Reproductive and Endocrine Biomarkers in Largemouth 

Bass (Micropterus salmoides) and Common Carp 

(Cyprinus carpio) from United States Waters

Steven L. Goodbred1, Stephen B. Smith2, Patricia S. Greene3, Richard H. Rauschenberger4, 

and Timothy M. Bartish5

1

 U.S. Geological Survey

 Biological Resources Discipline

 Sacramento, California

2

 U.S. Geological Survey

 Biological Resources Discipline

 Reston, Virginia

3

 U.S. Geological Survey

 Water Resources Discipline

 Reston, Virginia

4

 U.S. Fish and Wildlife Service

 Jacksonville, Florida

5

 U.S. Geological Survey

 Biomonitoring of Environmental Status and Trends

 Biological Resources Discipline

 Ft. Collins, Colorado

Data Series 2006-227

U.S. Department of the Interior

U.S. Geological Survey

Prepared in cooperation with the

U.S. Fish and Wildlife Service
ii

U.S. Department of the Interior

DIRK KEMPTHORNE, Secretary

U.S. Geological Survey

Mark D. Myers, Director

U.S. Geological Survey, Reston, Virginia: 2007

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Suggested Citation:

Goodbred, S.L., Smith, S.B., Greene, P.S., Rauschenberger, R.H., Bartish, T.M., Reproductive and Endocrine 

Biomarkers in Largemouth Bass (Micropterus salmoides) and Common Carp (Cyprinus carpio) from United 

States Waters: U.S. Geological Survey Data Series 2006, 99 p.

This report can be accessed online at:

USGS Publications Warehouse—http://pubs.usgs.gov/ds/ds2006-227

and

USGS Florida Integrated Science Center—http://fisc.er.usgs.gov/Endocrine_Data_Report.pdf
Contents

Abstract...........................................................................................................................................................1

Introduction.....................................................................................................................................................1

Acknowledgments and Dedication.............................................................................................................2

Materials and Methods.................................................................................................................................3

Site Locations........................................................................................................................................3

Sex Steroid Hormones .........................................................................................................................6

Vitellogenin ............................................................................................................................................6

Histological Determination of Sexual Maturation...........................................................................7

Results .............................................................................................................................................................7

References Cited..........................................................................................................................................98

Figure

1. Map showing Regional boundaries and collection sites for common carp, 

largemouth bass, and both common carp and largemouth bass .........................................3

Tables

1. Description of all study sites by region and species collected during 1994–97 ............4-6

2. National summary statistics (mean ± SE and ranges) for: fish size, biomarkers 

and reproductive stage for COMMON CARP and LARGEMOUTH BASS collected 

during non-reproductive (mid-August through March 1994–1997) and reproductive

 (April through mid-August 1995–1997) seasons .....................................................................8

3. Site and regional mean values for biomarker and ancillary data for (A) COMMON 

CARP and (B) LARGEMOUTH BASS collections made at all sites in U.S. waters 

from March 1994 through August 1997 ...............................................................................9-26

4. All individual COMMON CARP values for biomarker and ancillary data 

from collections made at sites in U.S. waters from August 1994 through 

October 1997..........................................................................................................................27-80

5. All individual LARGEMOUTH BASS values for biomarker and ancillary data 

from collections made at sites in U.S. waters from March 1994 through 

August 1997............................................................................................................................81-97

iii
Selected Abbreviations and Acronyms

°C degree Celsius

ELISA Enzyme-Linked Immunosorbent Assay

g gram

mL milliliter

mm millimeter

μg microgram

μL microliter

ng nanogram

nm nanometer

N number of

pg picogram

NaCl sodium chloride

NaPO4 sodium phosphate

TBST tris-buffered saline

BEST Biomonitoring of Environmental Status and Trends Program (of the USGS)

CERC Columbia Environmental Research Center (of the USGS)

FISC Florida Integrated Science Center (of the USGS)

NAWQA National Water Quality Assessment Program (of the USGS)

USGS U.S. Geological Survey

iv
Abstract

A nationwide reconnaissance investigation was initiated 

in 1994 to develop and evaluate a suite of reproductive and 

endocrine biomarkers for their potential to assess reproductive 

health and status in teleost (bony) fish. Fish collections were 

made at 119 sites, representing many regions of the country 

and land- and water-use settings. Collectively, this report will 

provide a national and regional benchmark and a basis for 

evaluating biomarkers of endocrine and reproductive function. 

Approximately 2,200 common carp (Cyprinus carpio) and 650 

largemouth bass (Micropterus salmoides) were collected from 

1994 through 1997. The suite of biomarkers used for these 

studies included: the plasma sex-steroid hormones, 17β-estradiol (E2) and 11 ketotestosterone (11KT); the ratio of E2 to 

11KT (E2:11KT); plasma vitellogenin (VTG); and stage of 

gonadal development. This data report provides fish size, stage 

and reproductive biomarker data for individual fish and for site 

and regional summaries of these variables.

Introduction

Freshwater ecosystems have been shown to be the 

ultimate sink for many anthropogenic chemicals that can 

potentially affect aquatic communities and the terrestrial 

species that depend on them (Schmitt and others, 1999; Gross 

and others, 2003; Hoffman and others, 2003). Excess nutrients, runoff of pesticides from agricultural fields, industrial 

effluents, wastewater/sewage discharge, and urban nonpointsource contamination may contribute to the deterioration of 

freshwater aquatic environments. Many field studies designed 

to assess the effects of environmental contaminants in aquatic 

ecosystems focus efforts at local “hot spots,” to evaluate 

the occurrence of endocrine disruption or the prevalence of 

external and internal anomalies (Baumann and others, 1991; 

Smith and others, 2002a,b). Although these focused studies 

have great value in determining cause-and-effect on select fish 

species at specific sites, a different approach is required to 

describe geographic differences, which is critical to determine 

if contaminant-induced effects might threaten populations 

of aquatic species nationally or regionally (Stahl and Clark, 

1998). These data and the development of a battery of biomarkers designed to monitor the health and/or reproductive 

status of fish populations contribute to our understanding the 

well-being of aquatic resources, and help generate hypotheses 

concerning cause-and-effect relationships between environmental stressors (e.g., chemical contaminants) and fish health. 

A variety of fish species are frequently used as sentinels 

or bioindicators for monitoring environmental quality in 

aquatic ecosystems. Teleost (bony fish) species are frequently 

preferred because of their wide geographic distribution, are 

relatively easy to sample, have a high reproductive rate, are 

integral components of the aquatic ecosystem, and are both 

commercially and recreationally important. Teleost fish in 

temperate regions undergo a seasonal reproductive cycle that 

has two distinct phases: (1) gonadal recrudescence/development, and (2) spawning. Cycling between these two phases is 

governed by both photoperiod and temperature, rainfall, water 

flow, as well as by other environmental cues (Kime, 1998; 

Gross and others, 2002, 2003). Most temperate fish in North 

America, such as black bass (Micropterus spp.), spawn during 

spring and early summer, but spawning time varies widely 

across a range of climatic conditions. Some species, such as 

common carp (Cyprinus carpio), may spawn as early as March 

in the southern U.S. to as late as mid-August in the northern 

U.S. (Panek, 1987). 

Reproductive and Endocrine Biomarkers in Largemouth 

Bass (Micropterus salmoides) and Common Carp 

(Cyprinus carpio) from United States Waters

By Steven L. Goodbred, Stephen B. Smith, Patricia S. Greene, Richard H. Rauschenberger, 

and Timothy M. Bartish
The two fish species selected for this study and described 

in this report, common carp and largemouth bass (Micropterus 

salmoides), are widely distributed across the United States. 

Although both species are oviparous (egg-laying), their reproductive strategies differ; common carp spawn by broadcasting 

eggs into the water column, whereas largemouth bass deposit 

eggs in nests. In addition, common carp are in the minnow 

family (Cyprinidae) and are omnivores, whereas largemouth 

bass are in the sunfish family (Centrarchidae) and are predators. Because they derive their food from different habitats, 

are in different trophic levels (Lee and others, 1980), and are 

in different families, they may also differ in their exposures 

to environmental contaminants at the same site and may have 

different physiological responses (Schmitt and others, 2002). 

The biomarkers in this reconnaissance investigation 

were selected to measure a range of reproductively significant 

factors. The endocrine system is comprised of a wide variety 

of hormone-secreting cells, tissues, and ductless glands (e.g., 

the pituitary, thyroid, adrenal, and gonads), and each part plays 

an important role in growth, development, reproduction, and 

homeostasis of an organism. Endocrine hormones influence 

virtually every stage of the life cycle, from gametogenesis 

and fertilization, through development into a sexually mature 

organism. Other studies have reported correlations between 

specific impairment of reproduction (reduced fertility, hatchability and viability of offspring, impaired reproductive 

hormone activity, and/or altered sexual development and 

behavior) and elevated tissue concentrations of environmental 

contaminants (Hose and others, 1989; Tilitt and others 

1992; Harrison and others, 1999; Gross and others, 2003). 

Abnormalities of these types may be caused by alteration of 

normal endocrine function either before or after hormone 

interactions with specific cellular receptors. 

Concentrations of sex steroid hormones regulate 

important aspects of reproduction, such as spermatogenesis, 

follicular growth, and vitellogenesis. Initiation of these processes begins in the pituitary gland, which regulates gonadal 

hormone production in response to external cues (Kime, 

1998; Gross and others, 2002, 2003). In most teleost fish, 17ßestradiol is the major estrogenic hormone in females responsible for ovarian growth. The concentration of 17ß-estradiol 

increases in the late winter to early spring, immediately 

preceding the spawning season, but its concentration is lower 

during the rest of the year (Norris and Jones, 1987; Down and 

others, 1990; Kime, 1998; Gross and others, 2002). In teleost 

fish, the primary sex steroid hormone in males is an androgen, 

11-ketotestosterone, which is responsible for spermatogenesis 

(Borg, 1994; Mylonas and others, 1997; Cavaco and others, 

1998; Cavaco and others, 1999). Concentrations of estrogens 

and androgens, and the ratio of two sex steroid hormones, 17β

estradiol/11-ketotestosterone (E2:11KT) have been utilized 

to assess the endocrine system in fish (Goodbred and others, 

1997; Gross and others, 2002; Schmitt and others, 2002). 

Vitellogenesis is the process of producing the protein 

vitellogenin in the liver in response to stimulation by estrogenic 

sex-steroid hormones. Vitellogenin is then processed by the 

 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Acknowledgments and Dedication

This study was conducted by several U.S. Geological Survey (USGS) programs and research centers, 

including: the National Water Quality Assessment (NAWQA) Program, Biomonitoring of Environmental 

Status and Trends Program (BEST), the Florida Integrated Science Center (FISC), the Columbia 

Environmental Research Center (CERC), and personnel from both the Biological and the Water Resources 

disciplines. All biomarker analyses were conducted by the Joint Ecotoxicology Program of FISC and the 

University of Florida, College of Veterinary Medicine, Department of Physiological Sciences, and we 

wish to thank Tim Gross, Carla Wieser, Jon Wiebe, Nancy Denslow, and Kevin Kroll for these analyses. 

We also wish to thank members of NAWQA’s biological team; Rachel Muir and Rod DeWeese for their 

field efforts and the review of this document and Wade Bryant for the expertise and field efforts. We 

also wish to thank the members of the BEST Program, Tim Bartish, Pat Anderson, and Jim Coyle for 

their expertise and support. In addition, we wish to acknowledge the critical support of the U.S. Fish and 

Wildlife Service in the field efforts nationally. The authors also want to thank all of the personnel from 

the various USGS NAWQA study units involved with the fish collections. Additional thanks to Steve 

Sorenson and Mike Yurewicz from the Northeast Region, and Christopher Schmitt from CERC. 

Finally, and most importantly, we dedicate this report to our close friend and colleague, Tim Bartish,

who is missed by all who knew him. 
ovary and transferred into developing follicles to form a large 

portion of the egg yolk. Male fish produce significant amounts 

of vitellogenin when exposed to estrogen or estrogenic chemicals (Purdom and others, 1994; Bromage and Cumaranatunga, 

1998). Therefore, concentrations of vitellogenin have been 

recently recognized as an important biomarker for assessment 

of the reproductive status or health of fish (Denslow and others, 

1999; Gross and others 2002, 2003; Schmitt and others, 2002). 

Vitellogenin provides an additional physiological biomarker 

for assessing reproductive health and predicting the ability of 

various compounds to alter the endocrine and reproductive 

systems. Males do not normally produce significant amounts 

of vitellogenin, but males treated with an estrogen produced 

significant vitellogenin concentrations. Therefore, it has 

been hypothesized that the elevated concentrations of plasma 

vitellogenin in males might serve as a reliable biomarker 

of exposure to environmental contaminants with estrogenic 

properties or endocrine effects (Heppell and others, 1995; 

Sumpter and Jobling, 1995; Palmer and Selcer, 1996; Denslow 

and others, 1996 a,b; 1997). Gonadal stage (stage of follicular 

or sperm maturation as determined by histological techniques) 

is also a potentially useful measure of the presence and effects 

of environmental stressors in teleost fish. 

The purpose of this data report is to present multiple 

biomarkers of reproductive/endocrine function in common 

carp and largemouth bass on a national and regional scale in 

the United States.

Materials and Methods

Site Locations

This report brings together data from sites sampled by 

a combination of two USGS monitoring/research efforts: 

(1) a national network of water-quality and bed-sediment and 

tissue sites of the USGS National Water Quality Assessment 

(NAWQA) Program (Gilliom and others, 1995); and (2) the 

sites in the Mississippi and Columbia River basins that were 

sampled by the USGS Biomonitoring of Environmental Status 

and Trends (BEST) Program (Schmitt and others, 1999; 2000).

The collection sites (fig. 1, table 1) represent a wide 

range of environmental settings and biologically important 

aquatic ecosystems across the contiguous United States; they 

are characterized by flow regime (riverine or lacustrine) and 

land use (agricultural, urban, or reference). The regional 

boundaries are rough estimates for the geographic separation 

of sites based on river basin and site proximity. There were no 

collection sites in the Southeast Region.

Figure 1. Regional boundaries and collection sites for common carp (magenta), largemouth bass (blue), and both common carp and 

largemouth bass (gold). 

Materials and Methods 

G u l f o f 

M e x i c o

A t l a n t i c O c e a n

Pa c i f i c O c e a n

Alabama

Delaware

Maryland

Maine

New York

Vermont

New

Hampshire

Massachusetts

Rhode Island

Connecticut

Virginia

North Carolina

South

Carolina

New

Jersey

Oklahoma

Kansas

Nebraska

South Dakota

North Dakota

Kentucky

Michigan

Indiana Ohio Illinois

Tennessee

Arkansas

Missouri

Iowa

Minnesota

Wisconsin

Pennsylvania

Georgia

Mississippi

Florida

Texas

Louisiana

New 

Mexico

Wyoming

Utah

Arizona

Nevada

California

Montana

Idaho

Washington

Oregon

Colorado

West

Virginia

120°N

120°N

115°N

115°N

110°N 105°N 95°N

110°N

100°N

100°N 90°N

90°N

85°N 80°N

80°N

75°N 70°N

105°N 95°N 85°N 75°N

30°W 30°W

40°W

45°W 45°W

40°W

35°W 35°W

0 100 200 300 400 MILES

0 100 200 300 400 500 600 KILOMETERS

N

NORTHEAST 

REGION

NORTHWEST 

REGION

SOUTHWEST 

REGION

NORTHERN 

MID-CONTINENT REGION

SOUTHERN 

MID-CONTINENT REGION

SOUTHEAST 

REGION
 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Site name and location Site code Latitude Longitude

Flow regime Land use Species

Riverine Lacustrine

ReferenceAgriculture Urban Common 

carp

Largemouth 

bass 

Northeast Region

Connecticut Riv @ Thompsonville, CT NECT-1 41 ̊ 59 ́ N 72 ̊ 36 ́ W R U C B

Potomac Riv @ Anacostia Riv NEDC-1 38 ̊ 52 ́ N 76 ̊ 59 ́ W R U C B

Woods Pond at Lenox, MA NEMA-1 42 ̊ 21 ́ N 73 ̊ 14 ́ W L U B

Hudson Riv @ L Lazume, NY NENY-1 43 ̊ 18 ́ N 73 ̊ 50 ́ W R R C

Mohawk Riv @ Frankfort, NY NENY-2 43 ̊ 03 ́ N 75 ̊ 04 ́ W R U C B

Swinging Bridge Res nr Monticello, NY NENY-3 41 ̊ 37 ́ N 74 ̊ 46 ́ W L R B

Hudson Riv @ Poughkeepsie, NY NENY-4 41 ̊ 42 ́ N 73 ̊ 55 ́ W R U C B

Hudson Riv @ Catskill, NY NENY-5 41 ̊ 13 ́ N 73 ̊ 50 ́ W R U B

Ohio Riv @ Marietta, OH NEOH-1 39 ̊ 24 ́ N 81 ̊ 26 ́ W R U C B

Allegheny Riv @ Natrona, PA NEPA-1 40 ̊ 39 ́ N 79 ̊ 41 ́ W R R C

Shenandoah Riv @ Melville, WV NEWV-1 39 ̊ 16 ́ N 77 ̊ 46 ́ W R A C

Leetown Res at Leetown, WV NEWV-2 39 ̊ 23 ́ N 77 ̊ 56 ́ W L R C B

Northern Mid-Continent Region

South Platte Riv @ Denver, CO NMCO-1 39 ̊ 44 ́ N 104 ̊ 58 ́ W R U C

South Platte Riv @ Kersey, CO NMCO-2 40 ̊ 25 ́ N 104 ̊ 38 ́ W R A C

Arkansas Riv @ J Martin Res, CO NMCO-3 38 ̊ 03 ́ N 102 ̊ 56 ́ W L A C B

Iowa Riv @ Morengo, IA NMIA-1 41 ̊ 50 ́ N 92 ̊ 11 ́ W R A C

South Fork at Iowa Riv @ Rowan, IA NMIA-2 42 ̊ 44 ́ N 93 ̊ 35 ́ W R A C

Iowa Riv nr New Providence, IA NMIA-3 42 ̊ 19 ́ N 93 ̊ 09 ́ W R A C

South Skunk River Oskaloosa, IA NMIA-4 41 ̊ 18 ́ N 92 ̊ 38 ́ W R A C

Cedar Riv @ St. Chas City, IA NMIA-5 43 ̊ 03 ́ N 92 ̊ 40 ́ W R A C

Des Moines Riv @ Keosauqua, IA NMIA-6 40 ̊ 44 ́ N 91 ̊ 59 ́ W R A C

Mississippi Riv @ Dubuque, IA NMIA-7 42 ̊ 26 ́ N 90 ̊ 35 ́ W R A C B

Mississippi Riv @ Guttenburg, IA NMIA-8 42 ̊ 43 ́ N 91 ̊ 01 ́ W R A C B

Illinois Riv @ Beardstown, IL NMIL-1 40 ̊ 07 ́ N 90 ̊ 20 ́ W R A C B

Sugar Creek nr New Palistine, IN NMIN-1 39 ̊ 43 ́ N 85 ̊ 53 ́ W R U C

Brushy Creek Res nr North Vernon, IN NMIN-2 39 ̊ 01 ́ N 85 ̊ 36 ́ W L R C

White Riv @ Hazelton, IN NMIN-3 38 ̊ 29 ́ N 87 ̊ 32 ́ W R U C

Wabash Riv @ N Harmony, IN NMIN-4 38 ̊ 11 ́ N 87 ̊ 58 ́ W R A C B

Kansas Riv @ Bonner Sp, KS NMKS-1 39 ̊ 02 ́ N 94 ̊ 47 ́ W R A C

Ottertail River nr Breckenridge, MN NMMN-1 46 ̊ 15 ́ N 96 ̊ 32 ́ W R R C

Red Riv nr Breckenridge, MN NMMN-2 46 ̊ 17 ́ N 96 ̊ 35 ́ W R A C

Mississippi Riv @ L City, MN NMMN-3 44 ̊ 22 ́ N 92 ̊ 07 ́ W L U C

Missouri Riv @ Hermann, MO NMMO-3 38 ̊ 42 ́ N 91 ̊ 26 ́ W R A C B

Big Horn Riv @ Hardin, MT NMMT-1 45 ̊ 52 ́ N 107 ̊ 34 ́ W R R C

Yellowstone Riv @ Sidney, MT NMMT-2 47 ̊ 34 ́ N 104 ̊ 13 ́ W R A C

South Platte Riv below No Platte, NE NMNE-1 41 ̊ 08 ́ N 100 ̊ 54 ́ W R A C

Missouri Riv @ Nebraska City, NE NMNE-3 40 ̊ 40 ́ N 95 ̊ 49 ́ W R A C

Red Riv @ Drayton, ND NMND-1 48 ̊ 17 ́ N 97 ̊ 12 ́ W R A C

Red Riv nr Grand Forks, ND NMND-2 47 ̊ 54 ́ N 97 ̊ 01 ́ W R A C

Missouri Riv @ Garrison Dam, ND NMND-3 47 ̊ 28 ́ N 101 ̊ 26 ́ W L R C

James Riv @ Olivet, SD NMSD-1 43 ̊ 13 ́ N 97 ̊ 41 ́ W R A C

Mullet Riv @ Plymouth, WI NMWI-1 43 ̊ 46 ́ N 88 ̊ 05 ́ W R A C

North Br Milwaukee at Random Lake, WI NMWI-2 43 ̊ 02 ́ N 87 ̊ 54 ́ W R A C

Wisconsin Riv @ Woodman, WI NMWI-3 43 ̊ 05 ́ N 90 ̊ 48 ́ W R R C

Table 1. Description of all study sites by region and species collected during 1994–97.
Site name and location Site code Latitude Longitude

Flow regime Land use Species

Riverine Lacustrine

ReferenceAgriculture Urban Common 

carp

Largemouth 

bass 

Southern Mid-Continent Region

Cache Riv @ Cotton Plant, AR SMAR-1 35 ̊ 00 ́ N 91 ̊ 11 ́ W R A C

Cache Riv @ Egypt, AR SMAR-2 35 ̊ 50 ́ N 90 ̊ 57 ́ W R A C

White River nr DeValls Bluff, AR SMAR-3 34 ̊ 44 ́ N 91 ̊ 18 ́ W R A C

Arkansas Riv @ Pine Bluff, AR SMAR-4 34 ̊ 16 ́ N 94 ̊ 57 ́ W R A C B

Ohio Riv @ Metropolis, IL SMIL-2 37 ̊ 07 ́ N 88 ̊ 39 ́ W R U C B

Tensas Riv @ Tendell, LA SMLA-1 32 ̊ 17 ́ N 91 ̊ 07 ́ W R A C

Mississippi Riv @ Luling, LA SMLA-2 29 ̊ 59 ́ N 90 ̊ 25 ́ W R U C

Red Riv @ Alexandria, LA SMLA-3 31 ̊ 20 ́ N 92 ̊ 27 ́ W R A C B

Little Riv Ditch at Moorehouse, MO SMMO-1 36 ̊ 50 ́ N 89 ̊ 43 ́ W R A C B

Mississippi Riv @ Cape Girardeau, MO SMMO-2 37 ̊ 18 ́ N 89 ̊ 31 ́ W R A C

Missouri Riv @ Hermann, MO SMMO-3 38 ̊ 42 ́ N 91 ̊ 26 ́ W R A C B

Big Sunflower Riv @ Anguilla, MS SMMS-1 32 ̊ 58 ́ N 90 ̊ 50 ́ W R A C

Boque Phalia at Leland, MS SMMS-2 33 ̊ 24 ́ N 90 ̊ 54 ́ W R A C

Steele Bayou at Rolling Fork, MS SMMS-3 32 ̊ 54 ́ N 90 ̊ 52 ́ W R A C

Yazoo Riv @ Redwood, MS SMMS-4 32 ̊ 55 ́ N 90 ̊ 57 ́ W R A C

Arkansas Riv @ Keystone Res, OK SMOK-1 36 ̊ 13 ́ N 96 ̊ 15 ́ W L R C B

Canadian Riv @ Eufaula, OK SMOK-2 35 ̊ 16 ́ N 95 ̊ 34 ́ W L A C B

Verdigris Riv @ Oologah, OK SMOK-3 36 ̊ 31 ́ N 95 ̊ 33 ́ W L A C B

Wolf River nr LaGrange, TN SMTN-1 35 ̊ 02 ́ N 89 ̊ 15 ́ W R A B

Mississippi Riv @ Memphis, TN SMTN-2 35 ̊ 08 ́ N 90 ̊ 03 ́ W R U C B

Real Foot L NWR nr Union City, TN SMTN-3 36 ̊ 21 ́ N 89 ̊ 58 ́ W L R B

Cumberland Riv @ Clarksville, TN SMTN-4 36 ̊ 32 ́ N 87 ̊ 22 ́ W R R C B

Tennessee Riv @ Savannah, TN SMTN-5 35 ̊ 12 ́ N 88 ̊ 18 ́ W R A C B

Chickasaw L NWR nr Ripley, TN SMTN-6 35 ̊ 44 ́ N 89 ̊ 32 ́ W L R B

Ray Roberts L nr Gainesville, TX SMTX-1 33 ̊ 45 ́ N 96 ̊ 40 ́ W L U B

Mountain Creek L nr Dallas, TX SMTX-2 32 ̊ 45 ́ N 96 ̊ 51 ́ W L U C B

Clear Creek nr Sanger, TX SMTX-3 33 ̊ 20 ́ N 97 ̊ 11 ́ W R A C

Trinity River below Dallas TX SMTX-4 32 ̊ 44 ́ N 96 ̊ 46 ́ W R U C

White Rock Lake, Dallas, TX SMTX-5 32 ̊ 50 ́ N 96 ̊ 43 ́ W L U B

Chambers L nr Corscama, TX SMTX-6 32 ̊ 13 ́ N 96 ̊ 33 ́ W L U B

Red Riv @ L Texoma, TX SMTX-7 33 ̊ 52 ́ N 96 ̊ 47 ́ W L A C B

Northwest Region

Snake Riv @ Hagerman, ID NWID-1 42 ̊ 47 ́ N 114 ̊ 56 ́ W R A C B

Snake Riv @ Lewiston, ID NWID-2 46 ̊ 24 ́ N 117 ̊ 02 ́ W R A C

Flathead Riv @ Creston, MT NWMT-1 48 ̊ 09 ́ N 114 ̊ 11 ́ W R A B

Willamette Riv below Portland, OR NWOR-1 45 ̊ 28 ́ N 122 ̊ 39 ́ W R U C B

North Fork Willamette Riv @ Spgfield, OR NWOR-2 44 ̊ 07 ́ N 123 ̊ 05 ́ W L U C

Columbia Riv @ Warrendale, OR NWOR-3 45 ̊ 38 ́ N 121 ̊ 58 ́ W R A C

Willamette Riv @ Oregon Cty, OR NWOR-4 45 ̊ 19 ́ N 122 ̊ 39 ́ W R A C

Billy Clapp L nr Stratford, WA NWWA-1 47 ̊ 16 ́ N 121 ̊ 01 ́ W L R C

Royal L nr Othello, WA NWWA-2 46 ̊ 54 ́ N 119 ̊ 54 ́ W L A C

Columbia Riv @ Pasco, WA NWWA-3 46 ̊ 31 ́ N 119 ̊ 16 ́ W R A C B

Columbia Riv @ Vernita Bg, WA NWWA-4 46 ̊ 37 ́ N 119 ̊ 51 ́ W R A C

Snake Riv @ Ice Harbor Dam, WA NWWA-5 46 ̊ 14 ́ N 118 ̊ 53 ́ W R A C

Yakima Riv @ Granger, WA NWWA-6 46 ̊ 20 ́ N 120 ̊ 12 ́ W R A C B

Table 1. (Continued) Description of all study sites by region and species collected during 1994–97.

Table 1 
 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Site name and location Site code Latitude Longitude

Flow regime Land use Species

Riverine Lacustrine

ReferenceAgriculture Urban Common 

carp

Largemouth 

bass 

Southwest Region

Colo Riv inflo to L Mead @ Pearce Ferry, AZ SWCZ-1 36 ̊ 15 ́ N 114 ̊ 00 ́ N R R C

Don Pedro Res nr La Grange, CA SWCA-1 37 ̊ 39 ́ N 120 ̊ 56 ́ N L R C B

San Joaquin Riv @ Salt Slough, CA SWCA-2 37 ̊ 34 ́ N 121 ̊ 09 ́ W R A C B

San Joaquin Riv @ Vernalis, CA SWCA-3 37 ̊ 36 ́ N 121 ̊ 10 ́ W R A C B

Virgin Riv inflo to L Mead, NV SWNV-1 36 ̊ 33 ́ N 114 ̊ 20 ́ W R R C

Muddy Riv inflo to L Mead, NV SWNV-2 36 ̊ 17 ́ N 114 ̊ 30 ́ W R A C

Las Vegas Bay nr Las Vegas Wash, NV SWNV-3 36 ̊ 06 ́ N 114 ̊ 48 ́ W L U C

Callville Bay, NV SWNV-4 35 ̊ 16 ́ N 114 ̊ 24 ́ W R R C

Lake Mohave nr Laughlin, NV SWNV-5 35 ̊ 28 ́ N 114 ̊ 40 ́ W L U C

Bear Riv @ Corinne, UT SWUT-1 41 ̊ 33 ́ N 112 ̊ 06 ́ W R R C

Jordon Riv @ 17th Street, UT SWUT-2 40 ̊ 41 ́ N 112 ̊ 06 ́ W R U C

Jordon Riv @ Cudahy, UT SWUT-3 40 ̊ 39 ́ N 111 ̊ 58 ́ W R A C

Jordon Riv @ Utah Lake, UT SWUT-4 40 ̊22 ́ N 111 ̊54 ́ W R A C

Weber Riv @ Ogden Bay, UT SWUT-5 41 ̊13 ́ N 112 ̊40 ́ W R U C

Table 1. (Continued) Description of all study sites by region and species collected during 1994–97.

of 17ß-estradiol or 11- ketotestosterone in 50 μL of charcoalstripped plasma. Values of R2 for correlations between actual 

and measured amounts during the calibration were 0.93 for 

17ß-estradiol and 0.88 for 11-ketotestosterone. Interassay and 

intraassay coefficients of variation were 7.3 and 9.5 percent, 

respectively, for plasma 17ß-estradiol; and 9.1 and 8.7 percent, 

respectively, for plasma 11-ketotestosterone (Bevans and others, 

1996; Goodbred and others, 1997; Schmitt and Dethloff, 2000).

Vitellogenin 

Vitellogenin concentrations in plasma of both species 

were assayed and quantified by capture Enzyme-Linked 

Immunosorbent Assay (ELISA) as previously described by 

Goodbred and others (1997) and summarized below. Initially, 

vitellogenin from bass was purified by ion-exchange chromatography (Denslow and others, 1999) for use in the standard. 

The monoclonal antibody Mab HL 1080 1C8-3C11 was used 

in the ELISA assay. This antibody reacts specifically to bass 

vitellogenin and not with other plasma proteins. Purified 

antibody was diluted to 10 μg/mL in phosphate-buffer saline 

(0.01 M NaPO4, 150 mm NaCl, pH 7.4) and coated onto 

96-well microlitre plates (50 μg/mL per well), and stored 

overnight at 4 °C. Plates were washed with tris-buffered 

saline (TBST) buffer (10 mM Tris, pH 7, 150 mM NaCl, 

0.5% Tween), blocked with 0.1 percent bovine serum albumin 

in TBST for 2 hours at room temperature, and thoroughly 

Sex Steroid Hormones

Plasma samples from both species were analyzed for 

17ß-estradiol and 11-ketotestosterone using radioimmunoassay procedures. Plasma samples (50 μL) were extracted 

twice with 5-mL diethyl ether prior to radioimmunoassay 

analysis. Each sample was analyzed in duplicate for both 17ßestradiol and 11-ketotestosterone and corrected for extraction 

efficiencies of 92 ±2.8 percent and 86 ± 3.3 percent, respectively. Standard curves were prepared in a buffer with known 

amounts of radioinert 17ß-estradiol or 11-ketotestosterone 

(1, 5, 10, 25, 50, 100, 250, 500, and 1,000 pg). The minimum 

detectable concentration was 6.4 pg/mL for 17ß-estradiol and 

8.1 pg/mL for 11-ketotestosterone. Cross-reactivities with other 

steroids for the 17ß-estradiol antiserum were as follows: 11.2 

percent for estrone, 1.7 percent for estriol, less than 1.0 percent 

for 17alpha-estradiol and androstenedione, and less than 0.1 

percent for all other steroids examined. Cross-reactivities of 

the 11-ketotestosterone antiserum with other steroids were: 9.7 

percent for testosterone, 3.7 percent for a-dihydrotestosterone, 

less than 1.0 percent for androstenedione, and less than 0.1 

percent for all other steroids examined. A pooled sample (approximately 275 pg of 17ß-estradiol/mL and 220 pg 11-ketotestosterone/mL) was assayed serially in 10, 20, 30, 40, and 50 μL 

volumes (final volume of 50 μL with charcoal-stripped plasma). 

Further characterization of the assays involved measurement 

of known amounts (1, 2, 5, 10, 25, 50, 100, 250, and 500 pg) 
washed again three times with TBST. Plasma samples were 

diluted from 1:100 to 1:5,000 in 0.1 percent bovine serum 

albumin in TBST and 50 μg was added in duplicate to microtitre 

plate wells and incubated overnight. Standard curves were from 

10 to 1,000 ng/mL, and were constructed in male plasma at 

the same dilution as the sample to be analyzed. Of this solution, only 50 μg were placed in the well. Although the assay 

itself is very sensitive, plasma samples must be diluted at 

least to 1:100 to eliminate interferences that affect the assay, 

thus making the assay sensitive to 0.001 mg/mL. Male control 

plasma was made from composites of plasma from fish 

collected at an uncontaminated site, which was shown by 

Western Blot assay to have no vitellogenin. The next day, 

plates were washed with TBST, incubated with 50 μg per well 

rabbit anti-vitellogenin polyclonal antibody UF114, produced 

and characterized by University of Florida, diluted to 1:500, 

and incubated for 2 hours at room temperature. This discloses 

the vitellogenin captured by the monoclonal antibody in the 

first step. The polyclonal antibody was, in turn, disclosed 

by a goat anti-rabbit immunoglobulin IgG linked to alkaline 

phosphatase and incubated for 2 hours at room temperature. 

After a final series of washes with TBST, 100 μg of 

p-nitro-phenyl-phosphate in carbonate buffer (pH 9.6) was 

added to each well and incubated for 30 minutes. The intensity of yellow color that developed was quantified at 405 nm 

with an automated ELISA reader. Vitellogenin concentrations 

were calculated from standard curves after subtracting the 

small nonspecific color reaction (about 0.2 A, 405 nm) with 

male control plasma. The ELISA assay used in this study can 

detect between 10 to 100 ng of vitellogenin per well, resulting 

in a sensitivity of about 0.001 mg/mL. Each ELISA assay 

included a positive control, which was plasma with a known 

vitellogenin concentration, to test for interassay and intraassay variation. The coefficient of variation was calculated for 

each duplicate sample and, if it exceeded 10 percent, samples 

were rerun. Standard curves fit by linear regression were 

used to calculate vitellogenin concentration, with R2 values 

usually between 0.95 and 0.99 (Denslow and others, 1996 a,b; 

1997; Schmitt and Dethloff, 2000).

Histological Determination of Sexual Maturation 

Samples of male and female gonads were taken, fixed 

in the field with Bouins’ solution or 10 percent buffered 

formalin, and transferred to 100 percent ethanol in the laboratory prior to processing. Testes were cut longitudinally and 

ovaries were cut transversely. Samples were embedded in 

paraffin, sectioned to 5 μm, and stained with hematoxylin and 

eosin for histological evaluation. All tissue slides were evaluated for reproductive stage, anomalies, and gonadal parasites. 

Gonads of female fish were classified according to four 

stages of sexual maturation, based on evaluation of histological slides. Ovaries containing mostly perinucleolar oocytes 

at various stages of pre-vitellogenic growth were classified 

as undeveloped (stage 0). Ovaries showing a mixture of both 

perinucleolar and cortical alveoli oocytes were classified as 

early-vitellogenic (stage 1). Ovaries classified as early-mid 

vitellogenic (stage 2) had some vitellogenic oocytes of various 

sizes and development, with few to moderate numbers of 

vitelline granules, and few fully developed oocytes. Late 

vitellogenisis was classified as stages 3 and 4, ovaries in 

which most oocytes were at or near maximum size and 

contained numerous, densely packed vitelline granules. Stage 

4 were considered vitellogenic but post-ovulatory. Stage 5 

females were post-vitellogenic.

Male gonads were classified according to three stages 

of sexual maturation. Undeveloped testes with no spermatogenesis were considered stage 0. Testes that were classified as 

early spermatogenic (stage 1) had thick germinal epithelium, 

with diffuse pronounced proliferation and maturation of 

spermatozoa. Mid-spermatogenic (stage 2) testes had germinal 

epithelium of moderate thickness, with diffuse moderate 

proliferation and maturation of sperm. Testes classified as late 

spermatogenic (stage 3) had mostly thin germinal epithelium, 

with only scattered spermatogenic activity characteristic of 

full-grown testes and the latest stage of maturity (Goodbred 

and others, 1997; Schmitt and Dethloff, 2000). 

Results

More than 2,200 common carp and nearly 650 bass were 

collected at over 100 sites nationally. This is a relatively large 

sample size, and data from all collected fish were utilized in 

the national summary. Common carp and largemouth bass 

were collected during both non-reproductive and reproductive seasons from 1994 though 1997. Largemouth bass 

were collected only from the northeastern region during the 

reproductive season.

Summary statistics are provided for fish size, biomarkers, 

and repro­ductive stage by species (table 2) and by region and 

site (tables 3a and 3b). Biomarkers and ancillary information 

for individual common carp and largemouth bass are listed in 

tables 4 and 5, respectively. This infor­mation may serve as a 

unique and useful set of baseline values for comparison and 

for planning future studies. 

Results 
 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 2. National summary statistics (mean ± SE and ranges) for: fish size, biomarkers and reproductive stage for COMMON CARP and 

LARGEMOUTH BASS collected during non-reproductive (mid-August through March 1994–1997) and reproductive (April through 

mid-August 1995–1997) seasons. 

[N, number of fish; E2, 17ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; mm, millimeter; g, grams; pg, picograms; mg, milligram; mL, milliliter; 

ND, nondetected]

Length

(millimeter) N Weight(gram) N

E2

(picogram per 

milliliter)

N

11KT

(picogram per 

milliliter)

N Ratio of 

E2 to KT N

VTG

(milligram per 

milliliter)

N Developmental

stage N

COMMON CARP

 Female non-reproductive season

535 ± 3.5

(278 – 920) 819 2380 ± 50.4(288 – 10000) 819 1479 ± 44.4(15 – 11401) 823 641 ± 27.5(13 – 9066) 823 3.8 ± 0.15(0.1 – 40.6) 823 12.6 ± 0.51(ND – 108) 821 2.7 ± 0.02(0 – 4) 822

 Female reproductive season

515 ± 6.3

(332 – 742) 233 2220 ± 93.1(479 – 6300) 233 1070 ± 49.3(17 - 4142) 234 659 ± 49.4(45 – 5066) 234 2.6 ± 0.18(0.0 – 24.8) 234 18.1 ± 1.64(ND – 124.8) 234 2.8 ± 0.04(0 – 4) 234

 Male non-reproductive season

507 ± 2.8 

(278 – 779) 831 1868 ± 31.4 (324 – 6407) 831 502 ± 15.8 (17 – 4546) 833 1232 ± 42.1(61 – 9699) 833 0.7 ± 0.04 (0.0 – 19.3) 833 0.1 ± 0.04  (ND – 25.4) 830 2.1 ± 0.03(0 – 3) 828

 Male reproductive season

501 ± 5.4

(297 – 763) 254 1991 ± 72.2(325 – 7300) 254 408 ± 22.1(51 – 2565) 255 1000 ± 63.8(74 – 6821) 255 0.7 ± 0.05(0.0 – 6.9) 255 0.01 ± 0.05(ND – 12.0) 255 2.5 ± 0.04(1 – 3) 253

LARGEMOUTH BASS

 Female non-reproductive season

369 ± 4.0

(231 – 595) 323 869 ± 30.8(140 – 2920) 320 717 ± 40.1(29 – 6330) 335 343 ±17.3(12 – 2203) 335 3.3 ± 0.20(0.0 – 27.8) 335 1.3 ± 0.38(ND – 77.9) 335 1.7 ± 0.04(0 – 3) 332

 Female reproductive season 

378 ± 13.7

(286 – 478) 18 872 ± 97.1(350 – 1648) 18 2710 ± 606.2(384 – 9670) 18 704 ±154.3(108 – 2306) 18 4.6 ± 0.68(1.6 – 12.4) 18 4.8 ± 0.84(ND – 12.4) 18 3.2 ± 0.15(2 – 5) 18

 Male non-reproductive season

338 ± 3.2

(226 – 515) 294 616 ± 19.5(114 – 1918) 283 375 ± 16.1(11 – 1823) 302 703 ± 41.8(15 – 7451) 302 1.0 ± 0.06(0.0 – 6.9) 302 0.0 ± 0.02(ND – 3.0) 302 2.3 ± 0.04(0 – 3) 302

 Male reproductive season

305 ± 7.8

(276 – 386) 14 450 ± 43.0(321 – 938) 14 781 ± 120.2(280 – 1621) 14 1195 ± 115.2(410 – 1706) 14 0.8 ± 0.21(0.2 – 3.0) 14 0.0 ± 0.02 (ND – 0.3) 14 3.0 ± 0.0(3 – 3) 14
Regional site 

code

(fig. 1)

N Length

(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental

stage

COMMON CARP

Northern Mid-Continent Region

 Female non-reproductive

NMCO-1 32 572 ± 11.9 2827 ± 178.7 1376 ± 148.3 668 ± 69.2 2 ± 0.4 19 ± 2.5 3 ± 0.1

(441-675) (1200-4588) (55-3249) (127-1500) (0-13) (ND-53.904) (1-3)

NMCO-2 25 533 ± 7.5 2214 ± 95.8 1519 ± 112.4 586 ± 67.6 3 ± 0.2 29 ± 1.4 3 ± 0

(480-620) (1559-3275) (559-2320) (185-1250) (2-6) (14.7-45.9) (3-3)

NMCO-3 11 404 ± 19.7 928 ± 147.4 967 ± 154.6 238 ± 48.9 6 ± 1.7 3 ± 0.9 3 ± 0.2

(342-555) (550-2150) (366-1901) (31-633) (2-22) (0.001-9.285) (1-3)

NMIA-1 10 461 ± 18.8 1353 ± 144.4 1819 ± 431.6 303 ± 42.2 6 ± 1 3 ± 0.5 3 ± 0.2

(360-530) (629-1970) (334-4819) (57-569) (2-13) (0.181-5.132) (1-3)

NMIA-2 10 500 ± 24.4 1928 ± 284.5 1578 ± 312.1 818 ± 81.1 2 ± 0.7 4 ± 0.6 3 ± 0.1

(402-595) (970-3102) (143-3377) (272-1155) (0-8) (ND-6.93) (2-3)

NMIA-3 3 645 ± 11.9 4166 ± 280.3 1521 ± 500.9 250 ± 117.5 7 ± 0.9 3 ± 0.4 3 ± 0

(624-665) (3751-4700) (819-2491) (112-484) (5-8) (2.736-4.159) (3-3)

NMIA-4 10 422 ± 11.2 969 ± 73.6 1468 ± 256.8 287 ± 25.4 6 ± 1 3 ± 0.4 3 ± 0.2

(381-482) (717-1355) (323-3276) (121-397) (1-11) (0.027-4.999) (1-3)

NMIA-5 10 490 ± 16.8 1534 ± 159 1846 ± 202.9 571 ± 108.7 6 ± 2.1 4 ± 0.7 3 ± 0

(400-570) (811-2398) (1198-3190) (135-1189) (1-24) (2.371-9.531) (3-3)

NMIA-6 10 538 ± 14.3 2164 ± 175 654 ± 71.8 280 ± 45.2 3 ± 0.4 2 ± 0.5 3 ± 0.2

(462-605) (1310-2900) (488-1220) (132-522) (1-5) (0.045-5.482) (1-3)

NMIA-7 10 510 ± 20.3 1964 ± 181.8 2262 ± 353.8 714 ± 75.4 3 ± 0.6 5 ± 1.6 3 ± 0.1

(340-560) (555-2500) (1114-4852) (351-1072) (1-8) (ND-15.47) (2-3)

NMIA-8 10 483 ± 7.5 1595 ± 69.6 1576 ± 276.7 461 ± 71.9 4 ± 1.2 6 ± 1.2 3 ± 0

(441-513) (1230-1895) (639-3172) (224-863) (1-14) (2.465-16.201) (3-3)

NMIL-1 10 421 ± 15.6 1053 ± 125.3 548 ± 92.2 471 ± 76.8 1 ± 0.3 4 ± 0.4 3 ± 0

(366-529) (616-1864) (171-1051) (101-846) (1-3) (2.316-6.344) (3-3)

NMIN-1 9 627 ± 23.3 3582 ± 448.2 809 ± 110.2 810 ± 97.9 1 ± 0.1 28 ± 1.9 3 ± 0

(524-742) (1775-5600) (504-1423) (383-1257) (1-2) (16.9-33.4) (3-3)

NMIN-2 10 633 ± 35.8 3691 ± 732.4 2367 ± 274.9 893 ± 162.8 3 ± 0.7 18 ± 1 3 ± 0.2

(510-855) (1610-8959) (828-3833) (248-1890) (1-9) (12.911-22.7) (2-3)

NMIN-3 17 569 ± 13.9 2722 ± 202 1722 ± 133 1177 ± 116.6 2 ± 0.3 30 ± 1.2 3 ± 0

(446-668) (1314-4530) (884-2799) (325-1969) (1-5) (21.4-41.1) (3-3)

NMIN-4 9 562 ± 24.8 3008 ± 542.7 570 ± 65.4 109 ± 11.1 6 ± 0.9 3 ± 0.7 2 ± 0.4

(447-718) (1800-7181) (351-920) (69-181) (3-12) (0.001-7.453) (0-3)

NMKS-1 10 619 ± 20.3 3238 ± 374.1 934 ± 109.9 837 ± 44.3 1 ± 0.1 2 ± 0.4 3 ± 0.2

(533-752) (2075-5450) (443-1583) (637-1070) (0-2) (ND-3.761) (1-3)

Table 3A. Site and regional mean values for biomarker and ancillary data for COMMON CARP collections made at all sites in U.S. 

waters from March 1994 through August 1997.

[N, number of fish; E2, 17ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; mm, millimeter; g, grams; pg, picograms; mg, milligram; mL, milliliter; 

ND, nondetected]

Table 3A 
10 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Regional site 

code

(fig. 1)

N Length

(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental

stage

NMMN-1 8 523 ± 3.3 1825 ± 64.5 3763 ± 1041.7 1198 ± 421.4 4 ± 0.8 31 ± 0.9 3 ± 0

(510-530) (1520-2020) (1086-9942) (187-3641) (3-8) (25.8-34.1) (3-3)

NMMN-2 14 533 ± 8.8 2032 ± 112.2 1968 ± 424.9 284 ± 43.5 11 ± 2.8 33 ± 1.3 3 ± 0.1

(485-610) (1467-3164) (462-6852) (63-540) (2-38) (23.4-41) (3-4)

NMMN-3 10 531 ± 14.1 1975 ± 139.8 1290 ± 149 870 ± 87 2 ± 0.2 4 ± 1.1 3 ± 0.2

(439-600) (1100-2800) (797-2058) (411-1221) (1-2) (0.054-11.701) (1-3)

NMMO-1 10 528 ± 12.8 2150 ± 180 1706 ± 149.3 936 ± 159.1 3 ± 0.7 5 ± 0.7 3 ± 0

(480-610) (1519-3474) (1143-2492) (286-2008) (1-7) (1.455-8.291) (3-3)

NMMO-4 6 494 ± 41.6 1758 ± 549.2 1472 ± 221.4 508 ± 122.6 4 ± 0.6 2 ± 0.2 3 ± 0.3

(375-640) (650-4000) (456-1923) (100-939) (2-5) (1.286-2.588) (1-3)

NMMT-1 12 572 ± 18.1 2603 ± 239.5 2410 ± 264.3 634 ± 133.2 6 ± 1.1 3 ± 0.6 3 ± 0

(474-681) (1400-3900) (1280-4285) (132-1644) (1-13) (ND-6.821) (3-3)

NMMT-2 12 479 ± 31.4 1538 ± 313.6 1919 ± 222.2 388 ± 94.5 9 ± 2.7 1 ± 0.2 2 ± 0.3

(328-672) (450-3850) (917-3350) (52-1274) (2-33) (ND-2.522) (1-3)

NMND-1 11 714 ± 25 4514 ± 430.5 1399 ± 289.3 342 ± 77.5 5 ± 1.2 27 ± 3.3 3 ± 0.1

(518-831) (1640-6000) (533-3530) (123-832) (2-11) (ND-44.1) (3-4)

NMND-2 10 488 ± 13.7 1575 ± 103.9 3543 ± 891 895 ± 250 6 ± 1.3 27 ± 7.2 3 ± 0.2

(424-534) (1182-2063) (443-8075) (57-2639) (1-13) (7.8-87.8) (2-3)

NMND-3 9 505 ± 16 1770 ± 164.1 361 ± 33 129 ± 10.8 3 ± 0.3 3 ± 0.8 2 ± 0.3

(409-565) (1050-2500) (227-502) (94-188) (2-4) (0.032-7.699) (1-3)

NMNE-1 17 533 ± 13.4 2035 ± 191.4 1077 ± 134.7 465 ± 109.9 3 ± 0.4 20 ± 3.3 3 ± 0.2

(435-680) (910-4000) (232-1959) (93-1988) (1-8) (ND-43.952) (1-4)

NMNE-2 17 542 ± 21.4 2336 ± 221.5 397 ± 72.8 534 ± 115.5 1 ± 0.4 19 ± 4.7 2 ± 0.3

(344-662) (649-3702) (15-1104) (51-1789) (0-6) (ND-72.5) (0-3)

NMNE-3 12 409 ± 20.5 1123 ± 220.9 445 ± 51.3 712 ± 161.5 2 ± 0.6 2 ± 0.5 3 ± 0.2

(345-620) (600-3500) (287-789) (124-1819) (0-6) (ND-5.774) (1-3)

NMSD-1 10 420 ± 27.7 1078 ± 300 937 ± 161.2 804 ± 152.7 2 ± 0.7 3 ± 1 2 ± 0.2

(316-588) (400-3600) (461-2112) (110-1461) (0-8) (ND-9.328) (1-3)

NMWI-1 10 441 ± 10.4 1242 ± 134.8 2370 ± 261 684 ± 118.2 5 ± 1.5 28 ± 1.5 3 ± 0.1

(398-522) (926-2405) (1276-3830) (72-1503) (1-18) (20.9-36.2) (2-4)

NMWI-2 17 511 ± 14.6 1809 ± 160.9 2584 ± 404.8 633 ± 130.2 9 ± 2.6 25 ± 2.3 3 ± 0

(402-630) (892-2994) (354-6147) (84-2020) (1-41) (16-47.5) (3-3)

NMWI-3 10 552 ± 9.3 2255 ± 108.9 1848 ± 297.6 463 ± 87.3 5 ± 0.8 3 ± 0.4 3 ± 0

(508-600) (1900-3000) (578-3449) (132-871) (1-10) (1.672-5.255) (3-3)

Summary 401 524 ± 4.5 2136 ± 57.4 1540 ± 60.9 600 ± 23.4 4 ± 0.2 14 ± 0.7 3 ± 0

(316-855) (400-8959) (15-9942) (31-3641) (0-41) (ND-87.8) (0-4)

Table 3A. (Continued) Site and regional mean values for biomarker and ancillary data for COMMON CARP collections made at all sites 

in U.S. waters from March 1994 through August 1997.

