NOAA Technical Memorandum NMFS-NWFSC-114

The 2003 to 2008 U.S. West Coast 

Bottom Trawl Surveys 

of Groundfish Resources

off Washington, Oregon, and California: 

Estimates of Distribution, Abundance, Length, 

and Age Composition

December 2011

U.S. DEPARTMENT OF COMMERCE

National Oceanic and Atmospheric Administration

National Marine Fisheries Service
NOAA Technical Memorandum

NMFS-NWFSC Series

The Northwest Fisheries Science Center of the National 

Marine Fisheries Service, NOAA, uses the NOAA Technical 

Memorandum NMFS-NWFSC series to issue scientific and 

technical publications. Manuscripts have been peer reviewed 

and edited. Documents published in this series may be cited 

in the scientific and technical literature.

The NMFS-NWFSC Technical Memorandum series of the 

Northwest Fisheries Science Center continues the NMFSF/NWC series established in 1970 by the Northwest & 

Alaska Fisheries Science Center, which has since been split 

into the Northwest Fisheries Science Center and the Alaska 

Fisheries Science Center. The NMFS-AFSC Technical 

Memorandum series is now used by the Alaska Fisheries 

Science Center.

Reference throughout this document to trade names does not 

imply endorsement by the National Marine Fisheries Service, 

NOAA.

This document should be referenced as follows:

Bradburn, M.J., A.A. Keller, and B.H. Horness. 2011. The 

2003 to 2008 U.S. West Coast bottom trawl surveys of 

groundfish resources off Washington, Oregon, and 

California: Estimates of distribution, abundance, length, and 

age composition. U.S. Dept. Commer., NOAA Tech. Memo. 

NMFS-NWFSC-114, 323 p.
NOAA Technical Memorandum NMFS-NWFSC-114

The 2003 to 2008 U.S. West Coast 

Bottom Trawl Surveys 

of Groundfish Resources

off Washington, Oregon, and California: 

Estimates of Distribution, Abundance, Length, 

and Age Composition

Mark J. Bradburn, Aimee A. Keller, and Beth H. Horness

Northwest Fisheries Science Center

Fishery Resource Analysis and Monitoring Division

2725 Montlake Boulevard East

Seattle, Washington 98112

December 2011

U.S. DEPARTMENT OF COMMERCE

National Oceanic and Atmospheric Administration

National Marine Fisheries Service
Most NOAA Technical Memorandums

NMFS-NWFSC are available at the 

Northwest Fisheries Science Center

Web site, http://www.nwfsc.noaa.gov

Copies are also available from the

National Technical Information Service

5285 Port Royal Road

Springfield, VA 22161

Phone orders: 1-800-553-6847

E-mail orders: orders@ntis.fedworld.gov

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iii

Table of Contents

List of Figures...............................................................................................................................................v

List of Tables.............................................................................................................................................xiii

Executive Summary....................................................................................................................................xv

Acknowledgments.....................................................................................................................................xvii

Introduction...................................................................................................................................................1

Methods and Results.....................................................................................................................................3

Survey Design...........................................................................................................................................3

Vessels......................................................................................................................................................4

Gear Description.......................................................................................................................................4

Trawling Protocol.....................................................................................................................................4

Net Performance and Trawl Analysis...........................................................................................................6

Sensors and Trawl Monitoring .................................................................................................................6

Sensor Data and Point Estimation ............................................................................................................6

Sensor Data Quality..................................................................................................................................7

Haul and Catch Data.................................................................................................................................8

Catch per Unit Effort and Biomass Estimates..........................................................................................8

Biological Data Collection .......................................................................................................................9

Length and Age Composition.................................................................................................................10

Weight-length Relationships ..................................................................................................................10

Fish Community Metrics........................................................................................................................11

Figures 1–235..............................................................................................................................................13

Tables 1–12...............................................................................................................................................244

References.................................................................................................................................................285

Appendix A: Primary and Alternative Station Selection for the Survey Cell Pool ..................................287

Primary Station Selection .....................................................................................................................288

Alternative Station Selection ................................................................................................................288

Search Times........................................................................................................................................288

Appendix B: Species Frequency of Occurrence, Depth Range, and Latitudinal Range ...........................289
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v

List of Figures

Figure 1. Total survey area and sampling stations during 2003–2008.......................................................14

Figure 2. Geographic extent of the surveys subdivided into INPFC areas................................................15

Figure 3. Schematic of Aberdeen-type sampling trawl used during the surveys.......................................16

Figure 4. Mean net width and height for trawls conducted as part of the surveys for years 2003–2008...17

Figure 5. Trends in total catch of 46 FMP species plus grooved tanner crab and giant grenadier in the 

surveys from 2003 to 2008 .........................................................................................................................18

Figure 6. Arrowtooth flounder distribution and relative abundance for 2003 and 2004 ...........................19

Figure 7. Arrowtooth flounder distribution and relative abundance for 2005 and 2006 ...........................20

Figure 8. Arrowtooth flounder distribution and relative abundance for 2007 and 2008 ...........................21

Figure 9. Aurora rockfish distribution and relative abundance for 2003 and 2004 ...................................22

Figure 10. Aurora rockfish distribution and relative abundance for 2005 and 2006 .................................23

Figure 11. Aurora rockfish distribution and relative abundance for 2007 and 2008 .................................24

Figure 12. Bank rockfish distribution and relative abundance for 2003 and 2004 ....................................25

Figure 13. Bank rockfish distribution and relative abundance for 2005 and 2006 ....................................26

Figure 14. Bank rockfish distribution and relative abundance for 2007 and 2008 ....................................27

Figure 15. Big skate distribution and relative abundance for 2003 and 2004............................................28

Figure 16. Big skate distribution and relative abundance for 2005 and 2006............................................29

Figure 17. Big skate distribution and relative abundance for 2007 and 2008............................................30

Figure 18. Blackgill rockfish distribution and relative abundance for 2003 and 2004..............................31

Figure 19. Blackgill rockfish distribution and relative abundance for 2005 and 2006..............................32

Figure 20. Blackgill rockfish distribution and relative abundance for 2007 and 2008..............................33

Figure 21. Bocaccio distribution and relative abundance for 2003 and 2004............................................34

Figure 22. Bocaccio distribution and relative abundance for 2005 and 2006............................................35

Figure 23. Bocaccio distribution and relative abundance for 2007 and 2008............................................36

Figure 24. Calico rockfish distribution and relative abundance for 2003 and 2004 ..................................37

Figure 25. Calico rockfish distribution and relative abundance for 2005 and 2006 ..................................38

Figure 26. Calico rockfish distribution and relative abundance for 2007 and 2008 ..................................39

Figure 27. California scorpionfish distribution and relative abundance for 2003 and 2004......................40

Figure 28. California scorpionfish distribution and relative abundance for 2005 and 2006......................41

Figure 29. California scorpionfish distribution and relative abundance for 2007 and 2008......................42
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Figure 30. California skate distribution and relative abundance for 2003 and 2004 .................................43

Figure 31. California skate distribution and relative abundance for 2005 and 2006 .................................44

Figure 32. California skate distribution and relative abundance for 2007 and 2008 .................................45

Figure 33. Canary rockfish distribution and relative abundance for 2003 and 2004.................................46

Figure 34. Canary rockfish distribution and relative abundance for 2005 and 2006.................................47

Figure 35. Canary rockfish distribution and relative abundance for 2007 and 2008.................................48

Figure 36. Chilipepper rockfish distribution and relative abundance for 2003 and 2004..........................49

Figure 37. Chilipepper rockfish distribution and relative abundance for 2005 and 2006..........................50

Figure 38. Chilipepper rockfish distribution and relative abundance for 2007 and 2008..........................51

Figure 39. Curlfin sole distribution and relative abundance for 2003 and 2004........................................52

Figure 40. Curlfin sole distribution and relative abundance for 2005 and 2006........................................53

Figure 41. Curlfin sole distribution and relative abundance for 2007 and 2008........................................54

Figure 42. Darkblotched rockfish distribution and relative abundance for 2003 and 2004.......................55

Figure 43. Darkblotched rockfish distribution and relative abundance for 2005 and 2006.......................56

Figure 44. Darkblotched rockfish distribution and relative abundance for 2007 and 2008.......................57

Figure 45. Dover sole distribution and relative abundance for 2003 and 2004 .........................................58

Figure 46. Dover sole distribution and relative abundance for 2005 and 2006 .........................................59

Figure 47. Dover sole distribution and relative abundance for 2007 and 2008 .........................................60

Figure 48. English sole distribution and relative abundance for 2003 and 2004.......................................61

Figure 49. English sole distribution and relative abundance for 2005 and 2006.......................................62

Figure 50. English sole distribution and relative abundance for 2007 and 2008.......................................63

Figure 51. Flathead sole distribution and relative abundance for 2003 and 2004 .....................................64

Figure 52. Flathead sole distribution and relative abundance for 2005 and 2006 .....................................65

Figure 53. Flathead sole distribution and relative abundance for 2007 and 2008 .....................................66

Figure 54. Giant grenadier distribution and relative abundance for 2003 and 2004..................................67

Figure 55. Giant grenadier distribution and relative abundance for 2005 and 2006..................................68

Figure 56. Giant grenadier distribution and relative abundance for 2007 and 2008..................................69

Figure 57. Greenspotted rockfish distribution and relative abundance for 2003 and 2004 .......................70

Figure 58. Greenspotted rockfish distribution and relative abundance for 2005 and 2006 .......................71

Figure 59. Greenspotted rockfish distribution and relative abundance for 2007 and 2008 .......................72

Figure 60. Greenstriped rockfish distribution and relative abundance for 2003 and 2004........................73

Figure 61. Greenstriped rockfish distribution and relative abundance for 2005 and 2006........................74

Figure 62. Greenstriped rockfish distribution and relative abundance for 2007 and 2008........................75

Figure 63. Grooved tanner crab distribution and relative abundance for 2003 and 2004..........................76

Figure 64. Grooved tanner crab distribution and relative abundance for 2005 and 2006..........................77
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Figure 65. Grooved tanner crab distribution and relative abundance for 2007 and 2008..........................78

Figure 66. Halfbanded rockfish distribution and relative abundance for 2003 and 2004..........................79

Figure 67. Halfbanded rockfish distribution and relative abundance for 2005 and 2006..........................80

Figure 68. Halfbanded rockfish distribution and relative abundance for 2007 and 2008..........................81

Figure 69. Lingcod distribution and relative abundance for 2003 and 2004 .............................................82

Figure 70. Lingcod distribution and relative abundance for 2005 and 2006 .............................................83

Figure 71. Lingcod distribution and relative abundance for 2007 and 2008 .............................................84

Figure 72. Longnose skate distribution and relative abundance for 2003 and 2004..................................85

Figure 73. Longnose skate distribution and relative abundance for 2005 and 2006..................................86

Figure 74. Longnose skate distribution and relative abundance for 2007 and 2008..................................87

Figure 75. Longspine thornyhead distribution and relative abundance for 2003 and 2004.......................88

Figure 76. Longspine thornyhead distribution and relative abundance for 2005 and 2006.......................89

Figure 77. Longspine thornyhead distribution and relative abundance for 2007 and 2008.......................90

Figure 78. Pacific cod distribution and relative abundance for 2003 and 2004.........................................91

Figure 79. Pacific cod distribution and relative abundance for 2005 and 2006.........................................92

Figure 80. Pacific cod distribution and relative abundance for 2007 and 2008.........................................93

Figure 81. Pacific flatnose distribution and relative abundance for 2003 and 2004..................................94

Figure 82. Pacific flatnose distribution and relative abundance for 2005 and 2006..................................95

Figure 83. Pacific flatnose distribution and relative abundance for 2007 and 2008..................................96

Figure 84. Pacific grenadier distribution and relative abundance for 2003 and 2004 ...............................97

Figure 85. Pacific grenadier distribution and relative abundance for 2005 and 2006 ...............................98

Figure 86. Pacific grenadier distribution and relative abundance for 2007 and 2008 ...............................99

Figure 87. Pacific hake distribution and relative abundance for 2003 and 2004.....................................100

Figure 88. Pacific hake distribution and relative abundance for 2005 and 2006.....................................101

Figure 89. Pacific hake distribution and relative abundance for 2007 and 2008.....................................102

Figure 90. Pacific ocean perch distribution and relative abundance for 2003 and 2004 .........................103

Figure 91. Pacific ocean perch distribution and relative abundance for 2005 and 2006 .........................104

Figure 92. Pacific ocean perch distribution and relative abundance for 2007 and 2008 .........................105

Figure 93. Pacific sanddab distribution and relative abundance for 2003 and 2004................................106

Figure 94. Pacific sanddab distribution and relative abundance for 2005 and 2006................................107

Figure 95. Pacific sanddab distribution and relative abundance for 2007 and 2008................................108

Figure 96. Petrale sole distribution and relative abundance for 2003 and 2004 ......................................109

Figure 97. Petrale sole distribution and relative abundance for 2005 and 2006 ......................................110

Figure 98. Petrale sole distribution and relative abundance for 2007 and 2008 ......................................111

Figure 99. Pygmy rockfish distribution and relative abundance for 2003 and 2004 ...............................112
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Figure 100. Pygmy rockfish distribution and relative abundance for 2005 and 2006 .............................113

Figure 101. Pygmy rockfish distribution and relative abundance for 2007 and 2008 .............................114

Figure 102. Redbanded rockfish distribution and relative abundance for 2003 and 2004.......................115

Figure 103. Redbanded rockfish distribution and relative abundance for 2005 and 2006.......................116

Figure 104. Redbanded rockfish distribution and relative abundance for 2007 and 2008.......................117

Figure 105. Redstripe rockfish distribution and relative abundance for 2003 and 2004 .........................118

Figure 106. Redstripe rockfish distribution and relative abundance for 2005 and 2006 .........................119

Figure 107. Redstripe rockfish distribution and relative abundance for 2007 and 2008 .........................120

Figure 108. Rex sole distribution and relative abundance for 2003 and 2004.........................................121

Figure 109. Rex sole distribution and relative abundance for 2005 and 2006.........................................122

Figure 110. Rex sole distribution and relative abundance for 2007 and 2008.........................................123

Figure 111. Rosethorn rockfish distribution and relative abundance for 2003 and 2004 ........................124

Figure 112. Rosethorn rockfish distribution and relative abundance for 2005 and 2006 ........................125

Figure 113. Rosethorn rockfish distribution and relative abundance for 2007 and 2008 ........................126

Figure 114. Rougheye rockfish distribution and relative abundance for 2003 and 2004 ........................127

Figure 115. Rougheye rockfish distribution and relative abundance for 2005 and 2006 ........................128

Figure 116. Rougheye rockfish distribution and relative abundance for 2007 and 2008 ........................129

Figure 117. Sablefish distribution and relative abundance for 2003 and 2004........................................130

Figure 118. Sablefish distribution and relative abundance for 2005 and 2006........................................131

Figure 119. Sablefish distribution and relative abundance for 2007 and 2008........................................132

Figure 120. Sharpchin rockfish distribution and relative abundance for 2003 and 2004 ........................133

Figure 121. Sharpchin rockfish distribution and relative abundance for 2005 and 2006 ........................134

Figure 122. Sharpchin rockfish distribution and relative abundance for 2007 and 2008 ........................135

Figure 123. Shortbelly rockfish distribution and relative abundance for 2003 and 2004........................136

Figure 124. Shortbelly rockfish distribution and relative abundance for 2005 and 2006........................137

Figure 125. Shortbelly rockfish distribution and relative abundance for 2007 and 2008........................138

Figure 126. Shortspine thornyhead distribution and relative abundance for 2003 and 2004...................139

Figure 127. Shortspine thornyhead distribution and relative abundance for 2005 and 2006...................140

Figure 128. Shortspine thornyhead distribution and relative abundance for 2007 and 2008...................141

Figure 129. Spiny dogfish distribution and relative abundance for 2003 and 2004 ................................142

Figure 130. Spiny dogfish distribution and relative abundance for 2005 and 2006 ................................143

Figure 131. Spiny dogfish distribution and relative abundance for 2007 and 2008 ................................144

Figure 132. Splitnose rockfish distribution and relative abundance for 2003 and 2004..........................145

Figure 133. Splitnose rockfish distribution and relative abundance for 2005 and 2006..........................146

Figure 134. Splitnose rockfish distribution and relative abundance for 2007 and 2008..........................147
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Figure 135. Spotted ratfish distribution and relative abundance for 2003 and 2004 ...............................148

Figure 136. Spotted ratfish distribution and relative abundance for 2005 and 2006 ...............................149

Figure 137. Spotted ratfish distribution and relative abundance for 2007 and 2008 ...............................150

Figure 138. Squarespot rockfish distribution and relative abundance for 2003 and 2004.......................151

Figure 139. Squarespot rockfish distribution and relative abundance for 2005 and 2006.......................152

Figure 140. Squarespot rockfish distribution and relative abundance for 2007 and 2008.......................153

Figure 141. Stripetail rockfish distribution and relative abundance for 2003 and 2004..........................154

Figure 142. Stripetail rockfish distribution and relative abundance for 2005 and 2006..........................155

Figure 143. Stripetail rockfish distribution and relative abundance for 2007 and 2008..........................156

Figure 144. Widow rockfish distribution and relative abundance for 2003 and 2004.............................157

Figure 145. Widow rockfish distribution and relative abundance for 2005 and 2006.............................158

Figure 146. Widow rockfish distribution and relative abundance for 2007 and 2008.............................159

Figure 147. Yellowtail rockfish distribution and relative abundance for 2003 and 2004........................160

Figure 148. Yellowtail rockfish distribution and relative abundance for 2005 and 2006........................161

Figure 149. Yellowtail rockfish distribution and relative abundance for 2007 and 2008........................162

Figure 150. Species specific CPUE by latitude for survey years 2003–2008 for 9 of 46 FMP species ..163

Figure 151. Species specific CPUE by latitude for survey years 2003–2008 for 9 of 46 FMP species ..164

Figure 152. Species specific CPUE by latitude for survey years 2003–2008 for 9 of 46 FMP species ..165

Figure 153. Species specific CPUE by latitude for survey years 2003–2008 for 9 of 46 FMP species ..166

Figure 154. Species specific CPUE by latitude for survey years 2003–2008 for 9 of 46 FMP species ..167

Figure 155. Species specific CPUE by latitude for survey years 2003–2008 for 1 of 46 FMP species 

plus grooved tanner crab and giant grenadier...........................................................................................168

Figure 156. Species specific biomass by year for 9 of 46 FMP species..................................................169

Figure 157. Species specific biomass by year for 9 of 46 FMP species..................................................170

Figure 158. Species specific biomass by year for 9 of 46 FMP species..................................................171

Figure 159. Species specific biomass by year for 9 of 46 FMP species..................................................172

Figure 160. Species specific biomass by year for 9 of 46 FMP species..................................................173

Figure 161. Species specific biomass by year for 1 of 46 FMP species plus grooved tanner crab and 

giant grenadier ..........................................................................................................................................174

Figure 162. Frequency of lengths by gender and stratum for arrowtooth flounder .................................175

Figure 163. Frequency of lengths by gender in stratum 2 for aurora rockfish.........................................176

Figure 164. Frequency of lengths by gender and stratum for bank rockfish ...........................................177

Figure 165. Frequency of lengths by gender and stratum for big skate...................................................178

Figure 166. Frequency of lengths by gender in stratum 2 for blackgill rockfish.....................................179

Figure 167. Frequency of lengths by gender and stratum for bocaccio ...................................................180
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Figure 168. Frequency of lengths by gender in stratum 1 for calico rockfish .........................................181

Figure 169. Frequency of lengths by gender in stratum 1 for California scorpionfish ............................182

Figure 170. Frequency of lengths by gender in stratum 1 for California skate........................................183

Figure 171. Frequency of lengths by gender in stratum 1 for canary rockfish ........................................184

Figure 172. Frequency of lengths by gender and stratum for chilipepper rockfish .................................185

Figure 173. Frequency of lengths by gender in stratum 1 for curlfin sole...............................................186

Figure 174. Frequency of lengths by gender and stratum for darkblotched rockfish ..............................187

Figure 175. Frequency of lengths by gender and stratum for Dover sole................................................188

Figure 176. Frequency of lengths by gender in stratum 1 for English sole .............................................189

Figure 177. Frequency of lengths by gender in stratum 1 for flathead sole.............................................190

Figure 178. Frequency of lengths by gender in stratum 1 for greenspotted rockfish...............................191

Figure 179. Frequency of lengths by gender and stratum for greenstriped rockfish................................192

Figure 180. Frequency of lengths by gender in stratum 1 for halfbanded rockfish .................................193

Figure 181. Frequency of lengths by gender and stratum for lingcod .....................................................194

Figure 182. Frequency of lengths by gender and stratum for longnose skate..........................................195

Figure 183. Frequency of lengths by gender and stratum for longspine thornyhead...............................196

Figure 184. Frequency of lengths by gender and stratum for Pacific cod ...............................................197

Figure 185. Frequency of lengths by gender and stratum for Pacific flatnose.........................................198

Figure 186. Frequency of lengths by gender in stratum 3 for Pacific grenadier......................................199

Figure 187. Frequency of lengths by gender and stratum for Pacific hake..............................................200

Figure 188. Frequency of lengths by gender and stratum for Pacific ocean perch..................................201

Figure 189. Frequency of lengths by gender in stratum 1 for Pacific sanddab........................................202

Figure 190. Frequency of lengths by gender in stratum 1 for petrale sole...............................................203

Figure 191. Frequency of lengths by gender in stratum 1 for pygmy rockfish........................................204

Figure 192. Frequency of lengths by gender and stratum for redbanded rockfish...................................205

Figure 193. Frequency of lengths by gender in stratum 1 for redstripe rockfish.....................................206

Figure 194. Frequency of lengths by gender and stratum for rex sole.....................................................207

Figure 195. Frequency of lengths by gender and stratum for rosethorn rockfish ....................................208

Figure 196. Frequency of lengths by gender and stratum for rougheye rockfish ....................................209

Figure 197. Frequency of lengths by gender and stratum for sablefish ...................................................210

Figure 198. Frequency of lengths by gender and stratum for sharpchin rockfish....................................211

Figure 199. Frequency of lengths by gender and stratum for shortbelly rockfish ...................................212

Figure 200. Frequency of lengths by gender and stratum for shortspine thornyhead..............................213

Figure 201. Frequency of lengths by gender and stratum for spiny dogfish............................................214

Figure 202. Frequency of lengths by gender and stratum for splitnose rockfish.....................................215
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Figure 203. Frequency of lengths by gender and stratum for spotted ratfish...........................................216

Figure 204. Frequency of lengths by gender in stratum 1 for squarespot rockfish..................................217

Figure 205. Frequency of lengths by gender and stratum for stripetail rockfish .....................................218

Figure 206. Frequency of lengths by gender and stratum for swordspine rockfish.................................219

Figure 207. Frequency of lengths by gender and stratum for widow rockfish ........................................220

Figure 208. Frequency of lengths by gender and stratum for yellowtail rockfish ...................................221

Figure 209. Age distribution for arrowtooth flounder from 2003 to 2006...............................................222

Figure 210. Age distribution for canary rockfish from 2003 to 2008......................................................223

Figure 211. Age distribution for chilipepper rockfish from the 2004 survey ..........................................224

Figure 212. Age distribution for darkblotched rockfish sampled from 2003 to 2008..............................225

Figure 213. Age distribution for Dover sole sampled from 2003 to 2004...............................................226

Figure 214. Age distribution for English sole sampled from 2003 to 2005.............................................227

Figure 215. Age distribution for greenstriped rockfish sampled from 2003 to 2008...............................228

Figure 216. Age distribution for lingcod sampled from 2003 to 2008 ....................................................229

Figure 217. Age distribution for Pacific ocean perch sampled from 2003 to 2008 .................................230

Figure 218. Age distribution for petrale sole sampled from 2003 to 2008 ..............................................231

Figure 219. Age distribution for sablefish sampled from 2003 to 2006 ..................................................232

Figure 220. Age distribution for splitnose rockfish sampled from 2003 to 2008 ....................................233

Figure 221. Age distribution for widow rockfish sampled from 2003 to 2008 .......................................234

Figure 222. Age distribution for yellowtail rockfish sampled during 2003 and 2004 .............................235

Figure 223. Weight-length relationship observed for Dover sole and petrale sole from 2003 to 2008...236

Figure 224. Weight-length relationship observed for sablefish and shortspine thornyhead from 2003 

to 2008 ......................................................................................................................................................237

Figure 225. Species richness per tow and per year by latitude for stratum 1 from 2003 to 2008............238

Figure 226. Species diversity by latitude for stratum 1 from 2003 to 2008.............................................238

Figure 227. Species eveness by latitude for stratum 1 from 2003 to 2008 ..............................................239

Figure 228. Species richness per tow and per year by latitude for stratum 2 from 2003 to 2008............239

Figure 229. Species diversity by latitude for stratum 2 from 2003 to 2008.............................................240

Figure 230. Species eveness by latitude for stratum 2 from 2003 to 2008 ..............................................240

Figure 231. Species richness per tow and per year by latitude for stratum 3 from 2003 to 2008............241

Figure 232. Species diversity by latitude for stratum 3 from 2003 to 2008.............................................241

Figure 233. Species eveness by latitude for stratum 3 from 2003 to 2008 ..............................................242

Figure 234. Patterns of variance of H′ in relation to latitude for each depth stratum for all survey 

years combined .........................................................................................................................................242

Figure 235. Relationship between species richness and diversity ...........................................................243
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List of Tables

Table 1. Coefficients for multiple linear regressions of net width as a function of net height and 

inverse scope for each fishing vessel by year ...........................................................................................244

Table 2. Area sampled and sampling density in each depth stratum for each INPFC statistical area 

based on successful tows during the surveys............................................................................................245

Table 3. Mean CPUE of 47 important FMP groundfish species plus grooved tanner crab caught by 

depth strata in the Vancouver INPFC area during the surveys.................................................................249

Table 4. Mean CPUE of 47 important FMP groundfish species plus grooved tanner crab caught by 

depth strata in the Columbia INPFC area during the surveys...................................................................254

Table 5. Mean CPUE of 47 important FMP groundfish species plus grooved tanner crab caught by 

depth strata in the Eureka INPFC area during the surveys .......................................................................259

Table 6. Mean CPUE of 47 important FMP groundfish species plus grooved tanner crab caught by 

depth strata in the Monterey INPFC area during the surveys...................................................................264

Table 7. Mean CPUE of 47 important FMP groundfish species plus grooved tanner crab caught by 

depth strata in the Conception INPFC area during the surveys................................................................269

Table 8. Number of individual length measurements and age structures collected by species during 

the surveys ................................................................................................................................................274

Table 9. Number of individual length measurements and age structures collected by species by year 

during the surveys.....................................................................................................................................275

Table 10. Number of length measurements collected by stratum for FMP species during the surveys ..278

Table 11. Weight-length relationships by year for the 2003–2008 surveys using a nonlinear least 

squares fit..................................................................................................................................................280

Table 12. Coefficients for linear regressions describing net width and height as a function of inverse 

scope for each fishing vessel, all years combined.....................................................................................284

Table B-1. Frequency of occurrence, depth range, and latitudinal range for fish species encountered 

during the surveys.....................................................................................................................................290

Table B-2. Frequency of occurrence, depth range, and latitudinal range for invertebrates encountered 

during the surveys.....................................................................................................................................305
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Executive Summary

The Northwest Fisheries Science Center’s Fishery Resource Analysis and Monitoring 

Division conducts an annual West Coast Groundfish Bottom Trawl Survey (WCGBTS) from 

May to October each year. The WCGBTS is a continuation of a number of transect-based 

surveys conducted between 1977 and 2002. This report summarizes the results of the annual 

WCGBTSs from 2003 to 2008.

WCGBTSs collects data on more than 90 species described in the Pacific Coast 

Groundfish Fishery Management Plan to fulfill the mandates included in the Magnuson-Stevens 

Fishery Conservation and Management Act. The surveys are designed to provide fisheriesindependent indices of stock abundance, which support stock assessment models for 

commercially and recreationally harvested groundfish species. The surveys also monitor longterm trends in distribution and abundance of West Coast groundfish species. Since 2003 the 

surveys have adopted a stratified-random sampling design and included the continental shelf 

(depth 55–183 m) and the continental slope (depth 184–1,280 m).

WCGBTSs have been conducted each year by chartered commercial fishing vessels from 

mid-May until late October. An Aberdeen-type net with a small mesh codend liner has been

used to sample fish biomass. Tows were made within randomly selected cells in three depth 

zones (55–183 m, 184–549 m, and 550–1,280 m) and five International North Pacific Fisheries 

Commission statistical areas. In 2004 the survey was subsequently grouped into two geographic 

strata, north and south of Point Conception, California.

From 2003 to 2008, 3,971 successful tows were completed along the U.S. Pacific Coast 

south of Cape Flattery (47°20′N), Washington, and north of the U.S.-Mexico border (32°40′N). 

During this time, 750,000 individual lengths and 136,000 age structures were collected from 

species of management concern. In this report, we document operations and results of the 2003–

2008 groundfish surveys with intent to summarize the data available for the creation of indices of 

abundance for stock assessment purposes.
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Acknowledgments

The West Coast Groundfish Bottom Trawl Survey is indebted to owners, captains, and 

crew of the fishing vessels B.J. Thomas, Blue Horizon, Captain Jack, Excalibur, Ms. Julie, 

Noah’s Ark, and Raven, as well as the numerous dedicated biologists who participated in the 

survey. Julia Clemons and Curt Whitmire created the GIS products for this report. Ian Taylor 

provided assistance with figure design.
xviii
Introduction

The U.S. West Coast groundfish fishery is supported by more than 90 commercially 

valuable species. It spans the area between Canada and Mexico along the U.S. Pacific Coast 

from nearshore to depths less than 2,000 m. Multiple vessel types, ranging in size from kayaks 

to trawlers, participate in the fishery. The fishery sectors deploy mobile and fixed gear including 

bottom trawls, midwater trawls, pots, longlines, and other hook and line gear; however, trawlers 

account for the majority of landed groundfish. Active management of the fishery began in the 

early 1980s with the establishment of optimum yields and trip limits for several managed 

species. Management measures currently include landings limits, size limits, gear restrictions, 

and time and area closures. The management measures are designed to avoid overfishing and 

rebuild overfished stocks.

The primary objective of the Northwest Fisheries Science Center’s (NWFSC) annual 

West Coast groundfish bottom trawl survey (WCGBTS) is to provide the fishery-independent 

data necessary to support the assessment of the status and trends of fish species inhabiting 

trawlable habitat along the upper continental slope (184–1,280 m) and shelf (55–183 m) of the 

U.S. West Coast. Coast-wide sampling cruises were undertaken by the NWFSC beginning in 

1998 to establish an ongoing time series of groundfish catch, fishing effort, and individual fish 

measurement data (Turk et al. 2001, Builder Ramsey et al. 2002, Keller et al. 2005, 2006a, 

2006b, 2007a, 2007b, 2008).

Although the NWFSC assumed responsibility for the slope portion of the groundfish 

survey starting in 1998, the time series began as the West Coast continental slope survey 

conducted by the Alaska Fisheries Science Center (AFSC) from 1984 to 2001. The AFSC slope 

survey was conducted triennially from 1984 until 1988, when it was conducted annually.

Beginning in 2003, the NWFSC expanded the depth coverage to include the continental 

shelf, replacing the continental shelf survey conducted triennially by the AFSC from 1977 to 

2001. Consequently, in the current sampling design, the WCGBTS now encompasses the area 

historically monitored by the continental shelf survey (1977–2001) and the continental slope 

survey (1984–2001) conducted by the AFSC. The NWFSC’s groundfish survey currently 

replaces the two AFSC time series and provides an annual snapshot of groundfish stock status.

Prior to 1998, surveys conducted by the AFSC were the principal source for fisheryindependent data about groundfish resources along the upper continental slope and shelf of the 

U.S. West Coast (Methot et al. 2000). The AFSC slope surveys were conducted with the NOAA 

research vessel Miller Freeman while the triennial survey used chartered Alaska fishing vessels. 

Both slope and shelf surveys used a transect-based sampling design until 2003. Spatial coverage 

of the West Coast surveys, as conducted by AFSC, varied between years due to constraints 

imposed by annual budget levels and availability of NOAA ship time (Lauth 2001).
2

Since their inception in 1998, the NWFSC groundfish surveys have used chartered 

fishing vessels from the West Coast commercial fishing industry. Chartered fishing vessels 

provide the surveys with the expertise of skilled fishing captains familiar with the challenges of 

fishing off the West Coast, and fulfill the cooperative research provisions of the MagnusonStevens Fishery Conservation and Management Act. The results of the surveys provide 

measures of change in relative abundance, distribution, and condition of groundfish stocks over 

time, which is of interest to fisheries managers, fishers, and concerned citizens. In this report, 

we document operations and results of the 2003–2008 groundfish surveys with intent to 

summarize the data available for the creation of indices of abundance for stock assessment 

purposes.
3

Methods and Results

Survey Design

The WCGBTS spans from 48°10′N to 32°30′N latitude and is geographically subdivided 

into the five International North Pacific Fisheries Commission (INPFC) statistical areas: U.S.-

Vancouver, Columbia, Eureka, Monterey, and Conception (Figure 1 and Figure 2). The survey 

area extends from northern Washington (U.S.-Canada border) to southern California (U.S.-

Mexico border) in waters ranging in depth from 55 to 1,280 m. The survey operations occur 

from mid-May to late October. The WCGBTS is conducted in two completed cycles of 

operations or passes. Four chartered West Coast bottom trawlers, two per pass, are used to 

conduct the survey. All vessels start operations from Newport, Oregon, heading north to Cape 

Flattery, Washington, and progress south along the coast, finishing south of San Diego, 

California.

Since 2003, the surveys have utilized a stratified random sampling design. The entire 

U.S. West Coast (55–1,280 m water depth) has been subdivided into 13,000 adjacent cells of 

equal area (1.5 nautical miles [nm] longitude by 2.0 nm latitude, Albers Equal Area projection) 

(Figure 1). The cells were stratified by geographic location and depth. The geographic 

allocation was based on a simple north-south division at lat 34°30′N (Point Conception, 

California), with 80% of effort in the northern portion of the survey and 20% in the southern 

range. The survey area was further divided into three depth strata (55–183 m, 184–549 m, and 

550–1,280 m). The percentage of sampling stations allocated to each depth range in the northern 

and southern geographic areas was based on the proportion of the area within each depth stratum. 

North of Point Conception, 40% of the area was located within the shallow depth stratum (55–

183 m) and 30% within each of the two remaining strata (mid depth and deep). South of Point 

Conception, 25% of the area was located within the shallow depth stratum, 45% within the mid

depth zone, and the remaining 30% within the deep zone.

Cells located on major shipping lanes, munition dump sites, and marine conservation 

areas were excluded. In addition, cells can be documented as untrawlable by survey personnel. 

When a cell cannot be successfully trawled because of rough bottom, shipwrecks, or other 

obstacles, it is designated untrawlable. A cell that has been designated untrawlable three times 

and has never been trawled successfully is also excluded.

Each year, the total number of sites targeted for the survey was apportioned across 

geographic area and depth categories according to the scheme described above. Primary stations 

were then drawn from the survey cell pool by strata, using a pseudorandom number generator. 

Each cell was sequentially assigned to an individual vessel. The process was repeated to identify 

two alternate sampling sites per location. Additional constraints were imposed to ensure that 

alternate sites were neither so close to an untrawlable primary site that they exhibited the same 

untrawlable features, nor were an impractical transit distance (see Appendix A for primary and 

alternative station selection). Each randomly selected cell was searched for trawlable ground. 
4

Once an acceptable site was found, a standard tow was made, targeted for 15-minutes bottom 

time.

Vessels

As a cooperative research effort, the WCGBTSs charter commercial trawlers to conduct 

survey operations. Chartered fishing vessels ranged in size from 19.8 to 28.0 m (65 to 92 feet) 

and in power from 450 to 1,200 horsepower. Each vessel was rigged as a stern trawler, with a 

rear gantry housing one or two net reels to set and retrieve trawl gear. Vessels were outfitted 

with split trawl winches and equipped with modern electronics including global positioning 

systems (GPS), multiple depth sounders, and radar. Prior to the start of the survey, NWFSC 

provided each vessel with two, 5⁄8′′ steel core trawl cables, each 2,288 m (1,250 fathoms) in 

length. Cables were measured side by side and marked at 25 fathoms increments while being 

spooled onto the vessel’s winches. The markings provided real-time verification of the release 

of equal warp length from both winches while setting a tow.

Gear Description

WCGBTS vessels were equipped with a standard four-panel, single-bridle, Aberdeentype trawl spread by 5 x 7 ft (1.5 x 2.1 m) steel V doors weighing 590 kg (Figure 3). The 

headrope and footrope measured 25.9 and 31.7 m, respectively. Each net had an additional liner 

(1.5 inch [3.81 cm] stretched measure, #24 twisted polypropylene) extending from the middle of 

the intermediate, through the codend, to retain smaller fish and invertebrates. The 85/104 

Aberdeen trawl nets were manufactured by Nor’Eastern Trawl Systems (Bainbridge Island, 

Washington) and certified by NWFSC personnel. Each vessel was outfitted with a primary and 

secondary net, and the NWFSC provided additional nets to vessels during survey if severe net 

damage occurred. To ensure continuity, fishing operations were conducted in compliance with 

national and regional protocols detailed in Stauffer (2004).

Trawling Protocol

Standard trawl operations are followed to minimize differences in sampling (fishing) 

efficiency across the range of conditions encountered during the survey. Trawling protocols are 

described in detail in Stauffer (2004). Trawling operations are limited to the daylight period. 

The initial tow each day begins (net on seafloor) following official sunrise, and the last tow of 

the day ends (net off seafloor) before official sunset.

Once a vessel is in the preselected sampling area (1.5 x 2.0 nm cell), the captain is 

instructed to observe the following search rules to identify a specific tow site: 1) search within 

the specified depth range, 2) remain fully within specified area, and 3) complete the search for 

trawlable ground within 1 hour. If no trawlable site is found within the 1-hour limit, the cell is 

designated as untrawlable and the vessel proceeds to the secondary cell. If the secondary cell 

also proves untrawlable, a search of the tertiary cell is attempted. If a tow is attempted but 

judged unsatisfactory or if the tow is aborted, an attempt is made to redo the tow before 

proceeding to secondary or tertiary cells. Rarely do the primary, secondary, and tertiary cells fail 

to produce an acceptable tow.
5

Vessel speed varies between 2.2 and 5 knots while the gear is set. When the net contacts 

the seafloor, the target towing speed is 2.2 ± 0.5 knots (nm hr–1) over ground. Two Wide Area 

Augmentation System (WAAS) GPS units (Garmin 152, Garmin International Inc., Olathe, 

Kansas) display speed instantaneously during a tow. Position, course over ground, and speed 

over ground are logged continuously. The captain, with oversight from the chief scientist, is 

responsible for maintaining the target speed during the tow. Tows are declared unsatisfactory 

and repeated if the target speed is not maintained throughout the tow.

The target tow duration is 15 minutes (15 minutes at 1.13 m sec–1 = 0.55 km). Tows 

begin when the footrope contacts the seafloor and end when the footrope lifts off the seafloor 

during retrieval. Initial bottom contact is determined in real time by a headrope height sensor, 

which detects the bottom as the net approaches. Once the footrope touches down, the vessel 

starts towing at the target speed.

A pair of bottom contact sensors (BCSs, AFSC, Seattle, Washington) are attached 4 feet 

from the center point of the footrope, on either side of the net. The BCS is a tilt meter housed in 

a heavy steel case that is vertical when the net is off the bottom and horizontal when the net is in 

contact with the bottom. Data from the BCSs are downloaded after the completion of each haul.
6

Net Performance and Trawl Analysis

Sensors and Trawl Monitoring

The performance and geometry of trawls are measured with Simrad Integrated Trawl 

Instrumentation System (ITI), Simrad PI44 Catch Monitoring System (Simrad Fisheries, 

Lynnwood, Washington), and BCSs. The Simrad ITI and PI44 systems are used to monitor and 

record net position and performance for each haul. Sensors mounted on the center of the 

headrope provide information on the vertical opening of the trawl, distance from the headrope to 

the seafloor, footrope clearance above the seafloor, ambient temperature, and depth. Paired 

spread sensors are attached on port and starboard wings to measure net width. The Simrad trawl 

instruments display gear performance in real time and provide georeferenced trawl positions 

relative to ship position, supplying means to track the trawl location along the seafloor 

throughout each tow. The ITI and PI44 data streams are monitored and continuously recorded 

during fishing operations.

Aspects of net performance of (e.g., spread between net wings and distance from the 

headrope to the footrope) are recorded using acoustic and bottom contact instruments attached to 

the net during each deployment. Adequate wingspread and door spread indicate that the net is 

making consistent bottom contact and that proper amount of scope was deployed. Extreme or 

prolonged periods of abnormal spread are indicative of net performance problems.

Sensor Data and Point Estimation

Upon completion of a trawl, the BCS data are downloaded. All features of the trawl 

event (from the commencement of net deployment to completion of net retrieval), including 

trawl mensuration data, are then synchronized based on time stamps.

BCS data indicate when the net landed on and lifted off bottom, and are used to 

determine tow duration and distance fished. The Simrad ITI and PI44 trawl instruments record 

net mensuration data and provide georeferenced trawl positions relative to ship position, 

supplying a means to track trawl location along the seafloor throughout each tow. Position data, 

collected at 2-second intervals, are used to monitor ground speed, track the vessel path, and 

estimate distance fished. Standard survey haul positions are estimated from GPS data—

generally the midpoint between the net touchdown and net liftoff positions. A Garmin WAAS 

GPS navigation unit is used to monitor towing speed during each haul. Average net speed over 

ground and distance fished is calculated from the position data and actual bottom time (Keller et 

al. 2008). Gear depth and bottom depth are estimated from Simrad ITI and PI44 sensor readings 

of headrope depth and headrope distance from bottom. Gear depth is provided by the headrope 

depth sensor and bottom depth was calculated as the sum of headrope depth and distance of the 

headrope from bottom.
7

Mean net widths (m) and distances fished (km) are calculated for each haul. When net 

mensuration data are available, the mean net width for each tow is calculated, excluding the 

initial and final 10% of the tow duration. The mean net width was 13.9 m, based on data from 

3,753 hauls that had available net mensuration measurements and exhibited good trawl 

performance. The mean net widths ranged from 9.8 to 17.6 m with a standard deviation of 0.96 

m (Figure 4). When the net mensuration instrumentation did not function properly, the mean of 

net width per tow was estimated using multiple linear regressions of net width as a function of 

net height and inverse scope for the individual chartered vessel (Table 1). Distances fished were 

calculated by estimating the length that the net traveled on the seafloor from the point where it 

touched down to the point where it lifted off (Wallace and West 2006).

Sensor Data Quality

Instrumentation played an important role in monitoring trawl performance, with 

mensuration data used to facilitate detection and correction of gear malfunction and identify 

deviation from standardized fishing procedures. In addition to their role in evaluating trawl 

performance, three sensors—BCS, ITI, and GPS—provided data used to estimate effort 

following the completion of the survey. Because of the occasional erratic readings inherent to 

acoustic data, sensor streams were reviewed prior to use.

Since sensor readings should be consistently present during a tow, recorded values of 

zero were treated as missing values and filtered prior to estimation of depth, net geometry, and 

temperature. Exclusion of extreme points was more difficult: large isolated spikes in depth, net 

geometry, and temperature readings were frequent and assumed to be the result of acoustic or 

electronic noise and were removed prior to processing. When multiple extreme points occurred 

in sequence, they were more difficult to evaluate because large swings in sensor data are 

expected during tows over sloped and irregular substrates. Trawl execution problems also 

produced data sets with large fluctuations in readings. Consequently, extreme values recorded 

where expected—either as part of a continuous variation in magnitude or during a particularly 

variable stretch of readings—were not excluded prior to analysis.

To ensure reliability of on-bottom readings, sensor data used to estimate depth, net width, 

and height were restricted to a subset of values collected from the center 80% of the tow 

duration. In the vast majority of tows, this criterion did not appreciably reduce the number of 

observations, but did effectively exclude small timing offsets between the BCS and ITI sensor 

systems and noise introduced by net touchdown and liftoff.

For some tows, only a few sensor readings (depth, net geometry, and temperature) fell 

within the estimation time interval. The extent to which these single or few point subsamples 

were representative of the entire tow was necessarily a subjective judgment. If the points seemed 

in alignment with the trajectory of points outside the subset time interval, they were used as the 

basis for estimation. Notations, hand recorded at sea during a tow, provided an additional level 

of data quality control. These notations were subsequently evaluated and potentially impacted 

the decision whether to accept or reject a tow.
8

Haul and Catch Data

The WCGBTSs have been progressively increasing the number of cells sampled since 

2003. In 2008 each vessel was assigned 188 randomly selected cells per vessel (752 total cells). 

Between 2003 and 2008, the survey sampled 3,791 cells. In a total area of 135,538 km2, the 

average sampling density per year is 4.66 hauls/1,000 km2. In the strata 1, 2, and 3, the average 

sampling densities are 7.22, 6.28, and 2.54 hauls/1,000 km2

, respectively. Table 2 describes the 

sampling effort by year, stratum, and INPFC area. The trend in total catch of 46 fisheries 

management plan (FMP) species plus grooved tanner crab (Chionoecetes tanneri) and giant 

grenadier (Albatrossia pectoralis) from 2003 to 2008 is negative (Figure 5).

Catch per Unit Effort and Biomass Estimates

Relative density was calculated as catch per unit effort (CPUE) for individual species in 

each INPFC area and depth stratum by dividing total catch weight (kg) per species by area swept 

(ha) per tow (j),

(1)

where CPUE is catch per unit effort in kg ha–1, Cj is catch per tow in kg for a given species, and 

aj is area swept (ha). Area swept was estimated by multiplying the mean net width for each tow 

by the distance fished. The distance fished was measured from the point the net touched down to 

the point the net lifted off the seafloor.

Mean estimates were initially calculated for each depth stratum within an INPFC area by 

averaging all tow-specific CPUE estimates, including those with zero CPUE, by species. Mean 

CPUE was calculated as:

(2)

where i is the INPFC-specific depth stratum and ni is the number of hauls within i.

The mean CPUE of 46 important FMP groundfish species plus grooved tanner crab and 

giant grenadier for each INFPC area and depth stratum is described in Table 3 through Table 7. 

Figure 6 through Figure 149 show the geographical distribution and relative abundance of select 

groundfish species. They were created with ArcGIS software (ESRI, Redlands, California). The 

maps show haul location where species were caught. Relative abundance is categorized as 

follows: 1) greater than zero but less than or equal to the mean CPUE, 2) greater than the mean 

CPUE but less than or equal to 1 SD from the mean, 3) between 1 and 2 SDs greater than the 

mean CPUE, 4) more than 2 SDs greater than the mean CPUE, and 5) no catch. During survey 

years 2003–2008, species-specific CPUE (kg ha–1) varied by latitude and depth stratum (Figure 

150 through Figure 155).
9

Mean biomass estimates (metric tons) were similarly calculated by multiplying the 

weighted mean CPUE for total area, depth strata, or INPFC region by the appropriate area of the 

stratum or region,

(3)

where b

ˆ is the mean biomass estimate in metric tons, CPUE is the mean CPUE in ha–1

calculated as noted above by weighting the initial mean by area, A is area of the stratum or 

region in ha, and n = 3 when depth strata (shallow, mid, and deep) were combined within an 

INPFC area, or n = 5 if individual INPFC areas were combined or if depth strata for all areas 

were combined (see above). For all depth strata and INPFC areas, the dominant FMP species 

varied from year to year. Species-specific biomass (mt) by year and depth stratum are displayed 

in Figure 156 through Figure 161. Variance for mean biomass estimates (within and among 

INPFC areas and depth strata) was calculated as

(4)

after first adjusting for differences in units and with symbols as defined in equation 3.

