FRANK N. YOL A Checklist of the

• American Bidessini

(Coleoptera:

Dytiscidae-Hydroporinae

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY • 1969 NUMBER 33
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SMITHSONIAN CONTRIBUTIONS TO

ZOOLOGY

NUMBER 33

Frank N. Young A Checklist of the

American Bidessini

(Coleoptera:

DytiscidaeHydroporinae)

ISSUE D

NQV25B89

SMITHSONIAN INSTITUTION PRESS

CITY OF WASHINGTON
ABSTRACT

Young, Frank N. A Checklist of the American Bidessini (Coleoptera: DytiscidaeHydroporinae). Smithsonian Contributions to Zoology, 33:1-5. 1969.—A list of

the American water beetles of tribe Bidessini (Dytiscidae-Hydroporinae). Typespecies are listed for each genus with references to the original designations.

Contribution No. 809 from the Zoological Laboratories of Indiana University aided

by GB 2768 from the National Science Foundation

Official publication date is handstamped in a limited number of initial copies and is recorded

in the Institution's annual report, Smithsonian Year

UNITED STATES GOVERNMENT PRINTING OFFICE

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For sale by the Superintendent of Documents, U.S. Government Printing Office

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Frank N. Toung A Checklist of the

American Bidessini

(Coleoptera:

DytiscidaeHydroporinae)

This checklist attempts to bring together all of the

names of water beetles of the family Dytiscidae, subfamily Hydroporinae, tribe Bidessini, which have been

proposed for species from the Americas and to assign

them as far as possible to currently accepted genera. In

large part, reliance has been placed upon the original

descriptions of the species, but wherever possible specimens have been examined. The latter include material

in the British Museum (Natural History), United

States National Museum, California Academy of Science, Museum of Comparative Zoology, University of

Michigan Museum of Zoology, American Museum of

Natural History, Zoologische Sammlung des Bayerischen Staates (Munich), and the collection of the

writer.

An asterisk (*) preceding a species name indicates

that I have personally examined the type or authentic

cotype or paratype material and confirmed the generic

assignment or synonymy. Those preceded by a question

mark (?) are tentatively assigned pending further

study. Those names considered to be synonyms, some

of which probably represent geographical subspecies,

are indicated by an "s-" preceding the name.

The arrangement adopted is that of the Leng Catalogue (Leng, 1920; Leng and Mutchler, 1927, 1933;

Blackwelder, 1939) and Blackwelder's Checklist of

Coleopterous Insects of Mexico, Central America, the

West Indies, and South America (1944-1957). The

Frank N. Young, Department of Zoology, Indiana University,

Bloomington, Indiana 47401.

citations to the literature are in the abbreviated form

of these works and refer to the bibliographies in them

with the exception of those followed by an exclamation

mark (!), which are cited in full at the end of the

present paper.

The references have been cross checked with the

Coleopterorum Catalogus (Zimmermann, 1920), Zoological Record, and the original books or papers.

The shortcomings of such a list as this are evident,

but some arrangement of the American species is necessary to provide a basis for further investigation.

There are dozens, possibly hundreds of Bidessines

awaiting study, particularly from Mexico and South

America. This work has been seriously hampered by

premature publication of incomplete descriptions. The

male genitalia prove to be especially valuable in the

diagnosis of species in many of the genera, and I urge

that all future descriptions of new species be accompanied by figures of these structures.

Platydessus perforatus Guignot (55-4), described as

a Bidessine, is not included because it was found to be a

Macrovatellus (Vatellini) and was transferred to that

genus by Spangler (66-57).

Amarodytes Regimbart 00-524

(Type-species Amarodytes percosioides Regimbart, 1900, subsequent designation of Guignot 39:53!)

boggianiiRegimbart00-526 (Paraguay)

duponti (Aube) 38-568 (Brazil)

guidiGuignot57-37! (Brazil)
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

o&erf/mri Regimbart00-526 (Brazil)

percosioides Regimbart 00-525 (Brazil)

plaumanni Gschwendtner 35-152 (Brazil)

pulchellus Guignot 55-273! (Colombia)

segrix Guignot 50-7! (Brazil)

testaceo-pictusRegimbart 00-527 (Brazil)

undulatus Gschwendtner 54-111! (Peru)

*peregrinus Balfour-Browne 47-444! (Panama)

pulicarius (Aube) 38-494 (U.S.A.)

