DEPARTMENT OF THE INTERIOR 

FRANKLIN K. LANE, Secretary 

UNITED STATES GEOLOGICAL SURVEY . 

GEORGE OTIS SMITH, Director 

Professional Paper 118 

SOME AMERICAN JURASSIC AMMONITES 

OF THE 

GENERA QUENSTEDTICERAS, CARDIOCERAS, AND AMOEBOCERAS 

FAMILY CARDIOCERATIDAE 

BY 

JOHN B. REESIDE, JR. 

WASIIINGTON 

GOVERNMENT PRINTING OFFICE 

1919 
_..,...;---
CONTENTS. 

Page. 

Introduction ..... · ..................... ~ ......... ·.· ........ · .......................... ~ .......... :...... 5 

Distribution of species. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5 

Corr~lation ................... . ·. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 10 

Works cited in this paper ................................... , ............... . ................... ·.......... 11 

Systematic descriptions ............................... . ......... . ................ : ........................ 12: 

Family Cardioceratidae H. Douville............... . .. . ................................................. 12: 

Genus Quenstedticeras Hyatt ............................................................ ......... H 

Genus Cardioceras N eumayr and Uhlig ...... : ........................... · .... : . . l . . . . . . . . . . . . . . . . . . . 18 

Genus Amoeboceras Hyatt. ......................... _·. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 38-

Index ... .............. _ ............................... .. ....... . ...... . ................. _............... 64 

ILLUSTRATIONS. 

PLATES I-XXIV. JuraSsic ammonites .. ................ : ............... . ........... · ........ : ............. p~9 FIGURE 1. Map of Wyoming showing localitie3 where Cardioceratidae were obtained ................... _: . . . . 8 

3 

SOME AMERICAN .JURASSIC AMMO 1 

ITES OF THE GEN RA QUENSTEDTICERAS, 

CARDIOCERAS, AND A~OEHOCERAS, FAMILY OA DIOCERATIDAE. 

By Joi N B. REESIDE, Jr. 

INTRODUCTION. ] p~ied, ev~n when here seemed to be no recogThe species cordiforme Meek and Har den, nizable difference between the American and 

distans ~itfield, canadense Whiteaves, and European species. Furthermore, 'the writer has 

dubium_ Hyatt ~probably ~eluding whitnkyi J. accepted the nam s as applied by the European 

P. Smith), variously assigned to · the g[jnera authors, althoug there is a wide variation in 

Amaltheus, Quenstedticeras, Amoeboceras.,. and their conceptions of the same species, particuOardioceras, and subtumidum Whitfiel~ and larly as to · the · its within which a given 

Hovey, assigned to Aegoceras, include all the specific name sho dbe applied. Some authors 

prev~ously described s~ecies o~ Jurassi9 . am..: have extended su h names as cordatum Sower by 

. monites that are considered In this ~aper .. and excavatum So erby to ·cover a considerable 

Material ac.cumulated in the National M seum range of form, w ereas· others have ~estricted at Washington, mainly through the effo · ts of these names . and pplied other na~es to varifield parties of the United ·states. Geolf gical ant~ from the ty e. ·It seems to the writer at 

Sl}l'vey, has shown, however, the presencr of a least a great conv nience to have names for the 

number of undescribed forms of . consid,rable distinguishable f rms, even though i~termedi~ . 

sc~entific i~terest. These new species wefe ob- ate ~orms cause oubts as to the exact systetained .m. amly Jrom the Sundance formatton of matic rank of su h names. 

Wyommg. One comes from the Ellis ffrma- The writer tak s this opportunity to thank 

tion of Montana, one from Jurassic bed~ near Mr. T. W. Stant :n for advice . and assistance, . 

Lillooet, British Columbia, and three from the and Mr. C. A. Re ds, of the American Museum 

Oardioceras-beari:pg beds near the base i f the of N a.tural His to y, for the loan of Whitfield 

Naknek formation of Alaska. and Hovey's figu ed specimens. He wishes to 

The interpretation of the data furnis~d by express his appre iation of the work of Mr. W. 

the study. of these fossils has been hi ered 0. Hazard, who hotographed the specimens 

somewhat because the exact stratigraphi loca- and of Miss Fra~ es Wieser, who retouched th~ tio.n of much of the m.aterial is unknoll .. ' It photographs and assembled the plates. 

was gathered with regard merely for the g . neral 

stratigraphic and geographic position. ow- . 

ever, as a knowledge of tlie gener_al relati nship · of the forms to those of other areas is i~itself The species dis ussed in this paper are placed 

f 1 

under 33 specifi and· 2 vari'etal names. Of 

o va ue to the stratigrapher and systema ist it h d d . bl d . these names, 30 renew and 5 are old names 

as ~eeme esira e to escr~be and fig e the 2 of which are re tricted in application. ' 

species. Th d' · · I . e Istr1but1 n of · the species by forman makmg comparisons with Euro'pean forms tions is as follow : 

the write~ has had little authentically identified 

material and has relied entirely upon the published descriptions and figures. For this reason 

in part the European names have not been apSundance formation northeastern Utah: 

Cardioceras hyat i Reeside. 

Sundance formation f northwestern Colorado: 

Cardioceras cf. C wyomingense Reeside. 

!l 
6 SOME AMERICAN JURASSIC AMMONITES. 

Sundance formation of southern Wyoming~ 

Quenstedticeras? hoveyi Reeside. 

suspectum Reeside. 

Cardioceras auroraense Reeside. 

cordiforme (Meek and Hayden). 

crassum Reeside. 

crookense Reeside. 

haresi Reeside. 

hyatti Reeside. 

plattense Reeside. 

russelli Reeside. 

schucherti Reeside. 

stantoni Reeside. 

stan.toni var. obesum Reeside. 

wyomingense Reeside. 

Cardioceras? albaniense Reeside. 

incertum Reeside. 

latum Reeside. 

sp. undet. 

Sundance formation of northeastern Wyoming and western South Dakota (Black Hills): 

Quenstedticeras? hoveyi Reeside. 

subtumidum (Whitfield and Hovey). 

suspectum Re~ide. 

tumidum Reeside. 

Cardioceras americanum Reeside. 

bellefourchense Reeside. 

canadense (Whiteaves). 

cordiforme (Meek and Hayden). 

crassum Reeside. 

crookense Reeside 

Sundance formation, etc.-Continued. 

Cardioceras americanum Reeside-Continued. 

distans (Wh,itfield). 

distans var. depressum Reeside. 

haresi Reeside. 

obtusum Reeside. 

russelli Reeside. 

stillwelli Reeside. 

sundancense Reeside. 

whitfieldi Reeside . 

. whiteavesi Re~ide. 

wyomingense Reeside. 

Cardioceraa? latum Reeside. 

Ellis formation of Montana: 

Quenstedticeras collieri Reeside. 

Mariposa slate of California: 

Amoeboceras dubium Hyat>t. 

Cardioceras-bearing beds of British Columbia: 

Cardioceras canaden~e (Whiteaves). 

lillooetense Reeside. 

whiteavesi Reeside. 

Cardioceras-bearing beds in the lower part of the Naknek 

formation of Alaska~ 

Cardioceras alaskense Rooside. 

distana (Whitfield). 

distana var. depressum Reeside. 

martini Reeside. 

spiniferum Reeside. 

The following table and list show the geographic distribution of the species: 
DISTRIBUTION OF SPECIES. 7 

D-iStribution of species by localities. 

[T, Type locality; X, occurrence.] 

0 biJ "'0 .s 

j e 

0 biJ s 

0 ~ 0 ..... .~ p 0 · s ~ "S 0 

~ >=l · I-< :>-. ~ 

a:> ~ .g I-< 

-1-> a:> 

·m -1-> 00 

~ ~ 00 a:> >=l 0 c3 ~ I-< a:> a:>· a:> ~ .8 .g ~ -B -B ~ .$ ,.!:.d 

~ ..... ·.o I-< ;::! ] .,... 0 0 0 0 0 ~ I-< 

z z w. z ~ 0 ~ -< 

~------------ -a-1 ~~~~~~~_!)_~~~~~~~~~~~~~~ Quenstedticeras collieri Reeside..... . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . T X ........... . Quenstedticeras? hoveyi Reeside ... - . . . . . . X . . . . . . . . . ·x X . . . . . . . . . T . . . . . . X .......... _._ ........ . 

subtumidum (Whitfield and - Hovey) ........................................... T ........•................................. suspect urn Reeside ....... ~ . . . . . . . .. . . . T . . . . . . . . . . . . X ................................... ...... . tumidum Reeside ................ · .............. . ................. : ........... T ................. . 

Cru;~~:rlca~~:R::s~~~s-i~_e_ .. · ." ~ : : : : :·: : : : : : : : : : : : : : : : : : : : : : : : : :J: : : . T. : : : : : : : : : : : : : : : : : : : : : : : : : : : · ~ · : ~ : 

~E.:ri'~~i!?v~~~~::_:_:_:_:::: :::::: :~: ::: ::::::::: :T: :J:: :~: ::::::::: :::r::: ::::::::: :~: :::::: 

cordiforme (Meek and Hayden).. . .. .. . .. . .. . .. . X .. . .. . X .. . ? .. • .. . . .. X .................... . crassum Reeside.......... . . . . . . . . . . . . . . . . . ... . . . T . . . X ......................................... . crookense Reeside. . . . . . . . . . . . . . . . . . . . . . . . . . . . . X . . . . . . T ...................................... . distans (Whitfield) .................... ~ .... , ......... X ... T .. , ............................ X .. . distans depressum Reeside ........ · ........ ~ .. . .. .. . .. . T X . .. X .. . X .. . .. . .. . .. . .. . .. . .. . X ... 

haresi Reeside ........................... X ...... 

1 

... X X ........ . -... · · T · · · · · · · · · · · · '· · · · · · · · 

fJi~~t~ne::i~:~~id~::::::::::::: -~- ::: ::: ::: -~- ::: ::: ::: ::: ::: ::: ::: : :: ::: ::: ::: ::: ::: ::: .T. ::: ::: 

martini Reeside...... . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . T X 

~!i~~~:-:-::::::::::::: ::: ::::~: ::::::::: :T: ::: :~: ::::::::::::::: -~- :::::::::::: :C:':::::: :~?ci£~:~! ~~~~~~~---.· _.:::::: :_::: ::: ~:: ::: ::: ::: -~ ::: ::: ::: :: ~ ::: ::: ::: ::: ::: ::: ::: ::: ::: ::: "rj:" ::: 

~d~~~~tJidj::.~::: j\[ \?!I: !ii ;fi [~[ :[[ ::: ~:; ~T:I ::; [:: [x :[: ::: ~~i :::::: :x;l::: ::: 

incertum Reeside. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . '1J . . . • • • • . . • . . . . . . . . . . . . . . • . • . . . . . . . . . • . . . . . . . . • .. latum Reeside ......... ~ .............. X X ......... T X ............... X ................... · .. . sp. undet ......... · ............. · . . . . . . . . . . . . . . . )<( •..•.. ~ ••••• 1 .•••..••••.••••••••.•••.•.••.••.•••• 

Amoeboceras dubium Hyatt ............... ·1-- .

1 

.................... --· ....... · .............. T ..... · --~ 

a Numbers indicate.numbers asslgned to localities in list (pp. 8-9). 

I 
·' 

8 SOME AMERICAN JURASSIQ AMMONITES. 

The localities which have furnished the material on which this paper is based and the 

species identified from each locality are listed 

below. The numbers are indicated in the table 

on page 7. The accompanying index map 

(fig. 1) shows the localities in the State of 

Wyoming from which these fossils have been 

obtained, the numbers corresponding to those 

in the following list; 

4. Sundance formation, SW. i sec. 13, T. 23 N., R. 79 W., 

Carbon County, Wyo.: 

Cardiocera;s haresi. 

Cardioceras? latum. 

5. Sundance formation, Difficulty Canyon, in sec. 31, T. 

. 24 N., R. 80 W., Carbon County, Wyo.: 

Cardioceras hyatti. 

stanton.i. 

stantoni var. obesum. 

6. Sundance fo.rmation, east side of Freezeout Hills in T. 

25 N. , R. 78 W., Carbon County, Wyo.: 

0 50 150 MILES W-UWU---~----~--~----L---~ 

25 75 100 125 

FIGURE 1.-Map of Wyoming showing localities where Cardioceratidae were obtained. 

List of localities and fossils. 

1. Sundance formation, near dinosaur quarry of Carnegie 

Museum at Jensen, Uinta County, Utah: 

Cardioceras hyatti. 

2. Sundance formation, near Yampa River, Moffat County, 

Colo.: · 

Cardioceras cf. C. wyomingense. 

3. Sundance formation, Como Bluff, near Aurora, Wyo. 

(now known as Ridge, on the Union Pacific 

Railroad, in T. 22 N., R, 77 W., Carbon 

County): 

Quenstedticeras? hoveyi. 

suspectum. 

Cardioceras wyomingense. 

stan toni. 

russelli. 

auroraense. 

Cardioceras? latum. 

From belemnite zone near middle of formation: 

Cardioceras cordiforme. 

wyomingense. 

schucherti. 

crassum. 

Cardioc(;lras? incertum. 

albani3nse. 

sp. undet. 

From the nodules above the belemnite zone: 

Cardioceras cordiforme. 

crookense. 

7. /·Sundance formation, near Grand Canyon of the North 

Platte, south of Alcova, Natrona County, 

Wyo.: 

Quenstedticeras? hoveyi. 

Cardioceras plattense. 
DISTRIBUTION OF SPECIES. 9 

8. Sundance formation, half a mile northeast of the Johnson horse ranch in T. 52 N., R. 65 W., Belle 

Fourche River, 25 miles west of Sundance, 

Wyo.: 

Quensted tic eras? suspectum. 

subtumidum. 

Cardioceras distans. 

distans var. depressum. 

haresi. 

crassum. 

bellefourchense. 

Cardiocer:as? latum. 

9. Sundance formation; 4 miles west of Devils Tower, 

Crook County, Wyo.: 

Cardioceras distans var. depressum. 

cordiform e. 

wyomingense. 

haresi. 

crookense. 

obtusum. 

Cardioceras? latum. 

10. Sundance formation, 2 miles south ot Belle Fourche 

River, near Bear Lodge Butte (Devils Tower), 

Crook County, Wyo.: · 

Cardioceras distans. 

whitfieldi. 

11. Sundance formation, 4 miles west of SundancP., Crook 

County, Wyo.: . 

Cardioceras distans var. depressum. 

cordiforme? 

stillwelli. 

canadense. 

whiteavesi. 

sundancense. 

american urn. 

12. Sundance formation, foothill~ west Qf Bear Lodge 

Mountains, Crook County, Wyo.: 

Quenstedticeras? hoveyi. 

13. Sundance formation, Sundance-Aladdin stage road at 

south fork of Redwater Creek, Crook County, 

Wyo.: 

Cardioceras distans var. depressum. 

14. Sundance formation, miles southwest of Barrett 

(Aladdin), Crook County, Wyo.: 

Cardioceras wyomingense. 

15. Sundance formation, 1! miles north of Redwr.ter 

Creek, near boundary between South Dakota 

and Wyoming, in Wyoming: 

Quenstedticeras? hoveyi. 

Cardioceras cordiforme. · 

haresi. 

russelli. 

obtusum. 

Cardioceras? latum. 

16. Sundance formation, Red Canyon, half a mile sbuth 

of Mathias Peak, in sec. 24, T. 3 S., R. 7 E., 

Fall River County, S. Dak.: 

Quenstedticeras? ttrmidum. 

17. Ellis formation, sec. 7, T. 25 N., R . . 26 E., Blaine 

County, Mont. (Little Rocky Mountains): 

Quenstedticeras collieri. 

18. Ellis formation, sec. 27,· T. 25 N., R. 26 E., Blaine 

County, Mont. (Little Rocky Mountains): 

Quenstedticeras collieri. 

19. Mariposa slate, Texas Ranch, Calaveras County, Cal.: 

Amoeboceras dubium. 

20. Cardioceras-bearing beds, head of Big Creek, tribu- · 

tary of Chilcotin River; about 90 miles northwest_ of Lillooet, Lillooet County, British 

· Columbia: 

· Cardioceras canadense. 

lillooetense. 

w hiteavesi. 

21. Cardioceras-bearing beds in lower part Of Naknek 

form~tion, east side of Oil Bay, .Cook Inlet 

region, Alaska:. 1 

Cardioceras distans. 

distans var. depressum. 

spiniferum. 

martini. 

alaskense . . 

22. Cardioceras-bearing beds near creek entering Boulder 

Creek, Matanuska Valley, Alaska: 

Cardioceras martini. · ' 

Comparati~e list of American and most closely related European species. 

American species. ., . . European species. 

Quenstedticeras collieri Reeside ........................ . ..... Quenstedticeras Iamberti (Sowei-by). 

Quenstedticeras? hoveyi Reeside ....... , .... _' . . .............. ~ Quenstedticeras? goliath (D'Orbigny). 

subtumidum (Whitfield and Hovey).... . ............... . . goliath (D'Orbigny) var. De Loriol. 

suspectum Reeside .... ~ ................ .- .... :. .. .......... goliath (D'Orbigny) . . 

tumidum Reeside ..................... . . . ............... Quenstedticeras carina tum Eichwald. 

Cardioceras alaskense Reeside .............. . . . .............. ; Cardioceras vagum Ilowaisky. 

americanum Reeside. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . e~cavatum (Sower by) Lahusen. 

auroraense Reeside.. .... . .. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . nikitinianum Lahusen. 

bellefourchense Reeside .................. : . . . . . . . . . . . . . . (?) 

canadense (Whiteaves) ................... . . . ......... : .·.. zenaidae Ilowaisky. 

cordiforme (Meek and Hayden).-.~ ........... ~ ......... .'.. · cordatum (Sowerby) var. b De LorioL 

crassum Reeside ........................... .. .. · . . ....... . 

crookense Reeside ........................ · ............... . 

distans (Whitfield), ........ ~ ......... .. . . ........... . .. . 

popilian'ense Boden .. 

(?) 

cordatum (Sowerby) var. d De Loriol. 

rouilleri (Nikitin) Lahusen. · 

1 Stanton, T. W., and Martin, G. C., Mesozoic section on Cook Inlet and Alaska Peninsula: Geol. Soc. America Bull., vol. 16, pp. 394, 395, 

406,1905. 

I 
10 SOME AMERICAN JURASSIC AMMONITES. 

· . • American species. European species 

Card10ceras • dlStans • var • depressum Ree Bl · d e.---.-----.----- ... C ar d' 10ceras vertebrale (Sowerby) · var. a Ilowaisky. hares1 ReeB1de .. _. _. _____ ... ___ . ________________ . _____ . __ . (?) · 

~yatti Reeside. -.---- · -·· ·-- ·---------.-.-.-.-.----- ~--.- cordatum (Sowerby) Lahusen BorissJ'ak. 

lillooetense Rees1de _ d (S ' · · t' . . - ------------- ·- ·--------- ·--------- ·- cor atum owerby) var. a De Lonol. 

mar illl Reesi~e----------- ·-----.---.-----.-.- ... ----- . . excavatum (Sowerby) Borissjak. obtusum Rees1de .... ___ .... _____ . ______ . _______ .... _ _ _ _ _ _ (?) 

plattense Reeside. ____ . __ . ________ . ____ . _____ . _ ... _. ___ . _ _ _ (?) 

russelli Reeside. _____ ... _. _________ ...... _ .. __ · _ _ _ _ _ _ _ _ _ _ _ (?) 

schucherti Reeside __ ... _____ . _ . _ . _ . _ ... __ . ___ ... ___ .. _ _ _ rouilleri ·(Nikitin) . if · R . . . spm erum ees1de .. ________ ~. _. __ . ____ . _. _ .. __ .. ____ . _.. (?) 

stantoni Reeside. -------------- ·.·---.- ... -.- ... --- ... --.-- vertebrale (Sowerby) Boden. 

stantoni var. obesum Reeside. _ . _____ . _ .. __ . ____________ . quadradtoides (Nikitin). 

vertebrale (l3owerby) Boden. 

quadradtoides (Nikitin). 

stillwelli Reeside .. ____ . _ . ·. _ .. _ . · · · ·-.---.- ..... --------- excavatum (Sowerby) var. laevigata Boden. 

sundancense Reeside ___ - __ . t (S b 

whiteavesi Reeside- _____ . ___ -_- _- _- _- _· _· _· _- _- .· .- _- ~· .· .· _- _· _- _·._- _- _- _- _- _- _- _- _· ~:r~:~:a~so::~Yf~ ru;:;~:r;~;ii!~:~Y whitfieldi Reeside .... -.- ~--- ------·_ . - ___ . __ .. ____ . _ ... __ . tenuicostatum Nikitin. 

wyomingense Reeside.--.-------- _-- ____ .... __ ... _. _. _.. nikitinianum Lahusen. 

Cardioceras? albaniense Reeside. ____ . _. ____ . ______ ..... _ _ _ _ _ _ _ (?) , .. 

incertum Reeside __________________ .. _____ . _ ..... _____ .... (?) 

latum Reeside. _- ................ _ .... _ .......... ·....... (?) 

sp. undet _ .. _ . ____________ . ___ . __ . _ . __ ._ . __ . _ . _______ . . . . (?) 

Amoeboceras . . dubium Hyatt .................................. A moe b. ocerasaternans(VonBuch). · 1 bauhini(Oppel). 

CORRELATION. 

Sundance formation of W.voming, Utah, and 

Oolorado.-The Sundance formation has been 

· considere4 by nearly all American authors &s 

a single naleontologic and stratigraphic unit of 

Lower Oxfordian age, in the sense in which that 

term is applied by the continental and particularly the Russian stratigraphers. The present 

study bears out this interpretation. The fauna 

of the localities listed in this paper is essen-

. tially a unit, even though the species are 

. widely separated in geographic range. Some 

of the species from .northeastern Wyoming 

show affinities with the genus Amoebocera8 in 

the tendency to obsolescenc.e of the ventral 

ribs and may indicat~ a late stage of the Sundance, but no true representatives of Amoeboceras are known from the formation. A single 

speci~s from the Sundance of South Dakota 

Quenstedticeras? tumidum Reeside, may indicat~ an older horizon than the Oardioceras · zone 

but this is still open to some doubt. The clos~ relationship between the cardioceratids of the 

Sundance and those of the zone of Oardioceras 

cordatum Sowerby, of the European Jurassic 

indic~tes their equivalence. It is true that 

some of the American species seem to have no 

close relatives in the European faunas so far 

as o.ne may judge from the literature of the 

subject, but the remaining species are sufficient to establish the correlation. Many European stratigraphers refer the cordalum zone to 

the Lower Oxfordian, others to the Upper Oxfordian or to the Lower Corallian, ·and still 

others drop the terms Oxfordian and Corallian 

and substitute Argovian or Sequanian. 

Ellis formatio.n of Montana.-The single 

ammonite described from the Ellis formation 

of Montana would indicate a lower strati-· 

graphic position than that of the Sundance. 

The number of species of inver-tebrates common to, both formations is sufficient to preclude a great difference in age, yet many of the 

most characteristic forms of the one are lacking 

in the other, particularly the ammonites. · 

Quenstedticeras of the type of Quenstedticeras 

collieri is characteristic of a zone referred by . 

many continental stratigraphers to the latest 

Callovian, by others to the Lower Oxfordian 

or Divesian. · 

Mariposa slate of California.-The one speqies from the Mariposa slate is a true Amoeboceras and indicates, as long ago pointed out 

by Smith 1 and· Hyatt, 2 a later age than that 

of the Sundance formation, a conclusion which 

is strengthened by the presence of Aucella 

with the ammonite. Amoeboceras is characteristic of a· zone referred by many authors to 

the Upper Oxfordian, by others to the Kimmeridgian. 

1 Smith, J. P., Age of the auriferous slates of the Sierra Nevada: Geol. 

Soc. America Bull., vol. 5, p. 254, 1894. 2 Hyatt, Alpheus, Trias and Jura in the Western States: Idem pp 

407-413. . ' • 
WORKS CITED. 11 

Cardioceras-bearing beds of British · Columbia.-The three species of Cardioceras identified from British Columbia are in one collection 

accompanied by a species of Aucell:a related to 

Aucella bronni Quenstedt and by fragments 

of a belemnite. As the matrix of the ammonites is very different from that of the specimens of A ucella it is quite justifiable to assume 

that the conditions prevalent elsewhere 0btain 

here and that there are two different. horizons 

represented in the collection. The beds containing the species of Cardioceras are of the 

same age as the Sundance formation, whereas 

The accompanying table presents these correlations in compact form. 

WORKS CITED IN TIDS PAPER. 