Biological Data Collection

The surveys encountered more than 380 species or groups of fish and at least 540 species 

or groups of invertebrates from 2003 to 2008. Frequency of occurrence, depth range, and 

latitudinal range for all identified fish and invertebrates are listed in Appendix B.

All fishes and invertebrates were sorted to species (or the lowest possible taxon), and 

then weighed using an electronic, motion-compensated scale (Marel, Reykjavik, Iceland). Any 

unidentified species were labeled, frozen, or preserved in formalin and retained for later 

identification. Subsamples of selected management species were randomly selected for 

individual measurements (length and weight) and biological sampling (age structure, sex 

determination, or stomach content analysis). Up to 100 sex determinations and length 

measurements (to the nearest cm) were collected per haul from each of these species. Although 

fork length (or total length) is generally measured for most species, anal length was recorded for 

spotted ratfish (Hydrolagus colliei) and Pacific grenadier (Coryphaenoides acrolepis). Carapace 

width was recorded for Dungeness crab (Cancer magister) and grooved tanner crab.

Otoliths were removed to determine age, except for lingcod (Ophiodon elongatus) (fin 

rays) and spiny dogfish (Squalus acanthias) (dorsal spines). A subset of fish selected for length 

measurement was also randomly selected for ageing. Age structures for up to 100 individuals 

per species were collected per haul. For other species, only total counts and weights were 

recorded. From 2003 to 2008, the surveys collected more than 750,000 individual length 

measurements and 136,000 age structures. The species with the greatest number of length 

measurements and age structures include Dover sole (Microstomus pacificus), English sole 

(Parophrys vetulus), longspine thornyhead (Sebastolobus altivelis), Pacific hake (Merluccius 
10

productus), Pacific sanddab (Citharichthys sordidus), rex sole (Glyptocephalus zachirus), 

sablefish (Anoplopoma fimbria), shortspine thornyhead (Sebastolobus alascanus), splitnose 

rockfish (Sebastes diploproa), and stripetail rockfish (Sebastes saxicola). Those 10 species 

represented more than 60% of total length measurements taken during the 2003–2008 surveys. 

Table 8 through Table 10 describe the length and age structure collection effort by species and 

year.

Length and Age Composition

Figure 162 through Figure 208 show frequency of lengths for FMP species encountered 

during the 2003–2008 surveys. Only FMP species with greater than 500 length measurements 

are shown. Length frequencies are subset by year, stratum, and sex. If the number of individuals 

within a depth stratum was less than 10% of the total, then the panel was omitted. Females (F) 

are represented by a solid line; males (M) by a dashed line; and unknown (U) by a bold gray line. 

The mean length for each gender classification (F, M, and U) is displayed in each figure panel. 

Unsexed individuals were included only when the number of individuals with unknown sex was 

more than 2% of the total. Lengths are fork length, except for skates and spiny dogfish (total 

length), and pacific grenadier and spotted ratfish (anal length). The length frequencies represent 

the sum of all measured individuals. They are unweighted and not adjusted for relative 

collection efforts between years or strata.

Age frequencies were analyzed for 14 species for each year collected. The relative 

frequency for each 1-year age bin is shown in Figure 209 through Figure 222. Ages were 

determined from otoliths, dorsal fin rays, or dorsal spines. The age frequencies represent the 

sum of all measured individuals. They are unweighted and not adjusted for relative collection 

efforts between years.

Weight-length Relationships

Individual measurements of weight (kg) and length (cm) were fit to the following weightlength relationship using a nonlinear least squares regression. The length-weight allometric 

relationship is expressed as

W = a × Lb (5)

where W is fish weight in kg, L is fish length in cm, and a and b are fitted parameters from 

nonlinear least squares regression. The results of these analyses, including number sampled and 

the coefficient of determination (r2), are reported in Table 11. As noted, fork length (or total 

length) was measured for most species; however, anal length was recorded for Pacific grenadier. 

The weight-length relationships for commercially important species (Dover sole, petrale sole

[Eopsetta jordani], sablefish, and shortspine thornyheads) varied during 2003–2008 (Figure 223

and Figure 224). Generally, the weight for a given length has decreased with time. In 23 of 33 

cases, the value of the exponent b was lower in 2008 than it was in 2003.
11

Fish Community Metrics

Fish community metrics were calculated for the latitudinal range (32.2°N to 49.2°N) of 

the survey in one degree increments for each year. Latitude increments are centered on a 

geographic break at Cape Mendocino (40.2°N) and progress toward the northern and southern 

bounds of the survey. Values are displayed at the midpoint of each latitude bin. Each latitude 

increment was divided into three depth strata (55–183 m, 184–549 m, and 550–1,280 m). 

Measurements of richness, diversity, and evenness were calculated based on the species 

encountered within each specific latitude and depth bin. Individual counts were estimated based 

on the species average weight (within bin).

Only fish identified to species were considered for the community analyses. Because 

genus and family level classifications overlap with species level identifications, individuals that 

were identified to a more general taxonomic classification could not be counted. For example, 

individuals identified only as Sebastes were discarded, so individuals identified to species within 

Sebastes could be included. The inclusion of overlapping classifications would lead to 

overestimation of species richness.

However, fish identified within the families Bathylagidae or Myctophidae were included 

in the diversity and richness analyses at the family level and not the species level. These families 

were encountered often and identified to the family level frequently. Identification to species 

within these families was less common. In the special cases of Bathylagidae and Myctophidae, 

discarding the more general classification in favor of species level classification would result in 

ignoring more than half of the encounters with members of these families.

Species diversity index (H′) was estimated according to Shannon’s diversity index,

(6)

where S is the total number of species, pi is the proportion S comprised of the ith species. 

Species evenness (E) was estimated according to the following equation,

(7)

where H′ is Shannon’s diversity index and S is the total number of species.

Measurements of fish community richness, diversity, and evenness vary by depth, 

latitude, and year (Figure 225 through Figure 233). Overall, species richness (the total number 

of species) varied more by year than latitude. In all strata, higher richness values were typically 

found south of Cape Mendocino (40.02°N). The lowest richness values were found in stratum 1. 

Patterns of species diversity and eveness demonstrated few trends. Stratum 3 exhibited the 

lowest species diversity and evenness. Variation between years was greater than variation 

between strata or latitudes, although variation between years decreased north of 44.2°N.
12

In stratum 1, the highest richness values were found off Cape Flattery, Washington

(48.2–49.2°N), and Cape Mendocino (40.2–41.2°N) and Point Conception (35.2–36.2°N), 

California. In 2008 species richness was 21 at 32.7°N. This is the largest value found in stratum 

1 (Figure 225). Species diversity varied among years. North of 44.2°N, species diversity and 

evenness increased with latitude and there was less variability among years (Figure 234).

In stratum 2, there was a consistent species richness peak between 46.2 and 47.2°N and a 

consistent depression between 34.2 and 35.2°N (Figure 228). Richness was also elevated 

between 36.2 and 40.2°N. The lowest species richness was found between 44.2 and 46.2°N. 

Areas of higher diversity were found from 44.2 to 45.2°N and from 47.2 to 48.2°N. A consistent 

depression in diversity was located between 46.2 and 47.2°N. As latitude decreased species 

evenness became more variable. Low values of evenness and diversity coincided between 33.2

and 34.2°N.

In stratum 3, the sampling stations off Cape Flattery (48.2–49.2°N) and south of Cape 

Mendocino (37.2–40.2°N) were the most species rich (Figure 231). Areas of high diversity and 

evenness were located from 48.2 to 49.2°N, from 42.2 to 43.2°N, and south of 37.2°N. Between 

41.2 and 42.2°N, diversity and evenness were consistently low in all years.

There was a negative relationship between species diversity and richness in stratum 1 

(P < 0.01, r2 = 0.13) and stratum 2 (P < 0.01, r2 = 0.278). In stratum 3 there was no relationship 

between diversity and richness, but diversity was negatively related to latitude (P < 0.01, r2

= 

0.1816) (Figure 235). There was no relationship between species diversity and evenness.
13

Figures 1–235
14

Figure 1. Total survey area and sampling stations during the 2003–2008 surveys. Color of sampling stations or cells is based on the number of 

times sampled. Green cells have been sampled once, yellow twice, orange three times, and red four times. Neutral cells were not sampled 

during the time period.
15

Figure 2. Geographic extent of the surveys subdivided into INPFC areas. Four main survey strata 

isobaths are shown (55, 183, 549, and 1,280 m).
16

Figure 3. Schematic of Aberdeen-type sampling trawl used during the surveys. Headrope, fishing line, and footrope dimensions are meters (and 

feet). The depth and width of net sections are measured by mesh count. Each section is composed of 3, 4, and 5 mm polyethylene line 

where indicated. For further details, see Stauffer (2004).
17

 

Excalibur

Inverse scope (m)

Net height and width (m)

0 5 10 15 20

 

Ms. Julie

Inverse scope (m)

Net height and width (m)

0 5 10 15 20

 

Noahs Ark

Inverse scope (m)

Net height and width (m)

0 5 10 15 20

0.000 0.002 0.004

 

Raven

Inverse scope (m)

Net height and width (m)

 

Blue Horizon

 

BJ Thomas

0.000 0.002 0.004

 

Captain Jack

Inverse scope (m)

Net height and width (m)

0.000 0.002 0.004

Figure 4. Mean net width and height for trawls conducted as part of the surveys for years 2003–2008. 

Width (●) and height (●) estimates are grouped by vessel and plotted relative to inverse scope 

(1/meters). Coefficients and intercepts of linear regressions are reported in Table 12.
18

 

0

500

1,000

1,500

2,000

2,500

3,000

3,500

4,000

4,500

2003 2004 2005 2006 2007 2008

Year

Total catch (thousands of metric tons)

Figure 5. Trends in total catch of 46 FMP species plus grooved tanner crab and giant grenadier in the 

surveys from 2003 to 2008.
19

Figure 6. Arrowtooth flounder (Atheresthes stomias) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five 

ranges of relative abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 

SD from the mean, between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
20

Figure 7. Arrowtooth flounder distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
21

Figure 8. Arrowtooth flounder distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
22

Figure 9. Aurora rockfish (Sebastes aurora) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges of 

relative abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the 

mean, between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
23

Figure 10. Aurora rockfish distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
24

Figure 11. Aurora rockfish distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
25

Figure 12. Bank rockfish (Sebastes rufus) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges of 

relative abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the

mean, between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
26

Figure 13. Bank rockfish distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
27

Figure 14. Bank rockfish distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
28

Figure 15. Big skate (Raja binoculata) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
29

Figure 16. Big skate distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
30

Figure 17. Big skate distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
31

Figure 18. Blackgill rockfish (Sebastes melanostomus) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five 

ranges of relative abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 

SD from the mean, between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
32

Figure 19. Blackgill rockfish distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
33

Figure 20. Blackgill rockfish distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
34

Figure 21. Bocaccio (Sebastes paucispinis) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges of 

relative abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the 

mean, between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
35

Figure 22. Bocaccio distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
36

Figure 23. Bocaccio distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
37

Figure 24. Calico rockfish (Sebastes dalli) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges of 

relative abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the 

mean, between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
38

Figure 25. Calico rockfish distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
39

Figure 26. Calico rockfish distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
40

Figure 27. California scorpionfish (Scorpaena guttata) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five 

ranges of relative abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 

SD from the mean, between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
41

Figure 28. California scorpionfish distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
42

Figure 29. California scorpionfish distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
43

Figure 30. California skate (Raja inornata) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges of 

relative abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the 

mean, between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
44

Figure 31. California skate distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
45

Figure 32. California skate distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
46

Figure 33. Canary rockfish (Sebastes pinniger) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges of 

relative abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the 

mean, between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
47

Figure 34. Canary rockfish distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
48

Figure 35. Canary rockfish distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
49

Figure 36. Chilipepper rockfish (Sebastes goodei) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges 

of relative abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from 

the mean, between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
50

Figure 37. Chilipepper rockfish distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
51

Figure 38. Chilipepper rockfish distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
52

Figure 39. Curlfin sole (Pleuronichthys decurrens) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges 

of relative abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from 

the mean, between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
53

Figure 40. Curlfin sole distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
54

Figure 41. Curlfin sole distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
55

Figure 42. Darkblotched rockfish (Sebastes crameri) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five 

ranges of relative abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 

SD from the mean, between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
56

Figure 43. Darkblotched rockfish distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
57

Figure 44. Darkblotched rockfish distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
58

Figure 45. Dover sole distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
59

Figure 46. Dover sole distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
60

Figure 47. Dover sole distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
61

Figure 48. English sole distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
62

Figure 49. English sole distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
63

Figure 50. English sole distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
64

Figure 51. Flathead sole (Hippoglossoides elassodon) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five 

ranges of relative abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 

SD from the mean, between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
65

Figure 52. Flathead sole distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
66

Figure 53. Flathead sole distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
67

Figure 54. Giant grenadier distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
68

Figure 55. Giant grenadier distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
69

Figure 56. Giant grenadier distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
70

Figure 57. Greenspotted rockfish (Sebastes chlorostictus) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five 

ranges of relative abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 

SD from the mean, between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
71

Figure 58. Greenspotted rockfish distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
72

Figure 59. Greenspotted rockfish distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
73

Figure 60. Greenstriped rockfish (Sebastes elongatus) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five

ranges of relative abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 

SD from the mean, between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
74

Figure 61. Greenstriped rockfish distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
75

Figure 62. Greenstriped rockfish distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
76

Figure 63. Grooved tanner crab distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
77

Figure 64. Grooved tanner crab distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
78

Figure 65. Grooved tanner crab distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
79

Figure 66. Halfbanded rockfish (Sebastes semicinctus) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five 

ranges of relative abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 

SD from the mean, between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
80

Figure 67. Halfbanded rockfish distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
81

Figure 68. Halfbanded rockfish distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
82

Figure 69. Lingcod distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
83

Figure 70. Lingcod distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
84

Figure 71. Lingcod distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
85

Figure 72. Longnose skate (Raja rhina) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
86

Figure 73. Longnose skate distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
87

Figure 74. Longnose skate distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
88

Figure 75. Longspine thornyhead distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
89

Figure 76. Longspine thornyhead distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
90

Figure 77. Longspine thornyhead distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
91

Figure 78. Pacific cod (Gadus macrocephalus) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges of 

relative abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the 

mean, between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
92

Figure 79. Pacific cod distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
93

Figure 80. Pacific cod distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
94

Figure 81. Pacific flatnose (Antimora microlepis) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges 

of relative abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from 

the mean, between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
95

Figure 82. Pacific flatnose distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
96

Figure 83. Pacific flatnose distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
97

Figure 84. Pacific grenadier distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
98

Figure 85. Pacific grenadier distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
99

Figure 86. Pacific grenadier distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
100

Figure 87. Pacific hake distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
101

Figure 88. Pacific hake distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
102

Figure 89. Pacific hake distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
103

Figure 90. Pacific ocean perch (Sebastes alutus) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges of 

relative abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the 

mean, between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
104

Figure 91. Pacific ocean perch distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
105

Figure 92. Pacific ocean perch distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
106

Figure 93. Pacific sanddab distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
107

Figure 94. Pacific sanddab distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
108

Figure 95. Pacific sanddab distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
109

Figure 96. Petrale sole distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
110

Figure 97. Petrale sole distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
111

Figure 98. Petrale sole distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
112

Figure 99. Pygmy rockfish (Sebastes wilsoni) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges of 

relative abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the 

mean, between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
113

Figure 100. Pygmy rockfish distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
114

Figure 101. Pygmy rockfish distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
115

Figure 102. Redbanded rockfish (Sebastes babcocki) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five 

ranges of relative abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 

SD from the mean, between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
116

Figure 103. Redbanded rockfish distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
117

Figure 104. Redbanded rockfish distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
118

Figure 105. Redstripe rockfish (Sebastes proriger) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges 

of relative abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from 

the mean, between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
119

Figure 106. Redstripe rockfish distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
120

Figure 107. Redstripe rockfish distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
121

Figure 108. Rex sole distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
122

Figure 109. Rex sole distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
123

Figure 110. Rex sole distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
124

Figure 111. Rosethorn rockfish (Sebastes helvomaculatus) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The 

five ranges of relative abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 

1 SD from the mean, between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
125

Figure 112. Rosethorn rockfish distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
126

Figure 113. Rosethorn rockfish distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
127

Figure 114. Rougheye rockfish (Sebastes aleutianus) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five 

ranges of relative abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 

SD from the mean, between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
128

Figure 115. Rougheye rockfish distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
129

Figure 116. Rougheye rockfish distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
130

Figure 117. Sablefish distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
131

Figure 118. Sablefish distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
132

Figure 119. Sablefish distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative abundance are

categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 and 2 

SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
133

Figure 120. Sharpchin rockfish (Sebastes zacentrus) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five 

ranges of relative abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 

SD from the mean, between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
134

Figure 121. Sharpchin rockfish distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
135

Figure 122. Sharpchin rockfish distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
136

Figure 123. Shortbelly rockfish (Sebastes jordani) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges 

of relative abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from 

the mean, between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
137

Figure 124. Shortbelly rockfish distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
138

Figure 125. Shortbelly rockfish distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
139

Figure 126. Shortspine thornyhead distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
140

Figure 127. Shortspine thornyhead distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
141

Figure 128. Shortspine thornyhead distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
142

Figure 129. Spiny dogfish distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
143

Figure 130. Spiny dogfish distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
144

Figure 131. Spiny dogfish distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
145

Figure 132. Splitnose rockfish distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
146

Figure 133. Splitnose rockfish distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
147

Figure 134. Splitnose rockfish distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
148

Figure 135. Spotted ratfish distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
149

Figure 136. Spotted ratfish distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
150

Figure 137. Spotted ratfish distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
151

Figure 138. Squarespot rockfish (Sebastes hopkinsi) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five 

ranges of relative abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 

SD from the mean, between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
152

Figure 139. Squarespot rockfish distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
153

Figure 140. Squarespot rockfish distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
154

Figure 141. Stripetail rockfish distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
155

Figure 142. Stripetail rockfish distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
156

Figure 143. Stripetail rockfish distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
157

Figure 144. Widow rockfish (Sebastes entomelas) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges 

of relative abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from 

the mean, between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
158

Figure 145. Widow rockfish distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
159

Figure 146. Widow rockfish distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative abundance 

are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, between 1 

and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
160

Figure 147. Yellowtail rockfish (Sebastes flavidus) distribution and relative abundance (kg ha–1) from the 2003 and 2004 surveys. The five ranges 

of relative abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from 

the mean, between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
161

Figure 148. Yellowtail rockfish distribution and relative abundance (kg ha–1) from the 2005 and 2006 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
162

Figure 149. Yellowtail rockfish distribution and relative abundance (kg ha–1) from the 2007 and 2008 surveys. The five ranges of relative 

abundance are categorized from top to bottom in the legend as follows: > 0 but ≤ mean CPUE, > mean CPUE but ≤ 1 SD from the mean, 

between 1 and 2 SDs > mean CPUE, between 2 and 3 SDs > mean CPUE, and more than 3 SDs > mean CPUE.
163

Figure 150. Species specific CPUE (kg ha–1) by latitude for survey years 2003–2008 for 9 of 46 FMP species. The dashed line is stratum 1,

55–183 m; the gray line is stratum 2, 184–549 m; and the black line is stratum 3, 550–1,280 m.

Arrowtooth flounder

0

20

40

60

32 34 36 38 40 42 44 46 48

Aurora rockfish

0

20

40

60

32 34 36 38 40 42 44 46 48

CPUE

Bank rockfish

0

20

40

60

80

32 34 36 38 40 42 44 46 48

Big skate

0

20

32 34 36 38 40 42 44 46 48

CPUE

Blackgill rockfish

0

10

20

30

32 34 36 38 40 42 44 46 48

Bocaccio

0

10

20

30

32 34 36 38 40 42 44 46 48

Calico rockfish

0

1

32 34 36 38 40 42 44 46 48

Calif. scorpionfish

0

2

4

32 34 36 38 40 42 44 46 48

Latitude o N

Calif. skate

0

2

4

6

8

10

32 34 36 38 40 42 44 46 48
164

Figure 151. Species specific CPUE (kg ha–1) by latitude for survey years 2003–2008 for 9 of 46 FMP species. Except for canary rockfish and 

chilipepper, indicated above, the dashed line is stratum 1, 55–183 m; the gray line is stratum 2, 184–549 m; and the black line is stratum 3, 

550–1,280 m.
165

Figure 152. Species specific CPUE (kg ha–1) by latitude for survey years 2003–2008 for 9 of 46 FMP species. The dashed line is stratum 1, 

55–183 m; the gray line is stratum 2, 184–549 m; and the black line is stratum 3, 550–1,280 m.
166

Figure 153. Species specific CPUE (kg ha–1) by latitude for survey years 2003–2008 for 9 of 46 FMP species. The dashed line is stratum 1,

55–183 m; the gray line is stratum 2, 184–549 m; and the black line is stratum 3, 550–1,280 m.
167

Figure 154. Species specific CPUE (kg ha–1) by latitude for survey years 2003–2008 for 9 of 46 FMP species. The dashed line is stratum 1, 

55–183 m; the gray line is stratum 2, 184–549 m; and the black line is stratum 3, 550–1,280 m.
168

Figure 155. Species specific CPUE (kg ha–1) by latitude for survey years 2003–2008 for 1 of 46 FMP species plus grooved tanner crab and giant 

grenadier. The dashed line is stratum 1, 55–183 m; the gray line is stratum 2, 184–549 m; and the black line is stratum 3, 550–1,280 m.
169

Figure 156. Species specific biomass (mt) by year for 9 of 46 FMP species. The dashed line is stratum 1, 55–183 m; the gray line is stratum 2, 

184–549 m; and the black line is stratum 3, 550–1,280 m.
170

Figure 157. Species specific biomass (mt) by year for 9 of 46 FMP species. Except for canary rockfish and chilipepper, indicated above, the 

dashed line is stratum 1, 55–183 m; the gray line is stratum 2, 184–549 m; and the black line is stratum 3, 550–1,280 m.
171

Figure 158. Species specific biomass (mt) by year for 9 of 46 FMP species. The dashed line is stratum 1, 55–183 m; the gray line is stratum 2, 

184–549 m; and the black line is stratum 3, 550–1,280 m.
172

Figure 159. Species specific biomass (mt) by year for 9 of 46 FMP species. The dashed line is stratum 1, 55–183 m; the gray line is stratum 2, 

184–549 m; and the black line is stratum 3, 550–1,280 m.
173

Figure 160. Species specific biomass (mt) by year for 9 of 46 FMP species. The dashed line is stratum 1, 55–183 m; the gray line is stratum 2, 

184–549 m; and the black line is stratum 3, 550–1,280 m.
174

Figure 161. Species specific biomass (mt) by year for 1 of 46 FMP species plus grooved tanner crab and giant grenadier. The dashed line is 

stratum 1, 55–183 m; the gray line is stratum 2, 184–549 m; and the black line is stratum 3, 550–1,280 m.
175

20 40 60 80

0 50 150

Stratum 1 (55-155 m)

 

F = 42.8

M = 34.4

0 50 150

2003

20 40 60 80

0 50 150

Stratum 2 (155-549 m)

 

F = 45.6

M = 36.2

20 40 60 80

0 50 150

Stratum 1 (55-155 m)

 

F = 40.9

M = 33.8

0 50 150

2004

20 40 60 80

0 50 150

Stratum 2 (155-549 m)

 

F = 45.7

M = 36.7

20 40 60 80

0 50 150

Stratum 1 (55-155 m)

 

F = 44.7

M = 35.6

0 50 150

2005

20 40 60 80

0 50 150

Stratum 2 (155-549 m)

 

F = 48.8

M = 38

20 40 60 80

0 50 150

Stratum 1 (55-155 m)

 

F = 45.1

M = 35.9

0 50 150

2006

20 40 60 80

0 50 150

Stratum 2 (155-549 m)

 

F = 48.7

M = 38.1

20 40 60 80

0 50 150

Stratum 1 (55-155 m)

 

F = 46.1

M = 34.7

0 50 150

2007

20 40 60 80

0 50 150

Stratum 2 (155-549 m)

 

F = 50.7

M = 39.8

20 40 60 80

0 50 150

Stratum 1 (55-155 m)

 

F = 40

M = 31.5

0 50 150

20 40 60 80

2008

20 40 60 80

0 50 150

Stratum 2 (155-549 m)

 

F = 48.4

M = 37.2

20 40 60 80

Length (cm)

Frequency

Figure 162. Frequency of lengths by gender in stratum 1 (left column) and stratum 2 (right column) for 

arrowtooth flounder from the 2003–2008 surveys. Females are represented by a solid line, males 

by a dashed line. Mean length for each gender classification (F, M) is displayed in each panel.

Stratum 1 (55-155 m) Stratum 2 (155-549 m)
176

10 20 30 40 50

0 40 80 120

Stratum 2 (155-549 m)

 

F = 26.6

M = 26.2

0 40 80 120

2003

10 20 30 40 50

0 40 80 120

Stratum 2 (155-549 m)

 

F = 27

M = 26.3

0 40 80 120

2004

10 20 30 40 50

0 40 80 120

Stratum 2 (155-549 m)

 

F = 25.4

M = 24.8

0 40 80 120

2005

10 20 30 40 50

0 40 80 120

Stratum 2 (155-549 m)

 

F = 25.6

M = 25.5

0 40 80 120

2006

10 20 30 40 50

0 40 80 120

Stratum 2 (155-549 m)

 

F = 25.6

M = 25.8

0 40 80 120

2007

10 20 30 40 50

0 40 80 120

Stratum 2 (155-549 m)

 

F = 25.6

M = 25.7

0 40 80 120

10 20 30 40 50

2008

Length (cm)

Frequency

Figure 163. Frequency of lengths by gender in stratum 2 for aurora rockfish from the 2003–2008 surveys. 

Females are represented by a solid line, males by a dashed line. Mean length for each gender 

classification (F, M) is displayed in each panel.

Stratum 2 (155-549 m)
177

20 30 40 50

0 5 10 15

Stratum 1 (55-182 m)

 

0 5 10 15

2003

20 30 40 50

0 5 10 15

Stratum 2 (183-549 m)

 

F = 32.4

M = 33

20 30 40 50

0 5 10 15

Stratum 1 (55-182 m)

 

F = 16.8

M = 14.4

0 5 10 15

2004

20 30 40 50

0 5 10 15

Stratum 2 (183-549 m)

 

F = 20.5

M = 20

20 30 40 50

0 5 10 15

Stratum 1 (55-182 m)

 

F = 17.5

M = 15.1

0 5 10 15

2005

20 30 40 50

0 5 10 15

Stratum 2 (183-549 m)

 

F = 37.6

M = 29

20 30 40 50

0 5 10 15

Stratum 1 (55-182 m)

 

F = 23.8

M = 22.8

0 5 10 15

2006

20 30 40 50

0 5 10 15

Stratum 2 (183-549 m)

 

F = 33.5

M = 34.2

20 30 40 50

0 5 10 15

Stratum 1 (55-182 m)

 

F = 25.7

M = 20.6

0 5 10 15

2007

20 30 40 50

0 5 10 15

Stratum 2 (183-549 m)

 

F = 30.7

M = 30.7

20 30 40 50

0 5 10 15

Stratum 1 (55-182 m)

 

F = 17

M = 18

0 5 10 15

20 30 40 50

2008

20 30 40 50

0 5 10 15

Stratum 2 (183-549 m)

 

F = 33.5

M = 33.1

20 30 40 50

Length (cm)

Frequency

Figure 164. Frequency of lengths by gender in stratum 1 (left column) and stratum 2 (right column) for 

bank rockfish from the 2003–2008 surveys. Females are represented by a solid line, males by a 

dashed line. Mean length for each gender classification (F, M) is displayed in each panel.

Stratum 1 (55-182 m) Stratum 2 (183-549 m)
178

0 50 100 150

0 2 4 6

Stratum 1 (55-182 m)

 

F = 62.1

M = 69.3

0 2 4 6

0 50 100 150

0 2 4 6

Stratum 2 (183-549 m)

 

F = 95

M = 110

0 50 100 150

0 2 4 6

Stratum 1 (55-182 m)

 

F = 64.1

M = 84.7

0 2 4 6

0 50 100 150

0 2 4 6

Stratum 2 (183-549 m)

 

F = 113.5

M = 114.5

0 50 100 150

0 2 4 6

Stratum 1 (55-182 m)

 

F = 68.1

M = 78.1

0 2 4 6

0 50 100 150

0 2 4 6

Stratum 2 (183-549 m)

 

F = 71.1

M = 95.3

0 50 100 150

0 2 4 6

Stratum 1 (55-182 m)

 

0 2 4 6

0 50 100 150

0 2 4 6

Stratum 2 (183-549 m)

 

0 50 100 150

0 2 4 6

Stratum 1 (55-182 m)

 

0 2 4 6

00

0 50 100 150

0 2 4 6

Stratum 2 (183-549 m)

 

0 50 100 150

0 2 4 6

Stratum 1 (55-182 m)

 

F = 60.6

M = 68

0 2 4 6

0 50 100 150 0 50 100 150

0 2 4 6

Stratum 2 (183-549 m)

 

F = 134

M = 38

0 50 100 150

Length (cm)

Frequency

Figure 165. Frequency of lengths by gender in stratum 1 (left column) and stratum 2 (right column) for 

big skate from the 2003–2008 surveys. Females are represented by a solid line, males by a 

dashed line. Mean length for each gender classification (F, M) is displayed in each panel.

2003

2007

2008

2006

2005

2004

Stratum 1 (55-182 m) Stratum 2 (183-549 m)
179

10 20 30 40 50 60

0 10 20

Stratum 2 (183-549 m)

 

F = 31.1

M = 31.5

0 10 20

2003

10 20 30 40 50 60

0 10 20

Stratum 2 (183-549 m)

 

F = 31

M = 31.5

0 10 20

2004

10 20 30 40 50 60

0 10 20

Stratum 2 (183-549 m)

 

F = 33.8

M = 32.2

0 10 20

2005

10 20 30 40 50 60

0 10 20

Stratum 2 (183-549 m)

 

F = 30.6

M = 30.4

0 10 20

2006

10 20 30 40 50 60

0 10 20

Stratum 2 (183-549 m)

 

F = 35.3

M = 35

0 10 20

2007

10 20 30 40 50 60

0 10 20

Stratum 2 (183-549 m)

 

F = 33.5

M = 33.9

0 10 20

10 20 30 40 50 60

2008

Length (cm)

Frequency

Figure 166. Frequency of lengths by gender in stratum 2 for blackgill rockfish from the 2003–2008 

surveys. Females are represented by a solid line, males by a dashed line. Mean length for each 

gender classification (F, M) is displayed in each panel.

Stratum 2 (183-549 m)
180

20 40 60 80

0 5 15

Stratum 1 (55-182 m)

 

F = 48.9

M = 41.4

0 5 15

2003

20 40 60 80

0 5 15

Stratum 2 (183-549 m)

 

F = 54.4

M = 49.5

20 40 60 80

0 5 15

Stratum 1 (55-182 m)

 

F = 44.2

M = 39.7

0 5 15

2004

20 40 60 80

0 5 15

Stratum 2 (183-549 m)

 

F = 44.5

M = 46.1

20 40 60 80

0 5 15

Stratum 1 (55-182 m)

 

F = 40.1

M = 32.1

0 5 15

2005

20 40 60 80

0 5 15

Stratum 2 (183-549 m)

 

F = 53.9

M = 51.8

20 40 60 80

0 5 15

Stratum 1 (55-182 m)

 

F = 47.1

M = 39.4

0 5 15

2006

20 40 60 80

0 5 15

Stratum 2 (183-549 m)

 

F = 61.1

M = 63.1

20 40 60 80

0 5 15

Stratum 1 (55-182 m)

 

F = 46.7

M = 46.1

0 5 15

2007

20 40 60 80

0 5 15

Stratum 2 (183-549 m)

 

F = 53.1

M = 52.2

20 40 60 80

0 5 15

Stratum 1 (55-182 m)

 

F = 56.6

M = 52.9

0 5 15

20 40 60 80

2008

20 40 60 80

0 5 15

Stratum 2 (183-549 m)

 

F = 59

M = 56.8

20 40 60 80

Length (cm)

Frequency

Figure 167. Frequency of lengths by gender in stratum 1 (left column) and stratum 2 (right column) for 

bocaccio from the 2003–2008 surveys. Females are represented by a solid line, males by a 

dashed line. Mean length for each gender classification (F, M) is displayed in each panel.

Stratum 1 (55-182 m) Stratum 2 (183-549 m)
181

10 15 20 25

0 20 40

Stratum 1 (55-182 m)

 

F = 14.4

M = 12

0 20 40

2003

10 15 20 25

0 20 40

Stratum 1 (55-182 m)

 

F = 15.3

M = 13.1

0 20 40

2004

10 15 20 25

0 20 40

Stratum 1 (55-182 m)

 

F = 12.2

M = 11.7

0 20 40

2005

10 15 20 25

0 20 40

Stratum 1 (55-182 m)

 

F = 11.8

M = 12.1

0 20 40

2006

10 15 20 25

0 20 40

Stratum 1 (55-182 m)

 

F = 12.1

M = 11.5

0 20 40

2007

10 15 20 25

0 20 40

Stratum 1 (55-182 m)

 

F = 13.3

M = 13

0 20 40

10 15 20 25

2008

Length (cm)

Frequency

Figure 168. Frequency of lengths by gender in stratum 1 for calico rockfish from the 2003–2008 surveys. 

Females are represented by a solid line, males by a dashed line. Mean length for each gender 

classification (F, M) is displayed in each panel.

Stratum 1 (55-182 m)
182

15 20 25 30 35

0 10 20 30

Stratum 1 (55-182 m)

 

F = 27.2

M = 25.6

0 10 20 30

2003

15 20 25 30 35

0 10 20 30

Stratum 1 (55-182 m)

 

F = 25

M = 23.4

0 10 20 30

2004

15 20 25 30 35

0 10 20 30

Stratum 1 (55-182 m)

 

F = 25.7

M = 24.2

0 10 20 30

2005

15 20 25 30 35

0 10 20 30

Stratum 1 (55-182 m)

 

F = 27

M = 24.7

0 10 20 30

2006

15 20 25 30 35

0 10 20 30

Stratum 1 (55-182 m)

 

F = 26.1

M = 24

0 10 20 30

2007

15 20 25 30 35

0 10 20 30

Stratum 1 (55-182 m)

 

F = 26.4

M = 24.2

0 10 20 30

15 20 25 30 35

2008

Length (cm)

Frequency

Figure 169. Frequency of lengths by gender in stratum 1 for California scorpionfish from the 2003−2008 

surveys. Females are represented by a solid line, males by a dashed line. Mean length for each 

gender classification (F, M) is displayed in each panel.

Stratum 1 (55-182 m)
183

20 30 40 50 60 70 80

0 4 8 12

Stratum 1 (55-182 m)

 

F = 40.4

M = 42.4

0 4 8 12

2003

20 30 40 50 60 70 80

0 4 8 12

Stratum 1 (55-182 m)

 

F = 46.5

M = 44.8

0 4 8 12

2004

20 30 40 50 60 70 80

0 4 8 12

Stratum 1 (55-182 m)

 

F = 44

M = 41.4

0 4 8 12

2005

20 30 40 50 60 70 80

0 4 8 12

Stratum 1 (55-182 m)

 

0 4 8 12

2006

20 30 40 50 60 70 80

0 4 8 12

Stratum 1 (55-182 m)

 

0 4 8 12

2007

20 30 40 50 60 70 80

0 4 8 12

Stratum 1 (55-182 m)

 

F = 38.8

M = 39.7

0 4 8 12

20 30 40 50 60 70 80

2008

Length (cm)

Frequency

Figure 170. Frequency of lengths by gender in stratum 1 for California skate from the 2003–2008 

surveys. Females are represented by a solid line, males by a dashed line. Mean length for each 

gender classification (F, M) is displayed in each panel.

Stratum 1 (55-182 m)
184

10 20 30 40 50 60

0 20 40

Stratum 1 (55-350 m)

 

F = 41.8

M = 39.2

0 20 40

2003

10 20 30 40 50 60

0 20 40

Stratum 1 (55-350 m)

 

F = 41.2

M = 38.8

0 20 40

2004

10 20 30 40 50 60

0 20 40

Stratum 1 (55-350 m)

 

F = 40.3

M = 41

0 20 40

2005

10 20 30 40 50 60

0 20 40

Stratum 1 (55-350 m)

 

F = 47.2

M = 45

0 20 40

2006

10 20 30 40 50 60

0 20 40

Stratum 1 (55-350 m)

 

F = 42.1

M = 41.8

0 20 40

2007

10 20 30 40 50 60

0 20 40

Stratum 1 (55-350 m)

 

F = 37.6

M = 34.3

U = 13.9

0 20 40

10 20 30 40 50 60

2008

Length (cm)

Figure 171. Frequency of lengths by gender in stratum 1 for canary rockfish from the 2003–2008 

surveys. Females are represented by a solid line, males by a dashed line, and individuals without 

gender determination by a thick gray line. Mean length for each gender classification (F, M, and 

U [unknown]) is displayed in each panel.

Frequency

Stratum 1 (55-350 m)
185

10 20 30 40 50

0 50 150

Stratum 1 (55-155 m)

 

F = 31.2

M = 27

U = 11

0 50 150

2003

10 20 30 40 50

0 50 150

Stratum 2 (155-400 m)

 

F = 32.1

M = 29.4

10 20 30 40 50

0 50 150

Stratum 1 (55-155 m)

 

F = 29.8

M = 23.4

U = 11.3

0 50 150

2004

10 20 30 40 50

0 50 150

Stratum 2 (155-400 m)

 

F = 35.1

M = 31.4

10 20 30 40 50

0 50 150

Stratum 1 (55-155 m)

 

F = 27.7

M = 24.4

U = 13.1

0 50 150

2005

10 20 30 40 50

0 50 150

Stratum 2 (155-400 m)

 

F = 36.2

M = 30.3

10 20 30 40 50

0 50 150

Stratum 1 (55-155 m)

 

F = 33.2

M = 29.6

0 50 150

2006

10 20 30 40 50

0 50 150

Stratum 2 (155-400 m)

 

F = 39.8

M = 31.4

10 20 30 40 50

0 50 150

Stratum 1 (55-155 m)

 

F = 34.2

M = 29.5

U = 11.5

0 50 150

2007

10 20 30 40 50

0 50 150

Stratum 2 (155-400 m)

 

F = 38.8

M = 32.8

10 20 30 40 50

0 50 150

Stratum 1 (55-155 m)

 

F = 34.8

M = 29.3

U = 14.8

0 50 150

10 20 30 40 50

2008

10 20 30 40 50

0 50 150

Stratum 2 (155-400 m)

 

F = 41.2

M = 32.2

10 20 30 40 50

Length (cm)

Frequency

Figure 172. Frequency of lengths by gender in stratum 1 (left column) and stratum 2 (right column) for 

chilipepper rockfish from the 2003–2008 surveys. Females are represented by a solid line, males 

by a dashed line, and individuals without gender determination by a thick gray line. Mean length 

for each gender classification (F, M, and U) is displayed in each panel.

Stratum 1 (55-155 m) Stratum 2 (155-400 m)
186

5 10 15 20 25 30 35

0 10 30

Stratum 1 (55-182 m)

 

F = 23

M = 20.2

0 10 30

2003

5 10 15 20 25 30 35

0 10 30

Stratum 1 (55-182 m)

 

F = 23.8

M = 20.1

0 10 30

2004

5 10 15 20 25 30 35

0 10 30

Stratum 1 (55-182 m)

 

F = 24.8

M = 21.8

0 10 30

2005

5 10 15 20 25 30 35

0 10 30

Stratum 1 (55-182 m)

 

F = 23.3

M = 20.5

0 10 30

2006

5 10 15 20 25 30 35

0 10 30

Stratum 1 (55-182 m)

 

F = 24.7

M = 21.6

0 10 30

2007

5 10 15 20 25 30 35

0 10 30

Stratum 1 (55-182 m)

 

F = 24.2

M = 20.5

0 10 30

5 10 15 20 25 30 35

2008

Length (cm)

Frequency

Figure 173. Frequency of lengths by gender in stratum 1 for curlfin sole from the 2003–2008 surveys. 

Females are represented by a solid line, males by a dashed line. Mean length for each gender 

classification (F, M) is displayed in each panel.

Stratum 1 (55-182 m)
187

10 20 30 40 50

0 20 60

Stratum 1 (55-182 m)

 

F = 24.7

M = 24.7

0 20 60

2003

10 20 30 40 50

0 20 60

Stratum 2 (183-549 m)

 

F = 32.1

M = 29.2

10 20 30 40 50

0 20 60

Stratum 1 (55-182 m)

 

F = 24

M = 23.7

0 20 60

2004

10 20 30 40 50

0 20 60

Stratum 2 (183-549 m)

 

F = 29

M = 29.1

10 20 30 40 50

0 20 60

Stratum 1 (55-182 m)

 

F = 22.2

M = 21.4

0 20 60

2005

10 20 30 40 50

0 20 60

Stratum 2 (183-549 m)

 

F = 32.8

M = 30.9

10 20 30 40 50

0 20 60

Stratum 1 (55-182 m)

 

F = 20.5

M = 20.2

0 20 60

2006

10 20 30 40 50

0 20 60

Stratum 2 (183-549 m)

 

F = 30.7

M = 30.1

10 20 30 40 50

0 20 60

Stratum 1 (55-182 m)

 

F = 21.6

M = 22.5

0 20 60

2007

10 20 30 40 50

0 20 60

Stratum 2 (183-549 m)

 

F = 31.2

M = 30.6

10 20 30 40 50

0 20 60

Stratum 1 (55-182 m)

 

F = 20.6

M = 20.6

0 20 60

10 20 30 40 50

2008

10 20 30 40 50

0 20 60

Stratum 2 (183-549 m)

 

F = 29.4

M = 28.4

10 20 30 40 50

Length (cm)

Frequency

Figure 174. Frequency of lengths by gender in stratum 1 (left column) and stratum 2 (right column) for 

darkblotched rockfish from the 2003–2008 surveys. Females are represented by a solid line,

males by a dashed line. Mean length for each gender classification (F, M) is displayed in each 

panel.

Stratum 1 (55-182 m) Stratum 2 (183-549 m)
188

10 20 30 40 50 60 70

0 200 500

Stratum 1 (55-182 

 

F = 34.8

M = 30.5

0 200 500

2003

10 20 30 40 50 60 70

0 200 500

Stratum 2 (183-549 

 

F = 36.3

M = 32.8

10 20 30 40 50 60 70

0 200 500

Stratum 3 (550-1,2 

 

F = 42.7

M = 38.7

10 20 30 40 50 60 70

0 200 500

Stratum 1 (55-182 

 

F = 32.6

M = 29.1

0 200 500

2004

10 20 30 40 50 60 70

0 200 500

Stratum 2 (183-549 

 

F = 34.4

M = 31.2

10 20 30 40 50 60 70

0 200 500

Stratum 3 (550-1,2 

 

F = 41.8

M = 37.9

10 20 30 40 50 60 70

0 200 500

Stratum 1 (55-182 

 

F = 32

M = 28.6

0 200 500

2005

10 20 30 40 50 60 70

0 200 500

Stratum 2 (183-549 

 

F = 34.8

M = 31.8

10 20 30 40 50 60 70

0 200 500

Stratum 3 (550-1,2 

 

F = 41.7

M = 37.2

10 20 30 40 50 60 70

0 200 500

Stratum 1 (55-182 

 

F = 33.6

M = 29.5

0 200 500

2006

10 20 30 40 50 60 70

0 200 500

Stratum 2 (183-549 

 

F = 34.8

M = 31.8

10 20 30 40 50 60 70

0 200 500

Stratum 3 (550-1,2 

 

F = 42.4

M = 37.6

10 20 30 40 50 60 70

0 200 500

Stratum 1 (55-182 

 

F = 33.7

M = 29.7

0 200 500

2007

10 20 30 40 50 60 70

0 200 500

Stratum 2 (183-549 

 

F = 34.6

M = 32.3

10 20 30 40 50 60 70

0 200 500

Stratum 3 (550-1,2 

 

F = 41.4

M = 37.6

10 20 30 40 50 60 70

0 200 500

Stratum 1 (55-182 

 

F = 34.2

M = 29.4

0 200 500

10 20 30 40 50 60 70

2008

10 20 30 40 50 60 70

0 200 500

Stratum 2 (183-549 

 

F = 35.4

M = 32.5

10 20 30 40 50 60 70 10 20 30 40 50 60 70

0 200 500

Stratum 3 (550-1,2 

 

F = 42.5

M = 37.9

10 20 30 40 50 60 70

Length (cm)

Frequency

Figure 175. Frequency of lengths by gender in stratum 1 (left column), stratum 2 (middle column), and 

stratum 3 (right column) for Dover sole from the 2003–2008 surveys. Females are represented by 

a solid line, males by a dashed line. Mean length for each gender classification (F, M) is 

displayed in each panel.

Stratum 1 (55-182 m) Stratum 2 (183-549 m) Stratum 3 (550-1,280 m)
189

10 20 30 40

0 200 500

Stratum 1 (55-182 m)

 

F = 30.3

M = 24.1

0 200 500

2003

10 20 30 40

0 200 500

Stratum 1 (55-182 m)

 

F = 30

M = 24.1

0 200 500

2004

10 20 30 40

0 200 500

Stratum 1 (55-182 m)

 

F = 29.1

M = 23.7

0 200 500

2005

10 20 30 40

0 200 500

Stratum 1 (55-182 m)

 

F = 29.2

M = 23.9

0 200 500

2006

10 20 30 40

0 200 500

Stratum 1 (55-182 m)

 

F = 29

M = 23.2

0 200 500

2007

10 20 30 40

0 200 500

Stratum 1 (55-182 m)

 

F = 27.7

M = 22.4

0 200 500

10 20 30 40

2008

Length (cm)

Frequency

Figure 176. Frequency of lengths by gender in stratum 1 for English sole from the 2003–2008 surveys. 

Females are represented by a solid line, males by a dashed line. Mean length for each gender 

classification (F, M) is displayed in each panel.

Stratum 1 (55-182 m)
190

15 20 25 30 35 40 45

0 50 150

Stratum 1 (55-182 m)

 

F = 31.3

M = 28.3

0 50 150

2003

15 20 25 30 35 40 45

0 50 150

Stratum 1 (55-182 m)

 

F = 32.2

M = 29.1

0 50 150

2004

15 20 25 30 35 40 45

0 50 150

Stratum 1 (55-182 m)

 

F = 32.1

M = 28.6

0 50 150

2005

15 20 25 30 35 40 45

0 50 150

Stratum 1 (55-182 m)

 

F = 32.1

M = 27.8

0 50 150

2006

15 20 25 30 35 40 45

0 50 150

Stratum 1 (55-182 m)

 

F = 32.8

M = 28.4

0 50 150

2007

15 20 25 30 35 40 45

0 50 150

Stratum 1 (55-182 m)

 

F = 31.6

M = 27.8

0 50 150

15 20 25 30 35 40 45

2008

Length (cm)

Frequency

Figure 177. Frequency of lengths by gender in stratum 1 for flathead sole from the 2003–2008 surveys. 

