A Checklist of the Aquatic Invertebrates of 

the Delaware River Basin, 1990-2000

By Michael D. Bilger, Karen Riva-Murray, and Gretchen L. Wall

U.S. Department of the Interior 

U.S. Geological Survey

Data Series 116
U.S. Department of the Interior

Gale A. Norton, Secretary

U.S. Geological Survey

Charles G. Groat, Director

U.S. Geological Survey, Reston, Virginia: 2005

For sale by U.S. Geological Survey, Information Services 

Box 25286, Denver Federal Center 

Denver, CO 80225

For more information about the USGS and its products: 

Telephone: 1-888-ASK-USGS 

World Wide Web: http://www.usgs.gov/

Any use of trade, product, or firm names in this publication is for descriptive purposes only and does not imply 

endorsement by the U.S. Government.

Although this report is in the public domain, permission must be secured from the individual copyright owners to reproduce any copyrighted materials contained within this report.

Suggested citation:

Bilger, M.D., Riva-Murray, Karen, and Wall, G.L., 2005, A checklist of the aquatic invertebrates of the Delaware River 

Basin, 1990-2000: U.S. Geological Survey Data Series 116, 29 p. 
 iii

FOREWORD

The U.S. Geological Survey (USGS) is committed to providing the Nation with accurate and timely scientific information that helps enhance and protect the overall quality of life and that facilitates effective management of water, biological, energy, and mineral resources (http://www.usgs.gov/). Information on the quality of the Nation’s water resources is critical to assuring the long-term availability of 

water that is safe for drinking and recreation and suitable for industry, irrigation, and habitat for fish 

and wildlife. Population growth and increasing demands for multiple water uses make water availability, now measured in terms of quantity and quality, even more essential to the long-term sustainability 

of our communities and ecosystems.

The USGS implemented the National Water-Quality Assessment (NAWQA) Program in 1991 to support 

national, regional, and local information needs and decisions related to water-quality management 

and policy (http://water.usgs.gov/nawqa). Shaped by and coordinated with ongoing efforts of other 

Federal, State, and local agencies, the NAWQA Program is designed to answer: What is the condition 

of our Nation’s streams and ground water? How are the conditions changing over time? How do natural 

features and human activities affect the quality of streams and ground water, and where are those 

effects most pronounced? By combining information on water chemistry, physical characteristics, 

stream habitat, and aquatic life, the NAWQA Program aims to provide science-based insights for current and emerging water issues and priorities. 

From 1991-2001, the NAWQA Program completed interdisciplinary assessments in 51 of the Nation’s 

major river basins and aquifer systems, referred to as Study Units (http://water.usgs.gov/nawqa/

studyu.html). Baseline conditions were established for comparison to future assessments, and longterm monitoring was initiated in many of the basins. During the next decade, 42 of the 51 Study Units 

will be reassessed so that 10 years of comparable monitoring data will be available to determine 

trends at many of the Nation’s streams and aquifers. The next 10 years of study also will fill in critical 

gaps in characterizing water-quality conditions, enhance understanding of factors that affect water 

quality, and establish links between sources of contaminants, the transport of those contaminants 

through the hydrologic system, and the potential effects of contaminants on humans and aquatic ecosystems.

The USGS aims to disseminate credible, timely, and relevant science information to inform practical 

and effective water-resource management and strategies that protect and restore water quality. We 

hope this NAWQA publication will provide you with insights and information to meet your needs, and 

will foster increased citizen awareness and involvement in the protection and restoration of our 

Nation’s waters. 

The USGS recognizes that a national assessment by a single program cannot address all waterresource issues of interest. External coordination at all levels is critical for a fully integrated understanding of watersheds and for cost-effective management, regulation, and conservation of our 

Nation’s water resources. The NAWQA Program, therefore, depends on advice and information from 

other agencies—Federal, State, interstate, Tribal, and local—as well as nongovernmental organizations, industry, academia, and other stakeholder groups. Your assistance and suggestions are greatly 

appreciated.

Robert M. Hirsch

Associate Director for Water
iv 
 v

Contents 

Abstract. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1

Introduction . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1

Description of the Basin . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1

Methods . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3

Aquatic Invertebrates of the Delaware River Basin . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3

Acknowledgments . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5

References Cited . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 27

Figures

1. Map showing ecoregions and locations and types of sampling sites within the 

Delaware River Basin. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .2

Tables 

1. Agencies and programs from which benthic-invertebrate data were obtained for 

checklist of taxa in the Delaware River Basin . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .4

2. Checklist of the aquatic invertebrates of the Delaware River Basin, 1990-2000 . . . . . . . . . . . . . . . . . . . . .6
vi 

Conversion Factors and Datum 

Multiply By To obtain

Area

square kilometer (km2) 0.3861 square mile (mi2)
A Checklist of the Aquatic Invertebrates of the Delaware 

River Basin, 1990-2000

By Michael D. Bilger, Karen Riva-Murray, and Gretchen L. Wall

Abstract

This paper details a compilation of aquatic-invertebrate 

taxa collected at 1,080 sites as part of 13 surface-water-quality 

studies completed by selected Federal, state, and local environmental agencies during 1990-2000, within the 32,893-km2 area 

of the Delaware River Basin. This checklist is intended to be a 

“working list” of aquatic invertebrates that can be applied successfully to the calculation and interpretation of various biological estimators to determine the status of water quality and can 

be used as a foundation to document the current state of biodiversity. It is not intended as a comprehensive historical inventory of the literature or of private and public holdings. A total of 

11 phyla comprising 20 classes, 46 orders, 196 families, 

685 genera, and 835 species were recorded.

Introduction 

Since the acceptance of the Saprobien system of Kolkwitz 

and Marsson (1908; 1909), the work of pioneering benthic scientists such as H.B. Noel Hynes (1960; 1970), and later 

researchers like Plafkin and others (1989), it was recognized 

that benthic-invertebrate assemblages were measurable indicators of localized conditions because of their complex life cycles 

of a year or more, their mostly sedentary nature, and their ability 

to react in a predictable way to human influences on aquatic 

ecosystems. These earlier efforts to document water-quality 

effects dealt mostly with organic pollutants. As the science has 

progressed, benthic assessments have evolved into multi-disciplined approaches that remain faunistic but also include habitat 

and geomorphic analyses of lotic systems. Newer efforts are 

concerned with understanding the native fauna and managing 

waters for maximized biological quality, including taking corrective action when monitoring indicates trouble (Cairns and 

Pratt, 1993). Barbour and others (1999) contains both multimetric and multivariate statistical methods for determining ecosystem effects on benthic-invertebrate communities. Accurate 

regional taxa lists and their accompanying pollution tolerance 

values are important to these assessments and interpretations 

and will gain increasing importance in the establishment of narrative or numeric regulatory values for biologically based criteria (U.S. Environmental Protection Agency, 1994). 

After the works of E.O. Wilson (1984; 1988; 2000), the 

subject of biodiversity has been identified to represent the basic 

fabric of life. Because biodiversity is in a rapidly diminishing 

phase, it is increasingly difficult to provide a full accountability 

of what sorts of organisms were present before human influences eliminated them. Even within the fauna of North America, it is estimated that less than 50 percent of the arthropod 

biodiversity has been described (Kosztarab and Schaefer, 1990; 

Rawlins and Bier, 1998). Taxonomic studies can record and 

describe new taxa, determine the phylogenetic relation and taxonomic status of various taxa, provide a sound classification, 

and produce taxonomic keys and other tools to aid in accurate 

identification (K.C. Kim, State College, Pa., unpubl. data, 

2001). Lists of taxa within the Delaware River Basin, as presented in this paper, will aid in the baseline effort to explain the 

status of the aquatic-invertebrate biodiversity even at the inconsistent level of taxonomic identification and assessment. 

Description of the Basin 

The non-tidal Delaware River Basin encompasses more 

than 32,893 km2 (fig. 1) and includes parts of the States of 

Pennsylvania, (16,744 km2), New Jersey (7,690 km2), New 

York (6,120 km2), and Delaware (2,507 km2) (Fischer, 2004). 

The headwaters of the Delaware River begin in the Catskill 

Mountains in the northern part of the basin. The river becomes 

tidally influenced in the area of Trenton, N.J., and is saline just 

south of Philadelphia, Pa. Two major tributaries of the Delaware River are the Lehigh (4,903 km2) and Schuylkill Rivers 

(3,520 km2). There are also a number of large reservoirs within 

the basin, including three operated by the City of New York for 

water supply and other uses.

The Delaware River Basin covers parts of eight ecoregions 

(Omernick, 1987; U.S. Environmental Protection Agency, 

1999): the Northeastern Highlands, Northern Appalachian Plateau and Uplands, North Central Appalachians, Middle Atlantic 

Coastal Plain, Northern Piedmont, Southeastern Plains, Ridge 

and Valley, and Atlantic Coastal Pine Barrens. A wide range of 
2 A Checklist of the Aquatic Invertebrates of the Delaware River Basin, 1990-2000

Figure 1. Ecoregions and locations and types of sampling sites within the Delaware River Basin. 
 3

topography describes the basin from nearly flat in the coastal 

area, to rolling hills and broad uplands, to ridges and valleys, to 

glaciated plateaus. Altitude varies from sea level in the south to 

more than 1,000 meters in the north. According to 1992 landuse data, about 60 percent of the basin is forested, 24 percent 

agricultural, 9 percent urban, and 7 percent surface-water bodies and other miscellaneous uses (Vogelmann and others, 

1998). Most of the population and urban land use is along the 

Delaware Estuary. Although the population within the basin has 

not increased significantly over the past 20 years, there has been 

a pattern of intense “urban flight” from the larger cities, creating 

an expansive suburbia. 

Methods 

Data were collected and compiled from Federal, state, and 

local environmental agencies active within the boundaries of 

the Delaware River Basin as defined by the U.S. Geological 

Survey (table 1). These agencies were selected on the basis of 

their common biomonitoring approach to document conditions 

and changes in surface-water quality from lotic habitats by 

using benthic-invertebrate community measures over roughly 

the past decade. Field-collection techniques were not standardized; only those data resulting from laboratory identifications 

were included (field-identified results from the Pennsylvania 

Department of Environmental Protection Unassessed Waters 

Program were not used). Historical records from extensive literature reviews and the examination of public and private specimen holdings were considered beyond the scope of this paper, 

as were data from distributional surveys of particular macroinvertebrate groups. It is hoped that this checklist be expanded in 

scope as a future endeavor and combined with other existing 

taxa lists such as those for Trichoptera (Masteller and others, 

2004; Masteller and Flint, 1992), Plecoptera (Masteller and others, 2004; J. Earle, Harrisburg, Pa., unpub. data, 2003), 

Ephemeroptera (G. Hoover, State College, Pa., unpub. data, 

2003), Simuliidae (Adler and Kim, 1986; D. Rebuck, Harrisburg, Pa., unpub. data, 2003), freshwater mussels (Strayer and 

Jirka, 1997; Lellis, 2001), Culicidae (M. Hutchinson, Harrisburg, Pa., unpub. data, 2003), Tabanidae (F. Fees, State College, Pa., unpub. data, 2003), Odonata (Donnelly, 2004a, 

2004b, 2004c; C. Schiffer, State College, Pa., unpub. data, 

2003), Tipulidae (Young and Gelhaus, 2000), Amphipoda and 

Isopoda (D. Glazier, Huntingdon, Pa., unpub. data, 2003) and 

crayfish (T. Nuttall, Lock Haven, Pa., unpub. data, 2003). 

The phylogenetic schemes employed in the checklist 

mostly followed those of Smith (2001) and Thorp and Covich 

(2001) for the non-insect arthropods and Merritt and Cummins 

(1996) for the insects. Some groups such as the annelids (Kathman and Brinkhurst, 1998; Wetzel, 2001), water mites (Smith, 

2001; Smith and Cook, 1991), mayflies (McCafferty, 2000), 

stoneflies (Stewart and Stark, 2002), caddisflies (Wiggins, 

1995), beetles (Epler, 1996) and midges (Epler, 1995, 2001; 

R. Bode, New York Department of Environmental Conservation, Albany, N.Y., written commun., 2001) required additional 

research to document the most current phylogeny. Although the 

senior author has extensively collected aquatic invertebrates 

within the Delaware River Basin, it was not possible to confirm 

all identifications of animals comprising the list. All taxa were 

reviewed for distributional accuracy (several taxa remain unresolved) and amended as necessary for synonomy and to correct 

“outdated taxonomy.” Any weaknesses within certain groups in 

the checklist likely resulted from the perceived difficulty in 

conducting identifications, especially for the oligochaetes, mollusks, chironomids, and several minor phyla. 

Aquatic Invertebrates of the Delaware 

River Basin

The compilation of a Delaware River Basin aquatic-invertebrate checklist incorporating multiple agency data (three 

county, four state, and four Federal) at 1,080 sites presented a 

challenge. Although these agencies have biomonitoring as a 

common goal, there was much latitude in the sampling designs, 

methods and seasons of collection, sample-site density, and levels of taxonomic resolution. Because this is a “working list” of 

taxa, the authors decided not to include data collected outside 

the 13 studies; however, the works of active taxonomists are 

mentioned in the methods section and their works will aid 

greatly in the assemblage of a comprehensive list of Delaware 

River Basin aquatic invertebrates.

At the end of 2000, the checklist consisted of 11 phyla 

comprising 20 classes, 46 orders, 196 families, 685 genera, and 

835 species (table 2). An examination of the checklist revealed 

a number of interesting points. The arthropods made up about 

83 percent of the total genera listed, and 95 percent of those 

were insects. Thirty percent of the total genera were dipterans; 

62 percent of those were chironomids, which themselves made 

up 18 percent of the total genera. The mayflies, stoneflies, and 

caddisflies, prominent animals in biomonitoring assessments, 

collectively made up 24 percent of the total genera listed. 

