Coral Zonation of the Mississippian System 

in the Western Interior Province 

of North America

U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 1334
Coral Zonation of the Mississippian System 

in the Western Interior Province 

of North America

By WILLIAM J. SANDO and E. WAYNE BAMBER

U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 1334

A biostratigraphic zonation of the Mississippian 

rocks of parts of the Western United States 

and Western Canada, based on corah

UNITED STATES GOVERNMENT PRINTING OFFICE, WASHINGTON: 1985
DEPARTMENT OF THE INTERIOR 

DONALD PAUL HODEL, Secretary

U.S. GEOLOGICAL SURVEY 

Dallas L. Peck, Director

Library of Congress Cataloging in Publication Data

Sando, William Jasper.

Coral zonation of the Mississippian System in the Western Interior Province of North America.

(Geological Survey professional paper; 1334)

Bibliography: p.

Includes index.

Supt. of Docs, no.: I 19.16:1334

1. Corals, Fossil. 2. Paleontology The West Mississippian. 3. Paleontology Canada, Western Mississippian. I. Bamber, E. W. II. Title. III. Series. 

QE778.S23 1984 563'.6'0978 84-600130

For sale by the Distribution Branch, U.S. Geological Survey 

604 South Pickett Street, Alexandria, VA 22304
CONTENTS

Page

Abstract _.___ ._______..__--___._.__.----___. ------- --. _- - -- ----- .~.~ .--- --- -- 1

Introduction ...........................................,................. ...--.....-- ----.- ------- -- 1

Acknowledgments ...........................................-........ .-....... - --- - ---. 2

Previous work ................................................-..-....-..... .-..-..... -... --- . 4

Procedure ..............................-..........-.............................- - --.------.- ----- 4

Definition of coral zones and subzones ...--........---.....-.... ......-.. --.....-.- ..---.--- 9

Influence of environment on zonation .........................-......--......--....-- - .. ---- 9

Description of Coral Zones I-VI -.....................................-.... ....---...---- ---. -----. 9

Applications of the coral zonation ______._.__._.._______- ._._. ---- -~~. ~ --~-~- . 14

Zonal taxa ......................................-.......-...........-..........-- ..---- - --- .- -----.. 14

Locality register .............................. -....---.............---... ... ..--- -. ------32

United States ............................................................ ... .. - 32

Canada ........................... ........................ ......---- ------.--- ----- -- --.44

References cited ........................................................... -... -- -- ---- -- -..56

ILLUSTRATIONS

	[Plates follow references cited]

PLATE 1. Acrocyathus, Amplexizaphrentis, Amplexocarinia, Amplexus, Ankhelasma, Aulostylus.

2. Barytichisma, Beaumontia, Canadiphyllum, Caninia, Guerichophyllum, Siphonophyllia.

3. Cleistopora, Clisiophyllum, Cyathaxonia, Dorlodotia?, "Diphyphyllum."

4. Duncanopora, Ekvasophyllum, Faberophyllum.

5. Hapsiphyllum, Koninckophyllum, Liardiphyllum, Siphonodendron.

6. Actinocyathus, Lophophyllum?, Metriophyllum, Siphonodendron.

1. Michelinia, Multithecopora, Neaxon?, Palaeacis, Petalaxis.

8. Pleurosiphonella, "Pseudodorlodotia," Rhopalolasma, Rotiphyllum, Schoenophyllum.

9. Sciophyllum, Stelechophyllum, Sychnoelasma, Syringopora.

10. "Thysanophyllum," Turbinatocaninia?, Vesiculophyllum, Zaphriphyllum.

Page

FIGURE 1. Map showing Mississippian coral localities in the Western Interior Province of Canada .............................................. 2

2. Map showing Mississippian coral localities in the Western Interior Province of the United States ................................ 3

3. Correlation chart comparing new Western Interior coral zonation with other coral zonations (since 1960) in the United States

and Canada and with foraminifer and conodont zones .- .._.____.._.__-_____.______._______.___._.______.._____._________.__..____.______..__ 4

4. Chart showing composite ranges of coral taxa in the Western Interior Province of the United States, and foraminifer, conodont, and coral zones in relation to North American Mississippian provincial series and western European Carboniferous 

stages ......................................................................................................................................................... 5

5. Chart showing composite ranges of coral taxa in the Western Interior Province of Canada, and foraminifer and coral zones

in relation to North American Mississippian provincial series and western European Carboniferous stages. 6

6. Chart showing combined composite ranges of coral taxa in the Western Interior Province of the United States and Canada, 

and foraminifer, conodont, and coral zones in relation to North American Mississippian provincial series and western 

European stages ................. --.................................................................................................................... 7

7. Composite range chart in which coral taxa are arranged in ascending order of bases of ranges _.-- __.__.___ --_._.__-_-. __ 8

8. Distribution of Mississippian lithofacies with respect to biostratigraphic zonations and generalized environmental model 10

9. Lithofacies distribution and shallow-water habitat index (SWHI) of coral taxa ............................................................ 11

10. Chart showing correlation of Mississippian stratigraphic units in selected areas of the Western Interior Province of the United

States and Canada by means of foraminifer, conodont, and coral zones __._.- ._____._._.__-_____._.._.._._____.___._________._________ 15

in

CORAL ZONATION OF THE MISSISSIPPIAN SYSTEM IN THE WESTERN 

INTERIOR PROVINCE OF NORTH AMERICA

By WILLIAM J. SANDO and E. WAYNE BAMBER'

ABSTRACT

Analysis of the distribution of coral taxa, principally genera and 

subgenera, in the Mississippian of the Western Interior Province of 

Canada and the conterminous United States permits recognition of 

a system of coral zones and subzones useful for correlating stratigraphic 

units throughout an area extending from the southwestern District 

of Mackenzie in the Northwest Territories of Canada to southern 

California in the United States. The zonation comprises six Oppel 

zones, four of which are divided into locally useful subzones. Coral 

Zone I corresponds approximately to the Kinderhookian Provincial 

Series and is divided into Subzones A, B, and C. Coral Zone II corresponds approximately to the Osagean Provincial Series and is divided 

into Subzones A and B. The Tournaisian Stage of western Europe is 

represented by Coral Zones I and II. Coral Zone III corresponds approximately to the lower and middle parts of the Meramecian Provincial Series and is divided into Subzones A, B, C, and D. Coral Zone 

IV corresponds approximately to the upper part of the Meramecian 

and is not divided. Coral Zone V corresponds approximately to the 

lower and middle parts of the Chesterian Provincial Series and is divided into Subzones A and B. Coral Zone VI corresponds approximately 

to the upper part of the Chesterian and is not divided. The Visean 

Stage of western Europe is represented by Coral Zones III and IV and 

Subzone VA. The lower part of the Namurian Stage of western Europe 

is represented by Coral Subzone VB and Zone VI. Deep-water and 

shallow-water coral biofacies are discussed and integrated into the 

zonation system.

The coral zones are correlated with foraminifer and conodont zones, 

which provide the primary bases for establishing ranges of coral taxa. 

Forty-five coral genera and subgenera used in the zonation are diagnosed, discussed, and illustrated. Correlations of Mississippian formations in the Western United States and Western Canada are presented 

as examples of the utility of the coral zonation.

INTRODUCTION

Although planktonic and nektonic groups such as 

conodonts and cephalopods are generally regarded as the 

best zonal fossils for biostratigraphy in the Carboniferous, corals are among the most widely used 

groups of sessile benthos for zonation and correlation 

because of their abundance in widespread shallow-water 

carbonate lithofacies, particularly in the Mississippian.

'E. W. Bamber: Geological Survey of Canada, 3303 33d St., N.W., Calgary, 

Alberta, Canada T2L 2A7.

The widespread distribution of certain coral biofacies is 

evidently the result of their free-swimming larval stage, 

which in some modern corals may be as long as 2 months 

(Sando, 1977b, p. 490). Within parts of North America, 

the geochronometric resolution of Mississippian coral 

zones compares favorably with that of zones based on 

foraminifers, conodonts, and cephalopods (Sando, 1977a, 

p. 10; 1977b, p. 494), but corals, because of their greater 

provinciality, are less useful than these more rapidly 

and widely dispersed groups of fossils for detailed 

worldwide zonation and correlation. North American 

Mississippian corals are more abundant and widely 

distributed than conodonts and cephalopods in shallowwater carbonate facies, and they are more abundant and 

widely distributed than foraminifers in deep-water carbonate facies. Corals are more practical than 

microfossils for field biostratigraphy because many taxa 

useful for zonation can be identified with a hand lens, 

whereas microfossils can seldom be identified in the 

field. Corals are also less susceptible to destruction by 

recrystallization and dolomitization than are 

foraminifers.

Studies of the distribution of North American 

Mississippian corals (Sando, Bamber, and Armstrong, 

1975,1977) indicated high degrees of similarity between 

the coral faunas of western Canada and those of the conterminous western United States throughout Mississippian time, owing to continuity of shallow-water environments conducive to gene flow. These studies led to 

recognition of a large zoogeographic province, the 

Western Interior Province, which includes the 

southwesternmost District of Mackenzie, northeastern 

and southeastern British Columbia, western and 

southern Alberta, and southern Saskatchewan in 

Canada (fig. 1), and northeastern Washington, Montana, 

western North and South Dakota, Wyoming, 

southeastern and south-central Idaho, eastern Nevada, 

Utah, western Colorado, southern California, and most 

of Arizona in the United States (fig. 2). Areas of coralliferous Mississippian rocks in northern and 

southeastern Alaska, the northern Yukon, northwestern 

Washington, central Oregon, and northern California 

were in different zoogeographic provinces during the
CORAL ZONATION, MISSISSIPPIAN SYSTEM, WESTERN INTERIOR PROVINCE

130° 100°

CANADA 

UNITED STATES

Fernie .»

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100 200 300 400 MILES

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0 100 200 300 400 500 KILOMETERS

FIGURE 1. Mississippian coral localities in the Western Interior Province of Canada. X, coralliferous stratigraphic section; dot, locality not 

in stratigraphic section; dashed line, boundaries of Western Interior Province; circled numbers, location of selected stratigraphic sequences 

shown on figure 10. Facies boundaries are from Macqueen and Bamber (1967) and Bamber, Macqueen, and Richards (1980).

Mississippian and are not considered in this report.

Preliminary tabulation of the stratigraphic ranges of 

genera and subgenera found in the United States and 

Canada in the Western Interior Province indicated that 

a unified Mississippian coral zonation is feasible for the 

entire area. A preliminary version of this new zonation 

was presented at the Ninth International Congress of 

Carboniferous Stratigraphy and Geology (Sando and 

Bamber, 1979, and in press). The present report is a 

more comprehensive account of the unified coral zonation for the Western Interior Province of North America.

ACKNOWLEDGMENTS

Stratigraphic ranges of coral taxa in this report are 

based mainly on correlations by means of foraminifer 

and conodont zones. We are greatly indebted to B. L. 

Mamet, Universite de Montreal, Montreal, P.Q., 

Canada, and C. A. Sandberg, U.S. Geological Survey, 

Denver, Colo., for zonal identifications of foraminifers 

and conodonts, respectively; many of these identifica-
INTRODUCTION

110° 100°

30°

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300 400 MILES

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FIGURE 2. Mississippian coral localities in the Western Interior Province of the United States. X, coralliferous stratigraphic section; dot, 

coral locality not in stratigraphic section. Circled numbers, locations of selected stratigraphic sequences shown on figure 10. Major paleotectonic units from Poole and Sandberg (1977, fig. 1). Craton margin indicated by dashed line. Western Interior Province includes Antler 

erogenic highland (part), foreland basin, and cratonic platform.

tions are unpublished. We are also grateful to A. E. H. 

Pedder, Geological Survey of Canada, Calgary, Alberta, and J. T. Dutro, Jr., U.S. Geological Survey.

Washington, B.C., for helpful comments on the 

manuscript.
CORAL ZONATION, MISSISSIPPIAN SYSTEM, WESTERN INTERIOR PROVINCE

PREVIOUS WORK

The coral zonation described in this paper is part of 

an evolution of zonal concepts based on earlier work. 

Previous zonations in the Western Interior Province 

were summarized by Sando (1977a,b). In figure 3, the 

new unified zonation for the Western Interior Province

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1977d, 1983) laid the taxonomic groundwork.

PROCEDURE

This analysis is based on the distribution of Mississippian corals at 360 localities in the United States (fig. 

2), of which 155 are detailed measured stratigraphic sections, and at 175 localities in Canada (fig. 1), of which 

157 are detailed measured stratigraphic sections. The 

localities are described in the locality register at the end 

of the report. In the United States, an estimated 10,000 

specimens were assigned to 45 genera or subgenera; in 

Canada, an estimated 8,000 specimens were assigned 

to 34 genera or subgenera. Zonal taxa are diagnosed and 

discussed in another part of this report.

The occurrences and ranges of coral taxa were plotted on lithic logs along with boundaries of stratigraphic 

units, foraminifer zones of B. L. Mamet (Mamet and 

Skipp, 1970a,b), and conodont zones of C. A. Sandberg 

(in Sando, Sandberg, and Gutschick, 1981). Correlation 

charts were prepared for principal areas of coral occurrences, and stratigraphic units were correlated by means 

of foraminifer and conodont zones and, to a limited extent, by physical stratigraphy.

Data on the tops and bottoms of coral ranges in each 

stratigraphic section with respect to lithostratigraphic 

units, foraminifer zones, and conodont zones were colFIGURE 3. Correlation chart comparing new Western Interior 

coral zonation with other coral zonations (since 1960) in 

the United States and Canada and with foraminifer and 

conodont zones.
PROCEDURE

FIGURE 4. Composite ranges of coral taxa in the Western Interior Province of the United States, and foraminifer, conodont, and coral zones 

in relation to North American Mississippian provincial series and western European Carboniferous stages. X, single occurrence; query, 

questionable occurrence; dotted line, interval of no occurrence joining intervals of occurrence; dashed line, continuous rare occurrence; 

solid line, continuous common to abundant occurrence.

lated and tabulated for each taxon, as were similar, less 

precise data on occurrences of coral taxa at localities not 

in detailed stratigraphic sections. Ranges were 

crosschecked by examining associations of taxa found 

at isolated localities.

Charts showing composite ranges of taxa in the United 

States (fig. 4) and Canada (fig. 5} were compiled by integrating the range data from individual stratigraphic 

sections. Variations in abundance (frequency of occurrence) at different stratigraphic levels were also noted
CORAL ZONATION, MISSISSIPPIAN SYSTEM, WESTERN INTERIOR PROVINCE

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PROCEDURE

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See explanation on figure 4 for definition of occurrence symbols.

and are shown on these composite charts. The composite 

range of each taxon established from measured sections 

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Composite range charts for the United States and 

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charts for the United States and Canada. Important 

discontinuities in range or frequency of occurrence of
CORAL ZONATION, MISSISSIPPIAN SYSTEM, WESTERN INTERIOR PROVINCE

taxa were given numerical values dependent on the 

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bases of ranges (fig. 7).

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FIGURE 7. Composite range chart in which coral taxa are arranged in ascending order of bases of ranges. See explanation on figure 4 for 

definition of occurrence symbols.
DEFINITION OF CORAL ZONES AND SUBZONES

DEFINITION OF CORAL ZONES AND SUBZONES

The coral zones are Oppel zones as described by 

Hedberg (1976, p. 57-58). Each zone is characterized by 

an association of selected taxa of restricted and largely 

concurrent range, and the boundaries of the zone are 

marked by first or last appearances or by changes in 

abundance (frequency of occurrence) of selected taxa. Index taxa are mostly genera and subgenera, but species 

are used to a limited extent where their distribution is 

well known.

The zones are divided into subzones that may or may 

not be recognizable in both the United States and 

Canada. The subzones are based on the same criteria 

as the zones but are generally characterized by fewer 

index taxa and are commonly of more limited areal 

extent.

Although the zones and subzones are established here 

as formal biostratigraphic units, the use of fossil names 

for these units has been avoided because the taxonomy 

is in a state of flux and the use of fossil names would 

create an unstable zonal nomenclature. Instead, zones 

are identified by Roman numerals; subzones are identified by letters.

INFLUENCE OF ENVIRONMENT ON ZONATION

The stratigraphic and geographic distribution of coral taxa is influenced by differences in the environments 

in which the corals lived. In a study of the paleoecology 

of Mississippian corals from the Western Interior Province of the United States, Sando (1980) determined the 

occurrences of 46 taxa in each of six lithofacies representing environments ranging from more than 100 m below 

sea level in deep basins far from shore to a few meters 

or less below sea level in shoals and tidal flats at the 

shoreline (fig. 8). The taxa analyzed in Sando's study are 

the same as those used in the zonation system described 

herein. Although no similar analysis has been made for 

Canada, we assume that the conclusions are valid for 

Canada as well.

Sando (1980) grouped the occurrences of coral taxa into 

deep-water and shallow-water facies and derived a 

shallow-water habitat index (SWHI) that represented 

the percentage of occurrences of each taxon in shallowwater lithofacies. Thus, taxa having SWHI less than 50 

were regarded as deep-water biofacies, and those having SWHI greater than 50 were regarded as shallowwater biofacies. The biofacies classification and 

lithofacies distribution of all the taxa used in zonation 

are summarized in figure 9.

Recognition of deep-water and shallow-water biofacies 

has an important bearing on zonation because some 

zones that represent the same time interval are

represented by different assemblages of taxa in different 

geographic areas. Lack of understanding of this relationship caused incorrect correlations of some stratigraphic 

units in the past. In the description of the coral zones 

presented below, biofacies factors are discussed and integrated into the zonation concepts.

DESCRIPTION OF CORAL ZONES I-VI 

CORAL ZONE I

The lowest zone is a generally poorly coralliferous unit 

that corresponds approximately to the Kinderhookian 

Provincial Series. Coral Zone I corresponds to five 

Siphonodella conodont zones of Sandberg and others 

(1978) and includes Foraminifer Zone pre-7 (fig. 6). 

Although the United States-Canada composite range 

charts (figs. 6 and 7) show ,17 genera in this zone, only 

3 genera have been found in the unit in Canada (fig. 5), 

and 10 genera are rare in the unit in the United States 

(fig. 4). Metriophyllum, although known from the older 

Devonian, is unknown immediately below Coral Zone 

I and is not known above it. Amplexocarinia may be confined to the zone. Other genera that are not known below 

this unit in the Western Interior Province are Amplexizaphrentis, Beaumontia, Guerichophyllum, Cyathaxonia, Lophophyllum?, Palaeacis, Neaxon?, and 

Rotiphyllum.

In the United States, the base of Coral Zone I is indistinctly marked by the first appearance of 

Lophophyllum?, which coincides with the base of the 

sulcata Conodont Zone. Amplexus, Vesiculophyllum?, 

and Syringopora are rare in the lower part of the coral 

zone but are also known from uppermost Devonian (upper Famennian) beds beneath the zone. For practical 

purposes of correlation, the base of Coral Zone I is identified by conodont correlation of the base of the Mississippian because of the rarity of corals immediately above 

and below the Devonian-Mississippian boundary.

Subzones A, B, and C of Zone I are recognizable in 

parts of the United States but not in Canada. Subzone 

IA includes rare Lophophyllum?, Amplexus, and 

Vesiculophyllum? found in the middle part of the Fitchville Formation (blue shaly limestone unit) of central 

Utah and possibly in the lower part of the Tin Mountain Limestone of southern California. Elsewhere in the 

Cordilleran region of the United States, this 

stratigraphic interval is missing or represented by noncoralliferous strata. In Canada, the interval is 

represented partly by hiatus and partly by noncoralliferous terrigenous beds in the Yohin, Besa River, 

Bakken, and Exshaw Formations. Subzone IA corresponds to the sulcata, duplicata, and sandbergi Conodont Zones and the lower part of the Lower crenulata 

Conodont Zone.
10 CORAL ZONATION, MISSISSIPPIAN SYSTEM, WESTERN INTERIOR PROVINCE

Subzone IB is marked by a distinctive assemblage consisting of Amplexizaphrentis, Amplexocarinia, 

Amplexus, Beaumontia, Guerichophyllum, Cyathaxonia, 

Lophophyllum?, Metriophyllum, Michelinia, Palaeacis, 

Neaxon?, and Syringopora, Vesiculophyllum does not occur with this assemblage but is present in the same interval. The base of this subzone is marked by the first

occurrence of eight genera. Metriophyllum and a new 

species of Palaeacis are confined to the subzone. The 

principal occurrence of this assemblage is in transgressive glauconitic crinoidal limestone that marks the 

base of the Paine Member of the Lodgepole Limestone 

in southwestern Montana, southeastern Idaho, and 

northern Utah. The subzone may be more widespread

ENVIRONMENTAL MODEL

LAND

Adetognathus 

unicornis

naviculus

VI

16s

15

14

13

12

11

10

Cavusgnathus

IV

texanus

anchoralislatus

typicus

isostichaUpper 

crenulata

sandbergi

duplicata

sulcata

HIATUS HIATUS HIATUS HIATUS HIATUS

FIGURE 8. Distribution of Mississippian lithofacies with respect to biostratigraphic zonations and generalized environmental model (modified from 

Sando, 1980). A, deep-water basinal terrigenous facies; B, deep-water basinal carbonate facies; C, deep-water bank carbonate facies; D, shallow-water basinal carbonate facies; E, shallow-water shelf carbonate facies; F, red-bed facies; G, evaporite and evaporitic carbonate facies; 

H, shallow-water terrigenous facies. Coralliferous lithofacies are stippled; noncoralliferous lithofacies are shaded. Nonpatterned intervals indicate absence of lithofacies. Environmental model not to scale; vertical dimension greatly exaggerated.
DESCRIPTION OF CORAL ZONES I-VI 11

than is presently known, because the small size of most 

of the index taxa makes recognition difficult except in 

silicified collections. In Canada, this interval is 

represented by noncoralliferous terrigenous beds in the 

Besa River, Clausen, and Banff Formations. Subzone IB 

corresponds to the upper part of the Lower crenulata 

Conodont Zone.

Subzone 1C is a poorly coralliferous interval between 

the top of subzone IB and the top of Coral Zone I. Nine 

genera range into this subzone from below, and only two 

taxa, Cleistopora placenta (White) and Sychnoelasma 

(very rare) first appear in the unit. In the United States, 

two biofacies may be distinguished a shallow-water 

biofacies characterized by rare Amplexizaphrentis, 

Lophophyllum?, Syringopora, and Vesiculophyllum, and 

a deep-water biofacies characterized by Amplexizaphrentis, Amplexus, Guerichophyllum, Cleistopora (rare), 

Cyathaxonia, Lophophyllum?, Rotiphyllum, and Syringopora (rare) (see Sando, 1980). The deep-water 

biofacies is found in dark, thin-bedded silty limestone 

of the Paine Member of the Lodgepole Limestone in 

Montana, Idaho, Utah, and western Wyoming. The 

shallow-water biofacies occurs in light-colored crinoidal 

dolomite of the Little Bighorn Member of the Madison 

Limestone in central and eastern Wyoming and in 

shallow-water carbonate rocks of the Whitmore Wash 

Member of the Redwall Limestone in Arizona and the 

lower part of the Pahasapa Limestone in South Dakota. 

In Canada, Subzone 1C is not distinguished but correlates with the upper part of the Banff Formation, 

which contains Amplexizaphrentis, Syringopora, and 

Vesiculophyllum; it also correlates with noncoralliferous 

terrigenous beds in the Besa River and Clausen Formations. Subzone 1C corresponds to all but the highest part 

of the isosticha-Upper crenulata Conodont Zone.

CORAL ZONE II

The second zone is a generally abundantly coralliferous unit that corresponds approximately to the 

Osagean Provincial Series. The top of the zone correlates 

with the top of the Tournaisian Stage of western Europe. 

Coral Zone II corresponds to Foraminifer Zones 7,8, and 

9 and to one conodont zone and parts of two others (fig. 

6). The United States-Canada composite range charts 

(figs. 6, 7) show 29 genera (6 questionable) in this zone, 

26 (5 questionable) of which occur in the United States 

(fig. 4) and 16 (1 questionable) in Canada (fig. 5). 

Aulostylus is the only genus confined to this unit, but 

three others are not known above it, and 13 others are 

not known below it.

The base of Coral Zone II is generally well marked in 

the United States by the first appearances of

Stelechophyllum microstylum (White) and Michelinia expansa White, and, in most places, the first appearances 

of Sychnoelasma and Vesiculophyllum. This 

stratigraphic level corresponds to the base of 

Foraminifer Zone 7. In Canada, the base of Coral Zone 

II is indistinctly marked by the first appearances of 

Cleistopora, Cyathaxonia, and Michelinia, all of which

TAXA 

(NUMBER OF LOCALITIES)

Solitary Rugosa (*dissepimented)

r ;'$i&^$$tim'&i' " ' ' " ' ' " : "

i ' : ifgax(mM:

KlwpaJ&tas^iia'Wty '.'.'""'""'

-. , ^^plexwarimk (75 s

. . ;Cya^t,mnm E(8$)

; . tAm^ismj&l) . '." ' ' ""'

'.. »Guerich^hyUwriM^:- ...

 &»%%Jfawaii)L , "". .. ..." " ..

*Koninckophyllum (2)

Canadiphyllum (16)

Amplexizaphrentis (70)

Lophophyllum ? (43)

*Caninia (8)

*Ekvasophyllum (14)

*Faberophyllum (14)

Sychnoelasma (62)

*Vesiculophyllum (84)

Siphonophyllia (25)

Ankhelasma (8)

Barytichisma (1)

*Clisiophyllum (1)

Hapsiphyllum (4)

*Liardiphyllmn (1)

*Turbinatocaninia? (8)

*Zaphriphyllum (17)

Colonial Rugosa

Acrocyathus (5)

Aulostylus (4)

"Diphyphyllum" (31)

Dorlodotia? (1)

Siphonodendron (22)

Actinocyathus (5)

Petalaxis (1)

'"Pseudodorlodotia" (7)

Schoenophyllmn (1)

Sciophyllum (1)

Stelechophyllum (27)

"Thysanophyllum " (2)

Tabulate

; ' B^mtAM&d'^, . . . ., : ,.. ,."! ..,....,......,

=' .^atimMt^Mr.'.. ....... ... ... ... ..

Cleistfrpora (9)

Michelinia (34)

Syringopora (78)

Duncanopora (11)

Multithecopora (12)

Pleurosiphonella (19)

I 

CO

0

5

6

17

18

29

33

33

50

66

74

77

91

95

96

97

98

99

100

100

100

100

100

100

100

100

100

100

100

100

100

100

100

100

100

100

100

0

11

64

86

95

100

100

100

LITHOFACIES

DEEP 

WATER

B

19

18

15

10

95

60

20

18

3

103

29

1

3

13

12

1

2

25

5

9

19

C

1

1

10

4

3

SHALLOW WATER

D

1

2

13

19

10 j

2

137

51

2

189

245

1

1

3

41

9

36

127

E

1

8

5

7

1

25

167

46

8

52

66

178

247

64

10

1

10

3

12

31

10

1

42

1

33

9

1

10

1

1

7

2

3

20

205

17

21

31

F

1

H

2

3

3

FIGURE 9. Lithofacies distribution and shallow-water habitat index 

(SWHD of coral taxa (modified from Sando, 1980). Number in parenthesis after each taxon is number of localities from which taxon 

has been recorded. Number in column under each lithofacies type 

is number of occurrences of taxon in that lithofacies. Taxa in each 

major group are arranged in increasing SWHI. Stippled taxa are 

predominantly deep-water forms (SWHI <50). Asterisks denote 

dissepimented solitary forms.
12 CORAL ZONATION, MISSISSIPPIAN SYSTEM, WESTERN INTERIOR PROVINCE

are rare. Stelechophyllum, Sychnoelasma, and abundant 

Vesiculophyllum do not appear until the middle of 

Foraminifer Zone 7, which marks the lowest level of 

favorable carbonate facies in the Canadian sequence.

Subzones IIA and IIB are recognized in both Canada 

and the United States. In Canada, the boundary between 

the subzones is marked by the first appearances of 

Aulostylus, Siphonodendron, Lophophyllum?, 

Stelechophyllum, Sychnoelasma, and in most places the 

first appearances of Syringopora and Vesiculophyllum. 

In the United States, this boundary is indistinctly 

marked by the first appearances of Aulostylus and Hapsiphyllum (very rare), and the disappearance of 

Beaumontia. A deep-water biofacies characterized by 

Amplexizaphrentis, Amplexus, Guerichophyllum, 

Cyathaxonia, Michelinia sp., Rhopalolasma?, and 

Rotiphyllum represents Subzone IIB in the phosphatic 

member of the Deseret Limestone in Utah and in 

equivalent beds in Idaho (see Sando, 1980). The more 

widely distributed shallow-water biofacies of Subzone 

IIB is characterized by Amplexizaphrentis, Aulostylus, 

Lophophyllum?, Michelinia expansa White, 

Stelechophyllum, Sychnoelasma, Syringopora, and 

Vesiculophyllum.

In the United States, Coral Zone II is best represented 

in the Woodhurst Member of the Lodgepole Limestone 

and the lower part of the Mission Canyon Limestone of 

Montana. In Canada, the zone includes the lower part 

of the Flett Formation in southwestern District of 

Mackenzie, the upper part of the Banff Formation, the 

lower part of the Livingstone Formation, the Pekisko 

and Shunda Formations of Alberta and British Columbia, and the Lodgepole and Mission Canyon Limestones 

of Saskatchewan. It correlates with noncoralliferous terrigenous beds in the upper part of the Besa River and 

Clausen Formations and the lower part of the Prophet 

Formation of northeastern British Columbia and 

southwestern District of Mackenzie.

CORAL ZONE III

The third zone is a thick unit that correlates approximately with the lower and middle parts of the Meramecian Provincial Series and the lower and middle parts 

of the Visean Stage of western Europe. Coral Zone III 

corresponds to Foraminifer Zones 10,11,12,13, and the 

lower half of 14 and to one conodont zone and parts of 

two others (fig. 6). The United States-Canada composite 

range charts (figs. 6 and 7) show 33 genera in this zone, 

31 of which occur in the United States (fig. 4) and 23 

in Canada (fig. 5). Ankhelasma is the only genus confined to the zone, but 11 others are not known above it, 

and 7 others are not known below it.

| The base of Coral Zone III in the United States is well 

marked in the shallow-water biofacies by the first appearances of Ankhelasma and "Diphyphyllum," in most 

places by the first appearances of Siphonodendron and 

Zaphriphyllum, and by the disappearance of 

Sychnoelasma at most localities. This stratigraphic level 

corresponds to the base of Foraminifer Zone 10, which 

marks the base of the Visean Stage and is the best approximation of the base of the Meramecian Provincial 

Series in the Western Interior Province. In Canada, the 

base of Coral Zone III is generally well marked by the 

first appearances of Ankhelasma and Siphonodendron 

oculinum Sando and by the disappearance of 

Sychnoelasma at most localities.

Zone III is divided into four subzones (IIIA-D). Subzone IIIA corresponds to Foraminifer Zones 10 and 11. 

In both the United States and Canada, it is characterized by Ankhelasma, "Diphyphyllum," Siphonodendron 

oculinum Sando, Syringopora, Vesiculophyllum, and 

Zaphriphyllum. In parts of Utah and Idaho, this interval is characterized by a deep-water biofacies similar to 

that in Subzone IIB (see Sando, 1980).

Subzone IIIB coincides with Foraminifer Zone 12. In 

Canada, the subzone is characterized by a continuation 

of most of the index taxa that occur in Subzone IIIA, but 

Subzone IIIB is marked by the absence of Ankhelasma, 

an abundance of Canadiphyllum, and the first appearance of Acrocyathus (cerioid). In the United States, 

this interval is missing, because of erosion, over most 

of the cratonic platform, except for a shallow-water 

biofacies in the Horseshoe Mesa Member of the Redwall 

Limestone in the Grand Canyon region of northern 

Arizona, where Dorlodotia?, Syringopora, and 

Vesiculophyllum characterize the unit. Ankhelasma, 

"Diphyphyllum," Siphonodendron, and Zaphriphyllm 

are unknown in Subzone IIIB in the United States. In 

parts of Utah and Idaho, the subzone is characterized 

by a deep-water biofacies that consists of Amplexizaphrentis, Canadiphyllum, Cyathaxonia, Rhopalolasma, and Rotiphyllum (see Sando, 1980).

Subzone IIIC coincides with the lower half of 

Foraminifer Zone 13. In the United States, the base of 

this subzone is marked by the first appearance of the 

abundant and widespread genus Ekvasophyllum and at 

most localities by the first appearance of Canadiphyllum 

(new species with widely flaring calice). In Canada, the 

base of Subzone IIIC is less distinctly marked by a break 

in the occurrence of cerioid A crocyathus, an abundance 

of "Diphyphyllum," and renewed occurrence of 

Dorlodotia?

Subzone HID corresponds to the interval from the middle of Foraminifer Zone 13 to the middle of Zone 14. In 

the United States, the base of this subzone is marked
DESCRIPTION OF CORAL ZONES I-VI 13

by the first appearance of Siphonodendron whitneyi of 

Meek and the first appearance of undoubted 

Pleurosiphonella virginica (Butts). Ekvasophyllum, 

Canadiphyllum, and Siphonodendron whitneyi of Meek 

are abundant and widespread, and Faberophyllum, Hapsiphyllum, and cerioid Acrocyathus are rare in this subzone. Rare Sciophyllum is confined to the subzone. In 

Canada, the base of the subzone is marked by the first 

appearances of fasciculate Acrocyathus and Siphonodendron whitneyi of Meek. Ekvasophyllum, Pleurosiphonella, and cerioid Acrocyathus become common at 

this level.

Coral Zone III is represented in a large number of formations in the United States, including the upper parts 

of the Mission Canyon and Redwall Limestones and 

parts of the Deseret Limestone and Little Flat Formation. In Canada, the zone is included in the Turner 

Valley, Mount Head, Prophet, and Debolt Formations, 

and the upper parts of the Livingstone and Flett 

Formations.

CORAL ZONE IV

The fourth zone is a relatively thin unit that correlates 

approximately with the upper part of the Meramecian 

Provincial Series and the lower part of the upper division of the Visean Stage of western Europe. Coral Zone 

IV corresponds to the upper half of Foraminifer Zone 14 

and all of Zone 15, and to part of the Cavusgnathus Conodont Zone. The United States-Canada composite range 

charts (figs. 6 and 7) show 19 genera in this zone, 14 of 

which occur in Canada (fig. 5) and 13 in the United 

States (fig. 4). Although Clisiophyllum is the only genus 

confined to the zone, 12 other genera or subgenera are 

not known above it, and one other subgenus is not 

known below it. Faberophyllum, although not confined 

to the zone, is particularly characteristic of it.

The base of Coral Zone IV is well marked in the United 

States by the disappearances of cerioid Acrocyathus, 

"Diphyphyllum," Dorlodotia?, Siphonodendron whitneyi 

of Meek, Palaeacis cuneiformis Milne-Edwards, 

Rhopalolasma, Sciophyllum, and Stelechophyllum, a 

decrease in the number of occurrences of Canadiphyllum 

and Ekvasophyllum, and the first appearance of 

Clisiophyllum. In Canada, the zone is characterized by 

abundant "Thysanophyllum," fasciculate Acrocyathus?, 

large Ekvasophyllum, and Faberophyllum. Its base is 

marked by the disappearances of Canadiphyllum, 

Caninia, Petalaxis, Stelechophyllum, Siphonodendron 

whitneyi of Meek, and by the first appearance of 

Faberophyllum.

In the United States, Coral Zone IV is best known in 

the lower part of the Monroe Canyon Limestone and

Scott Peak Formation of Idaho and in the Humbug Formation and lower part of the Great Blue Formation of 

Utah. In Canada, the zone is present in the upper part 

of the Mount Head Formation of Alberta and 

southeastern British Columbia and in the lower part of 

the Golata Formation and the upper part of the Prophet 

Formation of northeastern British Columbia. It is partly correlative with noncoralliferous terrigenous beds in 

the lower part of the Mattson Formation of southwestern 

District of Mackenzie and the upper part of the Golata 

Formation of northeastern British Columbia.

CORAL ZONE V

The fifth zone is a thick unit that correlates with the 

lower and middle parts of the Chesterian Provincial 

Series and the upper part of the Visean Stage and lower 

part of the Namurian Stage of western Europe. Coral 

Zone V corresponds to Foraminifer Zones 16i, 16s, 17, 

and the lower half of 18, and to part of the Cavusgnathus 

and lowermost naviculus Conodont Zones. The United 

States-Canada composite range charts (figs. 6 and 7) 

show 15 genera (one questionable) in this zone, all of 

which occur in the United States (fig. 4) and eight (one 

questionable) of which occur in Canada (fig. 5). Duncanopora, Actinocyathus and Schoenophyllum are confined to the zone. Seven other genera or subgenera are 

not known above it and two others are not known below 

it. Siphonophyllia is particularly characteristic.

The base of Coral Zone V is well marked in both the 

United States and Canada by the disappearance of the 

abundant and widespread genera Faberophyllum, 

Clisiophyllum, and Syringopora. In Canada, this 

stratigraphic level is distinguished further by the first 

appearance of Schoenophyllum and by the renewed occurrence of Siphonodendron.

Zone V is divided into Subzones VA and VB in the 

United States but not in Canada. The boundary between 

the subzones is marked by the first appearances ofDuncanopora, Actinocyathus, and Pleurosiphonella drummondi (Nelson), the first appearances of Siphonophyllia, 

and Multithecopora at most localities, and the disappearance of Rotiphyllum.

In the United States, Coral Zone V is best represented 

in the upper part of the Monroe Canyon Limestone and 

in the uppermost part of the Scott Peak Formation and 

South Creek and Surrett Canyon Formations of Idaho. 

In Canada, the zone is present in the Etherington Formation of Alberta and southeastern British Columbia 

and correlates with noncoralliferous terrigenous beds in 

the Mattson Formation of southwestern District of 

Mackenzie and the upper part of the Stoddart Group of 

northeastern British Columbia.
14 CORAL ZONATION, MISSISSIPPIAN SYSTEM, WESTERN INTERIOR PROVINCE

CORAL ZONE VI

The highest coral zone in the Mississippian of the 

Western Interior Province is a very poorly coralliferous 

unit that correlates with the upper part of the 

Chesterian Provincial Series and the lower part of the 

Lower Naniurian of western Europe. Coral Zone VI corresponds to the upper half of Foraminifer Zone 18 and 

all of Zone 19. It also includes the Adetognathus unicornis Conodont Zone and the upper part of the naviculus 

Conodont Zone. The United States-Canada composite 

range charts (figs. 6 and 7) show only six genera in this 

zone, most of which are rare. In the United States, the 

zone includes all six genera (fig. 4), but in Canada, only 

two genera are known (fig. 5). All six of the index genera 

range into the zone from below, and only two of these 

are known above the zone. Petalaxis may be present in 

the zone in the United States.

The base of Coral Zone VI is recognized in the United 

States by the disappearances of Duncanopora, Actinocyathus, Michelinia, and Multithecopora and in 

Canada by the disappearances of Siphonophyllia, Actinocyathus, and Michelinia.

The top of Coral Zone VI is poorly marked because the 

immediately overlying Pennsylvanian beds are generally noncoralliferous. The base of the Pennsylvanian is 

most commonly determined by foraminifers (Zone 20), 

conodonts, or brachiopods.

In the United States, Coral Zone VI is best represented 

in the uppermost part of the Surrett Canyon Formation 

and the Arco Hills and Bluebird Mountain Formations 

of Idaho and in Canada, in the uppermost part of the 

Etherington Formation of Alberta. At most localities, 

this interval is represented by noncoralliferous strata 

or by an hiatus.

APPLICATIONS OF THE CORAL ZONATION

The proposed coral zonation finds its main application 

in the correlation of significant Mississippian sequences 

from particular geographic areas within the Western Interior Province (fig. 10). The composite sections in figure 

10 are based on two or more individual sections for a 

given geographic area. The zonation attains maximum 

utility if used in conjunction with biostratigraphic data 

from other fossil groups. Thus, each zonal boundary, be 

it based on corals, foraminifers, or conodonts, provides 

a correlation datum for comparing stratigraphic sequences and for checking the temporal relationships of 

lithostratigraphic datum planes.

The coral zonation is also useful for determining the 

stratigraphic position of coral collections of unknown or 

uncertain stratigraphic position, such as those made by 

geologists mapping in areas where the local stratigraphy 

is poorly known. By using the range charts (figs. 4-7),

assemblages of corals can be placed stratigraphically, 

with varying degrees of accuracy depending on the taxa 

present. Such determinations are of particular value in 

analyzing the geology of structurally complex areas.

Recognition of the zones and subzones in a given 

stratigraphic section will depend on the abundance and 

diversity of coral taxa and discovery of all taxa actually present. Ranges of individual taxa and assemblages 

of taxa at a single locality are seldom the same as the 

composite ranges upon which the zonation is based. Unfavorable environmental and collecting factors combine 

to make recognition of all possible zonal and subzonal 

boundaries extremely improbable at most localities. No 

single locality affords a complete sequence of zones and 

subzones for the entire Mississippian sequence. At most 

localities, zonal and subzonal boundaries can only be approximated within known limits. Therefore, the coral 

zonation may be limited in its application to detailed 

correlation of individual stratigraphic sections. 

Nonetheless, the zonal scheme provides an important 

biostratigraphic framework, linked to the standard 

foraminiferal and conodont zonations, to which coralliferous Mississippian sequences throughout the North 

American Western Interior Province may be related.

ZONAL TAXA

This section presents diagnoses, discussions, and 

remarks on the distribution of genera and subgenera 

used in the zonation; for convenience, these taxa are 

listed alphabetically. No attempt is made to classify the 

taxa above the generic level. Although the distribution 

of genera and subgenera is generally well known, much 

descriptive work remains to be done on species. Nominal 

species are listed for each genus, and their stratigraphic 

and geographic distribution are given. Morphologic terminology is generally that of Hill (1981). Remarks on 

ecologic occurrence refer to environments recognized by 

Sando (1980). Paleobiogeographic provinces are those 

recognized by Sando, Bamber, and Armstrong (1975, 

1977).

Aulopora Goldfuss, 1829, Cladochonus McCoy, 1847, 

Aulina Smith, 1916, and Favosites Lamarck, 1816, are 

represented in the Western Interior Province but have 

not been included in the zonation. Aulopora and 

Cladochonus are seldom found except as fragments in 

silicified collections. To differentiate these genera from 

each other and from the young stage of Syringopora is 

commonly difficult, if not impossible. The only occurrences of Aulina are in residual chert overlying, and in 

solution cavities in, the Redwall Limestone of Arizona 

(Sando, 1969), where the chert was derived from a 

stratigraphic unit of uncertain age that was removed 

from the sequence by post-Redwall, pre-Supai erosion. 

Favosites is represented by F. whitfieldi White, 1874,
ZONAL TAXA 15

NORTH PROVINCIAL SERIES AMERICAN 

WESTERN EUROPEAN STAGES MAMET FORAMINIFER ZONES

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FIGURE 10. Correlation of Mississippian stratigraphic units in selected areas of the Western Interior Province of the United States and Canada 

by means of foraminifer, conodont, and coral zones. Vertical line pattern denotes hiatus. Closed circles in stratigraphic columns denote occurrences of coral-zone index taxa; open circles denote occurrences of coral-subzone taxa. Shaded parts of stratigraphic columns indicate intervals 

of deep-water lithofacies. Locations of selected areas are shown on figures 1 and2.

which was described from an obscure locality in Arizona. 

It has been found in the Redwall Limestone of Arizona 

and a locality in Nevada, but its exact stratigraphic position is unknown.