Boden, Karl, Die Fauna des unteren Oxford von Popilany 

in Litau: Geol. u. palaeont. Abh., N . . F., Band 10, 

Heft 2, 1911. 

Borissjak, A., Die Fauna des Donez-Jura: Com. geol. 

[Petrograd] Mem., new. ser., livraison 37, 1908. · 

Bukowski, Gejza, Uber die Jurabildungen von Czenstochau in Polen: Beitr. PaUiontologie Oesterr.-:-Ungarns u. 

des Orients, Band 5, pp. 75-171, 1887 . . 

Douville, Robert, Etude sur les Cardioceratides de Dives, 

Villers-sur-1\fer, et quelques autres gisements: Soc. geol: 

France, Paleontologie, val. 19, fasc. 2, Mem. 45, 1912. 

Correlation of European and North American strata. 

Europe. I 

' Most Alaska: British United 

Columbia. States . Buckman.a Haug.b :pe Lap- . continental 

parent.c 'stratigraphers. 

Kimmerid- Kimmerid- Kimmerid- (?) (?) (?) 

gian. gian. gian. 

Kimmerid- Zone of 

gian. Aucella Mariposa 

Zone of . Lusitanian. Sequanian . Upper Naknek cf. slate. 

.A moeboceras. Oxfordian . formation. A. bronni. 

Zone of Argovian. Upper Upper J,ower Zone of Sundance 

Cardioceras. Oxfordian. Oxfordian. Oxfordian. Ccrrdioceras. formation. 

(?) 

Zone of Divesian. Lower Ellis 

Quenstedticeras. Oxfordian. formation. e 

Callovian. Callovian. Chinitna (?) . shale.d 

Zone of Callovian. Callovian. Cosmdceras. 

a Buckman, S. S., Geol. Soc. London Quart. Jour., vol. 69, p. 160, 1913. b Haug, Emile, ,Traite de geologie, vol. 2, fasc. 2, p. 998, 1910. c De Lapparen~: A : A. C., Traite de geologie, pt. 2, livre 1, p. 1289, 5th ed., 1905. a May include Middle Jurassic. e Relations to Sundance formation uncertain. (Seep. 10.) 

those containing Aucella ·are probably of the 

age of the Mariposa ·slate. 

Cardioceras-bearing beds of the Naknek formation of Alaska.---:The four species of Cardioceras and one variety from the lower part of 

the Naknek formation warrant the correlation 

of the zone containing them with the Sundance 

and with the Cardioceras zone of British Columbia. As the ov~rlying part of the 'Naknek 

formation contains Aucella, it probably is 

equivalent at least in part to the Mariposa 

slate and the Aucella zone of British Columbia. 1 

1 Stanton, T. W., and Martin, G. C., Mesozoic section on Cook Inlet 

and Alaska Peninsula: Geol. Soc. America Bull., vol. 16, p. 407, 1905. 

Healey, Maud, Ammonites ( Cardioceras) excavatus Sower by 

Ammonites ( Cardioceras) vertebral is Sower by, Ammonites 

(Cardiociras) cordatus Sowerby: Paleontologia Universalis, pis. 92-94, 1905. 

Hyatt, Alpheus,· Jura and Trias at Taylorsville, Calif.: 

Geol. Soc. America Bull., .vol. 3, pp. 395-412, 1892. 

Hyatt, Alpheus, Tl'i~ and Jura in the Western States: 

Geol. Soc. America Bull., val. 5, pp. 395-434, 1894. 

Ilowaisky, David, L'Oxfordien et le Sequanien des gouvernements de Moscou et de Riasan: Soc. imp. nat. Moscou Buil., new ser., val. 17, pp. 223-292, 1903. 

Lahusen, Joseph, Die Fauna der jlir~ischen Bildungen 

des Rjasanschen Gouvernements: Com. geol. [Petrograd] 

Mem., vol. 1, No.1; 1883. 

Logan, W. N., The stratigraphy and.invertebrate faunas of 

the Jurassic formation in. the Freezeout Hills' of Wyoming: Kansas Univ. Quart., vol. 9, pp. 109-134, 1900. 
12 SOME AMERICAN JURASSIC AMMONITES. 

De Lorioi, Perceval, Etude sur les mollusques et brachiopodes de l'Oxfordien (zone.a Am. renggeri) du Jura bernois, I: Schweizer. paleont. Gesell. Abh., vol. 25, No.6, 

i898. 

Meek, F. B., and Hayden, F. V., Paleontology of the 

upper Missouri: Smithsonian Contr. Knowledge, vol. 14, 

No. 172, pt .. l, 1865. 

Neumayr, Melchior, 'Uber klimatische Zonen wahrend der 

Jura und Kreidezeit: K . .Akad. Wiss. Wien Denkschriften, Band 47, pp. 277-310; 1883. 

Nikitin, S. N., Die Jura-Ablagerungen zwischen Rybinsk, 

Mologa, und Mischkin an der oberen Wolga: Acad. sci. 

St.-Petersbourg Mem., 7th ser., vol. 28, No. 5, 1881. 

-- Die Cephalopodenfauna der Jurabildungen des 

Gouvernements Kostroma: Russ. k. mineral. Gesell., 

2d ser., Band 20, pp. 13-88, 1884. 

D'Orbigny, Alcide, Paleontologie fran~aise; Terrains 

jurassiques, vo~. 1, 1850-1860. 

Quenstedt, F. A., Die Ammoniten des schwabischen Jura, 

Band II, Der braune Jura, Stuttgart, 1886-87. 

Smith, J. 'p;, Age of the auriferous slates of the Sierra 

Nevada: Geol. Soc·: America Bull., vol. 5, pp. 243-258, 

1894. 

Stanton, T. W., and Martin, G. C.; Mesozoic section on 

Cook Inlet and Alaska Peninsula: Geol. Soc. America 

Bull., .vol. 16, pp. 391-410, 1905. 

Whitfield, R. P., and Hovey, E. 0., Remarks on and descriptions of Jurassic' fossils of the Black Hills: Am. Mus. 

Nat. Hist. Bull., vol. 22, art. 23, 1906. 

Whiteaves, J. F., Description of a species of Cardioceras 

from the Crows Nest coal fields: Ottawa Naturalist; vol. 

17, pp. 65-67, 1903. 

SYSTEMATIC DESCRIPTIONS. 

Fa]Qlly CARDIOCERATIDAE H. Douville. 

The term Cardioceratidae was attributed by 

Robert Douville 1 in his discussion of the group 

to Henri Douville, 2 who characterized' -the 

·family as a "polymorphic group, the origin 

of which is still uncertain, and which appears 

in the Callovian; aperture of the shell simple, 

without lappets or rostrum." 3 He · inCluded 

in the family · the genera Macrocephalites, 

Oadoceras, Stephanoceras ( = Stepheoceras), Oardioceras, and Reineckia (i:ncluding S piticeras). 

In 1892 Hyatt 4 applied the name to a group 

including Oardioceras, Oadoceras, Quenstedioceras ( =Quenstedticeras),. and Neumayria 

Nikitin (not Bayle) but gave no discussion. 

In l900 Hyatt 5 grouped under the name 

Cadoceratida~ the same genera which in 1892 

1 Douville, Robert, Etude sur les Cardioceratides de Dives, Villerssur-Mer, et quelques autres gisements: Soc. geol. France, Paleontologie, 

vol.19, fasc. 2, Mem. 45, p._ll, 1912. 

2 Douville, Henri, Notes pour le cours de paleontologie, professe a 

!'Ecole des mines en 1889-90, 1890. 

a Translated from quot~tion by R. Douville. 

4Hyatt, Alpheus, Jura and Trias at Taylorsville, Cal.: Geol. Soc. 

America Bull., vol. 3, p. 410, 1892. 

s Von Zittel, K. A., and Eastman; C. R., Textbook of paleontology, 

p. 580, 1896. 

he put into the family Cardioceratidae and 

added the genus Amoeboceras to incl1Jde the 

group of Ammonites alternans von Buch. 

- In 1912 Robert Douville 6 included under 

the term Cardiocera tidae the genera Macrocephalites, Oadoceras, Ohamoussetia, Stepheoceras, Pachyceras, Quenstedticeras, and Oardi- · 

oceras (including Amoeboceras . Hyatt), and · 

characterized the group as follows: 

Aperture of shell simple, without lateral lappets or rostrum. * * * Costae continuous on the ventral region; no external nor ocular tubercles; lateral tubercles 

may or may not be present; the keel is never really individualized (as it is, for example, in Amaltheus of the 

group margaritatus). 

The enrollment is subject to great variation. The living chamber of individuals of large si~e is normally scaphitoid in the genera Macrocephalites, Pachyceras, Cardioceras, 

and in certain early species of Qu,enstedticeras (Q. carinatum). The suture possesses generally archaic characters; 

in other words, the several elements are all similar, with 

parallel axes perpendicular to the radius, simply decreasing in size in approaching the umbilicus. Only in 

. Stepheoceras an~ Pachyceras do these different elements 

commence to be individualized. * * * On the contrary, the sutures of the genera Cadoceras, Quenstedticeras, 

and Cardioceras are almost identical. 7 

·It seems to ·the wri te:r that the keel of 

Amoeboceras Hyatt does become individualized, but otherwise the description applies 

very well. 

The following summary gives the principal 

diagnostic characters. of . the genera which 

Douville includes in the family. 

Macrocephalites von Sutner; type M. macrocephalus (Schlotheim): Shell large, involute; 

increases markedly in circumference, and the 

external side is broad and rounded; scaphitoid 

in adult. Umbilicus narrow and deep in 

typical forms but wide in some Asiatic species. 

Umbilical shoulders invariably rounded. Apertme semilunate. All whorls re~arly covered 

with numerous sharp ribs, which · divide one or 

more times near the umbilicus and pass 

straight ou:t and across the venter. In typical 

for:m.S the ribs tend to become attenuated .near 

the umbilicus; In others they tend to be . accentuated near the umbilicus. Sutural features not generic. 

Oadoceras. (Fischer) Niki~in; type 0. modioZare (Luidius): Youngest whorls of shell 

rolinded; next older whorls show elevated 

cross section; latest whorls enlarged in the 

umbilical region so that cross section is lower. 

6 Douville, Robert, op. cit., p. 11, 1912. 1 Translation. 
CARDIOCERATIDAE. 13 

Typical aspect of adult is globular. Cro~s usually bifurcate once (some of them twice) 

section of whorl may narrow ventrally In and usually show a sharp siphonal sinus. 

some species but is never pinched or keeled. Sutural featu,res not generic. 

LiVing chamber of adult smooth. Umbilicus Arrweboceras Hyatt; type A. alternans (von 

narrow. Umbilical shoulders of adult sharp. Buch): Like Oardioceras except that the ribs 

Ribs rather straight on younger whorls, bend disappear on the venter, leaving two smooth 

forward in an arc on older whorls but show no bands bordering the keel, whose crenulations 

siphonal sinus. Primary ribs usually biflir~ therefore appear to have no relation to the 

cate, though in some specimens simple pFi- ribs. The umbilical ribs are_ straight and 

mary ribs are present. No intercalated second- usually simple to the venter; the secondaries 

ary ribs. Umbilical tubercles do not appear arise between them independently. 

before a relatively advanced age. Sutural Stepheoceras Buckman; type S. coronatum 

characters not generic. (Bruguiere): Shell stout or much depressed; 

Oharrwussetia R. Douville; type 0. cha- venter broadly rounded; in no species sharp 

mousseti (D'Orbigny) :Shell flattened laterally. or keeled. Living chamber of adults in some 

Venter narrow; pinched above the region species ornamented by more or less distinct 

where the shell attains a diameter of 60 or 70 ribs; in other species smooth. Umbilicus 

millimeters. Umbilicus . narrow; umbilical invariab~y wide; some species have sharp 

sh9ulders overhang the line of involution at tuberculated umbilical shoulders. Young 

the stage where the shell has a diameter of whorls in nearly all species tuberculate; 

80 to 90 millimeters. Ribs well marked only ornamentation of young and adult whorls 

on young and quite straight; lacking on practically the same. The ·ribs are nearly 

flanks above the stage where the diameter of straight and regularly' bifurcated. Suture in 

the shell . is 30 to 40 millimeters. First and some species has a very wide second lateral 

second lateral saddles of the suture ~qual in saddle, but in others it is narrow. 

height. Pachyceras Bayle; type P. lalandei (D'Or-· 

Quenstedticeras Hyatt; . type Q. 1lamberti bigny): Shell variable in form, either elevated 

(Sower by) : Inner whorls rounded, next whorls or much depressed. Older whorls rounde~; 

more or less narrowed externally but not with age they become narrowed ventrally and 

pinched and keeled. In adults the. ventral in some species even sharp. Living chamber 

region is rounded again and in no species is is ornamented by ribs, feeble in some species, 

sharp. Living chamber of large .forms ~mooth. very stout in o~hers. Adult scaphitoid. 

Aperture has a ventral crest In young. In Sculpture consists of stiff, straight, blunt ribs 

young the ribs are invariably sickle-sh~ped starting from the umbilical shoulder and 

and on the external region -have a distinct passing over the venter without any trace of 

siphonal sinus. With greater age the ribs. a sinus. Some species show tubercles at the 

become straighter. The umbilical ribs usually umbilical shoulders. 

fork and there are usually intercalated sec- In recent papers Buckman 1 assigns Stepheoondary ribs which do not extend to the ceras (type Ammonites humphresianus Sowerby, 

umbilicus. Costae not .tuberculate. Sutural not A. coronatus Bruguiere, which is the type 

. characters not generic. of Erymnoceras) to the family Stepheoceratidae 

Oardioceras Neumayr and · Uhlig; type 0. Neumayr; the genera Pachyceras, Macrocephcordatum (S~werby): Older whoris rounded~ alites, with Tornquistes and Erymnoceras, to 

younger whorls sharp-ventered and in many ·the family Pachyceratidae Buckman; and the 

species pinched into a keel. Adult is invari- genera Oadoceras, Quenstedtoceras [ = Quenstedably sharp-ventered. Living chamber of ticeras], Ohamoussetia, Oardioceras, Amoebaadult smooth. Umbilicus wide· only in youth. ceras, with the new genera Pseudocadoceras, 

Umbilical shoulders of adult sharp and over- Longaeviceras, Prorsiceras, Eboraciceras, and 

hang the line of involution. Ribs well marked Vertumniceras, to the family Cadoceratidae 

on young of many species at .the stage where Hyatt. Buckman considers Cardioceratidae 

the shell has a diameter of 10 millimeters; H. Douville as of doubtful validity t~ough 

usually sickle-shaped. Older individuals have 

·straighter rip~, commonly tuberculate. Ribs 1 Buckman, S. S., Yorkshire type anpnonites, Part XV, pp. xi, xii; 

Part XVI, desc. 116; ,Part XVII, pp. xiii, xiv, desc. 117, 118r l918. 

/ 
14 SOME AMERICAN JURASSIC AMMONITES. 

technical lack of publication, but inasmuch 

as Hyatt himself used the term Cardioceratidae in 1892 (s.ee above) with the same scope 

·as his later term Cadoceratidae, the former 

should stand even if Douville's usage is not 

accepted as establishing it. None of the 

species considered in this paper will fall 

within the new genera, which Buckman 

characterizes as follows: 

Genus PSEUDOCADOCERAS Buckman. 

Type, Am. longaevus Bean, syntype=P. boreale, Y. T. A. 

CXXIb. 

Syn. Cadoceras pars, Auctt. The Grewingki series, S. 

Buckman, 1913, 162. 

A cadoceratid series like young Cadoceras, but not 

developing cadicone stage, only feeble inflation; the 

catagenetic feature of excentrumbilication, commencing 

almost as soon as swelling, would prevent any cadicone development. (Suture line (D'Orbigny, 1848, 

CLXXIX, 9) rather simple, .L1 same length as EL, L2 

short and broad.) 

Genus LONGAEVICERAS Buckman. 

Type, Am. longaevus Bean, lectotype, Y. T. A. CXXIa. 

A cadoceratid seriee like Pseudocadoceras but prorsiradiate, attaining only to feeble swollen stage. Dev~lops 

oxycone? ' 

R. Douville (1912, p. 21) reckoned Am. galdrynus as a 

Chamoussetia, but its lack of inflation indicates a different 

stock. It is an involute development either of the present 

genus or of a parallel series; for this one may end without 

developing keel. Am. junijerus Phillips, 1829, vi, 23, is 

prior name for Am. galdrynus. 

Genus EBORACICERAS Buckman. 

Type, A. dissimilis Brown, holotype, Y. T. A. CXVIIIa. 

· A cadoceratid series which deylops considerable 

inflation but does not attain to cad1cone. Versi-, subflexiradiate, ribs of low relief, primaries short, secondaries 

angulate on periphery but straightening out in swollen 

stage; obsolescence follows. L1 r~ther narrow stemmed, 

longer than. EL; L2 quite short, rather broad (cf. D'Orbigny, CLXXVI, 3). Eboracum, York.· 

Genus VERTU:MNICERAS Buckman. 

Type, Am. vertumnus Leckenby, holotype, Y. T. A. 

CXVIa. 

A cadoceratid series distinguished by coarse subflexicostae which are strong on venter, where they· form a 

herring-bone pattern rising to a slight discontinuous ridge~ 

(Suture line with long L1 and quite short L2, V. spatiatum, 

CXVIc.) 

The forms described in this paper belong 

largely to the genus Oardioceras. Two species 

have a rounded venter in the adult stage and 

are referred with doubt to Quenstedticeras. 

They differ from the normal representatives 

in the obscure keel that is present in the 

younger stages and in this respect are like 

Quenstedticeras? goliath (D'Orbigny). Three 

species have rounded whorls in the younger 

stages and a sharp venter in the adult but do 

not show the suture and sculpture ofPachyceras 

and are referred with doubt to Oardioceras. 

One species is a typical A moeboceras and one 

a typical Quenstedticeras. 

Genus QUENSTEDTICERAS Hyatt. 

Quenstedticera~ collieri Reeside, n. sp. 

Plate I, :figures 1-8. 

Shell compressed; venter quite narrow but 

not sharp or keeled. Cross section of whorl 

oval, higher than wide. Umbilicus wide; 

umbilical shoulders evenly rounded. Living 

chamber and aperture unlmown. 

Sculpture regular. Where the diameter of 

the sh!3ll is. less than 8 millimeters the exposed 

flanks of the whorl are smooth. Where the 

diameter is more than 8 millimeters fairly 

sharp ribs rise within the umbilicus, b~nd 

backward a little, then pass ou,t over the 

flanks in a shallow forward curve to the venter, 

where they bend forward to form chevrons. 

Some of the ribs fork at the middle of the flank, 

and at about the same position a few inter-

. calated secondary ribs rise. An entire rib 

Type, Am. gregarius Leckenby, lectotyPe, Y. T. A. has a sigmoid form. 

CXVIIa. The suture has a narrow siphonal lobe. 

A _cadoceratid series swelling in serpenticone stage, First lateral saddle a little broader than 

presumably not attaining inflation of Eboraciceras by some 

Genus PRORSICERAS Buckman. 

10 per cent. Marked prorsiradiation. Well-developed siphonal lobe; first lateral lobe about same 

arcuation of ribs on narrowish rounded venter. Whorls width as siphons] lobe' but longer; remaining 

convergent. Suture line with rather short, broad lobes, elements small. Suture little incised. Radius 

L1 about as long as EL, L2 fairly developed. cuts first lateral lobe. 

()hamoussetia, which is much earlier in date, has remains This species is based on young specimens of similar prorsiradiation. Apart from date it could not 

be involute development, for present genus gives no only but is worthy of description as a member 

sign of attaining the much swollen galeatiform whorl with of the group of Quenstedticeras lamberti Sowdist.l.nct carination. erby. It differs from this typical specieS . 
QUENSTEDTICERAS. 15 

mainly in the paucity of intercalated ribs. Its 

sculpture and the fo'rm of the whorl separate 

it from any other American species. 

The species is named for Mr. A. J. Collier, 

of the Dnited States Geological Survey, who 

collected the type. . 

All specimens from Ellis formation. 

Localities: Type, sec. 7, T. 25 N., R. 26 

E., Blaine County, Mont.; also a number of 

poorly preserved fragments. Sec. 27, T. 25 N., 

R. 26 E., Blaine County, Mont., one small specimen and a number of fragments. 

The suture has ·a wide siphonal lobe; first 

lateral saddle as wide as the siphonal lobe; 

first lateral lobe reiatively narrow and a little longer than the siphonal lobe; second 

lateral saddle about as wide as first lateral 

saddle; second lateral lobe relatively small; 

auxiliaries small. Suture deeply ·incised. 

Radius just cuts first lateral lobe. 

This species is marked by its narrow umbilicus, short umbilical ribs, and the wide superior 

sad.dle of the suture. It may include the form 

figured by Wlu tfield and Hovey in Plate 

LXII of their paper. The writer has had the 

Quenst.edticeras? hoveyi Reeside, n. sp. opport~nity of comparing the original speciPlate I, figures 9-14; ~late II, figure 1. men with his types and finds it to agree in 

1906. Ammonites (Aegoceras) subtumidum vVhitfield and form arid suture, though the lack of the earlier 

Hovey (part), Am. Mus. Nat. Hist. Bull., vol. 22, art. whorls and the crushing of Whitfield and 

23, p. 400, pl. 62 (not pis. 60, 61). Hovey's. specimen introduce some doubt. The 

Shell stout, venter rounded. Earliest whorls species differs from Quenstediiceras? suspectum 

depressed; at 16 millimeters diameter the Reeside in its narrow umbilicus and wide 

cross section is circular and there is a shallow superior saddle. It differs from all forffign 

pinching of the venter; where the diaJneter species in the form of ·the umbilicus but 

is more than 16 millimeters the whorls are seems to be related to Quenstedticeras? goliath 

well rounded, a little F"ider than high. Type (D'OrbignyJ in' the obscure angulation on the 

is entirely septate, but a specimen 190 milli.:. venter of 1Jhe young produced by the shallow 

meters in diameter referred to this species has pinching. rThe round-ventered adult poi:n.ts 

last three-eighths of final whorl unseptate; to the genus Quenstedticeras, but the young 

aperture is unknown. The whorls of the speci- stages are not normal for the genus, and the 

men last me'n.tioned are a little broader pro- reference is therefore questioned. 

portionately than those of the type. Umbilicus This species is named for Mr. E. 0. Hovey, 

narrow, in width one-half height of outer whorl. of the American Museum of Natural History. 

At extreme end of the large specimen width or' All specimens from Sundance formation. 

the umbilicus is only one-third height of outer Localities: Type, foothills west of Bear 

whorl. Umbilical shoulders well rounded to Lodge Mountains, Crook County, Wyo.; also,· . 

a diameter of perhaps 20 millimeters, then another specimen. Near Aurora (Ridge), 

inner slope is steep. In mature individuals Wyo.; large specimen figured in Plate II and 

umbilical shoulders overhang. · one small one. Belle Fourche River, opposite 

Sculpture consists of straight, short, broadly mouth of Inyr nkara. Cree~, Crook Conn~y, 

rounded umbilical ribs . that fork close to the Wyo.; one large speCimen figured by Whitumbilicus into two or three rounded seeond- field and Hov~y. (This locality is probably 

aries, which pass out to the venter ~nd bend identical With locality 8 of list on page 97 

forward to make · a shallow siphonal sinus. half a mile northeast of the Johnson horse 

Intercalated secondaries uncommon. On the ranch, Belle Fourche River, Crook County, 

larger specimen, at a diameter of 120 milli- Wyo.) Near the Grand Canyon of the North 

meters, the umbilical ribs have vanished:r but Platte, Natrona County, Wyo.; one small 

the ventral ribs are visible and show a fai~t ~ spec~men . . A locality 1! miles north of Redsinus. At a diameter of 1~0 millimeters and water Creek, near the south boundary between 

more the whorl is smooth. Dakota and Wyoming, in Wyoming . 

. . 

'). 
I 

l6 SOME AMERICAN JURASSIC AMMONITES. 

Quenstedticeras? subtumidum (Whitfield and Hovey) 

Reeside. 

Plate IV, figures 1-4. 

1906. Ammonites (Aegoceras) subtumidum ·whitfield and 7 Hovey (part), Am. Mus. Nat. Hist. Bull., vol. 22, 

art. 23, p. 400, pl. 60, figs. 3, 4. (Not pl. 60, fig. 

5; pl. 61, figs. 1, 2; pl. 62.) 