*regimbarti Balfour-Browne 47-430! (Mexico)

*sobrinus Balfour-Browne 47-445! (Panama)

tibialis Regimbart 95-337 (Brazil)

*truncatusBalfour-Browne 47^140! (Brazil)

*vicinus Balfour-Browne 47-428! (British Honduras)

Anodocheilus Babington 41-15

(Type-species Anodocheilus maculatus Babington, 1841,

monobasic)

exiguus (Aube) 38-490 (U.S.A.)

germanus (Sharp) 82-20 (Guatemala)

guatemalensis Zaitzev 10-223 (Guatemala)

*maculatus Babington 41-16 (Brazil)

silvestrii Regimbart 03^49 (Argentina)

Bidessodes Regimbart 00-528

(Type-species Bidessodes semistriatus Regimbart, 1900, present designation.)

*elongatus (Sharp) 82-25 (Guatemala)

fragilis Regimbart 00-530 (Paraguay)

knischi Zimmermann 21-198 (Brazil)

obscuripennis Zimmermann 21-198 (Brazil)

plicatus Zimmermann 21-198 (Brazil)

semistriatus Regimbart 00-529 (Brazil)

subsignatus Zimmermann 21-199 (Brazil)

Bidessonotus Regimbart 95-331

(Type-species Bidessonotus obtusatus Regimbart, 1895, subsequent designation of J. Balfour-Browne 47—427! to replace

Hydroporus adumbratus Clark, 1862, species incognitus

designated Guignot 46-114!)

bicolor Guignot 57-36! (Brazil)

*browneanus Balfour-Browne 47-441! (Jamaica)

*caraibus (Chevrolat) 63-199 (Cuba)

*championi Balfour-Browne 47-436! (Guatemala)

*fallax Balfour-Browne 47-429! (Cuba)

*inconspicuus (LeConte) 55-290! (Louisiana)

*longovalis (Blatchley) 19-310! (Florida)

melanocephalus Regimbart 95-332 (Brazil)

mexicanus Regimbart 95-333 (Mexico)

*mobilis Balfour-Browne 47-427! (Mexico)

*morosus Balfour-Browne 47-446! (Mexico)

*nepotinus Balfour-Browne 47-442! (Trinidad)

obtusatus Regimbart 95-336 (Brazil, Paraguay)

Brachyvatus Zimmermann 19-134 (as subgenus of

Bidessus)

(Type-species Hyphydrus acuminatus Steinheil, 1869, subsequent designation of Young 54-58!)

acuminatus (Steinheil) 69-249! (Argentina)

*apicatus (Clark) 62-184 (Mexico)

*hydrovatoides (Sharp) 82-20 (Guatemala)

*seminulum (LeConte) 78-377 (Florida)

*borrei (Sharp) 82-346 (Brazil)

Hemibidessus Zimmermann 21-196 (as subgenus of

Bidessus)

(Type-species Bidessus [Hemibidessus] conicus Zimmermann,

1921, subsequent designation of Guignot 46-114!)

bifasciatus Zimmermann 21-197 (Brazil)

celinoides Zimmermann 21-197 (Brazil)

*conicus Zimmermann 21-196 (Brazil)

plaumanni Gschwendtner 35-374; 36-19 (Brazil)

Hydrodessus J. Balfour-Browne 53-55!

(Type-species Hydrodessus siolii Balfour-Browne, 1953,

monobasic.)

Brinkius Guignot 57-38!

(Type-species Brinkius biguttatus Guignot, 1957, original

designation.)

*amazonensisSpangler 66-380! (Peru)

biguttatus (Guignot) 57-39! (Brazil)

brasiliensis (Guignot) 57-40! (Brazil)

*nanayensis Spangler 66-382! (Peru)

octospilus (Guignot) 57-39! (Brazil)

pereirai (Guignot) 57-41! (Brazil)

* siolii Balfour-Browne 53-56! (Brazil)

Hypodessus Guignot 39-52, 54!

(Type-species Bidessus cruciaius Regimbart, 1903, original

designation.)
NUMBER 3 3

Brachybidessus Gschwendtner 54-111!