[N, number of fish; E2, 17ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; mm, millimeter; g, grams; pg, picograms; mg, milligram; mL, milliliter; 

ND, nondetected]
Regional site 

code

(fig. 1)

N Length

(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental

stage

 Female reproductive

NMIN-1 6 574 ± 28.5 3677 ± 612.5 415 ± 63.3 651 ± 155.3 1 ± 0.3 6 ± 0.4 3 ± 0.3

(494-674) (1590-5600) (181-585) (215-1344) (0-2) (4.215-6.894) (2-4)

NMIN-3 11 588 ± 16.6 3389 ± 377.4 561 ± 66.1 290 ± 52.4 3 ± 0.7 64 ± 9 3 ± 0.2

(510-667) (1863-5239) (261-882) (93-646) (1-8) (27.875-124.8) (1-4)

Summary 17 583 ± 14.3 3490 ± 316.5 509 ± 50.1 417 ± 75 2 ± 0.5 43 ± 9 3 ± 0.2

(494-674) (1590-5600) (181-882) (93-1344) (0-8) (4.215-124.8) (1-4)

 Male non-reproductive

NMCO-1 31 569 ± 8 2464 ± 109 215 ± 27 1462 ± 279.6 0 ± 0.1 1 ± 0.9 2 ± 0.1

(480-633) (1400-3700) (20-591) (198-6796) (0-2) (ND-25.427) (1-3)

NMCO-2 15 516 ± 11.4 1922 ± 149.5 118 ± 14.9 675 ± 113.6 0 ± 0.1 0 ± 0 3 ± 0.1

(440-585) (1077-3107) (31-227) (152-1733) (0-1) (ND-0.001) (2-3)

NMCO-3 7 369 ± 4.2 663 ± 16.7 435 ± 14.9 783 ± 123.9 1 ± 0.5 0 ± 0 2 ± 0.2

(347-382) (600-720) (390-505) (138-1174) (0-4) (0.001-0.001) (2-3)

NMIA-1 10 470 ± 20.2 1412 ± 190.3 491 ± 116.3 680 ± 167.9 1 ± 0.5 0 ± 0 2 ± 0

(383-540) (650-2265) (235-1455) (164-1348) (0-5) (ND-0.001) (2-2)

NMIA-2 10 520 ± 21.4 1951 ± 215.1 162 ± 45.3 578 ± 177.2 1 ± 0.5 0 ± 0 2 ± 0.1

(440-615) (1190-2971) (27-421) (75-1872) (0-6) (ND-0.001) (1-2)

NMIA-3 5 563 ± 20.6 2572 ± 329 545 ± 323.9 596 ± 225.3 1 ± 0.5 0 ± 0 2 ± 0

(508-615) (1744-3512) (66-1790) (69-1391) (0-2) (ND-0.001) (2-2)

NMIA-4 10 432 ± 12.1 982 ± 123.6 347 ± 76.9 420 ± 97.7 1 ± 0.3 0 ± 0 2 ± 0

(385-523) (324-1768) (47-890) (204-1249) (0-3) (ND-0.001) (2-2)

NMIA-5 10 480 ± 15.4 1377 ± 135.1 217 ± 16.2 1479 ± 317.6 0 ± 0.1 0 ± 0 2 ± 0.1

(395-550) (740-1965) (156-296) (406-3034) (0-1) (ND-0.001) (1-2)

NMIA-6 10 460 ± 22 1269 ± 144.8 505 ± 15.1 2891 ± 328.4 0 ± 0 0 ± 0 2 ± 0.1

(360-528) (566-1760) (441-567) (1596-4301) (0-0) (ND-0.001) (1.5-2.5)

NMIA-7 10 510 ± 10 1827 ± 138.7 1019 ± 197.2 1633 ± 244.3 1 ± 0.2 0 ± 0.3 2 ± 0.1

(470-570) (1396-2900) (579-2713) (985-3402) (0-2) (ND-2.645) (1-2)

NMIA-8 10 458 ± 9.7 1277 ± 80.5 716 ± 86 2084 ± 511.3 0 ± 0.1 0 ± 0 1 ± 0.1

(395-496) (862-1672) (321-1020) (711-6341) (0-1) (0.001-0.001) (1-2)

NMIL-1 10 405 ± 8 806 ± 37.3 309 ± 22.3 1522 ± 465.1 1 ± 0.3 0 ± 0 1 ± 0.2

(376-441) (629-961) (220-442) (113-5401) (0-3) (ND-0.001) (1-3)

NMIN-1 7 586 ± 14.4 2347 ± 146.2 804 ± 140.6 1234 ± 126.1 1 ± 0.1 0 ± 0 1 ± 0.2

(551-665) (1797-3020) (436-1491) (726-1619) (0-1) (ND-0.001) (1-2)

NMIN-2 10 573 ± 24.3 2744 ± 469.8 230 ± 26.9 922 ± 166.3 0 ± 0 0 ± 0 3 ± 0

(446-722) (1229-6407) (50-364) (402-1961) (0-1) (ND-0.001) (3-3)

NMIN-3 19 540 ± 11.7 1970 ± 79.6 726 ± 84.6 1821 ± 147.4 0 ± 0.1 0 ± 0 1 ± 0.1

(457-638) (1389-2756) (176-1728) (755-2604) (0-1) (ND-0.83) (1-2)

Table 3A. (Continued) Site and regional mean values for biomarker and ancillary data for COMMON CARP collections made at all sites 

in U.S. waters from March 1994 through August 1997.

[N, number of fish; E2, 17ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; mm, millimeter; g, grams; pg, picograms; mg, milligram; mL, milliliter; 

ND, nondetected]

Table 3A 11
12  Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Regional site 

code

(fig. 1)

N Length

(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental

stage

NMIN-4 8 536 ± 18.1 2275 ± 181 411 ± 26.7 858 ± 261.2 1 ± 0.2 0 ± 0 2 ± 0.2

(450-592) (1300-2800) (310-544) (246-2202) (0-2) (0.001-0.001) (2-3)

NMKS-1 10 573 ± 15.8 2438 ± 172.1 702 ± 108.2 1022 ± 39.1 1 ± 0.1 0 ± 0 2 ± 0.2

(460-621) (1200-2950) (417-1445) (828-1206) (0-2) (ND-0.013) (1-3)

NMMN-1 9 490 ± 6.1 1435 ± 58.9 586 ± 141.5 1174 ± 339.3 1 ± 0.1 0 ± 0 3 ± 0.1

(460-520) (1253-1763) (57-1105) (77-2935) (0-1) (ND-0.09) (2-3)

NMMN-2 10 505 ± 15.4 1604 ± 170 759 ± 118.2 822 ± 258.2 2 ± 0.6 0 ± 0 3 ± 0

(450-620) (998-2918) (267-1573) (105-2261) (0-7) (ND-0.05) (3-3)

NMMN-3 10 504 ± 6.1 1975 ± 298.5 862 ± 78.7 1457 ± 111.5 1 ± 0.1 0 ± 0 1 ± 0.1

(470-536) (1450-4600) (445-1258) (1050-1963) (0-1) (ND-0.008) (1-1.5)

NMMO-1 10 487 ± 14 1590 ± 132.4 429 ± 135.2 772 ± 92.4 1 ± 0.2 0 ± 0 2 ± 0.1

(410-560) (1158-2467) (17-1457) (287-1389) (0-2) (ND-0.001) (2-3)

NMMO-4 9 524 ± 30.4 2078 ± 287.4 915 ± 107.9 1240 ± 302.3 3 ± 2.1 0 ± 0 2 ± 0.2

(346-610) (550-3050) (413-1431) (72-2954) (0-19) (ND-0.001) (1-2.5)

NMMT-1 8 529 ± 15.6 1775 ± 161.8 1096 ± 86.8 2267 ± 245 1 ± 0.1 0 ± 0.1 2 ± 0.2

(470-585) (1250-2600) (716-1418) (1184-2970) (0-1) (ND-0.958) (1-3)

NMMT-2 8 551 ± 26.3 2150 ± 220.6 1209 ± 255.8 2709 ± 371.8 0 ± 0.1 0 ± 0 1 ± 0.2

(390-619) (800-2800) (709-2918) (1191-4214) (0-1) (ND-0.001) (1-2)

NMND-1 10 589 ± 16.6 2352 ± 203 444 ± 95 965 ± 129 1 ± 0.1 1 ± 0.5 3 ± 0.2

(520-675) (1645-3470) (100-901) (508-1947) (0-1) (ND-5) (2-3)

NMND-2 4 471 ± 15.3 1252 ± 83.4 1431 ± 348.4 857 ± 238.1 2 ± 0.8 0 ± 0 3 ± 0.3

(436-510) (1079-1470) (465-2104) (387-1507) (1-4) (ND-0.07) (2-3)

NMND-3 11 489 ± 23.1 1625 ± 226.6 300 ± 13 2249 ± 513.2 0 ± 0 0 ± 0 1 ± 0.2

(299-595) (420-3400) (219-362) (887-6760) (0-0) (ND-0.007) (0-2)

NMNE-1 20 509 ± 11.1 1752 ± 88.3 308 ± 54.3 1432 ± 327.9 0 ± 0.1 1 ± 0.5 1 ± 0.1

(390-615) (838-2350) (39-881) (111-5303) (0-2) (ND-10.981) (1-2)

NMNE-2 18 462 ± 20 1331 ± 137 294 ± 83.5 600 ± 123.1 1 ± 0.2 0 ± 0 2 ± 0.3

(317-586) (464-2456) (17-1171) (76-2007) (0-2) (ND-0.01) (0-3)

NMNE-3 11 472 ± 34.5 1655 ± 331 363 ± 18.8 2205 ± 218.5 0 ± 0 0 ± 0 1 ± 0.1

(352-680) (600-3450) (283-482) (1142-3136) (0-0) (ND-0.001) (1-2)

NMSD-1 9 494 ± 32.1 1692 ± 262.1 821 ± 94.6 1504 ± 301.3 1 ± 0.3 0 ± 0 1 ± 0.2

(319-603) (325-2775) (446-1285) (256-3062) (0-3) (ND-0.008) (0.5-2)

NMWI-1 10 433 ± 8 980 ± 62.2 656 ± 51 707 ± 100 1 ± 0.2 0 ± 0 2 ± 0.3

(405-490) (712-1430) (365-859) (253-1306) (0-3) (ND-0.02) (1-3)

NMWI-2 23 486 ± 8.9 1384 ± 74.6 775 ± 65.3 621 ± 55.5 1 ± 0.2 1 ± 0.7 3 ± 0.1

(414-560) (868-2154) (214-1340) (108-1118) (1-4) (ND-15) (2-3)

NMWI-3 12 519 ± 8.3 1817 ± 81.7 582 ± 109.2 1661 ± 249.4 1 ± 0.6 0 ± 0 1 ± 0.1

(468-578) (1400-2400) (145-1474) (191-2893) (0-8) (ND-0.005) (1-2)

Summary 384 505 ± 3.7 1752 ± 38.1 531 ± 21.1 1288 ± 54.9 1 ± 0.1 0 ± 0.1 2 ± 0

(299-722) (324-6407) (17-2918) (69-6796) (0-19) (ND-25.427) (0-3)

Table 3A. (Continued) Site and regional mean values for biomarker and ancillary data for COMMON CARP collections made at all sites 

in U.S. waters from March 1994 through August 1997.

[N, number of fish; E2, 17ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; mm, millimeter; g, grams; pg, picograms; mg, milligram; mL, milliliter; 

ND, nondetected]
Regional site 

code

(fig. 1)

N Length

(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental

stage

 Male reproductive

NMIN-1 5 583 ± 4.9 2973 ± 95.8 173 ± 38.3 690 ± 148.2 0 ± 0 0 ± 0 3 ± 0

(572-599) (2675-3235) (101-308) (376-1092) (0-0) (ND-0.001) (3-3)

NMIN-3 19 539 ± 13.9 2343 ± 158.5 278 ± 21.4 573 ± 77.5 1 ± 0.1 1 ± 0.6 3 ± 0.1

(458-653) (1340-3722) (108-441) (151-1267) (0-2) (ND-11.956) (1-3)

Summary 24 548 ± 11.6 2475 ± 136.9 256 ± 20.3 597 ± 67.9 1 ± 0.1 1 ± 0.5 3 ± 0.1

(458-653) (1340-3722) (101-441) (151-1267) (0-2) (ND-11.956) (1-3)

Northeast Region

 Female non-reproductive

NECT-1 9 729 ± 19.8 6613 ± 581.2 3343 ± 644 1568 ± 191.3 2 ± 0.3 29 ± 4.2 3 ± 0.2

(618-810) (2423-8286) (1646-7703) (769-2740) (1-5) (ND-38.3) (3-4)

NEDC-1 10 591 ± 22.8 3493 ± 373.2 787 ± 124 552 ± 129 3 ± 1.2 24 ± 3 3 ± 0.1

(496-692) (2032-5477) (323-1765) (66-1125) (1-12) (6.64-39.6) (3-4)

NENY-1 7 556 ± 32 3023 ± 591.5 2694 ± 659.6 336 ± 101 11 ± 3 20 ± 2.5 3 ± 0

(487-727) (1569-6200) (1002-5780) (113-920) (4-22) (8-29) (3-3)

NENY-2 16 561 ± 9.3 2774 ± 129 1746 ± 342.1 586 ± 101.1 5 ± 1.7 14 ± 1.4 3 ± 0

(486-614) (1813-3516) (562-4741) (115-1508) (1-22) (6.4-26.6) (3-3)

NENY-4 11 635 ± 35.3 4104 ± 749.7 3329 ± 388.8 1102 ± 146.4 3 ± 0.5 35 ± 2.5 3 ± 0

(513-920) (2175-10000) (435-4499) (304-1702) (1-8) (19-50.2) (3-3)

NEOH-1 2 623 ± 87.5 3225 ± 1425 424 ± 145.5 137 ± 13.5 3 ± 0.8 3 ± 1 3 ± 0.5

(535-710) (1800-4650) (278-569) (123-150) (2-4) (2.18-4.23) (2-3)

NEPA-1 6 519 ± 35.9 1975 ± 463.3 1166 ± 452 467 ± 221.7 3 ± 0.7 2 ± 0.6 3 ± 0

(429-660) (1000-3900) (259-2768) (88-1504) (1-6) (0.314-4.671) (3-3)

NEWV-1 14 629 ± 14.1 3740 ± 220 1159 ± 125.7 278 ± 53.1 9 ± 2.5 14 ± 1.5 3 ± 0.1

(518-726) (1834-5335) (315-1760) (13-700) (1-28) (ND-20.4) (3-4)

NEWV-2 11 406 ± 8.7 875 ± 54.9 716 ± 92.8 262 ± 33.3 3 ± 0.4 1 ± 0.3 3 ± 0

(362-442) (584.4-1204.8) (309-1231) (103-481) (1-6) (ND-2.887) (3-3)

Summary 86 581 ± 11.8 3319 ± 211.2 1783 ± 162.1 620 ± 60.1 5 ± 0.6 17 ± 1.4 3 ± 0

(362-920) (584.4-10000) (259-7703) (13-2740) (1-28) (ND-50.2) (2-4)

 Female reproductive

NENY-1 3 575 ± 23.9 2990 ± 482.1 683 ± 147.3 799 ± 331.1 1 ± 0.6 57 ± 15.5 3 ± 0

(529-610) (2070-3700) (433-943) (167-1286) (1-3) (40.36-87.793) (3-3)

NENY-2 12 585 ± 18.1 3392 ± 279.5 2753 ± 297.6 3333 ± 307.1 1 ± 0.1 37 ± 5.3 3 ± 0

(455-689) (2100-5300) (1159-4142) (1456-5066) (0-2) (13.348-65.612) (3-3)

NENY-4 13 658 ± 14 4715 ± 346 1722 ± 234.2 1004 ± 195 3 ± 0.8 44 ± 6.1 3 ± 0

(576-735) (2800-6300) (467-3662) (199-2554) (1-12) (ND-89.456) (3-3)

Summary 28 618 ± 12.5 3963 ± 242.6 2053 ± 209.9 1980 ± 276 2 ± 0.4 42 ± 4 3 ± 0

(455-735) (2070-6300) (433-4142) (167-5066) (0-12) (ND-89.456) (3-3)

Table 3A. (Continued) Site and regional mean values for biomarker and ancillary data for COMMON CARP collections made at all sites 

in U.S. waters from March 1994 through August 1997.

[N, number of fish; E2, 17ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; mm, millimeter; g, grams; pg, picograms; mg, milligram; mL, milliliter; 

ND, nondetected]

Table 3A 13
14  Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Regional site 

code

(fig. 1)

N Length

(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental

stage

 Male non-reproductive

NECT-1 11 569 ± 37.9 3268 ± 632.8 744 ± 102.1 1349 ± 175.7 1 ± 0.1 0 ± 0 3 ± 0.1

(339-690) (601-6356) (114-1227) (442-2161) (0-1) (ND-0.001) (2-3)

NEDC-1 22 508 ± 10.3 2068 ± 123.8 391 ± 24.8 911 ± 113.3 1 ± 0.1 0 ± 0 1 ± 0.1

(401-615) (1073-3545) (204-601) (341-2038) (0-1) (ND-0.001) (1-3)

NENY-1 19 557 ± 14.1 2628 ± 250.9 506 ± 80.6 857 ± 91.8 1 ± 0.1 0 ± 0 3 ± 0.1

(475-674) (1356-5500) (72-1183) (254-1638) (0-1) (ND-0.57) (2-3)

NENY-2 24 539 ± 5 2212 ± 66.7 190 ± 29.6 837 ± 113.8 0 ± 0.1 0 ± 0 2 ± 0.2

(466-576) (1582-3037) (30-509) (144-2186) (0-1) (ND-0.18) (1-3)

NENY-4 18 607 ± 10.9 3461 ± 201 216 ± 45.5 1762 ± 142.5 0 ± 0 0 ± 0 3 ± 0

(508-673) (1946-5352) (29-857) (565-3012) (0-0) (ND-0.04) (3-3)

NEOH-1 3 523 ± 22.5 1624 ± 246.8 458 ± 47.4 1760 ± 1620.8 2 ± 1.2 0 ± 0 1 ± 0

(479-552) (1150-1981) (401-552) (97-5001) (0-4) (0.001-0.013) (1-1)

NEPA-1 5 472 ± 39.3 1870 ± 223.4 361 ± 23.6 2356 ± 564.7 0 ± 0 0 ± 0 2 ± 0.2

(373-594) (1500-2750) (300-407) (1319-4090) (0-0) (ND-0.027) (2-3)

NEWV-1 22 577 ± 6.7 2802 ± 111.5 260 ± 39.2 421 ± 62.9 1 ± 0.1 0 ± 0 1 ± 0.1

(507-637) (1832-3764) (50-872) (75-1187) (0-2) (ND-0.27) (1-2)

NEWV-2 8 389 ± 16.9 753 ± 97 301 ± 37.9 971 ± 196.5 0 ± 0.1 0 ± 0 3 ± 0

(337-464) (462.8-1129.5) (135-419) (371-1764) (0-1) (ND-0.001) (2.5-2.5)

Summary 132 543 ± 6.6 2490 ± 94.8 350 ± 22.8 1038 ± 69.2 1 ± 0 0 ± 0 2 ± 0.1

(337-690) (462.8-6356) (29-1227) (75-5001) (0-4) (ND-0.57) (1-3)

 Male reproductive

NENY-1 17 568 ± 17.7 2856 ± 306.5 827 ± 72 800 ± 102.8 1 ± 0.2 0 ± 0 3 ± 0.2

(473-702) (1375-5150) (427-1786) (222-1811) (0-4) (ND-0.001) (1-3)

NENY-2 17 555 ± 8.3 2371 ± 113.7 435 ± 40.4 3480 ± 388.8 0 ± 0 0 ± 0 3 ± 0.1

(445-595) (1200-3200) (83-686) (1409-6821) (0-0) (ND-0.001) (2-3)

NENY-4 13 648 ± 20.9 4585 ± 430.1 911 ± 219.5 1679 ± 349.6 1 ± 0.4 0 ± 0 2 ± 0.2

(496-763) (2500-7300) (89-2565) (205-4871) (0-6) (ND-0.202) (1-3)

Summary 47 585 ± 10.6 3158 ± 211.2 709 ± 72.7 2012 ± 241.5 1 ± 0.2 0 ± 0 3 ± 0.1

(445-763) (1200-7300) (83-2565) (205-6821) (0-6) (ND-0.202) (1-3)

Northwest Region

 Female non-reproductive

NWID-1 10 399 ± 12.1 892 ± 80.8 1094 ± 160.3 289 ± 52.4 5 ± 1.1 1 ± 0.3 1 ± 0.2

(355-464) (603-1289) (483-1990) (89-612) (2-12) (0.009-2.805) (0-2)

NWID-2 4 617 ± 11.8 3550 ± 266.1 1201 ± 48.7 535 ± 155.1 3 ± 1.1 4 ± 1.3 2 ± 0

(585-639) (2900-4200) (1115-1341) (186-844) (2-6) (1.437-7.368) (2-2)

NWOR-2 14 404 ± 3.7 843 ± 33.1 538 ± 122.8 386 ± 78.5 2 ± 0.2 9 ± 2.9 2 ± 0.2

(385-437) (681-1022) (67-1522) (22-1030) (1-3) (0.31-36.9) (1-3)

NWOR-3 10 573 ± 19.5 2724 ± 307.3 632 ± 83 431 ± 96.1 2 ± 0.6 0 ± 0.1 2 ± 0.1

(500-696) (1670-4500) (88-1031) (58-898) (1-6) (0-0.997) (2-3)

Table 3A. (Continued) Site and regional mean values for biomarker and ancillary data for COMMON CARP collections made at all sites 

in U.S. waters from March 1994 through August 1997.

[N, number of fish; E2, 17ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; mm, millimeter; g, grams; pg, picograms; mg, milligram; mL, milliliter; 

ND, nondetected]
Regional site 

code

(fig. 1)

N Length

(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental

stage

NWOR-4 4 677 ± 53.4 5313 ± 1252.1 578 ± 164 196 ± 58.9 3 ± 0.9 0 ± 0.2 2 ± 0

(562-771) (2650-8000) (183-899) (82-349) (1-5) (0.067-0.994) (2-2)

NWWA-3 11 518 ± 19.4 2041 ± 257.7 1241 ± 139.5 270 ± 44.9 6 ± 0.8 2 ± 0.2 2 ± 0

(439-626) (1105-3750) (512-1833) (53-613) (1-10) (1.355-2.986) (2-2)

NWWA-4 10 607 ± 19.3 3178 ± 303.8 512 ± 86.4 262 ± 51.6 3 ± 0.6 1 ± 0.2 2 ± 0.1

(475-661) (1376-4550) (102-855) (51-548) (1-7) (0.126-2.327) (1-2)

NWWA-5 10 654 ± 23.3 4219 ± 551.1 958 ± 108.3 260 ± 41.2 5 ± 1 2 ± 0.2 2 ± 0

(535-745) (2150-6738) (518-1462) (79-447) (1-10) (1.457-3.109) (2-2)

NWWA-6 10 520 ± 18.7 1818 ± 213.8 1063 ± 96.1 361 ± 67.5 4 ± 1 2 ± 0.2 2 ± 0.1

(428-615) (898-3150) (587-1418) (100-770) (2-12) (0.537-2.885) (1-2)

Summary 83 531 ± 12 2386 ± 178.5 854 ± 50.5 329 ± 24.6 4 ± 0.3 3 ± 0.6 2 ± 0.1

(355-771) (603-8000) (67-1990) (22-1030) (1-12) (0.009-36.9) (0-3)

 Female reproductive

NWOR-1 15 504 ± 9.4 1959 ± 129.4 823 ± 127.9 592 ± 87.8 2 ± 0.4 41 ± 8 3 ± 0.2

(428-565) (1052-2983) (249-2069) (168-1124) (0-5) (2.66-118.839) (1-3)

NWOR-2 15 402 ± 7 939 ± 48.3 443 ± 89.8 458 ± 48 1 ± 0.5 25 ± 3.2 2 ± 0.2

(345-445) (550-1216) (17-1105) (144-744) (0-6) (0.001-40.04) (2-4)

NWWA-1 10 505 ± 9.7 1945 ± 112.5 1289 ± 115.9 501 ± 77.3 3 ± 0.8 20 ± 5.1 3 ± 0

(453-542) (1454-2451) (464-1875) (45-934) (2-10) (4.596-62.448) (3-3)

NWWA-2 15 650 ± 14.3 4190 ± 237 1517 ± 153.7 741 ± 68 2 ± 0.1 44 ± 6.9 3 ± 0.1

(530-710) (2192-5320) (1005-3542) (476-1507) (2-3) (16.589-86.521) (3-4)

Summary 55 516 ± 13.6 2287 ± 184.7 993 ± 84.6 580 ± 38 2 ± 0.2 34 ± 3.4 3 ± 0.1

(345-710) (550-5320) (17-3542) (45-1507) (0-10) (0.001-118.839) (1-4)

 Male non-reproductive

NWID-1 10 364 ± 8.7 683 ± 54.1 304 ± 56.2 777 ± 176.9 1 ± 0.2 0 ± 0 2 ± 0.2

(321-406) (397-900) (135-575) (128-1503) (0-2) (0-0) (2-3)

NWID-2 9 540 ± 18.2 2300 ± 207.5 452 ± 75.1 778 ± 158.9 1 ± 0.3 0 ± 0 2 ± 0.2

(440-601) (1300-3300) (171-792) (260-1638) (0-3) (0-0) (2-3)

NWOR-2 15 385 ± 8.1 742 ± 57.4 169 ± 53 730 ± 107.3 0 ± 0.1 0 ± 0 2 ± 0.1

(339-482) (454-1476) (23-875) (84-1738) (0-2) (ND-0.68) (2-3)

NWOR-3 10 514 ± 20.3 1961 ± 221 384 ± 103.1 440 ± 103.6 1 ± 0.3 0 ± 0 3 ± 0

(383-625) (819-3400) (118-1117) (126-1156) (0-4) (0-0.035) (3-3)

NWOR-4 10 563 ± 21.3 2586 ± 287.6 374 ± 84.2 316 ± 36.7 1 ± 0.3 0 ± 0 3 ± 0

(465-677) (1351-3800) (63-882) (147-556) (0-3) (0-0.01) (3-3)

NWWA-3 9 501 ± 12.2 1473 ± 104.4 316 ± 70.8 663 ± 103 1 ± 0.1 0 ± 0 2 ± 0.2

(461-581) (1200-2200) (109-798) (195-1139) (0-1) (0-0) (2-3)

NWWA-4 11 523 ± 25.4 1853 ± 224.2 237 ± 34.1 423 ± 64.1 1 ± 0.2 0 ± 0 2 ± 0.2

(416-667) (917-2950) (102-432) (103-703) (0-2) (0-0.037) (2-3)

Table 3A. (Continued) Site and regional mean values for biomarker and ancillary data for COMMON CARP collections made at all sites 

in U.S. waters from March 1994 through August 1997.

[N, number of fish; E2, 17ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; mm, millimeter; g, grams; pg, picograms; mg, milligram; mL, milliliter; 

ND, nondetected]

Table 3A 15
16  Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Regional site 

code

(fig. 1)

N Length

(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental

stage

NWWA-5 11 595 ± 11.6 2891 ± 174.6 179 ± 32.7 582 ± 125.4 0 ± 0.1 0 ± 0 2 ± 0.2

(560-667) (2200-4000) (72-463) (185-1648) (0-1) (0-0.009) (2-3)

NWWA-6 10 489 ± 13.6 1437 ± 139.8 323 ± 77.2 651 ± 104 1 ± 0.2 0 ± 0 3 ± 0.1

(444-590) (1038-2500) (76-783) (123-1124) (0-2) (0-0) (2-3)

Summary 95 492 ± 9.3 1726 ± 94.7 293 ± 23.2 598 ± 40.1 1 ± 0.1 0 ± 0 3 ± 0.1

(321-677) (397-4000) (23-1117) (84-1738) (0-4) (0-0.68) (2-3)

 Male reproductive

NWOR-1 15 461 ± 9.2 1342 ± 75.9 561 ± 49.8 1473 ± 273.9 1 ± 0.1 0 ± 0 3 ± 0.1

(379-521) (690-1996) (169-818) (509-4310) (0-1) (ND-0.001) (2-3)

NWOR-2 15 369 ± 7.1 726 ± 35.2 263 ± 13.9 509 ± 59.3 1 ± 0.1 0 ± 0 3 ± 0.2

(327-446) (547-1096) (121-341) (149-821) (0-2) (ND-0.09) (1-3)

NWWA-1 10 511 ± 12.2 1713 ± 152 515 ± 101.2 1136 ± 139.1 0 ± 0.1 0 ± 0 2 ± 0.3

(464-571) (1056-2688) (201-1179) (549-2020) (0-1) (ND-0.001) (1-3)

NWWA-2 15 624 ± 7.5 3404 ± 118.1 554 ± 78.9 1038 ± 129.1 1 ± 0 0 ± 0 3 ± 0

(575-683) (2750-4012) (226-1129) (393-2181) (0-1) (ND-0.001) (3-3)

Summary 55 489 ± 13.7 1804 ± 148.6 469 ± 35.2 1030 ± 98.6 1 ± 0 0 ± 0 3 ± 0.1

(327-683) (547-4012) (121-1179) (149-4310) (0-2) (ND-0.09) (1-3)

Southern Mid-Continent Region

 Female non-reproductive 

SMAR-1 7 471 ± 28 1582 ± 236.7 2021 ± 554.3 194 ± 20.5 11 ± 2.9 4 ± 0.5 3 ± 0.1

(340-550) (523-2278) (821-5126) (99-268) (3-21) (2.155-5.763) (2-3)

SMAR-2 10 516 ± 12.6 2022 ± 187 1465 ± 214.9 615 ± 155 3 ± 0.9 4 ± 0.5 3 ± 0.2

(465-600) (1358-3472) (151-2192) (203-1956) (0-11) (0.421-5.804) (1-3)

SMAR-3 7 599 ± 21.3 3241 ± 337.6 1105 ± 255.4 565 ± 233.7 3 ± 0.8 1 ± 0.6 3 ± 0

(540-700) (2052-4690) (439-2265) (172-1925) (0-7) (ND-4.28) (3-3)

SMAR-4 9 525 ± 12.2 2026 ± 150.7 1755 ± 445.3 376 ± 65.7 4 ± 0.8 3 ± 0.5 3 ± 0

(473-570) (1480-2792) (147-4537) (166-779) (1-9) (1.122-5.995) (3-3)

SMIL-2 5 640 ± 36.9 3304 ± 744.7 462 ± 77.9 409 ± 152.9 3 ± 1.2 2 ± 0.4 3 ± 0

(510-736) (1700-5720) (349-769) (108-779) (0-7) (0.845-3.24) (3-3)

SMLA-2 7 577 ± 47.3 3790 ± 784.5 597 ± 68.9 290 ± 76.2 2 ± 0.3 2 ± 0.4 2 ± 0.4

(350-720) (720-6602) (288-907) (131-716) (1-4) (0.567-3.244) (1-3)

SMLA-3 8 531 ± 38.1 2129 ± 469.1 1211 ± 125.3 335 ± 24 4 ± 0.5 2 ± 0.3 3 ± 0.1

(435-750) (1097-5000) (591-1782) (260-429) (1-7) (0.245-2.897) (2-3)

SMMO-2 10 375 ± 7.7 663 ± 59.8 491 ± 94.8 226 ± 34.8 3 ± 0.6 2 ± 0.5 2 ± 0.3

(344-415) (304-990) (181-984) (123-461) (1-7) (0.006-3.998) (1-3)

SMMS-3 1 495 ± 0 1638 ± 0 1149 ± 0 305 ± 0 4 ± 0 0 ± 0 3 ± 0

(495-495) (1638-1638) (1149-1149) (305-305) (4-4) (0.14-0.14) (3-3)

SMOK-1 11 534 ± 17.7 2003 ± 282.4 1426 ± 218.6 419 ± 63.7 4 ± 0.6 1 ± 0.4 2 ± 0.4

(473-670) (1250-4350) (271-2489) (196-943) (0-6) (0.001-3.717) (0-3)

Table 3A. (Continued) Site and regional mean values for biomarker and ancillary data for COMMON CARP collections made at all sites 

in U.S. waters from March 1994 through August 1997.

[N, number of fish; E2, 17ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; mm, millimeter; g, grams; pg, picograms; mg, milligram; mL, milliliter; 

ND, nondetected]
Regional site 

code

(fig. 1)

N Length

(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental

stage

SMOK-2 10 490 ± 16.6 1603 ± 143.1 1453 ± 151.8 891 ± 86.4 2 ± 0.4 1 ± 0.3 3 ± 0.3

(420-580) (1000-2550) (741-2466) (464-1337) (1-5) (0.044-2.613) (0-3)

SMOK-3 10 540 ± 20.5 2150 ± 249.4 1622 ± 116.7 960 ± 143.8 2 ± 0.6 2 ± 0.2 3 ± 0

(470-656) (1400-3750) (1221-2303) (199-1746) (1-7) (0.712-3.109) (3-3)

SMTN-2 9 451 ± 15.5 1454 ± 165.2 444 ± 109.6 226 ± 36.6 2 ± 0.8 2 ± 0.7 2 ± 0.3

(379-504) (799-2200) (186-1008) (118-448) (0-7) (0.01-4.859) (0-3)

SMTN-4 2 545 ± 0 2184 ± 216 550 ± 4.5 94 ± 25 5 ± 0.7 3 ± 0.4 3 ± 0

(545-545) (1968-2400) (545-554) (69-119) (5-6) (2.304-3.113) (3-3)

SMTN-5 5 543 ± 38.7 2420 ± 469.5 1470 ± 327 712 ± 157 2 ± 0.4 0 ± 0.2 3 ± 0.2

(455-662) (1200-3600) (583-2582) (370-1249) (1-3) (0.001-1.24) (2-3)

SMTX-3 5 534 ± 17.7 2110 ± 152.8 382 ± 77.4 119 ± 31.2 3 ± 0.6 12 ± 1.8 3 ± 0

(472-575) (1589-2423) (110-558) (57-237) (2-5) (7-16.5) (3-3)

SMTX-4 5 626 ± 23.1 4106 ± 597.3 1708 ± 367.3 445 ± 63 4 ± 0.4 10 ± 1.2 3 ± 0

(559-690) (2690-5992) (947-2922) (265-654) (2-4) (7-12.8) (3-3)

SMTX-7 13 635 ± 15.4 3292 ± 239.8 733 ± 48.6 246 ± 86.1 8 ± 1.9 1 ± 0.2 3 ± 0

(535-730) (1800-4600) (538-1080) (32-1136) (1-26) (0.265-3.142) (3-3)

Summary 134 531 ± 8 2268 ± 107.3 1145 ± 69.4 441 ± 32.5 4 ± 0.3 3 ± 0.3 3 ± 0.1

(340-750) (304-6602) (110-5126) (32-1956) (0-26) (ND-16.5) (0-3)

 Female reproductive

SMLA-1 5 479 ± 29 1588 ± 286.5 977 ± 317.1 441 ± 195.7 3 ± 0.8 3 ± 1.1 3 ± 0

(404-554) (868-2380) (166-2122) (154-1209) (1-5) (0.202-5.641) (3-3)

SMMS-1 8 478 ± 21.6 1411 ± 189.2 1125 ± 207.7 299 ± 41.8 4 ± 1 0 ± 0 3 ± 0.3

(413-558) (907-2432) (23-1634) (165-479) (0-9) (ND-0.228) (1-3)

SMMS-2 10 445 ± 19.5 1157 ± 160.3 1676 ± 215.9 539 ± 108.2 5 ± 2.2 1 ± 0.3 3 ± 0.2

(356-528) (613-2316) (236-2318) (88-1114) (1-25) (0.127-2.238) (1-3)

SMMS-3 7 392 ± 16 843 ± 90.7 1534 ± 269.1 260 ± 23.2 6 ± 1.3 1 ± 0.4 3 ± 0

(332-456) (479-1175) (338-2317) (169-339) (2-12) (0.006-2.877) (3-3)

SMMS-4 5 609 ± 42.2 3605 ± 699.2 834 ± 197.1 635 ± 155.5 1 ± 0.5 0 ± 0.3 3 ± 0.2

(497-742) (2071-6200) (356-1391) (427-1246) (1-3) (ND-1.473) (2-3)

SMTX-2 16 499 ± 18.6 1681 ± 228.4 1158 ± 164.4 477 ± 79.7 3 ± 0.5 4 ± 2.9 2 ± 0.1

(420-710) (960-4521) (119-2397) (112-1291) (1-9) (ND-46.029) (2-3)

Summary 51 479 ± 12 1599 ± 147.8 1256 ± 94.6 443 ± 42.7 4 ± 0.5 2 ± 0.9 3 ± 0.1

(332-742) (479-6200) (23-2397) (88-1291) (0-25) (ND-46.029) (1-3)

 Male non-reproductive

SMAR-1 9 512 ± 15.1 1809 ± 146.5 626 ± 172 217 ± 19.3 3 ± 0.8 0 ± 0 2 ± 0

(460-570) (1189-2505) (61-1472) (158-317) (0-7) (ND-0.001) (2-2)

SMAR-2 10 465 ± 13.5 1317 ± 100.3 410 ± 97.9 990 ± 190.2 1 ± 0.3 0 ± 0 2 ± 0

(391-519) (882-1758) (83-946) (109-1815) (0-4) (ND-0.001) (2-2)

Table 3A. (Continued) Site and regional mean values for biomarker and ancillary data for COMMON CARP collections made at all sites 

in U.S. waters from March 1994 through August 1997.

[N, number of fish; E2, 17ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; mm, millimeter; g, grams; pg, picograms; mg, milligram; mL, milliliter; 

ND, nondetected]

Table 3A 17
18  Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Regional site 

code

(fig. 1)

N Length

(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental

stage

SMAR-3 10 566 ± 23.5 2773 ± 382.2 535 ± 55.3 1265 ± 317 1 ± 0.2 0 ± 0 2 ± 0.2

(430-674) (1147-4996) (358-945) (248-3226) (0-2) (ND-0.001) (1-3)

SMAR-4 10 545 ± 30.9 2016 ± 302.9 496 ± 116.1 3531 ± 924.1 0 ± 0.1 0 ± 0 1 ± 0.1

(330-700) (643-4200) (80-1228) (632-8492) (0-1) (0.001-0.014) (1-1.5)

SMIL-2 6 578 ± 14.8 2550 ± 282.3 300 ± 47.7 2564 ± 1379.1 1 ± 0.2 0 ± 0 1 ± 0.2

(515-620) (1500-3600) (170-469) (271-7985) (0-1) (0.001-0.001) (1-2)

SMLA-2 1 600 ± 0 3325 ± 0 491 ± 0 590 ± 0 1 ± 0 0 ± 0 2 ± 0

(600-600) (3325-3325) (491-491) (590-590) (1-1) (0.001-0.001) (2-2)

SMLA-3 4 554 ± 45.3 2192 ± 428.8 584 ± 63.5 870 ± 215 1 ± 0.2 0 ± 0 2 ± 0.3

(473-651) (1410-2950) (459-746) (493-1296) (0-1) (0.001-0.001) (2-3)

SMMO-2 10 401 ± 22.7 918 ± 167.6 369 ± 31 1767 ± 594.9 0 ± 0.1 0 ± 0 2 ± 0.2

(300-550) (336-2009) (259-527) (220-5991) (0-1) (0.001-0.001) (1-2)

SMOK-1 9 494 ± 14.6 1578 ± 134.7 339 ± 21.4 865 ± 176.1 0 ± 0.1 0 ± 0 3 ± 0.3

(432-575) (1000-2500) (246-461) (359-2016) (0-1) (0.001-0.001) (1-3)

SMOK-2 10 435 ± 10.1 1153 ± 73.3 1228 ± 146.2 1068 ± 152.1 1 ± 0.2 0 ± 0 2 ± 0.1

(356-470) (900-1700) (464-1957) (577-2348) (0-3) (0.001-0.001) (2-3)

SMOK-3 9 485 ± 11.1 1500 ± 117.2 775 ± 98.5 1536 ± 191.7 1 ± 0.1 0 ± 0 3 ± 0

(433-530) (1000-2175) (300-1148) (652-2116) (0-1) (0.001-0.009) (3-3)

SMTN-2 8 445 ± 14.5 1292 ± 115.5 413 ± 85.7 499 ± 159.3 1 ± 0.2 0 ± 0 2 ± 0.1

(370-498) (711-1778) (205-828) (167-1565) (0-2) (0.001-0.005) (1-2)

SMTN-4 2 538 ± 32.5 2065 ± 335.5 246 ± 85.5 651 ± 513.5 1 ± 1.1 0 ± 0 2 ± 0

(505-570) (1729-2400) (160-331) (137-1164) (0-2) (0.001-0.025) (2-2)

SMTN-5 10 505 ± 16.7 1745 ± 177.4 444 ± 99.8 1406 ± 284.4 1 ± 0.5 0 ± 0 2 ± 0.1

(395-570) (850-2500) (138-1276) (264-2783) (0-5) (0.001-0.001) (1.5-3)

SMTX-3 2 518 ± 7 1749 ± 33 78 ± 0.5 85 ± 11 1 ± 0.1 0 ± 0 1 ± 0

(511-525) (1716-1782) (77-78) (74-96) (1-1) (ND-0.001) (1-1)

SMTX-4 4 590 ± 22.6 2969 ± 107.4 303 ± 60 616 ± 47.8 0 ± 0.1 0 ± 0 2 ± 0.3

(550-655) (2718-3155) (128-388) (521-732) (0-1) (ND-0.001) (1-2)

SMTX-5 1 490 ± 0 1550 ± 0 636 ± 0 1415 ± 0 0 ± 0 0 ± 0 2 ± 0

(490-490) (1550-1550) (636-636) (1415-1415) (0-0) (0.008-0.008) (2-2)

SMTX-7 10 585 ± 15.4 2455 ± 202 632 ± 54.2 1046 ± 350.5 4 ± 1.5 0 ± 0 2 ± 0.1

(521-670) (1700-3700) (403-1037) (61-2983) (0-13) (0.001-0.093) (2-3)

Summary 125 506 ± 7 1816 ± 74.6 538 ± 32.7 1305 ± 137.9 1 ± 0.2 0 ± 0 2 ± 0.1

(300-700) (336-4996) (61-1957) (61-8492) (0-13) (ND-0.093) (1-3)

 Male reproductive

SMLA-1 9 518 ± 11.6 1730 ± 78.6 362 ± 88.7 243 ± 25.2 2 ± 0.7 0 ± 0 2 ± 0.1

(445-555) (1298-2065) (71-792) (111-355) (0-7) (ND-0.001) (1-2)

SMMS-1 8 447 ± 20.6 1036 ± 176.2 286 ± 77.4 438 ± 71.1 1 ± 0.2 0 ± 0 1 ± 0

(368-548) (325-1829) (78-786) (201-738) (0-1) (ND-0.178) (1-1)

Table 3A. (Continued) Site and regional mean values for biomarker and ancillary data for COMMON CARP collections made at all sites 

in U.S. waters from March 1994 through August 1997.

[N, number of fish; E2, 17ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; mm, millimeter; g, grams; pg, picograms; mg, milligram; mL, milliliter; 

ND, nondetected]
Regional site 

code

(fig. 1)

N Length

(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental

stage

SMMS-2 10 444 ± 16 1119 ± 126.5 197 ± 69.3 688 ± 73.9 0 ± 0.1 0 ± 0 2 ± 0.1

(383-546) (685-1977) (51-757) (372-1093) (0-1) (ND-0.001) (1-2)

SMMS-3 10 401 ± 10.6 824 ± 75 176 ± 46.1 553 ± 81.5 0 ± 0.1 0 ± 0 2 ± 0.1

(362-460) (649-1347) (62-539) (158-961) (0-1) (ND-0.001) (1-2)

SMMS-4 7 527 ± 41.8 2289 ± 367.1 526 ± 96 465 ± 64.3 1 ± 0.4 0 ± 0 2 ± 0.1

(297-657) (407-3693) (120-924) (199-721) (0-3) (ND-0.001) (1-2)

SMTX-2 9 434 ± 9.4 1066 ± 71.3 594 ± 139.6 1029 ± 204.9 1 ± 0.5 0 ± 0 2 ± 0.1

(389-474) (661-1400) (158-1442) (158-1945) (0-5) (ND-0.001) (2-3)

Summary 53 459 ± 9.7 1309 ± 91 345 ± 41 578 ± 53.3 1 ± 0.2 0 ± 0 2 ± 0.1

(297-657) (325-3693) (51-1442) (111-1945) (0-7) (ND-0.178) (1-3)

Southwest Region

 Female non-reproductive

SWCA-1 22 551 ± 22 2413 ± 265.6 2669 ± 334.1 1225 ± 193 3 ± 0.4 32 ± 4.1 3 ± 0.1

(410-766) (943-5176) (614-6398) (215-2868) (1-11) (2.3-108.266) (1-3)

SWCA-2 36 514 ± 18.1 2124 ± 244.8 1601 ± 257.3 1779 ± 403.2 2 ± 0.3 23 ± 2 3 ± 0.1

(323-742) (460-6356) (129-8333) (17-9066) (0-9) (1.04-42.6) (1-4)

SWCA-3 36 584 ± 17.7 3255 ± 296.2 2328 ± 376.7 1278 ± 232 3 ± 0.5 32 ± 2.8 3 ± 0.1

(411-795) (913-7290) (135-11401) (121-5973) (0-18) (0.001-75.235) (1-4)

SWUT-1 5 611 ± 41.4 3154 ± 611.8 1256 ± 240.4 464 ± 48.9 3 ± 0.6 4 ± 0.5 3 ± 0.2

(495-700) (1540-4572) (791-2176) (300-586) (2-5) (2.774-5.757) (2-3)

SWUT-2 5 415 ± 22.5 1063 ± 147.3 784 ± 152.2 417 ± 62.1 2 ± 0.4 2 ± 1.2 2 ± 0.2

(366-492) (702-1572) (366-1296) (268-579) (1-3) (0.025-6.396) (2-3)

SWUT-3 5 723 ± 29 5262 ± 708.2 939 ± 278.9 564 ± 111.2 2 ± 0.4 1 ± 0.3 3 ± 0

(625-785) (3397-7581) (316-1816) (311-838) (1-3) (0.581-2.198) (3-3)

SWUT-4 6 438 ± 52.3 1427 ± 440.5 654 ± 154 231 ± 48.9 4 ± 1 1 ± 0.5 2 ± 0.4

(278-560) (288-2869) (278-1207) (34-364) (1-8) (0.042-3.172) (1-3)

SWUT-5 4 604 ± 46.4 3082 ± 861 759 ± 166.8 400 ± 71 2 ± 0.1 2 ± 0.6 3 ± 0.3

(498-718) (1520-5500) (575-1259) (273-603) (2-2) (0.275-3.125) (2-3)

Summary 119 549 ± 10.8 2642 ± 153.9 1866 ± 161.9 1237 ± 150.6 2 ± 0.2 23 ± 1.7 3 ± 0.1

(278-795) (288-7581) (129-11401) (17-9066) (0-18) (0.001-108.266) (1-4)

 Female reproductive

SWAZ-1 10 422 ± 6.6 953 ± 31.5 450 ± 70.3 446 ± 59.1 1 ± 0.1 2 ± 0.5 3 ± 0.2

(389-450) (794-1118) (187-904) (171-857) (1-2) (0.317-5.816) (2-4)

SWCA-3 11 642 ± 26.7 3692 ± 358.9 607 ± 84.5 839 ± 173.3 1 ± 0.2 21 ± 4.6 3 ± 0.2

(415-720) (1020-5256) (115-1060) (375-2309) (0-3) (2.347-56.932) (1-4)

SWNV-1 19 444 ± 9.1 1237 ± 72.5 843 ± 112.8 424 ± 65 2 ± 0.3 1 ± 0.1 3 ± 0.3

(385-526) (719-1830) (139-1766) (112-992) (0-6) (0.15-2.423) (0-4)

SWNV-2 18 480 ± 6.7 1568 ± 91 944 ± 85.1 402 ± 53.8 3 ± 0.4 1 ± 0.3 3 ± 0.1

(441-533) (1110-2462) (163-1694) (122-936) (1-7) (0.001-4.816) (2-4)

Table 3A. (Continued) Site and regional mean values for biomarker and ancillary data for COMMON CARP collections made at all sites 

in U.S. waters from March 1994 through August 1997.

[N, number of fish; E2, 17ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; mm, millimeter; g, grams; pg, picograms; mg, milligram; mL, milliliter; 

ND, nondetected]

Table 3A 19
20 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Regional site 

code

(fig. 1)

N Length

(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental

stage

SWNV-3 24 471 ± 7.6 1427 ± 64.4 843 ± 76.5 311 ± 33.9 4 ± 0.7 3 ± 0.7 3 ± 0.1

(403-526) (745-2193) (231-1401) (56-707) (1-16) (0.001-16.124) (2-3)

SWUT-5 1 720 ± 0 5000 ± 0 1159 ± 0 687 ± 0 2 ± 0 1 ± 0 3 ± 0

(720-720) (5000-5000) (1159-1159) (687-687) (2-2) (0.672-0.672) (3-3)

Summary 83 487 ± 9.1 1700 ± 113.2 790 ± 44.1 447 ± 36 3 ± 0.3 4 ± 0.9 3 ± 0.1

(385-720) (719-5256) (115-1766) (56-2309) (0-16) (0.001-56.932) (0-4)

 Male non-reproductive

SWCA-1 20 477 ± 10.8 1547 ± 102.7 1338 ± 220.2 3010 ± 385.9 0 ± 0 0 ± 0 3 ± 0.1

(416-566) (1009-2310) (88-2542) (152-5009) (0-1) (ND-0.12) (2-3)

SWCA-2 28 455 ± 18.7 1340 ± 153.1 414 ± 98.8 1988 ± 460 0 ± 0.1 0 ± 0 3 ± 0.1

(302-640) (334-2905) (51-2388) (183-9699) (0-2) (ND-0.183) (2-3)

SWCA-3 25 533 ± 13.1 2182 ± 177.6 894 ± 205.1 1531 ± 276 1 ± 0.1 0 ± 0.1 3 ± 0.1

(420-660) (477-3806) (51-4546) (149-4978) (0-3) (ND-3.148) (2-3)

SWUT-1 5 472 ± 59.5 1585 ± 629.8 629 ± 55.5 875 ± 119.3 1 ± 0.1 0 ± 0 2 ± 0.2

(336-674) (552-3983) (531-841) (575-1263) (1-1) (ND-0.024) (1-2)

SWUT-2 5 455 ± 42.9 1361 ± 285.6 286 ± 62 776 ± 212.8 0 ± 0.1 0 ± 0 3 ± 0

(347-570) (762-2168) (168-468) (347-1569) (0-1) (ND-0.043) (3-3)

SWUT-3 5 559 ± 77.4 2635 ± 956 638 ± 97.2 1098 ± 343.2 1 ± 0.1 0 ± 0 2 ± 0.4

(391-779) (870-5700) (281-835) (329-2324) (0-1) (ND-0.019) (1-3)

SWUT-4 4 403 ± 58.3 1506 ± 432.7 486 ± 40.8 580 ± 46.2 1 ± 0 0 ± 0 2 ± 0.4

(278-515) (473-2591) (366-546) (447-660) (1-1) (ND-0.061) (1-3)

SWUT-5 5 465 ± 30.1 1291 ± 236.2 504 ± 68.8 944 ± 149.1 1 ± 0.1 0 ± 0 2 ± 0.4

(380-552) (740-2060) (263-690) (521-1317) (0-1) (ND-0.018) (1-3)

Summary 97 485 ± 9.5 1688 ± 97.3 752 ± 82.8 1804 ± 185.4 1 ± 0 0 ± 0 3 ± 0.1

(278-779) (334-5700) (51-4546) (149-9699) (0-3) (ND-3.148) (1-3)

 Male reproductive

SWAZ-1 10 430 ± 7.9 887 ± 53 95 ± 3.8 261 ± 60.9 1 ± 0.1 0 ± 0.1 3 ± 0.2

(404-490) (717-1294) (77-118) (74-731) (0-1) (ND-0.678) (2-3)

SWCA-3 5 600 ± 17.9 2584 ± 283.9 355 ± 96.9 982 ± 268.7 1 ± 0.3 0 ± 0 3 ± 0.4

(544-649) (1923-3360) (181-731) (191-1866) (0-1) (ND-0.037) (1-3)

SWNV-1 20 460 ± 8.3 1236 ± 70.6 404 ± 93.4 638 ± 127.3 1 ± 0.1 0 ± 0 3 ± 0.2

(403-518) (745-1830) (71-1298) (127-2481) (0-2) (ND-0.005) (1-3)

SWNV-2 19 469 ± 7.9 1306 ± 72.7 180 ± 30.9 545 ± 83.3 0 ± 0 0 ± 0 3 ± 0.1

(393-538) (666-1885) (63-519) (109-1363) (0-1) (ND-0.233) (2-3)

SWNV-3 22 467 ± 8.3 1542 ± 79 282 ± 27.1 1279 ± 166.4 0 ± 0 0 ± 0 3 ± 0.1

(378-526) (1033-2193) (71-585) (257-2804) (0-1) (0.001-0.128) (2-3)

Summary 76 470 ± 5.8 1385 ± 58.1 269 ± 29.7 773 ± 76 0 ± 0 0 ± 0 3 ± 0.1

(378-649) (666-3360) (63-1298) (74-2804) (0-2) (ND-0.678) (1-3)

Table 3A. (Continued) Site and regional mean values for biomarker and ancillary data for COMMON CARP collections made at all sites 

in U.S. waters from March 1994 through August 1997.

