if

Coefficient of Coefficient of
Conservation | Taxon Conservation | Taxon

Flatiron Lake Bog

Sarracenia purpurea

Silica Sand Quarry Bog }

|

Triadenum virginicum

Scirpus cyperinus

Toxicodendron vernix

Vaccinium
angustifolium

Triadenum virginicum

Vaccinium
corymbosum

Vaccinium macrocarpon

Vaccinium
macrocarpon

Vaccinium corymbosum

Viburnum dentatum var.
lucidum

Woodweardia virginica

R = 146;N = 31;1 = 26.22 |



avegOvaNv Ione
aVaOvVIdv
aVveOV INL
aVaOVaEdAD
aVaOVasdAD
aVaOVeadAD
avaOvaddAD
aVaOvasdAD
avaovasddAD
avaOvesadAD
aVeOVaEdAD
aVaOvaadAD
avaovasdAD
avaOvesdAD
avadovasdAD
avaovusdAD
AVEOVAadAD
aVaOVesadAD
aveovasdAD
avadovaddAD
aVeOvVeadAD
AVSOVaEdAD
aVaOVaadAD
aVaOVasdAD
avdovaddAD
aVeOVaEdAD
aVeOVaadAD

aVaOVaadAD
avaOvasdAD
avaOvasdAD
avaovasddAD
aVeOVaEdAD
aAvaOvaadAD

STUUOsIpI09

area

Bueruyore>

Hpoom

Twouapytia
vaprourdina “rea eapyourdina
(suajoouue *D) endiquie “rea voprourdna

BINpLYA
suaosalia
BEvOISOA

(erensos “D) eeqnomND

weyaqun
eurydAy

queso)

euuodsin

ediesoyouy
saprornquy

Buon
voreia
wOyNUD
rou
Tues
woaragns
wos
vouruens
eyedns
sis
esourenbs
Wesuards

eaployeydas “sea soproursseds
soproruedieds “sea soproruedieds
eyeda33e “sea soporuesieds
(eauady “D) BrED01s

euenioys

A15

JESS USSSA USSSSUESEAUUSSENUSSOLS

MANMOLSNANALATOCSMOoTrANnweoanrnnoeSoennnvtvoSd

555

Appendix A Checklist of Vascular Plants


aVaOVUaLSV
aVeOVaELSV
AVAOVOISSVud
SVEOVOISSVud
AVEOVOISS Vad
AVAOVOISSVad
AVEOVOISSVUd
AVAOVOISSVud
AVAOVOISSVud
AVAOVOISSVud
AVAOVOISSVud
SVAOVOISSVa
AVEOVOISSva
aVaOVOISsSvas
SVEOVOISSVaE
AVAOVOISSVUd
aVSOVAVNNVO
AVAOVINONOIG
aVSOVINNVdNVO
aVaOVINNVdNVO
AVIOVINNVdNVD
AVAOVINNVINVO
AVaOVOISSVad
AVEOVOISSVa
SVaOVITIT
aVIOVINATOANOD
AVEOVINATOANOD
AVEOVINATOANOD
AVEOVINONONVA
AVaOVdIHOYO
AVaOVHORLLITIVO
AVEOVHORILITTVO
SVEOVHOILITIVO
aVaOVaV

sueynu
sapjoupueoe

equip

ByogTpunjor

sysuajeid “18a sysuayeid
sunsnyed “rea sisuyead
eorueapAsuad

ejooruare “sea wlopiAsed
suapeduy

eynsayy

Hssey3nop

Cp eurequag) endydip
(eiequyoR] eLejuaq) BeUaZIUOD
‘esoqing

(eq[Aydormoy eLequaq) weisnsue
sy1o}sed-esing

wapes

sueoiper

eqfoyrpumar
sopyojnoundes

BiO[jIpueIs “sea soploutrede
euvopoure

wanes

edie01Dput

SOprOyTOS

voeweupids

uinidas

waoeiapay

sinsnyed

‘snsosaqny

susaua

sinsnyed

eqAydorayoy

stnsnyed

snnpie)
sanpie)
epepie)
ouyurepre
oupuepIeD
‘ouyurepse)
ouyurepse)
ourwepred
aupuBprez)
aupwepies
oururepsed)
oujurepreD
eururepred)
ULUTEpIeD)
oururepred)
eypsded
syqeuues
sisdue,
eynuedured
Binuedue
enuedures
ejnuedure>
‘BUypaUIEZ
‘BuUypaUtEZ)
BisseureD)
BIdasATeD
wsasheD
eBayskte)
. BRD
uosodoyeD
SHOUTED
SySENTTeD
SySENTTTeD

SmnSwmYSnononocoet romeo come ntnoomannowoood

Ait

Appendix A Checklist of Vascular Plants


avaovuddsAD
aVaOVaddAD
avaovuddAD
aVeOVaadAD
aVaOVaddAD
aAVaOVaEdAD
avadovod
aVaOVOd
AVAOVNIOVUOE
AVAOVNIOVUOE
AVAOVNIOVUOE
avaovod
aVaOVIAV INHdOUOS
VEOVIGOdONSHD
aVaOV.LNOSND
aWaOvV.LAOSND
avagov.nosnd
aVaOV.LNOSND
aVaOVLAOSND
aVaOV.LNOSND
avaovVLNOSND
SVAOVYHLAT
AVEOVIdV
avaOvIddOHdnd
aVaOVIddOHdNa
AVAOVUaLSV
aVEOVASLSV
AVAOVUALSV
avaovsod
avaovsoud
avaovsod
avaovsod
aAVaOVSOU
avgovsod

STUN

stquainosa
sozmsonypAsa

smupueip

(strejnay “D) seapurediq
snyeurumnse

smyeuppa

snyeyspo

UWINUETUISIIA “eA UINUEIUTSITA
‘[Basog “FeA WNUETUTSIIA
areupyyo

woop

syeanut
umnpopidins
wimuouos{jod
(ansadures “2 “Jout) euosejuad
Waouass

umutfipyda

umuyyda

Asoo

rynuereydao
BUIISSISOOSTA
sysuapeueo
snudZoyjueuoul
snsojnpucys
WN.10}99}

waypind

syreydeo

‘Byuginoons

weound

esourd

eudgouout

stjjour

wyeOEUT

umssojzous>
uinssojgouXkD

Bpejequik)
ewojophy
enosnd
eynasnD
wnosny
eynasny
BynosnZ
einosnd
wynosnd
woydng
eruanordér

uo}0ID

sidaay
s\daa

sndoues)
sndowesy
sndavyer)
sngaeies
sndovyer)

RmoameooooomenwmoomnneggosQ{rooknwmran

w

**

* a

see ee

Appendix A Checklist of Vascular Plants

A20


Expressed mathematically,

where
I = floristic quality assessment index
R = sum of valuation coefficients for all plants recorded in the area
N = number of different native species recorded

According to Wilhelm and Ladd (1988), “by treating diversity as the square
root of N, increasing extremes of diversity are dampened to allow lower-
diversity, specialized and often small areas of very high mean quality to rate
favorably in relation to larger, often more diverse areas with lower overall
mean qualities.”

Table 1 provides an example of a floristic quality assessment index for two
Ohio peatlands. In addition to the presence of a Sphagnum-dominated mat,
these two areas have in common that no alien taxa were recorded from within
either study area. Flatiron Lake Bog contains 11 state-listed rare plants,
whereas Silica Sand Quarry Bog contains 4. Flatiron Lake Bog (Andreas and
Bryan 1990) is a low diversity, high quality natural area. The floristic quality
assessment index value for Flatiron Lake Bog is I = 37.53. The second area,
Silica Sand Quarry Bog, has developed on the floor of a sandstone quarry
within the past 80 years (Andreas and Host 1983). The floristic quality
assessment index value for Silica Sand Quarry Bog is I = 26.22. The differ-
ence in the floristic index values between the undisturbed Flatiron Lake Bog
and the disturbed Silica Sand Quarry Bog are probably a result of human
disturbance and is reflected in the numerical values between the two sites.

The range of floristic index values can vary depending upon the quality of
the species composition occurring in an area. For example, Wilhelm and
Ladd (1988) reported values for woodlands ranging from as low as 10 to as
high as 80 (or more). When they compared three sites within the Chicago
tegion, each about 1 acre! in size, the index value for an old field was I =
8.4, for a degraded prairie, I = 28, and for a high quality prairie, I = 50.

Assigned values for a particular species can differ between physiographic
regions. For example, when Wilhelm and Ladd’s species list for the old field
(I = 8.4) was subjected to the coefficient of conservatism values presented in
this study, the result is 1 = 10.2 (Table 2). The major difference in the val-
ues for the two areas is the coefficient of conservatism for Aster drummondii.

! To convert acres to square meters, multiply by 4,046.873.

Chapter 3 Application of Coefficient of Conservatism to Floristic Quality Assessment System


avadoOvsod
aVAOVOISSVad
aVaOVOISSvad
AVEOVOISSvad
avaovdva
aVaOVava
avaovdva
aveOviadondng
gvVaOVRIvInssow
avgaoOvrvInssowD
SVaOVNIVINSsOND
avaoOvMvINssowD
avaovRivinssowl
avVeOVMVINSsOUD
avaovmivinssoud
aveOvrvINssow
aVaOVasdAD
aVaOVaadAD
aVaOVesdAD
SVEOVIGUVOVNY
FVFOVICUVOWNY
AVEOVIGUVOVNV
AVEOVIGUVOVNV
AVEOVIGUVOVNY
aVaOvVOrda
FVFOVLIVWOLSV TSW
aVEOVNNVHS
SVaOVNNVHE
aVaEOVNNVHY
aVAOVNNVHS
avaovaasaa
AVEOVUALSV
aVeOVOISsvad
aVaOVOIssvud

epurlq

sysaay{s

stnsnyed

Qjeupyyo wnpinse_) unopenbe-umpanyseu
BSODSTA
roeovopnasd

epidsy

‘syunumu0s

asin

umapes

WIngeI0po

unyeury

‘BLaBiNsso1d
uinsojnpueys

neqsouto

wnueououre
wepaudes

vaoertides

eqre

eurydy

vagerd

euyjedoo

voneWOIe “JeA BONEMIOIE
eueuare “Jea kOneWOE
(winasoa “rea usnuoLIpau “y) wnyTAydound
BOTUIBIEA

wqejooouRT

Bnsuey

wopseyyed

erosure

Bjoayny

eyeuud

snapes
wnysusydes

seqra
wiodsoyoudyy,
eodsoyoudyy
wiodsoyoudyy,

sayy -

snug
soy
sn
snug
Uospuapopouy,
Bony
snuureyy
snuweyy
snumeny
snuueyy
Spesey
epiqnea,
snueydey,
snueydey

SCOrnrowmoowvummSronnSanonSoloonmoocooxncenw

semen tae @

*

*

Appendix A Checklist of Vascular Plants

A50


AVFOVOVUAIXVS
aVeaOVIaaGALNOd
AVAOVOISSVad
SVIOVIdV
FVEDVINONANVa
@VEOVINONONVE
AVSOVesdAD
VaOVITIT
aVeOVITII
AVAOVNIOVUOE
aVeOVUALSY
AVaOVaALSV
AVAOVUALSV
AVIOVUALSV
aVSOVYALSV
AVSOVASLSV
FVEOVaaLSV
AVEOVUALSV
aVaOVaALSV
aVeOVaELSV
aVSOVUALSY
AVAOVAALSV
AVAOVYALSV
AVAOVASLSV
AVSOVUALSV
AVAOVISIO
AVFOVISIO
AVAOVUALSV
FVaOVUELSV
aVeOVIGNY
avasoviana
avaoviana
avaoviana
avaoviana

euRopoure

wiqnp

syeuojeur

wayeuey

vueouoUre

eqornnse

eypuesopur
snjepoydse-orny

wang

umeedoma
soplomueray
stuoynery x

snsosoqny

snsoumns

siejopad

suyequap1900

‘stout

snyeydaoouoru
queyqurpeur

smasity

smeuasassord
snoquesid

snyeouealp

snqeradeaep

sunuue

‘osuapeues

HewyIq

umsonxay,

‘syeuunyne

(-d equoysnoqy) easndind
(cu eruojsnoqy) suesndyu
CL equoisnogy) erfoyy3uoy
(-9 Bruo}sno#y) sisuapeueo
(19 eruoysnogy) vanae]

BOyONAy
BioyUeIIOH
spedsoH
wngpBay
vonedapyy
voneday
eydreorwaHy
Spieooaaurayy
Speoor8uiegy
wunydonoyayy
sisdonoH
snmpuey op
SnIUeHoH
snmpuetay
smyueIPH
SsayUeHeH
snmpueroH
snmpueyiay
SmIpUEL AE
snmpUeH aH
snUUeY oF
snyueHoH
snUpUe HE
SnUpUR TOE
snuQUIPH
wnWEYUET OH
WnWoUET aE
unyuapa
uunyua[eH
snoxpoy
snodpay
snofpoyy
snodpayy
snodpoy

TOMTOM TOAHOYTNHOTNHOTHNONnNTNOSDSOMNHNTOVWYS

es)

shee

Appendix A Checklist of Vascular Plants

A30


aVeOVOd
AVaOVOd
HVAOVINIdTVSaVO
SVEOVINVT
AVAOVINOWIOd
avsovsou
aVeOvsou
avaovsow
avasovsou
aVeovsON

AVEOVINVUAD
SVEOVINVUED
AVSOVINVUED
AVEOVINVAAD
AVIOVNVLLNAD
AVEOVNVLLNAD
AVEDVNVILNED
AVEOVNVLLNAD
SVAFOVNVLLNED
AVIOVNVLLNAD
AVAOVNVLLNAD:
VSOVNVLLNAD
EVER) Orit
AVIOVYOVNO
HVAOVYDVNO:
AVEOVYDVNO
avaovorg
avasovonig
avaoviand

stjeasoq

woynnoe

Sompueseny

(4 eur099[5) Baoesapay
(a sysdowiody) eaqna
‘wnuerUTaatA

wausaa

yea

wnyerUoET

‘esuapeues

umoddaye

umaujngues:
wnuenseqor

winyysnd

ayjour

wngernseur

umyassip

wnuefuyose

Heuyo1q

(-d vuenuan) esaz0id

(9 euenuap) eo

(‘b euenuap) exjoyonbuinb
eueuodes

Bjuamnsagnd

(eqre ‘p) epraey

esneyo

Hsmarpue

Bye20eq

. eiogfared
(B20 y{Suoy |) praysyid “zea squua}q
Stuuarg “rea suuoIq
suaquinsoid

emnprdsry

waa

eHa0KID
epaok{
PISHPAID
BWOIPIH
SID
unap

wap
wep
wnag
wna
umyue.sas)
tunpueran
umyue.1as)
umyuesas)
uunjuesaQ
umnypuezas)
wnjueap
UUnTURII
sisdouenuer
sisdouenuap
Byfeuenue_
vuenuep
euenuap
buenuen
‘buenuap
Buenuag
Broessnjxe
wanes
eineg
enep,
pate
BpoTNeD
wnyesy

“~2e

SknnocorwemSSSanwatreotrocomomanavvoo

gwmm

Appendix A Checklist of Vascular Plants

A28


AVaOVINNVdNVO
aVeOVINNVdNVO
AVIOVINNVdNVD
AVAOVNIOVUOT
FVAOVNIOVUOE
aAVSOVNIOVAOE
aVeOVNIOVAOE
FVEOVNIOVYOR
AVeOVAIHOYO
AVEOVITONOVI
AVAOVGIHONO
VEOVGIHDYO
AVSOVNIT
aVAOVNIT
AVAOVNIT
SVAOVNIT
AVAOVNIT
AVAOVNIT
AVSOVITONdVO
AVEOVNIVINNdOWS
aVaOVENVT
aVaOVNIVINHdOYS
AVAOVRIVINHdOUS
AVAOVNIVINHdOUOS
aVaOVITII
AVaOVITIT
SVSOVITIT
AVaOVITIT
aVaOvVaATIO
aVaOVeIO