Twenty-eight genera were listed as collected in at least 10 of the 

13 studies and 10 genera were collected in 12 studies—Baetis, 

Isonychia, Stenonema, Sialis, Glossosoma, Polycentropus, 

Cheumatopsyche, Lepidostoma, Antocha, and Tipula. 

Assessments of the status of the Delaware River Basin 

invertebrate assemblage are difficult to interpret at this time. As 

the diversity is better defined and other basins like the Susquehanna River are documented for comparison, a more complete 

understanding of how large rivers function in their benthic ecology will evolve. For now, it is hoped this updated taxa checklist 

will provide a consistent basis for current and future regional 

synthesis or comparison of biomonitoring data. 
Table 1. Agencies and programs from which benthic-invertebrate data were obtained for checklist of taxa in the Delaware River Basin. [Number of sites sampled, and years during which sampling was conducted, pertain only to those data used in this report. DNREC, Delaware Department of Natural ResourceSTORET, U.S. Environmental Protection Agency Storage and Retrieval System; NJDEP, New Jersey Department of Environmental Protection; NYSDEC, New York State Deservation; PaDEP, Pennsylvania Department of Environmental Protection; USGS, U.S. Geological Survey] Agency and program(s) Sites Year(s) Reference(s) Data source 

Chester County Water Resources Authority Stream Conditions of Chester County Program 39 1990-2000 Reif, 1999; 2000; written commun., 2001 

Delaware Department of Natural Resources and Environ- mental Control Macroinvertebrate Survey 52 1993 John Maxted, DNREC, written commun., 1998 

Delaware River Basin Commission Scenic Rivers Monitoring Program 19 1990-92 STORET 

Monroe County Planning Commission Water Quality Studies 92 1990-2000 Monroe County Planning Commission, 1990-2000 New Jersey Department of Environmental Protection Ambient Biomonitoring Network 328 1990-2000 John Kurtz, NJDEP, written commun., 1995; 1996; 2001 Ken

199199

New York State Department of Environmental Conservation Rotating Intensive Basins Study Program 156 1993-2000 Margaret Novak, NYSDEC, written commun., 1998, 2001 Bod

199200

Pennsylvania Department of Environmental Protection Surface Water Quality Monitoring Network Anti-Degradation Program 33 150 1991-96 1990, 1992-97 Rodney Kime, PaDEP, written commun., 1998 Rodney Kime, PaDEP, written commun., 1998 She

Pike County Conservation District Surface Water Quality Monitoring Program 44 1991-2000 Pike County Conservation District, 1991-2000 U.S. Environmental Protection Agency 

Environmental Monitoring and Assessment Program Region 3

3225

1993-961994-96

U.S. Environmental Protection Agency, 2000 Passmore and Green, 1997

Kle

U.S. Geological Survey, Biological Resources Discipline 

Hemlock/ Hardwood Study 28 1997 Snyder and others, 1999

U.S. Geological Survey, Water Resources Discipline 

National Water-Quality Assessment Program Neversink Watersheds Studies Other New York District Studies

352126

1999-2000

1991-92, 1995-99

1997

unpublished data Barry Baldigo, USGS, written commun., 1999 Barry Baldigo, USGS, written commun., 1999

FisLaw
 5

Acknowledgments

This checklist could not have been completed without the 

help of the biologists from the selected agencies, Robert W. 

Bode (colleague reviewer) and Margaret A. Novak and Larry E. 

Abele from the New York State Department of Environmental 

Conservation, Albany, N.Y.; Rodney Kime from the Pennsylvania Department of Environmental Protection, Harrisburg, 

Pa.; Robert M. Ross from the U.S. Geological Survey, Biological Resources Discipline, Wellsboro, Pa.; Andrew G. Reif from 

the U.S. Geological Survey, Water Resources Discipline, Malvern, Pa.; Barry P. Baldigo from the U.S. Geological Survey, 

Water Resources Discipline, Troy, N.Y.; Allison R. Brigham, 

U.S. Geological Survey, Denver, Colo.; Eric Bartolacci from 

the Monroe County Planning Commission, Stroudsburg, Pa.; 

Sally Jones and Hannelore Schanzenbacher from the Pike 

County Conservation District, Hawley, Pa.; John R. Maxted 

from the Auckland Regional Council, Auckland, New Zealand, 

formerly of the Delaware Department of Natural Resources and 

Environmental Control, Dover, Del.; Edward D. Santoro and 

Robert L. Limbeck from the Delaware River Basin Commission, Trenton, N.J.; Richard Evans from the National Park Service, Milford, Pa.; Al Korndoerfer and John Kurtz from the 

New Jersey Department of Environmental Protection, Trenton, 

N.J.; Ellyn Del Corso Campbell currently with Gannett Fleming, Camp Hill, Pa.; and Jim Green (retired) and Margaret Passmore from the U.S. Environmental Protection Agency, Wheeling, W.Va.

The authors especially thank Jonathan G. Kennen from the 

U.S. Geological Survey, Water Resources Discipline, Trenton, 

N.J., who served as a colleague reviewer and technical and taxonomic expert; Robin A. Brightbill, U.S. Geological Survey, 

Water Resources Discipline, New Cumberland, Pa., for data 

analysis, organization, and guidance; J. Kent Crawford, U.S. 

Geological Survey, New Cumberland, Pa., for editorial review; 

and Jeffrey M. Fischer, U.S. Geological Survey, Water 

Resources Discipline, Trenton, N.J., who served as Project 

Chief of the Delaware River Basin NAWQA Project for his 

support. 
6 A Checklist of the Aquatic Invertebrates of the Delaware River Basin, 1990-2000

Table 2. Checklist of the aquatic invertebrates of the Delaware 

River Basin, 1990-2000. 

[Agency codes: C1, Chester County Water Resources Authority; C2, Monroe 

County Planning Commission; C3,Pike County Conservation District; F1,U.S. 

Environmental Protection Agency (USEPA), EMAP program; F2, USEPA Region 3; F3, Delaware River Basin Commission; F4, U.S. Geological Survey 

(USGS) Biological Resources Discipline; F5, USGS Water Resources Discipline (WRD) National Water-Quality Assessment Program; F6, USGS WRD, 

New York District; S1, Delaware Department of Natural Resources and Environmental Control; S2, New Jersey Department of Environmental Protection; 

S3, New York State Department of Environmental Conservation; S4, Pennsylvania Department of Environmental Protection]

Taxonomic name Agency code

PORIFERA F5

DEMOSPONGIAE

HAPLOSCLERIDA

Spongillidae

Anheteromeyenia argyrosperma S2

Ephydatia fluviatilis S2

Eunapius fragilis S2

Eunapius ingloviformis S2

Heteromeyenia S2

Heteromeyenia latitenta S2

Heteromeyenia tubisperma S2

Spongilla S2

Spongilla lacustris S2

Trochospongilla S2

CNIDARIA F6

HYDROZOA

HYDROIDA

Clavidae

Cordylophora lacustris S2

Hydridae

Hydra C1,F5,S2

PLATYHELMINTHES S2

TURBELLARIA F1,F2,F5,F6,S2,S4

MACROSTOMIDA

Macrostomidae

Macrostomum C3

PRORHYNCHIDA

Prorhynchidae

Geocentrophora baltica S2

PROSERIATA

Plagiostomidae

Hydrolimax S2

Hydrolimax grisea S2

TRICLADIDA S3

Dendrocoelidae

Procotyla S2

Procotyla fluviatilis S2

Dugesiidae

Cura S4

Cura foremanii S1,S2

Dugesia S2,S4

Dugesia tigrina S2

Planariidae C1,S4

Hymanella retenuova S2

Phagocata S2

Phagocata gracilis S2

Phagocata morgani morgani F1,S2

Phagocata velata S2

Phagocata woodworthi S2

Planaria C2,S4

NEMERTEA C1,S2,S4

ENOPLA

HOPLONEMERTEA

Tetrastemmatidae

Prostoma C1,F5,S2

Prostoma graecense S2,S3

NEMATODA C1,C2,F1,F5,F6,S2,S4

NEMATOMORPHA C2,F3

GORDIOIDEA

GORDIOIDA

Gordiidae

Gordius S4

ECTOPROCTA F5

GYMNOLAEMATA

CTENOSTOMATA

Paludicellidae

Paludicella S2

Paludicella articulata S2

PHYLACTOLAEMATA

PLUMATELLIDA

Fredericellidae

Fredericella S2

Fredericella sultana S2

Pectinatellidae

Pectinatella magnifica S2

Plumatellidae

Hyalinella punctata S2

Plumatella casmiana S2

Plumatella fruticosa S2

Plumatella repens S2

ANNELIDA C2,F3,S4

POLYCHAETA C1

SABELLIDA

Sabellidae C1

Taxonomic name Agency code
 7

Manayunkia speciosa F5,S2,S3,S4

APHANONEURA

AEOLOSOMATIDA

Aeolosomatidae S1

Aeolosoma F6,S2

BRANCHIOBDELLAE

BRANCHIOBDELLIDA F1,F6

Branchiobdellidae F1,S3

Xironogiton S4

OLIGOCHAETA C1,C2,C3,F1,F2,F3,F6,

S4

LUMBRICULIDA S4

Lumbriculidae C1,C2,C3,F1,F5,S1,S2,

S3,S4

Eclipidrilus F1,S2

Lumbriculus C1,C2,F1,S2

Lumbriculus variegatus F1,F4,S1,S2

Stylodrilus heringianus C3,F1,S2,S3

HAPLOTAXIDA

Haplotaxidae

Haplotaxis S2

Haplotaxis gordioides S2

TUBIFICIDA

Enchytraeidae F1,F4,F5,S1,S2,S3

Lumbricillus S2

Naididae C1,F5,S2,S4

Arcteonais S2

Arcteonais lomondi S2

Chaetogaster S2

Chaetogaster diaphanus F1,S2,S3

Chaetogaster diastrophus S2

Chaetogaster limnaei S2

Dero S1,S2,S3

Dero flabelliger S2

Dero furcata S2

Dero nivea S2

Dero obtusa S2

Nais F1,S1,S2

Nais behningi F1,S2,S3

Nais bretscheri F1,S2,S3

Nais communis F1,S2,S3

Nais elinguis S2

Nais pardalis F1

Nais pseudobtusa F1,S2

Nais simplex F1,S2,S3

Nais variabilis F1,S2,S3

Ophidonais serpentina S2,S3

Pristina S2

Pristina foreli S2

Taxonomic name Agency code

Pristina leidyi S2,S3

Pristina osborni S2

Slavina S2

Slavina appendiculata S2,S3

Specaria josinae F1,S2

Stylaria C1,S2,S4

Stylaria lacustris S1,S2,S3

Vejdovskyella comata S2

Tubificidae C1,F1,F5,S1,S2,S4

Undet. Tubificidae w/capilliform setaeF1

Undet. Tubificidae w/o capilliform setaeF1,S3

Aulodrilus S1,S2

Aulodrilus limnobius F1

Aulodrilus pigueti S2

Aulodrilus pluriseta F1,S2

Bothrioneurum vejdovskyanum F1

Branchiura sowerbyi S2

Limnodrilus F1,S2

Limnodrilus claparedianus F1,S2

Limnodrilus hoffmeisteri F1,S2,S3

Limnodrilus udekemianus S2

Monopylephorus helobius S2

Quistradrilus multisetosus S2,S3

Rhyacodrilus S2

Spirosperma F1,S1,S2

Spirosperma beetoni S2

Spirosperma ferox F1,S2,S3

Spirosperma nikolskyi F4,S2

Telmatodrilus S2

Telmatodrilus vejdovskyi S2

Tubifex C2,S2

Tubifex tubifex S2,S3

LUMBRICINA C2,C3,F6,S2,S3

Lumbricidae C2,F1,F4,S2,S4

Megascolecidae F4,S2

HIRUDINEA C2,C3,F5,S2,S3,S4

RHYNCHOBDELLIDA F3

Glossiphoniidae F5

Actinobdella S2

Alboglossiphonia heteroclita S2

Desserobdella phalera S2

Desserobdella picta S2

Gloiobdella elongata S2

Glossiphonia S4

Glossiphonia paludosa S2

Helobdella C1,S2

Helobdella fusca S2

Taxonomic name Agency code
8 A Checklist of the Aquatic Invertebrates of the Delaware River Basin, 1990-2000

