UNITED STATES DEPARTMENT OF THE INTERIOR 

Harold L. Ickes, Secretary

GEOLOGICAL SURVEY 

W. C. Mendenhall, Director

Professional Paper 181

UPPER EOCENE FORAMINIFERA OF THE 

SOUTHEASTERN UNITED STATES

JOSEPH A. CUSHMAN

UNITED STATES

GOVERNMENT PRINTING OFFICE 

WASHINGTON : 1935

For salo by llio Superintendent of Documents, Washington, D.C. Price 25 cents
CONTENTS

Lituolidae---- _--____ ------ _____ __

Textulariidae..- .--__. _____ ._-- _ _ _

Valvulinidae---.- ______ - _-__.- ___ _____

Miliolidae __ --------- _ __ _-___-

Ophthalmidiidae.--. ... __ ,. ... .......

Lagenidae

Nonionidae____- __ _ _______ - _ _ _

Camerinidae___- __ _J - ____ _._

Page

6 Ellipsoidinidae _ _ '___:___ ______ ______

7 Rotaliidae. __ _. _________ _ _.__ _ ____

10 Chilostomellidae----- _.. _ - __._ __-_-_---___

11 Hantkeninidae _______ _ ______ ._ _ ___ ________

32 Index ______._ ___________-___-_____-____________.__

Page 

33 

34 

42 

42 

49 

49 

49 

50 

50 

54 

55 

85

ILLUSTRATIONS

PLATE 1. Lituolidae and Textulariidae . __.

2. Textulariidae, Verneuilinidae, Valvulinidae,

4. Miliolidae, Ophthalmidiidae, and Lagenidae _

9. Lagenidae and Polymorphinidae . . _ _ _

12. Nonionidae and Heterohelicidae. _ _ _ _ _ _ _ _ _

Distribution of Eocene Foraminifera in Florida _ __

Distribution of species of Foraminifera in Georgia. ____

Page

63 16. Buliminidae, Ellipsoidinidae, and Rotaliidae64 17-19. Rotaliidae___ ---------------------- ____

65 20. Rotaliidae, Cassidulinidae, and Chilostomel66-69 lidae

72 23. Anomalinidae, Planorbulinidae, and Ruper73 ' tiidae.. __ __ _----__-_----_-__-_-_____

INSERTS

Page 

60 Distribution of Eocene Foraminifera in Alabama- ____

60

Page

74 

75-76

77 

78-80

81 

82 

83

84

Pago 

60

II
UPPER EOCENE FORAMINIFERA OF THE SOUTHEASTERN UNITED STATES

By JOSEPH A. CUSHMAN

ABSTRACT

This paper describes the Foraminifera (except the Orbitoididae) of upper Eocene (Jackson) age from the States between 

North Carolina and Texas. About 203 species or varieties, 

including 47 new species, are described, and most of them are 

figured. Faunas of both shallow and deep water facies are 

recognized. As a whole, the shallow-water faunas are more 

closely related to the Recent shallow-water faunas of the 

Indo-Pacific region than to the Recent West Indian faunas, 

but the deeper-water species are most closely related to the 

Recent species of the Gulf of Mexico and the West Indies. 

The Cooper marl of South Carolina contains many species not 

found in the Gulf States.

INTRODUCTION

The uppe7r Eocene (Jackson formation) is represented in the Atlantic and Gulf Coastal Plain of the 

United States by a varying series of sediments. Some 

of these evidently represent very shallow water and 

conditions not unlike those which prevail today in the 

warm waters of southern Florida and the West Indies. 

Other sediments represent much deeper and somewhat 

cooler water. The fauna of the Jackson is therefore 

a varied one, as it covers a wide range of ecologic conditions. There is also a great range in the condition 

of preservation of the Foraminifera; many of those 

in the limestones are very difficult to determine, 

although they are in many places fairly common. On 

the other hand, numerous specimens from the finer 

clays, deposited in deeper water, are beautifully preserved in their minute details and are fully as satisfactory for study as living material.

There are very many undescribed species, and the 

Jackson as a whole has a fauna which is unlike most 

of those already described from the upper Eocene of 

other regions. As a whole the shallow-water faunas 

are most closely like the present-day faunas of shallow 

water in the Indo-Pacific, rather than the living West 

Indian fauna. The deeper-water group, however, is 

much more closely allied with forms still living in the 

present Gulf of Mexico and West Indian region. The 

shallow-water group has therefore migrated from this 

general region during the time interval since the Jackson, whereas the deeper-water representatives have 

remained.

The fauna of the Jackson includes many species 

which are identical with or very closely related to 

species in the Oligocene of the Gulf Coastal Plain.

Many of those which are not identical represent 

ancestral forms from which undoubtedly the Oligocene 

species were derived. There are, however, a great 

many species in the Jackson which apparently did not 

persist into the Oligocene. A few species are identical 

with or closely allied to species known from the 

Claiborne group.

Some of the larger Miliolidae, such as those from 

Jackson, Miss., seem to be closely allied to similar 

forms found in the Eocene of the Paris Basin. As 

a whole, however, this group is poorly preserved, 

although specimens are fairly common.

The fauna of the Jackson is economically important, as in the western portion of the area (Louisiana 

and Texas) it is often used in the correlation of sediments and in the determination of the age of surface outcrops and of well drillings. The Jackson 

is also brought up toward the surface about some 

of the coastal salt domes. For the general worker 

on the group it has been thought advantageous to 

describe and illustrate the fauna rather than to 

attempt any detailed discussion of vertical ranges 

of the particular species. With adequate definitions and illustrations it will be possible for the 

economic worker to plot for himself the vertical 

ranges in any particular area for which he has 

detailed information.

Some of the forms that are very common in the 

Jackson, such as Hmtkenina, are widely distributed 

in the areas about the Gulf of Mexico and Caribbean 

regions and serve as excellent index fossils for this 

particular period.

The species of the Cooper marl of South Carolina 

show perhaps less definite correlation with some of 

the other formations of this region than the species 

occurring in formations that crop out along the Gulf 

Coastal Plain. Although many species are common 

to the two areas, the Cooper marjl presents many 

species not found at all, so far as present collections 

show, in the Gulf Coastal Plain.

In the systematic descriptions the various genera 

and species are arranged according to the latest ideas 

in regard to the classification of the Foraminifera. 

This may make some difficulties in comparisons between 

this paper and those on the Oligocene Foraminifera 

(Professional Papers 129 and 133), but where the

1
UPPER EOCENE FOKAMINIFERA OP THE SOUTHEASTERN UNITED STATES

species are identical the older name will be found in 

the synonymy in the present paper.

There are two groups of the so-called larger Foraminifera. A report on one group has already been 

published.1 Descriptions of these species with additions will be found in their proper place in this paper, 

but the illustrations are not repeated. The other 

group, the Orbitoididae, includes several genera, especially Lepidocyclina and Discocyclina. As this group 

is now receiving a great deal of special attention, no 

attempt has been made to include members of this 

family in the present paper. Most of the species of 

this group occurring in the upper Eocene of the 

Coastal Plain region are described and illustrated in 

an earlier paper.2

Most of the species described in the present paper 

are illustrated on the accompanying plates. Plates 1 

to 6 are reproduced from photographs, and the other 

plates from drawings by Miss Margaret S. Moore.

A list of the stations from which material has been 

studied, with the names of the collectors, is given 

below. The data in somewhat abbreviated form are 

also given under each species.

NORTH CAROLINA

Castle Hayne marl

5611. Wilmington, New Honover County.

8165. Erroneously assigned to Trent marl (lower Miocene). 

Label reads: " Trent fm. From marl pits 2 miles north from 

creek near Maple Cypress, Craven County, farm of J. H. 

Stokes. J. A. Cushman, 1918."

8171. At bridge across New River on road to Jacksonville, 

4 miles from Richlands, Onslow County. J. A. Cusbman, 1918.

SOUTH CAROLINA

Cooper marl

5531. Navy yard, Charleston. Sloan's locality 444. Earle 

Sloan.

6319. Ingleside marl pit, Ingleside, Berkeley County. T. W. 

Vaughan and Earle Sloan.

6321. Mill Creek, Baldock, Allendale County, formerly Barnwell County. T. W. Vaughan.

11857. Pit on US Highway 17, 1 mile south of Moncks Corner, Berkeley County. C. W. Cooke, 1928.

11861. Steep bluff on west side of Biggin Creek, three-fourths 

mile below the coastal highway, Berkeley County. C. W. 

Cooke, 1928.

11862. Dump heap from Charleston aqueduct tunnel, about 1 

mile west of Givhans, Dorchester County. C. W. Cooke, 1928.

11863. At end of fill at west side of Santee Canal on US 

Highway 17, three-fourths mile west of old Biggin Church, 

Berkeley County. C. W. Cooke, 1928.

 . Baldock. C. W. Gooke (C-48-16). 

3595. Barhwell County.

 . Ashley marl works, Dorchester County.

1 Cushman, J. A., American species of Operculina and Heterostegina 

and their faunal relations: U.S. Geol. Survey Prof. Paper 128, 

pp. 125-146, 1921.

2 Cushman, J. A., The American species of Ortliophragmina and 

Lepidoeyclina: U.S. Geol. Survey Prof. Paper 125, pp.. 39-108, 1921.

GEORGIA

Ocala limestone

3387. Red Bluff, Flint River, 7 miles above Bainbridge, 

Decatur County. T. W. Vaughan.

3390. Below Plant System railroad wharf, Bainbridge, 

Decatur County. T. W. Vaughan, 1900.

3617. Just above railroad bridge on Muckafoonee Creek, 

near Albany, Dougherty County. T. W. Vaughan, 1902.

3626. Well 4% miles southwest of Philema, Lee County, 30 

feet down. T. W. Vaughan, 1902.

3760. Georgia Southern & Florida Railway, half a mile 

south of Tivola, Houston County. S. W. McCallie, 1903.

3966. 2 miles south of Perry, Houston County. S. W. McCallie.

6110. Steamboat landing, Bainbridge, Decatur County.

6160. Flint River, 4 miles below Bainbridge, Decatur County. 

L. W. Stephenson.

6161. Flint River, Bainbridge, Decatur County. L. W. 

Stephenson.

6171. Flint River, Bainbridge, Decatur County. L. W. 

Stephenson.

7097. East bank of Flint River at bend three-fourths mile 

northeast of Atlantic Coast Line railroad station at Bainbridge. C. W. Cooke and W. C. Mansfield.

7098. Red Bluff, west bank of Flint River 7 miles above 

Bainbridge, Decatur County. C. W. Cooke. ,

7099. East bank of Flint River about half a mile above 

Red Bluff, Decatur County. C. W. Cooke.

7108. Flint River at Seaboard Air Line Railway bridge, 

Sumter County. C. W. Cooke and J. E. Brantly.

7115. East bank of Flint River at bend to west 6 miles below Burke Ferry and about 6 miles above Georgia Southwestern & Gulf Railway bridge near Warwick, Worth County.

 7116. Flint River, Lee County, about 1% miles above Georgia Southwestern & Gulf Railway bridge, at base of bluff. 

0. W. Cooke.

7117. Flint River, Lee County, about 1% miles above Georgia Southwestern & Gulf Railway bridge, upper beds. C. W. 

Cooke.

7119. East bank of Flint River, Worth County, near 

Dougherty County line, about 16 miles above Albany. C. W. 

Cooke. .

7120. West bank of Flint River half a mile below Georgia 

Northern Railway bridge at Albany, Dougherty County. C. W. 

Cooke.

7123. East bank of Flint River 9% miles below Albany, 

Dougherty County, 4% miles below Blue Spring. 0. W. 

Cooke.

7126. Flint River, Mitchell County, at Dry Bread Shoals, 

8% or 9 miles below Newton, Baker County, from dumps 

blasted from the channels. C. W. Cooke.

7127. A quarter of a mile below Norman's Ferry, Flint River, 

Mitchell County. C. W. Cooke and J. E. Brantly.

7130. East bank of Flint River 1^ miles below Red Bluff, 

Decatur County. C. W. Cooke.

7189. Rich Hill, Crawford County. C. W. Cooke.

7235. East bank Flint River near northeast corner of 

Dougherty County. C. W. Cooke.

7237. Quarry half # mile east of Huguenen Ferry, Crisp 

County. C. W. Cooke and J. E. Brantly.

7689. Quarry on Perry-Fitzgerald road 1 mile south of 

Tivola, Houston County. C. W. Cooke.

7704. Chalk Hill, 3 miles west of Tartersville, Twigg County. 

C. W. Cooke and H. K. Shearer.

7719. Old Spring Mill, Spring Mill Branch, 5 miles north of 

Millen, Jenkins County. C. W. Cooke and H. K. Shearer.
INTRODUCTION

Banwoell formation

7755. Stevens pottery, Central of Georgia Railway, Baldwin 

County. C. W. Cooke.

FLORIDA

Probably all Ocala limestone

322. Newmansville, Alachua County. L. C. Johnson.

329. Deep wells at Padlock, Suwannee County. L. C. Johnson. 

. 365. Johnsons Sink, Levy County. L. C. Johnson.

8SO. Vicinity of Gainesville and Arredondo, Alachua County. 

L. C. Johnson.

3174. Edgar kaolin works, Edgar, Putnam County, at depth 

of 130 feet.

3629. Ocala limestone, Alachua County. T. W. Vaughan.

3636. Spring Hill schoolhouse, 6 miles north of Alachua. 

Alachua County. T. W. Vaughan.

3684. Pembertou Ferry (now Croom), Hernando County. 

Jos. Willcox (?).

3655. Martin, Marion County. W. I-I. Dall.

3656. Said to have come from Martin. Jos. Willcox (?).

4965. Half a mile southwest of Fort White, Columbia 

County. G. C. Matson.

4974. 6 miles southwest of Chipley, Washington County, 

near Duncan (near 1/48). G. C. Matson.

5030. Ocala, Marion County. G. C. Matson.

6754 (C-67-13). East bank of Choctawhatehee River at 

Alabama-Florida State line (Geneva County, Ala.-Holmes 

County, Fla.). C. W: Cooke.

0768. Wagon bridge over Chipola River east of Marianna, 

lowest bed of Ocala limestone exposed. C. W. Cooke (supplementary collection by C. W. Cooke and Stuart Mossoni).

6799. Alachua Sink, near Gainesville, Alachua County. T. W. 

Vnughan and C. W. Cooke.

OS04. Ocala, Marion County, quarry 1, Florida Lime Co. 

C. W. Cooke.

6805. Oakhurst quarry, 2 miles southeast of Ocala, Marion 

County, quarry 2, Florida Lime Co. C. W. Cooke.

6807. Quarry at Zuber, 6% miles north of Ocala, Marion 

County, quarry 3, Florida Lime Co. C. W. Cooke.

6808. Quarry at Martin, on Atlantic Coast Line Railroad, 9 

miles north of Ocala. C. W. Cooke and H. Gunter.

6810. Cummer Lumber Co.'s pit 10, 1 mile northwest of 

Newberry, Alachua County. O. W. Cooke.

6812. Cummer Lumber Co.'s pit 6, l3/4 miles south of Newberry, Alachua County. C. W. Cooke.

6814. Quarry 1% miles northwest of Newberry, Alachua 

County. C. W. Cooke.

6815. Clark, on Atlantic Coast Line Railroad, 5 miles south 

of High Springs, Alachua County. C. W. Cooke.

6817. Fort White, Columbia County. C. W. Cooke.

6823. East bank of Suwannee River, Suwannee County, near 

Ellaville; no. 2 of section. C. W. Cooke.

7191. Marianna, Jackson County, bed 3 of section at cavern. 

C. W. Cooke and W. C. Mansfield.

7194. Marianna, bed 4 of section at cavern. C. W. Cooke 

and W. C. Mansfield.

7337. Left bank of Suwannee River above bridge of Florida 

Railway, Suwannee County. C. W. Cooke.

7338. Suwannee River at Dowliug Springs, Suwannee County. 

C. W. Cooke.

7341. Left bank of Suwannee River at Brauford, Suwannee 

County. C. W. Cooke.

7345. Suwannee River 2% miles above Branford, Suwanuee 

County. C. W. Cooke.

7348. Suwannee River about iVfe miles above Troy Springs 

and 9 miles aibove Branford,, Suwannee County. O. W. 

Cooke.

7349. Suwannee River about half a mile below Fort McComb, 

Suwanuee County. C. W. Cooke.

7355. 3 miles northwest of Levyville, Levy County. C. W. 

Cooke.

7360. 3]/2 miles south of Floral City, Citrus County. C. W. 

Cooke.

7364. 1% miles west of Croom, Heruando County. C. W. 

Cooke.

7365. Piueola, 5 miles south of Floral City, Citrus County. 

C. W. Cooke.

7366. Old French phosphate mine east of Anthony, Marion 

County. C. W. Cooke.

7367. Willow Sink, about 2 miles west of Chiefland, Levy 

County. C. W. Cooke.

7378. 1^4.miles southwest of Inverness, Citrus County. C. 

W. Cooke.

1/19. Near Blue Springs, Jackson County, 6 miles northeast of Marianna. C. W. Cooke and Julia Gardner.

1/48. Duncan, 7 miles southwest of Chipley, Washington 

County. C. W. Cooke and Julia Gardner.

ALABAMA

Ocala Ivmestoiie

5567. Roadside up from Claibome Ferry, Mouroe County, 

bed S. T. W. Vaughan.

6707. Oven Bluff, Tombigbee River, Clarke County. C. AV. 

Cooke.

6719. Three-quarters of a mile north of Drewry, Mouroe 

County, upper part of cut on Louisville & Nashville Railroad. 

C. W. Cooke.

6720. Three-quarters of a mile north of Drewry, Monroe 

County, lower part of cut on Louisville & Nashville Railroad. 

C. W. Cooke. .

6722. Three-quarters of a niile northwest of Mouroeville, 

Monroe County. C. AV. Cooke.

6724. Road leading up from Claiborne Ferry, Monroe 

County, bed 5 of section, just above " Scutella bed." C. W. 

Cooke.

6725. Road leading up from Claiborne Ferry, Mouroe 

County, bed 7 of section. C. W. Cooke.

6726. Road leading up from Claiborne Ferry, Monroe 

County, bed 8 of section. C. W. Cooke.

6727. Road leading up from Claiborne Ferry, Monroe 

County, bed 10 of section. C. W. Cooke.

6730. Roadside 2% miles west of Monroeville, Mouroe 

County, west of Double Branches. C. W. Cooke.

6738. Sepulga River G'X: miles northeast of Brooklyn, Conecuh County, on J. B. Robinson's land, sec. 13 (?), T. 4 N.; 

R, 13 E., top of bed 3. C. W. Cooke.

6739. 4 miles northeast of Brooklyn, Conecuh County, mouth 

of Robinsons Mill Creek, bed 2, east branch .of Sepulga River. 

C. W. Cooke. " ' : : : . .:.

6740. 4 miles northeast of Brooklyn, Conecuh County, mouth 

of Robinsons Mill Creek, bed-3:.i,C. W. Cooke.

6741. East bank of Sepulga River 3Vj miles northeast of 

Brooklyn, Conecuh County, bed 4. C. W. Cooke.

6742. PowelFs Landing, Sepulga River, Covington County, 

sec. 35, T. 4 N., R. 13 E.,.beds 1 and 2. C. W. Cooke.

6746. East bank of Sepulga River opposite Ward's sawmill, 

sec. 8, T. 3 N., R. 13 E., Conecuh County,, bed 2. C. W. Cooke!

6747. Steamboat Point,, on .west side of Sepulga River, 

Escambia County, near .middle..of sec." 20, T. 3 N., R. 13 E. C. W. Cooke. ' - '" " " ; -'" - '-' -1 -- -"- - 
UPPER EOCENE FORAMINIFERA OF THE SOUTHEASTERN UNITED STATES

6754. East bank of Choctawhatchee River at Alabama-Florida State line, Geneva County, Ala., and Holmes County, Fla. 

C. W. Cooke.

6755. West bank of Choctawhatcb.ee River a quarter of a 

mile above Alabama-Florida State line. C. W. Cooke.

6756. Choctawhatchee River 3% miles below Geneva, Geneva 

County. C. W. Cooke.

7159. Near middle of EV2 sec. 35, T. 8 N., R. 3 E., Clarke 

County, 4 miles southeast of Whatley, " Zeuglodon beds ", No. 

2 of section. C. W. Cooke.

7161. 5 miles southeast of Whatley, Clarke County, east of 

Jones Field Branch, sec. 35, T. 8 N., R. 3 E. C. W. Cooke.

1/7. 1 mile east of Beck, Covington County, NE^ sec. 6, 

T. 3 N., R. 15 E. C. W. Cooke and Julia Gardner.

1/8. Conecuh River at Beck, Covington County. C. W. 

Cooke and Julia Gardner.

1/21. SEVi sec. 22, T. 1 N., R. 18 E., Covington County. C. 

W. Cooke and Julia Gardner.

1/23. About 8 feet above " Scutella bed " on road to Claiborne Ferry, Monroe County. C. W. Cooke.

1/25. South bank of Conecuh River at sharp bend 1 mile 

east of Harts Bridge, SWVi sec. 28, T. 3 N., R. 14 E., Covington 

County. C. W. Cooke and Julia Gardner.

1/26. Bottom bed at Drewry, Monroe County. C. W. Cooke.

1/33. 7% miles northwest of Geneva, Geneva County, 1.2 

miles north of Lytle. C. W. Cooke and Julia Gardner.

1/38. 3% miles north of Grove Hill, Clarke County. C. W. 

Cooke and Julia Gardner.

1/39. West bank of Pea River at Geneva, Geneva County, 

100 yards above mouth of river. C. W. Cooke and Julia Gardner.

1/43. 1% miles southwest of Perdue Hill, Monroe County, 

sec. 7, T. 6 N., R. 6 E. C. W. Cooke and Julia Gardner.

Jackson foi'mation

2637. " Zeuglodon beds", Jackson formation (middle zone), 

Dr. Brown's field, near Cocoa post office, Choctaw County 

(echinoderm horizon). Schuchert and Burns, 1894.

7203. About 3% miles southeast of Cullomburg, Washington 

County, bed 4. C. W. Cooke and W. C. Mansfield.

7220. Half a mile southeast of Melvin, Choctaw County, bed 

3 (top of formation). C. W. Cooke.

7227. Keysers Hill, a quarter of a mile west of Water Valley, Choctaw County, on road to Melvin. W. C. Mansfield.

MISSISSIPPI 

Jackson formation

6457. Chickasawhay River, a quarter of a mile below Mobile & Ohio Railway bridge, 1% miles south of Shubuta, Wayne 

County. C. W. Cooke.

6463. Jackson, hill above pumping station. C. W. Cooke.

6646. Bluff on Chickasawhay River at Hays Chapel, Wayue 

County, a quarter to half a mile above railroad bridge, 1% 

miles south of Shubutn, near base of bluff. C. W. Cooke.

TEXAS 

Jackson group

6143. Near Tordilla Mountain, Karnes County. A. Deussen. 

6148. Bluff on Atascosa Creek near old Whitsett ranch 

house, Atascosa County. A. Deussen. 

6217. 4% miles south of Smiley, Gonzales County. A. Deussen.

 . Upper Eocene near Cheapside, Gonzales County.

 . Haynes well 1, near Burkeville, Newton County.

 . Stovall Creek east of Diboll, Angelina County. (Is 

tliis same as " upper Eocene of Texas from 4 miles east of 

Diboll, Angelina County"?)

 . Tarkiln Creek, half a mile above Neches River, Trinity 

County.

 . Three-quarters of a mile below Robinson's Ferry, Sabine River, Sabine County.

 . Bridge Creek 1% miles above Angelina River, San 

Augustine County.

 . Upper Eocene north of Whitsett, Live Oak County. 

(About same location as 6148 but on other side of county 

line.)

 . Warren well 4, depth 4,250 feet, Texas Exploration Co., 

Hockley, Harris County.

 . Upper Eocene of Ohio and Red River well 2, Tyler 

County, depth 330 feet.

 . Same, depth 400 feet.

 . Same, depth 537 feet.

 . Same, depth 890 feet.

 . Same, depth 1,056 feet.

 . Same, depth 2,330 feet.

Since the completion of the manuscript for this 

paper several works have appeared with descriptions 

and figures of upper Eocene Foraminifera from the 

same general region. In order to give the reader the 

references so that descriptions and figures of these 

other species may be available, a list is here given of 

those species not included in the present paper. These 

species are noted below with references to publications containing the descriptions and to the State from 

which they have been described.

Angulogerina danvillensis Howe and Wallace, Louisiana Dept.

Conservation Geol. Bull. 2, p. 56, pi. 12, fig. 2, 1932. Upper

Eocene, Louisiana. 

Anomalina barrowi Cushman and Ellisor, Contr. Cushman

Lab. Foram. Research, vol. 7, p. 57, pi. 7, figs. lOa-c, 1931.

Upper Eocene, Texas. Ellisor, Am. Assoc. Petroleum Geologists Bull., vol. 17, no. 11, pi. 4, figs. 5a-c, 1933. 

Anomalina danvillensis Howe and Wallace, op. cit, p. 76, pi.

14, figs. 2a, b. Upper Eocene, Louisiana. 

Anomalina, jachsonensis (Cushman and Applin) var. limbosa

Cushman and Ellisor, op. cit., p. 58, pi. 7, figs, lla-c. Upper

Eocene, Texas. Ellisor, op. cit., pi. 5, figs, la-c, 1933. 

Bolivina danvillensis Howe and Wallace, op. cit., p. 56, pi. 11,

figs. 8a, b. Upper Eocene, Louisiana. 

Bolivina gracilis Cushman and Applin var. danvillensis Howe

and Wallace, op. cit., p. 57, pi. 11, fig. 7. Upper Eocene,

Louisiana. 

Bolivina mexicana Cushman var. liorisontalis Cushman.

Howe and Wallace, op. cit, p. 58, pi. 11, fig. 10. Upper

Eocene, Louisiana. 

Bolivina ouacliitaensis Howe and Wallace, op. cit., p. 59, pi. 11,

figs. 9a, b. Upper Eocene, Louisiana. 

Bulimina pyrula D'Orbigny. Howe and Wallace, op. cit., p.

60, pi. 11, fig. 6. Upper Eocene, Louisiana. Ellisor, op. cit.,

pi. 3, fig. 2. 

Buliminella ~basistriata Cushman and Jarvis var. nuda Howe

and Wallace, op. cit., p. 60, pi. 11, fig. 4. Upper Eocene,

Louisiana. 

Buliminella elegantissima, (D'Orbigny). Howe and Wallace,

op. cit., p. 61, pi. 11, fig. 3. Upper Eocene, Louisiana. 

Cancris danvillensis Howe and Wallace, op. cit., p. 67, pi. 13,

figs. 4, 5. Upper Eocene, Louisiana.
INTRODUCTION

Cassidulinoides braziliensis (Cushman). Howe and Wallace,

op. cit., p. 72, pi. 10; figs. 6a, b. Upper Eocene, Louisiana. 

Chilostomella oylindroides Reuss. Howe and Wallace, op.

cit., p. 72, pi. 14, figs, la, b. Upper Eocene, Louisiana. 

OMlostomelloides oviformis (Sherborn and Chapman). Howe

and Wallace, op. cit., p. 73, pi. 15, fig. 5. Upper Eocene,

Louisiana. 

Oilncides danvillensis Howe and Wallace, op. cit., p. 77, pi. 14,

figs. 5a-c. Upper Eocene, Louisiana. 

Cibicides oiiaohitaensis Howe and Wallace, op. cit, p. 78, pi.

14, figs. 6a-c. Upper Eocene, Louisiana. 

Darbyella danvillensis Howe and Wallace, op. cit., p. 24, pi. 4,

figs. 1, 2. Upper Eocene, Louisiana. 

Dcntalina oomiminis D'Orbigny. Howe and Wallace, op. cit.,

pi. 6, fig. 8. Upper Eocene, Louisiana. 

Dentalina communis D'Orbigny var. etnaoiata Reuss. Howe

and Wallace, op. cit, p. 25, pi. 7, fig. 7. Upper Eocene,

Louisiana. 

Dentalina filiformis (D'Orbigny). Howe and Wallace, op. cit.,

p. 25, pi. 6, figs. 2a, b. Upper Eocene, Louisiana. 

Dentalina mexicana (Cushman) var. danvillensis Howe and

Wallace, op. cit., p. 26, pi. 6, fig. 4. Upper Eocene, Louisiana. 

Dentalina sp. a Howe and Wallace, op. cit., p. 20, pi. 6, fig.

1. Upper Eocene, Louisiana. 

Dentalina sp. It Howe and Wallace, op. cit., p. 20, pi. 7, fig.

4. Upper Eocene, Louisiana. 

Dentalina sp. c Howe and Wallace, op. cit., p. 27, pi. 6, fig. 3.

Upper Eocene, Louisiana. 

DisoorMs iarisM Cushman and Ellisor, Contr. Cushman Lab.

Foram. Research, vol. S, p. 43, pi. 6, figs. 6a-c, 1932. Upper

Eocene, Louisiana. Ellisor, op. cit., pi. 7, figs. 6a-c. 

Dhnorphina danvillensis Howe and Wallace, op. cit., p. 44, pi. 8,

figs, la, b. Upper Eocene, Louisiana. 

Dyocibi aides'danvillensis Howe and Wallace, op. cit., p. 78,

pi. 15, figs. 1, 2. Upper Eocene, Louisiana. Ellisor, op. cit.,

pi. 6, fig. 3. 

Ellipsonodosariaf sp. i-Iowe and Wallace, op. cit, p. 67, pi. 11,

fig. 9. Upper Eocene, Louisiana. 

Elpliidium eocenitium, Cushman and Ellisor, Contr. Cushman

Lab. Foram. Research, vol. 7, p. 53, pi. 7, figs. 6a, b, 1931.

Upper Eocene, Texas. Ellisor, op. cit., pi. 2, figs. 13a, b. 

Jillphiditun sp. Howe and Wallace, op. cit., p. 50, pi. 9, fig. 6.

Upper Eocene, Louisiana. 

Epistomina elegans (D'Orbiguy). Howe and Wallace, op. cit.,

p. 68, pi. 13, fig. 2. Upper Eocene, Louisiana. 

Eponides ouachitaensis I-Iowe and Wallace, op. cit., p. 69,

pi. 13, figs. Sa-c. Upper Eocene, Louisiana. 

Glandulina dimorplia (Bornemann). Howe and Wallace, op.

cit., p. 44, pi. 8, fig. 7. Upper Eocene, Louisiana. 

Glandulina laevigata D'Orbigny. Howe and Wallace, op. cit.,

p. 45, pi. 8, figs. 5, 6. Upper Eocene, Louisiana. 

G-loMgei-ina bulloides D'Orbigny. Howe and Wallace, op. cit.,

p. 73, pi. 10, fig. 10. Upper Eocene, Louisiana. 

Globigerina danvillensis Howe and Wallace, op. cit., p. 74,

pi. 10, figs. 9a, b. Upper Eocene, Louisiana. 

G-lobiyerina oiiachitaensis Howe and Wallace, op. cit., p. 74,

pi. 10, figs. 7a, b. Upper Eocene, Louisiana. 

Globigerina sp. a Howe and Wallace, op. cit., p. 75, pi. 10,

figs. Sa, b. Upper Eocene, Louisiana. 

GloMgerina sp. & Howe and Wallace, op. cit., p. 75, pi. 10,

fig. 5. Upper Eocene, Louisiana. 

Glotiulina ampiMa (Jones). Howe and Wallace, op. cit., p. 45,

pi. S, fig. 9. Upper Eocene, Louisiana. 

G'ilnibelina sp. Howe and Wallace, op. cit., p. 53, pi. 9, fig. 8.

Upper Eocene, Louisiana. 

(htttulina austriaca D'Orbigny. Howe and Wallace, op. cit,

p. 47, pi. 8, fig. 3. Upper Eocene, Louisiana.

Guttulina sp. Howe and Wallace, op. cit., p. 49, pi. .7, fig. 6.

Upper Eocene, Louisiana. 

Gyroidina danvillensip Howe and Wallace, op. cit., p. 69, pi. 13,

figs. 3a-c. Upper Eocene, Louisiana. 

Hantkenina br&vispina Cushman. Howe and Wallace, op. cit.,

p. 54, pi. 10, fig. 4. Upper Eocene, Louisiana. 

Hantkenina longispma Cushman. Howe and Wallace, op. cit.,

p. 55, pi. 10, fig. 2. Upper Eocene, Louisiana. 

Hantkenina mecordi Howe and Wallace, op. cit, p. 55, pi. 10,

figs, la, b. Upper Eocene, Louisiana. 

Sopkinsina danvillensis Howe and Wallace, op. cit., p. 62,

pi. 12, figs. la^. Upper Eocene, Louisiana. 

Lagena globosa (Montagu). Howe and Wallace, op. cit., p. 27,

pi. 6, figs. 15, 16. Upper Eocene, Louisiana. 

Lagena hirspida, Reuss. Howe and Wallace, op. cit., p. 28, pi. 6,

fig. 13. Upper Eocene, Louisiana. 

Lagena ouachitaensis Howe and Wallace, op. cit., p. 29, pi. 6,

fig. 9. Upper Eocene, Louisiana. 

Lagena pleniluna Howe and Wallace, op. cit., p. 29, pi. 6, fig. 5.

Upper Eocene, Louisiana. 

Lagena striata (D'Orbigny) var. strumosa Reuss. Howe and

Wallace, op. cit., p. 30, pi. 6, fig. 11. Upper Eocene, Louisiana. 

Lagena sp. a Howe and Wallace, op. cit., p. 30, pi. 6, fig. 12.

Upper Eocene, Louisiana. 

Lagena sp. 6 Howe and Wallace, op. cit., p. 30, pi. 6, fig. 10.

Upper Eocene, Louisiana. 

Lagena sp. c Howe and Wallace, op. cit., p. 31, pi. 6, fig. 6.

Upper Eocene, Louisiana. 

Lagena sp. d Howe and Wallace, op. cit., p. 31, pi. 6, fig. 7.

Upper Eocene, Louisiana. 

Lentioulina sp. Howe and Wallace, op. cit., p. 32, pi. 7, figs.

2a, b. Upper Eocene, Louisiana. 

Margmulina triangularis D'Orbigny var. danvillensis Howe and

Wallace, op. cit., p. 34, pi. 5, figs. 6a, b. Upper Eocene,

Louisiana. 

Margimilina sp. a Howe and Wallace, op. cit., p. 34, pi. 7,

fig. 3. Upper Eocene, Louisiana. 

Marginuliaia sp. b Howe and Wallace, op. cit., p. 35, pi. 3,

fig. 5. Upper Eocene, Louisiana. 

Marginulinaf sp. c Howe and Wallace, op. cit., p. 35, pi. 3,

fig. 6. Upper Eocene, Louisiana. 

Massilina liumblei Cushman and Ellisor, Contr. Cushman Lab.

Foram. Research, vol. 8, p. 40, pi. 6, figs, la-c, 1932. Upper

Eocene, Texas. Ellisor, op. cit., pi. 6, figs. 8a-c. 

Massilina pratti Cushman and Ellisor, Contr. Cushman Lab.

Foram. Research, vol. 7, p. 53, pi. 7, figs. 4a-c, 1931. Upper

Eocene, Texas. Howe and Wallace, op. cit., p. 21, pi. 2,

fig. 5. Ellisor, op. cit., pi. 1, figs. lOa-e. 

Nodosaria longiscata D'Orbigny. Howe and Wallace, op. cit.,

p. 35, pi. 7, fig. 8. Upper Eocene, Louisiana. 

Nonion applini Howe and Wallace, op. cit., p. 51, pi. 9, figs.

4a, b. Upper Eocene, Louisiana. 

Nonion danvillensis Howe and Wallace, op. cit., p. 51, pi. 9,

fig. 3. Upper Eocene, Louisiana.

Nonion hantkeni (Cushman and Applin) var. fayettei Cushman and Ellisor, Contr. Cushman Lab. Foram. Research,

vol. 8, p. 41, pi. 6, figs. 3a, b, 1932. Upper Eocene, Louisiana. Ellisor, op. cit., pi. 7, figs. 9a, b. 

Nonion laevis (D'Orbigny) var. marginatunt Cushman and

Ellisor, Contr. Cushman Lab. Foram. Research, vol. 7, p. 52,

pi. 7, figs. 5a, b, 1931. Upper Eocene, Texas. Ellisor, op.

cit, pi. 2, figs. 8a, b. 

Nonion scaphum (Fichtel and Moll) var. infiatum Cushman

and Ellisor, Contr. Cushman Lab. Foram. Research, vol. 8,

p. 41, pi. 6, figs. 2a, b, 1932. Upper Eocene, Louisiana. 

Ellisor, op. cit., No. 11, pi. 7, figs. 7a, b,
UPPER EOCENE FORAMINIFERA OF THE SOUTHEASTERN UNITED STATES

Nonion sp. Howe and Wallace, op. cit, p. 52, pi. 9, flgs. la, b.

Upper Eocene, Louisiana. 

Nonionella damnllensis Howe and Wallace, op. cit., p. 52, pi. t\

figs. 5a-c. Upper Eocene, Louisiana. 

Nonionella turgida (Williamson). Howe and Wallace, op.

cit, p. 63, pi. 9, figs. 2a-c. Upper Eocene, Louisiana. 

Planularia catahoulaensis Howe and Wallace, op. cit., p. 30,

pi. 3, figs. 3, 4. Upper Eocene, Louisiana. 

Planularia danvillensis Howe and Wallace, op. cit., p. 36, pi. 3,

fig. 1. Upper Eocene, Louisiana. 

Planularia ouachitaensis Howe and Wallace, op. cit., p. 37,

pi. 3, fig. 7. Upper Eocene, Louisiana. Ellisor, op. cit., pi.

6, fig. 9. 

Pulvinulinella danvillensis Howe and Wallace, op. cit., p. 71,

pi. 13, figs. 7a-c. Upper Eocene, Louisiana. 

Pyrgo inornata (D'Orbigny) var. danvillensis Howe and Wallace, op. cit., p. 21, pi. 2, flgs. la, b. Upper Eocene, Louisiana. Ellisor, op. cit., pi. 7, fig. 13, 1933. 

Quinquelqoulina danvillensis Howe and Wallace, op. cit., p. 22.

pi. 2, flgs. 2a-c. Upper Eocene, Louisiana. 

Qiiinquelooulma seminula (Linn£); Howe and Wallace, op.

cit., p. 22, pi. 2, fig. 3. Upper Eocene, Louisiana. 

Robulus clericii (Fornasini). Howe and Wallace, op. cit., p.

38, pi. 15, fig. 3. Upper Eocene, Louisiana. 

Robulus cultratus Montfort. Howe and Wallace, op. cit., p.

38; .pi. 7, fig. 1. Upper Eocene, Louisiana. 

Robulus? danvillensis Howe and Wallace, op. cit., p. 39, pi. 5.

'fig. 7. Upper Eocene, Louisiana.

Robulus mayi Cushman and Parker. Howe and Wallace, op. 

.' cit, p. 40, pi. 2, fig. 7. Upper Eocene, Louisiana. 

Saracenaria danvillensis Howe and Wallace, op. cit., p. 41, pi.

4,' figs. 1' 3a-c> Upper Eocene, Louisiana. 

Saracenaria moresiana Howe and Wallace, op. cit., p. 42, pi. 2,

figs:'8a-c. .Upper Eocene, Louisiana. 

Siffmomorphina semitecta (Reuss). Howe and Wallace, op.

cit, p. 50, pi. 8, figs. 4a-c. Upper Eocene, Louisiana. 

Siphonina carltoni Cushman and Ellisor, Contr. Cushman Lab.

Foram. Research, vol. 8, p. 42, pi. 6, figs. 5a-c, 1932. Upper 

. Eocene, Louisiana. Ellisor, op. cit, pi. 6, flgs. la-c. 

Siphonina danvillensis Howe and Wallace,' op. cit., p. 70.

pi..; 13, figs, la, b. Upper Eocene; Louisiana. 

Textularia broussardi Howe and Wallace, op. cit., p. 18, pi.

1, flgs. 3a, b. Upper Eocene, Louisiana. 

Textularia* danvillensis Howe and Wallace, op. cit., p. 18, pi.

1, flgs. 2, 4. Upper Eocene, Louisiana.

Textularia mississippiensis Cushman var. rhomboidea Cushman and Ellisor, Contr. Cushman Lab. Foram. Research, 

."vol. 7, p. 51, pi. 7,, figs, la, b, 1931. Upper Eocene, Texas. 

Ellisor, op. cit., pl.-l, figs. 2a, b. 

Textularia ouaohitaensis Howe and Wallace, op. cit., p. 20, pi.

1, figs, la, b. Upper Eocene, Louisiana. 

Triloculina rotunda D'Orbigny yar. Howe and Wallace, op.

cit, p. 23, pi. 2, figs. 4a, b. Upper Eocene, Louisiana. 

Ellisor, op. cit., pi. 7, fig. 14, 1933.

Trochammina teasi Cushman and Ellisor, Contr. Cushman Lab. 

: . Foram. Research, vol. 7, p. 52, pi. 7, figs. 3a-c, 1931. Upper 

..; Eocene, Texas. Ellisor, op. cit., pi. 1, figs. 9a-c. 

Tubulogenerina eocenica Cushman and Ellisor, Contr. Cushman Lab. Foram. Research, vol. 8, p. 42, pi. 6, figs. 4a, b,

1932.: :.Upper Eocene, .Texas. Ellisor, op. cit., pi. 6, figs.

4a^b. ..: . /;;' ' . Uviger.iiia:':.danvillensiS''B.ow& and Wallace, op. cit., p. 62,

pi. 12, fig. 5. Upper Eocene, Louisiana.

Uvifferina ,farinosa Hantken. Howe and Wallace, op. cit., 

:; P. 63, ipl. 12, fig. 4. Upper Eocene, Louisiana.

Valvulineria texana Cushman and Ellisor, Contr. Cushman 

Lab. Foram. Research, vol. 7, p. 56, pi. 7, figs. 9a-c, 1931. 

Upper Eocene, Texas. Howe and Wallace, op. cit., p. 70, 

pi. 13, figs. 6a, b. Ellisor, op. cit., pi. 4,, figs. la-c.

Ventilabrella sp. Howe and Wallace, op. cit., p. 54, pi. 9, fig. 

7. Upper Eocene, Louisiana.

Virgulina danvillensis Howe and Wallace, op. cit., p. 65, pL 11 

figs 2a, b. Upper Eocene, Louisiana.

SYSTEMATIC DESCRIPTIONS

Family LITUOLIDAE

Genus HAPLOPHRAGMOIDES Cushman, 1910

Haplophragmoides dibollensis Cushman and Applin

Plate 1, figures la, b

Haplophragmoides dibollensis Cushman and Applin, Am. Assoc. 

Petroleum Geologists Bull., vol. 10, p. 163, pi. 6, figs, 

la, b, 1928.

Test planispiral, much compressed, completely involute, slightly depressed at the umbilical region on 

both sides, periphery rounded, in the later portions 

subacute; chambers rather indistinct, 8 or 9 in the 

last-formed coil; sutures indistinct, not depressed; 

apertural face convex; wall arenaceous but smoothly 

finished. Diameter 0.60 mm.

Originally described from the upper Eocene of 

Ohio and Red River well 2, Tyler County, Tex., 

at a depth of 1,056 feet. This species has later been 

found in several other wells of Texas that reached 

the upper Eocene. It has not been noted in outcrop 

material from the eastern Gulf Coastal Plain.

Genus AMMOBACULITES Cushman, 1910 

Ammobaculites hockleyensis Cushman and Applin

Plate 1, figures 2a, b

Ammobaculites cf. A. foliaceus Brady. Dumble, Am. Assoc.

Petroleum Geologists Bull., vol. 8, p. 443, 1924. 

Ammobaculites hockleyensis Cushman and Applin, Am. Assoc.

Petroleum Geologists Bull., vol. 10, p. 163, pi. 6, figs.

2a, b, 1926. 

Ellisor, Am. Assoc. Petroleum Geologists Bull., vol. 17,

no. 11, pi. 7, fig. 11, 1933.

Test nearly twice as long as broad, very much compressed, involute, early portion close coiled, later 1 

or 2 chambers tending to become uncoiled, periphery 

rounded; chambers rather indistinct; sutures indistinct; wall coarsely arenaceous, of angular black sand 

grains. Length 1.00 mm; breadth 0.65 mm; thickness 

0.10 mm.

The type of this species came from a sample from 

Ohio and Red River well 2, Tyler County, Tex., at a 

depth of 400 feet. The species has occurred in several 

well sections from the Texas upper Eocene but is not 

recorded from the eastern Gulf Coastal Plain.
TEXTULAKIDAE

Family TEXTULABJIDAE

Genus TEXTULABIA Defiance, 1824

Textularia mississippiensis Cushman

Plate 1, figures 3, 4

Textularia mississippieiisis Cushinan, U.S. Geol. Survey Prof.

Paper 129, pp. 90, 125, pi. 14, fig. 4, 1922; Prof.'Paper

133, p. 17, 1923. 

Cushinan and Applin, Am. Assoc. Petroleum Geologists

Bull., vol. 10, p. 166, pi. 6, figs. 10, 11, 1926. 

Gushman and Thomas, Jour. Paleontology, vol. 3, p. 177,

pi. 23, figs, la, b, 1929. 

Cushman, Contr. Cushman Lab. Foram. Research, vol. 5.

p. 79, pi. 12, fig. 5, 1929. 

Howe and Wallace, Louisiana Dept. Conservation Geol.

Bull. 2, p. 19, pi. 1, figs, 7a, b, 1932.

Test elongate, fairly broad, thickest in the middle, 

thence thinning toward the periphery, in end view 

biconvex, central portion curved; chambers low and 

broad, especially in the early stages, becoming somewhat higher in the adult; sutures covered by a coarsely 

arenaceous layer, meeting at the center and at the 

periphery, having the central portion of each chamber uncovered; periphery irregular, not definitely or 

regularly spinose, chamber walls smooth and finely 

perforate. Length 0.40-0.75 mm.

This species is known from the upper Eocene of 

Texas, where it is widely distributed in both outcrop 

tind well samples, and is common in the Alazan of 

Mexico, with a few records from the region east of 

Texas.

Textularia mississippiensis, which is found in most 

of the formations of the Vicksburg group (Oligocene) 

of the Gulf Coastal Plain of the United States, occurs 

at a very few of the U.S.G.S. stations in the upper 

Eocene.

Jackson formation: 6457, 1% miles south of Shubuta, Miss.; 

6472, 6473, Yazoo City, Miss.; three-fourths mile below Robinson's Ferry, on Sabiue River, Sabine County, Tex.; 4 miles 

east of Diboll, Angelina County, Tex.; Bridge Creek, 1% miles 

above Angelina River, San Augustine County, Tex.

Textularia mississippiensis Cushman var. alabamensis 

Cushman

Plate 1, figures 5, G

Textularia, wrississi-ppionsis Cuslnnun var. alabamcnsis Cushinan, U.S. Geol. Survey Prof. Paper 133, p. 17, pi. 1, 

fig. 4, 1923.

Ellisor, Am. Assoc. Petroleum Geologists Bull., vol. 17, 

no. 11, pi. 1, fig. 6, 1933.

Variety differing from the typical species in the 

more elongate form, somewhat thicker test, especially 

in the center, the less well-defined peripheral carina, 

and the sutures excavated instead of being covered by 

an arenaceous layer.

This 'variety is especially characteristic of the 

Marianna limestone of the Vicksburg group (Oligocene) . It is very much more abundant than the typical form in the upper Eocene of the Coastal Plain of 

the United States.

Cooper marl: 6319, Ingleside, S.C.; 6321, Balclock, S.C.

Ocala limestone: 7108, Flint River at Seaboard Air Line 

Ry. bridge, Sumter County, Ga.; 7237, half a mile east of 

Huguenen Ferry, Crisp County, Ga.; 6805, Oakhurst quarry, 

Ocala, Fla.; 7194, Marianna, Fla.; 6706, Oven Bluff, Tombigbee River, Ala.; 6719, three-fourths mile north of Drewry, 

Ala.; 6722, three-fourths mile northwest of Monroeville, Ala.; 

6724, 6725, 6726, Claiborne, Ala.; 6730, Double Branches 

Creek, Ala.; 6746, Ward's sawmill, Sepulga River, Ala.; 7159, 

4 miles southeast of Whatley, Ala.; 7161, 5 miles southeast 

of Whatley, Ala.; 1/8, Beck, Ala.; 1/21, Covington County, 

Ala.; 1/26, Drewry, Ala.; 1/38, 3% miles north of Grove Hill, 

Ala.; 1/43, 1% miles southwest of Perdue Hill, Ala.

Jackson formation: 7227, Water Valley, Ala.; 6463, Jackson, 

Miss.

Textularia ocalana Cushman

Plate 1, figures 7a, b

Textularia ocalana Cushinan, Contr. Cushman Lab. Foram. 

Research, vol. 2, p. 30, pi. 4, figs. 3a, b, 1926.

Test very much compressed, sides nearly flat, apical 

end bluntly pointed, apertural end broadly rounded, 

periphery even, subacute; chambers .few, low, and 

broad, not inflated; sutures very slightly depressed, 

rather indistinct; wall finely arenaceous, smoothly finished. Maximum length 0.70 mm.

This peculiar-shaped species is very rare, occurring 

only in the typical Ocala limestone at a few stations. 

Its. very broad form, with the sides very much compressed and but very slightly convex, distinguishes it 

from any other of the Vicksburg or Jackson species". 

It probably occurs at more stations, but the finer 

material of this phase of the Ocala is often not well 

preserved.

Ocala limestone: 6807, Zuber, Fla.; 6808, Martin, Fla.; 6812, 

1% miles south of Newberry, Fla., the type locality.

Textularia recta Cushman

Plate 1, figures 8, 9

Textularia recta Cushman, U.S. Geol. Survey Prof. Paper 

133, p. 17, pi. 1, fig. 2, 1923.

Test elongate, slightly compressed, early portion 

rapidly increasing in diameter, later portion in the. 

adult with the sides parallel for a large part of the 

test; chambers numerous; sutures distinct; wall thick, 

covered with agglutinated calcareous grains but when 

worn showing a coarsely perforated undertest; apertural end obliquely truncate; aperture in a deep depression, at the base of the last-formed chamber. 

Maximum length of specimens 1.25 mm.

Specimens similar to Textularia recta, which was 

described, from the Red Bluff clay of the Vicksburg 

group, occur in the upper Eocene at the following 

U.S.G.S. stations:

Ocala limestone: 6725, 6726, Claiborne, Ala.; 6754, Choctawhatchee. River at State line, Ala.rFla:; ..6756, Choctaw-
8 UPPER EOCENE FORAMINIFERA OP THE SOUTHEASTERN UNITED STATES

hatchee River 3% miles below Geneva, Ala.; 1/8, Beck, Ala.; 

1/39, Pea River at Geneva, Ala.

Jackson formation: 6463, Jackson, Miss.; 6471, Garlands 

Creek, Miss.

Textularia subhauerii Cushman

Plate 1, figure 10

Textularia suWiauerii Cushman, U.S. Geol. Survey Prof. Paper 

129, pp. 89, 126, pi. 14, figs. 2a, b, 1922; Prof. Paper 133, 

p. 16, 1923.

Test large, stout, elongate, early portion rapidly 

increasing in width with each newly added chamber, 

later adult portion with the sides nearly parallel, 

slightly lobulate; periphery rounded, but the median 

portion nearly flat; chambers 18 to 20, increasing in 

height as added, those of the later portion nearly as 

high as broad; sutures usually rather indistinct; wall 

coarsely arenaceous, aperture at the base of the inner 

margin of the chamber. Maximum length 2.00 mm.

Specimens very similar to this species, which was 

described from the Oligocene Byram marl, were found 

in the upper Eocene at the following U.S.G.S. 

stations :

Cooper marl: 6319, Ingleside, S.C.; 6321, Baldock, S.C.

Ocala limestone: 7108, Flint River at Seaboard Air Line 

Railway bridge, Sumter County, Ga.; 7189, Rich Hill, Ga.; 

7704, 3 miles west of Tarversville, Ga.; 6768, Marianna, Fla.; 

6738, 6% miles north of Brooklyn, Ala.; 6740, mouth of Robinsons Mill Creek, Ala.; 6741, 3y2 miles northeast of Brooklyn, 

Ala.; 6754, Choctawhatchee River at State line, Ala.-Fla.: 

1/7, 1 mile east of Beck, Ala.; 1/8, Beck, Ala.; 1/25, 1 mile 

east of Harts Bridge, Ala.; 1/48, Duncan, Fla.

Jackson formation: 6457, 1% miles south of Shubuta, Miss.

Textularia adalta Cushman

Plate 1, figures 11, 12

Textularia adalta Cushman, Contr. Cushman Lab. Foram.

Research, vol. 2, p. 29, pi. 4, figs. 2a, b, 1926. 

Ellisor, Am. Assoc. Petroleum Geologists Bull., vol. 17, 

no. 11, pi. 6, fig. 2, 1933.

Test elongate, slender, early portion tapering and 

compressed, adult portion thicker and with the sides 

nearly parallel, periphery subacute except in the last 

few chambers, which are rounded; chambers numerous, 

.the last 5 or 6 making up half the test, earlier ones 

indistinct, low and broad, later ones more inflated, 

higher; sutures distinct, especially in the later portion, 

where they are somewhat depressed, usually oblique; 

wall finely arenaceous, only slightly roughened; aperture a high, arched opening in the central part of the 

base of the apertural face. Maximum length 2.00 mm.

Ocala limestone: 6741, 3% miles northeast of Brooklyn, Ala., 

type and only locality (abundant).

Textularia dibollensis Cushman and Applin

Plate 1, figures 13-16

Textularia dibollensis Dumble (nomen nudum), Am. Assoc. 

Petroleum Geologists Bull., vol. 8, p. 443, 1924.

Textularia dibollensis Cushmau and Applin, Am. Assoc. Petroleum Geologists Bull., vol. 10, p. 165, pi. 6, figs. 12-14, 

1926.

Ellisor, Am. Assoc. Petroleum Geologists Bull., vol. 17, no. 

11, pi. 1, fig. 4, 1933.

Test small, short, and broad, moderately compressed, margin subacute, initial end rounded; apertural end broadly truncate; chambers few, rather indistinct, rapidly increasing in breadth as added, 

becoming high in the adult, with the sides nearly 

parallel, the last four usually making up at least half 

of the test; sutures indistinct, not depressed, at right 

angles to the peripheral margin; wall finely arenaceous; aperture an arched opening, low and broad, at 

the inner margin of the last-formed chamber, the 

apertural face of the chamber evenly rounded. Maximum length 0.80 mm, usually much shorter; breadth 

0.30 mm.

This is a well-characterized small species not easily 

confused with any other of the Coastal Plain Oligocene or upper Eocene species. It was originally described from the upper Eocene outcropping 4 miles 

cast of Diboll, Angelina County, Tex. It is known 

from the upper Eocene of samples from several wells 

in the eastern part of the Gulf Coastal Plain of 

Texas.

Ocala limestone: 3629, Alachua County, Fla.; 672:4, Claiborne, Ala.; 6754, Choctawhatchee River at State line, Ala.- 

Fla. ; 6756, Choctawhatchee River 3^ miles below Geneva, 

Ala.; 1/8, Beck, Ala.: 1/39, Pea River at Geneva, Ala.

Textularia dibollensis Cushman and Applin var. humble! 

Cushman and Applin

Plate 1, figures 17a, b

Textularia dibollensis var. humblei Dumble (nornen nudurn), 

Am. Assoc. Petroleum Geologists Bull., vol. 8, p. 443, 

1924.

Textularia dibollensis Cushman and Applin var. humblei Cushman and Applin, Am. Assoc. Petroleum Geologists Bull., 

vol. 10, p. 165, pi. 6, fig. 9, 1926.

Variety differing from the typical in the larger size, 

more elongate form, more tapering, the greatest width 

being near the apertural end; the sutures much more 

distinct and depressed, and the periphery somewhat 

more acute. Length 1.60 mm; breadth 1.00 mm.