Females are represented by a solid line, males by a dashed line. Mean length for each gender 

classification (F, M) is displayed in each panel.

Stratum 1 (55-182 m)
191

10 20 30 40

0 10 20 30

Stratum 1 (55-182 m)

 

F = 28.6

M = 29.5

0 10 20 30

10 20 30 40

0 10 20 30

Stratum 1 (55-182 m)

 

F = 26.3

M = 27.7

U = 15

0 10 20 30

10 20 30 40

0 10 20 30

Stratum 1 (55-182 m)

 

F = 24.9

M = 24.3

U = 11.6

0 10 20 30

10 20 30 40

0 10 20 30

Stratum 1 (55-182 m)

 

F = 27.1

M = 27.2

U = 12.9

0 10 20 30

10 20 30 40

0 10 20 30

Stratum 1 (55-182 m)

 

F = 27.4

M = 28.7

U = 11

0 10 20 30

10 20 30 40

0 10 20 30

Stratum 1 (55-182 m)

 

F = 24.1

M = 22.1

U = 11.3

0 10 20 30

10 20 30 40

Length (cm)

Frequency

Figure 178. Frequency of lengths by gender in stratum 1 for greenspotted rockfish from the 2003–2008 

surveys. Females are represented by a solid line, males by a dashed line, and individuals without 

gender determination by a thick gray line. Mean length for each gender classification (F, M, and 

U) is displayed in each panel.

2003

2004

2005

2006

2007

2008

Stratum 1 (55-182 m)
192

5 10 15 20 25 30 35 40

0 50 150

Stratum 1 (55-182 m)

 

F = 26.7

M = 24.1

0 50 150

2003

5 10 15 20 25 30 35 40

0 50 150

Stratum 2 (183-549 m)

 

F = 26.7

M = 24.6

5 10 15 20 25 30 35 40

0 50 150

Stratum 1 (55-182 m)

 

F = 26.1

M = 23.9

0 50 150

2004

5 10 15 20 25 30 35 40

0 50 150

Stratum 2 (183-549 m)

 

F = 27

M = 25.1

5 10 15 20 25 30 35 40

0 50 150

Stratum 1 (55-182 m)

 

F = 24.9

M = 22.9

0 50 150

2005

5 10 15 20 25 30 35 40

0 50 150

Stratum 2 (183-549 m)

 

F = 25.5

M = 24

5 10 15 20 25 30 35 40

0 50 150

Stratum 1 (55-182 m)

 

F = 25.2

M = 23.2

0 50 150

2006

5 10 15 20 25 30 35 40

0 50 150

Stratum 2 (183-549 m)

 

F = 25.1

M = 23.7

5 10 15 20 25 30 35 40

0 50 150

Stratum 1 (55-182 m)

 

F = 25.6

M = 23.6

0 50 150

2007

5 10 15 20 25 30 35 40

0 50 150

Stratum 2 (183-549 m)

 

F = 27.4

M = 25.1

5 10 15 20 25 30 35 40

0 50 150

Stratum 1 (55-182 m)

 

F = 25.2

M = 22.7

0 50 150

5 10 15 20 25 30 35 40

2008

5 10 15 20 25 30 35 40

0 50 150

Stratum 2 (183-549 m)

 

F = 26.2

M = 24.3

5 10 15 20 25 30 35 40

Length (cm)

Frequency

Figure 179. Frequency of lengths by gender in stratum 1 (left column) and stratum 2 (right column) for 

greenstriped rockfish from the 2003–2008 surveys. Females are represented by a solid line,

males by a dashed line. Mean length for each gender classification (F, M) is displayed in each 

panel.

Stratum 1 (55-182 m) Stratum 2 (183-549 m)
193

10 15 20

0 200 500

Stratum 1 (55-182 m)

 

0 200 500

2003

10 15 20

0 200 500

Stratum 1 (55-182 m)

 

F = 13.7

M = 11.2

U = 10.6

0 200 500

2004

10 15 20

0 200 500

Stratum 1 (55-182 m)

 

F = 11.9

M = 10.7

U = 9.7

0 200 500

2005

10 15 20

0 200 500

Stratum 1 (55-182 m)

 

F = 13.3

M = 11.5

U = 10.1

0 200 500

2006

10 15 20

0 200 500

Stratum 1 (55-182 m)

 

F = 13.8

M = 12.1

U = 11.4

0 200 500

2007

10 15 20

0 200 500

Stratum 1 (55-182 m)

 

F = 13.8

M = 12

U = 12.9

0 200 500

10 15 20

2008

Length (cm)

Frequency

Figure 180. Frequency of lengths by gender in stratum 1 for halfbanded rockfish from the 2003–2008 

surveys. Females are represented by a solid line; males by a dashed line; and individuals without 

gender determination by a thick gray line. Mean length for each gender classification (F, M, and 

U) is displayed in each panel.

Stratum 1 (55-182 m)
194

20 40 60 80 100

0 20 40

Stratum 1 (55-182 m)

 

F = 50.5

M = 45.8

U = 26

0 20 40

20 40 60 80 100

0 20 40

Stratum 2 (183-549 m)

 

F = 60.9

M = 53.6

U = 54.4

20 40 60 80 100

0 20 40

Stratum 1 (55-182 m)

 

F = 48.2

M = 48.2

U = 22.3

0 20 40

20 40 60 80 100

0 20 40

Stratum 2 (183-549 m)

 

F = 63.7

M = 55.5

20 40 60 80 100

0 20 40

Stratum 1 (55-182 m)

 

F = 54.6

M = 51.3

U = 19.8

0 20 40

20 40 60 80 100

0 20 40

Stratum 2 (183-549 m)

 

F = 70.9

M = 64.1

20 40 60 80 100

0 20 40

Stratum 1 (55-182 m)

 

F = 65.8

M = 59.3

0 20 40

20 40 60 80 100

0 20 40

Stratum 2 (183-549 m)

 

F = 73.8

M = 64.3

20 40 60 80 100

0 20 40

Stratum 1 (55-182 m)

 

F = 65.1

M = 54.2

U = 17.4

0 20 40

20 40 60 80 100

0 20 40

Stratum 2 (183-549 m)

 

F = 77.3

M = 62.2

20 40 60 80 100

0 20 40

Stratum 1 (55-182 m)

 

F = 44.4

M = 39.3

U = 18.2

0 20 40

20 40 60 80 100 20 40 60 80 100

0 20 40

Stratum 2 (183-549 m)

 

F = 75.3

M = 67.8

U = 59.8

20 40 60 80 100

Length (cm)

Frequency

Figure 181. Frequency of lengths by gender in stratum 1 (left column) and stratum 2 (right column) for 

lingcod from the 2003–2008 surveys. Females are represented by a solid line, males by a dashed 

line, and individuals without gender determination by a thick gray line. Mean length for each 

gender classification (F, M, and U) is displayed in each panel.

2003

2004

2008

2007

2006

2005

Stratum 1 (55-182 m) Stratum 2 (183-549 m)
195

0 20 40 60 80 100 120 140

0 10 20 30

Stratum 1 (55-182 m)

 

F = 59

M = 60.1

0 10 20 30

2003

0 20 40 60 80 100 120 140

0 10 20 30

Stratum 2 (183-549 m)

 

F = 59.5

M = 60

0 20 40 60 80 100 120 140

0 10 20 30

Stratum 1 (55-182 m)

 

F = 57.9

M = 56.2

0 10 20 30

2004

0 20 40 60 80 100 120 140

0 10 20 30

Stratum 2 (183-549 m)

 

F = 59.8

M = 58.7

0 20 40 60 80 100 120 140

0 10 20 30

Stratum 1 (55-182 m)

 

F = 57.8

M = 58.9

0 10 20 30

2005

0 20 40 60 80 100 120 140

0 10 20 30

Stratum 2 (183-549 m)

 

F = 58.3

M = 55.4

0 20 40 60 80 100 120 140

0 10 20 30

Stratum 1 (55-182 m)

 

0 10 20 30

2006

0 20 40 60 80 100 120 140

0 10 20 30

Stratum 2 (183-549 m)

 

0 20 40 60 80 100 120 140

0 10 20 30

Stratum 1 (55-182 m)

 

0 10 20 30

2007

0 20 40 60 80 100 120 140

0 10 20 30

Stratum 2 (183-549 m)

 

0 20 40 60 80 100 120 140

0 10 20 30

Stratum 1 (55-182 m)

 

F = 56.4

M = 57.3

0 10 20 30

0 20 40 60 80 100 120 140

2008

0 20 40 60 80 100 120 140

0 10 20 30

Stratum 2 (183-549 m)

 

F = 63

M = 62.7

0 20 40 60 80 100 120 140

Length (cm)

Frequency

Figure 182. Frequency of lengths by gender in stratum 1 (left column) and stratum 2 (right column) for 

longnose skate from the 2003–2008 surveys. Females are represented by a solid line, males by a 

dashed line. Mean length for each gender classification (F, M) is displayed in each panel.

Stratum 1 (55-182 m) Stratum 2 (183-549 m)
196

5 10 15 20 25 30 35

0 400 1000

Stratum 2 (183-549 m)

 

U = 21.7

0 400 1000

2003

5 10 15 20 25 30 35

0 400 1000

Stratum 3 (550-1,280 m)

 

U = 19.7

5 10 15 20 25 30 35

0 400 1000

Stratum 2 (183-549 m)

 

F = 20.4

M = 20.5

U = 17.4

0 400 1000

2004

5 10 15 20 25 30 35

0 400 1000

Stratum 3 (550-1,280 m)

 

F = 22.1

M = 21.8

U = 17.8

5 10 15 20 25 30 35

0 400 1000

Stratum 2 (183-549 m)

 

F = 21.5

M = 20.9

U = 12.9

0 400 1000

2005

5 10 15 20 25 30 35

0 400 1000

Stratum 3 (550-1,280 m)

 

F = 21.4

M = 21.4

U = 12.5

5 10 15 20 25 30 35

0 400 1000

Stratum 2 (183-549 m)

 

F = 21.7

M = 21.1

U = 12.7

0 400 1000

2006

5 10 15 20 25 30 35

0 400 1000

Stratum 3 (550-1,280 m)

 

F = 21.8

M = 22.2

U = 12.8

5 10 15 20 25 30 35

0 400 1000

Stratum 2 (183-549 m)

 

F = 21.8

M = 21.3

U = 14

0 400 1000

2007

5 10 15 20 25 30 35

0 400 1000

Stratum 3 (550-1,280 m)

 

F = 22.2

M = 22.2

U = 13.9

5 10 15 20 25 30 35

0 400 1000

Stratum 2 (183-549 m)

 

F = 21.6

M = 21.4

U = 14.2

0 400 1000

5 10 15 20 25 30 35

2008

5 10 15 20 25 30 35

0 400 1000

Stratum 3 (550-1,280 m)

 

F = 21.9

M = 22.2

U = 14

5 10 15 20 25 30 35

Length (cm)

Frequency

Figure 183. Frequency of lengths by gender in stratum 2 (left column) and stratum 3 (right column) for 

longspine thornyhead from the 2003–2008 surveys. Females are represented by a solid line,

males by a dashed line, and individuals without gender determination by a thick gray line. Mean 

length for each gender classification (F, M, and U) is displayed in each panel.

Stratum 2 (183-549 m) Stratum 3 (550-1,280 m)
197

20 30 40 50 60 70 80

0 10 20 30

Stratum 1 (55-182 m)

 

F = 43.7

M = 43.3

0 10 20 30

2003

20 30 40 50 60 70 80

0 10 20 30

Stratum 2 (183-549 m)

 

F = 47.5

M = 46.5

20 30 40 50 60 70 80

0 10 20 30

Stratum 1 (55-182 m)

 

F = 53.6

M = 52.3

0 10 20 30

2004

20 30 40 50 60 70 80

0 10 20 30

Stratum 2 (183-549 m)

 

F = 55

M = 54.3

20 30 40 50 60 70 80

0 10 20 30

Stratum 1 (55-182 m)

 

F = 37.1

M = 37.7

0 10 20 30

2005

20 30 40 50 60 70 80

0 10 20 30

Stratum 2 (183-549 m)

 

F = 52.1

M = 54.6

20 30 40 50 60 70 80

0 10 20 30

Stratum 1 (55-182 m)

 

F = 45

M = 43.8

0 10 20 30

2006

20 30 40 50 60 70 80

0 10 20 30

Stratum 2 (183-549 m)

 

F = 53

M = 47.5

20 30 40 50 60 70 80

0 10 20 30

Stratum 1 (55-182 m)

 

F = 30

M = 28.8

0 10 20 30

2007

20 30 40 50 60 70 80

0 10 20 30

Stratum 2 (183-549 m)

 

F = 52.6

M = 45.2

20 30 40 50 60 70 80

0 10 20 30

Stratum 1 (55-182 m)

 

F = 40.6

M = 41.7

0 10 20 30

20 30 40 50 60 70 80

2008

20 30 40 50 60 70 80

0 10 20 30

Stratum 2 (183-549 m)

 

F = 44

M = 46

20 30 40 50 60 70 80

Length (cm)

Frequency

Figure 184. Frequency of lengths by gender in stratum 1 (left column) and stratum 2 (right column) for 

Pacific cod from the 2003–2008 surveys. Females are represented by a solid line, males by a 

dashed line. Mean length for each gender classification (F, M) is displayed in each panel.

Stratum 1 (55-182 m) Stratum 2 (183-549 m)
198

10 20 30 40 50

0 40 80

Stratum 2 (183-549 m)

 

F = 28.9

M = 27.6

0 40 80

2003

10 20 30 40 50

0 40 80

Stratum 3 (550-1,280 m)

 

F = 29.3

M = 28.7

10 20 30 40 50

0 40 80

Stratum 2 (183-549 m)

 

0 40 80

2004

10 20 30 40 50

0 40 80

Stratum 3 (550-1,280 m)

 

10 20 30 40 50

0 40 80

Stratum 2 (183-549 m)

 

F = 28.1

M = 24.4

0 40 80

2005

10 20 30 40 50

0 40 80

Stratum 3 (550-1,280 m)

 

F = 31.5

M = 30.3

10 20 30 40 50

0 40 80

Stratum 2 (183-549 m)

 

F = 29.1

M = 27.7

0 40 80

2006

10 20 30 40 50

0 40 80

Stratum 3 (550-1,280 m)

 

F = 32.5

M = 32.4

10 20 30 40 50

0 40 80

Stratum 2 (183-549 m)

 

F = 29.1

M = 29.8

0 40 80

2007

10 20 30 40 50

0 40 80

Stratum 3 (550-1,280 m)

 

F = 34.4

M = 33.7

10 20 30 40 50

0 40 80

Stratum 2 (183-549 m)

 

F = 32.6

M = 31

0 40 80

10 20 30 40 50

2008

10 20 30 40 50

0 40 80

Stratum 3 (550-1,280 m)

 

F = 35.8

M = 35.4

10 20 30 40 50

Length (cm)

Figure 185. Frequency of lengths by gender in stratum 2 (left column) and stratum 3 (right column) for 

Pacific flatnose from the 2003–2008 surveys. Females are represented by a solid line, males by a 

dashed line. Mean length for each gender classification (F, M) is displayed in each panel.

Frequency

Stratum 2 (183-549 m) Stratum 3 (550-1,280 m)
199

0 5 10 15 20 25 30

0 200 400

Stratum 3 (550-1,280 m)

 

F = 11.5

M = 15

U = 3.6

0 200 400

2003

0 5 10 15 20 25 30

0 200 400

Stratum 3 (550-1,280 m)

 

F = 11.5

M = 14.6

U = 4.1

0 200 400

2004

0 5 10 15 20 25 30

0 200 400

Stratum 3 (550-1,280 m)

 

F = 11.8

M = 13.4

U = 5.3

0 200 400

2005

0 5 10 15 20 25 30

0 200 400

Stratum 3 (550-1,280 m)

 

F = 12.6

M = 15.7

U = 5.4

0 200 400

2006

0 5 10 15 20 25 30

0 200 400

Stratum 3 (550-1,280 m)

 

F = 12.1

M = 14.6

U = 5.9

0 200 400

2007

0 5 10 15 20 25 30

0 200 400

Stratum 3 (550-1,280 m)

 

F = 11.1

M = 13.9

U = 4.7

0 200 400

0 5 10 15 20 25 30

2008

Length (cm)

Frequency

Figure 186. Frequency of lengths by gender in stratum 3 for Pacific grenadier from the 2003–2008 

surveys. Females are represented by a solid line, males by a dashed line, and individuals without 

gender determination by a thick gray line. Mean length for each gender classification (F, M, and 

U) is displayed in each panel.

Stratum 3 (550-1,280 m)
200

20 40 60 80

0 100 200

Stratum 1 (55-182 m)

 

F = 44.9

M = 42.6

U = 10.8

0 100 200

2003

20 40 60 80

0 100 200

Stratum 2 (183-549 m)

 

F = 40.5

M = 38.5

U = 33.6

20 40 60 80

0 100 200

Stratum 1 (55-182 m)

 

F = 41.3

M = 39.6

U = 10.5

0 100 200

2004

20 40 60 80

0 100 200

Stratum 2 (183-549 m)

 

F = 34.4

M = 29.6

U = 10.5

20 40 60 80

0 100 200

Stratum 1 (55-182 m)

 

F = 41.6

M = 38.6

U = 14.2

0 100 200

2005

20 40 60 80

0 100 200

Stratum 2 (183-549 m)

 

F = 39.6

M = 35.9

U = 11.8

20 40 60 80

0 100 200

Stratum 1 (55-182 m)

 

F = 32.9

M = 31.8

U = 17.5

0 100 200

2006

20 40 60 80

0 100 200

Stratum 2 (183-549 m)

 

F = 38

M = 35

U = 14.5

20 40 60 80

0 100 200

Stratum 1 (55-182 m)

 

F = 38.2

M = 34.1

U = 18.6

0 100 200

2007

20 40 60 80

0 100 200

Stratum 2 (183-549 m)

 

F = 40.2

M = 36.4

U = 17

20 40 60 80

0 100 200

Stratum 1 (55-182 m)

 

F = 39.1

M = 34.1

U = 14.8

0 100 200

20 40 60 80

2008

20 40 60 80

0 100 200

Stratum 2 (183-549 m)

 

F = 40.8

M = 36.8

U = 25.3

20 40 60 80

Length (cm)

Frequency

Figure 187. Frequency of lengths by gender in stratum 1 (left column) and stratum 2 (right column) for 

Pacific hake from the 2003–2008 surveys. Females are represented by a solid line, males by a 

dashed line, and individuals without gender determination by a thick gray line. Mean length for 

each gender classification (F, M, and U) is displayed in each panel.

Stratum 1 (55-182 m) Stratum 2 (183-549 m)
201

20 30 40 50

0 40 80

Stratum 1 (55-182 m)

 

F = 22.8

M = 23.5

0 40 80

2003

20 30 40 50

0 40 80

Stratum 2 (183-549 m)

 

F = 36.1

M = 35.3

20 30 40 50

0 40 80

Stratum 1 (55-182 m)

 

F = 27.2

M = 25.5

0 40 80

2004

20 30 40 50

0 40 80

Stratum 2 (183-549 m)

 

F = 35.9

M = 35.4

20 30 40 50

0 40 80

Stratum 1 (55-182 m)

 

F = 35.2

M = 30.2

0 40 80

2005

20 30 40 50

0 40 80

Stratum 2 (183-549 m)

 

F = 37.3

M = 34.9

20 30 40 50

0 40 80

Stratum 1 (55-182 m)

 

F = 22

M = 24.4

0 40 80

2006

20 30 40 50

0 40 80

Stratum 2 (183-549 m)

 

F = 35.1

M = 34.8

20 30 40 50

0 40 80

Stratum 1 (55-182 m)

 

F = 25.8

M = 24.9

0 40 80

2007

20 30 40 50

0 40 80

Stratum 2 (183-549 m)

 

F = 32.9

M = 33.7

20 30 40 50

0 40 80

Stratum 1 (55-182 m)

 

F = 22.5

M = 22.7

0 40 80

20 30 40 50

2008

20 30 40 50

0 40 80

Stratum 2 (183-549 m)

 

F = 37.9

M = 37.4

20 30 40 50

Length (cm)

Frequency

Figure 188. Frequency of lengths by gender in stratum 1 (left column) and stratum 2 (right column) for 

Pacific ocean perch from the 2003–2008 surveys. Females are represented by a solid line, males 

by a dashed line. Mean length for each gender classification (F, M) is displayed in each panel.

Stratum 1 (55-182 m) Stratum 2 (183-549 m)
202

5 10 15 20 25 30 35

0 400

Stratum 1 (55-182 m)

 

F = 22.6

M = 19.8

0 400

2003

5 10 15 20 25 30 35

0 400

Stratum 1 (55-182 m)

 

F = 22.2

M = 19.5

0 400

2004

5 10 15 20 25 30 35

0 400

Stratum 1 (55-182 m)

 

F = 21.7

M = 18.9

0 400

2005

5 10 15 20 25 30 35

0 400

Stratum 1 (55-182 m)

 

F = 22.3

M = 19.3

0 400

2006

5 10 15 20 25 30 35

0 400

Stratum 1 (55-182 m)

 

F = 21.7

M = 19

0 400

2007

5 10 15 20 25 30 35

0 400

Stratum 1 (55-182 m)

 

F = 21.8

M = 19

0 400

5 10 15 20 25 30 35

2008

Length (cm)

Frequency

Figure 189. Frequency of lengths by gender in stratum 1 for Pacific sanddab from the 2003–2008 

surveys. Females are represented by a solid line, males by a dashed line. Mean length for each 

gender classification (F, M) is displayed in each panel.

Stratum 1 (55-182 m)
203

10 20 30 40 50 60

0 100 200

Stratum 1 (55-182 m)

 

F = 35.6

M = 31.2

0 100 200

2003

10 20 30 40 50 60

0 100 200

Stratum 1 (55-182 m)

 

F = 35.7

M = 30.4

0 100 200

2004

10 20 30 40 50 60

0 100 200

Stratum 1 (55-182 m)

 

F = 36.1

M = 31.2

0 100 200

2005

10 20 30 40 50 60

0 100 200

Stratum 1 (55-182 m)

 

F = 36.9

M = 31.4

0 100 200

2006

10 20 30 40 50 60

0 100 200

Stratum 1 (55-182 m)

 

F = 36.7

M = 30.9

0 100 200

2007

10 20 30 40 50 60

0 100 200

Stratum 1 (55-182 m)

 

F = 35.3

M = 30

0 100 200

10 20 30 40 50 60

2008

Length (cm)

Frequency

Figure 190. Frequency of lengths by gender in stratum 1 for petrale sole from the 2003–2008 surveys. 

Females are represented by a solid line, males by a dashed line. Mean length for each gender 

classification (F, M) is displayed in each panel.

Stratum 1 (55-182 m)
204

10 15 20

0 20 40 60

Stratum 1 (55-182 m)

 

0 20 40 60

2003

10 15 20

0 20 40 60

Stratum 1 (55-182 m)

 

F = 17.1

M = 13.9

0 20 40 60

2004

10 15 20

0 20 40 60

Stratum 1 (55-182 m)

 

F = 13.2

M = 11.1

U = 13.1

0 20 40 60

2005

10 15 20

0 20 40 60

Stratum 1 (55-182 m)

 

F = 18

M = 15.2

0 20 40 60

2006

10 15 20

0 20 40 60

Stratum 1 (55-182 m)

 

F = 16.9

M = 14.2

0 20 40 60

2007

10 15 20

0 20 40 60

Stratum 1 (55-182 m)

 

F = 15.1

M = 12.6

0 20 40 60

10 15 20

2008

Length (cm)

Frequency

Figure 191. Frequency of lengths by gender in stratum 1 for pygmy rockfish from the 2003–2008 

surveys. Females are represented by a solid line, males by a dashed line, and individuals without 

gender determination by a thick gray line. Mean length for each gender classification (F, M, and 

U) is displayed in each panel.

Stratum 1 (55-182 m)
205

10 20 30 40 50 60 70

0 4 8

Stratum 1 (55-182 m)

 

F = 26.1

M = 23.9

0 4 8

2003

10 20 30 40 50 60 70

0 4 8

Stratum 2 (183-549 m)

 

F = 27.5

M = 28

10 20 30 40 50 60 70

0 4 8

Stratum 1 (55-182 m)

 

M = 35.5

U = 12.5

0 4 8

2004

10 20 30 40 50 60 70

0 4 8

Stratum 2 (183-549 m)

 

F = 33.8

M = 25.5

10 20 30 40 50 60 70

0 4 8

Stratum 1 (55-182 m)

 

F = 18

M = 24

0 4 8

2005

10 20 30 40 50 60 70

0 4 8

Stratum 2 (183-549 m)

 

F = 26.8

M = 26.7

U = 15.7

10 20 30 40 50 60 70

0 4 8

Stratum 1 (55-182 m)

 

F = 18.7

M = 16

0 4 8

2006

10 20 30 40 50 60 70

0 4 8

Stratum 2 (183-549 m)

 

F = 25.2

M = 28.8

U = 15

10 20 30 40 50 60 70

0 4 8

Stratum 1 (55-182 m)

 

F = 20.4

M = 19.6

U = 11.8

0 4 8

2007

10 20 30 40 50 60 70

0 4 8

Stratum 2 (183-549 m)

 

F = 28.7

M = 30.8

10 20 30 40 50 60 70

0 4 8

Stratum 1 (55-182 m)

 

F = 21.3

M = 21

U = 11.5

0 4 8

10 20 30 40 50 60 70

2008

10 20 30 40 50 60 70

0 4 8

Stratum 2 (183-549 m)

 

F = 26.9

M = 28.5

10 20 30 40 50 60 70

Length (cm)

Frequency

Figure 192. Frequency of lengths by gender in stratum 1 (left column) and stratum 2 (right column) for 

redbanded rockfish from the 2003–2008 surveys. Females are represented by a solid line, males 

by a dashed line, and individuals without gender determination by a thick gray line. Mean length 

for each gender classification (F, M, and U) is displayed in each panel.

0

Stratum 1 (55-182 m) Stratum 2 (183-549 m)
206

10 15 20 25 30 35 40

0 50 100

Stratum 1 (55-182 m)

 

F = 28.6

M = 21.5

U = 11.1

0 50 100

2003

10 15 20 25 30 35 40

0 50 100

Stratum 1 (55-182 m)

 

F = 25.1

M = 22.5

U = 16.2

0 50 100

2004

10 15 20 25 30 35 40

0 50 100

Stratum 1 (55-182 m)

 

F = 22.5

M = 20.1

U = 15.5

0 50 100

2005

10 15 20 25 30 35 40

0 50 100

Stratum 1 (55-182 m)

 

F = 27.5

M = 22.2

0 50 100

2006

10 15 20 25 30 35 40

0 50 100

Stratum 1 (55-182 m)

 

F = 25.5

M = 26

U = 10.5

0 50 100

2007

10 15 20 25 30 35 40

0 50 100

Stratum 1 (55-182 m)

 

F = 30.4

M = 26.5

U = 8.5

0 50 100

10 15 20 25 30 35 40

2008

Length (cm)

Frequency

Figure 193. Frequency of lengths by gender in stratum 1 for redstripe rockfish from the 2003–2008 

surveys. Females are represented by a solid line, males by a dashed line, and individuals without 

gender determination by a thick gray line. Mean length for each gender classification (F, M, and 

U) is displayed in each panel.

Stratum 1 (55-182 m)
207

10 20 30 40 50

0 200 500

Stratum 1 (55-182 m)

 

F = 28.8

M = 26.5

0 200 500

2003

10 20 30 40 50

0 200 500

Stratum 2 (183-549 m)

 

F = 30.5

M = 29.1

10 20 30 40 50

0 200 500

Stratum 1 (55-182 m)

 

F = 27.4

M = 25.3

0 200 500

2004

10 20 30 40 50

0 200 500

Stratum 2 (183-549 m)

 

F = 29.4

M = 28.3

10 20 30 40 50

0 200 500

Stratum 1 (55-182 m)

 

F = 27.4

M = 25.7

0 200 500

2005

10 20 30 40 50

0 200 500

Stratum 2 (183-549 m)

 

F = 29.2

M = 27.9

10 20 30 40 50

0 200 500

Stratum 1 (55-182 m)

 

F = 27.8

M = 26.3

0 200 500

2006

10 20 30 40 50

0 200 500

Stratum 2 (183-549 m)

 

F = 29.5

M = 28.6

10 20 30 40 50

0 200 500

Stratum 1 (55-182 m)

 

F = 27.7

M = 26.2

0 200 500

2007

10 20 30 40 50

0 200 500

Stratum 2 (183-549 m)

 

F = 29.7

M = 28.4

10 20 30 40 50

0 200 500

Stratum 1 (55-182 m)

 

F = 27.9

M = 25.8

0 200 500

10 20 30 40 50

2008

10 20 30 40 50

0 200 500

Stratum 2 (183-549 m)

 

F = 30.5

M = 29.1

10 20 30 40 50

Length (cm)

Frequency

Figure 194. Frequency of lengths by gender in stratum 1 (left column) and stratum 2 (right column) for 

rex sole from the 2003–2008 surveys. Females are represented by a solid line, males by a dashed 

line. Mean length for each gender classification (F, M) is displayed in each panel.

Stratum 1 (55-182 m) Stratum 2 (183-549 m)
208

10 15 20 25 30 35

0 20 40 60

Stratum 1 (55-182 m)

 

F = 23.6

M = 24

0 20 40 60

2003

10 15 20 25 30 35

0 20 40 60

Stratum 2 (183-549 m)

 

F = 23.4

M = 23.9

10 15 20 25 30 35

0 20 40 60

Stratum 1 (55-182 m)

 

F = 23.8

M = 24.5

0 20 40 60

2004

10 15 20 25 30 35

0 20 40 60

Stratum 2 (183-549 m)

 

F = 23

M = 22.7

10 15 20 25 30 35

0 20 40 60

Stratum 1 (55-182 m)

 

F = 22.7

M = 23

0 20 40 60

2005

10 15 20 25 30 35

0 20 40 60

Stratum 2 (183-549 m)

 

F = 23.5

M = 24

10 15 20 25 30 35

0 20 40 60

Stratum 1 (55-182 m)

 

F = 21

M = 21.4

0 20 40 60

2006

10 15 20 25 30 35

0 20 40 60

Stratum 2 (183-549 m)

 

F = 22.5

M = 23.4

10 15 20 25 30 35

0 20 40 60

Stratum 1 (55-182 m)

 

F = 20.3

M = 20.4

0 20 40 60

2007

10 15 20 25 30 35

0 20 40 60

Stratum 2 (183-549 m)

 

F = 22.3

M = 23

10 15 20 25 30 35

0 20 40 60

Stratum 1 (55-182 m)

 

F = 21.2

M = 21.7

0 20 40 60

10 15 20 25 30 35

2008

10 15 20 25 30 35

0 20 40 60

Stratum 2 (183-549 m)

 

F = 22.8

M = 23.3

10 15 20 25 30 35

Length (cm)

Frequency

Figure 195. Frequency of lengths by gender in stratum 1 (left column) and stratum 2 (right column) for 

rosethorn rockfish from the 2003–2008 surveys. Females are represented by a solid line, males 

by a dashed line. Mean length for each gender classification (F, M) is displayed in each panel.

Stratum 1 (55-182 m) Stratum 2 (183-549 m)
209

10 20 30 40 50 60 70

0 5 10 15

Stratum 1 (55-182 m)

 

F = 22.6

M = 22.5

0 5 10 15

2003

10 20 30 40 50 60 70

0 5 10 15

Stratum 2 (183-549 m)

 

F = 31.3

M = 32.8

10 20 30 40 50 60 70

0 5 10 15

Stratum 1 (55-182 m)

 

F = 22.8

M = 23.6

0 5 10 15

2004

10 20 30 40 50 60 70

0 5 10 15

Stratum 2 (183-549 m)

 

F = 54.9

M = 52.1

10 20 30 40 50 60 70

0 5 10 15

Stratum 1 (55-182 m)

 

F = 23.2

M = 23.5

0 5 10 15

2005

10 20 30 40 50 60 70

0 5 10 15

Stratum 2 (183-549 m)

 

F = 48.9

M = 47.6

10 20 30 40 50 60 70

0 5 10 15

Stratum 1 (55-182 m)

 

F = 25.4

M = 27

0 5 10 15

2006

10 20 30 40 50 60 70

0 5 10 15

Stratum 2 (183-549 m)

 

F = 40.2

M = 42.4

10 20 30 40 50 60 70

0 5 10 15

Stratum 1 (55-182 m)

 

F = 23.1

M = 27.3

0 5 10 15

2007

10 20 30 40 50 60 70

0 5 10 15

Stratum 2 (183-549 m)

 

F = 34

M = 33.6

10 20 30 40 50 60 70

0 5 10 15

Stratum 1 (55-182 m)

 

F = 23.5

M = 24.8

0 5 10 15

10 20 30 40 50 60 70

2008

10 20 30 40 50 60 70

0 5 10 15

Stratum 2 (183-549 m)

 

F = 37.4

M = 41.8

10 20 30 40 50 60 70

Length (cm)

Figure 196. Frequency of lengths by gender in stratum 1 (left column) and stratum 2 (right column) for 

rougheye rockfish from the 2003–2008 surveys. Females are represented by a solid line, males 

by a dashed line. Mean length for each gender classification (F, M) is displayed in each panel.

Frequency

Stratum 1 (55-182 m) Stratum 2 (183-549 m)
210

20 40 60 80

0 50 150

Stratum 1 (55-182 

 

F = 50.7

M = 47.4

U = 21.8

0 50 150

2003

20 40 60 80

0 50 150

Stratum 2 (183-549 

 

F = 53.9

M = 49.8

U = 63

20 40 60 80

0 50 150

Stratum 3 (550-1,2 

 

F = 59.3

M = 54.7

U = 57.8

20 40 60 80

0 50 150

Stratum 1 (55-182 

 

F = 43.2

M = 42.2

0 50 150

2004

20 40 60 80

0 50 150

Stratum 2 (183-549 

 

F = 56

M = 52.2

U = 57.3

20 40 60 80

0 50 150

Stratum 3 (550-1,2 

 

F = 57.4

M = 53.5

U = 57

20 40 60 80

0 50 150

Stratum 1 (55-182 

 

F = 45.3

M = 43.8

0 50 150

2005

20 40 60 80

0 50 150

Stratum 2 (183-549 

 

F = 55.9

M = 50.8

U = 45

20 40 60 80

0 50 150

Stratum 3 (550-1,2 

 

F = 57.5

M = 53

U = 52.4

20 40 60 80

0 50 150

Stratum 1 (55-182 

 

F = 53.3

M = 48

0 50 150

2006

20 40 60 80

0 50 150

Stratum 2 (183-549 

 

F = 59.7

M = 54.5

20 40 60 80

0 50 150

Stratum 3 (550-1,2 

 

F = 59.6

M = 54.4

U = 62

20 40 60 80

0 50 150

Stratum 1 (55-182 

 

F = 54.1

M = 52.4

0 50 150

2007

20 40 60 80

0 50 150

Stratum 2 (183-549 

 

F = 60.3

M = 54.6

U = 59

20 40 60 80

0 50 150

Stratum 3 (550-1,2 

 

F = 59.6

M = 55.1

U = 49.4

20 40 60 80

0 50 150

Stratum 1 (55-182 

 

F = 44.8

M = 42.8

U = 23.5

0 50 150

20 40 60 80

2008

20 40 60 80

0 50 150

Stratum 2 (183-549 

 

F = 59.8

M = 54.7

U = 53.1

20 40 60 80 20 40 60 80

0 50 150

Stratum 3 (550-1,2 

 

F = 60.5

M = 55.2

U = 49.6

20 40 60 80

Length (cm)

Frequency

Figure 197. Frequency of lengths by gender in stratum 1 (left column), stratum 2 (middle column), and 

stratum 3 (right column) for sablefish from the 2003–2008 surveys. Females are represented by a 

solid line, males by a dashed line, and individuals without gender determination by a thick gray 

line. Mean length for each gender classification (F, M, and U) is displayed in each panel.

Stratum 1 (55-182 m) Stratum 2 (183-549 m) Stratum 3 (550-1,280 m)
211

5 10 15 20 25 30 35 40

0 40 80 120

Stratum 1 (55-182 m)

 

F = 23.6

M = 21

0 40 80 120

2003

5 10 15 20 25 30 35 40

0 40 80 120

Stratum 2 (183-549 m)

 

F = 28.5

M = 24.9

5 10 15 20 25 30 35 40

0 40 80 120

Stratum 1 (55-182 m)

 

F = 24.5

M = 21

0 40 80 120

2004

5 10 15 20 25 30 35 40

0 40 80 120

Stratum 2 (183-549 m)

 

F = 27.5

M = 23.4

5 10 15 20 25 30 35 40

0 40 80 120

Stratum 1 (55-182 m)

 

F = 23.7

M = 21.3

0 40 80 120

2005

5 10 15 20 25 30 35 40

0 40 80 120

Stratum 2 (183-549 m)

 

F = 25.9

M = 23.2

5 10 15 20 25 30 35 40

0 40 80 120

Stratum 1 (55-182 m)

 

F = 22.4

M = 19.9

0 40 80 120

2006

5 10 15 20 25 30 35 40

0 40 80 120

Stratum 2 (183-549 m)

 

F = 26.3

M = 22.9

5 10 15 20 25 30 35 40

0 40 80 120

Stratum 1 (55-182 m)

 

F = 22.3

M = 20.7

0 40 80 120

2007

5 10 15 20 25 30 35 40

0 40 80 120

Stratum 2 (183-549 m)

 

F = 26.3

M = 23

5 10 15 20 25 30 35 40

0 40 80 120

Stratum 1 (55-182 m)

 

F = 25

M = 22.2

0 40 80 120

5 10 15 20 25 30 35 40

2008

5 10 15 20 25 30 35 40

0 40 80 120

Stratum 2 (183-549 m)

 

F = 25.5

M = 22.4

5 10 15 20 25 30 35 40

Length (cm)

Frequency

Figure 198. Frequency of lengths by gender in stratum 1 (left column) and stratum 2 (right column) for 

sharpchin rockfish from the 2003–2008 surveys. Females are represented by a solid line, males 

by a dashed line. Mean length for each gender classification (F, M) is displayed in each panel.

Stratum 1 (55-182 m) Stratum 2 (183-549 m)
212

10 15 20 25 30

0 100 200

Stratum 1 (55-182 m)

 

F = 21.4

M = 15.6

U = 10.3

0 100 200

2003

10 15 20 25 30

0 100 200

Stratum 2 (183-549 m)

 

F = 21.4

M = 20.6

10 15 20 25 30

0 100 200

Stratum 1 (55-182 m)

 

F = 16

M = 15.1

U = 12.2

0 100 200

2004

10 15 20 25 30

0 100 200

Stratum 2 (183-549 m)

 

F = 19.4

M = 18.6

10 15 20 25 30

0 100 200

Stratum 1 (55-182 m)

 

F = 17.4

M = 16.5

U = 10.1

0 100 200

2005

10 15 20 25 30

0 100 200

Stratum 2 (183-549 m)

 

F = 20.9

M = 19.4

10 15 20 25 30

0 100 200

Stratum 1 (55-182 m)

 

F = 17.2

M = 16.8

U = 14.2

0 100 200

2006

10 15 20 25 30

0 100 200

Stratum 2 (183-549 m)

 

F = 15.7

M = 15.7

U = 20.3

10 15 20 25 30

0 100 200

Stratum 1 (55-182 m)

 

F = 18.2

M = 17.1

U = 11.4

0 100 200

2007

10 15 20 25 30

0 100 200

Stratum 2 (183-549 m)

 

F = 19.8

M = 19.6

10 15 20 25 30

0 100 200

Stratum 1 (55-182 m)

 

F = 19.3

M = 18.8

U = 12

0 100 200

10 15 20 25 30

2008

10 15 20 25 30

0 100 200

Stratum 2 (183-549 m)

 

F = 17.9

M = 17.8

10 15 20 25 30

Length (cm)

Frequency

Figure 199. Frequency of lengths by gender in stratum 1 (left column) and stratum 2 (right column) for 

shortbelly rockfish from the 2003–2008 surveys. F emales are represented by a solid line, males 

by a dashed line, and individuals without gender determination by a thick gray line. Mean length 

for each gender classification (F, M, and U) is displayed in each panel.

Stratum 1 (55-182 m) Stratum 2 (183-549 m)
213

0 20 40 60 80

0 200 400

Stratum 2 (183-549 m)

 

F = 24.8

U = 24

0 200 400

2003

0 20 40 60 80

0 200 400

Stratum 3 (550-1,280 m)

 

U = 39.6

0 20 40 60 80

0 200 400

Stratum 2 (183-549 m)

 

F = 26.7

M = 24.2

U = 22.1

0 200 400

2004

0 20 40 60 80

0 200 400

Stratum 3 (550-1,280 m)

 

F = 42

M = 38.7

U = 36

0 20 40 60 80

0 200 400

Stratum 2 (183-549 m)

 

F = 26.1

M = 25

U = 14.2

0 200 400

2005

0 20 40 60 80

0 200 400

Stratum 3 (550-1,280 m)

 

F = 38

M = 36.3

U = 23.5

0 20 40 60 80

0 200 400

Stratum 2 (183-549 m)

 

F = 26.1

M = 25.5

U = 14.7

0 200 400

2006

0 20 40 60 80

0 200 400

Stratum 3 (550-1,280 m)

 

F = 41.5

M = 38.4

0 20 40 60 80

0 200 400

Stratum 2 (183-549 m)

 

F = 26.1

M = 25.2

U = 15.2

0 200 400

2007

0 20 40 60 80

0 200 400

Stratum 3 (550-1,280 m)

 

F = 41.1

M = 38.8

U = 20.1

0 20 40 60 80

0 200 400

Stratum 2 (183-549 m)

 

F = 26.6

M = 25.8

U = 16.2

0 200 400

0 20 40 60 80

2008

0 20 40 60 80

0 200 400

Stratum 3 (550-1,280 m)

 

F = 43.1

M = 39.6

U = 40

0 20 40 60 80

Length (cm)

Frequency

Figure 200. Frequency of lengths by gender in stratum 2 (left column) and stratum 3 (right column) for 

shortspine thornyhead from the 2003–2008 surveys. Females are represented by a solid line,

males by a dashed line, and individuals without gender determination by a thick gray line. Mean 

length for each gender classification (F, M, and U) is displayed in each panel.

Stratum 2 (183-549 m) Stratum 3 (550-1,280 m)
214

20 40 60 80 100 120

0 40 80

Stratum 1 (55-182 m)

 

F = 57.8

M = 65.5

0 40 80

2003

20 40 60 80 100 120

0 40 80

Stratum 2 (183-549 m)

 

F = 54.2

M = 56.8

20 40 60 80 100 120

0 40 80

Stratum 1 (55-182 m)

 

F = 53.4

M = 58

0 40 80

2004

20 40 60 80 100 120

0 40 80

Stratum 2 (183-549 m)

 

F = 56.4

M = 62.2

20 40 60 80 100 120

0 40 80

Stratum 1 (55-182 m)

 

F = 52.4

M = 54.9

0 40 80

2005

20 40 60 80 100 120

0 40 80

Stratum 2 (183-549 m)

 

F = 54.4

M = 58.7

20 40 60 80 100 120

0 40 80

Stratum 1 (55-182 m)

 

F = 50.6

M = 53.5

0 40 80

2006

20 40 60 80 100 120

0 40 80

Stratum 2 (183-549 m)

 

F = 54.7

M = 56.5

20 40 60 80 100 120

0 40 80

Stratum 1 (55-182 m)

 

F = 48.1

M = 51.3

0 40 80

2007

20 40 60 80 100 120

0 40 80

Stratum 2 (183-549 m)

 

F = 50

M = 53.9

20 40 60 80 100 120

0 40 80

Stratum 1 (55-182 m)

 

F = 50.2

M = 53.2

0 40 80

20 40 60 80 100 120

2008

20 40 60 80 100 120

0 40 80

Stratum 2 (183-549 m)

 

F = 57.2

M = 58.4

20 40 60 80 100 120

Length (cm)

Frequency

Figure 201. Frequency of lengths by gender in stratum 1 (left column) and stratum 2 (right column) for 

spiny dogfish from the 2003–2008 surveys. Females are represented by a solid line, males by a 

dashed line. Mean length for each gender classification (F, M) is displayed in each panel.

Stratum 1 (55-182 m) Stratum 2 (183-549 m)
215

10 20 30 40

0 100 200

Stratum 1 (55-182 m)

 

F = 16.6

M = 15.4

U = 11.7

0 100 200

2003

10 20 30 40

0 100 200

Stratum 2 (183-549 m)

 

F = 20.9

M = 19.7

U = 12

10 20 30 40

0 100 200

Stratum 1 (55-182 m)

 

F = 14

M = 14.7

U = 10.4

0 100 200

2004

10 20 30 40

0 100 200

Stratum 2 (183-549 m)

 

F = 20

M = 19.3

U = 9.8

10 20 30 40

0 100 200

Stratum 1 (55-182 m)

 

F = 17.8

M = 16.9

U = 8.6

0 100 200

2005

10 20 30 40

0 100 200

Stratum 2 (183-549 m)

 

F = 19.4

M = 18.9

U = 10.1

10 20 30 40

0 100 200

Stratum 1 (55-182 m)

 

F = 16.1

M = 15.4

U = 9

0 100 200

2006

10 20 30 40

0 100 200

Stratum 2 (183-549 m)

 

F = 20.7

M = 20.9

U = 9.7

10 20 30 40

0 100 200

Stratum 1 (55-182 m)

 

F = 18.6

M = 18.3

U = 10

0 100 200

2007

10 20 30 40

0 100 200

Stratum 2 (183-549 m)

 

F = 21.6

M = 20.7

U = 10.1

10 20 30 40

0 100 200

Stratum 1 (55-182 m)

 

F = 15.7

M = 13.7

U = 10.8

0 100 200

10 20 30 40

2008

10 20 30 40

0 100 200

Stratum 2 (183-549 m)

 

F = 21.3

M = 20.5

U = 10.3

10 20 30 40

Length (cm)

Frequency

Figure 202. Frequency of lengths by gender in stratum 1 (left column) and stratum 2 (right column) for 

splitnose rockfish from the 2003–2008 surveys. Females are represented by a solid line, males by 

a dashed line, and individuals without gender determination by a thick gray line. Mean length for 

each gender classification (F, M, and U) is displayed in each panel.

Stratum 1 (55-182 m) Stratum 2 (183-549 m)
216

10 20 30 40 50 60

0 100 200

Stratum 1 (55-182 m)

 

F = 45.9

M = 42

0 100 200

2003

10 20 30 40 50 60

0 100 200

Stratum 2 (183-549 m)

 

F = 48.7

M = 43.7

10 20 30 40 50 60

0 100 200

Stratum 1 (55-182 m)

 

0 100 200

2004

10 20 30 40 50 60

0 100 200

Stratum 2 (183-549 m)

 

10 20 30 40 50 60

0 100 200

Stratum 1 (55-182 m)

 

F = 18.8

M = 16.6

0 100 200

2005

10 20 30 40 50 60

0 100 200

Stratum 2 (183-549 m)

 

F = 19.5

M = 16.6

10 20 30 40 50 60

0 100 200

Stratum 1 (55-182 m)

 

F = 19.6

M = 17.6

0 100 200

2006

10 20 30 40 50 60

0 100 200

Stratum 2 (183-549 m)

 

F = 19

M = 16.3

10 20 30 40 50 60

0 100 200

Stratum 1 (55-182 m)

 

F = 17.9

M = 16.8

0 100 200

2007

10 20 30 40 50 60

0 100 200

Stratum 2 (183-549 m)

 

F = 19.2

M = 16.6

10 20 30 40 50 60

0 100 200

Stratum 1 (55-182 m)

 

F = 18.3

M = 17.1

0 100 200

10 20 30 40 50 60

2008

10 20 30 40 50 60

0 100 200

Stratum 2 (183-549 m)

 

F = 20

M = 17

10 20 30 40 50 60

Length (cm)

Frequency

Figure 203. Frequency of lengths by gender in stratum 1 (left column) and stratum 2 (right column) for 

spotted ratfish from the 2003–2008 surveys. Females are represented by a solid line, males by a 

dashed line. Mean length for each gender classification (F, M) is displayed in each panel.