Helobdella stagnalis F1,S2

Helobdella triserialis S2

Oligobdella biannulata S2

Placobdella S2,S4

Placobdella hollensis S2

Placobdella multilineata S2

Placobdella ornata S2

Placobdella papillifera S2

Placobdella parasitica S2

Placobdella translucens S2

Piscicolidae F3

Myzobdella lugubris C3,S2

Piscicola punctata S2

ARHYNCHOBDELLIDA

Erpobdellidae C1,F5,F6,S2

Dina anoculata S2

Erpobdella C1,S4

Erpobdella punctata S1, S2

Mooreobdella S2

Mooreobdella fervida S2

Mooreobdella melanostoma S2

Mooreobdella microstoma S2

Mooreobdella tetragon S2

Haemopidae

Haemopis marmorata S2

MOLLUSCA

GASTROPODA C1,C2,F3,F5,S4

MESOGASTROPODA

Bithyniidae

Bithynia C2,F1

Bithynia tentaculata S3

Hydrobiidae C2,F3,F5,F6,S2,S4

Amnicola C1,S2,S4

Amnicola grana S2

Amnicola limosa S2,S3

Birgella S2

Fontigens S2

Gillia S2

Gillia altilis S2

Hydrobia S2

Paludestrema bottimeri S2

Somatogyrus S2

Pleuroceridae S3,S4

Elimia C1,F5,S2,S4

Elimia virginica S2

Pleurocera S4

Pleurocera acuta S2

Valvatidae C2,F2

Taxonomic name Agency code

Valvata F6,S2,S4

Valvata bicarinata S2

Valvata piscinalis F6

Valvata tricarinata S2

Viviparidae C1

Campeloma C1,C2,S2,S4

Campeloma decisum S2

Cipangopaludina chinensis malleataF5

Viviparus georgianus S2

BASOMMATOPHORA S1

Ancylidae C2,F2,F3,F5,S4

Ferrissia C1,C2,C3,F1,F3,F5,S1,

S2,S3,S4

Ferrissia parallela S2

Ferrissia rivularis F1,F6,S2,S3

Laevapex F5,S4

Laevapex fuscus C3,S2

Lymnaeidae C2,F5,S1,S2,S3,S4

Fossaria C2,F5,S2,S4

Fossaria obrussa S2

Lymnaea C1,C2

Pseudosuccinea S1

Pseudosuccinea columella S2

Stagnicola F5,S2,S4

Stagnicola caperata S2

Stagnicola catascopium S2

Physidae C2,F5,F6,S4

Physinae C3

Aplexa elongata S2

Physa C1,C2,C3,F3,S2,S4

Physella F4,F5,S1,S2,S3,S4

Physella ancillaria S3

Physella gyrina S2

Physella heterostropha S2

Physella integra S2

Physella vinosa S2

Planorbidae C1,C2,F5,S2,S3,S4

Gyraulus C1,C2,F5,F6,S2,S4

Gyraulus circumstriatus S2,S3

Gyraulus deflectus S2

Gyraulus parvus C3,F4,S2

Helisoma C1,F5,S2

Helisoma anceps anceps S2

Menetus C1,F5,S1,S2,S4

Menetus dilatatus S2

Planorbella C3,F5,S2,S4

Planorbella campanulata C3

Planorbella trivolvis S2

Taxonomic name Agency code
 9

Planorbula C1,S2,S4

Promenetus S2

Promenetus exacuous S2

STYLOMMATOPHORA

Discidae

Discus cronkhitei S2

Helicodiscidae

Helicodiscus parallelus S2

Zonitidae

Zonitoides S2

Zonitoides arboreus S2

BIVALVIA F1,F3,F5,F6

UNIONOIDA

Unionidae C2,F5,S2

Alasmidonta undulata S2

Anodonta S2

Anodonta imbecillis F5

Anodonta implicata S2

Elliptio F5

Elliptio complanata F3,S2

Pyganodon cataracta S2

VENEROIDA

Corbiculidae

Corbicula F5,S2,S4

Corbicula fluminea F3,S1,S2

Sphaeriidae C1,C2,C3,F1,F2,F5,F6,

S2,S3,S4

Musculium C1,C2,C3,F1,F3,F5,S1,

S2,S4

Musculium partumeium S2

Musculium securis S2

Musculium transversum S2

Pisidium C1,C2,C3,F1,F3,F5,S1,

S2,S4

Pisidium casertanum F4,S2

Pisidium compressum S2

Pisidium dubium F1,S2

Pisidium fallax S2

Pisidium punctatum S2

Pisidium variabile S2

Pisidium walkeri S2

Sphaerium C1,C2,C3,F3,F5,S2,S3,

S4

Sphaerium fabale S2

Sphaerium occidentale S2

Sphaerium rhomboideum S2

Sphaerium simile S2

Sphaerium striatinum S2

CHELICERATA

Taxonomic name Agency code

ARACHNIDA

ACARI F5,F6

HYDRACHNIDIA C1,F1,F6,S4

TROMBIDIFORMES

Eylaoidea

Limnocharidae

Limnochares F1

Hydryphantoidea

Hydryphantidae F4

Hydrodromidae

Hydrodroma despiciens S2

Lebertioidea

Sperchonidae

Sperchon S2,S4

Sperchonopsis verrucosa S2

Lebertiidae

Lebertia F1,S1,S2

Oxidae

Oxus C3

Torrenticolidae

Torrenticola F4

Hygrobatoidea

Limnesiidae

Limnesia S2

Hygrobatidae

Hygrobates S2

Unionicolidae

Neumania S2

Unionicola S2

Pionidae

Piona S2

Aturidae

Brachypoda S2

Arrenuroidea

Krendowskiidae

Geayia ovata S2

Arrenuridae

Arrenurus S2

ORIBATIDA

Hydrozetidae

Hydrozetes S1,S2

ARTHROPODA

CRUSTACEA C2,F6

BRANCHIOPODA

CLADOCERA C1

Chydoridae

Eurycercus lamellatus S2

Daphnidae

Taxonomic name Agency code
10 A Checklist of the Aquatic Invertebrates of the Delaware River Basin, 1990-2000

Daphnia S2

Simocephalus exspinosus S2

Simocephalus vetulus S2

Macrothricidae

Acantholeberis curvirostris S2

Ilyocryptus acutifrons S2

Ilyocryptus sordidus S2

Sididae

Latona setifera S2

Sida S2

OSTRACODA

PODOCOPA C1

Candoniidae

Candona S2

Cyprididae S2

Cypricercus S2

Eucypris virens S2

COPEPODA

CALANOIDA

Diaptomidae

Diaptomus S2

HARPACTICOIDA

Canthocamptidae

Attheyella illinoisensis S2

CYCLOPOIDA C1

Cyclopidae C1

Acanthocyclops vernalis S2

Cyclops C1,C2,S2

Eucyclops S2

Macrocyclops albidus S2

Megacyclops viridis S2

Orthocyclops modestus S2

MALACOSTRACA

ISOPODA C2,F1,S4

Asellidae C1,C2,F2,F5

Caecidotea C1,C2,C3,F1,F5,F6,S1,

S2,S3,S4

Caecidotea forbesi S2

Caecidotea nodulus S2

Caecidotea obtusus S2

Caecidotea racovitzai S2, S3

Conasellus communis S2

Lirceus C1,C2,F1,F5,S2,S4

Lirceus brachyurus S2

Lirceus fontinalis S2

Lirceus lineatus S2

Cylisticidae S2

Cylisticus S2

Oniscidae

Taxonomic name Agency code

Porcellionides S2

AMPHIPODA C2,F3,F5,S4

Crangonyctidae S4

Crangonyx C1,C3,F1,F5,S1,S2,S4

Crangonyx pseudogracilis S2

Crangonyx richmondensis S2

Crangonyx serratus S2

Crangonyx shoemakeri S2

Stygonectes C3,F1,F4,S1,S2,S4

Gammaridae C1,C2,F1

Gammarus C1,C2,C3,F1,F3,F5,F6,

S1,S2,S3,S4

Gammarus fasciatus S2

Gammarus lacustris C3

Synurella chamberlaini S2

Talitridae C2,S4

Hyalella C2,C3,S2,S4

Hyalella azteca C1,C3,F5,S2

DECAPODA F5,S4

Cambaridae C1,C2,C3,F1,F2,F4,F5,

F6,S2,S3,S4

Cambarus C1,C2,F5,S1,S2,S4

Cambarus bartonii bartonii C3,S2

Orconectes C1,C2,F5,S2,S4

Orconectes limosus S2

Orconectes rusticus F1

Procambarus C1,C2

Procambarus acutus acutus S2

Palaemonidae

Palaemonetes S2

Palaemonetes paludosus S2

INSECTA

COLLEMBOLA C2,C3,F5,S4

Entomobryidae S2

Cyphoderus S2

Entomobrya S2

Tomocerus S2

Hypogastruridae

Anurida S2

Neanura S2

Tafallia S2

Xynella F4,S2

Isotomidae S4

Agrenia F3,F4

Archisotoma S2

Folsomia S2

Isotoma S1,S2

Isotomurus S2

Isotomurus palustris S1,S2

Taxonomic name Agency code
 11

Onychiuridae

Onychiurus S2

Poduridae

Podura S4

Podura aquatica S2

EPHEMEROPTERA C2,F1,F3,F5,F6

CARAPACEA

Prosopistomatoidea

Baetiscidae

Baetisca C2,F3,F6,S2,S3

Baetisca carolina S2

Baetisca obesa S2

FURCATERGALIA

Leptophlebioidea

Leptophlebiidae C1,C2,F1,F2,F5,F6,S1,

S2,S3,S4

Choroterpes S2

Habrophlebia C1,S4

Habrophlebia vibrans F1,S2

Habrophlebiodes C3,F1,S4

Leptophlebia S1,S2,S4

Paraleptophlebia C1,C2,C3,F1,F3,F4,F5,

F6,S2,S3,S4

Paraleptophlebia guttata F6,S3

Paraleptophlebia mollis C3,F6,S3

Paraleptophlebia volitans F6,S3

Ephemeroidea

Potamanthidae

Anthopotamus C1,F3,S2,S4

Anthopotamus myops S1

Polymitarcyidae

Ephoron leukon F3,S2,S4

Ephemeridae C2,F1,F2

Ephemera C1,C3,F1,F3,S2,S3,S4

Ephemera guttulata C3

Ephemera varia F1

Hexagenia C2,F3,S2,S4

Litobrancha recurvata C3,F1

Pentagenia F3

Caenoidea

Caenidae C2,F1,F2

Brachycercus F1,F5,F6

Caenis C1,C2,F1,F3,F5,F6,S1,

S2,S3,S4

Caenis anceps F5

Caenis cf. hilaris F5

Caenis latipennis S3

Caenis macafferti S3

Ephemerelloidea

Taxonomic name Agency code

Ephemerellidae C2,C3,F1,F2,F3,F5,S1,

S3,S4

Attenella C3,F3,F6,S4

Attenella attenuata C3,F5,F6,S2,S3

Attenella margarita F6

Dannella F1,F3,F6,S4

Dannella lita F1,S2

Dannella simplex C3,F1,S2

Drunella C3,F1,F3,F6,S2,S4

Drunella allegheniensis C3

Drunella cornuta F1,F6,S3

Drunella cornuta/cornutella C3

Drunella cornutella F1,F6,S2,S3

Drunella lata C2,C3,F6,S2,S3

Drunella tuberculata C3,F1,F5,F6,S3

Drunella walkeri C3,S2,S3

Ephemerella C1,C2,C3,F1,F3,F5,F6,

S1,S2,S3,S4

Ephemerella aurivillii S2

Ephemerella dorothea C3,F1,F4,F6,S2,S3

Ephemerella excrucians C3

Ephemerella inconstans F1

Ephemerella invaria C3,F1,S2,S3

Ephemerella invaria/rotunda C3,F1

Ephemerella needhami C3,S2

Ephemerella rotunda C3,F1,F4,F6,S2

Ephemerella septentrionalis C3,F1,S2

Ephemerella subvaria C3,S2,S3

Ephemerella/Serratella F6

Eurylophella C1,C3,F1,F5,F6,S1,S2,

S3,S4

Eurylophella bicolor C3,F1,S2

Eurylophella doris C3

Eurylophella funeralis C3,F1,F4

Eurylophella poconoensis C3

Eurylophella temporalis C3,F1,S2

Eurylophella temporalis/doris C3

Eurylophella verisimilis C3,F1

Serratella C1,C3,F1,F3,F5,F6,S1,

S2,S3,S4

Serratella deficiens C3,F1,F5,F6,S2,S3

Serratella frisoni F5

Serratella serrata F1,F5,F6,S2,S3

Serratella serratoides F6,S2,S3

Leptohyphidae C1,C2,F1,F2,F5

Tricorythodes C1,C2,C3,F1,F3,F5,F6,

S1,S2,S3,S4

PISCIFORMA

Baetoidea

Taxonomic name Agency code
12 A Checklist of the Aquatic Invertebrates of the Delaware River Basin, 1990-2000