This variety was described from Haynes well 1, 

near Burkeville, Newton County, Tex., at a depth of 

3,175 to 3,270 feet.

Textularia distortio Cushman and Applin

Plate 1, figures 18, 19

Textulai-ia liockleyensis var. distortio Dumble (noinen 

nudum), Am. Assoc. Petroleum Geologists Bull., vol. 8, 

p. 443, 1924.

Textularia, distortio Cushman and Applin, Am. Assoc. Petroleum Geologists Bull., vol. 10, p. 164, pi. 6, figs. 7, 8, 

1926.
VERNEUILINIDAE 9

Test irregular in shape, variously distorted, much 

compressed, thickest in the central portion, periphery 

thin but rounded; chambers numerous, low, and 

broad; sutures very distinctly excavated, especially 

toward the median portion of each chamber; wall 

arenaceous, somewhat roughened; aperture curved 

in end view, the aperture making a very strong crescent. Length 1.00 mm or slightly more.

This peculiar-formed species was described from 

samples from Ohio and Eed Eiver well 2, Tyler 

County, Tex., at a depth of 890 feet.

Textularia hockleyensis Cushman and Applin

Plate 1, figures 20, 21

Textularia hockleycnsis Dumble (nomen nucluin), Am. Assoc.

Petroleum Geologists Bull., vol. 8, p. 443, 1924. 

Textularia hockleycnsis Cushman and Applin. Am. Assoc.

Petroleum Geologists Bull., vol. 10, p. 164, pi. 6, figs. 3-6,

1926. 

Ellisor, Am. Assoc. Petroleum Geologists Bull., vol. 17,

no. 11, pi. 1, tig. 7, 1933. 

Textulw-ia liocldeyensis Cushman and Applin var.? Howe

and Wallace, Louisiana Dept. Conservation Geol. Bull.

2, p. 19, pi. 1, figs. 5a, b, 1932.

Test comparatively large, tapering, compressed, the 

central portion thickest, thence with a depressed area 

between the center and the periphery, periphery thin 

but rounded; chambers numerous, distinct; sutures 

distinct and toward the central portion often somewhat limbate, strongly curved, especially toward the 

periphery; wall arenaceous but smoothly finished, in 

end view rhomboid; the aperture much curved, low. 

Length up to 3.00 mm; breadth up to 1.25 mm.

Type specimens came from Bridge Creek, iy2 miles 

above Angelina River, San Augustine County, Tex. 

Outside of the Texas region the only specimens seen 

are those noted below.

Barnwell formation: 7755, Stevens Pottery, Ga.

There are probably several other species of Textularia in the upper Eocene of the Coastal Plain. In 

the Ocala limestone specimens are frequently found, 

but the nature of the friable material is such that the 

surface characters are not well preserved. If the same 

fauna can be found in a less friable and better-preserved condition, these species can be worked out in 

specific details.

Genus VULVULINA D'Orbigny, 1826 

Vulvulina advena Cushman

Plate 2, figures la, b

Vuloulvna ad-vena Cushman, Contr. Cushman Lab. Fora in. Research, vol. 2, p. 32, pi. 4, figs. 9a, b, 1926; vol. 8, p. 

84, pi. 10, fig. 14, 1932.

Cole, Bull. Am. Paleontology, vol. 14, no. 53, p. 206 (6), 

pi. 1, fig. 24; pi. 3, fig. 17, 1928.

Test small, thin, much compressed, periphery acute 

but not keeled, early chambers alternating, later ones 

(as many as 5) uniserial; chambers rather high, gently 

sloping; sutures of the biserial portion flush with the 

surface, the whole early portion smooth, in the later 

portion the sutures slightly depressed; wall smoothly 

finished, each angle of the chambers with a short, 

spinose projection, even those of the uniserial portion; 

aperture elongate, terminal. Maximum length 1.25 

mm.

This is a very distinct species, having a very white 

test, smooth and shining, the material being very fine 

in texture with much cement.

Jackson formation: 7203, 3% miles southeast of Cullomburg, Ala., the type locality.

Ocala limestone: 1/43, 1% miles southwest of Perdue Hill, 

Ala.

Family VERNEUILINIDAE

Genus VERNEUILINA D'Orbigny, 1840 

Verneuilina propinqua H. B. Brady .

Plate 2, figures 3a, b

Vemetiilina propinqua H. B. Brady, Challenger Kept., Zoology,

vol. 9, p. 387, pi. 47, figs. 8-12 (not 13, 14), 1884. 

Goes, K. svensk. vet. akad. Handl., vol. 25, no. 9, p. 33,

pi. 7, figs. 264-266, 1895. 

Flint, U.S. Nat. Mus. Kept, for 1897, p. 285, pi. 31, fig. 2,

1899. 

Cushman, U.S. Nat. Mus. Bull. 104, pt. 3, p. 56, pi. 9,

figs. 10, 11, 1922.

Test free, pyramidal, triserial, the apical end bluntly 

rounded; chambers well inflated but closely set; wall 

arenaceous, the surface nearly smooth; aperture elongate, at the base of the inner margin of the chamber. 

Length 1.20 mm; breadth 0.75 mm.

Cooper marl: 11857,1 mile south of Moncks Corner, Berkeley 

County, S.C.

The one Cretaceous record for this species is unaccompanied by a figure, and the range is therefore given 

as from Eocene to Kecent.

Genus GAUDRYINA D'Orbigny, 1839. 

Gaudryina jacksonensis Cushman

Plate 2, figures 4-6

Gaudryina jacksonensis Cushman, Contr. Cushman Lab. 

Forarn. Research, vol. 2, p. 33, pi. 5, figs, la, b, 1926.

Test large, elongate, irregularly triangular in section, the angles subacute, triserial portion short, biserial portion triangular, angles subacute, almost carinate; chambers distinct, very slightly inflated; sutures 

distinct, slightly depressed; wall composed of fine 

sand grains with a large amount of cement, surface 

smoothly finished; aperture semicircular, in a reentrant at the base of the apertural face of the lastformed chamber. Maximum length 2.00 mm.
10 UPPER EOCENE FORAMINIFERA OF THE SOUTHEASTERN UNITED STATES

This- is a striking species of the deeper-water sediments of the upper Eocene. It is apparently the direct ancestral form of the recent species 6r. atlantica 

(Bailey), which is very abundant off the eastern coast 

of the United States.

It is a dominant species in the Albatross dredgings 

at depths ranging from 67 to 210 fathoms. The recent species has carried the development further in 

the loss of the angles in the last-formed chambers, 

the greater sharpness of the angles of earlier chambers, and the reduction of the early triserial stage. 

(r. atlantica, is also a larger species.

Gaudryina jacksonensis also occurs in the Alazan 

shale of Mexico.

Cooper marl: 6319, Ingleside, S.C., the type locality.

Ocala limestone: 1/38, sy^ miles north of Grove Hill, Ala.; 

1/43, l]/2 miles southwest of Perdue Hill, Ala.

Jackson formation : 7203, 3% miles southeast of Cullomburg, 

Ala.; 7220, half a mile southeast of Melvin, Ala.; 6457, 1% 

miles south of Shubuta, Miss.

Gaudryina gardnerae Cushman

Plate 2, figures 7a, b

Gaudryina gardnerae Cushman, Contr. Cushman Lab. Foram. 

Kesearch, vol. 2, p. 33, pi. 5, figs. 2a, b, 1926.

Test elongate, early portion triserial and forming a 

distinctly triangular test in section, the angles somewhat rounded, sides flattened or very slightly convex,, 

later portion biserial, with straight, nearly parallel 

sides, somewhat polygonal in section; chambers rather 

indistinct, especially in the earlier portion, in the 

later portion more distinct and the sutures somewhat 

depressed; wall rather coarsely arenaceous; aperture 

rounded, deep, at the inner border of the last-formed 

chamber. Length 0.85 mm.

This resembles G. triangularis Cushman in some 

respects, and the Oligocene specimens referred to that 

species may belong here.

Ocala limestone: 6785, three-fourths mile south of Herlongs, 

Fla.; 1/43, lJ/£ miles southwest of Perdue Hill, Ala., the type 

locality.

Gaudryina subquadrata Cushman

Plate 2, figures Sa-c

Oaudryina subquadrata Cushman, Contr. Cushman Lab. 

Foi-ain. Research, vol. 9, p. 2, pi. 1, figs, la-c, 1933.

Test elongate, slender, the biserial portion compressed, concave at the broader faces, margins 

rounded, truncate, early triserial portion greatly reduced; chambers of the biserial portion distinct, 7 or 

8 pairs; sutures strongly oblique, slightly depressed; 

wall arenaceous, exterior somewhat roughened; aperture at the base of the last-formed chamber at the 

inner edge, small. Length 1.00 mm; breadth 0.30 

mm; thickness 0.20 mm.

Holotype (no. 371526, U.S.N.M.) from Cooper 

marl, station 11861, Berkeley County, S.C.

This is a very elongate, tapering species, the triserial portion being greatly reduced. The broader 

faces are very distinctly concave.

Cooper marl: 11861, steep bluff on west side of Biggin 

Creek, Berkeley County, S.C.

Genus PSEUDOCLAVULINA Cushman, 1934 

Pseudoclavulina sp.?

Plate 2, figures lOa, b

Clavulina nodosaria D'Orbigny var. novangliae Cushman, U.S. 

Nat. Mus. Bull. 104, pt. 3, p. 82, pi. 15, figs. 3-5, 1922.

Test elongate, slender, slightly tapering, the early 

triserial portion much reduced, three-sided, the angles 

rounded, later portion circular in transverse section, 

increasing in size as the chambers are added; chambers 

very few in the triserial portion, 5 to 8 in the uniserial 

portion, more or less indistinct except the last-formed 

ones, slightly inflated; sutures indistinct except near 

the apertural end, where they are depressed; wall arenaceous, slightly roughened; aperture single, circular, 

terminal, often with a slight neck; color light gray. 

Length of figured specimen 0.90 mm; diameter 

0.30 mm.

The specimens seem to belong to Pseudoclavuttna 

but are not numerous enough to warrant a specific 

name.

Cooper marl: 11857,1 mile south of Moncks Corner, Berkeley 

County, S.C.

Family VAIVULINIDAE 

Genus VALVULINA D'Orbigny, 1826 

Valvulina ocalana Cushman

Plate 2, figures 11, 12

Valvulina, ocalana Cushman, Contr. Cushman Lab. Foram. 

Research, vol. 2, p. 34, pi. 5, figs. 4 a, b, 1926.

Test elongate, early portion triangular in section, 

the chambers closely set, later and larger portion 

loosely coiled; chambers numerous, those of the early 

triserial portion indistinct, later portion with the 

chambers inflated and very distinct; sutures indistinct 

in the early portion, depressed and distinct in the later 

portion; wall arenaceous but smoothly finished when 

the specimens are well preserved; aperture rounded, 

m a deep reentrant of the apertural face, with an 

inward-projecting toothlike plate above the opening. 

Maximum length 1.50 mm.

Valvulwa ocalana is a very characteristic species of 

the shallow-water phase of the upper Eocene of warm 

waters as developed in the Ocala limestone of Florida. 

Such deposits are not usually very well preserved as 

to their smaller fossils, but V, ocalana is a well-char-
MILIOLIDAE 11

acterized species, recognizable even when the preservation would make impossible the specific identification 

of many other species. It is related to such species 

as Valvulma triangularis D'Orbigny, of the Eocene 

of the Paris Basin.

Valwulina ocalana is the direct ancestral form of a 

species now very abundant in shallow, warm waters 

of coral-reef conditions in the general West Indian 

region, where it occurs in very shallow water of reefs 

of southern Florida, the Bahamas, Jamaica, etc. The 

Kecent species V. oviedoiana D'Orbigny is a shorter, 

stouter form but with the same characteristic apertural features.

Ocula limestone: 3629, Alachua County, Fla.; 6790, Alachua, 

Fla.; 6S05, Oakhurst quarry, Ocala, Fla., the type locality; 

6807, Zuber, Fla.; 6808, Martin, Fla.; 7345, Suwannee River 

2% miles above Branford, Fla.; 6823, Ellaville, Fla.; 7360, 

3% miles south of Floral City, Fla.; 7364, 1% miles west of 

Croom, Fla.; 7366, Anthony, Fla.

Genus TEXTULARIELLA Cushman, 1927 

Textulariella barrettii (Jones and Parker)

Plate 2, figures 2a, b

Tcmtularia batrettti Jones and Parker, British Assoc. Kept.

Newcastle meeting, pp. 50, 105. 1860; Soc. malacol.

Belgique Annales, vol. 11, p. 99, woodcut, 1876. 

H. B. Brady, Challenger Kept., Zoology, vol. 9, p. 367,

pi. 44, flgs. 6-8, 1884.

Woodward, New York Micr. Soc. Jour., 1885, p. 149. 

Flint, U.S. Nat. Mus. Kept, for 1897, p. 285, pi. 30, fig.

2, 1899. 

Cushman, Carnegie Inst. Washington Pub. 291, p. 31,

pi. 6, flgs. 5-7, 1919; U.S. Nat. Mus. Bull. 104, pt. 3, p.

20, pi. 3, figs. 3-6, 1922. 

Textulariella barrettii Cushman, Contr. Cushman Lab. Foram.

Research, vol. 3, p. 24, pi. 5, fig. 3, 1927; Special Pub.

1, p. 115, pi. 11, flgs. 6-8; pi. 13, fig. 3, 1928. 

TctBlularia oorvica Goes (not D'Orbigny), Harvard Coll. Mus.

Comp. Zoology Bull., vol. 29, p. 43, 1896.

Test tapering, about twice as long as broad, very 

slightly compressed, broadest near the apertural end, 

the apical end bluntly pointed, later portion of the 

test often with nearly straight sides; chambers distinct, numerous, labyrinthic; sutures very , clearly 

marked, not depressed; wall finely arenaceous with 

an abundance of cement, very smoothly finished; aperture a narrow slit at the base of the inner margin of 

the last-formed chamber, the sides of the chamber 

slightly projecting beyond it on each side, sometimes 

subdivided into one or more openings. Length of 

figured specimen 0.75 mm; diameter 0.65 mm.

The figured specimen is evidently not an adult. 

The species is known from the Miocene of Jamaica 

and is a common one in the West Indian region and 

off the southeastern coast of the United States in 

100 fathoms or more. It is interesting to see that 

this species has persisted in this region since the upper Eocene with but little change.

Cooper marl: 11861, steep bluff on west side of Biggen 

Creek, Berkeley County, S.C.

Genus IIEBUSELLA Cushman, 1933 

Liebusella byramensis (Cushman) var. turgida (Cushman)

Plate 2, figure 9

ClavuUna byramensis Cushman var. turgida Cushman, U.S. 

Geol. Survey Prof. Paper 133, p. 22, pi. 2, figs. 4, 5, 1923.

Test elongate, subcylindrical, early portion triserial, 

forming but a small portion of the test, later ones uniserial, both portions rounded in section; sutures usually indistinct or very slightly depressed; wall coarsely arenaceous but smoothly finished; aperture terminal, rounded, in well-preserved specimens with a 

slightly projecting, tapering neck. Maximum length 

3.00 mm.

This variety becomes common in the lower portions 

of the Oligocene, occurring in the Glendon limestone, 

the typical Marianna limestone, and abundantly in the 

Red Bluff clay. It is somewhat larger and stouter 

than the typical form of the Byram marl. This form 

is related to Liebusella cylindrica, described from the 

Eocene of Europe by Hantken, and L. eocaena (Giimb'el). Liebusella byramensis is probably the direct ancestor of L. flwtiana. Cushman, now living off the 

southeastern coast of the United States, the depths 

Avhere it is common ranging from 68 to 210 fathoms.

Liebusella byramensis var. turgida is characteristic 

of the deeper-water deposits of the upper Eocene of 

the Coastal Plain of the United States:

Cooper marl: 6319, Ingleside, S.C.

Ocala limestone: 7194, Marianna, Fla.; 1/21, Covington 

County, Ala.; 1/26, Drewry, Ala.; 1/43, 1% miles southwest 

of Perdue Hill, Ala.

Jackson formation: 7203, 3% miles southeast of Cullomburg, 

Ala.; 6457, 1% miles south of Shubuta, Miss.

Family MILIOLIDAE

Genus QUINftTTELOCULINA D'Orbigny, 1826 

Quinqueloculina laevigata D'Orbigny

Plate 2, figures 13-15

Quinqueloculinat laeviyata D'Orbigny, Annales sci. nat, vol. 7, 

p. 301, no. 6, 1826; in Barker-Webb and Berthelot, 

Histoire naturelle des iles Canaries, vol. 2, pt. 2, 

Foraminiferes, p. 143, pi. 3, figs. 31-33, 1839. 

Terquem, Soc. geol. France Mem., ser. 3, vol. 2, p. 173, 

pi. 18(26), figs. 14, 15, 18S2.

Test nearly twice as long as broad, slightly compressed, periphery rounded; chambers distinct, somewhat inflated, of nearly uniform diameter, the ends 

only slightly extended, apertural end slightly exserted; sutures distinct, slightly depressed; wall 

smooth; aperture nearly circular, terminal, with a 

simple tooth but usually without a lip. Length 0.35 

mm; breadth 0.20 mm; thickness 0.12 mm.

Jackson formation: 6463, Jackson, Miss.
UPPER EOCENE FORAMINlFERA OP THE SOUTHEASTERN UNITED STATES

Quinqueloculina longirostra D'Orbigny

Plate 2, figures 16a-c

Quinqueloculina longirostra D'Orbigny, Annales sci. nat., vol. 

7, p. 303, no. 46, 1826; Foraminiferes fossiles du bassin 

tertiaire de Vienne, p. 291, pi. 18, figs. 25-27, 1846.

Test nearly twice as long as broad, compressed, 

periphery subacute; chambers distinct, not inflated, 

gradually compressed to the subacute, almost keeled 

periphery, sides on the inner part convex, thence becoming slightly concave toward the periphery, each 

extending well beyond the previous ones at both ends, 

apertural end exserted so that the aperture is terminal; sutures distinct, slightly depressed; wall 

smooth; aperture circular, terminal, with a slight lip 

and a small, simple tooth. Length 0.90 mm; breadth 

0.45 mm; thickness 0.25 mm.

Jackson formation: 6463, Jackson, Miss.

Quinqueloculina anguina Terquem

Plate 2, figures 18, 19

Quinqueloculina anffuina Terquem, Soc. g£ol. France Mem., 

 ser. 3, vol. 1, p. 78, pi. 9 (14), figs. 20a-c, 1878; vol. 2, 

p. 180, pi. 19 (27), figs. 20a-c, 1882.

Test about 21/2 times as long as broad, compressed, 

periphery broadly rounded; chambers distinct, inflated, at the base somewhat enlarged and extending 

well beyond the previous chamber, at the apertural 

end narrowed and extended into a short cylindrical 

neck; sutures distinct, much depressed; wall smooth; 

aperture circular, terminal with a distinct lip and a 

small, narrow, simple tooth. Length 0.35 mm; 

breadth 0.15 mm; thickness 0.08 mm.

Our specimens agree very closely with specimens 

from the Eocene of the Paris Basin.

Jackson formation: 6463, Jackson, Miss.

Quinqueloculina hauerina D'Orbigny?

Plate 2, figures 17a-c

There are some specimens of a small quinqueloculine form which resemble somewhat the species described by D'Orbigny from the Miocene of the Vienna 

Basin. They are figured here for reference until 

larger series of well-preserved specimens are available for study.

Jackson formation: 6463, Jackson, Miss.

Quinqueloculina sp.?

Plate 4, figures 5, 6

Quinqueloculina sp.? Cushman and Applin, Am. Assoc. Petroleum Geologists Bull., vol. 10, p. 184, pi. 10, figs. 20, 21, 

1926.

Although not enough material is available for specific determination, these specimens from the upper

Eocene of Stovall Creek east of Diboll, Angelina 

County, Tex., are figured, that the record of this form 

may be available.

Quinqueloculina sp.?

Plate 4, figures 7, 8

Quinqiwloculina sp.? Cushman and Applin, Am. Assoc. Petroleum Geologists Bull., vol. 10, p. 184, pi. 10, figs. 22, 

23, 1926.

This species seems to be distinct from the preceding, especially in its apertural characters, the aperture 

being much smaller and the test itself comparatively 

broader. Without more material, it seems unwise to 

give this .a specific name.

The specimens came from Stovall Creek east oi 

Diboll, Angelina County, Tex.

Quinqueloculina sp.?

Plate 4, figures 9, 10

Quinqueloculina sp.? Cushman and Applin, Am. Assoc. Petroleum Geologists Bull., vol. 10, p. 184, pi. 10, fig. 24, 

1926.

. The peculiar form here figured occurs with the two 

preceding species on Stovall Creek east of Diboll, 

Angelina County, Tex. Much more material is 

needed before a specific determination can be made.

Genus MILIOLA Lamarck, 1801

By elimination of other species placed by Lamarck 

under this genus, the type of Miliola becomes Miliola 

saxorum Lamarck. The original figures given by 

Lamarck are crude and leave much to the imagination 

as to the exact character of the type. The model of 

D'Orbigny, no. 33, made in 1826, gives a very clear 

idea of what was probably Lamarck's species. It is 

the large common species of the Calcaire grossier of 

the Paris Basin. The aperture is cribrate, and this 

generic name Miliola may therefore be used for M. 

saxorum and its allies without disturbing the later 

Quinqueloculina of D'Orbigny.

Miliola saxorum Lamarck

Plate 3, figures 1-3

Miliola, (Miliolites) saxorum Lamarck, Annales du Museum, 

vol. 5, p. 352, no. 5, 1804; vol. 9, pi. 17, figs. 2a, b, 1807. 

Defrance, Dictionnaire des sciences naturelles, vol. 31, 

p. 69, 1824; Atlas de conchyliologie, pi. 15, fig. 1.

Quinqueloculina saxorum D'Orbigny, Annales sci. nat., vol. 7, 

p. 301, no. 1, pi. 16, figs. 10-14, 1826.

Test large, elongate, fusiform, quinqueloculine; 

chambers very numerous, distinct, the periphery flattened ; sutures distinct; wall nearly smooth, marked by 

very numerous small circular pits arranged in oblique 

rows across the chambers; aperture at the end of a
MILIOLIDAE 13

very short neck, cribrate, made up of a large number 

of pores. Length 2.50 mm; diameter 0.60 mm.

In the Paris Basin Eocene there are at least two 

large species with very different types of ornamentation, as in the Jackson. As no indication is given in 

the original figures or description as to the surface 

except that it is not indicated and therefore probably 

not prominent, this form may be taken as the typical.

Terquem 5 figures a specimen with the punctae arranged in longitudinal lines. In the largest specimens 

from the Paris Basin the ornamentation is similar to 

that in the American specimens, very fine punctae arranged nearly at right angles to the axis of the chamber. In both localities specimens with longitudinal 

lines of punctae are found similar to Terquem's figures.

Jackson formation: 6463, Jackson, Miss.; 6471, Garlands 

Creek, Miss.

Miliola jacksonensis Cushman

Plate 3, figures 4-6

Miliola jacksonensis Cushnian, Contr. Cushnmii Lab. Foram. 

Research, vol. 9, p. 2, pi. 1, figs. 2, 3, 1933.

Test elongate, elliptical or fusiform, large, quinqueloculine; chambers numerous, distinct, the periphery angled; sutures distinct; wall ornamented with 

numerous oblique longitudinal costae with a single 

row, occasionally a double one, of coarse round pits 

between each two costae; aperture at the end of a very 

short neck cribrate in the adult, in the young with a 

large number of fine teeth projecting in from the 

edge. Length 2.00 mm; diameter 0.55 mm.

Holotype (no. 371527, U.S.N.M.) from Jackson formation, Jackson, Miss.

The species occurs in some numbers at Jackson, 

Miss., in the upper Eocene. It differs from M. saxorwn in the shape of the chambers and the character 

of the ornamentation. In some respects it resembles 

the QuinquelocuLwia parisiensis of D'Orbigny, as figured by Terquem, but Fornasini's tracings of D'Orbigny's original plates show only the longitudinal 

costae with no sign of punctae.

Jackson formation: 6463, Jackson, Miss.; 6471, Garlands 

Creek, Miss.

?Ocala limestone: 6725, Claiborne, Ala.

The specimens from Claiborne are internal casts of 

large size which may be this species.

Genus MASSILINA Schlumberger, 1893

Massilina decorata Cushman

Plate 3, figures 14-4.6

Massilina decorata Cushman, U.S. Geol. Survey Prof. Paper 

129, p. 143, pi. 34, fig. 7,1922; Prof. Paper 133, p. 55,1923. 

I-Iowe and Wallace, Louisiana Dept. Conservation Geol. 

Bull. 2, p. 20, pi. 2, fig. 6, 1932.

8 Soc. g ol. France M m., sir. 3, vol. 2, pi. 19, figs. 22a, b, 1882.

Test much flattened, elliptical or oval, slightly 

longer than broad, basal and apertural ends projecting, the apertural end narrowing to a small cylindrical 

neck, nearly in the longitudinal axis of the test; 

sutures rather indistinct; surface dull white; periphery rounded, the wall ornamented by very fine pits, 

giving a finely granular, matte appearance to the test. 

Maximum length 1.00 mm.

This species was described from the lower members 

of the Oligocene, being especially abundant in the 

Red Bluff clay. It occurs in considerable numbers in 

the upper part of the Jackson.

Jackson formation: 6463, Jackson, Miss.; 6471, Garlands 

Creek, Miss.

Massilina cookei Cushman, n. sp.

Plate 3, figure 17

Test in the early stages quinqueloculine, later with 

the chambers in one plane and much compressed; periphery acute and slightly carinate; chambers of the 

early portion less flattened, those of the adult very 

much flattened, widest near the inner margin, which 

stands above the remainder of the surface; ends projecting, especially the apertural end, which has a short 

cylindrical neck; surface very finely costate; nearly 

parallel with the curvature of the chamber but more 

distinct and somewhat obliquely twisted toward the 

apertural end. Length 1.00 mm; breadth 0.55 mm.

Holotype (no. 371528, U.S.N.M.) from upper 

Eocene, Jackson, Miss.

Jackson formation: 6463, Jackson, Miss, (fairly common).

Massilina jacksonensis Cushmao

Plate 3, figures 7-10

Massilina jacksonensis Cushman, Contr. Cushman Lab. 

Foram. Research, vol. 9, p. 2, pi. 1, fig. 4, 1933.

Test broadly oval or elliptical, much compressed, 

periphery subacute, apertural end slightly projecting ; .early chambers quinqueloculine, later ones in a 

single plane and JSpirolocviina-likQ; sutures distinct, 

slightly depressed; wall beautifully ornamented by a 

series of punctae, nearly circular, arranged in longitudinal curved rows parallel to the sutures and also in 

somewhat definite curved diagonal lines across the 

chamber; apertural end with a short neck but without a distinct lip. Length 1.40 mm; breadth 1.00 

mm; thickness 0.20 mm.

Holotype (no. 371529, U.S.N.M.) from Jackson 

formation, station 6463, Jackson, Miss., where it is 

common.

The early stages are occasionally found represented 

by young individuals and strongly resemble Quinqueloculina. They remotely resemble some of the forms 

referred by authors to Q. prisca D'Orbigny, but they 

are sharply angled, and the numerous specimens show
14 UPPER EOCENE FORAMINIFERA OP THE SOUTHEASTERN UNITED STATES

that they are only the young stages of the larger 

Massilina.

Jackson formation: 6463, Jackson, Miss.; 6471, Garlands 

Creek, Miss.

Massilina Jacksonensis Cushman var. punctatocostata 

Cushman

Plate 3, figures 11-13

Massilina jacksonentfis Cushman var. punctatocostata, Cushman, Contr. Cushman Lab. Foram. Research, vol. 9, p. 

3, pi. 1, figs. 5, 6, 1933.

Variety differing from the typical form in the ornamentation of the wall of the test, which has a series 

of longitudinal curved costae between the rows of 

punctae, those near the periphery more strongly developed than those nearer the inner portion of the 

chamber; the neck is more strongly developed than 

in the typical.

Holotype (no. 371530, U.S.N.M.') from upper Eocene at Jackson, Miss., where it occurs somewhat more 

abundantly than the typical form.

Jackson formation: 6463, Jackson, Miss.

Massilina sp.?

Plate 4, figures 11-13

Massilina sp.? Cushman and Applin, Am. Assoc. Petroleum 

Geologists Bull., vol. 10, p. 185, pi. 10, figs. 25-27, 1926.

This is a fairly large species with the periphery of 

the chambers keeled in the adult. The apertural end 

has a definite projecting cylindrical neck and distinct 

lip.

These specimens came from a depth of 4QO feet in 

Ohio and Red River well 2, Tyler County, Tex.

Genus SPIROLOCULINA D'OrMgny, 1826 

Spiroloculina grateloupi D'Orbigny

Plate 3, figures 18-21

Spiroloculina grateloupi D'Orbigny, Annales sci. nat, vol. 7, 

p. 29$ 1826.

Terquem, Soc. g6ol. France M£m., ser. 3, vol. 1, p. 52, 

pi.. 5, figs. 5, 6, 1878; vol. 2, p. 155, pi. 16, figs. 6a, b, 

1882.

Weisner, Archiv Protistenkunde, vol. 25, p. 203, 1912.

Cushman, U.S. Nat. Mus. Bull. 71, pt. 6, p. 31, pi. 4, figs. 

4, 5, 1917; U.S.Geol. Survey Prof. Paper 129, p. 101, 

pi. 25, fig. 2, 1922; Prof. Paper 133, p. 50, 1923. 

Spiroloculina eacavata H. B. Brady (not D'Orbigny), Challenger Kept, Zoology, vol. 9, p. 151, pi. 9, figs. 5, <'<, 

1884.

Test with the length greater than the breadth, tht 

periphery flattened or convex, the angles somewhat 

carinate, flat faces of -the test much excavated; wal) 

smooth, matte; apertural end produced with a cylindrical neck. Specimens not exceeding 0.75 mm.

All the specimens of both the Vicksburg and the 

Jackson are smaller than Recent specimens referred to 

this species. In most other respects they are similar. 

Terquem figures specimens from the upper Eocene of 

the Paris Basin that are similar to some of the specimens from the Jackson.

Jackson formation: 6463, Jackson, Miss.; 6471, Garlands 

Creek, Miss.; 6457, iy2 miles south of Shubuta, Miss.

Spiroloculina sp.?

Plate 4, figures la-c

There are some very small specimens, one of which 

is here figured, which belong to this genus. Owing to 

their scarcity and rather poor state of preservation, it 

has seemed unwise to give them a specific name.

Jackson formation: 6463, Jackson, Miss.

Genus AKTICTTLINA D'Orbigny, 1826 

Articulina terquemi Cushman

Plate 4, figures 2, 3

ArtiGulina terquemi Cushman, Contr. Cushman Lab. Foram. 

Research, vol. 9, p. 3, pi. 1, figs. 7a-c, 1933.

Test small, somewhat larger than broad, compressed; 

periphery broadly rounded; chambers indistinct except the last two, nearly involute, inflated, ornamented 

by numerous rounded longitudinal costae, in general 

parallel to the periphery, often somewhat uneven, 

apertural end with a distinct lip; sutures indistinct; 

wall matte; aperture circular, with a very distinct, 

slightly flaring lip without a definite tooth. Length 

0.25 mm; breadth 0.13 mm; thickness 0.07 mm.

Holotype (no. 371531, U.S.N.M) from Jackson formation, Miss.

This species somewhat resembles one of the figures 

given by Terquem 6 as Articulina gibbosula D'Orbigny, 

from the Eocene of the Paris Basin. It is not the 

same as D'Orbigny's species of the Miocene of the 

Vienna Basin, and Terquem's figure 25 is much more 

like D'Orbigny's species. On the same plate, figure 24, 

referred by Terquem to Articulina nitid'd D'Orbigny, 

is somewhat similar but not identical with our form. 

A figure given by Sowerby 7 as " Triloculina striata 

Brown " also somewhat resembles our specimens, but 

the costae are not the same, and the species is probably 

a Massilina. Our species is somewhat similar to 

Articulina byramensis Cushman, but that species has 

a very distinctive shape as well as a very different 

ornamentation.

Jackson formation: 6463, Jackson, Miss.

°Soc. ffgol. Trance M6m., s£r. 3, vol. 2, pi. 15 (23), fig. 26, 1S82. 

7 Sowerby, G.B., Foraminifera from the Colne tidal river, pi., 

flgs. 5, 6, 3850.
LAGENIDAE 15

Genus TRILOCUIINA D'Orbigny, 1826 

Triloculina sp.?

Plate 4, figures 4a-c

The figured form is evidently a Triloculina, but a 

large enough series for specific determination is not 

available. The chambers are highly angled, and the 

aperture is circular and terminal, with a distinctly 

bifid tooth.

Jackson formation: 6463, Jackson, Miss.

Family OPHTHALMIDIIDAE

Genus CORNUSPIRA Schultze, 1854

Cornuspira olygogyra Hantken

Plate 4, figure 14

Cornuspira olygogyra Hantken, -Magy. kir. foldt. int. lilvkonyve,

vol. 4, p. 16, pi. 1, fig. 10, 1875 (1876) ; K. ungar. geol.

Anstalt Mitt. Jahrb., vol. 4, p. 20, pi. 1, fig. 10, 1875

(1881).

Liebus, Geol. Reichsanstalt, Jahrb., vol. 56, p. 360, 1906.

Test close-coiled, much compressed, sides flattened, 

periphery truncate; chamber rectangular in section, 

in end view the apertural end higher than broad, the 

sides in the last-formed chamber very slightly concave, sutural line distinct and the basal edge of the 

chamber slightly thickened; wall smooth, glossy. Diameter 0.60 mm.

There is but a single microspheric specimen in the 

collection from the Jackson formation at Jackson, 

Miss., but this is very typical in the shape of the 

test, the slightly concave sides, and the raised ridge at 

the base of the chamber as in Hantken's figure. His 

specimen was evidently megalospheric. Our specimen 

is probably not a full-grown one. Hantken's record 

is from the Clavulina szciboi beds of Hungary, and 

that of Liebus from the upper Eocene of Biarritz.

Jackson formation: 6463, Jackson, Miss.

Family LAGENIDAE 

Genus ROBULUS Montfort, 1808 

Robulus gutticostatus (GUmbel) 

Plate 5, figures 1, 2

Robulma gutticostata Giimbel, K. bayer. Akad. Wiss. Miinchen, 01. 2, Abh., vol. 10, p. 643, pi. 1, fig. 74, 1868 (1870).

Hantken, Magy. kir. foldt. int. fivkonyve, vol. 4, p. 48, pi.

6, fig. 10, 1875 (1876) ; K. ungar. geol. Anstalt Mitt.

Jahrb., vol. 4, p. 57, pi. 6, fig. 10, 1875 (1881).

Test close-coiled, periphery acute, keeled, compressed; chambers distinct but not inflated, 9 to 11 

chambers in the last-formed coil in the adult; sutures 

distinct, ornamented, limbate, a raised costa on the 

exterior broken into a row of rounded, beadlike protuberances, especially toward the umbilicus, the beads

62330 35 2

increasing in size, with the largest nearest the umbilicus; sutures very slightly curved; wall between the 

sutures smooth; aperture peripheral, radiate, slightly 

projecting. Diameter up to 1.50 mm.

The type figure of Giimbel shows but 7 chambers 

in the last-formed whorl; Hantken's figure shows 8. 

Most of the American specimens have 9, or 1 or 2 

more in adult specimens. Giimbel's types are from 

the Eocene of Bavaria, and Hantken's from the upper 

Eocene of Hungary.

Ocala limestone: 7159, 4 miles southeast of Whatley, Ala.; 

1/26, Drewry, Ala.; 1/3S, 3% miles north of Grove Hill, Ala.; 

1/48, Duncan, Fla.

Jackson formation: 7203, 3% miles southeast of Cullomburg, Ala.; 6457, 1% miles south of Shubuta, Miss.

Robulus gutticostatus (Giimbel) var. cocoaensis (Cushman)

Plate 5, figures 3, 4

Cristellaria gutticostata (Gumbel) var. coooaensis Cushman, 

Contr. Cushman Lab. Foram. Research, vol. 1, p. 67, 

pi. 10, fig. 11, 1925.

Variety with usually a larger number of chambers 

in each whorl than in the typical form; sutures with 

very distinct beading, the beads growing larger toward the inner end of the sutures, and the umbonal 

region occupied by a series of large bosses, last-formed 

suture in adults often unornamented and slightly 

depressed.

Jackson formation: 2637, Cocoa, Ala., type station (fairly 

abundant) ; also in the Jackson of well, samples in Louisiana.

Robulus gutticostatus (Gumbel) var. yazooensis Cushman

Plate 5, figure 5

Robulus gutticostatus (Giimbel) var. yazooensis Cushman, 

Contr. Cushman Lab. Foram. Research, vol. 9, p. 4, 

pi. 1, fig. 8, 1933.

Variety differing from the typical form in having a 

yery few beads on the proximal end of the chambers, 

the distal portion unornamented and depressed, in the 

adult with the last few sutures entirely without ornamentation and depressed throughout, the keel wanting, and the periphery of each chamber nearly 

straight, each slightly projecting beyond its immediate 

predecessor at the angle. Diameter 0.75 mm.

Holotype (no. 371532, U.S.N.M.) from Jackson 

formation at station 7220, half a mile southeast of 

Melvin, Ala.

Robulus alato-limbatus (GUmbel)

Plate 6, figures 2a, b

Robulina alato-limbata Gumbel, K. bayer. Akad. Wiss. Miinchen, Cl. 2, Abh., vol. 10, p. 641, pi. 2, figs. 70a, b, 1868 

(1870).

Robulus alato-limbata (Gumbel) Howe and Wallace, Louisiana Dept. Conservation Geol. Bull. 2, p. 37, pi. 3, figs. 2a, 

b, 1932.
16 UPPER EOCENE FORAMINIFERA OF THE SOUTHEASTERN UNITED STATES

Cristellaria alato-limbaUi (Gtimbel) Cushman arid Applin, Am. 

Assoc. Petroleum Geologists Bull., vol. 10, p. 171, pi. 8, 

fig. 8, 1926.

Test close-coiled, last-formed coil composed of few 

chambers, usually 7 in number, the central region 

with a large umbo not greatly projecting above the 

general surface but distinct, periphery with a narrow 

keel; chambers distinct, not inflated; sutures distinct, 

not depressed, strongly curved; aperture radiate; wall 

smooth. Diameter 0.65 mm.

This species is already recorded from the upper 

Eocene of Texas from Ohio and Red River well 2, at 

a depth of 400 feet, Tyler County. Giimbel described 

the species from the upper Eocene of central Europe.

Cooper marl: 5531, Cooper River, S.C.

Ocala limestone: 1/38, 3% miles north of Grove Hill, Ala.

Jackson formation: 6457, 1% miles south of Shubuta, Miss.

Robulus articulatus (Reuss) var. texanus (Cushman and

Applin)

Plate 4, figures 16, 17

Cristellaria articulata Reuss var. texana Cushman and Applin, Am. Assoc. Petroleum Geologists Bull., vol. 10, p. 

170, pi. 8, figs. 1, 2, 1926.

Robulus articulatus (Reuss) var. texanus Ellisor, Am. Assoc. 

Petroleum Geologists Bull., vol. 17, no. 11, pi. 2, fig. 3, 

1933.

L&nticulina articulata (Reuss) var. texana Howe and Wallace, Louisiana Dept. Conservation Geol. Bull.. 2, p. 

31, pi. 5, figs. 1, 2, 1932.

This variety is of large size, in the young with the 

chambers close-coiled, but in the adult with the central portion becoming visible, owing to the shortening 

of the chambers, which fail to cover the preceding 

whorl entirely, periphery with a distinct, rounded 

keel, in the adult with as many as 10 to 12 chambers. 

Diameter 1.60 mm or less.

This variety is related to the typical form, as it 

occurs in the Tertiary of Europe. " Type specimens 

of the variety are from Warren well 4, 4,250 feet, 

Texas Exploration Co., Hockley, Harris County, Tex." 

Jackson formation.

Cooper marl: 5531, Cooper River, Charleston, S.C.; 6319, 

Ingleside, S.C.

Robulus limbosus (Reuss)

Plate 6, figure 5

Robulina limbosa Reuss, Akad. Wiss. Wien Sitzungsber., vol.

48, pt. 1, p. 55, pi. 6, figs. 69a, b, 1863 (1864). 

Hantken, Magy. kir. foldt. int. Evkonyve, vol. 4, p. 48, pi. 6,

fig. 11, 1875 (1876) ; K. ungar. geol. Anstalt Mitt. Jahrb.,

vol. 4, p. 57, pi. 6, fig. 11, 1875 (1881). 

Robulus limbosus Ellisor, Am. Assoc. Petroleum Geologists

Bull., vol. 17, no. 11, pi. 2, figs, la, b, 1933.

Test close-coiled, umbonate, periphery with a broad, 

very thin, platelike transparent keel; chambers distinct, 9 or 10 in the last-formed coil; sutures distinct, 

curved, slightly limbate, of clear material, ending at

the umbo in a transparent mass of clear shell material; 

wall smooth; apertural face slightly concave, aperture 

on the peripheral angle, radiate. Maximum diameter 

1.50 mm.

The Coastal Plain specimens are very close indeed to 

the figures of R. limbosm, especially that of Hantken.

Ocala limestone: 1/38, 3% miles north of Grove Hill, Ala.

Jackson formation: 7203, 3% miles southeast of Cullomburg, 

Ala.; 6457, iy2 miles south of Shubuta, Miss.; 6473, Yazoo 

City, Miss.

Robulus limbosus (Reuss) var. hockleyensis (Cushman and

Applin)

Plate 4, figures 15a, b; plate 6, figures 3a, b

Cristellaria limbosa (Reuss) var. hockleyensis Cushman and

Applin, Am. Assoc. Petroleum Geologists Bull., vol. 10,

p. 171, pi. 8, figs. 3, 4, .1926. 

Robulus limbosus (Reuss) var. hocMeyensis Ellisor, Am. Assoc.

Petroleum Geologists Bull., vol. 17, no. 11, pi. 1, figs.

lla, b, 1933.

Test differing from the typical form in the fewer 

chambers and the broader form in apertural view, the 

keel perhaps not so broad and thin, and the central 

umbo not so strongly developed. Diameter 0.75 mm.

Type specimens from Bridge Creek, iy2 miles above 

Angelina River, San Augustine County, Tex.

This is evidently related to RobulMS limbosus, which 

is known from the upper Eocene of central Europe.

Specimens probably the same as the Texas variety 

occur in the upper Eocene of the Coastal Plain at 

numerous stations. The material is not always well 

preserved, and accurate identification is difficult.

Castle Hayne marl: 5611, Wilmington, N.C.

Cooper marl: Ashley marl works, Dorchester County, S.C.; 

6319, Ingleside, S.C.

Jackson formation : 7203, 3% miles southeast of Cullomburg, 

Ala.

Robulus propinquus (Hantken)

Plate 6, figures la, b

Cristellaria propinqua Hantken, Magy. kir. foldt. int. Evkonyve,

vol. 4, p. 45, pi. 5, fig. 4, 1875 (1876) ; K. ungar. geol.

Anstalt Mitt. Jahrb., vol. 4, p. 52, pi. 5, fig. 4, 1876

(1881). 

Cushman and Applin, Am. Assoc. Petroleum Geologists

Bull., vol. 10, p. 172, pi. 8, fig. 9, 1926. 

Robulus propinquus Ellisor, Am. Assoc. Petroleum Geologists

Bull., vol. 17, no. 11, pi. 7, figs. 12a, b, 1933.

Test with the early portion close-coiled, later tending to become uncoiled, the periphery subacute, later 

chambers increasing in width; chambers few, 6 or 7 

in the last-formed coil, distinct but not inflated; sutures distinct, curved, not depressed; wall smooth; 

apertural face somewhat tapering toward the aperture, 

which is radiate and slightly projecting. Diameter 

0.60 mm.

The only upper Eocene specimens from the Coastal 

Plain region are from the Ohio and Red River well 2, 

Tyler County, Tex., at a depth of 1,056 feet.
LAGENIDAE 17

These specimens are very similar to those figured 

and described by Hantken from the upper Eocene of 

central Europe.

Robulus arcuato-striatus (Hantken) var. carolinianus 

Cushman

Plate 6, figures 6a, b

Rotoilus arcuato-striatus (Hautken) var. carolinianus Cushman, Contr. Cushman Lab. Foram. Research, vol. 9, 

p. 4, pi. 1, figs. 9a, b, 1933.

Test close-coiled throughout, strongly umbonate, 

periphery keeled with a fairly wide thin carina; chambers very distinct, 8 or 9 in the last-formed coil, of 

uniform shape and increasing very slightly in size 

as added; sutures distinct, strongly limbate, slightly 

raised, very strongly curved, ending in the middle in a 

clear umbo; wall smooth except for the slightly raised 

sutures; aperture slightly protruding, at the peripheral angle, radiate, apertural face slightly concave, 

the sides thickened. Diameter 1.30 mm.

Holotype of variety (no. 371548, U.S.N.M.) from 

Ocala limestone, station 1/38, 3^4 miles north of 

Grove Hill, Ala.

Cooper marl: 6319, Ingleside, S.C.; 5531, Cooper River, S.C. 

Ocala limestone: 1/21, Covington County, Ala.; 1/38, 3*4 

miles north of Grove Hill, Ala. 

Jackson formation: 6457, 1% miles south of Shubuta, Miss.

Genus LENTICULINA Lamarck, 1804 

Lenticulina convergens (Bornemann)

Plate 6, figures 4a, b

Cristellaria convergens Bomemann, Deutsch. geol. Gesell. 

Zeitschr., vol. 7, p. 327, pi. 13, figs. 16, 17, 1855.

Test compressed, ovate, close-coiled, thickest in the 

umbonal region but without a distinct umbo; chambers 

fairly distinct, the later ones increasing somewhat in 

height; sutures fairly distinct, not depressed; apertural face strongly convex, aperture at the peripheral 

angle, radiate, slightly produced. Diameter 1.00 mm.

Bornemann originally described this species from 

the Oligocene of central Europe, and many fossil and 

Recent specimens have been assigned to this name.

This species has been recorded as far back as the 

Triassic, but the earlier records lack further confirmation.

Ocala limestone: 7097, Bainbridge,. Ga.; 7719, Spring Mill 

Branch, Jenkins County, Ga.

Genus SARACENARIA Defiance, 1824 

Saracenaria arcuata (D'Orbigny) var. hantkeni Cushman

Plate 5, figures 6, 7

Cristellaria arcuata Hantken (not D'Orbigny), Magy. kir. 

foldt. int. Evkb'nyve, vol. 4, p. 45, pi. 5, figs. 5a-c, 6, 1875 

(1876) ; K. ungar. geol. Anstalt Mitt. Jahrb., vol. 4, 

p. 53, pi. 5, figs. 5a-c, 6, 1875 (1881).

Saracenaria arcuata (D'Orbigny) var. hantkeni Cushman, 

Contr. Cushman Lab. Foram. Research, vol. 9, p. 4, pi. 1, 

figs. 11, 12, 1933.

Test longer than broad, periphery subacute, apertural face truncate, test triangular in transverse section, early portion somewhat close-coiled in the first 

few chambers, soon becoming uncoiled; chambers comparatively few, usually only 7 or 8, distinct but not 

inflated, angles subacute, almost keeled in the lastformed chambers; sutures distinct, very slightly if at 

all depressed; wall smooth and polished; aperture 

peripheral, radiate, slightly projecting. Maximum 

length 1.30 mm; breadth of final chamber 0.50 mm.

Holotype of variety (no. 371533, U.S.N.M.) from 

Cooper marl, station 6319, Ingleside, S.C.

This is, very similar to if not identical with the 

variety figured by Hantken from the Eocene of Hungary and referred to Cristellcvria arcwata D'Orbigny. 

It is usually much more acute than the Oligocene 

species of the Vienna Basin, and the apertural face is 

much more flattened. The Eocene variety is also 

straighter, the uncoiled portion being much less 

strongly curved on both faces, but especially in the 

inner concave margin.

Cooper marl: 5531, Cooper River, S.C.; 6319, Ingleside, 

S.C.; 11861, Steep bluff on west side of Biggin Creek, Berkeley County, S.C.; 11863, three-fourths mile west of Old Biggin 

Church, Berkeley County, S.C.; 11862, 1 mile west of Givhans, 

Dorchester County, S.C.; 11857, 1 mile south of Moncks Corner, Berkeley County, S.C.

Ocala limestone: 7719, Spring Mill Branch, Jenkins County, 

Ga.; 1/38, 3^ miles north of Grove Hill, Ala.; 1/43, 1^ miles 

southwest of Perdue Hill, Ala.

Jackson formation: 7203, 3% miles southeast of Cullomburg, 

Ala.; 6457, 1% miles south of Shubuta, Miss.

Genus PLANULARIA Defrance, 1824 

Planularia truncana (GUmbel)

Plate 6, figures 7a, b

Oristellaria truncana Giimbel, K. Akad. Wiss. Miinchen, Cl. 2, 

Abb.., vol. 10, p. 639, pi. 1, figs. 68a, b, 1868 (1870).

Test much compressed, periphery slightly keeled; 

the earliest chambers thicker than the others, forming 

a distinct umbo 5 chambers numerous, early ones short, 

making a close-coiled test, later ones increasing rapidly 

in length but not actually uncoiling, the inner margin 

extending back to the previous coil; sutures distinct, 

somewhat limbate; wall smooth; aperture at the peripheral angle. Maximum length 1.50 mm.

The specimens from the upper Eocene of the Coastal 

Plain are apparently identical with that described by 

Giimbel from the upper Eocene of the Bavarian Alps.

Castle Hayne marl: 5611, Wilmington, N.C.

Cooper marl: Baldock, S.C.

Ocala limestone: 7719, Spring Mill Branch, Jenkins County, 

Ga.; 6768, Marianna, Fla.

Jackson formation: 2637, Cocoa, Ala.; 7203, 3% miles southeast of Cullomburg, Ala.
18

Planularia cooperensis Cushman

Plate 6, figures 8a, b

Planularia. cooperensis Cushman, Contr. Cushman Lab. Foram. 

Research, vol. 9, p. 5, pi. 1, figs. 10 a, b, 1933.

Test large, strongly compressed, periphery rounded, 

not keeled, 15 or more chambers in the final coil, of 

nearly uniform shape, increasing gradually in size as 

added; sutures fairly distinct, very slightly limbate, 

slightly curved, later ones very slightly depressed, 

earlier ones flush with the surface; wall smooth, matte. 

Lenth 4.00 mm5 breadth 3.00 mm; thickness 0.60 mm.

Holotype (no. 371534, U.S.N.M.) from Cooper marl, 

station 11863, highway 17, three-fourths mile west of 

Old Biggin Church, Berkeley County, S.C.

This is a fine large species of the general form 

usually assigned to Cristellaria cassis (Fichtel and 

Moll), which is, however, a very different species. 

The Eocene species has no keel, and the wall is entirely 

smooth.

Cooper marl: 11SG1, Steep Bluff, on west side of Biggin 

Creek, Berkeley County, S.C.; 11863, three-fourths mile west 

of Old Biggin Church, Berkeley County, S.C.

Genus MARGINULINA D'Orbigny, 1826 

Margjnulina karreriana Cushman, n. sp.

Plate 7, figures 1, 2

Marginulina abbreviala Karrer (not Neugeboren, 1851), Akad. 

Wiss. Wien Sitzungsber., vol. 44, pt. 1, p. 445, pi. 1, flg. 1, 

1861 (1862).

Test elongate, somewhat compressed, periphery 

rounded, earliest chambers coiled, later ones uncoiled 

but with the sutures oblique, last-formed chambers 

much inflated; sutures of the earlier portion very 

slightly if at all depressed, later ones distinctly depressed; wall smooth; aperture at the dorsal margin 

radiate, somewhat produced. Length 0.90 mm; 

breadth 0.45 mm; thickness 0.20 mm.

Holotype (no. 371535, U.S.N.M.) from Cooper marl, 

station 11857, 1 mile south of Moncks Corner, Berkeley County, S.C.

Specimens from the Cooper marl are like the figures 

given by Karrer, but these are not the same as the 

early species of Neugeboren under the same name.

Cooper marl: 11863, three-fourths mile west of Old Biggin 

Church, Berkeley County, S.C.; 11862, 1 mile west of Givhans, 

Dorchester County, S.C.; 11857, 1 mile south of Moncks Corner, 

Berkeley County, S.C.

Marginulina subrecta Franke

Plate 7, figures 3a, b

Marginulina subrecta Franke, Danniarks geol. Unders0gelse, 

2. Raekke, Nr. 46, p. 19, pi. 1, fig. 28, 1927.

Test elongate, subcylindrical, not compressed, circular in transverse section, earliest chambers showing

traces of coiling, later ones in a rectilinear series, 

slightly inflated; sutures distinct, later ones slightly 

compressed; wall smooth; aperture in the adult becoming terminal, central, slightly exserted, radiate. 

Length 0.90 mm; diameter 0.35 mm.

Cooper marl: 11861, Steep bluff on west side of Biggin 

Creek, Berkeley County, S.C.

Marginulina jacksonensis (Cushman and Applin)

Plate 7, figures 5a, b

Cristellaria jacksonensis Cushman and Applin, Am. Assoc. 

Petroleum Geologists Bull., vol. 10, p. 172, pi. 8, fig. 10, 

1926.

Test much elongate, the greatest width being at the 

base, where there are a few close-coiled chambers, later 

portion consisting of 3 to 5 uniserial chambers, much 

inflated, especially toward the apertural end, early 

portion with the periphery acute, consisting of 4 or 5 

chambers in the coil, the later uncoiled chamber 

progressively increasing in thickness so that the lastformed chamber is often circular in transverse section; sutures distinct, those of the later portion depressed; wall smooth; aperture radiate, terminal. 

Maximum length nearly 1.00 mm; breadth at the base 

0.20 mm.

Type specimens were obtained -from the Jackson 

group in Ohio and Red River well 2, Tyler County, 

Tex., at a depth of 1,056 feet. It occurs also in the 

Alazan clays of Mexico.

Marginulina cocoaensis Cushman 

Plate 7, figures 6, 7

Marginulma cocoaensis Cushman, Coutr. Cushmau Lab. Foram.

Research, vol. 1, p. 67, pi. 10, figs. 9, 10, 1925. 

Howe and Wallace, Louisiana Dept. Conservation Geol.

Bull. 2, p. 33, pi. 7, fig. 5, 1932. 

Ellisor, Am. Assoc. Petroleum Geologists Bull., vol. 17, no.

11, pi. 6, fig. 6, 1933.

Test elongate, compressed, initial end composed of a 

few partly coiled chambers much compressed, later 

and major portion composed of more rounded chambers, 6 to 10 in number in adult specimens; sutures 

fairly distinct, of clear shell material; ornamentation 

consisting of 9 or 11 lamellate costae running from the 

initial end to the base of the last-formed chamber in 

adults, the last chamber being usually smooth in 

completely developed specimens, two of the costae 

forming keels on the compressed portion of the test; 

aperture at the peripheral side of the apertural face, 

radiate, at the end of a distinct projection. Maximum" 

length 1.60 mm; breadth 0.25 mm.

MarffinuUna cocoaensis is a very well characterized 

species with its strongly lamellate costae, compressed 

early portion, the smooth last chamber, and the peculiar apertural characters.
LAGENIDAE 19

Jackson formation: 2637, Cocoa, Ala., type locality; 7203, 

3 '/£ miles southeast of Cullomburg, Ala.; 6457, 1^ miles south 

of Shubuta, Miss.

Cooper marl: 6321, Baldock, S.C.

Ocala limestone: 7345, Suwannee River, 2^4 miles above 

Branford, Fla.; 7719, Spring Mill Branch, Jenkins County, Ga.; 

1/38, 3% miles north of Grove Hill, Ala.

Marginulina cooperi Cushman

Plate 7, figures 4a, b

Margmulina cooperi Cushman, Contr. Cushman Lab. Foram. 

