United States Department of Agriculture

Forest 

Service

International Institute

of Tropical Forestry

General Technical

Report IITF-GTR-45 January 

2015

Chemical and Physical Analyses 

of Selected Plants and Soils From 

Puerto Rico (1981–2000)
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Compilers 

Mary Jeane Sánchez and Edwin Lopez are chemists, and Ariel E. Lugo is an 

ecologist, U.S. Department of Agriculture, Forest Service, International Institute 

of Tropical Forestry, Jardín Botánico Sur, 1201, Calle Ceiba, Rio Piedras, PR 

00926-1119.
Chemical and Physical Analyses of 

Selected Plants and Soils From Puerto 

Rico (1981–2000)

Mary Jeane Sánchez, Edwin Lopez, and Ariel E. Lugo 

Compilers

U.S. Department of Agriculture

Forest Service

International Institute of Tropical Forestry

Jardín Botánico Sur

1201 Calle Ceiba

Rio Piedras, PR 00926-119

General Technical Report IITF-GTR-45

January 2015
ii

Abstract

Sánchez, Mary Jeane; Lopez, Edwin; Lugo, Ariel E., comps. 2015. Chemical 

and physical analyses of selected plants and soils of Puerto Rico (1981–2000). 

Gen. Tech. Rep. GTR-IITF-45. Rio Piedras, PR: U.S. Department of Agriculture, 

Forest Service, International Institute of Tropical Forestry. 85 p.

This report contains the results of analyses conducted at the chemistry laboratory 

of the International Institute of Tropical Forestry in Puerto Rico from 1981 to 2000. 

The data set includes 109,177 plant analyses and 70,729 soil analyses. We report 

vegetation chemical data by plant part, species, life zone, soil order, geology, or parent material. Soil data are reported by depth, soil order, geology or parent material, 

and life zone. These data are summarized to facilitate interpretation and highlight 

the mean and range of values obtained. 

Keywords: Plant, soil, chemical, element, tropical.

Acknowledgments

We thank Ernesto Medina, Michael Keller, Jorge Frangi, Elvira Cuevas, and Fred 

Scatena for their contributions to quality assurance and quality control in the 

laboratory. Carmen Marrero, Maribelis Santiago, and Luis Iglesias contributed to 

the processing and analysis of samples. Institute staff support both the functioning 

and maintaining of the infrastructure and laboratory facilities. All the work is done 

in collaboration with the University of Puerto Rico.
iii

Contents

1 Introduction

1 Description of the Contents

10 References

11 Appendix 1: Data Sources

35 Appendix 2: List of Species

43 Appendix 3: Elemental Composition by Species in Live Vegetation

47 Appendix 4: Elemental Composition by Species in Loose Litter

51 Appendix 5: Elemental Composition by Species in Litter Fall

55 Appendix 6: Elemental Composition by Soil Order

61 Appendix 7: Elemental Composition of Species From Contrasting

 Environments in Puerto Rico

 Ernesto Medina
iv
1

Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

Introduction

The International Institute of Tropical Forestry’s chemistry laboratory, established 

in 1981 in Puerto Rico, has the capacity to analyze total and available elements 

in soil and plant parts, including aluminum, calcium, carbon, chromium, iron, 

manganese, magnesium, nickel, nitrogen, phosphorus, potassium, and sulfur as 

well as ash, loss on ignition, and organic matter. In 1981 the laboratory conducted 

93 analyses for two investigators; in 1990, it analyzed samples in 10 projects for 

8 investigators for a total of 14,840 analyses. By 2000, the laboratory analyzed 

samples in 21 projects for 16 investigators, for a total of 47, 614 analyses.

This document contains the results of most analyses conducted at the laboratory 

from 1981 to 2000. Sanchez et al. (1997) reported results up to 1990, and here we 

update that report to the year 2000. Our objective continues to be making available 

in one place all the chemical and physical data that we develop in our laboratory. 

With the increased importance of the field of ecological stoichiometry (Sterner and 

Elser 2002), the relevance of data compendia such as this one increases. Most of 

these data are for vegetation (mostly forests) and soils of Puerto Rico. Not included 

here (but available in the laboratory) are data for other parts of the world, particularly from Argentina. The chemical composition of plant parts or soil is summarized by plant species, soil order, and geology, or parent material. 

Description of the Contents

The elemental concentration is described in terms of mean, standard deviation, and 

median, maximum, and minimum values. Codes used for the life zones and types 

of plant parts (tissues) are defined in tables 1 and 2, respectively. The results of 

the analyses are not discussed in this report. However, we summarize the data to 

facilitate interpretation and to synthesize the range of values obtained for tropical 

ecosystems. The range of values for elements in plants and soils are summarized in 

table 3 (plants) and tables 4 to 6 (soils). Methods of analysis are described in table 7. 

Table 8 contains the bulk of the vegetation data and table 9 contains the bulk of the 

soil data, both based on individual studies. Data sources for table 8 and 9 are listed 

in appendix 1, Tables 8 and 9 are published on the CD.

Study species and identification numbers are listed in appendix 2. Appendices 

3 to 5 consist of graphs that show elemental concentration in various plant parts 

by species. Appendix 6 contains graphs with data about soil samples. The figures 

show data on mean concentration of elements and other constituents in different 

types of soil. Data on other characteristics, such as bulk density and effective cation 

exchange capacity, are also shown. Appendix 7 contains an analysis of leaf chemistry data from different environments.
2

International Institute of Tropical Forestry II

Table 1—Ecological life zone codes

Code Ecological region

Lmrf Subtropical lower montane rain forest

Lmwf Subtropical lower montane wet forest

Sdf Subtropical dry forest

Smf Subtropical moist forest

Srf Subtropical rain forest

Swf Subtropical wet forest

Table 2—Codes for the identification of plant part samples (continued)

I. Soil

II. Vegetation

1. Trees A. Leaves i. Canopy a. Young b. Mature c. Other

ii. Shade a. Young b. Mature

iii. Other

B. Branches

C. Main bole, stem, trunk i. Bark

ii. Heartwood

iii. Sapwood

D. Twigs

E. Roots

F. Flowers, fruits

G. Miscellaneous

H. Leaves, stems

2. Shrubs A. Leaves

B. Branches

C. Main bole, stem, trunk i. Bark

D. Twigs

E. Roots

F. Flowers, fruits

G. Miscellaneous

H. Leaves, stems

The next and third version of this report will include analyses from 1981 to 

2010, and we welcome comments for improving the presentation of data. We ask for 

acknowledgment of any use of this information and for copies of reports, articles, or 

documents resulting from the use of this information.
3

Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

Table 2—Codes for the identification of plant part samples (continued)

3. Herbs A. Leaves i. Canopy

ii. Shade

B. Branches

C. Main bole, stem, trunk i. Bark

D. Twigs

E. Roots

F. Flowers, fruits

G. Miscellaneous

H. Leaves, stems

4. Climber A. Leaves

B. Branches

C. Main bole, stem, trunk i. Bark

D. Twigs

E. Roots

F. Flowers, fruits

5. Moss

6. Other A. Leaves

B. Branches

C. Main bole, stem, trunk

D. Twigs

E. Roots

F. Flowers, fruits

7. Coarse woody debris A. Bark

B. Heartwood

C. Sapwood

8. Loose litter A. Leaves

B. Wood

C. Flowers, fruits

D. Miscellaneous

9. Litter fall A. Leaves

B. Wood

C. Flowers, fruits

D. Miscellaneous
4

International Insitute ofTroicalForestry IITF-G

Table 3—Range of chemical concentration data for plants partsPlant part Aluminum Ash Calcium Carbon Cobalt Copper Chromium Iron Magnesium Manganese Nickel Nitrogen Phosphorus Potassium Sodium Sulfur Zinc

(mg/g) (%) (mg/g) (%) (μg/g) (μg/g) (μg/g) (mg/g) (mg/g) (mg/g) (μg/g) (%) (mg/g) (mg/g) (mg/g) (%) (μg/g) 

Bark 3.52-0.01 15.86-0.14 63.48-0.29 54.80-40.00 1.40-0.65 0.24-0.08 0.35-0.17 2.66-0.98

Bole 13.61-0.01 11.48-0.44 41.60-0.47 54.16-48.12 1.48-0.01 7.00-0.06 0.37-0.01 3.32-0.22 2.00-0.02 31.00-0.42 12.49-10.43 0.54-0.01

Branches 6.98-0.01 4.53 22.26-0.91 49-41 0.30 0.28-0.01 5.53-0.26 1.36-0.01 1.53-0.03 1.59-0.01 16.20-0.27 0.28-0.01

Climber 8.85-0.01 14.80-0.43 2.55-0.05 69.75 7.03-0.37 4.47-0.21 89.46-1.12 0.57-0.07

Coarse woody debris: 0.25-0.05 11.86--2.82 13.79-1.18 51.20-43.96 1.17-0.33 0.31-0.08 0.18-0.08 2.14-0.20

Bark 4.35-0.09 13.50-6.72 1.40-0.07 2.38-0.36 0.79-0.05 0.47-0.05 3.06-0.58

Heartwood 0.14-0.01 14.01-1.75 0.15-0.01 1.17-0.25 0.13-0.02 0.31-0.08 0.22-0.04 2.23-0.23

Sapwood 0.75-0.02 25.37-2.60 1.18-0.02 1.01-0.22 0.97-0.03 0.49-0.07 3.47-0.57

Flowers and fruits 0.07-0.02 10.05-2.65 21.00-0.04 4.86-0.77 1.01-0.01 2.92-0.36 3.80-0.37 34.34-2.68

Heartwood 34.80-0.01 21.94-1.15 71.55-0.71 57.24-42.00 5.80-0.30 32.20-1.88 0.08-0.02 1.13-0.32 0.13-0.03 0.24-0.11 0.15-0.03 1.64-0.37

Leaves: 34.80-0.01 57.55-0.50 49.90-0.36 61-16 4.13-0.10 7.55-2.00 178.00-1.00 5.97-0.01 35.05-0.35 2.10-0.01 181.45-2.35 4.57-0.21 13.87-0.01 61.15-0.43 185.41-0.28 1.67-0.04 52.43-4.84

Canopy 0.03 19.97-0.53 80.02-32.04 52 0.04 1.42 0.04 4.57-0.51 0.84-0.21 10.49-0.80 0.72 0.13

Mature-canopy 0.17-0.12 15.58-10.16 3.84-2.93 1.80-0.48 5.61-0.21 23.99-0.80

Mature-shade 17.55-11.83 4.50-3.45 0.06-0.04 1.71-1.42 0.88-0.60 8.73-4.99

Leaves-other: 0.08-0.03 5.65-4.19 12.13-5.65 54-41 0.21-0.05 4.45-2.04 0.15-0.04 5.08-0.65 1.50-0.86 12.79-6.95 0.66-0.21 0.52-0.31

Shade 2.07-0.09 17.55-14.28 97.98-46.54 5.94-3.36 1.59-0.93 3.64-0.77 26.38-4.99

Leaves and stems 7.89-0.06 22.07-4.02 44.95-2.72 55.80-37.80 0.39-0.11 5.67-1.19 0.72-0.05 1.92-1.36 1.98-0.66 41.35-7.25

Miscellaneous 33.37-2.68 14.57-5.51 7.39-0.09 2.84-1.11 0.40-0.05 2.80-0.87 1.51-0.22 12.42-0.93 0.74-0.28

Moss 17.07-2.09 27.47-1.82 3.09-0.53 1.42-0.04 1.43-0.24 1.79-0.20 1.79-0.20

Other 13.73-0.01 16.46-2.10 9.77-1.21 54.82-39.85 8.25-064 1.65-0.78 1.15-0.22 20.68-2.25

Roots 12.54-6.70 17.62-0.09 17.71-0.44 0.69-0.01 2.07-0.14 2.00-0.11 15.75-1.56 48.24-11.52 0.78-0.04

Shrubs 2.7-0.01 31.82-0.14 41.92-0.21 0.70-0.52 0.22-0.14 5.90-3.00Trunk: 63.48-10.47 0.27-0.01 4.53-0.05 0.33-0.01 2.43-0.06 2.48-0.01 44.13-0.30

Bark 0.46-0.18 16.19-3.24 0.70-0.20 22.28-5.02

Sapwood 22.42-0.77 0.50-0.13 1.53-0.039 0.45-0.05 3.39-2.59 0.28-0.12 2.38-0.99

Twigs 4.33-0.21 1.18-0.02 8.53-4.44

Loose litter:

Flowers andfruits 9.46-0.33 9.59-0.71 18.75-1.97 55.23-43.62 4.57-1.11 7.12-0.77 0.77-0.06 2.36-0.44 2.18-0.25 4.45-0.99 2.27-1.32 0.40-0.13

Leaves 15.09-0.07 22.66-3.14 36.69-1.38 57.14-37.86 6.01-0.09 6.77-1.10 1.69-0.03 3.96-0.50 1.18-0.01 12.84-1.09 1.55-0.11

Miscellaneous 20.90-0.15 43.13-5.94 103.03-1.81 46.39-30.71 22.02-0.26 4.87-1.42 1.56-0.05 8.80-0.91 0.95-0.05 9.14-0.23 0.38-0.15

Wood 6.63-0.13 9.82-1.98 26.99-2.10 54.07-42.23 2.16-0.03 5.05-0.70 1.05-0.03 1.45-0.32 0.85-0.10 7.57-0.46 0.35-0.11

Litter fall:

Flowers and fruits 2.17-0.10 6.01-3.80 8.73-1.13 54.21-43.50 3.32-0.25 3.56-0.09 0.46-0.02 2.33-0.41 5.63-0.27 11.62-0.68 9.77-6.72 0.57-0.21

Leaves 4.97-0.35 16.46-4.80 38.71-0.28 57.55-41.20 6.26-0.13 6.13-0.37 1.22-0.11 1.96-0.48 0.82-0.15 8.76-0.68 23.93-0.19 0.59-0.15

Miscellaneous 9.49-0.34 21.90-5.49 15.87-4.43 50.40-36.00 11.59-0.27 4.74-1.25 0.56-0.17 2.44-0.58 1.32-0.31 8.48-2.28 0.46-0.21

Wood 2.79-0.13 10.43-3.42 25.47-2.52 52.23-44.13 0.55-0.08 3.09-0.81 0.80-0.09 1.14-0.26 0.40-0.07 6.25-0.81 0.37-0.16
5

Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

Table 4—Chemical and physical characteristics of alfisols, entisols, and histosols by soil 

order or parent material (continued)

Alfisols Entisols Histosols

Element Depth Value Depth Value Depth Value

cm cm cm

Aluminum (cmol/kg-1) 0-30 0.12-0.05 0-40 0.39-0.01

Aluminum (mg/g) 0-30 0.16-0.03 0-20 3.24-0.01

100-150 0.13-0.05

Aluminum-total (mg/g) 0-30 23.07-12.44

100-150 18.25-9.17

Bulk density (g/cc) 0-18 1.96-0.45 0-18 1.32-1.11

Carbon/nitrogen 0-15 747-21 0-40 307-1

Calcium (cmol/kg-1) 0-15 49.13-1.24 0-40 19.66-2.56

15-80 4.99-0.67

Calcium (mg/g) 0-80 9.83-0.13 0-30 7.53-1.72 0-30 3.93-0.34

Calcium-total (mg/g) 0-150 7.01-4.06

Carbon (%) 0-15 45.17-28.58 0-30 32.16-7.15 0-30 28.06-0.40

Clay (%) 0-30 26-3 0-30 27-5

Cobalt-total (mg/g) 

Copper-total (mg/g) 

Chromium-total (mg/g) 

ECEC (cmol/kg-1) 0-30 65.79-4.00 0-30 23.95-4.09

Iron (mg/g) 0-30 0.20-0.01 0-20 0.07-0.01 0-40 0.19-0.01

Iron-total (%) 0-150 2.21-1.13

LOI (%) 0-30 74.90-33.39 0-40 55.91-1.42

Magnesium (mg/g) 0-30 1.85-0.05 0-30 13.19-0.28 0-40 1.82-0.01

Magnesium (cmol/kg-1) 0-80 15.44-0.58 0-40 3.34-0.11

Magnesium-total (mg/g) 0-30 14.01-1.63

Manganese (mg/kg) 0-30 350-16 0-30 250-6 0-30 84-1

Manganese-total (mg/g) 0-150 0.17-0.05

Nickel-total (mg/g)

Nitrogen (%) 0-15 1.35-0.06 0-30 1.19-0.28 0-30 3.00-0.01

Organic matter (%) 0-30 20.96-0.35 0-30 3.54-0.60

pH (H2O) 0-30 8.86-4.73 0-30 8.26-3.76 0-30 8.26-4.38

pH (KCl) 0-30 7.26-4.69 0-20 4.50-3.66 0-30 7.83-4.08

Phosphorus (mg/kg) 0-30 39-2 0-30 572-20 0-30 16-2

 30-120
6

International Institute of Tropical Forestry II

Table 4—Chemical and physical characteristics of alfisols, entisols, and histosols by soil 

order or parent material (continued)

Alfisols Entisols Histosols

Element Depth Value Depth Value Depth Value

cm cm cm

Phosphorus-total (mg/g) 0-30 0.29-0.09

Potassium (cmol/kg-1) 0-80 0.38-0.06 0-40 0.57-0.03

Potassium (mg/kg) 0-30 292-80 0-30 4850-50 0-30 220-11

Potassium-total (mg/g) 0-30 4.06-2.82

Sand (%) 0-30 97-41 0-30 86-11

Silt (%) 0-30 35-1 0-30 58-9

Sodium (cmol/kg-1) 0-15 0.76-0.03 0-30 2.28-0.06

Sodium (mg/kg) 0-30 130-7 0-30 830-12

Sodium-total (mg/g) 

Sulfur (mg/g) 0-150 76.90-16.00 0-40 19.00-0.04

ECEC = effective cation exchange capacity. LOI = loss on ignition. KCI = potassium chloride.

Table 5—Chemical and physical characteristics of inceptisols, limestone, and mollisols by soil 

order or parent material (continued)

Inceptisols Limestone Mollisols

Element Depth Value Depth Value Depth Value

cm cm cm

Aluminum (cmol/kg-1) -50 16.12-0.15 0-24 0.24-0.04 -30 19.42-0.01

0-30 49.82-7.30 0-10 19.34-1.18

Bulk density (g/cc) 0-30 1.63-0.07 0-23 1.52-0.53 0-30 1.42-0.25

30-75 1.20-0.65

Carbon/nitrogen 0-30 120-12 0-30 28-11

Calcium (cmol/kg-1) 0-25 50.96-0.02 0-30 97.06-0.10

25-50 6.47-0.01

Aluminum (mg/g) 0-30 9.70-0.01 0-30 10.99-1.37 0-30 19.41-0.02

0-30 84.47-0.78 0-23 25.06-7.86 0-23 346.58-1.91

Aluminum-total (mg/g) 

0-30 19.68-0.32 0-30 46.07-1.41

Clay (%) 0-30 85-5 0-30 77-1 0-30 61-1

Cobalt-total (mg/g) 

Copper-total (mg/g) 

Chromium-total (mg/g) 

ECEC (cmol/kg-1) 0-25 62.61-1.52 0-30 59.85-9.00 0-30 117.38-8.00

Iron (mg/g) 0-30 3.370-0.004 0-30 0.39-0.01 0-30 1.590-0.002
7

Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

Table 5—Chemical and physical characteristics of inceptisols, limestone, and mollisols by soil 

order or parent material (continued)

Inceptisols Limestone Mollisols

Element Depth Value Depth Value Depth Value

cm cm cm

Iron-total (%) 0-30 8.02-1.18 0-10 5.88-0.15

LOI (%) 0-30 98.18-1.46 0-30 15.25-8.98 0-30 93.18-0.24

Magnesium (mg/g) 0-30 1.48-0.01 0-30 0.77-0.12 0-30 2.52-0.02

Magnesium (cmol/kg-1) 0-30 12.37-0.01 0-30 19.72-0.18

Magnesium-total (mg/g) 0-30 31.57-1.34 0-23 9.99-6.96 0-23 14.06-0.21

Manganese (mg/kg) 0-30 195-1 0-30 310-10 0-30 573-7

Manganese-total (mg/g) 0-30 1.71-0.34 0-10 1.14-0.04

Nickel-total (mg/g) 

Nitrogen (%) 0-30 0.76-0.03 0-23 0.38-0.22 0-30 3.14-0.12

Organic matter (%) 0-30 28.16-0.85 0-30 21.01-0.60 0-30 41.32-2.36

30-100 3.39-0.12 30-100 1.26-0.25

pH (H2O) 0-30 9.07-3.78 0-30 7.80-4.23 0-30 8.42-3.45

pH (KCl) 0-30 8.02-3.18 0-30 7.13-4.65 0-30 7.48-3.13

Phosphorus (mg/kg) 0-30 36-1 0-30 95-29 0-30 1010-8

Phosphorus total (mg/g) 0-30 7.22-0.09 0-30 1.12-0.13 0-23 1.42-0.03

30-100 0.40-0.06 30-100 0.29-0.09

Potassium (cmol/kg-1) 0-30 1.60-0.01 0-30 18.93-0.10

Potassium (mg/kg) 0-30 609-16 0-30 371-90 0-30 7194-36

Potassium-total (mg/g) 0-30 7.96-1.28 0-23 3.72-2.63 0-23 13.89-0.76

Sand (%) 0-30 89-1 0-30 91-3 0-30 80-7

Silt (%) 0-30 64-4 0-30 77-3 0-30 68-11

Sodium (cmol/kg-1) 0-25 1.34-0.08 0-30 5.22-0.01

Sodium (mg/kg) 0-25 310-5 0-30 210-21 0-30 1201-4

Sodium-total (mg/g) 0-43 0.47-0.33 0-23 0.99-0.77

Sulfur (mg/g) 0-30 16.97-0.05 0-30 4.45-0.08

ECEC = effective cation exchange capacity. LOI = loss on ignition. KC1 = potassium chloride.
8

International Institute of Tropical Forestry II

Table 6—Chemical and physical characteristics of oxisols, ultisols, and vertisols by soil 

order or parent material (continued)

Oxisols Ultisols Vertisols

Element Depth Value Depth Value Depth Value

cm cm cm

Aluminum (cmol/kg-1) 0-15 0.23-0.09 0-50 53.01-0.01 0-30 0.06

50-680 15.42-0.01

Aluminum (mg/g) 0-40 0.81-0.01

Aluminum-total (mg/g) 0-30 180.20-6.57

50-120 228.20-

98.90

Bulk density (g/cc) 0-18 1.25-0.57 0-30 1.76-0.08

Carbon/nitrogen 0-15 148-42 0-35 37-6

25-120 62-9

Calcium (cmol/kg-1) 0-15 36.75-13.05 0-40 80.75-0.01

100-680 14.23-0.37

Calcium (mg/g) 0-15 7.35-2.61 0-50 62.000-

0.003

0-30 4.16

Calcium-total (mg/g) 0-30 16.93-0.03

50-120 9.80-0.90

Carbon (%) 0-30 42.18-34.45 0-35 27.42-0.34

Clay (%) 0-18 27-25 0-30 89-1 0-30 40-11

Cobalt-total (mg/g) 0-60 0.67-0.38

Copper-total (mg/g) 0-60 0.04-0.01

Chromium-total (mg/g) 0-60 8.33-5.10

ECEC (cmol/kg-1) 0-15 40.95-14.66 0-30 90.03-0.16 0-30 38.44

Iron (mg/g) 0-15 0.03-0.01 0-40 13.08-0.002 0-30 0.01

Iron-total (%) 0-30 15.9-0.89

LOI (%) 0-30 116.41-2.27

Magnesium (mg/g) 0-15 0.47-0.11 0-30 7.68-0.01 0-30 1.71

Magnesium (cmol/kg-1) 0-15 3.95-0.95 0-30 33.9-0.12

Magnesium-total (mg/g) 0-30 35.03-0.34

Manganese (mg/kg) 0-15 41-13 0-30 840-1 0-30 40

Manganese-total (mg/g) 0-30 10.22-0.01

Nickel-total (mg/g) 0-60 3.35-1.51

Nitrogen (%) 0-15 0.82-0.28 0-30 3.00-0.02

Organic matter (%) 0-18 14.23-3.34 0-30 44.23-0.25 0-30 3.14

Organic matter (%) 

pH (H2O) 0-18 7.89-5.43 0-30 4.45-3.64 0-30 8.05

pH (KCl) 0-15 7.37-6.51 0-30 7.68-3.11 0-30 6.99

Phosphorus (mg/kg) 0-15 97-18 0-30 1793-1 0-30 63

Phosphorus (mg/kg) 30-120 230.00-0.44
9

Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

Table 6—Chemical and physical characteristics of oxisols, ultisols, and vertisols by soil 

order or parent material (continued)

Oxisols Ultisols Vertisols

Element Depth Value Depth Value Depth Value

cm cm cm

Phosphorus total (mg/g) 0-30 4.32-0.04

Phosphorus total (mg/g) 30-120 0.59-0.10

Potassium (cmol/kg-1) 0-15 0.81-0.19 0-30 1.65-0.02

Potassium (mg/kg) 0-15 307-74 0-30 1905-1 0-30 420

Potassium-total (mg/g) 0-30 13.00-0.79

Sand (%) 0-18 36 0-30 78-1 0-30 11

Silt (%) 0-18 37 0-30 63-6 0-30 38

Sodium (cmol/kg-1) 0-15 0.33-0.04 0-30 2.46-0.01

Sodium (mg/kg) 0-15 76-10 0-30 567-1 0-30 520

Sodium-total (mg/g) 

Sulfur (mg/g) 0-20 1.80-0.01

ECEC = effective cation exchange capacity. LOI = loss on ignition. KC1 = potassium chloride.

Table 7—Methods of chemical analysis and referencesa

Sample Analysis Reference to method

Soil

Available-aluminium, calcium, magnesium, sodium Hunter 1974

Available-iron, manganese, phosphorus, potassium Hunter 1974

Total-aluminium, calcium, iron, magnesium Huang and Shulte 1985

manganese, phosphorus, potassium, sodium

Loss on ignition Wilde et al. 1979

Nitrogen (macro Kjeldahl) from Jan 1980–1995 Page 1982b

Nitrogen (macro dry combustion) from Feb 1995–2000 Leco Corp. 2000b form No. 203-821-165

Organic matter Page 1982

pH (H2O-KCl) Southern Coop. Series 1983

Vegetation

Aluminium, calcium, cobalt, iron, magnesium, Huang and Shulte 1985 and

manganese, nickel, phosphorus, potassium, sodium Chapman and Pratt 1979a

Ash Wilde et al. 1979

Nitrogen (macro Kjeldahl) from Jan 1995–1980 Chapman and Pratt 1979b

Nitrogen (macro dry combustion) from Feb 1995–2000 Leco Corp. 2000a form No. 203-821-172

Wood density ASTM 1968 a Details on variations in methodology are available upon request.
10

International Institute of Tropical Forestry II

References

American Society of Testing and Materials [ASTM]. 1968. Book of ASTM 

Standard. Part 16. Philadelphia, PA. 

Chapman, H.D.; Pratt, P.F. 1979a. Métodos de análisis para suelos, plantas y 

aguas. Métodos de acenización. Capítulo. 2: 47–49.

Chapman, H.D.; Pratt, P.F. 1979b. Métodos de análisis para suelos, plantas y 

aguas Nitrógeno. Capítulo. 17: 102–103.

Huang, C.Y.; Schulte, E.E. 1985. Digestion of plant tissue for analysis by ICP 

emission spectroscopy. Communications in Soil Science and Plant Analysis.

16: 943–958.

Hunter, A.H. 1974. International soil fertility evaluation and improvement: In: 

Chapters A1—B3. Laboratory procedures. Raleigh, NC: Department of Soils 

Science, North Carolina State University.

LECO Corp. 2000a. Nitrogen and sulfur in plant tissue. Organic Application Note. 

CNS-2000 (form No. 203-821-172 ). Carbon, St. Joseph, MI. 

LECO Corp. 2000b. Total/organic carbon and nitrogen in soils. Organic 

Application Note. CNS-2000 (form No. 203-821-165 ). St. Joseph, MI.

Page, A.E. 1982a. Methods of soil analysis. Part 2. Nitrogen. Madison, WI: 

American Society of Agronomy: 621–622.

Page, A.E. 1982b. Methods of soil analysis. Part 2. Nitrogen. Madison, WI: 

American Society of Agronomy: 565–571.

Sánchez, M.J.; López, E.; Lugo, A.E. 1997. Chemical and physical analyses 

of selected plants and soils from Puerto Rico (1981–1990). Research Note 

IITF-RN-1. Río Piedras, PR: U.S. Department of Agriculture, Forest Service, 

International Institute of Tropical Forestry. 112 p.

Southern Cooperative Series Bulletin 289, 1983. Reference soil test methods for 

the southern region of the United States. Athens, GA: The University of Georgia 

College of Agriculture Experiment Stations: 1–3.

Sterner, R.W.; Elser, J.J. 2002. Ecological stoichiometry: the biology of elements 

from molecules to the biosphere. Princeton, NJ: Princeton University Press.

437 p.

Wilde, S.A.; Corey, J.G.; Iyer, J.G.; Voight, J.G 1979. Soil and plant analysis for 

tree culture. 5th ed. New Delhi, India: Oxford and IBH Publishing Co: 184–185.
11

Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

Appendix 1: Data Sources

Data sources are arranged by general area of study. The numbers correspond to 

sources cited in the tables on the characteristics of plant and soil samples. The 

sources are cross referenced to species in appendix 2.

Soils

1. Nutrient dynamics of a Puerto Rican subtropical dry forest. Journal of

Tropical Ecology. 2: 55–72, 1986

Investigators: Ariel E. Lugo and P. Murphy 

Location: Guánica Forest, Guánica, Puerto Rico

Date of collection: 1981–1983

2. Guánica Study #2508

Investigator: Vicky Dunevitz (Michigan State University)

Location: Guánica Forest, Guánica, Puerto Rico

Date of collection: 1982

3. Guánica Study #2508

Investigator: Dan Nepstad (Michigan State University)

Location: Guánica Forest, Guánica, Puerto Rico

Date of collection: 1983

4. Estudio de la colonización en áreas de derrumbes en el Bosque

Experimental de Luquillo. In: Lugo, A.E., ed. Los Bosques de Puerto

Rico. U.S. Department of Agriculture, Department of Natural Resources; 

University of Puerto Rico, 110–122, 1983

Investigators: Carmen Varela, D. Ortiz, A. Berríos, and H.J. ÁlvarezScience Teacher Association Investigation

Location: Landslides along PR 191 in Luquillo Experimental Forest,

Puerto Rico

Date of collection: August 1981

5. Structure and dynamics in the Colorado Forest of the Luquillo Mountains

of Puerto Rico

Investigator: Peter Weaver (Michigan State University, Ph.D. dissertation,

1987)

Location: Luquillo Experimental Forest, Puerto Rico

Date of collection: December 1981–July 1982
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International Institute of Tropical Forestry II

6. Estudio comparativo de algunas propiedades químicas y físicas de suelos

de bosque bajo uso natural y plantación 

Investigator: Mary Jeane Sánchez (M.Sc. thesis University of Puerto Rico,

Mayagüez Campus)

Location: Luquillo Experimental Forest, Puerto Rico

Date of collection: 1985–1986

7. Landslide disturbance and forest regeneration in the Upper Luquillo

Mountains of Puerto Rico. Journal of Ecology. 78: 814–832, 1990

Investigator: Manuel Guariguata

Location: Luquillo Experimental Forest, Puerto Rico

Date of collection: June 1988

8. Growth studies of plantations of Pinus caribaea var. hondurensis in

Puerto Rico

Investigator: Mohammed Zakir Hussain (Yale University, Ph.D.

dissertation, 1987)

Location: Pine plantations, various sites, Puerto Rico

Date of collection: 1984–1985

9. Studies of ecological and geological factors controlling the pattern of

Tabonuco forests in the Luquillo Experimental Forest, Puerto Rico

Investigator: Khadga Basnet (State University of New Jersey, Ph.D.

dissertation, 1990)

Location: Bisley, Luquillo Experimental Forest, Puerto Rico

Date of collection: 1988

10. Structure, succession, and soil chemistry of palm forest in the Luquillo

Experimental Forest. Tropical forest: Management and Ecology:

142–177, 1995

Investigators: Ariel E. Lugo, A. Bokkestijn, and F.N. Scatena 

Location: Palm Slope Forest, El Verde (PS-2) and Juan Diego (PS-1):

Caribbean National Forest, Puerto Rico

Date of collection: June 1986

11. Land use and organic carbon content of some subtropical soils. Plant and 

Soil. 96: 185–196, 1986

Investigators: Ariel E. Lugo, M.J. Sánchez, and S. Brown

Location: Various sites, Puerto Rico

Date of collection: 1982–1984
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Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

12. Revegetation on landslides

Investigator: Jean Lodge

Location: Large slide next to El Verde Field Station, Puerto Rico

Date of collection: December 1988

13. Performance of Anthocephalus chinensis in Puerto Rico. Canadian Journal

of Forest Research. 15: 577–585, 1985

Investigators: Ariel E. Lugo and J. Figueroa

Location: Various sites, Puerto Rico

Date of collection: May 1983

14. Forest plantings in Puerto Rico and Virgin Islands/small leaf mahogany

survey

Investigator: John Francis

Location: Various sites, Puerto Rico

Date of collection: 1990

15. Restoration and rehabilitation of tropical forests. 

Investigator: Ariel E. Lugo (Environmental Sciences Course, University

of Puerto Rico, Río Piedras Campus)

Location: Río Mameyes (river basin), Luquillo Experimental Forest,

Puerto Rico

Date of collection: September–October 1989

16. Fauna oligotológica de suelos de la serie Nipe en el Bosque

Estatal de Maricao

Investigator: Hendriekje Hubers (M.Sc. thesis University of Puerto Rico,

Mayagüez Campus, 1993)

Location: Maricao Forest, Puerto Rico

Date of collection: August 1990

17. Productivity, nutrient cycling, and succession in single- and mixed-species

plantations of C. equisetifolia, E. robusta, and L.leucocephala in Puerto

Rico. Forest Ecology and Management. 124: 45–77, 1999

Investigator: John Parrotta

Location: Toa Baja Experimental Farm, Toa Baja, Puerto Rico

Date of collection: 1993
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International Institute of Tropical Forestry II

18. Protection and planting to accelerate increases in biodiversity on

deforested sites

Investigators: John Francis and J. Parrotta

Location: Camp Santiago, Salinas, Puerto Rico

Date of collection: 2000

19. Productivity, nutrient cycling, and succession in single- and mixed-species

plantations of C. equisetifolia, E. robusta, and L. leucocephala in Puerto

Rico. Forest Ecology and Management. 124: 45–77, 1999

Investigator: John Parrotta

Location: Toa Baja Experimental Farm, Toa Baja, Puerto Rico

Date of collection: December 1996

20. Former land use and tree species affect nitrogen oxide emissions from

a tropical dry forest. Oecologia. 130: 297–308, 2002

Investigator: Heather Erickson

Location: Guánica Forest, Guánica, Puerto Rico

Date of collection: May–December 1995

21. Nitrogen oxide fluxes and nitrogen cycling during post agricultural

succession and forest fertilization in the humid tropics. Ecosystems.

4: 67–84, 2001

Investigators: Heather Erickson, M. Keller, and E.A. Davidson

Location: El Verde Station, Sabana and Mameyes, Luquillo Experimental

Forest, Río Grande, Puerto Rico

Date of collection: 1995–1996

22. Eco-physiological evaluation of drought stress in montane and lowland

forests as a result of an unusual dry spell in June 1994

Investigators: Ernesto Medina and E. Cuevas

Location: Bisley and East Peak, Puerto Rico

Date of collection: June 1994

23. Pterocarpus-Punta Viento

Investigators: Ernesto Medina, E. Cuevas, and A.E. Lugo

Location: Pterocarpus Forest, Punta Viento; Patillas, Puerto Rico

Date of collection: August 1994
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Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

24. Dwarf mangrove project

Investigators: Ernesto Medina, E. Cuevas, and A.E. Lugo

Location: Ceiba, Puerto Rico (entrance through Marina del Rey,

about 2 km in the direction of Roosevelt Roads swamp)

Date of collection: March 1999

25. Pterocarpus soil collection

Investigators: Ernesto Medina, E. Cuevas, and M. Aide

Location: Sabana Seca, Puerto Rico

Date of collection: February 1997

26. Dwarf mangrove project

Investigators: Ernesto Medina, E. Cuevas, and A.E. Lugo 

Location: Ceiba, Puerto Rico (entrance through Marina del Rey, about 2 

km in the direction of Roosevelt Roads swamp)

Date of collection: December 1995 

27. The effects of small-scale and catastrophic disturbance on carbon and

nutrient cycling in a lower subtropical wet forest in Puerto Rico

Investigator: Whendee Silver

Location: Bisley Watersheds, Carribbean National Forest, Luquillo

Experimental Forest, Puerto Rico

Date of collection: 1993–1999

28. Bisley Gaps soil phosphorus project

Investigator: Whendee Silver

Location: In a lower subtropical wet forest, Puerto Rico

Date of collection: September 1994

29. The role of biodiversity in biogeochemical cycling in a moist

subtropical forest in the Luquillo Experimental Forest, Puerto Rico 

Investigator: Lara Kueppers (M.Sc. thesis Standford University,

Standford, California, 1996)

Location: Cubuy, Luquillo Experimental Forest, Puerto Rico

Date of collection: 1995

30. Archaeological study

Investigator: Edna Lee Ortiz

Location: Ensenada Breñas and Cerro Gordo, Vega Alta, Puerto Rico

Date of collection: November–December 1990
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International Institute of Tropical Forestry II

31. Hydrological process in a humid tropical rain forest: a combined experi

mental and modelling approach

Investigators: Japp Schellekens (Ph.D. dissertation Vrije University,

Amsterdam, 2000)

Location: Bisley Watersheds, Luquillo Experimental Forest, Puerto Rico

Date of collection: June 1997

32. Biogeochemical study of a tropical estuarine lagoon: carbon/nitrogen

fluxes

Investigator: Mayra E. Suárez (M.Sc. thesis University of Puerto Rico,

Mayagüez Campus)

Location: Joyuda Lagoon, Cabo Rojo, Puerto Rico

Date of collection: 1994–1995

33. Distribution and population ecology and reproductive biology of Goetzea

elegans Wydler

Investigator: Eugenio Santiago Valentín

Location: Quebradillas, Isabela, Puerto Rico

Date of collection: July 1993

34. Controls on spatial and temporal variability in nitrous oxide fluxes

across a tropical rain forest ecosystem in the Luquillo Experimental

Forest, Puerto Rico 

Investigator: Claire P. McSwiney (Ph.D. dissertation, University of 

Hampshire May 1999)

Location: Río Icacos basin, Luquillo Experimental Forest, Puerto Rico

Date of collection: January 1998

35. Long-term ecological research watersheds flood-plain study

Investigator: Don Durfee

Location: Río Mameyes, flood plains, Luquillo Experimental Forest,

Puerto Rico

Date of collection: August 1991

36. Estudio de la vegetación en afloraciones de serpentinita en el Bosque

Estatal de Maricao, Puerto Rico 

Investigator: Geraldino Caminero (M.Sc. thesis, University of Puerto

Rico, Mayagüez Campus)

Location: Maricao Forest, Maricao, Puerto Rico

Date of collection: February 1991
17

Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

37. Forest plantings in Puerto Rico and Virgin Islands. Study #2510 Prunus 

occidentalis (Amendrón)

Investigator: Salvador Alemañy

Location: Road 191 Km 19, Caribbean National Forest, Puerto Rico

Date of collection: January 1992

38. Influence of microenvironment of growth and nutrient dynamics of three 

herbaceous species in the Bisley Experimental Watersheds of Luquillo

Experiment Forest, Puerto Rico

Investigator: Amy Arnold (M.Sc. thesis Tennessee University)

Location: Bisley, Luquillo Experimental Forest, Puerto Rico

Date of collection: December 1995–January 1996

39. Performance of Hibiscus elatus in Puerto Rico. Commonwealth Forestry

Review. 67: 327–338, 1988

Investigators: John Francis and P. Weaver

Location: Puerto Rico

Date of collection: 1987

40. The performance of Tectona grandis in Puerto Rico. Commonwealth

Forestry Review. 69: 313–323, 1990

Investigators: Peter Weaver and J. Francis

Location: Puerto Rico

Date of collection: 1987

41. Tabonuco Project (Dacryodes excelsa Vahl)

Investigator: Migdalia Álvarez 

Location: Bisley, Vereda Cristal, Puerto Rico

Date of collection: 1994

42. Ecological aspects of earthworms from Laguna Cartagena, Puerto Rico.

Caribbean Journal of Science. 32: 406–412, 1996

Investigators: Mónica Alfaro and S. Borges

Location: Lajas, Puerto Rico

Date of collection: September 1990

43. A comparison of ten provenances of Eucalyptus deglupta and

E. Europhylla in Puerto Rico: growth and survival over 15 years

Investigators: Ariel. E. Lugo and J. Francis

Location: Yabucoa and Río Abajo, Puerto Rico

Date of collection: May–June 1987
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International Institute of Tropical Forestry II

44. Swietenia macrophylla and Swietenia macrophylla x S. Mahogoni

development and growth: the nursery phase and establishment phase

in line planning in the Caribbean Nacional Forest, Puerto Rico

Investigator: Gerald P. Bauer (M.Sc. thesis New York State University,

Syracuse)

Location: Sabana, Caribbean National Forest, Puerto Rico

Date of collection: 1985

45. Nutrients and mass in litter and top soil of ten tropical tree plantations.

Arboretum Study. Plant and Soil. 125: 263–280, 1990

Investigators: Ariel E. Lugo, E. Cuevas, and M.J. Sánchez

Location: Arboretum, Cienaga Alta, Luquillo Experimental Forest,

Puerto Rico

Date of collection: October 1986

46. Nutrient return accumulation in litter of a secondary forest in the coffee

region of Puerto Rico. Acta Científica. 13(1-3): 43–74, 1999

Investigator: Ariel E. Lugo, C. Domínguez, A. Santos, and E. Torres

Location: Torrecilla secondary forest, Bo. Hincado, Barranquitas,

Puerto Rico

Date of collection: May 1991

47. Comparison of two secondary forests in the coffee zone of central Puerto

Rico. Acta Científica. 13(1-3): 27–41, 1999

Investigators: Nathanial Popper, C. Domínguez, A. Santos et al.

Location: Bo. Caguana, Utuado, Puerto Rico

Date of collection: 1996

48. Comparative study of nutrient dynamics of two forest types in Luquillo

Experimental Forest

Investigators: Fu Sheinglei and A.E. Lugo

Location: El Verde, Luquillo Experimental Forest, Puerto Rico

Date of collection: July 1995

49. Soil oxygen availability and biogeochemistry along rainfall and

topographic gradients in upland wet tropical forest soils

Biogeochemistry. 44: 301–328, 1999

Investigator: Whendee Silver, A.E. Lugo and M. Keller

Location: Bisley, Puerto Rico

Date of collection: July 1993
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Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

50. Contribution of wood to soil organic matter

Investigators: Jean Lodge and N. Clum

Location: El Verde, Puerto Rico

Date of collection: 1999

51. Estudio comparativo de análisis de nutrientes entre una zona tabacalera

y una zona cafetalera que actualmente son bosques secundarios en

Naranjito

Investigator: Ángel Santiago (Francisco Morales High School, Naranjito,

Puerto Rico)

Location: Naranjito, Puerto Rico

Date of collection: August 2000

52. Luquillo Experimental Forest tree-uproots

Investigators: Melanie Lenart and F. Scatena

Location: Luquillo Experimental Forest, Puerto Rico

Date of collection: December 2000

53. Difference in extrametrical hypha in three coffee cultivars in Puerto Rico

Investigator: Ligia Lebrón

Location: Adjuntas, Puerto Rico

Date of collection: June 1998

54. Cerro de La Rosa soil collection

Investigators: María Rivera and J.L. Ramírez

Location: Cerro de la Rosa, Toro Negro Forest, Orocovis, Puerto Rico

Date of collection: August 2000

55. Monito Gecko (Sphaeerodactylus micropithecus) census

Investigator: Miguel García

Location: Monito Island, Puerto Rico

Date of collection: 1994/1998

56. Soil factors in a coffee plantation and a natural pasture

Investigators: Randall Myster and L. Lebrón

Location: El Verde, Sabana, Puerto Rico

Date of collection: May 1999

57. Assessment for available elements, total elements and pH of soils samples

Investigator: Pedro Robles

Location: Reserva Nacional JBNERR, Aguirre, Salinas, Puerto Rico

Date of collection: December 1999
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International Institute of Tropical Forestry II

58. Long-term recovery of a Caribbean dry forest after abandonment of

different land uses in Guánica, Puerto Rico

Investigator: Sandra Molina (Ph.D. dissertation, University

of Puerto Rico, 1998)

Location: El Maniel, Guánica, Puerto Rico

Date of collection: 1992–1993

59. Nutrient concentration and heavy metal accumulations on different

soils over serpentine

Investigators: Julio Figueroa and M.J. Sánchez

Location: Maricao Forest, Maricao, Puerto Rico

Date of collection: October 1989

60. Spatial patterns in forest nutrient pools, growth and disturbance-soil

collection

Investigator: Skip Van Bloem

Location: Guánica Forest, Guánica, Puerto Rico

Date of collection: September 1998

61. Earth worm project

Investigator: Sonia Borges

Location: Luquillo Experimental Forest, Puerto Rico

Date of collection: 1991–1992

62. Ecological studies of the termites of Puerto Rico

Investigator: Susan Jones

Location: Guánica forest, Guánica, Puerto Rico

Date of collection: 1992–1993

63. Soil macrofauna and litter nutrients in three tropical tree plantations on

a disturbed site in Puerto Rico. Forest Ecology and Management. 170:

161–171, 2002

Investigators: Matthew Warren and X. Zou

Location: Toa Baja Plantation, Puerto Rico

Date of collection: February 1998

64. Assessment of productivity and biological nitrogen fixation in mixedspecies forest plantation.

Investigator: John Parrotta

Location: Toa Baja Plantation, Puerto Rico

Date of collection: 1989-1992-1993-1996
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Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

65. La importancia de los cristales de oxalato de calcio en Sida rhombifolia

(Malvaceae) como mecanismo de defensa y producto del metabolismo

secundario 

Investigator: Brenda Molano (M.Sc. thesis University of Puerto Rico,

Rio Piedras Campus)

Location: Carolina-Canovanas, Puerto Rico

Date of collection: September 1989

66. Tropic venture

Investigator: Will Scott

Location: Patillas, Bo. Muñoz Rivera, adjacent to the Carite

Commonwealth Forest, Puerto Rico

Date of collection: February 1991

Wood decomposition

67. Wood decomposition of Cyrilla racemiflora in a tropical montane forest.

Biotropica. 26(2): 124–140, 1994

Investigator: Juan Torres

Location: Luquillo Experimental Forest, Puerto Rico

Date of collection: 1989

68. Wood decay in PS-2 and TS-2

Investigator: Ariel E. Lugo

Location: El Verde and Juan Diego, Luquillo Experimental Forest,

Puerto Rico

Date of collection: 1982–1986

69. Litter and root dynamics in a tropical pine plantation and an equal age

secondary forest-decomposition of woody debris

Investigator: Sandra Brown

Location: Guzmán Pine Plantation, Luquillo Experimental Forest,

Puerto Rico

Date of collection: April–October 1988

70. Hydrology, sediment, and nutrient budgets in three humid watersheds

in Puerto Rico

Investigators: Sandra Brown and A.E. Lugo

Location: Bisley, Luquillo Experimental Forest, Puerto Rico

Date of collection: September 1991–February 1994
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International Institute of Tropical Forestry II

71. Decomposition of leaves from general plant species and the role of fungi

in Luquillo Experimental Forest, Puerto Rico

Investigators: Jean Lodge and M. Santana

Location: El Verde, Sabana and Bisley; Luquillo Experimental Forest,

Puerto Rico

Date of collection: February 2000

Mangroves

72. Jobanes mangrove

Investigators: José López and M. Álvarez

Location: Jobanes Mangrove, Guayama, Puerto Rico

Date of collection: 1985

73. Jobanes Study—litter fall collection

Investigators: Ariel E. Lugo and G. Cintrón

Location: Jobos Bay, Aguirre, Guayama, Puerto Rico

Date of collection: May–July 1986

74. Jobos Bay—fresh leaves

Investigator: Ariel E. Lugo

Location: Jobos Bay, Aguirre, Guayama, Puerto Rico

Date of collection: December 1986

75. Características morfológicas y químicas de las hojas del Rhizophora

mangle en un bosque de franja en Las Mareas, Puerto Rico.

Acta Científica. I: 20–24, 1987

Investigators: Honora Serrano and I. Monefeldt

Location: Santuario Estuarino Jobanes, Las Mareas; Puerto Rico

Date of collection: 1983

76. Pterocarpus officinalis—litter fall

Investigator: Luis Negrón (Humacao College of University of Puerto Rico)

Location: Río Antón Ruiz Swamp, Humacao; Puerto Rico

Date of collection: 1986

77. Soil salinity, sun exposure, and growth of Acrostichum aureum, the

mangrove fern. Botanical Gazette. 151(1): 41–49, 1990

Investigators: Ernesto Medina, E. Cuevas, M. Popp, and A.E. Lugo 

Location: Río Grande and Vega Baja, Puerto Rico

Date of collection: 1986
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Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

78. Pterocarpus—Punta Viento Study

Investigators: Ernesto Medina, E. Cuevas, and A.E. Lugo

Location: Pterocarpus Forest, Punta Viento; Patillas, Puerto Rico

Date of collection: August 1994

79. Dwarf Mangrove Project—nutrient use efficiency collection

Investigators: Ernesto Medina, E. Cuevas, and A.E. Lugo

Location: Ceiba, Puerto Rico (entrance through Marina del Rey,

about 2 km in the direction of Roosevelt Roads)

Date of collection: March 1999

80. Dwarf Mangrove Project—fresh plant tissue collection

Investigators: Ernesto Medina, E. Cuevas, and A.E. Lugo

Location: Ceiba, Puerto Rico (entrance through Marina del Rey,

about 2 km in the direction of Roosevelt Roads)

Date of collection: December 1995

81. Dwarf Mangrove Project—nutrient use efficiency leaves collection

Investigators: Ernesto Medina, E. Cuevas, and A.E. Lugo

Location: Ceiba, Puerto Rico (entrance through Marina del Rey,

about 2 km in the direction of Roosevelt Roads)

Date of collection: December 1995

82. Dwarf Mangrove Project—biomass collection

Investigators: Ernesto Medina, E. Cuevas, and A.E. Lugo

Location: Ceiba, Puerto Rico (entrance through Marina del Rey,

about 2 km in the direction of Roosevelt Roads)

Date of collection: March 1999

83. Salt accumulation by Pterocarpus officinalis along a salinity gradient

Investigators: Ernesto Medina, E. Cuevas, and M. Aide

Location: Sabana Seca, Puerto Rico

Date of collection: February 1997

84. Litter nutrients in a Pterocarpus officinalis forest

Investigators: Mitch Aide and F. Scatena

Location: Sabana Seca, Puerto Rico

Date of collection: July 1995–July 1997

85. Materia orgánica y nutrientes en la hojarasca de un manglar

de franja. Acta Científica. I: 25–29, 1987

Investigators: Carmen Varela and A. Berrios

Location: Jobos, Las Mareas, Puerto Rico

Date of collection: 1984
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International Institute of Tropical Forestry II

86. Comparison of the nutrient status of dwarf and riverine mangrove and

other tropical forest types 

Investigator: Alicia Miñana (M.Sc. thesis Princeton University,

New Jersey)

Location: Roosevelt Road, Ceiba, Puerto Rico

Date of collection: September 1983

87. Susceptibility of mangrove wood to the arboreal termite (Nasutitermes

costalis) in the Piñones State Forest

Investigator: Teresa Carasquillo

Location: Piñones Forest, Loiza, Puerto Rico

Date of collection: August–September 1993

Vegetation

88. Biomass and nutrient accumulation in ten-year-old bryophyte communities inside a flood plain in the Luquillo Experimental Forest, Puerto

Rico. Biotropica. 24: 106–112, 1992

Investigators: Jorge Frangi and A.E. Lugo

Location: Montane Palm Floodplain Forest, Luquillo Experimental Forest,

Puerto Rico

Date of collection: 1990

89. Population dynamics of Manilkara bidentata (A.DC.) Cher. In the Luquillo

Experimental Forest, Puerto Rico

Investigator: Chengxia You (Ph.D. dissertation University of Tennessee) 

Location: El Verde, Luquillo Experimental Forest, Puerto Rico

Date of collection: February 1987

90. Nutrients and mass in litter and top soil of ten tropical tree plantations

Investigators: Ariel E. Lugo, E. Cuevas, and M.J. Sánchez

Location: Arboretum, El Verde, Luquillo Experimental Forest,

Puerto Rico

Date of collection: October 1986

91. Plumeria study

Investigator: Eva Dávila

Location: Guánica Forest, Maricao Forest, Ponce and Dorado, Puerto Rico

Date of collection: 1985
25

Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

92. Porciento de cenizas en la flora epifitica sobre Prestoea montana

Investigator: Carmen L. Báez (University of Puerto Rico High School)

Location: Luquillo Experimental Forest, Puerto Rico

Date of collection: 1985

93. Hurricane damage to a flood plain forest in the Luquillo Mountains

of Puerto Rico. Biotropica. 23(4a): 324–335, 1991

Investigators: Jorge Frangi and A.E. Lugo

Location: Montane palm floodplain forest, Luquillo Experimental Forest,

Puerto Rico

Date of collection: 1980/1990

94. Comparison of tropical tree plantations with secondary forest of similar

age. Ecological Monographs. 62(1): 1–41, 1992 Study 2501-Understory

study

Investigator: Ariel E. Lugo

Location: Guzmán, Cubuy, Sabana, El Verde, Luquillo Experimental

Forest, Puerto Rico

Date of collection: 1982–1984

95. Decayed roots collection

Investigators: Patrick Kangas and A.E. Lugo

Location: El Verde, Luquillo Experimental Forest, Puerto Rico

Date of collection: August 1986

96. The influence of density on stand development, biomass partitioning

and nutrient allocation in Albizia lebbeck (L.) Benth. Plantations in

Puerto Rico 

Investigator: John Parrotta (Ph.D. dissertation Yale University)

Location: Toa Baja Experimental Farm, Puerto Rico

Date of collection: 1985–1986

97. Wood density. SO-ITF-SM-15,32,35,42,44,45,47,48,58,60,69,75,77 in

IITF Silvics Manual

Investigator: John Francis

Location: Puerto Rico

Date of collection: 1985–1990
26

International Institute of Tropical Forestry II

98. Restoration and rehabilitation of tropical forests. Environmental Sciences

Course, University of Puerto Rico, Río Piedras Campus

Investigator: Ariel E. Lugo

Location: Río Mameyes (river basin), Luquillo Experimental Forest,

Puerto Rico

Date of collection: September–October 1989

99. Nutrient dynamics of a Puerto Rican subtropical dry forest. Journal of

Tropical Ecology. 2: 55–72, 1986

Investigators: Peter Murphy and A.E. Lugo

Location: Guánica Forest, Guánica, Puerto Rico

Date of collection: 1981–1983

100. Leaf decomposition study

Investigator: Yiovani Santiago Molina (Naranjito High School)

Location: Bisley Watersheds; Luquillo Experimental Forest, Puerto Rico

Date of collection: 1988

101. Comparison of tropical tree plantations with secondary forest of similar

ages. Ecological Monographs. 62(1): 1–41, 1992

Investigator: Ariel E. Lugo

Location: Guzmán, Cubuy, Sabana, El Verde, Luquillo Experimental

Forest, Puerto Rico

Date of collection: 1982–1984

102. Mineral content of leaves from trees growing on serpentine soils

under contrasting rainfall regimes in Puerto Rico. Plant and Soil.

158: 13–21, 1994

Investigators: Ernesto Medina, E. Cuevas, J. Figueroa, and A.E. Lugo

Location: Maricao Forest, Puerto Rico

Date of collection: June 1990

103. Comparative analysis of the nutritional status of Mahogany plantations

in Puerto Rico. In: Lugo, A. E.; Figueroa, J.; Alayón, M., ed. Big-leaf

mahogany genetics. Ecology and Management: 129–145

Investigators: Ernesto Medina and E. Cuevas

Location: Puerto Rico

Date of collection: 1986–1987
27

Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

104. Physiological ecology of dominant woody species in a dry coastal forest

on limestone (Guánica) forest

Investigator: Ernesto Medina

Location: Guánica Forest, Puerto Rico

Date of collection: 1986–1987

105. Plasticity of crassulacean acid metabolism (CAM) and biomass allocation

of the pantropical Kalanchoe pinnata (LAM.) Pers. Acta Cientifica.

12(1-3): 59–66, 1998

Investigator: Ernesto Medina

Location: IITF Nursery, Puerto Rico

Date of collection: 1987

106. Bisley Study—litter fall collection

Investigator: Fred Scatena

Location: Bisley, Luquillo Experimental Forest, Puerto Rico

Date of Collection: 1990–2000

107. MRCE foliar nutrients/fertilizer plots

Investigator: Heather Erickson

Location: El Verde and Pico del Este, Puerto Rico

Date of collection: March–October 1993

108. Nitrogen oxide fluxes and nitrogen cycling during post agricultural

succession and forest fertilization in the humid tropics. Ecosystems.

4: 67–84, 2001

Investigators: Michael Keller and H. Erickson

Location: LTER, Sabana, Mameyes, El Yunque, Puerto Rico

Date of collection: 1995–1996

109. Former land-use and tree species affect nitrogen oxide emissions from

a tropical dry forest. Oecologia. 130: 297–308, 2002

Investigator: Heather Erickson

Location: Guánica Forest, Guánica, Puerto Rico

Date of collection: March 1996–March 1997

110. Woods Hole Nasa study/clip plots

Investigator: Heather Erickson

Location: Sabana and Mameyes, Puerto Rico

Date of collection: April 1995–1996
28

International Institute of Tropical Forestry II

111. Long-term ecological research Hurricane Hugo loose litter collection

Investigators: Fred Scatena and J. Lodge

Location: East Peak, Caribbean National Forest, Puerto Rico

Date of collection: September 19, 1989

112. The metabolism of Río Mameyes, Puerto Rico: carbon fluxes in a tropical

rain forest river

Investigator: Jorge Ortiz (Ph.D. dissertation University of Colorado)

Location: Río Mameyes Watershed, Luquillo Experimental Forest,

Puerto Rico

Date of collection: February 1995–March 1996

113. Litter fall experiment: Sabana 60 years

Investigator: Michael Keller

Location: Sabana, Luquillo Experimental Forest, Puerto Rico

Date of collection: March–October 1997

114. Roots Sabana forest 60 years

Investigator: Michael Keller

Location: Sabana, Luquillo Experimental Forest, Puerto Rico

Date of collection: July 1998

115. Simple and inexpensive method for extracting wood density samples

from tropical hardwoods. Tree Planter’s Notes. 45(1): 10–12, 1994

Investigator: John Francis

Location: Puerto Rico

Date of collection: 1992–1993

116. Acromia media O.F. Cook. SO-ITF-SM-68

Investigator: John Francis 

Location: Puerto Rico

Date of collection: 1993

117. Prunus occidentalis S.W. SO-ITF-SM-79

Investigator: Salvador Alemañy 

Location: Puerto Rico

Date of collection: 1992

118. Pouteria multiflora (A.D.C) EYMA. SO-ITF-SM-62

Investigator: John Francis

Location: Puerto Rico

Date of collection: 1993
29

Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

119. Roystonea borinquena O.F. Cook. SO-ITF-SM-55

Investigator: John Francis

Location: Puerto Rico

Date of collection: 1990

120. Dacroydes excelsa

Investigator: John Francis

Location: El Verde, Luquillo Experimental Forest, Puerto Rico

Date of collection: October 1991

121. Studies of life history of three herbaceous species in the Bisley

Watersheds, Puerto Rico

Investigator: Amy Arnold (M.Sc. thesis Tennessee University)

Location: Bisley, Luquillo Experimental Forest, Puerto Rico

Date of collection: November 1995

122. Effects of nutrient availability and other elevational changes on bromeliad

populations and their invertebrate communities in a humid tropical forest

in Puerto Rico. Journal of Tropical Ecology. 16: 167–188, 2000

Investigators: Barbara Richardson, M.J. Richardson, F. Scatena,

and W.H. McDowell

Location: Tabonuco, Palo Colorado and Dwarf forests Luquillo 

Experimental Forest, Puerto Rico

Date of collection: 1996

123. Bryophyte communities on different stages of decaying wood

Investigator: Inés Sastre

Location: Bisley Watersheds, Luquillo Experimental Forest, Puerto Rico

Date of collection: 1990–1991

124. Tabonuco Project (Dacryodes excelsa)

Investigator: Migdalia Álvarez 

Location: Various sites, Puerto Rico

Date of collection: 1994

125. Contrasting light environments and response flexibility of trees in

the Luquillo Mountains of Puerto Rico 

Investigator: Denny Fernández (Ph.D. dissertation University of

Puerto Rico, Rio Piedras Campus)

Location: El Verde, Luquillo Experimental Station, Puerto Rico

Date of collection: May 1996
30

International Institute of Tropical Forestry II

126. Estimating biomass and carbon content of saplings in Puerto Rican

secondary forests. Caribbean Journal of Science. 26: 346–350, 2000

Investigator: John Francis

Location: Various sites, Puerto Rico

Date of collection: 1999

127. Arboretum root standing stock

Investigators: Ariel E. Lugo, E. Cuevas, and M.J. Sánchez

Location: Cienaga Alta, Luquillo Experimental Forest, Puerto Rico

Date of collection: December 1986

128. Acrocomia media O.F. Cook. SO-ITF-SM-68

Investigator: John Francis

Location: Punta Salinas, Toa Baja; Puerto Rico

Date of collection: October 1991

129. Nonindigenous bamboo along headwater streams of the Luquillo

Mountains, Puerto Rico: leaf fall, aquatic leaf decay and patterns

of invasion. Journal of Tropical Ecology. 16: 499–516, 2000

Investigators: Paul O’Connor, A.P. Covich, F.N. Scatena, and L.L. Loope

Location: Bisley watershed, Luquillo Experimental Forest, Puerto Rico

Date of collection: 1997

130. Nutrient return accumulation in litter of a secondary forest in the coffee

region of Puerto Rico. Acta Científica. 13(1-3): 43–74, 1999

Investigators: Ariel E. Lugo, C. Domínguez, A. Santos, and E. Torres

Location: Torrecilla secondary forest, Bo. Hincado, Barranquitas,

Puerto Rico

Date of collection: 1990–1993

131. Biomasa y nutrientes en raíces y brinzales de un bosque secundario

en la zona cafetalera de Utuado. Acta Científica. 13(1–3): 75–87, 1999

Investigators: Ariel E. Lugo, C. Domínguez, and N. Méndez

Location: Bo. Caguana, Utuado, Puerto Rico

Date of collection: February 1998

132. Ecological study of a secondary forest in Caguana, Utuado

Investigators: Ariel E. Lugo and N. Méndez

Location: Bo. Caguana, Utuado, Puerto Rico

Date of collection: 1998–1999
31

Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

133. Comparative study of nutrient dynamics of two forest types in Luquillo

Experimental Forest—loose litter

Investigators: Fu Sheinglei and A.E. Lugo

Location: El Verde, Luquillo Experimental Forest, Puerto Rico

Date of collection: July 1995

134. Seasonal dynamics and succession in a subtropical dry forest

Investigator: Ariel E. Lugo

Location: Guánica forest, Guánica, Puerto Rico

Date of collection: 1992

135. Bisley study—biomass

Location: Bisley, El Yunque, Puerto Rico

Investigator: Fred Scatena

Date of collection: 1989–1999

136. Insects population collection 1999–2000

Investigator: Barbara Richardson

Location: El Verde, Puerto Rico

Date of collection: 1999–2000

137. Nutrients, diversity, and community structure of two phytotelm systems

in a lower montane forest, Puerto Rico

Investigators: Barbara Richardson, C. Rogers, and M.J. Richardson

Location: El Verde, Puerto Rico

Date of collection: 1998

138. Physiological characteristic of two tree species growing on a pasture

Investigators: Adisel Montaña and N. Fetcher

Site location: Sabana, Luquillo, Puerto Rico

Date of collection: February 1998

139. Dynamics of organic matter and nutrient return from litter fall in stands

of ten tropical tree plantation species. Forest Ecology and Management.

112: 263–279, 1998

Investigators: Elvira Cuevas and A.E. Lugo

Location: Arboretum, Cienaga Alta, Luquillo Experimental Forest,

Puerto Rico

Date of collection: 1986–1987
32

International Institute of Tropical Forestry II

140. Hurricane Georges loose litter

Investigators: Rebeca Ostertag and F. Scatena

Location: Cubuy, Bisley and Guánica, Puerto Rico

Date of collection: 1998–1999

141. Effects of fertilization on plant growth in landslide in the Luquillo Forest

Investigator: Ned Fetcher

Location: Palo Hueco, Luquillo Experimental Forest, Puerto Rico

Date of collection: 1989

142. Changes in necromass and nutrients on the Forest floor of a palm

floodplain in forest in the Luquillo mountains of Puerto Rico.

Caribbean Journal of Science. 39: 265–272, 2003

Investigators: Ariel E. Lugo and J. Frangi

Location: Montane palm forest, Luquillo Experimental Forest,

Puerto Rico

Date of collection: October 2000

143. Reorganization of the palm forest in the Luquillo Experimental Forest,

Puerto Rico, 5 years after Hurricane Hugo

Investigator: Jorge Frangi

Location: Montane palm flood plain forest, Luquilo Experimental Forest,

Puerto Rico

Date of collection: June 1995

144. Sabana Seca climate study

Investigators: Michael Van Der Molen and F. Scatena

Location: Sabana Seca, Puerto Rico

Date of collection: 1998

145. Pivotal fungal and plant species and its role influencing the rates of

decomposition in a tropical forest

Investigators: Mirna Santana and J. Lodge

Location: El Verde, Sabana, Bisley, Luquillo Experimental Forest,

Puerto Rico

Date of collection: August 2000

146. Landslide litter decomposition

Investigators: Randall Myster and D. Shaffer

Location: El Verde Field Station, Puerto Rico

Date of collection: February–October 1996
33

Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

147. Moon phase influence over palm leaves nutrients

Investigators: Fred Scatena and S. Moya

Location: Bisley, El Yunque, Puerto Rico

Date of collection: July–September 2000

148. Mineral content of leaves from trees growing on serpentine soils in

Puerto Rico

Investigators: Ernesto Medina, E. Cuevas, J. Figueroa, and A.E. Lugo

Location: Susua Forest, Puerto Rico

Date of collection: August 1993

149. Comparison of the leaf nutrient composition of trees from contrasting

soil environments: Susua Forest and Guánica Forest

Investigators: Ernesto Medina, E. Cuevas, and J. Figueroa

Location: Susua forest and Guánica forest

Date of collection: October 1986–January 1992

150. Ontogenic comparison in leaf morphology and physiology of tropical

rainforest tree species

Investigator: Shiyun Wen

Location: El Verde, Luquillo Experimental Forest, Puerto Rico

Date of collection: 1999–2000

151. Spatial patterns in forest nutrient pools, growth and disturbance—plant

collection

Investigator: Skip Van Bloem

Location: Guánica Forest, Guánica, Puerto Rico

Date of collection: September 1998

152. Soil macro fauna and litter nutrients in three tropical tree plantations on

a disturbed site in Puerto Rico. Forest Ecology and Management.

170: 161–171, 2002

Investigators: Matthew Warren and X. Zou

Location: Toa Baja Plantation, Puerto Rico

Date of collection: February 1998

153. Exotic earthworms acelerate plant litter decomposition in a Puerto Rican

pasture and a wet Forest. Ecological Applications. 12(5): 1406–1417, 2002

Investigator: Z.G. Liu and X. Zou

Location: Secondary Forest and pine plantation in Guzmán, Luquillo

Experimental forest, Puerto Rico

Date of collection: May 1996
34

International Institute of Tropical Forestry II

154. Fine litterfall and related nutrient inputs resulting from Hurricane

Hugo in subtropical wet and lower montane rain forests of Puerto Rico.