Ameletidae

Ameletus C1,C2,C3,F1,F4,F6,S2,

S3,S4

Baetidae C2,F1,F2,F3,F5,S1,S3,

S4

Acentrella C3,F1,F3,F5,F6,S2,S3,

S4

Acentrella ampla C3,F1,F6,S1

Acentrella turbida C3,F5,F6,S2

Acerpenna C3,F5,S1,S4

Acerpenna macdunnoughi S3

Acerpenna pygmaea C3,F4,S2,S3

Baetis C1,C2,C3,F1,F3,F4,F5,

F6,S1,S2,S3,S4

Baetis brunneicolor C3,F6,S2,S3

Baetis flavistriga C3,F1,F5,F6,S2,S3

Baetis intercalaris C3,F1,F5,F6,S2,S3

Baetis pluto C3,S3

Baetis tricaudatus C3,F1,F4,F5,F6,S2,S3

Callibaetis C2,F1,F3,F5,S2

Centroptilum C1,C3,F1,F3,F5,S2,S4

Cloeon C2,C3,F1,F3,S2

Diphetor hageni C3,F6,S2

Heterocloeon C3,F3,F5,F6,S2,S4

Heterocloeon curiosum F5

Paracloeodes S2

Plauditus F5,S2

Plauditus cestus F6

Plauditus cingulatus C3,S2

Plauditus dubius S2

Plauditus parvulus S2

Plauditus punctiventris C3,S2

Procloeon F5,F6,S2

Pseudocloeon C1,C2,F3,F5,S2,S4

Pseudocloeon propinquum F5,S2,S3

Metretopodidae

Siphloplecton S2

Siphlonuridae C2,F1,F2,S1

Siphlonisca S4

Siphlonurus C1,C3,F3,S2,S3,S4

SETISURA

Heptagenioidea

Isonychiidae C1,C2,F1,F2

Isonychia C1,C2,C3,F1,F3,F4,F5,

F6,S1,S2,S3,S4

Isonychia arida S2

Isonychia bicolor S2,S3

Isonychia sayi S2

Taxonomic name Agency code

Heptageniidae C2,F1,F2,F3,F5,F6,S1,

S2,S3,S4

Cinygmula C3,F1,S2,S4

Cinygmula subaequalis C3

Epeorus C1,C2,C3,F1,F3,F5,F6,

S2,S4

Epeorus (Iron) F4,S3

Epeorus pleuralis/punctatus C3

Epeorus vitreus C3

Heptagenia C1,C2,C3,F3,F5,F6,S2,

S3,S4

Heptagenia flavescens S3

Heptagenia pulla S3

Leucrocuta C3,F1,F3,F5,F6,S2,S3,

S4

Leucrocuta hebe C3

Macdunnoa C3

Nixe C3,F1,F6,S4

Nixe lucidipennis S2

Rhithrogena C3,F4,F6,S4

Stenacron C1,C3,F1,F3,F5,S1,S2,

S4

Stenacron candidum S2

Stenacron carolina C3

Stenacron gildersleevi C3

Stenacron interpunctatum C3,F5,F6,S2,S3

Stenacron minnetonka C3,S2

Stenacron pallidum C3,S2

Stenonema C1,C2,C3,F1,F3,F5,F6,

S1,S2,S3,S4

Stenonema femoratum S3

Stenonema ithaca C3,F1,F5,F6,S2,S3

Stenonema luteum C3,S2

Stenonema mediopunctatum 

mediopunctatum

F5,S2

Stenonema meririvulanum F4

Stenonema mexicanum integrum S2,S3

Stenonema modestum C3,F1,F5,F6,S2,S3

Stenonema pudicum C3,F1,F4,F6,S2

Stenonema pulchellum F5

Stenonema terminatum F1,F6,S2,S3

Stenonema vicarium F4,F5,S2,S3

ODONATA C2

ANISOPTERA F1

Aeschnidae C2,F1,F2,F5

Aeschna C1,C2,F5,S2

Aeschna interrupta S2

Anax S2

Basiaeschna S2

Taxonomic name Agency code
 13

Basiaeschna janata S2

Boyeria C1,C2,C3,F1,F5,F6,S2,

S3,S4

Boyeria vinosa F1,F4,F5,S1,S2

Gomphaeschna furcillata S2

Cordulegastridae C2,F2

Cordulegaster C2,C3,F1,F4,S2,S3,S4

Cordulegaster diastatops S2

Cordulegaster maculata F1,S2

Corduliidae F2

Didymops transversa S2

Dorocordulia S2

Epicordulia S2

Epitheca F5

Helocordulia F1,S2

Helocordulia uhleri S2

Neurocordulia C3,F5,S2

Neurocordulia obsoleta S2

Somatochlora S2

Somatochlora forcipata S2

Somatochlora provocans S2

Somatochlora tenebrosa S2

Tetragoneuria S2

Gomphidae C1,C2,C3,F1,F2,F5,S1,

S2,S3,S4

Arigomphus C3,F1,S2,S4

Dromogomphus C3,F1,F5,S2,S4

Dromogomphus spinosus S2

Erpetogomphus F5,S4

Gomphus C1,C2,C3,F1,F5,S1,S2,

S4

Gomphus exilis S2

Gomphus spicatus S2

Hagenius C3

Hagenius brevistylus C2,F5,S2

Lanthus C1,C2,C3,F1,F5,F6,S2,

S3,S4

Lanthus parvulus F1,F4,F5,S2

Lanthus/Stylogomphus C2

Octogomphus specularis S2

Ophiogomphus C1,C2,C3,F5,F6,S2,S3,

S4

Progomphus S2

Progomphus obscurus S2

Stylogomphus C1,C2,C3,S4

Stylogomphus albistylus F1,F5,S1

Stylurus C3

Libellulidae F1,F2,S2

Erythemis F1,S2

Taxonomic name Agency code

Erythemis simplicicollis S2

Erythrodiplax S2

Libellula C2,F5,S1,S2

Libellula exusta S2

Libellula vibrans S2

Pachydiplax S2

Pachydiplax longipennis S2

Perithemis F5,S2

Perithemis domitia S2

Plathemis S2

Plathemis lydia S2

Sympetrum S2

Tramea carolina S2

Macromiidae C2,F5,S4

Macromia C1,C3,F5,S2

Macromia illinoisensis F5,S2

ZYGOPTERA

Calopterygidae C2,F2,F5,S2

Calopteryx C1,C2,C3,F1,F5,S1,S2,

S3,S4

Hetaerina C1,C2,S2,S3,S4

Hetaerina americana S2

Coenagrionidae C1,C2,F2,F5,S1,S2,S4

Amphiagrion C1,S2

Argia C1,C2,C3,F5,S1,S2,S4

Argia apicalis C3,S2

Argia bipunctulata S2

Argia fumipennis violacea C3,S2

Argia moesta F5,S2

Chromagrion S2,S4

Chromagrion conditum S2

Coenagrion S4

Enallagma C1,F5,S2

Enallagma exsulans S2

Ischnura C1,F4,S2

Ischnura posita S2

Ischnura verticalis S2

Nehalennia C1

Lestidae

Lestes S2

Lestes vigilax S2

HEMIPTERA C2

Belostomatidae

Belostoma F5,S2

Lethocerus americanus S2

Corixidae C1,C2,F5,S2,S4

Callicorixa S2

Hesperocorixa F4,S2

Palmacorixa C2,S2

Taxonomic name Agency code
14 A Checklist of the Aquatic Invertebrates of the Delaware River Basin, 1990-2000

Ramphocorixa S2

Sigara C1,F5,S2

Trichocorixa C1,F5,S2,S4

Trichocorixa verticalis S2

Gerridae C1,C2,F5,S2

Gerrinae F5

Aquarius S2

Aquarius remigis F5,S2

Gerris C1,C2,S2

Gerris marginatus S2

Limnoporus F1

Trepobatinae F5

Metrobates C1,F5,S2,S4

Metrobates hesperius F5,S2

Trepobates C1,F5,S2

Trepobates inermis F5

Trepobates pictus S2

Rhagadotarsinae

Rheumatobates C1,S2

Hebridae

Hebrus S2

Hydrometridae

Hydrometra S2

Mesoveliidae

Mesovelia C1,F5,S2

Mesovelia mulsanti S2

Naucoridae

Pelocoris S2

Nepidae

Nepa apiculata S2

Ranatra S2

Ranatra fusca F5

Ranatra protensa S2

Notonectidae

Notonecta F5,S2

Notonecta insulata S2

Notonecta irrorata S2

Notonecta uhleri S2

Pleidae

Neoplea striola S2

Saldidae F1

Veliidae C2

Microvelia C1,C3,F5,S1,S2,S4

Microvelia borealis S2

Rhagovelia C1,C2,F1,F5,S1,S2,S4

Rhagovelia obesa F5,S2

PLECOPTERA F1,F3,F5,S1,S4

EUHOLOGNATHA

Taxonomic name Agency code

Capniidae C1,C2,F1,F2,F6,S3,S4

Allocapnia C1,C2,F1,F4,S1,S2,S4

Paracapnia C1,F4,S2,S4

Paracapnia angulata S2

Paracapnia opis C3,S2

Leuctridae C2,F1,F2,F3,F5,F6,S3,

S4

Leuctra C1,C2,C3,F1,F3,F6,S2,

S3,S4

Leuctra ferruginea C3,F6,S3

Leuctra sibleyi F4

Leuctra tenuis F6,S2

Leuctra truncata S2

Paraleuctra C1,S2

Paraleuctra sara S2

Nemouridae C1,C2,F1,F2,S4

Amphinemura C2,C3,F1,F6,S2,S3,S4

Amphinemura delosa C3,F4,S2

Amphinemura wui F4

Nemoura C2,S2,S4

Nemoura trispinosa S2

Ostrocerca S4

Ostrocerca truncata S2

Paranemoura C1,S4

Paranemoura perfecta F4

Prostoia S4

Prostoia similis F4,S2

Shipsa rotunda S2

Soyedina F1

Taeniopterygidae C2,C3

Bolotoperla rossi F4

Oemopteryx contorta F4

Oemopteryx glacialis S2

Strophopteryx C1,S4

Strophopteryx fasciata S2

Taenionema S2,S4

Taeniopteryx C1,F1,S1,S2,S4

Taeniopteryx burksi S2

Taeniopteryx nivalis S2

Taeniopteryx parvula S2

SYSTELLOGNATHA

Chloroperlidae C1,C2,C3,F1,F2,F5,F6,

S4

Alloperla C3,F1,F6,S4

Haploperla C1,C2,F1,S4

Haploperla brevis S2

Rasvena C3

Suwallia C3,F1,S4

Taxonomic name Agency code
 15

Sweltsa C2,C3,F1,F4,F5,F6,S3,

S4

Peltoperlidae C2,C3,F1,F2,S4

Peltoperla C1,C2,F5,S2,S4

Tallaperla C3,F1,F4,F5,F6,S1,S2,

S3,S4

Perlidae C1,C2,C3,F1,F2,F3,F5,

F6,S3,S4

Acroneuria C1,C2,C3,F1,F3,F5,F6,

S1,S2,S3,S4

Acroneuria abnormis C2,C3,F1,F4,F6,S2,S3

Acroneuria carolinensis C2,C3,F1,F4,F6,S2,S3

Acroneuria internata C2,C3

Acroneuria lycorias S2

Agnetina C1,C2,C3,F1,F3,F5,F6,

S4

Agnetina annulipes S3

Agnetina capitata C2,C3,F4,F5,F6,S2,S3

Attaneuria S4

Attaneuria ruralis C2

Eccoptura C3,F1,S4

Eccoptura xanthenes C3,S2

Neoperla C1,C2,C3,F3,S4

Neoperla clymene C3,S2

Paragnetina C1,C2,C3,F1,F3,F5,F6,

S2,S3,S4

Paragnetina immarginata C2,C3,F5,F6,S2,S3

Paragnetina media C2,C3,F6,S2,S3

Perlesta C2,C3,F1,F3,F5,F6,S4

Perlesta placida C3,S2

Perlinella F1,S2

Perlinella drymo S2

Perlodidae C1,C2,C3,F1,F2,F6,S3,

S4

Clioperla S4

Cultus C3

Diploperla S4

Helopicus S4

Isogenoides C3,S4

Isoperla C1,C2,C3,F1,F6,S1,S2,

S3,S4

Isoperla bilineata C3,F1

Isoperla holochlora C3,S3

Isoperla lata C3,F1

Isoperla marlynia C3,F4,S2

Isoperla orata C3

Isoperla similis C3,F4,S2

Isoperla transmarina C3,S2

Malirekus F6,S4

Malirekus iroquois F4

Taxonomic name Agency code

Remenus S4

Remenus bilobatus C3

Yugus S4

Yugus bulbosus C2,C3,F1

Pteronarcyidae C2,F2

Pteronarcys C2,C3,F1,F5,F6,S2,S4

Pteronarcys biloba C3,F4,S3

Pteronarcys comstocki C3

Pteronarcys proteus C3,F4

COLEOPTERA F3,S4

ADEPHAGA

Gyrinidae C1,C2,F2

Dineutus C1,C2,C3,F3,F5,F6,S2,

S4

Dineutus discolor F5

Gyrinus C3,F1,F5,S2

Haliplidae

Haliplus C2,S2,S4

Peltodytes F1,F5,S2

Peltodytes lengi F4

Dytiscidae C1,C2,F1,F2,F6,S4

Acilius S2

Agabetes S2

Agabus C1,C2,F6,S2,S3

Bidessus S2

Copelatus S2

Cybister S2

Deronectes S2

Derovatellus S2

Dytiscus C3,S4

Eretes S2

Hydroporini F5

Hydroporus S1,S2

Hydroporus niger S2

Hydrovatus S2

Hygrotus C1

Ilybius S2

Laccophilus C1,C3,F5,S2,S4

Oreodytes F5,F6

Thermonectus S2

Uvarus S2

Noteridae C2

Hydrocanthus C2,C3

Hydrocanthus iricolor S2

POLYPHAGA

Helophoridae

Helophorus C1,F1,S2

Hydrochidae

Hydrochus C3,F5,S2

Taxonomic name Agency code
16 A Checklist of the Aquatic Invertebrates of the Delaware River Basin, 1990-2000