EVER) Ese)
AVEOVUALSV
AVAOVAELSV
AVEOVUaLSV

yuyey

egy

syreutpaeo
ayeupyyo
wayoyney
esuatuyjore>
‘SUSDSOUD
‘@SUIAIE
weps0d
wrayidym
Hleseo]
HOSEL
wunuepurgara,
WInWySssHEyysn
wnje[ns
umens
auuarad
unuexa) “sea WNIpous
stqeasoq

‘eiqnp

urozuaq
syredna
wopewyep

~ sisuapeued
wnquodns
umorydjepepryd
‘osuauediyoyur
asuopeugo
aredqna
winyfoytTe4o

wunyjogysnygo
‘esouenbs

Beards
‘BSOLIEIS

SCammoconnwmnwmoowrlvonwmonnawlSonanonna

Appendix A Checklist of Vascular Plants

A36


AVAOVOVUOTVH
AVEOVOVUOTVH
AVEOVOVUOTVH
AVIOVOVYAOTVH
aVeOVONAN
AVSOVNIOVUOE
SVAOVNIOVEOE
FVAOVNIOVUOE
AVAOVNIOVUOE
AVEOVNIOVUOE
AVAOVNIOVUOE
AVAOVNIDVUOE
aVaOVITIT
aVvaOVOd
aVaOVOd
aAVdOVOd
AVaIVOd
avaovod
aVaOVOd
avaovod
avaovod
aVeOVUON
aveOVUOW
avaovaOn
AVaDVdOULONOW
AVADVdONLONOW
AVEOVTIONAd
aVEOVINVI
AVEOVINVIT
aVaOVINVIT
AVEOVINVA?
AVaOVINVI
AVAOVOZIV
AVAOVOVAAIXVS

winjeqonsaa
umyeojds
(suaosaqyexe "Jy) WNOIIGIS
uinytAydorsyoy
eorueapésuad
BUIOA
wageayss:
sapjord.oos
(ByOEyS “A) BypUEZoFUE
Bxe]

40{09STp
‘sysuaare
sapjokn0q
‘eroyyinUa)
voneayés
wrayT]oqos
Nagaryos
eueorxour
weoWO[S
‘Bsopuoly
Byojpedse
eiqni

Basu

eqre

woyrun
shunidodéy
wioyiun
erpow x
wejound
Bsormsiy
eudpip
eipodouy]>
BEPPIAA
‘eqydip

uunytaydou ky,
wn dydop sy
wmnypAydousyy
wumnytAydouAy

BOL

snosokyW
sposokyy
Sposoky
Heosny]
Byszaquaryn,
BIss9quaTyN,
erdsoqua[yny
Bidsoquaryny
ByBroqueyynyA,
BIBoquaryN,
nBsoquayyNyL
BBraquaTUNy
STuO
sno
smo,
edonouo,
edonouow
sasouoj,
epreuoyy
epreuoy,
‘epreuoyy
epreuoyy,
‘epreuoyy
osnoW
INA

Bwos

RONRNRNERNOEDVOMANADOMODORNOOONRS

Appendix A Checklist of Vascular Plants

A40



aVaOVeadAD
aveovsou
avadovsou
VEOVOd
aVeOVOd
aVaOVOd
SVeEOVOd
aVeOVOd
aVeOVOVA
AVEOVNODA10d
AVIOVUALSV
SVAQVUALSV
avsoviauondng
AVaOVIGYOHdNA
aveOviduoHdng
aveoviaNondng
aVaOVIGyOHdNa
AvaOviauoHdng
avaoviddondng
avaoviadondng
aVaOviauoHana
avaoviadoHdng
aVaOVIddOHdNA
AVSDVIgNOHdng
AvaoviadoHdng
aVaOVIEdOHdNg
aVaOvVIadoOHdna
avaoviawoHdng
avaoOviawoHdna
AVaOVIEdOHdNg
AVEOVUELSV
AVAOVaRLSV
AVaOVUaLSV
SVEOVUALSV

stjeuwmnjne
Bpewyn
wiqny

(esmqo *) ere[TonFeAqns
equ
sysuajeid
‘BuyAo
(eaoeuypunie *3) s0nefa
eyoypued
umjuamnosa
{soprounradsouwAd o3epijog) wiowar
‘eyoyruuerd
‘ByEINITULIOA
suadias
Byexsoid
BrostuoSAjod
sopdyd (eid
snjdad
wresmgo
‘suginu
Byeupsseu
wemnoew
spidey
wyEoTey
Bnsa

Bye Quap
suyssyed {a
wsoydoyyesa
wreyjor09
‘Ryeynwwo0S
wmngoyrisses
wnunoses
uinsosn
wmaindind

sqAsuquiny
Bmpuadyy
emnpuadiEy
vONsay
vomsay
wonysay
eamysag
wonysay
sn3eq
umsXdoseq
Byung
erureyng,
eiquoydng
wiquoydng,
Biquoydng
eiquoydng,
Byquoydngy,
Biquoydng
eiquoydng,
eiquoydag
eiquoydng
eiquoydag,
Biquoydng
Biquoydng,
Biquoydng
Biquoydng
eiquoydng
Biqaoydng
viquoydng:
eiquoydng,
umpojedng
wnauojedng
unporedng
unyoredng

mrammntToooooocooowecOoLomnnnowcocoonnwod

Appendix A Checklist of Vascular Plants

A26


Corps of Engineers Research Report Summary, August 1995

Wetlands Vegetation

N

Floristic Index for Establishing Assessment Standards: A Case Study for

Northern Ohio (TR WRP-DE-8)

ISSUE:

The assemblage of plant species can indicate
various responses to environmental gradients
and disturbances. Information is needed about
the occurrence of species within natural and
disturbed plant communities for establishing
reference standards for use in the hydrogeomor-
phic approach used for evaluating wetland con-
ditions and natural places.

RESEARCH:

A floristic checklist was compiled for 31 coun-
ties in northern Ohio. Rankings of 1 to 10 were
assigned to native taxa based on their degree of
fidelity to a range of synecological parameters.
Plants found in a variety of plant communities,
including disturbed sites, were assigned rank-
ings of 1 to 3. Rankings of 4 to 6 were applied
to taxa that typically are associated with a spe-
cific plant comunity, but tolerate moderate dis-
turbance to that community. Rankings of 7 to 8
were applied to those taxa associated with a
plant community in an advanced successional
stage that has undergone minor disturbance.
Those plants with high degrees of fidelity to a
narrow range of synecological parameters were
assigned a value of 9 to 10.

SUMMARY:

The floristic quality index for 2,063 plant
species in northern Ohio provides a tool to
assess the quality of naturalness or presence
of conservative species. It allows for an ob-
jective numerical comparison of two or more
unrelated community types and reflects nu-
merically the impact of human disturbance by
taking into account the presence of alien taxa.
The ability to evaluate floristically and assign a
repeatable quantitative value has use in assess-
ing wetland restoration projects and in design-
ing and monitoring mitigation creations.

AVAILABILITY OF REPORT:

This report is available on Interlibrary Loan
Service from the U.S. Army Engineer Water- -
ways Experiment Station (WES) Library, 3909 -
Halls Ferry Road, Vicksburg, MS 39180-
6199, telephone (601) 634-2355.

To purchase a copy, call the National Technical
Information Service (NTIS) at (703) 487-4650.
For help in identifying a title for sale, call (703)
487-4780.

NTIS report numbers may also be requested
from the WES librarians.

Mr. Robert W. Lichvar is now at Cold Regions Research and Engineering

Laboratory (CRREL), phone: 603-646-4657.

(April 2001)
Please reproduce this page locally, as needed.


AVEOVTIAHdOLVaD
AVAOVTIAHAOANVO
AVAOVTIAHdOANVO
AVAOVTIAHAOAUVO
AVAOVTIAHdOAUVO
AVEOVTIAHAOAUVO
AVEOVTIAHdOAUVO

avaoviand
aVeOVINWTdd
AVAOVNVLLNED
SVSOVAALSV
aVEOVUALSV
AVAOVUALSV
AVaOVUELSV
AVSOVUELSV
AVaOVUELSV
aveOvOd
aVaOVNTN
avVaOvVW IN
AVEOVALSV TIO
SVEOVNNVHY
aVSOVNNVHS
avaovarsdsd
avaovargausd
AVAOVINONDIG
AVSOVINONDIG
AVAOVINONODIG
aVaOVNIV INHdOwOS
avadovovd
aveovaNv Ione
aVaOVGNVIONE
FVAOVGONV TONE
aVeOVaNV TONE
AVeOVONV IONE

uinssowop
(umuBjuoy *D) wiNgesnA
umMsoos{A

‘uInso}UsUIO}

‘sueinu

uInye.z2UL0[su09

OSUSATE

sifewuapisc0

snuryuu

wmnypaypynd

SHEPHS|OS

Bisqu

Bsomoeur

eaoef

eiqnp

snuet>

snutdsi3uoy

eyoprnuay

stfewapio00

suapuos

snaoeqiay

snugopoure

Soplonorfelpy “JeA Sapronoreypy
winaqued!s “sea saplonoyeyy
esopads

By8Ao

sapjojuousyq

‘BQUIDI00

‘Byejuap

BSO}UIWIO

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STTeAo

‘BSOLULOE]

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eka

MERNowvecwmocovwvvSmoamocooccooocoonnovooon

Appendix A Checklist of Vascular Plants

A16


AVeOVINONANVE
AVADVINONANVA
AVAOVINONANVE
AVEOVINONANVA
AVEOVINONANVE
AVEOVINONANVE
AVEOVINONANVA
AVFOVINONANVA
AVAOVINONANVA
AVEOVINONANVA
AVEOVINONANVE
AVEOVINONANVE
AVAOVINONANVa
VAOVINONNNVE
AVEOVINONANVA
AVEOVINONANVA
AVEOVINONANVaA
aVeOVOVsA
aVaOVOVA
aVaOVOVA
gavadovova
aVeOVOVA
gVeOVOVA
aVaOVOVA
aVeOVOVA
aVeOvVoOVA
aVESOVOVA
aVaOVOVd
avaovsou
avaovsou
aVeOVSOU

EVER) Ae):
AVeOVSON
AVEOVIONAd

snyEmnoysay
smyesapaos
suadar
smeamnoar
snoqueapAsuad
snypuezorur
sisonsuoy

(streuornuoidas -y) snpniu “sea snprdsty

snpidsty “sea snpydsty
sueyoqey

Bywoy

siemnorosey

snsoqing

suosiquie
Ssuaquoysore

supe

snantoge

eueay X

eunnyon

wiqny

snuud

sunsnyed
H8zoqueqyanu
edresosoeul
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40]091q

eqqe

(ejyund snjeyq) snyeur
‘sysusoy

(co snfeyy) eEeuOJ0D
syunuruzco

(ce snyeyy) eroynsnsue
epunses

snjnounuey
snjnounuey
snjnounuey
snjnounuey
snjnounuey
snqnounury
snnounuey
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snjnounuey
snjnounuey
snqnounuey
snynounuey
snjnounuey

SR omoom mM eMontTonnntTSONnmomoOENoROEMmMoNS
A49

es
x xe

Appendix A Checklist of Vascular Plants


Contents

Preface 2... 1 ete teens vi
1—Introduction 2... 6... eee 1
2—Methods .. 6... et eee 4
3—Application of Coefficient of Conservation to Floristic
Quality Assessment System .... 0.0... 0.0.00 eee eee eee 6

4—Conclusions 6.0... eee ee eee eee 9
References 2.0.0.0... eee ee ents 10 —
Tables 1-2
Appendix A: A Checklist of Vascular Plants for the Floristic

Quality Assessment for Northern Ohio .................000. Al

SF 298


aVEOVINVIT
AVAOVINVI
AVEOVIGOdONAHD
AVAOVOITVS
AVSOVOIIVS
AVAOVOITVS
VEOVOITVS
AVIOVOITIVS
AVAOVOITVS
AVAOVOITVS
SVAOVOITVS
AVEOVOITVS
AVSOVOITVS
AVSOVOrIVS
AVAOVOITVS
aVaOVOITVS
AVSOVOITVS
SVAOVOITVS
AVAOVOITVS
AVAOVOITVS
AVAOVOIIVS
AVEOVOITVS
SVAOVOITVS
AVAOVIGOdONaHD
AVSOVLVNSTIV
AVSOVLVWSTIV
AVAOVLVWSITV
AVEOVLVWSITV
AVAOVLVWSITV
AVEOVLVWSITV
AVSOVTIAHdOAYVO
AVEOVTIAHdOAUVO
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(9. sndreso0ydo) eurocteo
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Oe +

Rae *

Row

Appendix A Checklist of Vascular Plants

A52


AVAOVNODATOd
AVAOVNODATOd
AVADVNODATOd
AVFOVNOOAIOd
AVAOVNODAIOd
HVSOVNODATOd
AVAOVNODATOd
AVAOVNODATOd
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AVAOVUELSV
AVAOVUaLSV
aVaOVUALSV
aVAOVUELSV
avgovsod
avaovsou
avaovsou
avaovsod
avadovsod
avaOvsod
avaovsod
avgovsod
aVvaoOvsod
avgovsoa
SVAOVUHLAT
avaovsod
aVaovsod
aVaOvVSOu
avaOvsod
avaOvsou
avaoOvsou
avaOvsod
avaoOvsod
avaovsod
avaovsod

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smyETNOIqIO
Snyjoysnygo
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‘SHTEWAPIIO
SnyeUpe]
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(eowoureuurs “y) syefeur
Byajuesa

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euyueo

xoumny
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xouny,
xouUnYy,
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eee

eee

AS1

Appendix A Checklist of Vascular Plants


avaovod
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AVAOVINIdTVSaVO
AVaOVUaLSV
AVEOVUELSV
aVEOVUaLSV
AVaOVaELSV
aVaOVUALSV
AVSOVAaLSV
aVaOVUELSV
AVEOVUALSV
SVSOVTTANIOV TAS
SVIOVTTANIOV TAS
aVaOVINSSVaD
avasov1Inssvad
avaovINssvaD
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avaovod
AVEOVINVT
AVEOVINVT
aVEOVINVT
aVaOVINVT
AVEOVINVI
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AVeOVIaV INHdOUIS
SVaOVesdAD
aVaOVaadAD
aVaOVaRdAD
AVAOVTIAHdOAUVD
aVaOVeEdAD
aVeOVeadAD
avadoveddAD
AVaOVaadAD

Haqey

(Cw eisseD) eorpuepueur
(4 Bisse) edresaqay
syegna

snopeads

stsuoneyd

snqnazedned

‘smjeaogo

snyequis:

sname

snus{uoue

susodru

epode

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wmsojuaULes
(uinyydaja3 *g) winaandind
unas

ane

ayearaa

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BITOJLATED “TWA BSOAIOU
BOUL]

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(enogqorsda *S) erejnouopes
‘PoIpuEl eur
‘PyejOIOUR]

wenn
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(snuerysind “s) mprus

BETES
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Buus

opaueg
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Bue yang
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m ><

Appendix A Checklist of Vascular Plants

AS4


AVAOVSSOTOOIHdO
AVAOVSSOTOOIHdO
AVaOVNIOVUOE
AVAOVUELSV
aVaOVOLLEN
AVEOVINVT
aVEOVINVT
AVEOVUALSV
aVSOVUELSV
AVaOVUELSV
aVaOVaSLSV
AVEOVUELSV
AVaOVUELSV
AVIOVUALSV
aAVaOVaaLSV
aAVAOVUSLSV
AVAOVUELSV
avaOVINLad
aveOV INL
aVeOVINLIG
aVEOV INLAL
aVeOV INL
aVEOV INA
avadov Inada
AVAOVOISSVud
avaovarisagd
avaovanidadda
AVaOVaELSV
AVAOVNVLLNED
AVEOVOISSVud
AVEOVOISSVad
avaovava
avaovava
VAOVINIATVS

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winjsessip
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(q eiuopoyesayy) 1PID2q
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syuuazed
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wnyoknog
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suopig
suapid
supig
suepig
suapid
suapig
suepig
suapig
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visndeg
syozy

SCwmasowvscoocornoowrfSomEammannanweanttt aeons

AQ

Appendix A Checklist of Vascular Plants


Table 2

Index Values for Plants in an Old Field in Chicago Region Using
|| Coefficient of Conservatism from Wilhelm and Ladd (1988) and
| Present Study

Taxon

Acalypha rhomboidea

Achillea millefolium

Agrostis alba (= A. gigantea)

Ambrosia artemisiifolia

ll Asclepias syriace

Aster pilosus

Aster drummondii

Barbarea vulgaris

Carex laxiflora

Chrysanthemum leucanthemum

Cichorium intybus

Cirsium arvense

Cirsium vulgare

Crataegus mollis

Danthonia spicata

Daucus carota

Festuca elatior

Fragaria virginiana

Geum canadense

Geum laciniatum

Lonicera maackii

Medicago lupulina

w lo Jo |x [x [x Jo Jo Jw Jo Jw Jo Jo Jo Jo Jw Jo Ja

Panicum implicatum {= P. languinosum)

Parthenocissus inserta (= P. vitace

| R = Sum of valuation coefficients for all plants recorded in the area.
N = Number of different native species recorded.
| = Floristic quality assessment index.
‘ Wilhelm and Ladd did not assign values for alien taxa.
2 Considered an alien taxon in O!