[N, number of fish; E2, 17ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; mm, millimeter; g, grams; pg, picograms; mg, milligram; mL, milliliter; 

ND, nondetected]
Regional site 

code

(fig. 1)

N Length

(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental

stage

LARGEMOUTH BASS

Northern Mid-Continent Region

 Female non-reproductive

NMCO-3 8 332 ± 4.3 741 ± 32.7 977 ± 193.8 369 ± 107.2 3 ± 0.3 1 ± 1.1 2 ± 0

(309-348) (600-850) (403-1883) (111-1040) (2-4.3) (ND-8.833) (2-2)

NMIA-7 10 382 ± 7.3 972 ± 59.2 750 ± 46.9 558 ± 46.5 1 ± 0.2 3 ± 1.1 2 ± 0.1

(355-420) (682-1272) (604-1116) (304-771) (1-2.3) (0.116-11.276) (2-3)

NMIA-8 10 398 ± 10.4 1102 ± 96.3 997 ± 150.3 406 ± 66.5 3 ± 0.4 2 ± 1 2 ± 0

(337-456) (656-1726) (202-1371) (179-782) (1-4.6) (0.021-10.331) (2-2)

NMIL-1 10 319 ± 19.3 618 ± 117.6 236 ± 26.6 473 ± 95.9 1 ± 0.2 0 ± 0 2 ± 0.1

(251-408) (228-1173) (102-423) (128-906) (0-1.6) (ND-0.072) (1-3)

NMIN-4 4 297 ± 11.2 340 ± 59.4 681 ± 249.8 173 ± 55.1 4 ± 0.5 1 ± 1.3 2 ± 0.6

(266-316) (176-450) (369-1426) (97-337) (3-5.1) (ND-5.092) (0-3)

NMMO-3 5 339 ± 25.2 645 ± 157.6 494 ± 77.7 369 ± 76.6 2 ± 0.3 0 ± 0.2 2 ± 0

(296-426) (375-1225) (338-789) (176-622) (1-2.4) (ND-1.21) (2-2)

Summary 47 352 ± 7.4 796 ± 50.6 699 ± 65.1 422 ± 35.4 2 ± 0.2 1 ± 0.4 2 ± 0.1

(251-456) (176-1726) (102-1883) (97-1040) (0-5.1) (ND-11.276) (0-3)

 Male non-reproductive

NMCO-3 10 331 ± 4.7 759 ± 31.5 448 ± 28.4 1369 ± 237.5 0 ± 0.1 0 ± 0 2 ± 0

(295-349) (550-875) (329-544) (561-2812) (0-0.8) (ND-0.001) (2-2)

NMIA-7 10 337 ± 9.6 661 ± 59.6 551 ± 45.4 982 ± 87.3 1 ± 0.1 0 ± 0 2 ± 0.2

(275-387) (323-1030) (340-857) (432-1335) (0-1.2) (ND-0.026) (1-2)

NMIA-8 10 367 ± 7.8 817 ± 53.8 220 ± 20.3 654 ± 46.2 0 ± 0.1 0 ± 0 2 ± 0.1

(310-393) (449-1008) (99-291) (386-901) (0-0.8) (ND-0.006) (1-2)

NMIL-1 10 310 ± 17.3 539 ± 101.2 201 ± 11.8 750 ± 172.2 0 ± 0.1 0 ± 0 1 ± 0.1

(234-404) (192-1119) (153-251) (231-1755) (0-1) (ND-0.001) (1-2)

NMIN-4 5 301 ± 10 385 ± 44.5 566 ± 197.8 717 ± 334.5 2 ± 1.1 0 ± 0 2 ± 0.2

(265-318) (258-500) (315-1355) (129-1681) (0-6.2) (ND-0.008) (2-3)

NMMO-3 4 349 ± 31.8 800 ± 229.8 648 ± 108.4 1020 ± 438.7 1 ± 0.4 0 ± 0 2 ± 0

(287-435) (400-1450) (395-923) (191-2132) (0-2.1) (ND-0.001) (2-2)

Summary 49 334 ± 5.8 671 ± 36.9 400 ± 33.1 923 ± 83.7 1 ± 0.1 0 ± 0 2 ± 0.1

(234-435) (192-1450) (99-1355) (129-2812) (0-6.2) (ND-0.026) (1-3)

Northeastern Region

 Female non-reproductive

NECT-1 3 295 ± 39.2 446 ± 169.2 730 ± 326.5 133 ± 31.4 5 ± 1.1 0 ± 0 (1-0)

(254-373) (273-784) (307-1372) (93-195) (3-7) (ND-0.008) 1 ± 1

NEDC-1 5 340 ± 22.9 597 ± 148.9 228 ± 84.9 267 ± 122.5 1 ± 0.2 0 ± 0.2 (1-0)

(293-400) (354-1164) (51-546) (35-731) (1-1.8) (0.0016-0.88) 1 ± 1

Table 3B. Site and regional mean values for biomarker and ancillary data for LARGEMOUTH BASS collections made at all 

sites in U.S. waters from March 1994 through August 1997.

[N, number of fish; E2, 17ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; mm, millimeter; g, grams; pg, picograms; mg, milligram; mL, milliliter; 

ND, nondetected]

Table 3B  21
22  Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Regional site 

code

(fig. 1)

N Length

(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental

stage

NEMA -1 12 310 ± 9 479 ± 51.5 387 ± 85.4 376 ± 63.3 1 ± 0.3 0 ± 0 (1-0)

(264-375) (288-886) (33-958) (123-729) (0-2.5) (0.0008-0.008) 1 ± 1

NENY-2 4 307 ± 1.9 478 ± 21.4 375 ± 99.7 118 ± 84.6 9 ± 3.9 0 ± 0.1 (1-0)

(303-312) (430-534) (172-561) (18-371) (2-17.7) (ND-0.4) 1 ± 1

NENY-3 7 413 ± 18.9 890 ± 53.2 444 ± 87.1 116 ± 25.1 5 ± 1.8 0 ± 0 (2-0.2)

(359-495) (785-956) (124-741) (46-244) (1-15.2) (0.006-0.4) 1 ± 2

NENY-4 2 439 ± 45.5 1414 ± 399 350 ± 92.5 97 ± 76 7 ± 4.8 0 ± 0 (1-0)

(393-484) (1015-1813) (257-442) (21-173) (3-12.2) (0.097-0.139) 1 ± 1

NEOH-1 2 330 ± 2 602 ± 38 322 ± 42.5 119 ± 32 3 ± 0.4 1 ± 0.4 (3-0)

(328-332) (564-640) (279-364) (87-151) (2-3.2) (0.245-1.003) 3 ± 3

NEWV -2 10 308 ± 4.3 352 ± 15.1 683 ± 121 335 ± 60.5 2 ± 0.3 0 ± 0.1 (2-0.1)

(284-325) (278-418.2) (304-1403) (84-601) (1-3.6) (ND-0.512) 1 ± 2

Summary 45 334 ± 8.3 541 ± 47.4 461 ± 48.5 251 ± 30.7 3 ± 0.6 0 ± 0 (1-0.1)

(254-495) (273-1813) (33-1403) (18-731) (0-17.7) (ND-1.003) 1 ± 3

 Male non-reproductive

NECT-1 4 314 ± 26.5 389 ± 37.8 709 ± 282.6 757 ± 259.4 1 ± 0.2 0 ± 0 3 ± 0

(284-393) (314-494) (231-1500) (257-1206) (1-1.5) (ND-0.002) (3-3)

NEDC-1 7 339 ± 13.3 581 ± 65.5 121 ± 38 129 ± 42.2 1 ± 0.4 0 ± 0 3 ± 0

(274-386) (228-780) (21-294) (46-327) (0-3.7) (0.001-0.002) (3-3)

NEMA -1 7 296 ± 10.7 393 ± 42.2 526 ± 122.3 383 ± 85.1 2 ± 0.5 0 ± 0 2 ± 0.4

(251-335) (248-542) (107-1054) (176-807) (0-4.2) (0.001-0.004) (0-3)

NENY-2 6 292 ± 5.2 378 ± 15 200 ± 44.7 68 ± 15.2 3 ± 0.8 0 ± 0 3 ± 0

(277-310) (325-420) (44-373) (39-142) (1-6.6) (0.001-0.215) (3-3)

NENY-3 12 339 ± 15.4 716 ± 134.5 60 ± 18.9 151 ± 31 0 ± 0.1 0 ± 0.2 3 ± 0

(252-422) (581-850) (13-200) (63-446) (0-1) (ND-3) (3-3)

NENY-4 2 377 ± 7 874 ± 19 35 ± 23.5 41 ± 25.5 2 ± 1.9 0 ± 0 3 ± 0

(370-384) (855-893) (11-58) (15-66) (0-3.9) (ND-0.001) (3-3)

NEOH-1 2 395 ± 15.5 945 ± 155.5 359 ± 50.5 300 ± 141.5 2 ± 0.9 0 ± 0 2 ± 0

(379-410) (789-1100) (308-409) (158-441) (1-2.6) (ND-0.001) (2-2)

NEWV -2 10 300 ± 6.9 294 ± 16.6 313 ± 16.4 748 ± 64.7 0 ± 0 0 ± 0 2 ± 0.1

(263-330) (214-377.8) (201-387) (426-1087) (0-0.6) (ND-0.114) (1-3)

Summary 50 321 ± 6.2 466 ± 34 264 ± 39.8 340 ± 47.3 1 ± 0.2 0 ± 0.1 3 ± 0.1

(251-422) (214-1100) (11-1500) (15-1206) (0-6.6) (ND-3) (0-3)

 Female reproductive

NEMA -1 4 303 ± 7.9 420 ± 39.4 920 ± 120.6 531 ± 92.8 2 ± 0.1 4 ± 2 3 ± 0.4

(286-318) (350-497) (732-1243) (382-784) (2-1.9) (ND-8.242) (2-4)

NENY-3 14 399 ± 12.5 1001 ± 99.8 3221 ± 725.6 754 ± 196.4 5 ± 0.7 5 ± 0.9 3 ± 0.2

(325-478) (478-1648) (384-9670) (108-2306) (2-12.4) (ND-12.441) (3-5)

Summary 18 378 ± 13.7 872 ± 97.1 2710 ± 606.2 704 ± 154.3 5 ± 0.7 5 ± 0.8 3 ± 0.1

(286-478) (350-1648) (384-9670) (108-2306) (2-12.4) (ND-12.441) (2-5)

Table 3B. (Continued) Site and regional mean values for biomarker and ancillary data for LARGEMOUTH BASS collections made at all 

sites in U.S. waters from March 1994 through August 1997.

[N, number of fish; E2, 17ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; mm, millimeter; g, grams; pg, picograms; mg, milligram; mL, milliliter; 

ND, nondetected]
Regional site 

code

(fig. 1)

N Length

(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental

stage

 Male reproductive

NEMA -1 12 296 ± 4.6 401 ± 21 823 ± 136.9 1149 ± 129.9 1 ± 0.2 0 ± 0 3 ± 0

(276-328) (321-541) (280-1621) (410-1706) (0-3) (ND-0.245) (3-3)

NENY-3 2 359 ± 27.5 746 ± 192.5 526 ± 36.5 1475 ± 46 0 ± 0 0 ± 0 3 ± 0

(331-386) (553-938) (489-562) (1429-1521) (0-0.4) (ND-0.001) (3-3)

Summary 14 305 ± 7.8 450 ± 43 781 ± 120.2 1195 ± 115.2 1 ± 0.2 0 ± 0 3 ± 0

(276-386) (321-938) (280-1621) (410-1706) (0-3) (ND-0.245) (3-3)

Northwestern Region

 Female non-reproductive

NWID-1 9 365 ± 22.8 891 ± 170.3 642 ± 91.4 287 ± 42.2 2 ± 0.4 0 ± 0.2 1 ± 0.1

(257-492) (253-2000) (242-1134) (167-481) (1-4.2) (0.026-1.535) (1-2)

NWMT-1 10 407 ± 25.7 1320 ± 249.3 855 ± 125.4 383 ± 67.3 2 ± 0.2 1 ± 0.2 2 ± 0.2

(279-496) (280-2600) (398-1585) (112-805) (2-3.7) (0.158-2.355) (1-3)

NWOR-1 5 328 ± 13.2 571 ± 84.7 588 ± 80.5 266 ± 75.6 3 ± 0.8 0 ± 0 1 ± 0

(280-355) (306-758) (412-797) (101-532) (1-5.7) (ND-0.138) (1-1)

NWWA-3 7 355 ± 32.9 920 ± 316.7 684 ± 157.6 360 ± 110.5 2 ± 0.5 5 ± 4.8 1 ± 0.3

(269-522) (299-2600) (239-1465) (122-908) (1-4.5) (ND-33.665) (1-3)

NWWA-6 4 363 ± 40.8 944 ± 329.2 1123 ± 164 404 ± 121.2 3 ± 0.5 1 ± 0.5 2 ± 0.3

(263-450) (272-1803) (775-1406) (193-689) (2-4.3) (0.299-2.631) (1-2)

Summary 35 369 ± 12.7 980 ± 113.9 758 ± 61.1 340 ± 34.7 3 ± 0.2 2 ± 1 1 ± 0.1

(257-522) (253-2600) (239-1585) (101-908) (1-5.7) (ND-33.665) (1-3)

 Male non-reproductive

NWID-1 7 344 ± 12.6 673 ± 79.4 341 ± 98 1540 ± 987.6 1 ± 0.2 0 ± 0 2 ± 0.3

(314-397) (444-970) (131-815) (349-7451) (0-2) (ND-0.001) (1-3)

NWMT-1 10 339 ± 22.3 756 ± 198.3 251 ± 46.4 528 ± 88.6 1 ± 0.2 0 ± 0 3 ± 0.2

(252-461) (114-1880) (113-591) (121-976) (0-2.3) (ND-0.083) (1-3)

NWOR-1 3 286 ± 25.1 373 ± 110.9 641 ± 210.2 813 ± 354.3 2 ± 1.2 0 ± 0 3 ± 0.3

(255-336) (224-590) (310-1031) (260-1473) (0-4) (ND-0.001) (2-3)

NWWA-3 6 314 ± 35.7 534 ± 194.5 376 ± 107.2 661 ± 143.3 1 ± 0.1 0 ± 0.1 2 ± 0

(240-440) (144-1248) (131-848) (331-1307) (0-1.2) (ND-0.669) (2-2)

NWWA-6 2 387 ± 31 915 ± 223.5 143 ± 36.5 317 ± 118 0 ± 0.1 0 ± 0 3 ± 0.5

(356-418) (691-1138) (106-179) (199-435) (0-0.5) (ND-0.001) (2-3)

Summary 28 333 ± 12.1 658 ± 86.6 334 ± 46.7 825 ± 252.5 1 ± 0.2 0 ± 0 2 ± 0.1

(240-461) (114-1880) (106-1031) (121-7451) (0-4) (ND-0.669) (1-3)

Table 3B. (Continued) Site and regional mean values for biomarker and ancillary data for LARGEMOUTH BASS collections made at all 

sites in U.S. waters from March 1994 through August 1997.

[N, number of fish; E2, 17ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; mm, millimeter; g, grams; pg, picograms; mg, milligram; mL, milliliter; 

ND, nondetected]

Table 3B  23
24  Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Regional site 

code

(fig. 1)

N Length

(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental

stage

Southern Mid-Continent Region

 Female non-reproductive

SMAR-3 10 333 ± 10.1 618 ± 93.2 1405 ± 455 355 ± 176.8 9 ± 2.2 1 ± 0.7 2 ± 0.2

(294-379) (313-1255) (256-5221) (55-1838) (0-24.7) (ND-6.942) (1-2)

SMAR-4 10 419 ± 25.9 1286 ± 245.4 530 ± 71.8 288 ± 26.1 2 ± 0.3 0 ± 0 2 ± 0

(305-540) (391-2400) (235-845) (144-429) (1-4.1) (ND-0.197) (2-2)

SMIL-2 9 341 ± 9.4 488 ± 46.8 926 ± 244.6 779 ± 105.4 2 ± 0.5 0 ± 0 2 ± 0

(310-377) (343-721) (245-2434) (95-1312) (0-4.6) (ND-0.001) (2-2)

SMLA-3 16 348 ± 17.8 702 ± 110.7 689 ± 79.4 480 ± 53 2 ± 0.2 0 ± 0 1 ± 0.2

(231-462) (140-1493) (404-1591) (210-899) (0-3.8) (ND-0.001) (0-2)

SMOK-1 7 411 ± 20.9 1161 ± 179.6 1726 ± 368.1 282 ± 60.4 7 ± 1 2 ± 0.8 3 ± 0.1

(332-495) (500-1950) (705-3386) (114-513) (4-11.3) (ND-5.443) (2-3)

SMOK-2 12 419 ± 15.2 1110 ± 132.3 970 ± 147.2 925 ± 144.6 2 ± 0.5 3 ± 1.3 3 ± 0.1

(351-510) (600-1900) (462-2417) (87-1999) (0-6) (ND-15.897) (2-3)

SMOK-3 9 393 ± 29.3 1228 ± 255.1 1491 ± 204.8 675 ± 219.3 4 ± 0.9 1 ± 0.3 3 ± 0.2

(280-510) (350-2400) (403-2356) (151-2203) (0-8.6) (ND-3.169) (2-3)

SMTN-1 4 311 ± 32.9 426 ± 149.5 4283 ± 1408.4 1050 ± 303.5 9 ± 6.3 52 ± 18 3 ± 0.3

(250-400) (191-859) (121-6330) (228-1512) (0-27.8) (ND-77.918) (2-3)

SMTN-2 7 348 ± 32.6 763 ± 249.4 374 ± 79.5 183 ± 49.8 3 ± 0.8 0 ± 0 2 ± 0

(231-469) (151-1822) (196-690) (44-422) (1-5.4) (ND-0.341) (2-2)

SMTN-3 11 421 ± 23.5 1196 ± 185.4 386 ± 54.9 164 ± 23.9 3 ± 1.1 0 ± 0 1 ± 0.4

(285-543) (340-2381) (189-779) (58-314) (1-13.4) (0.005-0.415) (0-3)

SMTN-4 4 286 ± 22.4 295 ± 65.2 362 ± 71.1 65 ± 9.2 6 ± 1 0 ± 0.2 3 ± 0.3

(231-325) (142-425) (230-563) (49-89) (4-8) (ND-0.925) (2-3)

SMTN-5 10 375 ± 31.3 963 ± 256.8 657 ± 240.2 248 ± 80 3 ± 0.8 0 ± 0.1 2 ± 0.3

(260-573) (232-2920) (52-2261) (70-869) (0-7.6) (ND-0.863) (1-3)

SMTX-1 6 339 ± 32.5 308 ± 3 313 ± 59.9 178 ± 30.4 2 ± 0.4 0 ± 0 0 ± 0.2

(306-371) (305-311) (154-566) (113-314) (1-3.9) (ND-0.001) (0-1)

SMTX-5 6

<—— Not available——>

0 ± 0 1 ± 0.2

(ND-0.001) (0-1)

SMTX-6 1

<—— Not available ——>

0 ± 0 0 ± 0

(ND-0.001) (0-0)

SMTX-7 13 403 ± 19.8 961 ± 136.1 721 ± 68.3 223 ± 52.2 6 ± 1.2 1 ± 0.4 2 ± 0.1

(309-540) (366-2100) (391-1061) (57-648) (1-16.1) (ND-4.874) (2-3)

Summary 135 377 ± 6.9 897 ± 53.6 899 ± 86.9 416 ± 35.5 4 ± 0.4 2 ± 0.9 2 ± 0.1

(231-573) (140-2920) (52-6330) (44-2203) (0-27.8) (ND-77.918) (0-3)

Table 3B. (Continued) Site and regional mean values for biomarker and ancillary data for LARGEMOUTH BASS collections made at all 

sites in U.S. waters from March 1994 through August 1997.

[N, number of fish; E2, 17ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; mm, millimeter; g, grams; pg, picograms; mg, milligram; mL, milliliter; 

ND, nondetected]
Regional site 

code

(fig. 1)

N Length

(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental

stage

 Male non-reproductive

SMAR-3 9 313 ± 9.3 428 ± 39.8 555 ± 87.7 832 ± 151.1 1 ± 0.6 0 ± 0.3 2 ± 0.1

(275-370) (269-655) (277-1150) (181-1544) (0-6.4) (ND-2.817) (1-2)

SMAR-4 10 336 ± 17.5 563 ± 113.9 358 ± 36.3 1459 ± 464.4 0 ± 0.1 0 ± 0 2 ± 0.1

(262-411) (200-1251) (249-550) (319-5144) (0-1.1) (ND-0.001) (1-2)

SMIL-2 14 352 ± 16.9 637 ± 122.1 344 ± 28.9 1191 ± 156.4 0 ± 0 0 ± 0 2 ± 0.1

(249-515) (176-1918) (199-596) (418-2648) (0-0.6) (ND-0.005) (1-2)

SMLA-3 7 290 ± 20.2 361 ± 87.4 361 ± 37.5 500 ± 41.7 1 ± 0.1 0 ± 0 1 ± 0.2

(235-372) (149-733) (198-450) (340-689) (0-1.2) (ND-0.001) (1-2)

SMOK-1 8 379 ± 6.4 800 ± 47.2 355 ± 22.5 1192 ± 185 0 ± 0.1 0 ± 0 2 ± 0.1

(354-405) (650-1000) (280-427) (421-1942) (0-0.7) (ND-0.01) (2-3)

SMOK-2 10 394 ± 12.7 944 ± 105.1 645 ± 63.1 764 ± 141 1 ± 0.2 0 ± 0.2 2 ± 0.1

(313-453) (450-1500) (341-1022) (214-1420) (0-2.1) (ND-2.408) (1-3)

SMOK-3 8 365 ± 21.8 849 ± 145.2 765 ± 168 1454 ± 113.4 1 ± 0.1 0 ± 0 2 ± 0.2

(272-457) (300-1500) (329-1823) (1044-1951) (0-0.9) (ND-0.001) (2-3)

SMTN-1 7 320 ± 21 429 ± 114.8 415 ± 107.7 847 ± 199.5 1 ± 0.4 0 ± 0 2 ± 0

(260-425) (198-1075) (143-958) (333-1549) (0-2.9) (ND-0.003) (2-2)

SMTN-2 4 353 ± 12.9 667 ± 73.2 489 ± 155.5 239 ± 59.3 3 ± 1.5 0 ± 0 2 ± 0

(315-373) (463-802) (204-788) (115-389) (1-6.9) (ND-0.001) (2-2)

SMTN-3 4 414 ± 37.7 869 ± 242.9 431 ± 186.3 533 ± 141 1 ± 0.2 0 ± 0 2 ± 0

(330-484) (510-1332) (124-969) (208-852) (0-1.3) (ND-0.001) (2-2)

SMTN-4 5 284 ± 18.5 330 ± 60.3 348 ± 65.7 144 ± 16.4 3 ± 0.6 0 ± 0 2 ± 0.2

(230-335) (167-525) (214-594) (106-198) (1-4.9) (ND-0.102) (2-3)

SMTN-5 13 311 ± 17 458 ± 73.8 297 ± 19.7 907 ± 195.2 1 ± 0.2 0 ± 0 2 ± 0.2

(233-415) (170-964) (196-426) (88-2574) (0-2.2) (ND-0.008) (1-3)

SMTX-1 4 262 ± 0 285 ± 0 227 ± 13.5 646 ± 134.4 0 ± 0.1 0 ± 0 3 ± 0.3

(262-262) (285-285) (187-245) (291-931) (0-0.8) (ND-0.001) (2-3)

SMTX-5 2 387 ± 0 295 ± 0 188 ± 4 265 ± 33 1 ± 0.1 0 ± 0 3 ± 0.5

(387-387) (295-295) (184-192) (232-298) (1-0.8) (ND-0.001) (2-3)

SMTX-6 5 317 ± 71.5 302 ± 34.5 237 ± 22 452 ± 96.4 1 ± 0.1 0 ± 0 2 ± 0.4

(245-388) (267-336) (209-324) (274-813) (0-0.8) (ND-0.001) (1-3)

SMTX-7 13 381 ± 13.5 796 ± 92 657 ± 42.7 745 ± 168.9 1 ± 0.3 0 ± 0 2 ± 0.1

(305-470) (373-1467) (479-936) (205-2215) (0-3) (ND-0.001) (2-3)

Summary 123 345 ± 5.7 617 ± 33.2 443 ± 22.5 875 ± 62.7 1 ± 0.1 0 ± 0 2 ± 0

(230-515) (149-1918) (124-1823) (88-5144) (0-6.9) (ND-2.817) (1-3)

Table 3B. (Continued) Site and regional mean values for biomarker and ancillary data for LARGEMOUTH BASS collections made at all 

sites in U.S. waters from March 1994 through August 1997.

[N, number of fish; E2, 17ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; mm, millimeter; g, grams; pg, picograms; mg, milligram; mL, milliliter; 

ND, nondetected]

Table 3B  25
26  Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Regional site 

code

(fig. 1)

N Length

(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental

stage

Southwest Region

 Female non-reproductive

SWCA-1 23 398 ± 17.3 1068 ± 147.6 802 ± 77.1 221 ± 27.9 5 ± 0.9 0 ± 0 1 ± 0.1

(273-580) (222-2804) (289-1539) (38-662) (1-20.8) (ND-0.564) (0-1)

SWCA-2 19 392 ± 17.3 1046 ± 116.2 455 ± 130.7 101 ± 22.5 5 ± 1 0 ± 0.1 1 ± 0.1

(285-595) (320-2020) (54-2229) (12-362) (0-17.4) (ND-1.123) (1-2)

SWCA-3 31 376 ± 10.5 912 ± 80.8 374 ± 60.8 273 ± 36.6 2 ± 0.3 0 ± 0.1 1 ± 0.1

(270-482) (277-2010) (29-1345) (59-811) (0-9.3) (ND-3.541) (1-2)

Summary 73 387 ± 8.3 996 ± 64.9 530 ± 53.2 212 ± 20.3 4 ± 0.5 0 ± 0.1 1 ± 0.1

(270-595) (222-2804) (29-2229) (12-811) (0-20.8) (ND-3.541) (0-2)

 Male non-reproductive

SWCA-1 11 316 ± 20.1 514 ± 117.6 789 ± 119.4 904 ± 194.9 1 ± 0.3 0 ± 0.2 3 ± 0

(252-448) (197-1389) (207-1370) (98-1894) (0-3.4) (ND-1.8) (3-3)

SWCA-2 21 351 ± 9.4 709 ± 57.1 187 ± 51.1 202 ± 72 2 ± 0.3 0 ± 0 3 ± 0

(258-435) (207-1333) (19-821) (18-1131) (0-4.2) (ND-0.039) (2-3)

SWCA-3 20 354 ± 12.4 677 ± 56.4 197 ± 39 260 ± 37.9 1 ± 0.2 0 ± 0 3 ± 0

(226-475) (230-1263) (35-761) (97-656) (0-4) (ND-0.117) (3-3)

Summary 52 345 ± 7.6 656 ± 40.7 318 ± 48.9 373 ± 64 1 ± 0.1 0 ± 0 3 ± 0

(226-475) (197-1389) (19-1370) (18-1894) (0-4.2) (ND-1.8) (2-3)

Table 3B. (Continued) Site and regional mean values for biomarker and ancillary data for LARGEMOUTH BASS collections made at all 

sites in U.S. waters from March 1994 through August 1997.

[N, number of fish; E2, 17ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; mm, millimeter; g, grams; pg, picograms; mg, milligram; mL, milliliter; 

ND, nondetected]
Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Allegheny Riv @ Natrona, PA NE 1995 Oct NR F 660 3900 2379 522 4.56 1.326 3

Allegheny Riv @ Natrona, PA NE 1995 Oct NR F 589 2800 658 118 5.58 1.510 3

Allegheny Riv @ Natrona, PA NE 1995 Oct NR F 498 1450 584 465 1.26 4.671 3

Allegheny Riv @ Natrona, PA NE 1995 Oct NR F 475 1500 259 105 2.47 0.314 3

Allegheny Riv @ Natrona, PA NE 1995 Oct NR F 460 1200 347 88 3.94 1.247 3

Allegheny Riv @ Natrona, PA NE 1995 Oct NR F 429 1000 2768 1504 1.84 1.016 3

Allegheny Riv @ Natrona, PA NE 1995 Oct NR M 594 2750 300 1336 0.22 ND 2

Allegheny Riv @ Natrona, PA NE 1995 Oct NR M 498 1700 407 4090 0.10 0.027 2

Allegheny Riv @ Natrona, PA NE 1995 Oct NR M 495 1500 309 1319 0.23 ND 3

Allegheny Riv @ Natrona, PA NE 1995 Oct NR M 401 1700 407 1741 0.23 0.008 2

Allegheny Riv @ Natrona, PA NE 1995 Oct NR M 373 1700 383 3293 0.12 0.008 2

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR F 555 2150 1582 320 4.944 1.772 3

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR F 467 1375 1901 633 3.003 2.315 3

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR F 460 1250 1377 257 5.358 1.932 3

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR F 411 860 1380 221 6.244 8.369 3

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR F 385 850 589 221 2.665 2.921 3

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR F 382 680 489 213 2.296 9.285 3

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR F 373 690 743 117 6.350 1.800 3

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR F 364 598 538 158 3.405 ND 1

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR F 353 650 697 31 22.484 1.187 3

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR F 350 560 366 93 3.935 0.275 1

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR F 342 550 974 354 2.751 1.640 3

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR M 382 720 428 1174 0.365 ND 2

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR M 377 640 390 872 0.447 ND 3

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR M 373 700 424 639 0.664 ND 3

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR M 370 700 429 820 0.523 ND 2

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR M 368 650 403 985 0.409 ND 3

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR M 365 630 505 138 3.659 ND 2

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR M 347 600 469 856 0.548 ND 2

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR F 670 4350 365 294 1.241 3.586 3

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR F 570 2050 271 943 0.287 ND 0

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR F 541 2075 2489 482 5.164 1.996 3

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR F 537 2300 1536 249 6.169 0.143 3

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR F 533 1850 2317 497 4.662 ND 1

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR F 524 1700 1393 374 3.725 0.972 3

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR F 510 1700 1309 366 3.577 3.717 3

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR F 500 1250 2286 568 4.025 ND 0

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR F 481 1450 1277 423 3.019 1.130 3

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR F 473 1300 1239 196 6.321 0.477 3

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR F NA NA 1208 217 5.567 0.176 3

Table 4. All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. waters 

from August 1994 through October 1997.

Table 4  27
28  Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR M 575 2500 285 566 0.504 ND 3

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR M 547 1550 246 359 0.685 ND 1

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR M 500 1650 461 1002 0.460 ND 3

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR M 495 1600 325 2016 0.161 ND 3

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR M 487 1650 385 1234 0.312 ND 3

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR M 483 1550 356 654 0.544 ND 3

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR M 470 1400 331 545 0.607 ND 3

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR M 457 1300 287 381 0.753 ND 3

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR M 432 1000 372 1031 0.361 ND 1

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR F 570 2365 474 172 2.76 2.299 3

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR F 565 2582 1788 296 6.04 2.985 3

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR F 552 1990 1013 295 3.43 5.995 3

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR F 550 2792 4537 510 8.90 1.523 3

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR F 530 1785 2927 779 3.76 2.317 3

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR F 510 1861 1859 338 5.50 3.042 3

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR F 490 1762 1888 531 3.56 2.502 3

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR F 485 1615 147 166 0.89 1.122 3

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR F 473 1480 1159 297 3.90 5.195 3

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR M 700 4200 339 1281 0.26 0.014 1

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR M 622 2933 1228 1599 0.77 ND 1

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR M 610 1858 107 2337 0.05 ND 1

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR M 580 2250 585 2017 0.29 ND 1.5

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR M 555 1560 327 7227 0.05 ND 1

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR M 521 1600 892 8492 0.11 ND 1

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR M 520 1794 501 3992 0.13 0.008 1

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR M 510 1660 196 6801 0.03 ND 1

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR M 505 1662 706 930 0.76 ND 1

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR M 330 643 80 632 0.13 ND 1

Bear Riv @ Corine UT SW 1998 Aug R F 700 4572 2176 452 4.814 3.66 2

Bear Riv @ Corine UT SW 1998 Aug R F 682 4206 977 586 1.667 4.742 3

Bear Riv @ Corine UT SW 1998 Aug R F 647 3562 791 445 1.778 5.757 3

Bear Riv @ Corine UT SW 1998 Aug R F 530 1890 1156 539 2.145 4.108 3

Bear Riv @ Corine UT SW 1998 Aug R F 495 1540 1181 300 3.937 2.774 3

Bear Riv @ Corine UT SW 1998 Aug R M 674 3983 632 1263 0.500 ND 2

Bear Riv @ Corine UT SW 1998 Aug R M 524 1668 841 966 0.871 0.024 2

Bear Riv @ Corine UT SW 1998 Aug R M 442 1031 558 686 0.813 ND 2

Bear Riv @ Corine UT SW 1998 Aug R M 383 690 584 884 0.661 ND 2

Bear Riv @ Corine UT SW 1998 Aug R M 336 552 531 575 0.923 ND 1

Big Horn Riv @ Hardin, MT NMC 1995 Oct NR F 681 3900 3101 929 3.34 6.821 3

Big Horn Riv @ Hardin, MT NMC 1995 Oct NR F 640 3600 2564 195 13.15 5.826 3
Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Big Horn Riv @ Hardin, MT NMC 1995 Oct NR F 623 3600 1782 748 2.38 1.365 3

Big Horn Riv @ Hardin, MT NMC 1995 Oct NR F 610 3150 1743 206 8.46 3.289 3

Big Horn Riv @ Hardin, MT NMC 1995 Oct NR F 596 2800 3347 762 4.39 0.827 3

Big Horn Riv @ Hardin, MT NMC 1995 Oct NR F 592 2600 4285 929 4.61 1.661 3

Big Horn Riv @ Hardin, MT NMC 1995 Oct NR F 556 2440 1455 132 11.02 1.307 3

Big Horn Riv @ Hardin, MT NMC 1995 Oct NR F 546 2250 2773 341 8.13 1.271 3

Big Horn Riv @ Hardin, MT NMC 1995 Oct NR F 535 2000 1280 195 6.56 ND 3

Big Horn Riv @ Hardin, MT NMC 1995 Oct NR F 522 2000 1540 462 3.33 1.523 3

Big Horn Riv @ Hardin, MT NMC 1995 Oct NR F 487 1490 2086 1644 1.27 2.405 3

Big Horn Riv @ Hardin, MT NMC 1995 Oct NR F 474 1400 2961 1061 2.79 3.974 3

Big Horn Riv @ Hardin, MT NMC 1995 Oct NR M 585 1420 1038 2464 0.42 ND 2

Big Horn Riv @ Hardin, MT NMC 1995 Oct NR M 577 2600 1136 2594 0.44 ND 2

Big Horn Riv @ Hardin, MT NMC 1995 Oct NR M 565 2200 939 1184 0.79 0.007 3

Big Horn Riv @ Hardin, MT NMC 1995 Oct NR M 542 2000 716 2970 0.24 ND 1

Big Horn Riv @ Hardin, MT NMC 1995 Oct NR M 511 1730 899 1209 0.74 ND 1

Big Horn Riv @ Hardin, MT NMC 1995 Oct NR M 493 1500 1388 2368 0.59 0.958 2

Big Horn Riv @ Hardin, MT NMC 1995 Oct NR M 489 1500 1418 2884 0.49 ND 2

Big Horn Riv @ Hardin, MT NMC 1995 Oct NR M 470 1250 1237 2464 0.50 0.010 2.5

Big Sunflower Riv @ Anguilla, MS SMC 1995 Aug R F 558 2432 1564 264 5.924 0.012 3.0

Big Sunflower Riv @ Anguilla, MS SMC 1995 Aug R F 557 1976 1137 165 6.891 0.199 3.0

Big Sunflower Riv @ Anguilla, MS SMC 1995 Aug R F 518 1481 1630 479 3.403 0.130 3.0

Big Sunflower Riv @ Anguilla, MS SMC 1995 Aug R F 489 1302 1634 183 8.929 0.128 1.0

Big Sunflower Riv @ Anguilla, MS SMC 1995 Aug R F 436 991 484 227 2.132 ND 3.0

Big Sunflower Riv @ Anguilla, MS SMC 1995 Aug R F 430 1103 1412 315 4.483 0.023 2.0

Big Sunflower Riv @ Anguilla, MS SMC 1995 Aug R F 419 907 23 465 0.049 ND 3.0

Big Sunflower Riv @ Anguilla, MS SMC 1995 Aug R F 413 1095 1113 297 3.747 0.228 3.0

Big Sunflower Riv @ Anguilla, MS SMC 1995 Aug R M 548 1829 228 201 1.134 0.178 1.0

Big Sunflower Riv @ Anguilla, MS SMC 1995 Aug R M 505 1625 101 590 0.171 0.012 .

Big Sunflower Riv @ Anguilla, MS SMC 1995 Aug R M 455 1128 786 536 1.466 ND 1.0

Big Sunflower Riv @ Anguilla, MS SMC 1995 Aug R M 449 1062 303 584 0.519 ND 1.0

Big Sunflower Riv @ Anguilla, MS SMC 1995 Aug R M 443 930 78 269 0.290 ND .

Big Sunflower Riv @ Anguilla, MS SMC 1995 Aug R M 410 325 270 738 0.366 ND 1.0

Big Sunflower Riv @ Anguilla, MS SMC 1995 Aug R M 398 714 301 375 0.803 0.007 1.0

Big Sunflower Riv @ Anguilla, MS SMC 1995 Aug R M 368 671 223 213 1.047 ND 1.0

Billy Clapp L nr Stratford, WA NW 1995 July R F 542 2111 1287 313 4.112 7.113 3.0

Billy Clapp L nr Stratford, WA NW 1995 July R F 538 2451 1309 379 3.454 21.916 3.0

Billy Clapp L nr Stratford, WA NW 1995 July R F 531 2265 1402 934 1.501 19.221 3.0

Billy Clapp L nr Stratford, WA NW 1995 July R F 521 2367 1485 498 2.982 62.448 3.0

Billy Clapp L nr Stratford, WA NW 1995 July R F 513 2049 1295 570 2.272 24.392 3.0

Billy Clapp L nr Stratford, WA NW 1995 July R F 510 1795 1875 778 2.410 4.596 3.0

Table 4  29
30 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Billy Clapp L nr Stratford, WA NW 1995 July R F 490 1714 979 560 1.748 19.677 3.0

Billy Clapp L nr Stratford, WA NW 1995 July R F 489 1754 464 45 10.311 16.836 3.0

Billy Clapp L nr Stratford, WA NW 1995 July R F 462 1487 1309 518 2.527 11.563 3.0

Billy Clapp L nr Stratford, WA NW 1995 July R F 453 1454 1481 415 3.569 11.746 3.0

Billy Clapp L nr Stratford, WA NW 1995 July R M 571 2688 620 910 0.681 ND 3.0

Billy Clapp L nr Stratford, WA NW 1995 July R M 555 2091 226 591 0.382 ND 2.0

Billy Clapp L nr Stratford, WA NW 1995 July R M 546 1980 1179 2020 0.584 ND 3.0

Billy Clapp L nr Stratford, WA NW 1995 July R M 538 1928 846 1477 0.573 ND 3.0

Billy Clapp L nr Stratford, WA NW 1995 July R M 505 1817 730 1407 0.519 ND 3.0

Billy Clapp L nr Stratford, WA NW 1995 July R M 494 1464 201 975 0.206 ND 2.0

Billy Clapp L nr Stratford, WA NW 1995 July R M 491 1429 387 1272 0.304 ND 1.0

Billy Clapp L nr Stratford, WA NW 1995 July R M 480 1345 304 549 0.554 ND 3.0

Billy Clapp L nr Stratford, WA NW 1995 July R M 467 1056 301 961 0.313 ND 1.0

Billy Clapp L nr Stratford, WA NW 1995 July R M 464 1327 356 1194 0.298 ND 2.0

Boque Phalia @ Leland, MS SMC 1995 Aug R F 528 2316 2117 1114 1.900 2.238 3.0

Boque Phalia @ Leland, MS SMC 1995 Aug R F 525 1026 1266 601 2.106 0.127 3.0

Boque Phalia @ Leland, MS SMC 1995 Aug R F 500 1617 1179 592 1.992 0.143 1.0

Boque Phalia @ Leland, MS SMC 1995 Aug R F 485 1385 236 92 2.565 0.140 3.0

Boque Phalia @ Leland, MS SMC 1995 Aug R F 440 1132 1185 954 1.242 0.217 3.0

Boque Phalia @ Leland, MS SMC 1995 Aug R F 428 1055 2141 424 5.050 0.433 3.0

Boque Phalia @ Leland, MS SMC 1995 Aug R F 421 921 2270 383 5.927 0.278 3.0

Boque Phalia @ Leland, MS SMC 1995 Aug R F 386 762 1866 797 2.341 2.023 2.0

Boque Phalia @ Leland, MS SMC 1995 Aug R F 383 745 2180 88 24.773 1.392 3.0

Boque Phalia @ Leland, MS SMC 1995 Aug R F 356 613 2318 346 6.699 0.332 3.0

Boque Phalia @ Leland, MS SMC 1995 Aug R M 546 1977 51 950 0.054 ND 2.0

Boque Phalia @ Leland, MS SMC 1995 Aug R M 500 1547 161 1093 0.147 ND 2.0

Boque Phalia @ Leland, MS SMC 1995 Aug R M 460 1159 757 634 1.194 ND 2.0

Boque Phalia @ Leland, MS SMC 1995 Aug R M 450 1137 70 704 0.099 ND 2.0

Boque Phalia @ Leland, MS SMC 1995 Aug R M 443 1101 55 458 0.120 ND 2.0

Boque Phalia @ Leland, MS SMC 1995 Aug R M 441 1177 355 624 0.569 ND 2.0

Boque Phalia @ Leland, MS SMC 1995 Aug R M 425 921 224 921 0.243 ND 1.0

Boque Phalia @ Leland, MS SMC 1995 Aug R M 404 774 52 644 0.081 ND 2.0

Boque Phalia @ Leland, MS SMC 1995 Aug R M 385 709 95 482 0.197 ND 2.0

Boque Phalia @ Leland, MS SMC 1995 Aug R M 383 685 147 372 0.395 ND 2.0

Brushy Cr Res nr N Vernon, IN NMC 1995 Aug R F 855 8959 2572 1540 1.670 22.700 2.0

Brushy Cr Res nr N Vernon, IN NMC 1995 Aug R F 790 6577 1337 1050 1.273 17.091 2.0

Brushy Cr Res nr N Vernon, IN NMC 1995 Aug R F 667 3952 3833 1890 2.028 14.032 2.0

Brushy Cr Res nr N Vernon, IN NMC 1995 Aug R F 662 3175 1849 1101 1.679 18.749 3.0

Brushy Cr Res nr N Vernon, IN NMC 1995 Aug R F 622 2887 3187 627 5.083 12.911 3.0

Brushy Cr Res nr N Vernon, IN NMC 1995 Aug R F 558 3028 2208 248 8.903 19.131 3.0
Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Brushy Cr Res nr N Vernon, IN NMC 1995 Aug R F 557 2411 2503 672 3.725 13.777 3.0

Brushy Cr Res nr N Vernon, IN NMC 1995 Aug R F 555 2372 2733 728 3.754 20.559 3.0

Brushy Cr Res nr N Vernon, IN NMC 1995 Aug R F 551 1941 2620 744 3.522 19.590 3.0

Brushy Cr Res nr N Vernon, IN NMC 1995 Aug R F 510 1610 828 328 2.524 18.952 3.0

Brushy Cr Res nr N Vernon, IN NMC 1995 Aug R M 722 6407 218 831 0.262 ND 3.0

Brushy Cr Res nr N Vernon, IN NMC 1995 Aug R M 664 3981 272 553 0.492 ND 3.0

Brushy Cr Res nr N Vernon, IN NMC 1995 Aug R M 607 2818 50 402 0.124 ND 3.0

Brushy Cr Res nr N Vernon, IN NMC 1995 Aug R M 558 2495 226 614 0.368 ND 3.0

Brushy Cr Res nr N Vernon, IN NMC 1995 Aug R M 557 2515 173 988 0.175 ND 3.0

Brushy Cr Res nr N Vernon, IN NMC 1995 Aug R M 557 2414 364 716 0.508 ND 3.0

Brushy Cr Res nr N Vernon, IN NMC 1995 Aug R M 552 1960 321 1783 0.180 ND 3.0

Brushy Cr Res nr N Vernon, IN NMC 1995 Aug R M 551 1932 208 706 0.295 ND 3.0

Brushy Cr Res nr N Vernon, IN NMC 1995 Aug R M 511 1688 252 1961 0.129 ND 3.0

Brushy Cr Res nr N Vernon, IN NMC 1995 Aug R M 446 1229 211 665 0.317 ND 3.0

Cache Riv @ Cotton Plant, AR SMC 1995 Sept R F 550 2278 2122 99 21.43 2.155 3.0

Cache Riv @ Cotton Plant, AR SMC 1995 Sept R F 513 2018 1359 169 8.04 3.175 3.0

Cache Riv @ Cotton Plant, AR SMC 1995 Sept R F 511 1861 982 212 4.63 3.233 3.0

Cache Riv @ Cotton Plant, AR SMC 1995 Sept R F 501 1916 2220 179 12.40 4.512 3.0

Cache Riv @ Cotton Plant, AR SMC 1995 Sept R F 482 1497 5126 239 21.45 5.763 3.0

Cache Riv @ Cotton Plant, AR SMC 1995 Sept R F 400 982 1515 193 7.85 3.412 3.0

Cache Riv @ Cotton Plant, AR SMC 1995 Sept R F 340 523 821 268 3.06 5.192 2.0

Cache Riv @ Cotton Plant, AR SMC 1995 Sept R M 570 2505 1472 212 6.94 ND 2.0

Cache Riv @ Cotton Plant, AR SMC 1995 Sept R M 567 2257 61 291 0.21 ND 2.0

Cache Riv @ Cotton Plant, AR SMC 1995 Sept R M 551 2127 791 192 4.12 ND 2.0

Cache Riv @ Cotton Plant, AR SMC 1995 Sept R M 545 1987 942 164 5.74 ND 2.0

Cache Riv @ Cotton Plant, AR SMC 1995 Sept R M 490 1665 252 259 0.97 ND 2.0

Cache Riv @ Cotton Plant, AR SMC 1995 Sept R M 487 1367 267 187 1.43 ND 2.0

Cache Riv @ Cotton Plant, AR SMC 1995 Sept R M 480 1466 380 158 2.41 ND 2.0

Cache Riv @ Cotton Plant, AR SMC 1995 Sept R M 462 1715 1289 317 4.07 ND 2.0

Cache Riv @ Cotton Plant, AR SMC 1995 Sept R M 460 1189 181 175 1.03 ND 2.0

Cache Riv @ Egypt, AR SMC 1995 Sept R F 600 3472 1124 631 1.78 3.474 3.0

Cache Riv @ Egypt, AR SMC 1995 Sept R F 555 2506 1198 378 3.17 5.804 3.0

Cache Riv @ Egypt, AR SMC 1995 Sept R F 529 1992 950 406 2.34 2.574 3.0

Cache Riv @ Egypt, AR SMC 1995 Sept R F 523 1937 2148 639 3.36 3.139 3.0

Cache Riv @ Egypt, AR SMC 1995 Sept R F 521 1918 1435 549 2.61 5.804 3.0

Cache Riv @ Egypt, AR SMC 1995 Sept R F 502 1977 2112 555 3.81 2.412 3.0

Cache Riv @ Egypt, AR SMC 1995 Sept R F 499 1749 2192 1956 1.12 3.929 3.0

Cache Riv @ Egypt, AR SMC 1995 Sept R F 482 1612 2159 203 10.64 4.134 3.0

Cache Riv @ Egypt, AR SMC 1995 Sept R F 482 1695 151 361 0.42 0.421 1.0

Cache Riv @ Egypt, AR SMC 1995 Sept R F 465 1358 1180 471 2.51 3.905 3.0

Table 4  31
32  Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Cache Riv @ Egypt, AR SMC 1995 Sept R M 519 1758 101 1326 0.08 ND 2.0

Cache Riv @ Egypt, AR SMC 1995 Sept R M 506 1661 617 921 0.67 ND 2.0

Cache Riv @ Egypt, AR SMC 1995 Sept R M 498 1572 307 1520 0.20 ND 2.0

Cache Riv @ Egypt, AR SMC 1995 Sept R M 482 1430 83 109 0.76 ND 2.0

Cache Riv @ Egypt, AR SMC 1995 Sept R M 457 1186 861 1702 0.51 ND 2.0

Cache Riv @ Egypt, AR SMC 1995 Sept R M 454 1217 508 659 0.77 ND 2.0

Cache Riv @ Egypt, AR SMC 1995 Sept R M 445 1101 183 511 0.36 ND 2.0

Cache Riv @ Egypt, AR SMC 1995 Sept R M 431 1050 269 1815 0.15 ND 2.0

Cache Riv @ Egypt, AR SMC 1995 Sept R M 391 882 224 1088 0.21 ND 2.0

Cache Riv @ Egypt, AR SMC 1995 Sept R M NA NA 946 251 3.77 ND 2.0

Calville Bay, L Mead, NV SW 1997 May R F 507 1899 1694 936 1.810 1.057 3

Calville Bay, L Mead, NV SW 1997 May R F 466 1232 1342 811 1.655 0.414 2

Calville Bay, L Mead, NV SW 1997 May R F 493 1561 1400 607 2.306 1.307 3

Calville Bay, L Mead, NV SW 1997 May R F 509 2193 1327 217 6.115 1.307 3

Calville Bay, L Mead, NV SW 1997 May R F 472 1257 765 258 2.965 ND 2

Calville Bay, L Mead, NV SW 1997 May R F 449 1324 732 219 3.342 1.924 3

Calville Bay, L Mead, NV SW 1997 May R F 484 1392 1147 342 3.354 0.728 3

Calville Bay, L Mead, NV SW 1997 May R F 445 1260 1186 547 2.168 1.799 3

Calville Bay, L Mead, NV SW 1997 May R M 459 1239 303 528 0.574 ND 3

Calville Bay, L Mead, NV SW 1997 May R M 512 1885 299 744 0.402 ND 3

Calville Bay, L Mead, NV SW 1997 May R M 494 1704 236 792 0.298 0.175 3

Calville Bay, L Mead, NV SW 1997 May R M 491 1489 237 984 0.241 0.233 3

Calville Bay, L Mead, NV SW 1997 May R M 473 1338 519 1363 0.381 ND 3

Calville Bay, L Mead, NV SW 1997 May R M 443 1085 214 759 0.282 ND 3

Calville Bay, L Mead, NV SW 1997 May R M 457 1181 438 723 0.606 ND 3

Calville Bay, L Mead, NV SW 1997 May R M 511 1851 236 1015 0.233 ND 3

Calville Bay, L Mead, NV SW 1997 May R M 419 1118 136 917 0.148 ND 3

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR F 580 2550 2466 539 4.58 0.793 3

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR F 551 1950 1189 1037 1.15 0.940 3

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR F 522 1750 1258 785 1.60 0.322 3

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR F 513 1800 1368 1239 1.10 2.613 3

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR F 493 1450 1926 858 2.24 2.583 3

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR F 476 1400 1749 464 3.77 0.077 3

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR F 463 1120 741 901 0.82 0.044 0

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR F 457 1300 1366 802 1.70 1.989 3

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR F 422 1710 1306 1337 0.98 0.108 3

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR F 420 1000 1162 951 1.22 0.241 3

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR M 470 1250 948 1042 0.91 ND 2

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR M 464 1325 1957 908 2.16 ND 2

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR M 453 1100 1345 2348 0.57 ND 2
Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR M 450 1150 1718 1065 1.61 ND 2

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR M 443 1100 954 1062 0.90 ND 2

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR M 435 1050 785 673 1.17 ND 2

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR M 430 900 1448 988 1.47 ND 2

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR M 428 1000 1590 577 2.76 ND 3

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR M 425 1700 1070 1068 1.00 ND 2

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR M 356 950 464 953 0.49 ND 2

Cedar Riv @ St. Chas City, IA NMC 1995 Sept R F 570 2398 3190 135 23.63 7.045 3.0

Cedar Riv @ St. Chas City, IA NMC 1995 Sept R F 555 1935 1198 352 3.40 2.371 3.0

Cedar Riv @ St. Chas City, IA NMC 1995 Sept R F 525 2056 2205 558 3.95 3.067 3.0

Cedar Riv @ St. Chas City, IA NMC 1995 Sept R F 515 1844 1329 258 5.15 5.259 3.0

Cedar Riv @ St. Chas City, IA NMC 1995 Sept R F 495 1500 2171 346 6.27 9.531 3.0

Cedar Riv @ St. Chas City, IA NMC 1995 Sept R F 480 1282 1317 851 1.55 3.074 3.0

Cedar Riv @ St. Chas City, IA NMC 1995 Sept R F 460 1200 2163 1189 1.82 3.516 3.0