Biotropica. 23(4A): 336–342, 1991

Investigators: Jean Lodge, F.N. Scatena, C.E. Asbury, and M.J. Sánchez

Location: El Verde, Caribbean National Forest, Puerto Rico

Date of collection: September 1989

155. Prescottia oligantha

Investigator: Carlos Rodríguez

Location: Luquillo and Maricao, Puerto Rico

Date of collection: May 1989

156. Above and below organic matter storage and production in a tropical pine

plantation and a paired broadleaf secondary forest

Investigators: Elvira Cuevas, S. Brown, and A.E. Lugo

Location: Guzmán, Luquillo Experimental Forest, Puerto Rico

Date of collection: 1987–1989

157. Study of the termite species N. costalis in the Piñones Forest

Investigator: Verónica Carrasquillo

Location: Piñones Forest, Loiza, Puerto Rico

Date of collection: December 23, 1992
35

Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

Appendix 2: List of Species

Table 10—Species, family, growth habit, and source

No. Name Family Growth habit Source Group

1 Acrocomia media Arecaceae Tree 116, 128 Monocot

2 Acrostichum aureum Pteridaceae Forb/herb 78, 77 Fern

3 Adiantum latifolium Pteridaceae Forb/herb 135 Fern

4 Adiantum pyramidale Pteridaceae Forb/herb 135 Fern

5 Adiantum spp. Pteridaceae Forb/herb 135 Fern

6 Albizia lebbeck Fabaceae Tree 96 Dicot

7 Albizia procera Fabaceae Tree 126 Dicot

8 Alchornea latifolia Euphorbiaceae Tree 68, 97, 135 Dicot

9 Alchorneopsis floribunda Euphorbiaceae Tree 68 Dicot

10 Alchorneopsis portoricensis Euphorbiaceae Tree 135 Dicot

11 Aleurites moluccana Euphorbiaceae Tree 97 Dicot

12 Allamanda violacea Apocynaceae Shrub/vine 91 Dicot

13 Alsophila portoricensis Cyatheaceae Tree 135, 143 Fern

14 Amyris elemifera Rutaceae Tree/shrub 104, 149, 151 Dicot

15 Andira inermis Fabaceae Tree 94, 135 Dicot

16 Anetium citrifolium Vittariaceae Forb/herb 135 Fern

17 Anthocephalus chinensis Rubiaceae Tree 90, 115, 139 Dicot

18 Ardisia glauciflora Myrsinaceae Tree/shrub 102 Dicot

19 Ardisia solanacea Myrsinaceae Tree/shrub 126 Dicot

20 Artocarpus altilis Moraceae Tree 97 Dicot

21 Asplenium spp. Aspleniaceae Herb 135 Fern

22 Avicennia germinans Verbenaceae Tree/shrub 74, 78, 80, 87, 

157

Dicot

23 Bambusa spp. Poaceae Tree/shrub 129 Monocot

24 Bambusa vulgaris Poaceae Tree/shrub 94 Monocot

25 Beilschmiedia pendula Lauraceae Tree 68 Dicot

26 Blechnum occidentale Acanthaceae 135 Dicot

27 Bolbitis aliena Dryopteridaceae Forb/herb 135 Fern

28 Bolbitis nicotianifolia Dryopteridaceae Forb/herb 135 Fern

29 Bourreria succulenta Boraginaceae Tree/shrub 104, 149, 151 Dicot

30 Bromeliads Bromeliaceae 93, 135 Other

31 Bryophytes 88, 123

32 Buchenavia tetraphylla Combretaceae Tree 153 Dicot

33 Bucida buceras Combretaceae Tree 104, 126, 149 Dicot

34 Bursera simaruba Burseraceae Tree/shrub 97, 102, 104, 

149, 151

Dicot

35 Byrsonima lucida Malpighiaceae Tree/shrub 102, 149 Dicot
36

International Institute of Tropical Forestry II

No. Name Family Growth habit Source Group

36 Byrsonima spicata Malpighiaceae Tree/shrub 131 Dicot

37 Byrsonima wadsworthii Malpighiaceae Tree/shrub 93 Dicot

38 Calophyllum antillanum Clusiaceae Tree/shrub 59, 94, 101, 126 Dicot

39 Canna glauca Cannaceae Forb/herb 135 Monocot

40 Canna spp. Cannaceae Forb/herb 135 Monocot

41 Capparis cynophallophora Capparaceae Tree/shrub 104, 149 Dicot

42 Casearia arborea Flacourtiaceae Tree/shrub 94, 101, 135 Dicot

43 Casearia decandra Flacourtiaceae Tree/shrub 126, 148 Dicot

44 Casearia guianensis Flacourtiaceae Tree/shrub 126 Dicot

45 Casearia sylvestris Flacourtiaceae Tree/shrub 94, 148 Dicot

46 Cassine xylocarpa Celastraceae Tree/shrub 102, 104, 149 Dicot

47 Casuarina equisetifolia Casuarinaceae Tree 152 Dicot

48 Cecropia schreberiana Cecropiaceae Tree 107, 123, 129, 

135, 141, 146

Dicot

49 Cecropia sp. Cecropiaceae Tree 146 Dicot

50 Cissampelos pareira Menispermaceae Shrub/vine 135 Dicot

51 Cissus erosa Vitaceae Shrub/vine 135 Dicot

52 Cissus verticillata Vitaceae Shrub/vine 135 Dicot

53 Citrus sinensis Rutaceae Tree/shrub 97, 101 Dicot

54 Clibadium erosum Asteraceae Tree/shrub 135 Dicot

55 Clusia rosea Clusiaceae Tree/shrub 71, 97, 102, 104, 

149

Dicot

56 Cnemidaria horrida Cyatheaceae Shrub/vine 135 Fern

57 Coccoloba microstachya Polygonaceae Tree/shrub 102, 104, 

149,151

Dicot

58 Coccoloba diversifolia Polygonaceae Tree/shrub 104, 149, 151 Dicot

59 Coffea arabica Rubiaceae Tree/shrub 53, 94, 131 Dicot

60 Colubrina arborescens Rhamnaceae Tree/shrub 104, 149 Dicot

61 Colubrina elliptica Rhamnaceae Tree/shrub 104, 149 Dicot

62 Commelina diffusa Commelinaceae Forb/herb 135 Monocot

63 Commelina spp. Commelinaceae Forb/herb 135 Monocot

64 Conocarpus erectus Combretaceae Tree/shrub 87, 157 Dicot

65 Cordia borinquensis Boraginaceae Tree/shrub 68, 135 Dicot

66 Cordia sulcata Boraginaceae Tree/shrub 68, 97 Dicot

67 Croton poecilanthus Euphorbiaceae Tree 68, 71, 93, 145 Dicot

68 Ctenitis hirta Dryopteridaceae Forb/herb 135 Fern

69 Cupania americana Sapindaceae Tree 97 Dicot

70 Cyathea arborea Cyatheaceae Tree 135 Fern

71 Cyclopogon cranichoides Orchidaceae Forb/herb 135 Monocot

72 Cyrilla racemiflora Cyrillaceae Tree/shrub 67, 93 Dicot

Table 10—Species, family, growth habit, and source (continued)
37

Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

No. Name Family Growth habit Source Group

73 Dacryodes excelsa Burseraceae Tree 68, 71, 94, 100, 

101, 107, 120, 

123, 124, 135, 

153

Dicot

74 Dacryodes sp. Burseraceae Tree 146 Dicot

75 Danaea nodosa Marattiaceae Forb/herb 129, 135 Fern

76 Daphnopsis philippiana Thymelaeaceae Tree/shrub 93 Dicot

77 Delonix regia Fabaceae Tree 115 Dicot

78 Dendropanax arboreus Araliaceae Tree/shrub 71 Dicot

79 Dennstaedtia bipinnata Dennstaedtiaceae Forb/herb 135 Fern

80 Dennstaedtia obtusifolia Dennstaedtiaceae Forb/herb 135 Fern

81 Dennstaetdia spp. Dennstaedtiaceae Forb/herb 135 Fern

82 Dialium guineense Fabaceae 115 Dicot

83 Dieffenbachia seguine Araceae Forb/herb 121 Monocot

84 Dioscorea polygonoides Dioscoreaceae Vine/forb-herb 135 Monocot

85 Ditta myricoides Euphorbiaceae Tree/shrub 93 Dicot

86 Drypetes glauca Euphorbiaceae Tree/shrub 68 Dicot

87 Enterolobium cyclocarpum Fabaceae Tree 97 Dicot

88 Erithalis fruticosa Rubiaceae Tree/shrub 104, 149, 151 Dicot

89 Erythrodes plantaginea Orchidaceae Forb/herb 135 Monocot

90 Erythroxylum areolatum Erythroxylaceae Forb/herb 104, 149 Dicot

91 Erythroxylum rotundifolium Erythroxylaceae Tree/shrub 104, 149 Dicot

92 Eucalyptus patentinervis Myrtaceae Tree 90, 139 Dicot

93 Eucalyptus robusta Myrtaceae Tree 97, 152 Dicot

94 Eucalyptus saligna Myrtaceae Tree 90, 139 Dicot

95 Eugenia borinquensis Myrtaceae Tree/shrub 93 Dicot

96 Eugenia eggersii Myrtaceae Tree/shrub 93,135 Dicot

97 Eugenia foetida Myrtaceae Tree/shrub 149 Dicot

98 Eugenia maleolens Myrtaceae Tree/shrub 104 Dicot

99 Eugenia monticola Myrtaceae Tree/shrub 126 Dicot

100 Eugenia rhombea Myrtaceae Tree/shrub 104, 149 Dicot

101 Eugenia stahlii Myrtaceae Tree/shrub 94 Dicot

102 Exostema caribaeum Rubiaceae Tree/shrub 102, 104, 149, 

151

Dicot

103 Faramea occidentalis Rubiaceae Tree/shrub 126 Dicot

104 Ficus citrifolia Moraceae Tree/shrub 55, 97 Dicot

105 Forest floor-logs 95

106 Forest floor-roots 127

107 Genipa americana Rubiaceae Tree 97 Dicot

Table 10—Species, family, growth habit, and source (continued)
38

International Institute of Tropical Forestry II

No. Name Family Growth habit Source Group

108 Gleichenia spp. (has various names) Gleicheniaceae Forb/herb 135 Fern

109 Gomidesia lindeniana Myrtaceae Tree/shrub 58 Dicot

110 Gonzalagunia spicata Rubiaceae Tree/shrub 135, 146 Dicot

111 Guaiacum officinale Zygophyllaceae Tree 104, 149 Dicot

112 Guaiacum sanctum Zygophyllaceae Tree 104, 149 Dicot

113 Guanica forest 109, 134

114 Guapira obtusata Nyctaginaceae Tree/shrub 102 Dicot

115 Guararibaea turbinata Bombacaceae Tree/shrub 68 Dicot

116 Guarea glabra Meliaceae Tree/shrub 68, 93, 94, 101 Dicot

117 Guarea guidonia Meliaceae Tree/shrub 71, 125, 126, 

129, 135

Dicot

118 Guarea ramiflora Meliaceae Tree/shrub 125 Dicot

119 Guazuma ulmifolia Sterculiaceae Tree/shrub 97 Dicot

120 Guettarda pungens Rubiaceae Tree/shrub 149 Dicot

121 Guettarda scabra Rubiaceae Tree/shrub 102, 149 Dicot

122 Guzmania berteriana Bromeliaceae Forb/herb 135 Monocot

123 Gymnanthes lucida Euphorbiaceae Tree/shrub 104, 149, 151 Dicot

124 Heliconia caribaea Heliconiaceae Forb/herb 122, 135, 137 Monocot

125 Henriettea squamulosum Melastomataceae Tree 93 Dicot

126 Herbaceous 135

127 Hernandia sonora Hernandiaceae Tree/shrub 90, 139 Dicot

128 Heteropteris laurifolia Malpighiaceae Tree/shrub/vine 135 Dicot

129 Heterotrichum cymosum Melastomataceae Tree/shrub 146 Dicot

130 Hibiscus elatus Malvaceae Tree 90, 97, 139 Dicot

131 Hillia parasitica Rubiaceae Tree/shrub/vine 135 Dicot

132 Homalium racemosum Flacourtiaceae Tree 148 Dicot

133 Hyeronima clusioides Euphorbiaceae Tree/shrub 97 Dicot

134 Hypolepsis spp. 135

135 Ichnanthus pallens Poaceae Graminoid 135 Monocot

136 Ichnanthus spp. Poaceae graminoid 135 Monocot

137 Ilex sideroxyloides Aquifoliaceae Tree 93 Dicot

138 Inga fagifolia Fabaceae Tree 145 Dicot

139 Inga laurina Fabaceae Tree 70, 71, 126, 135 Dicot

140 Inga vera Fabaceae Tree 71, 100, 145, 146 Dicot

141 Jacquinia berteroi Theophrastaceae Tree/shrub 104, 149, 151 Dicot

142 Juglans jamaicensis Juglandaceae Tree 115 Dicot

143 Kalanchoe pinnata Crassulaceae Forb/herb 105 Dicot

144 Khaya grandifolia Meliaceae Tree 97 Dicot

145 Khaya nyasica Meliaceae Tree 90, 97, 135, 139 Dicot

Table 10—Species, family, growth habit, and source (continued)
39

Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

No. Name Family Growth habit Source Group

146 Khaya senegalensis Meliaceae Tree 97 Dicot

147 Krugiodendron ferreum Rhamnaceae Tree/shrub 104, 149 Dicot

148 Laguncularia racemosa Combretaceae Tree/shrub 72, 73, 74, 78, 

83, 87, 157

Dicot

149 Leucaena leucocephala Fabaceae Tree/shrub 126, 148, 152 Dicot

150 Lonchitis hirsuta Dennstaedtiaceae Forb/herb 135 Fern

151 Maesopsis eminii Rhamnaceae Tree 97 Dicot

152 Magnolia portoricensis Magnoliaceae Tree 115 Dicot

153 Magnolia splendens Magnoliaceae Tree 71, 93 Dicot

154 Mahogany plantation 133

155 Mangrove 79

156 Manilkara bidentata Sapotaceae Tree 68, 71, 89, 94, 

101, 107, 125, 

129, 145, 153

Dicot

157 Marcgravia rectiflora Marcgraviaceae 135 Dicot

158 Marcgravia spp. Marcgraviaceae 135 Dicot

159 Matayba domingensis Sapindaceae Tree 68 Dicot

160 Megalastrum subincisa Dryopteridaceae Forb/herb 135 Fern

161 Melicoccus bijugatus Sapindaceae Tree 97 Dicot

162 Miconia impetiolaris Melastomataceae Tree/shrub 129 Dicot

163 Miconia prasina Melastomataceae Tree/shrub 68, 94, 126, 131, 

138, 

Dicot

164 Miconia racemosa Melastomataceae Tree/shrub 129, 146 Dicot

165 Miconia sp. Melastomataceae Tree/shrub 146 Dicot

166 Miconia spp. Melastomataceae Tree/shrub 143 Dicot

167 Miconia tetrandra Melastomataceae Tree/shrub 92, 94 Dicot

168 Micropholis chrysophylloides Sapotaceae Tree 92 Dicot

169 Micropholis garciniifolia Sapotaceae Tree/shrub 92 Dicot

170 Mikania cordifolia Asteraceae Vine/forb/herb 135 Dicot

171 Mixed-species Guanica forest 99

172 Musa paradisiaca Musaceae Tree 135 Monocot

173 Musa spp. Musaceae Tree 135 Monocot

174 Myrcia deflexa Myrtaceae Tree/shrub 93, 94, 135 Dicot

175 Myrcia splendens Myrtaceae Tree/shrub 94, 101, 126 Dicot

176 Neolaugeria resinosa Rubiaceae ? 102 Dicot

177 Nephrolepsis portoricensis 135

178 Nephrolepis rivularis 135

179 Nepsera aquatica Melastomataceae Shrub-forb/herb 135 Dicot

180 Ocotea leucoxylon Lauraceae Tree 93, 94, 126, 135 Dicot

181 Ocotea moschata Lauraceae Tree 97 Dicot

Table 10—Species, family, growth habit, and source (continued)
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No. Name Family Growth habit Source Group

182 Ocotea spathulata Lauraceae Tree 93 Dicot

183 Odontosoria spp. Dennstaedtiaceae Forb/herb 135 Fern

184 Olyra latifolia Poaceae Graminoid 135 Monocot

185 Ormosia krugii Fabaceae Tree 68, 135 Dicot

186 Ouratea littoralis Ochnaceae Tree/shrub 102, 149 Dicot

187 Palicourea crocea Rubiaceae Tree/shrub 135 Dicot

188 Palicourea riparia Rubiaceae Tree/shrub 94, 135, 146 Dicot

189 Panicum maximum Poaceae Graminoid 144 Monocot

190 Pasture 110, 153

191 Paullinia pinnata Sapindaceae Vine 135 Dicot

192 Paullinia spp. Sapindaceae Vine 135 Dicot

193 Peperomia spp. Piperaceae Forb/herb 135 Dicot

194 Petitia domingensis Verbenaceae Tree/shrub 149 Dicot

195 Philodendron angustatum Araceae Vine/forb/herb 135 Monocot

196 Philodendron scandens Araceae Vine/forb/herb 135 Monocot

197 Philodendron spp. Araceae Vine/forb/herb 135 Monocot

198 Phoradendron racemosum Viscaceae Shrub 104, 149 Dicot

199 Phytolacca icosandra Phytolaccaceae Forb/herb 135, 141 Dicot

200 Phytolacca rivinoides Phytolaccaceae Forb/herb 135 Dicot

201 Pictetia aculeata Fabaceae Tree/shrub 102, 104, 149, 

151 

Dicot

202 Pilea inaequalis Urticaceae Forb/herb 135 Dicot

203 Pilea krugii Urticaceae Forb/herb 135 Dicot

204 Pimenta racemosa Myrtaceae Tree/shrub 126 Dicot

205 Pine plantation 153, 156

206 Pinus caribaea Pinaceae Tree 69, 90, 101, 115 Gimnosperm

207 Pinus caribaea hondurensis Pinaceae Tree 139 Gimnosperm

208 Pinus elliottii Pinaceae Tree 139 Gimnosperm

209 Pinus massoniana Pinaceae Tree 68 Gimnosperm

210 Pinus ocarpa Pinaceae Tree 115 Gimnosperm

211 Piper aduncum Piperaceae Tree/shrub 129 Dicot

212 Piper glabrescens Piperaceae Tree/shrub 135, 146 Dicot

213 Piper hispidum Piperaceae Tree/shrub 135, 146 Dicot

214 Pisonia albida Nyctaginaceae Tree 104, 149 Dicot

215 Pisonia subcordata Nyctaginaceae Tree 102 Dicot

216 Pithecellobium dulce Fabaceae Tree/shrub 97 Dicot

217 Plumeria alba Apocynaceae Tree/shrub 91 Dicot

218 Plumeria obtusa Apocynaceae Tree/shrub 91 Dicot

219 Plumeria rubra (red) Apocynaceae Tree/shrub 91 Dicot

Table 10—Species, family, growth habit, and source (continued)
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Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

No. Name Family Growth habit Source Group

220 Plumeria rubra (white) Apocynaceae Tree/shrub 91 Dicot

221 Polybotrya cervina Dryopteridaceae Forb/herb 135 Fern

222 Polypodium chnoodes Polypodiaceae Forb/herb 135 Fern

223 Polypodium crassifolium Polypodiaceae Forb/herb 135 Fern

224 Polypodium spp. Polypodiaceae Forb/herb 135 Fern

225 Pouteria multiflora Sapotaceae Tree 118 Dicot

226 Prescotia oligantha Orchidaceae Forb/herb 155 Monocot

227 Prestoea montana Arecaceae Tree 68, 69, 92, 93, 

107, 135, 147, 

150

Monocot

228 Prunus occidentalis Rosaceae Tree 117 Dicot

229 Psychotria berteriana Rubiaceae Tree/shrub 93, 135, 143, 146 Dicot

230 Psychotria maleolens Rubiaceae Tree/shrub 143 Dicot

231 Psychotria sp. Rubiaceae Tree/shrub 94 Dicot

232 Pteris altissima Pteridaceae Forb/herb 135 Fern

233 Pterocarpus macrocarpus Fabaceae Tree 97 Dicot

234 Pterocarpus officinalis Fabaceae Tree 76, 78, 83, 84, 97 Dicot

235 Rhizophora mangle Rhizophoraceae Tree 72, 73, 74, 75, 

77, 78, 80, 81, 

82, 85, 86, 87, 

157

Dicot

236 Rourea surinamensis Connaraceae Vine/shrub 135 Dicot

237 Roystonea borinquena Arecaceae Tree 119 Monocot

238 Sabicea hirsuta Rubiaceae Vine/shrub 135 Dicot

239 Sapindus saponaria Sapindaceae Tree/shrub 129 Dicot

240 Sapium laurocerasus Euphorbiaceae Tree 68, 71, 135, 145 Dicot

241 Sauvagesia erecta Ochnaceae Shrub-forb/herb 135 Dicot

242 Schefflera morototoni Araliaceae Tree 35, 68, 71, 94, 

126, 135

Dicot

243 Schizolobium parahybum Fabaceae Tree 97 Dicot

244 Schlegelia brachyantha Bignoniaceae Vine/shrub 135 Dicot

245 Scleria canescens Cyperaceae Graminoid 135 Monocot

246 Scleria spp. Cyperaceae Graminoid 135 Monocot

247 Secondary forest 106, 108, 111, 

112, 113, 114, 

124, 130, 131, 

132, 136, 140, 

142, 153, 154, 

156

248 Securidaca virgata Polygalaceae Vine/shrub 135 Dicot

249 Selaginella krugii Selaginellaceae Forb/herb 135 Lycopod

Table 10—Species, family, growth habit, and source (continued)
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No. Name Family Growth habit Source Group

250 Sloanea berteriana Elaeocarpaceae Tree 67, 70, 135, 150 Dicot

251 Solanum torvum Solanaceae Tree/shrub 146 Dicot

252 Spathodea campanulata Bignoniaceae Tree 97, 126 Dicot

253 Swietenia humilis Meliaceae Tree 103 Dicot

254 Swietenia macrophylla Meliaceae Tree 90, 94, 101, 103, 

104, 129, 139

Dicot

255 Swietenia macrophylla x mahagoni Meliaceae Tree 103 Dicot

256 Swietenia mahogoni Meliaceae Tree 102, 103, 149 Dicot

257 Swietenia sp. Meliaceae Tree 103 Dicot

258 Syzygium jambos Myrtaceae Tree 94, 101, 126, 

129, 131

Dicot

259 Tabebuia haemantha Bignoniaceae Tree/shrub 102, 149 Dicot

260 Tabebuia heterophylla Bignoniaceae Tree/shrub 58, 94, 101, 115, 

126, 138

Dicot

261 Tabebuia rigida Bignoniaceae Tree/shrub 93, 107 Dicot

262 Tabebuia spp. Bignoniaceae Tree/shrub 149 Dicot

263 Tabonuco forest 153

264 Tabonuco secondary forest 133

265 Tectaria trifoliata Dryopteridaceae Forb/herb 135 Fern

266 Thelypteris deltoidea Thelypteridaceae Gorb/herb 135 Fern

267 Terebraria resinosa Rubiaceae Shrub 58 Dicot

268 Terminalia ivorensis Combretaceae Tree 90, 139 Dicot

269 Ternstroemia stahlii Theaceae Tree/shrub 102, 149 Dicot

270 Tetragastris balsamifera Burseraceae Tree 135 Dicot

271 Thelypteris deltoidea Thelypteridaceae Forb/herb 135 Fern

272 Thespesia grandiflora Malvaceae Tree 126 Dicot

273 Thouinia striata Sapindaceae Tree/shrub 104, 149 Dicot

274 Torralbasia cuneifolia Celastraceae Tree/shrub 93 Dicot

275 Trema micranthum Ulmaceae Tree/shrub 94 Dicot

276 Trichilia pallida Meliaceae Tree/shrub 71 Dicot

277 Trichipteris borinquena Cyatheaceae Shrub 135 Fern

278 Trichipteris procera Cyatheaceae Tree 135 Fern

279 Trichostigma octandrum Phytolaccaceae Shrub/vine 135 Dicot

280 Turpinia occidentalis Staphyleaceae Tree 101 Dicot

281 Urera baccifera Urticaceae Tree/shrub 135 Dicot

282 Xylosma schwaneckeana Flacourtiaceae Vine/tree/shrub 93 Dicot

283 Young secondary forest 98

284 Zanthoxylum flavum Rutaceae Tree/shrub 115 Dicot

285 Zanthoxylum martinicense Rutaceae Tree/shrub 97 Dicot

Table 10—Species, family, growth habit, and source (continued)
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Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

Appendix 3: Elemental Composition by Species in Live 

Vegetation

This appendix contains charts of element and ash concentrations, and carbon to 

nitrogen (C/N) ratios of live vegetation parts by species. Species identification is by 

code number in appendix 2. The charts are intended to provide a visual idea of the 

range of element and ash concentration and C/N values and outlier species. Some 

charts have the Y-axis scale in log scale while others are linear. The charts are 

arranged in alphabetical order by the Y-axis label.
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Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

Appendix 4: Elemental Composition by Species in 

Loose Litter

This appendix contains charts of element and ash concentrations, and carbon to 

nitrogen (C/N) ratios of ground or loose litter (LL) by species. Loose litter is sorted 

by different components. Species identification is by code number in appendix 2. 

The charts are intended to provide a visual idea of the range of element and ash 

concentration and C/N values and outlier species. Some charts have the Y-axis scale 

in log scale while others are linear. The charts are arranged in alphabetical order by 

the Y-axis label.
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Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)
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Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

Appendix 5: Elemental Composition by Species

in Litter Fall

This appendix contains charts of element and ash concentrations, and carbon to 

nitrogen ratios (C/N) of ground or litter fall (LF) by species is sorted by different 

components. Species identification is by code number in appendix 2. The charts are 

intended to provide a visual idea of the range of element and ash concentration and 

C/N values and outlier species. Some charts have the Y-axis scale in log scale while 

others are linear. The charts are arranged in alphabetical order by the Y-axis label.
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Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)
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Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

Appendix 6: Elemental Composition by Soil Order

This appendix contains charts of mean values of element concentrations; cation 

exchange capacity; percentage clay, sand, and silt; loss on ignition; organic matter; 

and pH of soils by soil order. The number in parenthesis is the number of samples. 

The charts are intended to provide a visual idea of the range of element and other 

soil parameters by soil order. The charts are arranged in alphabetical order by the 

Y-axis label.
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Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

Appendix 7: Elemental Composition of Species From 

Contrasting Environments in Puerto Rico

Ernesto Medina

Adjunct Scientist International Institute of Tropical Forestry

Centro de Ecología, Instituto Venezolano de Investigaciones Científicas, Caracas

Introduction

The laboratory analyzed leaf samples obtained from a variety of sites in Puerto 

Rico including karstic areas in Guánica, Caguana, and Monito island; serpentine 

areas in Susúa and Maricao; several forested sites in the Luquillo mountains 

(Sabana, Bisley, El Verde); and mangrove and forested swamp sites in Punta 

Viento (Patillas), Jobos (Salinas), Humacao, and Sabana Seca. Frequently, species 

were repeatedly sampled within the same project on separate dates or in different 

projects. The samples were collected to satisfy descriptive requirements of specific 

projects and researchers, and were not designed to answer explicit questions or to 

test a priori hypotheses on nutrient relations. Therefore, the leaf data set does not 

represent a balanced set of families, species, or sites. In addition, there are no precise indications about leaf age, light exposure, or position in the canopy. I assumed 

that the sample set was constituted by adult leaves, not severely attacked by herbivores, and represented fully functional photosynthetic units. In spite of the obvious 

limitations, the data set is large enough to allow certain questions to be answered:

1. Are there common distribution patterns of ash and mineral elements in 

adult leaves obtained from contrasting ecosystems such as karst, serpentine, saline swamps, or wet volcanic sites?

2. Are families, and or species uniform in their patterns of accumulation of 

ash and mineral elements?

3. Are there significant correlations between element concentrations?

4. How do leaf nutrient ratios such as carbon/nitrogen, C/phosphorus, and N/P 

ratios compare with those reported from other tropical forests?

5. Are there patterns in the ratios of alkaline and earth alkaline metals?

Methods

The leaf samples were analyzed to determine the concentrations of ash, C, N, sulfur 

(S), potassium (K), magnesium (mg), calcium (Ca), aluminum (Al), and iron (Fe). 

However, not all the leaf samples have a complete set of analyses. For the purpose 

of this statistical analysis, all element concentrations are expressed on a molar basis 

(mmol/kg dry weight), so that element comparisons can be made on an atom instead 
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of weight basis. Transformations of the molar units into weight units are simply 

accomplished multiplying the value by the atomic weight of the element considered. 

Analytical methods are described in a separate section of this book.

The distribution of each parameter was determined for the leaf samples 

grouped by families and species, and for each group the sequence of average values 

was established. I did not attempt to establish if this sequence was statistically significant because there were not enough replicates from each family, species, or sites 

to do it properly. However, in each case, I tried to identify emerging ecological or 

phylogenetic patterns that may lead to further indepth research. For each element, 

I listed the five species with highest (H) and lowest (L) values. I did not attempt to 

detect patterns by site of collection, as the description of sites was insufficient for 

that purpose. However, I did select a group of species characteristic of certain forest 

types, with a complete set of analyses for N, S, P, K, Ca, and Mg, to explore potential relationships with elemental composition.

Results

The data set contains 143 species distributed among 57 families (table 11). Most 

species are native to Puerto Rico. Introduced species include trees naturalized 

in Puerto Rico such as Spathodea campanulata, and Albizia spp, cultivated for 

agricultural purposes (Coffea arabica, Citrus sinensis, and Syzygium jambos), and 

planted for forestry use such as Swietenia species and Khaya nyasica. Bambusa

species are planted along many roads of Puerto Rico for border protection. Other 

species were introduced as garden plants that later escaped and naturalized such as 

Kalanchoe pinnata. The data set used for the following analyses is provided in the 

addendum.

Patterns of Element Concentration

Ash concentration—

The percentage of ash per species averages 7.8 with a coefficient of variation of 

nearly 50 percent; the distribution is positively skewed and slightly leptokurtic

(fig. 1). The highest percentage of ash values (20 percent) were recorded for 

Bambusa vulgaris and Citrus sinensis, and the lowest (<3 percent) was measured in 

two species of the Colorado forest in the Luquillo mountains (Cyrilla racemiflora

and Micropholis chrysophylloides).

Potassium concentration—

The K concentration averages 296 μmol/kg and the variation coefficient is larger 

than that of percentage of ash, the highest values correspond to species of the

families Araceae and Urticaceae (>1000 mmol/kg), and the lowest values included 
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Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

Table 11—List of species analyzed in the present report, organized according to families

Family Species Family Species

Apocynaceae Allamanda violacea Euphorbiaceae Alchornea latifolia

Plumeria alba Croton poecilanthus

Plumeria obtusa Ditta myricoides

Plumeria rubra (red)* Gymnanthes lucida

Plumeria rubra (white)* Sapium laurocerasus

Aquifoliaceae Ilex sideroxyloides Fabaceae Albizia lebbeck*

Araceae Dieffenbachia seguine* Albizia procera*

Araliaceae Schefflera morototoni Andira inermis

Arecaceae Prestoea montana Inga laurina

Bignoniaceae Schlegelia brachyantha Inga vera

Spathodea campanulata* Leucaena leucocephala*

Tabebuia haemantha Ormosia krugii

Tabebuia heterophylla Pictetia aculeata

Tabebuia rigida Pterocarpus officinalis

Tabebuia spp. Flacourtiaceae Casearia arborea

Boraginaceae Bourreria succulenta Casearia decandra

Cordia borinquensis Casearia guianensis

Burseraceae Bursera simaruba Casearia sylvestris

Dacryodes excelsa Homalium racemosum

Tetragastris balsamifera Xylosma schwaneckeana

Capparaceae Capparis cynophallophora Heliconiaceae Heliconia caribaea

Cecropiaceae Cecropia schreberiana Hernandiaceae Hernandia sonora*

Celastraceae Cassine xylocarpa Lauraceae Ocotea leucoxylon

Clusiaceae Calophyllum antillanum Ocotea spathulata

Clusia rosea Magnoliaceae Magnolia splendens

Combretaceae Bucida buceras Malpighiaceae Byrsonima lucida

Laguncularia racemosa Byrsonima spicata

Crassulaceae Kalanchoe pinnata* Byrsonima wadsworthii

Cyrillaceae Cyrilla racemiflora Malvaceae Khaya nyasica*

Elaeocarpaceae Sloanea berteriana Thespesia grandiflora

Erythroxylaceae Erythroxylum areolatum Melastomataceae Henriettea squamulosum

Erythroxylum rotundifolium Miconia impetiolaris

Melastomataceae Miconia prasina Nyctaginaceae Pisonia subcordata

Miconia racemosa Ochnaceae Ouratea littoralis

Miconia sp. Orchidaceae Prescotia oligantha

Miconia spp. Phytolaccaceae Phytolacca icosandra

Miconia tetrandra Pinaceae Pinus caribaea

Meliaceae Guarea glabra Piperaceae Piper aduncum

Guarea guidonia Piper glabrescens

Guarea ramiflora Piper hispidum
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Table 11—List of species analyzed in the present report, organized according to families (continued)

Family Species Family Species

Swietenia humilis* Poaceae Bambusa spp.*

Swietenia macrophylla* Bambusa vulgaris*

Swietenia macrophylla xmahagoni* Polygonaceae Coccoloba diversifolia

Swietenia mahagoni* Coccoloba microstachya

Swietenia sp.* Pteridaceae Acrostichum aureum

Trichilia palida Rhamnaceae Colubrina arborescens

Colubrina elliptica

Moraceae Ficus citrifolia Krugiodendron ferreum

Myrsinaceae Ardisia glauciflora Rhizophoraceae Rhizophora mangle

Ardisia solanacea Rubiaceae Coffea arabica*

Myrtaceae Eugenia borinquensis Exostema caribaeum

Eugenia eggersii Faramea occidentalis

Eugenia foetida Guettarda pungens

Eugenia maleolens Guettarda scabra

Eugenia monticola Neolaugeria resinosa

Eugenia rhombea Palicourea riparia

Eugenia stahlii Psychotria berteriana

Gomidesia lindeniana Psychotria maleolens

Myrcia deflexa Psychotria sp.

Myrcia splendens Rutaceae Amyris elemifera

Pimenta racemosa Citrus sinensis*

Syzygium jambos* Sapindaceae Sapindus saponaria

Nyctaginaceae Guapira obtusata Thouinia striata

Pisonia albida

Sapotaceae Manilkara bidentata Families: 57

Micropholis chrysophylloides Species: 143

Staphyleaceae Turpinia occidentalis

Theaceae Ternstroemia stahlii

Theophrastaceae Jacquinia berteroi

Thymelaeaceae Daphnopsis philippiana

Erithalis fruticosa

Ulmaceae Trema micranthum

Urticaceae Urera baccifera

Verbenaceae Avicennia germinans

Petitia domingensis

Viscaceae Phoradendron racemosum

Zygophyllaceae Guaiacum officinale

Guaiacum sanctum

* Indicates introduced species.
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Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

C. sinensis, although this species has one of the highest percentage of ash values 

(fig. 1).

Calcium concentration—

Average calcium concentration is similar to that of K, but the levels of asymmetry, kurtosis, and coefficient of variation are considerably higher (fig. 1). Urera 

baccifera (Urticaceae) and Phoradendron racemosusm (Viscaceae) are within 

a group of species with the highest Ca concentration (>900 mmol/kg). Ardisia 

glauciflora (Myrsinaceae) lies within the high Ca concentration group (>700 

mmol/kg) contrasting with its position regarding K. In the data set, Kalanchoe pinnata (Crassulaceae) appears as an outlier regarding Ca concentration. This family is known for its calciotrophic character, and most of their Ca is in soluble form 

(Kinzel 1983).

Magnesium concentration—

The species of Viscaceae and Urticaceae appear again within the group with 

the highest element concentration as in the case of K and Ca, although they 

do not belong to the group with highest ash concentration. Miconia prasina 

(Melastomataceae), Micropholis chrysophylloides (Sapotaceae), and Cyrilla racemiflora (Cyrillaceae) show very low Mg concentrations (<40 mmol/kg). The distribution of Mg concentrations has similar levels of skewness, kurtosis, and coefficient 

of variation as those of Ca. Among the species analyzed, it is noteworthy that one 

Flacourtiaceae, Casearia sylvestris, appears in the high group, whereas another, 

Xylosma schwaneckeana, has extremely low concentrations of this element. 

Aluminum and Iron concentration—

Distributions of these elements are highly right-skewed, and strongly leptokurtic, 

indicating that a few species present rather high concentrations compared to the 

rest (fig. 2). Average Al concentration is 61 mmol/kg, and the variation coefficient 

amounts to more than 300 percent with high values well above 500 mmol/kg. The 

Miconia species (Melastomataceae) stand out as Al accumulators, as reported for 

several tropical plant communities (Jansen et al. 2002) (fig. 2). 

In the high Al concentration group the Flacourtiaceae, Rubiaceae, and Phytolaccaceae are also included; families known for containing many metal accumulating species. Rubiaceae species are found both in the high (P. maleolens) and low 

(Guettarda scabra) concentration groups.

The Miconia species are again within the Fe concentrating species >30 mmol/

kg), together with Ocotea spathulata (Lauraceae), and Phytolacca icosandra

(Phytolaccaceae). Species of the latter genus have been reported as accumulators of 

heavy metals, mainly manganese (Baker and Brooks 1989, Yuan et al. 2007).
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Figure 1—Distribution of concentrations of percentage of ash and major alkaline and earth alkaline metals. The species list includes the five highest and lowest values recorded in the data set. Statistical calculations obtained using JMP 8.
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Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

Figure 2—Distributions of concentrations of aluminum (Al) and iron (Fe). The species list includes 

the five highest and lowest values recorded in the data set. Statistical calculations obtained using 

JMP 8.
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Nonmetallic elements—

Phosphorus distribution was strongly asymmetric, because a few species have concentrations above the 100 mmol/kg level, resulting therefore in a large coefficient of 

variation (fig. 3). Among these species, Ficus citrifolia (Moraceae) and Kalanchoe 

pinnata (Crassulaceae) have surprisingly high values. The species A. glauciflora 

(Myrsinaceae), Eugenia rhombea (Myrtaceae), Clusia rosea (Clusiaceae), and 

Jacquinia berteroi (Theophrastaceae) are within the low P concentration group.

Distribution of C concentrations was nearly normal, with a small coefficient 

of variation, values ranged with narrow limits between 48 to 35 mol/kg (fig. 3). 

Nitrogen and S were much more variable, with CVs around 45 percent, and levels of 

skewness and kurtosis departing slightly from normality. The high N concentration 

group (>2500 mmol/kg) contains one species of Fabaceae (Leucacena leucocephala) and species with rather fleshy or thin leaves such as Phoradendron racemosum

(Viscaceae), P. icosandra (Phytolaccaceae), and Trema micranthum (Ulmaceae). 

Again C. racemiflora and C. rosea appear within the low concentration group. 

Average concentration of N (1254 mmol/kg) and S (90 mmol/kg) are above the 

average concentrations required for normal, healthy growth in cultivated plants, 

whereas average for P (27 mmol/kg) is about half of that value (see table 4, Epstein 

1965).

Inter-Element Correlations

Pair-wise correlations (Pearson product-moment correlations between each pair

of variables) were highly significant (p < 0.0001) and positive between percentage

of ash and Ca, Al, and Fe, and negative between percentage of ash and C (table 12). 

Other elements highly correlated are Fe and Al; P with K, Ca, and N. Nitrogen was 

significantly correlated with K, P and less so with S.

I applied also a test for nonparametric correlations (Spearman r) to the data set 

and obtained somewhat different results. As before, percentage of ash was highly 

correlated with Ca and C. Other significant correlations were obtained between Fe 

and Al, N with P and K. However, nonparametric coefficients for Ca and P, Ca and 

K, percentage of ash and Fe were not significant in contrast to the Pearson coefficients (table 12).

Element Ratios, Comparisons With Cultivated Plants and Other 

Tropical Forests

Element ratios calculated from the composition of organisms are useful for

estimating functional relationships that relate biological activity with environmental variables (Elser 2006, Elser et al. 2010, Redfield 1958). The study of element 
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Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000) Figure 3—Distribution of concentrations of nonmetallic elements. The species list includes the five highest and lowest values recorded in the data set. Statistical calculations obtained using JMP 8.
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proportions has been the subject of biological stoichiometry, and their variation 

associated with the environment is treated as ecological stoichiometry (Sterner and 

Elser 2002).

For tropical forest the C/N and N/P ratios have been frequently discussed 

within the framework of P limitation (McGroddy et al. 2004). In addition, the ratios 

of alkaline and earth alkaline metals have also implications for both functional 

properties of organisms, and ecological relationships between plants and soils 

(Horak and Kinzel 1971; Kinzel 1989).

Compared to the averages reported for tropical forests (McGroddy et al. 2004) 

the C/N, C/P and N/P ratios are within the range of variation expected (Table 13). 

However, the N/P ratio in the present data set is about 20 percent higher than the 

average for other tropical forests, with a coefficient of variation above 40 percent 

implying that a large number of samples are limited by P based on N/P ratio, probably affecting plant production capacity.

Depicting the sequence of N/P ratios of the species in the data set shows that 

at least 75 of the 120 species in the data set have N/P ratios above the assumed P 

sufficiency range in wild plants (Mc Groddy et al. 2004) (Figure 4).

Table 12—Significant (p < 0.01) pair-wise (Pearson product-moment) and

nonparametric (Spearman r) correlations among elements measured in

leaf samples 

V1 V2 Correlation Signif prob Spearman ρ Prob > |ρ| Count

Al Ash 0.465 <.0001 0.166 0.167 71

C Ash -0.558 <.0001 -0.551 <.0001 58

Ca P 0.337 0.0003 -0.044 0.645 113

Ca Ash 0.666 <.0001 0.706 <.0001 85

Ca K 0.296 0.001 0.211 0.023 116

Fe Al 0.732 <.0001 0.864 <.0001 80

Fe Ash 0.460 <.0001 0.107 0.377 71

K P 0.447 <.0001 0.419 <.0001 124

Mg K 0.373 <.0001 0.328 0.001 106

Mg Ash 0.268 0.019 0.327 0.004 76

N S 0.394 0.002 0.367 0.005 57

N P 0.552 <.0001 0.671 <.0001 119

N K 0.412 <.0001 0.447 <.0001 122

N Ash 0.253 0.011 0.344 <0.001 101

N Mg 0.240 0.015 0.264 0.007 102

Note: Ash in percentage, C in mol/kg, other elements in mmol/kg. In bold correlation coefficients that do not 

agree with each other at the given p level.
71

Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

Table 13—Elemental ratios on a molar basis for the species leaf data set

C/N x 10-3 C/P C/S N/P N/S S/P K/Ca Mg/Ca

Mean 38.6 2279 700 53.3 20.5 4.7 1.53 0.74

Std dev 18.1 1320 941 22.2 32.6 2.4 1.78 0.70

Std err mean 2.4 190 125 2.0 4.3 0.4 0.17 0.07

Median 35.1 1974 501 49.6 13.8 4.2 1.05 0.53

Skewness 1.5 2.3 5.4 1.6 6.2 1.0 5.12 2.32

Kurtosis 1.9 5.9 31.2 5.7 41.3 1.9 36.33 5.84

CV 46.9 57.9 134.4 41.6 159.1 52.2 115.9 94.7

n 59 48 57 119 57 47 116 106

 Mean 35.5 2457 43.4

McGroddy et al. 2004.

Figure 4—Species in the data set ordered according to their nitrogen/phosphorus (N/P) molar ratio.
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International Institute of Tropical Forestry II

I do not have data of C/S and N/S ratios for tropical ecosystems. Based on 

data from the agronomic literature, I estimate that the average C/S ratio indicates 

limitations of this element that may reduce its availability during decomposition of 

leaf litter. The N/S ratios, however, are within the range of sufficiency estimated 

for normal growth in cultivated plants (Epstein 1965) (table 14). The S/P ratio is 

about 10 times higher than the agronomic average, indicating high availability of S 

compared to that of P.

The K/Ca ratio average is very near to the average expected for higher plants 

(≈ 2). However, the variation coefficient is quite large, indicating strong departure 

from the rule of K/Ca >1. Ratios below 1 may indicate the presence of calciotrophic 

plants (sensu Horak and Kinzel 1989), or that the samples were obtained from 

forests on soils with high Ca availability (e.g., calcareous substrates). The potential 

calciotrophs or Ca accumulators are depicted in figure 5.

Mg/Ca ratios are usually around 0.6, and higher values are to be expected in 

areas with elevated Mg availability, such as serpentine derived soils. The data set 

has an average Mg/Ca of 0.7, well within the normal values. However, a number 

of species have markedly higher values indicating the presence of potential Mg 

accumulators (fig. 6).

Species Signatures and Variability Among Species and 

Ecological Groups.

The amount and proportion of mineral elements that constitute a living organism, 

or a given organ, is related to their physiology and biochemistry and is influenced 

by the availability of those elements in the environment. More recently the concept 

of ionome has been introduced and defined “as the mineral nutrient and trace 

Table 14—Average mineral concentrations of mineral 

nutrients in plant shoot dry matter that are sufficient for 

adequate growth (Epstein 1965)

Element Concentrations

mmol/kg mg/kg

Nitrogen 1000 14000

Potassium 250 9970

Calcium 125 5010

Magnesium 80 1944

Phosphorus 60 1858

Sulfur 30 962

Molar ratios 

N/K = 4, N/Ca = 8, N/Mg = 12.5, N/P = 17, N/S = 33, 

S/P = 0.5, K/Ca = 2, Mg/Ca = 0.6.
73

Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

Figure 5—Species in the data set ordered according to their potassium/calcium (K/Ca) molar ratio. The species with ratios <1 are listed.
74

International Institute of Tropical Forestry II

Figure 6—Species in the data set ordered according to their magnesium/calcium (Mg/Ca) molar ratio. The species with ratios

>1 are listed.

element composition of an organism and represents the inorganic component of 

cellular and organismal systems” (Salt et al. 2008). Using elemental composition it 

may be possible to characterize species, identify metabolic syndromes (e.g., cation 

accumulators), and gain insight into the influence of substrate mineral availability 

on the regulation of metabolic pathways of species that grow on them.

In higher plants, the proportion of major elements, (excluding C that constitutes 

about 50 percent of organic matter by weight) have been well studied in cultivated 

species. The sequence of major element concentrations may be used as a guideline 

to assess the interspecific and ecological variability of the data set developed by the 

International Institute of Tropical Forestry Laboratory (see “Addendum”).

As an example of the potential application of element profiles to characterize 

species and ecological sites, I selected a number of species that are major components of some forest types in Puerto Rico and put together in a stack column graph 

the values for the major elements in leaf tissues (fig. 7). Secondary species appear 

to be particularly rich in N, whereas species from wet forests have a tendency to 

lower levels of all the elements considered. Species from karst areas appear subdivided in two groups on the basis of Ca concentration. Mangrove swamp species are 
75

Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

Figure 7—Species element composition “signatures.” Species were separated according to their assumed common 

occurrence: calcareous soils, moist forests, high montane forest in the Luquillo mountain range, and species from coastal 

mangroves.
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International Institute of Tropical Forestry II

heterogeneous regarding element proportions. In this case, the addition of Na would 

have separated this group. This representation also allows detection of departure 

from the general pattern of concentration sequence of plant nutrients (table 14). 

From the group of species selected, some concentrations stand out: the low concentration of S in L. racemosa and P. officinalis in spite of their occurrence in S rich 

soils (mangrove swamps); the small porportion of Mg in M. prasina and D. excelsa; 

and the high concentration of Mg relative to Ca in A. germinans.

Concluding Remarks

The analysis of the leaf data set compiled by the International Institute of Tropical 

Forestry laboratory during the 1980–2000 period has given some relevant results 

regarding:

1. Average and range of variation in the mineral element composition of

species belonging to a wide array of families and from different ecological 

sites.

2. Confirmation of P limitation in the majority of samples analyzed based on 

the N/P ratios

3. Determination of the distribution of C/S and N/S ratios, parameters that 

have not been properly evaluated in the current ecological literature on 

nutrient cycling. The present data set for example, detects comparatively 

high levels of S in the majority of species analyzed.

4. Identification of Al and Fe accumulators within certain families such as the 

Melastomataceae, Flacourtiaceae, and Phytolaccaceae.

5. Detection of species departing strongly from the usual K/Ca and Mg/Ca 

ratios, revealing the probable existence of calciotrophic species. 

Literature Cited

Baker, A.J.M.; Brooks, R.R. 1989. Terrestrial higher plants which hyper 

accumulate metallic elements-a review of their distribution, ecology and 

phytochemistry. Biorecovery. 1: 81–126.

Elser, J. 2006. Biological stoichiometry: a chemical bridge between ecosystem 

ecology and evolutionary biology. American Naturalist. 168: S25-S35. 

Elser, J.J.; Fagan, W.F.; Kerkhoff, A.J.; Swenson, N.G.; Enquist, B.J. 2010.

Biological stoichiometry of plant production: metabolism, scaling and ecological 

response to global change. New Phytologist. 186: 593–608. 

Epstein, E. 1965. Mineral metabolism. In: Bonner, J.; Varner, J.E., eds. Plant 

biochemistry. London: Academic Press: 438–466. 
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Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

Han, W.; Fang, J.; Guo, D.; Zhang, Y. 2005. Leaf nitrogen and phosphorus 

stoichiometry across 753 terrestrial plant species in China. New Phytologist. 168: 

377–385.

Horak, O.; Kinzel, H. 1971. Typen des Mineralstoffwechsels bei den höheren 

Pflanzen. Österreichischer Botanische Zeitung. 111: 475–495.

Jansen S.; Watanabe, T.; Smets, E. 2002. Aluminium accumulation in leaves of 

127 species in Melastomataceae, with comments on the order Myrtales. Annals 

of Botany. 9: 53–64.

Kinzel, H. 1983. Influence of limestone, silicates and soil pH on vegetation. In: 

Lange, O.L.; Nobel, P.P.; Osmond, P.C.B.; Ziegler, P.H., eds. Physiological plant 

ecology III. Responses to chemical and biological Berlin, Environment. Berlin: 

Springer Verlag: 201–244.

Kinzel, H. 1989. Calcium in the vacuoles and cell walls of plant tissue: forms of 

deposition and their physiological and ecological significance. Flora. 182: 99–

125.

McGroddy, M.E.; Daufresne, T.; Hedin, L.O. 2004. Scaling of C:N:P 

stoichiometry in forests worldwide: implications of terrestrial Redfield-type 

ratios. Ecology. 85: 2390–2401.

Min, Y.; Tie, boqing: Meizhen, T.; Aoyama, I. 2007. Accumulation and uptake of 

manganese in a hyperaccumulator Phytolacca americana. Minerals Engineering. 

20: 188–190. 

Redfield, A.C. 1958. The biological control of chemical factors in the environment. 

American Scientist. 46: 205–221.

Salt, D.E.; Baxter, I.; Lahner, B. 2008. Ionomics and the study of the plant 

Ionome. Annual Review of Plant Biology. 59: 709–733.

Sterner R.W.; Elser, J.J. 2002. Ecological stoichiometry. New Jersey: Princeton 

University Press. 
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Addendum

Data set used for the analyses developed in the text. Ash in percentage, carbon in mol/kg, the other elements in mmol/kg

Species Ash C K Ca Mg Al Fe P C N S 

Acrostichum aureum 6.07 43.4 529 67 84 1 0.4 34.6 36.17 1664

Alchornea latifolia 226 177 111 7.8 2.9 27.2 1379

Albizia lebbeck 2371

Albizia procera 7.11 51.6 43.00 1564 69

Allamanda violacea 10.09 189 499 47.8 1171

Amyris elemifera 10.7 51.9 351 701 104 2.6 1.1 18.3 43.25 1393 100

Andira inermis 4.48 189 67 106 42.1 1650

Ardisia glauciflora 10.41 107 786 38 1.5 0.9 8.7 379

Ardisia solanacea 13.5 48.9 40.75 1279 59

Avicennia germinans 10 45 320 105 353 3.3 1.4 24.9 37.50 1536 100

Bambusa spp. 6.87 52.7 309 204 83 35 9.8 29.2 43.92 1593 122

Bambusa vulgaris 21.42 240 27.8 1593

Bourreria succulenta 10.56 49 485 583 186 1.7 0.7 15.4 40.83 1057 56

Bucida buceras 13.71 45 267 470 83 17.8 37.50 893 59

Bursera simaruba 6.11 53.2 277 218 106 2.25 1 25.4 44.33 1157 90

Byrsonima lucida 3.92 155 193 192 1.35 1.2 15 943

Byrsonima spicata 3.86 56.8 174 140 150 3.8 1.7 17.3 47.33 1779 137

Byrsonima wadsworthii 3.4 158 186 111 8.5 3.4 22 1286

Calophyllum antillanum 3.85 54.8 222 136 135 1.8 0.9 17 45.67 714 62

Capparis cynophallophora 623 16.5 1457

Casearia arborea 5.88 292 91 116 26.8 1143

Casearia decandra 6.63 52 73 315 303 2.5 2.5 25.1 43.33 1807 69

Casearia guianensis 7.68 50 41.67 1714 59

Casearia sylvestris 15.2 341 163 555 2.1 1 41.3 1257

Cassine xylocarpa 8.08 228 279 142 1.15 1.25 10 579

Cecropia schreberiana 9.26 46.5 423 259 167 24.733 14.88 35.2 38.75 1657 97

Citrus sinensis 20.68 58 415 807

Clusia rosea 5.87 54.8 165 240 144 1.4 6.5 7.8 45.67 471 56

Coccoloba diversifolia 10.68 50.9 278 710 126 2.2 1 17.5 42.42 1129 103

Coccoloba microstachya 4.73 52.5 178 273 96 1.25 0.8 15.1 43.75 600 87

Coffea arabica 10.34 51.1 466 249 160 3.5 2.15 35 42.58 1886 153

Colubrina arborescens 497 493 136 18.6 1029

Colubrina elliptica 608 27.2 1621

Cordia borinquensis 346 97 173 28.4 1386

Croton poecilanthus 5.64 315 168 20.1 2.6 32.8 729

Cyrilla racemiflora 2.79 239 41 28 3.7 1.7 21.4 493

Dacryodes excelsa 6.89 46.9 116 112 43 25.6 8.1 16.8 39.08 936 78

Daphnopsis philippiana 3.09 177 96 62 6.4 1.5 18.3 886
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Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

Species Ash C K Ca Mg Al Fe P C N S 

Dieffenbachia seguine 11.36 45.1 1093 285 131 3.05 1.6 41 37.58 1393 56

Ditta myricoides 183 141 182 5.2 2.5 25.5

Erithalis fruticosa 5.02 56.8 215 245 74 1.3 0.6 10.8 47.33 636 69

Erythroxylum areolatum 363 462 111 33 1386

Erythroxylum rotundifolium 250 359 55 11.1 1179

Eugenia borinquensis 2.88 227 74 66 6.7 1.3 17.8 586

Eugenia eggersii 5.25 263 206 126 8.1 2.5 21 971

Eugenia foetida 285 342 154 12.2 1014

Eugenia maleolens 285 342 154 12.2

Eugenia monticola 4.58 53.7 44.75 1057 50

Eugenia rhombea 317 8.4 693

Eugenia stahlii 7.34 142 14.9 543

Exostema caribaeum 3.91 56.1 232 220 101 1.967 1.63 17.7 46.75 1307 72

Faramea occidentalis 8.6 48.7 40.58 1293 81

Ficus citrifolia 13.98 317 505 118 0.7 0.7 148 2450

Gomidesia lindeniana 255 180 305 1.3 1.5 9.7 543

Guaiacum officinale 329 14.2 1293

Guaiacum sanctum 380 434 99 13.1 1229

Guapira obtusata 4.94 350 85 257 0.7 1.1 47.1 1907

Guarea glabra 6.58 389 245 57.2 2200

Guarea guidonia 5.88 50.5 232 236 117 16 7.8 41 42.08 2193 159

Guarea ramiflora 44.4 37.00 1729

Guettarda pungens 4.05 176 249 54 1.7 2.2 9.6 550

Guettarda scabra 4.85 220 89 138 0.7 0.95 17 707

Gymnanthes lucida 11.2 48.4 389 682 103 1.4 0.6 16.9 40.33 1229 59

Heliconia caribaea 5.1 48.5 582 37 76 2.233 0.67 11 40.42 529 75

Henriettea squamulosum 5.39 200 326 97 149.4 3.35 16.3 764

Hernandia sonora 13.09 82 286 200 24.1 1286

Homalium racemosum 6.05 70 304 213 2.3 1.1 14.9 871

Ilex sideroxyloides 3.25 263 119 89 8.9 1.8 24.9 721

Inga laurina 4.37 53.9 179 130 52 29.9 44.92 1757 97

Inga vera 73 257 131 27.2 1500

Jacquinia berteroi 9.02 54.2 167 359 80 1.5 0.6 6.2 45.17 600 81

Kalanchoe pinnata 530 1575 166 114 629

Khaya nyasica 229 140 93 29 1214

Krugiodendron ferreum 377 471 129 15.6 1671

Laguncularia racemosa 16.93 43.5 339 555 247 9.1 7.1 22 36.25 893 9

Leucaena leucocephala 9.27 51.1 42.58 2936 137

Magnolia splendens 5.93 57.2 301 171 98 5.2 2 23.9 47.67 1050 69

Manilkara bidentata 5.69 49.8 173 179 110 10.4 4.8 14.4 41.50 921 137
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International Institute of Tropical Forestry II

Species Ash C K Ca Mg Al Fe P C N S 

Miconia impetiolaris 15.98 46.3 95 457 93 748.8 100 27.2 38.58 1371 106

Miconia prasina 7.9 49.3 122 151 33 538.975 2.4 15.3 41.08 836 66

Miconia racemosa 11.13 47.9 106 544 77 492.167 32.63 18.4 39.92 1136 156

Miconia sp. 17.35 81 694 128 1251.95 61.3 15.4

Miconia spp. 3.76 49.8 91 119 52 245.3 8.03 6.4 41.50 457 37

Miconia tetrandra 9.67 102 18.6 1043

Micropholis chrysophylloides 2.68 150 107 32 3.7 1.8 12.6 721

Myrcia deflexa 166 171 108 15.6 943

Myrcia splendens 8.07 51.5 110 19.5 42.92 936 137

Neolaugeria resinosa 5.06 330 203 557 2.1 1.27 12.5 664

Ocotea leucoxylon 6.84 53.6 196 186 89 14.8 2.5 30.6 44.67 1543 62

Ocotea spathulata 477 160 21.5 33.7 50.4

Ormosia krugii 114 52 37 24.4 1514

Ouratea littoralis 4.71 143 298 118 1.5 1.2 14.9 764

Palicourea riparia 9.13 351 216 173 250.6 3.9 23.8 1779

Petitia domingensis 4.76 158 149 163 1.7 1.5 26.6 964

Phoradendron racemosum 863 651 465 2886

Phytolacca icosandra 358 163 329 85 29.6 77.8 2593

Pictetia aculeata 9.19 51.8 302 516 103 1.85 1.2 22.1 43.17 1393 84

Pimenta racemosa 7.92 52.6 43.83 643 50

Pinus caribaea 2.93 51 25.2 629

Piper aduncum 14.62 48.5 380 348 86 26.3 11 29.5 40.42 2300 156

Piper glabrescens 619 265 146 7 3.2 33.5 1521

Piper hispidium 686 302 170 5.1 3.1 52.7 1650

Pisonia albida 380 362 345 36.3 1521

Pisonia subcordata 4.66 330 71 251 1.1 0.8 25.2 1071

Plumeria alba 8.92 256 508 49.7 1250

Plumeria obtusa 6.1 207 317 10.3 957

Plumeria rubra (red) 10 673 531 9.7 1629

Plumeria rubra (white) 10.25 839 328 88.3 1914

Prescotia oligantha 469 276 99 0.1 49.1 1286

Prestoea montana 7.91 45 161 94 119 18.4 5.97 27.2 37.50 1150 197

Psychotria berteriana 9.75 46.3 491 288 220 66.4 5.43 29.4 38.58 1936 56

Psychotria maleolens 12.1 42 123 294 213 525 5.4 14.3 35.00 886 34

Psychotria sp. 12.67 532 39.1 1686

Pterocarpus officinalis 7.29 49.8 249 391 141 1.8 1.13 24.4 41.50 1521 6

Rhizophora mangle 11.35 45.9 172 310 206 1.233 0.6 13.7 38.25 764 119

Sapindus saponaria 3.69 55 66 177 32 17.6 8.6 22.6 45.83 2586 159

Sapium laurocerasus 7.22 51.3 525 189 186 10.1 6.6 36.5 42.75 1186 156

Schefflera morototoni 6.86 51.2 246 169 142 29.1 42.67 1321 109
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Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

Species Ash C K Ca Mg Al Fe P C N S 

Schlegelia brachyantha 0.7

Sloanea berteriana 3.81 46.6 210 139 65 3.3 1.3 20.1 38.83 986 97

Spathodea campanulata 6.7 51.1 42.58 1900 87

Swietenia humilis 283 445 110 4.25 26.7 1236

Swietenia macrophylla 7.01 52.7 242 360 77 11.2 25.2 24.9 43.92 1136 122

Swietenia macrophylla xmahagoni 295 338 81 5.8 24.1 1179

Swietenia mahagoni 6.51 203 225 85 1.9 1.2 21.8 864

Swietenia sp. 280 291 140 2.725 25.6 1307

Syzygium jambos 4.72 53.8 145 162 140 4.9 1.2 20.4 44.83 964 72

Tabebuia haemantha 4.67 297 82 83 0.5 0.75 19.1 764

Tabebuia heterophylla 4.53 51.5 275 133 105 0.2 0 28.3 42.92 1007 143

Tabebuia rigida 4.03 46.4 202 64 109 14.65 6.3 22.6 38.67 914 34

Tabebuia spp. 452 506 640 1250

Ternstroemia stahlii 5.85 230 176 125 1.667 0.97 15.9 621

Tetragastris balsamifera 184 142 68 21.7 1000

Thespesia grandiflora 9.34 53.3 44.42 1579 84

Thouinia striata 279 155 48 26.8 1329

Trema micrantha 9.37 459 69.1 2593

Trichilia palida 9.71 51.9 43.25 1014 90

Turpinia occidentalis 5.41 341 283 27.4 1043

Urera baccifera 1050 931 444 2.6 1.4 93.5 1921

Xylosma schwaneckeana 290 458 9 346.2 27.4 52.3
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International Institute of Tropical Forestry II

Species C/N C/P N/P C/S N/S S/P K/Ca Ca/g Mg/Ca

Acrostichum aureum 22 1045 48 7.90 0.80 1.25

Alchornea latifolia 51 1.28 1.59 0.63

Albizia lebbeck

Albizia procera 27 627 23

Allamanda violacea 25 0.38

Amyris elemifera 31 2363 76 433 14 5.45 0.50 6.74 0.15

Andira inermis 39 2.82 0.63 1.58

Ardisia glauciflora 44 0.14 20.68 0.05

Ardisia solanacea 32 688 22

Avicennia germinans 24 1506 62 376 15 4.01 3.05 0.30 3.36

Bambusa spp. 28 1504 55 361 13 4.17 1.51 2.46 0.41

Bambusa vulgaris 57

Bourreria succulenta 39 2652 69 727 19 3.65 0.83 3.13 0.32

Bucida buceras 42 2107 50 633 15 3.33 0.57 5.66 0.18

Bursera simaruba 38 1745 46 490 13 3.56 1.27 2.06 0.49

Byrsonima lucida 63 0.80 1.01 0.99

Byrsonima spicata 27 2736 103 345 13 7.93 1.24 0.93 1.07

Byrsonima wadsworthii 58 0.85 1.68 0.60

Calophyllum antillanum 64 2686 42 732 11 3.67 1.63 1.01 0.99

Capparis cynophallophora 88

Casearia arborea 43 3.21 0.78 1.27

Casearia decandra 24 1726 72 631 26 2.73 0.23 1.04 0.96

Casearia guianensis 24 703 29

Casearia sylvestris 30 2.09 0.29 3.40

Cassine xylocarpa 58 0.82 1.96 0.51

Cecropia schreberiana 23 1101 47 401 17 2.75 1.63 1.55 0.64

Citrus sinensis 0.14

Clusia rosea 97 5855 60 813 8 7.20 0.69 1.67 0.60

Coccoloba diversifolia 38 2424 64 412 11 5.88 0.39 5.63 0.18

Coccoloba microstachya 73 2897 40 501 7 5.78 0.65 2.84 0.35

Coffea arabica 23 1217 54 279 12 4.37 1.87 1.56 0.64

Colubrina arborescens 55 1.01 3.63 0.28

Colubrina elliptica 60

Cordia borinquensis 49 3.57 0.56 1.78

Croton poecilanthus 22 1.88

Cyrilla racemiflora 23 5.83 1.46 0.68

Dacryodes excelsa 42 2326 56 501 12 4.64 1.04 2.60 0.38

Daphnopsis philippiana 48 1.84 1.55 0.65

Dieffenbachia seguine 27 917 34 669 25 1.37 3.84 2.18 0.46

Ditta myricoides 1.30 0.77 1.29
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Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

Species C/N C/P N/P C/S N/S S/P K/Ca Ca/g Mg/Ca

Erithalis fruticosa 74 4383 59 690 9 6.35 0.88 3.31 0.30

Erythroxylum areolatum 42 0.79 4.16 0.24

Erythroxylum rotundifolium 106 0.70 6.53 0.15

Eugenia borinquensis 33 3.07 1.12 0.89

Eugenia eggersii 46 1.28 1.63 0.61

Eugenia foetida 83 0.83 2.22 0.45

Eugenia maleolens 0.83 2.22 0.45

Eugenia monticola 42 897 21

Eugenia rhombea 82

Eugenia stahlii 36

Exostema caribaeum 36 2641 74 652 18 4.05 1.05 2.18 0.46

Faramea occidentalis 31 500 16

Ficus citrifolia 0 0.63 4.28 0.23

Gomidesia lindeniana 0 1.42 0.59 1.69

Guaiacum officinale 91

Guaiacum sanctum 94 0.88 4.38 0.23

Guapira obtusata 40 4.12 0.33 3.02

Guarea glabra 38 1.59

Guarea guidonia 19 1026 53 265 14 3.88 0.98 2.02 0.50

Guarea ramiflora 21

Guettarda pungens 57 0.71 4.61 0.22

Guettarda scabra 42 2.47 0.64 1.55

Gymnanthes lucida 33 2387 73 681 21 3.51 0.57 6.62 0.15

Heliconia caribaea 76 3674 48 540 7 6.81 15.73 0.49 2.05

Henriettea squamulosum 47 0.61 3.36 0.30

Hernandia sonora 53 0.29 1.43 0.70

Homalium racemosum 58 0.23 1.43 0.70

Ilex sideroxyloides 29 2.21 1.34 0.75

Inga laurina 26 1502 59 465 18 3.23 1.38 2.50 0.40

Inga vera 55 0.28 1.96 0.51

Jacquinia berteroi 75 7285 97 557 7 13.08 0.47 4.49 0.22

Kalanchoe pinnata 0 0.34 9.49 0.11

Khaya nyasica 42 1.64 1.51 0.66

Krugiodendron ferreum 107 0.80 3.65 0.27

Laguncularia racemosa 41 1648 41 3874 95 0.43 0.61 2.25 0.45

Leucaena leucocephala 15 310 21

Magnolia splendens 45 1994 44 695 15 2.87 1.76 1.74 0.57

Manilkara bidentata 45 2882 64 302 7 9.53 0.97 1.63 0.61

Miconia impetiolaris 28 1419 50 364 13 3.90 0.21 4.91 0.20

Miconia prasina 49 2685 55 627 13 4.28 0.81 4.58 0.22
84

International Institute of Tropical Forestry II

Species C/N C/P N/P C/S N/S S/P K/Ca Ca/g Mg/Ca

Miconia racemosa 35 2169 62 256 7 8.48 0.19 7.06 0.14

Miconia sp. 0.12 5.42 0.18

Miconia spp. 91 6484 71 1109 12 5.85 0.76 2.29 0.44

Miconia tetrandra 56

Micropholis chrysophylloides 57 1.40 3.34 0.30

Myrcia deflexa 60 0.97 1.58 0.63

Myrcia splendens 46 2201 48 313 7 7.04

Neolaugeria resinosa 0 1.63 0.36 2.74

Ocotea leucoxylon 29 1460 50 716 25 2.04 1.05 2.09 0.48

Ocotea spathulata 2.98

Ormosia krugii 62 2.19 1.41 0.71

Ouratea littoralis 51 0.48 2.53 0.40

Palicourea riparia 75 1.63 1.25 0.80

Petitia domingensis 36 1.06 0.91 1.09

Phoradendron racemosum 1.33 1.40 0.71

Phytolacca icosandra 33 2.20 0.50 2.02

Pictetia aculeata 31 1953 63 513 17 3.81 0.59 5.01 0.20

Pimenta racemosa 68 878 13

Pinus caribaea 25

Piper aduncum 18 1370 78 259 15 5.29 1.09 4.05 0.25

Piper glabrescens 45 2.34 1.82 0.55

Piper hispidium 31 2.27 1.78 0.56

Pisonia albida 42 1.05 1.05 0.95

Pisonia subcordata 43 4.65 0.28 3.54

Plumeria alba 25 0.50

Plumeria obtusa 93 0.65

Plumeria rubra (red) 168 1.27

Plumeria rubra (white) 22 2.56

Prescotia oligantha 26 1.70 2.79 0.36

Prestoea montana 33 1379 42 191 6 7.22 1.71 0.79 1.27

Psychotria berteriana 20 1312 66 687 34 1.91 1.70 1.31 0.76

Psychotria maleolens 40 2448 62 1020 26 2.40 0.42 1.38 0.72

Psychotria sp. 43

Pterocarpus officinalis 27 1701 62 6652 244 0.26 0.64 2.77 0.36

Rhizophora mangle 50 2792 56 323 6 8.65 0.55 1.50 0.66

Sapindus saponaria 18 2028 114 288 16 7.04 0.37 5.53 0.18

Sapium laurocerasus 36 1171 32 274 8 4.27 2.78 1.02 0.98

Schefflera morototoni 32 1466 45 391 12 3.75 1.46 1.19 0.84

Schlegelia brachyantha

Sloanea berteriana 39 1932 49 402 10 4.81 1.51 2.14 0.47
85

Chemical and Physical Analyses of Selected Plants and Soils From Puerto Rico (1981—2000)

Species C/N C/P N/P C/S N/S S/P K/Ca Ca/g Mg/Ca

Spathodea campanulata 22 488 22

Swietenia humilis 46 0.64 4.05 0.25

Swietenia macrophylla 39 1764 46 361 9 4.89 0.67 4.68 0.21

Swietenia macrophylla xmahagoni 49 0.87 4.17 0.24

Swietenia mahagoni 40 0.90 2.65 0.38

Swietenia sp. 51 0.96 2.08 0.48

Syzygium jambos 46 2198 47 625 13 3.52 0.90 1.16 0.86

Tabebuia haemantha 40 3.62 0.99 1.01

Tabebuia heterophylla 43 250 0 299 7 0.83 2.07 1.27 0.79

Tabebuia rigida 42 1711 40 1127 27 1.52 3.16 0.59 1.70

Tabebuia spp. 0.89 0.79 1.26

Ternstroemia stahlii 39 1.31 1.41 0.71

Tetragastris balsamifera 46 1.30 2.09 0.48

Thespesia grandiflora 28 527 19

Thouinia striata 50 1.80 3.23 0.31

Trema micranthum 38

Trichilia palida 43 478 11

Turpinia occidentalis 38 1.20

Urera baccifera 21 1.13 2.10 0.48

Xylosma schwaneckeana 0.63 50.89 0.02
Pacific Northwest Research Station

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Mailing address Publications Distribution

	 	 Pacific Northwest Research Station

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Official Business

Penalty for Private Use, $300
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

17

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Acrocomia media Aluminum (mg/kg) II1F 165 7 2 165 170 160 128 Mfs Histosol Alluvial deposits pulp 1

II1F 14 18 2 14 26 1 128 Mfs Histosol Alluvial deposits 1

Ash (mg/g) II1F 5.45 1 128 Mfs Histosol Alluvial deposits pulp 1

II1F 1.95 1 128 Mfs Histosol Alluvial deposits 1

Calcium (mg/g) II1F 3.42 0.05 2 3.42 3.46 3.38 128 Mfs Histosol Alluvial deposits pulp 1

II1F 1.18 0.01 2 1.18 1.18 1.17 128 Mfs Histosol Alluvial deposits 1

Iron (mg/g) II1F 0.14 0.17 2 0.14 0.22 0.05 128 Mfs Histosol Alluvial deposits pulp 1

Magnesium (mg/g) II1F 2.23 0.02 2 2.23 2.25 2.22 128 Mfs Histosol Alluvial deposits pulp 1

II1F 1.79 0.06 2 1.79 1.83 1.74 128 Mfs Histosol Alluvial deposits 1

Manganese (mg/kg) II1F 14 1 2 14 14 13 128 Mfs Histosol Alluvial deposits pulp 1

II1F 15 3 2 15 17 13 128 Mfs Histosol Alluvial deposits 1

Nitrogen (%) II1F 0.74 0.02 2 0.74 0.75 0.72 128 Mfs Histosol Alluvial deposits pulp 1

II1F 1.80 0.03 2 1.80 1.82 1.78 128 Mfs Histosol Alluvial deposits 1

Phosphorus (mg/g) II1F 0.47 0.01 2 0.47 0.48 0.46 128 Mfs Histosol Alluvial deposits pulp 1

II1F 3.81 0.28 2 3.81 4.01 3.61 128 Mfs Histosol Alluvial deposits 1

Potassium (mg/g) II1F 19.81 0.47 2 19.81 20.14 19.48 128 Mfs Histosol Alluvial deposits pulp 1

II1F 5.26 0.36 2 5.26 5.51 5.00 128 Mfs Histosol Alluvial deposits 1

Wood density (g/cc) II1C 0.99 0.03 3 0.98 1.02 0.96 116 1

Acrostichum aureum Aluminum (mg/kg) II6A 28 1 2 28 29 26 78 Mfs Mangle Swamp 2

Ash (%) II6A 6.07 0.04 2 6.07 6.10 6.04 78 Mfs Mangle Swamp 2

C/N II6A 34 0 2 34 35 34 78 Mfs Mangle Swamp 2

Calcium (mg/g) II6A 1.84 0.66 17 1.82 3.16 0.78 77 Mfs Swamp Swamp 2

II6A 3.49 0.02 2 3.49 3.50 3.48 78 Mfs Mangle Swamp 2

Carbon (%) II6A 43 0 2 43 43 43 78 Mfs Mangle Swamp 2

Iron (mg/kg) II6A 20 20 2 20 30 10 78 Mfs Mangle Swamp 2

Magnesium (mg/g) II6A 2.12 0.56 17 2.14 3.08 1.18 77 Mfs Mangle Swamp 2

II6A 1.97 0.03 2 1.97 1.99 1.95 78 Mfs Mangle Swamp 2

Manganese (mg/kg) II6A 14 1 2 10 20 10 78 Mfs Mangle Swamp 2
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

18

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Nitrogen (%) II6A 1.46 0.28 17 1.41 2.25 0.90 77 Mfs Mangle Swamp 2

II6A 4.26 2.11 17 3.82 9.93 2.00 77 Mfs Mangle Swamp 2

II6A 1.27 0.02 2 1.27 1.28 1.26 78 Mfs Mangle Swamp 2

Phosphorus (mg/g) II6A 1.36 0.41 17 1.45 2.09 0.47 77 Mfs Mangle Swamp 2

II6A 0.78 0.01 2 0.78 0.79 0.78 78 Mfs Mangle Swamp 2

Potassium (mg/g) II6A 17.26 2.13 17 17.09 21.46 14.18 77 Mfs Mangle Swamp 2

II6A 24.15 0.15 2 24.15 24.25 24.05 78 Mfs Mangle Swamp 2

Sodium (mg/g) II6A 4.59 0.73 2 4.59 5.11 4.07 78 Mfs Mangle Swamp 2

Adiantum spp. Aluminum (mg/g) II6 0.59 0.46 10 0.42 1.79 0.23 135 Lmrf Ultisols Tuffac. sandstone 5

Calcium (mg/g) II6 4.15 0.65 10 4.35 4.96 2.91 135 Lmrf Ultisols Tuffac. sandstone 5

Iron (mg/g) II6 0.46 0.37 10 0.31 1.38 0.17 135 Lmrf Ultisols Tuffac. sandstone 5

Magnesium (mg/g) II6 3.48 0.28 5 3.55 3.78 3.07 135 Lmrf Ultisols Tuffac. sandstone 5

Manganese (mg/g) II6 3.60 0.32 10 3.67 4.17 3.07 135 Lmrf Ultisols Tuffac. sandstone 5

Nitrogen (%) II6 1.52 0.18 5 1.58 1.69 1.24 135 Lmrf Ultisols Tuffac. sandstone 5

Phosphorus (mg/g) II6 1.03 0.23 10 0.98 1.50 0.69 135 Lmrf Ultisols Tuffac. sandstone 5

Potassium (mg/g) II6 15.96 4.44 10 15.47 22.51 10.19 135 Lmrf Ultisols Tuffac. sandstone 5

Adiantum latifolium Aluminum (mg/g) II6 0.65 0.46 4 0.66 1.06 0.22 135 Lmrf Ultisols Tuffac. sandstone 3

C/N II6 23 2 3 23 24 21 135 Lmrf Ultisols Tuffac. sandstone 3

Calcium (mg/g) II6 3.61 1.62 4 3.73 5.02 1.98 135 Lmrf Ultisols Tuffac. sandstone 3

Carbon (%) II6 36 4 3 34 41 34 135 Lmrf Ultisols Tuffac. sandstone 3

Iron (mg/g) II6 0.38 0.11 4 0.40 0.48 0.25 135 Lmrf Ultisols Tuffac. sandstone 3

Magnesium (mg/g) II6 2.61 0.12 4 2.57 2.79 2.53 135 Lmrf Ultisols Tuffac. sandstone 3

Manganese (mg/g) II6 0.14 0.09 4 0.14 0.21 0.04 135 Lmrf Ultisols Tuffac. sandstone 3

Nitrogen (%) II6 1.51 0.23 5 1.43 1.90 1.34 135 Lmrf Ultisols Tuffac. sandstone 3

Phosphorus (mg/g) II6 0.71 0.15 4 0.72 0.85 0.56 135 Lmrf Ultisols Tuffac. sandstone 3

Potassium (mg/g) II6 15.12 4.00 4 16.50 18.21 9.25 135 Lmrf Ultisols Tuffac. sandstone 3

Sulfur (%) II6 0.48 0.14 3 0.53 0.58 0.33 135 Lmrf Ultisols Tuffac. sandstone 3
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

19

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Adiantum pyramidale Aluminum (mg/g) II6 0.49 0.41 2 0.49 0.77 0.20 135 Lmrf Ultisols Tuffac. sandstone 4

C/N II6 19 2 4 18 21 17 135 Lmrf Ultisols Tuffac. sandstone 4

Calcium (mg/g) II6 5.30 0.26 2 5.30 5.48 5.12 135 Lmrf Ultisols Tuffac. sandstone 4

Carbon (%) II6 38 1 4 39 39 37 135 Lmrf Ultisols Tuffac. sandstone 4

Iron (mg/g) II6 0.52 0.46 2 0.52 0.85 0.20 135 Lmrf Ultisols Tuffac. sandstone 4

Magnesium (mg/g) II6 4.37 0.20 2 4.37 4.51 4.23 135 Lmrf Ultisols Tuffac. sandstone 4

Manganese (mg/g) II6 0.12 0.03 2 0.12 0.14 0.10 135 Lmrf Ultisols Tuffac. sandstone 4

Nitrogen (%) II6 2.03 0.19 6 2.12 2.18 1.73 135 Lmrf Ultisols Tuffac. sandstone 4

Phosphorus (mg/g) II6 1.31 0.51 2 1.31 1.67 0.94 135 Lmrf Ultisols Tuffac. sandstone 4

Potassium (mg/g) II6 17.51 5.56 2 17.51 21.41 13.58 135 Lmrf Ultisols Tuffac. sandstone 4

Sulfur (%) II6 0.35 0.04 4 0.33 0.40 0.32 135 Lmrf Ultisols Tuffac. sandstone 4

Albizia lebbeck Nitrogen (%) II1Aib 1.93 1 96 Smf Inceptisols Alluvial deposits Lower canopy 6

II1Aib 3.27 1 96 Smf Inceptisols Alluvial deposits Mid canopy 6

II1Aib 4.57 1 96 Smf Inceptisols Alluvial deposits Upper canopy 6

II1Aiii 3.51 0.43 17 3.35 4.30 3.05 96 Smf Inceptisols Alluvial deposits 6

II1B 1.47 0.41 10 1.61 1.95 0.79 96 Smf Inceptisols Alluvial deposits >50mm 6

II1B 1.23 0.34 7 1.28 1.71 0.77 96 Smf Inceptisols Alluvial deposits 25-50mm 6

II1B 1.79 0.57 12 1.75 2.68 1.04 96 Smf Inceptisols Alluvial deposits l0mm 6

II1B 1.11 0.46 6 1.07 1.77 0.55 96 Smf Inceptisols Alluvial deposits >50mm 6

II1Ciii 2.59 0.01 2 2.59 2.60 2.59 96 Smf Inceptisols Alluvial deposits <10mm 6

II1Ciii 3.32 0.09 2 3.32 3.39 3.26 96 Smf Inceptisols Alluvial deposits l0-50mm 6

II1Ciii 3.08 0.25 2 3.08 3.25 2.90 96 Smf Inceptisols Alluvial deposits >50mm 6

II1E 1.20 0.32 9 1.15 1.65 0.76 96 Smf Inceptisols Alluvial deposits 10-25mm 6

II1E 1.64 0.19 8 1.64 1.93 1.33 96 Smf Inceptisols Alluvial deposits 2-10mm 6

II1E 1.08 0.24 7 1.15 1.38 0.77 96 Smf Inceptisols Alluvial deposits 25-50mm 6

II1E 2.07 0.33 9 2.02 2.63 1.68 96 Smf Inceptisols Alluvial deposits <2mm 6
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

20

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Albizia procera Ash (%) II1A 7.11 1.43 5 6.63 9.58 5.86 126 Ultisols Siltstone,sandstone Urban forest, RP 7

II1C 2.01 0.55 5 1.75 2.91 1.51 126 Ultisols Siltstone,sandstone Urban forest, RP 7

II1Ci 10.53 2.31 6 11.53 12.31 6.04 126 Ultisols Siltstone,sandstone Urban forest, RP 7

C/N II1A 24 3 5 24 28 20 126 Ultisols Siltstone,sandstone Urban forest, RP 7

II1C 186 110 5 186 297 66 126 Ultisols Siltstone,sandstone Urban forest, RP 7

II1Ci 33 10 6 35 42 22 126 Ultisols Siltstone,sandstone Urban forest, RP 7

Carbon (%) II1A 52 1 5 52 52 51 126 Ultisols Siltstone,sandstone Urban forest, RP 7

II1C 50 0 5 50 51 50 126 Ultisols Siltstone,sandstone Urban forest, RP 7

II1Ci 48 2 6 47 52 46 126 Ultisols Siltstone,sandstone Urban forest, RP 7

Nitrogen (%) II1A 2.19 0.29 5 2.15 2.60 1.81 126 Ultisols Siltstone,sandstone Urban forest, RP 7

II1C 0.39 0.26 5 0.27 0.76 0.17 126 Ultisols Siltstone,sandstone Urban forest, RP 7

II1Ci 1.58 0.51 6 1.48 2.16 1.12 126 Ultisols Siltstone,sandstone Urban forest, RP 7

Sulfur (%) II1A 0.22 0.02 5 0.23 0.24 0.20 126 Ultisols Siltstone,sandstone Urban forest, RP 7

II1C 0.08 0.02 5 0.07 0.10 0.06 126 Ultisols Siltstone,sandstone Urban forest, RP 7

II1Ci 0.28 0.13 6 0.22 0.47 0.18 126 Ultisols Siltstone,sandstone Urban forest, RP 7

Wood density (g/cc) II1C 0.40 0.11 5 0.34 0.55 0.30 126 Ultisols Siltstone,sandstone Urban forest, RP 7

Alchornea latifolia Aluminum (mg/g) II1A 0.21 0.31 17 0.14 1.39 0.02 135 Lmrf Ultisols Tuffac. sandstone 8

