connecting the 

two streams passes, along old road that intersects road through gap, and just north of 

the second cemetery at altitude of about 

1,210 ft above sea level. The Stoney Fork occurs just above Hindman coal bed but was 

found only as loose boulders and small rocks. 

C.L. Rice, colln. LC-8. Fusulinella pria. 

 Morgan County. Middle Pennsylvanian.

From north-facing wall about 14 ft above 

Nickell coal bed at 1,070-ft altitude, 7,725 ft 

west and 2,750 ft south of northeast corner 

of Cannel City Quadrangle, lat 37°52'03" N. 

long83°01'35"W. Limestone containing some 

silicified fossils. Breathitt Formation. C.L. 

Rice, collector, June 1976. Fusulinella pria.

f!3985 _Morgan County. Middle Pennsylvanian.

Stoney Fork Member of Breathitt Formation. 6,300 ft east and 11,000 ft north of 

southeast corner of West Liberty 7V2-min 

Quadrangle. C.L. Rice colln. KG-78-6. 

Fusulinella pria.

f!3986 Morgan County. Middle Pennsylvanian.

Stoney Fork Member of Breathitt Formation. 5,300 ft west and 1,100 ft south of 

northeast corner of Cannel City 7V2-min 

Quadrangle, about 20 ft up in high wall of 

strip mine. About 125 ft above Magoffin 

Member of Breathitt Formation. C.L. Rice 

colln. KG-78-6. Fusulinella pria.

f!4306 Morgan County. Middle Pennsylvanian.

See f!3767 for locality. Re-collection of C.L.

53 ___f!3767 __.

REGISTER OF LOCALITIES AND COLLECTIONSContinued

Locality Collection _____ _______

____________KENTUCKY - Continued____________

Rice, 1976, locality. C.L. Rice, R.C. 

Douglass, and R. Margerum colln. 9-9-80-1. 

Fusulinella pria.

54 ___f!4185 __Lawrence County. Upper Pensylvanian.

Divide between Lost Creek and Big Branch 

of North Fork of Kentucky River. Just north 

of gap through which road connecting the 

streams passes and just north of second 

cemetery, at altitude 1,210 ft. Carter coordinates sec. 11, L-74, 700 ft from north line 

and 50 ft from east line, Haddix 71/2-min 

Quadrangle. G. Merrill, collector, 1979. 

Triticites ohioensis.

f!4314 __Lawrence County. Upper Pennsylvanian.

Limestone about 6 in. thick, roadcut on west 

side of U.S. Highway 23 about % mi south of 

bridge over Bear Creek. Approx. same as 

locality f!4185 of G. Merrill. C.L. Rice, R.C. 

Douglass, and R. Margerum colln. 9-10-80. 

Re-collected by Rice. Triticites ohioensis.

f!4315 __Lawrence County. Upper Pennsylvanian.

Approx. same locality as f!4314. C.L. Rice, 

R.C. Douglass, and R. Margerum colln. 

9-10-80. Triticites ohioensis.

55 ___f!3990 __Carter County. Middle Pennsylvanian.

Vanport Limestone as used by Phalen (1912). 

7,500 ft west and 13,250 ft south of northeast corner of Willard 7V2-min Quadrangle. 

C.L. Rice colln. KG-78-6. Beedeina sp.

56 ___f!3991 __Carter County. Upper Pennsylvanian.

Carter coordinates sec. 20, U-79. 1,250 ft 

from south line and 1,950 ft from east line at 

altitude 950 ft, Webbville 7V2-min Quadrangle. Cores from depth of 37.5-38 ft 

through Brush Creek Limestone Member. 

C.L. Rice colln. KG-78-7. Triticites ohioenf!3992 __Carter County. Upper Pennsylvanian.

Same locality as f!3991, but depth of 41.7 ft. 

C.L. Rice colln. KG-76-7. Triticites ohioensis.

f!3993 __Carter County. Upper Pennsylvanian.

Same locality as f!3991, but depth of 33.7 ft 

in Brush Creek Limestone. C.L. Rice colln. 

KG-78-7. Triticites ohioensis.

57 f!3584 Boyd County. Upper Pennsylvanian.

Carter coordinates sec. 19, V-81, 3,000 ft 

from north line and 1,600 ft from east line, 

Rush 7V2-min Quadrangle. From rock quarry 

2,200 ft southwest of Long Branch School on 

southwest side of drainage running southeast into Long Branch and about 1,500 ft 

north of Clybur Cems [sic]. Brush Creek 

Limestone Member of Conemaugh Formation. C.L. Rice, collector, 1975. Triticites 

ohioensis.

58 f!3576 Boyd County. Upper Pennsylvanian.

"Fusulina" limestone, Boltsfork Quadrangle.
22

REGISTER OF LOCALITIES AND COLLECTIONSContinued

Locality___Collection

____________KENTUCKY - Continued____________

Left Fork of Friendship Creek, east of Mavity. J.W. Huddle, collector. Triticites ohioensis. 

f!4309 __Boyd County. Upper Pennsylvanian.

At 840-ft altitude in roadcut on west side of 

north/south road about IVz mi east of Mavity, 

connecting Dog Fork with Friendship Creek, 

Boltsfork 7V2-min Quadrangle. C.L. Rice, 

R.C. Douglass, and R. Margerum colln. 

9-10-80-1. Triticites ohoioensis.

59 ___f!3120 __Boyd County. Upper Pennsylvanian.

Lower Conemaugh Formation. Boltsfork 

7V2-min Quadrangle, along secondary road 

between Friendship Creek and Dog Fork, 

about ¥4 mi south of gas-field boundary, unit 

25 of measured section. E. Dobrovolny and 

T. Knous, collectors, 3-3-62. Triticites 

ohioensis.

f!3122 __Boyd County. Upper Pennsylvanian.

"Fusulina" Limestone. Boltsfork Quadrangle. Roadcut on left fork of Friendship 

Creek, east of Mavity. E. Dobrovolny and T. 

Knous, collectors, 3-3-62. Triticites ohioensis.

f24987 __Boyd County. Upper Pennsylvanian.

8,400 ft east of road junction 617 at Mavity, 

Boltsfork Quadrangle. D.E. Wolcott colln. 

BB-2, 1966. Triticites ohioensis.

60 ___f!4311 __Boyd County. Upper Pennsylvanian.

North side of Dog Fork just east of powerline 

crossing and about lllz mi east-northeast of 

Mavity. Boltsfork Quadrangle. C.L. Rice, 

R.C. Douglass, and R. Margerum colln. 

9-10-80-3. Triticites ohioensis.

f!4312 __Boyd County. Upper Pennsylvanian.

Brush Creek Limestone Member in abandoned quarry in ravine about 4/10 mi west of 

site of Long Branch School shown on Rush 

Quadrangle map. 6-in. limestone bed. 

Altitude approx. 760 ft. C.L. Rice, R.C. 

Douglass, and R. Margerum colln. 

9-10-80-4. Triticites ohioensis.

f!4313 __Boyd County. Upper Pennsylvanian.

Same locality as f!4312, but about 10 ft 

lower in section. C.L. Rice, R.C. Douglass, 

and R. Margerum colln. 9-10-80-5. 

Triticites ohioensis.

61 ___f!4308 __Greenup County. Middle Pennsylvanian.

Upper Allegheny Formation along powerline 

and just north of Ashland Coal Company office in Riverview area on bluff overlooking 

Ohio River. Southeast corner of Ironton 

Quadrangle. C.L. Rice, R.C. Douglass, and 

R. Margerum colln. 9-9-80-3. Beedeina 

ashlandensis

_________________OHIO_________________

62 ___f!3787 __Lawrence County. Middle Pennsylvanian.

REGISTER OF LOCALITIES AND COLLECTIONSContinued

Locality Collection ______________

_______________OHIO-Continued_______________

Upper Allegheny Formation, 3,200 ft west 

and 5,400 ft south of northeast corner 

Ashland 7V2-min Quadrangle, Kentucky and 

Ohio. About 4 ft above and 20 ft northeast of 

U.S. Highway 52. Limestone 3V2 ft thick. 

C.L. Rice, collector, 1976. Beedeina 

ashlandensis.

63 ___f!4307 __Lawrence County. Middle Pennsylvanian.

Upper Allegheny Formation north side of 

southbound lanes of U.S. Highway 52, 0.5 mi 

east of 12th Street bridge over Ohio River. 

Re-collection of Rice fl3787. Ashland 

Quadrangle. C.L. Rice, R.C. Douglass, and 

R. Margerum colln. 9-9-80-2. Beedeina 

ashlandensis.

64 ___f!4316 __Gallia County. Upper Pennsylvanian.

Southeast of Patriot along Hannan Trace in 

NWV4NEV4 sec. 2, T. 4 N., R. 16 W., Rodney 

7V2-min Quadrangle. Shaley limestone at 

base of 18-in. limestone bed in small roadcut 

on northeast side of road. C.L. Rice, R.C. 

Douglass, and R. Margerum colln. 

9-11-80-1. Triticites ohioensis.

f!4317 __Gallia County. Upper Pennsylvanian.

Southeast of Patriot on Patriot-Cadmus 

Road near center SEV4 sec. 35, T. 5 N., R. 16 

W., Rodney 7V2-min Quadrangle. Just south 

of road in streambed. Nodular limestone 

below on 18-20-in. limestone. Cambridge 

Limestone Member of Smyth (1957), locality 

7. C.L. Rice, R.C. Douglass, and R. 

Margerum colln. 9-11-80-2. Triticites ohioensis.

£14318 __Gallia County. Upper Pennsylvanian.

Near center sec. 35, T. 6 N., R. 16 W., 

Rodney 7V2-min Quadrangle. Small outcrop 

along Garners Cora Road northwest of 

Garners Ford Bridge over Raccoon Creek. 

C.L. Rice, R.C. Douglass, and R. Margerum 

colln. 9-11-80-5. Triticites ohioensis.

65 ___f!3997 __Gallia County. Upper Pennsylvanian.

Northeast bank of Hannan Trace Road, 750 

ft east and 5,000 ft north of southwest corner 

of Rodney 7V2-min Quadrangle. Near Smyth 

(1957) locality 7. Cambridge limestone and 

shale. C.L. Rice, collector, 1978. Triticites 

ohioensis.

66 ___f!4004 __Gallia County. Upper Pennsylvanian.

2,400 ft east and 5,050 ft south of northwest 

corner of Rodney 7V2-min Quadrangle. 4-ft 

limestone on north side of Garners Cora 

Road at locality 15 of Smyth (1957). C.L. 

Rice, collector, 1978. Brush Creek Limestone Member in two parts, top and bottom 

of bed. Triticites ohioensis. 

f!4005 __Gallia County. Upper Pennsylvanian.

4,300 ft south and 14,555 ft west of northeast corner of Vinton 7V2-min Quadrangle at 

about 855-ft altitude. 1-2 ft of Ames
REGISTER OF LOCALITIES AND COLLECTIONS 23

REGISTER OF LOCALITIES AND COLLECTIONSContinued

Locality Collection

OHIO-Continued

Limestone Member in roadbank and road at 

Smyth (1957) locality 5. C.L. Rice, collector, 

1978. Triticites ricei.

67 ___£14509 __Hocking County. Middle Pennsylvanian.

Boggs Limestone Member of Pottsville Formation from exposure on east side of U.S. 

Highway 93, approx. 0.4 mi north of Mt. 

Pleasant, SWV4 sec 34, Washington 

Township, New Plymouth 7V2-min Quadrangle. R. Hoare colln. Hwa-1. Profusulinella ohioensis, Fusulinella imprima.

68 ___f!4319 __Athens County. Upper Pennsylvanian.

East of Chauncey in NWV4SEV4 sec. 8, T. 10 

N., R. 14 W., Jacksonville 7V2-min Quadrangle. Ames? Limestone Member along 

tributary to Mill Creek along road to Brown 

Cemetery at about 770-ft altitude. Obscure 

outcrop in west bank of road. C.L. Rice, R.C. 

Douglass, and R. Margerum colln. 

9-11-80-4. Tricities skinneri, T. ohioensisl 

f!4320 __Athens County. Upper Pennsylvanian.

Approx. same locality as f!4319, but about 

18 ft higher stratigraphically and possibly 

representing Gaysport Limestone Member. 

C.L. Rice, R.C. Douglass, and R. Margerum 

colln. 9-11-80-5. Triticites skinneri.

69 ___f9719 ___Perry County. Upper Pennsylvanian.

Cambridge Limestone Member. Bearfield 

Township. West-central sec. 24. This locality 

was described by Condit (1912, p. 126-127). 

Pauline Smyth, collector, 9-8-53. Triticites 

smythi, T. ohioensis. 

f9720 ___Perry County. Upper Pennsylvanian.

Ames Limestone Member. Bearfield Township. West-central sec. 24. Locality was 

described by Condit (1912, p. 126-127). 

Ames Limestone Member here was source of 

type specimens for Triticites skinneri (Journal of Paleontology, v. 10, p. 683). Pauline 

Smyth, collector, 9-8-53. Triticites skinneri.

70 ___f!3833 __Muskingum County. Middle Pennsylvanian.

Boggs Limestone Member of Pottsville Formation from creek bottom of Symmes Creek 

just above junction with North Branch, 

13,050 ft east and 15,200 ft south of northwest corner of Adamsville 7V2-min 

Quadrangle. C.L. Rice colln. RO-11. Profusulinellal sp.

71 ___f!3817 __Muskingum County. Middle Pennsylvanian.

to North side of U.S. Highway 1-70, underpass, 

f!3822 18,700 ft west and 19,000 ft south of northeast corner of Zanesville West Quadrangle. 

Vanport Limestone Member as identified by 

R.M. Delong. T-l at base to T-24 at top of 

limestone. C.L. Rice colln. KG-76-8. Beedeina carmani.

72 ___f!4146 __Muskingum County. Upper Pennsylvanian.

REGISTER OF LOCALITIES AND COLLECTIONSContinued

Locality___Collection_______________________________________

_______________OHIO-Continued_______________

Ames Limestone Member. East side of Ohio 

Route 83, 0.3 mi south of junction with U.S. 

40 in New Concord. R.C. Douglass, collector, 

5-13-79. Triticites skinneri. 

f!4148 __Muskingum County. Middle Pennsylvanian.

Middle Mercer Limestone (local usage). At 

Cement Plant (Columbia) south of Zanesville 

off U.S. Route 22, about 3 mi southwest of 

junction with Ohio Route 93. R.C. Douglass, 

collector, 5-13-79. Millerella sp., 

Fusulinella iowensis.

73 ___f 14000 __Muskingum County. Middle Pennsylvanian.

2,400 ft south and 13,550 ft east of northwest corner of Dresden 7V2-min Quadrangle. 

1-2 in. of fossiliferous limestone "paves" entrance to small field and cemetery. Layer 

seems to grade into siltstone but may have 

been brought in for road ballast. All material 

seems to be similar. C.L. Rice colln. 2B. 

1978. Beedina sp. aff. B. leei. 

f!4001 __Muskingum County. Middle Pennsylvanian.

2,400 ft south and 13,550 ft east of northwest corner of Dresden 7V2-min Quadrangle. 

Northwest of road and opposite southtrending driveway in massive fossiliferous 

silty limestone in place. C.L. Rice colln. 2A, 

about 12 ft northeast of colln. 2B. 1978. Beedeina sp. 

f!4002 __Muskingum County. Middle Pennsylvanian.

1,350 ft south and 14,550 ft east of northwest corner of Dresden 7V2-min Quadrangle 

on southeast side of road and about 25 ft 

above Vanport locality 40 of Smyth (1957). 

Nodular limestone, broken and slumped into 

thick whitish underclay. C.L. Rice, collector, 

1978. Beedeina sp.

74 ___f!3823 __Muskingum County. Middle Pennsylvanian.

WV2 sec. 3, T. 2 N., R. 7 W., Dresden 7V2 -min 

Quadrangle. 12,400 ft north and 1,000 ft 

west of southeast corner of quadrangle. 

Lower Mercer Limestone Member at M.L. 

Thompson's locality on Blunt Run. Tl at base 

to T9 at top of limestone. C.L. Rice colln. 

RO-4-T1-T3. Fusulinella! sp.

f!3824 __Muskinghum County. Middle Pennsylvanian. 

Same locality as f!3823. C.L. Rice colln. 

RO-4T4-T5. Fusulinella iowensis.

f!3825 __Muskingum County. Middle Pennsylvanian.

Same locality as f!3823. C.L. Rice colln. 

RO-4-T6-T8. Fusulinella iowensis.

f!3826 __Muskingum County. Middle Pennsylvanian.

Same locality as f!3823. C.L. Rice colln. 

RO-4-T9. Fusulinella iowensis.

f!3829 __Muskingum County. Middle Pennsylvanian.

Boggs Limestone Member of Pottsville Formation in Blunt Run, 11,575 ft north and 300 

ft west of southeast corner of Dresden 

7V2-min Quadrangle. C.L. Rice colln. RO-8-1
REGISTER OF LOCALITIES AND COLLECTIONSContinued

Locality Collection

OHIO-Continued

at base through RO-8-11 at top. RO-8-3, 

RO-8-4. Fusulinella imprimal

f!3830 __Muskingum County. Middle Pennsylvanian.

Same locality as f!3829. C.L. Rice colln. 

RO-8-5. Fusulinella imprimal

f 13831 __Muskingum County. Middle Pennsylvanian.

Same locality as f!3829. C.L. Rice colln. 

RO-8-6-11. Fusulinella sp.

£13832 __Muskingum County. Middle Pennsylvanian.

Upper Mercer Limestone Member in Blunt 

Run. 12,700 ft north and 1,100 ft west of 

southeast corner of Dresden TVz-min 

Quadrangle. C.L. Rice colln. RO-8-9-b. 

Fusulinella iowensis.

75 ___f9717 ___Muskingum County. Middle Pennsylvanian.

Vanport Limestone Member. Hopewell 

Township. NE sec. 15 N. The place where 

Thompson collected the types for 

Fusulinella carmani. Pauline Smyth, collector, 9-8-53.

76 ___f!3815 __Muskingum County. Middle Pennsylvanian.

Same as locality 73 of Smyth (1957), in 

center partial sec. 18, T. 1 N., R. 9 W., 

Gratiot Quadrangle, in center roadside ditch 

about 1 mi north of Mt. Sterling. 1,050 ft east 

of west line and 2,025 ft south of north line of 

sec. 18, Hopewell Township. Lower Mercer 

Limestone Member of Pottsville Formation. 

C.L. Rice, colln. KG-76-8. Fusulinella 

iowensis. 

f!3816 __Muskingum County. Middle Pennsylvanian.

Same locality as f!3815. C.L. Rice colln. 

KG-76-8. Fusulinella iowensis.

77 ___f!4006 __Licking County. Middle Pennsylvanian.

950 ft east and 1,300 ft south of northwest 

corner sec. 20, Hopewell Township, Gratiot 

7V2-min Quadrangle. C.L. Rice, collector, 

1978. Vanport R-l in float blocks of chert. 

Beedeina sp.

f!4007 __Licking County. Middle Pennsylvanian.

1,800 ft west and 2,650 ft south of northeast 

corner sec. 11, Hopewell Township, Gratiot 

7V2-min Quadrangle. C.L. Rice, collector, 

1978. Vanport R-2 in cherty limestone. Beedeina sp.

f!4147 __Licking County. Middle Pennsylvanian.

Lower Mercer Limestone Member. U.S. 

Route 40 about 3.1 mi west of Brownsville. 

4-ft thick, exposed 10 ft above highway on 

north side of roadcut at roadside park. R.C. 

Douglass, collector, 5-13-79. Fusulinella 

iowensis.

78 ___f9718 ___Muskinghum County. Middle Pennsylvanian. 

Vanport Limestone Member. Cass Township 

on The Highlands, 1% mi north of Shannon. 

Pauline Smyth collector, 9-8-53. 

Wedekindellina euthysepta, Beedeina sp.

REGISTER OF LOCALITIES AND COLLECTIONSContinued

Locality Collection

______________OHIO-Continued______________

f!3847 __Muskingum County. Middle Pennsylvanian.

Vanport Limestone Member of Allegheny 

Formation at Smyth's (1957) locality 40, at

I,125 ft south and 15,250 ft east of northwest corner of Dresden 7V2-min Quadrangle 

on north side of road. C.L. Rice colln. 

RO-13-^- Wedekindellina sp.

f!3848 __Muskingum County. Middle Pennsylvanian.

Same locality as f!3847. C.L. Rice colln. 

RO-13-2 to RO-13-4. Wedekindellina sp.

f!3849 __Muskingum County. Middle Pennsylvanian.

Same locality as f!3847. C.L. Rice colln. 

RO-13-5. Wedekindellina euthysepta.

f!3850 __Muskingum County. Middle Pennsylvanian.

Same locality as f!3847. C.L. Rice colln. 

RO-13-6. Wedekindellina sp.

f!3851 __Muskingum County. Middle Pennsylvanian.

Same locality as f!3847. C.L. Rice collns. 

RO-13-7, RO-13-8. Wedekindellina sp.

f!3852 __Muskingum County. Middle Pennsylvanian.

Same locality as f!3847. C.L. Rice colln. 

RO-13-9. Wedekindellina euthysepta, 

Beedeina sp.

f!3853 __Muskingum County. Middle Pennsylvanian.

Same locality as f!3847. C.L. Rice colln. 

RO-13-10. Wedekindellina sp.

f 13854 __Muskingum County. Middle Pennsylvanian.

Columbiana Limestone Member of Allegheny Formation about 22 ft above Vanport 

Limestone Member, 4,900 ft south and

II,200 ft east of northwest corner of 

Dresden 7V2-min Quadrangle, on east side of 

road. C.L. Rice, colln. RO-12. Wedekindellina sp, Beedeina sp. 

79 ___f!3836 __Coshocton County. Middle Pennsylvanian.

Putnam Hill Limestone Member of Allegheny Formation at Smyth's (1957) locality 

54, at 6,050 ft east and 21,600 ft north of 

southwest corner of Trinway 7V2-min 

Quadrangle, on east side of road. C.L. Rice 

colln. RO-14, samples 1 at top to 11 at bottom. RO-14-1. Beedeina led.

f!3837 __Coshocton County. Middle Pennsylvanian.

Same locality as f!3836. C.L. Rice colln. 

RO-14-2. Beedeina leei.

f!3838 __Coshocton County. Middle Pennsylvanian.

Same locality as f!3836. C.L. Rice colln. 

RO-14-3. Beedeina leei.

f!3839 __Coshocton County. Middle Pennsylvanian.

Same locality as f!3836. C.L. Rice colln. 

RO-14-4. Beedeina leei.

f!3840 __Coshocton County. Middle Pennsylvanian.

Same locality as f!3836. C.L. Rice colln. 

RO-14-5. Beedeina leei.

f!3841 __Coshocton County. Middle Pennsylvanian.

Same locality as f!3836. C.L. Rice colln. 

RO-14-6. Beedeina leei.
REGISTER OF LOCALITIES AND COLLECTIONS 25

REGISTER OF LOCALITIES AND COLLECTIONSContinued

Locality Collection

OHIO-Continued

f!3842 __Coshocton County. Middle Pennsylvanian.

Same locality as f!3836. C.L. Rice colln.

RO-14-7. Beedeina leei. 

f!3843 __Coshocton County. Middle Pennsylvanian.

Same locality as f!3836. C.L. Rice colln.

RO-14-8. Beedeina leii. 

f!3844 __Coshocton County. Middle Pennsylvaniaa

Same locality as f!3836. C.L. Rice colln.

RO-14-9. Beedeina leei. 

f!3845 __Coshocton County. Middle Pennsylvanian.

Same locality as £13836. C.L. Rice colln.

RO-14-10. Beedeina leei. 

f!3846 __Coshocton County. Middle Pennsylvanian.

Same locality as f!3836. C.L. Rice colln.

RO-14-11. Beedeina leei. 

f9716 ___Coshocton County. Middle Pennsylvanian.

Putnam Hill Limestone Member. Pike

Township, east center sec. 20. This is

measured section No. 1656 in files of Ohio

Geol. Survey. Pauline Smyth, collector,

9-8-53. Beedeina leei.

80 ___f!4149 __Guernsey County. Upper Pennsylvanian.

Bloomfield 7V2-min Quadrangle. Cambridge 

Limestone Member. Small abandoned quarry 

approx. 1 mi northeast of New Concord in 

W%SEV4 sec. 25, Adams Township. R.C. 

Douglass, collector, 5-13-79. Triticites 

ohioensis.

81 ___f!3827 __Tuscarawas County. Middle Pennsylvaniaa

Center of partial sec. 3, T. 9 N., R. 4 W., 

Sugarcreek Quadrangle. 9,700 ft west and 

15,000 ft south of northeast corner of 

quadrangle. Smyth (1957) locality 60 in 

Boggs Limestone. C.L. Rice colln. KG-76-8. 

Fusulinella imprimal

82 f9715 ___Tuscarawas County. Middle Pennsylvanian.

Upper Mercer Limestone from Lawrence 

Township, lk mi southeast of Zoar in railroad 

cut. Pauline Smyth, collector, 9-8-53. 

Fusulinella iowensis. 

f!4003 ___Tuscarawas County. Middle Pennsylvanian.

1 mi southwest of Mineral City on north side 

of Ohio Route 800 at locality 9 of Smyth 

(1957). Road and railroad may have been 

relocated. C.L. Rice colln. MC-1, about 10 ft 

above railroad bed and overlying coal bed. 

Beedeina sp.

83 ___f9714 ___Stark County. Middle Pennsylvanian.

Lower Mercer Limestone Member from 

Bethlehem Township. SWV4WV4 sec. 16. 

Pauline Smyth, collector, 9-1-53. 

Fusulinella iowensis. 

f!3108 __Stark County. Middle Pennsylvanian.

Putnam Hill Limestone Member of 

Allegheny Formation. Pike Township, Dover 

15-min Quadrangle. SWV4NEViNEV4 sec. 27.

REGISTER OF LOCALITIES AND COLLECTIONS -

Continued

Locality Collection

______________OHIO-Continued______________

L.G. Henbest, collector, 10-30-61. Beedeina 

sp. 

f!3109 __Stark County. Middle Pennsylvanian.

Vanport Limestone Member of Allegheny 

Formation. Dover 15-min Quadrangle. 

SWV4NEV4NEV4 sec. 27. L.G. Henbest, collector, 10-30-61. Wedekindellina euthysepta, Beedeina henbesti.

84 ___f!3887 __Columbiana County. Upper Pennsylvanian.

NWV4NEV4 sec. 21, T. 13 N., R. 2 W., West 

Point 7-min. Quadrangle. Ames Limestone in 

roadbank on west side of Ohio Route 45 at 

crest of hill west of lookout tower. Locality 

108 of Smyth (1957). R.C. Douglass, R. Ingavat, and C.H. Cheong colln. 5-26-4. 

Triticites sp. aff. T. ricei.

85 ___f!3886 __Mahoning County. Middle Pennsylvanian.

Mouth of Turkey Broth Creek where it 

enters Berlin Lake at approx. locality 79 of 

Smyth (1957). Mercer Limestone Member. 

R.C. Douglass, R. Ingavat, and C.H. Cheong 

colln. 5-26-3. Fusulinella iowensis.

______________PENNSYLVANIA______________

86 ___f!3884 __Mercer County. Middle Pennsylvanian.

Abandoned quarry on small road between 

Irishtown and Filer Corners, 5 mi east of 

Mercer, about 2 mi north of Irishtown. Vanport Limestone Member float. R.C. 

Douglass, R. Ingavat, and C.H. Cheong colln. 

5-26-1. Beedeina sp. 

f 13885 __Mercer County. Middle Pennsylvanian.

Same general locality as f!3884, but only 1 

mi north of Irishtown in abandoned quarry. 

Vanport Limestone Member. R.C. Douglass, 

R. Ingavat, and C.H. Cheong colln. 5-26-2. 

Beedeina sp.

87 ___f9111 ___Armstrong County. Middle Pennsylvanian.

1 mi by road northeast of Mahoning. F. 

Swain, collector. Beedeina sp. 

f!3881 __Armstrong County. Middle Pennsylvanian.

About 1 mi by road (V2 mi airline) northeast 

of Mahoning. 6 ft of limestone outcrop on 

west side of road. Mahoning Limestone of 

Rogers (1858). R.C. Douglass, R. Ingavat, 

and C.H. Cheong colln. 5-24-1. Beedeina 

leei, B. sp.

88 ___f!3882 __Allegheny County. Upper Pennsylvanian.

Brilliant Cut on south bank of Allegheny 

River in Pittsburgh, just east of railroad 

bridge from Sharpsburg and south of 

highway. Ames Limestone. R. C. Douglass, 

R. Ingavat, and C.H. Cheong, with H.B. 

Rollins, colln. 5-25-1. Triticites cullomensis.
26 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS

SYSTEMATIC DESCRIPTIONS

Profusulinella kentuckyensis Thompson and Riggs, 1959

Plate 1, figures 1-31

Profusulinella kentuckyensis Thompson and Riggs, in Thompson, 

Shaver, and Riggs, 1959, p. 776, 777, pi. 104, figs. 1-15.

Diagnosis. - Shell elongate fusiform, as much as 3.5 

mm long and 1 mm wide at six volutions; nearly straight 

axis of coiling after first volution. Spirotheca thin, composed of protheca with irregularly developed outer tectorium, inner tectorium developed locally in parts of 

outer volutions.

Description. - Shell small, elongate fusiform attaining 

3.5 mm in length and 1 mm in diameter; six volutions in 

the largest specimens. Initial chambers commonly forming a rounded to elliptical shell, axis of coiling at an 

angle of several to as much as 90° to the axis of the 

outer volutions. The axis is nearly straight in many 

specimens but is curved in others. Measurements for the 

newly illustrated specimens are given in table 1 and 

figure 6. The proloculus varies in outer diameter from 

near 0.05 to 0.11 mm in the specimens studied. It is quite 

round except for a slight flattening in the area of the 

apertural pore. The prolocular wall is thin and has an indistinct structure.

The spirotheca is composed of a tectum and 

diaphanotheca, with only a hint of fibrous structure in 

the outer volutions of some specimens. An outer tectorium is deposited irregularly. The deposits appear to 

be thicker along the lateral slopes, as stated by Thompson and Riggs (in Thompson, Shaver, and Riggs, 1959), 

but they are only apparent along the base of the septa 

and are, therefore, neither tectoria nor chomata. An inner tectorium is present only in small areas and in some 

specimens. In the holotype (figs. 1-3), some inner tectorium can be seen in parts of two volutions, and, in an 

equatorial section of a paratype (figs. 10-12), it can be 

seen in places in the two outer volutions. The minimal 

development of inner tectorium on this species suggests 

the first stage of advancement toward Fusulinella, but 

the species is more appropriately assigned to Profusulinella.

The septa are plane to irregularly fluted even across 

the central region in some specimens. The fourth and 

fifth volutions contain 19-20 septa per volution.

The tunnel extends one-third to one-half the volution 

height, is narrow in the inner volutions, and widens irregularly in the outer whorls. It tends to wander in most 

specimens. The chomata are low and asymmetrical, with 

a gentle slope toward the poles.

Discussion. - The description is based on the Thompson and Riggs specimens and on specimens collected 

from the type locality and other surface and well-core 

samples from western Kentucky. The specimens illustrated are from the core samples in addition to the 

type material. In outcrop and in the cores, this species is 

found at more than one horizon, but the differences between specimens from the several horizons are minor 

and not systematic. The specimens from the cores are 

shown in approximate stratigraphic order; those 

depicted in figures 28-31 occur below those depicted in 

figures 23-26, which, in turn, are below those shown in 

figures 21-22. The specimen depicted in figure 27 occurs 

below those shown in figures 19-20. Specimens shown 

in figures 13-18 are from the same general level as those 

shown in figures 19-20, which are from a different well. 

The outcrop samples are from limestone beds less than 

10 ft apart. The core samples include one or more 

horizons through an interval of about 3 ft, followed by 

another set of horizons about 3 ft thick and 30-45 ft 

deeper in the wells. The surface samples correlate with 

the upper horizons in the wells; the lower horizons have 

not been recognized in the surface samples. The smallest 

specimens are dominant in the lower horizon, but some 

larger specimens were found even at the lowest horizon. 

The species is probably a late representative of the 

genus, as suggested by its elongate shape and the incipient development of the inner tectorium.

Distribution. - Surface samples containing this 

species include f 13632 at locality 14 in Hancock County 

and f 13630, f 13631, f!3900, f!3901, and f!3994 at locality 36 in Butler County, Ky. The core samples are from 

wells at locality 11 in Hancock County, 13 in Daviess 

County, 21 in Webster County, 31 in Christian County, 

33 in Muhlenberg County, 34 and 37 in Butler County, 

and 38 in Ohio County, Ky. The samples containing Profusulinella kentuckyensis also have abundant specimens 

of Millerella. Thompson and Riggs (in Thompson, 

Shaver, and Riggs, 1959, p. 776) noted the wide variety 

of forms present, from partly evolute to involute. 

Because of the apparent complete gradation between 

the end members of Millerella, I do not believe that they 

should be separated into species, let alone genera, even 

though Thompson and Riggs (in Thompson, Shaver, and 

Riggs, 1959) assigned the more involute forms to 

Paramillerella sp.

Types. - Some of the originally illustrated specimens 

are here reillustrated. The additional specimens are 

hypotypes.

Profusulinella ohioensis Douglass n. sp.

Plate 2, figures 1-12

Diagnosis. - Shell minute, subrounded, attaining 

lengths near 0.7 mm, widths near 0.5 mm in four to five 

volutions. Coiling of first volution commonly at angle to 

later volutions. Spirotheca thin and composed of tectum, 

diaphanotheca, and outer tectorium. Septa nearly 

straight throughout.
SYSTEMATIC DESCRIPTIONS 27

TABLE 1.-Measurements (in millimeters) for Profusulinella kentuckyensis Thompson andRiggs, 1959

[Leaders (__), data not available or character absent]

Character Measurements of fossils shown in plate 1, figures 13, 16, 18-22

18 19 20 21 22

Diameter of proloculus. 

Radius vector

0.10

1

2

3

4

5

Half length 

1

2

3

4

5

Volution height 

1

2

3

4

5

Wall thickness 

1

2

3

4

5

0.11

.17

.28

.37

0.12

.25

.49

.76

0.03

.06

.10

.10

0.02

0.11

0.13 

.20 

.29 

.40

0.21 

.42 

.67 

.90

0.05 

.06 

.10 

.10

0.05 

.04

0.08

0.09 

.15 

.23

0.14 

.30

.41

0.03 

.05

.05

0.09 

.14 

.20 

.30 

.41

0.10

.24

.44

.77

1.08

0.03 

.05 

.06 

.09 

.13

.04

.05

0.10

0.10 

.15 

.23 

.34

0.14 

.24

.44

0.025

.05

.07

.11

Tunnel width (axials) or 

septal spacing

Diameter of proloculus __ 0.09

Radius vector 

1

2

3

4

5

Half length 

1

2

3

4

5

Volution height 

1

2

3

4

5

Wall thickness 

1

2

3

4

5

0.10

.16

.23

.33

.46

0.14

.32

.58

1.14

1.52

0.03

.05

.07

.10

.13

.03

0.09

0.12 

.18 

.26

0.05 

.06

Tunnel width (axials) or 

septal spacing

1 ___________.

2 ___________.

3 _________________

4 ___________.