Stratum 1 (55-182 m) Stratum 2 (183-549 m)
217

10 15 20 25

0 10 30

Stratum 1 (55-182 m)

 

F = 20.2

M = 18.4

0 10 30

2003

10 15 20 25

0 10 30

Stratum 1 (55-182 m)

 

F = 23

M = 16.8

U = 24.1

0 10 30

2004

10 15 20 25

0 10 30

Stratum 1 (55-182 m)

 

F = 16.6

M = 13.7

U = 11

0 10 30

2005

10 15 20 25

0 10 30

Stratum 1 (55-182 m)

 

F = 19.4

M = 16.1

0 10 30

2006

10 15 20 25

0 10 30

Stratum 1 (55-182 m)

 

F = 17.3

M = 15.3

U = 13.6

0 10 30

2007

10 15 20 25

0 10 30

Stratum 1 (55-182 m)

 

F = 17.4

M = 15.5

0 10 30

10 15 20 25

2008

Length (cm)

Frequency

Figure 204. Frequency of lengths by gender in stratum 1 for squarespot rockfish from the 2003–2008 

surveys. Females are represented by a solid line, males by a dashed line, and individuals without 

gender determination by a thick gray line. Mean length for each gender classification (F, M, and 

U) is displayed in each panel.

Stratum 1 (55-182 m)
218

5 10 15 20 25 30 35

0 100 200

Stratum 1 (55-182 m)

 

F = 16.9

M = 14.4

U = 12

0 100 200

2003

5 10 15 20 25 30 35

0 100 200

Stratum 2 (183-549 m)

 

F = 20.6

M = 16.2

U = 13.3

5 10 15 20 25 30 35

0 100 200

Stratum 1 (55-182 m)

 

F = 15.6

M = 12.5

U = 11

0 100 200

2004

5 10 15 20 25 30 35

0 100 200

Stratum 2 (183-549 m)

 

F = 20.8

M = 15.5

5 10 15 20 25 30 35

0 100 200

Stratum 1 (55-182 m)

 

F = 14.8

M = 12.4

U = 10.3

0 100 200

2005

5 10 15 20 25 30 35

0 100 200

Stratum 2 (183-549 m)

 

F = 20.5

M = 16

U = 11.6

5 10 15 20 25 30 35

0 100 200

Stratum 1 (55-182 m)

 

F = 15.8

M = 14

U = 10.2

0 100 200

2006

5 10 15 20 25 30 35

0 100 200

Stratum 2 (183-549 m)

 

F = 21.6

M = 16.9

U = 11

5 10 15 20 25 30 35

0 100 200

Stratum 1 (55-182 m)

 

F = 15.8

M = 14.3

U = 10.8

0 100 200

2007

5 10 15 20 25 30 35

0 100 200

Stratum 2 (183-549 m)

 

F = 21.3

M = 16.4

U = 14

5 10 15 20 25 30 35

0 100 200

Stratum 1 (55-182 m)

 

F = 16.5

M = 14.2

U = 10.9

0 100 200

5 10 15 20 25 30 35

2008

5 10 15 20 25 30 35

0 100 200

Stratum 2 (183-549 m)

 

F = 20.8

M = 15.6

U = 12

5 10 15 20 25 30 35

Length (cm)

Frequency

Figure 205. Frequency of lengths by gender in stratum 1 (left column) and stratum 2 (right column) for 

stripetail rockfish from the 2003–2008 surveys. Females are represented by a solid line, males by 

a dashed line, and individuals without gender determination by a thick gray line. Mean length for 

each gender classification (F, M, and U) is displayed in each panel.

Stratum 1 (55-182 m) Stratum 2 (183-549 m)
219

10 15 20 25

0 20 40

Stratum 1 (55-182 m)

 

0 20 40

2003

10 15 20 25

0 20 40

Stratum 2 (183-549 m)

 

10 15 20 25

0 20 40

Stratum 1 (55-182 m)

 

F = 15.5

M = 14.9

U = 14

0 20 40

2004

10 15 20 25

0 20 40

Stratum 2 (183-549 m)

 

F = 17.3

M = 16

10 15 20 25

0 20 40

Stratum 1 (55-182 m)

 

F = 13.7

M = 14

0 20 40

2005

10 15 20 25

0 20 40

Stratum 2 (183-549 m)

 

10 15 20 25

0 20 40

Stratum 1 (55-182 m)

 

F = 15

M = 14.5

U = 11.8

0 20 40

2006

10 15 20 25

0 20 40

Stratum 2 (183-549 m)

 

F = 21.6

M = 20.4

10 15 20 25

0 20 40

Stratum 1 (55-182 m)

 

F = 15.9

M = 15.2

U = 10.7

0 20 40

2007

10 15 20 25

0 20 40

Stratum 2 (183-549 m)

 

F = 19

M = 15.5

10 15 20 25

0 20 40

Stratum 1 (55-182 m)

 

F = 15.1

M = 14.5

U = 8.8

0 20 40

10 15 20 25

2008

10 15 20 25

0 20 40

Stratum 2 (183-549 m)

 

F = 17.2

M = 16.5

U = 16

10 15 20 25

Length (cm)

Frequency

Figure 206. Frequency of lengths by gender in stratum 1 (left column) and stratum 2 (right column) for 

swordspine rockfish (Sebastes ensifer) from the 2003–2008 surveys. Females are represented by 

a solid line, males by a dashed line, and individuals without gender determination by a thick gray 

line. Mean length for each gender classification (F, M, and U) is displayed in each panel.

Stratum 1 (55-182 m) Stratum 2 (183-549 m)
220

20 30 40 50

0 5 10

Stratum 1 (55-182 m)

 

F = 39.6

M = 37

0 5 10

03

20 30 40 50

0 5 10

Stratum 2 (183-549 m)

 

F = 45

M = 42.9

20 30 40 50

0 5 10

Stratum 1 (55-182 m)

 

F = 24

M = 23.5

0 5 10

04

20 30 40 50

0 5 10

Stratum 2 (183-549 m)

 

F = 36.2

M = 37.3

20 30 40 50

0 5 10

Stratum 1 (55-182 m)

 

F = 31.4

M = 28.7

0 5 10

05

20 30 40 50

0 5 10

Stratum 2 (183-549 m)

 

F = 42.5

M = 40.5

20 30 40 50

0 5 10

Stratum 1 (55-182 m)

 

F = 37.9

M = 35.7

0 5 10

06

20 30 40 50

0 5 10

Stratum 2 (183-549 m)

 

F = 48

M = 42.6

20 30 40 50

0 5 10

Stratum 1 (55-182 m)

 

F = 42.1

M = 42.1

0 5 10

07

20 30 40 50

0 5 10

Stratum 2 (183-549 m)

 

F = 46.1

M = 41.8

20 30 40 50

0 5 10

Stratum 1 (55-182 m)

 

F = 41.8

M = 37.5

0 5 10

20 30 40 50

08

20 30 40 50

0 5 10

Stratum 2 (183-549 m)

 

F = 46.3

M = 43.3

20 30 40 50

Length (cm)

Frequency

Figure 207. Frequency of lengths by gender in stratum 1 (left column) and stratum 2 (right column) for 

widow rockfish from the 2003–2008 surveys. Females are represented by a solid line, males by a 

dashed line. Mean length for each gender classification (F, M) is displayed in each panel.

2003

2004

2008

2007

2006

2005

Stratum 1 (55-182 m) Stratum 2 (183-549 m)
221

10 20 30 40 50 60

0 20 60

Stratum 1 (55-182 m)

 

F = 43.5

M = 41.2

0 20 60

2003

10 20 30 40 50 60

0 20 60

Stratum 2 (183-549 m)

 

F = 47.6

M = 43.4

10 20 30 40 50 60

0 20 60

Stratum 1 (55-182 m)

 

F = 41.4

M = 39.9

0 20 60

2004

10 20 30 40 50 60

0 20 60

Stratum 2 (183-549 m)

 

10 20 30 40 50 60

0 20 60

Stratum 1 (55-182 m)

 

F = 40.4

M = 38.8

0 20 60

2005

10 20 30 40 50 60

0 20 60

Stratum 2 (183-549 m)

 

F = 49.2

M = 46.1

10 20 30 40 50 60

0 20 60

Stratum 1 (55-182 m)

 

F = 47.7

M = 43.4

0 20 60

2006

10 20 30 40 50 60

0 20 60

Stratum 2 (183-549 m)

 

F = 44

M = 44.8

10 20 30 40 50 60

0 20 60

Stratum 1 (55-182 m)

 

F = 44.9

M = 42.8

0 20 60

2007

10 20 30 40 50 60

0 20 60

Stratum 2 (183-549 m)

 

F = 47.4

M = 45

10 20 30 40 50 60

0 20 60

Stratum 1 (55-182 m)

 

F = 40.7

M = 39.6

0 20 60

10 20 30 40 50 60

2008

10 20 30 40 50 60

0 20 60

Stratum 2 (183-549 m)

 

F = 46.8

M = 44.3

10 20 30 40 50 60

Length (cm)

Frequency

Figure 208. Frequency of lengths by gender in stratum 1 (left column) and stratum 2 (right column) for 

yellowtail rockfish from the 2003–2008 surveys. Females are represented by a solid line, males 

by a dashed line. Mean length for each gender classification (F, M) is displayed in each panel.

Stratum 1 (55-182 m) Stratum 2 (183-549 m)
222

Figure 209. Age distribution for arrowtooth flounder sampled during the surveys from 2003 to 2006.

2003

 

0 5 10 15 20 25

0.0 0.2 0.4 0.6

2004

 

0 5 10 15 20 25

0.0 0.2 0.4 0.6

2005

 

0 5 10 15 20 25

0.0 0.2 0.4 0.6

2006

 

0 5 10 15 20 25

0.0 0.2 0.4 0.6

Age (yrs)

Proportion
223

2003

 

0 10 20 30 40 50 60

0.00 0.05 0.10 0.15 0.20

2004

 

0 10 20 30 40 50 60

0.00 0.05 0.10 0.15 0.20

2005

 

0 10 20 30 40 50 60

0.00 0.05 0.10 0.15 0.20

2006

 

0 10 20 30 40 50 60

0.00 0.05 0.10 0.15 0.20

2007

 

0 10 20 30 40 50 60

0.00 0.05 0.10 0.15 0.20

2008

 

0 10 20 30 40 50 60

0.00 0.05 0.10 0.15 0.20

Age (yrs)

Proportion

Figure 210. Age distribution for canary rockfish sampled during the surveys from 2003 to 2008.
224

2004

 

0 5 10 15 20

0.0 0.1 0.2 0.3 0.4

Age (yrs)

Proportion

Figure 211. Age distribution for chilipepper rockfish sampled during the 2004 survey.
225

2003

 

0 20 40 60 80

0.0 0.1 0.2 0.3 0.4

2004

 

0 20 40 60 80

0.0 0.1 0.2 0.3 0.4

2005

 

0 20 40 60 80

0.0 0.1 0.2 0.3 0.4

2006

 

0 20 40 60 80

0.0 0.1 0.2 0.3 0.4

2007

 

0 20 40 60 80

0.0 0.1 0.2 0.3 0.4

2008

 

0 20 40 60 80

0.0 0.1 0.2 0.3 0.4

Age (yrs)

Proportion

Figure 212. Age distribution for darkblotched rockfish sampled during the surveys from 2003 to 2008.
226

2003

 

0 10 20 30 40 50

0.00 0.05 0.10 0.15

2004

 

0 10 20 30 40 50

0.00 0.05 0.10 0.15

Age (yrs)

Proportion

Figure 213. Age distribution for Dover sole sampled during the surveys from 2003 to 2004.
227

2003

 

0 5 10 15

0.00 0.10 0.20 0.30

2004

 

0 5 10 15

0.00 0.10 0.20 0.30

2005

 

0 5 10 15

0.00 0.10 0.20 0.30

Age (yrs)

Proportion

Figure 214. Age distribution for English sole sampled during the surveys from 2003 to 2005.
228

2003

 

0 10 20 30 40 50

0.00 0.10 0.20 0.30

2004

 

0 10 20 30 40 50

0.00 0.10 0.20 0.30

2005

 

0 10 20 30 40 50

0.00 0.10 0.20 0.30

2006

 

0 10 20 30 40 50

0.00 0.10 0.20 0.30

2007

 

0 10 20 30 40 50

0.00 0.10 0.20 0.30

2008

 

0 10 20 30 40 50

0.00 0.10 0.20 0.30

Age (yrs)

Proportion

Figure 215. Age distribution for greenstriped rockfish sampled during the surveys from 2003 to 2008.
229

2003

 

0 2 4 6 8 10 12 14

0.0 0.1 0.2 0.3 0.4 0.5

2004

 

0 2 4 6 8 10 12 14

0.0 0.1 0.2 0.3 0.4 0.5

2005

 

0 2 4 6 8 10 12 14

0.0 0.1 0.2 0.3 0.4 0.5

2006

 

0 2 4 6 8 10 12 14

0.0 0.1 0.2 0.3 0.4 0.5

2007

 

0 2 4 6 8 10 12 14

0.0 0.1 0.2 0.3 0.4 0.5

2008

 

0 2 4 6 8 10 12 14

0.0 0.1 0.2 0.3 0.4 0.5

Age (yrs)

2003

 

0 2 4 6 8 10 12 14

0.0 0.1 0.2 0.3 0.4 0.5

2004

 

0 2 4 6 8 10 12 14

0.0 0.1 0.2 0.3 0.4 0.5

2005

 

0 2 4 6 8 10 12 14

0.0 0.1 0.2 0.3 0.4 0.5

2006

 

0 2 4 6 8 10 12 14

0.0 0.1 0.2 0.3 0.4 0.5

2007

 

0 2 4 6 8 10 12 14

0.0 0.1 0.2 0.3 0.4 0.5

2008

 

0 2 4 6 8 10 12 14

0.0 0.1 0.2 0.3 0.4 0.5

Age (yrs)

Proportion

Figure 216. Age distribution for lingcod sampled during the surveys from 2003 to 2008.
230

2003

 

0 10 20 30 40 50 60 70

0.00 0.05 0.10 0.15

2004

 

0 10 20 30 40 50 60 70

0.00 0.05 0.10 0.15

2005

 

0 10 20 30 40 50 60 70

0.00 0.05 0.10 0.15

2006

 

0 10 20 30 40 50 60 70

0.00 0.05 0.10 0.15

2007

 

0 10 20 30 40 50 60 70

0.00 0.05 0.10 0.15

2008

 

0 10 20 30 40 50 60 70

0.00 0.05 0.10 0.15

Age (yrs)

Proportion

Figure 217. Age distribution for Pacific ocean perch sampled during the surveys from 2003 to 2008.
231

2003

 

0 5 10 15 20 25 30 35

0.0 0.1 0.2 0.3 0.4

2004

 

0 5 10 15 20 25 30 35

0.0 0.1 0.2 0.3 0.4

2005

 

0 5 10 15 20 25 30 35

0.0 0.1 0.2 0.3 0.4

2006

 

0 5 10 15 20 25 30 35

0.0 0.1 0.2 0.3 0.4

2007

 

0 5 10 15 20 25 30 35

0.0 0.1 0.2 0.3 0.4

2008

 

0 5 10 15 20 25 30 35

0.0 0.1 0.2 0.3 0.4

Age (yrs)

Proportion

Figure 218. Age distribution for petrale sole sampled during the surveys from 2003 to 2008.
232

2003

 

0 10 20 30 40 50 60

0.00 0.15 0.30

2004

 

0 10 20 30 40 50 60

0.00 0.15 0.30

2005

 

0 10 20 30 40 50 60

0.00 0.15 0.30

2006

 

0 10 20 30 40 50 60

0.00 0.15 0.30

Age (yrs)

Proportion

Figure 219. Age distribution for sablefish sampled during the surveys from 2003 to 2006.
233

2003

 

0 20 40 60 80 100

0.00 0.05 0.10 0.15 0.20

2004

 

0 20 40 60 80 100

0.00 0.05 0.10 0.15 0.20

2005

 

0 20 40 60 80 100

0.00 0.05 0.10 0.15 0.20

2006

 

0 20 40 60 80 100

0.00 0.05 0.10 0.15 0.20

2007

 

0 20 40 60 80 100

0.00 0.05 0.10 0.15 0.20

2008

 

0 20 40 60 80 100

0.00 0.05 0.10 0.15 0.20

Age (yrs)

Proportion

Figure 220. Age distribution for splitnose rockfish sampled during the surveys from 2003 to 2008.
234

2003

 

0 5 10 15 20 25 30

0.0 0.1 0.2 0.3 0.4

2004

 

0 5 10 15 20 25 30

0.0 0.1 0.2 0.3 0.4

2005

 

0 5 10 15 20 25 30

0.0 0.1 0.2 0.3 0.4

2006

 

0 5 10 15 20 25 30

0.0 0.1 0.2 0.3 0.4

2007

 

0 5 10 15 20 25 30

0.0 0.1 0.2 0.3 0.4

2008

 

0 5 10 15 20 25 30

0.0 0.1 0.2 0.3 0.4

Age (yrs)

Proportion

Figure 221. Age distribution for widow rockfish sampled during the surveys from 2003 to 2008.
235

2003

 

0 10 20 30

0.00 0.10 0.20

2004

 

0 10 20 30

0.00 0.10 0.20

Age (yrs)

Proportion

Figure 222. Age distribution for yellowtail rockfish sampled during the 2003 and 2004 surveys.
236

0 10 20 30 40 50 60

0.0 0.5 1.0 1.5 2.0

Length (cm)

Weight (kg)

Year

2003

2004

2005

2006

2007

2008

48.0 48.5 49.0 49.5 50.0

1.10 1.20 1.30 1.40

0 10 20 30 40 50 60

0.0 0.5 1.0 1.5 2.0

Length (cm)

Weight (kg)

Year

2003

2004

2005

2006

2007

2008

40 42 44 46 48 50

1.0 1.1 1.2 1.3 1.4 1.5

Figure 223. Weight-length relationship observed for Dover sole (left) and petrale sole (right) during the surveys from 2003 to 2008. Inset shows 

an enlarged view of a subsection of the graph.
237

0 10 20 30 40 50 60

0.0 0.5 1.0 1.5 2.0

Length (cm)

Weight (kg)

Year

2003

2004

2005

2006

2007

2008

50.0 50.5 51.0 51.5 52.0

1.20 1.22 1.24 1.26 1.28 1.30

0 10 20 30 40 50 60

0.0 0.5 1.0 1.5 2.0

Length (cm)

Weight (kg)

Year

2003

2004

2005

2006

2007

2008

45.0 45.5 46.0 46.5 47.0

1.10 1.15 1.20 1.25

Figure 224. Weight-length relationship observed for sablefish (left) and shortspine thornyhead (right) during the surveys from 2003 to 2008. Inset 

shows an enlarged view of a subsection of the graph.
238

 

32

34

36

38

40

42

44

46

48

20 15 10 5 0

Species per haul

 

32

34

36

38

40

42

44

46

48

3 2 1 0

H'

Latitude oN

2003

2004

2005

2006

2007

2008

Mean

Figure 225. Species richness per tow and per year by 

latitude for stratum 1 (55–183 m) in the surveys from 

2003 to 2008.

Figure 226. Species diversity by latitude for stratum 1 (55–183 m) in the 

surveys from 2003 to 2008. The Shannon-Weaver diversity values are 

based on fish species encountered within each depth strata in 1°

intervals of latitude.
239

 

32

34

36

38

40

42

44

46

48

1 0

Species eveness

2003

2004

2005

2006

2007

2008

Mean

 

32

34

36

38

40

42

44

46

48

20 15 10 5 0

Species per haul

Latitude oN

Figure 227. Species eveness by latitude for stratum 1 (55–183 

m) in the surveys from 2003 to 2008. The eveness 

values are based on fish species encountered within each 

depth strata in 1° intervals of latitude.

Figure 228. Species richness per tow and per year by latitude for stratum 2 

(184–549 m) in the surveys from 2003 to 2008.
240

 

32

34

36

38

40

42

44

46

48

3 2 1 0

H'

2003

2004

2005

2006

2007

2008

Mean

 

32

34

36

38

40

42

44

46

48

1 0

Species eveness

Latitude oN

2003

2004

2005

2006

2007

2008

Mean

Figure 229. Species diversity by latitude for stratum 2 (184–549 m) in 

the surveys from 2003 to 2008. The Shannon-Weaver 

diversity values are based on fish species encountered within 

each depth strata in 1° intervals of latitude.

Figure 230. Species eveness by latitude for stratum 2 (184–549 m) in the 

surveys from 2003 to 2008. The eveness values are based on fish 

species encountered within each depth strata in 1° intervals of 

latitude.
241

 

32

34

36

38

40

42

44

46

48

20 15 10 5 0

Species per haul

 

32

34

36

38

40

42

44

46

48

3 2 1 0

H'

Latitude oN

2003

2004

2005

2006

2007

2008

Mean

Figure 231. Species richness per tow and per year by latitude for 

stratum 3 (550–1,280 m) in the surveys from 2003 to 

2008.

Figure 232. Species diversity by latitude for stratum 3 (550–1,280 m) in the 

surveys from 2003 to 2008. The Shannon-Weaver diversity values 

are based on fish species encountered within each depth strata in 1°

intervals of latitude.
242

 

32

34

36

38

40

42

44

46

48

1 0

Species eveness

Latitude oN

2003

2004

2005

2006

2007

2008

Mean

0

0.05

0.1

0.15

0.2

0.25

0.3

0.35

0.4

32 34 36 38 40 42 44 46 48

Latitude oN 

Variance

shallow

mid

deep

Figure 233. Species eveness by latitude for stratum 3 (550–1,280 

m) in the surveys from 2003 to 2008. The eveness values 

are based on fish species encountered within each depth 

strata in 1° intervals of latitude.

Figure 234. Patterns of variance of H′ in relation to latitude for each depth 

stratum for all survey years combined. Variance of H′ and latitude 

are plotted for each 1° latitude bin.
243

Stratum 2 (183 - 549 m)

y = -0.0584x + 2.4738

R2

 = 0.278

0

0.5

1

1.5

2

2.5

3

0 5 10 15 20 25

Richness

H'

Stratum 1 (55 - 182 m)

y = -0.0591x + 2.4242

R2

 = 0.134

0

0.5

1

1.5

2

2.5

3

0 5 10 15 20 25

Richness

H'

Stratum 3 (550 - 1,280 m)

y = -0.0005x + 1.3159

R2

 = 0.00

0

0.5

1

1.5

2

2.5

3

0 5 10 15 20 25

Richness

H'

Figure 235. Relationship between species richness and diversity. Richness and diversity values are 

plotted for each 1° latitude bin in each year of the 2003–2008 surveys.
244

Tables 1–12

Table 1. Coefficients for multiple linear regressions of net width as a function of net height and inverse 

scope for each fishing vessel (FV) by year. Net width = B0 + ax1 + bx2 where x1 is net height 

and x2 is inverse scope. For years with no data, the vessel did not participate in the survey.

Vessel and year B0 a b

FV Excalibur

2003 19.62 –1.10 –324.022

2004 18.69 –0.80 –337.449

2005 22.25 –1.46 –290.290

2006 20.78 –1.17 –353.574

2007 20.38 –1.21 –256.146

2008 17.13 –0.53 –415.877

FV Ms. Julie

2003 17.30 –0.73 –274.059

2004 19.31 –1.11 –270.333

2005 18.01 –0.86 –278.788

2006 17.29 –0.74 –210.001

2007 18.67 –1.08 –83.643

2008 18.63 –0.93 –227.672

FV Noah’s Ark

2003–2004 — — —

2005 20.24 –0.98 –426.917

2006 18.21 –0.62 –475.140

2007 19.58 –0.97 –230.452

2008 19.06 –0.97 –213.228

FV Raven

2003–2004 — — —

2005 18.05 –0.71 –320.346

2006 19.79 –1.14 –109.761

2007 20.56 –1.34 –151.477

2008 17.97 –0.64 –372.588

FV Captain Jack

2003 18.24 –0.78 –335.527

2004–2008 — — —

FV Blue Horizon

2003 19.68 –1.02 –308.055

2004–2008 — — —
245

Table 2. Area sampled and sampling density in each depth stratum for each INPFC statistical area based on successful tows during the surveys 

from 2003 to 2008.

Stratum 1 Stratum 2 Stratum 3 All strata

(55–183 m) (184–549 m) (550–1,280 m) (55–1,280 m)

INFPC area

Area 

(km2)

No. 

hauls

Hauls/ 

1,000 

km2

Area 

(km2)

No. 

hauls

Hauls/ 

1,000 

km2

Area 

(km2)

No. 

hauls

Hauls/ 

1,000 

km2

Area 

(km2)

No. 

hauls

Hauls/

1,000 

km2

2003–2008

U.S.-Vancouver 2,318 164 70.75 2,853 80 28.04 2,286 74 32.37 7,457 318 42.64

49o 00′–47o 30′N

Columbia 14,413 597 41.42 8,621 338 39.21 9,804 272 27.74 32,838 1,207 36.76

47o 30′–43o 00′N

Eureka 4,069 169 41.53 2,034 148 72.76 6,365 179 28.12 12,468 496 39.78

43o 00′–40o 30′N

Monterey 8,605 369 42.88 3,650 150 41.10 8,646 172 19.89 20,901 691 33.06

40o 30′–36o 00′N

Conception 6,994 307 43.89 12,839 414 32.25 42,041 358 8.52 61,874 1,079 17.44

36o 00′–32o 30′N

Total survey area 36,399 1,606 44.12 29,997 1,130 37.67 69,142 1,055 15.26 135,538 3,791 27.97

49o 00′–32o 30′N

2003

U.S.-Vancouver 2,318 48 20.71 2,853 16 5.61 2,286 23 10.06 7,457 87 11.67

49o 00′–47o 30′N

Columbia 14,413 47 3.26 8,621 40 4.64 9,804 61 6.22 32,838 148 4.51

47o 30′–43o 00′N

Eureka 4,069 35 8.60 2,034 36 17.70 6,365 33 5.18 12,468 104 8.34

43o 00′–40o 30′N

Monterey 8,605 45 5.23 3,650 30 8.22 8,646 26 3.01 20,901 101 4.83

40o 30′–36o 00′N

Conception 6,994 52 7.43 12,839 54 4.21 42,041 28 0.67 61,874 134 2.17

36o 00′–32o 30′N

Total survey area 36,399 227 6.24 29,997 176 5.87 69,142 171 2.47 135,538 574 4.23

49o 00′–32o 30′N
246

Table 2 continued. Area sampled and sampling density in each depth stratum for each INPFC statistical area based on successful tows during the 

surveys from 2003 to 2008.

Stratum 1 Stratum 2 Stratum 3 All strata

(55–183 m) (184–549 m) (550–1,280 m) (55–1,280 m)

INFPC area

Area 

(km2)

No. 

hauls

Hauls/ 

1,000 

km2

Area 

(km2)

No. 

hauls

Hauls/ 

1,000 

km2

Area 

(km2)

No. 

hauls

Hauls/ 

1,000 

km2

Area 

(km2)

No. 

hauls

Hauls/

1,000 

km2

2004

U.S.-Vancouver 2,318 29 12.51 2,853 8 2.80 2,286 6 2.62 7,457 43 5.77

49o 00′–47o 30′N

Columbia 14,413 84 5.83 8,621 50 5.80 9,804 28 2.86 32,838 162 4.93

47o 30′–43o 00′N

Eureka 4,069 20 4.92 2,034 12 5.90 6,365 25 3.93 12,468 57 4.57

43o 00′–40o 30′N

Monterey 8,605 59 6.86 3,650 17 4.66 8,646 21 2.43 20,901 97 4.64

40o 30′–36o 00′N

Conception 6,994 47 6.72 12,839 46 3.58 42,041 53 1.26 61,874 146 2.36

36o 00′–32o 30′N

Total survey area 36,399 239 6.57 29,997 133 4.43 69,142 133 1.92 135,538 505 3.73

49o 00′–32o 30′N

2005

U.S.-Vancouver 2,318 21 9.06 2,853 14 4.91 2,286 8 3.50 7,457 43 5.77

49o 00′–47o 30′N

Columbia 14,413 119 8.26 8,621 51 5.92 9,804 53 5.41 32,838 223 6.79

47o 30′–43o 00′N

Eureka 4,069 38 9.34 2,034 23 11.31 6,365 28 4.40 12,468 89 7.14

43o 00′–40o 30′N

Monterey 8,605 78 9.06 3,650 21 5.75 8,646 20 2.31 20,901 119 5.69

40o 30′–36o 00′N

Conception 6,994 54 7.72 12,839 75 5.84 42,041 72 1.71 61,874 201 3.25

36o 00′–32o 30′N

Total survey area 36,399 310 8.52 29,997 184 6.13 69,142 181 2.62 135,538 675 4.98

49o 00′–32o 30′N
247

Table 2 continued. Area sampled and sampling density in each depth stratum for each INPFC statistical area based on successful tows during the 

surveys from 2003 to 2008.

Stratum 1 Stratum 2 Stratum 3 All strata

(55–183 m) (184–549 m) (550–1,280 m) (55–1,280 m)

INFPC area

Area 

(km2)

No. 

hauls

Hauls/ 

1,000 

km2

Area 

(km2)

No. 

hauls

Hauls/ 

1,000 

km2

Area 

(km2)

No. 

hauls

Hauls/ 

1,000 

km2

Area 

(km2)

No. 

hauls

Hauls/

1,000 

km2

2006

U.S.-Vancouver 2,318 24 10.35 2,853 7 2.45 2,286 14 6.12 7,457 45 6.03

49o 00′–47o 30′N

Columbia 14,413 112 7.77 8,621 63 7.31 9,804 42 4.28 32,838 217 6.61

47o 30′–43o 00′N

Eureka 4,069 24 5.90 2,034 20 9.83 6,365 28 4.40 12,468 72 5.77

43o 00′–40o 30′N

Monterey 8,605 61 7.09 3,650 32 8.77 8,646 37 4.28 20,901 130 6.22

40o 30′–36o 00′N

Conception 6,994 48 6.86 12,839 72 5.61 42,041 70 1.67 61,874 190 3.07

36o 00′–32o 30′N

Total survey area 36,399 269 7.39 29,997 194 6.47 69,142 191 2.76 135,538 654 4.83

49o 00′–32o 30′N

2007

U.S.-Vancouver 2,318 25 10.79 2,853 17 5.96 2,286 12 5.25 7,457 54 7.24

49o 00′–47o 30′N

Columbia 14,413 124 8.60 8,621 72 8.35 9,804 45 4.59 32,838 241 7.34

47o 30′–43o 00′N

Eureka 4,069 28 6.88 2,034 28 13.77 6,365 33 5.18 12,468 89 7.14

43o 00′–40o 30′N

Monterey 8,605 56 6.51 3,650 18 4.93 8,646 29 3.35 20,901 103 4.93

40o 30′–36o 00′N

Conception 6,994 55 7.86 12,839 85 6.62 42,041 71 1.69 61,874 211 3.41

36o 00′–32o 30′N

Total survey area 36,399 288 7.91 29,997 220 7.33 69,142 190 2.75 135,538 698 5.15

49o 00′–32o 30′N
248

Table 2 continued. Area sampled and sampling density in each depth stratum for each INPFC statistical area based on successful tows during the 

surveys from 2003 to 2008.

Stratum 1 Stratum 2 Stratum 3 All strata

(55–183 m) (184–549 m) (550–1,280 m) (55–1,280 m)

INFPC area

Area 

(km2)

No. 

hauls

Hauls/ 

1,000 

km2

Area 

(km2)

No. 

hauls

Hauls/ 

1,000 

km2

Area 

(km2)

No. 

hauls

Hauls/ 

1,000 

km2

Area 

(km2)

No. 

hauls

Hauls/

1,000 

km2

2008

U.S.-Vancouver 2,318 17 7.33 2,853 18 6.31 2,286 11 4.81 7,457 46 6.17

49o 00′–47o 30′N

Columbia 14,413 111 7.70 8,621 62 7.19 9,804 43 4.39 32,838 216 6.58

47o 30′–43o 00′N

Eureka 4,069 24 5.90 2,034 29 14.26 6,365 32 5.03 12,468 85 6.82

43o 00′–40o 30′N

Monterey 8,605 70 8.13 3,650 32 8.77 8,646 39 4.51 20,901 141 6.75

40o 30′–36o 00′N

Conception 6,994 51 7.29 12,839 82 6.39 42,041 64 1.52 61,874 197 3.18

36o 00′–32o 30′N

Total survey area 36,399 273 7.50 29,997 223 7.43 69,142 189 2.73 135,538 685 5.05

49o 00′–32o 30′N
249

Table 3. Mean CPUE (kg ha–1

) of 47 important FMP groundfish species plus grooved tanner crab caught 

by depth strata in the Vancouver INPFC area during the 2003–2008 surveys. Stratum 1 typically 

is 54.9–182.9 m, stratum 2 is 182.9–548.6 m, and stratum 3 is 548.6–1,280.2 m; however, for 

arrowtooth flounder, stratum 1 is 54.9–155 m and stratum 2 is 155–548.6 m; and for canary 

rockfish, stratum 1 is 54.9–350 m and stratum 3 is defined as 350–1,280.2 m with no stratum 2. 

These exceptions are noted with an asterisk (*).

Stratum 1

2003 (number of hauls = 48) 2004 (number of hauls = 29) 2005 (number of hauls = 21)

Spiny dogfish 362.95 Spiny dogfish 138.46 Spiny dogfish 126.87

Pacific hake 88.94 Dover sole 24.48 Yellowtail rockfish 35.02

Sablefish 51.32 Arrowtooth flounder* 23.02 Arrowtooth flounder* 27.36

Dover sole 31.32 Canary rockfish* 21.25 Dover sole 19.42

Arrowtooth flounder* 25.29 Yellowtail rockfish 20.97 Greenstriped rockfish 10.06

Spotted ratfish 13.98 English sole 12.67 Longnose skate 8.83

Yellowtail rockfish 13.98 Greenstriped rockfish 5.58 English sole 6.30

Pacific sanddab 10.11 Sablefish 5.55 Spotted ratfish 6.29

Redstripe rockfish 10.08 Pacific cod 5.34 Rex sole 5.94

Greenstriped rockfish 9.51 Pacific sanddab 5.15 Lingcod 3.73

English sole 9.11 Longnose skate 4.65 Petrale sole 3.03

Lingcod 9.01 Petrale sole 4.27 Pacific hake 2.94

Pacific cod 8.49 Spotted ratfish 3.93 Pacific cod 2.84

Rex sole 8.20 Rex sole 3.68 Pacific sanddab 2.41

Longnose skate 7.59 Pacific hake 3.57 Sablefish 2.02

Petrale sole 5.88 Big skate 3.23 Canary rockfish* 1.46

Widow rockfish 3.95 Lingcod 3.06 Flathead sole 1.14

Canary rockfish* 3.71 Rosethorn rockfish 1.14 California skate 0.44

Flathead sole 2.62 Flathead sole 1.03 Rosethorn rockfish 0.44

Pygmy rockfish 1.73 Darkblotched rockfish 0.95 Big skate 0.33

Rosethorn rockfish 1.56 Redstripe rockfish 0.92 Darkblotched rockfish 0.24

Big skate 1.51 Sharpchin rockfish 0.57 Sharpchin rockfish 0.18

Sharpchin rockfish 0.64 Pacific ocean perch 0.44 Redstripe rockfish 0.11

Greenspotted rockfish 0.49 Redbanded rockfish 0.22 Rougheye rockfish 0.03

Darkblotched rockfish 0.29 Shortspine thornyhead 0.14 Pacific ocean perch 0.01

Bocaccio 0.24 Splitnose rockfish 0.10 Redbanded rockfish 0.01

Shortspine thornyhead 0.23 Greenspotted rockfish 0.09 Pygmy rockfish 0.01

Pacific ocean perch 0.08 Widow rockfish 0.08 Splitnose rockfish <0.01

Curlfin sole 0.07 Pygmy rockfish 0.03 Stripetail rockfish <0.01

Splitnose rockfish 0.02 Rougheye rockfish 0.01

Rougheye rockfish 0.01 Stripetail rockfish <0.01

Redbanded rockfish 0.01

Stripetail rockfish 0.01

Grooved tanner crab <0.01

Shortbelly rockfish <0.01
250

Table 3 continued. Mean CPUE (kg ha–1

) of 47 important FMP groundfish species plus grooved tanner 

crab caught by depth strata in the Vancouver INPFC area during the 2003–2008 surveys. Stratum 

1 typically is 54.9–182.9 m, stratum 2 is 182.9–548.6 m, and stratum 3 is 548.6–1,280.2 m; 

however, for arrowtooth flounder, stratum 1 is 54.9–155 m and stratum 2 is 155–548.6 m; and for 

canary rockfish, stratum 1 is 54.9–350 m and stratum 3 is defined as 350–1,280.2 m with no 

stratum 2. These exceptions are noted with an asterisk (*).

Stratum 1

2006 (number of hauls = 24) 2007 (number of hauls = 25) 2008 (number of hauls = 17)

Canary rockfish* 171.14 Yellowtail rockfish 51.37 Yellowtail rockfish 30.36

Yellowtail rockfish 26.94 Spiny dogfish 38.77 Spiny dogfish 15.13

Redstripe rockfish 25.73 Dover sole 16.52 Arrowtooth flounder* 15.08

Spotted ratfish 15.87 Arrowtooth flounder* 15.30 Spotted ratfish 11.97

Dover sole 12.70 Greenstriped rockfish 12.02 Greenstriped rockfish 9.60

Spiny dogfish 9.80 Longnose skate 11.05 Dover sole 6.35

Lingcod 8.11 Lingcod 8.06 Longnose skate 5.80

Longnose skate 6.23 Canary rockfish* 5.35 Lingcod 5.37

Arrowtooth flounder* 6.15 Spotted ratfish 4.49 English sole 5.37

Greenstriped rockfish 5.99 Rex sole 3.31 Pacific sanddab 4.88

Rex sole 5.51 Petrale sole 2.61 Redstripe rockfish 4.14

Petrale sole 4.03 English sole 2.27 Canary rockfish* 3.53

Pacific cod 4.00 Splitnose rockfish 1.92 Rosethorn rockfish 2.66

Bocaccio 3.26 Pacific hake 1.83 Petrale sole 2.40

English sole 2.85 Sharpchin rockfish 1.80 Rex sole 2.18

Flathead sole 1.91 Redstripe rockfish 1.72 Sharpchin rockfish 2.18

Pacific sanddab 1.83 Pacific cod 1.20 Pacific cod 1.33

Widow rockfish 1.34 Darkblotched rockfish 1.16 Bocaccio 0.71

Big skate 1.30 Rosethorn rockfish 0.95 Sablefish 0.35

Pacific hake 1.22 Bocaccio 0.71 Splitnose rockfish 0.32

Darkblotched rockfish 1.12 Pacific sanddab 0.70 Pygmy rockfish 0.16

Sharpchin rockfish 1.01 Sablefish 0.59 Rougheye rockfish 0.13

Sablefish 0.55 Pacific ocean perch 0.32 Flathead sole 0.05

Shortbelly rockfish 0.30 Big skate 0.20 Redbanded rockfish 0.03

Rosethorn rockfish 0.22 Flathead sole 0.17 Greenspotted rockfish 0.03

Shortspine thornyhead 0.11 Stripetail rockfish 0.12 Darkblotched rockfish 0.03

Pacific ocean perch 0.10 Greenspotted rockfish 0.04 Bank rockfish <0.01

Rougheye rockfish 0.05 Widow rockfish 0.03 Stripetail rockfish <0.01

Splitnose rockfish 0.03 Curlfin sole 0.02 Shortspine thornyhead <0.01

Greenspotted rockfish 0.03 Pygmy rockfish 0.02

Pygmy rockfish 0.02 Shortspine thornyhead 0.01

Stripetail rockfish 0.01 Redbanded rockfish 0.01

Redbanded rockfish <0.01 Longspine thornyhead <0.01
251

Table 3 continued. Mean CPUE (kg ha–1

) of 47 important FMP groundfish species plus grooved tanner 

crab caught by depth strata in the Vancouver INPFC area during the 2003–2008 surveys. Stratum 

1 typically is 54.9–182.9 m, stratum 2 is 182.9–548.6 m, and stratum 3 is 548.6–1,280.2 m; 

however, for arrowtooth flounder, stratum 1 is 54.9–155 m and stratum 2 is 155–548.6 m. These 

exceptions are noted with an asterisk (*).

Stratum 2

2003 (number of hauls = 16) 2004 (number of hauls = 8) 2005 (number of hauls = 14)

Spiny dogfish 133.75 Dover sole 57.47 Dover sole 64.93

Dover sole 63.98 Sablefish 29.56 Arrowtooth flounder* 58.43

Pacific ocean perch 39.52 Pacific ocean perch 29.13 Sablefish 21.89

Arrowtooth flounder* 38.17 Arrowtooth flounder* 26.9 Longnose skate 15.77

Longnose skate 13.11 Sharpchin rockfish 24.03 Pacific ocean perch 14.46

Shortspine thornyhead 11.57 Spiny dogfish 19.37 Spiny dogfish 10.87

Spotted ratfish 8.89 Spotted ratfish 11.47 Splitnose rockfish 10.43

Sablefish 8.19 Longnose skate 9.19 Spotted ratfish 8.47

Rex sole 7.83 Shortspine thornyhead 8.59 Rex sole 6.77

Splitnose rockfish 6.51 Redbanded rockfish 7.05 Shortspine thornyhead 4.56

Flathead sole 6.45 Rex sole 5.80 Flathead sole 3.80

Pacific hake 5.16 Rosethorn rockfish 2.60 Greenstriped rockfish 3.71

Sharpchin rockfish 4.88 Pacific hake 2.08 Yellowtail rockfish 3.38

Greenstriped rockfish 4.64 Rougheye rockfish 1.43 Pacific cod 1.87

Darkblotched rockfish 4.60 English sole 1.36 Rougheye rockfish 1.38

Pacific cod 3.34 Lingcod 1.09 Redstripe rockfish 1.24

Lingcod 2.02 Flathead sole 0.76 Pacific hake 1.19

Redbanded rockfish 1.49 Pacific cod 0.54 English sole 0.79

Rosethorn rockfish 1.24 Longspine thornyhead 0.30 Darkblotched rockfish 0.69

Yellowtail rockfish 1.17 Petrale sole 0.16 Redbanded rockfish 0.54

Petrale sole 1.11 Greenstriped rockfish 0.15 Lingcod 0.37

Rougheye rockfish 0.81 Giant grenadier 0.09 Sharpchin rockfish 0.26

English sole 0.35 Widow rockfish 0.08 Aurora rockfish 0.12

Redstripe rockfish 0.28 Pacific flatnose 0.07 Petrale sole 0.07

Grooved tanner crab 0.17 Grooved tanner crab 0.02 Rosethorn rockfish 0.07

Longspine thornyhead 0.11 Pacific grenadier 0.01 Widow rockfish 0.03

Widow rockfish 0.08 Splitnose rockfish <0.01 Grooved tanner crab 0.01

Aurora rockfish 0.06 Stripetail rockfish 0.01

Pacific flatnose 0.03

Stripetail rockfish 0.01

Pacific grenadier <0.01
252

Table 3 continued. Mean CPUE (kg ha–1

) of 47 important FMP groundfish species plus grooved tanner 

crab caught by depth strata in the Vancouver INPFC area during the 2003–2008 surveys. Stratum 

1 typically is 54.9–182.9 m, stratum 2 is 182.9–548.6 m, and stratum 3 is 548.6–1,280.2 m; 

however, for arrowtooth flounder, stratum 1 is 54.9–155 m and stratum 2 is 155–548.6 m. These 

exceptions are noted with an asterisk (*).

Stratum 2

2006 (number of hauls = 7) 2007 (number of hauls = 17) 2008 (number of hauls = 18)

Pacific ocean perch 88.58 Dover sole 55.69 Dover sole 96.86

Dover sole 50.74 Arrowtooth flounder* 32.54 Arrowtooth flounder* 34.79

Arrowtooth flounder* 34.74 Splitnose rockfish 18.02 Longnose skate 19.90

Longnose skate 20.47 Greenstriped rockfish 17.83 Rex sole 10.03

Darkblotched rockfish 12.90 Pacific ocean perch 15.25 Spiny dogfish 9.51

Sablefish 11.94 Longnose skate 10.64 Shortspine thornyhead 9.20

Sharpchin rockfish 8.96 Spotted ratfish 9.94 Sablefish 7.19

Splitnose rockfish 8.56 Sablefish 7.59 Greenstriped rockfish 4.97

Spotted ratfish 7.92 Rex sole 6.97 Pacific ocean perch 4.57

Shortspine thornyhead 7.83 Shortspine thornyhead 6.92 Pacific hake 4.54

Rex sole 5.45 Spiny dogfish 5.69 Spotted ratfish 4.34

Lingcod 2.30 Pacific hake 5.41 Splitnose rockfish 3.85

Spiny dogfish 1.84 Darkblotched rockfish 5.18 Lingcod 3.40

Grooved tanner crab 1.04 Sharpchin rockfish 3.80 Rosethorn rockfish 2.90

English sole 0.92 Redbanded rockfish 3.36 Rougheye rockfish 1.75

Rougheye rockfish 0.85 Lingcod 2.21 Flathead sole 1.42

Pacific hake 0.79 Yellowtail rockfish 1.91 Sharpchin rockfish 1.06

Pacific cod 0.68 Petrale sole 1.09 Redstripe rockfish 0.62

Rosethorn rockfish 0.57 Rosethorn rockfish 1.06 Redbanded rockfish 0.51

Longspine thornyhead 0.48 Pacific cod 0.89 Darkblotched rockfish 0.31

Bocaccio 0.42 Rougheye rockfish 0.48 Bocaccio 0.31

Greenstriped rockfish 0.41 Grooved tanner crab 0.45 Petrale sole 0.22

Yellowtail rockfish 0.31 Flathead sole 0.40 Aurora rockfish 0.12

Petrale sole 0.18 English sole 0.27 Widow rockfish 0.06

Redbanded rockfish 0.18 Redstripe rockfish 0.21 Blackgill rockfish 0.05

Aurora rockfish 0.16 Longspine thornyhead 0.08 English sole 0.05

Bank rockfish 0.12 Stripetail rockfish 0.04 Longspine thornyhead 0.02

Pacific grenadier 0.11 Aurora rockfish 0.02 Pacific sanddab 0.01

Flathead sole 0.08 Pacific flatnose 0.01 Stripetail rockfish <0.01

Stripetail rockfish 0.02 Pacific grenadier <0.01 Grooved tanner crab <0.01
253

Table 3 continued. Mean CPUE (kg ha–1

) of 47 important FMP groundfish species plus grooved tanner 

crab caught by depth strata in the Vancouver INPFC area during the 2003–2008 surveys. Stratum 

1 typically is 54.9–182.9 m, stratum 2 is 182.9–548.6 m, and stratum 3 is 548.6–1,280.2 m; 

however, for arrowtooth flounder, stratum 1 is 54.9–155 m and stratum 2 is 155–548.6 m. These 

exceptions are noted with an asterisk (*).