II1B 0.04 0.06 5 0.01 0.14 0.01 135 Lmrf Ultisols Tuffac. sandstone 8

II1C 0.04 0.04 12 0.03 0.10 0.01 135 Lmrf Ultisols Tuffac. sandstone 8

II1E 1.35 1.70 12 0.69 6.35 0.19 135 Lmrf Ultisols Tuffac. sandstone 8

II1H 0.10 0.06 6 0.11 0.19 0.01 135 Lmrf Ultisols Tuffac. sandstone 8

Ash (%) II1B 2.94 0.95 5 2.83 4.51 1.96 68 Swf Ultisols Tuffac. sandstone Class 3 LD 8

II1B 2.11 0.08 2 2.11 2.16 2.05 68 Swf Ultisols Tuffac. sandstone Class 1 SD 8

II1B 5.80 1 68 Swf Ultisols Tuffac. sandstone Class 3/4 LD 8

II1B 11.48 1 68 Swf Ultisols Tuffac. sandstone Class 3 ST 8

Calcium (mg/g) II1A 7.08 1.93 17 6.54 9.94 4.29 135 Lmrf Ultisols Tuffac. sandstone 8

II1B 8.38 2.82 5 7.50 13.17 5.73 68 Swf Ultisols Tuffac. sandstone Class 3 LD 8

II1B 4.90 1.19 2 4.90 5.74 4.06 68 Swf Ultisols Tuffac. sandstone Class 1 SD 8
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

21

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1B 13.62 1 68 Swf Ultisols Tuffac. sandstone Class 3/4 LD 8

II1B 41.60 1 68 Swf Ultisols Tuffac. sandstone Class 3 ST 8

II1B 2.82 0.84 5 2.41 4.21 2.22 135 Lmrf Ultisols Tuffac. sandstone 8

II1C 15.83 1 135 Lmrf Ultisols Tuffac. sandstone 8

II1C 7.23 1 135 Lmrf Ultisols Tuffac. sandstone 8

II1C 2.25 0.56 12 2.10 3.59 1.52 135 Lmrf Ultisols Tuffac. sandstone 8

II1Ci 5.44 0.05 2 5.44 5.47 5.40 135 Lmrf Ultisols Tuffac. sandstone 8

II1E 2.73 0.88 12 2.71 4.54 1.14 135 Lmrf Ultisols Tuffac. sandstone 8

II1H 5.73 1.19 6 5.64 7.40 4.26 135 Lmrf Ultisols Tuffac. sandstone 8

Iron (mg/g) II1A 0.16 0.06 17 0.15 0.25 0.09 135 Lmrf Ultisols Tuffac. sandstone 8

II1B 0.03 0.02 5 0.02 0.07 0.01 135 Lmrf Ultisols Tuffac. sandstone 8

II1C 0.09 0.05 12 0.08 0.20 0.03 135 Lmrf Ultisols Tuffac. sandstone 8

II1E 0.09 0.05 12 0.08 0.20 0.03 135 Lmrf Ultisols Tuffac. sandstone 8

II1H 0.12 0.03 6 0.11 0.17 0.09 135 Lmrf Ultisols Tuffac. sandstone 8

Magnesium (mg/g) II1A 2.69 0.54 17 2.67 3.90 1.83 135 Lmrf Ultisols Tuffac. sandstone 8

II1B 0.83 0.44 5 0.67 1.55 0.37 135 Lmrf Ultisols Tuffac. sandstone 8

 II1C 1.41 1 135 Lmrf Ultisols Tuffac. sandstone 8

II1C 0.79 1 135 Lmrf Ultisols Tuffac. sandstone 8

II1C 0.78 0.21 12 0.77 1.39 0.58 135 Lmrf Ultisols Tuffac. sandstone 8

II1Ci 1.18 0.00 2 1.18 1.18 1.18 135 Lmrf Ultisols Tuffac. sandstone 8

II1E 0.74 0.21 12 0.75 1.04 0.38 135 Lmrf Ultisols Tuffac. sandstone 8

II1H 2.19 0.33 6 2.20 2.75 1.81 135 Lmrf Ultisols Tuffac. sandstone 8

Manganese (mg/g) II1A 0.91 0.36 17 0.77 1.60 0.44 135 Lmrf Ultisols Tuffac. sandstone 8

II1B 0.42 0.31 5 0.29 0.95 0.21 135 Lmrf Ultisols Tuffac. sandstone 8

II1C 0.37 0.14 12 0.38 0.65 0.16 135 Lmrf Ultisols Tuffac. sandstone 8

II1E 0.18 0.10 12 0.16 0.46 0.11 135 Lmrf Ultisols Tuffac. sandstone 8

II1H 0.72 0.34 6 0.62 1.26 0.37 135 Lmrf Ultisols Tuffac. sandstone 8

Nitrogen (%) II1A 1.93 0.34 17 1.88 2.70 1.35 135 Lmrf Ultisols Tuffac. sandstone 8

II1B 0.48 0.15 5 0.42 0.68 0.31 68 Swf Ultisols Tuffac. sandstone Class 3 LD 8
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

22

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1B 0.30 1 68 Swf Ultisols Tuffac. sandstone Class 1 SD 8

II1B 0.74 1 68 Swf Ultisols Tuffac. sandstone Class 3/4 LD 8

II1B 0.60 1 68 Swf Ultisols Tuffac. sandstone Class 3 ST 8

II1B 0.68 0.18 5 0.78 0.84 0.48 135 Lmrf Ultisols Tuffac. sandstone 8

II1C 0.26 1 135 Lmrf Ultisols Tuffac. sandstone 8

II1C 0.22 1 135 Lmrf Ultisols Tuffac. sandstone 8

II1C 0.76 0.30 12 0.74 1.16 0.35 135 Lmrf Ultisols Tuffac. sandstone 8

II1Ci 0.66 0.02 2 0.66 0.67 0.64 135 Lmrf Ultisols Tuffac. sandstone 8

II1E 0.71 0.24 12 0.73 1.32 0.37 135 Lmrf Ultisols Tuffac. sandstone 8

II1H 1.48 0.26 6 1.47 1.75 1.23 135 Lmrf Ultisols Tuffac. sandstone 8

Phosphorus (mg/g) II1A 0.84 0.20 17 0.82 1.31 0.51 135 Lmrf Ultisols Tuffac. sandstone 8

II1B 0.10 0.10 5 0.10 0.30 0.09 68 Swf Ultisols Tuffac. sandstone Class 3 LD 8

II1B 0.14 0.02 2 0.14 0.16 0.13 68 Swf Ultisols Tuffac. sandstone Class 1 SD 8

II1B 0.43 1 68 Swf Ultisols Tuffac. sandstone Class 3/4 LD 8

II1B 0.24 1 68 Swf Ultisols Tuffac. sandstone Class 3 ST 8

II1B 0.45 0.17 5 0.46 0.63 0.27 135 Lmrf Ultisols Tuffac. sandstone 8

II1C 0.21 1 135 Lmrf Ultisols Tuffac. sandstone 8

II1C 0.19 1 135 Lmrf Ultisols Tuffac. sandstone 8

II1C 0.48 0.13 12 0.48 0.68 0.22 135 Lmrf Ultisols Tuffac. sandstone 8

II1Ci 0.31 0.00 2 0.31 0.31 0.31 135 Lmrf Ultisols Tuffac. sandstone 8

II1E 0.48 0.13 12 0.48 0.68 0.22 135 Lmrf Ultisols Tuffac. sandstone 8

II1H 0.76 0.14 6 0.71 1.03 0.66 135 Lmrf Ultisols Tuffac. sandstone 8

Potassium (mg/g) II1A 8.85 2.03 17 9.42 12.65 5.67 135 Lmrf Ultisols Tuffac. sandstone 8

II1B 0.94 0.28 5 0.86 1.30 0.86 68 Swf Ultisols Tuffac. sandstone Class 3 LD 8

II1B 5.03 0.78 2 5.03 5.58 4.48 68 Swf Ultisols Tuffac. sandstone Class 1 SD 8

II1B 3.13 1 68 Swf Ultisols Tuffac. sandstone Class 3/4 LD 8

II1B 1.48 1 68 Swf Ultisols Tuffac. sandstone Class 3 ST 8

II1B 7.30 2.18 5 6.91 10.98 5.48 135 Lmrf Ultisols Tuffac. sandstone 8

II1C 1.39 1 135 Lmrf Ultisols Tuffac. sandstone 8
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

23

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1C 2.38 1 135 Lmrf Ultisols Tuffac. sandstone 8

II1C 8.29 2.05 12 7.84 12.23 4.32 135 Lmrf Ultisols Tuffac. sandstone 8

II1Ci 4.56 0.23 2 4.56 4.72 4.40 135 Lmrf Ultisols Tuffac. sandstone 8

II1E 4.78 0.78 12 4.72 6.29 3.60 135 Lmrf Ultisols Tuffac. sandstone 8

II1H 10.42 3.55 6 10.14 17.05 7.03 135 Lmrf Ultisols Tuffac. sandstone 8

Wood density (g/cc) II1B 0.38 0.00 3 0.38 0.38 0.38 97 Wfs Inceptisols Tuffac. sandstone 8

Alchorneopsis floribunda Ash (%) II1B 1.63 0.16 2 1.63 1.74 1.51 68 Swf Ultisols Tuffac. sandstone Class 3 LD 9

II1B 1.72 0.57 8 1.57 2.68 1.11 68 Swf Ultisols Tuffac. sandstone Class 3/4LD 9

Calcium (mg/g) II1B 4.96 0.96 2 4.93 5.61 4.25 68 Swf Ultisols Tuffac. sandstone Class 3 LD 9

II1B 4.10 2.17 8 3.25 7.81 2.20 68 Swf Ultisols Tuffac. sandstone Class 3/4 LD 9

Nitrogen (%) II1B 0.70 0.11 2 0.70 0.77 0.62 68 Swf Ultisols Tuffac. sandstone Class 3 LD 9

II1B 0.82 0.10 8 0.79 1.03 0.73 68 Swf Ultisols Tuffac. sandstone Class 3/4 LD 9

Phosphorus (mg/g) II1B 0.21 0.06 2 0.21 0.26 0.17 68 Swf Ultisols Tuffac. sandstone Class 3 LD 9

II1B 0.21 0.03 8 0.21 0.27 0.16 68 Swf Ultisols Tuffac. sandstone Class 3/4 LD 9

Potassium (mg/g) II1B 0.40 0.34 8 0.30 1.22 0.17 68 Swf Ultisols Tuffac. sandstone Class 3/4 LD 9

II1B 1.17 0.17 2 1.12 1.29 1.05 68 Swf Ultisols Tuffac. sandstone Class 3 LD 9

Alchorneopsis portoricensis Calcium (mg/g) II1B 0.58 1 135 Lmrf Ultisols Tuffac. sandstone 10

II1Ci 12.88 1 135 Lmrf Ultisols Tuffac. sandstone 10

Magnesium (mg/g) II1B 1.01 1 135 Lmrf Ultisols Tuffac. sandstone 10

II1Ci 0.86 1 135 Lmrf Ultisols Tuffac. sandstone 10

Nitrogen (%) II1B 0.30 1 135 Lmrf Ultisols Tuffac. sandstone 10

II1Ci 0.64 1 135 Lmrf Ultisols Tuffac. sandstone 10

Phosphorus (mg/g) II1B 0.15 1 135 Lmrf Ultisols Tuffac. sandstone 10

II1Ci 0.23 1 135 Lmrf Ultisols Tuffac. sandstone 10

Potassium (mg/g) II1B 2.25 1 135 Lmrf Ultisols Tuffac. sandstone 10

II1Ci 3.96 1 135 Lmrf Ultisols Tuffac. sandstone 10
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

24

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Aleurites moluccana Wood density (g/cc) II1B 0.33 0.01 4 0.33 0.32 97 Wfs Inceptisols Tuffac. sandstone 11

Allamanda violacea Ash (%) II2A 10.09 1 91 Sdf Mollisols Alluvial deposits 12

Calcium (mg/g) II2A 20.00 1 91 Sdf Mollisols Alluvial deposits 12

Nitrogen (%) II2A 1.64 1 91 Sdf Mollisols Alluvial deposits 12

Phosphorus (mg/g) II2A 1.48 1 91 Sdf Mollisols Alluvial deposits 12

Potassium (mg/g) II2A 7.40 1 91 Sdf Mollisols Alluvial deposits 12

Alsophila portorricensis Aluminum (mg/g) II6A 1.85 0.16 2 1.85 1.96 1.74 143 Wflm Inceptisols Tuffac. sandstone 13

II6B 2.75 0.56 2 2.75 3.14 2.35 143 Wflm Inceptisols Tuffac. sandstone 13

II6 1.39 0.34 7 1.49 1.76 1.01 135 Lmrf Ultisols Tuffac. sandstone 13

Ash (%) II6A 6.63 0.10 2 6.63 6.70 6.56 143 Wflm Inceptisols Tuffac. sandstone 13

II6B 6.50 0.72 2 6.50 7.01 5.99 143 Wflm Inceptisols Tuffac. sandstone 13

C/N II6A 46 3 2 46 48 44 143 Wflm Inceptisols Tuffac. sandstone 13

II6B 460 551 2 460 849 70 143 Wflm Inceptisols Tuffac. sandstone 13

II6 25 3 13 25 32 21 135 Lmrf Ultisols Tuffac. sandstone 13

Calcium (mg/g) II6A 1.99 0.03 2 1.99 2.01 1.97 143 Wflm Inceptisols Tuffac. sandstone 13

II6B 1.51 0.11 2 1.51 1.58 1.43 143 Wflm Inceptisols Tuffac. sandstone 13

II6 1.89 0.50 7 1.78 2.61 1.24 135 Lmrf Ultisols Tuffac. sandstone 13

Carbon (%) II6A 48 0 2 48 48 48 143 Wflm Inceptisols Tuffac. sandstone 13

II6B 49 0 2 49 49 48 143 Wflm Inceptisols Tuffac. sandstone 13

II6 41 3 13 42 45 35 135 Lmrf Ultisols Tuffac. sandstone 13

Iron (mg/g) II6A 0.54 0.04 2 0.54 0.56 0.51 143 Wflm Inceptisols Tuffac. sandstone 13

II6B 1.00 0.04 2 1.00 1.02 0.97 143 Wflm Inceptisols Tuffac. sandstone 13

II6 0.13 0.03 7 0.12 0.19 0.10 135 Lmrf Ultisols Tuffac. sandstone 13

Magnesium (mg/g) II6A 3.11 0.26 2 3.11 3.30 2.93 143 Wflm Inceptisols Tuffac. sandstone 13

II6B 1.18 0.07 2 1.18 1.23 1.13 143 Wflm Inceptisols Tuffac. sandstone 13

II6 2.15 0.60 7 2.10 2.97 1.26 135 Lmrf Ultisols Tuffac. sandstone 13

Manganese (mg/g) II6A 0.10 0.02 2 0.10 0.12 0.09 143 Wflm Inceptisols Tuffac. sandstone 13
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

25

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II6B 0.06 0.00 2 0.06 0.06 0.06 143 Wflm Inceptisols Tuffac. sandstone 13

II6 0.05 0.01 7 0.05 0.06 0.02 135 Lmrf Ultisols Tuffac. sandstone 13

Nitrogen (%) II6A 1.05 0.07 2 1.05 1.10 1.00 143 Wflm Inceptisols Tuffac. sandstone 13

II6B 0.38 0.45 2 0.38 0.69 0.06 143 Wflm Inceptisols Tuffac. sandstone 13

II6 1.47 0.39 20 1.46 2.10 0.87 135 Lmrf Ultisols Tuffac. sandstone 13

Phosphorus (mg/g) II6A 0.54 0.05 2 0.54 0.57 0.51 143 Wflm Inceptisols Tuffac. sandstone 13

II6B 0.52 0.11 2 0.52 0.59 0.44 143 Wflm Inceptisols Tuffac. sandstone 13

II6 0.58 0.20 7 0.47 0.94 0.40 135 Lmrf Ultisols Tuffac. sandstone 13

Potassium (mg/g) II6A 11.72 0.15 2 11.72 11.82 11.62 143 Wflm Inceptisols Tuffac. sandstone 13

II6B 9.45 0.25 2 9.45 9.63 9.27 143 Wflm Inceptisols Tuffac. sandstone 13

II6 9.82 2.71 7 9.41 14.85 6.75 135 Lmrf Ultisols Tuffac. sandstone 13

Amyris elemifera Aluminum (mg/g) II1A 0.07 0.03 13 0.07 0.11 0.03 151 Sdf Mollisols Alluvial deposits 14

Ash (%) II1A 10.70 1.93 12 10.53 16.48 9.01 151 Sdf Mollisols Alluvial deposits 14

C/N II1A 23 2 12 23 25 20 151 Sdf Mollisols Alluvial deposits 14

Calcium (mg/g) II1A 28.08 3.47 13 27.82 35.43 23.25 151 Sdf Mollisols Alluvial deposits 14

Carbon (%) II1A 52 1 12 52 53 51 151 Sdf Mollisols Alluvial deposits 14

Iron (mg/g) II1A 0.06 0.02 13 0.06 0.09 0.04 151 Sdf Mollisols Alluvial deposits 14

Magnesium (mg/g) II1A 3.13 1 104 Sdf Mollisols Alluvial deposits Mature 14

II1A 3.13 1 149 Sdf Mollisols Alluvial deposits Guanica Forest 14

II1A 1.30 0.13 13 1.28 1.54 1.14 151 Sdf Mollisols Alluvial deposits 14

Manganese (mg/g) II1A 0.05 0.01 13 0.04 0.05 0.04 151 Sdf Mollisols Alluvial deposits 14

Nitrogen (%) II1A 1.59 1 149 Sdf Mollisols Alluvial deposits Guanica Forest 14

II1A 2.31 0.19 12 2.27 2.60 2.05 151 Sdf Mollisols Alluvial deposits 14

Phosphorus (mg/g) II1A 0.59 1 104 Sdf Mollisols Alluvial deposits Mature 14

II1A 0.59 1 149 Sdf Mollisols Alluvial deposits Guanica Forest 14

II1A 0.52 0.05 13 0.49 0.62 0.45 151 Sdf Mollisols Alluvial deposits 14

Potassium (mg/g) II1A 15.95 1 104 Sdf Mollisols Alluvial deposits Mature 14

II1A 15.95 1 149 Sdf Mollisols Alluvial deposits Guanica Forest 14
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

26

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 9.25 0.74 13 9.31 10.79 8.12 151 Sdf Mollisols Alluvial deposits 14

Sulfur (%) II1A 0.32 0.02 12 0.31 0.34 0.28 151 Sdf Mollisols Alluvial deposits 14

Andira inermis Ash (%) II1A 4.48 1 94 Wfs Ultisols Lava 15

II1B 3.48 1 94 Wfs Ultisols Lava 15

II1C 1.34 1 94 Wfs Ultisols Lava 15

Calcium (mg/g) II1A 2.70 1 135 Lmrf Ultisols Tuffac. sandstone 15

II1B 1.63 1 135 Lmrf Ultisols Tuffac. sandstone 15

II1C 0.78 1 135 Lmrf Ultisols Tuffac. sandstone 15

Magnesium (mg/g) II1A 2.58 1 135 Lmrf Ultisols Tuffac. sandstone 15

II1B 1.63 1 135 Lmrf Ultisols Tuffac. sandstone 15

II1C 0.35 1 135 Lmrf Ultisols Tuffac. sandstone 15

Nitrogen (%) II1A 1.95 1 94 Wfs Ultisols Lava 15

II1A 2.66 1 135 Lmrf Ultisols Tuffac. sandstone 15

II1B 1.72 1 94 Wfs Ultisols Lava 15

II1B 1.53 1 135 Lmrf Ultisols Tuffac. sandstone 15

II1C 0.95 1 94 Wfs Ultisols Lava 15

II1C 0.41 1 135 Lmrf Ultisols Tuffac. sandstone 15

Phosphorus (mg/g) II1A 1.47 1 94 Wfs Ultisols Lava 15

II1A 1.13 1 135 Lmrf Ultisols Tuffac. sandstone 15

II1B 1.02 1 94 Wfs Ultisols Lava 15

II1B 0.77 1 135 Lmrf Ultisols Tuffac. sandstone 15

II1C 0.52 1 94 Wfs Ultisols Lava 15

II1C 0.22 1 135 Lmrf Ultisols Tuffac. sandstone 15

Potassium (mg/g) II1A 6.48 1 94 Wfs Ultisols Lava 15

II1A 8.34 1 135 Lmrf Ultisols Tuffac. sandstone 15

II1B 6.72 1 94 Wfs Ultisols Lava 15

II1B 8.26 1 135 Lmrf Ultisols Tuffac. sandstone 15

II1C 2.30 1 94 Wfs Ultisols Lava 15
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

27

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1C 3.54 1 135 Lmrf Ultisols Tuffac. sandstone 15

Anetium citrifolium Aluminum (mg/g) II6 3.26 1 135 Lmrf Ultisols Tuffac. sandstone 16

C/N II6 19 1 135 Lmrf Ultisols Tuffac. sandstone 16

Calcium (mg/g) II6 5.17 1 135 Lmrf Ultisols Tuffac. sandstone 16

Carbon (%) II6 38 1 135 Lmrf Ultisols Tuffac. sandstone 16

Iron (mg/g) II6 2.08 1 135 Lmrf Ultisols Tuffac. sandstone 16

Magnesium (mg/g) II6 3.67 1 135 Lmrf Ultisols Tuffac. sandstone 16

Manganese (mg/g) II6 0.27 1 135 Lmrf Ultisols Tuffac. sandstone 16

Nitrogen (%) II6 2.04 1 135 Lmrf Ultisols Tuffac. sandstone 16

Phosphorus (mg/g) II6 1.67 1 135 Lmrf Ultisols Tuffac. sandstone 16

Potassium (mg/g) II6 30.51 1 135 Lmrf Ultisols Tuffac. sandstone 16

Sulfur (%) II6 0.66 1 135 Lmrf Ultisols Tuffac. sandstone 16

Anthocephalus chinensis Ash (%) II8A 6.03 0.15 2 6.03 6.13 5.92 90 Swf Ultisols Tuffac. sandstone Old 17

II8A 10.36 8.95 3 6.17 20.63 4.27 90 Swf Ultisols Tuffac. sandstone 17

II8B 3.33 0.84 3 3.42 4.12 2.44 90 Swf Ultiso1s Tuffac. sandstone 17

II8C 3.93 0.94 3 4.29 4.64 2.86 90 Swf Ultisols Tuffac. sandstone 17

II8D 14.78 0.20 2 14.78 14.92 14.64 90 Swf Ultisols Tuffac. sandstone 17

II8Bi 2.81 1.25 3 3.07 3.91 1.46 90 Swf Ultisols Tuffac. sandstone 17

Calcium (mg/g) II8A 15.30 2.14 2 15.30 16.81 13.79 90 Swf Ultisols Tuffac. sandstone 17

II8A 10.10 3.13 3 8.57 13.70 8.02 90 Swf Ultisols Tuffac. sandstone 17

II8B 7.16 0.80 3 6.92 8.05 6.50 90 Swf Ultisols Tuffac. sandstone 17

II8C 7.44 0.48 3 6.94 7.90 6.94 90 Swf Ultisols Tuffac. sandstone 17

II8D 11.16 2.87 2 11.16 13.19 9.13 90 Swf Ultisols Tuffac. sandstone 17

II8Bi 3.57 1.20 3 3.13 4.93 2.66 90 Swf Ultisols Tuffac. sandstone 17

II9A 10.43 2.99 22 10.72 16.64 4.42 139 Wfs Ultisols Tuffac. sandstone 17

II9A 7.63 0.81 22 7.77 8.95 6.21 139 Wfs Ultisols Tuffac. sandstone Other 17

II9B 6.17 2.07 5 6.89 8.75 3.48 139 Wfs Ultisols Tuffac. sandstone 17
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

28

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II9C 4.61 0.67 6 4.84 5.20 3.33 139 Wfs Ultisols Tuffac. sandstone 17

II9D 7.12 0.96 5 7.28 8.22 5.57 139 Wfs Ultisols Tuffac. sandstone 17

Magnesium (mg/g) II9A 4.20 1.03 22 3.97 6.52 2.48 139 Wfs Ultisols Tuffac. sandstone 17

II9A 2.85 0.64 22 2.84 4.56 1.94 139 Wfs Ultisols Tuffac. sandstone Other 17

II9B 2.36 0.73 5 1.98 3.48 1.81 139 Wfs Ultisols Tuffac. sandstone 17

II9C 2.07 0.53 6 2.07 2.64 1.18 139 Wfs Ultisols Tuffac. sandstone 17

II9D 2.83 0.25 5 2.84 3.18 2.48 139 Wfs Ultisols Tuffac. sandstone 17

Nitrogen (%) II8A 1.11 0.05 2 1.11 1.14 1.07 90 Swf Ultisols Tuffac. sandstone 17

II8A 1.19 0.15 3 1.15 1.36 1.07 90 Swf Ultisols Tuffac. sandstone 17

II8B 0.43 0.15 3 0.44 0.57 0.28 90 Swf Ultisols Tuffac. sandstone 17

II8C 0.80 0.17 3 0.72 1.00 0.69 90 Swf Ultisols Tuffac. sandstone 17

II8D 1.21 0.02 2 1.21 1.22 1.19 90 Swf Ultisols Tuffac. sandstone 17

II8Bi 0.39 0.15 3 0.37 0.54 0.25 90 Swf Ultisols Tuffac. sandstone 17

II9A 1.19 0.34 22 1.13 1.87 0.19 139 Wfs Ultisols Tuffac. sandstone 17

II9A 0.93 0.26 22 0.93 1.32 0.51 139 Wfs Ultisols Tuffac. sandstone Other 17

II9B 0.37 0.08 10 0.37 0.50 0.24 139 Wfs Ultisols Tuffac. sandstone 17

II9C 0.58 0.17 11 0.53 0.94 0.35 139 Wfs Ultisols Tuffac. sandstone 17

II9D 1.17 0.24 5 1.18 1.66 0.80 139 Wfs Ultisols Tuffac. sandstone 17

Phosphorus (mg/g) II8A 0.42 0.04 2 0.42 0.45 0.39 90 Swf Ultisols Tuffac. sandstone 17

II8A 0.53 0.05 3 0.48 0.58 0.48 90 Swf Ultisols Tuffac. sandstone 17

II8B 0.15 0.07 3 0.15 0.21 0.08 90 Swf Ultisols Tuffac. sandstone 17

II8C 0.32 0.12 3 0.34 0.43 0.19 90 Swf Ultisols Tuffac. sandstone 17

II8D 0.54 0.00 2 0.54 0.54 0.54 90 Swf Ultisols Tuffac. sandstone 17

II8Bi 0.12 0.05 3 0.13 0.17 0.07 90 Swf Ultisols Tuffac. sandstone 17

II9A 0.64 0.17 22 0.61 1.03 0.26 139 Wfs Ultisols Tuffac. sandstone 17

II9A 0.38 0.14 22 0.37 0.65 0.17 139 Wfs Ultisols Tuffac. sandstone 17

II9B 0.18 0.05 5 0.17 0.25 0.13 139 Wfs Ultisols Tuffac. sandstone 17

II9C 0.45 0.17 6 0.48 0.61 0.16 139 Wfs Ultisols Tuffac. sandstone 17

II9D 0.66 0.10 5 0.69 0.73 0.49 139 Wfs Ultisols Tuffac. sandstone 17
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

29

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Potassium (mg/g) II8A 2.25 0.45 2 2.25 2.57 1.93 90 Swf Ultisols Tuffac. sandstone 17

II8A 5.36 4.50 3 3.46 10.50 2.13 90 Swf Ultisols Tuffac. sandstone 17

II8B 1.98 0.92 3 1.47 3.04 1.42 90 Swf Ultisols Tuffac. sandstone 17

II8C 2.37 1.14 3 1.76 3.68 1.66 90 Swf Ultisols Tuffac. sandstone 17

II8D 2.95 1.96 2 2.95 4.34 1.56 90 Swf Ultisols Tuffac. sandstone 17

II8Bi 1.47 0.90 3 1.04 2.50 0.86 90 Swf Ultisols Tuffac. sandstone 17

II9A 4.67 1.16 22 4.72 6.29 2.56 139 Wfs Ultisols Tuffac. sandstone 17

II9A 3.81 0.62 22 3.77 5.21 2.91 139 Wfs Ultisols Tuffac. sandstone Other 17

II9B 3.09 0.97 5 2.82 4.44 1.89 139 Wfs Ultisols Tuffac. sandstone 17

II9C 4.62 1.18 6 4.65 5.92 3.28 139 Wfs Ultisols Tuffac. sandstone 17

II9D 4.04 1.52 5 3.73 6.29 2.14 139 Wfs Ultisols Tuffac. sandstone 17

Wood density (g/cc) II1C 0.26 0.01 20 0.26 0.28 0.25 115 17

Ardisia glauciflora Aluminum (mg/kg) II1A 41 1 102 Smf Limestone Tuffac. sandstone Adult 18

Ash (%) II1A 10.41 1 102 Smf Limestone Tuffac. sandstone Adult 18

Calcium (mg/g) II1A 31.52 1 102 Smf Limestone Tuffac. sandstone Adult 18

Cobalt (μg/g) II1A 1.70 1 102 Smf Limestone Tuffac. sandstone Adult 18

Iron (mg/kg) II1A 49 1 102 Smf Limestone Tuffac. sandstone Adult 18

Magnesium (mg/g) II1A 0.92 1 102 Smf Limestone Tuffac. sandstone Adult 18

Manganese (mg/kg) II1A 23 1 102 Smf Limestone Tuffac. sandstone Adult 18

Nickel (μg/g) II1A 7.50 1 102 Smf Limestone Tuffac. sandstone Adult 18

Nitrogen (%) II1A 0.53 1 102 Smf Limestone Tuffac. sandstone Adult 18

Phosphorus (mg/g) II1A 0.27 1 102 Smf Limestone Tuffac. sandstone Adult 18

Potassium (mg/g) II1A 4.17 1 102 Smf Limestone Tuffac. sandstone Adult 18

Ardisia solanacea Ash (%) II1A 13.50 0.60 6 13.23 14.36 12.86 126 Ultisols Siltstone,sandstone Botanical G., RP 19

II1C 1.77 0.31 6 1.74 2.21 1.35 126 Ultisols Siltstone,sandstone Botanical G., RP 19

II1Ci 7.49 1.46 5 7.85 8.81 4.99 126 Ultisols Siltstone,sandstone Botanical G., RP 19

C/N II1A 27 1 6 28 28 26 126 Ultisols Siltstone,sandstone Botanical G., RP 19
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

30

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1C 228 78 6 218 319 134 126 Ultisols Siltstone,sandstone Botanical G., RP 19

II1Ci 105 8 5 102 118 99 126 Ultisols Siltstone,sandstone Botanical G., RP 19

Carbon (%) II1A 49 1 6 49 49 48 126 Ultisols Siltstone,sandstone Botanical G., RP 19

II1C 51 0 6 51 51 50 126 Ultisols Siltstone,sandstone Botanical G., RP 19

II1Ci 49 2 5 49 52 48 126 Ultisols Siltstone,sandstone Botanical G., RP 19

Nitrogen (%) II1A 1.79 0.07 6 1.78 1.90 1.71 126 Ultisols Siltstone,sandstone Botanical G., RP 19

II1C 0.25 0.09 6 0.24 0.38 0.16 126 Ultisols Siltstone,sandstone Botanical G., RP 19

II1Ci 0.47 0.02 5 0.47 0.49 0.44 126 Ultisols Siltstone,sandstone Botanical G., RP 19

Sulfur (%) II1A 0.19 0.01 6 0.19 0.21 0.18 126 Ultisols Siltstone,sandstone Botanical G., RP 19

II1C 0.05 0.01 6 0.05 0.05 0.03 126 Ultisols Siltstone,sandstone Botanical G., RP 19

II1Ci 0.08 0.01 5 0.08 0.09 0.07 126 Ultisols Siltstone,sandstone Botanical G., RP 19

Wood density (g/cc) II1C 0.48 0.03 5 0.49 0.51 0.43 126 Ultisols Siltstone,sandstone Botanical G., RP 19

Artocarpus altilis Wood density (g/cc) II1B 0.32 0.01 3 0.32 0.33 0.32 97 Wfs Inceptisols Tuffac. sandstone 20

Asplenium spp. Aluminum (mg/g) II6 0.16 1 135 Lmrf Ultisols Tuffac. sandstone 21

Calcium (mg/g) II6 18.22 1 135 Lmrf Ultisols Tuffac. sandstone 21

Iron (mg/g) II6 0.15 1 135 Lmrf Ultisols Tuffac. sandstone 21

Magnesium (mg/g) II6 3.52 1 135 Lmrf Ultisols Tuffac. sandstone 21

Manganese (mg/g) II6 1.16 1 135 Lmrf Ultisols Tuffac. sandstone 21

Nitrogen (%) II6 1.02 1 135 Lmrf Ultisols Tuffac. sandstone 21

Phosphorus (mg/g) II6 0.98 1 135 Lmrf Ultisols Tuffac. sandstone 21

Potassium (mg/g) II6 27.82 1 135 Lmrf Ultisols Tuffac. sandstone 21

Avicennia germinans Aluminum (mg/g) II1A 0.17 0.02 2 0.17 0.18 0.16 78 Mfs Mangle Swamp 22

II1B 0.03 0.01 4 0.03 0.05 0.03 87 Mfs Entisols Compound dunes 22

II1B 0.03 0.03 3 0.02 0.07 0.02 157 Mfs Entisols Compound dunes 22

Aluminum (mg/kg) II1A 7 3 4 6 10 4 80 Mfs Entisols Swamps 22

Ash (%) II1A 10.21 0.14 6 10.21 10.42 10.04 80 Mfs Entisols Swamps 22
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

31

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 9.80 0.07 2 9.80 9.85 9.75 78 Mfs Mangle Swamp 22

II1B 1.48 0.35 4 1.45 1.93 1.08 87 Mfs Entisols Compound dunes 22

II1C 8.84 0.16 4 8.88 8.98 8.62 80 Mfs Entisols Swamps 22

C/N II1A 22 0 6 22 22 22 80 Mfs Entisols Swamps 22

II1A 20 0 2 20 20 20 78 Mfs Mangle Swamp 22

II1B 155 42 4 157 192 114 87 Mfs Entisols Compound dunes 22

II1C 35 1 4 35 36 34 80 Mfs Entisols Swamps 22

Calcium (mg/g) II1A 5.77 0.20 6 5.66 6.06 5.60 80 Mfs Entisols Swamps 22

II1A 2.82 1 74 Dfs Mangle Dune deposits Old Mature 22

II1A 4.21 1 74 Dfs Mangle Dune deposits Senescent 22

II1A 3.32 1 74 Dfs Mangle Dune deposits Yellow 22

II1A 2.66 1 74 Dfs Mangle Dune deposits Young Mature 22

II1A 6.46 0.19 2 6.46 6.60 6.33 78 Mfs Mangle Swamp 22

II1B 1.76 0.31 4 1.63 2.21 1.56 87 Mfs Entisols Compound dunes 22

II1B 6.56 0.60 3 6.37 7.23 6.24 157 Mfs Entisols Compound dunes 22

II1C 4.99 0.42 4 4.93 5.55 4.56 80 Mfs Entisols Swamps 22

Carbon (%) II1A 48 0 6 48 48 48 80 Mfs Entisols Swamps 22

II1A 42 0 2 42 42 42 78 Mfs Mangle Swamp 22

II1C 45 0 4 45 46 45 80 Mfs Entisols Swamps 22

Iron (mg/g) II1A 0.14 0.00 2 0.14 0.14 0.14 78 Mfs Mangle Swamp 22

II1B 0.05 0.03 4 0.05 0.09 0.02 87 Mfs Entisols Compound dunes 22

II1B 0.06 0.07 3 0.03 0.01 0.13 157 Mfs Entisols Compound dunes 22

II1A 0.02 0.01 6 0.02 0.03 0.01 80 Mfs Entisols Swamps 22

II1C 0.01 0.00 6 0.01 0.02 0.01 80 Mfs Entisols Swamps 22

Magnesium (mg/g) II1A 7.97 0.18 6 7.96 8.30 7.79 80 Mfs Entisols Swamps 22

II1A 9.14 1 74 Dfs Mangle Dune deposits Old Mature 22

II1A 10.77 1 74 Dfs Mangle Dune deposits Senescent 22

II1A 10.46 1 74 Dfs Mangle Dune deposits Yellow 22

II1A 5.70 1 74 Dfs Mangle Dune deposits Young Mature 22
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

32

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 7.48 0.09 2 7.48 7.54 7.41 78 Mfs Mangle Swamp 22

II1B 0.70 0.16 4 0.77 0.80 0.46 78 Mfs Mangle Swamp 22

II1B 0.82 0.09 3 0.81 0.91 0.73 157 Mfs Entisols Compound dunes 22

II1C 1.49 0.12 4 1.48 1.64 1.36 80 Mfs Entisols Swamps 22

Manganese (mg/kg) II1A 200 10 2 200 210 200 78 Mfs Mangle Swamp 22

II1A 67 2 6 67 71 65 80 Mfs Entisols Swamps 22

II1B 7 2 4 7 10 6 78 Mfs Mangle Swamp 22

II1B 7 2 3 6 6 9 157 Mfs Entisols Compound dunes 22

II1C 14 2 4 14 17 12 80 Mfs Entisols Swamps 22

Nitrogen (%) II1A 2.22 0.02 6 2.23 2.24 2.19 80 Mfs Entisols Swamps 22

II1A 2.08 0.01 2 2.08 2.09 2.08 78 Mfs Mangle Swamp 22

II1A 1.53 1 74 Dfs Mangle Dune deposits Old Mature 22

II1A 0.60 1 Dfs Mangle Dune deposits Senescent 22

II1A 0.57 1 Dfs Mangle Dune deposits Yellow 22

II1A 1.82 1 Dfs Mangle Dune deposits Young Mature 22

II1B 0.31 0.09 4 0.31 0.40 0.24 87 Mfs Entisols Compound dunes 22

II1C 1.29 0.02 4 1.29 1.31 1.27 80 Mfs Entisols Swamps 22

Phosphorus (mg/g) II1A 1.18 0.04 6 1.18 1.22 1.11 80 Mfs Entisols Swamps 22

II1A 0.70 1 74 Dfs Mangle Dune deposits Old Mature 22

II1A 0.16 1 74 Dfs Mangle Dune deposits Senescent 22

II1A 0.31 1 74 Dfs Mangle Dune deposits Yellow 22

II1A 1.11 1 74 Dfs Mangle Dune deposits Young Mature 22

II1A 1.16 0.01 2 1.16 1.17 1.16 78 Mfs Mangle Swamp 22

II1B 0.32 0.09 4 0.28 0.45 0.25 87 Mfs Entisols Compound dunes 22

II1B 0.23 0.11 3 0.28 0.31 0.11 157 Mfs Entisols Compound dunes 22

II1C 1.08 0.08 4 1.07 1.17 0.99 80 Mfs Entisols Swamps 22

Potassium (mg/g) II1A 9.35 0.33 6 9.40 9.80 8.84 80 Mfs Entisols Swamps 22

II1A 12.40 1 74 Dfs Mangle Dune deposits Old Mature 22

II1A 12.61 1 74 Dfs Mangle Dune deposits Senescent 22
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

33

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 14.31 1 74 Dfs Mangle Dune deposits Yellow 22

II1A 12.19 1 74 Dfs Mangle Dune deposits Young Mature 22

II1A 14.23 0.83 2 14.23 14.82 13.64 78 Mfs Mangle Swamp 22

II1B 3.53 0.94 4 3.45 4.75 2.47 87 Mfs Entisols Compound dunes 22

II1B 2.88 1.53 3 3.53 3.97 1.33 157 Mfs Entisols Compound dunes 22

II1C 15.97 1.00 4 16.01 17.07 14.90 80 Mfs Entisols Swamps 22

Sodium (mg/g) II1A 16.88 0.43 6 16.74 17.51 16.34 80 Mfs Entisols Swamps 22

II1A 22.46 1 74 Dfs Mangle Dune deposits Old Mature 22

II1A 22.34 1 74 Dfs Mangle Dune deposits Senescent 22

II1A 24.87 1 74 Dfs Mangle Dune deposits Yellow 22

II1A 42.50 1 74 Dfs Mangle Dune deposits Young Mature 22

II1A 24.70 0.47 2 24.70 25.03 24.36 78 Mfs Mangle Swamp 22

II1C 12.49 1.07 4 12.38 13.82 11.37 80 Mfs Entisols Swamps 22

Bambusa spp. Aluminum (mg/g) II3A 0.94 0.03 2 0.94 0.96 0.93 129 Lmrf Ultisols Tuffac. sandstone 23

II3 5.96 1.81 21 6.10 9.57 1.96 129 Lmrf Ultisols Tuffac. sandstone Decomp. 23

II9A 0.78 0.27 34 0.75 2.01 0.33 129 Lmrf Ultisols Tuffac. sandstone 23

Ash (%) II3A 6.87 0.02 2 6.87 6.88 6.85 129 Lmrf Ultisols Tuffac. sandstone 23

II3 24.98 4.58 21 24.83 37.46 17.08 129 Lmrf Ultisols Tuffac. sandstone Decomp. 23

II9A 16.46 2.94 34 16.68 21.08 6.15 129 Lmrf Ultisols Tuffac. sandstone 23

C/N II3A 24 0 2 24 24 24 129 Lmrf Ultisols Tuffac. sandstone 23

II3 24 3 21 25 29 19 129 Lmrf Ultisols Tuffac. sandstone Decomp. 23

II9A 67 24 34 62 169 42 129 Lmrf Ultisols Tuffac. sandstone 23

Calcium (mg/g) II3A 8.16 0.05 2 8.16 8.20 8.12 129 Lmrf Ultisols Tuffac. sandstone 23

II3 1.36 0.31 21 1.29 1.98 0.80 129 Lmrf Ultisols Tuffac. sandstone Decomp. 23

II9A 4.26 1.32 34 4.38 6.63 0.86 129 Lmrf Ultisols Tuffac. sandstone 23

Carbon (%) II3A 53 0 2 53 53 52 129 Lmrf Ultisols Tuffac. sandstone 23

II3 39 2 21 40 43 33 129 Lmrf Ultisols Tuffac. sandstone Decomp. 23

II9A 44 1 34 44 49 42 129 Lmrf Ultisols Tuffac. sandstone 23
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

34

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Iron (mg/g) II3A 0.55 0.01 2 0.55 0.56 0.54 129 Lmrf Ultisols Tuffac. sandstone 23

II3 5.07 2.08 21 5.07 10.70 1.00 129 Lmrf Ultisols Tuffac. sandstone Decomp. 23

II9A 0.40 0.19 34 0.37 1.39 0.19 129 Lmrf Ultisols Tuffac. sandstone 23

Magnesium (mg/g) II3A 2.02 0.17 2 2.02 2.14 1.90 129 Lmrf Ultisols Tuffac. sandstone 23

II3 0.95 0.44 21 0.77 2.30 0.52 129 Lmrf Ultisols Tuffac. sandstone Decomp. 23

II9A 0.88 0.26 34 0.82 1.67 0.38 129 Lmrf Ultisols Tuffac. sandstone 23

Manganese (mg/g) II3A 1.01 0.00 2 1.01 1.01 1.00 129 Lmrf Ultisols Tuffac. sandstone 23

II3 0.12 0.05 21 0.12 0.28 0.05 129 Lmrf Ultisols Tuffac. sandstone Decomp. 23

II9A 1.02 0.34 34 0.91 1.65 0.33 129 Lmrf Ultisols Tuffac. sandstone 23

Nitrogen (%) II3A 2.23 0.01 2 2.23 2.23 2.22 129 Lmrf Ultisols Tuffac. sandstone 23

II3 1.63 0.14 21 1.59 1.88 1.39 129 Lmrf Ultisols Tuffac. sandstone Decomp. 23

II9A 0.70 0.15 34 0.69 1.02 0.29 129 Lmrf Ultisols Tuffac. sandstone 23

Phosphorus (mg/g) II3A 0.90 0.00 2 0.90 0.90 0.90 129 Lmrf Ultisols Tuffac. sandstone 23

II3 0.26 0.09 21 0.24 0.42 0.11 129 Lmrf Ultisols Tuffac. sandstone Decomp. 23

II9A 0.18 0.07 34 0.16 0.36 0.06 129 Lmrf Ultisols Tuffac. sandstone 23

Potassium (mg/g) II3A 12.09 0.05 2 12.09 12.12 12.05 129 Lmrf Ultisols Tuffac. sandstone 23

II3 0.63 0.18 21 0.59 0.93 0.38 129 Lmrf Ultisols Tuffac. sandstone Decomp. 23

II9A 1.46 0.41 34 1.44 2.66 0.62 129 Lmrf Ultisols Tuffac. sandstone 23

Sulfur (%) II3A 0.39 0.02 2 0.39 0.40 0.38 129 Lmrf Ultisols Tuffac. sandstone 23

II3 0.17 0.02 21 0.16 0.21 0.14 129 Lmrf Ultisols Tuffac. sandstone Decomp. 23

II9A 0.24 0.03 34 0.23 0.31 0.17 129 Lmrf Ultisols Tuffac. sandstone 23

Bambusa vulgaris Ash (%) II3A 21.42 1 94 Wfs Ultisols Lava < 1 cm dbh 24

II3C 13.33 1 94 Wfs Ultisols Lava < 1 cm dbh 24

Nitrogen (%) II3A 2.23 1 94 Wfs Ultisols Lava < 1 cm dbh 24

II3C 0.65 1 94 Wfs Ultisols Lava < 1 cm dbh 24

Phosphorus (mg/g) II3A 0.86 1 94 Wfs Ultisols Lava < 1 cm dbh 24

II3C 0.26 1 94 Wfs Ultisols Lava < 1 cm dbh 24

Potassium (mg/g) II3A 9.39 1 94 Wfs Ultisols Lava < 1 cm dbh 24
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

35

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II3C 3.95 1 94 Wfs Ultisols Lava < 1 cm dbh 24

Beilschmiedia pendula Ash (%) II1B 3.85 2.66 26 2.99 12.35 1.70 68 Swf Ultisols Tuffac. sandstone Decay-class 3/4 LD 25

Calcium (mg/g) II1B 8.35 7.34 26 6.09 34.25 2.40 68 Swf Ultisols Tuffac. sandstone Decay-class 3/4 LD 25

Nitrogen (%) II1B 0.70 0.27 26 0.65 1.10 0.32 68 Swf Ultisols Tuffac. sandstone Decay-class 3/4 LD 25

Phosphorus (mg/g) II1B 0.15 0.12 26 0.13 0.47 0.02 68 Swf Ultisols Tuffac. sandstone Decay-class 3/4 LD 25

Potassium (mg/g) II1B 1.17 0.89 26 1.09 4.04 0.12 68 Swf Ultisols Tuffac. sandstone Decay-class 3/4 LD 25

Blenchnum occidentale Aluminum (mg/g) II6 0.53 1 135 Lmrf Ultisols Tuffac. sandstone 26

Calcium (mg/g) II6 6.67 1 135 Lmrf Ultisols Tuffac. sandstone 26

Iron (mg/g) II6 0.33 1 135 Lmrf Ultisols Tuffac. sandstone 26

Magnesium (mg/g) II6 4.20 1 135 Lmrf Ultisols Tuffac. sandstone 26

Manganese (mg/g) II6 0.085 1 135 Lmrf Ultisols Tuffac. sandstone 26

Nitrogen (%) II6 2.45 1 135 Lmrf Ultisols Tuffac. sandstone 26

Phosphorus (mg/g) II6 1.41 1 135 Lmrf Ultisols Tuffac. sandstone 26

Potassium (mg/g) II6 48.62 1 135 Lmrf Ultisols Tuffac. sandstone 26

Bolbitis aliena Aluminum (mg/g) II6 1.26 1 135 Lmrf Ultisols Tuffac. sandstone 27

Calcium (mg/g) II6 7.05 1 135 Lmrf Ultisols Tuffac. sandstone 27

Iron (mg/g) II6 3.03 1 135 Lmrf Ultisols Tuffac. sandstone 27

Magnesium (mg/g) II6 4.85 1 135 Lmrf Ultisols Tuffac. sandstone 27

Manganese (mg/g) II6 0.18 1 135 Lmrf Ultisols Tuffac. sandstone 27

Phosphorus (mg/g) II6 1.11 1 135 Lmrf Ultisols Tuffac. sandstone 27

Potassium (mg/g) II6 58.54 1 135 Lmrf Ultisols Tuffac. sandstone 27

Bolbitis nicotianifolia Aluminum (mg/g) II6 2.39 1.70 17 1.98 7.43 0.87 135 Lmrf Ultisols Tuffac. sandstone 28

C/N II6 16 2 16 15 19 14 135 Lmrf Ultisols Tuffac. sandstone 28

Calcium (mg/g) II6 6.36 1.33 17 6.14 8.94 4.49 135 Lmrf Ultisols Tuffac. sandstone 28

Carbon (%) II6 40 2 16 41 43 34 135 Lmrf Ultisols Tuffac. sandstone 28
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

36

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Iron (mg/g) II6 1.28 1.61 17 0.58 6.39 0.14 135 Lmrf Ultisols Tuffac. sandstone 28

Magnesium (mg/g) II6 5.00 1.10 17 4.55 7.37 3.85 135 Lmrf Ultisols Tuffac. sandstone 28

Manganese (mg/g) II6 0.21 0.17 17 0.15 0.80 0.09 135 Lmrf Ultisols Tuffac. sandstone 28

Nitrogen (%) II6 2.52 0.36 23 2.55 3.01 1.67 135 Lmrf Ultisols Tuffac. sandstone 28

Phosphorus (mg/g) II6 1.17 0.22 17 1.12 1.85 0.92 135 Lmrf Ultisols Tuffac. sandstone 28

Potassium (mg/g) II6 42.00 5.58 17 29.61 42.23 22.18 135 Lmrf Ultisols Tuffac. sandstone 28

Sulfur (%) II6 0.45 0.12 16 0.43 0.66 0.26 135 Lmrf Ultisols Tuffac. sandstone 28

Bourreria succulenta Aluminum (mg/g) II1A 0.05 0.02 14 0.05 0.07 0.02 151 Sdf Mollisols Alluvial deposits 29

Ash (%) II1A 10.56 2.70 14 11.64 13.16 3.95 151 Sdf Mollisols Alluvial deposits 29

C/N II1A 28 3 14 29 31 23 151 Sdf Mollisols Alluvial deposits 29

Calcium (mg/g) II1A 14.84 1.74 2 14.85 16.67 12.99 104 Sdf Mollisols Alluvial deposits New 29

II1A 24.51 3.07 6 23.03 29.16 20.23 104 Sdf Mollisols Alluvial deposits Mature 29

II1A 28.98 1.39 2 28.96 30.25 27.74 104 Sdf Mollisols Alluvial deposits Old 29

II1A 23.35 5.58 18 23.03 30.25 12.99 149 Sdf Mollisols Alluvial deposits Guanica Forest 29

II1A 25.07 3.03 14 25.40 30.72 18.80 151 Sdf Mollisols Alluvial deposits 29

II1Ci 28.09 2.11 2 28.09 29.58 26.60 104 Sdf Mollisols Alluvial deposits 29

II1Ci 28.09 1 149 Sdf Mollisols Alluvial deposits Guanica Forest 29

Carbon (%) II1A 49 0 14 49 50 49 151 Sdf Mollisols Alluvial deposits 29

Iron (mg/kg) II1A 38 11 14 34 53 23 151 Sdf Mollisols Alluvial deposits 29

Magnesium (mg/g) II1A 3.50 0.60 2 3.51 4.08 2.90 104 Sdf Mollisols Alluvial deposits New 29

II1A 5.91 1.07 6 5.52 8.04 4.98 104 Sdf Mollisols Alluvial deposits Mature 29

II1A 4.58 0.73 2 4.48 5.41 3.96 104 Sdf Mollisols Alluvial deposits Old 29

II1A 5.17 1.34 20 3.20 8.04 2.90 149 Sdf Mollisols Alluvial deposits Guanica Forest 29

II1A 3.42 0.51 14 3.36 4.51 2.75 151 Sdf Mollisols Alluvial deposits 29

II1Ci 3.47 0.04 2 3.47 3.50 3.44 104 Sdf Mollisols Alluvial deposits 29

II1Ci 3.47 1 149 Sdf Mollisols Alluvial deposits Guanica Forest 29

Manganese (mg/kg) II1A 32 6 14 33 40 21 151 Sdf Mollisols Alluvial deposits 29

Nitrogen (%) II1A 1.25 0.13 2 1.26 1.38 1.10 104 Sdf Mollisols Alluvial deposits New 29
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

37

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 1.50 0.09 6 1.47 1.66 1.40 104 Sdf Mollisols Alluvial deposits Mature 29

II1A 1.44 0.02 2 1.44 1.46 1.41 104 Sdf Mollisols Alluvial deposits Old 29

II1A 1.45 0.14 20 1.45 1.66 1.10 149 Sdf Mollisols Alluvial deposits Guanica Forest 29

II1A 1.77 0.18 14 1.68 2.17 1.56 151 Sdf Mollisols Alluvial deposits 29

II1Ci 0.71 1 149 Sdf Mollisols Alluvial deposits Guanica Forest 29

Phosphorus (mg/g) II1A 0.55 0.16 2 0.55 0.70 0.41 104 Sdf Mollisols Alluvial deposits New 29

II1A 0.45 0.06 6 0.46 0.56 0.36 104 Sdf Mollisols Alluvial deposits Mature 29

II1A 0.44 0.01 2 0.44 0.45 0.42 104 Sdf Mollisols Alluvial deposits Old 29

II1A 0.47 0.09 20 29.96 0.70 0.36 149 Sdf Mollisols Alluvial deposits Guanica Forest 29

II1A 0.48 0.07 14 0.46 0.61 0.37 151 Sdf Mollisols Alluvial deposits 29

II1Ci 0.21 0.05 2 0.21 0.26 0.15 104 Sdf Mollisols Alluvial deposits 29

II1Ci 0.10 1 149 Sdf Mollisols Alluvial deposits Guanica Forest 29

Potassium (mg/g) II1A 24.23 1.30 2 24.31 25.43 22.87 104 Sdf Mollisols Alluvial deposits New 29

II1A 19.40 3.07 6 18.23 26.17 16.94 104 Sdf Mollisols Alluvial deposits Mature 29

II1A 16.18 0.76 2 16.17 16.95 15.43 104 Sdf Mollisols Alluvial deposits Old 29

II1A 19.72 3.57 20 30.96 26.17 15.43 149 Sdf Mollisols Alluvial deposits Guanica Forest 29

II1A 15.29 3.53 14 17.71 23.21 9.43 151 Sdf Mollisols Alluvial deposits 29

II1Ci 5.61 0.25 2 5.61 5.79 5.44 104 Sdf Mollisols Alluvial deposits 29

II1Ci 5.61 1 149 Sdf Mollisols Alluvial deposits Guanica Forest 29

Sulfur (%) II1A 0.18 0.02 14 0.18 0.21 0.15 151 Sdf Mollisols Alluvial deposits 29

Bromeliads Aluminum (mg/g) II6 0.81 0.04 2 0.81 0.84 0.79 135 Lmrf Ultisols Tuffac. sandstone 30

II6 0.50 1 93 Wflm Inceptisols Tuffac. sandstone 30

Ash (%) II6 4.53 1 93 Wflm Inceptisols Tuffac. sandstone 30

C/N II6 24 0 2 24 24 24 135 Lmrf Ultisols Tuffac. sandstone 30

Calcium (mg/g) II6 3.61 0.26 2 3.61 3.80 3.43 135 Lmrf Ultisols Tuffac. sandstone 30

II6 1.97 1 93 Wflm Inceptisols Tuffac. sandstone 30

Carbon (%) II6 46 0 2 46 46 46 135 Lmrf Ultisols Tuffac. sandstone 30

Iron (mg/g) II6 0.67 0.05 2 0.67 0.71 0.64 135 Lmrf Ultisols Tuffac. sandstone 30
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

38

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II6 0.33 1 93 Wflm Inceptisols Tuffac. sandstone 30

Magnesium (mg/g) II6 2.02 0.10 2 2.02 2.09 1.94 135 Lmrf Ultisols Tuffac. sandstone 30

II6 1.66 1 93 Wflm Inceptisols Tuffac. sandstone 30

Manganese (mg/g) II6 0.18 0.01 2 0.18 0.19 0.17 135 Lmrf Ultisols Tuffac. sandstone 30

II6 0.12 1 93 Wflm Inceptisols Tuffac. sandstone 30

Nitrogen (%) II6 1.95 0.02 2 1.95 1.96 1.93 135 Lmrf Ultisols Tuffac. sandstone 30

II6 0.39 1 93 Wflm Inceptisols Tuffac. sandstone 30

Phosphorus (mg/g) II6 1.07 0.06 2 1.07 1.11 1.02 135 Lmrf Ultisols Tuffac. sandstone 30

II6 0.19 1 93 Wflm Inceptisols Tuffac. sandstone 30

Potassium (mg/g) II6 12.05 0.74 2 12.05 12.57 11.52 135 Lmrf Ultisols Tuffac. sandstone 30

II6 6.33 1 93 Wflm Inceptisols Tuffac. sandstone 30

Sulfur (%) II6 0.57 0.01 2 0.57 0.58 0.57 135 Lmrf Ultisols Tuffac. sandstone 30

Bryophytes Aluminum (mg/g) II5 26.73 6.71 6 28.06 33.37 14.64 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 19.84 8.85 6 23.95 29.63 2.68 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 12.91 3.04 8 12.48 16.84 6.83 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 6.28 3.32 10 5.10 12.23 2.69 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 0.58 1.28 137 0.15 8.13 0.01 123 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 2.93 2.90 88 1.98 12.64 0.41 123 Wflm Inceptisols Tuffac. sandstone Humus 31

II5 2.60 2.88 51 1.90 18.77 0.22 123 Wflm Inceptisols Tuffac. sandstone Roots 31

II5 1.55 1.62 45 0.89 8.52 0.25 123 Wflm Inceptisols Tuffac. sandstone Bark and sapwood 31

Ash (%) II5 68.38 5.09 4 69.51 73.03 61.47 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 60.64 22.85 6 72.62 77.89 13.40 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 50.96 20.67 8 51.11 85.86 20.31 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 37 28 18.79 8 37.32 66.13 12.98 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

Calcium (mg/g) II5 1.52 1.14 6 1.17 3.72 0.64 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 1.02 0.40 6 0.94 1.74 0.59 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 1.53 0.81 8 1.29 2.81 0.41 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 2.38 1.15 10 2.27 4.44 0.84 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

39

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II5 7.66 4.82 137 6.93 23.52 1.21 123 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 9.21 3.95 93 9.18 22.05 1.92 123 Wflm Inceptisols Tuffac. sandstone Humus 31

II5 9.82 4.66 51 8.42 20.12 2.06 123 Wflm Inceptisols Tuffac. sandstone Roots 31

II5 8.24 5.36 45 6.68 26.63 2.09 123 Wflm Inceptisols Tuffac. sandstone Bark and sapwood 31

Iron (mg/g) II5 22.61 3.72 6 22.96 27.47 17.12 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 17.63 7.68 6 21.01 23.20 1.96 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 11.17 4.39 8 11.20 16.06 4.63 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 5.05 3.57 10 4.52 14.51 1.82 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 0.68 2.08 137 0.12 14.49 0.01 123 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 3.14 4.23 85 1.55 18.59 0.07 123 Wflm Inceptisols Tuffac. sandstone Humus 31

II5 2.98 4.13 51 1.72 27.31 0.07 123 Wflm Inceptisols Tuffac. sandstone Roots 31

II5 1.89 1.73 40 1.26 5.87 0.03 123 Wflm Inceptisols Tuffac. sandstone Bark and sapwood 31

Magnesium (mg/g) II5 2.51 0.45 6 2.59 3.09 1.72 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 2.07 0.35 6 2.17 2.35 1.31 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 1.34 0.32 8 1.35 1.71 0.72 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 1.30 0.57 10 1.19 2.14 0.53 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 1.79 1.38 137 1.41 5.29 0.10 123 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 4.33 2.42 90 4.04 9.75 0.28 123 Wflm Inceptisols Tuffac. sandstone Humus 31

II5 3.06 2.03 51 2.483 9.32 0.42 123 Wflm Inceptisols Tuffac. sandstone Roots 31

II5 3.68 1.67 45 3.54 8.77 0.77 123 Wflm Inceptisols Tuffac. sandstone Bark and sapwood 31

Manganese (mg/g) II5 0.62 0.16 6 0.61 0.86 0.39 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 0.49 0.25 6 0.50 0.92 0.08 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 0.23 0.09 8 0.24 0.34 0.08 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 0.33 0.41 10 0.18 1.42 0.04 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 0.25 0.27 137 0.18 1.49 0.01 123 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 0.65 0.72 85 0.35 2.84 0.01 123 Wflm Inceptisols Tuffac. sandstone Humus 31

II5 0.74 0.99 51 0.42 4.88 0.03 123 Wflm Inceptisols Tuffac. sandstone Roots 31

II5 0.62 0.88 44 0.21 3.59 0.02 123 Wflm Inceptisols Tuffac. sandstone Bark and sapwood 31

Nitrogen (%) II5 0.70 0.35 6 0.61 1.31 0.30 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

40

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II5 0.63 0.32 6 0.57 1.11 0.29 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 0.78 0.40 8 0.85 1.37 0.24 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 1.03 0.23 9 1.00 1.43 0.76 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 1.09 0.54 127 0.98 2.50 0.32 123 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 1.65 0.59 66 1.69 2.83 0.58 123 Wflm Inceptisols Tuffac. sandstone Humus 31

II5 1.85 0.58 40 1.78 3.08 0.80 123 Wflm Inceptisols Tuffac. sandstone Roots 31

II5 1.73 0.49 35 1.65 2.90 0.60 123 Wflm Inceptisols Tuffac. sandstone Bark and sapwood 31

Phosphorus (mg/g) II5 0.47 0.12 6 0.43 0.63 0.32 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 0.36 0.10 6 0.33 0.52 0.24 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 0.48 0.17 8 0.48 0.74 0.20 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 0.52 0.24 10 0.48 1.17 0.30 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 0.28 0.13 137 0.25 0.69 0.05 123 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 1.74 0.98 93 1.55 5.61 0.15 123 Wflm Inceptisols Tuffac. sandstone Humus 31

II5 0.94 0.41 51 0.83 2.26 0.28 123 Wflm Inceptisols Tuffac. sandstone Roots 31

II5 1.21 0.56 45 1.04 2.91 0.40 123 Wflm Inceptisols Tuffac. sandstone Bark and sapwood 31

Potassium (mg/g) II5 2.29 0.52 6 2.22 3.16 1.56 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 3.42 1.99 6 2.37 6.31 1.87 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 2.10 0.30 8 2.06 2.57 1.61 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 2.73 1.03 10 2.79 4.16 1.25 88 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 1.17 0.91 137 0.86 6.09 0.28 123 Wflm Inceptisols Tuffac. sandstone Bryophytes 31

II5 10.17 4.81 94 9.11 23.13 2.35 123 Wflm Inceptisols Tuffac. sandstone Humus 31

II5 2.50 1.81 51 1.75 9.09 0.61 123 Wflm Inceptisols Tuffac. sandstone Roots 31

II5 9.83 6.26 45 9.33 30.16 1.36 123 Wflm Inceptisols Tuffac. sandstone 31

Buchenavia tetraphylla Aluminum (mg/g) II9A 1.06 0.41 6 0.94 1.72 0.68 153 Swf Ultisols Tuffac. sandstone 32

Ash (%) II9A 4.80 0.46 5 4.71 5.46 4.21 153 Swf Ultisols Tuffac. sandstone 32

C/N II9A 42 2 6 42 45 40 153 Swf Ultisols Tuffac. sandstone 32

Calcium (mg/g) II9A 8.56 1.17 6 8.34 10.31 7.28 153 Swf Ultisols Tuffac. sandstone 32

Carbon (%) II9A 55 1 6 56 56 54 153 Swf Ultisols Tuffac. sandstone 32
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

41

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Iron (mg/g) II9A 0.87 0.34 6 0.77 1.42 0.56 153 Swf Ultisols Tuffac. sandstone 32

Magnesium (mg/g) II9A 1.12 0.04 6 1.12 1.18 1.08 153 Swf Ultisols Tuffac. sandstone 32

Manganese (mg/g) II9A 0.41 0.02 6 0.40 0.44 0.39 153 Swf Ultisols Tuffac. sandstone 32

Nitrogen (%) II9A 1.32 0.08 6 1.34 1.40 1.21 153 Swf Ultisols Tuffac. sandstone 32

Phosphorus (mg/g) II9A 0.18 0.02 6 0.18 0.22 0.17 153 Swf Ultisols Tuffac. sandstone 32

Potassium (mg/g) II9A 2.04 0.07 6 2.01 2.14 1.97 153 Swf Ultisols Tuffac. sandstone 32

Sodium (mg/g) II9A 0.19 0.04 6 0.20 0.24 0.14 153 Swf Ultisols Tuffac. sandstone 32

Bucida buceras Ash (%) II1A 13.71 0.68 5 13.49 14.80 13.04 126 Ultisols Siltstone,sandstone Urban forest, RP 33

II1C 1.20 0.06 5 1.18 1.26 1.11 126 Ultisols Siltstone,sandstone Urban forest, RP 33

II1Ci 15.76 0.83 6 15.40 17.38 15.23 126 Ultisols Siltstone,sandstone Urban forest, RP 33

C/N II1A 22 1 5 22 23 21 126 Ultisols Siltstone,sandstone Urban forest, RP 33

II1C 303 15 5 300 318 282 126 Ultisols Siltstone,sandstone Urban forest, RP 33

II1Ci 59 1 6 59 61 57 126 Ultisols Siltstone,sandstone Urban forest, RP 33

Calcium (mg/g) II1A 10.05 1.17 2 10.27 11.21 8.42 104 Sdf Mollisols Alluvial deposits 33

II1A 20.62 6.49 7 19.45 33.53 13.18 104 Sdf Mollisols Alluvial deposits New 33

II1A 25.19 4.82 2 25.07 29.82 20.80 104 Sdf Mollisols Alluvial deposits Mature 33

II1A 19.42 7.39 20 19.45 33.53 8.42 149 Sdf Mollisols Alluvial deposits Guanica Forest 33

II1Ci 52.11 0.87 2 51.97 53.26 51.25 104 Sdf Mollisols Alluvial deposits 33

II1Ci 52.11 0.87 4 51.97 53.26 51.25 149 Sdf Mollisols Alluvial deposits Guanica Forest 33

Carbon (%) II1A 45 0 5 45 45 45 126 Ultisols Siltstone,sandstone Urban forest, RP 33

II1C 51 0 5 51 51 51 126 Ultisols Siltstone,sandstone Urban forest, RP 33

II1Ci 43 1 6 43 44 42 126 Ultisols Siltstone,sandstone Urban forest, RP 33

Magnesium (mg/g) II1A 1.28 0.17 2 1.27 1.47 1.11 104 Sdf Mollisols Alluvial deposits 33

II1A 2.30 0.41 7 2.28 2.95 1.49 104 Sdf Mollisols Alluvial deposits New 33

II1A 2.39 0.19 2 2.46 2.54 2.12 104 Sdf Mollisols Alluvial deposits Mature 33

II1A 2.13 0.53 22 2.24 2.95 1.11 149 Sdf Mollisols Alluvial deposits Guanica Forest 33

II1Ci 0.97 0.16 2 0.98 1.12 0.80 104 Sdf Mollisols Alluvial deposits 33

II1Ci 0.97 0.16 4 0.98 1.12 0.80 149 Sdf Mollisols Alluvial deposits Guanica Forest 33
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

42

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Nitrogen (%) II1A 2.03 0.05 5 2.02 2.09 1.97 126 Urban forest, RP 33

II1A 0.80 0.12 2 0.80 0.91 0.68 104 Sdf Mollisols Alluvial deposits New 33

II1A 1.22 0.10 7 1.24 1.35 1.03 104 Sdf Mollisols Alluvial deposits Mature 33

II1A 1.10 0.06 2 1.10 1.16 1.03 104 Sdf Mollisols Alluvial deposits Old 33

II1A 1.11 0.18 22 1.15 1.35 0.68 149 Sdf Mollisols Alluvial deposits Guanica Forest 33

II1C 0.17 0.01 5 0.17 0.18 0.16 126 Ultisols Siltstone,sandstone Urban forest, RP 33

II1Ci 0.73 0.02 6 0.73 0.77 0.71 126 Ultisols Siltstone,sandstone Urban forest, RP 33

II1Ci 0.57 0.03 4 0.57 0.60 0.54 149 Sdf Mollisols Alluvial deposits Guanica Forest 33

Phosphorus (mg/g) II1A 0.64 0.16 2 0.64 0.79 0.47 104 Sdf Mollisols Alluvial deposits 33

II1A 0.53 0.12 7 0.54 0.66 0.30 104 Sdf Mollisols Alluvial deposits New 33

II1A 0.50 0.02 2 0.49 0.52 0.48 104 Sdf Mollisols Alluvial deposits Mature 33

II1A 0.54 0.12 22 0.53 0.79 0.30 149 Sdf Mollisols Alluvial deposits Guanica Forest 33

II1Ci 0.21 0.05 2 0.21 0.26 0.15 104 Sdf Mollisols Alluvial deposits 33

II1Ci 0.21 0.05 4 0.21 0.26 0.15 149 Sdf Mollisols Alluvial deposits Guanica Forest 33

Potassium (mg/g) II1A 10.16 1.08 2 10.26 11.21 8.92 104 Sdf Mollisols Alluvial deposits 33

II1A 10.86 2.47 7 10.51 15.45 6.91 104 Sdf Mollisols Alluvial deposits New 33

II1A 10.16 0.67 2 10.15 10.78 9.57 104 Sdf Mollisols Alluvial deposits Mature 33

II1A 10.60 2.03 22 10.51 15.45 6.91 149 Sdf Mollisols Alluvial deposits Guanica Forest 33

II1Ci 9.95 2.51 2 9.97 12.33 7.52 104 Sdf Mollisols Alluvial deposits 33

II1Ci 9.95 2.51 4 9.97 12.33 7.52 149 Sdf Mollisols Alluvial deposits Guanica Forest 33

Sulfur (%) II1A 0.19 0.01 5 0.19 0.20 0.18 126 Ultisols Siltstone,sandstone Urban forest, RP 33

II1C 0.04 0.06 5 0.04 0.05 0.03 126 Ultisols Siltstone,sandstone Urban forest, RP 33

II1Ci 0.09 0.01 6 0.09 0.10 0.08 126 Ultisols Siltstone,sandstone Urban forest, RP 33

Wood density (g/cc) II1C 0.66 0.03 5 0.67 0.69 0.62 126 Ultisols Siltstone,sandstone Urban forest, RP 33

Bursera simaruba Aluminum (mg/kg) II1A 42 11 10 43 62 29 151 Sdf Mollisols Alluvial deposits 34

II1A 77 1 102 Smf Limestone Tuffac. sandstone New Adult leaves 34

Ash (%) II1A 4.76 1 102 Smf Limestone Tuffac. sandstone New Adult leaves 34

II1A 7.47 0.84 8 7.66 8.60 6.04 151 Sdf Mollisols Alluvial deposits 34
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

43

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

C/N II1A 22 1 8 22 24 21 151 Sdf Mollisols Alluvial deposits 34

Calcium (mg/g) II1A 6.34 0.97 2 6.31 7.27 5.47 104 Sdf Mollisols Alluvial deposits 34

II1A 10.43 3.01 7 10.98 14.77 5.28 104 Sdf Mollisols Alluvial deposits 34

II1A 3.50 1 102 Smf Limestone Tuffac. sandstone New Adult leaves 34

II1A 9.41 3.19 16 6.80 14.77 5.28 149 Sdf Mollisols Alluvial deposits Guanica Forest 34

II1A 14.02 2.86 11 14.27 17.78 9.06 151 Sdf Mollisols Alluvial deposits 34

II1Ci 34.17 0.90 1 34.17 34.80 33.53 104 Sdf Mollisols Alluvial deposits 34

II1Ci 34.17 1 149 Sdf Mollisols Alluvial deposits Guanica Forest 34

Carbon (%) II1A 53 1 8 53 54 52 151 Sdf Mollisols Alluvial deposits 34

Cobalt (μg/g) II1A 1.00 1 102 Smf Limestone Tuffac. sandstone New Adult leaves 34

Iron (mg/g) II1A 0.06 0.02 11 0.06 0.08 0.01 151 Sdf Mollisols Alluvial deposits 34

Magnesium (mg/g) II1A 1.96 0.29 2 1.98 2.22 1.66 104 Sdf Mollisols Alluvial deposits Mature 34

II1A 2.09 0.59 7 2.23 2.68 0.80 104 Sdf Mollisols Alluvial deposits 34

II1A 4.35 1 102 Smf Limestone Tuffac. sandstone New Adult leaves 34

II1A 2.06 0.53 17 2.22 2.68 0.80 149 Sdf Mollisols Alluvial deposits Guanica Forest 34

II1A 2.46 0.29 11 2.44 3.05 1.92 151 Sdf Mollisols Alluvial deposits 34

II1Ci 0.64 0.01 1 0.64 0.65 0.63 104 Sdf Mollisols Alluvial deposits 34

II1Ci 0.64 1 149 Sdf Mollisols Alluvial deposits Guanica Forest 34

Manganese (mg/kg) II1A 31 9 10 28 46 16 151 Sdf Mollisols Alluvial deposits 34

II1A 40 1 102 Smf Limestone Tuffac. sandstone New Adult leaves 34

Nickel (μg/g) II1A 5.90 1 102 Smf Limestone Tuffac. sandstone New Adult leaves 34

Nitrogen (%) II1A 1.58 0.04 2 1.58 1.62 1.53 104 Sdf Mollisols Alluvial deposits Mature 34

II1A 1.79 0.22 7 1.87 1.96 1.30 104 Sdf Mollisols Alluvial deposits 34

II1A 0.53 1 102 Smf Limestone Tuffac. sandstone New Adult leaves 34

II1A 1.77 0.22 18 1.84 2.07 1.30 149 Sdf Mollisols Alluvial deposits Guanica Forest 34

II1A 2.44 0.13 8 2.46 2.54 2.14 151 Sdf Mollisols Alluvial deposits 34

II1Ci 0.31 1 149 Sdf Mollisols Alluvial deposits Guanica Forest 34

Phosphorus (mg/g) II1A 0.19 0.01 1 0.19 0.20 0.19 104 Sdf Mollisols Alluvial deposits New 34

II1A 0.82 0.07 2 0.82 0.88 0.76 104 Sdf Mollisols Alluvial deposits Mature 34
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

44

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 0.92 0.15 7 0.95 1.20 0.72 104 Sdf Mollisols Alluvial deposits 34

II1A 1.10 1 102 Lmrf Ultisols Serpentinite New Adult leaves 34

II1A 0.90 0.14 17 0.87 1.20 0.72 149 Sdf Mollisols Alluvial deposits Guanica Forest 34

II1A 0.79 0.12 11 0.74 1.01 0.63 151 Sdf Mollisols Alluvial deposits 34

II1Ci 0.19 1 149 Sdf Mollisols Alluvial deposits Guanica Forest 34

Potassium (mg/g) II1A 10.33 0.84 2 10.32 11.07 9.59 104 Sdf Mollisols Alluvial deposits Mature 34

II1A 12.15 2.29 7 11.23 17.13 10.11 104 Sdf Mollisols Alluvial deposits 34

II1A 10.26 1 102 Smf Limestone Tuffac. sandstone New Adult leaves 34

II1A 11.75 2.18 18 11.05 17.13 9.59 149 Sdf Mollisols Alluvial deposits Guanica Forest 34

II1A 9.76 2.30 11 9.07 15.70 7.48 151 Sdf Mollisols Alluvial deposits 34

II1Ci 22.28 0.46 1 22.28 22.60 21.96 104 Sdf Mollisols Alluvial deposits 34

II1Ci 22.28 1 149 Sdf Mollisols Alluvial deposits Guanica Forest 34

Sulfur (%) II1A 0.29 0.02 8 0.29 0.31 0.25 151 Sdf Mollisols Alluvial deposits 34

Wood density (g/cc) II1B 0.25 0.03 3 0.26 0.26 0.22 97 Wfs Inceptisols Tuffac. sandstone 34

Byrsonima lucida Aluminum (mg/kg) II1A 59 1 149 Sdf Mollisols Alluvial deposits Susua forest 35