0.04 

.09 

.14

0.02

.035

0.07

0.07 

.12

.17 

.24

0.09 

.21 

.38 

.58

0.02 

.04 

.05 

.07

.04

0.02 

.04 

.09

0.05

0.07 

.14 

.21 

.28

0.10 

.25 

.44 

.73

0.03 

.06 

.08 

.08

0.02

0.02 

.05 

.07 

.08

0.09

0.10 

.15 

.23 

.34 

.46

0.16 

.33

.54

.94

1.22

0.03 

.05 

.08 

.11

.12

0.02

0.02 

.06 

.11 

.18

0.06

0.06 

.10 

.24 

.34

0.04 

.06 

.07 

.09

.01 

.02

0.02 

.03 

.04 

.05

0.08

0.11 

.17 

.22 

.32

0.03 

.04 

.07 

.09

0.02 

.02 

.02

0.07

0.06 

.10 

.15 

.23

0.09 

.21 

.42 

.56

0.02 

.04 

.05 

.07

.02 

.05

0.07

0.07 

.12 

.17 

.27

0.08 

.17 

.36 

.54

0.02 

.03 

.05 

.09

.025

1

2

3

4

Character

0.03

.04

.06

.10

.07 

.11

.17

0.02 

.04 

.08

0.03 ____ 

.06

.11

.15

0.025 

.03 

.04 

.06

0.03 

.04 

.05

Measurements of fossils shown in plate 1, figures 23-30

23 24 25 26 27 28 29 30

0.11

0.11 

.15 

.23

0.11 

.21

.44

0.02 

.04 

.07

0.02 

.03 

.06

Description. -Measurements are presented in table 2 

and on figure 7. Coiling of the first volution is commonly 

at a large angle to the outer volutions. The shape is quite 

rounded, expanding only a little faster in length than in

width; maximum form ratios are near 1.3 in the outer 

volutions. Only four to five volutions are present. The 

observed prolocular diameters range from 0.06 to 0.12 

mm, with a mean of 0.09 mm in six specimens.
28 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS

0.3

|f 

Z D 0.2

2* 

0.1

CO

_iuj 0.10

Xh1-

1 0.3

I

Z 0.2 

O

D 0.1 

0

3.5

3.0

I

§ 2.0

LJ_

< 1.5 

1.0 

0.5

Figure 6.- 

Riggs, 19

each attn

The sp 

protheca 

The septs 

fluting a1 

tunnel is 

regularly 

only well 

Compaf 

one of th< 

copiosa 1 

about hal 

is about t 

has a smj 

Distrib 

f!4509 at

 ***""* *

 S* 

: ....

t .* >r %V 

-

;

-

0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 0.9 1.0 

RADIUS VECTOR

Graphs for Projusulinella kentuckyensis Thompson and 

59. All measurements are in millimeters, and the values for 

bute are plotted against the radius vector.

irotheca is thin and profusulinellid, having 

and an outer tectorium but no inner tectorium. 

i are nearly plane but show slight and irregular 

; the poles. They are irregularly spaced. The 

fairly straight in the outer whorls and widens 

. It is bordered by low broad chomata that are 

developed in the vicinity of the septa. 

rison and remarks. -Profusulinella ohioensis is 

. smallest species of the genus. It resembles P 

'hompson, 1948, in general shape but is only 

f its size at comparable volutions. P. ohioensis 

;he same size as P. fittsi Thompson, 1935, but 

aller form ratio and less-developed chomata. 

ution. -P. ohioensis is known only from sample 

localitv 67 from Hockinsr Countv. Ohio, where

it occurs in the Boggs Limestone Member of the Pottsville Formation. It is associated with the intermediate 

form Fusulinella imprima n. sp. 

Specimens studied. -A large number of isolated 

specimens were studied, and seven were prepared as 

oriented thin sections for detailed study. 

Designation of types. -The specimen illustrated on 

plate 2 as figures 9 and 10 is designated the holotype. 

The other illustrated specimens are paratypes.

Fusulinella imprima Douglass n. sp.

Plate 2, figures 13-24

Name derivation. - From the Latin imprimis, among the first. 

Diagnosis. - Shell elongate fusiform attaining lengths 

near 4 mm and widths more than 1 mm in six volutions. 

Coiling regular, showing rapid increase in length with 

each volution. Spirotheca is profusulinellid through most 

of shell but develop inner tectorium in outermost volutions. Septa are irregularly fluted. 

Description. - Measurements are presented in table 3 

and on figure 8. Coiling is regular from the beginning, 

the shell expanding regularly in diameter and rapidly in 

axial length. The form ratio increases from near 2 in the 

inner volutions to more than 3 in the outer volutions.

TABLE 2. -Measurements (in millimeters) for Profusulinella ohioensis 

Douglass n. sp.

[Leaders ( __ ), data not available or character absent]

Character Measurements of fossils shown in plate 2, figures 1, 7, 10, 3-5

1 7 10 3 4 5

Diameter of 

proloculus ____ 0.08 0.07 0.12 0.08 0.10 0.06 

Radius vector 

1 0.09 0.06 0.08 0.12 0.13 0.07

2 .15 .15 .12 .18 .19 .10

3 .20 .20 .18 .26 .30 .16

4 .27 .25 .27 .24

5

6

Half length 

1 0.10 0.10 0.14

2 .16 .14 .18

3 .22 .21 .25

4 .32 .28 .36

5 .35

6

Volution height 

1 0.03 0.015 0.02 0.04 0.04 0.03

2 .06 .035 .03 .07 .07 .03

3 .05 .05 .06 .08 .10 .06

4 .07 .06 .08 .08

5

Wall thickness 

1 0.018 _ 0.015

2 0.02 .02 .02

3 .035 .02 .03 .02 .03

4 .04 .02

5

Tunnel width (axials) 

or septal spacing 

1 0.03 0.02 0.04 0.02 0.03 0.01

2 .05 .04 .05 .025 .05 .03

3 .07 .04 .07 .035 .05 .03

4 .06 .10 _ .04
SYSTEMATIC DESCRIPTIONS 29

0.4

0.3 

9 0.2

0.1 

| 0.10

y o.o5

y 0.3

LJJ

X

z 0.2

0.1

3.5 

3.0

2.0

< 1.5

1.0 

0.5

*** .-» 

**

0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 0.9 1.0 

RADIUS VECTOR

Figure 7.-Graphs for Profusulinella ohioensis Douglass n. sp. All 

measurements are in millimeters, and the values for each attribute 

are plotted against the radius vector.

The axis of coiling is nearly straight to irregularly 

curved. The maximum outer diameter of the proloculus 

ranges from 0.08 to 0.14 mm, averaging 0.11 in nine 

specimens.

The spirotheca is thin and is intermediate between 

profusulinellid and fusulinellid. Throughout most of the 

shell, the two layers of the protheca have only an outer 

tectorium, but in the outermost volutions, an inner tectorium is deposited. The inner tectorium is not continuous in some of the specimens, suggesting an early 

stage of development of the fusulinellid wall. The septa 

are slightly and irregularly fluted across the middle of 

the shell and irregularly fluted at the poles. Septal pores 

are common in some specimens. The septa are irregularly spaced and number 24 to 28 per volution in the outer

whorls. The tunnel extends less than half the volution 

height. It widens gradually in the inner volutions and 

more rapidly in the outer whorls. Its path tends to be irregular in many of the specimens. The chomata are 

asymmetrical and tend to be high and broad. At and 

near the septa, they overhang the tunnel.

Comparison and remarks -Fusulinella imprima is 

one of the species intermediate between Profusulinella 

and Fusulinella. It is similar to F. primotina but differs 

in being larger, more elongate, and in having more 

chambers per volution. It is most similar to F. pria n. sp. 

but differs in being smaller, and in having fewer volutions, fewer chambers per volution, and a smaller proloculus. The assignment of this form to Fusulinella 

rather than Profusulinella is arbitrary and is based on 

this form's greater similarity to the several transitional 

forms assigned here, and previously assigned to 

Fusulinella. The wall structure is profusulinellid except 

in the outermost volutions, so immature specimens do 

not develop the fusulinellid stage.

Distribution-F. imprima is known from one sample 

in the Boggs Limestone Member of the Pottsville Formation at locality 67 in Hocking County, Ohio. Other 

samples of the Boggs Limestone Member yield 

specimens suggestive of this form, but all I saw were so 

poorly preserved that they did not justify description or 

illustration.

Specimens studied.-Twenty oriented sections were 

prepared from isolated specimens, many of which were 

abraded. Other sections of possibly conspecific forms 

from other samples of the Boggs Limestone Member 

were studied but not used in the description.

Designation of types.-The specimen illustrated on 

plate 2 as figures 19 and 20 is designated the holotype. 

The other illustrated specimens are paratypes.

Fusulinella primotina Douglass n. sp.

Plate 3, figures 1-23 

Name derivation-From the Latin primotinus, early or first.

Diagnosis. - Shell irregularly fusiform, attaining 

lengths near 2.5 mm and widths near 1 mm in about six 

volutions. Coiling regular after the first volution, which 

may be at a large angle to subsequent volutions. Wall is 

profusulinellid through most of shell, but develops an inner tectorium in outermost volutions. Septa are straight 

across middle of shell and irregularly fluted at poles.

Description. -Measurements are presented in table 4 

and on figure 9. The coiling, after the first volution, is 

regular, each volution tending to parallel the previous 

one, with little shape change throughout the ontogeny. 

The ratio of length to diameter (fig. 9) increases gradually from 1.3 in the inner volutions to approximately 2.8 at 

maturity. The axis of coiling is commonly irregular, so
30 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS

TABLE 3. -Measurements (in millimeters) for Fusulinella imprima Douglass n. sp.

[Leaders (__), data not available or character absent]

Character

Diameter of proloculus __ 

Radius vector 

1

2

3

4

5

6

Half length 

1

2

3

4

5

6

Volution height 

1

2

3

4

5

6

Wall thickness

1

2

3

4

5

6

Tunnel width (axials) or 

septal spacing 

1

2

3

4

5

6

Measurements of fossils shown in plate 2, figures 15-17, 13, 18, 19, 21-23

15

0.13

0.11 

.18 

.24 

.33 

.45

0.22

.42 

.55 

1.25

0.04 

.06 

.06 

.09 

.12

0.04 

.06 

.10

16

0.13

0.09 

.13 

.22 

.29 

.41 

.57

0.21 

.37 

.48 

.96 

1.38

0.03 

.08 

.08 

.11 

.16

 

0.04 

.06 

.09

____

17

0.13

0.10 

.14 

.18 

.25 

.32 

.44

0.02 

.03 

.035 

.06 

.08 

.11

0.01 

.015 

.02 

.02

0.02 

.025 

.03 

.04 

.04

13

0.10

0.11 

.16

.24 

.32 

.45

0.02 

.04 

.07 

.09 

.11

.01 

.02 

.03 

.04

0.02 

.03 

.05 

.06 

.09

18

0.09

0.08 

.14 

.22 

.31 

.42

0.17 

.34 

.65 

1.00 

1.66

0.02 

.06 

.07 

.09 

.11

.04

0.03 

.04 

.06 

.07 

.15

19

0.08

0.10 

.14 

.20 

.27 

.40 

.56

0.18 

.32 

.52 

1.02 

1.51 

2.11

0.03 

.04 

.05 

.07 

.13 

.16

____

0.03

.07 

.08 

.11

21

0.11

0.12 

.18 

.26 

.39 

.52

0.27 

.51 

1.03 

1.68 

2.09

0.03 

.06 

.08 

.12 

.14

.02 

.03

0.06 

.08 

.12 

.18

____

22

0.11

0.13 

.18 

.26 

.36 

.51 

.69

0.16 

.31 

.71 

1.15 

1.59 

2.08

0.04 

.05 

.07 

.10 

.15 

.18

0.03 

.04 

.06 

.09 

.20 

.21

23

0.14

0.12 

.18 

.29 

.41

.57

0.24 

.46 

.86 

1.14 

1.80 

2.35

0.02 

.06 

.10 

.12 

.16

~~02

0.04 

.08 

.11 

.18

many axial sections deviate from the axis toward one of 

the poles. The proloculus is variable in outer diameter 

from about 0.06 to 0.12 mm but is commonly close to 

0.10 mm and tends to be round.

The spirotheca is of special interest in this form 

because it shows attributes intermediate between Profusulinella and Fusulinella. The inner volutions have a 

spirotheca composed of the protheca (tectum and 

diaphanotheca) and an outer tectorium. This structure is 

typical of Profusulinella. The outer volutions show some 

development of an inner tectorium. More inner tectorium is present in some specimens than in others, but 

the overall sample available shows that an inner tectorium is common in the last two volutions. The septa 

are nearly straight across the middle of the shell and are 

irregularly fluted at the poles. The septa are evenly 

spaced, and there are commonly 19-20 septa per volution from the fourth volution out. The tunnel is narrow 

and tends to be straight, but it is irregular in many of 

the specimens. It is generally less than half the chamber 

height. The chomata are generally low and broad but 

often appear higher or even overhanging where the section passes near a septum.

Comparison and remarks.-F. primotina is a form

whose attributes are intermediate between those of Profusulinella and Fusulinella. Similar previously described species are F. fugax Thompson (1948), the 

specimens of F. acuminata Thompson (1936) that 

Thompson (1948) illustrated from Powwow Canyon, 

Tex., and F. searighti Thompson, 1953, from Missouri. 

F. primotina is also of the same general nature as F. imprima and F. pria, described herein from eastern Kentucky and from Ohio. F. primotina differs from F. 

fugax, which expands more rapidly in volution height 

and less rapidly in axial length and has a thicker 

spirotheca. Comparisons with F. searighti Thompson, 

1953, are difficult because the seven specimens of that 

species illustrated show such variation. Several of the 

specimens have a straight axis of coiling and a regular 

fusiform shape, in contrast to the irregularities of F. 

primotina. The two adult axials illustrated by Thompson 

are both larger than any specimens of F. primotina having the same number of volutions. The specimens illustrated as F. acuminata by Thompson (1948, pi. 34) 

from Powwow Canyon, Tex., and from the Mud Springs 

Mountains, N. Mex., are similar in several attributes to 

F. primotina, but they are larger, have a larger proloculus, and have a straighter axis of coiling.
SYSTEMATIC DESCRIPTIONS 31

0.4

0.3 -

in

1 9 0.2

0.1 -

uj 0.10 

0.05

§ 0-3

Z 0.2 

O

3 °' 1 

d

3.5 

3.0 

2.5 

2.0

< 1.5

1.0 

0.5

V

0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 0.9 1.0 

RADIUS VECTOR

Figure 8.-Graphs for Fusulinella imprima Douglass n. sp. All 

measurements are in millimeters, and the values for each attribute 

are plotted against the radius vector.

Three species of this transitional group are described 

in this paper. F. primotina is the smallest and the most 

irregular in form and in axis and has the fewest 

chambers per volution.

Distribution. -F. primotina is known only from sample f!3927 at locality 21 in Webster County, Ky. It was 

found in the core of Gil-31 at a depth of 471.5-471.7 ft, 

just 2 feet below the occurrence of F. iowensis in the 

core. No specimens of this species were found in the 

base of the Curlew Limestone Member at its type locality or in other beds below the F. iowenis horizon in the 

Eastern Interior Basin.

Specimens studied. -Thirty thin sections were prepared, each containing at least one oriented section of a 

specimen and one or more random sections of other

specimens of this species. The types of the closely 

related species were studied for comparison.

Designation of types.-The specimen illustrated on 

plate 3 as figures 20 and 21 is designated the holotype. 

The other illustrated specimens are paratypes.

Fusulinella pria Douglass n. sp.

Plate 4, figures 1-18 

Name derivation-From the Latin prius, earlier.

Diagnosis. - Shell elongate, fusiform, attaining 

lengths near 5.5 mm, widths near 1.5 mm in six volutions. Coiling is regular after first volution, with rapid 

increases in length but with irregular axis of coiling. 

Wall is composed of protheca and minimum epithecal 

deposits; inner tectorium restricted to parts of outer 

volutions. Septa are irregularly fluted throughout.

Description. -Measurements are presented in table 5 

and on figure 10. Coiling of the first volution may be at 

an obtuse angle to the outer whorls. The shell expands 

regularly in diameter and rapidly in length, and form 

ratios expanding from near 1.5 in the inner volutions to 

more than 3 in the outer volutions (see fig. 10). The axis 

of coiling is irregularly curved, and the lateral slopes 

tend to be irregular. The proloculus ranges in maximum 

outer diameter from 0.09 mm to 0.22 mm with a mean of 

0.15 mm in 16 specimens. The older specimens (lower 

stratigraphically) have the larger proloculi. The 

spirotheca is thin and of special interest because, in 

many parts of most specimens, it is composed solely of 

the protheca (tectum and diaphanotheca). A thin outer 

tectorium is present along parts of the wall, and an inner tectorium is deposited in the outer volutions. The 

daphanotheca has what Dunbar (in Thomas, 1931, p. 32) 

called fibrous structure, a structure resembling ultrafine 

keriotheca (fig. 11). The septa are irregularly fluted 

throughout the length of the shell and are irregularly 

spaced, about 32 per volution in the outer volutions. The 

tunnel is narrow and either straight or wandering in the 

equatorial area. It is bounded by low broad chomata that 

appear higher to overhanging in the vicinity of the septa.

Comparison and remarks. -Fusulinella pria is one of 

the forms that have attributes intermediate between 

those of Profusulinella and Fusulinella. The wall structure is more like that of Profusulinella than that of 

Fusulinella through most of its growth, but an inner 

tectorium is deposited in the outer volutions. F. pria is 

the largest of the three species of this group described in 

this paper. It has more volutions and larger dimensions, 

more chambers per volution, and more fluting across the 

middle of the shell. Some smaller specimens are similar 

in size to the largest specimens of F. imprima but still 

differ in septal count, septal fluting, and details of the 

wall.
32

TABLE 4. -Measurements (in millimeters) for Fusulinella primotina Douglass n. sp.

[Leaders (__), data not available or character absent]

Character Measurements of fossils shown in plate 3, figures 1, 2, 4, 6-8, 10

1

Diameter of proloculus ____ 0.09 

Radius vector

1 _______________ 0.08

2 _______________ .10

3 _______________ .15

4 _______________ .21

5 _______________ .28 

Half length

1 _______________ 0.09

2 _______________ .15

3 _______________ .27

4 _______________ .42

5 _______________ .78 

Volution height

1 _______________ 0.02

2 _______________ .05

3 _______________ .06

4 _______________ .09

5 _______________ .09 

Wall thickness

1 _______________ 0.015

2 _______________ .02

3 _______________ .02

4 _______________ .025

5 ______________________

Tunnel width (axials) or 

septal spacing

1 _______________ 0.02

2 _______________ .04

3 _______________ .05

4 _______________ .08

5 ______________________

Character ________

___________________________12

Diameter of proloculus ____ 0.12 

Radius vector

1 _______________ 0.10

2 _______________ .15

3 _______________ .21

4 _______________ .31

5 ______________________

Half length

1 ______________________

2 ______________________

3 ______________________

4 ______________________

5 ______________________

Volution height

1 __,._____________ 0.03

2 _______________ .05

3 _______________ .065

4 _______________ .09

5 ______________________

6 ______________________

Wall thickness

1 _______________ 0.005

2 _______________ .01

3 _______________ .02

4 _______________ .03

5 ______________________

Tunnel width (axials) or 

septal spacing

1 _______________ 0.02

2 _______________ .03

3 _______________ .04

4 _______________ .04

5 ______________________

0.10

0.10 

.15 

.20 

.28 

.39

0.11

0.09 

.15 

.22 

.31 

.36

0.04

.045

.05

.08

.10

.01 

.01 

.02 

.03

0.02 

.03 

.03 

.04 

.04

0.03

.06

.07

.075

.09

0.01

.01

.015

.02

0.02 

.03 

.04 

.04 

.05

0.10

0.08 

.14 

.22 

.29 

.38

0.15 

.31 

.53

.77 

1.20

0.02 

.06 

.06 

.08 

.10

.03 

.03 

.05

0.03 

.05 

.08

.12

0.11

0.08 

.12 

.16 

.28 

.40

0.085

.23

.44

.69

.93

0.04

.045

.08

.10

.12

.02 

.02

0.03 

.05 

.06 

.10 

.14

0.08

0.07 

.11 

.15 

.23

0.10 

.20 

.39

.72

0.02 

.04 

.04 

.09

0.02 

.02

0.03 

.04

0.10

0.085

.12

.17

.25

.34

0.03 

.03 

.05 

.08 

.09

.02 

.02 

.025

0.02 

.03 

.03 

.04 

.05

Measurements of fossils shown in plate 3, figures 12, 11, 14, 16, 19, 20, 22

14 16 19 20

0.06

0.08 

.15 

.26 

.39 

.51

0.11

.24

.49

.75

1.09

0.03 

.05 

.06 

.07 

.11 

.13

.025

.03

.045

0.025

.04

.06

.10

.12

0.07

0.08 

.13 

.19 

.26

0.13 

.23 

.41 

.59

0.02 

.05 

.06 

.07

.03 

.04

0.02 

.03 

.05

0.07

0.08 

.09 

.13 

.20 

.26

0.14 

.29 

.49 

.74 

.87

0.03 

.05 

.06 

.07 

.11

.01 

~02

0.03 

.03

.13

0.09

0.09 

.15 

.22 

.30

0.11 

.24

0.02 

.05 

.07 

.08

.03

0.03 

.05 

.07

0.085

0.11 

.16 

.23 

.32

0.17 

.35 

.56 

.76

0.03 

.05

.09

0.03 

.03 

.05

.06 

.12

0.11

0.13 

.21 

.29 

.38

.47

0.29

.48

.74

1.01

1.35

0.04 

.07 

.09 

.09 

.09

0.02 

.03

.03

0.03 

.06 

.12 

.15

Distribution. -F. pria is common in the Stoney Fork 

Member of the Breathitt Formation in eastern Kentucky 

from Bell County in the south to Morgan County and is

present in collections from localities 45 through 53.

Specimens studied.-The description is based on 

specimens from the samples at locality 46 in Bell Coun-
SYSTEMATIC DESCRIPTIONS 33

0.4

0.3 

Q 0.2

0.1

ui 0.10

0.05

i

*2 0.3

LU

X

Z 0.2

O

H

D 0.1

_i 

O

3.5 

3.0

I 2 ' 5 

^ 2.0

< 1-5

1.0 

0.5

-.*

 *»« 

0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 0.9 1.0 

RADIUS VECTOR

Figure 9.-Graphs for Fusidinella primotina Douglass n. sp. All 

measurements are in millimeters, and the values for each attribute 

are plotted against the radius vector.

ty, supplemented by information from 32 other samples 

found in six counties. 115 thin sections were prepared, 

and other material was studied in etched slices.

Designation of types.-The specimen illustrated as 

figures 14 and 16 is designated the holotype, and the 

other illustrated specimens are paratypes.

Fusulinella stouti Thompson, 1936

Plate 5, figures 1-26 

Fusulinella iowensis var. stouti Thompson, 1936, p. 676-677, pi. 90,

figs. 5-11; Dunbar and Henbest, 1942, p. 95, pi. 3, figs. 7-9;

Smyth, 1957, p. 263, pi. 1, figs. 8-9. 

Fusulinella stouti Thompson and Riggs in Thompson, Shaver, and

Riggs, 1959, p. 780, pi. 105, figs. 12-24.

Diagnosis. - Shell variable from fusiform to elongate

fusiform, attaining a length of as much as 3.7 mm and a 

width of as much as 1.5 mm in eight to nine volutions. 

Proloculus variable in outer diameter. Spirotheca thin, 

composed of protheca and inner and outer tectoria. 

Septa numerous and irregularly fluted.

Description. -Measurements for the specimens here 

illustrated are given in table 6 and on figure 11. Thompson's original description and illustrations indicate a 

variable species with considerable latitude in inflation, 

septal fluting, and development of chomata. The suite of 

specimens from the Curlew Limestone Member of the 

Tradewater Formation in western Kentucky shows a 

similar wide range of variability. The shell is generally 

fusiform to elongate fusiform, with lateral slopes that 

range from convex through straight to slightly concave. 

Lengths to 3.7 mm and widths to 1.5 mm are attained in 

eight to nine volutions. Form ratios range from 1.5 to 

near 2.5 in mature specimens (fig. 11). The early volutions commonly have low ratios. The proloculus ranges 

in outer diameter from 0.06 to 0.17 mm, averaging 0.10 

mm in 26 specimens.

The spirotheca is thin but includes the protheca and 

varying amounts of epitheca. Many specimens have an 

outer tectorium about equal in thickness to the rest of 

the wall, whereas others have a relatively thin outer tectorium. An inner tectorium is developed throughout the 

outer four to five volutions. The septa are irregularly 

fluted and, contrary to the original description, show 

fluting even in the middle of the shell of some specimens. 

The septa are numerous and are closely, but often irregularly, spaced.

The tunnel is relatively narrow and though fairly 

straight in some specimens, tends to wander in the 

equatorial area. It is generally less than half the 

chamber height. The chomata are high and broad and 

are overhanging in the vicinity of the septa.

Comparison and remarks.-F. stouti, despite its 

variability, is a relatively distinctive form. The dense 

look of the interior whorls caused by the nearly straight 

septa and the extended chomata is characteristic of only 

a few species. F. iowensis is similar but is more inflated, 

producing a smaller form ratio. The specimens from the 

Curlew Limestone Member illustrated here and in 

Thompson, Shaver, and Riggs (1959) show a little more 

of the inner density than the Ohio specimens illustrated 

by Thompson (1936).

Distribution. -F. stouti was first described from beds 

referred to the Boggs Limestone and Mercer Limestone 

Members of the Pottsville Formation in Ohio. It was 

subsequently described from the Seville Limestone 

Member of the Spoon Formation in Illinois and from the 

Curlew Limestone Member of the Tradewater Formation in western Kentucky. All the present illustrations 

are from the type locality of the Curlew Limestone
34 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS

TABLE 5. -Measurements (in millimeters) for Fusulinella pria Douglass n. sp.

[Leaders (__), data not available or character absent]

Character

Diameter of proloculus ___ 

Radius vector 

1

2

3

4

5

6

7

Half length 

1

2 _

3

4

5

6

7

Volution height

1

2

3

4

5

6 _ _

7

Wall thickness 

1

2

3

4

5

6

Tunnel width (axials) or 

septal spacing 

1

2_

3

4

5

6

Measurements of fossils shown in plate 4, figures 1-8

1

0.09

0.07 

.10 

.15 

.22 

.32 

.41 

.52

0.07 

.12 

.26 

.48 

.76 

1.20 

1.58

0.02 

.03 

.05 

.07 

.10 

.10 

.12

0.01

.02

.04

0.02 

.03 

.06

.14

2

0.09

0.06 

.11 

.16 

.23 

.30 

.42

 

0.02 

.05 

.045 

.07 

.08 

.11

.02 

.03 

.03

0.02 

.02 

.03 

.05 

.06

3

0.11

0.09 

.16 

.23 

.32

.40 

.55

0.12 

.26 

.50 

.71 

.96 

1.54

0.02 

.06 

.08 

.09 

.10 

.12

.02 

.02

0.01 

.02 

.05 

.09 

.11

4

0.12

0.10 

.15 

.21 

.31 

.39 

.52

0.16 

.30 

.50 

.78 

1.25 

1.48

0.02 

.04 

.06 

.10 

.08 

.14

.02

.03 

.04

0.02 

.03 

.06

.13

5

0.09

0.10 

.15 

.26 

.35

.47

0.10 

.22 

.36 

.64 

1.03 

1.32

0.03 

.04 

.05 

.07 

.09 

.12

0.01

.02

.02 

.05 

.08 

.10 

.19

6

0.13

0.12 

.18 

.26 

.35

 

 

0.02 

.05 

.08 

.10

.02

.02

0.02 

.03 

.03 

.04

7

0.14

0.22 

.30 

.41 

.51 

.63

0.24 

.46 

.83 

1.17 

1.54 

2.07 

2.58

0.04 

.07 

.08 

.10 

.11 

.12

 

.03 

.035

0.04 

.07 

.10 

.18 

.23

8

0.16

0.14 

.20 

.30 

.41 

.52 

.69 

.86

 

0.03 

.05 

.09 

.11 

.12 

.16 

.17

0.01 

.01

.03 

.04 

.05

0.025 

.04 

.05 

.05 

.06 

.06

Member in Union County, Ky. Other specimens were 

recognized in other surface and subsurface samples 

from this horizon in western Kentucky.

Specimens studied.-Specimens originally described 

by Thompson (1936) were studied, in addition to 

specimens from the localities described by Smyth (1957). 

Most of the specimens studied were from locality 17 in 

Union County; supplemental specimens were from locality 20 in Crittenden County and locality 32 in Christian 

County, Ky.

Designation of types.-The specimen illustrated by 

Thompson (1936) as figure 10 on his plate 90 is 

designated the lectotype.

Fusulinella iowensis Thompson, 1934

Plate 6, figures 1-16; plate 15, figures 14-26

Fusulinella iowensis Thompson, 1934, p. 296-297, pi. 20 figs. 28-30; 

Thompson, 1936, p. 675-676, pi. 90, figs. 12-16; Dunbar and 

Henbest, 1942, p. 93-95, pi. 3, figs. 10-25; Thompson, 1948, pi. 26, 

fig. 12, designated the holotype (lectotype); Smyth, 1957, p. 

262-263, pi. 1, figs. 5-7, 10, 11; Thompson and Riggs, in Thompson, Shaver, and Riggs 1959, p. 777-780, pi. 105, figs. 1-11

Diagnosis. - Shell short and obese, as much as 3.5 mm 

in length and 2.5 mm in width in 10 volutions; form ratio 

near 1.4 in mature specimens. Proloculus ranges from 

0.05 to 0.13 mm in outer diameter. Wall consists of protheca and commonly thick epithecal deposits. Tunnel 

straight to irregular with high broad chomata.

Descriptions. - Measurements for the specimens illustrated on plates 6 and 15 are given in table 7 and on 

figure 12, except for specimens 1-7 on plate 6, for which 

data were tabulated by Thompson and Riggs (in Thompson, Shaver, and Riggs, 1959, p. 778). The shell is small 

and highly inflated fusiform with nearly straight to concave or convex lateral slopes and broadly rounded to 

pointed poles. The first whorl may be at a large angle to 

the rest of the specimen, as illustrated by Dunbar and 

Henbest (1942, pi. 3, figs. 15, 25) and by Thompson and 

Riggs (in Thompson, Shaver, and Riggs, 1959), 

reproduced here on plate 6, figure 1. Subsequent growth 

is quite regular, the shell expanding rapidly in both 

length and width, maintaining a small form ratio (fig. 

12). The proloculus ranges from about 0.04 to near 0.13 

mm in outer diameter. The specimens having the largest
SYSTEMATIC DESCRIPTIONS

TABLE 5. -Measurements (in millimeters) for Fusulinella pria Douglass n. sp. - Continued

35

Character

Diameter of proloculus ___ 

Radius vector 

1

2

3

4

5

6

7

Half length 

1

2

3

4

5

6

7

Volution height 

1

2

3

4

5

6

7

Wall thickness 

1

2

3

4

5

6

Tunnel width (axials) or 

septal spacing

1

2

3

4

5

6

Measurements of fossils shown in plate 4, figures 9, 10, 12-15, 17, 18

9

0.16

0.14 

.20 

.30 

.41 

.52 

.69 

.86

0.03 

.05 

.09 

.11 

.12 

.16 

.17

0.01

.04

0.025 

.04 

.05 

.05 

.06 

.06

10

0.14

0.14 

.22 

.32 

.43 

.55

 

 

 

0.03 

.08 

.10 

.11 

.13

0.01 

.01 

.02 

.05 

.03

0.02 

.03 

.05 

.05

12

0.19

0.14 

.21 

.29 

.40 

.54

0.31 

.48 

.81 

1.34 

2.00

0.04 

.07 

.08 

.10 

.14

.03

.04

0.03 

.05 

.09 

.14 

.21

13

0.22

0.20 

.28 

.40 

.54 

.73 

.89

0.30 

.44 

.97 

1.20 

2.38 

2.83

0.05 

.08 

.12 

.14 

.18 

.17

~~03

____

0.04 

.09 

.11

.28

14

0.20

0.16 

.22 

.33 

.46 

.60 

.75

0.23 

.43 

.86 

1.32 

2.00 

2.55

0.05 

.07 

.10 

.12 

.15 

.15

~~03 

.04 

.03 

.04

0.05 

.06 

.08 

.16 

.24

15

0.12

0.15 

.22 

.36 

.48 

61 

.75

 -

0.06 

.07 

.13 

.12 

.14 

.14

0.02

.03 

.04

0.02 

.03 

.05 

.07 

.08

17

0.21

0.20 

.28 

.37 

.50 

.65

0.37 

.58 

1.02 

1.46 

1.87

0.07 

.08 

.09 

.12 

.15

~03 

.04 

.06

0.04 

.06 

.11 

.20 

.25

18

0.15

0.18 

.25 

.34 

.45 

.60

 

 -

 -

0.04 

.06 

.09 

.11 

.15

0.01 

.01 

.02 

.03 

.04

0.02 

.03 

.04 

.05 

.07

proloculi are from Illinois and Kentucky, but the 

specimens from Ohio also have larger proloculi than 

those illustrated from Iowa. The spirotheca is composed 

of protheca and, in many specimens, an unusually thick 

outer tectorium. The inner tectorium is present 

throughout most of the shell. The septa are numerous 

and closely but irregularly spaced. They are irregularly 

fluted throughout the shell in the outer volution and 

toward the poles in the inner volutions. The tunnel is 

narrow, and, in most specimens, it wanders irregularly 

in the equatorial area. It is commonly about half the 

chamber height. The chomata are high and broad, 

decreasing in breadth in the outermost volutions.

Comparisons and remarks.-The inflated form with 

tight coiling and the broad chomata combined with nearly straight septa in the inner volutions make Fusulinella 

iowensis easy to recognize and distinctive among the 

fusulinellas.

Distribution. - The original specimens of this species 

were described from a limestone 20 ft above rocks of 

Mississippian age and about 90 ft below the White 

Breast coal bed in Jefferson County, Iowa. The species

subsequently has been described from the Mercer 

Limestone Member of the Pottsville Formation in Ohio, 

the Seville Limestone Member of the Spoon Formation 

in Illinois, and the Curlew Limestone Member of the 

Tradewater Formation in Kentucky. No representatives 

have been found in eastern Kentucky.

Specimens studied.-The types from Iowa and 

hypotypes from Illinois, Ohio, and Kentucky were 

studied, in addition to specimens from localities 17 in 

Union County, 21 in Webster County, and 31 and 32 in 

Christian County, Ky., and from localities 82 in Tuscarawas County, 74 and 76 in Muskingum County, and 

85 in Mahoning County, Ohio.

Fusulinella sp.

Plate 6, figures 17-22

Description.-Several specimens in collection f 13635 

at locality 32 and in collection f!3741B at locality 31, 

both in Christian County, Ky., are quite distinct from 

the Fusulinella iowensis Thompson, with which they are 

associated. They are much more elongate and have low
36

0.4

0.3 -

0.2 -

0.1 -

<f)

uj 0.10 h

^ 0.05 -

E °' 3

I

0.2 -

0.1

3.5 

3.0 

2.5 

2.0 

1.5 

1.0 

0.5

 v

 * 

»*.

r

0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 0.9 1.0 

RADIUS VECTOR

Figure 10.-Graphs forFusulinellapriaDouglsiSS n. sp. All measurements are in millimeters, and the values for each attribute are 

plotted against the radius vector.

narrow chomata. They bear some resemblance to F. 

acuminata Thompson, 1936, but have a smaller proloculus and are more tightly coiled. Similar forms were 

not found associated with F. iowensis in the other 

samples. Measurements are presented in table 8 and 

figure 13.

Wedekindellina euthysepta (Henbest), 1928

Plate 7, figures 1-42; plate 16, figures 32-35

Fusulinella euthusepta Henbest, 1928, p. 80-81, pi. 8, fig. 6-8, pi. 9,

figs. 1,2.

Wedekindella euthysepta Dunbar and Henbest, 1930, p. 357-364. 

Wedekindia euthysepta Dunbar and Henbest, 1931, p. 458. 

Wedekindellina euthysepta Dunbar and Henbest, 1933, p. 134, key pi.

10, figs. 13-15; Thompson, 1934, p. 282-285, pi. 20, figs. 1, 2, 7, 9,

12, 13, 17, 22, 24-27; Dunbar and Henbest, 1942, p. 98-100, pi. 8, 

figs. 1-23, pi. 9, fig. 1-4; Smyth, 1957, p. 269-270, pi. 2, figs. 15, 

16, 20; St. Jean, 1957, p. 50-53, pi. 5, figs. 18-32.

Wedekindellina dunbari Thompson, 1934, p. 285-287, pi. 20, figs. 3, 6, 

15, 16, 20, 21.

Wedekindellina uniformis Thompson, 1934, p. 289, 290, pi. 20, fig. 5.

Wedekindellina elfina Thompson, 1934, p. 287-289, pi. 20, figs. 8, 10, 

11.

Diagnosis. - Small slender shell with straight to slightly convex or concave lateral slopes and nearly straight 

septa. Proloculus small and coiling tight. Narrow irregular tunnel and broad chomata. Wall of thin protheca, inner tectorium, and variable amounts of outer 

tectorium.

Description. -This form has been described several 

times, commonly with more stringent limitations than 

those given in the following description. Measurements 

for this species are provided in table 9 and on figure 14. 

The samples studied and the previously described 

specimens show a large size and shape variation. Some 

of the differences reflect growth stage, but some 

variability also exists in rate of expansion in both the 

equatorial and axial directions. The shapes tend to be 

elongate fusiform with straight to slightly curved lateral 

slopes and pointed poles. Rare specimens start with an 

endothyrid juvenarium, but most coil regularly and 

maintain a nearly uniform chamber height well out onto 

the lateral slopes.

The proloculus varies widely in maximum outer 

diameter but rarely exceeds 0.10 mm; it had a mean of 

0.07 mm in 40 specimens. Only rare examples of 

microspheric proloculi have been found. One such 

specimen illustrated by Thompson (1934, pi. 20, fig. 5) 

was assigned to a new species, but, because it is 

associated with abundant Wedekindellina euthysepta 

and does not differ in other ways, I do not consider it 

distinct.