Cedar Riv @ St. Chas City, IA NMC 1995 Sept R F 460 1244 1418 759 1.87 3.084 3.0

Cedar Riv @ St. Chas City, IA NMC 1995 Sept R F 435 1068 2175 334 6.51 2.899 3.0

Cedar Riv @ St. Chas City, IA NMC 1995 Sept R F 400 811 1292 923 1.40 3.923 3.0

Cedar Riv @ St. Chas City, IA NMC 1995 Sept R M 550 1965 210 1648 0.13 ND 1.0

Cedar Riv @ St. Chas City, IA NMC 1995 Sept R M 535 1923 294 2699 0.11 ND 2.0

Cedar Riv @ St. Chas City, IA NMC 1995 Sept R M 515 1848 184 556 0.33 ND 2.0

Cedar Riv @ St. Chas City, IA NMC 1995 Sept R M 505 1424 226 589 0.38 ND 2.0

Cedar Riv @ St. Chas City, IA NMC 1995 Sept R M 495 1472 204 3034 0.07 ND 2.0

Cedar Riv @ St. Chas City, IA NMC 1995 Sept R M 470 1245 159 2664 0.06 ND 2.0

Cedar Riv @ St. Chas City, IA NMC 1995 Sept R M 455 1135 296 406 0.73 ND 2.0

Cedar Riv @ St. Chas City, IA NMC 1995 Sept R M 450 1083 156 1494 0.10 ND 2.0

Cedar Riv @ St. Chas City, IA NMC 1995 Sept R M 430 933 182 1131 0.16 ND 2.0

Cedar Riv @ St. Chas City, IA NMC 1995 Sept R M 395 740 258 572 0.45 ND 2.0

Clear Creek nr Sanger, TX SMC 1994 Dec NR F 575 2423 415 115 3.61 14.000 3.0

Clear Creek nr Sanger, TX SMC 1994 Dec NR F 555 2379 491 237 2.07 12.800 3.0

Clear Creek nr Sanger, TX SMC 1994 Dec NR F 545 2188 337 79 4.27 7.000 3.0

Clear Creek nr Sanger, TX SMC 1994 Dec NR F 522 1971 110 57 1.93 8.000 3.0

Clear Creek nr Sanger, TX SMC 1994 Dec NR F 472 1589 558 109 5.12 16.500 3.0

Clear Creek nr Sanger, TX SMC 1994 Dec NR M 525 1716 77 74 1.04 ND 1.0

Clear Creek nr Sanger, TX SMC 1994 Dec NR M 511 1782 78 96 0.81 ND 1.0

Colo R inflo L Mead @ Pearce Fy, AZ SW 1996 May R F 450 969 383 354 1.082 0.642 3.0

Colo R inflo L Mead @ Pearce Fy, AZ SW 1996 May R F 444 1071 702 425 1.652 5.816 3.0

Colo R inflo L Mead @ Pearce Fy, AZ SW 1996 May R F 443 1118 391 401 0.975 1.752 2.0

Colo R inflo L Mead @ Pearce Fy, AZ SW 1996 May R F 428 995 311 296 1.051 1.047 3.0

Colo R inflo L Mead @ Pearce Fy, AZ SW 1996 May R F 428 902 307 541 0.567 3.016 2.0

Colo R inflo L Mead @ Pearce Fy, AZ SW 1996 May R F 428 949 626 435 1.439 3.504 4.0

Table 4  33
34  Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Colo R inflo L Mead @ Pearce Fy, AZ SW 1996 May R F 411 995 402 596 0.674 3.129 3.0

Colo R inflo L Mead @ Pearce Fy, AZ SW 1996 May R F 402 834 187 171 1.094 3.096 3.0

Colo R inflo L Mead @ Pearce Fy, AZ SW 1996 May R F 401 794 290 381 0.761 0.317 3.0

Colo R inflo L Mead @ Pearce Fy, AZ SW 1996 May R F 389 907 904 857 1.055 2.341 4.0

Colo R inflo L Mead @ Pearce Fy, AZ SW 1996 May R M 490 1294 95 255 0.373 0.678 3.0

Colo R inflo L Mead @ Pearce Fy, AZ SW 1996 May R M 449 1038 87 132 0.659 ND 2.0

Colo R inflo L Mead @ Pearce Fy, AZ SW 1996 May R M 433 793 95 279 0.341 ND 2.0

Colo R inflo L Mead @ Pearce Fy, AZ SW 1996 May R M 431 878 89 731 0.122 ND 3.0

Colo R inflo L Mead @ Pearce Fy, AZ SW 1996 May R M 428 754 107 406 0.264 ND 3.0

Colo R inflo L Mead @ Pearce Fy, AZ SW 1996 May R M 427 828 118 259 0.456 0.350 3.0

Colo R inflo L Mead @ Pearce Fy, AZ SW 1996 May R M 423 899 85 154 0.552 ND 3.0

Colo R inflo L Mead @ Pearce Fy, AZ SW 1996 May R M 410 826 77 74 1.041 ND 3.0

Colo R inflo L Mead @ Pearce Fy, AZ SW 1996 May R M 408 847 89 95 0.937 ND 2.0

Colo R inflo L Mead @ Pearce Fy, AZ SW 1996 May R M 404 717 105 229 0.459 ND 3.0

Columbia Riv @ Warrendale, OR NW 1997 Nov NR M 383 819 725 1156 0.63 ND 3

Columbia Riv @ Pasco, WA NW 1997 Oct NR F 626 3750 556 178 3.12 1.731 2

Columbia Riv @ Pasco, WA NW 1997 Oct NR F 613 3100 1535 208 7.38 1.603 2

Columbia Riv @ Pasco, WA NW 1997 Oct NR F 560 2500 748 613 1.22 2.634 2

Columbia Riv @ Pasco, WA NW 1997 Oct NR F 555 2450 512 53 9.66 1.882 2

Columbia Riv @ Pasco, WA NW 1997 Oct NR F 541 2350 1407 338 4.16 2.004 2

Columbia Riv @ Pasco, WA NW 1997 Oct NR F 492 1536 1067 241 4.43 2.242 2

Columbia Riv @ Pasco, WA NW 1997 Oct NR F 490 1619 1833 351 5.22 2.986 2

Columbia Riv @ Pasco, WA NW 1997 Oct NR F 468 1462 1186 118 10.05 2.526 2

Columbia Riv @ Pasco, WA NW 1997 Oct NR F 459 1351 1524 362 4.21 1.355 2

Columbia Riv @ Pasco, WA NW 1997 Oct NR F 459 1223 1672 246 6.80 2.972 2

Columbia Riv @ Pasco, WA NW 1997 Oct NR F 439 1105 1608 260 6.18 2.317 2

Columbia Riv @ Pasco, WA NW 1997 Oct NR M 581 2200 170 1139 0.15 ND 2

Columbia Riv @ Pasco, WA NW 1997 Oct NR M 515 1600 465 820 0.57 ND 3

Columbia Riv @ Pasco, WA NW 1997 Oct NR M 515 1520 798 588 1.36 ND 2

Columbia Riv @ Pasco, WA NW 1997 Oct NR M 506 1200 381 419 0.91 ND 2

Columbia Riv @ Pasco, WA NW 1997 Oct NR M 504 1600 205 195 1.05 ND 2

Columbia Riv @ Pasco, WA NW 1997 Oct NR M 487 1287 253 1049 0.24 ND 3

Columbia Riv @ Pasco, WA NW 1997 Oct NR M 471 1250 292 432 0.68 ND 2

Columbia Riv @ Pasco, WA NW 1997 Oct NR M 467 1254 109 548 0.20 ND 3

Columbia Riv @ Pasco, WA NW 1997 Oct NR M 461 1350 172 776 0.22 ND 3

Columbia Riv @ Vernita Bg, WA NW 1997 Oct NR F 661 3800 102 119 0.86 0.476 2

Columbia Riv @ Vernita Bg, WA NW 1997 Oct NR F 659 3600 184 212 0.87 0.714 2

Columbia Riv @ Vernita Bg, WA NW 1997 Oct NR F 645 4250 320 238 1.34 1.614 2

Columbia Riv @ Vernita Bg, WA NW 1997 Oct NR F 645 3400 711 548 1.30 1.319 2

Columbia Riv @ Vernita Bg, WA NW 1997 Oct NR F 641 3100 610 405 1.51 0.126 1
Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Columbia Riv @ Vernita Bg, WA NW 1997 Oct NR F 630 4550 832 231 3.60 1.701 2

Columbia Riv @ Vernita Bg, WA NW 1997 Oct NR F 604 3000 261 51 5.12 2.327 2

Columbia Riv @ Vernita Bg, WA NW 1997 Oct NR F 558 2400 652 205 3.18 1.340 2

Columbia Riv @ Vernita Bg, WA NW 1997 Oct NR F 548 2300 594 481 1.23 2.259 2

Columbia Riv @ Vernita Bg, WA NW 1997 Oct NR F 475 1376 855 131 6.53 1.954 2

Columbia Riv @ Vernita Bg, WA NW 1997 Oct NR M 667 2950 408 335 1.22 ND 3

Columbia Riv @ Vernita Bg, WA NW 1997 Oct NR M 617 2950 225 103 2.18 ND 2

Columbia Riv @ Vernita Bg, WA NW 1997 Oct NR M 610 2800 432 329 1.31 ND 3

Columbia Riv @ Vernita Bg, WA NW 1997 Oct NR M 575 2100 102 383 0.27 ND 3

Columbia Riv @ Vernita Bg, WA NW 1997 Oct NR M 523 1531 183 696 0.26 ND 2

Columbia Riv @ Vernita Bg, WA NW 1997 Oct NR M 516 1750 223 703 0.32 ND 2

Columbia Riv @ Vernita Bg, WA NW 1997 Oct NR M 500 1576 316 361 0.88 0.018 2

Columbia Riv @ Vernita Bg, WA NW 1997 Oct NR M 457 1246 118 695 0.17 0.007 3

Columbia Riv @ Vernita Bg, WA NW 1997 Oct NR M 440 1477 219 335 0.65 ND 2

Columbia Riv @ Vernita Bg, WA NW 1997 Oct NR M 435 1086 272 562 0.48 0.037 3

Columbia Riv @ Vernita Bg, WA NW 1997 Oct NR M 416 917 105 151 0.70 ND 2

Columbia Riv @ Warrendale, OR NW 1997 Nov NR F 696 4500 882 782 1.13 0.997 2

Columbia Riv @ Warrendale, OR NW 1997 Nov NR F 635 4250 658 663 0.99 0.390 2

Columbia Riv @ Warrendale, OR NW 1997 Nov NR F 607 2600 791 567 1.40 ND 2

Columbia Riv @ Warrendale, OR NW 1997 Nov NR F 592 2700 727 124 5.86 0.139 2

Columbia Riv @ Warrendale, OR NW 1997 Nov NR F 582 3100 518 898 0.58 0.777 2

Columbia Riv @ Warrendale, OR NW 1997 Nov NR F 550 2400 1031 468 2.20 0.191 2

Columbia Riv @ Warrendale, OR NW 1997 Nov NR F 541 2250 416 486 0.86 0.367 2

Columbia Riv @ Warrendale, OR NW 1997 Nov NR F 520 1948 88 58 1.52 0.349 2

Columbia Riv @ Warrendale, OR NW 1997 Nov NR F 511 1821 661 149 4.44 0.120 3

Columbia Riv @ Warrendale, OR NW 1997 Nov NR F 500 1670 550 115 4.78 0.305 2

Columbia Riv @ Warrendale, OR NW 1997 Nov NR M 625 3400 118 396 0.30 ND 3

Columbia Riv @ Warrendale, OR NW 1997 Nov NR M 582 2600 197 126 1.56 0.008 3

Columbia Riv @ Warrendale, OR NW 1997 Nov NR M 539 2400 196 189 1.04 0.010 3

Columbia Riv @ Warrendale, OR NW 1997 Nov NR M 520 1878 119 294 0.40 0.035 3

Columbia Riv @ Warrendale, OR NW 1997 Nov NR M 517 1918 214 336 0.64 0.009 3

Columbia Riv @ Warrendale, OR NW 1997 Nov NR M 515 1679 1117 311 3.59 ND 3

Columbia Riv @ Warrendale, OR NW 1997 Nov NR M 495 1548 588 911 0.65 0.011 3

Columbia Riv @ Warrendale, OR NW 1997 Nov NR M 490 1736 291 335 0.87 0.007 3

Columbia Riv @ Warrendale, OR NW 1997 Nov NR M 478 1630 274 349 0.79 ND 3

Conn Riv @ Thompsonville, CT NE 1994 Sept NR F 810 8286 4605 2740 1.68 36.000 3.0

Conn Riv @ Thompsonville, CT NE 1994 Sept NR F 765 7264 4358 1621 2.69 38.000 4.0

Conn Riv @ Thompsonville, CT NE 1994 Sept NR F 764 8172 7703 1663 4.63 26.600 4.0

Conn Riv @ Thompsonville, CT NE 1994 Sept NR F 745 7037 1646 769 2.14 ND 3.0

Conn Riv @ Thompsonville, CT NE 1994 Sept NR F 740 5902 2729 1962 1.39 25.600 3.0

Table 4  35
36  Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Conn Riv @ Thompsonville, CT NE 1994 Sept NR F 738 7037 2564 1532 1.67 38.000 4.0

Conn Riv @ Thompsonville, CT NE 1994 Sept NR F 732 6356 1982 1368 1.45 21.300 3.0

Conn Riv @ Thompsonville, CT NE 1994 Sept NR F 650 7037 1996 924 2.16 38.300 3.0

Conn Riv @ Thompsonville, CT NE 1994 Sept NR F 618 2423 2505 1537 1.63 37.700 3.0

Conn Riv @ Thompsonville, CT NE 1994 Sept NR M 690 6356 923 2161 0.43 ND 3.0

Conn Riv @ Thompsonville, CT NE 1994 Sept NR M 670 5902 995 1271 0.78 ND 3.0

Conn Riv @ Thompsonville, CT NE 1994 Sept NR M 652 4313 556 1043 0.53 ND 3.0

Conn Riv @ Thompsonville, CT NE 1994 Sept NR M 648 878 610 1937 0.31 ND 3.0

Conn Riv @ Thompsonville, CT NE 1994 Sept NR M 638 4540 869 1462 0.59 ND 3.0

Conn Riv @ Thompsonville, CT NE 1994 Sept NR M 632 4086 857 1593 0.54 ND 3.0

Conn Riv @ Thompsonville, CT NE 1994 Sept NR M 610 4313 1120 2109 0.53 ND 3.0

Conn Riv @ Thompsonville, CT NE 1994 Sept NR M 582 2918 1227 1394 0.88 ND 3.0

Conn Riv @ Thompsonville, CT NE 1994 Sept NR M 420 1167 327 803 0.41 ND 3.0

Conn Riv @ Thompsonville, CT NE 1994 Sept NR M 383 873 114 629 0.18 ND 2.0

Conn Riv @ Thompsonville, CT NE 1994 Sept NR M 339 601 586 442 1.33 ND 3.0

Cumberland Riv @ Clarksville, TN SMC 1995 Oct NR F 545 2400 554 69 5.91 2.304 3

Cumberland Riv @ Clarksville, TN SMC 1995 Oct NR F 545 1968 545 119 4.58 3.113 3

Cumberland Riv @ Clarksville, TN SMC 1995 Oct NR M 570 2400 160 1164 0.14 ND 2

Cumberland Riv @ Clarksville, TN SMC 1995 Oct NR M 505 1729 331 137 2.42 0.025 2

Des Moines Riv @ Keosauqua, IA NMC 1995 Oct NR M 360 667 567 4301 0.13 ND 2

Des Moines Riv @ Keosauqua, IA NMC 1995 Oct NR F 605 2900 557 135 4.13 1.861 3

Des Moines Riv @ Keosauqua, IA NMC 1995 Oct NR F 600 2900 541 224 2.42 1.317 3

Des Moines Riv @ Keosauqua, IA NMC 1995 Oct NR F 555 2500 850 157 5.41 4.055 3

Des Moines Riv @ Keosauqua, IA NMC 1995 Oct NR F 548 2350 517 446 1.16 1.900 3

Des Moines Riv @ Keosauqua, IA NMC 1995 Oct NR F 547 2050 652 282 2.31 0.989 3

Des Moines Riv @ Keosauqua, IA NMC 1995 Oct NR F 541 2100 1220 522 2.34 3.673 3

Des Moines Riv @ Keosauqua, IA NMC 1995 Oct NR F 536 2400 488 163 2.99 1.967 3

Des Moines Riv @ Keosauqua, IA NMC 1995 Oct NR F 497 1571 556 429 1.30 0.474 3

Des Moines Riv @ Keosauqua, IA NMC 1995 Oct NR F 491 1555 673 305 2.21 5.482 3

Des Moines Riv @ Keosauqua, IA NMC 1995 Oct NR F 462 1310 490 132 3.71 0.045 1

Des Moines Riv @ Keosauqua, IA NMC 1995 Oct NR M 528 1756 498 2490 0.20 ND 2

Des Moines Riv @ Keosauqua, IA NMC 1995 Oct NR M 522 1760 467 1613 0.29 ND 2

Des Moines Riv @ Keosauqua, IA NMC 1995 Oct NR M 513 1563 533 2518 0.21 ND 2

Des Moines Riv @ Keosauqua, IA NMC 1995 Oct NR M 511 1610 567 1596 0.36 ND 1.5

Des Moines Riv @ Keosauqua, IA NMC 1995 Oct NR M 504 1556 441 1780 0.25 ND 1.5

Des Moines Riv @ Keosauqua, IA NMC 1995 Oct NR M 480 1317 538 3068 0.18 ND 2

Des Moines Riv @ Keosauqua, IA NMC 1995 Oct NR M 445 1115 452 3936 0.11 ND 2

Des Moines Riv @ Keosauqua, IA NMC 1995 Oct NR M 369 777 459 4001 0.11 ND 2

Des Moines Riv @ Keosauqua, IA NMC 1995 Oct NR M 365 566 525 3611 0.15 ND 2.5

Don Pedro Res nr La Grange, CA SW 1994 Oct NR F 766 5176 2929 1416 2.07 26.000 3.0
Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Don Pedro Res nr La Grange, CA SW 1994 Oct NR F 485 1536 5922 1854 3.19 30.400 3.0

Don Pedro Res nr La Grange, CA SW 1994 Oct NR F 484 1487 3121 2168 1.44 38.000 3.0

Don Pedro Res nr La Grange, CA SW 1994 Oct NR F 484 1398 3532 1691 2.09 22.900 3.0

Don Pedro Res nr La Grange, CA SW 1994 Oct NR F 457 1348 3623 1757 2.06 30.700 3.0

Don Pedro Res nr La Grange, CA SW 1994 Oct NR F 454 1310 6398 2851 2.24 34.900 3.0

Don Pedro Res nr La Grange, CA SW 1994 Oct NR F 437 1082 3156 2868 1.10 30.000 3.0

Don Pedro Res nr La Grange, CA SW 1994 Oct NR F 430 1066 4553 2542 1.79 41.300 3.0

Don Pedro Res nr La Grange, CA SW 1994 Oct NR F 415 1023 4344 2470 1.76 31.000 3.0

Don Pedro Res nr La Grange, CA SW 1994 Oct NR F 410 943 2493 1048 2.38 2.300 1.0

Don Pedro Res nr La Grange, CA SW 1994 Oct NR M 526 1853 2253 4141 0.54 ND 3.0

Don Pedro Res nr La Grange, CA SW 1994 Oct NR M 470 1448 2412 4421 0.55 ND 3.0

Don Pedro Res nr La Grange, CA SW 1994 Oct NR M 455 1272 2319 4478 0.52 ND 3.0

Don Pedro Res nr La Grange, CA SW 1994 Oct NR M 450 1294 2066 5009 0.41 ND 3.0

Don Pedro Res nr La Grange, CA SW 1994 Oct NR M 445 1127 1338 3589 0.37 ND 3.0

Don Pedro Res nr La Grange, CA SW 1994 Oct NR M 440 1264 2220 4722 0.47 ND 3.0

Don Pedro Res nr La Grange, CA SW 1994 Oct NR M 434 1183 2123 4121 0.52 ND 3.0

Don Pedro Res nr La Grange, CA SW 1994 Oct NR M 430 1153 2542 3882 0.65 0.010 3.0

Don Pedro Res nr La Grange, CA SW 1994 Oct NR M 426 1129 1376 3945 0.35 ND 3.0

Don Pedro Res nr La Grange, CA SW 1994 Oct NR M 425 1051 2218 3702 0.60 0.120 3.0

Don Pedro Res nr La Grange, CA SW 1994 Oct NR M 423 1009 2219 4978 0.45 ND 3.0

Don Pedro Res nr La Grange, CA SW 1994 Oct NR M 416 1009 1950 4476 0.44 ND 3.0

Don Pedro Res nr La Grange, CA SW 1995 Oct NR F 733 4638 1032 405 2.55 19.166 3.0

Don Pedro Res nr La Grange, CA SW 1995 Oct NR F 671 3676 1809 1020 1.77 20.444 3.0

Don Pedro Res nr La Grange, CA SW 1995 Oct NR F 662 3769 1112 396 2.81 28.680 3.0

Don Pedro Res nr La Grange, CA SW 1995 Oct NR F 648 3848 2397 701 3.42 19.295 3.0

Don Pedro Res nr La Grange, CA SW 1995 Oct NR F 642 3592 1232 646 1.91 23.568 3.0

Don Pedro Res nr La Grange, CA SW 1995 Oct NR F 581 2760 2437 968 2.52 36.916 3.0

Don Pedro Res nr La Grange, CA SW 1995 Oct NR F 579 2734 1381 477 2.90 21.310 3.0

Don Pedro Res nr La Grange, CA SW 1995 Oct NR F 571 1990 614 245 2.51 24.846 3.0

Don Pedro Res nr La Grange, CA SW 1995 Oct NR F 560 2478 2289 215 10.65 108.266 3.0

Don Pedro Res nr La Grange, CA SW 1995 Oct NR F 555 2424 1505 354 4.25 33.650 3.0

Don Pedro Res nr La Grange, CA SW 1995 Oct NR F 554 2498 1443 235 6.14 44.584 3.0

Don Pedro Res nr La Grange, CA SW 1995 Oct NR F 540 2311 1398 615 2.27 30.526 3.0

Don Pedro Res nr La Grange, CA SW 1995 Oct NR M 566 2310 227 1553 0.15 ND 3.0

Don Pedro Res nr La Grange, CA SW 1995 Oct NR M 535 2088 267 744 0.36 ND 3.0

Don Pedro Res nr La Grange, CA SW 1995 Oct NR M 535 2132 132 705 0.19 ND 3.0

Don Pedro Res nr La Grange, CA SW 1995 Oct NR M 532 2054 167 682 0.24 ND 2.0

Don Pedro Res nr La Grange, CA SW 1995 Oct NR M 525 2117 422 615 0.69 ND 2.0

Don Pedro Res nr La Grange, CA SW 1995 Oct NR M 515 2098 88 152 0.58 ND 2.0

Don Pedro Res nr La Grange, CA SW 1995 Oct NR M 511 1882 152 1394 0.11 ND 2.0

Table 4  37
38  Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Don Pedro Res nr La Grange, CA SW 1995 Oct NR M 481 1475 260 2885 0.09 ND 3.0

Hudson Riv @ L Lazurne, NY NE 1994 Sept NR F 727 6200 3964 920 4.31 20.000 3.0

Hudson Riv @ L Lazurne, NY NE 1994 Sept NR F 583 3497 5780 261 22.15 29.000 3.0

Hudson Riv @ L Lazurne, NY NE 1994 Sept NR F 580 3174 1002 217 4.62 23.000 3.0

Hudson Riv @ L Lazurne, NY NE 1994 Sept NR F 515 2047 1372 272 5.04 24.000 3.0

Hudson Riv @ L Lazurne, NY NE 1994 Sept NR F 503 1922 1281 346 3.70 22.000 3.0

Hudson Riv @ L Lazurne, NY NE 1994 Sept NR F 500 2753 2216 113 19.61 17.000 3.0

Hudson Riv @ L Lazurne, NY NE 1994 Sept NR F 487 1569 3244 220 14.75 8.000 3.0

Hudson Riv @ L Lazurne, NY NE 1994 Sept NR M 674 5500 889 913 0.97 ND 3.0

Hudson Riv @ L Lazurne, NY NE 1994 Sept NR M 648 4007 409 578 0.71 ND 3.0

Hudson Riv @ L Lazurne, NY NE 1994 Sept NR M 645 3667 76 319 0.24 ND 3.0

Hudson Riv @ L Lazurne, NY NE 1994 Sept NR M 622 3743 446 510 0.87 ND 3.0

Hudson Riv @ L Lazurne, NY NE 1994 Sept NR M 601 3322 468 1211 0.39 0.240 2.0

Hudson Riv @ L Lazurne, NY NE 1994 Sept NR M 600 3150 1070 1426 0.75 0.020 3.0

Hudson Riv @ L Lazurne, NY NE 1994 Sept NR M 584 2554 1103 1638 0.67 ND 3.0

Hudson Riv @ L Lazurne, NY NE 1994 Sept NR M 575 2408 405 528 0.77 ND 3.0

Hudson Riv @ L Lazurne, NY NE 1994 Sept NR M 554 2436 742 1180 0.63 ND 3.0

Hudson Riv @ L Lazurne, NY NE 1994 Sept NR M 553 3367 654 729 0.90 ND 3.0

Hudson Riv @ L Lazurne, NY NE 1994 Sept NR M 532 2097 339 651 0.52 0.150 3.0

Hudson Riv @ L Lazurne, NY NE 1994 Sept NR M 530 1919 182 254 0.72 0.160 3.0

Hudson Riv @ L Lazurne, NY NE 1994 Sept NR M 518 2334 72 1410 0.05 0.090 2.0

Hudson Riv @ L Lazurne, NY NE 1994 Sept NR M 508 1716 277 659 0.42 0.500 3.0

Hudson Riv @ L Lazurne, NY NE 1994 Sept NR M 495 1356 1183 1081 1.09 0.080 3.0

Hudson Riv @ L Lazurne, NY NE 1994 Sept NR M 490 1535 253 458 0.55 0.380 2.0

Hudson Riv @ L Lazurne, NY NE 1994 Sept NR M 490 1519 623 995 0.63 0.180 2.0

Hudson Riv @ L Lazurne, NY NE 1994 Sept NR M 488 1707 93 1125 0.08 0.570 3.0

Hudson Riv @ L Lazurne, NY NE 1994 Sept NR M 475 1591 323 626 0.52 ND 3.0

Hudson Riv @ L Lazurne, NY NE 1995 May R F 610 3700 433 167 2.593 87.793 3.0

Hudson Riv @ L Lazurne, NY NE 1995 May R F 585 3200 943 944 0.999 42.469 3.0

Hudson Riv @ L Lazurne, NY NE 1995 May R F 529 2070 673 1286 0.523 40.360 3.0

Hudson Riv @ L Lazurne, NY NE 1995 May R M 702 5000 846 1245 0.680 ND 2.0

Hudson Riv @ L Lazurne, NY NE 1995 May R M 684 5150 760 1281 0.593 ND 3.0

Hudson Riv @ L Lazurne, NY NE 1995 May R M 662 4650 699 1811 0.386 ND 3.0

Hudson Riv @ L Lazurne, NY NE 1995 May R M 636 4100 759 784 0.968 ND 3.0

Hudson Riv @ L Lazurne, NY NE 1995 May R M 615 3920 644 468 1.376 ND 3.0

Hudson Riv @ L Lazurne, NY NE 1995 May R M 600 3300 805 483 1.667 ND 3.0

Hudson Riv @ L Lazurne, NY NE 1995 May R M 587 2950 1067 322 3.314 ND 3.0

Hudson Riv @ L Lazurne, NY NE 1995 May R M 568 2550 634 426 1.488 ND 2.0

Hudson Riv @ L Lazurne, NY NE 1995 May R M 564 2550 817 222 3.680 ND 2.0

Hudson Riv @ L Lazurne, NY NE 1995 May R M 534 1950 1786 1378 1.296 ND 1.0
Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Hudson Riv @ L Lazurne, NY NE 1995 May R M 531 2100 839 479 1.752 ND 2.0

Hudson Riv @ L Lazurne, NY NE 1995 May R M 530 2100 909 807 1.126 ND 2.0

Hudson Riv @ L Lazurne, NY NE 1995 May R M 497 1800 427 584 0.731 ND 3.0

Hudson Riv @ L Lazurne, NY NE 1995 May R M 497 1750 885 765 1.157 ND 3.0

Hudson Riv @ L Lazurne, NY NE 1995 May R M 494 1650 432 753 0.574 ND 3.0

Hudson Riv @ L Lazurne, NY NE 1995 May R M 480 1650 832 949 0.877 ND 2.0

Hudson Riv @ L Lazurne, NY NE 1995 May R M 473 1375 924 836 1.105 ND 3.0

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR F 920 10000 3696 1702 2.17 19.000 3.0

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR F 762 7500 4426 1594 2.78 26.000 3.0

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR F 693 4858 3773 1007 3.75 33.900 3.0

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR F 623 3650 3626 1391 2.61 39.600 3.0

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR F 616 3737 2429 318 7.64 35.000 3.0

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR F 598 2492 435 304 1.43 31.000 3.0

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR F 578 3045 1756 916 1.92 43.500 3.0

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR F 574 2685 4499 1231 3.65 35.800 3.0

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR F 557 2407 3364 1168 2.88 35.600 3.0

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR F 554 2590 4476 1640 2.73 50.200 3.0

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR F 513 2175 4141 846 4.89 31.000 3.0

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR M 673 5352 37 1602 0.02 ND 3.0

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR M 667 4521 105 1806 0.06 ND 3.0

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR M 666 4623 105 1996 0.05 ND 3.0

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR M 664 4278 183 1145 0.16 ND 3.0

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR M 642 3449 158 1435 0.11 ND 3.0

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR M 631 3998 186 2144 0.09 ND 3.0

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR M 628 3797 186 1671 0.11 ND 3.0

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR M 615 3498 857 1994 0.43 ND 3.0

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR M 608 3627 503 3012 0.17 ND 3.0

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR M 603 2786 29 565 0.05 0.020 3.0

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR M 602 3181 195 1381 0.14 0.040 3.0

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR M 587 3080 178 1809 0.10 ND 3.0

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR M 578 2913 59 1049 0.06 ND 3.0

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR M 577 2960 161 1120 0.14 ND 3.0

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR M 573 3139 213 2128 0.10 ND 3.0

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR M 560 2607 262 1754 0.15 ND 3.0

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR M 545 2538 160 2572 0.06 ND 3.0

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR M 508 1946 304 2528 0.12 ND 3.0

Hudson Riv @ Poughkeepsie, NY NE 1995 May R F 735 6100 1334 1868 0.714 45.034 3.0

Hudson Riv @ Poughkeepsie, NY NE 1995 May R F 729 6300 1390 485 2.866 89.456 3.0

Hudson Riv @ Poughkeepsie, NY NE 1995 May R F 692 6200 1286 715 1.799 35.256 3.0

Hudson Riv @ Poughkeepsie, NY NE 1995 May R F 685 6100 467 199 2.347 ND 3.0

Table 4  39
40 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Hudson Riv @ Poughkeepsie, NY NE 1995 May R F 685 5100 1839 1589 1.157 40.491 3.0

Hudson Riv @ Poughkeepsie, NY NE 1995 May R F 682 5400 1787 922 1.938 43.649 3.0

Hudson Riv @ Poughkeepsie, NY NE 1995 May R F 679 4600 3662 312 11.737 38.154 3.0

Hudson Riv @ Poughkeepsie, NY NE 1995 May R F 651 4500 1646 1235 1.333 78.525 3.0

Hudson Riv @ Poughkeepsie, NY NE 1995 May R F 624 3900 3182 1431 2.224 45.337 3.0

Hudson Riv @ Poughkeepsie, NY NE 1995 May R F 613 3500 1231 902 1.365 30.064 3.0

Hudson Riv @ Poughkeepsie, NY NE 1995 May R F 609 3700 1257 541 2.323 29.717 3.0

Hudson Riv @ Poughkeepsie, NY NE 1995 May R F 598 2800 1384 303 4.568 49.750 3.0

Hudson Riv @ Poughkeepsie, NY NE 1995 May R F 576 3100 1917 2554 0.751 47.024 3.0

Hudson Riv @ Poughkeepsie, NY NE 1995 May R M 763 7300 1331 231 5.762 ND 3.0

Hudson Riv @ Poughkeepsie, NY NE 1995 May R M 740 7100 160 714 0.224 0.202 3.0

Hudson Riv @ Poughkeepsie, NY NE 1995 May R M 713 6000 1701 1002 1.698 ND 3.0

Hudson Riv @ Poughkeepsie, NY NE 1995 May R M 705 5600 1370 2313 0.592 ND 1.0

Hudson Riv @ Poughkeepsie, NY NE 1995 May R M 665 4700 1403 947 1.482 ND 3.0

Hudson Riv @ Poughkeepsie, NY NE 1995 May R M 657 4100 191 1375 0.139 ND 1.0

Hudson Riv @ Poughkeepsie, NY NE 1995 May R M 654 4000 378 2658 0.142 ND 3.0

Hudson Riv @ Poughkeepsie, NY NE 1995 May R M 651 4200 253 2380 0.106 ND 3.0

Hudson Riv @ Poughkeepsie, NY NE 1995 May R M 625 3400 89 1838 0.048 ND 2.0

Hudson Riv @ Poughkeepsie, NY NE 1995 May R M 624 4800 476 2243 0.212 ND 1.0

Hudson Riv @ Poughkeepsie, NY NE 1995 May R M 592 3300 310 205 1.512 ND 2.0

Hudson Riv @ Poughkeepsie, NY NE 1995 May R M 542 2600 2565 1047 2.450 ND 2.0

Hudson Riv @ Poughkeepsie, NY NE 1995 May R M 496 2500 1621 4871 0.333 ND 2.0

Illinois Riv @ Beardstown, IL NMC 1995 Sept R F 529 1864 633 541 1.17 6.344 3

Illinois Riv @ Beardstown, IL NMC 1995 Sept R F 466 1421 767 846 0.91 3.885 3

Illinois Riv @ Beardstown, IL NMC 1995 Sept R F 443 1214 171 218 0.78 6.264 3

Illinois Riv @ Beardstown, IL NMC 1995 Sept R F 436 1368 390 547 0.71 3.129 3

Illinois Riv @ Beardstown, IL NMC 1995 Sept R F 411 819 1051 382 2.75 4.557 3

Illinois Riv @ Beardstown, IL NMC 1995 Sept R F 403 797 355 671 0.53 4.513 3

Illinois Riv @ Beardstown, IL NMC 1995 Sept R F 399 877 809 270 3.00 2.316 3

Illinois Riv @ Beardstown, IL NMC 1995 Sept R F 387 811 221 101 2.19 3.453 3

Illinois Riv @ Beardstown, IL NMC 1995 Sept R F 373 745 738 757 0.97 2.477 3

Illinois Riv @ Beardstown, IL NMC 1995 Sept R F 366 616 346 378 0.92 3.853 3

Illinois Riv @ Beardstown, IL NMC 1995 Sept R M 441 956 393 1948 0.20 ND 2

Illinois Riv @ Beardstown, IL NMC 1995 Sept R M 434 961 286 5401 0.05 ND 1

Illinois Riv @ Beardstown, IL NMC 1995 Sept R M 428 922 316 1389 0.23 ND 1

Illinois Riv @ Beardstown, IL NMC 1995 Sept R M 428 865 254 857 0.30 ND 1

Illinois Riv @ Beardstown, IL NMC 1995 Sept R M 397 684 291 605 0.48 ND 1

Illinois Riv @ Beardstown, IL NMC 1995 Sept R M 392 734 367 113 3.25 ND 3

Illinois Riv @ Beardstown, IL NMC 1995 Sept R M 390 754 442 1090 0.41 ND 1

Illinois Riv @ Beardstown, IL NMC 1995 Sept R M 381 833 279 1887 0.15 ND 1
Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Illinois Riv @ Beardstown, IL NMC 1995 Sept R M 379 721 220 1046 0.21 ND 1

Illinois Riv @ Beardstown, IL NMC 1995 Sept R M 376 629 246 881 0.28 ND 1

Iowa Riv @ Rowan, IA NMC 1995 Sept R F 595 3102 2681 944 2.84 5.685 3.0

Iowa Riv @ Rowan, IA NMC 1995 Sept R F 592 2984 2113 773 2.73 3.184 3.0

Iowa Riv @ Rowan, IA NMC 1995 Sept R F 590 2960 912 651 1.40 2.697 3.0

Iowa Riv @ Rowan, IA NMC 1995 Sept R F 570 2753 1219 844 1.44 3.454 3.0

Iowa Riv @ Rowan, IA NMC 1995 Sept R F 470 1444 143 1155 0.12 6.930 3.0

Iowa Riv @ Rowan, IA NMC 1995 Sept R F 458 1388 1472 830 1.77 3.567 3.0

Iowa Riv @ Rowan, IA NMC 1995 Sept R F 455 1495 1092 640 1.71 4.528 3.0

Iowa Riv @ Rowan, IA NMC 1995 Sept R F 435 1192 3377 1093 3.09 5.139 3.0

Iowa Riv @ Rowan, IA NMC 1995 Sept R F 432 970 651 980 0.66 ND 2.0

Iowa Riv @ Rowan, IA NMC 1995 Sept R F 402 994 2117 272 7.78 2.072 3.0

Iowa Riv @ Rowan, IA NMC 1995 Sept R M 615 2720 57 310 0.18 ND 1.0

Iowa Riv @ Rowan, IA NMC 1995 Sept R M 598 2971 142 672 0.21 ND 2.0

Iowa Riv @ Rowan, IA NMC 1995 Sept R M 576 2618 151 295 0.51 ND 2.0

Iowa Riv @ Rowan, IA NMC 1995 Sept R M 563 2303 214 296 0.72 ND 2.0

Iowa Riv @ Rowan, IA NMC 1995 Sept R M 543 2119 421 75 5.61 ND 2.0

Iowa Riv @ Rowan, IA NMC 1995 Sept R M 515 1688 46 158 0.29 ND 2.0

Iowa Riv @ Rowan, IA NMC 1995 Sept R M 452 1205 401 261 1.54 ND 1.0

Iowa Riv @ Rowan, IA NMC 1995 Sept R M 451 1407 27 1872 0.01 ND 2.0

Iowa Riv @ Rowan, IA NMC 1995 Sept R M 451 1292 98 1192 0.08 ND 2.0

Iowa Riv @ Rowan, IA NMC 1995 Sept R M 440 1190 61 644 0.09 ND 2.0

Iowa Riv @ S Fk nr N Providence, IA NMC 1995 Sept R F 665 4700 819 112 7.31 3.228 3.0

Iowa Riv @ S Fk nr N Providence, IA NMC 1995 Sept R F 647 4047 1253 155 8.08 2.736 3.0

Iowa Riv @ S Fk nr N Providence, IA NMC 1995 Sept R F 624 3751 2491 484 5.15 4.159 3.0

Iowa Riv @ S Fk nr N Providence, IA NMC 1995 Sept R M 615 3512 66 69 0.96 ND 2.0

Iowa Riv @ S Fk nr N Providence, IA NMC 1995 Sept R M 602 3119 179 1391 0.13 ND 2.0

Iowa Riv @ S Fk nr N Providence, IA NMC 1995 Sept R M 560 2441 1790 717 2.50 ND 2.0

Iowa Riv @ S Fk nr N Providence, IA NMC 1995 Sept R M 528 2043 116 488 0.24 ND 2.0

Iowa Riv @ S Fk nr N Providence, IA NMC 1995 Sept R M 508 1744 575 314 1.83 ND 2.0

Iowa River nr Murengo, IA NMC 1995 Sept R F 530 1970 4819 569 8.47 3.760 3.0

Iowa River nr Murengo, IA NMC 1995 Sept R F 510 1635 1298 249 5.21 3.927 3.0

Iowa River nr Murengo, IA NMC 1995 Sept R F 500 1724 1208 378 3.20 4.899 3.0

Iowa River nr Murengo, IA NMC 1995 Sept R F 500 1694 468 268 1.75 2.952 3.0

Iowa River nr Murengo, IA NMC 1995 Sept R F 480 1476 1319 380 3.47 3.051 3.0

Iowa River nr Murengo, IA NMC 1995 Sept R F 468 1370 3267 258 12.66 5.132 3.0

Iowa River nr Murengo, IA NMC 1995 Sept R F 465 1363 1266 208 6.09 2.474 3.0

Iowa River nr Murengo, IA NMC 1995 Sept R F 435 1033 2473 314 7.88 3.631 2.0

Iowa River nr Murengo, IA NMC 1995 Sept R F 360 629 334 57 5.86 0.181 1.0

Iowa River nr Murengo, IA NMC 1995 Sept R F 360 631 1735 346 5.01 1.980 2.0

Table 4  41
42  Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Iowa River nr Murengo, IA NMC 1995 Sept R M 540 2265 307 361 0.85 ND 2.0

Iowa River nr Murengo, IA NMC 1995 Sept R M 530 1621 430 255 1.69 ND 2.0

Iowa River nr Murengo, IA NMC 1995 Sept R M 520 1985 235 263 0.89 ND 2.0

Iowa River nr Murengo, IA NMC 1995 Sept R M 507 1860 746 1348 0.55 ND 2.0

Iowa River nr Murengo, IA NMC 1995 Sept R M 505 1797 312 1253 0.25 ND 2.0

Iowa River nr Murengo, IA NMC 1995 Sept R M 505 1598 249 1251 0.20 ND 2.0

Iowa River nr Murengo, IA NMC 1995 Sept R M 410 838 1455 268 5.43 ND 2.0

Iowa River nr Murengo, IA NMC 1995 Sept R M 405 772 417 312 1.34 ND 2.0

Iowa River nr Murengo, IA NMC 1995 Sept R M 390 736 378 164 2.30 ND 2.0

Iowa River nr Murengo, IA NMC 1995 Sept R M 383 650 383 1320 0.29 ND 2.0

James Riv @ Olivet, SD NMC 1995 Sept NR F 588 3600 2112 1461 1.45 9.328 3

James Riv @ Olivet, SD NMC 1995 Sept NR F 510 1600 741 1270 0.58 3.716 2

James Riv @ Olivet, SD NMC 1995 Sept NR F 426 950 855 568 1.51 5.683 2

James Riv @ Olivet, SD NMC 1995 Sept NR F 420 1000 1299 640 2.03 6.678 3

James Riv @ Olivet, SD NMC 1995 Sept NR F 404 800 1281 463 2.77 0.390 1

James Riv @ Olivet, SD NMC 1995 Sept NR F 388 675 503 152 3.31 3.122 2

James Riv @ Olivet, SD NMC 1995 Sept NR F 384 650 907 110 8.25 0.059 1

James Riv @ Olivet, SD NMC 1995 Sept NR F 347 500 716 988 0.72 2.531 2

James Riv @ Olivet, SD NMC 1995 Sept NR F 316 400 492 1129 0.44 0.142 1

James Riv @ Olivet, SD NMC 1995 Sept NR F NA 600 461 1255 0.37 ND 2

James Riv @ Olivet, SD NMC 1995 Sept NR M 603 2775 602 1186 0.51 ND 2

James Riv @ Olivet, SD NMC 1995 Sept NR M 570 2300 1285 1230 1.04 ND 1.5

James Riv @ Olivet, SD NMC 1995 Sept NR M 541 1900 589 1203 0.49 ND 2

James Riv @ Olivet, SD NMC 1995 Sept NR M 535 1975 616 1652 0.37 0.008 1.5

James Riv @ Olivet, SD NMC 1995 Sept NR M 517 1750 1084 2805 0.39 ND 1.5

James Riv @ Olivet, SD NMC 1995 Sept NR M 509 1800 862 759 1.14 ND 1

James Riv @ Olivet, SD NMC 1995 Sept NR M 504 1875 1092 3062 0.36 ND 1.5

James Riv @ Olivet, SD NMC 1995 Sept NR M 348 525 811 256 3.17 ND 0.5

James Riv @ Olivet, SD NMC 1995 Sept NR M 319 325 446 1387 0.32 ND 1

Jordan Riv @ 17th St, UT SW 1998 Aug R F 492 1572 782 500 1.564 6.396 2

Jordan Riv @ 17th St, UT SW 1998 Aug R F 421 1110 845 276 3.062 1.149 2

Jordan Riv @ 17th St, UT SW 1998 Aug R F 421 1074 633 579 1.093 0.399 3

Jordan Riv @ 17th St, UT SW 1998 Aug R F 374 858 1296 460 2.817 0.025 3

Jordan Riv @ 17th St, UT SW 1998 Aug R F 366 702 366 268 1.366 2.163 2

Jordan Riv @ 17th St, UT SW 1998 Aug R M 570 1885 225 768 0.293 ND 3

Jordan Riv @ 17th St, UT SW 1998 Aug R M 532 2168 168 1569 0.107 ND 3

Jordan Riv @ 17th St, UT SW 1998 Aug R M 447 1195 170 347 0.490 0.023 3

Jordan Riv @ 17th St, UT SW 1998 Aug R M 378 796 399 720 0.554 0.023 3

Jordan Riv @ 17th St, UT SW 1998 Aug R M 347 762 468 477 0.981 0.043 3

Jordan Riv @ Cudahy, UT SW 1998 Aug R F 785 7581 888 822 1.080 1.057 3
Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Jordan Riv @ Cudahy, UT SW 1998 Aug R F 779 5963 316 311 1.016 0.581 3

Jordan Riv @ Cudahy, UT SW 1998 Aug R F 715 4600 1816 838 2.167 1.728 3

Jordan Riv @ Cudahy, UT SW 1998 Aug R F 710 4767 1270 466 2.725 2.198 3

Jordan Riv @ Cudahy, UT SW 1998 Aug R F 625 3397 404 385 1.049 0.926 3

Jordan Riv @ Cudahy, UT SW 1998 Aug R M 779 5700 610 678 0.900 ND 3

Jordan Riv @ Cudahy, UT SW 1998 Aug R M 694 4000 776 2324 0.334 0.019 3

Jordan Riv @ Cudahy, UT SW 1998 Aug R M 524 1725 281 329 0.854 ND 2

Jordan Riv @ Cudahy, UT SW 1998 Aug R M 405 881 690 875 0.789 ND 1

Jordan Riv @ Cudahy, UT SW 1998 Aug R M 391 870 835 1286 0.649 ND 1

Jordan Riv @ Utah L, UT SW 1998 Aug R F 560 2089 1040 364 2.857 0.825 3

Jordan Riv @ Utah L, UT SW 1998 Aug R F 560 2869 278 34 8.176 1.927 3

Jordan Riv @ Utah L, UT SW 1998 Aug R F 535 2165 1207 269 4.487 3.172 3

Jordan Riv @ Utah L, UT SW 1998 Aug R F 362 660 546 151 3.616 0.042 2

Jordan Riv @ Utah L, UT SW 1998 Aug R F 331 493 428 313 1.367 1.114 1

Jordan Riv @ Utah L, UT SW 1998 Aug R F 278 288 424 257 1.650 0.678 1

Jordan Riv @ Utah L, UT SW 1998 Aug R M 515 2591 546 660 0.827 ND 3

Jordan Riv @ Utah L, UT SW 1998 Aug R M 489 1455 527 615 0.857 ND 2

Jordan Riv @ Utah L, UT SW 1998 Aug R M 331 473 366 447 0.819 0.061 1

Jordan Riv @ Utah L, UT SW 1998 Aug R M 278 1504 504 597 0.844 0.024 2

Kansas Riv @ Bonner Sp, KS NMC 1995 Nov NR F 752 5450 853 879 0.97 1.039 3

Kansas Riv @ Bonner Sp, KS NMC 1995 Nov NR F 657 3250 979 637 1.54 ND 2

Kansas Riv @ Bonner Sp, KS NMC 1995 Nov NR F 657 4850 1177 814 1.45 1.509 3

Kansas Riv @ Bonner Sp, KS NMC 1995 Nov NR F 653 3850 1583 847 1.87 2.518 3

Kansas Riv @ Bonner Sp, KS NMC 1995 Nov NR F 628 3450 1207 750 1.61 2.044 3

Kansas Riv @ Bonner Sp, KS NMC 1995 Nov NR F 603 2800 1131 1025 1.10 3.761 3

Kansas Riv @ Bonner Sp, KS NMC 1995 Nov NR F 586 2275 611 649 0.94 ND 2

Kansas Riv @ Bonner Sp, KS NMC 1995 Nov NR F 568 2250 443 1070 0.41 1.677 3

Kansas Riv @ Bonner Sp, KS NMC 1995 Nov NR F 555 2075 614 821 0.75 1.005 1

Kansas Riv @ Bonner Sp, KS NMC 1995 Nov NR F 533 2125 739 875 0.84 3.207 3

Kansas Riv @ Bonner Sp, KS NMC 1995 Nov NR M 621 2800 979 979 1.00 ND 1

Kansas Riv @ Bonner Sp, KS NMC 1995 Nov NR M 621 2750 612 1090 0.56 ND 2

Kansas Riv @ Bonner Sp, KS NMC 1995 Nov NR M 620 2950 1445 845 1.71 ND 1.5

Kansas Riv @ Bonner Sp, KS NMC 1995 Nov NR M 596 2800 422 1034 0.41 ND 1.5

Kansas Riv @ Bonner Sp, KS NMC 1995 Nov NR M 593 2700 417 1206 0.35 ND 1.5

Kansas Riv @ Bonner Sp, KS NMC 1995 Nov NR M 565 2750 592 1124 0.53 0.013 1.5

Kansas Riv @ Bonner Sp, KS NMC 1995 Nov NR M 563 2000 417 953 0.44 ND 1

Kansas Riv @ Bonner Sp, KS NMC 1995 Nov NR M 557 2375 1027 1137 0.90 ND 2

Kansas Riv @ Bonner Sp, KS NMC 1995 Nov NR M 533 2050 466 828 0.56 ND 1.5

Kansas Riv @ Bonner Sp, KS NMC 1995 Nov NR M 460 1200 645 1026 0.63 ND 3

Lake Mojave, NV SW 1997 May R F 483 1232 1003 316 3.174 1.122 3

Table 4  43
44  Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Lake Mojave, NV SW 1997 May R F 506 1239 1401 516 2.715 3.818 3

Lake Mojave, NV SW 1997 May R F 436 1561 1373 707 1.942 3.348 3

Lake Mojave, NV SW 1997 May R F 403 1885 610 255 2.392 4.091 3

Lake Mojave, NV SW 1997 May R F 486 2193 742 193 3.845 3.063 3

Lake Mojave, NV SW 1997 May R F 474 1257 651 62 10.500 3.257 3

Lake Mojave, NV SW 1997 May R F 501 1704 727 199 3.653 2.565 2

Lake Mojave, NV SW 1997 May R F 468 1394 1307 340 3.844 2.537 3

Lake Mojave, NV SW 1997 May R F 411 1035 1286 139 9.252 0.487 3

Lake Mojave, NV SW 1997 May R F 475 1033 873 424 2.059 1.549 3

Lake Mojave, NV SW 1997 May R F 493 1830 881 56 15.732 16.124 2

Lake Mojave, NV SW 1997 May R F 512 1620 922 103 8.951 2.818 3

Lake Mojave, NV SW 1997 May R F 509 1575 1301 356 3.654 1.265 3

Lake Mojave, NV SW 1997 May R F 472 1620 1313 483 2.718 1.497 3

Lake Mojave, NV SW 1997 May R F 505 1575 424 363 1.168 6.171 3

Lake Mojave, NV SW 1997 May R F 483 1276 231 140 1.650 ND 3

Lake Mojave, NV SW 1997 May R F 434 745 913 498 1.833 2.154 3

Lake Mojave, NV SW 1997 May R F 416 1430 619 516 1.200 1.025 3

Lake Mojave, NV SW 1997 May R F 526 1339 1220 249 4.900 3.256 3

Lake Mojave, NV SW 1997 May R F 483 1495 408 300 1.360 0.141 3

Lake Mojave, NV SW 1997 May R F 506 1481 930 416 2.236 1.427 3

Lake Mojave, NV SW 1997 May R F 436 987 479 419 1.143 0.248 3

Lake Mojave, NV SW 1997 May R F 403 1357 366 234 1.564 0.016 3

Lake Mojave, NV SW 1997 May R F 486 1394 260 173 1.503 ND 3

Lake Mojave, NV SW 1997 May R M 472 1035 211 1290 0.164 ND 2

Lake Mojave, NV SW 1997 May R M 494 1033 404 2211 0.183 ND 3

Lake Mojave, NV SW 1997 May R M 491 1830 585 2494 0.235 ND 3

Lake Mojave, NV SW 1997 May R M 473 1620 418 2802 0.149 ND 3

Lake Mojave, NV SW 1997 May R M 443 1575 391 2804 0.139 ND 3

Lake Mojave, NV SW 1997 May R M 449 1232 342 1677 0.204 0.128 3

Lake Mojave, NV SW 1997 May R M 457 1239 356 1698 0.210 0.065 3

Lake Mojave, NV SW 1997 May R M 378 1561 273 1800 0.152 ND 2

Lake Mojave, NV SW 1997 May R M 435 1885 187 257 0.728 ND 3

Lake Mojave, NV SW 1997 May R M 512 2193 282 751 0.375 0.004 3

Lake Mojave, NV SW 1997 May R M 493 1257 71 263 0.270 ND 3

Lake Mojave, NV SW 1997 May R M 512 1704 281 522 0.538 ND 3

Lake Mojave, NV SW 1997 May R M 509 1232 192 1327 0.145 0.026 3

Lake Mojave, NV SW 1997 May R M 472 1239 198 652 0.304 0.034 3

Lake Mojave, NV SW 1997 May R M 434 1561 497 1692 0.294 ND 2

Lake Mojave, NV SW 1997 May R M 416 1885 100 369 0.271 ND 3

Lake Mojave, NV SW 1997 May R M 526 2193 205 680 0.301 ND 3
Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Lake Mojave, NV SW 1997 May R M 483 1257 179 970 0.185 ND 3