Stratum 3

2003 (number of hauls = 23) 2004 (number of hauls = 6) 2005 (number of hauls = 8)

Longspine thornyhead 26.91 Dover sole 24.91 Longspine thornyhead 29.88

Grooved tanner crab 9.72 Sablefish 21.2 Grooved tanner crab 11.71

Giant grenadier 7.75 Longspine thornyhead 21.17 Giant grenadier 9.58

Pacific grenadier 4.99 Grooved tanner crab 13.55 Sablefish 5.12

Sablefish 4.56 Pacific grenadier 11.88 Pacific grenadier 4.79

Shortspine thornyhead 2.81 Shortspine thornyhead 9.80 Shortspine thornyhead 1.71

Dover sole 1.92 Giant grenadier 5.99 Pacific flatnose 0.55

Pacific flatnose 0.64 Pacific flatnose 0.62 Dover sole 0.23

Pacific hake 0.15 Rex sole 0.58 Longnose skate 0.15

Longnose skate 0.12 Pacific hake 0.21 Pacific ocean perch 0.12

Longnose skate 0.17

Rougheye rockfish 0.09

Arrowtooth flounder* 0.05

2006 (number of hauls = 14) 2007 (number of hauls = 12) 2008 (number of hauls = 11)

Longspine thornyhead 27.86 Longspine thornyhead 24.69 Longspine thornyhead 32.95

Sablefish 11.34 Grooved tanner crab 13.50 Sablefish 8.52

Giant grenadier 10.69 Dover sole 11.50 Grooved tanner crab 7.72

Grooved tanner crab 7.87 Sablefish 11.32 Giant grenadier 6.67

Dover sole 6.04 Shortspine thornyhead 6.14 Pacific grenadier 5.13

Pacific grenadier 5.40 Pacific grenadier 5.47 Dover sole 4.78

Shortspine thornyhead 2.47 Giant grenadier 5.13 Shortspine thornyhead 3.34

Longnose skate 1.35 Arrowtooth flounder* 0.67 Pacific flatnose 0.59

Pacific flatnose 0.55 Pacific flatnose 0.29 Spiny dogfish 0.09

Arrowtooth flounder* 0.34 Longnose skate 0.25

Spiny dogfish 0.08 Pacific hake 0.19

Rex sole 0.05 Spiny dogfish 0.08

Redbanded rockfish 0.08

Aurora rockfish 0.06
254

Table 4. Mean CPUE (kg ha–1

) of 47 important FMP groundfish species plus grooved tanner crab caught 

by depth strata in the Columbia INPFC area during the 2003–2008 surveys. Stratum 1 typically 

is 54.9–182.9 m, stratum 2 is 182.9–548.6 m, and stratum 3 is 548.6–1,280.2 m; however, for 

arrowtooth flounder, stratum 1 is 54.9–155 m and stratum 2 is 155–548.6 m; and for canary 

rockfish, stratum 1 is 54.9–350 m and stratum 3 is defined as 350–1,280.2 m with no stratum 2. 

These exceptions are noted with an asterisk (*).

Stratum 1

2003 (number of hauls = 47) 2004 (number of hauls = 84) 2005 (number of hauls = 119)

Pacific hake 87.79 Pacific hake 65.83 Pacific hake 27.73

Pacific sanddab 30.31 Dover sole 28.44 Dover sole 25.01

Dover sole 28.37 Pacific sanddab 19.44 Pacific sanddab 12.58

Sablefish 17.26 English sole 11.14 Rex sole 10.05

Yellowtail rockfish 14.37 Rex sole 11.07 English sole 6.15

Rex sole 9.30 Longnose skate 8.29 Yellowtail rockfish 6.07

Longnose skate 9.27 Sablefish 5.91 Arrowtooth flounder* 5.82

English sole 7.36 Lingcod 4.57 Longnose skate 5.28

Arrowtooth flounder* 5.61 Arrowtooth flounder* 4.40 Petrale sole 4.84

Widow rockfish 5.55 Yellowtail rockfish 3.96 Greenstriped rockfish 4.36

Spiny dogfish 5.27 Petrale sole 3.72 Sharpchin rockfish 4.34

Lingcod 5.08 Spotted ratfish 3.47 Big skate 3.62

Spotted ratfish 4.20 Greenstriped rockfish 3.05 Lingcod 2.62

Petrale sole 3.24 Spiny dogfish 2.93 Spotted ratfish 2.61

Sharpchin rockfish 2.76 Big skate 2.59 Spiny dogfish 2.36

Big skate 2.57 Pacific cod 2.34 Sablefish 2.01

Redstripe rockfish 1.49 Redstripe rockfish 2.33 Darkblotched rockfish 1.23

Greenstriped rockfish 1.39 Sharpchin rockfish 2.29 Canary rockfish* 0.57

Flathead sole 1.31 Darkblotched rockfish 2.09 Pacific ocean perch 0.54

Pacific cod 1.23 Canary rockfish* 1.26 Splitnose rockfish 0.51

Canary rockfish* 0.91 Flathead sole 0.83 Flathead sole 0.42

Splitnose rockfish 0.83 Pygmy rockfish 0.55 Rosethorn rockfish 0.23

Rosethorn rockfish 0.68 Rosethorn rockfish 0.39 Pacific cod 0.20

Bocaccio 0.30 Greenspotted rockfish 0.22 Widow rockfish 0.20

Darkblotched rockfish 0.27 Curlfin sole 0.15 Greenspotted rockfish 0.15

Shortspine thornyhead 0.22 Splitnose rockfish 0.06 Shortspine thornyhead 0.11

Curlfin sole 0.18 Stripetail rockfish 0.05 Stripetail rockfish 0.08

Redbanded rockfish 0.08 Shortspine thornyhead 0.05 Curlfin sole 0.07

Pygmy rockfish 0.05 Rougheye rockfish 0.02 Redstripe rockfish 0.07

Stripetail rockfish 0.04 Redbanded rockfish 0.01 Rougheye rockfish 0.05

Rougheye rockfish 0.04 Halfbanded rockfish 0.01 Bocaccio 0.02

Grooved tanner crab 0.04 Pacific ocean perch <0.01 Redbanded rockfish 0.01

Greenspotted rockfish 0.01 Pygmy rockfish 0.01

Halfbanded rockfish 0.01 Grooved tanner crab <0.01

Pacific ocean perch <0.01 California skate <0.01

Shortbelly rockfish <0.01 Halfbanded rockfish <0.01
255

Table 4 continued. Mean CPUE (kg ha–1

) of 47 important FMP groundfish species plus grooved tanner 

crab caught by depth strata in the Columbia INPFC area during the 2003–2008 surveys. Stratum 

1 typically is 54.9–182.9 m, stratum 2 is 182.9–548.6 m, and stratum 3 is 548.6–1,280.2 m; 

however, for arrowtooth flounder, stratum 1 is 54.9–155 m and stratum 2 is 155–548.6 m; and for 

canary rockfish, stratum 1 is 54.9–350 m and stratum 3 is defined as 350–1,280.2 m with no 

stratum 2. These exceptions are noted with an asterisk (*).

Stratum 1

2006 (number of hauls = 112) 2007 (number of hauls = 124) 2008 (number of hauls = 111)

Pacific hake 45.76 Dover sole 36.52 Dover sole 25.17

Dover sole 26.78 Pacific hake 29.45 Pacific hake 23.19

Pacific sanddab 15.32 Pacific sanddab 9.24 Pacific sanddab 15.38

Rex sole 9.16 Rex sole 9.10 Arrowtooth flounder* 8.03

Longnose skate 6.27 Longnose skate 8.61 Rex sole 6.45

English sole 5.57 Yellowtail rockfish 6.06 Longnose skate 6.22

Greenstriped rockfish 4.29 Arrowtooth flounder* 5.92 Greenstriped rockfish 5.76

Lingcod 4.25 English sole 5.20 English sole 3.96

Petrale sole 4.23 Greenstriped rockfish 4.21 Canary rockfish* 3.89

Arrowtooth flounder* 3.30 Petrale sole 3.37 Spiny dogfish 3.25

Redstripe rockfish 3.11 Lingcod 3.10 Petrale sole 3.13

Spotted ratfish 2.60 Big skate 2.61 Spotted ratfish 2.89

Big skate 2.34 Spotted ratfish 2.12 Lingcod 2.87

Sablefish 1.90 Spiny dogfish 1.90 Darkblotched rockfish 2.49

Spiny dogfish 1.46 Sablefish 1.74 Yellowtail rockfish 2.37

Pygmy rockfish 1.15 Canary rockfish* 1.05 Big skate 1.31

Sharpchin rockfish 0.85 Redstripe rockfish 0.91 Sablefish 0.72

Rosethorn rockfish 0.74 Splitnose rockfish 0.91 Sharpchin rockfish 0.60

Darkblotched rockfish 0.58 Rosethorn rockfish 0.87 Splitnose rockfish 0.44

Yellowtail rockfish 0.36 Darkblotched rockfish 0.32 Rosethorn rockfish 0.35

Flathead sole 0.29 Stripetail rockfish 0.32 Redstripe rockfish 0.17

Canary rockfish* 0.21 Shortspine thornyhead 0.18 Stripetail rockfish 0.14

Stripetail rockfish 0.18 Sharpchin rockfish 0.14 Pacific cod 0.13

Greenspotted rockfish 0.16 Greenspotted rockfish 0.14 Pacific ocean perch 0.12

Pacific cod 0.13 Widow rockfish 0.14 Curlfin sole 0.10

Curlfin sole 0.13 Flathead sole 0.11 Greenspotted rockfish 0.07

Splitnose rockfish 0.08 Curlfin sole 0.08 Shortspine thornyhead 0.07

Shortspine thornyhead 0.07 Pygmy rockfish 0.04 Flathead sole 0.04

Rougheye rockfish 0.06 Pacific ocean perch 0.04 Rougheye rockfish 0.02

Pacific ocean perch 0.02 Pacific cod 0.03 Widow rockfish 0.02

Widow rockfish 0.01 Rougheye rockfish 0.03 Redbanded rockfish 0.01

Redbanded rockfish <0.01 Redbanded rockfish 0.01 Pygmy rockfish <0.01

California skate <0.01 Halfbanded rockfish <0.01 Bocaccio <0.01

Shortbelly rockfish <0.01
256

Table 4 continued. Mean CPUE (kg ha–1

) of 47 important FMP groundfish species plus grooved tanner 

crab caught by depth strata in the Columbia INPFC area during the 2003–2008 surveys. Stratum 

1 typically is 54.9–182.9 m, stratum 2 is 182.9–548.6 m, and stratum 3 is 548.6–1,280.2 m; 

however, for arrowtooth flounder, stratum 1 is 54.9–155 m and stratum 2 is 155–548.6 m. These 

exceptions are noted with an asterisk (*).

Stratum 2

2003 (number of hauls = 40) 2004 (number of hauls = 50) 2005 (number of hauls = 51)

Dover sole 38.10 Dover sole 30.15 Dover sole 34.23

Pacific hake 32.97 Sharpchin rockfish 16.61 Darkblotched rockfish 17.76

Sharpchin rockfish 27.99 Sablefish 13.37 Sablefish 14.04

Darkblotched rockfish 27.13 Pacific hake 13.04 Shortspine thornyhead 9.76

Pacific ocean perch 25.24 Arrowtooth flounder* 12.84 Arrowtooth flounder* 9.63

Sablefish 24.41 Rex sole 11.64 Longnose skate 9.57

Arrowtooth flounder* 17.32 Shortspine thornyhead 10.90 Pacific ocean perch 9.40

Rex sole 12.40 Splitnose rockfish 8.95 Pacific hake 8.26

Spiny dogfish 9.50 Longnose skate 7.54 Rex sole 7.45

Shortspine thornyhead 9.17 Pacific ocean perch 2.48 Splitnose rockfish 7.29

Longnose skate 8.02 Stripetail rockfish 2.45 Sharpchin rockfish 4.88

Splitnose rockfish 6.39 Rougheye rockfish 2.43 Greenstriped rockfish 3.67

Redstripe rockfish 4.05 Aurora rockfish 1.42 Rougheye rockfish 3.48

Rosethorn rockfish 2.91 English sole 1.33 Longspine thornyhead 1.84

Lingcod 2.12 Longspine thornyhead 1.29 Spotted ratfish 1.24

Spotted ratfish 1.43 Spotted ratfish 1.25 Rosethorn rockfish 0.72

Greenstriped rockfish 0.94 Darkblotched rockfish 1.24 Spiny dogfish 0.63

Longspine thornyhead 0.87 Spiny dogfish 1.11 Lingcod 0.61

Redbanded rockfish 0.54 Rosethorn rockfish 1.05 Aurora rockfish 0.52

Pacific cod 0.50 Greenstriped rockfish 0.89 Grooved tanner crab 0.47

Petrale sole 0.47 Lingcod 0.87 Petrale sole 0.30

Stripetail rockfish 0.45 Blackgill rockfish 0.82 English sole 0.28

Rougheye rockfish 0.39 Big skate 0.39 Redbanded rockfish 0.24

Grooved tanner crab 0.38 Redbanded rockfish 0.24 Big skate 0.22

English sole 0.36 Pacific cod 0.23 Blackgill rockfish 0.20

Yellowtail rockfish 0.28 Grooved tanner crab 0.18 Yellowtail rockfish 0.09

Big skate 0.27 Petrale sole 0.13 Flathead sole 0.09

Aurora rockfish 0.26 Redstripe rockfish 0.05 Stripetail rockfish 0.04

Blackgill rockfish 0.15 Pacific flatnose 0.04 Widow rockfish 0.04

Flathead sole 0.09 Widow rockfish 0.04 Pacific grenadier 0.04

Pacific flatnose 0.07 Flathead sole 0.02 Pacific flatnose 0.03

Widow rockfish 0.06 Giant grenadier 0.02 Pacific cod 0.02

Bank rockfish 0.03 Shortbelly rockfish 0.01 Redstripe rockfish 0.01

Greenspotted rockfish 0.03 Pacific grenadier <0.01 Bank rockfish <0.01

Giant grenadier 0.02 Bank rockfish <0.01 Bocaccio <0.01

Pacific grenadier 0.01 Bocaccio <0.01 Calico rockfish <0.01

Shortbelly rockfish 0.01 Calico rockfish <0.01 Calif. scorpionfish <0.01
257

Table 4 continued. Mean CPUE (kg ha–1

) of 47 important FMP groundfish species plus grooved tanner 

crab caught by depth strata in the Columbia INPFC area during the 2003–2008 surveys. Stratum 

1 typically is 54.9–182.9 m, stratum 2 is 182.9–548.6 m, and stratum 3 is 548.6–1,280.2 m; 

however, for arrowtooth flounder, stratum 1 is 54.9–155 m and stratum 2 is 155–548.6 m. These 

exceptions are noted with an asterisk (*).

Stratum 2

2006 (number of hauls = 63) 2007 (number of hauls = 72) 2008 (number of hauls = 62)

Dover sole 50.26 Dover sole 46.14 Dover sole 44.28

Arrowtooth flounder* 26.51 Sablefish 15.96 Pacific hake 15.14

Sablefish 15.18 Shortspine thornyhead 12.20 Splitnose rockfish 10.57

Rex sole 11.13 Rex sole 11.91 Shortspine thornyhead 10.14

Longnose skate 9.50 Arrowtooth flounder* 11.5 Arrowtooth flounder* 10.02

Pacific hake 8.92 Splitnose rockfish 10.35 Longnose skate 9.57

Shortspine thornyhead 8.82 Longnose skate 9.35 Sablefish 8.69

Splitnose rockfish 8.41 Pacific hake 5.25 Rex sole 8.11

Sharpchin rockfish 3.35 Darkblotched rockfish 2.53 Pacific ocean perch 6.21

Darkblotched rockfish 2.96 English sole 1.72 Lingcod 5.13

English sole 2.50 Spotted ratfish 1.60 Darkblotched rockfish 3.92

Pacific ocean perch 2.16 Stripetail rockfish 1.34 Spiny dogfish 2.27

Lingcod 1.71 Longspine thornyhead 1.28 Sharpchin rockfish 2.16

Longspine thornyhead 1.31 Sharpchin rockfish 1.18 Yellowtail rockfish 1.91

Spotted ratfish 1.16 Spiny dogfish 0.97 Stripetail rockfish 1.86

Stripetail rockfish 0.85 Pacific ocean perch 0.91 Longspine thornyhead 1.54

Greenstriped rockfish 0.82 Lingcod 0.87 Spotted ratfish 1.27

Aurora rockfish 0.74 Rougheye rockfish 0.46 English sole 1.00

Spiny dogfish 0.72 Aurora rockfish 0.45 Greenstriped rockfish 0.95

Rougheye rockfish 0.55 Grooved tanner crab 0.43 Rosethorn rockfish 0.53

Grooved tanner crab 0.48 Rosethorn rockfish 0.36 Rougheye rockfish 0.52

Rosethorn rockfish 0.47 Greenstriped rockfish 0.31 Aurora rockfish 0.35

Petrale sole 0.23 Redbanded rockfish 0.20 Grooved tanner crab 0.35

Big skate 0.21 Widow rockfish 0.18 Redbanded rockfish 0.33

Redbanded rockfish 0.21 Petrale sole 0.10 Blackgill rockfish 0.24

Flathead sole 0.14 Bocaccio 0.04 Petrale sole 0.19

Widow rockfish 0.11 Blackgill rockfish 0.03 Widow rockfish 0.08

Yellowtail rockfish 0.02 Yellowtail rockfish 0.02 Bocaccio 0.04

Pacific flatnose 0.02 Pacific flatnose 0.01 Pacific cod 0.04

Redstripe rockfish 0.01 Bank rockfish 0.01 Giant grenadier 0.02

Blackgill rockfish <0.01 Redstripe rockfish <0.01 Flathead sole 0.02

Greenspotted rockfish <0.01 Greenspotted rockfish <0.01 Greenspotted rockfish 0.01

Shortbelly rockfish <0.01 Pygmy rockfish <0.01 Pacific grenadier <0.01

Pacific grenadier <0.01 Big skate <0.01 Bank rockfish <0.01

Bank rockfish <0.01 Calico rockfish <0.01 Pacific flatnose <0.01

Bocaccio <0.01 California scorpionfish <0.01

Calico rockfish <0.01 California skate <0.01
258

Table 4 continued. Mean CPUE (kg ha–1

) of 47 important FMP groundfish species plus grooved tanner 

crab caught by depth strata in the Columbia INPFC area during the 2003–2008 surveys. Stratum 

1 typically is 54.9–182.9 m, stratum 2 is 182.9–548.6 m, and stratum 3 is 548.6–1,280.2 m; 

however, for arrowtooth flounder, stratum 1 is 54.9–155 m and stratum 2 is 155–548.6 m. These 

exceptions are noted with an asterisk (*).

Stratum 3

2003 (number of hauls = 61) 2004 (number of hauls = 28) 2005 (number of hauls = 53)

Longspine thornyhead 26.11 Longspine thornyhead 22.71 Longspine thornyhead 21.55

Dover sole 10.70 Sablefish 9.55 Grooved tanner crab 19.59

Sablefish 10.44 Pacific grenadier 8.86 Giant grenadier 14.12

Grooved tanner crab 9.61 Grooved tanner crab 6.82 Sablefish 12.81

Pacific grenadier 8.84 Giant grenadier 6.73 Dover sole 9.82

Giant grenadier 6.13 Dover sole 5.95 Pacific grenadier 9.59

Shortspine thornyhead 3.55 Shortspine thornyhead 3.50 Shortspine thornyhead 3.94

Pacific flatnose 0.83 Pacific flatnose 0.54 Pacific flatnose 0.90

Pacific hake 0.57 Pacific hake 0.22 Pacific hake 0.22

Rex sole 0.32 Longnose skate 0.16 Longnose skate 0.18

Longnose skate 0.29 Rex sole 0.07 Arrowtooth flounder* 0.14

Arrowtooth flounder* 0.06 Aurora rockfish 0.01 Rex sole 0.12

Rougheye rockfish 0.02 Spotted ratfish 0.01 Aurora rockfish 0.02

Aurora rockfish 0.02 Pacific ocean perch 0.01

2006 (number of hauls = 42) 2007 (number of hauls = 45) 2008 (number of hauls = 43)

Longspine thornyhead 23.49 Longspine thornyhead 28.58 Longspine thornyhead 32.15

Pacific grenadier 17.46 Sablefish 11.75 Sablefish 14.59

Sablefish 12.39 Giant grenadier 10.55 Pacific grenadier 12.39

Grooved tanner crab 7.90 Grooved tanner crab 10.54 Grooved tanner crab 9.77

Giant grenadier 6.32 Pacific grenadier 10.16 Dover sole 8.65

Dover sole 6.18 Dover sole 9.83 Giant grenadier 8.00

Shortspine thornyhead 3.52 Shortspine thornyhead 4.56 Shortspine thornyhead 3.87

Pacific flatnose 0.79 Pacific flatnose 1.29 Pacific flatnose 0.89

Longnose skate 0.38 Longnose skate 0.09 Longnose skate 0.67

Pacific hake 0.10 Pacific hake 0.07 Rex sole 0.23

Rex sole 0.04 Rex sole 0.04 Pacific hake 0.22

Spiny dogfish 0.01 Rougheye rockfish 0.02 Arrowtooth flounder* 0.08

Aurora rockfish 0.01 Aurora rockfish 0.02
259

Table 5. Mean CPUE (kg ha–1

) of 47 important FMP groundfish species plus grooved tanner crab caught 

by depth strata in the Eureka INPFC area during the 2003–2008 surveys. Stratum 1 typically is 

54.9–182.9 m, stratum 2 is 182.9–548.6 m, and stratum 3 is 548.6–1,280.2 m; however, for 

arrowtooth flounder and chilipepper rockfish, stratum 1 is 54.9–155 m and stratum 2 is 155–

548.6 m; and for canary rockfish, stratum 1 is 54.9–350 m and stratum 3 is defined as 350–

1,280.2 m with no stratum 2. These exceptions are noted with an asterisk (*).

Stratum 1

2003 (number of hauls = 35) 2004 (number of hauls = 20) 2005 (number of hauls = 38)

Sharpchin rockfish 215.89 Pacific hake 43.71 Pacific hake 30.38

Pacific hake 96.75 English sole 29.27 Dover sole 19.61

Redstripe rockfish 52.8 Dover sole 25.98 English sole 16.36

Dover sole 34.04 Rex sole 20.68 Stripetail rockfish 15.18

Lingcod 32.56 Chilipepper rockfish* 14.15 Rex sole 12.24

Pacific sanddab 20.17 Petrale sole 11.21 Pacific sanddab 11.2

English sole 14.62 Longnose skate 9.77 Canary rockfish* 10.57

Rex sole 13.33 Pacific sanddab 8.58 Lingcod 9.93

Sablefish 8.54 Big skate 7.98 Petrale sole 7.98

Longnose skate 7.22 Stripetail rockfish 6.42 Longnose skate 6.49

Greenstriped rockfish 6.29 Lingcod 5.53 Spiny dogfish 6.05

Spotted ratfish 5.24 Sablefish 4.83 Big skate 4.86

Stripetail rockfish 5.02 Spotted ratfish 2.53 Chilipepper rockfish* 4.74

Petrale sole 3.78 Arrowtooth flounder* 1.11 Sablefish 4.35

Yellowtail rockfish 3.55 Greenstriped rockfish 1.07 Greenstriped rockfish 3.06

Big skate 2.77 California skate 0.66 Yellowtail rockfish 1.75

Darkblotched rockfish 2.55 Curlfin sole 0.63 Spotted ratfish 1.63

Rosethorn rockfish 2.11 Spiny dogfish 0.44 Arrowtooth flounder* 1.14

Pygmy rockfish 1.61 Canary rockfish* 0.26 Darkblotched rockfish 0.54

Greenspotted rockfish 1.27 Darkblotched rockfish 0.16 Curlfin sole 0.5

Canary rockfish* 1.08 Shortbelly rockfish 0.06 Splitnose rockfish 0.18

Arrowtooth flounder* 1.03 Sharpchin rockfish 0.03 Redstripe rockfish 0.12

Spiny dogfish 0.86 Splitnose rockfish 0.02 California skate 0.1

Widow rockfish 0.66 Shortspine thornyhead 0.01 Greenspotted rockfish 0.07

Splitnose rockfish 0.59 Flathead sole <0.01 Bocaccio 0.03

Pacific cod 0.55 Shortbelly rockfish 0.02

Curlfin sole 0.37 Shortspine thornyhead 0.02

Bocaccio 0.19 Pacific ocean perch 0.01

Shortbelly rockfish 0.11 Widow rockfish 0.01

Chilipepper rockfish* 0.04 Sharpchin rockfish <0.01

Pacific ocean perch 0.03 Redbanded rockfish <0.01

Grooved tanner crab 0.03

Rougheye rockfish 0.02

California skate 0.02

Redbanded rockfish 0.01

Shortspine thornyhead <0.01
260

Table 5 continued. Mean CPUE (kg ha–1

) of 47 important FMP groundfish species plus grooved tanner 

crab caught by depth strata in the Eureka INPFC area during the 2003–2008 surveys. Stratum 1 

typically is 54.9–182.9 m, stratum 2 is 182.9–548.6 m, and stratum 3 is 548.6–1,280.2 m; 

however, for arrowtooth flounder and chilipepper rockfish, stratum 1 is 54.9–155 m and stratum 2 

is 155–548.6 m; and for canary rockfish, stratum 1 is 54.9–350 m and stratum 3 is defined as 

350–1,280.2 m with no stratum 2. These exceptions are noted with an asterisk (*).

Stratum 1

2006 (number of hauls = 24) 2007 (number of hauls = 28) 2008 (number of hauls = 24)

Lingcod 84.99 Pacific hake 37.76 Dover sole 16.41

Pacific hake 41.46 Dover sole 22.31 Pacific sanddab 10.92

Spiny dogfish 19.95 Lingcod 20.4 Greenstriped rockfish 9.97

Dover sole 16.97 Longnose skate 11 Stripetail rockfish 8.77

Canary rockfish* 13.88 Spiny dogfish 9.37 Longnose skate 7.12

Chilipepper rockfish* 12.4 English sole 8.7 English sole 5.81

Longnose skate 8.78 Spotted ratfish 7.16 Big skate 5.64

Spotted ratfish 8.14 Canary rockfish* 7.12 Petrale sole 4.7

English sole 7.53 Rex sole 6.72 Rex sole 3.43

Pacific sanddab 7.34 Petrale sole 5.95 Spotted ratfish 2.42

Petrale sole 5.56 Stripetail rockfish 5.59 Spiny dogfish 2.3

Rex sole 4.98 Pacific sanddab 4.71 Pacific hake 1.64

Big skate 3.97 Big skate 3.7 Arrowtooth flounder* 1.21

Stripetail rockfish 3.09 Darkblotched rockfish 2.25 Redstripe rockfish 1.02

Greenstriped rockfish 1.28 Redstripe rockfish 2.21 Lingcod 0.79

Sablefish 1.28 Yellowtail rockfish 2.06 Sablefish 0.5

Rosethorn rockfish 0.97 Sablefish 1.63 Splitnose rockfish 0.23

Widow rockfish 0.95 Greenstriped rockfish 1.62 Canary rockfish* 0.18

Splitnose rockfish 0.82 Greenspotted rockfish 1.22 Curlfin sole 0.16

Redstripe rockfish 0.31 Chilipepper rockfish* 1.1 Darkblotched rockfish 0.12

Greenspotted rockfish 0.3 Widow rockfish 0.52 Chilipepper rockfish* 0.09

Sharpchin rockfish 0.26 Arrowtooth flounder* 0.47 Pacific cod 0.06

Darkblotched rockfish 0.26 Bocaccio 0.19 Widow rockfish 0.04

Arrowtooth flounder* 0.17 Rosethorn rockfish 0.13 California skate 0.04

Yellowtail rockfish 0.02 Curlfin sole 0.12 Blackgill rockfish 0.03

Curlfin sole 0.01 Sharpchin rockfish 0.07 Sharpchin rockfish 0.02

Shortbelly rockfish <0.01 Splitnose rockfish 0.07 Rosethorn rockfish <0.01

Pygmy rockfish <0.01 Pacific ocean perch 0.02 Bocaccio <0.01

Pygmy rockfish 0.02

Shortbelly rockfish 0.02

Shortspine thornyhead 0.01
261

Table 5 continued. Mean CPUE (kg ha–1

) of 47 important FMP groundfish species plus grooved tanner 

crab caught by depth strata in the Eureka INPFC area during the 2003–2008 surveys. Stratum 1 

typically is 54.9–182.9 m, stratum 2 is 182.9–548.6 m, and stratum 3 is 548.6–1,280.2 m; 

however, for arrowtooth flounder and chilipepper rockfish, stratum 1 is 54.9–155 m and stratum 2 

is 155–548.6 m. These exceptions are noted with an asterisk (*).

Stratum 2

2003 (number of hauls = 36) 2004 (number of hauls = 12) 2005 (number of hauls = 23)

Dover sole 94.25 Pacific hake 43.71 Dover sole 117.87

Rex sole 25.5 English sole 29.27 Spotted ratfish 42.55

Pacific hake 22.27 Dover sole 25.98 Lingcod 39.68

Sharpchin rockfish 19.56 Rex sole 20.68 Splitnose rockfish 37.5

Sablefish 18.04 Petrale sole 11.21 Sharpchin rockfish 35.65

Longnose skate 14.87 Longnose skate 9.77 Rex sole 30.24

Stripetail rockfish 10.5 Pacific sanddab 8.58 Chilipepper rockfish* 21.36

Splitnose rockfish 9.01 Big skate 7.98 Sablefish 18.16

Spiny dogfish 5.69 Stripetail rockfish 6.42 Longnose skate 18.07

Chilipepper rockfish* 4.31 Lingcod 5.53 Pacific hake 17.7

Lingcod 3.79 Sablefish 4.83 Darkblotched rockfish 6.65

Darkblotched rockfish 3.65 Spotted ratfish 2.53 Shortspine thornyhead 6.27

Shortspine thornyhead 3.65 Arrowtooth flounder* 1.11 Spiny dogfish 3.85

Spotted ratfish 2.73 Greenstriped rockfish 1.07 Arrowtooth flounder* 3.25

Arrowtooth flounder* 2.53 California skate 0.66 Stripetail rockfish 3.14

English sole 2.23 Curlfin sole 0.63 Greenstriped rockfish 2.01

Greenstriped rockfish 1.36 Chilipepper rockfish* 0.44 Rosethorn rockfish 1.62

Aurora rockfish 1.09 Spiny dogfish 0.44 Longspine thornyhead 1.39

Longspine thornyhead 0.57 Darkblotched rockfish 0.16 Aurora rockfish 1.01

Big skate 0.34 Shortbelly rockfish 0.06 Redbanded rockfish 0.78

Redbanded rockfish 0.31 Sharpchin rockfish 0.03 English sole 0.57

Petrale sole 0.28 Splitnose rockfish 0.02 Pacific ocean perch 0.51

Rosethorn rockfish 0.18 Shortspine thornyhead 0.01 Big skate 0.48

Grooved tanner crab 0.16 Flathead sole <0.01 Widow rockfish 0.45

Pacific ocean perch 0.12 Bocaccio 0.31

Bocaccio 0.12 Petrale sole 0.1

Pacific cod 0.07 Redstripe rockfish 0.04

Pacific flatnose 0.05 Greenspotted rockfish 0.03

Widow rockfish 0.05 Pacific flatnose 0.03

Rougheye rockfish 0.04 Blackgill rockfish 0.01

Giant grenadier 0.02

Greenspotted rockfish 0.01

Pacific grenadier <0.01

Redstripe rockfish <0.01

Pacific sanddab <0.01

Shortbelly rockfish <0.01
262

Table 5 continued. Mean CPUE (kg ha–1

) of 47 important FMP groundfish species plus grooved tanner 

crab caught by depth strata in the Eureka INPFC area during the 2003–2008 surveys. Stratum 1 

typically is 54.9–182.9 m, stratum 2 is 182.9–548.6 m, and stratum 3 is 548.6–1,280.2 m; 

however, for arrowtooth flounder and chilipepper rockfish, stratum 1 is 54.9–155 m and stratum 2 

is 155–548.6 m. These exceptions are noted with an asterisk (*).

Stratum 2

2006 (number of hauls = 20) 2007 (number of hauls = 28) 2008 (number of hauls = 29)

Dover sole 105.94 Dover sole 107.75 Dover sole 105.39

Sablefish 67.55 Splitnose rockfish 58.5 Rex sole 22.63

Rex sole 29.68 Chilipepper rockfish* 51.1 Pacific hake 20.82

Pacific hake 22.21 Rex sole 23.46 Sablefish 16.36

Longnose skate 14.88 Sablefish 15.15 Longnose skate 15.75

Spiny dogfish 14.16 Pacific hake 14.67 Spotted ratfish 7.39

Splitnose rockfish 6.45 Stripetail rockfish 13.71 Lingcod 6.13

Arrowtooth flounder* 4.66 Longnose skate 12.89 Splitnose rockfish 5.78

Chilipepper rockfish* 4.65 Spiny dogfish 8.68 Arrowtooth flounder* 3.02

Spotted ratfish 4.55 Shortspine thornyhead 3.68 Shortspine thornyhead 2.96

Stripetail rockfish 4.41 Spotted ratfish 3.41 Stripetail rockfish 2.02

Shortspine thornyhead 2.58 Darkblotched rockfish 2.81 Darkblotched rockfish 1.96

Lingcod 2.18 Arrowtooth flounder* 2.71 Spiny dogfish 1.95

Darkblotched rockfish 1.38 Longspine thornyhead 0.94 Grooved tanner crab 1.94

Aurora rockfish 0.99 Lingcod 0.78 Longspine thornyhead 1.17

Redbanded rockfish 0.54 Aurora rockfish 0.71 Sharpchin rockfish 1.14

English sole 0.53 Grooved tanner crab 0.54 Aurora rockfish 0.76

Longspine thornyhead 0.26 Greenstriped rockfish 0.38 Greenstriped rockfish 0.64

Greenstriped rockfish 0.17 Redbanded rockfish 0.17 English sole 0.31

Rougheye rockfish 0.15 Sharpchin rockfish 0.17 Chilipepper rockfish* 0.31

Petrale sole 0.12 Widow rockfish 0.16 Redbanded rockfish 0.3

Pacific ocean perch 0.08 Shortbelly rockfish 0.14 Pacific ocean perch 0.16

Sharpchin rockfish 0.07 Pacific ocean perch 0.09 Blackgill rockfish 0.09

Grooved tanner crab 0.05 Pacific grenadier 0.06 Petrale sole 0.05

Rosethorn rockfish 0.03 Petrale sole 0.05 Rosethorn rockfish 0.05

Bocaccio 0.02 English sole 0.03 Greenspotted rockfish 0.05

Blackgill rockfish <0.01 Bank rockfish 0.02 Bank rockfish 0.03

Blackgill rockfish 0.02 Giant grenadier 0.02

Pacific flatnose 0.01 Big skate 0.01

Rosethorn rockfish 0.01 Pacific flatnose <0.01

Rougheye rockfish 0.01 Pacific grenadier <0.01
263

Table 5 continued. Mean CPUE (kg ha–1

) of 47 important FMP groundfish species plus grooved tanner 

crab caught by depth strata in the Eureka INPFC area during the 2003–2008 surveys. Stratum 1 

typically is 54.9–182.9 m, stratum 2 is 182.9–548.6 m, and stratum 3 is 548.6–1,280.2 m; 

however, for arrowtooth flounder, stratum 1 is 54.9–155 m and stratum 2 is 155–548.6 m. These 

exceptions are noted with an asterisk (*).

Stratum 3

2003 (number of hauls = 33) 2004 (number of hauls = 25) 2005 (number of hauls = 28)

Longspine thornyhead 35.72 Dover sole 39.07 Longspine thornyhead 48.90

Dover sole 22.89 Longspine thornyhead 31.91 Dover sole 32.91

Sablefish 18.48 Pacific grenadier 23.41 Sablefish 17.91

Giant grenadier 12.97 Giant grenadier 14.32 Grooved tanner crab 14.17

Grooved tanner crab 8.52 Sablefish 13.83 Shortspine thornyhead 5.56

Pacific grenadier 6.05 Shortspine thornyhead 6.34 Rex sole 3.52

Shortspine thornyhead 5.07 Grooved tanner crab 3.14 Pacific grenadier 3.03

Rex sole 1.46 Rex sole 2.28 Giant grenadier 2.41

Pacific hake 0.87 Pacific flatnose 1.25 Longnose skate 1.00

Pacific flatnose 0.62 Pacific hake 0.69 Pacific flatnose 0.42

Longnose skate 0.05 Longnose skate 0.42 Pacific hake 0.38

Spotted ratfish 0.02 Arrowtooth flounder* 0.03

Arrowtooth flounder* <0.01

2006 (number of hauls = 28) 2007 (number of hauls = 33) 2008 (number of hauls = 32)

Longspine thornyhead 45.77 Longspine thornyhead 53.13 Longspine thornyhead 63.14

Dover sole 31.17 Dover sole 36.58 Dover sole 36.65

Sablefish 16.04 Sablefish 14.49 Pacific grenadier 15.36

Grooved tanner crab 12.86 Grooved tanner crab 12.52 Sablefish 11.39

Shortspine thornyhead 6.19 Shortspine thornyhead 10.08 Grooved tanner crab 8.99

Giant grenadier 4.59 Giant grenadier 7.89 Giant grenadier 5.00

Pacific grenadier 3.86 Pacific grenadier 3.87 Shortspine thornyhead 4.79

Rex sole 2.11 Rex sole 1.85 Pacific flatnose 1.23

Pacific flatnose 0.48 Pacific flatnose 1.73 Rex sole 0.79

Longnose skate 0.38 Longnose skate 0.26 Pacific hake 0.09

Pacific hake 0.10 Pacific hake 0.03 Longnose skate 0.02
264

Table 6. Mean CPUE (kg ha–1

) of 47 important FMP groundfish species plus grooved tanner crab caught 

by depth strata in the Monterey INPFC area during the 2003–2008 surveys. Stratum 1 typically is 

54.9–182.9 m, stratum 2 is 182.9–548.6 m, and stratum 3 is 548.6–1,280.2 m; however, for 

chilipepper rockfish, stratum 1 is 54.9–155 m and stratum 2 is 155–548.6 m; and for canary 

rockfish, stratum 1 is 54.9–350 m and stratum 3 is defined as 350–1,280.2 m with no stratum 2. 

These exceptions are noted with an asterisk (*).

Stratum 1

2003 (number of hauls = 45) 2004 (number of hauls = 59) 2005 (number of hauls = 78)

Pacific sanddab 77.44 Spiny dogfish 76.38 Chilipepper rockfish* 83.35

Chilipepper rockfish* 71.30 Chilipepper rockfish* 40.31 Pacific hake 27.42

Spiny dogfish 60.52 Pacific sanddab 24.91 Shortbelly rockfish 18.9

Pacific hake 21.51 Pacific hake 18.02 Spiny dogfish 13.89

English sole 12.41 Shortbelly rockfish 17.40 Petrale sole 12.49

Stripetail rockfish 5.11 Lingcod 12.63 Pacific sanddab 11.25

Shortbelly rockfish 4.98 English sole 10.03 English sole 9.80

Petrale sole 4.95 Longnose skate 9.47 Longnose skate 6.20

Longnose skate 4.61 Bocaccio 8.03 Dover sole 5.11

Rex sole 3.36 Petrale sole 7.00 Stripetail rockfish 4.24

Greenstriped rockfish 3.17 Stripetail rockfish 6.86 Lingcod 3.71

Lingcod 2.83 Dover sole 6.24 Spotted ratfish 3.49

Spotted ratfish 2.20 Big skate 6.09 Rex sole 3.30

Big skate 1.81 Spotted ratfish 3.62 Big skate 2.91

California skate 1.71 California skate 3.52 Sablefish 2.29

Sablefish 1.59 Sablefish 3.50 California skate 1.90

Yellowtail rockfish 1.26 Rex sole 3.32 Greenstriped rockfish 1.21

Dover sole 1.03 Greenstriped rockfish 1.29 Bocaccio 0.90

Greenspotted rockfish 1.01 Halfbanded rockfish 1.28 Widow rockfish 0.81

Curlfin sole 0.81 Greenspotted rockfish 1.22 Halfbanded rockfish 0.76

Canary rockfish* 0.52 Yellowtail rockfish 0.87 Curlfin sole 0.53

Bocaccio 0.44 Canary rockfish* 0.77 Yellowtail rockfish 0.27

Pacific cod 0.12 Curlfin sole 0.49 Canary rockfish* 0.13

Widow rockfish 0.11 Squarespot rockfish 0.29 Greenspotted rockfish 0.12

Darkblotched rockfish 0.09 Widow rockfish 0.28 Darkblotched rockfish 0.03

Squarespot rockfish 0.05 Rosethorn rockfish 0.04 Sharpchin rockfish 0.02

Rosethorn rockfish 0.01 Darkblotched rockfish 0.04 Rosethorn rockfish 0.01

Shortspine thornyhead 0.01 Calico rockfish 0.03 Pygmy rockfish 0.01

Sharpchin rockfish 0.01 Sharpchin rockfish <0.01 Splitnose rockfish <0.01

Calico rockfish <0.01 Shortspine thornyhead <0.01

Blackgill rockfish <0.01
265

Table 6 continued. Mean CPUE (kg ha–1

) of 47 important FMP groundfish species plus grooved tanner 

crab caught by depth strata in the Monterey INPFC area during the 2003–2008 surveys. Stratum 

1 typically is 54.9–182.9 m, stratum 2 is 182.9–548.6 m, and stratum 3 is 548.6–1,280.2 m; 

however, for chilipepper rockfish, stratum 1 is 54.9–155 m and stratum 2 is 155–548.6 m; and for 

canary rockfish, stratum 1 is 54.9–350 m and stratum 3 is defined as 350–1,280.2 m with no 

stratum 2. These exceptions are noted with an asterisk (*).

Stratum 1

2006 (number of hauls = 61) 2007 (number of hauls = 56) 2008 (number of hauls = 70)

Chilipepper rockfish* 57.56 Pacific hake 17.56 Spiny dogfish 15.54

Spiny dogfish 27.94 Chilipepper rockfish* 12.69 Pacific sanddab 11.92

Pacific hake 10.80 Pacific sanddab 7.21 Pacific hake 11.50

Longnose skate 10.59 Petrale sole 6.48 Stripetail rockfish 11.28

Petrale sole 8.80 Spiny dogfish 5.76 Chilipepper rockfish* 5.48

Pacific sanddab 8.28 Yellowtail rockfish 5.65 Petrale sole 4.71

Dover sole 5.94 Spotted ratfish 4.96 Longnose skate 4.34

Lingcod 4.54 Longnose skate 4.24 Halfbanded rockfish 3.79

Stripetail rockfish 4.45 Dover sole 3.35 Greenstriped rockfish 3.36

Shortbelly rockfish 4.44 English sole 2.79 Dover sole 3.15

English sole 3.91 Rex sole 2.22 Spotted ratfish 2.81

Rex sole 2.92 Stripetail rockfish 2.17 English sole 2.47

Halfbanded rockfish 2.83 Big skate 1.89 Squarespot rockfish 1.56

Big skate 2.17 Lingcod 1.61 Bocaccio 1.44

California skate 1.66 California skate 1.48 Big skate 1.41

Spotted ratfish 1.57 Shortbelly rockfish 0.90 Lingcod 1.20

Canary rockfish* 1.54 Sablefish 0.75 Rex sole 1.15

Greenstriped rockfish 1.41 Greenstriped rockfish 0.51 California skate 1.02

Sablefish 0.73 Bocaccio 0.48 Shortbelly rockfish 0.89

Bocaccio 0.53 Canary rockfish* 0.34 Greenspotted rockfish 0.83

Yellowtail rockfish 0.29 Darkblotched rockfish 0.24 Pygmy rockfish 0.67

Greenspotted rockfish 0.23 Halfbanded rockfish 0.20 Sablefish 0.50

Curlfin sole 0.19 Widow rockfish 0.18 Yellowtail rockfish 0.48

Widow rockfish 0.19 Curlfin sole 0.14 Canary rockfish* 0.21

Darkblotched rockfish 0.06 Greenspotted rockfish 0.06 Curlfin sole 0.17

Squarespot rockfish 0.04 Grooved tanner crab 0.01 Redstripe rockfish 0.12

Rosethorn rockfish 0.03 Sharpchin rockfish <0.01 Rosethorn rockfish 0.11

Calico rockfish <0.01 Rosethorn rockfish <0.01 Widow rockfish 0.09

Sharpchin rockfish <0.01 Splitnose rockfish <0.01 Sharpchin rockfish 0.03

Splitnose rockfish <0.01 Aurora rockfish 0.01

Darkblotched rockfish <0.01

Splitnose rockfish <0.01
266

Table 6 continued. Mean CPUE (kg ha–1

) of 47 important FMP groundfish species plus grooved tanner 

crab caught by depth strata in the Monterey INPFC area during the 2003–2008 surveys. Stratum 

1 typically is 54.9–182.9 m, stratum 2 is 182.9–548.6 m, and stratum 3 is 548.6–1,280.2 m; 

however, for chilipepper rockfish, stratum 1 is 54.9–155 m and stratum 2 is 155–548.6 m. These 

exceptions are noted with an asterisk (*).

Stratum 2

2003 (number of hauls = 30) 2004 (number of hauls = 17) 2005 (number of hauls = 21)

Chilipepper rockfish* 137.71 Chilipepper rockfish* 148.16 Chilipepper rockfish* 102.64

Shortbelly rockfish 101.17 Sablefish 68.78 Shortbelly rockfish 126.41

Dover sole 79.36 Splitnose rockfish 64.72 Stripetail rockfish 52.94

Spotted ratfish 20.78 Dover sole 51.64 Dover sole 38.45

Splitnose rockfish 19.51 Pacific hake 41.00 Spiny dogfish 24.81

Pacific hake 17.91 Stripetail rockfish 40.27 Pacific hake 15.85

Rex sole 15.99 Shortbelly rockfish 19.64 Splitnose rockfish 12.83

Darkblotched rockfish 12.47 Rex sole 17.46 Rex sole 9.61

Stripetail rockfish 11.31 Longnose skate 15.50 Sablefish 9.51

Sablefish 9.90 Lingcod 5.77 Longnose skate 7.84

Longnose skate 6.98 Spiny dogfish 3.84 Bocaccio 6.89

Spiny dogfish 5.44 Petrale sole 3.73 Petrale sole 2.90

Shortspine thornyhead 4.29 Spotted ratfish 3.47 English sole 2.90

English sole 4.23 Darkblotched rockfish 2.79 Spotted ratfish 2.25

Lingcod 2.57 Sharpchin rockfish 2.41 Lingcod 2.10

Aurora rockfish 1.46 Bocaccio 2.37 Shortspine thornyhead 1.96

Petrale sole 1.43 Shortspine thornyhead 2.18 Greenstriped rockfish 1.72

Longspine thornyhead 0.94 Aurora rockfish 1.95 Darkblotched rockfish 1.06

Widow rockfish 0.85 English sole 1.82 Aurora rockfish 0.94

Bocaccio 0.72 Big skate 1.01 Longspine thornyhead 0.56

Blackgill rockfish 0.42 Greenstriped rockfish 0.46 Pacific sanddab 0.16

Grooved tanner crab 0.41 Greenspotted rockfish 0.35 Widow rockfish 0.13

Big skate 0.29 Blackgill rockfish 0.24 Blackgill rockfish 0.10

Redbanded rockfish 0.25 Widow rockfish 0.18 Redbanded rockfish 0.09

Rosethorn rockfish 0.21 Redbanded rockfish 0.12 Rosethorn rockfish 0.09

Greenstriped rockfish 0.10 Pacific sanddab 0.11 Grooved tanner crab 0.07

Giant grenadier 0.09 Longspine thornyhead 0.09 Bank rockfish 0.02

Pacific ocean perch 0.07 Giant grenadier 0.05 Greenspotted rockfish 0.02

Pacific sanddab 0.06 Bank rockfish 0.02 Pacific flatnose 0.02

Bank rockfish 0.03 Grooved tanner crab 0.02 Giant grenadier 0.01

Pacific flatnose 0.02 Halfbanded rockfish <0.01 California skate <0.01

Greenspotted rockfish 0.01 Pacific flatnose <0.01 Sharpchin rockfish <0.01

Sharpchin rockfish <0.01

Pygmy rockfish <0.01
267

Table 6 continued. Mean CPUE (kg ha–1

) of 47 important FMP groundfish species plus grooved tanner 

crab caught by depth strata in the Monterey INPFC area during the 2003–2008 surveys. Stratum 

1 typically is 54.9–182.9 m, stratum 2 is 182.9–548.6 m, and stratum 3 is 548.6–1,280.2 m; 

however, for chilipepper rockfish, stratum 1 is 54.9–155 m and stratum 2 is 155–548.6 m. These 

exceptions are noted with an asterisk (*).