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AVEOVNVISVOOddIH
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AVeOVeaOV
AVEOVeEOV
aVaOVEROV
AVeOVeaOV
aVEOVAROV
aVaOVUsOV
aVaOVUEOV
avaoviduoudna
aVeOVIaNOHdNA
AVeOvVIddOHdNa
AVEOVATVN

ATINVA

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snmosay
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SQNZD 9409 SLNAWNOD

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wm

*

Appendix A Checklist of Vascular Plan


Wetlands Research Program Technical Report WRP-DE-8
. August 1995

Floristic Index for Establishing
Assessment Standards: A Case
Study for Northern Ohio

by Barbara K. Andreas

Cuyahoga Community College
Division of Natural Sciences
Cleveland,OH 44122

and

Department of Biological Sciences
Kent State University

Kent,OH 44242

Robert W. Lichvar

U.S. Army Corps of Engineers
Waterways Experiment Station
3909 Halis Ferry Road
Vicksburg, MS 39180-6199

Final report
Approved for public release; distribution is unlimited

Prepared for U.S. Army Corps of Engineers
Washington, DC 20314-1000


aVdOVOd

aVaOVOd
AVOVOISSVa
aVaOVUSLSV
aVSOVaALSV
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avaovxnd
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aVaOVMV INHdOWOS
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avadovod
avdovod

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avaOvOd

avaovod

avaOvod
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aVaOVOd
AVAOVSSOTOOIHdO
HVADVSSOTOOIHdO
AVFOVSSOTOOINdO
AVAOVSSOTOOIHdO
AVAOVSSOTOOIHdO

(esuedxaut ‘D) wioins
sisuapeueo
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snuloig
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snuloig
snwoig
snuioig,
snwioig.
snwiosg:
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se

“eee eee

ee

Appendix A Checklist of Vascular Plants

A10


12

Sears, P. B. (1926). “The natural vegetation of Ohio. II. The prairies,”
Ohio J. Sci 26, 128-146.

Smith, R. D. “An approach for assessing wetland functions using hydrogeo-
morphic classification, reference wetlands, and functional indices,”
Technical Report WRP in preparation, U.S. Army Engineer Waterways
Experiment Station, Vicksburg, MS.

Soule, M. E. (1990). “The onslaught of alien species, and other challenges in
the coming decade,” Conservation Biology 4, 233-239.

Swink, F., and Wilhelm, G. (1979). Plants of the Chicago region. Morton
Arboretum, Lisle, IL.

. (1994). Plants of the Chicago region. Indiana Academy of
Science, Indianapolis, IN.

Transeau, E. N. (1935). “The prairie peninsula,” Ecology 16, 423-437.

Wilhelm, G., and Ladd, D. (1988). “Natural areas assessment in the Chicago
region,” Trans. 53rd N.A. Wildl. and Nat. Res. Conf. 361-375.

References


avadovasddAD
AVEOVUAdAD
avaovdddAD
aVEOVUEdAD
aVEOVUEdAD
aVaOVasdAD
aVaOvaadAD
AVAOVaEdAD
aVaOVITIT
aVaOVOd
avVaovOd
aVaOVIIEZHONEHOS
AVAOVOVAAIXVS
AVAOVOVAAIXVS
avadovananvs
SVaOVINVT
aVEOVINVIT
AVAOVINVI
avaovanvt
AVAOVINIOVUAVS
FVIOVTIAHAOAAVOD
AVAOVIdV
aVaOvidv
aVeOVIdV
aVeOVIdV
avaovsou
aAVaOVaaA Vdd
aVaOV TINWId
aVeOVITOURIdVO
AVeOVITONNIdVO
AVSOVINVT
aVEOVINVT
aVaOVINVT
aVaOVINVI

snAyddjod

‘snjnpued

symerany

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suaoseindind

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sisuopUTgatA

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sanuzo

(a wnpodourfD) suedyna
sysuay10y

(euesueyze *S) erjoynsndue “sea eyjoqels
wnpiqre

vaindind

streupyyo

wrefoyin

eorpueyieur

euesad

sisuopeue>

sisuopeued

sisuapeuvo

(stuoyazed *s) snpunquoy
(suagnd ‘g) esowaoes
sisuapeueo

wquedns x

exayjar

sysuayed

syeupyyo

SIPS
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eyes

SCOTOMENHHNeHYYNNHOSYMOMrOVMASSoonnananodr

eeee

Appendix A Checklist of Vascular Plants


AVIOVATVAN
AVIOVATVA
AVEOVGIHOYO
aAVeEOVITIT
AVAOVITONOVA
aveovuOW
AVSOVAHLAT
AVAOVUHLAT
AVAOVAHLAT
aVeOV INN
aVeOV INN
AVEOV INN
aVaoOvINNNd
aveOVINWNNd
aVeOV INN
aVaOV'INWTd
aVaOV INN
AVaOVINWId
aVaOV INNTd
SVEOVITII
AVeOVINVI
AVEOVINVI
AVEOVINVT
aVaEOVINVT
AVEOVINVI
SVSOVINVIT
AVAOVIGOdODAT
AVFOVIGOdODAT
AVFOVIGOdODAT
AVFOVIGOdODAT
AVEOVIGOdODAT
AVAOVIGOdODAT
HVAOVIdGOdODAT
AVEDVIGOdODAT

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‘wyeypsoul
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‘snopurauya
stuoyiun
snqjequ
snasdoma
dedse
snueoproure
Haraqey x
uindyoeisiy
uinyrydosod
ummosqo
uunjprony
umyepunut
(euOsTTEqeYy “T) wINAENSIp
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BARA,
Bae
SIXEE,
uNWaIUEEAL
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wna hy
wna kt
wun}
eqyoeuys<]
Brpeuyshy
eqyoeunysé]
eqyoeunisé]
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‘Byypeurshy
‘eyypeunysiy
eqyoeunséy
Byyoeunsé]
spooky
sndook]
sndookq
sndook]
sndook]
sndook]
sndook]
wintpodooky
‘uwntpodoss]
uuntpodook']
winrpodoak]
umrpodoosy
unrpodook]
umrpodook]
uinypodooky

MMADMDNAOMMOSOONMNTOTMOSCHMNOYVONMROSCORRwAOOS

Appendix A Checklist of Vascular Plants

A38


In an attempt to make more objective evaluations and assessments of open
land areas, Wilhelm (Swink and Wilhelm 1979) and Wilhelm and Ladd (1988)
devised an index of conservatism, a component of their Natural Area Assess-
ment. Their evaluation is based on the fundamental character of the native
flora of a region. A numerical quality rating, called the coefficient of conser-
vatism, is assigned to each plant. Each numerical value is an expression of
the taxon’s autecological value with respect to all other taxa in the flora. The
higher the numerical rating, the more conservative is the taxon. Species
conservatism reflects the ecological specializations that a plant displays to a
specific habitat or set of environmental conditions. The natural quality of an
area is reflected by its richness in conservative species.

The coefficient of conservatism is independent of frequency. A plant may
be widely distributed in Ohio, but occur in only a limited number of habitats.
Viburnum acerifolium, primarily found in rich mesic forests, is an example of
this situation. Conversely, a plant species may be somewhat uncommon, but
occur in various habitats throughout the study range. Habenaria flava var.
herbiola, which grows in wet woods, fens, weedy fields, and margins of
pools, is an example. Both species have a value of 6 (Appendix A).

Chapter 1 Introduction



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(sraurogzount “y) steazoq
snyeuyWNe
souewoyouy
CA stuosoidwreg) snyjAydoziys
uomauhred
winpyneuuid
wnugjuOul
steupyyo
eqonn
BIOUIPHIA
wqeyONIOA
eyesouen
esoxoqy
mueaqyas
eoenks
eyjoyupenb
suaosesndind
wqeUseoUT
Upeury
wyeyexo
syineorxoidure
‘esuapeurs
voquesys
sngiorp

syre3 [na

A7

CHOSORRMmaNAROALDASORLCSMmaNnMmMDSAONNALNHDON

Be 2228928222, 855258
HU

Appendix A Checklist of Vascular Plants


AVSOVNOOATOd
SVEOVNODATOd
AVAOVNODA10d
AVSOVNODATOd
AVSOVNODATOd
AVAOVNODATOd
SVAOVITIT
AVSOVITIT
AVAOVTIVOATOd
AVEOVTIVDATOd
AVAOVTIVOATOd
AVEO TIVOATOd
AVAOVIVOAT0d
AVAOVTVOAI0d
AVAOVINOWS10d
AVAOVINOWA10d
aVaOVaVddVO
aVdOVaVddVO
AVEOVGIHOUO
AVEOVWELSOdOd
avaovahiseusd
aVSOVOd
aVadOVvOd
aVvaOvOd
avaovod
aVaOVOd
avaOvOd
avadovod
avaovod
aVdOVOd
avaovod
avdovod
aVaOVOd
AVFOVNVLV Td

umsoyidsaeo
‘arBNIAS
wnyoyuE
wnyseuare
wniqrydure
winazoIpe
‘suaosaqnd
(wNyEINUIWIOD “g “[OUT) WNOYIG
wreponsen
e8auas
vourndues
euresfjod
eqjogtoned
we}
suejdaa
umajniee
CfenereispD) nsawef
Bapuesepop
soplossojgorydo
uiny[Aydoyes99
weed
syria
sisoayhs
‘stsuony[es
ssuaqeid
sinsnyed
syesowau
epmnguey
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essaadui0a
Bsoqing
enuue
‘soposye
‘stfewapts00

wmnuos{jog
winuossjog
winuo8{jog
wmuo3fjog
umuosAjog
wmuosfjog
wmyeuosAjog
umjeuosAjod
Resco
vpesttod
Byesjod
vpesctod

EEZSSSE22828

5
i

mYroCOoOoCKnwmoNnowmonNneSSmnocowwmaServy~nnonortos

A4S

Appendix A Checklist of Vascular Plants


avaovsou

avaovsou

aVaOvsou

aVaOVSOU

aVeOvsou
avaOvsod
aveOvsou

avaovsor

aVaOVSOU

avgovsoud

aVaOVSOu

avaovsoa
AVAOVNOLADOWVLOd
FVFOVNOLYDONVLOd
AVFOVNOLADOWV.LOd
AVAIVNOLADOWV.LOd
AVIOVNOLADOWV.LOd
AVEOVNOLADONV.LOd
FVIJOVNOLADOWV.LOd
AVAOVNOLAIDOWV.LOd
AVADVNOLADOWNV.LOd
AVIJOVNOLADOWV.LOd
FVAOVNOLADOWV.LOd
AVAOVNOLADOWNV.LOd
AVIOVNOLADOWV.LOd
AVAOVNOLADOWV.LOd
aVFOVNOLEADOWV.LOd
AVEOVNOLADOWV.LOd
AVEOVNOLADOWV.LOd
AVAOVNOLADONV.LOd
AVFOVNOLADOWVLOd
AVADVNOLEDOWV.LOd
AVAOVNOLADOWV.LOd
AVEOVOVINLAOd

suejdar
Byal
exopered
sinsnyed
‘eordaAlou
BypaULIajuy
ByeuPUy
Rsoonnuy
sisuapeugo
ange
‘Baquadie
BuLosue
SHULIOJTOISOZ
1hasea,
snyjoynomns
soqrunds
surqqoa
quospreyons
(Hproiyoreq “d “Tout) snyyisnd
yeyomnd
snZuojoeid
smeunood
snsopou
sueyeu
sISUOOUTTTE
a4
snouyweld
Hsouy
SnsOToy
stupfyida
SNYTOFISIOAID
sndspo
snyjosrdure
Ba0819/0

BIpUAI0
BIpUI0g
eynuaed
eynuaod
enue
BIpUaod
Byipuajod
eyAuaiog
eynuaiog
eymnusiog
BUA
eynuaiog
uoyedourej0g
uoladourelog
uojedourelog
uojSourej0g
uoradoureiog
uojesourri0g
uojadourni0g
uoyadourei0g
uojsdourRj0g
uojadourejog
uoyaSoureiog
uordourejog
uojaSourrj0g
uojadoureiog
uojadourej0g
uoiadourniog
uoladourgjog
uoyasoure10g
uoyedourRjog
uojsadoursjog
uoladourrj0g
soumnyz0g

ecaomBoonS Soo

+ beee

em

WK

me

aw

A47

Appendix A Checklist of Vascular Plants


AVIOVIONAd
aVaOVTIOUAd
aVEOVTOUAd
AVaOVINVIT
AVEOVINVIT
aVEOVINVIT
SVEOVINVI
aVeOVINVI
aVeOVOd
aVdOVOd
AVSOVILGAV.ISNNAG
aVeOVINA
avaovava
avaOvsod
avasoOvsod
avaOvsod
avaovsod
avaovsow
avdoOvsod
aVaOvVsON
avaovsou
avaovsod
avgsovsod
avadovsod
avaovsou
AVSOVINVI
AVAOVOVUOTVH
AVaOVITVdad
aVeOVaALSV
AVEOVAELSV
AVAOVUELSV
AVEOVUaLSV
AVHSOVUALSVY
avaOvsod

‘elfoyrpunyos
vondyye
‘equRIo[yo
wuNTeETUISITA
uinsopid sea wiNye| TINA
winyoyrnuay
wnonnw

wnueouy

epured

‘suBysIp

umuypmbe
weyOSN
soprorfexosd
‘eueyUyasta
‘BS0]UaUIO}
eunoses
geueyonbsns “sea epuind
eyruind “sea epund
woysuad
eorueapAsuad
edi

qareyeut

snsei30

ume

‘eueououre
suesna,

sunsnyed
‘BUBySMO]
‘esoulgo8
voutpidass

wodse

BUSSE

eq

xajduns

man Swmovoenscocomev oS SmonwomornramamnQon

ee

Appendix A Checklist of Vascular Plants

A48


avaov INWidd
aVaOVOd
avaovod

EVER Oc
AVAOVNIDVAO
AVAOVNIOVUO
AVEOVAALSV
aVaOVINWIad
avaoOvavs
AVEOVLIA
AVEOVAVIdaTOSV
aVaOVavA
aVaOVOd
AVSOVEHLAT
aVaOVSOU
aVaOVSOU
aVaOVSON
aVaOvsod

AVEOVOISSVad
AVAOVATVN
aVaOVOd
aVaOVOd
avdovod
aVaOV INLad

stfeiuapt920
snoqurguta

preso8

epAydooneys

BaINZze

(8 sysdoo47J) sysuaare
Baowyesew

ssuaAre

weaowq
eyepnounpadyaaiq
(eaae] wnyoueudD) snpiqre
esoonny

weyers paar

wsnqor

wreoids

eourngues

StAE]

Baroque

epyin

ehypessoyisd
eyoysTMoLe
smyemozeqny
(snupsjreures “y) sypnt
snxayoxyar

snppiqhy

(snpyaq “y) wnt
‘SPIO

snqre

sapyossdye

syeupyyo

sysuayead

snuequyores

sijenboe

wepnusas

wnoocoocoosoooooooonfwmuwweSanamonovooSuomsS

Appendix A Checklist of Vascular Plants

A4


AVIOVAALSV
aVSOVUALSY
AVEOVORIVAVL
AVEOVIdV
aVaOvVaTO
avdovav
AVEOVNIOVUO
AVFOVNIOVUOE
AVAOVITOddvO
aVaOVITONNdvO
AVEOVITONdVD
aVeOvVIIOAnNdvo
AVAOVIGOdONAHO
AVAOVAAA Vdd
avaovava
AVSOVITIT
AVdOVOd
AVEOVITIT
SVEOVTIAHdOANVD
AVSOVTIAHdOAUVO
AVEOVTIAHdOANVD
AVFOVTIAHdOAYVD
AVIOVTIAHdOAUVO
SVAOVATAHdV.LS
AVEOVINVT
AVEOVINVT
SVEOVINVI
AVEOVINVT
AVEOVINVT
aVdOVOd

aVaOVOd

aVaOVOd

avaovod
AVaOVNWGT

uInjedtAaey
asesina
Boqes
eULasaUT
sida
snpneoy
ayeupyyo
wmnuadse
smejnoiqio
syequapps0
snawspaauy “eA suqye
snqje “sea snqye
SpuLoys]0a9]89
uinyAydip
Bjoapay

snasox

voueds
winourweld
esaqnd

eppaut
eyopzuoy
vaurwesd

(7B UOJOSOAIA) BOHENbE
ByOyLN
eyopnua
stnsnyed
woqueuLiad
(trent g) erepio9
wradse
‘stuoyursea
snyo9j8ou
stupumdézs
Jodse
ezqyscjod

wngexeI8 |,
umyjaoeuey,
xpeurel,
viprusey,
edupitg
sndsesojduiks
umptydurkg
wing fydurkg
sodreopoyduikg
sodzeapoydurtg
sodagoproydurfg
sodrespoyduikg
epeeng
umoydoyAig
saffasoydong
sndoydang
edng
winpueUag
PHETAS
BpEras
eueyag
Byers
eels
eaAydeig
skyoug
sdyoug
shipeyg
skyong
skipeig
snjoqouods
‘snjoqorods
snjoqorods
snjoqoiods
eleponds

mmMoemMNoOnroetwCocononalSmoecSoorocwvowccnse

Mee

Rea

Appendix A Checklist of Vascular Plants

ASS


This report should be cited as follows:

Andreas, B. K., and Lichvar, R. W. (1995). “Floristic index for
establishing assessment standards: A case study for northem
Ohio,” Technical Report WRP-DE-8, U.S. Army Engineer
Waterways Experiment Station, Vicksburg, MS.