Whitfield and B~ovey's description is as 

follows: 

Shell attaining more than medium size, and extremely 

variable in different individuals and at different stages of 

growth. Sectibn decidedly cordiform when. not co~­

pressed, the volutions imbedded about o~e~thrrd o~ th~Ir 

diameter, giving a moderately large umb1hcus wh1ch m 

the larger (older) specimens haS' vertical sides. At the 

period represented by the individual figured on Plate LX, 

figures 3 and 4 (two views of the same individual), the 

shell is generally very strongly annulated, or marked by 

very strong transverse and deeply marked ridg~s. crossing 

the volution, beginning at the edge of the umbilicus as a 

single protuberant transverse node or ridge, dividing 

Inidway of the side into two or three sharply elevated 

ridges, having a strongly forward curvature in crossing the 

dorsum, with an additional intercalated ridge added to 

fill up the space after crossing the outer, half of the shell. 

Shell moderately stout. Venter concealed 

in type where diameter is less than 50 millimeters; at the diameter of 50 millimeters it is 

quite _sharp but not pinched; it becomes ~ore 

rounded 'vith increase in size and at the diameter of 70 millimeters is broadly rounded and a 

little wider than high. Last half-whorl preserved unseptate; aperture unknown. Umbilicus narrow, in width a little more ~han 

half the height of the o~ter whorl. Umbilical 

·shoulders evenly rounded in younger whorls to 

a diameter of perhaps 30 millimeters; then 

inner slope is abrupt. 

Sculpture consists of rather coarse, straight, 

rounded umbilical, ribs which fork at about 

one-third the distance to the middle of the 

venter into two, rarely three, rounded secondaries. Intercalated seconrlary ribs are common. The secondaries pass outward .. to the 

venter and swing forward to form a chevron, 

sharp at 5o millimeters diameter but rounded 

at and above the diameter of 70 millimeters. 

The sculpture is persistent on the unseptate 

part of the type. There are 16 primary and 

48 secondary ribs to each whor I on the type. 

The suture has a moderately wide siphonal 

· lobe; wider first lateral sadd)e; first lateral 

lobe narrower than the first lateral saddle; remaining elements small. Radius just touches 

the first lateral saddle. 

This species is ~haracterized by its_ coarse 

sculpture · and sharp but unpinched· venter, 

which becomes rounded with increase in size. 

It seems to be a normal Q·uenstedticeras in the 

lack of pinching and in the rounding of the 

venter with aO'e, but as the innermost whorls 

have not beenl::>seen the reference is questioned. 

The writer has at hand the four specimens figured by Whitfield and Hovey... The origin&! 

of Plate LX, figures 3 and 4, of their paper, 

should receive the name, though · inappropriate, of s·ubtumidurn, as the ·originals of the 

remaining figures are distinct from it and from 

one another. .. Th~ species is easily distinguished from Q·uenstedticeras? hoveyi Reeside 

and Q. suspecturn Reeside by the form of the 

whorls. It differs from Quenstedticeras? goliath 

(D'Orbigny) De Loriol 1 only in the number of 

ribs to tho whorl and · the sh&rper umbilical 

shoulders. 

The type is in the American Museum of 

Natural History, New York City. · 

Locality: Type, Sundance formation, Belle 

Fourche River, opposite the mouth of Inyankara Creek, Crook County, Wyo. This locality 

is :apparently ·the same· as that half a mile 

northeas-t of the Johnson horse ranch, on Belle 

Fourche River." (See No. 8 in list of localities, 

p. 9.) 

Quenstedticeras? suspectum Reeside, n. sp. 

Plate II, figures 2-4; Plate III; Plate IV, ·figures 5-7; 

Plate V, figures 1-2. 

Shell .robust, stout; venter rounded. Early 

whorls up to s·millimeters diameter, broad and 

·depressed; from 8 to 12 millimeters in diameter broadly ·rounded with venter faintly 

pinched, and sub angular; above 30 ~limeters 

whorls are very stout, rounded, wider than 

high. The specimen shown in Plate II,_ fi~­

ures 2-4, is entirely septate, but another mdividual 260 millimeters in diameter1referred to 

this species (Pl. III, figs. 1, 2) shows half a whorl 

Uiiseptate, very stout, and scaphitoid; a per~ 

ture unknown. Umbilicus moderately wide, 

in width three-fifths the height of the outer 

whorl. tJmbilical shoulders gently rmmded in 

early who~ls to the stage where the diameter is 

'20 ~r 25 millimeters, then inner slope is steep, 

straight, and eventually becomes very high 

and finally overhangs the umbilicus. 

1 De Loriol Perceval, Etude s~ les mollusques et brachiopodes de 

l'Oxfordien (~one a," Am. renggeri) du Jura bemois, I: Schweizer. paleont. Gesell. Abh., vol. 25, No.6, p. 24, pl. 2, fig. 15, 1898. 
QUENSTEDTICERAS. 17 

Sculpture of early whorls regular. Where Wyo.; specimen 300 millimeters ~n diameter 

the diameter is less than 8 millimeters, the doubtfully r_eferred to this species and the origiwhorls are smooth. Where the diameter is nal of Plate III. 

greater than 8 millimeters, distinct ribs are 

present on the exposed area of the whorls.-

These high, rounded ribs rise ~ithin the umbilicus, pass outward with little curve about one~ 

third the distance to the venter, fork, pass 

almost to the median line of the venter, then 

bend ·forward to form a shallow, rounded sinus. 

Intercalated secondary.ribs are common. On 

the younger subangular whorls the sinus is 

quite angular and is really a broad chevron. 

Where .the diameter is greater than 80 millimeters, the ribs of the specime;n shown in 

Plate IV, figures 5-7, gradually become lower 

and more rounded,. and are much less distinct 

where the diameter is 120 millimeters~ Other 

specimens, which agree very well in every 

other particular With the last, have throughout 

the younger stages somewhat coarser and more 

rounded ribs. On the large specimen preserving part of the living chamber it is marked 

. only by faint _ striae. There are from 22 to 30 

primary ribs to the whorl. 

The suture has a relatively wide siphonal 

lobe; first lateral saddle and first lateral lobe 

narrow, subequal in width; first lateral lobe a 

little longer than the siphonal lobe; second 

lateral saddle and lobe small; auxiliaries small. 

Suture deeply incised. The radius just cuts 

the first lateral lobe. 

The chief characters of this species ·are the 

stout, persistently rounded whorls, the -moderately wide umbilicus, and· the regular sculpture, which is clearly visible to the region 

where the diameter is at least 120 millimeters. 

It is. related. to the form described as Quenstedticeras? hoveyi Reeside but differs in having lower whorls and a much wider umbilicus. 

It differs from all the European species, 

judged by published figures, in the form of the 

umbilicus, ~hough it is related to Quenstedticeras? goliath (D'Orbigny) in the obtuse angulation of the younger whorls. As in the species 

hoveyi Reeside, · the adult is like a normal 

· Quenstedticeras, but the young are not. 

·All specimens from Sundance formation. 

Localities: Type . locality, near Aurora 

(Ridge), Wyo.; also two other specimens and 

a number of fragments. Locality half a mile 

northeast or the. Johnson horse ranch, Belle 

Fourche River,· 25 miles west of Sundance, 

Quenstedticeras? tumidum Reeside, n. sp. 

· Plate IV, figures 8-10; Plate V, figures 3, 4. 

1906. Ammonites (Aegoceras) subtumidum Whitfield and 

. Hovey (part), Am. Mus. Nat. Hist. Bull., vol. 22, 

art. 23, p. 400, pl. 60, fig. 5. (Not pl. 60, figs·. 

3, 4; pl. 61; pl. 62.) 

· Shell very stout. Innermost · whorls not 

seen. Where diameter is more than 30 millimeters, cross section, of whorls is broad, evenly 

rounded. Type entirely separate; aperture· 

unknown. Umbilicus broad in young hut narrow in adult; its width is three-fourths the 

height of the outer whorl. Umbilical shoulders 

well rounded· at the diameter of 30 millimeters 

but become fairly sharp and have steep inner 

slope at perhaps 40 millimeters. · 

The sculpture at the diameter of 30 millimeters consists- of coarse, rounded. primary 

ribs which begin within the umbilicus and rise 

into a moderately high elongated node · at the 

middle · of the flank. From each node three, 

rarely two, rounded secondary ribs pass out 

across the broad venter and bend forward to 

form a· shallow rounded smus. Intercalated 

secondaries are common. There would be 18 

to 20 primary ribs and 60 to 65 secondaries on 

a complete whorl at the diameter of 30 millimeters. At the diameter of 50 millimeters 

the primary ribs show very 'faintly on the high 

umbilical wall and dq not rise into a node on 

the flank; the s.econdaries are as on the younger 

whorls, though practically straight. At the 

diameter of 7 5 millimeters the umbilical ribs 

show as low, broad swellings near the umbilical 

shoulders and the secondaries' pass as -faint 

straight · ridges across the venter~ At the 

diameter of 100 millimeters the whorl is smooth. 

The suture has a siphonallobe; firstlateral 

saddle and first lateral lobe of equal width. 

The first lateral lobe and the siphonal lobe 

equal in length. The second I a teral saddle and 

other elements small. The radius barely cuts 

the first lateral lobe. Suture moderately 

incised. 

The swollen form, fine ventral ribs, and 

nodose younger whorls mark this species. The 

whorls visible have the characterS of a normal Quenstedticeras, but uutil the absence of 

pinching on the whorls where the . diameter is · / 
18 SOME AMERICAN JURASSIC AMMONITES. 

less than ~0 millimeters is known the refer- 1 The youngest part of the type, at the diameter 

ence should be questioned. The forni and of perhaps 100 millimeters for the entire shell, 

sculpture distinguish this species from related is practically smooth and has a cr~nulated ~eel. 

American species. The closest foreign rela- Th~re would be 60 or 70 secondanes on a comtive, at least in form, is Quenstedticeras carina- plete whorl at the diameter of 80 millimeters. 

tum Eichwald. · If the inner ·whorls of this The suture is much incised; the siphonallobe 

specimen are not pinched, i{ is the only . true is wide, the first lateral saddle narrower; the first 

Quenstedticeras from the . Sutidance formation lateral lobe is longer than. tpe siphonallobe and 

that the writer has seen and would indicate as wide as the first lateral saddle; other elements 

that the Ellis formation of Montana has in small. The radial line cuts the first lateral lobe. 

some . places an equivalent in the lower part of This species is best recognized by its peculiar 

the Sundance~ compressed form with pinched venter and narThe type and only specimen is in the Ameri- row umbilicus. In form Oardioceras · vagum 

can Museum of Natural History, New York Ilowaisky 1 is quite like this species but is a 

City. little more compressed, has fewer secondaries, 

· Locality: Type, Sundance formation, Red and wider umbilicus. 

Canyon, south of Mathias Peak, South Dakota Locality: Lower part of Naknek formati9n, 

{probably sec. 34, T. 3 S., R. 7 E., Fall River eastshoreofOilBay,Cookinletregion,Alaska. 

County) . . 

~enus CARDIOCERAS Neumayr and Uhlig. 

Cardioceras alaskense Reeside, n. sp. 

Plate VI, figures .7-10. 

Shell much. compressed, venter ~inched a~d 

keeled. Cross section of whorls at the diameter 

of 15 millimeters oval, a little higher than wide, 

sharp-ventered; at the diameter of 40 millimeters much compressed, flanks subparallel, 

venter pinched, width about four-sevenths the 

height;· la tet . whorls more triangular, much 

higher than wide. Type entirely septate. 

Umbilicus narrow, in width one-fourth the 

height of the outer whorl. . Umbilical s:P,oulders 

gently rounded to the diameter of 20 millimeters, then sharper; the ~teep inner slope increases in height and finally overhangs the 

umbilicus. 

Ventral sculpture of whorls where the diameter is less than 10 millimeters not seen; umbilical ribs visible at the diameter of 7 or 8 millimeters. These low, rounded umbilical ribs pass 

first backward, then radially to the middle of 

flank, where they fork. The secondar~es pass to 

the venter and curve sharply forward to cross the 

keel as crenulations. At the diameter of 15 millimeters· there are 26 umbilical ribs to the whorl. 

At the diameter of about 50 millimeters the umbilical ribs are still distmct. At the diameter of 

80 millimeters the. umbilical ribs are practically 

absent; but at the middle of the flank the low, 

rounded secondaries rise, pass outward and 

· then forward, making crenulations on the keel. 

Cardioceras americanum Reeside, n. sp. 

Plate VI, figures 15-20. 

Shell compressed, discoidal. Type has maximum diameter of 50 millimeters. Whorl shows 

a triangular cro~s section, higher than wide; · 

venter slightly pinched, sharp; sides almost 

straight from umbilicus to keeL Younger 

whorls concealed in type, but in another specimen are· seen to be first depressed, then almost 

circula~ in cross section. Venter sharp at the 

diameter of about 8 millimeters, and where the 

diameter is more than 8 millimeters the whorls 

are high, compressed, and sharp ventered. 

Living chamber and aperture unknown. Umbilicus narrow; in widthabout half the height of 

the' outer whorl. Umbilical sh~ulders gently 

rounded to the diameter of about 25 millimeters, then umbilical shoulders become sharp 

and umbilical walls steep. 

Sculpture feeble but ·very irregular. On the 

type low, broad ribs rise at the umbilical shoulders and pass to the middle of the flank, where 

they fork into three low, rounded secondaries. 

Between these bundles intercalated secondaries 

ri~e. All bend evenly forward to cross the 

sharp venter as crenulations, and here and th'ere 

a secondary forks just before it reaches the middle of the venter. The last whorl of the type 

has 17 primary ribs and about 75 crenulations 

on ,the sharp venter. Other specimens show 

fine ventral ribs at the diameter of 10 milli1 Ilowaisky, David, L'Oxfordien et le Sequanien des gouvernements 

de Moscou et de Riasan: Soc. imp. nat. Moscou Bull., new ser., vol. 

17, P·, 270, pl. 11, fig. 1, 1903. 

---
CARDIOCERAS. 19 

meters, but the umbilical ribs do not begin 

until the diameter reaches 20 to 25 millimeters. 

The · suture has a very wide siphonal lobe, 

narrower first lateral saddle, and still narrower first lateral lobe. The second lateral 

saddle is about as wide as the first lateral lobe, 

the second lateral lobe -is quite narrow, and the 

auxiliaries feeble. The siphonal lobe is the 

.longest and the radius to it does not touch the 

other lobes. 

Oardioceras . americanum is best recognized 

by its high, triangular, flat-sided cross section, 

regular but low sculpture, and by the sutural 

proportions. It is distinguished from Oardioceras cord~forme (Meek and Hayden) in its 

more compressed form, finer and less elevated 

ribs, more abrupt umbilical shoulders, and long 

siphonal lobe of the suture; from Oardioceras 

whitfieldi Reeside by the coarser and more 

persistent sculpture, relatively coarse crenulations on the venter, and by the suture. It 

differs from Oardioceras excavatum (Sowerby) 

Lahusen 1 in the smaller number of umbilical 

ribs and the wider and steeper-walled umbilicus but is otherwise very close; from Oardioceras 

excavatum (Sowerby) Bukowski 2 by the apparently weaker sculpture, flatter-sided whorls, 

and steep umbilical shoulders; from Oardioceras 

cordatum (Sowerby) D'Orbigny 3 in the relative length of the siphonal lobe and form of 

umbilical shoulder. 

Locality. Type and four other specimens 

from Sundance formation, 4 miles west of 

Sundance, Wyo. 

Cardioceras auroraense ;Reeside, n. sp. 

Plate X, figures 1-5. 

Shell compressed, keeled. Cross section of 

adult whorl triangular, wider than high; earlier 

whorls fairly stout. Living chamber and a,perture not preserved. - Umbilicus narrow, in 

width about half the height of outer whorl of 

adult; in earlier whorls umbilicus apparently 

wider~ · Umbilical shoulders sharp and high 

on later whorls. 

1 Lahusen, Joseph, Die Faunader jurassischen Bildungen des Rjasan-. 

schen Gouvernements: Com. geol. [Petrograd] Mem., vol. 1, No.1, p. 48, 

pl. 5, fig. 2, 1883. 

2 Bukowski, Gejza, Uber die Jurabildungen von Czenstochau in 

Polen: Beitr. Palii.ontologie Oesterr.-Ungarns u . des Orients, Band s; p. 

130, pl. 26, figs. 21, 22, .1887. 

a D'Orbigny, Alcide, PaieontologiefranQaise; Terrainsjurassiques, vol. 

1, p. 514, pl. 193, 1850-1860. 

Sculpture of whorls where the diameter is less 

tha:ri 80 · millimeters unknown. On. younger 

whorls it consists of high, sharp umbilical ribs 

which rise to a maximum at the middle of the 

flank, bifurcate, pass to the venter, when~ they 

are· raised slightly, then bend forward to cross 

the keel as crenulations. Intercalated secondaries ate common. On the last whorl of the type 

the ribs decrease in distinctness and are almost 

imperceptible at the end. On the fragment 

figured the youngest part preserved is perfectly smooth. The type · has 15 primary ribs 

and about 45 secondaries to the whorl. 

The suture shows a relatively· wide siphonal 

lobe, wide first lateral saddle; short and narrow 

first lateral lobe, small second lateral saddle 

and second lateral lobe; auxiliaries small. The 

radius just touches the first lateral lobe~ 

This species is based on a · partly crushed, 

weathered individual, which is about 120 millimeters in maximum diameter, and on an apparently normal mold of part of another, 

which shows part of three' whorls. The material is poor, but the great 'lateral compression 

of the . whorls, much more than the crushing 

would account for, together with the high, 

sharp sculpture and the suture, separate it 

from the other species at hand. · Oardioceras 

· plattense R~eside ·and Oardioceras schucherti 

Reesid·e are related but are both less compressed. Oardioceras plattense has coarse primaries which divide nearer the umbilicus; Oardioceras schucherti has perhaps twice as many 

secondary ribs for the same number of primary 

ribs. Oardioceras cordatum (Sowerby) Lahusen4 

is related but has a somewhat different cross 

section and more ribs; Oardioceras cordatum 

(Sowerby) var. a De Loriol 5 has more ribs; 

Oardioceras nikitin'ianum Lahusen 6 is very 

close in form to· our species. Oardioceras cordatum (Sowerby) as refigured by Healey 7 is 

strikingly like this species except that it is 

much stouter. · 

Localities: Type; Sundance formation, Aurora (Ridge), yYyo. (Hyatt and Russell); Como 

Bluff, near Aurora (Ridge), · Wyo. (C. 

Schuchert). 

4 Lahusen, Joseph, op. cit., p. 49, pl. 5, fig. 3, 1883. 

5 De Lorio!, Perceval, Etude sur les mollusques et brachiopodes de 

l'Oxfardien (zone a Am. renggeri) du Jura bernois, I: Schweizer. paleont. 

Gesell. Abh., vol. 25, No.6, p. 16, pl. 2, figs. 1-3, 1898. 

6 Lahusen, Joseph, op. cit. , p. 49, pl. 5, fig~ 8. 7 Healey, Maud, Paleontologia Universalis, pis. 94, 94a, 1905. 
20 SOME AMERICAN JURASSIC AMMONITES. 

Cardioceras bellefourchense Reeside, n. sp. 

Plate XI, figures 1, 2; Plate XII, figures 1, 2. 

lobe-; first lateral lobe longer than the siphonal; 

second lateral saddle of medium size; second 

lateral lobe and' auxiliaries sm~ll. The radius 

1906. Ammonites (Amaltheus) cordiformis Meek- and Hay- cuts the first lateral lobe and touches the 

den (part). Whitfield and Hovey, Am. Mus. Nat. second. 

Hist. Bull., vol. 22, art. 23, P· 401, pis. 58' 59· Oardioceras bellefourchense is best recognized (Not pls. 52-56, 60.) - by its broad, rounded whorls with sharp venter, 

Shell stout; venter . angular in all stages coarse, rounded ribs, and the proportions of the 

above 10 millimeters in diameter, which are suture. It differs from Oardioceras crassum 

the only stages seen; not pinched where the Reeside in the form of the whorls, especially 

diameter is more than 40 millimeters and not the inner· whorls, and in the relative widths of 

much so below that diameter. Cross section the superior saddle and amount of incision of 

of early whorls oval, keeled, higher than wide; the suture; from Oardioceras plattense Reeside 

at the diameter of 35 millimeters well rounded, in lacking the second bifurcation; from Oardiosubequal in,width and height; at the diameter ceras russelli Reeside in the coarser sculpture 

of 50 millimeters broadly cordiform with ob- and different suture. The ·writer knows no 

scure ventral shoulders; at the diameter of 70 foreign forms like this species. 

millimeters an:d more rounded cordiform, width Locality: Type, Sundance formation, half a 

great_er than height, ventral angle obtuse. mile northeast of the Johnson horse ranch, 

Last one-eighth whorl of type unseptate, aper- Belle Fourche River, 25 miles east of Sundance, 

ture unknown. Umbilicus wide in · young Wyo. 

stages, narrow in adult; width about half the Cardioceras canadense Whiteaves. 

height of outer whorl. Umbilical shoulders Plate XVII, figures 5-11. 

gently rounded where the diameter is less than 1903. Cardioceras canadense Whiteaves, Ottawa Naturalist, 

40 millimeters; where it is more than 40 milli- vol. 17, pp. 65-67, figs. 1, 1a. 

meters the inner slope is flat. At the diameter Whiteaves de~cribed this spec-ies as follows: 

of 90 millimeters the inner slope is parallel to . Shell, at least in its immature .:stage, compressed, shal~ 

the axes, but at the maximum diameter of the lowly and rather widely umbilicated, with a small and 

specimen, 130 millimeters, it overhangs the minutely crenulated keel. Wkorls about five, increasing 

umbilicus. rather rapidly in size and rather strongly embracing; about 

Sculpture regular. Umbilical part of earlier one-half the sides of the inner ones being covered by the 

overlap of those that succeed them. Umbilicus occupying whorls shows high rounded ribs. Last whorl about one-third of the entire diameter, oneach side, though 

of type has coarse, high, rounded umbilical ribs · its margin is rounded and very indistinctly defined; . 

on the older half, but on the 'younger half these peripheral carina neither very prominent nor distinctly 

gradually decrease in height and distinctness compressed. 

Surface of each side of the outer volution marked with a· ~nd are practically lacking on _ the unseptate few comparatively large and distant but narrow and acute 

part. At one-third the distance from urn- primary radiating ribs, that commence at the suture and 

b1licus to venter, these primary ribs divide into terminateabouthalfwayacross, ina small pointed tubercle. 

two, rarely three, rounded secondary ·ribs Of these ribs there are about ten in the specimen figured. 

which first pass outward and then bend rather Besides them there are rather more than twice as many 

:Sharply forward to form chevrons on the sharp small short secondary ribs, that are little more than narrow, 

V enter. Toward the end of the last whorl transversely elongated, compressed and acute tubercles, on the outer half of each side. The primary ribs almost 

these secondary ribs also become less. distinct, bifurcate from a median tubercle, and seem to occasionally 

though still clearly visible. There are some . alternate with an intercalated secondary rib, but the secintercalated secondaries and a few seconda- · ondary ribs are not quite continuous with any of the 

· f k th t 'It· b bl th t th primaries. Between the secondary ribs, a'lso, and parallel r.le~ _or on e .ven er · IS pro a e a e to them there are a few fine radiating raised lines. 

liVIng chamber IS smooth finally. There are 15 Sutural line unknown, as are also the ~xact shape and 

umbilical ribs and about 45 secondaries on the sculpture of the adult and the contour of the outer lip. 

last whorl of the type, though the inner whorls Shell compressed, venter shouldered and 

show more pr~maries. keeled. Cross section of whorl rudely hexThe suture shows a wide siphonal-lobe; first agonal, widerthanhigh. Whorls not much overlateral saddle and first lateral lobe about equal lapping Final one-eighth of lastwho~l of best 

in width, both' narrower than the siphonal specimen preserved unseptate. Aperture un-
CARDIOCERAS. 21 

known. Umbilicus wide, four-fifths the height 

of outer whorl in width. Umbilical shoulders 

gently rounded. 

Sculpture very characteristic. Earlier whorls 

smooth on flanks where the shell is less than 15 

millimeters in diameter; distinct ribs show on 

venter wher~ the diameter is mo.re than 8 millimeters. These . ribs appear first as a faint, 

crenulated keel and extend outward from it 

with growth of shell. Where the diameter is 

more than 15 millimeters umbilical ribs are 

present. These ribs begin at line of involution, 

pass first backward, then outward in a shallow 

forward curve, increasing in height till they 

reach a point ·beyond the middle of the flank, 

where each rises into a high node and forks into 

two or three short secondary ribs·. Some intercalated se~ondaries are present. Each short 

secondary rises into a node and divides, the 

branches bending forward and passing across 

the keel as sharp crenulations. There are 16 

·· umbilical ribs on the last whorl of the larger 

specimen (Pl. XVII, fig._ 5) and there would be 

about 90 crenulations on the complete keel. 