Lake Mojave, NV SW 1997 May R M 506 1704 163 859 0.190 ND 3

Lake Mojave, NV SW 1997 May R M 436 1232 235 934 0.252 ND 2

Lake Mojave, NV SW 1997 May R M 403 2193 363 1139 0.319 0.108 3

Lake Mojave, NV SW 1997 May R M 486 1257 265 950 0.279 ND 3

Las Vegas Bay, L Mead, NV SW 1997 May R F 465 1162 1297 749 1.732 1.047 0

Las Vegas Bay, L Mead, NV SW 1997 May R F 444 1394 541 754 0.718 1.078 3

Las Vegas Bay, L Mead, NV SW 1997 May R F 526 1830 139 527 0.264 1.069 0

Las Vegas Bay, L Mead, NV SW 1997 May R F 468 1481 1484 758 1.958 1.105 3

Las Vegas Bay, L Mead, NV SW 1997 May R F 411 987 1469 547 2.686 1.038 3

Las Vegas Bay, L Mead, NV SW 1997 May R F 475 1357 1593 917 1.737 1.548 4

Las Vegas Bay, L Mead, NV SW 1997 May R F 451 1022 679 482 1.409 0.206 1

Las Vegas Bay, L Mead, NV SW 1997 May R F 451 1344 546 219 2.493 0.804 3

Las Vegas Bay, L Mead, NV SW 1997 May R F 515 1822 1766 992 1.780 0.15 3

Las Vegas Bay, L Mead, NV SW 1997 May R M 494 1608 1199 1121 1.070 ND 3

Las Vegas Bay, L Mead, NV SW 1997 May R M 434 1035 616 366 1.683 ND 3

Las Vegas Bay, L Mead, NV SW 1997 May R M 416 1033 118 725 0.163 ND 3

Las Vegas Bay, L Mead, NV SW 1997 May R M 483 1620 372 1352 0.275 ND 3

Las Vegas Bay, L Mead, NV SW 1997 May R M 506 1575 913 1248 0.732 ND 3

Las Vegas Bay, L Mead, NV SW 1997 May R M 436 1276 151 671 0.225 ND 3

Las Vegas Bay, L Mead, NV SW 1997 May R M 403 745 459 765 0.600 ND 3

Las Vegas Bay, L Mead, NV SW 1997 May R M 486 1430 876 503 1.742 ND 3

Las Vegas Bay, L Mead, NV SW 1997 May R M 474 1339 980 609 1.609 ND 3

Las Vegas Bay, L Mead, NV SW 1997 May R M 501 1495 1298 2481 0.523 ND 3

Leetown Res @ Leetown, WV NE 1996 Oct NR F 442 1204.8 822 244 3.37 1.400 3

Leetown Res @ Leetown, WV NE 1996 Oct NR F 441 1050 904 225 4.02 ND 3

Leetown Res @ Leetown, WV NE 1996 Oct NR F 429 986.2 309 103 3.00 1.685 3

Leetown Res @ Leetown, WV NE 1996 Oct NR F 424 982.3 319 248 1.29 1.762 3

Leetown Res @ Leetown, WV NE 1996 Oct NR F 421 922.1 1160 187 6.20 2.875 3

Leetown Res @ Leetown, WV NE 1996 Oct NR F 416 908.2 602 183 3.29 2.887 3

Leetown Res @ Leetown, WV NE 1996 Oct NR F 388 797.3 1231 411 3.00 ND 3

Leetown Res @ Leetown, WV NE 1996 Oct NR F 387 798.4 543 285 1.91 1.446 3

Leetown Res @ Leetown, WV NE 1996 Oct NR F 386 742.2 622 179 3.48 ND 3

Leetown Res @ Leetown, WV NE 1996 Oct NR F 367 649.5 505 334 1.51 1.950 3

Leetown Res @ Leetown, WV NE 1996 Oct NR F 362 584.4 854 481 1.78 1.549 3

Leetown Res @ Leetown, WV NE 1996 Oct NR M 464 1129.5 419 1764 0.24 ND 2.5

Leetown Res @ Leetown, WV NE 1996 Oct NR M 448 1122 389 1366 0.29 ND 2.5

Leetown Res @ Leetown, WV NE 1996 Oct NR M 414 957.2 416 643 0.65 ND 2.5

Leetown Res @ Leetown, WV NE 1996 Oct NR M 381 677 135 489 0.28 ND 2.5

Leetown Res @ Leetown, WV NE 1996 Oct NR M 368 602.8 293 1646 0.18 ND 2.5

Table 4  45
46  Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Leetown Res @ Leetown, WV NE 1996 Oct NR M 353 560.4 223 501 0.45 ND 2.5

Leetown Res @ Leetown, WV NE 1996 Oct NR M 346 513 197 371 0.53 ND 2.5

Leetown Res @ Leetown, WV NE 1996 Oct NR M 337 462.8 335 990 0.34 ND 2.5

Little Riv Ditch @ Moorehouse, MO NMC 1995 Oct NR F 610 3474 2114 419 5.05 4.834 3.0

Little Riv Ditch @ Moorehouse, MO NMC 1995 Oct NR F 580 2571 1143 2008 0.57 1.684 3.0

Little Riv Ditch @ Moorehouse, MO NMC 1995 Oct NR F 548 2559 1401 1102 1.27 6.222 3.0

Little Riv Ditch @ Moorehouse, MO NMC 1995 Oct NR F 525 1995 1735 1083 1.60 4.464 3.0

Little Riv Ditch @ Moorehouse, MO NMC 1995 Oct NR F 524 1828 1406 868 1.62 1.455 3.0

Little Riv Ditch @ Moorehouse, MO NMC 1995 Oct NR F 510 1986 1979 286 6.92 6.870 3.0

Little Riv Ditch @ Moorehouse, MO NMC 1995 Oct NR F 510 1912 1162 1366 0.85 5.837 3.0

Little Riv Ditch @ Moorehouse, MO NMC 1995 Oct NR F 500 1793 1409 982 1.43 8.291 3.0

Little Riv Ditch @ Moorehouse, MO NMC 1995 Oct NR F 495 1865 2217 675 3.28 7.039 3.0

Little Riv Ditch @ Moorehouse, MO NMC 1995 Oct NR F 480 1519 2492 575 4.33 7.607 3.0

Little Riv Ditch @ Moorehouse, MO NMC 1995 Oct NR M 560 2467 1457 610 2.39 ND 2.0

Little Riv Ditch @ Moorehouse, MO NMC 1995 Oct NR M 530 1953 229 287 0.80 ND 3.0

Little Riv Ditch @ Moorehouse, MO NMC 1995 Oct NR M 520 1921 47 823 0.06 ND 2.0

Little Riv Ditch @ Moorehouse, MO NMC 1995 Oct NR M 510 1748 62 565 0.11 ND 2.0

Little Riv Ditch @ Moorehouse, MO NMC 1995 Oct NR M 480 1293 604 836 0.72 ND 2.0

Little Riv Ditch @ Moorehouse, MO NMC 1995 Oct NR M 475 1399 364 646 0.56 ND 2.0

Little Riv Ditch @ Moorehouse, MO NMC 1995 Oct NR M 475 1400 335 731 0.46 ND 2.0

Little Riv Ditch @ Moorehouse, MO NMC 1995 Oct NR M 470 1302 632 826 0.77 ND 2.0

Little Riv Ditch @ Moorehouse, MO NMC 1995 Oct NR M 440 1158 17 1389 0.01 ND 2.0

Little Riv Ditch @ Moorehouse, MO NMC 1995 Oct NR M 410 1256 539 1010 0.53 ND 2.0

Mississippi R @ Cape Girardeau, MO SMC 1995 Oct NR F 415 990 181 149 1.21 3.821 3

Mississippi R @ Cape Girardeau, MO SMC 1995 Oct NR F 413 915 984 285 3.45 2.132 3

Mississippi R @ Cape Girardeau, MO SMC 1995 Oct NR F 390 723 361 175 2.06 3.753 2

Mississippi R @ Cape Girardeau, MO SMC 1995 Oct NR F 375 664 520 207 2.51 2.219 3

Mississippi R @ Cape Girardeau, MO SMC 1995 Oct NR F 370 607 976 147 6.64 0.006 1

Mississippi R @ Cape Girardeau, MO SMC 1995 Oct NR F 370 575 212 246 0.86 0.058 1

Mississippi R @ Cape Girardeau, MO SMC 1995 Oct NR F 362 648 660 344 1.92 3.972 3

Mississippi R @ Cape Girardeau, MO SMC 1995 Oct NR F 360 603 505 123 4.11 3.998 2

Mississippi R @ Cape Girardeau, MO SMC 1995 Oct NR F 350 601 283 461 0.61 0.010 1

Mississippi R @ Cape Girardeau, MO SMC 1995 Oct NR F 344 304 232 124 1.87 2.476 3

Mississippi R @ Cape Girardeau, MO SMC 1995 Oct NR M 550 2009 500 3805 0.13 ND 2

Mississippi R @ Cape Girardeau, MO SMC 1995 Oct NR M 480 1678 291 2774 0.10 ND 1

Mississippi R @ Cape Girardeau, MO SMC 1995 Oct NR M 415 861 396 5991 0.07 ND 2

Mississippi R @ Cape Girardeau, MO SMC 1995 Oct NR M 413 968 268 478 0.56 ND 2

Mississippi R @ Cape Girardeau, MO SMC 1995 Oct NR M 400 858 465 1440 0.32 ND 1

Mississippi R @ Cape Girardeau, MO SMC 1995 Oct NR M 390 746 527 513 1.03 ND 2

Mississippi R @ Cape Girardeau, MO SMC 1995 Oct NR M 375 698 318 342 0.93 ND 2
Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Mississippi R @ Cape Girardeau, MO SMC 1995 Oct NR M 360 556 353 879 0.40 ND 2

Mississippi R @ Cape Girardeau, MO SMC 1995 Oct NR M 330 466 259 220 1.18 ND 2

Mississippi R @ Cape Girardeau, MO SMC 1995 Oct NR M 300 336 314 1225 0.26 ND 1

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR F 560 2500 2831 952 2.97 6.376 3

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR F 560 2200 1422 351 4.05 ND 3

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR F 540 2500 1114 606 1.84 4.767 3

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR F 540 2300 2473 491 5.04 3.488 3

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR F 540 2300 2082 717 2.90 10.725 3

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR F 510 2000 2261 935 2.42 0.270 3

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR F 510 1688 1576 1072 1.47 1.272 3

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR F 510 1783 2877 895 3.21 2.266 3

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR F 490 1809 4852 604 8.03 15.470 3

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR F 340 555 1128 513 2.20 3.778 2

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR M 570 2900 2713 1167 2.32 2.645 2

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR M 550 2100 761 1618 0.47 ND 2

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR M 530 1966 579 1097 0.53 ND 1

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR M 510 1781 1045 1838 0.57 ND 1

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR M 505 1744 598 2506 0.24 ND 2

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR M 500 1693 1081 1321 0.82 ND 2

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR M 495 1771 913 985 0.93 ND 1.5

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR M 485 1396 616 1065 0.58 ND 1.5

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR M 480 1483 908 3402 0.27 ND 2

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR M 470 1436 972 1327 0.73 ND 1.5

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR F 513 1648 1422 273 5.21 9.090 3

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR F 508 1895 639 312 2.05 4.662 3

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR F 497 1732 2799 494 5.67 2.465 3

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR F 496 1872 913 343 2.66 3.892 3

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR F 495 1685 2231 863 2.59 5.829 3

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR F 485 1458 827 556 1.49 6.049 3

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR F 470 1634 1515 836 1.81 4.534 3

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR F 465 1472 3172 224 14.16 16.201 3

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR F 455 1230 1341 325 4.13 3.546 3

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR F 441 1328 901 379 2.38 6.719 3

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR M 496 1672 555 3052 0.18 ND 1

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR M 490 1571 719 2108 0.34 ND 1

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR M 485 1487 1019 1558 0.65 ND 2

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR M 473 1391 906 1385 0.65 ND 1

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR M 470 1335 932 1561 0.60 ND 1

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR M 448 1119 345 1227 0.28 ND 1

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR M 445 1144 1020 6341 0.16 ND 1

Table 4  47
48  Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR M 442 1091 867 1482 0.59 ND 1

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR M 440 1101 479 711 0.67 ND 1

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR M 395 862 321 1418 0.23 ND 1

Mississippi Riv @ L City, MN NMC 1995 Sept NR F 600 2800 1697 785 2.16 2.953 3

Mississippi Riv @ L City, MN NMC 1995 Sept NR F 564 2200 1007 1006 1.00 4.490 3

Mississippi Riv @ L City, MN NMC 1995 Sept NR F 562 2250 1654 958 1.73 0.054 1

Mississippi Riv @ L City, MN NMC 1995 Sept NR F 542 2070 2058 1147 1.79 3.332 3

Mississippi Riv @ L City, MN NMC 1995 Sept NR F 536 2000 989 980 1.01 3.169 3

Mississippi Riv @ L City, MN NMC 1995 Sept NR F 533 2000 1859 994 1.87 11.701 3

Mississippi Riv @ L City, MN NMC 1995 Sept NR F 531 1850 797 411 1.94 0.689 2

Mississippi Riv @ L City, MN NMC 1995 Sept NR F 524 1900 907 777 1.17 6.734 3

Mississippi Riv @ L City, MN NMC 1995 Sept NR F 482 1580 1049 1221 0.86 3.787 3

Mississippi Riv @ L City, MN NMC 1995 Sept NR F 439 1100 881 421 2.09 0.157 2

Mississippi Riv @ L City, MN NMC 1995 Sept NR M 536 4600 642 1050 0.61 ND 1.5

Mississippi Riv @ L City, MN NMC 1995 Sept NR M 523 1900 1089 1241 0.88 ND 1.5

Mississippi Riv @ L City, MN NMC 1995 Sept NR M 515 2050 843 1389 0.61 ND 1.5

Mississippi Riv @ L City, MN NMC 1995 Sept NR M 511 1800 1047 1154 0.91 ND 1.5

Mississippi Riv @ L City, MN NMC 1995 Sept NR M 510 1600 827 1799 0.46 ND 1

Mississippi Riv @ L City, MN NMC 1995 Sept NR M 503 1800 1258 1915 0.66 ND 1.5

Mississippi Riv @ L City, MN NMC 1995 Sept NR M 491 1600 599 1116 0.54 ND 1

Mississippi Riv @ L City, MN NMC 1995 Sept NR M 490 1500 864 1706 0.51 ND 1

Mississippi Riv @ L City, MN NMC 1995 Sept NR M 488 1450 1008 1963 0.51 0.008 1

Mississippi Riv @ L City, MN NMC 1995 Sept NR M 470 1450 445 1235 0.36 ND 1.5

Mississippi Riv @ Luling, LA SMC 1995 Nov NR F 720 6602 907 716 1.27 0.567 1

Mississippi Riv @ Luling, LA SMC 1995 Nov NR F 680 5400 547 155 3.53 1.260 3

Mississippi Riv @ Luling, LA SMC 1995 Nov NR F 650 5300 651 298 2.18 3.244 3

Mississippi Riv @ Luling, LA SMC 1995 Nov NR F 585 3475 593 156 3.80 2.213 3

Mississippi Riv @ Luling, LA SMC 1995 Nov NR F 550 2900 631 269 2.35 1.092 3

Mississippi Riv @ Luling, LA SMC 1995 Nov NR F 505 2131 562 302 1.86 0.628 3

Mississippi Riv @ Luling, LA SMC 1995 Nov NR F 350 720 288 131 2.20 1.782 1

Mississippi Riv @ Luling, LA SMC 1995 Nov NR M 600 3325 491 590 0.83 ND 2

Mississippi Riv @ Memphis, TN SMC 1995 Oct NR F 504 2200 284 139 2.04 0.010 3

Mississippi Riv @ Memphis, TN SMC 1995 Oct NR F 500 1950 186 448 0.42 0.119 2

Mississippi Riv @ Memphis, TN SMC 1995 Oct NR F 483 1700 228 118 1.93 0.013 3

Mississippi Riv @ Memphis, TN SMC 1995 Oct NR F 480 1800 204 183 1.11 2.786 3

Mississippi Riv @ Memphis, TN SMC 1995 Oct NR F 460 1450 225 232 0.97 1.168 3

Mississippi Riv @ Memphis, TN SMC 1995 Oct NR F 455 1300 652 350 1.86 0.507 2

Mississippi Riv @ Memphis, TN SMC 1995 Oct NR F 402 1020 926 176 5.26 4.859 3

Mississippi Riv @ Memphis, TN SMC 1995 Oct NR F 400 799 280 250 1.12 0.136 0

Mississippi Riv @ Memphis, TN SMC 1995 Oct NR F 379 870 1008 139 7.25 4.834 3
Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Mississippi Riv @ Memphis, TN SMC 1995 Oct NR M 498 1650 346 305 1.13 ND 2

Mississippi Riv @ Memphis, TN SMC 1995 Oct NR M 490 1778 389 1565 0.25 ND 2

Mississippi Riv @ Memphis, TN SMC 1995 Oct NR M 458 1300 205 167 1.23 ND 2

Mississippi Riv @ Memphis, TN SMC 1995 Oct NR M 450 1355 311 378 0.82 ND 2

Mississippi Riv @ Memphis, TN SMC 1995 Oct NR M 447 1203 828 399 2.08 ND 2

Mississippi Riv @ Memphis, TN SMC 1995 Oct NR M 422 1135 219 451 0.49 0.005 2

Mississippi Riv @ Memphis, TN SMC 1995 Oct NR M 422 1200 251 172 1.46 ND 1

Mississippi Riv @ Memphis, TN SMC 1995 Oct NR M 370 711 754 551 1.37 ND 2

Missour Riveri @ Nebraska City, NE NMC 1995 Oct NR F 620 3500 789 124 6.36 1.616 3

Missour Riveri @ Nebraska City, NE NMC 1995 Oct NR F 422 975 348 1092 0.32 0.768 3

Missour Riveri @ Nebraska City, NE NMC 1995 Oct NR F 419 1050 379 231 1.64 4.993 3

Missour Riveri @ Nebraska City, NE NMC 1995 Oct NR F 418 1200 355 735 0.48 1.473 3

Missour Riveri @ Nebraska City, NE NMC 1995 Oct NR F 405 1025 287 1452 0.20 2.715 1

Missour Riveri @ Nebraska City, NE NMC 1995 Oct NR F 400 1000 298 949 0.31 0.097 1

Missour Riveri @ Nebraska City, NE NMC 1995 Oct NR F 385 850 506 133 3.80 2.435 3

Missour Riveri @ Nebraska City, NE NMC 1995 Oct NR F 384 900 404 989 0.41 0.040 3

Missour Riveri @ Nebraska City, NE NMC 1995 Oct NR F 380 850 730 152 4.80 3.349 3

Missour Riveri @ Nebraska City, NE NMC 1995 Oct NR F 375 750 632 347 1.82 1.705 3

Missour Riveri @ Nebraska City, NE NMC 1995 Oct NR F 352 775 309 1819 0.17 5.774 3

Missour Riveri @ Nebraska City, NE NMC 1995 Oct NR F 345 600 302 526 0.57 ND 2

Missour Riveri @ Nebraska City, NE NMC 1995 Oct NR M 680 3350 335 2635 0.13 ND 1

Missour Riveri @ Nebraska City, NE NMC 1995 Oct NR M 620 3450 408 1432 0.28 ND 1

Missour Riveri @ Nebraska City, NE NMC 1995 Oct NR M 575 2800 424 2476 0.17 ND 2

Missour Riveri @ Nebraska City, NE NMC 1995 Oct NR M 530 2150 283 3136 0.09 ND 1.5

Missour Riveri @ Nebraska City, NE NMC 1995 Oct NR M 472 1350 371 2298 0.16 ND 2

Missour Riveri @ Nebraska City, NE NMC 1995 Oct NR M 457 1375 359 3108 0.12 ND 1

Missour Riveri @ Nebraska City, NE NMC 1995 Oct NR M 412 975 402 1485 0.27 ND 2

Missour Riveri @ Nebraska City, NE NMC 1995 Oct NR M 370 800 290 2294 0.13 ND 1.5

Missour Riveri @ Nebraska City, NE NMC 1995 Oct NR M 365 725 296 2841 0.10 ND 1.5

Missour Riveri @ Nebraska City, NE NMC 1995 Oct NR M 363 600 482 1142 0.42 ND 1

Missour Riveri @ Nebraska City, NE NMC 1995 Oct NR M 352 625 346 1403 0.25 ND 1.5

Missouri Riv @ Garrison Dam, ND NMC 1995 Sept NR F 565 2500 403 188 2.14 2.486 1

Missouri Riv @ Garrison Dam, ND NMC 1995 Sept NR F 562 2400 299 137 2.18 3.551 3

Missouri Riv @ Garrison Dam, ND NMC 1995 Sept NR F 535 1680 502 154 3.26 0.176 3

Missouri Riv @ Garrison Dam, ND NMC 1995 Sept NR F 518 1950 246 160 1.54 3.007 3

Missouri Riv @ Garrison Dam, ND NMC 1995 Sept NR F 502 2050 418 97 4.31 2.360 3

Missouri Riv @ Garrison Dam, ND NMC 1995 Sept NR F 498 1550 342 94 3.64 0.826 1

Missouri Riv @ Garrison Dam, ND NMC 1995 Sept NR F 485 1450 490 114 4.30 7.699 2

Missouri Riv @ Garrison Dam, ND NMC 1995 Sept NR F 475 1300 322 118 2.73 4.561 3

Missouri Riv @ Garrison Dam, ND NMC 1995 Sept NR F 409 1050 227 102 2.23 0.032 2

Table 4  49
50 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Missouri Riv @ Garrison Dam, ND NMC 1995 Sept NR M 595 3400 356 887 0.40 ND 0

Missouri Riv @ Garrison Dam, ND NMC 1995 Sept NR M 541 1850 292 923 0.32 ND 1

Missouri Riv @ Garrison Dam, ND NMC 1995 Sept NR M 539 1850 323 3554 0.09 ND 1

Missouri Riv @ Garrison Dam, ND NMC 1995 Sept NR M 524 1950 219 1526 0.14 0.006 2

Missouri Riv @ Garrison Dam, ND NMC 1995 Sept NR M 522 2000 283 2143 0.13 ND 2

Missouri Riv @ Garrison Dam, ND NMC 1995 Sept NR M 495 1450 316 1181 0.27 ND 1

Missouri Riv @ Garrison Dam, ND NMC 1995 Sept NR M 488 1400 296 6760 0.04 ND 0.5

Missouri Riv @ Garrison Dam, ND NMC 1995 Sept NR M 478 1300 280 1730 0.16 0.006 1

Missouri Riv @ Garrison Dam, ND NMC 1995 Sept NR M 465 1200 323 2631 0.12 ND 1

Missouri Riv @ Garrison Dam, ND NMC 1995 Sept NR M 432 1050 362 1145 0.32 ND 1

Missouri Riv @ Garrison Dam, ND NMC 1995 Sept NR M 299 420 245 2263 0.11 0.007 1

Missouri Riv @ Hermann, MO NMC 1995 Oct NR F 640 4000 1845 397 4.65 2.588 3

Missouri Riv @ Hermann, MO NMC 1995 Oct NR F 592 2800 1325 655 2.02 1.286 3

Missouri Riv @ Hermann, MO NMC 1995 Oct NR F 472 750 1734 939 1.85 1.390 3

Missouri Riv @ Hermann, MO NMC 1995 Oct NR F 460 1250 1548 301 5.14 2.490 3

Missouri Riv @ Hermann, MO NMC 1995 Oct NR F 422 1100 1923 657 2.93 1.820 3

Missouri Riv @ Hermann, MO NMC 1995 Oct NR F 375 650 456 100 4.56 2.023 1

Missouri Riv @ Hermann, MO NMC 1995 Oct NR M 610 3050 964 2954 0.33 ND 2

Missouri Riv @ Hermann, MO NMC 1995 Oct NR M 603 2950 848 1065 0.80 ND 1.5

Missouri Riv @ Hermann, MO NMC 1995 Oct NR M 572 2500 1431 1332 1.07 ND 2

Missouri Riv @ Hermann, MO NMC 1995 Oct NR M 572 2600 413 989 0.42 ND 2

Missouri Riv @ Hermann, MO NMC 1995 Oct NR M 566 2000 838 1510 0.55 ND 2

Missouri Riv @ Hermann, MO NMC 1995 Oct NR M 550 2400 1392 72 19.33 ND 1

Missouri Riv @ Hermann, MO NMC 1995 Oct NR M 490 1650 763 2109 0.36 ND 1

Missouri Riv @ Hermann, MO NMC 1995 Oct NR M 409 1000 913 1030 0.89 ND 2.5

Missouri Riv @ Hermann, MO NMC 1995 Oct NR M 346 550 676 99 6.83 ND 1

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR F 614 3210 834 972 0.86 9.200 3.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR F 606 3516 1559 145 10.75 9.400 3.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR F 600 3311 4741 1508 3.14 9.500 3.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR F 598 3147 1544 441 3.50 7.700 3.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR F 586 3357 2512 115 21.84 26.600 3.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR F 586 3405 1425 969 1.47 15.600 3.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR F 570 2686 730 608 1.20 17.400 3.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR F 563 2834 562 435 1.29 19.900 3.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR F 560 2925 625 153 4.08 10.800 3.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR F 547 2480 594 771 0.77 13.100 3.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR F 542 2439 1062 320 3.32 12.200 3.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR F 541 2443 4347 213 20.41 6.400 3.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR F 540 2432 656 804 0.82 11.600 3.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR F 538 2322 1279 419 3.05 19.300 3.0
Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR F 497 2068 3713 377 9.85 19.600 3.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR F 486 1813 1753 1130 1.55 11.500 3.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR M 576 3037 377 638 0.59 ND 3.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR M 574 2826 449 377 1.19 ND 1.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR M 570 2601 290 978 0.30 0.100 3.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR M 561 2264 59 365 0.16 0.050 1.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR M 560 2452 172 275 0.63 0.010 3.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR M 558 2206 509 526 0.97 ND 2.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR M 557 2465 116 639 0.18 ND 2.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR M 557 2414 77 1259 0.06 0.170 2.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR M 554 2310 189 2186 0.09 ND 3.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR M 540 2160 198 1904 0.10 ND 3.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR M 539 2119 149 1653 0.09 ND 3.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR M 537 2264 114 679 0.17 ND 2.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR M 536 2245 292 610 0.48 0.060 1.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR M 536 2207 508 850 0.60 ND 3.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR M 534 2242 47 167 0.28 0.180 1.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR M 531 1989 39 1685 0.02 ND 2.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR M 530 2236 216 1071 0.20 ND 1.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR M 529 2013 30 144 0.21 ND 1.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR M 524 1886 69 319 0.22 ND 3.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR M 520 1954 91 728 0.13 ND 2.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR M 518 2011 218 431 0.51 ND 3.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR M 514 1924 166 1143 0.15 ND 2.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR M 513 1688 89 895 0.10 ND 3.0

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR M 466 1582 99 558 0.18 ND 2.0

Mohawk Riv @ Frankfort, NY NE 1995 May R F 689 5200 3375 3425 0.985 53.761 3.0

Mohawk Riv @ Frankfort, NY NE 1995 May R F 685 5300 3661 3108 1.178 15.682 3.0

Mohawk Riv @ Frankfort, NY NE 1995 May R F 620 3700 3590 3823 0.939 14.507 3.0

Mohawk Riv @ Frankfort, NY NE 1995 May R F 596 3300 4142 3797 1.091 50.535 3.0

Mohawk Riv @ Frankfort, NY NE 1995 May R F 590 3000 2818 5066 0.556 48.860 3.0

Mohawk Riv @ Frankfort, NY NE 1995 May R F 585 3200 3004 4860 0.618 14.430 3.0

Mohawk Riv @ Frankfort, NY NE 1995 May R F 580 3500 2873 2315 1.241 51.156 3.0

Mohawk Riv @ Frankfort, NY NE 1995 May R F 570 3200 1582 2704 0.585 13.348 3.0

Mohawk Riv @ Frankfort, NY NE 1995 May R F 551 2900 2014 1456 1.383 65.612 3.0

Mohawk Riv @ Frankfort, NY NE 1995 May R F 548 2500 1159 3110 0.373 32.139 3.0

Mohawk Riv @ Frankfort, NY NE 1995 May R F 547 2800 1172 4045 0.290 44.579 3.0

Mohawk Riv @ Frankfort, NY NE 1995 May R F 455 2100 3651 2291 1.594 40.484 3.0

Mohawk Riv @ Frankfort, NY NE 1995 May R M 595 3200 561 5211 0.108 ND 3.0

Mohawk Riv @ Frankfort, NY NE 1995 May R M 585 2700 521 6821 0.076 ND 3.0

Table 4 51
52 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Mohawk Riv @ Frankfort, NY NE 1995 May R M 584 2400 83 3560 0.023 ND 2.0

Mohawk Riv @ Frankfort, NY NE 1995 May R M 583 3200 414 2096 0.198 ND 3.0

Mohawk Riv @ Frankfort, NY NE 1995 May R M 579 2500 454 4287 0.106 ND 3.0

Mohawk Riv @ Frankfort, NY NE 1995 May R M 570 2600 470 5762 0.082 ND 3.0

Mohawk Riv @ Frankfort, NY NE 1995 May R M 567 2500 273 1692 0.161 ND 3.0

Mohawk Riv @ Frankfort, NY NE 1995 May R M 565 2200 369 3697 0.100 ND 3.0

Mohawk Riv @ Frankfort, NY NE 1995 May R M 560 2100 511 1409 0.363 ND 3.0

Mohawk Riv @ Frankfort, NY NE 1995 May R M 555 2500 112 2725 0.041 ND 3.0

Mohawk Riv @ Frankfort, NY NE 1995 May R M 552 2400 386 5048 0.076 ND 3.0

Mohawk Riv @ Frankfort, NY NE 1995 May R M 546 2000 662 3719 0.178 ND 2.0

Mohawk Riv @ Frankfort, NY NE 1995 May R M 545 2300 433 1642 0.264 ND 2.0

Mohawk Riv @ Frankfort, NY NE 1995 May R M 540 2500 447 1866 0.240 ND 3.0

Mohawk Riv @ Frankfort, NY NE 1995 May R M 535 2100 620 4106 0.151 ND 3.0

Mohawk Riv @ Frankfort, NY NE 1995 May R M 530 1900 388 3520 0.110 ND 3.0

Mohawk Riv @ Frankfort, NY NE 1995 May R M 445 1200 686 2001 0.343 ND 3.0

Mountain Crk Lake, Dallas, TX SMC 1995 June R M 389 661 726 1562 0.465 ND 2.0

Mountain Crk Lake, Dallas, TX SMC 1995 June R F 710 4521 1322 599 2.207 46.029 2.0

Mountain Crk Lake, Dallas, TX SMC 1995 June R F 570 1550 571 159 3.591 ND 2.0

Mountain Crk Lake, Dallas, TX SMC 1995 June R F 543 2133 1250 711 1.758 0.299 3.0

Mountain Crk Lake, Dallas, TX SMC 1995 June R F 525 1680 574 315 1.822 0.150 2.0

Mountain Crk Lake, Dallas, TX SMC 1995 June R F 505 1481 204 202 1.010 ND 2.0

Mountain Crk Lake, Dallas, TX SMC 1995 June R F 500 1789 1208 518 2.332 0.360 2.0

Mountain Crk Lake, Dallas, TX SMC 1995 June R F 490 1510 1461 290 5.038 9.150 2.0

Mountain Crk Lake, Dallas, TX SMC 1995 June R F 485 1405 1100 129 8.527 1.456 3.0

Mountain Crk Lake, Dallas, TX SMC 1995 June R F 480 1014 1239 622 1.992 0.332 3.0

Mountain Crk Lake, Dallas, TX SMC 1995 June R F 480 1352 2397 631 3.799 0.420 3.0

Mountain Crk Lake, Dallas, TX SMC 1995 June R F 470 1310 1892 569 3.325 ND 2.0

Mountain Crk Lake, Dallas, TX SMC 1995 June R F 448 1152 1501 518 2.898 ND 3.0

Mountain Crk Lake, Dallas, TX SMC 1995 June R F 435 964 119 112 1.063 0.058 2.0

Mountain Crk Lake, Dallas, TX SMC 1995 June R F 425 960 1945 831 2.341 ND 2.0

Mountain Crk Lake, Dallas, TX SMC 1995 June R F 420 2392 1458 1291 1.129 0.295 2.0

Mountain Crk Lake, Dallas, TX SMC 1995 June R F NA NA 280 133 2.105 ND 2.0

Mountain Crk Lake, Dallas, TX SMC 1995 June R M 474 1400 183 391 0.468 ND 2.0

Mountain Crk Lake, Dallas, TX SMC 1995 June R M 460 1169 158 158 1.000 ND 3.0

Mountain Crk Lake, Dallas, TX SMC 1995 June R M 455 1120 907 1945 0.466 ND 2.0

Mountain Crk Lake, Dallas, TX SMC 1995 June R M 445 1224 493 1031 0.478 ND 2.0

Mountain Crk Lake, Dallas, TX SMC 1995 June R M 435 1126 1442 311 4.637 ND 2.0

Mountain Crk Lake, Dallas, TX SMC 1995 June R M 430 1000 747 1233 0.061 ND 2.0

Mountain Crk Lake, Dallas, TX SMC 1995 June R M 420 1033 183 1391 0.132 ND 2.0

Mountain Crk Lake, Dallas, TX SMC 1995 June R M 398 860 511 1241 0.412 ND 2.0
Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Muddy Riv inflo to L Mead, NV SW 1996 May R F 533 2121 732 438 1.671 0.679 2.0

Muddy Riv inflo to L Mead, NV SW 1996 May R F 529 2462 561 514 1.091 0.831 3.0

Muddy Riv inflo to L Mead, NV SW 1996 May R F 502 1697 909 373 2.437 0.300 4.0

Muddy Riv inflo to L Mead, NV SW 1996 May R F 497 1695 831 282 2.947 1.198 4.0

Muddy Riv inflo to L Mead, NV SW 1996 May R F 485 1628 1047 149 7.027 4.816 3.0

Muddy Riv inflo to L Mead, NV SW 1996 May R F 466 1532 721 184 3.918 0.131 3.0

Muddy Riv inflo to L Mead, NV SW 1996 May R F 462 1309 922 528 1.746 0.498 3.0

Muddy Riv inflo to L Mead, NV SW 1996 May R F 453 1120 163 210 0.776 0.402 4.0

Muddy Riv inflo to L Mead, NV SW 1996 May R F 449 1431 670 122 5.492 1.446 3.0

Muddy Riv inflo to L Mead, NV SW 1996 May R F 441 1110 841 491 1.713 1.436 3.0

Muddy Riv inflo to L Mead, NV SW 1996 May R M 538 1591 112 468 0.239 ND 2.0

Muddy Riv inflo to L Mead, NV SW 1996 May R M 492 1539 98 248 0.395 ND 2.0

Muddy Riv inflo to L Mead, NV SW 1996 May R M 491 1520 70 341 0.205 0.019 2.0

Muddy Riv inflo to L Mead, NV SW 1996 May R M 464 1198 78 109 0.716 ND 3.0

Muddy Riv inflo to L Mead, NV SW 1996 May R M 463 1181 65 179 0.363 ND 3.0

Muddy Riv inflo to L Mead, NV SW 1996 May R M 462 959 67 135 0.496 0.015 3.0

Muddy Riv inflo to L Mead, NV SW 1996 May R M 452 1076 89 290 0.307 ND 3.0

Muddy Riv inflo to L Mead, NV SW 1996 May R M 449 1130 63 362 0.174 0.010 3.0

Muddy Riv inflo to L Mead, NV SW 1996 May R M 445 1070 91 142 0.641 0.005 3.0

Muddy Riv inflo to L Mead, NV SW 1996 May R M 393 666 64 256 0.250 ND 2.0

Mullet River nr Plymouth, WI NMC 1994 Sept NR F 522 2405 2976 590 5.04 29.600 4.0

Mullet River nr Plymouth, WI NMC 1994 Sept NR F 450 1223 3830 503 7.61 26.200 3.0

Mullet River nr Plymouth, WI NMC 1994 Sept NR F 448 1260 2419 441 5.49 30.800 3.0

Mullet River nr Plymouth, WI NMC 1994 Sept NR F 445 1111 2398 686 3.50 24.600 3.0

Mullet River nr Plymouth, WI NMC 1994 Sept NR F 444 1184 1837 779 2.36 32.300 3.0

Mullet River nr Plymouth, WI NMC 1994 Sept NR F 436 1265 1276 72 17.72 28.600 2.0

Mullet River nr Plymouth, WI NMC 1994 Sept NR F 422 966 1725 782 2.21 36.200 3.0

Mullet River nr Plymouth, WI NMC 1994 Sept NR F 421 1020 1410 948 1.49 24.600 3.0

Mullet River nr Plymouth, WI NMC 1994 Sept NR F 421 1056 2580 1503 1.72 22.400 3.0

Mullet River nr Plymouth, WI NMC 1994 Sept NR F 398 926 3245 532 6.10 20.900 3.0

Mullet River nr Plymouth, WI NMC 1994 Sept NR M 490 1430 715 253 2.83 ND 1.0

Mullet River nr Plymouth, WI NMC 1994 Sept NR M 450 1012 735 882 0.83 ND 1.0

Mullet River nr Plymouth, WI NMC 1994 Sept NR M 445 1103 399 255 1.57 0.020 1.0

Mullet River nr Plymouth, WI NMC 1994 Sept NR M 438 1070 820 940 0.87 ND 3.0

Mullet River nr Plymouth, WI NMC 1994 Sept NR M 430 985 701 1306 0.54 ND 1.0

Mullet River nr Plymouth, WI NMC 1994 Sept NR M 428 838 365 829 0.44 ND 2.0

Mullet River nr Plymouth, WI NMC 1994 Sept NR M 423 907 620 691 0.90 ND 2.0

Mullet River nr Plymouth, WI NMC 1994 Sept NR M 410 891 638 549 1.16 ND 3.0

Mullet River nr Plymouth, WI NMC 1994 Sept NR M 409 855 859 726 1.18 0.020 2.0

Mullet River nr Plymouth, WI NMC 1994 Sept NR M 405 712 704 634 1.11 ND 1.0

Table 4 53
54 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR F 630 2994 4466 323 13.83 29.500 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR F 582 2408 6147 2020 3.04 30.500 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR F 570 2717 354 223 1.59 22.500 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR F 560 2818 2233 985 2.27 43.000 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR F 550 2186 2877 114 25.24 19.000 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR F 543 1964 4128 254 16.25 16.000 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR F 536 1838 1707 364 4.69 18.700 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR F 520 1773 5719 141 40.56 16.000 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR F 519 1866 1926 373 5.16 22.000 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR F 510 1822 1254 564 2.22 20.500 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR F 502 1854 3094 192 16.11 16.500 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR F 470 1344 1178 731 1.61 16.000 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR F 462 1163 2253 1062 2.12 29.500 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR F 452 1085 876 84 10.43 47.500 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR F 441 979 1117 984 1.14 19.500 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR F 438 1048 1853 945 1.96 31.500 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR F 402 892 2742 1400 1.96 32.000 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR M 560 2154 1032 699 1.48 ND 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR M 548 1889 759 732 1.04 ND 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR M 541 1860 751 628 1.20 0.010 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR M 537 1740 690 609 1.13 ND 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR M 530 1763 811 484 1.68 0.060 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR M 517 1490 464 108 4.30 ND 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR M 517 1610 822 985 0.83 ND 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR M 509 1643 222 423 0.52 0.030 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR M 508 1518 302 518 0.58 ND 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR M 494 1524 951 225 4.23 ND 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR M 491 1392 653 942 0.69 ND 2.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR M 490 1354 899 792 1.14 ND 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR M 480 1312 483 422 1.14 ND 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR M 478 1294 214 262 0.82 ND 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR M 475 1242 862 668 1.29 ND .

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR M 465 1090 1340 1118 1.20 ND 2.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR M 452 1044 571 267 2.14 ND 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR M 450 1250 708 779 0.91 ND 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR M 443 1075 1064 977 1.09 0.020 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR M 441 868 1285 736 1.75 0.020 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR M 430 950 1296 449 2.89 ND 2.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR M 418 892 828 619 1.34 ND 3.0

N Br Milwaukee R @ Random L, WI NMC 1994 Sept NR M 414 880 821 842 0.98 15.000 3.0
Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

N F Willamette Riv @ Spgfield, OR NW 1994 Oct NR F 437 1022 670 398 1.68 7.829 1.0

N F Willamette Riv @ Spgfield, OR NW 1994 Oct NR F 424 1022 335 191 1.75 6.900 3.0

N F Willamette Riv @ Spgfield, OR NW 1994 Oct NR F 416 908 67 22 3.05 2.300 1.0

N F Willamette Riv @ Spgfield, OR NW 1994 Oct NR F 410 908 215 311 0.69 12.043 2.0

N F Willamette Riv @ Spgfield, OR NW 1994 Oct NR F 404 1022 860 603 1.43 3.600 3.0

N F Willamette Riv @ Spgfield, OR NW 1994 Oct NR F 403 795 222 72 3.08 36.900 1.0

N F Willamette Riv @ Spgfield, OR NW 1994 Oct NR F 403 681 1522 752 2.02 1.500 2.0

N F Willamette Riv @ Spgfield, OR NW 1994 Oct NR F 402 795 279 258 1.08 22.345 1.0

N F Willamette Riv @ Spgfield, OR NW 1994 Oct NR F 400 681 199 61 3.26 1.000 1.0

N F Willamette Riv @ Spgfield, OR NW 1994 Oct NR F 400 795 1232 670 1.84 3.214 2.0

N F Willamette Riv @ Spgfield, OR NW 1994 Oct NR F 396 908 227 414 0.55 0.570 1.0

N F Willamette Riv @ Spgfield, OR NW 1994 Oct NR F 390 795 1044 1030 1.01 20.400 3.0

N F Willamette Riv @ Spgfield, OR NW 1994 Oct NR F 390 681 536 445 1.20 3.093 1.0

N F Willamette Riv @ Spgfield, OR NW 1994 Oct NR F 385 795 125 174 0.72 0.310 1.0

N F Willamette Riv @ Spgfield, OR NW 1994 Oct NR M 482 1476 875 1738 0.50 ND 2.0

N F Willamette Riv @ Spgfield, OR NW 1994 Oct NR M 411 795 213 886 0.24 ND 3.0

N F Willamette Riv @ Spgfield, OR NW 1994 Oct NR M 394 795 135 321 0.42 ND 2.0

N F Willamette Riv @ Spgfield, OR NW 1994 Oct NR M 394 795 217 590 0.37 ND 2.0

N F Willamette Riv @ Spgfield, OR NW 1994 Oct NR M 393 681 124 869 0.14 ND 3.0

N F Willamette Riv @ Spgfield, OR NW 1994 Oct NR M 387 795 124 783 0.16 ND 3.0

N F Willamette Riv @ Spgfield, OR NW 1994 Oct NR M 380 681 185 1218 0.15 0.680 2.0

N F Willamette Riv @ Spgfield, OR NW 1994 Oct NR M 379 681 132 84 1.57 ND 3.0

N F Willamette Riv @ Spgfield, OR NW 1994 Oct NR M 378 681 174 1122 0.16 ND 2.0

N F Willamette Riv @ Spgfield, OR NW 1994 Oct NR M 373 681 30 382 0.08 ND 3.0

N F Willamette Riv @ Spgfield, OR NW 1994 Oct NR M 372 681 23 723 0.03 ND 2.0

N F Willamette Riv @ Spgfield, OR NW 1994 Oct NR M 370 681 79 751 0.11 ND 3.0

N F Willamette Riv @ Spgfield, OR NW 1994 Oct NR M 368 681 46 558 0.08 ND 2.0

N F Willamette Riv @ Spgfield, OR NW 1994 Oct NR M 362 568 135 649 0.21 0.160 3.0

N F Willamette Riv @ Spgfield, OR NW 1994 Oct NR M 339 454 47 279 0.17 ND 2.0

N F Willamette Riv @ Spgfield, OR NW 1995 July R F 445 1216 40 416 0.096 32.565 2.0

N F Willamette Riv @ Spgfield, OR NW 1995 July R F 442 1213 846 627 1.349 29.971 2.0

N F Willamette Riv @ Spgfield, OR NW 1995 July R F 436 1079 33 374 0.088 32.622 2.0

N F Willamette Riv @ Spgfield, OR NW 1995 July R F 416 1103 129 321 0.402 22.857 2.0

N F Willamette Riv @ Spgfield, OR NW 1995 July R F 411 920 17 404 0.042 19.372 2.0

N F Willamette Riv @ Spgfield, OR NW 1995 July R F 411 991 405 432 0.938 28.112 3.0

N F Willamette Riv @ Spgfield, OR NW 1995 July R F 411 1006 514 297 1.731 ND 2.0

N F Willamette Riv @ Spgfield, OR NW 1995 July R F 403 1049 288 737 0.391 29.958 3.0

N F Willamette Riv @ Spgfield, OR NW 1995 July R F 400 931 314 744 0.422 24.994 3.0

N F Willamette Riv @ Spgfield, OR NW 1995 July R F 396 901 1105 190 5.816 33.366 4.0

N F Willamette Riv @ Spgfield, OR NW 1995 July R F 389 873 356 707 0.504 40.040 2.0

Table 4 55
56 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

N F Willamette Riv @ Spgfield, OR NW 1995 July R F 382 783 940 494 1.903 33.082 2.0

N F Willamette Riv @ Spgfield, OR NW 1995 July R F 376 795 795 144 5.521 ND 2.0

N F Willamette Riv @ Spgfield, OR NW 1995 July R F 373 676 597 528 1.131 39.614 3.0

N F Willamette Riv @ Spgfield, OR NW 1995 July R F 345 550 259 459 0.564 11.924 3.0

N F Willamette Riv @ Spgfield, OR NW 1995 July R M 446 1096 284 756 0.376 ND 3.0

N F Willamette Riv @ Spgfield, OR NW 1995 July R M 385 865 305 532 0.573 ND 3.0

N F Willamette Riv @ Spgfield, OR NW 1995 July R M 382 773 265 327 0.810 0.090 3.0

N F Willamette Riv @ Spgfield, OR NW 1995 July R M 380 714 269 149 1.805 ND 3.0

N F Willamette Riv @ Spgfield, OR NW 1995 July R M 377 774 341 379 0.900 ND 2.0

N F Willamette Riv @ Spgfield, OR NW 1995 July R M 374 714 262 241 1.087 ND 3.0

N F Willamette Riv @ Spgfield, OR NW 1995 July R M 374 749 241 662 0.364 ND 3.0

N F Willamette Riv @ Spgfield, OR NW 1995 July R M 371 719 340 804 0.423 ND 3.0

N F Willamette Riv @ Spgfield, OR NW 1995 July R M 369 677 252 488 0.516 ND 1.0

N F Willamette Riv @ Spgfield, OR NW 1995 July R M 363 621 208 391 0.532 0.028 3.0

N F Willamette Riv @ Spgfield, OR NW 1995 July R M 362 666 299 458 0.653 ND 3.0

N F Willamette Riv @ Spgfield, OR NW 1995 July R M 354 611 261 671 0.389 ND 3.0

N F Willamette Riv @ Spgfield, OR NW 1995 July R M 341 547 121 186 0.651 ND 3.0

N F Willamette Riv @ Spgfield, OR NW 1995 July R M 336 553 231 775 0.298 ND 2.0

N F Willamette Riv @ Spgfield, OR NW 1995 July R M 327 806 265 821 0.323 ND 3.0

Ohio Riv @ Marietta, OH NE 1995 Oct NR F 710 4650 278 123 2.260 2.180 3

Ohio Riv @ Marietta, OH NE 1995 Oct NR F 535 1800 569 150 3.793 4.230 2

Ohio Riv @ Marietta, OH NE 1995 Oct NR M 552 1981 421 5001 0.084 0.013 1

Ohio Riv @ Marietta, OH NE 1995 Oct NR M 539 1740 401 97 4.134 ND 1

Ohio Riv @ Marietta, OH NE 1995 Oct NR M 479 1150 552 181 3.050 ND 1

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR F 736 5720 387 774 0.500 1.302 3

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR F 665 4050 349 267 1.307 2.220 3

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR F 660 1850 374 779 0.480 3.240 3

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR F 630 3200 769 115 6.687 0.845 3

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR F 510 1700 431 108 3.991 1.442 3

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR M 620 3600 469 841 0.558 ND 2

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR M 604 2800 170 7985 0.021 ND 1

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR M 580 2450 292 5635 0.052 ND 1

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR M 580 2700 397 337 1.178 ND 1

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR M 566 2250 285 316 0.902 ND 1

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR M 515 1500 185 271 0.683 ND 1.5

Ottertail Riv nr Breckenridge, MN NMC 1994 Sept NR F 530 2020 1086 372 2.92 33.200 3.0

Ottertail Riv nr Breckenridge, MN NMC 1994 Sept NR F 530 2008 2154 471 4.57 32.400 3.0

Ottertail Riv nr Breckenridge, MN NMC 1994 Sept NR F 530 1960 5162 1810 2.85 29.600 3.0

Ottertail Riv nr Breckenridge, MN NMC 1994 Sept NR F 530 1926 9942 3641 2.73 32.100 3.0

Ottertail Riv nr Breckenridge, MN NMC 1994 Sept NR F 525 1654 3213 978 3.29 30.000 3.0
Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Ottertail Riv nr Breckenridge, MN NMC 1994 Sept NR F 515 1520 1288 187 6.89 34.100 3.0

Ottertail Riv nr Breckenridge, MN NMC 1994 Sept NR F 510 1775 2143 262 8.18 25.800 3.0

Ottertail Riv nr Breckenridge, MN NMC 1994 Sept NR F 510 1735 5114 1863 2.75 30.900 3.0

Ottertail Riv nr Breckenridge, MN NMC 1994 Sept NR M 520 1763 57 77 0.74 ND 3.0

Ottertail Riv nr Breckenridge, MN NMC 1994 Sept NR M 510 1640 123 383 0.32 ND 3.0

Ottertail Riv nr Breckenridge, MN NMC 1994 Sept NR M 500 1420 1097 1017 1.08 ND 3.0

Ottertail Riv nr Breckenridge, MN NMC 1994 Sept NR M 490 1540 308 599 0.51 ND 3.0

Ottertail Riv nr Breckenridge, MN NMC 1994 Sept NR M 490 1265 682 1988 0.34 ND 2.0

Ottertail Riv nr Breckenridge, MN NMC 1994 Sept NR M 482 1370 1096 2404 0.46 0.050 2.0

Ottertail Riv nr Breckenridge, MN NMC 1994 Sept NR M 480 1306 376 329 1.14 ND 3.0

Ottertail Riv nr Breckenridge, MN NMC 1994 Sept NR M 475 1356 426 837 0.51 ND 3.0

Ottertail Riv nr Breckenridge, MN NMC 1994 Sept NR M 460 1253 1105 2935 0.38 0.090 3.0