The contents of this report are not to be used for advertising, publication,
or promotional purposes. Citation of trade names does not constitute an
official endorsement or approval of the use of such commercial products.

vii


vi

Preface

The work described in this report was authorized by Headquarters,
U.S. Army Corps of Engineers (HQUSACE), as part of the Wetlands Evalua-
tion Task Area of the Wetlands Research Program (WRP). The work was
performed under Work Unit 32755, for which Mr. Dan Smith was the Tech-
nical Manager. Mr. Sam Collinson (CECW-OR) was the WRP Technical
Monitor for this work.

Mr. Dave Mathis (CERD-C) was the WRP Coordinator at the Directorate
of Research and Development, HQUSACE; Dr. William L. Klesch
(CECW-PO) served as the WRP Technical Monitor’s Representative;

Dr. Russell F. Theriot, Environmental Laboratory (EL), U.S. Army Engineer
Waterways Experiment Station (WES), was the Wetlands Program Manager.
Mr. Ellis J. Clairain, Jr., EL, WES, was the Task Area Manager.

The work was performed at Cuyahoga Community College and Kent State
University, OH, by Dr. Barbara K. Andreas and at WES by Mr. Robert W.
Lichvar, Wetlands Branch (WB), Ecological Research Division (ERD), EL.
The preparation of the report was under the direct supervision of Mr. E. Carl
Brown, Chief, WB; Dr. Conrad J. Kirby, Chief, ERD; and Dr. John W.
Keeley, Director, EL.

Grateful appreciation is extended to Mr. Aaron R. Andreas, Mr. Gary R.
Bryan, Ms. Kim D. Herman, and Mr. Jeffrey D. Knoop for their assistance in
the preparation of the manuscript. Special thanks are extended to Dr. Gerould
Wilhelm for giving much advice and leadership in the development of this
project.

At the time of publication of this report, Director of WES was
Dr. Robert W. Whalin. Commander was COL Bruce K. Howard, EN.


Floristic Quality Assessment for Two Peatlands in Portage
jeune. Ohio

Coefficient of

Coefficient of
Conservation

re

jlica Sand Quarry Bog

a Ls — —J
H Flatiron Lake Bog Si

Acer rubrum

Aronia melanocarpa

Amelanchier arborea

Betula alleghaniensis

Andropogon virginicus

Bidens coronata

Aronia melanocarpa

Calla palustris

Bartonia virginica

Carex atlantica var. capillacea

Betula populifolia

Carex canescens

Carex canescens

Carex trisperma

Carex lacustris

Cephalenthus occidentalis

Danthonia spicata

Chamaedaphne calyculata

Drosera rotundifolia

Decodon verticillatus

Gaylussacia baccata

Drosera rotundifolia

aia [Jo la fo lo lola fo la

Juncus canadensis

Dulichium arundinaceum

Juncus effusus

Gaylussacia baccata

Leersia oryzoides

Glyceria striata

Lycopodium clavatum

Hex verticillata

Lycopodium inundatum

Juncus effusus

Lycopodium
tristachyum

Larix laricina

Nyssa sylvatica

Leersia oryzoides

Populus grandidentata

Lycopus virginicus

Populus tremuloides

Nemopanthus mucronatus

Prunus pensylvanica

Nyssa sylvatica

Quercus palustris

Osmunda cinnamomea

Scirpus cyperinus

Polygonum arifolium

Spiraea tomentosa

Rhynchospora alba

Note:

Rubus hispidus var. obovalis

R = Sum of valuation coefficients for all plants recorded in the area.
N = Number of different native species recorded.

| = Floristic quality assessment index.



by focusing on large tracts of contiguous forest types. Forsyth (1970) corre-
lated Gordon’s vegetation types to edaphic factors such as the availability of
moisture, parent geologic material, topography, and direction of slope.
Forsyth found that the distribution of these vegetation types, or plant commu-
nities, is predictable on the basis of climate, geology, and topography.

Through time, native taxa adapted to a specific set of biotic and abiotic
factors of natural disturbance such as the local extremes of drought, inunda-
tion, fires, storms, and faunal interactions (Wilhelm and Ladd 1988; Hobbs
and Huenneke 1992). Because of periodic natural disturbances, a vegetation
seldom maintains a constant species composition for more than a few centuries
(Noss 1985).

The arrival of European settlers had a profound and permanent effect on
the native landscape by changing its physical character (clearing, plowing, and
draining) and by the introduction, both deliberate and unwittingly, of alien
taxa, creating what Pielou (1979) has called “man-made disjunctions.” The
terms “alien,” “non-native,” and “exotic” are used to refer to taxa believed to
have been introduced into the flora either with or after the arrival of European
settlers. A “native” taxon is one that has maintained historical integrity and
ecological processes since some time prior to European settlement (Maser
1990).

The native plant communities observed by the early surveyors and
explorers now include a large number of non-native (alien) taxa. Cooperrider
(1982) estimated that approximately one-third of the Ohio flora is composed
of these alien (mostly Eurasian) species. By contrast, the Hawaiian Islands
(one-sixth the size of Ohio) may have as many as 4,600 species of exotic
plants, which is about three times the number of native plant species (Soule
1990). The flood of exotic species, along with anthropogenic disturbances,
has tended to make more uniform natural landscapes by providing an opportu-
nity for alien taxa to replace native plant species. With the abundance of alien
taxa, natural places (natural areas) with intact native floras are becoming
rarer.

The surviving undisturbed natural areas dominated by native flora, or those
containing remnants of rare plant communities, are often sought out as special
places or significant natural areas. To date, there is no adequate way to pro-
vide meaningful comparisons of the flora of the different types of plant com-
munities found in these natural places. However, field biologists frequently
are asked to evaluate their quality. Herrick (1974), with the help of numerous
individuals, compiled preliminary data on 580 Ohio natural areas. In the
early 1980s, the Ohio Chapter of The Nature Conservancy, with the help of
regional experts, organized a list (scorecard) of the 100 best natural areas
remaining in Ohio. Assessing the ecological value of these areas was done
visually with the only criterion often being the presence of rare or unusual
plant species.

Chapter 1 Introduction


AVEOVUALSV
aVaOVITIT
AVFOVNId
AVEOVINONNNVa
avdovod
avaovod
SVeOVAIHOYO
aVeOVIIOddvo
aVeOVIIOIdvo
aVaOVINNVdNVD
aVaOVITIT
AVAOVITIT
SVIOVITIT
AVAOVITIT
aVaOVITIT
aVaOVIIT
AVEOVNIOVONNE
AVAOVNIOVONNL
avaovavs
avaovava
aVaOVEVA
avaovava
avaovdva
avaovava

EVER EAL!
avadovavd
aVaOvavd
aVaOV TINATd
avdovod
AVEOVINVT
SVEOVINVT
AVAOVTIAHAODAZ
aVEOVISNTD
aVaOVISNTD

BIB IBY
Bprouses
sisuapeue
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unnusad
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(areaury “L) winaoejas
wauio}oyoIp
‘SHSeL19}

(a wmnapadéyy) unoquyaita
(J unowad4yy) poseay

odeyjssny,
edyny
eansy,
sayjoay,
wmpHL
syseduy,
wouduL
wnaIsouL
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snnquy,
uunuapenL,
uunuapenyy,

WEHOWDANASTDSOSOSOSCSOCHOADALSr rer amNnnaAnoOwHDOS

*

a

Appendix A Checklist of Vascular Plants

A60


AVAOVIOIA
AVSOVIOIA
AVAOVIOIA
AVAOVIOIA
AVAOVIOIA
AVEOVIOIA
AVAOVTIOIA
AVAOVIOIA
AVSOVIOIA
AVEOVIOIA
AVEOVIOLA
AVEOVIOIA
AVAOVIOIA
AVSOVIOIA
SVAOVIOIA
AVEOVIOIA
AVAOVIOIA
AVEOVAVIddTOSV
SVSJOVNAOOdV
avaovdvd
avaovdvd
avgovavd
avaovavs

EVER EAE
avaovavs
avaovavd
AVaOVITONdVO
AVEOVITOSRIdVD
aVaOVIIONdvO
AVEOVITONadVO
AVeaOVIIOsadvO
aVaOvITOdNdvo
aVaOviIosdddvo
AVaOVIIOARIdVD

BqOsIpunjOI

wensor

mnbsouyer

suaosoqnd

eyosyqurd

wrepod

(eqoyin “A “[our) erewed
eye10po

eAydosydou

suajred sea 1Xaysopoeut
BqE|OIOUE]

eyeisey

ereynond

‘esiadsu0o

stsuapeues

(enugoouy “A “fouy) epuelq
sjsuaare

(u umypueud)) und
Jouju

esojta

BApEs

eynsayy

82081)

euerurjore>

Byoynsnsue

eueonoure

umueimbsouryes “sea wnuembsouryes

ouyye “sea unueInbsoures
uunyjoyyunud

snyndo ‘zea snjndo
umueopoure “sea snjndo

(saproursseo “A) sapfoursseo “zea umpaut

o8eua]
suejuEy

EOLA
BIOL,
IOLA,
RIOLA
BIOLA
PIOLA
RIOIA
SIOIA
PIOLA
BIOA
POA
POA
ROI,
BIOLA,
BIOL
BOI,
SIOIA
‘WINIKOJIIUT A

ePIA
SPIA
ePIA
BPIA
PDIA
SpIA
PIA
‘WINUINGIA,
‘UInUINgLA
wnWNgiA,
umuIngyA
umnumngiA,
umuingtA
wWnWINgrA
WIMNUING]A

SCenwmonwmmaNneOnsDooCoOSooONnNenwmawQownnnnwonw

eee ee ee

*

A63

Appendix A Checklist of Vascular Plants



Contents

Preface 2... 1 ete teens vi
1—Introduction 2... 6... eee 1
2—Methods .. 6... et eee 4
3—Application of Coefficient of Conservation to Floristic
Quality Assessment System .... 0.0... 0.0.00 eee eee eee 6

4—Conclusions 6.0... eee ee eee eee 9
References 2.0.0.0... eee ee ents 10 —
Tables 1-2
Appendix A: A Checklist of Vascular Plants for the Floristic

Quality Assessment for Northern Ohio .................000. Al

SF 298


US Army Corps
of Engineers

Waterways Experiment
Station

Wetlands Research Program Technical Report WRP-DE-8

Floristic Index for Establishing Assessment Standards:
A Case Study for Northern Ohio

by Barbara K. Andreas, Robert W. Lichvar

August 1995 — Final Report 7

Approved For Public Release; Distribution Is Unlimited


AVAOVITIT
aVaOVIdV
avaovod
avaovod
avaoviny
AVAOVITTSHOINNVZ
AVAOVAVOV
AVAOVGRIAX
AVAOVGIAX
AVAOVUALSV
AVEOVUaLSV
AVSOVNHOSTE
AVSOVNHO1E
AVHOVINaTSV
AVAOVNNET
AVAOVNINGT
AVSOVNWET
AVAOVNNGT
AVHOVSOU
avaovod
AVIOVINVIGV
AVHOVLIA
AVAOVLIA
AVOVLIA
AVAOVIIA
AVAOVIOIA
AVAOVIOIA
AVEOVIOIA
AVAOVIOIA
AVaOVIOIA
AVOVIOIA

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paine

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(eqnansayy “A) BSOTTIA
4010913

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(eImepquiy “A Jour) eeNIses

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wnnowovatranSnnwelSovwmnwmncocolwownowns

*

ee Whe

Appendix A Checklist of Vascular Plants

AG4


aVaOV INN
aVSOVITIT
aVaOVAV TAWAHL
aVeOVOVSdIG
aVEOVOVSdIG
aVeOVOVSdId
aVaOvoIssvud
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aVaovOd

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avaovava
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winyedyaaxy,
‘asua0Urt
umsounnys
umepidsno
(wunsmigo *q) area
susosourd
asuapeueo

mneoewnFantowmveccomnrnnweocscascormocosc orange

ee eee

Appendix A Checklist of Vascular Plants


of natural areas, often creating extensive monocultures (for example, Phrag-
mites australis). A ranking of 0 also was assigned to those native taxa that are
typically part of a ruderal community (for example, Ambrosia artemisiifolia).

Rankings of 1 to 10 were assigned to native taxa based on their degree of
fidelity to a range of synecological parameters. Plants found in a variety of
plant communities, including disturbed sites, were assigned rankings of 1 to 3.
Rankings of 4 to 6 were applied to taxa that typically are associated with a
specific plant community, but tolerate moderate disturbance to that community.
Rankings of 7 to 8 were applied to those taxa associated with a plant commu-
nity in an advanced successional stage that has undergone minor disturbance.
Those plants with high degrees of fidelity to a narrow range of synecological
parameters were assigned a value of 9 to 10.

All alien (non-native) taxa were assigned the value of 0. These plants are
preceded with an asterisk (*) in the “Comments” column on the checklist, and
their scientific name is printed in bold type.

Plants listed as “threatened,” “endangered,” or “extirpated” in the Ohio rare
plant list (Division of Natural Areas and Preserves 1992) are noted in the
“Comments” column on the checklist (Appendix A). While Ohio’s rare plant
list is updated every 2 years and the status of a taxon may change with the
discovery of new sites, the majority of the “rare” taxa are inherently a rare part
of the Ohio flora and generally have coefficient of conservatism rankings of
7-10.

Some taxa on the checklist are preceded by a double asterisk (**) in the
“Comments” column. These plants fall into the following conditions: (a) taxa
considered to be native in another region of Ohio, but adventive or naturalized
within the study area (Aralia spinosa, Campsis radicans, Cercis canadensis,
Gymnocladus dioica, Hydrangea arborescens, llex opaca, Napaea dioica,
Robinia pseudoacacia, Sagina decumbens, Thuja occidentalis), and (b) taxa
that include both native and non-native populations within the study area
(Physostegia virginiana, Pinus strobus, Prunella vulgaris). For the latter
group, the coefficient of conservatism ranking is based on native populations.