(Type-species Amarodytes [Brachybidessus] titschacki Gschwendtner, 1954, monobasic)

cruciatus (Regimbart) 03-50 (Argentina)

cruciferGxxignot39-60! (Cayenne)

cruvilineatus (Zimmermann) 21-195 (Brazil)

dasythrix Guignot 54—40! (Cayenne)

*frustrator Spangler 66-3 78! (Peru)

titschacki (Gschwendtner) 54-112! (Peru)

Liodessus Guignot 39-53! (as subgenus of Bidessonotus, sensu Guignot)

(Type-species Hydroporus affinis Say, 1823, original designation. )

*abjectus (Sharp) 82-24 (Mexico)

acollensis Guignot 55-272! (Peru)

affinis (Say) 23-104 (North America)

*charlotti (Clark) 62-182 (Mexico)

*emilianus (Clark) 62-183 (Mexico)

s-erythrostomus (Mannerheim) 52-305 (Alaska)

*macularis (LeConte) 52-206! (Colorado)

nanus (Aube) 38-496 (U.S.A.)

nigrinus (Casey) 84-80 (California)

*obscurellus (LeConte) 52-206! (California)

microreticulatus (Hatch) 28-219 (Washington)

andinus Guignot 58-3! (Peru)

atomus Guignot 57-36! (Brazil)

bogotensis Guignot 53-111! (Bolivia)

bonariensis (Steinheil) 69-249! (Argentina)

*cantralli (Young) 53-111! (Michigan)

chilensis (Solier) 49-189 (Chili)

*crassus (Sharp) 82-365 (Brazil)

dilatatus (Regimbart) 95-330 (Brazil)

delfini (Regimbart) 99-1 (Chili)

emaciatus Guignot 53-112!(Brazil)

flavofasciata (Steinheil) 69-249! (Argentina)

*flavicollis (LeConte) 55-291 (New York)

*fuscatus (Crotch) 73-391 (Lake Superior to Florida)

*hobbsi (Young) 50-4! (Florida)

involucer (Brinck) 48-4! (Tristan da Cunha)

microscopicus (Zimmermann) 21-93 (Brazil)

miersi (White) 47-33 (Brazil)

burrus (Guignot) 53-111 (Brazil)

*nitidus (Babington 41-14 (Brazil)

*obscurus (Babington) 41-14 (Brazil)

patagonicus (Zimmermann) 22-35 (Patagonia)

strobeli (Steinheil) 69-249! (Chili)

lusculentus Guignot 48-164! (Argentina)

*uruguensis (Sharp) 82-365 (Uruguay)

Microdessus Young 67-79!

(Type-species Bidessus atomarius Sharp, 1882, original

designation)

* atomarius (Sharp) 82-366 (Brazil)

Neobidessus Young 67-79!

(Type-species Hydroporus pullus LeConte, 1855, original

designation)

alternatus (Regimbart) 89-384 (Venezuela)

*pescheti (Zimmermann) 21-194 (Brazil)

brasiliensis (Regimbart) 95-238 (Brazil)

Jcancellosus (Guignot) 57-34! (Peru)

*corumbensis (Zimmermann) 21-195 (Brazil)

curticornis (Regimbart) 03-54 (Argentina)

*discoidalis (Sharp) 82-369 (Brazil)

duplicatus (Guignot) 53-111! (Brazil)

liliputanus (Aube) 38-591 (Brazil)

*obtusus (Sharp) 82-24 (Mexico)

persimilis (Regimbart) 95-329 (Mexico)

* pullus (LeConte) 55-290 (Louisiana)

s-*floridanus (Fall) 17-168 (Florida)

*shermani (Mutchler) 18-82 (Florida)

*subvittatus (Zimmermann) 21-193 (Brazil)

surinamensis (Regimbart) 89-390 (Surname)

trilineatus (Zimmermann) 25-255 (Brazil)

*vittatipennis (Zimmermann) 21-194 (Brazil)

*youngi (Leech) 48-392! (Lower California)

Neoclypeodytes Young 67-77, 78

(Type-species Hydroporus plicipennis Crotch, 1873, original

designation.)

americanus (Guignot) 36-144 (Mexico)

*centralis (Sharp) 87-753 (Mexico)

*cinctellus (LeConte) 52-206 (Arizona)

*curtulus (Sharp) 87-753 (Mexico)

*decoratus (Fall) 17-167 (Arizona)

* disc e dens (Sharp) 82-19 (Mexico)

*discretus (Sharp) 82-350 (Texas)

*fryi (Clark) 62-421 (Mexico)