Platte Riv @ Louisville, NE NMC 1994 Sept R F 662 3702 166 295 0.56 26.000 3.0

Platte Riv @ Louisville, NE NMC 1994 Sept R F 635 2386 271 547 0.50 40.500 1.0

Platte Riv @ Louisville, NE NMC 1994 Sept R F 606 3603 385 992 0.39 23.000 3.0

Platte Riv @ Louisville, NE NMC 1994 Sept R F 600 3137 1104 515 2.14 20.000 3.0

Platte Riv @ Louisville, NE NMC 1994 Sept R F 595 3094 40 180 0.22 ND 3.0

Platte Riv @ Louisville, NE NMC 1994 Sept R F 592 2948 243 536 0.45 72.500 2.0

Platte Riv @ Louisville, NE NMC 1994 Sept R F 588 3004 550 87 6.32 24.500 3.0

Platte Riv @ Louisville, NE NMC 1994 Sept R F 587 2984 129 152 0.85 30.000 3.0

Platte Riv @ Louisville, NE NMC 1994 Sept R F 581 2581 321 323 0.99 18.500 3.0

Platte Riv @ Louisville, NE NMC 1994 Sept R F 545 1738 132 214 0.62 0.240 1.0

Platte Riv @ Louisville, NE NMC 1994 Sept R F 539 1948 505 200 2.53 ND 3.0

Platte Riv @ Louisville, NE NMC 1994 Sept R F 515 2181 538 986 0.55 28.000 3.0

Platte Riv @ Louisville, NE NMC 1994 Sept R F 485 1520 862 1789 0.48 28.000 3.0

Platte Riv @ Louisville, NE NMC 1994 Sept R F 362 834 15 51 0.29 0.040 0.0

Platte Riv @ Louisville, NE NMC 1994 Sept R F 344 649 208 166 1.25 0.120 0.0

Platte Riv @ Louisville, NE NMC 1994 Sept R M 586 2456 53 2007 0.03 ND 3.0

Platte Riv @ Louisville, NE NMC 1994 Sept R M 584 2344 19 76 0.25 ND 3.0

Platte Riv @ Louisville, NE NMC 1994 Sept R M 563 1378 22 615 0.04 ND 3.0

Platte Riv @ Louisville, NE NMC 1994 Sept R M 536 1845 17 262 0.06 ND 3.0

Platte Riv @ Louisville, NE NMC 1994 Sept R M 505 1690 127 123 1.03 0.010 3.0

Platte Riv @ Louisville, NE NMC 1994 Sept R M 502 1758 36 132 0.27 ND 3.0

Platte Riv @ Louisville, NE NMC 1994 Sept R M 470 1130 21 987 0.02 ND 2.0

Platte Riv @ Louisville, NE NMC 1994 Sept R M 464 1069 19 166 0.11 ND 3.0

Platte Riv @ Louisville, NE NMC 1994 Sept R M 421 952 59 1109 0.05 ND 3.0

Platte Riv @ Louisville, NE NMC 1994 Sept R M 370 880 19 968 0.02 ND 3.0

Platte Riv @ Louisville, NE NMC 1994 Sept R M 317 464 79 99 0.80 ND 3.0

Platte Riv @ Louisville, NE NMC 1995 Oct NR F 512 2050 685 1033 0.66 2.087 3

Platte Riv @ Louisville, NE NMC 1995 Oct NR F 462 1350 587 1017 0.58 1.628 3

Table 4 57
58 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Platte Riv @ Louisville, NE NMC 1995 Oct NR M 543 1750 642 512 1.25 0.008 0

Platte Riv @ Louisville, NE NMC 1995 Oct NR M 517 1850 607 428 1.42 0.008 2

Platte Riv @ Louisville, NE NMC 1995 Oct NR M 440 1000 444 424 1.05 ND 1.5

Platte Riv @ Louisville, NE NMC 1995 Oct NR M 404 1100 709 1416 0.50 ND 1

Platte Riv @ Louisville, NE NMC 1995 Oct NR M 396 1000 1171 517 2.26 ND 1

Platte Riv @ Louisville, NE NMC 1995 Oct NR M 362 750 505 664 0.76 ND 2

Platte Riv @ Louisville, NE NMC 1995 Oct NR M 343 550 742 299 2.48 ND 0

Potomac R @ Anacostia R, DC NE 1994 Nov NR F 692 5477 1765 1089 1.62 17.200 3.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR F 689 4703 633 359 1.76 20.600 4.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR F 670 5038 323 338 0.96 6.640 3.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR F 605 3356 540 944 0.57 19.700 3.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR F 584 3280 631 1125 0.56 32.200 3.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR F 573 2853 605 159 3.81 16.300 3.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR F 557 3210 696 90 7.73 28.400 3.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR F 521 2636 1018 746 1.36 31.800 3.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR F 519 2343 782 66 11.85 24.300 3.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR F 496 2032 877 605 1.45 39.600 3.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR M 615 3545 484 851 0.57 ND 1.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR M 574 2660 528 1380 0.38 ND 1.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR M 555 2346 589 1579 0.37 ND 1.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR M 544 2335 226 895 0.25 ND 1.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR M 540 2870 309 358 0.86 ND 1.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR M 540 2345 204 1663 0.12 ND 1.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR M 535 2902 325 495 0.66 ND 1.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR M 530 2286 430 395 1.09 ND 1.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR M 530 2224 323 737 0.44 ND 1.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR M 521 2126 218 341 0.64 ND 2.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR M 516 1878 450 996 0.45 ND 1.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR M 515 2222 421 434 0.97 ND 1.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR M 509 1858 318 788 0.40 ND 1.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR M 498 1692 601 1375 0.44 ND 2.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR M 488 1498 425 440 0.97 ND 1.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR M 477 1713 320 407 0.79 ND 1.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR M 472 1720 298 514 0.58 ND 1.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR M 472 1711 511 758 0.67 ND 1.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR M 463 1526 411 2038 0.20 ND 1.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR M 455 1446 526 1877 0.28 ND 1.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR M 435 1525 369 1220 0.30 ND 1.0

Potomac R @ Anacostia R, DC NE 1994 Nov NR M 401 1073 317 495 0.64 ND 3.0

Red Riv @ Draton, ND NMC 1994 Sept NR F 831 6000 670 231 2.90 31.200 3.0
Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Red Riv @ Draton, ND NMC 1994 Sept NR F 782 6000 2740 248 11.05 24.800 3.0

Red Riv @ Draton, ND NMC 1994 Sept NR F 770 >6000 1352 123 10.99 44.100 3.0

Red Riv @ Draton, ND NMC 1994 Sept NR F 768 6000 3530 311 11.35 23.500 3.0

Red Riv @ Draton, ND NMC 1994 Sept NR F 730 4506 723 410 1.76 31.400 3.0

Red Riv @ Draton, ND NMC 1994 Sept NR F 720 5180 1290 825 1.56 21.200 3.0

Red Riv @ Draton, ND NMC 1994 Sept NR F 710 4035 1822 832 2.19 35.000 3.0

Red Riv @ Draton, ND NMC 1994 Sept NR F 698 4025 1212 206 5.88 22.000 3.0

Red Riv @ Draton, ND NMC 1994 Sept NR F 665 3690 533 124 4.30 30.200 3.0

Red Riv @ Draton, ND NMC 1994 Sept NR F 660 4060 552 127 4.35 ND 4.0

Red Riv @ Draton, ND NMC 1994 Sept NR F 518 1640 970 327 2.97 29.400 3.0

Red Riv @ Draton, ND NMC 1994 Sept NR M 675 3470 901 826 1.09 ND 2.0

Red Riv @ Draton, ND NMC 1994 Sept NR M 665 3160 641 706 0.91 ND 3.0

Red Riv @ Draton, ND NMC 1994 Sept NR M 615 2640 233 1245 0.19 0.060 3.0

Red Riv @ Draton, ND NMC 1994 Sept NR M 610 2585 342 658 0.52 ND 2.0

Red Riv @ Draton, ND NMC 1994 Sept NR M 585 2298 812 868 0.94 0.400 3.0

Red Riv @ Draton, ND NMC 1994 Sept NR M 580 2509 199 1077 0.18 ND 3.0

Red Riv @ Draton, ND NMC 1994 Sept NR M 560 1755 750 1947 0.39 ND 2.0

Red Riv @ Draton, ND NMC 1994 Sept NR M 547 1652 308 508 0.61 5.000 2.0

Red Riv @ Draton, ND NMC 1994 Sept NR M 535 1810 156 771 0.20 ND 2.0

Red Riv @ Draton, ND NMC 1994 Sept NR M 520 1645 100 1048 0.10 ND 3.0

Red Riv @ Grand Forks, ND NMC 1994 Sept NR F 534 1783 4811 563 8.55 8.500 3.0

Red Riv @ Grand Forks, ND NMC 1994 Sept NR F 533 1888 673 57 11.81 23.500 3.0

Red Riv @ Grand Forks, ND NMC 1994 Sept NR F 530 2063 4249 1329 3.20 19.000 2.0

Red Riv @ Grand Forks, ND NMC 1994 Sept NR F 526 1825 4697 806 5.83 28.000 3.0

Red Riv @ Grand Forks, ND NMC 1994 Sept NR F 507 1707 1774 307 5.78 26.000 3.0

Red Riv @ Grand Forks, ND NMC 1994 Sept NR F 485 1599 8075 1656 4.88 30.000 2.0

Red Riv @ Grand Forks, ND NMC 1994 Sept NR F 455 1296 443 149 2.97 87.800 2.0

Red Riv @ Grand Forks, ND NMC 1994 Sept NR F 453 1193 7763 596 13.03 7.800 3.0

Red Riv @ Grand Forks, ND NMC 1994 Sept NR F 436 1209 1226 848 1.45 25.000 3.0

Red Riv @ Grand Forks, ND NMC 1994 Sept NR F 424 1182 1720 2639 0.65 16.000 3.0

Red Riv @ Grand Forks, ND NMC 1994 Sept NR M 510 1470 1457 387 3.76 ND 3.0

Red Riv @ Grand Forks, ND NMC 1994 Sept NR M 472 1280 1696 1507 1.13 ND 3.0

Red Riv @ Grand Forks, ND NMC 1994 Sept NR M 464 1180 465 868 0.54 ND 2.0

Red Riv @ Grand Forks, ND NMC 1994 Sept NR M 436 1079 2104 665 3.16 0.070 2.0

Red Riv nr Breckenridge, MN NMC 1994 Sept NR F 610 3164 1208 467 2.59 35.300 4.0

Red Riv nr Breckenridge, MN NMC 1994 Sept NR F 580 2338 6852 178 38.49 36.600 3.0

Red Riv nr Breckenridge, MN NMC 1994 Sept NR F 550 2315 1475 117 12.61 31.000 3.0

Red Riv nr Breckenridge, MN NMC 1994 Sept NR F 550 2187 1013 445 2.28 41.000 3.0

Red Riv nr Breckenridge, MN NMC 1994 Sept NR F 540 2215 2291 240 9.55 31.900 3.0

Red Riv nr Breckenridge, MN NMC 1994 Sept NR F 535 2040 462 143 3.23 29.900 3.0

Table 4 59
60 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Red Riv nr Breckenridge, MN NMC 1994 Sept NR F 525 2040 2609 117 22.30 27.800 3.0

Red Riv nr Breckenridge, MN NMC 1994 Sept NR F 520 1954 2953 208 14.20 38.500 3.0

Red Riv nr Breckenridge, MN NMC 1994 Sept NR F 520 1900 808 443 1.82 23.400 3.0

Red Riv nr Breckenridge, MN NMC 1994 Sept NR F 520 1728 1728 190 9.09 28.600 3.0

Red Riv nr Breckenridge, MN NMC 1994 Sept NR F 520 1758 2534 481 5.27 35.000 3.0

Red Riv nr Breckenridge, MN NMC 1994 Sept NR F 512 1730 1090 540 2.02 28.800 3.0

Red Riv nr Breckenridge, MN NMC 1994 Sept NR F 490 1609 1154 337 3.42 35.100 3.0

Red Riv nr Breckenridge, MN NMC 1994 Sept NR F 485 1467 1368 63 21.71 33.500 3.0

Red Riv nr Breckenridge, MN NMC 1994 Sept NR M 620 2918 489 427 1.15 ND 3.0

Red Riv nr Breckenridge, MN NMC 1994 Sept NR M 535 1887 714 303 2.36 ND 3.0

Red Riv nr Breckenridge, MN NMC 1994 Sept NR M 520 1689 267 264 1.01 ND 3.0

Red Riv nr Breckenridge, MN NMC 1994 Sept NR M 510 1757 778 404 1.93 ND 3.0

Red Riv nr Breckenridge, MN NMC 1994 Sept NR M 495 1497 694 105 6.61 ND 3.0

Red Riv nr Breckenridge, MN NMC 1994 Sept NR M 495 1370 1112 466 2.39 ND 3.0

Red Riv nr Breckenridge, MN NMC 1994 Sept NR M 495 1468 1573 1340 1.17 0.050 3.0

Red Riv nr Breckenridge, MN NMC 1994 Sept NR M 480 1351 925 2221 0.42 ND 3.0

Red Riv nr Breckenridge, MN NMC 1994 Sept NR M 450 1106 463 431 1.07 ND 3.0

Red Riv nr Breckenridge, MN NMC 1994 Sept NR M 450 998 579 2261 0.26 ND 3.0

Red Riv @ Alexandria, LA SMC 1995 Oct NR F 750 5000 918 260 3.531 1.801 3

Red Riv @ Alexandria, LA SMC 1995 Oct NR F 619 3000 1190 365 3.260 1.613 2

Red Riv @ Alexandria, LA SMC 1995 Oct NR F 549 2322 1189 323 3.681 2.140 3

Red Riv @ Alexandria, LA SMC 1995 Oct NR F 500 1605 1782 261 6.828 2.490 3

Red Riv @ Alexandria, LA SMC 1995 Oct NR F 495 1540 591 399 1.481 2.023 3

Red Riv @ Alexandria, LA SMC 1995 Oct NR F 454 1265 1198 264 4.538 1.602 3

Red Riv @ Alexandria, LA SMC 1995 Oct NR F 442 1205 1374 382 3.597 2.897 3

Red Riv @ Alexandria, LA SMC 1995 Oct NR F 435 1097 1445 429 3.368 0.245 3

Red Riv @ Alexandria, LA SMC 1995 Oct NR M 651 2950 618 493 1.254 ND 2

Red Riv @ Alexandria, LA SMC 1995 Oct NR M 610 2918 746 1296 0.576 ND 3

Red Riv @ Alexandria, LA SMC 1995 Oct NR M 480 1490 459 507 0.905 ND 2

Red Riv @ Alexandria, LA SMC 1995 Oct NR M 473 1410 511 1185 0.431 ND 2

Red Riv @ L Texoma, TX SMC 1995 Nov NR F 730 4300 611 49 12.47 0.265 3

Red Riv @ L Texoma, TX SMC 1995 Nov NR F 692 4000 907 237 3.83 1.428 3

Red Riv @ L Texoma, TX SMC 1995 Nov NR F 683 4600 538 257 2.09 1.065 3

Red Riv @ L Texoma, TX SMC 1995 Nov NR F 680 4100 657 1136 0.58 1.365 3

Red Riv @ L Texoma, TX SMC 1995 Nov NR F 660 3150 588 70 8.40 0.946 3

Red Riv @ L Texoma, TX SMC 1995 Nov NR F 645 3400 755 89 8.48 1.263 3

Red Riv @ L Texoma, TX SMC 1995 Nov NR F 630 3600 1080 81 13.33 0.632 3

Red Riv @ L Texoma, TX SMC 1995 Nov NR F 628 3000 817 32 25.53 0.883 3

Red Riv @ L Texoma, TX SMC 1995 Nov NR F 621 3400 860 113 7.61 2.463 3

Red Riv @ L Texoma, TX SMC 1995 Nov NR F 610 3000 583 56 10.41 0.561 3
Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Red Riv @ L Texoma, TX SMC 1995 Nov NR F 590 2600 956 227 4.21 1.137 3

Red Riv @ L Texoma, TX SMC 1995 Nov NR F 553 1800 626 229 2.73 2.875 3

Red Riv @ L Texoma, TX SMC 1995 Nov NR F 535 1850 557 622 0.90 3.142 3

Red Riv @ L Texoma, TX SMC 1995 Nov NR M 670 3700 506 2718 0.19 ND 3

Red Riv @ L Texoma, TX SMC 1995 Nov NR M 660 3000 558 90 6.20 ND 2

Red Riv @ L Texoma, TX SMC 1995 Nov NR M 612 2750 1037 107 9.69 ND 2

Red Riv @ L Texoma, TX SMC 1995 Nov NR M 580 2500 403 416 0.97 ND 2

Red Riv @ L Texoma, TX SMC 1995 Nov NR M 580 2500 571 1084 0.53 ND 2

Red Riv @ L Texoma, TX SMC 1995 Nov NR M 566 2800 626 2983 0.21 0.093 2

Red Riv @ L Texoma, TX SMC 1995 Nov NR M 565 1900 769 61 12.61 ND 3

Red Riv @ L Texoma, TX SMC 1995 Nov NR M 555 1800 659 1632 0.40 ND 3

Red Riv @ L Texoma, TX SMC 1995 Nov NR M 540 1900 609 75 8.12 ND 2.5

Red Riv @ L Texoma, TX SMC 1995 Nov NR M 521 1700 583 1292 0.45 ND 2.5

Red Riv @ L Texoma, TX SMC 1995 Nov NR M 490 1550 636 1415 0.45 0.008 2

Royal Lake, nr Othello, WA NW 1995 July R F 710 5190 1211 628 1.928 55.468 3.0

Royal Lake, nr Othello, WA NW 1995 July R F 705 5320 1487 564 2.637 57.608 3.0

Royal Lake, nr Othello, WA NW 1995 July R F 704 4557 1209 541 2.235 23.570 3.0

Royal Lake, nr Othello, WA NW 1995 July R F 700 4915 1162 622 1.868 16.589 3.0

Royal Lake, nr Othello, WA NW 1995 July R F 698 4859 1284 716 1.793 21.934 3.0

Royal Lake, nr Othello, WA NW 1995 July R F 660 4438 1005 476 2.111 25.553 3.0

Royal Lake, nr Othello, WA NW 1995 July R F 660 4349 1475 619 2.383 57.608 3.0

Royal Lake, nr Othello, WA NW 1995 July R F 652 4542 1392 695 2.003 82.643 3.0

Royal Lake, nr Othello, WA NW 1995 July R F 648 3913 1717 543 3.162 26.206 3.0

Royal Lake, nr Othello, WA NW 1995 July R F 643 4011 1418 916 1.548 17.480 3.0

Royal Lake, nr Othello, WA NW 1995 July R F 642 4625 1245 601 2.072 86.521 4.0

Royal Lake, nr Othello, WA NW 1995 July R F 638 3980 1688 1041 1.622 21.701 3.0

Royal Lake, nr Othello, WA NW 1995 July R F 631 3666 1299 778 1.670 60.555 3.0

Royal Lake, nr Othello, WA NW 1995 July R F 535 2296 1621 874 1.855 86.521 3.0

Royal Lake, nr Othello, WA NW 1995 July R F 530 2192 3542 1507 2.350 18.485 3.0

Royal Lake, nr Othello, WA NW 1995 July R M 683 4012 731 1426 0.513 ND 3.0

Royal Lake, nr Othello, WA NW 1995 July R M 654 3825 243 558 0.435 ND 3.0

Royal Lake, nr Othello, WA NW 1995 July R M 649 3692 265 409 0.648 ND 3.0

Royal Lake, nr Othello, WA NW 1995 July R M 648 3810 944 2181 0.433 ND 3.0

Royal Lake, nr Othello, WA NW 1995 July R M 634 3955 565 1239 0.456 ND 3.0

Royal Lake, nr Othello, WA NW 1995 July R M 632 3395 383 1488 0.257 ND 3.0

Royal Lake, nr Othello, WA NW 1995 July R M 630 3560 325 820 0.396 ND 3.0

Royal Lake, nr Othello, WA NW 1995 July R M 628 3196 930 1161 0.801 ND 3.0

Royal Lake, nr Othello, WA NW 1995 July R M 623 3448 645 830 0.777 ND 3.0

Royal Lake, nr Othello, WA NW 1995 July R M 619 3870 572 989 0.578 ND 3.0

Royal Lake, nr Othello, WA NW 1995 July R M 612 3020 249 937 0.266 ND 3.0

Table 4 61
62 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Royal Lake, nr Othello, WA NW 1995 July R M 595 2794 226 393 0.575 ND 3.0

Royal Lake, nr Othello, WA NW 1995 July R M 592 2750 850 1308 0.650 ND 3.0

Royal Lake, nr Othello, WA NW 1995 July R M 583 2960 1129 1429 0.790 ND 3.0

Royal Lake, nr Othello, WA NW 1995 July R M 575 2770 249 396 0.629 ND 3.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR F 625 3186 1400 855 1.64 21.900 3.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR F 620 3295 333 769 0.43 34.500 3.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR F 614 2998 1996 1172 1.70 42.600 3.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR F 599 2442 896 864 1.04 31.200 3.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR F 548 2028 456 856 0.53 34.800 3.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR F 531 2148 2299 778 2.96 38.000 3.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR F 513 1726 1379 970 1.42 33.800 3.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR F 500 1355 411 441 0.93 35.600 3.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR F 495 1488 1390 861 1.61 31.300 3.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR F 473 1386 1847 1079 1.71 31.200 3.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR F 470 1410 1451 634 2.29 30.800 3.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR F 460 1208 1006 1085 0.93 36.000 3.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR F 442 1121 875 391 2.24 39.600 4.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR F 404 970 287 414 0.69 38.500 3.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR F 350 635 253 224 1.13 22.900 3.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR F 338 499 392 189 2.07 29.800 3.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR F 323 460 204 177 1.15 2.000 2.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR M 640 2665 90 983 0.09 ND 3.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR M 595 2657 279 2040 0.14 ND 3.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR M 537 2234 136 1451 0.09 ND 2.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR M 529 1865 189 2008 0.09 ND 3.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR M 523 1689 336 787 0.43 ND 2.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR M 503 1232 126 517 0.24 ND 3.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR M 475 1328 129 981 0.13 ND 2.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR M 475 1472 309 1100 0.28 ND 3.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR M 445 986 151 2010 0.08 ND 3.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR M 437 1066 295 687 0.43 ND 3.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR M 350 580 157 677 0.23 ND 3.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR M 330 480 122 1035 0.12 ND 3.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR M 315 408 130 287 0.45 ND 3.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR M 311 396 139 462 0.30 ND 3.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR M 304 334 454 355 1.28 ND 3.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR M 302 407 272 951 0.29 ND 3.0

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR F 742 6356 3234 1745 1.85 17.842 3.0

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR F 700 5549 2019 1113 1.81 27.223 3.0

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR F 660 4637 1654 1390 1.19 16.576 3.0
Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR F 656 5142 1739 5152 0.34 11.662 3.0

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR F 627 3026 2596 7511 0.35 20.941 3.0

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR F 625 2915 3117 9066 0.34 13.306 3.0

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR F 598 3365 2344 1582 1.48 37.459 3.0

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR F 580 2680 8333 2004 4.16 16.670 3.0

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR F 550 2287 1164 1405 0.83 16.462 3.0

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR F 532 2203 3530 8965 0.39 15.631 3.0

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR F 525 1713 1101 2204 0.50 22.072 3.0

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR F 475 1382 4047 6369 0.64 12.021 3.0

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR F 472 959 1110 488 2.27 8.600 3.0

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR F 467 1248 861 152 5.66 14.166 3.0

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR F 463 1459 2353 1187 1.98 15.999 3.0

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR F 445 1221 303 698 0.43 NA NA

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR F 423 984 970 1024 0.95 5.812 2.0

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR M 598 2905 337 1223 0.28 0.093 3.0

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR M 580 2700 1241 9699 0.13 NA NA

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR M 553 2263 188 1703 0.11 ND 3.0

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR M 516 1745 431 9379 0.05 0.183 2.0

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR M 450 1318 356 6440 0.06 ND 2.0

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR M 443 1090 199 2538 0.08 ND 2.0

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR M 435 1038 1695 1011 1.68 NA NA

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR M 427 1015 641 1164 0.55 ND 3.0

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR M 422 882 285 1637 0.17 ND 3.0

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR M 401 787 51 183 0.28 ND 2.0

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR M 388 641 454 2084 0.22 0.011 2.0

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR M NA NA 2388 2284 1.05 NA NA

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 755 6840 1334 2093 0.64 14.341 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 745 5765 5214 284 18.36 75.235 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 700 5489 518 4603 0.11 20.790 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 697 5701 5496 1675 3.28 52.152 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 684 5365 3586 1520 2.36 30.321 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 680 4154 1797 1371 1.31 11.662 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 617 3519 11401 5973 1.91 53.811 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 616 3606 315 1606 0.20 2.943 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 616 3367 4312 3054 1.41 16.541 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 615 2992 3403 3921 0.87 21.187 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 600 3039 5102 3025 1.69 16.431 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 596 2963 5777 3124 1.85 62.377 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 586 3148 3448 2030 1.70 45.384 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 479 1561 449 706 0.64 35.445 3.0

Table 4 63
64 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 411 919 1952 274 7.12 13.544 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 May R F 720 5256 577 800 0.721 56.932 2.0

San Joaquin Riv @ Vernalis, CA SW 1995 May R F 707 4836 763 2309 0.330 9.472 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 May R F 692 4846 951 375 2.536 19.341 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 May R F 690 3923 206 385 0.535 34.561 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 May R F 684 4050 640 1270 0.504 5.624 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 May R F 675 4110 583 466 1.251 25.678 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 May R F 664 3108 115 427 0.269 2.347 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 May R F 658 3634 738 1114 0.662 12.540 2.0

San Joaquin Riv @ Vernalis, CA SW 1995 May R F 588 3040 1060 689 1.538 24.359 4.0

San Joaquin Riv @ Vernalis, CA SW 1995 May R F 569 2789 506 909 0.557 14.924 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 May R F 415 1020 542 487 1.113 22.934 1.0

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR M 618 2928 4546 3652 1.24 0.041 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR M 613 3300 51 686 0.07 0.310 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR M 585 2498 373 3894 0.10 ND 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR M 574 2348 721 1871 0.39 ND 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR M 570 2736 1484 2081 0.71 0.183 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR M 558 2344 250 656 0.38 ND 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR M 541 2164 301 423 0.71 ND 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR M 533 2163 2605 1356 1.92 ND 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR M 530 2420 2326 1087 2.14 ND 2.0

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR M 514 1658 463 486 0.95 ND 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR M 510 1795 2233 1604 1.39 ND 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR M 479 477 457 4978 0.09 ND 2.0

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR M 445 1239 198 2667 0.07 ND 2.0

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR M 424 1085 177 1327 0.13 3.148 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR M 420 1022 801 4569 0.18 ND 2.0

San Joaquin Riv @ Vernalis, CA SW 1995 May R M 649 3360 731 826 0.885 ND 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 May R M 619 2992 326 942 0.346 ND 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 May R M 610 2690 181 1866 0.097 ND 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 May R M 580 1923 280 191 1.466 ND 3.0

San Joaquin Riv @ Vernalis, CA SW 1995 May R M 544 1955 257 1087 0.236 0.037 1.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR F 340 486 146 17 8.59 6.300 1.0

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR F 328 498 129 209 0.62 1.040 1.0

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 795 7290 2735 994 2.75 ND 3.0

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 718 5175 2384 971 2.46 34.800 3.0

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 695 5175 3196 1144 2.79 37.400 3.0

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 632 4608 2778 807 3.44 37.900 3.0

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 623 3329 1159 659 1.76 40.700 3.0

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 610 3880 2503 808 3.10 42.200 3.0
Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 588 3035 2298 514 4.47 42.600 3.0

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 584 3207 1445 497 2.91 40.200 3.0

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 575 2967 2784 520 5.35 32.500 3.0

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 522 1975 2126 834 2.55 44.700 3.0

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 518 1978 377 152 2.48 41.400 1.0

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 510 1773 500 407 1.23 25.700 2.0

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 502 1979 939 439 2.14 28.600 3.0

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 495 1817 658 238 2.76 36.300 3.0

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 485 1557 135 121 1.12 47.900 4.0

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 447 1242 210 161 1.30 9.200 1.0

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 438 1277 808 354 2.28 34.300 3.0

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 436 1294 1032 401 2.57 46.400 3.0

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 429 1043 377 232 1.63 39.600 3.0

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 425 913 473 169 2.80 3.600 1.0

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F NA NA 784 325 2.41 31.400 3.0

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR M 660 3650 601 739 0.81 ND 3.0

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR M 605 3559 596 1511 0.39 ND 3.0

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR M 590 2825 804 1354 0.59 ND 3.0

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR M 586 2790 563 473 1.19 ND 3.0

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR M 580 3806 871 790 1.10 ND 2.0

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR M 508 1993 242 762 0.32 ND 2.0

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR M 500 1676 434 408 1.06 ND 3.0

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR M 498 1360 514 305 1.69 ND 2.0

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR M 455 1493 307 458 0.67 ND 3.0

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR M 440 1221 431 149 2.89 ND 3.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR F 726 5335 880 183 4.81 12.900 3.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR F 705 3942 1298 86 15.09 11.800 4.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR F 661 4320 1580 85 18.59 13.100 3.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR F 654 4151 1639 520 3.15 14.800 3.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR F 648 4192 1294 443 2.92 18.000 3.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR F 646 4028 315 271 1.16 13.600 3.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR F 643 4272 1366 385 3.55 18.300 3.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR F 618 3333 1760 430 4.09 19.300 3.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR F 615 3720 1448 700 2.07 14.300 3.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR F 602 3287 536 250 2.14 20.400 3.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR F 596 3878 348 13 26.77 ND 3.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR F 594 3276 1440 237 6.08 18.500 3.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR F 574 2787 1045 37 28.24 8.900 3.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR F 518 1834 1275 245 5.20 5.800 3.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR M 637 3764 50 75 0.67 ND 1.0

Table 4 65
66 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Shenandoah Riv @ Melville, WV NE 1994 Nov NR M 630 3744 158 249 0.63 ND 1.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR M 609 3530 201 869 0.23 ND 1.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR M 608 2751 113 226 0.50 ND 1.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR M 607 3418 468 259 1.81 ND 1.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR M 599 3010 219 133 1.65 ND 1.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR M 587 2855 872 682 1.28 ND 1.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR M 585 3126 479 565 0.85 ND 2.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR M 585 2891 234 672 0.35 0.020 1.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR M 583 2777 433 323 1.34 ND 2.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR M 580 2811 206 646 0.32 ND 1.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR M 577 3041 111 344 0.32 ND 2.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR M 574 2846 69 138 0.50 ND 1.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR M 570 2361 190 161 1.18 ND 1.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR M 564 2446 204 140 1.46 ND 1.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR M 563 2120 203 307 0.66 ND 1.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR M 561 2584 301 427 0.70 0.270 1.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR M 555 2406 326 277 1.18 ND 1.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR M 548 2242 217 166 1.31 ND 1.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR M 548 3003 393 1187 0.33 ND 1.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR M 520 2086 157 842 0.19 0.210 1.0

Shenandoah Riv @ Melville, WV NE 1994 Nov NR M 507 1832 107 577 0.19 ND 1.0

Snake @ Ice Harbor Dam, WA NW 1997 Oct NR F 745 5817 777 79 9.84 1.626 2

Snake Riv @ Hagerman, ID NW 1997 Oct NR F 464 1270 562 264 2.13 0.111 1

Snake Riv @ Hagerman, ID NW 1997 Oct NR F 451 1289 701 251 2.79 2.805 2

Snake Riv @ Hagerman, ID NW 1997 Oct NR F 425 1090 797 227 3.51 0.009 1

Snake Riv @ Hagerman, ID NW 1997 Oct NR F 423 1017 483 94 5.14 1.490 1

Snake Riv @ Hagerman, ID NW 1997 Oct NR F 386 681 852 286 2.98 0.009 0

Snake Riv @ Hagerman, ID NW 1997 Oct NR F 382 701 1292 491 2.63 0.016 1

Snake Riv @ Hagerman, ID NW 1997 Oct NR F 372 841 1990 191 10.42 1.406 2

Snake Riv @ Hagerman, ID NW 1997 Oct NR F 368 708 1458 381 3.83 0.184 1

Snake Riv @ Hagerman, ID NW 1997 Oct NR F 368 717 1094 89 12.29 0.013 1

Snake Riv @ Hagerman, ID NW 1997 Oct NR F 355 603 1712 612 2.80 0.962 1

Snake Riv @ Hagerman, ID NW 1997 Oct NR M 406 882 239 1397 0.17 ND 2

Snake Riv @ Hagerman, ID NW 1997 Oct NR M 390 900 145 1440 0.10 ND 2

Snake Riv @ Hagerman, ID NW 1997 Oct NR M 380 829 575 1503 0.38 ND 3

Snake Riv @ Hagerman, ID NW 1997 Oct NR M 373 669 135 846 0.16 ND 2

Snake Riv @ Hagerman, ID NW 1997 Oct NR M 370 707 453 1174 0.39 ND 2

Snake Riv @ Hagerman, ID NW 1997 Oct NR M 366 800 152 495 0.31 ND 2

Snake Riv @ Hagerman, ID NW 1997 Oct NR M 365 600 498 215 2.32 ND 3

Snake Riv @ Hagerman, ID NW 1997 Oct NR M 340 538 495 279 1.77 ND 2
Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Snake Riv @ Hagerman, ID NW 1997 Oct NR M 325 506 185 128 1.45 ND 3

Snake Riv @ Hagerman, ID NW 1997 Oct NR M 321 397 164 290 0.57 ND 2

Snake Riv @ Lewiston, ID NW 1997 Oct NR F 639 4200 1168 367 3.183 5.043 2

Snake Riv @ Lewiston, ID NW 1997 Oct NR F 630 3600 1115 743 1.501 7.368 2

Snake Riv @ Lewiston, ID NW 1997 Oct NR F 615 3500 1341 844 1.589 1.437 2

Snake Riv @ Lewiston, ID NW 1997 Oct NR F 585 2900 1181 186 6.349 2.527 2

Snake Riv @ Lewiston, ID NW 1997 Oct NR M 601 2500 792 601 1.318 ND 2

Snake Riv @ Lewiston, ID NW 1997 Oct NR M 600 3300 733 260 2.819 ND 2

Snake Riv @ Lewiston, ID NW 1997 Oct NR M 565 2500 642 277 2.318 ND 2

Snake Riv @ Lewiston, ID NW 1997 Oct NR M 562 2500 493 619 0.796 ND 3

Snake Riv @ Lewiston, ID NW 1997 Oct NR M 550 2300 262 709 0.370 ND 3

Snake Riv @ Lewiston, ID NW 1997 Oct NR M 547 2700 401 1214 0.330 ND 3

Snake Riv @ Lewiston, ID NW 1997 Oct NR M 530 2200 268 1638 0.164 ND 3

Snake Riv @ Lewiston, ID NW 1997 Oct NR M 468 1300 171 458 0.373 ND 2

Snake Riv @ Lewiston, ID NW 1997 Oct NR M 440 1400 302 1230 0.246 ND 2

Snake R @ Ice Harbor Dam, WA NW 1997 Oct NR F 745 6055 1241 133 9.33 1.676 2

Snake R @ Ice Harbor Dam, WA NW 1997 Oct NR F 707 6738 913 272 3.36 2.945 2

Snake R @ Ice Harbor Dam, WA NW 1997 Oct NR F 704 5634 1411 447 3.16 1.711 2

Snake R @ Ice Harbor Dam, WA NW 1997 Oct NR F 659 4200 834 427 1.95 2.983 2

Snake R @ Ice Harbor Dam, WA NW 1997 Oct NR F 645 4000 1172 295 3.97 1.457 2

Snake R @ Ice Harbor Dam, WA NW 1997 Oct NR F 633 2300 658 81 8.12 2.391 2

Snake R @ Ice Harbor Dam, WA NW 1997 Oct NR F 613 2900 1462 257 5.69 1.857 2

Snake R @ Ice Harbor Dam, WA NW 1997 Oct NR F 551 2150 589 261 2.26 3.109 2

Snake R @ Ice Harbor Dam, WA NW 1997 Oct NR F 535 2400 518 348 1.49 1.635 2

Snake R @ Ice Harbor Dam, WA NW 1997 Oct NR M 667 4000 141 696 0.20 ND 3

Snake R @ Ice Harbor Dam, WA NW 1997 Oct NR M 644 3450 131 294 0.45 ND 2

Snake R @ Ice Harbor Dam, WA NW 1997 Oct NR M 635 3600 133 261 0.51 ND 3

Snake R @ Ice Harbor Dam, WA NW 1997 Oct NR M 617 3200 252 746 0.34 ND 3

Snake R @ Ice Harbor Dam, WA NW 1997 Oct NR M 586 2750 127 470 0.27 ND 2

Snake R @ Ice Harbor Dam, WA NW 1997 Oct NR M 572 2600 72 472 0.15 ND 3

Snake R @ Ice Harbor Dam, WA NW 1997 Oct NR M 569 2500 127 735 0.17 ND 2

Snake R @ Ice Harbor Dam, WA NW 1997 Oct NR M 568 2600 463 694 0.67 ND 2

Snake R @ Ice Harbor Dam, WA NW 1997 Oct NR M 567 2500 249 1648 0.15 ND 2

Snake R @ Ice Harbor Dam, WA NW 1997 Oct NR M 563 2400 124 185 0.67 0.009 3

Snake R @ Ice Harbor Dam, WA NW 1997 Oct NR M 560 2200 152 197 0.77 ND 2

So Platte R below No Platte, NE NMC 1994 Sept R F 573 2708 1074 343 3.13 30.100 3.0

So Platte R below No Platte, NE NMC 1994 Sept R F 570 2571 561 424 3.12 31.000 3.0

So Platte R below No Platte, NE NMC 1994 Sept R F 566 2415 1005 383 2.62 19.200 3.0

So Platte R below No Platte, NE NMC 1994 Sept R F 542 2511 1356 338 4.01 23.600 3.0

So Platte R below No Platte, NE NMC 1994 Sept R F 540 1869 1889 352 5.37 27.600 3.0

Table 4 67
68 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

So Platte R below No Platte, NE NMC 1994 Sept R F 536 2105 1873 421 4.45 28.600 3.0

So Platte R below No Platte, NE NMC 1994 Sept R F 510 1729 838 427 1.96 29.600 3.0

So Platte R below No Platte, NE NMC 1994 Sept R F 505 1457 1682 217 7.75 14.000 3.0

So Platte R below No Platte, NE NMC 1994 Sept R F 500 1607 813 227 3.58 37.400 3.0

So Platte R below No Platte, NE NMC 1994 Sept R F 495 1441 719 433 1.66 24.000 3.0

So Platte R below No Platte, NE NMC 1994 Sept R F 490 1129 484 167 2.90 0.720 1.0

So Platte R below No Platte, NE NMC 1994 Sept R F 480 1142 1469 409 3.59 23.300 4.0

So Platte R below No Platte, NE NMC 1994 Sept R F 435 910 745 278 2.68 1.760 3.0

So Platte R below No Platte, NE NMC 1994 Sept R M 615 2074 396 254 1.56 ND 2.0

So Platte R below No Platte, NE NMC 1994 Sept R M 540 1942 627 629 1.00 ND 2.0

So Platte R below No Platte, NE NMC 1994 Sept R M 528 2152 466 723 0.64 ND 1.0

So Platte R below No Platte, NE NMC 1994 Sept R M 429 1041 451 538 0.84 ND 1.0

So Platte R below No Platte, NE NMC 1994 Sept R M 390 838 691 352 1.63 ND 1.0

So Platte Riv @ Denver, CO NMC 1994 Sept R F 675 4588 1787 694 2.57 34.000 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R F 655 4535 2851 1500 1.90 16.000 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R F 651 4030 3249 1270 2.56 33.500 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R F 650 4200 952 182 5.23 39.500 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R F 646 3908 1943 970 2.00 14.700 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R F 640 4175 1477 829 1.78 26.000 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R F 640 3200 1983 1040 1.91 ND 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R F 632 3827 1666 808 2.06 15.800 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R F 622 3640 1863 674 2.76 41.500 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R F 615 3400 971 436 2.23 12.200 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R F 615 3720 2863 1127 2.54 16.000 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R F 612 3099 1312 1192 1.10 ND 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R F 604 2900 1449 343 4.22 0.700 1.0

So Platte Riv @ Denver, CO NMC 1994 Sept R F 600 3158 2643 1243 2.13 ND 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R F 566 2800 1523 708 2.15 15.200 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R F 542 1949 1249 440 2.84 ND 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R F 537 2300 1724 610 2.83 28.000 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R F 504 1700 1536 296 5.19 46.500 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R F 500 2000 1416 517 2.74 12.700 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R F 498 1920 791 360 2.20 30.700 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R F 468 1400 1829 1092 1.67 14.600 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R F 441 1200 1297 992 1.31 31.500 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R M 633 3243 118 894 0.13 ND 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R M 628 3700 20 805 0.03 ND 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R M 625 3200 25 494 0.05 ND 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R M 616 2971 419 1528 0.27 ND 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R M 582 2500 42 913 0.05 ND 3.0
Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

So Platte Riv @ Denver, CO NMC 1994 Sept R M 578 2719 231 1339 0.17 ND 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R M 574 2800 413 316 1.31 ND 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R M 570 2766 306 804 0.38 ND 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R M 570 2500 189 1093 0.17 ND 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R M 566 2200 247 221 1.12 ND 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R M 535 2200 102 399 0.26 0.020 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R M 523 1995 235 818 0.29 ND 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R M 522 2316 448 1249 0.36 ND 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R M 514 1700 362 585 0.62 ND 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R M 508 1900 591 902 0.66 ND 3.0

So Platte Riv @ Denver, CO NMC 1994 Sept R M 496 1700 368 550 0.67 0.030 2.0

So Platte Riv @ Denver, CO NMC 1995 Sept R F 636 3700 539 265 2.03 17.494 3.0

So Platte Riv @ Denver, CO NMC 1995 Sept R F 587 2950 286 242 1.18 7.334 3.0

So Platte Riv @ Denver, CO NMC 1995 Sept R F 580 2700 393 365 1.08 11.958 2.0

So Platte Riv @ Denver, CO NMC 1995 Sept R F 577 2700 1038 661 1.57 12.911 3.0

So Platte Riv @ Denver, CO NMC 1995 Sept R F 527 2120 219 1213 0.18 53.904 3.0

So Platte Riv @ Denver, CO NMC 1995 Sept R F 515 2200 112 253 0.44 18.631 2.0

So Platte Riv @ Denver, CO NMC 1995 Sept R F 513 1800 2143 171 12.53 9.380 2.0

So Platte Riv @ Denver, CO NMC 1995 Sept R F 505 1550 55 127 0.43 3.158 2.0

So Platte Riv @ Denver, CO NMC 1995 Sept R F 496 1750 545 346 1.58 17.824 2.0

So Platte Riv @ Denver, CO NMC 1995 Sept R F 457 1350 330 423 0.78 18.631 2.0

So Platte Riv @ Denver, CO NMC 1995 Sept R M 625 3050 23 4536 0.01 ND 3.0

So Platte Riv @ Denver, CO NMC 1995 Sept R M 615 2700 52 745 0.07 14.787 2.0

So Platte Riv @ Denver, CO NMC 1995 Sept R M 610 2600 146 2387 0.06 ND 2.0

So Platte Riv @ Denver, CO NMC 1995 Sept R M 605 3700 487 198 2.46 25.427 1.0

So Platte Riv @ Denver, CO NMC 1995 Sept R M 603 2750 126 6796 0.02 0.052 2.0

So Platte Riv @ Denver, CO NMC 1995 Sept R M 602 2700 169 5735 0.03 ND 2.0

So Platte Riv @ Denver, CO NMC 1995 Sept R M 601 2600 200 633 0.32 0.013 2.0

So Platte Riv @ Denver, CO NMC 1995 Sept R M 595 3150 259 1702 0.15 0.263 2.0

So Platte Riv @ Denver, CO NMC 1995 Sept R M 567 2100 300 680 0.44 ND 2.0

So Platte Riv @ Denver, CO NMC 1995 Sept R M 562 2100 113 740 0.15 ND 2.0

So Platte Riv @ Denver, CO NMC 1995 Sept R M 555 1990 86 1949 0.04 ND 1.0

So Platte Riv @ Denver, CO NMC 1995 Sept R M 545 1800 225 2488 0.09 ND 2.0

So Platte Riv @ Denver, CO NMC 1995 Sept R M 536 1820 88 1451 0.06 ND 1.0

So Platte Riv @ Denver, CO NMC 1995 Sept R M 490 1500 151 378 0.40 0.013 1.0

So Platte Riv @ Denver, CO NMC 1995 Sept R M 480 1400 113 2009 0.06 ND 1.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R F 620 3275 2038 1250 1.63 27.000 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R F 598 3028 819 244 3.36 39.900 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R F 580 2775 559 205 2.73 38.800 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R F 572 2510 1329 748 1.78 27.700 3.0

Table 4 69
70 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

So Platte Riv @ Kersey, CO NMC 1994 Sept R F 570 3176 1177 261 4.51 25.700 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R F 564 2466 951 365 2.61 38.000 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R F 554 2444 2163 780 2.77 21.400 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R F 554 2598 2195 910 2.41 32.700 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R F 545 2284 1438 383 3.75 21.000 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R F 543 2186 1094 302 3.62 25.100 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R F 540 2186 2288 1111 2.06 29.100 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R F 533 2075 2150 958 2.24 32.500 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R F 529 2133 1546 536 2.88 14.700 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R F 526 2092 1976 649 3.04 32.700 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R F 522 2030 2152 478 4.50 34.000 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R F 519 2010 851 198 4.30 25.800 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R F 515 2044 1428 363 3.93 29.300 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R F 510 2041 1464 875 1.67 23.800 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R F 500 2098 1885 332 5.68 25.300 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R F 494 1683 762 253 3.01 45.900 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R F 493 1559 1518 394 3.85 28.700 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R F 492 1676 1891 895 2.11 21.600 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R F 484 1733 1349 782 1.73 33.800 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R F 483 1576 2320 1198 1.94 37.400 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R F 480 1676 633 185 3.42 25.200 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R M 585 2933 112 454 0.25 ND 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R M 573 3107 196 326 0.60 ND 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R M 565 2556 85 450 0.19 ND 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R M 560 2164 227 1136 0.20 ND 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R M 540 2033 129 788 0.16 ND 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R M 532 1972 91 701 0.13 ND 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R M 515 1837 75 612 0.12 ND 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R M 510 1846 31 723 0.04 ND 2.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R M 510 1824 211 1733 0.12 ND 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R M 502 1678 118 609 0.19 ND 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R M 496 1611 131 183 0.72 ND 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R M 492 1623 103 614 0.17 ND 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R M 460 1312 149 284 0.53 ND 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R M 455 1262 51 152 0.34 ND 3.0

So Platte Riv @ Kersey, CO NMC 1994 Sept R M 440 1077 66 1353 0.05 ND 3.0

So Platte Riv below No Platte, NE NMC 1995 Sept R F 680 4000 347 251 1.38 9.296 3.0

So Platte Riv below No Platte, NE NMC 1995 Sept R F 591 3000 1959 1988 0.99 ND 3.0

So Platte Riv below No Platte, NE NMC 1995 Sept R F 555 2300 1258 1158 1.09 43.952 2.0

So Platte Riv below No Platte, NE NMC 1995 Sept R F 500 1700 232 93 2.49 2.935 2.0
Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

So Platte Riv below No Platte, NE NMC 1995 Sept R M 577 2350 179 1581 0.11 ND 2.0

So Platte Riv below No Platte, NE NMC 1995 Sept R M 550 2150 276 2543 0.11 ND 1.0

So Platte Riv below No Platte, NE NMC 1995 Sept R M 544 2100 153 1658 0.09 ND 2.0

So Platte Riv below No Platte, NE NMC 1995 Sept R M 535 2200 103 744 0.14 ND 2.0

So Platte Riv below No Platte, NE NMC 1995 Sept R M 525 1600 58 271 0.21 10.981 1.0

So Platte Riv below No Platte, NE NMC 1995 Sept R M 521 1950 881 5303 0.17 ND 2.0

So Platte Riv below No Platte, NE NMC 1995 Sept R M 520 1650 101 707 0.14 ND 2.0

So Platte Riv below No Platte, NE NMC 1995 Sept R M 513 1700 92 2296 0.04 ND 1.0

So Platte Riv below No Platte, NE NMC 1995 Sept R M 512 2000 124 998 0.12 ND 1.0

So Platte Riv below No Platte, NE NMC 1995 Sept R M 495 1500 301 1052 0.29 ND 1.0

So Platte Riv below No Platte, NE NMC 1995 Sept R M 491 1450 303 647 0.47 ND 1.0

So Platte Riv below No Platte, NE NMC 1995 Sept R M 481 1750 239 618 0.39 ND 2.0

So Platte Riv below No Platte, NE NMC 1995 Sept R M 477 1500 77 111 0.69 ND 1.0

So Platte Riv below No Platte, NE NMC 1995 Sept R M 471 1400 39 4892 0.01 ND 1.0

So Platte Riv below No Platte, NE NMC 1995 Sept R M 471 1700 609 2714 0.22 ND 1.0

So Skunk Riv @ Oskaloosa, IA NMC 1995 Sept R F 482 1355 1245 223 5.58 2.683 3.0

So Skunk Riv @ Oskaloosa, IA NMC 1995 Sept R F 472 1336 1241 274 4.53 1.962 3.0

So Skunk Riv @ Oskaloosa, IA NMC 1995 Sept R F 437 1013 1383 397 3.48 2.074 3.0

So Skunk Riv @ Oskaloosa, IA NMC 1995 Sept R F 430 991 893 262 3.41 0.027 1.0

So Skunk Riv @ Oskaloosa, IA NMC 1995 Sept R F 423 1071 3276 336 9.75 4.999 3.0

So Skunk Riv @ Oskaloosa, IA NMC 1995 Sept R F 414 820 323 386 0.84 2.873 3.0

So Skunk Riv @ Oskaloosa, IA NMC 1995 Sept R F 410 883 1295 121 10.70 2.587 3.0

So Skunk Riv @ Oskaloosa, IA NMC 1995 Sept R F 385 772 1341 310 4.33 3.238 2.0

So Skunk Riv @ Oskaloosa, IA NMC 1995 Sept R F 382 727 2385 295 8.08 4.101 2.0

So Skunk Riv @ Oskaloosa, IA NMC 1995 Sept R F 381 717 1302 263 4.95 2.355 3.0

So Skunk Riv @ Oskaloosa, IA NMC 1995 Sept R M 523 1377 517 310 1.67 ND 2.0

So Skunk Riv @ Oskaloosa, IA NMC 1995 Sept R M 460 1109 343 1249 0.27 ND 2.0

So Skunk Riv @ Oskaloosa, IA NMC 1995 Sept R M 445 1081 366 341 1.07 ND 2.0

So Skunk Riv @ Oskaloosa, IA NMC 1995 Sept R M 432 324 890 343 2.59 ND 2.0

So Skunk Riv @ Oskaloosa, IA NMC 1995 Sept R M 425 932 47 252 0.19 ND 2.0

So Skunk Riv @ Oskaloosa, IA NMC 1995 Sept R M 420 1768 252 254 0.99 ND 2.0

So Skunk Riv @ Oskaloosa, IA NMC 1995 Sept R M 418 803 447 243 1.84 ND 2.0

So Skunk Riv @ Oskaloosa, IA NMC 1995 Sept R M 408 907 174 487 0.36 ND 2.0

So Skunk Riv @ Oskaloosa, IA NMC 1995 Sept R M 404 805 85 204 0.42 ND 2.0

So Skunk Riv @ Oskaloosa, IA NMC 1995 Sept R M 385 717 346 517 0.67 ND 2.0

Steele Bayou @ Rolling Fork, MS SMC 1995 Aug R F 495 1638 1149 305 3.767 0.140 3.0

Steele Bayou @ Rolling Fork, MS SMC 1995 Aug R F 456 1175 1165 339 3.437 0.006 3.0

Steele Bayou @ Rolling Fork, MS SMC 1995 Aug R F 422 1027 2104 169 12.450 1.349 3.0

Steele Bayou @ Rolling Fork, MS SMC 1995 Aug R F 400 821 338 208 1.625 0.017 3.0

Steele Bayou @ Rolling Fork, MS SMC 1995 Aug R F 396 848 1612 271 5.948 0.115 3.0

Table 4 71
72 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Steele Bayou @ Rolling Fork, MS SMC 1995 Aug R F 394 946 2317 301 7.698 2.877 3.0

Steele Bayou @ Rolling Fork, MS SMC 1995 Aug R F 347 603 2111 308 6.854 2.003 3.0

Steele Bayou @ Rolling Fork, MS SMC 1995 Aug R F 332 479 1090 224 4.866 0.975 3.0