The spirotheca is generally quite thin and composed of 

a protheca and varying amounts of tectoria. An inner 

tectorium is common, and an outer tectorium along the 

flanks-especially at the base of the septa-develops an 

axial filling. Dunbar and Henbest (1942, p. 98) discussed 

the porosity in the protheca that is evident in some 

specimens. Many of the specimens show the fine porosity especially well in equatorial section.

The septa are essentially straight in most specimens 

but show some weak fluting-especially in the poles. The 

septa are closely and fairly regularly spaced in the inner 

volutions; more irregularity appears in the outer volutions. The tunnel is narrow and tends to wander in the 

equatorial area. It commonly extends more than half the 

chamber height. The chomata are low and broad, often 

appearing to extend laterally where epitheca are 

deposited along the base of the septa, forming axial filling.
SYSTEMATIC DESCRIPTIONS 37

TABLE 6. -Measurements (in millimeters) for Fusulinella stouti Thompson, 1936

[Leaders (__), data not available or character absent"]

Character 

123456789

Measurements of fossils shown in plate 5, figures 1-9

Diameter of proloculus __ 0.08 0.09 0.12 0.09 

Radius vector 

1 0.07 0.11 0.11 0.09

2 .13 .15 .16 .15

3 .20 .21 .24 .21

4 .30 .29 .34 .31

5 .45 .39 .45 .39

6 .59 .50 .60 .56

7 .75 .64 .74 .69

8 .80

0.14 0.08 0.09 0.10 0.07

0.11 0.08 0.10 0.075 0.07 

.16 .11 .15 .12 .11 

.26 .19 .22 .17 .17 

.36 .25 .34 .26 .26 

.48 .35 .45 .38 .36 

.59 .48 .58 .53 .50 

.75 .62 .72 .71 .66 

.89 .79 .90 .83

Half length 

1 0.10 0.12 0.11 0.16 0.10 0.09

2 .23 .23 .24 .25 .17 .13

3 .41 .34 .39

4 .58 .51 .58

5 .77 .79 .99

6 1.00 1.02 1.39

7 1.34 1.15 1.60

8 1.31

Volution height 

1 0.03 0.04 0.04 0.02

2 .05 .04 .05 .03

3 .07 .06 .08 .04

4 .11 .07 .10 .10

5 .16 .11 .11 .08

6 .15 .11 .15 .17

7 .16 .14 .15 .16

8 16

Wall thickness 

1 0.01

.41 __ __ .36 .27 

.68 .55 .41

.86 __ __ .79 .55 

1.05 1.12 .79

1.34 __ 1.43 1.01 

1.58 __ __ __ 1.27

0.03 0.03 0.03 0.02 0.03 

.05 .03 .05 .05 .04 

.09 .07 .08 .05 .06 

.11 .03 .11 .02 .08 

.12 .10 .11 .125 .11 

.12 .13 .13 .145 .14 

.16 .16 .14 .185 .16 

.18 .16 .18 __ .17

2 .01 .02

3 .01 .02 .01 .01 .02 .012

4 .025 .02 .015 .03 .02

5 .025 .03 .02 .03 .025

6 .045 .035 .05

7

8 .05

Tunnel width (axials) or 

septal spacing 

1 0.02 0.02 0.03

2 .03 .03 .02

3 .03 .02 .04 .02

4 .06 .04 .05 .04

5 .07 .06 .06 .06

6 .08 .05 .14

7 .12 ... ... . .

.03 .02 .02 .02 .015 

.025 .03 .035 .02

.05 .04 .03 .035 .03 

.07 .04 .03 .06 .05 

__ .05 .03 .12 .07 

07 .05 .15 .11

8 .07 .20

Comparison and remarks. - W. euthy septa is quite 

variable in size and shape within any one sample. Some 

specimens have been assigned to other species on the 

basis of minor differences in size, shape, or wall 

thickness. Most of the size differences are based on 

maturity, the smaller specimens having fewer volutions. 

Large and small forms are present in most samples. 

Dunbar and Henbest (1942, p. 99) pointed out the differences between W. euthysepta and W. henbesti (Skinner, 1931): W. henbesti is larger at 10 volutions than W. 

euthysepta at 11 volutions. W. euthysepta is also smaller 

than W. ellipsoides Dunbar and Henbest. W. ellipsoides 

is considered sufficiently different from W. euthysepta 

because it has a different shape and several more volutions than do the large specimens of W. euthysepta 

Distribution. - W. euthysepta was described from the 

Stonefort Limestone Member of the Spoon Formation in 

Illinois. In Kentucky, it was found in outcrop samples of

the Yeargins limestone member (informal usage) of the 

Tradewater Formation and in core samples correlated 

with those beds. In Ohio, it is found in the Vanport 

Limestone Member of the Allegheny Formation. 

Specimens identified as Wedekindellina sp. that might 

be assignable to this species were found in rocks assigned to the Vanport in western Pennsylvania. Thompson (1934) described this species from 35 ft below the 

"White Breast Coal" in Iowa. 

Specimens studied. - The types of this species from Illinois and of conspecific forms from Iowa were studied, 

in addition to surface and core samples from western 

Kentucky at localities 18, 21, 31, 32, 39, 40, and 41; and 

from Ohio at locality 83. 

Designation of types. -Dunbar and Henbest (1942) 

designated as a lectotype the specimen illustrated by 

Henbest (1928, pi. 8, fig. 6), which they reillustrated 

(1942, pi. 8, figs. 15-17).
38 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS 

TABLE 6.-Measurements (in millimeters) for Fusulinella stouti Thompson, 1936-Continued

Character Measurements of fossils shown in plate 5, figures 10-18

10 11 12 13 14 15 16 17 18

Diameter of proloculus __ 0.09 0.06 0.09 0.12 

Radius vector 

1 0.09 0.09 0.10 0.11

2 .15 .13 .14 .16

3 .22 .23 .22 23

4 .30 .31 .34 .33

5 .41 .45 .49 .46

6 .54 .60 .64 .60

7 .69 .75 .95 .75

0.07 0.08 0.08 0.095 0.12

0.07 0.09 0.08 0.09 0.10 

5 .10 .15 .12 .13 .15 

.17 .22 .21 .19 .23 

.23 .34 .28 .26 .31 

.32 .43 .38 .34 .43 

.43 .55 .49 .48 .58 

58 .69 .63 .61

8 .87 .76

Half length 

1 0.09 0.14 0.20

2 .18 .23 .26

3 .35 .40 .45

4 .48 .56 .62

5 .67 .94 .94

0.09 0.06 0.13 0.16 

__ .19 .15 .27 .31 

.30 .31 .36 .53

.43 .48 .46 .80 

.58 .71 .65 1.15

6 .91 1.05 1.07 .76 1.00 .98 1.60

7 1.33 1.41 1.24

8

Volution height 

1 0.04 0.03 0.03 0.05

2 .05 .04 .04 .06

3 .07 .09 .08 .07

4 .08 .03 .12 .09

5 .12 .14 .15 .14

6 .12 .15 .15 .15

7 .16 .16 .20

1.30 1.23 1.38 ____ 

1.55

0.03 0.04 0.03 0.03 0.03 

.04 .06 .04 .04 .05 

.07 .065 .08 .06 .07 

.05 .04 .07 .07 .09 

.11 .10 .11 .08 .11 

.11 .12 .11 .14 .15 

.14 .13 .14 .13

8 .19 .13

Wall thickness 

1 0.01

2 .01 .015

3 .01 .. .. .

.01 .02 .01 __ ____ 

.02 .015 .01 .02

4 .02 .012 .02 .02 .02 .02

5 .02 .025 .03

6 .03 .035 .025

7 .03

8

Tunnel width (axials) or 

septal spacing 

1 0.01 0.02 003 0.03 0.01 0.02

2 .02 .03 .05 .05

3 .03 .04 .05 .05

4 .04 .08 .06 .07

5 .05 .09 .12

6 .05 .15 .17 .16

7 .06 .21 . _ .

.02 .03 .035 .02 .025 

.04 .04 .05 .03 .04 

.04 .05 .05 .05 .09 

.05 .07 .10 .075 .15 

.06 .11 .145 .10 .19 

.13 .16 .16

8 .10

Wedekindellina sp.

Plate 17, figures 7-9

Discussion. -Limestone attributed to the Vanport 

Limestone Member of the Allegheny Formation in 

Mercer County, Pa., has scattered immature fusulinids. 

The three specimens figured suggest the inner volutions 

of Wedekindellina and are at the proper stratigraphic 

level to represent W. euthysepta (Henbest). Their 

presence is worth noting, but a positive identification 

cannot be made. The specimens are from locality 86. 

Measurements are given in table 10.

Beedeina sp. aff. B. novamexicana (Needham),1937

Plate 8, figures 1-3, 5, 6; plate 9, figures 3-10; plate 10, 

figures 1-8, 10, 11

Description. - Measurements for these specimens are 

given in table 11 and on figure 15. The specimens are all

small relative to the types of Beedeina novamexicana 

and to many of the specimens assigned to the species by 

Dunbar and Henbest (1942, p. 113-115). Most of the 

specimens from Kentucky have five to six volutions, 

whereas specimens from New Mexico and Illinois commonly have eight. Most specimens have the inflated 

fusiform shape with convex to irregular lateral slopes, a 

narrow irregular tunnel bordered by high narrow 

chomata. The septa are numerous but irregularly 

spaced, especially in the outer volutions. They are tightly and irregularly fluted throughout. The proloculus is 

relatively large, measuring 0.10-0.21 mm in maximum 

outer diameter, most specimens showing 0.15 mm or 

more. One of the specimens studied (pi. 9, fig. 3) has a 

more elongate fusiform shape in the outer volutions and 

suggests a similarity to B. illinoisensis (Dunbar and 

Henbest, 1942); however, it has less regular septal 

fluting.
SYSTEMATIC DESCRIPTIONS

TABLE 6. -Measurements (in mittimeters) for Fusulinella stouti Thompson, 1936- Continued

39

Character Measurements of fossils shown in plate 5, figures 19-26

19 20 21 22 23 24 25 26

Diameter of proloculus __________ 0.08 

Radius vector

1 _____________________ 0.09

2 _____________________ .14

3 _____________________ .20

4 _____________________ .29

5 _____________________ .38

6 _____________________________

7 _____________________________

8 _____________________ ____ 

Half length

1 _______________________________

2 ______________________________

3 _____________________ ____

4 _______________________________

5 ______________________________

6

7 ______________________________

8 ______________________________

Volution height

1 _____________________ 0.03

2 _____________________ .05

3 _____________________ .05

4 _____________________ .09

5 _____________________ .09

6 _____________________________

7 _____________________________

8 ____________________ ____ 

Wall thickness

1 _____________________ 0.005

2 _____________________ .01

3 _____________________ .01

4 _____________________ .015

5 ______________________________

6 ______________________________

7 _______________________________

8 ______________________________

Tunnel width (axials) or 

septal spacing

1 _____________________ 0.02

2 _____________________ .025

3 _____________________ .04

4 _____________________ .03

5 _____________________ .04

6 _____________________________

7 ____________________

0.10

0.12 

.18 

.25 

.33 

.43 

.53

0.12

.24

.38

.65

.80

1.12

0.06

.06

.07

.08

.105

.10

.02 

.03 

.025 

.02

.02 

.05 

.07 

.09 

.11

0.17

0.08 

.13 

.19 

.26 

.36 

.45 

.58 

.72

0.11

.23

.42

.61

.94

1.10

1.33

1.78

0.02 

.05 

.06 

.07 

.10 

.09 

.13 

.15

0.02

.02

.03

.035

.03

.03

.05

.04

0.04 

.04 

.07 

.11 

.16 

.23

0.14

0.06 

.11 

.15 

.21 

.31 

.46 

.60

0.08

.17

.31

.46

.70

1.13

1.41

0.02 

.04 

.05 

.07 

.10 

.13 

.16

.01 

.02 

.02 

.03 

.04

.03

.035

.05

.08

.10

0.08

0.06

.105

.17

.24

.34

.46

0.02

.045

.065

.06

.10

.12

.015

.015

.02

.03

.02 

.035 

.05 

.05

0.10

0.08 

.14 

.20 

.30 

.39 

.52 

.73

0.14

.29

.41

.59

.86

1.28

1.73

0.03 

.05 

.06 

.10 

.08 

.14 

.20

0.02 

.02 

.02 

.03 

.02 

.04

0.04 

.04 

.06 

.08 

.11 

.14

0.09

0.08 

.15 

.19 

.29 

.41 

.55

0.13 

.24

0.03 

.04 

.06 

.10 

.13 

.14

0.01

.015

.03

.03

0.03 

.04 

.06 

.09 

.12

0.08

0.07 

.10 

.17 

.23 

.32 

.44 

.56 

.71

0.025

.03

.05

.06

.08

.12

.12

.16

.015

.015

.02

.025

.035

0.02

"~02 

.03 

.045 

.05

The spirotheca of B. sp. aff. B. novamexicana differs from 

that of most of the species of Beedeina seen in Kentucky in 

that it has a thin protheca but thick deposits of both inner and 

outer tectoria. The tectoria make the wall twice to three 

times the thickness of the protheca.

Distribution. -B. sp. aff. B. novamexicana is common 

in the Yeargins limestone member of the Trade water 

Formation in both surface and subsurface samples. It 

has been described from localities 18 in Union County, 

39 in Butler County, and 40 and 41 in Ohio County, Ky. 

The specimens from Illinois were described from the 

Stonefort Limestone Member of the Spoon Formation.

Beedeina leei (Skinner), 1931

Plate 8, figures 4, 7-9; plate 9, figures 1, 2; plate 10,figures 9, 12-17; 

plate 15, figures 7-13; plate 17, figures 10-15 24, 25

Fusulina leei Skinner, 1931, p. 257, 258, pi. 30, figs. 4-6; non 

Thompson, 1934, p. 301-303, pi. 21, figs. 3, 7, 10, 18; non Thompson, 1935, p. 305, pi. 26, figs. 17-19; Dunbar and Henbest, 1942, p.

109-111, pi. 5, figs. 1-8, pi. 6, figs. 1-10; Smyth, 1957, p. 266-267,

pi. 2, figs. 17-19. 

Fusulina aff. F. leei Thompson and Riggs, in Thompson, Shaver, and

Riggs, 1959, p. 781, pi. 106, figs. 25-28, 33. 

Fusulinella serotina Thompson, 1936, p. 677-678, pi. 90, figs. 1-4,

pi. 21, figs. 9. 

Fusulina serotina (Thompson and Riggs), in Thompson, Shaver, and

Riggs, 1959, p. 780-781, pi. 106, figs. 29-32.

Description. - Measurements for this species are given 

in table 12 and on figure 16. This species of Beedeina is a 

relatively primitive form, in which the inner volutions 

are fusulinellid and only the outer volutions have septal 

fluting regular and intense enough to qualify for assignment to Beedeina. The specimens are fusiform to 

elongate fusiform and attain lengths of about 5 mm in 

7V2-8 volutions. The proloculus is small, averaging 

about 0.10 mm and ranging from 0.08 to 0.16 mm in 

maximum outside diameter.
40 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS

0.4

0.3 

0 0.2

0.1

uj 0.10 

^ 0.05

1 0-3

I

Z 0.2

O

3 °- 1 

o

3.5 

3.0 

2.5 

2.0

< 1.5

1.0 

0.5

. > :

. !?£ ' / ft*****"

 .% *% « *" 

y*

0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 0.9 1.0 

RADIUS VECTOR

Figure 11.-Graphs for Fusulinella stouti Thompson, 1936. All 

measurements are in millimeters, and the values for each attribute 

are plotted against the radius vector.

The spirotheca is thin and composed of protheca and 

variable amounts of tectoria. Dunbar and Henbest 

(1942, p. 109) reported a thin inner tectorium and 

thicker outer tectorium. The specimens from Kentucky, 

Ohio, and Pennsylvania show a thick inner tectorium in 

the inner volutions, thinning in the outer volutions. The 

outer tectorium is thin in the inner volutions and absent 

in the outer volutions. The septa are closely but irregularly spaced. They are relatively straight in the 

inner volutions but become tightly fluted in the poles 

and across the shell in the outer volutions. The tunnel is 

generally straight and is bounded by high broad 

chomata in the inner volutions and by lower chomata in

the outer volutions. The tunnel is high, extending more 

than half the volution height in most chambers.

Comparison and remarks, B. leei is easily recognized 

by its fusulinellid initial volutions and Beedeina outer 

volutions, by its numerous, regularly expanding volutions, and by its thin spirotheca. The forms illustrated by 

Thompson from Iowa (1934) and Oklahoma (1935) have 

fewer, more loosely coiled volutions and do not show the 

typical Fusulinella-Beedeina transition. Specimens 

described by Thompson (1936) from Ohio as Fusulinella 

serotina fall completely within the range of variation of 

B. leei. B. leei differs from B. carmani in having a 

smaller proloculus, more volutions, and less widely 

spaced septa.

Distribution. -B. leei is a common form in beds of early Desmoinesian age from Oklahoma to Pennsylvania. 

The Kentucky specimens are described from the 

Yeargins limestone member of the Tradewater Formation from localities 18 in Union County, 32 in Christian 

County, 35 in Bulter County, and 40 in Ohio County. The 

Ohio specimens are described from the Putnam Hill 

Limestone Member of the Allegheny Formation at 

locality 79 in Coshocton County, and the Pennsylvania 

specimens are described from the Mahoning Limestone 

of Rogers (1858) at locality 87 in Armstrong County. 

Dunbar and Henbest (1942, p. Ill) reported this species 

from their Curlew Limestone Member of the Spoon Formation. That unit is younger than the Curlew Limestone 

Member of Kentucky.

Beedeina sp. aff. B. leei (Skinner), 1931

Plate 8, figures 10, 13, 15, 17; plate 12, figures 8-10

Several specimens from cores in western Kentucky 

bear a resemblance to Beedeina leei but cannot be 

assigned to this species with any confidence. 

Measurements for these specimens are given in table 12. 

A specimen from locality 21 in Webster County (pi. 8, 

fig. 10) shows inner volutions similar to those of B. leei, 

but the outer volutions do not expand in length like the 

typical form. Three specimens from locality 31 in Christian County are illustrated as plate 8, figures 13, 15, and 

17. The microspheric axial shown in figure 13 is probably the same form as those shown in figures 15 and 17. 

These forms remain fusiform throughout their growth 

and do not elongate as in typical B. leei. The specimens 

illustrated on plate 12 as figures 8-10 may represent B. 

leei but are also less elongate and cannot be assigned 

with any certainty. They occur in a surface sample 

associated with Wedekindellina. All these specimens are 

found in association with Wedekindellina in beds assigned to the Yeargins limestone member of the Tradewater Formation.
SYSTEMATIC DESCRIPTIONS 41

TABLE 7.-Measurements (in millimeters) for Fusulinella iowensis Thompson, 1934

[Leaders (__), data not available or character absent]

Diameter of proloculus ___ 

Radius vector 

1

2

3

4

5

6

7

8

9

Half length 

1

2

3

4

5

6

7

8 _ _

9

Volution height 

1

2

3

4

5

6

7

8

9

Wall thickness 

1

2 _

3

4

5 _

6

7

8

9

Tunnel width (axials) or 

septal spacing 

1

2

3

4

5

6

7

8

Measurements of fossils shown in plate 6,

8

0.05

0.08 

.16 

.23 

.33 

.45 

.57 

.71 

.88

0.09 

.14 

.24 

.33 

.43 

.56 

.73 

.95 

1.22

0.02 

.02 

.06 

.07 

.10 

.12 

.12 

.14 

.17

0.02 

.02

0.02 

.025 

.03 

.04 

.05 

.07 

.07 

.14

9

0.10

0.10 

.16 

.25 

.34 

.42 

.56 

.71

0.15 

.23 

.41 

.53 

.67 

.88 

1.17

0.03 

.06 

.08 

.09 

.09 

.15

 

0.02 

.02 

.03 

.06 

.08 

.08 

.11

10

0.08

0.09 

.13 

.19 

.26 

.36 

.47 

.58 

.68

0.11 

.19 

.25 

.35 

.48 

.70 

.82 

1.07

0.03 

.04 

.05 

.07 

.07 

.10 

.11 

.10

 

0.02 

.02 

.02 

.04 

.07

11

0.11

0.15 

.25 

.36 

.46 

.62 

.76

0.07 

.09 

.11 

.10 

.10 

.15

0.01 

.02 

.02

0.02 

.04 

.06 

.07 

.08

15

0.06

0.10 

.15 

.22 

.29 

.40 

.51 

.65

0.04 

.05 

.06 

.07 

.07 

.11 

.14

.02 

.02

0.02 

.04 

.04 

.04 

.07 

.07

figures 8-11, 15, 12-14, 16

12

0.10

0.06 

.10 

.19 

.28 

.38 

.49 

.65 

.77 

.95

0.08 

.14 

.22 

.33 

.48 

.64 

.84 

1.14 

1.33

0.02 

.04 

.06 

.11 

.11 

.13 

.16 

.16 

.18

0.02 

.015 

.03

.02 

.03 

.04 

.05 

.07 

.11 

.15

13

0.10

0.07 

.12 

.19 

.26 

.35 

.46 

.60 

.78 

.95

0.07 

.12 

.18 

.27 

.37 

.53 

.72 

.97 

1.18

0.02 

.05 

.07 

.07 

.095 

.11 

.14 

.18 

.19

.02 

'.03

0.02 

.04 

.05

.08 

.11 

.17

.21

14

0.12

0.08 

.13 

.19 

.30 

.39 

.51 

.65

0.02 

.05 

.06 

.10 

.10 

.12 

.13

0.02 

.02

0.01 

.025 

.03 

.04 

.04

16

0.07

0.10 

.16 

.24 

.33 

.44 

.56 

.72

0.14 

.25 

.40 

.54 

.78 

.93 

1.22

0.04 

.06 

.08 

.10 

.10 

.12 

.155

 -

0.03 

.04 

.04 

.06 

.06

Beedeina sp. aff. B. spissiplicata (Dunbar and Henbest), 1942

Plate 8, figures 11, 12, 14, 16, 18-21; plate 9, figs. 11-16

Specimens from cores in wells resemble some of the 

specimens illustrated by Dunbar and Henbest (1942, pi. 

7, figs. 1-12). The specimen they selected as the 

holotype (their figs. 3, 4) is more regular in coiling and 

fluting than the other specimens. The specimens from 

the Kentucky cores are similar to the specimen they illustrated as figure 5. The axis of coiling is apparently 

not straight, the tunnel is less regular, and the shape is 

not as constant as in the holotype.

Measurements for the Kentucky specimens are given 

in table 13 and on figure 17. These measurements agree, 

in general, with the data provided by Dunbar and 

Henbest (1942), although some of the Kentucky 

specimens are a little larger at comparable volutions.

The Kentucky specimens were found associated with 

Wedekindettina at the following locations: 31 in Christian County in core DDK, G16 at a depth of 36-138 ft; 33 

in Muhlenberg County in core DDK, G10 at a depth of 

150 ft; 35 in Butler County in core Gil-23 at a depth of 

177.6 ft; 40 in Ohio County in core Gil-9 at a depth of 

332 ft.
42 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS 

TABLE 7. -Measurements (in millimeters) for Fusulinella iowensis Thompson, 1934 -Continued

Character

14

Measurements of fossils shown in plate 15, figures 14-18

16 18

Diameter of proloculus _. 

Radius vector 

1 _________________

Half length

1 _______

Volution height 

1 ______.

Wall thickness

1 ________

2 _____.

3 ______.

Tunnel width (axials) or 

septal spacing

0.065

0.08 

.12 

.17 

.22 

.32 

.44 

.60 

.74

0.07

.11

.19

.30

.47

.66

.90

1.09

0.025

.04

.05

.05

.09

.12

.16

0.09

0.08

.125

.18

.25

.35

.46

.63

0.10

.16

.28

.40

.71

.94

1.11

0.018

.04

.05

.07

.09

.11

.16

.03

0.10

0.10

.17

.26

.35

.485

.62

.91

0.03

.065

.075

.09

.13

.135

.185

.02

.015

.035

.04

.065

0.06

0.07

.13

.20

.29

.425

.54

.68

0.02 

.06 

.07 

.08 

.14 

.12 

.14

.02

.035

.06

0.06

0.08

.12

.18

.26

.365

.52

.665

0.12

.21

.34

.45

.55

.94

1.03

0.03 

.04 

.06 

.08 

.11 

.15 

.15

0.015

.018

.025

1

2

3

4

5

6

7

8

0.03

.04

.04

.06

.07

.08

.10

0.025

.045

.04

.07

.10

.12

0.02

.03

.025

.035

.04

.07

0.025

.025

.04

.05

.05

.06

0.03

.04

.055

.055

.08

.10

*~

The Illinois specimens were described from a 

limestone above the Colchester No. 2 coal bed. In Kentucky, the specimens come from the lower part of the 

Yeargins limestone member.

Beedeina henbesti Douglass n. sp.

Plate 16, figures 29-31

Name derivation.-Named for Lloyd G. Henbest, who collected the 

sample.

Diagnosis. - Shell small, fusiform, with nearly straight 

to concave lateral slopes and pointed poles. Septa tightly 

and regularly fluted throughout the shell. Proloculus 

relatively large.

Description. -Measurements are presented in table 14 

and on figure 18. The shell develops regularly, maintaining a fusiform shape from its early volutions, expanding 

only slightly more rapidly along the axis than equatorially. The lateral slopes tend to be concave in most volutions. The proloculus averages 0.17 mm in four 

specimens, two specimens attaining a maximum outer 

diameter just greater than 0.20 mm. The spirotheca is 

thin and is composed of protheca and variable amounts 

of inner tectorium. A thin outer tectorium is deposited 

locally along the lateral slopes. The septa are intensely 

and regularly fluted throughout the shell. The septa are 

numerous and relatively evenly spaced, about 25 septa 

per volution in the fourth volution. The tunnel wanders a
SYSTEMATIC DESCRIPTIONS 

TABLE 7. -Measurements (in millimeters) for Fusulinella iowensis Thompson, 1934 -Continued

43

Character Measurements of fossils shown in plate 15, figures 19-26

19 20 21 22 23 24 26

Diameter of proloculus __________ 0.13 0.065 0.06 0.18 0.095 0.07 0.09 0.09 

Radius vector

1 _____________________ 0.11 0.09 0.065 0.13 0.10 0.09 0.09 0.12

2 _____________________ .20 .12 .13 .21 .18 .14 .15 .195

3 _____________________ .31 .165 .21 .315 .27 .20 .21 .28

4 _____________________ .42 .26 .30 .45 .39 .31 .29 .39

5 _____________________ .56 .375 .41 .58 .54 .40 .41 .51

6 _____________________ .725 .51 .54 .76 .69 .51 .52 .645

7 _____________________ __ .67 .71 .96 __ .67 __ .84

8 _____________________ __ __ __ __ __ __ __ ____

9 _____________________ __ __ __ __ __ __ __ __

Half length

1 ____________________ 0.14 __ __ 0.19 0.10 0.10 __ ____

2 _____________________ .23 __ __ .31 .16 .17 __ ____

3 _____________________ .36 __ __ .445 .29 .29 __ __

4 _____________________ .465 __ __ .62 .46 .40 __ ____

5 _____________________ .635 __ __ .90 .68 .55 __ ____

6 _____________________ .94 __ __ 1.10 .965 .755 __ ____

7 _____________________ 1.05 __ __ 1.45 1.11 .95 __ ____

8 _____________________ __ __ __ __ __ __ __ ____

9 _____________________ __ __ __ __ __ __ __ ____

Volution height

1 _____________________ 0.03 0.02 0.03 0.04 0.03 0.02 0.035 0.04

2 _____________________ .09 .04 .04 .09 .07 .04 .05 .07

3 _____________________ .11 .04 .075 .10 .09 .06 .06 .08

4 _____________________ .12 .10 .09 .13 .12 .10 .08 .09

5 _____________________ .14 .11 .10 .14 .14 .10 .11 .12

6 _____________________ .17 .125 .13 .18 .15 .11 .12 .13

7 _____________________ __ .16 .16 .20 __ .16 __ .19

8 _____________________ __ __ __ __ __ __ __ ____

9 ________..____________ __ __ __ __ __ __ __ ____

Wall thickness

1

2 _____________________ __ __ __ __ __ __ __ 0.02

3 _____________________ __ __ __ __ __ __ .01 .02

4 _____________________ __ __ .025 __ __ __ .012 .025

5 _____________________ __ .035 .04 .03 __ __ .035 .04

6 _____________________ __ .05 .04 .06 __ __ __ .04

7 _____________________ __ .07 .06 __ __ __ __ ____

8 _____________________ __ __ __ __ __ __ __ ____

9 _____________________ __ __ __ __ __ __ __ ____

Tunnel width (axials) or 

septal spacing

1 _____________________ 0.03 0.025 0.02 0.02 0.03 0.03 0.025 0.02

2 _____________________ .05 .04 .02 .03 .04 .04 .03 .035

3 _____________________ .07 .04 .03 .06 .065 .06 .03 .04

4 _____________________ .07 .05 .03 .09 .09 .08 .05 .04

5 _____________________ .10 .055 .04 .11 .125 .12 .07 .05

6 _____________________ .14 .105 .04 .17 .19 __ __ .05

7 _____________________ __ .125 __ __ __ __ __ __

little through the equatorial plane. It extends about half 

the volution height and is bordered by high narrow 

chomata.

Comparison and remarks. -Beedeina henbesti resembles some specimens of B. leei but has more regularly intensely fluted septa, more concave lateral slopes, and a 

large proloculus. It differs from B. carmani in shape and 

especially in the septal fluting; the prolocular diameter 

is comparable.

Distribution.-B. henbesti occurs in the Vanport 

Limestone Member of the Allegheny Formation in Stark 

County, Ohio, at locality 83. It is associated with the 

genus Wedekindellina.

Specimens studied.-Eleven specimens from collection

f 13109 were studied. The species was not recognized in 

other samples of the Vanport Limestone.

Designation of types.-The specimen illustrated on 

plate 16 as figure 29 is designated the holotype. The 

other specimens are paratypes.

Beedeina ashlandensis Douglass n. sp.

Plate 11, figures 1-19

Name derivation.-Named for Ashland, Ky., where outcrops on the 

Kentucky and Ohio sides of the Ohio River contain this species.

Fusulina pumila Thompson, 1934 (part) of Dunbar and Henbest, 1942, 

pi. 5, figs. 13-15.

Diagnosis. - Shell relatively small and rhombodial, expanding regularly and symmetrically. Chomata high and
44 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS

_u °-3

i 9 0.2

0.1

_1 UJ 0.10

^ y 0.05

tt- 

<2 0.3

LU

I

Z 0.2 

O

3 °- 1 

o >

3.5 

3.0

Z 

^ 2.0 

i_i_

< 1.5 

1.0 

0.5

Figure 12. 

measuren 

are plotte

narrow, 

spaced ai 

Descrij. 

and on fi 

boidal sh< 

addition < 

little elor 

the poles 

0.13mm, 

to 0.19 m 

The spi 

generally 

torium is 

and close

recnilarlv

1 1 1 1 

V*.J*

00

**t</iO-»* .M***** 

 J* .

-

 ! 

« * * ? *

yfcl- » *

«^* III ....

1 1 1 1

 *:; . " . -

 * 

J'a Tr ^^ ^^ ^

P ' 00 

f 0

. ... ....

1 1 1 1

0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 0.9 1.0 

RADIUS VECTOR

-Graphs for Fusulinella iowensis Thompson, 1934. All 

nents are in millimeters, and the values for each attribute 

id against the radius vector.

bordering a narrow tunnel. Septa closely 

id fluted throughout the shell. 

)twn. - Measurements are presented in table 15 

gure 19. Most specimens start with a rhomape and maintain a similar shape through the 

rf whorls that tend to parallel each other with 

igation or increase in chamber height toward 

. The proloculus averages a little more than 

ranging in maximum outer diameter from 0.08 

m.

rotheca is thin and composed of a protheca and 

well developed inner tectorium. An outer tecpresent only sporadically. Septa are numerous 

sly spaced. They are fluted strongly but irfnrmincr pVmirtVwarlofc i-n fVio in'/iini-fir r»4? fV»/» -t-nv>

nel. The tunnel tends to be straight and widens slowly. 

It is bordered by high narrow chomata. The tunnel extends about half the volution height. 

Comparison and remarks. -Beedeina ashlandensis is 

similar to some of the specimens included in Fusulina 

pumila Thompson by Dunbar and Henbest (1942, pi. 5, 

figs. 13 and 14.). It is more regular in its growth form 

from inner to outer volutions than most species of Beedeina, but it fits in the general group within the genus 

that has attributes of Fusulinella in the inner volutions 

and those of Beedeina in the outer volutions. The change 

is primarily in the fluting of the septa but also includes 

the chomata shape and the spirothecal structure with little outer tectorium in the adult chambers. 

Distribution. -This species is described from localities 

61, 62, and 63 in Greenup County, Ky., and just across 

the Ohio River in Lawrence County, Ohio. The beds 

represent the upper part of the Allegheny Formation 

and probably are most closely tied to the Columbiana 

Limestone Member as mapped in Ohio. The specimens

TABLE 8. -Measurements (in millimeters) for Fusulinella sp.

[Leaders ( __ ), data not available or character absent]

Character Measurements of fossils shown in plate 6, figures 20, 17-19, 21, 22

20 17 18 19 21 22

Diameter of 

proloculus 0.09 0.11 0.09 0.07 0.09 0.10

Radius vector 

1 0.07 0.06 0.09 0.06 0.06 0.11

2 .11 .10 .15 .09 .12 .16

3 .16 .15 .21 .13 .18 .24

4 .24 .21 .28 .20 .28 .34

5 .32 .33 .38 .28 .36 .46

6 .43 .42 .52 .35

7

Half length 

1 0.05 0.06

2 .14 .11

3 .26 .21

4 .54 .38

5 .80 .63

6 1.16 1.01

7 1.43

Volution height 

1 0.02 0.02 0.02 0.02 0.02 0.03

2 .04 .04 .03 .03 .04 .05

3 .05 .05 .05 .04 .06 .08

4 .08 .06 .06 .06 .10 .10

5 .08 .11 .07 .08 .09 .13

6 .10 .11 .10 .08

7 .11

Wall thickness 

1 0.01 0.01

2 .01 .02

3 .02 .01 .02 .02

4 .02 .03 .02 .02 .02

5 .03 .02 .02 .04

6 .05 .02 .03

7 .04

Tunnel width (axials) 

or septal spacing 

1 0.02 0.01 0.02 0.02 0.03

2 .02 .02 ' .03 .02 .03 .04

3 .03 .03 .02 .03 .04

4 .04 .06 .03 .03 .045

5 .04 .08 .04 .04 .11

6 .05 .14 .05 .06

7
SYSTEMATIC DESCRIPTIONS 45

<J.--t

0.3

Q 0.2

0.1

ui 0.10

:=: 0.05

H

H 

X

X

Z 0.2 

O

1 

D 0.1

_J 

O

3.5

3.0

H 2.5

§ 2.0

u. 

< 1.5

1.0

0.5

I I I

-

 

_ 

 

 

-

 

 

 * *

-

. /v **

-

-

-

-

 

_

 ? **! I

I I I I

-

-

-

 

-

-

-

-

-

I I I

0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 0.9 1.0 

RADIUS VECTOR

Figure 13.-Graphs for Fusulinella sp. All measurements are in 

millimeters, and the values for each attribute are plotted against 

the radius vector.

illustrated by Dunbar and Henbest (1942) are from the 

limestone tentatively assigned to the Seahorn? 

Limestone in Randolph County, 111.

Specimens studied.-The description is based on 53 

oriented thin sections from three collections. Comparative material was studied from the Dunbar and 

Henbest and the Thompson collections.

Designation of types.-The specimen illustrated as 

figure 18 on plate 11 is designated the holotype. The 

other illustrated specimens are paratypes.

Beedeina girtyi (Dunbar and Condra), 1927

Plate 12, figures 1-7; plate 13, figures 6, 9, 10

Fusulina ventricosa Meek and Worthen, 1873, p. 560, pi. 24, figs. 6, 

8a.

Girtyina ventricosa Staff, 1912, p. 164, pi. 18, figs. 2, 5, 7. 

Fusulinella girtyi Dunbar and Condra, 1927, p. 76-78, pi. 2, figs. 1-5. 

Beedeina girtyi Galloway, 1933; p. 401, pi. 36, fig. 17; Ishii, 1958,

p. 39, pi. 1, fig. 9. 

Fusulina girtyi Thompson, 1934, p. 314-316, pi. 22, figs. 1, 5, 7, 20;

Dunbar and Henbest, 1942, p. 115-117, pi. 11, figs. 1-17, pi. 12,

figs. 2-8, 10, 11 (designated lectotype).