II1A 14 1 102 Smf Limestone Tuffac. sandstone Adult 35

Ash (%) II1A 3.51 1 102 Smf Limestone Tuffac. sandstone Adult 35

II1A 4.33 1 149 Sdf Mollisols Alluvial deposits Susua forest 35

Calcium (mg/g) II1A 8.22 1 102 Smf Limestone Tuffac. sandstone Adult 35

II1A 7.23 1 149 Sdf Mollisols Alluvial deposits Susua forest 35

Cobalt (μg/g) II1A 1.00 1 102 Smf Limestone Tuffac. sandstone Adult 35

Iron (mg/kg) II1A 99 1 149 Sdf Mollisols Alluvial deposits Susua forest 35

II1A 35 1 102 Smf Limestone Tuffac. sandstone Adult 35

Magnesium (mg/g) II1A 3.54 1 102 Smf Limestone Tuffac. sandstone Adult 35

II1A 5.80 1 149 Sdf Mollisols Alluvial deposits Susua forest 35

Manganese (mg/kg) II1A 19 1 149 Sdf Mollisols Alluvial deposits Susua forest 35

II1A 19 1 102 Smf Limestone Tuffac. sandstone Adult 35

Nickel (μg/g) II1A 3.15 1 102 Smf Limestone Tuffac. sandstone Adult 35
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

45

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Nitrogen (%) II1A 1.24 1 102 Smf Limestone Tuffac. sandstone Adult 35

II1A 1.40 1 149 Sdf Mollisols Alluvial deposits Susua forest 35

Phosphorus (mg/g) II1A 0.47 1 102 Smf Limestone Tuffac. sandstone Adult 35

II1A 0.46 1 149 Sdf Mollisols Alluvial deposits Susua forest 35

Potassium (mg/g) II1A 7.59 1 102 Smf Limestone Tuffac. sandstone Adult 35

II1A 4.54 1 149 Sdf Mollisols Alluvial deposits Susua forest 35

Zinc (μg/g) II1A 4.84 1 149 Sdf Mollisols Alluvial deposits Susua forest 35

Byrsonima spicata Aluminum (mg/g) II1A 0.10 0.01 2 0.10 0.11 0.10 131 Wfs Mollisols Plutonic rocks 36

II1B 0.09 0.00 3 0.09 0.84 0.09 131 Wfs Mollisols Plutonic rocks 36

II1B 0.06 0.02 4 0.06 0.07 0.04 131 Wfs Mollisols Plutonic rocks 36

II1C 0.09 0.11 2 0.09 0.17 0.02 131 Wfs Mollisols Plutonic rocks 36

II1E 0.63 0.33 2 0.63 0.87 0.40 131 Wfs Mollisols Plutonic rocks 36

Ash (%) II1A 3.86 0.35 2 3.86 4.10 3.61 131 Wfs Mollisols Plutonic rocks 36

II1B 3.10 0.46 3 3.25 3.46 2.58 131 Wfs Mollisols Plutonic rocks 36

II1B 3.27 0.09 4 3.30 3.34 3.15 131 Wfs Mollisols Plutonic rocks 36

II1C 1.42 0.35 2 1.42 1.66 1.17 131 Wfs Mollisols Plutonic rocks 36

II1E 2.87 1.58 2 2.87 3.99 1.75 131 Wfs Mollisols Plutonic rocks 36

C/N II1A 23 1 2 23 23 22 131 Wfs Mollisols Plutonic rocks 36

II1B 52 0 3 52 53 52 131 Wfs Mollisols Plutonic rocks 36

II1B 35 3 4 36 37 31 131 Wfs Mollisols Plutonic rocks 36

II1C 84 8 2 84 90 79 131 Wfs Mollisols Plutonic rocks 36

II1E 88 4 2 88 90 85 131 Wfs Mollisols Plutonic rocks 36

Calcium (mg/g) II1A 5.62 0.82 2 5.62 6.20 5.04 131 Wfs Mollisols Plutonic rocks 36

II1B 5.52 1.45 3 6.29 6.42 3.85 131 Wfs Mollisols Plutonic rocks 36

II1B 4.96 0.16 4 4.95 5.14 4.79 131 Wfs Mollisols Plutonic rocks 36

II1C 2.38 0.39 2 2.38 2.65 2.10 131 Wfs Mollisols Plutonic rocks 36

II1E 3.88 1.06 2 3.88 4.62 3.13 131 Wfs Mollisols Plutonic rocks 36

Carbon (%) II1A 57 0 2 57 57 57 131 Wfs Mollisols Plutonic rocks 36
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

46

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1B 54 0 3 54 54 54 131 Wfs Mollisols Plutonic rocks 36

II1B 52 0 4 52 52 52 131 Wfs Mollisols Plutonic rocks 36

II1C 53 0 2 53 53 53 131 Wfs Mollisols Plutonic rocks 36

II1E 53 0 2 53 53 53 131 Wfs Mollisols Plutonic rocks 36

Iron (mg/g) II1A 0.10 0.01 2 0.10 0.10 0.09 131 Wfs Mollisols Plutonic rocks 36

II1B 0.11 0.01 3 0.10 0.12 0.10 131 Wfs Mollisols Plutonic rocks 36

II1B 0.05 0.02 4 0.04 0.08 0.03 131 Wfs Mollisols Plutonic rocks 36

II1C 0.07 0.04 2 0.07 0.10 0.04 131 Wfs Mollisols Plutonic rocks 36

II1E 0.53 0.22 2 0.53 0.68 0.38 131 Wfs Mollisols Plutonic rocks 36

Magnesium (mg/g) II1A 3.64 0.61 2 3.64 4.07 3.21 131 Wfs Mollisols Plutonic rocks 36

II1B 1.79 0.34 3 1.64 2.17 1.55 131 Wfs Mollisols Plutonic rocks 36

II1B 1.45 0.22 4 1.51 1.63 1.16 131 Wfs Mollisols Plutonic rocks 36

II1C 0.82 0.04 2 0.82 0.85 0.80 131 Wfs Mollisols Plutonic rocks 36

II1E 0.94 0.12 2 0.94 1.03 0.85 131 Wfs Mollisols Plutonic rocks 36

Manganese (mg/g) II1A 0.09 0.05 2 0.09 0.13 0.06 131 Wfs Mollisols Plutonic rocks 36

II1B 0.25 0.08 3 0.28 0.30 0.16 131 Wfs Mollisols Plutonic rocks 36

II1B 0.07 0.03 4 0.06 0.11 0.05 131 Wfs Mollisols Plutonic rocks 36

II1C 0.07 0.03 2 0.07 0.09 0.04 131 Wfs Mollisols Plutonic rocks 36

II1E 0.03 0.00 2 0.03 0.03 0.03 131 Wfs Mollisols Plutonic rocks 36

Nitrogen (%) II1A 2.49 0.06 2 2.49 2.53 2.44 131 Wfs Mollisols Plutonic rocks 36

II1B 1.04 0.01 3 1.03 1.05 1.03 131 Wfs Mollisols Plutonic rocks 36

II1B 1.49 0.11 4 1.46 1.64 1.40 131 Wfs Mollisols Plutonic rocks 36

II1C 0.64 0.06 2 0.64 0.68 0.59 131 Wfs Mollisols Plutonic rocks 36

II1E 0.61 0.02 2 0.61 0.62 0.59 131 Wfs Mollisols Plutonic rocks 36

Phosphorus (mg/g) II1A 0.54 0.04 2 0.54 0.57 0.51 131 Wfs Mollisols Plutonic rocks 36

II1B 0.26 0.02 3 0.26 0.28 0.25 131 Wfs Mollisols Plutonic rocks 36

II1B 0.23 0.01 4 0.22 0.25 0.22 131 Wfs Mollisols Plutonic rocks 36

II1C 0.11 0.01 2 0.11 0.12 0.10 131 Wfs Mollisols Plutonic rocks 36

II1E 0.11 0.01 2 0.11 0.12 0.10 131 Wfs Mollisols Plutonic rocks 36
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

47

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Potassium (mg/g) II1A 6.81 0.37 2 6.81 7.07 6.55 131 Wfs Mollisols Plutonic rocks 36

II1B 4.12 0.12 3 4.13 4.24 4.00 131 Wfs Mollisols Plutonic rocks 36

II1B 7.01 0.41 4 6.88 7.59 6.68 131 Wfs Mollisols Plutonic rocks 36

II1C 2.01 0.85 2 2.01 2.61 1.41 131 Wfs Mollisols Plutonic rocks 36

II1E 2.23 0.58 2 2.23 2.65 1.82 131 Wfs Mollisols Plutonic rocks 36

Sulfur (%) II1A 0.44 0.03 2 0.44 0.46 0.42 131 Wfs Mollisols Plutonic rocks 36

II1B 0.28 0.04 3 0.30 0.31 0.24 131 Wfs Mollisols Plutonic rocks 36

II1B 0.20 0.02 4 0.20 0.22 0.18 131 Wfs Mollisols Plutonic rocks 36

II1C 0.09 0.00 2 0.09 0.09 0.09 131 Wfs Mollisols Plutonic rocks 36

II1E 0.30 0.14 2 0.30 0.40 0.21 131 Wfs Mollisols Plutonic rocks 36

Byrsonima wadsworthii Aluminum (mg/g) II1A 0.23 1 93 Wflm Inceptisols Tuffac. sandstone 37

II1B 0.06 1 93 Wflm Inceptisols Tuffac. sandstone 37

II1C 0.08 1 93 Wflm Inceptisols Tuffac. sandstone 37

Ash (%) II1A 3.40 1 93 Wflm Inceptisols Tuffac. sandstone 37

II1B 6.33 1 93 Wflm Inceptisols Tuffac. sandstone 37

II1C 1.00 1 93 Wflm Inceptisols Tuffac. sandstone 37

C/N II1A 25 1 93 Wflm Inceptisols Tuffac. sandstone 37

II1B 56 1 93 Wflm Inceptisols Tuffac. sandstone 37

IIIC 286 1 93 Wflm Inceptisols Tuffac. sandstone 37

Calcium (mg/g) II1A 7.47 1 93 Wflm Inceptisols Tuffac. sandstone 37

II1B 24.69 1 93 Wflm Inceptisols Tuffac. sandstone 37

II1C 3.81 1 93 Wflm Inceptisols Tuffac. sandstone 37

Iron (mg/g) II1A 0.19 1 93 Wflm Inceptisols Tuffac. sandstone 37

II1B 0.08 1 93 Wflm Inceptisols Tuffac. sandstone 37

II1C 0.06 1 93 Wflm Inceptisols Tuffac. sandstone 37

Magnesium (mg/g) II1A 2.70 1 93 Wflm Inceptisols Tuffac. sandstone 37

II1B 1.01 1 93 Wflm Inceptisols Tuffac. sandstone 37

II1C 1.41 1 93 Wflm Inceptisols Tuffac. sandstone 37
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

48

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Manganese (mg/g) II1A 0.09 1 93 Wflm Inceptisols Tuffac. sandstone 37

II1B 0.06 1 93 Wflm Inceptisols Tuffac. sandstone 37

II1C 0.01 1 93 Wflm Inceptisols Tuffac. sandstone 37

Nitrogen (%) II1A 1.80 1 93 Wflm Inceptisols Tuffac. sandstone 37

II1B 0.78 1 93 Wflm Inceptisols Tuffac. sandstone 37

II1C 0.16 1 93 Wflm Inceptisols Tuffac. sandstone 37

Phosphorus (mg/g) II1A 0.68 1 93 Wflm Inceptisols Tuffac. sandstone 37

II1B 0.32 1 93 Wflm Inceptisols Tuffac. sandstone 37

II1C 0.12 1 93 Wflm Inceptisols Tuffac. sandstone 37

Potassium (mg/g) II1A 6.18 1 93 Wflm Inceptisols Tuffac. sandstone 37

II1B 3.89 1 93 Wflm Inceptisols Tuffac. sandstone 37

II1C l.54 1 93 Wflm Inceptisols Tuffac. sandstone 37

Calophyllum antillanum Aluminum (μg/g) II1A 48 1 58 Lmrf Ultisols Serpentinite 38

Ash (%) II1A 3.54 0.56 5 3.32 4.20 3.03 94 Swf Ultisols Tuffac. sandstone Understory 38

II1A 4.87 0.22 2 4.87 5.02 4.71 101 Wfs Ultisols Lava Overstory 38

II1A 3.16 0.11 6 3.17 3.27 3.02 126 Mfs Alfisol Alluvial deposits Vega Alta 38

II1B 4.18 0.99 3 4.34 5.09 3.12 94 Swf Ultisols Tuffac. sandstone 38

II1B 2.87 0.65 2 2.87 3.33 2.41 101 Wfs Ultisols Lava 38

II1C 3.11 1.08 4 3.41 3.86 1.52 94 Swf Ultisols Tuffac. sandstone 38

II1C 1.06 0.39 6 0.89 1.72 0.68 126 Mfs Alfisol Alluvial deposits Vega Alta 38

II1Ci 8.76 2.37 5 7.97 12.81 6.77 126 Mfs Alfisol Alluvial deposits Vega Alta 38

C/N II1A 48 2 6 48 51 47 126 Mfs Alfisol Alluvial deposits Vega Alta 38

II1C 636 199 6 612 870 428 126 Mfs Alfisol Alluvial deposits Vega Alta 38

II1Ci 145 20 5 152 160 109 126 Mfs Alfisol Alluvial deposits Vega Alta 38

Calcium (mg/g) II1A 5.58 1 101 Wfs Ultisols Lava Overstory 38

II1A 5.32 1 58 Lmrf Ultisols Serpentinite 38

II1B 5.89 2.07 2 5.89 7.36 4.43 101 Wfs Ultisols Lava 38

Carbon (%) II1A 55 0 6 55 55 54 126 Mfs Alfisol Alluvial deposits Vega Alta 38
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

49

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1C 52 0 6 52 52 51 126 Mfs Alfisol Alluvial deposits Vega Alta 38

II1Ci 54 2 5 54 56 52 126 Mfs Alfisol Alluvial deposits Vega Alta 38

Cobalt (μg/g) II1A 4.13 1 58 Lmrf Ultisols Serpentinite 38

Copper (μg/g) II1A 3.60 1 58 Lmrf Ultisols Serpentinite 38

Chromium (μg/g) II1A 2.33 1 58 Lmrf Ultisols Serpentinite 38

Iron (μg/g) II1A 51.38 1 58 Lmrf Ultisols Serpentinite 38

Magnesium (mg/g) II1A 3.28 1 58 Lmrf Ultisols Serpentinite 38

Manganese (μg/g) II1A 65.63 1 58 Lmrf Ultisols Serpentinite 38

Niquel (μg/g) II1A 24.34 1 58 Lmrf Ultisols Serpentinite 38

Nitrogen (%) II1A 1.21 0.03 2 1.21 1.23 1.19 101 Wfs Ultisols Lava Overstory 38

II1A 0.62 0.07 5 0.64 0.71 0.56 94 Swf Ultisols Tuffac. sandstone Understory 38

II1A 1.13 0.04 6 1.15 1.18 1.06 126 Mfs Alfisol Alluvial deposits Vega Alta 38

II1A 1.03 1 58 Lmrf Ultisols Serpentinite 38

II1B 0.61 0.31 3 0.46 0.96 0.40 94 Swf Ultisols Tuffac. sandstone 38

II1B 0.45 0.03 2 0.45 0.48 0.41 101 Wfs Ultisols Lava 38

II1C 0.56 0.34 4 0.44 1.08 0.14 94 Swf Ultisols Tuffac. sandstone 38

II1C 0.09 0.03 6 0.09 0.12 0.06 126 Mfs Alfisol Alluvial deposits Vega Alta 38

II1Ci 0.38 0.06 5 0.35 0.48 0.34 126 Mfs Alfisol Alluvial deposits Vega Alta 38

Phosphorus (mg/g) II1A 0.83 0.07 2 0.83 0.88 0.78 101 Wfs Ultisols Lava Overstory 38

II1A 0.62 0.07 5 0.64 0.71 0.55 94 Swf Ultisols Tuffac. sandstone 38

II1A 12.94 1 58 Lmrf Ultisols Serpentinite 38

II1B 0.42 0.10 3 0.38 0.54 0.35 94 Swf Ultisols Tuffac. sandstone 38

II1B 0.65 0.46 2 0.65 0.97 0.32 101 Wfs Ultisols Lava 38

II1C 0.61 0.44 4 0.48 1.16 0.10 94 Swf Ultisols Tuffac. sandstone 38

Potassium (mg/g) II1A 14.35 4.03 2 14.35 17.21 11.50 101 Wfs Ultisols Lava Overstory 38

II1A 5.10 0.80 5 5.40 6.00 4.30 94 Swf Ultisols Tuffac. sandstone Understory 38

II1A 6.58 1 58 Lmrf Ultisols Serpentinite 38

II1B 4.70 0.10 3 4.70 4.80 4.70 94 Swf Ultisols Tuffac. sandstone 38

II1B 8.87 1.65 2 8.87 10.04 7.71 101 Wfs Ultisols Lava 38
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

50

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1C 5.10 2.10 4 5.30 7.00 2.10 94 Swf Ultisols Tuffac. sandstone 38

Sulfur (%) II1A 0.20 0.02 6 0.19 0.24 0.18 126 Mfs Alfisol Alluvial deposits Vega Alta 38

II1C 0.02 0.04 6 0.02 0.03 0.02 126 Mfs Alfisol Alluvial deposits Vega Alta 38

II1Ci 0.11 0.02 5 0.11 0.13 0.09 126 Mfs Alfisol Alluvial deposits Vega Alta 38

Wood density (g/cc) II1C 0.53 0.05 5 0.52 0.61 0.49 126 Mfs Alfisol Alluvial deposits Vega Alta 38

Canna glauca Aluminum (mg/g) II3 0.28 1 135 Lmrf Ultisols Tuffac. sandstone 39

Calcium (mg/g) II3 3.61 1 135 Lmrf Ultisols Tuffac. sandstone 39

Iron (mg/g) II3 0.19 1 135 Lmrf Ultisols Tuffac. sandstone 39

Magnesium (mg/g) II3 4.19 1 135 Lmrf Ultisols Tuffac. sandstone 39

Manganese (mg/g) II3 2.64 1 135 Lmrf Ultisols Tuffac. sandstone 39

Nitrogen (%) II3 1.86 1 135 Lmrf Ultisols Tuffac. sandstone 39

Phosphorus (mg/g) II3 1.09 1 135 Lmrf Ultisols Tuffac. sandstone 39

Potassium (mg/g) II3 18.73 1 135 Lmrf Ultisols Tuffac. sandstone 39

Canna spp. Aluminum (mg/g) II3 0.17 0.04 2 0.17 0.20 0.14 135 Lmrf Ultisols Tuffac. sandstone 40

Calcium (mg/g) II3 4.82 0.38 2 4.82 5.09 4.56 135 Lmrf Ultisols Tuffac. sandstone 40

Iron (mg/g) II3 0.27 0.06 2 0.27 0.31 0.23 135 Lmrf Ultisols Tuffac. sandstone 40

Magnesium (mg/g) II3 3.32 0.76 2 3.32 3.85 2.78 135 Lmrf Ultisols Tuffac. sandstone 40

Manganese (mg/g) II3 0.90 0.72 2 0.90 1.41 0.39 135 Lmrf Ultisols Tuffac. sandstone 40

Nitrogen (%) II3 1.70 0.63 2 1.70 2.14 1.25 135 Lmrf Ultisols Tuffac. sandstone 40

Phosphorus (mg/g) II3 0.75 0.32 2 0.75 0.98 0.53 135 Lmrf Ultisols Tuffac. sandstone 40

Potassium (mg/g) II3 15.00 3.28 2 15.00 17.32 12.68 135 Lmrf Ultisols Tuffac. sandstone 40

Capparis cynophallophora Nitrogen (%) II1A 2.04 1 149 Sdf Mollisols Alluvial deposits Guanica Forest 41

Phosphorus (mg/g) II1A 0.51 1 104 Sdf Mollisols Alluvial deposits Mature 41

II1A 0.51 1 149 Sdf Mollisols Alluvial deposits Guanica Forest 41

Potassium (mg/g) II1A 24.37 1 104 Sdf Mollisols Alluvial deposits Mature 41

II1A 24.37 1 149 Sdf Mollisols Alluvial deposits Guanica Forest 41
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

51

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Casearia arborea Ash (%) II1A 7.05 3.31 8 5.68 11.20 3.52 94 Swf Ultisols Tuffac. sandstone Understory 42

II1A 4.72 0.59 2 4.72 5.13 4.30 101 Wfs Ultisols Lava Overstory 42

II1B 4.38 1.71 7 4.00 7.06 2.71 94 Swf Ultisols Tuffac. sandstone Understory 42

II1C 3.43 1.72 7 2.55 6.06 1.74 94 Swf Ultisols Tuffac. sandstone Understory 42

Calcium (mg/g) II1A 3.64 1.26 9 3.95 6.06 1.95 135 Lmrf Ultisols Tuffac. sandstone 42

II1B 2.46 1.08 11 2.33 4.25 0.51 135 Lmrf Ultisols Tuffac. sandstone 42

II1B 1.85 2.47 10 0.59 6.89 0.42 135 Lmrf Ultisols Tuffac. sandstone 42

II1F 2.00 1 135 Lmrf Ultisols Tuffac. sandstone 42

Magnesium (mg/g) II1A 2.83 0.67 9 2.89 3.79 1.74 135 Lmrf Ultisols Tuffac. sandstone 42

II1B 1.86 0.61 11 1.86 2.53 0.50 135 Lmrf Ultisols Tuffac. sandstone 42

II1B 0.95 0.71 10 0.63 2.60 0.48 135 Lmrf Ultisols Tuffac. sandstone 42

II1Ci 3.49 0.26 6 3.49 3.83 3.11 135 Lmrf Ultisols Tuffac. sandstone 42

II1F 2.43 1 135 Lmrf Ultisols Tuffac. sandstone 42

Nitrogen (%) II1A 2.18 0.14 2 2.18 2.28 2.08 101 Wfs Ultisols Lava Overstory 42

II1A 0.87 0.46 8 0.95 1.43 0.04 94 Swf Ultisols Tuffac. sandstone Understory 42

II1A 1.74 0.70 9 1.94 2.40 0.51 135 Lmrf Ultisols Tuffac. sandstone 42

II1B 1.03 0.16 7 1.06 1.26 0.77 94 Swf Ultisols Tuffac. sandstone Understory 42

II1B 1.30 0.62 11 1.35 2.13 0.41 135 Lmrf Ultisols Tuffac. sandstone 42

II1B 0.38 0.41 10 0.16 1.22 0.16 135 Lmrf Ultisols Tuffac. sandstone 42

II1C 0.74 0.20 7 0.78 0.94 0.33 94 Swf Ultisols Tuffac. sandstone Understory 42

II1Ci 1.04 1.06 6 1.03 1.13 0.98 135 Lmrf Ultisols Tuffac. sandstone 42

II1F 1.85 1 135 Lmrf Ultisols Tuffac. sandstone 42

Phosphorus (mg/g) II1A 0.86 0.13 2 0.86 0.95 0.76 101 Wfs Ultisols Lava Overstory 42

II1A 0.87 0.46 8 0.95 1.43 0.39 94 Swf Ultisols Tuffac. sandstone 42

II1A 0.76 0.36 9 0.68 1.34 0.26 135 Lmrf Ultisols Tuffac. sandstone 42

II1B 0.76 0.20 7 0.73 1.11 0.50 94 Swf Ultisols Tuffac. sandstone Understory 42

II1B 0.57 0.41 11 0.46 1.30 0.09 135 Lmrf Ultisols Tuffac. sandstone 42

II1B 0.20 0.20 10 0.10 0.67 0.09 135 Lmrf Ultisols Tuffac. sandstone 42
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

52

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1C 0.62 0.21 7 0.56 0.88 0.26 94 Swf Ultisols Tuffac. sandstone Understory 42

II1Ci 0.48 0.03 6 0.49 0.51 0.43 135 Lmrf Ultisols Tuffac. sandstone 42

II1F 1.21 1 135 Lmrf Ultisols Tuffac. sandstone 42

Potassium (mg/g) II1A 9.34 1.61 2 9.34 10.48 8.20 101 Wfs Ultisols Lava Overstory 42

II1A 18.68 12.60 8 16.00 33.49 1.49 94 Swf Ultisols Tuffac. sandstone Understory 42

II1A 6.23 2.56 9 5.598 11.28 3.32 135 Lmrf Ultisols Tuffac. sandstone 42

II1B 11.97 8.80 7 9.22 25.30 0.99 94 Swf Ultisols Tuffac. sandstone Understory 42

II1B 5.49 2.09 11 6.04 9.43 2.39 135 Lmrf Ultisols Tuffac. sandstone 42

II1B 4.11 2.02 10 3.15 8.42 2.66 135 Lmrf Ultisols Tuffac. sandstone 42

II1C 11.00 6.66 7 1.05 21.76 3.96 94 Swf Ultisols Tuffac. sandstone Understory 42

II1Ci 9.67 0.51 6 9.56 10.54 9.21 135 Lmrf Ultisols Tuffac. sandstone 42

II1F 8.56 1 135 Lmrf Ultisols Tuffac. sandstone 42

Casearia decandra Aluminum (mg/g) II1A 0.07 1 148 Smf Ultisols Tuffac. sandstone 43

Ash (%) II1A 7.73 0.24 5 7.66 8.12 7.53 126 Ultisols Siltstone,sandstone Urban forest, RP 43

II1A 5.54 1 148 Smf Ultisols Tuffac. sandstone 43

II1C 2.82 0.88 5 2.76 3.85 1.50 126 Ultisols Siltstone,sandstone Urban forest, RP 43

II1Ci 15.86 3.04 6 16.39 20.58 12.34 126 Ultisols Siltstone,sandstone Urban forest, RP 43

C/N II1A 16 1 5 16 17 14 126 Ultisols Siltstone,sandstone Urban forest, RP 43

II1A 24 1 148 Smf Ultisols Tuffac. sandstone 43

II1C 155 16 5 159 172 129 126 Ultisols Siltstone,sandstone Urban forest, RP 43

II1Ci 43 8 6 42 58 36 126 Ultisols Siltstone,sandstone Urban forest, RP 43

Calcium (mg/g) II1A 12.61 1 148 Smf Ultisols Tuffac. sandstone 43

Carbon (%) II1A 52 1 5 52 53 51 126 Ultisols Siltstone,sandstone Urban forest, RP 43

II1C 52 1 5 52 53 51 126 Ultisols Siltstone,sandstone Urban forest, RP 43

II1Ci 46 2 6 45 49 43 126 Ultisols Siltstone,sandstone Urban forest, RP 43

Cobalt (mg/kg) II1A 4 1 148 Smf Ultisols Tuffac. sandstone 43

Copper (mg/kg) II1A 3 1 148 Smf Ultisols Tuffac. sandstone 43

Chromium (mg/kg) II1A 2 1 148 Smf Ultisols Tuffac. sandstone 43
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

53

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Iron (mg/g) II1A 0.14 1 148 Smf Ultisols Tuffac. sandstone 43

Magnesium (mg/g) II1A 7.37 1 148 Smf Ultisols Tuffac. sandstone 43

Manganese (mg/g) II1A 0.08 1 148 Smf Ultisols Tuffac. sandstone 43

Niquel (mg/kg) II1A 11.02 1 148 Smf Ultisols Tuffac. sandstone 43

Nitrogen (%) II1A 3.22 0.20 5 3.19 3.55 3.02 126 Ultisols Siltstone,sandstone Urban forest, RP 43

II1A 1.83 1 148 Smf Ultisols Tuffac. sandstone 43

II1C 0.34 0.04 5 0.32 0.40 0.30 126 Ultisols Siltstone,sandstone Urban forest, RP 43

II1Ci 1.09 0.22 6 1.07 1.36 0.74 126 Ultisols Siltstone,sandstone Urban forest, RP 43

Phosphorus (mg/g) II1A 0.78 1 148 Smf Ultisols Tuffac. sandstone 43

Potassium (mg/g) II1A 2.86 1 148 Smf Ultisols Tuffac. sandstone 43

Sodium (mg/g) II1A 65.23 1 148 Smf Ultisols Tuffac. sandstone 43

Sulfur (%) II1A 0.22 0.01 5 0.23 0.24 0.20 126 Ultisols Siltstone,sandstone Urban forest, RP 43

II1C 0.05 0.01 5 0.05 0.06 0.03 126 Ultisols Siltstone,sandstone Urban forest, RP 43

II1Ci 0.10 0.02 6 0.09 0.18 0.14 126 Ultisols Siltstone,sandstone Urban forest, RP 43

Wood density (g/cc) II1C 0.63 0.07 5 0.62 0.72 0.55 126 Ultisols Siltstone,sandstone Urban forest, RP 43

Zinc (mg/kg) II1A 13 1 148 Smf Ultisols Tuffac. sandstone 43

Casearia guianensis Ash (%) II1A 7.68 0.24 6 7.67 8.06 7.32 126 Ultisols Siltstone,sandstone Urban forest, RP 44

II1C 3.03 0.38 6 2.91 3.68 2.66 126 Ultisols Siltstone,sandstone Urban forest, RP 44

II1Ci 16.65 2.57 5 16.23 20.62 13.86 126 Ultisols Siltstone,sandstone Urban forest, RP 44

C/N II1A 21 1 6 21 22 20 126 Ultisols Siltstone,sandstone Urban forest, RP 44

II1C 115 9 6 118 123 99 126 Ultisols Siltstone,sandstone Urban forest, RP 44

II1Ci 29 3 5 27 34 27 126 Ultisols Siltstone,sandstone Urban forest, RP 44

Carbon (%) II1A 50 1 6 50 51 49 126 Ultisols Siltstone,sandstone Urban forest, RP 44

II1C 51 0 6 51 52 51 126 Ultisols Siltstone,sandstone Urban forest, RP 44

II1Ci 45 1 5 45 47 43 126 Ultisols Siltstone,sandstone Urban forest, RP 44

Nitrogen (%) II1A 2.40 0.05 6 2.42 2.45 2.31 126 Ultisols Siltstone,sandstone Urban forest, RP 44

II1C 0.45 0.04 6 0.44 0.52 0.42 126 Ultisols Siltstone,sandstone Urban forest, RP 44

II1Ci 1.58 0.15 5 1.66 1.68 1.32 126 Ultisols Siltstone,sandstone Urban forest, RP 44
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

54

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Sulfur (%) II1A 0.19 0.01 6 0.19 0.20 0.18 126 Ultisols Siltstone,sandstone Urban forest, RP 44

II1C 0.07 0.01 6 0.07 0.08 0.06 126 Ultisols Siltstone,sandstone Urban forest, RP 44

II1Ci 0.13 0.01 5 0.13 0.14 0.11 126 Ultisols Siltstone,sandstone Urban forest, RP 44

Wood density (g/cc) II1C 0.60 0.07 5 0.60 0.70 0.51 126 Ultisols Siltstone,sandstone Urban forest, RP 44

Casearia sylvestris Aluminum (mg/g) II1A 0.06 1 148 Smf Ultisols Tuffac. sandstone 45

Ash (%) II1A 21.94 10.94 6 23.52 32.81 4.37 94 Swf Ultisols Tuffac. sandstone 45

II1A 8.46 1 148 Smf Ultisols Tuffac. sandstone 45

II1B 5.77 1.34 7 6.05 7.36 3.48 94 Swf Ultisols Tuffac. sandstone 45

II1C 3.55 1.05 7 3.09 5.22 2.64 94 Swf Ultisols Tuffac. sandstone 45

C/N II1A 21 1 148 Smf Ultisols Tuffac. sandstone 45

Calcium (mg/g) II1A 6.54 1 148 Smf Ultisols Tuffac. sandstone 45

Cobalt (mg/kg) II1A 2 1 148 Smf Ultisols Tuffac. sandstone 45

Copper (mg/kg) II1A 2 1 148 Smf Ultisols Tuffac. sandstone 45

Chromium (mg/kg) II1A 3 1 148 Smf Ultisols Tuffac. sandstone 45

Iron (mg/kg) II1A 54 1 148 Smf Ultisols Tuffac. sandstone 45

Magnesium (mg/g) II1A 13.50 1 148 Smf Ultisols Tuffac. sandstone 45

Manganese (mg/kg) II1A 16 1 148 Smf Ultisols Tuffac. sandstone 45

Niquel (mg/kg) II1A 9 1 148 Smf Ultisols Tuffac. sandstone 45

Nitrogen (%) II1A 1.48 0.47 6 1.39 2.21 0.95 94 Swf Ultisols Tuffac. sandstone 45

II1A 2.04 1 148 Smf Ultisols Tuffac. sandstone 45

II1B 1.34 0.30 7 1.31 1.77 0.89 94 Swf Ultisols Tuffac. sandstone 45

II1C 0.65 0.22 7 0.65 1.08 0.44 94 Swf Ultisols Tuffac. sandstone 45

Phosphorus (mg/g) II1A 1.48 0.48 6 1.39 2.21 0.95 94 Swf Ultisols Tuffac. sandstone 45

II1A 1.08 1 148 Smf Ultisols Tuffac. sandstone 45

II1B 1.09 0.41 7 1.01 1.86 0.68 94 Swf Ultisols Tuffac. sandstone 45

II1C 0.64 0.29 7 0.77 1.05 0.30 94 Swf Ultisols Tuffac. sandstone 45

Potassium (mg/g) II1A 21.94 10.94 6 23.52 32.81 4.37 94 Swf Ultisols Tuffac. sandstone 45

II1A 4.73 1 148 Smf Ultisols Tuffac. sandstone 45
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

55

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1B 14.28 5.46 7 14.62 22.40 7.98 94 Swf Ultisols Tuffac. sandstone 45

II1C 7.30 3.90 7 6.13 15.50 3.98 94 Swf Ultisols Tuffac. sandstone 45

Sodium (mg/g) II1A 82.16 1 148 Smf Ultisols Tuffac. sandstone 45

Zinc (mg/kg) II1A 41 1 148 Smf Ultisols Tuffac. sandstone 45

Cassine xylocarpa Aluminum (mg/kg) II1A 28 1 149 Sdf Mollisols Alluvial deposits Susua forest 46

II1A 35 1 102 Smf Limestone Tuffac. sandstone Adult 46

Ash (%) II1A 7.46 1 102 Smf Limestone Tuffac. sandstone Adult 46

II1A 8.71 1 149 Sdf Mollisols Alluvial deposits Susua forest 46

Calcium (mg/g) II1A 7.50 1 104 Sdf Mollisols Alluvial deposits 46

II1A 13.54 1 102 Smf Limestone Tuffac. sandstone Adult 46

II1A 16.19 1 149 Sdf Mollisols Alluvial deposits Susua forest 46

II1A 7.50 1 149 Sdf Mollisols Alluvial deposits Guanica Forest 46

II1Ci 28.19 1 104 Sdf Mollisols Alluvial deposits 46

II1Ci 28.19 1 149 Sdf Mollisols Alluvial deposits Guanica Forest 46

Cobalt (μg/g) II1A 1.10 1 102 Smf Limestone Tuffac. sandstone Adult 46

Iron (mg/kg) II1A 100 1 149 Sdf Mollisols Alluvial deposits Susua forest 46

II1A 39 1 102 Smf Limestone Tuffac. sandstone Adult 46

Magnesium (mg/g) II1A 0.99 1 104 Sdf Mollisols Alluvial deposits 46

II1A 7.35 1 102 Smf Limestone Tuffac. sandstone Adult 46

II1A 4.51 1 149 Sdf Mollisols Alluvial deposits Susua forest 46

II1A 0.99 1 149 Sdf Mollisols Alluvial deposits Guanica Forest 46

II1Ci 0.54 1 104 Sdf Mollisols Alluvial deposits 46

II1Ci 0.54 1 149 Sdf Mollisols Alluvial deposits Guanica Forest 46

Manganese (mg/kg) II1A 10 1 149 Sdf Mollisols Alluvial deposits Susua forest 46

II1A 8 1 102 Smf Limestone Tuffac. sandstone Adult 46

Nickel (μg/g) II1A 4.45 1 102 Smf Limestone Tuffac. sandstone Adult 46

Nitrogen (%) II1A 0.60 1 102 Smf Limestone Tuffac. sandstone Adult 46

II1A 0.79 1 149 Sdf Mollisols Alluvial deposits Susua forest 46
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

56

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 1.03 1 149 Sdf Mollisols Alluvial deposits Guanica Forest 46

II1Ci 0.60 1 149 Sdf Mollisols Alluvial deposits Guanica Forest 46

Phosphorus (mg/g) II1A 0.31 1 149 Sdf Mollisols Alluvial deposits Susua forest 46

II1Ci 0.08 1 104 Sdf Mollisols Alluvial deposits 46

II1Ci 78.00 1 149 Sdf Mollisols Alluvial deposits Guanica Forest 46

Potassium (mg/g) II1A 11.70 1 104 Sdf Mollisols Alluvial deposits 46

II1A 6.04 1 102 Smf Limestone Tuffac. sandstone Adult 46

II1A 6.25 1 149 Sdf Mollisols Alluvial deposits Susua forest 46

II1A 11.70 1 149 Sdf Mollisols Alluvial deposits Guanica Forest 46

II1Ci 6.03 1 104 Sdf Mollisols Alluvial deposits 46

II1Ci 6.03 1 149 Sdf Mollisols Alluvial deposits Guanica Forest 46

Zinc (μg/g) II1A 6.51 1 149 Sdf Mollisols Alluvial deposits Susua forest 46

Casuarina equisetifolia Aluminum (mg/g) II8 2.88 2.59 20 1.74 7.81 0.49 152 Mfs Histosol Alluvial deposits 47

Ash (%) II8 23.03 21.02 20 13.90 76.13 5.27 152 Mfs Histosol Alluvial deposits 47

C/N II8 34 9 20 32 50 21 152 Mfs Histosol Alluvial deposits 47

Calcium (mg/g) II8 15.92 3.88 20 16.07 21.48 9.80 152 Mfs Histosol Alluvial deposits Organic layer 47

Carbon (%) II8 45 13 20 51 54 17 152 Mfs Histosol Alluvial deposits 47

Iron (mg/g) II8 4.49 4.84 20 2.19 14.17 0.51 152 Mfs Histosol Alluvial deposits 47

Magnesium (mg/g) II8 2.54 0.91 20 2.27 40.01 1.34 152 Mfs Histosol Alluvial deposits 47

Manganese (mg/g) II8 0.20 0.12 20 0.14 0.41 0.07 152 Mfs Histosol Alluvial deposits 47

Nitrogen (%) II8 1.33 0.39 20 1.31 1.99 0.68 152 Mfs Histosol Alluvial deposits 47

Phosphorus (mg/g) II8 0.23 0.07 20 0.22 0.34 0.11 152 Mfs Histosol Alluvial deposits 47

Potassium (mg/g) II8 1.46 0.18 20 1.48 1.79 1.03 152 Mfs Histosol Alluvial deposits 47

Sulfur (%) II8 0.30 0.09 20 0.32 0.45 0.15 152 Mfs Histosol Alluvial deposits 47

Cecropia schreberiana Aluminum (mg/g) II1A 0.19 0.45 29 0.09 2.52 0.04 135 Lmrf Ultisols Tuffac. sandstone 48

II1A 0.55 0.50 4 0.47 1.11 0.13 141 Wflm Ultisols Tuffac. sandstone 48

II1A 2.69 1 129 Lmrf Ultisols Tuffac. sandstone 48
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

57

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 0.17 1 129 Lmrf Ultisols Tuffac. sandstone Petiole 48

II1A 0.23 0.33 13 0.13 1.32 0.07 107 Swf Ultisols Tuffac. sandstone 48

II1A 0.17 0.16 11 0.10 0.49 0.03 135 Lmrf Ultisols Tuffac. sandstone 48

II1C 1.09 0.19 49 0.32 8.13 0.02 123 Wflm Inceptisols Tuffac. sandstone Wood decay 48

II1C 0.07 0.08 30 0.04 0.38 0.02 135 Lmrf Ultisols Tuffac. sandstone 48

II1C 0.15 1 141 Wflm Ultisols Tuffac. sandstone 48

II1C 0.04 0.05 8 0.02 0.14 0.01 135 Lmrf Ultisols Tuffac. sandstone 48

II1E 1.40 0.79 19 1.42 3.31 0.23 135 Lmrf Ultisols Tuffac. sandstone 48

II1E 2.48 2.08 3 1.88 4.80 0.77 141 Wflm Ultisols Tuffac. sandstone 48

II1H 0.13 0.08 2 0.13 0.19 0.08 135 Lmrf Ultisols Tuffac. sandstone 48

II8A 0.41 0.25 2 0.41 0.59 0.23 146 Swf Ultisols Tuffac. sandstone 48

II8B 0.18 0.18 2 0.18 0.31 0.06 146 Swf Ultisols Tuffac. sandstone 48

Ash (%) II1A 9.43 1 129 Lmrf Ultisols Tuffac. sandstone 48

II1A 7.95 1 129 Lmrf Ultisols Tuffac. sandstone Petiole 48

II1A 10.41 1.51 9 10.17 13.58 9.12 107 Swf Ultisols Tuffac. sandstone 48

II8A 9.16 2.05 2 9.16 10.61 7.71 146 Swf Ultisols Tuffac. sandstone 48

II8B 4.40 1.93 2 4.40 5.76 3.03 146 Swf Ultisols Tuffac. sandstone 48

C/N II1A 19 1 129 Lmrf Ultisols Tuffac. sandstone 48

II1A 60 1 129 Lmrf Ultisols Tuffac. sandstone Petiole 48

II1A 13 4 13 13 21 9 107 Swf Ultisols Tuffac. sandstone 48

II8A 19 6 2 19 23 15 146 Swf Ultisols Tuffac. sandstone 48

II8B 51 16 2 51 62 39 146 Swf Ultisols Tuffac. sandstone 48

Calcium (mg/g) II1A 8.22 3.13 31 8.22 16.04 2.53 135 Lmrf Ultisols Tuffac. sandstone 48

II1A 11.95 0.88 4 11.74 13.17 11.12 141 Wflm Ultisols Tuffac. sandstone 48

II1A 12.16 1 129 Lmrf Ultisols Tuffac. sandstone 48

II1A 14.15 1 129 Lmrf Ultisols Tuffac. sandstone Petiole 48

II1A 10.41 2.86 13 9.74 17.21 6.00 107 Swf Ultisols Tuffac. sandstone 48

II1A 5.41 2.69 11 5.83 9.25 2.01 135 Lmrf Ultisols Tuffac. sandstone 48

II1B 3.45 0.00 2 3.45 3.46 3.45 135 Lmrf Ultisols Tuffac. sandstone 48
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

58

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1B 0.90 0.14 9 0.84 1.15 0.77 135 Lmrf Ultisols Tuffac. sandstone 48

II1C 7.39 3.14 49 7.63 14.68 2.03 123 Wflm Inceptisols Tuffac. sandstone Wood decay 48

II1C 2.57 0.77 30 2.39 4.35 1.30 135 Lmrf Ultisols Tuffac. sandstone 48

II1C 5.10 1 141 Wflm Ultisols Tuffac. sandstone 48

II1C 2.91 2.28 8 1.98 8.06 0.99 135 Lmrf Ultisols Tuffac. sandstone 48

II1Ci 7.66 1.09 9 7.46 9.15 6.12 135 Lmrf Ultisols Tuffac. sandstone 48

II1E 1.61 0.66 19 1.49 2.89 0.75 135 Lmrf Ultisols Tuffac. sandstone 48

II1E 2.07 1.21 3 1.37 3.47 1.36 141 Wflm Ultisols Tuffac. sandstone 48

II1H 7.56 1.32 2 7.56 8.49 6.62 135 Lmrf Ultisols Tuffac. sandstone 48

II8A 18.68 0.06 2 18.68 18.72 18.64 146 Swf Ultisols Tuffac. sandstone 48

II8B 9.11 0.34 2 9.11 9.35 8.87 146 Swf Ultisols Tuffac. sandstone 48

Carbon (%) II1A 51 1 129 Lmrf Ultisols Tuffac. sandstone 48

II1A 50 1 129 Lmrf Ultisols Tuffac. sandstone Petiole 48

II1A 43 1 27 43 46 42 150 Swf Ultisols Tuffac. sandstone 48

II1A 42 3 13 43 46 37 107 Swf Ultisols Tuffac. sandstone 48

II8A 51 4 2 51 54 48 146 Swf Ultisols Tuffac. sandstone 48

II8B 53 2 2 53 54 51 146 Swf Ultisols Tuffac. sandstone 48

Iron (mg/g) II1A 0.20 0.58 29 0.07 3.19 0.02 135 Lmrf Ultisols Tuffac. sandstone 48

II1A 1.18 1.83 4 0.36 3.91 0.10 141 Wflm Ultisols Tuffac. sandstone 48

II1A 3.00 1 129 Lmrf Ultisols Tuffac. sandstone 48

II1A 0.18 1 129 Lmrf Ultisols Tuffac. sandstone Petiole 48

II1A 0.27 0.32 13 0.16 1.32 0.12 107 Swf Ultisols Tuffac. sandstone 48

II1A 0.15 0.05 11 0.14 0.27 0.07 135 Lmrf Ultisols Tuffac. sandstone 48

II1C 1.48 3.26 49 0.30 14.49 0.05 123 Wflm Inceptisols Tuffac. sandstone Wood decay 48

II1C 0.12 0.23 30 0.05 1.08 0.03 135 Lmrf Ultisols Tuffac. sandstone 48

II1C 0.12 1 141 Wflm Ultisols Tuffac. sandstone 48

II1C 0.11 0.05 8 0.10 0.21 0.05 135 Lmrf Ultisols Tuffac. sandstone 48

II1E 1.42 0.86 19 1.37 3.54 0.20 135 Lmrf Ultisols Tuffac. sandstone 48

II1E 1.83 1.38 3 1.49 3.35 0.65 141 Wflm Ultisols Tuffac. sandstone 48
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

59

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1H 0.12 0.08 2 0.12 0.17 0.06 135 Lmrf Ultisols Tuffac. sandstone 48

II8A 0.27 0.21 2 0.27 0.42 0.12 146 Swf Ultisols Tuffac. sandstone 48

II8B 0.13 0.12 2 0.13 0.21 0.04 146 Swf Ultisols Tuffac. sandstone 48

Magnesium (mg/g) II1A 4.69 1.33 31 5.00 7.40 1.99 135 Lmrf Ultisols Tuffac. sandstone 48

II1A 5.68 2.04 4 5.92 7.51 3.39 141 Wflm Ultisols Tuffac. sandstone 48

II1A 1.89 1 129 Lmrf Ultisols Tuffac. sandstone 48

II1A 3.41 1 129 Lmrf Ultisols Tuffac. sandstone Petiole 48

II1A 6.43 1.49 13 6.08 9.55 4.59 107 Swf Ultisols Tuffac. sandstone 48

II1A 2.23 0.95 11 1.85 4.05 0.93 135 Lmrf Ultisols Tuffac. sandstone 48

II1B 3.72 0.03 2 3.72 3.74 3.70 135 Lmrf Ultisols Tuffac. sandstone 48

II1B 1.43 0.19 9 1.39 1.75 1.22 135 Lmrf Ultisols Tuffac. sandstone 48

II1C 2.24 1.20 49 2.76 5.28 1.14 123 Wflm Inceptisols Tuffac. sandstone Wood decay 48

II1C 2.88 1.18 30 2.57 4.85 1.08 135 Lmrf Ultisols Tuffac. sandstone 48

II1C 6.93 1 141 Wflm Ultisols Tuffac. sandstone 48

II1C 1.71 0.93 8 1.431 3.68 0.64 135 Lmrf Ultisols Tuffac. sandstone 48

II1Ci 3.69 0.41 9 3.67 4.26 3.10 135 Lmrf Ultisols Tuffac. sandstone 48

II1E 3.49 0.90 19 3.33 5.24 1.70 135 Lmrf Ultisols Tuffac. sandstone 48

II1E 5.27 1.16 3 5.71 6.17 3.98 141 Wflm Ultisols Tuffac. sandstone 48

II1H 4.73 1.07 2 4.73 5.49 3.98 135 Lmrf Ultisols Tuffac. sandstone 48

II8A 6.77 0.83 2 6.77 7.36 6.19 146 Swf Ultisols Tuffac. sandstone 48

II8B 5.05 4.28 2 5.05 8.08 2.03 146 Swf Ultisols Tuffac. sandstone 48

Manganese (mg/g) II1A 0.19 0.10 29 0.18 0.44 0.05 135 Lmrf Ultisols Tuffac. sandstone 48

II1A 0.65 0.14 4 0.67 0.78 0.49 141 Wflm Ultisols Tuffac. sandstone 48

II1A 0.11 1 129 Lmrf Ultisols Tuffac. sandstone 48

II1A 0.27 1 129 Lmrf Ultisols Tuffac. sandstone Petiole 48

II1A 0.25 0.10 13 0.29 0.40 0.08 107 Swf Ultisols Tuffac. sandstone 48

II1A 0.10 0.03 11 0.10 0.15 0.05 135 Lmrf Ultisols Tuffac. sandstone 48

II1C 0.30 0.20 49 0.24 1.00 0.06 123 Wflm Inceptisols Tuffac. sandstone Wood decay 48

II1C 0.24 0.11 30 0.24 0.51 0.06 135 Lmrf Ultisols Tuffac. sandstone 48
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

60

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1C 0.49 1 141 Wflm Ultisols Tuffac. sandstone 48

II1C 0.14 0.11 8 0.11 0.38 0.02 135 Lmrf Ultisols Tuffac. sandstone 48

II1E 0.16 0.08 19 0.18 0.31 0.06 135 Lmrf Ultisols Tuffac. sandstone 48

II1E 0.14 0.07 3 0.10 0.22 0.10 141 Wflm Ultisols Tuffac. sandstone 48

II1H 0.32 0.19 2 0.32 0.46 0.18 135 Lmrf Ultisols Tuffac. sandstone 48

II8A 0.24 0.10 2 0.24 0.31 0.17 146 Swf Ultisols Tuffac. sandstone 48

II8B 0.30 0.11 2 0.30 0.38 0.23 146 Swf Ultisols Tuffac. sandstone 48

Nitrogen (%) II1A 2.52 0.50 30 2.52 3.44 1.60 135 Lmrf Ultisols Tuffac. sandstone 48

II1A 2.21 0.19 4 2.22 2.42 1.96 141 Wflm Ultisols Tuffac. sandstone 48

II1A 2.74 1 129 Lmrf Ultisols Tuffac. sandstone 48

II1A 0.83 1 129 Lmrf Ultisols Tuffac. sandstone Petiole 48

II1A 3.11 0.52 27 3.01 4.42 2.43 150 Swf Ultisols Tuffac. sandstone 48

II1A 3.38 0.82 13 3.30 4.72 1.99 107 Swf Ultisols Tuffac. sandstone 48

II1A 1.44 0.87 11 1.72 2.62 0.34 135 Lmrf Ultisols Tuffac. sandstone 48

II1B 0.42 0.02 2 0.42 0.43 0.40 135 Lmrf Ultisols Tuffac. sandstone 48

II1B 0.11 0.02 9 0.11 0.13 0.08 135 Lmrf Ultisols Tuffac. sandstone 48

II1C 1.41 0.51 39 1.29 2.50 0.63 123 Wflm Inceptisols Tuffac. sandstone Wood decay 48

II1C 0.62 0.21 30 0.61 1.15 0.23 135 Lmrf Ultisols Tuffac. sandstone 48

II1C 0.67 1 141 Wflm Ultisols Tuffac. sandstone 48

II1C 0.67 0.58 8 0.46 2.04 0.21 135 Lmrf Ultisols Tuffac. sandstone 48

II1Ci 0.73 0.06 9 0.70 0.82 0.65 135 Lmrf Ultisols Tuffac. sandstone 48

II1E 0.59 0.13 19 0.56 0.84 0.39 135 Lmrf Ultisols Tuffac. sandstone 48

II1E 0.59 0.11 3 0.54 0.71 0.51 141 Wflm Ultisols Tuffac. sandstone 48

II1H 1.43 0.07 2 1.43 1.48 1.38 135 Lmrf Ultisols Tuffac. sandstone 48

II8A 2.80 1.03 2 2.80 3.52 2.07 146 Swf Ultisols Tuffac. sandstone 48

II8B 1.11 0.41 2 1.11 1.40 0.82 146 Swf Ultisols Tuffac. sandstone 48

Phosphorus (mg/g) II1A 1.50 0.57 31 1.34 3.04 0.69 135 Lmrf Ultisols Tuffac. sandstone 48

II1A 1.49 0.27 4 1.38 1.90 1.32 141 Wflm Ultisols Tuffac. sandstone 48

II1A 0.62 1 129 Lmrf Ultisols Tuffac. sandstone 48
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

61

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 0.41 1 129 Lmrf Ultisols Tuffac. sandstone Petiole 48

II1A 1.82 0.84 13 1.77 3.57 0.58 107 Swf Ultisols Tuffac. sandstone 48

II1A 0.69 0.36 11 0.73 1.33 0.22 135 Lmrf Ultisols Tuffac. sandstone 48

II1B 0.35 0.01 2 0.35 0.36 0.34 135 Lmrf Ultisols Tuffac. sandstone 48

II1B 0.07 0.01 9 0.07 0.08 0.06 135 Lmrf Ultisols Tuffac. sandstone 48

II1C 0.26 0.09 49 0.27 0.58 0.14 123 Wflm Inceptisols Tuffac. sandstone Wood decay 48

II1C 0.39 0.15 30 0.36 0.87 0.18 135 Lmrf Ultisols Tuffac. sandstone 48

II1C 0.65 1 141 Wflm Ultisols Tuffac. sandstone 48

II1C 0.41 0.27 8 0.30 0.93 0.17 135 Lmrf Ultisols Tuffac. sandstone 48

II1Ci 0.28 0.03 9 0.28 0.33 0.24 135 Lmrf Ultisols Tuffac. sandstone 48

II1E 0.44 0.10 19 0.43 0.60 0.27 135 Lmrf Ultisols Tuffac. sandstone 48

II1E 0.69 0.24 3 0.60 0.96 0.50 141 Wflm Ultisols Tuffac. sandstone 48

II1H 0.82 0.16 2 0.83 0.94 0.71 135 Lmrf Ultisols Tuffac. sandstone 48

II8A 1.05 0.14 2 1.05 1.14 0.95 146 Swf Ultisols Tuffac. sandstone 48

II8B 0.65 0.00 2 0.65 0.65 0.64 146 Swf Ultisols Tuffac. sandstone 48

Potassium (mg/g) II1A 28.60 11.25 31 28.50 61.15 7.96 135 Lmrf Ultisols Tuffac. sandstone 48

II1A 12.44 2.83 4 12.52 15.81 8.90 141 Wflm Ultisols Tuffac. sandstone 48

II1A 5.48 1 129 Lmrf Ultisols Tuffac. sandstone 48

II1A 15.79 1 129 Lmrf Ultisols Tuffac. sandstone Petiole 48

II1A 24.78 9.68 13 20.75 47.00 9.35 107 Swf Ultisols Tuffac. sandstone 48

II1A 12.20 3.58 11 11.33 17.04 7.42 135 Lmrf Ultisols Tuffac. sandstone 48

II1B 7.43 0.06 2 7.43 7.47 7.38 135 Lmrf Ultisols Tuffac. sandstone 48

II1B 2.13 0.73 9 1.85 3.91 1.60 135 Lmrf Ultisols Tuffac. sandstone 48

II1C 0.94 0.56 49 0.76 3.22 0.28 123 Wflm Inceptisols Tuffac. sandstone 48

II1C 17.24 9.78 30 13.82 49.34 5.87 135 Lmrf Ultisols Tuffac. sandstone 48

II1C 9.19 1 141 Wflm Ultisols Tuffac. sandstone 48

II1C 9.47 3.40 8 8.70 16.92 5.85 135 Lmrf Ultisols Tuffac. sandstone 48

II1Ci 8.21 0.64 9 8.03 9.36 7.44 135 Lmrf Ultisols Tuffac. sandstone 48

II1E 10.89 3.99 19 12.78 16.95 5.03 135 Lmrf Ultisols Tuffac. sandstone 48
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

62

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1E 7.15 0.26 3 7.27 7.32 6.85 141 Wflm Ultisols Tuffac. sandstone 48

II1H 33.07 0.72 2 33.07 33.58 32.56 135 Lmrf Ultisols Tuffac. sandstone 48

II8A 7.06 7.24 2 7.06 12.18 1.95 146 Swf Ultisols Tuffac. sandstone 48

II8B 7.57 7.94 2 7.57 13.18 1.96 146 Swf Ultisols Tuffac. sandstone 48

Sulfur (%) II1A 0.37 1 129 Lmrf Ultisols Tuffac. sandstone 48

II1A 0.29 1 129 Lmrf Ultisols Tuffac. sandstone Petiole 48

II1A 0.38 0.07 17 0.36 0.52 0.28 150 Swf Ultisols Tuffac. sandstone 48

Cecropia sp. Aluminum (mg/g) II8A 1.73 3.16 65 0.66 16.87 0.13 146 Swf Ultisols Tuffac. sandstone

Ash (%) II8A 9.41 3.14 63 8.69 24.83 4.47 146 Swf Ultisols Tuffac. sandstone

Calcium (mg/g) II8A 18.17 4.80 65 17.79 27.89 2.18 146 Swf Ultisols Tuffac. sandstone

Iron (mg/g) II8A 1.71 3.96 65 0.49 24.09 0.08 146 Swf Ultisols Tuffac. sandstone

Magnesium (mg/g) II8A 3.18 0.75 65 3.08 5.21 1.68 146 Swf Ultisols Tuffac. sandstone

Manganese (mg/g) II8A 0.17 0.10 65 0.14 0.48 0.07 146 Swf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II8A 0.56 0.22 65 0.51 1.04 0.23 146 Swf Ultisols Tuffac. sandstone

Potassium (mg/g) II8A 3.74 2.20 65 2.87 12.77 1.48 146 Swf Ultisols Tuffac. sandstone

Cissampelos pareira Aluminum (mg/g) II4 0.03 1 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II4 6.39 1 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II4 0.08 1 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II4 2.18 1 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II4 0.04 1 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II4 0.90 1 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II4 0.48 1 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II4 11.63 1 135 Lmrf Ultisols Tuffac. sandstone

Cissus erosa Aluminum (mg/g) II4 0.19 0.25 7 0.10 0.75 0.01 135 Lmrf Ultisols Tuffac. sandstone

II4 0.20 0.10 2 0.20 0.27 0.13 135 Lmrf Ultisols Tuffac. sandstone

C/N II4 27 5 2 27 31 23 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

63

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Calcium (mg/g) II4 13.84 0.10 7 13.54 17.28 10.74 135 Lmrf Ultisols Tuffac. sandstone

II4 12.56 1.78 2 12.56 13.81 11.30 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II4 43 1 2 43 44 42 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II4 0.05 0.06 7 0.02 0.16 0.01 135 Lmrf Ultisols Tuffac. sandstone

II4 0.17 0.05 2 0.17 0.21 0.14 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II4 3.69 2.18 7 2.81 8.58 2.55 135 Lmrf Ultisols Tuffac. sandstone

II4 2.51 0.17 2 2.51 2.63 2.39 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II4 0.37 0.22 7 0.37 0.61 0.08 135 Lmrf Ultisols Tuffac. sandstone

II4 0.28 0.13 2 0.28 0.37 0.19 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II4 2.07 0.41 7 2.28 2.39 1.44 135 Lmrf Ultisols Tuffac. sandstone

II4 1.72 0.49 4 1.66 2.30 1.25 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II4 2.34 0.43 7 2.45 2.81 1.63 135 Lmrf Ultisols Tuffac. sandstone

II4 1.70 0.02 2 1.70 1.71 1.68 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II4 25.57 7.65 7 28.48 32.38 13.56 135 Lmrf Ultisols Tuffac. sandstone

II4 21.20 0.54 2 21.20 21.58 20.82 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II4 0.32 0.00 2 0.32 0.32 0.31 135 Lmrf Ultisols Tuffac. sandstone

Cissus verticillata Aluminum (mg/g) II4 0.20 0.10 2 0.20 0.27 0.13 135 Lmrf Ultisols Tuffac. sandstone

C/N II4 27 5 2 27 31 23 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II4 12.56 1.78 2 12.56 13.82 11.30 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II4 43 1 2 43 44 42 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II4 0.17 0.05 2 0.17 0.21 0.14 135 Lmrf Ultisols Tuffac. sandstone

Magnessium (mg/g) II4 2.51 0.17 2 2.51 2.63 2.39 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II4 0.28 0.13 2 0.28 0.37 0.19 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II4 1.72 0.49 4 1.66 2.30 1.25 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II4 1.70 0.02 2 1.70 1.71 1.68 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II4 21.20 0.54 2 21.20 21.58 20.82 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II4 0.32 0.00 2 0.32 0.32 0.31 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

64

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Citrus sinensis Ash (%) II1A 20.68 1 101 Wfs Ultisols Lava Sabana

II1B 3.02 0.58 2 3.02 3.43 2.61 101 Wfs Ultisols Lava Sabana

Calcium (mg/g) II1A 16.63 1.18 2 16.63 17.46 15.79 101 Wfs Ultisols Lava Sabana

II1B 10.69 2.20 2 10.69 12.24 9.13 101 Wfs Ultisols Lava Sabana

Nitrogen (%) II1A 1.13 0.34 2 1.13 1.37 0.89 101 Wfs Ultisols Lava Sabana

II1B 0.54 0.13 2 0.54 0.64 0.45 101 Wfs Ultisols Lava Sabana

Phosphorus (mg/g) II1B 0.51 0.16 2 0.51 0.62 0.40 101 Wfs Ultisols Lava Sabana

II1B 0.68 0.01 3 0.68 0.69 0.67 97 Wfs Inceptisols Tuffac. sandstone Sabana

Potassium (mg/g) II1A 2.26 0.17 2 2.26 2.38 2.14 101 Wfs Ultisols Lava Sabana

II1B 6.45 0.18 2 6.45 6.58 3.32 101 Wfs Ultisols Lava Sabana

Clibadium erosum Aluminum (mg/g) II2H 0.12 1 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II2H 11.61 1 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II2H 0.11 1 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II2H 2.72 1 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II2H 0.05 1 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II2H 1.92 1 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II2H 1.98 1 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II2H 41.35 1 135 Lmrf Ultisols Tuffac. sandstone

Clusia rosea Aluminum (mg/kg) II1A 38 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 39 1 102 Smf Limestone Tuffac. sandstone Old 

Ash (%) II1A 5.91 1 71 Wfs Inceptisols Tuffac. sandstone

II1A 5.83 1 149 Sdf Mollisols Alluvial deposits Susua forest

Calcium (mg/g) II1A 7.28 1 102 Smf Limestone Tuffac. sandstone Old 

II1A 11.98 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1Ci 25.35 1 104 Sdf Mollisols Alluvial deposits

II1Ci 25.35 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Carbon (%) II1A 55 1 71 Wfs Inceptisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

65

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Cobalt (μg/g) II1A 1.00 1 102 Smf Limestone Tuffac. sandstone Old 

Iron (mg/g) II1A 0.54 1 102 Smf Limestone Tuffac. sandstone Old 

II1A 0.19 1 149 Sdf Mollisols Alluvial deposits Susua forest

Magnesium (mg/g) II1A 1.96 1 102 Smf Limestone Tuffac. sandstone Old 

II1A 5.06 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1Ci 0.55 1 104 Sdf Mollisols Alluvial deposits

II1Ci 0.55 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Manganese (mg/kg) II1A 166 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 80 1 102 Smf Limestone Tuffac. sandstone Old 

Nickel (μg/g) II1A 7.70 1 102 Smf Limestone Tuffac. sandstone Old 

Nitrogen (%) II1A 0.60 1 102 Smf Limestone Tuffac. sandstone Old 

II1A 1.00 1 71 Wfs Inceptisols Tuffac. sandstone

II1A 0.52 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 0.50 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 0.07 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Phosphorus (mg/g) II1A 0.22 1 104 Sdf Mollisols Alluvial deposits Mature

II1A 0.29 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 0.22 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 0.07 1 104 Sdf Mollisols Alluvial deposits

II1Ci 0.07 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Potassium (mg/g) II1A 7.75 1 104 Sdf Mollisols Alluvial deposits Mature

II1A 3.11 1 102 Smf Limestone Tuffac. sandstone Old

II1A 7.22 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 7.75 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 6.94 1 104 Sdf Mollisols Alluvial deposits

II1Ci 6.94 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Sulfur (%) II1A 0.18 1 71 Wfs Inceptisols Tuffac. sandstone

Wood density (g/cc) II1A 0.68 0.01 3 0.68 0.69 0.67 97 Wfs Inceptisols Tuffac. sandstone

Zinc (μg/g) II1A 24 1 149 Sdf Mollisols Alluvial deposits Susua forest
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

66

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Cnemidaria horrida Aluminum (mg/g) II6 4.67 1.42 11 4.97 6.81 2.22 135 Lmrf Ultisols Tuffac. sandstone

C/N II6 23 4 9 23 33 19 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II6 2.47 1.15 11 2.30 5.59 1.38 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II6 41 1 9 41 43 39 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II6 0.17 0.09 11 0.17 0.28 0.05 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II6 2.96 0.82 11 3.11 4.26 1.98 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II6 0.10 0.09 11 0.06 0.32 0.02 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II6 1.73 0.30 19 1.68 2.19 1.23 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II6 0.89 0.31 11 0.90 1.33 0.41 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II6 19.79 7.90 11 20.18 31.20 0.68 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II6 0.34 0.09 9 0.316 0.44 0.20 135 Lmrf Ultisols Tuffac. sandstone

Coccoloba microstachya Aluminum (mg/kg) II1A 40 1 102 Smf Limestone Tuffac. sandstone Adult 

Ash (%) II1A 2.70 1 102 Smf Limestone Tuffac. sandstone Adult 

II1A 13.80 1.31 4 13.64 15.26 12.65 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1A 8.21 5.46 18 6.90 16.45 2.05 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1A 14.91 1 104 Sdf Mollisols Alluvial deposits

II1A 12.69 1 104 Sdf Mollisols Alluvial deposits Mature

II1A 4.53 3.72 5 3.77 14.24 2.05 104 Sdf Mollisols Alluvial deposits New

II1A 16.29 1 104 Sdf Mollisols Alluvial deposits Old

II1A 2.21 1 102 Smf Limestone Tuffac. sandstone Adult 

Cobalt (μg/g) II1A 0.60 1 102 Smf Limestone Tuffac. sandstone Adult 

Iron (mg/kg) II1A 52 1 102 Smf Limestone Tuffac. sandstone Adult 

Magnesium (mg/g) II1A 1.98 0.15 4 1.97 2.16 1.81 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1A 1.88 0.78 18 1.72 3.05 0.57 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1A 1.97 1 104 Sdf Mollisols Alluvial deposits

II1A 1.99 1 104 Sdf Mollisols Alluvial deposits Mature

II1A 1.88 0.92 5 1.67 3.05 0.57 104 Sdf Mollisols Alluvial deposits New
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

67

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 3.00 1 104 Sdf Mollisols Alluvial deposits Old

II1A 3.47 1 102 Smf Limestone Tuffac. sandstone Adult 

Manganese (mg/kg) II1A 50 1 102 Smf Limestone Tuffac. sandstone Adult 

Nickel (μg/g) II1A 25 1 102 Smf Limestone Tuffac. sandstone Adult 

Nitrogen (%) II1A 0.62 0.11 4 0.63 0.71 0.49 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1A 0.85 0.23 18 0.84 1.24 0.54 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1A 0.52 1 104 Sdf Mollisols Alluvial deposits

II1A 0.71 1 104 Sdf Mollisols Alluvial deposits Mature

II1A 1.01 0.20 5 0.95 1.27 0.71 104 Sdf Mollisols Alluvial deposits New

II1A 0.55 1 104 Sdf Mollisols Alluvial deposits Old

II1A 1.02 1 102 Smf Limestone Tuffac. sandstone Adult 

Phosphorus (mg/g) II1A 0.37 0.07 4 0.38 0.43 0.30 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1A 0.50 0.17 18 0.42 0.35 0.31 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1A 0.31 1 104 Sdf Mollisols Alluvial deposits

II1A 0.43 1 104 Sdf Mollisols Alluvial deposits Mature

II1A 0.61 1 102 Smf Limestone Tuffac. sandstone Adult 

II1A 0.61 0.16 5 0.63 0.84 0.40 104 Sdf Mollisols Alluvial deposits New

II1A 0.32 1 104 Sdf Mollisols Alluvial deposits Old

Potassium (mg/g) II1A 6.80 0.15 4 6.78 6.99 6.64 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1A 7 1 18 7 8 6 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1A 6.68 1 104 Sdf Mollisols Alluvial deposits

II1A 6.92 1 104 Sdf Mollisols Alluvial deposits Mature

II1A 6.87 0.66 5 6.85 7.90 5.54 104 Sdf Mollisols Alluvial deposits New

II1A 6.82 1 104 Sdf Mollisols Alluvial deposits Old

II1A 10.06 1 102 Smf Limestone Tuffac. sandstone Adult 

Coccoloba diversifolia Aluminum (mg/g) II1A 0.06 0.01 10 0.07 0.07 0.05 151 Sdf Mollisols Alluvial deposits

Ash (%) II1A 10.68 1.01 9 10.84 11.81 8.18 151 Sdf Mollisols Alluvial deposits

C/N II1A 30 2 10 31 33 27 151 Sdf Mollisols Alluvial deposits
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

68

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Calcium (mg/g) II1A 28.45 7.26 10 30.61 33.96 8.73 151 Sdf Mollisols Alluvial deposits

II1Ci 3.21 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 43.21 1 104 Sdf Mollisols Alluvial deposits

Carbon (%) II1A 51 0 10 51 51 50 151 Sdf Mollisols Alluvial deposits

Iron (mg/g) II1A 0.05 0.01 10 0.05 0.08 0.04 151 Sdf Mollisols Alluvial deposits

Magnesium (mg/g) II1A 3.05 0.44 10 3.13 3.69 2.23 151 Sdf Mollisols Alluvial deposits

II1Ci 2.04 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 2.04 1 104 Sdf Mollisols Alluvial deposits

Manganese (mg/g) II1A 0.03 0.01 10 0.04 0.04 0.02 151 Sdf Mollisols Alluvial deposits

Nitrogen (%) II1A 1.47 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1A 1.69 0.11 10 1.67 1.88 1.55 151 Sdf Mollisols Alluvial deposits

II1Ci 0.39 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Phosphorus (mg/g) II1A 0.47 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1A 0.68 0.07 10 0.68 0.79 0.58 151 Sdf Mollisols Alluvial deposits

II1A 0.47 1 104 Sdf Mollisols Alluvial deposits

II1Ci 0.27 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 0.27 1 104 Sdf Mollisols Alluvial deposits

Potassium (mg/g) II1A 12.31 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1A 7.94 1.00 10 9.22 9.66 6.12 151 Sdf Mollisols Alluvial deposits

II1A 12.31 1 104 Sdf Mollisols Alluvial deposits

II1Ci 7.48 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 7.48 1 104 Sdf Mollisols Alluvial deposits

Sulfur (%) II1A 0.33 0.05 10 0.31 0.42 0.27 151 Sdf Mollisols Alluvial deposits

Cocoloba microstachya Aluminum (mg/g) II1A 0.03 0.02 15 0.03 0.06 0.01 151 Sdf Mollisols Alluvial deposits

Ash (%) II1A 6.76 3.50 15 5.75 19.26 5.13 151 Sdf Mollisols Alluvial deposits

C/N II1A 36 2 15 35 40 33 151 Sdf Mollisols Alluvial deposits

Calcium (mg/g) II1A 14.74 4.24 15 14.19 29.04 10.11 151 Sdf Mollisols Alluvial deposits

Carbon (%) II1A 52 1 15 52 55 51 151 Sdf Mollisols Alluvial deposits
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

69

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Iron (mg/g) II1A 0.04 0.01 15 0.04 0.07 0.03 151 Sdf Mollisols Alluvial deposits

Magnesium (mg/g) II1A 2.58 0.17 15 2.59 2.84 2.31 151 Sdf Mollisols Alluvial deposits

Manganese (mg/g) II1A 0.01 0.01 15 0.01 0.04 0.01 151 Sdf Mollisols Alluvial deposits

Nitrogen (%) II1A 1.46 0.11 15 1.47 1.60 1.30 151 Sdf Mollisols Alluvial deposits

Phosphorus (mg/g) II1A 0.59 0.05 15 0.58 0.67 0.51 151 Sdf Mollisols Alluvial deposits

Potassium (mg/g) II1A 4.57 0.42 15 4.57 5.25 4.02 151 Sdf Mollisols Alluvial deposits

Sulfur (%) II1A 0.28 0.03 15 0.29 0.32 0.22 151 Sdf Mollisols Alluvial deposits

Coffea arabica Aluminum (mg/g) II1A 0.12 0.08 44 0.10 0.48 0.05 53 Wflm Ultisols Tuffac. sandstone

II1A 0.07 0.01 3 0.07 0.08 0.06 131 Wfs Mollisols Plutonic rocks

II1C 0.08 0.08 26 0.06 0.35 0.01 53 Wflm Ultisols Tuffac. sandstone

II1C 0.05 0.02 3 0.04 0.07 0.04 131 Wfs Mollisols Plutonic rocks

II1E 0.87 0.47 3 0.63 1.42 0.57 131 Wfs Mollisols Plutonic rocks

Ash (%) II1A 14.21 0.37 2 14.21 14.47 13.95 94 Wfs Ultisols Lava

II1A 9.40 0.71 44 9.55 10.57 7.79 53 Wflm Ultisols Tuffac. sandstone

II1A 7.42 0.35 3 7.60 7.64 7.02 131 Wfs Mollisols Plutonic rocks

II1B 3.54 0.28 2 3.54 3.74 3.34 94 Wfs Ultisols Lava

II1C 2.31 1 94 Wfs Ultisols Lava

II1C 4.19 0.82 26 4.08 5.62 2.57 53 Wflm Ultisols Tuffac. sandstone

II1C 1.80 0.10 3 1.76 1.91 1.72 131 Wfs Mollisols Plutonic rocks

II1E 3.21 1.40 3 2.84 4.75 2.03 131 Wfs Mollisols Plutonic rocks

C/N II1A 14 2 44 14 18 10 53 Wflm Ultisols Tuffac. sandstone

II1A 16 1 3 16 17 16 131 Wfs Mollisols Plutonic rocks

II1C 37 8 26 36 55 25 53 Wflm Ultisols Tuffac. sandstone

II1C 64 7 3 62 72 58 131 Wfs Mollisols Plutonic rocks

II1E 47 6 3 49 52 40 131 Wfs Mollisols Plutonic rocks

Calcium (mg/g) II1A 12.54 3.36 44 11.96 22.25 7.15 53 Wflm Ultisols Tuffac. sandstone

II1A 7.41 1.58 3 7.07 9.13 6.02 131 Wfs Mollisols Plutonic rocks

II1C 6.45 0.87 26 6.35 8.29 4.84 53 Wflm Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

70

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1C 2.95 0.29 3 3.06 3.17 2.62 131 Wfs Mollisols Plutonic rocks

II1E 1.67 0.30 3 1.77 1.91 1.33 131 Wfs Mollisols Plutonic rocks

Carbon (%) II1A 51 1 44 51 53 48 53 Wflm Ultisols Tuffac. sandstone

II1A 52 1 3 51 57 51 131 Wfs Mollisols Plutonic rocks

II1C 51 1 26 51 52 49 53 Wflm Ultisols Tuffac. sandstone

II1C 52 0 3 52 52 51 131 Wfs Mollisols Plutonic rocks

II1E 53 0 3 53 53 53 131 Wfs Mollisols Plutonic rocks

Iron (mg/g) II1A 0.17 0.09 44 0.13 0.45 0.05 53 Wflm Ultisols Tuffac. sandstone

II1A 0.07 0.01 3 0.07 0.08 0.06 131 Wfs Mollisols Plutonic rocks

II1C 0.12 0.09 26 0.85 0.49 0.03 53 Wflm Ultisols Tuffac. sandstone

II1C 0.04 0.02 3 0.03 0.07 0.03 131 Wfs Mollisols Plutonic rocks

II1E 0.80 0.54 3 0.58 1.41 0.41 131 Wfs Mollisols Plutonic rocks

Magnesium (mg/g) II1A 3.33 0.63 4 3.26 4.81 2.30 53 Wflm Ultisols Tuffac. sandstone