Rarely encountered interspecific hybrids, as included in Andreas (1989),

Cooperrider (1995), and Furlow (1991), were eliminated from the list. Taxa
rarely collected from landfills or gardens were deleted from the checklist.

Chapter 2 Methods



2 Methods

A floristic checklist was compiled for 31 Ohio counties (Appendix A).
Data for 20 counties (Ashland, Ashtabula, Columbiana, Cuyahoga, Geauga,
Holmes, Knox, Lake, Licking, Lorain, Mahoning, Medina, Morrow, Perry,
Portage, Richland, Stark, Summit, Trumbull, and Wayne) were taken from The
Vascular Flora of the Glaciated Allegheny Plateau (Andreas 1989). These
data were collected from extensive field collections by the author as well as
from surveys of major Ohio herbaria with specimens from the region (Cleve-
land Museum of Natural History, Kent State University, Oberlin College, The
Ohio State University, Ohio University, and the University of Akron).

Additional records were obtained for Erie, Defiance, Fulton, Henry, Huron,
Lucas, Ottawa, Sandusky, Seneca, Williams, and Wood counties by examining
county dot-distribution maps prepared by Braun (1967), Cooperrider (1995),
Fisher (1988), and Furlow (1991). Additional county records for three species,
Carex longii, Panicum spretum, and Utricularia geminiscapa, were obtained
from the Division of Natural Areas and Preserves, Ohio Department of Natural
Resources. In all, 2,063 species and 30 interspecific hybrids are included on
the checklist.

The arrangement of the checklist is alphabetical by genus and species; the
family name for each taxon is given in the right column. Nomenclature and
circumscription follow Gleason and Cronquist (1991). Where a name differs
from the one used by Andreas (1989), the latter is given in synonymy. The
native status of taxa was determined from Fernald (1950), Braun (1967),
Cooperrider (1995), Furlow (1991), and Gleason and Cronquist (1991).

Following Wilhelm and Ladd (1988), each taxon included in the checklist
was assigned a numerical value. The assignment of these values by the
authors was based on (a) the senior author’s extensive field experience (over
25 years) with the flora of Ohio, (b) descriptions of habitat preferences in local
and regional manuals, (c) a survey of information on herbarium labels, and
(d) published abstracts of state-listed taxa (McCance and Bums 1984). The
values assigned become less valid when applied beyond the study area.

Native species were given numerical ranks, or coefficients of conservatism,
between 0 and 10. The ranking of 0 was given to those native taxa that,
primarily as a result of human disturbance, have become opportunistic invaders

Chapter 2. Methods



AVEOVIGOdODAT
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AVEOVNV TOS

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avaOvVITOdddvo

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aVaOVITON advo

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eee

A37

Appendix A Checklist of Vascular Plants


aVaOvITIT
AVAOVOISSVad
AVFJOVHONV ONO
aVaOVIdV
aVaOVIdV
AVEOVOAN
SVFOVNITANWOD
AVEOVNITANWOD
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aVeOVNIVINHdOUOS
avaOvITIT
aVaOVITT
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aVeOVINONANVA
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AVIOVOVINLYOd
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AVAOVAELSV
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MOLTAANNNECMSMOSLMAMOEOMOLwmEeNnRooMSOrnOoS

Appendix A Checklist of Vascular Plant

A18


AVEOVRIV INHdOWS
aVaOVITIT
AVaOVLINIVHDOUGAH
AVEOVNVIGATIVA
FVIOVNVRIATVA
AVEOVNVINA TVA
AVEOVNVA TVA
AVEOVNVINA TVA
AVAOVNVINA TVA
FVEOVNVINA IVA
aVaOVORg
gvaovora
aVvaovord
aveOvoda
avaovorna

EVES tcl

EVER tcl
AVAOVTIAHdOAUVD
AVSOVITIT
aVaOVITIT
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AVEOVIAV INELLNAT
AVAOVIAVINGLLNAT
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AVAOVOLLEN
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ONnonnwmanlSSSwmarnnnornwoSanaororlrscoemaSo

A61

Appendix A Checklist of Vascular Plants


avgovsou
AVAOVOISSvUd
aVvgeOvVoIssvud
AVEOVIHOOTOLSTUY
aVdoOvOd

aveovod

aVdOVOd

avdovod

aVeOVaV

avadovav

aVaOvVaV
AVEOVARAVdVd
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AVIOVTIAHdOAUVO,
AVEOVTIAHdOAUVO
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aVaOVoIssvua
AVAOVOISSvua
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spmuayq

snuue

uwnyQUySqE

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Ter anwscoclwmolQonavalouwnwmonsoccolos

See Mee

Appendix A Checklist of Vascular Plants

AG


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me

A23

Appendix A Checklist of Vascular Plants


AVEOVOISSVad
AVAOVOISSVud
AVEOVOISSVUa
AVEOVOISSvud
AVAOVOISSVa
AVeOVINONNNVA
AVEOVINONANVA
AVAOVNADOdV
AVADVNADOdV

_ SWHOVNADOdV
AVSOVNADOdV
AVAOVCIHOYO
avaovavs
avaovod
AVEOVNIV INHdOWS
AVeOVIdV
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AVAOVaALSV
AVAOVAELSV
AVEOVAELSV
AVIOVAELSV
AVeOVaSELSV
SVaOVaALSV
AVEOVATVA
aVaOVIdV
aVaOVIdV
aVaOVIdV
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AVEOVTINONNNVS
AVEOVINONONVS
AVEOVTINONONVS
aVEOVINONNNVa

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MANMOSOOCHBONNASOOCOCOCSCSCOOTMOMAMNOEOSrGaAM

AS

Appendix A Checklist of Vascular Plants


AVOVNIOV.INV 1d
AVIOVNIOVINV 1d
AVAOVNIOVINV Td
AVIOVNIOVING 1d
AVAOVNIOVINV 1d
AVAOVNIOV.INV Td
AVAOVNIOVINV Id
AVIOVNIOVINV 1d
AVAOVNId
AVAOVNId
AVAOVNId
AVaOVOLLEN
aVaOVOLLEN
AVAOVUALSV
SVSOVUELSV
AVEOVOOV IOLAHA
AVEOVINVI
aVaOVSON
SVIOVNVIOS
AVAOVNVTIOS
AVSOVNVIOS
AVAOVNVIOS
AVSOVNVIOS
aVAOVNaGaaA
AVEOVNAGaA
avaovOd
AVASOVINOWS10d
VAOVINOWA 10d
SVAOVINOWA 10d
AVAaOVINOWA 10d
AVAOVINOWNA 10d
AVAOVINOWA 10d
avaOvOd
AVAOVAONVACAH

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(1 eyddry) wiejooouey
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(stunusu09 “g) sijensne
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wyernoqued
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ecoenatnwonvonnootencortTSoVooQgcoooose

ee Me eae

*
ee

Appendix A Checklist of Vascular Plants

A44


1. Introduction

The U.S. Army Corps of Engineers is developing a procedure for assess-
ing wetland functions using functional indices (Smith 1995). This procedure
compares wetlands using functional indices calibrated to regional reference
wetlands. Reference standards are conditions exhibited by a group of refer-
ence wetlands that correspond to the highest level of functioning (highest
sustainable capacity) across the suite of functions of a regional wetland sub-
class. The quality of species occurrences at regional reference wetlands can
be used to assist in the calibration of the vegetation components of functional
indices.

The purpose of this report was to adapt the existing Wilhelm method
(Swink and Wilhelm 1979, 1994) for evaluating the reference standard for
species occurrences at reference wetlands and other vegetated habitats as a
method to evaluate natural places by providing a floristic quality assessment
index. This report contains a floristic checklist that is applicable to 31 coun-
ties in northern Ohio. The quality index ratings presented here are intended
to both assist regional efforts to establish reference standards for species
occurrence in wetlands and evaluate natural places in this region.

The modern native flora of northern Ohio is composed of a mixture of taxa
that became established after the melting of the last Wisconsinan ice advance,
about 16,000 BP (Goldthwait 1959). The native flora of this part of glaciated
Ohio resulted from (a) the northward migration of species that survived south
of the glacial moraine (Delcourt and Delcourt 1981), (b) the establishment in
suitable habitats of northern plants that had migrated southward into Ohio in
front of the glacial advance, (c) the eastward extension of prairie plants and
plants more typical of drier areas that occurred during the Xerothermic Period
8,000 - 5,000 years BP (Benninghoff 1964), and (d) the westward migration
of coastal species via eastward drainage channels that formed in the
St. Lawrence lowlands as the ice front retreated (Andreas 1989).

At the time of the arrival of the European settlers, it is estimated that about
96 percent of Ohio was forested (Gordon 1966; Cooperrider 1982). The
remaining 4 percent of the land surface was open areas of freshwater marshes,
peatlands, prairies, and barrens (Sears 1926; Transeau 1935, Gordon 1966,
1969). Through historical accounts written by early land surveyors, Gordon
(1969) was able to reconstruct the original (presettlement) vegetation of Ohio

Chapter 1 Introduction


Wilhelm and Ladd’ —_| Present Study Values
Values for Northern Ohio

|| Phieum pratense

Plantago lanceolata

Poa pratensis

Polygonatum canaticulatum

Potentilla simplex

| Prunella vulgaris

Prunus serotina

Prunus virginiana

Pyrus ioensis?

Rhamnus carthartica

Rosa multiflora

Rubus occidentalis

Solanum dulcamara

Solidago altissima (= S. canadensis)
Solidago nemoralis

Taraxacum officinale

Trifolium pratense

Uimus americana

Viola papilionacea (= V. sororia)

Vitis riparia
= 42;N = 25;1= | R = 50;N = 24;1 =
8.4



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aVSOVITONdVO
aVaOVITOAIdvO
AVAOVITIOANIdVO
AVeOVMIV TNHdOYOS
aVaOVNIVINHdOUOS
AVaOVNIV TNHdOWOS
aVeOVMIV NHdOWOS
AVAOVMIVINHdOUDS
AVaOVIV INHdOWOS
aVaOVRIVINHdOUOS
aVaOVMIVINHdOUOS
aVeOVNIVINHdOUOS
AVEOVINV TNHdOUOS
aVaOVMV INHdOUOS
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AVEOVINV TNHdOwOS
aVEOVNIV INHdOWOS
aVaOVIaV INHdOWOS
aVaOVAaLSV
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AVAOVUaLSV
AVaOVUaLSV
AVSOVUALSV
AVAOVNEPAA
AVFOVNAGHSA
AVAOVNAGAA
AVSOVNEGUAA
AVSOVNECAA
AVEOVNAGAA
AVEOVNAGaAA
AVaOVNIVINHdOWS

(uimyu3001 *A) uMproNy “zea WnyeUEpP
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eu2qiaA
euaqiea
wuasqiaA
umaseq73A,

ecoortnurvttoOnmMromeomeooooonootroOoaM fan

Appendix A Checklist of Vascular Plants

A62


GVaDVSOU
AVFOVHINVOV
avaovssaudnd
aVaOvssaadndD
@VSOVONNL
AVEOVONNE
SVAOVONNE
@VEOVONNE
AVeOVONNL
AVEOVONNL
aVESOVONNE
SVaOVONNE
AVSOVONNE
@VSOVONNE
aVaOVONNE
AVEOVONNE
SVaOVONNL
AVAOVONNE
aVEOVONNE
AVEOVONNE
AVEOVONNE
AVaOVONNAL
AVAOVONNL
@VeOVGNV TONE
aVaOvVaNV TONE
avaovahigauaa
AVAOVAELSV
SVaOVdIHDNO
AVeOVINONANVA
AVEOVLAOSI
AVIOVLAOSI
@VEOVINVI
avaovanil
avaovanil

eoquodef

eugououre

euRrUaA
syunurai0o

Vhauioy

(Jouauy “fF “Jour) sinus “sea sinua
1Aorpnp “sea sinus
(snypAydAreyd *p) snuojoypip “zea smu)
snpunses

snsopou

smgeuraeur

rousad

Hpse1a3

snsnyja

stsuapeueo

snquoyng
snpeydaokyorg
sndseokyourq
sTuoyIq

smejnone

(snonqeq “f) snonore
(snuidye “f) smejnonreourdye
snyeuyumnoe

wary

BaroulD

epdydip

ByoypuEX
wreyonaa
wyeWwsG
Wuuewyosua
euodsouyoo
smperyselq

yoaarys “3ea worUTsatA
JOTODISIOA.

BELRY
wonsng

snuadrung
snoung
snoung
snoung
snoung
snoung

snount
snoune
snount
snoung
snoung
snoung
snoung

snoung
snoung
snoung

snoung
suepange
sue[ang
Byuossayjor

wnsfdosy
sara0s]
So190S]
snipuesy

wer SsSracrSumGarernomntrnonoernntamaomns

F

A33

Appendix A Checklist of Vascular Plants


AVAOVAELSV
aVeOVUaLSY
AVAOVAALSV
VEOVNITSANWOD
AVEOVNITANNOD
AVEOVNITAWINOD
AVEOVIGUVOVNV
AVSOVIGUVOVNV
AVEOVIGUVOVNV
AVAOVIdV
AVaOVITIT
aVaOVGIHOYO
AVAOVITIL
AVEOVOVAAIXVS
AVaOVINVT
avaovssaddnd
AVAOVOISSVad
aAVeOVOIsSsvad
SVEOVCINaLIA TIL
EVER) IREA AG CIAR
aVeOVdIaLdA TL
AVAOVOELIA TAH
aVaOVIdV
aVaOVIdv
AVEOVINONNNVA
AVEOVINONANVA
AVeOVINONNNVa
FVEOVINONNNVA
AVEOVINVT
aVaOVINVI
avaovava
HVAOVXVL
AVEOVIGOXVL
aAVAOVUELSV

sysuayeid
snyjoyyizod
snqnp

BURTUTS IA
sysuaryo
Byeaj08.1q

(CA sayy) xaquzan
(sueoypes sayy) W1eqpAa
(3 snyy) sueorpes
woquodef
esounnys
Jojoostp
Buvououre
BtOsIp109

uwinyy fdas
stfequap1990
winjeyojiad
asuaare
suandosoyd
sunsnyed
sisuaaeioqonou
wiojdouosexay
wunyeroyuy
‘Spourqieq
wnynjOAar
suaosagnd
uunsyorp
windreaXsep
‘9[e1UEpII90 “Fea asuapeuEd
asuapeued “Fea osuapeuo
BUBLUISIIA
sisuapeuro
umippsyp
aeupyyo

uododosery,
uodododeay,
uododose11,
enueosopesl
enueosapelL,
BAUBSepeLy,
uoIpuspoolxoy
wospuapoorxo],
wospuaposrxoy,
SHHOL
eIpPuOL
ependyy,

ULL

Laue,

efnyy,
idseruL
sdseru,
suadcpoyy,
spadAqouL
suardépouL,

‘wnidseyy,
umidseyy,

wnnoreyL
unNTey,
unyonay,
twnyonay,

unypoxey,
umaexeiey,

eonemntnoernranrnnacsccononmQonrwmonnmnooo

A59

Appendix A Checklist of Vascular Plants


REPORT DOCUMENTATION PAGE OMe No ovoa-or88

‘reporting burden for this collection of information ts estimated to avert | hour per response, including the time tor: Instructions, ing existing data
ournes ‘he data ‘and completing and ng the collection of “comments regarding this burden estimate or any athet arsecton ors

Ee a eae ret oti ‘of Managenton and Balgel Paperwork Redettion rojecs OTOa OvaaT Wemarstees Be Soeia TS Jetferion
T, AGENCY USE ONLY (Leave blank) [2. REPORT DATE ‘3. REPORT TYPE AND DATES COVERED

August 1995 Final report
‘4. TITLE AND SUBTITLE ‘5. FUNDING NUMBERS
Floristic Index for Establishing Assessment
Standards: A Case Study for Northern Ohio

6. AUTHOR(S)
Barbara K. Andreas, Robert W. Lichvar

7, PERFORMING ORGANIZATION NAME(S) AND ADDRESS(ES) 8. PERFORMING [ORGANIZATION
Cuyahoga Community College, Cleveland, OH 44122; ‘Feennical port
Kent State University, Kent, OH 44242; WRP-DE-8

U.S. Army Engineer Waterways Experiment Station
3909 Halls Ferry Road, Vicksburg, MS 39180-6199

‘9. SPONSORING / MONITORING AGENCY NAME(S) AND ADDRESS(ES) 10. SPONSORING / MONITORING
AGENCY REPORT NUMBER

U.S. Army Corps of Engineers, Washington, DC 20314-1000

11. SUPPLEMENTARY NOTES
Available from National Technical Information Service, 5285 Port Royal Road, Springfield, VA 22161.