Description. -Measurements are given in table 16 and 

on figure 20. Shell inflated fusiform of moderate size, attaining lengths of about 5 mm and widths of 3 mm in 

eight to nine volutions. Some specimens attain 10 volutions and reach a length of 5.5 mm and a width of 3.5 

mm. The inner volutions tend to be fusiform and to have 

concave lateral slopes, and the general shape is commonly maintained throughout, but some specimens have 

straight to slightly convex lateral slopes in the outer 

volutions. The proloculus in megalospheric forms ranges 

from 0.16 to 0.21 mm in maximum outer diameter. One 

microspheric form was sectioned, but the size of the proloculus could not be determined accurately.

The spirotheca is composed of a thin protheca and 

commonly heavy tectoria. In better preserved 

specimens, a fine fibrous structure can be determined in 

the diaphanotheca. The septa are strongly and regularly 

fluted in the outer volutions. They show less fluting in 

the inner volutions. The appearance of closed loops of 

the septa in equatorial section is characteristic of the 

outer volutions in this species. The loops are formed by 

the touching of opposing folds in the septa high up in the 

chamber. They do not indicate the presence of cuniculi. 

The tunnel is narrow, and its path ranges from straight 

to quite irregular. It is bordered by high broad chomata 

in the inner volutions and high narrow chomata in the 

outer volutions. No axial filling, as such, is recognized, 

but the chomata and epithecal deposits in the inner volutions do limit the open spaces in the inner chambers.

Comparison and remarks. -Beedeina girtyi is the type 

species for the genus and is an easily recognized form, 

but the limits of its variability are not well known. The 

type specimen was designated by Dunbar and Henbest 

(1942) in a footnote to a table of measurements. The 

specimen selected is more elongate than many of the 

other specimens illustrated for this species. It has a form 

ratio of about 1.7 and is at the high end of the form 

ratios reported for the species. This fact is only significant because B. girtyi is generally reported to occur 

with B. illinoisensis and (or) B. haworthi or B. stookeyi, 

all of which are reported to differ from B. girtyi primarily in being more elongate forms. Both Thompson (1934) 

and Dunbar and Henbest (1942) suggest the possibility 

that these specimens are variants and not distinct 

species. Perhaps collections from Kansas or Illinois 

might permit a study of the range of variability to help 

decide this issue. The collections available from Kentucky contain some elongate specimens and many in-
46

TABLE 9. -Measurements (in millimeters) for Wedekindellina euthysepta (Henbest), 1928

[Leaders (__), data not available or character absent]

Character

Diameter of proloculus _ 

Radius vector 

1

2

3

4

5

6

7

8

9

10

Half length 

1

2

3

4

5

6

7

8

9

Volution height 

1

2

3

4

5

6

7

8

9

10

Wall thickness 

1

2

3

4

5

6

7

8

9

Tunnel width (axials) or 

septal spacing 

1

2

3

4 _

5

6

7

8

9

10

Measurements of fossils shown in plate 7, figures 1, 4

1

0.11

0.08 

.11 

.15 

.20 

.24 

.30 

.34 

.40

0.12 

.19 

.29 

.40 

.52 

.67 

.83 

1.04

0.02 

.04 

.04 

.05 

.04 

.06 

.04 

.09

0.01 

.01 

.02 

.02

.02

.02 

.03 

.04 

.05 

.06 

.08

4

0.09

0.07 

.10 

.13 

.18 

.23 

.28

0.14 

.23 

.32

.44 

.59 

.78

0.02 

.03 

.03 

.06 

.05 

.06

"61

 

 

"Toi

.04 

.05 

.07

 

6

0.03?

0.06 

.09 

.12 

.19 

.24 

.31 

.40

0.08 

.16 

.26 

.36 

.58

 

0.02 

.03 

.03 

.03 

.04 

.07 

.09

 

.01 

.02 

.02 

.02 

.04

 

.015 

.02 

.04 

.09 

.08

2

0.11

0.08 

.11 

.16 

.21 

.26 

.32

0.17 

.30 

.43 

.63 

.78 

.88

0.02 

.03 

.05 

.05 

.05 

.06

 

"61

.015

.025

 

"02 

.03 

.035 

.04 

.06

7

0.07

0.06 

.09 

.13 

.17 

.22 

.28 

.36 

.44 

.55

 

0.01 

.02 

.04 

.04 

.045 

.06 

.07 

.08 

.11

 

.01

"62 

.02

~T6i

0.02

.02 

.04 

.03 

.045 

.05 

.045 

.065

, 6, 2, 7, 3, 8, 5

3

0.10

0.06 

.10 

.13 

.18 

.24 

.31 

.40

0.14 

.21 

.37 

.53 

.74 

1.09 

1.40

0.015 

.03 

.03 

.045 

.065 

.07 

.09

 

.02 

.02

.025 

.04

"6l 

.02 

.04 

.06 

.07 

.09

 

8

0.08

0.06 

.08 

.11 

.15 

.21 

.28 

.37 

.45 

.55

0.12 

.21 

.33

.45 

.58 

.75 

.89 

1.15 

1.37

0.01 

.01 

.025 

.03 

.06 

.07 

.09 

.09 

.10

.02

.02

 

"6l 

.02 

.03 

.06 

.07 

.08 

.11 

.14

5

0.06

0.06 

.09 

.13 

.17 

.28 

.39 

.49 

.60

 

0.02 

.02 

.04 

.04 

.10 

.10 

.10 

.11

"005 

.01 

.01 

.01 

.02 

.04

"6l5 

.02 

.03 

.04 

.05 

.07

 

flated but shorter specimens. The problem cannot be 

resolved with the specimens available from Kentucky.

Distribution. -B. girtyi is present in the Providence 

Limestone Member of the Sturgis Formation in western 

Kentucky. It is common in the Herrin Limestone 

Member of the Carbondale Formation in Illinois and the 

Fort Scott Limestone of Kansas.

Specimens studied.-The specimens illustrated by 

Dunbar and Condra (1927) and Dunbar and Henbest 

(1942) were studied along with specimens from locality 

27 in Hopkins County and locality 29 in Muhlenberg 

County, Ky.

Beedeina sp. aff. B. haworthi (Beede), 1916

Plate 13, figures 1-5, 7, 8

Discussion. - Several specimens from the Providence 

Limestone Member of the Sturgis Formation in 

Muhlenberg County, Ky., resemble specimens illustrated by Dunbar and Henbest (1942) as Fusulina 

haworthi and as F. illinoisensis. These specimens tend 

to be more elongate and to have smaller chomata in the 

inner volutions than do the specimens that Dunbar and 

Henbest assigned to F. girtyi. Beedeina girtyi (as it is
SYSTEMATIC DESCRIPTIONS

TABLE 9. -Measurements (in millimeters) for Wedekindellina euthysepta (Henbest), 1928 -Continued

47

Diameter of proloculus _ 

Radius vector 

1

2

3

4

5

6

7

8

9

10

Half length 

1

2

3

4

5

6

7

8

9

Volution height 

1

2

3

4

5

6

7

8

9

10

Wall thickness 

1

2

3

4

5

6

7

8

9

Tunnel width (axials) or 

septal spacing 

1

2

3

4

5

6

7

8

9

10

Measurements of fossils shown in plate 7, figures 10, 11, 9, 13, 14, 12, 16, 18

10

0.06

0.06 

.09 

.13 

.17 

.28 

.39 

.49 

.60

0.02 

.02 

.04 

.04 

.10 

.10 

.10 

.11

.005 

.01 

.01 

.01 

.02 

.04

.015 

.02 

.03 

.04 

.05 

.07

11

0.09

0.06 

.08 

.11 

.14 

.17 

.25 

.32 

.39 

.48

0.11 

.21 

.30 

.38 

.52 

.69 

.82 

1.01 

1.34

0.01 

.02 

.03 

.03 

.03 

.06 

.08 

.08 

.09

0.075 

.01 

.02

.04

0.02 

.02

.06

.06 

.07 

.09

____

9

0.08

0.06 

.09 

.13 

.19 

.25 

.31 

.37 

.46

0.14 

.24 

.37 

.55

1.40

0.015 

.025 

.04 

.06 

.055 

.06 

.06 

.09

0.01 

.015 

.02

~02 

.035

.05 

.07

 

13

0.06

0.06 

.10 

.14 

.21 

.28 

.35

 

0.10 

.20 

.34 

.50 

.76 

1.14

0.02 

.03 

.05 

.07 

.07

 

.01 

.015 

.02

0.02 

.02 

.03 

.04 

.07 

1.05

 

14

0.08

0.065 

.095 

.14 

.18 

.26 

.35 

.41

 

0.16 

.29 

.41 

.60 

.87 

1.11 

1.53

0.015 

.045 

.09 

.13 

.21 

.30 

.36

 

0.018 

.02 

.015 

.02

0.015 

.02 

.03

.04

 

12

0.05

0.06 

.09 

.12 

.15 

.20 

.26 

.34 

.40 

.51

0.11 

.20 

.31 

.42 

.51 

.81 

1.01 

1.19 

1.30

0.01 

.02 

.03 

.025 

.05 

.06 

.08 

.06 

.11

.01

.01 

.015 

.02

.04 

.04

::::

16

0.05

0.07 

.09 

.11 

.15 

.20 

.28 

.34 

.41 

.49

0.08 

.17 

.29 

.49

~78

Tio

1.58

0.01 

.02 

.02 

.04 

.05 

.07 

.07 

.07 

.08

0.01 

.01 

.01

.015

 

0.01 

.012 

.018 

.03 

.05

.06 

.08 

.08

18

0.07

0.08 

.12 

.16 

.21 

.26 

.34 

.41 

.50 

.60 

.74

 

 

0.02 

.04 

.04 

.05 

.045 

.07 

.075 

.09 

.11 

.13

.02

.02

.05

0.015 

.02 

.025 

.03 

.04 

.05 

.055 

.05 

.09 

.04

now known) occurs in the Kentucky collections of the 

Providence Limestone Member but was not seen in the 

same samples as the forms here discussed. The 

specimens illustrated on plate 13, figures 5 and 8, are 

considered juveniles. Figures 3 and 4 show the extremes 

of shape difference in the inner volutions, and figures 1 

and 2 show well-developed mature specimens that are 

more robust than most of the specimens illustrated by 

Dunbar and Henbest (1942).

Measurements for these specimens are provided in 

table 17 and on figure 21. The specimens are from locality 29 in Muhlenberg County, Ky.

Beedeina acme (Dunbar and Henbest), 1942

Plate 14, figures 1-7

Fusulina acme Dunbar and Henbest, 1942, p. 122, pi. 15, figs 1-18- pi

16, fig. 14.

Description.-The specimens from the Madisonville 

Limestone Member of the Sturgis Formation in Kentucky bear a striking resemblance to those described by 

Dunbar and Henbest (1942) from the Lonsdale 

Limestone Member of the Modesto Formation in Illinois. Measurements for the Kentucky specimens are 

given in table 18 and on figure 22. The specimens are 

elongate fusiform and expand relatively rapidly in
48

TABLE 9. -Measurements (in millimeters) for Wedekindellina euthysepta (Henbest), 1928 -Continued

19 15

Diameter of proloculus __ 0.06 0.06 

Radius vector 

1 0.07 0.06

2 _ .11 .09

3 _ .16 .12

4 .21 .18

5 .27 .24

6 .35 .32

7 .46 .44

8

9

10

Half length 

1 0.12

2 _ .28

3 .38

4 .51

5 _ .80

6 _ 1.20

7 _ 1.81

8

9

Volution height 

1 0.02 0.02

2 .04 .02

3 .05 .03

4 .05 .05

5 .06 .06

6 _ .08 .08

7 .10 .11

8

9

10

Wall thickness 

1 0.01

2 .02

3 .01 .015

4

5 .02 .03

6 .03

7

8

9

Tunnel width (axials) or 

septal spacing 

1 0.02 0.01

2 .02 .02

3 .03 .04

4 _ .05 .07

5 .05 .08

6 .05

7 .055

8

9

10

Measurements of fossils shown in plate 7, figures 19, 15, 17, 20-24

17

0.04

0.06 

.09 

.13

.18 

.24 

.32 

.44 

.59

 

 

0.02 

.03 

.04 

.055 

.06 

.08 

.12 

.15

"01

.012 

.018 

.02 

.025 

.05

0.018 

.02 

.03 

.028 

.04 

.07 

.07

 

diameter and length. The proloculus is commonly close 

to 0.16 mm in maximum outer diameter, but some specimens are almost 0.19 mm. The spirotheca is thin and 

composed primarily of protheca. An inner tectorium is 

present in parts of the shell, and an outer tectorium is 

present in parts of the inner volutions. Septa are 

numerous and closely spaced. They are strongly and 

regularly to irregularly fluted, forming chamberlets 

throughout the outer volutions. The tunnel is narrow 

and irregular, bordered by small chomata that are broad 

in the inner volutions and narrow in the outer volutions. 

Comparison and remarks. -Beedeina acme comes 

closer to the concept of Fusulina than any other

20 21 22 23 24

0.06 0.08 0.06 0.07 0.09

0.06 0.06 0.07 0.06 0.09 

.06 .09 .09 .07 .13 

.08 .12 .12 .13 .17 

.11 .16 .15 .16 .21 

.15 .21 .22 .21 .28 

.20 .28 .29 .27

.25 .36 .37 .32 __

.45 _ _ 

0.10 0.16 0.11 0.12 0.14 

.19 .35 .21 .25 .29 

.36 .50 .34 .40 .42 

.54 .64 .56 .51 .61 

.78 .84 .70 .65 .82 

1.04 1.32 .90 .89 ____ 

1.68 1.15 1.10 ____

0.02 0.02 0.015 0.01 0.02 

.02 .03 .015 .03 .04 

.02 .04 .03 .035 .04 

.04 .05 .06 .04 .05 

.05 .07 .06 .05 .06 

.05 .08 .08 .065 ____ 

.09 .05

0.02 __ ____

.02 .015

.01 .015 .02

.02 _ .02 .02 .018 

.03 .02

.04 .04

0.02 0.02 0.02

.03 .03 .02 .03

.04 .04 .04 .045 .04 

.06 .055 .05 .05 .04 

.08 .08 .09 .06 __

.09 .09 ____

fusulinid found in Kentucky. The thin spirotheca 

primarily composed of protheca, the elongate shape, the 

tightly fluted septa, and the small chomata all suggest a 

tendency toward Fusulina. The presence of chomata 

out to the last volution, the absence of axial filling, and 

the weak septal fluting in the inner volutions all show 

the greater affinity to the genus Beedeina. Most of the 

late Desmoinesian beedeinas are more fusiform and 

have more massive chomata than does B. acme. 

Distribution. - This species is the youngest Beedeina, 

found in Kentucky and among the youngest reported 

from Illinois. In Kentucky, it was found only in the 

Madisonville Limestone Member of the Sturgis Forma-
SYSTEMATIC DESCRIPTIONS 49

TABLE 9. -Measurements (in millimeters) for Wedekindellina euthysepta (Henbest), 1928- Continued

Character

Diameter of proloculus ___ 

Radius vector 

1

2

3

4

5

6

7

8

9

10

Half length

1

2

3

4

5

6

7

8

9

Volution height 

1

2

3

4

5

6

7

8

9

10

Wall thickness 

1

2

3

4

5

6

7

8

9

Tunnel width (axials) or 

septal spacing

1 __________________ 

2

3

4

5

6

7

8

9

10

Measurements of fossils shown in plate 7, figures 31, 27, 25, 33, 34, 28, 37, 29, 26

31

0.07

0.09 

.13 

.18 

.25 

.32

0.18 

.04 

.045 

.07 

.08

.01 

.01 

.015 

.03

0.025 

.04 

.04 

.045 

.05

27

0.10

0.06 

.10 

.14 

.19 

.26 

.33 

.45 

.56

0.11 

.28 

.43 

.58 

.79 

1.00 

1.32 

1.72

0.01 

.035 

.045 

.05 

.07 

.07 

.12 

.11

.03 

.035 

.07

.04 

.065 

.08 

.11 

.13

 

25

0.05

0.04 

.065 

.09 

.14 

.18 

.24 

.42 

.52

 

0.015 

.02 

.03 

.04 

.04 

.07 

.08 

.10

___

.005 

.01 

.01 

.015 

.01 

.02

0.03 

.03 

.05 

.05 

.05 

.06 

.09 

.13

___

33

0.05

0.06 

.07 

.12 

.19 

.25 

.32 

.45

___

____

0.02 

.02 

.03 

.06 

.06 

.07 

.12

.015 

.018 

.02 

.03 

.04

0.018 

.03 

.04 

.04 

.06 

.08 

.05

___

34

0.06

0.06 

.11 

.16 

.22 

.30 

.39 

.52

___

0.02 

.04 

.04 

.07 

.07 

.09 

.13

____

.015 

.02 

.02 

.025 

.04

.015 

.025 

.04 

.05 

.07 

.08

28

0.06

0.04 

.06 

.10 

.16 

.22

___

0.06 

.15 

.26 

.34 

.52

0.015 

.02 

.04 

.06 

.06

____

.01 

.018

 

0.02 

.03 

.04

.06

____

37

0.06

0.08 

.11 

.15 

.195 

.265 

.39

0.18 

.30 

.50 

.76 

1.13

0.02 

.03 

.04 

.045 

.07 

.10

 

E

0.025

.06 

.075 

.10

 

____

29

0.08

0.065 

.10 

.15 

.19 

.28

0.09 

.23 

.37 

.55 

.74

0.02 

.03 

.05 

.04 

.09

0.01 

.01 

.02

 

.028 

.03 

.04 

.05

____

____

26

0.095

0.07 

.15 

.20 

.28 

.36 

.44

____

0.12 

.22 

.37 

.59 

.875 

1.17 

1.44

0.02 

.03 

.045 

.045 

.08 

.08 

.09

____

____

0.04 

.045 

.06 

.08 

.105

 

tion at locality 26 in Hopkins County. It has been 

reported from the Lonsdale, Cutler (now included in the 

Piasa), and Piasa Limestone Members of Illinois.

Specimens studied.-The types from Illinois were 

studied in addition to 22 specimens from Kentucky.

Beedeina sp. aff. B. acme (Dunbar and Henbest), 1942

Plate 14, figures 8-12

Description. - One sample from beds mapped as Providence Limestone Member of the Sturgis Formation in 

Hopkins County, Ky., contains fusulinids more elongate 

than any found in the other samples of Providence. The

specimens bear some resemblance to Beedeina haworthi 

(Beede) but are more like B. acme (Dunbar and 

Henbest), 1942. Measurements for these specimens are 

given in table 18. The specimens are elongate fusiform 

but do not elongate as rapidly as do specimens of B. 

acme. The proloculus ranges from 0.15 to 0.21 mm in 

outside diameter. The spirotheca is thin and composed 

of protheca and variable amounts of epitheca, principally outer tectorium. The protheca in some specimens 

shows clearly the fine keriothecal or fibrous structure. 

The septa are tightly and regularly fluted. Septal pores 

are common and especially visible at the poles. The tunnel is narrow and irregular, bordered by low narrow
50 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS 

TABLE 9. -Measurements (in millimeters) for Wedekindellina euthysepta (Henbest), 1928 -Continued

Character

Diameter of proloculus

Radius vector 

1

2

3

4

5

6

7

8

9

Half length 

1

2

3

4

5

6

7

8 _

9

Volution height 

1

2

3

4

5 _ _

6

7

8

9 _ _

Wall thickness 

1

2

3

4

5 _

6

7

8

9 _

Tunnel width (axials) or 

septal spacing 

1

2 _ _

3

4

5

6

7

8

9 .

Measurements of fos

32

0.07

0.07

.095

.18

.23

.31

.40

0.16

.25

.38

.62

.875

1.24

0.02

.025

.025

.05

.055

.078

.085

.03

.03

.06

.075

34

0.10

0.06 

.09 

.13 

.17 

.24 

.31 

.39 

.50

0.15 

.26 

.485 

.70 

.96 

1.16 

1.41

0.02 

.02 

.04 

.05 

.06 

.07 

.08 

.11

.025

.03 

.04 

.055 

.07 

.10 

.11 

.14

35

0.05

0.06 

.09 

.12 

.16 

.20 

.28 

.36 

.46

0.018 

.025 

.03 

.035 

.04 

.08 

.08 

.09

0.01 

.012 

.02 

.02

0.018 

.018 

.018 

.038 

.04 

.05 

.06

ssils shown in plate 16, figures 32, 34, 35; plate 7, fig

30

0.02

0.04 

.06 

.09 

.13 

.17 

.22 

.29

0.01 

.02 

.025 

.03 

.04 

.07

.01 

.01

.015

.018 

.02 

.03 

.03

38

0.08

0.06 

.08 

.11 

.15 

.22 

.30 

.40

0.14 

.20 

.26 

.50 

.88 

1.12 

1.32

0.015 

.02 

.03 

.04 

.07 

.08 

.10

.025

.025 

.04 

.05 

.06 

.08 

.09

39

0.09

0.08 

.11 

.14 

.18 

.25 

.34 

.42

0.10 

.16 

.25 

.38

1.19

0.02 

.03 

.03 

.05 

.07 

.09 

.09

.015

.02 

.02

.02 

.03 

.06 

.08 

.105

32

0.06

0.06 

.09 

.13 

.18 

.25 

.32 

.41 

.52

0.02 

.03 

.04 

.05 

.07 

.08 

.085

.005 

.015 

.03

.01 

.028 

.04 

.045 

.05

ures 30, 38, 39. 32. 35. 36. 41. 42

35

0.05

0.05 

.08 

.12 

.17 

.21 

.29 

.39 

.47

0.018 

.03 

.03 

.055 

.05 

.06 

.07 

.10

.015

0.01 

.015 

.02 

.05 

.06 

.06

.09

36

0.07

0.07 

.11 

.145 

.20

0.01 

.03 

.04 

.05

 

.01

0.01 

.02 

.025 

.03

 

41

0.07

0.06 

.10 

.13 

.17 

.23 

.31 

.39

0.12 

.22 

.38 

.63 

.79 

1.05 

1.42

0.02 

.03 

.03 

.04 

.06 

.07 

.08

0.01

~~6i

.02

.02

0.02 

.03 

.05 

.06

.08

42

0.07

0.07 

.10 

.14 

.19 

.22 

.28 

.36 

.46 

.58

 

0.018 

.03 

.04 

.05 

.035 

.05 

.075 

.10 

.12

.01 

.02

0.018 

.02 

.02 

.03 

.03 

.05 

.07 

.06 

.09

chomata that are commonly obscured by the septal 

loops.

Distribution. - These forms were found at locality 28 

in Hopkins County, Ky. They are associated with rare 

specimens of a form more properly assigned to B. girtyi 

(Dunbar and Condra). 1927.

Beedeina carmani (Thompson), 1936

Plate 15, figs. 1-6

Fusulinella carmani Thompson, 1936, p. 678, 679, pi. 91, figs. 10-12 

Fusulina carmani (Thompson) Smyth, 1957, p. 267, 268, pi. 2, figs. 13, 

14, 20.

Description. - Measurements are presented in table 19

and on figure 23. The specimens are elongate fusiform 

after early volutions that are rhomboidal to fusiform. 

The shell expands rapidly and elongates along a straight 

to irregularly curved axis. The proloculus is moderately 

large, averaging more than 0.15 mm in maximum outer 

diameter, some specimens attaining 0.20 mm.

The spirotheca is thin and is composed of protheca and 

inner tectorium, with minor amounts of outer tectorium 

in limited areas. The septa are widely and fairly evenly 

spaced. They are weakly fluted in the inner volutions 

and more tightly fluted in the outer volutions, forming 

chamber-lets all the way to the tunnel area. The tunnel is 

wide and about half the volution height. It is bordered by
SYSTEMATIC DESCRIPTIONS 51

high narr 

lower chc 

Compa 

bles B. le 

with a la 

not as la 

Fusuline 

recognize 

the septa 

Distrib 

Vanport 

tion in 01 

specimen 

localities

0.4

six °'3

Z 9 0.2

(- ^ 

0.1

CO 

_l LU 0.10

^jj: 0.05

1 0-3

I

Z 0.2 

O

D 0.1 _j

O

3.5

3.0

I

5 2 - 5

z 

^ 2.0 

u.

< 1.5 

1.0 

0.5

Figure 14. 

All measi 

bute are

ow chomata in the inner volutions and smaller 

>mata at maturity. 

rison and remarks. -Beedeina carmani resemei in general stage of development but starts 

rger proloculus, is more loosely coiled, and is 

rge. The species was originally described as a 

lla because of its inner volutions but was later 

>d as an early form of Beedeina on the basis of 

1 fluting in the outer volutions. 

ution.-B. carmani was described from the 

Limestone Member of the Allegheny Formalio and has not been recognized elsewhere. The 

s described and illustrated here are from 

71 and 75 in Muskingum County, Ohio.

1 1 1 1

 4

 £ 

^^9 

_ . - «fc-

. ^*/.*

*r i i i

1 1 1 1

 -

^ 

1 1 i 1

0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 0.9 1.0 

RADIUS VECTOR

-Graphs for Wedekindellina euthysepta (Henbest), 1928. 

irements are in millimeters, and the values for each attriplotted against the radius vector.

TABLE 10. -Measurements (in millimeters) for Wedekindellina sp.

[Leaders ( __ ), data not available or character absent]

Measurements of fossils shown in plate 17, 

Character 

789

figures 7-9

Diameter of 

proloculus _ 0.07 0.10 0.10

Radius vector 

1 _ 0.06 0.07 0.07

2 .09 .10 .095

3 .12 .14 .14

4 _ _ .16 .19 .19

5 _ _ .21 .29

Half length 

1 _ 0.07 0.15

2 .18 .32

3 .31 .49

4 .45 .68

5 .60

Volution height 

1 0.02 0.02 0.02

2 _ .025 .03 .02

3 .04 .04 .04

4 .04 .04 .05

5 .05 .10

Wall thickness 

1

2 0.01

3 .025

4 .025

5 .022

Tunnel width (axials) 

or septal spacing 

1 0.02 0.03

2 .025 .025 .04

3 .03

4 .055 .04

Beedeina sp.

Plate 17, figures 5, 6, 16-23

Description. - Measurements for these specimens are 

given in table 20. The specimens are small and inflated 

fusiform. They grow only to about four volutions, starting from a proloculus that has a maximum outer 

diameter near 0.10 mm. The innermost volutions are 

globular; later chambers become more elongate. The 

spirotheca is composed of protheca and variable 

amounts of inner and outer tectoria. No porous structure was seen in the diaphanotheca. The septa are weakly fluted. The tunnel is nearly straight, bordered by low 

narrow chomata that increase in height in the third volution. 

Comparison and remarks. -These specimens may 

represent the juveniles of Beedeina leei (Skinner), a 

species that is found with them in one of the samples. In 

the other sample, they occur only with juvenaria of 

Wedekindellina sp. The nearly spherical chambers in the 

specimens represented as figures 5, 16, 17, 21, and 22 on 

plate 17 are not typical of the juvenaria of the associated 

Beedeina adults. 

Distribution. - These specimens are from localities 86 

and 87 in Mercer and Armstrong Counties, Pa. The 

rocks are assigned to the Vanport Limestone Member of 

the Allegheny Formation.
52 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS 

TABLE 11.-Measurements (in millimeters) for Beedeina sp. off. B. novamexicana (Needham), 1937

[Leaders (__), data not available or character absent]

Character

Diameter of proloculus

Radius vector 

1

2

3

4

5

6

7

8

Half length 

1

2

3

4

5

6

7

8

Volution height 

1

2

3

4

5

6

7

Wall thickness 

1

2

3

4

5

6

7

Tunnel width (axials) or 

septal spacing 

1

2

3

4

5

6

7

8

Measurements of fossils shown in plate 8, figures 1, 2, 5, 6, 3; plate 9, figures 3-10

1

0.10

0.12

.18

.30

.44

.61

.84

0.16

.37

.56

.87

1.15

1.45

0.03

.07

.08

.12

.15

.20

.02

0.04

.07

.06

.11

.20

.24

2

0.20

0.18 

.30 

.45 

.63 

.81

0.18 

.35 

.64 

.90 

1.21

0.05 

.12 

.14 

.17 

.18

.015 

.02

0.045 

.07 

.10 

.14

5

0.155

0.16 

.23 

.37 

.48 

.65 

.82

0.20 

.46 

.62 

.79 

.99 

1.16

0.03 

.04 

.06 

.08 

.11 

.13

0.02 

.025 

.03

.04

0.04 

.04 

.06 

.08 

.11 

.13

6

0.15

0.10 

.19 

.31 

.46 

.59

0.24 

.37 

.57 

.85 

1.16

0.03 

.08 

.12 

.13 

.16 

.16

0.02

.03

 

0.03 

.05 

.07 

.10 

.14

3

0.11

0.11 

.19 

.32 

.45 

.59 

.76

0.03 

.07 

.13 

.13 

.14

.015 

.02 

..03 

.07 

.09

0.025 

.03 

.04 

.06 

.06

3

0.20

0.17 

.25 

.35 

.51 

.68 

.97 

1.15

0.22 

.39 

.75 

1.28 

1.80 

2.33 

2.93

0.07 

.08 

.10 

.16 

.17 

.26 

.21

.02

.04

.07

0.04 

0.07 

.08 

.10 

.19 

.26 

.28

4

0.14

0.17 

.27 

.39 

.54 

.76 

.98 

1.20

 

0.06 

.095 

.12 

.16 

.21 

.22 

.22

0.01

.04 

.045

.10

0.03 

.03 

.05 

.07

5

0.10

0.09 

.15 

.23 

.31

.45 

.70

0.15 

.27 

.45 

.68 

1.00 

1.43

0.03 

.065 

.07 

.08 

.13 

.25

.03 

.05 

.06 

.085 

.145

6

0.15

0.16 

.27 

.40 

.54 

.78 

1.00

0.22 

.48 

.66 

.855 

1.21 

1.63

0.05 

.11 

.13 

.14 

.22 

.25

.03

.06

0.03 

.05 

.07 

.12 

.15 

.21

7

0.19

0.17 

.26 

.40 

.595 

.815 

1.07

0.21 

.36 

.58 

.95 

1.39 

2.08

0.06 

.095 

.13 

.20 

.22 

.25

0.01

.04

0.03 

.05 

.07 

.11 

.18 

.25

8

0.17

0.17 

.27 

.42 

.585 

.78

0.06 

.10 

.16 

.16 

.19

0.01 

.02 

.035 

.04 

.06

0.03 

.04 

.05 

.07

9

0.15

0.11 

.18 

.28 

.39 

.54 

.73 

.96

 

0.02 

.07 

.09 

.10 

.15 

.19 

.23

0.005 

.01 

.015 

.02 

.045 

.07 

.08

0.04 

.05 

.06

-M

10

0.10

0.09 

.15 

.23 

.31 

.45 

.70

0.17 

.26 

.58 

.87 

1.24 

1.82

0.04 

.05 

.10 

.13 

.19 

.20

.02 

.025

~~04

0.03 

.04 

.05 

.08 

.11 

.16

Triticites skinneri Thompson, 1936

Plate 16, figures 1-10

Triticites skinneri Thompson, 1936, p. 682, pi. 91, figs. 4-7; Smyth, 

1957, p. 274, pi. 3, figs. 21, 22.

Diagnosis. - Shell small, inflated fusiform, with convex lateral slopes. Proloculus small, averaging 0.05 mm 

in maximum outer diameter. Tightly coiled, with seven 

to eight volutions, attaining length of 4-5 mm and width 

of as much as 2 mm.

Description. -Measurements are presented in table 21 

and on figure 24. This species has a small proloculus that 

ranges in maximum outer diameter from about 0.03 to 

0.07 mm, averaging 0.05 mm. The first volution in some 

specimens is nearly lenticular and at a large angle to the 

rest of the shell. The inner volutions are tightly coiled 

and elongate rapidly, but the outer volutions are more 

loosely coiled and more inflated in the equatorial area.

The lateral slopes are nearly straight in the inner volutions but tend to be convex in the outer volutions.

The spirotheca is composed of protheca with distinct 

keriothecal structure in the outer volutions. The septa 

are strongly but irregularly fluted, forming chamberlets 

across the entire shell in the outer volutions. The septa 

are closely but irregularly spaced, 35-38 in the outer 

volutions. The tunnel is nearly straight and quite narrow. It ranges in height from less to more than half the 

volution height. It is bordered by low broad chomata in 

the inner volutions and higher but narrower chomata in 

the outer volutions.

Comparison and remarks. - Triticites skinneri resembles T. moorei Dunbar and Condra (1927) in size but is 

more tightly fluted, has a smaller proloculus, is more 

tightly coiled, and has a narrower tunnel angle. Smyth 

(1957, p. 276) compares T. skinneri with T. cullomensis
SYSTEMATIC DESCRIPTIONS

TABLE 11.-Measurements (in millimeters) for Beedeinasp. off. B. novamexicana (Needham), 1937 -Continued

53

Diameter of proloculus ______ 

Radius vector 

1

2

3

4

5

6

7

8

Half length

1

2

3

4

5

6

7

8

Volution height 

1

2

3

4

5

6

7

Wall thickness

1

2

3

4

5

6

7

8

Tunnel width (axials) or 

septal spacing 

1

2

3

4

5

6

7

8

Measurements of fossils shown in plate 10, figures ]

1

0.16

0.07 

.19 

.32 

.43 

.62 

.83 

1.08

0.24 

.43 

.57 

.80 

1.18 

1.78 

2.13

0.04 

.08 

.12 

.12 

.18 

.21 

.24

0.02 

.03 

.03 

.03 

.04 

.05 

.04

0.03 

.03 

.09 

.14 

.14 

.23

3

0.19

0.10 

.18 

.27 

.42 

.56 

.73 

.90

0.14 

.29 

.48 

.76 

1.06 

1.3L 

1.58

0.02 

.07 

.09 

.15 

.15 

.17 

.16

0.02 

.03 

.03 

.02 

.04 

.03 

.02

0.02 

.04 

.06 

.11 

.12 

.12 

.20

2

0.16

0.12 

.22 

.34 

.52 

.70 

.92 

1.13

0.16 

.29 

.56 

.77 

1.06 

1.55 

1.86

0.04 

.09 

.12 

.17 

.19 

.21 

.21

.02 

.03 

.02 

.03 

.02 

.03

0.04 

.07 

.10 

.15

.26

4

0.19

0.10 

.16 

.25 

.41 

.59 

.78

0.14 

.30 

.46 

.78 

1.03 

1.24

0.03 

.06 

.07 

.16 

.18 

.19

0.02 

.02 

.02 

.015

0.03 

.04 

.06 

.09

.20

5

0.21

0.11 

.21 

.33 

.51 

-.6.7 

.89

0.30 

.50 

.67 

.98 

1.40 

2.02

0.02 

.10 

.11 

.17 

.16 

.22

0.02 

.02 

.04 

.04 

.03 

.04

0.03 

.05 

.06 

.10 

.13

6

0.19

0.16 

.24 

.38 

.58 

.80 

1.04

0.28 

.42 

.62 

.96 

1.25 

1.62

0.07 

.08 

.14 

.19 

.22 

.24

0.01 

.01 

.02 

.02 

.03 

.03

0.06 

.07 

.10 

.12 

.16 

.18

[, 3, 2, 4-6, 10, !

10

0.18

0.011 

.18 

.27 

.39 

.58 

.75 

.95

0.11 

.24 

.44 

.64 

.99 

1.21 

1.57

0.03 

.06 

.10 

.12 

.18 

.18 

.20

0.02 

.02 

.03 

.02 

.03 

.04 

.04

0.02 

.03 

.06 

.08 

.10 

.14 

.18

8, 11, 7

8

0.12

0.13 

.20 

.29 

.45 

.61 

.82 

1.02

0.09 

.20 

.33 

.54 

.85 

1.11 

1.60

0.02 

.05 

.08 

.10 

.16 

.18 

.22

0.005 

.005 

.01 

.01 

.02 

.03 

.03

0.03 

.03 

.06 

.07 

.12

11

0.15

0.12 

.19 

.27 

.39 

.55 

.75 

.96

0.24 

.43 

.57 

.96 

1.16 

1.81 

2.21

0.02 

.04 

.07 

.08 

.12 

.16 

.20

0.015 

.02 

.03 

.03 

.03 

.03

0.02 

.03 

.05 

.09 

.13

7

0.11

0.12 

.20 

.37 

.51 

.66 

.90

0.07 

.08 

.16 

.15 

.15 

.23

0.10 

.02 

.02 

.04 

.04

0.02 

.04 

.04 

.05 

.06 

.09

Dunbar and Condra (1927) but noted the smaller proloculus and narrower tunnel in T. skinneri. Except for 

the smaller proloculus, some juvenaria of T, ricei could 

be confused with those of T. skinneri, but the nature of 

the tunnel and the septal fluting distinguish them easily 

beyond the juvenarium.

Distribution.-T. skinneri has only been reported 

from the Ames Limestone Member of the Conemaugh 

Formation in Ohio.

Specimens studied.-Thompson's original material, including the types, was studied, in addition to topotypes 

from locality 69 in Perry County. Ohio.

Designation of types.-The specimen illustrated by 

Thompson (1936, pi. 91, fig. 6) is hereby designated the 

lectotype. It is specimen 1046A in the collections of the 

State University of Iowa. The specimens here illustrated 

are topotypes.