The suture shows a very wide siphonal saddle, 

wide first lateral saddle, narrow first lateral 

lobe, relatively wide second lateral saddle, and 

small second - lateral lobe. The radius· just 

touches the tip of ~he first la~erallobe. 

The nodose ribs with double forking, the fine 

secondaries, and_ the sutural charact'1rs distinguish this species. The figures published 

by Whiteaves are not particularly go-~UI , but 

the details shown and the descriptio~ given 

tally so well with the Wyoming specimens at 

hand that there seems to be little doubt of their 

identity with the Canadian form. 

Oardioceras caJ~;adense Whiteaves is most like 

Oardioceras whiteavesi Reeside but differs 

sharply in having forked secondaries. It differs 

from Oardioceras sundancense Ree~ide in the 

presence of . high nodes, more vigorous sculpture, and narrower umbilicus. It is distinguished from Oardioceras vertebrale (Sower by) 

var. densiplicatum Boden 1 by the persistence 

of smooth flanks on the inner whorls to a diameter of 15 millimeters, fewer umbilical ribs, 

much finer ventral ribs, and stouter whorls; 

from Oardioceras zenaidae Ilowai~ky2 only by 

1 Boden, Karl, Die Fauna des unteren Oxford von Popilany in Litau: 

Geol. u. palaeont. Abh., N. F., Band 10, Heft 2, p. 3i, pl. 1, fig. 14, 1911. 

2 Ilowaisky, David, L'Oxfordien et le Sequanien des ·gouvemements 

de Moscou et de Riasan: Soc. imp. Nat. Moscou. Bull., new ser., vol. 

17, p. 269, pl. 10, figs. 33-36, 1903. 

the slightly greater number of ventral ribs; 

from Oardioceras sp. Borissjak3 by ·the lack of 

a second bifurcation of the ribs. . 

Lecalities: Type, Oardioceras-bearing beds, 

2! miles northeast of Fernie, British Columbia. 

Sundance formation, 4 miles west of Sundance, 

Wyo;; ·also four other-specimens. Oardiocerasbearing beds Lillooet, British Columbia, one 

fragment, indistinguishable from this species. 

Cardioceras cordiforme ~Meek and Hayden) Neumayr. 

Plate VII, figures 1-6; Plate VIII, figtires 1-7; Plate IX, 

figure 1. 

1859. Ammonites cordijormis Meek and Hayden, Acad. 

j Nat. Sci. Philadelphia Proc. for 1858, p. 57. 

1860. Ammonites cordiform·is Meek and Hayden, Acad .. 

Nat. Sci. Philadelphia Proc., October, 1860, p. 418. 

1865. Ammonites cordfformis Meek and Hayden. Meek 

and Hayden, Paleontology of the upper Missouri: · 

Smithsonian Contr. Knowledg~, vol. 14, No; 172) 

pt. 1, pp. 122-123, pl. 5, figs. 2a-e. 

1880. Ammonites cordijormis Meek and Hayden (part). 

Whitfield, Report on the geology and resources of 

·the Black Hills of Dakota, U. S. GeoL and Geog. 

.Survey Rocky. Mtn. region, pp. 378-380. (Not 

· I pl. 6, figs. 20-24.) 

1S83. Cardioceras cordijorme (Meek and Hayden). Neumayr, K. Akad. Wiss. Wien Denkschr., Math.-

Naturwiss. Classe, Band 47, p. 302. 

1894., Cardioceras cordijorme (Meek and Hayden). Smith, 

Geol. Soc. America Bull., vol. 5, p. 253. 

1894. Quenstedioceras cordij~rme (Meek and Hayden). 

Hyatt, Geol. Soc. America Bull., vol. 5, p. 409. 

1900. Cardioceras cordijor1'M ,(Meek and Hayden) (part?). 

. J Logan, Kansas Univ. Quart., Series A., vol. 9, 

· No.2, p. 124. (Not pl. 27, figs. 1-12?.) 

1906.1 Ammonites (Amaltheus) cordiformis Meek and Hayden (part). Whitfield and Hovey, Am: Mus. Nat~ 

Hist. Bull., vol. 22, art: 23, p. 401. (Not pls.52-59; 

j not pl. 60, figs; 1, ~-) . , 

1910. _Cardiqceras cordifprme (Meek and Hayden) (pa,rt). 

Grabau andShimer, North American index fossils, 

vol. 2, p. 185, :fig. 1442, septum. (Not fig. 1442, 

entire _ammonite.) · 

Meek and Hayden's original description is as 

follows: -

Shell lenticular, adult specimens being m"Q.ch 'm6re convrx than the young; umbilicus rather small,- "or .froin 

one-~hird to one-half the breadth of the outer whorl; ' dorsUm carinate; volutions increaf?ing ·so as to more than 

double their diameter every turn, each of the. inner ones 

fro~ 1 one-half io three~ fourths hidden within the ventral 

groove of the succeeding whorl. Surface-ornaruented by 

numerous small flexuous costae, which, in crossing the 

sides, increase by division and intercalation so as to number 

two or three times as many at the periphery as around the 

umbVicus. In approaching the dorsum they curve f<?r3 BJissjak, ~~., Die F'auna des Donez-Jura: Co~. geol. (Petrograd) 

Mem.,l new s~r., livraiso~ 37, p. 66, pl: 1, fig. 11, 1908. 
22 SOME AMERICAN jURASSIC AMMONITES. 

ward, and all cross the dorsal carina, to' which, in yonng 

specimens, they impart. a distinctly crenate outline. 

Greatest diameter of a specimen divested of its outer 

whorls, 3.30 inches; diameter of its last turn, from umbilicus to dorsum, 1.6-3 inch; breadth of same, 1.46 inch. 

Shell compressed, _discoidal throughout developme~t .. The venter is distinctly sharp at 

a diameter of 7 or 8 millimeters and continues 

so throughout life. Where the diameter is 

greater than 40 millimeters the cross section is 

triangular, .expanding laterally in the umbilical 

region as the size increases. One specimen preserves five-eighths of a whorl unseptate, with 

some constriction toward the outer end, though 

this may be accidental. Aperture not seen. 

Umbilicus wide in young (two-thirds height of 

whorl), narrow and deep in adult; umbilical 

shoulders rounded where the diameter is less 

than ·20 to 30 millimeters, then the straight 

inner slope is parallel to the axis of the shell 

· until the diameter reaches 70 to 80 millim~ters; 

youngest whorls have overhanging umbilical 

walls. 

· The sculpture differs somewhat in coarseness 

in individuals of same size. Where the diameter is less than 6 millimeters the whorls are 

smooth and broadly rounded. At the diameter 

of 7 or 8 millimeters faint but distinct ventral 

ribs and a keel appear. These become more 

distmct with increase in ·size, and at the diameter of 25 millimeters there are 25 to 30 fairly 

sharp ribs to the whorl at the umbilical shoulders and 60 to 70 on the venter. Each umbilical rib passes outward straight or faintly 

curved, bifurcates at about the middle of the 

flank, and bends forward to cross the venter. 

There are a few intercalated sec.ondaries and a 

few· simple primaries. The . outer- whorls, preserved as internal casts, have in medium-sized 

individuals relatively fine rounded ribs of the 

same type as on the inner whorls. These ribs 

become fainter with age, and the largest specimens have only striae on the ovter whorl. 

Meek's type at the diameter of 85 millimeters 

has 22 umbilical and 65 ventral ribs to the 

whorl. 

-The suture of this species has the siphonal 

lobe broader than the first lateral saddle and 

the first lateral lobe, the , first lateral lobe 

broader than the first lateral saddle, and the 

second lateral lobe about half the width of the 

first lateral lobe. The siphonallobe is shorter 

than the first lateral lobe. The radius cuts the 

first lateral lobe and touches the second. 

Meek and Hayden's original description is 

sufficiently general to include more than the 

typical Oardioceras cordiforme, and in ,this he 

has been followed by later workers. The type 

specimen, however, represents a quite distinct 

form and to it the writer would restrict the 

name. The diagnostic characters of Oardiocerds . 

cordiforme are the evenly curved, unshouldered 

:flanks of the whorls; the narrow umbilicus; 

fine, low, shallow sigmoid ribs, persisting in 

some individuals to a diameter of perhaps 200 

millimeters; wide siphonal lobe an~ narrow 

superior saddle of the suture. · 

Oardioceras cordiforme (Meek and Hayden) 

differs from Oardioceras whitfieldi Reeside in its 

stronger and more persistent sculpture, and its 

relatively lower cross section; from Oardioceras 

americanum .Reeside in its stouter form, stronger 

sculpture, and less abrupt umbilical shoulders 

in .the young, and in the ·sutural details; from 

Oardioceras crookense Reeside in its less swollen 

venter, more regular sculpture, and narrower 

first lateral lobe of the suture. It is distinguished from Oardioceras excavatum (Sowerby) 

Nik~tin 1 by the lack of pinching near the keel 

and by its regular sculpture; from Oardioceras 

cordatum (Sowerby) var. b. Loriol 2 by its 

narrower umbilicus and by sutural details. 

Oardioceras popilianense Boden. 3 differs in sut"Qre, finer ribs, and in the height of the whorl, 

though very like Meek and Hayden's species in 

general expression. 

All specimens from Sundance formation. 

Localities: Type, "southwest base of Black 

Hills," Wyo. i also several small specimens. 

Locality in T. 25 N., R. 78 W., Freezeout Hills, -

Crook County, Wyo., 1 larg~ specimen and ·18. 

small ones. ' Locality, 1! miles north of Redwater Creek, near the bolindary between Wyoming and South Dakota in Wyoming; 14 -small 

specimens. Locality, 4 miles west of Devils 

Tower, Crook County, Wyo.; 10 small speci~ 

mens. 

1 Nikitin, S. N., Die Jura-Ablagerungen zwischen Rybinsk, Mologa, 

und Mischken an der oberen Wolga: Acad. sci. St .• Petersbourg Mem., 

7th ser., vol. 28, No.5, p. 52-54, pl. -2, fig. 13, 1881. 

2 De Loriol, Perceval, Etude sur les mollusques et brachiopodes de 

l'Oxfordien (zone a Am. renggeri) du Jura bemois, 1: Schweizer. paleont. 

Gesell. Abh., vol. 25, No.6, p. 16, pl. 2, figs. 4-7, 1898. 

a Boden, Karl, Die Fauna des unteren Oxford von Popilany in Litau: 

Geol. u. paleont. Abh., N. F., Band. 10, Heft 2, p . 41, pl. 2, figs, 4, 5, 1911, 
' I CARDIOCERAS. 2 3 

Cardioceras crassum Reeside, n. sp. deeply in-cised, the second la~eral saddle broad, 

and the second lateral lobe small. Theradius Plate XII, figures 3-4; Plate XIII, figure 1; Plftte XIV, 

fi 2 • cuts the ·first lateral lo_be and touches . the gures 1; . 

1906. Ammonites (Amaltheus) cordijormis Meek and Hayden (part). Whitfield and Hovey, Am. Mus. 

Nat. Hist. Bull., vol. 22, art. 23, p.' 401, pia. 52, 

55, 56. (Not pia. 53, 54, 58, 59, 60.) 

second. 

Oardioceras crassum is marked by its vigorous 

sculpture, stout form, sharp but unpinched 

venter, and. the sutural proportions. It is possible that large specimens of Oardioceras hyatti 

Reeside, Oardioceras stantoni Reeside, and 

Oardioceras wyomingense Reeside would be like 

this species in form, but all of these have a distinctly pinched venter at the diameter of 40 

millimeters, and the suture differs. It differs 

from Oardioceras russelli Reeside in the; sharper 

venter, more vigorous and more persistent 

sculpture, and the form of the younger who:ds. 

All speCimens from Sundance formation. 

Localities: .Type, T. 25 N., R. 78 W., Freezeout Hills, Carbon County, Wyo. Locality half 

a mile northeast of the Johnson horse ranch, 

Belle Fourche River, 25 miles west of Sundance, 

Crook County, Wyo., 2 specimens. 

Cardioceras crookense Reeside, n. sp. 

shell stout; venter sharp at all stages seen 

but not pinched. Cross section of whorl at the 

diameter of 4;0 millimeters is wider than high; 

fiat subparallel sides and swollen venter; width 

three-fourths the height. Where the diameter 

is greater than 7 5 millimeters the cross section 

is rounded-triangular and width and height are 

subequal. The type at the dia:r_neter or' 150 

millimeters preserves nearly one-fourth whorl 

unseptate; aperture unknown. Umbilicus 

narrow, in width half the height of the outer 

whorl. Umbilical shoulders at the dia~eter of 

40 milliweters rounded; at the diameter of 55 

millimeters they' have short steep inner slope; 

at the diameter of 75 millimeters they are sharp 

and have high steep inner slope; in older stages 

they overhang. · 

Sculpture regular, relatively coarse through- Plate IX, figures 2

-4. 

out stages seen. - From the diameter of 40 Shell compressed; venter somewhat swollen, 

millimeters to the diameter of 75 millimeters sharp, and but little pinched; flanks well 

irly high rounded ribs rise at the umbilic l rounded. Cross section of whorl higher than 

shoulders, pass nearly straight to the middle of wide. Type entirely septate; living chamber 

the flank, bifurcate, pass·to the venter, and then and aperture unknown. Umbilicus relatively 

bend forward to form sharp chevrons at the narrow, width one-half the height of the outer 

middle line. There are few intercalated ribs whorl. Umbilical shoulders well rotinded and 

on these earlier whorls. Between the diameters inner slope short and steep. 

of 75 and 125 millimeters the primary ribs are Sculpture .is characteristic in its irregularity. 

at first high and coarse, the point of bifurcation Rou~ded but distinct ribs rise within the 

is closer to the ill:nbilicus, and there are single umbilicus. A few pass to the keel as single 

intercalated secondary ribs between the p~irs ribs, but most pa8s about one-third the dis-: 

that branch from the primaries. Toward the tance toward the keel and bi~urcate. Here and 

end of the septate portion the umbilical ribs there a secondary divides again immediately, 

gradually decrease in height until they dis- forming an irregular trifid bu:o.dle.· A few inappear. The secondaries continue distinct to tercalated secondaries are presen;t; and· rarely 

the end of the part of the living chamber pre- a ·seconda_ry _bifurcates at the keel. Most of the · 

served, though progressively lower . in height. ribs bend gently forward on the venter. The 

The last septate whorl of the type has 15 um- whole system of ribs has an 'irreghlar wavy ap~ 

bilical ribs and 45 secondary ribs. A specimen pearance. There are 29 umbilical ribs on the 

230 millimeters in diameter preserves nearly . last whorl of the type and about 60 at the keel. 

three-fourths of . a whorl unseptate. The The suture has a widesiphonal ·lobe; wide 

youngest part is practically smooth. first lateral saddle; rather narrow triangular 

The sutural' details of the type are difficult first lateral lobe, which is about equal in leng~h 

to see, though the proportions are clear. The to siphonal-lobe; wide, shallow second lateral 

siphonal lobe is broad . and short, the first saddle; short second' lateral lobe; and very 

lateral saddle about as broad as the siphonal small ·auxiliaries. The radius just cuts the 

lobe, the first lateral lobe long, narrow, ·and first lateral lobe. 
24 SOME AMERICAN JURASSIC AMMONITES. · 

. Oar,dioceras crvokense is b~st marked by the 

rounded flanks of its whorls, the peculiar sculpture, and the suture. Among related forms it 

differs from Oardioc~ras cordiforme (Meek and 

Hayden) in its, relatively swollen whorls, less 

regular sculpture, and the. wide first lateral 

saddle of its suture; from Oardioceras wyomingense Reeside in the narrow umbilicus and 

irregular sculpture. It is distinguished from 

Oardioceras excavatum (Sowerby) Nikitin 1 by 

the lesser pinching of the v~n ter and narrow 

umbilicus; from Oardioceras excavatum (Sowerby) Lahusen,Z Oardioceras excavatum (Sowerby) Bukowski,S and from Oardioceras excavatum · (Sowerby) Borissjak 4 by its irregular 

sculpture and less pinched venter; from Oardioceras cordatum (Sowerby) var. c De Loriol 5 

by its irregular sculpture. From Oardioceras 

cordatum (Sowerby) var. d De Loriol 6 it is 

scarcely distinguishable in form and sculpture. 

All specimens from Sundance formation. 

· Localities: Type and one other small specimen from locality 4 miles west of "Devils 

Tower, Crook County, Wyo. Fragmen_t from 

T. 25 N., R. 78 W., · Freezeout Hills, Carbon 

County, Wyo. 

Cardioceras distans (Whitfield) Neumayr. 

Plate XV, figures 18-21; Plate XVI, figures 1-6. 

1880. Ammonites cordijormis var. distans Whi,tfield, Report on the geology and resources of the Black 

Hills of Dak;ota: U. S. Geog. and Geol. Survey 

Rocky Mtn. Region, pp. 380-381, pl. 6, fig. 25. 

1883. · Cardioceras distans (WhitfiE;lld) Neumayr, K. Akad. 

Wiss. Wien Denkschr., Math.-Naturwiss. Classe, 

Band 47, p. 302. 

Whitfield's description is as follows: 

Shell of moderate size, slightly cordiform in a transverse 

section of the volution, or approaching subquadrate, and 

distinctly keeled on the back. Volutions few in number, 

largely embracing, leaving a broad, open umbilicus equal 

to nearly one-third of the diameter of the entire shell, and 

in which is exposed less than one-third of the breadth of 

each volution. Umbilical margin of the volution ab1 Nikitin, S. N., Die Jura-Ablagerungen zwischen Rybinsk, Mologa, 

und Mischkin an der oberen Wolga: Acad. sci. St.-Petersbourg Mem., 

7thser., vol. 28, No.5, p. 52, pl. 2, fig.13, 1881. 

2 Lahusen, Joseph, Die Fauna der jurassischen Bildungen des Rjasanschen Gouvemements: Com._geol. [Petrograd) Mem., vol. 1, No.1, p. 48, 

pl. 5, fig. 2, 1883. 

3 Bukowski, Gejza, Uberdie Jurabildungen von Czenstochauin Polen: 

Beitr. Paliiontologie Oesterr.-U.agams u. des Orients, Band 5, p. 130, 

pl. 26, figs. 21, 22, 1887. 

4 Borissjak, A., Die Fauna des Donez-Jura: Com. geol. [Petrograd] 

Mem., new ser., livraison 37, p. 7, pl. 1, fig. 6, 1908. 

6 De Lorioi, Perceval, Etude sur les mollusques et brachiopodes de 

l'Oxfordien (zone a Am. renggeri) du Jura bemois, I: Schweizer. paleont. 

Gesell. Abh., vol. 25, No. 6, p. 16, pl. 2, fig. 8, 1898. 

e Idem, p. 17, pl. 2, fig. 9. 

ruptly r6unded, sides depressed convex, somewhat flattened in the middle, slightly decreasing in convexity and 

thickness outward to about the outer fourth of the width, 

beyond w · ch point the surface abruptly deClines to the 

thickened, rounded, and prominent dorsal keel. 

Surface marked by strong, distant, and abruptly elevated flexuous ribs or ridges, with concave interspaces; 

originating a little outside of the umbilical .margin, they 

are directed, with a slight sigmoidal curvature, nearly 

across the volution, and are then directed abruptly forward along the dorsal slope-for a distance nearly equal to 

one-half the width of the volution at the point of their 

occurrence, and form strong, wavelike' ridges in crossing 

the dorsal carina. On the dorsal slopes there are also one 

and sometimes two additional ridges between the primary 

ones but usually of a little less strength. The primary 

ridges sometimes form pointed spinelike nodes at their· 

intersection with the intermediate or secondary ones, but 

this is not a constant feature, even on the same shell, and 

only occu.rs when the primary ridge dies out or bifurcates, 

forming two secondaries on the slope, which is an exceptional feature. Lines of growth apparently coincident 

with the iidges or nearly so. 

Shell compressed, venter obscurely shoul~ 

· dered and with pinched keel. Cross section of 

whorl about as high as wide, subquadrate. 

Living chamber and aperture unknown. Umbilicus moderately wide with rounded shoulders; width of umbilicus four-fifths the height 

of last whorl. 

Sculpture consists of high, sharp umbilical 

ribs (18 on last whorl of type), which bifurcate 

at about the middle of the flanks and bend 

sharply forward on the venter, producing a 

coarsely nodose keel. 

The suture of the type shows a fairly wide 

ventral lobe, wide first lateral saddle, relatively · 

narrow first lateral lobe, narrow second lateral 

saddle, and small second late~:al lobe. The 

radius just cuts the first lateral lobe. 

The most characteristic features are the stout 

form, coarse sculpture, and moderately wide 

umbilicus. 

Most of the specimens referred to this species 

show more ribs to the whorl than the type but 

otherwise agree very well. A large specimen 

(Pl. XVI, figs. 1-3) has very pronounced sculpture, more secondaries than the type, and rather 

abrupt umbilical shoulders. The suture of 

this specimen has .a first lateral saddle but 

little wider than the first lateral lobe. 

Oardioceras distans (Whitfield) is distinguished from 'Oardioceras wyomi'T}gense Rooside by its stouter form, coarser sculpture, and · 

more · rounded umbilical shoulders; from Oardioceras stantoni Reeside by its stout form and 
CARDIOCERAS. 25 

narrower umbilicus; from Oardioceras auroraense Reeside by its more compressed form; 

from Oardioceras russelli Reeside by its coarser 

sculpture and pinched venter. · .It differs from 

Oardioceras vertebrale (Sowerby) Boden1 in it~ 

less quadrate cross section and more vigorou~ 

sculpture; from Oardioceras rouilleri (Nikitin) 

Lahusen2 in the more rounded ribs and lesser 

anterior inclination of the secondaries but is 

very close; from Oardioceras cordatum (Sowerby) Lahusen3 in having fewer ribs and le~s 

compressed form; from Oardioceras rouilleri 

· · Nikitin4 in having a wider umbilicus and 

fewer secondaries. Oardioce'ras cordatumnikitinianum Bukowski5 has less vigorous 

sculpture. Oardioceras cordatum (Sowerby) 

var. f De Loriol6 has more quadrate whorls 

and finer sculpture; Oardioceras vertebrale 

(Sowerby) var. a Ilowaisky7 is hardly separable except that the whorls are stouter. 

Oardioceras cordatum (Sowerby) refigured by 

Maud Healy8 is ve-ry close to Oardioceras 

distans. 

Cardioceras distans (Whitfield) var. depressum Reeside, 

n. var. 

·Plate XV, figures 22..:..24 ; Plate XVI, figures 7-11. 

. The · variety here named depressum differs 

from the typical Cardioceras distans only in t~e 

distinct depression of the whorls. T~e rat~o 

of height to width in Cardioceras d~stan,s IS 

about one to one; in the variety it is four to 

fivB. 

Cardioceras vertebrale (Sowerby) var. a Dowaisky 9 is scarcely separable. · 

Localities: Type, Sundance formation, half a 

mile northeast of the Johnson horse ranch, 

Belle Fourche River, 25 miles west of Sundance, 

Wyo.; also five. small specimens. and many , 

fragments. Sundance formation, 4 miles west 

of Devils Tower, Crook County, Wyo.; three 

small pec~ens. Sundance formation, 4 miles 

west of Sundance, Crook County, Wyo.; one 

specimen. _Sundance formation, SundanceAladdin stage road at south fork of Redwater 

Creek, Crook County, Wyo.; 11 sma? specimens. Lower part of Naknek formatiOn, east 

Localities: Type, Sundance formation, 2 side of Oil Bay, Cook Inlet region, Alaska; one 

miles south of Belle Fourche River, near Bear specimen. 