Research, vol. 9, p. 5, pi. 1, fig. 13, 1933.

Test of medium size, the earlier chambers closecoiled, later ones becoming uncoiled, periphery broadly 

rounded, without a keel but with 2 or 3 short, stout 

spines at the curve of the test, 1 to a chamber; chambers distinct, about 6 in a volution in the earlier part, 

not inflated, entirely involute, in the later uncoiled 

part with more inflated chambers; sutures very distinct, not depressed, slightly limbate; aperture peripheral throughout, rather large, radiate. Length 

1.50 mm.

Holotype (no. 371536, U.S.N.M.) from Cooper marl, 

Cooper River, S.C.

This is a very distinct species in the American 

Eocene. It perhaps most closely resembles M. 

hauerina (D'Orbigny) and M. alazanensis (Cushman) 

but is distinct in many of its characters.

Cooper marl: 5531, Cooper River, Charleston County, S.C.

Marginulina fragaria (Giimbel) var. texasensis (Cushman 

and Applin)

Plate 7, figures 8-10

Cristellaria fragaria Giimbel var. texasensis Cushman and 

Applin, Am. Assoc. Petroleum Geologists Bull., vol. 10, 

p. 171, pi. 8, figs. 5-7, 1926.

Lenticulina fragaria (Giimbel) var. texasensis Howe and Wallace, Louisiana Dept. Conservation Geol. Bull. 2, p. 32, 

pi. 5, figs. 3-5, 1932.

Marginulina fragaria (Giimbel) var. texasensis Ellisor, Am. 

Assoc. Petroleum Geologists Bull., vol. 17, no. 11, pi. 2, 

fig. 4, 1933.

Test differing from the typical form in the more 

elongate character of the test, the more coarsely beaded 

ornamentation, and the more deeply excavated sutures, 

the periphery often decidedly lobulate. Maximum 

length 1.25 mm; breadth 0.50 mm; thickness 0.25 mm.

A specimen from the Jackson formation at Cocoa, 

Ala., station 2637, is very close to if not identical with 

the Texas variety. The type specimens are from 

Haynes well 1, near Burkeville, Newton County, Tex., 

at a depth of 3,175 to 3,270 feet.

Marginulina cooperensis Cushman

Plate 7, figures 11, 12

Maryinuli'iia cooperensis Cushman, Contr. Cushman Lab. 

Forara. Research, vol. 9, p. 8, pi. 1, figs. 14,15,1933.

Test elongate, tapering, the early portion much compressed, later portion circular in transverse section; 

chambers distinct, earlier ones forming part of a coil; 

last 3 or 4 subglobular; sutures somewhat hidden by 

the ornamentation, which is composed of longitudinal 

costae, numerous, about 15 to 18 on the visible half 

of the last chamber, becoming somewhat produced at 

the proximal margin of each chamber; aperture terminal with a definite cylindrical neck and thickened 

lip. Length 1.40 mm; maximum breadth 0.35 mm.

Holotype (no. 371537, U.S.N.M.) from station 5531, 

Cooper marl, Cooper River, Charleston County, S.C.

This has not been noted at any of the other stations 

from which material was examined.

Genus DENTALINA D'Orbigny, 1826

Dentalina cocoaensis (Cushman)

Plate 8, figures 1, 2

Nodosaria sp. Cushman and Applin, Am. Assoc. Petroleum

Geologists Bull., vol. 10, p. 170, pi. 7, fig. 17, 1926. 

Nodosaria cocoaensis Cushman, Contr. Cushman Lab. Foram.

Research, vol. 1, p. 66, pi. 10, figs. 5, 6, 1925; Jour.

Paleontology, vol. 1, p. 153, pi. 24, fig. 1, 1927. 

Dentalina cocoaensis Ellisor, Am. Assoc. Petroleum Geologists

Bull., vol. 17, no. 11, pi. 2, fig. 5, 1933.

Test elongate, very slender, slightly curved, gradually tapering from the acute or spinose initial end, 

early portions with the sides straight, toward the 

apertural end with the chambers slightly inflated; 

chambers in the adult 11 to 14, in the very early portion only slightly longer than broad, becoming as much 

as 2y2 to 3 times as long as broad in the last-formed 

ones; sutures very distinct, of clear shell material but 

not depressed; wall very smooth, slightly glossy in 

well-preserved specimens, apertural end produced. 

Maximum length, 2.25 mm; maximum breadth, 

0.20 mm.

This species is allied to Nodosaria fliformis 

D?Orbigny and other species. It is very close to 

Hantken's figure of Dentalina consobrina D'Orbigny, 

from the Eocene of the Clavulina szdboi beds of 

Hungary, but the early portion is different. It is 

close to the Recent material figured by Flint from off 

the coast of the United States under the name Nodosaria filiformis D'Orbigny. The initial end is ornamented by a single stout, short spine, the sutures are 

of clear shell material and broad, only the later chambers show any swelling, and the form is very constant. 

The best-preserved specimens have 12, 13, or 14 chambers, and the number is very constant. The species 

occurs in the upper Eocene of Texas, in the Ohio and 

Red River well 2, Tyler County, at a depth of 537 

-feet, and in the Alazan clays of Mexico.

Jackson formation: 2637, Cocoa, Ala., the type locality; 

7203, 3V2 miles southeast of Cullomburg, Ala.; 6457, 1% miles 

south of Shubuta, Miss.
20 UPPER EOCENE FORAMINIFERA OF THE SOUTHEASTERN UNITED STATES

Cooper marl: 5531, Cooper River, S.C.; 11861, steep bluff on 

west side of Biggin Creek, Berkeley County, S.C.; 11862, 1 

mile west of Givhans, Dorchester County, S.O.

Ocala limestone: 7159, 4 miles southeast of Whatley, Ala.; 

7719, Spring Mill Branch, Jenkins County, Ga.; 1/8, Beck, 

Ala.; 1/38, S^A miles north of Grove Hill, Ala.; 1/43, iy2 

miles southwest of Perdue Hill, Ala.

Dentalina cooperensis Cushman 

Plate 8, figures 3, 4

Dentalina cooperensis Cushman, Contr. Cushman Lab. Foram. 

Research, vol. 9, p. 8, pi. 1", fig. 17, 1933.

Test elongate, slightly compressed, very slightly 

tapering, gently curved, periphery only slightly sinuate, apical end pointed or with a single small spine; 

chambers few, usually about 10 in the adult specimen, 

often indistinct; sutures fairly distinct, oblique; wall 

smooth, matte; aperture at the periphery of the 

chamber slightly projecting. Length 2.00-2.50 mm; 

breadth 0.30 mm.

Holotype (no. 371538, U.S.N.M.) from Cooper 

marl, Cooper Eiver, Charleston County, S.C.

This species is allied to some of the forms referred 

to Nodosaria com/munis D'Orbigny, but in the upper 

Eocene of the Coastal Plain it seems to be distinct. 

The margins are nearly entire, only slightly sinuous; 

occasionally, near the apertural end, the sutures are 

distinctly oblique, and the number of chambers is 

usually not more than 10.

Cooper marl: 5531, Cooper River, S.C.; 11861, steep bluff on 

west side of Biggin Creek, Berkeley County, S.C.

Ocala limestone: 6768, Marianna, Fla.; 7159, 4 miles southeast of Whatley, Ala.; 7719, Spring Mill Branch, Jenkins 

County, Ga.; 1/38, 3^ miles north of Grove Hill, Ala.; 1/43, 

iy2 miles southwest of Perdue Hill, Ala.

Jackson formation: 6457, 1% miles south of Shubuta, Miss.

Dentalina hantkeni Cushman

Plate 8, figures 5, 6

Dentalina hantkeni Cushman, Contr. Cushman Lab. Foram.

Research, vol. 9, p. 9, pi. 1, figs. 18, 19, 1933. 

Dentalina, budensis Hantken, Magy. kir. foldt. fivkonyve, vol.

4, p. 28, pi. 3, fig. 12, 1875 (1876). [Not Nodosaria

budensis Hantken.]

Test elongate, arcuate, somewhat compressed, composed of a few chambers, initial end rounded; chambers distinct, increasing in length as added, outer curve 

sinuate, inner curve nearly uniform; sutures fairly 

distinct, somewhat oblique; aperture near the inner 

curve with a slightly produced neck. Length 1.25 

mm; greatest diameter 0.15 mm.

Holotype (no. 371539, U.S.N.M.) from Ocala limestone, station 6741, 3% miles northeast of Brooklyn, 

Ala.

Hantken's figure and description of Dentalina^ 

' budensis are identical with this species from the upper 

Eocene. Hantken's specimens were from the Eocene 

of Hungary.

Ocala limestone: 6740, mouth of Robinsons Mill Creek, Ala.; 

6741, 3% miles northeast of Brooklyn, Ala.

Dentalina jacksonensis (Cushman and Applin)

Plate 8, figures 7-9

Nodosaria jacksonensis Cushman and Applin, Am. Assoc. Petroleum Geologists Bull., vol. 10, p. 170, pi. 7, figs. 14-16,

1926. 

Cushman, Jour. Paleontology, vol. 1, p. 153, pi. 24, flg. 3,

1927.

Cole, Bull. Am. Paleontology, vol. 14, no. 53, p. 208, pi. 3, 

fig. 12, 1928.

Test elongate, tapering, gently curved, initial end 

rounded, ornamented with one to several spines, sides 

lobulate throughout, more strongly so in later growth; 

chambers subglobular, fairly numerous, usually 10 in 

well-developed specimens, inflated, length and breadth 

about equal except the last 1 or 2 in the adult, which 

are slightly longer than broad; sutures distinct, somewhat depressed, of clear shell material, surface smooth, 

glossy to dull; aperture with a cylindrical neck, the 

aperture itself not well preserved. Maximum length 

2.50 mm; maximum breadth 0.35 mm.

In its character of a broadly rounded initial end, 

with occasionally a group of spines, and in the apertural neck, this species resembles Nodosaria abyssorum 

H. B. Brady. The Eocene species has more chambers, 

is more slender, and is always curved. Specimens are 

very constant in their characters. There is some resemblance to N. insecta Schwager, but our species is 

curved, much more tapering, and has more chambers. 

The types were from the upper Eocene of Texas, from 

Haynes well 1, Newton County. The species is also 

common in the Alazan clays of Mexico.

Cooper marl: 5531, Cooper River, S.C.; 6321, Baldock, S.C.; 

11861, steep bluff on west side of Biggin Creek, Berkeley 

County, S.C.

Ocala limestone: 7719, Spring Mill Branch, Jenkins County, 

Ga.; 1/38, 3V2 miles north of Grove Hill, Ala.; 1/43, 1% miles 

southwest of Perdue Hill, Ala.

Jackson formation: 2637, Cocoa,- Ala.; 7203, 3% miles southeast of Cullomburg, Ala.; 6457, 1% miles south of Shubuta, 

Miss.

Dentalina halkyardi Cushman

Plate 8, figure 10

Dentalma halkyardi Cushman, Contr. Cushman Lab. Foram.

Research, vol. 9, p. 9, pi. 1, fig. 20, 1933. 

Nodosaria (Dentalina) spinulosa Halkyard (not Montagu),

Manchester Lit. Philos. Soc. Mem. and Proc., vol. 62,

p. 79, 1919.

Test elongate, tapering, initial end rounded, composed of a few chambers, 7 in the type specimen, 

globular, the earlier ones more overlapping than the 

later ones; sutures distinct and depressed; wall ornamented in the earliest chambers by longitudinal costae, 

about 7 or 8 visible on each side of the chambers, the 

proximal end of each costa somewhat projecting, in
LAQENIDAB 21

the next-formed chambers the costae are broken into 

short bits, which become spinose at the proximal end, 

and in the last-formed chambers the ornamentation 

consists of a series of fine, almost hispid spines in 

longitudinal lines; apertural end with a cylindrical 

neck with a thickened lip, the neck with annular corrugations and the outer border with a series of teeth. 

Length 1.40 mm or more; breadth of last-formed 

chamber 0.30-0.40 mm.

From the description given by Halkyard in the reference above, this is the same as his material from the 

upper Eocene Blue marl of Cote des Basques, Biarritz. 

His description is as follows: " The earlier chambers 

of the shells are invariably ornamented with longitudinal costae, which later are interrupted or broken up 

into short lengths, and later still into spines or prickles 

pointing backward. The prickles first make their appearance on the basal portion of each segment."

Cooper inarl: 5531, Cooper River, S.C.; 6319, Ingleside, S.O.; 

11861, steep bluff on west side of Biggin Creek, Berkeley 

County, S.C.; 11863, three-fourths mile west of Old Biggin 

Church, Berkeley County, S.C.; 11862, 1 mile west of Givhans, 

Dorchester County, S.C.

Dentalina cf. D. adolphina D'Orbigny

Plate 8, figures 11, 12

There are a very few incomplete specimens from the 

Cooper marl, station 11857, pit on U.S. Highway IT, 

about 1 mile south of Moncks Corner, Berkeley 

County, S.C., which may be referred to this species. 

Figures of two of these specimens are given.

Dentalina cf. D. vertebralis (Batsch)

Plate 8, figures 13, 14

Specimens similar to the figures given here are common in the Cooper marl of South Carolina. They are 

in some respects close to the species described and 

figured by Batsch. A more slender form is also figured here. It is very rare but occurs with the other 

form. It has less pronounced chambers.

Cooper marl: 11861, Steep bluff on west side of Biggin 

Creek, Berkeley County, S.C.; 11863, three-fourths mile west 

of Old Biggin Church, Berkeley County, S.C.; 11862, 1 mile 

west of Givhans, Dorchester County, S.C.

Dentalina sp.? 

Plate 8, figures 15, 16

In the Cooper marl of South Carolina there is a 

large species which may be referred to Dentalina. 

The initial end has a short terminal spine, and the 

chambers are well separated and rounded. Although 

fragments occur at several stations, there are no complete specimens to give the full adult characters of 

the species.

Cooper marl: 11861, Steep bluff on west side of Biggin 

Creek, Berkeley County, S.C.; 11863, three-fourths mile west

of Old Biggin Church, Berkeley County, S.C.; 11862, 1 mile 

west of Givhans, Dorchester County, S.C.; 11857, 1 mile south 

of Moncks Corner, Berkeley County, S.C.

Genus NODOSARIA Lamarck, 1812 

Nodosaria latejugata Gumbel

Nodosaria latejugata, Gumbel, K. bayer. Akad. Wiss. Miinchen,

Cl. 2, Abb.., vol. 10, p. 619, pi. 1, fig. 32, 1868 (1870). 

Hantken, Magy. kir. fb'ldt. int. fivkb'nyve, vol. 4, p. 21,

pi. 2, figs. 6a-d, 1875 (1876). 

Cushman, Contr. Cushman Lab. Foram. Research, vol. 1,

p. 66, pi. 10, fig. 7, 1925.

Nodosaria budensis Hantken, Magy. kir. fo'ldt. int. liivkonyve, 

vol. 4, p. 23, pi. 2, fig. 10;-pi. 16, fig. 4, 1875 (1876).

Test elongate, subcylindrical, initial end with a single spine; chambers distinct, slightly inflated; sutures 

distinct, of clear shell material, slightly depressed; 

surface ornamented with a few very prominent longi-. 

tudinal costae, averaging about 10, continuous from 

initial to apertural ends, except occasionally the final 

chamber smooth; apertural end slightly prolonged; 

aperture radiate. Maximum length of American 

specimens 8.00 mm; breadth 0.50 mm.

Gumbel originally described this species from the 

upper Eocene of Bavaria, and Hantken figures it from 

the upper Eocene of Hungary. Our upper Eocene 

specimens are very similar, except that the microspheric form occurs as well. The species belongs 

to the general Nodosaria raphamw group, which 

occurs as many species from the Cretaceous to the 

present seas. A very large species belonging to this 

group is now living in the Philippine region. Nodosaria bacilloides Hantken is probably the same species.

Castle Hayne marl: 5611, Wilmington, N.C.

Cooper marl: 5531, Cooper River, S.C.; 6319, Ingleside, S.C.

Ocala limestone: 7097, Bainbridge, Ga.; 7719, Spring Mill 

Branch, Jenkius County, Ga.; 6768, Marianna, Fla.; 7378, 1^ 

miles southwest of Inverness, Fla.; 6722, three-fourths mile 

northwest of Monroeville, Ala.; 1/7, 1 mile east of Beck, Ala.; 

1/21, Covington County, Ala.; 1/26, Drewry, Ala.; 1/38, 3% 

miles north of Grove Hill, Ala.; 1/43, 1% miles southwest of 

Perdue Hill, Ala.; 7159, 4 miles southeast of Whatley, Ala.

Jackson formation: 2637, Cocoa, Ala.; 6457, 1% miles south 

of Shubuta, Miss.; 7203, 3V2 miles southeast of Cullomburg, Ala.

The only Cretaceous record for this species probably should be assigned to Nodosaria a/finis Beuss.

Nodosaria latejugata Gtimbel var. carolinensis Cushman

Plate 5, figures 10-13

Nodosaria latejugata Gtimbel var. carolinensis Cushman, Contr. 

Cushman Lab, Foram. Research, vol. 9, p. 5, pi. 1, fig. 

16, 1933.

Variety differing from the typical form in having 

the chambers more distinct and more inflated, the 

costae similar, but double the number in the typical, 

and the whole test larger.

Holotype of variety (no. 371540, U.S.N.M.) from 

Cooper marl, Cooper River, Charleston County, S.C.
22 UPPER EOCENE FOKAMINIFERA OF THE SOUTHEASTERN UNITED STATES

In the Carolina material this variety seems to be 

more characteristic than the typical form.

Cooper marl: 5531, Cooper River, S.C.; 11861, steep bluff on 

west side of Biggin Creek, Berkeley County, S.C.; 11863, threefourths mile west of Old Biggin Church, Berkeley County, 

S.C.; 11862, 1 mile west of Givhans, Dorchester County, S.C.; 

11857, 1 mile south of Moncks Corner, Berkeley County, S.C.

Jackson formation: 7203, 3% miles southeast of Cullomburg, 

Ala.

Ocala limestone: 1/38, 3% miles north of Grove Hill, Ala.

Nodosaria cookei Cushman 

Plate 9, figure 1

Nodosaria cookei Cushman, Contr. Cushman Lab. Foram. 

Research, vol. 9, p. 9, pi. 1, fig. 21, 1933.

Test small, composed of few chambers, well separated from one another; chambers very distinct, 

slightly longer than wide, circular in diameter; sutures 

indistinct; wall covered by very fine hispid spines, both 

on the body of the chambers and on the connections; 

aperture at the end of an elongate cylindrical neck. 

Length 0.75 mm; diameter 0.25 mm.

Holotype (no. 371541, U.S.N.M.) from Ocala limestone, station 1/21, Covington County, Ala.

This seems to be identical with a species figured 

without name by Von Schlicht,8 and apparently not 

named by Reuss.

The newly added chamber is set onto the neck of 

the preceding one near the apertural end, and a long, 

narrow connection is thus left between adjacent chambers. The surface ornamentation is very delicate. 

The species resembles somewhat N. conspurcata Reuss, 

but it has a finer ornamentation, and the chambers are 

more remote.

Ocala limestone: 1/21, Covington County, Ala.

Nodosaria ewaldi Reuss?

Plate 9, figure 2

There are fragments showing two chambers each 

from two stations, which from'their very cylindrical 

character and peculiar lack of incurving at the ends 

of the chambers, much resemble the figures of Reuss' 

N. ewaldi? They strongly resemble the figures given 

by Von Schlicht 10 and referred by Reuss to N. ewaldi.

Ocala limestone: 7719, Spring Mill Branch, Jenkins County, 

Ga.

Jackson formation: 6457, 1% miles south of Shubuta, Miss.

Cooper marl: 11861, steep bluff on west side of Biggin Creek, 

Berkeley County, S.C.

Nodosaria fissicostata (Gtimbel) 

Plate 5, figures 8, 9

Dentalina fissicostata Giimbel, K. bayer. Akad. Wiss. Miinchen, 

Cl. 2, Abh., vol. 10, p. 626, pi. 1, fig. 46, 1868 (1870).

8 Von Schlicht, E., Die Foraminiferen des Septarienthones von 

Pletzpuhl, pi. 6, flg. 28, 1870.

9 Deutsch. geol. Gesell. Zeitschr., vol. 3, pi. 3, flg. 2, 1851.

10 Von Schjicht, E., op. clt., pi. 7. figs. 8, 9.

Hantken, Magy. kir. foldt. int. fivkonyve, vol. 4, p. 37, pi. 3,

fig. 19, 1875 (1876); K. ungar. geol. Anstalt Mitt. Jahrb.,

vol. 4, p. 37, pi. 3, flg. 19, 1875 (1881). 

Nodosaria fissicostata, Cushman, Contr. Cusliman Lab. Foram.

Research, vol. 1, p. 66, pi. 10, fig. 8, 1925. 

Chapman and Parr, Linuean Soc. London Jour., Zoology,

vol. 36, p. 384, pi. 19, flg. 45, 1926. 

Cushman, Jour. Paleontology, vol. 1, p. 154, pi. 24, figs. 10,

11, 1927.

Test large, slightly curved, especially in the earlier 

portion, slightly tapering, initial end rounded; chambers fairly numerous, about as long as broad, earlier 

ones somewhat hidden by the ornamentation, later 

ones somewhat inflated and distinct; sutures between 

the later chambers slightly depressed and distinct, 

earlier ones indistinct; surface ornamented with numerous fine longitudinal costae, continuous over the 

length of the test except for a slight anastomosing or 

interpolation of new costae as the diameter of the test 

increases, about 20 to 25 costae in the last-formed 

chamber of adult specimens; aperture radiate, at the 

end of a conical projection. Maximum length, 6.00 

mm; maximum breadth, 0.60 mm.

Giimbel described this species originally from the 

Eocene of Bavaria, and Hantken records it from the 

Eocene of central Europe and Italy, with numerous 

localities. Hantken gives as measurements 3-6 mm 

for the length and 0.60-1.00 mm for the diameter of 

the last-formed chamber. It is an easily broken species, complete specimens being rare in the collections. 

Specimens occur in the Alazan clay of Mexico.

Cooper marl: 3595, Barnwell County, S.C.; 5531, Cooper 

River, S.C.; 6321, Baldock, S.C.

Ocala limestone: 6768, Marianna, Fla.; 7719, Spring Mill 

Branch, Jenkins County, Ga.; 1/7, 1 mile east of Beck, Ala.; 

1/43, 1% miles southwest of Perdue Hill, Ala.

Jackson formation: 2637. Cocoa, Ala.; 7203, 3% miles southeast of Cullomburg, Ala.

Genus LAGENA Walker and Jacob, 1798

Lagena laevis (Montagu)?

Plate 9, figures 3, 4

" Serpula .(Lagena) laevis ovalis " Walker and Boys, Testacea

minuta, p. 3, pi. 1, fig. 9, 1784.

Vermiculum laeve Montagu, Testacea Britannica, p. 524, 1803. 

Lagena laevis Williamson, Annals and Mag. Nat. Hist., 2d ser.,

vol. 1, p. 12, pi. 1, figs. 1, 2, 1848.

The specimen here figured is referred to the above 

pecies provisionally. The ornamentation if ever present has been eroded, and more specimens are necessary 

to place it more definitely. The specimen is from station 3595, the Cooper marl of Barnwell County, S.C. 

An additional specimen from the Cooper marl, station 

11857, Berkeley County, S.C., also figured, seems, to 

show traces of the typical costae at the base.

Cooper marl: 3595, Barnwell County, S.C.; 11857, 1 mile 

south of Moncks Corner, Berkeley County, S.C.
POLYMORPHINIDAE 23

Lagena acuticosta Reuss 

Plate 9, figures 5, 6

Lagena acuticosta Reuss, Akad. Wiss. Wien Sitzungsber., vol. 

44, pt. 1, p. 305, pi1. 1, fig. 4, 1861 (1862).

There are single specimens which may be referred 

to this species, at least until further material is 

obtained.

Ocala limestone: 7719, Spring Mill Branch, Jenkins County, 

Ga.

Cooper marl: 11S61, Steep bluff on west side of Biggin Creek, 

Berkeley County, S.C.; 11863, three-fourths mile west of Old 

Biggin Church, Berkeley County, S.C.; 11862, 1 mile west of 

Givhaus, Dorchester County, S.C.; 11857, 1 mile south of 

Moncks Corner, Berkeley County, S.C.

Lagena costata (Williamson)

Plate 9, figures 7, 8

Entosolenia costata Williamson, Recent Foraminifera of Great

Britain, p. 9, pi. 1, fig. 18, 1858. 

Lagena oostata Reuss, Akad. Wiss. Wien Sitzungsber., vol. 46,

pt. 1, p. 329, pi. 4, fig. 54. 1862 (1863).

The costate specimens figured may be referred to 

L. costata.

Ocala limestone: 7719, Spring Mill Branch, Jenkins County, 

Ga.

Cooper marl: 11861, Steep bluff on west side of Biggin Creek, 

Berkeley County,. S.C.; 11863, three-fourths mile west of Old 

Biggin Church, Berkeley County, S.C.; 11862, 1 mile west of 

Givhans, Dorchester County, S.C.; 11857, 1 mile south of 

Moncks Corner, Berkeley County, S.C.

The only record for this species from the Cretaceous 

is without a figure.

Lagena orbignyana (Seguenza) var. semiconcentrica 

Cushman

Plate 9, figure 9

Lagena orbignyana, (Seguenza) var. semiconcentrica Cushman,. 

Contr. Cushman Lab. Foram. Research, vol. 9, p. 10, 

pi. 1, fig. 22, 1933.

This variety differs from var. concentrica Sidebottom in having the central part of the faces of the test 

without ornamentation. There are numerous concentric ridges at the sides, but they are slightly interrupted at the base and are wanting in the center. 

Length 0.45 mm.

Holotype (no. 371542, U.S.N.M.) from Castle 

Hayne marl, station 5611, Wilmington, N.C.

Castle Hayne marl: 5611, Wilmington, N.C.

Lagena hexagona (Williamson)

Plate 9, figure 10

The single specimen figured here may be referred to 

Williamson's species.

Cooper marl: 11863, three-fourths mile west of Old Biggin 

Church, Berkeley County, S.C.

Lagena sp.? 

Plate 9, figure 11

The figured specimen shows a form with several 

basal spines, that occurs in the Cooper marl of South 

Carolina, but the single specimen is not enough to 

place the species with certainty.

Cooper marl: 5531, Charleston, S.C.

Family POLYMORPHINIDAE

Genus GUTTULINA D'Orbigny, 1826

Guttulina problema D'Orbigny

Plate 9, figure 12

Guttulina problema D'Orbigny, Annales sci. nat., vol. 7, p. 266,

no: 14, 1826. 

Cushman and Schenck, California Univ. Dept. Geol. Sci.

Bull., vol. 17, p. 310, pi. 43, figs. 9-11, 1928. 

Cushman and Ozawa, U.S. Nat. Mus. Proc., vol. 77, art.

6, p. 19, pi. 2, figs. 1-6; pi. 3, figs. la-cr 1930. 

Howe and Wallace, Louisiana Dept. Conservation Geol.

Bull. 2, p. 49, pi. 7, fig. 9, 1932. 

Polymorphina problema/ D'Orbigny, Modeles, no. 61, 1826.

H. B. Brady, Challenger Rept, Zoology, vol. 9, p. 568, pi.

72, fig. 20; pi. 73, fig. 1, 1884. 

Jones, Foraminifera of the Crag, pt. 3, p. 267, pi. 1, fig.

64; pi. 5, fig. 23; pi. 6, figs. 12a, b, 1896. 

Burrows and Holland, Geol. Assoc. Proc., vol. 15, p. 46,

pi. 2, fig. 17, 1897. 

Weller, New Jersey Geol. Survey, Paleontology, vol. 4, p.

253, pi. 3, figs. 27, 28, 1907. 

Cushman, U.S. Nat. Mus. Bull. 100, vol. 4, p. 264, pi. 54,

figs. 3, 4, 1921; U.S. Geol. Survey Prof. Paper 133, p.

33, pi. 5, fig. 6, 1923. 

Chapman, New Zealand Geol. Survey Pal. Bull. 11, p. 68,

pi. 5, figs. 10-12, 1926. 

Polymorphina (Guttulina) problema Andreae, GeoJ. SpecialKarte Elsass-Lothringen Abh., vol. 2, pt. 3, p. 118, pi. 9,

figs. 21a-c, 1884. 

PolymorpMna problema var. Hosius, Naturhist. Ver. preuss.

Rheinlande Verb., vol. 50, p. 103, pi. 2, figs. 2a-c (not

figs. 3-5), 1893. 

Guttulina communis D'Orbigny, Annales sci. nat., vol. 7, p.

266, no. 15, pi. 12, figs. 1-4, Modeles, no. 62, 1826. 

Reuss, in Geinitz, Grundriss der Versteinerungskunde, p.

669, pi. 24, fig. 82, 1845-46. 

PolymorpMna communis Roemer, Neues Jahrb., 1838, p. 385,

pi. 3, fig. 29. 

H. B. Brady, Parker, and Jones, Linnean Soc. London

Trans., vol. 27, p. 224, pi. 39, figs. lOa, b, 1869. 

H. B. Brady, Challenger Rept., Zoology, vol. 9, p. 568, pi.

72, fig. 19, 1884. 

Flint, U.S. Nat. Mus. Rept. for 1897, p. 319, pi. 67, fig. 6

(part), 1899.

Bagg, U.S. Geol. Survey Bull. 88, p. 60, pi. 6, fig. 2, 1898. 

Fornasini, Accad. sci. 1st. Bologna Mem., ser. 5, vol. 8,

p. 33, fig. 37, 1900. 

Weller, New Jersey Geol. Survey, Paleontology, vol. 4, p.

248, pi. 3, fig. 18, 1907. 

Bagg, U.S. Geol. Survey Bull. 513, p. 68, pi. 21, figs. 7a, b,

13-15, 1912. 

Cushman, U.S. Nat. Mus. Bull. 71, pt. 3, p. 87, pi. 37, fig. 7,

1913.
24 UPPER EOCENE FORAMINTFERA OF THE SOUTHEASTERN UNITED STATES

Polymorphina lactea Walker and Jacob var. communis Williamson, Recent Foraminifera of Great Britain, p. 72,

pi. 6, figs. 153-155, 1858. 

Polymorphma (Guttulina) lata Egger, Neues Jahrb., 1857, p.

288, pi. 13, figs. 22-24. 

Guttulina cretacea Alth, Haidinger's Naturwiss. Abh., vol. 3,

p. 262, pi. 13, fig. 14, 1850. 

Reuss, Haidinger's Naturwiss. Abh., vol. 4, p. 28, pi. 4, fig.

10, 1851. 

Polymorphma cretacea Egger, K. bayer. Akad. Wiss. Munchen,

01. 2, Abh., vol. 21, pt. 1, p. 127, pi. 17, figs. 12, 13, 1899. 

Globulina irregularis Terquem, Soc. g£ol. France M6m., s6r. 3,

vol. 1, p. 44, pi. 4 (9), figs. 13, 14, 1878. 

Polymorphina lactea Sidebottom, Manchester Lit. Philos. Soc.

Mem. and Proc., vol. 5, no. 9, p. 9, pi. 2, fig. 11, 1907. 

Franke, Geol. palaeont. Inst. Univ. Greifswald Abh., vol. 6,

p. 77, pi. 6, fig. 18, 1925; Danmarks geol. Unders0gelse, 2.

Raekke, Nr. 46, p. 34, pi. 3, fig. 13, 1927.

Globulina gibba var. glomula Fornasini, Accad. sci. 1st. Bologna Mem., ser. 5, vol. 9, p. 68, fig. 20 (in text), 1902.

Test broadly fusiform, acute at the apertural end, 

more or less rounded at the initial end in the megalospheric form, rather rounded at the base in the 

microspheric form; chambers elongated, more or less 

inflated, arranged in a clockwise, quinqueloculine 

series, each succeeding chamber slightly removed from 

the base; sutures depressed, very distinct; wall rather 

thick, smooth; aperture radiate. Length 0.50-1.25 

mm; breadth 0.40-1.25 mm; thickness 0.28-0.70 mm.

For further notes in regard to this species, see Gushman and Ozawa in the paper cited above.

Cooper marl: 5531, Charleston, S.C.

Jackson formation: Tarkiln Creek, half a mile above Neches 

River, Trinity County, Tex.

Guttulina irregularis (D'Orbigny)

Plate 9, figures 13-16

Globulina irregularis D'Orbigny, Foraminiferes fossiles du

bassin tertiaire de Vienne, p. 226, pi. 13, figs. 9, 10, 1846. 

Cushman and Thomas, Jour. Paleontology, vol. 3, p. 177,

pi. 23, figs. 2a-c, 1929. 

Quttulma irregularis Cushman and Ozawa, U.S. Nat. Mus.

Proc., vol. 77, art. 6, p. 25, pi. 3, figs. 4, 5; pi. 7, figs.

1, 2, 1930. 

Howe and Wallace, Louisiana Dept. Conservation Geol.

Bull. 2, p. 48, pi. 8, fig. 8, 1932. 

Guttulina dilatata Reuss, K. Akad. Wiss. Wien Denkschr., vol.

1, p. 378, pi. 48, fig. 11, 1850. 

Guttulina problems D'Orbigny (not D'Orbigny, 1826), Foraminif£res fossiles du bassin tertiaire de Vienne, p. 224,

pi. 12, figs. 26-28, 1846. 

Reuss, in Geinitz, Grundriss der Versteinerungskunde, p.

669, pi. 24, fig. 83, 1845-46. 

Globulina guttula Reuss, Deutsch. geol. Gesell. Zeitschr., vol. 3,

p. 82, pi. 6, fig. 46, 1851. 

Guttulina semiplana Reuss, Deutsch. geol. Gesell. Zeitschr., vol.

3, p. 82, pi. 6, fig. 48, 1851. 

Guttulina centrata Terquem, Soc. g6ol. France M6m., se'r. 3,

vol. 1, p. 46, pi. 4 (9), figs. 25a-26, 1878. 

Polymorphina byramensis Cushman, U.S. Geol. Survey Prof.

Paper 129, p. 94, pi. 17, figs. 2a, b, 1922; Prof. Paper 133,

p. 31, pi. 5, figs. 1-5, 1923.

Guttulina byramensis Cushman and Schenck, California Univ. 

Dept. Geol. Sci. Bull., vol. 17, p. 309, pi. 43, figs. 6-8,1928.

Test oval to subdeltoidal, equilaterally triangular 

with rounded sides and angles, except the acute apertural end; chambers more or less angular, elongated, 

arranged in a clockwise, quinqueloculine series, each 

succeeding chamber except the last one or two chambers in full-grown specimens coming down to the base; 

sutures depressed, distinct; wall smooth but in fullgrown specimens often having the last small chamber 

with spines or covered with fistulose tubes; aperture 

radiate. Length 0.45-1.40 mm; breadth 0.30-1.20 mm; 

thickness 0.20-0.75 mm.

This is a common species in both the lower Oligocene and the Eocene of the Gulf Coastal Plain of the 

United States. It is one of the most common species 

of the whole family and is very widely distributed in 

many parts of the.world.

Castle Hayne marl: 5611, Wilmington, N.C.

Ocala limestone: 7098, Red Bluff, Flint River, Ga.; 7189, 

Rich Hill, Ga.; 7719, Spring Mill Branch, Jenkins County, 

Ga.; 6768, Marianna, Fla.; 6799, Alachua Sink, Gainesville, 

Fla.; 1/19, near Blue Springs, Jackson County, Fla.; 6706, 

Oven Bluff, Tombigbee River, Ala.; 6722, three-fourths mile 

northwest of Monroeville, Ala.; 6739, 6740, mouth of Robinsons Mill Creek, Ala.; 6741, 3% miles northeast of Brooklyn, 

Ala.; 7159, 4 miles southeast of Whatley, Ala.; 1/21, Covington County, Ala.; 1/25, 1 mile east of Harts Bridge, Ala.; 1/26, 

Drewry, Ala.; 1/39, Pea River at Geneva, Ala.; 1/43, 1% miles 

southwest of Perdue Hill, Ala.

Jackson formation: 2637, Cocoa, Ala.; 7203, 3% miles southeast of Cullomburg, Ala.; 6457, 1% miles south of Shubuta, 

Miss.; 6463, pumping station, Jackson, Miss.; 6472, Yazoo City, 

Miss.

Guttulina spicaeformis (Roemer)

Plate 9, figure 17; plate 10, figures 9, 10

Polymorphina spicaeformis Roemer, Neues Jahrb., 1838, p. 386,

pi. 3, fig. 31. 

Guttulina spicaeformis Cushman and Ozawa, U.S. Nat. Mus.

Proc., vol. 77, art. 6, p. 31, pi. 5, figs. 1, 2, 1930. 

Ellisor, Am. Assoc. Petroleum Geologists Bull., vol. 17,

no. 11, pi. 7, fig. 3, 1933. 

Polymorphina austriaca D'Orbigny var. io Cushman and

Applin, Am. Assoc. Petroleum Geologists Bull., vol. 10,

p. 174, pi. 9, figs. 6, 7, 1926. 

Guttulina plancii D'Orbigny, Voyage dans J'Amerique m6ridionale, vol. 5, pt. 5, For am in if fires, p. 60, pi. 1, fig. 5,

1839. 

Polymorphma uviformis Reuss, Deulsch. geol. Gesell. Zeitschr.,

vol. 7, p. 289, pi. 11, fig. 5, 1855.

Test fusiform, initial end rounded, apertural end 

acute, margin slightly lobulate; chambers clavate, but 

little embracing, arranged in a contraclockwise, quinqueloculine series, each succeeding chamber removed 

from the base; sutures depressed, distinct; wall 

smooth; aperture radiate. Length 0.35-0.75 mm; 

breadth 0.20-0.35 mm; thickness 0.12-0.25 mm.

This is a common species in the upper Eocene of 

the Coastal Plain region of the United States but is 

not as abundant as the preceding species.
POLYMOBPHINIDAB 25

Castle Hayne marl: 5611, Wilmington, N.C.

Cooper marl: 11861, steep bluff on west side of Biggin 

Creek, Berkeley County, S.C.; 6321, Baldock, S.C.

Ocala limestone: 6768, Marianna, Fla.; 6730, Double 

Branches Creek, Ala.; 6738, 6% miles north of Brooklyn, Ala.

Jackson formation: 6457,1% miles south of Shubuta, Miss.; 

6463, pumping station, Jackson, Miss.; 6471, Garlands Creek, 

Miss.; 6646, Hays Chapel, Wayne County, Miss.; 4787, threefourths mile below Robinsons Ferry, Sabine County, Tex.; on 

Stoval Creek, east of Diboll, and in oil-well drillings, Tex.

Genus QLOBUIINA D'Orbigny, 1826 

Globulina gibba D'Orbigny

Plate 9, figure 18

Globulina g-ibba D'Orbigny, Annales sci. nat, vol. 7, p. 266, no.

10, Modeles, no. 63, 1826; Foraminiferes fossiles du

bassin tertiaire de Vienne, p. 227, pi. 13, figs. 13, 14, 1846. 

Terquem, Essai sur le classement des animaux de Dunkerque, p. 38, pi. 5, fig. 15,1875; Soc. g£ol. France Mem., ser.

3, vol. 1, p. 43, pi. 4 (9), figs. 1-5, 1878; vol. 2, p. 130,

pi. 13 (21), figs. 22-27, 1882. 

Jones and Chapman, Linnean Soc. London Jour., Zoology,

vol. 25, pp. 509, 515, figs. 6, 7, 40 (in text), 1896. 

Cushman and Ozawa, U.S. Nat. Mus. Proc., vol. 77, art 6,

p. 60, pi. 16, figs. 1-4, 1930. 

Howe and Wallace, Louisiana Dept. Conservation Geol.

Bull. 2, p. 46, pi. 8, figs, lla, b, 1932. 

Polymorphism (Globulina) gibba Egger, Neues Jahrb., 1857, p.

289, pi. 13, figs. 1-4. 

Andreae, Geol. Special-Karte Elsass-Lothringen Abb.., vol.

2, pt. 3, pp. 117, 141, pi. 9, figs. 10-12, 13a, b, 1884. 

Polymorphina gibba H. B. Brady, Parker, and Jones, Linnean

Soc. London Trans., vol. 27, p. 216, pi. 39, figs. 2a, b,

1870. 

Reuss, Akad. Wiss. Wien Sitzungsber., vol. 62, pt. 1, p.

485, 1870. 

Von Schlicht, Die Foraminiferen des Septarienthones von

Pietzpuhl, pi. 27, figs. 1-6, 16-18; pi. 26, figs. 31-34,

1870. 

H. B. Brady, Challenger Rept, Zoology, vol. 9, p. 561, pi.

71, figs. 12a, b, 1884. 

Sherborn and Chapman, Roy. Micr. Soc. Jour., ser. 2, vol.

6, p. 755, pi. 16, fig. 5, 1886. 

Goes, K. svensk. vet. akad. Handl., vol. 25, no. 9, p. 55,

pi. 9, figs. 520-526, 1894. 

Jones, Foraminifera of the Crag, pt. 3, p. 253, pi. 1, figs.

49-51, 1896. 

Jones and Chapman, Linnean Soc. London Jour., Zoology,

vol. 25, pp. 513, 514, figs. 31-33 (in text), 1896. 

Bagg, Maryland Geol. Survey, Eocene, p. 248, pi. 63, fig. 12,

1901; Miocene, p. 477, pi. 133, fig. 4, 1904. 

Weller, New Jersey Geol. Survey, Paleontology, vol. 4, p.

250, pi. 3, fig. 21, 1907. 

Sidebottom, Manchester Lit. Philos. Soc. Mem. and Proc.,

no. 9, vol. 51, p. 10, pi. 2, figs. 15-17, 1907. 

Paalzow, Offenbacher Ver. Naturkunde Ber., 1912, p. 67, p.

1, fig. 10. 

Cushman, U.S. Geol. Survey Bull. 676, pp. 10, 52, pi. 2, fig.

4; pi. 11, fig. 5, 1918; Prof. Paper 129, pp. 93, 94; pi. 17,

fig. 3; pi. 18, figs. 3a, b, 1922. 

Plummer, Texas Univ. Bull. 2644, p. 122, pi. 6, figs. Sa, b,

1927. 

Stadnichenko, Jour. Paleontology, vol. 1, p. 230, pi. 38, figs.

20-22, 1928.

Guttulina (Globulina) gibba Cushman, Soc. scl. Seine-et-Oise

Bull., s<§r. 2, vol. 9, p. 50, pi. 1, figs. 4a-c, 5a, b, 1928

(separate, p. 4, pi. 1, figs. 4a-c, 5a, b). 

Globulina globosa Reuss (not Minister), in Geinitz, Grundriss

der Versteinerungskunde, p. 669, pi. 24, fig. 85, 1845-46;

Akad. Wiss. Wien Sitzungsber., vol. 44, pt. 1, p. 318, pi. 3,

fig. 3, 1861 (1862). 

Globulina tubulosa D'Orbigny, Foraminiferes fossiles du bassin

tertiaire de Vienne, p. 228, pi. 13, figs. 15, 16, 1846. 

Polymorphina (Globulvna) gibba D'Orbigny var. ovoidea Egger,

Neues Jahrb., 1857, p. 289, pi. 13, figs. 5-7. 

PolymorpMna (Globulina) gibba D'Orbigny var. subgibba

Egger, Neues Jahrb., 1857, p. 289, pi. 13, figs. 8-10. 

PolymorpMna (Globulina) gibba D'Orbigny var. pirula Egger,

Neues Jahrb., 1857, p. 290, pi. 13, figs. 11, 12. 

Globulina amplectens Bornemann (not Reuss), Deutsch. geol.

Gesell. Zeitschr., vol. 12, p. 160, pi. 6, figs. 12o-o, 1860. 

Polymorphina gibba (D'Orbigny) var. orbicularis Karrer, Akad.

Wiss. Wien Sitzungsber., vol. 58, Abt. 1, p. 174, pi. 4,

fig. 8, 1868. 

Globulina subgibba Gurnbel, K. bayer. Akad. Wiss. Miinchen,

Cl. 2, Abh., vol. 10, p. 645, pi. 2, fig. 79, 1870. 

Globulina ovalis Terquem, Essai sur le classement des animaux

de Dunkerque, p. 77, pi. 10, fig. 2, 1876. 

Globulina oviformis Terquem, Soc. geol. France Mem., ser. 3,

vol. 1, p. 44, pi. 4(9), figs. 9-12, 1878. 

Globulina transversa Terquem, Soc. geol. France Mem., s6r. 3,

vol. 2, p. 129, pi. 13 (21), figs. 17-21, 1882. 

Guttulina, ponderosa Terquem, Soc. geol. France Mem., se'r. 3,

vol. 2, p. 135, pi. 14 (22), figs, la, b, 1882. 

Guttulina gravida Terquem and Terquem (not Terquem, 1878),

Soc. zool. France Bull., vol. 11, p. 334, pi. 11, fig. 21, 1886. 

Polymorphina inflata Terquem and Terquem (not Terquem,

1878), Soc. zool. France Bull., vol. 11, p. 335, pi. 11,

fig. 23, 1886.

PolymorpMna lactea (Walker and Jacob) var. diffusa Cushman, U.S. Nat Mus. Bull. 71, pt. 3, p. 84, pi. 41, fig. 8,

1913.

Test globular to subglobular, transverse section 

almost circular; chambers few, inflated, rounded, arranged in a nearly triserial series; sutures not depressed, generally clear; wall smooth, translucent, 

often with fistulose tubes, especially at apertural end; 

aperture radiate. Length 0.45-1.10 mm; breadth 0.40- 

0.90 mm; thickness 0.40-0.90 mm.

This is a very common species widely distributed in 

the Coastal Plain, especially in the lower Oligocene 

and the Eocene.

Cooper marl: 6319, Ingleside, S.C.; 11861, steep bluff on 

west side of Biggin Creek, Berkeley County, S.C.; 11863, threefourths mile west of Old Biggin Church, Berkeley County, S.C.; 

11862, 1 mile west of Givhans, Dorchester County, S.C.; 11857, 

1 mile south of Moncks Corner, Berkeley County, S.C.; 6321, 

Baldock, S.C.

Ocala limestone: 7097, Bainbridge, Ga.; 7189, Rich Hill, Ga.; 

7719, Spring Mill Branch, Ga.; 6768, Marianna, Fla.; 6804, 

Ocala, Fla.; 7364, 1% miles west of Groom, Fla.; 1/19, near 

Blue Springs, Jackson County, Fla.; 1/8, Beck, Ala.; 1/38, 

3% miles north of Grove Hill, Ala.; 6730, Double Branches 

Creek, Ala.; 6740, mouth of Robinsons Mill Creek, Ala.; 6742, 

Powells Landing, Sepulga River, Ala.

Jackson formation: 6463, Jackson, Miss., near pumping station ; 6471, Garlands Creek, Miss.
26 UPPER EOCENE FORAMINIFERA OP THE SOUTHEASTERN UNITED STATES

Globulina gibba D'Orbigny var. globosa (Yon Minister)

Plate 9, figure 21

Polymorphina globosa Von Munster, in Roemer, Neues Jahrb.,

1838, p. 386, pi. 3, flg. 33. 

Reuss, Versteinerungen der bohmischen Kreideformation,

p. 40, pi. 13, fig. 82, 1845. 

Bgger, K. bayer. Akad. Wiss. Miinchen, Cl. 2, Abh., vol. 21,

pt. 1, p. 129, pi. 17, fig. 26, 1899. 

Globulina ffibba D'Orbigny var. globosa Cushrnan and Ozawa,

U.S. Nat. Mus. Proc., vol. 77, art. 6, p. 64, pi. 17, figs.

8, 9, 1930. 

Howe and Wallace, Louisiana Dept. Conservation Geol.

Bull. 2, p. 46, pi. 8, fig. 10, 1932. 

PolymorpMna aouta Roemer, Neues Jahrb., 1838, p. 386, pi. 3,

fig. 36. 

Globulina acuta Reuss, Akad. Wiss. Wien Sitzungsber., vol.

18, p. 245, pi. 6, flg. 62, 1855 (1856). 

Globulina aequalis D'Orbigny, Foraminiferes fossiles du bassin

tertiaire de Vienne, p. 227, pi. 13, figs. 11, 12, 1846. 

Polymorphina equalis Cushman, U.S. Geol. Survey Prof. Paper

129, p. 132, pi. 31, fig. 3, 1922; U.S. Nat. Mus. Bull. 104,

pt. 4, p. 149, pi. 40, fig. 3, 1923. 

Polymorphina gibba var. aequalis H. B. Brady, Parker, and

Jones, Linnean Soc. London Trans., vol. 27, p. 216, pi.

39, figs. 2c, d, 1870. 

Globulina tivrbinata Terquem. Soc. g6ol. France M<5in., ser. 3,

vol. 1, p. 43, pi. 4 (9), figs. 6-8, 1898.

Test subglobular to broadly oval, rather convex on 

one side in the microspheric form, more or less equally 

compressed in the megalospheric form, slightly produced at the apertural end; chambers more or less 

compressed, generally as many as 1 in the microspheric, 4 in the megalospheric form, the later chambers more embracing on one side, arranged in an 

almost triserial series, each succeeding chamber more 

or less above the base; sutures not depressed, distinct; 

wall smooth, thick, often with fistulose tubes; aperture 

radiate. Length 0.6S-0.90 mm; breadth 0.40-0.65 mm; 

thickness 0.20-0.35 mm.

This is a compressed modification of Globulina 

gib^a, and there is a certain range of variation in its 

test.

This variety seems to be rare in the Eocene, so far 

as the material examined is concerned.

Castle Hayne marl: 5611, Wilmington, N.C.

Cooper marl: Ashley marl works, Dorchester County, S.C.; 

11861, steep bluff on west side of Biggin Creek, Berkeley 

County, S.C.

Ocala limestone: 7189, Rich Hill, Ga.

Globulina gibba D'Orbigny var. tuberculata D'Orbigny 

Plate 9, figures 19, 20

Globulina tuberculata D'Orbigny, Foraminiferes fossiles du

bassin tertiaire de Vienne, p. 230, pi. 13, figs. 21, 22,

1846. 

Terquem, Soc. g6ol. France Mem., s6r. 3,. vol. 2, p. 132, pi.

13 (21), figs. 33, 34, 1882. 

Globulina gibba D'Orbigny var. tuberculata Cushman and

Ozawa, U.S. Nat. Mus. Proc., vol. 77, art. 6, p. 68, pi. 17,

figs. 6, 7, 1930.

Polymorphina (Globulina) tubertfulata Egger, Neues Jahrb.,

1857, p. 292, pi. 14, figs. 7, 8. 

Polymorphina tuberculata H. B. Brady, Parker, and Jones,

Linnean Soc. London Trans., vol. 27, p. 242, pi. 41, figs.

35a-d, 1870. 

Jones, Foraminifera of the Crag, pt. 3, p. 273, pi. 5, fig. 29,

1896. 

Cushman, Am. Assoc. Petroleum Geologists Bull., vol. 10,

p. 603, pi. 20, fig. 5, 1926. 

Globulina sp'inosa D'Orbigny, Foraminiferes fossiles du bassin

tertiaire de Vienne, p. 230, pi. 13, figs. 23, 24, 1846. 

Terquem, Soc. g<5ol. France Mem., s6r. 3, vol. 2, p. 132, pi.

13 (21), fig. 35, 1882. 

Polymorphina (Globulina) spinosa Egger, Neues Jahrb., 1857,

p. 292, pi. 14, figs. 9, 10.

Polymorphina spinosa H. B. Brady, Parker, and Jones, Linnean Soc. London Trans., vol. 27, p. 243, pi. 42, figs. 36a,

b, 1870. 

Balkwill and Wright, Roy. Irish Acad. Sci. Trans., vol.

28, p. 347, pi. 12, fig. 27, 1885. 

Cushman, U.S. Geol. Survey Prof. Paper 129, p. 133, pi.

31, fig. 5, 1922; U.S. Nat. Mus. Bull. 104, pt. 4, p. 157,

pi. 41, figs. 1, 13, 1923. 

Globulina* asperula Giirnbel, K. bayer. Akad. Wiss. Munchen,

Cl. 2, Abb., vol. 10, p. 646, pi. 2, figs. 81a, b, 1870. 

Polymorphina frondiformis Searles Wood var. brevis Jones,

Foraminifera of the Crag, pt. 3, p. 271, pi. 7, fig. 20, 1896.

Variety ornamented with strong spines of unequal 

size and irregularly spaced. Length with spines 0.70- 

0.90 mm; breadth 0.60-0.80 mm; thickness 0.50-0.70 

mm.

Hare specimens occur in the Jackson, which may be 

referred to this variety. The spines are variable.

Cooper marl: 6319, Ingleside, S.C.

Ocala limestone: 1/21, Covington County, Ala.

Globulina inaequalis Reuss

Plate 9, figure 22

Globulina inaequalis Reuss, K. Akad. Wiss. Wien Denkschr.,

vol. 1, p. 377, pi. 48, fig. 9, 1850. 

Cushman and Ozawa, U.S. Nat. Mus. Proc., vol. 77, art. 6,

p. 73, pi. 18, figs. 2-A, 1930. 

Globulina discreta Reuss, K. Akad. Wiss. Wien Denkschr., vol.

1, p. 378, pi. 48, fig. 10, 1850. 

Globulina amplectens Reuss, Deutsch. geol. Gesell. Zeitschr.,

vol. 3, p. 81, pi. 6, fig. 44, 1851. 

Globulina inftata Reuss, Deutsch. geol. Gesell. Zeitschr., vol. 3,

p. 81, pi. 6, fig. 45, 1851. 

Globulina amygdaloides Reuss, Deutsch. geol. Gesell. Zeitschr.,

vol. 3, p. 82, pi. 6, fig. 47, 1851. 

Polymorphina amygdaloides Reuss (not Reuss, 1851), Akad.

Wiss. Wien Sitzungsber., vol. 62, pt. 1, p. 486, 1870. 

Von Schlicht, Die Foraminiferen des Septarienthones von

Pietzpuhl, pi. 27, figs. 7-9, 10-12, 1870.. 

Guttulina diluta Bornemann, Deutsch. geol. Gesell. Zeitschr.,

vol. 12, p. 160, pi. 6, fig. lla-c, 1860. 

Polymorphina pauperata Terquem (part), Soc. g6ol. France

Mem., ser. 3, vol. 1, p. 38, pi. 3 (8), figs. 12, 13 (not

figs. 11, 14-19), 1878. 

Globulina translueida D'Orbigny, Annales sci. nnt., vol. 7, p.

267, no. 25, 1826. 

Terquem, Soc. ggol. France M6in., s£r. 3, vol. 2, p. 131, pi.

13 (21), fig. 31a, b, 1882. 

Polymorphina translueida Fornasini, Accad. sci. 1st. Bologna

Mem., ser. 5, vol. 10, p. 52, fig. 51, 1902.
POLYMOBPHINIDAB 27

Test ovate, more or less compressed, broadly rounded 

at the base, tapering toward the apex; chambers few, 

inflated, much overlapping, arranged in a nearly triserial series; sutures very slightly if at all depressed, 

distinct; wall smooth, translucent; aperture radiate. 

Length 0.50-0.85 mm; breadth 0.45-0.85 mm; thickness 0.25-0.50 mm.

This species is not common in the Eocene of the 

Coastal Plain area and also occurs rarely in the lower 

Oligocene and in the Miocene.

Cooper marl: 11857, 1 mile south of Moncks Corner, Berkeley 

County, S.C.

Jackson formation: 4787, three-fourths mile below Robinsons 

Ferry, Sabine River, Sabine County, Tex.

Globulina minuta (Roemer)

Plate 9, figure 23

Polymorphina, minu:a Roemer, Neues Jahrb., 1838, p. 386, pi. 3,

flg. 35. 

Reuss, Akad. Wiss. Wien Sitzungsber., vol. 62, pt. 1, p. 486,

1870. 

. Von Schlicht, Die Foraminiferen cles Septarienthones von

Pietzpuhl, pi. 27, figs. 13-15; pi. 25, figs. 51-56, 1870. 