Triticites ricei Douglass n. sp.

Plate 16, figures 11-18

Name derivation. - Named for Charles L. Rice, who collected the 

sample containing this species and also many other samples for 

this study.

Diagnosis. - Shell elongate fusiform to elliptical, commonly one side flat or concave and the other convex, 

proloculus small, commonly less than 0.10 mm; shell expands slowly but uniformly, attaining nine volutions in 

some specimens.

Description. - Measurements are given in table 22 and 

on figure 25. The proloculus averages 0.09 mm in maximum outer diameter, ranging from 0.05 to 0.12 mm in 

14 specimens. The shell is tightly coiled and tends to 

establish and maintain a uniform shape, each volution 

closely paralleling the others. The lateral slopes are 

generally convex, but many specimens have one flat-
54 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS

u.o 

0.4

I

Q 0.3

0.2

0.1

C/D 

tf\ 

LU 0.10

^

y 0.05

tt1 0.3

I

| °'2

3 o.i

o

3.5

3.0

£ 2.5

d

^ 2.0

LL

< 1.5

I

1.0

0.5

_

. ,' . *" V'

9

. . -. .' % V

-

 <* ? *

s^*?" '

-

_

9 i

j ..' ; 

.. 4:%*" * * i i i

. . '. "." 

» * 

 :

 .**r*% * 

** . 

, 

w

-. "" '

>**' * *

i i i i

* *

 ..

.

*

i i i

-

-

-

-

_

-

_

-

i i i i

0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 0.9 1.0 1.1 1.2 

RADIUS VECTOR

1.3 1.4 1.5 1.6 1.7 1.8 1.9 2.0

Figure 15. - Graphs for Beedeina sp. aff. B. novamexicana (Needham), 1937. All measurements are in millimeters, and the values for each

attribute are plotted against the radius vector.

tened or even concave side. The spirotheca is composed 

of protheca with fine keriothecal structure. It is quite 

thin in the inner volutions and thickens gradually 

through the outer volutions. The septa are weakly fluted 

throughout most of the shell. Septa are numerous but irregularly spaced, some chambers being more than twice 

as wide as others. About 22 chambers are common in the 

seventh volution and 30 in the eighth. The tunnel is 

straight or wanders in a narrow equatorial zone; it extends less than half the chamber height and is limited by 

narrow chomata. 

Comparison and remarks.-Triticites ricei differs

from described species of Triticites in its tight uniform 

coiling, chambers maintaining nearly uniform height 

throughout much of their length. Some specimens of T. 

cullomensis bear a superficial resemblance to T. ricei, 

but they expand more rapidly, start with a larger proloculus, and have a thicker spirotheca.

Distribution. - T. ricei is known from the Ames Limestone Member of the Conemaugh Formation at locality 

66 in Gallia County, Ohio.

Specimens studied.-Thirty specimens of T. ricei from 

collection f!4005 at locality 66 were studied. Topotypes 

of T. cullomensis were sectioned for comparison.
SYSTEMATIC DESCRIPTIONS 55

TABLE 12.-Measurements (in millimeters) for Beedeina leei (Skinner), 1931, and B. sp. aff. B. leei (Skinner), 1931

[Leaders (__), data not available or character absent]

Character

Diameter of proloculus ___ 

Radius vector 

1

2

3

4

5

6

7

8

Half length

1

2

3

4

5

6

7

8

Volution height 

1

2

3

4

5

6

7

Wall thickness 

1

2

3

4

5

6

Tunnel width (axials) or 

septal spacing

1

2

3

4

5

6

7

8

Measurements of fossils shown in plate 8, figures 4, £

4

0.15

0.18 

.28 

.36 

.56 

.75

0.07 

.10 

.10 

.19 

.19

0.02 

.02 

.02 

.03 

.04

0.04 

.045 

.05 

.05 

.06

8

0.14

0.12 

.19 

.30 

.44 

.61 

.76

0.13 

.46 

.79 

1.10 

1.73 

2.23

0.03 

.06 

.10 

.17 

.17 

.20

~~02 

.06 

.08

0.04 

.05 

.10 

.14 

.22

7

0.16

0.13 

.20 

.31 

.48 

.62 

.82

0.18 

.38 

.74 

1.05 

1.44 

1.79

0.05 

.08 

.10 

.16 

.16 

.20

 

.045 

.05

0.04 

.05 

.09 

.14 

.23

9

0.16

0.11 

.19 

.27 

.41 

.59 

.85

0.13 

.26 

.44 

.85 

1.42 

1.88

0.04 

.07 

.08 

.17 

.17 

.27

.025

0.03 

.05 

.07 

.16 

.16 

.22

10

0.10

0.10 

.14 

.22 

.33 

.63 

.72

0.18 

.35 

.52 

.71 

.97 

1.22

0.02 

.04 

.07 

.11 

.21 

.22

.02 

.03

0.03 

0.04 

.06 

.08

 

!, 7, 9, 10, 13, 17,

13

0.04

0.05 

.09 

.14 

.22 

.32 

.45 

.62

0.09 

.16 

.24 

.45 

.62 

.84 

1.20

0.01 

.03 

.05 

.08 

.10 

.13 

.17

"6l

 

.03 

.04 

.05 

.08 

.10

15; plate 9, Figure 2

17

0.13

0.12 

.195 

.29 

.42 

.60 

.85

 

0.03 

.07 

.09 

.13 

.18 

.245

0.02 

.02 

.025 

.05 

.05 

.10

0.03 

.04 

.05 

.07 

.08 

.08

15

0.15

0.15 

.23 

.32 

.45 

.63 

.80

0.24 

.38 

.61 

.985 

1.45 

1.80

0.03 

.07 

.09 

.13 

.17 

.18

0.01 

.02

.04

0.03 

.05 

.06 

.10 

.10

2

0.10

0.10 

.15 

.20 

.27 

.39 

.51 

.66 

.82

0.13 

.25 

.48 

.70 

1.00 

1.26 

1.45 

1.72

0.04 

.05 

.07 

.12 

.12 

.15 

.16

.01

.02 

.02

0.02 

.03 

.05 

.06 

.07 

.09 

.16 

.17

Designation of types.-The specimen illustrated on 

plate 16 as figure 11 is designated the holotype. The 

other specimens are paratypes.

Triticites ohioensis Thompson, 1936

Plate 16, figures 19-24; plate 18, figures 1-45

Fusulina secalica Condit, 1912, p. 44, 88

Triticites ohioensis Thompson, 1936, p. 680-682, pi. 91, figs. 1-3; 

Dunbar and Henbest, 1942, p. 130-132, pi. 19, figs. 1-22, pi. 20, 

figs. 20-24; Thompson, 1948, pi. 8, fig. 5; Smyth, 1957, p. 

272-273, pi. 3, figs. 25-28; Thompson, 1957, p. 313-315, pi. 27, 

figs. 1-8; Ross, 1963, p. 108-109, pi. 7, figs. 7, 9-14.

Diagnosis.-Shell elongate fusiform with broadly 

rounded poles, attaining lengths of about 7.7 mm and 

widths of about 2 mm in 6V2-7 volutions. Coiling tight 

and regular, but axis curved to irregular. Proloculus 

minute. Septa irregularly fluted and closely spaced.

Description. - Measurements are given in table 23 and 

on figure 26. The shape in the inner volutions is fusiform

with sharply pointed poles, but the specimens elongate 

rapidly, and the poles are broadly rounded in the outer 

volutions. The volution height increases regularly 

throughout the shell, and the length increases regularly 

and rapidly, resulting in form ratios of 3-4 in most 

specimens and 4-4Y£ in some mature specimens. Commonly, the last volution or so has a lower form ratio.

The proloculus ranges from 0.05 to 0.15 mm in outer 

diameter, with a mean of 0.09 mm, and the shape is 

spherical. The thin wall is composed of a tectum and indistinct keriotheca, especially in the inner volutions. The 

maximum thickness approaches 0.06 mm in the mature 

specimens. The thickness is maintained throughout 

most of the length of the shell, thinning only at the 

poles.

The septa are closely and regularly spaced, except in 

the outer volutions of some specimens. The number of 

septa increases regularly from 9 in the first volution to 

about 25 in the last volution, but as many as 31 were
56

TABLE 12. -Measurements (in millimeters) for Beedeina leei (Skinner), 1931, and B. sp. aff. B. leei (Skinner), 1931 -Continued

Character

Diameter of proloculus _ 

Radius vector 

1 _

2 _

3 _

4

5

6

7

8

Half length 

1

2 _

3

4 _

5 _

6 _

7 _

Volution height 

1

2

3

4

5 _

6 _

7

8

Wall thickness 

1

2

3

4

5 _

6

7

Tunnel width (axials) or 

septal spacing 

1

2

3 _

4

5 _

6 _

7 _

Measurements of fossils shown in plate 10, figures 9, 12,

9

0.07

0.08 

.13 

.19 

.29 

.39 

.57 

.79

0.03 

.05 

.06 

.09 

.10 

.18 

.21

.02 

.02 

.02 

.02 

.04

0.02 

.03 

.03 

.03 

.05 

.05

12

0.05

0.09 

.14 

.21 

.31 

.42 

.54 

.70 

.86

 -

0.03 

.04 

.06 

.10 

.10 

.13 

.16 

.16

.01 

.02 

.02 

.03 

.03

0.02 

.02 

.03 

.03 

.04 

.04

15

0.08

0.06 

.12 

.17 

.25 

.35 

.48 

.63

 

0.02 

.06 

.05 

.08 

.10 

.12 

.15

.015 

.02 

.02 

.025 

.07

0.02 

.02

.03 

.03 

.04 

.05

13

0.17

0.13 

.20 

.30 

.41 

.56

0.29 

.435 

.72 

1.00 

1.34

0.05 

.07 

.095 

.11 

.14

.02 

.03

.05

0.03 

.05 

.08 

.10

14

0.10

0.07 

.11 

.15 

.21 

.31 

.45 

.60

0.09 

.19 

.39 

.64 

1.13 

1.65 

2.35

0.02 

.04 

.04 

.06 

.09 

.14 

.15

0.02 

.03 

.03 

.04

.04

0.03 

.05 

.08 

.11 

.19

, 15, 13, 14, 16, 17; plate 12, figures 8, 10

16

0.11

0.12 

.18 

.27 

.39 

.53 

.68 

.94

0.14 

.23 

.34 

.53 

.79 

1.27 

1.69

0.04 

.06 

.08 

.13 

.14 

.16 

.26

.02 

.03 

.02

0.04 

.06 

.08 

.13 

.14 

.16 

.26

17

0.17

0.15 

.22 

.30 

.39 

.49 

.62 

.78

0.28 

.42 

.77 

1.00 

1.21 

1.44 

2.10

0.07 

.08 

.08 

.09 

.10 

.13 

.15

0.01 

.02 

.02

0.04 

.05 

.08 

.11 

.14 

.19 

.30

8

0.09

0.11 

.18 

.23 

.33

.42 

.59 

.77

0.10 

.22 

.33 

.63 

.89 

1.15 

1.70

0.02 

.02 

.05 

.07 

.09 

.17 

.17

0.02 

.02 

.03

.04

0.02 

.04 

.05 

.08 

.13 

.18

10

0.18

0.09 

.13 

.20 

.26 

.39 

.52

0.10 

.20 

.37 

.63 

1.09 

1.75

0.03 

.04 

.06 

.06 

.12 

.12 

.13

0.02 

.02 

.03 

.02

0.02 

.03 

.04 

.07

observed in the last volution of one specimen. Septal 

pores are small but abundant and commonly visible not 

only in the poles but also across the middle of the shell in 

the outer volutions. Fluting is weak and irregular across 

the middle of the shell; the presence of chamberlets is 

rare, except in the polar areas.

The tunnel widens rapidly but irregularly, attaining 

widths near 1 mm in the outer volutions. The chomata 

are low and narrow but often appear to be high, broad, 

or even overhanging when the plane of a septum is included in the section. In tangential views, the extension 

of secondary deposits along the septa can be seen in 

some specimens. No axial filling or other epithecal 

deposits are present.

Comparison and remarks. - Plate 18 shows the full 

range of variation seen in this species from its early occurrences in the lower bed of the Brush Creek 

Limestone Member, through the Cambridge Limestone 

Member of the Conemaugh Formation in eastern Kentucky. The smallest forms at the bottom of the plate are 

from the oldest beds, and the changes through the units 

are illustrated by fusulinids from the younger units 

toward the top. The specimens from the upper beds are 

similar to the type and others from the Cambridge 

Limestone Member in Ohio, as illustrated on plate 16, 

figures 19-24.

At first glance, the specimens from the lower Brush 

Creek Limestone Member look quite different from 

those of the Cambridge Limestone Member, but the difference is principally the addition of volutions in the 

younger specimens. Plots of all the commonly measured 

attributes indicate a complete paralleling of growth 

through the fifth or sixth volution, where growth stops 

in the older specimens. The younger forms added one or 

more volutions, attaining 7V2-8 volutions.

Distribution. -This species is common in rocks of early 

Missourian age. Its typical specimens are from the Cambridge Limestone Member of the Conemaugh Formation.
SYSTEMATIC DESCRIPTIONS 57 

TABLE 12. -Measurements (in millimeters) for Beedeina leei (Skinner), 1931, and B. sp. aff. B. leei (Skinner), 1931 - Continued

Character Measurements of fossils shown in plate 15, figures 7-13

7 8 9 10 11 12 13

Diameter of proloculus 0.10 0.11 0.14

Radius vector 

1 _ 0.12 0.11 0.11

2 .16 .15 .19

3 .23 .20 .27

4 .35 .30 .39

5 .48 .41 .60

6 .68 .55 .79

7 .88 .73

8

Half length 

1 0.19 0.16 0.25

0.10 0.10 0.15 0.12

0.13 0.12 0.11 0.12 

.17 .19 .175 .18 

.24 .28 .26 .27 

.35 .39 .38 .40 

.48 .55 .55 .56 

.63 .72 .75 .81 

.82

2 .40 .33 .445

3 .66 .50 .725

4 1.01 .695 1.02

5 1.52 .73 1.54

6 1.98 1.38 1.93

7 2.40 1.845

Volution height 

1 0.045 0.04 0.04

2 .05 .045 .07

3 .06 .05 .08

4 .12 .095 .11

5 .13 .115 .21

6 .20 .14 .20

7 .20 .18

8

Wall thickness 

1

2

3

4 .03 .025

5 .05 .07

6 .06

0.04 0.03 0.02 0.04 

.04 .07 .06 .06 

.06 .09 .08 .10 

.10 .11 .12 .12 

.13 .15 .165 .17 

.15 .18 .19 .15 

.19

0.02 0.015 0.015 0.018 

.02 .015 .03 .025 

.03 .04 .05 .045 

.035 __ .08 .06 

.07 .08

7

Tunnel width (axials) or 

septal spacing 

1 0.05

2 .05 .06

3 .07 .06 .095

4 .11 .14 .14

5 .20 21 20

0.02 0.025 0.03 0.03 

.04 .03 .04 .04 

.05 .04 .04 .04 

.03 .05 .05 .04 

03 .035 .02

6 .20 .25 .02 .03

Specimens studied. -Several hundred thin sections 

from 15 samples were studied to recognize the lateral 

and vertical variation in this species. 

Designation of types. -The specimen illustrated by 

Thompson in 1936 (his pi. 91, fig. 1) was reillustrated as 

a syntype by Thompson in 1948 (his pi. 8, fig. 5). In 1957 

(p.314), Thompson referred to "the holotype of T. 

ohioensis but did not indicate the specimen he considered to be the type. I hereby designate the above 

specimen as the lectotype of T. ohioensis. It is here 

reillustrated on plate 16 as figure 19. It is specimen 

S.U.1. 1044B in the State University of Iowa collections. 

The specimen here illustrated on plate 16 as figure 20 is 

a paralectotype S.U.I. 1044 A.

Triticites smythi Douglass n. sp.

- Plate 16, figures 25-28

Name derivation. - Named for Pauline Smyth, who collected the 

sample and who has studied the fusulinids of Ohio.

Triticites venustus (part) Dunbar and Henbest, 1942, pi. 20, figs. 12, 

13, 18, 19.

Diagnosis. -Elliptical to fusiform, relatively loosely 

coiled form, attaining length of 2.5-3 mm and width of 

about 1 mm in six volutions. Proloculus about 0.12 mm 

in maximum outer diameter. Tunnel narrow. 

Description. -Measurements are presented in table 24 

and on figure 27. The shell of this form is small and 

fusiform to elliptical, with convex lateral slopes and 

rounded poles. The shape is relatively constant 

throughout growth. The small proloculus is about 0.12 

mm in maximum outer diameter. The spirotheca is thin 

and composed of protheca with very fine keriothecal 

structure. A suggestion of outer tectorium is found 

along parts of the lateral slopes. The septa are weakly 

and irregularly fluted. They are closely but irregularly 

spaced, forming about 20 chambers in the fifth volution. 

The tunnel is narrow and straight, bordered by low 

broad chomata. It extends more than half the height of 

the chambers in the outer volutions.
58 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS 

TABLE 12.-Measurements (in millimeters) for Beedeina leei (Skinner), 1931, and B. sp. aff. B. leei (Skinner), 1931)-Continued

Character

Diameter of proloculus __ 

Radius vector 

1

2

3

4

5

6

7

8

Half length 

1

2

3

4

5

6

7

Volution height 

1

2

3

4

5

6

7

8

Wall thickness 

1

2

3

4

5

6

7

Tunnel width (axials) or 

septal spacing 

1

2

3

4

5

6

7

Measurements of fossils shown in plate 17, figures 10-15, 24, 25

10

0.16

0.15 

.23 

.33 

.44 

.58 

.77 

.92

0.22 

.41 

.65 

1.04 

1.34 

1.82 

2.19

0.06 

.07 

.10 

.12 

.14 

.14 

.20

.05 

.07

0.035 

0.08 

.11 

.16 

.22 

.25

11

0.15

0.13 

.21 

.32 

.47 

.63 

.89

0.21 

.42 

.74 

1.12 

1.91 

2.04

0.04 

.08 

.10 

.15 

.16 

.25

0.018 

.02

.04

.11

0.04 

.07 

.08 

.10 

.16 

.24

12

0.11

0.10 

.15 

.23 

.34 

.49 

.67

0.20 

.41 

.75 

1.13 

1.42 

1.78

0.03 

.05 

.08 

.10 

.15 

.20

 

 

0.035 

.04 

.09 

.11 

.17 

.28

13

0.12

0.11 

.17 

.275 

.43 

.60

 

0.035 

.06 

.11 

.16 

.17

0.01 

.015 

.015 

.03 

.04

0.03 

.035 

.04 

.06 

.05

 

14

0.08

0.09 

.12 

.18 

.24 

.35 

.47 

.60

 

0.02 

.04 

.06 

.11 

.12 

.14

 

.015

~~03 

.03

0.03 

.03 

.035 

.05 

.07 

.06 

.03

15

0.11

0.11 

.18 

.26 

.38 

.55 

.74

 

 

0.04 

.06 

.08 

.12 

.17 

.20

 

.015 

.015 

.02 

.04 

.065

0.02 

.025 

.04 

.06

 

24

0.12

0.12 

.21 

.30 

.39 

.54

0.31 

.63 

.82 

1.17 

1.56

0.02 

.085 

.09 

.098 

.145

 

.045

0.04 

.07 

.10 

.12

 

25

0.09

0.11 

.17 

.24 

.33

.47

0.11 

.32 

.57 

.92 

1.23

0.04 

.06 

.06 

.09 

.135

.02 

.03 

.07

0.05 

.05 

.08 

.10 

.17

 

Comparison and remarks. - Triticites smythi is small 

and elliptical to fusiform with a narrow tunnel. It is easily distinguished from T. ohioensis, with which it occurs 

in Ohio, and from T. venustus in Illinois, both of which 

are more elongate and larger forms having a wider tunnel and more closely spaced septa. Three specimens illustrated as T. venustus by Dunbar and Henbest (1942, 

pi. 20, figs. 12, 13, 18, 19) resemble T. smythi quite 

closely; those three specimens are from the Livingston 

(?) Limestone, according to the authors. The other illustrated and described specimens assigned by Dunbar 

and Henbest to T. vinustus are from other localities.

Distribution. - T. smythi is recognized from the Cambridge Limestone Member of the Conemaugh Formation in Perry County, Ohio, at locality 69. The sample 

was collected by Pauline Smyth at her locality 33, where 

it is associated with abundant T. ohioensis. The 

specimens illustrated by Dunbar and Henbest, as mentioned above, are from their locality 490 from Christian

County, 111. where the specimens are also associated 

with T. ohioensis.

Designation of types. - The specimen illustrated on 

plate 16 as figure 25 is designated the holotype; the 

other illustrated specimens are paratypes.

Triticites cullomensis Dunbar and Condra, 1927

Plate 17, figures 1-4

Triticites cullomensis Dunbar and Condra, 1927, p. 93-95, pi. 5, figs. 

5-10; Dunbar and Henbest, 1942, p. 135, pi. 23, figs. 13-18; 

Thompson, Verville and Bissell, 1950, p. 457-460, pi. 62, figs. 

7-17, pi. 63, figs. 1-3; non Smyth, 1957, p. 273-274, pi. 3, figs. 

23, 24; Slade, 1961, p. 70, pi. 10, figs. 5, 6; Sabins and Ross, 1963, 

p. 339, pi. 35, figs. 13-16.

Discussion. -The two specimens illustrated by Dunbar 

and Henbest (1942, pi. 23, figs. 17, 18) from Brilliant 

Cut, Pittsburgh, Pa. (locality 88), and two additional 

specimens from the same locality are illustrated here.
SYSTEMATIC DESCRIPTIONS 59

The exact horizon from which the specimens were obtained is not known. The original collection was made by 

Dr. William Darrah. With the help of H. B. Rollins of the 

University of Pittsburgh and colleagues Rucha Ingavat 

and C. H. Cheong, I was able to find some float material 

at Brilliant Cut containing Triticites. The source bed 

was not located.

Pauline Smyth (1957) reported T. cnllomensis from 

the Ames Limestone Member of the Conemaugh Formation in Gallia County, Ohio. The two specimens that she 

illustrated have a small proloculus and are tightly coiled; 

these features suggest that the specimens are probably 

more closely related to T. skinneri than to T. cullomensis.

0.4

0.3

I1 *-

z 9 °- 2

0.1

CO

uj 0.10

0.05

1 0-3

X

0.2 

0.1

3.5 

3.0 

2.5 

2.0

1.0 

0.5

 

 t'

 . - t '-* 

0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 0.9 

RADIUS VECTOR

1.0

Figure 16.-Graphs for Beedeina leei (Skinner), 1931. All measurements are in millimeters, and the values for each attribute are plotted against the radius vector.

Triticites kehni Douglass n. sp.

Plate 19, figures 1-11

Name derivation.-Named for Thomas M. Kehn, who provided many 

samples and stratigraphic data from western Kentucky for this 

study.

Diagnosis. - Shell of moderate size, elongate fusiform 

to subcylindrical with nearly straight to slightly convex 

lateral slopes and rounded poles. Proloculus fairly large, 

0.09-0.16 mm in maximum outer diameter. Spirotheca 

with fairly fine keriotheca. Septa irregularly spaced and 

irregularly fluted, chomata broad in inner volutions, 

high and narrow in outer volutions.

Description. - Measurements are presented in table 25 

and on figure 28. Specimens elongate rapidly after the 

first volution, developing an elongate fusiform to subcylindrical shape, and attaining a length greater than 8 

mm and a diameter of about 2.5 mm in about seven volutions.

The lateral slopes tend to be straight or slight convex, 

and the poles range from bluntly pointed in the inner 

volutions to broadly rounded at maturity. The proloculus 

ranges from 0.09 to 0.16 mm in maximum outer 

diameter, with a mean near 0.11 mm. The spirotheca is 

composed of protheca with relatively thin and fine 

keriotheca. The septa are strongly but irregularly 

fluted. They are irregularly spaced; about 24 chambers 

are found in the fourth volution, 28 in the fifth, and 

25-32 in the sixth. The tunnel is nearly straight and 

widens rapidly in the outer volutions. It is bordered by 

low broad chomata in the inner volutions and high narrow chomata in the outer volutions. The tunnel is commonly less than half the chamber height.

Comparisons and remarks.-Triticites kehni has attributes intermediate between those of T. ohioensis 

Thompson and T. iatensis Thompson (1957). The fluting, 

though irregular, is more intense than in T. ohioensis 

and about the same as in T. iatensis. The elongate shape 

is more like that of T. ohioensis than that of T. iatensis. 

T. kehni is more loosely coiled than T. iatensis, and it 

commonly is more elongate and subcylindrical and 

larger. The number of septa and their fluting are 

similar. The proloculus in T. kehni is generally larger 

than that in T. iatensis, almost all the proloculi in T. 

kehni being larger than the mean for T. iatensis. T. 

kehni bears some resemblance to Kansanella tennis 

(Merchant and Keroher, 1939), but K. tennis is more 

slender and has more pointed poles throughout its 

growth.

Distribution.-T. kehni is found in the Carthage 

Limestone Member of the Sturgis Formation in Union 

and Webster Counties, western Kentucky, at localities 

16 and 22. The rocks are of Missourian age.

Specimens studied.-Only 16 oriented sections of this 

species were prepared from the three samples available.
60 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS

TABLE 13. -Measurements (in millimeters) for Beedeinasp. off. B. spissiplicata (Dunbar and Henbest), 1942

[Leaders (__), data not available or character absent]

Character

Diameter of proloculus ___ 

Radius vector 

1

2

3

4

5

6

7

8

Half length 

1

2

3

4

5

6

7

8

Volution height 

1

2

3

4

5

6

7

8

Wall thickness 

1

2

3

4

5

6

7

Tunnel width (axials) or 

septal spacing 

1

2

3

4

5

6

7

Measurements of fossils shown in plate 8, figures 11

11

0.09

0.10 

.18 

.33 

.47 

.71 

.95

0.14 

.26 

.52 

.76 

1.06 

1.45

0.03 

.14 

.14 

.16 

.23 

.26

.07

0.02 

.05 

.11 

.12 

.17

12

0.11

0.08 

.12 

.17 

.22 

.27 

.37 

.50 

.62

0.19 

.29 

.44 

.54 

.71 

.98 

1.23 

1.32

0.02 

.04 

.05 

.05 

.07 

.09 

.12 

.13

0.01 

.02

.03 

.03

0.02 

.03 

.04 

.04 

.07 

.10 

.10

14

0.125

0.12 

.18 

.26 

.33 

.43 

.60

0.18 

.37 

.58 

.85 

1.16 

1.93

0.04 

.06 

.08 

.09 

.10 

.17

.02 

.025

.04

0.03 

.035 

.05 

.07 

.08

16

0.10

0.08 

.12 

.20 

.30 

.43

0.17 

.28 

.59 

.88 

.16

0.02 

.03 

.08 

.10 

.13

.02

~To2

.03

0.03 

.04 

.07 

.08

20

0.10

0.10 

.14 

.18 

.25 

.36 

.53

0.13 

.26 

.42 

.64 

1.08 

1.40

0.02 

.04 

.05 

.07 

.11 

.16

0.01 

~03

 

0.02

.05 

.08 

.10 

.14

18

0.11

0.12 

.20 

.30 

.40 

.56 

.73

 

0.05 

.08 

.09 

.10 

.15 

.19

.01 

.025

.03 

.05

0.02 

.03 

.03 

.06 

.06

, 12, 14, 16, 20, 18, 21, 19;

21

0.075

0.09 

.13 

.19 

.27 

.43 

.64 

.79

0.03 

.04 

.05 

.09 

.16 

.18 

.19

0.01

.015 

.025 

.045 

.05

0.02 

.03 

.035 

.05 

.06 

.07

19

0.11

0.10 

.15 

.25 

.37 

.52

0.22 

.40 

.69 

1.10 

1.40

0.02 

.05 

.095 

.13 

.15

.025 

.06

.04 

.06 

.11

plate 9, figures 11, 13, 15

11

0.11

0.10 

.165 

.25 

.35 

.48

0.22 

.39 

.58 

.84 

1.15

0.03 

.06 

.09 

.10 

.14

0.01 

.02 

.025

0.02 

.03 

.05 

.09 

.15

13

0.095

0.10 

.19 

.28 

.44 

.59

0.21 

.43 

.88 

1.40 

1.61

0.03 

.08 

.09 

.15 

.15

0.02

.03

0.05 

.06 

.09

.17 

.22

14

0.10

0.13 

.21 

.30 

.46 

.60

0.21 

.45 

.76 

1.28 

1.90

0.05 

.07 

.10 

.145 

.16

.035 

.04

.05

15

0.09

0.11 

15 

.22 

.31 

.48

0.03 

.035 

.07 

.09 

.165

.02 

.02 

.05 

.05

0.015 

.02 

.03 

.04 

.04

The specimens are sparse in a limestone rich in other 

organic debris.

Designation of types.-The specimen illustrated on 

plate 19 as figure 5 is designated the holotype. The other 

specimens are paratypes.

Triticites calloms Dunbar and Henbest, 1942

Plate 20, figures 1-3, 5, 6

Triticites callosus Dunbar and Henbest, 1942, p. 136-138, pi. 21, 

figs. 15, pi. 22, figs. 1-11.

Description- Measurements for these specimens are 

shown in table 26 and on figure 29. Large inflated 

fusiform to rhomboidal specimens attaining 6-8 mm in 

length and 3-3.5 mm in width in seven to eight volutions. The inner volutions are tightly coiled and expand 

regularly. Two specimens illustrated by Dunar and 

Henbest (1942, pi. 22, figs. 2 and 5) attain lengths 

greater than 9 mm in 7V2-8 volutions. The proloculus is

large, having a range from 0.13 to 0.26 mm and a mean 

about 0.20 mm in seven specimens. The spirotheca 

thickens rapidly in the outer volutions and is composed 

of protheca with fine keriothecal structure. The septa 

are numerous, fairly evenly spaced, and irregularly 

fluted. Septal pores are common. The tunnel is narrow 

and wanders irregularly in the equatorial area. It is 

bordered by high narrow chomata. The tunnel extends 

to about half the chamber height.

Comparison and remarks.-The above description is 

based on the types and on topotypes. Most of the 

specimens found do not attain the large size described 

by Dunbar and Henbest, nor do they start with a proloculus as large as these authors reported. The 

specimens are more robust, larger, and more fluted than 

Triticites cullomensus Dunar and Condra, 1927, as 

noted by Dunbar and Henbest. The forms are associated 

with T. mediocris Dunbar and Henbest, 1942. 

Specimens of T. mediocris show many similarities to T.
SYSTEMATIC DESCRIPTIONS 61

callosus 

not attai 

Distril 

Greenup 

in Cumb 

of Virgil 

Specirr 

topotype

5

Triticites i 

figs. 2 5, 7

Descri^

0.4

0.3 

ull 

| Q 0.2

£* 

0.1

</) 

_iuj 0.10

gssy o.o5

HI1 0-3

I

Z 0.2 

O

3 °' 1 

o >

3.5 

3.0

1 " 

y 2.0

u. 

< 1.5

1.0 

0.5

Figure 17 

and Hen 

values fc

Dut are more elongate at each volution and 

n a comparable size. 

mtion.-This species is only known from t 

Limestone Member of the Matoon Formati< 

erland County, 111., at locality 1. The rocks a 

ian age. 

,ens studied. -The types and three samples 

s were studied.

"riticites mediocris Dunbar and Henbest, 1942

Plate 20, figures 4, 7-9

mediocris Dunbar and Henbest, 1942, p. 134-135, pi. ! 

, 8, 16.

ption. - Measurements for these specimens a

\ \ \ \

. * * 

 ; ». " *

 

 

.-. *' 

-

 % **,

-u !** < 

«HA

;: -i . v ! '

"fill

\ \ \ \

 

 

 

 ~~ * '.

 

r i

i i i i

io

he 

Dn 

re

of

21,

re

0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 0.9 1.0 

RADIUS VECTOR

-Graphs for Beedeina sp. aff. B. spissiplicata (Dunbar 

best), 1942. All measurements are in millimeters, and the 

>r each attribute are plotted against the radius vector.

TABLE 14. -Measurements (in millimeters) for Beedeina henbesti 

Douglass n. sp.

[Leaders ( _ _), data not available or character absent]

Measurements of fossils shown in plate 16, figures 

Character 29-31

29 30 31

Diameter of proloculus 0.16 0.21 0.11

Radius vector 

1 0.14 0.15 0.10

2 .23 .23 .14

3' .31 .34 .20

4 .43 .48 .30

5 .61 .66 .45

6 .83 .84 .59

7 _

8

Half length 

1 0.22 0.20

2 .375 .32

3 .80 .64

4 1.20 1.03

5 1.62 1.49

fi /Ml 1 QQ

7

8

Volution height 

1 0.03 0.03 0.02

2 .075 .07 .04

3 .08 .11 .06

4 .12 .135 .10

5 .18 .17 .14

6 .22 .19 .15

7

8

Wall thickness 

1

2 0.025

3 .02 .05 .015

4 .03 .015

5 .05 .04

6 .04

7

Tunnel width (axials) 

or septal spacing 

1 0.04 0.04 0.025

2 .05 .06 .035

3 .08 .07 .05

4 .12 .10 .05

5 .19 .03

6 _ .32 .24 .03

7

8

given in table 27 and on figure 30. The shell is elongate 

fusiform with irregular to convex lateral slopes and 

bluntly pointed poles. The proloculus ranges in maximum outer diameter from 0.07 to 0.20 mm, with a mean 

of 0.14 mm. The spirotheca is composed of protheca with 

fine but distinct keriotheca. It thickens rapidly in the 

outer whorls. The septa are numerous, fairly regularly 

spaced, and irregularly fluted throughout the shell. The 

tunnel is narrow and fairly straight, bordered by high 

narrow chomata. The tunnel is low, commonly less than 

half the chamber height. 

Comparison and remarks. - The above description is 

based on the types and on topotypes. Some of the dimensions differ a little from those in the original description. 

Dunbar and Henbest (1942, p. 134) described the proloculus as ranging from 0.075 to 0.10 mm in diameter, 

but their table indicated correctly a range in their three
62 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS

0.3 

0.2

0.1 

0.10

0.05 

0.3

0.2

3.5

3.0

2.0

1.5

1.0

0.5

-

. - '*

-

. / *

 *

 fc

-

-

~ t

-

 i

 '

-

.* * *

-

-

*.

* 9 _

-

-

0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 0.9 1.0 

RADIUS VECTOR

Figure 18.-Graphs for Beedeina hcnbesti Douglass n. sp. All measurements are in millimeters, and the values for each attribute are 

plotted against the radius vector.

specimens from 0.07 to 0.13 mm. Topotypes extend the 

range to 0.20 mm, and the specimen that they illustrated 

on their plate 21 as figure 7 is at the high end of this 

range.

Specimens of Tritidtes mediocris are more elongate 

at each volution than those of T. callosus with which 

they are associated, but they show similarity in most 

other attributes.

Distribution.-T. mediocris is known only from the 

Greenup Limestone Member of the Matoon Formation 

at locality 1 in Cumberland County, 111. It is associated 

with T. callosus Dunbar and Henbest.

Specimens studied.-The types and topotype 

specimens from three samples were studied.

Designation of types.-The specimen illustrated by 

Dunbar and Henbest (1942, pi. 21, fig. 3) is hereby 

designated the lectotype.

Tritidtes beardi Douglass n. sp.

Plate 20, figures 10-15

Name derivation.-Named for John Beard, who collected many of the 

fusulinid samples from western Kentucky and Illinois for this study.

Diagnosis. - Shells rhomboidal, attaining lengths of 7 

mm and widths of 3.6 mm in nine volutions. Proloculus 

large, averaging 0.17 mm. Spirotheca with coarse 

keriotheca. Tunnel straight and narrow with prominent 

high narrow chomata.

Description. - Measurements are presented in table 28 

and on figure 31. Specimens expand regularly from a 

large proloculus, attaining an inflated fusiform to rhomboidal shape that elongates only slightly more rapidly 

than it expands equatorially. The proloculi range from 

0.12 to 0.23 mm in maximum outer diameter and have a 

mean of 0.17 mm. The spirotheca is composed of protheca with coarse keriothecal structure. The septa are 

strongly but irregularly fluted across the entire shell. 

The tunnel is narrow and straight. It is bordered by 

prominent high narrow chomata.

Comparison and remarks. - Tritidtes beardi is similar 

in some ways to Dunbarinella eoextenta Thompson, 

1954. It has the same general size of proloculus and 

overall size, and the development of the chomata is 

similar. The septa are more tightly fluted than in most of 

the specimens of D. eoextenta, including the type, and 

the keriotheca in T. beardi is coarser than that of D. 

eoextenta. These specimens are assigned to Tritidtes instead of Dunbarinella because they do not fit the definition of Dunbarinella Thompson, 1942 or 1954, and are 

more like the types of Tritidtes than the types of Dunbarinella.