II1A 4.46 1.14 3 4.40 5.63 3.34 131 Wfs Mollisols Plutonic rocks

II1C 1.32 0.30 26 1.24 2.06 0.87 53 Wflm Ultisols Tuffac. sandstone

II1C 0.54 0.04 3 0.52 0.58 0.51 131 Wfs Mollisols Plutonic rocks

II1E 0.87 0.38 3 0.80 1.27 0.53 131 Wfs Mollisols Plutonic rocks

Manganese (mg/g) II1A 0.38 0.18 44 0.34 0.89 0.10 53 Wflm Ultisols Tuffac. sandstone

II1A 0.14 0.05 3 0.12 0.20 0.11 131 Wfs Mollisols Plutonic rocks

Manganese (mg/g) II1C 0.10 0.05 6 0.09 0.26 0.04 53 Wflm Ultisols Tuffac. sandstone

II1C 0.02 0.01 3 0.02 0.02 0.01 131 Wfs Mollisols Plutonic rocks

II1E 0.02 0.02 3 0.01 0.04 0.01 131 Wfs Mollisols Plutonic rocks

Nitrogen (%) II1A 1.12 0.13 2 1.12 1.21 1.03 94 Wfs Ultisols Lava

II1A 3.66 0.44 44 3.57 4.80 2.87 53 Wflm Ultisols Tuffac. sandstone

II1A 3.13 0.19 3 3.15 3.31 2.93 131 Wfs Mollisols Plutonic rocks

II1B 0.92 0.05 2 0.92 0.95 0.88 94 Wfs Ultisols Lava

II1C 0.61 1 94 Wfs Ultisols Lava

II1C 1.44 0.28 26 1.44 2.05 0.95 53 Wflm Ultisols Tuffac. sandstone

II1C 0.82 0.08 3 0.84 0.88 0.73 131 Wfs Mollisols Plutonic rocks
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

71

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1E 1.14 0.16 3 1.09 1.32 1.02 131 Wfs Mollisols Plutonic rocks

Phosphorus (mg/g) II1A 1.63 0.32 44 1.61 2.59 1.05 53 Wflm Ultisols Tuffac. sandstone

II1A 0.54 0.06 3 0.54 0.60 0.48 131 Wfs Mollisols Plutonic rocks

II1B 0.71 0.04 2 0.71 0.74 0.68 94 Wfs Ultisols Lava

II1C 1.01 0.45 26 0.86 2.20 0.35 53 Wflm Ultisols Tuffac. sandstone

II1C 0.15 0.02 3 0.15 0.17 0.14 131 Wfs Mollisols Plutonic rocks

II1E 0.23 0.03 3 0.22 0.26 0.21 131 Wfs Mollisols Plutonic rocks

Potassium (mg/g) II1A 14.21 0.37 2 14.21 14.47 13.95 94 Wfs Ultisols Lava

II1A 21.00 6.85 44 19.48 37.45 2.92 53 Wflm Ultisols Tuffac. sandstone

II1A 19.48 2.61 3 20.30 21.57 16.56 131 Wfs Mollisols Plutonic rocks

II1B 6.88 0.06 2 6.88 6.92 6.84 94 Wfs Ultisols Lava

II1C 3.67 1 94 Wfs Ultisols Lava

II1C 7.84 2.74 26 6.98 14.39 3.29 53 Wflm Ultisols Tuffac. sandstone

II1C 3.77 0.28 3 3.66 4.08 3.55 131 Wfs Mollisols Plutonic rocks

II1E 4.38 1.01 3 4.43 5.36 3.34 131 Wfs Mollisols Plutonic rocks

Sulfur (%) II1A 0.51 0.20 44 0.50 0.94 0.26 53 Wflm Ultisols Tuffac. sandstone

II1A 0.47 0.04 3 0.47 0.51 0.43 131 Wfs Mollisols Plutonic rocks

II1C 0.32 0.06 26 0.30 0.48 0.23 53 Wflm Ultisols Tuffac. sandstone

II1C 0.10 0.02 3 0.09 0.12 0.09 131 Wfs Mollisols Plutonic rocks

II1E 0.22 0.08 3 0.19 0.31 0.17 131 Wfs Mollisols Plutonic rocks

Colubrina arborescens Calcium (mg/g) II1A 19.74 4.22 8 19.62 25.35 13.57 104 Sdf Mollisosls Alluvial deposits Guanica Forest

Magnesium (mg/g) II1A 3.31 2.68 8 2.61 10.00 0.92 104 Sdf Mollisosls Alluvial deposits Guanica Forest

Nitrogen (%) II1A 1.39 0.44 8 1.40 2.08 0.21 104 Sdf Mollisosls Alluvial deposits Guanica Forest

Phosphorus (mg/g) II1A 0.58 0.14 8 0.53 0.83 0.40 104 Sdf Mollisosls Alluvial deposits Guanica Forest

Potassium (mg/g) II1A 19.45 3.80 8 18.44 26.06 14.12 104 Sdf Mollisosls Alluvial deposits Guanica Forest

Colubrina elliptica Calcium (mg/g) II1Ci 17.95 5.12 2 18.06 22.60 13.10 104 Sdf Mollisosls Alluvial deposits Guanica Forest

Magnesium (mg/g) II1Ci 1.98 0.56 2 2.00 2.52 1.41 104 Sdf Mollisosls Alluvial deposits
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

72

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1Ci 1.98 0.56 4 2.00 2.52 1.41 149 Sdf Mollisols Alluvial deposits Guanica Forest

Nitrogen (%) II1A 2.27 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 1.45 0.29 4 1.41 1.79 1.21 149 Sdf Mollisols Alluvial deposits Guanica Forest

Phosphorus (mg/g) II1A 0.84 0.03 2 0.84 0.86 0.82 104 Sdf Mollisosls Alluvial deposits Mature

II1Ci 0.33 0.05 2 0.33 0.38 0.28 104 Sdf Mollisosls Alluvial deposits Guanica Forest

Potassium (mg/g) II1A 23.78 0.28 2 23.78 23.98 23.59 104 Sdf Mollisosls Alluvial deposits Mature

II1A 23.78 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 14.16 7.30 2 14.16 20.60 7.73 104 Sdf Mollisosls Alluvial deposits Guanica Forest

Commelina diffusa Aluminum (mg/g) II3 1.10 1.13 2 1.10 1.90 0.30 135 Lmrf Ultisols Tuffac. sandstone

C/N II3 20 4 11 19 28 15 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II3 6.40 0.73 2 6.40 6.92 5.89 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II3 40 1 11 40 42 38 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II3 0.78 0.84 2 0.78 1.38 0.19 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II3 3.21 0.28 2 3.21 3.41 3.01 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II3 0.81 0.21 2 0.81 0.96 0.66 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II3 2.00 0.45 12 2.08 2.72 1.38 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II3 2.05 1.79 2 2.05 3.31 0.79 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II3 32.72 8.28 2 32.72 38.57 26.87 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II3 0.65 0.20 11 0.58 0.96 0.47 135 Lmrf Ultisols Tuffac. sandstone

Commelina spp. Aluminum (mg/g) II3 0.88 0.61 19 0.77 2.01 0.84 135 Lmrf Ultisols Tuffac. sandstone

II3 0.44 0.50 9 0.27 1.67 0.08 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II3 5.79 0.75 19 5.75 7.56 4.51 135 Lmrf Ultisols Tuffac. sandstone

II3 6.08 1.18 9 5.67 7.85 4.32 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II3 0.84 0.67 19 0.72 2.30 0.10 135 Lmrf Ultisols Tuffac. sandstone

II3 0.43 0.49 0.21 1.54 0.09 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II3 3.11 0.39 19 3.03 3.85 2.50 135 Lmrf Ultisols Tuffac. sandstone

II3 3.33 0.52 9 3.11 4.12 2.61 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

73

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Manganese (mg/g) II3 0.85 0.30 19 0.81 1.55 0.46 135 Lmrf Ultisols Tuffac. sandstone

II3 0.82 0.27 9 0.80 1.13 0.43 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II3 1.56 0.28 19 1.61 2.00 0.93 135 Lmrf Ultisols Tuffac. sandstone

II3 1.84 0.52 9 1.88 2.51 1.05 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II3 0.90 0.36 19 0.72 1.60 0.51 135 Lmrf Ultisols Tuffac. sandstone

II3 1.22 0.66 9 1.01 2.87 0.61 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II3 35.73 7.99 19 33.63 51.06 25.22 135 Lmrf Ultisols Tuffac. sandstone

II3 38.29 3.92 9 39.37 43.46 31.04 135 Lmrf Ultisols Tuffac. sandstone

Conocarpus erectus Aluminum (mg/kg) II1B 39 13 4 44 47 20 87 Mfs Entisols Compound dunes

II1B 21 4 3 20 17 25 157 Mfs Entisols Compound dunes

Ash (%) II1B 2.24 0.30 4 2.26 2.53 1.92 87 Mfs Entisols Compound dunes

C/N II1B 301 21 4 295 329 285 87 Mfs Entisols Compound dunes

Calcium (mg/g) II1B 6.10 1.12 4 5.73 7.75 5.21 87 Mfs Entisols Compound dunes

II1B 9.20 0.84 3 9.35 8.30 9.95 157 Mfs Entisols Compound dunes

Iron (mg/g) II1B 0.06 0.02 4 0.07 0.08 0.03 87 Mfs Entisols Compound dunes

II1B 0.03 0.02 3 0.03 0.01 0.05 157 Mfs Entisols Compound dunes

Magnesium (mg/g) II1B 0.96 0.14 4 0.99 1.10 0.77 87 Mfs Entisols Compound dunes

II1B 0.64 0.11 3 0.58 0.57 0.76 157 Mfs Entisols Compound dunes

Manganese (mg/kg) II1B 8 3 4 8 11 6 87 Mfs Entisols Compound dunes

II1B 5 2 3 6 2 6 157 Mfs Entisols Compound dunes

Nitrogen (%) II1B 0.15 0.01 4 0.15 0.16 0.14 87 Mfs Entisols Compound dunes

Phosphorus (mg/g) II1B 0.24 0.07 4 0.26 0.30 0.16 87 Mfs Entisols Compound dunes

II1B 0.18 0.04 3 0.19 0.14 0.22 157 Mfs Entisols Compound dunes

Potassium (mg/g) II1B 2.36 0.49 4 2.32 2.92 1.90 87 Mfs Entisols Compound dunes

II1B 1.60 1.42 3 0.79 0.77 3.23 157 Mfs Entisols Compound dunes

Cordia borinquensis Ash (%) II1B 5.82 1 68 Swf Ultisols Tuffac. sandstone Class 1 LD

II1B 4.58 3.12 3 3.28 8.14 2.33 68 Swf Ultisols Tuffac. sandstone Class 3 LD
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

74

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Calcium (mg/g) II1A 3.89 1 135 Lmrf Ultisols Tuffac. sandstone

II1B 14.85 1 68 Swf Ultisols Tuffac. sandstone Class 1 LD

II1B 6.28 0.59 3 5.94 6.97 5.94 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 4.72 0.51 3 4.49 5.30 4.37 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II1A 4.20 1 135 Lmrf Ultisols Tuffac. sandstone

II1B 1.35 0.34 3 1.55 1.55 0.96 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II1A 1.94 1 135 Lmrf Ultisols Tuffac. sandstone

II1B 0.74 1 68 Swf Ultisols Tuffac. sandstone Class 1 LD

II1B 0.72 0.16 3 0.74 0.86 0.55 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 0.80 0.16 3 0.86 0.91 0.62 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.88 1 135 Lmrf Ultisols Tuffac. sandstone

II1B 0.23 1 68 Swf Ultisols Tuffac. sandstone Class 1 LD

II1B 0.48 0.15 3 0.54 0.60 0.31 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 0.46 0.02 3 0.46 0.48 0.45 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II1A 13.54 1 135 Lmrf Ultisols Tuffac. sandstone

II1B 2.43 0.52 3 2.46 2.94 1.90 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 4.21 1 68 Swf Ultisols Tuffac. sandstone Class 1 LD

II1B 6.96 1.22 3 7.59 7.74 5.56 135 Lmrf Ultisols Tuffac. sandstone

Cordia sulcata Ash (%) II1B 3.31 1.24 8 3.21 5.13 2.11 68 Swf Ultisols Tuffac. sandstone Class 3 LD

Calcium (mg/g) II1B 2.74 1.37 8 2.17 5.42 1.67 68 Swf Ultisols Tuffac. sandstone Class 3 LD

Nitrogen (%) II1B 0.29 0.09 8 0.28 0.43 0.22 68 Swf Ultisols Tuffac. sandstone Class 3 LD

Phosphorus (mg/g) II1B 0.07 0.03 8 0.08 0.11 0.02 68 Swf Ultisols Tuffac. sandstone Class 3 LD

Potassium (mg/g) II1B 0.83 0.30 8 0.74 1.46 0.59 68 Swf Ultisols Tuffac. sandstone Class 3 LD

Wood density (g/cc) II1B 0.48 0.00 3 0.48 0.48 0.48 97 Wfs Inceptisols Tuffac. sandstone

Croton poecilanthus Aluminum (mg/g) II1A 0.54 1 93 Wflm Inceptisols Tuffac. sandstone New

II1B 2.36 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.01 1 93 Wflm Inceptisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

75

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8A 0.42 0.03 3 0.41 0.45 0.41 145 Wfs Ultisols Tuffac. sandstone

Ash (%) II1A 5.64 1 93 Wflm Inceptisols Tuffac. sandstone New

II1B 1.34 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.49 0.12 4 0.46 0.63 0.39 68 Swf Ultisols Tuffac. sandstone Class 2 LD

II1B 2.80 0.62 2 2.80 3.24 2.36 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1C 0.71 1 93 Wflm Inceptisols Tuffac. sandstone

II8A 8.35 0.35 3 8.17 8.76 8.13 145 Wfs Ultisols Tuffac. sandstone

II8A 8.58 1 71 Wfs Ultisols Tuffac. sandstone

C/N II1A 26 1 93 Wflm Inceptisols Tuffac. sandstone New

II1B 59 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 287 65 4 287 327 191 68 Swf Ultisols Tuffac. sandstone Class 2 LD

II1B 67 10 2 67 83 70 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1C 199 1 93 Wflm Inceptisols Tuffac. sandstone

II8A 68 1 3 68 69 68 145 Wfs Ultisols Tuffac. sandstone

Calcium (mg/g) II1A 6.72 1 93 Wflm Inceptisols Tuffac. sandstone New

II1B 12.50 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 1.45 0.39 4 1.33 2.02 1.12 68 Swf Ultisols Tuffac. sandstone Class 2 LD

II1B 5.62 1.87 2 5.62 6.94 4.30 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1C 0.84 1 93 Wflm Inceptisols Tuffac. sandstone

II8A 14.32 0.63 3 14.18 15.01 13.78 145 Wfs Ultisols Tuffac. sandstone

Carbon (%) II8A 52 1 3 52 53 52 145 Wfs Ultisols Tuffac. sandstone

II8A 54 1 71 Wfs Ultisols Tuffac. sandstone

Iron (mg/g) II1A 0.15 1 93 Wflm Inceptisols Tuffac. sandstone New

II1B 0.13 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.01 1 93 Wflm Inceptisols Tuffac. sandstone

II8A 0.10 0.01 3 0.10 0.10 0.09 145 Wfs Ultisols Tuffac. sandstone

Magnesium (mg/g) II8A 3.32 0.14 3 3.26 3.49 3.23 145 Wfs Ultisols Tuffac. sandstone

Manganese (mg/g) II1A 1.70 1 93 Wflm Inceptisols Tuffac. sandstone New

II1B 1.36 1 93 Wflm Inceptisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

76

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1C 0.13 1 93 Wflm Inceptisols Tuffac. sandstone

II8A 1.69 0.12 3 1.75 1.76 1.55 145 Wfs Ultisols Tuffac. sandstone

Nitrogen (%) II1A 1.02 1 93 Wflm Inceptisols Tuffac. sandstone New

II1B 0.77 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.17 0.05 4 0.15 0.24 0.14 68 Swf Ultisols Tuffac. sandstone

II1B 0.59 0.07 2 0.59 0.64 0.54 68 Swf Ultisols Tuffac. sandstone

II1C 0.13 1 93 Wflm Inceptisols Tuffac. sandstone

II8A 0.76 0.01 3 0.76 0.77 0.76 145 Wfs Ultisols Tuffac. sandstone

II8A 0.83 1 71 Wfs Ultisols Tuffac. sandstone

Phosphorus (mg/g) II1A 1.02 1 93 Wflm Inceptisols Tuffac. sandstone New

II1B 0.40 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.10 0.05 4 0.09 0.16 0.06 68 Swf Ultisols Tuffac. sandstone Class 2 LD

II1B 0.16 0.03 2 0.16 0.18 0.14 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1C 0.16 1 93 Wflm Inceptisols Tuffac. sandstone

II8A 0.25 0.02 3 0.25 0.26 0.23 145 Wfs Ultisols Tuffac. sandstone

Potassium (mg/g) II1A 12.31 1 93 Wflm Inceptisols Tuffac. sandstone New

II1B 5.49 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.31 0.14 4 0.29 0.50 0.18 68 Swf Ultisols Tuffac. sandstone Class 2 LD

II1B 0.61 0.04 2 0.61 0.64 0.58 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1C 0.67 1 93 Wflm Inceptisols Tuffac. sandstone

II8A 4.40 0.13 3 4.42 4.52 4.26 145 Wfs Ultisols Tuffac. sandstone

Sulfur (%) II8A 0.11 0.01 3 0.11 0.12 0.11 145 Wfs Ultisols Tuffac. sandstone

II8A 0.15 1 71 Wfs Ultisols Tuffac. sandstone

Ctenitis hirta C/N II6 17 1 135 Lmrf Ultisols Tuffac. sandstone Ferms

Carbon (%) II6 43 1 135 Lmrf Ultisols Tuffac. sandstone Ferms

Nitrogen (%) II6 2.56 1 135 Lmrf Ultisols Tuffac. sandstone Ferms

Sulfur (%) II6 0.42 1 135 Lmrf Ultisols Tuffac. sandstone Ferms
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

77

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Cupania americana Wood density (g/cc) II1B 0.52 1 97 Wfs Inceptisols Tuffac. sandstone

Cyathea arborea Aluminum (mg/g) II6 0.64 0.30 6 0.73 0.88 0.06 135 Lmrf Ultisols Tuffac. sandstone

C/N II6 35 6 3 38 39 28 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II6 1.94 0.39 6 2.04 2.38 1.26 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II6 47 1 3 46 49 46 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II6 0.37 0.26 6 0.33 0.75 0.13 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II6 2.39 2.60 6 1.39 7.63 0.88 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II6 0.08 0.04 6 0.08 0.13 0.04 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II6 1.08 0.31 8 1.07 1.64 0.76 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II6 0.48 0.12 6 0.46 0.68 0.34 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II6 10.68 6.55 6 9.42 21.78 1.74 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II6 0.21 0.05 3 0.22 0.25 0.15 135 Lmrf Ultisols Tuffac. sandstone

Cyclopogon cranichoides Aluminum (mg/g) II6 0.91 1 135 Lmrf Ultisols Tuffac. sandstone

C/N II6 13 1 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II6 7.61 1 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II6 43 1 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II6 0.50 1 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II6 4.76 1 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II6 0.34 1 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II6 3.28 1 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II6 3.10 1 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II6 28.80 1 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II6 0.45 1 135 Lmrf Ultisols Tuffac. sandstone

Cyrilla racemiflora Aluminum (mg/g) II1A 0.10 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 9.25 1 93 Wflm Inceptisols Tuffac. sandstone Fresh-1980

II1B 0.01 1 93 Wflm Inceptisols Tuffac. sandstone Fresh-1990
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

78

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1C 0.03 0.01 3 0.03 0.04 0.03 93 Wflm Inceptisols Tuffac. sandstone

Ash (%) II1A 2.79 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 8.12 1 93 Wflm Inceptisols Tuffac. sandstone Fresh-1980

II1B 0.73 1 93 Wflm Inceptisols Tuffac. sandstone Fresh-1990

II1B 2.30 0.12 65 2.09 6.06 1.06 67 Smf Ultisols Tuffac. sandstone Bark sound

II1B 3.37 0.97 22 1.67 20.36 0.31 67 Smf Ultisols Tuffac. sandstone Exposed surface

II1B 0.37 0.05 17 0.36 0.76 0.03 67 Smf Ultisols Tuffac. sandstone Sapwood sound

II1B 0.52 0.06 19 0.59 1.03 0.01 67 Smf Ultisols Tuffac. sandstone Heartwood sound

II1B 0.46 0.04 64 0.43 1.21 0.01 67 Smf Ultisols Tuffac. sandstone Xylem sound

II1B 0.44 0.07 32 0.31 1.61 0.02 67 Smf Ultisols Tuffac. sandstone Xylem-cerambycid

II1B 0.64 0.12 28 0.38 2.50 0.05 67 Smf Ultisols Tuffac. sandstone Xylem-(hard)

II1B 1.38 0.23 9 1.41 2.26 0.41 67 Smf Ultisols Tuffac. sandstone Xylem-(friable)

II1B 3.69 1.00 35 1.81 3.22 0.06 67 Smf Ultisols Tuffac. sandstone Humus,soil-like

II1C 0.66 0.73 3 0.24 1.50 0.24 93 Wflm Inceptisols Tuffac. sandstone Fresh

C/N II1B 52 1 93 Wflm Inceptisols Tuffac. sandstone Fresh-1980

II1B 765 1 93 Wflm Inceptisols Tuffac. sandstone Fresh-1990

II1C 639 225 3 769 769 378 93 Wflm Inceptisols Tuffac. sandstone Fresh

Calcium (mg/g) II1A 1.65 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 3.26 0.15 65 3.12 7.21 0.57 67 Smf Ultisols Tuffac. sandstone Bark sound

II1B 2.00 0.14 22 1.98 3.28 0.99 67 Smf Ultisols Tuffac. sandstone Exposed surface

II1B 0.59 0.09 17 0.51 1.82 0.25 67 Smf Ultisols Tuffac. sandstone Sapwood sound

II1B 0.59 0.08 19 0.50 1.62 0.14 67 Smf Ultisols Tuffac. sandstone Heartwood sound

II1B 0.67 0.05 64 0.54 2.82 0.16 67 Smf Ultisols Tuffac. sandstone Xylem sound

II1B 0.76 0.06 32 0.65 1.76 0.22 67 Smf Ultisols Tuffac. sandstone Xylem with cerambycid

II1B 1.04 0.15 28 0.88 3.95 0.01 67 Smf Ultisols Tuffac. sandstone Xylem-(hard)

II1B 1.50 0.30 9 1.01 2.62 0.44 67 Smf Ultisols Tuffac. sandstone Xylem-(friable)

II1B 1.24 0.23 35 0.73 6.23 0.11 67 Smf Ultisols Tuffac. sandstone Humus,soil-like

II1B 19.10 1 93 Wflm Inceptisols Tuffac. sandstone Fresh-1980

II1B 0.91 1 93 Wflm Inceptisols Tuffac. sandstone Fresh-1990
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

79

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1C 0.23 0.03 3 0.22 0.27 0.21 93 Wflm Inceptisols Tuffac. sandstone

Iron (mg/g) II1A 0.10 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.11 1 93 Wflm Inceptisols Tuffac. sandstone Fresh-1980

II1B 0.11 1 93 Wflm Inceptisols Tuffac. sandstone Fresh-1990

II1C 0.06 0.04 3 0.04 0.10 0.04 93 Wflm Inceptisols Tuffac. sandstone Fresh

Magnesium (mg/g) II1A 0.69 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 1.85 1 93 Wflm Inceptisols Tuffac. sandstone Fresh-1980

II1B 0.17 1 93 Wflm Inceptisols Tuffac. sandstone Fresh-1990

II1B 1.83 0.11 65 1.58 4.20 0.55 67 Smf Ultisols Tuffac. sandstone Bark sound

II1B 1.21 0.12 22 1.08 3.01 0.67 67 Smf Ultisols Tuffac. sandstone Exposed surface

II1B 0 41 0 04 17 0.37 0.90 0.26 67 Smf Ultisols Tuffac. sandstone Sapwood sound

II1B 0.43 0.07 19 0.32 1.31 0.09 67 Smf Ultisols Tuffac. sandstone Heartwood sound

II1B 0.35 0.04 64 0.26 1.46 0.06 67 Smf Ultisols Tuffac. sandstone Xylem sound

II1B 0.39 0.03 32 0.37 0.90 0.09 67 Smf Ultisols Tuffac. sandstone Xylem-cerambycid

II1B 0 69 0.10 28 0.53 2.73 0.13 67 Smf Ultisols Tuffac. sandstone Xylem-(hard)

II1B 0.69 0.11 9 0.60 1.26 0.28 67 Smf Ultisols Tuffac. sandstone Xylem-(friable)

II1B 0.68 0.06 35 0.61 1.68 0.16 67 Smf Ultisols Tuffac. sandstone Humus,soil-like

II1C 0.14 0.08 3 0.10 0.23 0.08 93 Wflm Inceptisols Tuffac. sandstone Fresh

Nitrogen (%) II1A 0.69 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.82 1 93 Wflm Inceptisols Tuffac. sandstone Fresh-1980

II1B 0.06 1 93 Wflm Inceptisols Tuffac. sandstone Fresh-1990

II1B 0.60 0.02 65 0.58 1.11 0.35 67 Smf Ultisols Tuffac. sandstone Bark sound

II1B 0.46 0.05 22 0.44 1.07 0.12 67 Smf Ultisols Tuffac. sandstone Exposed surface

II1B 0.09 0.01 17 0.08 0.18 0.05 67 Smf Ultisols Tuffac. sandstone Sapwood sound

II1B 0.06 0.01 19 0.06 0.09 0.02 67 Smf Ultisols Tuffac. sandstone Heartwood sound

II1B 0.07 0.00 64 0.07 0.18 0.03 67 Smf Ultisols Tuffac. sandstone Xylem sound

II1B 0.13 0.01 32 0.12 0.26 0.08 67 Smf Ultisols Tuffac. sandstone Xylem-cerambycid

II1B 0.13 0.03 28 0.09 0.83 0.02 67 Smf Ultisols Tuffac. sandstone Xylem-(hard)

II1B 0.17 0.03 9 0.16 0.36 0.08 67 Smf Ultisols Tuffac. sandstone Xylem-(friable)
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

80

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1B 0.69 0.06 35 0.64 1.69 0.27 67 Smf Ultisols Tuffac. sandstone Humus,soil-like

II1C 0.08 0.04 3 0.06 0.12 0.06 93 Wflm Inceptisols Tuffac. sandstone Fresh

Phosphorus (mg/kg) II1A 662 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 582 1 93 Wflm Inceptisols Tuffac. sandstone Fresh-1980

II1B 12 1 93 Wflm Inceptisols Tuffac. sandstone Fresh-1990

II1B 140 10 65 126 360 140 67 Smf Ultisols Tuffac. sandstone Bark sound

II1B 90 14 22 82 332 7 67 Smf Ultisols Tuffac. sandstone Exposed surface

II1B 16 2 17 19 31 4 67 Smf Ultisols Tuffac. sandstone Sapwood sound

II1B 6 1 19 4 17 3 67 Smf Ultisols Tuffac. sandstone Heartwood sound

II1B 7 1 64 5 42 2 67 Smf Ultisols Tuffac. sandstone Xylem sound

II1B 18 2 32 15 53 5 67 Smf Ultisols Tuffac. sandstone Xylem-cerambycid

II1B 22 3 28 20 89 3 67 Smf Ultisols Tuffac. sandstone Xylem-(hard)

II1B 28 5 9 27 65 14 67 Smf Ultisols Tuffac. sandstone Xylem-(friable)

II1B 143 14 35 123 490 34 67 Smf Ultisols Tuffac. sandstone Humus,soil-like

II1C 20 20 3 10 40 10 93 Wflm Inceptisols Tuffac. sandstone

Potassium (mg/g) II1A 9.34 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 7.45 1 93 Wflm Inceptisols Tuffac. sandstone Fresh-1980

II1B 1.82 1 93 Wflm Inceptisols Tuffac. sandstone Fresh-1990

II1B 2.42 0.19 65 2.32 7.37 0.31 67 Smf Ultisols Tuffac. sandstone Bark sound

II1B 0.69 0.07 22 0.62 1.41 0.10 67 Smf Ultisols Tuffac. sandstone Exposed surface

II1B 0.98 0.12 17 0.93 1.87 0.03 67 Smf Ultisols Tuffac. sandstone Sapwood sound

II1B 1.28 0.14 19 1.20 2.49 0.10 67 Smf Ultisols Tuffac. sandstone Heartwood sound

II1B 1.04 0.11 64 0.84 3.58 0.02 67 Smf Ultisols Tuffac. sandstone Xylem sound

II1B 0.82 0.20 32 0.23 4.69 0.05 67 Smf Ultisols Tuffac. sandstone Xylem-cerambycid

II1B 1.00 0.12 28 0.97 3.17 0.09 67 Smf Ultisols Tuffac. sandstone Xylem-(hard)

II1B 1.36 0.56 9 0.42 4.19 0.06 67 Smf Ultisols Tuffac. sandstone Xylem-(friable)

II1B 0.48 0.07 35 2.77 0.20 67 Smf Ultisols Tuffac. sandstone Humus,soil-like

II1C 2.02 0.06 3 0.36 5.38 0.33 93 Wflm Inceptisols Tuffac. sandstone Fresh
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

81

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Dacryodes sp. Aluminum (mg/g) II8A 2.69 5.44 47 0.96 23.48 0.25 146 Swf Ultisols Tuffac. sandstone New

II8A 0.99 1.47 22 0.39 6.23 0.26 146 Swf Ultisols Tuffac. sandstone Old

Ash (%) II8A 13.71 4.24 46 12.59 32.65 8.64 146 Swf Ultisols Tuffac. sandstone New

II8A 11.68 3.19 21 10.39 18.69 8.70 146 Swf Ultisols Tuffac. sandstone Old

Calcium (mg/g) II8A 7.90 1.40 47 7.52 11.21 5.74 146 Swf Ultisols Tuffac. sandstone New

II8A 6.50 1.05 22 6.20 9.08 4.97 146 Swf Ultisols Tuffac. sandstone Old

Iron (mg/g) II8A 2.52 6.54 47 0.56 29.87 0.14 146 Swf Ultisols Tuffac. sandstone New

II8A 0.70 1.10 22 0.25 4.68 0.17 146 Swf Ultisols Tuffac. sandstone Old

Magnesium (mg/g) II8A 1.27 0.23 47 1.26 1.90 0.81 146 Swf Ultisols Tuffac. sandstone New

II8A 1.37 0.18 22 1.40 1.62 0.94 146 Swf Ultisols Tuffac. sandstone Old

Manganese (mg/g) II8A 0.83 0.16 47 0.78 1.27 0.60 146 Swf Ultisols Tuffac. sandstone New

II8A 0.88 0.13 22 0.91 1.09 0.55 146 Swf Ultisols Tuffac. sandstone Old

Phosphorus (mg/g) II8A 0.23 0.10 47 0.18 0.52 0.12 146 Swf Ultisols Tuffac. sandstone New

II8A 0.16 0.06 22 0.14 0.34 0.09 146 Swf Ultisols Tuffac. sandstone Old

Potassium (mg/g) II8A 1.34 0.23 47 1.32 1.86 0.83 146 Swf Ultisols Tuffac. sandstone New

II8A 1.18 0.22 22 1.17 1.56 0.74 146 Swf Ultisols Tuffac. sandstone Old

Dacryodes excelsa Aluminum (mg/g) II1A 0.69 0.91 15 0.47 3.61 0.06 107 Swf Ultisols Tuffac. sandstone

II1C 0.19 0.43 40 0.08 2.57 0.00 123 Wflm Inceptisols Tuffac. sandstone Wood decay

II1F 0.06 1 120 Swf Ultisols Tuffac. sandstone

II9A 0.36 0.04 6 0.37 0.41 0.32 153 Swf Ultisols Tuffac. sandstone

II8A 1.74 1.30 12 1.49 4.02 0.37 124 Lmrf Ultisols Tuffac. sandstone

II8A 1.70 1.03 11 1.54 4.31 0.47 124 Lmrf Ultisols Tuffac. sandstone

Ash (%) II1A 8.76 1 94 Wfs Ultisols Lava Understory biomass

II1A 6.50 1 101 Wfs Ultisols Tuffac. sandstone Overstory biomass

II1A 6.26 0.28 8 6.27 6.61 5.76 100 Lmrf Ultisols Tuffac. sandstone Decomp. (1month)

II1A 6.07 2.35 13 5.45 13.28 3.78 107 Swf Ultisols Tuffac. sandstone

II1B 0.88 0.48 45 0.78 2.75 0.40 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 2.22 1.35 13 1.55 5.37 0.63 68 Swf Ultisols Tuffac. sandstone Class 4 LD
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

82

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1B 2.94 1.71 2 2.94 4.15 1.73 68 Swf Ultisols Tuffac. sandstone Class 5 LD

II1B 2.68 1.39 5 2.00 4.63 1.42 68 Swf Ultisols Tuffac. sandstone Class 3/4 LD

II1B 6.96 6.37 3 5.03 14.08 1.78 68 Swf Ultisols Tuffac. sandstone Class 4/5 LD

II1B 1.63 0.99 8 1.51 3.79 0.66 68 Swf Ultisols Tuffac. sandstone Class 3 SD

II1B 6.66 1 68 Swf Ultisols Tuffac. sandstone Class 3/4 ST

II1C 4.66 1 94 Wfs Ultisols Lava Understory biomass

II1F 3.48 1 120 Swf Ultisols Tuffac. sandstone

II8A 8.04 1 71 Wfs Ultisols Tuffac. sandstone

II9A 10.07 0.81 6 10.26 10.84 8.85 153 Swf Ultisols Tuffac. sandstone

II8A 13.40 1.74 12 13.55 15.75 9.74 124 Lmrf Ultisols Tuffac. sandstone

II8A 12.62 1.16 11 12.30 14.38 11.22 124 Wflm Ultisols Tuffac. sandstone

C/N II1A 29 5 14 2 39 21 107 Swf Ultisols Tuffac. sandstone

II9A 66 4 6 67 69 59 153 Swf Ultisols Tuffac. sandstone

II8A 47 4 12 47 55 42 124 Lmrf Ultisols Tuffac. sandstone

II8A 51 5 11 51 59 44 124 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II1A 3.55 0.53 5 3.60 4.26 2.81 100 Lmrf Ultisols Tuffac. sandstone

II1A 5.46 2.07 5 6.01 7.25 2.23 100 Lmrf Ultisols Tuffac. sandstone Decomp. (1week)

II1A 4.32 0.85 5 4.34 5.50 3.09 100 Lmrf Ultisols Tuffac. sandstone Decomp. (2 week)

II1A 6.26 0.72 8 6.75 7.12 4.90 100 Lmrf Ultisols Tuffac. sandstone Decomp. (1 month)

II1A 4.66 0.81 5 5.13 5.35 3.55 135 Lmrf Ultisols Tuffac. sandstone

II1A 2.68 0.53 15 2.58 3.70 1.90 107 Swf Ultisols Tuffac. sandstone

II1B 1.38 0.65 45 1.39 3.32 0.30 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 1.94 1.50 13 1.40 4.68 0.50 68 Swf Ultisols Tuffac. sandstone Class 4 LD

II1B 1.00 0.14 2 1.00 1.10 0.90 68 Swf Ultisols Tuffac. sandstone Class 5 LD

II1B 4.61 3.23 5 3.16 9.44 1.84 68 Swf Ultisols Tuffac. sandstone Class 3/4 LD

II1B 0.47 0.32 3 0.60 0.70 0.10 68 Swf Ultisols Tuffac. sandstone Class 4/5 LD

II1B 1.57 0.62 8 1.75 2.33 0.62 68 Swf Ultisols Tuffac. sandstone Class 3 SD

II1B 10.59 1 68 Swf Ultisols Tuffac. sandstone Class 3/4 ST

II1B 5.18 2.24 6 5.50 8.37 2.28 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

83

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1B 1.32 2.66 8 0.41 7.89 0.28 135 Lmrf Ultisols Tuffac. sandstone

II1C 6.42 3.16 40 6.45 15.94 2.22 123 Wflm Inceptisols Tuffac. sandstone Wood decay

II1Ci 3.45 0.64 6 3.56 4.00 2.24 135 Lmrf Ultisols Tuffac. sandstone

II1F 2.59 1 120 Swf Ultisols Tuffac. sandstone

II9A 5.77 0.62 6 5.79 6.43 5.04 153 Swf Ultisols Tuffac. sandstone

II8A 7.07 1.47 12 6.90 10.68 4.65 124 Lmrf Ultisols Tuffac. sandstone

II8A 10.60 0.61 11 10.61 11.92 9.80 124 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II1A 47 1 92 48 49 45 150 Swf Ultisols Tuffac. sandstone

II1A 47 2 14 47 49 43 107 Swf Ultisols Tuffac. sandstone

II8A 51 1 71 Wfs Ultisols Tuffac. sandstone

II9A 50 1 6 49 51 49 153 Swf Ultisols Tuffac. sandstone

II8A 40.05 0.80 12 39.98 41.75 38.97 124 Lmrf Ultisols Tuffac. sandstone

II8A 40.41 0.54 11 40.56 41.06 39.60 124 Wflm Ultisols Tuffac. sandstone

Iron (mg/kg) II1A 450 610 15 270 2490 90 107 Rfs Inceptisols Tuffac. sandstone

II1C 170 400 40 70 2470 3 123 Wflm Inceptisols Tuffac. sandstone Wood decay

II1F 21 1 120 Swf Ultisols Tuffac. sandstone

II9A 340 30 6 339 378 299 153 Swf Ultisols Tuffac. sandstone

II8A 1730 1508 12 1338 4658 250 124 Lmrf Ultisols Tuffac. sandstone

II8A 1123 701 11 979 2901 319 124 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II1A 0.86 0.13 5 0.81 1.05 0.74 100 Lmrf Ultisols Tuffac. sandstone

II1A 1.15 0.48 5 0.97 1.91 0.73 100 Lmrf Ultisols Tuffac. sandstone Decomp. (1 week)

II1A 0.84 0.09 5 0.85 0.95 0.73 100 Lmrf Ultisols Tuffac. sandstone Decomp (2 week)

II1A 0.87 0.06 5 0.89 0.94 0.81 100 Lmrf Ultisols Tuffac. sandstone

II1A 1.22 0.17 5 1.16 1.41 1.00 135 Lmrf Ultisols Tuffac. sandstone

II1A 1.29 0.31 15 1.30 2.14 0.80 107 Swf Ultisols Tuffac. sandstone

II1B 0.69 0.23 6 0.73 1.04 0.41 135 Lmrf Ultisols Tuffac. sandstone

II1B 0.26 0.40 8 0.12 1.25 0.11 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.82 0.40 40 0.79 1.76 0.10 123 Wflm Inceptisols Tuffac. sandstone Wood decay

II1Ci 0.70 0.04 6 0.70 0.76 0.64 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

84

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1F 1.76 1 120 Swf Ultisols Tuffac. sandstone

II9A 1.44 0.07 6 1.44 1.53 1.34 153 Swf Ultisols Tuffac. sandstone

II8A 1.51 0.07 12 1.50 1.65 1.41 124 Lmrf Ultisols Tuffac. sandstone

II8A 1.10 0.06 11 1.08 1.22 1.00 124 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II1A 0.35 0.12 15 0.37 0.58 0.11 107 Swf Ultisols Tuffac. sandstone

II1C 0.23 0.15 40 0.23 0.61 0.04 123 Wflm Inceptisols Tuffac. sandstone Wood decay

II1F 0.29 1 120 Swf Ultisols Tuffac. sandstone

II9A 1.22 0.17 6 1.24 1.38 0.97 153 Swf Ultisols Tuffac. sandstone

II8A 0.47 0.08 12 0.48 0.60 0.28 124 Lmrf Ultisols Tuffac. sandstone

II8A 0.95 0.14 11 0.97 1.17 0.78 124 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II1A 1.01 0.18 5 1.00 1.20 0.77 100 Lmrf Ultisols Tuffac. sandstone Decomp (1 week)

II1A 1.10 0.14 5 1.13 1.30 0.92 100 Lmrf Ultisols Tuffac. sandstone Decomp (2 week)

II1A 2.04 1 94 Wfs Ultisols Lava Understory biomass

II1A 1.32 1 101 Wfs Ultisols Tuffac. sandstone Overstory biomass

II1A 0.12 0.06 8 0.13 0.16 0.05 100 Lmrf Ultisols Tuffac. sandstone Decomp (1 month)

II1A 1.84 0.46 92 1.68 3.10 1.00 150 Swf Ultisols Tuffac. sandstone

II1A 1.35 0.11 5 1.38 1.47 1.23 135 Lmrf Ultisols Tuffac. sandstone

II1A 1.67 0.29 14 1.64 2.19 1.20 107 Swf Ultisols Tuffac. sandstone

II1B 0.20 0.12 45 0.16 0.68 0.08 68 Swf Ultisols Tuffac. sandstone Class 3LD

II1B 0.50 0.24 13 0.42 0.94 0.23 68 Swf Ultisols Tuffac. sandstone Class 4 LD

II1B 0.78 0.18 2 0.78 0.91 0.65 68 Swf Ultisols Tuffac. sandstone Class 5 LD

II1B 0.51 0.27 5 0.41 0.96 0.29 68 Swf Ultisols Tuffac. sandstone Class 3/4 LD

II1B 0.92 0.39 3 0.99 1.27 0.50 68 Swf Ultisols Tuffac. sandstone Class 4/5 LD

II1B 0.29 0.17 8 0.25 0.64 0.11 68 Swf Ultisols Tuffac. sandstone Class 3 SD

II1B 0.58 1 68 Swf Ultisols Tuffac. sandstone Class 3/4 ST

II1B 0.41 0.13 6 0.42 0.61 0.23 135 Lmrf Ultisols Tuffac. sandstone

II1B 0.15 0.19 8 0.08 0.61 0.07 135 Lmrf Ultisols Tuffac. sandstone

II1C 1.51 1 94 Wfs Ultisols Lava Understory biomass

II1C 0.67 0.28 40 0.59 1.57 0.32 123 Wflm Inceptisols Tuffac. sandstone Wood decay
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

85

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1Ci 0.31 0.02 6 0.32 0.34 0.28 135 Lmrf Ultisols Tuffac. sandstone

II1F 1.02 1 120 Swf Ultisols Tuffac. sandstone

II8A 1.06 1 71 Wfs Ultisols Tuffac. sandstone

II9A 0.75 0.06 6 0.73 0.86 0.70 153 Swf Ultisols Tuffac. sandstone

II8A 0.85 0.06 12 0.84 0.97 0.76 124 Lmrf Ultisols Tuffac. sandstone

II8A 0.80 0.06 11 0.80 0.91 0.70 124 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.21 0.05 5 0.23 0.24 0.13 100 Lmrf Ultisols Tuffac. sandstone

II1A 0.22 0.12 5 0.18 0.40 0.10 100 Lmrf Ultisols Tuffac. sandstone Decomp. (1 week)

II1A 0.27 0.09 5 0.28 0.38 0.14 100 Lmrf Ultisols Tuffac. sandstone Decomp. (2 week)

II1A 0.83 1 94 Wfs Ultisols Lava Understory biomass

II1A 0.70 1 101 Wfs Ultisols Tuffac. sandstone Overstory biomass

II1A 0.64 0.04 5 0.63 0.70 0.60 135 Lmrf Ultisols Tuffac. sandstone

II1A 0.77 0.15 15 0.75 1.10 0.56 107 Swf Ultisols Tuffac. sandstone

II1B 0.05 0.06 45 0.02 0.32 0.01 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 0.13 0.08 13 0.09 0.30 0.03 68 Swf Ultisols Tuffac. sandstone Class 4 LD

II1B 0.22 0.05 2 0.22 0.25 0.19 68 Swf Ultisols Tuffac. sandstone Class 5 LD

II1B 0.14 0.10 5 0.11 0.26 0.05 68 Swf Ultisols Tuffac. sandstone Class 3/4 LD

II1B 0.35 0.17 3 0.43 0.45 0.15 68 Swf Ultisols Tuffac. sandstone Class 4/5 LD

II1B 0.09 0.09 8 0.06 0.30 0.02 68 Swf Ultisols Tuffac. sandstone Class 3 Sd

II1B 0.09 1 68 Swf Ultisols Tuffac. sandstone Class 3/4 ST

II1B 0.27 0.08 6 0.29 0.39 0.14 135 Lmrf Ultisols Tuffac. sandstone

II1B 0.09 0.12 8 0.06 0.39 0.03 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.38 1 94 Wfs Ultisols Lava Understory biomass

II1C 0.22 0.10 40 0.20 0.52 0.07 123 Swf Ultisols Tuffac. sandstone Wood decay

II1Ci 0.16 0.01 6 0.16 0.18 0.15 135 Lmrf Ultisols Tuffac. sandstone

II1F 0.57 1 120 Swf Ultisols Tuffac. sandstone

II9A 0.13 0.01 6 0.13 0.14 0.12 153 Swf Ultisols Tuffac. sandstone

II8A 0.20 0.05 12 0.19 0.34 0.13 124 Lmrf Ultisols Tuffac. sandstone

II8A 0.17 0.03 11 0.16 0.22 0.12 124 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

86

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Potassium (mg/g) II1A 1.61 0.13 5 1.56 1.81 1.50 100 Lmrf Ultisols Tuffac. sandstone

II1A 1.54 0.16 5 1.54 1.68 1.29 100 Lmrf Ultisols Tuffac. sandstone Decomp. (l week)

II1A 1.35 0.20 5 1.34 1.67 1.10 100 Lmrf Ultisols Tuffac. sandstone Decomp. (2 week)

II1A 4.54 1 94 Wfs Ultisols Lava Understory biomass

II1A 7.22 1 101 Wfs Ultisols Tuffac. sandstone Overstory biomass

II1A 6.26 0.72 8 6.45 7.12 4.90 100 Lmrf Ultisols Tuffac. sandstone Decomp. (1 month)

II1A 5.80 1.40 5 5.73 7.24 3.59 135 Lmrf Ultisols Tuffac. sandstone

II1A 7.84 2.96 15 7.10 12.89 4.99 107 Swf Ultisols Tuffac. sandstone

II1B 0.62 0.42 45 0.59 1.66 0.03 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 0.47 0.42 13 0.23 1.24 0.08 68 Swf Ultisols Tuffac. sandstone Class 4 LD

II1B 0.20 0.11 4 0.17 0.35 0.10 68 Swf Ultisols Tuffac. sandstone Class 5 LD

II1B 0.92 0.52 5 0.74 1.71 0.41 68 Swf Ultisols Tuffac. sandstone Class 3/4 LD

II1B 0.27 0.12 3 0.21 0.41 0.20 68 Swf Ultisols Tuffac. sandstone Class 4/5 LD

II1B 1.23 0.13 8 1.22 1.41 1.04 68 Swf Ultisols Tuffac. sandstone Class 3 SD

II1B 0.71 1 68 Swf Ultisols Tuffac. sandstone Class 3/4 ST

II1B 3.91 1.78 6 3.63 7.28 1.94 135 Lmrf Ultisols Tuffac. sandstone

II1B 1.74 2.10 8 1.02 6.89 0.73 135 Lmrf Ultisols Tuffac. sandstone

II1C 5.51 1 94 Wfs Ultisols Lava Understory biomass

II1C 1.19 0.91 40 1.05 6.09 0.41 123 Wflm Inceptisols Tuffac. sandstone

II1Ci 4.23 0.29 6 4.13 4.81 4.02 135 Lmrf Ultisols Tuffac. sandstone

II1F 2.68 1 120 Swf Ultisols Tuffac. sandstone

II9A 2.12 0.09 6 2.14 2.21 1.96 153 Swf Ultisols Tuffac. sandstone

II8A 1.55 0.82 12 1.39 4.08 0.87 124 Lmrf Ultisols Tuffac. sandstone

II8A 1.71 0.22 11 1.66 2.25 1.39 124 Lmrf Ultisols Tuffac. sandstone

Sodium (mg/g) II9A 3.97 0.22 6 4.00 4.23 3.68 153 Swf Ultisols Tuffac. sandstone

Sulfur (%) II1A 0.32 0.10 59 0.29 0.69 0.19 150 Swf Ultisols Tuffac. sandstone

II8A 0.38 1 71 Wfs Ultisols Tuffac. sandstone

Danaea nodosa Aluminum (mg/g) II6A 7.71 1 129 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

87

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II6 4.37 2.01 69 3.98 10.63 0.03 135 Lmrf Ultisols Tuffac. sandstone

Ash (%) II6A 7.70 1 129 Lmrf Ultisols Tuffac. sandstone

C/N II6A 7 1 129 Lmrf Ultisols Tuffac. sandstone

II6 26 30 26 39 18 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II6A 10.49 1 129 Lmrf Ultisols Tuffac. sandstone

II6 3.77 1.41 70 3.69 7.55 0.89 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II6A 49 1 129 Lmrf Ultisols Tuffac. sandstone

II6 43 1 30 43 45 39 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II6A 1.43 1 129 Lmrf Ultisols Tuffac. sandstone

II6 0.37 0.68 7 0.20 4.37 0.02 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II6A 3.29 1 129 Lmrf Ultisols Tuffac. sandstone

II6 3.72 1.41 70 3.57 10.22 1.68 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II6A 0.13 1 129 Lmrf Ultisols Tuffac. sandstone

II6 0.15 0.08 70 0.14 0.54 0.01 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II6A 22.95 1 129 Lmrf Ultisols Tuffac. sandstone

II6 1.64 0.32 92 1.60 2.42 0.96 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II6A 1.19 1 129 Lmrf Ultisols Tuffac. sandstone

II6 0.93 0.24 70 0.93 1.54 0.41 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II6A 7.89 1 129 Lmrf Ultisols Tuffac. sandstone

II6 19.34 8.15 70 18.75 37.61 3.08 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II6A 0.68 1 129 Lmrf Ultisols Tuffac. sandstone

II6 0.41 0.17 30 0.35 0.83 0.17 135 Lmrf Ultisols Tuffac. sandstone

Daphnopsis philippiana Aluminum (mg/g) II1A 0.17 1 93 Wflm Inceptisols Tuffac. sandstone

Ash (%) II1A 3.09 1 93 Wflm Inceptisols Tuffac. sandstone

C/N II1A 36 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 67 1 93 Wflm Inceptisols Tuffac. sandstone

Calcium (mg/g) II1A 3.86 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 4.97 1 93 Wflm Inceptisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

88

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Iron (mg/g) II1A 0.09 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.09 1 93 Wflm Inceptisols Tuffac. sandstone

Magnesium (mg/g) II1A 1.51 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.47 1 93 Wflm Inceptisols Tuffac. sandstone

Manganese (mg/g) II1A 0.14 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.22 1 93 Wflm Inceptisols Tuffac. sandstone

Nitrogen (%) II1A 1.24 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.68 1 93 Wflm Inceptisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.57 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.42 1 93 Wflm Inceptisols Tuffac. sandstone

Potassium (mg/g) II1A 6.93 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 4.84 1 93 Wflm Inceptisols Tuffac. sandstone

Delonix regia Wood density (g/cc) II1C 0.29 0.02 3 0.30 0.31 0.27 115

Dendropanax arboreus Ash (%) II8A 9.05 1 71 Wfs Ultisols Tuffac. sandstone

Carbon (%) II8A 53 1 71 Wfs Ultisols Tuffac. sandstone

Nitrogen (%) II8A 1.09 1 71 Wfs Ultisols Tuffac. sandstone

Sulfur (%) II8A 0.25 1 71 Wfs Ultisols Tuffac. sandstone

Dennstaedtia bipinnata Aluminum (mg/g) II6 0.29 0.15 7 0.29 0.54 0.03 135 Lmrf Ultisols Tuffac. sandstone

C/N II6 20 4 4 19 26 18 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II6 1.15 0.35 7 1.09 1.55 0.69 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II6 43 1 4 44 45 41 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II6 0.24 0.06 7 0.21 0.37 0.19 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II6 3.33 2.91 7 2.17 9.80 1.67 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II6 0.06 0.03 7 0.05 0.11 0.03 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II6 2.23 0.36 7 2.39 2.54 1.58 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II6 0.85 0.23 7 0.88 1.12 0.52 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

89

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Potassium (mg/g) II6 17.59 8.18 7 18.17 27.72 2.94 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II6 0.79 0.11 4 0.80 0.89 0.66 135 Lmrf Ultisols Tuffac. sandstone

Dennstaedtia obstusifolia Aluminum (mg/g) II6 0.28 0.18 10 0.32 0.56 0.06 135 Lmrf Ultisols Tuffac. sandstone

C/N II6 24 5 10 22 34 18 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II6 4.71 2.20 10 5.45 8.75 0.69 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II6 44 2 10 45 47 42 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II6 0.18 0.11 10 0.18 0.33 0.05 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II6 3.06 1.05 10 2.77 4.95 1.82 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II6 0.07 0.03 10 0.07 0.14 0.02 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II6 1.87 0.34 16 1.82 2.42 1.33 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II6 0.90 0.28 10 0.79 1.48 0.54 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II6 23.84 7.47 10 24.16 37.05 13.43 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II6 0.63 0.51 10 0.40 1.68 0.24 135 Lmrf Ultisols Tuffac. sandstone

Dennstaetdia spp. Aluminum (mg/g) II6 0.47 0.09 3 0.15 0.23 0.06 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II6 6.00 3.60 3 6.97 9.00 2.01 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II6 0.18 0.06 3 0.19 0.23 0.12 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II6 2.91 1.39 3 2.77 4.37 1.59 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II6 0.05 0.02 3 0.04 0.07 0.03 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II6 1.85 0.24 3 1.79 2.12 1.65 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II6 1.00 0.50 3 1.26 1.32 0.42 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II6 33.80 2.02 3 33.10 36.08 32.23 135 Lmrf Ultisols Tuffac. sandstone

Dialium guineense Wood density (g/cc) II1C 0.88 0.06 20 0.88 0.97 0.78 115

Dieffenbachia seguine Aluminum (mg/kg) II3A 128 1 121 Lmrf Ultisols Tuffac. sandstone

II3A 39 1 121 Lmrf Ultisols Tuffac. sandstone Petiole

II3C 43 1 121 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

90

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Ash (%) II3A 13.72 1 121 Lmrf Ultisols Tuffac. sandstone

II3A 9.01 1 121 Lmrf Ultisols Tuffac. sandstone Petiole

II3C 7.50 1 121 Lmrf Ultisols Tuffac. sandstone

C/N II3A 15 1 121 Lmrf Ultisols Tuffac. sandstone

II3A 51 1 121 Lmrf Ultisols Tuffac. sandstone

II3C 147 1 121 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II3A 14.17 1 121 Lmrf Ultisols Tuffac. sandstone

II3A 8.66 1 121 Lmrf Ultisols Tuffac. sandstone Petiole

II3C 8.47 1 121 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II3A 47 1 121 Lmrf Ultisols Tuffac. sandstone

II3A 43 1 121 Lmrf Ultisols Tuffac. sandstone Petiole

II3C 42 1 121 Lmrf Ultisols Tuffac. sandstone

Iron (mg/kg) II3A 150 1 121 Lmrf Ultisols Tuffac. sandstone

II3A 28 1 121 Lmrf Ultisols Tuffac. sandstone Petiole

II3C 29 1 121 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II3A 4.37 1 121 Lmrf Ultisols Tuffac. sandstone

II3A 1.98 1 121 Lmrf Ultisols Tuffac. sandstone Petiole

II3C 3.27 1 121 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II3A 0.34 1 121 Lmrf Ultisols Tuffac. sandstone

II3A 0.90 1 121 Lmrf Ultisols Tuffac. sandstone Petiole

II3C 1.78 1 121 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II3A 3.05 1 121 Lmrf Ultisols Tuffac. sandstone

II3A 0.85 1 121 Lmrf Ultisols Tuffac. sandstone Petiole

II3C 0.29 1 121 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II3A 1.68 1 121 Lmrf Ultisols Tuffac. sandstone

II3A 0.86 1 121 Lmrf Ultisols Tuffac. sandstone Petiole

II3C 0.46 1 121 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II3A 44.85 1 121 Lmrf Ultisols Tuffac. sandstone

II3A 40.60 1 121 Lmrf Ultisols Tuffac. sandstone Petiole
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

91

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II3C 24.35 1 121 Lmrf Ultisols Tuffac. sandstone

Dioscorea polygonoides Aluminum (mg/g) II3 0.09 0.04 14 0.09 0.17 0.05 135 Lmrf Ultisols Tuffac. sandstone

C/N II3 44 28 5 38 92 23 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II3 5.18 2.43 14 5.20 8.99 1.89 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II3 45 3 5 45 48 42 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II3 0.12 0.06 14 0.11 0.20 0.03 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II3 4.16 1.93 14 4.54 6.50 0.79 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II3 0.22 0.21 14 0.15 0.69 0.07 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II3 1.53 0.36 18 1.56 1.94 0.51 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II3 1.08 0.27 14 1.04 1.70 0.78 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II3 18.46 5.23 14 17.73 27.74 7.60 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II3 0.30 0.08 5 0.33 0.39 0.17 135 Lmrf Ultisols Tuffac. sandstone

Ditta myricoides Aluminum (mg/g) II1A 0.14 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.31 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.04 1 93 Wflm Inceptisols Tuffac. sandstone

Ash (%) II1B 2.50 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.14 1 93 Wflm Inceptisols Tuffac. sandstone

C/N II1B 35 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 420 1 93 Wflm Inceptisols Tuffac. sandstone

Calcium (mg/g) II1A 5.63 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 3.63 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.89 1 93 Wflm Inceptisols Tuffac. sandstone

Iron (mg/kg) II1A 140 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 70 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 70 1 93 Wflm Inceptisols Tuffac. sandstone

Magnesium (mg/g) II1A 4.42 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 1.05 1 93 Wflm Inceptisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

92

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1C 0.39 1 93 Wflm Inceptisols Tuffac. sandstone

Manganese (mg/g) II1A 0.52 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.15 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.13 1 93 Wflm Inceptisols Tuffac. sandstone

Nitrogen (%) II1B 1.27 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.11 1 93 Wflm Inceptisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.79 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.52 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.20 1 93 Wflm Inceptisols Tuffac. sandstone

Potassium (mg/g) II1A 7.15 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 5.31 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 1.54 1 93 Wflm Inceptisols Tuffac. sandstone

Drypetes glauca Ash (%) II1B 1.53 0.87 3 1.09 2.53 0.97 68 Swf Ultisols Tuffac. sandstone Class 3 Sd

II1B 1.83 1 68 Swf Ultisols Tuffac. sandstone Class 2

Calcium (mg/g) II1B 3.33 1.37 3 2.70 4.90 2.40 68 Swf Ultisols Tuffac. sandstone Class 3 Sd

II1B 4.47 1 68 Swf Ultisols Tuffac. sandstone Class 2

Nitrogen (%) II1B 0.24 0.17 3 0.14 0.44 0.14 68 Swf Ultisols Tuffac. sandstone Class 3 Sd

II1B 0.31 1 68 Swf Ultisols Tuffac. sandstone Class 2

Phosphorus (mg/g) II1B 0.14 0.11 3 0.09 0.28 0.07 68 Swf Ultisols Tuffac. sandstone Class 3 Sd

II1B 0.22 1 68 Swf Ultisols Tuffac. sandstone Class 2

Potassium (mg/g) II1B 0.32 0.18 3 0.28 0.52 0.17 68 Swf Ultisols Tuffac. sandstone Class 3 Sd

II1B 0.30 1 68 Swf Ultisols Tuffac. sandstone Class 2

Enterolobium cyclocarpum Wood density (g/cc) II1B 0.37 0.01 5 0.37 0.38 0.37 97 Wfs Inceptisols Tuffac. sandstone

Erithalis fructicosa Aluminum (mg/g) II2A 0.04 0.04 10 0.02 0.15 0.01 151 Sdf Mollisols Alluvial deposits

Ash (%) II2A 5.02 0.55 10 4.98 6.12 4.03 151 Sdf Mollisols Alluvial deposits

C/N II2A 48 8 10 45 61 41 151 Sdf Mollisols Alluvial deposits
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

93

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Calcium (mg/g) II2A 9.63 4.62 2 9.71 13.68 13.68 104 Sdf Mollisols Alluvial deposits New

II2A 11.08 2.90 11 11.53 14.87 2.88 104 Sdf Mollisols Alluvial deposits Mature

II2A 7.68 1.58 10 8.39 9.61 4.97 151 Sdf Mollisols Alluvial deposits

II2A 10.85 3.16 25 11.53 14.87 2.88 149 Sdf Mollisols Alluvial deposits Guanica Forest

Carbon (%) II2A 57 1 10 57 58 56 151 Sdf Mollisols Alluvial deposits

Iron (mg/kg) II2A 35 34 10 25 132 16 151 Sdf Mollisols Alluvial deposits

Magnesium (mg/g) II2A 0.85 0.46 2 0.83 1.34 0.42 104 Sdf Mollisols Alluvial deposits New

II2A 2.72 0.63 11 2.60 4.09 1.66 104 Sdf Mollisols Alluvial deposits Mature

II2A 1.18 0.22 10 1.23 1.50 0.75 151 Sdf Mollisols Alluvial deposits

II2A 2.42 0.92 25 2.57 4.09 0.42 149 Sdf Mollisols Alluvial deposits Guanica Forest

Manganese (mg/g) II2A 0.03 0.01 10 0.03 0.06 0.02 151 Sdf Mollisols Alluvial deposits

Nitrogen (%) II2A 0.87 0.08 2 0.87 0.94 0.80 104 Sdf Mollisols Alluvial deposits New

II2A 0.72 0.09 11 0.71 0.91 0.62 104 Sdf Mollisols Alluvial deposits Mature

II2A 1.20 0.17 10 1.26 1.37 0.92 151 Sdf Mollisols Alluvial deposits

II2A 0.75 0.10 26 0.76 0.94 0.62 149 Sdf Mollisols Alluvial deposits Guanica Forest

Phosphorus (mg/g) II2A 0.54 0.24 2 0.54 0.75 0.33 104 Sdf Mollisols Alluvial deposits New

II2A 0.25 0.06 11 0.26 0.34 0.15 104 Sdf Mollisols Alluvial deposits Mature

II2A 0.26 0.06 10 0.24 0.34 0.19 151 Sdf Mollisols Alluvial deposits

II2A 0.29 0.15 26 0.27 0.75 0.15 149 Sdf Mollisols Alluvial deposits Guanica Forest

Potassium (mg/g) II2A 8.34 0.73 2 8.29 9.13 7.65 104 Sdf Mollisols Alluvial deposits New

II2A 9.36 1.97 11 8.46 12.79 6.76 104 Sdf Mollisols Alluvial deposits Mature

II2A 6.78 1.58 10 6.88 8.93 4.39 151 Sdf Mollisols Alluvial deposits

II2A 9.20 1.86 26 8.46 12.79 6.76 149 Sdf Mollisols Alluvial deposits Guanica Forest

Sulfur (%) II2A 0.22 0.03 10 0.22 0.27 0.17 151 Sdf Mollisols Alluvial deposits

Erythrodes plantaginea Aluminum (mg/g) II6 3.03 1 135 Lmrf Ultisols Tuffac. sandstone

C/N II6 17 5 2 17 21 14 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II6 5.21 1 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II6 43 2 2 43 44 42 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

94

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Iron (mg/g) II6 3.59 1 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II6 2.31 1 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II6 0.21 1 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II6 2.54 0.62 2 2.54 2.97 2.10 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II6 2.85 1 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II6 18.63 1 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II6 0.20 0.09 2 0.20 0.27 0.13 135 Lmrf Ultisols Tuffac. sandstone

Erythroxylum aerolatum Calcium (mg/g) II1A 15.73 1 104 Sdf Mollisols Alluvial deposits New

II1A 20.26 2.11 6 19.95 23.50 17.06 104 Sdf Mollisols Alluvial deposits

II1Ci 20.70 1 104 Sdf Mollisols Alluvial deposits

Magnesium (mg/g) II1A 1.63 1 104 Sdf Mollisols Alluvial deposits New

II1A 3.33 0.51 7 3.34 4.12 2.32 104 Sdf Mollisols Alluvial deposits

II1Ci 0.48 1 104 Sdf Mollisols Alluvial deposits

Nitrogen (%) II1A 2.26 1 104 Sdf Mollisols Alluvial deposits New

II1A 1.69 0.21 6 1.76 1.92 1.40 104 Sdf Mollisols Alluvial deposits

Phosphorus (mg/g) II1A 1.38 1 104 Sdf Mollisols Alluvial deposits New

II1A 0.81 0.15 7 0.78 1.07 0.63 104 Sdf Mollisols Alluvial deposits

II1Ci 0.26 1 104 Sdf Mollisols Alluvial deposits

Potassium (mg/g) II1A 15.06 1 104 Sdf Mollisols Alluvial deposits New

II1A 13.68 2.60 7 13.65 18.53 9.29 104 Sdf Mollisols Alluvial deposits

II1Ci 7.83 1 104 Sdf Mollisols Alluvial deposits

Calcium (mg/g) II1A 19.61 2.55 14 19.57 23.50 15.40 149 Sdf Mollisols Alluvial deposits Guanica Forest

Magnesium (mg/g) II1A 3.12 0.75 16 3.33 4.12 1.61 149 Sdf Mollisols Alluvial deposits Guanica Forest

Nitrogen (%) II1A 1.87 0.39 16 1.85 2.63 1.40 149 Sdf Mollisols Alluvial deposits Guanica Forest

Phosphorus (mg/g) II1A 0.88 0.24 16 0.83 1.38 0.63 149 Sdf Mollisols Alluvial deposits Guanica Forest

Potassium (mg/g) II1A 13.85 2.47 16 13.74 18.53 9.29 149 Sdf Mollisols Alluvial deposits Guanica Forest

Erythroxylum rotundifolium Calcium (mg/g) II1A 14.40 1 104 Sdf Mollisols Alluvial deposits
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

95

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 14.40 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 20.70 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 23.24 1 104 Sdf Mollisols Alluvial deposits

II1Ci 23.24 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Magnesium (mg/g) II1A 1.34 1 104 Sdf Mollisols Alluvial deposits

II1A 1.34 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 0.48 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 0.19 1 104 Sdf Mollisols Alluvial deposits

II1Ci 0.19 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Nitrogen (%) II1A 1.65 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 0.89 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 0.85 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Phosphorus (mg/g) II1A 0.34 1 104 Sdf Mollisols Alluvial deposits

II1A 0.34 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 0.26 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 0.21 1 104 Sdf Mollisols Alluvial deposits

II1Ci 0.21 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Potassium (mg/g) II1A 9.76 1 104 Sdf Mollisols Alluvial deposits

II1A 9.76 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 7.83 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 5.86 1 104 Sdf Mollisols Alluvial deposits

II1Ci 5.86 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Eucalyptus pathentinervis Ash (%) II8A 6.28 1.65 4 6.28 8.74 5.31 90 Swf Ultisols Tuffac. sandstone Torn

II8A 4.67 0.42 3 4.61 5.12 4.29 90 Swf Ultisols Tuffac. sandstone Recently fall

II8B 3.92 0.50 3 3.68 4.50 3.58 90 Swf Ultisols Tuffac. sandstone

II8C 4.60 0.29 3 4.51 4.92 4.36 90 Swf Ultisols Tuffac. sandstone

II8D 10.49 2.43 3 10.66 12.84 7.98 90 Swf Ultisols Tuffac. sandstone

Calcium (mg/g) II8A 15.13 1.27 4 15.13 16.12 13.44 90 Swf Ultisols Tuffac. sandstone Torn
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

96

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8A 14.84 0.40 3 14.90 15.20 14.41 90 Swf Ultisols Tuffac. sandstone Recently fall

II8B 12.95 2.52 3 11.93 15.82 11.09 90 Swf Ultisols Tuffac. sandstone

II8C 11.57 0.43 3 11.34 12.07 11.30 90 Swf Ultisols Tuffac. sandstone

II8D 14.52 1.23 3 14.53 15.75 13.29 90 Swf Ultisols Tuffac. sandstone

II9A 9.14 2.20 22 9.51 12.01 1.38 139 Wfs Ultisols Tuffac. sandstone

II9A 10.68 2.66 22 10.31 17.23 6.73 139 Wfs Ultisols Tuffac. sandstone Other

II9B 9.24 2.03 5 9.04 12.49 7.04 139 Wfs Ultisols Tuffac. sandstone

II9C 6.94 0.91 6 7.41 7.70 5.42 139 Wfs Ultisols Tuffac. sandstone

II9D 5.97 0.94 5 5.79 7.54 5.11 139 Wfs Ultisols Tuffac. sandstone

Magnesium (mg/g) II9A 3.09 0.68 22 3.00 5.58 2.17 139 Wfs Ultisols Tuffac. sandstone

II9A 3.34 0.85 22 3.31 5.60 2.23 139 Wfs Ultisols Tuffac. sandstone Other

II9B 3.27 0.83 5 2.88 4.62 2.58 139 Wfs Ultisols Tuffac. sandstone

II9C 3.56 0.43 6 3.65 3.96 2.80 139 Wfs Ultisols Tuffac. sandstone

II9D 2.70 0.40 5 2.80 3.12 2.04 139 Wfs Ultisols Tuffac. sandstone

Nitrogen (%) II8A 0.78 0.26 4 0.78 1.16 0.56 90 Swf Ultisols Tuffac. sandstone Torn

II8A 0.50 0.07 3 0.50 0.56 0.43 90 Swf Ultisols Tuffac. sandstone Recently fall

II8B 0.32 0.05 3 0.34 0.35 0.26 90 Swf Ultisols Tuffac. sandstone

II8C 0.73 0.06 3 0.73 0.79 0.68 90 Swf Ultisols Tuffac. sandstone

II8D 1.23 0.20 3 1.15 1.46 1.09 90 Swf Ultisols Tuffac. sandstone

II9A 0.52 0.16 22 0.47 0.98 0.34 139 Wfs Ultisols Tuffac. sandstone

II9A 1.67 0.29 22 1.59 2.40 1.32 139 Wfs Ultisols Tuffac. sandstone Other

II9B 0.35 0.26 10 0.27 0.82 0.13 139 Wfs Ultisols Tuffac. sandstone

II9C 0.41 0.06 11 0.39 0.51 0.31 139 Wfs Ultisols Tuffac. sandstone

II9D 0.95 0.23 10 0.97 1.25 0.56 139 Wfs Ultisols Tuffac. sandstone

Phosphorus (mg/g) II8A 0.26 0.09 4 0.26 0.39 0.18 90 Swf Ultisols Tuffac. sandstone Torn

II8A 0.14 0.01 3 0.14 0.15 0.13 90 Swf Ultisols Tuffac. sandstone Recently fall

II8B 0.09 0.03 3 0.11 0.11 0.06 90 Swf Ultisols Tuffac. sandstone

II8B 0.10 0.03 3 0.11 0.11 0.06 90 Swf Ultisols Tuffac. sandstone

II8C 0.30 0.03 3 0.31 0.32 0.26 90 Swf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

97

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8D 0.48 0.11 3 0.44 0.60 0.39 90 Swf Ultisols Tuffac. sandstone

II9A 0.19 0.08 22 0.17 0.36 0.08 139 Wfs Ultisols Tuffac. sandstone

II9A 0.62 0.26 22 0.52 1.54 0.42 139 Wfs Ultisols Tuffac. sandstone Other

II9B 0.10 0.43 5 0.08 0.17 0.07 139 Wfs Ultisols Tuffac. sandstone

II9C 0.27 0.09 6 0.26 0.39 0.19 139 Wfs Ultisols Tuffac. sandstone

II9D 0.55 0.07 5 0.54 0.62 0.46 139 Wfs Ultisols Tuffac. sandstone

Potassium (mg/g) II8A 3.14 0.85 4 3.14 3.97 2.15 90 Swf Ultisols Tuffac. sandstone Torn

II8A 2.68 0.57 3 2.45 3.33 2.26 90 Swf Ultisols Tuffac. sandstone Recently fall

II8B 1.87 0.48 3 1.88 2.34 1.38 90 Swf Ultisols Tuffac. sandstone

II8C 2.47 0.74 3 2.11 3.32 1.99 90 Swf Ultisols Tuffac. sandstone

II8D 3.19 0.90 3 2.70 4.23 2.71 90 Swf Ultisols Tuffac. sandstone

II9A 3.48 0.97 22 3.33 5.17 0.80 139 Wfs Ultisols Tuffac. sandstone

II9A 4.84 1.49 22 4.67 8.76 2.06 139 Wfs Ultisols Tuffac. sandstone Other

II9B 2.91 0.98 5 2.59 4.64 2.28 139 Wfs Ultisols Tuffac. sandstone

II9C 3.56 0.21 6 3.50 3.84 3.28 139 Wfs Ultisols Tuffac. sandstone

II9D 3.26 0.56 5 3.20 4.19 2.71 139 Wfs Ultisols Tuffac. sandstone

Eucalyptus robusta Aluminum (mg/g) II8 2.76 2.53 19 1.58 7.92 0.19 152 Mfs Histosol Alluvial deposits

Ash (%) II8 29.27 24.65 19 17.42 83.48 5.63 152 Mfs Histosol Alluvial deposits

C/N II8 39 17 19 35 86 18 152 Mfs Histosol Alluvial deposits

Calcium (mg/g) II8 28.05 8.39 19 26.30 41.38 16.60 152 Mfs Histosol Alluvial deposits

Carbon (%) II8 42 14 19 50 53 12 152 Mfs Histosol Alluvial deposits

Iron (mg/g) II8 4.78 4.58 19 2.49 13.18 0.26 152 Mfs Histosol Alluvial deposits

Magnesium (mg/g) II8 3.49 1.20 19 2.863 6.59 2.32 152 Mfs Histosol Alluvial deposits

Manganese (mg/g) II8 0.30 0.11 19 0.25 0.55 0.16 152 Mfs Histosol Alluvial deposits

Nitrogen (%) II8 1.16 0.45 19 1.13 2.21 0.45 152 Mfs Histosol Alluvial deposits

Phosphorus (mg/g) II8 0.32 0.11 19 0.27 0.59 0.17 152 Mfs Histosol Alluvial deposits

Potassium (mg/g) II8 2.04 0.39 19 1.89 2.78 1.45 152 Mfs Histosol Alluvial deposits

Sulfur (%) II8 0.26 0.10 19 0.26 0.47 0.09 152 Mfs Histosol Alluvial deposits
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

98

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Wood density (g/cc) II1B 0.85 0.03 3 0.85 0.87 0.82 97 Wfs Inceptisols Tuffac. sandstone

Eucalyptus saligna Ash (%) II8A 3.14 1 90 Swf Ultisols Tuffac. sandstone New

II8A 3.79 1 90 Swf Ultisols Tuffac. sandstone Old

II8A 5.15 0.61 5 5.16 5.79 4.19 90 Swf Ultisols Tuffac. sandstone Seeds

II8B 4.44 1.25 3 5.08 5.23 3.00 90 Swf Ultisols Tuffac. sandstone

II8C 4.52 1.66 3 4.27 6.29 3.00 90 Swf Ultisols Tuffac. sandstone Torn

II8D 12.39 4.66 3 11.33 17.49 8.35 90 Swf Ultisols Tuffac. sandstone

Calcium (mg/g) II8A 6.13 1 90 Swf Ultisols Tuffac. sandstone New

II8A 11.14 1 90 Swf Ultisols Tuffac. sandstone Old

II8C 12.45 2.04 5 13.37 13.82 9.62 90 Swf Ultisols Tuffac. sandstone Seeds

II8B 12.96 4.05 3 15.23 15.37 8.28 90 Swf Ultisols Tuffac. sandstone

II8C 7.12 1.22 3 7.20 8.29 5.86 90 Swf Ultisols Tuffac. sandstone Torn

II8D 11.51 0.56 3 11.66 11.98 10.89 90 Swf Ultisols Tuffac. sandstone

II9A 7.97 0.92 22 7.98 10.14 6.08 139 Wfs Ultisols Tuffac. sandstone

II9A 12.16 3.82 21 10.72 22.77 7.90 139 Wfs Ultisols Tuffac. sandstone Other

II9B 5.35 1.86 6 5.87 7.23 2.38 139 Wfs Ultisols Tuffac. sandstone

II9C 4.87 0.53 6 4.80 5.70 4.28 139 Wfs Ultisols Tuffac. sandstone

II9D 7.07 0.81 5 7.47 7.88 6.14 139 Wfs Ultisols Tuffac. sandstone

Magnesium (mg/g) II8A 2.28 1 90 Swf Ultisols Tuffac. sandstone New

II8A 2.69 1 90 Swf Ultisols Tuffac. sandstone Old

II8C 2.31 0.42 5 2.37 2.94 1.85 90 Swf Ultisols Tuffac. sandstone Seeds

II8B 1.75 0.47 3 1.93 2.10 1.22 90 Swf Ultisols Tuffac. sandstone

II8C 1.70 0.48 3 1.54 2.23 1.31 90 Swf Ultisols Tuffac. sandstone Torn

II8D 2.52 0.63 3 2.33 2.81 2.01 90 Swf Ultisols Tuffac. sandstone

II9A 2.66 0.40 22 2.66 3.51 1.94 139 Wfs Ultisols Tuffac. sandstone

II9A 4.38 1.06 21 4.23 6.69 2.44 139 Wfs Ultisols Tuffac. sandstone Other

II9B 1.89 0.70 6 2.03 2.42 0.52 139 Wfs Ultisols Tuffac. sandstone

II9C 2.29 0.37 6 2.30 2.87 1.82 139 Wfs Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