Hydrophilidae C1,C2,F2,S3,S4

Anacaena S2

Berosus C1,C2,F5,S1,S2,S4

Cercyon S2

Chaetarthria S2

Crenitis F4

Derallus C3

Enochrus F4,S2

Helochares S1,S2

Helocombus S2

Helocombus bifidus S2

Hydrobius C1,F1,S2,S3,S4

Hydrochara C1

Laccobius F1,S2

Paracymus F5,S2

Sperchopsis F1,S2

Tropisternus C3,F5,S2

Staphylinidae S2

Bledius S2

Diglotta S2

Stenus S2

Salpingidae S2

Hydraenidae C2

Limnebius C1

Psephenidae C2,F2

Ectopria C1,C2,C3,F1,F5,S2,S3,

S4

Ectopria nervosa C1,F4,S1,S2,S3

Psephenus C1,C2,C3,F1,F3,F6,S2,

S3,S4

Psephenus herricki C3,F1,F4,F5,S1,S2,S3

Dryopidae C1

Helichus C1,F4,S4

Helichus basalis F5,S2

Helichus fastigiatus C1,F1,S2

Helichus lithophilus S2

Scirtidae F5

Cyphon S2

Elodes S2

Scirtes S2

Elmidae C2,F1,F2,F3,F5,F6,S1,

S2,S4

Ancyronyx C1,C3,S2,S4

Ancyronyx variegata C1,F5,S1,S2,S3

Dubiraphia C1,C3,F1,F5,F6,S1,S2,

S3,S4

Dubiraphia bivittata S2

Dubiraphia quadrinotata S2

Dubiraphia vittata S3

Taxonomic name Agency code

Gonielmis S4

Macronychus C1,C3,F3,S2,S4

Macronychus glabratus C1,F1,F4,F5,F6,S1,S2,

S3

Microcylloepus C1,C3,F3,S4

Microcylloepus pusillus F5,S1,S2

Optioservus C1,C2,C3,F1,F3,F5,F6,

S1,S2,S3,S4

Optioservus fastiditus S3

Optioservus immunis F1,S2,S3

Optioservus ovalis F1,F5,F6,S2,S3

Optioservus trivittatus F1,F5,F6,S2,S3

Oulimnius C1,C3,F1,F3,F6,S2,S3,

S4

Oulimnius latiusculus F1,F4,F5,F6,S1,S2

Promoresia C1,C2,C3,F1,F3,F5,F6,

S1,S2,S4

Promoresia elegans F5,F6,S2,S3

Promoresia tardella F1,F4,F6,S2,S3

Stenelmis C1,C2,C3,F1,F3,F5,F6,

S1,S2,S3,S4

Stenelmis bicarinata S3

Stenelmis cheryl S3

Stenelmis concinna F1,F5,S2,S3

Stenelmis crenata F1,F5,S2,S3

Stenelmis decorata S2

Stenelmis humerosa S2

Stenelmis lateralis S2

Stenelmis markeli S2

Stenelmis mera F1,S2

Stenelmis mirabilis S2

Stenelmis musgravei F5

Stenelmis sandersoni F5,S2

Stenelmis vittipennis S2

Ptilodactylidae C2

Anchytarsus C2,C3,F1,S2,S4

Anchytarsus bicolor C1,F5,S1,S2

Lutrochidae

Lutrochus C3

Chrysomelidae C1

Agasicles S1

Donacia F1,S2

Galerucella S2

Curculionidae C1,C3,F4,F5,S2,S4

Hyperodes S2

Onychylis S2

Stenopelmus rufinasus S2

Lampyridae

Photurus S2

Taxonomic name Agency code
 17

MEGALOPTERA C2,C3

Corydalidae C2,F2

Chauliodes C1,C2,C3,F3,S2

Chauliodes pectinicornis S2

Corydalus C1,C2,C3,F3,S2,S4

Corydalus cornutus F5,F6,S1,S2,S3

Nigronia C1,C2,C3,F1,F6,S1,S2,

S4

Nigronia fasciatus F4

Nigronia serricornis C3,F1,F4,F5,S2,S3

Sialidae C2,F2

Sialis C1,C2,C3,F1,F3,F4,F5,

F6,S1,S2,S3,S4

Sialis hasta S2

Sialis iola S2

Sialis joppa S2

Sialis mohri S2

Sialis velata S2

NEUROPTERA

Sisyridae C1,C2

Climacia C1

Climacia areolaris C1,C3,S2

Sisyra F5

TRICHOPTERA F1,F3,F5,F6,S4

SPICIPALPIA

Rhyacophilidae C2,F2,F3

Rhyacophila C1,C2,C3,F1,F3,F5,F6,

S1,S2,S3,S4

Rhyacophila acutiloba C3,F5

Rhyacophila amicus S2

Rhyacophila carolina C3,F6,S3

Rhyacophila carpenteri F6,S3

Rhyacophila fenestra S2

Rhyacophila formosa C3,F5

Rhyacophila fuscula C1,C2,C3,F1,F4,F5,F6,

S2,S3

Rhyacophila glaberrima C3,F4,S2

Rhyacophila invaria S2

Rhyacophila manistee S2

Rhyacophila melita C2,C3,F6,S2,S3

Rhyacophila minora F4

Rhyacophila nigrita C3,F4,S2

Rhyacophila sp. 4 S3

Rhyacophila sp. 5 S2

Rhyacophila torva C3,F4,S3

Hydroptilidae C2,F2,F5,F6,S3,S4

Ptilocolepinae

Palaeagapetus S4

Palaeagapetus celsus F1,F4

Taxonomic name Agency code

Hydroptilinae

Agraylea C1,S2,S3

Alisotrichia S4

Hydroptila C1,C2,F1,F3,F4,F5,F6,

S1,S2,S3,S4

Hydroptila nr. albicornis sp. 1 S3

Hydroptila consimilis S3

Hydroptila nr. hamata S2

Hydroptila nr. hamata sp. 1 S3

Hydroptila nr. waubesiana S2

Leucotrichia C1,C3,F5,F6,S2,S3,S4

Leucotrichia pictipes F5,S1,S2

Neotrichia C2

Ochrotrichia C1,S2,S4

Orthotrichia S2

Oxyethira F6,S2,S4

Oxyethira pallida F5

Stactobiella C1

Glossosomatidae F1,F2,F3,F5,F6,S4

Glossosomatinae

Glossosoma C1,C2,C3,F1,F3,F4,F5,

F6,S1,S2,S3,S4

Agapetinae

Agapetus C3,F1,F6,S2,S4

Protoptilinae

Culoptila C1,F3,F5,F6

Protoptila C1,F6,S2,S3,S4

ANNULIPALPIA

Philopotamoidea

Philopotamidae C1,C2,F1,F2,F3,S4

Philopotaminae

Dolophilodes C1,C2,C3,F1,F5,F6,S1,

S2,S3,S4

Dolophilodes distinctus C3,F4,F5

Dolophilodes major C3

Wormaldia C1,C2,C3,F1,S2,S3,S4

Wormaldia moesta C3

Wormaldia shawnee C3

Chimarrinae

Chimarra C1,C2,C3,F1,F3,F5,F6,

S1,S2,S3,S4

Chimarra aterrima C3,S2,S3

Chimarra obscura C3,S2,S3

Chimarra socia S2,S3

Hydropsychoidea

Psychomyiidae S2

Psychomyiinae

Lype C1,C3,S1,S2,S4

Lype diversa F1,F4,S2,S3

Taxonomic name Agency code
18 A Checklist of the Aquatic Invertebrates of the Delaware River Basin, 1990-2000

Psychomyia C1,F1,F5,S2,S4

Psychomyia flavida F5,F6,S3

Psychomyia nomada F5,S2

Tinodes C2,S2

Dipseudopsidae

Phylocentropus C1,F1,S1,S2

Polycentropodidae C2,F1,F2,F5,S1,S3,S4

Polycentropodinae

Cernotina F1,S4

Cyrnellus C1,S4

Neureclipsis C1,C3,F3,F5,F6,S2,S3,

S4

Nyctiophylax C1,F1,F3,S2,S4

Nyctiophylax moestus S1

Polycentropus C1,C2,C3,F1,F3,F4,F5,

F6,S1,S2,S3,S4

Hydropsychidae C2,C3,F1,F2,F3,F5,F6,

S1,S3,S4

Arctopsychinae

Arctopsyche F6

Parapsyche C1,C2,F1,F6,S4

Parapsyche apicalis F4,F6

Diplectroninae

Diplectrona C1,C3,F1,F6,S2,S4

Diplectrona modesta C3,F1,F4,F5,S1,S2

Hydropsychinae

Ceratopsyche C1,C3,F1,F3,F5,S2,S4

Ceratopsyche aenigma F5

Ceratopsyche alhedra F1,F5,S2

Ceratopsyche bifida gr. C2,F1

Ceratopsyche bronta C3,F1,F5,F6,S2,S3

Ceratopsyche cheilonis F5

Ceratopsyche morosa C3,F1,F5,F6,S2,S3

Ceratopsyche slossonae C3,F1,F5,S2,S3

Ceratopsyche sparna C2,C3,F1,F4,F5,F6,S2,

S3

Ceratopsyche ventura C3,F1,F4

Ceratopsyche vexa C3

Ceratopsyche walkeri C3,F1,F5

Cheumatopsyche C1,C2,C3,F1,F3,F4,F5,

F6,S1,S2,S3,S4

Cheumatopsyche campyla S2

Hydropsyche C1,C2,C3,F1,F3,F5,F6,

S1,S2,S3,S4

Hydropsyche betteni C2,C3,F1,F5,S2,S3

Hydropsyche decalda S2

Hydropsyche demora F5,S2

Hydropsyche depravata gr. F5

Hydropsyche leonardi F5,S2,S3

Taxonomic name Agency code

Hydropsyche scalaris F5,S2,S3

Hydropsyche valanis S2

Hydropsyche venularis C3,F5,S2,S3

Potamyia C1,F3

Macronematinae

Macrostemum C1,C2,C3,F3,F5,F6,S2,

S3,S4

Macrostemum carolina S2,S3

Macrostemum zebratum F5

INTEGRIPALPIA

Phryganeoidea

Phryganeidae C2,F2

Phryganeinae

Agrypnia S2

Banksiola S2

Oligostomis F5,F6,S2,S3

Phryganea S2

Ptilostomis S2

Limnephiloidea

Brachycentridae C2,F2,S2,S3

Adicrophleps hitchcocki F4,F6,S4

Brachycentrus C1,C2,C3,F1,F3,F5,F6,

S2,S4

Brachycentrus americanus S2

Brachycentrus appalachia F5,F6,S3

Brachycentrus lateralis C3,F5,S2,S3

Brachycentrus nigrosoma F1

Brachycentrus numerosus F5,F6,S2,S3

Brachycentrus solomoni F5,S3

Micrasema C1,C2,C3,F1,F3,F5,F6,

S2,S3,S4

Micrasema rusticum S2

Micrasema sprulesi F4

Micrasema wataga C3,F5,S2

Micrasema sp. 1 S3

Micrasema sp. 2 S3

Lepidostomatidae C2,F2,S2

Lepidostomatinae

Lepidostoma C1,C2,C3,F1,F3,F4,F5,

F6,S1,S2,S3,S4

Theliopsychinae

Theliopsyche S2

Limnephilidae C1,C2,F1,F2,F6,S1,S2,

S3,S4

Dicosmoecinae

Ironoquia F1,F4,S2

Onocosmoecus C2,F1,F6

Pseudostenophylacinae

Pseudostenophylax F1,F3,F6,S2,S4

Taxonomic name Agency code
 19

Limnephilinae

Frenesia F5,S2

Hesperophylax F1,F6,S2,S4

Hesperophylax argus F1

Hydatophylax C1,C2,C3,F6,S1,S4

Ironoquia C3

Lenarchus S2

Limnephilus C2,F6,S2,S4

Platycentropus C3,S2

Psychoglypha subborealis F5

Pycnopsyche C2,C3,F1,F4,F5,F6,S2,

S3,S4

Pycnopsyche gentilis C3

Apataniidae C1

Apatania C1,C2,F1,F3,F5,F6,S2,

S3,S4

Uenoidae C2

Thremmatinae

Neophylax C1,C2,C3,F1,F3,F5,F6,

S2,S3,S4

Neophylax concinnus F4

Neophylax consimilis F5,S1

Neophylax nacatus F4

Goeridae

Goerinae F1

Goera C1,F1,S2,S3,S4

Goerita S2

Leptoceroidea

Leptoceridae C1,C2,F1,F2,F3,F5,F6,

S2,S4

Leptocerinae

Ceraclea C1,C2,F1,F3,F5,F6,S2,

S4

Leptocerus S2

Mystacides C1,C2,F1,F6,S2,S3,S4

Mystacides sepulchralis F5,S2

Nectopsyche C1,F1,S2

Oecetis C1,C2,C3,F1,F5,F6,S2,

S3,S4

Oecetis avara C3,F5,F6

Oecetis cinerascens F5

Oecetis persimilis F5,F6

Oecetis sp. A F5

Setodes S4

Triaenodes C1,F1,S2,S4

Molannidae C2

Molanna C1,C2,F1,F3,S2,S4

Molanna blenda F4

Calamoceratidae

Taxonomic name Agency code

Anisocentropus pyraloides C1

Heteroplectron S2

Heteroplectron americanum S2

Odontoceridae C2,F2

Odontocerinae

Psilotreta C1,C2,C3,F1,F3,F5,F6,

S2,S3,S4

Psilotreta frontalis F4,S2

Psilotreta labida C3,F5

Sericostomatoidea

Sericostomatidae

Agarodes S2,S4

Beraeidae

Beraea C2

Helicopsychidae C2

Helicopsyche C1,C3,F3,F6,S2,S4

Helicopsyche borealis C3,F5,S2,S3

LEPIDOPTERA C3,F4,S1,S3

Nepticulidae

Nepticula S2

Noctuidae C1

Archanara C1

Pyralidae C1,C2,C3,F1,F2,F6,S2

Acentria C2,S2

Crambus F5

Elophila S2

Nymphula C1

Nymphuliella S2

Parapoynx S2,S4

Petrophila C1,F5,S1,S2,S3,S4

Synclita C1,S2

HYMENOPTERA

APOCRITA

Ichneumonoidea

Ichneumonidae C1

DIPTERA C2,C3,F1,F3,F6,S1

NEMATOCERA

Blephariceridae C1,C2,S3

Blepharicera C3,F1,F4,F5,F6,S2,S4

Ceratopogonidae C1,C3,F1,F2,F3,F5,F6,

S2,S3,S4

Dasyheleinae F1

Dasyhelea F1,F5,S1,S4

Forcipomyiinae

Atrichopogon S1,S2

Ceratopogoninae F1,F6

Alluaudomyia S2

Alluaudomyia needhami S2

Bezzia C3,F1,F4,F5,S1,S2,S4

Taxonomic name Agency code
20 A Checklist of the Aquatic Invertebrates of the Delaware River Basin, 1990-2000