Boruemann', Deutsch. geol. Gesell. Zeitschr., vol. 7, p. 344,

pi. 17, flg. 3, 1855. 

Glob'itiina m'vnuta Cushman and O/awa, U.S. Nat. Mus. Proc.,

vol. 77, art. 6, p. 83, pi. 20, Jigs. 3, 4, 1930. 

Polymorphina (Guttulina) aff. P. minima Audreae, Geol.

Special-Karte Elsass-Lothringen Abh., vol. 2, pt. 3, p.

118, pi. 9, flg. 16, 1884. 

PolyrnorpMna similis Reuss, Akad. Wiss. Wien Sitzungsber.,

vol. 18, p. 249, pi. 7, fig. 79, 1855 (1856). 

Polymorphina sororia Reuss, Akad. Wiss. Wien Sitzungsber.,

vol. 48, pt. 1, p. 57, pi. 7, figs. 72(7), 73, 74, 1863; Acad.

roy. sci. Belgique Bull., s6r. 2, vol. 15, p. 151, pi. 2,

flg. 25 (not figs. 26-29), 1863. 

Polymorphina gracilis Reuss, Akad. Wiss. Wien Sitzungsber.,

vol. 62, pt. 1, p. 486, 1870. 

Von Schlicht, Die Foramiuiferen des Septarienthones von

Pietzpuhl, pi. 31, figs. 34-37, 42, 45-49; pi. 32, figs. 5-16,

27, 28, 1870. 

Polymorphina acuta Reuss, Akad. Wiss. Wien Sitzungsber.,

vol. 62, pt. 1, p. 486, 1870. 

Voii Schlicht, Die Foraminiferen des Septarienthones von

Pietzpuhl, pi. 27, figs. 19-21; pi. 29,. figs. 15, 16, 43-46,

1870. 

Polymorphina lactea Burrows, Sherborn, and Bailey (not

Walker and Jacob), Roy. Micr. Soc. Jour., 1890, p. 561,

pi. 11, ligs. 9, 10. 

ISgger, K. buyer. Akad. Wiss. Munchen, Cl. 2, Abh., vol. 18,

p. 308, pi. 9, figs. 8, 14, 15, 1893. 

Mills, Hull Sci. and Field Nat. Club Trans., vol. 1, p. 149,

pi. 11, flg. 33, 1900. 

Polymorphina lactea elongate variety, Weller, New Jersey Geol.

Survey, Paleontology, vol. 4, p. 251, pi. 3, figs. 24-25,

1907.

Test fusiform, pointed at both ends, circular to elliptical in section; chambers few, elongate, each some 

distance from the base; sutures distinct, not depressed; 

wall smooth; aperture radiate, pointed. Length 0.50- 

0.70 mm; breadth 0.20-0.35 mm; thickness 0.20- 

0.35 mm.

Ocala limestone: 6738, 6% miles north of Brooklyn, Ala.

Globulina miinsteri (Reuss)

Plate 9, figure 25

Polymorphina mumteri Reuss, Akad. Wiss. Wien Sitzungsber.,

vol. 18, p. 249, pi. 8, fig. 80, 1855 (1856). 

Globulina miinsteri Cushman and Ozawa, U.S. Nat. Mus. Proc.,

vol. 77, art. 6, p. 85, pi. 22, figs. 3a-c, 1930. 

Polymorphina ovulum Reuss, Akad. Wiss. Wien Sitzungsber.,

vol. 18, p. 250, pi. 8, fig. 83, 1855 (1856). 

Polymorphina* amyydaloides Reuss (not Reuss, 1851), Akad.

Wiss. Wien Sitzungsber., vol. 18, p. 250, pi. 8, fig. 84,

1855 (1856). 

Burrows and Holland, Geol. Assoc. Proc., vol. 15, p. 46, pi.

2, fig. 18, 1897. 

Polymorphina oommunis D'Orbigny var. etrusca Fornasini,

Accad. sci. 1st. Bologna Mem., ser. 5, vol. 9, p. 70, fig.

23 (in text), 1900-1902. 

Polymorphina obtusa D'Orbigny, Annales sci. nat, vol. 7, p.

265, no. 1, 1826. 

Fornasini, Soc. geol. italiana Boll., vol. 19, p. 146, fig. 4

(in text), 1900. 

PolymorpJiina gibba Burrows, Sherborn, and Bailey, Roy. Micr.

Soc. Jour., 1890, p. 561, pi. 11, fig. 13.

Specimens occur at several stations in the Coastal 

Plain region, but the species is not at all common.

Ocala limestone: 7719, Spring Mill Branch, Jenkins County, 

Ga.; 6730, Double Branches Creek, Ala.; 6741, 3V2 miles northeast of Brooklyn, Ala.; 1/7, 1 mile east of Beck, Ala.

Globulina rotundata (Bornemann)

Plate 9, figures 24a-c

Guttulina rotundata Bornemann, Deutsch. geol. Gesell. Zeitschr., vol. 7, p. 346, pi. 18, fig. 3, 1855. 

Globulina rotundata Cushman and Ozawa, U.S. Nat. Mus. Proc.,

vol. 77, art. 6, p. 86, pi. 21, figs. 3, 4, 1930. 

Howe and Wallace, Louisiana Dept. Conservation Geol.

Bull. 2, p. 47, pi. 15, fig. 4, 1932. 

Polymorphina rotundata H. B. Brady, Parker, and Jones, Linnean Soc. London Trans., vol. 27, p. 234, pi. 40, figs.

19a-e, woodcuts k, 1, m, 1870. 

H. B. Brady, Annals and Mag. Nat. Hist., 5th ser., vol. 1,

p. 435, pi. 20, figs. 6a, b, 1878. 

Sidebottom, Manchester Lit. Philos. Soc. Mem. and Proc.,

vol. 151, no. 9, p. 11, pi. 2, fig. 18, 1907. 

Guttulma fracta Bornemann, Deutsch. geol. Gesell. Zeitschr.,

vol. 7, p. 344, pi. 17, fig. 4, 1855. 

Guttulma incurva Bornemann, Deutsch. geol. Gesell. Zeitschr.,

vol. 7, p. 345, pi. 17, fig. 6, 1855. 

Guttulina globosa Bornemann, Deutsch. geol. Gesell. Zeitschr.,

vol. 7, p. 346, pi. 18, fig. 1, 1855. 

Guttulina obtusa Bornemann, Deutsch. geol. Gesell. Zeitschr.,

vol. 7, p. 346, pi. 18, fig. 2, 1855. 

Guttulina turgida Reuss, Akad. Wiss. Wien Sitzungsber., vol.

18, p. 246, pi. 6, fig. 66, 1855 (1856). 

Polymorphina turgida Reuss, Akad. Wiss. Wien Sitzungsber.,

vol. 62, pt. 1, p. 487, 1870. 

Von Schlicht, Die Foraminiferen des Septarienthones von

Pietzpuhl, pi. 28, figs. 6-10; pi. 29, figs. 1-5,1870. 

Globulina subalpina Gttmbel, K. bayer. Akad. Wiss. Munchen,

Cl. 2, Abh., vol. 10, p. 646, pi. 2, figs. 80a, b, 1870. 

Polymorphina subcruoiata Terquem, Soc. g6ol. France Me"m.,

ser. 3, vol. 2, p. 140, pi. 14 (22), figs. 26, 27, 1882.

Test ellipsoidal or ovoid to cylindrical, rounded at 

the base, slightly produced at the apertural end; 

chambers rounded, almost as long as broad, arranged
28 UPPER EOCENE FOEAMINIFEEA OP THE SOUTHEASTERN UNITED STATES

in a nearly triserial series, each succeeding chamber 

removed much farther from the base, rarely becoming 

almost uniserial in the last chamber; sutures but little 

depressed, generally distinct; wall smooth, thick, often 

with fistulose tubes; aperture radiate. Length, 0.45- 

0.90 mm; breadth, 0.30-0.60 mm; thickness, 0.25-0.55 

mm.

Cooper marl: 11862, 1 mile west of Givhans, Dorchester 

County, S.C.; 11857, 1 mile south of Moncks Corner, Berkeley 

County, S.C.

Genus SIGMOMORPHINA Cushman and Ozawa, 1928 

Sigmomorphina jacksonensis (Cushman)

Plate 10, figures \-A

Polymorphina jacksonensis Cushman, Contr. Cushman Lab.

Foram. Research, vol. 2, p. 36, pi. 5, figs. 5a, b, 1926. 

Sigmomorphina jacksonensis Cushman and Ozawa, U.S. Nat.

Mus. Proc., vol. 77, art. 6, p. 123, pi. 32, figs. 2a, b, 1930. 

Howe and Wallace, Louisiana Dept. Conservation Geol.

Bull. 2, p. 49, pi. 8, fig. 2, 1932. 

Polymorphina compressa Nuttall (not D'Orbigny), Geol. Soc.

London Quart. Jour., vol. 84, p. 93, pi. 6, figs. 18, 19,

1928.

Test broad and compressed, periphery broadly 

rounded, apertural and narrowed to a slightly produced aperture; chambers elongate, slightly inflated, 

embracing, arranged in a contraclockwise sigmoid 

series, each succeeding chamber removed farther from 

the base; sutures very slightly depressed, distinct, almost straight; wall thick, smooth; aperture radiate. 

Maximum length 1.10 mm; breadth 0.65 mm; thickness 0.25 mm.

While this species and the following variety are 

characteristic of the Jackson Eocene of the Gulf 

Coastal Plain of the United States, it also occurs in 

the Claiborne group and in the Eocene of Trinidad.

Cooper marl: 6321, Baldock, S.C.

Ocala limestone: 1/19, near Blue Springs, Jackson County, 

Fla., the type locality; 1/7, 1 mile east of Beck, Ala.; 1/43, 

l1/^ miles southwest of Perdue Hill, Ala.

Jackson formation: 2-637, Cocoa, Ala.; 7203, 3% miles southeast of Cullomburg, Ala.; 6457, 1% miles south of Shubuta, 

Miss.; 6463, Jackson, Miss.

Sigmomorphina jacksonensis (Cushman) var. costifera 

Cushman and Ozawa

Plate 10, figures 5, 6

Polymorphina jacksonensis Cushman var. costifera Cushman, 

Contr. Cushman Lab. Foram. Research, vol. 2, p. 35, 

1926.

Sigmomorphina jacksonensis (Cushman) var. costifera Cushman and Ozawa, U.S. Nat. Mus. Proc., vol. 77, art. 6, 

p. 123, pi. 32, figs. 3 a, b, 1930.

Ellisor, Am. Assoc. Petroleum Geologists Bull., vol. 17, 

no. 11, pi. 7, fig. 1, 1933.

Variety differing1 from the typical form in the ornamentation of the test, which has several rounded longitudinal costae on the basal portion, sometimes covering 

a large part of the surface. As a rule the variety and

the typical form do not occur together, although both 

occur at Jackson, Miss., and at a station 1 mile east of 

Beck, Covington County, Ala. Maximum length 1.20 

mm; breadth 0.50 mm; thickness 0.13 mm.

Cooper marl: 5531, Barnwell County, S.C.

Ocala limestone: 7189, Rich Hill, Ga.; 6730, Double Branches 

Greek, Ala.; 6740, mouth of Robinsons Mill Creek, Ala.; 1/7, 

1 mile east of Beck, Ala., type locality.

Jackson formation: 6463, Jackson, Miss.; 6471, Garlands 

Creek, Miss.; 6472, Yazoo City, Miss.

Sigmomorphina regularis (Yon Minister)

The only American records for this species seem to 

be from the Cooper marl of South Carolina. The 

other records are mostly from the German Oligocene. 

Figures and synonymy of this species are given by 

Cushman and Ozawa.11

Cooper marl: 5531, Charleston, S.C.; 11857, 1 mile south of 

Moncks Corner, S.C.

Sigmomorphina vaughani Cushman and Ozawa

Plate 10, figures 7a-c

Sigmomorphina vaughani Cushman and Ozawa, U.S. Nat. Mus. 

Proc., vol. 77, art. 6, p. 137, pi. 38, figs. 2a-c, 1930.

Test almost rhombic with three rounded angles and 

acute apertural angle, compressed, and more or less 

twisted, margin of the lower half wavy; chamber compressed, elongated, arranged in a clockwise sigmoid 

series, each succeeding chamber farther removed from 

the base; sutures slightly depressed, distinct; wall 

smooth; aperture radiate. Length of holotype 0.80 

mm; breadth 0.62 mm; thickness 0.30 mm.

Cooper marl: 6319, Ingleside, S.C., type locality; 11857, 1 

mile south of Moncks Corner, Berkeley County, S.C.

Sigmomorphina semitecta (Reuss) var. terquemiana 

(Fornasini)

Polymorphina amygdaloides Reuss var. terquemiana Fornasini, Accad. sci. 1st. Bologna Mem., ser. 5, vol. 9, p. 72, 

fig. 25 (in text), 1902.

Sigmomorphina semitecta (Reuss) var. terquemiana Cushman and Ozawa, U.S. Nat. Mus. Proc., vol. 77, art. 6, p. 

129, pi. 33, figs. 4, 5; pi. 34, figs. 2, 3; pi. 35, fig. 1, 

1930.

Polymorphina, amygdaloides Terquem (not Reuss, 1851), Soc. 

gjeol. France Mem., ser. 3, vol. 2, p. 141, pi. 14 (22), 

figs. 30, 31, 1882.

Polymorphina pauperata Terquem (part), Soc. g6ol. France 

Mem., ser. 3, vol. 1, p. 38, pi. 3 (8), figs, lla, b (not 

figs. 12-19), 1878.

Olo'bulina varians Terquem, Soc. g6ol. France Mem., s6r. 3, 

vol. 2, p. 128, pi. 13 (21), figs. 9-16, 1882.

This variety occurs in the Jackson Eocene of the 

Gulf Coastal Plain but only rarely. Other records are 

mostly from the European Tertiary.

Ocala limestone: 7189, Rich Hill, Ga.; 6741, 3^ miles northeast of Brooklyn, Ala.

11 U.S. Nat. Mus. Proc., vol. 77, art. 6, p. 126, pi. 33, flgs. la, b, 

1930.
POLYMORPHINIDAE 29

Genus PSETJDOPOLYMOKPHINA Cushman and Ozawa, 1928 

Pseudopolymorphina dumblei (Cushman and Applin)

Plate 10, figures 14,15

Polymorphina compressa D'Orbigny var. dumblei Cushman and

Applin, Am. Assoc. Petroleum Geologists Bull., vol. iO,

p. 173, pi. 9, figs. 4, 5, 1926. 

Pseudopolymorphina diirnblei Cushman and Ozawa, U.S. Nat.

Mus. Proc., vol. 77, art. 6, p. 97, pi. 25, figs, la, b, 1930. 

Blllsor, Am. Assoc. Petroleum Geologists Bull., vol. 17,

no. 11, pi. 7, fig. 2, 1933.

Test compressed, elongated, obtuse at the base, more 

or less acute at the apertural end; chambers slightly 

longer than wide, more or less compassed, much embracing, arranged at first in a nearly triserial series, 

becoming biserial later; sutures but little depressed, 

distinct; wall smooth; aperture radiate. Length 

0.70-1.10 mm.; breadth 0.35-0.40 mm; thickness 0.16- 

0.20 mm.

This seems to be a definite species originally described from the Jackson of Bridge Creek, 1% miles 

above the Angelina River, San Augustine County, 

Tex., but since found rather widely distributed in the 

upper Eocene of the Gulf Coastal Plain and in 

Mexico.

Ocala limestone: 7189, Rich Hill, Ga.; 7719, Spring Mill 

Branch, Jenkins County, Ga.; 6742, Powells Landing, Sepulga 

River, Ala.; 1/8, Beck, Ala.

Pseudopolymorphina decora (Reuss)

Plate 10, figures 11, 12

Polymorphina decora Reuss, Acad. roy. sci. Belgique Bull., 

s6r. 2, vol. 15, p. 152, pi. 3, fig. 41, 1863.

Ps&udopolymorphina decora Cushman and Ozawa, U.S. Nat. 

Mus. Proc., vol. 77, art. 6, p. 96, pi. 24, figs. 6-8, 1930.

Polymorphina texana Cushman and Applin, Am. Assoc. Petroleum Geologists Bull., vol. 10, p. 173, pi. 9, figs. 1, 2, 

1926.

Test elongated, more or less compressed, periphery 

rounded, sides nearly parallel; chambers somewhat 

longer than wide, slightly embracing, alternating, arranged in a nearly biserial series from the start; 

sutures not depressed, distinct; wall smooth; aperture 

radiate. Length 0.75-1.10 mm; breadth 0.32-0.36 

mm; thickness 0.15-0.18 mm.

This species occurs in the upper Eocene of Texas, 

and the specimens figured came from the Jackson 

group on Tarkiln Creek, half a mile above the Neches 

River, Trinity County, Tex.

Genus POLYMORPHINA D'Orbigny, 1826 

Polymorphina advena Cushman

Plate 10, figure 8

Polymorphina advena Cushman, U.S. Geol. Survey Prof. Paper 

129, p. 132, pi. 31, fig. 4, 1922; Contr. Cushman Lab. 

Forarn. Research, vol. 5, p. 41, pi. 7, fig. 5, 1929.

Cushman and Ozawa, U.S. Nat. Mus. Proc., vol. 77, art. 6, 

p. 118, pi. 30, figs. lOa, b, 1930.

Test much compressed, broadly ovate; chambers 

numerous, elongate, alternating, much the broadest 

near the apertural end; sutures slightly depressed, 

very oblique; surface ornamented with numerous fine 

longitudinal costae, often obscure, except the lastformed one or two chambers, which are smooth, at 

least at the apertural end; aperture radiate. Length 

0.40-0.65 mm; breadth 0.22-0.30 mm; thickness 0.05- 

0.06 mm.

This is a characteristic species of the lower Oligocene of the Gulf Coastal Plain of the United States 

but occurs also in the upper Eocene of the same region.

Cooper marl: 11S63, three-fourths mile west of Old Biggin 

Church, Berkeley County, S.C.; 11S57, 1 mile south of Moncks 

Corner, Berkeley County, S.C.

Ocala limestone: 7189, Rich Hill, Ga.; 7237, half a mile east 

of Huguenen Ferry, Crisp County, Ga.; 6720, three-fourths mile 

north of Drewry, Ala.; 6725, Claiborne, Ala.; 6730, Double 

Branches Creek, .Ala.

Jackson formation: 6463. Jackson, Miss.

Genus SIGMOIDELLA Cushman and Ozawa, 1928 

Sigmoidella plummerae Cushman and Ozawa

Plate 10, figures 13a, b

Sigmoidella plummerae Cushman and Ozawa, U.S. Nat. Mus. 

Proc., vol. 77, art. 6, p. 142, pi. 39, figs. 3a, b, 1930.

Test compressed, ovate, broadly rounded at the base, 

acute toward the aperture, periphery angular; chambers elongated, more or less compressed, arranged in a 

contraclockwise sigmoid series, each succeeding chamber embracing the earlier one, but often the last chamber not extending down to the initial end; sutures not 

depressed, usually distinct; wall rather thin, smooth; 

aperture radiate. Length of the holotype 0.70 mm; 

breadth 0.35 mm; thickness 0.22 mm.

The types are from the Eocene Cook Mountain formation of Texas, but there are also records from the 

Cooper marl of South Carolina.

Cooper marl: 5531, Cooper River, S.C.; 6319, Ingleside, S.C.; 

11861, steep bluff on west side of Biggin Creek, Berkeley 

County, S.C.; 11863, three-fourths mile west of Old Biggin 

Church, Berkeley County, S.C.; 11862, 1 mile west of Givhans, 

Dorchester County, S.C.; 11857, 1 mile south of Moncks Corner, Berkeley County, S.C.

Genus GLANDULINA D'Orbigny, 1828

Glandulina laevigata D'Orbigny var. ovata Cushman and

Applin

Plate 10, figures 16, 17

Nodosaria (Glandulina) laevigata D'Orbigny var. ovata Cushman and Applin, Am. Assoc. Petroleum Geologists Bull., 

vol. 10, p. 169, pi. 7, figs. 12, 13, 1926.

Glandulina laevigata D'Orbigny var. ovata Ellisor, Am. Assoc. 

Petroleum Geologists Bull., vol. 17, no. 11, pi. 2, fig. 6, 

1933.
30 UPPER EOCENE FOKAMINIFERA OF THE SOUTHEASTERN UNITED STATES -

Test ovate, longer than broad, circular in transverse 

section, widest toward the apertural end; chambers 

overlapping, few, indistinct, earlier ones irregularly 

biserial, especially in the microspberic form, later ones 

uniserial; sutures indistinct; wall smooth, matte; aperture radiate, slightly projecting. Average length 

0.60 mm; breadth 0.40 mm.

This somewhat shorter and broader variety of the 

typical form occurs in the Jackson of Texas, and the 

types were collected 4 miles east of Diboll, Angelina 

County.

Family NONIONIDAE 

Genus NONION Montfort, 1808 

Nonion advenum (Cushman)

Plate 11, figures 1- 4

Nonionina advena Cushman, U.S. Geol. Survey Prof. Paper 129, 

p. 139, pi. 32, fig. 8, 1922; Prof. Paper 133, p. 50, 1923. 

Cushman and Applin, Am. Assoc. Petroleum Geologists 

Bull., vol. 10, p. 181, pi. 10, figs. 16, 17, 1926.

Test small, circular, in side view biconvex; periphery 

rounded, 9 to 11 chambers in the last-formed coil, inflated ; sutures curved, slightly sigmoid, the inner portion excavated and broadened; umbilical region at 

each side of the test occupied by a large projecting 

knob of clear shell material; aperture at the base of 

the last-formed chamber. Maximum length 0.75 mm.

This species was originally described from the Mint 

Spring marl member of the Marianna limestone 

(lower Oligocene), from Vicksburg and Boice, Miss. 

It occurs in the upper Eocene of Texas near Cheapside, Gonzales County. The species also occurs in the 

lower Oligocene and upper Eocene of the Atlantic 

Coastal Plain of Mexico. It is allied to some of the 

species described by Terquem from the Eocene of the 

Paris Basin.

Cooper marl: 6321, Baldock, S.C.

Ocala limestone: 7189, Rich Hill, Ga.; 7704, 3 miles west of 

Taryersville, Ga.; 6804, Ocala, Fla.; 6805, Oakhurst quarry, 

Ocala, Fla.; 6807, Zuber, Fla.; 6808, Martin, Fla.; 7366, Anthony, Fla.; 6719, three-fourths mile north of Drewry, Ala.: 

6725, 6727, Claiborne, Ala.; 6730, Double Branches Creek, Ala.; 

6741, 3% miles northeast of Brooklyn, Ala.; 6742, Powells 

Landing, Sepulga River, Ala.; 7159, 4 miles southeast of 

Whatley, Ala.; 7161, 5 miles southeast of Whatley, Ala.

Jackson formation: 7227, Water Valley, Ala.; 6463, Jackson, Miss.; 6471, Garlands Creek, Miss.

Barnwell formation: 7755, Stevens pottery, Ga.

Nonion inexcavatum (Cushman and Applin) 

Plate 11, figures 5-8

No-nionina advena Cushman var. ineticavata Cushman and

Applin, Am. Assoc. Petroleum Geologists Bull., vol. 10,

p. 182, pi. 10, figs. 18, 19, 1926. 

Nonion inexcavatum Bllisor, Am. Assoc. Petroleum Geologists

Bull., vol. 17, no. 11, pi. 2', fig. 7, 1933. 

Nonion inexcavatum (Cushman and Applin) var. Ellisor, op.

cit, pi. 7, figs. lOa, b.

Test of medium size, circular in side view, biconvex, 

periphery subacute, 12 to 15 chambers in the lastformed coil in adults, distinct, slightly inflated; sutures very slightly curved, very slightly if at all 

depressed; umbilical region at each side of the test 

occupied by a slightly projecting knob of clear shell 

material, with the area toward the aperture from this 

knob covered with a pustulose ornamentation, especially marked in adults; aperture at the base of the 

last-formed chamber. Maximum breadth 0.85 mm.

This was originally described as a variety of Nonion 

advenum (Cushman), but a study of the more numerous specimens of .the present series seems to show that 

it is specifically different. In adults there are traces 

in the last two or three chambers of retral processes, 

making the relation to Elphidium very marked.

The type specimens are from the upper Eocene of 

Texas, 4 miles east of Diboll, Angelina County.

Cooper marl: 6321, Baldock, S.C. 

Ocala limestone: 1/8, Beck, Ala.

Jackson formation: 6463, Jackson, Miss.; 7227, Water Valley, 

Ala.

Nonion chapapotense Cole

Plate 11, figures 9-13

Nonion chapapotensis Cole, Bull. Am. Paleontology, vol. 14, 

no. 53, p. 210 (10), pi. 1, figs. 18, 19, 1928.

Test close-coiled planispirally, the umbilical area on 

each side very slightly open, but filled with a central 

.boss, about which is a narrow, depressed, grooved area, 

the outer edge of which is slightly thickened, periphery rounded; chambers distinct, numerous, usually 

about 10 in the adult coil, of uniform shape, gradually 

increasing in size as added; sutures distinct, somewhat 

limbate, slightly curved, fusing at the inner end into 

the thickened ridge at the outside edge of the grooved 

area about the umbilical boss; wall of the chamber 

smooth, conspicuously perforate; aperture along the 

base of the apertural face in the median line. Diameter 0.40 mm; thickness 0.20 mm.

This species was described by Cole from the Eocene 

Chapapote formation of Mexico. In the Eocene material of the present paper it shows considerable variation, with many intermediate stages between the 

forms with an open umbilicus and those with a large 

boss.

Ocala limestone: 3966, 2 miles south of Perry, Ga.; 6726, 

Claiborne, Ala.; 6738, 6% miles north of Brooklyn, Ala.; 7161, 

5 miles southeast of Whatley, Ala.; 1/8, Beck, Ala.

Jackson formation: 6472, 6473, Yazoo City, Miss.

Nonion micrum Cole

Plate 11, figures 14, 15

Nonion micrus Cole, Bull. Am. Paleontology, vol. 14, no. 51, 

p. 22, pi. 5, fig. 12, 1927; vol 14, no. 53, p. 211 (11), 

1928.

Weinzierl and Applin, Jour. Paleontology, vol. 3, p. 400, 

pi. 43, figs. 6a-c, 1929.
NONTONIDAE 31

Test small, compressed, bilaterally symmetrical, 

periphery subacute; chambers distinct, somewhat involute, rapidly increasing in size as added, about 6 

making up the adult coil, sutures distinct, depressed, 

slightly curved; wall smooth, finely perforate; aperture median, at the base of the apertural face of the 

chamber, with a slight lip. Diameter 0.25-0.32 mm; 

thickness 0.10-0.13 mm.

This species has been recorded from the Claiborne 

Eocene of Mexico and Texas. It seems to occur in 

the Jackson but is evidently very rare.

Jackson formation: 6646, Hays Chapel, Wayne County, Miss.

Nonion whitsettense (Cushman and Applin)

Plate 11, figures 16-18

Nonionina whitsettensis Cushman and Applin, Am. Assoc. Petroleum Geologists Bull., vol. 10, p. 183, pi. 10, figs. 

4-6, 1926.

Nondon wliitsettensis Ellisor, Am. Assoc. Petroleum Geologists 

Bull., vol. 17, no. 11, pi. 2, fig. 14, 1933.

Test much compressed, sides nearly parallel, the 

periphery broadly rounded; chambers long, narrow, 

distinct, usually 9 in the adult coil; sutures very 

strongly curved, especially in the later portion, slightly 

depressed; wall smooth; aperture a very narrow, 

elongate slit at the base of the apertural face. Maximum diameter 0.50 mm; thickness 0.10-0.12 mm.

The type specimens of this species came from the 

Whitsett beds (Frio clay) of the Jackson of Texas, 

from Ohio and Ked River well 2, Tyler County, at a 

depth of 330 feet, where it is very common and holds 

its characters very closely. There seem to be occasionally traces of what may be retral processes, but 

they are not definite. The species has not been found 

in the collections from the more easterly portion of the 

Gulf Coastal Plain.

Genus NONIONELLA Cushman, 1926 

Nonionella hantkeni (Cushman and Applin)

Plate 12, figures 1, 2

Nonionina Iwmtkeni Cushman and Applin, Am. Assoc. Petroleum Geologists Bull., vol. 10, p. 182, pi. 10, figs. 10, 11, 

1926.

Nonionella hantkeni Ellisor, Am. Assoc. Petroleum Geologists 

Bull., vol. 17, no. 11, pi. 2, fig. 9, 1933.

Pullenda elongata Hantken, Magy. kir. fb'ldt. int. Evkonyve, 

p. 50, pi. 10, fig. 10, 1875 (1876) ; K. ungar. geol. Anstalt 

Mitt. Jahrb., vol. 4, p. 59, pi. 10, fig. 10, 1875 (1881).

Test somewhat longer than broad, consisting of 

numerous chambers, as many as 12 in the last-formed 

coil, periphery broadly rounded, the umbilical region 

slightly exposed, showing the inner ends of the earlier 

chambers; sutures distinct, very slightly and evenly 

curved; wall smooth; aperture arched in the base of 

the apertural face of the last-formed chamber. Maximum length, 0.60 mm.

62330 36 3

Type specimen from Ohio and Red River well 2, 

Tyler County, Tex., at a depth of 2,330 feet.

In the regions east of Texas this species is apparently replaced by the following variety.

Nonionella hantkeni (Cushman and Applin) var. spissa 

Cushman

Plate 12, figures 6a-c

Nonionella hantkeni (Cushman and Applin) var. spissa Cushman, Contr. Cushman Lab. Foram. Research, vol. 7, p. 58, 

pi. 7, figs. 13a-c, 1931.

Ellisor, Am. Assoc. Petroleum Geologists Bull., vol. 17, no. 

11, pi. 2, figs. 10, 12, 1933.

Variety differing from the typical form in the much 

thicker test and the usually smaller number of chambers. Some of the broader specimens are close to the 

typical in outline, but there is a very considerable 

amount of variation in any large series of specimens. 

The typical form occurs in the Eocene of Texas, but 

the material from regions to the east seems all to 

belong to the variety.

Holotype (no. 371677, U.S.N.M.) from Cooper marl, 

station 11863, Berkeley County, S.C.

Cooper marl: 11861, steep bluff on west side of Biggin 

Creek, Berkeley County, S.C.; 11863, three-fourths mile west 

of Old Biggin Church, Berkeley County, S.C.; 11862, 1 mile 

west of Givhans, Dorchester County, S.C.; 11857, 1 mile south 

of Moncks Corner, Berkeley County, S.C.

Ocala limestone: 6754, Choctawhatchee River at State line, 

Ala.-Fla.; 6755, Choctawhatchee River above State line, Ala.; 

6720, three-fourths mile north of Drewry, Ala.; 6724, 6725, 6726, 

6727, Claiborne, Ala.; 6730, Double Branches Creek, Ala.; 6741, 

3% miles northeast of Brooklyn, Ala.; 6742, Powells Landing, 

Sepulga v River, Ala.; 1/7, 1 mile east of Beck, Ala.; 1/8, 

Beck, Ala.

Jackson formation: 6463, Jackson, Miss.; 6471, Garlands 

Creek, Miss.

Nonionella jacksonensis Cushman

Plate 12, figures 3, 4

Nonionella jacJcsonensis Cushman, Contr. Cushman Lab. Foram. 

Research, vol. 9, p. 10, pi. i, figs. 23a-c, 1933.

Test longer than broad, periphery rounded, ventral 

side involute and the chambers extending over the umbilical region, dorsal side with the chambers ending at 

the umbilical region; chambers distinct, about 8 in the 

final whorl, becoming increasingly elongate in the 

adult, the inner end of the final chamber extending 

across the umbilical area nearly to the periphery on 

the ventral side, inflated; sutures distinct, slightly if 

at all depressed; wall smooth, finely perforate; aperture peripheral at the base of the apertural face, low. 

Length 0.35 mm; breadth 0.20 mm; thickness 0.10 mm.

Holotype (no. 371678, U.S.N.M.) from Ocala lime 

stone, station 6726, Claiborne, Ala.

This species resembles some of the forms often assigned to Nonionella turgida (Williamson), but the
32 UPPER EOCENE FOEAMINIPEEA OP THE SOUTHEASTERN UNITED STATES

general shape is different, and the chambers of our 

species are broader and usually fewer.

Cooper marl: 5531, Charleston, S.C.

Ocala limestone: 5567, 6726, 6727, Claiborne, Ala.; 6738, 6% 

miles north of Brooklyn, Ala.; 6741, 3% miles northeast of 

Brooklyn, Ala.

Genus ELPHIDIUM Montfort, 1808 

Elphidium texanum (Cushman and Applin)

Plate 12, figures 5a-c

Polystomella texana Cushman and Applin, Am. Assoc. Petroleum Geologists Bull., vol. 10, p. 184, pi. 10, figs. 7-9, 

1926.

Test nearly circular in outline, the periphery 

broadly rounded, slightly umbonate; chambers distinct, usually 11 in the adult coil, of nearly uniform 

size; sutures distinct, slightly depressed, strongly 

curved, retral processes only slightly developed; wall, 

smooth; aperture a low arched slit at the base of the 

ventral face of the last-formed chamber. Diameter 

0.45 mm; thickness 0.20 mm.

The type specimens of this species came from the 

Jackson Eocene from Ohio and Red River well 2, 

Tyler County, Tex., at a depth of 537 feet.

The retral processes are not highly developed, and 

young specimens are difficult to distinguish from 

Nonion. It has not been recorded from the area east 

of Texas. Specimens were collected by Alexander 

Deussen in outcrop material of the Jackson group in 

a bluff on Atascosa Creek near the old Whitsett ranch 

house in Atascosa County, Tex., U.S.G.S. station 6148.

Family CAMERINIDAE

Genus ASSILINA D'Orbigny, 1826

Assilina floridensis (Heilprin)

Nummulites floridensis Heilprin, Acad. Nat. Sci. Philadelphia 

Proc. for 1884, pp. 321, 322, text fig., 1885.

Operculina floridensis Cushman, U.S. Geol. Survey Prof. Paper 

128, p. 130, pi. 20, fig. 12, 1921.

Test planispiral, much compressed, composed of 3 

or 4 coils, gradually increasing in breadth; chambers 

very numerous, 30 or more in the last-formed coil of 

the adult, the length about five times the height, somewhat irregular in size, each chamber very gradually 

increasing in size toward the greatest height, which is 

near the periphery; sutures slightly curved at first but 

gradually increasing in convexity in the adult; the 

height of the coil not increasing, and in the specimens 

there is actually a decrease, indicating senescence 

toward the end of the development. Maximum diameter 8.00 mm.

This species probably belongs in the genus AssiMna 

of D'Orbigny. The outer whorls are not involute; 

otherwise it is close to Camerina.

Ocala limestone: 3636, Spring Hill schoolhouse, 6 miles 

north of Alachua, Fla.; 3684, Pemberton Ferry (now Groom), 

Hernando County, Fla., the type locality; 3686(7), Martin, 

Fla.

Genus OPERCULINA D'Orbigny, 1826 

Operculina cookei Cushman

Operculina, cookei Cushinaii, U.S. Geol. Survey Prof. Paper 

128, p. 127, pi. 18, flgs. 1, 2, 1921.

Test planispiral, much compressed, composed of 2% 

to 3 coils rapidly increasing in breadth; .chambers very 

numerous, 28 to 32 in the last-formed coil, fairly constant in size and shape, in the adult the length of the 

chamber 10 to 12 times the height, each chamber gradually widening toward the center or slightly beyond 

the center; sutures starting in a slightly tangential 

curve that rapidly increases in convexity toward the 

periphery, near which it bends back at a sharp angle, 

joining the peripheral margin of the test; surface of 

the test in the umbonal region with numerous rounded 

bosses irregularly placed, the sutures in the lastformed coil ornamented by raised ridges, in the earlier 

part broken into separate knobs and bosses, toward the 

later part fused into continuous, raised ridges; surface of the chambers between the sutures somewhat 

concave, smooth, translucent. Length of adult specimens as much as 10.00 mm.

Ocala limestone: 7116, Flint River near Oakfield, Ga., the 

type locality; 3626(7), 4y2 miles southwest of Philema, Ga. 

Jackson formation: 6458, Moodys Branch, Jackson, Mis-s.

Operculina ocalana Cushman

Operculina ocalana Cushman, U.S. Geol. Survey Prof. Paper 

128, p. 129, pi. 19, figs. 4, 5, 1921.

Test planispiral, much compressed, composed of 

2 to 3 coils, the last with 16 to 18 chambers; sutures 

raised, confluent in the center, somewhat rounded, 

the area between concave and smooth; chambers 3 to 

4 times as long as wide; central area of the test umbonate; periphery somewhat raised by a thickening 

in which the raised sutures terminate. Maximum 

length 6.00 mm.

Ocala limestone: 3626, 4^ miles southwest of Philema, Ga.; 

3760, half a mile south of Tivola, Ga.; 3387, Red Bluff, Flint 

River, Ga.; 3390, 7097, Bainbridge, Ga.; 7123, Flint River, 4% 

miles below Blue Spring, Ga.; 7126, Flint River, Sy2 to 

9 miles below Newton, Ga.; 7127, a quarter of a mile below 

Normans Ferry, Flint River, Ga.; 4965, half a mile southwest 

of Fort White, Fla.; 6785, three-fourths of a mile south of 

Herlongs, Fla.; 7337, Suwannee River above Florida Railway, 

Fla.; 7348, Suwannee River 1% miles above Troy Springs, 

Fla.; 7349, Suwannee River half a mile below Fort McComb, 

Fla.; 6747, Steamboat Point, Sepulga River, Ala., the type 

locality.

Operculina vaughani Cushman

Operculina vaughani Cushman, U.S. Geol. Survey Prof. Paper 

128, p. 128, pi. 19, figs. 6, 7, 1921.

Test planispiral, much compressed, composed of 

about 3 coils rapidly increasing in breadth; chambers 

numerous, 20 to 22 in the last-formed coil, fairly constant in size and shape, in the adult the length of the 

chamber 5 to 6 times the height, each chamber grad-
HETEBOHELICIDAE 33

ually increasing in height from the proximal end to 

the maximum height near the distal end; sutures usually starting from the proximal end in a nearly 

straight radial line and keeping so for about one-third 

the length, then gently rounded backward toward the 

periphery, near which the rate of curvature is abruptly 

increased. Length of adult specimens 7.00-8.00 mm. 

Ocala limestone: 3617, Muckafoonee Creek, Albany, Ga.; 

7115, Flint River near Warwick, Ga.; 7117, Flint River, Lee 

County, Ga.; 7119, Flint River 16 miles above Albany, Ga.; 

7120, Flint River at Albany, Ga.; 7237, half a mile east of 

Huguenen Ferry, Crisp County, Ga.

Operculina mariannensis Vaughan

Operoulina mariannensis Vaughan, Florida Geol. Survey 19tli 

Ann. Rept., p. 158, pi. 1, figs. 1-4, 1928.

Vaughan gives the following description:

Test small, compressed, thin. Diameter from outer edge of 

aperture through the center 2.00 mm; diameter at right angle 

to preceding 1.70 mm; thickness through the center about 0.30 

mm, but only about half as thick outside the central area.

The number of chambers in the last coil is 11 or 12. The 

chamber walls are marked externally by costae and rather 

coarse granules, as shown in the figures. At the center there 

is a knob which is larger than the costal granulations, its 

diameter ranging between 0.10 and 0.20 mm.

There are almost 4 coils in a specimen 2.00 mm in diameter. 

The chamber walls curve rather uniformly and bend backward 

near the periphery.

Operculwa mariannensis is closely related to O. ocalana 

Cushman, with which it is associated and from which it is 

distinguished by its more delicate test and its smaller number 

of chambers. The number of chambers in even small specimens 

of 0. ocalana is between 16 and 18, whereas in 0. mariannensis 

the number is between 11 and 13. The costae corresponding to 

the chamber walls in O. ocalana are usually sinuous and not 

regularly curved as in 0. mariannensis, and their sides slope. 

The intercostal areas in 0: mariannensis are flatter and more 

sharply bounded. The center of the test of O. ocalana is more 

conical than in 0. mariannensis.

Vaughan described this species from " Chipola 

River below wagon bridge east of Mananna, Fla., 

lowest bed of Ocala limestone exposed " at that locality; collected by C. Wythe Cooke and D. S. Mossom 

(U.S.G.S. station 6768).

Genus OPERCULINELLA Yabe, 1918 

Operculinella willcoxi (Heilprin)

Nummulites willcoxi Heilprin, Acad. Nat. Sci. Philadelphia 

Proc. for 1882, p. 191, text figs. 1, 2, 1S82; idem for 

1884, pp. 321, 322, text figs. 1, 2, 1885.

Operoulina willcoxi Cushman, U.S. Geol. Survey Prof. Paper 

128, p. 129, pi. 20, figs. 9-11, 1921.

Test planispiral, nearly involute, especially in the 

young, compressed, about 5 coils in the adult, 35 to 

45 chambers in the last-formed coil; sutures raised, 

irregularly curved, somewhat wavy near the middle, 

central part of the test umbonate. Diameter about 

8.00 mm.

Ocala limestone: 322, Newmansville, Fla.; 329, Padlock, Fla.; 

4965, half a mile southwest of Fort White, Fla.; 6790, Alachua,

Fla.; 6804, Ocala, Fla.; 6805, Oakhurst quarry, Ocala, Fla.; 

6S08, Martin, Fla.; 6817, Fort White, Fla.; 7337, Suwannee 

River above Florida Railway, Fla.

Genus HETEROSTEGINA D'Orbigny, 1826 

Heterostegina ocalana Cushman

Heterostegina, ocalana Cushman, U.S. Geol. Survey Prof. Paper 

128, p. 130, pi. 21, figs. 15-18, 1921.

Test broadly complanate, central portion thickest, 

biconvex, rapidly thinning to the periphery, which is 

rounded; central portion with irregularly radial 

costae fusing near the base of the convex portion with 

the raised ribs of the sutures; in well-preserved specimens the lines between the chambers often marked by 

raised ribs, giving that portion of the test a reticulate 

appearance. Maximum length 8.00 mm.

Ocala limestone: 322, Newmansville, Fla.; 365, Johnsons 

Sink, Levy County, Fla.; 380, near Gainesville, Fla.; 3174, 

Edgar, Fla.; 3685, Martin, Fla.; 5030, Ocala, Fla.; 6785, threefourths of a mile south of Herlongs, Fla.; 6787, 2 miles north 

of High Springs, Fla.; 6789, Alachua, Fla.; 6804, Ocala, Fla.; 

6805, Oakhurst quarry, Ocala, Fla.; 6808, Martin, Fla.; 6810, 1 

mile northwest of Newberry, Fla.; 6812, 1% miles south of 

Newberry, Fla., the type locality; 6814, 1% miles northwest of 

Newberry, Fla.; 817, Fort White, Fla.; 7338, Bowling Springs, 

Fla.; 7341, Brauford, Fla.; 7365, Pineola, Fla.; 7367, Wiiiow 

Sink, Fla.; 7098, Red Bluff, Flint River, Ga.

Heterostegina ocalana Cushman var. glabra Cushman

Heterostegina ocalana Cushman var. glab)-a Cushman, U.S. 

Geol. Survey Prof. Paper'l28, p. 131, pi. 21, fig. 19, 1921.

Variety differing from the typical form mainly in 

the smoother surface of the test.

Ocala limestone: 7345, Suwannee River 2% miles above 

Branford, Fla.; 7348, Suwannee River 1% miles above Troy 

Springs, Fla., the type locality; 7349, Suwannee River half a 

mile below Fort McComb, Fla.

Family HETEROHELICIDAE

Genus SPIROPLECTOIDES Cushman, 1927

Spiroplectoides curta Cushman

Plate 12, figures 7a, b

Spiroplectoides curta Cushman, Contr. Cushman Lab. Foram. 

Research, vol. 9, p. 11, pi. 1, figs. 24a, b, 1933.

Test minute, about three times as long as broad, only 

slightly compressed in the later biserial portion, early 

planispiral portion much compressed, sides of the test 

nearly parallel; chambers distinct, slightly inflated, 

usually four pairs in the biserial portion, periphery in 

the later chambers broadly rounded; sutures distinct, 

very slightly if at all depressed, nearly at right angles 

to the periphery; wall distinctly perforate, otherwise 

smooth; aperture narrow on the terminal face, median. 

Length 0.25 mm; breadth 0.10 mm; thickness 0.06 mm.

Holotype (no. 371543, U.S.N.M.) from Cooper marl, 

station 11857, Berkeley County, S.C., where the species 

is common. It was not found in any of the other 

samples examined.
34 UPPER EOCENE FORAMINIFERA OF THE SOUTHEASTERN UNITED STATES

This is a very short, stout species of this genus and 

evidently has a restricted range. The planispiral early 

portion consists of about a single complete coil, and 

the chambers show best when the specimen is wet.

Cooper marl: 11857,1 mile south of Moncks Corner, Berkeley 

County, S.C.

Genus PLECTOFRONDICTILARIA Ljebus, 1903 

Plectofrondicularia-cf. P. vaughani Cushman

Plate 12, figures 8, 9

There are but two specimens of smooth, broad 

Plectofrondicularia in all the Eocene material examined. These are both from the Copper marl. One 

of these shows the early chambers fairly well when 

wet and seems to show that both probably belong to 

this species, described from the Alazan clays of 

Mexico.

Cooper marl: 5531, Charleston, S.C.; 11861, steep bluff on 

west side of Biggin Creek, Berkeley County, S.C.

Plectofrondicularia trinitatensis Cushman and Jarvis

Plate 12, figure 10

Plectofrondioularia trimtatensis Cushman and Jarvis, Contr. 

Cushman Lab. Foram. Research, vol. 5, p. 11, pi. 2, fig. 

16, 1929. i

Test elongate, sides nearly parallel except at the 

initial end, which is slightly tapering, initial end subacute; chambers in the earliest portion biserial, quickly 

becoming uniserial in all the later development, numerous, increasing in height as added, early ones with 

the sutures sloping to the sides and the chambers 

much broader than high, in the adult the sutures 

more nearly transverse and the chambers becoming 

higher than broad; sutures slightly depressed, distinct ; wall ornamented by several distinct, low, sharp 

costae, continuous from the initial to the apertural 

end, one at each periphery with 4 or 6 on each side 

of the test and occasionally another in the median 

line; aperture elliptical. Maximum length 1.00 mm 

or slightly more; breadth 0.18 mm; thickness 0.08 mm.

This species was originally described from the 

Eocene Mount Moriah beds of the Vistabella quarry, 

Trinidad.

The specimens from the Cooper marl have been 

compared with the holotype and seem to be identical.

Cooper marl: 11861, steep bluff on west side of Biggin Creek, 

Berkeley County, S.C.; 11857, 1 mile south of Moncks Corner, 

Berkeley County, S.C.

Plectofrondicularia cookei Cushman

Plate 12, figures 11, 12

Plectofrondicularia cookei Cushman, Contr. Cushman' Lab. 

' Foram. Research, vol. 9, p. 11, pi. 1, fig. 26, 1933.

Test elongate, tapering, much compressed, slightly 

keeled, early chambers biserial, later ones uniserial; 

chambers distinct, especially the later ones, earlier ones 

somewhat obscured by the ornamentation of the wall,

the uniserial ones extending back farther and farther 

at the sides as added; sutures of the later portion distinct and slightly depressed, somewhat limbate 

throughout; wall of the earlier one-third or one half 

ornamented by 'very distinct, sharp longitudinal 

costae, as many as 15 in some specimens, later portion 

of the test smooth; aperture elliptical, terminal. 

Maximum length 2.00 mm; diameter 0.15-0.25 mm.

Holotype (no. 371679, U.S.N.M.) from Cooper marl, 

Cooper River, S.C.

This species resembles some of those described by 

Eeuss and by Karrer from the Miocene and Oligocene 

of Europe and the Philippines, but it differs .from all 

of them. It is abundant at several stations in the 

Cooper marl but was not found in the other Jackson 

material examined. The biserial chambers can be 

observed only when the specimen is wet or mounted 

in balsam or other transparent media. Some specimens are much wider than those shown in the figures.

Cooper marl: 5531, Charleston, S.C.; 11861, steep bluff on 

west side of Biggin Creek, Berkeley County, S.C.; 11863, 

three-fourths of a mile west of Old Biggin Church, Berkeley 

County, S.C.; 11857, 1 mile south of Moncks Corner, Berkeley 

County, S.C.

Genus NODOGENEBJNA Cushman, 1927 

Nodogenerina cooperensis Cushman

Plate 12, figure 13

Nodogenerina cooperensis Cushman, Contr. Cushman Lab. 

Foram. Research, vol. 9, p. 11, pi. 1, fig. 27, 1933.

Test small, slender, tapering, final chamber usually 

the largest but occasionally reduced somewhat in size, 

uniserial throughout, or the initial end showing traces 

of a triserial arrangement; chambers distinct, subglobular or slightly pyriform, the greatest diameter 

slightly below the middle; sutures distinct, much depressed; wall very finely spinose; aperture terminal, 

circular, with a very short neck and distinct but narrow lip. Length 0.75 mm; diameter 0.13 mm.

Holotype (no. 371680, U.S.N.M.) from Cooper marl. 

1 mile south of Moncks Corner, Berkeley County, S.C., 

where it is abundant. This is a small but distinctive 

species with its finely spinose surface. The basal portion of the chamber is riot cut under as distinctly as in 

some of the other species of the genus, and the neck is 

very short.

Cooper marl: 6319, Ingleside, S.C.; 11857, 1 mile south of 

Moncks Corner, Berkeley County, S.C.

Family BULIMINIDAE

Genus BULIMINELIA Cushman, 1911

Buliminella alabamensis Cushman

Plate 13, figures 6a, b

Buliminella alabamensis Cushman, Contr. Cushman Lab. 

Foram. Research, vol. 2, p. 32, pi. 4, figs. 8a, b, 1926.

Test small, obovate, broadest in front view slightly 

above the middle, initial end pointed, apertural end
BULIMINIDAE 35

broadly rounded, somewhat obliquely truncated, whole 

test of iy2 to 2 coils; chambers fairly distinct, not inflated; sutures distinct but not depressed; apertural 

face with a very large open area somewhat broadening toward the base, which reaches to at least the 

middle of the test; wall very thin and transparent. 

Length 0.22 mm.

This species differs from others of the genus in the 

peculiar large open area on the apertural side of the 

test, broadening toward the base, which reaches to or 

below the middle line of the test. The apical end is 

acute. It is very distinct from the species recorded 

from the lower Oligocene.

Jackson formation: 7227, Water Valley, Choctaw County, 

Ala., type locality.

Genus BUIIMINA D'Orbigny, 1826

Bulimina jacksonensis Cushman

Plate 13, figures 7-9

Buliminch jacksonensis Cushman, Contr. Cushman Lab. Foram.

Research, vol. 1, pt. 1, p. 6, pi. 1, figs. 6, 7, 1925. 

Cushman and Applin, Am. Assoc. Petroleum Geologists

Bull., vol. 10, p. 168, pi. 7, figs. 8a, b, 1926. 

Howe and Wallace, Louisiana Dept. Conservation Geol.

Bull. 2, p. 59, pi. 11, fig. 5, 1932. 

Ellisor, Am. Assoc. Petroleum Geologists Bull., vol. 17, no.

11, pi. 7, fig. 5, 1933.

Test elongate, tapering, the initial end acute, 

broadly rounded at the apertural end and in adults 

somewhat contracted; chambers numerous, fairly distinct; sutures not depressed; surface ornamented by 

very prominent longitudinal costae, usually 6 to 8, 

platelike, much raised above the general surface, continuous from the apical end to the base of the lastformed chamber in adults; the outer margin in wellpreserved specimens serrate; aperture elongate, 

comma-shaped. Maximum length 1.40 mm.

This species is evidently the ancestor of the species 

of the Red Bluff clay, B. sculptiUs Cushman. The 

Jackson species is larger, the platelike costae higher, 

the sutures not depressed, and the costae in well-preserved specimens decidedly serrate as each new portion is added in growth. It occurs also in the Alazai 

clays of Mexico and the upper Eocene of Texas.

Both species belong to the group represented in the 

Eocene of Europe by B. tnmcana Giimbel, which has 

the costae more numerous and less prominent than 

in the American species. Related to these is B. 

buchiana D'Orbigny, known from the later Tertiary 

and in the present oceans. In this species the sculpture is still further reduced, but it is evidently a derivative from ancestral forms such as B. jacksonensis, 

B. sculptiUs, and B. truncana.

B. jacksonensis is found in,1 that part of the upper 

Eocene Jackson formation characterized by the 

" Zeuglodon bed " of station 2637, Cocoa, Ala., which

is representative of offshore conditions. Similar conditions are indicated in the Texan and Mexican 

material.

Cooper marl: Ashley marl works, Dorchester County, S.C.

Ocala limestone: 7378, 1% miles southwest of Inverness, 

Fla.; 7159, 4 miles southeast of, Whatley, Ala.; 1/38, 3% miles 

north of Grove Hill, Ala.; 1/43, V/2 miles southwest of Perdue 

Hill, Ala.

Jackson formation: 2637, Cocoa, Ala.; 7203, 3% miles southeast of Cullomburg, Ala.; 6457, 1% miles south of Shubuta, 

Miss.; 6646, Hays Chapel, Wayne County, Miss.

Bulimina jacksonensis Cushman var. cuneata Cushman

Plate 13, figures 10, 11

Bulimina jacksonensis Cushman var. cuneata Cushman, Contr. 

Cushman Lab. Foram. Research, vol. 2, p. 35, 1926.

Variety differing from the typical in the larger number of costae (10 to 12), the more tapering form, and 

especially the very serrate character of the edge of the 

costae. Length 1.00 mm.

This variety, which is distinct from the typical form, 

has been found at only two stations but occurs abundantly at the Alabama station.

Cooper marl: 11857,1 mile south of Moncks Corner, Berkeley 

County, S.C.

Jackson formation: 7220, half a mile southeast of Melvin, 

Ala., type locality.

Bulimina cooperensis Cushman

Plate 13, figures 12-14

Bulimina cooperensis Cushman, Contr. Cushman Lab. Foram. 

Research, vol. 9, p. 12, pi. 1, figs. 32a, b, 1933.

Test elongate, tapering, 2,y2 to 3 times as long as 

wide, greatest breadth toward the apertural end; 

chambers distinct, inflated, considerably overlapping; 

sutures deep, distinct; wall of the basal half of the 

chambers with platelike costae ending in sharp points, 

the initial end of the test often with a spine; aperture 

elongate with a slightly depressed border. Length 

0.40-0.50 mm; breadth 0.18-0.20 mm.

Holotype (no. 3T1684, U.S.N.M.) from Cooper marl, 

station 11857, Berkeley County, S.C.

This belongs in the general group of BuliTWna inflcKta but is quite different from the typical form of 

that species in the Pliocene of Italy. It was common 

at the type station but not found at any other in the 

material examined.

Cooper marl: 11857,1 mile south of Moncks Corner, Berkeley 

County, S.C.

Bulimina ovata D'Orbigny

Plate 13, figures 15, 16

Bulimina ovata D'Orbigny, ForaminifSres fossiles du bassin 

tertiaire de Vienne, p. 185, pi. 11, figs, 13, 14,1846.

Test narrowly ovate in front view, circular or nearly 

so in end view, the apex broadly rounded; chambers
36 TJPPEK EOCENE FORAMINIFERA OF THE SOUTHEASTERN UNITED STATES

elongate, distinct, slightly inflated; sutures distinct, 

slightly depressed; wall smooth; aperture narrow, 

with a platelike tooth visible in well-preserved specimens. Length 0.35-0.50 mm; breadth 0.15-0.18 mm.

The specimens from the Cooper marl are very close 

indeed to those figured by D'Orbigny from the Vienna 

Basin.

Possibly the specimen figured from the Eocene Ohio 

and Bed Kiver well 2, Tyler County, Texas, at a depth 

of 1,056 feet,12 may belong here.