Stratum 2

2006 (number of hauls = 32) 2007 (number of hauls = 18) 2008 (number of hauls = 32)

Chilipepper rockfish* 52.14 Chilipepper rockfish* 198.46 Chilipepper rockfish* 88.47

Dover sole 41.44 Dover sole 72.34 Dover sole 47.45

Splitnose rockfish 24.62 Splitnose rockfish 48.40 Splitnose rockfish 24.06

Rex sole 14.23 Spiny dogfish 41.35 Longnose skate 13.91

Spiny dogfish 13.70 Pacific hake 28.01 Stripetail rockfish 11.99

Sablefish 13.18 Rex sole 10.85 Pacific hake 11.14

Pacific hake 13.04 Stripetail rockfish 10.16 Rex sole 8.49

Stripetail rockfish 11.50 Sablefish 8.71 Spiny dogfish 7.64

Longnose skate 9.77 Longnose skate 8.57 Sablefish 5.16

Bank rockfish 8.68 Darkblotched rockfish 3.09 Grooved tanner crab 4.57

Blackgill rockfish 3.71 Shortspine thornyhead 2.69 Spotted ratfish 4.50

Shortspine thornyhead 3.54 Spotted ratfish 2.57 Shortspine thornyhead 2.32

Grooved tanner crab 2.72 Aurora rockfish 1.57 Petrale sole 1.23

Sharpchin rockfish 2.71 Grooved tanner crab 1.53 Aurora rockfish 1.13

Darkblotched rockfish 2.05 Longspine thornyhead 0.63 Longspine thornyhead 1.01

Spotted ratfish 1.80 Petrale sole 0.49 Darkblotched rockfish 0.90

English sole 1.72 Big skate 0.46 Bocaccio 0.88

Lingcod 1.67 English sole 0.40 Lingcod 0.85

Aurora rockfish 1.09 Greenspotted rockfish 0.25 Blackgill rockfish 0.73

Longspine thornyhead 0.99 Blackgill rockfish 0.23 English sole 0.41

Bocaccio 0.88 Greenstriped rockfish 0.21 Bank rockfish 0.17

Petrale sole 0.83 Lingcod 0.19 Widow rockfish 0.14

Greenspotted rockfish 0.70 Sharpchin rockfish 0.17 Redbanded rockfish 0.09

Widow rockfish 0.56 Bocaccio 0.16 Greenstriped rockfish 0.08

Rosethorn rockfish 0.52 Rosethorn rockfish 0.05 Rosethorn rockfish 0.02

Greenstriped rockfish 0.51 Redbanded rockfish 0.04 Pacific flatnose 0.02

Redbanded rockfish 0.32 Pacific flatnose 0.03 Shortbelly rockfish 0.01

Pacific flatnose 0.05 Pacific grenadier 0.03 Greenspotted rockfish 0.01

Shortbelly rockfish <0.01 Bank rockfish 0.02 Pacific grenadier 0.01

California skate <0.01 Sharpchin rockfish 0.01
268

Table 6 continued. Mean CPUE (kg ha–1

) of 47 important FMP groundfish species plus grooved tanner 

crab caught by depth strata in the Monterey INPFC area during the 2003–2008 surveys. Stratum 

1 typically is 54.9–182.9 m, stratum 2 is 182.9–548.6 m, and stratum 3 is 548.6–1,280.2 m.

Stratum 3

2003 (number of hauls = 26) 2004 (number of hauls = 21) 2005 (number of hauls = 20)

Dover sole 42.52 Dover sole 48.51 Dover sole 52.5

Pacific grenadier 32.96 Longspine thornyhead 39.11 Longspine thornyhead 32.41

Longspine thornyhead 28.86 Grooved tanner crab 11.64 Grooved tanner crab 17.78

Grooved tanner crab 20.39 Sablefish 9.93 Sablefish 16.71

Sablefish 16.43 Shortspine thornyhead 5.37 Pacific grenadier 14.87

Giant grenadier 12.32 Pacific grenadier 2.88 Shortspine thornyhead 6.60

Shortspine thornyhead 7.69 Giant grenadier 2.23 Giant grenadier 1.62

Pacific flatnose 1.48 Longnose skate 1.38 Longnose skate 1.44

Longnose skate 0.67 Aurora rockfish 0.45 Pacific flatnose 1.20

Pacific hake 0.65 Rex sole 0.39 Aurora rockfish 0.38

Aurora rockfish 0.49 Pacific flatnose 0.23 Pacific hake 0.07

Rex sole 0.37 Pacific hake 0.20 Spiny dogfish 0.04

California skate 0.02 Blackgill rockfish 0.04

Spiny dogfish 0.02

2006 (number of hauls = 37) 2007 (number of hauls = 29) 2008 (number of hauls = 39)

Dover sole 28.55 Dover sole 33.94 Dover sole 44.53

Longspine thornyhead 27.80 Longspine thornyhead 27.28 Longspine thornyhead 37.17

Grooved tanner crab 20.35 Pacific grenadier 21.34 Grooved tanner crab 15.92

Pacific grenadier 16.53 Grooved tanner crab 15.36 Pacific grenadier 14.15

Sablefish 14.96 Sablefish 12.10 Sablefish 10.57

Shortspine thornyhead 6.58 Shortspine thornyhead 6.05 Shortspine thornyhead 6.79

Giant grenadier 5.83 Giant grenadier 4.48 Giant grenadier 3.57

Pacific flatnose 1.93 Pacific flatnose 1.76 Longnose skate 0.74

Longnose skate 0.97 Rex sole 0.87 Pacific flatnose 0.70

Rex sole 0.37 Longnose skate 0.70 Pacific hake 0.14

Aurora rockfish 0.24 Aurora rockfish 0.09 Rex sole 0.06

Pacific hake 0.20 Pacific hake 0.07 Aurora rockfish 0.05

Blackgill rockfish 0.04
269

Table 7. Mean CPUE (kg ha–1

) of 47 important FMP groundfish species plus grooved tanner crab caught 

by depth strata in the Conception INPFC area during the 2003–2008 surveys. Stratum 1 typically 

is 54.9–182.9 m, stratum 2 is 182.9–548.6 m, and stratum 3 is 548.6–1,280.2 m; however, for 

chilipepper rockfish, stratum 1 is 54.9–155 m and stratum 2 is 155–548.6 m; and for canary 

rockfish, stratum 1 is 54.9–350 m and stratum 3 is defined as 350–1,280.2 m with no stratum 2. 

These exceptions are noted with an asterisk (*).

Stratum 1

2003 (number of hauls = 52) 2004 (number of hauls = 47) 2005 (number of hauls = 54)

Pacific sanddab 14.67 Halfbanded rockfish 12.58 Halfbanded rockfish 31.99

Spotted ratfish 2.16 Spotted ratfish 5.80 Spotted ratfish 15.23

English sole 1.65 Pacific sanddab 5.37 Pacific sanddab 7.01

Petrale sole 0.88 Spiny dogfish 1.21 English sole 2.68

California skate 0.79 English sole 1.20 Spiny dogfish 2.62

Halfbanded rockfish 0.76 Stripetail rockfish 0.94 Shortbelly rockfish 2.08

California scorpionfish 0.64 California skate 0.93 Squarespot rockfish 1.94

Lingcod 0.63 California scorpionfish 0.92 Chilipepper rockfish* 1.85

Spiny dogfish 0.61 Pacific hake 0.87 California scorpionfish 1.47

Stripetail rockfish 0.60 Lingcod 0.84 Canary rockfish* 1.40

Big skate 0.56 Petrale sole 0.80 Rosethorn rockfish 1.25

Sablefish 0.38 Bocaccio 0.77 Pygmy rockfish 1.04

Longnose skate 0.31 Greenspotted rockfish 0.68 Greenspotted rockfish 0.96

Curlfin sole 0.27 Longnose skate 0.61 Big skate 0.89

Bocaccio 0.24 Big skate 0.56 Stripetail rockfish 0.80

Greenspotted rockfish 0.23 Canary rockfish* 0.53 Pacific hake 0.67

Pacific hake 0.21 Widow rockfish 0.32 California skate 0.67

Chilipepper rockfish* 0.16 Chilipepper rockfish* 0.25 Petrale sole 0.66

Shortbelly rockfish 0.13 Curlfin sole 0.21 Lingcod 0.53

Rex sole 0.05 Shortbelly rockfish 0.16 Bocaccio 0.49

Aurora rockfish 0.03 Greenstriped rockfish 0.15 Yellowtail rockfish 0.30

Canary rockfish* 0.03 Yellowtail rockfish 0.14 Widow rockfish 0.29

Dover sole 0.03 Rex sole 0.13 Greenstriped rockfish 0.18

Greenstriped rockfish 0.01 Splitnose rockfish 0.09 Calico rockfish 0.16

Rosethorn rockfish 0.01 Sablefish 0.05 Longnose skate 0.14

Yellowtail rockfish 0.01 Dover sole 0.05 Bank rockfish 0.12

Squarespot rockfish <0.01 Rosethorn rockfish 0.03 Curlfin sole 0.09

Calico rockfish <0.01 Squarespot rockfish 0.03 Dover sole 0.07

Darkblotched rockfish <0.01 Darkblotched rockfish 0.02 Sablefish 0.06

Splitnose rockfish <0.01 Calico rockfish 0.02 Rex sole 0.05

Bank rockfish 0.01 Redbanded rockfish 0.03

Splitnose rockfish <0.01

Sharpchin rockfish <0.01

Flathead sole <0.01
270

Table 7 continued. Mean CPUE (kg ha–1

) of 47 important FMP groundfish species plus grooved tanner 

crab caught by depth strata in the Conception INPFC area during the 2003–2008 surveys. 

Stratum 1 typically is 54.9–182.9 m, stratum 2 is 182.9–548.6 m, and stratum 3 is 548.6–1,280.2 

m; however, for chilipepper rockfish, stratum 1 is 54.9–155 m and stratum 2 is 155–548.6 m; and 

for canary rockfish, stratum 1 is 54.9–350 m and stratum 3 is defined as 350–1,280.2 m with no 

stratum 2. These exceptions are noted with an asterisk (*).

Stratum 1

2006 (number of hauls = 48 2007 (number of hauls = 55) 2008 (number of hauls = 51)

Pacific sanddab 14.06 Halfbanded rockfish 16.10 Pacific sanddab 8.58

Halfbanded rockfish 7.03 Pacific sanddab 10.49 Spotted ratfish 5.47

Pacific hake 4.66 Spotted ratfish 7.59 Halfbanded rockfish 5.37

English sole 2.02 Greenspotted rockfish 2.54 Chilipepper rockfish* 2.18

Bocaccio 1.60 Chilipepper rockfish* 1.46 English sole 1.34

Spotted ratfish 1.52 Bocaccio 1.43 Spiny dogfish 0.77

Greenspotted rockfish 1.39 Shortbelly rockfish 1.42 Canary rockfish* 0.68

Spiny dogfish 1.37 English sole 1.21 Greenspotted rockfish 0.68

Stripetail rockfish 1.30 Rosethorn rockfish 1.10 California skate 0.66

Shortbelly rockfish 0.95 California scorpionfish 1.05 Lingcod 0.65

Petrale sole 0.88 Spiny dogfish 0.99 Petrale sole 0.64

Big skate 0.77 Greenstriped rockfish 0.81 Big skate 0.38

Longnose skate 0.66 Lingcod 0.75 Greenstriped rockfish 0.31

California skate 0.65 Petrale sole 0.63 Stripetail rockfish 0.29

Lingcod 0.63 Pacific hake 0.61 California scorpionfish 0.24

California scorpionfish 0.40 Squarespot rockfish 0.59 Curlfin sole 0.21

Greenstriped rockfish 0.40 California skate 0.55 Shortbelly rockfish 0.18

Rosethorn rockfish 0.37 Big skate 0.53 Longnose skate 0.16

Rex sole 0.23 Longnose skate 0.43 Squarespot rockfish 0.11

Chilipepper rockfish* 0.19 Stripetail rockfish 0.42 Bocaccio 0.09

Squarespot rockfish 0.14 Pygmy rockfish 0.24 Dover sole 0.08

Curlfin sole 0.12 Bank rockfish 0.19 Sablefish 0.05

Dover sole 0.12 Rex sole 0.17 Rex sole 0.04

Bank rockfish 0.05 Splitnose rockfish 0.14 Pacific hake 0.04

Pygmy rockfish 0.02 Dover sole 0.11 Calico rockfish 0.02

Calico rockfish 0.01 Curlfin sole 0.09 Pygmy rockfish <0.01

Canary rockfish* 0.01 Calico rockfish 0.02 Bank rockfish <0.01

Splitnose rockfish <0.01 Sablefish 0.01 Rosethorn rockfish <0.01

Sablefish <0.01 Canary rockfish* 0.01 Redbanded rockfish <0.01

Darkblotched rockfish <0.01
271

Table 7 continued. Mean CPUE (kg ha–1

) of 47 important FMP groundfish species plus grooved tanner 

crab caught by depth strata in the Conception INPFC area during the 2003–2008 surveys. 

Stratum 1 typically is 54.9–182.9 m, stratum 2 is 182.9–548.6 m, and stratum 3 is 548.6–1,280.2 

m; however, for chilipepper rockfish, stratum 1 is 54.9–155 m and stratum 2 is 155–548.6 m. 

These exceptions are noted with an asterisk (*).

Stratum 2

2003 (number of hauls = 54 2004 (number of hauls = 46) 2005 (number of hauls = 75)

Chilipepper rockfish* 17.99 Splitnose rockfish 17.21 Splitnose rockfish 14.95

Dover sole 11.05 Shortbelly rockfish 16.09 Dover sole 14.23

Splitnose rockfish 10.72 Dover sole 13.06 Shortbelly rockfish 8.17

Pacific hake 9.11 Pacific hake 13.00 Sablefish 7.31

Shortbelly rockfish 7.19 Longnose skate 5.35 Pacific hake 6.61

Longnose skate 6.41 Sablefish 4.16 Longnose skate 6.48

Rex sole 4.95 Rex sole 4.12 Chilipepper rockfish* 6.12

Sablefish 2.85 Stripetail rockfish 3.52 Rex sole 5.04

Stripetail rockfish 2.77 Lingcod 3.51 Shortspine thornyhead 4.09

Longspine thornyhead 2.25 Chilipepper rockfish* 3.08 Aurora rockfish 2.50

Aurora rockfish 2.07 Blackgill rockfish 2.75 Longspine thornyhead 2.38

Shortspine thornyhead 2.06 Shortspine thornyhead 2.06 Blackgill rockfish 2.08

Spiny dogfish 2.03 Aurora rockfish 1.71 Stripetail rockfish 1.67

Spotted ratfish 1.55 Pacific sanddab 1.46 Spotted ratfish 1.18

Blackgill rockfish 0.69 Spotted ratfish 1.22 Rosethorn rockfish 0.93

Petrale sole 0.62 English sole 1.03 Spiny dogfish 0.54

Lingcod 0.61 Longspine thornyhead 0.95 Lingcod 0.46

English sole 0.58 Petrale sole 0.90 Petrale sole 0.36

Bocaccio 0.30 Spiny dogfish 0.55 Grooved tanner crab 0.18

Pacific sanddab 0.20 Rosethorn rockfish 0.32 Big skate 0.09

Greenspotted rockfish 0.18 Bocaccio 0.32 Bank rockfish 0.06

Greenstriped rockfish 0.12 Greenstriped rockfish 0.21 Bocaccio 0.05

Rosethorn rockfish 0.07 Greenspotted rockfish 0.15 Darkblotched rockfish 0.02

Redbanded rockfish 0.03 Bank rockfish 0.15 English sole 0.01

Darkblotched rockfish 0.02 Darkblotched rockfish 0.07 Redbanded rockfish 0.01

California skate 0.02 Grooved tanner crab 0.03 Greenstriped rockfish 0.01

California scorpionfish 0.01 California skate 0.02 Pacific sanddab 0.01

Curlfin sole 0.01 Curlfin sole 0.02 Pacific grenadier 0.01

Grooved tanner crab 0.01 Redbanded rockfish 0.01 Greenspotted rockfish 0.01

Halfbanded rockfish <0.01 Rougheye rockfish 0.01 Widow rockfish <0.01

Bank rockfish <0.01 Widow rockfish <0.01 Sharpchin rockfish <0.01

Halfbanded rockfish <0.01
272

Table 7 continued. Mean CPUE (kg ha–1

) of 47 important FMP groundfish species plus grooved tanner 

crab caught by depth strata in the Conception INPFC area during the 2003–2008 surveys. 

Stratum 1 typically is 54.9–182.9 m, stratum 2 is 182.9–548.6 m, and stratum 3 is 548.6–1,280.2 

m; however, for chilipepper rockfish, stratum 1 is 54.9–155 m and stratum 2 is 155–548.6 m. 

These exceptions are noted with an asterisk (*).

Stratum 2

2006 (number of hauls = 72 2007 (number of hauls = 85) 2008 (number of hauls = 82)

Dover sole 14.36 Dover sole 14.65 Dover sole 16.46

Pacific hake 6.54 Splitnose rockfish 11.66 Splitnose rockfish 15.67

Longnose skate 5.81 Pacific hake 10.60 Longnose skate 14.03

Splitnose rockfish 4.95 Longnose skate 7.16 Pacific hake 12.42

Shortbelly rockfish 4.86 Chilipepper rockfish* 4.77 Shortbelly rockfish 8.23

Sablefish 4.61 Sablefish 4.68 Sablefish 4.91

Shortspine thornyhead 4.00 Rex sole 4.25 Rex sole 3.71

Rex sole 3.11 Shortspine thornyhead 3.20 Shortspine thornyhead 3.47

Blackgill rockfish 2.91 Aurora rockfish 2.58 Chilipepper rockfish* 3.09

Chilipepper rockfish* 2.58 Spotted ratfish 2.17 Aurora rockfish 2.85

Aurora rockfish 2.35 Longspine thornyhead 2.07 Longspine thornyhead 2.50

Longspine thornyhead 2.20 Stripetail rockfish 1.96 Spiny dogfish 2.10

Stripetail rockfish 2.11 Blackgill rockfish 1.43 Spotted ratfish 1.82

Spotted ratfish 1.21 Shortbelly rockfish 0.70 Stripetail rockfish 1.47

Bank rockfish 0.95 Petrale sole 0.49 Blackgill rockfish 1.23

Spiny dogfish 0.32 Spiny dogfish 0.38 Pacific sanddab 0.37

English sole 0.27 English sole 0.37 Halfbanded rockfish 0.36

Lingcod 0.23 Pacific sanddab 0.26 Grooved tanner crab 0.33

Greenstriped rockfish 0.23 Bocaccio 0.21 English sole 0.33

Petrale sole 0.17 Grooved tanner crab 0.11 Petrale sole 0.19

Grooved tanner crab 0.13 Lingcod 0.11 Big skate 0.11

Pacific sanddab 0.12 California skate 0.06 Lingcod 0.08

Greenspotted rockfish 0.09 Greenstriped rockfish 0.06 Bocaccio 0.07

Big skate 0.09 Halfbanded rockfish 0.03 Bank rockfish 0.05

Sharpchin rockfish 0.05 Rosethorn rockfish 0.03 California skate 0.03

Rosethorn rockfish 0.03 Widow rockfish 0.03 Greenstriped rockfish 0.03

Bocaccio 0.02 Redbanded rockfish 0.02 Greenspotted rockfish 0.02

Curlfin sole 0.01 Bank rockfish 0.02 Darkblotched rockfish 0.02

Giant grenadier 0.01 Greenspotted rockfish 0.01 Redbanded rockfish 0.01

Halfbanded rockfish 0.01 Darkblotched rockfish 0.01 Rosethorn rockfish <0.01

Darkblotched rockfish <0.01 Curlfin sole 0.01 Curlfin sole <0.01

Pacific grenadier <0.01 Squarespot rockfish <0.01

Redbanded rockfish <0.01 Redstripe rockfish <0.01

California skate <0.01

Rougheye rockfish <0.01
273

Table 7 continued. Mean CPUE (kg ha–1

) of 47 important FMP groundfish species plus grooved tanner 

crab caught by depth strata in the Conception INPFC area during the 2003–2008 surveys. 

Stratum 1 typically is 54.9–182.9 m, stratum 2 is 182.9–548.6 m, and stratum 3 is 548.6–1,280.2 

m.

Stratum 3

2003 (number of hauls = 28) 2004 (number of hauls = 53) 2005 (number of hauls = 72)

Longspine thornyhead 16.27 Longspine thornyhead 19.57 Longspine thornyhead 13.51

Dover sole 10.23 Dover sole 13.31 Dover sole 13.18

Shortspine thornyhead 6.89 Sablefish 7.67 Sablefish 7.53

Sablefish 4.38 Shortspine thornyhead 6.82 Shortspine thornyhead 5.54

Giant grenadier 0.82 Pacific grenadier 1.79 Pacific grenadier 1.79

Grooved tanner crab 0.74 Grooved tanner crab 1.57 Grooved tanner crab 1.73

Pacific grenadier 0.66 Giant grenadier 1.32 Longnose skate 0.78

Pacific hake 0.64 Longnose skate 0.49 Giant grenadier 0.74

Longnose skate 0.60 Pacific flatnose 0.37 Pacific flatnose 0.36

Aurora rockfish 0.43 Pacific hake 0.22 Aurora rockfish 0.23

Pacific flatnose 0.30 Aurora rockfish 0.18 Pacific hake 0.14

Rex sole 0.03 Rex sole 0.01 Spotted ratfish 0.13

California skate 0.01 Rex sole 0.01

California skate 0.01

2006 (number of hauls = 70) 2007 (number of hauls = 71) 2008 (number of hauls = 64)

Longspine thornyhead 15.65 Dover sole 18.86 Longspine thornyhead 16.97

Dover sole 13.66 Longspine thornyhead 16.05 Dover sole 14.25

Sablefish 7.24 Sablefish 11.44 Sablefish 5.82

Pacific grenadier 6.71 Shortspine thornyhead 6.92 Shortspine thornyhead 4.61

Shortspine thornyhead 5.95 Pacific grenadier 3.46 Pacific grenadier 1.41

Grooved tanner crab 1.75 Grooved tanner crab 1.67 Grooved tanner crab 1.30

Giant grenadier 1.02 Giant grenadier 1.64 Giant grenadier 0.88

Pacific flatnose 0.78 Pacific flatnose 0.78 Longnose skate 0.50

Longnose skate 0.58 Longnose skate 0.35 Pacific flatnose 0.36

Pacific hake 0.14 Aurora rockfish 0.12 Aurora rockfish 0.33

Aurora rockfish 0.08 Pacific hake 0.08 Pacific hake 0.26

Rex sole 0.06 Rex sole 0.07 Rex sole 0.01

Spotted ratfish 0.01 Spotted ratfish 0.01 Blackgill rockfish <0.01

Blackgill rockfish <0.01

Splitnose rockfish <0.01
274

Table 8. Number of individual length measurements and age structures collected by species during the surveys from 2003 to 2008. Dorsal spines

were collected for spiny dogfish, dorsal finrays for lingcod, and otoliths for all other species.

Species Lengths Ages Species Lengths Ages Species Lengths Ages

Arrowtooth flounder 21,138 5,466 Greenblotched r’fish 308 207 Rosy rockfish 280 103

Aurora rockfish 9,113 3,407 Greenspotted rockfish 2,954 1,262 Rougheye rockfish 814 599

Bank rockfish 571 264 Greenstriped rockfish 17,678 3,809 Sablefish 29,690 14,562

Big skate 1,326 — Grooved tanner crab 18,410 — Sand sole 252 62

Black rockfish 1 1 Halfbanded rockfish 8,286 1,519 Sharpchin rockfish 7,602 1,718

Blackgill rockfish 2,394 1,119 Honeycomb rockfish 191 95 Shortbelly rockfish 10,527 2,780

Blue rockfish 44 19 Kelp greenling 285 126 Shortraker rockfish 19 16

Bocaccio 1,458 853 Lingcod 6,835 4,921 Shortspine thornyhead 39,094 8,270

Bronzespotted rockfish 1 1 Longnose skate 19,218 — Silvergray rockfish 153 148

Brown rockfish 165 86 Longspine thornyhead 60,356 5,796 Soupfin shark 5 —

Butter sole 401 — Mexican rockfish 1 — Southern rock sole 954 —

Cabezon 1 1 Olive rockfish 3 3 Speckled rockfish 123 42

Calico rockfish 510 330 Pacific cod 1,720 — Spiny dogfish 19,474 4,636

California scorpionfish 1,055 364 Pacific flatnose 7,510 270 Splitnose rockfish 26,202 4,381

California skate 2,112 — Pacific grenadier 19,238 3,857 Spotted ratfish 18,063 —

Canary rockfish 3,690 2,047 Pacific hake 36,380 4,100 Squarespot rockfish 1,427 365

Chilipepper 18,044 4,388 Pacific ocean perch 4,350 1,939 Starry flounder 166 143

Copper rockfish 242 186 Pacific sanddab 47,840 5,820 Starry rockfish 17 16

Cowcod 176 176 Petrale sole 21,384 8,541 Stripetail rockfish 22,888 2,620

Curlfin sole 2,646 265 Pink rockfish 44 43 Swordspine rockfish 1,642 242

Darkblotched rockfish 11,078 4,864 Pinkrose rockfish 39 29 Tiger rockfish 9 7

Dover sole 90,417 13,396 Pygmy rockfish 1,196 306 Tree rockfish 1 1

English sole 39,000 6,967 Quillback rockfish 65 43 Vermilion rockfish 363 205

Flag rockfish 213 129 Redbanded rockfish 1,077 963 Widow rockfish 885 349

Flathead sole 3,348 — Redstripe rockfish 3,031 902 Yelloweye rockfish 241 241

Freckled rockfish 79 44 Rex sole 70,145 1,571 Yellowmouth rockfish 164 57

Gopher rockfish 7 3 Rosethorn rockfish 7,247 2,429 Yellowtail rockfish 4,722 1,648
275

Table 9. Number of individual length measurements and age structures collected by species by year during the surveys from 2003 to 2008. Dorsal 

spines were collected for spiny dogfish, dorsal finrays for lingcod, and otoliths for all other species.

2003 2004 2005 2006 2007 2008

Species Lengths Ages Lengths Ages Lengths Ages Lengths Ages Lengths Ages Lengths Ages

Arrowtooth flounder 4,542 1,386 2,765 704 3,991 867 3,036 735 3,553 895 3,251 879

Aurora rockfish 1,176 575 1,087 354 1,699 577 1,749 609 1,694 586 1,708 706

Bank rockfish 6 4 106 60 96 50 257 64 75 55 31 31

Big skate 203 — 267 — 343 — 154 — 197 0 162 —

Black rockfish — — — — — — — — — — — —

Blackgill rockfish 146 104 449 175 398 191 764 212 298 194 339 243

Blue rockfish 19 — 2 2 21 15 — — — — 2 2

Bocaccio 113 113 493 222 319 196 265 138 157 98 111 86

Bronzespotted rockfish — — — — — — 1 1 — — — —

Brown rockfish 56 — 43 43 48 25 13 13 1 1 4 4

Butter sole 89 — 7 — 132 — 4 — 121 — 48 —

Cabezon — — — — — — — — — — 1 1

Calico rockfish 12 — 84 84 258 119 51 51 55 47 50 29

California scorpionfish 142 — 180 — 362 147 99 73 217 89 55 55

California skate 279 — 489 — 457 — 316 — 273 — 298 —

Canary rockfish 423 278 550 289 622 277 623 247 673 497 799 459

Chilipepper 2,658 716 3,802 855 4,102 892 2,754 626 2,519 596 2,209 703

Copper rockfish 29 — 76 76 70 43 10 10 13 13 44 44

Cowcod 13 13 65 65 32 32 25 25 25 25 16 16

Curlfin sole 700 — 629 — 518 — 299 — 201 — 299 265

Darkblotched rockfish 2,375 772 1,062 595 1,983 804 1,925 944 2,086 987 1,647 762

Dover sole 23,330 3,019 17,878 2,569 18,008 2,572 13,615 2,008 11,322 2,151 6,264 1,077

English sole 8,380 1,840 8,697 994 8,780 1,332 5,529 1,001 4,088 955 3,526 845

Flag rockfish 78 — 29 29 17 17 26 26 55 49 8 8

Flathead sole 1,521 — 519 — 593 — 421 — 137 — 157 —

Freckled rockfish — — — — 79 44 — — — — — —

Gopher rockfish 3 — 1 — 1 1 2 2 — — — —

Greenblotched 

rockfish

43 — 59 59 30 30 74 40 31 27 71 51

Greenspotted rockfish 424 — 567 283 276 153 475 273 655 309 557 244

Greenstriped rockfish 3,557 625 2,718 541 4,008 722 3,167 687 2,362 665 1,866 569

Grooved tanner crab — — — — 5,441 — 5,911 — 3,560 — 3,498 —
276

Table 9 continued. Number of individual length measurements and age structures collected by species by year during the surveys from 2003 to 

2008. Dorsal spines were collected for spiny dogfish, dorsal finrays for lingcod, and otoliths for all other species.

2003 2004 2005 2006 2007 2008

Species Lengths Ages Lengths Ages Lengths Ages Lengths Ages Lengths Ages Lengths Ages

Halfbanded rockfish — — 1,693 439 2,749 418 1,549 220 1,213 234 1,082 208

Honeycomb rockfish 1 — 16 16 91 33 14 14 68 31 1 1

Kelp greenling 61 — 97 — 46 45 23 23 39 39 19 19

Lingcod 1,381 931 1,436 907 1,182 944 1,022 721 650 557 1,164 861

Longnose skate 2,673 — 2,644 — 3,328 — 3,323 — 3,867 — 3,383 —

Longspine thornyhead 15,595 970 11,703 772 13,975 1,149 9,180 946 6,249 962 3,654 997

Mexican rockfish — — 1 — — — — — — — — —

Olive rockfish — — 1 1 2 2 — — — — — —

Pacific cod 785 — 304 — 199 — 166 — 177 — 89 —

Pacific flatnose 1,973 — — — 1,206 — 1,743 — 1,689 270 899 —

Pacific grenadier 4,463 1,141 2,048 350 2,643 548 4,334 599 3,157 641 2,593 578

Pacific hake 5,947 1,714 3,893 — 9,140 — 7,372 — 5,465 1,233 4,563 1,153

Pacific ocean perch 1,426 434 565 219 533 264 659 255 628 439 539 328

Pacific sanddab 9,690 838 11,935 1,565 10,957 1,122 6,130 744 4,559 768 4,569 783

Petrale sole 2,868 1,631 3,509 1,973 4,732 1,919 3,745 986 3,461 995 3,069 1,037

Pink rockfish 1 — — — — — 22 22 21 21 — —

Pinkrose rockfish — — — — — — 12 12 27 17 — —

Pygmy rockfish — — 171 46 196 53 405 81 248 77 176 49

Quillback rockfish 3 — — — 2 2 3 3 20 16 37 22

Redbanded rockfish 272 179 158 144 171 171 152 145 178 178 146 146

Redstripe rockfish 1,068 246 527 179 240 113 507 131 356 102 333 131

Rex sole 11,047 — 14,388 — 16,173 — 11,641 — 9,646 805 7,250 766

Rosethorn rockfish 1,604 382 1,003 353 931 325 1,578 496 1,320 507 811 366

Rosy rockfish — — 3 3 14 14 — — 41 16 222 70

Rougheye rockfish 112 56 115 78 259 140 99 97 108 107 121 121

Sablefish 5,938 2,402 4,671 2,204 5,721 2,946 4,904 2,696 4,477 2,123 3,979 2,191

Sand sole 81 — 29 — 31 — 12 — 37 — 62 62

Sharpchin rockfish 2,355 553 1,176 199 890 158 1,283 239 975 204 923 365

Shortbelly rockfish 2,133 638 2,920 479 2,154 572 1,445 387 1,034 376 841 328

Shortraker rockfish 3 — 8 8 8 8 — — — — — —

Shortspine thornyhead 7,712 1,269 6,766 1,719 8,152 1,415 6,240 1,261 5,506 1,298 4,718 1,308

Silvergray rockfish 34 34 19 19 51 46 7 7 22 22 20 20
277

Table 9 continued. Number of individual length measurements and age structures collected by species by year during the surveys from 2003 to 

2008. Dorsal spines were collected for spiny dogfish, dorsal finrays for lingcod, and otoliths for all other species.

2003 2004 2005 2006 2007 2008

Species Lengths Ages Lengths Ages Lengths Ages Lengths Ages Lengths Ages Lengths Ages

Soupfin shark 1 — — — 3 — — — — — 1 —

Southern rock sole 25 — 174 — 165 — 168 — 175 — 247 —

Speckled rockfish — — — — — — 17 17 106 25 — —

Spiny dogfish 3,847 679 2,616 582 3,727 931 3,903 789 2,479 763 2,902 892

Splitnose rockfish 6,633 1,068 5,229 437 4,467 543 3,592 619 3,363 884 2,918 830

Spotted ratfish 4,871 — — — 3,736 — 3,293 — 3,524 — 2,639 —

Squarespot rockfish 30 — 166 26 416 117 152 49 424 93 239 80

Starry flounder 6 — 51 45 54 43 7 7 25 25 23 23

Starry rockfish 1 — 3 3 4 4 1 1 6 6 2 2

Stripetail rockfish 5,945 — 4,852 499 4,647 591 3,330 538 2,302 555 1,812 437

Swordspine rockfish — — 305 49 51 — 524 55 453 74 309 64

Tiger rockfish 2 — — — — — 3 3 4 4 — —

Tree rockfish — — — — 1 1 — — — — — —

Vermilion rockfish 63 — 20 19 58 37 27 27 121 57 74 65

Widow rockfish 216 10 182 58 197 83 172 89 92 83 26 26

Yelloweye rockfish 68 68 21 21 40 40 43 43 26 26 43 43

Yellowmouth rockfish 141 34 — — 23 23 — — — — — —

Yellowtail rockfish 849 275 703 203 1,134 365 388 171 958 279 690 355
278

Table 10. Number of length measurements collected by stratum for FMP species during the surveys from 

2003 to 2008.

Stratum 1 Stratum 2 Stratum 3 All strata

Species (55–183 m) (184–549 m) (550–1,280 m) (55–1,280 m)

Arrowtooth flounder 13,988 7,118 32 21,138

Aurora rockfish 6 8,606 501 9,113

Bank rockfish 185 386 — 571

Big skate 1,297 29 — 1,326

Black rockfish 1 — — 1

Blackgill rockfish 2 2,387 5 2,394

Blue rockfish 44 — — 44

Bocaccio 1,160 298 — 1,458

Bronzespotted rockfish — 1 — 1

Brown rockfish 165 — — 165

Butter sole 401 — — 401

Cabezon 1 — — 1

Calico rockfish 510 — — 510

California scorpionfish 1,052 3 — 1,055

California skate 2,088 22 2 2,112

Canary rockfish 3,511 179 — 3,690

Chilipepper 13,619 4,425 — 18,044

Copper rockfish 240 2 — 242

Cowcod 100 76 — 176

Curlfin sole 2,618 28 — 2,646

Darkblotched rockfish 4,419 6,659 — 11,078

Dover sole 34,399 38,491 17,527 90,417

English sole 35,723 3,277 — 39,000

Flag rockfish 188 25 — 213

Flathead sole 2,730 618 — 3,348

Freckled rockfish 79 — — 79

Gopher rockfish 7 — — 7

Greenblotched rockfish 103 205 — 308

Greenspotted rockfish 2,692 262 — 2,954

Greenstriped rockfish 14,230 3,448 — 17,678

Halfbanded rockfish 8,114 172 — 8,286

Honeycomb rockfish 190 1 — 191

Kelp greenling 285 — — 285

Lingcod 5,792 1,043 — 6,835

Longnose skate 9,038 9,689 491 19,218

Longspine thornyhead 1 9,707 50,648 60,356

Mexican rockfish — 1 — 1

Olive rockfish 3 — — 3

Pacific cod 1,532 188 — 1,720

Pacific flatnose — 170 7,340 7,510

Pacific grenadier — 102 19,136 19,238

Pacific hake 18,111 17,748 521 36,380

Pacific ocean perch 554 3,792 4 4,350

Pacific sanddab 47,061 779 — 47,840

Petrale sole 20,257 1,127 — 21,384

Pink rockfish 15 29 — 44
279

Table 10 continued. Number of length measurements collected by stratum for FMP species during the 

surveys from 2003 to 2008.

Stratum 1 Stratum 2 Stratum 3 All strata

Species (55–183 m) (184–549 m) (550–1,280 m) (55–1,280 m)

Pinkrose rockfish — 39 — 39

Pygmy rockfish 1,195 1 — 1,196

Quillback rockfish 65 — — 65

Redbanded rockfish 111 965 1 1,077

Redstripe rockfish 2,730 301 — 3,031

Rex sole 40,739 27,849 1,557 70,145

Rosethorn rockfish 4,013 3,234 — 7,247

Rosy rockfish 275 5 — 280

Rougheye rockfish 220 591 3 814

Sablefish 5,424 11,527 12,739 29,690

Sand sole 252 — — 252

Sharpchin rockfish 3,418 4,184 — 7,602

Shortbelly rockfish 7,077 3,450 — 10,527

Shortraker rockfish — 13 6 19

Shortspine thornyhead 507 27,812 10,775 39,094

Silvergray rockfish 58 95 — 153

Soupfin shark 5 — — 5

Southern rock sole 942 12 — 954

Speckled rockfish 123 — — 123

Spiny dogfish 14,709 4,760 5 19,474

Splitnose rockfish 2,565 23,636 1 26,202

Spotted ratfish 11,726 6,299 38 18,063

Squarespot rockfish 1,426 1 — 1,427

Starry flounder 166 — — 166

Starry rockfish 17 — — 17

Stripetail rockfish 12,425 10,463 — 22,888

Swordspine rockfish 1,343 299 — 1,642

Tiger rockfish 9 — — 9

Tree rockfish 1 — — 1

Vermilion rockfish 357 6 — 363

Widow rockfish 744 141 — 885

Yelloweye rockfish 198 43 — 241

Yellowmouth rockfish 7 157 — 164

Yellowtail rockfish 4,455 267 — 4,722
280

Table 11. Weight-length relationships by year for the 2003–2008 surveys using a nonlinear least squares fit for the equation: Weight = a ×

Lengthb (weight in kg; length in cm).

Species and 

year

Weight-length coeffcients Species and 

year

Weight-length coeffcients

a b r

2 Number a b r2 Number

Arrowtooth flounder Blackgill rockfish

2003 2.19E-06 3.39 0.98 921 2003 1.63E-05 2.99 0.98 118

2004 2.44E-06 3.36 0.98 567 2004 9.32E-06 3.14 0.99 182

2005 3.78E-06 3.24 0.98 875 2005 1.57E-05 3.00 0.99 181

2006 4.38E-06 3.20 0.98 739 2006 1.19E-05 3.07 0.99 223

2007 5.99E-06 3.11 0.98 905 2007 1.49E-05 3.00 0.97 189

2008 6.25E-06 3.10 0.99 895 2008 1.28E-05 3.05 0.99 299

Aurora rockfish Bocaccio

2003 9.67E-06 3.15 0.96 533 2003 6.66E-06 3.15 0.99 120

2004 8.49E-06 3.19 0.98 365 2004 5.96E-06 3.17 0.99 226

2005 9.23E-06 3.16 0.98 574 2005 8.39E-06 3.09 0.99 208

2006 1.05E-05 3.11 0.97 602 2006 7.94E-06 3.09 0.99 160

2007 1.06E-05 3.11 0.97 575 2007 6.16E-06 3.16 0.99 98

2008 9.31E-06 3.16 0.97 745 2008 1.04E-05 3.03 1.00 86

Bank rockfish Canary rockfish

2004 6.76E-06 3.22 0.97 77 2003 8.45E-06 3.18 0.99 264

2005 9.11E-06 3.13 0.97 50 2004 8.97E-06 3.17 0.99 291

2006 6.74E-06 3.21 0.99 70 2005 1.03E-05 3.14 0.99 278

2007 7.21E-06 3.21 0.98 54 2006 1.27E-05 3.08 0.98 272

2008 6.03E-06 3.25 0.99 33 2007 1.06E-05 3.11 0.99 502

2008 1.26E-05 3.07 0.99 470

Chilipepper Dover sole

2003 6.01E-06 3.23 0.97 693 2003 2.24E-06 3.41 0.97 2,989

2004 6.09E-06 3.23 0.99 856 2004 2.40E-06 3.39 0.97 2,368

2005 9.02E-06 3.12 0.99 898 2005 2.76E-06 3.36 0.98 2,588

2006 8.34E-06 3.14 0.99 643 2006 2.34E-06 3.39 0.97 2,242

2007 8.78E-06 3.13 0.99 607 2007 2.49E-06 3.38 0.97 2,193

2008 9.91E-06 3.09 0.99 685 2008 3.18E-06 3.31 0.98 1,092
281

Table 11 continued. Weight-length relationships by year for the 2003–2008 surveys using a nonlinear least squares fit for the equation: Weight = 

a × Lengthb (weight in kg; length in cm).

Species and 

year

Weight-length coeffcients Species and 

year

Weight-length coeffcients

a b r

2 Number a b r2 Number

Cowcod English sole

2003 8.16E-06 3.19 0.99 13 2003 7.04E-06 3.07 0.95 1,863

2004 6.68E-06 3.26 0.99 70 2004 7.28E-06 3.06 0.95 940

2005 8.37E-06 3.19 0.99 32 2005 1.01E-05 2.96 0.96 1,363

2006 5.19E-06 3.30 0.98 25 2006 9.11E-06 2.98 0.97 1,017

2007 9.63E-06 3.12 0.99 25 2007 8.14E-06 3.02 0.96 1,008

2008 8.34E-06 3.17 0.99 18 2008 8.68E-06 3.01 0.96 856

Darkblotched rockfish Greenstriped rockfish

2003 9.50E-06 3.18 0.97 679 2003 6.38E-06 3.20 0.94 383

2004 1.22E-05 3.11 0.98 565 2004 6.34E-06 3.21 0.97 484

2005 1.16E-05 3.12 0.99 799 2005 1.09E-05 3.06 0.97 735

2006 1.17E-05 3.11 0.98 944 2006 8.35E-06 3.13 0.97 707

2007 1.04E-05 3.14 0.98 983 2007 7.76E-06 3.15 0.97 584

2008 1.36E-05 3.07 0.99 814 2008 1.05E-05 3.06 0.97 561

Lingcod Pacific hake

2003 1.84E-06 3.39 0.99 866 2003 8.24E-06 2.93 0.97 1,152

2004 1.92E-06 3.39 0.99 860 2005 5.90E-06 3.02 0.98 416

2005 2.44E-06 3.32 0.99 970 2007 4.52E-06 3.08 0.99 1,816

2006 2.41E-06 3.32 0.99 743 2008 5.16E-06 3.06 0.99 1,227

2007 3.17E-06 3.25 0.99 559

2008 3.00E-06 3.26 0.99 897

Longspine thornyhead Pacific ocean perch

2003 1.04E-05 3.03 0.89 956 2003 5.44E-06 3.28 0.97 368

2004 1.90E-05 2.84 0.94 792 2004 1.71E-05 2.94 0.98 163

2005 1.75E-05 2.88 0.95 1,145 2005 1.19E-05 3.05 0.98 264

2006 1.23E-05 2.97 0.96 954 2006 7.66E-06 3.18 0.99 230

2007 4.53E-06 3.30 0.94 943 2007 9.97E-06 3.10 0.99 467

2008 4.96E-06 3.28 0.96 1,009 2008 9.67E-06 3.10 0.99 381
282

Table 11 continued. Weight-length relationships by year for the 2003–2008 surveys using a nonlinear least squares fit for the equation: Weight = 

a × Lengthb (weight in kg; length in cm).

Species and 

year

Weight-length coeffcients Species and 

year

Weight-length coeffcients

a b r

2 Number a b r2 Number

Pacific grenadier Pacific sanddab

2003 0.00030 2.61 0.92 836 2003 4.71E-06 3.23 0.92 872

2004 0.00025 2.69 0.96 380 2004 5.34E-06 3.19 0.92 1,512

2005 0.00024 2.73 0.96 545 2005 9.39E-06 3.01 0.94 1,145

2006 0.00018 2.82 0.97 603 2006 4.54E-06 3.23 0.94 856

2007 0.00014 2.90 0.97 640 2007 5.38E-06 3.19 0.95 818

2008 0.00019 2.81 0.97 569 2008 4.54E-06 3.25 0.94 794

Petrale sole Rex sole

2003 1.60E-06 3.55 0.97 1,583 2006 2.33E-06 3.28 0.92 1,699

2004 1.87E-06 3.50 0.97 1,821 2007 2.66E-06 3.24 0.93 823

2005 2.49E-06 3.42 0.98 1,970 2008 2.88E-06 3.23 0.94 773

2006 2.14E-06 3.45 0.98 1,079

2007 2.29E-06 3.44 0.98 1,043

2008 3.18E-06 3.35 0.98 1,047

Redbanded rockfish Rosethorn rockfish

2003 7.88E-06 3.22 0.99 189 2003 4.21E-06 3.38 0.96 425

2004 4.65E-06 3.38 0.99 128 2004 8.73E-06 3.14 0.92 315

2005 7.26E-06 3.24 0.99 173 2005 1.51E-05 2.97 0.95 323

2006 9.42E-06 3.15 0.99 144 2006 6.21E-06 3.24 0.96 511

2007 9.19E-06 3.16 0.99 169 2007 8.16E-06 3.15 0.96 502

2008 1.15E-05 3.10 0.99 149 2008 1.01E-05 3.10 0.96 386

Redstripe rockfish Rougheye rockfish

2003 8.75E-06 3.10 0.98 201 2003 7.09E-06 3.22 0.99 52

2004 1.61E-05 2.94 0.96 167 2004 4.52E-06 3.31 0.99 63

2005 4.07E-06 3.35 0.98 113 2005 1.01E-05 3.11 0.99 140

2006 3.38E-06 3.41 0.98 136 2006 1.09E-05 3.09 0.99 82

2007 5.86E-06 3.23 0.95 96 2007 7.12E-06 3.20 0.99 104

2008 2.30E-05 2.83 0.99 156 2008 1.00E-05 3.12 0.98 129
283

Table 11 continued. Weight-length relationships by year for the 2003–2008 surveys using a nonlinear least squares fit for the equation: Weight = 

a × Lengthb (weight in kg; length in cm).