Repositories for figured specimens (pis. 1-10) are as 

follows: USNM, U.S. National Museum of Natural 

History, Washington, B.C.; MNA, Museum of Northern

Arizona, Flagstaff, Ariz.; UM, University of Missouri, 

Columbia, Mo.; GSC, Geological Survey of Canada, Ottawa, Ontario. Collections of the U.S. Geological Survey 

are designated by the prefix USGS and the suffix -PC, 

which indicates the upper Paleozoic catalog. Collections 

of the Geological Survey of Canada are designated GSC 

loc.
16 CORAL ZONATION, MISSISSIPPIAN SYSTEM, WESTERN INTERIOR PROVINCE

Genus ACROCYATHUS d'Orbigny, 1849 

Plate 1, figures 5, 6, 10, 11

Diagnosis. Cerioid and fasciculate colonial rugose 

corals. Major septa thin, seldom extending across 

tabularium to columella and ordinarily discontinuous 

or absent in dissepimentarium. Cardinal septum short 

and situated in a fossula formed by downwarped tabulae 

in some species. Counter septum long and extending to 

columella. Minor septa ordinarily absent or poorly 

developed. Columella ranging from a simple axial plate 

joined to counter septum to a complex spiderweb structure made up of axial plate, septal lamellae, and axial 

tabellae or upturned edges of tabulae. Tabulae ordinarily complete, conical, without well-defined shoulders, but 

may be nearly horizontal in some species. Peripheral 

tabellae rare. Dissepimentarium lonsdaleoid, commonly 

showing traces or crests of discontinuous septa in 

transverse section. Increase peripheral.

Nominal species in Western Interior Province. Cerioid: 

A. girtyi (Hayasaka, 1936), A. pennsylvanicus (Shimer, 

1926), A. ? shimeri (Crickmay, 1955). Fasciculate: A. ? 

arizelum (Crickmay, 1955), A. subcaespitosus (Meek, 

1873).

Discussion. This genus includes some of the corals 

previously referred to Lithostrotionella Yabe and 

Hayasaka, 1915. The taxonomy and world distribution 

of Acrocyathus have been discussed by Sando (1983). 

Acrocyathus was probably derived from Stelechophyllum 

in the Late Mississippian. It is the probable ancestor of 

Petalaxis and Sciophyllum.

Distribution. The principal occurrence of Acrocyathus 

in North America is in the St. Louis Limestone and its 

equivalents in the Southeastern Province. In the 

Western Interior Province of the United States, cerioid 

forms are rare in the Little Flat and Aspen Range Formations and Monroe Canyon Limestone of Idaho, the 

Humbug Formation of Utah, the Banner Formation and 

Monte Cristo Limestone of Nevada, and possibly in the 

Perdido Formation of California. The cerioid species A. 

girtyi occurs in the Monroe Canyon Limestone of Utah. 

The fasciculate species A. subcaespitosus is known only 

from the Monroe Canyon Limestone in Idaho. The 

fasciculate form is also represented by undetermined 

species in the Monroe Canyon Limestone of Utah and 

in the Red wall Limestone of Arizona. The occurrences 

of the genus in the western United States are in Zones 

IIIC and D.

In Canada, A. pennsylvanicus and A. ? shimeri occur 

in the Mount Head Formation in Alberta and 

southeastern British Columbia and in the Prophet Formation of northeastern British Columbia. A. ? arizelum 

occurs in the Mount Head Formation of Alberta. The

Canadian occurrences are in Zones IIIB-D and IV. The 

genus occurs exclusively in the shallow-water shelfcarbonate facies.

Genus ACTINOCYATHUS d'Orbigny, 1849 

Plate 6, figures 9, 10

Diagnosis. Cerioid colonial rugose corals. Columella 

complex, composed of a median plate, septal lamellae, 

and conically arranged, shallowly curved tabellae. Major septa may or may not extend to columella. Minor septa short. Tabularium composed of flat or sagging 

tabulae. Dissepimentarium lonsdaleoid.

Nominal species in Western Interior Province. A. 

stelcki (Nelson, 1960).

Discussion. All specimens belonging to this subgenus 

in the Western Interior Province are referred to the 

nominal species. See Sando (1975b, p. C21-C23) for a 

detailed analysis of the species.

Distribution. Actinocyathus is a relatively rare coral 

in both Canada and the United States. It is restricted 

to Zone VB in the United States and occurs in the upper part of Zone V in Canada. The subgenus is known 

from the Amsden Formation of Wyoming, the Surrett 

Canyon Formation of Idaho, the Green Ravine Formation of Utah, and the Etherington Formation of Alberta and British Columbia. This taxon occurs exclusively 

in the shallow-water shelf-carbonate lithofacies.

Genus AMPLEXIZAPHRENTIS Vaughan, 1906

Plate 1, figures 7, 8

Diagnosis. Small, solitary, ceratoid or trochoid rugose 

corals with a conspicuous cardinal fossula bounded 

laterally by cardinal lateral septa and axially by a wall 

consisting of fused axial ends of rhopaloid major septa 

of counter quadrants. Cardinal fossula ordinarily on concave side of corallum but may vary greatly in position 

in some species, generally expanded axially but may be 

parallel sided or axially constricted. Alar pseudofossulae 

commonly well developed. Septal plan ordinarily pinnate, although septal withdrawal in later growth stages 

may superimpose a radial plan in some species. Minor 

septa absent or poorly developed. Tabulae arched at axis 

of corallum and depressed in cardinal fossula. 

Dissepiments absent.

Nominal species in Western Interior Province. A. 

durhami (Langenheim and Tischler), 1960; A. elliptica 

(White, 1862); A. enniskilleni (Milne-Edwards and 

Haime, 1851); A.? eumetria (Crickmay, 1955); A. excavata (Girty, 1899); A. persimilis (Easton and 

Gutschick, 1953); A. spinulosa (Milne-Edwards and
ZONAL TAXA 17

Haime, 1851); A. taylori Sutherland, 1958; A. tumida 

(Easton, 1962); A. indifferens Sutherland, 1958.

A. elliptica includes the junior synonym Zaphrentis 

carinatus Worthen, 1890. A. spinulosa includes junior 

synonyms Zaphrentis spinulifera Hall in Hall and 

Whitney, 1858; Z. spergenensis Worthen, 1890; Z. 

chesterensis Worthen, 1890; andZ. pellaensis Worthen, 

1890.

Discussion. This genus, as presently recognized, includes corals previously assigned to Zaphrentites Hudson, 1941, and Menophyllum Milne-Edwards and Haime, 

1850 (e.g., Sando, 1960b, 1969). The nominal species are 

in need of restudy, and large collections assigned to this 

taxon have never been studied at the species level. 

Detailed work on these collections would probably result 

in recognition of stratigraphically useful species.

Distribution. Amplexizaphrentis is one of the commonest corals throughout the Mississippian of the 

Western Interior Province. It is represented in almost 

every coralliferous formation studied. Its stratigraphic 

range is from the base of Zone IB to the top of Zone VI. 

The genus occurs in a wide variety of lithofacies 

representing shallow- to deep-water habitats; it is most 

common in shallow-water environments.

Genus AMPLEXOCARINIA Soshkina, 1928 

Plate 1, figures 1, 2

Diagnosis. Small, solitary, ceratoid to cylindrical 

rugose corals in which a wide aulos is formed by 

downwardly bent edges of axial tabulae, dividing the 

lumen into axial and periaxial regions. Major septa 

short, abutting against aulos. Minor septa absent or 

poorly developed. Tabulae in aulos horizontal. Periaxial tabulae sloping downward toward periphery. 

Dissepiments ordinarily absent but may be present in 

some species.

Nominal species in Western Interior Province. None.

Discussion. Specimens from Montana were described 

by Sando (1960b) under the name of Amplexocarinia aff. 

A. heimoi Heritsch, a Permian species.

Distribution. Amplexocarinia is a rare coral with a 

short stratigraphic range in the Western Interior Province. It is known only from the Lodgepole Limestone of 

Montana and Utah, where it is in Zones IB and 1C and 

may also be present in IIA. Although it is found in both 

deep-water and shallow-water lithofacies, the known occurrences are mainly in deep-water habitats. The apparent rarity of this taxon may be due partly to its 

diminutive size, which may cause it to go unnoticed except in silicified collections.

Genus AMPLEXUS Sowerby, 1814 

Plate 1, figures 3, 4

Diagnosis. Small to large, solitary, cylindrical to vermiform rugose corals. Major septa thin, short, amplexoid. Minor septa absent or poorly developed in mature 

stages. Tabulae widely spaced, flat, complete, sometimes 

with downturned edges. Dissepiments absent.

Nominal species in Western Interior Province. None.

Discussion. This genus is easily confused with 

Guerichophyllum, whose principal stratigraphic occurrence is about the same. Guerichophyllum differs from 

Amplexus by having a zaphrentoid early stage and by 

having dissepiments. Although nominal species are 

unknown, study of large collections should prove rewarding in establishing stratigraphically useful species.

Distribution. Amplexus is common in Zones IB and 

1C but is known in Zone IA and in Zone IIB. It is very 

rare above Zone IIB, but sporadic single occurrences are 

known in Zones IIIB, VA and B, and VI. It is found in 

both shallow-water and deep-water lithofacies, but its 

principal occurrence is in deep-water habitats.

The genus is known mainly from the Lodgepole 

Limestone of Montana, Wyoming, Utah, and Idaho; the 

Allan Mountain Limestone of Montana; and the Deseret 

Limestone of Utah. It is rare in the Amsden Formation 

of Wyoming; the Great Blue Limestone, Surrett Canyon 

Formation, and Scott Peak Formation of Idaho; the 

Brazer Dolomite, Gardison Limestone, Joana Limestone, 

and Fitchville Formation of Utah; the Mission Canyon 

Limestone of Montana; the Monte Cristo Limestone, 

Eleana Formation, Joana Limestone, and Bristol Pass 

Limestone of Nevada; and the Tin Mountain Limestone 

of California. Amplexus has not been found in the 

Western Interior Province of Canada.

Genus ANKHELASMA Sando, 1961 

Plate 1, figure 9

Diagnosis. Small, solitary, ceratoid or trochoid rugose 

corals having a well-developed cardinal fossula on the 

convex side. Fossula bounded laterally by fused axial 

segments of cardinal lateral septa and axially by fused 

ends of counter and counter lateral septa. Septal plan 

in ephebic stage characterized by retreat of most major 

septa, leaving only five plates extending from corallum 

axis to calicular wall in ephebic calice: swollen counter 

septum, two plates formed by fusion of axial segments 

of cardinal lateral septa, and two plates formed by fusion of counter lateral septa. Minor septa present. 

Tabulae absent, open spaces in earlier part of corallum
18 CORAL ZONATION, MISSISSIPPIAN SYSTEM, WESTERN INTERIOR PROVINCE

filled with stereoplasm to floor of calice. Dissepiments 

absent.

Nominal species in Western Interior Province. A. 

typicum Sando, 1961.

Discussion. Ankhelasma is distinguished from 

Sychnoelasma, from which it was probably derived, by 

the presence of five septal plates in the ephebic stage.

Distribution. Although somewhat rare, Ankhelasma 

has a relatively restricted stratigraphic range, being 

known only from Zone IIIA. The nominal species occurs 

in the Brazer Dolomite of Utah and the Castle Reef 

Dolomite of Montana. Similar forms have been found 

in the Mission Canyon Limestone of Wyoming, Idaho, 

and Montana; the Monte Cristo Limestone of Nevada; 

and in the Turner Valley and Debolt Formations of 

northeastern British Columbia. The genus occurs exclusively in the shallow-water shelf-carbonate lithofacies 

near the shelf margin.

Genus AULOSTYLUS Sando, 1976 

Plate 1, figures 12, 13

Diagnosis. Cerioid colonial rugose corals that have 

an aulos formed by union of deflected axial ends of major septa. Columella ordinarily present, a vertically 

discontinuous axial plate formed by prolongation of one 

or two major septa. Major and minor septa noncarinate 

or weakly carinate (zigzag type). Tabulae in aulos ordinarily horizontal, flat, commonly turned up at intersection with columella; tabulae in tabularium ordinarily concave upward, horizontal to slightly inclined 

toward periphery of corallite. Dissepimentarium 

lonsdaleoid.

Nominal species in Western Interior Province. A. 

tubiferus (Hayasaka, 1936) and A. bamberi Sando, 1976.

Discussion. Aulostylus is distinguished from 

Stelechophyllum, from which it was probably derived, 

by the presence of an aulos.

Distribution. Aulostylus is a relatively rare coral that 

has a narrow stratigraphic range. It is confined to the 

lower half of Zone IIB. Aulostylus tubiferus occurs in the 

Lodgepole and Mission Canyon Limestones of Montana 

and in the Pekisko Formation of British Columbia. A. 

bamberi occurs in the Pekisko Formation of British Columbia. Undetermined species are also known from the 

Lodgepole Limestone and the Gardison Limestone of 

Utah. The genus occurs exclusively in shallow-water 

lithofacies.

Genus BARYTICHISMA Moore and Jeffords, 1945 

Plate 2, figures 4, 5

Diagnosis. Slightly curved, conico-cylindrical solitary 

rugose corals that have a thick theca. Major septa pinnately arranged and axially confluent in early stages 

but become radially arranged and amplexoid in mature 

stages. Tabulae generally flat but with downturned 

margins. Dissepiments absent.

Nominal species in Western Interior Province. B. 

amsdenense (Branson and Greger, 1918).

Discussion. Barytichisma is a descendent ofAmplexizaphrentis. It is distinguished from Amplexizaphrentis 

by its thick theca and amplexoid septa in mature stages.

Distribution. Barytichisma is a rare coral in the 

Mississippian of the Western Interior Province. Four of 

the five species known in North America are from beds 

of Pennsylvanian age outside the Western Interior Province (see discussion by Sando, 1975b). The nominal 

Mississippian species B. amsdenense occurs in the 

Amsden Formation of Wyoming. Undetermined species 

are known from the Surrett Canyon Formation of Idaho 

and the Chainman Shale of Utah. The occurrences are 

in the upper part of Zone VB and the lower part of Zone 

VI. The genus occurs exclusively in shallow-water 

lithofacies. It has not been found in Canada.

Genus BEAUMONTIA Milne-Edwards and Haime, 1851 

Plate 2, figure 3

Diagnosis. Cerioid tabulate corals in which some of 

the corallites become fasciculate and have rounded cross 

sections. Tabulae convex, incomplete. Mural pores present. Septal spines may be present.

Nominal species in Western Interior Province. None.

Discussion. Beaumontia is distinguished from 

Michelinia by its partly phaceloid growth form. It probably was derived from Michelinia.

Distribution. The genus is very rare in the Western 

Interior Province, where it is restricted to Zones IB and 

C, and DA. Undetermined species occur in the Lodgepole 

Limestone of Utah and the Tin Mountain Limestone of 

California. The genus has not been found in Canada. Its 

ecologic occurrence is exclusively in the deep-water 

basinal-carbonate facies.

Genus CANADIPHYLLUM Sutherland, 1954 

Plate 2, figures 1, 2

Diagnosis. Small to large, solitary trochoid rugose 

corals in which the major septa of the cardinal quadrants 

are pinnate and generally almost at right angles to a 

prominent cardinal fossula. Septa of the counter 

quadrants radially arranged to subparallel with the 

cardinal-counter plane. Minor septa short or absent. 

Tabulae complete, slightly convex, with downturned 

edges. Dissepiments absent.
ZONAL TAXA 19

Nominal species in Western Interior Province. C. knoxi 

Sutherland, 1954, and C. californicum (Tischler, 1956).

Discussion. In addition to the nominal species, at 

least two undescribed species are useful for stratigraphic 

zonation. A large form having a widely flaring calice is 

very common in Zones IIIC and D.

Canadiphyllum seems to have been derived from 

Amplexizaphrentis. The flaring form has a primitive columella at some localities, suggesting that the genus may 

be the ancestor of Ekvasophyllum.

Distribution. The total stratigraphic range of 

Canadiphyllum is from the upper part of Zone IIB (single 

occurrence) well into Zone IV, but it is abundant only 

in Zones IIIC and D. C. knoxi occurs in the Prophet Formation of British Columbia, and C. californicus occurs 

in the Perdido Formation of California. Undetermined 

species are known from the Mission Canyon Limestone 

of Wyoming; the Little Flat Formation, Aspen Range 

Formation, Great Blue Limestone, and Scott Peak Formation of Idaho; the Humbug Formation, Little Flat Formation, and Deseret Limestone of Utah; the Perdido Formation equivalent, Banner Formation, Monte Cristo 

Limestone, and Goughs Canyon Formation of Nevada; 

the Perdido Formation of California; the Turner Valley, 

Livingstone, and Mount Head Formations of Alberta; 

and the Debolt and Prophet Formations of northeastern 

British Columbia. The genus occurs in both shallow- and 

deep-water lithofacies but is most common in shallowwater habitats.

Genus CANINIA Michelin in Gervals, 1840 

Plate 2, figures 8, 9

Diagnosis. Trochoid to cylindrical solitary rugose corals. Major septa numerous, radially arranged, and 

amplexoid in later growth stages. Minor septa ordinarily 

short. Cardinal fossula well marked and on convex side 

of corallum, bounded axially by downturned edges of 

tabulae. Tabulae numerous, flat axially, and turned 

down at margins. Dissepimentarium variable in width, 

composed of regular or herringbone dissepiments.

Nominal species in Western Interior Province. None.

Discussion. Western Interior specimens assigned 

here are similar to corals identified as Caninia s.l. in 

most other parts of the world. This genus is the probable 

ancestor of Siphonophyllia.

Distribution. Caninia has a sporadic occurrence 

beginning at the top of Zone 1C and ranging to the top 

of Zone HID. The genus is rare throughout its range. It 

occurs in the Lodgepole Limestone, Allan Mountain 

Limestone, Mission Canyon Limestone, and Charles Formation of Montana; Madison Limestone of Wyoming; 

Scott Peak Formation of Idaho; Brazer Dolomite of Utah;

Monte Cristo Limestone of Nevada; Redwall Limestone 

of Arizona; Tin Mountain Limestone of California; Flett 

Formation of southwestern District of Mackenzie; and 

Debolt Formation of northeastern British Columbia. It 

occurs almost exclusively in shallow-water lithofacies.

Genus CLEISTOPORA Nicholson, 1888 

Plate 3, figure 5

Diagnosis. Small, discoid, adherent tabulate corals 

having basal epitheca. Corallites short, prismatic, having dense walls pierced by numerous mural pores and 

lined with reticulate skeletal tissue. Tabulae rare, thin.

Nominal species in Western Interior Province. C. 

placenta (White, 1880).

Discussion. All specimens of this genus found in the 

Western Interior Province are referred to C. placenta, 

which was originally described from the Chouteau 

Limestone of Missouri. C. placenta is distinguished from 

other North American species by having a larger corallum and larger number of corallites (see discussion by 

Sando, 1960b, p. 182). The Mississippian species probably were derived from species that occurred in the Late 

Devonian.

Distribution. C. placenta is a relatively rare coral that 

has a short stratigraphic range from the base of Zone 

1C into the lower part of Zone IIB. It occurs in the 

Lodgepole Limestone of Wyoming, Montana, and Utah; 

the Joana Limestone of Nevada; and the Banff Formation of Alberta. It is found in both deep-water and 

shallow-water lithofacies but is slightly more common 

in shallow-water habitats.

Genus CLISIOPHYLLUM Dana, 1846

Plate 3, figures 1, 2

Diagnosis. Solitary, ceratoid or trochoid to cylindrical 

rugose corals. Axial structure wide, composed of short 

medial plate and septal lamallae about half as numerous 

as major septa and commonly twisted spirally. Major 

septa radially arranged and not curved about small 

fossula. Minor septa short. Tabular floors conical. Two 

series of tabellae; taballae of periaxial series fewer and 

less steep than those of axial series. Dissepimentarium 

narrow, composed of numerous small concentric 

dissepiments.

Nominal species in Western Interior Province. None.

Discussion. This taxon includes corals previously 

referred to Dibunophyllum (see Sando, Mamet, and 

Dutro, 1969), from which it is distinguished by the 

nature of its axial structure.

Distribution. Clisiophyllum is a rare coral restricted 

to Zone IV in the Western Interior Province. It occurs
20 CORAL ZONATION, MISSISSIPPIAN SYSTEM, WESTERN INTERIOR PROVINCE

in the Otter Formation of Montana; the Monroe Canyon 

Limestone, Aspen Range Formation, Great Blue 

Limestone, and Scott Peak Formation of Idaho; the 

Monroe Canyon Limestone, Humbug Formation, and 

Great Blue Limestone of Utah; and the Mount Head Formation of Alberta. It is found exclusively in the shallowwater shelf-carbonate lithofacies.

Genus CYATHAXONIA Michelin, 1847 

Plate 3, figures 3, 4

Diagnosis. Small, solitary ceratoid to cylindrical 

rugose corals. Minor septa long, contratingent, inserted 

alternately with metasepta according to metriophyllid 

septal plan. Columella tall, developed independently of 

major septa but in contact with them. Tabulae inclined 

downwards toward epitheca. Dissepiments absent. Sides 

of septa may have rows of tubercles steeply inclined 

downward from the horizontal toward the corallum axis.

Nominal species in Western Interior Province. C. tantilla (Miller, 1891). C. tantilla includes junior synonyms 

C. minor Weller, 1909, and C. cordillerensis Easton, 

1958.

Discussion. Sando (1977d) amended the genus to include tuberculated forms referred to Cyathocarinia 

Soshkina, 1928, and discussed the distinguishing 

characters of the North American species. Although only 

one nominal species is known, other undescribed species 

probably are in collections from the Western Interior 

Province.

Distribution. Cyathaxonia is abundant in Zones IB 

and C and rare in Zones IIA and B, IIIA-C; it may also 

be present in Zones HID and IV. The nominal species 

is in the Lodgepole Limestone of Montana and Utah. 

Undetermined species occur in the Lodgepole Limestone 

of Wyoming; the Lodgepole Limestone, Mission Canyon 

Limestone, Allan Mountain Limestone, and Castle Reef 

Dolomite of Montana; the Mission Canyon Limestone 

of North Dakota; the Lodgepole Limestone, Mission Canyon Limestone, Middle Canyon Formation, and Scott 

Peak Formation of Idaho; the Lodgepole Limestone, 

Brazer Dolomite, Deseret Limestone, Joana Limestone, 

and Gardison Limestone of Utah; the Tin Mountain 

Limestone and Perdido Formation of California; the Redwall Limestone of Arizona; the Monte Cristo Limestone, 

Joana Limestone, Diamond Peak Formation, Eleana 

Formation, and Goughs Canyon Formation of Nevada; 

the Banff Formation of Alberta and northeastern British 

Columbia; and the Turner Valley Formation of northeastern British Columbia. The genus is found 

predominantly in deep-water lithofacies.

Genus DIPHYPHYLLUM Lonsdale, 1845

Plate 3, figures 6, 7

Diagnosis. Fasciculate colonial rugose corals. Septa 

short. Columella absent or impersistent. Tabulae flat or 

convex, with downturned edges. Dissepimentarium 

regular, composed of one or more rows of small 

dissepiments.

Nominal species in Western Interior Province. None.

Discussion. This genus is very similar to Siphonodendron, from which it differs by having flat or convex 

tabulae and by lacking a columella or having a thin, vertically discontinuous one. The genomorph concept of 

Smith and Lang (1930) was proposed to explain the relationship between Diphyphyllum and phaceloid Lithostrotion; this concept and its attendant nomenclatural notation is no longer accepted (see Wilson, 1963). Although 

some workers have questioned the biologic validity of 

Diphyphyllum as a distinct genus, the concept is useful 

at least in recognizing distinct morphotypes. The name 

is used in this report in quotes to denote the uncertainty of its usage.

Distribution. In the United States, Diphyphyllum is 

a common coral in Zone IIIA and is rare in Zone HID. 

In Canada, it is common in the upper half of Zone IIB 

and the lower half of Zone IIIA, rare in the upper half 

of Zone IIIA and in Zone IIIB, and common again in 

Zones IIIC and D, and in the lower part of Zone IV. It 

occurs in the Madison Limestone and Mission Canyon 

Limestone of Wyoming; the Charles Formation and Mission Canyon Limestone of Montana; the Mission Canyon 

Limestone and Little Flat Formation of Idaho; the Monte 

Cristo Limestone of Nevada; the Deseret Limestone of 

Utah; the Charles Formation of Saskatchewan(?); the 

Shunda and Mount Head Formations of Alberta; the 

Turner Valley and Prophet Formations of northeastern 

British Columbia; and the Flett Formation of 

southwestern District of Mackenzie. It is found exclusively in the shallow-water carbonate lithofacies.

Genus DORLODOTIA Salee, 1920 

Plate 3, figures 8, 9

Diagnosis. Fasciculate colonial rugose corals. Major 

septa short, ordinarily confined to tabularium. Minor 

septa ordinarily absent or consist of ridges on theca. Columella composed of a single axial plate that may be 

joined to one or more septa. Tabulae tent-shaped. 

Dissepimentarium composed of lonsdaleoid 

dissepiments.
ZONAL TAXA 21

Nominal species in Western Interior Province. D. ? inconstans (Easton and Gutschick, 1953).

Discussion. Corals from the Western Interior Province are referred to Dorlodotia with a query because they 

have a weakly lonsdaleoid dissepimentarium and an impersistent columella.

Distribution. The nominal species occurs in Zones 

IIIA and B in the Redwall Limestone of Arizona. 

Undescribed species are rare from the top of Zone IIB 

to the top of Zone HID. The genus is represented in the 

Little Flat Formation of Idaho; the Banner Formation 

and Monte Cristo Limestone of Nevada; the Rundle 

Group of British Columbia; and the Turner Valley Formation of Alberta. It occurs exclusively in the shallowwater shelf-carbonate lithofacies.

Genus DUNCANOPORA Sando, 1975 

Plate 4, figures 5, 6

Diagnosis. Phaceloid and cerioid tabulate corals composed of cylindrical corallites joined at regular intervals 

in the corallum by lateral expansion or by connecting 

structures. Regular connections within the corallum 

may be by simple increase in diameter of corallites, 

which crowds the corallites together to produce 

polygonal cross sections, or by connecting tubules or encircling deposits of sclerenchyme perforated by pores 

connecting adjacent corallites. Tabulae sparse, ordinarily complete, horizontal, and slightly concave upward. 

Septal spines rare.

Nominal species in Western Interior Province. D. duncanae Sando, 1975.

Discussion. Corals belonging to this taxon were 

previously referred to Tetraporinus? (Dutro and Sando, 

1963) and Hayasakaia? (Sando, Mamet, and Dutro, 

1969). Duncanopora was probably derived from 

Multithecopora?

Distribution. All specimens of this genus known from 

the Western Interior Province are referred to D. duncanae, which is confined to Zone VB. The species occurs 

in the Amsden Formation of Wyoming; the Monroe Canyon Limestone, Aspen Range Formation, and Surrett 

Canyon Formation of Idaho; and the Green Ravine Formation of Utah. It has not been found in Canada. Duncanopora occurs exclusively in the shallow-water shelfcarbonate lithofacies.

Genus EKVASOPHYLLUM Parks, 1951

Plate 4, figures 3, 4

Diagnosis. Medium, sized, trochoid, solitary rugose 

corals having numerous septa tending toward radial

symmetry, except near prominent cardinal fossula on 

convex side of corallum, where they are pinnately arranged. Columella solid, slightly laterally compressed, 

rodlike. Tabulae incomplete, arched upwards to join columella. Dissepimentarium composed of small regular 

dissepiments.

Nominal species in Western Interior Province. E. banffense (Warren, 1927); E. cascadense (Warren, 1927); E. 

enclinotabulatum Sutherland, 1958; E.? harkeri 

Sutherland, 1958; E. inclinatum Parks, 1951. E. 

cascadense includes the junior synonym E. turbineum 

Parks, 1951.

Discussion. Ekvasophyllum is similar to 

Lophophyllum? but differs mainly in having a welldeveloped dissepimentarium and a thicker solid columella. Ekvasophyllum may have been derived from 

Canadiphyllum and is the ancestor of Faberophyllum.

Distribution. Ekvasophyllum is a common coral in 

Zone HID. In the United States, it is rare in Zone IIIC 

and in the lower half of Zone IV. In Canada, the genus 

ranges from the lower half of Zone IIIA into the upper 

half of Zone IV. It occurs in the Little Flat Formation, 

Monroe Canyon Limestone, Middle Canyon Formation, 

Aspen Range Formation, and Scott Peak Formation of 

Idaho; the Humbug Formation and Deseret Limestone 

of Utah; the Redwall Limestone of Arizona; the Banner 

Formation(?) and Chainman Formation(?) of Nevada; the 

Perdido Formation of California(?); the Mount Head and 

Livingstone Formations of Alberta; the Mount Head Formation and Milford Group of southeastern British Columbia; the Golata(?), Prophet, and Debolt Formations 

of northeastern British Columbia; and the Flett Formation of the southwestern District of Mackenzie. 

Ekvasophyllum occurs almost exclusively in the shallowwater shelf-carbonate facies.

Genus FABEROPHYLLUM Parks, 1951

Plate 4, figures 1, 2

Diagnosis. Large trochoid to subcylindrical solitary 

rugose corals having numerous major septa tending 

toward radial symmetry, except near prominent closed 

cardinal fossula on convex side of corallum, where they 

are pinnately arranged. Minor septa long. Among 

species, axial structure ranges from a complex of one or 

more lamellae and tabellae to no axial structure and sagging tabulae. Tabulae incomplete, nearly horizontal, except for upturned edges at dissepimentarium. 

Dissepimentarium wide, composed of regular 

dissepiments.

Nominal species in Western Interior Province. F. 

araneosum Parks, 1951; F. languidum Parks, 1951; F.
22 CORAL ZONATION, MISSISSIPPIAN SYSTEM, WESTERN INTERIOR PROVINCE

leathamense Parks, 1951; F. minnewankensis (Shimer, 

1926); F. occultum Parks, 1951; F. pisgahense Parks, 

1951; F. stansburyi (Hall, 1852). F. stansburyi includes 

the junior synonym Zaphrentis? multilamellata Hall, 

1852.

Discussion. In 1956, Hill (p. F290) regarded 

Faberophyllum as a junior synonym of Turbophyllum 

Parks, 1951, but later, she (1981, p. F374) considered 

the two genera as distinct. Although the two genera are 

very similar, we regard them as distinct. However, we 

have not referred any specimens other than the types 

to Turbophyllum and have not used this genus in the 

zonation because of its apparent rarity.

The nominal species of Faberophyllum are in need of 

restudy because they are based on fragmentary coralla 

that do not include the early growth stages. Study of 

abundant specimens in our collections might provide a 

satisfactory basis for revising nominal species.

Complete specimens of Faberophyllum have an early 

stage that has a solid styliform columella like that of 

Ekvasophyllum(see Sando, 1965, p. E21). This observation supports Parks' (1951) conclusion that 

Ekvasophyllum is the ancestor of Faberophyllum.

Distribution. Faberophyllum is a very common coral 

in Zone IV; it is rare in the upper half of Zone HID. The 

genus occurs in the Little Flat Formation, Monroe Canyon Limestone, Great Blue Limestone, and Scott Peak 

Formation of Idaho; the Monroe Canyon Limestone, 

Aspen Range Formation, Humbug Formation, and 

Great Blue Limestone of Utah; the Diamond Peak and 

Eleana Formations of Nevada; the Mount Head Formation of Alberta; and the Milford Group of southeastern 

British Columbia. It occurs almost exclusively in the 

shallow-water shelf-carbonate facies.

Genus GUERICHOPHYLLUM Rozkowska, 1969

Plate 2, figures 6, 7

Diagnosis. Solitary trochoid to vermiform rugose corals. Major septa united at axis in early growth stages, 

withdrawn from axis in later stages. Cardinal fossula 

variable in position with respect to curvature of corallum, variably developed, bounded axially by 

downturned edges of tabulae. Minor septa short or absent. Tabulae ordinarily complete, flat axially, and 

turned down at margins. Dissepimentarium narrow, 

regular to lonsdaleoid.

Nominal species in Western Interior Province. None.

Discussion. Corals here referred to Guerichophyllum 

were previously identified as Caninia (Caninia), which 

does not have lonsdaleoid dissepiments. The occurrence 

of Guerichophyllum is mainly Devonian (Hill, 1981, p. 

F196), but Bartzsh and Weyer (1981) illustrated

specimens from the Gattendorfia Zone of Saalfeld in 

Thuringia. The Western Interior specimens are similar 

to those reported to Bartzsh and Weyer.

Guerichophyllum is similar to Amplexus but differs 

from the latter by having a zaphrentoid early stage and 

by having dissepiments.

Distribution. Guerichophyllum is a common coral in 

Zones IB and C, IIA, and the lower part of IIB; it is rare 

in the upper part of Zone IIB. It is most common in the 

Lodgepole Limestone of Montana, Idaho, and Utah, and 

the Allan Mountain Limestone of Montana; it is rare 

in the Gardison and Deseret Limestones of Utah and in 

the Tin Mountain Limestone of California. It has not 

been found in Canada. Although it occurs in shallowwater lithofacies, it is most common to deep-water 

habitats.

Genus HAPSIPHYLLUM Simpson, 1900 

Plate 5, figure 3

Diagnosis. Small, curved, conical, solitary rugose corals having a prominent key-hole-shaped cardinal fossula 

ordinarily on the concave side of the corallum. Major septa pinnate. Minor septa contratingent. Tabulae present. 

Dissepiments absent.

Nominal species in Western Interior Province. None.

Discussion. Some authors have used this taxon in a 

broad sense to include simple Carboniferous corals 

formerly referred to Zaphrentis. Actually, the genus is 

distinguished by its contratingent minor septa, a rare 

feature in Carboniferous corals, and it is here used in 

a restricted sense for corals similar to Amplexizaphrentis but having contratingent minor septa. Hapsiphyllum 

was probably derived from Amplexizaphrentis.

Distribution. Hapsiphyllum is a common coral in the 

Southeastern Province, where it occurs in the Warsaw 

and Salem Limestones. It is rare in the Western Interior 

Province, where it occurs mainly in Zone HID; a single 

occurrence is known in Zone IIA. Undetermined species 

occur in the Allan Mountain Limestone of Montana, the 

Deseret Limestone of Utah, the Aspen Range Formation 

of Idaho, and an unknown formation in Nevada. The 

genus has not been found in Canada. It occurs exclusively in the shallow-water shelf-carbonate lithofacies.

Genus KONINCKOPHYLLUM Thomson and Nicholson, 1876 

Plate 5, figures 1, 2

Diagnosis. Solitary trochoid to cylindrical or dendroid 

colonial rugose corals. Columella small, compressed, 

styliform, commonly vertically discontinuous. Major septa thin, radial, do not extend to columella. Cardinal
ZONAL TAXA

fossula poorly defined. Minor septa long. Tabulae tentshaped. Dissepimentarium wide, regular.

Nominal species in Western Interior Province. None.

Distribution. This genus is rare in the Western Interior Province. In Canada, its main occurrence is in the 

lower half of Zone IV, but sporadic occurrences are 

known in Zones IIB and IIIA. In the United States, forms 

referred questionably to the genus occur in Zone HID. 

The occurrences are in the Deseret Limestone of Utah, 

an unknown formation in Nevada, the Prophet and 

Debolt Formations of northeastern British Columbia, 

and the Mount Head Formation of Alberta. The genus 

occurs in both deep-water and shallow-water lithofacies.

Genus LIARDIPHYLLUM Sutherland, 1954

Plate 5, figure 4

Diagnosis. Small to medium-sized trochoid to cylindrical solitary rugose corals. Major septa amplexoid. 

Tabulae complete and incomplete, inclined downward 

axially to form a series of irregular inverted cones with 

cup-shaped depressions in axial region. Dissepimentarium lonsdaleoid.

Nominal species in Western Interior Province. L. hagei 

Sutherland, 1954.

Distribution. This very rare genus is known with certainty only from Zones IIIA and C in the Flett Formation of the southwestern District of Mackenzie. It has 

been identified questionably in Zone IIB in the Allan 

Mountain Limestone of Montana. It occurs in the 

shallow-water shelf-carbonate facies.

Genus LOPHOPHYLLUM Milne-Edwards and Haime, 1850

Plate 6, figure 4

Diagnosis. Trochoid solitary rugose corals having a 

prominent cardinal fossala on convex side of corallum. 

Columella simple, styliform. Major septa long, may or 

may not extend to columella, pinnate to radial. Minor 

septa short. Tabulae arched upward at axis. 

Dissepiments absent or sparse in mature stage only.

Nominal species in Western Interior Province. L. ? columnirotata (Langenheim and Tischler, 1960); L. eastoni 

(Langenheim and Tischler, 1960); L. panamintense 

(Langenheim and Tischler, 1960); L. ? proteus 

(Sutherland, 1958); and L. ? teres (Girty, 1899).

Discussion. This taxon includes corals formerly referred to Rylstonia (e.g., Sando, 1960b); they are here 

referred to Lophophyllum with a query because the type 

species of that genus is in need of further study before 

the genus can be identified with certainty. Some Soviet 

coral students (e.g., Sayutina, 1973) placed corals having similar morphology in Cyathoclisia, but the type 

species of that genus has a well-developed dissepimentarium and a complex axial structure, whereas the corals from the Western Interior Province lack 

dissepiments or have a poorly developed dissepimentarium and a simple axial structure.

Undescribed specimens assigned to Lophophyllum? 

have not been studied in detail, and how many species 

are present in the Western Interior Province is not clear. 

Some of the nominal species may be synonyms. Forms 

that occur in the middle and upper parts of the 

stratigraphic range of the genus are characterized by 

a subdued columella. These forms may have given rise 

to Zaphriphyllum by reduction of the columella and addition of dissepiments.

Distribution. In the United States, Lophophyllum? is 

rare in Zone IA, common from the base of Zone IB to 

about the middle of Zone IIB, and then rare again from 

about the middle of Zone IIB to the middle of Zone IIIA. 

It occurs in the Lodgepole Limestone, Mission Canyon 

Limestone, and Madison Limestone of Wyoming; the 

Allan Mountain Limestone, Mission Canyon Limestone, 

and Lodgepole Limestone of Montana; the Lodgepole and 

Mission Canyon Limestones of Idaho; the Lodgepole 

Limestone, Gardison Limestone, Joana Limestone, and 

Fitchville Formation of Utah; the Tin Mountain 

Limestone of California; the Redwall Limestone of 

Arizona; the Pahasapa Limestone of South Dakota; and 

the Joana Limestone, Mercury Limestone, Goughs Canyon Formation, and Monte Cristo Limestone of Nevada.

In Canada, the genus is common from the base to the 

top of Zone IIB and rare from the top of Zone IIB to the 

lower half of Zone HID. It occurs in the Flett Formation 

of the southwestern District of Mackenzie; the Banff, 

Pekisko, Shunda, Prophet, and Debolt Formations of 

northeastern British Columbia; the Banff, Pekisko, 

Shunda, Turner Valley, Livingstone, and Mount Head 

Formations of Alberta; and the Mission Canyon 

Limestone of Saskatchewan.

Lophophyllum? occurs in both deep- and shallow-water 

lithofacies but is more common in the shallow-water 

habitats.

Genus METRIOPHYLLUM Milne-Edwards and Haime, 1850 

Plate 6, figure 3

Diagnosis. Small solitary trochoid or ceratoid rugose 

corals. Major septa extend to axis, where they form a 

simple columella. Minor septa absent or short, ordinarily 

contratingent when present. Cardinal fossula may be 

present at maturity. Sides of septa have horizontal or 

nearly horizontal flanges that have upturned outer 

edges. Tabulae very thin, steeply inclined from axis. 

Dissepiments absent.
24 CORAL ZONATION, MISSISSIPPIAN SYSTEM, WESTERN INTERIOR PROVINCE

Nominal species in Western Interior Province. 

Specimens in the Western Interior Province may be conspecific with M. deminutivum Easton, 1944, which was 

described from the Chouteau Limestone of Missouri.

Discussion. Metriophyllum is one of the few Devonian 

genera that extend into the Mississippian of North 

America. Specimens are so small that they are seldom 

collected except in etched silicified collections, so that 

the genus may be more common than is indicated by the 

number of existing specimens and localities.

Distribution. This rare genus is restricted to Zone IB 

in the United States. It has not been found in Canada. 

It occurs in the Allan Mountain Limestone of Montana 

and the Lodgepole Limestone of Montana, Idaho, and 

Utah. The genus is found exclusively in the deep-water 

basinal-carbonate facies.

Genus MICHELINIA de Koninck, 1841

Plate 7, figures 6, 7

Diagnosis. Cerioid tabulate corals of variable corallum shape. Walls pierced by mural pores. Septal spines 

or ridges present. Tabulae numerous, incomplete, 

convex.

Nominal species in Western Interior Province. M. expansa White, 1880.

Discussion. Michelinia is one of the few Devonian 

genera that range into the Mississippian of western 

North America. Existing collections have not been 

studied in detail in order to determine how many species 

are present in the Western Interior Province. In addition to the nominal species, there is a species similar 

to M. meekana Girty, 1910, in Zone V.

Distribution. Michelinia has a long stratigraphic 

range beginning at the base of Zone IB and extending 

to the top of Zone VB, but the occurrences are sporadic. 

The nominal species is most common from the base of 

Zone II to about the middle of Zone II. Michelinia occurs 

in the Lodgepole and Mission Canyon Limestones of 

Wyoming; the Lodgepole Limestone, Mission Canyon 

Limestone, Allan Mountain Limestone, Castle Reef 

Dolomite, and Big Snowy Formation of Montana; the 

Lodgepole Limestone, Surrett Canyon Formation, Scott 

Peak Formation, and South Creek Formation of Idaho; 

the Lodgepole Limestone, Gardison Limestone, Deseret 

Limestone, and Doughnut Formation of Utah; the Tin 

Mountain Limestone and Perdido Formation(?) of 

California; the Redwall Limestone of Arizona; the Diamond Peak Formation, Joana Limestone, Goughs Canyon Formation, and Monte Cristo Limestone of Nevada; 

the Mission Canyon Limestone of Saskatchewan; the 

Banff, Pekisko, and Shunda Formations of Alberta; the 

Etherington Formation of southeastern British Columbia; and the Banff, Pekisko, Shunda, and Debolt Formations of northeastern British Columbia. Michelinia 

occurs predominantly in the shallow-water shelfcarbonate lithofacies.

Genus MULTITHECOPORA Yoh, 1927 

Plate 7, figures 1, 2

Diagnosis. Fasciculate tabulate corals. Corallite 

walls variable in thickness, commonly composed of concentric layers of sclerenchyme that reduce visceral cavity to a third to a fifth of the corallite diameter. Connecting tubes very thin and sparse or absent. Tabulae thin, 

sparse, complete, concave upward. Septal spines absent 

or sparse.

Nominal species in Western Interior Province. M. 

amsdenensis Sando, 1975b.

Discussion. Specimens from the Western Interior 

Province have walls that are not as thick as those of the 

type species, and connecting tubes are moderately abundant in these corals. Probably more than one species is 

present in the Western Interior Province, but the 

specimens collected have not been thoroughly studied. 

This genus was probably derived from Syringopora and 

gave rise to Duncanopora. Multithecopora differs from 

Syringopora by having sparse, nearly flat tabulae and 

thicker walls.

Distribution. Multithecopora is rare in Zones HID, IV, 

and VA and B. Most occurrences are in Zone VB, where 

the genus is represented by the nominal species. The 

genus is known from the Amsden Formation of Wyoming; the Big Snowy Formation of Montana; the Monroe 

Canyon Limestone and the Surrett Canyon, Aspen 

Range, and Scott Peak Formations of Idaho; the Deseret 

Limestone, Chainman Shale, and the Green Ravine and 

Doughnut Formations of Utah; and the Banner Formation of Nevada. It has not been found in Canada. The 

genus occurs exclusively in the shallow-water shelfcarbonate lithofacies.

Genus NEAXON Kullmann, 1965

Plate 7, figure 8

Diagnosis. Small, trochoid solitary rugose corals. Major septa radially arranged and ordinarily rhopaloid, 

joined axially to form an aulos. Minor septa contratingent or free, longer in counter quadrants. Outer 

tabulae slope up from epitheca. Inner tabulae horizontal where present. Dissepiments absent.

Nominal species in Western Interior Province. None.

Discussion. Corals previously identified as Permia 

Stuckenberg, 1895 (as interpreted by Hudson, 1943),
ZONAL TAXA 25

from the Western Interior Province are here referred to 

Neaxon. According to J. Fedorowski (oral commun., 

1973) and N. V. Kabakovich (oral commun., 1975), who 

have examined Stuckenberg's types ofPermia, the types 

are early stages of Aulophyllum. Although the Western 

Interior corals seem to belong to Neaxon, the name is 

used with a query because of the lack of detailed study 

of these North American specimens.