Bezzia glabra S2

Bezzia opaca S2

Bezzia setulosa S2

Bezzia varicolor S2

Ceratopogon F1,F4

Culicoides S2,S4

Culicoides sangusuigus S2

Palpomyia F4,F5,S2

Palpomyia lineata S2

Palpomyia tibialis S2

Probezzia C3,F1,F4,S1,S2,S4

Stilobezzia S2

Stilobezzia antennalis S2

Chaoboridae

Chaoborus C1,S2,S4

Chaoborus punctipennis S2

Chironomidae C1,C2,C3,F1,F2,F3,F5,

F6,S1,S2,S4

Tanypodinae C2,C3,F1,F5,F6,S2,S4

Coelotanypodini

Clinotanypus F1,F5,S2

Clinotanypus pinguis S2

Coelotanypus tricolor S2

Macropelopiini

Apsectrotanypus F1,S2

Apsectrotanypus trifascipennis S2

Brundiniella F4

Brundiniella eumorpha F1

Macropelopia F1,F6,S2

Macropelopia decedens S2

Psectrotanypus S2

Psectrotanypus dyari S2

Natarsiini

Natarsia F1,S1,S2

Natarsia baltimoreus F1,F4

Natarsia fastuosa S2

Pentaneurini

Ablabesmyia C1,F1,F5,F6,S1,S2

Ablabesmyia annulata S2

Ablabesmyia aspera S2

Ablabesmyia janta S2

Ablabesmyia mallochi F1,S2

Ablabesmyia monilis S2

Ablabesmyia peleensis S2

Ablabesmyia simpsoni S2

Cantopelopia aleta S2

Conchapelopia C1,F1,F4,F6,S1,S2

Conchapelopia americana F1,S2

Conchapelopia currani S2

Taxonomic name Agency code

Conchapelopia fasciata S2

Conchapelopia flavifrons F1,S2

Conchapelopia pallens S2

Conchapelopia rurika S2

Conchapelopia telema S2

Hayesomyia F1

Hayesomyia senata S2

Helopelopia F1

Helopelopia cornuticaudata S2

Hudsonimyia karelena S3

Labrundinia F5,S2,S4

Labrundinia pilosella S2

Larsia F1,F4

Meropelopia F1,F4,S1

Meropelopia americanus F1

Meropelopia flavifrons F1

Nilotanypus C1,F1,F6,S4

Nilotanypus fimbriatus F1,S2

Paramerina F1,F5

Pentaneura F5,F6,S2,S3,S4

Pentaneura carnea S2

Pentaneura inconspicua S3

Rheopelopia F1,F4,S1,S2

Rheopelopia perda S2

Telopelopia F1

Telopelopia okoboji F6,S2

Thienemannimyia F1,S1,S2,S4

Thienemannimyia gr. F1,F5,F6,S2,S3

Thienemannimyia norena S2

Trissopelopia F1,F5,F6,S2

Zavrelimyia F1,F4,F6,S1,S3

Procladiini

Djalmabatista S2

Procladius C1,F1,F5,F6,S2,S4

Procladius (Psilotanypus) bellus F1,S2

Procladius culiciformis S2

Procladius curtus S2

Procladius riparius S2

Tanypodini

Tanypus F1,S2

Tanypus neopunctipennis S2

Tanypus punctipennis S2

Tanypus stellatus S2

Diamesinae F5

Diamesini

Diamesa C1,F1,F4,F5,F6,S1,S2,

S3,S4

Diamesa nivoriunda S2

Pagastia F1,F5,F6

Taxonomic name Agency code
 21

Pagastia orthogonia S2

Pagastia sp. A S3

Potthastia C1,F1,F5,F6

Potthastia gaedii F6,S1,S3

Potthastia gaedii gr. F1

Potthastia longimana F6,S2,S3

Pseudodiamesa C1,F1

Pseudodiamesa pertinax S2

Sympotthastia F6,S2,S4

Sympotthastia sp. 2 S2

Protanypini

Protanypus S2

Prodiamesinae

Monodiamesa bathyphila S2

Odontomesa F1,S2

Prodiamesa S2

Prodiamesa olivacea F1,S2

Orthocladiinae C2,C3,F1,F5,F6,S2,S4

Corynoneurini

Corynoneura C1,F1,F4,F5,F6,S1,S2,

S3,S4

Corynoneura taris S2

Orthocladiini F1

Acamptocladius S2

Acricotopus S2

Brillia C1,F1,F5,F6,S1,S2,S3,

S4

Brillia flavifrons F1,F4,S2

Brillia flavifrons gr. F1

Brillia modesta S2

Brillia parva F1,F4,S2,S3

Brillia sera F4

Bryophaenocladius S2

Cardiocladius C1,F1,F5,F6,S2,S4

Cardiocladius albiplumus F6,S2

Cardiocladius obscurus S2,S3

Chaetocladius C1,F1,F6,S1

Chaetocladius dentiforceps F4

Chaetocladius piger F4

Cricotopus C1,F1,F5,F6,S1,S2,S4

Cricotopus albiforceps F1,S2

Cricotopus algarum S2

Cricotopus bicinctus F1,F4,F6,S2,S3

Cricotopus bicinctus gr. F1,F5

Cricotopus curtus S2

Cricotopus cylindraceus S2

Cricotopus festivellus F1,S2

Cricotopus flavocinctus S2

Cricotopus fugax S2

Taxonomic name Agency code

Cricotopus fuscatus S2

Cricotopus fuscus S2

Cricotopus infuscatus S2

Cricotopus intersectus S2,S3

Cricotopus junus S2

Cricotopus laetus S2

Cricotopus laricomalis S2

Cricotopus nostocicola S3

Cricotopus pirifer S2

Cricotopus reversus S2

Cricotopus reversus gr. S2,S3

Cricotopus slossonae S2

Cricotopus sylvestris S2

Cricotopus sylvestris gr. F1,S3

Cricotopus tibialis S2

Cricotopus tremulus S2

Cricotopus tremulus gr. F1,S2,S3

Cricotopus triannulatus S2

Cricotopus tricinctus S2

Cricotopus trifascia F1,S2

Cricotopus trifascia gr. F1,F5,S3

Cricotopus trifasciatus S2

Cricotopus vierriensis S2,S3

Cricotopus vierriensis gr. F1

Diplocladius C1,F1,F4,S2

Diplocladius cultriger S1,S2

Doithrix F4,S2

Eukiefferiella C1,F1,F5,F6,S1,S2,S4

Eukiefferiella brehmi F4

Eukiefferiella brehmi gr. F1,S3

Eukiefferiella brevicalcar F1,F4,S2

Eukiefferiella brevicalcar gr. F1,S3

Eukiefferiella claripennis F1,F4,S2

Eukiefferiella claripennis gr. F1,S3

Eukiefferiella devonica F4,F6,S2

Eukiefferiella devonica gr. F1,S3

Eukiefferiella gracei F4,S2

Eukiefferiella gracei gr. F1

Eukiefferiella pseudomontana F4,F6,S2

Eukiefferiella pseudomontana 

gr.

S3

Eukiefferiella rectangularis F4,S2

Eukiefferiella rectangularis gr. F1

Georthocladius F4

Halocladius S2

Heleniella F4

Heterotrissocladius C1,F1,F6,S2

Heterotrissocladius hirtapex F4

Heterotrissocladius marcidus S2

Taxonomic name Agency code
22 A Checklist of the Aquatic Invertebrates of the Delaware River Basin, 1990-2000

Heterotrissocladius marcidus gr. F1,S3

Hydrobaenus F4,S1,S2

Hydrobaenus johannseni S2

Krenosmittia F6,S2

Limnophyes F1,F4,F6,S3

Lopescladius F1,S2

Metriocnemus C1,F6,S2

Metriocnemus knabi S2

Nanocladius C1,F1,F4,F5,F6,S1,S2,

S3

Nanocladius (Plecopteracoluthus)C1,S3

Nanocladius (Plecopteracoluthus) downesiS3

Nanocladius crassicornus F1,S2

Nanocladius distinctus S2

Nanocladius minimus S2

Nanocladius parvulus F1

Nanocladius parvulus gr. F1

Nanocladius rectinervis F1,S2

Orthocladius C1,F1,F5,F6,S1,S2,S4

Orthocladius (Eudactylocladius) F1

Orthocladius (Euorthocladius) F1

Orthocladius (Euorthocladius) 

rivulorum

S2

Orthocladius (Symposiocladius) 

lignicola

F1,F6,S2,S3

Orthocladius annectens S2,S3

Orthocladius carlatus F1,S3

Orthocladius clarkei F4

Orthocladius curtiseta gr. F1

Orthocladius nr. dentifer S3

Orthocladius dorenus F1,F4,S2

Orthocladius lignicola S2

Orthocladius mallochi F4

Orthocladius nigritus gr. F1

Orthocladius obumbratus F1,F4,S2,S3

Orthocladius thienemanni S2

Parachaetocladius F1,F4,F6,S1,S2,S3,S4

Parachaetocladius hudsoni S2,S3

Paracladius F1

Paracricotopus F1,S2

Parakiefferiella F1,S1,S2,S4

Parakiefferiella coronata S2

Parakiefferiella sp. 1 S2

Paralimnophyes S3

Parametriocnemus C1,F1,F4,F5,F6,S1,S2,

S4

Parametriocnemus graminicola S2

Taxonomic name Agency code

Parametriocnemus lundbecki F1,S2,S3

Parametriocnemus stylatus S2

Paraphaenocladius F1,F4,F6,S2

Paraphaenocladius sp. 1 S2

Paratrichocladius F1,F4,S1,S2

Psectrocladius C1,F1,F5,F6,S2,S3

Psectrocladius barbimanus S2

Psectrocladius dilatatus gr. S3

Psectrocladius elatus S2

Psectrocladius flavus S2

Psectrocladius nigrus S2

Psectrocladius octomaculatus S2

Psectrocladius pilosus S2

Psectrocladius psilopterus S2

Psectrocladius simulans S2

Psectrocladius sordidellus S2

Psectrocladius vernalis S2

Psectrocladius sp. 1 S2

Psectrocladius sp. 4 S2

Pseudorthocladius F1,F4,S2

Pseudosmittia F4

Psilometriocnemus F4

Rheocricotopus F1,F5,F6,S1,S2,S4

Rheocricotopus eminellobus F4

Rheocricotopus pauciseta F1

Rheocricotopus robacki F1,S2,S3

Rheocricotopus tuberculatus F4,S2

Rheocricotopus sp. 1 S2

Rheocricotopus sp. 2 S2

Rheosmittia F1,F4,S2

Smittia C1,F1,F5,S1,S2

Stilocladius F1,F4

Stilocladius clinopecten S1

Symbiocladius S2

Synorthocladius C1,F1,F4,F6,S1,S4

Synorthocladius semivirens F5,S2

Synorthocladius nr. semivirens S3

Thienemannia S1

Thienemanniella C1,F1,F4,F5,F6,S1,S2

Thienemanniella fusca S3

Thienemanniella xena S2,S3

Thienemanniella sp. 2 S2

Tvetenia F1,F5,F6,S1,S2

Tvetenia bavarica F4,F6,S2

Tvetenia bavarica gr. F1,S3

Tvetenia discoloripes S2

Tvetenia discoloripes gr. F1

Tvetenia vitracies F6,S2,S3

Taxonomic name Agency code
 23

Unniella S2,S3

Unniella multivirga S2

Xylotopus S2

Xylotopus par F1,S2

Zalutschia S2

Zalutschia zalutschicola S2

Chironominae C2,C3,F1,F5,F6,S4

Chironomini C3,F1,F5,F6,S4

Chironomus C1,F1,F5,F6,S2,S3,S4

Chironomus decorus S2

Chironomus riparius S2

Chironomus tentans S2

Cladopelma F5,F6,S2

Cladopelma amachaerus S2

Cryptochironomus C1,F1,F5,F6,S1,S2,S4

Cryptochironomus argus S2

Cryptochironomus fulvus S2

Cryptochironomus fulvus gr. S3

Cryptochironomus sp. 1 S2

Cryptochironomus sp. 3 S2

Cryptotendipes F1,F5,S2

Cryptotendipes emorsus S2

Cryptotendipes pseudotener S2

Demicryptochironomus C3,F1,F6

Demicryptochironomus vulneratusS2

Demicryptochironomus sp. 3 S3

Dicrotendipes C1,F1,F5,F6,S1,S2,S4

Dicrotendipes fumidus F1,S2

Dicrotendipes modestus S2

Dicrotendipes neomodestus F1,S2,S3

Dicrotendipes nervosus S2

Einfeldia F1,S2

Einfeldia natchitocheae S2

Endochironomus C1,F5,S2

Endochironomus nigricans S2

Endochironomus subtendens S2,S3

Glyptotendipes C1,F5,S1,S2,S4

Glyptotendipes lobiferus S2,S3

Glyptotendipes senilis S2

Goeldichironomus C1

Harnischia S2

Harnischia curtilamellata S2

Harnischia sp. 2 S2

Hyporhygma S2

Kiefferulus C1,S2

Lauterborniella F1

Lauterborniella agrayloides F1,S2

Lauterborniella varipennis S2

Taxonomic name Agency code

Microtendipes C1,F1,F5,F6,S2,S4

Microtendipes caducus S2

Microtendipes pedellus F4,F6,S1,S2

Microtendipes pedellus gr. F1,S3

Microtendipes rydalensis F6

Microtendipes rydalensis gr. F1,S3

Microtendipes tarsalis S2

Nilothauma C1,F1,F6,S3,S4

Nilothauma babiyi S2

Pagastiella F1,F6

Pagastiella ostansa S2

Parachironomus C1,F5,S2

Parachironomus abortivus S2

Parachironomus frequens S2

Paracladopelma C1,F1,F5,F6

Paralauterborniella nigrohalteraleF1,F5,S2

Paratendipes F1,F4,F5,F6,S1,S2,S4

Paratendipes albimanus S2

Phaenopsectra C1,F1,F5,F6,S1,S2

Phaenopsectra dyari F1,S3

Phaenopsectra flavipes S2

Phaenopsectra obediens S2

Phaenopsectra obediens gr. F1

Phaenopsectra punctipes gr. F1

Phaenopsectra sp. 1 S2

Phaenopsectra sp. 2 S2

Polypedilum C1,F1,F5,F6,S1,S2,S4

Polypedilum angulum F1

Polypedilum aviceps F1,F4,S3

Polypedilum convictum gr. F1

Polypedilum digitifer S3

Polypedilum fallax F1,S2

Polypedilum flavum S2,S3

Polypedilum halterale S2

Polypedilum halterale gr. F1

Polypedilum illinoense F1,F4,S2,S3

Polypedilum laetum F1,S3

Polypedilum ontario S2

Polypedilum ophioides S2

Polypedilum scalaenum S2

Polypedilum scalaenum gr. F1,S3

Polypedilum tritum F1,F4,S2

Polypedilum sp. C F1

Robackia F1

Robackia demeijerei F6,S2

Robackia demeijerei gr. F1

Saetheria F1,S2,S3,S4

Saetheria tylus S1,S2

Taxonomic name Agency code
24 A Checklist of the Aquatic Invertebrates of the Delaware River Basin, 1990-2000