Lodge Butte (Devils Tower, Crook County), 

Black Hills, Wyo. _ Sundance formation, half Cardioceras haresi Reeside, n. sp. 

a mile northeast of the Johnson horse ranch, Plate XIX, figures 4-12. · 

Belle Fourche River; 25 miles west of Sun- Shell stout, whorls depressed, venter keeled. 

dance Mountain, Crook County, 'Wyo. · Sev- Oldest whorls not seen. Cross section of 

erallarge and a number of fragmentary small whorls where the shell is less than 15 millispecimens. . meters' in diameter, oval, higher than wide; 

Lower part of Naknek formation, east side where the diameter is 15 to 25 millimeters, 

of Oil Bay, Cook Inlet region, Alaska. Four rudely hexagonal s-q.bequal in width ~n~ 

small complete and a number .of fragmentary height; where it is more than 25 milhspecimens. meters depressed, wide:t: than high, quad1 Boden, Karl, Die Fauna des unteren Oxford von Popilany in Litau: rate. Small part only of living cha~ber preGeol. u. palaeont. Abh., N. F., Band 10, Heft2, p. 37, pl.1, fig. 13,. 1911 . . served in type but another specrmen (Pl. · 2 Lahusen, Joseph, Die Fauna der jurassischen Bildungen des RJ!l,S~II·. · ') h b · t half of who:fl schen Gouvernements: Com. geol. [Petrograd) Mem., vol. 1, No. 1, p. XIX, figs. 9-12 S OWS a OU .. 

so, pl. s, figs. s, 6,1883. unseptate. Aperture unknown. Umbilicus 

: ~:~· :\t ~i!g;~~-~blagerungeii zwischen Rybinsk. Mologa, -wide, in width about 1 i times the height of 

und Mischkin an der oberen Wolga: Acad. sci. St.-Petersbourg, 7thser., the outer whorl. Umbilical sho~lders gently vol. 28, No.5, p. 56, pl. 2, fig. 17, 1881. _ d d 6 Bukowski, Gejza, Uber die Jurabildungen von Czenstoc:nau in roun e . 

Polen: Beitr. Palaontologie Oesterr.-Ungarns u. des Orients, Band 5, p. Sculpture regular. Exposed part of whorls 

~·J;· ~~'!t ;~r!::l, :Etude sur Ies mollusques et brachiopooes de- where the diameter is less than 7 or 8 m~ll~­ l'Oxfordien (zone a Am. renggeri) du Jura bernois,I: Schweizer. pale- meters smooth· where it is more than 8 IDifiiont. Gesell. Aoh .. vol. 25, No.6, p. 17, pl. 2, figs. 10, 11, 1898. ' h h'. - h hili' 1 'b :Ut 1 IIowaisky, D~vid ; L'Oxfordien et le Sequanien des gouvernements meters, S arp, Ig· UID Ca ri S are prese · de Mouscou et de Riasan: Soc. rtnp. Nat. Moscou Bull., new ser., vol. 

17, p. 268, pl. 10, fig. 31,1903. 9Ilowaisky, David, L'Oxfordien et le Sequanien des gouvernemen:s SHealey, Maud, Ammonites (Cardioceras) cordatus Sowerby: Paleon- de MoscouetdeRiasan: Soc. imp.nat.MoscouBull.,newser.,vol.11, tologia Universalis, .Pl<>. 94, 94a, 1905~ p. 268, pl. 10, figs. 31, 32, 1903. 
26 SOME AMERICAN JURASSIC AMMONITES. 

These ribs rise within the umbilicus, bend 

backward, emerge from the umbilicus, and 

pass outward in a shallow forward curve to 

the middle of the flanks. On the outer whorls 

each primary rib, at the middle of the flanks, 

is raised into a fairly high, elongated node. 

The primaries bifurcate, pass outward on the 

venter, and then bend sharply forward, decreasing in distinctness near t~e middle of the 

venter but rising in a distinct node when they 

reach the median line. The chevron formed is 

comparatively sharp, making an angle of perhaps · 50°. Intercalated secondaries are uncommon on the inner whorls but are numerous 

on the outer whorls. The last complete whorl 

of the type has 22 primary ribs and about 50 

secondaries. The unseptate specimen shows 

on the inner surface a mold of sculpture like 

that on the venter of the outer whorl of the 

type, but the venter of this specimen shows 

the keel almost continuous and like that of 

Oardioceras obtusum Reeside, with the secondaries quite distinct to the keel. There 

would be apparently 18 to 20 primary ribs 

and 50 to 60 secondary ribs on a complete 

whorl. 

The suture has a broad siphonal lobe, subequal first lateral saddle and lobe, first lateral 

lobe a little longer than siphonal lobe, relatively wide second lateral saddle, small second 

lateral lobe, and small auxiliaries. 

Oardioceras haresi is best recognized by the 

depressed, quadrate cross section of the outer 

whorls, which has the gre-atest width external, 

the sharp high primary ribs, and acute ventral 

chevrons. It is distinguished from Oardioceras 

obtusum Reeside by the sharper sculpture, 

quadrate cross section, and different sutural 

proportions; from Oardioceras ? latum Reeside 

by the presence of a keel on the younger whorls, 

stronger sculpture, and quadrate cross section. 

The writer knows no similar ·foreign· species. · 

The species is named for Mr. C. J. Hares, who 

collected the type. 

All specimens from Sundance formation. 

Localities: Type, 1! miles·north of Redwater 

Cre~k in Wyoming, near the boundary between 

South Dakota .and . Wyoming. Also several 

small specimens and fragments of larger ones. 

Locality 4 miles west of Dev~.ls Tower, Wyo.; 

one specimen. Locality in SW.i sec. 13, T. 23 

N., R. 79 W., Carbon County, Wyo.; one 

specimen. Locality half a mile east of the 

Johnson horse ranch, Belle Fourche River, 25 

miles west of Sundance, Wyo. 

Cardioceras hyatti Reeside, n. sp. 

. Plate XV, :figures 1-4. 

Shell compressed, venter pinched, keel elevated. Cross section of whorl quadrate, higher 

than wide. tast half whorl of type unseptate, 

aperture unknown. Umbilicus relatively narrow, a little more in width than half the 

height of the outer whorl. 'Umbilical wall 

rounded on inner whorls, abrupt on last 

whorl of type. 

Sculpture regular. The ribs begin at . the 

umbilical sho.ulder, pass outward in a shallow 

curve to about the middle of the flank, bifurcate and pass to the venter, where they bend 

forward and cross the keel. Some intercalated 

secondaries are present. The ribs are rather 

coarse and fairly sharp but · of even height 

from umbilicus to venter. Last whorl of type 

has 21 umbilical ribs and 55 secondaries. The 

earlier whorls appear to be smooth to the 

region where the shell, attains a diameter of 7 

or 8 millimeters. 

Suture has relatively narrow dorsal lobe, 

wide shallow first lateral saddle, triangularly 

arranged first lateral lobe, wide second lateral 

saddle, and small second lateral· lobe. Auxiliaries very small. Radius just cuts tip of first 

lateral lobe. . · 

The most diagnostic characters of this 

species are the narrow umbilicus, compressed 

form, and regular, even sculpture. 

Oardioceras hyatii Reeside differs from Oardioceras stantoni Reeside in the narrower umbilicus and coarser sculpture. · It differs from 

Oardioceras cordatum (Sowerby) Lahusen1 in 

the lack of prominences on the ribs, and in its 

stouter, more quadrate whorls but is quite 

similar; from Oardioceras cordatum-quadratoides Bukowski 2 in its coarser SGulpture and 

less quadrate cross section; from Oardioceras 

· cordatum (Sowerby) Borissjak 3 only in its 

mo~e compressed whorl; from Oardioceras 

1 Lahusen, Joseph, Die Fauna der jurassischen ·Bildungen des Rj:isanschen Gouvemements: Com. geol. [Petrograd] Mem., vol. 1, No. ,1, 

p. 49, pl. 5, fig. 3, 1883. . 

2 Bukowski, Gejza, Uber die Jurabildungen von Czenstochau in 

Polen: Beitr. Palli.ontologie Oesterr.-Ungams u. des Orients, Band 

5, p. 130, pl. 26, fig. 23, 1887. 

s Borissjak, A., Die Fauna des Donez-Jura: Com. geol. [Petrograd] 

Mein., new ser., livraison 37, p. 8, pl. i, fig. 8, 1908. 
CARDIOCERAS. 27 

cordatum (Sowerby) var. b De .Loriol 1 in its 

·stouter whorls, different suture, and abrupt 

umbilical shoulders. 

This species is named in honor of Mr. Alpheus. 

Hyatt. 

All speciinens from Sundance formation. 

Locality: Type, from Difficulty Canyon, se.c. 

31, T. 24 N., R. 80 W., Carbon County, Wyo., 

one specimen. ·Two specimens from locality 

near the Carnegie Museum dinosaur quarry, 

Jensen, Utah. · 

Cardioceras Iillooet~nse Reeside, · n. sp. 

Plate XVII, figur s 20...:..23. 

Shell compressed, venter pinched and keeled. 

Cross section of whorls at first broader than 

high, cordiform; at -the diameter of 25 millimeters, compressed, oval, with well-rounded 

flanks, higher than wide; at and above the 

, diameter of 35 millimeters; subquadrate with 

obscure ventral shoulders and flattened flanks; 

higher than wide. Three-eighths of the last 

whorl of the type unseptate; aperture 

unknown. Umbilicus moderately wide, in 

width nearly equal to the height of the outer 

whorl of type; umbilical shoulders gently 

rounded to the diameter of 25 millimeters; 

then the shoulders become sharper and the 

inner slope becomes oblique. 

Sculpture regular. Low, fairly sharp ribs 

rise within the umbilicus, bend backward, 

· emerge, ·and pass nearly straight outward, 

rising to a faint node at the middle of the 

flank. Then each primary bifurcates, one or 

two intercalated ribs rise between the bundles, 

and all bend forward and cross the keel as 

crenulations on it. · Many of the secondaries 

fork at the base of the keel. · The last whorl of 

the type has 20 primary ribs and about 80 

crenulations on the keel. On a fragment of 

a larger specimen of this species than the type~ 

the umbilical ribs end abruptly at the middle 

of the flank and are only . faintly connected 

with the three secondaries. which rise abruptly 

on the venter but are much fainter at the 

base of the keel. 

The suture is deeply incised. Siphonal 

lobe and first lateral saddle about equal in 

width; first lateral lobe narrower than first 

lateral saddle and a trifle lo~ger than the 

siphonallobe; seconq .lateral saddle wide and 

shallow; ·se ond lateral lobe and auxiliaries 

small. The radial line just cuts the first'latera1 

lobe. 

Cardiocert lillooetense is marked by its 

compressed orm, obscurely shouldered venter, 

straight pr" ary ribs, and fine secondaries. 

The large . specimen mentioned above approaches in several respects the genus Amoeboceras, but the secondary ribs are distinct to 

the keel thtugh faint and the umbilical ribs 

are rarely imple to the keel. It is distinguished fro Cardioceras americanum Reeside 

by its more distinct sculpture and swollen 

venter; fr~~ Cardioceras hyatti Reeside by 

its more roJnded ribs, more numerous secondaries and straighter p:rimaries; from Cardioceras sundancense Reeside by its more compressed form and more distinct sculpture. It 

differs from Cardioceras cordatum (Sowerby) 

var. a De Loriol 2 in the greater number of 

umbilical costae but is otherwise very close; 

from Cardioceras ·cordatum (Sowerby) Boden,S 

as :figur·ed, in the relative independence of the 

primaries and secondaries. 

Locality: Cardioceras-bearing beds at head 

of Big Creek, tributary to Chilcotin River, 

Lillooet, British Columbia; one entire and 

four broken specimens. 

Cardioceras martini Reeside, n. sp. 

Plate IX, figures 5-8. 

Shell comp.ressed, venter pinched and keeled. 

Cross section of whorl oval; flanks flattened,-

higher than wide. · Last three-eighths of whorl 

unseptate; aperture unknown. Umbilicus narrow, in width a little more than half the 

height of the outer whorl. Umbilical shoulders 

rounded to the stage where the shell attains a 

diameter of 35 1 

millimeters, and where the 

diameter js more than 35 millimeters the inner 

slope is short and steep. 

Sculpture somewhat irregular. Numerous 

low but sharp ribs rise within the umbilicus 

and run almost straight outward to the middle 

of the flank. Many of the ribs fork at this 

point; others· continue simple to the keel 

and show an intercalated secondary that 

rises at the line of forking between each pair 

of primaries. Some of the secondaries fork 

on the venter. The secondaries alLbend for1 De Lorio!, Perceval, Etude sur les mollusques et brachiopodes de 2 De Lorio!, Perceval, op. cit., p. 15, pl. 2, figs. 1-3. 

l'Oxfordien (zone a Am. renggeri) du Jura bernois, I: Schweizer. paleont. · a Boden, Karl, Die Fauna des unteren Ox!.ord von Popilany in Litau: 

Gesell. Abh., vol. 25. No. 6, p. 16, pl. 2, figs. 6, 7, 1898. Geol. u : palaeont. A'!:>h., N. F., Band 10, Heft 2, p. 35,pl.1, figs.6-8, 1911. 
28 SOME AMERICAN JURASSIC AMMONITES. 

ward to cross-the keel, which is evenly but 

finely serrate. The type has 33 primary ribs 

and . 80 secondary ribs on the last whorl. 

The suture has a moderately wide siphc;mal 

lobe; wide first lateral saddle; first lateral lobe 

relatively narrow and equal in length to the 

siphonal lobe; second lateral saddle nearly 

as wide as first lateral lobe; other lobes small. 

This species is marked . by its· compressed 

form, pinched venter, fine, sharp ribs; and 

the prqportions of the suture. It differs from 

Oardio.ceras -crookense Reeside in the finer, 

sharper ribs and greater pinching of the venter; 

from Oardioceras cordiforme (Meek and Hayden) in its finer, straighter, and less elevated 

umbilical ribs and in sutural detail, though in 

general aspect very close to this species. It is 

distinguished from Oardioceras cordatum (Sowerby) var. c de Loriol 1 in the pinching of the 

- venter and _the finer ribs; from Oardioceras 

cordatum (Sowerby) var. d De Loriol 2 in the 

niore compressed form and finer ribs; from 

Qardioceras excavatum (Sowerby) Lahusen 3 

in that the ribs are sharp and distinct; from 

Oardioceras excavatum (Sowerby) Bukowski 4 

it is scarcely separable. 

The species is named for Mr. G. C. Martin, of 

the United States Geological Survey, who collected the type. 

Localities: Type, Oardioceras-bearing beds 

along creek entering Boulder Creek from north, 

Matanuska Valley, Alaska; also four other 

specimens. Lower part of Naknek formation, east side of Oil Bay, Cook Inlet region, 

AJ.aska; four specimens. 

Cardioceras obtusum Reeside, -n. sp. 

Plate XX, figures 1-6. 

Shell swollen, depressed, venter obtusely 

keeled. Oldest whorls not seen. Cross section of whorl oval, longer than wide, where the 

diameter is less than 12 millimeters; where it is 

20 to 25 millimeters, almost circular; where-it 

is 25 millimeters, oval, wider than high. Living 

1 De Loriol, Perceval, Etude sur les mollusques· et brachiopodes de 

I' Oxfordian (zone a Am. renggeri) du Jura bemois, I: Schweizer. paleont. 

Gesell. Abh., vol. 25, No.6, p. 16, pl. 2,Jlg. 8, 1898. 

'Idem, p. 17, pl. 2, fig. 9. · 

a Lahusen, Joseph, Die Fauna der jurassischen Bildungen des Rjasanschen Gouvemements: Com. geol. [Petrograd) Mem., vol.1, No.1, p. 48, 

pl. 5, fig. 2, 1883. 

f Bukowski, Gejza, Uber die Jura bildungen von Czenstochau in 

Polen: Beitr. Palaontologie Oesterr.-Ungams u. aes Orients, Band 5, 

p. 130, pl. 26, figs. 21, 22, 1887. · 

chamber and aperture unKnown. Umbilicus 

wide, about 1! times the height of outer whorl. 

Umbilical shoulders gently rounded. 

Sculpture regular. W:Q.ere the diameter is 8 

millimeters, the exposed part of the whorls is 

smooth;. where the diameter is greater distinct, 

rounded umbilical ribs are present. These ribs 

become more elevated as the shell increases in 

size but are invariably rounded. They begin 

within the umbilicus, pass outward with a shallow forward curve to about the middle of the 

flank, where· they reach the maximum height, 

then bifurcate into. low, flattened seconda~ies. 

The secondaries bend forward to meet the keel, 

which, however, is quite smooth and even and 

practically lacks nodes. A few intercalated 

secondaries occur. There would be 25 umbilical ribs and about 60 secondaries on the complete last whorl of the type. The keel is obscure 

where the diameter is less than 20 millimeters. 

The suture shows a wide, short siphonallobe; 

first lateral saddle and first lateral lobe subequal in width and the first lateral lobe is a 

trifle longer than the siphonal lobe; second 

lateral saddle and second lateral lobe small; 

auxiliaries very small. The radius cuts the tip 

of the first lateral lobe. 

Oardioceras obtusum is best recognized by its 

depressed oval whorls; rounded primary and 

close, flattened secondary ribs; smooth rounded 

keel; and the suture. It differs from Oardioceras 

haresi Reeside in the greater distinctness of the 

ventral ribs, oval cross section, and in the wider 

superior saddle of the suture; from Oardioceras? . lat.um Reeside in the sharp venter and 

lesser depression of the younger whorls. No 

foreign forms are known to the writer which are 

quite like this species. 

All specimens from Sundance formation. 

Localities: Type, locality in Wyoming 1! 

miles north of Redwater Creek, near t:Q.e boundary between South Dakota and Wyoming; also 

one· other specimen. Locality 4 miles west of 

Devils Tower, Crook County, Wyo.; five small 

specimens. 

Cardioceras plattense Reeside, n. sp. 

Plate IX, figures 9-12. 

Shell fairly stout; venter swollen, obtusely 

shouldered, and keeled; early whorls very · 

stout, distinctly keeled. Cross section of last 

whorl about as wide as high, broadly cordiform; 

inner whorl depressed ova!. · Umbilical part of 
CARDIOCERAS. 29 

Cardioceras russelli Reeside, n~ sp. 

Plate XIII, .figures 2, 3; Plate XIV, figures 3-5. 

living chamber preserved for half whorl; aperture _ unknown. Umbili~us · moderately wide; 

in width three-fifths the height of the outer 

whorl in - type. Umbilical shoulders gently ' 

rounded to the place where the shell attains a Shell moderately compressed, discoidal; ven- · 

diameter of 25 to 30 millimeters, then rounded , ter sharp throughout life. Cross section of 

with steep inner wall. whorls invariably cordiform, higher than wide . 

Sculpture consists of low,' rounded umbilical in youth; width and height subequal in 

ribs on the exposed part of the inner whorls later stages. Only fragment of)iving chamber 

where the diameter of the shell is greater than of type known -(not shown in figures); a perabout 20 millimeters, though better preserva- ture unknown.! Umbilicus narrow; width 

tion might show them to have begun earlier. equals two-fifths height of last whorl. . UmbiliAt this qiameter distinct ventral nodes and a cal shoulde~ gently . rotinded to the region 

crenulated keel are present. The umbilical where the diameter of the shell is. about 40 

ribs continue low and rounded on the whole of millimeters; then the umbilical wall becomes 

the type,_ though on the unseptate part they steep and at a diameter of about 70 millimebecome less distinct. These umbilical ribs.. ters begins to overhang. . 

pass about one.;.fourth the distance toward the Sculpt~e . of youngest whorls unknown. 

keel and divideinto two (rarely three) second- From a diameter of 50 millimeters to a diamearies. Intercalated secondaries rise between ter of 90 millimeters fairly high rounded ribs 

the bundles. The sec9ndaries pass outwa~d to b.egin at the umbilical shoulder, pass .about 

a point three-fourths of the way from umbilicus one-third the distance toward the keel, and split 

to keel, are raised into low, elongated nodes, into two or three secondaries. A few inter!.. 

bifurcate and bend forward a little to cross the calated secondaries are present. The secondsharpened venter as crenulations. There would aries continue outward, curving gently forprobably be 15 primary ribs on the last whorl ward, to the sharp venter. On the last whorl 

of the type and 70 to 80 crenulations on the keel. · of th~ typ,e the ribs gradually become in disThe suture shows a long, narrow siphonal tinct, and above a diameter of 90 millim.eters 

lobe, wide first ·lateral saddle, relatively short the whorl is practically smooth. Umbilical 

first lateral lobe, small second lateral saddle ribs number about 20 to the whorl and secondand lobe, and a few small auxiliaries. aries about 50. 

This species is marked by its stout form, dis- The suture shows a short siphonallobe and 

tinct regular sculpture, twice-forked ribs and long first lateral lobe. The · siphoilal lobe, 

low ventral nodes, and by the-sutura:-1 propor- first.lateral saddle, first lateral lobe, and second 

tions. It differs from Oardioceras sundancense lateral saddle are about equal in width. The 

Reeside in its coarser andmore persistent sculp- second lateral lobe is small; the auxiliaries 

ture, stouter form, and in the relative length of feeble. The radius cuts the first lateral lobe 

the siphon31 lobe of the suture; from Cardia- and barely cuts the second. 

ceras wyomingense Reeside in its. nar~ower urn- Oardioceras russelli -is best marked by its 

bilicus, stou,ter form, second bifurcation of the stout form, relatively strong regular sculpture, 

ribs, and sutural detail;, fro~ Oardioceras rus- and the sutural proportions. It differs from ~ 

selli Reeside in its stouter Jorm, second bifur- Oardioceras wyomingense Reeside in the more 

cation of the ribs, and .in the relativelengths of triangular cross section of the whorl, the much 

the lobes of the suture; from Oardioceras narrower umbilicus, and in the relative length 

auroraense Reeside in·.its stouter form, ..second of the lobes of the suture-; from Oardioceras ' 

bifurca~ion of the ribs, and in sutural propor- sundancense Reeside in its vigorous and more 

tions. From related foreign species it is persistent sculpture, blunter unshouldered vansharply distinguished by the second. bifurcation. ter; and in suturat details; from Oardioceras 

Locality: Type specimen is from Sundance cordiforme Meek and Hayden , in the blunter · 

formation, "near the Grand Canyon of the venter, more elevated ribs, and in sutural 

North Platte," near Alcova, Natrona County, detail. The writer ·knows no close European 

Wyo. species. 
30 'soME MERICAN JURASSIC AMMONITES., 

The species is named for Mr. I. C. Russell, 

who with Alpheus Hyatt collected part of the 

material on which this pap.er is based. 

All specimens from Sundance formation. 

Localities: Type, in Wyoming 1! miles 

north of Redwater Creek, near the boundary. between South Dakota and Wyoming. 

ComG Bluff, near Aur()ra (Ridge), Wyo.; one 

specimen. 

Cardioceras schucherti Reeside, n. sp. 

Plate XI, figures 3-5. 

Shell stout, venter sharp and pinched. 

Oldest whorls unknown. Cross section cordiform, about as wide as high; in all stages to a 

diameter of 30 millimeters. · Living. chamber 

and aperture unknown. Umbilicus narrow 

in type, in width a little over half the height 

of the outer · whorl. Umbilical shoulders 

rounded on innermost whorl preserved, abrupt 

on outer. 

Sculpture of type a bit obscured by weathering but yet distinct. Inner whorl shows fairly 

high umbilical ribs. Outer whorl has umbilical ribs that are first stout and high but become 

low and rounded. At about middle of flank 

each rib is raised and divides into four secondaries; between some of the bundles an intercalated secondary rises. The youngest part of 

the last whorl of the type had apparently no 

secondaries. The secondaries bend gently 

forward and cross the keel as crenulations. 

There would be about 15 umbilical ribs and 80 

secondaries on a complete whorl of the type. 

The suture shows a short, broad siphonal 

lobe; broad first lat.eral saddle; long, narrow 

first lateral lobe; and wide second lateral 

saddle. Apparently there are few small auxiliaries. The radius cuts the first lateral lobe. 

Oardioceras schucherti is distinguished by the 

combination of stout whorls with cordiform 

section, subdivision of each primary into four 

secondaries, and a suture that has a short, 

' broad siphonal lobe and a long, narrow first 

lateral lobe. It differs from . Oardioceras plattense Reeside in suture and in lacking a second 

forking of the ribs; from Oardioceras wyomingense Reeside in its narrow umbilicus, stout 

form, and in the suture; from Oardioceras 

auroraense Reeside in its stouter form, more 

numerous secondaries, and sutural details; 

from Oardiocerascordiforme (Meek and Hayden) 

in fine secondaries and in sutural detail. It is 

distiri.guished from Oardioceras rouilleri Nikitin 1 by its less pinched venter an~ less persistent sculpture, though apparently very 

close; froni Oardioceras nikitinianum Lahusen 2 

by finer secondaries and stouter form. 

This species is named for the collector of the 

type, Prof. Charles Schuchert. 

Locality: Type specimen from Sundance 

formation, in T. 25 N., R. 78 W., Freezeout 

·Hills, Carbon Coun~y, Wyo. 

Cardioceras spiniferum Reeside, n. sp. 

Plate XVIII, figure 4; Plate XIX, figures 1-3. 

Shell compressed; venter much pinched, 

sharp. Earlier whorls unknown. Cross section · of whorls sharp, cordiform. Last half 

whorl unseptate; aperture unknown. Umbilicus narrow, in width about half the height 

of the outer whorl. Unbilical shoulders sharp 

and inner slope steep on the part of type preserved. 