Cooper marl: 11861, steep bluff on west side of Biggin 

Creek, Berkeley County, S.C.

Genus VIRGULINA D'Orbigny, 1826 

Virgulina dibollensis Cushman and Applin 

Plate 14, figures 1-3

Virgulina dibollensis Cushman and Applin, Am. Assoc. Petroleum Geologists Bull., vol. 10, p. 168, pi. 7, figs. 7a-c, 

1926.

Howe, Jour. Paleontology, vol. 2, p. 175, 1928.

Howe and Wallace, Louisiana Dept. Conservation Geol. 

Bull. 2, p. 66, pi. 11, fig. 1, 1932.

Test much compressed, very slender, tapering, about 

41/2 times as long as broad, the whole test twisted 

through about 180°, initial end rounded, greatest 

breadth at about three-quarters of the length from the 

initial end; periphery rounded; chambers distinct, 

15 or more, early ones spiral, later ones biserial, the 

last three chambers making up half the test; sutures 

oblique, distinct, slightly depressed; wall thin, punctate, the punctae in longitudinal rows; aperture elongate, elliptical. Length, 0.60 mm; maximum breadth, 

0.12-0.15 mm.

This is a much slenderer species than V. sohreibersiana Czjzek, from the Vienna Basin. It has many 

more chambers, and the ornamentation of punctae 

arranged in linear series is distinctive.

The type specimens were from Haynes well 1, near 

Burkeville, Newton County, Tex., at a depth of 3,175 

to 3,270 feet.

Ocala limestone: 6722, three-fourths mile northwest of Monroeville, Ala.; 6730, Double Branches Creek, Ala.; 6741, 3% 

miles northeast of Brooklyn, Ala.

Virgulina recta Cushman

Plate 14, figures 4a, b

Virgulina, recta Cushman, Contr. Cushman Lab. Foram. Research, vol. 9, p. 12, pi. 1, figs. 31a, b, 1933.

Virgulina sp.?, Cushman, U.S. Geol. Survey Prof. Paper'129, 

p. 92, pi. 16, figs. 2, 3, 1922.

Test elongate, not much compressed, tapering at 

the initial end, sides for the remainder of the test 

nearly parallel, apertural end broadly truncate; chambers distinct, high, 6 biserial ones making up the 

larger part of the test; sutures distinct, slightly de12 Cushman, ,T.C A., and Applin, E. K., Am. Assoc. Petroleum Geologists Bull., vol. 10, pi. 7, flgs. 10, 11, 1926.

pressed, nearly at right angles to the periphery; wall 

smooth; aperture rather broad and large for the genus. 

Length 0.55 mm; breadth 0.15 mm; thickness 0.08 mm.

Holotype (no. 371544, U.S.N.M.) from Cooper marl, 

station 11857, 1 mile south of Moncks Corner, Berkeley County, S.C.

This is the same as the species figured without name 

from the Byram marl of Mississippi. It resembles 

somewhat Virgulina mearicana Cole, from the Guayabal .formation of Mexico, but specimens of that species 

examined show a smaller, much more delicate and 

more compressed form.

Cooper marl: 11861, steep bluff on west side of Biggin 

Creek, Berkeley County, S.C.; 11863, three-fourths mile west 

of Old Biggin Church, Berkeley County, S.C.; 11862, 1 mile 

west of Givhans, Dorchester County, S.C.; 11857, 1 mile south 

of Moncks Corner, Berkeley County, S.C.

Genus BOLIVINA D'Orbigny, 1839 

Bolivina attenuata Cushman

Plate 14, figure 5

Jlolivina attenuata Cushman, Contr. Cushman Lab. Foram. 

Research, vol. 2, p. 30, pi. 4, fig. 4, 1926.

Test very strongly compressed, broad, rapidly increasing in width with newly added chambers, in the 

type specimen about as broad as long; central portion 

slightly thickened; chambers comparatively few, distinct, elongate, curved backward strongly, the inner 

end of each with a slight backward-pointing projection; sutures very distinct, limbate, the periphery of 

the chamber with a slight'keel continuous with the 

suture; earliest chambers with a reticulate ornamentation, later ones smooth. Length 0.40 mm.

This species has a very broad test, much broader 

than any of the others of the upper Eocene except 

perhaps Bolivina frondea Cushman, which it resembles 

in no other particular. The early ornamentation is 

suggestive of B. caelata Cushman, and the later portion with its peculiar backward-projecting inner portion of the chamber is somewhat like B. jacJcsonensis 

Cushman.

Ocala limestone: 1/43, 1% miles southwest of Perdue Hill, 

Ala., the type locality.

Bolivina caelata Cushman var.

At two stations in the deeper-water deposits of the 

upper Eocene there are numerous specimens of a form 

of Bolivina caelata Cushman which more closely resemble var. byramensis than the typical form of the 

species.

All these forms are related to Bolivina reticulata 

Hantken, from the Eocene of Europe, but have a more 

carinate, thinner test. A similar form occurs in the 

Alazan clays of Mexico.

Ocala limestone: 1/38, 3% miles north of Grove Hill, Ala.; 

1/43, 1% miles southwest of Perdue Hill, Ala.
BULIMINIDAE 37

Bolivinu gardnerac Cushman

Plate 14, figures 6, 7

Uolivina cf. 13. punctata Cushman, U.S. Geol. Survey Prof.

Paper 133, p. 19, pi. 2, flg. 1, 1923. 

Uolivina gwdnerae Oushman, Contr. Cushman Lab. Foram.

Research, vol. 2, p. 31, pi. 4, figs. 7a, b, 1926.

Test elongate, compressed, slightly tapering from 

the rounded initial end, periphery rounded, test 

broadly oval in transverse section, sides nearly parallel; chambers numerous, slightly inflated, distinct; 

sutures distinct, depressed; wall coarsely perforate, 

without a definite arrangement of the perforations; 

aperture elongate. Maximum length 0.4C mm.

Bolivma gardtnerae occurs also in tha. lower Vicksburg. It is more compressed, shorter, and broader 

than B. gracttis, described next, and the perforations 

have no definite arrangement.

Ocala limestone: 7719, Spring Mill Branch, Jenkins County, 

Ga., the type locality; 6742, Powells Landing, Sepulga River, 

Ala.; 7159, 4 miles southeast of Whatley, Ala.; 1/38, 3% 

miles north of Grove Hill, Ala.

Bolivina gracilis Cushman and Applin

Plate 14, figures 8-10

Uolivina, gracilis Cushman and Applin, Am. Assoc. Petroleum

Geologists Bull., vol. 10, p. 167, pi. 7, figs. 1, 2, 1926. 

Howe and Wallace, Louisiana Dept. Conservation Geol. 

Bull. 2, p. 57, 1932.

Test small, slender, slightly curved, very little compressed, composed of numerous chambers, distinct, 

slightly inflated; sutures distinct, oblique, slightly 

depressed; wall coarsely perforate, the perforation?; 

often falling into a linear arrangement lengthwise of 

the test; aperture comma-shaped, with a slight trace 

of a lip. Maximum length 0.45 mm.

Bolivina graciUs is one of the group of elongate 

punctate or perforate species represented by B. punctata D'Orbigny. Its nearly circular transverse section distinguishes it from B. g'ardnerae, described 

above. It was originally described from the upper 

Eocene, 4 miles east of Diboll, Angelina County, Tex.

Ocala limestone: 6722, three-fourths of a mile northwest: 

of Monroeville, Ala.; 1/38, 3% miles north of Grove Hill, Ala.

Jackson formation: 7227, Water Valley, Ala.; 6472, 6473, 

Ynzoo City, Miss.

Bolivina jacksonensis Cushman and Applin 

Plate 14, figures 11-13

HoUvina sp.? Cushman, U.S. Geol. Survey Prof. Paper 133,

p. 19, pi. 3, flg. 2, 1923.

Bolivina jacksonensis Cushman and Applin, Am. Assoc. Petroleum Geologists Bull., vol. 10, p. 167, pi.'7, figs. 3, 4, 

1926. 

Howe and Wallace, Louisiana Dept. Conservation Geol.

Bull. 2, p. 59, pi. 11, flg. 11, 1932.

Ellisor, Am. Assoc. Petroleum Geologists Bull., vol. 17, 

no. 11, pi. 3, fig. 3, 1933.

Test much compressed, thickest along the median 

line, periphery slightly keeled or with the keel wanting ; chambers numerous, distinct, both the median and 

peripheral portions extending backward; sutures distinct, slightly depressed; wall thin, translucent, 

smooth, very finely punctate; aperture elongate. Maximum length 0.50 mm.

This is the species figured in the first reference given 

above. The specimen was from the Ked Bluff clay, the 

lowest formation of the Oligocene of Mississippi and 

Alabama. The species is common at numerous stations in the upper Eocene of the Coastal Plain of the 

United States and Mexico. It is related to Bolivina 

mississippiensis Cushman, of the Oligocene Byram 

marl. The type specimens are from Haynes well 1, 

near Burkeville, Newton County, Tex., at a depth of 

a,lT5 to 3>270 feet.

Cooper uurl: 11861, steep bluff on west side of Biggin Creek, 

Berkeley County, S.C.; 6321, Baldock, S.C.

Ocala limestone: 7719, Spring Mill Branch, Jenkins County, 

Ga.; 6723, 6726, 6727, Claiborne, Ala.; 6730, Double Branches 

Creek, Ala.; 6740, mouth of Robinsons Mill Creek, Ala.; 1/8, 

Beck, Ala.; 1/38, 3% miles north of Grove Hill, Ala.; 1/43, 

1% miles southwest of Perdue Hill, Ala.

Jackson formation: 7203, 3% miles southeast of Cullomburg, 

Ala.; 7227, Water Valley, Ala.; 6457, 1% miles south of Shubuta, Miss.; 6463, Jackson. Miss.; 6473, Garlands Creek, Miss.

Bolivina jacksonensis Cushman and Applin var. striatella 

Cushman and Applin

Plate 14, figures 14-18

o Bolivina jacksonensis Cushman and Applin var. striatella

Cushman and Applin, Am. Assoc. Petroleum Geologists 

Bull., vol. 10, p. 167, pi, 7, figs. 5, 6, 1926. 

Ellisor, Am. Assoc. Petroleum Geologists Bull., vol..17, no. 

11, pi. 3, fig. 4, 1933.

Variety differing from the typical form in having 

the early portion with a number of very fine longitudinal costae, without a definite carina; sutures in the 

median portion limbate.

This variety is known from the upper Eocene of the 

Coastal Plain of the United States and Mexico. The 

type specimens were found 4 miles east of Diboll, 

Angelina County, Tex.

Cooper marl: 11861, steep bluff on west side of Biggin 

Creek, Berkeley County, S.C.; 11S63, three-fourths mile west 

of Old Biggin Creek, Berkeley County, S.C.; 11862, 1 mile 

west of Givhans, Dorchester County, S.C.; 11857, 1 mile south 

of Moncks Corner, Berkeley County, S.C.; 6321, Baldock, S.C.

Ocala limestone: 6742, Powells Landing, Sepulga River, Ala.; 

7159, 4 miles southeast of Whatley, Ala.; 7161, 5 miles southeast of Whatley, Ala.; 1/38, 3^ miles north of Grove Hill, 

Ala.; 1/43, 1% miles southwest of Perdue Hill, Ala.

Jackson formation: 6457, 1% miles south of Shubuta, Miss.; 

6463, Jackson, Miss.; 6471, Garlands Creek, Miss.; 6646, Hays 

Chapel, Wayne County, Miss.

Bolivina spiralis Cushman

Plate 15, figures 1, 2

Bolivina spiralis Cushman, Contr. Cushman Lab. Foram. Research, vol. 2, p. 31, pi. 4, figs. 6a, b, 1926.

Test small, only slightly compressed, spirally 

twisted, periphery rounded; chambers numerous, dis-
38 UPPEB EOCENE FOBAMINIFEBA OP THE SOUTHEASTERN UNITED STATES

tinct, last 5 chambers making up half of the test; 

sutures strongly depressed, distinct; wall in the earlier 

chambers roughened and slightly spinose, later ones 

coarsely punctate. Length 0.50 mm.

This species in some respects resembles the Recent 

Indo-Pacific B. tortuosa H. B. Brady but is much 

less compressed and has a roughened exterior, almost 

spinose in the young.

Cooper marl: 6319, Ingleside, S.C., the type locality; 11862, 

1 mile west of Givhans, Dorchester County, S.C.; 11857, 1 

mile south of Moncks Corner, Berkeley County, S.C.

Genus LOXOSTOMUM Ehrenberg, 1854 

Loxostomum dalli (Cushman)

Plate 15, figure 3

Bifarina dalli Cushman, Contr. Cushman Lab. Foram. Research, vol. 2, p. 31, pi. 4, figs. 5a, b, 1926.

Test slightly compressed, elongate, slightly tapering 

from the rounded initial end, apertural end truncate; 

chambers few, distinct, inflated, especially toward the 

apertural end; sutures distinct, compressed; wall ornamented with numerous fine but sharp costae on the 

earlier two-thirds, the last-formed chambers smooth, 

finely punctate; aperture becoming terminal in the 

adult, elliptical or elongate oval. Maximum length 

0.55 mm.

This is a beautifully ornamented species occurring 

at only two stations. The later chambers place the 

species in the genus Loxostonwm, having the aperture 

terminal and the chamber occupying the entire width 

of the test.

Jackson formation: 7220, half a mile southeast of Melvin, 

Ala., the type locality. 

Ocala limestone: 1/38, 3*4 miles north of Grove Hill, Ala.

Genus REUSSELLA Galloway, 1933 

Reussella eocena (Cushman)

Plate 15, figures 4, 5

Reussia, eoceruu Cushman, Contr. Cushman Lab. Foram. Research, vol. 9, p. 13, pi. 1, fig. 25, 1933.

Test short and broad, pyramidal, three-sided, widest 

above the middle, triangular in transverse section, the 

sides in the adult deeply concave, in the young stages 

nearly flat; angles in the young sharp, in the adult 

becoming thick and rounded; surface smooth; aperture at the inner border of the last-formed chamber. 

Maximum length 0.80 mm.

Holotype (no. 371545, U.S.N.M.) from Ocala limestone, station 7365, Pineola, Fla.

This species differs from the two common Vicksburg 

species, Reussella reotimargo (Cushman) and R. spinulosa (Keuss) var. gldbrata (Cushman), in the short, 

broad form and especially the much deeper concavity 

of the sides and the rounded angles. It seems to be 

most abundant in the shallower-water phase, such as is 

represented by the Ocala limestone.

In its very early stages the sides are flat and suggest Reussella spinulosa (Reuss); later it assumes the 

form of var. glabrata (Cushman) but in its adult 

characters becomes very broad, with deeply concave 

sides. The early stages might easily be confused with 

the other forms mentioned, but the adult is distinctive.

Cooper marl: 6321, Baldock, S.C.

Ocala limestone: 7097, Bainbridge, Ga.; 7098, Red Bluff, 

Flint River, Ga.; 7099, half a mile above Red Bluff, Ga.; 7127, 

a quarter of a mile below Normans Ferry, Flint River, Ga.; 

7130, Flint River, 1% miles below Red Bluff, Ga.; 7189, Rich 

Hill, Ga.; 3629, Alachua County, Fla.; 6768, Marianna, Fla.; 

6785, three-fourths mile south of Herlongs, Fla.; 6790, Alachua, 

Fla.; 6790, Alachua Sink, Gainesville, Fla.; 6804, Ocala, Fla.; 

6805, Oakhurst quarry, Ocala, Fla.; 6807, Zuber, Fla.; 6808, 

Martin, Fla.; 6812, 1% miles south of Newberry, Fla.; 7194, 

Marianna, Fla.; 7338, Bowling Springs, Fla.; 7345, Suwannee 

River 2^4 miles above Branford, Fla.; 7348, Suwannee River 

V/z miles above Troy Springs, Fla.; 7349, Suwannee River half 

a mile below Fort McComb, Fla.; 7355, 3 miles northwest of 

Levyville, Fla.; 7364, 1% miles west of Groom, Fla.; 7365, 

Pineola, Fla.; 7366, Anthony, Fla.; 7378, l*/4 miles southwest 

of Inverness, Fla.; 1/48, Duncan, Fla.; 6719, three-fourths mile 

north of Drewry, Ala.; 6722, three-fourths mile northwest of 

Monroeville, Ala.; 6725, Claiborne, Ala.; 6730, Double Branches 

Creek, Ala.; 6738, 6% miles north of Brooklyn, Ala.; 6740, 

mouth of Robinsons Mill Creek, Ala.; 6746, Ward's sawmill, 

Sepulga River, Ala.; 6754, Choctawhatchee River at State line, 

Ala.-Fla.; 6755, Choctawhatchee River above State line, Ala.; 

6756, Choctawhatchee River 3% miles below Geneva, Ala.; 

7159, 4 miles southeast of Whatley, Ala.; 7161, 5 miles southeast of Whatley, Ala.; 1/8, Beck, Covington County, Ala.; 1/21, 

Covington County, Ala.; 1/25, 1 mile east of Harts Bridge, Ala.

Jackson formation: 6463, Jackson, Miss.; 6471, Garlands 

Creek, Miss.

Reussella rectimargo (Cushman)

VerneuUina, rectimargo Cushman, U.S. Geol. Survey Prof. 

Paper 129, p. 127, pi. 29, figs. 4, 5, 1922; Prof. Paper 

133, p. 21, 1923.

Test elongate, triangular in cross section, early portion tapering, adult portion with the sides nearly 

parallel and straight; chambers numerous, arranged 

triserially; sutures not depressed, often slightly limbate; sides of the test flattened or very slightly concave; peripheral angles rounded; apertures slightly 

elongate at the base of the inner margin of the lastformed chamber; wall finely punctate. Length 1.00 

mm or less.

With the one exception noted below, Reussella rectimargo has been found only in the Vicksburg Oligocene. It is related to R. sculptilis and R. eocena, described in this paper.

Ocala limestone: 1/7, 1 mile east of Beck, Ala.

Reussella sculptilis (Cushman)

Plate 15, figures 6, 7

VerneuUina sculptilis Cushman, Contr. Cushman Lab. Foram.

Research, vol. 2, p. 34, pi. 5, fig. 3, 1926. 

Test somewhat longer than broad, pyramidal, threesided, widest at about two-thirds its length, triangular
BULIMINIDAB 39

in transverse section, sides flattened or even slightly 

convex, apical-end tapering, in well-preserved specimens ending in a short point, angles of the test acute; 

wall sculptured, the sutural lines strongly raised, the 

central line of each side of the test marked by a 

strongly raised costa; aperture on the inner border of 

the last-formed chamber. Maximum length 0.50 mm.

This is a beautifully sculptured species. The apical 

end is prolonged into a short spine.

Ocala limestone: 1/39, Pea River at Geneva, Ala.

Genus UVIGERINA D'Orbigny, 1826 

Uvigerina alata Cushman and Applin

Plate 15, figures 8-10

Uvigerina alata Cusliman and Applin, Am. Assoc. Petroleum

Geologists Bull., vol. 10, p. 176, pi. 8, figs. 11-13, 1926. 

Bllisor, Am. Assoc. Petroleum Geologists Bull., vol. 17, 

no. 11, pi. 3, figs. 11, 12,1933.

Test elongate, about twice as long as broad; chambers distinct and inflated; sutures distinct, depressed; 

wall ornamented with a few very prominent, thin, 

high, platelike costae, usually extending posteriorly 

beyond the limits of the chambers; apertural end of 

the last-formed chamber truncate, with a very short, 

cylindrical neck somewhat set down into a depression 

in the chamber wall. Maximum length 0.75 mm; 

breadth 0.30-0.35 mm.

Type specimens from Jackson group in Ohio and 

Red River Avell 2, Tyler County, Tex., at a depth of 

1,056 feet.

Uvigerina cocoaensis Cushman

Plate 15, figures 11-13

Uvigerina cocoaensis Cushman, Contr. Cushman Lab. Foram.

Research, vol. 1, p. 68, pi. 10, fig. 12, 1925. 

Cushman and Applin, Am. Assoc. Petroleum Geologists

Bull., vol. 10, p. 174, pi. 8, fig. 15, 1926. 

Ellisor, Am. Assoc. Petroleum Geologists Bull., vol. 17, 

no. 11, pi. 3, fig. 13, 1933.

Test of medium size for the genus, elongate, fusiform, greatest width slightly above the middle, periphery very slightly lobulate; chambers rather few, 

inflated, evenly rounded; sutures slightly depressed; 

wall ornamented with coarse longitudinal costae, not 

usually confluent with those of the chambers above or 

below, becoming lower and less conspicuous in later 

chambers, the last-formed chamber in the adult usually 

smooth, about 12 to 16 costae in the complete circumference in the widest region; wall finely punctate; 

apertural end with a short cylindrical neck and phialine lip. Maximum length 0.80 mm; width 0.30-0.35 

mm.

This species is nearest to U. jacksonensis, of the 

lower Eocene, but is somewhat smaller, more slender, 

and has fewer costae. It is especially characteristic 

of the Jackson horizon and occurs in the upper Eocene 

of Texas, in the Ohio and Red River well 2, Tyler

County, at a depth of 1,056 feet, and in the Alazan 

clays of Mexico. The type specimen is from Cocoa, 

Ala.

Ocala limestone: 6720, three-fourths mile north of Drewry, 

Ala.; 7159, 4 miles southeast of Whatley, Ala.; 1/38, 3% miles 

north of Grove Hill, Ala.; 1/43, 1% miles southwest of 

Perdue Hill, Ala.

Jackson formation: 2637, Cocoa, Ala.; 7203, 3% miles southeast of Cullomburg, Ala.; 6457, 1% miles south of Shubuta, 

Miss.; 6646, Hays Chapel, Wayne County, Miss.

Uvigerina cookei Cushman, n. sp.

Plate 15, figures 14-16

Test of large size for the genus, elongated, subcylindrical, slightly fusiform, greatest width above 

the middle, periphery slightly lobulate; chambers numerous, inflated, evenly rounded; sutures slightly depressed; wall ornamented with sharp, low, longitudinal costae, in part confluent with those above and 

below, usually reduced on the last-formed chamber, 

about 24 to 30 costae in the complete circumference 

in the widest region; wall finely punctate; apertural 

end with a very short cylindrical neck and wide phialine lip. Maximum length 1.20 mm; width 0.35-0.40 

mm.

Holotype (no. 371546, U.S.N.M.) from Ocala limestone, station 1/38, 3% miles north of Grove Hill, 

Ala.

This species is the largest of those of the upper Eocene; the costae are very numerous but not highly 

developed, and the neck is very short and stout for 

the size of the test. It is related to such species as 

U. tenuistriata Reuss and may be the ancestral form 

of the living U. peregrina var. l>radyana Cushman of 

our eastern coast.

Castle Hayne marl: 5611, Wilmington, N.C.

Cooper marl: 6319, Ingleside, S.C.; Ashley marl works, Dorchester County, S.C.

Ocala limestone: 7719, Spring Mill Branch, Jenkins County. 

Ga.; 7237, half a mile east of Huguenen Ferry, Crisp County, 

Ga.; 6706, Oven Bluff, Tombigbee River, Ala.; 6720, threefourths of a mile north of Drewry, Ala.; 1/21, Covington 

County, Ala.; 1/38, 3% miles north of Grove Hill, Ala.; 1/43, 

iy2 miles southwest of Perdue Hill, Ala.

Jackson formation : 7203, 3% miles southeast of Cullomburg, 

Ala.

Uvigerina dumblei Cushman and Applin

Plate 15, figure 17

Uvigerina dwriblei Cushman and Applin, Am. Assoc. Petroleum 

Geologists Bull., vol. 10, p. 177, pi. 8, fig. 19, 1926.

Test large, periphery rounded, the sides nearly parallel for most of its length, nearly twice as long as 

broad; chambers distinct; sutures slightly depressed; 

wall ornamented by very numerous fine longitudinal 

costae, often 10 to 12' to a single chamber, extending 

backward, making the sutures crenulate; the apertural 

end with a slightly projecting cylindrical neck. Maximum length 1.00 mm.
40 UPPER EOCENE POBAMINIFEEA OP THE SOUTHEASTERN UNITED STATES

Type specimens from Haynes well 1, near Burkeville, Newton County, Tex., at a depth of 3,175 to 

3.270 feet.

Uvigerina gardnerae Cushman

Plate 15, figures 18, 19

Uvigerina gardnerae Cushman, in Cushman and Applin, Am. 

Assoc. Petroleum Geologists Bull., vol. 10, p. 175, pi. 8, 

figs. 16, 17, 1926.

Howe and Wallace, Louisiana Dept. Conservation Geol. 

Bull. 2, p. 63, pi. 12, fig. 6, 1932.

Test of medium size for the genus, much elongated, 

slender, early portion fusiform, later portion with 

the chambers somewhat loosely arranged, peripheiy 

somewhat lobulate; chambers numerous, inflated, especially the later ones, earlier ones with the basal end 

of the chambers tending to overhang the preceding 

ones; wall ornamented with longitudinal costae in the 

earlier portion, the costae not confluent with those of 

chambers above or below, costae later tending to 

break up into lines of spines, and the later portion 

of the test in adults generally hispid, over 20 costae 

in the complete circumference before breaking into 

spines; apertural end with a slightly tapering subcylindrical neck and slight phialine lip. Maximum 

length 0.80 mm; width 0.25 mm.

This species can hardly be confused with any other 

of the upper Eocene species of the Coastal Plain. Its 

characters of ornamentation and the slender form 

distinguish it from all but U. ocalana, which can be 

identified by the very small size and the triangular 

shape of the later chambers.

UiMgerina gardnerae is related to and probably the 

ancestral form of the living U. auberiana D'Orbigny.

Besides the following records for the eastern and 

Gulf Coastal Plain, U. gardnerae occurs in the upper 

Eocene of Texas and in the Alazan clays of Mexico. 

The type specimens are from Ohio and Red River 

well 2, Tyler County, Tex., at a depth of 1,056 feet.

Cooper marl: Ashley marl works, Dorchester County, S.C.

Ocala limestone: 7378, 1% miles southwest of Inverness, 

Fla.; 6706, Oven Bluff, Tombigbee River, Ala.; 7159, 4 miles 

southeast of Whatley, Ala.; 1/26, Drewry, Ala.; 1/38, 3% 

miles north of Grove Hill, Ala.

Jackson formation: 7203, 3% miles southeast of Cullomburg, 

Ala.; 7227, Water Valley, Ala.; 6457, iy2 miles south of Shubuta, Miss.; 6463, Jackson, Miss.; 6646, Hays Chapel, Wi.yne 

County, Miss.

Uvigerina gardnerae Cushman var. texana Cushman and

Applin

Plate 15, figure 20

Uvigerina gar&nerae Cushman var. texana Cushman and Applin, Am. Assoc. Petroleum Geologists Bull., vol. 10, p. 

175, pi. 8, fig. 18, 1926. 

Howe and Wallace, Louisiana Dept. Conservation Geol.

Bull. 2, p. 64, pi. 12, figs. 3, 9, 1932.

Bllisor, Am. Assoc. Petroleum Geologists Bull., vol. 17, no. 

11, pi. 3, fig. 15, 1933.

Variety differing from the typical form in being 

much more elongate; the later portion is somewhat 

less in diameter than the earlier part. Length 0.60 

mm; breadth 0.20 mm.

This variety occurs in the upper Eocene of Texas 

but has not been recognized from the eastern portion 

of the Coastal Plain. The type specimens came from 

a depth of 1,056 feet in Ohio and Red River well 2, 

Tyler County, Tex.

Uvigerina glabrans Cushman 

Plate 15, figure 21

Uvigerina glabrans Cushman, Contr. Cushman Lab. Foram. 

Research, vol. 9, p. 13, pi. 1, fig. 28, 1933.

Test of medium size for the genus, elongated, subcylindrical, or slightly fusiform, greatest width usually below the middle, periphery only slightly lobulate; chambers comparatively few, inflated, evenly 

rounded; sutures very slightly depressed; wall smooth, 

or with faint traces of costae near the initial end, 

finely perforate; apertural end truncate, with a short 

delicate cylindrical neck and phialine lip, the neck 

often broken. Maximum length, 0.75 .mm; width, 

0.30-0.35 mm.

Holotype (no. 371686, U.S.N.M.) from Jackson 

formation, station 7203, Sy2 miles southeast of 

Cullomburg, Ala.

This is the only smooth species of the Coastal Plain 

upper Eocene. It is related to U. cocoaensis and is 

usually found associated with that species but in fewer 

numbers. It is also related to certain of the living 

species of the western Atlantic, usually referred to 

U. canariensis D'Orbigny.

Cooper marl: 11861, steep bluff on west side of Biggin Creek, 

Berkeley County, S.C.; 11862, 1 mile west of Givhans, 

Dorchester County, S.C.

Ocala limestone: 7159, 4 miles southeast of Whatley, Ala.; 

1/26, Drewry, Ala.; 1/38, 3% miles north of Grove Hill, Ala.; 

1/43, li/2 miles southwest of Perdue Hill, Ala.

Jackson formation: 7203, 3V& miles southeast of Cullomburg, 

A.la.

Uvigerina jacksonensis Cushman 

Plate 16, figures 1-3

Ur/igerina jacksonensis Cushman, Contr. Cushmau Lab. Foram.

Research, vol. 1, p. 67, pi. 10, fig. 13, 1925. 

Howe and Wallace, Louisiana Dept. Conservation Geol. 

Bull. 2, p. 65, pi. 12, figs. 7, 8, 1932.

Test large for the genus, stout, broadly fusiform, 

greatest width at about the middle, periphery slightly 

lobulate; chambers rather few, inflated; sutures 

somewhat depressed, basal part of chamber not conspicuously overhanging, evenly curved; wall ornamented with coarse longitudinal costae, in the early 

portion usually limited to the individual chamber, in 

the adult portion usually becoming confluent with
BTJLIMINIDAE 41

those of the adjacent chambers above and below, outer 

edge of the costae entire; about 18 to 22 costae in the 

complete circumference in the widest region; wall 

rather coarsely punctate; apertural end with a tendency in the last-formed chamber to lose or reduce the 

costae, with a cylindrical neck of medium length and 

phialine lip. Maximum length 0.90 mm; width 

0.45 mm.

Uvigerina jacksonensis is a larger, stouter species 

than U. yazooensis. The costae, although nearly as 

numerous, seem to be much fewer on account of the 

larger size of the test. It seems to be characteristic 

of the middle part of the upper Eocene represented by 

the " Zeuglodon bed" of the Jackson formation, 

U.S.G.S. station 2637. It is not found in any numbers 

in the eastern material representing the shallower 

phase of the Ocala limestone, being replaced there by 

Angulogerina ocalana, which is closely related to 

Angulogerina octidentalis Cushman, a species of. shallow, warm waters of the Tortugas region and elsewhere, which also occurs in the Miocene of Florida. 

Uvigerina jacksonensis occurs also in the Alazan 

clays of Mexico.

Cooper marl: 6319, Inglesicle, S. C.; Asbley marl works, Dorchester County, S.C.

Ocala limestone: 7719, Spring Mill Branch, Jenkins County, 

Ga.; 6730, Double Branches Creek, Ala.; 7159, 4 miles southeast of Whatley, Ala.; 1/26, Drewry, Ala.; 1/38, 3% miles 

north of Grove Hill, Ala.; 1/43, 1% miles southwest of Perdue 

Hill, Aln.

Jackson formation: 2637, Cocoa, Ala.; 7203, 3% miles southeast of Cullornburg, Ala.; 6457, 1% miles south of Shubuta, 

Miss.; 6646, Hays Chapel, Wayne County, Miss.

Uvigerina topilensis Cushman

Plate 16, figure 4

Uviyerina toi>ilensis Cushman, Contr. Cusbman Lab. Forarn.

"Research, vol. 1, p. 5, pi. 1, figs. 5a, b, 1925. 

Cnsliman and Applin, Am. Assoc. Petroleum Geologists

Bull., vol. 10, p. 176, pi. 8, fig. 14, 1926. 

Ellisor, Am. Assoc. Petroleum Geologists Bull., vol. 17, no.

11, pi. 3, fig. 14, 1033.

Test generally .fusjform, broadest in the middle, 

initial and apertural ends both rounded; chambers 

irregularly spiral, inflated; sutures distinct, depressed; wall ornamented with a very few costae, progressively decreasing in height toward the apertural 

end of the test and usually continuous from one chamber to another, the last-formed chamber usually 

smooth; wall finely punctate, the costae on the earliest portion sometimes projecting backward into platelike processes; aperture with a very narrow cylindrical neck. Length 0.70 mm; breadth 0.30 mm.

This species, described from the upper Eocene 

clays of Mexico, occurs in the upper Eocene of Texas 

at a depth of 1,056 feet in Ohio and Bed River well 

2, Tyler County.

Uvigerina yazooensis Cushman 

Plate 16, figures 5, 6

Uvigerina yazooensis Cushman, Contr. Cushman Lab. Foram. 

Research, vol. 9, p. 13, pi. 1, fig. 29, 1933.

Test small, elongate, fusiform, greatest width toward the apertural end, periphery strongly lobulate; 

chambers numerous, inflated; sutures strongly depressed, the basal portion of the chamber overhanging 

the preceding ones; wall ornamented with sharp, longitudinal costae, limited to the individual chamber, 

those preceding the suceeding chambers not usually 

in the same line, the outer edge of the costae often 

serrate, about 22 to 26 costae in the complete circumference in the widest region; wall rather coarsely 

punctate; apertural end with a short, narrow cylindrical neck and phialine lip. Maximum length 0.70 

mm; width 0.28 mm.

Holotype (no. 371549, U.S.N.M.) from Jackson formation, station 7220, half a mile southeast of Melvin, 

Ala.

This species is distinct from the others of the upper 

Eocene of the Coastal Plain in the sharp costae, limited to the individual chambers, the very deeply indented sutural regions making a very lobulate periphery. There is apparently no tendency for the 

costae to become obsolete in the last-formed chambers.

Uvigerina yazooensis is apparently the direct ancestral form of some of the species now living in the 

eastern Atlantic, especially U. peregrina Cushman. 

This Recent species has occurred below the 100-f athom 

mark only twice in a large number of stations from 

which it is recorded.

Cooper marl: 11861, steep bluff on west side of Biggin 

Creek, Berkeley County, S.C.; 11863, three-fourths of a mile 

west of Old Biggin Church, Berkeley County, S.C.; 11862, 1 

mile west of Givhans, Dorchester County, S.C.; 11857, 1 mile 

south of Moncks Corner, Berkeley County, S.C.

Jackson formation: 7220, half a mile southeast of Melvin, 

Ala.; 6472, Yazoo City, Miss.; 6463, Jackson, Miss.

Genus ANGULOGEBJNA Cushman, 1927 

Angulogerina ocalana Cushman

Plate 16, figures 7, 8

Angulofferina ocalana Cushman, Contr. Cusbman Lab. Foram. 

Research, vol. 9, p. 14, pi. 1, fig. 30, 1933.

Test small for the genus, elongate, fusiform, periphery very slightly lobulate, somewhat triangular 

in section, the angles rounded, especially in the early 

portion; wall ornamented with numerous very fine, 

slightly raised costae, the outer edge broken into a 

finely serrate line; apertural end with the chambers 

somewhat loosely arranged, the costae less prominent 

or nearly wanting, the chambers more definitely triangular, angles sharper; apertural end extended into
42 UPPER EOCENE POEAMINIFERA OP THE SOUTHEASTERN UNITED STATES

a short neck with a slight lip. Maximum length 0.35 

mm; width 0.15 mm.

Holotype (no. 371550, U.S.N.M.) from upper 

Eocene at station 7719, Spring Hill Branch, Jenkins 

County, Ga.

This is probably the ancestral form of A. byramensis Cushman, so abundant in the Byram marl of the 

lower Oligocene, and these are again ancestral forms 

of the Miocene and living A. occidentalis Cushman 

of the shallow water of the Florida region.

A. ocalana occurs in the lower part of the upper 

Eocene close to the " Scutella bed" in the western 

part of the area, which probably represents shallower 

conditions than the succeeding members. It is also 

found in the Ocala limestone or its equivalent in the 

shallower phase of the eastern part of the area. In 

the Alazan clays of Mexico it occurs very sparingly.

Cooper marl: 11861, steep bluff on west side of Biggin 

Creek, Berkeley County, S.C.; 11863, three-fourths mile west 

of Old Biggin Church, Berkeley County, S.C.; 11862, 1 mile 

west of Givhans, Dorchester County, S.C.; 11857, 1 mile south 

of Moncks Corner, Berkeley County, S.C.

Ocala limestone: 7237, half a mile east of Huguenen Ferry, 

Crisp County, Ga.; 7719, Spring Mill Branch, Jenkins County, 

Ga.; 7378, 1% miles southwest of Inverness, Fla.; 6724, Claiborne, Ala.; 6740, mouth of Robinson Mill Creek, Ala.; 6756, 

Choctawhatchee River S% miles below Geneva, Ala.; 7161, 

5 miles southeast of Whatley, Ala.; 1/33, 7% miles northwest 

of Geneva, Ala.; 1/39, Pea River at Geneva, Ala.; 1/48, Duncan, Fla.

Jackson formation: 6471, Garlands Creek, Miss.

Angulogerina cooperensis Cushman, n. sp.

Plate 16, figures 9a, b

Test small, slender, initial end tapering to a sharp 

point, triangular in section throughout, periphery 

slightly keeled at the three angles; chambers distinct 

in the later part but not inflated or loosely arranged, 

strongly angled; sutures distinct, very slightly if at 

all depressed; wall ornamented with longitudinal costae, usually broken at the sutures, last chambers often 

becoming' somewhat smooth; aperture with a short 

neck and a distinct lip. Length 0.40-0.45 mm; breadth 

0.18-0.20 mm.

Holotype (no. 371551, U.S.N.M.) from Cooper marl, 

station 11863, three-fourths of a mile west of Old 

Biggin Church, Berkeley County, S.C.

This is distinct from Angulogerina ocalana in the 

angular test throughout, A. ocalana having very 

rounded, inflated chambers in the early series. In 

A. oGalan® the last chambers are very loosely arranged 

in the adult; in A. cooperensis they are fairly compact 

throughout. The costae of A. oGalana are finer and 

more numerous.

Cooper marl: 11863, three-fo.urths of a mile west of Old 

Biggin Church, Berkeley County, S.C.; 11857, 1 mile south of 

Moncks Corner, Berkeley County, S.C.

Genus TRIFARINA Cushman, 1923 

Trifarina bradyi Cushman var. advena Cushman

Plate 16, figures lOa, b

Trifarina, bradyi Cushman var. advena Cushman, Contr. Cushman Lab. Foram. Research, vol. 1, pt. 4, p. 87, 1926.

Variety differing from the typical form in being 

slightly smaller and stouter; the sides especially being 

much less concave, and the carinae of the sides much 

less prominent, the surface rougher and the wall 

thicker. Length 0.30 mm.

This variety is most nearly allied to T. ~bradyi from 

the Pacific Ocean.

This is apparently the first record of the typical 

form of the genus from the Eocene. It is a small, 

rather easily overlooked form, but has occurred at 

several stations in the upper Eocene.

Ocala limestone: 6790, Alachua, Fla.; 6739, 6740, mouth of 

Robinsons Mill Creek, Ala.; 1/33, 7% miles northwest of 

Geneva, Ala.

Jackson formation: 6463, Jackson, Miss., type locality.

Family ELLIFSOIDINIDAE 

Genus PLEUROSTOMELLA Reuss, 1860 

Pleurostomella jacksonensis Cushman and Applin

Plate 16, figures lla-c

Pleurostomella jacksonensis Cushman and Applin, Am. Assoc. 

Petroleum Geologists Bull., vol. 10, p. 168, pi. 7, figs. 

9'a-c, 1926.

Test elongate, tapering, broadest at about the 

middle of the last-formed chamber, circular in transverse section; chambers comparatively few, alternating, distinct; sutures slightly depressed; wall marked 

by series of depressions in longitudinal lines; aperture of the usual type in this genus. Length 0.50 mm..

The only record of this species is from the upper 

Eocene of the Coastal Plain of Texas. The specimens are from a depth of 3,175 to 3,270 feet in Haynes 

well 1, near Burkeville, Newton County.

Family ROTALIIDAE

Genus SPIRILLINA Ehrenberg, 1841

Spirillina vivipara Ehrenberg

Plate 16, figure 12 -

Spirillina vivipwa Ehrenberg, Akad. Wiss. Berlin Abh., 1841,

p. 442, pi. 3, fig. 41. 

Moebius, Beitrage zur Meeresfauna der Insel Mauritius,

p. 88, pi. 8, figs. 1, 2, 1880. 

H. B. Brady, Challenger Rept, Zoology, vol. 9, p. 630, pi.

85, figs. 1-5, 1884. 

Flint, U.S. Nat. Mus. Rept. for 1897, p. 326, pi. 71, fig. 4,

1899.

Millett, Roy. Micr. Soc. Jour., 1903, p. 693. 

Rhumbler, Zool. Jahrb., Abt. Syst, vol. 24, p. 32, pi. 2,

fig. 7, 1906. 

Cushman, U.S. Nat. Mus. Bull. 71, pt. 5, p. 3, pi. 1, figs.

1, 2, fig. 1 (in text), 1915.
ROTALIIDAE 43

Test planispiral, composed of a few coils not divided into chambers, coils flattened and somewhat 

broader in the later portion, both sides somewhat 

flattened, the periphery rounded; wall conspicuously 

perforate. Diameter of single specimen 0.30 mm.

There is a single specimen from U.S.G.S. station

6725. Ocala limestone, Claiborne, Ala., which probably 

can best be referred to this species, although the coils 

are not as much flattened as in the Kecent specimens, 

nor are the perforations as large. 

There are specimens of Spirillina also from stations

6726. Claiborne, Ala., and 1/8, Beck, Ala., Ocala limestone, but they are not sufficiently well preserved to 

warrant specific determination.

Genus DISCORBIS Lamarck, 1804 

Discorbis globulo-spinosa Cushman

Plate 16, figures 14a-c

DiscorMs globulo-spinosa Cushman, Contr. Cushman Lab. 

For'am. Research, vol. 9, p. 14, pi. 2, figs, la-c, 1933.

Test small, ventral side flat, dorsal side strongly 

convex, composed of several whorls, last-formed one 

with 5 chambers, ventral peripheral angle sharp and 

somewhat keeled; chambers of the early whorls somewhat/indistinct, later ones more so, narrow and high, 

and inner portion on the dorsal side produced into a 

distinct, raised ridge, which often becomes spinose in 

the central portion; sutures fairly distinct, slightly 

depressed and very oblique on the dorsal side, on the 

ventral side nearly radial; wall coarsely perforate 

both on the dorsal and ventral sides, ventral side 

smooth; aperture a curved, arched opening on the 

ventral side of the test, extending in a curve toward 

the umbilical area. Diameter 0.30 mm; thickness 

0.20-0.25 mm.

Holotype (no. 371553, U.S.N.M.) from upper 

Eocene, Jackson, Miss.

Castle Hayne marl: 5611, Wilmington, N.C.

Ocala limestone: 6725, Claiborne, Ala.; 7161, 5 miles southeast of Whatley, Ala.; 1/23, Claiborne, Ala.

Jackson formation: 7227, Water Valley, Ala.; 6463, Jackson, 

Miss.; 6471, Garlands Creek, Miss.

This species is related to those of the Indo-Pacific 

region, more especially to some of those described 

from the late Tertiary of Australia.

Discorbis hemisphaerica Cushman 

Plate 16, figures 13a-c

Disoorlis hemisphaerica Cushman, Contr. Cushman Lab.

Foram. Research, vol. 7, p. 59, pi. 7, figs. 14a-c, 1931. 

Bllisor, Am. Assoc. Petroleum Geologists Bull., vol. 17, 

no. 11, pi. 3, figs. 17, 18, 1933.

Test small, hemispherical, the ventral side flattened, dorsal side strongly convex, composed of several whorls, last-formed one with 4 chambers, ventral 

peripheral portions rounded; chambers of the early

whorls somewhat covered by a secondary clear 

growth, later chambers more distinct, somewhat narrowed, smooth; sutures distinct, oblique, slightly depressed, on the ventral side radial; wall coarsely and 

conspicuously perforate, less distinctly so on the ventral side, which is peculiar, as the last-formed whorl 

of chambers only partly covers the earlier ones; aperture a curved arched opening on the ventral side of 

the test from near the periphery, well back from the 

umbilical region, developing a slight raised lip and 

apertural face. Diameter 0.35 mm; thickness 0.20- 

0.30 mm.

Holotype (no. 371552, U.S.N.M.) from upper Eocene, station 6463, Jackson, Miss.

Ocala limestone: 7161, 5 miles southeast of Whatley, Ala.; 

1/7, 1 mile east of Beck, Ala.; 1/8, Beck, Ala.

Jackson formation: 7227, Water Valley, Ala.; 6463, Jackson, Miss.; 6471, Garlands Creek, Miss.

The ventral side of the test is very peculiar in the 

slight overlapping of the last-formed whorl, leaving 

the earlier chambers exposed in the umbilical region.

Discorbis subaraucana Cushman

Plate 18, figures la-c

DiscorMs subaraucana Cushman, Carnegie Inst. Washington 

Pub. 311, p. 41, pi. 7, figs. 1, 2, 1922.

Test unequally biconvex, dorsal side somewhat convex, ventral side very slightly convex, flattened, or 

even somewhat concave, umbilicate, periphery 

rounded, entire except the last part, which may be 

very slightly lobulate; chambers about 6 in the adult 

whorl, rapidly increasing in size as added, very 

slightly inflated; sutures oblique, curved, earlier ones 

strongly limbate; wall smooth, finely but conspicuously perforate; aperture at the umbilical end of the 

chamber, with a slight overhanging lip. Diameter 

0.50-0.60 mm; thickness 0.15-0.20 mm.

This species was described from off the southern 

coast of Florida. The types are very close indeed to 

the specimen figured here, and they seem specifically 

identical.

Jackson formation: 7220, half a mile southeast of Melvin, 

Ala.

Ocala limestone: 7704, 3 miles west of Tarversville, Ga.; 

6742, Powells Landing, Sepulga River, Ala.; 1/38, 3% miles 

north of Grove Hill, Ala.

Discorbis bulla Cushman

Plate 17, figures 6a-c

Discorbis bulla Cushman, Contr. Cushman Lab. Foram. Research, vol. 9, p. 16, pi. 2, figs. 6a-c, 1933.

Test much compressed except at the umbo, which 

is distinctly elevated, ventral side convex but umbilicate, periphery acute; chambers few, much elongate, 

narrow, strongly curved, three completing a whorl, 

the last-formed one making up more than half the 

periphery; sutures distinct, very strongly curved on
UPPER EOCENE PORAMINIFEEA OF THE SOUTHEASTERN UNITED STATES

the dorsal side, somewhat limbate, ventrally slightly 

depressed, curved; wall on the dorsal side smooth, 

ventrally with traces of radial markings; aperture 

opening into the umbilical area. Diameter 0.50-0.60 

mm; height 0.15-0.18 mm.

Holotype (no. 371554, U.S.N.M.) from U.S.G.S. 

station 1/39, Ocala limestone, Pea Kiver at Geneva, 

Ala.

This is a peculiar-shaped species with a very prominent umbo, convex ventral side, and very long curved 

chambers. It belongs in the group usually assigned 

to Discorbis orbicularis (Terquem).

Ocala limestone: 1/39, Pea River, Geneva, Ala.; 6727, Claiborne, Ala.

Discorbis ocalana Cushman

Plate 16, figures 15a-c

Dtscorbis ocalana Cushman, Contr. Gushrnan Lab. Foram. Research, vol. 9, p. 15, pi. 2, figs. 5a-c, 1933.

Test biconvex, dorsal side slightly more convex than 

the ventral, which is slightly umbilicate, periphery 

rounded; chambers usually 5 in the adult whorl, those 

of the last whorl distinct, earlier ones obscure, gradually increasing in size as added, slightly inflated; 

sutures gently curved dorsally, ventrally nearly 

radial, slightly depressed on both sides; wall thick, 

smooth; aperture ventral, toward the umbilicus. 

Diameter 0.60 mm; height 0.20 mm.

Holotype (no. 371555, U.S.N.M.) from Ocala limestone, station 6768, Marianna, Fla.

This is a thick-walled species which occurs in the 

Ocala limestone but differs from numerous other 

species of this formation in not being usually well 

preserved.

Ocala limestone: 6768, Marianna, Fla.

Discorbis assulata Cushman

Plate 17, figures 1, 2

Discorbis assulata Cushman, Contr. Cushman Lab. Foram. Research, vol. 9, p. 15, pi. 2, figs. 2a-c, 1933.

Test very much compressed, planoconvex, ventral 

side flat or concave, dorsal side very slightly convex, 

periphery subacute, slightly keeled; chambers usually 

6 in the final whorl, of uniform shape but increasing 

rather rapidly in size as added; sutures distinct, 

limbate, about evenly curved on the dorsal and ventral sides, the later one or two on the ventral side 

with a liplike projection; wall smooth, finely perforate; aperture ventral toward the umbilicus beneath 

a slight overhanging lip. Diameter 0.50 mm; height 

0.10 mm.

Holotype (no. 371556, U.S.N.M.) from U.S.G.S. station 3966, Ocala limestone, 2 miles south of Perry, 

Ga.

This is a very thin scalelike species with distinctly 

limbate sutures, which are evenly curved on the two 

sides.

Ocala limestone: 3966, 2 miles south of Perry, Ga.; 6727,

Claiborne, Ala.; 1/8, Beck, Ala.

Discorbis alabamensis Cushman

Plate 17, figures 3a-c

Disoorbis alabamensis Cushman, Contr. Cushman Lab. Foram. 

Research, vol. 9, p. 16, pi. 2, figs. 3a-c, 1933.

Test small, unequally biconvex, ventral side slightly 

convex, dorsal side more strongly so, periphery 

rounded; chambers distinct, usually about 6 in the 

final whorl, slightly inflated, of uniform shape, increasing very slightly in size as added, on the ventral side 

with the inner end of the chambers lobed; sutures distinct, slightly depressed, gently curved; wall smooth; 

aperture near the umbilicus, with a slight overhanging 

lip. Diameter 0.25 mm; height 0.06 mm.

Holotype (no. 371557, U.S.N.M.) from station 1/8, 

Ocala limestone, Beck, Ala.

This is a very small but distinctive species, especially when seen from the ventral side.

Ocala limestone: 1/8, Beck, Ala.

Discorbis alveata Cushman

Plate 17, figures 4a-c

Discorbis alveata Cushman, Contr. Cushman Lab. Foram. Research, vol. 9, p. 16, pi. 2, figs. 4a-c, 1933.

Test planoconvex, dorsal side raised in a low spire, 

ventral side nearly flat, periphery acute and slightly 

keeled; chambers distinct, about 5 in the adult whorl, 

of uniform shape, increasing gradually in size as 

added, the ventral side with the inner portion broken 

up into a series of channels radiating from the umbilicus; sutures distinct, slightly limbate on the dorsal 

side, strongly curved, flush with the surface, on the 

ventral side nearly radial, slightly curved and distinctly depressed; wall smooth except for the channeling of the umbilical area on the ventral side; aperture 

narrow, at the umbilical end of the chamber. Diameter 0.35 mm; height 0.12 mm.

Holotype; (no. 371558, U.S.N.M.) from U.S.G.S. 

station 6471, Jackson formation. Garlands Creek, Miss.

This is a small, distinctive species of few chambers, 

one of the main characteristics being the channeling of 

the ventral side. It is common at the type' locality.

Jackson formation: 6471, Garlands Creek, Miss.

Genus VAIVULINERIA Cushman, 1926

Valvulineria jacksonensis Cushman

Plate 18, figures 2a-c

Valvulineria jacksonensis Cushman, Contr. Cushmau Lab. 

Foram. Research, vol. 9, p. 18, pi. 2, figs. 9a-c, 1933.

Test biconvex, compressed, dorsal side with a very 

low spire, ventrally convex toward the periphery, but 

depressed at the umbilicus, which is somewhat finely
EOTALIIDAE 45

papillate, periphery rounded; chambers distinct, about 

8 in the adult whorl, of uniform shape, gradually 

increasing in size as added, not inflated; sutures distinct, on the dorsal side gently curved, limbate, not 

depressed, ventrally nearly straight, oblique, very 

slightly depressed; wall smooth; aperture ventral beneath the umbilicate lobe of the last-formed chamber. 

Length 0.50 mm; breadth 0.40 mm; thickness 0.18 mm.

Holotype (no. 371559, U.S.N.M.) from Ocala limestone, U.S.G.S. station 6741, 3% miles northeast of 

Brooklyn, Ala.

This is a characteristic species, probably occurring 

at more stations than here recorded, but the preservation of the material does not always allow certain 

determination.

Ocala limestone: 7189, Rich Hill, Ga.; 6741, 3% miles northeast of Brooklyn, Ala.

Genus LAMARCKINA Berthelin, 1881

Lamarckina ocalana Cushman

Plate 17, figures 5a-c

Lamarckinot ocalana Cushman, Contr. Cushman Lab. Forani. 

Research, vol. 2, p. 11, pi. 1, figs. 4a-c, 1926.

Test subcircular, periphery slightly lobulate, thin, 

slightly biconvex, periphery very thin, slightly keeled, 

composed of about iy2 coils, 7 chambers in the lastformed coil; dorsal side smooth and very finely perforate, the sutures barely depressed, not limbate; 

ventral side flattened or slightly convex, umbilicate, 

smooth and highly polished, sutures barely depressed; 

aperture large with a broadly rounded projecting lip: 

Diameter about 1.00 mm; thickness 0.35 mm.

This is the largest of the species of the genus and 

may be distinguished by its smooth, unornamented 

dorsal surface, tendency to become keeled, and compressed form.

Ocala limestone: 1/19, near Blue Springs, Jackson County, 

Pla.

Lamarckina jacksonensis Cushman

Plate 20, figures 14a-c

Lamarckina jacksonensis Cushman, Contr. Cushman Lab. 

Forum. Research, vol. 9, p. 20, pi. 2, figs. lOa-c, 1933.

Test about equally biconvex, periphery acute, 

slightly keeled; chambers about 5 in the adult whorl, 

rapidly increasing in size as added, ventrally somewhat inflated; sutures on the dorsal side slightly 

curved, very oblique, limbate and raised, on the ventral side curved, either flush with the surface or 

slightly depressed; walls smooth except for the raised 

sutures on the dorsal side, on the ventral side smooth 

and polished; aperture a large opening in a deep reentrant of the ventral side. Length 0.45-0.50 mm; 

breadth 0.35-0.40 mm; thickness 0.15 mm.

Holotype (no. 371571, U.S.N.M.) from U.S.G.S. 

station 1/39, Ocala limestone, Geneva, Ala.

This species is most closely related in the shape of 

its chambers to Lamarcldna ocalana Cushman but differs from that species in the much more oblique sutures, fewer chambers, and less convex form.

Ocala limestone: 1/39, west bank of Pea River at Geneva, 

Ala.

Genus GYKOIDINA D'Orbigny, 1826

Gyroidina soldanii D'Orbigny var. octocamerata Cushman 

and G. D. Hanna

Plate 18, figures 4a-c

Gyroidina, soldanii D'Orbigny var. octocamerata Cushman and

G. D. Hauna, California Acad. Sci. Proc.,~ser. 4, vol. 16,

p. 223, pi. 14, figs. 16-18, 1927. 

Cole, Bull. Am. Paleontology, vol. 14, no. 51, p. 29, pi. 2,

figs. 22-24, 1927. 

Cushman and Schenck, California Univ. Dept. Geol. Sci.

Bull., vol. 17, p. 312, pi. 44, figs. 3-5, 1928. 

Weinzierl and Applin, Jour. Paleontology, vol. 3, no. 4,

p. 406, 1929. 

Cushman and Thomas, Jour. Paleontology, vol. 4, p. 40,

pi. 4, figs. 2, 3, 1930.