Species and 

year

Weight-length coeffcients Species and 

year

Weight-length coeffcients

a b r

2 Number a b r2 Number

Sablefish Shortspine thornyhead

2003 5.30E-06 3.15 0.93 2,367 2003 4.79E-06 3.26 0.98 1,249

2004 3.02E-06 3.30 0.98 2,080 2004 5.35E-06 3.23 0.99 1,673

2005 3.12E-06 3.29 0.98 2,963 2005 6.17E-06 3.19 0.99 1,425

2006 3.50E-06 3.26 0.97 2,702 2006 6.98E-06 3.14 0.98 1,272

2007 4.08E-06 3.22 0.96 2,131 2007 4.68E-06 3.26 0.99 1,293

2008 4.54E-06 3.20 0.99 2,224 2008 4.73E-06 3.27 0.99 1,337

Sharpchin rockfish Silvergray rockfish

2003 1.18E-05 3.05 0.90 506 2003 6.76E-06 3.18 0.99 32

2004 1.67E-05 2.94 0.94 168 2004 2.09E-05 2.89 0.96 19

2005 8.19E-06 3.16 0.97 157 2005 4.39E-05 2.70 0.84 46

2006 8.53E-06 3.14 0.96 252 2006 4.79E-06 3.27 0.98 7

2007 8.42E-06 3.15 0.95 218 2007 1.12E-05 3.04 0.99 22

2008 9.98E-06 3.10 0.95 387 2008 6.12E-06 3.18 0.92 28

Shortbelly rockfish Spiny dogfish

2003 1.82E-06 3.55 0.89 540 2003 2.50E-06 3.11 0.98 689

2004 4.57E-06 3.27 0.92 477 2004 2.87E-06 3.08 0.99 490

2005 7.30E-06 3.13 0.95 585 2005 3.83E-06 3.02 0.98 953

2006 3.33E-06 3.38 0.93 384 2006 3.33E-06 3.05 0.98 818

2007 8.17E-06 3.09 0.93 331 2007 2.90E-06 3.09 0.98 788

2008 3.18E-06 3.40 0.96 343 2008 3.50E-06 3.04 0.98 901

Splitnose rockfish Yellowmouth rockfish

2003 1.05E-05 3.13 0.93 888 2003 5.72E-06 3.27 0.99 33

2004 1.95E-05 2.93 0.96 415 2005 7.83E-06 3.20 0.99 23

2005 2.57E-05 2.85 0.96 543

2006 2.42E-05 2.86 0.97 632

2007 1.77E-05 2.97 0.97 778

2008 1.71E-05 2.98 0.97 825
284

Table 11 continued. Weight-length relationships by year for the 2003–2008 surveys using a nonlinear least squares fit for the equation: Weight = 

a × Lengthb (weight in kg; length in cm).

Species and 

year

Weight-length coeffcients Species and 

year

Weight-length coeffcients

a b r

2 Number a b r2 Number

Widow rockfish Yellowtail rockfish

2003 5.16E-06 3.29 0.93 12 2003 8.95E-06 3.14 0.99 218

2004 4.01E-06 3.37 0.99 62 2004 8.29E-06 3.16 0.99 171

2005 6.03E-06 3.24 0.99 83 2005 9.56E-06 3.13 0.99 381

2006 2.29E-05 2.88 0.98 87 2006 2.07E-05 2.92 0.95 192

2007 7.56E-05 2.56 0.89 81 2007 2.53E-05 2.86 0.94 276

2008 5.59E-06 3.27 0.99 32 2008 1.87E-05 2.94 0.98 387

Yelloweye rockfish

2003 6.56E-06 3.26 0.98 73

2004 3.87E-06 3.42 0.99 18

2005 1.04E-05 3.15 0.99 40

2006 5.93E-06 3.29 0.98 45

2007 3.48E-06 3.44 0.99 24

2008 6.43E-06 3.28 0.98 46

Table 12. Coefficients for linear regressions describing net width and height as a function of inverse 

scope for each fishing vessel, all years combined. Net Width = Bw + ax where x is inverse scope. 

Net Height = Bh + ax where x is inverse scope.

Vessel Bw a Bh a

FV Excalibur –474.22 14.86 4.59 188.58

FV Ms. Julie –404.34 14.21 4.43 215.15

FV Noah’s Ark –463.32 14.84 4.73 226.25

FV Raven –445.52 15.09 4.21 229.45

FV Blue Horizon –392.01 14.97 4.36 179.83

FV B.J. Thomas –623.04 15.56 4.30 396.28

FV Captain Jack –404.32 14.59 4.49 159.15
285

References

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Kamikawa, L. C. Hufnagle Jr., and K. Piner. 2002. The 1999 Northwest Fisheries Science 

Center Pacific West Coast upper continental slope trawl survey of groundfish resources off 

Washington, Oregon, and California: Estimates of distribution, abundance, and length 

composition. U.S. Dept. Commer., NOAA Tech. Memo. NMFS-NWFSC-55.

Keller, A. A., E. L. Fruh, K. L. Bosley, D. J. Kamikawa, J. R. Wallace, B. H. Horness, V. H. Simon, and 

V. J. Tuttle. 2006a. The 2001 West Coast upper continental slope trawl survey of groundfish 

resources off Washington, Oregon, and California: Estimates of distribution, abundance, and 

length composition. U.S. Dept. Commer., NOAA Tech. Memo. NMFS-NWFSC-72.

Keller A. A., B. H. Horness, E. L. Fruh, V. H. Simon, V. J. Tuttle, K. L. Bosley, J. C. Buchanan, D. J. 

Kamikawa, and J. R. Wallace. 2008. The 2005 U.S. West Coast bottom trawl survey of 

gjroundfish resources of Washington, Oregon, and California: Estimates of distribution, 

abundance, and length composition. U.S. Dept. Commer., NOAA Tech. Memo. NMFS-NWFSC93.

Keller, A. A., B. H. Horness, V. H. Simon, V. J. Tuttle, J. R. Wallace, E. L. Fruh, K. L. Bosley, D. J. 

Kamikawa, and J. C. Buchanan. 2007b. The 2004 West Coast upper continental slope trawl 

survey of groundfish resources off Washington, Oregon, and California: Estimates of distribution, 

abundance, and length composition. U.S. Dept. Commer., NOAA Tech. Memo. NMFS-NWFSC87.

Keller, A. A., B. H. Horness, V. J. Tuttle, J. R. Wallace, V. H. Simon, E. L. Fruh, K. L. Bosley, and D. J.

Kamikawa. 2006b. The 2002 West Coast upper continental slope trawl survey of groundfish 

resources off Washington, Oregon, and California: Estimates of distribution, abundance, and 

length composition. U.S. Dept. Commer., NOAA Tech. Memo. NMFS-NWFSC-75.

Keller, A. A., V. H. Simon, B. H. Horness, J. R. Wallace, V. J. Tuttle, E. L. Fruh, K. L. Bosley, D. J. 

Kamikawa, and J. C. Buchanan. 2007a. The 2003 West Coast upper continental slope trawl 

survey of groundfish resources off Washington, Oregon, and California: Estimates of distribution, 

abundance, and length composition. U.S. Dept. Commer., NOAA Tech. Memo. NMFS-NWFSC86.

Keller, A. A., T. L. Wick, E. L. Fruh, K. L. Bosley, D. J. Kamikawa, J. R. Wallace, and B. H. Horness.

2005. The 2000 West Coast upper continental slope trawl survey of groundfish resources off 

Washington, Oregon, and California: Estimates of distribution, abundance, and length 

composition. U.S. Dept. Commer., NOAA Tech. Memo. NMFS-NWFSC-70.

Lauth, R. R. 2001. The 2000 Pacific West Coast upper continental slope trawl survey of groundfish 

resources off Washington, Oregon, and California: Estimates of distribution, abundance, and 

composition. U.S. Dept. Commer., NOAA Tech. Memo. NMFS-AFSC-120.

Methot, R. D., J. R. Wallace, and C. W. West. 2000. Introducing a new trawl survey for U.S. West Coast 

slope groundfish. Presented at ICES Annual Science Conference, Brugge, Belgium, September 
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2000. (Available from R. Methot, NWFSC, Fishery Resource Analysis and Monitoring Division, 

2725 Montlake Blvd. E., Seattle, WA 98112.)

Stauffer, G. 2004. NOAA protocols for groundfish bottom trawl surveys of the nation’s fishery 

resources. U.S. Dept. Commer., NOAA Tech. Memo. NMFS-F/SPO-65.

Turk, T. A., T. L. Builder, C. W. West, D. J. Kamikawa, J. R. Wallace, R. D. Methot, A. R. Bailey, K. L. 

Bosley, A. J. Cook, E. L. Fruh, B. H. Horness, K. Piner, H. R. Sanborn, and W. W. Wakefield.

2001. The 1998 Northwest Fisheries Science Center Pacific West Coast upper continental slope 

trawl survey of groundfish resources off Washington, Oregon, and California: Estimates of 

distribution, abundance, and length composition. U.S. Dept. Commer., NOAA Tech. Memo. 

NMFS-NWFSC-50.

Wallace, J. R., and C. W. West. 2006. Measurements of distance fished during the trawl retrieval period. 

Fish. Res. 77:285-292.
287

Appendix A: Primary and Alternative Station 

Selection for the Survey Cell Pool

The survey pool of cells were picked to be:

1. Within U.S. waters.

2. On the Pacific Ocean only, no straits or inlets.

3. Between 30 and 700 fathoms (fm) deep.

4. Only within the natural fathom contours, that is, seamounts outside the 700 fm 

contour line were not included. In the California Bight, a small extremely western 

seamount was excluded, but all other larger areas less than or equal to 700 fm were 

included.

5. Outside the Cowcod Exclusion Area.

6. Outside the 3-mile limit of the Channel Island Marine Protected Area.

7. Only those cells with greater than 50% of the cell area within the political and 

geographical borders defined above.

For stratum apportionment, each cell needs a unique INPFC area and depth designation. 

For area, cells spanning more than one area were assigned the INPFC area that was greater by 

surface area. Note that cells below the classic Conception area boundary of 32°30′N and above 

the U.S.-Mexico exclusive economic zone are assigned the Conception Area designation.

Likewise, each cell was given a single classifying depth designation from the following 

list:

• <30 fm

• 30–100 fm

• 100–300 fm

• 300–700 fm

• 700–1,000 fm

• >1,000 fm

Cells spanning two depth zones were assigned the one with the greatest surface area. 

Cells spanning three depth zones were assigned the one with the largest area, regardless of 

whether it was greater than 50% of the total. However, cells on any border still need to first 

qualify using item 7 above. For example, if a cell has 25% of its area greater than 1,000 m, 35% 

of its area 700–1,000 m, and 40% of its area 300–700 m, then the cell would not be included in 

the pool. For even though the 300–700 depth stratum is the largest, 60% of the area is outside of 
288

700 fm. Note that in this example, if a new survey includes the 700–1,000 fm depth strata, then 

the majority of the cell would be within the survey boundaries (75%), but the majority depth 

strata would be the 300–700 strata. So adding the 700–1,000 strata transforms a previously 

unselected cell to a 300–700 cell in the new survey. The percent of cells that span three depth 

strata, but have no depth strata greater than 50%, is two-tenths of 1% of the survey pool of cells.

Primary Station Selection

For 2003, 720 cells (one tow per cell) were apportioned to the strata by multiplying 720 

times the strata proportions found above. Rounding to the nearest whole number caused the 

grand total to miss the mark, so unary adjustments were made to the strata numbers to obtain a 

grand total of 720. The primary stations were then picked from the survey cell pool by strata, 

using a pseudorandom number generator. These 720 selected primary station cells were assigned 

a vessel number of 1–4, using a simple sequence (1,2,3,4, 1,2,3,4, ...)

Alternative Station Selection

For each primary station, two alternative stations were chosen as follows.

By strata, a secondary random selection of the nonprimary cells was performed. Using a 

great circle distance measurement, the closest site of the available alternative site pool was 

matched to a primary site within a stratum. These were then removed from the secondary pool 

for that stratum and the process was repeated for the second alternative site. This was done for 

each of the strata, in turn. The number of cells selected for this secondary pick is set greater than 

the first and hence more densely distributed geographically than the primary cells. This density 

was selected by strata to achieve a good balance between travel distance between sites (distance 

from primary to first alternative and first alternative to second alternative) and to ensure to a 20–

30% chance that the alternative sites would not be positioned directly adjacent to the primary 

site.

Where nearby alternative sites are at a premium, such as in the shallow Conception Area, 

a few alternative sites are more than 20 nautical miles away from their primary station. In the 

event this proves to be a problem, the following rule was developed: If the primary station is 

deemed untrawlable and the secondary station is greater than 2 hours “out of your way” (in the 

wrong direction, that is, in relation to your next primary station), then it is at the field party 

chief’s discretion to postpone or skip that secondary station.

Search Times

All sites, whether primary or alternative, will be searched for a towable location for up to 

a maximum of 1 hour each. If no towable location is found after 3 hours, then the vessel will 

move on to the next primary location on its list.
289

Appendix B: Species Frequency of Occurrence, 

Depth Range, and Latitudinal Range

Appendix B consists of two tables. Table B-1 lists frequency of occurrence, depth range, 

and latitudinal range for fish species and Table B-2 lists them for invertebrates.
290

Table B-1. Frequency of occurrence, depth range, and latitudinal range for fish species, grouped by family (or higher taxonomic classification), 

encountered during the 2003–2008 surveys (unident. = unidentified).

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal degrees) Depth (m)

South North Min. Max.

Myxinidae

Myxinidae Hagfish unident. 357 32.1 48.1 60 1,297

Eptatretus sp. 1 47.6 47.6 986 986

E. deani Black hagfish 332 32.3 48.3 52 1,271

E. stouti Pacific hagfish 128 32.6 48.2 66 1,155

Petromyzontidae Lamprey unident. 3 34.3 47.6 99 920

Lampetra tridentata Pacific lamprey 3 41.2 44.5 171 308

Hexanchiformes 

Hexanchus griseus Sixgill shark 11 33.6 45.7 79 406

Squalidae Dogfish sharks 1 33.6 33.6 1,188 1,188

Squalus acanthias Spiny dogfish 1,386 32.6 48.5 36 1,143

Somniosus pacificus Pacific sleeper shark 17 33.0 47.3 463 1,166

Scyliorhinidae

Scyliorhinidae Cat shark unident. 11 33.4 44.8 335 1,209

Apristurus brunneus Brown cat shark 1,333 32.1 48.3 82 1,241

A. brunneus egg case Cat shark egg case 77 32.7 48.2 74 1,147

A. kampae Longnose cat shark 78 32.2 47.6 230 1,268

A. sp. 2 34.3 34.3 512 571

Cephaloscyllium ventriosum Swell shark 21 33.0 34.7 59 117

Parmaturus xaniurus Filetail cat shark 363 32.7 37.3 113 1,224

Triakididae Smoothhounds 40 32.6 38.1 52 208

Galeorhinus galeus Soupfin shark 10 33.7 41.5 69 175

Mustelus californicus Gray smoothhound 27 32.7 38.8 27 247

M. henlei Brown smoothhound 52 32.6 38.4 35 243

Carcharhinidae

Prionace glauca Blue shark 1 40.6 40.6 173 173

Squatinidae

Squatina californica Pacific angel shark 20 32.8 35.1 27 184

Dalatiidae

Centroscyllium nigrum Combtooth dogfish 15 32.6 33.2 871 1,212
291

Table B-1 continued. Frequency of occurrence, depth range, and latitudinal range for fish species, grouped by family (or higher taxonomic 

classification), encountered during the 2003–2008 surveys (unident. = unidentified).

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal degrees) Depth (m)

South North Min. Max.

Rhinobatidae Guitarfishes 2 34.3 34.3 85 174

Rhionbatos productus Shovelnose guitarfish 1 34.0 34.0 27 27

Zapteryx exasperata Banded guitarfish 1 33.8 33.8 60 60

Rajidae unident. 2 33.8 48.4 313 648

Bathyraja abyssicola Deepsea skate 19 32.3 47.5 780 1,428

B. aleutica Aleutian skate 12 36.2 47.3 206 1,241

B. kincaidii Bering skate 1,258 32.4 48.5 52 1,173

B. kincaidii egg case Bering skate egg case 1 44.9 44.9 1,007 1,007

Bathyraja sp. 17 32.6 47.7 209 1,206

Bathyraja sp. egg case 6 44.9 47.7 69 1,011

Bathyraja trachura Roughtail skate 500 32.2 48.2 107 1,428

Bathyraja trachura egg case Roughtail skate egg case 2 42.6 46.0 1,186 1,243

Raja binoculata Big skate 489 32.6 48.4 36 302

R. binoculata egg case Big skate egg case 66 34.1 48.2 54 1,169

R. inornata California skate 432 32.6 48.0 24 792

R. rhina Longnose skate 2,239 32.4 48.5 43 1,162

R. sp. egg case Raja sp. egg case 46 34.3 48.3 59 1,011

R. stellulata Starry skate 77 33.3 48.1 54 982

Rajiformes egg case Skate egg case unident. 306 32.6 48.5 35 1,214

Torpedinidae Electric rays 8 33.5 38.1 69 379

Torpedo californica Pacific electric ray 237 32.6 48.4 27 1,079

Dasyatidae Whiptail Stingrays 1 34.6 34.6 59 59

Myliobatidae Eagle rays 2 32.8 39.1 359 488

Myliobatis californicus Bat ray 25 32.6 34.4 27 176

Chimaeridae

Hydrolagus colliei Spotted ratfish 1,983 32.6 48.5 44 1,241

Nettastomatidae Duckbilled eels 1 32.7 32.7 1,096 1,096

Facciolella gilbertii Dogface witch-eel 11 32.7 34.4 371 507

Venefica sp. Whipsnout eels 5 32.7 46.3 916 1,148

Acipenseridae
292

Table B-1 continued. Frequency of occurrence, depth range, and latitudinal range for fish species, grouped by family (or higher taxonomic 

classification), encountered during the 2003–2008 surveys (unident. = unidentified).

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal degrees) Depth (m)

South North Min. Max.

Acipenser medirostris Green sturgeon 1 44.7 44.7 53 53

Anguilliformes Eel unident. 1 34.6 34.6 743 743

Nemichthyidae Snipe eel unident. 10 32.7 35.8 196 972

Avocettina infans Blackline snipe eel 46 32.4 47.9 71 1,186

Nemichthys larseni Pale snipe eel 7 32.9 47.4 366 935

N. scolopaceus Slender snipe eel 13 32.7 33.9 482 1,241

Serrivomeridae Sawtooth eels 23 32.3 40.6 290 1,246

Serrivomer jesperseni Crossthroat snipe eel 2 36.1 36.2 1,139 1,173

S. sector Sawtooth eel 21 32.7 47.7 516 1,206

Saccopharyngidae Whiptail gulpers 5 32.6 33.2 1,075 1,160

Clupeidae 

Alosa sapidissima American shad 418 34.0 48.4 52 440

Clupea pallasi Pacific herring 281 34.7 48.4 47 723

Sardinops sagax Pacific sardine 82 32.8 48.4 35 982

Salmonidae

Oncorhynchus kisutch Coho salmon 2 38.3 38.5 89 101

O. tshawytscha Chinook salmon 52 34.9 48.3 58 171

Osmeridae Smelt unident. 37 34.2 48.3 57 579

Allosmerus elongatus Whitebait smelt 60 34.4 47.9 53 170

Hypomesus pretiosus Surf smelt 1 41.5 41.5 113 113

Osmerus mordax Rainbow smelt 6 45.8 48.4 61 156

Spirinchus starksi Night smelt 21 34.0 48.1 53 594

S. thaleichthys Longfin smelt 6 33.9 46.6 55 64

Thaleichthys pacificus Eulachon 165 34.0 48.4 51 374

Engraulidae Anchovies 14 33.4 46.1 53 276

Engraulis mordax Northern anchovy 166 32.7 47.8 35 806

Argentinidae Argentine unident. 7 32.6 42.1 96 847

Nansenia candida Bluethroat argentine 4 40.1 48.0 437 968

Argentina sialis Pacific argentine 127 32.6 42.6 60 907

Bathylagidae Deepsea smelt unident. 657 32.2 48.2 247 1,341
293

Table B-1 continued. Frequency of occurrence, depth range, and latitudinal range for fish species, grouped by family (or higher taxonomic 

classification), encountered during the 2003–2008 surveys (unident. = unidentified).

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal degrees) Depth (m)

South North Min. Max.

Bathylagus milleri Robust blacksmelt 14 32.9 46.9 562 1,215

B. pacificus Pacific blacksmelt 5 33.3 46.5 867 1,215

B. sp. Blacksmelt unident. 84 32.6 48.0 562 1,428

Leuroglossus schmidti Northern smoothtongue 18 33.3 42.1 397 898

Leuroglossus sp. 5 33.4 35.2 399 554

L. stilbius California smoothtongue 91 32.5 43.4 390 1,174

Opisthoproctidae Spookfish unident. 5 32.4 41.7 431 885

Bathylychnops exilis Javelin spookfish 1 32.7 32.7 452 452

Macropinna microstoma Barreleye 22 32.6 47.7 279 1,201

Gonostomatidae Bristlemouth unident. 13 32.4 47.9 566 1,186

Gonostoma sp. 1 36.7 36.7 805 805

Sternoptychidae Hatchetfish unident. 21 32.6 43.3 185 1,148

Argyropelecus affinis Slender hatchetfish 64 32.1 43.6 303 1,212

A. lychnus Tropical hatchetfish 1 33.0 33.0 968 968

A. sp. 20 32.6 43.8 190 1,205

Sternoptyx diaphana Longspine hatchetfish 20 32.5 41.7 328 1,230

S. sp. 10 32.9 47.6 543 1,079

Stomiiformes Dragonfish unident. 1 33.2 33.2 893 893

Stomiidae Barbled dragonfishes 5 32.7 47.6 83 1,184

Stomias atriventer Blackbelly dragonfish 92 32.6 48.0 177 1,241

Chauliodontidae Viperfish unident. 20 32.9 47.9 207 1,197

Chauliodus macouni Pacific viperfish 454 32.5 48.2 106 1,297

Astronesthidae

Borostomias panamensis Panama snaggletooth 4 32.6 37.1 666 1,098

Malacosteidae Loosejaw unident. 1 32.9 32.9 1,096 1,096

Aristostomias scintillans Shining loosejaw 53 32.1 47.9 176 1,237

Idiacanthus antrostomus Pacific blackdragon 56 32.5 46.1 311 1,240

Melanostomiidae Scaleless dragonfish unident. 6 32.7 47.7 307 763

Bathophilus flemingi Highfin dragonfish 1 38.4 38.4 603 603

Tactostoma macropus Longfin dragonfish 301 32.8 48.2 156 1,341
294

Table B-1 continued. Frequency of occurrence, depth range, and latitudinal range for fish species, grouped by family (or higher taxonomic 

classification), encountered during the 2003–2008 surveys (unident. = unidentified).

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal degrees) Depth (m)

South North Min. Max.

Alepocephalidae Slickhead unident. 2 35.3 45.0 867 1,196

Alepocephalus tenebrosus California slickhead 846 32.2 48.2 477 1,279

Bajacalifornia burragei Sharpchin slickhead 12 32.6 40.0 805 1,169

B. erimoensis 2 33.0 33.0 814 1,212

Talismania bifurcata Threadfin slickhead 242 32.3 48.1 601 1,341

Platytroctidae Tubeshoulder unident. 7 32.7 40.1 885 1,029

Maulisia mauli 7 32.8 47.7 723 1,172

Maulisia sp. 1 35.3 35.3 934 934

Sagamichthys abei Shining tubeshoulder 30 32.3 47.6 381 1,197

Synodontidae Lizardfish unident. 4 32.7 34.3 79 85

Synodus lucioceps California lizardfish 61 32.6 35.5 24 122

Alepisauridae

Alepisaurus ferox Longnose lancetfish 1 32.8 32.8 1,041 1,041

Scopelarchidae

Benthalbella dentata Northern pearleye 7 35.8 47.8 710 1,205

Paralepididae

Arctozenus risso Ribbon barracudina 5 32.9 47.8 589 1,102

Lestidiops ringens Slender barracudina 5 33.4 44.9 71 805

Magnisudis atlantica Duckbill barracudina 4 40.2 47.6 520 808

Scopelosauridae

Scopelosaurus harryi Scaly paperbone 1 37.1 37.1 667 667

Myctophidae Lanternfish unident. 729 32.3 48.2 86 1,268

Diaphus theta California headlightfish 91 32.3 48.4 157 1,215

D. sp. 1 35.6 35.6 1,002 1,002

Lampanyctus ritteri Broadfin lanternfish 9 35.2 47.9 397 903

L. sp. 523 32.4 48.4 115 1,428

Stenobrachius leucopsarus Northern lampfish 33 32.4 46.4 386 1,297

Symbolophorus californiensis California lanternfish 15 33.3 47.7 490 1,197

Tarletonbeania crenularis Blue lanternfish 30 34.5 47.7 119 1,095

T. sp. 9 34.7 47.7 151 1,068
295

Table B-1 continued. Frequency of occurrence, depth range, and latitudinal range for fish species, grouped by family (or higher taxonomic 

classification), encountered during the 2003–2008 surveys (unident. = unidentified, YOY = young of year).

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal degrees) Depth (m)

South North Min. Max.

Neoscopelidae Blackchin unident. 1 45.9 45.9 1,132 1,132

Scopelengys tristis Blackchin 6 32.6 42.0 723 972

Gadidae

Gadus macrocephalus Pacific cod 216 39.2 48.5 53 285

Microgadus proximus Pacific tomcod 133 35.1 48.4 53 117

Theragra chalcogramma Walleye pollock 23 47.1 48.5 73 270

Merlucciidae

Merluccius productus Pacific hake 2,180 32.1 48.4 52 1,213

M. productus YOY Pacific hake YOY 103 32.9 47.6 66 1,015

Moridae Codlings unident. 2 32.7 47.3 498 713

Antimora microlepis Pacific flatnose 783 32.2 48.2 262 1,428

Halargyreus johnsoni Slender codling 2 41.8 43.5 753 1,130

Physiculus rastrelliger Hundred fathom mora 33 32.7 35.1 142 498

Melanonidae

Melanonus zugmayeri Arrowtail 9 32.7 42.0 766 1,124

Macrouridae Grenadier unident. 2 34.4 46.1 388 565

Albatrossia pectoralis Giant grenadier 685 32.3 48.2 443 1,428

Coelorinchus scaphopsis Shoulderspot grenadier 2 34.5 36.3 166 343

Coryphaenoides acrolepis Pacific grenadier 797 32.2 48.2 313 1,428

C. cinereus Popeye grenadier 52 33.4 48.1 395 1,297

C. filifer Filamented grenadier 2 40.0 40.5 1,064 1,196

Nezumia liolepis Smooth grenadier 172 32.1 46.2 390 1,246

N. stelgidolepis California grenadier 200 32.1 45.0 377 1,201

Ophidiidae Cusk-eel unident. 1 48.4 48.4 117 117

Chilara taylori Spotted cusk-eel 121 32.8 47.9 27 366

Dicrolene filamentosa Threadfin cusk-eel 2 32.9 37.1 879 1,096

Lamprogrammus niger Paperbone cusk-eel 10 32.6 33.7 876 1,184

Bythitidae

Brosmophycis marginata Red brotula 2 32.6 41.2 138 190

Cataetyx rubrirostris Rubynose brotula 40 32.7 41.8 322 874
296

Table B-1 continued. Frequency of occurrence, depth range, and latitudinal range for fish species, grouped by family (or higher taxonomic 

classification), encountered during the 2003–2008 surveys (unident. = unidentified).

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal degrees) Depth (m)

South North Min. Max.

C. sp. 1 34.3 34.3 402 402

Zoarcidae Eelpout unident. 9 32.7 45.9 104 1,132

Bothrocara brunneum Twoline eelpout 700 32.2 48.2 450 1,428

B. molle Soft eelpout 6 34.5 46.6 803 1,341

Lycenchelys camchatica Kamchatka eelpout 25 35.0 47.7 524 1,238

L. crotalinus Snakehead eelpout 601 32.7 48.2 219 1,297

Lycodapus dermatinus Looseskin eelpout 1 41.0 41.0 646 646

L. endemoscotus Deepwater eelpout 29 34.3 47.8 386 1,197

L. fierasfer Blackmouth eelpout 16 32.7 46.6 743 1,200

L. mandibularis Pallid eelpout 24 32.6 48.1 399 1,173

L. sp. 5 33.2 39.0 371 1,111

Lycodema barbatum Bearded eelpout 3 34.2 40.6 144 290

Lycodes brevipes Shortfin eelpout 1 41.5 41.5 171 171

L. cortezianus Bigfin eelpout 1,086 32.7 48.2 57 1,162

L. diapterus Black eelpout 686 32.6 48.3 82 1,162

L. pacificus Blackbelly eelpout 664 32.6 48.4 52 729

L. palearis Wattled eelpout 2 47.6 47.9 111 187

Maynea californica Persimmon eelpout 2 34.3 34.4 70 349

Melanostigma pammelas Midwater eelpout 3 33.9 41.0 713 806

Batrachoididae Midshipmen 1 35.2 35.2 252 252

Porichthys notatus Plainfin midshipman 500 32.7 47.8 24 464

Oneirodidae Dreamer unident. 16 32.8 45.4 691 1,188

Oneirodes thompsoni 1 32.4 32.4 1,040 1,040

Oneirodes sp. 12 32.6 47.8 528 1,268

Chaenophryne longiceps 1 45.7 45.7 84 84

Chaenophryne draco Smooth dreamer 6 32.9 47.3 766 1,186

Melanocetidae

Melanocetus johnsonii Common blackdevil 2 34.5 45.2 503 797

Ceratiidae

Cryptopsaras couesii Triplewart sea devil 1 33.4 33.4 844 844
297

Table B-1 continued. Frequency of occurrence, depth range, and latitudinal range for fish species, grouped by family (or higher taxonomic 

classification), encountered during the 2003–2008 surveys (unident. = unidentified).

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal degrees) Depth (m)

South North Min. Max.

Gigantactinidae

Gigantactis vanhoeffeni Whipnose 1 32.6 32.6 1,056 1,056

Trachipteridae

Trachipterus altivelis King-of-the-salmon 6 32.7 45.3 158 1,214

Melamphaidae Bigscale unident. 9 32.3 41.7 489 1,124

Melamphaes lugubris Highsnout bigscale 23 32.6 45.8 741 1,214

Poromitra crassiceps Crested bigscale 131 32.5 47.9 126 1,428

Anoplogastridae Fangtooths 3 33.2 43.0 848 1,049

Anoplogaster cornuta Fangtooth 83 32.4 48.2 641 1,246

Scorpaenidae Scorpionfish/rockfish unident. 3 33.0 34.0 27 117

Scorpaena guttata California scorpionfish 98 32.6 34.4 24 190

Sebastolobus alascanus Shortspine thornyhead 1,989 32.1 48.4 67 1,341

S. altivelis Longspine thornyhead 1,426 32.1 48.3 163 1,428

Sebastes aleutianus Rougheye rockfish 200 32.7 48.4 125 798

S. alutus Pacific ocean perch 250 40.2 48.4 87 715

S. auriculatus Brown rockfish 33 32.8 41.7 58 94

S. aurora Aurora rockfish 530 32.1 48.2 129 814

S. babcocki Redbanded rockfish 316 33.7 48.4 71 550

S. borealis Shortraker rockfish 9 44.5 48.2 301 555

S. brevispinis Silvergray rockfish 38 37.4 48.2 66 283

S. carnatus Gopher rockfish 5 32.7 35.5 58 76

S. caurinus Copper rockfish 46 32.8 40.5 36 408

S. chlorostictus Greenspotted rockfish 245 32.6 48.4 65 320

S. constellatus Starry rockfish 14 32.6 38.0 62 141

S. crameri Darkblotched rockfish 686 33.6 48.4 84 538

S. dalli Calico rockfish 57 32.6 37.6 52 146

S. diploproa Splitnose rockfish 844 32.3 48.4 68 559

S. elongatus Greenstriped rockfish 992 32.6 48.5 64 447

S. emphaeus Puget Sound rockfish 7 34.4 48.0 36 173

S. ensifer Swordspine Rockfish 26 32.6 34.2 106 308
298

Table B-1 continued. Frequency of occurrence, depth range, and latitudinal range for fish species, grouped by family (or higher taxonomic 

classification), encountered during the 2003–2008 surveys.

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal degrees) Depth (m)

South North Min. Max.

S. entomelas Widow rockfish 135 34.4 48.4 64 399

S. eos Pink rockfish 20 33.2 37.2 83 344

S. flavidus Yellowtail rockfish 248 33.8 48.4 36 231

S. gilli Bronzespotted rockfish 1 32.7 32.7 382 382

S. goodei Chilipepper 509 32.6 47.5 36 464

S. helvomaculatus Rosethorn rockfish 329 32.6 48.5 65 447

S. hopkinsi Squarespot rockfish 67 32.6 38.0 52 169

S. jordani Shortbelly rockfish 328 32.6 48.3 26 406

S. lentiginosus Freckled rockfish 8 32.6 33.8 60 103

S. levis Cowcod 87 32.6 42.3 74 308

S. macdonaldi Mexican rockfish 1 33.9 33.9 269 269

S. maliger Quillback rockfish 12 40.5 48.3 61 125

S. melanops Black rockfish 2 37.4 40.8 70 71

S. melanostictus Blackspotted rockfish 5 44.4 45.7 367 408

S. melanostomus Blackgill rockfish 195 32.3 47.9 133 595

S. miniatus Vermilion rockfish 57 32.6 38.0 58 478

S. mystinus Blue rockfish 5 34.4 37.4 64 132

S. nigrocinctus Tiger rockfish 6 38.2 48.3 114 178

S. ovalis Speckled rockfish 6 32.6 33.9 106 151

S. paucispinis Bocaccio 233 32.6 48.3 56 326

S. pinniger Canary rockfish 283 32.6 48.4 54 260

S. proriger Redstripe rockfish 105 33.5 48.4 66 271

S. reedi Yellowmouth rockfish 7 43.0 48.3 114 313

S. rosaceus Rosy rockfish 20 32.7 43.0 60 328

S. rosenblatti Greenblotched rockfish 69 32.6 38.3 63 413

S. ruberrimus Yelloweye rockfish 90 36.7 48.4 66 250

S. rubrivinctus Flag rockfish 63 32.6 36.9 55 431

S. rufus Bank rockfish 73 32.6 48.1 108 499

S. saxicola Stripetail rockfish 801 32.3 48.4 36 436

S. semicinctus Halfbanded rockfish 312 32.6 44.6 52 395
299

Table B-1 continued. Frequency of occurrence, depth range, and latitudinal range for fish species, grouped by family (or higher taxonomic 

classification), encountered during the 2003–2008 surveys (unident. = unidentified).

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal degrees) Depth (m)

South North Min. Max.

S. serranoides Olive rockfish 2 37.4 37.7 103 134

S. serriceps Tree rockfish 1 33.8 33.8 60 60

S. simulator Pinkrose rockfish 3 33.8 34.2 205 325

S. umbrosus Honeycomb rockfish 12 32.6 34.4 52 241

S. wilsoni Pygmy rockfish 81 32.6 48.4 64 268

S. zacentrus Sharpchin rockfish 236 33.7 48.5 86 404

S. sp. Rockfish unident. 33 32.7 48.1 54 512

S. sp. Rockfish unident. 3 33.3 43.9 114 240

Triglidae

Prionotus stephanophrys Lumptail sea robin 13 32.9 33.8 52 184

Bellator xenisma Splitnose Searobin 1 33.4 33.4 66 66

Anoplopoma fimbria Sablefish 2,549 32.1 48.5 26 1,428

Hexagrammidae

Ophiodon elongatus Lingcod 1,170 32.6 48.5 26 410

Hexagrammos lagocephalus Rock greenling 1 34.0 34.0 27 27

H. decagrammus Kelp greenling 59 34.4 48.4 60 149

Zaniolepididae

Zaniolepis latipinnis Longspine combfish 363 32.6 45.6 26 288

Z. frenata Shortspine combfish 144 32.6 38.2 59 411

Cottidae Sculpin unident. 15 33.3 48.1 73 1,015

Bolinia euryptera Broadfin sculpin 1 33.8 33.8 126 126

Chitonotus pugetensis Roughback sculpin 30 32.9 45.6 56 240

Clinocottus acuticeps Sharpnose sculpin 6 33.3 44.5 61 96

Dasycottus setiger Spinyhead sculpin 2 37.8 48.4 58 169

Enophrys bison Buffalo sculpin 8 33.3 48.2 56 194

E. taurina Bull sculpin 28 33.9 37.9 58 95

Hemilepidotus hemilepidotus Red Irish lord 10 42.4 48.4 70 151

H. spinosus Brown Irish lord 5 38.2 48.3 63 126

H. sp. Irish lord 2 37.4 45.7 65 122

Icelinus burchami Dusky sculpin 11 33.5 48.0 61 435
300

Table B-1 continued. Frequency of occurrence, depth range, and latitudinal range for fish species, grouped by family (or higher taxonomic 

classification), encountered during the 2003–2008 surveys (unident. = unidentified).

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal degrees) Depth (m)

South North Min. Max.

I. filamentosus Threadfin sculpin 347 32.9 48.5 54 482

I. fimbriatus Fringed sculpin 23 32.5 46.0 90 1,158

I. tenuis Spotfin sculpin 5 33.3 48.0 67 165

I. sp. Sculpin unident. 5 32.7 42.5 75 382

Jordania zonope Longfin sculpin 3 33.7 44.3 121 497

Leptocottus armatus Pacific staghorn sculpin 61 33.4 48.2 47 335

Malacocottus kincaidi Blackfin sculpin 8 45.1 47.9 293 437

Nautichthys oculofasciatus Sailfin sculpin 2 37.8 44.4 58 88

Paricelinus hopliticus Thornback sculpin 1 33.3 33.3 234 234

Psychrolutes phrictus Blob sculpin 14 39.1 47.8 988 1,341

Radulinus asprellus Slim sculpin 57 33.3 48.0 62 380

R. taylori Spinynose sculpin 1 38.6 38.6 212 212

Rhamphocottus richardsoni Grunt sculpin 5 33.7 43.1 61 150

Scorpaenichthys marmoratus Cabezon 2 34.0 43.1 27 61

Triglops macellus Roughspine sculpin 3 48.2 48.4 113 117

Zesticelus profundorum Flabby sculpin 7 34.2 47.7 941 1,186

Agonidae Poacher unident. 9 32.7 48.3 70 302

Agonopsis vulsa Northern spearnose poacher 19 34.4 48.2 61 260

Bathyagonus nigripinnis Blackfin poacher 221 35.2 48.3 116 1,008

B. pentacanthus Bigeye poacher 55 32.7 48.3 53 611

Chesnonia verrucosa Warty poacher 14 37.9 47.9 53 89

Odontopyxis trispinosa Pygmy poacher 1 34.4 34.4 70 70

Podothecus acipenserinus Sturgeon poacher 2 48.1 48.2 65 88

Xeneretmus latifrons Blacktip poacher 114 32.7 48.2 83 1,064

X. leiops Smootheye poacher 7 34.8 48.0 149 399

Liparidinae Snailfish unident. 19 32.6 47.9 63 1,240

Careproctus colletti Alaska snailfish 2 35.6 42.5 331 946

C. cypselurus Blackfin snailfish 113 33.4 47.8 317 1,341

C. gilberti Smalldisk snailfish 38 34.8 44.9 248 672

C. melanurus Blacktail snailfish 770 32.3 48.2 163 1,215
301

Table B-1 continued. Frequency of occurrence, depth range, and latitudinal range for fish species, grouped by family (or higher taxonomic 

classification), encountered during the 2003–2008 surveys.

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal degrees) Depth (m)

South North Min. Max.

C. sp. 7 32.5 47.8 737 1,115

Elassodiscus caudatus Humpback snailfish 19 36.8 47.8 399 879

E. tremebundus Blacklip snailfish 1 33.4 33.4 611 611

Liparidae n. gen. (Orr) Hardhead snailfish 1 33.9 33.9 1,224 1,224

Liparis fucensis Slipskin snailfish 1 42.1 42.1 77 77

L. pulchellus Showy snailfish 1 41.8 41.8 88 88

Nectoliparis pelagicus Tadpole snailfish 1 39.2 39.2 476 476

Paraliparis cephalus Swellhead snailfish 77 32.6 47.8 504 1,285

P. dactylosus Red snailfish 48 32.6 47.6 422 1,162

P. pectoralis Broadfin snailfish 14 32.6 47.7 893 1,241

P. rosaceus Rosy snailfish 8 32.4 46.2 972 1,215

P. sp. 2 32.9 35.6 302 1,029

Rhinoliparis attenuatus Slim snailfish 6 32.8 40.8 546 1,139

R. barbulifer Longnose snailfish 13 32.6 40.1 795 1,184

R. sp. 2 32.7 35.3 867 961

Percichthyidae

Howella sherborni Sherborn’s pelagic bass 1 45.1 45.1 331 331

Chiasmodontidae Swallowers 3 32.9 33.2 848 933

Chiasmodon niger Black swallower 24 32.8 42.3 379 1,241

Kali indica Shortnose swallower 1 32.9 32.9 1,028 1,028

K. normani Needletooth swallower 1 33.4 33.4 1,178 1,178

Caristiidae

Caristius macropus Manefish 3 32.4 42.1 365 1,086

Serranidae

Paralabrax nebulifer Barred sand bass 26 32.7 34.4 24 101

Carangidae

Trachurus symmetricus Jack mackerel 90 32.7 46.9 27 981

Sciaenidae Croakers 5 32.7 34.6 62 107

Genyonemus lineatus White croaker 268 32.6 41.4 27 132

Seriphus politus Queenfish 4 32.7 33.4 35 81
302

Table B-1 continued. Frequency of occurrence, depth range, and latitudinal range for fish species, grouped by family (or higher taxonomic 

classification), encountered during the 2003–2008 surveys (unident. = unidentified).

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal degrees) Depth (m)

South North Min. Max.

Embiotocidae Surfperch unident. 7 33.3 36.1 31 119

Amphistichus argenteus Barred surfperch 2 34.4 37.1 53 80

A. rhodoterus Redtail surfperch 2 34.0 39.2 60 73

Cymatogaster aggregata Shiner perch 161 32.6 47.2 26 114

Damalichthys vacca Pile perch 4 32.8 37.1 60 86

Embiotoca lateralis Striped surfperch 2 34.3 34.3 63 64

Hyperprosopon anale Spotfin surfperch 23 34.0 37.6 36 80

Phanerodon atripes Sharpnose surfperch 1 34.0 34.0 83 83

P. furcatus White surfperch 5 33.9 38.0 62 86

Rhacochilus toxotes Rubberlip surfperch 2 32.9 34.4 52 80

Zalembius rosaceus Pink sea perch 507 32.6 39.9 26 276

Sphyraenidae 

Sphyraena argentea California barracuda 1 33.3 33.3 35 35

Bathymasteridae

Ronquilus jordani Northern ronquil 16 38.7 48.4 61 191

Uranoscopidae 

Kathetostoma averruncus Smooth stargazer 16 32.6 34.3 59 225

Stichaeidae

Poroclinus rothrocki Whitebarred prickleback 7 43.0 48.2 88 171

Cryptacanthodidae

Lyconectes aleutensis Dwarf wrymouth 1 43.8 43.8 109 109

Cryptacanthodes giganteus Giant wrymouth 8 45.8 48.2 107 309

Tetragonuridae

Tetragonurus cuvieri Smalleye squaretail 1 37.1 37.1 491 491

Anarhichadidae

Anarrhichthys ocellatus Wolf-eel 24 32.7 48.1 36 145

Ammodytidae

Ammodytes hexapterus Pacific sand lance 2 42.8 44.4 64 83

Trichiuridae

Aphanopus carbo Black scabbardfish 3 32.1 44.3 578 669
303

Table B-1 continued. Frequency of occurrence, depth range, and latitudinal range for fish species, grouped by family (or higher taxonomic 

classification), encountered during the 2003–2008 surveys (unident. = unidentified).

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal degrees) Depth (m)

South North Min. Max.

Lepidopus xantusi Scabbardfish 20 32.7 41.0 163 1,080

Scombridae

Scomber japonicus Chub mackerel 38 32.6 47.6 60 752

Stromateidae Butterfishes 3 33.6 34.3 78 109

Peprilus simillimus Pacific butterfish 242 32.6 47.4 26 550

Centrolophidae

Icichthys lockingtoni Medusafish 9 34.3 46.5 65 775

Icosteidae Ragfish unident. 1 32.8 32.8 1,201 1,201

Icosteus aenigmaticus Ragfish 3 39.4 46.8 598 713

Pleuronectiformes Flatfish unident. 1 34.0 34.0 78 78

Pleuronectiformes larvae Flatfish larvae 1 45.0 45.0 1,196 1,196

Bothidae unident. Lefteye flounder unident. 2 44.3 47.9 63 463

Citharichthys sordidus Pacific sanddab 1,213 32.6 48.4 24 451

C. stigmaeus Speckled sanddab 2 33.3 44.8 35 65

C. xanthostigma Longfin sanddab 42 32.6 34.7 26 125

C. sp. Sanddab unident. 3 35.0 37.8 58 135

Hippoglossina stomata Bigmouth sole 174 32.6 35.5 24 478

Paralichthys californicus California halibut 47 32.7 42.1 24 317

Xystreurys liolepis Fantail sole 25 32.7 34.8 24 104

Pleuronectidae

Atheresthes stomias Arrowtooth flounder 1,261 37.8 48.5 52 1,111

Embassichthys bathybius Deepsea sole 877 32.4 48.2 276 1,428

Eopsetta jordani Petrale sole 1,543 32.9 48.4 36 538

Glyptocephalus zachirus Rex sole 2,406 32.3 48.5 47 937

G. zachirus larvae Rex sole larvae 1 45.4 45.4 433 433

Hippoglossoides elassodon Flathead sole 244 42.0 48.4 62 374

Hippoglossus stenolepis Pacific halibut 300 37.8 48.4 53 492

Isopsetta isolepis Butter sole 44 40.2 47.9 47 178

Lepidopsetta bilineata Southern rock sole 179 32.7 48.4 47 236

Lepidopsetta sp. Rock sole unident. 16 36.8 48.2 36 141
304

Table B-1 continued. Frequency of occurrence, depth range, and latitudinal range for fish species, grouped by family (or higher taxonomic 

classification), encountered during the 2003–2008 surveys (unident. = unidentified).

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal degrees) Depth (m)

South North Min. Max.

Lyopsetta exilis Slender sole 1,929 32.3 48.5 52 680

Microstomus pacificus Dover sole 3,214 32.3 48.5 52 1,246

Parophrys vetulus English sole 1,541 32.6 48.5 24 480

Platichthys stellatus Starry flounder 43 36.8 44.8 36 95

Pleuronichthys decurrens Curlfin sole 401 32.6 48.2 26 380

P. ritteri Spotted turbot 23 32.8 37.7 26 219

P. verticalis Hornyhead turbot 155 32.6 38.1 24 496

Psettichthys melanostictus Sand sole 50 36.3 48.4 36 110

Cynoglossidae Tonguefish unident. 1 36.9 36.9 60 60

Symphurus atricauda California tonguefish 5 34.0 34.4 70 110

Molidae

Mola mola Ocean sunfish 1 37.0 37.0 125 125
305

Table B-2. Frequency of occurrence, depth range, and latitudinal range for invertebrates (grouped by taxonomic classification) encountered during 

the 2003–2008 surveys (unident. = unidentified).