Sculpture of inner whorl marked. Distant, 

sharp umbilical ribs rise within the umbilicus, 

pass about halfway to the keel, rise into a . 

high, sharp node, and then divide into two or 

three secondaries. Between each of these 

groups two or three .intercalated secondaries 

are present. All bend forward on the venter 

to the· base of the keel and subdivide into fine 

striae-like ribs on the keel. The unseptate 

part shows low, rounded, distant swellings 

near the middle of the flank, which have 

coarse striae on and between them. 

Entire suture not seen. Siphonal lobe 

apparently long and narrow, first lateral 

saddle narrow, first lateral lobe longer than 

the siphonal lobe and wide, second lateral 

saddle and lobe relatively short and narrow. 

Suture deeply incised. 

The compressed form, pinched venter, and 

high nodes at point of division of the primary 

ribs mark this species. It is closely related 

to Cardioceras canadense Whiteaves but so far · 

as may be judged differ~ in the pinching of the · 

venter, the position of the second forking, 

and the proportions of the suture. However, 

as the spe9imens of Cardioceras canadense 

available for comparison are all small, the 

1 Nikitin, S. N., Die Jura-Ablagerungen zwischen Rybinsk, Mologa, 

und Mischkin an der oberen Wolga: Acad. sci. St.-Petersbourg Mem., 

7th ser., vol. 28, No.5, p. 56, pl. 2, fig. 17, 1881. 

2 Lahusen, Joseph, Die Fauna der jurassischen Bildungen des Rjasanschen Gouvemements: Com. geol. [Petrograd] Mem., vol. 1; No. 1, p. 

50, pl. 5, fig. 7, 1883. 
·CARDIOCERAS. 31 

differences inay not hold. The coarsely sharper sculptur-e . and less numerous secondstriated keel and high nodes 'separate Oardio- aries; froni Oardio(}eras vertebrale (Sowerby) 

ceras spiniferum from Oardioceras distans with var. alta Boden 3 by its relatively finer sculpwhich it is associated. The writer knows no ture. . . 

close European species: This species is named for Mr. T. W.· Stanton, 

. Locality: Type, lower part of Naknek for-· who collected the type. 

mation, east side of Oil Bay, Cook Inlet All specimens from Sundance' formation. 

region, Alaska. Localities: Type, Difficulty Canyon, sec. 31, 

Cardioceras stantoni Reeside, n. sp. 

Plate XV, figures 5-8. 

1906. Ammonites (Amaltheus) cordijormis Meek and Hayden (part). Whitfield and Hovey, Am:. Mus. 

Nat. Hist. Bull.,· vol. 22, art. 23, p. 401, pl. 60, 

·figs. i, 2; pl. 53. 

T. 24 N., R. 80 W., Carbon County; Wyo.; 

also five other specimens. Aurora (Ridge), 

Wyo.; Alpheus Hyatt and I. C. Russell, collectors; one specimen. Como Bluff, near 

Aurora, Wyo.; Charles Schuchert, collector; 

one specimen. 

Cardioceras stantoni var. obesum Reeside, n. var. 

_ Plate XV, figures 9-11. 

Shell much compressed, discoidal; keel sharp 

and high. Qross section . of whorl · rudely 

hexagonal, higher than wide. Last half whorl 

of type unseptate, aperture unknown. Umbili- . This variety differs from the typical form 

cus wide, a little greater . in widt_h. than the only in its stouter whorls. 

height of the outer whorl. Umbilical shoulders Locality: Sundance formation, Difficulty 

gently rounded. Canyon, sec. 31, T. 24 N., R. 80 W., Carbon 

Sculpture consists of high, s~arp ribs, County, Wyo. 

which bend backward slightly on the umbilical 

wall, then pass outward, curving a little forward, to the middle of the flank, increasing in 

. elevation. Then each rib forks, the branches 

passing outward to the venter where they are 

elevated and bend forward to form crenulations on the keel. Some intercalated secondaries are present. The iast whorl of the type 

has 24 umbilical ribs and 55 ventral ribs. 

The suture shows a relatively narrow 

siphonal lobe, wide superior saddle, relatively 

narrow triangular first lateral lobe, small second lateral lobe, and feeble auxiliaries. 

The compressed form, wide umbilicus with 

rounded shoulders, ·and sharp sculpture best 

mark this species. 

Oardioceras stantoni · Reeside differs from 

Oardioceras wyomingense Reeside in its coarser 

and more elevated ribs and rounded umbilical 

shoulders; from Oardioceras hyatti Reeside in 

, its wider umbilicus. It is separable from 

Oardioceras vertebrale Sowerby var. densiplicata Boden 1 by its greater compression and 

.less nodose sculpture; from Oardioceras quadradtoides Nikitin 2 by its coarser, apparently 

1 Boden, Karl, Die Fauna des unteren Oxford von Popilany in Lftau: 

Geol. u. palaeont. Abh., N _F., Band 10, Heft 2, p. 37, pl. 1, fig. 14, 1911. 

2 Nikitin, S. N., Die Jura-Ablagerungen zwischen Rybinsk, Mologa, 

und Mischkin an der oberen Wolga: Acad. Sci. St.-Peters bourg Mem., 

7th. ser,l vol. 28, No.5, p. 58, pl. 2, fig. 20, 1883. 

Cardioceras stillwelli Reeside, n. sp. 

Plate VI, figures 11-14 . 

Shell compressed, ven te:r pinched slightly 

and keeled, flanks rounded. Cross· section of 

whorl gently rounded, higher than wide. Living chamber and aperture unknown. Umbilicus narrow, about half the height of outer 

whorl in width; umbilical shoulders gently 

rounded. . 

Sculpture feeble. Exposed part · of early 

whorls smooth. Last who:d of type entirely 

septate and has 10 hroad, low umbilical ribs, 

which are obscure at the beginning of the last 

whorl but become quite distinct. The interspaces are smooth or show fine striae. The 

umbilical ribs reach the iniddle of the flank and 

split into four or five fine, low, . crowded 

secondaries, between which other fine intercalated secondaries rise, so that the venter 

shows about 12 secondaries to each primary 

rib. Many of the secondaries divide before 

reaching the keel so that it shows very fine 

crenulations. 

The suture shows a very broad siphonallobe, · 

first lateral saddle wider than the first lateral 

lobe, which is relatively narrow, small second 

a Boden, Karl, op. cit., p. 38, pl. 1, fig. 15. 
SOME AMERICAN JVRASSIC AMMONITES. 

lateral saddle and lobe, and feebly developed 

auxiliaries. The radius just cuts the tip of the 

first lateral lobe. 

This species is based on a single broken specimen but is quite distinct in its peculiar sculpture, rounded umbilical shoulders and flanks, 

and sutural characters. 

. Oardioceras stillwelli differs from Oardioceras 

whitjieldi Reeside, its nearest American relative, by its rounded flanks, fine sculpture, and 

in sutural detail. It · differs from Oardioce'ras 

sche71wieni Boden 1 in the rounded umbilical 

shoulders, less 'defined umbilical ribs, and par-

. ticularly in the absence of nodes on the venter; 

from Oardio~eras excavatum (Sowerby) var. 

. laevigata Boden 2 in the feeble sculpture, 

rounded umbilical shoulders, and narrower 

umbilicus. 

The species is named for Mr. L. W. Stillwell, 

who collected the type. 

Locality: Type, Sundance formation, 4 miles 

west of Sundance, Crook County, Wyo.; one 

speCimen. 

Card.ioceras sundancense Reeside, n. sp. , 

Plate XVII, figures 1~-19; Plate XVIII, figures 1-3. 

the line of involution but become higher with 

age; from a diameter of 20 millimeters to a 

diameter of 40 millimeters the umbilical wall 

is parallel to the axis of the shell; where the 

shell is more than 40 millimeters in diameter 

the umbilical wall overhangs the line of involution. In the plane passing through the last 

septum of the type the width of the umbilicus 

is one-third the height of the outer whorl. 

Sculpture of younger whorls of type seen 

only on umbilical shoulders to a diameter of 75 

millimeters and consists of low, rounded 

primary _ribs, about 20 to the whorl. The 

first half of the last septate whorl of type 

shows on the flanks nine broad, low, rounded, 

evenly spaced ribs, or rather swellings, which· 

decrease in distinctness toward the living 

chamber. These primary ribs· seem to divide 

into two or three very low secondary ribs, and 

between each group one or two intercalated 

secondary ribs rise. The secondary ribs pass 

to the venter, are raised into blunt, elongated 

nodes, and bifurcate agai.n, the resulting 

tertiary ribs passing forward and across the 

keel as crenulations. The younger half of the 

last septate whorl 8Jld the living chamber are 

smooth except for fine sigmoid striae and 

'S:P.ell compressed, discoidal. Where the di- crenulations on the keel. On young indi vidameter is 10 to 16 millimeters the shell is uals of t:q.is species the keel and the ribs seem 

much compressed, keeled, and has quadrate to arise at a diameter of about 10 millimeters 

cross section of whorls; where the diameter is and to show above this diameter about the 

25 to 35 millimeters the whorl is stout, subequal same characters. Low but distinct umbilical 

in width and height, and has ventral shoulders. ribs within the umbilicus bend backward 

At 35 millimeters in · diameter the venter is slightly, then emerge and pass almost radially 

distinctly pinched near the keel. At 50 milli- to the middle of the flank, where they are · 

meters in diameter the whorl is cordiform, and elevated a .trifle and bifurcate. . Many interthe ventral shoulders are still discernible. calated ribs rise at the middle line of the flanks. 

Where the ·shell is more than 60 to 70 milli- All the secondaries pass to the venter, are 

meters in diameter the whorl is cordiform, and raised into blunt nodes, and many then divide. 

width and height are subequal; sides evenly All bend forward and cross the keel. On a 

curved to the keel, which is sharp and crenate. specimen 40 millimeters in diameter there are 

A specimen preserving the early whorls where 20 primary ribs to the whorl and about 100 

the diameter is less than 10 millimeters shows crenulations on the keel. 

them to be first much depressed, then ·almost The sutur·e is marked by a short, broad 

circular in cross section. · siphonallobe; first lateral saddle broader than 

The type preserves three-eighths of the last the first lateral lobe; first lateral lobe long and 

whorl unseptate and scaphitoid. Aperture un- relatively narrow; second lateral lobe and 

known. saddle small;· auxiliaries feebly developed. 

Umbilicus in the younger whorls moderately The radius cuts both the first and second 

wide. Umbilical shoulders gently rounded to lateral lobes. _ Oardioceras sundancense Reeside is character- 1 Boden, Karl, Die Fauna des unteren Oxford von Popilany in Litau: ized by the regular but rather weak sculpture Geol. u. palaeont". Abh., N. F., Band 10, Heft 2, p. 36, pl. 2, fig. 3, 1911. 2 Idem, p. 40, pl. 2, fig. 2. , of the younger whorls, in particular by the 
CARDIOCERAS. 33 

second bifurcation and the fine ribs on the 

venter, by the form of the adult whorl, and by 

the short siphonal lobe of the suture. It 

differs from dardioceras canadense Whi tea ves 

by its less elevated ribs and by its different 

sutural proportions; from Oardioceras whiteavesi 

Reeside by its finer ventral ribs and different 

sutural proportions. It is distinguished from 

Oardioceras sc~ellwieni Boden1 by the more 

numerous primary and secondary ribs and by 

sutural details ; from Oardioceras kokeni Boden 2 

by the presence of relatively large primary 

ribs; from Oardioceras cordatum (Sowerby) 

Boden 3 by the fine, even ventral ribs; from 

Oardioceras excavatum (Sower by) Boden 4 by 

the fine ventral ribs and crenulated keel. 

·aardioceras dieneri Neu+ann-Borissjak5 seems 

to agree in form with some very small individ-

·uals of our species. pardioceras excavatum 

(Sowerby) as refigured by Healey6 is practically 

indistinguishable. I 

Locality: Type and ~ve smaller ~pecimens 

from Sundance format~on, 4 miles west of 

Sundance, Crook Co~nt , Wyo. 

Cardioceras whiteavesi Reeside, n.. sp. 

I . 

Plate XVII~ figures 1-4. 

Shell moderately compressed, venter shouldered and keeled. Cross section of whorl 

rudely hexagonal, wider than high. Final fiveeighths of last whorl unseptate; aperture unknown. Umbilicus wide, about equal in width 

to the height of the outer whorl. Umbilical 

shoulders rounded but have a steep inner slppe. 

Sculpture relatively coarse: Keel appears at · 

diameter of about 8 or 9 millimeters. Earlier 

whorls smooth on exposed flanks to <fiameter 

of 15 millimeters, then umbilical ribs appear, 

which soon become high and sharp and rise into 

a sharp node within the middle line of the 

1 Boden, Karl, Die Fauna des unteren Oxford von Popilany in Litau: 

Geol. u. palaeont. Abh., N. F., Band 10, Heft 2, p. 36, pl. 2, fig. 3, 1911. 

2 Idem, pl. 1, figs. 9, 10. 

a Idem, p. 35, pl. 1, figs. 6, 7. 

• Idem, p. 38,.pl. 2, fig. 1. 

o Borissjak, A., Die Fauna des Done1.-Jura: Com. geol. [Petrograd) 

Mem., new ser., livraison 37, p. 9, pl. 1, fig. 10, 1908. 

6 Healey, Maud, A mmonues ( Cardioceras) excavatus Sowerby: Paleontologia Universalis, pls . . 92, .92a, 92b, 1905. 

flank. Each of the primary ribs then divides 

into two low secondaries, which then rise intb 

high nodes on the· venter. and continue from 

these obliquely for)Vard to the keel, upon which 

they produce coarse crenulations. Some intercalated secondaries · are present, and some of 

these divide on the venter. The ribs are 

clearly visible to the keel. The .last whorl of 

the type has 15 umbilical ribs, and there would 

be about 56 crenulations on the keel. 

The suture has a relatively long, wide 

siphonallobe; wide first lateral saddle; short, 

narrower first lateral lobe, small second lateral 

saddle and lobe, and very feebly developed 

auxiliaries. 

Oardioceras whiteavesi Reeside is best characterized by its coarse nodose sculpture and the 

very long, wide siphonallobe of the suture. It 

is .closest to Oardioceras canadense Whiteaves 

but is sharply differentiated by its coarse ventral ribs, coarsely crenula ted keel, and r long 

siphorial lobe. It differs from · Oardioceras , 

vertebrale (Sowerby) var. duplicata Boden 7 in 

having fewer umbilical ribs and more pronounced nodes; from Oardioceras vertebrale 

(Sowerby) Nikitin 8 in the more inward position 

of the primary nodes, the finer secondaries, due 

to greater forking and more numerous intercalation of secondaries; from Oardioceras 

zenaidae Ilowaisky 9 in its coarser ventral ribs. 

Oardioceras vertebrale (Sowerby), Tefigured by 

Healey/0 is stouter but, is very like Oardioceras 

whiteavesi in general expression. 

This species is named for Mr. J. F. Whiteaves, Canadian paleontologist. 

Localities: Type, Sundance formation, 4 

miles west· of Sundance, Wyo.; one specimen. 

Oardioceras-bearing beds, Lillooet, British 

Columbia; one fragment indistinguishable 

from this species. · · 

7 Boden, Karl, p. 37, pl. 1, fig. 14. 

s Nikitin, S. N., Die Jura-Ablagerungen zwischen Rybinsk, M.ologa, 

tmd :Mischkin an der oberen Wolga: Acad. sci. St.-Petersbonrg Mem., 

7thser., vol. 28, No.5, p. 57, pl. 2, fig.18, 1881. 

9 Ilowaisky, David, L'Oxfordien et le Sequanien des gouvernements 

de Moscou et de Riasan: Soc. imp. nat. Moscou Bull., new ser ., vol. 17, 

p. 269, pl. 10, figs. 33-36, 1903. 

10 Healey, Maud, Ammonites ( Cardioceras) vertebrale Sower by: Paleontologia Uruversalis, pis. 93, 93a, 93b, 1905. ' 
34 SOME AMERICAN JURASSIC AMMONITES. 

Cardiocera8 whltfieidi ·Reeside, n. sp·. 

Plate VI, figures 1-6. 

'referred the type . specimen . . It differs from 

Oardioceras americanum Reeside in its steeper 

walled, narrow umbilicus and finer sculpture 1880· Ammonites cordijormis Meek and Hayden (part). on flanks and keel; from Oardioceras stillwelli 

Whitfield, Report on the geology and resources of 

the Black Hills of Dakota, u. s. Geog. and Geol. Reeside in its more vigorous sculpture, steeper 

Survey Rocky Mtn. Region, pp. 378-380, pl. 6, walled Umbilicus, and the less oval cross section 

figs. 20-24. of the whorls._ It may be distinguished from 

1910. Cardioceras cordijorme (Meek and Hayden) (part). Oardioceras tenuicostatum Nikitin 1 by the 

Grabau and Shimer, NorthAmericanindexfossils·, · · d' 1 · I 2 185 fi 1442 t

. 't (N ··t g~eater compressiOn at same 1ameter, ess vo . , p. , g. , en Ire ammon1 e. o ' . . fig. 1442, suture.) ·'; pillching of the venter, and more finely crenush 11 d d. 'd 1 t . .. . Iated keel; from Oardioceras excavatum (Sowere compresse , 1scm a, ven er .some- b ) Za · t B d 2 b th · h · · h d d h 1 k 1 d Fl . k f Y var. emga um o en y e nonper- w at p1nc e an s arp y ee e . an s o · . t f th · 'b f n d · h 1 · · 1 1 · d SIS ence o e pnmary r1 s; rom var wceras 

w or s ill younger stages near y pane an ··7 : B d 3 nl b 't 

b ll l

. • 1 1 1 f pOJ!1;tfWnense o en o y y 1 s narrower 

su para e, ill ater stages neary pane rom · bil' I · h d t d t f 

umbilical shoulder to keel. ·cross section of ~a: ICus, etss pl~c tae t ven(Ner!k~nt. )suEur~; _rok~ 

h l · 1 h h' h h oar wceras enu~cos um 1 1 ill onssJa . type s arp y tr1angu ar, muc 1g er t an b th :fui 1 ul t d k I d ll 

wide. . Living chamber and aperture unkp.own. Y t e e Y cren a e ee an more swo en 

U b m 1 .li b fif h h · h f ven er. · cus narrow, a out two- t s mg t o . . 

t h 1 · 'd.th U b'l' 1 h ld t Locality: Type, Sundance formation, tops of ou er w or ill W1 . · m 1 ICa s ou ers o . h . . . 

d. t f 30 'llim t f th. h II hig est hills, 2 miles south of Belle Fourche a 1ame er o m1 . e ers or e s e . . 

d d 'th t b ·t · 1 Wh R1ver, near Bear Lodge Butte (Devils Tower), roun e _ W1 ou a rup Inner s ope. ere W · 

the diameter is more than 30 millinleters the yo. 

inner slope ~s abrupt and the shoulder quite 

sharp; it begins to overhang slightly at a diameter of about 65 millimeters. 

Sculpture feeble. Early whorls at a diameter of perhaps 10 millimeters show distinct 

ribs on the flanks. These ribs are visible to a 

diameter of about 45 millimeters and number 

25 to the whorl. The final third of the last 

whorl of ·the type preserved shows only striae 

on the umbilical shoulders and inner part of 

the flank At one-third the distance out from 

the umbilicus the primary ribs divide into two 

or three secondaries,. which pass radially for 

another third, bend forward to the base of the 

keel, and there bifurcate. The keel is crenulated by these fine, crowded tertiary ribs. 

All the secondary ribs are irregular ail.d indistinct on the final third of the last whorl of the 

type. 

The sut~e has a wide, short siphonallobe; 

first lateral saddle and first lateral lobe subequal; second lateral saddle and second. lateral 

lobe small; auXiliaries feebly developed. The 

radius just touches the tips of the first and 

·second lateral lobes. 

Oardioceras whitfieldi Reeside is marked by 

its triangular cross section, narrow, deep umbilicus, suppressed sculpture, and its suture. It 

is quite distinct from Oardioceras cordijorme 

Meek ~nd Hayden to which species Whitfield 

Cardioceras wyomingense Reeside, n. sp. 

Plate XV, figures 12-17. 

Shell much compressed, venter pinched, keel 

elevated. Flanks of whorls flattened; venter a 

little swollen, producing an obscure shoulder. 

Cross section of whorl higher than wide. Living 

chamber and aperture unknown. Umbilicus 

about equal in width to height of outer whorl. 

Umbilical wall rather abrupt. 

Sculpture very regular. Distinct sharp ribs 

'start within the umbilicus, pass straight or 

slightly curved to the middle of the flank, 

bifurcate there and pass to the venter, then 

bend forward and cross the keel as low crenulations. A few intercalated secondaries are 

present, and some of the secondaries bifurcate 

again near the keel. There are about 25 

umbilical ribs and about 60 secondaries to the 

whorl at a diameter of 45 millimeters. 

The inner whorls of the type are not well 

preserved. On another specimen referred to 

'this species (Pl. XV, fig. 17) the exposed parts 

of the flanks are smooth to a diameter of more 

1 Nikitin, S. N., Die Jura-Ablagerungen zwischen Rybinsk, Mologa, 

und Mischkin an der oberen Wolga: Acad. sci. St.-Petersbourg Mem., 

7th ser., vol. 28, No. 5, p. 57, pl. 2, fig. 19, 1881. 

2 Boden, Karl, Die Fauna des unteren Oxford von Popilany in Litau: 

Geol. u. palaeont. Abh., N. F., Band 10, Heft 2, p. 40, pl. 2, fig. 2, 1911. 

a Idem, p. 41, pl. 2, figs. 4, 5. 

4 Borissjak, A., Die Fauna des Donez-Jura: Com. geol. .[Petrograd] 

Mem., new ser., livraison 37, p. 10, pl. 1, fig. 12, 1908. 
CARDIOCERAS. . 35 

than 6 millimeters for the shell. No keel is from Cardioceras cordatum (Sowerby) Boriss: 

visible on the cross section until a diameter of jak 6 in wider umbilicus and more compressed' 

9 millimeters is reached, where the ribs on the form. Cardioceras excavatU'lfJ (Sowerby) var. a 

flanks are also plain. These earlier whorls are ~orissjak 7 is scarcely separable, according to 

almost round in cross section, At the diam- the figure. · 

eter of about 13. -millimeters the~ cross section All specimens from Slindance formation. 

of the whorl is .higher than wide and the Localities: Type, · Aurora (Ridge), Wyo.; 

umbilical wall is narrow but .distinct . .. ,4- large also several other fragments. Locality 3 :pailes 

septate fragment referred to this species .(PL southwest of Barrett .(Aladdin), Crook County, 

XV, fig. 14) shows the same sort of sculpture, Wyo.; one small and fragment of large specirather high umbilical ribs but more inter- men. Locality 4 miles west of Devils Tower 7 

calated secondaries than occur on the type: Crook County, Wyo.; two small ·specimens. 

The suture of the septate fragment (Pl. XV, Locality in T. 25 N.,. R. 78 W., Freezeout Hillsj. · 

fig. 14) shows a long, relatively narrow siphonal Carbon ·County, Wyo.; five fragments of larg~ · 

lobe; wide superior saddle; narrow first lateral specimens. Locality 4 miles west of Sun-~ 

lobe, shorter than the siphonallobe; small sec..; dance, Wyo.; one small specimen. Locality 

ond lateral saddle and second lateral lobe; and near Yampa River 0), Colo.; fragments ~ 

poorly developed au.~iliaries. The radius just · 1 

cuts the first lateral lobe. Cardioceras? albanitmse Reeside, n. sp. 