Distribution. Neaxon? is a rare coral in the Western 

Interior Province, ranging sporadically from the base of 

Zone IB into the lower part of Zone IIIA. It is represented 

by undetermined species in the Lodgepole Limestone of 

Wyoming; the Allan Mountain and Lodgepole 

Limestones of Montana; the Lodgepole and Joana 

Limestones of Utah; the Goughs Canyon Formation of 

Nevada; and the Perdido Formation of California. It occurs almost exclusively in the deep-water basinalcarbonate facies.

Genus PALAEACIS Haime in Milne-Edwards, 1857 

Plate 7, figures 3-5

Diagnosis. Small wedge-shaped or conical tabulate 

corals, adherent in young stages, having few corallites. 

Thecae very thick, spongy, traversed by numerous 

anastomosing canals that open on the surface of the corallum (between the cups) and inside the cups via scattered or oriented pores. Septa absent or represented by 

longitudinal carinae. Tabulae absent.

Nominal species in Western Interior Province. P. 

cuneiformis Milne-Edwards, I860, P. elongata McGugan, 

1983. P. cuneiformis includes the junior synonym 

Sphenopoterium cuneatum Meek and Worthen, 1860.

Discussion. Although this genus does not have 

tabulae, it is referred to the Order Tabulata because its 

other morphological features are similar to those of other 

tabulates.

Distribution. Palaeacis is a rare coral in the Western 

Interior Province and has a sporadic range from the base 

of Zone IB into the lower part of Zone VB in the United 

States. An undescribed new species is restricted to Zone 

IB. The nominal species, P. cuneiformis, has been found 

in Zones IIIA and D. Undetermined species occur in 

Zones IIB and VB.

P. elongata, recently found in Zone IV or the lower part 

of Zone V, is the only representative known from 

Canada.

The genus occurs in the Lodgepole Limestone, Castle 

Reef Dolomite, and Big Snowy Formation of Montana; 

the Lodgepole Limestone, Little Flat Formation, and 

Scott Peak Formation of Idaho; the Lodgepole Limestone 

and Deseret Limestone of Utah; the "Trinity Limestone"

of Nevada; and at an undetermined level (talus 

specimen) in the upper part of the Mount Head Formation or lower part of the Etherington Formation in 

Alberta. It occurs in both deep-water and shallow-water 

carbonate lithofacies.

Genus PETALAXIS Milne-Edwards and Haime, 1852 

Plate 7, figures 9, 10

Diagnosis. Cerioid colonial rugose corals having 

tabular to hemispherical corallum shape. Major septa 

thin, ordinarily extending into tabularium but seldom 

reaching columella, and ordinarily discontinuous or absent in dissepimentarium, but species with only weakly lonsdaleoid dissepimentarium are known. Minor septa 

absent to well developed. Columella ordinarily a simple axial plate joined to counter and/or cardinal septum 

but composed of axial plates, impersistent vertical axial tabellae, and a few axial lamellae in some species. 

Tabulae complete and incomplete, essentially horizontal, flat, concave, or convex, turned up at columella in 

some species. Peripheral clinotabellae may be present. 

Dissepimentarium ordinarily lonsdaleoid but may be 

mostly regular in some species.

Nominal species in Western Interior Province. P. 

bailliei (Nelson, 1960); P. simplex (Hayasaka, 1936); P. 

tabulatus (Hayasaka, 1936); P. wyomingensis Sando(?).

Discussion. Sando (1983) recently revised the genus 

to include the first three nominal species listed above 

from the Mississippian of the Western Interior Province. 

P. wyomingensis Sando, another nominal species, occurs 

in beds of uncertain age and identity that may belong 

to the Mississippian. The genus ranges from Late 

Mississippian to Permian, but Pennsylvanian and Permian species do not occur in the Western Interior Province. Petalaxis was probably derived from Acrocyathus 

in the Late Mississippian.

Distribution. Petalaxis is a rare coral in the Mississippian of the Western Interior Province. Its principal occurrence is in Zones HID and IV, but it may also be present in Zone V or Zone VI, on the basis of a single occurrence of uncertain stratigraphic position. It occurs in the 

Scott Peak and Aspen Range Formations of Idaho; the 

Monroe Canyon Limestone equivalent of Utah; an 

unknown formation in Wyoming; and the Mount Head 

Formation of Alberta. It occurs exclusively in the 

shallow-water shelf-carbonate lithofacies.

Genus PLEUROSIPHONELLA Chudinova, 1970

Plate 8, figures 7, 8

Diagnosis. Fasciculate tabulate corals composed of 

relatively thick walled, cylindrical corallites joined at
26 CORAL ZONATION, MISSISSIPPIAN SYSTEM, WESTERN INTERIOR PROVINCE

irregular intervals and connected by mural pores or very 

short tubules. Corallites commonly joined to form short 

chains or clusters at some levels in the corallum. 

Tabulae mostly vesicular, ordinarily vertically disposed along one side of corallite wall in such a way as to 

leave a visceral canal along the other side, or tabulae 

may be infundibular, having the axis of the funnels 

closer to one side of the corallite than the other. Septal 

spines rare or absent in some species, present in others. 

Wall structure fibrous. Increase lateral, commonly 

multiple.

Nominal species in Western Interior Province. P. 

drummondi (Nelson, 1962); P. magnussoni (Nelson, 

1962); P. virginica (Butts, 1941).

Discussion. This genus is differentiated from Syringopora, from which it was probably derived, by its excentric tabulae, thick fibrous walls, and corallites seldom 

joined by connecting tubules. We regard Oharaia 

Nelson, 1977, as a junior synonym of Pleurosiphonella.

Distribution. An undetermined species of 

Pleurosiphonella occurs in Zones IIB and IIIA in the 

United States; P. virginica is common in Zones HID and 

IV, and P. drummondi is rare in Zones VB and VI. In 

Canada, P. virginica and P. magnussoni range from near 

the base of Zone IIIA into Zone V, and P. drummondi 

is found in the lower part of Zone VI.

The genus occurs in the Mission Canyon Limestone 

and Amsden Formation of Wyoming; the Castle Reef 

Dolomite, Charles Formation, and Mission Canyon 

Limestone of Montana; the Monroe Canyon Limestone, 

Great Blue Limestone, Aspen Range Formation, Scott 

Peak Formation, and Surrett Canyon Formation of 

Idaho; the Little Flat Formation, Monroe Canyon 

Limestone, Deseret Limestone, Humbug Formation, and 

Great Blue Limestone of Utah; an unnamed formation 

in Nevada; and the Mount Head, Livingstone, and 

Etherington Formations of Alberta and British 

Columbia.

Pleurosiphonella occurs exclusively in the shallowwater shelf-carbonate lithofacies.

Genus PSEUDODORLODOTIA Minato, 1955

Plate 8, figures 3, 4

Diagnosis. Phaceloid colonial rugose corals. Major 

septa do not reach axis. Minor septa short. Cardinal 

fossula may be present. Columella weak, impersistent, 

or absent, consisting of a simple axial plate commonly 

connected to counter septum. Tabulae flat, with 

downturned edges. Dissepimentarium lonsdaleoid.

Nominal species in Western Interior Province. None.

Discussion. Just as Diphyphyllum may be a noncolumellate variant of Siphonodendron, so

Pseudodorlodotia may be a noncolumellate variant of 

Dorlodotia. However, corals identified as 

"Pseudodorlodotia" from the Western Interior Province 

may actually represent a noncolumellate variant of 

phaceloid species of Acrocyathus; the ranges of these two 

taxa are almost identical. These corals also differ from 

the type species of Dorlodotia and Pseudodorlodotia by 

having an only weakly lonsdaleoid dissepimentarium. 

The name is used in quotes for Western Interior Province corals because of these taxonomic uncertainties.

Distribution. "Pseudodorlodotia" is a rare coral in the 

Western Interior Province. Its principal range in the 

United States is from about the middle of Zone HID to 

the top of Zone IV; a questionable occurrence is in the 

upper half of Zone VA. In Canada, the genus ranges 

nearly through the upper half of Zone IV.

"Pseudodorlodotia" occurs in the Monroe Canyon 

Limestone, Scott Peak Formation, Aspen Range Formation, and possibly the Surrett Canyon Formation of 

Idaho; the Deseret Limestone of Utah; the Mount Head 

Formation of Alberta; and the Milford Group of 

southeastern British Columbia. The genus occurs exclusively in the shallow-water shelf-carbonate 

lithofacies.

Genus RHOPALOLASMA Hudson, 1936 

Plate 8, figure 5

Diagnosis. Small, curved, trochoid solitary rugose 

corals having rhopaloid septa and tachylasmoid septal 

development. Major septa approximately straight, 

bilaterally symmetrical. Cardinal septum on convex side 

of corallum. Cardinal and counter septa short. Alar septa 

and KL1 strongly rhopaloid. KL4, KL5, CL3, CL4 may 

be strongly rhopaloid. KL2, KL3, CL2, CL3 not fully 

developed. Minor septa short. Tabulae conical, incomplete. Dissepiments absent.

Nominal species in Western Interior Province. None.

Discussion. This genus is commonly regarded as a 

junior synonym of Ufimia, a Late Carboniferous and Permian form. We regard Ufimia as a homeomorph of 

Rhopahlasma and retain the latter name for Early Carboniferous forms.

Distribution. Rhopahlasma is represented by questionable occurrences in Zone IIB and is rare from the 

base of Zone IIIA to the top of Zone HID. It occurs in 

the Deep Creek Formation of Idaho; the Deseret 

Limestone, Little Flat Formation, and Humbug Formation of Utah; and the Chainman Formation, Goughs 

Canyon Formation, and "Trinity Limestone" of Nevada. 

The genus has not been found in Canada. It occurs 

almost exclusively in the deep-water basinal-carbonate 

lithofacies.
ZONAL TAXA 27

Genus ROTIPHYLLUM Hudson, 1942 

Plate 8, figure 6

Diagnosis. Small ceratoid solitary rugose corals having a cardinal fossula on the convex side of the corallum. 

Major septa radially arranged, peripherally dilated, and 

rhopaloid axially, where they are joined by stereoplasm 

to form a columella. Counter septum may be longer than 

other septa. Minor septa long, contratingent. Tabulae 

incomplete, conical. Dissepiments absent.

Nominal species in Western Interior Province. None. 

Sando (1960b, p. 171) referred some corals from Montana questionably to R. hians Easton, a Mississippi 

Valley species.

Distribution. Rotiphyllum is a rare coral having a 

long stratigraphic range from the base of Zone 1C to the 

top of Zone VA in the United States. It occurs in the 

Madison Limestone of Wyoming; the Lodgepole, Mission 

Canyon, and Allan Mountain Limestones of Montana; 

the Aspen Range, Deep Creek, Middle Canyon, Scott 

Peak, South Creek, and McGowan Creek Formations of 

Idaho; the Little Flat Formation , Lodgepole Limestone, 

Deseret Limestone, Humbug Formation, Gardison 

Limestone, and Great Blue Limestone of Utah; and in 

an undetermined formation in Nevada. The genus is 

very rare in Zone IIIB in Canada. It occurs in both deepwater and shallow-water lithofacies but is more common 

in deep-water habitats.

Genus SCHOENOPHYLLUM Simpson, 1900 

Plate 8, figures 1, 2

Diagnosis. Phaceloid colonial rugose corals having 

slender connecting tubules. Major septa few, mostly not 

reaching axis. Minor septa short or absent. Columella 

solid, compressed, styliform, formed by prolongation of 

counter septum. Tabulae complete, almost flat. 

Dissepimentarium consists of a single row of impersistent dissepiments.

Nominal species in Western Interior Province. S. aggregatum Simpson, 1900. S. aggregatum includes the 

junior synonym Lithostrotion [SiphonodendronJ 

genevievensis Easton, 1957.

Discussion. Schoenophyllum is similar to 

Siphonodendron, from which it was probably derived, 

but it differs in having a strong counter septum prolonged to form the columella, connecting processes, and nearly flat tabulae. Easton (1957) concluded that the type 

species belongs in Siphonodendron and proposed the 

name genevievensis to replace Simpson's (1900) aggregatum because of the prior existence of Siphonodendron aggregatum McCoy. We now return to Simpson's 

name because we regard Schoenophyllum as distinct

from Siphonodendron. Schoenophyllum aggregatum is 

common in the Ste. Gene vie ve Limestone and its 

equivalents of Kentucky, Indiana, and Tennessee, where 

it was called Lithostrotion harmodites by earlier authors. 

Distribution. Schoenophyllum is a rare coral having 

a limited stratigraphic range in the Western Interior 

Province, where it is confined to Zone VA. It occurs in 

the Great Blue Limestone and Scott Peak Formation of 

Idaho and in the Etherington Formation of Alberta and 

southeastern British Columbia. The genus occurs exclusively in the shallow-water shelf-carbonate 

lithofacies.

Genus SCIOPHYLLUM Harker and McLaren, 1950

Plate 9, figures 3, 4

Diagnosis. Cerioid colonial rugose corals. Septa absent or reduced to fine vertical striae on inner side of 

theca or inside inner wall, rarely both in one corallite. 

Columella absent. Tabulae complete, flat, or slightly 

arched. Dissepimentarium consists of one or two rows 

of lonsdaleoid dissepiments.

Nominal species in Western Interior Province. S. adjunctivum (White, 1880).

Discussion. This genus is similar to "Thysanophyllum" but differs in its complete reduction of the septa and lack of even a vestige of a columella. It was probably derived from Acrocyathus in the Late Mississippian.

Distribution. Sciophyllum is rare in the upper part 

of Zone HID and the lower part of Zone IV. It has been 

found in the Scott Peak Formation and Monroe Canyon 

Limestone of Idaho, in an unnamed formation in 

Nevada, and in the Mount Head Formation of Alberta. 

It occurs exclusively in the shallow-water shelflimestone lithofacies.

Genus SIPHONODENDRON McCoy, 1849

Plate 5, figures 5-8; plate 6, figures 1, 2, 5-8

Diagnosis. Phaceloid colonial rugose corals. Columella simple, styliform. Major septa ordinarily long, 

may or may not extend to columella. Minor septa short. 

Tabulae large, conical, ordinarily supplemented by outer 

smaller and nearly horizontal tabulae. Dissepimentarium regular, continuous and wide in large species but 

may be narrow and discontinuous in small species.

Nominal species in Western Interior Province. S. 

mutabile (Kelly, 1942); S. oculinum (Sando, 1963); S. 

sinuosum (Kelly, 1942); S. warreni (Nelson, 1960); S. 

whitneyi of Meek, 1877 (not White, 1875).

Discussion. Similarities between Siphonodendron
28 CORAL ZONATION, MISSISSIPPIAN SYSTEM, WESTERN INTERIOR PROVINCE

and "Diphyphyllum" have been discussed under the latter. The nominal species are well defined, and their 

distribution in Canada is well known. Specimens from 

the United States have not been so thoroughly studied 

as the material from Canada. The problem of S. whitneyi 

of Meek has been discussed by Sando (1965, p. E17-E18); 

a new name is needed for this species.

Distribution. In Canada, Siphonodendron has a 

discontinuous range from the base of Zone IIB into the 

lower half of Zone V. It is represented by S. mutabile 

in Zone IIB, S. oculinum in Zones IIIA and B, and S. 

whitneyi of Meek in Zone HID. S. sinuosum ranges from 

Zone IIB to Zone IIIC, and S. warreni ranges from Zone 

HID into lowermost Zone IV. The genus occurs in the 

Banff, Livingstone, Shunda, Turner Valley, Mount 

Head, and Etherington Formations of Alberta; the 

Debolt, Prophet, Turner Valley, Mount Head, and Etherington Formations and the Milford and Rundle Groups 

of British Columbia; and the Flett Formation of the 

southwestern District of Mackenzie.

In the United States, the genus has a more restricted 

stratigraphic range, and fewer species are recognized. 

Many specimens have not been identified specifically. 

S. oculinum is very rare at the top of Zone IIB and is 

common in Zone IIIA. S. whitneyi of Meek is common 

in Zone HID and rare in Zone IV. The genus occurs in 

the Mission Canyon Limestone of Wyoming; the Mission 

Canyon Limestone and Castle Reef Dolomite of Montana; the Mission Canyon Limestone, and the Little Flat, 

Aspen Range, and Scott Peak Formations of Idaho; the 

Brazer Dolomite, Deseret Limestone, Humbug Formation, and Great Blue Limestone of Utah; the Leadville 

Limestone of Colorado; the Redwall Limestone of 

Arizona; the Banner Formation, Diamond Peak Formation, Goughs Canyon Formation, and Monte Cristo 

Limestone of Nevada; and, possibly, the Perdido Formation of California. The genus occurs exclusively in the 

shallow-water shelf-carbonate lithofacies.

Genus SIPHONOPHYLLIA Scouler in McCoy, 1844

Plate 2, figures 10-11

Diagnosis. Large trochoid to cylindrical solitary 

rugose corals. Major septa numerous, amplexoid in later 

growth stages. Minor septa short. Cardinal fossula well 

marked and on convex side of corallum, bounded axially by downturned edges of tabulae. Tabulae numerous, 

ordinarily complete, flat axially, and turned down at 

margins. Dissepimentarium broad, ordinarily 

lonsdaleoid.

Nominal species in Western Interior Province. 

Siphonophyllia enormis (Easton, 1945); S. excentrica 

(Meek, 1873); S. montanensis (Easton, 1962); S. nevadensis (Meek, 1877).

Discussion. Corals of this group in the Western Interior Province were previously identified as Caninia 

(Siphonophyllia?) because they have a dissepimentarium 

that is weakly lonsdaleoid. They differ from typical 

Siphonophyllia by having lonsdaleoid dissepiments in 

a medial zone between inner and outer zones of regular 

dissepiments. Among the nominal species, S. nevadensis, a cylindrical form, and S. excentrica, a conical form, 

may be different growth forms of the same species (see 

Sando, 1965, p. E24; 1975b. p. C19). S. enormis may be 

conspecific with S. excentrica. Siphonophyllia was probably derived from Caninia.

Distribution. Siphonophyllia is abundant in and 

characteristic of Zone V and has been found recently 

near the top of Zone VI at two localities in southwest 

Montana in the Big Snowy Formation (B. R. Wardlaw, 

oral commun., 1982). This genus is known from the 

Amsden Formation of Wyoming; the Big Snowy, Otter, 

and Heath Formations of Montana; the Monroe Canyon 

Limestone, the Surrett Canyon, Scott Peak, and Aspen 

Range Formations, and Great Blue Limestone of Idaho; 

the Green Ravine Formation, Great Blue Limestone, 

and Doughnut Formation of Utah; and the Etherington 

Formation of Alberta. The genus occurs almost exclusively in shallow-water lithofacies.

Genus STELECHOPHYLLUM Tolmachev, 1933 

Plate 9, figures 6-11

Diagnosis. Cerioid colonial rugose corals. Major septa thin, ordinarily extending across tabularium to columella but discontinuous or absent in dissepimentarium. Minor septa absent to variably developed. 

Columella ordinarily a simple axial rod or plate that 

may be derived from one or more major septa. 

Tabularium consisting of an axial series of bell-shaped 

to tent-shaped complete tabulae resting one upon the 

other and a weak peripheral series of horizontal or inclined concave-upward tabellae. Dissepimentarium 

lonsdaleoid.

Nominal species in Western Interior Province. S. banffense (Warren, 1927); S.? birdi Armstrong, 1970; S. circinatum (Easton and Gutschick, 1953); S. lochmanae 

(Armstrong, 1962); S.? macouni (Lambe, 1899); S.? 

mclareni (Sutherland, 1958); S.microstylum (White, 

1880); S. micrum (Kelly, 1942).

S. microstylum includes junior synonyms Lithostrotionella jasperensis Kelly, 1942; L confluens Easton, 

1958; and L. multiradiata Hayasaka, 1936. S. banffense 

includes junior synonyms Lithostrotionella floriformis 

Hayasaka, 1936, and L. vesicularis Hayasaka, 1936.

Discussion. This genus includes some of the corals 

previously referred to Lithostrotionella Yabe and 

Hayasaka, 1915. The taxonomy and world distribution
ZONAL TAXA 29

of Stelechophyllum have been discussed by Sando (1983). 

Stelechophyllum is the probable ancestor of Acrocyathus 

and Aulostylus.

Distribution. The principal occurrence of 

Stelechophyllum in the Western Interior Province of the 

United States is in the lower half of Zone II. S. 

microstylum, a Mississippi Valley species, occurs in the 

Lodgepole Limestone of Wyoming; the Lodgepole and 

Mission Canyon Limestones of Montana; the Lodgepole, 

Gardison, and Joana Limestones of Utah; the Lodgepole 

and Mission Canyon Limestones of Idaho; and the Joana 

and Monte Cristo Limestones of Nevada. S. circinatum 

is found at a slightly higher level in the upper half of 

Zone II in the Redwall Limestone of Arizona. Undetermined species of Stelechophyllum are rare in Zone IIIA 

in the Castle Reef Dolomite of Montana and in Zone HID 

in the Little Flat Formation of Idaho.

In Canada, the genus occurs in the same zones as it 

does in the United States. S. microstylum is found in the 

uppermost part of the Banff Formation and the lowermost part of the Pekisko Formation of Alberta and 

northeastern British Columbia. S. micrum is slightly 

higher, in the upper part of the Pekisko Formation and 

lower part of the Shunda Formation of Alberta and the 

lower part of the Mission Canyon Limestone of Saskatchewan. S. lochmanae occurs in the Pekisko Formation of 

Alberta. S. banffense is found in the Mount Head Formation of Alberta and the Prophet and Debolt Formations of northeastern British Columbia. S. ? macouni is 

in the Prophet Formation of British Columbia and the 

Mount Head Formation of Alberta. S. ? mclareni occurs 

in the Flett Formation of southwestern District of 

Mackenzie, the Prophet and Debolt Formations of northeastern British Columbia, the Mount Head Formation 

of Alberta, and the Milford Group of southeastern 

British Columbia. S. ? birdi is found in the Prophet Formation of northeastern British Columbia. Unnamed 

species occur in the Flett Formation of southwestern 

District of Mackenzie and in the Prophet, Pekisko, and 

Debolt Formations of northeastern British Columbia.

Stelechophyllum occurs exclusively in shallow-water 

carbonate lithofacies. Its principal occurrence is in the 

shallow-water basinal-carbonate facies.

Genus SYCHNOELASMA Lang, Smith, and Thomas, 1940

Plate 9, figure 5

Diagnosis. Ceratoid to trochoid, slightly curved 

solitary rugose corals having a slit-shaped cardinal 

fossula on the convex side of the corallum. Major septa 

long, radially arranged, axial ends fused to form axial 

stereozone. Minor septa long, peripheral ends fused with 

major septa to form peripheral stereozone. Tabulae upwardly arched. Dissepiments absent.

Nominal species in Western Interior Province. S. subcrassum (Easton and Gutschick, 1953), S. ulrichanum 

(Girty, 1903). Triplophyllites (Homalophyllites) circularis 

Easton, 1958, may be a junior synonym of S. subcrassum. S. ulrichanum includes the junior synonym 

Triplophyllites (Homalophyllites) paucicinctus Easton 

and Gutschick, 1953.

Discussion. Sychnoelasma includes corals previously referred to Homalophyllites Easton, 1944 (e.g., Sando, 1960b, 1969). We agree with Sayutina's (1976) placement of Homalophyllites as a junior synonym of 

Sychnoelasma. We do not agree with Sayutina's (1976) 

conclusion that Menophyllum Milne-Edwards and 

Haime, 1850, is identical with Sychnoelasma. 

Sychnoelasma is a probable derivative of Amplexizaphrentis and is regarded as the ancestor of 

Ankhelasma.

Sychnoelasma is one of the most common corals in the 

Mississippian of the Western Interior Province. Species 

in addition to the nominal species may be present, 

because large collections of this genus have not been 

studied in detail.

Distribution. In the United States, the principal 

range of the genus is from the base to the top of Zone 

II, but it is rare in Zone 1C and in the lowermost part 

of Zone IIIA. The nominal species are in the Redwall 

Limestone of Arizona and the Leadville Limestone of 

Colorado. Undetermined species occur in the Madison, 

Mission Canyon, and Lodgepole Limestones of Wyoming; the Lodgepole Limestone, Mission Canyon 

Limestone, Allan Mountain Limestone, and Castle Reef 

Dolomite of Montana; the Lodgepole and Mission Canyon Limestones of Idaho; the Lodgepole Limestone, 

Brazer Dolomite, Gardison Limestone, and Joana 

Limestone of Utah; the Monte Cristo and Joana 

Limestones of Nevada; and the Tin Mountain Limestone 

of California.

In Canada, Sychnoelasma is common from the base to 

near the top of Zone II and is rare in uppermost Zone 

II and lowermost Zone IIIA. It occurs in the Lodgepole 

and Mission Canyon Limestones of Saskatchewan; the 

Banff, Pekisko, Shunda, and Livingstone Formations of 

Alberta; and the Banff, Pekisko, and Shunda Formations of northeastern British Columbia.

Sychnoelasma is found in both shallow-water and deepwater lithofacies, but its predominant occurrence is in 

shallow-water habitats.

Genus SYRINGOPORA Goldfuss, 1826 

Plate 9, figures 1, 2

Diagnosis. Phaceloid tabulate corals having axial infundibular tabulae, randomly arranged connecting 

tubes, and septal spines.
30 CORAL ZONATION, MISSISSIPPIAN SYSTEM, WESTERN INTERIOR PROVINCE

Nominal species in Western Interior Province. 

Recognition of nominal species in Syringopora depends 

on a choice between different systems of classification 

(see discussion below). For the present, no nominal 

species are recognized because of the difficulty in using 

existing species concepts in zonation.

Discussion. Studies by Sando (1960b, 1969) of Syringopora in the United States resulted in recognition 

of four "species" in the Madison Group of Montana and 

three "species" in the Redwall Limestone of Arizona. 

These "species" are intergrading morphogroups that 

have no apparent value in zonation because each morphogroup has about the same stratigraphic range.

In Canada, Nelson (1962a) recognized nine species and 

two formae in the Mississippian of Alberta. Nelson 

(1977) revised his earlier work and proposed a system 

that lumped the Canadian representatives of Syringopora in two species and recognized four subspecies 

that he thought had stratigraphic significance. Nelson's 

taxa are based on rather subtle morphologic distinctions 

of questionable biologic and stratigraphic significance. 

Inasmuch as we have not had time to test Nelson's concepts, we have not accepted his system of classification.

Syringopora is similar to Pleurosiphonella and 

Multithecopora; differences between Syringopora and 

these taxa are discussed under the other taxa.

Distribution. Syringopora is one of the most common 

corals in the Mississippian of the Western Interior Province. It occurs in both deep-water and shallow-water 

lithofacies but is much more common in shallow-water 

habitats. Syringopora is one of the few genera that range 

from the Devonian into the Mississippian.

In the United States, Mississippian occurrences of the 

genus range from the base of Zone IA to the top of Zone 

IV, but the genus is most common from the base of Zone 

IB to the top of Zone IIIA. It occurs in the Madison, 

Lodgepole, Mission Canyon, and Pahasapa Limestones 

of Wyoming; the Allan Mountain Limestone, Castle Reef 

Dolomite, Lodgepole Limestone, Charles Formation, and 

Mission Canyon Limestone of Montana; the Lodgepole 

Limestone, Mission Canyon Limestone, Little Flat Formation, and Scott Peak Formation of Idaho; the 

Pahasapa Limestone of South Dakota; the Mission Canyon Limestone of North Dakota; the Leadville 

Limestone of Colorado; the Lodgepole Limestone, Brazer 

Dolomite, Gardison Limestone, Deseret Limestone, 

Humbug Formation, Joana Limestone, and Fitchville 

Formation of Utah; the Redwall Limestone of Arizona; 

the Monte Cristo Limestone, Joana Limestone, Mercury 

Limestone, Goughs Canyon Formation, and Bristol Pass 

Limestone of Nevada; and the Tin Mountain Limestone 

of California.

In Canada, Mississippian Syringopora ranges from the 

upper part of Zone I to the top of Zone IV and is most

common in Zone IIB. The genus occurs in the Banff, 

Pekisko, Shunda, Turner Valley, and Mount Head Formations of Alberta; the Banff Formation, Rundle Group, 

and Milford Group of British Columbia; and the 

Lodgepole and Mission Canyon Limestones of 

Saskatchewan.

Genus THYSANOPHYLLUM Nicholson and Thomson, 1876 

Plate 10, figures 1, 2

Diagnosis. Cerioid colonial rugose corals. Major septa do not extend to axis of corallum. Minor septa may 

be present. Columella absent or poorly developed and 

impersistent, may be attached to counter septum. 

Tabulae complete, ordinarily flat or sagging, having 

downturned edges. Dissepimentarium lonsdaleoid.

Nominal species in Western Interior Province. "T." 

astreiforme (Warren, 1927).

Discussion. Corals included in this taxon may be noncolumellate variants of Stelechophyllum or Acrocyathus. 

The name "Thysanophyllum" is used in quotes to denote 

taxonomic uncertainty.

Distribution. "Thysanophyllum" is a rare coral in the 

Western Interior Province. In Canada, the nominal 

species is found in the upper part of Zone HID and in 

Zone IV; an undetermined species occurs in Zone IIIA. 

The nominal species is in the Mount Head Formation 

of Alberta and the Prophet Formation of northeastern 

British Columbia; an undetermined species occurs in the 

Debolt Formation of northeastern British Columbia.

In the United States, the taxon is known from single 

occurrences in Zones IIB, IIIA, and HID (questionable). 

It is found in the Mission Canyon Limestone of Wyoming, the Castle Reef Dolomite of Montana, the Perdido 

Formation of California (questionable), and the Scott 

Peak Formation of Idaho. It occurs exclusively in the 

shallow-water shelf-carbonate lithofacies.

Genus TURBINATOCANINIA Dobrolyubova, 1970 

Plate 10, figures 5, 6

Diagnosis. Solitary, curved or straight, conical rugose 

corals. Major septa well developed, sometimes thickened 

in cardinal quadrants in tabularium. Minor septa thin, 

confined to dissepimentarium. Young stages have a complex columella composed of axial plate, radial lamellae, 

and tabellae, but adult stages do not have a columella. 

Tabulae horizontal except at axis where they are turned 

up. Dissepimentarium regular.

Nominal species in Western Interior Province. None.

Discussion. Sando (1975b, p. C19) referred fragmentary corals having a columella in young stages and the 

morphology of Caninia in adult stages to this genus. The
ZONAL TAXA 31

name is used with a query because the fragmentary 

nature of the specimens leaves doubts concerning their 

identity.

Distribution. Turbinatocaniniat is a rare coral found 

only in Zones V and VI of the Western Interior Province 

of the United States. It occurs in the Amsden Formation of Wyoming and the Heath Formation of Montana. 

It may also be present in the Monroe Canyon Limestone 

of Idaho. This taxon occurs exclusively in the shallowwater shelf-carbonate lithofacies.

Genus VESICULOPHYLLUM Easton, 1944 

Plate 10, figures 7, 8

Diagnosis. Large solitary cylindrical rugose corals 

having numerous major septa that ordinarily closely approach the axis of the corallum but may be short. Septal plan bilateral, palmate, or radial. Cardinal septum 

long or short, may or may not be in a fossula. Counter 

septum ordinarily short. Septa ordinarily dilated, dilation confined to axial region of corallum in later growth 

stages. Minor septa short and impersistent or absent. 

Dissepimentarium broad, consisting of an outer zone of 

lonsdaleoid dissepiments and an inner zone of regular 

dissepiments. Tabulae numerous, incomplete, steeply inclined axially or bowlshaped, gradational with 

dissepiments.

Nominal species in Western Interior Province. V. 

bilateralis (Easton, 1945), V. burlingi (Sutherland, 1958), 

V. dux (Sutherland, 1954), V. incrassatum (Easton and 

Gutschick, 1953). V. incrassatum includes junior 

synonyms Caninophyllum sonorense Easton, 1958, 

Enygmophyllum californiense Langenheim and Tischler, 

1960, and Vesiculophyllum sutherlandi Armstrong, 

1962.

Discussion. Vesiculophyllum is characterized by extreme ontogenetic and individual variation (see Sando, 

1969, p. 298-305). It has a weak outer zone of 

dissepiments that was commonly eroded away prior to 

deposition; most specimens are fragmentary. Because 

these characteristics cause great difficulty in discrimination of species, the number of species present in the 

Western Interior Province is unknown. Some of the 

nominal species listed above may be synonyms in addition to synonyms already determined. Ultimately, the 

large number of specimens from many localities may 

have to be referred to a single species because of difficulties in discrimination resulting from such great 

variation in a single collection.

Vesiculophyllum is a member of the Family 

Uraliniidae, a group of solitary rugose corals 

characterized by their peculiar septal plan, concave 

tabulae, and well-developed dissepimentarium. In North

America, Kakwiphyllum Sutherland, 1954, is a junior 

synonym. Clinophyllum Grove, 1935, may be merely a 

young stage of Vesiculophyllum. Among Russian 

representatives of the family, Enygmophyllum 

Fomichev, 1931, Humboldtia Stuckenberg, 1895, and 

Keyserlingpphyllum Stuckenberg, 1895, may be senior 

synonyms; these genera have not been investigated in 

sufficient detail for a decision on synonymy.

Distribution. Vesiculophyllum is the most common 

coral in the Mississippian of the Western Interior Province. In the United States, specimens questionably referred to the genus were found in the uppermost Devonian (Famennian) and in Zone IA of the Mississippian. 

The genus is rare in Zones IB and C, then becomes common from the base of Zone IIA to the top of Zone IIIA. 

It is rare in Zones IIIB and C. The genus is found in the 

Madison, Lodgepole, and Mission Canyon Limestones of 

Wyoming; the Allan Mountain Limestone, Castle Reef 

Dolomite, Lodgepole Limestone, Mission Canyon 

Limestone, and Charles Formation of Montana; the Mission Canyon Limestone of North Dakota; the Lodgepole 

and Mission Canyon Limestones of Idaho; the Lodgepole 

Limestone, Brazer Dolomite, Gardison Limestone, Joana 

Limestone, and Fitchville Formation of Utah; the 

Pahasapa Limestone of South Dakota; the Tin Mountain Limestone of California; the Redwall Limestone of 

Arizona; the Leadville Limestone of Colorado; and the 

Joana, Monte Cristo, Mercury, and Bristol Pass 

Limestones of Nevada.

In Canada, Vesiculophyllum ranges from the upper 

part of Zone I into Zone IIIC. It occurs in the Lodgepole 

Limestone, Mission Canyon Limestone, and Charles Formation of Saskatchewan; the Banff, Pekisko, Shunda, 

Turner Valley, and Livingstone Formations of Alberta; 

the Banff, Prophet, Pekisko, Shunda, Debolt, and Turner 

Valley Formations of British Columbia; and the Flett 

Formation of the southwestern District of Mackenzie. 

Vesiculophyllum occurs predominantly in shallow-water 

lithofacies.

Genus ZAPHRIPHYLLUM Sutherland, 1954 

Plate 10, figures 3, 4

Diagnosis. Small to medium-sized trochoid solitary 

rugose corals having numerous long major septa that 

tend to be radially arranged. Cardinal fossula well 

developed on convex side of corallum. Alar fossulae present. Minor septa moderately long. Tabulae mostly incomplete, arched periaxially and downturned into cardinal fossula. Dissepimentarium regular, well developed 

in ephebic stage.

Nominal species in Western Interior Province. Z. 

disseptum Sutherland, 1954.
32 CORAL ZONATION, MISSISSIPPIAN SYSTEM, WESTERN INTERIOR PROVINCE

Discussion. This taxon includes some corals previously referred to Canadiphyllum with a query (see Sando, 

Dutro, and Gere, 1959). Zaphriphyllum is like the 

"zaphrentoid" corals in having a simple skeleton but differs in having a well-developed dissepimentarium at 

maturity. The genus may have been derived from 

Lophophyllum?.

Distribution. In the United States, Zaphriphyllum 

has a sporadic range from the lower part of Zone IIB to 

the top of Zone IIIA; it is most common in the lower half 

of Zone IIIA. It occurs in the Madison and Mission Canyon Limestones of Wyoming; the Mission Canyon 

Limestone, Charles Formation, and Castle Reef 

Dolomite of Montana; the Mission Canyon Limestone 

of Idaho; and the Brazer Dolomite of Utah.

In Canada, the genus ranges from the middle of Zone 

IIB into Zone IIIC; it is most common from the upper 

part of Zone IIB to the middle of Zone IIIB. It occurs in 

the Banff, Pekisko, Shunda, Turner Valley, Livingstone, 

and Mount Head Formations of Alberta; the Prophet, 

Debolt, Pekisko, Shunda, and Turner Valley Formations 

of northeastern British Columbia; and the Flett Formation of southwestern District of Mackenzie. 

Zaphriphyllum occurs exclusively in the shallow-water 

shelf-carbonate lithofacies.

LOCALITY REGISTER 

UNITED STATES

The following list includes (1) localities represented by 

detailed measured sections (in plate captions, referred 

to as "sections") and (2) localities not represented by 

such sections (in plate captions, referred to as 

"localities"). Localities not represented by sections are 

mostly taken from unpublished reports on referred collections by Sando and other members of the U.S. 

Geological Survey. Many of these localities are composite, segregated by township in the land grid system. 

Localities in both categories are listed alphabetically by 

State. Geographic location, published or unpublished 

references, coralliferous formations, coral zones identified, and a list of genera and subgenera present are 

given for each locality.

DETAILED MEASURED SECTIONS 

ARIZONA

A-l. North Kaibab Trail, 0.75 mi east of Grand Canyon Lodge, 

Coconino County (see Sando, 1969). Redwall Limestone. Zone 

II. Sychnoelasma, Syringopora, Vesiculophyllum.

A-2. Havasu Canyon, 2 mi northwest of Supai Indian Village, 

Coconino County (see Sando, 1969). Redwall Limestone. 

Zones II, IIIB. Dorlodotia?, Sychnoelasma, Syringopora, 

Vesiculophyllum.

A-3. Whitmore Wash, 16 mi southeast of Mt. Trumbull P.O. in 

T. 32 N., R. 8 W., Coconino County (see Sando, 1969). Redwall Limestone. Zones II, IIIB. Dorlodotia?, Sychnoelasma, 

Syringopora, Vesiculophyllum.

A-4. Parashant Canyon, 13 mi south of Mt. Trumbull P.O. in 

T. 32 N., R. 9 W., Coconino County (see Sando, 1969). Redwall Limestone. Zones II, IIIB. Dorlodotia?, Syringopora, 

Vesiculophyllum.

A-5. Grand Wash Cliffs, south fork of Pigeon Wash, 6 mi north 

of Snap Point in T. 34 N., R. 14 W., Coconino County (see 

Sando, 1969). Redwall Limestone. Zones II, mC. Dorlodotia?, 

Ekvasophyllum, Vesiculophyllum.

A-6. Pakoon, Whitney Ridge, Virgin Mountains in T. 36 N.(?), 

R. 16 W.,Coconino County (see Sando, 1969). Redwall 

Limestone. Zones II, IIIB, IIIC. Amplexizaphrentis, Caninia, 

Dorlodotia?, Ekvasophyllum, Syringopora, Vesiculophyllum. 

A-7. Iceberg Canyon, west side of Iceberg Ridge, Lake Mead 

Recreation Area in T. 32 N., R. 16 W., Mohave County (see 

Sando, 1969). Redwall Limestone. Zones II, IIIB, IIIC?. 

Amplexizaphrentis, Cyathaxonia, Dorlodotia?, Syringopora, 

Vesiculophyllum.

A-8. Quartermaster Canyon, Hualapai Indian Reservation in 

T. 30 N., R. 14 W., Mohave County (see Sando, 1969). Redwall Limestone. Amplexizaphrentis, Dorlodotia?, 

Sychnoelasma, Syringopora, Vesiculophyllum. 

A-9. Bridge Canyon, east side of canyon, near corner of sees. 10 

and 14, T. 27 N., R. 12 W., Mohave County (see Sando, 1969). 

Redwall Limestone. Zones II, IIIA, IIIB. Amplexizaphrentis, 

Dorlodotia?, Siphonodendron, Sychnoelasma, Syringopora, 

Vesiculophyllum.

A-10. Hindu Canyon, sec. 23, T. 27 N., R. 12. W., Mohave County (see Sando, 1969). Redwall Limestone. Zones II, IIIA. 

Amplexizaphrentis, Siphonodendron, Sychnoelasma, 

Vesiculophyllum.

A-ll. Diamond Creek, sees. 4, 5, T. 27, N., R. 10 W., Mohave 

County (see Sando, 1969). Redwall Limestone. Zone IIIA. 

Siphonodendron, Syringopora.

A-12. Metuck Canyon, 3 to 4 mi north of Metuck Tank, Coconino 

County (see Sando, 1969). Redwall Limestone. Zones IIIA, 

IIIB. Dorlodotia?, Siphonodendron, Syringopora, 

Vesiculophyllum.

A-13. Peach Springs-Nelson, east side of gully 0.3 mi east of 

Nelson Station and quarry at Nelson station, Mohave County (see Sando, 1969. Redwall Limestone. Zones II, IIIA, IIIB. 

Dorlodotia?, Siphonodendron, Sychnoelasma, Syringopora, 

Vesiculophyllum.

A-14. Ring Cone, near center sec. 20, T. 23 N., R. 11 W., Mohave 

County (see Sando, 1969). Redwall Limestone. Zone II. 

Amplexizaphrentis, Vesiculophyllum.

A-15. Seligman Field, sec. 4, T. 22 N., R. 8 W., Yavapai County 

(see Sando, 1969). Redwall Limestone. Zone II. Sychnoelasma, Vesiculophyllum.

A-16. Chino Point, sec. 3, T. 22 N., R. 7 W., Yavapai County (see 

Sando, 1969). Redwall Limestone. Zone II. Sychnoelasma?, 

Vesiculophyllum.

A-17. Pichacho Butte, sec. 18, T. 21 N., R. 4 W., Yavapai County 

(see Sando, 1969). Redwall Limestone. Zone II. Syringopora.

A-18. Black Mesa composite, along Black Mesa north of Chino 

Valley from U.S. Highway 89 West to South Butte, Yavapai 

County (see Sando, 1969). Redwall Limestone. Zone II.
LOCALITY REGISTER

Amplexizaphrentis, Michelinia, Sychnoelasma, Syringopora, 

Vesiculophyllum.

A-19. Drake quarry, sec. 32, T. 19 N., R. 1 W., Yavapai County (see 

Sando, 1969). Redwall Limestone. Zone II. Amplexizaphrentis, Michelinia, Stelechophyllum, Sychnoelasma, Syringopora, Vesiculophyllum.

A-20. Simmons, sec. 1, T. 16 N., R. 3 W., Yavapai County (see Sando, 1969). Redwall Limestone. Zone II. Vesiculophyllum.

A-21. Lonesome Valley, sec. 14, T. 16. N., R. 1 W., Yavapai 

County (see Sando, 1969). Redwall Limestone. Zone II. 

Amplexizaphrentis.

A-22. Sycamore Canyon, sees. 31, 32, T. 18 N., R. 3 E., Yavapai 

County (see Sando, 1969). Redwall Limestone. Zone II. 

Sychnoelasma, Syringopora, Vesiculophyllum.

A-23. Mingus Pass composite, in roadcut on north side of 

U.S. Highway 89, about 5 mi west of Jerome and in sec. 12, 

T. 16. N., R. 1 E., Yavapai County (see Sando, 1969). Redwall Limestone. Zone II. Amplexizaphrentis, Michelinia, Stelechophyllum, Sychnoelasma, Syringopora, Vesiculophyllum.

A-24. Jerome composite, sec. 21, T. 16 N., R. 2 E., Yavapai County (see Sando, 1969). Redwall Limestone. Zone II. Amplexizaphrentis, Michelinia, Sychnoelasma, Syringopora, Vesiculophyllum.