Distribution. T. beardi is known only from core 

samples at locality 19 in Union County, Ky. It 

represents a horizon younger than any other known 

fusulinid-bearing bed in the Eastern Interior Basin. The 

species is more advanced than T. callosus Dunbar and 

Henbest, 1942, from the Greenup Limestone Member of 

the Matoon Formation. The fossiliferous zone in the core 

is from unnamed beds of the Mauzy Formation of Early 

Permian age.

Specimens studied.-Ten specimens were obtained 

from the section of core at about 194-ft depth. Eight 

oriented sections were prepared.

Designation of types.-The specimen illustrated on 

plate 20 as figures 14 and 15 is designated the holotype. 

The other specimens are paratypes.
SYSTEMATIC DESCRIPTIONS 63

TABLE 15. -Measurements (in millimeters) for Beedeina ashlandensis Douglass n. sp.

[Leaders (__), data not available or character absent]

Character

Diameter of proloculus

Radius vector 

1

2

3

4

5

6

7

8

9

Half length

1

2

3

4

5

6

7

8

9

Volution height 

1

2

3

4

5

6

7

8

Wall thickness 

1

2

3

4

5

6

7

8

Tunnel width (axials) or 

septal spacing

1

2

3

4

5

6

7

Measurements of fossils shown in plate 11, figures 1,

1

0.17

0.09

.14

.19

.27

.37

.51

.67

.88

0.11

.23

.47

.62

.82

1.20

1.55

1.81

0.02

.04

.06

.08

.10

.13

.15

.22

0.02

.02

.03

0.03

.05

.05

.12

.15

.15

.22

2

0.15

0.11 

.19 

.28 

.43 

.53 

.71 

.88

0.13 

.30 

.50 

.72 

1.08 

1.41 

1.98

0.03 

.08 

.09 

.14 

.11 

.18 

.17

~T6l 

.02

~05

0.03 

.04 

.05 

.07

.11 

.19

4

0.12

0.10 

.17 

.27 

.39 

.56 

.75

 

0.14 

.34 

.48 

.81 

1.00 

1.71

0.04 

.07 

.10 

.12 

.17 

.19

0.02 

.02 

.03

.04

0.03 

.04 

.05 

.06 

.13 

.28

5

0.19

0.09 

.15 

.21 

.31 

.48 

.63

___

0.14 

.29 

.41 

.69 

1.16 

1.60

0.03 

.05 

.07 

.09 

.16 

.16

.18 

0.02

.02 

.03

 

.04 

.05 

.10

7

0.12

0.08 

.12 

.17 

.24 

.34 

.36 

.61 

.80

0.09 

.19 

.33 

.57 

.89 

1.13 

1.45 

1.89

0.03 

.04 

.06 

.07 

.09 

.11 

.16

"T02

~03

0.02 

.04 

.06 

.06 

.09 

.17 

.17

8

0.15

0.08 

.12 

.17 

.23 

.32 

.40 

.56

0.09 

.24 

.41 

.75 

1.08 

1.50 

1.87

0.02 

.04 

.05 

.06 

.09 

.08 

.16

~02 

.02

.03

0.03 

.04 

.06 

.08 

.14

.19

2, 4, 5, 7, 8, 3,

3

0.10

0.13 

.21 

.28 

.39 

.52 

.73

 -

0.04 

.07 

.07 

.11 

.14 

.20

.02 

.03 

.04 

.06

0.02 

.02 

.03 

.05 

.05

6, 9, 10

6

0.16

0.09 

.13 

.21 

.31

.44

____

0.02 

.04 

.08 

.11 

.13

 

____

0.04 

.05 

.06 

.10

9

0.12

0.08 

.14 

.20 

.30 

.42 

.61

____

 

0.03 

.06 

.07 

.10 

.12 

.18

.01

.02 

.04 

.06

0.01 

.02 

.03 

.04 

.05 

.05

10

0.17

0.10 

.15 

.21 

.29 

.42 

.54 

.67 

.89 

1.15

0.15 

.26 

.38 

.60 

.83 

1.01 

1.19 

1.42 

1.77

0.06 

.06 

.09 

.11 

.12 

.13 

.22 

.26

.03 

.03 

.05

0.03 

.04 

.05 

.09 

.13 

.18 

.20
64

TABLE 15. -Measurements (in millimeters) for Beedeina ashlandensis Douglass n. sp. -Continued

Character

Diameter of proloculus ___ 

Radius vector 

1

2

3

4

5

6

7

8

9

Half length 

1

2

3

4

5

6

7

8

9

Volution height 

1

2

3

4

5

6

7

8

Wall thickness 

1

2

3

4

5

6

7

8

Tunnel width (axials) or 

septal spacing 

1

2

3

4

5

6

7

8

Measurements of fossils shown in plate 11, figures 11, 17, 18, 16

11

0.18

0.11 

.15 

.22 

.32 

.42 

.60 

.77 

.95

0.14 

.19 

.27 

.40 

.68 

.87 

1.09 

1.27

0.03 

.04 

.07 

.10 

.13 

.15 

.17 

.17

.02 

.03

0.02 

.03 

.04 

.06 

.08 

.12 

.16 

.26

17

0.13

0.11 

.15 

.22 

.31 

.42 

.61 

.79

0.15 

.27 

.45 

.76 

.94 

1.29 

1.61

0.03 

.05 

.07 

.09 

.11 

.13 

.16

.01 

.03 

.03 

.03

0.03 

.04 

.07 

.11 

.16 

.20 

.30

18

0.10

0.09 

.13 

.20 

.29 

.39 

.55 

.69 

.83

0.11 

.23 

.34 

.50 

.69 

.93 

1.17 

1.45

0.03 

.04 

.06 

.09 

.10 

.15 

.13 

.15

0.01

~6i

.03 

.03

0.02 

.03 

.04 

.06 

.10 

.14 

.20 

.20

16

0.12

0.12 

.21 

.28

.44 

.59 

.77 

.97

 

0.05 

.08 

.08 

.14 

.18 

.18 

.21

"02

.03 

.07 

.06

0.03 

.03 

.04 

.05 

.05 

.06

19

0.16

0.11 

.16 

.24 

.35 

.44 

.57 

.74 

.91

____

0.02 

.05 

.07 

.10 

.09 

.13 

.17 

.17

"61 

.02 

.02

.03 

.04 

.07

0.01 

.04 

.05 

.05 

.07 

.08

13

0.15

0.12 

.18 

.25 

.35 

.47 

.63

 

0.04 

.05 

.06 

.11 

.12 

.15

0.01

~02 

.03 

.03

0.03 

.03 

.02 

.03 

.06

, 19, 13-15, 12

14

0.09

0.08 

.13 

.18 

.28 

.38 

.52 

.74

0.10 

.23 

.32 

.53 

.95 

1.33 

1.75

0.02 

.05 

.06 

.10 

.11 

.14 

.22

.02 

"02

.04

0.03 

.04 

.04 

.05 

.13 

.16 

.25

15

0.08

0.10 

.17 

.21 

.29 

.40 

.56 

.68 

.86

0.15 

.24 

.33 

.52 

.68 

.96 

1.50 

2.15

0.04 

.04 

.05 

.09 

.10 

.15 

.13 

.18

~02 

.03 

.04 

.04 

.04

0.03

.05 

.07

".13

.17

12

0.09

0.08 

.15 

.21 

.30 

.40 

.56

-- 

0.02 

.06 

.07 

.09 

.09 

.15

"61

.02 

.04 

.05

0.01 

.02 

.02 

.03 

.04 

.04
SYSTEMATIC DESCRIPTIONS 65

U.3

0.4

_i

LU X

z ^ t f~} n \J.O^

D >

K ^ 0.2

WALL 

THICKNESS HEIGHT 

0

P b^o L. OJ O <J1 

§ °' 2

3 0.1

0 

3.5

3.0

JE 2.5

Z 

y 2.0

LL

< 1.5

X

1.0

0.5

t 

_ 

~~ + 

^tx.-V**"

 * *

- ^'X1. ""

-f1**! 1

*

 ^ «

 . 

»% 

* * 

 . 

 *. 

f * *

1 1 1 1 1 1

-

-

-

-

-

:

-

i i

0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 0.9 1.0 1.1 1.2 1.3 1.4 1.5 1.6 1.7 1.8 1.9 2.0

RADIUS VECTOR

Figure 19. - Graphs for Beede,ina, ashlandensis Douglass n. sp. All measurements are in millimeters, and the values for each attribute are

plotted against the radius vector.
66 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS

TABLE 16. -Measurements (in millimeters) for Beedeina girtyi (Dunbar and Condra), 1927

[Leaders (__), data not available or character absent]

Character Measurements of fossils shown in plate 12, figures 1, 3, 4, 2; plate 13, figures 9, 6, 10

1 10

Diameter of proloculus ___ 0.21 

Radius vector

1 ______________ 0.20

2 ______________ .29

3 ______________ .42

4 __________ ___ .55

5 ______________ .76

6 ______________ .96

7 ______________ 1.16

8 ______________ 1.43

9 ______________ 1.78

10 ______________ ____ 

Half length

1 ______________ 0.30

2 ______________ .49

3 ______________ .76

4 ______________ 1.10

5 ______________ 1.35

6 ______________ 1.78

7 ______________ 2.28

8 ______________ 2.73 

Volution height

1 ______________ 0.08

2 ______________ .09

3 ______________ .12

4 ______________ .14

5 ______________ .20

6 ______________ .20

7 ______________ .20

8 ______________ .27

9 ______________ .35

10 ______________ __ 

Wall thickness

1 ______________ 0.02

2 ______________ .02

3 ______________ .02

4 ___ __________ __

5 ______________ .03

6 ______________ .04

7 ______________ .05

8 ______________ __

9 ______________ __

10 ______________ ____ 

Tunnel width (axials) or 

septal spacing

1 ______________ 0.04

2 ______________ .05

3 ______________ .06

4 ______________ .10

5 ______________ ____

6 ______________ .14

7 ______________ ,21

8 ______________ .35

9 __________ ___ __

0.18

0.12

.19

.29

.40

.58

.76

.98

1.15

0.18

.31

.43

.63

.98

1.35

1.74

2.39

0.05 

.07 

.09 

.11 

.17 

.19 

.21 

.17

0.02 

.02 

.03 

.02 

.03 

.03 

.04

0.03 

.03 

.09 

.12 

.18 

.26

0.08

0.11

.17

.25

.35

.48

.60

.79

.93

1.15

0.12

.16

.27

.46

.70

.96

1.22

1.57

0.03 

.05 

.08 

.10 

.12 

.12 

.19 

.15 

.22

0.02 

.03 

.02 

.01 

.02

.04 

.05 

.06 

.07 

.08 

.10 

.12 

.21

0.12

0.09

.15

.24

.36

.62

.83

1.07

1.88

0.03 

.05 

.09 

.12 

.26 

.21 

.23 

.31

0.02 

.02

.06

0.03 

.07

"08 

.08 

.08 

.12

0.19

0.18

.25

.37

.49

.65

.85

1.07

1.36

0.23

.39

.59

.81

1.00

1.27

1.56

1.97

0.06 

.07 

.12 

.12 

.18 

.19 

.21 

.29

0.03 

.02 

.02 

.02 

.02 

.02 

.03 

.04

0.03 

.06 

.06 

.10 

.12 

.15

0.21

0.10

.16

.24

.37

.50

.65

.86

1.07

1.31

0.13

.27

.49

.61

.78

.95

1.24

1.53

0.03 

.06 

.07 

.12 

.13 

.15 

.20 

.21 

.24

0.02 

.02 

.02 

.03 

.02 

.04 

.03 

.03 

.03

0.02 

.03 

.07 

.09 

.14 

.14 

.16

0.16

0.15

.19

.26

.38

.48

.64

.77

.97

1.20

1.40

0.04 

.05 

.06 

.12 

.09 

.16 

.14 

.20 

.22 

.21

0.02 

.03 

.02 

.03 

.04 

.04 

.05 

.11 

.08 

.06

0.03 

.03 

.04 

.05 

.06 

.12 

.16 

.14 

.20
SYSTEMATIC DESCRIPTIONS 67

u.o 

0.4

_i

LLJ I

|o 0.3

?l o,

0.1

_JUJ 0.10

-iZ

^^ 0.05

1-

1- 

1 0-3

I

§ 0.2

3 0.1 

o

3.5

3.0

|E 2.5

0 

Z 

y 2.0

LL

< 1.5

X

1.0

0.5

_

_

i

.. *.' V ' ** ' *

-

w . ...

-

n- ."* "*

. 4 * .

_

..; ' ""

. * .

 . 

.

. . .

. . t. '/ 

. * 

 »

 

 \

 X

»

i i i

..

 . "

.

* I* *

*

.

*

i i

-

-

-

i i i

0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 0.9 1.0 1.1 1.2 1.3 1.4 1.5 1.6 1.7 1.8 1.9 2.0

RADIUS VECTOR

Figure 20. - Graphs for Beedeina girtyi (Dunbar and Condra), 1927. All measurements are in millimeters, and the values for each attribute are

plotted against the radius vector.
68 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS

TABLE 17. -Measurements (in millimeters) for Beedeina sp. off. B. haworthi (Beede), 1916

[Leaders (__), data not available or character absent]

Character

1

Measurements of fossils shown in plate 13, figures 1, 4, 2, 3, 5, 8, 7

Diameter of proloculus ___ 0.32 

Radius vector

1 ______________ 0.15

2 ______________ .21

3 ______________ .31

4 ______________ .44

5 ______________ .59

6 ______________ .76

7 ______________ .95

8 ______________ 1.15

9 ______________ __

10 ______________ __ 

Half length

1 ______________ 0.19

2 ______________ .34

3 ______________ .72

4 ______________ 1.05

5 ______________ 1.42

6 ______________ 1.97

7 ______________ 2.32

8 ______________ 2.93 

Volution height

1 ______________ 0.03

2 ______________ .05

3 ______________ .09

4 _________ ____ .13

5 ______________ .15

6 ______________ .17

7 ______________ .20

8 ______________ .20

9 ______________ __

10 ______________ __ 

Wall thickness

1 ______________ ____

2 ______________ .03

3 ______________ .04

4 ______________ .03

5 ______________ __

6 ______________ ____

7 ______________ .07

8 ______________ __

9 ____________________

10 ______________ ____

Tunnel width (axials) or 

septal spacing

1 ______________ 0.06

2 ______________ .07

3 ______________ .10

4 ______________ .12

5 ______________ .15

6 ______________ .19

7 ______________ .26

8 ______________ ____

9 _____________________

0.21

0.15

.26

.35

.52

.72

.92

1.18

0.32

.49

.65

.91

1.19

1.62

2.26

0.05 

.11 

.09 

.17 

.19 

.21 

.25

.04 

.04

0.03 

.06 

.08 

.10 

.12 

.21 

.30

0.21

0.19

.27

.39

.51

.69

.89

1.40

0.04 

.08 

.11 

.13 

.17 

.19 

.25

0.02 

.02 

.03 

.04 

.08 

.10 

.08

0.03 

.03 

.04 

.04 

.07 

.07

0.14

0.11

.17

.24

.35

.49

.69

.90

1.09

0.16

.33

.49

.67

.93

1.26

1.68

0.03 

.06 

.07 

.11 

.13 

.21 

.21 

.19

0.01 

.02 

.03

.03 

.03

0.03 

.04 

.06 

.07 

.10 

.14 

.23 

.25

0.14

0.09 

.14 

.23 

.38 

.52

0.15

.34

.55

.89

1.28

0.03 

.06 

.10 

.11 

.16

0.01 

.02 

.03 

.02

0.02 

.04 

.04 

.09 

.17

0.23

0.13 

.22 

.32 

.49 

.66

0.19

.39

.67

1.21

1.57

0.04 

.09 

.10 

.17

.17

.02

.03 

.05

0.03 

.04 

.06 

.07 

.12

0.16

0.17

.24

.34

.45

.62

.81

1.04

0.05 

.07 

.10 

.11 

.16 

.19 

.22

0.02 

.01 

.03 

.03 

.04 

.07 

.07

0.03 

.04 

.04 

.05 

.06 

.13 

.14
SYSTEMATIC DESCRIPTIONS 69

0.5

0.4

i 11

' a °- 3

0.2 

0.1

<LU 0.10 

: = 0.05

I

^ 0.3

LU

I

§ 0.2

I3 0.1

3.5 

3.0

f 2.5 

O

y 2.0

LL.

< 1.5

I

1.0 

0.5

 * * * . I 

« . 

0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 0.9 1.0 1.1 1.2 1.3 1.4 1.5 1.6 1.7 1.8 1.9 2.0

RADIUS VECTOR

Figure 21. -Graphs for Beedeina sp. aff. B. haworthi (Beede), 1916. All measurements are in millimeters, and the values for each attribute are plotted against the radius vector.
70 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS

TABLE 18.-Measurements (in millimeters) for Beedeina acme (Dunbar andHenbest), 1942, and B. sp. off. B. acme (Dunbar andHenbest), 1942

[Leaders (__), data not available or character absent]

Character

Diameter of proloculus ___ 

Radius vector

1

2

3

4

5

6

7

Half length

1

2

3

4

5

6

7

Volution height 

1

2

3

4

5

6

7

Wall thickness

1

2

3

4

5

6

7

Tunnel width (axials) or 

septal spacing 

1

2

3

4

5

6

Measurements of fossils shown in plate 14, figures 1-4, 6, 5, 7, 8

1

0.16

0.12 

.17 

.23 

.32 

.43 

.62 

.84

.34 

.66 

.99 

1.47 

2.30 

2.69

0.04 

.05 

.06 

.08 

.12 

.20 

.23

.01 

.02 

.04 

.05 

.03

0.01 

.06 

.04 

.12 

.20 

.29

2

0.16

0.15 

.23 

.34 

.50 

.69 

.89

0.05 

.06 

.08 

.14 

.20 

.20

0.02 

.02 

.03 

.04 

.06 

.07

0.02 

.04 

.05 

.05 

.04

3

0.12

0.08 

.12 

.26 

.38 

.52 

.74 

.98

0.18 

.32 

.52 

1.00 

1.34 

2.35 

2.87

0.04 

.05 

.08 

.12 

.14 

.22 

.24

.015 

.02 

.03 

.02 

.03

0.02 

.04 

.05 

.10 

.12 

.20

4

0.14

0.12 

.19

.28 

.42 

.60 

.78

0.23 

.54 

1.07 

1.66 

2.39 

3.54

0.04 

.06 

.09 

.12 

.16 

.21

0.01 

.02 

.03 

.03 

.03

0.04 

.06 

.11 

.12

6

0.19

0.12

.17 

.24 

.34 

.50 

.71 

.95

0.22 

.41 

.61 

1.13 

1.37 

1.75 

2.27

0.04 

.06 

.06 

.10 

.16 

.20 

.24

.02 

.025 

.03 

.03 

.02 

.03

0.04 

.05 

.07 

.10 

.12 

.22

5

0.16

0.12 

.18 

.27 

.36 

.49 

.70 

1.00

0.30 

.52 

.70 

1.20 

1.52 

1.86 

2.48

0.05 

.08 

.09 

.10 

.13 

.22 

.30

0.02 

.03 

.05 

.04 

.04 

.04 

.04

0.06 

.10 

.03 

.06 

.10

7

0.16

0.11 

.14 

.20

.28 

.42 

.58

0.15 

.31 

.58 

.89 

1.47 

2.34

0.04 

.05 

.06 

.08 

.12 

.17

0.03 

.03 

.02 

.02 

.02 

.01

0.05 

.10

.14 

.18

8

0.19

0.11 

.16 

.25 

.33 

.47 

.66 

.92

0.17 

.25 

.48 

.81 

1.17 

1.45 

2.15

0.03 

.04 

.08 

.08 

.15 

.19 

.26

0.02 

.04 

.06 

.03 

.03 

.04 

.04

0.04 

.07 

.11

, 11, 12,9, 10

11

0.21

0.11 

.16 

.26 

.41 

.59 

.79 

.89

0.21 

.41 

.59 

.80 

1.19 

2.02 

3.04

0.03 

.05 

.10 

.14 

.17 

.21 

.17

0.02 

.02

.03 

.02 

.04 

.02

0.02 

.03 

.05 

.08 

.14

12

0.15

0.10 

.18 

.26 

.41 

.59 

.78 

.90

0.10 

.26 

.37 

.63 

1.23 

1.70 

2.05

0.03 

.07 

.08 

.15 

.18 

.20 

.13

.02 

.02 

.02 

.03 

.03 

.02

0.03 

.04 

.05

9

0.19

0.16 

.25 

.36 

.48 

.65 

.90

____

 

0.06 

.08 

.11 

.12 

.19 

.25

0.02 

.03 

.03 

.04 

.07 

.08

0.03 

0.3 

.03 

.04 

.05

in

0.8 

.13 

.21 

.34

.47 

.62 

.88

 -

0.04 

.05 

.08 

.13 

.15 

.25

.01 

.02 

.03 

.04 

.06

0.02 

.03 

.03 

.04 

.05 

.06
SYSTEMATIC DESCRIPTIONS 71

0.5

0.4

_i

UJ I

Z t- n o ZQ 0.3

^ 0,

0.1

0.10

0.05

t0.3

UJ

I

^ 0.2

d 0.1 

o

3.5 

3.0

fE 2.5

Z

"j 2.0

LL.

< 1.5 I

1.0 

0.5

< .« .- *

 jt 

0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 0.9 1.0 1.1 1.2 1.3 1.4 1.5 1.6 1.7 1.8 1.9 2.0

RADIUS VECTOR

Figure 22. - Graphs for Beedeina acme (Dunbar and Henbest), 1942. All measurements are in millimeters, and the values for each attribute

are plotted against the radius vector.
72 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS

TABLE 19. -Measurements (in millimeters) for Beedeina carmani 

(Thompson), 1936

[Leaders ( __ ), data not available or character absent]

Character

Diameter of 

proloculus ______ 

Radius vector 

1

2 _

3

4 _

5

6

Measurements of fossils shown in plate 15, figures 1-6

1

0.19

0.14 

.23 

.38 

.515 

.69

2

0.20

0.11 

.185 

.28 

.42 

.62

3

0.12

0.12 

.19 

.29 

.43 

.58

4

0.12

0.12 

.205 

.305 

.46 

.65

5

0.12

0.12 

.18 

.25 

.38 

.515

6

0.16

0.13 

.205 

.29 

.41 

.57

7

Half length 

1

2

3

4

5

6

7

Volution height 

1

2

3

4

5

6

0.27 

.55 

.88 

1.31 

1.95

0.055 

.09 

.135 

.15 

.175

0.16 

.33 

.82 

1.19

1.78

0.02 

.06 

.10 

.14 

.20

0.03 

.07 

.09 

.14 

.15

0.04 

.08 

.10 

.15 

.19

0.21 

.45 

.62 

.92 

1.35

0.04 

.05 

.06 

.12 

.13

0.26 

.42 

.68 

.99 

1.24 

1.78

0.04 

.07 

.09 

.12 

.15

7

Wall thickness 

1

2

3

4

5

6

Tunnel width (axials) 

or septal spacing 

1

2

3

4

5

6

.04

.07

0.055 

.09 

.14 

.18 

.32

0.025 

.025 

.06

0.04 

.06 

.12 

.215

0.01 

.015 

.03 

.05

0.03 

.04 

.05 

.05 

.05

0.02 

.05 

.04 

.08

0.02 

.04 

.04 

.04 

.04

____

0.06 

.07 

.11 

.12 

.18

0.015

.05 

.08

0.045 

.07 

.12 

.15 

.32

U.4

_j 0 u ' 1 d

Z* 

Z 0 0.2

?*

0.1

CO

_i uj 0.10 -J Z

< *

^ 0.05

K

K

T

V 0.3

LU

X

Z 0.2

O

D 0.1

_i

O

>

3.5

3.0

X

fc 25 C3

2

y 2.0

LL

_J

< 1 R

I ' b

1.0

0.5

.* :

** '

 m .\ m

. * *

w

- v *"*

M 

tf

r

-

-

-

 

s v ,

.

I

0 0

 

I 41

-

-

 -

 .

1

0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 0.9 

RADIUS VECTOR

1.0

Figure 23.-Graphs for Beedeina carmani (Thompson), 1936. All 

measurments are in millimeters, and the values for each attribute 

are plotted against the radius vector.
SYSTEMATIC DESCRIPTIONS 73

TABLE 20. -Measurements (in millimeters) for Beedeina sp.

[Leaders (__), data not available or character absent]

Character

Diameter of proloculus _ 

Radius vector 

1

2

3 __________________

4

5

6

Half length 

1

2

3

4

5

Volution height 

1

2

3

4

5

6

Wall thickness 

1

2

3

4

5

6

Tunnel width (axials) or 

septal spacing 

1

2

3

4

5

Measurements of fossils shown in plate 17, figures 16-19, 21, 22, 5, 6

16

0.10

0.095 

.16 

.25

0.12 

.24 

.40

0.03 

.07 

.09

.02

0.03 

.06 

.10

17

0.10

0.09 

.16 

.26

0.125 

.21 

.37

0.03 

.07 

.10

0.028 

.04 

.09

 

18

0.10

0.10 

.15 

.21 

.345 

.51

0.13 

.25 

.51 

.87

0.02 

.04 

.06 

.13 

.16

____

.04

~Toe

.04 

.07 

.11 

.20

19

0.09

0.11 

.15 

.27

0.17 

.38 

.68

0.04 

.05 

.12

0.025 

.025 

.06

0.045 

.06 

.09

____

21

0.12

0.10 

.165 

.27

0.143 

.23 

.39

0.03 

.06 

.10

0.02 

.03 

.03

0.03 

.06 

.08

____

22

0.13

0.12 

.19 

.27 

.41

0.12 

.23 

.43 

.525

0.05 

.06 

.09 

.13

____

____

0.05 

.07 

.10 

.14 

.14

5

0.10

0.10 

.16 

.225 

.34

0.13 

.24 

.38 

.51

0.04 

.06 

.07 

.115

~Toi

.025

0.02 

.04 

.07

6

0.105

0.115 

.225 

.32

0.19 

.34 

.61

0.04 

.10 

.10

 

0.02 

.05 

.09

 
74 FUSUUNID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS

TABLE 21. -Measurements (in millimeters) for Triticites skinneri Thompson, 1936

[Leaders (__), data not available or character absent]

Character

Diameter of proloculus

Radius vector 

1

2

3

4

5

6

7 _ _

Half length

1

2

3

4

5

6

7

Volution height 

1

2

3

4

5

6

7

Wall thickness 

1

2

3

4

5

6

7

Tunnel width (axials) or 

septal spacing

1

2

3

4

5

6

7

Measurements of fossils shown in plate 16, figures 1-10

1

0.06

0.07 

.13 

.21 

.29 

.42 

.62

0.13 

.29 

.51 

.82 

1.25 

1.90

0.02 

.05 

.09 

.07 

.14 

.17

0.02 

.04 

.07 

.11 

.18

2

0.06

0.10 

.17 

.28 

.43 

.63

0.17 

.29 

.49 

.99

1.47

0.02 

.07 

.11 

.17 

.20

 -

.03 

.04

0.02 

.04 

.07 

.12 

.18

3

0.05

0.07 

.125 

.21 

.23 

.33 

.50 

.69

0.11 

.28 

.45 

.73 

.95 

1.18 

1.61

0.015 

.05 

.08 

.12 

.17 

.20 

.21

0.015 

.025

0.02 

.05 

.08 

.11 

.13

4

0.045

0.07 

.12 

.18 

.31 

.49 

.73 

1.00

 

0.02 

.04 

.06 

.14 

.18 

.24 

.26

.005 

.03 

.035 

.03

0.02 

.03 

.04 

.04 

.05 

.05 

.06

5

0.03

0.06 

.10 

.17 

.25 

.38 

.53

0.02 

.03 

.06 

.08 

.12 

.14

.018 

.03 

.04

0.025 

.03 

.04 

.05 

.05

6

0.06

0.07 

.11 

.17 

.24 

.34 

.50 

.73

0.15 

.04 

.06 

.07 

.10 

.16 

.23

.015 

.02 

.05

0.01 

.025 

.03 

.04 

.05 

.09

7

0.07

0.10 

.16 

.26 

.42 

.63 

.89

0.13 

.31 

.73 

1.23 

2.03 

2.54

0.04 

.05 

.11 

.16 

.20 

.27

.025

.06

.07 

.08 

.16 

.20

8

0.06

0.10 

.14 

.23 

.37 

.58 

.82 

1.09

0.15 

.34 

.50 

.83 

1.19 

1.76 

2.45

0.04 

.05 

.09 

.14 

.21 

.23 

.27

.08

0.04 

.08 

.13 

.23

9

0.05

0.08 

.12 

.20 

.30 

.44 

.68 

.91

0.9 

.21 

.435 

.77 

1.24 

1.76 

2.15

0.02 

.04 

.07 

.10 

.14 

.23 

.23

 

 

.06 

.10 

.10

10

.13 

.21 

.32 

.52 

.78 

1.03

~~06 

.10 

.20 

.25 

.25

0.015 

.02 

.035 

.05 

.06

~~04 

.05 

.06 

.065 

.06
SYSTEMATIC DESCRIPTIONS 75

0.5 

0.4

UJ I

z Z o I- n t U - J

*~ 0.2 

0.1

c/) 

_j| 0.10

^y 0.05

a °-3

i

£ 0.2

0.1

3.5 

3.0 

2.5

CD

2.0

 \ - * "

i A v

I.""'

0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 0.9 1.0 1.1 1.2 1.3 1.4 1.5 1.6 1.7 1.8 1.9 2.0

RADIUS VECTOR

Figure 24.-Graphs for Triticites skinneri Thompson, 1936. All measurements are in millimeters, and the values for each attribute are

plotted against the radius vector.
76 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS

TABLE 22.-Measurements (in millimeters) for Triticites ricei Douglass n. sp.

[Leaders (__), data not available or character absent]

Diameter of proloculus ___ 

Radius vector 

1 _ _

2

3

4

5

6

7

8

Half length 

1

2

3

4

5

6

7

8

Volution height 

1

2

3

4

5

6

7

8

Wall thickness 

1

2

3

4 _ _

5

6

Tunnel width (axials) or 

septal spacing 

1

2

3

4

5

6

7

Measurements of fossils shown in plate 16, figures 11-18

11

0.12

0.13 

.19 

.28 

.42 

.58 

.76 

.94 

1.13

0.30 

.52 

.97 

1.33 

1.72 

2.16 

2.60 

3.18

0.05 

.07 

.09 

.13 

.16 

.18 

.18 

.18

.015 

.03

0.07 

.12 

.24 

.31

12

0.21

0.14 

.20 

.29 

.41 

.57 

.71 

.93 

1.10

0.32 

.46 

.69 

.98 

1.34 

1.75 

2.21 

2.73

0.03 

.06 

.09 

.12 

.15 

.15 

.20 

.17

~03 

.08

.10 

.13 

.23 

.34 

.42

13

0.11

0.15 

.21 

.31 

.47 

.64 

.86 

1.08

0.05 

.06 

.10 

.16 

.17 

.22 

.23

.01 

.01 

.04 

.05 

.05

0.02 

.03 

.04 

.06 

.065 

.07 

.06

14

0.08

0.11 

.18 

.28 

.42 

.58 

.79

 

0.04 

.05 

.10 

.13 

.16 

.21

0.02 

.02 

.04 

.04 

.04

0.02 

.02 

.04 

.04 

.04

15

0.16

0.12 

.18 

.26 

.37 

.57

0.31 

.56 

.89 

1.41 

1.82

0.04 

.06 

.08 

.11 

.19

 

.04

.06 

.10 

.12 

.21

16

0.09

0.10 

.15 

.22 

.34 

.51 

.68

0.18 

.36 

.66 

.99 

1.50 

1.79

0.03 

.05 

.07 

.12 

.16 

.17

~02 

.04

0.02 

.04 

.08 

.12 

.27

17

0.10

0.11 

.17 

.26 

.38 

.58 

.78

 

0.04 

.06 

.09 

.12 

.19 

.21

.02

.04 

.055

0.02 

.03 

.05 

.07 

.07

 

18

0.08

0.095 

.14 

.20 

.31 

.43

0.03 

.05 

.06 

.10 

.12

 

.04

0.02 

.03 

.04 

.05 

.06

 
SYSTEMATIC DESCRIPTIONS 77

U.D

0.4

O °-3

0.2

0.1

C/3

ft 0.10

y 0.05

hh1 0.3

I

3 0.1

0 

3.5

3.0

jE 2.5

0 

Z 

y 2.0

LL. 

< 1.5 I

1.0

0.5

. *****

-

 * * * *

V ******

-

_

-

. .

.' 

. %**

1 %* 1 1

1

.;... 

i i i

*

i i i

-

-

-

-

-

-

-

-

_

-

-

III!