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal 

degree) Depth (m)

South North Min. Max.

Invertebrata Invertebrate unident. 62 32.3 48.2 59 1,241

Invertebrata Invertebrate eggs unident. 3 32.4 40.5 53 1,064

Annelida (phylum) Worm unident. 52 32.8 48.1 71 1,238

Annelid spp. Tube worm unident. 7 32.6 48.4 121 1,169

Polychaeta (class) Polychaete worm unident. 37 32.6 48.4 68 1,239

Aphroditidae 131 32.8 48.4 95 1,201

Aphrodita sp. Sea mouse unident. 313 32.7 48.4 58 1,428

Nereis sp. 1 46.6 46.6 1,002 1,002

Polynoidae (family) Scale worm unident. 1 44.9 44.9 302 302

Arthropoda (phylum)

Isopoda (order) Sea cockroaches unident. 94 32.9 48.3 48 1,253

Anuropsus bathypelagica Giant isopod 1 43.1 43.1 233 233

Pycnogonida (class) Sea spiders 4 33.2 47.6 451 1,141

Crustacea (sub-phylum) Crustacean unident. 1 39.3 39.3 596 596

Malacostraca (class)

Decapoda (order)

Sicyonia ingentis Razor-back prawn 130 32.6 42.7 55 1,127

Malacostraca Shrimp unident. 34 32.6 47.8 24 1,105

Eualus sp. Eualid unident. 30 32.6 48.1 239 1,239

Eualus macrophthalmus Big eyed eualid 212 33.0 48.2 301 1,285

E. biunguis Deepsea eualid 18 34.8 47.8 391 1,341

Notostomus japonicus Japanese spinyridge 11 33.0 48.2 210 1,201

Acanthephyra sp. 1 33.1 33.1 1,186 1,186

Acanthephyra curtirostris Peaked shrimp 36 32.6 47.9 382 1,241

Nematocarcinus sp. Red sword shrimps 1 35.8 35.8 900 900

Hemisquilla californiensis Mantis shrimp 15 32.6 34.0 27 86

Sergestidae (family) Sergestid shrimp unident. 4 33.5 45.1 431 1,010

Sergestes sp. 88 32.9 48.1 118 1,184

Sergestes similis Pacific sergestid 137 32.7 47.9 129 1,140

Euphausiacea Deep sea red shrimps 3 34.3 39.6 92 498
306

Table B-2 continued. Frequency of occurrence, depth range, and latitudinal range for invertebrates (grouped by taxonomic classification) 

encountered during the 2003–2008 surveys (unident. = unidentified).

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal 

degree) Depth (m)

South North Min. Max.

Mysidacea Crested red mysids 22 32.6 47.5 524 1,186

Neognathophausia sp. Red mysids 38 32.3 41.6 483 1,230

Neognathophausia gigas 19 32.4 43.7 756 1,206

N. ingens Giant red mysid 106 32.5 46.1 564 1,271

Neognathophausia sp. A 1 47.7 47.7 991 991

Pandalidae (family) Pandalid shrimp unident. 7 33.2 48.1 108 1,109

Pandalus sp. 5 34.5 44.1 79 787

P. danae Dock shrimp 2 43.1 44.3 61 73

P. jordani Ocean shrimp 578 32.7 48.4 67 1,056

P. tridens Yellowleg pandalid 4 42.9 47.5 250 340

P. platyceros Spot prawn 249 32.7 48.4 70 397

P. hypsinotus Coonstriped shrimp 8 37.1 47.9 53 259

Pandalopsis dispar Sidestripe shrimp 63 32.9 48.4 147 1,214

P. ampla Smooth shrimp 184 32.2 48.0 230 1,341

Crangonidae (family) Crangonid shrimp unident. 1 42.5 42.5 96 96

Crangon sp. 93 34.2 48.4 62 676

Crangon alba Stout crangon 1 35.3 35.3 77 77

C. communis Twospine crangon 42 34.4 47.8 53 215

C. dalli Ridged crangon 5 34.2 41.5 76 204

C. septemspinosa Sand shrimp 10 34.4 47.9 63 156

Pasiphaeidae (family) Pasiphaeid shrimp unident. 5 33.4 41.5 451 854

Pasiphaea pacifica Glass shrimp 531 32.7 48.4 84 1,156

P. tarda Crimson pasiphaeid 485 32.1 47.9 181 1,341

Brachyura Crab unident. 19 32.6 36.3 24 343

Moloha faxoni Pacific carrier crab 9 32.6 33.8 112 151

Cancridae (family) Rock crabs 1 32.7 32.7 83 83

Cancer sp. Cancer crab unident. 13 33.0 38.5 24 455

Cancer branneri Furrowed rock crab 2 39.4 43.2 54 68

C. anthonyi Yellow rock crab 75 32.6 38.7 27 451

C. magister Dungeness crab 1,189 34.5 48.3 36 835
307

Table B-2 continued. Frequency of occurrence, depth range, and latitudinal range for invertebrates (grouped by taxonomic classification) 

encountered during the 2003–2008 surveys (unident. = unidentified).

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal 

degree) Depth (m)

South North Min. Max.

C. productus Red rock crab 127 32.8 39.0 31 437

C. gracilis Graceful cancer crab 30 34.0 47.9 26 97

Calappidae (family) Box crab 14 33.7 48.2 66 287

Platymera gaudichaudii Shame faced crab 263 32.6 44.5 24 480

Majidae (family) Spider crab unident. 1 34.8 34.8 48 48

Oregonia gracilis Graceful decorator crab 4 33.3 44.5 67 101

Macroregonia macrochira Long-armed spider crab 2 32.6 32.8 151 630

Loxorhynchus crispatus Masking crab 21 32.8 44.3 35 464

L. grandis Sheep crab 25 32.7 35.0 27 514

Chorilia longipes Long horned decorator crab 309 32.3 48.4 35 1,230

Chionoecetes sp. Tanner crab unident. 64 33.6 48.2 280 1,197

C. tanneri Grooved tanner crab 1,122 32.6 48.2 64 1,428

C. bairdi Baird’s tanner crab 25 33.0 48.4 124 1,142

C. angulatus Triangle tanner crab 5 44.9 47.7 421 1,428

Hyas lyratus Pacific lyre crab 7 43.2 48.4 73 777

Paguridae Hermit crab unident. 385 32.4 48.4 61 1,268

Paguristes turgidus Hermit crab 38 32.8 48.2 77 968

Pagurus sp. 29 32.7 47.6 99 1,186

Pagurus aleuticus Aleutian hermit crab 2 41.7 41.7 254 273

P. confragosus Knobbyhand hermit crab 1 41.7 41.7 254 254

P. rathbuni Longfinger hermit crab 2 42.3 47.2 180 385

P. tanneri Tanners hermit crab 10 33.5 48.4 168 1,184

P. capillatus Hairy hermit crab 5 41.2 48.0 108 185

Lithodidae unident. Stone crab unident. 2 33.3 34.0 455 823

Lopholithodes sp. Box crab unident. 10 37.0 48.3 64 206

Lopholithodes foraminatus Brown box crab 220 32.7 48.4 67 519

Acantholithodes hispidus Fuzzy crab 24 33.8 48.4 86 502

Lithodes sp. 1 33.9 33.9 1,224 1,224

Lithodes couesi Scarlet king crab 210 32.1 48.1 492 1,246

L. aequispina Golden king crab 1 34.4 34.4 301 301
308

Table B-2 continued. Frequency of occurrence, depth range, and latitudinal range for invertebrates (grouped by taxonomic classification) 

encountered during the 2003–2008 surveys (unident. = unidentified).

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal 

degree) Depth (m)

South North Min. Max.

Rhinolithodes wosnessenskii Rhinoceros crab 1 37.2 37.2 112 112

Paralithodes sp. 2 35.0 45.5 255 267

Paralithodes californiensis California king crab 116 32.6 38.3 82 1,098

P. rathbuni Spiny king crab 76 32.6 38.7 86 1,184

Paralomis sp. 9 32.8 33.5 846 1,246

Paralomis verrilli Vermillion crab 1 34.0 34.0 496 496

P. manningi Deep-sea spider crab 3 33.2 45.9 1,101 1,200

P. multispina Hair crab 168 32.2 47.8 149 1,341

Neolithodes diomedeae Spiky king crab 18 32.6 38.3 302 1,229

Glyptolithodes cristatipes 37 32.7 34.0 269 588

Pugettia sp. Kelp crab 2 33.3 44.3 130 184

Pugettia richii Cryptic kelp crab 8 33.2 44.2 92 147

Galatheidae (family) 16 33.1 48.2 133 1,188

Galatheoidea Squat lobsters and pinch bugs 4 35.6 47.5 1,035 1,241

Janetogalathea californiensis California pinch bug 2 33.2 33.5 379 876

Munidopsis sp. Thorny pinch bugs 28 32.6 47.4 105 1,201

Munidopsis sp. A Thorny pinch bugs sp. A 6 32.3 33.2 972 1,157

Chirostylus sp. Spiny pinch bugs 5 33.5 47.8 344 1,170

Munida hispida Deep water squat lobster 26 32.1 36.2 147 1,230

M. quadrispina Pinch bug 60 32.5 48.3 78 1,215

Panulirus interruptus California spiny lobster 1 34.0 34.0 27 27

Polycheles sculptus Deep-sea lobster 68 32.2 46.0 251 1,264

Pleuroncodes planipes Pelagic red crab 2 34.2 34.2 179 312

Calocarides sp. Deep-sea shrimps 25 33.9 47.5 324 1,170

Callianassa sp. Ghost shrimp 6 34.3 47.9 454 1,095

Callianassa californiensis Bay ghost shrimp 1 44.3 44.3 69 69

Munidopsis quadrata Pale rough pinch 4 33.4 44.6 729 1,166

M. hystrix 3 33.0 33.7 756 826

Thoracica (order)

Thoracica Barnacle unident. 3 36.1 47.5 60 633
309

Table B-2 continued. Frequency of occurrence, depth range, and latitudinal range for invertebrates (grouped by taxonomic classification) 

encountered during the 2003–2008 surveys (unident. = unidentified).

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal 

degree) Depth (m)

South North Min. Max.

Lepas sp. 1 35.6 35.6 1,002 1,002

Brachiopoda (phylum)

Thetys vagina Rabbit-eared salp 21 32.9 48.2 52 1,181

Styela rustica Sea potato 126 32.7 48.3 59 1,224

Cnidaria (phylum)

Anthozoa (class)

Scleractinia unident. Stony coral unident. 7 33.4 48.2 242 1,115

Coenocyathus bowersi Colonial cup coral 1 32.9 32.9 74 74

Polymyces cf montereyensis Cup coral 10 32.7 44.6 78 421

Caryophyllia alaskensis Alaska cup coral 1 32.6 32.6 72 72

Alcyonacea (order) Soft coral unident. 28 32.5 47.9 60 1,215

Alcyonaria unident. Octocoral unident. 3 32.8 45.1 419 915

Anthomastus ritteri Mushroom coral 63 32.6 47.9 293 1,107

Anthomastus sp. 29 32.6 46.9 347 1,094

Anthomastus sp. A Red anthomastus 1 33.7 33.7 447 447

Halocynthia sp. Sea peach unident. 3 36.8 46.8 36 181

Actinaria (order)

Actiniaria Purple striated anemone 207 33.3 47.8 189 1,176

Actiniaria Red striated anemone 179 32.3 48.3 59 1,235

Actiniaria Sea anemone unident. 678 32.5 48.4 36 1,341

Actinostolidae 146 32.4 47.8 53 1,259

Actinostola sp. 37 32.6 48.3 70 1,235

Sicyonis spp. Orange actinistolids 89 33.0 47.8 138 1,238

Hormathiidae (family) Hormathiid anemones unident. 410 32.3 48.2 64 1,271

Hormathiid sp. A Pink hormathiid anemones 4 32.9 47.7 858 1,034

Actinauge verrilli Reticulated anemone 359 32.4 48.2 131 1,428

Actinernus spp. Lava anemones 82 32.4 47.7 352 1,259

Actinoscyphia sp. Sea whip anemones 49 32.6 47.7 63 1,268

Anthopleura xanthogrammica Green anemone 37 37.5 47.9 60 176

Corallimorphus sp. Club tipped anemones 48 32.7 48.2 331 1,205
310

Table B-2 continued. Frequency of occurrence, depth range, and latitudinal range for invertebrates (grouped by taxonomic classification) 

encountered during the 2003–2008 surveys (unident. = unidentified).

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal 

degree) Depth (m)

South North Min. Max.

Liponema brevicornis Pom pom anemone 814 32.2 48.2 83 1,428

Metridium farcimen Giant anemone 1,068 32.6 48.4 53 1,153

M. senile Colonial plumose sea anemone 2 42.4 47.9 64 304

Metridium sp. 8 32.6 44.0 64 380

Oractis diomedeae Grape anemone 5 43.5 47.6 484 969

Paractinostola faeculenta Rough anemone 1,308 32.3 48.2 89 1,428

Stomphia coccinea Swimming anemone 115 32.7 48.4 63 1,240

Stomphia sp. 34 33.5 48.3 66 1,157

Urticina columbiana Columbian anemone 30 33.6 47.8 59 147

U. crassicornis Painted anemone 101 33.5 48.3 56 581

Urticina sp. 90 33.8 48.4 53 521

Gorgonacea (order) Gorgonian coral unident. 42 32.6 47.0 52 1,184

Amphilaphis sp. 1 33.8 33.8 114 114

Antipatharia Black coral 64 33.2 48.1 103 1,285

Antipathes sp. 26 33.7 46.6 82 1,162

Bathypathes sp. Quill black corals 20 32.8 47.8 136 1,243

Calcigorgia sp. 1 47.9 47.9 604 604

Callogorgia kinoshitae Golden coral 19 33.3 48.1 114 1,152

Callogorgia sp. 4 32.1 33.7 129 820

Chrysopathes sp. 18 33.1 47.8 93 1,235

Euplexaura marki Red licorice coral 4 42.8 48.2 181 398

Isidella sp. Articulated bamboo corals 9 33.1 44.8 84 1,092

Keratoisis sp. Forked bamboo corals 3 40.3 47.5 1,063 1,285

Lillipathes sp. Long stemmed black corals 12 43.4 47.6 449 1,243

Lophogorgia chilensis Red gorgonian 4 32.8 47.1 73 710

Paragorgia pacifica 11 34.2 48.3 122 1,166

Paragorgia sp. Peppermint corals 8 33.2 47.6 185 1,092

Pennatulacea Sea pen unident. 132 32.6 48.1 61 1,264

Swiftia pacifica 5 32.4 46.7 78 1,186

S. simplex 28 32.6 48.0 52 1,206
311

Table B-2 continued. Frequency of occurrence, depth range, and latitudinal range for invertebrates (grouped by taxonomic classification) 

encountered during the 2003–2008 surveys (unident. = unidentified).

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal 

degree) Depth (m)

South North Min. Max.

Swiftia sp. Red sea fans 51 32.6 47.7 52 1,184

Pennatulacea (order)

Virgulariidae (family) Sea whip unident. 175 32.7 48.2 59 1,325

Virgularia sp. Smoothstem sea whip 1 42.1 42.1 96 96

Anthoptilum grandiflorum Fleshy sea pen 293 32.6 48.2 80 1,241

Anthoptilum murrayi Murray’s sea pen 22 32.6 47.7 345 1,239

Distichoptilum gracile Deep-sea sea whip 17 32.6 47.6 80 1,229

Halipteris californica 66 36.7 48.3 84 1,237

Halipteris sp. 29 33.6 47.9 81 922

Pennatula phosphorea Branched sea pen 3 34.2 48.0 465 1,179

P. phosphorea californica Deep-sea sea pen 4 34.2 47.7 418 982

Ptilosarcus gurneyi Orange sea pen 66 33.9 47.0 47 798

Stylatula gracile Slender sea whip 146 33.0 47.9 64 1,243

Stylatula sp. Slender sea whips 68 32.7 47.8 63 1,173

Umbellula sp. Flower sea pens 99 32.7 48.1 441 1,428

Hydrozoa (class)

Stylasterina unident. Hydrocoral unident. 2 33.5 35.1 344 544

Stylaster sp. 1 44.4 44.4 102 102

Stylaster alaskanus 1 46.1 46.1 172 172

S. californicus California hydrocoral 2 34.0 34.0 68 94

Distichopora sp. 1 32.9 32.9 1,140 1,140

Scyphozoa (class) Jellyfish unident. 754 32.6 48.3 26 1,428

Chrysaora sp. Chrysaora jellyfish 101 32.6 48.1 52 1,235

Periphylla sp. 2 40.2 44.7 583 1,214

Chrysaora fuscens Sea nettle 49 36.4 47.9 47 728

Periphylla periphylla Purple cone jelly 274 32.6 48.2 317 1,268

Chrysaora melanaster Sunrise jelly 31 37.1 48.0 53 918

Phacellophora camtschatica Egg-yolk jellyfish 165 32.7 48.4 59 1,184

Aequorea sp. 8 35.1 46.9 82 1,130

Phacellophora sp. 1 43.2 43.2 54 54
312

Table B-2 continued. Frequency of occurrence, depth range, and latitudinal range for invertebrates (grouped by taxonomic classification) 

encountered during the 2003–2008 surveys (unident. = unidentified).

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal 

degree) Depth (m)

South North Min. Max.

Atolla sp. Wheel jellies 275 32.1 48.2 64 1,428

Aurelia sp. Moon jellies 105 34.0 47.3 47 1,228

Aurelia labiata 122 34.1 47.9 53 1,162

Pelagia colorata Purple striped jellyfish 9 32.9 36.9 52 464

Cyanea capillata Lion’s mane 3 48.1 48.2 129 248

Ctenophora (phylum) Comb jelly unident. 14 33.9 48.2 60 1,132

Beroe sp. 1 42.6 42.6 1,235 1,235

Echinodermata (phylum)

Asteroidea (class)

Asteroidea unident. Sea star unident. 493 32.6 48.5 26 1,341

Forcipulatida (order)

Ampheraster marianus Pink star 180 32.7 48.1 143 1,185

Ampheraster sp. 10 37.1 47.5 607 998

Anteliaster sp. Soft stars 25 32.7 45.6 123 1,188

Asterias sp. 1 45.6 45.6 115 115

Astrometis sertulifera Fragile rainbow star 1 36.8 36.8 94 94

Brisingella exilis Lacy-armed star 88 32.6 47.9 420 1,240

Brisingella sp. 19 33.5 45.3 344 1,148

Myxoderma platyacanthum Red star 556 32.1 47.9 84 1,241

M. sacculatum Snakehead star 152 32.2 47.7 154 1,297

Orthasterias koehleri Rainbow star 34 33.3 48.4 60 981

Orthasterias sp. 8 41.2 47.5 60 181

Pedicellaster magister Majestic sea star 4 41.0 47.2 171 688

Pedicellaster sp. 4 42.3 45.7 253 957

Pisaster brevispinus Short-spined pink star 267 33.4 48.3 24 723

P. ochraceus 11 43.5 48.1 53 155

Pisaster sp. 3 43.8 44.2 60 312

Pycnopodia helianthoides Sunflower star 504 33.9 48.4 48 1,169

Rathbunaster californicus Deep-sea sunflower star 569 32.3 48.4 53 995

Sclerasterias heteropaes False rainbow star 4 34.0 39.8 87 139
313

Table B-2 continued. Frequency of occurrence, depth range, and latitudinal range for invertebrates (grouped by taxonomic classification) 

encountered during the 2003–2008 surveys.

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal 

degree) Depth (m)

South North Min. Max.

Stylasterias forreri Fish-eating star 392 32.7 48.4 53 1,214

Stylasterias sp. 2 34.9 35.7 74 499

Zoroaster ophiurus 2 41.5 43.8 968 1,019

Z. evermanni Slender star 452 32.6 48.2 97 1,428

Zoroasteridae 1 41.8 41.8 484 484

Paxillosida (order)

Astropecten armatus Spiny sand star 21 33.7 43.0 69 832

A. californicus California sand star 92 32.6 44.5 53 960

A. ornatissimus Ornate sand star 11 32.6 38.5 61 881

Cheiraster dawsoni Fragile star 44 38.9 47.9 72 1,012

Ctenodiscus crispatus Mud star 62 33.2 48.3 163 1,271

Ctenodiscus sp. 5 33.0 45.2 295 1,179

Dipsacaster borealis Northern sand star 7 33.0 45.7 408 1,166

D. eximius Broad sand star 221 32.5 47.5 64 1,246

Dipsacaster sp. 14 33.1 41.9 60 1,145

Leptychaster arcticus Arctic sand star 2 44.7 48.1 133 383

L. pacificus Pacific sand star 4 38.7 48.0 273 991

Leptychaster sp. 2 43.3 44.9 448 451

Luidia asthenosoma Pretty mud star 2 42.8 45.3 164 173

L. foliolata Flat mud star 1,553 32.6 48.4 47 1,125

Luidia sp. 3 32.9 34.4 65 507

Nearchaster aciculosus Deep-sea fragile star 398 32.6 48.2 168 1,428

N. pedicellaris 1 35.8 35.8 1,241 1,241

Nearchaster sp. 26 32.6 47.6 391 1,229

Thrissacanthias penicillatus Carpet star 926 32.6 48.2 60 1,428

Spinulosida (order)

Asthenactis fisheri Slimy deep sea sun star 3 33.2 46.3 1,148 1,184

Crossaster borealis Grooved sun star 605 32.6 48.4 79 1,428

C. papposus Rose star 70 33.6 48.4 66 618

Crossaster sp. 2 33.2 47.3 432 1,089
314

Table B-2 continued. Frequency of occurrence, depth range, and latitudinal range for invertebrates (grouped by taxonomic classification) 

encountered during the 2003–2008 surveys.

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal 

degree) Depth (m)

South North Min. Max.

Diplopteraster multipes Pincushion sea star 105 32.6 48.2 192 1,241

Diplopteraster sp. Pincushion star 3 40.2 46.8 95 906

Henricia aspera Smooth henricia 6 33.4 45.8 129 699

H. clarki Serpent-arm henricia 9 32.7 47.6 280 1,246

H. leviuscula Blood star 10 32.6 46.9 72 182

Henricia sp. 63 32.6 48.3 67 1,230

Heterozonias alternatus Pink sun star 845 32.2 48.2 89 1,428

Lophaster furcilliger Pink crested star 86 32.6 48.0 92 1,428

Lophaster sp. 9 32.9 40.2 489 1,229

L. vexator Crested star 18 33.4 47.6 73 1,184

Peribolaster biserialis 1 34.8 34.8 477 477

Poraniopsis flexilis Flexible thorny star 24 32.6 48.2 67 1,157

P. inflata Thorny star 117 32.6 48.4 70 1,259

Pteraster jordani Jordans slime star 198 32.1 48.2 235 1,428

P. militaris Wrinkled slime star 58 32.8 48.3 92 1,239

Pteraster sp. 68 32.6 48.3 72 1,271

Pteraster sp. cf. temnochiton 1 35.1 35.1 546 546

P. tesselatus Slimy cushion star 35 33.6 48.4 61 1,148

P. trigonodon Triangle-toothed cushion star 13 32.7 45.5 395 1,230

Solaster dawsoni Morning sun star 10 43.2 48.4 54 433

S. endeca Northern sun star 3 44.4 48.3 98 346

S. exiguus Deep-sea sunstar 61 32.7 47.9 255 1,279

S. paxillatus Evening sun sea star 5 48.1 48.3 100 166

Solaster sp. Orange sun stars 65 32.7 48.4 60 1,241

S. stimpsoni Striped sun star 8 43.1 48.2 61 235

Valvatida (order)

Asterina miniata Bat star 19 32.7 38.2 58 755

Ceramaster arcticus Arctic bat sea star 2 36.2 43.2 722 1,166

C. leptoceramus California cookie star 81 32.7 48.0 53 1,188

C. patagonicus Orange cookie star 9 33.3 36.8 201 756
315

Table B-2 continued. Frequency of occurrence, depth range, and latitudinal range for invertebrates (grouped by taxonomic classification) 

encountered during the 2003–2008 surveys (unident. = unidentified).

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence

(no. hauls)

Latitude (decimal 

degree) Depth (m)

South North Min. Max.

Ceramaster sp. 43 32.8 48.0 130 1,208

Cryptopeltaster lepidonotus Grainy star 66 32.4 48.3 72 1,201

Dermasterias imbricata Leather star 52 34.7 48.4 53 173

Hippasteria californica Deep-sea spiny star 536 32.4 48.4 55 1,428

Hippasteria sp. 50 32.1 48.2 60 1,197

H. spinosa Spiny star 436 32.6 48.4 54 1,214

Mediaster aequalis Equal armed star 376 32.6 48.4 54 1,279

Mediaster sp. 42 34.0 48.4 66 604

M.r tenellus Pale equal armed star 18 32.7 47.6 186 1,163

Pseudarchaster alascensis Alaskan pseudarchaster 64 33.1 47.9 69 1,148

P. dissonus Deep-sea pseudarchaster 11 34.9 45.7 70 1,268

P. parelii Scarlet sea star 31 33.5 48.1 95 520

P. pusillus Little pseudarchaster 13 34.2 45.3 133 700

Pseudarchaster sp. 90 32.6 48.4 61 1,243

Crinoidea (class) Crinoid unident. 29 32.6 48.0 86 1,238

Florometra serratissima Feather star 45 32.6 48.3 72 1,184

Florometra sp. 15 33.5 47.7 97 1,104

Psathyrometra fragilis Fragile crinoid 2 32.6 40.3 559 769

Echinoidea (class) Sea urchins/sand dollars unident. 33 32.4 48.3 100 1,158

Allocentrotus fragilis Fragile red sea urchin 1,271 32.3 48.4 61 1,224

Allocentrotus sp. 5 33.6 45.2 328 795

Brisaster latifrons Mud urchin 1,041 32.6 48.4 72 1,268

Brisaster sp. 148 32.9 48.3 85 1,224

B. townsendi Giant mud urchin 31 33.3 48.1 122 1,013

Brissopsis pacifica Oval sea biscuit 126 32.6 39.1 76 937

Echinacea unident. 258 32.7 48.5 63 1,157

Lytechinus anamesus White urchin 30 32.6 48.4 60 248

Spatangoida Heart sea urchin unident. 60 32.3 48.2 113 972

Spatangus californicus Giant sea biscuit 109 32.6 39.1 73 498
316

Table B-2 continued. Frequency of occurrence, depth range, and latitudinal range for invertebrates (grouped by taxonomic classification) 

encountered during the 2003–2008 surveys (unident. = unidentified).

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal 

degree) Depth (m)

South North Min. Max.

Strongylocentrotus 

droebachiensis

Green sea urchin 8 32.6 48.4 91 370

S. echinoides 1 34.2 34.2 115 115

S. franciscanus Red sea urchin 10 32.7 34.4 70 163

S. pallidus Crowned sea urchin 24 32.7 48.4 62 346

Strongylocentrotus sp. 8 32.6 48.4 76 211

Brisaster spp. Brisaster spp/brissopsis pacifica

unident.

45 32.7 36.2 155 1,048

Crushed urchin 141 32.6 48.4 100 999

Holothuroidea (class)

Holothuroidea unident. Sea cucumber unident. 224 32.6 48.5 35 1,285

Cucumaria sp. 3 34.4 48.4 227 503

Dromalia alexandri Pineapple benthic siphonophore 278 32.3 36.9 90 916

Molpadia intermedia Purple sea potato 193 32.8 48.4 60 1,271

Molpadia sp. 1 33.2 33.2 1,241 1,241

Paelopatides confundus Deep sea swimming sea cucumber 4 32.2 39.3 1,086 1,188

Pannychia moseleyi Sloppy cucumber 281 32.4 48.2 145 1,268

Parastichopus californicus California cucumber 277 32.6 48.4 43 1,178

P. leukothele Giant soft cucumber 566 32.6 48.4 54 1,241

P. parvimensis Warty sea cucucumber 15 32.7 34.5 259 1,028

Parastichopus sp. 9 32.6 48.3 77 1,205

Pseudostichopus mollis Sandy sea cucumber 261 32.3 48.2 95 1,268

Pseudostichopus sp. 1 47.8 47.8 397 397

Psolidae 1 33.2 33.2 879 879

Psolus fabricii Brownscaled sea cucumber 1 36.4 36.4 1,050 1,050

P. squamatus White-scaled cucumber 137 32.3 48.1 235 1,197

Scotoplanes spp. Sea pigs 146 32.2 48.1 589 1,428

Synallactidae (family) Deep sea papillose sea cucumbers 47 32.1 34.5 72 1,246

Synallactes challengeri 7 32.8 33.6 200 1,188

Ophiuroidea (class)
317

Table B-2 continued. Frequency of occurrence, depth range, and latitudinal range for invertebrates (grouped by taxonomic classification) 

encountered during the 2003–2008 surveys (unident. = unidentified).

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal 

degree) Depth (m)

South North Min. Max.

Ophiuroida unident. Ophuiroid unident. 69 32.7 48.4 35 1,279

Amphiophiura sp. Southern armored brittle stars 26 32.7 47.6 820 1,428

Amphiura diomedeae Tan brittle star 9 32.9 48.1 727 1,271

Amphiuridae unident. 2 33.0 46.9 968 1,111

Asteronyx longifissus Long-slit serpent star 88 32.5 47.8 132 1,241

A. loveni Giant serpent star 121 33.0 47.9 71 1,271

Asteronyx sp. 56 32.6 47.8 76 1,238

Gorgonocephalus eucnemis Basket star 234 32.6 48.4 54 1,206

Ophiacantha diplasia Lacy brittle star 22 32.7 45.6 123 910

O. enneactis 1 45.2 45.2 371 371

Ophiacantha sp. 43 32.6 46.4 68 1,297

Ophiomusium jolliensis Red brittle star 45 32.6 47.9 215 1,200

O. lymani Lymans brittle star 9 32.6 46.0 547 1,243

Ophiopholis aculeata Ubiquitous brittle star 1 47.7 47.7 1,095 1,095

O. longispina Longspined brittle star 8 33.8 45.4 114 948

Ophiopholis sp. 52 32.7 48.4 85 998

Ophiopthalmus normani Rosy brittle star 6 32.9 46.2 1,098 1,157

Ophioscolex corynetes Blob disc serpent star 5 43.5 45.5 449 594

Ophioscolex sp. 2 45.4 45.4 408 467

Ophiura sarsi Notched brittle star 176 34.7 48.4 64 923

Ophiura sp. 32 32.7 47.8 61 1,160

Ophiurida Brittlestars unident. 55 32.7 48.2 70 1,246

Stegophiura ponderosa Giant armored brittle star 106 32.8 47.9 322 1,241

Aplacophora (class) Aplacophora unident. 16 34.2 48.1 74 1,186

Alexandromenia agassizi Sponge dwelling aplacophoran 2 32.9 33.0 347 395

Neomenia sp. Solenogasters 81 32.7 48.3 69 1,264

Bivalvia (class)

Acesta sphoni Sphons giant file clam 10 32.7 33.8 493 1,114

Acharax johnsoni 6 42.1 48.1 449 1,143
318

Table B-2 continued. Frequency of occurrence, depth range, and latitudinal range for invertebrates (grouped by taxonomic classification) 

encountered during the 2003–2008 surveys (unident. = unidentified).

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal 

degree) Depth (m)

South North Min. Max.

Bivalvia (empty shells) Empty bivalve shells 1 33.4 33.4 491 491

Bivalvia unident. Bivalve unident. 48 33.0 48.2 66 1,224

Calyptogena pacifica Cold seep clam 3 32.6 46.5 458 1,271

Chlamys hastata hericia 2 36.5 44.5 72 97

C. rubida Reddish scallop 1 32.8 32.8 59 59

Cyclopecten davidsoni Salmon glass-scallop 1 45.0 45.0 743 743

Delectopecten vancouverensis Glass scallop 67 32.7 47.8 95 1,271

Gastropod eggs Snail eggs 94 32.6 47.9 60 1,235

Halicardia perplicata Deep-sea heart clam 2 33.0 44.5 968 1,129

Mytilidae (family) Mussel unident. 2 33.6 46.3 216 1,056

Panopea abrupta Pacific geoduck 3 37.1 42.6 58 80

Parvamussium alaskense Alaska glass-scallop 12 33.0 47.8 611 1,197

Patinopecten caurinus Weather vane scallop 13 40.9 47.4 72 141

Pectinid unident. Scallop unident. 5 33.2 48.1 70 1,154

Solemya sp. Awning clam 1 46.3 46.3 957 957

Teredinidae Shipworm unident. 1 32.7 32.7 76 76

Tochuina tetraquetra Giant orange tochui 2 34.0 43.0 95 235

Tritonia sp. 6 33.7 45.9 129 1,094

Vesicomya pacifica Cold seep clam 9 33.9 48.1 505 1,224

Yoldia scissurata Crisscrossed yoldia 1 43.7 43.7 103 103

Yoldia sp. 3 34.4 46.5 820 885

Cephalopoda (class)

Abraliopsis felis 2 47.2 47.5 174 562

Benthoctopus leioderma Smooth octopus unident. 15 33.9 48.4 98 1,197

Benthoctopus spp. 379 32.2 48.4 70 1,428

Berryteuthis magister Magistrate armhook squid 21 32.7 48.4 163 1,127

Cephalopoda unident. Cephalopod unident. 1 47.5 47.5 903 903

Chiroteuthis calyx 32 33.5 47.8 104 1,155

Cranchia scabra Sandpaper squid 7 34.0 40.1 455 1,105

Cranchiidae Glass squids 10 32.1 47.7 443 1,032
319

Table B-2 continued. Frequency of occurrence, depth range, and latitudinal range for invertebrates (grouped by taxonomic classification) 

encountered during the 2003–2008 surveys (unident. = unidentified).

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal 

degree) Depth (m)

South North Min. Max.

Dosidicus gigas Rojo diablo humboldt squid 126 32.3 47.4 66 1,153

Enteroctopus dofleini Giant pacific octopus 153 32.8 48.4 35 734

Galiteuthis phyllura Arrow squid 11 33.5 47.8 176 1,235

Gonatopsis borealis North Pacific armhook squid 77 32.7 48.0 156 1,235

Gonatopsis sp. 1 37.0 37.0 737 737

Gonatus onyx Clawed armhook squid 203 32.4 48.0 58 1,228

Gonatus sp. 18 32.7 47.9 138 1,063

Graneledone boreopacifica Ghost octopus 20 32.7 47.5 1,040 1,285

Graneledone sp. Deep-sea octopus unident. 10 32.6 47.8 1,125 1,428

Grimpoteuthis albatrossi Albatross’ flapjack 1 47.6 47.6 1,076 1,076

Histioteuthis heteropsis Jewel squid 171 32.7 47.8 102 1,201

H. hoylei Long armed jewel squid 15 35.6 47.7 387 1,098

Histioteuthis sp. Jewel squids 3 32.8 33.5 493 1,184

Japetella diaphana Yellow ringed octopus 67 32.6 48.1 461 1,285

Loligo opalescens California market squid 706 32.6 48.2 26 888

Moroteuthis robusta Big squid 18 34.3 47.3 273 700

Octopodidae Octopus unident. 41 32.6 48.4 43 1,285

Octopoteuthis deletron Octopus squid 513 32.6 48.2 273 1,297

Octopus californicus North Pacific bigeye octopus 175 32.6 47.0 64 754

O. rubescens Red octopus 40 32.7 47.8 47 516

Octopus sp. 15 32.8 48.4 59 1,251

Onychoteuthidae 1 32.6 32.6 897 897

Onychoteuthis borealijaponicus Boreal clubhook squid 3 34.5 41.5 380 948

Opisthoteuthis californiana Flapjack devilfish 313 32.8 48.2 60 1,241

Rossia pacifica Bobtail squid 304 32.8 48.3 56 1,151

Taonius pavo Cone squid 46 34.2 47.7 75 1,251

Teuthida Squid unident. 40 32.6 48.1 84 1,237

Teuthida eggs Squid eggs unident. 1 33.4 33.4 1,206 1,206

Vampyroteuthis infernalis Vampire squid 238 32.4 48.2 94 1,297

Gastropoda (class)
320

Table B-2 continued. Frequency of occurrence, depth range, and latitudinal range for invertebrates (grouped by taxonomic classification) 

encountered during the 2003–2008 surveys (unident. = unidentified).

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal 

degree) Depth (m)

South North Min. Max.

Heteropoda (order) 38 32.3 47.7 68 1,243

Carinaria cristata 11 33.2 47.6 86 1,184

Heteropod/pteropod unident. 7 32.7 46.7 73 1,193

Aforia goodei Goode’s aforia 3 33.2 46.7 947 1,271

Anisodoris nobilis Pacific sea lemon 22 32.7 48.4 61 564

Antiplanes perversa Lefthanded turrid 6 39.5 45.0 84 1,049

Antiplanes sp. A Green lefthanded turrids 1 43.5 43.5 554 554

Archidoris odhneri White night doris 3 44.3 45.8 98 180

Armina californica Striped tongue nudibranch 17 41.8 48.4 53 211

Bathybembix bairdii Green top snail 599 32.1 48.2 285 1,246

Boreotrophon spp. 1 45.3 45.3 384 384

Buccinum sp. eggs 1 45.3 45.3 384 384

Buccinum spp. 35 33.2 47.7 366 1,144

Buccinum strigillatum Striated whelk 24 33.0 47.6 276 1,215

B. viridum Turban whelk 59 34.5 48.0 355 1,297

Bulbus fragilis Fragile moonsnail 91 32.6 48.4 67 337

Calinaticina oldroydii Oldroyd’s moonsnail 98 33.3 48.2 60 419

Calliostoma platinum Silvery top snail 4 34.4 45.1 341 828

Colus aphelus Oblique whelk 10 35.1 47.4 371 1,169

C. georgianus Georgias whelk 4 34.4 48.0 820 1,008

Colus spp. 45 33.3 47.9 218 1,238

C. trophius Ribbed whelk 3 34.4 47.7 805 982

Cryptonatica russa Rusty moonsnail 4 34.4 46.2 550 1,210

Dorididae Dorid nudibranch unident. 19 32.7 47.7 67 1,088

Euspira lewisii Moon snail 22 33.8 47.9 79 347

Exilioidea rectirostris Spindle shell 2 35.2 43.1 496 1,048

Fusitriton oregonensis Oregon triton 82 40.3 48.4 91 785

Gastropod unident. Snail unident. 232 32.7 48.4 48 1,325

Gastropoda (empty shells) Empty gastropod shells 3 34.6 43.8 204 1,176

Lepeta sp. Deep-sea limpets unident. 1 44.6 44.6 70 70
321

Table B-2 continued. Frequency of occurrence, depth range, and latitudinal range for invertebrates (grouped by taxonomic classification) 

encountered during the 2003–2008 surveys (unident. = unidentified).

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal 

degree) Depth (m)

South North Min. Max.

Megasurcula spp. 1 39.0 39.0 125 125

Megasurcula stearnsiana Stearns’ turrid 1 34.4 34.4 86 86

Natica sp. 1 46.2 46.2 1,125 1,125

Neptunea amianta Deep-sea whelk 416 32.6 48.2 102 1,279

N. eulimata 1 44.1 44.1 446 446

N. humboldtiana Humboldt whelk 17 39.1 48.0 307 1,008

N. ithia Slender whelk 2 40.5 46.0 114 359

N. phoenicia Phoenicean whelk 2 46.9 47.7 521 1,068

N. pribiloffensis Pribilof whelk 8 39.1 47.0 333 1,193

N. smirnia Chocolate whelk 3 45.5 48.2 90 1,152

N. sp. 459 32.3 48.2 134 1,241

N. stilesi Stile’s whelk 2 42.3 45.7 253 280

N. tabulata Tabled whelk 4 33.1 46.0 93 850

Nudibranchia unident. Nudibranch unident. 254 32.6 48.4 26 1,325

Philine bakeri Ocean won ton 79 32.3 38.5 77 531

Pleurobranchaea californica California sea slug 499 32.6 44.8 55 1,209

Plicifusus (formerly Colus) sp. 2 37.4 42.2 734 969

Plicifusus griseus Gray whelk 16 33.0 48.0 689 1,011

Polinices sp. 2 38.0 38.0 60 62

Puncturella rothi Keyhole limpet 4 34.0 48.4 66 211

Solariella nuda Naked solarelle 5 36.2 42.3 467 906

Solariella sp. 1 33.4 33.4 960 960

Tritonia diomedea Rosy tritonia 915 32.4 48.1 55 1,279

Nudibranch sp.a 22 33.3 48.3 65 1,056

Polyplacophora

Polyplacophora unident. Chiton unident. 2 34.4 48.1 70 1,152

Brachiopoda (phylum)

Brachiopod unident. Lampshells unident. 43 32.8 47.7 85 1,170

Laqueus californianus California lamp shell 12 33.4 45.8 91 504

Terebratalia transversa Common brachiopod 9 40.6 46.8 109 449
322

Table B-2 continued. Frequency of occurrence, depth range, and latitudinal range for invertebrates (grouped by taxonomic classification) 

encountered during the 2003–2008 surveys (unident. = unidentified).

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal 

degree) Depth (m)

South North Min. Max.

Nematoda

Nematoda Nematode worm unident. 18 32.6 47.7 65 1,188

Platyhelminthes

Platyhelminthes Flatworm unident. 1 45.0 45.0 143 143

Porifera Sponge unident. 797 32.1 48.5 59 1,285

Acanthascus sp. Chimney sponges 16 32.6 47.3 123 1,088

Aphrocallistes vastus Clay pipe sponge 311 32.6 48.4 77 1,268

Chonelasma calyx Goblet sponge 8 32.7 47.6 275 1,024

Farrea convolulus Crusty tube sponge 8 32.9 47.7 307 1,083

Hexactinellida Glass sponge unident. 18 33.2 48.0 114 1,097

Hyalonema sp. Fiber optic sponges 117 32.3 48.0 35 1,285

Leucandra heathi Spiny vase sponge 13 32.6 47.7 133 1,239

Mycale sp. 1 34.8 34.8 477 477

Polymastia pachymastia Black-orange spud sponge 1 45.7 45.7 112 112

Rhabdocalyptus sp. Cloud sponges 42 32.7 47.9 129 1,125

Staurocalyptus sp. Spiny vase sponges unident. 53 32.6 48.2 122 1,184

Suberites ficus Hermit sponge 2 32.7 33.8 87 130

Suberites sp. Deep sea free living sponges 5 32.6 34.4 121 395

Tethya sp. Ball sponge 11 32.6 45.1 66 826

Vase sponge 43 32.6 48.0 70 1,147

White claypipe sponge 5 33.5 48.2 130 1,193

Soft green sponge 4 46.8 48.2 285 1,197

Mushroom sponge 2 35.3 46.3 528 889

Firm yellow green sponge 1 32.6 32.6 133 133

Sipuncula (phylum) Peanut worm unident. 18 33.1 47.4 59 1,193

Gymnolaemata Bryozoan unident. 1 33.3 33.3 846 846

Cellepora ventricosa Coral bryozoan 1 47.5 47.5 819 819

Sipuncula

Tunicata Tunicate unident. 62 32.8 47.7 52 1,246

Urochordata
323

Table B-2 continued. Frequency of occurrence, depth range, and latitudinal range for invertebrates (grouped by taxonomic classification) 

encountered during the 2003–2008 surveys (unident. = unidentified).

Taxonomic group and 

scientific name Common name

Frequency of 

occurrence 

(no. hauls)

Latitude (decimal 

degree) Depth (m)

South North Min. Max.

Thaliacea unident. Salps unident. 878 32.3 48.4 53 1,268

Pyrosoma atlanticum Sea tongue 7 33.5 36.2 87 1,154

Halocynthia igaboja Spiny tunicate 28 34.2 47.5 67 947

H. aurantium Sea peach 2 32.9 33.2 882 1,157

Cnemidocarpa finsmarkiensis Solitary heart tunicate 1 33.2 33.2 882 882

Halocynthia hispidus Hairy tunicate 1 34.1 34.1 95 95

Molgula griffithsii Sea grape 53 32.9 48.2 69 1,239

Recent NOAA Technical Memorandums

published by the

Northwest Fisheries Science Center

NOAA Technical Memorandum NMFS-NWFSC113 Ford, M.J. (ed.). 2011. Status review update for Pacific salmon and steelhead listed under the 

Endangered Species Act: Pacific Northwest. U.S. Dept. Commer., NOAA Tech. Memo. NMFS-NWFSC113, 281 p. NTIS number pending.

112 Skidmore, P.B., C.R. Thorne, B.L. Cluer, G.R. Pess, J.M. Castro, T.J. Beechie, and C.C. Shea. 

2011. Science base and tools for evaluating stream engineering, management, and restoration proposals. 

U.S. Dept. Commer., NOAA Tech. Memo. NMFS-NWFSC-112, 255 p. NTIS number pending.

111 Leonard, J., and P. Watson. 2011. Description of the input-output model for Pacific Coast fisheries. 

U.S. Dept. Commer., NOAA Tech. Memo. NMFS-NWFSC-111, 64 p. NTIS number PB2011-113405.

110 Ainsworth, C.H., I.C. Kaplan, P.S. Levin, R. Cudney-Bueno, E.A. Fulton, M. Mangel, P. TurkBoyer, J. Torre, A. Pares-Sierra, and H.N. Morzaria Luna. 2011. Atlantis model development for the 

northern Gulf of California. U.S. Dept. Commer., NOAA Tech. Memo. NMFS-NWFSC-110, 293 p. 

NTIS number PB2011-113404.

109 Levin, P.S., and F.B. Schwing (eds.). 2011. Technical background for an integrated ecosystem 

assessment of the California Current: Groundfish, salmon, green sturgeon, and ecosystem health. U.S. 

Dept. Commer., NOAA Tech. Memo. NMFS-NWFSC-109, 330 p. NTIS number PB2011-112724.

108 Drake, J.S., E.A. Berntson, J.M. Cope, R.G. Gustafson, E.E. Holmes, P.S. Levin, N. Tolimieri, 

R.S. Waples, S.M. Sogard, and G.D. Williams. 2010. Status review of five rockfish species in Puget 

Sound, Washington: Bocaccio (Sebastes paucispinis), canary rockfish (S. pinniger), yelloweye rockfish 

(S. ruberrimus), greenstriped rockfish (S. elongatus), and redstripe rockfish (S. proriger). U.S. Dept. 

Commer., NOAA Tech. Memo. NMFS-NWFSC-108, 234 p. NTIS number PB2011-107576.

107 Lian, C.E. 2010. West Coast limited entry groundfish trawl cost earnings survey protocols and results for 

2004. U.S. Dept. Commer., NOAA Tech. Memo. NMFS-NWFSC-107, 35 p. NTIS number PB2011-

102712.

106 Harvey, C.J., K.K. Bartz, J. Davies, T.B. Francis, T.P. Good, A.D. Guerry, B. Hanson, 

K.K. Holsman, J. Miller, M.L. Plummer, J.C.P. Reum, L.D. Rhodes, C.A. Rice, J.F. Samhouri, 

G.D. Williams, N. Yoder, P.S. Levin, and M.H. Ruckelshaus. 2010. A mass-balance model for 

evaluating food web structure and community-scale indicators in the central basin of Puget Sound. U.S. 

Dept. Commer., NOAA Tech. Memo. NMFS-NWFSC-106, 180 p. NTIS number PB2011-102711.

Most NOAA Technical Memorandums NMFS-NWFSC are available at the

Northwest Fisheries Science Center Web site, http://www.nwfsc.noaa.gov