A-25. Clemenceau quarry, sec. 7, T. 15 N., R. 3 E., Yavapai 

County (see Sando, 1969). Redwall Limestone. Zone II. 

Amplexizaphrentis, Vesiculophyllum.

A-26. Natural Bridge, sec. 8, T. 11 N., R. 9 E., Gila County (see 

Sando, 1969). Redwall Limestone. Zone II. Amplexizaphrentis, Syringopora.

A-27. Pine composite, sees. 6, 7, T. 11 N., R. 10 E., Gila County (see 

Sando, 1969). Redwall Limestone. Zone II. Amplexizaphrentis, Cyathaxonia, Michelinia, Sychnoelasma, Syringopora, 

Vesiculophyllum.

A-28. Colcord Canyon, sec. 34, T. 10 1/2 N., R. 14 E., Gila County (see Sando, 1969). Redwall Limestone. Zone II. 

Stelechophyllum, Sychnoelasma, Syringopora, Vesiculophyllum.

A-29. O. W. Ranch 2, sec. 33, T. 10 1/2 N., R. 15 E., Gila County 

(see Sando, 1969). Redwall Limestone. Zone II. 

Sychnoelasma, Syringopora, Vesiculophyllum.

A-30. Brush Mountain N.W., sec. 5, T. 7 N., R. 16 E., Navajo 

County (see Sando, 1969). Redwall Limestone. Zone II. 

Michelinia, Vesiculophyllum.

A-31. Salt River Draw 1, sec. 24, T. 7 N., R. 16. E., Gila County 

(see Sando, 1969). Redwall Limestone. Zone II. Michelinia, 

Sychnoelasma, Vesiculophyllum.

A-32. Salt River Draw 2, about 1 mi north of Rock House Butte, 

Gila County (see Sando, 1969). Redwall Limestone. Zone II. 

Michelinia.

A-33. Salt River U.S. 60, sec. 20, T. 5 N., R. 18. E., Gila County 

(see Sando, 1969). Redwall Limestone. Zone II. Amplexizaphrentis, Caninia?, Michelinia, Sychnoelasma, Syringopora, Vesiculophyllum.

A-34. Black River Crossing, sec. 11, T. 4 N., R. 20 E., Gila County (see Sando, 1969). Redwall Limestone. Zones I, II. Amplexizaphrentis, Amplexus, Lophophyllum?, Michelinia, Syringopora, Vesiculophyllum.

CALIFORNIA

CA-1. Tin Mountain type section, southern slope of hills about 

2.5 mi southeast of Quartz Spring and about 3,000 ft north

of road to Rest Spring, Inyo County (see Langenheim and 

Tischler, 1960). Tin Mountain Limestone. Zone II. Amplexizaphrentis, Amplexus, Lophophyllum?, Vesiculophyllum. 

CA-2. Funeral Mountains, Hill 2997, northeast end of Funeral 

Mountains, Ryan quadrangle, Inyo County (see McAllister, 

1974). Tin Mountain Limestone. Zones I?, II. Amplexizaphrentis, Amplexus, Caninia?, Cyathaxonia, Guerichophyllum, Lophophyllum?, Michelinia, Sychnoelasma, Syringopora, Vesiculophyllum.

IDAHO

1-1. Black Mountain, sees. 23, 24, T. 3 N., R. 43 E., Bonneville 

County (see Sando, 1977c). Lodgepole and Mission Canyon 

Limestones. Zones 1C, II, IIIA. Amplexizaphrentis, Amplexus?, Cyathaxonia, "Diphyphyllum," Guerichophyllum, Stelechophyllum, Sychnoelasma, Syringopora, Vesiculophyllum, 

Zaphriphyllum.

1-2. Sheep Creek, sec. 28, T. 1 N., R. 45 E., Bonneville County 

(see Sando, 1977c). Lodgepole and Mission Canyon 

Limestones. Zones II, IIIA, Amplexizaphrentis, Ankhelasma, 

Cyathaxonia, Lophophyllum?, Michelinia, Siphonodendron, 

Stelechophyllum, Sychnoelasma, Syringopora, Vesiculophyllum, Zaphriphyllum.

1-3. Little Flat Canyon, sees. 17, 20, T. 7 S., R. 40 E., Bannock 

County (see Dutro and Sando, 1963). Lodgepole Limestone, 

Little Flat Formation, and Monroe Canyon Limestone. 

Zones IB, 1C, II, IIID, IV, VA, VB. Acrocyathus (cerioid), Amplexizaphrentis, Amplexus, Canadiphyllum, Clisiophyllum, 

Cyathaxonia, "Diphyphyllum," Duncanopora, Ekvasophyllum, Faberophyllum, Guerichophyllum, Lophophyllum?, 

Metriophyllum, Michelinia, Multithecopora, Palaeacis, Pleurosiphonella, "Pseudodorlodotia," Siphonodendron, Siphonophyllia, Sychnoelasma, Syringopora, Turbinatocaninia?, 

Vesiculophyllum.

1-4. Hunter Canyon, sec. 35, T. 9 S., R. 31 E., Power County (see 

Trimble and Carr, 1976). Lodgepole Limestone, Deep Creek 

Formation, and Great Blue Limestone. Zones II, IV, V. 

Amplexizaphrentis, Amplexus, Canadiphyllum, Clisiophyllum, "Diphyphyllum," Faberophyllum, Lophophyllum?, 

Pleurosiphonella, Rhopalolasma, Rotiphyllum, Schoenophyllum, Siphonophyllia, Stelechophyllum, Sychnoelasma, Syringopora, Vesiculophyllum.

1-5. East Canyon, sec. 1, T. 6 N., R. 29 E., Butte County (see Huh, 

1968). Scott Peak and Surrett Canyon Formations. 

Zones IIIB, IIIC, HID, IV, VA, VB, VI. Amplexizaphrentis, 

Barytichisma, Duncanopora, Ekvasophyllum, Faberophyllum, Multithecopora, Pleurosiphonella, "Pseudodorlodotia," 

Siphonodendron, Siphonophyllia.

1-6. Hawley Mountain, sees. 24, 25, T. 9 N., R. 26 E., Butte 

County (see Mamet and others, 1971). Middle Canyon, Scott 

Peak, and Surrett Canyon Formations. Zones IIIB, IIIC, IIID, 

IV, VA, VB, VI. Acrocyathus (cerioid), Actinocyathus, 

Amplexizaphrentis, Amplexus, Barytichisma, Canadiphyllum, Caninia, Cyathaxonia, Duncanopora, Ekvasophyllum, 

Faberophyllum, Multithecopora, Palaeacis, Petalaxis, Pleurosiphonella, "Pseudodorlodotia," Rotiphyllum, Siphonodendron, Siphonophyllia, Syringopora?.

1-7. Devils Canyon, sec. 6, T. 12 N., R. 21 E., Custer County (see 

Mamet and others, 1971). Scott Peak Formation. Zones IIID, 

IV. Amplexizaphrentis, Canadiphyllum, Faberophyllum, 

Pleurosiphonella, "Pseudodorlodotia."
34 CORAL ZONATION, MISSISSIPPIAN SYSTEM, WESTERN INTERIOR PROVINCE

1-8. McGowan Creek, sec. 28, T. 12 N., R. 21 E., Custer County 

(see Mamet and others, 1971). Scott Peak Formation. 

Zones HID, IV. Amplexizaphrentis, Cyathaxonia, Faberophyllum, Rotiphyllum, Syringopora.

1-9. Christian Gulch, sees. 32, 33, T. 12 N., R. 22 E. (see Mamet 

and others, 1971). Scott Peak and Surrett Canyon Formations. Zones HID, IV, VA, VB. Actinocyathus, Amplexizaphrentis, Duncanopora, Faberophyllum, Multithecopora, 

Ple.urosiphone.lla, "Pseudodorlodotia," Siphonodendron, 

Siphonophyllia, Syringopora.

1-10. Van Dorn Canyon, Sec. 11, T. 8 N., R. 26 E., Butte County 

(see Mamet and others, 1971). Surrett Canyon Formation. 

Zone VB. Duncanopora, Siphonophyllia.

1-11. Cherry Creek, sees. 32, 33, T. 5 N., R. 24 E., Custer County (see Skipp, 1961). Scott Peak Formation. Zone IV. Clisiophyllum, Faberophyllum, Syringopora.

1-12. Timbered Dome, sec. 3, T. 4 N., R. 25 E., Butte County (see 

Skipp, 1961). Surrett Canyon Formation. ZoneV. Duncanopora, "Pseudodorlodotia," Siphonophyllia.

1-13. Cabin Creek, sees. 13,14,15, T. 6 N., R. 22 E., Custer County (see Skipp and Mamet, 1970). White Knob Limestone. 

Zone indeterminate. Michelinia, Rotiphyllum^.

1-14. Wood Canyon, sees. 29, 30, T. 5 N., R. 25 E. Custer County 

(Betty Skipp, unpublished data, 1970). Scott Peak and Surrett Canyon Formations. Zones IV, VA, VB. Amplexizaphrentis, Ekvasophyllum?, Faberophyllum, Multithecopora, Pleurosiphonella, Rotiphyllum?, Siphonophyllia.

1-15. North Georgetown Canyon, sees. 16, 21, T. 10 S., R. 44 E., 

Bear Lake County (Sando, Sandberg, and Gutschick, 1981). 

Aspen Range Formation and Mission Canyon Limestone. 

Zones IIB, IIIC, HID, IV, VB. Acrocyathus (cerioid) Amplexizaphrentis, Canadiphyllum, Clisiophyllum, Cyathaxonia, 

Duncanopora, Ekvasophyllum, Faberophyllum, Hapsiphyllum, Lophophyllum?, Michelinia, Multithecopora, Pleurosiphonella, "Pseudodorlodotia," Rotiphyllum, Siphonodendron, Siphonophyllia, Sychnoelasma, Syringopora, 

Vesiculophyllum.

MONTANA

M-l. Diversion Dam and Hannan Gulch, sees. 35, 36, T. 22 N., 

R. 9 W., Teton County (see Mudge, Sando, and Dutro, 1962). 

Allan Mountain Limestone and Castle Reef Dolomite. 

Zones II, IIIA. Amplexizaphrentis, Ankhelasma, Caninia, Cyathaxonia, Lophophyllum?, Siphonodendron, Stelechophyllum, Sychnoelasma, Syringopora, "Thysanophyllum," Vesiculophyllum, Zaphriphyllum.

M-2. Gibson Dam and Gibson Reservoir, sees. 4, 6, T. 21 N., 

R. 9 W., and sec. 31, T. 22 N., R. 9 W., Teton County (see 

Mudge, Sando, and Dutro, 1962). Allan Mountain Limestone 

and Castle Reef Dolomite. Zones IB, 1C, II, IIIA. Amplexizaphrentis, Amplexus, Cyathaxonia, Hapsiphyllum, Liardiphyllum, Lophophyllum?, Metriophyllum, Michelinia, 

Neaxon?, Pleurosiphonella, Rotiphyllum, Sychnoelasma, Syringopora, Vesiculophyllum, Zaphriphyllum..

M-3. Mission Canyon, sec. 32, T. 26 N., R. 24 E., Blaine County 

(see Sando and Dutro, 1974). Mission Canyon Limestone. 

Zone II. Lophophyllum?, Michelinia, Sychnoelasma, Syringopora, Vesiculophyllum.

M-4. Little Chief Canyon, sees. 19, 30, T. 26 N., R. 25 E., Blaine 

County (see Sando and Dutro, 1974). Lodgepole and Mission

Canyon Limestones. Zones I, II. Amplexizaphrentis, Amplexocarinia, Amplexus, Caninia, Cleistopora, Cyathaxonia, Lophophyllum?, Michelinia, Stelechophyllum, Sychnoelasma, 

Syringopora, Vesiculophyllum.

M-5. California Co., Grimm No. 1 well core, sec. 13, T. 32 N., 

R. 49 E., Roosevelt County (see Sando, 1978). Mission Canyon Limestone. Zones II, IIIA. Siphonodendron, Vesiculophyllum.

M-6. Hodge, Smith, and Hodge Co., No. 1 Eggebrecht well core, 

sec. 3, T. 23 N., R. 49 E., McCone County (see Sando, 1978). 

Mission Canyon Limestone and Charles Formation. Zones II, 

IIIA. Amplexizaphrentis, Cyathaxonia, "Diphyphyllum,"Lophophyllum?, Michelinia, Siphonodendron, Sychnoelasma, 

Syringopora, Vesiculophyllum.

M-7. Shell Oil Co., Richey area, Northern Pacific Railroad No. 1 

well core, sec. 19, T. 23 N., R. 50 E., Dawson County (see 

Sando, 1978). Lodgepole Limestone, Mission Canyon Limestone, and Charles Formation. Zones II, IIIA. Amplexizaphrentis, "Diphyphyllum," Lophophyllum?, Michelinia, 

Sychnoelasma, Syringopora, Vesiculophyllum, Zaphriphyllum?.

M-8. C. H. Murphy Co., East Poplar Unit No. 1 well core, sec. 2, 

T. 28 N., R. 51 E., Roosevelt County (see Sando, 1978). 

Lodgepole and Mission Canyon Limestones. Zones II, IIIA. 

Amplexizaphrentis, "Diphyphyllum," Lophophyllum?, Rotiphyllum, Siphonodendron, Sychnoelasma, Syringopora, Vesiculophyllum.

M-9. Shell Oil Co., Pine Unit No. 1 well core, sec. 30, T. 12 N., 

R. 57 E., Wibaux County (see Sando, 1978). Lodgepole and 

Mission Canyon Limestones and Charles Formation. Zones I, 

II, IIIA. Amplexizaphrentis, Amplexocarinia, Amplexus, Caninia, Cleistopora, Cyathaxonia, "Diphyphyllum,"Lophophyllum?, Pleurosiphonella, Sychnoelasma, Syringopora, Vesiculophyllum, Zaphriphyllum.

M-10. Damm No. F-33-23-P well core, sec. 23, T. 29 N., R. 54 E., 

Roosevelt County (see Sando, 1978). Mission Canyon 

Limestone. Zones II, IIIA?. Siphonodendron?, Syringopora, 

Vesiculophyllum.

M-ll. Dry Fork, sec. 36, T. 16 N., R. 7 E., Cascade County (see 

Sando and Dutro, 1974). Lodgepole and Mission Canyon 

Limestones. Zones I, II. Amplexizaphrentis, Amplexus, Aulostylus, Cyathaxonia, Guerichophyllum, Lophophyllum?, 

Michelinia, Stelechophyllum, Sychnoelasma, Syringopora, 

Vesiculophyllum.

M-12. Monarch-U.S. 89, sees. 22, 27, T. 16 N., R. 7 E., Cascade 

County (see Sando and Dutro, 1974). Mission Canyon 

Limestone. Zones II, IIIA. Amplexizaphrentis, "Diphyphyllum"?, Lophophyllum?, Stelechophyllum, Sychnoelasma, 

Syringopora, Vesiculophyllum.

M-13. Indian Creek, sec. 32, T. 7 N., R. 1 E., Broadwater County 

(W. J. Sando, unpublished data, 1962). Big Snowy Formation. Zone V. Siphonophyllia?.

M-14. Stonehouse Canyon, sec. 25, T. 11 N., R. 20 E., Golden 

Valley County (see Easton, 1962). Heath Formation. Zone V. 

Amplexizaphrentis, Siphonophyllia, Turbinatocaninia?.

M-15. Hopley Creek, sees. 5, 8, T. 10 N., R. 13 E., Wheatland 

County (see Gardner and others, 1946). Heath Formation. 

Zone V. Siphonophyllia, Turbinatocaninia?.

M-16. Moss Agate, sec. 6, T. 7 N., R. 8 E., Meagher County (see 

Gardner and others, 1946). Otter Formation. Zone V. 

Siphonophyllia.

M-17. Delpine, sec. 14, T. 9 N., R. 10 E., Meagher County (see 

Easton, 1962). Otter Formation. Zones IV, V. Amplexizaphrentis, Siphonophyllia, Clisiophyllum?.
LOCALITY REGISTER 35

M-18. Potter Creek, sees. 5, 8, T. 13 N., R. 21 E., Fergus County 

(see Easton, 1962). Otter Formation?. Zone V?. Amplexizaphrentis.

M-19. Logan, sec. 25, T. 2 N., R. 2 E., Gallatin County (see Sando 

and Dutro, 1974). Lodgepole and Mission Canyon 

Limestones. Zone IB, 1C, II, IIIA. Amplexizaphrentis, Amplexocarinia, Amplexus, Cyathaxonia, "Diphyphyllum," 

Guerichophyllum, Lophophyllum?, Metriophyllum, Michelinia, Neaxon?, Stelechophyllum, Sychnoelasma, Syringopora, 

Vesiculophyllum.

M-20. Lombard, sec. 7, T. 4 N., R. 3 E., Broadwater County 

(J. T. Dutro, Jr., unpublished data, 1957). Big Snowy Formation. Zone V. Siphonophyllia.

M-21. Sacajawea Peak, sec. 27, T. 2 N., R. 6 E., Gallatin County 

(W. J. Sando, unpublished data, 1962). Lodgepole and Mission Canyon Limestones. Zones IB, 1C, II, IIIA. Amplexizaphrentis, Amplexus, Cyathaxonia, "Diphyphyllum," 

Lophophyllum?, Metriophyllum, Michelinia, Neaxon?, Palaeacis, Stelechophyllum, Sychnoelasma, Syringopora, Vesiculophyllum.

M-22. South Angler Lake, sec. 15, T. 2 N., R. 6 E., Gallatin County (W. J. Sando, unpublished data, 1962). Big Snowy Formation. Zone V. Multithecopora, Siphonophyllia.

M-23. Baldy Mountain, sees. 26, 27, 35, T. 7 S., R. 3 W., Madison 

County (see Sando and Dutro, 1980). Lodgepole Limestone, 

Mission Canyon Limestone, and Big Snowy Formation. 

Zones IB, 1C, II, IIIA, V. Amplexizaphrentis, Amplexus, Aulostylus, Cyathaxonia, "Diphyphyllum," Guerichophyllum, 

Michelinia, Multithecopora, Neaxon?, Palaeacis, Siphonophyllia, Stelechophyllum, Sychnoelasma, Syringopora, Vesiculophyllum, Zaphriphyllum.

M-24. White Peak, sees. 1, 2, T. 11 S., R. 4 E., Gallatin County (see 

Witkind, 1969). Lodgepole and Mission Canyon Limestones. 

Zones IB, 1C, II. Amplexizaphrentis, Amplexus, Cleistopora, 

Cyathaxonia, Lophophyllum?, Metriophyllum, Michelinia, 

Stelechophyllum, Sychnoelasma, Syringopora, Vesiculophyllum.

M-25. Squaw Creek, sec. 34, T. 4 S., R. 4 E., Gallatin County 

(W. J. Sando, unpublished data, 1962). Lodgepole and Mission Canyon Limestones. Zones IB, 1C, II, IIIA. Amplexizaphrentis, Amplexocarinia, Cyathaxonia, Guerichophyllum, 

Lophophyllum?, Michelinia, Pleurosiphonella, Stelechophyllum, Sychnoelasma, Syringopora, Vesiculophyllum.

M-26. Livingston, sees. 1, 2, T. 3 S., R. 9 E., and sec. 35, T. 2 S., 

R. 9 E., Park County (see Sando, 1972b). Lodgepole and Mission Canyon Limestones. Zones IB, 1C, II. Amplexizaphrentis, Amplexus, Caninia, Cyathaxonia, Guerichophyllum, 

Lophophyllum?, Neaxon?, Rotiphyllum?, Sychnoelasma, Syringopora, Vesiculophyllum.

M-27. Baker Mountain, sees. 34, 35, T. 3 S., R. 12 E., and sec. 29, 

T. 3 S., R. 13 E., Park and Sweet Grass Counties (see Sando, 1972b). Lodgepole and Mission Canyon Limestones. 

Zones IB, 1C, II, III. Amplexizaphrentis, Caninia?, "Diphyphyllum," Lophophyllum?, Palaeacis, Stelechophyllum, Sychnoelasma, Syringopora, Vesiculophyllum.

M-28. Picket Pin Ridge, sec. 33, T. 4 S., R. 14 E., Sweet Grass County (see Sando, 1972b). Lodgepole Limestone. Zone 1C. 

Amplexizaphrentis, Cyathaxonia, Cleistopora, Guerichophyllum, Lophophyllum?, Michelinia, Syringopora, Vesiculophyllum.

M-29. West Stillwater River, sees. 1, 2, T. 5 S., R. 14 E., Sweet 

Grass County (see Sando, 1972b). Madison Limestone.

Zones 1C, II. Amplexizaphrentis, Amplexus, Cleistopora, Cyathaxonia, Lophophyllum?, Sychnoelasma, Syringopora, Vesiculophyllum.

M-30. Ben Bow Mine Road, sees. 20,29, T. 5 S., R. 16 E., Stillwater 

County (see Sando, 1972b). Madison Limestone. Zones 1C, 

II, IIIA. Amplexizaphrentis, Cyathaxonia, "Diphyphyllum," 

Lophophyllum?, Michelinia, Syringopora, Vesiculophyllum.

M-31. West Red Lodge Creek, sec. 10, T. 7 S., R. 18 E., Carbon 

County (see Sando, 1972b). Madison Limestone. Zone II. 

Lophophyllum?, Michelinia, Stelechophyllum, Sychnoelasma, 

Syringopora, Vesiculophyllum.

M-32. Point of Rocks, sec. 17, T. 8 S., R. 20 E., Carbon County (see 

Sando, 1972b). Madison Limestone. Zone II. Michelinia, Vesiculophyllum.

M-33. South Grove Creek, sec. 35, T. 8 S., R. 20 E., Carbon County (see Sando, 1972b). Madison Limestone. Zones 1C, II. Syringopora, Vesiculophyllum.

M-34. Devils Canyon, sees. 25, 36, T. 9 S., R. 28 E., Big Horn 

County (W. J. Sando, unpublished data, 1966). Madison 

Limestone. Zones 1C, II, IIIA. Lophophyllum?, Stelechophyllum, Sychnoelasma, Syringopora, Vesiculophyllum.

M-35. Little Dryhead Canyon, sec. 23, T. 7 S., R. 27 E., Carbon 

County (W. J. Sando, unpublished data, 1966). Madison 

Limestone. Zone II. Syringopora, Vesiculophyllum.

NORTH DAKOTA

ND-1. Texas Co., Donahue No. 1 well core, sec. 23, T. 154 N., 

R. 100 W., Williams County (see Sando, 1978). Mission Canyon Limestone. Zone II. Amplexizaphrentis?, Cyathaxonia, 

Syringopora.

UTAH

U-l. Old Laketown Canyon, sec. 32, T. 13 N., R. 6 E., Rich County (see Sando, Dutro, and Gere, 1959). Lodgepole Limestone, 

Little Flat Formation, and Monroe Canyon Limestone. 

Zones II, IIIA, HID, IV. Amplexizaphrentis, Amplexus, Canadiphyllum, Clisiophyllum, Dorlodotia?, Ekvasophyllum, Faberophyllum, Michelinia, Rhopalolasma, Rotiphyllum?, 

Pleurosiphonella, Stelechophyllum, Sychnoelasma, Syringopora, Vesiculophyllum.

U-2. Emma Canyon, sec. 17, T. 11 N., R. 8 E., Rich County (see 

Sando, Dutro, and Gere, 1959). Lodgepole Limestone and 

Brazer Dolomite. Zones IB, 1C, II, IIIA. Amplexizaphrentis, 

Amplexocarinia?, Amplexus, Ankhelasma, Cleistopora, Cyathaxonia, Guerichophyllum, Lophophyllum?, Michelinia?, 

Stelechophyllum, Sychnoelasma, Syringopora, Vesiculophyllum, Zaphriphyllum.

U-3. Brazer Canyon, sec. 20, T. 11 N., R. 8 E., Rich County (see 

Sando, Dutro, and Gere, 1959). Lodgepole Limestone and 

Brazer Dolomite. Zones IB, 1C, II, IIIA. Amplexizaphrentis, 

Amplexocarinia, Amplexus, Ankhelasma, Beaumontia, 

Cleistopora, Cyathaxonia, Guerichophyllum, Lophophyllum?, 

Metriophyllum, Michelinia, Neaxon?, Palaeacis, Stelechophyllum, Syringopora, Vesiculophyllum, Zaphriphyllum.

U-4. Rex Peak, sec. 31, T. 11 N., R. 8 E., Rich County (see Sando, Dutro, and Gere, 1959). Lodgepole Limestone and Brazer 

Dolomite. Zones II, IIIA. Amplexizaphrentis, Ankhelasma,
36 CORAL ZONATION, MISSISSIPPIAN SYSTEM, WESTERN INTERIOR PROVINCE

Sychnoelasma, Syringopora, Vesiculophyllum, Zaphriphyllum.

U-5. Ogden Canyon, sec. 17, T. 6 N., R. 1 E., Weber County 

(W. J. Sando, unpublished data, 1961). Lodgepole Limestone, 

Brazer Dolomite, and Deseret Limestone. Zone II. Amplexizaphrentis, Amplexus, Cyathaxonia, Michelinia, Neaxon?, 

Stelechophyllum, Sychnoelasma, Syringopora, Vesiculophyllum.

U-6. Flux, sec. 31, T. 1 S., R. 6 W., Tooele County (see Sandberg 

and Gutschick, 1979). Deseret Limestone and Humbug Formation. Zones II, HID. Acrocyathus (cerioid), Canadiphyllum, 

Cyathaxonia, Ekvasophyllum, Palaeacis, Pleurosiphonella, 

Siphonodendron.

U-7. Ophir Canyon, sec. 22, T. 5 S., R. 4 W., Tooele County 

(R. C. Gutschick and Mackenzie Gordon, Jr., unpublished 

data, 1953 and 1976). Gardison Limestone, Deseret 

Limestone, and Humbug Formation. Zones II, IIIA, IIIB, 

IIIC, HID, IV. Amplexizaphrentis, Amplexus?, 

Canadiphyllum, Cyathaxonia, Ekvasophyllum, Michelinia?, 

Pleurosiphonella, Rhopalolasma, Sychnoelasma, Vesiculophyllum.

U-8. Rock Canyon, sec. 28, T. 6 S., R. 3 E., Utah County (see Sandberg and Gutschick, 1979). Gardison Limestone and Deseret 

Limestone. Zones II, HID. Amplexizaphrentis, Amplexus, 

Canadiphyllum, Cyathaxonia, Rhopalolasma, Rotiphyllum. 

U-9. Twelvemile Pass, sec. 22, T. 8 S., R. 3 W., Tooele County 

(Mackenzie Gordon, Jr., unpublished data, 1962). Deseret 

Limestone and Humbug Formation. Zones HID, IV. Amplexizaphrentis, Canadiphyllum, Clisiophyllum?, Ekvasophyllum, Faberophyllum, Pleurosiphonella, "Pseudodorlodotia," 

Rotiphyllum?, Syringopora?.

U-10. Gardison Ridge, sec. 1, T. 9 S., R. 3 W., Utah County 

(Mackenzie Gordon, Jr., unpublished data, 1953). Gardison 

Limestone. Zone II. Amplexizaphrentis, Amplexus, Cyathaxonia, Lophophyllum?, Michelinia, Neaxon?, Stelechophyllum, 

Sychnoelasma, Syringopora, Vesiculophyllum.

U-ll. Buckhorn Canyon, sees. 1, 2, T. 10 S., R. 12 W., Tooele 

County (see Gutschick, 1976). Joana Limestone and Woodman Formation. Zones I, II, III. Amplexizaphrentis, Amplexus, Cyathaxonia, Lophophyllum?, Neaxon?, Vesiculophyllum.

U-12. Mammoth Peak-Sioux Creek, sees. 19, 30, T. 10 S., R. 2 W., 

Tooele County (R. C. Gutschick, unpublished data, 1976). 

Gardison Limestone and Deseret Limestone. Zones II, IIIA, 

IIIC, IIID. Amplexizaphrentis, Canadiphyllum, Cyathaxonia, 

Guerichophyllum, Hapsiphyllum, Michelinia, Rhopalolasma?, Rotiphyllum, Vesiculophyllum?.

U-13. North Gillson Mountains, sec. 33, T. 13 S., R. 3 W., Juab 

County (R. C. Gutschick, unpublished data, 1976). Gardison 

Limestone, Deseret Limestone, and Humbug Formation. 

Zones II, IDA. Amplexizaphrentis, Amplexus, Aulostylus, Cyathaxonia, Guerichophyllum, Rhopalolasma, Rotiphyllum, 

Stelechophyllum, Sychnoelasma, Syringopora, Vesiculophyllum.

U-14. Dog Valley Mountain, sec. 21, T. 24 S., R. 6 W., Millard 

County (R. C. Gutschick, unpublished data, 1976). Gardison 

Limestone and Deseret Limestone. Zones II, IIIB. Amplexizaphrentis, Amplexus, Cyathaxonia, Michelinia.

U-15. Dog Valley Peak, sec. 17, T. 25 S., R. 6 W., Millard County 

(R. C. Gutschick, unpublished data, 1976). Gardison 

Limestone and Deseret Limestone. Zones II, IIID. 

Acrocyathus (cerioid), Amplexizaphrentis, Amplexus, 

Aulostylus, Canadiphyllum, Cyathaxonia, Ekvasophyllum, 

Hapsiphyllum, Koninckophyllum?, Lophophyllum?, Michelinia, Palaeacis?, Pleurosiphonella, Rotiphyllum, Siphonodendron, Sychnoelasma, Vesiculophyllum.

U-16. Elephant Canyon, sees. 24, 25, T. 28 S., R. 12 W., Beaver 

County (D. W. Lemmon, unpublished data, 1962). Gardison 

Limestone and Deseret Limestone. Zones II, IIID. Amplexizaphrentis, Canadiphyllum, Cyathaxonia, "Diphyphyllum," 

Ekvasophyllum, Multithecopora, Siphonodendron, Sychnoelasma, Syringopora, Vesiculophyllum.

U-17. Shauntie Hills, sec. 13. T. 29 S., R. 13 W., Beaver Co. 

(D. W. Lemmon, unpublished data, 1962). Zones IIID, IV. 

Deseret Limestone, Humbug Formation, and Great Blue 

Limestone. Acrocyathus (cerioid), Amplexizaphrentis, 

Canadiphyllum, Cyathaxonia, Ekvasophyllum, Faberophyllum, Koninckophyllum?, Rhopalolasma, Siphonodendron, 

Syringopora.

U-18. Green Ravine, sec. 1, T. 2 S., R. 4 W., and sec. 6, T. 2 S., 

R. 3 W., Tooele County (see Tooker and Roberts, 1970). Green 

Ravine Formation. Zone VB. Actinocyathus, Amplexizaphrentis, Duncanopora, Multithecopora, Siphonophyllia.

U-19. Fitchville Ridge, sec. 19, T. 10 S., R. 2 W., Juab County 

(R. C. Gutschick, unpublished data, 1977). Fitchville 

Formation. Zone IA. Amplexus, Lophophyllum?, Syringopora, Vesiculophyllum?.

U-20. Broad Canyon, sec. 23, T. 1 N., R. 6 W., Tooele County 

(R. C. Gutschick, unpublished data, 1977). Lodgepole 

Limestone, Deseret Limestone, Great Blue Limestone(?). 

Zones IB, 1C, IIA, IIB, IIID, IV. Amplexizaphrentis, 

Amplexocarinia, Canadiphyllum, Cleistopora, Cyathaxonia, 

Ekvasophyllum, Faberophyllum, Guerichophyllum, Lophophyllum?, Metriophyllum, Michelinia, Palaeacis, Siphonodendron, Siphonophyllia, Sychnoelasma, Syringopora, Vesiculophyllum.

U-21. Silveropolis Hill, sec. 30, T. 5 S., R. 3 W., and sec. 25, T. 5 S., 

R. 4 W., Tooele County (Mackenzie Gordon, Jr., unpublished 

data, 1962). Great Blue Limestone. Zones IV, VA, VB. Amplexizaphrentis, Canadiphyllum, Clisiophyllum, Faberophyllum, Pleurosiphonella, Rotiphyllum, Siphonophyllia.

U-22. Cedar Valley Hills, sees. 29, 32, T. 6 S., R. 1 W., Utah 

County (R. C. Gutschick, unpublished data, 1976, 1977). 

Deseret Limestone. Zone IIID. Canadiphyllum, Ekvasophyllum, Hapsiphyllum, Palaeacis.

U-23. Lakeside Mountains, sees. 6, 7, T. 1 N., R. 8 W., Tooele 

County (R. C. Gutschick, unpublished data, 1977). Lodgepole 

Limestone and Woodman Formation. Zone II. Amplexizaphrentis, Cyathaxonia, Vesiculophyllum.

U-24. Big Canyon, sees. 32, 33, T. 6 S., R. 6 W., Tooele County 

(R. C. Gutschick, unpublished data, 1977). Humbug Formation^). Zone IIID. Ekvasophyllum, Siphonodendron, Stelechophyllum.

U-25. Samak, sec. 25, T. 2 S., R. 6 E., Summit County (see Dockal, 

1980). Lodgepole Limestone and Brazer Dolomite. Zones II, 

IIIA. Amplexizaphrentis, Amplexus, Ankhelasma?, Caninia, 

Cyathaxonia, Lophophyllum?, Michelinia, Rotiphyllum, Siphonodendron, Sychnoelasma, Syringopora, Vesiculophyllum.

U-26. Trail Hollow, sec. 33, T. 2 N., R. 9 W., Wasatch County (see 

Dockal, 1980). Brazer Dolomite. Zone ULA. Amplexizaphrentis, Ankhelasma, Caninia, Siphonodendron, Sychnoelasma, 

Syringopora, Vesiculophyllum, Zaphriphyllum.

U-27. Rhodes Canyon, sec. 3, T. 1 N., R. 9 W., Duchesne County 

(J. A. Dockal, unpublished data, 1976). Doughnut Formation. 

Zone VB. Amplexizaphrentis, Michelinia, Multithecopora, Siphonophyllia.

U-28. Duchesne River, sees. 6,7, T. 1 N., R. 8 W., Duchesne County 

(see Dockal, 1980). Lodgepole Limestone and Brazer Dolo-
LOCALITY REGISTER 37

mite. Zones II, IIIA. Amplexizaphrentis, Lophophyllum?, 

Siphonodendron, Stelechophyllum, Sychnoelasma, Syringopora, Vesiculophyllum.

U-29. Whiterocks, sec. 1, T. 2 N., R. 1 W., and sec. 6, T. 2 N., 

R. 1E., Uintah County (see Dockal, 1980). Lodgepole 

Limestone and Brazer Dolomite. Zone II. Stelechophyllum, 

Sychnoelasma, Syringopora, Vesiculophyllum.

U-30. Little Brush Creek, sec. 24, T. 1 S., R. 21 E., and sec. 30, 

T. 1 S., R. 22 E., Uintah County (see Dockal, 1980). 

Lodgepole Limestone. Zone II. Amplexizaphrentis, Michelinia, Sychnoelasma, Syringopora, Vesiculophyllum.

U-31. Holiday Park, sec. 35, T. 1 N., R. 9 E., Summit County (see 

Dockal, 1980). Lodgepole Limestone. Zone II. Sychnoelasma.

U-32. East Fork of Black's Fork, sec. 24, T. 2 N., R. 13 E., Summit County (see Dockal, 1980). Lodgepole Limestone. Zone II. 

Amplexizaphrentis, Sychnoelasma.

WYOMING

WY-1. Darby Canyon, sees. 14, 15, 23, T. 43 N., R. 118 W., Teton 

County (see Sando and Dutro, 1960). Lodgepole and Mission 

Canyon Limestones. Zones IB, 1C, II. Amplexizaphrentis, 

Amplexus, Cleistopora, Cyathaxonia, Guerichophyllum, Lophophyllum?, Michelinia, Neaxon?, Stelechophyllum, Sychnoelasma, Syringopora, Vesiculophyllum.

WY-2. Hoback Canyon, sees. 2, 3, T. 38 N., R. 115 W., Teton County 

(see Sando, 1977c; Sando, Bamber, and Armstrong, 1975). 

Mission Canyon Limestone and Amsden Formation. 

Zones II, IIIA. Amplexizaphrentis, "Diphyphyllum," Duncanopora, Multithecopora, Rotiphyllum?, Siphonophyllia, 

Sychnoelasma, Syringopora, Turbinatocaninia?, Vesiculophyllum.

WY-3. Haystack Peak, sec. 19, T. 34 N., R. 117 W., Lincoln County 

(see Sando, 1977c; Sando, Bamber, and Armstrong, 1975). 

Lodgepole and Mission Canyon Limestones and Amsden 

Formation. Zones IB, 1C, II, IIIA, VB. Actinocyathus, 

Amplexizaphrentis, Amplexus, Ankhelasma, Canadiphyllum?, Caninia, Cyathaxonia, Duncanopora, Multithecopora, Pleurosiphonella?, Siphonodendron, Siphonophyllia, 

Stelechophyllum, Sychnoelasma, Syringopora, "Thysanophyllum," Turbinatocaninia?, Vesiculophyllum, Zaphriphyllum.

WY-4. Covey Cutoff Trail, sec. 27, T. 34 N., R. 117 W., Lincoln 

County (see Sando, Bamber, and Armstrong, 1975). Amsden 

Formation. Zone VB. Actinocyathus, Duncanopora, 

Siphonophyllia, Turbinatocaninia?.

WY-5. Moffat Trail, sec. 3, T. 33 N., R. 117 W., Lincoln County (see 

Sando, Bamber, and Armstrong, 1975). Amsden Formation. 

Zone VB. Amplexizaphrentis, Duncanopora, Multithecopora, 

Pleurosiphonella, Siphonophyllia, Turbinatocaninia?.

WY-6. South Indian Creek, sec. 14, T. 38 N., R. 118 W., Lincoln 

County (see Sando, Bamber, and Armstrong, 1975). Amsden 

Formation. Zone VB. Duncanopora, Pleurosiphonella, Siphonophyllia, Turbinatocaninia?.

WY-7. Soda Creek, sec. 5, T. 45 N., R. 110 W., Teton County (see 

Sando, Bamber, and Armstrong, 1975). Amsden Formation. 

Zone VA? Amplexus.

WY-8. Horse Creek, sec. 19, T. 43 N., R. 106 W., Fremont County 

(see Sando, 1967, 1977c). Madison Limestone. Zones I, II, 

IIIA. "Diphyphyllum,"Lophophyllum?, Sychnoelasma, Syringopora, Zaphriphyllum.

WY-9. Wiggins Fork, sec. 7, T. 42 N., R. 105 W., Fremont County 

(see Sando, 1967). Madison Limestone. Zones II, IIIA.

"Diphyphyllum," Sychnoelasma, Syringopora, Vesiculophyllum.

WY-10. Warm Spring Canyon, sec. 31, T. 42 N., R. 107 W., Fremont 

County (see Sando, 1967, 1977c). Madison Limestone. 

Zones I, II, IIIA. Amplexizaphrentis, "Diphyphyllum," Siphonodendron, Sychnoelasma, Syringopora, Vesiculophyllum.

WY-11. Dinwoody Canyon, sees. 1,11,12, T. 4 N., R. 6 W., Fremont 

County (see Sando, 1967). Madison Limestone. Zones I, II, 

IIIA. Amplexizaphrentis, "Diphyphyllum," Lophophyllum?, 

Michelinia, Sychnoelasma, Syringopora, Vesiculophyllum, 

Zaphriphyllum.

WY-12. Bull Lake Creek, sees. 2, 3, T. 2 N., R. 4 W., Fremont County 

(see Sando, 1967). Madison Limestone. Zones I, II, IIIA. 

"Diphyphyllum," Sychnoelasma, Syringopora, Vesiculophyllum, Zaphriphyllum.

WY-13. Washakie Reservoir, sec. 18, T. 1 S., R. 2 W., Fremont 

County (see Sando, 1967). Madison Limestone. Zones I, II, 

IIIA. Amplexizaphrentis, "Diphyphyllum," Lophophyllum?, 

Sychnoelasma, Syringopora, Vesiculophyllum, Zaphriphyllum.

WY-14. Sinks Canyon, sec. 18, T. 32 N., R. 100 W., and sec. 24, 

T. 32 N., R. 101 W., Fremont County (see Sando, 1967). 

Madison Limestone. Zones II, IIIA. Amplexizaphrentis, "Diphyphyllum"?, Lophophyllum?, Michelinia, Sychnoelasma, 

Syringopora, Vesiculophyllum.

WY-15. Little Popo Agie, sees. 13 and 24, T. 31 N., R. 100 W., 

Fremont County (see Sando, 1967). Madison Limestone. 

Zone II. Amplexizaphrentis, Sychnoelasma, Syringopora, 

Vesiculophyllum..

WY-16. Sweetwater Canyon, sees. 27, 34, T. 29 N., R. 97 W., 

Fremont County (see Sando, 1967). Madison Limestone. 

Zone II. Syringopora, Vesiculophyllum.

WY-17. Clarks Fork Canyon, sees. 5, 6, 7, T. 56 N., R. 103 W., Park 

County (see Sando, 1975a). Madison Limestone. Zones I, II, 

IIIA. Amplexizaphrentis, "Diphyphyllum," Lophophyllum?, 

Stelechophyllum, Syringopora, Vesiculophyllum.

WY-18. Shoshone Canyon, sees. 4 and 5, T. 52 N., R. 102 W., Park 

County (see Sando, 1975a). Madison Limestone. Zones I, II, 

IIIA. "Diphyphyllum,"Stelechophyllum, Sychnoelasma, Syringopora, Vesiculophyllum.

WY-19. Red Creek, sec. 28, T. 8 N., R. 1 E., Hot Springs County (see 

Sando, 1967). Madison Limestone. Zones I, II, IIIA. 

"Diphyphyllum," Sychnoelasma, Syringopora, Vesiculophyllum.

WY-20. Wind River Canyon, sec. 4, T. 6 N., R. 6 E., Hot Springs 

County (see Sando, 1967). Madison Limestone. Zones I, II, 

IIIA. Syringopora, Vesiculophyllum.

WY-21. Brown Spring Draw, sec. 35, T. 41 N., R. 92 W., Hot Springs 

County (W. J. Sando, unpublished data, 1965). Madison 

Limestone. Zones I, II. Lophophyllum?, Syringopora, Vesiculophyllum.

WY-22. Sheep Mountain, sec. 35, T. 54 N., R. 94 W., and sec. 2, 

T. 53 N., R. 94 W., Big Horn County (W. J. Sando, unpublished data, 1966). Madison Limestone. Zones II, IIIA. 

Sychnoelasma, Syringopora, Vesiculophyllum.

WY-23. Cottonwood Canyon, sees. 33, 34, T. 57 N., R. 93 W., Big 

Horn County (W. J. Sando, unpublished data, 1966). Madison 

Limestone. Zones I, II, IIIA. Amplexizaphrentis, "Diphyphyllum, " Lophophyllum?, Sychnoelasma, Syringopora, Vesiculophyllum.

WY-24. Shell Canyon, sees. 9, 17, T. 53 N., R. 90 W., Big Horn 

County (W. J. Sando, unpublished data, 1966). Madison 

Limestone. Zones I, II, IIIA. Lophophyllum?, Syringopora, 

Vesiculophyllum.
38 CORAL ZONATION, MISSISSIPPIAN SYSTEM, WESTERN INTERIOR PROVINCE

WY-25. Paintrock Creek, sec. 20, T. 50 N., R. 88 W., and sees. 25 and 

26, T. 50 N., R. 89 W., Big Horn County (W. J. Sando, unpublished data, 1966). Madison Limestone. Zones I, IIIA. "Diphyphyllum," Pleurosiphonella, Syringopora, Vesiculophyllum.

WY-26. Tensleep Canyon, sec. 27, T. 48 N., R. 87 W., Washakie 

County (W. J. Sando, unpublished data, 1965). Madison 

Limestone. Zones I, II, IIIA. Amplexus?, "Diphyphyllum," 

Lophophyllum?, Syringopora, Vesiculophyllum.

WY-27. Little Bighorn Canyon, sec. 20, T. 58 N., R. 89 W., Sheridan 

County (see Sando, 1976b). Madison Limestone. Zone II. 

Lophophyllum?, Sychnoelasma, Syringopora, Vesiculophyllum.