0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 0.9 1.0 1.1 1.2 1.3 1.4 1.5 1.6 1.7 1.8 1.9 2.0

RADIUS VECTOR

Figure 25.-Graphs for Triticites ricei Douglass n. sp. All measurements are in millimeters, and the values for each attribute are plotted

against the radius vector.
78 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS

TABLE 23. -Measurements (in millimeters) for Triticites ohioensis 

Thompson, 1986

[Leaders ( _ _), data not available or character absent]

Character

Diameter of proloculus ___ 

Radius vector 

1

2 _ __

3

4

5 _

6

7

8 _ _

Half length 

1 _

2

3

4 _ _

5

6 _ _ _

7

Volution height 

1

2

3

4 _

5,

6 __________________

7

8

Wall thickness 

1

2

3

4

5

6 _

7

8

Tunnel width (axials) 

or septal spacing 

1 _ _ _

2 _ _

3

4

5

6

7

8

Measurements of fossils shown in plate 16

21

0.13

0.10 

.16 

.30 

.50 

.68 

.94

0.16 

.34 

.73 

1.31 

2.16 

3.33

0.03 

.06 

.14 

.20 

.19 

.26

.02

.04

0.04 

.07 

.11 

.21 

.48

22

0.12

0.10 

.16 

.26 

.42 

.64 

.89 

1.17

0.14 

.30 

.57 

.96 

1.64 

2.49 

3.38

0.025 

.06 

.10 

.16 

.22 

.25 

.29

.03 

.07 

.12

0.03 

.05 

.09 

.20 

.34 

.65

23

0.11

0.09 

.14 

.19 

.30 

.47 

.68 

.89

 

 -

0.01 

.04 

.05 

.10 

.16 

.20 

.21

0.02 

.02 

.04 

.04 

.04 

.06

0.03 

.03 

.04 

.05 

.05 

.04

figures 21-24

24

0.10

0.09 

.14 

.21 

.32 

.47 

.70 

.92

____

0.02 

.04 

.07 

.11 

.15 

.22 

.23

.02 

.04 

.06 

.05 

.06 

.05

.02 

.02 

.04 

.04
SYSTEMATIC DESCRIPTIONS

TABLE 23. -Measurements (in millimeters) for Triticites ohioensis Thompson, 1986 -Continued

79

Character

Diameter of proloculus __ 

Radius vector 

1

2

3

4

5

6

7

8

Half length 

1

2

3

4 _

5

6

7

Volution height 

1

2

3

4

5

6

7

8

Wall thickness 

1

2

3

4

5

6

7

8

Tunnel width (axials) or 

septal spacing 

1

2

3

4

5

6

7

Measurements of fossils shown in plate 18, figures 1-9

1

0.10

0.07 

.15 

.25 

.41 

.60 

.88 

1.10

0.11 

.34 

.56 

.89 

1.75 

3.09 

3.96

0.04 

.07 

.10 

.16 

.19 

.27 

.21

.02

.08

0.03 

.04 

.08 

.13 

.23 

.41

2

0.10

0.09 

.15 

.23 

.37 

.62 

.84 

1.15

0.15 

.31 

.84 

1.04 

1.79 

2.89- 

3.91

0.04 

.06 

.08 

.14 

.25 

.22 

.30

 

____

0.03 

.06 

.07 

.10 

.19

3

0.10

0.10 

.15 

.24 

.38 

.54 

.75 

.98

0.02 

.04 

.09 

.14 

.15 

.21 

.23

0.01

.04

.04

0.05 

.05 

.08 

.12 

.28 

.32

4

0.09

0.11 

.17 

.28 

.40 

.60 

.85 

1.08

0.03 

.06 

.10 

.12 

.20 

.24 

.24

.01 

.02 

.03 

.04 

0.5 

.07

0.03 

.04 

.04 

.05 

.08 

.04

5

0.13

0.11 

.16 

.25 

.42 

.64 

.85 

1.07

0.24 

.41 

.77 

1.24 

1.75 

2.24 

3.16

0.04 

.05 

.08 

.16 

.22 

.20 

.23

.02

.04 

.07

0.02 

.04 

.05

.17 

.37

6

0.09

0.09 

.15 

.19 

.30 

.45 

.70 

1.02 

1.26

0.02 

.05 

.05 

.10 

.15 

.25 

.33 

.24

.02 

.02 

.04 

.06 

.08

0.02 

.03 

.05 

.07 

.08 

.06 

.07

7

0.075

0.065 

.11 

.18 

.27 

.40 

.58 

.83 

1.05

0.09 

.21 

.37 

.61 

1.00 

1.90 

2.70

0.03 

.035 

.06 

.10 

.12 

.19 

.24

.02 

.04 

.06 

.08 

.05

.02 

.04 

.05 

.16 

.38 

.79

8

0.065

0.06 

.10 

.16 

.25 

.38 

.53 

.75

0.12 

.25 

.43 

.64 

1.07 

1.89 

2.92

0.35 

.04 

.05 

.09 

.13 

.15 

.23

"T02 

.02 

.03 

.04 

.05 

.045 

.05

0.045 

.065 

.09 

.15 

.34 

.63 

1.12

9

0.060

0.065 

.11 

.18 

.27 

.39 

.61 

.83

0.14 

.29 

.48 

.86 

1.53 

2.64 

3.50

0.04 

.05 

.07 

.09 

.14 

.19 

.22

.015 

.02 

.025 

.02 

.05 

.05 

.05

0.04 

.06 

.14 

.26 

.55 

.93
80

TABLE 23.-Measurements (in millimeters) for Triticites ohioensis Thompson, 1936- Continued

Character

Diameter of proloculus ___ 

Radius vector 

1

2

3

4

5

6

7

8

Half length 

1

2

3

4

5

6

7

Volution height 

1

2

3

4

5

6

7

8

Wall thickness 

1

2

3

4

5

6

7

8

Tunnel width (axials) or 

septal spacing 

1

2

3

4

5

6

7

8

Measurements of fossils shown in plate 18, figures 10-18

10

0.06

0.065 

.13 

.19 

.29 

.47 

.60 

.87

0.08 

.19 

.31 

.56 

.89 

1.52

0.03 

.06 

.06 

.09 

.14 

.18 

.22

.03 

.025 

.03 

.04 

.05 

.06

.04 

.05 

.13 

.32 

.54

11

0.06

0.055 

.10 

.155 

.23 

.38 

.57 

.87 

.93

0.025 

.04 

.05 

.08 

.14 

.19 

.25

.015 

.015 

.03 

.04 

.06 

.05 

.05

.045 

.055 

.065 

.105 

.14 

.18 

.34

12

0.06

0.09 

.15 

.26 

.38 

.56 

.78 

1.03

0.055 

.06 

.10 

.12 

.15 

.22 

.25

.025 

.03 

.025 

.045 

.055 

.05

.06 

.07 

.13 

.15 

.30

13

0.06

0.07 

.18 

.25 

.28 

.40 

.59 

.82 

.92

0.05 

.055 

.06 

.09 

.12 

.19 

.22

"02 

.03 

.03 

.03 

.06 

.06

0.03 

.045 

.06 

.08 

.10 

.13 

.16

14

0.05

0.055 

.095 

.145 

.25 

.39 

.55 

.76 

.81

 

0.03 

.04 

.07 

.08 

.13 

.17 

.21

.01 

.02 

.025 

.025 

.04 

.04 

.04

0.03 

.04 

.08 

.07 

.115 

.17 

.20 

.20

15

0.08

0.07 

.13 

.22 

.34 

.54 

.75 

.99

0.02 

.04 

.09 

.12 

.20 

.20 

.24

0.01 

.02 

.025 

.04 

.05 

.05

0.03 

.03 

.04 

.08 

.05 

.06

16

0.10

0.11 

.16 

.26 

.41 

.62 

.85 

1.09

0.17 

.34 

.67 

1.02 

2.21 

2.87 

3.71

0.04 

.05 

.10 

.15 

.21 

.22 

.25

 

.03

0.02 

.05 

.10 

.18

 

17

0.09

0.10 

.17 

.27 

.42 

.62 

.84

0.20 

.39 

.68 

1.19 

1.99 

2.66

0.04 

.08 

.10 

.15 

.20 

.23

 

.02

.04

0.02 

.03 

.06 

.10 

.18

 

18

0.09

0.08 

.15 

.22 

.34 

.48 

.68 

.88

0.13 

.25 

.45 

.75 

1.33 

2.18 

2.95

0.02 

.07 

.08 

.12 

.14 

.21 

.21

.02 

.03

.03

0.03 

.05 

.08 

.13 

.21

 
SYSTEMATIC DESCRIPTIONS 81

TABLE 23. -Measurements (in millimeters) for Triticites ohioensis Thompson, 1936- Continued

Character

Diameter of proloculus _ 

Radius vector 

1

2

3 __________________

4

5

6

7

8

Half length 

1

2

3

4

5

6

7

Volution height 

1

2

3

4

5

6

7

8

Wall thickness 

1

2

3

4

5

6

7

8

Tunnel width (axials) or 

septal spacing 

1

2

3

4

5

6

7

Measurements of fossils shown in plate 18, figures 19-27

19

0.12

0.12 

.21 

.31 

.46 

.62 

.85 

1.03

0.22 

.47 

.83 

1.46 

2.38 

3.08

0.05 

.08 

.10 

.15 

.17 

.22 

.20

.04

.09

0.04 

.07 

.11

20

0.10

0.09 

.17

.27 

.40 

.58

.77

G.17 

.33 

.70 

1.12 

1.91 

2.76

0.03 

.08 

.10 

.14 

.17 

.20

~03 

.03 

.04 

.05 

.06

.03 

.04 

.04 

.08 

.16 

.23 

.33

21

0.08

0.08 

.13 

.20 

.31 

.45 

.65 

.89 

1.05

 -

0.02 

.05 

.07 

.11 

.13 

.21 

.23 

.16

.02 

.02 

.03 

.03 

..07 

.07

0.02 

.025 

.04 

.04 

.05 

.08 

.05

22

0.06

0.10 

.17 

.26 

.39 

.56 

.77 

1.00

 

0.03 

.07 

.09 

.12 

.17 

.21 

.23

.018 

.02 

.025 

.04 

.07 

.08

0.02 

.03 

.05 

.06 

.06 

.04 

.04

23

0.08

0.08 

.13 

.20 

.45 

.62

 

 

0.02 

.05 

.07 

.08 

.17 

.11

0.02

.09

 

0.04 

.08 

.12 

.28

24

0.07

0.08 

.13 

.19 

.31 

.44 

.66 

.85

0.10 

24 

.45 

1.02 

1.89 

2.38 

2.72

0.03 

.05 

.06 

.11 

.14 

.21 

.19

~03

 

____

.03 

.04 

.12 

.19

25

0.06

0.10 

.14 

.21 

.32 

.45 

.66

;:::

0.03 

.04 

.07 

.10 

.14 

.20

____

.015 

.02 

.02 

.08 

.08

0.02 

.05 

.04 

.06 

.05 

.05

26

0.08

0.08 

.14 

.19 

.30 

.60 

.78

 

0.03 

.06 

.06 

.11 

.14 

.16 

.18

.02 

.025 

.06 

.06

0.02 

.02 

.03 

.035 

.06 

.04 

.05

27

0.09

0.12 

.18 

.24 

.35 

.46 

.62

 

 

 

0.04 

.05 

.07 

.11 

.12 

.15

0.01 

.02 

.02 

.035 

.04

0.02 

.03 

.05 

.04 

.04 

.05
82 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS 

TABLE 23. -Measurements (in millimeters) for Triticites ohioensis Thompson, 1936- Continued

Character

Diameter of proloculus _ 

Radius vector 

1

2

3

4

5

6_

7

8

Half length 

1 _ __

2

3

4 __

5_ _ _

6

7

Volution height 

1

2

3

4

5 _

6

7

8

Wall thickness

1 _ _

2

3

4 _ _

5

6

7 _ _

8

Tunnel width (axials) or 

septal spacing 

1

2

3

4_

5 _

6

7

8

Measurements of fossils shown in plate 18, figures 28-36

28

0.09

0.09 

.14 

.22 

.54

0.12 

.33 

.50 

.90 

2.25 

2.92

0.02 

.05 

.08 

.10 

.22

.04

.04 

.07 

.14

29

0.08

0.08 

.15 

.24 

.37 

.57

0.12 

.25 

.42 

.72 

1.25 

2.08

0.03 

.07 

.08 

.14 

.20

0.02 

.05 

.07

 

0.04 

.06 

.11 

.20

30

0.08

0.10 

.16 

.24 

.34 

.46 

.62

 

0.04 

.05 

.08 

.10 

.12 

.16

~0~01 

.01 

.01 

.04 

.05

0.02 

0.3 

.04 

.05 

.04 

.04

31

0.09

0.08 

.14 

.22 

.34 

.47

0.14 

.30 

.42 

.82 

1.32 

2.07

0.01 

.03 

.05 

.08 

.12 

.13

 

 

~Tii

.15

32

0.11

0.08 

.13 

.20 

.32 

.45 

.64

0.20 

.40 

.86 

1.12 

1.68 

2.45

0.02 

.04 

.07 

.12 

.12 

.20

____

.04

 -

.05 

.10 

.17

 

33

0.09

0.11 

.16 

.23 

.37 

.52 

.70

0.16 

.31 

.67 

1.34 

2.08 

3.07

0.04 

.04 

.08 

.13 

.16 

.18

.01

.05 

.06

0.03 

.05

.11

34

0.07

0.10 

.17 

.26 

.41 

.59 

.79

0.14 

.25 

.37 

.63 

1.25 

1.81 

2.31

0.04 

.07 

.09 

.15 

.18 

.20

 

.02 

.03 

.04

0.035 

.04 

.06 

.09

 

35

0.14

0.12 

.18 

.26 

.37 

.53 

.75

0.26 

.48 

.86 

1.44 

1.97 

2.68

0.04 

.06 

.08 

.11 

.16 

.21

 

 -

0.02 

.04 

.065 

.14 

.17 

.22

36

0.08

0.09 

.15 

.24 

.39 

.55 

.77

0.10 

.24 

.44 

.77 

1.28 

2.16 

2.86

0.03 

.06 

.09 

.16 

.19 

.20

____

.03

 

0.02 

.06 

.11 

.21 

.32

 
SYSTEMATIC DESCRIPTIONS

TABLE 23. -Measurements (in millimeters) for Triticites ohioensis Thompson, 1936 -Continued

83

Character

Diameter of proloculus ___ 

Radius vector 

1

2

3

4

5 _ _

6

7 _ _

8 _

Half length 

1 _ _

2

3

4

5

6

7

Volution height 

1

2

3

4

5

6

7

8

Wall thickness 

1

2

3

4

5_ _

6

7 _ _

8

Tunnel width (axials) or 

septal spacing 

1

2

3

4

5_ _

6

7

Measurements of fossils shown in plate 18, figures 37-45

37

0.07

0.09 

.13 

.20 

.29 

.44 

.63

0.03 

.04 

.06 

.07 

.15 

.19

0.01 

.01 

.02 

.04 

.05

0.02 

.03 

.04 

.05 

.07 

.07

38

0.07

0.075 

.15 

.36 

.55

 

0.04 

.07 

.08 

.11 

.18

"b~oi5

.015 

.035 

.05

0.03 

.04 

.06 

.06 

.09

39

0.06

0.08 

.14 

.20 

.48 

.67

 

0.03 

.06 

.06 

.11 

.16 

.19

~0~6l 

.02 

.025 

.03

0.02 

.03 

.04 

.05 

.06 

.06

40

0.155

0.10 

.16 

.25 

.40 

.59

 

0.28 

.43 

1.05 

1.56 

2.32

0.03 

.07 

.08 

.15 

.19

~06 

.115

____

.07 

.12 

.24

41

0.14

0.11 

.19 

.36

.57

 

0.28 

.60 

1.63 

2.38

0.05 

.08 

.16 

.21

oToi

.06

0.05 

.08 

.14

 

42

0.12

0.12 

.20 

.32 

.45

 

0.23 

.51 

1.06 

1.66

0.06 

.08 

.12 

.13

 

.05

0.045 

.08 

.14

43

0.15

0.12 

.20 

.31 

.47 

.675

0.33 

.62 

1.03 

1.87 

2.27

0.05 

.07 

.11 

.16 

.205

 

____

.04 

.06 

.045 

.06 

.065

0.02 

.07 

.15 

.25

44

0.08

0.09 

.15 

.23 

.34 

.50 

.69 

.91

____

 

0.03 

.06 

.08 

.11 

.16 

.21

 

.025 

.05

0.025 

.025 

.04 

.045 

.06 

.05 

.05

45

0.10

0.11 

.18 

.28 

.46 

.66

::::

 

0.05 

.07 

.11 

.17 

.21

0.01 

.015 

.025

0.02 

.02 

.04 

.05 

.03
84

1.1

1.0 

0.9 

0.8 

0.7

Q

> 0.6

0.5 

0.4 

0.3 

0.2 

0.1

_j[{j 0.10 

^ 

$% 0.05

hh1 0.3

X

0.2

0.1

o

3.5 

3.0 

2.5 - 

2.0 

< 1.5 I 

 

 . * 

vO *

f

0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 0.9 1.0 1.1 1.2 1.3 1.4 1.5 1.6 1.7 1.8 1.9 2.0

RADIUS VECTOR

Figure 26.-Graphs for Triticites ohioensis Thompson, 1936. All measurements are in millimeters, and the values for each attribute are

plotted against the radius vector.
SYSTEMATIC DESCRIPTIONS 85

IABLE Z4.- measurements (in millimeters) /or Triticites smytm 

Douglass n. sp.

[Leaders ( __ ), data not available or character absent]

Character

Diameter of proloculus ___ 

Radius vector

1

2

3

4

5

Half length

1

2

3

4

5

Volution height 

1

2

3

4

5

Wall thickness 

1

2

3

4

5

Tunnel width (axials) or 

septal spacing

1

2

3

4

5

Measurements of fossils shown in plate 16,

25

0.13

0.12 

.17 

.26 

.37 

.53

0.19 

.36 

.59 

.81 

1.22

0.04 

.06 

.08 

.11 

.16

.05

.055

0.03 

.04 

.10 

.17 

.23

26

0.11

0.09 

.15 

.24 

.38 

.54

0.20 

.32 

.56 

1.01 

1.43

0.03 

.05 

.09 

.14 

.16

0.03 

.05 

.10 

.17

0.03 

.04 

.08 

.15

27

0.11

0.11 

.15 

.22 

.33 

.45

 

0.03 

.04 

.07 

.10 

.13

.01 

.04 

.05

0.02 

.03 

.04 

.05 

.06

figures 25-28

28

0.13

0.13 

.19 

.29 

.41

 

0.03 

.07 

.10 

.12

.025 

.028 

.03

0.028 

.04 

.05 

.07

WALL TUNNEL 

ICKNESS WIDTH

HALFEIGHT LNGTH VOLUTION TH

0.3 

0.2

0.1

0.10 

0.05

0.3 

0.2 

0.1

3.5 

3.0 

2.5 

2.0 

1.5 

1.0 

0.5

I I I

-

.* ********

-

V**i i I

I I I I

-

-

-

I I I I

0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 0.9 1.0 

RADIUS VECTOR

Figure 27. - Graphs for Triticites smythi Douglass n. sp. All measurements are in millimeters, and the values for each attribute are plotted against the radius vector.
86 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS

TABLE 25.-Measurements (in millimeters) for Triticites kehni Douglass n. sp.

[Leaders (__), data not available or character absent]

Character

Diameter of proloculus ___ 

Radius vector 

1

2

3

4

5

6

7

Half length 

1

2

3

4

5

6

7

Volution height 

1

2

3

4

5

6

7

Wall thickness 

1

2

3

4

5

6

7_ _

Tunnel width (axials) or 

septal spacing 

1

2 _ _

3

4

5

6

Measurements of fossils shown in plate 19, figures 1-i

1

0.11

0.16 

.25 

.38 

.55 

.80 

1.14

0.21 

.44 

.90 

1.69 

2.40 

3.58

0.09 

.09 

.12 

.16 

.25 

.32

.02 

.02 

.05

0.04 

.07 

.08 

.17

2

0.13

0.11 

.18 

.29 

.41 

.67 

1.01

0.21 

.40 

.69 

1.06 

2.01 

3.49

0.03 

.06 

.10 

.12 

.25 

.33

.04 

.08 

.16 

.31

3

0.09

0.12 

.22 

.32 

.45 

.66 

1.00

0.04 

.09 

.10 

.13 

.21 

.35

.18 

.02 

.02 

.04 

.07

0.02 

.03 

.04

.06

4

0.12

0.14 

.25 

.33 

.47 

.68 

.98

0.23 

.42 

.56 

.83 

1.38 

1.84 

2.75

0.05 

.09 

.09 

.14 

.20 

.30

.07

0.04 

.08 

.11 

.22

____

5

0.13

0.11 

.18 

.25 

.36 

.53 

.81 

1.17

0.22 

.40 

.73 

1.13 

1.74 

3.07 

3.95

0.03 

.06 

.07 

.12 

.16 

.30 

.33

.04 

.16 

.14

0.04 

.08 

.13 

.23 

.48 

.63

6

0.16

0.18 

.27 

.42 

.59 

.86 

1.17

0.03 

.07 

.10 

.12 

.20 

.25 

.28

.03 

.03 

.02 

.05 

.06

0.02 

.05 

.06 

.06 

.09

3, 10

7

0.09

0.12 

.19 

.28 

.40 

.60 

.85 

1.14

 

0.06 

.09 

.14 

.17 

.25 

.30

.01 

.02 

.04 

.045 

.055

0.02 

.025 

.03 

.05 

.06 

.06

8

0.10

0.14 

.23 

.31 

.52 

.75 

1.12

0.34 

.66 

1.16 

1.65 

2.48 

3.44 

4.45

0.05 

.09 

.09 

.21

 

0.03 

.07 

.09 

.16 

.17

10

0.15

0.16 

.23 

.40 

.63 

.94

0.34 

.69 

1.26 

2.37 

2.94

0.05 

.07 

.16 

.22 

.30

.05 

.07

0.06 

.08 

.15

 
SYSTEMATIC DESCRIPTIONS 87

UJ I

Z h0.5 

0.4 

0.3 

0.2

0.1

CO

,m 0.10 

g

0.05

^ 0.3

LU

I

0.2

0.1

3.5

3.0 -

2.5 -

2.0 -

1.5 -

1.0 -

0.5 -

I* * 

0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 0.9 1.0 1.1 1.2 1.3 1.4 1.5 1.6 1.7 1.8 1.9 2.0

RADIUS VECTOR

Figure 28.-Graphs for Triticites kehni Douglass n. sp. All measurements are in millimeters, and the values for each attribute are

plotted against the radius vector.
88 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS

TABLE 26. -Measurements (in millimeters) for Triticites callosus Dunbar and Henbest, 1942

[Leaders ( __ ), data not available or character absent]

Diameter of 

proloculus

Radius vector 

1

2

3

4 ________________ 

5

6 ________________ 

7

8

9

Half length 

1

2

3

4

5

6

7

8

9

Volution height 

1

2

3

4

5

6 _

7

8

9

Wall thickness 

1

2

3

4

5

6

Tunnel width (axials) 

or septal spacing 

1

2

3

4

5

6

7

Measurements of fossi

1

0.22

0.23 

.38 

.63 

.73 

1.24 

1.61

0.41 

.84 

1.44 

2.12 

3.05 

3.90

0.09 

.14 

.24 

.30 

.32 

.36

0.03 

.04 

.10

0.07 

.105

2

0.15

0.14 

.22 

.35 

.54 

.835 

1.155 

1.48

0.24 

.47 

.68 

1.07 

1.70 

2.16 

3.34

0.05 

.05 

.12 

.20 

.29 

.32 

.33

.05 

.09 

.09

0.04 

.06 

.095 

.12 

.21 

.23

s shown in plate 20, figures 1-3, 5, 6

3

0.13

0.13 

.21 

.35 

.48 

.70 

.99 

1.34 

1.74

0.18 

.31 

.54 

.75 

1.12 

1.65 

2.29 

3.21

0.04 

.08 

.14 

.14 

.22 

.28 

.35 

.39

.04

.05

0.04 

.06 

.15

.41

5

0.16

0.19 

.32 

.50 

.71 

1.00 

1.32

 

0.06 

.13 

.17 

.20 

.29 

.31

.03 

.05 

.06 

.08

0.04 

.05 

.05 

.06 

.10

6

0.15

0.18 

.26 

.50 

.60 

.645 

.90 

1.26

 

 

0.07 

.09 

.135 

.23 

.25 

.36

.02

.07

0.03 

.04 

.06 

.07 

.10

8
SYSTEMATIC DESCRIPTIONS 89

u.o 

0.4

iiJ I

IQ 0.3

*~ 0.2

0.1

C/D

_jft 0.10

^^ 0.05

IIi 0.3 LJJ

I

I 0.2

3 0.1

O 

3.5

3.0

f 2.5

0

z

y 2.0

LL. 

< 1.5

1.0

0.5

_

. « "

 i 

<

-

4

 «

 

*

*

i 

 ^

* 

, 

 

 *

i i

~

-

-

-

*

-

_

-

i

0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 0.9 1.0 1.1 1.2 1.3 1.4 1.5 1.6 1.7 1.8 1.9 2.0

RADIUS VECTOR

Figure 29.-Graphs for Triticites callosus Dunbar and Henbest, 1942. All measurements are in millimeters, and the values for each attribute

are plotted against the radius vector.
90 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS

TABLE 27. -Measurements (in millimeters) for Triticites mediocris 

Dunbar and Henbest, 1942

[Leaders ( __ ), data not available or character absent]

Diameter of 

proloculus

Radius vector 

1

2

3

4

5

6

7

Half length 

1

2 _

3

4

5

6

7

Volution height 

1

2

3

4

5 ___________________ 

6

7 _

Wall thickness 

1

2

3

4 ___________________ 

5

Tunnel width (axials) 

or septal spacing 

1

2

3

4

5

Measurements of fossils shown in plate 20, figures 4, 7-9

4

0.18

0.16 

.25 

.41 

.64 

.90

0.22 

.50 

1.10 

1.91 

2.68

0.04 

.10 

.16 

.22 

.25

.03

 

0.03 

.075 

.11 

.19

7

0.20

0.13 

.21 

.335 

.50 

.705 

.98 

1.20

0.22 

.40 

.66 

.97 

1.56 

2.32 

3.11

0.035 

.06 

.13 

.15 

.21 

.27 

.49

.03

 

0.04 

.07 

.135 

.23 

.34

8

0.14

0.15 

.24 

.39 

.62 

.84 

1.13 

1.40

0.06 

.08 

.15 

.22 

.22 

.29 

.27

.02 

.02 

.03 

.055

0.015 

.025 

.05 

.075 

.09

9

0.15

0.14 

.25 

.41 

.60 

.89 

1.12

0.17 

.35 

.51 

.70 

1.04 

1.51

0.035 

.11 

.16 

.18 

.28 

.24

.05

0.035 

.06 

.08 

.14 

.14
SYSTEMATIC DESCRIPTIONS 91

U.D

0.4

UJ I

z I- n o z Q °-3

H 0.2

0.1

_,| 0.10

^^ 0.05

1-

1- 

1 0.3

I

| 0-2

3 0.1

o

3.5

3.0

JE 2.5

O 

Z 

y 2.0

LL.

< 1.5

I

1.0

0.5

_

_ *

». " *'

-

 * .

-

1 *

-

 * ,

l % " I** I I

*

.

* *

S

.

 * 

.

.

-

-

-

-

-

_

-

_

-

I I

0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 0.9 1.0 1.1 1.2 1.3 1.4 1.5 1.6 1.7 1.8 1.9 2.0

RADIUS VECTOR

Figure 30. - Graphs for Triticites mediocris Dunbar and Henbest, 1942. All measurements are in millimeters, and the values for each attribute are plotted against the radius vector.
92 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS

TABLE 28. -Measurements (in millimeters) for Triticites beardi 

Douglass n. sp.

[Leaders (__), data not available or character absent]

Measurements of fossils shown in plate 20, figures 10, 

12-15 Character

10

Diameter of proloculus ______ 0.21 0.23 0.20 

Radius vector

1 _________________ 0.14 0.19 0.17

2 _________________ .20 .29 .26

3 _________________ .23 .425 .44

4 _________________ .33 .61 .62

5 _________________ .46 .845 .88

6 _________________ .67 1.03 1.15

7 _________________ .70 __ 1.39

8 _________________ __ __ 1.64

9 _________________ __ __ 1.86 

Half length

1 _________________ 0.21 0.33 0.31

2 _________________ .37 .62 .44

3 _________________ .55 .96 .60

4 _________________ .80 1.52 .90

5 _________________ 1.09 2.27 1.09

6 _________________ 1.48 __ 1.41

7 _________________ __ __ 2.06

8 _________________ __ __ 2.79

9 _________________ __ __ 3.59 

Volution height

1 _________________ 0.03 0.07 0.05

2 _________________ .06 .10 .09

3 _________________ .13 .13 .18

4 _________________ .13 .18 .19

5 _________________ .20 .23 .25

6 _________________ .23 .29 .27

7 _________________ __ __ .24

8 _________________ __ __ .25

9 _________________ __ __ .24 

Wall thickness

1 _________________ 0.02 0.02 0.02

2 _________________ .03 .02 .03

3 _________________ .04 .03 .04

4 _________________ .03 .05 .05

5 _________________ .04 .05 .06

6 _________________ .05 __ .06 

Tunnel width (axials) 

or septal spacing

1 _________________ __ 0.045 0.07

2 _________________ __ .055 .08

3 _________________ __ .08 .08

4 _________________ __ .13 .12

5 _________________ __ __ .20

6 _________________ __ __ .28

7 _________________ __ __ .25

8 _________________ __ __ .29
SYSTEMATIC DESCRIPTIONS 93

LU X

0.5 

0.4

0.2

0.1

^ 0.10 

= 0.05

1 0-3

=J 0.1

o

3.5 

3.0

h- 2.5

z

y 2.0

LJ_

< 1.5

x

1.0 

0.5

0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 0.9 1.0 1.1 1.2 1.3 1.4 1.5 1.6 1.7 1.8 1.9 2.0

RADIUS VECTOR

Figure 31.-Graphs for Triticites beardi Douglass n. sp. All measurements are in millimeters, and the values for each attribute are

plotted against the radius vector.
94 FUSULINID BIOSTRATIGRAPHY AND CORRELATIONS BETWEEN THE APPALACHIAN AND EASTERN INTERIOR BASINS

REFERENCES CITED

Beede, J.W., 1916, New species of fossils from the Pennsylvanian and 

and Permian rocks of Kansas and Oklahoma: Indiana University 

Studies, v. 3, no. 29, 15 p.

Condit, D.D., 1912, Conemaugh formation in Ohio: Ohio Geological 

Survey, 4th ser., Bulletin 17, 363 p.

Crider, A.F., 1913, Economic geology of Tell City and Owensboro 

quadrangles: Kentucky Geological Survey, 4th ser., v. 1, p. 

263-316.

Douglass, R.C., 1977, The development of fusulinid biostatigraphy, 

in Kauffman, E.G., and Hazel, J.E., Concepts and methods of 

biostratigraphy: Stroudsburg, Pa., Dowden, Hutchinson, & Ross, 

p. 463-481, 6 figs.

_____1979, The distribution of fusulinids and their correlation between the Illinois Basin and the Appalachian Basin, in Palmer, 

J.E., and Dutcher, R.R., eds. Depositional and structural history of 

the Pennsylvanian System of the Illinois Basin. Part 2, Invited 

papers-Field trip 9, Ninth International Congress of Carboniferous Stratigraphy and Geology: Illinois State Geological 

Survey [Guidebook Series, no. 15], p. 15-20, 3 figs.

_____1981, Evolution of some primitive fusulinids [abs.]: Geological Society of America, Abstracts with Programs, v. 13, no. 7, 

p. 441. 

___1982, Fusulinids of the Atoka Formation, south-central

Oklahoma [abs.]: Geological Society of America, Abstracts with 

Programs, v. 14, no. 3, p. 110.

Douglass, R.C., and Nestell, M.K., 1984, Fusulinids of the Atoka Formation, lower Middle Pennsylvanian, south-central Oklahoma, in 

Sutherland, P.K., and Manger, W.L., eds., The Atokan Series 

(Pennsylvanian) and its boundaries-A symposium: Oklahoma 

Geological Survey Bulletin 136, p. 19-39, 5 pis., 7 figs.

Dunbar, C.O., and Condra, G.E., 1927, The Fusulinidae of the Pennsylvanian Systems in Nebraska: Nebraska Geological Survey, 2d ser., 

Bulletin 2, 135 p., 15 pis., 13 figs.

Dunbar, C.O., and Henbest, L.G., 1930, The fusulinid genera 

Fusulina, Fusulinella,, and Wedekindella: American Journal of 

Science, 5th ser., v. 20, p. 357-364, 1 fig.

_____1931, Wedekindia, a new fusulinid name: American Journal 

of Science, 5th ser., v. 21, p. 458.

___1933, Genus Wedekindellina Dunbar and Henbest, new 

name, in Cushman, J.A., Foraminifera, their classification and 

economic use (2d ed.): Cushman Laboratory for Foraminiferal 

Research Special Publication 4, p. 134.

___1942, Pennsylvanian Fusulinidae of Illinois: Illinois State

Geological Survey Bulletin 67, 218 p., 23 pis., 13 figs.

Galloway, J.J., 1933, A manual of Forminifera: Bloomington, Ind., 

Principia Press, 483 p., 42 pis.

Henbest, L.G., 1928, Fusulinellas from the Stonefort limestone member of the Tradewater formation (Illinois]: Journal of Paleontology 

v. 2, no. 1, p. 70-85, 3 pis., 2 figs.

Ishii, Ken-ichi, 1958, On the phylogeny, morphology and distribution 

of Fusulina, Beedeina and allied fusulinid genera: Osaka City 

University, Institute of Polytchnics Journal, ser. G., Geoscience, v. 

4, p. 29-70, 4 pis., 5 figs.

Kehn, T.M., Beard, J.G., and Williamson, A.D., 1982, Mauzy Formation, a new stratigraphc unit of Permian age in western Kentucky: U.S. Geological Survey Bulletin, 1529-H, p. H73-H86, 3 

figs.

Meek, F.B., and Worthen, A.H., 1873, Paleontology [of Illinois]. 

Descriptions of invertebrates from Carboniferous System: Illinois 

Geological Survey, v. 5 (Geology and palaeontology), pt. 2, p. 

321-619, 32 pis.

Merchant, F.E., and Keroher, R.P., 1939, Some fusulinids from the 

Missouri series of Kansas: Journal of Paleontology, v. 13, no. 6, p. 

594-614, 1 pi., 10 figs.

Morse, W.C., 1931, Pennsylvanian invertebrate fauna [of Kentucky]: 

Kentucky Geological Survey, ser. 6, v. 36, p. 293-348, 10 pis, 2 

figs.

Needham, C.E., 1937, Some New Mexico Fusulinidae: New Mexico 

School of Mines Bulletin 14, 88 p., 13 pis., 1 fig.

Phalen, W.C., 1912, Description of the Kenova quadrangle [KentuckyWest Virginia-Ohio]: U.S. Geological Survey Geologic Atlas, Folio 

184, 16 p.

Ping, R.G., and Rice, C.L., 1979,Stoney Fork Member (new name) of 

the Breathitt Formation in southeasternmost Kentucky, in Sohl, 

N.F., and Wright, W.B., Changes in stratigraphic nomenclature by 

the U.S. Geological Survey, 1978: U.S. Geological Survey Bulletin 

1482-A, p. A70-A76.
REFERENCES CITED 95

Rice, C.L., Kehn, T.M., and Douglass, R.C., 1979, Pennsylvanian 

correlations between the Eastern Interior and Appalachian 

Basins, in Palmer, J.E., and Butcher, R.R., eds., Depositional and 

structural history of the Pennsylvanian Systems of the Illinois 

Basin. Part 2, Invited papers-Field trip 9, Ninth International 

Congress of Carboniferous Stratigraphy and Geology: Illinois 

State Geological Survey [Guidebook Series, no. 15], p. 103-105, 2 

figs.

Rogers, H.D., 1858, The geology of Pennsylvania; a government survey: Philadelphia, J.B. Lippincott, 2 v. in 3. (See v. 2, pt. 1)

Ross, C.A., 1963, Standard Wolfcampian series (Permian), Glass 

Mountains, Texas: Geological Society of America Memoir, 88, 205 

p., 29 pis., 11 figs.

Sabins, F.F., Jr., and Ross, C.A., 1963, Late Pennsylvanian-Early 

Permian fusulinids from southeast Arizona: Journal of Paleontology, v. 37, no. 2, p. 323-365, 6 pis., 4 figs.

St. Jean, Joseph, Jr., 1957, A Middle Pennsylvanian foraminiferal 

fauna from Dubois County, Indiana: Indiana Geological Survey 

Bulletin 10, 66 p., 5 pis., 3 figs.

Shaver, R.H., and Smith, S.G., 1974, Some Pennsylvanian kirkbyacean 

ostracods of Indiana and Midcontinent Series terminology: Indiana 

Geological Survey Report of Progress 31, 59 p., 3 pis., 5 figs.

Skinner, J.W., 1931, Primitive fusulinids of the Mid-Continent region: 

Journal of Paleontology, v. 5, no. 3, p. 253-259, 1 pi., 1 fig.

Skinner, J.W., and Wilde, G.L., 1967, Eowaeringella, new generic 

designation for fusulinids of the groups of Wedekindellina 

ultimata, Newell and Keroher: Journal of Paleontology, v. 41, no. 

4, p. 1004-1005.

Slade, M.L., 1961, Pennsylvanian and Permian fusulinids of the 

Ferguson Mountain area, Elko County, Nevada: Brigham Young 

University Geology Studies, v. 8, p. 55-92, 10 pis., 2 figs.

Smyth, Pauline, 1957, Fusulinids from the Pennsylvanian rocks of 

Ohio: Ohio Journal of Science, v. 57, no. 5, p. 257-283, 3 pis., 4 figs.

_____1974, Fusulinids in the Appalachian Basin: Journal of Paleontology, v. 48, no. 4, p. 856-858, 2 figs.

Staff, Hans von, 1912, Monographic de Fusulinen (Geplant und begonnen von E. Schellwien). Teil 3-Die Fusulinen (Schellwienien) 

Nordamerikas: Palaeontographica, v. 59, Lieferung 3 and 4, p. 

157-192, 6 pis., 17 figs.

Sutherland, P.K., ed., 1982, Lower and Middle Pennsylvanian 

stratigraphy in south-central Oklahoma: Oklahoma Geological 

Survey Guidebook 20, 44 p., 58 figs.

Thomas, N.L., 1931, New early fusulinids from Texas: Texas University Bulletin 3101, p. 27-33, 1 pi., 1 fig.

Thompson, M.L., 1934, The fusulinids of the Des Moines Series of 

Iowa: Iowa University Studies in Natural History, v. 16, no. 4, p. 

274-332, 4 pis.

_____1935, Fusulinids from the lower Pennsylvanian Atoka and 

Boggy formations of Oklahoma: Journal of Paleontology, v. 9, no. 

4, p. 291-306, 1 pi.

_____1936, Pennsylvanian fusulinids from Ohio: Journal of Paleontology, v. 10, no. 8, p. 673-683, 2 pis. 

___1942, New genera of Pennsylvanian fusulinids: American

Journal of Science, v. 240, no. 6, p. 403-420, 3 pis.

.1948, Studies of American fusulinids: Kansas University

Paleontological Contributions, Article 4 (Protozoa 1), 184 p., 38 

pis., 8 figs. 

_____1953, Primitive Fusulinella from southern Missouri:

Journal of Paleontology, v. 27, no. 3, p. 321-327, 2 pis.; correction, 

no. 5, p. 748.

.1954, American Wolfcampian fusulinids: Kansas University Paleontological Contributions, Article 14 (Protozoa 5), 225 p., 

52 pis., 14 figs. 

___1957, Northern Midcontinent Missourian fusulinids: Journal

of Paleontology v. 31, no. 2, p. 289-328, 10 pis., 2 figs.

Thompson, M.L., and Shaver, R.H., 1964, Early Pennsylvanian microfaunas of the Illinois Basin: Illinois State Academy of Science 

Transactions, v. 57, no. 1, p. 3-23, 1 pi., 1 fig.

Thompson, M.L., Shaver, R.H., and Riggs, E.A., 1959, Early Pennsylvanian fusulinids and ostracods of the Illinois Basin: Journal of 

Paleontology, v. 33, no. 5, p. 770-792, 4 pis., 2 figs.

Thompson, M.L., Verville, G.J., andBissell, H.J., 1950, Pennsylvanian 

fusulinids of the south-central Wasatch Mountains, Utah: Journal 

of Paleontology, v. 24, no. 4, p. 430-465, 8 pis., 2 figs.