Sergentia S3

Stenochironomus F1,F4,F5,S1,S2,S3

Stictochironomus F1,F5,S2

Stictochironomus devinctus S2

Stictochironomus flavicingula F1

Stictochironomus marmoreus F1

Stictochironomus sp. 1 S2

Stictochironomus sp. 2 S2

Tribelos F1,F5,S2

Tribelos jucundus F1,F4,F6,S2

Xenochironomus F6,S2

Xenochironomus xenolabis F5,S2

Zavreliella marmorata S2

Pseudochironomini

Pseudochironomus F5,S1,S2,S3

Pseudochironomus fulviventris S2

Pseudochironomus prasinatus S2

Pseudochironomus richardsoni S2

 Tanytarsini C3,F1,F5,S4

Cladotanytarsus F1,F5,F6,S1,S2,S3,S4

Cladotanytarsus atridorsum S2

Cladotanytarsus dispersopilosus S2

Cladotanytarsus mancus S2

Cladotanytarsus nr. mancus S3

Cladotanytarsus mancus gr. F1

Cladotanytarsus van der wulpi F1,S2

Cladotanytarsus sp. 2 S2

Cladotanytarsus sp. 3 S2

Cladotanytarsus sp. 4 S3

Constempellina F1,F4

Constempellina brevicosta S2

Micropsectra C1,F1,F4,F5,F6,S1,S2,

S3,S4

Micropsectra curvicornis S2

Micropsectra nr. curvicornis S3

Micropsectra deflecta S2

Micropsectra nr. deflecta S3

Micropsectra dives S2,S3

Micropsectra junci S2

Micropsectra nigripila S2

Micropsectra polita S2,S3

Micropsectra sp. 7 S2

Micropsectra sp. E S2

Neozavrelia C1,F1

Paratanytarsus C1,F1,F5,S1,S2,S4

Paratanytarsus dimorphis S3

Paratanytarsus dissimilis S2

Rheotanytarsus C1,C3,F1,F4,F5,F6,S1,

S2,S4

Taxonomic name Agency code

Rheotanytarsus distinctissimus F1,F6,S2

Rheotanytarsus distinctissimus 

gr.

S3

Rheotanytarsus exiguus F6,S2

Rheotanytarsus exiguus gr. F1,S2,S3

Rheotanytarsus pellucidus S2

Stempellina C1,F1,F4,F5,F6,S2

Stempellina bausei gr. F1

Stempellina sp. 1 S3

Stempellinella F1,F4,F5,S1,S3

Sublettea F1,F6,S2

Sublettea coffmani F1,F5,S1,S2,S3

Tanytarsus C1,F1,F4,F5,F6,S1,S2,

S4

Tanytarsus confusus S2,S3

Tanytarsus curticornis gr. F1

Tanytarsus glabrescens F1,S2

Tanytarsus glabrescens gr. S2,S3

Tanytarsus guerlus S2

Tanytarsus guerlus gr. S2,S3

Tanytarsus varelus S2

Tanytarsus sp. 1 F1

Tanytarsus sp. 2 F1

Tanytarsus sp. 4 F1,S2

Tanytarsus sp. 5 F1,S2

Tanytarsus sp. 6 F1

Tanytarsus/Micropsectra F6

Zavrelia F6,S2,S4

Zavrelia pentatoma S2

Culicidae F5,S2

Aedes S2

Aedes fitchii S2

Anopheles F5,S2

Anopheles punctipennis S2

Culex S2

Dixidae C2

Dixa C1,C2,F1,F4,S2,S4

Pachyneuridae S1

Psychodidae S2,S3

Maruina S2

Pericoma S2,S4

Psychoda C2,F1,S1,S2

Psychoda alternata S2

Telmatoscopus C1,S2

Telmatoscopus albipunctatus S2

Ptychopteridae

Bittacomorpha S2

Bittacomorpha clavipes S2

Ptychoptera quadrifasciata S2

Taxonomic name Agency code
 25

Simuliidae C2,F1,F2,F3,F5,S1,S2,

S4

Cnephia S2

Cnephia dacotensis C3,S2

Greniera abdita S2

Prosimulium C1,C3,F1,S1,S2,S4

Prosimulium arvum C3,F4

Prosimulium fontanum C3

Prosimulium hirtipes S2

Prosimulium magnum C3,S2

Prosimulium rhizophorum C3

Simulium C1,C2,C3,F1,F3,F5,F6,

S1,S2,S3,S4

Simulium aureum S2,S3

Simulium decorum S2,S3

Simulium fibrinflatum F5,S3

Simulium gouldingi S2

Simulium impar C3

Simulium jenningsi S2,S3

Simulium parnassum C3

Simulium pictipes C3,S2,S3

Simulium tuberosum C3,S2,S3

Simulium venustum C3,S2,S3

Simulium venustum/verecundum complexF1

Simulium vittatum C3,F4,S2,S3

Stegopterna S4

Stegopterna mutata C3,F4, S2

Tanyderidae

Protoplasa F6

Thaumaleidae F6

Tipulidae C1,C2,F1,F2,F5,F6,S1,

S4

Tipulinae

Holorusia S2

Tipula C1,C2,C3,F1,F3,F4,F5,

F6,S1,S2,S3,S4

Tipula abdominalis S2

Tipula ignobilis S2

Limoniinae F1

Antocha C1,C2,C3,F1,F3,F4,F5,

F6,S1,S2,S3,S4

Cryptolabis C3

Dactylolabis S2

Dicranota C1,C2,C3,F1,F4,F6,S1,

S2,S3,S4

Elliptera S2

Erioptera F4

Erioptera chlorophylla S2

Taxonomic name Agency code

Gonomyia S2

Helius C1,S2

Hesperoconopa S2

Hexatoma C1,C2,C3,F1,F4,F5,F6,

S1,S2,S3,S4

Hexatoma fultonensis S2

Hexatoma megacera S2

Hexatoma spinosa S2

Limnophila C3,F1,F6,S2,S4

Limonia F1,F4,S1,S2,S4

Limonia rostrata S2

Lipsothrix C3,S4

Molophilus F4,S2,S4

Molophilus hirtapennis S1

Ormosia S1,S2,S4

Pedicia C2,C3,F1,F4,S2,S3,S4

Pilaria F1,F4,S2

Pilaria tenuipes S2

Pseudolimnophila C3,F1,F4,S1,S2,S4

Rhabdomastix C3

Rhipidia S1

BRACHYCERA-ORTHORRHAPHA

Athericidae C2,F2

Atherix C1,C2,C3,F1,F6,S2,S3,

S4

Atherix lantha F5

Atherix variegata S2

Chrysopilus F4

Dolichopodidae C3,S2,S3,S4

Argyra S2

Dolichopus S2

Hydrophorus S2

Empididae C1,C2,C3,F1,F2,F5,F6,

S1,S4

Chelifera C1,F1,F4,F6,S2,S4

Clinocera C1,C3,F1,F4,F6,S2,S4

Clinocera stagnalis S2

Hemerodromiinae F5

Hemerodromia C1,C2,C3,F1,F5,F6,S1,

S2,S3,S4

Hemerodromia praecatoria S2

Hemerodromia rogatoris S2

Oreogeton F1,F4,F6

Rhamphomyia C2

Wiedemannia F1,F6,S4

Stratiomyidae C1,C3,F4,S1

Nemotelus S2

Odontomyia S2

Taxonomic name Agency code
26 A Checklist of the Aquatic Invertebrates of the Delaware River Basin, 1990-2000

Stratiomys C1,S2

Stratiomys discalis S2

Tabanidae C2,F1,F2,S2,S4

Chrysops C2,C3,F1,F4,F5,S2,S3,

S4

Tabanus C1,C2,F4,F5,S2

BRACHYCERA-CYCLORRHAPHA

Ephydridae C1,C2,F6,S2

Ephydra S2

Discomyzinae

Psilopa S2

Ephydrinae

Brachydeutera S2

Brachydeutera argentata S2

Dimecoenia spinosa S2

Ephydra C2,S2

Scatella S2

Taxonomic name Agency code

Hydrelliinae

Hydrellia S2

Muscidae C1,C2,F1

Limnophora C1,S2

Lispe S2

Lispoides aequifrons S2

Phoridae

Diplonevra S2

Sciomyzidae

Dictya S2

Tetanocera S4

Syrphidae C1,F4

Taxonomic name Agency code
 27

References Cited

Adler, P., and Kim, K.C., 1986, The black flies of Pennsylvania 

(Simuliidae, Diptera) bionomics, taxonomy, and 

distribution: The Pennsylvania State University, College of 

Agriculture, Agricultural Experiment Station, University 

Park, Pa., Bull. 856, 88 p. 

Barbour, M.T., Gerrittsen, Jeroen, Snyder, B.D., and Stribling, 

J.B., 1999, Rapid bioassessment protocols for use in streams 

and wadeable rivers—Periphyton, benthic macroinvertebrates and fish (2nd ed.): Washington, D.C., U.S. Environmental Protection Agency, Office of Water, EPA 841-B-99-

002, variously paged. 

Bode, R.W., Novak, M.A., and Abele, L.E., 1993a, Biological 

stream assessment, Willowemoc Creek, 1993 survey: New 

York State Department of Environmental Conservation 

Technical Report, 40 p.

Bode, R.W., Novak, M.A., and Abele, L.E., 1993b, Biological 

stream assessment, Beaver Kill, 1993 survey: New York 

State Department of Environmental Conservation Technical 

Report, 34 p.

Bode, R.W., Novak, M.A., and Abele, L.E., 1995a, Biological 

stream assessment, Willowemoc Creek, 1994 survey: New 

York State Department of Environmental Conservation 

Technical Report, 34 p.

Bode, R.W., Novak, M.A., and Abele, L.E., 1995b, Biological 

stream assessment, Little Beaver Kill, 1994 survey: New 

York Department of Environmental Conservation Technical 

Report, 35 p.

Bode, R.W., Novak, M.A., and Abele, L.E., 1995c, Biological 

stream assessment, Vly Creek, 1995 survey: New York State 

Department of Environmental Conservation Technical 

Report, 25 p. 

Bode, R.W., Novak, M.A., Abele, L.E., Heitzman, D.L., and 

Smith, A.J., 2004, Thirty-year trends in water quality of 

rivers and streams in New York state based on macroinvertebrate data 1972-2002: New York State Department of Environmental Conservation Report, 384 p. 

Cairns, J., Jr., and Pratt, J.R., 1993, A history of biological monitoring using benthic macroinvertebrates in Rosenberg, 

D.M., and Resh, V.H., (eds.), Freshwater biomonitoring and 

benthic macroinvertebrates: New York, N.Y., Chapman and 

Hall, p. 10-27.

Donnelly, Thomas, 2004a, Distribution of North American 

Odonata, Part I—Aeshnidae, Petaluridae, Gomphidae, 

Cordulegastridae: Bulletin of American Odonatology, v. 7, 

no. 4, p. 61-90. 

Donnelly, Thomas, 2004b, Distribution of North American 

Odonata, Part II—Macromiidae, Corduliidae, and 

Libellulidae: Bulletin of American Odonatology, v. 8, no. 1, 

p. 1-32. 

Donnelly, Thomas, 2004c, Distribution of North American 

Odonata, Part III—Calopterygidae, Lestidae, Coenagrionidae, Protoneuridae, Platystictidae, with data sources and 

bibliography, Parts I–III: Bulletin of American Odonatology, v. 8, no. 2-3, p. 33-99. 

Epler, J.H., 1995, Identification manual for the larval Chironomidae (Diptera) of Florida: Orlando, Fla., Florida Department of Environmental Regulation, 302 p.

Epler, J.H., 1996, Identification manual for the water beetles of 

Florida (Coleoptera: Dryopidae, Dytiscidae, Elmidae, Gyrinidae, Haliplidae, Hydraenidae, Hydrophilidae, Noteridae, 

Psephenidae, Ptilodactylidae, Scirtidae): Tallahassee, Fla., 

Florida Department of Environmental Protection. 