*latifrons (Sharp) 82-22 (Guatemala)

*leachi (Leech) 48-11! (California)

luctuosus (Guignot) 49-43! (Central America)

*lynceus (Sharp) 82-368 (Guatemala)
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

*obesus (Sharp) 82-349 (Texas)

*ornatellus (Fall) 17-167 (Oregon)

*pictodes (Sharp) 82-348 (North America)

*plicipennis (Crotch) 73-388 (California)

*quadrinotatus (Sharp) 82-21 (Mexico)

*quadrisignatus (Sharp) 82-21 (Mexico, Guatemala)

rugulosus (Guignot) 36-146 (Mexico)

*substriatus (Sharp) 82-23 (Mexico)

Trogloguignotus Sanfilippo 58-160!

(Type-species Trogloguignotus concii Sanfilippo, 1958, monobasic)

concii Sanfilippo 58—161! (Venezuela)

Uvarus Guignot 39-53! (as subgenus of Amarodytes)

(Type-species Hydroporus lacustris Say, 1823, original

designation)

?adumbratus (Clark) 62-183 (Mexico)

*amandus (LeConte) 52-207! (Arizona)

costaricensis (Guignot) 39-59! (CostaRica)

*falli (Young) 40-30! (Florida)

granarius (Aube) 38-501 (U.S.A.)

*inflatus (Young) 50-1! (Florida)

lacustris (Say) 23-103 (U.S.A.)

*magensis (Clark) 62-182 (Mexico)

?omichlodes (Guignot) 57-37! (Brazil)

*rogersi (Young) 41-29! (Florida)

*spretus (Sharp) 82-24 (Mexico)

Jsubtilis (LeConte) 52-206 (California)

*subornatus (Sharp) 82-23 (Mexico)

?subplicatus (Hatch) 53-197! (Oregon)

*suburbanus (Fall) 17-169 (New York, Pennsylvania)

*texanus (Sharp) 82-366 (Texas)

Literature Cited

Balfour-Browne, J.

1947. A Revision of the Genus Bidessonotus Regimbart

(Coleoptera: Dytiscidae). Transactions of the

Royal Entomological Society of London, 98:425-

448, 12 figures.

1953. Hydrodessus Gen. N. Hydroporinarum (Coleoptera: Dytiscidae). Proceedings of the Royal Entomological Society of London, (B) 22:55-56,

1 figure.

Blackwelder, R. E.

1939. Catalogue of the Coleoptera of America, North of

Mexico. Fourth Supplement, 1933-1938:16-18.

1944. Checklist of the Coleopterous Insects of Mexico,

Central America, the West Indies, and South

America. United States National Museum Bulletin, 185 (1):72-80.

1957. Ibid. (Bibliography), 185 (6) =927-1492.

Blatchley, W. S.

1919. Insects of Florida. Va. Supplementary Notes on

the Water Beetles. Bulletin of the American

Museum of Natural History, 41:305-322.

Brinck, P.

1948. Coleoptera of Tristan da Cunha, Results of the

Norwegian Scientific Expedition to Tristan da

Cunha, 1937-1938, 17:1-122.

Gschwendtner, L.

1954. Dytiscidae (Col.). In Beitrage zur Fauna Peru:

Nach der Ausbeute der Hamburger Sudperu-Expedition 1936, anderer Sammlungen, wie auch auf

Grund von Liter aturangab en. 4:109—115, 24

figures. Jena: Fischer.

Guignot, F.

1939. Contribution a l'fitude des Bidessus. Bulletin

Societe d'Etude Science Naturel de Vaucluse,

1939:51-61, 10 figures.

1946. G6notypes des Dytiscoidea et des Gyrinoidea.

Revue Frangaise d'Entomologie, 13:112—118.

1948. Vingt-septieme Note sur les Hydrocanthans. Bulletin Mensuel de la SocUti Linneenne de Lyon,

17:163-171, 1 figure.

1949. Vingt-huitieme Note sur les Hydrocanthares. Bulletin de la Societe Entomologique de France,

54:42-47.

1950. Trente-troisieme Note sur les Hydrocanthares.

Bulletin Institute Science Naturel Belgique, Brussels, 26:1-10, 3 figures.

1953. Trente-neuvieme Note sur les Hydrocanthares.

Revue Frangaise d'Entomologie, 20:109-117, 2

figures.