‘12a. DISTRIBUTION / AVAILABILITY STATEMENT 12b. DISTRIBUTION CODE

Approved for public release; distribution is unlimited.

‘13. ABSTRACT (Maximum 200 words)

The Floristic Quality Assessment System was developed as a tool to provide a numerical value (Floristic Quality
Assessment Index) for a natural area evaluation based on plant species present. The index allows for objective
numerical comparison of two unrelated plant community types.

A numerical rating, called the coefficient of conservation was assigned to 2,063 species of plants and 30 inter-
specific hybrids (Appendix A). Appendix A contains a checklist of the vascular flora of 31 Ohio counties, including
those counties present within the Buffalo District of the U.S. Army Corps of Engineers.

Native species were assigned coefficient of conservatism values of 0 to 10. The rank of 0 was assigned to
native taxa that are opportunistic invaders of natural areas and those that are typically part of ruderal communities.
Rankings of 9 to 10 were used for those taxa that exhibit relatively high degrees of fidelity to a narrow Tange of
synecological parameters. All alien (nonnative) taxa were assigned a value of 0.

The Floristic Quality Assessment Index (1) can be determined for any natural area from the tabulation of the
coefficient of conservatism values. A higher index value expresses a natural area containing mostly native species,
whereas a lower index value reflects human disturbance by taking into account the presence of alien (nonnative)
taxa.

14. SUBJECT TERMS 15. NUMBER OF PAGES
Coefficient of conservatism Exotic species 87
Disturbance Natural areas assessment 16. PRICE CODE
17. SECURITY CLASSIFICATION | 16. SECURITY CLASSIFICATION ] 19. SECURITY CLASSIFICATION | 20, LIMITATION OF ABSTRACT
‘OF REPORT OF THIS PAGE OF ABSTRACT
UNCLASSIFIED UNCLASSIFIED
NSN 7540-01-260-5500 Standard Form 298 (Rev. 2-89)

Prescribed by ANS! Std. 239-18
298-102


aveovod
avdOvod
avaovod
aVadovOd
aVvaovOd
avaovod
aVveOvod
avaovod
aVvaOvOd
AVaOVOd
avadovod
VaIVOd
avaOvOd
avadovod
aVaOVOd
avaovOd
AVEOVOd
AVaOVOd
avaOvOd
aVaOVITVAV
aVeOVITVUY
SVeOVIdv
SVEOVaITVXO
aVaOVAITVXO
AVAOVAITVXO
aVeOVAITVXO
AVaOVAITVXO
SVaOVdITvxO

SVaDVGNNWSO
SVEOVGNNWSO
SVEOVGNNWSO
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aVeOVIdV

uodseso1oeyds
(wnendns “g pue saproisouse “g “[out) wnynpi3u
wnorydrepepyd
saypuesodiyo

umaoe yur
uodresoso1ut
uinyjoyueauty
uunyoyney

winsourSnuey
wnwojoyorp
wndoprUoIOYoIp
umesodnedop
wnyenTWOD
wnueiquinjoo
wmnupsapueys
(pesumes “g “ToUu) aseyyides
wnqyAydyeo

Ttos0q

(HT}9u 491g “q “Tour) afearog
unyjoyuy
uinyoyonbuinb

Jo;pIsy

‘BODRIOIA

moms

sipuesd

quamp

‘ByengyUI0D

(gugjuoul “C) efjesorz08
eueyuyaata

styeSax

eueuoiceyo
vawoureuuys
syAs}uo]

Huowkelo

wnotueg
wnotueg
umnoqueg
wnotueg
winoyued
umnoqued
wnoqueg
‘undtued
wnojued
wnoqueg
wnoqued
wnopueg
umorueg
unayueg

MnweowmnQoonrnovwmmnanQumanannanrnonads

Appendix A Checklist of Vascular Plants

A42


Wyvomuaavgy
BIOL XE,
spusjnorxey
edreoojse]
ByeUIseATASE],
sEnsnoey
tsouref
suaosoummqut
Jouaquy
eurouaisky
sidgjourre<y
BUERPDOOYOIY
Bron
MuopAey
1herd
suepiued
euoRs
susosaioed
Wyuey
‘PSOULIOJ
‘Byenoyoy
Parl

(eapoone[g *D) eunadsoooey

BaIeONysay
ours

(eyjuepeydao *D) eyeutyse

eoulnga
Buuadstp
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wupuelp
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Ye8pny “ea sepigap

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A13

wokSrSumennSnnworaonrnnSame

COEELEL ELLE ELE EE ELE CL ELE EEL ELEED

Appendix A Checklist of Vascular Plants


avaovanl
avaovanl
aVvaovarl
avaovarl
avaovani
aVeOVOISsvad
aVaOVOIssvad
aVEOVOISSvad
aVaOVOISsvad
AVEOVAELSV
AVEOVUALSV
aVaOVaaLSV
aAVaOVUALSV
aVaOVUaLSV
AVAOVTIAHdOAYVO
AVAOVTIAHdOANVD
AVAOVTIAHdOANVO
AVEOVTIAHAOAYVO
AVEOVTIAHdOAUVOD
AVAOVTIAHdOAYVO
AVAOVTIAHdOAUVD
AVEOVTIAHAOAAVO
AVAOVTIAHAOAUVO
AVAOVTIANdOAAVO
SVEOVTIAHdOAUVD
AVEOVTIAHdOANVO
AVEOVA TVA
avaovilawnond
aVaoOvIEna
AVEOVNOVAV TA
avdovOd

avadovod

aVdOVOd

avadovod

winjeuaronut
twnuejuour

umonuepe
winyoynsnsue

winpiqre

ayeupyyo

uMUTSSBIs

(cequy Bojsseig) ssussrE
("8 waysseig) Bare
umuepreut

wnyeTosN
unaoeUTyUIqQaIA
winyeroy3ed

umequroey

suas,

BOTUISITA

wreyas

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(sisuajead *s) epone]
(cP spuya47q) eojo;p
Buo;OyDIpP

poraso

‘woyu09

eorueajAsuad “sea BuRTUTOIRD
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euryunue

esousds

smejndue

sjsuaare

sisuapeuvo

syPHA

ByePpAIA

BCI

vanes

wnryounAsig
uunqyouAsis
uuntyoupAsTS
uunyyoupsAsis
uunqyoupAsis

tunpaquixsts
sydeuys
sydeuys
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uunmydis
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SOSCSOMONSNTDASCSCOSCSOSOVNOALVAMDOSSOOVSY

see wx wm

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*

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eee

A55

Appendix A Checklist of Vascular Plants


AVaOVesdAD
AVEOVAddAD
AVaOVasdAD
AVeOVeadAD
AVEOVUEdAD
AVIOVAEdAD
AVAOVaadAD
avaovaadAD
aVAOVaadAD
aVeOVaedAD
aVaOVaadAD
aVaOVeadAD
AVEOVaEdAD
aVaOVeadAD
aVaOvVeadAD
aveoOvaddAD
aveOvesdAD
EVER) ECA Ce)
avaOvasdAD
aVaOVaEdAD
aVaOVasdAD
aVeOVaadAD
HvdOVAddAD
aVaOVAEdAD
aVaOvaadAD
aVeaOVesdAD
aVaOVaEdAD
aveovasdAD
aVeOVUadAD
AVaOVaEdAD
aVeOVaadAD
aVaOVaadAD
aVadOVeadAD
AVEOVUadAD

estos
euedoos
weiqeos
Temes
euuadsosny
easor
fuospxeyou
esionar
exayonal
wyerper
wooford
vuiseid
vaneid
syperdovad
eAydAveyd
voursejueyd
eowuealAsuod
(esourgnuey “D) ented
eyejnounpad
susasajqed
euuadsostjo
edreo03tj0
stjeuuou
sisuoum3upysnur
HBroquoyyour
Hpeow
epuny
eugndny
stuuogyndny
BOPURISTNOT
voy
‘BSOUT]
BAsquoida,

vapeday

558

Appendix A Checklist of Vascular Plants

Bem

NnNnmowowraoellSromareonnwnwmmnSnnanrta

mm

CECE EE ECEECCEEEEEEEECECEEEEEEEE

A14


aVSOVINVT
AVEOVINVT
aVEOVGITANVNVH
AVAOVNIOVUOE
AVEOVGIHOYO
aVaOVdIHOYO
AVAOVGIHOYO
AVaOVGIHOYO
aVaOVGIHDYO
SWaOVdIHOYO
VEOVGIHOYUO
AVaOVdIHOUO
AVaOVGIHOYO
avaOVdIHOYUO
aVaOVGIHOYO
aVeDVGIHOYO
AVaOVGIHOUO
AVAOVTIAHdOAUVD
AVAOVINIdTVSavO
AVAOVINA SV
AVeaOVaRLSV
aVaOVNIVINHdOYOS
AVAOVGIHOYO
AVaOVGIHOYO
AVaOVUaLSV
AVSOVAELSV
AVAOVAELSV
AVAOVAELSV
avaovdvd
aVaOVOd
avaOvOd
aVaOVOd
avdOVvOd
GVIOVOd

sapjorsaind

wnprdsty

eueyUTaaA

euerUrsItA

(Ca umnssop3oj909) sIputA
(-d eraypueyetg) sapoodsd “sea sopoodsd
(cd exoyuryeyg) exopjipuesd “sea sopooksd
(Cd esampueyetg) eusouresod
(Co esoupueyeyg) BreINDIqQIO
CL exoqpuereyd) eseydoonay
(CL esompueyejd) 82208]

(Cy exampuryeyd) earoquedAy
(Cy exapuereyg) Uayooy

(Cy exoypuejetg) BAe

(19 elayjuere|g) B1e[[aAeO.
(C9 eompuere|a) sueHO

(-q erompueretg) smoydtreydaq
ByJoJLaUOZI0NS

BoIOIp

suadotip

esouenbs

eoajsou

Byeyassay

suaoseqnd

wnsouyayn

winaindind

wingjoyismqo

(winsodsiA “D) Hunoseur
xeut

Brens

stjeuoiuaides

‘BEeoTour

sypued

stsuopeues

winteydeup,
wunyeydeup
wnyqeydeup
winyeydeup
aupiip
BHI
BuBOKID,
BpeOKIO
eHa0KID
eusodI

monrnacSamnmveStroncoS Sumo SSouSrrGaganan

x sete me

*

A29

Appendix A Checklist of Vascular Plants


AVAOVNVLISVOOddIH
AVEOVNV.LSVOOddIH
SVaOVIdV
avdOvOd
aveOvrivNNns
AVaOVINVIGV
AVEOVTINONONVS
AVEOVINONNNVA
AVEOVYOOV
AVeOVINONANVS
AVAOVUALSV
aVaOVesOV
AVaOVesOV
aVaOVaROV
AVSOVUIOV
aVeOVasOV
aVeOVUFOV
aVaDVasOV
aVaOVIddOHdng
aVaOVIddOHdNa
aVaOVIadOHdNg
AVAOVA TWN

ATINVA

wnueysesoddyy
eiqeis
wpreasepod
woppuytd
esosuny
winjepod

eign
(epodAyoed “y) eqre
snurepes
‘esuaae10qeaou
uumyposay TU
wimeoids
wnreyoors
wnupeyooes
umqny
sapjourzed
umorueayAsuod
opungou
‘ROTUTBITA
vaproquioyy
eyoyaehyso
Asesydosyy

LAB LIdd DID adS

snnasoy
snqnosoy

*

sdojygay
eyaunpy
winjUuEIpy

BR Bee

<
cocomSonmweweSrravonoows
*

SQNID 9409 SLINAWWOD

Appendix A Checklist of Vascular Plants


avaovaral
avaovaral
aVaOVvaral
avaovaral
AVOVINATOANOD
AVEOVINATOANOD
AVAOV INA TOANOD
AVSOVINATOANOD
aVaOVOISSvUud
aVdOVUSLSVY
AVEOVNINVS IVa
AVAOVNINVSTVE
AVSOVNINVS IVa
SVEOVITOSINOV
AVAOVITOSINOV
aVaOvoIssvua
AVIOVITIT
aVEOVUaLSVY
AVaOVISNTIO
aVaOVISNTO
AVaOVISNTO
AVSOVISNTO
aVaOVISNTO
AVaOVISNIO
aVaOVISNIO
aVaOVISNTO
AVAOVISNIO
avaovIsn1o
aVaOVISNTO
aVaOVISNTO
aVaOVISNTO
aVvaOVISNTO
aVEOVULSVY
AVAOVTIAHdOUGAH

sniosepnesd
eoqueuLied
wIeISsHO
sypnestacaq
waandand
eyemnpued
Baowrapay
aupo09
snpyneuurd
wnyusay,
eprted
stsuode>
supuesyeq
wreyonea
voedo
ByeypPquin
wnsiy
‘BywoyPBT
umdseoorseyds
wmepyueskd
‘umnyeyound
umortord
umnyerojied
wnypnur
snfeu
winery
wintjueuws
sepyournua’
wnondio
Hpuowumup
‘asuapeues
gfeaiog
paoeqioy
cunuemursit4

SHI

su

SUT

sul
waoulody
eoowody
waouody
waouody
snyjuepoy
jnuy
suapeduyy
suapeduyy
suapedwy
xoll

xoll

speq
srxodéyy
spaoypodé
wnouad Ay
wmoned«R
wnouedéE
wnouad Ay
uunopadéy
winoyod AE
winsuedAy
wmnouedéE
winouodAyy
wnowadéy
umnouedAy
wmnouadAy
umopadkH
umopiodAH
skxouawAyy
uunytydospaH

neESwmowtrSSrnotmnrnrorscoraenmoveconefunos

oe

mame ee

Appendix A Checklist of Vascular Plants

A382


AVIOVIONVYGAH
avaovava
avaovava

avaovod

avdovod

SVEOV TIAHdOUGAH
AVIOVTIAHdOUGAH
AVAOVNVIOS
AVFIOVAELSV
AVEOVINVI
aVaOVIdv
AVAOVOVUAIXVS
aVeOVIVTINHdOUOS
AVaOVIaV TNHdOUOS
aVaOVMV INHdOUOS
AVAOVMIVINHdOUOS
avaovav
AVEOVIVTNHdONOS
aVeOVNV INHdOWOS
aVeOVIdV

avaovod

AVAOVLIA

SVEOV IIA
AVIOVTIAHdOAUVO
AVIOVTIAHAOAUVO
AVEOVOVUAIXVS
AVEOVOLLEN
AVEOVUEA Vdd
AVAOVUFAVdVd
AVEOVUAA Vdd
aVEOVUEAVdWd
avaovOd

aVaOVOd

aVaOVOd

snypxeu0l0>
syzedna
soryoeiskjod
sjsuapieugd
eaoeurpune
nysind

eignp

epyiqdy x
snppqdy
suaosapnyy
‘euROLOWe
septopas

snpyred
(snsoodyeo “q “Jout) smaeBtAgEy
smnsaty

sienaip

‘BOIUIaIIA
ByBIOVOUE]
sisuopeued

Bayes
UINTPOIHeIID “FEA wNAdEIes
(euasut “g) BaoeIA
eroyanbuinb
viersnsey
ssuapeues

eone[s
vorueatAsuad
uinaayyuuies
‘seoout

umyqnp
auowasie
umeaita
UINUTTSS|SOTTYA
winjaids

snydyepeyd
‘snjoaseyd
snjoeseyd
speed
suepeud
eyoRyd
eypoRy
equnjag
sayseyed
Bid
eipopuad
mowed
uoura)suad
uowssuag
uowajsuag
vows)suad
eupuesiad
sue[noipad
suejnoipad
wouupsed
umyedseg
snsstsousyed:
snss}oouayped
eryoduored
eryoAuoreg
eisseuled
ByEeNed
qoandeg
sanded
sasdeg
saaudeg
uunopued
wunoqued
uunoyued

Savrooccoer Serr mamovwmemonnmSoooSnoowecod

Kee eK

Oe

ee ee

oe

43

Appendix A Checklist of Vascular Plants


Corps of Engineers Research Report Summary, August 1995

Wetlands Vegetation

N

Floristic Index for Establishing Assessment Standards: A Case Study for

Northern Ohio (TR WRP-DE-8)

ISSUE:

The assemblage of plant species can indicate
various responses to environmental gradients
and disturbances. Information is needed about
the occurrence of species within natural and
disturbed plant communities for establishing
reference standards for use in the hydrogeomor-
phic approach used for evaluating wetland con-
ditions and natural places.