WY-28. Little Tongue River, sees. 15, 26, 27, T. 56 N., R. 87 W., 

Sheridan County (see Sando, 1976b). Madison Limestone. 

Zones II, IIIA. "Diphyphyllum," Sychnoelasma, Syringopora, 

Vesiculophyllum.

WY-29. Wolf Creek, sec. 5, T. 55 N., R. 86 W., Sheridan County 

(W. J. Sando, unpublished data, 1966). Madison Limestone. 

Zone IIIA. "Diphyphyllum," Syringopora.

WY-30. Big Goose Creek, sees. 2, 3, T. 54 N., R. 86 W., and sec. 35, 

T. 55 N., R. 86 W., Sheridan County (see Sando, 1976b). 

Madison Limestone. Zones I, II. Lophophyllum?, Sychnoelasma, Syringopora, Vesiculophyllum.

WY-31. South Rock Creek, sec. 25, T. 52 N., R. 84 W., Johnson 

County (see Sando, 1976b). Madison Limestone. Zones I, II. 

Syringopora, Vesiculophyllum.

WY-32. North Crazy Woman Creek, sees. 27, 28, T. 49 N., R. 83 W., 

Johnson County (see Sando, 1976b). Madison Limestone. 

Zones I, II, IIIA. "Diphyphyllum," Syringopora, Vesiculophyllum.

WY-33. Middle Buffalo Creek, sees. 20, 21, T. 40 N., R. 86 W., 

Natrona County (see Sando, 1976b). Madison Limestone. 

Zone II. Sychnoelasma, Syringopora, Vesiculophyllum.

WY-34. Blue Creek, sec. 3, T. 42 N., R. 85 W., Johnson County (see 

Sando, 1976b). Madison Limestone. Zones I, II. 

Sychnoelasma?, Syringopora, Vesiculophyllum.

WY-35. Cottonwood Creek, sees. 27, 34, T. 27 N., R. 88 W., Carbon 

County (M. W. Reynolds, unpublished data, 1968). Madison 

Limestone. Zone II. Syringopora, Vesiculophyllum.

WY-36. Casper Mountain, sees. 8, 9, T. 32 N., R. 79 W., Natrona 

County (W. J. Sando, unpublished data, 1974). Madison 

Limestone. Zone IIIA. Lophophyllum?, Vesiculophyllum.

LOCALITIES NOT REPRESENTED BY MEASURED SECTIONS

ARIZONA

A-9. Grand Wash Cliffs, sec. 2, T. 30 N., R. 16 W., Mohave County (report on shipment 0-60-55). Redwall Limestone. Zone II. 

Amplexizaphrentis, Syringopora, Vesiculophyllum.

A-10. New Water Spring, west of Hualapai Indian Reservation and 

north of Diamond Bar Ranch (report on shipment 0-60-55). 

Redwall Limestone. Zone II. Amplexizaphrentis, Cyathaxonia, Michelinia, Vesiculophyllum.

A-13. Tonto Basin dam site, Gila County (Ransome, 1917). Redwall Limestone. Zone II. Amplexizaphrentis, Caninia, Michelinia, Sychnoelasma, Syringopora, Vesiculophyllum.

A-16. Queen Creek mine, Superior, Florence quadrangle, Final 

County (USGS loc. 9156-PC). Redwall Limestone. Zone I. 

Amplexizaphrentis, Lophophyllum?, Syringopora.

CALIFORNIA

CA-6. Santa Rosa Hills, in T. 17 S., R. 40 E., Inyo County (reports 

on shipments MD-52-15 and MD-52-29). Tin Mountain 

Limestone and Perdido Formation or Lee Flat Limestone. 

Zones II, HID. Acrocyathus, Amplexizaphrentis, Canadiphyllum?, Caninia?, Ekvasophyllum?, Lophophyllum?, Siphonodendron, Syringopora, "Thysanophyllum," Vesiculophyllum. 

CA-7. Lost Burro, northeast corner of Ubehebe Peak quadrangle, 

Inyo County (report on shipment MD-54-15 and 

USGS loc. 24353-PC and 24354-PC). Tin Mountain 

Limestone. Zone II. Caninia?, Cyathaxonia, Lophophyllum?, 

Syringopora.

CA-8. Argus Range, southwest corner of Panamint Butte 

quadrangle, Inyo County (reports on shipments MD-56-1M 

and MD-56-77). Tin Mountain Limestone. Zone II. Amplexizaphrentis, Guerichophyllum, Sychnoelasma, Syringopora, 

Vesiculophyllum.

CA-9. Towne Pass, crest of Panamint Range, Inyo County (reports 

on shipments MD-56-1M and MD-56-77). Tin Mountain 

Limestone. Zone II. Amplexizaphrentis, Amplexus, Caninia, 

Cyathaxonia, Lophophyllum?, Stelechophyllum, Sychnoelasma, Syringopora, Vesiculophyllum.

CA-10. Lake Hill, Panamint Butte quadrangle, Inyo County (reports 

on shipments MD-56-1M, MD-56-77, and MD-57-31). Tin 

Mountain Limestone. Zone II. Amplexizaphrentis, Beaumontia?, Caninia, Cyathaxonia, Lophophyllum?, Stelechophyllum, 

Sychnoelasma, Syringopora, Vesiculophyllum.

CA-11. Tucki Mountain, sec. 24, T. 16 S., R. 45 E., Inyo County 

(reports on shipments GG-58-3D, GG-60-7, and PC-61-2). 

Tin Mountain Limestone. Zone II. Beaumontia, Cyathaxonia, 

Lophophyllum?, Sychnoelasma, Syringopora, Vesiculophyllum.

CA-12. Tucki Mountain, sec. 31, T. 16 S., R. 45 E., Inyo County 

(report on shipment PC-61-2). Perdido Formation. 

Zone HID?. Canadiphyllum?.

CA-13. Quartz Spring area, Panamint Range, Inyo County 

(McAllister, 1952). Tin Mountain Limestone and Perdido Formation. Zones I, II, HID?. Amplexus, Canadiphyllum?, Cyathaxonia, Sychnoelasma, Syringopora, Vesiculophyllum.

CA-14. Rest Spring Gulch, Quartz Spring area, Inyo County 

(Tischler, 1956, and USGS loc. 17668-PC, 17669-PC, 

17675-PC). Perdido Formation. Zone III?. Amplexizaphrentis, Canadiphyllum?, Cyathaxonia, Michelinia, Neaxon?.

CA-15. Unnamed locality, about 2.5 mi N. 70 W. from northeast corner of Panamint Butte quadrangle, Inyo County (report on 

shipment MD-58-26). Tin Mountain Limestone. Zone II?. 

Syringopora.

CA-16. Hill 2997, southeast end of Funeral Mountains, Ryan 

quadrangle, Inyo County (report on shipments LM-64-1 and 

GG-58-3). Tin Mountain Limestone. Zone II?. Cyathaxonia, 

Lophophyllum?, Syringopora.

CA-17. Knob 4756, Ash Meadows quadrangle, Inyo County (report 

on shipment GG-58-3). Tin Mountain Limestone. Zone II?. 

Amplexus?, Cyathaxonia, Vesiculophyllum.

CA-18. Lemoigne Canyon, 1.5 mi N. 38 E. of Lemoigne mine, Inyo 

County (report on shipment MD-58-26). Tin Mountain 

Limestone. Zone II?. Caninia, Syringopora.
LOCALITY REGISTER 39

COLORADO

C-l. Tincup, Mt. Harvard quadrangle, Gunnison County (report 

on shipment SR-63-3). Leadville Limestone. Zone II. 

Lophophyllum?, Sychnoelasma, Syringopora, Vesiculophyllum.

C-2. Rico, Rico District, Rico quadrangle, Dolores County (report 

on shipment MD-57-41). Leadville Limestone. Zone II. 

Sychnoelasma, Vesiculophyllum.

C-3. Durango, 12.5 mi north of town at switchback in Highway 

550, La Plata County (see Sando, 1972a). Leadville 

Limestone. Zone II. Amplexizaphrentis?, Sychnoelasma, Vesiculophyllum.

C-4. Cement Creek, Crested Butte quadrangle, Gunnison County (see Girty, 1903). Leadville Limestone. Zone II. 

Sychnoelasma, Syringopora, Vesiculophyllum.

C-5. Monarch, east side of trail to town, below office, Chaffee 

County (see Girty, 1903). Leadville Limestone. Zone II?. 

Syringopora.

C-6. Unnamed locality, sec. 18, T. 10 S., R. 84 W., Pitkin County (USGS Iocs. 8464-PC, 19484-PC). Leadville Limestone. 

Zone II. Sychnoelasma, Vesiculophyllum.

C-7. Unnamed locality, sec. 28, T. 7 S., R. 83 W., Eagle County 

(USGS loc. 8458-PC). Leadville Limestone. Zone II. 

Sychnoelasma, Syringopora, Vesiculophyllum.

C-8. Express Creek, sec. 32, T. 11 S., R. 84 W., Pitkin County 

(USGS Iocs. 8460-PC, 8462-PC, 8463-PC, 8466-PC, 

8468-PC, 8474-PC). Leadville Limestone. Zone II. 

Siphonodendron?, Sychnoelasma, Syringopora, Vesiculophyllum.

C-9. Lime Creek, at mouth of creek where it meets Frying Pan 

River, Mt. Jackson quadrangle, Pitkin County 

(USGS loc. 8470-PC). Leadville Limestone. Zone II. Vesiculophyllum.

C-10. Meredith quarry, sec. 14, T. 8 S., R. 83 W., Pitkin County 

(USGS loc. 8473-PC). Leadville Limestone. Zone II. 

Sychnoelasma, Vesiculophyllum.

C-ll. Duncan Hill, Garfield quadrangle, Gunnison County 

(USGS Iocs. 19339-PC, 19489-PC, 19490-PC). Leadville 

Limestone. Zone II. Amplexizaphrentis, Sychnoelasma, Syringopora, Vesiculophyllum.

C-12. Cascade Creek, about 0.5 mi below bridge on Silverton Road, 

San Juan County (see Sando, 1972a). Leadville Limestone. 

Zone II. Sychnoelasma, Vesiculophyllum.

C-13. Coalbank Hill, Engineer Mountain quadrangle, San Juan 

County (see Sando, 1972a). Leadville Limestone. Zone II. 

Sychnoelasma.

C-14. Canyon Creek, Needle Mountains quadrangle, La Plata 

County (see Sando, 1972a). Leadville Limestone. Zone II. 

Sychnoelasma.

C-15. Rockwood, near town, Durango quadrangle, La Plata County 

(see Sando, 1972a). Leadville Limestone. Zone II. Amplexizaphrentis, Sychnoelasma, Vesiculophyllum. 

C-16. Uncompaghre River, 4,000 ft southeast of Canyon Creek, 

Ouray County (Sando, 1972a). Leadville Limestone. Zone II. 

Sychnoelasma, Syringopora, Vesiculophyllum. 

C-17. Silverton, east side of wagon road, 2.5 mi south of town, 

San Juan County. Leadville Limestone. Zone II. 

Vesiculophyllum.

C-18. Irish Canyon, sec. 27, T. 10 N., R. 101 W., Moffat County 

(USGS Iocs. 5117-PC, 5117a-PC, 5117b-PC). Madison 

Limestone. Zone II. Amplexizaphrentis, Cyathaxonia?, 

Sychnoelasma, Syringopora, Vesiculophyllum.

IDAHO

1-1. Unnamed locality, sec. 31, T. 4 N., R. 27 E., Butte County 

(report on shipment CEG-77-3). Unnamed unit above Surrett Canyon Formation. Zone VI?. Amplexizaphrentis. 

1-2. Unnamed locality, sec. 33, T. 6 N., R. 27 E., Butte County 

(report on shipment CEG-75-18). Scott Peak and Surrett 

Canyon Formations. Zones IV, V. Amplexizaphrentis. 

1-3. Unnamed locality, sees. 5, 8,9, T. 5 N., R. 28 E., and sees 32, 

33, T. 6 N., R. 28 E., Butte County (report on shipment 

CEG-76-1). Scott Peak and Surrett Canyon Formations. 

Zones IV, V. Amplexizaphrentis, Faberophyllum, Multithecopora, "Pseudodorlodotia", Siphonodendron, Siphonophyllia.

1-4. Unnamed locality, sec. 5, T. 5 N., R. 27 E., and sec. 6, 

T. 6 N., R. 27 E., Butte County (report on shipment 

CEG-76-1). Scott Peak and Surrett Canyon Formations. 

Zones IV, V. Amplexizaphrentis, Faberophyllum, 

Siphonophyllia.

1-5. Unnamed locality, sec. 11, T. 7 N., R. 29 E., Butte County 

(report on shipment CEG-76-1). Scott Peak Formation. 

Zone IV. Faberophyllum.

1-6. Unnamed locality, sec. 19, T. 7 N., R. 29 E., Butte County 

(report on shipment CEG-76-1). Scott Peak and Surrett Canyon Formations. Zones IV, V., Faberophyllum, 

"Pseudodorlodotia," Siphonophyllia.

1-7. Unnamed locality, sec. 7, T. 7 N., R. 29 E., Butte County 

(report on shipment CEG-76-1). Scott Peak Formation. 

Zone HID. Ekvasophyllum.

1-8. Unnamed locality, sees. 20, 29, T. 5 N., R. 24 E., Custer 

County (reports on shipments CEG-73-7 and CEG-73-13). 

Scott Peak and Surrett Canyon Formations. Zones IV, V. 

Duncanopora, Pleurosiphonella, Schoenophyllum, 

Siphonophyllia.

1-9. Unnamed locality, sec. 3, T. 11 N., R. 32 E., Clark County 

(report on shipment CEG-74-7). McGowan Creek Formation. Zone II?. Rotiphyllum.

1-10. Unnamed locality, sec. 30, T. 9 N., R. 32 E., Clark County 

(report on shipment CEG-74-7). Scott Peak Formation. 

Zone IV. Faberophyllum, Siphonodendron.

1-11. Unnamed locality, sec. 11, T. 11 N., R. 32 E., Clark County (report on shipment CEG-74-7). Scott Peak Formation. 

Zones HID, IV. Ekvasophyllum, Faberophyllum, 

Pleurosiphonella.

1-12. Unnamed locality, sees. 5, 6, 7, T. 11 N., R. 32 E., Clark 

County (report on shipment CEG-74-7). Scott Peak Formation. Zones HID, IV. Canadiphyllum, Ekvasophyllum, 

Faberophyllum, Pleurosiphonella.

1-13. Unnamed locality, sec. 12, T. 13 N., R. 32 E., Clark County 

(report on shipment CEG-74-7). Scott Peak Formation. 

Zone IV. Clisiophyllum, Faberophyllum.

1-14. Unnamed locality, sec. 25, T. 10 N., R. 31 E., Clark County 

(report on shipment CEG-74-7). Surrett Canyon Formation. 

Zone V. Michelinia?.

1-15. Unnamed locality, sees. 3,11, T. 4 N., R. 24 E., Butte County 

(report on shipment CEG-73-13). Scott Peak and Surrett 

Canyon Formations. Zones IV, V. Faberophyllum, 

Siphonophyllia.

1-16. Unnamed locality, sec. 7, T. 3 N., R. 24 E., Butte County 

(report on shipment CEG-73-13). Scott Peak Formation?. 

Zone IV. Amplexizaphrentis?.

1-17. Unnamed locality, sec. 5, T. 2 N., R. 24 E., Butte County 

(report on shipment CEG-73-13). Surrett Canyon Formation. Zone V. Siphonophyllia.
40 CORAL ZONATION, MISSISSIPPIAN SYSTEM, WESTERN INTERIOR PROVINCE

1-18. Unnamed locality, sec. 7, T. 4 S., R. 37 E., Bingham County (report on shipment NR-66-6). Monroe Canyon 

Limestone?. Zone IV. Faberophyllum.

1-19. Unnamed locality, sec. 30, T. 4 S., R. 37 E., Bingham County 

(report on shipment NR-66-6). Monroe Canyon Limestone?. 

Zone IV. Faberophyllum.

1-20. Bannock Peak, sees. 27, 28, 33, 34, T. 9 S., R. 35 E., Power 

County (report on shipment EG-58-9). Lodgepole Limestone. 

Zone II. Amplexizaphrentis, Michelinia, Palaeacis, 

Sychnoelasma, Vesiculophyllum.

1-21. Unnamed locality, sec. 5, T. 9 S., R. 33 E., Power County 

(report on shipment EG-58-2). Lodgepole Limestone. 

Zone II. Sychnoelasma, Vesiculophyllum.

1-22. Unnamed locality, sec. 21, T. 5 S., R. 35 E., Power County 

(report on shipment EG-58-2). Lodgepole Limestone. 

Zone II. Amplexizaphrentis?, Sychnoelasma.

1-23. Unnamed locality, sec. 36, T. 8 S., R. 33 E., Power County 

(report on shipment EG-58-2). Great Blue Limestone. 

Zone IV. Faberophyllum.

1-24. Unnamed locality, sees. 25, 30, 36, T. 17 N., R. 26 E., Lemhi 

County (report on shipments NR-64-6 and NR-61-24). Scott 

Peak Formation?. Zones IV, V. Dorlodotia?, Ekvasophyllum?, 

Faberophyllum, Pleurosiphonella, Schoenophyllum.

1-25. Broken Wagon Creek, sec. 25, T. 11 N., R. 20 E., Custer 

County (report on shipment CEG-74-2). Scott Peak Formation?. Zone IV. Faberophyllum, "Pseudodorlodotia,"

1-26. Unnamed locality, sec. 15, T. 6 N., R. 30 E., Butte County 

(report on shipment WR-52-18). Scott Peak Formation?. 

Zone HID. "Diphyphyllum."

1-27. Unnamed locality, sec. 9, T. 12 N., R. 19 E., Custer County 

(report on shipment CEG-72-13D). Unknown formation. 

Zone indeterminate. Caninia, Siphonodendron.

1-28. Peck's Canyon, sees. 19, 30, T. 9 N., R. 21 E., Custer County (report on shipments 0-72-30 and 0-72-82). Scott Peak(?) 

and Surrett Canyon(?) Formations. Zones IV, V. Amplexizaphrentis, Faberophyllum, Multithecopora, Schoenophyllum, Siphonodendron, Siphonophyllia, "Thysanophyllum."

1-29. Unnamed locality, sec. 22, T. 11 N., R. 16 E., Custer County (report on shipment RME-71-1). McGowan Creek Formation?. Zone indeterminate. Amplexizaphrentis?.

1-30. Unnamed locality, sec. 13, T. 16 N., R. 43 E., Fremont County (report on shipment NR-67-3). Amsden Formation?. Zone 

indeterminate. A mplexizaphrentis.

1-31. Montpelier, sec. 15, T. 13 S., R. 44 E., Bear Lake County 

(report on shipment NR-65-6D). Little Flat Formation?. 

Zone HID. Canadiphyllum, Ekvasophyllum?.

1-32. Unnamed locality, sees. 26, 32, 35, 36, T. 9 S., R. 39 E., Bannock and Caribou Counties (report on shipment NR-62-9). 

Lodgepole Limestone, Little Flat Formation, and Monroe 

Canyon Limestone. Zones II, HID, IV. Acrocyathus (cerioid), 

Amplexizaphrentis, Dorlodotia?, Ekvasophyllum, Faberophyllum, Palaeacis, Petalaxis, Sychnoelasma, Syringopora, Vesiculophyllum.

1-33. Unnamed locality, sees. 7,12,13, T. 16 N., R. 27 E., Lemhi 

County (report on shipment MD-60-13). Lodgepole 

Limestone and Scott Peak Formation?. Zones II, IV. Amplexizaphrentis, Faberophyllum, Lophophyllum?, Sychnoelasma, 

Syringopora, Vesiculophyllum.

1-34. Unnamed locality, 43°35' N., Ill °20' W., Bonneville County 

(report on shipment CD-60-6) Lodgepole and Mission Canyon Limestones. Zone II. Stelechophyllum, Sychnoelasma, 

Syringopora, Vesiculophyllum.

1-35. Unnamed locality, sec. 20, T. 7 S., R. 44 E., Caribou County (report on shipment CD-59-2). Monroe Canyon

Limestone?. Zone IV. Faberophyllum, Multithecopora, 

Pleurosiphonella, "Pseudodorlodotia,"

1-36. Unnamed locality, sees. 16, 17, 18, 29, T. 8 S., R. 41 E., 

Caribou County (report on shipment MD-57-44). Lodgepole 

and Monroe Canyon Limestones. Zones II, IV. 

Faberophyllum, Michelinia, Pleurosiphonella, Siphonodendron, Stelechophyllum, Sychnoelasma, Syringopora, 

Vesiculophyllum.

1-37. Unnamed locality, sec. 11, T. 8 S., R. 40 E., Bannock County (report on shipment MD-57-44). Lodgepole Limestone. 

Zone II. Sychnoelasma.

1-38. Dry Ridge, sec. 36, T. 9 S., R. 44 E., Caribou County (report 

on shipment MD-52-24). Monroe Canyon Limestone. 

Zones HID, IV. Clisiophyllum, "Diphyphyllum," 

Ekvasophyllum, Faberophyllum, Multithecopora.

1-39. Unnamed locality, sec. 22, T. 8 N., R. 26 E., Butte County 

(report on shipment NR-64-14). Scott Peak Formation. 

Zone HID. Acrocyathus, Pleurosiphonella, Siphonodendron.

1-40. Unnamed locality, sees. 2, 3, 11, 12, T. 3 S., R. 39 E., 

Bingham County (Sando, 1965). Monroe Canyon Limestone. 

Zone HID?. Sciophyllum.

M-l.

M-2.

M-3.

M-4.

M-5.

M-6.

M-7.

M-8.

M-9.

MONTANA

Unnamed locality, sec. 36, T. 1 S., R. 10 W., Beaverhead 

County (report on shipment EGM-75-1). Lodgepole 

Limestone. Zone II. Sychnoelasma, Vesiculophyllum. 

Little Blackfoot River, sec. 29, T. 11 N., R. 6 W., Lewis and 

Clark County (report on shipment 0-63-66). Madison 

Limestone?. Zone IIIA. Ankhelasma, Vesiculophyllum. 

Canyon Creek (report on shipment EGM-73-1). Mission 

Canyon Limestone. Zone II. Lophophyllum?, Sychnoelasma, 

Syringopora, Vesiculophyllum.

Stockette, 0.25 mi north of town, Cascade County (report on 

shipment 0-66-27). Mission Canyon Limestone. Zone IIIA?. 

Vesiculophyllum.

Unnamed locality, sec. 12, T. 15 N., R. 9 E., Judith Basin 

County (report on shipment NR-65-2). Lodgepole Limestone. 

Zone II. Amplexizaphrentis, Caninia, Lophophyllum?, 

Sychnoelasma, Stelechophyllum.

Unnamed locality, sees. 7, 17, T. 15 N., R. 10 E., Judith 

Basin County (report on shipment NR-65-2). Lodgepole 

Limestone. Zone II. Amplexizaphrentis, Cyathaxonia, Lophophyllum?, Stelechophyllum, Sychnoelasma, Syringopora, Vesiculophyllum.

Unnamed locality, sees. 22, 27, T. 15 N., R. 9 E., Judith 

Basin County (report on shipment NR-65-2). Lodgepole 

Limestone. Zone II. Amplexizaphrentis, Amplexus, Cyathaxonia, Lophophyllum?, Sychnoelasma, Syringopora, Vesiculophyllum.

The Arch and Mixes Baldy, sees. 9, 21, 22, 33, T. 16 N., 

R. 9 E., Judith Basin County (report on shipment 

NR-64-15). Lodgepole Limestone. Zone II. Cyathaxonia, 

Michelinia, Stelechophyllum, Sychnoelasma, Syringopora, 

Vesiculophyllum.

Paddy's Run, sees. 29, 36, T. 6 N., R. 5 E., Gallatin and 

Meaghes Counties (report on shipment NR-64-2). Lodgepole 

and Mission Canyon Limestones. Zone II. Amplexus, 

Rotiphyllum?, Sychnoelasma, Syringopora, Vesiculophyllum. 

M-10. Unnamed locality, sec. 17, T. 5 N., R. 6 E., Broadwater County (report on shipment NR-63-3D). Lodgepole Limestone. 

Zone 1C. Amplexizaphrentis, Amplexus, Cyathaxonia, 

Rotiphyllum.
LOCALITY REGISTER 41

M-ll. Wall Mountain, sec. 10, T. 5 N., R. 5 E., Broadwater County (report on shipment NR-63-3D). Lodgepole Limestone. 

Zones 1C, II. Amplexizaphrentis, Amplexus, Caninia, 

Lophophyllum?, Sychnoelasma, Vesiculophyllum.

M-12. Wolf Creek, sec. 23, T. 16 N., R. 5 W., Lewis and Clark County (report on shipment G and P-59-3). Lodgepole Limestone. 

Zone II. Lophophyllum?.

M-13. Swift Reservoir, sec. 29, T. 28 N., R. 10 W., Pondera County. Castle Reef Dolomite?. Zone IIIA. "Diphyphyllum," 

Siphonodendron, Stelechophyllum?, Vesiculophyllum.

M-14. Unnamed locality, sec. 12, T. 2 N., R. 1 E., Gallatin County (report on shipment GG-53-10D). Madison Limestone. 

Zone II. Sychnoelasma.

M-15. Unnamed locality, sec. 36, T. 3 N., R. 1 E., Broadwater County (report on shipment GG-53-10D). Lodgepole Limestone. 

Zone II. Sychnoelasma.

M-16. Shell Mountain, T. 3 S., R. 11 E., Park County (report by 

Helen Duncan to Paul Richards, January 27, 1950). Lodgepole Limestone. Zone II. Lophophyllum?, Stelechophyllum, 

Sychnoelasma, Syringopora, Vesiculophyllum.

M-17. Devils Lane, 1.5 mi southeast of Crockett Lake, Varney 

quadrangle, Madison County (same report as M-16). Mission Canyon Limestone. Zone IIIA. Amplexizaphrentis, 

"Diphyphyllum," Syringopora, Vesiculophyllum.

M-18. Beaver Creek, sec. 34, T. 21 N., R. 9 W., Lewis and Clark 

County (report on shipment EG-58-10). Allan Mountain 

Limestone. Zone 1C. Cyathaxonia, Guerichophyllum, 

Neaxon?.

NEVADA

N-l. San Antonio Mountains, sec. 32, T. 6 N., R. 42 E., Nye County (report on shipment CMR-76-1). Unnamed formation. 

Zone HID?. "Diphyphyllum," Siphonodendron, Syringopora.

N-2. Gates Mill, sec. 8, T. 3 N., R. 36 E., Esmeralda County 

(reports on shipments RMM-72-3, CMR-74-3, CMR-74-5, 

and CMR-76-1). Unnamed formation. Zone HID. Amplexizaphrentis, Canadiphyllum, Caninia, "Diphyphyllum," 

Ekvasophyllum, Hapsiphyllum?, Koninckophyllum?, 

Sciophyllum, Siphonodendron.

N-3. Tierney Creek, sec. 28, T. 14 N., R. 42 E., Nye County 

(reports on shipments CMR-74-5 and CMR-76-1). 

Unnamed formation. Zone IV. Faberophyllum, Syringopora.

N-4. Rays townsite, sec. 14, T. 4 N., R. 42 E., Nye County (reports 

on shipments RMM-69-9 and CMR-76-1). Unnamed formation. Zones HID, IV. Acrocyathus (cerioid), Canadiphyllum, Caninia, Faberophyllum, Koninckophyllum?, Multithecopora, Pleurosiphonella, Siphonodendron, Syringopora.

N-6. Unnamed locality, sec. 35, T. 30 N., R. 66 E., Elko County 

(report on shipment PM-71-16). Unnamed formation. 

Zone II. Stelechophyllum, Sychnoelasma, Syringopora, 

Vesiculophyllum.

N-8. Kane Springs Wash, sec. 4, T. 10 S., R. 65 E., Lincoln County 

(report on shipment 0-71-37). Joana Limestone. Zone II. 

Sychnoelasma, Vesiculophyllum.

N-9. Unnamed locality, 36°41'10"N., 115°52'25"W., Mercury 

quadrangle, Clark County (report on shipment SP-65-1). 

"Trinity Limestone." Zone III. Palaeacis, Rhopalolasma?. 

N-10. Indian Springs, northwest corner of T. 16 S., R. 55% E., 

Clark County (report on shipment SP-65-7). Monte Cristo 

Limestone. Zones II, III. Amplexizaphrentis, Amplexus, 

Canadiphyllum, Caninia, Cyathaxonia, Stelechophyllum, 

Sychnoelasma, Syringopora, Vesiculophyllum.

N-ll. Indian Springs, sec. 31, T. 16 S., R. 56 E., Clark County 

(report on shipment SP-65-7). Bird Spring Formation. 

Zone V. Amplexizaphrentis.

N-12. Enright Hill, sec. 8, T. 46 N., R. 55 E., Elko County (report 

on shipment SW-62-23). Banner Formation. Zone HID. 

Acrocyathus (cerioid), Canadiphyllum, Multithecopora, 

Siphonodendron.

N-13. Unnamed locality, on road from Newark Summit to Pinto 

Creek Ranch, Pinto Summit quadrangle, White Pine County 

(report on shipment SW-69-11). Diamond Peak Formation. 

Zone V. Amplexizaphrentis?, Michelinia.

N-14. Diamond Peak, southwest shoulder, Eureka quadrangle, 

Eureka County (report on shipment SW-69-11). Diamond 

Peak Formation. Zone V. Amplexizaphrentis.

N-15. Spring Canyon, sec. 11, T. 32 N., R. 53 E., Elko County 

(report on shipment SW-67-1). Diamond Peak Formation. 

Zone IV. Faberophyllum.

N-16. Dixie Creek, sec. 24, T. 32 N., R. 54, Elko County (report on 

shipment SW-67-1). Diamond Peak Formation. Zone V. 

Amplexizaphrentis.

N-17. Unnamed locality, sees. 9,11, T. 45 N., R. 53 E., Elko County 

(reports on shipments MD-57-38 and SW-67-2). Banner 

Formation. Zone HID. Canadiphyllum, Dorlodotia?, 

Ekvasophyllum?, Siphonodendron.

N-18. Black Point, sec. 12, T. 16 N., R. 54 E., White Pine County 

(report on shipment SW-68-2). Joana Limestone. Zone I?. 

Cyathaxonia?.

N-19. Pancake Range, sec. 26, T. 17 N., R. 55 E., White Pine County (reports on shipments SW-66-20 and SW-68-2). Joana 

Limestone. Zone I. Cyathaxonia, Lophophyllum?, Michelinia, 

Syringopora.

N-20. Secret Canyon, sec. 31, T. 18 N., R. 54 E., Eureka County 

(report on shipment SW-68-2). Diamond Peak Formation. 

Zone V. Faberophyllum.

N-21. Pogues Hill, sec. 17, T. 15 N., R. 55 E., White Pine County 

(reports on shipments SW-69-12 and SW-61-40). Chainman 

Shale. Zone HID?. Amplexizaphrentis, Ekvasophyllum?, 

Rhopalolasma.

N-22. Schell Creek Range, sec. 16, T. 11 N., R. 65 E., White Pine 

County (report on shipment SW-68-1). Joana Limestone. 

Zone II. Michelinia, Vesiculophyllum.

N-23. Grant Range, sec. 33, T. 9 N., R. 50 E., Nye County (report 

on shipment SW-65-13). Joana Limestone. Zone II. 

Sychnoelasma, Vesiculophyllum.

N-24. Unnamed locality, sees. 25, 36, T. 20 S., R. 62 E., Clark 

County (report on shipments HM-66-4 and SP-66-1). Unnamed formation. Zone 1C or II. Amplexizaphrentis, Cyathaxonia, Lophophyllum?, Michelinia, Rotiphyllum, Vesiculophyllum.

N-25. Mountain Springs, sec. 19, T. 22 S., R. 58 E., Clark County 

(report on shipment SP-66-1). Monte Cristo Limestone. 

Zone HID?. Acrocyathus (cerioid), Canadiphyllum, 

Dorlodotia?.

N-26. Carlin Canyon, sec. 21, T. 33 N., R. 53 E., Elko County 

(report on shipment GG-58-19). "Tonka Formation." 

Zone II. Amplexizaphrentis, Cyathaxonia.

N-27. Newark Mountain, canyon near southwest edge of mountain, Eureka quadrangle, Eureka County (report on shipment SW-66-20). "White Pine Shale." Zone II?. 

Amplexizaphrentis?.

N-28. Quartzite Mountain, Tippipah Spring NE quadrangle, Nye 

County (report on shipment SP-64-4). Eleana Formation. 

Zone III?. Amplexizaphrentis, Amplexus, Cyathaxonia.
42 CORAL ZONATION, MISSISSIPPIAN SYSTEM, WESTERN INTERIOR PROVINCE

N-29. Grouse Canyon, Tippipah Spring NW quadrangle, Nye 

County (report on shipment SP-64-4). Eleana Formation. 

Zone IV. Cyathaxonia, Faberophyllum.

N-30. Frenchman Flat, Cane Springs NE quadrangle, Nye County (report on shipment SP-64-4). Eleana Formation. Zone V. 

Amplexizaphrentis.

N-31. Narrow Canyon, Mercury NE quadrangle, Nye County 

(report on shipment SP-64-4). Mercury Limestone. Zone II?. 

Lophophyllum?, Syringopora, Vesiculophyllum.

N-32. Unnamed locality, 10 mi north of Tonopah, Nye County 

(report on shipment SW-63-2). Unnamed formation. 

Zone III. Siphonodendron.

N-33. Big Spring, sec. 18, T. 9 N., R. 55 E., Nye County (report 

on shipment SW-63-17). Joana Limestone. Zone II?. 

Cyathaxonia, Syringopora.

N-34. Unnamed locality, sec. 17, T. 6 N., R. 59 E., Nye County 

(reports on shipments SW-62-7 and SW-61-8). Joana 

Limestone. Zone II. Amplexizaphrentis, Amplexus, Cyathaxonia, Lophophyllum?, Vesiculophyllum.

N-35. Diamond Peak, sec. 12, T. 20 W., R. 54 E., Eureka County 

(report on shipment SW-61-10). Diamond Peak Formation. 

Zone III. Amplexizaphrentis, Cyathaxonia, Siphonodendron.

N-36. Seaman Range, north end, Lund quadrangle, Nye County 

(report on shipment SW-63-7). Joana Limestone. Zone II. 

Sychnoelasma, Vesiculophyllum.

N-37. Unnamed locality, sec. 34, T. 4 N., R. 60 E., Nye County 

(report on shipment SW-62-7). Joana Limestone. Zone II. 

Sychnoelasma, Vesiculophyllum.

N-38. Cooper Canyon, sees. 20, 28, T. 15 N., R. 65 E., White Pine 

County (report on shipment SW-61-4). Joana Limestone. 

Zone II. Amplexizaphrentis, Stelechophyllum, Sychnoelasma, 

Vesiculophyllum.

N-39. Unnamed locality, sec. 32, T. 27 N., R. 53 E., Elko County 

(report on shipment GG-56-10). Unnamed formation. 

Zone II?. Syringopora.

N-40. Unnamed locality, sec. 35, T. 19 S., R. 59 E., Clark County 

(report on shipment GG-58-21). Monte Cristo Limestone. 

Zone IIIA. "Diphyphyllum," Syringopora, Vesiculophyllum.

N-41. Unnamed locality, sec. 5, T. 28 N., R. 54 E., Elko County 

(report on shipment GG-57-12). Unnamed formation. 

Zone IV?. Faberophyllum?.

N-42. Unnamed locality, sec. 13, T. 19 S., R. 56 E., Clark County 

(report on shipment 0-55-26). Dawn Limestone [Member of 

Monte Cristo Limestone]. Zone II. Sychnoelasma.

N-43. Deer Creek Basin, sec. 18, T. 19 S., R. 57 E., Clark County 

(report on shipment 0-55-26). Unnamed formation. Zone II. 

Sychnoelasma.

N-44. Pahroc Range, sec. 26, T. 5 N., R. 62 E., Lincoln County 

(report on shipment MD-56-65). Unnamed formation. 

Zone II?. Sychnoelasma?, Vesiculophyllum?.

N-45. South Schell Creek Range, sec. 13, T. 5 N., R. 63 E., Lincoln 

County (report on shipment MD-56-65). Unnamed formation. Zone II. Stelechophyllum, Sychnoelasma, Vesiculophyllum.

N-46. Unnamed locality, sees. 11, 22, 27, 28, T. 38 N., R. 41 E., 

Humboldt County (reports on shipments MD-52-18, 52-23, 

50-13, and 51-26). Goughs Canyon Formation. Zone III. 

Amplexizaphrentis, Canadiphyllum, Cyathaxonia, Lophophyllum?, Michelinia, Neaxon?, Rhopalolasma, Siphonodendron, Syringopora.

N-47. Dutch John Mountain, sees. 3,10, T. 6 N., R. 65 E., Lincoln 

County (see Stensaas and Langenheim, 1960). Bristol Pass 

Limestone. Zone II. Stelechophyllum, Vesiculophyllum.

N-48. 

N-49. 

N-50. 

N-51. 

N-52. 

N-53. 

N-54.

N-55. 

N-56.

N-57. 

N-58.

N-59.

N-60. 

N-61.

N-62.

Unnamed locality, sec. 26, T. 15 N., R. 62 E., White Pine 

County (see Stensaas and Langenheim, 1960). Joana 

Limestone. Zone II. Stelechophyllum, Vesiculophyllum. 

Unnamed locality, sec. 33, T. 10 N., R. 63 E., White Pine 

County (see Stensaas and Langenheim, 1960). Joana 

Limestone. Zone II. Stelechophyllum. 

Unnamed locality, sec. 33, T. 10 N., R. 63 E., White Pine 

County (see Stensaas and Langenheim, 1960). Joana 

Limestone. Zone II. Stelechophyllum, Vesiculophyllum. 

Unnamed locality, sec. 1, T. 14 N., R. 62 E., White Pine 

County (see Stensaas and Langenheim, 1960). Joana 

Limestone. Zone II. Stelechophyllum, Vesiculophyllum. 

Unnamed locality, sees. 16, 21, T. 23 S., R. 58 E., Clark 

County (see Hewett, 1931). Monte Cristo Limestone. Zone II. 

Amplexizaphrentis, Sychnoelasma, Vesiculophyllum. 

Unnamed locality, sec. 34, T. 23 S., R. 57 E., Clark County 

(see Hewett, 1931). Monte Cristo Limestone. Zone II. Amplexizaphrentis, Cyathaxonia, Lophophyllum?, Sychnoelasma. 

Unnamed locality, sec. 1, T. 24 S., R. 57 E., Clark County 

(USGS locality 4293-PC). Monte Cristo Limestone. Zone II?. 

Lophophyllum?, Michelinia, Sychnoelasma, Syringopora, 

Vesiculophyllum.

Unnamed locality, sees. 5, 8, 9, 31, 33, T. 24 S., R. 58 E., 

Clark County (see Hewett, 1931). Monte Cristo Limestone. 

Zone IIIA. Caninia, "Diphyphyllum,"Siphonodendron, Syringopora, Vesiculophyllum.

Unnamed locality, sees. 2,19, 23, 34, T. 25 S., R. 58 E., Clark 

County (see Hewett, 1931). Monte Cristo Limestone. Zones II, 

IIIA. "Diphyphyllum,"Stelechophyllum, Sychnoelasma, Syringopora, Vesiculophyllum.

West Range, Bristol Pass quadrangle, Lincoln County (see 

Westgate and Knopf, 1932). Bristol Pass Limestone. Zone II. 

Amplexus?, Sychnoelasma, Syringopora, Vesiculophyllum. 

Valley Ridge and Green Monster mine, sees. 1, 8, T. 24 S., 

R. 56 E., Clark County (see Hewett, 1956). Monte Cristo 

Limestone. Zones II, IIIA. Amplexizaphrentis, Caninia, 

"Diphyphyllum," Lophophyllum?, Syringopora. 

North slope of Dutch John Mountain, 40 mi northeast of 

Pioche, Lincoln County (see Westgate and Knopf, 1932). 

Bristol Pass Limestone. Zone II. Amplexizaphrentis, Stelechophyllum, Sychnoelasma, Syringopora, Vesiculophyllum. 

Unnamed locality, sec. 18, T. 16 N., R. 63 E., White Pine 

County. Joana Limestone. Zone II. Michelinia. 

Unnamed locality, Owyhee quadrangle, Elko County (report 

on shipment MD-59-25). Banner Formation. Zone HID. 

Canadiphyllum, Siphonodendron.

Unnamed locality, Wheeler Peak quadrangle, White Pine 

County (report on shipment MD-59-3). Joana Limestone. 

Zone II. Cleistopora, Stelechophyllum, Syringopora.

SOUTH DAKOTA

SD-1.

SD-2.

Unnamed locality, ridge northwest of Deadwood, Sturgis 

quadrangle, Lawrence County (USGS loc. 1899-PC). 

Pahasapa Limestone. Zone II. Syringopora. 

Unnamed locality, east side of Whitewood Creek Canyon, 

Sturgis quadrangle, Lawrence County (USGS Iocs. 4325-PC, 

4326-PC). Pahasapa Limestone. Zone II. Syringopora, 

Vesiculophyllum.

SD-3. Unnamed locality, west slope of Crook Mountain, Sturgis 

quadrangle, Lawrence County (USGS loc. 4328-PC). 

Pahasapa Limestone. Zone II. Syringopora.
LOCALITY REGISTER 43

SD-4. Meadow Creek, Sturgis quadrangle, Lawrence County

(USGS Iocs. 4334-PC, 4337-PC). Pahasapa Limestone.

Zone II. Syringopora, Vesiculophyllum. 

SD-5. Runkel, Sturgis quadrangle, Meade County

(USGS Iocs. 4339-PC, 4340-PC, 4347-PC, 4350-PC).

Pahasapa Limestone. Zone II. Syringopora, Vesiculophyllum?. 

SD-6. Pillar Peak, Sturgis quadrangle, Lawrence County

(USGS loc. 4341-PC). Pahasapa Limestone. Zone II.

Syringopora. 

SD-7. Bear Butte Canyon and Butcher Gulch, Sturgis quadrangle,

Lawrence County (USGS Iocs. 4342-PC, 4343-PC).

Pahasapa Limestone. Zone II. Amplexizaphrentis,

Syringopora. 

SD-8. Whiterock, east of Deadwood, Sturgis quadrangle, Lawrence

County (USGS loc. 4344-PC). Pahasapa Limestone. Zone II.

Lophophyllum?. 

SD-9. Tollgate Flats, Spearfish quadrangle, Lawrence County

(USGS loc. 4348-PC). Pahasapa Limestone. Zone II.

Vesiculophyllum. 

SD-10. Terry Peak, Spearfish quadrangle, Lawrence County

(USGS loc. 4349-PC). Pahasapa Limestone. Zone II.

Syringopora. 

SD-11. Unnamed locality, sec. 19, T. 4 N., R. 3 E., Lawrence County

(USGS Iocs. 5704-PC, 5704a-PC, 5704b-PC, 5704d-PC).

Pahasapa Limestone. Zone II. Lophophyllum?, Syringopora,

Vesiculophyllum?.

SD-12. Unnamed locality, sec. 16, T. 1 N., R. 1 E., Pennington County (report on shipment NR-65-6). Pahasapa Limestone.

Zone II. Syringopora, Vesiculophyllum. 

SD-13. Unnamed locality, sec. 33, T. 3 N., R. 1 E., Lawrence County

(report on shipment NR-65-6). Pahasapa Limestone. Zone II.

Vesiculophyllum.

UTAH

U-l. Unnamed locality, sec. 19, T. 9 N., R. 1 E., Cache County 

(report on shipment WMR-76-1M). Deseret Limestone or 

Humbug Formation. Zone HID. Ekvasophyllum, 

S iphonodendron.

U-2. Unnamed locality, sees. 2, 24, 31, 35, 36, T. 9 N., R. 1 W., 

Box Elder County (report on shipment WMR-76-1M). Gardison Limestone and Humbug Formation. Zones II, IV. 

Cleistopora, Cyathaxonia, Faberophyllum, Lophophyllum?, 

Sychnoelasma, Vesiculophyllum.