99

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II9D 2.82 0.34 5 2.93 3.27 2.40 139 Wfs Ultisols Tuffac. sandstone

Nitrogen (%) II8A 0.92 1 90 Swf Ultisols Tuffac. sandstone New

II8A 0.79 1 90 Swf Ultisols Tuffac. sandstone Old

II8C 0.73 0.06 5 0.75 0.78 0.67 90 Swf Ultisols Tuffac. sandstone Seeds

II8B 0.85 0.10 3 0.81 0.97 0.75 90 Swf Ultisols Tuffac. sandstone

II8C 1.57 0.16 3 1.59 1.71 1.40 90 Swf Ultisols Tuffac. sandstone Torn

II8D 1.20 0.09 3 1.25 1.25 1.09 90 Swf Ultisols Tuffac. sandstone Loose litter

II9A 0.84 0.23 22 0.78 1.32 0.55 139 Wfs Ultisols Tuffac. sandstone

II9A 1.59 0.43 22 1.73 2.41 0.86 139 Wfs Ultisols Tuffac. sandstone Other

II9B 0.26 0.09 11 0.26 0.42 0.14 139 Wfs Ultisols Tuffac. sandstone

II9C 0.49 0.10 11 0.46 0.72 0.38 139 Wfs Ultisols Tuffac. sandstone

II9D 1.17 0.17 10 1.21 1.37 0.76 139 Wfs Ultisols Tuffac. sandstone

Phosphorus (mg/g) II8A 0.35 1 90 Swf Ultisols Tuffac. sandstone New

II8A 0.17 1 90 Swf Ultisols Tuffac. sandstone Old

II8C 0.27 0.04 5 0.26 0.32 0.23 90 Swf Ultisols Tuffac. sandstone Seeds

II8B 0.14 0.01 3 0.14 0.15 0.14 90 Swf Ultisols Tuffac. sandstone

II8C 0.33 0.02 3 0.33 0.35 0.31 90 Swf Ultisols Tuffac. sandstone Torn

II8D 0.40 0.02 3 0.40 0.43 0.39 90 Swf Ultisols Tuffac. sandstone

II9A 0.29 0.12 22 0.27 0.56 0.12 139 Wfs Ultisols Tuffac. sandstone

II9A 0.61 0.23 21 0.63 1.14 0.25 139 Wfs Ultisols Tuffac. sandstone Other

II9B 0.07 0.04 6 0.55 0.15 0.03 139 Wfs Ultisols Tuffac. sandstone

II9C 0.30 0.04 6 0.30 0.37 0.26 139 Wfs Ultisols Tuffac. sandstone

II9D 0.58 0.07 5 0.63 0.63 0.46 139 Wfs Ultisols Tuffac. sandstone

Potassium (mg/g) II8A 2.89 1 90 Swf Ultisols Tuffac. sandstone New

II8A 1.37 1 90 Swf Ultisols Tuffac. sandstone Old

II8C 2.34 1.08 5 1.67 3.53 1.38 90 Swf Ultisols Tuffac. sandstone Seeds

II8B 1.78 0.95 3 1.59 2.81 0.94 90 Swf Ultisols Tuffac. sandstone

II8C 1.42 0.42 3 1.29 1.89 1.08 90 Swf Ultisols Tuffac. sandstone Torn

II8D 1.95 0.96 3 1.71 3.01 1.13 90 Swf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

100

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II9A 3.68 1.14 22 3.36 7.48 2.49 139 Wfs Ultisols Tuffac. sandstone

II9A 5.53 1.87 21 5.27 9.93 2.72 139 Wfs Ultisols Tuffac. sandstone Other

II9B 1.87 0.91 6 1.54 3.68 1.28 139 Wfs Ultisols Tuffac. sandstone

II9C 3.03 0.68 6 2.76 4.24 2.42 139 Wfs Ultisols Tuffac. sandstone

II9D 3.44 1.27 5 2.66 4.91 2.30 139 Wfs Ultisols Tuffac. sandstone

Eugeia eggersii Calcium (mg/g) II1A 7.45 1 135 Lmrf Ultisols Tuffac. sandstone

II1B 4.97 1 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.69 1 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II1A 2.15 1 135 Lmrf Ultisols Tuffac. sandstone

II1B 0.77 1 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.19 1 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II1A 1.47 1 135 Lmrf Ultisols Tuffac. sandstone

II1B 0.54 1 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.25 1 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.67 1 135 Lmrf Ultisols Tuffac. sandstone

II1B 0.32 1 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II1A 8.61 1 135 Lmrf Ultisols Tuffac. sandstone

II1B 4.22 1 135 Lmrf Ultisols Tuffac. sandstone

II1C 3.47 1 135 Lmrf Ultisols Tuffac. sandstone

Eugenia borinquensis Aluminum (mg/g) II1A 0.18 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.11 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.13 1 93 Wflm Inceptisols Tuffac. sandstone

Ash (%) II1A 2.88 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 3.73 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.78 1 93 Wflm Inceptisols Tuffac. sandstone

C/N II1A 55 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 66 1 93 Wflm Inceptisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

101

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1C 200 1 93 Wflm Inceptisols Tuffac. sandstone

Calcium (mg/g) II1A 2.97 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 7.09 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 2.71 1 93 Wflm Inceptisols Tuffac. sandstone

Iron (mg/g) II1A 0.07 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.11 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.11 1 93 Wflm Inceptisols Tuffac. sandstone

Magnesium (mg/g) II1A 1.60 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 1.92 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.57 1 93 Wflm Inceptisols Tuffac. sandstone

Manganese (mg/g) II1A 0.16 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.12 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.03 1 93 Wflm Inceptisols Tuffac. sandstone

Nitrogen (%) II1A 0.82 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.67 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.23 1 93 Wflm Inceptisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.55 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.93 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 2.00 1 93 Wflm Inceptisols Tuffac. sandstone

Potassium (mg/g) II1A 8.87 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 9.16 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 3.30 1 93 Wflm Inceptisols Tuffac. sandstone

Eugenia eggersii Aluminum (mg/g) II1A 0.22 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.06 0.03 2 0.06 0.08 0.04 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.45 1 93 Wflm Inceptisols Tuffac. sandstone

Ash (%) II1A 5.25 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 3.15 0.08 2 3.15 3.20 3.09 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.45 1 93 Wflm Inceptisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

102

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

C/N II1A 35 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 96 12 2 96 104 88 93 Wflm Inceptisols Tuffac. sandstone

II1C 382 1 93 Wflm Inceptisols Tuffac. sandstone

Calcium (mg/g) II1A 9.08 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 4.91 2.80 2 4.91 6.89 2.93 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.86 1 93 Wflm Inceptisols Tuffac. sandstone

Iron (mg/g) II1A 0.14 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.09 0.03 2 0.09 0.11 0.07 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.05 1 93 Wflm Inceptisols Tuffac. sandstone

Magnesium (mg/g) II1A 3.96 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 1.23 0.16 2 1.23 1.35 1.12 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.32 1 93 Wflm Inceptisols Tuffac. sandstone

Manganese (mg/g) II1A 0.11 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.04 0.02 2 0.36 0.05 0.03 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.18 1 93 Wflm Inceptisols Tuffac. sandstone

Nitrogen (%) II1A 1.24 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.47 0.06 2 0.47 0.51 0.43 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.12 1 93 Wflm Inceptisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.65 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.37 0.15 2 0.37 0.48 0.26 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.15 1 93 Wflm Inceptisols Tuffac. sandstone

Potassium (mg/g) II1A 10.28 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 6.21 1.97 2 6.21 7.61 4.82 93 Wflm Inceptisols Tuffac. sandstone

II1C 1.86 1 93 Wflm Inceptisols Tuffac. sandstone

Eugenia foetida Calcium (mg/g) II1A 13.69 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 19.73 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Magnesium (mg/g) II1A 3.74 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 1.66 1 149 Sdf Mollisols Alluvial deposits Guanica Forest
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

103

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Nitrogen (%) II1A 1.42 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 0.82 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Phosphorus (mg/g) II1A 0.38 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 0.26 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Potassium (mg/g) II1A 11.13 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 7.28 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Eugenia maleolens Calcium (mg/g) II1A 13.69 1 104 Sdf Mollisols Alluvial deposits

II1Ci 19.73 1 104 Sdf Mollisols Alluvial deposits

Magnesium (mg/g) II1A 3.74 1 104 Sdf Mollisols Alluvial deposits

II1Ci 1.66 1 104 Sdf Mollisols Alluvial deposits

Phosphorus (mg/g) II1A 0.38 1 104 Sdf Mollisols Alluvial deposits

II1Ci 0.26 1 104 Sdf Mollisols Alluvial deposits

Potassium (mg/g) II1A 11.13 1 104 Sdf Mollisols Alluvial deposits

II1Ci 7.28 1 104 Sdf Mollisols Alluvial deposits

Eugenia monticola Ash (%) II1A 4.58 0.38 5 4.76 4.92 3.96 126 Lmrf Ultisols Tuffac. sandstone Mameyes, Luquillo

II1C 1.63 0.29 5 1.67 2.01 1.24 126 Lmrf Ultisols Tuffac. sandstone Mameyes, Luquillo

II1Ci 6.85 1.38 6 6.85 9.04 4.70 126 Lmrf Ultisols Tuffac. sandstone Mameyes, Luquillo

C/N II1A 36 1 5 36 37 36 126 Lmrf Ultisols Tuffac. sandstone Mameyes, Luquillo

II1C 143 28 5 141 178 104 126 Lmrf Ultisols Tuffac. sandstone Mameyes, Luquillo

II1Ci 49 3 6 49 53 45 126 Lmrf Ultisols Tuffac. sandstone Mameyes, Luquillo

Carbon (%) II1A 54 0 5 54 54 54 126 Lmrf Ultisols Tuffac. sandstone Mameyes, Luquillo

II1C 52 0 5 52 53 52 126 Lmrf Ultisols Tuffac. sandstone Mameyes, Luquillo

II1Ci 51 1 6 52 52 49 126 Lmrf Ultisols Tuffac. sandstone Mameyes, Luquillo

Nitrogen (%) II1A 1.48 0.02 5 1.49 1.51 1.45 126 Lmrf Ultisols Tuffac. sandstone Mameyes, Luquillo

II1C 0.38 0.08 5 0.37 0.50 0.29 126 Lmrf Ultisols Tuffac. sandstone Mameyes, Luquillo

II1Ci 1.05 0.04 6 1.05 1.10 0.98 126 Lmrf Ultisols Tuffac. sandstone Mameyes, Luquillo

Sulfur (%) II1A 0.16 0.01 5 0.15 0.17 0.15 126 Lmrf Ultisols Tuffac. sandstone Mameyes, Luquillo
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

104

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1C 0.07 0.01 5 0.06 0.08 0.06 126 Lmrf Ultisols Tuffac. sandstone Mameyes, Luquillo

II1Ci 0.18 0.03 6 0.19 0.22 0.13 126 Lmrf Ultisols Tuffac. sandstone Mameyes, Luquillo

Wood density (g/cc) II1C 0.71 0.03 5 0.72 0.74 0.66 126 Lmrf Ultisols Tuffac. sandstone Mameyes, Luquillo

Eugenia rhombea Calcium (mg/g) II1Ci 25.00 1 104 Sdf Mollisols Alluvial deposits

II1Ci 25.00 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Magnesium (mg/g) II1Ci 0.49 1 104 Sdf Mollisols Alluvial deposits

II1Ci 0.49 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Nitrogen (%) II1A 0.97 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 0.55 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Phosphorus (mg/g) II1A 0.26 1 104 Sdf Mollisols Alluvial deposits

II1A 0.26 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 0.13 1 104 Sdf Mollisols Alluvial deposits

II1Ci 0.13 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Potassium (mg/g) II1A 12.41 1 104 Sdf Mollisols Alluvial deposits

II1A 12.41 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 5.88 1 104 Sdf Mollisols Alluvial deposits

II1Ci 5.88 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Eugenia stahlii Ash (%) II1A 7.34 0.90 3 7.00 8.36 6.67 94 Swf Ultisols Tuffac. sandstone

II1B 8.59 0.59 3 8.52 9.19 8.02 94 Swf Ultisols Tuffac. sandstone

II1C 1.58 0.11 2 1.58 1.65 1.50 94 Swf Ultisols Tuffac. sandstone

Nitrogen (%) II1A 0.76 0.07 3 0.76 0.83 0.69 94 Swf Ultisols Tuffac. sandstone

II1B 0.46 0.03 3 0.47 0.48 0.42 94 Swf Ultisols Tuffac. sandstone

II1C 0.22 0.01 2 0.22 0.22 0.21 94 Swf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.46 0.07 3 0.43 0.53 0.42 94 Swf Ultisols Tuffac. sandstone

II1B 0.42 0.08 3 0.40 0.51 0.34 94 Swf Ultisols Tuffac. sandstone

II1C 0.19 0.00 2 0.19 0.19 0.19 94 Swf Ultisols Tuffac. sandstone

Potassium (mg/g) II1A 5.56 1.58 3 4.96 7.36 4.37 94 Swf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

105

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1B 0.50 0.26 3 0.37 0.80 0.32 94 Swf Ultisols Tuffac. sandstone

II1C 1.19 0.15 2 1.19 1.29 1.08 94 Swf Ultisols Tuffac. sandstone

Exostema caribaea Aluminum (mg/kg) II1A 44 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 55 20 11 63 77 19 151 Sdf Mollisols Alluvial deposits

II1A 63 1 102 Smf Limestone Tuffac. sandstone Adult 

Ash (%) II1A 3.93 1 102 Smf Limestone Tuffac. sandstone Adult 

II1A 3.97 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 3.83 0.09 11 3.81 3.98 3.70 151 Sdf Mollisols Alluvial deposits

C/N II1A 21 1 11 21 22 20 151 Sdf Mollisols Alluvial deposits

Calcium (mg/g) II1A 14.02 6.68 14 11.24 23.11 5.60 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1A 5.79 1 104 Sdf Mollisols Alluvial deposits New

II1A 15.39 6.19 6 14.95 23.11 8.36 104 Sdf Mollisols Alluvial deposits

II1A 4.46 1 102 Smf Limestone Tuffac. sandstone Adult 

II1A 5.56 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 7.61 0.66 11 7.74 8.56 6.44 151 Sdf Mollisols Alluvial deposits

II1Ci 16.30 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 16.30 1 104 Sdf Mollisols Alluvial deposits

Carbon (%) II1A 56 1 11 56 57 55 151 Sdf Mollisols Alluvial deposits

Cobalt (μg/g) II1A 0.43 1 102 Smf Limestone Tuffac. sandstone Adult 

Iron (mg/g) II1A 0.10 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 0.07 0.02 11 0.08 0.10 0.04 151 Sdf Mollisols Alluvial deposits

II1A 0.10 1 102 Smf Limestone Tuffac. sandstone Adult 

Magnesium (mg/g) II1A 2.07 1.14 16 1.67 4.73 1.05 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1A 1.07 1 104 Sdf Mollisols Alluvial deposits New

II1A 2.21 1.15 7 1.93 4.73 1.27 104 Sdf Mollisols Alluvial deposits

II1A 1.37 1 102 Smf Limestone Tuffac. sandstone Adult 

II1A 7.04 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 0.92 0.07 11 0.90 1.05 0.81 151 Sdf Mollisols Alluvial deposits
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

106

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1Ci 0.28 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 0.28 1 104 Sdf Mollisols Alluvial deposits

Manganese (mg/g) II1A 0.01 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 0.02 0.01 11 0.02 0.03 0.01 151 Sdf Mollisols Alluvial deposits

II1A 0.04 1 102 Smf Limestone Tuffac. sandstone Adult 

Nickel (μg/g) II1A 12.80 1 102 Smf Limestone Tuffac. sandstone Adult 

Nitrogen (%) II1A 1.57 0.14 14 1.54 1.77 1.35 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1A 1.77 1 104 Sdf Mollisols Alluvial deposits New

II1A 1.54 0.13 6 1.52 1.77 1.35 104 Sdf Mollisols Alluvial deposits

II1A 2.08 1 102 Smf Limestone Tuffac. sandstone Adult 

II1A 1.38 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 2.66 0.10 11 2.67 2.82 2.52 151 Sdf Mollisols Alluvial deposits

II1Ci 0.63 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Phosphorus (mg/g) II1A 0.48 0.11 16 0.45 0.70 0.36 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1A 0.63 1 104 Sdf Mollisols Alluvial deposits New

II1A 0.46 0.11 7 0.44 0.70 0.36 104 Sdf Mollisols Alluvial deposits

II1A 0.86 1 102 Smf Limestone Tuffac. sandstone Adult 

II1A 0.43 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 0.43 0.04 11 0.43 0.50 0.39 151 Sdf Mollisols Alluvial deposits

II1Ci 0.14 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 0.14 1 104 Sdf Mollisols Alluvial deposits

Potassium (mg/g) II1A 10.25 1.79 16 9.43 13.20 7.88 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1A 12.55 1 104 Sdf Mollisols Alluvial deposits New

II1A 9.92 1.66 7 9.35 13.20 7.88 104 Sdf Mollisols Alluvial deposits

II1A 11.97 1 102 Smf Limestone Tuffac. sandstone Adult 

II1A 5.32 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 4.52 0.37 11 4.51 5.41 4.10 151 Sdf Mollisols Alluvial deposits

II1Ci 7.48 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 7.48 1 104 Sdf Mollisols Alluvial deposits
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

107

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Sulfur (%) II1A 0.23 0.02 11 0.23 0.25 0.17 151 Sdf Mollisols Alluvial deposits

Zinc (μg/g) II1A 21.90 1 149 Sdf Mollisols Alluvial deposits Susua forest

Faramea occidentalis Ash (%) II1A 8.60 0.69 6 8.55 9.51 7.73 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 2.30 0.57 6 2.48 2.72 1.24 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 8.49 0.67 5 8.60 9.39 7.61 126 Ultisols Siltstone,sandstone Urban forest, RP

C/N II1A 27 1 6 27 28 26 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 135 25 6 143 164 98 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 32 6 5 32 38 22 126 Ultisols Siltstone,sandstone Urban forest, RP

Carbon (%) II1A 49 0 6 49 49 48 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 51 0 6 51 52 50 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 49 1 5 49 50 48 126 Ultisols Siltstone,sandstone Urban forest, RP

Nitrogen (%) II1A 1.81 0.04 6 1.81 1.86 1.75 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 0.39 0.08 6 0.36 0.52 0.31 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 1.60 0.37 5 1.57 2.22 1.27 126 Ultisols Siltstone,sandstone Urban forest, RP

Sulfur (%) II1A 0.26 0.02 6 0.25 0.28 0.24 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 0.07 0.01 6 0.06 0.09 0.05 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 0.22 0.07 5 0.19 0.34 0.17 126 Ultisols Siltstone,sandstone Urban forest, RP

Wood density (g/cc) II1C 0.62 0.08 5 0.62 0.72 0.53 126 Ultisols Siltstone,sandstone Urban forest, RP

Ficus citrifolia Aluminum (mg/kg) II1A 20 1 55 Dfs Mangle Dune deposits

II8A 73 63 3 54 143 22 55 Dfs Mangle Dune deposits

II8A 141 1 55 Dfs Mangle Dune deposits Mona Island

II8D 145 100 2 145 216 74 55 Dfs Mangle Dune deposits

Ash (%) II1A 13.98 0.27 5 13.95 14.37 13.63 55 Dfs Mangle Dune deposits

II8A 16.98 0.33 3 16.79 17.37 16.79 55 Dfs Mangle Dune deposits

II8A 8.48 1 55 Dfs Mangle Dune deposits Mona Island

II8D 39.33 14.21 2 39.33 49.38 29.28 55 Dfs Mangle Dune deposits

Calcium (mg/g) II1A 20.26 1.60 5 20.08 22.78 18.68 55 Dfs Mangle Dune deposits
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

108

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8A 36.69 6.21 3 35.15 43.53 31.40 55 Dfs Mangle Dune deposits

II8A 21.88 1 55 Dfs Mangle Dune deposits Mona Island

II8D 103.03 29.12 2 103.03 123.62 82.44 55 Dfs Mangle Dune deposits

Iron (mg/g) II1A 0.04 0.01 5 0.04 0.04 0.03 55 Dfs Mangle Dune deposits

II8A 0.09 0.02 3 0.10 0.11 0.07 55 Dfs Mangle Dune deposits

II8A 0.12 1 55 Dfs Mangle Dune deposits Mona Island

II8D 0.26 0.00 2 0.26 0.26 0.25 55 Dfs Mangle Dune deposits

Magnesium (mg/g) II1A 2.87 0.13 5 2.82 3.07 2.72 55 Dfs Mangle Dune deposits

II8A 3.65 0.32 3 3.61 3.99 3.35 55 Dfs Mangle Dune deposits

II8A 1.88 1 55 Dfs Mangle Dune deposits Mona Island

II8D 4.85 2.10 2 4.85 6.33 3.36 55 Dfs Mangle Dune deposits

Manganese (mg/kg) II1A 28 5 5 27 36 25 55 Dfs Mangle Dune deposits

II8A 51 13 3 45 65 42 55 Dfs Mangle Dune deposits

II8A 30 1 55 Dfs Mangle Dune deposits Mona Island

II8D 52 5 2 52 55 48 55 Dfs Mangle Dune deposits

Nitrogen (%) II1A 3.43 0.67 5 3.89 3.95 2.65 55 Dfs Mangle Dune deposits

II8A 3.95 0.77 3 4.39 4.40 3.06 55 Dfs Mangle Dune deposits

II8A 0.60 1 55 Dfs Mangle Dune deposits Mona Island

II8D 8.80 2.28 2 8.88 10.49 7.26 55 Dfs Mangle Dune deposits

Phosphorus (mg/g) II1A 4.58 0.94 5 4.92 5.20 2.93 55 Dfs Mangle Dune deposits

II8A 10.61 4.01 3 12.11 13.65 6.07 55 Dfs Mangle Dune deposits

II8A 0.19 1 55 Dfs Mangle Dune deposits Mona Island

II8D 53.79 20.27 2 53.79 68.12 39.46 55 Dfs Mangle Dune deposits

Potassium (mg/g) II1A 12.39 0.05 5 12.42 12.44 12.33 55 Dfs Mangle Dune deposits

II8A 6.74 1.81 3 7.61 7.93 4.70 55 Dfs Mangle Dune deposits

II8A 0.97 1 55 Dfs Mangle Dune deposits Mona Island

II8D 9.14 1.92 2 9.14 10.49 7.78 55 Dfs Mangle Dune deposits

Sodium (mg/g) II1A 1.96 0.19 5 1.87 2.21 1.75 55 Dfs Mangle Dune deposits

II8A 2.27 0.71 3 2.32 2.96 1.54 55 Dfs Mangle Dune deposits
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

109

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8A 1.32 1 55 Dfs Mangle Dune deposits Mona Island

II8D 5.16 1.82 2 5.16 6.44 3.87 55 Dfs Mangle Dune deposits

Wood density (g/cc) II1B 0.42 0.01 3 0.42 0.43 0.41 97 Wfs Inceptisols Tuffac. sandstone

Forest Floor-Logs Calcium (mg/g) II6 4.74 3.74 2 4.74 7.38 2.09 95 Wfs Ultisols Tuffac. sandstone Class 3

II6 6.65 3.96 3 5.52 10.81 3.06 95 Wfs Ultisols Tuffac. sandstone Class 3--4

II6 6.47 3.96 3 5.52 10.81 3.06 95 Wfs Ultisols Tuffac. sandstone Class 4

II6 6.48 2.64 6 6.19 9.41 3.93 95 Wfs Ultisols Tuffac. sandstone Class 4--5

II6 7.87 4.55 17 6.50 17.07 2.55 95 Wfs Ultisols Tuffac. sandstone Class 5

Magnesium (mg/g) II6 3.78 1.80 2 3.78 5.06 2.51 95 Wfs Ultisols Tuffac. sandstone Class 3

II6 2.56 0.14 3 2.64 2.64 2.41 95 Wfs Ultisols Tuffac. sandstone Class 3--4

II6 2.56 0.14 3 2.64 2.64 2.41 95 Wfs Ultisols Tuffac. sandstone Class 4

II6 1.84 1.38 5 1.37 4.20 0.64 95 Wfs Ultisols Tuffac. sandstone Class 4--5

II6 3.09 2.02 17 2.56 8.25 0.78 95 Wfs Ultisols Tuffac. sandstone Class 5

Nitrogen (%) II6 1.26 0.37 2 1.26 1.52 1.00 95 Wfs Ultisols Tuffac. sandstone Class 3

II6 1.10 0.30 3 1.16 1.36 0.78 95 Wfs Ultisols Tuffac. sandstone Class 3--4

II6 1.10 0.30 3 1.16 1.36 0.78 95 Wfs Ultisols Tuffac. sandstone Class 4

II6 1.16 0.22 5 1.15 1.43 0.94 95 Wfs Ultisols Tuffac. sandstone Class 4--5

II6 1.19 0.34 17 1.15 1.65 0.80 95 Wfs Ultisols Tuffac. sandstone Class 5

Phosphorus (mg/g) II6 0.81 0.48 2 0.81 1.15 0.47 95 Wfs Ultisols Tuffac. sandstone Class 3

II6 0.67 0.35 3 0.50 1.06 0.43 95 Wfs Ultisols Tuffac. sandstone Class 3--4

II6 0.67 0.35 3 0.50 1.06 0.43 95 Wfs Ultisols Tuffac. sandstone Class 4

II6 0.44 0.11 5 0.43 0.57 0.33 95 Wfs Ultisols Tuffac. sandstone Class 4--5

II6 0.40 0.13 17 0.47 0.56 0.22 95 Wfs Ultisols Tuffac. sandstone Class 5

Potassium (mg/g) II6 6.62 6.18 2 6.62 10.99 2.25 95 Wfs Ultisols Tuffac. sandstone Class 3

II6 7.14 6.12 3 3.70 14.21 3.51 95 Wfs Ultisols Tuffac. sandstone Class 3--4

II6 7.14 6.12 3 3.70 14.21 3.51 95 Wfs Ultisols Tuffac. sandstone Class 4

II6 8.79 6.33 6 6.90 20.68 2.95 95 Wfs Ultisols Tuffac. sandstone Class 4--5

II6 5.03 1.39 17 5.32 6.71 2.98 95 Wfs Ultisols Tuffac. sandstone Class 5
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

110

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Forest Floor-Roots Calcium (mg/g) II6 4.66 3.14 14 2.93 11.25 2.20 127 Wfs Ultisols Tuffac. sandstone Dead roots

II6 3.20 0.76 20 3.25 4.50 1.67 127 Wfs Ultisols Tuffac. sandstone roots <2mm

II6 5.11 2.78 18 4.54 9.86 1.28 127 Wfs Ultisols Tuffac. sandstone roots >=5mm

II6 4.97 2.47 11 5.33 10.24 0.84 127 Wfs Ultisols Tuffac. sandstone >=2mm<5mm

Magnesium (mg/g) II6 2.06 0.76 14 0.18 3.93 1.06 127 Wfs Ultisols Tuffac. sandstone Dead roots

II6 2.36 1.01 20 2.30 4.82 1.01 127 Wfs Ultisols Tuffac. sandstone roots <2mm

II6 2.06 1.95 18 1.61 7.20 0.34 127 Wfs Ultisols Tuffac. sandstone roots >=5mm

II6 2.92 1.60 11 2.31 6.34 0.79 127 Wfs Ultisols Tuffac. sandstone >=2mm<5mm

Nitrogen (%) II6 0.78 0.35 11 0.68 1.60 0.37 127 Wfs Ultisols Tuffac. sandstone Dead roots

II6 1.05 0.37 10 1.01 1.83 0.52 127 Wfs Ultisols Tuffac. sandstone roots <2mm

II6 0.43 0.23 10 0.40 1.03 0.16 127 Wfs Ultisols Tuffac. sandstone roots >=5mm

II6 0.67 0.29 8 0.63 1.26 0.27 127 Wfs Ultisols Tuffac. sandstone >=2mm<5mm

Phosphorus (mg/g) II6 0.33 0.11 14 0.29 0.59 0.19 127 Wfs Ultisols Tuffac. sandstone Dead roots

II6 0.48 0.13 20 0.46 0.80 0.28 127 Wfs Ultisols Tuffac. sandstone Roots <2mm

II6 0.22 0.07 18 0.21 0.36 0.10 127 Wfs Ultisols Tuffac. sandstone Roots >=5mm

II6 0.30 0.13 11 0.26 0.52 0.15 127 Wfs Ultisols Tuffac. sandstone >=2mm<5mm

Potassium (mg/g) II6 1.91 1.18 14 1.70 4.72 0.77 127 Wfs Ultisols Tuffac. sandstone Dead roots

II6 1.83 0.79 20 1.38 3.19 0.76 127 Wfs Ultisols Tuffac. sandstone Roots <2mm

II6 2.32 0.91 18 2.33 3.97 0.65 127 Wfs Ultisols Tuffac. sandstone Roots >=5mm

II6 2.17 1.08 11 1.70 4.61 1.29 127 Wfs Ultisols Tuffac. sandstone >=2mm<5mm

Genipa americana Wood density (g/cc) II1B 0.59 0.02 3 0.59 0.61 0.58 97 Wfs Inceptisols Tuffac. sandstone

Gleichenia spp. Aluminum (mg/g) II6 6.11 1 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II6 1.07 1 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II6 0.33 1 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II6 1.28 1 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/kg) II6 68 1 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

111

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Nitrogen (%) II6 1.01 1 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II6 0.43 1 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II6 7.87 1 135 Lmrf Ultisols Tuffac. sandstone

Gomidesia lindeniana Aluminum (μg/g) II1A 36 1 58 Lmrf Ultisols Serpentinite

Calcium (mg/g) II1A 7.22 1 58 Lmrf Ultisols Serpentinite

Cobalt (μg/g) II1A 0.84 1 58 Lmrf Ultisols Serpentinite

Copper (μg/g) II1A 7.55 1 58 Lmrf Ultisols Serpentinite

Chromium (μg/g) II1A 4.18 1 58 Lmrf Ultisols Serpentinite

Iron (μg/g) II1A 82.75 1 58 Lmrf Ultisols Serpentinite

Magnesium (mg/g) II1A 7.42 1 58 Lmrf Ultisols Serpentinite

Manganese (μg/g) II1A 13.70 1 58 Lmrf Ultisols Serpentinite

Niquel (μg/g) II1A 181 1 58 Lmrf Ultisols Serpentinite

Nitrogen (%) II1A 0.76 1 58 Lmrf Ultisols Serpentinite

Phosphorus (mg/g) II1A 7.88 1 58 Lmrf Ultisols Serpentinite

Potassium (mg/g) II1A 9.96 1 58 Lmrf Ultisols Serpentinite

Gonzalagunia spicata Aluminum (mg/g) II2 0.19 1 135 Lmrf Ultisols Tuffac. sandstone

II8A 0.57 0.17 2 0.57 0.86 0.27 146 Swf Ultisols Tuffac. sandstone

II8B 0.17 0.15 2 0.17 0.27 0.06 146 Swf Ultisols Tuffac. sandstone

Ash (%) II8A 6.62 0.98 2 6.62 7.31 5.93 146 Swf Ultisols Tuffac. sandstone

II8B 1.98 1.57 2 1.98 3.09 0.87 146 Swf Ultisols Tuffac. sandstone

C/N II8A 30 12 2 30 38 21 146 Swf Ultisols Tuffac. sandstone

II8B 94 9 2 94 100 87 146 Swf Ultisols Tuffac. sandstone

Calcium (mg/g) II2 6.20 1 135 Lmrf Ultisols Tuffac. sandstone

II8A 10.24 3.32 2 10.24 12.59 7.89 146 Swf Ultisols Tuffac. sandstone

II8B 5.94 4.91 2 5.94 9.41 2.47 146 Swf Ultisols Tuffac. sandstone

Carbon (%) II8A 53 2 2 53 55 51 146 Swf Ultisols Tuffac. sandstone

 II8B 54 0 2 54 54 51 146 Swf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

112

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Iron (mg/g) II2 0.14 1 135 Lmrf Ultisols Tuffac. sandstone

II8A 0.37 0.32 2 0.37 0.60 0.14 146 Swf Ultisols Tuffac. sandstone

II8B 0.09 0.07 2 0.09 0.14 0.03 146 Swf Ultisols Tuffac. sandstone

Magnesium (mg/g) II2 3.30 1 135 Lmrf Ultisols Tuffac. sandstone

II8A 3.34 0.83 2 3.34 3.93 2.76 146 Swf Ultisols Tuffac. sandstone

II8B 0.83 0.42 2 0.83 1.13 0.54 146 Swf Ultisols Tuffac. sandstone

Manganese (mg/g) II2 0.10 1 135 Lmrf Ultisols Tuffac. sandstone

II8A 0.21 0.09 2 0.21 0.27 0.14 146 Swf Ultisols Tuffac. sandstone

II8B 0.07 0.02 2 0.07 0.08 0.05 146 Swf Ultisols Tuffac. sandstone

Nitrogen (%) II2 1.14 1 135 Lmrf Ultisols Tuffac. sandstone

II8A 1.96 0.88 2 1.96 2.58 1.34 146 Swf Ultisols Tuffac. sandstone

II8B 0.58 0.06 2 0.58 0.62 0.54 146 Swf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II2 0.72 1 135 Lmrf Ultisols Tuffac. sandstone

II8A 0.56 0.06 2 0.56 0.60 0.52 146 Swf Ultisols Tuffac. sandstone

II8B 0.23 0.08 2 0.23 0.29 0.18 146 Swf Ultisols Tuffac. sandstone

Potassium (mg/g) II2 16.80 1 135 Lmrf Ultisols Tuffac. sandstone

II8A 6.94 7.39 2 6.94 12.16 1.72 146 Swf Ultisols Tuffac. sandstone

II8B 3.08 3.20 2 3.08 5.34 0.82 146 Swf Ultisols Tuffac. sandstone

Guaiacum officinale Calcium (mg/g) II1Ci 26.74 1 104 Sdf Mollisols Alluvial deposits

II1Ci 26.74 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Magnesium (mg/g) II1Ci 1.73 1 104 Sdf Mollisols Alluvial deposits

II1Ci 1.73 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Nitrogen (%) II1A 1.81 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 2.17 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Phosphorus (mg/g) II1A 0.44 1 104 Sdf Mollisols Alluvial deposits

II1A 0.44 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 0.13 1 104 Sdf Mollisols Alluvial deposits

II1Ci 0.13 1 149 Sdf Mollisols Alluvial deposits Guanica Forest
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

113

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Potassium (mg/g) II1A 12.85 1 104 Sdf Mollisols Alluvial deposits

II1A 12.85 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 8.23 1 104 Sdf Mollisols Alluvial deposits

II1Ci 8.23 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Guaiacum sanctum Calcium (mg/g) II1A 17.37 1 104 Sdf Mollisols Alluvial deposits

II1A 17.37 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 63.48 1 104 Sdf Mollisols Alluvial deposits

II1Ci 63.48 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Magnesium (mg/g) II1A 2.41 1 104 Sdf Mollisols Alluvial deposits

II1A 2.41 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 3.30 1 104 Sdf Mollisols Alluvial deposits

II1Ci 3.30 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Nitrogen (%) II1A 1.72 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 1.18 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Phosphorus (mg/g) II1A 0.41 1 104 Sdf Mollisols Alluvial deposits

II1A 0.41 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 0.19 1 104 Sdf Mollisols Alluvial deposits

II1Ci 0.19 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Potassium (mg/g) II1A 14.87 1 104 Sdf Mollisols Alluvial deposits

II1A 14.87 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 14.62 1 104 Sdf Mollisols Alluvial deposits

II1Ci 14.62 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Guanica Forest Aluminum (mg/g) II1A 0.10 0.05 60 0.10 0.24 0.02 134 Sdf Mollisols Alluvial deposits

II1B 0.14 0.08 55 0.14 0.50 0.03 134 Sdf Mollisols Alluvial deposits

II1C 0.03 0.02 18 0.02 0.07 0.01 134 Sdf Mollisols Alluvial deposits

II1D 0.17 0.14 51 0.13 0.80 0.01 134 Sdf Mollisols Alluvial deposits

II1G 1.75 2.41 3 0.70 4.51 0.04 134 Sdf Mollisols Alluvial deposits Epiphyte
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

114

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1G 0.13 0.25 22 0.07 1.26 0.04 134 Sdf Mollisols Alluvial deposits Leaves + branches

II1G 7.89 3.94 6 6.98 13.90 2.55 134 Sdf Mollisols Alluvial deposits

II1G 0.06 0.02 10 0.05 0.10 0.04 134 Sdf Mollisols Alluvial deposits Vines

II8A 1.18 0.98 12 0.97 3.37 0.25 134 Sdf Mollisols Alluvial deposits

II8B 0.63 0.48 11 0.47 1.56 0.20 134 Sdf Mollisols Alluvial deposits

II8C 1.93 2.31 5 0.92 5.94 0.34 134 Sdf Mollisols Alluvial deposits

II8D 9.35 9.25 10 7.31 33.22 1.30 134 Sdf Mollisols Alluvial deposits

Ash (%) II1A 9.74 0.82 59 9.85 11.57 7.80 134 Sdf Mollisols Alluvial deposits

II1B 7.35 1.45 46 7.45 10.65 3.52 134 Sdf Mollisols Alluvial deposits

II1C 4.34 1.85 18 4.49 7.95 1.34 134 Sdf Mollisols Alluvial deposits

II1D 4.17 0.75 34 4.18 6.17 2.28 134 Sdf Mollisols Alluvial deposits

II1G 5.48 2.83 3 3.85 8.75 3.83 134 Sdf Mollisols Alluvial deposits Epiphyte

II1G 9.29 1.75 22 8.93 14.54 5.74 134 Sdf Mollisols Alluvial deposits Leaves + branches

II1G 22.07 6.93 6 19.48 32.71 14.04 134 Sdf Mollisols Alluvial deposits

II1G 8.19 3.43 8 7.34 15.82 4.19 134 Sdf Mollisols Alluvial deposits Vines

II8A 12.97 3.21 12 12.96 18.93 9.64 134 Sdf Mollisols Alluvial deposits

II8B 9.82 4.87 11 9.14 17.01 3.64 134 Sdf Mollisols Alluvial deposits

II8C 6.17 1 134 Sdf Mollisols Alluvial deposits

II8D 20.75 6.76 6 23.22 26.56 10.96 134 Sdf Mollisols Alluvial deposits

II9A 9.83 1.54 64 10.17 12.69 4.08 109 Sdf Mollisols Alluvial deposits Guanica

II9B 6.83 2.71 56 6.05 14.21 1.24 109 Sdf Mollisols Alluvial deposits Guanica

II9C 5.70 1.89 39 5.26 10.00 2.41 109 Sdf Mollisols Alluvial deposits Guanica

II9D 6.00 2.13 28 5.98 9.82 1.04 109 Sdf Mollisols Alluvial deposits Guanica

C/N II1A 53 8 60 53 69 31 134 Sdf Mollisols Alluvial deposits

II1B 56 9 55 55 86 43 134 Sdf Mollisols Alluvial deposits

II1C 114 39 18 106 189 62 134 Sdf Mollisols Alluvial deposits

II1D 41 12 50 41 64 20 134 Sdf Mollisols Alluvial deposits

II1G 69 27 4 76 92 34 134 Sdf Mollisols Alluvial deposits Epiphyte

II1G 36 7 22 35 49 25 134 Sdf Mollisols Alluvial deposits Leaves + branches
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

115

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1G 22 1 6 2 24 20 134 Sdf Mollisols Alluvial deposits

II1G 42 18 10 37 77 17 134 Sdf Mollisols Alluvial deposits Vines

II8A 37 8 12 37 50 23 134 Sdf Mollisols Alluvial deposits

II8B 61 25 11 53 97 38 134 Sdf Mollisols Alluvial deposits

II8C 39 6 5 41 46 29 134 Sdf Mollisols Alluvial deposits

II8D 28 11 9 25 52 17 134 Sdf Mollisols Alluvial deposits

II9A 36 12 68 35 65 17 109 Sdf Mollisols Alluvial deposits Guanica

II9B 49 16 60 49 110 25 109 Sdf Mollisols Alluvial deposits Guanica

II9C 28 15 53 23 85 11 109 Sdf Mollisols Alluvial deposits Guanica

II9D 50 59 34 31 357 17 109 Sdf Mollisols Alluvial deposits Guanica

Calcium (mg/g) II1A 29.20 4.43 60 29.32 42.30 18.16 134 Sdf Mollisols Alluvial deposits

II1B 25.46 7.10 55 26.13 45.04 3.98 134 Sdf Mollisols Alluvial deposits

II1C 14.40 6.17 18 15.57 24.05 4.14 134 Sdf Mollisols Alluvial deposits

II1D 9.97 4.35 51 9.08 33.47 4.64 134 Sdf Mollisols Alluvial deposits

II1G 12.06 5.65 3 11.08 18.13 6.96 134 Sdf Mollisols Alluvial deposits Epiphyte

II1G 27.31 8.07 22 25.58 56.97 18.47 134 Sdf Mollisols Alluvial deposits Leaves + branches

II1G 44.95 13.82 6 41.53 63.56 27.25 134 Sdf Mollisols Alluvial deposits

II1G 23.00 5.27 10 25.97 28.13 16.01 134 Sdf Mollisols Alluvial deposits Vines

II8A 43.13 10.32 12 41.40 69.13 32.65 134 Sdf Mollisols Alluvial deposits

II8B 24.09 16.50 11 36.32 57.89 12.99 134 Sdf Mollisols Alluvial deposits

II8C 19.70 8.66 5 16.63 30.63 9.60 134 Sdf Mollisols Alluvial deposits

II8D 44.05 13.37 10 40.88 61.78 20.42 134 Sdf Mollisols Alluvial deposits

Carbon (%) II1A 51 1 60 51 53 49 134 Sdf Mollisols Alluvial deposits

II1B 50 1 55 50 54 48 134 Sdf Mollisols Alluvial deposits

II1C 52 1 18 52 54 50 134 Sdf Mollisols Alluvial deposits

II1D 53 1 50 52 56 50 134 Sdf Mollisols Alluvial deposits

II1G 52 1 4 52 52 50 134 Sdf Mollisols Alluvial deposits Epiphyte

II1G 52 1 22 52 54 50 134 Sdf Mollisols Alluvial deposits Leaves + branches

II1G 46 4 6 47 52 39 134 Sdf Mollisols Alluvial deposits
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

116

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1G 52 2 10 51 54 47 134 Sdf Mollisols Alluvial deposits Vines

II8A 51 1 12 51 42 48 134 Sdf Mollisols Alluvial deposits

II8B 51 2 11 51 53 48 134 Sdf Mollisols Alluvial deposits

II8C 52 2 5 53 55 49 134 Sdf Mollisols Alluvial deposits

II8D 45 6 9 46 52 32 134 Sdf Mollisols Alluvial deposits

II9A 50 1 68 51 52 44 109 Sdf Mollisols Alluvial deposits Guanica

II9B 50 1 60 51 52 46 109 Sdf Mollisols Alluvial deposits Guanica

II9C 52 2 53 52 56 48 109 Sdf Mollisols Alluvial deposits Guanica

II9D 50 2 34 51 55 46 109 Sdf Mollisols Alluvial deposits Guanica

Iron (mg/g) II1A 0.11 0.15 60 0.06 1.08 0.01 134 Sdf Mollisols Alluvial deposits

II1B 0.13 0.14 55 0.10 1.04 0.04 134 Sdf Mollisols Alluvial deposits

II1C 0.05 0.03 18 0.05 0.10 0.01 134 Sdf Mollisols Alluvial deposits

II1D 0.16 0.16 51 0.11 0.89 0.02 134 Sdf Mollisols Alluvial deposits

II1G 1.56 1.98 3 0.78 3.80 0.09 134 Sdf Mollisols Alluvial deposits Epiphyte

II1G 0.11 0.18 22 0.08 0.90 0.03 134 Sdf Mollisols Alluvial deposits Leaves + branches

II1G 7.39 3.88 6 4.00 13.45 2.02 134 Sdf Mollisols Alluvial deposits

II1G 0.09 0.04 10 0.09 0.14 0.04 134 Sdf Mollisols Alluvial deposits Vines

II8A 0.98 0.82 12 0.88 2.89 0.18 134 Sdf Mollisols Alluvial deposits

II8B 0.60 0.56 11 0.35 1.85 0.15 134 Sdf Mollisols Alluvial deposits

II8C 1.60 1.75 5 0.71 4.49 0.26 134 Sdf Mollisols Alluvial deposits

II8D 8.78 8.37 10 7.10 30.17 1.01 134 Sdf Mollisols Alluvial deposits

Magnessium (mg/g) II1A 3.31 0.89 60 3.15 7.04 1.63 134 Sdf Mollisols Alluvial deposits

II1B 0.86 0.28 55 0.84 1.46 0.43 134 Sdf Mollisols Alluvial deposits

II1C 0.56 0.21 18 0.55 1.17 0.32 134 Sdf Mollisols Alluvial deposits

II1D 1.51 0.69 51 1.37 3.61 0.43 134 Sdf Mollisols Alluvial deposits

II1G 1.85 0.94 3 1.51 2.91 1.13 134 Sdf Mollisols Alluvial deposits Epiphyte

II1G 2.54 0.97 22 2.42 3.87 0.97 134 Sdf Mollisols Alluvial deposits Leaves + branches

II1G 2.84 0.62 6 2.86 3.44 1.77 134 Sdf Mollisols Alluvial deposits

II1G 2.06 2.49 10 1.11 8.62 0.41 134 Sdf Mollisols Alluvial deposits Vines
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

117

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8A 1.79 0.65 12 1.72 2.96 0.75 134 Sdf Mollisols Alluvial deposits

II8B 0.92 0.36 11 0.97 1.37 0.32 134 Sdf Mollisols Alluvial deposits

II8C 1.75 0.56 5 1.78 2.45 0.88 134 Sdf Mollisols Alluvial deposits

II8D 2.68 1.22 10 2.40 5.22 1.18 134 Sdf Mollisols Alluvial deposits

Manganese (mg/kg) II1A 40 10 60 39 64 25 134 Sdf Mollisols Alluvial deposits

II1B 18 9 53 17 54 1 134 Sdf Mollisols Alluvial deposits

II1C 11 7 18 10 24 1 134 Sdf Mollisols Alluvial deposits

II1D 20 7 51 21 29 3 134 Sdf Mollisols Alluvial deposits

II1G 49 34 3 30 89 29 134 Sdf Mollisols Alluvial deposits Epiphyte

II1G 41 15 22 45 63 13 134 Sdf Mollisols Alluvial deposits Leaves + branches

II1G 234 78 6 233 326 122 134 Sdf Mollisols Alluvial deposits

II1G 40 21 10 37 69 14 134 Sdf Mollisols Alluvial deposits Vines

II8A 78 40 12 65 171 36 134 Sdf Mollisols Alluvial deposits

II8B 39 22 11 39 83 12 134 Sdf Mollisols Alluvial deposits

II8C 63 55 5 37 160 23 134 Sdf Mollisols Alluvial deposits

II8D 242 180 10 238 638 43 134 Sdf Mollisols Alluvial deposits

Nitrogen (%) II1A 0.99 0.19 60 0.95 1.69 0.73 134 Sdf Mollisols Alluvial deposits

II1B 0.91 0.13 55 0.90 1.19 0.60 134 Sdf Mollisols Alluvial deposits

II1C 0.51 0.16 18 0.49 0.83 0.28 134 Sdf Mollisols Alluvial deposits

II1D 1.41 0.50 50 1.28 2.80 0.83 134 Sdf Mollisols Alluvial deposits

II1G 0.87 0.44 4 0.70 1.50 0.57 134 Sdf Mollisols Alluvial deposits Epiphyte

II1G 1.48 0.30 2 1.52 2.09 1.07 134 Sdf Mollisols Alluvial deposits Leaves + branches

II1G 2.07 0.24 6 2.12 2.34 1.63 134 Sdf Mollisols Alluvial deposits

II1G 1.47 0.64 10 1.44 2.80 0.66 134 Sdf Mollisols Alluvial deposits Vines

II8A 1.43 0.33 12 1.40 2.06 1.00 134 Sdf Mollisols Alluvial deposits

II8B 0.95 0.34 11 0.93 1.37 0.55 134 Sdf Mollisols Alluvial deposits

II8C 1.36 0.29 5 1.26 1.88 1.16 134 Sdf Mollisols Alluvial deposits

II8D 1.75 0.43 9 1.91 2.32 1.00 134 Sdf Mollisols Alluvial deposits

II9A 1.55 0.52 68 1.45 2.88 0.78 109 Sdf Mollisols Alluvial deposits Guanica
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

118

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II9B 1.14 0.35 60 1.03 1.92 0.47 109 Sdf Mollisols Alluvial deposits Guanica

II9C 2.33 1.00 53 2.23 4.87 0.59 109 Sdf Mollisols Alluvial deposits Guanica

II9D 1.54 0.71 34 1.60 2.94 0.14 109 Sdf Mollisols Alluvial deposits Guanica

Phosphorus (mg/g) II1A 0.19 0.05 60 0.18 0.40 0.12 134 Sdf Mollisols Alluvial deposits

II1B 0.20 0.07 55 0.20 0.52 0.10 134 Sdf Mollisols Alluvial deposits

II1C 0.15 0.04 18 0.15 0.24 0.08 134 Sdf Mollisols Alluvial deposits

II1D 0.54 0.24 51 0.51 1.31 0.10 134 Sdf Mollisols Alluvial deposits

II1G 0.40 0.19 3 0.32 0.61 0.26 134 Sdf Mollisols Alluvial deposits Epiphyte

II1G 0.41 0.10 22 0.40 0.63 0.24 134 Sdf Mollisols Alluvial deposits Leaves + branches

II1G 0.43 0.04 6 0.42 0.48 0.39 134 Sdf Mollisols Alluvial deposits

II1G 0.43 0.29 10 0.35 1.20 0.15 134 Sdf Mollisols Alluvial deposits Vines

II8A 0.26 0.06 12 0.24 0.37 0.17 134 Sdf Mollisols Alluvial deposits

II8B 0.17 0.07 11 0.17 0.27 0.08 134 Sdf Mollisols Alluvial deposits

II8C 0.33 0.04 5 0.32 0.40 0.30 134 Sdf Mollisols Alluvial deposits

II8D 0.42 0.13 10 0.38 0.65 0.23 134 Sdf Mollisols Alluvial deposits

Potassium (mg/g) II1A 5.98 1.91 60 5.84 10.25 2.35 134 Sdf Mollisols Alluvial deposits

II1B 2.81 1.16 55 2.68 5.05 0.66 134 Sdf Mollisols Alluvial deposits

II1C 3.84 0.86 18 3.96 5.80 2.55 134 Sdf Mollisols Alluvial deposits

II1D 5.06 1.92 51 4.79 10.93 1.62 134 Sdf Mollisols Alluvial deposits

II1G 8.71 7.32 3 5.30 17.11 3.72 134 Sdf Mollisols Alluvial deposits Epiphyte

II1G 10.78 3.20 22 10.24 17.66 3.46 134 Sdf Mollisols Alluvial deposits Leaves + branches

II1G 3.88 0.67 6 3.68 5.00 3.15 134 Sdf Mollisols Alluvial deposits

II1G 11.28 9.08 10 9.41 35.09 1.93 134 Sdf Mollisols Alluvial deposits Vines

II8A 3.42 0.55 12 3.40 4.19 2.60 134 Sdf Mollisols Alluvial deposits

II8B 2.63 0.86 11 2.60 3.80 1.34 134 Sdf Mollisols Alluvial deposits

II8C 2.60 0.72 5 2.79 3.58 1.73 134 Sdf Mollisols Alluvial deposits

II8D 0.23 1.35 10 4.04 7.05 2.20 134 Sdf Mollisols Alluvial deposits

Sulfur (%) II1A 0.34 0.05 60 0.33 0.45 0.23 134 Sdf Mollisols Alluvial deposits

II1B 0.28 0.05 55 0.28 0.42 0.19 134 Sdf Mollisols Alluvial deposits
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

119

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1C 0.13 0.04 18 0.13 0.19 0.08 134 Sdf Mollisols Alluvial deposits

II1D 0.28 0.05 50 0.27 0.43 0.21 134 Sdf Mollisols Alluvial deposits

II1G 0.31 0.04 4 0.31 0.34 0.27 134 Sdf Mollisols Alluvial deposits Epiphyte

II1G 0.33 0.08 22 0.31 0.54 0.22 134 Sdf Mollisols Alluvial deposits Leaves + branches

II1G 0.38 0.10 6 0.37 0.56 0.26 134 Sdf Mollisols Alluvial deposits

II1G 0.28 0.12 10 0.25 0.46 0.13 134 Sdf Mollisols Alluvial deposits Vines

II8A 0.33 0.04 12 0.33 0.40 0.26 134 Sdf Mollisols Alluvial deposits

II8B 0.24 0.06 11 0.24 0.36 0.15 134 Sdf Mollisols Alluvial deposits

II8C 0.27 0.06 5 0.24 0.38 0.22 134 Sdf Mollisols Alluvial deposits

II8D 0.36 0.04 9 0.38 0.41 0.29 134 Sdf Mollisols Alluvial deposits

Guapira obtusata Aluminum (mg/kg) II1A 19 1 102 Smf Limestone Tuffac. sandstone Adult 

Ash (%) II1A 4.94 1 102 Smf Limestone Tuffac. sandstone Adult 

Calcium (mg/g) II1A 3.39 1 102 Smf Limestone Tuffac. sandstone Adult 

Cobalt (μg/g) II1A 0.40 1 102 Smf Limestone Tuffac. sandstone Adult 

Iron (mg/kg) II1A 59 1 102 Smf Limestone Tuffac. sandstone Adult 

Magnesium (mg/g) II1A 6.25 1 102 Smf Limestone Tuffac. sandstone Adult 

Manganese (mg/kg) II1A 38 1 102 Smf Limestone Tuffac. sandstone Adult 

Nickel (μg/g) II1A 2.35 1 102 Smf Limestone Tuffac. sandstone Adult 

Nitrogen (%) II1A 2.67 1 102 Smf Limestone Tuffac. sandstone Adult 

Phosphorus (mg/g) II1A 1.46 1 102 Smf Limestone Tuffac. sandstone Adult 

Potassium (mg/g) II1A 13.69 1 102 Smf Limestone Tuffac. sandstone Adult 

Guararibaea turbinada Ash (%) II1B 2.14 1 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 1.20 1 68 Swf Ultisols Tuffac. sandstone Class 4 LD

II1B 1.08 1 68 Swf Ultisols Tuffac. sandstone Class 2/3 LD

Calcium (mg/g) II1B 5.81 1 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 1.88 1 68 Swf Ultisols Tuffac. sandstone Class 4 LD

II1B 3.58 1 68 Swf Ultisols Tuffac. sandstone Class 3 LD
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

120

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Nitrogen (%) II1B 0.34 1 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 0.30 1 68 Swf Ultisols Tuffac. sandstone Class 4 LD

II1B 0.16 1 68 Swf Ultisols Tuffac. sandstone Class 2/3 LD

Phosphorus (mg/g) II1B 0.18 1 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 0.06 1 68 Swf Ultisols Tuffac. sandstone Class 4 LD

II1B 0.03 1 68 Swf Ultisols Tuffac. sandstone Class 2/3 LD

Potassium (mg/g) II1B 0.92 1 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 0.32 1 68 Swf Ultisols Tuffac. sandstone Class 4 LD

II1B 0.25 1 68 Swf Ultisols Tuffac. sandstone Class 2/3 LD

Guarea glabra Aluminum (mg/g) II1B 0.10 1 93 Wflm Inceptisols Tuffac. sandstone

Ash (%) II1A 5.88 1 94 Swf Ultisols Tuffac. sandstone

II1A 7.28 0.09 2 7.28 7.34 7.21 101 Wfs Ultisols Lava

II1B 4.06 1 94 Swf Ultisols Tuffac. sandstone

II1B 6.43 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 4.50 0.26 3 4.40 4.80 4.30 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 1.57 1 68 Swf Ultisols Tuffac. sandstone Class 3 ST

II1C 2.14 1 94 Swf Ultisols Tuffac. sandstone

C/N II1B 44 1 93 Wflm Inceptisols Tuffac. sandstone

Calcium (mg/g) II1A 9.82 1 101 Wfs Ultisols Lava

II1B 21.06 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 3.21 0.36 3 3.12 3.61 2.91 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 4.26 1 68 Swf Ultisols Tuffac. sandstone Class 3 ST

Iron (mg/g) II1B 0.02 1 93 Wflm Inceptisols Tuffac. sandstone

Magnesium (mg/g) II1B 1.39 1 93 Wflm Inceptisols Tuffac. sandstone

Manganese (mg/g) II1B 0.04 1 93 Wflm Inceptisols Tuffac. sandstone

Nitrogen (%) II1A 2.84 1 94 Swf Ultisols Tuffac. sandstone

II1A 3.32 0.01 2 3.32 3.33 3.31 101 Wfs Ultisols Lava

II1B 1.23 1 94 Swf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

121

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1B 0.97 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.34 0.06 3 0.31 0.41 0.29 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 0.46 1 68 Swf Ultisols Tuffac. sandstone Class 3 ST

II1C 0.40 1 94 Swf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II1A 1.48 1 94 Swf Ultisols Tuffac. sandstone

II1A 2.06 0.36 2 2.06 2.31 1.80 101 Wfs Ultisols Lava

II1B 0.77 1 94 Swf Ultisols Tuffac. sandstone

II1B 0.65 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.15 0.06 3 0.13 0.22 0.10 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 0.19 1 68 Swf Ultisols Tuffac. sandstone Class 3 ST

II1C 0.29 1 94 Swf Ultisols Tuffac. sandstone

Potassium (mg/g) II1A 12.36 1 94 Swf Ultisols Tuffac. sandstone

II1A 18.07 0.10 2 18.07 18.14 18.00 101 Wfs Ultisols Lava

II1B 9.83 1 94 Swf Ultisols Tuffac. sandstone

II1B 8.43 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.43 0.15 3 0.44 0.58 0.28 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 0.76 1 68 Swf Ultisols Tuffac. sandstone Class 3 ST

II1C 3.39 1 94 Swf Ultisols Tuffac. sandstone

Guarea guidonia Aluminum (mg/g) II1A 0.43 1 129 Lmrf Ultisols Tuffac. sandstone

Ash (%) II1A 5.87 0.57 5 5.66 6.55 5.34 126 Ultisols Siltstone,sandstone Urban forest, RP

II1A 5.89 1 129 Lmrf Ultisols Tuffac. sandstone

II1C 1.62 0.39 5 1.55 2.14 1.12 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 7.20 1.94 6 6.77 10.09 5.50 126 Ultisols Siltstone,sandstone Urban forest, RP

II8A 8.40 1 71 Wfs Ultisols Tuffac. sandstone

C/N II1A 20 2 5 21 22 17 126 Ultisols Siltstone,sandstone Urban forest, RP

II1A 16 1 129 Lmrf Ultisols Tuffac. sandstone

II1C 213 70 5 197 314 127 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 42 2 6 43 43 38 126 Ultisols Siltstone,sandstone Urban forest, RP
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

122

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Calcium (mg/g) II1A 13.74 1 129 Lmrf Ultisols Tuffac. sandstone

II1A 5.20 0.90 14 5.29 6.49 3.82 135 Lmrf Ultisols Tuffac. sandstone

II1B 6.27 2.16 14 6.00 9.73 3.44 135 Lmrf Ultisols Tuffac. sandstone

II1C 1.28 0.48 27 1.06 2.37 0.87 135 Lmrf Ultisols Tuffac. sandstone

II1Ci 18.01 2.86 25 17.57 23.99 13.93 135 Lmrf Ultisols Tuffac. sandstone

II1F 2.85 0.35 12 2.79 3.53 2.42 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II1A 55 1 5 54 56 54 126 Ultisols Siltstone,sandstone Urban forest, RP

II1A 57 1 129 Lmrf Ultisols Tuffac. sandstone

II1Aiii 46 1 126 46 48 43 125 Wfs Ultisols Tuffac. sandstone Saplings

II1Aiii 45 1 95 45 48 41 125 Wfs Ultisols Tuffac. sandstone Seedlings

II1C 53 1 5 53 53 52 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 52 2 6 52 54 49 126 Ultisols Siltstone,sandstone Urban forest, RP

II8A 55 1 71 Wfs Ultisols Tuffac. sandstone

Iron (mg/g) II1A 0.43 1 129 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II1A 1.78 1 129 Lmrf Ultisols Tuffac. sandstone

II1A 3.93 0.74 14 4.00 4.92 2.75 135 Lmrf Ultisols Tuffac. sandstone

II1B 2.16 0.22 14 2.07 2.55 1.92 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.72 0.12 27 0.73 0.93 0.51 135 Lmrf Ultisols Tuffac. sandstone

II1Ci 2.87 0.83 25 2.62 5.06 1.76 135 Lmrf Ultisols Tuffac. sandstone

II1F 2.63 0.57 12 2.54 3.97 1.76 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/kg) II1A 52 1 129 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II1A 2.70 0.31 5 2.57 3.26 2.52 126 Ultisols Siltstone,sandstone Urban forest, RP

II1A 3.50 1 129 Lmrf Ultisols Tuffac. sandstone

II1A 2.68 0.22 14 2.63 3.04 2.41 135 Lmrf Ultisols Tuffac. sandstone

II1Aiii 3.19 0.89 126 3.13 5.08 1.63 125 Wfs Ultisols Tuffac. sandstone Saplings

II1Aiii 3.26 0.86 95 3.39 5.03 1.40 125 Wfs Ultisols Tuffac. sandstone Seedlings

II1B 0.89 0.25 14 0.83 1.10 0.60 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.27 0.09 5 0.27 0.41 0.17 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 0.20 0.05 27 0.20 0.33 0.10 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

123

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1Ci 1.23 0.04 6 1.24 1.28 1.17 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 1.10 0.11 25 1.11 1.38 0.90 135 Lmrf Ultisols Tuffac. sandstone

II1F 2.55 0.31 12 2.59 3.01 2.06 135 Lmrf Ultisols Tuffac. sandstone

II8A 1.83 1 71 Wfs Ultisols Tuffac. sandstone

Phosphorus (mg/g) II1A 1.00 1 129 Lmrf Ultisols Tuffac. sandstone

II1A 1.54 0.11 14 1.54 1.74 1.35 135 Lmrf Ultisols Tuffac. sandstone

II1B 0.45 0.13 14 0.42 0.68 0.30 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.18 0.07 27 0.16 0.31 0.09 135 Lmrf Ultisols Tuffac. sandstone

II1Ci 0.48 0.09 25 0.47 0.68 0.31 135 Lmrf Ultisols Tuffac. sandstone

II1F 1.45 0.15 12 1.44 1.72 1.22 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II1A 6.28 1 129 Lmrf Ultisols Tuffac. sandstone

II1A 11.87 1.17 14 11.64 13.88 10.06 135 Lmrf Ultisols Tuffac. sandstone

II1B 5.94 1.36 14 6.08 8.08 3.70 135 Lmrf Ultisols Tuffac. sandstone

II1C 1.60 0.79 27 1.48 2.94 0.30 135 Lmrf Ultisols Tuffac. sandstone

II1Ci 5.78 1.90 25 5.41 8.67 2.90 135 Lmrf Ultisols Tuffac. sandstone

II1F 13.71 3.44 12 14.65 15.17 2.87 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II1A 0.34 0.01 5 0.34 0.35 0.32 126 Ultisols Siltstone,sandstone Urban forest, RP

II1A 0.67 1 129 Lmrf Ultisols Tuffac. sandstone

II1C 0.10 0.04 5 0.08 0.14 0.05 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 0.44 0.05 6 0.45 0.49 0.33 126 Ultisols Siltstone,sandstone Urban forest, RP

II8A 0.43 1 71 Wfs Ultisols Tuffac. sandstone

Wood density (g/cc) II1C 0.52 0.09 5 0.54 0.60 0.38 126 Ultisols Siltstone,sandstone Urban forest, RP

Guarea ramiflora Carbon (%) II1Aiii 44 1 110 44 46 42 125 Wfs Ultisols Tuffac. sandstone

Nitrogen (%) II1Aiii 2.42 0.70 110 2.44 4.30 1.26 125 Wfs Ultisols Tuffac. sandstone

Guazuma ulmifolia Wood density (g/cc) II1B 0.52 0.03 4 0.52 0.55 0.49 97 Wfs Inceptisols Tuffac. sandstone

Guettarda pungens Aluminum (mg/kg) II1A 46 1 149 Sdf Mollisols Alluvial deposits Susua forest
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

124

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Ash (%) II1A 4.05 1 149 Sdf Mollisols Alluvial deposits Susua forest

Calcium (mg/g) II1A 9.96 1 149 Sdf Mollisols Alluvial deposits Susua forest

Iron (mg/g) II1A 0.12 1 149 Sdf Mollisols Alluvial deposits Susua forest

Magnesium (mg/g) II1A 1.30 1 149 Sdf Mollisols Alluvial deposits Susua forest

Manganese (mg/g) II1A 0.11 1 149 Sdf Mollisols Alluvial deposits Susua forest

Nitrogen (%) II1A 0.77 1 149 Sdf Mollisols Alluvial deposits Susua forest

Phosphorus (mg/g) II1A 0.30 1 149 Sdf Mollisols Alluvial deposits Susua forest

Potassium (mg/g) II1A 6.88 1 149 Sdf Mollisols Alluvial deposits Susua forest

Zinc (μg/g) II1A 40.09 1 149 Sdf Mollisols Alluvial deposits Susua forest

Guettarda scabra Aluminum (mg/kg) II1A 26 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 11 1 102 Smf Limestone Tuffac. sandstone New Adult 

Ash (%) II1A 3.42 1 102 Smf Limestone Tuffac. sandstone

II1A 6.28 1 149 Sdf Mollisols Alluvial deposits Susua forest

Calcium (mg/g) II1A 2.40 1 102 Smf Limestone Tuffac. sandstone New Adult 

II1A 4.77 1 149 Sdf Mollisols Alluvial deposits Susua forest

Cobalt (μg/g) II1A 0.10 1 102 Smf Limestone Tuffac. sandstone New Adult 

Iron (mg/kg) II1A 81 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 22 1 102 Smf Limestone Tuffac. sandstone New Adult 

Magnesium (mg/g) II1A 1.79 1 102 Smf Limestone Tuffac. sandstone New Adult 

II1A 4.92 1 149 Sdf Mollisols Alluvial deposits Susua forest

Manganese (mg/kg) II1A 26 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 12 1 102 Smf Limestone Tuffac. sandstone New Adult 

Nickel (μg/g) II1A 15 1 102 Smf Limestone Tuffac. sandstone New Adult 

Nitrogen (%) II1A 0.81 1 102 Smf Limestone Tuffac. sandstone New Adult 

II1A 1.17 1 149 Sdf Mollisols Alluvial deposits Susua forest

Phosphorus (mg/g) II1A 0.64 1 102 Smf Limestone Tuffac. sandstone New Adult 

II1A 0.41 1 149 Sdf Mollisols Alluvial deposits Susua forest

Potassium (mg/g) II1A 13.72 1 102 Smf Limestone Tuffac. sandstone New Adult 
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

125

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 3.47 1 149 Sdf Mollisols Alluvial deposits Susua forest

Zinc (μg/g) II1A 15.67 1 149 Sdf Mollisols Alluvial deposits Susua forest

Guzmania berteroniana Aluminum (mg/g) II6 0.61 1 135 Lmrf Ultisols Tuffac. sandstone

C/N II6 39 6 2 39 43 34 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II6 2.86 1 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II6 47 3 2 47 49 45 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II6 0.50 1 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II6 2.58 1 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II6 0.24 1 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II6 1.24 0.27 2 1.24 1.43 1.05 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II6 0.74 1 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II6 11.50 1 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II6 0.30 0.09 2 0.30 0.37 0.24 135 Lmrf Ultisols Tuffac. sandstone

Gymnanthes lucida Aluminum (mg/g) II3A 0.04 0.02 16 0.04 0.06 0.01 151 Sdf Mollisols Alluvial deposits

Ash (%) II3A 11.20 1.02 16 11.07 12.76 9.03 151 Sdf Mollisols Alluvial deposits

C/N II3A 27 1 16 26 28 24 151 Sdf Mollisols Alluvial deposits

Calcium (mg/g) II3A 27.32 3.91 16 27.03 32.83 16.02 151 Sdf Mollisols Alluvial deposits

II3Ci 28.53 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II3Ci 28.53 1 104 Sdf Mollisols Alluvial deposits

Carbon (%) II3A 48 1 16 48 50 47 151 Sdf Mollisols Alluvial deposits

Iron (mg/g) II3A 0.04 0.01 16 0.04 0.05 0.02 151 Sdf Mollisols Alluvial deposits

Magnesium (mg/g) II3A 2.94 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II3A 1.63 0.24 16 1.60 2.00 1.28 151 Sdf Mollisols Alluvial deposits

II3A 2.94 1 104 Sdf Mollisols Alluvial deposits

II3Ci 1.14 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II3Ci 1.14 1 104 Sdf Mollisols Alluvial deposits

Manganese (mg/kg) II3A 27 7 16 26 39 17 151 Sdf Mollisols Alluvial deposits
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

126

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Nitrogen (%) II3A 1.62 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II3A 1.82 0.10 16 1.82 2.04 1.67 151 Sdf Mollisols Alluvial deposits

II3Ci 1.06 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Phosphorus (mg/g) II3A 0.56 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II3A 0.46 0.03 16 0.45 0.53 0.42 151 Sdf Mollisols Alluvial deposits

II3A 0.56 1 104 Sdf Mollisols Alluvial deposits

II3Ci 0.24 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II3Ci 0.24 1 104 Sdf Mollisols Alluvial deposits

Potassium (mg/g) II3A 19.30 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II3A 7.04 0.53 16 7.04 8.40 6.16 151 Sdf Mollisols Alluvial deposits

II3Ci 10.22 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II3A 19.30 1 104 Sdf Mollisols Alluvial deposits

II3Ci 10.22 1 104 Sdf Mollisols Alluvial deposits

Sulfur (%) II3A 0.19 0.01 16 0.19 0.20 0.17 151 Sdf Mollisols Alluvial deposits

Heliconia caribaea Aluminum (mg/g) II3 0.20 0.00 2 0.20 0.20 0.20 135 Lmrf Ultisols Tuffac. sandstone

II3A 0.06 0.02 3 0.06 0.08 0.05 122 Wflm Inceptisols Tuffac. sandstone Palo Colorado

II3A 0.07 0.01 3 0.07 0.08 0.06 122 Lmrf Ultisols Tuffac. sandstone Tabonuco forest

II3A 0.05 0.02 4 0.05 0.07 0.03 122 Wfs Ultisols Tuffac. sandstone El Verde forest

II3F 0.12 0.12 13 0.07 0.38 0.02 129 Lmrf Ultisols Tuffac. sandstone Dwarf forest

II3F 0.12 0.12 13 0.07 0.38 0.02 137 Wfs Ultisols Tuffac. sandstone

II3G 4.78 4.06 13 2.80 10.90 0.35 122 Wfs Ultisols Tuffac. sandstone El Verde forest

II3G 1.80 1.97 5 1.47 5.16 0.20 122 Wflm Inceptisols Tuffac. sandstone Palo Colorado

II3G 2.42 2.06 3 1.73 6.68 0.26 122 Lmrf Ultisols Tuffac. sandstone Tabonuco forest

II3G 4.85 1.90 13 5.76 6.94 1.53 122 Smf Ultisols Tuffac. sandstone Dwarf forest

Ash (%) II3A 2.86 0.81 3 3.22 3.43 1.93 122 Wflm Inceptisols Tuffac. sandstone Palo Colorado

II3A 6.46 0.13 3 6.45 6.60 6.34 122 Lmrf Ultisols Tuffac. sandstone Tabonuco forest

II3A 5.99 0.07 4 6.01 6.04 5.90 122 Wfs Ultisols Tuffac. sandstone El Verde forest

II3F 15.30 1.19 13 15.21 17.03 13.53 129 Lmrf Ultisols Tuffac. sandstone Dwarf forest
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

127

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II3G 9.29 6.62 13 6.92 18.15 2.07 122 Wfs Ultisols Tuffac. sandstone El Verde forest

II3G 6.87 2.21 5 6.40 10.40 4.63 122 Wflm Inceptisols Tuffac. sandstone Palo Colorado

II3G 5.51 4.02 12 3.82 15.02 1.72 122 Lmrf Ultisols Tuffac. sandstone Tabonuco forest

II3G 8.59 2.58 13 8.39 14.02 5.11 122 Smf Ultisols Tuffac. sandstone Dwarf forest

II3F 15.30 1.19 13 15.21 17.03 13.53 137 Wfs Ultisols Tuffac. sandstone

C/N II3 23 3 3 21 26 20 135 Lmrf Ultisols Tuffac. sandstone

II3A 67 2 3 68 68 65 122 Wflm Inceptisols Tuffac. sandstone Palo Colorado

II3A 63 5 3 65 68 57 122 Lmrf Ultisols Tuffac. sandstone Tabonuco forest

II3A 67 5 4 66 73 62 122 Wfs Ultisols Tuffac. sandstone El verde Forest

II3F 20 4 13 20 27 14 129 Lmrf Ultisols Tuffac. sandstone Dwarf forest

II3G 36 13 13 36 62 14 122 Wfs Ultisols Tuffac. sandstone El Verde forest

II3G 40 14 5 31 57 29 122 Wflm Inceptisols Tuffac. sandstone Palo Colorado

II3G 41 12 13 45 59 24 122 Lmrf Ultisols Tuffac. sandstone Tabonuco forest

II3G 24 4 13 22 30 18 122 Smf Ultisols Tuffac. sandstone Dwarf forest

Calcium (mg/g) II3 2.81 0.08 2 2.81 2.86 2.75 135 Lmrf Ultisols Tuffac. sandstone

II3A 0.71 0.06 3 0.72 0.77 0.65 122 Wflm Inceptisols Tuffac. sandstone Palo Colorado

II3A 1.80 0.06 3 2.82 1.86 1.74 122 Lmrf Ultisols Tuffac. sandstone Tabonuco forest

II3A 1.88 0.14 4 1.88 2.04 1.72 122 Wfs Ultisols Tuffac. sandstone El Verde forest

II3F 10.23 2.51 13 9.79 15.38 6.79 129 Lmrf Ultisols Tuffac. sandstone Dwarf forest

II3F 10.23 2.51 13 9.79 15.38 6.79 137 Smf Ultisols Tuffac. sandstone

II3G 4.20 1.12 13 4.30 5.81 2.66 122 Wfs Ultisols Tuffac. sandstone El Verde forest

II3G 9.77 2.15 5 9.04 13.54 8.13 122 Wflm Inceptisols Tuffac. sandstone Palo Colorado

II3G 5.12 1.99 13 4.59 9.76 3.06 122 Lmrf Ultisols Tuffac. sandstone Tabonuco forest

II3G 7.93 2.17 13 8.19 10.93 5.19 122 Smf Ultisols Tuffac. sandstone Dwarf forest

Carbon (%) II3 46 0 3 46 46 46 135 Lmrf Ultisols Tuffac. sandstone

II3A 50 0 3 51 51 50 122 Wflm Inceptisols Tuffac. sandstone Palo Colorado

II3A 47 0 3 47 48 47 122 Lmrf Ultisols Tuffac. sandstone Tabonuco forest

II3A 48 0 4 48 48 48 122 Wfs Ultisols Tuffac. sandstone El verde Forest.