1954. Quarantieme Note sur les Hydrocanthares. Bulletin et Annals Societe Entomologique de Belgique,

90:40-45.

1955. Quarante-deuxieme Note sur les Hydrocanthares.

Revue Frangaise d'Entomologie, 22:270-276, 1

figure.

1955. Description de Nouveaux Dytiscidae Principalement de l'Amerique du Sud. Bulletin Institute

Royale Naturel Belgique (Brussels), 31:1-12.

1957. Contribution a la Connaissance des Dytiscides SudAmericains (Coleoptera). Revue Frangaise d'Entomologie, 24:33-45, 10 figures.

1957. Beitrage zur Kenntnis der Insektenfauna Boliviens,

Teil II. Coleoptera I. Dytiscidae. (2. Contribution a l'fitude des Dytiscides Sud-Americains),

Opuscula Zoologica, 6:1-10.

Leech, H. B.

1948. Contributions Toward a Knowledge of the Insect

Fauna of Lower California, No. 4, Coleoptera:

Haliplidae, Dytiscidae, Gyrinidae, Hydrophilidae,

Limnebiidae. Proceedings of the California Academy of Sciences, 24 (4) 375-483, 2 plates.
NUMBER 3 3

1948. Bidessus Leachi, a New Species of Dytiscid Water

Beetle from Northern California. Proceedings of

the Entomological Society of British Columbia,

44:11-12, 3 figures.

Leng, C. W.

1920. Catalogue of the Coleoptera of America, North of

Mexico. Pages 75-86. New York: J. D. Sherman.

Leng, C. W. and A. J. Mutchler.

1927. Ibid. Supplement, 1919-1924: 16-19.

1933. Ibid. Second and third supplements, 1925-1932:

14-16 and 81.

Sanfilippo, N.

1958. Viaggio in Venezuela di Nino Sanfilippo, V: Descrizione di Trogloguignotus Concii N. Gen. N. Sp.

di Dytiscidae Freatobio. Annali del Museo Civico

di Storia Naturale di Genova, 70: 159-164, 2

figures.

Spangler, P. J.

1966. Changes in Nomenclature and Reassignment of

Platydessus Perforatus (Coleoptera: Dytiscidae)

Coleopterists Bulletin, 20: 57-58, 1 figure.

1966. The Catherwood Foundation Peruvian-Amazon Expedition XIII—Aquatic Coleoptera (Dytiscidae;

Noteridae; Gyrinidae; Hydrophilidae; Dascillidae;

Helodidae; Psephenidae; Elmidae). Monographs

of the Academy of Natural Science, Philadelphia,

14: 377-443, 8 plates.

Hatch, M. H.

1953. The Beetles of the Pacific Northwest, Part I: Introduction and Adephaga. University of Washington Publications in Biology, 16:1-340, 37 plates.

LeConte, J. L.

1852. Descriptions of New Species of Coleoptera from

California. Annals of the Lyceum of Natural History of New York, 5:185-216.

Steinheil, E.

1869. Symbolae ad Historiam Coleopterorum Argentiniae

Meridionalis. . . . Atti della Societa Italiana di

Scienze Naturali, 12:238-260 (Centuria I).

Young, F. N.

1940. Description of a New Dytiscid from Florida (Coleoptera : Dytiscidae). Florida Entomologist, 23:30.

1941. Bidessus Rogersi, New Water Beetle from Florida

(Coleoptera: Dytiscidae). Florida Entomologist

24:29-31, 2 figures.

1950. Two New Species of Bidessus from the Apalachicola

Flat woods of Florida (Coleoptera: Dytiscidae).

Occasional Papers of the Museum of Zoology, University of Michigan, 526:1-6, 2 figures.

1953. A New Species of Bidessus from Southern Michigan

(Coleoptera: Dytiscidae). Bulletin of the Brooklyn Entomological Society, 48:111-112.

1954. The Water Beetles of Florida. University of Florida Biology Series, 5:1-238, illustrated.

1967. A Key to the Genera of American Bidessine Water

Beetles, with Descriptions of Three New Genera

(Coleoptera: Dytiscidae, Hydroporinae). Coleopterists Bulletin, 21:75-84.

Zimmermann, A.

1920. Coleopterorum Catalogus, Pars 71: Dytiscidae,

Haliplidae, Hygrobiidae, Amphizoidae, pages

1-426.

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