The material on which this species is founded Plate XXIII, figures:l-3;. Plate XXIV, figures 1, 2. 

is fragmentary, but the great compression of A single large specimeil, practically complete, 

the shell, the flat flanks, the width ·of the urn- has a robust form and the venter· harrowly 

bilicus, and the sculpture form a. group of char- rounded. Cross section of outer whorls .,about 

acters deserving of a distinctive name. as wide as high, cordiforme. Whorls t ·not 

Cardioceras wyomingense Reeside is distin- . known below 80 millim.eters. Living chamber· 

guished from Cardioceras stantoni Reeside by occupies three-fourths <?f last whorl, scaphitoid; 

its sharper umbilical shoulders, finer ribs, outer part of aperture missing. ,,. Umbilicus' 

obscure ventral swelling, and less elevated keel; narrow, steep-walled, in width about one-half 

from Cardioceras cordiforme (Meek and Hayden) the height of the whorl on inner turns. Urnby its wider umbilicus, coarser sculpture, and by bilical shoulder fairly sharp and overhanging~ 

its wide superior. saddle and narrow first lateral Sculpture regular. At the diameter of 120 · 

lobe. It differs from Oardioceras popilianense millimeters broad, rounded U:mbilical ribs (l6 : 

Boden 1 in its coarser and more regular ribs and to the whorl) split at about one-fourth the di~ V 

wider umbilicus; from Oardioceras rouilleri tance toward the ven.ter into three or fdtit l<?#~· · Nikitin 2 in its wider umbilicus, more numerous rounded secondaries. Intercalated ribs Tise:-b~~ 

primary and coarser secondary ribs, and less tween · these bundles. The secondaries·[ bknd • 

pinched venter; from Oardioceras nikitinianum forward on the venter to form ari. obtuse ~inu~( 

Lahusen 3 in its wider umbilicus and more There would be 60 siphonal ribs on a comp'lete 

numerQus ribs;- from Cardioceras cordatum whorl at the diameter of 120 to 150 ·millimeters. ·' 

(Sowerby) Lahusen 4 in the lower cross section The older part of the living chamber shows the 

of the whorl, more obtuse and nonnodose ribs; siphonal ribs very faintly; the remainder of the ·· · 

from Cardioceras cordatum (So~erby) Ilo- living chamber has only broad,· lowJ obsctir~ .. 

waisky 5 in wider umbilicus and coarser. ribs; swellings on the flanks. 

1 Boden, Karl, Die Fauna des unteren Oxford von Popilany in Litau: 

Geol. u . palaeont. Abh., N. , Band 10, Heft-2,p.41, pl. 2, figs.4, 5, 1911. 

2Nikitin, S. N., Die Jura-Ablagerungen zwischen· Rybinsk, Mologa, 

und Mischkin an der oberen Wolga: Acad. sci. St.-Petersbourg Mem., 

7th ser., vol. 28, No. 5, p. 56, pl. ·2, fig. 17, 1881. 

a Lahu.Sen, Joseph, Die Fauna der jurassischen Bildungen des Rjasanschen Gouvemements: Com. geol. [Petrograd] Mem., vol.1, No.1, p. 5G, 

pl. 5, figs. 7-9, 1883. 

~Idem, p. 49, pl. 5, fig. 3. 

6 Ilowaisky, David, L'Oxfordien et le Sequanien des gouvernements 

de Moscou et de Riasan: Soc. imp. nat. Moscou Bull., new ser., vol. 17, 

p. 266, pl. 10, figs. 28-30, 1903. · 

The last suture has a moderately wide, sh'drt 

siphoriallobe;l first lateral s·addle of about same 

.width;, first lateral lobe relatively narroW' and 

.longer than tln.e ·siphonallobe; other eleinents 

~mall. Suture not deeply incised. Radial line 

'cuts both thelfirst· and. second 1atera116'bes. 

s Borissjak, A., Die Fauna des Donez-Jura: Com. geol. [Petrograd] 

Mem. , new ser. , livraison 37, p. 8, pl. 1, fig. 8, 1908. 

7 Idem, p. 8, pl. 1, fig. 7. 
36 SOME AMERICAN JURASSIC AMMONITES. 

This specimen is very close to Oardioceras? 

incertumReeside in the narrow venter, in suture, 

and in the type of sculpture. It differs in its 

proportionately wider whorls, larger size, and 

more persistent and finer sculpture. The inner 

whorls are not preserved, and it is not possible 

to say whether they were rounded or had a ventral angle. Despite the uncertainty attached 

to this specimen it should be recognized in 

futlire collections and it deserves a distinct 

name. The writer knows .no closely. related 

foreign species. 

Locality: Type, Sundance formation, in T. 25 

N.-;;R . . 78 W., Freezeout Hills, Carbon County, 

Wyo. 

~ardioceras? incertum Reeside, n. sp. 

Plate XX, figures 17-20; Plate XXI; Plate XXII, 

· · ·figures'!, 2'.' · · 

imperceptible. The secondaries are plainly 

visible on the early part of the living chamber 

but grow fainter and finally disappear. There 

are 17 umbilical ribs and 50 secondaries on the 

last septate whorl of the type. 

The suture shows a short; wide siphonallobe;. 

first lateral saddle equal in width to thesiphonal 

lobe; long, narrow first lateral lobe; second 

lateral saddle equal in width to first laterallobe; 

small second lateral lobe; and auxiliaries. The 

radius cuts the first lateral lobe. Suture deeply 

incised. 

Oardioceras? incertum is marked by the 

rounded venter of the older whorls and the 

narrowed venter of the younger ones, the low 

sculpture, and the suture. The generic reference of this· species is open to question. The 

sculpture and suture might occur with Oardioceras or Q.ruenstedticeras. Some . species of 

Shell stout, venter round in youth but narrow Quenstedticeras have sharp-ventered young, but 

· and almost sharp in adult. Whorls unknown · the adults are all round-ventered. Oardioceras 

where the shell is less than 2S millimeters in acquires a keel or a sharpened venter at an 

diameter; from a diameter of 25 millimeters early stage and keeps it throughout life. As 

t~ a diameter of 55 mi~limeters cross section of the adult of this form is as sharp-ventered as 

whorl broadly rounded and height about three- some undoubted species of Oardioceras, it is 

fourths the width; where the shell is more than tentatively referred to that genus, though cer55 millimeters in diameter whorl narrowed tainly it is not ~ typical Oardioceras. The 

ventrally and cross section cordiform, subequal adult suggests Oardioceras bellefourchense Reein width and height. The type is 130 milli- side, though it is not so stout nor so strongly 

meters in diameter and preserves nearly half sculptured, and Gardioceras crassum Reeside, 

the last whorl unseptate, scaphitoid; aperture but it is less strongly sculptured. The younger 

whorls differ from Quenstedticeras? subtumidum unknown. Umbilicus moderately wide, in 

width about three-fourths the height of the (Whitfield and Hovey) in form and suture. 

outer whorl in the plane of the last septum. Besides the type specimen there are five other 

Umbilical shoulders rounded and inner slope large specimens from the type locality which 

short and steep until the shell attains a diam- preserve the outer whorls only. These specimens all show a rounded venter in the older eter of 90 millimeters. · part, which narrows markedly in the younger Sculpture of moderate strength. Rather part. All show a persistence of the ventral 

low, rounded umbilical ribs pass one-third the ribs on the living chamber and essentially the 

distance from the umbilicus to the venter, split same suture. There is some variation in the 

into two o:r three low, rounded secondaries, width of the whorls, and the stoutest specimen 

which pass almost to tlie median line of the is shown in figure 1, Plate XXI. It has seemed 

venter and then bend forward to form shallow inadvisable to ascribe these specimens to differchevrons. The ventral sculpture was not seen ent species until material ·with the younger 

where the diameter is less than 80 millimeters whorls preserved is available. 

and chevrons may be lacking. Some inter- Locality: Type specimen from _Sundance forcalated secondaries are present. Toward the mation, T. 25 N., R. 78 W., Freezeout Hills, 

end of the septate part of the type the umbilicai Carbon County, Wyo. Also the outer whorls 

ribs become faint and on the living chamber are ?f five large, poorly preserved specimens. 
CARDIOCERAS. ·37 

Cardioceras? Iatum Reeside, n. sp. 

flate XX, figures 7-16. 

Shell stout, whorls depressed; vertter 

rounded except in larg_e individuals. Cross 

section of whorls where the diameter is less 

than 15 mi~limeters , not seen in type, but inc 

other specimens is practically circular; 'Yhere 

the diameter is more than 15 millimeters cross 

section is oval, much wider than high. In a 

part of a whorl (Pl. XX, figs. 13-16) belo~ging 

to a shell that had a diameter of 80 to 90 millimeters the venter is broadly angular, ' but 

otherwise the form is the same as that of the 

younger whorls. . Last half whorl of type is 

unseptate, as is a~so large fragment. Aperture 

unknown. Umbilicw;; wide, in width 1! times 

height of outer whorl. Umbilical shoulders 

gently rounded through all stages seen. \ 

Sculpture regUlar. Whorls smooth to diameter of 8 or 9 millimeters, when distinct u·mbilical ribs begin. On the type these are at 

first rounded, then sharp; they rise within the 

umbilicus, pass outward with a shallow, for 

ward curve to the middle of the flank, and there 

reach the maximum height. Each primary 

then forks into two, rarely three, low, round'ed 

secondaries, which pass nearly to the middle 

line of the venter and bend forward to make a 

shallow, rounded chevron. On some specimens 

the ribs are faint at the median line of ·the 

venter, but there is never an elevated keel. 

• The large fragment mentioned above has a 

mold on the inner side that shows the characteristic round ' venter and shallow, rounded 

chevrons of the type, but on the ~xterior it has 

only obscure traces of the secondaries on the 

older end and none on the younger end. The, 

primaries, however, are well marked. 

The suture of the type has a broad siphonal 

lobe; equally broad first lateral saddle; relatively narrow first lateral lobe of same length as 

siphonallobe; short, wide second lateral saddle; small second lateral lobe; and auxiliaries. 

The radius just touches the first lateral lobe. 

This species is characterized by its depressed 

whorls that have a flattened, unkeeled venter; 

rounded, shallow ventral chevrons; and the 

acquisition of an obtusely angled venter at 

maturity. The writer is in doubt as to its 

generic reference. The form, sculpture, and 

suture place it with the Cardioceratidae beyond 

question. The suture differs from that of 

Stepheoceras, Pachyceras, and . Ohariwussetia, 

and the sculpture differs from that of .Maerocephalites and Oadoceras. None of these '•genera, moreover, are known from the horizon of 

true Oardioceras with · which this species is 

associated. It would seem to belong to either 

Quenstedticeras or Oardioceras. The main distinction between these genera is that -the adult 

of Quenstedticeras is round-ventered, whereas 

the admt of Oardioceras is sharp-ventered. · 

As this species has at first a round venter and 

later an angular venter, it is referred to Oardioceras, with the reservation that it is certainly not a typical representative of the genus. 

The writer knows no species close enough to 

deserve comparison. 

All specimens from Sundal).ce formation. 

Localities: Type, half a ·mile northeast of the 

Johnson horse ranch, Belle Fourche River, 25 

miles west of Sundance, Crook County, Wyo.; 

also another crushed specimen: SW.! sec. 13, 

T. 23 N., R. 79 W., Carbon County, Wyo.-; one 

small and part o£ la large specimen. Locality 4 

miles west of Devils Tower, Crook County, 

Wyo.; two specimens . . Locality 1! miles north 

of Redwater Creek, near the boundary between South Dakota and Wyoming, in Wyoming; 1 small specimen. Como Bluff, near 

Aurora (Ridge), Wyo.; 1 sm·all specimen. 

Cardioceras? sp. undet. 

Plate XXIV, figures 3, 4. 

Single specimen, poorly preserved. Shell 

compressed, venter pinched where the diameter 

is more than 30 millimeters. Cross section 

higher than ~ide. Living chamber partly preserved (three-eighths of last whorl) but 

crushed; ~ scaphitoid. Umbilicus narrow, in 

width one-fourth height of outer whorl. Umbilical shoulders sharp and steep, overhanging 

inner slope on last whorl. 

Whorls entirely smooth, so far a...;; can be observed; where the diameter is more than 50 or 

60 millimeters. Sculpture unknown where the 

diameter is less than 50 millimeters. 

Suture crowded, little incised. Siphonal 

lobe, short, narrow; first and second lateral 

saddles wide, subequal; first lateral lobe about 

as wide as siphonal lobe and a little longer; 

second -lateral lobe relatively narrow, short; 

other elements small. 

The specimen described above is too poor for 

definite reference, but ~he general form is that 
SOME AMERICAN JURASSIC AMMONITES. 

of Gardioceras. It might be a larger specimen 

of fotn;is like, Gardiorerashyatti Reeside ann Oardi ocera~·, alli~~clliium Reeside, though the proportions of the suture m'ake this doubtful. The 

co~pressed form, pinched venter, and the 

nearly equal size of the first and second lateral 

saddles a:re ·yery suggestive of Ohamoussetia R. 

Douville 1 and it would be within reason to 

ref~r . the' ·'specimen to that genus, though the 

A.merican specimen is associated with Car .. 

dioceras. 

Locality: One specimen from Sundance 

. formation, T. 2.5 N., R. 78 W., Freezeout 

Hills, Carbon County, Wyo. 

Genus AMOEBOCERAS Hyatt. 

Amoeboceras dubium (Hyatt) Reeside. 

Plate XXIV, figures 5-8. 

1894. Cardioceras dubium Hyatt, Geol. Soc. America Bull., 

vol: 5, p. 420. · 

1894. ?Cardioceras whitneyi Smith, Geol. Soc. America 

Bull., vol. 5, p. 253. 

Hyatt's description is as follows: 

The young are-imperfect, but it can be readily seen that 

the earliest stages were smooth for a long period, as in some 

speciesof the genus Cardioceras, and that they differ entirely from the heavily tuberculated young of the genus 

Amaltheus. 

1 Douville, Robert, Soc. geol. France, Paleontologie, vol. 19, fasc. 2, 

Mem. 45, pp. 19-23, 1912 

The costae· arise abruptly and do not appear to differ 

materially, so far as seen on the sides, from those of later 

stages. This is also more consistent with the characteristi~ of Cardioceras than with those of Amaltheus. 

The costae on the sides and genicular ridges are mimerous, linear and single. There are no lateral tubercules, but 

occasionally the costae disappear on the sides just before 

reaching the geniculae. The geniculae are very prominent and tuberculated. · 

The whorls are compressed, with sides flattened. The 

venter is very narrow. The inclusion varies from about 

one-third the diameter of the inner to considerably less, 

and there may be two species, one including more involute 

and the other the less involute forms, but both have what · 

would be called open umbilici. The keel is prominent 

and deeply crenulated. It is separated from the geniculae 

by narrow channels, which are not crossed by bifurcations 

of the costae, as in more generalized species of. this genus. 

The serrations of the keel appear to be separated by the 

channels from the costae. · 

The sutures were present, , and, although much distorted, it was ascertained that they had large first lateral 

saddles and lobes, with smaller second lateral saddles and 

small second lateral lobes. There are auxiliary saddles on 

the umbilical shoulder, and the abdominal lobe is shorter 

than the first lateral lobes. These sutures are decidedly 

cardioceran. 

This species is marked by the straight, simple ribs and the smooth channels on the 

venter and is apparently a true Amoeboceras. 

Oardioceras whitneyi ·smith from the same locality may be the same species. 

Locality: Type, ·Mariposa slate, Texas 

ranch, Calaveras Co~ty,, Calif. 
. . . 

PLATES. 

39 
PLATE I. 

Page. 

FiGURES 1-4. Quenstedticeras collieri Reeside, n. sp., from Ellis formation in sec. 7, T. 25 N., R. 26 E., Little 

Rocky Mountains, Mont....... . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 14 

1-3. Side, front, and back views of type (U.S. Nat. Mus. catalogue No. 32298). 

4. Side view of another specimen (U. S. Nat. Mus. catalogue No. 32299). 

FiGURES 5-8. Quenstedticeras collieri Reeside, n. sp., from Ellis formation in sec. 27, T. 25 N., R. 26 E., Little 

Rocky Mountains, Mont ........ . ........ .............................................. 14 

5, 6. Side and back views of another specimen (U.S. Nat. Mus. catalogue No. 32300). 

7, 8. Side view and suture (X 2) of another specimen (U. S. Nat. Mus. catalogue No. 32301). 

FIGURES 9-12. Quenstedticeras? hoveyi Reeside, n. sp., from Sundance formation in foothills west of Bear Lodge 

Mountains, Wyo. Side and back views, cross section, and suture of type (U. S. Nat. Mus. 

catalogue No. 29328)...................................................... . .......... . . 15 

' FIGURES 13; 14. Quenstedticeras? hoveyi Reeside, n. sp., from Sundance formation in Como Bluff, near Aurora, 

Wyo. Section and suture of the large specimen (U. S. Nat. Mus. catalogue No. 32302) 

shown in Plate :II...................................................................... 15 

40 
U S. GEOLOGICAL SURVEY PROFESSIO)<AL pAPER liS PLATE I 

I. ' 4 3 7 

2 

~X2 8 

11 

10 

12 

13 

14 9 

JURASSIC AMMONITES. 
U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 118 PLATE II 

2 

JURASSIC AMMONITES. 
PLATE II. 

FIGURE 1. Quenstedticeras'! hove~Ji Reeside, n. sp., from Sundance formation in Como Bluff, near Aurora, Wyo. 

Side view of a large specimen (U.S, Nat. Mus. catalogue No. 32302)....................... 15 

FIGURES 2-4. Quenstedticeras? suspectum Reeside, n. sp., from Sundance formation in Como Bluff, near Aurora, 

Page. · 

Wyo ......................... .. _ . . . . . _ . .. _._ ..........................•... _._._ ... _._._. 16 

2, 3. Side and front views of a small specimen (U.S. Nat. Mus. catalogue No. 32303). 

4. Suture of type (U. S. Nat. Mus. catalogue No. 32304). · , 

41 
PLATE III . . 

FIGURES 1. 2. Quenstedticeras? suspectum Reeside, n. sp., from Sundance formation in locality half a mile northPage. 

east of the Johnson horse ranch, Belle Fourche River, 25 miles west of Sundance, Wyo. 

Side and back views (X ~)of a large individual (U.S. Nat. Mus. catalogue No. 32359)...... 16 

42 

-
"C". S. GEOLOGICAL SL'"RVEY PROFESSIONAL PAPER 118 PLATE III 

.2 

JURASSIC AMMONITES. 
U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 118 PLATE IV 

2 

10 7 

JURASSIC AMMONITES. 
PLATE IV. 

FIGURES 1-4. Qu,enstedticeras? subtumidum (Whitfield and Hovey) Reeside, from undance formation on Inyankara Creek, Wyo. Side and back views; cross section, and suture of type specimen ... -.. 

FIGURES 5-7. Quenstedticeras? suspectum Reeside, n. sp., from Sundance formati Ii in Como Bluff, near Aurora, 

Wyo ............................ . ................ -.-.: -•".-.--. -- ----------------------- · • 

7. Side view of type with part of outer whorl removed. (U·. s.' ~t. Mus. catalogue No. 32304). 

5, 6. Side and back views of another specimen (U. S. Nat. Mus. catalogue No~ 32305). 

FIGURES 8-10. Quenstedticeras? tumidum Reeside, n. sp., from Sundance format'on in Red Canyon, half a mile 

south of Matias Peak, in T. 3 S., R. 7 E., Fall River County, S. Dak ..... -.--- ... -.. --.-.-

8, 9. Side and back views of an inner whorl of the type . . (See l. V.) 

10. Suture of type specll;nen. 

43 

Page. 

16 

J:'6 

17 

• 

. ' 
PLATE V. 

FIGURES 1, 2. Quenstedticeras? suspectum Reeside, from Sundance formation in Como Bluff, near Aurora, Wyo. 

Side view and cross section of type specimen (U. S. Nat. Mus. catalogue No. 32304).-------

FIGURES 3, 4. Quenstedticeras? tumidum Reeside, n. sp., from Sundance formation in Red Canyon, half a mile 

south of Matias Peak, in T. 3 S., R. 7 E., Fall River County, S. Dak. Side view and cross 

section of type. _________________ .......... _ ............ _ ............... _ .............. .' 

44 

Page. 

16 

17 
U.S. GEOLOGICAL SURVEY 

3 

JURASSIC AMMONITES. 

PROFESSIONAL PAPER 118 PLATE V 

I 

I 

I 

\ // 

,~J· 

',, ,,' .......... ,' .................. _____ ,,, 

2 
U.S. GEOLOGICAL SURVEY 

• 

8 16 15 

JURASSIC AMMONITES. 

PROFESSIONAL PAPER 118 PLATE VI 

17 

4 

~-6 

~ 14 

h I §:1A ~ MAA'-- r 

__ .<qV_-__ 5tf -~- ~r~r

l 18 
PLATE VI. 

FIGURES 1-6. Cardioceras whitfieldi Reeside, n. sp., from Sundance formation in locality 2 miles south of Belle 

Page. 

'Fourche River, near Bear Lodge Butte, S. Dak...... . . . . . . . . . . . . . . . . . . . . . . • . . . . . . . . . . . . . . 34 

1-4. Sideandfrontviews, crossse_ction, andsutureoftype (U.S. Nat. Mus. catalogueNo.l2294.) 

5, 6. Side view and suture of another specimen (U.S. Nat. Mus. catalogue No. 12345). 

FIGURES 7-10. Cardioceras ~laskense Reeside, from lower part of Naknek formation on east side of Oil Bay, Cook 

Inlet region, Alaska. Side and front views, cross section, and suture of type (U. S. Nat. 

Mus. catalogue No. 32306) ............... __ . _. _ ... _ ......... _ ......... • .. _ .. _ ........ -.-.- · 18 

FIGURES 11-14. cdrdioceras stillwelli Reeside, n. sp., from Sundance formation in locality 4 miles west of Sundance, Wyo. Side and back views, cross section, and suture of type (U.S. Nat. Mus. catalogue No. 32307) ....... · ...........•...•.............. _. _. _ ... _. _ ... _. _ ... _. _. _......... 31 

FIGURES 15-20. Cardioceras americanum Reeside, from Sundance formation in locality 4 miles west of Sundance, 

Wyo.-................... : .· ................................. _ ............ _ .. _._......... 18 

15-18. Side and back views, cross section, andsutureoftype(U. S. Nat. Mus. catalogueNo.32308). 

19. Side view of another specimen (U. S. Nat. Mus. catalogue No. 32309). 

20-. Crosssectionofinnerwhorlsofanotherspecimen(X·6) (U.S. Nat. Mus. catalogue No. 32310). 

45 
PLATE VII. 

FIGURES 1-6. 

Page. 

Cardioceras cordiforme (Meek and Hayden) N eumayr .............. ·. . . . . . . . . . . . . . . . . . . . . . . . . . . . . 21 

1-4. Side view with outer whorl, side view >yithout outer whorl, cross section, and suture of 

type (U. S. Nat. Mus. catalogue No. 203), from Sundance formation in locality at "southwest 

base of Black Hills," Wyo. ·· 

5, 6. Side view and cross section of small individual (U. S. _N_at. ¥us. catalogue No. 32311), 

from locality 4 miles west of Devils Tower, Wyo. · 

46 

, . 
U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 118 PLATE VII 

JURASSIC AMMONITES. 
U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 118 PLATE VIII 

4 5 

2 

JURASSIC AMMONITES. · 
• 

PLATE VIII. 

Page. 

FIGURES 1-7. Cardioceras cordiforme(Meekan<l Hayden) Neuml yr ... .' ............. ."......................... 21 

1-3. Side and front views (X ~) and suture (X 1) of a specimen (U. S. Nat. Mus. catalogue 

No. 32312) differingsomewhatinformfrom th~type, from Freezeout Hills,AlbanyCounty,Wyo. 

4, 5. Side and back views of small individual '(U. S. Nat. Mus. catalogue No. 32313), from 

loca~ty 1! miles north of Redwater, Wyo. 

6.' Side view of small individual (U. S. Nat. Mus. catalogue No. 32314), from locality 1! miles 

nort of Redwater, Wyo. . . . 

7. Side view of small individual (U.S. Nat. Mus. catalogue No. 02315), from locality 4 miles west 

of Devils Tower, Wyo. 

All specimens from Sundance formation. 

47 

/ 
PLATE IX. 

FlGURE- 1. Cardioceras cordiforme (Meek and Hayden) Neumayr, from Sundance formation in Freezeout Hills, 

Albany County, Wyo. Cross section of the large individual shown in Plate VIII (U. S. 

Nat. Mus. catalogue No. 32312) ........................................................ . 

FIGURES 2-4. Cardioceras crookense Reeside, n. sp., from Sundance formation, in locality 4 niiles west of Devils 

Tower, Wyo. Side view, cross section, and suture of type (U.S. Nat. Mus. catalogue No. 32316). 

FIGURES 5-8. Cardioceras martini Reeaide, n. sp., from Cardioceras-bearing beds on creek entering Boulder Creek 

from north, Matanuska Valley, Alaska. Side and front views, cross section, and suture (X 2) 

of type (U. S. Nat. Mus. catalogue No. 32317) .......................................... . 

FIGURES 9-12. Cardioceras plattense Reeside, n. sp., from Sundanc.e formation in Grand Canyon of the North 

Phtte, near Alcova, Wyo. Side and back views, cross section, and suture of type (U.S. Nat. 

M \ls. catalogue No. 32318) ............... ~ .......•...••••.•••.••..........•••.•....•...•• 

48 

Page. 

21 

23 

. '27 

28 
U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 118 PLATE IX 

C) 

7 
U. S. GEOLOGICAL SURVEY 

JURASSIC AMMONITES. 