II3F 44 1 13 44 46 43 129 Smf Ultisols Tuffac. sandstone Dwarf forest
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

128

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II3F 44 1 13 44 47 46 137 Wfs Ultisols Tuffac. sandstone

II3G 49 5 13 51 55 43 122 Wfs Ultisols Tuffac. sandstone El Verde forest

II3G 52 2 5 52 55 49 122 Wflm Inceptisols Tuffac. sandstone Palo Colorado

II3G 56 5 13 56 63 48 122 Lmrf Ultisols Tuffac. sandstone Tabonuco forest

II3G 51 2 13 51 55 49 122 Smf Ultisols Tuffac. sandstone Dwarf forest

Iron (mg/g) II3 0.08 0.01 2 0.08 0.09 0.08 135 Lmrf Ultisols Tuffac. sandstone

II3A 0.01 0.01 3 0.01 0.03 0.01 122 Wflm Inceptisols Tuffac. sandstone Palo Colorado

II3A 0.06 0.00 3 0.06 0.06 0.06 122 Lmrf Ultisols Tuffac. sandstone Tabonuco forest

II3A 0.04 0.01 4 0.05 0.05 0.04 122 Wfs Ultisols Tuffac. sandstone El Verde Forest

II3F 0.16 0.11 13 0.11 0.33 0.05 129 Lmrf Ultisols Tuffac. sandstone Dwarf forest

II3F 0.16 0.11 13 0.11 0.33 0.05 137 Wfs Ultisols Tuffac. sandstone

II3G 2.73 1.99 13 2.13 5.49 0.39 122 Wfs Ultisols Tuffac. sandstone El Verde forest

II3G 0.21 0.03 5 0.20 0.25 0.18 122 Wflm Inceptisols Tuffac. sandstone Palo Colorado

II3G 1.11 1.14 13 0.68 3.95 0.16 122 Lmrf Ultisols Tuffac. sandstone Tabonuco forest

II3G 2.20 0.79 13 1.93 3.59 1.11 122 Smf Ultisols Tuffac. sandstone Dwarf forest

Magnesium (mg/g) II3 2.16 0.04 2 2.16 11.27 2.19 135 Lmrf Ultisols Tuffac. sandstone

II3A 1.01 0.08 3 0.99 1.09 0.94 122 Wflm Inceptisols Tuffac. sandstone Palo Colorado

II3A 2.41 0.04 3 2.40 2.46 2.38 122 Lmrf Ultisols Tuffac. sandstone Tabonuco forest

II3A 2.16 0.16 4 2.13 2.37 2.02 122 Wfs Ultisols Tuffac. sandstone El Verde Forest

II3F 4.86 0.53 13 4.84 5.73 3.88 129 Smf Ultisols Tuffac. sandstone Dwarf forest

II3F 4.86 0.53 13 4.84 5.73 3.88 137 Wfs Ultisols Tuffac. sandstone

II3G 1.75 0.65 13 1.38 2.78 1.06 122 Wfs Ultisols Tuffac. sandstone El Verde forest

II3G 1.95 0.53 5 1.70 2.80 1.54 122 Wflm Inceptisols Tuffac. sandstone Palo Colorado

II3G 1.11 0.41 13 1.08 1.81 0.20 122 Lmrf Ultisols Tuffac. sandstone Tabonuco forest

II3G 1.28 0.21 13 1.27 1.66 0.98 122 Smf Ultisols Tuffac. sandstone Dwarf forest

Manganese (mg/g) II3 0.88 0.02 2 0.88 0.89 0.86 135 Lmrf Ultisols Tuffac. sandstone

II3A 0.07 0.00 3 0.07 0.08 0.07 122 Wflm Inceptisols Tuffac. sandstone Palo Colorado

II3A 0.25 0.01 3 0.25 0.27 0.24 122 Lmrf Ultisols Tuffac. sandstone Tabonuco forest

II3A 0.09 0.01 4 0.09 0.10 0.09 122 Wfs Ultisols Tuffac. sandstone El Verde Forest
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

129

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II3F 1.01 0.61 13 0.95 2.48 0.36 129 Smf Ultisols Tuffac. sandstone Dwarf forest

II3F 1.01 0.61 13 0.95 2.48 0.36 137 Wfs Ultisols Tuffac. sandstone

II3G 0.05 0.01 13 0.50 0.07 0.03 122 Wfs Ultisols Tuffac. sandstone El Verde forest

II3G 0.40 0.15 5 0.37 0.66 0.27 122 Wflm Inceptisols Tuffac. sandstone Palo Colorado

II3G 0.39 0.20 13 0.41 0.70 0.07 122 Lmrf Ultisols Tuffac. sandstone Tabonuco forest

II3G 0.22 0.16 13 0.16 0.61 0.08 122 Smf Ultisols Tuffac. sandstone Dwarf forest

Nitrogen (%) II3 2.04 0.27 3 2.18 2.22 1.73 135 Lmrf Ultisols Tuffac. sandstone

II3A 0.75 0.02 3 0.74 0.78 0.74 122 Wflm Inceptisols Tuffac. sandstone Palo Colorado

II3A 0.75 0.07 3 0.73 0.83 0.70 122 Lmrf Ultisols Tuffac. sandstone Tabonuco forest

II3A 0.72 0.05 4 0.72 0.78 0.66 122 Wfs Ultisols Tuffac. sandstone El verde Forest

II3F 2.31 0.49 13 2.21 3.04 1.65 129 Smf Ultisols Tuffac. sandstone Dwarf forest

II3F 2.31 0.49 13 2.21 3.04 1.65 137 Wfs Ultisols Tuffac. sandstone

II3G 1.53 0.55 13 1.41 3.07 0.90 122 Wfs Ultisols Tuffac. sandstone El Verde forest

II3G 1.40 0.39 5 1.60 1.78 0.96 122 Wflm Inceptisols Tuffac. sandstone Palo Colorado

II3G 1.46 0.37 13 1.26 2.24 1.02 122 Lmrf Ultisols Tuffac. sandstone Tabonuco forest

II3G 2.21 0.33 13 2.30 2.94 1.70 122 Smf Ultisols Tuffac. sandstone Dwarf forest

Phosphorus (mg/g) II3 0.73 0.00 2 0.73 0.73 0.72 135 Lmrf Ultisols Tuffac. sandstone

II3A 0.19 0.02 3 0.18 0.20 0.17 122 Wflm Inceptisols Tuffac. sandstone Palo Colorado

II3A 0.35 0.10 3 0.35 0.36 0.35 122 Lmrf Ultisols Tuffac. sandstone Tabonuco forest

II3A 0.48 0.02 4 0.48 0.50 0.45 122 Wfs Ultisols Tuffac. sandstone El Verde Forest

II3F 2.22 0.66 13 2.19 3.32 1.29 129 Smf Ultisols Tuffac. sandstone Dwarf forest

II3F 2.22 0.66 13 2.19 3.32 1.29 137 Wfs Ultisols Tuffac. sandstone

II3G 0.22 0.08 13 0.20 0.34 0.11 122 Wfs Ultisols Tuffac. sandstone El Verde forest

II3G 0.24 0.05 5 0.22 0.31 0.19 122 Wflm Inceptisols Tuffac. sandstone Palo Colorado

II3G 0.26 0.09 13 0.24 0.43 0.14 122 Lmrf Ultisols Tuffac. sandstone Tabonuco forest

II3G 0.68 0.16 13 0.68 0.95 0.46 122 Smf Ultisols Tuffac. sandstone Dwarf forest

Potassium (mg/g) II3 11.35 0.12 2 11.35 11.44 11.27 135 Lmrf Ultisols Tuffac. sandstone

II3A 15.56 0.97 3 15.45 16.58 14.65 122 Wflm Inceptisols Tuffac. sandstone Palo Colorado

II3A 27.25 0.65 3 27.21 27.92 26.62 122 Lmrf Ultisols Tuffac. sandstone Tabonuco forest
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

130

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II3A 25.51 1.26 4 25.33 27.21 24.17 122 Wfs Ultisols Tuffac. sandstone El Verde Forest

II3F 34.34 5.39 13 36.27 42.52 24.28 129 Smf Ultisols Tuffac. sandstone Dwarf forest

II3F 34.34 5.39 13 36.27 42.52 24.28 137 Wfs Ultisols Tuffac. sandstone

II3G 1.13 0.50 13 0.86 1.94 0.64 122 Wfs Ultisols Tuffac. sandstone El Verde forest

II3G 1.86 0.64 5 2.20 2.26 0.76 122 Wflm Inceptisols Tuffac. sandstone Palo Colorado

II3G 0.93 0.28 13 0.93 1.33 0.52 122 Lmrf Ultisols Tuffac. sandstone Tabonuco forest

II3G 1.09 0.12 13 1.09 1.29 0.91 122 Smf Ultisols Tuffac. sandstone Dwarf forest

Sulfur (%) II3 0.33 0.09 3 0.29 0.43 0.26 135 Lmrf Ultisols Tuffac. sandstone

II3A 0.22 0.03 3 0.21 0.25 0.20 122 Wflm Inceptisols Tuffac. sandstone Palo Colorado

II3A 0.26 0.02 3 0.27 0.28 0.24 122 Lmrf Ultisols Tuffac. sandstone Tabonuco forest

II3A 0.24 0.03 4 0.24 0.27 0.21 122 Wfs Ultisols Tuffac. sandstone El Verde Forest

II3F 0.16 0.04 13 0.16 0.22 0.10 129 Smf Ultisols Tuffac. sandstone Dwarf forest

II3F 0.16 0.04 13 0.16 0.22 0.10 137 Wfs Ultisols Tuffac. sandstone

II3G 0.49 0.11 13 0.51 0.63 0.33 122 Wfs Ultisols Tuffac. sandstone El Verde forest

II3G 0.40 0.07 5 0.37 0.48 0.33 122 Wflm Inceptisols Tuffac. sandstone Palo Colorado

II3G 0.47 0.22 13 0.50 0.79 0.17 122 Lmrf Ultisols Tuffac. sandstone Tabonuco forest

II3G 0.74 0.12 13 0.77 0.89 0.50 122 Wfs Ultisols Tuffac. sandstone El Verde Forest

Henriettea squamulosum Aluminum (mg/g) II1A 7.74 1 93 Wflm Inceptisols Tuffac. sandstone

II1A 0.32 1 93 Wflm Inceptisols Tuffac. sandstone New

II1B 3.16 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.30 1 93 Wflm Inceptisols Tuffac. sandstone

Ash (%) II1A 5.95 1 93 Wflm Inceptisols Tuffac. sandstone New

II1A 4.83 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 3.64 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.94 1 93 Wflm Inceptisols Tuffac. sandstone

C/N II1A 29 1 93 Wflm Inceptisols Tuffac. sandstone New

II1A 70 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 53 1 93 Wflm Inceptisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

131

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1C 218 1 93 Wflm Inceptisols Tuffac. sandstone

Calcium (mg/g) II1A 14.18 1 93 Wflm Inceptisols Tuffac. sandstone New

II1A 11.92 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 10.96 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 2.66 1 93 Wflm Inceptisols Tuffac. sandstone

Iron (mg/g) II1A 0.21 1 93 Wflm Inceptisols Tuffac. sandstone New

II1A 0.17 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.12 1 93 Wflm Inceptisols Tuffac. sandstone

Magnesium (mg/g) II1A 3.57 1 93 Wflm Inceptisols Tuffac. sandstone New

II1A 1.17 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 1.94 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.10 1 93 Wflm Inceptisols Tuffac. sandstone

Manganese (mg/g) II1A 1.11 1 93 Wflm Inceptisols Tuffac. sandstone New

II1A 0.82 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.47 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.05 1 93 Wflm Inceptisols Tuffac. sandstone

Nitrogen (%) II1A 1.50 1 93 Wflm Inceptisols Tuffac. sandstone New

II1A 0.63 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.85 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.21 1 93 Wflm Inceptisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.54 1 93 Wflm Inceptisols Tuffac. sandstone New

II1A 0.47 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.45 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.09 1 93 Wflm Inceptisols Tuffac. sandstone

Potassium (mg/g) II1A 7.38 1 93 Wflm Inceptisols Tuffac. sandstone New

II1A 8.24 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 5.24 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 1.10 1 93 Wflm Inceptisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

132

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Herbaceous Aluminum (mg/g) II3 0.93 1.73 11 0.21 4.57 0.03 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II3 7.28 2.39 11 8.42 10.27 2.71 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II3 1.10 1.99 11 0.27 5.89 0.07 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II3 3.22 1.08 11 2.75 5.01 1.33 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II3 0.20 0.11 11 0.19 0.48 0.06 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II3 1.71 0.47 11 1.77 2.41 1.10 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II3 1.24 0.32 11 1.05 1.85 0.84 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II3 29.36 6.80 11 31.81 36.94 18.64 135 Lmrf Ultisols Tuffac. sandstone

Hernandia sonora Ash (%) II1A 14.58 3.96 3 13.88 18.85 11.02 90 Swf Ultisols Tuffac. sandstone Torn

II1A 11.60 4.27 3 13.82 14.30 6.67 90 Swf Ultisols Tuffac. sandstone Whole

II1C 26.64 7.55 3 30.11 31.82 17.98 90 Swf Ultisols Tuffac. sandstone

II8A 7.46 0.17 2 7.46 7.58 7.34 90 Swf Ultisols Tuffac. sandstone Recently fall

II8B 4.75 0.72 3 4.53 5.55 4.17 90 Swf Ultisols Tuffac. sandstone

II8C 6.50 2.55 3 7.45 8.44 3.61 90 Swf Ultisols Tuffac. sandstone

Calcium (mg/g) II1A 11.11 3.64 3 9.32 15.30 8.70 90 Swf Ultisols Tuffac. sandstone Torn

II1A 11.82 3.57 3 10.43 15.88 9.16 90 Swf Ultisols Tuffac. sandstone Whole

II1C 8.61 4.22 3 6.26 13.48 6.09 90 Swf Ultisols Tuffac. sandstone

II8A 8.35 1.32 3 9.02 9.21 6.83 90 Swf Ultisols Tuffac. sandstone Recently fall

II8B 6.20 3.62 3 4.31 10.38 3.92 90 Swf Ultisols Tuffac. sandstone

II8C 1.97 0.23 3 1.93 2.22 1.77 90 Swf Ultisols Tuffac. sandstone

II9A 9.16 0.96 22 8.95 11.82 7.91 139 Wfs Ultisols Tuffac. sandstone

II9A 6.95 2.19 22 6.38 15.15 3.94 139 Wfs Ultisols Tuffac. sandstone Other

II9B 4.15 2.14 7 3.63 8.45 2.00 139 Wfs Ultisols Tuffac. sandstone

II9C 1.37 0.96 6 1.18 3.08 0.41 139 Wfs Ultisols Tuffac. sandstone

II9D 7.05 3.58 5 5.61 11.77 3.70 139 Wfs Ultisols Tuffac. sandstone

Magnesium (mg/g) II1A 4.61 0.53 3 4.85 4.99 4.01 90 Swf Ultisols Tuffac. sandstone Torn

II1A 5.14 1.04 3 5.10 6.20 4.12 90 Swf Ultisols Tuffac. sandstone Whole

II1C 3.17 1.24 3 2.86 4.53 2.10 90 Swf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

133

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8A 6.26 0.91 3 6.51 7.02 5.24 90 Swf Ultisols Tuffac. sandstone Recently fall

II8B 2.19 0.70 3 2.08 2.93 1.55 90 Swf Ultisols Tuffac. sandstone

II8C 2.70 0.35 3 2.66 3.07 2.37 90 Swf Ultisols Tuffac. sandstone

II9A 5.13 0.65 22 4.92 6.63 3.93 139 Wfs Ultisols Tuffac. sandstone

II9A 4.32 1.61 22 4.36 7.99 2.03 139 Wfs Ultisols Tuffac. sandstone Other

II9B 2.26 1.18 7 1.86 4.38 0.59 139 Wfs Ultisols Tuffac. sandstone

II9C 2.46 0.80 6 2.17 3.62 1.68 139 Wfs Ultisols Tuffac. sandstone

II9D 3.77 1.12 5 4.31 4.70 1.96 139 Wfs Ultisols Tuffac. sandstone

Nitrogen (%) II1A 1.94 0.07 3 1.91 2.02 1.90 90 Swf Ultisols Tuffac. sandstone Torn

II1A 1.65 0.01 3 1.65 1.72 1.59 90 Swf Ultisols Tuffac. sandstone Whole

II1C 1.99 0.38 3 1.77 2.43 1.76 90 Swf Ultisols Tuffac. sandstone

II8A 1.09 0.09 3 1.06 1.19 1.01 90 Swf Ultisols Tuffac. sandstone Recently fall

II8B 1.03 0.46 3 0.84 1.56 0.70 90 Swf Ultisols Tuffac. sandstone

II8C 2.07 0.01 3 2.06 2.08 2.06 90 Swf Ultisols Tuffac. sandstone

II9A 1.65 0.28 22 1.58 2.56 1.29 139 Wfs Ultisols Tuffac. sandstone

II9A 1.55 0.65 22 1.39 2.83 0.56 139 Wfs Ultisols Tuffac. sandstone Other

II9B 0.80 0.28 12 0.73 1.46 0.40 139 Wfs Ultisols Tuffac. sandstone

II9C 2.31 0.91 11 2.17 4.19 1.24 139 Wfs Ultisols Tuffac. sandstone

II9D 2.44 0.71 9 2.35 3.42 1.55 139 Wfs Ultisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.85 0.06 3 0.87 0.90 0.78 90 Swf Ultisols Tuffac. sandstone Torn

II1A 0.64 0.08 3 0.69 0.69 0.54 90 Swf Ultisols Tuffac. sandstone Whole

II1C 0.90 0.10 3 0.89 1.00 0.80 90 Swf Ultisols Tuffac. sandstone

II8A 0.45 0.04 3 0.45 0.49 0.40 90 Swf Ultisols Tuffac. sandstone Recently fall

II8B 0.46 0.20 3 0.41 0.68 0.28 90 Swf Ultisols Tuffac. sandstone

II8C 2.18 0.49 3 1.94 2.74 1.86 90 Swf Ultisols Tuffac. sandstone

II9A 0.74 0.16 22 0.70 1.20 0.55 139 Wfs Ultisols Tuffac. sandstone

II9A 0.67 0.30 22 0.63 1.18 0.14 139 Wfs Ultisols Tuffac. sandstone Other

II9B 0.28 0.15 7 0.27 0.58 0.09 139 Wfs Ultisols Tuffac. sandstone

II9C 1.69 0.42 6 1.72 2.28 1.09 139 Wfs Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

134

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II9D 1.32 0.54 5 1.27 2.10 0.81 139 Wfs Ultisols Tuffac. sandstone

Potassium (mg/g) II1A 3.22 1.19 3 2.59 4.59 2.47 90 Swf Ultisols Tuffac. sandstone Torn

II1A 3.21 1.20 3 2.64 4.59 2.40 90 Swf Ultisols Tuffac. sandstone Whole

II1C 2.78 0.64 3 2.87 3.38 2.10 90 Swf Ultisols Tuffac. sandstone

II8A 3.76 3.75 3 1.85 8.08 1.34 90 Swf Ultisols Tuffac. sandstone Recently fall

II8B 1.92 0.36 3 2.08 2.18 1.50 90 Swf Ultisols Tuffac. sandstone

II8C 4.45 0.79 3 4.56 5.18 3.61 90 Swf Ultisols Tuffac. sandstone

II9A 4.69 1.06 22 4.45 7.01 2.87 139 Wfs Ultisols Tuffac. sandstone

II9A 4.68 2.24 22 4.10 12.04 1.68 139 Wfs Ultisols Tuffac. sandstone Other

II9B 2.02 1.30 7 1.90 4.30 0.61 139 Wfs Ultisols Tuffac. sandstone

II9C 10.42 3.74 3 9.62 16.57 5.70 139 Wfs Ultisols Tuffac. sandstone

II9D 8.09 4.70 5 7.53 15.78 3.74 139 Wfs Ultisols Tuffac. sandstone

Heteropteris laurifolia Aluminum (mg/g) II4 0.19 0.01 2 0.19 0.20 0.19 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II4 6.53 3.45 2 6.53 8.97 4.09 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II4 48 1 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II4 0.16 0.06 2 0.16 0.20 0.12 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II4 5.05 0.93 2 5.05 5.70 4.39 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II4 0.71 0.87 2 0.71 1.32 0.09 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II4 1.26 0.35 2 1.26 1.50 1.01 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II4 0.69 0.50 2 0.69 1.05 0.33 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II4 8.29 10.14 2 8.29 15.46 1.12 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II4 0.14 1 135 Lmrf Ultisols Tuffac. sandstone

Heterotrichum cymosum Aluminum (mg/g) II8A 10.97 3.83 3 8.79 15.39 8.73 146 Swf Ultisols Tuffac. sandstone

II8B 4.13 2.72 2 4.13 6.05 2.20 146 Swf Ultisols Tuffac. sandstone

Ash (%) II8A 10.48 2.38 3 9.48 13.19 8.76 146 Swf Ultisols Tuffac. sandstone

II8B 3.81 1.38 2 3.81 4.78 2.83 146 Swf Ultisols Tuffac. sandstone

C/N II8A 97 9 3 32 33 17 146 Swf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

135

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8B 95 49 2 95 129 60 146 Swf Ultisols Tuffac. sandstone

Calcium (mg/g) II8A 20.37 3.44 3 18.94 24.29 17.87 146 Swf Ultisols Tuffac. sandstone

II8B 6.72 2.88 2 6.72 8.75 4.68 146 Swf Ultisols Tuffac. sandstone

Carbon (%) II8A 50 1 3 50 50 49 146 Swf Ultisols Tuffac. sandstone

II8B 52 1 2 52 53 52 146 Swf Ultisols Tuffac. sandstone

Iron (mg/g) II8A 0.55 0.59 3 0.21 1.24 0.21 146 Swf Ultisols Tuffac. sandstone

II8B 1.09 1.49 2 1.09 2.15 0.04 146 Swf Ultisols Tuffac. sandstone

Magnesium (mg/g) II8A 2.92 0.10 3 2.88 3.03 2.85 146 Swf Ultisols Tuffac. sandstone

II8B 1.18 0.75 2 1.18 1.71 0.66 146 Swf Ultisols Tuffac. sandstone

Manganese (mg/g) II8A 0.51 0.21 3 0.40 0.75 0.38 146 Swf Ultisols Tuffac. sandstone

II8B 0.55 0.30 2 0.55 0.76 0.34 146 Swf Ultisols Tuffac. sandstone

Nitrogen (%) II8A 1.96 0.73 3 1.54 2.81 1.54 146 Swf Ultisols Tuffac. sandstone

II8B 0.64 0.32 2 0.64 0.86 0.41 146 Swf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II8A 0.74 0.30 3 0.57 1.09 0.57 146 Swf Ultisols Tuffac. sandstone

II8B 0.40 0.24 2 0.40 0.56 0.23 146 Swf Ultisols Tuffac. sandstone

Potassium (mg/g) II8A 7.34 3.97 3 9.51 9.74 2.76 146 Swf Ultisols Tuffac. sandstone

II8B 2.97 1.71 2 2.97 4.18 1.76 146 Swf Ultisols Tuffac. sandstone

Hibiscus elatus Ash (%) II8A 8.48 2.17 4 8.48 11.50 6.35 90 Swf Ultisols Tuffac. sandstone Recently fall

II8A 8.32 2.82 6 8.32 10.70 2.93 90 Swf Ultisols Tuffac. sandstone Torn

II8B 4.99 2.23 4 4.99 7.25 3.60 90 Swf Ultisols Tuffac. sandstone

II8C 9.24 1 90 Swf Ultisols Tuffac. sandstone

II8C 3.86 1.69 4 3.86 5.91 1.91 90 Swf Ultisols Tuffac. sandstone

II8D 13.57 5.41 4 13.57 20.12 7.19 90 Swf Ultisols Tuffac. sandstone

Calcium (mg/g) II8A 17.19 2.40 3 18.10 19.01 14.47 90 Swf Ultisols Tuffac. sandstone Recently fall

II8A 21.95 4.99 5 21.95 29.26 16.48 90 Swf Ultisols Tuffac. sandstone Torn

II8B 10.37 4.38 4 8.49 15.39 7.24 90 Swf Ultisols Tuffac. sandstone

II8C 18.75 1 90 Swf Ultisols Tuffac. sandstone

II8C 6.62 3.35 3 6.50 10.03 3.33 90 Swf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

136

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8D 17.91 5.96 3 17.97 23.84 11.93 90 Swf Ultisols Tuffac. sandstone

II9A 13.32 2.67 22 12.75 21.52 10.17 139 Wfs Ultisols Tuffac. sandstone

II9A 4.63 1.97 22 3.82 8.82 2.26 139 Wfs Ultisols Tuffac. sandstone Other

II9B 6.55 3.73 7 6.04 12.53 1.40 139 Wfs Ultisols Tuffac. sandstone

II9C 6.04 1.05 6 5.73 8.04 5.11 139 Wfs Ultisols Tuffac. sandstone

II9D 8.08 1.61 6 7.63 10.82 6.54 139 Wfs Ultisols Tuffac. sandstone

Magnesium (mg/g) II8A 5.25 1.90 3 5.19 7.19 3.38 90 Swf Ultisols Tuffac. sandstone Recently fall

II8A 5.96 0.48 5 6.10 6.25 5.12 90 Swf Ultisols Tuffac. sandstone Torn

II8B 2.94 0.50 3 2.88 3.47 2.48 90 Swf Ultisols Tuffac. sandstone

II8C 7.12 1 90 Swf Ultisols Tuffac. sandstone

II8C 2.46 0.37 3 2.63 2.72 2.04 90 Swf Ultisols Tuffac. sandstone

II8D 5.34 0.21 3 5.26 5.58 5.18 90 Swf Ultisols Tuffac. sandstone

II9A 7.05 1.47 22 6.91 12.21 4.88 139 Wfs Ultisols Tuffac. sandstone

II9A 3.07 1.04 22 2.68 5.38 1.79 139 Wfs Ultisols Tuffac. sandstone Other

II9B 2.77 1.58 7 3.16 4.85 0.38 139 Wfs Ultisols Tuffac. sandstone

II9C 4.72 0.41 6 4.79 5.19 4.02 139 Wfs Ultisols Tuffac. sandstone

II9D 4.38 0.29 6 4.44 4.80 3.95 139 Wfs Ultisols Tuffac. sandstone

Nitrogen (%) II8A 0.91 0.13 4 0.91 1.02 0.76 90 Swf Ultisols Tuffac. sandstone Recently fall

II8A 1.14 0.19 6 1.14 1.36 0.89 90 Swf Ultisols Tuffac. sandstone Torn

II8B 0.68 0.09 4 0.68 0.80 0.61 90 Swf Ultisols Tuffac. sandstone

II8C 1.67 0.35 2 1.67 1.91 1.42 90 Swf Ultisols Tuffac. sandstone

II8C 0.70 0.31 4 0.70 1.08 0.32 90 Swf Ultisols Tuffac. sandstone

II8D 1.46 0.13 4 1.46 1.58 1.27 90 Swf Ultisols Tuffac. sandstone

II9A 1.00 0.26 22 0.94 1.68 0.72 139 Wfs Ultisols Tuffac. sandstone

II9A 0.77 0.33 22 0.66 1.37 0.37 139 Wfs Ultisols Tuffac. sandstone Other

II9B 0.53 0.23 12 0.59 0.77 0.12 139 Wfs Ultisols Tuffac. sandstone

II9C 1.51 0.32 11 1.51 2.18 1.08 139 Wfs Ultisols Tuffac. sandstone

II9D 1.62 0.27 11 1.58 2.14 1.32 139 Wfs Ultisols Tuffac. sandstone

Phosphorus (mg/g) II8A 0.35 0.05 3 0.32 0.41 0.32 90 Swf Ultisols Tuffac. sandstone Recently fall
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

137

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8A 0.47 0.11 5 0.44 0.66 0.39 90 Swf Ultisols Tuffac. sandstone Torn

II8B 0.85 1 90 Swf Ultisols Tuffac. sandstone

II8C 0.42 0.17 3 0.45 0.56 0.23 90 Swf Ultisols Tuffac. sandstone

II8C 0.63 0.06 3 0.59 0.70 0.59 90 Swf Ultisols Tuffac. sandstone

II8D 0.37 0.06 3 0.35 0.44 0.35 90 Swf Ultisols Tuffac. sandstone

II9A 0.48 0.15 22 0.45 0.88 0.30 139 Wfs Ultisols Tuffac. sandstone

II9A 0.54 0.29 11 0.44 1.19 0.21 139 Wfs Ultisols Tuffac. sandstone Other

II9B 0.18 0.11 7 0.19 0.29 0.03 139 Wfs Ultisols Tuffac. sandstone

II9C 1.10 0.22 6 1.11 1.38 0.76 139 Wfs Ultisols Tuffac. sandstone

II9D 0.99 0.20 6 0.97 1.29 0.76 139 Wfs Ultisols Tuffac. sandstone

Potassium (mg/g) II8A 6.12 0.28 4 6.28 6.28 5.80 90 Swf Ultisols Tuffac. sandstone Recently fall

II8A 5.20 2.00 5 5.91 7.15 2.97 90 Swf Ultisols Tuffac. sandstone Torn

II8B 2.27 0.72 3 2.67 2.70 1.44 90 Swf Ultisols Tuffac. sandstone

II8C 4.31 1 90 Swf Ultisols Tuffac. sandstone

II8C 2.61 0.53 3 2.46 3.19 2.17 90 Swf Ultisols Tuffac. sandstone

II8D 4.78 1.80 3 4.96 6.48 2.90 90 Swf Ultisols Tuffac. sandstone

II9A 7.62 1.82 22 7.78 10.58 4.38 139 Wfs Ultisols Tuffac. sandstone

II9A 5.05 2.97 22 4.02 13.27 1.60 139 Wfs Ultisols Tuffac. sandstone Other

II9B 2.23 1.44 7 2.05 4.71 0.41 139 Wfs Ultisols Tuffac. sandstone

II9C 11.62 3.21 6 11.23 15.66 8.31 139 Wfs Ultisols Tuffac. sandstone

II9D 5.84 1.33 6 6.17 7.10 3.51 139 Wfs Ultisols Tuffac. sandstone

Wood density (g/cc) II2B 0.44 0.11 173 0.44 0.72 0.18 97 Wfs Inceptisols Tuffac. sandstone Heartwood

II2B 0.52 0.10 173 0.53 1.06 0.21 97 Wfs Inceptisols Tuffac. sandstone Sapwood

Hilia paraitica Aluminum (mg/g) II4 0.08 1 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II4 6.66 1 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II4 0.14 1 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II4 3.28 1 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II4 0.05 1 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

138

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Nitrogen (%) II4 0.82 1 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II4 0.53 1 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II4 14.39 1 135 Lmrf Ultisols Tuffac. sandstone

Homalium racemoso Aluminum (mg/kg) II1A 62 1 148 Smf Ultisols Tuffac. sandstone

Ash (%) II1A 6.05 1 148 Smf Ultisols Tuffac. sandstone

C/N II1A 36 1 148 Smf Ultisols Tuffac. sandstone

Calcium (mg/g) II1A 12.20 1 148 Smf Ultisols Tuffac. sandstone

Cobalt (mg/kg) II1A 5.80 1 148 Smf Ultisols Tuffac. sandstone

Copper (mg/kg) II1A 2.85 1 148 Smf Ultisols Tuffac. sandstone

Chromium (mg/kg) II1A 1.49 1 148 Smf Ultisols Tuffac. sandstone

Iron (mg/g) II1A 0.06 1 148 Smf Ultisols Tuffac. sandstone

Magnesium (mg/g) II1A 5.19 1 148 Smf Ultisols Tuffac. sandstone

Manganese (mg/g) II1A 0.04 1 148 Smf Ultisols Tuffac. sandstone

Niquel (mg/kg) II1A 37.04 1 148 Smf Ultisols Tuffac. sandstone

Nitrogen (%) II1A 1.22 1 148 Smf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.46 1 148 Smf Ultisols Tuffac. sandstone

Potassium (mg/g) II1A 2.73 1 148 Smf Ultisols Tuffac. sandstone

Sodium (mg/g) II1A 185.41 1 148 Smf Ultisols Tuffac. sandstone

Zinc (mg/kg) II1A 13 1 148 Smf Ultisols Tuffac. sandstone

Hyeronima clusioides Wood density (g/cc) II1B 0.77 0.03 3 0.78 0.79 0.73 97 Wfs Inceptisols Tuffac. sandstone

Hypolepsis spp Aluminum (mg/g) II6 0.20 0.16 3 0.29 0.30 0.01 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II6 2.09 0.60 3 1.75 2.78 1.73 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II6 0.13 0.02 3 0.12 0.13 0.09 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II6 0.14 0.08 3 1.44 1.50 1.34 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II6 0.16 0.22 3 0.04 0.42 0.04 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II6 1.19 0.21 3 1.08 1.43 1.07 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

139

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Phosphorus (mg/g) II6 0.62 0.16 3 0.53 0.80 0.53 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II6 18.28 12.05 3 11.51 32.19 11.12 135 Lmrf Ultisols Tuffac. sandstone

Ichnanthus pallens Aluminum (mg/g) II3 0.67 1.08 24 0.32 4.77 0.05 135 Lmrf Ultisols Tuffac. sandstone

C/N II3 19 4 21 19 27 14 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II3 1.91 0.80 24 1.84 3.52 0.38 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II3 39 1 21 38 41 36 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II3 0.63 0.95 24 0.25 4.01 0.09 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II3 3.54 2.59 24 3.04 13.29 1.14 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II3 0.48 0.33 24 0.43 1.78 0.15 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II3 1.75 0.47 40 1.67 2.77 0.83 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II3 0.66 0.18 24 0.70 1.08 0.34 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II3 21.27 8.07 24 23.24 39.53 1.84 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II3 0.51 0.16 21 0.48 0.86 0.31 135 Lmrf Ultisols Tuffac. sandstone

Ichnanthus spp. Aluminum (mg/g) II3 1.35 1.30 12 0.74 4.23 0.30 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II3 2.26 0.72 12 2.11 3.71 1.33 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II3 1.16 1.20 12 0.82 4.35 0.20 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II3 2.74 0.64 12 2.76 4.34 1.68 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II3 0.38 0.12 12 0.34 0.58 0.19 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II3 1.53 0.36 12 1.57 2.00 0.86 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II3 0.66 0.15 12 0.63 0.91 0.46 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II3 22.18 3.93 12 23.77 28.94 15.68 135 Lmrf Ultisols Tuffac. sandstone

Ilex sideroxyloides Aluminum (mg/g) II1A 0.24 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.22 1 93 Wflm Inceptisols Tuffac. sandstone

Ash (%) II1A 3.25 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 4.43 1 93 Wflm Inceptisols Tuffac. sandstone

C/N II1A 44 1 93 Wflm Inceptisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

140

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1B 76 1 93 Wflm Inceptisols Tuffac. sandstone

Calcium (mg/g) II1A 4.75 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 11.65 1 93 Wflm Inceptisols Tuffac. sandstone

Iron (mg/g) II1A 0.10 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.07 1 93 Wflm Inceptisols Tuffac. sandstone

Magnesium (mg/g) II1A 2.16 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 1.14 1 93 Wflm Inceptisols Tuffac. sandstone

Manganese (mg/g) II1A 0.57 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.72 1 93 Wflm Inceptisols Tuffac. sandstone

Nitrogen (%) II1A 1.01 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.58 1 93 Wflm Inceptisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.77 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.47 1 93 Wflm Inceptisols Tuffac. sandstone

Potassium (mg/g) II1A 10.27 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 8.16 1 93 Wflm Inceptisols Tuffac. sandstone

Inga fagifolia Aluminum (mg/g) II8A 0.17 0.12 6 0.12 0.41 0.09 145 Wfs Ultisols Tuffac. sandstone

Ash (%) II8A 6.31 0.36 6 6.35 6.73 5.69 145 Wfs Ultisols Tuffac. sandstone

C/N II8A 29 2 6 29 31 26 145 Wfs Ultisols Tuffac. sandstone

Calcium (mg/g) II8A 10.76 2.23 6 10.15 14.32 8.16 145 Wfs Ultisols Tuffac. sandstone

Carbon (%) II8A 52 1 6 52 53 51 145 Wfs Ultisols Tuffac. sandstone

Iron (mg/g) II8A 0.12 0.02 6 0.12 0.17 0.10 145 Wfs Ultisols Tuffac. sandstone

Magnesium (mg/g) II8A 1.64 0.19 6 1.62 1.88 1.43 145 Wfs Ultisols Tuffac. sandstone

Manganese (mg/g) II8A 0.17 0.05 6 0.15 0.23 0.13 145 Wfs Ultisols Tuffac. sandstone

Nitrogen (%) II8A 1.82 0.11 6 1.79 2.00 1.71 145 Wfs Ultisols Tuffac. sandstone

Phosphorus (mg/g) II8A 0.26 0.06 6 0.24 0.36 0.21 145 Wfs Ultisols Tuffac. sandstone

Potassium (mg/g) II8A 2.75 0.29 6 2.70 3.27 2.49 145 Wfs Ultisols Tuffac. sandstone

Sulfur (%) II8A 0.19 0.01 6 0.19 0.20 0.18 145 Wfs Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

141

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Inga laurina Aluminum (mg/g) II1E 0.63 0.17 2 0.63 0.75 0.51 135 Lmrf Ultisols Tuffac. sandstone

II1H 0.15 0.11 7 0.14 0.36 0.05 135 Lmrf Ultisols Tuffac. sandstone

II7A 0.37 0.33 3 0.25 0.74 0.12 70 Lmrf Ultisols Tuffac. sandstone 1year

II7A 0.41 0.49 5 0.21 1.28 0.09 70 Lmrf Ultisols Tuffac. sandstone 2year

II7B 0.02 0.01 4 0.02 0.04 0.01 70 Lmrf Ultisols Tuffac. sandstone 1year

II7B 0.04 0.02 5 0.05 0.06 0.02 70 Lmrf Ultisols Tuffac. sandstone 2year

II7C 0.18 0.14 3 0.20 0.32 0.03 70 Lmrf Ultisols Tuffac. sandstone 1year

II7C 0.23 0.29 4 0.13 0.66 0.02 70 Lmrf Ultisols Tuffac. sandstone 2year

Ash (%) II1A 4.55 0.86 6 4.51 5.87 3.46 126 Ultisols Siltstone,sandstone Urban forest, RP

II1A 4.20 1 71 Lmrf Ultisols Tuffac. sandstone

II1C 1.41 0.48 6 1.41 2.10 0.66 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 5.58 0.89 5 5.70 6.64 4.21 126 Ultisols Siltstone,sandstone Urban forest, RP

C/N II1A 24 1 6 24 26 23 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 95 20 6 90 134 78 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 37 5 5 37 43 32 126 Ultisols Siltstone,sandstone Urban forest, RP

Calcium (mg/g) II1A 5.21 1.45 9 4.90 8.04 2.96 135 Lmrf Ultisols Tuffac. sandstone

II1B 7.04 1.87 10 7.60 9.20 3.76 135 Lmrf Ultisols Tuffac. sandstone

II1C 1.38 0.26 17 1.34 1.84 0.99 135 Lmrf Ultisols Tuffac. sandstone

II1Ci 8.51 1.28 18 8.00 11.42 6.96 135 Lmrf Ultisols Tuffac. sandstone

II1E 2.48 0.21 2 2.48 2.62 2.33 135 Lmrf Ultisols Tuffac. sandstone

II1H 4.10 0.84 7 4.33 5.05 2.93 135 Lmrf Ultisols Tuffac. sandstone

II7A 12.04 0.68 3 12.07 12.69 11.34 70 Lmrf Ultisols Tuffac. sandstone 1year

II7A 8.29 1.09 5 8.87 9.15 6.76 70 Lmrf Ultisols Tuffac. sandstone 2year

II7B 2.65 0.91 4 2.48 3.91 1.75 70 Lmrf Ultisols Tuffac. sandstone 1year

II7B 2.69 0.91 5 2.40 3.67 1.66 70 Lmrf Ultisols Tuffac. sandstone 2year

II7C 4.86 1.09 3 5.06 5.84 3.68 70 Lmrf Ultisols Tuffac. sandstone 1year

II7C 5.45 2.24 4 4.91 8.39 3.60 70 Lmrf Ultisols Tuffac. sandstone 2year

Carbon (%) II1A 53 0 6 54 54 53 126 Ultisols Siltstone,sandstone Urban forest, RP

II1A 54 1 71 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

142

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1C 53 0 6 53 53 53 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 52 1 5 3 53 51 126 Ultisols Siltstone,sandstone Urban forest, RP

Iron (mg/g) II1E 0.96 0.35 2 0.96 1.21 0.71 135 Lmrf Ultisols Tuffac. sandstone

II1H 0.16 0.08 7 0.14 0.32 0.10 135 Lmrf Ultisols Tuffac. sandstone

II7A 0.47 0.44 3 0.34 0.97 0.11 70 Lmrf Ultisols Tuffac. sandstone 1year

II7A 0.42 0.55 5 0.20 1.40 0.07 70 Lmrf Ultisols Tuffac. sandstone 2year

II7B 0.02 0.02 4 0.02 0.05 0.01 70 Lmrf Ultisols Tuffac. sandstone 1year

II7B 0.06 0.01 5 0.06 0.07 0.04 70 Lmrf Ultisols Tuffac. sandstone 2year

II7C 0.13 0.10 3 0.14 0.22 0.03 70 Lmrf Ultisols Tuffac. sandstone 1year

II7C 0.29 0.41 4 0.12 0.89 0.02 70 Lmrf Ultisols Tuffac. sandstone 2year

Magnesium (mg/g) II1A 1.26 0.44 9 1.48 1.68 0.42 135 Lmrf Ultisols Tuffac. sandstone

II1B 1.19 0.31 10 1.32 1.72 0.75 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.18 0.04 17 0.17 0.29 0.12 135 Lmrf Ultisols Tuffac. sandstone

II1Ci 0.63 0.07 18 0.64 0.74 0.43 135 Lmrf Ultisols Tuffac. sandstone

II1E 1.08 0.16 2 1.08 1.19 0.97 135 Lmrf Ultisols Tuffac. sandstone

II1H 1.19 0.14 7 1.14 1.44 0.03 135 Lmrf Ultisols Tuffac. sandstone

II7A 0.83 0.09 3 0.86 0.90 0.73 70 Lmrf Ultisols Tuffac. sandstone 1year

II7A 0.65 0.20 5 0.69 0.81 0.36 70 Lmrf Ultisols Tuffac. sandstone 2year

II7B 0.32 0.05 4 0.31 0.39 0.28 70 Lmrf Ultisols Tuffac. sandstone 1year

II7B 0.38 0.14 5 0.33 0.57 0.25 70 Lmrf Ultisols Tuffac. sandstone 2year

II7C 0.42 0.12 3 0.38 0.55 0.32 70 Lmrf Ultisols Tuffac. sandstone 1year

II7C 0.49 0.35 4 0.36 1.01 0.25 70 Lmrf Ultisols Tuffac. sandstone 2year

Manganese (mg/g) II1E 0.06 0.02 2 0.06 0.07 0.04 135 Lmrf Ultisols Tuffac. sandstone

II1H 0.13 0.03 7 0.15 0.16 0.10 135 Lmrf Ultisols Tuffac. sandstone

II7A 0.12 0.01 3 0.13 0.13 0.12 70 Lmrf Ultisols Tuffac. sandstone 1year

II7A 0.08 0.03 5 0.07 0.13 0.05 70 Lmrf Ultisols Tuffac. sandstone 2year

II7B 0.03 0.01 4 0.03 0.05 0.02 70 Lmrf Ultisols Tuffac. sandstone 1year

II7B 0.03 0.01 5 0.03 0.04 0.02 70 Lmrf Ultisols Tuffac. sandstone 2year

II7C 0.05 0.03 3 0.05 0.08 0.03 70 Lmrf Ultisols Tuffac. sandstone 1year
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

143

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II7C 0.08 0.03 4 0.07 0.11 0.05 70 Lmrf Ultisols Tuffac. sandstone 2year

Nitrogen (%) II1A 2.25 0.08 6 2.27 2.34 2.10 126 Ultisols Siltstone,sandstone Urban forest, RP

II1A 2.25 0.93 9 2.74 2.90 0.66 135 Lmrf Ultisols Tuffac. sandstone

II1A 2.87 1 71 Lmrf Ultisols Tuffac. sandstone

II1B 1.48 0.83 10 1.11 2.86 0.74 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.57 0.10 6 0.59 0.67 0.40 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 0.25 0.02 17 0.26 0.30 0.21 135 Lmrf Ultisols Tuffac. sandstone

II1Ci 1.45 0.19 5 1.39 1.66 1.22 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 1.47 0.14 18 1.48 1.69 1.09 135 Lmrf Ultisols Tuffac. sandstone

II1E 1.19 0.12 2 1.19 1.27 1.10 135 Lmrf Ultisols Tuffac. sandstone

II1H 1.75 0.28 7 1.79 2.10 1.44 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.93 0.35 9 1.09 1.29 0.31 135 Lmrf Ultisols Tuffac. sandstone

II1B 0.62 0.42 10 0.41 1.44 0.29 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.06 0.02 17 0.05 0.09 0.03 135 Lmrf Ultisols Tuffac. sandstone

II1Ci 0.27 0.03 18 0.27 0.33 0.24 135 Lmrf Ultisols Tuffac. sandstone

II1E 0.34 0.03 2 0.34 0.36 0.32 135 Lmrf Ultisols Tuffac. sandstone

II1H 0.66 0.08 7 0.64 0.82 0.56 135 Lmrf Ultisols Tuffac. sandstone

II7A 0.24 0.05 3 0.23 0.30 0.20 70 Lmrf Ultisols Tuffac. sandstone 1year

II7A 0.17 0.03 5 0.16 0.22 0.13 70 Lmrf Ultisols Tuffac. sandstone 2year

II7B 0.05 0.02 4 0.06 0.71 0.03 70 Lmrf Ultisols Tuffac. sandstone 1year

II7B 0.03 0.01 5 0.03 0.04 0.03 70 Lmrf Ultisols Tuffac. sandstone 2year

II7C 0.12 0.05 3 0.14 0.16 0.07 70 Lmrf Ultisols Tuffac. sandstone 1year

II7C 0.13 0.04 4 0.15 0.16 0.07 70 Lmrf Ultisols Tuffac. sandstone 2year

Potassium (mg/g) II1A 6.99 2.13 9 7.31 10.44 3.86 135 Lmrf Ultisols Tuffac. sandstone

II1B 5.71 2.47 10 4.84 11.55 3.43 135 Lmrf Ultisols Tuffac. sandstone

II1C 2.74 0.59 17 2.60 3.94 1.74 135 Lmrf Ultisols Tuffac. sandstone

II1Ci 5.84 0.56 18 5.95 6.80 4.90 135 Lmrf Ultisols Tuffac. sandstone

II1E 5.39 0.65 2 5.39 5.85 4.93 135 Lmrf Ultisols Tuffac. sandstone

II1H 7.25 1.34 7 7.33 9.47 5.73 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

144

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II7A 2.66 0.37 3 2.65 3.03 2.29 70 Lmrf Ultisols Tuffac. sandstone 1year

II7A 1.19 0.25 5 1.17 1.50 0.94 70 Lmrf Ultisols Tuffac. sandstone 2year

II7B 1.64 0.53 4 1.60 2.20 1.15 70 Lmrf Ultisols Tuffac. sandstone 1year

II7B 1.08 0.42 5 1.19 1.54 0.61 70 Lmrf Ultisols Tuffac. sandstone 2year

II7C 2.38 0.99 3 2.11 3.47 1.55 70 Lmrf Ultisols Tuffac. sandstone 1year

II7C 1.11 0.13 4 1.12 1.23 0.97 70 Lmrf Ultisols Tuffac. sandstone 2year

Sulfur (%) II1A 0.26 0.02 6 0.26 0.29 0.25 126 Ultisols Siltstone,sandstone Urban forest, RP

II1A 0.36 1 71 Lmrf Ultisols Tuffac. sandstone

II1C 0.06 0.01 6 0.06 0.07 0.04 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 0.12 0.02 5 0.12 0.14 0.09 126 Ultisols Siltstone,sandstone Urban forest, RP

Wood density (g/cc) II1C 0.61 0.05 5 0.60 0.69 0.56 126 Ultisols Siltstone,sandstone Urban forest, RP

Inga vera Aluminum (mg/g) II8A 0.24 0.07 7 0.24 0.32 0.15 145 Wfs Ultisols Tuffac. sandstone

II8A 0.97 0.65 3 1.25 1.43 0.23 146 Wfs Ultisols Tuffac. sandstone

II8B 0.27 0.03 2 0.27 0.28 0.25 146 Wfs Ultisols Tuffac. sandstone

Ash (%) II8A 6.68 1.64 6 6.47 9.37 4.70 145 Wfs Ultisols Tuffac. sandstone

II8A 6.81 1 71 Wfs Ultisols Tuffac. sandstone

II8A 6.54 1.03 3 7.07 7.20 5.36 145 Wfs Ultisols Tuffac. sandstone

II8B 2.36 1.40 2 2.36 3.38 1.34 145 Wfs Ultisols Tuffac. sandstone

C/N II8A 27 1 7 27 28 26 145 Wfs Ultisols Tuffac. sandstone

II8A 15 1 3 15 16 15 145 Wfs Ultisols Tuffac. sandstone

II8B 42 4 2 42 44 39 145 Wfs Ultisols Tuffac. sandstone

Calcium (mg/g) II1A 8.80 2.57 5 7.80 11.81 5.98 100 Lmrf Ultisols Tuffac. sandstone Fresh

II1A 11.63 3.94 5 11.75 16.92 6.91 100 Lmrf Ultisols Tuffac. sandstone 7 day Decomp.

II1A 10.50 3.00 5 10.85 13.24 7.06 100 Lmrf Ultisols Tuffac. sandstone 14 day Decomp.

II8A 9.52 1.32 7 10.24 11.05 8.05 145 Wfs Ultisols Tuffac. sandstone

II8A 15.36 2.52 3 16.85 17.90 11.34 145 Wfs Ultisols Tuffac. sandstone

II8B 6.03 2.59 2 6.03 7.86 4.20 145 Wfs Ultisols Tuffac. sandstone

Carbon (%) II8A 52 1 7 52 53 50 145 Wfs Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

145

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8A 54 1 71 Wfs Ultisols Tuffac. sandstone

II8A 53 1 3 53 54 53 145 Wfs Ultisols Tuffac. sandstone

II8B 53 2 2 53 54 52 145 Wfs Ultisols Tuffac. sandstone

Iron (mg/g) II8A 0.21 0.06 7 0.19 0.30 0.15 145 Wfs Ultisols Tuffac. sandstone

II8A 0.59 0.07 3 0.56 0.67 0.54 145 Wfs Ultisols Tuffac. sandstone

II8B 0.18 0.07 2 0.18 0.23 0.13 145 Wfs Ultisols Tuffac. sandstone

Magnesium (mg/g) II1A 3.16 0.74 5 2.78 4.25 2.49 100 Lmrf Ultisols Tuffac. sandstone Fresh

II1A 3.41 1.10 5 3.31 4.72 2.07 100 Lmrf Ultisols Tuffac. sandstone 7 day Decomp.

II1A 2.99 0.77 5 3.11 3.71 1.75 100 Lmrf Ultisols Tuffac. sandstone 14 day Decomp.

II8A 1.24 0.16 7 1.26 1.47 1.02 145 Wfs Ultisols Tuffac. sandstone

II8A 1.56 0.32 3 1.48 1.91 1.30 145 Wfs Ultisols Tuffac. sandstone

II8B 0.70 0.09 2 0.70 0.76 0.63 145 Wfs Ultisols Tuffac. sandstone

Manganese (mg/g) II8A 0.26 0.06 7 0.28 0.32 0.15 145 Wfs Ultisols Tuffac. sandstone

II8A 0.15 0.06 3 0.16 0.20 0.09 145 Wfs Ultisols Tuffac. sandstone

II8B 0.05 0.00 2 0.05 0.05 0.04 145 Wfs Ultisols Tuffac. sandstone

Nitrogen (%) II1A 2.08 0.09 5 2.07 2.17 1.95 100 Lmrf Ultisols Tuffac. sandstone Fresh

II1A 2.04 0.14 5 2.01 2.22 1.89 100 Lmrf Ultisols Tuffac. sandstone 7 day Decomp.

II1A 2.17 0.16 5 2.21 2.33 1.91 100 Lmrf Ultisols Tuffac. sandstone 14 day Decomp.

II8A 1.94 0.07 7 1.91 2.03 1.85 145 Wfs Ultisols Tuffac. sandstone

II8A 1.99 1 71 Wfs Ultisols Tuffac. sandstone

II8A 3.46 0.03 3 3.47 3.48 3.42 145 Wfs Ultisols Tuffac. sandstone

II8B 1.29 0.09 2 1.29 1.35 1.22 145 Wfs Ultisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.85 0.19 5 0.80 1.06 0.61 100 Lmrf Ultisols Tuffac. sandstone Fresh

II1A 0.82 0.23 5 0.81 1.07 0.49 100 Lmrf Ultisols Tuffac. sandstone 7 day Decomp.

II1A 0.86 0.17 5 0.88 1.05 0.60 100 Lmrf Ultisols Tuffac. sandstone 14 day Decomp.

II8A 0.35 0.03 7 0.36 0.39 0.30 145 Wfs Ultisols Tuffac. sandstone

II8A 0.77 0.21 3 0.68 1.02 0.62 145 Wfs Ultisols Tuffac. sandstone

II8B 0.42 0.18 2 0.40 0.53 0.27 145 Wfs Ultisols Tuffac. sandstone

Potassium (mg/g) II1A 3.20 0.73 5 2.82 4.11 2.55 100 Lmrf Ultisols Tuffac. sandstone Fresh
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

146

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 2.79 0.56 5 2.66 3.55 2.16 100 Lmrf Ultisols Tuffac. sandstone 7 day Decomp.

II1A 2.57 0.45 5 2.61 3.04 1.85 100 Lmrf Ultisols Tuffac. sandstone 14 day Decomp.

II8A 3.43 0.38 7 3.39 3.89 2.88 145 Wfs Ultisols Tuffac. sandstone

II8A 4.10 4.12 3 1.92 8.85 1.52 145 Wfs Ultisols Tuffac. sandstone

II8B 3.32 3.04 2 3.32 5.47 1.17 145 Wfs Ultisols Tuffac. sandstone

Sulfur (%) II8A 0.17 0.01 7 0.17 0.18 0.14 145 Wfs Ultisols Tuffac. sandstone

II8A 0.28 1 71 Wfs Ultisols Tuffac. sandstone

II8A 0.50 0.06 3 0.47 0.58 0.46 145 Wfs Ultisols Tuffac. sandstone

II8B 0.25 0.01 2 0.25 0.26 0.25 145 Wfs Ultisols Tuffac. sandstone

Jacquinia berterii Aluminum (mg/kg) II1A 40 58 10 21 201 2 151 Sdf Mollisols Alluvial deposits

Ash (%) II1A 9.02 1.99 9 8.68 11.60 5.56 151 Sdf Mollisols Alluvial deposits

C/N II1A 60 6 9 60 69 53 151 Sdf Mollisols Alluvial deposits

Calcium (mg/g) II1A 7.76 1 104 Sdf Mollisols Alluvial deposits

II1A 7.76 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1A 27.65 8.71 10 25.53 42.43 17.25 151 Sdf Mollisols Alluvial deposits

II1Ci 30.24 1 104 Sdf Mollisols Alluvial deposits

II1Ci 30.24 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Carbon (%) II1A 54 2 9 55 56 51 151 Sdf Mollisols Alluvial deposits

Iron (mg/kg) II1A 33 26 10 30 105 13 151 Sdf Mollisols Alluvial deposits

Magnesium (mg/g) II1A 2.30 1 104 Sdf Mollisols Alluvial deposits

II1A 2.30 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1A 1.20 0.34 10 1.17 1.70 0.51 151 Sdf Mollisols Alluvial deposits

II1Ci 0.59 1 104 Sdf Mollisols Alluvial deposits

II1Ci 0.59 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Manganese (mg/kg) II1A 13 7 10 11 26 5 151 Sdf Mollisols Alluvial deposits

Nitrogen (%) II1A 0.78 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1A 0.90 0.07 9 0.91 0.99 0.82 151 Sdf Mollisols Alluvial deposits

II1Ci 0.67 1 149 Sdf Mollisols Alluvial deposits Guanica Forest
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

147

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Phosphorus (mg/g) II1A 0.18 1 104 Sdf Mollisols Alluvial deposits

II1A 0.18 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1A 0.22 0.05 10 0.23 0.28 0.16 151 Sdf Mollisols Alluvial deposits

II1Ci 0.15 1 104 Sdf Mollisols Alluvial deposits

II1Ci 0.15 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Potassium (mg/g) II1A 7.28 1 104 Sdf Mollisols Alluvial deposits

II1A 7.28 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1A 4.98 0.69 10 4.97 6.00 4.19 151 Sdf Mollisols Alluvial deposits

II1Ci 7.12 1 104 Sdf Mollisols Alluvial deposits

II1Ci 7.12 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Sulfur (%) II1A 0.26 0.03 9 0.27 0.31 0.21 151 Sdf Mollisols Alluvial deposits

Juglans jamaicensis Wood density (g/cc) II1C 0.49 0.03 3 0.50 0.51 0.46 115

Kalanchoe pinnata Calcium (mg/g) II3Ai 54.72 20.43 12 50.10 80.02 32.04 105

II3Aii 71.55 25.23 12 70.47 97.98 46.54 105 Shade

II3C 41.92 16.25 8 36.08 68.81 27.34 105 Shade

II3C 27.22 5.74 8 27.06 36.62 20.76 105 Sun

II3E 15.67 1.49 8 15.97 17.15 13.61 105 Shade

II3E 16.50 1.83 8 15.73 19.78 14.96 105 Sun

Magnesium (mg/g) II3Ai 3.38 0.35 12 3.36 3.84 2.93 105

II3Aii 4.70 1.11 12 4.69 5.94 3.36 105 Shade

II3C 2.28 0.73 8 1.90 3.49 1.83 105 Shade

II3C 2.03 0.14 8 2.01 2.21 1.84 105 Sun

II3E 2.09 0.66 8 1.83 3.10 1.42 105 Shade

II3E 2.29 0.24 8 2.35 2.56 1.92 105 Sun

Nitrogen (%) II3Ai 0.70 0.17 7 0.77 0.90 0.48 105

II3Aii 1.06 0.09 7 1.08 1.19 0.93 105 Shade

II3C 0.33 0.05 5 0.32 0.40 0.27 105 Shade
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

148

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II3C 0.29 0.05 5 0.28 0.36 0.22 105 Sun

II3E 0.33 0.05 5 0.32 0.42 0.28 105 Shade

II3E 0.35 0.02 5 0.34 0.37 0.33 105 Sun

Phosphorus (mg/g) II3Ai 4.51 0.88 12 4.40 5.61 3.42 105

II3Aii 3.42 0.12 12 3.38 3.64 3.27 105 Shade

II3C 2.16 0.20 8 2.12 2.48 1.92 105 Shade

II3C 2.14 0.06 8 2.14 2.22 2.05 105 Sun

II3E 1.31 0.10 8 1.28 1.48 1.20 105 Shade

II3E 1.26 0.04 8 1.25 1.33 1.21 105 Sun

Potassium (mg/g) II3Ai 19.83 3.01 12 19.15 23.99 13.49 105

II3Aii 21.65 2.81 12 21.02 26.38 17.38 105 Shade

II3C 23.29 10.72 8 19.38 44.13 14.27 105 Shade

II3C 15.44 2.36 8 14.53 19.74 13.15 105 Sun

II3E 10.79 1.51 8 10.39 13.60 8.96 105 Shade

II3E 9.21 0.45 8 9.24 9.96 8.62 105 Sun

Khaya grandifolia Wood density (g/cc) II1B 0.57 0.02 5 0.58 0.58 0.54 97 Wfs Inceptisols Tuffac. sandstone

Khaya nyasica Ash (%) II8A 7.21 1.65 4 7.21 9.52 5.62 90 Swf Ultisols Tuffac. sandstone

II8A 13.35 5.50 4 13.35 21.90 8.42 90 Swf Ultisols Tuffac. sandstone Torn

II8A 8.37 0.69 4 8.37 9.34 7.86 90 Swf Ultisols Tuffac. sandstone Whole

II8B 3.94 1.43 4 3.94 5.49 2.15 90 Swf Ultisols Tuffac. sandstone

II8C 1.91 1 90 Swf Ultisols Tuffac. sandstone

II8D 21.85 6.92 4 21.85 29.40 13.68 90 Swf Ultisols Tuffac. sandstone

Calcium (mg/g) II1A 5.63 1.42 8 5.46 7.76 3.49 135 Lmrf Ultisols Tuffac. sandstone

II1B 5.39 1.11 8 5.40 7.53 3.89 135 Lmrf Ultisols Tuffac. sandstone

II1C 1.35 0.26 15 1.34 2.10 1.08 135 Lmrf Ultisols Tuffac. sandstone

II1Ci 7.46 1.01 15 7.68 9.29 5.95 135 Lmrf Ultisols Tuffac. sandstone

II8A 12.23 1.36 4 12.23 14.24 11.32 90 Swf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

149

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8A 11.02 1.70 4 11.02 12.18 8.63 90 Swf Ultisols Tuffac. sandstone Torn

II8A 11.94 1.02 4 11.94 13.18 11.06 90 Swf Ultisols Tuffac. sandstone Whole

II8B 8.03 3.15 4 8.03 11.09 4.15 90 Swf Ultisols Tuffac. sandstone

II8C 2.23 1 90 Swf Ultisols Tuffac. sandstone

II8D 7.49 2.97 4 7.94 11.19 4.01 90 Swf Ultisols Tuffac. sandstone

II9A 7.58 1.14 22 7.70 9.31 4.98 139 Wfs Ultisols Tuffac. sandstone

II9A 4.44 2.29 21 3.67 10.84 2.64 139 Wfs Ultisols Tuffac. sandstone Other

II9B 6.29 1.59 6 5.80 9.30 4.80 139 Wfs Ultisols Tuffac. sandstone

II9C 1.41 0.77 5 1.59 2.41 0.68 139 Wfs Ultisols Tuffac. sandstone

II9D 6.94 1.01 5 7.20 8.36 5.75 139 Wfs Ultisols Tuffac. sandstone

Magnesium (mg/g) II1A 2.26 0.42 8 2.28 2.98 1.52 135 Lmrf Ultisols Tuffac. sandstone

II1B 1.12 0.19 8 1.09 1.50 0.90 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.13 0.06 15 0.10 0.29 0.07 135 Lmrf Ultisols Tuffac. sandstone

II1Ci 0.73 0.19 15 0.75 1.00 0.48 135 Lmrf Ultisols Tuffac. sandstone

II8A 2.79 0.84 4 2.79 4.03 2.26 90 Swf Ultisols Tuffac. sandstone

II8A 2.37 0.18 4 2.37 2.52 2.12 90 Swf Ultisols Tuffac. sandstone Torn

II8A 2.31 0.08 4 2.31 2.38 2.20 90 Swf Ultisols Tuffac. sandstone Whole

II8B 1.85 0.51 4 1.85 2.50 1.40 90 Swf Ultisols Tuffac. sandstone

II8C 0.77 1 90 Swf Ultisols Tuffac. sandstone

II8D 1.82 0.36 4 1.82 2.27 1.43 90 Swf Ultisols Tuffac. sandstone

II9A 2.34 0.38 22 2.41 3.12 1.74 139 Wfs Ultisols Tuffac. sandstone

II9A 1.52 0.59 21 1.36 3.38 0.76 139 Wfs Ultisols Tuffac. sandstone Other

II9B 2.29 0.68 6 2.27 3.10 1.11 139 Wfs Ultisols Tuffac. sandstone

II9C 0.91 0.56 5 1.15 1.39 0.01 139 Wfs Ultisols Tuffac. sandstone

II9D 2.22 0.18 5 2.20 2.41 1.99 139 Wfs Ultisols Tuffac. sandstone

Nitrogen (%) II1A 1.70 0.46 8 1.62 2.53 0.90 135 Lmrf Ultisols Tuffac. sandstone

II1B 0.69 0.12 8 0.67 0.90 0.54 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.08 0.03 15 0.08 0.14 0.05 135 Lmrf Ultisols Tuffac. sandstone

II1Ci 0.33 0.05 15 0.31 0.45 0.24 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

150

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8A 1.24 0.22 4 1.24 1.53 0.99 90 Swf Ultisols Tuffac. sandstone

II8A 1.29 0.12 4 1.29 1.45 1.18 90 Swf Ultisols Tuffac. sandstone Tom leaves

II8A 1.05 0.17 4 1.05 1.29 0.91 90 Swf Ultisols Tuffac. sandstone Whole

II8B 0.63 0.15 4 0.63 0.77 0.41 90 Swf Ultisols Tuffac. sandstone

II8C 0.44 1 90 Swf Ultisols Tuffac. sandstone

II8D 1.15 0.20 4 1.15 1.40 0.91 90 Swf Ultisols Tuffac. sandstone

II9A 1.04 0.18 22 1.06 1.30 0.70 139 Wfs Ultisols Tuffac. sandstone

II9A 0.73 0.34 22 0.61 1.66 0.36 139 Wfs Ultisols Tuffac. sandstone Other

II9B 0.51 0.19 11 0.54 0.77 0.16 139 Wfs Ultisols Tuffac. sandstone

II9C 0.72 0.16 9 0.69 1.07 0.54 139 Wfs Ultisols Tuffac. sandstone

II9D 1.26 0.18 10 1.20 1.57 1.02 139 Wfs Ultisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.90 0.25 8 0.84 1.44 0.67 135 Lmrf Ultisols Tuffac. sandstone

II1B 0.43 0.15 0.40 0.68 0.24 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.04 0.02 15 0.03 0.08 0.02 135 Lmrf Ultisols Tuffac. sandstone

II1Ci 0.13 0.02 15 0.12 0.18 0.10 135 Lmrf Ultisols Tuffac. sandstone

II8A 0.40 0.21 4 0.40 0.63 0.13 90 Swf Ultisols Tuffac. sandstone

II8A 0.35 0.07 4 0.35 0.40 0.24 90 Swf Ultisols Tuffac. sandstone Torn

II8A 0.24 0.10 4 0.24 0.36 0.11 90 Swf Ultisols Tuffac. sandstone Whole

II8B 0.19 0.09 4 0.19 0.24 0.05 90 Swf Ultisols Tuffac. sandstone

II8C 0.25 1 90 Swf Ultisols Tuffac. sandstone

II8D 0.36 0.07 4 0.36 0.42 0.27 90 Swf Ultisols Tuffac. sandstone

II9A 0.35 0.12 22 0.36 0.66 0.17 139 Wfs Ultisols Tuffac. sandstone

II9A 0.27 0.15 21 0.24 0.70 0.08 139 Wfs Ultisols Tuffac. sandstone Other

II9B 0.18 0.10 6 0.18 0.31 0.06 139 Wfs Ultisols Tuffac. sandstone

II9C 0.66 0.10 5 0.62 0.78 0.55 139 Wfs Ultisols Tuffac. sandstone

II9D 0.65 0.21 5 0.58 0.89 0.44 139 Wfs Ultisols Tuffac. sandstone

Potassium (mg/g) II1A 8.95 1.10 8 9.16 10.67 7.03 135 Lmrf Ultisols Tuffac. sandstone

II1B 5.90 1.54 8 6.07 7.92 2.86 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.57 0.48 15 0.37 1.60 0.10 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

151

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1Ci 2.65 1.28 15 2.91 4.46 0.88 135 Lmrf Ultisols Tuffac. sandstone

II8A 4.89 2.80 4 4.89 8.33 1.78 90 Swf Ultisols Tuffac. sandstone

II8A 1.92 0.72 4 1.92 2.88 1.12 90 Swf Ultisols Tuffac. sandstone Torn

II8A 2.05 0.77 4 2.05 3.09 1.23 90 Swf Ultisols Tuffac. sandstone Whole

II8B 1.65 0.91 4 1.65 2.71 0.62 90 Swf Ultisols Tuffac. sandstone

II8C 0.99 1 90 Swf Ultisols Tuffac. sandstone

II8D 1.77 0.63 4 1.77 2.42 0.95 90 Swf Ultisols Tuffac. sandstone

II9A 5.37 1.02 22 4.36 7.09 2.78 139 Wfs Ultisols Tuffac. sandstone

II9A 2.43 1.15 21 2.28 5.43 0.89 139 Wfs Ultisols Tuffac. sandstone Other

II9B 2.65 0.88 6 2.49 3.95 1.77 139 Wfs Ultisols Tuffac. sandstone

II9C 3.53 1.00 5 4.04 4.62 2.35 139 Wfs Ultisols Tuffac. sandstone

II9D 3.84 0.88 5 4.18 4.93 2.86 139 Wfs Ultisols Tuffac. sandstone

Wood density (g/cc) II1B 0.49 0.03 4 0.49 0.52 0.46 97 Wfs Inceptisols Tuffac. sandstone

Khaya senegalensis Wood density (g/cc) II1B 0.50 0.03 5 0.50 0.52 0.46 97 Wfs Inceptisols Tuffac. sandstone

Krugiodendron ferreum Calcium (mg/g) II1A 18.89 1 104 Sdf Mollisols Alluvial deposits

II1A 18.89 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 31.47 1 104 Sdf Mollisols Alluvial deposits

II1Ci 31.47 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Magnesium (mg/g) II1A 3.13 1 104 Sdf Mollisols Alluvial deposits

II1A 3.13 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 1.98 1 104 Sdf Mollisols Alluvial deposits

II1Ci 1.98 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Nitrogen (%) II1A 2.34 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 0.87 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Phosphorus (mg/g) II1A 0.48 1 104 Sdf Mollisols Alluvial deposits

II1A 0.48 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 0.21 1 104 Sdf Mollisols Alluvial deposits
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

152

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1Ci 0.21 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Potassium (mg/g) II1A 14.73 1 104 Sdf Mollisols Alluvial deposits

II1A 14.73 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 7.02 1 104 Sdf Mollisols Alluvial deposits

II1Ci 7.02 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Laguncularia racemosa Aluminum (mg/g) II1A 0.04 0.01 4 0.04 0.05 0.04 78 Mfs Mangle Swamp

II1Aiiib 0.45 0.02 6 0.04 0.07 0.03 83 Mfs Histosol Alluvial deposits Adult 

II1B 0.03 0.01 3 0.04 0.04 0.03 87 Mfs Entisols Compound dunes

II1B 0.01 0.00 2 0.01 0.01 0.01 157 Mfs Entisols Compound dunes

Ash (%) II1A 19.17 3.51 13 19.03 22.71 12.50 72 Dfs Mangle Mangle

II1A 14.70 0.92 4 14.75 15.53 13.75 78 Mfs Mangle Swamp

II1B 2.27 0.43 3 2.09 2.76 1.96 87 Mfs Entisols Compound dunes

II1Aiiib 12.11 2.77 6 13.12 14.66 8.55 83 Mfs Histosol Alluvial deposits Adult 

C/N II1A 32 3 4 32 34 29 78 Mfs Mangle Swamp

II1Aiiib 29 3 6 29 32 26 83 Mfs Histosol Alluvial deposits Adult 

II1B 279 92 3 306 355 177 87 Mfs Entisols Compound dunes

Calcium (mg/g) II1A 33.40 10.60 12 27.20 49.90 23.00 72 Dfs Mangle Mangle

II1A 21.87 1 74 Dfs Mangle Dune deposits Old

II1A 21.38 1 74 Dfs Mangle Dune deposits Senescent

II1A 23.32 1 74 Dfs Mangle Dune deposits Yellow

II1A 11.66 1 74 Dfs Mangle Dune deposits Youg Mature

II1A 25.09 5.21 4 24.99 30.06 20.30 78 Mfs Mangle Swamp

II1Aiiib 19.08 8.20 6 16.48 29.77 11.44 83 Mfs Histosol Alluvial deposits Adult 

II1B 4.31 0.46 3 4.17 4.83 3.94 87 Mfs Entisols Compound dunes

II1B 6.76 0.07 2 6.76 6.71 6.81 157 Mfs Entisols Compound dunes

II9A 21.86 6.22 12 21.86 31.26 11.73 73 Dfs Mangle Dune deposits

II9C 7.36 3.19 10 7.36 13.75 3.03 73 Dfs Mangle Dune deposits

Carbon (%) II1A 39 0 4 39 40 39 78 Mfs Mangle Swamp
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

153

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1Aiiib 47 2 6 47 50 46 83 Mfs Histosol Alluvial deposits Adult 

Iron (mg/g) II1A 0.05 0.04 4 0.04 0.11 0.02 78 Mfs Mangle Swamp

II1Aiiib 0.74 0.26 6 0.77 1.02 0.40 83 Mfs Histosol Alluvial deposits Adult 

II1B 0.06 0.02 3 0.06 0.08 0.04 87 Mfs Entisols Compound dunes

II1B 0.01 0.01 2 0.01 0.01 0.02 157 Mfs Entisols Compound dunes

Magnesium (mg/g) II1A 7.39 1 74 Dfs Mangle Dune deposits Old

II1A 7.35 1 74 Dfs Mangle Dune deposits Senescent

II1A 8.10 1 74 Dfs Mangle Dune deposits Yellow

II1A 4.33 1 74 Dfs Mangle Dune deposits Youg Mature

II1A 5.84 0.41 4 5.82 6.24 5.48 78 Mfs Mangle Swamp

II1B 0.59 0.12 3 0.65 0.66 0.45 87 Mfs Entisols Compound dunes

II1B 0.72 0.13 2 0.72 0.63 0.81 157 Mfs Entisols Compound dunes

II9A 5.13 1.00 12 5.13 6.26 3.33 73 Dfs Mangle Dune deposits

II9C 1.89 0.47 10 1.89 2.41 1.25 73 Dfs Mangle Dune deposits

II1Aiiib 3.07 0.27 6 3.14 3.37 2.68 83 Mfs Histosol Alluvial deposits Adult 

Manganese (mg/g) II1A 0.02 0.01 4 0.02 0.03 0.01 78 Mfs Mangle Swamp

II1Aiiib 0.08 0.04 6 0.07 0.14 0.05 83 Mfs Histosol Alluvial deposits Adult 

II1B 0.03 0.02 3 0.02 0.05 0.01 87 Mfs Entisols Compound dunes

II1B 0.01 0.00 2 0.01 0.01 0.01 157 Mfs Entisols Compound dunes

Nitrogen (%) II1A 0.83 0.30 6 0.68 1.27 0.60 72 Dfs Mangle Mangle

II1A 1.26 0.13 4 1.26 1.38 1.14 78 Mfs Mangle Swamp

II1A 0.64 1 74 Dfs Mangle Dune deposits Old

II1A 0.37 1 74 Dfs Mangle Dune deposits Senescent

II1A 0.29 1 74 Dfs Mangle Dune deposits Yellow

II1A 1.07 1 74 Dfs Mangle Dune deposits Youg Mature

II1Aiiib 1.65 0.12 6 1.58 1.82 1.55 83 Mfs Histosol Alluvial deposits Adult 

II1B 0.18 0.07 3 0.15 0.25 0.13 87 Mfs Entisols Compound dunes

II9A 0.61 0.20 12 0.61 1.12 0.46 73 Dfs Mangle Dune deposits

II9C 0.73 0.11 10 0.73 0.99 0.65 73 Dfs Mangle Dune deposits
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

154

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Phosphorus (mg/g) II1A 0.44 0.09 13 0.42 0.67 0.32 72 Dfs Mangle Mangle