Test small, dorsal side flattened, ventral side much 

convex, composed of about 3 coils, the last-formed one 

consisting of 8 chambers, periphery broadly rounded, 

ventral side with the umbilical region strongly depressed; chambers distinct; sutures distinct, slightly 

depressed, on the dorsal side somewhat oblique, on the 

ventral side radial; wall finely perforate, smooth and 

polished; aperture elongate, arched, from the periphery at least halfway to the umbilicus along the 

ventral border of the last-formed chamber, with a 

very slightly developed lip. Length 0.50 mm; 

breadth 0.45 mm; thickness 0.35 mm. The 8 chambers and small size are characteristic of all the specimens, and there seems to be very little variation.

The type specimen is from U.S.G.S. station 1/39, 

Ocala limestone, Geneva, Ala.

Castle Hayne marl: 5611, Wilmington, N.C.; 8171, Richlands, N.C.

Ocala limestone: 1/21, southeastern Covington County, Ala.; 

1/39, Pea River at Geneva, Ala.

This variety occurs also in the Claiborne group, as 

well as in the equivalent Eocene of Mexico and California.

Gyroidina orbicularis D'Orbigny var. planata Cushman, 

n. var.

Plate 18, figures 3a-c

Test planoconvex, the dorsal side very flat, the ventral very strongly convex; periphery forming a blunt 

angle, the last-formed coil either at the same level 

as the previous ones or falling slightly below, occasionally rising very slightly above, ventrally distinctly
46 UPPEE EOCENE FOBAMINIFEEA OP THE SOUTHEASTERN UNITED STATES

umbilicate ; chambers fairly distinct on the dorsal side, 

especially those of the last-formed coil, which has 10 

or 11 chambers ; in peripheral view with the periphery 

broadly rounded, the dorsal angle blunt, the ventral 

side strongly curved, sometimes projecting slightly to 

form a slight angle ; sutures on the dorsal side oblique, 

only those of the last-formed coil distinct ; on the ventral side radial and depressed; wall smooth and polished; aperture extending from the peripheral angle 

nearly to the umbilicus, low, without a lip. Diameter 

0.50 mm; thickness 0.30-0.35 mm.

Holotype (no. 371560, U.S.N.M.) from Cooper marl, 

Ingleside, S.C.

Cooper marl: 6319, Ingleside, S.C. (common) ; 11861, steep 

bluff on west side of Biggin Creek, Berkeley County, S.C. ; 

11863, three-fourths of a mile west of Old Biggin Church, Berkeley County, S.C. ; 11862, 1 mile west of Givhans, Dorchester 

County, S.C. ; 11857, 1 mile south of Moncks Corner, Berkeley 

County, S.C.

Genus ROTALIA Lamarck, 1804 

Rotalia sp.?

Plate 19, figures lla-c

A fairly large sized species of Rotalia occurs commonly in the Ocala limestone. The dorsal side has a 

high conical form; the ventral side is flattened. Although this species is common, it is not well preserved, 

and it has not been deemed advisable to make a specific 

determination.

OcaLa limestone : 6812, 

County, Ma.

miles south of Newberry, Alachua

Genus EPONIDES Montfort, 1808 

Eponides jacksonensis (Cushman and Applin)

Plate 19, figures 4-8

Pulvmulvna jacksonensis Cushman and Applin, Am. Assoc. Petroleum Geologists Bull., vol. 10, p. 181, pi. 9, figs. 24, 

25, 1926.

Test with a fairly high spire, the periphery not 

keeled but only slightly rounded, the dorsal side much 

more convex than the ventral; chambers numerous, 6 

or 7 in the last-formed coil; sutures distinct but on 

the dorsal side very oblique, on the ventral side only 

slightly curvecl and somewhat depressed; wall 

smooth; aperture on the ventral side, forming a distinct angle in the border of the test and extending to 

the umbilicus. Diameter 1.00 mm.

The types of this species are from the upper Eocene 

4 miles east of Diboll, Angelina County, Tex. It 

also occurs in Texas on Tarkiln Creek, half a mile 

above the Neches River, Trinity County, and in well 

samples. The species is widely distributed in the 

upper Eocene of the Coastal Plain of the United

States eastward from Texas and in the Alazan clays 

of Mexico.

Eponides jacksonensis is related to E. byramensis 

Cushman, from the lower Oligocene of the Coastal 

Plain, but has more whorls and fewer chambers, as 

well as being of smaller size on the average. It is 

also related to E. mexicana Cushman, which has more 

chambers and is a much more highly ornamented 

form of larger size. All these species are related to 

E. haidingeri D'Orbigny, from the Tertiary of 

Europe, but are all distinct.

Cooper marl: 5531, Charleston, S.C.; 6321, Baldock, S.C.

Ocala limestone: 7189, Rich HiU, Ga.; 7237, half a mile 

east of Huguenen Ferry, Ga.; 7719, Spring Mill Branch, Jenkins County, Ga.; 4974, Duncan, Fla.; 6768, Marianna, Fla.; 

6790, Alachua, Fla..; 6805, Oakhurst quarry, Ocala, Fla.; 6807, 

Zuber, Fla.; 6808, Martin, Fla.; 6815, Clark, Fla.; 7194, Marianna, Fla.; 6720, three-quarters mile north of Drewry, Ala.; 

6738, Qy2 miles north of Brooklyn, Ala.; 7159, 4 miles southeast of Whatley, Ala.; 7161, 5 miles southeast of Whatley, 

Ala.; 1/7, 1 mile east of Beck, Ala.; 1/39, Pea River at 

Geneva, Geneva County, Ala.; 1/48, Duncan, Fla.

Jackson formation: 7203, 3% miles southeast of Cullomburg, Ala.; 6457, 1% miles south of Shubuta, Miss.; 6463, 

Jackson, Miss.; 6471, Garlands Creek, Miss.

Some specimens from the Cooper marl of South 

Carolina that are referred to this species have a very 

low spire and a subacute periphery and may be found 

worthy of varietal distinction with further collecting. 

They came from the following U.S.G.S. stations:

Cooper marl: 11863, three-quarters mile west of Old Biggin 

Church, Berkeley County, S.C.; 11861, steep bluff on west side 

of Biggin Creek, Berkeley County, S.C.; 11857, 1 mile south 

of Moncks Corner, Berkeley County, S.C.

Eponides carolinensis Cushman, n. sp.

Plate 17, figures 7a-c

Test comparatively large, biconvex, umbonate, periphery with a rounded thickened margin; chambers 

numerous, about 12 in the last-formed coil in the adult, 

the last few more distinct than the earlier ones; sutures 

indistinct except in the last portion, strongly curved 

on the dorsal side, slightly so on the ventral, sometimes 

limbate and fusing with the thickened peripheral 

margin; wall distinctly perforate, thickened with an 

outer heavy wall showing little structure but when 

broken away revealing the distinct perforations and 

the sutures below; chambers in the thickened specimens ending in a ring of thickened beadlike projections about the umbilical area, which is also filled to 

form a distinct cross; aperture an elongate opening 

on the ventral side at the inner margin of the lastformed chamber, sometimes with a slightly thickened 

lip and occasionally with secondary rounded apertures in the apertural face. Diameter 1.85 mm; 

thickness 1.15 mm.
EOTALIIDAE 47

Holotype (no. 371562, U.S;N.M.) from the upper 

Eocene Castle Hayne marl, station 5611, Wilmington, 

N.C., where the species is common.

It is the largest of the upper Eocene forms of the 

Coastal Plain area, and is peculiar in the thickened 

outer layers, which largely conceal the structures of 

the test.

Castle Hayne marl: 5611, Wilmington, N.C.

Eponides cocoaensis Cushman

Plate 19, figures 1, 2

Eponides cocoaensis Cushrnan, Contr. Cushrnan Lab. Foram. 

Research, vol. 4, p. 73, pi. 10, figs. 2a-c, 1928.

Test planoconvex, dorsal side flattened or very 

slightly umbonate, ventral side very strongly convex; 

chambers few, usually four in the last-formed coil, the 

last one large and occupying nearly half the surface 

on the ventral side; periphery subacute or even with 

a blunt keel; sutures distinct and depressed, on the 

ventral side nearly radiate, on the dorsal curved; wall 

in well-preserved specimens rather distinctly perforate, although the perforations are not coarse, and 

occasionally with small spines or papillae scattered 

over the surface; aperture elongate, semielliptica-1, at 

about the middle of the inner margin of the chamber 

on the ventral side. Diameter 0.50 mm; thickness 

0.45 mm.

The type specimens are from the Jackson formation 

al station 2637, Cocoa, Ala.

Ocala limestone: 7191, Marianna, Fla.; 7159, 4 miles southeast of Whatley, Ala.

Jackson formation: 2637, Cocoa, Ala.; 6457, iy2 miles south 

of Shubuta, Miss.

Cooper marl: 6321, Baldock, S.C.; 11863, three-fourths mile 

west of Old Biggin Church, Berkeley County, S.C.; 11857, 

1 ruile south of Moncks Corner, Berkeley County, S.C.

Eponides budensis (Hantken) var. planata Cushman, n. var.

Plate 18, figures 6a-c

Test planoconvex, dorsal side moderately convex, 

ventral side flat or even slightly concave, periphery 

acute, slightly keeled; chambers 5 or 6 in the final 

whorl, of uniform shape, gradually increasing in size 

as added, not inflated; sutures on the dorsal side flush 

with the surface, strongly oblique, ventrally curved, 

and slightly depressed; wall smooth; aperture on the 

ventral side midway between the umbilicus and periphery. Diameter 0.45-0.50 mm; thickness 0.12- 

0.15 mm. »

Holotype (no. 371563, U.S.N.M.) from Castle 

Hayne marl, station 5611, Wilmington, N.C.

This variety in many respects resembles Hantken's 

species, but that is biconvex, whereas our form is very 

decidedly planoconvex or even slightly concave on the 

ventral side.

Castle Hayne marl: 5611, Wilmington, N.C.

G2330--35 4

Eponides ocalana Cushman

Plate 18, figures 5a-c

Eponides ocalana Cushman, Contr. Cushman Lab. Foram. 

Research, vol. 9, p. 17, pi. 2, figs. 7a-c, 1933.

Test comparatively large, strongly planoconvex, periphery rounded, dorsal side flat, ventral side very 

convex; chambers 5 or 6 in the last-formed coil, distinct; sutures on the dorsal side curved, limbate, 

raised above the surface and often connecting with a 

raised ridge about the periphery, on the ventral side 

slightly depressed, very slightly curved, ending in the 

rather large umbilical cavity; wall thick, matte; 

aperture a short semielliptical opening near the umbilical end of the ventral margin of the chamber. 

Length 1.00-1.25 mm; breadth 0.75-0.85 mm; thickness 0.50-0.60 mm.

Holotype (no. 371561, U.S.N.M.) from Ocala limestone at U.S.G.S. station 1/21, Covington County, 

Ala. This rather large, heavy-walled species seems to 

occur only in the Ocala limestone.

Ocala limestone: 7194, Marianna, Fla.; 1/48, Duncan, Fla.; 

3/21, Covington County, Ala.; 1/25, 1 mile east of Harts 

Bridge, Ala.; 1/26, Drewry, Ala.

Eponides pygmaea (Hantken)

Plate 19, figures 9a-c

There are rare specimens from the upper Eocene 

Jackson group north of Whitsett, Live Oak County, 

Tex., one of which is here figured and which may be 

referred to the above-named species of Hantken. 

These have already been noted elsewhere.13 They 

have not occurred in the material from the eastern 

part of the region.

Eponides minima Cushman

Plate 19, figures 3a-c

Eponides minima Cushman, Contr. Cushman Lab. Foram. 

Research, vol. 9, p. 17, pi. 2, figs. 8a-c, 1933.

Test small, trochoid, spire low, periphery bluntly 

keeled, lobulate; chambers typically 6 or 7 in the 

adult whorl, of uniform shape, gradually increasing 

in size as added; sutures on the dorsal side obliquely 

curved, slightly limbate, flush with the surface, on 

ventral side slightly curved, slightly depressed; wall 

smooth dorsally, on the ventral side with papillae 

somewhat obscuring the sutures over the umbilical 

portion; aperture ventral, between the periphery and 

the umbilical area. Diameter 0.28-0.30 mm; height 

0.12 mm.

Holotype (no. 371564, U.S.N.M.) from Cooper 

marl, station 11863, Berkeley County, S.C.

10 Cushman, J. A., and Applln, B. R., Am. Assoc. Petroleum Geologists Bull., vol. 10, p. 179, pi. 9. flg. 14, 1926.
48 UPPEK EOCENE FORAMINIFEFvA OF THE SOUTHEASTERN UNITED STATES

This is a very small species, probably nearest to the 

Eecent Eponides easigiM (H. B. Brady) but differing 

from that species in the sutures of the dorsal side, 

which in Brady's species are strongly limbate, especially the spiral suture, and on the ventral side, which 

is smooth in Brady's species but in ours is very 

markedly papillate.

Cooper marl: 11863, three-fourths mile west of Old Biggin 

Church, Berkeley County, S.C.; 11862, 1 mile west of Givhans, 

Dorchester County, S.C.

Eponides umbonata (Reuss)

Plate 19, figures lOa-c

RotaUna umbonato, Reuss, Deutsch. geol. Gesell. Zeitschr., 

vol. 3, p. 75, pi. 5, fig. 35, 1851.

Wvonides utinbonata Cole, Bull. Am. Paleontology, vol. 14, p.

15, pi. 2, fig. 6, 1928.

Cushman, Contr. Cushman Lab. Forain. Research, vol. 5, 

p. 98, pi. 14, figs. 8a-c, 1929.

Rotalia umbonata, Galloway and Morrey, Bull. Am. Paleontology, vol. 15, no. 55, p. 26, pi. 4, figs, la-c, 1929.

Rotalia ecitadorensis Galloway and Morrey, idem, p. 26, pi. 3, 

figs. 13a-c.

Test trochoid, biconvex, periphery acute; chambers 

usually 5 or 6 in the adult whorl, of uniform shape, 

slightly increasing in size as added, not inflated; 

sutures on the dorsal side straight and at right angles 

to the periphery, slightly limbate, flush with the surface, on the ventral side typically with a sigmoid 

curve most pronounced near the inner end; wall 

smooth; aperture elongate, on the ventral side between the periphery and the umbilical area. Diameter about 0.50 mm.

This species, which has a general range from Oligocene to Recent, has occurred in the Eocene collections at but one locality.

Cooper marl: 11863, three-fourths mile west of Old Biggin 

Church, Berkeley County, S.C.

Gemis CANCRIS Montfort, 1808 

Cancris brongniartii (D'Orbigny)

Plate 20, figures la-c

Rotalia brongniartii D'Orbigny, Annales sci. nat, vol. 7, p.

273, Modeles, no. 27, 1826. 

Rotalwia brongniartii Reuss, in Geinitz, Grundriss der Versteinerungskunde, p. 673, pi. 24, fig. 55, 1845-^6. 

D'Orbigny, Foraminiferes fossiles du bassin tertiaire de

Vienne, p. 158, pi. 8, figs. 22-24, 1846.

Test biconvex, periphery subacute, longer than 

broad; chambers increasing rapidly in size and length 

as added in the adult whorl, all visible from the dorsal side, from the ventral side only those of the final 

whorl, the last-formed chamber making a large proportion of the surface; wall smooth, the last-formed 

chamber with an oval thinner area toward the base 

on the ventral side; aperture narrow, beneath the inner end of the chamber on the ventral side. Length 

0.85 mm; breadth 0.60 mm; thickness 0.30 mm.

Specimens of this species are rare in the Eocene 

collections examined. They seem very close to 

D'Orbigny's species.

Castle Hayne marl: 5611, Wilmington, N.C.; 8165, near 

Maple Cypress, Craven County, N.C.

Jackson formation: 6457, 1% miles south of Shubuta, Miss.; 

6646, Hays Chapel, Wayne County, Miss.

Cooper marl: 11861, steep bluff on west side of Biggiu 

Creek, Berkeley County, S.C.

Genus SIPHONINA Reuss, 1849 

Siphonina jacksonensis Cushman and Applin

Plate 20, figures 2-8

Siphonina jacksoncwsis Cushman and Applin, Am. Assoc 

Petroleum Geologists Bull., vol. 10, p. 180, pi. 9, figs. 

20-23, 1926.

Ellisor, Am. Assoc. Petroleum Geologists Bull., vol. 17, 

no. 11, pi. 3, fig. 21, 1933.

Test much compressed, chambers slightly projecting 

at the posterior angle, at the periphery, keeled, 5 

chambers in. the last-formed coil, fairly distinct; 

sutures very slightly limbate on the dorsal side, spiral 

suture not prominent; wall ornamented by very numerous small spinose processes, in some specimens distinctly developed so that the periphery of the test is 

itself spinose, central portion strongly reticulate; 

aperture elongate, with a slightly'projecting neck and 

lip. Diameter 0.50 mm.

The type specimens of this species came from the 

upper Eocene 4 miles east of Diboll, Angelina County, 

Tex. It has proved to be abundant at the type locality 

of the Jackson formation at Jackson, Miss., and is 

characteristic of the upper Eocene of the general Gulf 

Coastal Plain region. It also occurs in the Alazan 

clays of Mexico and in Trinidad.

The distinctive characters are the very much compressed form and the peculiar spinose and reticulate 

ornamentation of the surface.

Castle Hayne marl: 5611, Wilmington, N.C.

Ocala limestone: 7235, Flint Rivei1 near northeast corner of Dougherty County, Ga.; 7237, half a mile east of 

Huguenen Ferry, Ga.; 6768, Mariauna, Fla.; 6785, threefourths mile south of Herlongs, Fla.; 6789, 6790, Alachua, 

Fla.; 6799, Alachua Sink, Gainesville, Fla.; 6804, Ocala, 

Fla.; 6805, Oakhurst quarry, Ocala, Fla.; 7348, Suwannee 

River iy2 miles above Troy Springs, Fla.; 7360, 3% miles 

south of Floral City, Fla.; 6755, Choctawhatchee River above 

State line, Ala.; 5567, Claiborne, Ala.; 6706, Oven Bluff, 

Tombigbee River, Ala.; 6720, three-fourths mile north of 

Drewry, Ala.; 6722, three-fourths mile northwest of 

Monroeville, Ala; 6725, 6726, 6727, Claiborne, Ala.; 6730, 

Double Branches Creek, Ala.; 6738, 6y2 miles north of Brooklyn, Ala.; 6741, 3^ miles northeast of Brooklyn, Ala.; 6742, 

Powells Landing, Sepulga River, Ala.; 7159, 4 miles southeast of Whatley, Ala.; 1/7, 1 mile east of Beck, Ala.; 1/8, 

Beck, Ala.; 1/33, 7% miles northwest of Geneva, Ala.; 1/39, 

Pea River at Geneva, Ala.
HANTKENINIDAE 49

Jackson formation : 2637, Cocoa, Ala. ; 72U3, 3% miles southeast of Cullomburg, Ala. ; 7220, half a mile southeast of Melvin, Ala. ; 7227, Water Valley, Ala. ; 6457, iy2 miles south of 

Shubuta, Miss.; 6463, Jackson, Miss.; 6471, Garlands Creek, 

Miss.; 6473, Yazoo City, Miss.; 6646, Hays Chapel, Wayne 

County, Miss.

Cooper marl: 11861, steep bluff on west side of Biggin 

Creek, Berkeley County, S.C. ; 11S63, three-fourths mile 

west of Old Biggin Church, Berkeley County, S.C. ; 11862, 1 

mile west of Givhnns, Dorchester County, S.C. ; 11857, 1 mile 

south of Moncks Corner, Berkeley County, S.C.

Siphonina advena Cushman var. eocenica Cushman and 

Applin

Plate 20, figures 9-11

Ki'lihanriui' advGtM Cushman var. coccnica Cuslmiaii anil Applin, Am. Assoc. Petroleum Geologists Bull., vol. 10, p. 

ISO, pi. 9, figs. 16-19, 1926.

Test differing from the typical form of the species, 

which Avas described from the lower Oligocene, in the 

less prominent spiral suture on the dorsal side and the 

more entire periphery, the chambers not showing as 

definitely as in the typical form. Diameter 0.40 mm.

The type specimens of this variety came from 

Tarkiln Creek half a mile above the Neches River, 

Trinity County, Tex. It is very rare in the eastern 

portion of the Coastal Plain.

Cooper marl: 3595, Barnwell County, S.C. 

Family CASSIDULINIDAE 

Genus CASSIDULINA D'Orbigny, 1826 

Cassidulina globosa Hantken

Plate 20, figures 12; i, b

fflobosu Hautken, Magy. kir. foldt. int. iSvkonyve, 

vol. 4, p. 54, pi. 16, fig. 2, 1875 (1S76).

Liebus, Neues Jahrb., 1901, p. 125; Geol. Reichsanstalt 

Jahrb., vol. 56, p. 357, figs. 5a-c (in text), 1906.

Martinotti, Soc. italiana sci. nat. Atti, vol. 62, p. 328, 1923.

Cushman, Contr. Cushman Lab. Foram. Research, vol. 1, 

p. 56, pi. 9, figs. 25, 26, 1925.

Cole, Bull. Am. Paleontology, vol. 14, no. 51, p. 32, 1927.

Cushman, Jour. Paleontology, vol. 1, p. 167, pi. 26, fig. 13,

1927. 

Cole, Bull. Am. Paleontology, vol. 14, no. 63, p. 216 (16),

1928.

Test small, subglobose, periphery broadly rounded, 

about 5 pairs of chambers making up the adult whorl, 

only a very small triangular point of the chambers of 

the reverse side showing at the periphery; sutures 

slightly and evenly curved, distinct but not depressed; 

wall smooth; aperture small, narrow. Length 0.25- 

0.30 mm; breadth 0.20-0.25 mm; thickness 0,12-0.15 

mm.

This species, described by Hantken from Hungary, 

is found in the upper Eocene both of Europe and 

America. It is one of the smallest species of the genus 

but is common in the Cooper marl. There is a considerable amount of variation in the rotundity of the

test.

Cooper marl: 11861, steep bluff on west side of Biggin Creek, 

Berkeley County, S.C.; 11863, three-fourths mile west of Old 

Biggin Church, Berkeley County, S.C.; 11862, 1 mile west of 

Givhans, Dorchester County, S.C.; 11857, 1 mile south of 

Moncks Corner, Berkeley County, S.C.

Family CHILOSTOMELLIDAE

Genus SPHAEROIDINA D'Orbigny, 1826

Sphaeroidina austriaca D'Orbigny

Plate 20, figures 13a, b

Spliaeroidina austriaca D'Orbigny, Foraniinife'res fossiles du 

bassin tertiaire de Vienne, p. 284, pi. 20, figs. 19-21, 1846.

Test globose, on the dorsal side several chambers 

visible, on the ventral side only 3 chambers visible, one 

occupying at least half the ventral surface, periphery 

broadly rounded; sutures distinct, depressed; wall 

smooth; aperture small, at the inner margin of the 

last-formed chamber on the ventral side. Diameter 

0.45-0.50 mm.

Species of Sphaeroidina are rare in the Eocene but 

become increasingly more common in the later Tertiary. Specimens are fairly common in the Cooper 

marl of South Carolina but were not found elsewhere 

in the collections examined. They are of the typical 

form seen in the Miocene of the Vienna Basin and elsewhere in the Tertiary of Europe, with the chambers 

less completely involute than in S. bulloides D'Orbigny, and more chambers visible on the dorsal side.

Cooper marl: 11861, steep bluff on west side of Biggin Creek, 

Berkeley County, S.C.; 11863, three-fourths mile west of Old 

Biggin Church, Berkeley County, S.C.; 11862, 1 mile west of 

Givhaus, Dorchester County, S.C.

Family HANTKENINIDAE

Genus HANTKENINA Cushman, 1924

Hantkenina alabamensis Cushman

Plate 13, figures 1-5

Hantkenina alabamensis Cushmau, U.S. Nat. Mus. Proc., vol.

66, art. 3, p. 3, pi. 1, figs. 1-6, pi. 2, fig. 5, 1924; Contr.

Cushman Lab. Foram. Research, vol. 1, p. 7, pi. 1, fig. 11 ;

p. 68, 1925. 

Cushman and Applin, Am. Assoc. Petroleum Geologists

Bull., vol. 10, p. 177, pi. 10, fig. 3, 1926. 

Howe and Wallace, Louisiana Dept. Conservation Geol.

Bull. 2, p. 54, pi. 10, fig. 3, 1932. 

Ellisor, Am. Assoc. Petroleum Geologists Bull., vol. 17,

no. 11, pi. 6, fig. 5, 1933.

Test planispiral, compressed, adult coil with 5 or 6 

chambers, periphery very slightly if at all lobulate; 

wall very finely punctate, smooth, granular near the 

aperture, each chamber with a hollow, slender, acicular 

spine at the periphery, pointing somewhat anteriorly; 

aperture tripartite, with an elongate projection along
50 UPPER EOCENE FOBAMINIFERA OF THE SOUTHEASTERN" UNITED STATES

each side at the base of the apertural face, and the 

third, median, extending1 peripherally from the base of 

the apertural face. Diameter without spines 0.45 mm; 

with spines 0.75 mm.

I described this species from the sand in the Jackson 

formation at U.S.G.S. station 2637, Cocoa, Ala. It is 

abundant at this station and is characteristic of this 

horizon of the upper Jackson.

Ocala limestone: 6722, three-fourths mile northwest of Monroeville, Ala.; 6746, Ward's sawmill, Sepulga River, Ala.; 7159, 

4 miles southeast of Whatley, Ala.; 7719, Spring Mill Branch, 

Jenkins County, Ga.; 1/38, 3*4 miles north of Grove Hill, Ala.; 

1/43, iy2 miles southwest of Perdue Hill, Ala.

Jackson formation: 2637, Cocoa, Ala.; 7203, 3% miles southeast of Cullomburg, Ala.; 6457, 1% miles south of Shubuta, 

Miss.; 6473, Yazoo City, Miss.; 6646, Hays Chapel, Wayne 

County, Miss.

Hantkenina ailabamensis also occurs in well samples 

in Texas and is known from the Alazan clays of 

Mexico.

Related species, Hantkenina brevispina Cushman, 

H. longispina Cushman, and H. mexicana Cushman, 

occur in the Eocene of Mexico. H. mexicana var. 

aragonensis Nuttall, H. alabamensis var. primitivn 

Cushman and Jarvis, and H. lehneri Cushman and 

Jarvis occur in the Eocene of Trinidad. H. kocM 

(Hantken) is known from Europe.

Family GLOBOROTALIIDAE

Genus GLOBOROTALIA Cushman, 1927

Globorotalia cocoaensis Cushman

Plate 21, figures 1-3

Glo'borotaUa cocoaensis Cushman, Contr. Cushman Lab. Foram.

Research, vol. 4, p. 75, pi. 10, figs. 3a-c, 1928. 

Howe and Wallace, Louisiana Dept. Conservation Geol.

Bull. 2, p. 75, pi. 14, fig. 4, 1932. 

Ellisor, Am. Assoc. Petroleum Geologists Bull., vol. 17,

no. 11, pi. 4, figs. 6a, b, 1933.

Test small, with a low, rounded spire, ventral side 

strongly convex, periphery rounded; chambers very 

distinct, 4 usually making up the last-formed whorl, 

inflated, earlier ones more globular than the later ones; 

sutures very distinct on the dorsal side, oblique and 

curved, on ventral side nearly straight, radiate, deeper, 

depressed, wall in the early chambers rough and almost 

spinose, in the last ones nearly smooth, calcareous, perforate ; aperture about midway between the umbilicus 

and periphery, an elongate, arched opening. Diameter 0.35 mm; height 0.25 mm.

This species is closely related to Crloborotaliacanariensis (D'Orbigny) and G. crassa, (D'Orbigny), but it 

greatly differs from these in the relative shape of the 

test and the form of the individual chambers. It is 

very abundant at the type locality, Cocoa, Ala., but 

occurs at a few other stations.

Cooper marl: 6321, Baldock, S.C.

Ocala limestone: 7191, Marianna, Fla.; 7159, 4 miles southeast of Whatley, Ala.

Jackson formation: 2637, Cocoa, Ala.; 6457, 1% miles south 

of Shubuta, Miss.

Family ANOMALINIDAE

Genus ANOMALINA D'Orbigny, 1826

Anomalina bilateralis Cushman

Plate 21, figures 4, 5

Anomalina bilateralis Cushman, U.S. Geol. Survey Prof. Paper 

129, pp. 97, 137, pi. 21, figs. 1, 2, 1922; Prof. Paper 133, 

p. 42, 1923.

Ellisor, Am. Assoc. Petroleum Geologists Bull., vol. 17, 

no. 11, pi. 4, fig. 7, 1933.

Test of about 4 coils, bilateral or nearly so, composed of numerous chambers, 10 or more in the lastformed whorl, umbilical region on both sides with a 

knob of clear shell material, more pronounced on the 

dorsal side; chambers smooth but coarsely punctate, 

more coarsely so on the ventral side; sutures broad 

and somewhat limbate with clear shell material; aperture a narrow curved opening at the base of the final 

chamber. Maximum diameter 1.00 mm.

This species, which is widely distributed throughout the various members of the Vicksburg group of 

the lower Oligocene, also occurs in the upper Eocene, 

but in much less numbers. The type of this species is 

from the lower Oligocene of the United States.

Ocala limestone: 6706, Oven Bluff, Tombigbee River, Ala.; 

6719, three-fourths mile north of Drewry, Ala.; 6725, Claiborne, Ala.; 7159, 4 miles southeast of Whatley, Ala.; 7161, 5 

miles southeast of Whatley, Ala.; 1/21, Covington County, 

Ala.; 1/26, Drewry, Ala.; 3966, 2 miles south of Perry. Ga.

Jackson formation: 7203, 3% miles southeast of Cullomburg, 

Ala.; 6457, iy2 miles south of Shubuta, Miss.; 64613, Jackson, 

Miss.; 6473, Yazoo City, Miss.; 6646, Hays Chapel, Wayne 

County, Miss.

Anomalina granosa (Hantken) var. dibollensis Cushman and

Applin

Plate 21, figures 6. 7

Anomalina granosa (Hantken) var. dibollensis Cushman and 

Applin, Am. Assoc. Petroleum Geologists Bull., vol. 10, 

p. 179, pi. 9, fig. 15, 1926.

Test nearly bilaterally symmetrical, umbilicate on 

both sides, periphery rounded; chambers about 10 in 

the final coil, of nearly uniform size and shape, increasing slowly in size as added; sutures slightly 

curved, depressed; wall smooth but distinctly perforate; aperture at the base of the periphery of the 

chamber. Diameter 0.40 mm; thickness 0.10 mm.

The type specimens of this variety were collected 

4 miles east of Diboll, Angelina County, Tex.

Jackson formation: 6457, 1% miles south of Shubuta, Miss.
ANOMALINIDAE 51

Anomalina affinis (Hantken)

Plate 21, figures 11, 12

Pulviniulina afllnis Hantken, K. ungar. geol. Anstalt Mitt.

Jahrb., vol. 4, p. 78, pi. 10, figs. 6a, b, 1875. 

Anomalina afllnis Cusliman and Applin, Am. Assoc. Petroleum

Geologists Bull., vol. 10, p. 180, pi. 10, figs. 1, 2, 1926.

Test nearly planispiral in the adult, the dorsal side 

slightly evolute, periphery in the earlier portion subacute, later rounded; chambers about 12 in the final 

coil, of nearly uniform size and shape, increasing 

slightly in size as added; sutures curved, slightly depressed; wall smooth, finely but distinctly perforate; 

aperture peripheral at the base of the chamber. Maximum diameter 0.90 mm; thickness 0.25 mm.

The specimens came from the Jackson Eocene 4 

miles east of Diboll, Angelina County, Tex.

The chambers are more numerous and more elongate than those of A. granosa var. dibollensis,

Anomalina cocoaensis Cushman

Plate 21, figures 13a-c

Anomalina cocoaensis Cushmau, Contr. Cushman Lab. Foram. 

Research, vol. 4, p. 75, pi. 10, figs. 4a-c, 1928.

Test planoconvex, ventral side broadly convex, 

dorsal side flattened or even somewhat concave; the 

ventral side with a series of irregular-shaped bosses 

in a cluster in the umbilical region; the dorsal side 

with a low spire appearing almost as a low smooth 

boss, the adjacent area depressed and then rising to 

the raised, rounded border, periphery broadly 

rounded; chambers 10 to 12 in the last-formed coil, 

fairly distinct; sutures on the dorsal side limbate 

and slightly raised, oblique and curved, sutures on the 

ventral side limbate, not raised or depressed, broadest 

near the inner end, nearly radial; wall calcareous, 

coarsely perforate, especially on the ventral side; 

aperture peripheral, small, arched, with a slight lip. 

Diameter 0.50 mm; height 0.20 mm.

The types of this species are from the Jackson formation at Cocoa, Ala., where it is abundant. It occurs in typical form at other stations, and the characters are remarkably constant.

Ocala limestone: 1/38, 3% miles north of Grove Hill, Ala.

Jackson formation: 2637, Cocoa, Ala.; 7203, 3% miles southeast of Cullomburg, Ala.; 6457, 1% miles south of Shubuta, 

Miss.

Anomalina jacksonensis (Cushman and Applin)

Plate 21, figures 8a-c

Discorbis jacksonensis Cushman and Applin, Am. Assoc. Petroleum Geologists Bull., vol. 10, p. 178, pi. 9, figs. 8, 9, 

1926.

Anomalina jacksonensis Ellisor, Am. Assoc. Petroleum Geologists Bull., vol. 17, no. 11, pi. 4, figs. lOa, b, 1933.

Test small, planoconvex, the ventral side flattened, 

dorsal side only slightly convex, with a distinct, clear

umbonal mass, occupying the central region, periphery subacute; chambers numerous, 10 or more in the 

last-formed coil, elongate, distinct; sutures distinct, 

very slightly depressed and somewhat limbate, on the 

dorsal side nearly straight until toward the outer half, 

where they become strongly recurved; on the ventral 

side sutures much depressed, especially toward the 

umbilicus, which becomes somewhat open in welldeveloped specimens; wall smooth, finely punctate. 

Length 0.50 mm; breadth 0.40 mm.

The type specimens of this species came from the 

Jackson of Bridge Creek, iy2 miles above the Angelina River, San Augustine County, Tex. It occurs 

in numerous well sections of Texas.

Anomalma jacksonensis is represented also by several varieties.

Cooper marl: 3595, Barnwell County, S.C.

Ocala limestone: 6720, three-fourths mile north of Drewry, 

Ala.; 6722, three-fourths mile northwest of Monroeville, Ala.; 

6724, 6726, Claiborne, Ala.; 6738, 6% miles north of Brooklyn, 

Ala.; 6741, 3% miles northeast of Brooklyn, Ala.; 6754, Choctawhatchee River at State line, Ala.-Fla.; 7159, 4 miles southeast of Whatley, Ala.; 1/8, Beck, Ala.

Jackson formation: 7227, Water Valley, Ala.; 6457, 1% miles 

south of Shubuta, Miss.

Anomalina jacksonensis (Cushman and Applin) var. dibollensis (Cushman and Applin)

Plate 21, figure 9

DiscorMs jacksonensis Cushman and Applin var. dibollensis 

Cushman and Applin, Am. Assoc. Petroleum Geologists 

Bull., vol. 10, p. 178, pi. 9, fig. 10, 1926.

Variety differing from the typical form in being 

slightly more rounded and having fewer chambers; 

chambers shorter and wider; sutures more evenly 

curved and on the dorsal side slightly less developed, 

an umbonate thickening over the central region; wall 

finely punctate, smooth. Maximum length 0.50 mm; 

breadth 0.35 mm.

Type specimens from the upper Eocene 4 miles east 

of Diboll, Angelina County, Tex.

Cooper marl: 5531, Charleston, S.C.; 6321, Baldock, S.C.

Ocala limestone: 5567, Claiborne, Ala.; 6730, Double 

Branches Creek, Ala.; 6742, Powells Landing, Sepulga River, 

Ala.; 7161, 5 miles southeast of Whatley, Ala.; 1/39, Pea 

River at Geneva, Ala.

Jackson formation: 6463, Jackson, Miss.; 6471, Garlands 

Creek, Miss.

Anomalina jacksonensis (Cushman and Applin) var. texana 

(Cushman and Applin)

Plate 21, figure 10

Discorbina jacksonensis Cushman and Applin var. teaiana

Cushman and Applin, Am. Assoc. Petroleum Geologists

Bull., vol. 10, p. 178, pi. 9, fig. 11, 1926. 

Howe and Wallace, Louisiana Dept. Conservation Geol.

Bull. 2, p. 77, pi. 14, figs. 3a-c, 1932. 

Ellisor, Am. Assoc. Petroleum Geologists Bull., vol. 17,

no. 11, pi. 4, fig. 8, 1933.
52 UPPER EOCENE FORAMINIFERA OF THE SOUTHEASTERN UNITED STATES

Variety differing from the typical form in the 

slightly larger size and the more open coil, resulting 

in shorter, broader chambers. Length 0.60 mm; 

breadth 0.40 mm.

The type specimens came from the Jackson of Ohio 

and Red River well 2, Tyler County, Tex., at a depth 

of 400 feet. The variety has not been noted except at 

this locality.

Genus PLANULINA D'Orbigny, 1826 

Planulina cocoaensis Cushman

Plate 22, figures 7a-c

Planulina cocoaensis Cushman, Contr. Cushman Lab. Foram. 

Research, vol. 4, p. 76, pi. 10, figs, la-c, 1928.

Test much compressed, periphery acute and slightly 

keeled, central region of the ventral side occupied by 

a distinct, smooth, rounded boss, on the dorsal side 

the very low spire with numerous small rounded protuberances; chambers usually 10 to 12 in the lastformed whorl, distinct, the last-formed ones slightly 

inflated, especially on the ventral side; sutures very 

distinct, somewhat limbate on both sides, very slightly 

if at all depressed, curved on both sides; wall calcareous, coarsely perforate; aperture in the adult 

extending over onto the dorsal side of the test, elongate, narrow. Length 0.50 mm; breadth 0.40 mm; 

thickness 0.15 mm.

This species is very abundant at Cocoa, Ala., the 

type locality. It is a very finely marked species and 

is usually very well preserved.

Ocala limestone: 1/38, 3% miles north of Grove Hill, Ala.; 

7159, 4 miles southeast of Whatley, Ala.

Jackson formation: 2637, Cocoa, Ala.; 7203, 3% miles southeast of Cullomburg, Ala.; 7220, half a mile southeast of Melvin, Ala.; 6457, 1% miles south of Shubuta, Miss.

Planulina cocoaensis Cushman var. cooperensis Cushman

Plate 22, figures Sa-c

Planulina cocoaensis Cushman var. cooperensis Cushman, 

Contr. Cushman Lab. Foram. Research, vol. 9, p. 20, 

pi. 2, figs. 12a-c, 1933.

Variety differing from the typical form in the 

somewhat larger size, smaller number of chambers, 

typically .8, and a much smoother, less ornamented 

surface. Maximum diameter 1.50 mm.

Holotype (no. 371565, U.S.N.M.) from Cooper 

marl of station 11861, Berkeley County, S.C.

This variety so far as known seems to be characteristic of the Cooper marl and has not been found 

elsewhere in the material examined.

Cooper marl: 11861, steep bluff on west side of Biggin 

Creek, Berkeley County, S.C.; 11863, three-fourths mile west 

of Old Biggin Church, Berkeley County, S.C.; 11862, 1 mile 

west of Givhans, Dorchester County, S.C.; 11857, 1 mile south 

of Moncks Corner, Berkeley County, S.C.

Genus CIBICIDES Montfort, 1808 

Cibicides lobatulus (Walker and Jacob)

Plate 22, figures 4-6

Nautilus lobatula Walker and Jacob, Adam's Essays on the

. microscope, Kanmacher's ed., p. 642, pi. 14, fig. 36, 1798.

Cil)icides lobatulus Cushman, Jour. Paleontology, vol. 1, p.

170, pi. 27, figs. 12, 13, 1927.

Hanna and Church, Jour. Paleontology, vol. 1, p. 201, 1928. 

Truncatttlvna lobatula (Walker and Jacob) D'Orbigny, in 

Barker-Webb and Berthelot, Histoire naturelle des lies 

Canaries, vol. 2, pt. 2, Foraminiferes, p. 134, pi. 2, figs. 

22-24, 1839; Foraminiferes fossiles clu bassin tertiaire. 

de Vienne, p. 168, pi. 9, figs. 18-23, 1846. 

H. B. Brady, Challenger Rept, Zoology, vol. 9, p. 660, pi.

92, fig. 10; pi. 93, fig. 1, 1884.

Cushman, U.S. Geol. Survey Bull. 676, p. 16, pi. 1, fig. 10; 

p. 60, pi. 17, figs. 1-3, 1918; Carnegie Inst. Washington 

Pub. 291, p. 41, 1919; U.S. Geol. Survey Prof. Paper 

129, pp. 96, 135, pi. 20, figs. 1-3, 1923; Prof. Paper 133, 

p. 40, 1923.

Test planoconvex, flattened on the ventral face, 

moderately convex dorsally or nearly flat, sometimes 

slightly concave, peripheral margin rounded; chambers numerous, 7 or 8 in the last-formed whorl; sutures depressed, especially on the dorsal face; wall 

smooth, conspicuously punctate.

The specimens referred to this species were apparently attached in their growth, as is the species today. 

The dorsal face in some specimens is concave, apparently to conform to the surface of attachment. For 

this reason the dorsal face varies considerably in general shape. The species is common in the various 

members of the earlier Tertiary.

This is one of the species to which many forms have 

been referred, but it is questionable whether the records from older formations than the Eocene should be 

included under this name.

Cooper marl: 6319, Ingleside, S.C.; 11863, three-fourths mile 

west of Old Biggin Church, Berkeley County, S.C.; 11857, 

1 mile south of Moncks Corner, Berkeley County, S.C.; 6321, 

Baldock, S.C.

Castle Hayne marl: 8171, Richlands, N.C.

Ocala limestone: 3966, 2 miles south of Perry, Ga.; 7097, 

Baiubridge, Ga.; 7689, 1 mile south of Tivola, Ga.; 77.19, 

Spring Mill Branch, Jenkins County, Ga.; 6722, three-fourths 

mile northwest of Monroeville, Ala.; 1/7, 1 mile east of Beck, 

Ala.; 1/21, Covington County, Ala.; 1/26, Drewry, Ala.; 1/39, 

Pea River at Geneva, Ala.

Barnwell formation: 7755, Stevens Pottery, Ga.

Jackson formation: 7227, Water Valley, Ala.

Cibicides pseudoungerianus (Cushman)

Plate 23, figures la-c

Truncatulina ungeriana H. B. Brady, Challenger Rept., Zoology, vol. 9, pi. 94, figs. 9a-c, 1884 [not Rotti.lina. iingeriaiia 

D'Orbigny]. 

Cushman, U.S. Nat. Mus. Bull. 103, p. 09, pi. 24, fig. 1, 1918.

Truncatulina, pseudowngeriana Cushman, U.S. Geol. Survey 

Prof. Paper 129, pp. 97, 136. pi. 20, fig. 9, 1922; Prof. 

Paper 133, p. 40, 1923.
Cibiciilcs pseudounycrianus Cole and Gillespie, Bull. Am. Paleontology, vol. 15, no. 57b, p. 15, pi. 3, figs. 10, 11, 1930. 

Mllisor, Am. Assoc. Petroleum Geologists Bull., vol. 17, 

no. 11, pi. 5, figs. 3, 4, 1933.

Test almost equally biconvex, periphery subacute; 

chambers 9 to 11 in the last-formed whorl, those of 

the earlier whorls not visible on either the ventral 

or the dorsal side, being hidden on the dorsal side by 

the roughness of the surface and on the ventral side 

by the involute character of the chambers; periphery 

lobulate; sutures distinct in the last whorl on the 

dorsal side, on the ventral even more so; umbilical 

region filled by clear shell material nearly flush with 

the chambers, last few chambers on the dorsal side 

slightly above the surface on the inner margin; surface on the dorsal side coarsely punctate, below smooth 

and finely perforate; aperture peripheral. Diameter 

1.00 mm or less.

This species occurs in the upper members of the 

Tertiary and seems to show relatively little variation.

Castle Hayne marl: 5611, W.ilmington, N.C.

Cooper marl: 5531, Charleston, S.C.; 6319, Ingleside, S.C.; 

11863, three-fourths mile west of Old Biggin Church, Berkeley County, S.C.; 11862, 1 mile west of Givhans, Dorchester 

County, S.C.; 11857, 1 mile south of Moncks Corner, Berkeley 

County, S.O.

Ocala limestone: 7237, half a mile east of Huguenen Ferry, 

Ga.; 6706, Oven Bluff, Tombigbee River, Ala.; 7191, Marianna, 

Fla.; 1/21, Covington County, Ala.; 1/26, Drewry, Ala.; 1/39, 

Pea River at Geneva, Ala.; 6725, Claiborne, Ala.; 6726, Claiborne, Ala.; 6727, Claiborne, Ala.

Jackson formation: 6463, Jackson, Miss.; 6471, Garlands 

Creek, Miss.

Cibicides americanus (Cushman) var. antiquus (Cushman 

and Applin)

Plate 22; figures 1, 2

Truncatuli'iM amcricima Cushman var. antiqua Cushman and 

Applin, Am. Assoc. Petroleum Geologists Bull., vol. 10, 

p. 179, pi. 9, figs. 12, 13, 1926.

Variety differing from the typical form in the more 

Limbate, much more curved sutures and fewer chambers. Length 0.40 mm; breadth 0.28 mm.

Type specimens came from the upper Eocene 4 miles 

east of Diboll, Angelina County, Tex.

This variety possibly should have specific standing 

and is closely related to Cibicides yazooensis Cushman.

Cooper marl: 11863, three-fourths mile west of Old Biggin 

Church, Berkeley County, S.C.; 11S62, 1 mile west of Givhans, 

Dorchester County, S.C.; 11857, 1 mile south of Moncks 

Corner, Berkeley County, S.C.

Ocala limestone: 5567, Claiborne, Ala.; 6741, 3% miles 

northeast of Brooklyn, Ala.; 6738, 6% miles north of Brooklyn, 

Ala.

Jackson formation : 7203, 3% miles southeast of Cullomburg, 

Ala.; 7220, half a mile southeast of Melvin, Ala.; 7227, Water 

Valley, Ala.; 6457, 1% miles south of Shubuta, Miss.; 6463, 

Jackson, Miss.; 6471, Garlands Creek, Miss.

53

Cibicides yazooensis Cushman

Plate 23, figures 2a-c

Cibicides yasooensi'S Cusliman, Contr. Cushman Lab. Fornui.

Research, vol. 7, p. 59, pi. 7, figs. 12a-c, 1931. 

Bllisor, Am. Assoc. Petroleum Geologists Bull., vol. 17, 

no. 11, pi. 5, figs. 2, 5, 1933.

Test slightly longer than broad, compressed, periphery angled, slightly lobulate, ventral side convex, 

dorsal side flattened or slightly convex, nearly involute 

on both sides; chambers distinct, usually 8 in the lastformed coil; sutures distinct, curved, very strongly 

limbate and somewhat raised, increasing in thickness 

toward the inner end; wall distinctly but finely perforate, with a clouding of the surface except at the 

sutures, which are clear and transparent; aperture a 

curved, somewhat arched opening near the periphery 

on the ventral side. Length 0.60 mm; breadth 0.50 

mm; thickness 0.25 mm.

Holotype (no. 371566, U.S.N.M.) from Jackson formation, station 6473, Yazoo City, Miss.

This species is related to Cibicides amemcaniivs 

(Cushman) var. witiqitus (Cushman and Applin), but 

the sutures are much more limbate, the chambers are 

broader, and the whole test is heavier and coarser.

Ocala limestoue: 1/38, 3% miles north of Grove Hill, Ala. 

Jackson formation : 6472, 6473, Yazoo City, Miss.

Cibicides cooperensis Cushman

Plate 23, figures 3a-c

Gilicklcs cooperensis Cushman, Contr. Cushman Lab. Forain. 

Research, vol. 9, p. 20, pi. 2, figs, lla-c, 1933.

Test coiled, nearly planispiral in the adult, unequally biconvex, dorsal side slightly convex, deeply 

umbilicate, both sides almost completely involute, periphery rounded, more broadly so in the later portion, 

very slightly lobulate; chambers distinct, 7 or 8 in 

the final whorl, increasing rapidly in size as added, 

on the dorsal side with a slight lip along the umbilical end; sutures distinct, somewhat limbate, later 

ones slightly depressed, earlier ones flush with the 

surface, gently curved; wall smooth, finely but distinctly perforate; aperture extending from the periphery over onto the dorsal side, somewhat covered 

by the liplike projection from the umbilical end of 

the chamber. Length 0.80 mm; breadth 0.65 mm; 

thickness 0.25-0.30 mm.

Holotype (no. 371568, U.S.N.M.) from Cooper 

marl, station 11857, Berkeley County, S.C. This is. a 

very abundant and distinctive species in the Cooper 

marl.

Cooper marl: 11861, steep bluff; on west side of! Biggin 

Creek, Berkeley County, S.C.; 11863, three-fourths mile west 

of Old Biggin Church, Berkeley County, S.C.; 11862, 1 mile 

west of Givhans, Dorchester County, S.C.; 11857, 1 mile south 

of Moncks Corner, Berkeley County, S.C.
54 UPPER EOCENE FORAMINIFERA OP THE SOUTHEASTERN" UNITED STATES

Cibicides mississippiensis (Cushman)

Plate 22, figures 3a-c

Anomalina mississippiensis Cushman, U.S. Geol. Survey Prof. 

Paper 129, pp. 98, 137, pi. 21, figs. 6-8, 1922; Prof. Paper 

133, p. 43, 1923.

Cibicides mississippiensis Ellisor, Am. Assoc. Petroleum Geologists Bull., vol. 17, no. 11, pi. 5, figs. 6, 7, 1933.

Test small, planoconvex, of about 2i/£ coils, periphery slightly lobulate, bluntly rounded, dorsal side very 

much flattened, even slightly concave, ventral side 

very convex; chambers comparatively few, 6 to 8 in 

the last-formed coil; sutures curved, on the dorsal 

side broad and limbate, even with the surface of clear 

shell material, on the ventral side narrower and depressed ; the last-formed 2 or 3 chambers on the inner 

margin on the dorsal side slightly above the general 

surface; wall thin and translucent, especially on the 

dorsal side, smooth, on the ventral side finely punctate and not so clear; aperture a curved opening at 

the inner margin at the periphery, extending to the 

dorsal side. Length 0.25-0.35 mm; breadth 0.20- 

0.30 mm.

This small but very definite species occurs in all the 

divisions of the lower Oligocene of the eastern Gulf 

Coastal Plain. In the upper Eocene of the same 

region it is not abundant as to numbers but is widely 

distributed. At least one related species may be distinguished which is more abundant than this.

Castle Hayne marl: 5611, Wilmington, N.C.; 8165, near 

Maple Cypress, Craven County, N.C.

Cooper marl: 6321, Baldock, S.C.

Ocala limestone: 7097, Bainbridge, Ga.; 7108, Flint River 

at Seaboard Air Line Ry. bridge, Sumter County, Ga.; 7237, 

half a mile east of Huguenen Ferry, Ga.; 6768, Marianna, 

Fla.; 6785, three-fourths mile south of Herlongs, Fla.; 6790, 

Alachua, Fla.; 7191, Marianna, Fla.; 6720, three-fourths mile 

north of Drewry, Ala.; 6722, three-fourths mile northwest of 

Monroeville, Ala.; 6738, 6% miles north of Brooklyn, Ala.; 

6741, 3% miles northeast of Brooklyn, Ala.; 7159, 4 miles 

southeast of Whatley, Ala.

Jackson formation: 2637, Cocoa, Ala.; 6463, Jackson, Miss.; 

6457, 1% miles south of Shubuta, Miss.

Cibicides mississippiensis (Cushman) var. ocalanus 

Cushman, n. var.

Test planoconvex, of about 2^ coils, periphery entire, sharply angled, dorsal side very much flattened, 

even slightly concave, ventral side very convex, especially the central portion, near the periphery thinning 

rapidly; chambers 10 to 12 in the last-formed coil; 

sutures strongly curved, on the dorsal side somewhat 

limbate, not depressed, on the ventral side narrow and 

depressed, the last-formed chambers on the inner margin on the dorsal side very slightly above the general 

surface; wall smooth; aperture a curved opening at 

the inner margin at the periphery. Maximum diameter 0.40 mm.

Holotype (no. 371567, TJ.S.N.M.) from Ocala limestone, station 6815, Alachua County, Fla.

This variety is close to Cibicides mississippiensis 

(Cushman) -but may be distinguished by the larger 

number of chambers, the broader form, and the peculiar reversed curve of the ventral side, concave near 

the periphery, then strongly convex.

The variety ocalanus seems to be possibly the ancestral form of G. mississippiensis, occurring in the upper Eocene in much greater numbers, and replaced by 

G. mississippiensis in the lower Oligocene,

Ocala limestone: 3966, 2 miles south of Perry, Ga.; 6768, 

Marianna, Fla.; 6785, three-fourths mile south of Herlongs, 

Fla.; 6789, 6790, Alachua, Fla.; 6799, Alachua Sink, Gainesville, Fla.; 6804, Ocala, Fla.; 6805, Oakhurst quarry, Ocala, 

Fla.; 6807, Zuber, Fla.; 6808, Martin, Fla.; 6812, 1% miles 

south of Newberry, Fa.; 6815, Clark, Fla.; 7191, Marianna, 

Fla.; 7345, Suwannee River 2% miles above Branford, Fla.; 

7366, Anthony, Fla.; 7348, Suwannee River 1% miles above 

Troy Springs, Fla.; 7360, 3% miles south of Floral City, Fla.; 

6754, Choctawhatehee River at State line, Ala.-Fla.; 6755, 

Choctawhatchee River above State line, Ala.; 6724, Claiborne, 

Ala.; 1/8, Beck, Ala.; 1/39, Pea River at Geneva, Ala.

Cooper marl: 3595, Barnwell County, S.C.

Family PIANORBULINIDAE

Genus GYPSINA Carter, 1877

Gypsina globula (Reuss)

Plate 23, figures 4, 5

Ceriopora, glofiukis Reuss, Haidinger's Naturwiss. Abh., vol. 2, 

p. 23, pi. 5, fig. 7, 1847.

Test globular, usually spherical, consisting of numerous chambers in irregular concentric layers, surface 

pustulose, as all the chambers are not added at the 

same time, the last-formed chambers protruding beyond earlier-formed ones; wall coarsely perforate, 

between adjacent chambers somewhat thickened but 

not raised. Maximum diameter 2.00 mm.

This species was described by Reuss from the middle 

Tertiary of central Europe. It has been recorded 

from numerous Tertiary formations and from Recent 

seas. The Recent species is, however, distinct from 

this early Tertiary one in the details of its structure. 

The very large form from the Bowden marl of Jamaica has already been distinguished by Brady as var. 

pilatris.

From the material examined the species seems to be 

most common in the Ocala limestone phase of the 

upper Eocene. The Recent species are usually common in shallow water of the Tropics.

Ocala limestone: 7097, Bainbridge, Ga.; 6790, Alachua, 

Fla.; 7338, Bowling Springs, Fla.; 7360, 3% miles south of 

Floral City, Fla.; 7364, 1^ miles west of Groom, Fla.; 7365, 

Pineola, Fla.; 7366, Anthony, Fla.; 7161, 5 miles southeast of 

Whatley, Ala.

Jackson formation: 6463, Jackson, Miss.; 6471, Garlands 

Creek, Miss.
BUPEBTHDAE 55

Family RUPERTIIDAE

Genus KTTPERTIA Wallich, 1877

Rupertia floridana Cushman

Plate 23, figures 6, 7

Rupertiat floridana Cushman, Contr. Cushman Lab. Foram. 

Research, vol. 9, p. 21, pi. 2, figs. 13, 14, 1933.

Test attached, elongate, chambers in a loose spiral in 

a column, gradually increasing in diameter toward the 

upper, apertural end; chambers fairly distinct, slightly 

inflated; sutures fairly distinct, slightly depressed; 

wall smooth, calcareous,.perforate; aperture multiple, 

in a slight depression of the outer end of the test. 