Wanless, H.R., 1975, Illinois Basin region, in McKee, E.D., and others, 

Paleotectonic investigations of the Pennsylvanian System in the 

United States: U.S. Geological Survey Professional Paper 853, pt. 

1, p. 71-95, 3 figs.

PLATES 1-20

Contact photographs of the plates in this report are available, at cost, from 

U.S. Geological Survey Library, Federal Center, Denver, Colorado 80225
PLATE 1

Figures 1-31. Profusulinella kentuckyemis Thompson and Riggs, 1959 (p. 26) 

1-12. Refigured types from surface samples.

1-3. Holotype, X20, X50, and X100.

Specimen shows twisting of coiling axis through first volutions and traces of inner tectorium in fourth 

and fifth volutions. ISGS (Illinois State Geological Survey) specimen 13P-1.

4. Axial section, X20. Axis does not rotate in this specimen. Paratype, ISGS 13P-6.

5. Axial section, X20. Small paratype, ISGS 13P-3. 

6-7. Equatorial section, X20 and X50. Paratype, ISGS 13P-14. 

8-9. Equatorial sections, X20, X50, and X100. Paratype, ISGS 13P-12. 

10-12. Equatorial section, X20, X50, and X100. This specimen shows some fibrous structure in diaphanotheca

in outer volutions. Paratype, ISGS 13P-13. 

13-31. Specimens from core samples.

13-14. Axial section, X20 and X50, from locality 31, sample f!3740B-4, at depth of 181.9 ft. Hypotype,

USNM 368648. 

15-16. Axial section, X50 and X20, from locality 31, sample f!3740A-l, at depth of 184.2 ft. Hypotype,

USNM 368649.

17-18. Axial section, X50 and X20, from locality 31, sample f!3740A-3, at depth of 184.2 ft. Hypotype, USNM 

368650.

19. Axial section from locality 13, sample f!3923-l, at depth of 46.3 ft. Hypotype, USNM 368651.

20. Axial section, X20, from same locality as fig. 19. Hypotype, USNM 368652.

21. Equatorial section, X20, from locality 11, sample f!3908-4, at depth of 120 ft. Hypotype, USNM 

368653.

22. Axial section, X20, from same locality as fig. 21. Hypotype, USNM 368654. 

23-26. Equatorial and axial sections, X20, from same locality as fig. 21 but from depth of 122.6 ft. Sample

f!3909. Hypotypes, USNM 368655-368658.

27. Axial section, X20, from locality 13, sample f!3924-2, at depth of 76.5 ft. Hypotype, USNM 368659. 

28, 29. Equatorial and axial sections, X20, from locality 37, sample f!3746, at depth of 168 ft. Hypotypes,

USNM 368660, 368661.

30, 31. Axial section, X20 and X50, from locality 11, sample f!3911-2, at depth of 148.5 ft. Hypotype, USNM 

368662.
DEPARTMENT OF THE INTERIOR 

U.S. GEOLOGICAL SURVEY

PROFESSIONAL PAPER 1451 

PLATE 1

31
PLATE 2

[All specimens illustrated are from locality 67, collection f!4509, Boggs Limestone Member of Pottsville Formation]

Figures 1-12. Profusulinella ohioensis Douglass n. sp. (p. 26).

1, 2. Axial section, X20 and X50. Paratype, USNM 368663.

3. 11. Equatorial section, X50 and X20. Paratype, USNM 368667.

4. 12. Equatorial section, X50 and X20. Paratype, USNM 368668.

5, 6. Equatorial section, X20 and X50. Paratype, USNM 368664.

7, 8. Axial section, X20 and X50. Paratype, USNM, 368665.

9, 10. Axial section, X50 and X20. Holotype, USNM, 368666. 

13-24. Fusulinella imprima Douglass n. sp. (p. 28)

13, 14. Equatorial section, X20 and X50. Paratype, USNM 368669.

15. Axial section, X20. Paratype, USNM 368670.

16. Axial section, X20. Paratype, USNM 368671.

17. Equatorial section, X20. Paratype, USNM 368672.

18. Axial section, X20. Paratype, USNM 368673.

19, 20. Axial section, X20 and X50. Holotype, USNM 368674.

21. Axial section, X20. Paratype, USNM 368675.

22. Axial section, X20, showing septal pores exceptionally well in outer volution. Paratype USNM 368676.

23, 24. Axial section, X20 and X50. Paratype, USNM 368677.
DEPARTMENT OF THE INTERIOR 

U.S. GEOLOGICAL SURVEY

PROFESSIONAL PAPER 1451 

PLATE 2

24

PROFUSULINELLA, FUSULINELLA
PLATE 3

[All specimens illustrated are from locality 21, collection f!3927, at depth of 471.5-471.7 ft]

Figures 1-23. Fusulinella primotina Douglass n. sp. (p. 29).

1. Axial section, X20. Paratype, USNM 368678. 

2, 3. Equatorial section, X20 and X50. Paratype, USNM 368679. 

4, 5. Equatorial section, X20 and X50. Paratype, USNM 368680.

6. Axial section, X20. Paratype, USNM 368681.

7. Axial section, X20. Paratype, USNM 368682. 

8, 9. Axial section, X20 and X50. Paratype, USNM 368683.

10. Equatorial section, X20. Paratype, USNM 368684.

11. Axial section, X20. Paratype, USNM 368685. 

12, 13. Equatorial section, X20 and X50, showing some development of inner tectorium in outer volutions. Paratype, USNM

368686.

14, 15. Axial section, X20 and X50. Paratype, USNM 368687. 

16, 17. Axial section, X20 and X50. Paratype, USNM 368688. 

18, 19. Axial section, X50 and X20, Paratype, USNM 368689. 

20, 21. Axial section, X50 and X20. Holotype, USNM 368690. 

22, 23. Axial section, X20 and X50. Paratype, USNM 368691.
DEPARTMENT OF THE INTERIOR 

U.S. GEOLOGICAL SURVEY

PROFESSIONAL PAPER 1451 

PLATE 3

23

FUSULINELLA
PLATE 4

[All specimens illustrated are from locality 46, from samples through 30 ft of Stoney Fork Member of Breathitt Formation. Samples are in

stratigraphic order from collection f!3788 below to f!3794 at top]

Figures 1-18. Fusidinella pria Douglass n. sp. (p. 31).

1. Axial section, X20, sample f!3793-l. Paratype, USNM 368692.

2. Equatorial section, X20, sample f!3793-7. Paratype, USNM 368693.

3. Axial section, X20, sample f!3794-l. Paratype, USNM 368694.

4. Axial section, X20, sample f!3792-l. Paratype USNM 368695.

5. Axial section, X20, sample f!3792-2. Paratype, USNM 368696.

6. Equatorial section, X20, sample f!3792-10. Paratype, USNM 368697.

7. Axial section, X20, sample f!3791-l. Paratype, USNM 368698.

8. Equatorial section, X20, sample f!3791-4. Paratype, USNM 368699. 

9, 11. Equatorial section, X20 and X50, sample f!3789-6. Paratype, USNM 368700. 

10. Equatorial section, X20, sample f!3790-6. Paratype, USNM 368701.

12. Axial section, X20, sample f!3790-2. Paratype USNM 368702.

13. Axial section, X20, sample f!3789-l. Paratype USNM 368703. 

14, 16. Axial section, X20 and X50, sample f!3788-l. Holotype, USNM 368704. 

15. Equatorial section, X20, sample f!3788-5. Paratype, USNM 368705.

17. Axial section, X20, sample f!3788-2. Paratype, USNM 368706.

18. Equatorial section, X20, sample f!3788-7. Paratype, USNM 368707.
DEPARTMENT OF THE INTERIOR 

U.S GEOLOGICAL SURVEY

PROFESSIONAL PAPER 1451 

PLATE 4

18 

FUSULINELLA
PLATE 5

[All specimens illustrated are from the type Curlew Limestone Member of the Tradewater Formation at locality 17, but in stratigraphic order 

from top to bottom of the unit shown from top to bottom of the plate. All figures X20]

Figures 1-26. Fiisulinella stouti Thompson, 1936 (p. 33).

1-5. Specimens from sample f!3899 from upper beds on Indian Hill, showing tighter coiling and more fluting than most of 

the specimens from lower beds.

1. Axial section, f!3899-5. Hypotype, USNM 369708.

2. Axial section, f!3899-ll. Hypotype, USNM 368709.

3. Equatorial section, f!3899-14. Hypotype.USNM 368710.

4. Axial section, f!3899-l. Hypotype, USNM 368711.

5. Axial section, f!3899-10. Hypotype, USNM 368712. 

6-15. Specimens from sample f!3898 from 3 ft limestone considered principal bed of Curlew.

6. Equatorial section, f!3898-ll. Hypotype, USNM 368713.

7. Equatorial section, f!3898-23. Hypotype, USNM 368714.

8. Axial section, f!3898-8. Hypotype, USNM 368715.

9. Axial section, f!3898-15. Hypotype, USNM 368716.

10. Equatorial section, f!3898-40. Hypotype, USNM 368717.

11. Axial section, f!3989-19. Hypotype, USNM 368718.

12. Axial section, f!3898-28. Hypotype, USNM 368719.

13. Axial sectioin, f!3898-27. Hypotype, USNM 368720.

14. Equaorial section, f!3898-10. Hypotype, USNM 368721.

15. Axial section, f!3898-2. Hypotype, USNM 368722. 

16-20. Specimens from sample f 13629 from the 3 ft limestone.

16. Axial section, f!3629-12. Hypotype USNM 368723.

17. Axial section, f!3629-l. Hypotype, USNM 368724.

18. Axial section, f!3629-17. Hypotype, USNM 368725.

19. Equatorial section, fL3629-18. Hypotype, USNM 368726.

20. Axial section, f!3629-16. Hypotype, USNM 368727. 

21-26. Specimens from sample f 13897 from flaggy limestone, 6-8 in. below the 3-ft limestone.

21. Axial section, f!3897-l. Hypotype, USNM 368728.

22. Axial section, f!3897-3. Hypotype, USNM 368729.

23. Equatorial section, f!3897-7. Hypotype, USNM 368730.

24. Axial section, f 13897-2. Hypotype, USNM 368731.

25. Axial section, f!3897-3. Hypotype, USNM 368732.

26. Equatorial section, f!3897-5. Hypotype USNM 368733.
DEPARTMENT OF THE INTERIOR 

U.S. GEOLOGICAL SURVEY

PROFESSIONAL PAPER 1451 

PLATE 5

FUSULINELLA
PLATE 6

[All figures X20]

Figures 1-16. Fusulinella iowensis Thompson, 1934 (p. 34) (See also pi. 15).

1-7. Reillustration of specimens illustrated by Thompson and Riggs (in Thompson, Shaver, and Riggs, 1959) from type 

Curlew Limestone Member of Tradewater Formation.

1. Axial section. Hypotype, ISGS (Illinois State Geological Survey) 13P-21.

2. Axial section. Hypotype, ISGS 13P-20.

3. Equatorial section. Hypotype, ISGS 13P-16.

4. Axial section. Hypotype, ISGS 13P-22.

5. Axial section. Hypotype, ISGS 13P-25.

6. Equatorial section. Hypotype, ISGS 13P-19.

7. Equatorial section. Hypotype, ISGS 13P-18. 

8-11, 15. Specimens from Curlew equivalent at locality 31, sample f!3741B, at depth of 74.5 ft.

8. Axial section. Hypotype, USNM 368734.

9. Axial section. Hypotype, USNM 368735.

10. Axial section. Hypotype, USNM 368736.

11. Equatorial section. Hypotype, USNM 368737. 

15. Equatorial section. Hypotype, USNM 368741. 

12-14. Specimens from Curlew equivalent at locality 21, sample f!3926, at depth of 469 ft.

12. Axial section. Hypotype, USNM 368738.

13. Axial section. Hypotype, USNM 368739.

14. Equatorial section. Hypotype, USNM 368740.

16. Axial section from locality 32, sample f!3635-l. Hypotype, USNM 368745. 

17-22. FusuLinella sp. (p. 35).

17-19. Specimens from locality 32, sample f!3635.

17. Axial section. USNM 368742. 

18, 19. Equatorial sections. USNM 368743, 368744.

20. Equatorial section from locality 21, sample f!3926-5, from depth of 469 ft. USNM 368746. 

21, 22. Specimens from locality 31, sample f!3741B, at 74.5 ft.

21. Axial section. USNM 368747.

22. Equatorial section. USNM 368748.
DEPARTMENT OF THE INTERIOR 

U.S GEOLOGICAL SURVEY

PROFESSIONAL PAPER 1451 

PLATE 6

19

FUSULINELLA
PLATE 7

[All figures X20]

Figures 1-42. Wedekindellina euthysepta (Henbest), 1928 (p. 36) (see also pi. 16).

Surface samples

1-4, 6-8. Specimens from locality 39, sample f!3906, from Yeargins limestone member (informal usage) of 

Tradewater Formation, Butler County, Ky.

1-4. Axial sections. Hypotypes, USNM 368749-368752.

6. Axial section. Hypotype, USNM 368754.

7. Equatorial section. Hyopotype, USNM 368755.

8. Axial section. Hypotype, USNM 368756.

5, 10, 11. Specimens from locality 39, sample f!3905, from 3.5 ft below f!3906. 

5. Axial section. Hypotype, USNM 368753.

10. Equatorial section. Hypotype, USNM 368758.

11. Axial section. Hypotype, USNM 368759. 

9, 13, 14. Specimens from locality 41, sample f!3902, Yeargins limestone member in Ohio County, Ky. Axial sections.

Hypotypes, USNM 368757, 368761, 368762.

12, 16, 18, 19. Specimens from locality 32, sample f!3636, Yeargins limestone member in Christian County, Ky. 

12, 16. Axial sections. Hypotypes, USNM 368760, 368764. 

18, 19. Equatorial sections. Hypotypes, USNM 368766, 36867.

Subsurface samples

15, 17. Specimens from locality 18, sample f!3745, at depth of 265 ft, near top of Yeargins limestone member. 

15. Axial section. Hypotype.USNM 368763. 

17. Equatorial section. Hypotype, USNM 368765.

20-22. Specimens from locality 31, sample f!3742, at depth of 4.1-4.3 ft in Yeargins limestone member. Axial sections. Hypotypes, USNM 368768, 368770.

23-27, 31, 33, 34. Specimens from locality 18, sample f!3744, at depth of 267 ft, near base of Yeargins limestone member. 

23, 24. Axial sections. Hypotypes, USNM 368771, 368772.

25. Equatorial section. Hypotype, USNM 368773. 

26, 27. Axial sections. Hypotype, USNM 368774, 368775. 

31, 33, 34. Equatorial sections. Hypotypes, USNM 368779, 368781, 368782. 

28-30, 32, 

35, 36, 38, 

39, 41, 42. Specimens from locality 40, sample f!3752, from Yeargins limestone member at depths of 313-316 ft.

28, 29. Axial sections of small specimens. Hypotypes, USNM 368776, 368777. 

30, 32, 35, 36. Equatorial sections. Hypotypes, USNM 368778, 368780, 368783, 368784. 

38, 39. Axial sections. Hypotypes, USNM 368786, 368787.

41. Axial section. Hypotype, USNM 368789.

42. Equatorial section. Hyoptype, USNM 368790.

37. Specimen from locality 21, sample f!3925, at depth of 398 ft, in Yeargins limestone member. Axial section. 

Hypotype, USNM 368785.

40. Tangential section from locality 31, sample f!3741, at depth of 37 ft in Yeargins limestone member. Hypotype, USNM 368788.
DEPARTMENT OF THE INTERIOR 

U.S. GEOLOGICAL SURVEY

PROFESSIONAL PAPER 1451 

PLATE 7

42

WEDEKINDELLINA
PLATE 8

[Beedeina from drill cores in Yeargins limestone member (informal usage) of Tradewater Formation, western Kentucky]

[All figures X20]

Figures 1-3, 5, 6. Beedeina sp. aff. B. novamexicana (Needham), 1937 (p. 38) (see also pis. 9, 10). 

From locality 18, sample f!3745, at depth of 265 ft.

1, 2, 5, 6. Axial sections showing varieties of shape. USNM 368791, 368792, 368795, 368796.

3. Equatorial section. USNM 368793.

4, 7-9. Beedeina leei (Skinner), 1931 (p. 39) (see also pis. 9, 10, 15, 17). 

From locality 18, sample f!3744, at depth of 267 ft.

4. Equatorial section. Hypotype, USNM 368794. 

7-9. Axial sections showing variation in shape.

Hypotypes, USNM 368797-368799.

10. 13, 15, 17. Beedeina sp. aff. B. leei (Skinner), 1931 (p. 40) (see also pi. 12).

10. Axial section from locality 21, sample f!3925-3, at depth of 398.1 ft. USNM 368800. 

13, 15, 17. Specimens from locality 31, sample f!3742, at depth of 4.1-4.3 ft. 

13. Axial section of microspheric specimen. USNM 368803. 

15. Equatorial section. USNM 368805. 

17. Axial section. UNMN 368807.

11. 12, 14, 16,

18-21. Beedeina sp. aff. B. spissiplicata (Dunbar and Henbest), 1942 (p. 41) (see also pi. 9).

11. Axial section from locality 35, sample f!3919-3, at depth of 177.6 ft. USNM 368801. 

12, 14. Axial sections from locality 33, sample f!3750, at depth of 150 ft. USNM 368802, 368804. 

16, 18-21. Specimens from locality 31, sample f!3741, at depth of 36-38 ft. 

16. Axial section. USNM 368806. 

18. Equatorial section. USNM 368808. 

19, 20. Axial sections. USNM 368809, 368810. 

21. Equatorial section. USNM 368811.
DEPARTMENT OF THE INTERIOR 

U.S. GEOLOGICAL SURVEY

PROFESSIONAL PAPER 1451 

PLATE 8

21
PLATE 9

[Beedeina from locality 40, representing depths from 185 through 332 ft, in drill core Gil No. 9]

[All figures X20]

Figures 1, 2. Beedeina leei (Skinner), 1931 (p. 39) (see also pis. 8, 10, 15, 17). 

Axial sections from sample f!3753 at depth of 185 ft.

Hypotypes, USNM 368812, 368813.

3-10. Beedeina sp. aff. B. novamexicana (Needham), 1937 (p. 38) (see also pis. 8, 10). 

Specimens from sample f!3752, at depth of 313-316 ft.

3. Large axial section resembling some of specimens illustrated from Stonefort Limestone Member of Spoon Formation 

in Illinois. USNM 368814.

4. Large equatorial section of similar form. USNM 368815.

5. Axial section of microspheric specimen. USNM 368816. 

6, 7. Axial sections of smaller megalospheric specimens. USNM 368817, 368818. 

8, 9. Equatorial sections. USNM 368819, 368820.

10. Axial section. USNM 368821.

11-16. Beedeina sp. aff. B spissiplicata (Dunbar and Henbest), 1942 (p. 41) (see also pi. 8). 

Specimens from sample f 13751, at depth of 332 ft. 

11-14. Axial sections showing variety of shapes. Figure 12 has double proloculus but otherwise is similar to specimen

illustrated by Dunbar and Henbest on pi. 7 as fig. 5. Specimens are UNSM 368822-368825. 

15, 16. Equatorial sections. USNM 368826, 368826A.
DEPARTMENT OF THE INTERIOR 

U.S. GEOLOGICAL SURVEY

PROFESSIONAL PAPER 1451 

PLATE 9

15

BEEDEINA
PLATE 10

[Beedeina from surface samples of Yeargins limestone member (informal usage) of Tradewater Formation, western Kentucky]

[All figures X20]

Figures 1-8, 10, 11. Beedeina sp. aff. B. novamexicana (Needham), 1937 (p. 38) (see also pis. 8, 9).

1-5. Axial sections from locality 39, sample f!3904. USNM 368827-368831. 

6-8, 10, 11. Specimens from locality 41, sample f!3902.

6. Axial section. USNM 368832.

7. Equatorial section. USNM 368833. 

8, 10, 11. Axial sections. USNM 368834, 368836, 368837. 

9, 12-17. Beedeina leei (Skinner), 1931 (p. 39) (see also pis. 8, 9, 15, 17). Specimens from locality 32, sample f!3636.

9, 12,15. Equatorial sections. USNM 368835, 368838, 368841. 

13, 14, 16, 17. Axial sections. USNM 368839, 368840, 368842, 368843.
DEPARTMENT OF THE INTERIOR 

U.S. GEOLOGICAL SURVEY

PROFESSIONAL PAPER 1451 

PLATE 10

17

BEEDEINA
PLATE 11

[All figures X20]

Figures 1-19. Beedeina ashlandensis Douglass n. sp. (p. 43).

1-9. Specimens from locality 61, sample f!4308, showing variation in shape. 

1, 2. Axial sections. Paratypes, USNM 368844, 368845.

3. Equatorial section. Paratype, USNM 368846. 

4, 5. Axial sections. Paratypes, USNM 368847, 368848.

6. Equatorial section. Paratype, USNM 368849. 

7, 8. Axial sections. Paratypes, USNM 368850, 368851.

9. Equatorial section. Paratype, USNM 368852. 

10, 16-19. Specimens from locality 62, sample f!3787.

10. Axial section. Paratype, USNM 368853.

16. Equatorial section. Paratype, USNM 368859.

17. Axial section. Paratype, USNM 368860.

18. Axial section. Holotype, USNM 368861.

19. Equatorial section. Paratype, USNM 368862. 

11-15. Specimens from locality 63, sample f 14307.

11. Axial section. Paratype, USNM 368862. 

12, 13. Equatorial sections. Paratypes, USNM 368855, 368856. 

14, 15. Axial sections. Paratypes, USNM 368857, 368858.
DEPARTMENT OF THE INTERIOR 

U.S. GEOLOGICAL SURVEY

PROFESSIONAL PAPER 1451 

PLATE 11

19

BEEDEINA
PLATE 12

[All figures X20]

Figures 1-7. Beedeina girtyi (Dunbar and Condra), 1927 (p. 45) (see also pi. 13).

1-4. Specimens from Providence Limestone Member of Sturgis Formation at locality 27, sample f!3785.

1. Axial section of unusually large specimen having 10 volutions. Hypotype, USNM 368863.

2. Equatorial section. Hypotype, USNM 368864.

3. Axial section. Hypotype, USNM 368865.

4. Axial section of microspheric specimen. Hypotype, USNM 368866.

5-7. Reillustration of specimens from Herrin Limestone Member of Carbondale Formation in Illinois, illustrated 

by Dunbar and Henbest (1942, pi. 11).

5. Axial section of specimen illustrated by Dunbar and Henbest as fig. 11.

6. Equatorial section of paratype illustrated by Dunbar and Henbest as fig. 15.

7. Axial section of specimen illustrated by Dunbar and Henbest as fig. 14. 

8-10. Beedeina sp. aff. B. leei (Skinner), 1931 (p. 40) (see also pi. 8). 

Specimens from locality 39, sample f!3906.

8. Axial section. USNM 368867.

9. Tangential section. USNM 368867A.

10. Axial section. USNM 368868.
DEPARTMENT OF THE INTERIOR 

U.S. GEOLOGICAL SURVEY

PROFESSIONAL PAPER 1451 

PLATE 12

10

BEEDEINA
PLATE 13

[Beedeina from Providence Limestone Member of Sturgis Formation at locality 29 in Muhlenberg County, Ky]

[All figures X20]

Figures 1-5, 7, 8. Beedeina sp. aff. B. haworthi (Beede), 1916 (p. 46). 

1, 2, 4. Specimens from sample f!3784.

1. Axial section of large elongate specimen. USNM 368869.

2. Equatorial section. USNM 368870. 

4. Axial section of more fusiform specimen. USNM 368872. 

3, 5, 7, 8. Specimens from sample f!3783.

3. Incomplete axial section. USNM 368871. 

5, 8. Axial sections of immature specimens. USNM 368873, 368876.

7. Equatorial section. USNM 368875.

6, 9, 10. Beedeina girtyi (Dunbar and Condra), 1927 (p. 45) (see also pi. 12). 

Specimens from sample f 13782.

6. Axial section of subrounded specimen. Hypotype, USNM 368874.

9. Axial section. Hypotype, USNM 368877.

10. Equatorial section. Hypotype, USNM 368878.
DEPARTMENT OF THE INTERIOR 

U.S. GEOLOGICAL SURVEY

PROFESSIONAL PAPER 1451 

PLATE 13

10

BEEDEINA
PLATE 14

[All figures X20]

Figures 1-7. Beedeina acme (Dunbar and Henbest), 1942 (p. 47).

Specimens from Madisonville Limestone Member of Sturgis Formation at locality 26, sample f 13872.

1. Axial section Hypotype, USNM 368879.

2. Equatorial section. Hypotype, USNM 368880. 

3-7. Axial sections, showing shape variation.

Hypotypes, USNM 368881-368885. 

8-12. Beedeina sp. aff. B. acme (Dunbar and Henbest), 1942 (p. 49).

Specimens from Providence Limestone Member of Sturgis Formation at locality 28, sample f!3874.

8. Axial section. USNM 368886. 

9, 10. Equatorial sections. USNM 368887, 368888. 

11, 12. Axial sections. USNM 368889, 368890.
DEPARTMENT OF THE INTERIOR 

U.S. GEOLOGICAL SURVEY

PROFESSIONAL PAPER 1451 

PLATE 14

BEEDEINA
PLATE 15

[Beedeina and Fusulinella from Ohio] 

[All figures X20]

Figures 1-6. Beedeina carmcmi (Thompson), 1936 (p. 50).

1-4. Specimens from locality 75, sample f9717, in Vanport Limestone Member of Allegheny Formation. 

1, 2. Axial section. Topotypes, USNM 368891, 368892. 

3, 4. Equatorial sections. Typotypes, USNM 368893, 368894. 

5, 6. Axial sections from locality 71, sample f!3819, in Vanport Limestone Member. Hypotypes, USNM 368895,

368896. 

7-13. Beedeina leei (Skinner), 1931 (p. 39) (see also pis. 8-10, 17).

All specimens from locality 79 in Putnam Hill Limestone Member of Allegheny Formation. 

7, 13. Specimens from sample f!3837 near top of unit.

7. Axial section. Hypotype, USNM 368897. 

13. Equatorial section. Hypotype, USNM 368903. 

8-12. Specimens from sample f!3841 in middle of unit.

8, 9. Axial sections. Hypotypes, USNM 368898, 368899. 

10-12. Equatorial sections. Hypotypes, USNM 368900-368902. 

14-26. Fusulinella iowensis Thompson, 1934 (p. 34) (see also pi. 6).

14-16. Specimens from locality 82, sample f9715, in upper Mercer Limestone Member of Pottsville Formation. 

14, 15. Axial sections. Hypotypes, USNM 368904, 368905.

16. Equatorial section. Hypotype, USNM 368906.

17-21. Specimens from locality 74 from Mercer Limestone Member at Blunt Run. 

17, 18, 20, 21. Specimens from sample f 13832 in upper limestone.

17. 21. Equatorial sections. Hypotypes, USNM 368907, 368911.

18. 20. Axial sections. Hypotypes, USNM 368908, 368910.

19. Axial section from sample f!3824. Hypotype, USNM 368909.

22. Axial section from locality 85, sample f!3886. Hypotype, USNM 368912.

23. Axial section from locality 74, sample f!3825. Hypotype, USNM 368913.

24, 25. Axial and equatorial sections from locality 76, sample f!3816. Hypotypes, USNM 368914, 368915. 

26. Equatorial section from locality 76, sample f!3815. Hypotype, USNM 368916.
DEPARTMENT OF THE INTERIOR 

U.S. GEOLOGICAL SURVEY

PROFESSIONAL PAPER 1451 

PLATE 15

26

BEEDEINA, FUSULINELLA
PLATE 16

[Triticites, Beedeina, and Wedekindellinafrom Ohio]

Figures 1-10. Triticites skinneri Thompson, 1936 (p. 52).

All topotypes from locality 69, sample f9720, in Ames Limestone Member of Conemaugh Formation. All figures X10. 

1-3, 7-9. Axial sections, showing variability of shape and coiling. USNM 368917-1368919, 368923-368925. 

4-6, 10. Equatorial sections. USNM 368920-368922, 368926. 

11-18. Triticites ricei Douglass n. sp. (p. 53)

All specimens from locality 66, sample f!4005, in Ames Limestone Member. All figures X10.

11. Axial section. Holotype, USNM 368927.

12. Axial section. Paratype, USNM 368928. 

13, 14, 17, 18. Equatorial sections. Paratypes, USNM 368929, 368930, 368933, 368934.

15, 16. Axial sections. Immature paratypes, USNM 368931, 368932. 

19-24. Triticites ohioensis Thompson, 1936 (p. 55) (see also pi. 18). All figures X10.

19. Axial section of specimen illustrated by Thompson (1936, pi. 91, fig. 1), here designated lectotype. S.U.I. 

(State University of Iowa) 1044B.

20. Axial section of paralectotype illustrated by Thompson (1936, pi. 91, fig. 2). S.U.I. 1044A. 

21-24. Specimens from locality 65, sample f!3997, in Cambridge Limestone Member of Conemaugh Formation. 

21, 22. Axial sections. Hypotypes, USNM 368935, 368936. 

23, 24. Equatorial sections. Hyoptypes, USNM 368937, 368938. 

25-28. Triticites smythi Douglass n. sp. (p. 57).

Specimens from locality 69, sample f9719, from Cambridge Limestone Member of Conemaugh Formation. All figures 

X20.

25. Axial section. Holotype, USNM 368939.

26. Axial section. Paratype, USNM 368940. 

27-28. Equatorial sections. Paratypes, USNM 368941, 368942. 

29-31. Beedeina henbesti Douglass n. sp. (p. 42).

Specimens from locality 83, sample f!3109, from Vanport Limestone Member of Allegheny Formation. All figures X20.

29. Axial section. Holotype, USNM 368943.

30. Axial section. Paratype, USNM 368944.

31. Equatorial section. Paratype, USNM 368945. 

32-35. Wedekindellina euthysepta (Henbest), 1928 (p. 36) (see also pi. 7).

Specimens from locality 83, sample f!3109, from Vanport Limestone Member. All figures X20.

32. Axial section. Hypotype, USNM 368946.

33. Tangential section. Hypotype, USNM 368947.

34. Axial section. Hypotype, USNM 368948.

35. Equatorial section. Hypotype, USNM 368949.
DEPARTMENT OF THE INTERIOR 

U.S. GEOLOGICAL SURVEY

PROFESSIONAL PAPER 1451 

PLATE 16

TRITICITES, BEEDEINA, WEDEKINDELLINA
PLATE 17

Fusulinids from western Pennsylvania

Figures 1-4. Triticites cuttomensis Dunbar and Condra, 1927 (p. 58).

Specimens from locality 88 at Brilliant Cut, Pittsburgh, Pa. All figures X10.

1, 2. Axial sections illustrated by Dunbar and Henbest, (1942, pi. 23, figs. 18, 17). YPM (Yale Peabody Museum)

16501. 

3, 4. Axial and tangential sections of two additional specimens collected by William Darrah. Hypotypes USNM

368950, 368951. 

5, 6, 16-23. Beedeina sp. (p. 51).

5, 6. Axial sections of probably immature specimens, X20, from locality 86, sample f!3884, in float pieces of

Vanport Limestone Member of Allegheny Formation. USNM 368952, 368953.

16-23. Specimens from locality 87, from the Mahoning Limestone of Rogers (1858), but representing Vanport 

Limestone Member of Allegheny Formation.

16, 17. Axial sections of probably immmature specimens, X20, from sample f 13881. USNM 368963,

368964.

18. Axial section, X20, from sample £9111. USNM 368965. 

19, 20. Axial section X20 and X50, sample £9111. USNM 368966.

21. Axial section of immature specimen, X20, sample £13881. USNM 368967. 

22, 23. Axial section, X20 and X50, sample £9111. USNM 368968. 

7-9. Wedekindellina sp. (p. 38).

Specimens from locality 86, sample f!3884, in float pieces of Vanport Limestone Member of Allegheny Formation. 

7, 9. Axial sections. USNM 368954, 368956.

8. Equatorial section. USNM 368955. 

10-15, 24, 25. Beedeina leei (Skinner), 1931 (p. 39) (see also pis. 8-10, 15).

Specimens from locality 87, sample f 13881, from the Mahoning Limestone of Rogers (1858), but representing Vanport 

Limestone member of Allegheny Formation. All figures X20. 

10-12. Axial sections. Hypotypes, USNM 368957-368959. 

13-15. Equatorial sections. Hypotypes, USNM 368960-368962. 

24, 25. Axial sections. Hypotypes, USNM 368969, 368970.
DEPARTMENT OF THE INTERIOR 

U.S. GEOLOGICAL SURVEY

PROFESSIONAL PAPER 1451 

PLATE 17

25

TRITICITES, BEEDEINA, WEDEKINDELLINA
PLATE 18

[All figures X20]

Figures 1-45. Triticites ohioensis Thompson, 1936 (p. 55) (see also pi. 16).

All specimens from Brush Creek Limestone Member of Conemaugh Formation, eastern Kentucky. 

1-14. Specimens from locality 59, Boyd County.

1. Axial section from sample f!3122.

Hypotype USNM 368971. 

2-6. Specimens from sample f!3120.

2, 3, 5. Axial sections. Hypotypes, USNM 368972, 368973, 368975. 

4, 6. Equatorial sections. Hypotypes, USNM 368974, 368976. 

7-14. Specimens from sample f24987.

7-10. Axial sections. Hypotypes USNM 368977-368980. 

11-14. Equatorial sections. Hypotypes, USNM 368981-368984. 

15-22. Specimens from locality 58, sample f!4309, Boyd County.

15. Equatorial section. Hyoptype, USNM 368985. 

16-20. Axial sections. Hyoptypes, USNM 368986-368990. 

21, 22. Equatorial sections. Hyoptypes, USNM 368991, 368992. 

23-31. Specimens from locality 56, Carter County.

23-27. Specimens from sample f!3991.

23, 24. Axial sections. Hypotypes, USNM 368993, 368994. 

25-27. Equatorial sections. Hypotypes, USNM 368995-368997. 

28-31. Specimens from sample f 13992.

28, 29, 31. Axial sections. Hypotypes, USNM 368998, 368999, 369001.

30. Equatorial section. Hypotype, USNM 369000. 

32-39. Specimens from locality 57, sample f!3584, Boyd County.

32-36. Axial sections. Hypotypes, USNM 369002-369006. 

37-39. Equatorial sections. Hypotypes, USNM 369007-369009. 

40-45. Specimens from locality 60, sample f!4312, Boyd County.

40-43. Axial sections. Hypotypes, USNM 369010-369013. 

44-45. Equatorial sections. Hypotypes, USNM 369014, 369015.
DEPARTMENT OF THE INTERIOR 

U.S. GEOLOGICAL SURVEY

PROFESSIONAL PAPER 1451 

PLATE 18

44
PLATE 19

[All figures X10, except figures 9 and 11, X50]

Figures 1-11. Triticites kehni Douglass n. sp. (p. 59).

All specimens from Carthage Limestone Member of Sturgis Formation. 

1-4. Specimens from locality 22, sample f!3984.

1, 2. Axial sections. Paratypes, USNM 369016, 369017.

3. Equatorial section. Paratype, USNM 369018.

4. Axial section. Paratype, USNM 369019. 

5-7. Specimens from locality 16, sample f 13982.

5. Axial section. Holotype, USNM 369020. 

6, 7. Equatorial sections. Paratypes, USNM 369021, 369022. 

8-11. Specimens from locality 22, sample f 13977.

8, 9. Axial section. Paratype, USNM 369023. 

10, 11. Axial section. Paratype, USNM 369024.
DEPARTMENT OF THE INTERIOR 

U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 1451 

PLATE 19

TRITICITES
PLATE 20

Triticites from Illinois and western Kentucky

Figures 1-3, 5, 6. Triticites callosm Dunbar and Henbest, 1942 (p. 60).

Specimens from locality 1, sample f 13995; topotypes from Greenup Limestone Member of Matoon Formation. All figures 

X10.

1-3. Axial sections. Topotypes, USNM 369025-369027. 

5, 6. Equatorial sections. Typotypes, USNM 369029, 369030. 

4, 7-9. Triticites mediocris Dunbar and Henbest, 1942 (p. 61). 

Specimens from locality 1. All figures X10.

4. Axial section from sample f!3995 at type locality. Topotype, USNM 369028. 

7. Axial section from sample f 14013. USNM 369031.

8, 9. Equatorial and axial sections from sample f!4012. Topotypes, USNM 369032, 369033. 

10-15. Triticites beardi Douglass n. sp. (p. 62).

All specimens from locality 19, sample f!3876, from unnamed bed of Early Permian age in the Mauzy Formation. 

10-11. Partial axial section, X20 and X10. Paratype, USNM 369034. 

12-13. Axial section, X20 and X10. Paratype, USNM 369035. 

14-15. Axial section, X10 and X20. Holotype, USNM 369036.
DEPARTMENT OF THE INTERIOR 

U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 1451 

PLATE 20

15

TRITICITES