U-3. Unnamed locality, sees. 28, 29, T. 6 N., R. 9 W., Box Elder 

County (report on shipment 0-75-5). Great Blue Limestone 

and Humbug Formation?. Zone IV. Faberophyllum.

U-4. Unnamed locality, sec. 16, T. 1 N., R. 8 W., Tooele County 

(report on shipment 0-75-5). Great Blue Limestone. Zone IV. 

Faberophyllum.

U-5. Unnamed locality, sees. 4 and 15, T. 3 S., R. 4 E., Wasatch 

County (report on shipments RMM-71-1 and RMM-71-2). 

Gardison Limestone and Brazer Dolomite(?). Zones II, IIIA. 

Amplexizaphrentis, Lophophyllum?, S iphonodendron, 

Sychnoelasma, Syringopora, Vesiculophyllum.

U-6. Unnamed locality, sec. 14, T. 14 S., R. 16 W., Juab County 

(report on shipment WMR-72-4). Joana Limestone. Zone II. 

Stelechophyllum, Sychnoelasma, Syringopora, Vesiculophyllum.

U-7. Unnamed locality, sec. 3, T. 3 S., R. 4 E., Wasatch County 

(report on shipment RMM-71-1). Gardison Limestone. 

Zone II. Syringopora.

U-8. Unnamed locality, sec. 11, T. 2 N., R. 18 E., Daggett County (reports on shipments 0-65-23 and CD-66-5). Madison 

Limestone and Doughnut Formation. Zones II, V. Amplexizaphrentis, Sychnoelasma, Syringopora.

U-9. Unnamed locality, sees. 1, 10, 13, T. 7 N., R. 3 E., Weber 

County (reports on shipments CD-67-1 and CD-69-5). Humbug Formation. Zone IV. Amplexizaphrentis, Faberophyllum, 

Siphonodendron.

LJ-10. Unnamed locality, sees. 34 and 35, T. 13 S., R. 3 W., and 

sec. 2, T. 14 S., R. 3 W., Juab County (report on shipment 

BFM-68-1). Gardison Limestone and Humbug Formation. 

Zones II, HID. Acrocyathus? (cerioid), Canadiphyllum, 

Ekvasophyllum, Pleurosiphonella, Sychnoelasma, Syringopora, Vesiculophyllum.

LJ-11. Beirdneau Peak, sees. 11,14, T. 12 N., R. 2 E., Cache County 

(report on shipment 0-65-23). Lodgepole Limestone and 

"Brazer Limestone." Zones II, HID. Siphonodendron, 

Sychnoelasma.

U-12. Smith and Morehouse Canyon, T. 1 S., R. 7 E., Summit 

County (report on shipment 0-65-23). Madison Limestone. 

Zone II. Amplexizaphrentis, Sychnoelasma.

U-13. Duchesne River, T. 1 N., R. 8 W., Duchesne County (report 

on shipment 0-65-23). Madison Limestone. Zone II. 

Lophophyllum?, Vesiculophyllum.

U-14. Deadman Pass, sec. 33, T. 2 N., R. 11 E., Summit County 

(report on shipment 0-65-23). Madison Limestone. Zone II. 

Sychnoelasma, Vesiculophyllum.

U-15. Promontory Range, Box Elder County (report on shipment 

0-58-32). Madison Limestone, Humbug Formation, and 

Great Blue Limestone. Zones II, HID, IV. Dorlodotia?, 

Ekvasophyllum, Faberophyllum, Lophophyllum?, 

Pleurosiphonella, Sychnoelasma, Vesiculophyllum.

U-16. Unnamed locality, sec. 28, T. 18 S., R. 16 W., Millard County 

(report on shipment F-59-2). Joana Limestone. Zone II. 

Amplexizaphrentis, Stelechophyllum, Syringopora, 

Vesiculophyllum?.

U-17. Unnamed locality, north end of Beaver Lake Mountains, 

Beaver County (report on shipment 0-58-26). "Topache 

Limestone"?. Zone II?. Vesiculophyllum.

U-18. Unnamed locality, sec. 24, T. 4 S., R. 2 E., Utah County 

(report on shipment MD-54-10). Deseret Limestone. 

Zone III. Cyathaxonia?, Michelinia.

U-19. Unnamed locality, sec. 35, T. 23 S., R. 18 W., Millard County 

(report on shipment 0 and G-77-5). Chainman Shale. 

Zone V. Amplexizaphrentis, Barytichisma?, Michelinia.

U-20. Unnamed locality, sees. 30, 31, T. 1 S., R. 24 E., Uintah 

County (see Kinney, 1955). Madison Limestone. Zone II. 

Michelinia, Sychnoelasma, Syringopora, Vesiculophyllum.

U-21. Unnamed locality, sec. 30, T. 10 N., R. 2 E., Cache County 

(Sando, 1983). Monroe Canyon Limestone. Zone HID. 

Acrocyathus (cerioid).

U-22. Paradise Canyon, sec. 5 or 6, T. 9 N., R. 2 E., Cache County (Sando, 1983). Monroe Canyon Limestone. Zone IV. 

Petalaxis.

U-23. East Canyon, sec. 7, T. 9 N., R. 2 E., Cache County (Sando, 

1983). Monroe Canyon Limestone. Zone VA. Schoenophyllum.

WYOMING

WY-1. Unnamed locality, sec. 8, T. 45 N., R. 104 W., Park and Fremont Counties (report on shipment EEG-76-31). Madison
44 CORAL ZONATION, MISSISSIPPIAN SYSTEM, WESTERN INTERIOR PROVINCE

Limestone. Zone II. Lophophyllum?, Sychnoelasma?, 

Vesiculophyllum.

WY-2. Crowfoot Ridge and Bighorn Pass, Yellowstone Park (report 

on shipment NR-67-2). Madison Limestone. Zone II. 

Cleistopora, Sychnoelasma, Syringopora, Vesiculophyllum.

WY-3. Bannock Peak, 44°53'N. 110°52.5'W., Mt. Holmes 

quadrangle, Yellowstone Park (report on shipment 

NR-68-1). Lodgepole and Mission Canyon Limestones. 

Zone n. Amplexizaphrentis, Sychnoelasma, Vesiculophyllum.

WY-4. Unnamed locality, sees. 12, 18, T. 42 N., R. 115 W., Teton 

County (report on shipment RME-69-6). Madison 

Limestone. Zone II. Caninia, Lophophyllum?, Syringopora.

WY-5. Unnamed locality, sec. 3, T. 42 N., R. 118 W., Teton County (report on shipment CD-66-1). Mission Canyon 

Limestone? Zone IIIA. "Diphyphyllum."

WY-6. Unnamed locality, sees. 11,24,27, T. 41 N., R. 118 W., Teton 

County (report on shipment CD-66-1). Mission Canyon 

Limestone. Zones II, IIIA. "Diphyphyllum,"Lophophyllum?, 

Vesiculophyllum.

WY-7. Unnamed locality, sec. 11, T. 42 N., R. 115 W., Teton County 

(report on shipment NR-63-17D). Unnamed formation. 

Zone I or II. Lophophyllum?.

WY-8. Unnamed locality, sec. 30, T. 38 N., R. 118 W., Lincoln County (report on shipment CD-63-10). Lodgepole Limestone. 

Zone II. Sychnoelasma, Vesiculophyllum.

WY-9. Unnamed locality, sec. 28, T. 45 N., R. 60 W., Weston County (report on shipment MD-57-21D). Pahasapa Limestone. 

Zone II. Syringopora, Vesiculophyllum.

WY-10. Unnamed locality, sec. 9, T. 28 N., R. 92 W., Fremont County (report on shipment F-54-7D). Madison Limestone. 

Zone II?. Syringopora.

WY-11. Darby Mountain, sec. 8, T. 29 N., R. 115 W., Sublette County 

(report on shipment from P. E. Cloud, July 2, 1955). Mission 

Canyon Limestone. Zone II. Sychnoelasma, Syringopora, 

Vesiculophyllum.

WY-12. Unnamed locality, sec. 29, T. 43 N., R. 106 W., Fremont 

County (report on shipment F-53-33). Madison Limestone. 

Zone IIIA. "Diphyphyllum," Siphonodendron, Syringopora, 

Vesiculophyllum, Zaphriphyllum.

WY-13. Waterdog Lake, sec. 15, T. 38 W., R. Ill W., Sublette County (report on shipment F-55-19). Madison Limestone. 

Zone IIIA. Syringopora, Vesiculophyllum, Zaphriphyllum. 

WY-14. Unnamed locality, sec. 19, T. 41 N., R. 114 W., Teton County 

(report on shipment F-53-43). Madison Limestone. Zone II. 

Stelechophyllum.

WY-15. Unnamed locality, sec. 3, T. 49 N., R. 83 W., Johnson County 

(report on shipment F-50-34). Madison Limestone. Zone n?). 

Syringopora.

WY-16. Cherry Creek, sees. 19, 30, T. 31 N., R. 99 W., Fremont 

County (see Sando, 1975b). Amsden Formation. Zone VI. 

Barytichisma.

CANADA

The following list includes (1) localities represented by 

detailed measured sections (in plate captions, referred 

to as "sections") and (2) localities not represented by 

such sections (in plate captions, referred to as 

"localities"). The list is based mainly on data from internal Geological Survey of Canada (GSC) 

biostratigraphic reports by Bamber but also includes information from many publications on the Mississippian 

of western Canada. Geographic location, published or 

unpublished references, GSC locality numbers (where 

applicable), stratigraphic level, coral zones identified, 

and coral genera present are given for each locality.

DETAILED MEASURED SECTIONS

SOUTHWEST DISTRICT OF MACKENZIE

SWDM-1. Sheaf Creek, 61°12'23"N., 124°33'32"W. (GSC internal 

report), all measurements above base of Flett Formation. 

Amplexizaphrentis, GSC Iocs. C-52462, C-52463, 786 to 

837 ft, Zone IIIB (foraminiferal Zone 12); 

"Diphyphyllum,"GSC loc. C-47942,1,014 ft, Zone HID; 

Ekvasophyllum, GSC Iocs. C-52472, C-52485, 917 to 

1,081ft, Zone IIIB or IIIC, HID; Siphonodendron, 

GSC Iocs. C-47942, C-52479, C-52489, C-52490, 963 to 

1,122ft, Zone IIIB or IIIC, HID; Syringopora, 

GSC loc. C-47492, 1,014 ft, Zone HID.

SWDM-2. Twisted Mountain, ei 0 !!^, 123°27'38"W. (GSC internal report), all measurements above base of Flett Formation. Amplexizaphrentis, GSC Iocs. C-59036, 

C-59040, 284 to 289 ft, Zone IIIA; Caninia, 

GSC loc. C-59049, 627 ft, Zone IIIB; Siphonodendron, 

GSC Iocs. C-59028, C-59036, C-59049, 155 to 627 ft, 

Zones IIB?, IIIA, IIIB; Lophophyllum?, 

GSC Iocs. C-59039, C-59043, C-59049, 280 to 627 ft, 

Zone IIIA, IIIB; Vesiculophyllum, GSC loc. C-59043, 323 

to 337 ft, Zone IIIA; Zaphriphyllum, GSC Iocs. C-59039, 

C-59040, C-59043, 280 to 337 ft, Zone IIIA.

SWDM-3. North end Mattson anticline, 61°06'38"N., 

123°45'54"W. (GSC internal report), all measurements 

above base of Flett Formation. Amplexizaphrentis, 

GSC Iocs. C-47925, C-52234, C-52239, 80 to 1,123 ft, 

Zones IIB, IIIA, IIIB; "Diphyphyllum," 

GSC loc. C-52268, 1,504 to 1,513 ft, Zone IIIC or lower 

Zone HID (foraminiferal Zone 13); Siphonodendron, 

GSC Iocs. C-47929, C-47930, C-47933, C-52205, 

C-52238, 443 to 1,095 ft, Zone IIIA, mB; Lophophyllum?, 

GSC Iocs. C-47948, C-52234, C-52239, C-52240,1,041 

to 1,394 ft, Zone IIIB, IIIC, or lower HID (foraminiferal 

Zone 13); Syringopora, GSC loc. C-52268, 1,504 to 

1,513 ft, Zone IIIC or lower HID (foraminiferal Zone 13): 

Vesiculophyllum, GSC Iocs. C-47926, C-47929, C-52238, 

517 to 1,095 ft, Zone IIIA, IIIB; Zaphriphyllum, 

GSC Iocs. C-47928 to C-47930, 717 to 1,077 ft, Zone IHA, 

IIIB.

SWDM-4. Jackfish Gap, type section of Flett Formation, 

61°05'54"N., 123°59'26"W. (see Bamber and Mamet, 

1978), all measurements above base of Flett Formation. 

Amplexizaphrentis, GSC Iocs. C-52108, C-52111, 

C-52534, C-52538, C-52541, C-52544, C-52545, 

C-52549, 68813, 68819, 68820, 267 to 1,437 ft, Zones HE? 

(foraminiferal Zone 9?), IIIA, IIIB, IIIC, HID; 

"Diphyphyllum," GSC Iocs. C-52124, C-52546,1,140 to 

1,248 ft, Zone IIIB?; Ekvasophyllum, GSC Iocs. C-47950, 

C-52113, C-52119, C-52543, C-52544, C-52546, 68816, 

68820, 965 to 1,437 ft, Zone IIIA, IIIB, IIIC, HID; Liardiphyllum, GSC Iocs. 68811, 68817, 782 and 1,273ft, 

Zone IIIA, IIIC; Siphonodendron, GSC Iocs. 68814,
LOCALITY REGISTER 45

68816, 68889, 928 to 1,336 ft, Zone IIIA, IIIB?, HID; 

Lophophyllum?, GSC Iocs. C-52126, C-52157, C-52538,

68803. 68806, 68811, 68813, 380 to 1,552 ft, Zones IIB, 

IIIA, IIIB, HID; Syringopora, GSC Iocs. C-52120, 

C-52546, 68802, 304 to 1,248 ft, Zones IIB, IIIB?, IIIC 

or HID; Vesiculophyllum, GSC Iocs. C-47950, C-52087,

68804. 432 to 625 ft, Zones IIB, IIIA; Zaphriphyllum, 

GSC Iocs. C-47950, C-52541, 625 to 658 ft, Zone IIIA.

SWDM-5. Flett Mountain, west of Liard River at Flett Rapids, 

60°42'N., 123°37W. (see Sutherland, 1958, section 11, 

fossil localities 37, 38, p. 34, 40). Amplexizaphrentis, 

Lophophyllum?, GSC loc. 16605, unit 5, section 11, upper Flett Formation, Zone IIIA or IIIB; Siphonodendron, 

GSC loc. 16606, unit 5, section 11, upper Flett Formation, Zone IIIA or IIIB.

SWDM-6. Sawmill Mountain, exact location of collecting locality 

in doubt, approximately eO'^S'N., 123°40'W. (see 

Sutherland, 1958, section 11, fossil locality 40, p. 34, 40). 

Ekvasophyllum, GSC loc. 16627, unit 4, section 11, middle Flett Formation, Zone IIIA.

SWDM-7. West flank Flett anticline, 60°33'50"N., 123°48'20"W. 

(GSC internal report), all measurements above base of 

Flett Formation. Amplexizaphrentis, GSC Iocs. C-47938, 

C-52396, 722 to 932 ft, Zone IIB (foraminiferal Zone 9); 

Siphonodendron, GSC loc. C-47941, 1,391 to 1,394 ft, 

Zone HID; Stelechophyllum, GSC loc. C-52434, 994 ft, 

Zone HID.

SWDM-8. Liard Range, 5 mi west of Liard River, 60°29'46"N., 

123°38'53"W. (GSC internal report), all measurements 

above base of Flett Formation. Siphonodendron, 

GSC Iocs. C-52361, 68829, 1,117 to 1,392 ft, Zone IIIC?, 

IHD; Stelechophyllum, GSC Iocs. 68829,1,388 to 1,392 ft, 

Zone HID.

SWDM-9. Liard Range South, 5 mi west of Liard River at Big 

Island, 60°29'N., 123°38 rW. (see Sutherland, 1958, 

section 11, fossil localities 30, 32, p. 34, 39). Amplexizaphrentis, GSC loc. 16617, unit 6, section 11, upper 

Flett Formation, Zone IIIC or HID; Vesiculophyllum, 

GSC loc. 16615, unit 5, section 11, upper Flett Formation, Zone IIIB or IIIC.

3WDM-10. Liard River area, 2 mi west of Liard River at Big Island, 

60°29'N., 123°33'30"W. (see Sutherland, 1958, 

section 11, fossil localities 28, 29, p. 34, 39). Amplexizaphrentis, Ekvasophyllum, Liardiphyllum, 

Lophophyllum?, Zaphriphyllum, GSC loc. 16622, upper 

60ft of unit 4, section 11, middle Flett Formation, 

Zone IIIA.

NORTHEASTERN BRITISH COLUMBIA

NEBC-1. Gulf States Chuatse Cr. (1) b-14-I/94-J-16, subsurface 

(see Bamber and Mamet, 1978), all measurements below 

top of Debolt Formation, Zone IIIA. Siphonodendron, 

GSC Iocs. 55772, 55775, 1 to 8 ft.

NEBC-2. Imperial Kahntah (1) b-29-C/94-I-7, subsurface (see 

Bamber and Mamet, 1978). Sychnoelasma, 

GSC loc. 43467, 223 ft below top of Shunda Formation, 

Zone IIB; Syringopora, GSC Iocs. 43456, 43458, 222 to 

230ft below top of Debolt Formation, Zone IIIA? 

(foraminiferal Zone 10); Vesiculophyllum, 

GSC Iocs. 43459, 43463, 227 to 239 ft below top of Debolt 

Formation, Zone IIIA? (foraminiferal Zone 10).

NEBC-3. Imperial Sikanni Chief (1) b-92-D/94-I-4, subsurface 

(see Bamber and Mamet, 1978), Ekvasophyllum?, 

GSC loc. 55794, 237 ft below top of lower Debolt Formation, Zone IIIA; Siphonodendron, GSC loc. 43420, 242 ft 

below top of lower Debolt Formation, Zone IIIA; 

Vesiculophyllum, GSC loc. 43429, 108 ft below top of 

Shunda Formation, Zone IIB.

NEBC-4. HB-Imperial-Union Paddy (1) a-49-B/94-H-16, subsurface (see Bamber and Mamet, 1978), all measurements 

below top of Debolt Formation, Zone IIIA. Siphonodendron, Vesiculophyllum, GSC loc. 43509, 277 ft; 

Zaphriphyllum, GSC loc. 43512, 282 ft.

NEBC-5. West Canadian Lily Lake C-81-F/94-G-2, subsurface 

(see Bamber and Mamet, 1978), all measurements below 

top of Debolt Formation, Zone HID. Amplexizaphrentis, 

GSC loc. 59375,685 ft; ?Ekvasophyllum, GSC Iocs. 59358, 

59376, 566 to 575 ft; Siphonodendron, GSC Iocs. 59359, 

59364, 59366, 59367, 59379, 513 to 572ft; 

Lophophyllum?, GSC loc. 59371, 105 ft; Stelechophyllum, 

GSC Iocs. 59362, 59368, 59377, 115 to 203ft; Syringopora, GSC Iocs. 59366, 59376, 59379, 545 to 567 ft; 

Zaphriphyllum, GSC loc. 59372, 681 ft.

NEBC-6. Pacific Highway (2) a-47-I/94-B-16, subsurface (see 

Bamber and Mamet, 1978), all measurements below top 

of Debolt Formation, Zone IIIC or lower HID 

(foraminiferal Zone 13). Lophophyllum?, GSC Iocs. 43446, 

43452, 43453, 54 to 337 ft; Siphonodendron, 

GSC loc. 43447, 310 ft; Syringopora, GSC loc. 43454, 

309 ft.

NEBC-7. Pacific Highway (1) b-25-I/94-B-16, subsurface (see 

Bamber and Mamet, 1978), all measurements below top 

of lower Debolt Formation, Zone IIIB. Amplexizaphrentis, GSC Iocs. 43355, 43371, 43375, 43391, 50 to 134 ft; 

Caninia, GSC Iocs. 43370, 43387, 45 to 55 ft; Koninckophyllum?, GSC loc. 43384, 39 ft; Siphonodendron, 

GSC Iocs. 43355, 43357, 43361, 43363, 43379, 43387, 55 

to 245 ft; Lophophyllum?, GSC Iocs. 43361, 43374, 43378, 

72 to 188 ft; Syringopora, GSC Iocs. 43357,43359,43375, 

77 to 105 ft; Vesiculophyllum, GSC Iocs. 43357, 43360, 

43367, 43368, 43370, 43371, 43380, 43384, 24 to 81 ft; 

Zaphriphyllum, GSC Iocs. 43350, 43357 to 43359,43366, 

43370, 43379, 43386, 45 to 245 ft.

NEBC-8. Pacific Highway (3) a-69-I/94-B-16, subsurface (see 

Bamber and Mamet, 1978). Ekvasophyllum, 

GSC Iocs. 43393, 43400, 65.5 to 83 ft below top of lower 

Debolt Formation, Zone IIIB; Siphonodendron, 

GSC Iocs. 43394, 43409, 75 to 110 ft below top of lower 

Debolt Formation, Zone IIIB; Lophophyllum?, 

GSC loc. 43399, 93 ft below top of Debolt Formation, 

Zone HID? (foraminiferal Zone 13 (upper)); Syringopora, 

GSC Iocs. 43402, 43405, 43406, 87 ft below top of lower 

Debolt Formation to 95.5 ft below top of Debolt Formation, Zone IIIB, HID; Vesiculophyllum, GSC Iocs. 43395, 

43401, 43407, 40 to 88 ft below top of lower Debolt Formation, Zone IIIB; Zaphriphyllum, GSC Iocs. 43397, 

43414,43417, 44038, 70 to 93 ft below top of lower Debolt 

Formation, Zone IIIB.

NEBC-9. West Nat. Gundy Creek b-69-A/94-B-16, subsurface 

(see Bamber and Mamet, 1978), all measurements below 

top of Debolt Formation, Zone HID. Ekvasophyllum, 

GSC Iocs. 43497, 43501, 52 to 84 ft; Siphonodendron, 

GSC loc. 43490, 52 to 56 ft; Syringopora, 

GSC Iocs. 43496, 43502, 50.5 to 53 ft.
46 CORAL ZONATION, MISSISSIPPIAN SYSTEM, WESTERN INTERIOR PROVINCE

NEBC-10. West Nat. et al. Blueberry C-65-D/94-A-13, subsurface 

(see Bamber and Mamet, 1978). Siphonodendron, Syringopora, GSC loc. 55771, 33 ft below top of Debolt Formation, Zone HID?.

NEBC-11. Decalta Blueberry d-57-D/94-A-13, subsurface (see 

Bamber and Mamet, 1978), all measurements below top 

of Debolt Formation. Siphonodendron, GSC Iocs. 70452 

to 70454, 19 to 79 ft, Zone IIIA; Lophophyllum?, Syringopora, GSC loc. 70455, 117ft, Zone IIIC or lower 

HID.

NEBC-12. West Nat. et al. Blueberry d-46-D/94-A-13, subsurface 

(see Bamber and Mamet, 1978). Siphonodendron, 

GSC loc. 55793, 51 to 56 ft below top of Debolt Formation, Zone HID.

NEBC-13. West Nat. et al. Blueberry (13) d-50-K/94-A-12, subsurface (see Bamber and Mamet, 1978), all measurements 

below top of Debolt Formation. Siphonodendron, 

GSC Iocs. 43471, 43472, 146 to 168 ft, Zone HID; Syringopora, GSC Iocs. 43470, 43474, 145 to 161ft, 

Zone HID.

NEBC-14. West Nat. et al. Bernadet 8-1-88-25W6, subsurface (see 

Bamber and Mamet, 1978), all measurements below top 

of Debolt Formation. Siphonodendron, GSC Iocs. 70446 

to 70449, 266 to 288 ft, Zone HID; Lophophyllum?, Syringopora, GSC loc. 70447, 267 ft, Zone HID.

NEBC-15. West Nat. et al. Halfway (3) 8-11-87-25W6, subsurface 

(see Bamber and Mamet, 1978). Lophophyllum?, 

GSC loc. 55787, 122 ft below top of Debolt Formation, 

Zone HID? (foraminiferal Zone 13 or 14).

NEBC-16. Texaco NFA Boundary Lake (1) 6-6-86-13W6, subsurface (see Bamber and Mamet, 1978). Stelechophyllum?, 

GSC loc. 55780, 24 ft below top of Debolt Formation, 

Zone HID; Syringopora, GSC Iocs. 55777, 55778, 31 to 

58 ft below top of Debolt Formation, Zone HID.

NEBC-17. South Chlotopecta Creek, 3V2 mi south of Chlotopecta 

Creek, 58°27'N., 123°55W., all measurements below top 

of Prophet Formation (GSC internal report), Zone IIIC 

or HID (lower). Canadiphyllum, GSC Iocs. 66680, 66681, 

66684, 66689, 43 to 195 ft; Ekvasophyllum?, 

GSC loc. 66682,165 ft; Stelechophyllum, GSC loc. 66680, 

192 to 195 ft.

NEBC-18. Bat Creek (type section of Prophet Formation), cliffs north 

of Bat Creek waterfall, 57°47'N., 123°37W. (see Bamber 

and Mamet, 1978), all measurements below top of Prophet Formation. Acrocyathus?, GSC loc. 66587, 1 to 15 ft, 

Zone lllD;Amplexizaphrentis?, GSC Iocs. 66584, 66589, 

1 to 66 ft, Zone IIB? (foraminiferal Zone 9 or 10), HID; 

Canadiphyllum, GSC Iocs. 16045, 66585, 66586, 66587, 

1 to 195ft, Zone IIIB?, HID; Ekvasophyllum, 

GSC Iocs. C-11670, 66584, 66587,1 to 50? ft, Zone HID; 

Siphonodendron, GSC Iocs. 16030, 66588, 325 to 345 ft, 

Zone IIIB (foraminiferal Zone 12); Stelechophyllum?, 

GSC Iocs. 16034, 66584, 66586, 66587, 0 to 68 ft, 

Zone HID.

NEBC-19. Trimble Lake, 3.5 mi north of west end of Trimble Lake, 

57°19'37"N., 123°43'30"W. (see Bamber and Mamet, 

1978). Ekvasophyllum, GSC loc. 66524, 48 ft below top 

of Prophet Formation, Zone HID or lower Zone IV 

(foraminiferal Zone 14?).

NEBC-20. Trimble Lake East, 3 mi northeast of east end of Trimble Lake, 57 °17 TST., 123 °31 W., all measurements below 

top of Prophet Formation (GSC internal report). 

Acrocyathus?, GSC Iocs. 66582, 66583, 5 to 14 ft, 

Zone HID; Canadiphyllum, GSC Iocs. 66573, 66581, 16

to 509 ft, Zone IIIC?, HID; Ekvasophyllum, 

GSC loc. C-11684, 30 ft, Zone HID; Siphonodendron, 

GSC loc. 66579, 190ft, Zone HID; Stelechophyllum, 

GSC loc. 66582, 14 ft, Zone HID.

NEBC-21. Halfway River, north side of river valley, 56°59'N., 

123°29W. (see Sutherland, 1958, section 7, unit 6, p. 24, 

40). Canadiphyllum, Ekvasophyllum, GSC loc. 16038, upper 20 ft of member B Prophet Fm., Zone IIIB or IIIC.

NEBC-22. Halfway River South, 3 mi southwest of Halfway River, 

56°54'N., 123°27'30"W.; corals from upper part of Prophet Formation, faulted section, footage uncertain (GSC internal report). Ekvasophyllum, GSC loc. C-11649, 

Zone HID?, Siphonodendron, GSC loc. C-11647, 

Zone HID.

NEBC-23. Nabesche River, 24 mi north of Peace River, 56°20'N., 

123° 15'W. (see Bamber and Mamet, 1978), 

measurements below top of Prophet Formation, unless 

indicated otherwise. Acrocyathus, GSC loc. 66411, 327 

to 333 ft, Zone HID; Amplexizaphrentis, GSC Iocs. 66375, 

66421, 9 to 2,360 ft, Zone IIIB, IIIC; Canadiphyllum, 

GSC Iocs. 66376, 66378 to 66380, 66388, 66389, 66391, 

66401,897 to 2,186 ft, Zone Ilffi to HID; "Diphyphyllum," 

GSC Iocs. 66391 to 66392, 66396, 66399, 66413, 238 to 

1,456 ft, Zone IIIC, HID; Ekvasophyllum, 

GSC Iocs. C-11554, C-11558, 66389, 66397, 66401 to 

66403, 66405 to 66411, 66413, 66417 to 66420, 66422, 

66427,1,621 ft below top of Prophet Formation to 239 ft 

above base of Golata Formation, Zone HID, lower 

Zone IV?; Koninckophyllum, GSC Iocs. C-11554, 66421, 

9 to 10 ft, lower Zone IV?; Siphonodendron, 

GSC Iocs. C-11556, C-11562, C-11563, 66390, 66392, 

66395, 66404, 66406, 66411, 327 to 1,597 ft, Zone HID; 

Stelechophyllum, GSC Iocs. C-11557, C-11560, 66411, 

170 to 333 ft, Zone HID; Zaphriphyllum, GSC Iocs. 66376, 

66378, 66380 to 66386, 66390,66391, 66394, 66396,1,351 

to 2,186 ft, Zone IIIB, IIIC.

NEBC-24. Mount Greene, east side of mountain, 56°05'N, 

123° 15'W. (GSC internal report). Ekvasophyllum, 

GSC Iocs. C-11806, C-11811, top of Prophet Formation 

to 475 ft below top, Zone HID.

NEBC-25. Peace River, 4 mi southwest of Mt. Greene, north side 

of Peace River, 56°02'N., 123°12W., all measurements 

below top of Prophet Formation (GSC internal report). 

Acrocyathus, GSC loc. C-11741, 130ft, Zone HID; 

Ekvasophyllum, GSC loc. C-11738,15 to 25 ft, Zone HID; 

Siphonodendron, GSC loc. C-11752,1,425 ft, Zone IIIC?; 

Lophophyllum?, GSC loc. C-11742, 160 ft, Zone HID.

NEBC-26. Clearwater Lake, 8 mi north-northwest of Clearwater 

Lake, 55°40'30"N., 122°57'W., all measurements below 

top of Prophet Formation, Zone IIIC (GSC internal 

report). Canadiphyllum, GSC loc. C-4532, 190 ft; 

Ekvasophyllum, GSC Iocs. C-4528 to C-4531, 196 to 

320 ft; Siphonodendron, GSC loc. C^530, 264.5 to 266 ft; 

Lophophyllum?, GSC loc. C-4531, 196ft; Stelechophyllum?, GSC loc. C-4529, 303 to 309 ft.

NEBC-27. Solitude Mountain, roadcut on Hart Highway at north 

end of Solitude Mountain, 55 ° 30 '30 "N., 122°38'W., all 

measurements below top of Rundle Group, Zone IIIC 

(foraminiferal Zone 13) (GSC internal report). 

Siphonodendron, GSC Iocs. C-4513 to C-4517, 233 to 

381ft; Michelinia, GSC loc. C-4509, 79ft; Zaphriphyllum, GSC Iocs. C-4507, C-4510, 25 to 149 ft.

NEBC-28. Peck Creek, 1 mi northwest of headwaters of Peck Creek, 

55°44.5'N, 122°56W., all measurements below top of
LOCALITY REGISTER 47

Prophet Formation (GSC internal report). Acrocyathus, 

GSC Iocs. C-4542, C-4544, C-4546, 20 to 82 ft, 

Zone IIID; Ekvasophyllum, GSC Iocs. C-4341, C-4543 to 

C-4545, 7 to 96 ft, Zone IIIC?, IIID; Siphonodendron, 

GSC loc. C-4544, 22 to 24ft, Zone IIID; "Thysanophyllum," GSC loc. C-11666, top of Prophet Formation, 

Zone IIID.

NEBC-29. East Mountain Creek, ridge 1.5 mi east of Mountain 

Creek, SS^S'N., 122°25'W., all measurements below top 

of Debolt Formation (GSC internal report). Amplexizaphrentis, GSC Iocs. C-4562, C-4565, C-4568, C-4572, 

95 to 492ft, Zone IIIA to IIIC; Ankhelasma, 

GSC Iocs. C-4562, C-4567, 364? to 492 ft, Zone IIIA; 

Canadiphyllum?, GSC loc. C-4570, 221ft, Zone IIIB? 

(foraminiferal Zone 12); Caninia, GSC loc. C-4562, 

492ft, Zone IIIA; Ekvasophyllum, GSC Iocs. C-4558, 

C-4565, C-4573, 68 to 566ft, Zone IIIA to IIIC; 

Siphonodendron, GSC Iocs. C-4561, C-4569, 353 to 

526ft, Zone IIIA, IIIB?; Lophophyllum?, 

GSC Iocs. C-4557 to C-4560, C-4562, 490 to 575 ft, 

Zone IIIA; Vesiculophyllum, GSC Iocs. C-4557 to 

C-4560, C-4566, 379 to 575 ft, Zone IIIA; Zaphriphyllum, 

GSC Iocs. C-4557, C-4558, 566 to 575 ft, Zone IIIA.

NEBC-30. North Burnt River, 5 mi northwest of big bend of Burnt 

River, 55°14'30"N., 122°05'W. (GSC internal report). 

Amplexizaphrentis, GSC loc. C-4611, 296 to 311 ft above 

base of Shunda Formation, Zone IIB; Aulostylus, 

GSC loc. C-4603, 81 ft below top of Pekisko Formation, 

Zone IIB; Siphonodendron, GSC Iocs. C-4613 to C-4616, 

214 to 387 ft above base of Debolt Formation, Zone IIIA; 

Lophophyllum?, GSC Iocs. C-4600, C-4604, C-4611, 

161 ft below top of Pekisko Formation to 311 ft above 

base of Shunda Formation, Zone IIB; Sychnoelasma, 

GSC Iocs. C-4605 to C-4608, C-4610,132 to 270 ft above 

base of Shunda Formation, Zone IIB; Vesiculophyllum, 

GSC Iocs. C-4600 to C-4602, C-4604, C-4605, C-4607, 

C-4610, C-4611,161 ft below top of Pekisko Formation 

to 311 ft above base of Shunda Formation, Zone IIB; 

Zaphriphyllum, GSC Iocs. C-4600 to C-4602, C-4605, 

C-4611 to C-4613, 161 ft below top of Pekisko Formation to 214 ft above base of Debolt Formation.

NEBC-31. Burnt River, 2 mi north of big bend in Burnt River, 

55°12'N., 122°02'30"W. (GSC internal report). Amplexizaphrentis, GSC Iocs. C-4580 to C-4582, C-4585 to 

C-4587, C-4592, C-4593, C-4596, C-4598, 29 ft below 

top of Banff Formation to 358 ft above base of Debolt Formation, Zones IIB, IIIA; Aulostylus, GSC Iocs. C-11766, 

C-37921, Pekisko Formation?, Zone IIB; Cyathaxonia, 

GSC loc. C-4579, 38 to 41 ft below top of Banff Formation, Zone IIB?; Koninckophyllum, GSC loc. C-4595, 

248 ft above base of Debolt Formation, Zone IIIA; 

Lophophyllum?, GSC Iocs. C-4580, C-4585, C-4586, 

C-4590, C-4599, 29 ft below top of Banff Formation to 

381 ft above base of Debolt Formation, Zones IIB, IIIA; 

Michelinia, GSC Iocs. C-4582, C-4584,13 to 105 ft above 

base of Pekisko Formation, Zone IIB; Sychnoelasma, 

GSC Iocs. C-4580, C-4583 to C-4585, C-4587, 29 ft 

below top of Banff Formation to 32 ft above base of Debolt 

Formation, Zones IIB, IIIA (basal) (foraminiferal Zone 10 

(basal)); Vesiculophyllum, GSC Iocs. C-4585, C-4586, 

C-4589, C-4592 to C-4596, 142 ft above base of Shunda Formation to 258 ft above base of Debolt Formation, 

Zones IIB, IIIA; Zaphriphyllum, GSC Iocs. C-4584,

C-4594 to C-4596,105 ft above base of Pekisko Formation to 258 ft above base of Debolt Formation, Zones IIB, 

IIIA.

NEBC-32. North Sukunka River, 1 mi northeast of Sukunka River, 

55°05.5'N., 121°50'W., all measurements below top of 

Rundle Group (GSC internal report). Siphonodendron, 

GSC Iocs. C-l 1537, C-11538, 60 to 95ft, Zone IIIA; 

Stelechophyllum, GSC loc. C-11536, 870 ft, Zone IIB; 

Syringopora, GSC Iocs. C-11535, C-11537, 95 to 872 ft, 

Zones IIB, IIIA; Vesiculophyllum, GSC loc. C-11537, 

95 ft, Zone IIIA.

NEBC-33. Sukunka River, 2 mi east of Sukunka River, 

55°01'30"N., 121°42'W. (GSC internal report). Amplexizaphrentis, GSC loc. C-4619, 55 to 60 ft above base of 

Pekisko Formation, Zone IIB?; Aulostylus, 

GSC Iocs. C-4626, C-4627, 352 to 372 ft above base of 

Shunda Formation, Zone IIB; Michelinia, 

GSC loc. C-4619, 55 to 60 ft above base of Pekisko Formation, Zone IIB?; Stelechophyllum, GSC Iocs. C-4622 

to C-4625, 134 to 296 ft above base of Shunda Formation, Zone IIB; Sychnoelasma, GSC Iocs. C-4621, C-4627, 

C-4628,102 to 492 ft above base of Shunda Formation, 

Zone IIB; Vesiculophyllum, GSC Iocs. C-4621, C-4627, 

C-4628, 102 to 372 ft above base of Shunda Formation, 

Zone IIB.

NEBC-34. East Sukunka River, 4 mi east of Sukunka River, 

55°01'30"N., 121°38'W.; all measurements above base 

of Rundle Group (GSC internal report). Ankhelasma, 

GSC loc. C-4629, 1,471 ft, Zone IIIA; Aulostylus, 

GSC loc. C-7379, 365 ft, Zone IIB; Stelechophyllum, 

GSC Iocs. C-7380, C-7381, 280 to 298ft, Zone IIB; 

Sychnoelasma, GSC loc. C-7379, 365 ft, Zone IIB; Syringopora, GSC Iocs. C-7379, C-7380, 298 to 365 ft, 

Zone IIB; Vesiculophyllum, GSC Iocs. C-4631, C-4633, 

C-7380, C-7381, 280 to 995 ft, Zone IIB; Zaphriphyllum, 

GSC Iocs. C-4629 to 4631,995 to 1,471 ft, Zones IIB, IIIA.

NEBC-35. Hook Creek North, ridge east of Hook Creek, 54 °49'N., 

121°21'W. (GSC internal report). Siphonodendron, 

GSC loc. C-l 1637, 1 ft below top of Debolt Formation, 

Zone IIIA.

NEBC-36. Hook Creek, 5 mi southeast of Hook Lake and 1 mi east 

of Hook Creek, 54°48'N., 121°19'W. (GSC internal 

report). Amplexizaphrentis, Ankhelasma, Ekvasophyllum, 

GSC loc. C-7316, 94 ft above base of Debolt Formation, 

Zone IIIA; Siphonodendron, GSC Iocs. C-7313, C-7314, 

C-7317, C-7321, C-7322, C-11544, 9 to 214 ft above base 

of Debolt Formation, Zone IIIA; Lophophyllum?, 

GSC loc. C-11546, 84 to 94 ft above base of Debolt Formation, Zone IIIA; Michelinia, GSC loc. C-7309, 124 to 

129ft above base of Shunda Formation, Zone IIB; 

Sychnoelasma, GSC Iocs. C-7308, C-7309, C-7310, 82 

to 212 ft above base of Shunda Formation, Zone IIB; 

"Thysanophyllum," GSC Iocs. C-7318, C-7319, C-7320, 

126 to 131 ft above base of Debolt Formation, Zone IIIA; 

Vesiculophyllum, GSC Iocs. C-7304, C-7307, C-7308, 

C-7310, C-7313, C-7317, C-7322, 138 ft below top of 

Banff Formation to 214 ft above base of Debolt Formation, Zones II, IIIA; Zaphriphyllum, GSC Iocs. C-7312, 

C-7316, 341 ft above base of Shunda Formation to 94 ft 

above base of Debolt Formation, Zones IIB, IIIA.

NEBC-37. East Fellers Creek, 3 mi northwest of Bone Mountain, 

54°42.5'N., 120°54'W. (GSC internal report). 

Ankhelasma, GSC Iocs. C-7347, C-7350, C-7351, 2 to 

38.5 ft above base of Debolt Formation, Zone IIIA;
48 CORAL ZONATION, MISSISSIPPIAN SYSTEM, WESTERN INTERIOR PROVINCE

Siphonodendron, GSC Iocs. C-7349 to C-7357, 37 to 

496 ft above base of Debolt Formation, Zone IIIA; 

Lophophyllum?, GSC loc. C-7342, 161 ft above base of 

Shunda Formation, Zone IIB; Sychnoelasma, 

GSC Iocs. C-7341 to C-7344, 156 to 1,053 ft above base 

of Shunda Formation, Zone IIB; Syringopora, 

GSC Iocs. C-7345, C-7347, C-7352, 284 ft above base of 

Shunda Formation to 173 ft above base of Debolt Formation, Zones IIB, IIIA; Vesiculophyllum, 

GSC Iocs. C-7340 to C-7343, C-7345 to C-7348, C-7350, 

C-7352,122 ft above base of Shunda Formation to 173 ft 

above base of Debolt Formation, Zones IIB, IIIA; 

Zaphriphyllum, GSC Iocs. C-7343, C-7345 to C-7348, 

C-7355,171 ft above base of Shunda Formation to 471 ft 

above base of Debolt Formation, Zones II, IIIA.

NEBC-38. Murray River, 3 mi east of Murray River, 54°39'N., 

120°59W. (GSC internal report). Siphonodendron, 

GSC loc. C-11798, 515 ft below top of Rundle Group, 

Zone IIIA?.

NEBC-39. Mount Becker, 1 mi southwest of Mount Becker, 

54°31'N., 120°39'W. (GSC internal report). Ankhelasma, 

GSC loc. C-7377, 257 to 259 ft above base of Debolt Formation, Zone IIIA; Dorlodotia?, GSC Iocs. C-7372 to 

C-7374, 124 to 140 ft above base of Debolt Formation, 

Zone mA; Siphonodendron, GSC Iocs. C-7364 to C-7370, 

393 ft above base of unnamed carbonate-bank facies of 

Rundle Group to 90 ft above base of Debolt Formation, 

Zone IIIA; Lophophyllum?, GSC Iocs. C-7358, C-7363, 

C-7363A, C-7365, 9 ft below top of Banff Formation to 

556 ft above base of unnamed carbonate-bank facies of 

Rundle Group, Zones II, IIIA; Sychnoelasma, 

GSC Iocs. C-7363, C-7363A, 161 to 182 ft above base of 

unnamed carbonate-bank facies of Rundle Group, 

Zone IIB; Syringopora, GSC Iocs. C-7368, C-7372, 

C-7374, 614 ft above base of unnamed carbonate-bank 

facies of Rundle Group to 140 ft above base of Debolt Formation, Zone IIIA; Vesiculophyllum, GSC Iocs. C-7363, 

C-7366, C-7367, C-7374,161 ft above base of unnamed 

carbonate-bank facies of Rundle Group to 169 ft above 

base of Debolt Formation, Zones IIB, IIIA.