RESEARCH:

A floristic checklist was compiled for 31 coun-
ties in northern Ohio. Rankings of 1 to 10 were
assigned to native taxa based on their degree of
fidelity to a range of synecological parameters.
Plants found in a variety of plant communities,
including disturbed sites, were assigned rank-
ings of 1 to 3. Rankings of 4 to 6 were applied
to taxa that typically are associated with a spe-
cific plant comunity, but tolerate moderate dis-
turbance to that community. Rankings of 7 to 8
were applied to those taxa associated with a
plant community in an advanced successional
stage that has undergone minor disturbance.
Those plants with high degrees of fidelity to a
narrow range of synecological parameters were
assigned a value of 9 to 10.

SUMMARY:

The floristic quality index for 2,063 plant
species in northern Ohio provides a tool to
assess the quality of naturalness or presence
of conservative species. It allows for an ob-
jective numerical comparison of two or more
unrelated community types and reflects nu-
merically the impact of human disturbance by
taking into account the presence of alien taxa.
The ability to evaluate floristically and assign a
repeatable quantitative value has use in assess-
ing wetland restoration projects and in design-
ing and monitoring mitigation creations.

AVAILABILITY OF REPORT:

This report is available on Interlibrary Loan
Service from the U.S. Army Engineer Water- -
ways Experiment Station (WES) Library, 3909 -
Halls Ferry Road, Vicksburg, MS 39180-
6199, telephone (601) 634-2355.

To purchase a copy, call the National Technical
Information Service (NTIS) at (703) 487-4650.
For help in identifying a title for sale, call (703)
487-4780.

NTIS report numbers may also be requested
from the WES librarians.

Mr. Robert W. Lichvar is now at Cold Regions Research and Engineering

Laboratory (CRREL), phone: 603-646-4657.

(April 2001)
Please reproduce this page locally, as needed.


AVAOVSOWIN
AVaOVOISSvad
AVAOVOISSVUd

aVadOvOd

aVaOVOd
AVAOVLLGAVLSNNAG
SVEOVINONONVA
AVAOVINONNNVa
AVEOVINONONVA

AVSOVUHLAT

aVeOVIdV

AVAOVNVIOS

AVADVNVIOS

AVIOVIaV INHdOUOS
avaovOd
avadovod

avaovsou
avaovdvd
avaovod
AVAOVINATdSV
AVAOVINATdSV
AVAOVINATdSV
aVaADVGIHOYO
AVEOVGIHDYO
aVaOVdIHDUO
AVaDVdIHOYO
aVaOVGIHOUO
aVaOVaadAD
aveOveddAD
aVaOVesdAD
aVaOvVeadAD
aVeOVUadAD
aveOVvesdAD
aVaOVasdAD

sIsuaour[T:

eyydos

eyeuutd

esonxeyy

esoydsao

eingoypound

‘@usoomn

wnjEyexo

(‘8 epyosuoD) winndyqure
snqeypEonIaA

Boe

uwinuoweys

Bpxouy

eytAydosseur

Beards

essaidwioo

suedax

waandind

ByBIIWIO[S

esnnoid

sisexy

Bsogiqing

oeulsar

winpipues

suaosaqnd “sea snjoaayeo
wnoytared “sea snjoso[eo
gineoe

sniyjoyinuay

snsosins

(snxayjut ‘D) snsouenbs
Hzyuyamyos
(suaosoujsnuudy “D pue Huew[aBue “2 “[out) smeI0po
Huowygnoy

susosoaryy

snijuewseq
Byuyemosoq
Bjuyemosaq
eisdureyosoq:
eisdureyosoq:
enpewsuusd
wunturydieg:
wnurydjeq
wnyuyydpq
woposeq
snoneq
eimed
Banged
euloIsiseq
ByuouuEg
eruowpueg
eprequed
wae
sudpeg
suadoysk>
suardoisk)
suardoish
uungpadydé
uunypadpd
umppedidé)
uunypadudé
wunipaddé,

mMNNneocLamamaanoovonmyrnQoron

bee

A21

Appendix A Checklist of Vascular Plants



avaovsod
avadovsod
avaovsou
aVaOVSOU
avaovsod
avaovsou
AVAOVIGUVOVNV
AVaOVUELSV
avVaOVINLad
aVSOV INL
avaOVNIVNN
aVvaeoVRIVANS
avgovava
AVAOVNYOD
AVFOVNYOD
AVAOVNYOOD,
AVAOVNUOD
AVAOVNYOD
SVSOVNYOD
AVaOVNAOD
AVFOVNYOD
AVAOVIGOdONSHD
AVAOVIGOdONSHD
AVAOVUELSV
AVSOVUELSV
AVeOVaELSV
AVAOVUELSV
AVSOVGIHOYO
aVaDVdIHOUO
AVEOVCIHOYO
SVaOVINONONVa
AVaOVUELSV
AVEOVUELSV
AVaOV TINA TOANOD

weeqey
Wyyed-stua
‘BaUtI09
(etoytpumios *D) edresosAsyo
uospuopodyes
wprouresq
‘Bpsk3d0o
snyeuuydyq
‘eynw0.
eueopoue
suartsodwes
vmnaey

Bye
(esaytuojoIs “D) Baoes
esosn
PsOWaIe!
epuoy
Hpuowwmup
sisuopeued
wnwouwe
rosWaIe
umnppu
‘umyjoyydosshy
suadin
BpopUR
‘ByBjosoUR]
Br0yypuess
ep
eZIyOUOpO
eyejnoews
Bros
eUNISSTSOUrEL
sisuapeueo
ssuaare

corer unncoooncoonnaruarvornansooQorilm:

xe eM

*

Meee

A19

Appendix A Checklist of Vascular Plants


avaovsod
avaovsou
AVEOVOrIVS
AVEOVOITVS
aVEOVOIIVS
AVAOVOITVS
AVAOVOITVS
AVEOVOITVS
AVaOVOIIvs
HVAOVOITVS
AVFOVOIIvS
AVEOVIECSLNOd
SVFOVINA TdSV
AVAOVIGOdATOd
AVAOVUELSV
aVeOVaELSV
AVIJOVNODATOd
AVAOVNODATOd
SVEOVNOOATOd
AVEOVNODATOd
AVFOVNOOATOd
AVEOVNODATOd
FVFDVNODATOd
SVEOVNODATOd
HVEOVNODAIOd
AVFDVNODATOd
AVAOVNODATOd
AVAOVNODATOd
AVAOVNODATOd
AVAOVNODATOd
AVFOVNODATOd
AVFIVNODATOd
AVFOVNODATOd
AVFDVNODATOd

smyeroyEn
smejndys
Ippefx
suaosoued x
soprojnman
ead
eyAydornoy
Byeuapipuels
soploiep
Bsoyrures[eq
eqre

wepi09
saployonsosse
umueIUrgutA
trjepaan
sisuapeues
unueruraita
nua)

‘Suapueos “eA Suspueos
WINIEISHD “JeA SuapuLdS
wnenises
snysnqos
twnyeound
Byreojsuad
winojueayAsued
ayequapio
wmnyosnpedey
soprosadidospAy
sodidorpAy
wnjoa10
wmnyepydsna
snynajoauca
‘pours

¥harea

snupuesayog
snijuesayiod
snyndog
snndog
sngndog
snqndog
snindog
snndog
snindog
snyndog
snndog
Buapaquod
winyonsjog
uinypodhjog
eyuusAlod
‘eyuurdyog
umnuo3hjod
uinuosAjog
winuosfjog
umuoshjog
umuosijog
winuo3fjog
umuosfjog
umuo3{jog
winuoséjog
umuo3fjog
winuo3fjog
umuosAjog
umuosAjog
wnuoshjog
umuoséjog
umnuos{jog
umuo3sjog
wmnuosjog

SaocorannanSnowomanunsrnor sn onnnwonoowen

mse

Appendix A Checklist of Vascular Plants

A46


This plant is relatively rare in Ohio and is listed as endangered on Ohio’s rare
plant list (Division of Natural Areas and Preserves 1992). Therefore, the
coefficient of conservatism values presented here will probably vary for
another geographic region outside of northern Ohio.

Overall, Wilhelm and Ladd found that natural areas with ranking above 35
are significant from a regional perspective. Areas rating above 50 were
extremely rare. It should be noted that Wilhelm and Ladd assigned special
values (15 and 20) to those taxa considered threatened or endangered within
the Chicago region. As a result, their Natural Areas Index values for rare
communities would be higher than is possible under a strict 0-10 ranking
system.

The floristic quality assessment index can be used in establishing reference
standards for regional wetland subclass. The index can also provide a method
to measure the response of the vegetation community to mitigation from inva-
sion of non-native to native species. This measurement provides a numerical
method to rate the results from various mitigation methods from either
enhancement, restoration, or creation.

Chapter 3. Application of Coefficient of Conservatism to Floristic Quality Assessment System


SVEOVTIAHdOUGAH
AVIOVTIAHdOUGAH
AVEOVTIAHdOUdAH
aVAOVIdY
AVAOVIdV
aVaOVIdV
aVeOVINONANVa
AVAOVAONVAGAH
AVAOVIOIA
AVAOVEAVNNVD
aVAOVEYNNVO
aVEOVISID
SVeOVITIT
aVadOvOd

SVeOVOd

avaovod
AVEOVTIAHdOAUVO
aVAOVOd

aVaOVOd
SVeEOVAELSV
AVEOVUaLSY
SVaOVaSLSV
AVAOVUSLSV
aVeaOVASLSV
aVaOVaaLSV
AVEOVYALSV
AVAOVAELSV
AVAOVAALSV
AVaOVaELSV
AVaOVaALSV
AVAOVUELSV
AVIOVATVN
AVEOVATVN
AVOVATVN

uiny[Aydosoews
‘osuapeues
winyejnarpuodde
‘weqjequin
sapjojnounuex
euespoure
‘sysuapeueo
suaosazoqie
Jojoou0a
soqndny
snojuodef
BSO}UaUIO
ByoypuEy
ase3na
tnyysnd
umyeqnf
umjejpqun
snjeugy
welopo
winsous

nen

umiqeos
(umupuasoy “1) seprortesopd
Blpsoyd
winjejnoyued
wunpydiduoy
(@suapeueo “}) HuUpey
waouol3
winpunqyoy
uinsojjdsaeo
winoepnuEIne
umu0py
somnayosour
SsyAae[

wnqAqdospéyy
wnyAydospAHy
winytAydospéy
‘opMoo0spéH
a14oo01p4H
aphioooupAy
snseupAH
easuelpéy
snyuegkyy
snjnuiny,
snnumy
equospny
ISOH
wmnapsoyy
winapsoy]
wnapioH
wina}sojoH
sndj0H

20] 4OOs81H
wnpoRIaH
wmIesatH
wnpoRIaH
wpe
wpe
wnpORIaH
WnIORIOI
WINTOBIOTEL
uuNoRIaTEY
wnpelayy
wunpesay
umpezany
snosiqiH
SnOStqtH
SNOSTqTH

aAwooocowvlSrvcsconwmrwmocoococlQonrSraoQowonr

eee

A311

Appendix A Checklist of Vascular Plants


avaoviand
aVaOvOd
AVAOVNIOV.IOAN
AVAOVNIDV.LOAN
AVAOVINV INHdONOS
AVEOVMV INHdOWS
avaovod
aVAOVNIOVUOE
AVEOVHINVANAN
SVEOVINVT
AVAFOVINVT
aVEOVINVIT
AVeOVINVT
AVEOVINVAT
AVEOVINVIT
AVEOVINVT
aVEOVINVI
SVAOVNAdSININ
AVEOVINVT
aVaOvava
aVsovava
aVaOVEva
aVaOVITIT
HVaOVRV INHdOYS
avaoOvevd
avaovavd
AVEOVITIT
AVIOVATOONO
SVEDVaALSV
aVAOVaALSV
AVAOVAELSV
AVeSOVINVIT
AVEOVA TVW
AVFOVA TV

suadar

sysuaus
wausseyodu
edejef
suoSuu
smeye
wnsnyya
‘BOIUISITA
weyosN
BSOUIIA x
Byloj;punjos x
wipadyd x
sypuas x
Byeyp x
Byeayds
oyyauoy
sisuaare
asuapeueo
syeupyyo
syeupyyo
BUNSspE
eq
uNoTUTatA,
aueouy]
eapes
eugndny
euerUraata
spadonpans
epNoaT
sapjopeopyeur
sup peur
aredina
‘spysoayts
Byloyypunjor

BEY
SMUUBISTAL
SIGE
STUGETAL
snynuntyy
snnU
wr
SUSHI
sompueduapy
eyUa
BIUay
BYyUAW
‘ByUay
eypUa
ByUaTAL
BUUay
ea
wnuuodstuay
SSSI
STONE
STONE
STOHPIAL
WONHUEaA
wnukdurepayy
O8BopaW
O8eo1paW1
FlOapa
BponaneyA
BpEDEW
BHBOEBEA
BpEIyEW
wnjqnueyy
BAR
Bae

ecoscosconneooaseoosconneoooco coc onmamonooon

[ *

bee ee

o#

eee eee

A39

Appendix A Checklist of Vascular Plants


Fisher, T. R. (1988). The Dicotyledoneae of Ohio. Part 3: Asteraceae.
Ohio State University Press, Columbus, OH.

Forsyth, J. L. (1970). “A geologist looks at the natural vegetation map of
Ohio,” Ohio J. Sci. 70, 180-191.

Furlow, J. J. (1991). “The vascular flora of Ohio. Volume 2, Part 1,
Dicotyledoneae: Sauraceae through Fabaceae,” Checklist and distribution
maps, Reprographic manuscript, Ohio State University, Columbus, OH.

Gleason, H. A., and Cronquist, A. (1991). Manual of the vascular plants of
Northeastern United States and adjacent Canada. 2nd ed., New York
Botanical Garden, Bronx, NY.

Goldthwait, R. P. (1959). “Scenes in Ohio during the last Ice Age,” Ohio J.
Sci. 59, 193-216.

Gordon, R. B. (1966). Natural vegetation map of Ohio at the time of the
earliest land surveys. Ohio Biol. Surv., Columbus, OH.

. (1969). “The natural vegetation of Ohio in pioneer days,”
Ohio Biol. Surv. Bull. New Series 3(2), Columbus, OH.

Herman, K. D., Penskar, M. R., Reznicek, A. A., Brodowicz, W. M.,
Wilhelm, G., and Wetstein, L. (1993). “Michigan floristic assessment
system with wetland categories (Draft version),” Reprographic manuscript,
Michigan Natural Features Inventory, Natural Heritage Program, Lansing,
MI. :

Herrick, J. A. (1974). “The natural areas project. A summary of data to
date,” Ohio Biological Survey, Informative Circular No. 1, Columbus,
OH.

Hobbs, R. J., and Huenneke, L. F. (1992). “Disturbance, diversity and
invasion: Implications for conservation,” Conservation Biology 6,
324-337.

Maser, C. (1990). “On the “naturalness” of natural areas: A perspective for
the future,” Natural Areas Journal 10, 129-133.

McCance, R. M., Jr., and Burns, J. F., ed. (1984). Ohio endangered and
threatened vascular plants: Abstracts of state-listed taxa. Division of
Natural Areas and Preserves, Ohio Department of Natural Resources,
Columbus, OH.

Noss, R. F. (1985). “On characterizing presettlement vegetation: How and
why,” Natural Areas Journal 5, 5-19.

Pielou, E. C. (1979). Biogeography. John Wiley & Sons, New York.

References

11


US Army Corps
of Engineers

Waterways Experiment
Station

|
|

‘
7

Waterways Experiment Station Cataloging-in-Publication Data

Andreas, Barbara K.