PROFESSIONAL PAPER 118 PLATE X 

r I 

I 

' ' I 

' ' I 

I 

I 

\ 

' ' ' ' ' ' ' ' 

5 

\ 

\ 

\ 

\ 

\ 

\ 

\ . _______ _ 

,/ 

....... "' 

I 

I 

I 

I 

I 

' I 

I 

I 

I ' 

I 

! 

j 

I 

I 

I 
· PLATE X. 

FIGUR'B 1-5. Cctrdim>!ras auror(lenS< Reeside, fr~m sunL nce formation in Como Bl!uff, near Aurora, Wyo .. 

1-3.· Side and back views and suture of type (U.S, Nat. Mus. catalogue No. 32319). 

4, 5. Side view and cross section (restofed) of cast of mold of par1; of two whorls (U. S. 

Mus. catalogue No. 32320). 

114216°--19----4 . 49 

J 

Page. 

19 
PLATE XI. PaJe. 

FIGURES 1, 2. Cardioceras bellefourchense Reeside, from Sundance formation on Belle Fourche River, 25 miles 

west of Sundance, Wyo. Side view and suture Gf type (U. S. Nat. Mus. catalogue No. 32321) _ _ 20 · 

FIGURES 3-5. Cardioceras schucherti Reeside, n. sp., from Sundance formation in Freezeout Hills, Albany 

County, Wyo. Side and end views and suture of type (U.S. Nat. Mus." catalogue No. 32322).. 30 

50 
I 

U. S. GEOLOGICAL R:JRVEY PROFESSIONAL PAPER 118 PLATE XI 

4 
U. S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 118 PLATE XII 

JURASSIC AMMONITES. 
PLATE II. 

FIGURES 1, 2. Cardioceras bellefourchense Reeside, n. sp., from Sundance formation on Belle Fourche River, 

25 miles west of Sundance, Wyo. Front view and suture of. type (U. S. Nat. Mus. catalogue 

Page. 

No. 32321). ________ . _ ... _ . _ . ______ ... ________ . ______ ·" __________ ........•.•. __ , _______ . _ . . . 20 

FIGURES 3, 4. Cardioceras crassumReeside, n. sp., from Sundance formation ,in Freezeout Hills, Albany County, 

Wyo. Side view and suture of type (U.S. Nat. Mus. catalo~e No. 32323) ... -- ~ .... -.r ...... 23 

. ' 51 
PLATE. XIII. 

FIGURE 1. Ca:rdioceras crassum Reeside, n. sp., from Sundance formation in Freez\~out Hills, Albany County,Wyo. 

Side view of type specimen (U. S. Nat. Mus. catalogue No. 32323) _ . -- ___ .•• ____ -- _____ ..... _- _-

Fli:GURES 2, 3. CardiOceras russelli Reeside, n. sp., from Sundance formaticn in l11cality 1~ n::.iles north of Redwater, Wyo. Side view and cross section of type (U. S. Nat. Mm·. catalogue No. 32324) .••.... 

52 

Page. 

23 

29 
U.S. GEOLOGICAL SURVEY PROFESSIONAL rAPER 118 PI,AT:& XIII 

3 

JURASSIC AMMONITES. 
U. S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 118 PLATE XIV 

+ 

4 

JURASSIC AMMONITES. 
·PLATE XIV. 

FIGURES 1, 2. Cardioceras crassum Reeside, n. sn. , from Sundance formation in Freezeout Hills, Albany County; 

. . Wyo. Back view and cross +ction of type (U. S. Nat. ·Mus. catalogUe No. 32323):. _ _ _ _ _ _ _ _ 23 

FIGURES 3, 4. Cardioceras russelli Reeside, n. s:n., from Sundance formation in locality H miles north of Red-

... -. . water, Wyo. Front view and suture of type (U. S. Nat. Mus. catalogue No. 32324). _______ ~ 29 

FIGURE 5. Cardiocer~ ru~selli Reeside, n. s~., lfrom Sundance formation in somo Bluff, near Aurora, Wyo. 

S1de VIew of another specrmen (U. S. Nat. Mus. catalogue No. 32325). _. _ . _ ~ _ . _. ____ . _ _ _ _ _ 29 

Page. 

53 
I 

PLATE XV. 

FIGURES 1-4. Cardioceras hyatti Reeside, n. sp., from Difficulty Canyon, sec. 31, T. 24 N., R. 80 W., Carbon 

County, Wyo. Side and back views, cross section, and suture.( X 2) of type (U.S. Nat. Mus. 

catalogue No. 32326) .......................•............... ... ...................••.... 

· FIGURES 5-8. Cardioet5ras stantoni Reeside, n. sp., from Difficulty Canyon, sec. 31, T. 24 N., R. 80 W., Carbon 

County, Wyo. Side and back :views, cross sect~on, and suture (X 2) of type (U. S. Nat. 

Mus. catalogue 32327) ... . .................. : ....................................... : ... . 

FIGURES 9-11. Cardiocuas stantoni var. obesum Reeside, n. var., from DifficlJ.lty Canyon, sec. 31, T. 24 N., 

R. 80 W., Carbon County, Wyo. Side and back views and cross section of another speci-

. _ . _. men Fith stouter whorla (U. S. Nat. Mus. catalogue No. 32328) .......................... . 

]'IGURES 12, 13. Cardfoceras wyomingen_se Reeside, n. sp., from Como Bhdf, near Aurora, Wyo. Side view and 

_. . . cross section of type (U. S. Nat. Mus. catalogue No. 32329) .... · ..... ~ .................... . 

FIGURES 14-16. Cardioceras wyomingense Reeside, n. sp., from Freezeput Hills, Albany County, Wyo. Side 

view, cross section, and suture of another specimen (U. S. Nat. Mus. catalogue No. 32330). 

FIGURE 17. Cardioceras wyomingense Reeside, n. sp., from locality 4 miles west of Sundance, Wyo. Side 

view of another specimen (U. S. Nat. Mus. catalogue No. 32331) . .. ....................... . 

FIGURES 18-21. Cardioceras distans (Whitfield) Neumayr, from locality 2 miles south of Belle Fourche River, 

near Bear Lodge Butte, S. Dak Side and back views, cross section, and suture· (X 2) of 

type (U.S. Nat. Mus. catalogue No.12308) ...............•.............................. 

FIGURES 22-24. Cardioceras distans (Whitfield) var. depressum Reeside, n. var., from locality half a mile northeast of the Johnson horse ranch, Belle Fourche River, 25 miles west of Sundance, Wyo. 

Side and back views and cross section of type (U.S. Nat. Mus. catalogue No. 32332) ...... . 

All specimens from Sundance formation. 

54 

Page. 

26 ' 

31 

31 

34 

34 

34 

24 

25 
. SURVEY u.s. GEOLOGICAL 

0 

3 

21 

24 

18 19 23 22 

JURASSIC A MMONITES. 

PROFESSIO~AL P. o\PER 118 PLATE. XV 

~ 

G 

0 

20 

9 

4 

8 

15 
U.S. GEOLOGICAL SURVEY 

4 5 

8 

JURASSIC AMMONITES. 

PROFESSIONAL PAPER 118 PLATt: XVI 

6 

10 

11 
PLATE XVI. 

Page. 

FrG URE s 1-4. Cardioceras dis tans· (Whitfield) N euma. yr, from Sundanc,e fmrmat.ion, in locality half a mile- north- _ 

east of the Johnson. horse ranch, Belle Fourche RiverJ 25 mil es west of Sundance, Wyo.... "24 

. 1-~23~~ and bac.k ~e~s and s~ture of an~ther . sp~~im~~ (U .. s_. N.at. Mus. catalogue No. 

4. Side view of another specimen (U. S. Nat. Mus. ca alogue No. 32334). · 

FIGURES 5-6. Card;~;~~~~~a~~~~~~:!~~~:~:~~-r: -f~~~~ ·l·o-~~~ ~-~r~-~f ~~~ ~ -f~~~~~i-0~, -~~ -~~-t-s~~~-~f- ~-i~ . 24 

5. Side view of a small specimen (U. S. Nat. Mus. catlogue No. 323~5). · · 

6. Side view of a larger fragment (U. S. Nat. Mus. cat logue No. 32336). 

FIGURES 7-8. Cardioceras distans var. depressum Reeside, n . var., from S ndance formation in locality 4 miles 

;:;~e o~~-u~~~~~~:. ~:~: _ -~~~~-~~~ ~c-~ -~i·e-~~ _o_f_ ~-l r~~~-~~~~~~~~ _C~-- ~-· -~-~~: -~~~-- _c_a_t~~ Fraur..Es 9-10. Cardioceras distans var. depre.~sum· Reeside,. n. var., fro~ lower part ot Naknek formation on 

east side of Oil Bay, Alaska. Side and back views of a small specimen (G. S. Nat. Mus. 

FIGuRE 11. Ca,dio::l"£'~! ~~~~ 2~~;;,:,;,;; ·R~~;rid~: ;; .. ~~~:: j;_,;,;; . 4;;.j~~~~. i~~ ti~~. ;;; j~~~jit; b;;li. ~ . mile northeast of the Johnson horse ran-ch, Belle Four he River, 25 miles west of Sundance, 

Wyo. Side view of another specimen (U. S. Nat. Mus .. catalogue No. 32339) ............... . 

55 

• 

25 

25 

25 
PLATE XVII. 

FIGURES 1-4. Cardioceras whiteavesi Reeside, n. sp., from Sundance formation in locality west of Sundance, 

Wyo. Side and back vie~s, cross section, and suture (X 2) of type (U. S. Nat. Mus. cataPage. 

logue No. 32340). _ .. _____ . ____________________________ . _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ 33 

FIGURES 5-11. Cardioceras canadense Whiteaves from Sundance formation in locality 4 miles west of Sundance, Wyo.- _____________________ . _________ .- _________________________________________ .- _____ . 20 

5-8. Side and back views, cross section, and suture of small individual (U. S. Nat. Mus. catalogue No. 32341). 

9,10. Side and back views of another specimen (U. S. Nat. Mus. catalogue No. 32342). 

11. Side view of a small spe.cimen (U. S. Nat. Mus. catalogue No. 32343). 

FIGURES 12-19. Cardioceras sundancense Reeside, n. sp., from Sundance formation in locality 4, miles west of Sundance~ Wyo _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ 32 

12. Side vi~w of a small specimen (U. S. Nat. Mus. catalogue No. 32344). 

13-15. Cross section (X 1), suture (X 2), and cross section (X 6) of the inner whorls of a small 

specimen (U.S. Nat. Mus. catalogue No. 32345) . . 

16. Side view of a medium-sized individual (U.S. Nat. Mus. catalogue No. 32346). 

17-19. Cross section and two sutures of type (U. S. Nat. Mus. catalogue No. 29314). (See 

Pl. XVIII.) I 

FIGURES 20-23. Cardioceras lillooetense Reeside, n. sp., from Cardioceras-bearing beds at head of Big Creek, 

tributary to Chilcotin River; Lillooet, British Columbia. Side and. front views, cross section, and·suture of type (U. S. Nat. Mus. catalogue No. 32347) •••••••••••••••••••• -....... 27 

56 
U.S. GEOLOGICAL SURVEY 

12 

17 

PROFESSIONAL PAPER 118 PLATE XVII 

7 

3 

11 10 

~; 

+--~~-- 18 I 

JURASSIC .AMMONITES. 

-~-

-~+~~~Do~- 8 

16 

19 
U.S. GEOLQGICAL SURVEY 

2 

JURASSIC AMMONITES. 

PROFESSIONAL PAPER 118 PLATE XVIII 
• 

PLATE ~XVIII. 

Page. 

FIGURES 1-3. Card~::~ ;;:~~~~-~~~s~~~~ -~·- ~~~~- ~0-~- ~~~~~~~-f_o_r~ ~~~~ -i~. ~~~~~~~-~ -~~s- ~e~:-~~ ~~- 32 

1, 2. Front and side views of type specimen (U.S. Nat. Mu~l catalogue No. 29314). 

3. Same, with part of outer whorl removed. ~ ·· 

FIGURE 4. Cardioceras spinijerum Reeside, n. sp., from lower part of Nakn k formation, on Oil Bay, Cook Inlet 

region, 4J.aska. Cross section of type speeimen (U. S. . at. Mus. catalogue No. 32348). . . . . 3e 

57 

• 
• 

PLATE XIX. 

Page. 

FIGURES 1-3. Cardioceras spiniferum Reeside, n. sp., from lower part of Naknek formation, on east side of Oil 

Bay, Cook Inlet region, Alaska ................................................. _ . . . . . . . . 30 

1, 2. Side and end views, cross section of type (U. S.· Nat. Mus. catalogue No. 32348). 

3. Side view of same with part of outer whorl removed. 

FIGURES 4-8. Cardioceras haresi Reeside, n. sp., from Sundance formation in: locality 1! miles north of Redwater, Wyo. Side, front, and back views, cross section, and suture of type (U. S. Nat. 

Mus. catalogue No. 32349) ............... _. _ ... _ ... _ ~ .......... -... -................... 25 

FIGURES 9-12. Cardioceras haresi Reeside, n. sp., from Sundance formation in SW.! sec.13, T. 23 N., R. 79 W., 

Carbon County, Wyo. Side, front, and back views, cross section of another specimen (U.S. 

Nat. Mus. catalogue No. 32350).......................... ... . . . . . . . . . . . . . . . . . . . . . .. . . . . . • 25 

58 
U. S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 118 PLATE XIX 

~~ 8 X .2. 

JURASSIC AMMONITES. 
U.S. GEOLOGICAL SURVEY 

8 

9 

6 

6 

PROFESSIONAL PAPER 118 PLATE XX 

2 ~ 4 

JURASSIC AMMONITES. 
PLATE XX. 

FIGURES 1-4. Cardioceras obtusum Reeside, 'n. sp., from locality 1~ miles north of Redwater, Wyo. Side and 

Page. 

back views, cross section, and suture of type (U.S. Nat. Mus. catalogue No. 32351)_ _ _ _ _ _ _ 28 

FIGURES 5, 6. Cardioceras obtusum Reeside, n . sp~, from locality 4 miles west of Devils Tower, Wyo. Side view 

and cross section of another specimen (U. S. Nat. Mus. catalogue No. 32352) _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ 28 

FIGURES 7, 8. Cardioceras? latum Reeside, n. sp., from locality 4 miles west of Devils Tower, Wyo. Side and 

back views of another specimen (U.S. Nat. Mus. catalogue No. 32353) _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ 37 

FIGURES 9-12. Cardioceras? latum Reeside, n. sp., from locality half a mile northeast of the Johnson horse ranch, 

Belle Fourche River, .2::> miles west of Sundance, Wyo. Side and front views, cross section, 

and suture (X 2) of type (U.S. Nat. Mus. catalogue No. 32354)----------- _________ - _ _ _ _ _ _ _ 37 

FIGURES 13-16. Cardioceras? latum Reeside, n. sp., from locality in SW.! sec. 13, T. 23 N ., R. 79 W., Carbon 

County, Wyo. Side, end, and inside views, and cross section of part of a large whorl (U.S. 

Nat. Mus. catalogue No. 32355) ...... _ .... .. .................... _. _ ... ___ .............. _. 37 

FIGURES 17-20. Cardioceras? incertum Reeside, n. sp., from Freezeout Hills, Albany County, Wyo. ________ -___ 36 

17. Front view of type with part of outer whorl removed. 

18, 19. Cross section and'suture of type (U.S. Nat. Mus. catalogue No. 32356). 

20. Cross section of a largQ specimen referred to this species (U.S. Nat. Mus. catalogue No. 

32357), figured in Plate XXI. . 

All specimens from Sundance formation. 

59 
PLATE XXI. 

Cardioceras? incer-tum Reeside, n. sp.', from Sundance forniation in Freezeout Hills, Albany County, Wyo. Side 

view of a large specimen referred tO this species (U. S. Nat. Mus. catalogue No. 33257) •...• 

60 

Page. 

36 
U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 118 PLATE XXI 

JURASSIC AMMONITES. 
U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 118 PLATE XXII 

'' 

2 

JURASSIC AMMONITES. 
PLATE XXII. 

Page. 

FIGURES 1, 2. Cardioceras? incertum Reesid~, n. sp ., from Sundance formation in Freezeout Hills, Albany 

County, Wyo. Side views of the type specimen (U.S. Nat. Mus. catalogue No. 32356).. . . 36 

61 
.... 

PLATE XXIII. 

FIGURES 1-3. Cardioceras? albaniense Reeside, n. sp., from Sundance formation in Free~eout Hills, Albany 

62 

County, Wyo ............................................. . .......................... ~. 

1, 2. Side and back views (X!) of type (U.S. Nat. Mus. catalogue No. 28448). 

3. End view of part of an inner whorl of same. 

w I ... ~ ;" ... ,, ... "', .... 

I 

Page. 

35 
lJ. S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 118 PLATE XXIII 

JURASSIC AMMONITES. 
U, S. GEOLOGICAL SURVEY 

PROFESSIONAL PAPER 118 PLATE XXIV 

6 

7 

8 

JURASSIC AMMONITES. 
PLATE XXIV. 

Page. 

FIGURES 1, 2. Cardioceras? albaniense Reeside, n. sp., from Sundance formation in Freezeout Hills, Albany 

County, Wyo .........• : ....................... _·. _ ....... _ . _ . . . . . . . . . . . . . . . . . . . . . . . . . . . 35 

1. Suture of type specimen (U~ S. Nat. Mus. catalogue No. 28448). 

2. Side view of part of an inner whorl of the type. 

FIGURES 3, 4. Cardioceras? sp., from Sundance formation in ·Freezeout Hills, Albany County, Wyo. Side and 

back views (U.S. Nat. Mus. catalogue No. 32358) .............................. . -... , ..... · 37 

FIGURES 5-8. Amoeboceras dubium (Hyatt) Reeside, from Mariposa slate on Texas ranch, Calaveras County, 

Calif. Side views of four cotypes from Hyatt's type lot (U.S. Nat. Mus. catalogue No. 30201). 38 

63 

\, 
INDEX 

Names in italic are synonyms; figures in black face indicate descriptions; figures in italic indicate illustrations. 

Page. 

Acknowledgments for aid. . . . . . . . . . . . . . . . . . . ... . . . . . . . . . . . . . . . . . . . 5 

Ammonites cordiformis Meek and Hayden ................ _. ....... 21,34 

cordiformis var. distans Whitfield............................. 24 

(Aegoceras) subtumidum Whitfield and Hovey (part) ...... 15, 16,17 

(Amaltheus) cordiformis Meek and Hayden (part) ....... 201 21,23,31 

Amoeboceras Hyatt.............................................. 18 

dubi~ (Hyatt) Reeside ....... _ ......... _ ........ _ ...... _ ... 88,63 

Buckman, S. S., cited ........ : ......................... ,......... 14 

Cadoceras (Fischer) Nikitin ........ : .......................... · ... 12-18 

Cardioceras N eumayr and Uhlig ................... __ ....... . . . . . . 13 

alaskense Reeside, n. sp ...................................... 18,45 

americanum Reeside, n. sp ....... _ ...................... 18-19,45 

auroraense Ree~ide, n. s·p ......................... _ ........... 19,49 

bellefourchense Reeside, n. sp ............................ 20,50,51 

canadense Whiteaves .......... _ ......................... 20-21, 56 

cordiforme (Meek and Hayden) Neumayr ..... ..... . 21-22,46,47,48 

crassum Reeside, n. sp ................................ 23,51,52,53 

crookense Reeside, n. sp ......... _ ............ _ .... _ ...... 23-24,48 

distans (Whitfield) Neumayr .......... _ ..•.. _ .. _ ..... 24-25,54,55 

(Whitfield) var. depressum Reeside, n. var ............ 25, 54,55 

dubium Hyatt ....... :............. . . . . . . . . . . . . . . . . . . . . . . . . . . . 38 

haresi Reeside, n. sp ........ _ .. _ ............. _ ............ 25-26, 58 

hyatti Reeside, n. sp ......... ~ ........ _ ........... ... ..... 26-27, 54 

lillooetense Reeside, n. sp .................................. 27,56 

martini Reeside, n. sp .................................... 27-28,48 

obtusum Reeside, n. sp ... : ............. ... _ . __ ....... .. ..... 28, 59 

plattense Reeside, n. sp .................................. 28-29,48 

russelli Reeside, n. sp ......... -....................... 29-30,52,5~ 

schucherti Reeside, n. sp' .............. _ ..................... 30,50 

spiniferum Reeside, n. sp .................. . . . ., ...... 3()-31,57,58 

stan toni Reeside, n. sp ....................................... 31,54 

var. obesum Reeside, n. var .............................. 31,54 

stillwelli Reeside, n. sp ....... _ ............................ 31-32,45 

sundancense Reeside, n. sp ............................ 32-33,56,57 

vertebrale (Sower by) var. a Ilowaisky .. , .............. _...... 25 

whiteavesi Reeside, n. sp .................................... 33,56 

whitfieldi Reeside, n. sp ..................................... 34, 45 

wyomingense Reeside, n. sp .............................. 34-35, 54 

Cardioceras? albaniense Reeside, n. sp ............... : .... 31)-36, 62,63 

incertum Reeside, n. sp ....................... .' ....... 36,59,60,61 

latum Reeside, n. sp ........................ . .................. 37, 59 

sp. undet ............................................. , . . 37-38, 63 · 

whitneyi Smith ...................................... ,........ 38 

Cardioceras-bearing beds, age of. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 11 

Cardioceratidae H. Douville, genera included in, descrip ions of... 12· 

Chamoussetia R. Douville ....................... ·...... . .. . . .. . .. 13 

Comparative list of American and European species.... 9-10 

Correlation of formations containing the fossils.......... . . . . . . . . . lG-11 

64 

Page. 

Distribution of species by formations............................. 5-.,6 

by localities ....................... . ................. ---~----. 7-9 

Douville, Robert, cited ......................................... :. 12 

Eboraciceras Buckman........................................... 14 

Ellis formation, age of ............. :; . .. . . .. . . . . .. .. .. . .. . .. .. .. . .. 10 

European and American species, comparative list of.............. 9-10 

Hayden, F. V., Meek, F. B., and, cited .......................... 21-22 

Hovey, E. 0., Whitfield, R. P., and, cited ........ .'.............. 16 

Hyatt, Alpheus, cited............................................ 38 

Localities, distribution of species by ............. __ ............... 7 

list of. . . . . .. . . . . . . . . .. . . .. .. . . .. . .. . . . . . . . . . . . . . . . . . . . . . . . . . . 8-9 

Longaeviceras Buckman......................................... 14 

Macrocephalites von Sutner............ . . . . . . . . . . . . . . . . . . . . . . . . . . 12 

Mariposa slate, age of..... . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 10 

Meek, F. B., and Hayden, F. V., cited ............................ 21-22 

Names, application of. ............................ _ .. ~ ......... : . .5 

Pachyceras Bayle............................ . . . . . . . . .. . . . . .. . . .. 13 

Prorsicer[l.S Buckman........................................... . 14 

Pseudocadoceras Buckman...................................... 14 

Publications cited ................................................ 11-12 

Quenstedticeras Hyatt................... . . . . . . . . . . . . . . . . . . . . . . . . . 13 

collieri Reeside, n. sp ................. ___ ................. 14-15,40 

cordiforme (Meek and Hayden) Hyatt........................ 21 

hoveyi Reeside, n. sp ..................................... 15,40,41 

subtumidum (Whitfield and Hovey) Reeside ............... 16,43 

suspectum Reeside, n. sp ....................... 16-17,41,42,43,44 

tumidum Reeside, n. sp .............................. 17-18,43,44 

Reeds, C. A., acknowledgment to. . .. .. .. .. .. .. .. . .. . .. . . . .. .. .. . 5 

Stanton, T. W., acknowledgment to.............................. 5 

Stepheoceras Buckman..................... . . . . .. . . . . . . . . . . . .. . . . 13 

Sundance formation, age of................ . . . . . . . . . . . . . . . . . . . . . . . 18 

Vertumniceras Buckman .. .. .. . . . . . .. .. . . .. . .. . .. . .. .. .. . .. . .. .. 14 

Whiteaves, J. F., cited .......... · ................ ,................ 20 

Whitfield, R. P.,cited........................................... 24 

· and Hovey, E. o., cited ......... _ ........... _................ 16 

Wyoming, map of, showing localities where Cardioceratidae were 

obtained ............... ~............................. 8 

ADDITIONAL COPIES 

OF TmS PUBLICATION MAY BE PROCURED FROM 

THE SUPERINTENDENT OF DOCUMENTS 

GOVERNMENT PRINTING OFFICE 

WASIDNGTON, D. C. 

AT 

25 CENTS PER COPY 

.6.