Epler, J.H., 2001, Identification manual for the larval Chironomidae (Diptera) of North and South Carolina. jhepler@concentric.net

Fischer, J.M., Riva-Murray, Karen, Hickman, R.E., Chichester, 

D.C., Brightbill, R.A., Romanok, K.M., and Bilger, M.D., 

2004, Water quality in the Delaware River Basin, Pennsylvania, New Jersey, New York, and Delaware, 1998-2001: U.S. 

Geological Survey Circular 1227, 38 p. 

Hynes, H.B.N.,1960, The biology of polluted waters: 

Liverpool, England, Liverpool University Press, 199 p.

Hynes, H.B.N., 1970, The biology of running waters: Toronto, 

Canada, University of Toronto Press, 555 p.

Kathman, R.D., and Brinkhurst, R.O., 1998, Guide to the freshwater oligochaetes of North America: College Grove, Tenn., 

Aquatic Resources Center, 264 p. 

Kennen, J.G., 1999, Relation of macroinvertebrate community 

impairment to catchment characteristics in New Jersey 

streams: Journal of the American Water Resources Association, v. 35, no. 4, p. 939-955. 

Klemm, D.J., and Lazorchak, J.M., eds., 1994, Environmental 

and assessment program—Surface water and Region 3 

regional environmental monitoring and assessment program 

1994—Pilot field operation and methods manual for 

streams: Cincinnati, Ohio, U.S. Environmental Protection 

Agency, Environmental Monitoring Systems Lab, Office of 

Research and Development, EPA/620/R-94/004. 

Kolkwitz, R., and Marsson, M., 1908, Okologie der pflanzlichen Saprobien. Berichte der Deutschen Botanischien 

Gesellschaft 26A: p. 505-519.

Kolkwitz, R., and Marsson, M., 1909. Okologie der tierischen 

Saprobien. Beitrage zur Lehre von des bioloischen Gewasserbeurteilung. Internationale Revue der Gesamten Hydrobiologie und Hydrographie 2, p. 126-152.

Kosztarab, M., and Schaefer, C.W., eds., 1990, Systematics of 

North American insects and arachnids—Status and needs: 

Blacksburg, Va., Information Series 90-1, VA Agric. Exp. 

Sta., VA Polytechnic Institute and State University. 

Kurtenbach, J., 1991, A method for rapid bioassessment of 

streams in New Jersey using benthic macroinvertebrates: 

Bulletin of the North American Benthological Society, v. 8, 

no. 1, p. 129. 

Lawrence, G.B., Burns, D.A., Murdoch, P.S., Baldigo, B., and 

Baevsky, Y.H., 1994, Workplan of the Neversink watershed 

study in the Catskill Mountains of southeastern New 

York: U.S. Geological Survey Open-File Report 94-368, 

42 p. 
28 A Checklist of the Aquatic Invertebrates of the Delaware River Basin, 1990-2000

Lellis, W.A., 2001, Freshwater mussel survey of the Upper Delaware Scenic Recreational River—Qualitative survey 2000, 

Report to the National Park Service, February 20, 

2001: Wellsboro, Pa., U.S. Geological Survey, 57 p.

Masteller, E.C., Lee, S., Simons, J., and Magee, D., 2004, 

Searchable database of the caddisflies and stoneflies of 

Pennsylvania: Pennsylvania Sea Grant, accessed Jan. 31, 

2004, at http://www.pserie.psu.edu/seagrant/extension/

other/database.html

Masteller, E.C., and Flint, O.S., Jr., 1992, The Trichoptera (caddisflies) of Pennsylvania—An annotated checklist: Journal 

of Pennsylvania Academy of Science 66(2): 68-78.

McCafferty, W.P., 2000, Higher classification of the 

Ephemeroptera: Mayfly Central, Purdue University, Department of Entomology, accessed Jan. 31, 2004, at 

http://www.entm.purdue.edu/entomology/mayfly/.

Merritt, R.W., and Cummins, K.W., 1996, An introduction to 

the aquatic insects of North America (3d ed.): Dubuque, 

Iowa, Kendall/Hunt Publishing Company, 862 p.

Monroe County Planning Commission, 1990, Water quality 

study, 1990: Stroudsburg, Pa., Monroe County Planning 

Commission Report, 50 p.

Monroe County Planning Commission, 1991, Water quality 

study, 1991: Stroudsburg, Pa., Monroe County Planning 

Commission Report, 47 p.

Monroe County Planning Commission. 1992. Water Quality 

Study, 1992: Stroudsburg, Pa., Monroe County Planning 

Commission Report, 54 p.

Monroe County Planning Commission, 1993, Water Quality 

Study, 1993: Stroudsburg, Pa., Monroe County Planning 

Commission Report, 84 p.

Monroe County Planning Commission, 1994, Water Quality 

Study, 1994: Stroudsburg, Pa., Monroe County Planning 

Commission Report, 93 p.

Monroe County Planning Commission, 1995, Water Quality 

Study, 1995: Stroudsburg, Pa., Monroe County Planning 

Commission Report, 95 p.

Monroe County Planning Commission, 1996, Water Quality 

Study, 1996: Stroudsburg, Pa., Monroe County Planning 

Commission Report, 101 p.

Monroe County Planning Commission, 1997, Water Quality 

Study, 1997: Stroudsburg, Pa., Monroe County Planning 

Commission Report, 100 p.

Monroe County Planning Commission, 1998, Water Quality 

Study, 1998: Stroudsburg, Pa., Monroe County Planning 

Commission Report, 104 p.

Monroe County Planning Commission, 1999, Water Quality 

Study, 1999: Stroudsburg, Pa., Monroe County Planning 

Commission Report, 75 p.

Monroe County Planning Commission, 2000, Water Quality 

Study, 2000: Stroudsburg, Pa., Monroe County Planning 

Commission Report, 82 p. 

New Jersey Department of Environmental Protection, 1994a, 

Ambient biomonitoring network—Delaware River drainage 

basin, 1992 benthic macroinvertebrate data: New Jersey 

Department of Environmental Protection Report, 206 p.

New Jersey Department of Environmental Protection, 1994b, 

The establishment of ecoregion biological reference sites for 

New Jersey streams—Incorporating habitat quality and 

benthic macroinvertebrate communities, 1989-1993 monitoring data: New Jersey Department of Environmental Protection Report, 114 p.

New Jersey Department of Environmental Protection. 1996. 

Ambient biomonitoring network—Lower Delaware River 

drainage basin, 1995-96 benthic macroinvertebrate data: 

New Jersey Department of Environmental Protection Report, 

133 p.

New Jersey Department of Environmental Protection, 1999a, 

Ambient biomonitoring network—Watershed management 

areas 1, 2, and 11—Upper Delaware region, 1998 benthic 

macroinvertebrate data: New Jersey Department of Environmental Protection Report, 171 p.

New Jersey Department of Environmental Protection, 1999b, 

Ambient biomonitoring network—Watershed management 

areas 19 and 20, Delaware Region, upper tidal portion, 1998 

benthic macroinvertebrate data: New Jersey Department of 

Environmental Protection Report, 113 p.

New York State Department of Environmental Conservation, 

1996, The Delaware River Drainage Basin Biennial Report, 

1993-94: New York State Department of Environmental 

Conservation Technical Report, 102 p. + appendixes. 

Omernick, J.M., 1987, Ecoregions of the conterminous United 

States: Annals of the Association of American Geographers 

v. 77, no. 1, p. 118-125.

Passmore, M., and Green, J., 1997, The development of a 

stream reference condition in Monroe County, Pennsylvania: 

U.S. Environmental Protection Agency, Ecological Assessment and Protection Division Report, 61 p.

Pike County Conservation District, 1991, Annual Report, surface water quality monitoring program: Hawley, Pa., Pike 

County Conservation District Report, 81 p.

Pike County Conservation District, 1992, Annual Report, surface water quality monitoring program: Hawley, Pa., Pike 

County Conservation District Report, 110 p.

Pike County Conservation District, 1993, Annual Report, surface water quality monitoring program: Hawley, Pa., Pike 

County Conservation District Report, 104 p.

Pike County Conservation District, 1994, Annual Report, surface water quality monitoring program: Hawley, Pa., Pike 

County Conservation District Report., 117 p.

Pike County Conservation District, 1995, Annual Report, surface water quality monitoring program: Hawley, Pa., Pike 

County Conservation District Report, 142 p.

Pike County Conservation District, 1996, Annual Report, surface water quality monitoring program: Hawley, Pa., Pike 

County Conservation District Report, 138 p.

Pike County Conservation District, 1997, Annual Report, surface water quality monitoring program: Hawley, Pa., Pike 

County Conservation District Report.

Pike County Conservation District, 1998, Annual Report, surface water quality monitoring program: Hawley, Pa., Pike 

County Conservation District Report, 136 p. 
 29

Pike County Conservation District, 1999, Environmental quality of Pike County streams using bioassessment 

techniques: Hawley, Pa., Pike County Conservation District 

Report, 55 p. 

Pike County Conservation District, 2000, Environmental quality of Pike County streams using bioassessment 

techniques: Hawley, Pa., Pike County Conservation District 

Report, 56 p. 

Plafkin, J.L., Barbour, M.T., Porter, K.D., Gross, S.K., and 

Hughes, R.M., 1989, Rapid bioassessment protocols for use 

in streams and rivers—Benthic macroinvertebrates and 

fish: Washington, D.C., U.S. Environmental Protection 

Agency, Office of Water, EPA/444/4-89-001, variously 

paged. 

Rawlins, J.E., and Bier, C.W., 1998, Invertebrates—Review of 

status in Pennsylvania, in Hassinger, J.D., Hill, R.J., Storm, 

G.L., and Yahner, R.H., eds., Inventory and monitoring of 

biotic resources in Pennsylvania—Current ecological and 

landscape topic, Vol. 1., Pennsylvania Biological Survey: 

University Park, Pa., The Pennsylvania State University, 

Center for Biodiversity Research, 220 p.

Reif, A.G., 1999, Physical, chemical, and biological data for 

selected streams in Chester County, Pennsylvania, 1981-

94: U.S. Geological Survey Open-File Report 99-216, 

607 p.

Reif, A.G., 2000, Physical, chemical, and biological data for 

selected streams in Chester County, Pennsylvania, 1995-97: 

U.S. Geological Survey Open-File Report 00-238, 147 p. 

Shertzer, R.H., and Schreffler, T.L., 1996, Pennsylvania’s surface water quality monitoring network: Harrisburg, Pa., 

Pennsylvania Department of Environmental Protection, 

Report No. 3600-BK-DEP0636. 

Smith, D.G., 2001, Pennak’s freshwater invertebrates of the 

United States—Porifera to Crustacea: New York, John 

Wiley and Sons, Inc., 638 p.

Smith, I.M., and Cook, D.R., 1991, Water mites, in Thorp, J.H., 

and A.P. Covich, A.P., eds., Ecology and classification of 

North American freshwater invertebrates: San Diego, Calif., 

Academic Press, Inc., p. 523-592. 

Snyder, C., Young, J., Smith, D., Ross, R., and Bennett, R., 

1999, Influence of eastern hemlock on aquatic biodiversity in 

the Delaware Water Gap National Recreation Area: 

Kearneysville, W.Va., U.S. Geological Survey, Leetown Science Center. 

Stewart, K.W., and Stark, B.P., 2002, Nymphs of North American Stonefly Genera (Plecoptera) (2d ed.): Columbus, 

Ohio, The Caddis Press, 510 p. 

Strayer, D.L., and Jirka, K.J., 1997, The pearly mussels of New 

York State: New York State Museum Memoir 26: 1-113. 

Thorp, J.H., and Covich, A.P., 1991, Ecology and classification 

of North American freshwater invertebrates: San Diego, 

Calif., Academic Press, Inc., 911 p.

Thorp, J.H., and Covich, A.P., 2001, Ecology and classification 

of North American freshwater invertebrates (2d ed.): San 

Diego, Calif., Academic Press, Inc., 1056 p. 

U.S. Environmental Protection Agency, 1994, Guidance on 

implementation of biological criteria—Draft: Washington, 

D.C., U.S. Environmental Protection Agency, Office of Science and Technology.

U.S. Environmental Protection Agency, 1999, Level III Ecoregions of the Conterminous United States (revision of Omernick 1987): Corvallis, Oreg., U.S. Environmental Protection 

Agency, National Health and Environmental Effects Laboratory, Western Ecology Division. 

U.S. Environmental Protection Agency, 2000, Environmental 

monitoring and Assessment Program EMAP: accessed 

November 2001 at http://www.epa.gov/emap

Vogelmann, J.E., Sohl, T., and Howard, S.M., 1998, Regional 

characterization of land cover using multiple sources of data: 

Photogrammetric Engineering and Remote Sensing, v. 64, 

no. 1, p. 45-57.

Wetzel, M.J., 2001, The aquatic Annelida occurring in or adjacent to the Great Smokey Mountains National Park—An 

annotated checklist of species: Champaign, Ill., Illinois Natural History Survey, Center for Biodiversity. 

Wiggins, G.B., 1995, Larvae of the North American caddisfly 

genera (Trichoptera) (2d ed.): Toronto, Canada, University 

of Toronto Press, 457 p.

Wilson, E.O., 1984, Biophila: Cambridge, Mass., Harvard 

University Press, 111 p. 

Wilson, E.O., ed., 1988, Biodiversity: Washington, D.C., 

National Academy Press, 621 p. 

Wilson, E.O., 2000, A global biodiversity map: Science, 

v. 289, p. 2279.

Young, C., and Gelhaus, J., 2000, Crane flies of Pennsylvania—Preliminary checklist and database development with 

emphasis on aquatic species: Report to the Wild Resources 

Conservation Fund and the Pennsylvania Fish and Boat 

Commission, 256 p.