Height 2.00 mm; diameter 1.00 mm.

Holotype (no. 371569, U.S.N.M.) from Ocala limestone of Alachua, Fla.

This has many of the characters of Rupertia, but 

further specimens are needed so that sections may be 

made for comparison with some of the other Eocene 

forms of somewhat similar structure.

Ocala limestone: 6790, Alachua, Fla.

Genus CARPENTERIA Gray, 1859

Carpenteria monticularis Carter

Plate 23, figure 9

Carp&nteria monticularis Carter, Annals and Mag. Nat. Hist.,

ser. 4, vol. 19, p. 211, pi. 13, figs. 9-12, 1877. 

H. B. Brady, Cliallenger Kept., Zoology, vol. 9, p. 677, pi.

99, figs. 1-5, 1884. 

Hgger, K. bayer. Akad. Wiss. Miinchen, Cl. 2, Abh., vol. 18,

p. 439, pi. 21, fig. 12, 1893. 

Chapman, Linnean Soc. London Jour:, Zoology, vol. 28, p.

14, pi. 2, fig. 5; pi. 4, figs. 5, 6, 1899. 

Millett, Roy. Micr. Soc. Jour., 1904, p. 496. 

Heron-Alien and Earland, Zool. Soc. London Trans., vol.

20, p. 713, 1915. 

Cushman, U.S. Nat. Mus. Bull. 71, pt. 5, p. 48, pi. 20, figs.

3a-c, 1915; Bull. 100, vol. 4, p. 362, 1921.

Test attached, in the early stages resembling Cibioides, usually with about 4 chambers in a coil, in the

adult spreading, the chambers compressed but rising 

toward the center and ending in a necldike protuberance, at the open end of which is the aperture; wall 

smooth, very finely perforate. Diameter of Eocene 

specimens 3.00-4.00 mm.

The records for this species are almost entirely from 

Recent material of the general Indo-Pacific region. 

Brady recorded it from off the Falkland Islands and 

off Bermuda, although these records may possibly be 

based on the young of another species.

It is surprising to find this species in the upper 

Eocene of America, but several specimens confirm its 

presence. It is attached to valves of Pecten and to 

Lepidocyclina ocalana Cushman. There is little to 

distinguish it from the Recent Indo-Pacific species.

Castle Hayne marl; 5611, Wilmington, N.C.

Carpenteria carolinensis Cushman, n. sp.

Plate 23, figure 8

Test attached, somewhat spreading, but the whole 

much more compact and higher than in G. monticularis; chambers somewhat fused, the sutures indistinct, aperture central; wall smooth and somewhat 

polished, with a few large, deep, circular, porelike 

depressions. Diameter 2.00 mm.

Holotype (no. 371570, U.S.N.M.) from Castle 

Hayne marl, station 5611, Wilmington, N.C.

This species differs from the other species of the 

same locality in the punctations of the surface and the 

more compact higher form. It is unlike any described species. It was attached to a species of 

brachiopod.

Castle Hayne marl: 5611, Wilmington, N.C.

Fragments of other attached forms make it apparent that under the warm, shallow-water conditions, 

of the Ocala especially, there were probably others of 

the large attached Foraminifera present. A careful 

search of the surface of Mollusca and Lepidocyclina 

will undoubtedly reveal additional species.
56 UPPER EOCENE FORAMINIFERA OF THE SOUTHEASTERN UNITED STATES 

Distribution of species of Foraminifera in Texas

Haplophragmoides dibollensis 

Cushman and Applin. 

Ammobaculites hockleyensis 

Cushman and Applin. 

Textularia mississippiensis Cushman and Applin. 

dibollensis Cushman and 

Applin. 

dibollensis Cushman and 

Applin v.ar. humblei 

Cushman and Applin____ 

distortio Cushman and Applin. 

hockleyensis Cushman and 

Applin. 

Quinqueloculina sp.? Cushmau 

and Applin. 

sp.? Cushman and Applin .

sp.? Cushman and Applin. _ Massilina sp.? Cushman and ApRobulus alato-limbatus (Giimbel). 

articulatus (Reuss) var. texanus (Cushman and Applin) . 

limbosus (Reuss) var. hockleyensis (Cushman and 

Applin) . 

propinquus (Hantken) _ _ _ _

Marginulina jacksonensis Cushman and Applin. 

fragaria (Giimbel) var. texasensis (Cushman and 

Applin.) 

Dentalina cocoaensis (Cushman. 

jacksonensis (Cushman and 

Applin) . 

Guttulina problema D'Orbigny -

spicaef ormis (Roemer) _____

Globulina inaequalis Reuss___._

Pseudomorphina dumblei (Cushman and Applin). 

decora (Reuss) --_-________

Glandulina laevigata D'Orbigny 

var. ovata Cushman and Applin. 

Nonion advenum (Cushman) _ _

inexcavatum (Cushman and 

Applin) . 

whitsettense (Cushman and 

Applin) . 

Nonionella hantkeni (Cushman 

and Applin). 

Elphidium texanum (Cushman 

and Applin). 

Hautkenina alabamensis Cushman. 

Bulimina jacksonensis Cushman. 

ovata D'Orbigny-- __ ___.

Virgulina dibollensis Cushman 

and Applin. 

Bolivina gracilis Cushman and 

Applin. 

jacksonensis Cushman and 

Applin. 

jacksonensis Gush man and 

Applin var. striutella 

Cushman.

4787. 

Below Robinson.'s 

Ferry, 

Sabine 

County

X 

X

Bridge 

Creek, 

San Augustine 

County

X

X

X

Stovall 

Creek, 

4 miles 

east of 

Diboll, 

Angelina 

County

X

X

X 

X

X

X

X

X

X

Tarkiln 

Creek, 

Trinity 

County

X

X

Ohio 

and Red 

River 

well 2, 

Tyler 

County

X 

X

X

X 

X

X 

X

X

X 

X 

X

X

Warren 

well 4, 

Harris 

County

X

Near Cheapside, 

Gonzales 

County

X

6148. Old 

Whitsett 

ranch, 

Atascosa 

Creek, 

Atascosa 

County

X

North o Whitsett, 

Live Oak 

County

Other 

upper 

Eocene 

localities

X 

X 

X

X

X

X 

X

Known stratigraphic range

Earliest

Eocene____ 

_-.do------

-_-do-.-___

-_.do_____.

-_-do_.__._

-_-do______

__-do______

___do._ _ _

..-do- ._

-..do- _.

-.do -.,

 do 

__-do._____

_--do--___

-_-do______

_--do______

---do_--.__

 do -_

___do______

Cretaceous. 

Eocene____ 

 do _

 do _

 do _

 _do-__--.

-_-do-_-_.

 do _

 do _

_-.do-- .

--_do---_.

--_do---_.

---do--...

Cretaceous. 

Eocene.___

_-_do-_ .

 do__ _

_--do._ ..__

Latest

Eocene. 

Do. 

Miocene. 

Eocene. 

Do.

Do. 

Do. 

Do.

Do. 

Do. 

Do.

Do. 

Do.

Do.

Oligocene. 

Eocene.

Do.

Do. 

Do.

Recent. 

Do. 

Do. 

Miocene.

Do.

Eocene.

Oligocene. 

Eocene.

Do. 

Oligocene. 

Eocene. 

Do.

Do. 

Recent. 

Eocene.

Do. 

Do. 

Do.
UPPER EOCENE FORAMINIFERA OP THE SOUTHEASTERN UNITED STATES 

Distribution of species of Foraminifera in Texas Continued

57

Uvigerina alata Cushman and 

Applin.

dumblei Cushman and Applin. 

gardnerae Cushman and Applin. 

gardnerae Cushman and 

Applin var. texana Cushman and Applin.

Pleurostomella jacksonensis 

Cushman and Applin. 

Eponides jacksonensis Cushman 

and Applin.

Siphonina jacksonensis Cushman and Applin. 

advena Cushman var. eocenica Cushman and Applin. 

Anonmlina granosa (Hantken) 

var. dibollensis Cushman 

and Applin.

jacksonensis (Cushman and 

Applin. 

jacksonensis Cushman and 

Applin var. dibollensis 

(Cushman and Applin). 

jacksonensis (Cushman and 

Applin) var. texana (Cushman and Applin). 

Cibicides americanus (Cushman) 

var. antiquus (Cushman and 

Applin).

4787. 

Below Robinson's 

Ferry, 

Sabine 

County

Bridge 

Creek, 

San A ugustine 

County

X

S to vail 

Creek, 

4 miles 

east of 

Diboll, 

Angelina 

County

X

X

X 

X

X

X

Tarkiln 

Creek, 

Trinity 

County

X

X

Ohio 

and Red 

River 

well 2, 

Tyler 

County

X 

X

X 

X

X

X

Warren 

well 4, 

Harris 

County

Near Cheapside, 

Oonzales 

County

6148. Old 

Whitsett 

ranch, 

Atascosa 

Creek, 

A tasocsa 

County

Northof Whitsett, 

Live Oak 

County

,

X

Other 

upper 

Eocene 

localities

X

X

Known stratigraphic range

Earliest,

Eocene___. 

---do-._---

___do------

...do......

--Ido..-.--

-,-do------

_-_do_-.---

_-_do,_--_.

___do_-_-i_

_-_do-_._--

__.do_.....

___do_.----

_-_do_.----

__-do--.---

_--do.--.--

__.do_._...

..-do..--..

Latest

Eocene.

Do. 

Do.

Do. 

Do.

Do. 

Do.

Do.

Recent. 

Eocene.

Do. 

Do.

Oligocene. 

Eocene.

Do. 

Do. 

Do."

Distribution of Eocene Foraminifera in North Carolina

Lagena orbignyana (Seguenza) var. semiconcentrica Cushman. _______

Uvigerina cookei Cushman. n. sp_ _- .. .._-_ . ._--._ ._-__ __

Gyroidina soldanii D'Orbigny var. octocamerata Cushman and G. D. 

Hanna.

budensis (Hantken) var. planata Cushman, n. var_... ...

Cancris brongniarti (D'Orbigny). ..-- . .-_ _______ ...- ------

Siphonina jacksonensis Cushman and Applin _ ___-_ _________

Cibicides lobatulus (Walker and Jacob) D'Orbignv. .._.._..___. _

Carpentaria nionticularis Carter __ .__.__- _. __-_ .. _ __.

carolinensis Cushman. ________ ._ _ . __ ._._ ________ __

8105

X

X

8171

X

X

5611

X

xX 

. X 

X 

X 

X 

X 

X 

X

X 

X 

X 

X

X 

X 

X 

X

Known stratigraphic range

Earliest

...do...---.. -------

.-. do. _----.--------

-..do...-.----....-.

..-do... ....... -----

...do... .........--_

-.-do-.---.... ------

__-do------------.--

-_-do-----.---------

...do-.-. -.....----.

-.-do---.---....---.

... do........ .......

. -.do---... ---------

...do--....-..-.-..

...do----..........

...do-....----......

...do-------.-------

Latest

Eocene. 

Do. 

Oligocene. 

Eocene. 

Recent. 

Do. 

Do. 

Eocene. 

Do. 

Oligocene.

Eocene. 

Do. 

Miocene. 

Eocene. 

Recent. 

Do. 

Oligocene. 

Recent. 

Eocene.
58 UPPER EOCENE FORAMINIFERA OP THE SOUTHEASTERN UNITED STATES 

Distribution of species of Foraminifera in Mississippi

Textularia mississippiensis Cushman ___ __________ _

mississippiensis Cushman var. alabamensis Cushman. 

recta Cushman____________.. _ ________

subhauerii Cushman._________ _________ ______

Gaudryina jacksonensis Cushman __ ____ ______

Liebusella byramensis (Cushman) var. turgida (Cushman). 

Quinqueloculina laevigata D'Orbigny _____ _ _ _ _ _

anguina Terquem____._ ___________ __

hauerina? D'Orbigny __ __ .___ _________ __

Miliola saxorum Lamarck _ _ ___ _

jacksonensis Cushman. _____ ._ _ __.

Massilina decorata Cushman _ _ ___ ___ _______

cookei Cushman, n. sp____ __ _--__-________

jacksonensis Cushman. __________ .._ _________

jacksonensis Cushman var. punctato-costata Cushman. 

Spiroloculina grateloupi D'Orbigny _ ________ ______

Articulina terquemi Cushman _________ ________

Triloculina sp.?__ ______________ _ ______ ________

Cornuspira olygogyra Hantken __________ _ ____

Robulus gutticostatus (Gtimbel) _ __ _______________

alato-limbatus (Gumbel)___ _ ------ __-._

limbosus (Reuss)__------- -____----__ _ ______

arcuato-striatus (Hantken) var. carolinianus Cushman. 

Saracenaria arcuata (D'Orbigny) var. hantkeni Cushman. 

Marginulina cocoaensis Cushman__________ _ _______

Dentalina cocoaensis (Cushman) ___________ ________

cooperensis Cushman __ __________ __ ________

jacksonensis (Cushman and Applin) _____________

Nodos'aria latejugata Giimbel __ ___-____--____ ___

Guttulina irregularis (D'Orbigny) __________________

spicaeformis (Roemer) _____________ ___________

Globulina gibba D'Orbigny _______-_---__-__--___

Sigmomorphina jacksonensis (Cushman) _ ___________

jacksonensis (Cushman) var. costifera (Cushman) _ 

Polymorphina advena Cushman. ____ _______________

Nonion advenum (Cushman) --____-_ ___________ _

inexcavatum (Cushman and Applin) _____ __ _

micrum Cole. __^___--_- _____--_--_-___--.___

Nonionella hantkeni (Cushman and Applin) var. spissa 

Cushman. 

Operculina cookei Cushman. -_________-_---______

Hantkenina alabamensis Cushman ____' ____________

Bolivina gracilis Cushman and Applin ____________ __

jacksonensis Cushman and Applin ___________

jacksonensis Cushman and Applin var. striatella 

Cushman.

gardnerae Cushman. __________________________

jacksonensis Cushman. ______ _________________

Angulogerina ocalana Cushman _ ________ i ______ ___

Trifarina bradyi Cushman var. advena Cushman _ ___

Discorbis globulo-spinosa Cushman _ _______________

alveata Cushman___ _ ___________________

Eponides jacksonensis (Cushman and Applin) ________

Siphonina jacksonensis Cushman and Applin _ ________ 

Globorotalia cocoaensis Cushman ______________ _ _

6472

X

X

X

X

v

 

6473

X

V

X

X

X

X

Jackson

V

V

V

V

v

V

V

V

v

v

X

'

v

6463

X

X

V

X

X

X

v

X

v

v

X

X

X

6458

X

 

6457

v

V

X

. V

v

v

X

X

X

X

X

v

v

v

v

X

v

X

X 

X

X

X 

v

v

X 

V

6471

V

v

v

v

X

X 

X

X

V

X

X 

X

v

X

X

X

X

X

6646

X

X

X

v

X

v

X

v

X

Known stratigraphic range

Earliest

.___ _do__. 

_____do----_--.___

.-...do---. _______

_____do_-.________

.....do ________

--___do___________

._-.. do . _ ..

__--_do---_____-_-

__-__do-.-_-_____-

___-_do-----______

__--_do__---______

___._do__. ________

_____do------_-.__

__--_do-_-.__-_-__

__--_do___-____.__

_____do-----______

__--_do-----._____

_____do----____.__

_____dO---_-_.____

_____do__.________

___-_do----_._____

_____do---___--__.

___ __do __ _._

... __do - 

_____do----_--_.__

__.__do_-__-_-_.__

-__-_do------_.___

_____do__--__-____

_____do-_----_-.__

---_-do___-----.__

Cretaceous. ______

___-_do_____-_--__

--___do.__---_--__

-____do-_-_-_--.__

_____do-------____

_____do_--________

_____do-_----_.__-

 do . __

_____do_---__._-__

_____do-__--_---_-

 do ...

_____dO 

--___do-_----.__._

 do 

____.do__-. -_-_.__

_____do--.-_______.

_____ do _

_____do-_--_---__-

_____do--__-___--_

. .do---.-- _ -.

_____do-.--____-__

-_ _ do.. __ -_-__

_ _ '.do 

 do 

 do 

 __do 

 ...do 

_-___do__-_---_-_-

--__-_do 

.....do_. 

.....do ... . .

_____do__-_-----_-

Latest

Miocene. 

Oligocene.

Eocene. 

Oligocene. 

Eocene. 

Oligocene.

Recent. 

Do. 

Do. 

Do. 

Eocene. 

Do. 

Oligocene. 

Eocene. 

Do. 

Do.

Recent. 

Eocene. 

Do. 

Do. 

Oligocene. 

Eocene. 

Do. 

Oligocene. 

Eocene.

Do.

Do. 

Do. 

Do. 

Do. 

Oligocene. 

Pliocene. 

Recent. 

Do. 

Do. 

Do. 

Do. 

Oligocene. 

Do. 

Eocene. 

Do. 

Do. 

Do.

Do. 

Do. 

Do. 

Do. 

Do. 

Do.

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do.
UPPER EOCENE FORAMINIFERA OF THE SOUTHEASTERN UNITED STATES

Distribution of species of Foraminifera in Mississippi Continued

59

granosa (Hantken) var. dibollensis Cushman and 

Applin. 

cocoaensis Cushman ___________________________

jacksonensis (Cushman and Applin) _-__.___.____

jacksonensis (Cushman and Applin) var. dibollensis. 

(Cushman and Appliu) 

Planulina cocoaensis Cushman. _____________________

Cibicides pseudoungerianus (Cushman) ______________

arnericanus (Cushman) var. antiquus (Cushman 

and Applin).

mississippiensis (Cushman) _____________________

Gypsina globula (Reuss) __-_____-___________-__---_

6472

X

6473

X

Jackson

X

X

X

6463

X

X

6458

6457

V

X

X

v

X

X

6471

V

X

X

6646

X

Known stratigraphic range

Earliest

-____do_----______

_.___do___________

___._do_____ __ __

_____do__.________

_____do__--_-____.

do

_____do.__________

_____do_----._____

_____do_ -___ 

_____do._________-

Latest

Oligocene. 

Eocene.

Do. 

Do. 

Do.

Do.

Recent. 

Eocene.

Do. 

Oligocene. 

Recent.

Distribution of Eocene Foraminifera in South Carolina

Textularia mississippiensis Cushman var. alabamensis Cushman. 

subhauerii Cushman. ___________________

barrettii (Jones and Parker) ________ ____

Verneuilina propinqua H. B. Brady___________

Gaudryina jacksonensis Cushman. ___________

subquadrata Cushman. _________________

Liebusella byramensis (Cushman) var. turgida 

(Cushman) . 

Pseudoclavulina sp. ?____.___________!___

Robulus alato-limbatus Giimbel. _____________

articulatus (Reuss) var. texanus Cushman. _

limbosus (Reuss) var. hockleyensis (Cushman and Applin) . 

arcuato-striatus (Hantken) var. carolinianus Cushman. 

Saraceneria arcuata D'Orbigny var. hantkeni 

Cushman. 

Planularia truncana (Giimbel) ________________

cooperensis Cushman. ____ _____________

Marginulina karreriana Cushman, n.sp. ________

subrecta Franke. ._ _ ____________________

cocoaensis Cushman ___________________

cooperi Cushman. _ _____________________

cooperensis Cushman. _______ __________

Dentalina cocoaensis (Cushman) _____________

cooperensis Cushman. _ __________________

jacksonensis (Cushman and Applin) ______

halkyardi Cushman ____________________

cf. D. adolphina D'Orbigny _ ___________

cf. D. vertebralis (Batsch). __ __________

sp.? __ __ _ ___________ _. _ _ _____

Nodosaria latej ugata Giimbel _____________ _.

latejugata Gtimbel var. carolinensis Cushman. 

ewaldi Reuss? _ _ ______________________

fissicostata (Cushman ___________________

Lagena laevis (Montagu) ?___________________

acuticosta Reuss_ _______ _ _ __________

costata ( Williamson) _ _________________

hexagona (Williamson) ________ _________

sp.?. __________________________________

Guttuliua problema D'Orbigny _ ____________

spicaef ormis (Roemer) ______ __________

Globulina gibba D'Orbigny. ______________ __

gibba D'Orbigny var. globosa (Von Miinster) . 

gibba D'Orbigny var. tuberculata D'Orbigny. 

inaequalis Reuss _____________ __________

rotundata (Bornemann) _________________

3595

V

v

Baldock

X

6321

X

X

X

v

X

11862

V

X

X

X

v

X

X

X

X

X

X

Ashley 

marl 

works, Dorchester 

County

X

X

11857

X

X

X

X

X

X

X

X

X 

X

X

X

X

11861

v

X

X

v

X

v

X

X 

X

X 

v

X 

X

X

X 

X

11863

X

X 

v

X

X 

X

X

X 

X 

X

X

6319

X

X

X

X

X 

X

X

X

X

X

X

5531

X

X

X

X

X

X

X

X

X

X 

X

X

X

X

Known stratigraphic range

Earliest

Eocene.______

do

_____do_______

_____do.______

____-do.______

__ _do_ ______

_____do.______

_____do_______

_ __ do. ______

_.__.do_______

_____do_____._

_____do_______

____.do.______

do

_____do_______

_ __ do ___ _

_____do_______

_____do_______

.....do. ______

__ _ do. ___

___ do.. _____

__---do._..___

_ _ _do.__ ____

_____do._..___

Cretaceous? _ 

Eocene _ ____

_____do____._.

_ __ do. ______

-_-_-do_______

Cretaceous. __ 

Eocene _ ___

.____do.______

Cretaceous. __ 

Eocene _ ____

_____do_______

__.._do_____._

Cretaceous. __ 

Eocene. ______

_____do. _ ...

_____do. ______

_____do_______

_._._do_______

___ do_. _____

Latest

Oligocene.

 Do. 

Recent. 

Do. 

Eocene. 

Do. 

Oligocene.

Recent. 

Eocene. 

Do. 

Do.

Do. 

Do.

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Miocene. 

Recent. 

Do. 

Oligocene. 

Eocene.

Pliocene. 

Do. 

Recent. 

Do. 

Do. 

Do. 

Eocene. 

Recent. 

Do. 

Do. 

Do.

Pliocene.

Recent. 

Do.
60 UPPER EOCENE FORAMINIFERA OF THE SOUTHEASTERN UNITED STATES 

Distribution of Eocene Foraminifera in South Carolina Continued

jacksonensis (Cushman), var. costifera 

(Cushman) .

Nonionella hantkeni (Cushman and Applin) 

var. spissa Cushman.

Spiroplectoides curta Cushman______________

Plectofrondicularia cf. P. vaughani (Cushman) _ 

trinitatensis Cushman and Jarvis __ _____

jacksonensis Cushman var. cuneata Cushman. 

cooperensis Cushman ________ ___ ____

Virgulina recta Cushman. ______

Bolivina jacksonensis Cushman and Applin___

jacksonensis Cushman and Applin var. 

striatella (Cushman and Applin). 

spiralis Cushman_ ___________ _ ___

Reussella eocena (Cushman) _ ____ ___ _____

Uvigerina cookei Cushman, n. sp__ ______

glabrans Cushman__ _____________ ___

jacksonensis Cushman____ ___ _____

yazooensis Cushman. __ ______ ____ ___

Angulogerina ocalana Cushman_____ _ _____

cooperensis Cushman, n. sp____ __ __ ___

Gyroidina orbicularis D'Orbigny var. planata 

Cushman, n. var. 

Eponides jacksonensis (Cushman and Applin) __

jacksonensis (Cushman and Applin) var.?__

cocoaensis Cushman ______ _ __ _____

minima Cushman. ___ __ ___ ______

umbonata (Reuss)_________ __ __

Cancris brongniartii (D'Orbigny) __ ___

Siphonina jacksonensis Cushman and Applin- _

advena Cushman var. eocenica Cushman 

and Applin. v 

Cassidulina globosa Hantken______ __ __

Sphaeroidina austriaca D'Orbigny _ _ ____ !_

Globorotalia cocoaensis Cushman ____ ___

Anomalina jacksonensis (Cushman and Applin) _ 

jacksonensis (Cushman and Applin) var. 

dibollensis (Cushman and Applin). 

Planulina cocoaensis Cushman var. cooperensis Cushman. 

Cibicides lobatulus (Walker and Jacob) _____

pseudouugerianus (Cushman) __ ___ ___

americanus (Cushman) var. antiquus 

(Cushman and Applin). 

cooperensis Cushman _ _ _ _ ___________

mississippiensis (Cushman) __ __ ___ ___

mississippiensis (Cushman) var. ocalanus 

Cushman, n. var.

3595

X

X

X

__--

Baldock

____

6321

V

v

v

v

X

v

v

v

V

X

X

____

v

11862

V

X

X

X 

X

X

X

X

X

X

X

v

X

X

X

X 

X

Ashley 

marl 

works, Dorchester 

County

X

X 

X

X

------

11857

X

X

X

X

X

X 

X 

X 

v

X

X

X

X 

v

X 

X 

X

X

X 

X

X

X

X 

X

X 

X

11861

X

X

X 

X 

X

X

X 

X

X

X

X 

X

X

X

X

X

X 

X

X

X

"

11863

 X

X

X

X

X

X

X 

 X 

X

X

X 

v

v

X

X

X 

X

X

X 

X 

X

X

6319

X

X

X

X

X

v

X

X 

X

5531

X?

X

X

X

X

X

X

Known stratigraphic range

Earliest

Eocene.______

_ _ _do

do

do

- _ - do _ _ _

_ _ do

_ _ do

do

do

do

do

do

Cretaceous.--.

do

do

do

_ _ _ _ _do_ ______

Latest

Eocene. 

Do.

Oligocene. 

Eocene. 

Oligocene. 

Eocene. 

Oligocene. 

Eocene. 

Do.

Do. 

Do. 

Miocene. 

Eocene. 

Do, 

Do. 

Do. 

Do.

Do. 

Recent. 

Eocene. 

Do. 

Do.

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do.

Do. 

Do. 

Do. 

Do. 

Recent. 

Miocene. 

Eocene. 

Do.

Oligocene. 

Miocene. 

Eocene. 

Do. 

Do.

Do.

Recent. 

'Do. 

Eocene.

Do. 

Oligocene. 

Eocene.
Distribution of Eocene Foraminifera in Florida

Textularia mississippiensis Cushman var. alabamensis 

Cushman.. ___.__ _ _ __- _______ _______ __

ocalana Cushman. __________ __-_____---________.

recta Cushman.--. ____ __-- __- __-_ __ -_ ___

subhauerii Cushman..- _____-_- _- __________ .

dibollensis Cushman and Applin _____ _____________

Gaudryina gardnerae Cushman _______________________

Liebusella byramensis (Cushman) var. turgida (Cushman) _ 

Valvulina ocalana Cushman___ _ _ __________________ 

Robulus gutticostatus (Giimbel) _________ ____________

Planularia truncana (Gumbel) _____ _ __ _ ______ ____ 

Margin ulina cocoaensis Cushman___ __ _______________

Dentalina cooperensis Cushman.... ______________ ____ 

Nodosaria latejugata Gumbel _-_--____ ____ ______

fissicostata (Gumbel)__ _______ __ _____ ______ _ 

Guttulina irregularis (D'Orbigny) _______ __ _ ________

spicaeformis (Roemer)__ ________ ____________

Globulina gibba D'Orbigny. ________________________

Sigmomorphina jacksonensis (Cushman) ____ _ _ ____

Nonion advenum (Cushman). ________________ ____

Nonionella hantkeni (Cushman and Applin) var. spissa 

Cushman_________________ ___ __ _____ ___

Assilina floridensis (Heilprin) ______ ________________

Operculina ocalana Cushman_ _ _- ________________

mariannensis Vaughan________ _ _______________

Operculinella willcoxi (Heilprin) ____ _______________

Heterostegina ocalana Cushman ______ _____ __ __ _

ocalana Cushman var. glabra Cushman _ _ ____ ____

Bulimina jacksonensis Cushman ___ ____ _______

Reussella eocena (Cushman) ______ ___________

Uvigerina gardnerae Cushman_____ _ ______________ _

Angulogerina ocalana Cushman. _____ ____ _______ _

Trifarina bradyi Cushman var. advena Cushman.- _____ _

Discorbis ocalana Cushman

Lamarckina ocalana Cushman _

Rotalia sp.?___. ____________ _

Eponides jacksonensis (Cushman and Applin) ___________

cocoaensis Cushman____ _______ ________ ______

ocalana Cushman________________________ __

Siphonina jacksonensis Cushman and Applin _ ___ ____

Globorotalia cocoaensis Cushman __ ___ ______

Anomalina jacksonensis (Cushman and Applin) _ _

Ctbicides pseudoungerianus (Cushman)

mississippiensis (Cushman) _

mississippiensis (Cushman) var. ocalanus Cushman, 

n. var_____ __ _ _ _

Gypsina globula (Reuss) _ __ ___

Rupertia floridana Cushman

6754

X 

X XXXXX

1/19 X

X XX

7191 XX

X XX

7194 XXXXX

6768 XX

X 

X 

X 

X 

X XXXXXX

X X

4974 X1/48 XXXXXX

6823 X

329 X7349 XXX

7338 XX

7337 XX

X 1

7348 XXXXX7345v

X XXXX

7341 X

6817

X X

4965 XX

6785 XXXXX

X X

6815 XX3636 X

6787 X

6789 XXX

380 X

322

X X

3629 XXX

6799 XXXX

6810 X

6812 XXXXX

6814 X

6790 XXXXXXX

X 

X X

3174 X

365 X

7355 X

7367 X

3685 X

3686 X

7366 XXX

X X

5030 X

______

6808 XXX

X XXx

v

X

6807 XXXXXX

6804 XX

X XXXX

6805 XXX

X XXXXX

7378 X

X 

X 

X X

7360 XX

X X

7365 XXXX 1

3684 X7364 XXXX

Known stratigraphic range

Earliest

Eocene..___ ___

_____do-._-------

 _do 

.____do.__. - .

_ do 

_ __do 

_____do --------

 do - ._ 

_ ..do ._ 

..___do - 

. do 

__ ___do 

 do _ 

 do.. 

_ do 

. _. do 

 do_ _

_. __do-__ __-__-_

_____do ------ -_

 _do~_ _.

 do. ---- 

. do __

____ .do 

_____ dO 

_ __do 

....-do. ---------

._ ...do ..

..--.do. ----------

 do 

.. .do 

 _do 

____ do._ - --

_ ..do 

_ ..do. _ -------

.._.-do._--- _ -

-.-.-do-. . _ -

_____do---------

 do 

. do 

_ ..do-. _ . -

 do 

 do_ 

._ ...do 

_____do -------

 do 

Latest

Oligocene. 

Eocene. 

Do. 

Oligocene. 

Eocene. 

Do. 

Oligocene. 

Eocene. 

Do. 

Do. 

Do. 

Do. 

Oligocene. 

Pliocene. 

Recent. 

Do. 

Do. 

Eocene. 

Oligocene.

Eocene. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Recent. 

Oligocene.

Eocene. 

Recent. 

Do.

62330 34. (Face p. 60.) No, 1.
Distribution of species of Foraminifera in Georgia

mississippiensis Cushman var. alabamensis Cushman__________

Lenticulina convergens (Bornemann) ____________________________

Saracenaria arcuata (D'Orbigny) var. hantkeni Cushman. _____ ____

Planularia truncana (Gumbel)----__----------_-_-_-------_ -___

Dentalina cocoaensis (Cushman)____----___-_____-___--------_-_

cooperensis Cushman. _________________________ ___________

.Nodosaria latejugata GumbeL----------------------------------

ewaldi Reuss?. _ __________________________________________

Guttulina irregularis (D'Orbignv). ____._-_-- -____-_-_-_--_---_-_

 Globulina gibba D'Orbigny-. __"__--_-.--_ ._____------_---___.-

gibba D'Orbigny var. globosa (Von Minister).. _____-_-______-

miinsteri (Reuss) _ _^_.___-____ __-__-_______-_-_------_-_

Polymorphina advena Cushman_______.__ _ _ _ _.---_-_-____ __

Nonion advenum (Cushman) _____ _____ _______ ____ __________

chapapotense Cole___- ___________________ ________________

Operculina cookei Cushman__--__-._----_. _____________________

ocalana Cushman. ______--_------_____--___-_-___---_-__-_

vaughani Cushman

Hantkenina alabamiensis Cushman.. . _-____-__--_-_--________ __

Bolivina gardnerae Cushman _____-_----___-_-_____--_-_-_--_--_

Reussella eocena (Cushman) ._--_____ -__-_-_____-___--.-_---_--

Valvulineria jacksonensis Cushman_____-___-____ _ ___-___--_---

Eponides jacksonensis (Cushman and Applin) _-___________-_-__--

Anomalina bilateralis Cushman. _____-_____________-_--_--------

Cibicides lobatulus (Walker and Jacob) ... ___ _.__ ____-_-____-_

mississippiensis (Cushman) var. ocalanus Cushman, n. var __

Gypsina globula (Reuss)-__----____ ______ __ __ __ _____---____

Barnwell formation

7755 XXX

Ocala limestone

7719 ! 7704 '

XXXXXXXXXXXXXX XX

X 

X X

X 

X XXX

XXX

7189 X3966

X 

X X

xxxxx X

X X

XX

X XX

3760 X7689 X

7237 XXXXX

X X

X X

7108 XX

7115 7119 ! X X X3626

X X

7116

--__-_ X

7117 X

7235 X

3617 X7120 7123 i X X

i 7126 X

7127 XX7099 X 1

3387 X

7098 XXX

7130 X3390 X7097 XXXXXXXX

Known stratigraphic range

Earliest

Eocene. - ---

do

Eocene .___-___

... __do 

 do 

_. do 

do __-__-__--

do __________

Eocene.----.-.---

Cretaceous. .__-.. -

__---do___--_----_

._---do_-_--------

_____do___--------

_____do___-_--_-_-

do ________--

do - - - - _ _

 ..do 

C/l 1 frr\f*{*Y\ A

do

-....do __________

_ _ _ __do _

-- do

- - do

_ - - _ do _ _ _

_ _ _ _ _ do

Latest

Oligocene. 

Eocene. 

Oligocene. 

Do. 

Eocene. 

Do. 

Do. 

Do. 

Do. 

Do. 

Oligocene. 

Pliocene. 

Do. 

Recent. 

Do. 

Do. 

Do. 

Do. 

Miocene. 

Eocene. 

Recent. 

Miocene. 

Oligocene. 

Do. 

Eocene. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Do. 

Recent. 

Eocene. 

Do. 

Do. 

Do. 

Oligocene. 

Recent. 

Do. 

Oligocene. 

Eocene. 

Recent.

. (Face p. 60.) No. 2.

PLATES

01
PLATE 1 

Haplophragmoides dibollensis Cushman and Applin (p. 6)

FIGURE 1. Holotype. a, Side view; b, apertural view, X28. From well sample, .Tyler County, Tex.

Ammobaculites hockleyensis Cushman and Applin (p. 6)

FIGURE 2. Holotype. a, Side view; b, apertural view, X35. From well sample, Tyler County, Tex.

Textularia mississippiensis Cushman (p. 7)

FIGURES. Side view, X 50. From well sample, Tyler County, Tex. 

FIGURE 4. Side view, X50. U.S.G.S. station 6473.

Textularia mississippiensis Cushman var. alabamensis Cushman (p. 7)

FIGURE 5. Side view, X50. U.S.G.S. station 7227. 

FIGURE 6. Side view, X35. U.S.G.S. station 6321.

Textularia ocalana Cushman (p. 7)

FIGURE 7. Holotype. a, Side view; b, apertural view, X50. U.S.G.S. station 6812.

Textularia recta Cushman (p. 7)

FIGURE 8. Side view, X50. U.S.G.S. station 6726. 

FIGURE 9. Side view, X50. U.S.G.S. station 1/8.

Textularia subhauerii Cushman (p. 8)

FIGURE 10. Side view, X45. U.S.G.S. station 1/38.

Textularia adalta Cushman (p. 8)

FIGURE 11. a, Front view; b, apertural view, X30. U.S.G.S. station 6741. 

FIGURE 12. Front view, X30. U.S.G.S. station 6741.

Textularia dibollensis Cushman and Applin (p. 8)

FIGURE 13. Front view, X50. 4 miles east of Diboll, Angelina County, Tex.

FIGURE 14. Front view, X50. U.S.G.S. station 1/8.

FIGURE 15. Front view, X50. U.S.G.S. station 1/8.

FIGURE 16. Front view, X50. East of Diboll, Angelina County, Tex.

Textularia dibollensis Cushman and Applin var. humblei Cushman and Applin (p. 8)

FIGURE 17. a, Front view; b, apertural view, X50. From well sample, Newton County, Tex.

Textularia distortio Cushman and Applin (p. 8)

FIGURE 18. a, Front view; b, apertural view, X35. From well sample, Tyler County, Tex. 

FIGURE 19. Front view, X50. Alazan clay, Mexico.

Textularia hockleyensis Cushman and Applin (p. 9)

FIGURE 20. a, Side view; b, apertural view, X35. Bridge Creek, San Augustine County, Tex. 

FIGURE 21. a, Side view; b, apertural view, X35. Bridge Creek, San Augustine County, Tex.

62
U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 181 PLATE '

LITUOLTDAE AND TEXTULARIIDAE.

62330 35 8
0.S. GEOLOGICAL STJHVEY PROFESSIONAL PAPER 181 PLATE 2

TEXTULARIIDAE, VERNEUILINIDAE. VALVUUNIDAE, AND MILJOLJDAE.
PLATE 2

Vulvulina advena Cushman (p. 9)

FIGURE 1. a, Side view; b, apertural view, X65. U.S.G.S. station 7203.

Textulariella barrettii (Jones and Parker) (p. 11) 

FIGURE 2. a, Side view; b, apertural view, X60. U.S.G.S. station 289.

Verneuilina propinqua H. B. Brady (p. 9) 

FIGURE 3. a, Side view; b, apertural view, X50. U.S.G.S. station 292.

Gaudryina jacksonensis Cushman (p. 9)

FIGURE 4. a, Side view; b, apertural view, X50. U.S.G.S. station 6319. 

FIGURE 5. Side view, X 50. U.S.G.S. station 6319. 

FIGURE 6. Side view, X50. Alazan clay, Mexico.

Gaudryina gardnerae Cushman (p. 10)

FIGURE 7. a, Side view; b, apertural view, X65. U.S.G.S. station 1/43.

Gaudryina subquadrata Cushman (p. 10) 

FIGURE 8. a, Front view; b, side view; C, apertural view, X70. U.S.G.S. station 289.

Liebusella byramensis (Cushman) var. turgida (Cushman) (p. 11) 

FIGURE 9. Side view, X35. U.S.G.S. station 6319.

Pseudoclavulina sp.? (p. 10) 

FIGURE 10. a, Side view; b, apertural view, X60. U.S.G.S. station 292.

Valvulina ocalana Cushman (p. 10)

FIGURE 11. a, Side view; b, apertural'view, X65. U.S.G.S. station 6805. 

FIGURE 12. Side view, larger, more angled form, X65. U.S.G.S. station 6805.

Quinqueloculina laevigata D'Orbigny (p. 11) . . .

FIGURE 13. a, b, Opposite sides; c, apertural view, X70. Jackson, Miss.

FIGURE 14. a, b, Opposite sides; c, apertural view, X70. Showing variations. Jackson, Miss.

FIGURE 15. a, b, Opposite sides; c, apertural view, X70. Jackson, Miss.

Quinqueloculina longirostra D'Orbigny (p. 12)

FIGURE 16. a, b, Opposite sides; c, apertural view, X70. Jackson, Miss.

Quinqueloculina hauerina D'Orbigny? (p. 12) 

FIGURE 17. a, b, Opposite sides; c, apertural view, X70. Jackson, Miss.

Quinqueloculina anguina Terquem (p. 12)

FIGURE 18. Side view, X70. Jackson, Miss.

FIGURE 19. a, b, Opposite sides; c, apertural view, X70. Jackson, Miss.

03
PLATE 3 

Miliola saxorum Lamarck (p. 12)

FIGURE 1. Side view, X35. Jackson, Miss.

FIGURE 2. Side view, X35. Jackson, Miss.

FIGURE 3. Side view, X35. Showing the eroded form very common at Jackson, Miss.

Miliola jacksonensis Cushman (p. 13)

FIGURE 4. Holotype, side view, X35. Jackson, Miss.

FIGURES. Side view, X35. Jackson, Miss.

FIGURE 6. Side view, X35. Jackson, Miss.

Massilina jacksonensis Cushman (p. 13)

FIGURE 7. Holotype. Side view, X35. Jackson, Miss.

FIGURES. Side view, X35. Jackson, Miss.

FIGURE 9. Side view, early stages of micrbspheric form, X35. Jackson, Miss.

FIGURE 10. Side view, early stages of microspheric form, X35. Jackson, Miss.

Massilina jacksonensis Cushman var. punctatocostata Cushman (p. 14)

FIGURE 11. Holotype. Side view, X35. Jackson, Miss.

FIGURE 12. Side view, early stages of microspheric form, X35. Jackson, Miss.

FIGURE 13. Side view, X35. Jackson, Miss.

Massilina decorata Cushman (p. 13)

FIGURE 14. Side view, X35. U.S.G.S.-station 6457.

FIGURE 15. Side view, X35. U.S.G.S. station 6457.

FIGURE 16. Side view, X35. U.S.G.S. station 6457.

Massilina cookei Cushman, n. sp. (p. 13)

FIGURE 17. Side view, X35. Jackson, Miss. ,

Spiroloculina grateloupi D'Orbigny (p. 14)

FIGURE 18. Side view, X35. Jackson, Miss. .

FIGURE 19. Side view, X35. Jackson, Miss.

FIGURE 20. Side view, X35. Jackson, Miss.

FIGURE 21. Side view, X35. Jackson, Miss. . .

64
U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 181 PLATE 3

MILIOLIDAE.
U.S. GEOLOGICAL SURVEY PEOFESSIONAL PAPER 181 PLATE 4

MILIOLIDAE, OPHTHALMIDIIDAE, AND LAGENIDAE.
PLATE 4

Spiroloculina sp.? (p. 14) 

FIGURE 1. a, b, Opposite sides; c, apertural view, X120. Jackson, Miss.

Articulina terquemi Cushman (p. 14)

FIGURE 2. Holotype. a, b, Opposite sides; c, apertural view, X75. Jackson, Miss. 

FIGURE 3. a, b, Opposite sides; c, apertural view, X75. Jackson, Miss.

Triloculina sp.? (p. 15)

FIGURE 4. a, b, Opposite sides; c, apertural view, X70. Jackson, Miss. . .

Quinqueloculina sp.? (p. 12)

FIGURE 5. Side view, X35. Stovall Creek, east of Diboll, Angelina County, Tex. 

FIGURE 6. Side view, X35. Stovall Creek, east of Diboll, Angelina County, Tex.

Quinqueloculina sp.? (p. 12)

FIGURE 7. Side view, X35. Stovall Creek, east of Diboll, Angelina County, Tex. 

FIGURE S. Side view, X35.. Stovall Creek, east of Diboll, Angelina County, Tex.

Quinqueloculina sp.? (p. 12)

FIGURE 9. Side view, X35. Stovall Creek, east of Diboll, Angelina County, Tex. 

FIGURE 10. Side view, X35. Stovall Creek, east of Diboll, Angelina County, Tex.

Massilina sp.? (p. 14)

FIGURE 11. Side view, X35. Well sample, Tyler County, Tex.

FIGURE .12. Side view, X35. Well sample, Tyler County, Tex.

FIGURE 13. Side view, X35. Well sample, Tyler County, Tex. . .

Cornuspira olygogyra Hantkeh (p. 15)

FIGURE 14. Side view, X70. Jackson, Miss.

Robulus limbosus (Reuss) var. hockleyensis (Cushman and Applin) (p. 16) ,,.-. 

FIGURE 15. a, Side view; b, peripheral view, X35. Bridge Creek, San Augustine County, Tex.

Robulus articulatus (Reuss) var. texanus (Cushman and Applin) (p. 16)

FIGURE 16. Adult, a, Side view; b, peripheral view, X17. Well sample, Harris County, Tex. . 

FIGURE 17. Young. Side view, X35. Well sample, Harris County, Tex.

65
PLATE 5

Robulus gutticostatus (Giimbel) (p. 15)

FIGURE 1. Side view, X40. U.S.G.S. station 1/38.

FIGURE 2. Side view, X 40. U.S.G.S. station 1/38. . . ' '. . ,

Robulus gutticostatus (Giimbel) var. cocoaensis (Cushman) (p. 15)

FIGURE 3. Side view, X40. Cocoa, Ala. 

FIGURE 4. Side view, X40. Cocoa, Ala.

Robulus gutticostatus (Giimbel) var. yazooensis Cushman (p. 15) 

FIGURES. Side view, X 40. U.S.G.S. station .7220. ;

Saracenaria arcuata (D'Orbigny) var. hantkeni Cushman (p. 17)

FIGURE 6. Holotype. Side view, X50. U.S.G.S. station 6319.

FIGURE 7. Side view, X50. U.S.G.S. station.6319. . ;

Nodosaria fissicostata (Giimbel) (p. 22)

FIGURE 8. Side view, X50. U.S.G.S. station 1/43. 

FIGURE 9. Side view, X 50. U.S.G.S. station 1/7.

Nodosaria latejugata Giimbel var. carolinensis Cushman (p. 21)

FIGURE 10. Holotype.Side view, X35. U.S.G.S. station 7203. 

FIGURE 11. Side view, X35. Cooper River, S.C. 

FIGURE 12. Side view, X35. U.S.G.S. station 7203. 

FIGURE 13. Side view, X35. Cooper River, S.C.

66
U.8. GEOLOGICAL SURVEY PROFESSIONAL PAPER 181 PLATE 5

LAGENIDAE.
U.S. GEOLOGICAL SURVEY PBOPESSIONAL PAPER 181 PLATE 6

LAGENIDAE.
PLATE 6

Robulus propinquus (Hantken) (p. 16)

FIGURE 1. a, Side view; b, peripheral view, X35. Well sample, Tyler County, Tex.

Robulus alato-limbatus (Gurnbel) (p. 15) 

FIGURE 2. a, Side view; b, peripheral view, X35. Well sample, Tyler County, Tex.

Robulus limbosus (Reuss) var. hockleyensis (Cushman and Applin) (p. 16) 

FIG u HE 3. a, Side view; b, peripheral view, X35. Bridge Creek, San Augustine County, Tex.

Lenticulina convergens (Bornemann) (p. 17) 

FIGURE 4. a, Side view; b, peripheral view, X35. U.S.G.S. station 7719.

Robulus limbosus (Reuss) (p. 16) 

FIGURE 5. Side view, X30. U.S.G.S. station 1/38.

Robulus arcuato-striatus (Hantken) var. carolinianus Cushman (p. 17) 

FIGUHE 6. a, Side view; b, peripheral view, X30 U.S.G.S. station 1/38.

Planularia truncana (Giimbel) (p. 17) 

FIGURE 7. a, Side view; b, peripheral view, X40. U.S.G.S. station 7203.

Planularia cooperensis Cushman (p. 18) 

FiGUitis S. a, Side view; b, peripheral view, X 15. U.S.G.S. station 11863.

67
PLATE 7

Marginulina karreriana Cushman, n. sp. (p. 18)

FIGURE 1. Holotype. a, Side view; b, peripheral view, X40. U.S.G.S. station 11857. 

FIGURE 2. a, Side view; b, peripheral view, X40. U.S.G.S. station 11862.

Marginulina subrecta Franke (p. 18) 

FIGURES, a, Side view; b, peripheral view, X 40. U.S.G.S. station 11861.

Marginulina cooper! Cushman (p. 19) 

FIGURE 4. a,-Side view; b, peripheral view, X40. Cooper River, S.C.

Marginulina jacksonensis (Cushman and Applin) (p. 18)

FIGURE 5. a, Side view; b, peripheral view, X35. Well sample, Tyler County, Tcx.

,, Marginulina cocoaensis Cushman (p. 18)

FIGURE 6. Side view, X50. Megalospheric form. Cocoa, Ala. 

FIGURE 7. Side view, X 50. Paratype. Microspheric form. Cocoa, Ala.

Marginulina fragaria (Giimbel) var. texasensis (Cushman and Applin) (p. 19)

FIGURES, a, Side view; b, peripheral view, X 35. Well sample, Newton County, Tex. 

FIGURE 9. a, Side view; b, peripheral view, X35. Well sample, Newton County, Tex. 

FIGURE 10. a, Side view; b, peripheral view, X35. Well sample, Newton County, Tex.

Marginulina cooperensis Cushman (p. 19)

FIGURE 11. Holotype. Microspheric form. X50. Cooper River, S.C. 

FIGURE .12. Paratype. Megalospheric form. X50. Cooper River, S.C.
U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 181 PLATE 7

LAGENIDAE.
U.S. GEOLOGICAL STTBVEY PROFESSIONAL PAPER 181 PLATE

LAGENIDAE.
PLATE 8

Dentalina cocoaensis (Cushman) (p. 19)

FIGURE 1. Side view, X65. Cocoa, Ala.

FIGURE 2. Side view, X65. U.S.G.S. station 7203.

Dentalina cooperensis Cushman (p. 20)

FIGURE 3. Side view, X65. Cocoa, Ala.

FIGURE 4. Holotype. Side view, X65. Cooper River, S.C.

Dentalina hantkeni Cushman (p. 20)

FIGURE 5. Side view, X 65. U.S.G.S. station 6740.

FIGURE 6. Holotype. Side view, X65. U.S.G.S. station 6741.

Dentalina jacksonensis (Cushman and Applin) (p. 20)

FIGURE 7. Side view, X65. Alazan clay, Mexico.

FIGURES. Side view, X 65. Cocoa, Ala.

FIGURE 9. Side view, X65. Alazan clay, Mexico.

Dentalina halkyardi Cushman (p. 20)

FIGURE 10. Side view, X65. Cooper River, S.C.

Dentalina cf. D. adolphina D'Orbigny (p. 21)

FIGURE 11. Side view, X50. U.S.G.S. station 11857. 

FIGURE 12. Side view, X50. U.S.G.S. station 11857.

Dentalina cf. D. vertebralis (Batsch) (p. 21)

FIGURE 13. Side view, X35. U.S.G.S. station 11861. 

FIGURE 14. Side view, X35. U.S.G.S. station 11862.

Dentalina sp.? (p. 21)

FIGURE 15. Side view, X35. U.S.G.S. station 11861. 

FIGURE 16. Side view, X35. .U.S.G.S. station 11862.

69
PLATE 9

Nodosaria cookei Cushman (p. 22) 

FIGURE 1. Side view, X65. U.S.G.S. station 1/21.

Nodosaria ewaldi Reuss? (p. 22) 

FIGURE 2. Side view, X65. U.S.G.S. station 2457.

Lagena laevis (Montagu)? (p. 22)

FIGURE 3. Side view, X90. U.S.G.S. station 11857.

FIGURE 4. Side view, X65. Barnwell County, S.C. '

Lagena acuticosta Reuss (p. 23)

FIGURE 5. Side view, X90. U.S.G.S. station 11857. 

FIGURE 6. Side view, X65. U.S.G.S. station 7719.

Lagena costata (Williamson) (p. 23)

FIGURE 7. Side view, X90. U.S.G.S. station 11857. . 

FIGURE 8. Side view, X90. U.S.G.S. station 11857.

Lagena orbignyana (Seguenza) var. semiconcentrica Cushman (p. 23)

FIGURE 9. Side view, X65. U.S.G.S. station 5611. . . .

Lagena hexagona (Williamson) (p. 23) 

FIGURE 10. Side view, X90. U.S.G.S. station 11863.

Lagena sp.? (p. 23) 

FIGURE 11. Side view, X 90. U.S.G.S. station 11861.

Guttulina problema D'Orbigny (p. 23)

FIGURE 12. Side view, X 65. U.S.G.S. station 6321. . .:

Guttulina irregularis (D'Orbigny) (p. 24)

FIGURE 13. Side view, X65. Alazan clay, Mexico.

FIGURE 14. Side view, X65. U.S.G.S. station 1/21. ' " /-

FIGURE 15. Side view, X65. Cocoa, Ala. 

FIGURE 16. Side view, X 65. U.S.G.S. station 1/21. . . ... .

Guttulina spicaeformis (Roemer) (p. 24) 

FIGURE 17. Side view, X65. U.S.G.S. station 6319.

Globulina gibba D'Orbigny (p. 25) 

FIGURE 18. Side view, X65. U.S.G.S. station 6463.

Globulina glbba D'Orbigny var. tuberculata D'Orbigny (p. 26)

FIGURE 19. Side view, X65. U.S.G.S. station 1/21. 

FIGURE 20. Side view, X65. U.S.G.S. station 6319.

Globulina gibba D'Orbigny var. globosa (Yon Miinster) (p. 26)

FIGURE 21. Side view, X 65. U.S.G.S. station 7364.

Globulina inaequalis Reuss (p. 26) 

FIGURE 22. Side view, X90. Well sample, Tyler County, Tex.

Globulina minuta (Roemer) (p. 27) 

FIGURE 23. Side view, X90. U.S.G.S. station 6738.

Globulina rotundata (Bornemann) (p. 27) 

FIGURE 24. a, b, Opposite sides; c, basal view, X35. Specimen from Hermsdorf, Germany.

Globulina miinsteri (Reuss) (p. 27) 

FIGURE 25. Side view, X90. U.S.G.S. station 7719. 

70
U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 181 PLATE 9

LAGENIDAE AND POLYMORPHINIDAE

G2330- 35 G
U.S. GEOLOGICAL STJRVEY PROFESSIONAL PAPER 181 PLATE 10

POLYMORPHINIDAE.
PLATE 10 

Sigmomorphina jacksonensis (Cushman) (p. 28)

FIGURE 1. Side view, X65. Jackson, Miss.

FIGURE 2. Side view, X 05. U.S.G.S. station 1/19.

FIGURE 3. Side view, X65. U.S.G.S. station 1/19.

FIGURE 4. a, Side view; b, basal view, X65. Blue Springs, Jackson County, Fla.

Sigmomorphina jacksonensis (Cushman) var. costifera (Cushman) (p. 28)

FIGURE 5. Side view, X65. U.S.G.S. station 1/7.

FIGURE 6. a, Side view; b, apertural view, X65. Barnwell County, S.C.

Sigmomorphina vaughani Cushman and Ozawa (p. 28)

FIGURE 7. a, b, Opposite sides; c, basal view, X35. .Cooper River, S.C.

Polymorphina advena Cushman (p. 29) 

FIGURE S. Side view, X90. U.S.G.S. station 11857.

Guttulina spicaeformis (Roemer) (p. 24)

FIGURE 9. a, Side view; b, apertural view, X35. Well sample, Newton County, Tex. 

FIGURE 10. a, Side view; b, apertural view, X35. Well sample, Newton County, Tex.

Pseudopolymorphina decora (Reuss) (p. 29)

FIGURE 11. a, Side view; b, apertural view, X35. Tarkiln Creek, Trinity County, Tex. 

FIGURE 12. a, Side view; b, apertural view, X35. Tarkiln Creek, Trinity County, Tex.

Sigmoidella plummerae Cushman and Ozawa (p. 29)

FIGURE 13. a, Side view; b, basal view, X60. Specimen from Cook Mountain formation, Tex.

Pseudopolymorphina dumblei (Cushman and Applin) (p. 29)

FIGURE 14. a, Side view; b, apertural view, X35. Bridge Creek, San Augustine County, Tex. 

FIGURE 15. a, Side view; b, apertural view, -X35. Bridge Creek, San Augustine County, Tex.

Glandulina laevigata D'Orbigny var. ovata Cushman and Applin (p. 29)

FIGURE 16. Side view, X35. 4 miles east of Diboll, Angelina County, Tex. 

FIGURE 17. Side view, X35. 4 miles east of Diboll, Angelina County, Tex.

71
. . PLATE 11

Nonion advenum (Cushman) (p. 30)

FIGURE 1. a, Side view; b, apertural view, X35. U.S.G.S. station 6321.