NEBC-40. Belcourt Creek, 4 mi east-northeast of Muinok Mountain, 

54°22'N., 120°29.5'W. (GSC internal report). Ankhelsma, 

GSC Iocs. C-7398, C-7399, 102 to 181 ft above base of 

Turner Valley Formation, Zone IIIA; Cyathaxonia, 

GSC loc. C-7404, 446 to 450 ft above base of unnamed 

carbonate-bank facies of Rundle Group, Zone IIIA; 

Siphonodendron, GSC Iocs. C-7395, C-7396, C-7404, 

446 ft above base of unnamed carbonate-bank facies of 

Rundle group to 264 ft above base of Turner Valley Formation, Zone IIIA; Lophophyllum?, GSC Iocs. C-7401, 

C-7412 to C-7414, 50ft below top of Banff Formation 

to 39 ft above base of unnamed carbonate-bank facies of 

Rundle Group, Zones IIB, IIIA; Michelinia, 

GSC loc. C-7413, 47 ft below top of Banff Formation, 

Zone IIB; Sychnoelasma, GSC Iocs. C-7406, C-7410 to 

C-7412,158 ft above base of Shunda Formation to 436 ft 

above base of unnamed carbonate-bank facies of Rundle 

Group, Zone IIB; Syringopora, GSC Iocs. C-7410, 

C-7413, 47 ft below top of Banff Formation to 49 ft above 

base of unnamed carbonate-bank facies of Rundle Group, 

Zone IIB; Vesiculophyllum, GSC Iocs. C-7394, C-7397 to 

C-7399, C-7402, C-7407, C-7409, C-7412,158 ft above 

base of Shunda Formation to 361 ft above base of Turner 

Valley Formation, Zones HB, IIIA, ?mB; Zaphriphyllum,

GSC Iocs. C-7396, C-7398, C-7409, 99 ft above base of 

unnamed carbonate-bank facies of Rundle Group to 197 ft 

above base of Turner Valley Formation, Zones IIB, IIA.

NEBC-41. West Belcourt Creek, 4 mi west of Muinok Mountain, 

54°20'30"N., 120°30'W. (GSC internal report). 

Siphonodendron, GSC loc. C-80126, 10 ft below top of 

Shunda Formation, Zone IIB?.

NEBC-42. North Hanington, 5.5 mi northwest of Mount Hanington, 

54°10'30"N., 120°15'W. (GSC internal report), all 

measurements above base of Turner Valley Formation. 

Amplexizaphrentis, GSC loc. C-79628, 384 ft, Zone IIIA; 

Ankhelasma, GSC Iocs. C-79613, C-79627, C-79632, 20 

to 485 ft, Zone IIIA; "Diphyphyllum," GSC loc. C-79632, 

485 ft, Zone IIIA; Siphonodendron, GSC loc. C-79628, 

384 ft, Zone IIIA; Vesiculophyllum and Zaphriphyllum, 

GSC loc. C-79613, 20 ft, Zone IIIA.

NEBC-43. Ridge west of Mt. Hanington, 1.5 mi north of Jarvis 

Lakes, 54°07'N., 120°12'30"W. (GSC internal report). 

Amplexizaphrentis, GSC loc. C-7423, 353 ft above base 

of Turner Valley Formation, Zone IIIA; Ankhelasma, 

GSC Iocs. C-7421, C-7423, 353 to 473 ft above base of 

Turner Valley Formation, Zone IIIA; Aulostylus, 

GSC loc. C-79554,64 ft above base of Shunda Formation, 

Zone IIB; Cyathaxonia, GSC loc. C-7421, 464 to 473 ft 

above base of Turner Valley Formation, Zone IIIA; 

Dorlodotia, GSC loc. C-7424, 336 to 338 ft above base of 

Turner Valley Formation, Zone IIIA; Sychnoelasma, 

GSC Iocs. C-79548, C-79559, 8 to 210 ft above base of 

Shunda Formation, Zone IIB; Syringopora, 

GSC Iocs. C-7418, C-7420, C-79548, 210 ft above base 

of Shunda Formation to 697 ft above base of Turner 

Valley Formation, Zones IIB, IIIA, IIIB?; 

Vesiculophyllum, GSC Iocs. C-7418, C-7422 to C-7424, 

C-79548,210 ft above base of Shunda Formation to 697 ft 

above base of Turner Valley Formation, Zones IIB, IIIA, 

IHB?; Zaphriphyllum, GSC loc. C-7419,519 ft above base 

of Turner Valley Formation, Zone IIIA.

ALBERTA

AL-1. Brewster's wall, 2 mi southwest of South Sulphur River, 

53°28'N., 118°54'W. (see Mountjoy, 1962, p. 32, 108). 

Siphonodendron, GSC loc. 40252, top of Shunda Formation, Zone IIIA; Lophophyllum?, Syringopora, 

GSC loc. 40255, 80 ft below top of Shunda Formation, 

Zone IIB.

AL-2. Folding Mountain, 0.75 mi south of Highway 16, 

53°14'30"N., 117°45'30"W. (GSC internal report; E. W. 

Bamber, unpublished data). Amplexizaphrentis, 

GSC loc. 55414, basal 10 ft of Turner Valley Formation, 

Zone IIIA; Dorlodotia?, GSC Iocs. 36772, 55416, 55417, 

38 to 43 ft above base of Turner Valley Formation, 

Zone IIIA; Siphonodendron, GSC Iocs. 36772, 36813, 

55413 to 55415, 55418, 290 ft above base of Shunda Formation to 30 ft above base of Turner Valley Formation, 

Zones IIB, IIIA; Lophophyllum?, GSC Iocs. 36772, 55412, 

55414, 52 ft above base of Pekisko Formation and basal 

10 ft of Turner Valley Formation, Zone IIIA; 

Sychnoelasma, GSC loc. 55409, 4 to 6 ft above base of 

Pekisko Formation, Zone IIB; Syringopora, 

GSC Iocs. 55413, 55414, 55416, 55418, 290 ft above base 

of Shunda Formation to 10 ft above base of Turner Valley
LOCALITY REGISTER 49

Formation, Zones IIB, IIIA; Vesiculophyllum, 

GSC Iocs. 55409, 55413 to 55415, 55418, 4 ft above base 

of Pekisko Formation to 30 ft above base of Turner Valley 

Formation, Zones IIB, IIIA; Zaphriphyllum, 

GSC loc. 55414, basal 10 ft of Turner Valley Formation, 

Zone IIIA.

AL-3. Roche De Smet, southeast end of De Smet Range, 

53°05'30"N., 118°04'30"W. (see Mountjoy, 1962, 

section 5, p. 32, 100; E. W. Bamber, unpublished data. 

Cleistopora, GSC loc. C-57119, 165 ft below top of Banff 

Formation, Zone IIA; Siphonodendron, GSC Iocs. 55419 

to 55423, 55431, 31 ft below top of Pekisko Formation 

to 137 ft below top of Turner Valley Formation, 

Zones IIB, IIIA; Lophophyllum?, GSC Iocs. C-57117, 

C-57119, 36756,155 ft below top of Banff Formation to 

90 ft below top of Shunda Formation; Stelechophyllum, 

GSC Iocs. 55424 to 55428, 55430, 4 ft above base of 

Pekisko Formation to 168 ft above base of Pekisko Formation, Zone IIB; Sychnoelasma, GSC Iocs. 55427 to 

55429, 55431, 55435, 55436,16 ft above base of Pekisko 

Formation to 85 ft above base of Shunda Formation, 

Zone IIB; Syringopora, GSC Iocs. C-57117, C-57119, 

55426 to 55429, 55431, 55432, 55434, 155 ft below top 

of Banff Formation to 58 ft above base of Shunda Formation; Vesiculophyllum, GSC Iocs. 36756, 36800, 55426 

to 55428, 55431 to 55436,16 ft above base of Pekisko Formation to 90 ft below top of Shunda Formation, Zone IIB; 

Zaphriphyllum, GSC loc. 55434, 53 to 58 ft above base 

of Shunda Formation, Zone IIB.

AL-4. Jasper lake, on south side of Highway 16, between Talbot 

Lake and Jasper Lake, 53°05'N., 118°01'30"W. (see 

Mountjoy, 1965, p. 342; E. W. Bamber, unpublished data). 

Siphonodendron, GSC Iocs. 55439 to 55442, uppermost 

Shunda Formation or lowermost Turner Valley Formation, Zone IIIA.

AL-5. Cadomin, along railroad cuts west of town, 52°59'N., 

117°20'W. (see Macqueen, 1966; GSC internal report). 

Syringopora, GSC Iocs. 74857, 74864,125 ft below top of 

Banff Formation to 28 ft above base of Turner Valley Formation, Zones IIB, IIIA; Vesiculophyllum, 

GSC loc. 74863, 31 ft above base of Shunda Formation, 

Zone IIB; Siphonodendron, GSC loc. 74864, 28 ft above 

base of Turner Valley Formation, Zone IIIA.

AL-6. Mount Merlin, 52°57'N., 117 0 43W. (GSC internal 

report). Siphonodendron, GSC Iocs. 74964 to 74966, 118 

to 162 ft above base of Turner Valley Formation, 

Zone IIIA; Stelechophyllum, GSC loc. 74961, 49 ft above 

base of Pekisko Formation, Zone IIB; Syringopora, 

GSC Iocs. 74962 to 74964, top of Shunda Formation to 

121 ft above base of Turner Valley Formation, Zone IIIA; 

Vesiculophyllum, GSC Iocs. 74960, 74961, 74964, 74965, 

5 ft above base of Pekisko Formation to 140 ft above base 

of Turner Valley Formation, Zones IIB, IIIA.

AL-7. Sirdar Mountain, 52°54'N., 117°48'W. (GSC internal 

report). Syringopora, GSC loc. 74912, 435 ft above base 

of Turner Valley Formation, Zone HI A; Vesiculophyllum, 

GSC loc. 74913, 7 ft above base of Mount Head Formation, Zone IIIA; Zaphriphyllum, GSC loc. 74914, 39 ft 

above base of Mount Head Formation, Zone IIIA.

AL-8. Deception Creek, 52°51'N., 117°21W. (GSC internal 

report). Siphonodendron, GSC Iocs. 74942, 74945, 74946, 

42 to 119ft above base of Turner Valley Formation, 

Zone mA; Michelinia, Stelechophyllum, GSC Iocs. 74937, 

74939, lower 29 ft of Pekisko Formation, Zone IIB; Syringopora, GSC Iocs. 74939, 74943, 74944, 29 ft above 

base of Pekisko Formation to 76 ft above base of Turner 

Valley Formation, Zones IIB, IIIA; Vesiculophyllum, 

GSC Iocs. 74933, 74937 to 74941, 74948, 418 ft above 

base of Banff Formation, to 91 ft above base of Mount 

Head Formation, Zones IIA or IIB, IIIA; ^Zaphriphyllum, 

GSC loc. 74938, 16 ft above base of Pekisko Formation, 

Zone IIB.

AL-9. Rocky River, 52 °47 'N., 117 ° 26 W. (GSC internal report). 

Siphonodendron, GSC Iocs. 74976 to 74979, 74981, 

74983, 61 ft above base of Turner Valley Formation to 

54 ft above base of Mount Head Formation, Zone IIIA; 

Michelinia, GSC Iocs. 74974, 74975,135 to 146 ft above 

base of Pekisko Formation, Zone IIB; Stelechophyllum, 

GSC Iocs. 74971, 74973, 3 to 9 ft above base of Pekisko 

Formation, Zone IIB; Syringopora, GSC Iocs. 74971, 

74982, 9 ft above base of Pekisko Formation to 62 ft 

above base of Mount Head Formation, Zones IIB, IIIA; 

Vesiculophyllum, GSC Iocs. 74971, 74980, 9 ft above base 

of Pekisko Formation to 32 ft above base of Mount Head 

Formation, Zones IIB, IIIA; Zaphriphyllum?, 

GSC loc. 74982, 62 ft above base of Mount Head Formation, Zone IIIA.

AL-10. Southesk-Cairn, 52°43'N., 117°12W. (GSC internal 

report). Siphonodendron, GSC Iocs. 74951 to 74954, 69 

to 232 ft above base of Turner Valley Formation, 

Zone IIIA; Syringopora, GSC Iocs. 74950, 74953, 96 ft 

above base of Turner Valley Formation to 35 ft above 

base of Mount Head Formation, Zone IIIA; 

?Vesiculophyllum, GSC loc. 74953, 96 ft above base of 

Turner Valley Formation, Zone IIIA.

AL-11. Nordegg, along David Thompson Highway, composite 

section, 52°29'N., 116°04'W. (GSC internal report). 

Stelechophyllum, GSC Iocs. 74867, 74869, 74870, 74871 

to 74874, lower part and uppermost part of Pekisko Formation, contacts not exposed, Zone IIB; Vesiculophyllum, 

GSC Iocs. 74867, 74873, lower part of Pekisko Formation, 

contact not exposed, Zone IIB.

AL-12. Job Creek, 52°23'N., 116°43'W. (GSC internal report). 

Michelinia, GSC loc. 74919,154 ft below top of Banff Formation, Zone IIA or IIB; Siphonodendron, 

GSC Iocs. 74925, 74926, 160 to 181 ft above base of 

Turner Valley Formation, Zone IIIA; Lophophyllum?, 

GSC loc. 74920, 126 ft below top of Banff Formation, 

Zone IIB; Stelechophyllum, GSC loc. 74922, base of 

Pekisko Formation?, Zone IIB; Syringopora, 

GSC loc. 74916, 308 ft below top of Banff Formation, 

Zone IIA or IIB; Vesiculophyllum, GSC Iocs. 74916, 

74924, 74928, 308 ft below top of Banff Formation, to 

16 ft above base of Mount Head Formation, Zones IIA 

or IIB, IIB, IIIA.

AL-13. Sentinel Mountain, 52°05'N., 116°26'W. (GSC internal 

report). Siphonodendron, GSC loc. 68537, 219 to 224 ft 

above base of Turner Valley Formation, Zone IIIA; Syringopora, GSC loc. 68521, 359 to 365 ft above base of 

Banff Formation, Zone IIA?.

AL-14. Forbidden Creek, 51 °48'N.) 115^OW. (see Macqueen 

and Bamber, 1967, 1968). Faberophyllum, 

GSC loc. 68592, 335 ft above base of Mount Head Formation, Zone IV; Siphonodendron, GSC Iocs. 68589, 

68591,133 to 404 ft above base of Turner Valley Formation, Zone IIIA; Stelechophyllum, GSC loc. 68575, 7 ft 

above base of Pekisko Formation, Zone IIB.
50 CORAL ZONATION, MISSISSIPPIAN SYSTEM, WESTERN INTERIOR PROVINCE

AL-15. Clearwater River, 51°47'N., 116°04'W. (see Macqueen 

and Bamber, 1967, 1968). Ekvasophyllum, 

GSC Iocs. 68506, 68510, 68512, 358 to 518 ft above base 

of Mount Head Formation, Zone HID; Siphonodendron, 

GSC Iocs. 68504, 68511, 68513, 595ft above base of 

Turner Valley Formation, to 518 ft above base of Mount 

Head Formation, Zones IIIA, HID; ?Petalaxis, 

GSC loc. 68511, 482 ft above base of Mount Head Formation, Zone HID; Syringopora, GSC loc. 68512, 518 ft 

above base of Mount Head Formation, Zone HID.

AL-16. Divide Creek, approximately 4 mi north of Mt. Tyrrell, 

51°45'N., 115°53'W. (see Macqueen and Bamber, 1967, 

1968). Amplexizaphrentis, GSC loc. 68562, 136 ft above 

base of Mount Head Formation, Zone HID?; Dorlodotia?, 

Ekvasophyllum, GSC loc. 68563,161 to 169 ft above base 

of Mount Head Formation, Zone HID; Stelechophyllum, 

GSC Iocs. 68551, 68563, 38 ft below top of Banff Formation to 169 ft above base of Mount Head Formation, 

Zones IIB, HID; Syringopora, GSC Iocs. 68545, 68547, 

68550 to 68552, 327 to 12 ft below top of Banff Formation; Vesiculophyllum, GSC loc. 68553,161 ft above base 

of Pekisko Formation, Zone IIB.

AL-17. Mount Tyrrell-Mount White, 51°40'N., 115°50'W. (see 

Macqueen and Bamber, 1967,1968). Amplexizaphrentis, 

GSC loc. 66136, 30 ft above base of Etherington Formation, Zone V?; Ekvasophyllum, GSC loc. 66135, top beds 

of Carnarvon Member, Mount Head Formation, Zone IV; 

Siphonodendron, Syringopora, GSC loc. 66132, uppermost Shunda Formation, Zone IIIA.

AL-18. McConnell Creek, north side of Red Deer River, 51 °39'N., 

115°58'W. (GSC internal report). Dorlodotia, 

GSC loc. C-22, 5 ft above base of Shunda Formation, 

Zone IIB; Siphonodendron, GSC loc. C-21, 20 ft below 

top of Shunda Formation, basal Zone IIIA; 

Lophophyllum?, GSC Iocs. C-17, C-21, 915 ft above base 

of Exshaw Formation, (5 ft above base of "middle 

member" of Banff Formation) to 20 ft below top of Shunda Formation, Zone IIB; Sychnoelasma, GSC loc. C-17, 

5 ft above base of "middle member" of Banff Formation, 

Zone IIB.

AL-19. Pagan's Lake, 51°36'N., 115°46'W. (see Macqueen and 

Bamber, 1967,1968). Siphonodendron, GSC loc. 68469, 

308 to 311ft above base of Turner Valley Formation, 

Zone IIIA; Lophophyllum?, GSC loc. 68456, 760 to 765 ft 

above base of Banff Formation, Zone IIB?; Syringopora, 

GSC Iocs. 68450, 68454, 68457, 545 to 873 ft above base 

of Banff Formation, Zones IIA?, IIB; Vesiculophyllum, 

GSC loc. 68464,172 ft above base of Pekisko Formation, 

Zone IIB.

AL-20. Sheep Creek, 51°35'N., 115°29'W. (see Macqueen and 

Bamber, 1967, 1968). Lophophyllum?, Syringopora, 

GSC loc. 66104, 41 ft below top of Banff Formation, 

Zone IIB; Stelechophyllum, GSC Iocs. 66105, 66106, 6 ft 

below top of Banff Formation to 12 ft above base of Shunda Formation, Zone IIB.

AL-21. Panther River, 51 °34'N., 115 °43W. (see Macqueen and 

Bamber, 1967, 1968). Amplexizaphrentis, GSC loc. 62081, 

74 ft below top of Loomis Member?, Mount Head Formation; Zone IIIC or HID; Siphonodendron, GSC loc. 62083, 

Opal Member, Mount Head Formation, talus, Zone HID; 

Lophophyllum?, GSC loc. 66137, 153.5 ft below top of 

Banff Formation, Zone IIB; Syringopora, 

GSC Iocs. 62080, 62082, 40 to 80 ft below top of Loomis 

Member?, Mount Head Formation, Zone IIIC or HID;

Vesiculophyllum, GSC Iocs. 66141, 66144, 66147,164 ft 

below top of Banff Formation, to 507? ft above base of 

Livingstone Formation, Zone IIB?.

AL-22. Cascade River, 51°25'N., 115°45'W. (see Macqueen and 

Bamber, 1967). Amplexizaphrentis, GSC Iocs. 73526, 

73527, 224 to 269 ft below top of Banff Formation, 

Zone IIA or EB; Michelinia, GSC loc. 73525, 286 to 289 ft 

below top of Banff Formation, Zone IIA or IIB; 

Stelechophyllum, GSC loc. 73532, 821 ft above base of 

Livingstone Formation, Zone IIB?; Sychnoelasma, 

GSC loc. 73531, 811 ft above base of Livingstone Formation Zone IIB; Syringopora, GSC loc. 73519, 761 ft below 

top of Banff Formation, (in lower half of formation), 

Zone I?

AL-23. Ghost River, near headwaters on northeast fork, 

51°23'30"N., 115°24'W. (see Macqueen and Bamber, 

1967, 1968). Siphonodendron, GSC Iocs. 68408, 68409, 

381 to 419 ft above base of Turner Valley Formation, 

Zone IIIA; Stelechophyllum, GSC Iocs. 68403, 68405, 

780 ft above base of Banff Formation to 189 ft above base 

of Pekisko Formation, Zone IIB; Sychnoelasma, 

GSC loc. 68404, 645 to 650 ft above base of Banff Formation, Zone IIB; Syringopora, GSC loc. 68394, 530 to 

545 ft above base of Banff Formation, Zone IIA or IIB; 

Vesiculophyllum, GSC Iocs. 68401, 68410, 739 ft above 

base of Banff Formation, to 62 ft above base of Mount 

Head Formation, Zones IIB, IIIA.

AL-24. Mount Aylmer, 2.5 mi northwest of mountain, 

51°20.5'N., 115°28W. (see Macqueen and Bamber, 1967, 

1968). Acrocyathus, GSC Iocs. 68389, 68391,104 ft above 

base of Loomis Member to top of member, Mount Head 

Formation, Zone HID; Clisiophyllum, GSC loc. 68392, 

72 ft above base of Carnarvon Member, Mount Head Formation, Zone IV; Ekvasophyllum, GSC loc. 68389, 47 ft 

above base of Loomis Member, Mount Head Formation, 

Zone IIIC or HID; Faberophyllum, GSC loc. 68392, 72 ft 

above base of Carnarvon Member, Mount Head Formation, Zone IV; Siphonodendron, GSC loc. 68387, 35 ft 

above base of Livingstone Formation, Zone IIB?; Syringopora, Vesiculophyllum, GSC Iocs. 68384, 68385, 880 

to 1,010 ft above base of Banff Formation, Zone IIB?.

AL-25. Stoney Creek, 51°22'N., 115°34'W., Mount Head Formation (see Macqueen and Bamber, 1968). Acrocyathus, 

GSC Iocs. 66156, 66157, 66160, 66 ft below top of Loomis 

Member to 25 ft above base of Opal Member, Zone HID; 

Ekvasophyllum, GSC Iocs. 66155, 66157, 66160, 86 ft 

below top of Loomis Member to 25 ft above base of Opal 

Member, Zone HID; Faberophyllum, GSC loc. 66163, 25 

to 26 ft above base of Carnarvon Member, Zone IV; 

Siphonodendron, GSC Iocs. 66153, 66156, 66157, 42 ft 

below top of Salter Member to 37 ft below top of Loomis 

Member, Zone IIIB?, HID; Syringopora, GSC Iocs. 66156, 

66157, 37 to 66 ft below top of Loomis Member.

AL-26. Camp Creek, 51°20'N., 115°29'W. (see Macqueen and 

Bamber, 1967,1968). Siphonodendron, GSC loc. 68374, 

248 ft above base of Livingstone Formation, Zone IIB?; 

Lophophyllum?, GSC loc. 68369, 652 ft below top of Banff 

Formation, Zone IIB?; Syringopora and Vesiculophyllum, 

GSC loc. 68367, 742 ft below top of Banff Formation, 

Zone IIA or IIB.

AL-27. Mount Costigan, north side of Lake Minnewanka, 

51°16'N., 115°17'W. (see Macqueen and Bamber, 1967). 

?Sychnoelasma, Vesiculophyllum, GSC loc. 74901, 964 ft
LOCALITY REGISTER 51

above base of Banff Formation, Zone IIB; Syringopora, 

GSC Iocs. 74900, 74902, 920 to 1,032 ft above base of 

Banff Formation, Zone IIB.

AL-28. Lake Minnewanka, north shore of lake, 51°15'N., 

115°28'W. (see Shimer, 1926; Crickmay, 1955; Macqueen 

and Bamber, 1967,1968). Acrocyathus, GSC Iocs. 62020, 

62026, 62028 to 62030, Mount Head Formation, 141 ft 

above base of Salter Member to 9 ft below top of Loomis 

Member, Zones IIIB, IIIC, HID; tAmplexizaphrentis, 

GSC Iocs. 62014, 62016, bed 14 of Crickmay (1955), 55 

to 63 ft below top of Livingstone Formation, Zone IIIA?; 

?Canadiphyllum, GSC loc. 62012, 88 to 89 ft below top 

of Livingstone Formation, Zone IIIA?; Dorlodotia?, 

GSC loc. 62017,22 ft below top of Livingstone Formation, 

Zone IIIA?; Ekvasophyllum, GSC Iocs. 62019, 62026, 

62032, 62035, 62036, Mount Head Formation, 45 ft above 

base of Salter Member to 19 ft above base of Carnarvon 

Member, Zones IIIB, IIIC, HID, IV; Faberophyllum, 

Mount Head Formation, Carnarvon Member, bed 3 of 

Crickmay (1955), Zone IV; Siphonodendron, 

GSC Iocs. 62002, 62004, 62009, 62026 to 62029, lowermost 10 ft of Livingstone Formation to 27 ft below top 

of Loomis Member, Mount Head Formation, Zones IIB, 

HID; Lophophyllum?, GSC Iocs. 60995, 62007, 282ft 

below top of Banff Formation to 451 ft above base of Livingstone Formation, Zone IIB; Petalaxis, GSC loc. 62031, 

5 to 9 ft below top of Loomis Member, Mount Head Formation, Zone HID; Sciophyllum, section 2, bed 8 of 

Shimer (1926), upper Opal Member, Mount Head Formation, Zone IV; Sychnoelasma, GSC loc. 62003,109 to 

110 ft above base of Livingstone Formation, Zone IIB; 

Syringopora, GSC Iocs. 62000, 62002, 62007, 62009, 

62015, 62017, 62026, 62028, 62031, 120 ft below top of 

Banff Formation to 5 ft below top of Loomis Member, 

Mount Head Formation, Zones IIB, IIIA, HID; 

"Thysanophyllum," GSC loc. 62032, 38 ft above base of 

Opal Member, Mount Head Formation, Zone HID; 

Vesiculophyllum, GSC loc. 62000,118 to 120 ft below top 

of Banff Formation, Zone IIB; Zaphriphyllum, 

GSC Iocs. 60996, 62015, 239 ft below top of Banff Formation to 60 ft below top of Livingstone Formation, 

Zones IIB?, IIIA.

AL-29. Tunnel Mountain, north side of Bow River, 51°10'N., 

115°33'W. (see Macqueen and Bamber, 1967, 1968). 

Acrocyathus, GSC loc. 58183, 62 ft above base of Opal 

Member, Mount Head Formation, Zone HID; 

Ekvasophyllum?, GSC loc. 58182, 57 ft above base of Opal 

Member, Mount Head Formation, Zone HID; 

Faberophyllum, GSC loc. 58185, 328 ft above base of Opal 

Member, Mount Head Formation, Zone IV; Siphonodendron, GSC loc. 58177, 49 ft below top of Livingstone Formation, Zone IIIA?. "Thysanophyllum," GSC type 

no. 8911, ca. 1,640 ft above base of Rundle Group, 

Zone IV.

AL-30. Carrot Creek, 51°10'N., 115°23'W. (see Macqueen and 

Bamber, 1967, 1968). Faberophyllum, GSC Iocs. 66171, 

66172,118 to 148 ft above base of Opal Member, Mount 

Head Formation, Zone IV; Vesiculophyllum, 

GSC loc. 60994, upper Banff Formation, Zone IIB?.

AL-31. Upper Exshaw Creek, head of creek, 51°09'N., 

115°13'W., Mount Head Formation (see Macqueen and 

Bamber, 1968). Acrocyathus, GSC Iocs. 58160 to 58164, 

36 ft below top of Salter Member to 72 ft above base of 

Loomis Member, Zone IIIB, IIIC, HID; Ekvasophyllum,

GSC Iocs. 58165, 58171, 82 ft above base of Loomis 

Member to 53 ft above base of Marston Member, 

Zone HID; Syringopora, GSC loc. 58163, 44.5 ft above 

base of Loomis Member, Zone HID.

AL-32. Upper Jura Creek, 51°08'N., IIS^IW. (see Macqueen 

and Bamber, 1967). Siphonodendron, Vesiculophyllum, 

GSC loc. 74892, top 10 ft of Banff Formation, Zone IIB; 

Michelinia, GSC loc. 74889, 543 ft below top of Banff Formation, Zone HA?; Syringopora, GSC loc. 74888, 599 ft 

below top of Banff Formation, Zone HA?.

AL-33. Grotto Creek, 51°06'N., 115°19'W. (see Macqueen and 

Bamber, 1967, 1968). Acrocyathus, GSC loc. 62064, lower 

Carnarvon Member, Mount Head Formation, Zone IV; 

Ekvasophyllum, GSC Iocs. 62045, 62046, 62056, 62057, 

62060, 62061, 62063,47 ft below top of Livingstone Formation to 74 ft above base of Opal Member, Mount Head 

Formation, Zone IIIA? to IV; Faberophyllum, 

GSC loc. 62063, 49 to 74 ft above base of Opal Member, 

Mount Head Formation, Zone IV; Syringopora, 

GSC loc. 62062, 23 ft below top of Loomis Member, 

Mount Head Formation, Zone HID; ?Zaphriphyllum, 

GSC loc. 62046, 13ft above base of' Salter Member, 

Mount Head Formation, Zone IIIA or IIIB.

AL-34. Pigeon Mountain, along south side of Trans-Canada 

Highway, 51°02'N., 115°13'W. (see Macqueen and 

Bamber, 1967). Sychnoelasma, GSC loc. 69573, 107 ft 

below top of Banff Formation, Zone HA; Syringopora, 

GSC 69572, 199ft below top of Banff Formation, 

Zone IIB?.

AL-35. Lower Jura Creek, 51°05'N., U5°lOrW. (see Macqueen 

and Bamber, 1967). Stelechophyllum, GSC loc. 62109, 

50ft below top of Banff Formation, Zone IIB; Syringopora, GSC Iocs. 62103, 62106 to 62109, 50 to 555 ft 

below top of Banff Formation, Zones I?, IIB; 

Vesiculophyllum, GSC Iocs. 62109, 62110, 29 to 50 ft 

below top of Banff Formation, Zone IIB.

AL-36. Brewster Creek, 51 "OOTSL, 115 °36 rW., all measurements 

above base of Opal Member, Mount Head Formation (see 

Macqueen and Bamber, 1968). Ekvasophyllum, 

GSC Iocs. 68429, 68431, 68432, 68435, 68436 to 68438, 

68441, 25 to 430ft, Zones HID, IV; Faberophyllum, 

GSC Iocs. 68440 to 68442, 320 to 452ft, Zone IV; 

Siphonodendron, GSC loc. 68430, 40ft, Zone HID; 

Sciophyllum, GSC loc. 68439, 275 ft, Zone IV.

AL-37. Mount Lorette, south side of mountain, 50°58'30"N., 

115°07'W. (see Macqueen and Bamber, 1967). 

Acrocyathus, GSC loc. 58150, middle Loomis Member, 

Mount Head Formation, Zone IHC or HID; Syringopora, 

GSC loc. 58030, lower Livingstone Formation, Zone HB?.

AL-38. Mount McDougall, 1.25 mi north-northwest of peak, east 

of mouth of Evans-Thomas Creek, 50°55 rN., ll5°05 rW., 

Mount Head Formation (see Macqueen and Bamber, 

1968). Acrocyathus?, GSC loc. 58140, 233 to 234 ft above 

base of Opal Member, Zone IV; ?Clisiophyllum, 

GSC loc. 58145, uppermost Carnarvon Member, Zone IV; 

Ekvasophyllum and Syringopora, GSC Iocs. 58122, 

58126, 0.5 to 60 ft below top of Loomis Member, 

Zone HID; Siphonodendron and Stelechophyllum, 

GSC loc. 58126, 0.5 ft below top of Loomis Member, 

Zone HID.

AL-39. Canyon Creek, east and west flanks of "Moose Dome," 

50°54'N., 114°53'W. (see Macqueen and Bamber, 1967; 

Bamber, Macqueen, and Ollerenshaw, 1981). 

Siphonodendron, GSC loc. 66346, 117 ft below top of
52 CORAL ZONATION, MISSISSIPPIAN SYSTEM, WESTERN INTERIOR PROVINCE

Turner Valley Formation, Zone IIIA; Lophophyllum?, uppermost Banff Formation, Zone IIB; Stelechophyllum, 

10 ft below top of Banff Formation to 3 ft above base of 

Pekisko Formation, Zone IIA; Sychnoelasma, 

GSC loc. 66336, lower Shunda Formation, Zone IIB; Syringopora, GSC Iocs. 66326, 66328, 66329, 66336, 66342, 

upper Banff Formation to lower Turner Valley Formation, Zones IIB, IIIA; Vesiculophyllum, upper Banff Formation to lower Turner Valley Formation, Zones IIB, 

IIIA; Zaphriphyllum, lower Turner Valley Formation, 

Zone IIIA.

AL-40. Grizzly Creek, 50°46'N., 115°07'30"W. (see Macqueen 

and Bamber, 1968). fCanadiphyllum, GSC loc. 58039, 

42 ft above base of Salter Member, Mount Head Formation, Zone IIIB?; Siphonophyllia, GSC loc. 58083, 34 ft 

above base of Etherington Formation, Zone V; 

Faberophyllum, GSC loc. 58078,326 ft above base of Opal 

Member, Mount Head Formation, Zone IV; Petalaxis, 

GSC loc. 58044, 205 ft above base of Loomis Member, 

Mount Head Formation, Zone HID; Syringopora, 

GSC Iocs. 58039, 58080, 42 ft above base of Salter 

Member to 21 ft above base of Carnarvon Member, 

Mount Head Formation, Zones IIIB?, IV.

AL-41. Middle Mountain, 2.5 mi west of Little Elbow River, 

50°44'N., 115°03'W., Mount Head Formation unless 

otherwise indicated (see Macqueen and Bamber, 1967, 

1968). Acrocyathus, GSC Iocs. 66283, 66289, 66303, 31 ft 

below top of Loomis Member to 109 ft above base of Opal 

Member, Zones HID, IV?; Ekvasophyllum, 

GSC Iocs. 66282 to 66284, 66299, 66302, 66303, 38 ft 

below top of Livingstone Formation to 13 ft below top 

of Loomis Member, Zones IIIB?, HID; Siphonodendron, 

GSC Iocs. 66282, 66283, 16 to 29 ft below top of Loomis 

Member, Zone HID; Syringopora GSC Iocs. 66282, 66289, 

25 ft below top of Loomis Member to 109 ft above base 

of Opal Member, Zones HID, IV?.

AL-42. Rover Mountain-Beaver Ridge, composite section approximately 1 mi west of Little Elbow River, 50°44'N., 

115°01'W., Mount Head Formation (see Macqueen and 

Bamber, 1967,1968). Acrocyathus and Siphonodendron, 

GSC loc. 66268, 14 ft below top of Loomis Member, 

Zone HID; Ekvasophyllum, GSC Iocs. 66260, 66261, 

66264, 27 ft above base of Baril Member to 47 ft above 

base of Loomis Member, Zones IIIA?, IIIC?; Syringopora, 

GSC Iocs. 66261, 66264, 66268, top bed of Baril Member 

to 14 ft below top of Loomis Member, Zones IIIA?, IIIC?, 

HID.

AL-43. Mount Hood, 50°44'N., 115°04'W. (see Macqueen and 

Bamber, 1967, 1968). Acrocyathus, GSC Iocs. 66235, 

66245, 66249, 66251, Mount Head Formation, 50 ft below 

top of Salter Member to 18 ft below top of Opal Member, 

Zones IIIB, HID, IV; ?Canadiphyllum, GSC loc. 66252, 

83 to 87 ft below top of Livingstone Formation, 

Zone IIIA?; Ekvasophyllum, GSC loc. 66250, 23 to 33 ft 

below top of Loomis Member, Mount Head Formation, 

Zone HID; Siphonodendron, GSC loc. 66255, 65 ft above 

base of Livingstone Formation, Zone IIB; Syringopora, 

GSC Iocs. 66232, 66249, 66250, Mount Head Formation, 

33 ft below top of Loomis Member to 7 ft below top of Opal 

Member, Zones HID, IV.

AL-44. King Creek, measured in canyon approximately 1 mi 

upstream from Kananaskis Road, 50°43'N., H5°Q6 rW., 

all measurements above base of Opal Member, Mount 

Head Formation, unless otherwise indicated (see Macqueen and Bamber, 1968). Acrocyathus, GSC Iocs. 57997,

58000, 76 to 140 ft, Zone HID; Ekvasophyllum, 

GSC Iocs. 57998, 58002 to 58005, 84 to 307 ft, Zones HID, 

IV; Faberophyllum, GSC loc. 58007, 470ft, Zone IV; 

?Koninckophyllum, GSC loc. 58001, 168 ft, Zone HID; 

Siphonodendron, GSC Iocs. 57998, 58001, 58005, 84 to 

307 ft, Zone IED; Schoenophyllum, GSC loc. 58023,238 ft 

above base of Etherington Formation, Zone V; 

Stelechophyllum, GSC loc. 58001, 168 ft, Zone HID; Syringopora, GSC loc. 57999, 96 ft, Zone HID. 

AL-45. Opal Range, west slope, on east side of KananaskisColeman Highway, 50°43'N., 115 "OSW., lower Etherington Formation, Zone V (GSC internal report; see Scott, 

1964, Mount Wintour). Siphonophyllia, GSC loc. 53423; 

Schoenophyllum, GSC loc. 53422.

AL-46. Elpoca Creek, approximately 0.75 mi upstream from 

Kananaskis Forestry Road, 50°39'N., 115°04'W., all 

measurements above base of Opal Member, Mount Head 

Formation, unless otherwise indicated (see Macqueen and 

Bamber, 1968). Acrocyathus, GSC Iocs. 58095, 58098, 

58101, 58105, 52 to 313ft, Zones HID, IV; Amplexizaphrentis, GSC loc. 58118, 9 ft above base of Etherington Formation, Zone V?; Ekvasophyllum, 

GSC Iocs. 58093, 58095, 58100 to 58102, 58108, 58109, 

26 to 384.5 ft, Zones HID, IV; Faberophyllum, 

GSC Iocs. 58102, 58103, 58110, 58111, 244 to 515.5 ft, 

Zone IV; Siphonodendron, GSC Iocs. 58096, 58099, 54 to 

104 ft, Zones HID, IV; Petalaxis, GSC loc. 58094, 38 ft, 

Zone HID; Stelechophyllum, GSC Iocs. 58099, 58101, 

58104, 101.5 to 204ft, Zone IV; Syringopora, 

GSC Iocs. 58095, 58099, 58101, 58103, 58104, 58107, 52 

to 325ft, Zones HID, IV; "Thysanophyllum," 

GSC loc. 58109, 383.5 to 384.5 ft, Zone IV.

AL-47. Elbow Lake, about 0.5 mi east of Kananaskis Road, 

50°38'N., 115"OOW., Mount Head Formation, unless 

otherwise indicated (see Macqueen and Bamber, 1968). 

Acrocyathus, GSC Iocs. 66186, 66200, 66204, 66208, 95 

to 530 ft above base of Opal Member, Zones HID, IV; 

Amplexizaphrentis, GSC loc. 66184, 13 ft above base of 

Etherington Formation, Zone V?; ?Clisiophyllum, 

GSC loc. 66185, 9 to 12 ft above base of Carnarvon 

Member, Zone IV; Ekvasophyllum, GSC Iocs. 66200, 

66204, 66206, to 66208, 95 to 212 ft above base of Opal 

Member, Zones HID, IV; Siphonodendron, 

GSC Iocs. 66204, 66206, 78 to 147 ft above base of Opal 

Member, Zone HID; Stelechophyllum, GSC loc. 66200, 

212 ft above base of Opal Member, Zone IV; Syringopora, 

GSC Iocs. 66185, 66186, 5 ft below top of Opal Member 

to 12 ft above base of Carnarvon Member, Zone IV; 

Schoenophyllum, GSC loc. 66179, 110 to 111 ft above 

base of Etherington Formation, Zone V.

AL-48. Boulder Creek, 1.5 mi south of Mt. Burns, 50°37'N., 

114°52'W., Mt. Head Formation (see Macqueen and 

Bamber, 1967,1968). Acrocyathus?, GSC loc. 66325, upper Carnarvon Member, Zone IV; ?Canadiphyllum, 

GSC loc. 66314, upper 4 ft of Baril Member, Zone IIIA?; 

Ekvasophyllum, GSC Iocs. 66309, 66313, 66317, 19 ft 

below top of Wileman Member to 18 ft below top of 

Loomis Member, Zones IIIA?, HID; Siphonodendron, 

GSC loc. 66317,18 to 21 ft below top of Loomis Member, 

Zone HID; Syringopora, GSC Iocs. 66312, 66317, 66325, 

44 ft below top of Baril Member to upper Carnarvon 

Member, Zones IIIA?, IIIC?, IV; Zaphriphyllum, 

GSC loc. 66312, 44 to 46 ft below top of Baril Member, 

Zone IIIA?.
LOCALITY REGISTER 53

AL-49. Storm Creek, cirque at head of creek, 50°37'N., 

114 0 59 rW. (GSC internal reports; see Scott, 1964; Macqueen and Bamber, 1968). Amplexizaphrentis, 

GSC loc. 57934,15 ft above base of Etherington Formation, Zone V?; Acrocyathus, GSC Iocs. 57938, 57947, 

57953, 57968, 57971,57973, 57976, 57977, 57987,74876, 

74877, 74878, 74879, 10 to 524.5 ft above base of Opal 

Member, Mount Head Formation, Zones HID, IV; 

Siphonophyllia, GSC loc. 53475, lower Etherington Formation, Zone V; Clisiophyllum, GSC loc. 53462, Carnarvon Member?, Mount Head Formation, Zone IV; 

Ekvasophyllum, GSC Iocs. 57943, 57955, 57972, 57977, 

57983, 57985, 57987, 57988, 57992, 57993, 10 ft below 

top of Loomis Member to 500 ft above base of Opal 

Member, Mount Head Formation, Zones HID, IV; 

Faberophyllum, GSC Iocs. 57939 to 57942, 57945, 57947, 

57949, 57951, 57953, 57954, 390 to 519.5 ft above base 

of Opal Member, Mount Head Formation, Zone IV; 

Koninckophyllum, GSC loc. 53462, Carnarvon Member?, 

Mount Head Formation, Zone IV; Petalaxis, 

GSC loc. 57985,19 ft above base of Opal Member, Mount 

Head Formation, Zone HID; Pleurosipkonella, uppermost 

Etherington Formation, Zone VI; Syringopora, 

GSC Iocs. 57933, 57934, 57937, 57938, 57940, 57944, 

57965, 57979, 57982, 57988, 57992, 57993, 45 ft below 

top of Loomis Member to 531 ft above base of Opal 

Member, Mount Head Formation, Zones HID, IV; 

"Thysanophyllum," GSC Iocs. 57969, 57974,166 to 180 ft 

above base of Opal Member, Mount Head Formation, 

Zone HID or lower IV.

AL-50. South Misty Range, 2 mi north-northwest of Mist Mountain, 50°34.5'N., 114 0 56 rW., Mount Head Formation, 

unless otherwise indicated (see Macqueen and Bamber, 

1968). Amplexizaphrentis, GSC loc. 57872, lower Etherington Formation, Zone V?; Ekvasophyllum, 

GSC Iocs. 57905 to 57907, 57911, lllft below top of 

Loomis Member to 6 ft above base of Opal Member, 

Zone HID; ?Faberophyllum, GSC loc. 57874, 94 ft below 

top of Opal Member, Zone IV; Siphonodendron, 

GSC Iocs. 57902, 57904, 27 to 102 ft above base of Opal 

Member, Zone HID; Petalaxis, GSC loc. 57905, 6 ft above 

base of Opal Member, Zone HID; ?Syringopora, 

GSC loc. 57909, 31 ft below top of Loomis Member, 

Zone HID.

AL-51. Picklejar Lakes, north side of valley, at head of Picklejar Creek, 50°32'N., 114 0 46 rW. (see Macqueen and 

Bamber, 1967,1968). Ekvasophyllum, GSC Iocs. 57839, 

57846,15 ft below top of Baril Member to 16 ft above base 

of Marston Member, Mount Head Formation, 

Zones IHA?, HID; Siphonodendron, GSC loc. 57846,16 ft 

above base of Marston Member, Mount Head Formation, 

Zone HID; ?Zaphriphyllum, GSC loc. 57856,19 ft below 

top of Livingstone Formation, Zone IIIA?.

AL-52. Lantern Creek headwaters, Highwood Range, 

50°31'30"N., 114°47'30"W. (GSC internal report; see 

Scott, 1964). Amplexizaphrentis, GSC Iocs. 53455, 53456, 

53457,170 to 200 ft below top of Etherington Formation, 

Zone V?.