Floristic index for establishing assessment standards : a case study for northem Ohio
/by Barbara K. Andreas, Robert W. Lichvar ; prepared for U.S. Army Corps of Engineers.

87 p. : ill. ; 28 cm. — (Technical report ; WRP-DE-8) (Wetlands Research Program

_ technical report ; WRP-DE-8)

Includes bibliographic references.

1, Wetland plants - Ohio — Indexes. 2. Exotic plants. 3. Plants — Indexes. 1. Lichvar,
Robert Wayne. Il. United States. Army. Corps of Engineers. II. U.S. Army Engineer
‘Waterways Experiment Station. IV. Wetlands Research Program (U.S.) V. Title. VI. Series:
Wetlands Research Program technical report ; WRP-DE-8. VII. Series: Technical report
(U.S. Army Engineer Waterways Experiment Station) ; WRP-DE-8.

TA7 W34 no.WRP-DE-8



sidajondAso

Taloo-stup

eras

uD

Teme

wNjOAuOD

eaprouod

Brounfuoa

(effainsary *D) ereuejdwoo
Psow0d

stunuui0o

esoydoreydeo

euETUT}OeS

euefare

suaosoues

Humeqxng

sugosouuniq

seprouloiq

(eisajour *D “Jput) JoKAauq
epurlq

qqeq

ame

(fomoy *D) eaoejtideo “rea vonuepe
vonuepe “sea eonuEpe
soposaye

‘eyuerXare

eyejore

sypenbe

epidiny “rea ejoqrydure
eursingye

suaosainjoqye

Hsuourwsa “rea sueoiqye
(e1ooipre “D) suwotgye ‘sea sueotqye
wee

Appendix A Checklist of Vascular Plants

Oe

wmaowormalnawnananntenalonoennannonaoanmSa
BR

Ai2


SVEOVAELSV
AVSOVAALSV
AVEOVUELSV
AVEOVUALSV
aAVaOVUaLSV
AVEOVUELSV
AVaOVUSLSV
AVEOVUALSV
AVaOVUALSV
FVAOVAALSV
AVAOVUALSV
AVAOVUALSV
SVEOVUELSV
AVEOVUSLSV
AVEOVAELSV
AVAOVAELSV
FVAOVaELSV
AVaOVUALSV
AVeAOVUALSV
aVAOVNVIOS
AVAOVNVIOS
AVAOVNVTOS
AVAOVNVIOS
AVAOVNVIOS
AVAOVNVTOS
AVEOVOVIINS
AVIOVOVIINS
AVAOVOVIINS
AVEOVOVIINS
AVAOVOVTIINS
SVeOVITIT
aVaOVITIT
aVaOVITIT
HVaOVIdV

eyopuyn
esourgyn
‘esourenbs
esorseds
suasyaxeduras
esosn
Epis
neppa
seproorunresd
Bed
‘sISUsOIYyO
sqpexourou
eaount
epidsiy
voquesis
‘SHINVOTKATT
sisuepeueo
eIsaeo
4010919
winsozaqny
sapjoIpees
umnje.3jS01
umusqu
ereurejnp
asuauyjore.
BxosIpunjor
eprdsty
BeoRqIay
voneys
wmequoe
ByOSIN
Beles
esouraoer
earns

odepyos
odepyos
odepyos
ogepyos
o8epyios
o8eprog
osepros
o8eplos
osepros
o8eptfos
oeplos
o8epros
o8epyos
o8epyos
odepitos
odepios
oBeprfos
ogepyos
ogepyos
winuejog
umueyog
umnuEloS
uunuejos
umuejog
wmnuBlos
eps
xeqWS
epg
xeqs
xeq}UIS
BUpDETWIS
PUTSETTS
UPSETS
unis

munankloortrntseooncoocotrnanonnannmlwalaonmonnawonw

Appendix A Checklist of Vascular Plants

AS6


Appendix A

A Checklist of Vascular Plants for
the Floristic Quality Assessment
for Northern Ohio

Key: CofC = Coefficient of Conservatism
* and bold = Alien Taxon .
** = Native to another region of Ohio, or includes both
native and nonnative populations
X = Extirpated!
E= Endangered!
T = Threatened!

1 Division of Natural Areas and Preserves 1992, References cited in this appendix are listed at
the end of the main text.

Appendix A Checklist of Vascular Plants - Al


Wetlands Research Program Technical Report WRP-DE-8
. August 1995

Floristic Index for Establishing
Assessment Standards: A Case
Study for Northern Ohio

by Barbara K. Andreas

Cuyahoga Community College
Division of Natural Sciences
Cleveland,OH 44122

and

Department of Biological Sciences
Kent State University

Kent,OH 44242

Robert W. Lichvar

U.S. Army Corps of Engineers
Waterways Experiment Station
3909 Halis Ferry Road
Vicksburg, MS 39180-6199

Final report
Approved for public release; distribution is unlimited

Prepared for U.S. Army Corps of Engineers
Washington, DC 20314-1000


VdOVOd
avaovod
AVaOVIASINO’
AVAOVIASINOS
aVaOvIaSINO
SVaOV.LaSINO’
aVSOV.LasInod
AVSOV.LASINOS
AVAOVLASINOT
AVaOVLaSINOA
AVEOVAIHOYO
AVaOVYOVNO
AVdOVYOVNO
VaDVAOVNO
VaDVAOVNO
SVEOVADVNO
SVEOVAOVNO
VIOVADVNO
aVaOVOIaa
AVAOVHONVEOUO
aVaOVOd
SVaOVOd
SVaOVOd
avaOvod
avaovod
aVdOVOd
aVOVOd
aVaOVOd
AVAOVLRIVHOOUGAH
AVAOV.LRIVHOOUGAH
AVSOVOd
aVaOVaddAD
aVaOVUIdAD
AVAOVYEdAD

sysuausyy

suerpides

Huosjou x

Tsstsay x

uinjeSouea

umoneayfs

uimyedtaaey

oyeurady

arperany

asuaare

aups0gai1ey

wNjONS

umuoypared
umnypAydorday
wingnsayy

winy210[09

uumgerfo

wntjoynsnsue

suadar

wuerUrsitA

(s uoskdoa3y) jyuyppus
(‘1 uo1kdoa3 y) suader
SNOTUTSIIA

snsoq[ta

(LuosXdosdy) snjneokyorn
snuedg

(eimed xinsXyqq) xinsky
sisuapeues

Ayrennu

sisuapeuo

woypuy

(eondyts"a) streaiog “sea synuay
weyasor
‘wyepnsuespenb

spsodery
snsoudesg
winjasinby
uimastnbg
uungstnby
winasinby
wmasinby
uinjesinby:
uumgastnby,
cumesnby,
spoedydg
uuniqordg
uunyqorydy
uumiqopidg
umyqoyydg
uunigonida
winygondy
unigopidg
voedidg
snseyidg,

ALWPCONNMNNAETMODMDMDODYTNONOASCORNMROETTNS

Appendix A Checklist of Vascular Plants

A24



avaovod
aVaOVOd
AVIOVNV INHdONOS
aVeOVNIV INHdOWS
AVAOVRIVINHdOUOS
aVFOVIGOdONaHOD
AVAOVIGOdONSHO
@VaDVIGOdONAHO
@VEOVIGOdONaHO
AVAOVINA TdSV
AVAOVINA TdSV
VEOVINATdSV
avadovava
avaovava
AVaOVUALSV
aVaOVAaLSV
AVAOVUAELSV
AVAOVUALSV
AVSOVUELSV
aVeOVAaLSV
aVeOVaALSV
AVIOVUALSV
SVSOVUALSV
AVEOVUALSV
aVaOVUALSV
SVAOVUELSV
aVeOVUaLSV
AVEOVURLSV
AVEOVUALSV
AVEOVAALSV
AVAOVUALSV
SVEOVUELSV
AVEOVUALSV
AVaOVUALSV

wanes

ene

BOTUTaITA

‘suadiquie “sea eLrejnoipad
pay

wasor

eqnyed

(ereafdsqns -y) syes07sH1
Bajuasie

soploadxjouy
uodsesouakd
wuyway-xT}oy

smoaj3ou

sisuapeues

smejnpun

smejjaquin

snyejnqns

iuoys

Hoqaryos

snyojnudes

(snouruita “y) snsowaoes
snaotund

soproyjueuard

smjeoed

toyduud “sea snsopid
snsoyid “sea snsopid
snusajed

snyjoyrsoyyd “sea suared
suoqed “38a suojed
(snaimze “y) stsuai3ueyuajoo
‘oer[Sue-seaou
snyjAydosseus
‘SNUBOTIMOT

smoyuarey

PEEEEEEEEEE

aenuanranaamnmoarnmntonrnnSnacocconSace

Appendix A Checklist of Vascular Plants

A8


aVaOVIdV
AVaOVaaLSV
AVAOVOVAIXVS
AVAOVUELSV
aVEOVUALSV
AVEOVUALSV
AVAOVUALSV
aAVEOVAELSV
SVEOVUALSV
AVEOVOISSVUd
SVEOVIONAd
AVAOV1ONAd
AVEOVIGOdONAHD
AVAOVIGOdONaHO
AVAOVIGOdONAHO
HVAOVIGOdONaHO
AVAOVIGOdONSHD
VEOVIGOdONSHD
YVEOVIGOdONAHO
AVAOVIGOdONSHD
HVAOVIGOdONSHD
AVEOVIGOdONEHD
VEOVIGOdONSHO
aVEOVNIVINHdOYDS
AVEOVUAAVdVd
aVeOVITIT
avaovorg
AVAOVINIdTVSaVO
SVEOVIdV

. aVaOVIdv
AVEOVIAV INHdOUS
avaovsou
AVAOVINIdTVSaVO
AVAOVTIAHdOLVaaO

BIOFGING

snqAyuy

wnuespoure

uwmusodure>

unuerurgita

winjuaipsed

umnuwpxeur

wauwaljuEonay

Bypuesyeq

‘BypaUa

Heyjequin

wejnoew

Bpreayna

‘umoiqan

uinueAa[puels

ayeanut

winyjAydorday

umones

(uinpugdy *D) wnuadsoyuedi3
: winyendes
skayoq

sapjoysoquie

wings

eiqeis

snfeur

wmnainy

wyeynacyeo

(eisuoseureyo visseD) BIeINOTOSey
Muoys “rea suaquinsoid
suaquinsoid “sea suaquinooid
snupu

Byeuade]

sisuapeues

uinjeuTyoo

mynd
umpopy)
umruadsosAsy,
sysdost1y
wmnuoSosA1y
umuayjuesfiyD
umuayquesiiys
winweayjuestiyy
wawarjuestiys
wiodspi0yD
epydeany
epydewry
uimnypodouayp,
winypodousy
winypodousyy
umypodouay)
umnipodouayD
uinypodouaya
winipodouayp
uwmypodouay
umypodouay)

wsuooeureyD
uinyjAydosaey)
uunyt<ydoxoey
urnuyyLioUseY
Sapsurouaey)
stNaQ)
uinyyAydoyesaz

moocotvamnZwmomeseolnmowowcannccocowvowocry

Ree eee ee oe

oe

A17

Appendix A Checklist of Vascular Plants



aVaOV INLad
aVaOV INA
EVES) ela
aVaOVITIT
AVSOVITIT
aVaOVITIT
aVSOVITIT
avaOviIlT
aAVeOVOISSvad
AVEOVLVWSTIV
AVAOVLVASITV
aVaOVITIT
AVEOVATVN
AVEOVINVI
avaOvanowvNis
avaovOd
aVaOVOd
avaOvOd
aVaOVOd
avaovod
AVSOVTIAHdOAUVO
avadoOvsou
avadovsou
avaOvsod
avaOvsou
avaovsod

aVeOVINVIT .

aVaOVINVT
AVIOVNIV TNHdOADS

AVEOVINVINHOWOS

AVeOVNIV INHdOWOS
aVEOVNIV INHdOUDS
AVEOVRIV INHdONS

aVaOVIdV

(esonu “y) eueout
esoupnnys

ayeauya

ums000m
umsesdous0yps
umapes

winnuJeo

asuopeues

wyeoped

area
(vonenbe-odeiueyd “y) wimepsooqns
BsoUey

wasor

suejder

wuNsspIe

sueuuazed

waqeos “sea stfeuraky
syewoxy “sea stpewady
eaqued]3

(smnuay y) spxeinydeo
odes

‘eyoyaeurernydosss
sopioyedou

Byoyuay

BuepoUUTyS

vamdind “sea vamdind
‘eiopiased “rea vandind
(2 wayURWIO]) BIeTMOUNE
urnydeuds

snury
snuqy
wnTTy
wanly
wnTy
wnyty
unity
wanry
epeny
RUSTY
oust
smnery
wary
wanfy
snupueyyy

cwmoSaoSntry~mrannnnr scoonmntoooqonnwmomnoonoonw

A3

Appendix A Checklist of Vascular Plants



3 Application of Coefficient of
Conservatism to Floristic
Quality Assessment System

Following Swink and Wilhelm (1979) and Wilhelm and Ladd (1988), the
coefficients of conservatism can be used to arrive at a numerical value called
the Floristic Quality Assessment Index (I). This numerical value provides a
floristic based assessment of the natural area related to the degree of artificial
disturbance indicated by the presence of non-native or opportunistic native
taxa. The floristic quality assessment indices from different types of vegeta-
tion can be objectively compared. The index value does not imply that one
type of vegetation is “better” than another; it simply provides a way of mea-
suring the degree of naturalness of the species found there. The floristic
quality assessment index is also useful in comparing how vegetation changes
over time, either from natural succession or from management. In this situa-
tion, a repeatable vegetation sampling method would be used in conjunction
with the floristic quality assessment index.

The application of this method requires field sampling by an experienced
field biologist able to discern the subtle differences in the floristic elements.
Following Wilhelm and Ladd (1988), the floristic quality assessment is con-
structed in the following manner:

a. Compile a list of the plants growing in the area to be assessed, inde-
pendent of community types.

b. Assign coefficients of conservatism to each plant listed (Appendix A).

c. Determine the mean coefficient value by adding the coefficients of
native plants recorded from the area, and dividing the sum by the total
number of native plants.

d. Multiply the mean coefficient by the square root of the total number of
native species.

e. The product obtained is the floristic quality assessment index (1).

Chapter 3 Application of Coefficient of Conservatism to Floristic Quality Assessment System


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A41

Appendix A Checklist of Vascular Plants



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AS7

Appendix A Checklist of Vascular Plants


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SOMNTONONMOVANDA

Appendix A Checklist of Vascular Plants

A25


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avaoviana
avaoviana
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avaoviana
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avaoviana
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me

ee ee ee

A27

Appendix A Checklist of Vascular Plants


4 Conclusions

The floristic quality assessment index (index of conservatism) for northern
Ohio was developed as a tool to assess the nativeness of an area based on the
presence of conservative species. The floristic quality assessment index
allows for an objective numerical comparison of two or more unrelated com-
munity types for the occurrence of higher quality assemblages of species,
impacts by human disturbance reflected in the presence of alien species, or the
capability to assist with calibration of the vegetation component of wetland
functional indices. ‘It allows for an objective numerical comparison of two
unrelated community types and reflects numerically the impact of human
disturbance by taking into account the presence of alien taxa.

Numerical values included in this report become less valid outside of the
study area for several reasons. These include changes in species distribution
patterns, abundance, and changes in habitat. Values for coefficient of conser-
vatism are available for other areas outside of northern Ohio, including the
state of Michigan (Herman et al. 1993) and northern Illinois (Swink and
Wilhelm 1979, 1994). Michigan (Herman et al. 1993) has compiled for pub-
lication a Floristic Quality Assessment Index applicable to the entire state.

The floristic quality assessment index does provide a repeatable method for
monitoring changes in species composition over time, evaluating wetland
functions, natural area acquisition, selection of land management techniques,
assessing the success of restoration efforts, designing and monitoring mitiga-
tion, and in evaluating wetlands. The results of land management, whether it
be for mitigation or for restoration, require monitoring and evaluation. This
report presents the background, the coefficient of conservatism values, and the
steps to follow in order to establish a numerical rating for the floristic quality
of plant communities in northern Ohio.

Chapter 4 Conclusions