Steele Bayou @ Rolling Fork, MS SMC 1995 Aug R M 460 1347 104 158 0.658 ND 2.0

Steele Bayou @ Rolling Fork, MS SMC 1995 Aug R M 430 959 62 892 0.070 ND 2.0

Steele Bayou @ Rolling Fork, MS SMC 1995 Aug R M 411 779 137 281 0.488 ND 1.0

Steele Bayou @ Rolling Fork, MS SMC 1995 Aug R M 410 910 135 961 0.140 ND 2.0

Steele Bayou @ Rolling Fork, MS SMC 1995 Aug R M 400 658 100 566 0.177 ND 2.0

Steele Bayou @ Rolling Fork, MS SMC 1995 Aug R M 395 711 92 501 0.184 ND 2.0

Steele Bayou @ Rolling Fork, MS SMC 1995 Aug R M 380 731 539 369 1.461 ND 2.0

Steele Bayou @ Rolling Fork, MS SMC 1995 Aug R M 362 669 321 589 0.545 ND 2.0

Steele Bayou @ Rolling Fork, MS SMC 1995 Aug R M 362 649 111 753 0.147 ND 2.0

Steele Bayou @ Rolling Fork, MS SMC 1995 Aug R M NA NA 156 460 0.339 ND 2.0

Steele Bayou @ Rolling Fork, MS NMC 1994 Aug R F 742 5600 1423 925 1.538 28.400 3.0

Sugar Ck nr New Palistine, IN NMC 1994 Aug R F 682 5160 601 383 1.569 16.900 3.0

Sugar Ck nr New Palistine, IN NMC 1994 Aug R F 672 3426 656 1257 0.522 NA 3.0

Sugar Ck nr New Palistine, IN NMC 1994 Aug R F 670 4500 827 670 1.234 31.600 3.0

Sugar Ck nr New Palistine, IN NMC 1994 Aug R F 618 3600 1245 797 1.562 33.300 3.0

Sugar Ck nr New Palistine, IN NMC 1994 Aug R F 600 3750 525 737 0.712 33.400 3.0

Sugar Ck nr New Palistine, IN NMC 1994 Aug R F 567 2280 504 436 1.156 24.100 3.0

Sugar Ck nr New Palistine, IN NMC 1994 Aug R F 565 2151 924 1146 0.806 29.200 3.0

Sugar Ck nr New Palistine, IN NMC 1994 Aug R F 524 1775 577 942 0.613 27.100 3.0

Sugar Ck nr New Palistine, IN NMC 1994 Aug R M 665 3020 891 1004 0.887 ND 1.0

Sugar Ck nr New Palistine, IN NMC 1994 Aug R M 590 2200 488 726 0.672 ND 1.0

Sugar Ck nr New Palistine, IN NMC 1994 Aug R M 588 2340 946 986 0.959 ND 1.0

Sugar Ck nr New Palistine, IN NMC 1994 Aug R M 580 2540 871 1524 0.572 ND 1.0

Sugar Ck nr New Palistine, IN NMC 1994 Aug R M 575 2455 436 1474 0.296 ND 1.0

Sugar Ck nr New Palistine, IN NMC 1994 Aug R M 554 2080 502 1306 0.384 ND 2.0

Sugar Ck nr New Palistine, IN NMC 1994 Aug R M 551 1797 1491 1619 0.921 ND 2.0

Sugar Ck nr New Palistine, IN NMC 1995 June R F 674 5600 566 1344 0.421 5.971 4.0

Sugar Ck nr New Palistine, IN NMC 1995 June R F 617 3515 462 215 2.149 5.928 2.0

Sugar Ck nr New Palistine, IN NMC 1995 June R F 587 3360 314 701 0.448 5.241 3.0

Sugar Ck nr New Palistine, IN NMC 1995 June R F 573 2800 380 505 0.752 5.952 2.0

Sugar Ck nr New Palistine, IN NMC 1995 June R F 497 5195 585 659 0.888 6.894 3.0

Sugar Ck nr New Palistine, IN NMC 1995 June R F 494 1590 181 479 0.378 4.215 3.0

Sugar Ck nr New Palistine, IN NMC 1995 June R M 599 3110 308 1003 0.307 ND 3.0

Sugar Ck nr New Palistine, IN NMC 1995 June R M 588 2890 146 376 0.388 ND 3.0

Sugar Ck nr New Palistine, IN NMC 1995 June R M 583 3235 204 1092 0.187 ND 3.0

Sugar Ck nr New Palistine, IN NMC 1995 June R M 574 2675 108 518 0.208 ND 3.0

Sugar Ck nr New Palistine, IN NMC 1995 June R M 572 2955 101 463 0.218 ND 3.0
Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Tennessee Riv @ Savannah, TN SMC 1995 Oct NR F 662 3600 1485 554 2.681 ND 2

Tennessee Riv @ Savannah, TN SMC 1995 Oct NR F 604 3400 1142 370 3.086 ND 3

Tennessee Riv @ Savannah, TN SMC 1995 Oct NR F 505 2200 2582 1249 2.067 1.240 3

Tennessee Riv @ Savannah, TN SMC 1995 Oct NR F 490 1700 1559 517 3.015 0.624 3

Tennessee Riv @ Savannah, TN SMC 1995 Oct NR F 455 1200 583 869 0.671 ND 2

Tennessee Riv @ Savannah, TN SMC 1995 Oct NR M 570 2500 555 2783 0.199 ND 2

Tennessee Riv @ Savannah, TN SMC 1995 Oct NR M 560 2300 138 317 0.435 ND 2

Tennessee Riv @ Savannah, TN SMC 1995 Oct NR M 545 2500 419 1333 0.314 ND 3

Tennessee Riv @ Savannah, TN SMC 1995 Oct NR M 540 2000 319 1767 0.181 ND 1.5

Tennessee Riv @ Savannah, TN SMC 1995 Oct NR M 510 1600 467 1720 0.272 ND 2

Tennessee Riv @ Savannah, TN SMC 1995 Oct NR M 500 1600 374 2767 0.135 ND 2

Tennessee Riv @ Savannah, TN SMC 1995 Oct NR M 492 1500 209 1249 0.167 ND 2

Tennessee Riv @ Savannah, TN SMC 1995 Oct NR M 478 1400 365 523 0.698 ND 2

Tennessee Riv @ Savannah, TN SMC 1995 Oct NR M 460 1200 319 1337 0.239 ND 2

Tennessee Riv @ Savannah, TN SMC 1995 Oct NR M 395 850 1276 264 4.833 ND 2

Tensas Riv @ Tendell, LA SMC 1995 Aug R F 554 2380 925 258 3.585 0.202 3.0

Tensas Riv @ Tendell, LA SMC 1995 Aug R F 519 2047 166 203 0.818 0.786 3.0

Tensas Riv @ Tendell, LA SMC 1995 Aug R F 500 1592 2122 1209 1.755 4.256 3.0

Tensas Riv @ Tendell, LA SMC 1995 Aug R F 420 1054 851 379 2.245 4.960 3.0

Tensas Riv @ Tendell, LA SMC 1995 Aug R F 404 868 822 154 5.338 5.641 3.0

Tensas Riv @ Tendell, LA SMC 1995 Aug R M 555 1941 792 355 2.231 ND 2.0

Tensas Riv @ Tendell, LA SMC 1995 Aug R M 545 1706 415 236 1.758 ND 2.0

Tensas Riv @ Tendell, LA SMC 1995 Aug R M 540 1792 770 111 6.937 ND 2.0

Tensas Riv @ Tendell, LA SMC 1995 Aug R M 535 1903 121 188 0.644 ND 1.0

Tensas Riv @ Tendell, LA SMC 1995 Aug R M 532 1704 361 171 2.111 ND 2.0

Tensas Riv @ Tendell, LA SMC 1995 Aug R M 518 1690 71 264 0.269 ND 2.0

Tensas Riv @ Tendell, LA SMC 1995 Aug R M 514 2065 161 263 0.612 ND 2.0

Tensas Riv @ Tendell, LA SMC 1995 Aug R M 482 1475 191 292 0.654 ND 2.0

Tensas Riv @ Tendell, LA SMC 1995 Aug R M 445 1298 376 308 1.221 ND 2.0

Trinity Riv below Dallas, TX SMC 1994 Dec NR F 690 5992 1527 413 3.70 7.100 3.0

Trinity Riv below Dallas, TX SMC 1994 Dec NR F 650 4950 1036 265 3.91 12.300 3.0

Trinity Riv below Dallas, TX SMC 1994 Dec NR F 640 3466 2110 480 4.40 7.000 3.0

Trinity Riv below Dallas, TX SMC 1994 Dec NR F 590 3434 947 412 2.30 12.800 3.0

Trinity Riv below Dallas, TX SMC 1994 Dec NR F 559 2690 2922 654 4.47 10.500 3.0

Trinity Riv below Dallas, TX SMC 1994 Dec NR M 655 3155 128 521 0.25 ND 2.0

Trinity Riv below Dallas, TX SMC 1994 Dec NR M 580 3140 388 732 0.53 ND 2.0

Trinity Riv below Dallas, TX SMC 1994 Dec NR M 575 2861 324 654 0.50 ND 1.0

Trinity Riv below Dallas, TX SMC 1994 Dec NR M 550 2718 373 557 0.67 ND 1.0

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR F 656 3750 1471 1044 1.409 2.138 3

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR F 609 2800 2028 1187 1.709 3.109 3

Table 4 73
74 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR F 555 2200 1237 1176 1.052 1.397 3

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR F 554 2350 2303 942 2.445 1.462 3

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR F 538 1950 1401 199 7.040 1.973 3

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR F 505 1750 1965 1746 1.125 1.134 3

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR F 495 1700 1607 754 2.131 2.012 3

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR F 480 1450 1313 285 4.607 2.421 3

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR F 470 1400 1221 1153 1.059 1.679 3

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR F NA NA 1674 1115 1.501 0.712 3

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR M 530 2175 355 1765 0.201 ND 3

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR M 530 1850 689 1905 0.362 ND 3

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR M 505 1700 1148 2116 0.543 ND 3

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR M 494 1400 900 813 1.107 0.006 3

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR M 480 1400 982 2028 0.484 ND 3

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR M 480 1400 763 1944 0.392 ND 3

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR M 462 1275 1076 1683 0.639 0.009 3

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR M 452 1300 300 652 0.460 ND 3

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR M 433 1000 759 914 0.830 ND 3

Virgin Riv inflo to L Mead, NV SW 1996 May R F 469 1390 861 265 3.249 2.423 2.0

Virgin Riv inflo to L Mead, NV SW 1996 May R F 461 1417 921 318 2.896 0.381 3.0

Virgin Riv inflo to L Mead, NV SW 1996 May R F 453 1582 1120 192 5.833 1.697 3.0

Virgin Riv inflo to L Mead, NV SW 1996 May R F 441 1224 761 268 2.840 1.740 3.0

Virgin Riv inflo to L Mead, NV SW 1996 May R F 427 1129 512 112 4.571 1.374 2.0

Virgin Riv inflo to L Mead, NV SW 1996 May R F 417 947 219 212 1.033 0.581 3.0

Virgin Riv inflo to L Mead, NV SW 1996 May R F 402 947 694 175 3.966 0.585 4.0

Virgin Riv inflo to L Mead, NV SW 1996 May R F 392 870 321 179 1.793 0.815 3.0

Virgin Riv inflo to L Mead, NV SW 1996 May R F 388 719 257 171 1.503 2.071 3.0

Virgin Riv inflo to L Mead, NV SW 1996 May R F 385 886 831 227 3.661 0.871 2.0

Virgin Riv inflo to L Mead, NV SW 1996 May R M 518 1830 117 361 0.324 ND 2.0

Virgin Riv inflo to L Mead, NV SW 1996 May R M 491 1263 71 229 0.310 ND 3.0

Virgin Riv inflo to L Mead, NV SW 1996 May R M 491 1256 73 182 0.401 0.005 1.0

Virgin Riv inflo to L Mead, NV SW 1996 May R M 487 1383 107 217 0.493 ND 3.0

Virgin Riv inflo to L Mead, NV SW 1996 May R M 460 1066 88 174 0.506 ND 2.0

Virgin Riv inflo to L Mead, NV SW 1996 May R M 454 1376 134 268 0.500 ND 3.0

Virgin Riv inflo to L Mead, NV SW 1996 May R M 426 946 139 235 0.591 ND 3.0

Virgin Riv inflo to L Mead, NV SW 1996 May R M 422 849 94 326 0.288 ND 1.0

Virgin Riv inflo to L Mead, NV SW 1996 May R M 421 815 205 798 0.257 0.005 2.0

Virgin Riv inflo to L Mead, NV SW 1996 May R M 406 771 76 127 0.598 0.005 3.0

Wabash Riv @ New Harmony, IN NMC 1995 Oct NR M 450 1700 405 246 1.65 ND 3

Wabash Riv @ N Harmony, IN NMC 1995 Oct NR F 718 7181 589 101 5.83 2.924 3

Wabash Riv @ N Harmony, IN NMC 1995 Oct NR F 600 2700 463 92 5.03 3.270 0
Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Wabash Riv @ N Harmony, IN NMC 1995 Oct NR F 585 3300 358 131 2.73 7.453 1

Wabash Riv @ N Harmony, IN NMC 1995 Oct NR F 574 2600 765 100 7.65 ND 2

Wabash Riv @ N Harmony, IN NMC 1995 Oct NR F 563 2800 537 181 2.97 1.230 3

Wabash Riv @ N Harmony, IN NMC 1995 Oct NR F 539 2500 719 123 5.85 2.726 3

Wabash Riv @ N Harmony, IN NMC 1995 Oct NR F 530 2190 920 77 11.95 1.505 3

Wabash Riv @ N Harmony, IN NMC 1995 Oct NR F 506 2000 429 104 4.13 2.139 3

Wabash Riv @ N Harmony, IN NMC 1995 Oct NR F 447 1800 351 69 5.09 3.606 3

Wabash Riv @ N Harmony, IN NMC 1995 Oct NR M 592 2800 544 1200 0.45 ND 2

Wabash Riv @ N Harmony, IN NMC 1995 Oct NR M 570 2600 310 339 0.91 ND 2

Wabash Riv @ N Harmony, IN NMC 1995 Oct NR M 565 2600 430 534 0.81 ND 2

Wabash Riv @ N Harmony, IN NMC 1995 Oct NR M 561 2400 348 2202 0.16 ND 2

Wabash Riv @ N Harmony, IN NMC 1995 Oct NR M 544 2500 450 391 1.15 ND 3

Wabash Riv @ N Harmony, IN NMC 1995 Oct NR M 542 2300 347 302 1.15 ND 3

Wabash Riv @ N Harmony, IN NMC 1995 Oct NR M 465 1300 455 1646 0.28 ND 2

Weber Riv @ Ogden Bay UT SW 1998 Aug R F 720 5000 1159 687 1.687 0.672 3

Weber Riv @ Ogden Bay UT SW 1998 Aug R F 718 5500 591 370 1.597 1.723 3

Weber Riv @ Ogden Bay UT SW 1998 Aug R F 627 3000 1259 603 2.088 3.125 3

Weber Riv @ Ogden Bay UT SW 1998 Aug R F 572 2306 575 273 2.106 1.625 3

Weber Riv @ Ogden Bay UT SW 1998 Aug R F 498 1520 612 353 1.734 0.275 2

Weber Riv @ Ogden Bay UT SW 1998 Aug R M 552 2060 263 521 0.505 ND 1

Weber Riv @ Ogden Bay UT SW 1998 Aug R M 501 1575 690 736 0.938 ND 1

Weber Riv @ Ogden Bay UT SW 1998 Aug R M 472 1123 515 914 0.563 ND 3

Weber Riv @ Ogden Bay UT SW 1998 Aug R M 421 957 505 1317 0.383 0.018 2

Weber Riv @ Ogden Bay UT SW 1998 Aug R M 380 740 546 1231 0.444 ND 2

White Riv @ Hazelton, IN NMC 1994 Sept R F 668 4530 2125 1474 1.44 32.900 3.0

White Riv @ Hazelton, IN NMC 1994 Sept R F 638 3834 948 1342 0.71 41.100 3.0

White Riv @ Hazelton, IN NMC 1994 Sept R F 620 3227 1559 791 1.97 34.700 3.0

White Riv @ Hazelton, IN NMC 1994 Sept R F 617 3287 1521 325 4.68 30.800 3.0

White Riv @ Hazelton, IN NMC 1994 Sept R F 612 3225 2087 1152 1.81 24.400 3.0

White Riv @ Hazelton, IN NMC 1994 Sept R F 600 3300 1939 1429 1.36 23.800 3.0

White Riv @ Hazelton, IN NMC 1994 Sept R F 589 2866 884 862 1.03 27.000 3.0

White Riv @ Hazelton, IN NMC 1994 Sept R F 587 2219 2799 794 3.53 32.500 3.0

White Riv @ Hazelton, IN NMC 1994 Sept R F 575 3300 1328 1665 0.80 31.000 3.0

White Riv @ Hazelton, IN NMC 1994 Sept R F 564 2208 1283 681 1.88 26.100 3.0

White Riv @ Hazelton, IN NMC 1994 Sept R F 550 2292 2485 1126 2.21 28.900 3.0

White Riv @ Hazelton, IN NMC 1994 Sept R F 539 2463 1957 1881 1.04 21.400 3.0

White Riv @ Hazelton, IN NMC 1994 Sept R F 538 2614 2313 1969 1.17 29.200 3.0

White Riv @ Hazelton, IN NMC 1994 Sept R F 530 2122 1890 1621 1.17 31.500 3.0

White Riv @ Hazelton, IN NMC 1994 Sept R F 508 1800 1130 440 2.57 33.900 3.0

White Riv @ Hazelton, IN NMC 1994 Sept R F 484 1671 1248 1091 1.14 37.200 3.0

Table 4 75
76 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

White Riv @ Hazelton, IN NMC 1994 Sept R F 446 1314 1774 1373 1.29 29.700 3.0

White Riv @ Hazelton, IN NMC 1994 Sept R M 638 1909 176 1140 0.15 ND 2.0

White Riv @ Hazelton, IN NMC 1994 Sept R M 610 2200 1145 2604 0.44 ND 1.0

White Riv @ Hazelton, IN NMC 1994 Sept R M 598 2530 430 2441 0.18 ND 2.0

White Riv @ Hazelton, IN NMC 1994 Sept R M 589 2045 655 1417 0.46 0.010 1.0

White Riv @ Hazelton, IN NMC 1994 Sept R M 573 2756 546 1515 0.36 0.830 1.0

White Riv @ Hazelton, IN NMC 1994 Sept R M 568 2120 733 2514 0.29 ND 1.0

White Riv @ Hazelton, IN NMC 1994 Sept R M 565 2350 693 1284 0.54 0.050 1.0

White Riv @ Hazelton, IN NMC 1994 Sept R M 553 2050 1728 2075 0.83 ND 2.0

White Riv @ Hazelton, IN NMC 1994 Sept R M 551 2135 659 2288 0.29 ND 1.0

White Riv @ Hazelton, IN NMC 1994 Sept R M 550 1658 564 1314 0.44 ND 1.0

White Riv @ Hazelton, IN NMC 1994 Sept R M 543 2130 652 2580 0.25 0.020 1.0

White Riv @ Hazelton, IN NMC 1994 Sept R M 519 1928 389 2448 0.16 ND 1.0

White Riv @ Hazelton, IN NMC 1994 Sept R M 519 1736 737 2446 0.30 ND 1.0

White Riv @ Hazelton, IN NMC 1994 Sept R M 510 1827 1049 1067 0.98 ND 1.0

White Riv @ Hazelton, IN NMC 1994 Sept R M 500 1877 869 1720 0.51 0.060 1.0

White Riv @ Hazelton, IN NMC 1994 Sept R M 490 1691 443 2416 0.18 ND 1.0

White Riv @ Hazelton, IN NMC 1994 Sept R M 466 1556 598 836 0.72 ND 1.0

White Riv @ Hazelton, IN NMC 1994 Sept R M 465 1547 396 755 0.53 ND 2.0

White Riv @ Hazelton, IN NMC 1994 Sept R M 457 1389 1324 1743 0.76 ND 1.0

White Riv @ Hazelton, IN NMC 1995 June R F 667 4760 369 225 1.640 46.332 3.0

White Riv @ Hazelton, IN NMC 1995 June R F 642 5239 862 212 4.066 27.875 1.0

White Riv @ Hazelton, IN NMC 1995 June R F 636 3916 882 215 4.102 36.650 2.0

White Riv @ Hazelton, IN NMC 1995 June R F 631 3568 575 318 1.808 41.443 3.0

White Riv @ Hazelton, IN NMC 1995 June R F 625 3185 774 93 8.323 63.292 2.0

White Riv @ Hazelton, IN NMC 1995 June R F 579 5175 687 553 1.242 74.175 3.0

White Riv @ Hazelton, IN NMC 1995 June R F 576 2496 340 294 1.156 33.309 4.0

White Riv @ Hazelton, IN NMC 1995 June R F 551 2184 261 355 0.735 124.800 3.0

White Riv @ Hazelton, IN NMC 1995 June R F 526 2924 383 160 2.394 84.320 3.0

White Riv @ Hazelton, IN NMC 1995 June R F 524 1863 601 646 0.930 83.153 3.0

White Riv @ Hazelton, IN NMC 1995 June R F 510 1964 434 117 3.709 85.648 2.0

White Riv @ Hazelton, IN NMC 1995 June R M 653 3718 229 924 0.248 ND 3.0

White Riv @ Hazelton, IN NMC 1995 June R M 648 3252 359 191 1.880 ND 3.0

White Riv @ Hazelton, IN NMC 1995 June R M 616 3722 267 234 1.141 ND 3.0

White Riv @ Hazelton, IN NMC 1995 June R M 605 2935 441 660 0.668 ND 3.0

White Riv @ Hazelton, IN NMC 1995 June R M 581 2217 357 617 0.579 ND 3.0

White Riv @ Hazelton, IN NMC 1995 June R M 581 2509 255 484 0.527 11.956 3.0

White Riv @ Hazelton, IN NMC 1995 June R M 570 2595 232 1267 0.183 ND 3.0

White Riv @ Hazelton, IN NMC 1995 June R M 564 2645 391 1176 0.332 ND 3.0

White Riv @ Hazelton, IN NMC 1995 June R M 517 2180 306 335 0.913 ND 2.0
Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

White Riv @ Hazelton, IN NMC 1995 June R M 512 2180 351 347 1.012 ND 3.0

White Riv @ Hazelton, IN NMC 1995 June R M 511 2240 271 881 0.308 ND 3.0

White Riv @ Hazelton, IN NMC 1995 June R M 504 1990 108 151 0.715 ND 1.0

White Riv @ Hazelton, IN NMC 1995 June R M 504 2490 182 436 0.417 1.389 3.0

White Riv @ Hazelton, IN NMC 1995 June R M 491 2020 198 844 0.235 ND 3.0

White Riv @ Hazelton, IN NMC 1995 June R M 490 1635 363 377 0.963 ND 3.0

White Riv @ Hazelton, IN NMC 1995 June R M 486 1586 295 228 1.294 ND 3.0

White Riv @ Hazelton, IN NMC 1995 June R M 485 1747 143 919 0.156 ND 3.0

White Riv @ Hazelton, IN NMC 1995 June R M 471 1524 159 432 0.368 ND 3.0

White Riv @ Hazelton, IN NMC 1995 June R M 458 1340 368 380 0.968 ND 1.0

White Riv nr DeValls Bluff, AR SMC 1995 Sept R F 700 4690 2265 611 3.71 2.364 3

White Riv nr DeValls Bluff, AR SMC 1995 Sept R F 645 4084 855 1925 0.44 ND 3

White Riv nr DeValls Bluff, AR SMC 1995 Sept R F 600 3142 822 274 3.00 ND 3

White Riv nr DeValls Bluff, AR SMC 1995 Sept R F 590 3264 439 187 2.35 0.011 3

White Riv nr DeValls Bluff, AR SMC 1995 Sept R F 566 2845 1735 369 4.70 ND 3

White Riv nr DeValls Bluff, AR SMC 1995 Sept R F 553 2611 465 419 1.11 ND 3

White Riv nr DeValls Bluff, AR SMC 1995 Sept R F 540 2052 1151 172 6.69 4.280 3

White Riv nr DeValls Bluff, AR SMC 1995 Sept R M 674 4996 551 248 2.22 ND 2

White Riv nr DeValls Bluff, AR SMC 1995 Sept R M 657 4417 501 3226 0.16 ND 2

White Riv nr DeValls Bluff, AR SMC 1995 Sept R M 632 3470 358 949 0.38 ND 1

White Riv nr DeValls Bluff, AR SMC 1995 Sept R M 581 2860 367 2036 0.18 ND 2

White Riv nr DeValls Bluff, AR SMC 1995 Sept R M 571 2673 945 1657 0.57 ND 2

White Riv nr DeValls Bluff, AR SMC 1995 Sept R M 560 2428 474 311 1.52 ND 2

White Riv nr DeValls Bluff, AR SMC 1995 Sept R M 526 2003 574 2338 0.25 ND 1

White Riv nr DeValls Bluff, AR SMC 1995 Sept R M 523 1867 461 769 0.60 ND 1

White Riv nr DeValls Bluff, AR SMC 1995 Sept R M 510 1867 428 532 0.80 ND 3

White Riv nr DeValls Bluff, AR SMC 1995 Sept R M 430 1147 689 579 1.19 ND 2

Willamette Riv @ Oregon Cty, OR NW 1997 Nov NR F 771 8000 899 222 4.05 0.110 2

Willamette Riv @ Oregon Cty, OR NW 1997 Nov NR F 765 6800 439 82 5.35 0.994 2

Willamette Riv @ Oregon Cty, OR NW 1997 Nov NR F 611 3800 183 129 1.42 0.735 2

Willamette Riv @ Oregon Cty, OR NW 1997 Nov NR F 562 2650 789 349 2.26 0.067 2

Willamette Riv @ Oregon Cty, OR NW 1997 Nov NR M 677 3800 63 419 0.15 ND 3

Willamette Riv @ Oregon Cty, OR NW 1997 Nov NR M 645 3650 257 305 0.84 0.010 3

Willamette Riv @ Oregon Cty, OR NW 1997 Nov NR M 604 3700 789 257 3.07 ND 3

Willamette Riv @ Oregon Cty, OR NW 1997 Nov NR M 580 2750 273 276 0.99 ND 3

Willamette Riv @ Oregon Cty, OR NW 1997 Nov NR M 570 2800 179 194 0.92 ND 3

Willamette Riv @ Oregon Cty, OR NW 1997 Nov NR M 555 2350 426 382 1.12 ND 3

Willamette Riv @ Oregon Cty, OR NW 1997 Nov NR M 535 2100 414 302 1.37 ND 3

Willamette Riv @ Oregon Cty, OR NW 1997 Nov NR M 520 1867 256 147 1.74 ND 3

Willamette Riv @ Oregon Cty, OR NW 1997 Nov NR M 481 1351 197 556 0.35 ND 3

Table 4 77
78 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Willamette Riv @ Oregon Cty, OR NW 1997 Nov NR M 465 1494 882 324 2.72 ND 3

Willamette Riv below Portland, OR NW 1995 July R F 565 2983 822 215 3.823 118.839 3.0

Willamette Riv below Portland, OR NW 1995 July R F 554 2724 1176 1042 1.129 52.296 3.0

Willamette Riv below Portland, OR NW 1995 July R F 533 2354 804 168 4.786 20.342 3.0

Willamette Riv below Portland, OR NW 1995 July R F 533 2254 505 843 0.599 23.752 3.0

Willamette Riv below Portland, OR NW 1995 July R F 521 2150 621 309 2.010 11.689 2.0

Willamette Riv below Portland, OR NW 1995 July R F 512 1910 577 241 2.394 73.179 2.0

Willamette Riv below Portland, OR NW 1995 July R F 509 1844 249 695 0.358 60.231 2.0

Willamette Riv below Portland, OR NW 1995 July R F 507 1706 2069 410 5.046 45.768 3.0

Willamette Riv below Portland, OR NW 1995 July R F 501 2209 560 681 0.822 56.516 3.0

Willamette Riv below Portland, OR NW 1995 July R F 496 1760 325 1124 0.289 13.902 3.0

Willamette Riv below Portland, OR NW 1995 July R F 487 1846 1156 384 3.010 40.971 3.0

Willamette Riv below Portland, OR NW 1995 July R F 486 1646 1514 1036 1.461 17.885 2.0

Willamette Riv below Portland, OR NW 1995 July R F 482 1573 982 280 3.507 17.202 3.0

Willamette Riv below Portland, OR NW 1995 July R F 448 1375 687 487 1.411 66.651 3.0

Willamette Riv below Portland, OR NW 1995 July R F 428 1052 295 972 0.303 2.660 1.0

Willamette Riv below Portland, OR NW 1995 July R M 521 1996 659 3140 0.210 ND 2.0

Willamette Riv below Portland, OR NW 1995 July R M 505 1581 718 639 1.124 ND 3.0

Willamette Riv below Portland, OR NW 1995 July R M 486 1596 818 1087 0.753 ND 3.0

Willamette Riv below Portland, OR NW 1995 July R M 484 1375 392 986 0.398 ND 3.0

Willamette Riv below Portland, OR NW 1995 July R M 484 1494 448 1248 0.359 ND 3.0

Willamette Riv below Portland, OR NW 1995 July R M 477 1360 215 2178 0.099 ND 3.0

Willamette Riv below Portland, OR NW 1995 July R M 477 1492 635 1554 0.409 ND 3.0

Willamette Riv below Portland, OR NW 1995 July R M 462 1271 169 625 0.270 ND 3.0

Willamette Riv below Portland, OR NW 1995 July R M 450 1261 754 1177 0.641 ND 3.0

Willamette Riv below Portland, OR NW 1995 July R M 447 1384 627 975 0.643 ND 3.0

Willamette Riv below Portland, OR NW 1995 July R M 443 1156 642 2076 0.309 ND 2.0

Willamette Riv below Portland, OR NW 1995 July R M 442 1220 771 4310 0.179 ND 3.0

Willamette Riv below Portland, OR NW 1995 July R M 440 1252 471 695 0.678 ND 3.0

Willamette Riv below Portland, OR NW 1995 July R M 416 999 560 895 0.626 ND 3.0

Willamette Riv below Portland, OR NW 1995 July R M 379 690 538 509 1.057 ND 3.0

Wisconsin Riv @ Woodman, WI NMC 1995 Sept NR F 600 2600 2439 871 2.80 4.650 3

Wisconsin Riv @ Woodman, WI NMC 1995 Sept NR F 585 3000 1777 305 5.83 1.724 3

Wisconsin Riv @ Woodman, WI NMC 1995 Sept NR F 578 2200 855 621 1.38 3.101 3

Wisconsin Riv @ Woodman, WI NMC 1995 Sept NR F 556 2500 3449 869 3.97 4.537 3

Wisconsin Riv @ Woodman, WI NMC 1995 Sept NR F 555 2200 2632 668 3.94 3.447 3

Wisconsin Riv @ Woodman, WI NMC 1995 Sept NR F 548 2100 1134 447 2.54 2.806 3

Wisconsin Riv @ Woodman, WI NMC 1995 Sept NR F 535 2050 2782 285 9.76 2.690 3

Wisconsin Riv @ Woodman, WI NMC 1995 Sept NR F 533 2050 1214 226 5.37 1.672 3

Wisconsin Riv @ Woodman, WI NMC 1995 Sept NR F 520 1900 1617 208 7.77 3.458 3
Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Wisconsin Riv @ Woodman, WI NMC 1995 Sept NR F 508 1950 578 132 4.38 5.255 3

Wisconsin Riv @ Woodman, WI NMC 1995 Sept NR M 578 2400 1151 2688 0.43 0.005 1

Wisconsin Riv @ Woodman, WI NMC 1995 Sept NR M 544 1900 502 1345 0.37 ND 2

Wisconsin Riv @ Woodman, WI NMC 1995 Sept NR M 538 1900 395 2167 0.18 ND 1.5

Wisconsin Riv @ Woodman, WI NMC 1995 Sept NR M 535 2000 583 2893 0.20 ND 1.5

Wisconsin Riv @ Woodman, WI NMC 1995 Sept NR M 533 1900 1474 191 7.72 ND NA

Wisconsin Riv @ Woodman, WI NMC 1995 Sept NR M 518 1950 456 1291 0.35 ND 1

Wisconsin Riv @ Woodman, WI NMC 1995 Sept NR M 514 1950 293 1196 0.24 ND 1.5

Wisconsin Riv @ Woodman, WI NMC 1995 Sept NR M 510 1850 578 2770 0.21 ND 1

Wisconsin Riv @ Woodman, WI NMC 1995 Sept NR M 502 1550 295 694 0.43 ND 2

Wisconsin Riv @ Woodman, WI NMC 1995 Sept NR M 495 1450 694 2182 0.32 ND 1

Wisconsin Riv @ Woodman, WI NMC 1995 Sept NR M 494 1550 145 1244 0.12 ND 1

Wisconsin Riv @ Woodman, WI NMC 1995 Sept NR M 468 1400 416 1267 0.33 ND 1

Yakima Riv @ Granger, WA NW 1997 Oct NR F 615 2500 623 100 6.23 0.537 1

Yakima Riv @ Granger, WA NW 1997 Oct NR F 591 3150 1251 107 11.69 2.030 2

Yakima Riv @ Granger, WA NW 1997 Oct NR F 535 2150 1418 358 3.96 2.245 2

Yakima Riv @ Granger, WA NW 1997 Oct NR F 535 1678 587 167 3.51 1.741 2

Yakima Riv @ Granger, WA NW 1997 Oct NR F 534 1850 1146 440 2.60 2.642 2

Yakima Riv @ Granger, WA NW 1997 Oct NR F 524 1457 772 384 2.01 1.538 1

Yakima Riv @ Granger, WA NW 1997 Oct NR F 521 1900 1142 596 1.92 0.919 2

Yakima Riv @ Granger, WA NW 1997 Oct NR F 476 1635 1244 277 4.49 1.944 2

Yakima Riv @ Granger, WA NW 1997 Oct NR F 440 898 1037 415 2.50 1.944 2

Yakima Riv @ Granger, WA NW 1997 Oct NR F 428 966 1407 770 1.83 2.885 2

Yakima Riv @ Granger, WA NW 1997 Oct NR M 590 2500 207 123 1.68 ND 3

Yakima Riv @ Granger, WA NW 1997 Oct NR M 519 1889 88 612 0.14 ND 2

Yakima Riv @ Granger, WA NW 1997 Oct NR M 501 1370 683 712 0.96 ND 3

Yakima Riv @ Granger, WA NW 1997 Oct NR M 497 1450 113 851 0.13 ND 3

Yakima Riv @ Granger, WA NW 1997 Oct NR M 490 1339 401 711 0.56 ND 3

Yakima Riv @ Granger, WA NW 1997 Oct NR M 475 1255 211 220 0.96 ND 2

Yakima Riv @ Granger, WA NW 1997 Oct NR M 464 1273 783 1101 0.71 ND 3

Yakima Riv @ Granger, WA NW 1997 Oct NR M 462 1038 374 497 0.75 ND 3

Yakima Riv @ Granger, WA NW 1997 Oct NR M 446 1150 76 555 0.14 ND 3

Yakima Riv @ Granger, WA NW 1997 Oct NR M 444 1107 296 1124 0.26 ND 3

Yazoo Riv @ Redwood, MS SMC 1995 Aug R F 742 6200 356 478 0.745 1.473 3

Yazoo Riv @ Redwood, MS SMC 1995 Aug R F 660 3571 1391 427 3.258 ND 3

Yazoo Riv @ Redwood, MS SMC 1995 Aug R F 583 3380 673 593 1.135 ND 3

Yazoo Riv @ Redwood, MS SMC 1995 Aug R F 563 2805 1198 1246 0.961 0.820 2

Yazoo Riv @ Redwood, MS SMC 1995 Aug R F 497 2071 551 433 1.273 ND 2

Yazoo Riv @ Redwood, MS SMC 1995 Aug R M 657 3693 496 347 1.429 ND 2

Yazoo Riv @ Redwood, MS SMC 1995 Aug R M 566 2623 438 721 0.607 ND 2

Table 4 79
80 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 4. (Continued) All individual COMMON CARP values for biomarker and ancillary data from collections made at sites in U.S. 

waters from August 1994 through October 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Yazoo Riv @ Redwood, MS SMC 1995 Aug R M 560 2401 688 199 3.457 ND 2

Yazoo Riv @ Redwood, MS SMC 1995 Aug R M 550 2411 120 612 0.196 ND 2

Yazoo Riv @ Redwood, MS SMC 1995 Aug R M 542 2279 627 425 1.475 ND 2

Yazoo Riv @ Redwood, MS SMC 1995 Aug R M 520 2206 390 466 0.837 ND 2

Yazoo Riv @ Redwood, MS SMC 1995 Aug R M 297 407 924 487 1.897 ND 1

Yellowstone R @ Sidney, MT NMC 1995 Sept NR F 672 3550 1981 1274 1.55 1.661 3

Yellowstone R @ Sidney, MT NMC 1995 Sept NR F 662 3850 2939 297 9.90 1.857 3

Yellowstone R @ Sidney, MT NMC 1995 Sept NR F 556 1750 2119 91 23.29 0.810 3

Yellowstone R @ Sidney, MT NMC 1995 Sept NR F 519 1700 1184 286 4.14 2.522 3

Yellowstone R @ Sidney, MT NMC 1995 Sept NR F 473 1300 1563 501 3.12 0.947 3

Yellowstone R @ Sidney, MT NMC 1995 Sept NR F 468 1300 3350 674 4.97 1.676 3

Yellowstone R @ Sidney, MT NMC 1995 Sept NR F 458 1150 2505 240 10.44 ND 1

Yellowstone R @ Sidney, MT NMC 1995 Sept NR F 442 1050 2399 261 9.19 1.318 3

Yellowstone R @ Sidney, MT NMC 1995 Sept NR F 419 950 917 385 2.38 0.074 1

Yellowstone R @ Sidney, MT NMC 1995 Sept NR F 415 900 1691 52 32.52 0.571 2

Yellowstone R @ Sidney, MT NMC 1995 Sept NR F 339 500 1296 408 3.18 ND 2

Yellowstone R @ Sidney, MT NMC 1995 Sept NR F 328 450 1083 188 5.76 ND 1

Yellowstone R @ Sidney, MT NMC 1995 Sept NR M 619 2600 2918 4214 0.69 ND 1

Yellowstone R @ Sidney, MT NMC 1995 Sept NR M 619 2800 967 3811 0.25 ND 1

Yellowstone R @ Sidney, MT NMC 1995 Sept NR M 596 2450 758 2754 0.28 ND 1.5

Yellowstone R @ Sidney, MT NMC 1995 Sept NR M 566 2150 1198 2532 0.47 ND 1

Yellowstone R @ Sidney, MT NMC 1995 Sept NR M 562 2400 709 3389 0.21 ND 1.5

Yellowstone R @ Sidney, MT NMC 1995 Sept NR M 539 2200 1255 2136 0.59 ND 1

Yellowstone R @ Sidney, MT NMC 1995 Sept NR M 519 1800 736 1191 0.62 ND 2

Yellowstone R @ Sidney, MT NMC 1995 Sept NR M 390 800 1129 1644 0.69 ND 2
Table 5. (Continued) All individual LARGEMOUTH BASS values for biomarker and ancillary data from collections made at sites in U.S. 

waters from March 1994 through August 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR M 405 1000 280 421 0.6651 ND 2

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR M 400 1000 427 1563 0.2732 ND 2

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR M 380 800 396 661 0.5991 ND 2

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR M 375 800 327 1515 0.2158 ND 2

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR M 386 750 301 1942 0.1550 ND 2

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR M 354 700 402 1365 0.2945 ND 2

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR M 357 650 281 1313 0.2140 ND 3

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR M 378 700 422 755 0.5589 0.01 2

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR F 332 500 1094 149 7.3423 ND 3

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR F 464 1400 1268 337 3.7626 0.641 2

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR F 410 1150 2252 439 5.1298 0.939 3

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR F 495 1950 3386 299 11.3244 2.117 3

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR F 378 900 2396 513 4.6706 2.3 3

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR F 421 1400 705 114 6.1842 4.697 3

Arkansas Riv @ Keystone Res, OK SMC 1995 Dec NR F 378 825 978 121 8.0826 5.443 3

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR F 335 750 1432 488 2.9344 ND 2

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR F 309 600 1453 460 3.1587 0.01 2

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR F 333 775 475 111 4.2793 0.021 2

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR F 344 850 403 179 2.2514 0.043 2

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR F 330 600 758 278 2.7266 0.047 2

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR F 323 800 899 213 4.2207 0.131 2

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR F 336 750 514 181 2.8398 2 2

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR F 348 800 1883 1040 1.8106 8.833 2

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR M 349 875 544 1491 0.3649 ND 2

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR M 337 850 343 2541 0.1350 ND 2

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR M 343 825 330 561 0.5882 ND 2

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR M 336 825 482 1081 0.4459 ND 2

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR M 332 800 329 766 0.4295 ND 2

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR M 337 752 388 1187 0.3269 ND 2

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR M 327 750 535 1171 0.4569 ND 2

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR M 322 700 494 1368 0.3611 ND 2

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR M 328 660 493 2812 0.1753 ND 2

Arkansas Riv @ J Martin Res, CO NMC 1995 Oct NR M 295 550 537 711 0.7553 ND 2

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR F 392 980 498 429 1.1608 ND 2

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR F 305 397 308 144 2.1389 ND 2

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR F 315 391 235 329 0.7143 ND 2

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR F 405 970 749 319 2.3480 0.008 2

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR F 370 794 719 330 2.1788 0.009 2

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR F 525 2321 470 213 2.2066 0.018 2

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR F 415 1156 845 205 4.1220 0.019 2

Table 5 81

Table 5. All individual LARGEMOUTH BASS values for biomarker and ancillary data from collections made at sites in U.S. 

waters from March 1994 through August 1997.
82 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 5. (Continued) All individual LARGEMOUTH BASS values for biomarker and ancillary data from collections made at sites in U.S. 

waters from March 1994 through August 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR F 415 1145 249 285 0.8737 0.031 2

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR F 505 2308 458 277 1.6534 0.061 2

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR F 540 2400 766 348 2.2011 0.197 2

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR M 411 1251 550 1491 0.3689 ND 2

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR M 395 825 434 2597 0.1671 ND 2

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR M 380 783 523 622 0.8408 ND 2

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR M 358 640 249 1447 0.1721 ND 2

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR M 330 423 265 727 0.3645 ND 2

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR M 305 346 261 484 0.5393 ND 2

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR M 308 328 414 5144 0.0805 ND 2

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR M 278 273 347 319 1.0878 ND 2

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR M 262 200 263 448 0.5871 ND 2

Arkansas Riv @ Pine Bluff, AR SMC 1995 Oct NR M NA NA 277 1306 0.2121 ND 1

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR M 453 1500 466 1206 0.3864 ND 2

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR M 430 1300 842 1393 0.6045 ND 2

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR M 370 1250 469 252 1.8611 ND 1

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR M 410 1000 704 1420 0.4958 ND 2

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR M 410 950 745 909 0.8196 ND 2

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR M 380 800 683 641 1.0655 ND 3

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR M 410 750 341 214 1.5935 ND 2

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR M 357 550 574 619 0.9273 ND 2

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR M 313 450 604 503 1.2008 ND 2

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR M 410 889 1022 484 2.1116 2.408 2

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR F 455 1500 1198 398 3.0101 ND 3

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR F 390 800 1064 1402 0.7589 ND 3

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR F 385 750 2417 509 4.7485 ND 2

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR F 388 600 978 1069 0.9149 ND 3

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR F 361 800 518 87 5.9540 0.287 3

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR F 385 775 605 1999 0.3027 0.675 3

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR F 473 1800 918 904 1.0155 1.581 3

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR F 351 650 945 881 1.0726 2.804 3

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR F 490 1400 462 1045 0.4421 3.414 2

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR F 510 1900 1022 1201 0.8510 4.244 3

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR F 400 1000 833 1001 0.8322 4.276 3

Canadian Riv @ Eufaula, OK SMC 1995 Nov NR F 444 1350 678 599 1.1319 15.897 3

Chambers L nr Croscama, TX SMC 1996 Sept NR F NA NA 298 272 1.0956 ND 0

Chambers L nr Croscama, TX SMC 1996 Sept NR M 388 336 209 439 0.4761 ND 1

Chambers L nr Croscama, TX SMC 1996 Sept NR M 245 267 226 433 0.5219 ND 2

Chambers L nr Croscama, TX SMC 1996 Sept NR M NA NA 211 274 0.7701 ND 3

Chambers L nr Croscama, TX SMC 1996 Sept NR M NA NA 324 813 0.3985 ND 3
Table 5. (Continued) All individual LARGEMOUTH BASS values for biomarker and ancillary data from collections made at sites in U.S. 

waters from March 1994 through August 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Chambers L nr Croscama, TX SMC 1996 Sept NR M NA NA 214 299 0.7157 ND 1

Chickasaw NWR, TN SMC 1996 Jan NR M 405 964 247 218 1.1330 ND 3

Chickasaw NWR, TN SMC 1996 Jan NR M 361 624 314 186 1.6882 ND 1

Chickasaw NWR, TN SMC 1996 Jan NR M 351 595 277 135 2.0519 ND 3

Chickasaw NWR, TN SMC 1996 Jan NR M 341 567 196 88 2.2273 ND 3

Chickasaw NWR, TN SMC 1996 Jan NR F 282 454 173 101 1.7129 0.005 .

Chickasaw NWR, TN SMC 1996 Jan NR F 395 851 535 70 7.6429 0.023 1

Chickasaw NWR, TN SMC 1996 Jan NR F 573 2920 322 219 1.4703 0.034 3

Chickasaw NWR, TN SMC 1996 Jan NR F 465 1616 90 79 1.1392 0.141 1

Chickasaw NWR, TN SMC 1996 Jan NR F 365 680 214 104 2.0577 0.141 1

Chickasaw NWR, TN SMC 1996 Jan NR F 440 1304 132 115 1.1478 0.213 3

Chickasaw NWR, TN SMC 1996 Jan NR F 354 737 52 174 0.2989 0.412 .