FIGURE 2.'a, Side view; b, apertural view, X35. U.S.G.S. station 6321.

FIGURE 3. a, Side view; b, apertural view, X35. Cheapside, Gonzales County, Tex. 

FIGURE 4. Side view, X 35. Cheapside, Gonzales County, Tex.

Nonion inexcavatum (Cushman and Applin) (p. 30)

FIGURE 5. a, Side view; b, apertural view, X60. U.S.G.S. station 6321.

FIGURE 6. a, Side view; b, apertural view, X60. U.S.G.S. station 6321.

FIGURE 7. Side view, X 60. East of Diboll, Angelina County, Tex.

FIGURE 8. a, Side view; b, apertural view, X60. East of Diboll, Angelina County, Tex.

Nonion chapapotense Cole (p. 30)

FIGURE 9. Side view, X90. U.S.G.S. station 6472.

FIGURE 10. Side view, X90. U.S.G.S. station 6472. . '"

FIGURE 11. a, Side view; ,b, apertural view, X90. U.S.G.S. station 6472.

FIGURE 12. Side view, X 90. U.S.G.S. station 6473..

FIGURE 13. a, Side view; bj apertural view, X90. U.S.G.S. station 6473.

. Nonion micrum Cole (p; 30)

FIGURE 14. a, Side view; b, apertural view,. X90., U.S.G.S. station 6646. 

FIGURE 15. a, Side view; b, apertural view, X 90. U.S.G.S: station 6646.

Nonion whitsettense (Cushman and Applin) (p. 31)

FIGURE 16. Side view, X35. Well sample, Tyler County, Tex.

FIGURE 17. Side view, X35. Well sample, Tyler County, Tex. . ' .

FIGURE 18. a, Side view; b, apertural view, X35. Well sample, Tyler County, Tex.

72 . ' .
U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 181 PLATE 11

NONIONIDAE.
U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 181 PLATE 12

NONIONIDAE AND HETEROHELICIDAE.
PLATE 12 

Nonionella hantkeni (Cushman and Applin) (p. 31)

FIGURE 1. a, Side view; b, peripheral view, X35. Well sample, Tyler County, Tex. 

FIGURE 2. Side view, X35. Well sample, Tyler County, Tex.

Nonionella jacksonensis Cushman (p. 31)

FIGURE 3. a, Dorsal view; b, ventral view; c, apertural view, X90. U.S.G.S. station 11861.

FIGURE 4. Holotype. a, Dorsal view; b, ventral view; c, apertural view, X90. U.S.G.S. station 6720.

Elphidium texanum (Cushman and Applin) (p. 32)

FIGURE 5. a, b, Opposite sides; c, apertural view, X35. Well sample, Tyler County, Tex.

Nonionella hantkeni (Cushman and Applin) var. spissa Cushman (p. 31) 

FIGURE 6. a, Dorsal view; b, ventral view; c, apertural view, X70. U.S.G.S. station 11863.

Spiroplectoides curta Cushman (p. 33) 

FIGURE 7. a, Side view; b, peripheral view, X200. U.S.G.S. station 11857.

Plectofrondicularia cf. P. vaughani Cushman (p. 34)

FIGURE 8. Side view, X 50. U.S.G.S. station 11861. 

FIGURE 9. Side view, X50. Cooper River, S.C.

Plectofrondicularia trinitatensis Cushman and Jarvis (p. 34)

FIGURE 10. Side view, X90. U.S.G.S. station 11857.

Plectofrondicularia cookei Cushman (p. 34)

FIGURE 11. Side view, X65. U.S.G.S. station 6319. 

FIGURE 12. Holotype. Side view, X65. Cooper River, S.C.

Nodogenerina cooperensis Cushman (p. 34)

FIGURE 13. Side view, X90. U.S.G.S. station 11857.

73
PLATE 13

Hantkenina alabamensis Cushman (p. 49)

FIGURE 1. a, Side view; b, apertural view, X50. Cocoa, Ala.

FIGURE 2. "a, Side view; b, apertural view, X 50. Cocoa, Ala.

FIGURE 3. Side view, X50. Cocoa, Ala.

FIGURE 4. Side view, X50. Cocoa, Ala.

FIGURE 5. Side view, X 35. Sab ine River", Sabirie County, Tex.

Buliminella alabamensis Cushman (p. 34)

FIGURE 6. a, b, Opposite sides, X 100. U.S.G.S. station 7227.

Bulimina jacksonensis Cushman (p. 35)

FIGURE 7. a, Side view; b, apertural view, X50. 'Well sample, Newton County, Tex.

FIGURE 8. a, Side view; b, apertural view, X65. Cocoa, Ala.

FIGURE 9. a,' Side view; b, apertural view, X65. Cocoa, Ala.

Bulimina jacksonensis Cushman var. cuneata Cushman (p. 35)

FIGURE 10. Side view, X65. U.S.G.S. station 7220. 

FIGURE 11. Side view, X 65. U.S.G.S. station.7220.

Bulimina cooperensis Cushman (p. 35)

FIGURE 12. Holotype: a, Side view; b, apertural view, X 90. U.S.G.S. station 11857.

. FIGURE 13. Side view, X90. U.S.G.S. station 11857.

FIGURE 14. Side view, X90. U.S.G.S.-station 11857.

Bulimina ovata D'Orbigny (p. 35)

FIGURE 15. a, b, Opposite sides; c, apertural view, X90. Well sample, Tyler County, Tex. 

FIGURE 16. a, b, Opposite sides; c, apertural view, X50. U.S.G.S. station 11861.

74
U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 181 PLATE 13

HANTKENINIDAE AND BULIMINIDAE.
tJ.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 181 PLATE

BULIMINIDAE.
PLATE 14 

Virgulina dibollensis Cushman and Applin (p. 36)

B 1. a, b, Opposite sides; c, apertural view, X50. Well sample, Newton County, Tex.

FIGURE 2. Side view, X100. U.S.G.S. station 6741.

FIGURE 3. Side view, X100. U.S.G.S. station 6730.

Virgulina recta Cushman (p. 36)

FIGURE 4. a, Front view; b, apertural view, X90. U.S.G.S. station 11857.

Bolivina attenuata Cushman (p. 36) 

FIGURE 5. Side view, X65. U.S.G.S. station 1/43.

Bolivina gardnerae Cushman (p. 37)

FIGURE 6. Side view, X 65. U.S.G.S. station 7719.

FIGURE 7. a, Side view; b, apertural view, X65. U.S.G.S. station 7719.

Bolivina gracilis Cushman and Applin (p. 37)

FIGURE S. a, Side view; b, apertural view, X50. East of Diboll, Angelina County^ Tex. 

FIGURE 9. a, Side view; b, apertural view, X50. East of Diboll, Angelina County, Tex. 

FIGURE 10. Side view, X50. .U.S.G.S. station 6473. .

Bolivina jacksonensis Cushman and Applin (p. 37)

FIGURE 11. a, Side view; b, apertural view, X50. Well sample, Newton County, Tex.

FIGURE 12. Side view, X65. U.S.G.S. station 7227.

FIGURE 13. a, Side view; b, apertural view, X50. Well sample, Newton County, Tex.

Bolivina jacksonensis Cushman and Applin var. striatella Cushman and Applin (p. 37)

FIGURE 14. Side view, X65. U.S.G.S. station 6463.

FIGURE 15. a, Side view; b, apertural view, X50. East of Diboll, Angelina County, Tex.

FIGURE 1.6. a, Side view; b, apertural view, X50. East of Diboll, Angelina County, Tex.

FIGURE 17. Side view, X65. U.S.G.S. station 7159.

FIGURE IS. Side view, X65. U.S.G.S. station 6457.

7o
PLATE 15

Bolivina spiralis Cushman (p. 37)

FIGURE 1. Side view, X65. U.S.G.S. station 6319. 

FIGURE 2. Side view, X90. U.S.G.S. station 11857.

Loxostomum dalli (Cushman) (p. 38)

FIGURE 3. Side view, X65. U.S.G.S. station 7220. 

Reussella eocena (Cushman) (p. 38)

FIGURE 4. Holotype. Side view. U.S.G.S. station 7365. . 

FIGURE 5. Side view, X65. U.S.G.S. station 7365.

Reussella sculptilis (Cushman) (p. 38)

FIGURE 6.. Side view, X65. U.S.G.S. station 1/39. 

FIGURE 7. Side view,'X 65. U.S.G.S. station 1/39.

Uvigerina alata Cushman and Applin (p. 39)

FIGURE 8. Side view, X50. Well sample, Tyler County, Tex. 

FIGURE 9. Side view, X50. Well sample, Tyler County, Tex. 

FIGURE 10. Side view, X50. Well sample, Tyler County, Tex.

Uvigerina cocoaensis Cushman (p. 39)

FIGURE 11. Side view, X65. Cocoa, Ala.

FIGURE 12. Side view, X65. Cocoa, Ala.

FIGURE 13. Side view, X65. Alazan clay, Mexico.

Uvigerina cookei Cushman, n. sp. (p..39)

FIGURE 14. Holotype. Side view. X 65.. U.S.G.S. station 1/38.

FIGURE 15. Side view, X65. U.S.G.S. station 6319.

FIGURE 16. Side view, X65. U.S'.G.S. station 6319.

Uvigerina dumblei .Cushman and Applin (p. 39)

FIGURE 17. Side view,'-X 35. Well sample, Newton County, Tex.

Uvigerina gardnerae Cushman (p. 40)

FIGURE 18. Side view, X35. Well sample, Tyler County, Tex. 

FIGURE 19. Side view, X35. Well sample, Tyler County, Tex.

Uvigerina gardnerae Cushman var. texana Cushman and Applin (p. 40)

FIGURE 20. Side view, X35. Well sample, Tyler County, Tex.

Uvigerina glabrans Cushman (p. 40)

FIGURE 21. Side view, X65. U.S.G.S. station 7203. 

76
CT.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 181 PLATE l5

BULIMINIDAE.
tT.S. GEOLOGICAL SUBVEY PROFESSIONAL PAPER 181 PLATE 16

BULIMINIDAE, ELLIPSOIDINIDAE, AND ROTALIIDAE
PLATE 16 

Uvigerina jacksonensis Cushman (p. 40)

FIGURE 1. Side view, X65. Ala/an clay, Mexico. 

FIGURE 2. Side view, X65. 'U.S.G.S. station 7203. 

FIGURE 3. Side view, X65. Cocoa, Ala.

Uvigerina topilensis Cushman (p. 41)

FIGURE 4. Side view, X35. Well sample, Tyler County, Tex.

Uvigerina yazooensis Cushman (p. 41)

FIGURES. Holotype. Side view, X 65. U.S.G.S. station 7220. 

FIGURE 6. Side view, X65. U.S.G.S. station 6472.

Angulogerina ocalana Cushman (p. 41)

FIGURE 7. Holotype. Side view, X65. U.S.G.S. station 7719. 

FIGURE 8. Side view, X65. U.S.G.S. station 7719.

Angulogerina cooperensis Cushman, n. sp. (p. 42)

FIGURE 9. a, Side view; b, apertural view, X90. U.S.G.S. station 11863.

 Trifarina bradyi Cushman var. advena Cushman (p. 42) 

FIGURE 10. a, Side view; b, apertural view, X100. Jackson Miss.

Pleurostomella jacksonensis Cushman and Applin (p. 42) 

FIGURE 11. a, Side view; b, front view; c, end view, X50. Well sample, Newton County, Tex.

Spirillina vivipara Ehrenberg (p. 42) 

FIGURE 12. Side view, X90. U.S.G.S. station 6725.

Discorbis hemisphaerica Cushman (p. 43) 

FIGURE 13. a, Dorsal view; b, ventral view; c, peripheral view, X120. Jackson, Miss.

Discorbis globulo-spinosa Cushman (p. 43) 

FIGURE 14. a, Dorsal view; b, ventral view; c, peripheral view, X120. Jackson, Miss.

Discorbis ocalana Cushman (p. 44) 

FIGURE 15. a, Dorsal view; b, ventral view; c, peripheral view, X90. U.S.G.S. station 6768.

77
PLATE 17 

Discorbis assulata Cushman (p. 44)

FIGURE 1. a, Dorsal view; b, ventral view; c, peripheral view, X90. U.S.G.S. station 6727.

FIGURE 2. Holotype. a, Dorsal view; b, ventral view; c, peripheral view, X90. U.S.G.S. station 3966.

Discorbis alabamensis Cushman (p. 44) 

FIGURE 3. a, Dorsal view; b, ventral view; c, peripheral view, X90. U.S.G.S. station 1/8.

Discorbis alveata Cushman (p. 44) 

FIGURE 4. a, Dorsal view; b, ventral view; c, peripheral view, X90, U.S.G.S. station 6471.

Lamarckina ocalana Cushman (p. 45) 

FIGURE 5. a, Dorsal view; b, ventral view; c, peripheral view, X90. U.S.G.S. station 1/19.

Discorbis bulla Cushman (p. 43)

FIGURE 6. a, Dorsal view; b, ventral view; c, peripheral view, X90. U.S.G.S. station 1/39.

Eponides carolinensis Cushman, n. sp. (p. 46)

FIGURE 7. a, Dorsal view; b, ventral view; c, peripheral view, 'X40. U.S.G:S. station 5611. 

78 .
U.S. GEOLOGICAL STJBVEY PROFESSIONAL PAPER 181 PLATE 17

ROTATMTPATC

62330 35 7
0.8. GEOLOGICAL SURVEY PROFESSIONAL PAPER 181 PLATE 18

HQTALIIDAE.
PLATE 18

Discorbis subaraucana Cushman (p. 43) 

FIGURE 1. a, Dorsal view; b, ventral view; c, peripheral view, X90. U.S.G.S. station 7220.

Valvulineria jacksonensis Cushman (p. 44) 

FIGURE 2. a, Dorsal view; b, ventral view; c, peripheral view, X90. U.S.G.S. station 6741.

Gyroidina orbicularis D'Orbigny var. planata Cushman, n. var. (p. 45) 

FIGURE 3. a, Dorsal view; b, ventral view; c, peripheral view, X65. Three figures of different specimens. U.S.G.S. station 6319.

Gyroidina soldanii D'Orbigny var. octocamerata Cushman and G. D. Hanna (p. 45) 

FIGURE 4. a, Dorsal view; b, ventral view; c, peripheral view, X90. U.S.G.S. station 8572.

Eponides ocalana Cushman (p. 47)

FIGURE 5. a, Dorsal view; b, ventral view; c, peripheral view; X50: U.S.G.S. station 1/21. : -

Eponides budensis (Hantken) var. planata Cushman, n. var. (p. 47)

FIGURE 6. a, Dorsal view; b, ventral view; c, peripheral view, X90. U.S.G.S. station 56li. '" ' ' 

79
PLATE 19 

Eponides cocoaensis Cushman (p. 47)

FIGURE 1. a, Dorsal view; b, ventral view; c, peripheral view, X65. U.S.G.S. station 6457. 

FIGURE 2. a, Dorsal view; b, ventral view; c, peripheral view, X65. U.S.G.S. station 6457.

Eponides minima Cushman (p. 47)

FIGURE 3. a, Dorsal view; b, ventral view; c, peripheral view, X90. U.S.G.S. station 11863.

Eponides jacksonensis (Cushman and Applin) (p. 46)

FIGURE 4. Dorsal view, X35. East of Diboll, Angelina County, Tex. 

-FIGURE 5. Ventral view, X 65. U.S.G.S. station 1/7. 

FIGURE 6. Ventral view, X40. U.S.G.S. station 1/38. 

FIGURE 7. Peripheral view, X 50. U.S.G.S. station 1/7. 

FIGURE 8. Ventral view, X 35. East of Diboll, Angelina County, Tex.

Eponides pygmaea (Hantken) (p. 47) 

FIGURE 9. a, Dorsal view; b, ventral view; c, peripheral view, X90. North of Whitsett, Live Oak County, Tex.

Eponides umbonata (Reuss) (p. 48)

FIGURE 10. a, Dorsal view; b, ventral view; c, peripheral view, X90. U.S.G.S. station 11863.

Rotalia sp.? (p. 46)

FIGURE 11. a, Dorsal view; b, ventral view; c, peripheral view, X70. U.S.G.S. station 6812. 

80
U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 181 PLATE 19

ROTAL1IDAE,
tJ.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 181 PLATE 20

ROTALIIDAE, CASSIDULINIDAE, AND CHILOSTOMELLIDAE.
PLATE 20 

Cancris brongniartii (D'Orbigny) (p. 48)

FIGURE 1. a, Dorsal view; b, ventral view; c, peripheral view, X 50. U.S.G.S. station 11861.

Siphonina jacksonensis Cushman and Applin (p. 48)

FIGURE 2. a, Dorsal view; b, ventral view; c, peripheral view, X 65. U.S.G.S. station 1/7.

FIGURE 3. a, Dorsal view; b, ventral view; c, peripheral view, X65. U.S.G.S. station 6457.

FIGURE 4. a, Dorsal view; b, ventral view; c, peripheral view, X 65. U.S.G.S. station 6473.

FIGURES. Ventral view, X 65. U.S.G.S. station 1/7.

FIGURE 6. Ventral view, X 35. East of Diboll, Angelina County, Tex.

FIGURE 7. Ventral view, X 35. East of Diboll, Angelina County, Tex.

FIGURES. Dorsal view, X 35. East of Diboll, Angelina County, Tex.

Siphonina advena Cushman var. eocenica Cushman and Applin (p. 49)

FIGURE 9. Ventral view, X35. Tarkiln Creek, Trinity County, Tex. - ' 

FIGURE 10. Ventral view, X35. Tarkiln Creek, Trinity County, Tex. 

FIGURE 11. Dorsal view, X35. Tarkiln Creek, Trinity County, Tex.

Cassidulina globosa Hantken (p. 49)

FIGURE 12. a, Side view; b, peripheral view, X 90. U.S.G.S. station 11857. <

Sphaeroidina austriaca D'Orbigny (p. 49) 

FIGURE 13. a, Dorsal view; b, ventral view, X90. U.S.G.S. station 11863.

Lamarckina jacksonensis Cushman (p. 45) 

FIGURE 14. a, Dorsal view; b, ventral view; c, peripheral view, X 90. U.S.G.S. station 1/39.
PLATE 21

Globorotalia cocoaensis Cushman (p. 50)

FIGURE 1. a, Dorsal view; b, ventral view; c, peripheral view, X45. Cocoa, Ala.

FIGURE 2. Ventral view, X45. U.S.G.S. station 6457.

FIGURES, a, Dorsal view; b, ventral view; c, peripheral view, X 45. Cocoa, Ala.

Anomalina bilateralis Cushman (p. 50)

FIGURE 4. a, Dorsal view; b, ventral view; c, peripheral view, X 90. U.S.G.S. station 6725. 

FIGURE 5. a, Dorsal view; b, ventral view; c, peripheral view, X90. Jackson, Miss.

Anomalina granosa (Hantken) var. dibollensis Cushman and Applin (p. 50)

FIGURE 6. Ventral view, X35. East of Diboll, Angelina County, Tex.

FIGURE 7. a, Dorsal view; b, ventral view; c, peripheral view, X35. East of Diboll, Angelina County, Tex.

Anomalina jacksonensis (Cushman and Applin) (p. 51) 

FIGURES, a, Dorsal view; b, ventral view; c, peripheral view, X 35. Bridge Creek, San Augustine County, Tex.

Anomalina jacksonensis (Cushman and Applin) var. dibollensis (Cushman and Applin) (p. 51) 

FIGURE 9. Dorsal view, X35. East of Diboll, Angelina County, Tex.

Anomalina jacksonensis (Cushman and Applin) var. texana (Cushman and Applin) (p. 51) 

FIGURE 10. Dorsal view, X35. Well sample, Tyler County, Tex.

Anomalina affinis (Hantken) (p. 51)

FIGURE 11. Dorsal view, X 35. East of Diboll, Angelina County, Tex. 

FIGURE 12. Ventral view, X35. East of Diboll, Angelina County, Tex.

Anomalina cocoaensis Cushman (p. 51)

FIGURE 13. a, Dorsal view; b, ventral view; c, peripheral view, X 90. Cocoa, Ala.

82 
U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 181 PLATE 21

GLOBOROTALJIDAE AND ANOMALINIDAE.
tT.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 181 PLATE 22

ANOMALINIDAE.
PLATE 22 

Cibicides americanus (Cushman) var. antiquus (Cushman and Applin) (p. 53).

FIGURE 1. Ventral view, X35. East'of Diboll, Angelina County, Tex. 

FIGURE 2. Dorsal view, X35. East of Diboll, Angelina County, Tex.

Cibicides mississippiensis (Cushman) (p. 54)

FIGURE 3. a, Dorsal view; b, ventral view; c, peripheral view, X45. U.S.G.S. station 6321.

Cibicides lobatulus (Walker and Jacob) (p. 52)

FIGURE 4. a, Dorsal view; b, ventral view; c, peripheral view, X45. U.S.G.S. station 6319.

FIGURE 5. a, Dorsal view; b, ventral view; c, peripheral view, X45. U.S.G.S. station 7755.

FIGURE 6. Ventral view, X45. U.S.G.S. station 7755.

Planulina cocoaensis Cushman (p. 52) 

FIGURE 7. a, Dorsal view; b, ventral view; c, peripheral view, X45. Cocoa, Ala.

Planulina cocoaensis Cushman var. cooperensis Cushman (p. 52)

FIGURE 8. a, Dorsal view; b, ventral view; c, peripheral view, X50. U.S.G.S. station 11861.
PLATE 23

Cibicides pseudoungerianus (Cushman) (p. 52) 

FIGURE 1. a, Dorsal view; b, ventral view; c, peripheral view, X65. U.S.G.S. station 6319.

Cibicides yazooensis Cushman (p. 53)

FIGURE 2. a, Dorsal view; b, ventral view; c, peripheral view, X90. U.S.G.S. station 6473.

Cibicides cooperensis Cushman (p. 53)

FIGURE 3. a, Dorsal view; b, ventral view; c, peripheral view, X70. U.S.G.S. station 11857.

Gypsina globula (Reuss) (p. 54)

FIGURE 4. X30. U.S.G.S. station 6810. 

FIGURE 5. X30. U.S.G.S. station 6810.

Rupertia floridana Cushman (p. 55)

FIGURE 6. Holotype. Side view, X 18. U.S.G.S..station 6790. 

FIGURE 7. Apertural view, X 18. U.S.G.S. station 6790.

Carpenteria carolinensis Cushman, n. sp. (p. 55)

FIGURE 8. Viewed from above, X 18. U.S.G.S. station 5611.

Carpenteria monticularis Carter (p. 55)

FIGURE 9. Viewed from above, X18. U.S.G.S. station 5611. 

84
tT.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 181 PLATE 23

ANOMALINIDAE, PIANORBULINIDAE, AND RUPERTIIDAE.
INDEX

Page

Alubnmn, species of Foraminifera found in, table sliowing___.._ 60 

stations from which material lias been collected in_ 3-4

Ammobaculltes liockleyensls Cushman and Applln________ 6, pi. 1 

cf. A. follacens Brady _ _ _ _ ___ _ 6

Angulogerlmi byrnmensis Cushman-_ _ _-_ 42 

coopcrensis Cushman, n. sp____ __________-- 42, pi. 16 

danvlllensls Howc and Wallace-- _____ 4 

ocalann Cushnitin________________________ 41-42, pi. 16 

occidentnlis Cushman _ _ _ _ 41, 42

Anomallnu nflinis (l-fantken) ____ __________ 51, pi. 21 

barrowi Cushniau and Ellisor_ _ _______ __ 4

bllatcralis Cushman _ _ _____ 50, pi. 21

cocoacnsis Cushman ___ _____________ 51, pi. 21

danvillensis Mowe and Wallace__- ________ 4 

granosa (H'antkcn) var. dibollensis Cushman and

Applin_____________________________ 50, pi. 21

jacksonensls (Cushman and Appliu) ____ 51, pi. 21

var. dibollensis (Cushman and Applin) _______ 51, pi. 21

var. limbosa Cushman and Ellisor___________ 4

var. texana (Cushman and Applin) ______ 51-52, pi. 21

mlssisslppiensls Cushman __ __ ____ _ 54

Anomallnldae__ __________________ 50-54, pis. 21-23

Artlcullna byramensis Cusliman.___ _____________ 14

gibbosula D'0rblgny__________________________ 14

nltlda D'Orbigny_____________________________ 14 

terquoml Cushman __ ________ 14, pi. 4

Assllina floridcnsis (Hcllprin)______________________ 32

Barnwell formation, collections from, list of stations_______ 3 

Blfarlna dalli Cushman_____________________________ 38

Bollvlna attenuata Cushman______________________ 36, pi. 14

cadata Cushman var. byramensis________________ 36 

caelata Cushman var___ _____ _ _________ 36

danvillensis Howc and Wallace_______________ 4

gardnerae Cushman__ _ _ ________ 37, pi. 14 

gracllis Cushman and Applin________________ 37, pi. 14

var. danvillensis Howe and Wal.'ace_______-__ 4

jacksonensis Cushman and Applin-_ __________ 37, pi. 14 

var. slrlatclla Cushman and Applln__________ 37, pi. 14

mexicana Cushman var. horizontalis Cushman______ 4 

mlssisslppiensls Cushman_______________________ 37 

ouachltocnsis Howe and Wallace_______________ 4

punctata D'Orbigny _______________________ 37 

rctlculata Hantkcn___________________________ 36 

spiralis Cushman ________________________ 37-38, pi. 15 

tortuosa H. B. Brady_________________________ 38 

cf. B. punctata Cushman_______________________ 37 

Bnliinlna buchiana D'Orbigny.___________________ 35

coopcrensis Cushman________________________ 35, pi. 13

in data____________________________,_ 35

jacksoncnsis Cushman _____________________ 35, pi. 13

var. cuncnta Cushman________________..__ 35, pi. 13

ovata D'Orbigny _____________________ 35-36, pi. 13

pyrula D'Orbigny ________________________,_ 4

sculptllis Cushman _________________________ 35 

truncana G umbel________________________ 35

Bullminella alabamensis Cusliman_______________ 34-35, pi. 13 

baslstriata Cushman and .Tarvis var. nuda Howe and Wallace __ _____..______ 4 

elcgantissima (D'Orbigny) ______________________ 4 

Bulimlnidae __________________ 34-42, pis. 13-16

Camerlntdae _ ____________________________ 32-33

Cancrls brongniartil (D'Orbigny)__________________ 48, pi. 20

danvillensis Howe and Wallace_________________ 4

 Carpentaria carollnensis Cushman, n. sp_____________ 55, pi. 23

montlcularls Carter _______________________ 55, pi. 23

Cassldulina globosa Hantkcn____________________ 49, pi. 20

Cassldullnidae ____________________________ 49, pi. 20

Cassldullnoldes braziliensis (Cushman)________________ 5

Castle Hayne marl, collections from, list of stations_______ 2

Page

Ceriopora globulus Reuss 54 

Chilostoinella cylindroides Reuss _ 5 

Chilostomellidae ______ ___________ - ________ 49, pi. 20 

Chilostomelloides oviformis (Sherborn and Chapman) _ 5 

Cibicides americanus (Cushman) var. antiquus (Cushman and

Applin) ____________________________ 53, pi. 22 

cooperensis Cushman _ _ _ __ __ _ _ _ 53, pi. 23

danvillensis Howe and Wallace 5

lobatulus (Walker and Jacob) _ __ __ ___ 52, pi. 22

mississippiensis (Cushman) _ _ 54, pi. 22

var. ocalanus Cushman, n. var _ 54

ouachitaensis Howe and Wallace- __ _ __ 5

pseudoungerianus (Cushman) _ 62-53, pi. 23

yazooensis Cushmau _ _ 53, pi. 23

Clavulina nodosaria D'Orbigny var. uovangliae Cushman 10

szaboi __ : _ 19

Cooper marl, collections from, list of stations 2

fauual relationships of ______ 1

Cornuspira olygogyra Hantken 15, pi. 4

Cristellaria alato-limbata (Giimbel) Cushman and Appliu __ 16

arcuata Hantken 17

arcuata D'Orbigny _ 17 

articulata Reuss var. texana Cushman and Applin _ 16 

cassis (Fichtel and Moll) ________ ___ _ 18

convergens Bornemann__ _ _ _ _ _ 17 

gutticostata (Giimbel) var. cocoaensis ( Cushman )_ 15 

jacksonensis Cushman and Applin 18 

limbosa (Reuss) var. hockleyensis Cushman and Applin 16 

propinqua Hantken 16 

truncana Giimbel _____________ _ _ __ _ _ __ _ 17

Darbyella danvillensis Howe and Wallace _ 5

Dcntalina budensis Hantken ____ _ _____ ______ 20

cocoaensis (Cushman) __ _ ____ _ ____ 19-20, pi. 8

communis D'Orbigny . 5

var. ernaciata Reuss _ _ _______ __ _ __ 5

consobrina D'Orbigny _ __ _ _ 19

cooperensis Cushman _________ ._ _ _ 20, pi. 8

flliformis (D'Orbigny)- _ _ 5 

halkyardi Cushman _______________ _____ 20-21, pi. 8 

hantkeni Cushman __ _________ _ 20, pi. 8 

jacksonensis (Cushman and Applin) ______ _ ______ _ 20, pi. 8

mexicana (Cushman) var. danvillensis Howe and Wallace- 5 

cf. D. adolphina D'Orbigny ____________________ 21, pi. 8 

cf. D. vertebralis (Batsch) ___________________ 21, pi. 8 

sp., Howe and Wallace ____ __' __ ___ 5

sp - __ - ____ 21, pi. 8

Dimorphina danvillensis Howe and Wallace __ ______ 5 

Discorbina jacksonensis Cushman and Applin var. texana Cushman and Applin__ __ _______ ____ _ 51

Discorbis alabameusis Cushman __ _ ____ ____ 44, pi. 17 

alveata Cushman __ ______________________ 44, pi. 17 

assulata Cushman __ _ _ ___________ ___ __ __ 44, pi. 17

bulla Cushman ___________________________ 43-44, pl.17 

farishi Cushman and Ellisor __ _ _____ _ __ ___ __ 5 

globulo-spinosa Cushman ____________________ 43, pi. 16 

hemisphaerica Cushman _____________ _____ 43, pi. 16 

jacksonensis Cushman and Applin ______ ________ _ 51 

var. dibollensis Cushman and Appliu _ _ 51 

ocalana Cushman _______________ __ _ __ ___ 44, pi. 16

orbicularis (Terquem) ______________ ________ 44 

subaraucana Cushman ____ _ _ __ ___ _ _______ 43, pi. 18

Distribution of species, tables showing _________ __ _ 56-60 

Dyocibicides danvillensis Howe and Wallace _______ __ __ 5

Ellipsoidimdae __________ _ __________________ 42, pi. 16

Ellipsonodosaria ? sp ____ - ___ - ______ - __________ 5

Elphidium eocenicum Cushman and Ellisor ____ _ ____ 5

texanum (Cushman and Applin) _______ __ _ _ __ 32, pi. 12

sp.

Entosolenia costata Williamson. 23

85
86 INDEX

Page

Bpistomina elegans (D'Orbigny)-________________ _ 5

Eponides budensis (I-Iantken) var. planata Cushman, n. var__ 47, pi. 18

byramensis Cushman__ ___________ _____ 46

carolinensis Cushman, n. sp________ ____ 46-47, pi. 17

cocoaensis Cushman___________ _____________ 47, pi. 19

exigua (H. B. Brady)________________________ 48 

haidingeri D'Orbigny_________________________ 46

jacksonensis (Cushman and Applin)______ _ 46, pi. 19 

mexicana Cushman___________________________ 46

minima Cushman_ ____________________ 47 48, pi. 19

ocalana Cushman ____________________ 47, pi. 18

ouachitaensis Howe and Wallace-_______-_ ____ 5

pygmaea (Hantkeu)____________________ 47, pi. 19

umbonata (Reuss) ______________________ 48, pi. 19

Florida, species of Foraminifera found in, table showing - 60 

stations from which material has been collected in__ __ 3

Foraminifera, Oligocene, publications on__ ________ _ 1

upper Eocene, list of species described in other publications_______________________________ 4-6

Gaudryina garclnerae Cushman_______'____________- 10, pi. 2

jacksonensis Cushman______________________ 9-10, pi. 2

subquadrata Cushman _______________________ 10,pl. 2

triangularis Cushman_________________ _ _ 10

Georgia, species of Foraminifera found in, table showing- 60 

stations from which material has been collected in__ _ 2 3

G'andulina dimorpha (Borncinann)_____ ____ 5

laevigata D'Orbigny____ _______________ 5

var. ovata Cushman and Applin_________ 29-30, pi. 10

Globigerina bulloides D'Orbigny________________ _ 5

danvillensis Howc and Wallace______ _______ _ 5

ouachitaensis Howe and Wallace___________ _ 5

sp_ ____ ____________________ 5

Globorotalia canariensis (D'Orbigny)_____________ 50

cocoaens's Cushmaii-____________________ 50, pi. 21

crassa D'Orbigny___________.________________ 50

Globorotaliidae __ ____________________________ 50, pi. 21

Globulina acuta Reuss____________________________ 26

aequalis D'Orbigny __ _______________ _____ 26

amplectens Bornemann__________________:____ 25

amplectens Reuss______________________ 26

amygdaloides Reuss-___________________-__ 26

asperula Giimbel______________________________ 26 

discreta Reuss_____________________________ 26

gibba D'Orbigny_____________________________ 25, pi. 9 

var. globosa (Von Minister)--__________ 26, pi. 9

var. gloimila Fornasini_____________ .____ 24

var. tuberculata D'Orbigny____ ____ ____ 26, pi. 9 

globosa Reuss___________________-______ 25

guttula Reuss____________________________ 24 

inaequalis Reuss____..____________ -___ 26-27, pi. 9

inflata Reuss_____________________________ 26

irregularis D'Orbigny_______________________ 24

irregularls Terquem_____________________ 24

minuta (Roemer) __________________________ 27, pi. 9 

minister! (Reuss)________________________ 27, pi. 9

. ovalis Terquem__________________________ 25

oviformis Terquem__..________________________ 25 

rotundata (Bornemann) ____________________ 27-28, pi. 9

spinosa D'Orbigny_ ________

subalpina Gtimbel___-______

subgibba Giimbel-____ _____.

translucida D'Orbigny-________ 

transversa Terquem____________

tuberculata 0'Orbigny________.

tubulosa D'Orbigny_ _________

turbinata Terquem_________

varians Terquem________________

Giimbelina sp _______________.

Guttulina austriaca D'Orbigny_____. 

byramensis Cushman and Schenck. 

centrata Terquem _________

communis D'0rbigny_._________

cretacea Alth ______________

dilatata Reuss ______________

diluta Bornemann____________

fracta Boruemann ___________

globosa Bornemann _________. 

gravida Terquem and Terquem___ 

incurva Bornemann__________

____________ 26 

____________ 27 

_____________ 25 

-________ 26 

_____________ 25 

_____________ 26 

_____________ 25 

_____________ 26 

_____________ 28 

_____________ 5 

_______ _ 5 

_____________ 24 

_____________ 24 

_____________ 23 

	24

_____________ 24

_____________ 26

_____________ 27

__'___________ 27

_____________ 25

_____________ 27 

irregularis D'Orbigny_________________________ 24, pi. 9

obtusa Bornemann_________________________ 27

Page

Guttulina plancii D'Orbigny- __ __________________ 24 

ponderosa Terquem __ _ _____________ 25 

problenia D'Orbigny__ ______________ 23-24, pi. 9 

rotundata Bornemann_ _____________________ 27

semiplana Reuss____ ___________ 24 

spicaeformis (Roemer)_________________ 24-25, pis. 9-10 

turgida Reuss ___ _ _ ___________ 27

sp.

(Globulina) gibba Cushman__.___________________ 25

Gypsina globula (Reuss)--_ _ _ __________ 54, pi. 23

Gyroidina danvillensis Howe and Wallace_______ ____ 5

orbicularis D'Orbigny var. planata Cushman, n. var_ 45-46, pi. 18

soldanii D'Orbigny var. octocamerata Cushmaii and G. D.

Hanna___________________________ 45, pi. 18

liantkenina alabamensis Cushman___ ______ 49-50, pi. 13 

alabamensis Cushman var. primitiva Cushman and Jarvis_ 50 

brevispina Cushman _ _ _______________ 50 

kochi (Hantken)_____________________________ 50 

lehneri Cushman and Jarvis___________________ 50

longispina Cushman ___ ___________ 5, 50

mccordi Howe and Wallace__ _____________ 5

mexicana Cushman _ ____.._____ 50

var. aragonensis Nuttall_ ___________ 50 

Hantkeninidae _ _ _ ________ 49-50, pi. 13 

Haplophragmoides dibollensis Cushman and Applin ______ 6, pi. 1

Heterohelicidae ____ __ _ _ - ________ 33 34, pi. 12

Heterostegina ocalana Cushman__ _______________ 33

ocalana Cushman var. glabra Cushman ___ __ _ 33

Hopkinsina danvillensis Howe and Wallace-__ __ _ 5

Jackson formation, collections from, list of stationsfaunal characteristics of__ _ ___.

Lagena acuticosta Reuss__ ___ ___ __ 23, pi. 9 

costata (Williamson) __ _________ _ ____ __ 23, pi. 9

globosa (Montague)_____ _ _____ 5 

hexagona (Williamson)__________________ __ 23, pi. 9 

hispida Reuss______ _ _ _ __.._ ___ . 5 

laevis (Montagu) ?_____________ ____ _ _ 22, pi. 9 

orbiguyana (Seguenza) var. covicentrica Sidebottom _ 23 

var. semiconcentrica Cushman________ ____ 23, pi. 9

ouachitaensis Howe and Wallace __ 5 

pleniluna Howe and Wallace_ __ ______ 5

striata (D'Orbigny) var. strumosa Reuss_ _ _ 5 

sp., Howe and Wallace- ____ ____ __ _ 5 

sp.? _________________________ 23, pi. 9

Lagenidae - ______ ____ ________ 15-23, pis. 4-9 

Larnarckina jacksonensis Cushman_ ___________ 45, pi. 20

ocalana Cushman _ ___ _ __ 45, pi. 17

Lenticulina articulata (Reuss) var. texana Howe and Wallace- 16

convergens (Bornemann) 17, pi. 6

fragaria (Giimbel) var. texasensis Howe and Wallace.- 19

sp________ _ _ - -___ _ __ 5

Lepidocyclina ocalana Cushman____ _ ___ _ ____ 55

Liebusella byramensis (Cushman) var. turgida (Cushman) _ 11, pi. 2 

cylindrica Hantken ___ _ _ _ ___ 11 

eocaena (Giimbel) _ _ 11 

flintiana Cushmaii _ ____ 11

Lituolidae________________ __.._____ __ _._ 6, pi. 1

Loxostomum clalli (Cushniau) ___ 38, pi. 15

Marginulina abbreviata Karrer __ ___ 

alazanensis (Cushman)_ _____ ______________

cocoaensis Cushman_ _ _ _ __ _ 18-19, 

cooperensis Cushman __ 19,

cooperi Cushman_____ _ _ _ __ __ 19, 

fragaria Giimbel var. texasensis Cushman and Applin__ 19, 

hauerina (D'Orbigny) _ ____________________________

jacksonensis (Cushman and Applin) _ __ 18, 

karreriana Cushman, n. sp__ ____ ___ __ 18,

subrecta Franke_____-_________-__-______-_ 18, 

triangularis D'Orbigny .var. danvillensis Howe and Wallacesp.

Marginulina? sp____ _ _ _______ _ _

Massilina cookei Cushman, n. sp__________ ---__-- 13, 

decorata Cushman-.__ __ _ 13, 

 humblei Cushman and Ellisor _ _ _ _ _____

jacksonensis Cushman______ ___________ 13-14,

var. punctatocostata Cushman- __ _ 14, 

pratti Cushman and Ellisor____ _ ____ _____

sp. ?_____________________________________ 14,

18

19

pi. 7

pi. 7

pi. 7

pi. 7

19

pi. 7

pi. 7

pi. 7

5

5

5

pi. 3

pi. 3

5

pi. 3

pi. 3

5

pi. 4
INDEX 87

Page

Millolu jacksonensis Cushman _ 13, pi. 3

saxorum Lamarck.___ _- 12 13, pi. 3

(Mlllolil:es) saxorum Lamarck 12

Mlliolidac_____________ ______________ 11-15, pis. 2-4

Mississippi, species ol! Foraminifera found in, table showing 58-59

stations from which material has been collected in 4

Moore, Margaret S., acknowledgments to _ _ 2

Nautilus lobatula Walker and Jacob 52

Nodogcneriim cooperensis Cushman _ 34, pi. 12

Nodosaria abyssorum H. B. Brady 20

afllnis Keuss_____ _ __ 21

bacllloides Hantkcn____ __ 21

budensis Hantken 20 

cocoacnsis Cushman____ _ _ 19 

communis D'Orbigny. 20 

coiispurcata Keuss . _ 22 

cookel Cushman ____ _ __ 22, pi. 9 

cwaldi Reuss? _ ___ . _ 22, pi. 9 

fill form Is D'Orbigny ______ __ _ _ ______ 19

flssicostata (Giimbel) ______________ _ 22, pi. 5

insecta Schwager __ _ 20

jacksonensis Cushman and Applin____ 20

latejugata Giimbel 21

via*, carolinensis Cushman_______ __ _ 21-22, pi. 5

longiscata D'Orbigny 5 

raphanus 21 

sp__'_____________ - 19 

(Dentalina) spinulosa Halkyard 20 

(Glandulina) laevigata D'Orbigny var. ovata Cushman and

Applin _______ __ 29

Nonion advenum (Cushman) _ 30, pi. 11 

appllnl Howc and Wallace _ _ 5 

chapapotcnsc Cole _ 30, pi. 11 

danvillensls Howe and Wallace 5 

hantkcni (Cushman and Applin) var. fayettei Cushman

and Ellisor_ _ __ 5 

inexcavatum (Cushman and Applin)_ __ 30, pi. 11 

laevls (D'Orbigny) var. margiuatum Cushman and Ellisor. '5 

micrum Cole_____________________ __ 30-31, pi. 13 

scaphum (Flchtel and Moll) var. inflatum Cushman and

(Ellisor_ _ 5 

whltscttenso (Cushman and Applin)__ _ __ _ 31,

sp.

pi. 11 

(5 

6

pi. 12 

pi. 12 

pi. 12 

6

11-12 

30

30

31

3.1

57

2

32

33

2-4 

32

32

33 

ocalnua Cushman__________________________ 32, 33

vaughani Cushman____ ___ ______ ______ 32-33 

willcoxi Cushman______ ______ ___ _ _ 33

Opercullnella willcoxi (Heilprin) __ __ ______ 33 

Ophthalmidildae _____ __________________ 15, pi. 4

Planorbulluidae _____ ______ ____ 54, pi. 23

Planularla catahoulaensis Howe and Wallace___________ 6

cooperensis Cushman__ ___ _ __ ____ 18, pi. 6

danvillensis Howe and Wallace____:____________ 6

ouachitaensis Howe and Wallace________________ 6

truncana (Giimbel) ______ ______ ________ 17, pi. 6

Plauullna cocoaensls Cushman_________ ______ 52, pi. 22

cocoaensis Cushman var. cooperensis Cushmau______ 52, pi. 22

Ploctofrondicularla cookei Cushman_______________ 34, pi. 12

trinitatensis Cushman and Jarvis_______ __ __ 34, pi. 12

cf. P. vaughani Cushman_ ______ ______ 34, pi. 12

Pleurostomella jacksonensis Cushman and Applin______ 42, pi. 16

Polymorphina acuta Reuss___ _______ _ ____ 27

acuta Roomer _ _ _ ___ _ _________ 20

Nonionella danvillensis Howc and Wallace.- _ _

hantkeni (Cushman and Applin) 31

var. splssa Cushman__ _ ___ _ 31

jacksonensis Cushman _ __ _____ _ 31-32

turgida (Williamson) __ ______ _ .._______ 

Nonionidae____________________________ 30-32, pis.

Nonion ina ad vena Cushman_ _ _ __ _ 

advena Cushman var. inexcavata Cushmau and Applin-_ 

hantkeni Cushman and Applin__ __ ________

whil:settcnsis Cushman and Applin _ 

North Carolina, species of Foraminifera found in, table showingStations from which material has been collected in____

Nummulitcs floridensis Heilprin ____ 

willcoxi Heilprin ____ _______________

Ocala limestone, collections from, list of stations.

Operculina cookei Cushman 

florldensis Cushman ___ 

mariannensis Vaughan _ 

Page

Polymorphina advena Cushman __ ___________ 29, pi. 10

amygdaloides Reuss (1856) _______ __________ 27

amygdaloides Reuss (1870)____________________ 26

var. terquemiana Fornasini_______________ 28

amygdaloides Terquem _ _ ________ 28

austriaca D'Orbigny var. io Cushman- and Applin______ 24

byramensis Cushman_ __ _ _________ 24

communis D'Orbigny var. etrusca Fornasini__________ 27

communis Roemer _ _______ ______________ 23

compressa D'Orbigny var. dumblei Cushman and Applin__ 29

compressa Nuttall ____ ___________________ 28

cretacea Egger__ __ _ _ 24

decora Reuss____ _ _ ____ ___________ 29

equalis Cushman_____________________________ 26

frondiformis Searles Wood var. brevis Jones_________ 26

gibba H. B. Brady, Parker, and Jones______________ 25

var. aequalis H. B. Brady, Parker, and Jones___-- 26

gibba Burrows, Sherborn, and Bailey____________ 27

gibba (D'Orbiguy) var. orbicularis Karrer__________ 25

globosa Von Miinster__ _____________________ 26

gracilis Ueuss _ ________ 27

inflata Terquem and Terqueiu___ _____________ 25

jacksouensis Cushman__ ______________________ 28

var. costifera Cushman_____ __ _ _______ 28

lactea Burrows, Sherborn, and Bailey_ _____ _ 27

lactea Sidebottom___________________________ 24

lactea Walker and Jacob var. communis Williamson____ 24

var. diffusa Cushman-_____ ____________ 25

lactea elongate variety, Weller______ _________ 27

minuta Roemer__________________________ 27

miinsteri Reuss__ _ ____________________ 27

obtusa D'Orbigny _ __ __ _ ______ 27

ovulum Reuss ____________________________ 27

pauperata Terquem _ _______ _________ 26, 28

problema D'Orbigny ______ _ ___________ 23

var. Hosius _______ _ _______ 23

rotundata H. B. Brady, Parker, and Jones__-___-___ 27

sinailis Reuss___ ____________ ________ 27

sororia Reuss___________________________ 27

spicaeformis Roemer__ _____________________ 24

spinosa H. B. Brady, Parker, and Jones__ _ __ _ 26

subcruciata Terquem_ _ ___________ 27

texana Cushman and Applin- ___ ___________ 29

translucida Fornasini _ _____ ___________ 26

tuberculata H. B. Brady, Parker, and Jones__________ 26

turgida Reuss-__ ____ ____________ 27

uviformis Reuss_ _ _ _ __ ____________ 24

(Globulina) gibba D'Orbigny var. ovoidea Egger_______ 25

var. pirula Egger ____________________ 25

var. subgibba Egger ____________________ 25 

(Globulina) gibba Egger _________ 25

(Globulina) spinosa Egger _____ ___ _____ 26

(Globulina) tuberculata Egger__________________ 26

(Guttulina) lata Egger_________________________ 24

(Guttulina) problema Andreae______________ 23

(Guttulina) aff. P. minima Andreae- _-__ _______ 27 

Polymorphinidae---_______________________ 23-30, pis. 9-10

Polystomella texana Cushman and Applin___________ _ 32

Pseucloclavulina sp.?___________________________ 10, pi. 2 

Pseudopolymorphina decora (Reuss) _______________ 29, pi. 10

dumblei (Cushman and Applin)__ ___ ____ __ 29, pi. 10 

Pullenia elongata Hantken_________ _ _________ 31 

Pulvinulina afBnis Hantken_____________ ________ 51 

jacksonensis Cushman and Applin_ _________ 46 

Pulvinulinella danvillensis Howe and Wallace___________ 6 

Pyrgo inornata (D'Orbigny) var. danvillensis Howe and Wallace____ _____- ____ _____ 6

Quinqueloculina anguina Terquem__ ______ 12, pi. 2 

danvillensis Howe and Wallace__ --_______ 6 

hauerina D'Orbigny? 12, pt. 2 

laevigata D'Orbigny____ ___________________ 11, pi. 2 

longirostra D'Orbigny _ __ 12, pi. 2 

saxorum D'Orbigny _ __ _ _____ _ 12 

seminula (Linng)_______________ ____________ 6 

sp.? _________________________________ 12, pi. 4

Reussella eocena (Cushman) _ __ __ 38, pi. 15

rectimargo (Cushman)__ __ _ _ ______ 38

sculptilis (Cushman)____________ ________ 38, pi. 15

spinulosa (Reuss) _____ _--___ _____ 38

var. glabrnta (Cushman)____ ____ 38

Reussia eocena Cushman___ ___________ __,.,._ ..,. 38
INDEX

. - : Page 

Robulina alato-limbata Gumbel 15

; gutticostata Giimbel 15

limbosa Reuss : 16

Robulus alato-limbatus (Gumbel) - 15-16, pi. 6

; arcuato-striatus (Hantken) var. carolinianus Cushmari 17, pi. 6

 articulatus (Reuss) var. texanus (Cushman and Applin)- 16, pi. 4

clericii (Fornasini) _ 6

. cultratus Montfort _ _ 6

; gutticostatus (Gumbel) 15, pi. 5

var. cpcoaensis (Cushman) ___ __ 15, pi. 5

; . var. yazooensis Cushman___________ _ __ 15, pi. 5

limbosus (Reuss) _ . 16, pi. 6 

var. hockleyensis (Cushman and Applin)____ 16, pi. 4

mayi Cushman and Parker_ _ 6

 .; propinquus (Hantken) 16-17, pi. 6

Robulus? danvillensis Howe and Wallace _ 6

Rotalia brongniartii D'Orbigny __ , 48

.: ecuadorensis Galloway and Morrey ___ 48

umbonata Galloway and Morrey- ______ _ 48

sp.?_________________^__ ______________ 46, pi. 19

Rotaliiclae___________-________-_______ 42-49, pis. 16-20 

Rotalina brougniartii Reuss______ __ ____ 48

umbonata Reuss _ _ . 48 

Rupertla floridana Cushman__ _ _ _ 55, pi. 23 

Rupertiidae ______ - __ 55, pi. 23

Saracenaria arcuata (D'Orbigny) var. hantkeni Cushman 17, pi. 5 

: danvillensis Howe and Wallace 6

 . moresiana Howe and Wallace 6 

" Scutella bed " ' - _ 1 42

" Serpula (Lagena) laevis ovalis " Walker and Boys_ 22 

Sigmoidella plummerae Cushman and Ozawa __ _ 29, pi. 10 

Sigmomorphina jacksonensis (Cushman) _________ 28, pi. 10

jacksonensis (Cushman) var. costifera Cushman and 

; Ozawa__ ______: ___________ ____ 28, pi. 10

; regularls (Von Miinster)___________ ____________ 28

.. semitecta (Reuss) 6

var. terquemiana (Fornasini)_ T._ ___ 28

vaughani Cushman and Ozawa _ 28, pi. 10

Siphonina advena Cushman var. eocenlca .Cushman and Applin

: . . 49, pi. 20

 : carltoni Cushman and Ellisor _ _ _ '_ 6

danvillensis Howe and Wallace_ _ __ 6

jacksonensis Cushman and Applin__ _ _____ 48-49, pi. 20

South Carolina, species of Foraminifera found in, table showing- 59-60

stations from which material has been collected in_ _ 2

Sphaeroidina austriaca D'Orbigny __ _____ 49, pi. 20

Spirillina vivipara Bhrenberg___ __ 42-43, pi. 16

Spiroloculina excavata H. B. Brady____ ________ 14

grateloupi D'Orbigny _ ____ 14, pi. 3

sp. ?_______________________--___________ 14, pi. 4

Spiroplectoides curta Cushman________________ 33-34, pi. 12

Stations from which material has been collected, list of _ 2-4

: -tables showing _ 56-60

Texas, species of Foraminifera found in, table showing_____ 56-57

stations- from which material has been collected in_____ 4

Textularia adalta Cushman__________ __________ 8, pi. 1

barrettii Jones and Parker_________ _________ 11

broussardi Howe and Wallace__ ___. ___ 6

conica Goes _ __________ ________ 11

danvillensis Howe and Wallace _ ____ 6

Page

Textularia dibollensis Cushmau and Applin_____-_________; 8, pi. 1.

var. humblei Cushman and Applin____________ 8, pi. 1 

distortio Cushman and Applin__ _ _ _____ 8-9, pi. 1 

hockleyensis Cushman and Applin___.__________ 9, pi. 1

mississippiensis Cushman _ _ _ _ ______ 7, pi. 1

var. alabamensis Cushman_________________ 7, pi. 1

var. rhomhoidea Cushman and Ellisor__________ 6

ocalana Cushman _ _ __ ____ 7, pi. 1

ouachitaensis Howe and Wallace______ _ ______ 6

recta Cushman. _ _ _________________ 7-8, pi. 1

subhauerii Cushman _ __ _________ 8, pi. 3

Textulariella barrettii (Jones and Parker)____________ 11, pi. 2

Textulariidae________ ___________ 7-9, pis. 1-2

Tri farina bradyi Cushman var. advena Cushman_______ 42, pi. 16 

Triloculina rotunda D'Orbigny var__________________ 6 

sp. ? __ _ _ _ 15, pi. 4 

" Triloculina striata Brown " __________________ 14 

Trochammina teasi Cushman and Ellisor__ __________ 6 

Truncatulina americana Cushman var. antiqua Cushman and

Applin _ __ ______ 53

lobatula (Walker and Jacob) D'Orbigny__________ 52

pseudoungeriana Cushman ___________________ 52

ungeriana H. B. Brady _____________________ 52

Tubulogenerina eocenica Cushman and Ellisor__________:_ ' 6

Uvigerina alata Cushman and Applin________________ 39, pi. 15 

auberiana D'Orbigny _ ________________ 40 

canariensis D'Orbigny _________________ 40 

cocoaensis Cushman _ ____ ___ 39, 40, pi. 15 

cookei Cushman, n. sp____ ________________ 39, pi. 15 

danvillensis Howe and Wallace_________________ 6

dumblei Cushman and Applin_______________ 39-40, pi. 15 

farinosa Hantken _ _______________ 6

gardnerae Cushman __, ______________ 40, pi. 15

var. texana Cushman and Applin___________ 40, pi. 15 

glabrans Cushman _ _____________- 40, pi. 15 

jacksonensis Cushman- _____________ 39, 40-41, pi. 16 

ocalana _________ _____._____________ 40

peregrina var. bradyana Cushman_______________ 39, 4J 

tenuistriata Reuss__ ______________________ 39 

topilensis Cushman _ _ _______ 41, pi. 16 

yazooensis Cushman ________________ 41, pi. 16

Valvulina ocalana Cushman- _ __ __ ____ 10-11, pi. 2 

Valvulineria jacksonensis Cushman__________'._________ 44, pi. 18

texana Cushman and Ellisor___________________ Q 

Valvulinidae _____ __ __________________ 10-11, pi. 2

Ventilabrella sp______________________________ 6

Vermiculum laeve Montagu _ __________________ 22

Verneuilina propinqua H. B. Brady_________________ 9, pi. 2

rectimargo Cushman __ _________________ 38

sculptilis Cushman__ _ _______'-,____________ 38

Verneuilinidae _ _______________ 9-10, pi. 2

Virgulina danvillensis Howe and Wallace______________ 6

dibollensis Cushman and Applin________________ 36, pi. 14

mexicana Cole__ __________________ 36

recta Cushman _________________________ 36, pi. 14

schreibersiana Czjzek- _ ______ __________ 36

Vulvulina advena Cushman _ ____ ________ 9, pi. 2

" Zeuglodon bed "_______________________'______ 35, 41

o