AL-53. Flat Creek, 50"SOW., 114°40'W. (see Douglas, 1958; Macqueen and Bamber, 1967, 1968). Ekvasophyllum, 

GSC loc. 57774, upper Livingstone Formation, 

Zone HIA?; Siphonodendron, GSC loc. 57773, upper Livingstone Formation, Zone IIIA?; Syringopora, 

GSC Iocs. 57773, 69581, 770 ft above base of Banff Formation to upper part of Livingstone Formation,

Zones IIB?, IIIA?; Vesiculophyllum, GSC loc. 69589, 

990 ft above base of Livingstone Formation, Zone IIIA?; 

Zaphriphyllum, GSC loc. 57773, upper Livingstone Formation, Zone IIIA?.

AL-54. Highwood River, north side of valley, 50°24'N., 

114°37'W. (see Douglas, 1958; Macqueen and Bamber, 

1968). Ekvasophyllum, GSC loc. 57751, 48 ft below top 

of Marston Member, Mount Head Formation, Zone HID 

or lower IV.

AL-55. Palliser Range, east side of Cascade River, approximately 

10 mi above its mouth, 50°24'N.,115°37'W. (GSC internal report; see Scott, 1964). Amplexizaphrentis, 

GSC loc. 53468, basal bed of Etherington Formation, 

Zone V?.

AL-56. Mount Burke, 50°17'N., 114 0 32 rW. (see Macqueen and 

Bamber, 1968). Ekvasophyllum, ?Stelechophyllum, Syringopora, GSC loc. 57803,17 ft below top of Livingstone 

Formation, Zone IIIA?.

AL-57. Mount Inglismaldie, ridgetop southwest of peak, 

50°14'30"N., 115°26'30"W. (GSC internal report; see 

Scott, 1964). Amplexizaphrentis, GSC loc. 53467, upper 

Mount Head Formation (Carnarvon Member?), Zone IV?; 

Actinocyathus, GSC loc. 53468, upper? Etherington Formation, Zone V.

AL-58. Plateau Mountain, south end of mountain, 50°11'30"N., 

114°32'W. (see Macqueen and Bamber, 1968). 

Ekvasophyllum, GSC Iocs. 57815, 57816, 57839, 42 ft 

below top of Livingstone Formation to 15 ft below top 

of Baril Member, Mount Head Formation, Zone IIIA; 

Siphonodendron, GSC loc. 57818,12 ft below top of Livingstone Formation, Zone IIIA; Lophophyllum? and 

Stelechophyllum, GSC loc. 57819, 9 ft below top of Livingstone Formation, Zone IIIA.

AL-59. Oldman River Gap, west side of gap, north side of river, 

Livingstone Range, 49°52'30"N., 114°22'30"W. (GSC internal report; see Scott, 1964). Amplexizaphrentis, 

GSC loc. 55604, 78 ft below top of Etherington Formation, Zone V?.

AL-60. Alexander Creek headwaters, Highrock Range, 

49°50'30"N., 114°41 rW. (GSC internal report; see Scott, 

1964). Amplexizaphrentis, GSC loc. 51584, lower Etherington Formation, Zone V?.

AL-61. Daisy Creek headwaters, west side of Livingstone Range, 

49°46'N., 114°23'W. (GSC internal report; see Scott, 

1964). Amplexizaphrentis, GSC loc. 48317, 172 ft above 

base of Etherington Formation, Zone V.

AL-62. Turtle Mountain, 49°36'N., 114°25'W. (GSC internal 

report; see Price, 1962). Siphonodendron, GSC loc. 69590, 

153 to 158 ft above base of Livingstone Formation, 

Zone IIIA; Lophophyllum?, GSC loc. 47489, 19 ft above 

base of Baril Member, Mount Head Formation, 

Zone IIIA?.

AL-63. Mt. Ptolemy, northwest ridge top, 49°34'30"N., 

114°40'W. (GSC internal report; see Scott, 1964). 

Schoenophyllum, GSC Iocs. 48881, 48886, lower Etherington Formation, Zone V.

AL-64. West side Mt. Ptolemy, approximately 2 mi N.45 °W. of 

main peak, 49°34'N., 114°39'W. (GSC internal report; 

see Scott, 1964). Siphonophyllia, Actinocyathus, 

GSC loc. 49388,180 ft below top of Etherington Formation, Zone V.

AL-65. Amoco G-l Equity 11-27-32-24(4), subsurface, Zone HB 

(GSC internal report; see Boreski, 1978). 

Stelechophyllum, GSC loc. C-46466, 4.5 ft above base of 

Pekisko Formation; Sychnoelasma, GSC loc. C-46463,
54 CORAL ZONATION, MISSISSIPPIAN SYSTEM, WESTERN INTERIOR PROVINCE

37 ft above base of Pekisko Formation; Syringopora, 

GSC Iocs. C-46465, C-46470, C-46472,10 to 20 ft below 

top of Banff; Vesiculophyllum, GSC Iocs. C-46462 to 

C-46464, 12 to 87 ft above base of Pekisko Formation.

AL-66. Amoco E-l Equity 6-21-32-24(4), subsurface, all 

measurements above base of Pekisko Formation unless 

otherwise indicated, Zone IIB (GSC internal report; see 

Boreski, 1978). Amplexizaphrentis, GSC loc. C-46529, 

16ft; Lophophyllum?, GSC loc. C-46517, 71ft; 

Stelechophyllum, GSC loc. C-46528,19 ft; Sychnoelasma, 

GSC Iocs. C-46520, C-46523 to C-46526, C-46530, 10 

to 64 ft; Vesiculophyllum, GSC Iocs. C-46515, C-46516, 

C-46518, C-46519, C-46521, C-46525, C-46527, 

C-46528, 19 ft above base of Pekisko Formation to 1 ft 

above base of Shunda Formation.

AL-67. Pacific Twining 10-23-32-24(4), subsurface, all 

measurements above base of Pekisko Formation, 

Zone IIB (GAS internal report; see Boreski, 1978). 

Stelechophyllum, GSC Iocs. C-46473, C-46478, 11.4 to 

33 ft; Sychnoelasma, GSC Iocs. C-46477, C-46479, 2.5 

to 35 ft; Vesiculophyllum, GSC Iocs. C-46474, C-46475, 

C-46477, 30 to 35 ft.

AL-68. Pan Am CPR-1 Equity 14-7-32-23(4), subsurface, 

Zone IIB? (GSC internal report; see Boreski, 1978). Syringopora, GSC Iocs. C-46482, C-46484, 3.5 to 61ft 

above base of Pekisko Formation; Vesiculophyllum, 

GSC loc. C-46483, 35.5 ft above base of Pekisko 

Formation.

AL-69. Pacific Twining 14-6-32-24(4), subsurface, all 

measurements above base of Pekisko Formation, 

Zone IIB (GSC internal report; see Boreski, 1978). 

Amplexizaphrentis, GSC Iocs. C-46512, C-46513, 32 to 

33ft; Aulostylus, GSC loc. C-46506, 58ft; Stelechophyllum, GSC loc. C-46509, 38 ft; Sychnoelasma, 

GSC Iocs. C-46504, C-46508, C-46510 to C-46512, 

C-46514, 31 to 117 ft; Vesiculophyllum, 

GSC Iocs. C-46502, C-46503, C-46505, 74 to 140 ft.

AL-70. MicMac Twining 6-4-32-24(4), subsurface, all 

measurements above base of Pekisko Formation, 

Zone IIB (GSC internal report; see Boreski, 1978). 

Amplexizaphrentis, GSC Iocs. C-46488, C-46489, 

C-46491, C-46499, C-46500, C-46501, 9 to 93ft; 

Aulostylus, GSC loc. C-46494, 79 ft; Sychnoelasma, 

GSC Iocs. 46491, C-46498, C-46499, 35 to 89 ft; Syringopora, GSC loc. C-46484, 3.5 ft; Vesiculophyllum 

GSC Iocs. 46486 to C-46488, C-46490, C-46492 to 

C-46496, 57 to 99 ft.

AL-71. Mon Santo Twining 6-35-31-24(4), subsurface (GSC internal report; see Boreski, 1978). Sychnoelasma, 

GSC loc. C-46577, 46 ft above base of Pekisko Formation, Zone IIB.

AL-72. Mobil Twining 6-20-31-24W4, subsurface, all 

measurements above base of Pekisko Formation unless 

otherwise indicated, Zone IIB (GSC internal report; see 

Boreski, 1978). Amplexizaphrentis, GSC Iocs. C-46535, 

C-46549, C-46555, C-46557, C-46559, C-46569, 

C-46573 to C-46576, 1 ft above base to top of Pekisko 

Formation; Aulostylus, GSC loc. C-46565, 38ft; 

Lophophyllum?, GSC loc. C-46547, 67 ft; Michelinia, 

GSC Iocs. C-46545, C-46560, 56.5 to 69 ft; Stelechophyllum, GSC loc. C-46571,21.5 ft; Sychnoelasma, 

GSC Iocs. C-46542 to C-46544, C-46554, C-46560, 

C-46563, C-46565 to C-46568, C-46570, C-46572, 21 

to 72ft; Syringopora, GSC Iocs. C-46533, C-46534, 

C-46537, C-46538, C-46560, C-46572, 21 ft above base

of Pekisko Formation to 2 ft above base of Shunda Formation; Vesiculophyllum, GSC Iocs. C-46539 to C-46541, 

C-46543 to C-46545, C-46547 to C-46552, C-46554, 

C-46556, C-46561, C-46562, C-46564, C-46568, 24 to 

102 ft.

AL-73. Andex et al. Twining 5-23-31-24(4), subsurface, 

Zone IIB (GSC internal report; see Boreski, 1978). 

Stelechophyllum, GSC Iocs. C-46458, C-46460, C-46461, 

7.5 ft below top of Banff Formation to 1.5 ft above base 

of Pekisko Formation; Sychnoelasma, GSC Iocs. C-46455, 

C-46456, C-46458,1.5 to 36 ft above base of Pekisko Formation; Syringopora, GSC Iocs. C-46457, C-46459,1 ft 

below top of Banff Formation to 2.5 ft above base of 

Pekisko Formation.

AL-74. Andex et al. Twining 16-16-31-24(4), subsurface (GSC 

internal report; see Boreski, 1978). Amplexizaphrentis, 

GSC loc. C-46452, 19 ft below top of Banff Formation, 

Zone IIA or IIB.

SOUTHEASTERN BRITISH COLUMBIA

SEBC-1. Mount Thompson, 1.6 mi S. 60°W. from peak, 50°43TST., 

117°39'30"W., all measurements above base of Milford 

Group (GSC internal report). Amplexizaphrentis, 

GSC loc. C-236, talus from lower 15 ft, Zone HID or IV; 

Ekvasophyllum, GSC Iocs. C-237, C-241, C-242, C-243, 

C-244, 10 to 35 ft, Zone HID; ?Faberophyllum, 

GSC loc. 68709, talus from lower 100ft, Zone IV?; 

Siphonodendron, GSC Iocs. 68709, C-237, C-246, C-248, 

10 to 80 ft, talus from lower 100 ft, Zones HID, IV, V; 

?"Pseudodorlodotia", GSC loc. C-248, 80ft, Zone IV; 

Stelechophyllum?, GSC Iocs. C-240, C-242, C-245;/ 

C-249, 30 to 40 ft, Zone HID; Syringopora, 

GSC loc. C-237, 10 ft, Zone HID.

SEBC-2. Mount Storelk, southwest corner, Elk Range, 

50°32'30"N, IM'SgW. (GSC internal report; see Scott, 

1964). Amplexizaphrentis, GSC Iocs. 53401,53406, 53426, 

190 ft below top of Mt. Head Formation to upper Ether - 

ington Formation, Zones IV, V; Siphonophyllia, GSC Iocs. 

53409, 53410, 110 to 153 ft above base of Etherington 

Formation, Zone V; ?Clisiophyllum, GSC loc. 53407, 75 

ft below top of Mount Head Formation, Zone IV; ?Actinocyathus, GSC loc. 53408, 355 ft above base of Etherington Formation, Zone V; Schoenophyllum, 

GSC loc. 53402, lower Etherington Formation?, Zone V.

SEBC-3. Mount Loomis, south side, Elk Range, 50°22'30"N, 

114°55'W. (GSC internal report). Amplexizaphrentis, 

GSC loc. 53438,144 ft below top of Etherington Formation, Zone V; Actinocyathus, GSC loc. 53435, 313 ft below 

top of Etherington Formation, Zone V.

SEBC-4. Ewin Creek, ridgetop 2.2 mi west of Beehive Mountain, 

50°04'30"N, 114°42'30"W., all measurements below top 

of Etherington Formation, Zone V (GSC internal report; 

see Scott, 1964). Amplexizaphrentis, GSC Iocs. 48333, 

48347, 490 to 523 ft; Siphonodendron, GSC loc. 48334, 

between 540 and 570 ft; Pleurosiphonella, GSC Iocs. 

48334, 48342, between 540 and 570 ft; Schoenophyllum, 

GSC Iocs. 48338, 48342, 540 to 612 ft.

SEBC-5. Tornado Mountain, 2.75 mi northwest of mountain, north 

side of Line Creek, Highrock Range, 49°59'30"N, 

114°42'W., Mount Head Formation (GSC internal report, 

see Price, 1962). Acrocyathus, GSC loc. 40700,12 ft above 

base of Carnarvon Member, Zone IV; ?Acrocyathus and
LOCALITY REGISTER 55

?Ekvasophyllum, GSC loc. 40716,133.5 ft above base of 

Marston Member, Zone HID (upper) or IV (lower); 

Amplexizaphrentis, GSC loc. 40715,170 ft above base of 

Carnarvon Member, Zone IV; Faberophyllum, GSC Iocs. 

40697, 40703, 40711, 71 to 123 ft above base of Carnarvon Member, Zone IV; Siphonodendron, GSC loc. 40696, 

343 ft above base of Loomis Member, Zone HID.

SEBC-6. Mount Hosmer, northeast spur, 49°43'N., H5°59 rW. 

(GSC internal report; see Scott, 1964). Amplexizaphrentis, GSC Iocs. 49379, 49380, 49385, lower and upper 

Etherington Formation, Zone V; Siphonophyllia, 

GSC loc. 49385, lower Etherington, Zone V; Actinocyathus, GSC loc. 49354, upper Etherington, Zone V; 

Michelinia, GSC Iocs. 49370,49371,49384, lower Etherington, Zone V; Pleurosiphonella, GSC Iocs. 49364,49365, 

49374, lower Etherington, Zone V; ?Syringopora, 

GSC loc. 49386, upper Mount Head Formation (Carnarvon Member?), Zone IV.

SEBC-7. Mount Broadwood, east slope of eastern peak, 49°18TST., 

114°58'W. (GSC internal report, see Price, 1962). 

Acrocyathus, GSC Iocs. 40698,40702, 297.5 ft above base 

of Loomis Member to 36.5 ft above base of Marston 

Member, Mount Head Formation, Zone HID; ?Amplexizaphrentis, GSC loc. 40687, 273 ft above base of Etherington Formation, Zone V?; Siphonophyllia, 

GSC loc. 40688,182 ft above base of Etherington, Zone V; 

Ekvasophyllum, GSC Iocs. 40694, 40698, 36.5 to 106 ft 

above base of Marston Member, Mount Head Formation, 

Zone HID or IVdower); Schoenophyllum, GSC loc. 40688, 

182 ft above base of Etherington Formation, Zone V.

SEBC-8. Mount Broadwood, southeast corner, 49°17'30"N., 

114°58'W. (GSC internal report). Amplexizaphrentis, 

GSC loc. 49362, 275 ft below top of Etherington Formation, Zone V?; Siphonophyllia, GSC Iocs. 49355, 49391, 

358 to 360ft below top of Etherington, Zone V; 

Ekvasophyllum?, GSC loc. 49361, upper Mount Head Formation (Carnarvon Member?), Zone IV?; Faberophyllum, 

GSC Iocs. 49359, 49360,49369, upper Mount Head (Carnarvon Member?), Zone IV; Schoenophyllum, GSC Iocs. 

49358, 49363, 55586, 340 to 380 ft below top of Etherington Formation, Zone V.

SASKATCHEWAN

S-l. Husky-Mic Mac-Canadian Superior Bellegarde No. 1, 

Lsd 16-19-6-30W1, Tilston beds, Zone IIB? (see Brindle, 1960). Vesiculophyllum, 3,249 ft below surface.

S-2. Canadian Export Gas C. P. Bellegarde No. 5-27, Lsd 

5-27-6-31W1, Tilston beds, Zone EB, (see Brindle, 1960), 

all measurements in feet below surface. Amplexizaphrentis, 3,393.25; Sychnoelasma, 3,385; Vesiculophyllum, 

3,381.75, 3,382, 3,396.75, 3,403.

S-3. McCarty and Coleman Mobil Crown No. 16-28, Lsd 

16-28-6-32W1, Tilston beds, Zone IIB (see Brindle, 

1960), all measurements in feet below surface. 

Michelinia, 3,620; Sychnoelasma, 3,613.5, 3,615.5; Syringopora, 3,623.25;Vesiculophyllum,3,612.75, 3,615.5, 

3,623, 3,636, 3,636.25, 3,643.

S-4. McCarty and Coleman Pederson No. 4-35, Lsd 

4-35-7-33W1, Tilston beds, Zone EB (see Brindle, 1960), 

all measurements in feet below surface. Amplexizaphrentis, 3,630, 3,632.75; Stelechophyllum, 3,640; 

Sychnoelasma, 3,630; Syringopora, 3,636, 3,637; 

Vesiculophyllum, 3,684.

S-5. White Rose-Canadian Superior-Souris Cayuga No. 

14-31, Lsd 14-31-3-30W1, Frobisher-Alida beds, 

Zone IIB (see Brindle, 1960), all measurements in feet 

below surface. Amplexizaphrentis, 3,464, 3,474.5; 

Sychnoelasma, 3,463.75, 3,466.5, 3,481; Vesiculophyllum, 

3,471, 3,473, 3,480.75, 3,482, 3,486.5.

S-6. Imperial Rosebank No. 16-1M, Lsd 16-1-4-32W1, 

Frobisher-Alida beds, Zone IIB (see Brindle, 1960), all 

measurements in feet below subsurface. Amplexizaphrentis, 3,712, 3,717; Sychnoelasma, 3,716.5; 

Vesiculophyllum, 3,706, 3,725, 3,744.5. 

S-7. Imperial Seymour No. 4-19-3-31, Lsd 4-19-3-31W1 (see 

Brindle, 1960). Vesiculophyllum, 3,774.75 ft below surface, Frobisher-Alida beds, Zone IIB?.

S-8. Imperial-Canadian Superior Carnduff No. 3-27-2-33, 

Lsd 3-27-2-33W1, Frobisher-Alida beds, Zone IIB (see 

Brindle, 1960), all measurements in feet below surface. 

Sychnoelasma, 4,216, 4,216.5; Vesiculophyllum, 4,215.25, 

4,216, 4,224, 4,226.5.

S-9. Imperial-Royalite-Hudson's Bay Auburnton No. 

l_2-5-l, Lsd 1-2-5-1W2, Zone IIB (see Brindle, 1960), 

all measurements in feet below surface. Amplexizaphrentis, 4,334.5, 4,343.25, 4,389.25, Tilston beds; 

Lophophyllum?, 4,332.25,, 4,336, Tilston beds; 

Michelinia, 4,334, 4,338.5, Tilston beds; Sychnoelasma, 

3,957, Frobisher-Alida beds, 4,332, 4,341.5, 4,380, 

4,389.25, Tilston beds; Syringopora, 4,346.5, Tilston beds; 

Vesiculophyllum, 4,329,4,333,4,340,4,343, Tilston beds.

S-10. Imperial-Tidewater Carlyle Crown No. 1, Lsd 

16-23-7-3W2, Zone IIB? (see Brindle, 1960), all 

measurements in feet below surface. Amplexizaphrentis, 

3,891.75, 4,604.5, Souris Valley and Frobisher-Alida 

beds; Vesiculophyllum, 3,876.5, 3,885, 3,889.5, 

Frobisher-Alida beds.

S-ll. British American-Canadian Devonian Quinn No. 1-34, 

Lsd 1-34-4-3W2 (see Brindle, 1960). Vesiculophyllum, 

4,610 ft below surface, Midale beds, Zone IIB or IIIA.

S-12. British American-Union Pickard No. 16-27, Lsd 

16-27-3-4W2 (see Brindle, 1960). Amplexizaphrentis, 

4,761, 4,763.25, 4,763.5, 4,764, and 4,765 ft below surface, Frobisher-Alida beds, Zone IIB?.

S-13. Imperial Dalesboro No. 4-9M-6-3, Lsd 4-9-6-3W2 (see 

Brindle, 1960). Vesiculophyllum, 4,159 ft below surface, 

Frobisher-Alida beds, Zone IIB?.

S-14. California Standard Frobisher No. 5-21, Lsd 

5-21-2-4W2 (see Brindle, 1960). Amplexizaphrentis, 

?Vesiculophyllum, 5,270ft below surface, FrobisherAlida beds, Zone IIB?.

S-15. Tidewater-Imperial Lampman Crown No. 1, Lsd 

5-28-6-6W2 (see Brindle, 1960). Sychnoelasma, 

4,321.5 ft below surface, Frobisher-Alida beds, Zone IIB.

S-16. Sohio Stoughton No. 4-13, Lsd 13-4-9-8W2, FrobisherAlida beds, Zone IIB (see Brindle, 1960), all 

measurements in feet below surface. Sychnoelasma, 

4,080.5, 4,083.5; Vesiculophyllum, 4,079.5, 4,080.5.

S-17. Western Leaseholds North Midale No. 3-5, Lsd 

3_5_7_10W2 (see Brindle, 1960). Syringopora, 4,438 ft 

below surface, Midale beds, Zone IIB or IIIA.

S-18. Shell Martin No. 13-5, Lsd 13-5-6-10W2 (see Brindle, 

1960). Syringopora, 4,677 ft below surface, Midale beds, 

Zone IIB?.
56 CORAL ZONATION, MISSISSIPPIAN SYSTEM, WESTERN INTERIOR PROVINCE

S-19. Gridoil Colgate No. 12-27, Lsd 12-27-5-14W2 (see Brindle, 1960). Vesiculophyllum, Syringopora, 5,008 ft below 

surface, Frobisher-Alida beds, Zone IIB?.

S-20. Central-Del Rio Ratcliffe No. 4-20, Lsd 4-20-1-15W2 

(see Brindle, 1960). Vesiculophyllum, 6,512,6,530.5, and 

6,543.5 ft below surface, Ratcliffe beds, Zone IIB?.

S-21. Socony-Central-Del Rio Ratcliffe No. 2, Lsd 

12-30-1-15W2 (see Brindle, 1960), all measurements in 

feet below surface. Amplexizaphrentis, 6,538, Midale 

beds, Zone IIB or ILIA; Syringopora, 6,486, 6,498.5, 6,506 

to 6,506.5, 6,526, Ratcliffe beds, 6,536, Midale beds, 

Zone IIB or IHA; Vesiculophyllum, 6,498.5,6,514, 6,521, 

Ratcliffe beds, 6,545.5, Midale beds, Zone IIB or IIIA.

S-22. White Rose-Compagnie Francaise-Mobil Lake Alma No. 

8-22, Lsd 8-22-2-17W2 (see Brindle, 1960). Syringopora, 

6,456 ft below surface, Ratcliffe beds, Zone IIB or IIIA.

LOCALITIES NOT REPRESENTED BY MEASURED SECTIONS

SOUTHWEST DISTRICT OF MACKENZIE

SWDM-1. Liard River, east bank, 5 mi above Flett Rapids, 

60°37'30"N., 123°30'30"W. (see Sutherland, 1958, fossil 

locality 34, p. 39). Amplexizaphrentis, Siphonodendron, 

GSC loc. 16603, Flett Formation, stratigraphic level 

unknown, Zone IIB?.

NORTHEASTERN BRITISH COLUMBIA

NEBC-1. Chlotopecta Creek, 58°28'N., 123°57W. Ekvasophyllum, 

GSC Iocs. 66691, 66692, upper 5 ft of Prophet Formation, 

Zone IIIC or HID (lower).

NEBC-2. Mount Brewster East, northeast ridge, 56°16'N., 

123° 12 W. (see Irish, 1963, p. 375). Ekvasophyllum, 

GSC loc. 46993, middle Prophet Formation, Zone IIIB or 

IIIC.

NEBC-3. Mount Brewster West, northwest ridge, 56°15'N., 

123°16'W. (see Irish, 1963, p. 375). Siphonodendron, 

Zaphriphyllum, GSC loc. 46883, middle Prophet Formation, Zone IIIB or IIIC.

NEBC-4. Mount Brewster, 3 mi north of Mount Brewster Peak, 

56°14TST., 123°10W. Ekvasophyllum, GSC loc. C-11706, 

top 10 ft of Prophet Formation, Zone HID or lower 

Zone IV.

NEBC-5. Black Bear Valley, 2 mi west of highest point on Black 

Bear Ridge, Se^STST., 123°06W. (see Irish, 1963, p. 376). 

Ekvasophyllum?, GSC loc. 36936, within upper 400 ft of 

Prophet Formation (member C?), Zone HID?.

NEBC-6. Fossil Point, north bank of Peace River, 56°01'N., 

123°08W. (see Sutherland, 1958, p. 39,94; Lambe, 1899, 

p. 220). Stelechophyllum?, upper Prophet Formation, 

member C?, Zone HID?.

NEBC-7. Narraway River, ridge 3 mi north of Narraway River, 

54°13'N., 120°23'W. Dorlodotia?, Sychnoelasma, 

Vesiculophyllum, GSC loc. C-7425, upper Shunda Formation, Zone IIB.

ALBERTA

AL-1. The Rajah, 2 mi south of Snake Indian River, 53°17'N., 

118 "SIW. (GSC internal report). Stelechophyllum, 

GSC loc. 42144, uppermost Banff Formation, Zone IIB.

AL-2. Rocky Pass, 52°51'N., 117°16'30"W. (see Kelly, 1942, 

p. 357). Stelechophyllum, probably from upper Pekisko 

Formation, Zone IIB.

AL-3. Mount Rae, northwest slope, 50°37'40"N., 114°59'35"W. 

(see McGugan, 1983). Palaeacis, talus specimen from upper Mount Head Formation or lower Etherington Formation, Zone IV or V.

SOUTHEASTERN BRITISH COLUMBIA

SEBC-1. Alexander Creek headwaters, 4 mi on bearing N.22 ° W. 

from Race Horse Mountain, all occurrences in upper 

Mount Head Formation (GSC internal report; see Price, 

1962). Ekvasophyllum, GSC loc. 42820, Zone HID (upper) 

or IV (lower); Faberophyllum, GSC loc. 42811, Zone IV; 

Siphonodendron, GSC Iocs. 42821, 42822, Zone HID; Syringopora, GSC loc. 42822, Zone HID.

SEBC-2. Playsoo Ridge, 3.5 mi on bearing 253° from mouth of 

Harvey Creek, MacDonald Range (GSC internal report; 

see Price, 1962). Siphonodendron, GSC loc. 36605, lower 

Etherington Formation, Zone V.

SEBC-3. Lodgepole Creek, ridge between headwaters of Lodgepole 

Creek (south fork) and Twentynine Mile Creek, Macdonald Range (GSC internal report; see Price, 1962). 

Ekvasophyllum, Acrocyathus?, GSC loc. 36602, 55 ft 

below top of Carnarvon Member, Mount Head Formation, Zone IV.

SEBC-4. Taylor Range, 12,000 ft on bearing 230 ° from Flathead 

Pass (GSC internal report; see Price, 1962). Siphonodendron; GSC loc. 47495, approximately 15 ft above base of 

Etherington Formation, Zone V.

SEBC-5. Wilson Creek, north side of creek, 6,500 ft downstream 

from mouth of Telford Creek (GSC internal report; see 

Price, 1962). Siphonophyllia, GSC loc. 42813, middle 

Etherington Formation, Zone V.

SEBC-6. Bighorn Creek, 8.15 mi from mouth of creek on bearing 

81°, MacDonald Range (GSC internal report; see Price, 

1962). Faberophyllum, GSC loc. 40699, approximately 

50 ft below top of Carnarvon Member, Mount Head Formation, Zone IV.

SEBC-7. Mount Darrah, west slope, Flathead Range, 49°28'30"N., 

114°37W. (see Nelson, 1960, p. 122). Schoenophyllum, 

160 ft above base of Etherington Formation, Zone V.

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PLATES 1-10

Contact photographs of the plates in this report are available, at cost, from the U.S. Geological 

Survey Library, Federal Center, Denver, CO 80225.
PLATE 1

FIGURES 1,2. Amplexocarinia sp. (p.17).

USGS loc. 13872-PC, Section M-9, Lodgepole Limestone, Montana, Zone 1C.

1. Side view of broken etched silicified mature specimen, x2, USNM 119780.

2. Calicular view of etched silicified neanic specimen, x2, USNM 119779. 

3,4. Amplexus sp. (p.17).

USGS loc. 13872, Section M-9, Lodgepole Limestone, Montana, Zone 1C.

3. Transverse thin section, x2, USNM 119771.

4. Longitudinal thin section, x2, USNM 119771. 

5,6. Acrocyathus arizelum (Crickmay) (p. 16).

GSC loc. 57938, Section AL-49, Mount Head Formation, Alberta, Zone IV.

5. Transverse thin section, x2, GSC type no. 67849a.

6. Longitudinal thin section, x2, GSC type no. 67849b. 

7,8. Amplexizaphrentis persimilis (Easton and Gutschick) (p. 16). 

Redwall Limestone, Arizona, Zone II.

7. Longitudinal thin section, x3, MNA G2.4075, Section A-18.

8. Transverse thin section, x3, holotype, MNA G2.4078, Section A-23. 

9. Ankhelasma typicum Sando (p. 17).

USGS loc. 17722-PC, Section U-4, Brazer Dolomite, Utah, Zone IIIA. Calicular view of etched silicified specimen, holotype,

USNM 120201. 

10,11. Acrocyathus girtyi (Hayasaka) (p. 16).

USGS loc. 4801H-PC (green label), Locality U-21, Monroe Canyon Limestone equivalent, Utah, Zone HID.

10. Longitudinal thin section, x3, holotype, USNM 120243b.

11. Transverse thin section, x3, holotype, USNM 120243a. 

12,13. Aulostylus tubiferus tubiferus (Hayasaka) (p. 18).

USGS loc. 5894-PC, Section M-23, Mission Canyon Limestone, Montana, Zone IIB.

12. Longitudinal thin section, x5, holotype, USNM 120247b.

13. Transverse thin section, x5, holotype, USNM 120247a.
U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 1334 PLATE 1

11 12 13 

ACROCYATHUS, AMPLEXIZAPHRENTIS, AMPLEXOCARINIA, AMPLEXUS, ANKHELASMA, AULOSTYLUS
PLATE 2

FIGURES 1,2. Canadiphyllum sp. (p. 18).

USGS loc. 22939-PC, Section U-17, Deseret Limestone, Utah, Zone HID.

1. Calicular view of etched silicified specimen having broken calice, xl, USNM 309845.

2. Alar view of same specimen, xl. 

3. Beaumontia sp. (p. 18).

USGS loc. 16811-PC, Section U-3, Lodgepole Limestone, Utah, Zone IB. Side view of etched silicified specimen, x2,

USNM 309842. 

4,5. Barytichisma amsdenense (Branson and Greger) (p. 18).

Locality WY-16, Amsden Formation, Wyoming, Zone VI.

4. Longitudinal thin section, x2, paralectotype, UM 2663.

5. Transverse thin section, x2, lectotype, UM 2663. 

6,7. Guerichophyllum sp. (p. 22).

USGS loc. 17996-PC, Locality M-18, Allan Mountain Limestone, Montana, Zone 1C.

6. Longitudinal thin section, x2, USNM 309852b.

7. Transverse thin section, x2, USNM 309852a. 

8,9. Caninia sp. (p. 19).

USGS loc. 13903-PC, Section M-9, Charles Formation, Montana, Zone IIIA.

8. Transverse thin section, x2, USNM 119760.

9. Longitudinal thin section, x2, USNM 119760. 

10,11. Siphonophyllia excentrica (Meek) (p. 28).

USGS loc. 17498-PC, Section M-23, Big Snowy Formation, Montana, Zone VB.

10. Transverse thin section, xl, topotype, USNM 144795a.

11. Longitudinal thin section, xl, topotype, USNM 144796A.
U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 1334 PLATE 2

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BARYTICHISMA, BEAUMONTIA, CANADIPHYLLUM, CANINIA, GUERICHOPHYLLUM, SIPHONOPHYLLIA
PLATE 3

FIGURES 1,2. Clisiophyllum sp. (p. 19).

USGSloc. 16952-PC, Section U-l, Monroe Canyon Limestone, Utah, Zone IV.

1. Longitudinal thin section, x2, USNM 309853b.

2. Transverse thin section, x2, USNM 309853a. 

3,4. Cyathaxonia tantilla (Miller) (p.20).

USGS loc. 16803-PC, Section U-3, Lodgepole Limestone, Utah, Zone IB.

3. Transverse thin section, xlO, USNM 222532b.

4. Longitudinal thin section, x7, USNM 222530. 

5. Cleistopora placenta (White) (p. 19).

USGS loc. 13872-PC, Section M-9, Lodgepole Limestone, Montana, Zone 1C. Top view, xl, USNM 119790. 

6,7. "Diphyphyllum" sp. (p. 20).

USGSloc. 20683-PC, SectionM-21, Mission Canyon Limestone, Montana, Zone IIIA.

6. Longitudinal thin section, x2, USNM 309854b.

7. Transverse thin section, x2, USNM 309854a. 

8,9. Dorlodotia? inconstans (Easton and Gutschick) (p. 20).

Section A-13, Redwall Limestone, Arizona, Zone IIIB.

8. Longitudinal thin section, x2, USNM 142240a.

9. Transverse thin section, x2, USNM 142240b.
U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 1334 PLATE 3

CLEISTOPORA, CLISIOPHYLLUM, CYATHAXONIA, DORLODOTIA?, "DIPHYPHYLLUM
PLATE 4

FIGURES 1,2. Faberophyllum sp. (p. 21).

USGS loc. 18700-PC, Section 1-3, Monroe Canyon Limestone, Idaho, Zone IV.

1. Transverse thin section, x2, USNM 309855a.

2. Longitudinal thin section, x2, USNM 309855b. 

3,4. Ekvasophyllum cf. E. inclinatum Parks (p. 21).

Section A-6, Redwall Limestone, Arizona, Zone IIIC.

3. Transverse thin section, x2, USNM 142227c.

4. Longitudinal thin section, x2, USNM 142227a. 

5,6. Duncanopora duncanae Sando (p. 21).

USGS loc. 22982-PC, Section WY-5, Amsden Formation, Wyoming, Zone VB.

5. Transverse thin section, holotype, x4, USNM 165184a.

6. Longitudinal thin section, holotype, x4, USNM 165184b.
U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 1334 PLATE 4

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DUNCANOPORA, EKVASOPHYLLUM, FABEROPHYLLUM
PLATE 5

FIGURES 1,2. Koninckophyllum cf. K. magnificum Thomson and Nicholson (p.22)-

GSC loc. 66421, Section NEBC-23, Prophet Formation, British Columbia, Zone IV.

1. Transverse thin section, x2, GSC type no. 67850a.

2. Longitudinal thin section, x2, GSC type no. 67850b.

3. Hapsiphyllum sp. (p.22).

USGS loc. 27264-PC, Section 1-15, Aspen Range Formation, Idaho, Zone HID. Transverse thin section, x4, USNM 309851.

4. Liardiphyllum hagei Sutherland (p. 23).

GSC loc. 16622, Section SWDM-10, Flett Formation, British Columbia, Zone IIIA. Transverse thin section, x2, holotype,

GSC type no. 10571. 

5,6. Siphonodendron "whitneyi" of Meek (p. 27).

USGS loc. 21982-PC, Locality 1-39, Scott Peak Formation, Idaho, Zone HID.

5. Longitudinal thin section, x2, USNM 309858b.

6. Transverse thin section, x2, USNM 309858a. 

7,8. Siphonodendron oculinum Sando (p. 27).

Section A-13, Redwall Limestone, Arizona, Zone IIIA.

7. Transverse thin section, x2, holotype, USNM 143242a.

8. Longitudinal thin section, x2, holotype, USNM 143242b.
U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 1334 PLATE 5

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HAPSIPHYLLUM, KONINCKOPHYLLUM, LIARDIPHYLLUM, SIPHONODENDRON
PLATE 6

FIGURES 1,2. Siphonodendron warreni Nelson (p. 27).

GSC loc. 58126, Section AL-38, Mount Head Formation, Alberta, Zone HID.

1. Transverse thin section, x2, GSC type no. 67851a.

2. Longitudinal thin section, x2, GSC type no. 67851b.

3. Metriophyllum sp. (p. 23)-

USGS loc. 18635-PC, Section 1-3, Lodgepole Limestone, Idaho, Zone IB. Transverse thin section, x6, USNM 309849.

4. Lophophyllum? sp. (p. 23).

USGS loc. 15265-PC, Section M-4, Lodgepole Limestone, Montana, Zone II. Transverse thin section, x2, USNM 309846. 

5,6. Siphonodendron sinuosum (Kelly) (p. 27).

Section AL-29, Mount Head Formation, Alberta, Zone IIIA or IIIB.

5. Longitudinal thin section, x2, lectotype, GSC type no. 8913.

6. Transverse thin section, x2, same specimen. 

7,8. Siphonodendron mutabile (Kelly) (p. 27).

GSC loc. 62002, Section AL-28, Livingstone Formation, Alberta, Zone IIB.

7. Longitudinal thin section, x2, GSC type no. 67852a.

8. Transverse thin section, x2, GSC type no. 67852b. 

9,10. Actinocyathus stelcki (Nelson) (p. 16).

USGS loc. 17906-PC, Section WY-3, Amsden Formation, Wyoming, Zone VB.

9. Longitudinal thin section, x2, USNM 165171b.

10. Transverse thin section, x2, USNM 165171a.
U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 1334 PLATE 6

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ACTINOCYATHUS, LOPHOPHYLLUM?, METRIOPHYLLUM, SIPHONODENDRON
PLATE 7

FIGURES 1,2. Multithecopora amsdenensis Sando (p. 24).

USGS loc. 17907-PC, Section WY-3, Amsden Formation, Wyoming, Zone VB.

1. Transverse thin section, x4, USNM 165177a.

2. Longitudinal thin section, x4, USNM I65177b. 

3,4. Palaeacis cuneiformis Haime (p. 25).

USGS loc. 18683-PC, Section 1-3, Little Flat Formation, Idaho, Zone HID.

3. Side view of etched silicified specimen, x 2, USNM 309844.

4. Top view of same specimen, x2. 

5. Palaeacis sp. (p. 25).

USGS loc. 16813-PC, Section U-3, Lodgepole Limestone, Utah, Zone IB. Top view of etched silicified specimen, x4,

USNM 309843. 

6,7. Michelinia expansa White (p. 24).

Section A-30, Kedwall Limestone, Arizona, Zone II.

6. Transverse thin section, x2, USNM 42244b.

7. Longitudinal thin section, x2, USNM 42244a. 

8. Neaxon? sp. (p.24).

USGS loc. 17356-PC, Section M-19, Lodgepole Limestone, Montana, Zone IB. Calicular view of etched silicified broken

specimen, x4, USNM 309859. 

9,10. Petalaxis simplex (Hayasaka) (p. 25).

USGS loc. 5893-PC, Locality U-22, Monroe Canyon Limestone, Utah, Zone IV.

9. Longitudinal thin section, x2, USNM 120249b.

10. Transverse thin section, x2, USNM 120249a.
U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 1334 PLATE 7

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MICHELINIA, MULTITHECOPORA, NEAXON?, PALAEACIS, PETALAXIS
PLATE 8

FIGURES 1,2. Schoencphyllum sp. (p.27).

USGS loc. 20017-PC, Locality U-23, Monroe Canyon Limestone, Utah, Zone VA.

1. Longitudinal thin section, x2, USNM 309857b.

2. Transverse thin section, x2, USNM 309857a. 

3,4. "Pseudodorlodotia" sp. (p. 26).

USGS loc. 19895-PC, Section 1-9, Scott Peak Formation, Idaho, Zone IV.

3. Transverse thin section, xl, USNM 309856a.

4. Longitudinal thin section, x 1, USNM 309856b.

5. Rhopalolasma sp. (p. 26).

USGS loc. 26625-PC, Section U-7, Deseret Limestone, Utah, Zone IIIA. Transverse thin section, x2, USNM 309850.

6. Rotiphyllum sp. (p.27).

USGS loc. 13926-PC, Section M-8, Lodgepole Limestone, Montana, Zone II. Transverse thin section, x6, USNM 119772. 

7,8. Pleurosiphonella drummondi (Nelson) (p.25).

USGS loc. 23002-PC, Section WY-6, Amsden Formation, Wyoming, Zone VB.

7. Transverse thin section, x4, USNM 165174a.

8. Longitudinal thin section, x4, USNM 165174c.
U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 1334 PLATE 8

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PLEUROSIPHONELLA, "PSEUDODORLUDOTIA," RHOPALOLASMA, ROTIPHYLLUM, SCHOENOPHYLLUM
PLATE 9

FIGURES 1,2. Syringopora sp. (p. 29).

Section A-3, Redwall Limestone, Arizona, Zone II.

1. Transverse thin section, x2, USNM 142245b.

2. Longitudinal thin section, x2, USNM 142245a. 

3,4. Sciophyllum adjunctivum (White) (p. 27).

Locality 1-40, Monroe Canyon Limestone, Idaho, Zone HID?.

3. Transverse thin section, x2, lectotype, USNM 144786a.

4. Longitudinal thin section, x2, lectotype, USNM 144786b. 

5. Sychnoelasma subcrassum (Easton and Gutschick) (p. 29).

Section A-24, Redwall Limestone, Arizona, Zone II. Transverse thin section, x3, USNM 142217c. 

6,7. Stelechophyllum banffense (Warren) (p. 28).

GSC loc. 59368, Section NEBC-5, Debolt Formation, British Columbia, Zone HID.

6. Transverse thin section, x3, GSC type no. 67853a.

7. Longitudinal thin section, x3, GSC type no. 67853b. 

8,9. Stelechophyllum micrum (Kelly) (p.28).

Locality AL-2, Pekisko Formation, Alberta, Zone IIB.

8. Longitudinal thin section, x4, holotype, GSC type no. 9648.

9. Transverse thin section, x4, holotype, GSC type no. 9648. 

10,11. Stelechophyllum microstylum (White) (p.28).

USGS loc. 490-PC, Section U-l, Lodgepole Limestone, Utah, Zone II.

10. Longitudinal thin section, x2, USNM 120244b.

11. Transverse thin section, x2, USNM 120244a.
U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 1334 PLATE 9

10 11

SCIOPHYLLUM, STELECHOPHYLLUM, SYCHNOELASMA, SYRINGOPORA
PLATE 10

FIGURES 1,2. "Thysanophyllum" astreiforme (Warren) (p.30).

Section AL-29, Mount Head Formation, Alberta, Zone IV.

1. Transverse thin section, x2, lectotype, GSC type no. 8911.

2. Longitudinal thin section, x2, lectotype, GSC type no. 8911. 

3,4. Zaphriphyllum sp. (p. 31).

USGS loc. 21674-PC and 21677-PC, Section WY-12, Madison Limestone, Wyoming, Zone IIIA.

3. Longitudinal thin section, x2, USNM 309848.

4. Transverse thin section, x2, USNM 309847. 

5,6. Turbinatocaninia? sp. (p. 30).

USGS Iocs. 17907-PC and 6960-PC, Sections WY-3 and WY-5, Amsden Formation, Wyoming, Zone VB.

5. Transverse thin section, x2, USNM 165142a.

6. Longitudinal thin section, x2, USNM 165159a. 

7,8. Vesiculophyllum incrassatum (Easton and Gutschick) (p. 31). 

Section A-18, Redwall Limestone, Arizona, Zone II.

7. Longitudinal thin section, x2, MNA G2.4143.

8. Transverse thin section, x2, USNM 142231.
U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 1334 PLATE 10

"THYSANOPHYLLUM," TURBINATOCANINIA?, VESICULOPHYLLUM, ZAPHRIPHYLLUM