Columbia Riv @ Pasco, WA NW 1997 Oct NR F 323 556 468 141 3.3191 ND 1

Columbia Riv @ Pasco, WA NW 1997 Oct NR F 364 744 1465 387 3.7855 0.006 1

Columbia Riv @ Pasco, WA NW 1997 Oct NR F 269 299 881 908 0.9703 0.155 1

Columbia Riv @ Pasco, WA NW 1997 Oct NR F 320 452 833 581 1.4337 0.172 1

Columbia Riv @ Pasco, WA NW 1997 Oct NR F 283 337 239 173 1.3815 0.172 1

Columbia Riv @ Pasco, WA NW 1997 Oct NR F 522 2600 351 208 1.6875 0.61 1

Columbia Riv @ Pasco, WA NW 1997 Oct NR F 407 1450 553 122 4.5328 33.665 3

Columbia Riv @ Pasco, WA NW 1997 Oct NR M 440 1248 298 654 0.4557 ND 2

Columbia Riv @ Pasco, WA NW 1997 Oct NR M 261 282 182 331 0.5498 ND 2

Columbia Riv @ Pasco, WA NW 1997 Oct NR M 245 250 131 504 0.2599 ND 2

Columbia Riv @ Pasco, WA NW 1997 Oct NR M 240 250 306 750 0.408 ND 2

Columbia Riv @ Pasco, WA NW 1997 Oct NR M 290 144 491 422 1.1635 ND 2

Columbia Riv @ Pasco, WA NW 1997 Oct NR M 407 1032 848 1307 0.6488 0.669 2

Connecticut R @ Thompsonville, CT NE 1994 Sept NR F 254 273 1372 195 7.0359 ND 1

Connecticut R @ Thompsonville, CT NE 1994 Sept NR F 257 280 510 111 4.5946 0.0036 1

Connecticut R @ Thompsonville, CT NE 1994 Sept NR F 373 784 307 93 3.3011 0.008 1

Connecticut R @ Thompsonville, CT NE 1994 Sept NR M 295 379 1500 1203 1.2469 ND 3

Connecticut R @ Thompsonville, CT NE 1994 Sept NR M 284 314 718 1206 0.5954 ND 3

Connecticut R @ Thompsonville, CT NE 1994 Sept NR M 284 370 231 361 0.6399 0.0014 3

Connecticut R @ Thompsonville, CT NE 1994 Sept NR M 393 494 386 257 1.5019 0.002 3

Cumberland Riv @ Clarksville, TN SMC 1995 Oct NR F 268 235 563 70 8.0429 ND 3

Cumberland Riv @ Clarksville, TN SMC 1995 Oct NR F 231 142 230 49 4.6939 ND 2

Cumberland Riv @ Clarksville, TN SMC 1995 Oct NR F 325 425 313 89 3.5169 0.156 3

Cumberland Riv @ Clarksville, TN SMC 1995 Oct NR F 320 379 341 52 6.5577 0.925 2

Cumberland Riv @ Clarksville, TN SMC 1995 Oct NR M 335 525 318 162 1.9630 ND 2

Cumberland Riv @ Clarksville, TN SMC 1995 Oct NR M 300 370 265 106 2.5 ND 2

Cumberland Riv @ Clarksville, TN SMC 1995 Oct NR M 256 250 594 121 4.9091 ND 3

Cumberland Riv @ Clarksville, TN SMC 1995 Oct NR M 230 167 214 198 1.0808 ND 2

Table 5 83
84 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 5. (Continued) All individual LARGEMOUTH BASS values for biomarker and ancillary data from collections made at sites in U.S. 

waters from March 1994 through August 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Cumberland Riv @ Clarksville, TN SMC 1995 Oct NR M 300 336 348 131 2.6565 0.102 2

Don Pedro Res nr La Grange, CA SW 1995 Oct NR F 526 2231 1198 265 4.5208 ND 1

Don Pedro Res nr La Grange, CA SW 1995 Oct NR F 429 1337 329 294 1.1190 ND 1

Don Pedro Res nr La Grange, CA SW 1995 Oct NR F 391 939 775 307 2.5244 ND 1

Don Pedro Res nr La Grange, CA SW 1995 Oct NR F 408 938 916 175 5.2343 ND 1

Don Pedro Res nr La Grange, CA SW 1995 Oct NR F 375 749 479 238 2.0126 ND 1

Don Pedro Res nr La Grange, CA SW 1995 Oct NR F 333 636 365 193 1.8912 ND 1

Don Pedro Res nr La Grange, CA SW 1995 Oct NR F 356 629 532 194 2.7423 ND 1

Don Pedro Res nr La Grange, CA SW 1995 Oct NR F 338 579 1142 232 4.9224 ND 1

Don Pedro Res nr La Grange, CA SW 1994 Oct NR F 276 245 720 72 10 ND 1

Don Pedro Res nr La Grange, CA SW 1994 Oct NR F 293 322 1183 304 3.8914 0.0014 0

Don Pedro Res nr La Grange, CA SW 1994 Oct NR F 546 2603 461 38 12.1316 0.0016 1

Don Pedro Res nr La Grange, CA SW 1994 Oct NR F 433 1104 523 123 4.2520 0.0016 1

Don Pedro Res nr La Grange, CA SW 1994 Oct NR F 309 406 1539 74 20.7973 0.0016 1

Don Pedro Res nr La Grange, CA SW 1994 Oct NR F 273 222 551 189 2.9153 0.0016 0

Don Pedro Res nr La Grange, CA SW 1994 Oct NR F 492 1710 1155 274 4.2153 0.002 1

Don Pedro Res nr La Grange, CA SW 1994 Oct NR F 580 2804 548 74 7.4054 0.003 1

Don Pedro Res nr La Grange, CA SW 1994 Oct NR F 354 600 1089 662 1.6450 0.021 1

Don Pedro Res nr La Grange, CA SW 1994 Oct NR F 440 1293 1219 409 2.9804 0.026 1

Don Pedro Res nr La Grange, CA SW 1995 Oct NR F 374 959 923 182 5.0714 0.085 1

Don Pedro Res nr La Grange, CA SW 1995 Oct NR F 387 901 289 159 1.8176 0.282 1

Don Pedro Res nr La Grange, CA SW 1995 Oct NR F 448 1455 954 145 6.5793 0.292 1

Don Pedro Res nr La Grange, CA SW 1995 Oct NR F 423 1265 1243 323 3.8483 0.326 1

Don Pedro Res nr La Grange, CA SW 1995 Oct NR F 362 644 318 161 1.9752 0.564 1

Don Pedro Res nr La Grange, CA SW 1995 Oct NR M 356 788 1028 1221 0.8419 ND 3

Don Pedro Res nr La Grange, CA SW 1995 Oct NR M 334 638 301 1501 0.2005 ND 3

Don Pedro Res nr La Grange, CA SW 1994 Oct NR M 279 272 777 227 3.4229 ND 3

Don Pedro Res nr La Grange, CA SW 1994 Oct NR M 275 260 1370 662 2.0695 ND 3

Don Pedro Res nr La Grange, CA SW 1994 Oct NR M 255 217 1119 1543 0.7252 ND 3

Don Pedro Res nr La Grange, CA SW 1994 Oct NR M 414 970 207 98 2.1122 0.001 3

Don Pedro Res nr La Grange, CA SW 1994 Oct NR M 448 1389 1300 1894 0.6864 0.0012 3

Don Pedro Res nr La Grange, CA SW 1994 Oct NR M 271 273 565 551 1.0254 0.0014 3

Don Pedro Res nr La Grange, CA SW 1994 Oct NR M 252 197 612 527 1.161 0.0018 3

Don Pedro Res nr La Grange, CA SW 1994 Oct NR M 326 420 446 188 2.372 0.002 3

Don Pedro Res nr La Grange, CA SW 1994 Oct NR M 268 235 958 1527 0.6274 1.8 3

Flathead Riv @ Creston, MT NW 1997 Oct NR F 315 441 574 297 1.9327 0.158 1

Flathead Riv @ Creston, MT NW 1997 Oct NR F 279 280 1143 495 2.3091 0.158 1

Flathead Riv @ Creston, MT NW 1997 Oct NR F 290 341 1092 292 3.7397 0.334 2

Flathead Riv @ Creston, MT NW 1997 Oct NR F 421 1281 1585 805 1.9689 0.398 1

Flathead Riv @ Creston, MT NW 1997 Oct NR F 426 1050 598 318 1.8805 0.837 2
Table 5. (Continued) All individual LARGEMOUTH BASS values for biomarker and ancillary data from collections made at sites in U.S. 

waters from March 1994 through August 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Flathead Riv @ Creston, MT NW 1997 Oct NR F 440 1577 1210 608 1.9901 1.047 3

Flathead Riv @ Creston, MT NW 1997 Oct NR F 470 1779 557 185 3.0108 1.117 2

Flathead Riv @ Creston, MT NW 1997 Oct NR F 475 2100 398 112 3.5536 1.469 2

Flathead Riv @ Creston, MT NW 1997 Oct NR F 461 1750 446 230 1.9391 1.516 2

Flathead Riv @ Creston, MT NW 1997 Oct NR F 496 2600 942 488 1.9303 2.355 2

Flathead Riv @ Creston, MT NW 1997 Oct NR M 430 1756 242 191 1.2670 ND 2

Flathead Riv @ Creston, MT NW 1997 Oct NR M 410 1171 113 456 0.2478 ND 2

Flathead Riv @ Creston, MT NW 1997 Oct NR M 330 619 326 976 0.3340 ND 2

Flathead Riv @ Creston, MT NW 1997 Oct NR M 330 540 371 607 0.6112 ND 3

Flathead Riv @ Creston, MT NW 1997 Oct NR M 315 486 146 623 0.2343 ND 3

Flathead Riv @ Creston, MT NW 1997 Oct NR M 306 399 182 695 0.2619 ND 3

Flathead Riv @ Creston, MT NW 1997 Oct NR M 280 299 127 121 1.0496 ND 3

Flathead Riv @ Creston, MT NW 1997 Oct NR M 279 291 251 849 0.2956 ND 3

Flathead Riv @ Creston, MT NW 1997 Oct NR M 252 114 161 501 0.3214 ND 3

Flathead Riv @ Creston, MT NW 1997 Oct NR M 461 1880 591 256 2.3086 0.083 1

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR F 393 1015 257 21 12.2381 0.097 1

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR F 484 1813 442 173 2.5549 0.139 1

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR M 384 893 11 66 0.1667 ND 3

Hudson Riv @ Poughkeepsie, NY NE 1994 Sept NR M 370 855 58 15 3.8667 ND 3

Illinois Riv @ Beardstown, IL NMC 1995 Sept NR F 400 1160 223 249 0.8956 ND 1

Illinois Riv @ Beardstown, IL NMC 1995 Sept NR F 302 492 239 683 0.3499 ND 2

Illinois Riv @ Beardstown, IL NMC 1995 Sept NR F 280 345 168 138 1.2174 ND 2

Illinois Riv @ Beardstown, IL NMC 1995 Sept NR F 258 267 102 475 0.2147 ND 2

Illinois Riv @ Beardstown, IL NMC 1995 Sept NR F 251 228 205 128 1.6016 0.008 2

Illinois Riv @ Beardstown, IL NMC 1995 Sept NR F 408 1173 423 869 0.4868 0.015 3

Illinois Riv @ Beardstown, IL NMC 1995 Sept NR F 275 375 220 735 0.2993 0.048 2

Illinois Riv @ Beardstown, IL NMC 1995 Sept NR F 285 410 305 204 1.4951 0.057 2

Illinois Riv @ Beardstown, IL NMC 1995 Sept NR F 391 1026 248 906 0.2737 0.064 2

Illinois Riv @ Beardstown, IL NMC 1995 Sept NR F 343 708 231 343 0.6735 0.072 2

Illinois Riv @ Beardstown, IL NMC 1995 Sept NR M 389 1119 251 261 0.9617 ND 1

Illinois Riv @ Beardstown, IL NMC 1995 Sept NR M 404 1058 244 295 0.8271 ND 1

Illinois Riv @ Beardstown, IL NMC 1995 Sept NR M 340 643 243 715 0.3399 ND 2

Illinois Riv @ Beardstown, IL NMC 1995 Sept NR M 320 548 180 231 0.7792 ND 1

Illinois Riv @ Beardstown, IL NMC 1995 Sept NR M 313 516 173 1755 0.0986 ND 1

Illinois Riv @ Beardstown, IL NMC 1995 Sept NR M 287 398 229 1488 0.1539 ND 2

Illinois Riv @ Beardstown, IL NMC 1995 Sept NR M 274 342 153 391 0.3913 ND 1

Illinois Riv @ Beardstown, IL NMC 1995 Sept NR M 284 291 197 1127 0.1748 ND 1

Illinois Riv @ Beardstown, IL NMC 1995 Sept NR M 258 285 184 404 0.4554 ND 1

Illinois Riv @ Beardstown, IL NMC 1995 Sept NR M 234 192 159 828 0.1920 ND 1

Leetown Res @ Leetown, WV NE 1996 Oct NR F 325 418.2 304 84 3.619 ND 2

Table 5 85
86 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 5. (Continued) All individual LARGEMOUTH BASS values for biomarker and ancillary data from collections made at sites in U.S. 

waters from March 1994 through August 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Leetown Res @ Leetown, WV NE 1996 Oct NR F 322 403 1237 544 2.274 ND 2

Leetown Res @ Leetown, WV NE 1996 Oct NR F 310 387 801 466 1.719 ND 2

Leetown Res @ Leetown, WV NE 1996 Oct NR F 312 379 723 399 1.812 ND 2

Leetown Res @ Leetown, WV NE 1996 Oct NR F 315 372.8 542 378 1.434 ND 2

Leetown Res @ Leetown, WV NE 1996 Oct NR F 305 341.8 304 84 3.619 ND 2

Leetown Res @ Leetown, WV NE 1996 Oct NR F 295 305.5 1403 601 2.334 ND 2

Leetown Res @ Leetown, WV NE 1996 Oct NR F 284 296.7 304 84 3.619 ND 1

Leetown Res @ Leetown, WV NE 1996 Oct NR F 318 335.6 520 316 1.646 0.173 2

Leetown Res @ Leetown, WV NE 1996 Oct NR F 292 277.5 688 390 1.764 0.512 2

Leetown Res @ Leetown, WV NE 1996 Oct NR M 306 377.8 270 504 0.536 ND 2

Leetown Res @ Leetown, WV NE 1996 Oct NR M 318 363.8 295 849 0.347 ND 2

Leetown Res @ Leetown, WV NE 1996 Oct NR M 321 331.4 387 665 0.582 ND 2

Leetown Res @ Leetown, WV NE 1996 Oct NR M 305 306.4 335 837 0.4 ND 2

Leetown Res @ Leetown, WV NE 1996 Oct NR M 304 297 296 693 0.427 ND 2

Leetown Res @ Leetown, WV NE 1996 Oct NR M 290 277.8 359 618 0.581 ND 1

Leetown Res @ Leetown, WV NE 1996 Oct NR M 268 267.2 335 1087 0.308 ND 2

Leetown Res @ Leetown, WV NE 1996 Oct NR M 330 238.5 312 960 0.325 ND 2

Leetown Res @ Leetown, WV NE 1996 Oct NR M 263 213.8 201 426 0.472 ND 2

Leetown Res @ Leetown, WV NE 1996 Oct NR M 296 265.6 341 841 0.405 0.114 3

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR M 360 816 569 676 0.8417 0.026 1

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR M 382 971 236 697 0.3386 0.006 2

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR F 365 777 640 583 1.0978 0.116 2

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR F 370 968 788 376 2.0957 0.157 2

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR F 375 994 806 462 1.7446 0.609 2

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR F 405 1272 1116 604 1.8477 1.045 3

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR F 355 682 604 771 0.7834 1.048 2

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR F 355 846 802 704 1.1392 1.304 2

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR F 390 1021 659 507 1.2998 1.529 2

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR F 375 864 742 584 1.2705 3.964 NA

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR F 420 1223 711 304 2.3388 4.688 3

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR F 410 1072 633 681 0.9295 11.276 2

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR M 387 1030 554 1056 0.5246 ND 2

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR M 350 720 857 1335 0.6419 ND 2

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR M 350 706 638 1024 0.6230 ND 2

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR M 343 670 634 1272 0.4984 ND 2

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR M 332 635 400 942 0.4246 ND 2

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR M 340 620 461 1229 0.3751 ND 2

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR M 325 620 546 978 0.5583 ND 2

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR M 309 471 508 432 1.1759 ND 1

Mississippi Riv @ Dubuque, IA NMC 1995 Oct NR M 275 323 340 875 0.3886 ND 1
Table 5. (Continued) All individual LARGEMOUTH BASS values for biomarker and ancillary data from collections made at sites in U.S. 

waters from March 1994 through August 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR F 337 656 1266 782 1.6189 0.021 2

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR F 425 1435 357 199 1.7940 0.059 2

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR F 392 1016 1268 306 4.1438 0.09 2

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR F 375 900 1277 477 2.6771 0.192 2

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR F 376 910 380 320 1.1875 0.778 2

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR F 411 1174 202 179 1.1285 0.874 2

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR F 423 1223 1285 463 2.7754 1.025 2

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR F 390 1043 1371 314 4.3662 1.108 2

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR F 390 940 1289 739 1.7442 1.471 2

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR F 456 1726 1279 277 4.6173 10.331 2

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR M 382 1008 199 536 0.3713 ND 2

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR M 393 931 266 753 0.3533 ND 2

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR M 387 918 132 736 0.1793 ND 2

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR M 375 867 291 386 0.7539 ND 2

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR M 372 843 210 719 0.2921 ND 1

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR M 365 783 289 675 0.4281 ND 2

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR M 363 771 99 627 0.1579 ND 2

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR M 342 630 215 507 0.4241 ND 2

Mississippi Riv @ Guttenburg, IA NMC 1995 Oct NR M 310 449 266 901 0.2952 ND 2

Mississippi Riv @ Memphis, TN SMC 1995 Oct NR F 301 328 196 132 1.4848 ND 2

Mississippi Riv @ Memphis, TN SMC 1995 Oct NR F 295 318 367 422 0.8697 ND 2

Mississippi Riv @ Memphis, TN SMC 1995 Oct NR F 231 151 199 44 4.5227 ND 2

Mississippi Riv @ Memphis, TN SMC 1995 Oct NR F 447 1822 690 127 5.4331 0.061 2

Mississippi Riv @ Memphis, TN SMC 1995 Oct NR F 469 1555 269 119 2.2605 0.077 2

Mississippi Riv @ Memphis, TN SMC 1995 Oct NR F 372 720 651 130 5.0077 0.103 2

Mississippi Riv @ Memphis, TN SMC 1995 Oct NR F 320 447 249 305 0.8164 0.341 2

Mississippi Riv @ Memphis, TN SMC 1995 Oct NR M 373 802 204 271 0.7528 ND 2

Mississippi Riv @ Memphis, TN SMC 1995 Oct NR M 362 734 788 115 6.8522 ND 2

Mississippi Riv @ Memphis, TN SMC 1995 Oct NR M 361 667 238 389 0.6118 ND 2

Mississippi Riv @ Memphis, TN SMC 1995 Oct NR M 315 463 727 182 3.9945 ND 2

Missouri Riv @ Hermann, MO NMC 1995 Oct NR F 426 1225 436 289 1.5087 ND 2

Missouri Riv @ Hermann, MO NMC 1995 Oct NR F 300 375 789 447 1.7651 0.205 2

Missouri Riv @ Hermann, MO NMC 1995 Oct NR F 366 725 338 309 1.0939 0.352 2

Missouri Riv @ Hermann, MO NMC 1995 Oct NR F 307 500 493 622 0.7926 0.562 2

Missouri Riv @ Hermann, MO NMC 1995 Oct NR F 296 400 416 176 2.3636 1.21 2

Missouri Riv @ Hermann, MO NMC 1995 Oct NR M 435 1450 395 191 2.0681 ND 2

Missouri Riv @ Hermann, MO NMC 1995 Oct NR M 353 775 662 2132 0.3105 ND 2

Missouri Riv @ Hermann, MO NMC 1995 Oct NR M 320 575 612 463 1.3218 ND 2

Missouri Riv @ Hermann, MO NMC 1995 Oct NR M 287 400 923 1295 0.7127 ND 2

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR F 303 430 561 371 1.5121 ND 1

Table 5 87
88 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 5. (Continued) All individual LARGEMOUTH BASS values for biomarker and ancillary data from collections made at sites in U.S. 

waters from March 1994 through August 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR F 306 470 237 18 13.1667 0.064 1

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR F 305 479 172 54 3.1852 0.075 1

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR F 312 534 531 30 17.7 0.4 1

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR M 310 409 373 142 2.6268 0.001 3

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR M 286 378 154 59 2.6102 0.001 3

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR M 285 345 44 61 0.7213 0.001 3

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR M 288 388 256 39 6.5641 0.0012 3

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR M 305 420 190 59 3.2203 0.0016 3

Mohawk Riv @ Frankfort, NY NE 1994 Sept NR M 277 325 180 48 3.75 0.215 3

Ohio Riv @ Marietta, OH NE 1995 Oct NR F 332 564 364 151 2.4106 0.245 3

Ohio Riv @ Marietta, OH NE 1995 Oct NR F 328 640 279 87 3.2069 1.003 3

Ohio Riv @ Marietta, OH NE 1995 Oct NR M 410 1100 308 441 0.6984 ND 2

Ohio Riv @ Marietta, OH NE 1995 Oct NR M 379 789 409 158 2.5886 ND 2

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR F 377 721 2434 788 3.0888 ND 2

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR F 374 615 245 816 0.3002 ND 2

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR F 373 613 1384 731 1.8933 ND 2

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR F 345 520 391 856 0.4568 ND 2

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR F 344 499 1313 957 1.3720 ND 2

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR F 320 368 469 754 0.6220 ND 2

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR F 316 366 355 702 0.5057 ND 2

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR F 310 346 1310 1312 0.9985 ND 2

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR F 310 343 435 95 4.5789 ND 2

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR M 515 1918 320 2009 0.1593 ND 2

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR M 415 1120 302 1138 0.2654 ND 2

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR M 420 1090 269 594 0.4529 ND 2

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR M 360 657 391 641 0.6100 ND 2

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR M 354 611 261 2648 0.0986 ND 2

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR M 340 470 297 1077 0.2758 ND 1

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR M 340 424 297 1338 0.2220 ND 1

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR M 321 409 509 1397 0.3644 ND 2

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR M 325 405 396 653 0.6064 ND 2

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR M 315 378 596 1039 0.5736 ND 2

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR M 325 377 226 1185 0.1907 ND 2

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR M 305 321 385 1236 0.3115 ND 1

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR M 249 176 199 418 0.4761 ND 1

Ohio Riv @ Metropolis, IL SMC 1995 Oct NR M 350 555 365 1298 0.2812 0.005 2

Potomic Riv @ Anacostia Riv NE 1994 Nov NR F 293 354 235 232 1.0129 0.0016 1

Potomic Riv @ Anacostia Riv NE 1994 Nov NR F 298 365 172 97 1.7732 0.056 1

Potomic Riv @ Anacostia Riv NE 1994 Nov NR F 320 503 546 731 0.7469 0.117 1

Potomic Riv @ Anacostia Riv NE 1994 Nov NR F 400 1164 136 240 0.5667 0.18 1
Table 5. (Continued) All individual LARGEMOUTH BASS values for biomarker and ancillary data from collections made at sites in U.S. 

waters from March 1994 through August 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Potomic Riv @ Anacostia Riv NE 1994 Nov NR F 390 598 51 35 1.4571 0.88 1

Potomic Riv @ Anacostia Riv NE 1994 Nov NR M 386 780 205 247 0.8300 0.001 3

Potomic Riv @ Anacostia Riv NE 1994 Nov NR M 357 658 36 46 0.7826 0.001 3

Potomic Riv @ Anacostia Riv NE 1994 Nov NR M 360 651 77 55 1.4 0.001 3

Potomic Riv @ Anacostia Riv NE 1994 Nov NR M 338 619 151 87 1.7356 0.001 3

Potomic Riv @ Anacostia Riv NE 1994 Nov NR M 328 607 294 79 3.7215 0.0012 3

Potomic Riv @ Anacostia Riv NE 1994 Nov NR M 330 524 60 59 1.0169 0.0012 3

Potomic Riv @ Anacostia Riv NE 1994 Nov NR M 274 228 21 327 0.0642 0.0016 3

Ray Roberts L nr Gainesville, Tx SMC 1996 Sept NR F 371 311 386 206 1.8738 ND 0

Ray Roberts L nr Gainesville, Tx SMC 1996 Sept NR F 306 305 249 123 2.0244 ND 0

Ray Roberts L nr Gainesville, Tx SMC 1996 Sept NR F NA NA 219 314 0.6975 ND 1

Ray Roberts L nr Gainesville, Tx SMC 1996 Sept NR F NA NA 566 144 3.9306 ND 1

Ray Roberts L nr Gainesville, Tx SMC 1996 Sept NR F NA NA 154 113 1.3628 ND 0

Ray Roberts L nr Gainesville, Tx SMC 1996 Sept NR F NA NA 304 169 1.7988 ND 0

Ray Roberts L nr Gainesville, Tx SMC 1996 Sept NR M 262 285 243 291 0.8351 ND 2

Ray Roberts L nr Gainesville, Tx SMC 1996 Sept NR M NA NA 231 740 0.3122 ND 3

Ray Roberts L nr Gainesville, Tx SMC 1996 Sept NR M NA NA 245 622 0.3939 ND 3

Ray Roberts L nr Gainesville, Tx SMC 1996 Sept NR M NA NA 187 931 0.2009 ND 3

Real Foot NWR, TN SMC 1996 Jan NR M 470 1332 124 661 0.1876 ND 2

Real Foot NWR, TN SMC 1996 Jan NR M 370 765 261 208 1.2548 ND 2

Real Foot NWR, TN SMC 1996 Jan NR M 330 510 969 852 1.1373 ND 2

Real Foot NWR, TN SMC 1996 Jan NR M 484 368 412 0.8932 ND 2

Real Foot NWR, TN SMC 1996 Jan NR F 508 1928 193 138 1.3986 0.005 3

Real Foot NWR, TN SMC 1996 Jan NR F 415 1021 451 314 1.4363 0.0056 3

Real Foot NWR, TN SMC 1996 Jan NR F 377 680 406 94 4.3191 0.0056 1

Real Foot NWR, TN SMC 1996 Jan NR F 420 1049 529 243 2.1770 0.023 0

Real Foot NWR, TN SMC 1996 Jan NR F 385 936 219 243 0.9012 0.023 0

Real Foot NWR, TN SMC 1996 Jan NR F 285 340 323 112 2.8839 0.024 1

Real Foot NWR, TN SMC 1996 Jan NR F 543 2381 554 120 4.6167 0.045 3

Real Foot NWR, TN SMC 1996 Jan NR F 466 1446 189 129 1.4651 0.061 3

Real Foot NWR, TN SMC 1996 Jan NR F 325 539 279 125 2.232 0.215 1

Real Foot NWR, TN SMC 1996 Jan NR F 414 1162 779 58 13.4310 0.217 0

Real Foot NWR, TN SMC 1996 Jan NR F 495 1673 319 224 1.4241 0.415 0

Red Riv @ Alexandria, LA SMC 1995 Oct NR F 462 1493 989 260 3.8038 ND 1

Red Riv @ Alexandria, LA SMC 1995 Oct NR F 425 1380 998 500 1.996 ND 2

Red Riv @ Alexandria, LA SMC 1995 Oct NR F 439 1301 745 236 3.1568 ND 1

Red Riv @ Alexandria, LA SMC 1995 Oct NR F 414 1108 411 365 1.1260 ND 0

Red Riv @ Alexandria, LA SMC 1995 Oct NR F 402 995 459 260 1.7654 ND 1

Red Riv @ Alexandria, LA SMC 1995 Oct NR F 386 899 794 439 1.8087 ND 2

Red Riv @ Alexandria, LA SMC 1995 Oct NR F 373 720 722 582 1.2405 ND 2

Table 5 89
90 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 5. (Continued) All individual LARGEMOUTH BASS values for biomarker and ancillary data from collections made at sites in U.S. 

waters from March 1994 through August 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Red Riv @ Alexandria, LA SMC 1995 Oct NR F 350 615 408 210 1.9429 ND 1

Red Riv @ Alexandria, LA SMC 1995 Oct NR F 347 556 404 801 0.5044 ND 1

Red Riv @ Alexandria, LA SMC 1995 Oct NR F 340 520 589 263 2.2395 ND 2

Red Riv @ Alexandria, LA SMC 1995 Oct NR F 292 395 735 422 1.7417 ND 0

Red Riv @ Alexandria, LA SMC 1995 Oct NR F 310 345 526 669 0.7862 ND 1

Red Riv @ Alexandria, LA SMC 1995 Oct NR F 290 344 1591 556 2.8615 ND 1

Red Riv @ Alexandria, LA SMC 1995 Oct NR F 262 254 819 698 1.1734 ND 0

Red Riv @ Alexandria, LA SMC 1995 Oct NR F 244 174 405 518 0.7819 ND 2

Red Riv @ Alexandria, LA SMC 1995 Oct NR F 231 140 422 899 0.4694 ND 0

Red Riv @ Alexandria, LA SMC 1995 Oct NR M 372 733 445 689 0.6459 ND 1

Red Riv @ Alexandria, LA SMC 1995 Oct NR M 330 590 433 464 0.9332 ND 2

Red Riv @ Alexandria, LA SMC 1995 Oct NR M 320 416 254 435 0.5839 ND 2

Red Riv @ Alexandria, LA SMC 1995 Oct NR M 295 316 450 492 0.9146 ND 1

Red Riv @ Alexandria, LA SMC 1995 Oct NR M 240 164 391 340 1.15 ND 1

Red Riv @ Alexandria, LA SMC 1995 Oct NR M 241 158 198 502 0.3944 ND 1

Red Riv @ Alexandria, LA SMC 1995 Oct NR M 235 149 355 579 0.6131 ND 1

Red Riv @ L Texoma, TX SMC 1995 Nov NR M 470 1467 936 313 2.9904 ND 2

Red Riv @ L Texoma, TX SMC 1995 Nov NR M 405 1148 825 409 2.0171 ND 2

Red Riv @ L Texoma, TX SMC 1995 Nov NR M 420 1044 479 346 1.3844 ND 2

Red Riv @ L Texoma, TX SMC 1995 Nov NR M 400 1025 597 812 0.7352 ND 2

Red Riv @ L Texoma, TX SMC 1995 Nov NR M 419 959 562 209 2.6890 ND 2

Red Riv @ L Texoma, TX SMC 1995 Nov NR M 415 929 788 1235 0.6381 ND 2

Red Riv @ L Texoma, TX SMC 1995 Nov NR M 395 750 534 205 2.6049 ND 2

Red Riv @ L Texoma, TX SMC 1995 Nov NR M 375 658 496 273 1.8168 ND 2

Red Riv @ L Texoma, TX SMC 1995 Nov NR M 354 602 649 1569 0.4136 ND 3

Red Riv @ L Texoma, TX SMC 1995 Nov NR M 352 577 584 848 0.6887 ND 3

Red Riv @ L Texoma, TX SMC 1995 Nov NR M 320 426 896 2215 0.4045 ND 2

Red Riv @ L Texoma, TX SMC 1995 Nov NR M 318 388 654 441 1.4830 ND 2

Red Riv @ L Texoma, TX SMC 1995 Nov NR M 305 373 544 809 0.6724 ND 2

Red Riv @ L Texoma, TX SMC 1995 Nov NR F 497 1200 560 91 6.1538 ND 2

Red Riv @ L Texoma, TX SMC 1995 Nov NR F 330 460 466 497 0.9376 ND 2

Red Riv @ L Texoma, TX SMC 1995 Nov NR F 320 432 965 170 5.6765 ND 2

Red Riv @ L Texoma, TX SMC 1995 Nov NR F 309 366 842 648 1.2994 ND 2

Red Riv @ L Texoma, TX SMC 1995 Nov NR F 540 2100 721 182 3.9615 0.006 2

Red Riv @ L Texoma, TX SMC 1995 Nov NR F 340 494 391 123 3.1789 0.047 2

Red Riv @ L Texoma, TX SMC 1995 Nov NR F 400 1103 1061 168 6.3155 0.119 3

Red Riv @ L Texoma, TX SMC 1995 Nov NR F 425 1011 419 57 7.3509 0.165 2

Red Riv @ L Texoma, TX SMC 1995 Nov NR F 380 817 1031 64 16.1094 0.202 3

Red Riv @ L Texoma, TX SMC 1995 Nov NR F 370 769 858 71 12.0845 0.238 2

Red Riv @ L Texoma, TX SMC 1995 Nov NR F 410 1004 512 309 1.6570 0.622 2
Table 5. (Continued) All individual LARGEMOUTH BASS values for biomarker and ancillary data from collections made at sites in U.S. 

waters from March 1994 through August 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Red Riv @ L Texoma, TX SMC 1995 Nov NR F 472 1469 975 412 2.3665 0.977 3

Red Riv @ L Texoma, TX SMC 1995 Nov NR F 440 1265 569 102 5.5784 4.874 2

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR F 441 1459 1181 362 3.2624 ND 2

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR F 383 937 69 167 0.4132 ND 2

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR F 393 1000 129 12 10.75 0.024 1

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR F 405 1061 87 71 1.2254 0.045 1

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR F 285 320 207 79 2.6203 0.099 1

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR F 467 1515 135 35 3.8571 0.135 1

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR F 412 1200 100 35 2.8571 0.137 1

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR F 595 2009 152 86 1.7674 0.2 1

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR F 385 972 199 85 2.3412 0.207 2

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR F 300 474 224 285 0.7860 0.31 1

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR F 343 619 704 117 6.0171 0.321 1

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR F 306 548 54 21 2.5714 0.344 1

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR F 322 468 295 45 6.5556 0.355 1

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR F 481 2020 1083 101 10.7228 0.42 2

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR F 417 1260 112 17 6.5882 0.43 2

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR F 380 947 1076 62 17.3548 0.437 2

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR F 451 1667 514 52 9.8846 0.724 2

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR F 342 749 96 38 2.5263 0.9546 2

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR F 335 647 2229 254 8.7756 1.123 2

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR M 435 1333 168 562 0.2989 ND 3

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR M 410 1018 118 85 1.3882 ND 3

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR M 370 807 715 1131 0.6322 ND 3

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR M 374 796 106 245 0.4327 ND 3

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR M 380 769 74 18 4.1111 ND 3

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR M 360 678 80 46 1.7391 ND 3

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR M 336 665 284 93 3.0538 ND 3

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR M 344 650 19 42 0.4524 ND 3

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR M 335 641 66 52 1.2692 ND 3

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR M 322 623 88 39 2.2564 ND 3

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR M 335 594 72 117 0.6154 ND 3

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR M 323 570 640 152 4.2105 ND 3

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR M 304 485 131 65 2.0154 ND 3

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR M 302 307 66 140 0.4714 ND 3

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR M 375 830 32 47 0.6809 0.001 3

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR M 390 826 85 69 1.232 0.001 3

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR M 258 207 56 48 1.167 0.001 3

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR M 295 419 120 47 2.5532 0.005 3

San Joaquin Riv @ Salt Slough, CA SW 1994 Oct NR M 383 1021 27 31 0.935 0.016 3

Table 5 91
92 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 5. (Continued) All individual LARGEMOUTH BASS values for biomarker and ancillary data from collections made at sites in U.S. 

waters from March 1994 through August 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR M 402 1063 821 1131 0.7259 0.036 2

San Joaquin Riv @ Salt Slough, CA SW 1995 Oct NR M 328 596 151 91 1.6593 0.039 3

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 442 1401 377 165 2.2848 ND 1

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 410 1048 29 109 0.2661 ND 1

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 384 947 418 619 0.6753 ND 2

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 357 627 187 148 1.2635 ND 1

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 324 522 1345 241 5.5809 ND 1

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 320 457 281 132 2.1288 0.002 1

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 376 800 284 111 2.5586 0.004 1

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 290 326 321 190 1.6895 0.004 1

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 270 277 629 202 3.1139 0.018 1

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 455 1609 156 59 2.6441 0.029 1

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 320 419 116 127 0.9134 0.039 1

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 350 678 200 114 1.7544 0.046 1

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 270 809 170 137 1.2409 0.049 1

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 482 2010 93 117 0.7949 0.05 1

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR F 349 643 175 204 0.8578 0.052 1

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 311 463 95 159 0.5975 0.209 2

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 422 1043 405 527 0.7685 0.21 2

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 361 605 1243 133 9.3459 0.21 2

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 465 1583 344 310 1.1097 0.217 1

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 478 1986 206 516 0.3992 0.223 1

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 390 864 171 519 0.3295 0.244 2

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 375 875 421 699 0.6023 0.331 2

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 387 817 477 811 0.5882 0.379 2

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 355 733 236 89 2.6517 0.455 2

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 350 628 747 269 2.7770 0.566 2

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 426 1125 310 85 3.6471 0.597 1

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 373 851 259 353 0.7337 0.853 1

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 356 773 208 429 0.4848 0.956 2

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 448 1292 326 515 0.6330 1.259 2

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 346 674 1245 267 4.6629 1.288 2

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR F 428 1395 114 110 1.0364 3.541 2

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR M 378 870 141 144 0.9792 ND 3

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR M 356 761 243 403 0.6030 ND 3

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR M 354 732 131 181 0.7238 ND 3

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR M 354 678 761 190 4.0053 ND 3

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR M 375 654 77 323 0.2384 ND 3

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR M 357 627 439 618 0.7104 ND 3

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR M 328 553 77 128 0.6016 ND 3
Table 5. (Continued) All individual LARGEMOUTH BASS values for biomarker and ancillary data from collections made at sites in U.S. 

waters from March 1994 through August 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR M 354 549 50 279 0.1792 ND 3

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR M 311 467 339 197 1.7208 ND 3

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR M 291 360 209 656 0.3186 ND 3

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR M 433 1263 109 97 1.1237 0.001 3

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR M 475 914 85 176 0.4830 0.001 3

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR M 406 993 154 573 0.2688 0.0016 3

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR M 355 692 148 152 0.9737 0.0016 3

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR M 356 646 35 181 0.1934 0.0018 3

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR M 355 789 257 103 2.4951 0.002 3

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR M 273 267 116 160 0.725 0.002 3

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR M 330 530 387 231 1.6753 0.024 3

San Joaquin Riv @ Vernalis, CA SW 1995 Oct NR M 416 958 49 208 0.2356 0.025 3

San Joaquin Riv @ Vernalis, CA SW 1994 Oct NR M 226 230 141 207 0.6812 0.117 3

Snake Riv @ Hagerman, ID NW 1997 Oct NR F 430 1150 743 179 4.1508 0.026 1

Snake Riv @ Hagerman, ID NW 1997 Oct NR F 492 2000 412 167 2.4671 0.031 1

Snake Riv @ Hagerman, ID NW 1997 Oct NR F 318 484 1134 481 2.3576 0.031 1

Snake Riv @ Hagerman, ID NW 1997 Oct NR F 334 714 591 454 1.3018 0.058 1

Snake Riv @ Hagerman, ID NW 1997 Oct NR F 367 914 413 377 1.0955 0.059 2

Snake Riv @ Hagerman, ID NW 1997 Oct NR F 394 1125 797 208 3.8317 0.069 1

Snake Riv @ Hagerman, ID NW 1997 Oct NR F 330 531 242 191 1.2670 0.071 1

Snake Riv @ Hagerman, ID NW 1997 Oct NR F 257 253 871 342 2.5468 0.408 1

Snake Riv @ Hagerman, ID NW 1997 Oct NR F 359 850 576 188 3.0638 1.535 2

Snake Riv @ Hagerman, ID NW 1997 Oct NR M 397 970 215 867 0.2480 ND 3

Snake Riv @ Hagerman, ID NW 1997 Oct NR M 385 900 815 414 1.9686 ND 1

Snake Riv @ Hagerman, ID NW 1997 Oct NR M 341 781 159 349 0.4556 ND 3

Snake Riv @ Hagerman, ID NW 1997 Oct NR M 335 606 177 424 0.4175 ND 2

Snake Riv @ Hagerman, ID NW 1997 Oct NR M 324 510 131 565 0.2319 ND 2

Snake Riv @ Hagerman, ID NW 1997 Oct NR M 314 500 310 7451 0.0416 ND 2

Snake Riv @ Hagerman, ID NW 1997 Oct NR M 315 444 582 711 0.8186 ND 3

Swinging Brg Res nr Monticello, NY NE 1994 Oct NR F 448 NA 446 244 1.8279 0.006 2

Swinging Brg Res nr Monticello, NY NE 1994 Oct NR F 391 956 211 129 1.6357 0.09 1

Swinging Brg Res nr Monticello, NY NE 1994 Oct NR F 383 930 503 75 6.7067 0.136 2

Swinging Brg Res nr Monticello, NY NE 1994 Oct NR F 359 NA 701 46 15.2391 0.196 2

Swinging Brg Res nr Monticello, NY NE 1994 Oct NR F 442 NA 380 75 5.0667 0.2 2

Swinging Brg Res nr Monticello, NY NE 1994 Oct NR F 495 NA 124 92 1.3478 0.24 2

Swinging Brg Res nr Monticello, NY NE 1994 Oct NR F 370 785 741 148 5.007 0.4 1

Swinging Brg Res nr Monticello, NY NE 1994 Oct NR M 374 850 63 76 0.8289 ND 3

Swinging Brg Res nr Monticello, NY NE 1994 Oct NR M 345 581 20 63 0.3175 ND 3

Swinging Brg Res nr Monticello, NY NE 1994 Oct NR M 350 NA 14 149 0.0940 ND 3

Swinging Brg Res nr Monticello, NY NE 1994 Oct NR M 343 NA 52 212 0.2453 ND 3

Table 5 93
94 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 5. (Continued) All individual LARGEMOUTH BASS values for biomarker and ancillary data from collections made at sites in U.S. 

waters from March 1994 through August 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Swinging Brg Res nr Monticello, NY NE 1994 Oct NR M 327 NA 30 210 0.1429 ND 3

Swinging Brg Res nr Monticello, NY NE 1994 Oct NR M 322 NA 191 446 0.4283 ND 3

Swinging Brg Res nr Monticello, NY NE 1994 Oct NR M 252 NA 15 78 0.1923 ND 3

Swinging Brg Res nr Monticello, NY NE 1994 Oct NR M 350 NA 55 98 0.5612 0.0014 3

Swinging Brg Res nr Monticello, NY NE 1994 Oct NR M 420 NA 33 82 0.4024 0.0016 3

Swinging Brg Res nr Monticello, NY NE 1994 Oct NR M 305 NA 29 116 0.25 0.0016 3

Swinging Brg Res nr Monticello, NY NE 1994 Oct NR M 422 NA 13 92 0.1413 0.0018 3

Swinging Brg Res nr Monticello, NY NE 1994 Oct NR M 254 NA 200 194 1.0309 3 3

Swinging Brg Res nr Monticello, NY NE 1995 May R F 424 1127 776 228 3.4035 ND 3

Swinging Brg Res nr Monticello, NY NE 1995 May R F 417 1139 1734 230 7.5391 0.631 3

Swinging Brg Res nr Monticello, NY NE 1995 May R F 325 478 3229 261 12.3716 0.776 3

Swinging Brg Res nr Monticello, NY NE 1995 May R F 362 643 384 108 3.5556 0.907 3

Swinging Brg Res nr Monticello, NY NE 1995 May R F 390 800 9670 2087 4.6334 5.023 3

Swinging Brg Res nr Monticello, NY NE 1995 May R F 468 1637 2694 1437 1.8747 5.05 3

Swinging Brg Res nr Monticello, NY NE 1995 May R F 342 664 1682 372 4.5215 5.176 4

Swinging Brg Res nr Monticello, NY NE 1995 May R F 441 1269 8074 2306 3.5013 5.506 3

Swinging Brg Res nr Monticello, NY NE 1995 May R F 348 584 1789 319 5.6082 5.9 3

Swinging Brg Res nr Monticello, NY NE 1995 May R F 404 1025 5143 1185 4.3401 6.864 3

Swinging Brg Res nr Monticello, NY NE 1995 May R F 362 688 3575 720 4.9653 7.206 3

Swinging Brg Res nr Monticello, NY NE 1995 May R F 422 1218 3281 905 3.6254 7.358 3

Swinging Brg Res nr Monticello, NY NE 1995 May R F 478 1648 1215 211 5.7583 8.474 3

Swinging Brg Res nr Monticello, NY NE 1995 May R F 400 1098 1846 188 9.8191 12.441 5

Swinging Brg Res nr Monticello, NY NE 1995 May R M 386 938 489 1521 0.3215 ND 3

Swinging Brg Res nr Monticello, NY NE 1995 May R M 331 553 562 1429 0.3933 ND 3

Tennessee Riv @ Savannah, TN SMC 1995 Oct NR F 260 300 2261 501 4.5130 ND 2

Tennessee Riv @ Savannah, TN SMC 1995 Oct NR F 355 534 1309 869 1.5063 0.098 2

Tennessee Riv @ Savannah, TN SMC 1995 Oct NR F 260 232 1479 251 5.8924 0.863 2

Tennessee Riv @ Savannah, TN SMC 1995 Oct NR M 415 900 354 1390 0.2547 ND 2

Tennessee Riv @ Savannah, TN SMC 1995 Oct NR M 300 400 227 871 0.2606 ND 2

Tennessee Riv @ Savannah, TN SMC 1995 Oct NR M 272 281 426 921 0.4625 ND 2

Tennessee Riv @ Savannah, TN SMC 1995 Oct NR M 271 281 314 614 0.5114 ND 1

Tennessee Riv @ Savannah, TN SMC 1995 Oct NR M 265 231 341 2574 0.1325 ND 2

Tennessee Riv @ Savannah, TN SMC 1995 Oct NR M 242 216 395 1327 0.2977 ND 2

Tennessee Riv @ Savannah, TN SMC 1995 Oct NR M 233 184 196 773 0.2536 ND 2

Tennessee Riv @ Savannah, TN SMC 1995 Oct NR M 252 170 272 1306 0.2083 ND 2

Tennessee Riv @ Savannah, TN SMC 1995 Oct NR M 335 540 299 1385 0.2159 0.008 2

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR M 457 1500 653 1470 0.4442 ND 3

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR M 407 1215 331 1499 0.2208 ND 2

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR M 403 1000 329 1140 0.2886 ND 2

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR M 370 950 578 1664 0.3474 ND 3
Table 5. (Continued) All individual LARGEMOUTH BASS values for biomarker and ancillary data from collections made at sites in U.S. 

waters from March 1994 through August 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR M 367 825 947 1712 0.5532 ND 2

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR M 355 650 739 1149 0.6432 ND 2

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR M 287 350 722 1044 0.6916 ND 2

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR M 272 300 1823 1951 0.9344 ND 3

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR F 280 350 1015 2203 0.4607 ND 2

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR F 285 400 1728 671 2.5753 0.012 2

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR F 308 450 1651 907 1.8203 0.05 2

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR F 505 2150 2319 1056 2.1960 0.09 3

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR F 455 1800 2356 334 7.0539 0.136 3

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR F 403 1150 1370 271 5.0554 0.194 3

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR F 510 2400 403 233 1.7296 0.513 3

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR F 395 1000 1295 151 8.5762 1.129 3

Verdigris Riv @ Oologah, OK SMC 1995 Dec NR F 394 1350 1286 251 5.1235 3.169 3

Wabash Riv @ N Harmony, IN NMC 1995 Nov NR F 310 450 369 132 2.7955 ND 0

Wabash Riv @ N Harmony, IN NMC 1995 Nov NR F 295 335 431 127 3.3937 ND 2

Wabash Riv @ N Harmony, IN NMC 1995 Nov NR F 266 176 496 97 5.1134 ND 2

Wabash Riv @ N Harmony, IN NMC 1995 Nov NR F 316 398 1426 337 4.2315 5.092 3

Wabash Riv @ N Harmony, IN NMC 1995 Nov NR M 314 500 315 1372 0.2296 ND 2

Wabash Riv @ N Harmony, IN NMC 1995 Nov NR M 314 435 389 183 2.1257 ND 2

Wabash Riv @ N Harmony, IN NMC 1995 Nov NR M 318 424 1355 218 6.2156 ND 3

Wabash Riv @ N Harmony, IN NMC 1995 Nov NR M 265 258 393 129 3.0465 ND 3

Wabash Riv @ N Harmony, IN NMC 1995 Nov NR M 293 306 378 1681 0.2249 0.008 2

White Riv @ Devall’s Bluff, AR SMC 1995 Sept NR F 379 816 463 1838 0.2519 ND 1

White Riv @ Devall’s Bluff, AR SMC 1995 Sept NR F 365 791 1322 96 13.7708 ND 1

White Riv @ Devall’s Bluff, AR SMC 1995 Sept NR F 294 313 496 125 3.968 ND 1

White Riv @ Devall’s Bluff, AR SMC 1995 Sept NR F 321 427 807 68 11.8676 0.032 2

White Riv @ Devall’s Bluff, AR SMC 1995 Sept NR F 348 702 256 86 2.9767 0.162 2

White Riv @ Devall’s Bluff, AR SMC 1995 Sept NR F 312 399 927 71 13.0563 0.311 2

White Riv @ Devall’s Bluff, AR SMC 1995 Sept NR F 302 358 5221 721 7.2413 0.911 1

White Riv @ Devall’s Bluff, AR SMC 1995 Sept NR F NA 1255 1169 169 6.9172 2.604 2

White Riv @ Devall’s Bluff, AR SMC 1995 Sept NR F 360 715 1361 55 24.7455 3.177 2

White Riv @ Devall’s Bluff, AR SMC 1995 Sept NR F 320 407 2027 322 6.2950 6.942 2

White Riv @ Devall’s Bluff, AR SMC 1995 Sept NR M 370 655 583 649 0.8983 ND 2

White Riv @ Devall’s Bluff, AR SMC 1995 Sept NR M 332 538 481 1544 0.3115 ND 1

White Riv @ Devall’s Bluff, AR SMC 1995 Sept NR M 320 464 277 295 0.9390 ND 2

White Riv @ Devall’s Bluff, AR SMC 1995 Sept NR M 310 460 398 1189 0.3347 ND 2

White Riv @ Devall’s Bluff, AR SMC 1995 Sept NR M 320 441 279 618 0.4515 ND 2

White Riv @ Devall’s Bluff, AR SMC 1995 Sept NR M 300 345 545 958 0.5689 ND 2

White Riv @ Devall’s Bluff, AR SMC 1995 Sept NR M 275 269 644 1279 0.5035 ND 2

White Riv @ Devall’s Bluff, AR SMC 1995 Sept NR M 285 324 639 775 0.8245 0.007 2

Table 5 95
96 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

Table 5. (Continued) All individual LARGEMOUTH BASS values for biomarker and ancillary data from collections made at sites in U.S. 

waters from March 1994 through August 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

White Riv @ Devall’s Bluff, AR SMC 1995 Sept NR M 302 359 1150 181 6.3536 2.817 2

White Rock L, Dallas, TX SMC 1996 Sept NR F NA NA 400 327 1.2232 ND 1

White Rock L, Dallas, TX SMC 1996 Sept NR F NA NA 402 289 1.3910 ND 1

White Rock L, Dallas, TX SMC 1996 Sept NR F NA NA 390 311 1.2540 ND 1

White Rock L, Dallas, TX SMC 1996 Sept NR F NA NA 292 225 1.2978 ND 0

White Rock L, Dallas, TX SMC 1996 Sept NR F NA NA 248 267 0.9288 ND 1

White Rock L, Dallas, TX SMC 1996 Sept NR F NA NA 284 284 1 ND 1

White Rock L, Dallas, TX SMC 1996 Sept NR M 387 295 184 232 0.7931 ND 2

White Rock L, Dallas, TX SMC 1996 Sept NR M NA NA 192 298 0.6443 ND 3

Willamette Riv @ Portland, OR NW 1997 Nov NR F 280 306 576 101 5.7030 ND 1

Willamette Riv @ Portland, OR NW 1997 Nov NR F 345 739 743 191 3.8901 0.045 1

Willamette Riv @ Portland, OR NW 1997 Nov NR F 321 472 413 326 1.2669 0.052 1

Willamette Riv @ Portland, OR NW 1997 Nov NR F 355 758 412 182 2.2637 0.116 1

Willamette Riv @ Portland, OR NW 1997 Nov NR F 338 582 797 532 1.4981 0.138 1

Willamette Riv @ Portland, OR NW 1997 Nov NR M 336 590 1031 260 3.9654 ND 2

Willamette Riv @ Portland, OR NW 1997 Nov NR M 268 306 582 1473 0.3951 ND 3

Willamette Riv @ Portland, OR NW 1997 Nov NR M 255 224 310 705 0.4397 ND 3

Wolf River nr LaGrange, TN SMC 1995 Oct NR F 250 191 121 1505 0.0804 ND 2

Wolf River nr LaGrange, TN SMC 1995 Oct NR F 320 379 5501 1512 3.6382 54.137 3

Wolf River nr LaGrange, TN SMC 1995 Oct NR F 275 274 6330 228 27.7632 74.905 2

Wolf River nr LaGrange, TN SMC 1995 Oct NR F 400 859 5181 956 5.4195 77.918 3

Wolf River nr LaGrange, TN SMC 1995 Oct NR M 425 1075 143 1116 0.1281 ND 2

Wolf River nr LaGrange, TN SMC 1995 Oct NR M 340 462 483 335 1.4418 ND 2

Wolf River nr LaGrange, TN SMC 1995 Oct NR M 300 329 355 1549 0.2292 ND 2

Wolf River nr LaGrange, TN SMC 1995 Oct NR M 260 198 235 853 0.2755 ND 2

Wolf River nr LaGrange, TN SMC 1995 Oct NR M 305 276 958 333 2.8769 0.002 2

Wolf River nr LaGrange, TN SMC 1995 Oct NR M 340 449 555 1409 0.3939 0.003 2

Wolf River nr LaGrange, TN SMC 1995 Oct NR M 270 211 173 333 0.5195 0.003 2

Woods Pond @ Lenox, MA NE 1994 Sept NR F 264 288 171 178 0.9607 0.0008 1

Woods Pond @ Lenox, MA NE 1994 Sept NR F 295 375 228 400 0.57 0.0012 1

Woods Pond @ Lenox, MA NE 1994 Sept NR F 375 886 57 162 0.3519 0.0014 1

Woods Pond @ Lenox, MA NE 1994 Sept NR F 290 382 94 183 0.5137 0.0014 1

Woods Pond @ Lenox, MA NE 1994 Sept NR F 301 409 401 705 0.5688 0.0016 1

Woods Pond @ Lenox, MA NE 1994 Sept NR F 282 342 570 536 1.0634 0.0016 1

Woods Pond @ Lenox, MA NE 1994 Sept NR F 351 718 33 729 0.0453 0.002 1

Woods Pond @ Lenox, MA NE 1994 Sept NR F 302 419 692 272 2.5441 0.002 1

Woods Pond @ Lenox, MA NE 1994 Sept NR F 293 341 958 456 2.1009 0.003 1

Woods Pond @ Lenox, MA NE 1994 Sept NR F 311 460 405 201 2.0149 0.0032 1

Woods Pond @ Lenox, MA NE 1994 Sept NR F 320 489 720 564 1.2766 0.004 1

Woods Pond @ Lenox, MA NE 1994 Sept NR F 335 635 309 123 2.5122 0.008 1
Table 5. (Continued) All individual LARGEMOUTH BASS values for biomarker and ancillary data from collections made at sites in U.S. 

waters from March 1994 through August 1997.

[Region: NE, northeast; NW, northwest; SE, southeast; SW, southwest; NMC, northern mid-continent; SMC, southern mid-continent. 

Season: NR, non-reproductive. R, reproductive. Sex: F, female, M, male. E2, 17 ß Estradiol; 11KT, 11 ketotestosterone; VTG, vitellogenin; 

mm, millimeter; g, grams; pg, picograms; mL, milliliter; mg, milligram; ND, not detected; NA, not available]

Site name 

(fig. 1) Region Year Month Season Sex Length(mm)

Weight

(g)

E2

(pg/mL)

11KT

(pg/mL)

Ratio of 

E2 to KT

VTG

(mg/mL)

Developmental 

stage

Woods Pond @ Lenox, MA NE 1994 Sept NR M 308 440 1054 412 2.5583 0.001 3

Woods Pond @ Lenox, MA NE 1994 Sept NR M 251 248 795 190 4.184 0.0012 3

Woods Pond @ Lenox, MA NE 1994 Sept NR M 335 542 107 217 0.4931 0.0014 3

Woods Pond @ Lenox, MA NE 1994 Sept NR M 294 351 347 373 0.9303 0.0016 3

Woods Pond @ Lenox, MA NE 1994 Sept NR M 292 358 503 176 2.8580 0.0017 3

Woods Pond @ Lenox, MA NE 1994 Sept NR M 318 519 275 509 0.5403 0.002 0

Woods Pond @ Lenox, MA NE 1994 Sept NR M 272 290 601 807 0.745 0.0036 2

Woods Pond @ Lenox, MA NE 1995 May R F 286 354 732 402 1.8209 ND 3

Woods Pond @ Lenox, MA NE 1995 May R F 293 350 965 554 1.7419 0.718 2

Woods Pond @ Lenox, MA NE 1995 May R F 318 479 1243 784 1.5855 5.972 4

Woods Pond @ Lenox, MA NE 1995 May R F 315 497 739 382 1.9346 8.242 3

Woods Pond @ Lenox, MA NE 1995 May R M 328 541 763 1248 0.6114 ND 3

Woods Pond @ Lenox, MA NE 1995 May R M 307 510 449 583 0.7702 ND 3

Woods Pond @ Lenox, MA NE 1995 May R M 315 453 560 684 0.8187 ND 3

Woods Pond @ Lenox, MA NE 1995 May R M 293 437 1209 1545 0.7825 ND 3

Woods Pond @ Lenox, MA NE 1995 May R M 290 408 379 1706 0.2222 ND 3

Woods Pond @ Lenox, MA NE 1995 May R M 308 405 280 1117 0.2507 ND 3

Woods Pond @ Lenox, MA NE 1995 May R M 305 389 1621 845 1.9183 ND 3

Woods Pond @ Lenox, MA NE 1995 May R M 280 345 1246 410 3.0390 ND 3

Woods Pond @ Lenox, MA NE 1995 May R M 276 335 531 1679 0.3163 ND 3

Woods Pond @ Lenox, MA NE 1995 May R M 282 331 1372 1114 1.2316 ND 3

Woods Pond @ Lenox, MA NE 1995 May R M 283 338 1184 1690 0.7006 0.043 3

Woods Pond @ Lenox, MA NE 1995 May R M 289 321 286 1165 0.2455 0.245 3

Yakima Riv @ Granger, WA NW 1997 Oct NR F 263 272 910 212 4.2925 0.299 1

Yakima Riv @ Granger, WA NW 1997 Oct NR F 403 1065 775 193 4.0155 1.255 1

Yakima Riv @ Granger, WA NW 1997 Oct NR F 334 634 1406 689 2.0406 1.767 2

Yakima Riv @ Granger, WA NW 1997 Oct NR F 450 1803 1399 521 2.6852 2.631 2

Yakima Riv @ Granger, WA NW 1997 Oct NR M 418 1138 106 199 0.5327 ND 3

Yakima Riv @ Granger, WA NW 1997 Oct NR M 356 691 179 435 0.4115 ND 2

Table 5 97
98 Reproductive and Endocrine Biomarkers in Largemouth Bass and Common Carp from United States Waters 

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