II1A 0.50 1 74 Dfs Mangle Dune deposits Old

II1A 0.22 1 74 Dfs Mangle Dune deposits Senescent

II1A 0.14 1 74 Dfs Mangle Dune deposits Yellow

II1A 0.93 1 74 Dfs Mangle Dune deposits Youg Mature

II1A 1.22 0.47 4 1.21 1.64 0.81 78 Mfs Mangle Swamp

II1Aiiib 1.33 0.05 6 1.31 1.39 1.26 83 Mfs Histosol Alluvial deposits Adult 

II1B 0.35 0.15 3 0.26 0.53 0.26 87 Mfs Entisols Compound dunes

II1B 0.14 0.03 2 0.14 0.12 0.17 157 Mfs Entisols Compound dunes

II9A 0.29 0.22 12 0.29 0.88 0.07 73 Dfs Mangle Dune deposits

II9C 0.75 0.19 10 0.75 1.04 0.43 73 Dfs Mangle Dune deposits

Potassium (mg/g) II1A 4.40 3.20 12 3.00 9.20 1.00 72 Dfs Mangle Mangle

II1A 16.71 1 74 Dfs Mangle Dune deposits Old

II1A 15.97 1 74 Dfs Mangle Dune deposits Senescent

II1A 14.13 1 74 Dfs Mangle Dune deposits Yellow

II1A 11.47 1 74 Dfs Mangle Dune deposits Youg Mature

II1A 20.79 13.72 4 20.57 33.15 8.86 78 Mfs Mangle Swamp

II1Aiiib 9.34 5.53 6 7.01 16.35 4.55 83 Mfs Histosol Alluvial deposits Adult 

II1B 2.31 0.32 3 2.31 2.63 2.00 87 Mfs Entisols Compound dunes

II1B 1.99 0.17 2 1.99 1.86 2.11 157 Mfs Entisols Compound dunes

II9A 5.33 0.98 12 5.33 6.58 3.68 73 Dfs Mangle Dune deposits

II9C 7.97 2.18 10 7.97 11.37 4.87 73 Dfs Mangle Dune deposits

Sodium (mg/g) II1A 49.57 1 74 Dfs Mangle Dune deposits Old

II1A 56.34 1 74 Dfs Mangle Dune deposits Senescent

II1A 52.58 1 74 Dfs Mangle Dune deposits Yellow

II1A 27.47 1 74 Dfs Mangle Dune deposits Youg Mature

II1A 26.37 16.24 4 26.34 40.52 12.28 78 Mfs Mangle Swamp

II1Aiiib 14.15 4.58 6 12.87 19.89 9.62 83 Mfs Histosol Alluvial deposits Adult 

II9A 22.93 6.72 12 22.93 30.63 12.10 73 Dfs Mangle Dune deposits
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

155

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II9C 6.72 1.96 10 6.72 8.77 3.32 73 Dfs Mangle Dune deposits

Sulfur (mg/g) II1Aiiib 0.31 0.08 6 0.31 0.43 0.22 83 Mfs Histosol Alluvial deposits Adult 

Leucaena leucocephala Aluminum (mg/g) II8 3.29 2.56 20 2.11 8.46 0.48 152 Mfs Histosol Alluvial deposits

Ash (%) II1A 9.27 1.19 5 9.87 10.11 7.29 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 1.57 0.31 5 1.55 2.01 1.15 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 7.76 1.42 6 7.24 10.34 6.28 126 Ultisols Siltstone,sandstone Urban forest, RP

II8 33.59 21.40 20 25.62 72.26 8.77 152 Mfs Histosol Alluvial deposits

C/N II1A 12 1 5 12 14 12 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 95 27 5 107 123 60 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 19 3 6 20 21 13 126 Ultisols Siltstone,sandstone Urban forest, RP

II8 22 7 20 20 33 14 152 Mfs Histosol Alluvial deposits

Calcium (mg/g) II8 26.38 13.88 20 23.51 67.87 11.80 152 Mfs Histosol Alluvial deposits

Carbon (%) II1A 51 1 5 51 52 51 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 52 1 5 52 52 51 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 51 2 6 52 53 49 126 Ultisols Siltstone,sandstone Urban forest, RP

II8 38 12 20 42 52 13 152 Mfs Histosol Alluvial deposits

Iron (mg/g) II8 5.23 4.60 20 3.12 15.11 0.57 152 Mfs Histosol Alluvial deposits

Magnesium (mg/g) II8 3.64 0.90 20 3.88 5.11 2.00 152 Mfs Histosol Alluvial deposits

Manganese (mg/g) II8 0.20 0.13 20 0.15 4.61 0.04 152 Mfs Histosol Alluvial deposits

Nitrogen (%) II1A 4.11 0.33 5 4.27 4.37 3.58 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 0.59 0.19 5 0.48 0.85 0.42 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 2.80 0.51 6 2.63 3.81 2.41 126 Ultisols Siltstone,sandstone Urban forest, RP

II8 1.76 0.45 20 1.66 2.45 0.80 148 Smf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II8 0.50 0.17 20 0.53 1.01 0.26 148 Smf Ultisols Tuffac. sandstone

Potassium (mg/g) II8 2.50 0.73 20 2.57 4.61 1.65 148 Smf Ultisols Tuffac. sandstone

Sulfur (%) II1A 0.44 0.03 5 0.44 0.47 0.39 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 0.08 0.02 5 0.09 0.10 0.05 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 0.42 0.16 6 0.46 0.60 0.23 126 Ultisols Siltstone,sandstone Urban forest, RP
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

156

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8 0.42 0.11 2 0.43 0.65 0.20 148 Smf Ultisols Tuffac. sandstone

Wood density (g/cc) II1C 0.69 0.08 5 0.70 0.77 0.60 126 Ultisols Siltstone,sandstone Urban forest, RP

Lonchitis hirsuta Aluminum (mg/g) II6 1.26 0.66 5 1.57 1.77 0.22 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II6 2.94 0.89 5 2.24 4.18 1.97 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II6 0.38 0.40 5 0.30 1.07 0.10 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II6 3.00 0.99 5 2.58 4.76 2.51 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II6 0.06 0.03 5 0.05 0.10 0.04 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II6 1.52 0.54 5 1.33 2.47 1.11 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II6 1.00 0.74 5 0.74 2.32 0.50 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II6 20.60 9.88 5 21.67 33.73 5.92 135 Lmrf Ultisols Tuffac. sandstone

Maesopsis eminii Wood density (g/cc) II1B 0.46 0.04 5 0.44 0.52 0.43 97 Wfs Inceptisols Tuffac. sandstone

Magnolia portorricensis Wood density (g/cc) II1C 0.49 0.02 4 0.49 0.51 0.46 115

Magnolia splendens Aluminum (mg/g) II1A 0.14 0.04 2 0.14 0.12 0.11 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.53 0.04 3 0.05 0.10 0.02 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.05 1 93 Wflm Inceptisols Tuffac. sandstone

Ash (%) II1A 5.85 0.61 2 5.85 6.20 5.15 93 Wflm Inceptisols Tuffac. sandstone

II1A 6.02 1 71 Lmrf Ultisols Tuffac. sandstone

II1B 4.00 0.47 3 3.86 4.53 3.62 93 Wflm Inceptisols Tuffac. sandstone

II1C 1.27 1 93 Wflm Inceptisols Tuffac. sandstone

C/N II1A 33 2 2 33 34 31 93 Wflm Inceptisols Tuffac. sandstone

II1B 57 11 3 52 69 49 93 Wflm Inceptisols Tuffac. sandstone

II1C 245 1 93 Wflm Inceptisols Tuffac. sandstone

Calcium (mg/g) II1A 6.86 1.77 2 7.75 8.01 4.83 93 Wflm Inceptisols Tuffac. sandstone

II1B 2.82 0.77 3 2.90 3.56 2.01 93 Wflm Inceptisols Tuffac. sandstone

II1C 1.27 1 93 Wflm Inceptisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

157

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Carbon (%) II1A 57 1 71 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II1A 0.11 0.00 2 0.11 0.12 0.11 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.06 0.01 3 0.06 0.06 0.04 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.06 1 93 Wflm Inceptisols Tuffac. sandstone

Magnesium (mg/g) II1A 2.37 0.01 2 2.37 2.37 2.36 93 Wflm Inceptisols Tuffac. sandstone

II1B 1.28 0.67 3 1.62 1.71 0.50 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.45 1 93 Wflm Inceptisols Tuffac. sandstone

Manganese (mg/g) II1A 0.35 0.08 2 0.35 0.45 0.31 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.11 0.05 3 0.08 0.17 0.07 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.06 1 93 Wflm Inceptisols Tuffac. sandstone

Nitrogen (%) II1A 1.33 0.09 2 1.33 1.43 1.26 93 Wflm Inceptisols Tuffac. sandstone

II1A 1.60 1 71 Lmrf Ultisols Tuffac. sandstone

II1B 0.80 0.13 3 0.85 0.90 0.65 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.19 1 93 Wflm Inceptisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.74 0.02 2 0.74 0.77 0.73 93 Wflm Inceptisols Tuffac. sandstone

II1B 1.59 0.68 3 1.46 2.32 0.98 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.07 1 93 Wflm Inceptisols Tuffac. sandstone

Potassium (mg/g) II1A 11.75 1.72 2 11.75 12.90 9.77 93 Wflm Inceptisols Tuffac. sandstone

II1B 12.23 1.54 3 12.67 13.51 10.52 93 Wflm Inceptisols Tuffac. sandstone

II1C 1.67 1 93 Wflm Inceptisols Tuffac. sandstone

Sulfur (%) II1A 0.22 1 71 Lmrf Ultisols Tuffac. sandstone

Mahohany Plantation Aluminum (mg/g) II8A 3.03 2.51 11 1.24 7.28 0.58 133 Wfs Ultisols Tuffac. sandstone

II8B 1.80 1.88 9 0.76 4.84 0.43 133 Wfs Ultisols Tuffac. sandstone

II8C 1.13 1.10 11 0.65 3.25 0.22 133 Wfs Ultisols Tuffac. sandstone

II8D 13.07 5.21 9 14.19 20.61 5.18 133 Wfs Ultisols Tuffac. sandstone

Ash (%) II8A 11.58 3.59 11 10.25 17.60 8.04 133 Wfs Ultisols Tuffac. sandstone

II8B 7.38 4.18 9 4.90 13.80 4.01 133 Wfs Ultisols Tuffac. sandstone

II8C 4.28 2.01 11 3.10 7.96 2.68 133 Wfs Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

158

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8D 21.75 6.53 9 22.02 31.42 13.07 133 Wfs Ultisols Tuffac. sandstone

C/N II8A 39 11 11 32 57 26 133 Wfs Ultisols Tuffac. sandstone

II8B 77 26 9 78 124 43 133 Wfs Ultisols Tuffac. sandstone

II8C 63 10 11 61 79 44 133 Wfs Ultisols Tuffac. sandstone

II8D 24 3 9 23 30 20 133 Wfs Ultisols Tuffac. sandstone

Calcium (mg/g) II8A 23.91 4.64 11 22.93 34.62 16.27 133 Wfs Ultisols Tuffac. sandstone

II8B 13.71 2.75 9 13.12 19.37 11.11 133 Wfs Ultisols Tuffac. sandstone

II8C 7.02 1.14 11 7.50 8.50 4.84 133 Wfs Ultisols Tuffac. sandstone

II8D 19.32 2.08 9 19.46 21.96 14.84 133 Wfs Ultisols Tuffac. sandstone

Carbon (%) II8A 49 2 11 50 52 44 133 Wfs Ultisols Tuffac. sandstone

II8B 49 2 9 50 51 46 133 Wfs Ultisols Tuffac. sandstone

II8C 51 2 11 51 53 48 133 Wfs Ultisols Tuffac. sandstone

II8D 42 4 9 42 47 37 133 Wfs Ultisols Tuffac. sandstone

Iron (mg/g) II8A 2.76 2.57 11 0.95 7.50 0.45 133 Wfs Ultisols Tuffac. sandstone

II8B 2.16 2.71 9 0.61 7.21 0.25 133 Wfs Ultisols Tuffac. sandstone

II8C 1.11 1.17 11 0.67 3.47 0.04 133 Wfs Ultisols Tuffac. sandstone

II8D 14.37 5.93 9 16.04 22.28 5.12 133 Wfs Ultisols Tuffac. sandstone

Magnesium (mg/g) II8A 2.56 0.39 11 2.51 3.56 2.20 133 Wfs Ultisols Tuffac. sandstone

II8B 2.12 0.65 9 2.06 3.61 1.52 133 Wfs Ultisols Tuffac. sandstone

II8C 1.75 0.28 11 1.66 2.22 1.39 133 Wfs Ultisols Tuffac. sandstone

II8D 3.07 0.58 9 3.18 4.22 2.06 133 Wfs Ultisols Tuffac. sandstone

Manganese (mg/g) II8A 0.27 0.11 11 0.28 0.43 0.12 133 Wfs Ultisols Tuffac. sandstone

II8B 0.22 0.10 9 0.19 0.38 0.07 133 Wfs Ultisols Tuffac. sandstone

II8C 0.06 0.04 11 0.05 0.13 0.01 133 Wfs Ultisols Tuffac. sandstone

II8D 0.73 0.21 9 0.76 0.98 0.41 133 Wfs Ultisols Tuffac. sandstone

Nitrogen (%) II8A 1.32 0.31 11 1.45 1.72 0.92 133 Wfs Ultisols Tuffac. sandstone

II8B 0.70 0.24 9 0.64 1.14 0.41 133 Wfs Ultisols Tuffac. sandstone

II8C 0.83 0.13 11 0.82 1.14 0.65 133 Wfs Ultisols Tuffac. sandstone

II8D 1.80 0.14 9 1.74 2.01 1.58 133 Wfs Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

159

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Phosphorus (mg/g) II8A 0.41 0.11 11 0.39 0.58 0.27 133 Wfs Ultisols Tuffac. sandstone

II8B 0.28 0.06 9 0.27 0.40 0.18 133 Wfs Ultisols Tuffac. sandstone

II8C 0.43 0.17 11 0.45 0.80 0.26 133 Wfs Ultisols Tuffac. sandstone

II8D 0.62 0.10 9 0.60 0.77 0.45 133 Wfs Ultisols Tuffac. sandstone

Potassium (mg/g) II8A 2.45 1.12 11 2.26 5.18 1.26 133 Wfs Ultisols Tuffac. sandstone

II8B 1.96 0.83 9 2.03 3.36 0.90 133 Wfs Ultisols Tuffac. sandstone

II8C 4.40 2.86 11 2.95 10.67 2.29 133 Wfs Ultisols Tuffac. sandstone

II8D 1.79 0.83 9 1.29 3.23 0.92 133 Wfs Ultisols Tuffac. sandstone

Mangrove Aluminum (mg/kg) II1A 10 10 13 10 20 10 79 Mfs Entisols Swamps Adult 

II1A 90 170 13 20 650 20 79 Mfs Entisols Swamps Dead

II1A 30 20 13 20 70 10 79 Mfs Entisols Swamps Yellow

II1A 30 30 13 20 130 10 79 Mfs Entisols Swamps Young

Ash (%) II1A 9.30 0.48 13 9.44 9.98 8.29 79 Mfs Entisols Swamps Adult 

II1A 13.01 0.91 13 13.22 14.40 11.54 79 Mfs Entisols Swamps Dead

II1A 13.25 1.26 12 13.48 15.28 11.24 79 Mfs Entisols Swamps Yellow

II1A 7.88 0.48 12 7.69 8.66 7.31 79 Mfs Entisols Swamps Young

C/N II1A 39 1 13 40 41 37 79 Mfs Entisols Swamps Adult 

II1A 77 7 13 76 87 66 79 Mfs Entisols Swamps Dead

II1A 73 7 12 73 84 61 79 Mfs Entisols Swamps Yellow

II1A 50 3 12 49 55 46 79 Mfs Entisols Swamps Young

Calcium (mg/g) II1A 8.99 1.43 13 8.39 12.13 7.03 79 Mfs Entisols Swamps Adult 

II1A 11.39 2.57 13 10.66 14.98 8.47 79 Mfs Entisols Swamps Dead

II1A 10.32 2.34 13 9.23 15.67 7.82 79 Mfs Entisols Swamps Yellow

II1A 8.47 1.59 13 7.85 11.94 5.99 79 Mfs Entisols Swamps Young

Carbon (%) II1A 51 0 13 51 51 50 79 Mfs Entisols Swamps Adult 

II1A 49 1 13 49 51 48 79 Mfs Entisols Swamps Dead

II1A 49 1 13 48 51 47 79 Mfs Entisols Swamps Yellow

II1A 51 0 12 51 52 51 79 Mfs Entisols Swamps Young
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

160

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Iron (mg/kg) II1A 20 10 13 20 50 10 79 Mfs Entisols Swamps Adult 

II1A 32 31 12 24 130 14 79 Mfs Entisols Swamps Dead

II1A 50 110 13 20 400 10 79 Mfs Entisols Swamps Yellow

II1A 30 30 13 30 80 10 79 Mfs Entisols Swamps Young

Magnesium (mg/g) II1A 4.76 0.65 13 4.62 6.09 3.94 79 Mfs Entisols Swamps Adult 

II1A 5.99 0.76 13 5.93 7.53 4.99 79 Mfs Entisols Swamps Dead

II1A 5.66 0.95 13 5.38 7.51 4.18 79 Mfs Entisols Swamps Yellow

II1A 4.36 0.71 13 4.06 6.05 3.46 79 Mfs Entisols Swamps Young

Manganese (mg/g) II1A 0.06 0.02 13 0.05 0.09 0.03 79 Mfs Entisols Swamps Adult 

II1A 0.07 0.03 13 0.06 0.14 0.04 79 Mfs Entisols Swamps Dead

II1A 0.06 0.03 13 0.05 0.14 0.04 79 Mfs Entisols Swamps Yellow

II1A 0.05 0.02 13 0.04 0.09 0.03 79 Mfs Entisols Swamps Young

Nitrogen (%) II1A 1.29 0.04 13 1.27 1.38 1.25 79 Mfs Entisols Swamps Adult 

II1A 0.64 0.05 13 0.64 0.73 0.57 79 Mfs Entisols Swamps Dead

II1A 0.68 0.06 12 0.66 0.78 0.61 79 Mfs Entisols Swamps Yellow

II1A 1.03 0.07 12 1.04 1.12 0.92 79 Mfs Entisols Swamps Young

Phosphorus (mg/g) II1A 0.37 0.04 13 0.37 0.41 0.31 79 Mfs Entisols Swamps Adult 

II1A 0.08 0.03 13 0.07 0.13 0.05 79 Mfs Entisols Swamps Dead

II1A 0.16 0.11 13 0.12 0.37 0.07 79 Mfs Entisols Swamps Yellow

II1A 0.35 0.13 13 0.39 0.46 0.07 79 Mfs Entisols Swamps Young

Potassium (mg/g) II1A 5.91 1.05 13 5.93 7.74 3.94 79 Mfs Entisols Swamps Adult 

II1A 7.21 1.23 13 7..32 8.70 4.82 79 Mfs Entisols Swamps Dead

II1A 6.45 1.08 13 6.45 7.90 4.89 79 Mfs Entisols Swamps Yellow

II1A 5.50 0.86 13 5.29 6.70 4.08 79 Mfs Entisols Swamps Young

Sulfur (%) II1A 0.54 0.09 13 0.56 0.76 0.44 79 Mfs Entisols Swamps Adult 

II1A 0.75 0.13 13 0.76 0.97 0.56 79 Mfs Entisols Swamps Dead

II1A 0.76 0.09 12 0.74 0.91 0.64 79 Mfs Entisols Swamps Yellow

II1A 0.26 0.06 12 0.25 0.36 0.19 79 Mfs Entisols Swamps Young
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

161

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Manilkara bidentata Aluminum (mg/g) II1A 0.12 0.05 20 0.12 0.26 0.05 107 Swf Ultisols Tuffac. sandstone

II1A 0.44 1 129 Lmrf Ultisols Tuffac. sandstone

II8A 0.11 0.02 3 0.11 0.13 0.10 145 Wfs Ultisols Tuffac. sandstone

II9A 1.00 0.20 6 0.99 1.29 0.76 153 Swf Ultisols Tuffac. sandstone

Ash (%) II1A 6.85 1 94 Wfs Ultisols Tuffac. sandstone <1cm dbh

II1A 6.71 1 101 Wfs Ultisols Tuffac. sandstone Fresh

II1A 4.94 0.76 17 4.94 6.29 3.72 107 Swf Ultisols Tuffac. sandstone

II1A 4.27 1 129 Lmrf Ultisols Tuffac. sandstone

II1B 3.50 1 94 Wfs Ultisols Tuffac. sandstone <1cm dbh

II1B 3.62 1 68 Swf Ultisols Tuffac. sandstone Wood decay

II1C 3.26 1 94 Wfs Ultisols Tuffac. sandstone <1cm dbh

II8A 4.91 0.23 3 4.90 5.14 4.68 145 Wfs Ultisols Tuffac. sandstone

II8A 5.60 1 71 Wfs Ultisols Tuffac. sandstone

II9A 4.84 0.26 6 4.90 5.10 4.52 153 Swf Ultisols Tuffac. sandstone

C/N II1A 43 4 20 43 48 37 107 Swf Ultisols Tuffac. sandstone

II1A 34 1 129 Lmrf Ultisols Tuffac. sandstone

II8A 90 4 3 90 94 87 145 Wfs Ultisols Tuffac. sandstone

II9A 68 16 6 68 86 43 153 Swf Ultisols Tuffac. sandstone

Calcium (mg/g) II1A 6.05 1.15 7 6.58 7.63 3.73 89 Swf Ultisols Tuffac. sandstone

II1A 5.71 1.51 20 5.22 8.60 3.44 107 Swf Ultisols Tuffac. sandstone

II1A 9.80 1 129 Lmrf Ultisols Tuffac. sandstone

II1B 1.34 1 68 Swf Ultisols Tuffac. sandstone Wood decay

II1C 2.11 0.83 13 1.73 3.55 1.12 89 Swf Ultisols Tuffac. sandstone

II1C 1.24 0.47 24 1.22 2.17 0.48 89 Swf Ultisols Tuffac. sandstone

II1D 11.41 9.32 4 11.23 22.42 0.77 89 Swf Ultisols Tuffac. sandstone

II1F 3.37 1 89 Swf Ultisols Tuffac. sandstone

II8A 8.08 1.05 3 8.48 8.87 6.89 145 Wfs Ultisols Tuffac. sandstone

II9A 8.34 1.45 6 8.16 9.97 6.89 153 Swf Ultisols Tuffac. sandstone

Carbon (%) II1A 49 1 20 49 51 46 107 Swf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

162

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 55 1 129 Lmrf Ultisols Tuffac. sandstone

II1Aiii 45 1 116 45 48 41 125 Wfs Ultisols Tuffac. sandstone

II8A 55 1 3 55 56 54 145 Wfs Ultisols Tuffac. sandstone

II8A 57 1 71 Wfs Ultisols Tuffac. sandstone

II9A 58 1 6 58 58 57 153 Swf Ultisols Tuffac. sandstone

Iron (mg/g) II1A 0.11 0.03 20 0.11 0.15 0.06 107 Swf Ultisols Tuffac. sandstone

II1A 0.42 1 129 Lmrf Ultisols Tuffac. sandstone

II8A 0.11 0.01 3 0.11 0.12 0.09 145 Wfs Ultisols Tuffac. sandstone

II9A 0.97 0.20 6 0.95 1.30 0.75 153 Swf Ultisols Tuffac. sandstone

Magnesium (mg/g) II1A 3.29 0.75 7 3.54 3.99 1.89 89 Swf Ultisols Tuffac. sandstone

II1A 2.81 0.58 20 2.76 4.10 1.89 107 Swf Ultisols Tuffac. sandstone

II1A 1.89 1 129 Lmrf Ultisols Tuffac. sandstone

II1C 0.68 0.34 13 0.71 1.44 0.31 89 Swf Ultisols Tuffac. sandstone

II1C 0.27 0.05 24 0.28 0.37 0.18 89 Swf Ultisols Tuffac. sandstone

II1D 3.22 2.01 4 4.18 4.33 0.21 89 Swf Ultisols Tuffac. sandstone

II1F 2.17 1 89 Swf Ultisols Tuffac. sandstone

II8A 2.37 0.69 3 2.12 3.15 1.83 145 Wfs Ultisols Tuffac. sandstone

II9A 3.22 0.45 6 3.32 3.67 2.40 153 Swf Ultisols Tuffac. sandstone

Manganese (mg/g) II1A 0.03 0.01 20 0.03 0.05 0.02 107 Swf Ultisols Tuffac. sandstone

II1A 0.04 1 129 Lmrf Ultisols Tuffac. sandstone

II8A 0.03 0.02 3 0.02 0.05 0.02 145 Wfs Ultisols Tuffac. sandstone

II9A 0.04 0.01 6 0.04 0.05 0.03 153 Swf Ultisols Tuffac. sandstone

Nitrogen (%) II1A 1.46 1 94 Wfs Ultisols Tuffac. sandstone <1cm dbh

II1A 1.03 1 101 Wfs Ultisols Tuffac. sandstone Fresh

II1A 1.15 0.12 20 1.13 1.37 0.99 107 Swf Ultisols Tuffac. sandstone

II1A 1.63 1 129 Lmrf Ultisols Tuffac. sandstone

II1Aiii 1.18 0.30 116 1.21 2.01 0.65 125 Wfs Ultisols Tuffac. sandstone

II1B 1.05 1 94 Wfs Ultisols Tuffac. sandstone <1cm dbh

II1B 0.57 1 68 Swf Ultisols Tuffac. sandstone Wood decay
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

163

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1C 1.05 1 94 Wfs Ultisols Tuffac. sandstone <1cm dbh

II8A 0.61 0.03 3 0.62 0.64 0.58 145 Wfs Ultisols Tuffac. sandstone

II8A 0.67 1 71 Wfs Ultisols Tuffac. sandstone

II9A 0.89 0.25 6 0.85 1.35 0.67 153 Swf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.46 0.11 7 0.50 0.61 0.30 89 Swf Ultisols Tuffac. sandstone

II1A 0.58 1 94 Wfs Ultisols Tuffac. sandstone <1cm dbh

II1A 0.49 1 101 Wfs Ultisols Tuffac. sandstone Fresh

II1A 0.47 0.06 20 0.45 0.59 0.39 107 Swf Ultisols Tuffac. sandstone

II1A 0.24 1 129 Lmrf Ultisols Tuffac. sandstone

II1B 0.32 1 94 Wfs Ultisols Tuffac. sandstone <1cm dbh

II1B 0.15 1 68 Swf Ultisols Tuffac. sandstone Wood decay

II1C 0.16 0.08 13 0.14 0.30 0.07 89 Swf Ultisols Tuffac. sandstone

II1C 0.37 1 94 Wfs Ultisols Tuffac. sandstone <1cm dbh

II1C 0.05 0.02 24 0.05 0.10 0.01 89 Swf Ultisols Tuffac. sandstone

II1D 0.68 0.48 4 0.75 1.18 0.02 89 Swf Ultisols Tuffac. sandstone

II1F 0.78 1 89 Swf Ultisols Tuffac. sandstone

II8A 0.12 0.03 3 0.10 0.16 0.10 145 Wfs Ultisols Tuffac. sandstone

II9A 0.16 0.11 6 0.13 0.37 0.08 153 Swf Ultisols Tuffac. sandstone

Potassium (mg/g) II1A 5.86 2.05 7 5.86 8.30 2.03 89 Swf Ultisols Tuffac. sandstone

II1A 6.70 1 94 Wfs Ultisols Tuffac. sandstone <1cm dbh

II1A 9.30 1 101 Wfs Ultisols Tuffac. sandstone Fresh

II1A 7.72 2.94 20 8.42 12.10 3.57 107 Swf Ultisols Tuffac. sandstone

II1A 4.16 1 129 Lmrf Ultisols Tuffac. sandstone

II1B 5.04 1 94 Wfs Ultisols Tuffac. sandstone <1cm dbh

II1B 1.21 1 68 Swf Ultisols Tuffac. sandstone Class 4 LD

II1C 1.69 0.69 13 2.01 2.46 0.69 89 Swf Ultisols Tuffac. sandstone

II1C 2.84 1 94 Wfs Ultisols Tuffac. sandstone <1cm dbh

II1C 1.08 0.48 24 0.94 3.04 0.71 89 Swf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

164

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1D 6.56 2.23 4 6.64 8.53 4.44 89 Swf Ultisols Tuffac. sandstone

II1F 11.68 1 89 Swf Ultisols Tuffac. sandstone

II8A 3.79 0.19 3 3.84 3.95 3.59 145 Wfs Ultisols Tuffac. sandstone

II9A 3.60 1.41 6 3.15 6.36 2.59 153 Swf Ultisols Tuffac. sandstone

Sodium (mg/g) II9A 2.69 0.22 6 2.68 3.01 2.39 153 Swf Ultisols Tuffac. sandstone

Sulfur (%) II1A 0.64 1 129 Lmrf Ultisols Tuffac. sandstone

II8A 0.31 0.03 3 0.32 0.34 0.28 145 Wfs Ultisols Tuffac. sandstone

II8A 0.37 1 71 Wfs Ultisols Tuffac. sandstone

Marcgravia retiflora Aluminum (mg/g) II4 0.38 1 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II4 4.01 1 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II4 47 3 13 48 49 41 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II4 0.29 1 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II4 1.54 1 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II4 0.06 1 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II4 0.88 1 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II4 15.85 1 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II4 0.30 0.14 13 0.28 0.57 0.12 135 Lmrf Ultisols Tuffac. sandstone

Marcgravia spp. Aluminum (mg/g) II4 1.01 1.70 20 0.36 6.98 0.04 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II4 4.53 2.23 20 4.06 9.08 1.24 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II4 1.10 2.10 20 0.31 8.85 0.07 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II4 2.53 1.78 20 1.85 7.93 0.79 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II4 0.14 0.12 20 0.21 0.51 0.02 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II4 0.91 0.20 20 0.90 1.25 0.50 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II4 0.85 0.31 20 0.84 1.60 0.40 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II4 16.08 5.53 20 16.41 32.58 9.75 135 Lmrf Ultisols Tuffac. sandstone

Matayba dominguensis Ash (%) II1B 2.00 1 68 Swf Ultisols Tuffac. sandstone Wood decay
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

165

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Calcium (mg/g) II1B 5.64 1 68 Swf Ultisols Tuffac. sandstone Wood decay

Nitrogen (%) II1B 0.27 1 68 Swf Ultisols Tuffac. sandstone Wood decay

Phosphorus (mg/g) II1B 0.12 1 68 Swf Ultisols Tuffac. sandstone Wood decay

Potassium (mg/g) II1B 1.99 1 68 Swf Ultisols Tuffac. sandstone Wood decay

Megalastrum subincisa Aluminum (mg/g) II3 1.89 0.88 5 1.63 2.91 1.04 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II3 4.81 4.27 5 3.86 11.36 1.10 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II3 1.39 1.82 5 0.09 3.63 0.06 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II3 3.01 1.10 5 3.40 4.22 1.58 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II3 0.08 0.06 5 0.06 0.17 0.03 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II3 1.53 0.88 5 1.11 2.91 0.86 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II3 0.98 0.72 5 0.64 1.98 0.38 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II3 17.57 12.79 5 15.71 35.97 3.81 135 Lmrf Ultisols Tuffac. sandstone

Meliccocus bijugatus Wood density (g/cc) II1B 0.79 0.02 3 0.80 0.81 0.77 97 Wfs Inceptisols Tuffac. sandstone

Miconia impetiolaris Aluminum (mg/g) II2A 20.20 1 129 Lmrf Ultisols Tuffac. sandstone

Ash (%) II2A 15.98 1 129 Lmrf Ultisols Tuffac. sandstone

C/N II2A 24 1 129 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II2A 18.33 1 129 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II2A 46 1 129 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II2A 5.58 1 129 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II2A 2.26 1 129 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II2A 0.26 1 129 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II2A 1.92 1 129 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II2A 0.84 1 129 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II2A 3.72 1 129 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II2A 0.34 1 129 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

166

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Miconia prasina Aluminum (mg/g) II2A 25.83 1 131 Wfs Mollisols Plutonic rocks

II2A 10.58 1 131 Wfs Mollisols Plutonic rocks

II2A 8.57 1 131 Wfs Mollisols Plutonic rocks

II2A 13.19 1 131 Wfs Mollisols Plutonic rocks

Ash (%) II2A 11.50 2.73 7 11.68 16.07 8.25 94 Swf Ultisols Tuffac. sandstone

II2A 10.28 1.19 6 10.99 11.11 8.52 126 Ultisols Siltstone,sandstone Urban forest, RP

II2A 10.99 1 131 Wfs Mollisols Plutonic rocks

II2A 9.23 2.35 54 8.77 14.40 5.86 138 Wfs Ultisols Lava

II2A 7.05 0.95 6 7.05 7.94 5.28 94 Swf Ultisols Tuffac. sandstone

II2A 5.97 2.67 4 6.26 8.82 2.56 68 Swf Ultisols Tuffac. sandstone Wood decay

II2A 5.63 1 131 Wfs Mollisols Plutonic rocks

II2A 4.57 1.32 7 4.60 6.29 2.98 94 Swf Ultisols Tuffac. sandstone

II2A 3.72 1.28 6 4.19 5.13 2.02 126 Ultisols Siltstone,sandstone Urban forest, RP

II2A 4.61 1 131 Wfs Mollisols Plutonic rocks

II2A 12.52 1.30 5 11.92 14.22 11.29 126 Ultisols Siltstone,sandstone Urban forest, RP

II2A 4.55 1 131 Wfs Mollisols Plutonic rocks

II2A 12.12 0.42 2 12.12 12.42 11.82 94 Swf Ultisols Tuffac. sandstone

C/N II2A 26 1 6 26 28 25 126 Ultisols Siltstone,sandstone Urban forest, RP

II2A 23 1 131 Wfs Mollisols Plutonic rocks

II2A 49 1 131 Wfs Mollisols Plutonic rocks

II2A 203 100 6 159 339 119 126 Ultisols Siltstone,sandstone Urban forest, RP

II2A 61 1 131 Wfs Mollisols Plutonic rocks

II2A 38 17 5 30 65 23 126 Ultisols Siltstone,sandstone Urban forest, RP

II2A 74 1 131 Wfs Mollisols Plutonic rocks

Calcium (mg/g) II2A 12.40 1 131 Wfs Mollisols Plutonic rocks

II2A 3.28 0.80 4 3.44 4.02 2.24 68 Swf Ultisols Tuffac. sandstone Wood decay

II2A 8.00 1 131 Wfs Mollisols Plutonic rocks

II2A 4.48 1 131 Wfs Mollisols Plutonic rocks

II2A 2.10 1 131 Wfs Mollisols Plutonic rocks
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

167

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Carbon (%) II2A 50 1 6 50 52 49 126 Ultisols Siltstone,sandstone Urban forest, RP

II2A 48 1 131 Wfs Mollisols Plutonic rocks

II2A 49 3 54 49 53 44 138 Wfs Ultisols Lava

II2A 50 1 131 Wfs Mollisols Plutonic rocks

II2A 50 0 6 50 51 50 126 Ultisols Siltstone,sandstone Urban forest, RP

II2A 50 1 131 Wfs Mollisols Plutonic rocks

II2A 46 3 5 47 49 43 126 Ultisols Siltstone,sandstone Urban forest, RP

II2A 50 1 131 Wfs Mollisols Plutonic rocks

Iron (mg/g) II2A 0.10 1 131 Wfs Mollisols Plutonic rocks

II2A 0.09 1 131 Wfs Mollisols Plutonic rocks

II2A 0.06 1 131 Wfs Mollisols Plutonic rocks

II2A 0.29 1 131 Wfs Mollisols Plutonic rocks

Magnesium (mg/g) II2A 1.48 1 131 Wfs Mollisols Plutonic rocks

II2A 0.79 1 131 Wfs Mollisols Plutonic rocks

II2A 0.46 1 131 Wfs Mollisols Plutonic rocks

II2A 0.45 1 131 Wfs Mollisols Plutonic rocks

Manganese (mg/g) II2A 0.29 1 131 Wfs Mollisols Plutonic rocks

II2A 0.81 1 131 Wfs Mollisols Plutonic rocks

II2A 0.65 1 131 Wfs Mollisols Plutonic rocks

II2A 0.52 1 131 Wfs Mollisols Plutonic rocks

Nitrogen (%) II2A 2.12 1 131 Wfs Mollisols Plutonic rocks

II2A 2.03 0.41 7 2.26 2.42 1.44 94 Swf Ultisols Tuffac. sandstone

II2A 1.91 0.11 6 1.93 2.04 1.77 126 Ultisols Siltstone,sandstone Urban forest, RP

II2A 1.68 0.30 54 1.68 2.29 1.01 138 Wfs Ultisols Lava

II2A 1.02 1 131 Wfs Mollisols Plutonic rocks

II2A 0.82 0.19 6 0.82 1.13 0.58 94 Swf Ultisols Tuffac. sandstone

II2A 0.41 0.17 4 0.44 0.57 0.20 68 Swf Ultisols Tuffac. sandstone Wood decay

II2A 0.81 1 131 Wfs Mollisols Plutonic rocks

II2A 0.45 0.29 7 0.28 0.83 0.18 94 Swf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

168

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II2A 0.30 0.12 6 0.33 0.42 0.15 126 Ultisols Siltstone,sandstone Urban forest, RP

II2A 1.44 0.61 5 1.63 2.10 0.67 126 Ultisols Siltstone,sandstone Urban forest, RP

II2A 0.68 1 131 Wfs Mollisols Plutonic rocks

II2A 1.54 0.01 2 l.54 1.55 1.53 94 Swf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II2A 1.06 0.35 7 0.82 1.49 0.65 94 Swf Ultisols Tuffac. sandstone

II2A 0.45 1 131 Wfs Mollisols Plutonic rocks

II2A 0.56 0.34 6 0.56 1.23 0.30 94 Swf Ultisols Tuffac. sandstone

II2A 0.16 0.10 4 0.17 0.26 0.03 68 Swf Ultisols Tuffac. sandstone Wood decay

II2A 0.19 1 131 Wfs Mollisols Plutonic rocks

II2A 0.36 0.20 7 0.36 0.66 0.16 94 Swf Ultisols Tuffac. sandstone

II2A 0.12 1 131 Wfs Mollisols Plutonic rocks

II2A 0.12 1 131 Wfs Mollisols Plutonic rocks

II2A 1.26 0.34 2 1.26 1.50 1.02 94 Swf Ultisols Tuffac. sandstone

Potassium (mg/g) II2A 7.43 2.30 7 7.10 11.93 5.12 94 Swf Ultisols Tuffac. sandstone

II2A 4.69 1 131 Wfs Mollisols Plutonic rocks

II2A 5.31 1.15 6 0.53 0.69 0.37 94 Swf Ultisols Tuffac. sandstone

II2A 1.24 0.21 4 1.31 1.35 0.94 68 Swf Ultisols Tuffac. sandstone Wood decay

II2A 3.90 1 131 Wfs Mollisols Plutonic rocks

II2A 4.60 4.36 7 3.32 13.96 1.09 94 Swf Ultisols Tuffac. sandstone

II2A 3.39 1 131 Wfs Mollisols Plutonic rocks

II2A 3.14 1 131 Wfs Mollisols Plutonic rocks

II2A 9.23 3.88 2 9.23 1.20 0.65 94 Swf Ultisols Tuffac. sandstone

Sulfur (%) II2A 0.19 0.01 6 0.19 0.20 0.18 126 Ultisols Siltstone,sandstone Urban forest, RP

II2A 0.48 1 131 Wfs Mollisols Plutonic rocks

II2A 0.16 1 131 Wfs Mollisols Plutonic rocks

II2A 0.04 0.01 6 0.04 0.07 0.04 126 Ultisols Siltstone,sandstone Urban forest, RP

II2A 0.14 1 131 Wfs Mollisols Plutonic rocks

II2A 0.20 0.11 5 0.20 0.31 0.08 126 Ultisols Siltstone,sandstone Urban forest, RP

II2A 0.21 1 131 Wfs Mollisols Plutonic rocks
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

169

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Wood density (g/cc) II2A 0.63 0.05 5 0.62 0.68 0.58 126 Ultisols Siltstone,sandstone Urban forest, RP

Miconia racemosa Aluminum (mg/g) II2A 20.61 1 129 Lmrf Ultisols Tuffac. sandstone

II2A 17.01 2.04 2 17.01 18.46 15.57 146 Swf Ultisols Tuffac. sandstone

II2A 2.21 0.70 3 1.83 3.02 1.79 146 Swf Ultisols Tuffac. sandstone

Ash (%) II2A 16.49 1 129 Lmrf Ultisols Tuffac. sandstone

II2A 13.39 0.61 2 13.39 13.82 12.96 146 Swf Ultisols Tuffac. sandstone

II2A 3.53 1.05 3 4.10 4.17 2.32 146 Swf Ultisols Tuffac. sandstone

C/N II2A 23 1 129 Lmrf Ultisols Tuffac. sandstone

II2A 22 4 2 22 25 19 146 Swf Ultisols Tuffac. sandstone

II2A 85 27 3 70 116 69 146 Swf Ultisols Tuffac. sandstone

Calcium (mg/g) II2A 18.47 1 129 Lmrf Ultisols Tuffac. sandstone

II2A 40.32 4.47 2 40.32 43.49 37.16 146 Swf Ultisols Tuffac. sandstone

II2A 6.62 0.68 3 6.28 7.41 6.19 146 Swf Ultisols Tuffac. sandstone

Carbon (%) II2A 45 1 129 Lmrf Ultisols Tuffac. sandstone

II2A 47 3 2 47 49 44 146 Swf Ultisols Tuffac. sandstone

II2A 52 0 3 51 52 51 146 Swf Ultisols Tuffac. sandstone

Iron (mg/g) II2A 4.96 1 129 Lmrf Ultisols Tuffac. sandstone

II2A 0.47 0.43 2 0.47 0.77 0.16 146 Swf Ultisols Tuffac. sandstone

II2A 0.05 0.01 3 0.05 0.61 0.04 146 Swf Ultisols Tuffac. sandstone

Magnesium (mg/g) II2A 1.75 1 129 Lmrf Ultisols Tuffac. sandstone

II2A 1.93 0.61 2 1.93 2.36 1.51 146 Swf Ultisols Tuffac. sandstone

II2A 1.93 0.08 3 1.94 2.01 1.86 146 Swf Ultisols Tuffac. sandstone

Manganese (mg/g) II2A 0.39 1 129 Lmrf Ultisols Tuffac. sandstone

II2A 0.10 0.01 2 0.10 0.11 0.09 146 Swf Ultisols Tuffac. sandstone

II2A 0.05 0.04 3 0.03 0.09 0.03 146 Swf Ultisols Tuffac. sandstone

Nitrogen (%) II2A 1.94 1 129 Lmrf Ultisols Tuffac. sandstone

II2A 2.18 0.59 2 2.18 2.60 1.76 146 Swf Ultisols Tuffac. sandstone

II2A 0.65 0.17 3 0.74 0.75 0.45 146 Swf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

170

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Phosphorus (mg/g) II2A 0.53 1 129 Lmrf Ultisols Tuffac. sandstone

II2A 0.75 0.21 2 0.75 0.89 0.60 146 Swf Ultisols Tuffac. sandstone

II2A 0.44 0.13 3 0.51 0.52 0.29 146 Swf Ultisols Tuffac. sandstone

Potassium (mg/g) II2A 3.51 1 129 Lmrf Ultisols Tuffac. sandstone

II2A 4.63 2.68 2 4.63 6.53 2.73 146 Swf Ultisols Tuffac. sandstone

II2A 4.30 2.50 3 5.57 5.90 1.41 146 Swf Ultisols Tuffac. sandstone

Sulfur (%) II2A 0.25 1 129 Lmrf Ultisols Tuffac. sandstone

Miconia sp. Aluminum (mg/g) II2A 32.76 7.05 30 31.72 46.00 21.65 146 Swf Ultisols Tuffac. sandstone New

II2A 34.80 8.22 35 36.19 52.82 22.31 146 Swf Ultisols Tuffac. sandstone Old

Ash (%) II2A 16.81 4.35 29 16.28 34.46 10.14 146 Swf Ultisols Tuffac. sandstone New

II2A 17.89 2.86 35 17.70 26.84 12.79 146 Swf Ultisols Tuffac. sandstone Old

Calcium (mg/g) II2A 27.37 7.85 30 23.38 43.42 18.08 146 Swf Ultisols Tuffac. sandstone New

II2A 28.30 5.04 35 28.95 39.91 18.17 146 Swf Ultisols Tuffac. sandstone Old

Iron (mg/g) II2A 2.70 4.21 30 1.23 21.14 0.19 146 Swf Ultisols Tuffac. sandstone New

II2A 4.15 3.46 35 3.14 15.62 1.10 146 Swf Ultisols Tuffac. sandstone Old

Magnesium (mg/g) II2A 3.68 1.09 30 3.35 6.66 2.36 146 Swf Ultisols Tuffac. sandstone New

II2A 2.54 0.47 35 2.71 3.80 1.64 146 Swf Ultisols Tuffac. sandstone Old

Manganese (mg/g) II2A 2.10 1.18 30 1.80 4.20 0.73 146 Swf Ultisols Tuffac. sandstone New

II2A 1.16 0.31 35 1.09 1.78 0.53 146 Swf Ultisols Tuffac. sandstone Old

Phosphorus (mg/g) II2A 0.45 0.17 30 0.46 0.81 0.19 146 Swf Ultisols Tuffac. sandstone New

II2A 0.51 0.11 35 0.51 0.86 0.31 146 Swf Ultisols Tuffac. sandstone Old

Potassium (mg/g) II2A 3.21 0.77 30 3.30 4.59 1.95 146 Swf Ultisols Tuffac. sandstone New

II2A 3.14 0.67 35 3.12 4.23 1.75 146 Swf Ultisols Tuffac. sandstone Old

Miconia spp. Aluminum (mg/g) II2A 9.51 7.46 5 6.23 22.81 5.62 143 Wflm Inceptisols Tuffac. sandstone

II2A 4.65 3.78 6 3.25 12.27 2.57 143 Wflm Inceptisols Tuffac. sandstone <1cm

II2A 5.69 2.53 4 6.59 7.55 2.04 143 Wflm Inceptisols Tuffac. sandstone >1cm

Ash (%) II2A 5.96 2.47 5 4.80 10.37 0.68 143 Wflm Inceptisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

171

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II2A 3.27 1.19 6 2.98 5.52 2.35 143 Wflm Inceptisols Tuffac. sandstone <1cm

II2A 2.07 0.72 4 2.34 2.59 1.02 143 Wflm Inceptisols Tuffac. sandstone >1cm

C/N II2A 53 5 5 53 59 44 143 Wflm Inceptisols Tuffac. sandstone

II2A 94 13 6 98 105 72 143 Wflm Inceptisols Tuffac. sandstone <1cm

II2A 121 25 4 121 151 92 143 Wflm Inceptisols Tuffac. sandstone >1cm

Calcium (mg/g) II2A 7.97 5.06 5 5.77 16.95 4.95 143 Wflm Inceptisols Tuffac. sandstone

II2A 4.44 2.78 6 3.57 9.52 2.31 143 Wflm Inceptisols Tuffac. sandstone <1cm

II2A 1.92 0.81 4 2.04 2.70 0.88 143 Wflm Inceptisols Tuffac. sandstone >1cm

Carbon (%) II2A 50 2 5 51 51 46 143 Wflm Inceptisols Tuffac. sandstone

II2A 50 1 6 50 50 48 143 Wflm Inceptisols Tuffac. sandstone <1cm

II2A 50 1 4 50 51 49 143 Wflm Inceptisols Tuffac. sandstone >1cm

Iron (mg/g) II2A 0.28 0.22 5 0.24 0.65 0.07 143 Wflm Inceptisols Tuffac. sandstone

II2A 0.14 0.34 6 0.08 0.91 0.02 143 Wflm Inceptisols Tuffac. sandstone <1cm

II2A 0.92 0.99 4 0.81 2.05 0.02 143 Wflm Inceptisols Tuffac. sandstone >1cm

Magnesium (mg/g) II2A 2.32 1.21 5 1.79 4.47 1.64 143 Wflm Inceptisols Tuffac. sandstone

II2A 1.10 0.68 6 0.82 2.46 0.68 143 Wflm Inceptisols Tuffac. sandstone <1cm

II2A 0.35 0.12 4 0.36 0.46 0.20 143 Wflm Inceptisols Tuffac. sandstone >1cm

Manganese (mg/g) II2A 0.18 0.09 5 0.15 0.34 0.12 143 Wflm Inceptisols Tuffac. sandstone

II2A 0.11 0.10 6 0.06 0.29 0.05 143 Wflm Inceptisols Tuffac. sandstone <1cm

II2A 0.09 0.05 4 0.10 0.14 0.03 143 Wflm Inceptisols Tuffac. sandstone >1cm

Nitrogen (%) II2A 0.95 0.06 5 0.93 1.03 0.86 143 Wflm Inceptisols Tuffac. sandstone

II2A 0.54 0.08 6 0.51 0.69 0.48 143 Wflm Inceptisols Tuffac. sandstone <1cm

II2A 0.43 0.09 4 0.42 0.54 0.33 143 Wflm Inceptisols Tuffac. sandstone >1cm

Phosphorus (mg/g) II2A 0.32 0.05 5 0.32 0.39 0.25 143 Wflm Inceptisols Tuffac. sandstone

II2A 0.17 0.03 6 0.15 0.23 0.14 143 Wflm Inceptisols Tuffac. sandstone <1cm

II2A 0.11 0.03 4 0.11 0.15 0.07 143 Wflm Inceptisols Tuffac. sandstone >1cm

Potassium (mg/g) II2A 4.87 0.95 5 4.87 5.89 3.69 143 Wflm Inceptisols Tuffac. sandstone

II2A 3.31 1.40 6 2.90 6.01 2.09 143 Wflm Inceptisols Tuffac. sandstone <1cm

II2A 2.52 1.21 4 2.50 3.77 1.32 143 Wflm Inceptisols Tuffac. sandstone >1cm
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

172

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Miconia tetranda Aluminum (mg/g) II1B 13.61 1 93 Smf Ultisols Tuffac. sandstone

Ash (%) II1A 9.67 0.31 2 9.67 9.89 9.45 94 Swf Ultisols Tuffac. sandstone

II1B 8.88 1 93 Smf Ultisols Tuffac. sandstone

II1B 6.47 0.83 2 6.47 7.05 5.88 94 Swf Ultisols Tuffac. sandstone

II1C 5.52 0.13 2 5.52 5.61 5.43 94 Swf Ultisols Tuffac. sandstone

II1F 11.86 1 94 Swf Ultisols Tuffac. sandstone

C/N II1B 47 1 93 Smf Ultisols Tuffac. sandstone

Calcium (mg/g) II1B 15.26 1 93 Smf Ultisols Tuffac. sandstone

Iron (mg/g) II1B 0.07 1 93 Smf Ultisols Tuffac. sandstone

Magnesium (mg/g) II1B 1.88 1 93 Smf Ultisols Tuffac. sandstone

Manganese (mg/g) II1B 0.44 1 93 Smf Ultisols Tuffac. sandstone

Nitrogen (%) II1A 1.46 0.03 2 1.46 1.48 1.44 94 Swf Ultisols Tuffac. sandstone

II1B 0.90 1 93 Smf Ultisols Tuffac. sandstone

II1B 0.67 0.02 2 0.67 0.68 0.65 94 Swf Ultisols Tuffac. sandstone

II1C 0.37 0.08 2 0.37 0.42 0.31 94 Swf Ultisols Tuffac. sandstone

II1F 1.54 1 94 Swf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.58 0.05 2 0.58 0.61 0.54 94 Swf Ultisols Tuffac. sandstone

II1B 0.63 1 93 Smf Ultisols Tuffac. sandstone

II1B 0.32 0.01 2 0.32 0.33 0.32 94 Swf Ultisols Tuffac. sandstone

II1C 0.22 0.06 2 0.22 0.27 0.18 94 Swf Ultisols Tuffac. sandstone

II1F 1.24 1 94 Swf Ultisols Tuffac. sandstone

Potassium (mg/g) II1A 4.00 0.11 2 4.00 4.08 3.92 94 Swf Ultisols Tuffac. sandstone

II1B 14.68 1 93 Smf Ultisols Tuffac. sandstone

II1B 3.48 0.00 2 3.48 3.48 3.48 94 Swf Ultisols Tuffac. sandstone

II1C 2.67 0.00 2 2.67 2.67 2.67 94 Swf Ultisols Tuffac. sandstone

II1F 4.49 1 94 Swf Ultisols Tuffac. sandstone

Micropholis chrysophylloides Aluminum (mg/g) II1A 0.10 0.07 4 0.10 0.21 0.05 93 Smf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

173

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1B 0.08 1 93 Smf Ultisols Tuffac. sandstone

II1C 0.19 1 93 Smf Ultisols Tuffac. sandstone

Ash (%) II1A 2.68 0.32 4 2.68 3.16 2.47 93 Smf Ultisols Tuffac. sandstone

II1B 3.08 1 93 Smf Ultisols Tuffac. sandstone

II1C 0.53 1 93 Smf Ultisols Tuffac. sandstone

C/N II1A 45 5 4 45 50 38 93 Smf Ultisols Tuffac. sandstone

II1B 70 1 93 Smf Ultisols Tuffac. sandstone

II1C 184 1 93 Smf Ultisols Tuffac. sandstone

Calcium (mg/g) II1A 4.27 0.75 4 4.27 5.29 3.62 93 Smf Ultisols Tuffac. sandstone

II1B 10.23 1 93 Smf Ultisols Tuffac. sandstone

II1C 0.87 1 93 Smf Ultisols Tuffac. sandstone

Iron (mg/g) II1A 0.10 0.05 4 0.10 0.17 0.06 93 Smf Ultisols Tuffac. sandstone

II1B 0.10 1 93 Smf Ultisols Tuffac. sandstone

II1C 0.27 1 93 Smf Ultisols Tuffac. sandstone

Magnesium (mg/g) II1A 1.4 0.18 4 1.40 1.66 1.25 93 Smf Ultisols Tuffac. sandstone

II1A 0.14 0.12 4 0.14 0.28 0.03 93 Smf Ultisols Tuffac. sandstone

II1B 0.61 1 93 Smf Ultisols Tuffac. sandstone

II1C 0.13 1 93 Smf Ultisols Tuffac. sandstone

II1C 0.05 1 93 Smf Ultisols Tuffac. sandstone

Nitrogen (%) II1A 1.01 0.12 4 1.01 1.18 0.09 93 Smf Ultisols Tuffac. sandstone

II1B 0.64 1 93 Smf Ultisols Tuffac. sandstone

II1C 0.25 1 93 Smf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.39 0.08 4 0.39 0.48 0.31 93 Smf Ultisols Tuffac. sandstone

II1B 0.34 1 93 Smf Ultisols Tuffac. sandstone

II1C 0.12 1 93 Smf Ultisols Tuffac. sandstone

Potassium (mg/g) II1A 5.88 1.32 4 5.88 7.85 5.07 93 Smf Ultisols Tuffac. sandstone

II1B 4.1 1 93 Smf Ultisols Tuffac. sandstone

II1C 1.05 1 93 Smf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

174

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Micropholis garcinifolia Aluminum (mg/g) II1B 0.07 0.05 3 0.10 0.11 0.01 93 Smf Ultisols Tuffac. sandstone

Ash (%) II1B 2.16 1.04 3 2.67 2.84 0.96 93 Smf Ultisols Tuffac. sandstone

C/N II1B 87 69 3 49 167 45 93 Smf Ultisols Tuffac. sandstone

Calcium (mg/g) II1B 4.26 2.09 3 3.71 6.56 2.49 93 Smf Ultisols Tuffac. sandstone

Iron (mg/kg) II1B 70 60 3 100 110 4 93 Smf Ultisols Tuffac. sandstone

Magnesium (mg/g) II1B 0.79 0.29 3 0.74 1.11 0.54 93 Smf Ultisols Tuffac. sandstone

Manganese (mg/g) II1B 0.02 0.01 3 0.02 0.03 0.01 93 Smf Ultisols Tuffac. sandstone

Nitrogen (%) II1B 0.73 0.40 3 0.93 0.99 0.27 93 Smf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II1B 0.34 0.24 3 0.33 0.58 0.09 93 Smf Ultisols Tuffac. sandstone

Potassium (mg/g) II1B 3.82 2.43 3 4.59 5.78 1.10 93 Smf Ultisols Tuffac. sandstone

Mikania cordifolia Aluminum (mg/g) II4 0.06 1 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II4 3.61 1 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II4 0.23 1 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II4 2.31 1 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II4 19.34 1 135 Lmrf Ultisols Tuffac. sandstone

Mixed Species Guanica Forest Ash (%) II1A 16.67 3.16 7 16.97 21.57 12.69 99 Sdf Mollisols Alluvial deposits Decomp. cut plot

II1A 19.58 3.20 2 19.58 21.84 17.32 99 Sdf Mollisols Alluvial deposits Decomp.

II1Ai 13.72 3.90 17 13.11 19.97 8.61 99 Sdf Mollisols Alluvial deposits Canopy

II1C 7.14 2.34 10 7.49 10.95 2.22 99 Sdf Mollisols Alluvial deposits Canopy

II1F 5.36 1 99 Sdf Mollisols Alluvial deposits Canopy

II2 8.75 2.59 4 7.89 12.54 6.70 99 Sdf Mollisols Alluvial deposits Decomp. control 

II8A 12.62 0.30 2 12.62 12.83 12.40 99 Sdf Mollisols Alluvial deposits Decomp.

II8A 13.11 1.26 2 13.11 14.00 12.22 99 Sdf Mollisols Alluvial deposits Control plot

II8C 5.71 0.94 22 5.83 7.48 4.22 99 Sdf Mollisols Alluvial deposits Cut plot

II9A 9.76 0.83 9 9.73 11.05 8.65 99 Sdf Mollisols Alluvial deposits Control plot

II9A 10.08 1.33 10 9.91 12.56 8.22 99 Sdf Mollisols Alluvial deposits Cut plot

II9B 8.28 1.62 10 8.66 10.58 6.06 99 Sdf Mollisols Alluvial deposits Control plot
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

175

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II9B 10.43 4.55 8 10.52 15.75 4.89 99 Sdf Mollisols Alluvial deposits Cut plot

II9C 4.94 0.90 6 5.24 6.02 3.58 99 Sdf Mollisols Alluvial deposits Decomp.-control

II9C 6.01 0.29 3 6.00 6.31 5.73 99 Sdf Mollisols Alluvial deposits Decomp.-cut plot

Nitrogen (%) II1A 2.21 0.05 7 2.21 2.26 2.16 99 Sdf Mollisols Alluvial deposits Decomp. cut plot

II1A 1.98 1 99 Sdf Mollisols Alluvial deposits Decomp.

II1Ai 1.53 0.30 17 1.64 1.80 0.51 99 Sdf Mollisols Alluvial deposits Canopy

II1C 0.84 0.22 10 0.84 1.08 0.36 99 Sdf Mollisols Alluvial deposits Canopy

II1F 0.36 1 99 Sdf Mollisols Alluvial deposits Canopy

II2 0.63 0.09 4 0.64 0.70 0.52 99 Sdf Mollisols Alluvial deposits Decomp. Control

II8A 2.17 0.13 2 2.17 2.26 2.07 99 Sdf Mollisols Alluvial deposits Control plot

II8A 1.86 0.05 2 1.86 1.89 1.82 99 Sdf Mollisols Alluvial deposits Cut plot

II9A 1.01 0.11 9 0.98 1.21 0.88 99 Sdf Mollisols Alluvial deposits Control plot

II9A 0.88 0.14 8 0.92 1.01 0.56 99 Sdf Mollisols Alluvial deposits Cut plot

II9B 0.91 0.08 8 0.92 0.99 0.77 99 Sdf Mollisols Alluvial deposits Control plot

II9B 0.80 0.16 6 0.84 1.01 0.61 99 Sdf Mollisols Alluvial deposits Cut plot

II9C 1.44 0.72 4 1.16 2.50 0.95 99 Sdf Mollisols Alluvial deposits Decomp.-control

II9C 1.38 0.11 2 1.38 1.45 1.30 99 Sdf Mollisols Alluvial deposits Decomp.-cut

Phosphorus (mg/g) II1A 0.66 0.03 7 0.65 0.70 0.60 99 Sdf Mollisols Alluvial deposits Decomp. cut plot

II1A 0.69 0.03 2 0.69 0.71 0.66 99 Sdf Mollisols Alluvial deposits Decomp.

II1Ai 0.41 0.20 17 0.28 0.73 0.21 99 Sdf Mollisols Alluvial deposits Canopy

II1C 0.14 0.05 10 0.14 0.26 0.08 99 Sdf Mollisols Alluvial deposits Canopy

II1F 0.92 1 99 Sdf Mollisols Alluvial deposits Canopy

II2 0.18 0.04 4 0.17 0.22 0.14 99 Sdf Mollisols Alluvial deposits Decomp. Control

II8A 0.70 0.01 2 0.70 0.71 0.69 99 Sdf Mollisols Alluvial deposits Decomp.-control

II8A 0.64 0.07 2 0.64 0.69 0.59 99 Sdf Mollisols Alluvial deposits Decomp.-cut

II8C 1.00 0.28 22 0.97 1.86 0.59 99 Sdf Mollisols Alluvial deposits Decomp.

II9A 0.19 0.03 9 0.18 0.26 0.15 99 Sdf Mollisols Alluvial deposits Control plot

II9A 0.19 0.03 10 0.19 0.24 0.15 99 Sdf Mollisols Alluvial deposits Cut plot

II9B 0.20 0.02 10 0.19 0.25 0.17 99 Sdf Mollisols Alluvial deposits Control plot
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

176

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II9B 0.17 0.03 8 0.16 0.24 0.15 99 Sdf Mollisols Alluvial deposits Cut plot

II9C 0.84 0.40 6 0.68 1.59 0.49 99 Sdf Mollisols Alluvial deposits Control plot

II9C 1.01 0.35 3 0.92 1.39 0.71 99 Sdf Mollisols Alluvial deposits Cut plot

Potassium (mg/g) II1A 1.26 0.24 7 1.20 1.70 0.90 99 Sdf Mollisols Alluvial deposits Decomp. Cut

II1A 1.60 0.66 3 1.70 2.20 0.90 99 Sdf Mollisols Alluvial deposits Decomp.

II1Ai 5.44 5.73 17 2.70 15.50 0.80 99 Sdf Mollisols Alluvial deposits Canopy

II1C 0.70 0.25 9 0.70 1.10 0.30 99 Sdf Mollisols Alluvial deposits Canopy

II1F 11.10 1 99 Sdf Mollisols Alluvial deposits Canopy

II2 3.78 1.42 4 3.10 5.90 3.00 99 Sdf Mollisols Alluvial deposits Decomp.-control

II8A 3.00 0.57 2 3.00 3.40 2.60 99 Sdf Mollisols Alluvial deposits Decomp.

II8A 5.45 1.06 2 5.45 6.20 4.70 99 Sdf Mollisols Alluvial deposits Control plot

II8C 1.49 0.48 22 1.54 2.98 0.59 99 Sdf Mollisols Alluvial deposits Cut plot

II9A 0.68 0.19 9 0.70 0.97 0.28 99 Sdf Mollisols Alluvial deposits Control plot

II9A 3.13 0.77 10 2.95 4.40 2.30 99 Sdf Mollisols Alluvial deposits Cut plot

II9B 2.39 0.71 10 2.40 3.90 1.60 99 Sdf Mollisols Alluvial deposits Control plot

II9B 3.41 3.16 7 2.20 9.90 0.20 99 Sdf Mollisols Alluvial deposits Cut plot

II9C 0.68 0.22 6 0.60 0.95 0.49 99 Sdf Mollisols Alluvial deposits Decomp.-control

II9C 7.85 1.20 2 7.85 8.70 7.00 99 Sdf Mollisols Alluvial deposits Decomp.-cut plot

Musa paradisiaca C/N II3 25 9 2 25 32 19 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II3 45 0 2 45 45 45 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II3 1.89 0.66 2 1.89 2.35 1.42 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II3 0.30 0.10 2 0.30 0.38 0.23 135 Lmrf Ultisols Tuffac. sandstone

Musa spp. Aluminum (mg/g) II3 0.19 0.12 4 0.16 0.35 0.09 135 Lmrf Ultisols Tuffac. sandstone

II3A 0.09 0.06 4 0.12 0.13 0.01 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II3 3.35 1.83 4 2.82 5.82 1.95 135 Lmrf Ultisols Tuffac. sandstone

II3A 8.05 3.05 4 7.28 12.00 5.63 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II3 0.20 0.11 4 0.16 0.36 0.12 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

177

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II3A 0.12 0.04 4 0.13 0.14 0.07 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II3 2.36 0.44 4 2.23 2.99 1.97 135 Lmrf Ultisols Tuffac. sandstone

II3A 3.67 0.58 4 3.63 4.26 3.16 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II3 0.76 0.48 4 0.60 1.46 0.38 135 Lmrf Ultisols Tuffac. sandstone

II3A 1.81 1.21 4 1.80 2.89 0.77 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II3 1.37 0.42 4 1.55 1.64 0.74 135 Lmrf Ultisols Tuffac. sandstone

II3A 1.53 0.79 4 1.75 2.21 0.41 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II3 1.01 0.25 4 1.12 1.16 0.64 135 Lmrf Ultisols Tuffac. sandstone

II3A 0.75 0.47 4 0.75 1.16 0.35 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II3 31.13 7.81 4 28.55 42.40 25.03 135 Lmrf Ultisols Tuffac. sandstone

II3A 25.58 3.25 4 24.34 30.37 23.24 135 Lmrf Ultisols Tuffac. sandstone

Myrcia deflexa Aluminum (mg/g) II1B 0.14 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.07 1 93 Wflm Inceptisols Tuffac. sandstone

Ash (%) II1A 8 .47 1 94 Wfs Ultisols Tuffac. sandstone

II1B 2.97 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 3 .66 1 94 Wfs Ultisols Tuffac. sandstone

II1C 0.21 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 2.75 1 94 Wfs Ultisols Tuffac. sandstone

C/N II1B 83 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 420 1 93 Wflm Inceptisols Tuffac. sandstone

Calcium (mg/g) II1A 6.86 0.98 3 6.38 7.99 3.20 135 Lmrf Ultisols Tuffac. sandstone

II1B 9.39 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 3.93 1.57 2 3.93 5.04 2.82 135 Lmrf Ultisols Tuffac. sandstone

II1C 1.18 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 1.36 1 135 Lmrf Ultisols Tuffac. sandstone

II1F 4.56 1 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II1B 0.15 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.06 1 93 Wflm Inceptisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

178

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Magnesium (mg/g) II1A 2.62 0.24 3 2.71 2.80 2.35 135 Lmrf Ultisols Tuffac. sandstone

II1B 1.02 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.58 0.31 2 0.58 0.80 0.36 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.27 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.32 1 135 Lmrf Ultisols Tuffac. sandstone

II1F 1.32 1 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II1B 1.32 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.33 1 93 Wflm Inceptisols Tuffac. sandstone

Nitrogen (%) II1A 1.53 1 94 Wfs Ultisols Tuffac. sandstone

II1A 1.11 0.09 3 1.06 1.21 1.05 135 Lmrf Ultisols Tuffac. sandstone

II1B 0.54 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 1.05 1 94 Wfs Ultisols Tuffac. sandstone

II1B 0.38 0.14 2 0.38 0.48 0.28 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.11 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.95 1 94 Wfs Ultisols Tuffac. sandstone

II1C 0.16 1 135 Lmrf Ultisols Tuffac. sandstone

II1F 1.10 1 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.48 1 94 Wfs Ultisols Tuffac. sandstone

II1A 0.48 0.04 3 0.47 0.53 0.45 135 Lmrf Ultisols Tuffac. sandstone

II1B 0.48 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.43 1 94 Wfs Ultisols Tuffac. sandstone

II1B 0.23 0.10 2 0.23 0.30 0.15 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.07 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.35 1 94 Wfs Ultisols Tuffac. sandstone

II1C 0.08 1 135 Lmrf Ultisols Tuffac. sandstone

II1F 1.24 1 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II1A 6.09 1 94 Wfs Ultisols Tuffac. sandstone

II1A 6.91 0.52 3 7.11 7.29 6.32 135 Lmrf Ultisols Tuffac. sandstone

II1B 5.59 1 93 Wflm Inceptisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

179

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1B 2.89 1 94 Wfs Ultisols Tuffac. sandstone

II1B 2.94 1.36 2 2.94 3.90 1.98 135 Lmrf Ultisols Tuffac. sandstone

II1C 2.32 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 2.34 1 94 Wfs Ultisols Tuffac. sandstone

II1C 1.10 1 135 Lmrf Ultisols Tuffac. sandstone

II1F 19.15 1 135 Lmrf Ultisols Tuffac. sandstone

Myrcia splendens Ash (%) II1A 9.98 2.14 16 10.48 12.90 5.72 94 Swf Ultisols Tuffac. sandstone Understory

II1A 10.05 1 101 Lmwf Inceptisols Tuffac. sandstone Overstory

II1A 4.20 0.27 5 4.22 4.54 3.78 126 Ultisols Siltstone,sandstone Urban forest, RP

II1B 3.67 1.35 11 3.49 6.34 1.31 94 Swf Ultisols Tuffac. sandstone Understory

II1C 1.91 1.24 15 1.48 4.62 0.75 94 Swf Ultisols Tuffac. sandstone Understory

II1C 0.92 0.27 5 0.81 1.29 0.66 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 5.36 1.44 6 5.43 6.95 3.17 126 Ultisols Siltstone,sandstone Urban forest, RP

C/N II1A 32 1 5 32 33 31 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 261 85 5 293 347 132 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 66 9 6 67 74 49 126 Ultisols Siltstone,sandstone Urban forest, RP

Carbon (%) II1A 51 0 5 51 52 51 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 53 0 5 53 53 52 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 52 1 6 52 53 51 126 Ultisols Siltstone,sandstone Urban forest, RP

Nitrogen (%) II1A 1.09 0.14 16 1.14 1.33 0.82 94 Swf Ultisols Tuffac. sandstone Understory

II1A 1.22 1 101 Lmwf Inceptisols Tuffac. sandstone Overstory

Nitrogen (%) II1A 1.61 0.05 5 1.60 1.68 1.56 126 Ultisols Siltstone,sandstone Urban forest, RP

II1B 0.60 0.19 11 0.68 0.79 0.24 94 Swf Ultisols Tuffac. sandstone Understory

II1C 0.39 0.17 15 0.37 0.68 0.15 94 Swf Ultisols Tuffac. sandstone Understory

II1C 0.23 0.10 5 0.18 0.40 0.15 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 0.80 0.12 6 0.78 1.05 0.72 126 Ultisols Siltstone,sandstone Urban forest, RP

Phosphorus (mg/g) II1A 0.58 0.15 16 0.56 0.85 0.38 94 Swf Ultisols Tuffac. sandstone Understory

II1A 0.63 1 101 Lmwf Inceptisols Tuffac. sandstone Overstory
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

180

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1B 0.45 0.17 11 0.43 0.70 0.15 94 Swf Ultisols Tuffac. sandstone Understory

II1C 0.32 0.15 15 0.29 0.61 0.11 94 Swf Ultisols Tuffac. sandstone Understory

Potassium (mg/g) II1A 4.11 1.90 16 4.90 6.50 0.70 94 Swf Ultisols Tuffac. sandstone Understory

II1A 4.50 1 101 Lmwf Inceptisols Tuffac. sandstone Overstory

II1B 0.28 0.15 11 0.34 0.46 0.03 94 Swf Ultisols Tuffac. sandstone Understory

II1C 1.28 1.28 15 1.60 4.30 0.70 94 Swf Ultisols Tuffac. sandstone Understory

Sulfur (%) II1A 0.44 0.05 5 0.43 0.51 0.36 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 0.15 0.08 5 0.13 0.29 0.09 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 0.54 0.09 6 0.52 0.69 0.44 126 Ultisols Siltstone,sandstone Urban forest, RP

Wood density (g/cc) II1C 0.61 0.07 5 0.61 0.71 0.53 126 Ultisols Siltstone,sandstone Urban forest, RP

Neolaugeria resinosa Aluminum (mg/kg) II1A 27 1 102 Smf Limestone Tuffac. sandstone New Adult 

Ash (%) II1A 5.06 1 102 Smf Limestone Tuffac. sandstone New Adult 

Calcium (mg/g) II1A 6.66 1 102 Smf Limestone Tuffac. sandstone New Adult 

Cobalt (μg/g) II1A 0.95 1 102 Smf Limestone Tuffac. sandstone New Adult 

Iron (mg/kg) II1A 47 1 102 Smf Limestone Tuffac. sandstone New Adult 

Magnesium (mg/g) II1A 3.84 1 102 Smf Limestone Tuffac. sandstone New Adult 

Manganese (mg/kg) II1A 127 1 102 Smf Limestone Tuffac. sandstone New Adult 

Nickel (μg/g) II1A 30 1 102 Smf Limestone Tuffac. sandstone New Adult 

Nitrogen (%) II1A 1.10 1 102 Smf Limestone Tuffac. sandstone New Adult 

Phosphorus (mg/g) II1A 0.40 1 102 Smf Limestone Tuffac. sandstone New Adult 

Potassium (mg/g) II1A 13.89 1 102 Smf Limestone Tuffac. sandstone

Nephrolepsis portoricensis Aluminum (mg/g) II6 4.91 3.09 21 4.73 10.90 0.05 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II6 2.74 1.60 21 2.46 8.54 1.07 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II6 41 1 9 42 42 38 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II6 0.69 1.59 21 0.25 7.47 0.04 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II6 4.04 1.55 21 4.02 7.77 1.41 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II6 0.10 0.04 21 0.09 0.20 0.04 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

181

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Nitrogen (%) II6 1.50 0.38 25 1.48 2.43 0.80 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II6 0.83 0.37 21 0.68 1.91 0.47 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II6 13.83 8.40 21 11.16 27.50 1.46 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II6 0.50 0.18 9 0.56 0.73 0.16 135 Lmrf Ultisols Tuffac. sandstone

Nephrolepsis rivlaris Aluminum (mg/g) II6 0.30 0.16 4 0.26 0.51 0.14 135 Lmrf Ultisols Tuffac. sandstone

C/N II6 3 7 4 36 39 23 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II6 10.75 2.34 4 10.49 13.32 8.70 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II6 44 1 4 44 46 44 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II6 0.18 0.05 4 0.19 0.24 0.13 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II6 5.60 1.08 4 5.71 6.79 4.17 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II6 0.23 0.07 4 0.24 0.30 0.14 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II6 1.17 0.36 8 1.16 1.91 0.72 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II6 0.56 0.20 4 0.80 0.94 0.50 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II6 15.79 6.68 4 16.11 21.82 9.13 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II6 0.14 0.03 4 0.19 0.24 0.17 135 Lmrf Ultisols Tuffac. sandstone

Nepsera aquatica Aluminum (mg/g) II3 0.64 1 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II3 2.52 1 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II3 0.15 1 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II3 1.61 1 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II3 0.04 1 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II3 0.66 1 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II3 0.37 1 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II3 8.90 1 135 Lmrf Ultisols Tuffac. sandstone

Ocotea leucoxylan Aluminum (mg/g) II1A 0.40 1 93 Wflm Inceptisols Tuffac. sandstone

II1H 0.41 0.15 2 0.41 0.51 0.30 135 Lmrf Ultisols Tuffac. sandstone

Ash (%) II1A 4.52 1 93 Wflm Inceptisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

182

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 9.16 2.26 6 8.84 12.82 6.92 126 Smf Ultisols Tuffac. sandstone Luquillo Exp. Forest

II1C 1.63 0.24 6 1.62 1.97 1.28 126 Smf Ultisols Tuffac. sandstone Luquillo Exp. Forest

II1Ci 12.14 1.65 5 11.47 14.05 10.51 126 Smf Ultisols Tuffac. sandstone Luquillo Exp. Forest

C/N II1A 21 1 93 Wflm Inceptisols Tuffac. sandstone

II1A 27 2 6 28 28 24 126 Smf Ultisols Tuffac. sandstone Luquillo Exp. Forest

II1C 178 39 6 189 214 108 126 Smf Ultisols Tuffac. sandstone Luquillo Exp. Forest

II1Ci 32 4 5 33 37 27 126 Smf Ultisols Tuffac. sandstone Luquillo Exp. Forest

Calcium (mg/g) II1A 7.61 2.70 5 8.08 10.31 4.60 135 Lmrf Ultisols Tuffac. sandstone

II1A 7.31 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 6.09 1.42 4 5.60 8.17 4.98 135 Lmrf Ultisols Tuffac. sandstone

II1C 1.38 0.37 8 1.25 2.12 1.01 135 Lmrf Ultisols Tuffac. sandstone

II1Ci 10.45 1.77 9 10.42 12.67 8.66 135 Lmrf Ultisols Tuffac. sandstone

II1F 3.98 1.12 4 3.72 5.46 3.00 135 Lmrf Ultisols Tuffac. sandstone

II1H 7.90 0.51 2 7.90 8.25 7.54 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II1A 54 2 6 53 56 52 126 Smf Ultisols Tuffac. sandstone Luquillo Exp. Forest

II1C 53 1 6 53 54 53 126 Smf Ultisols Tuffac. sandstone Luquillo Exp. Forest

II1Ci 49 1 5 49 50 47 126 Smf Ultisols Tuffac. sandstone Luquillo Exp. Forest

Iron (mg/g) II1A 0.14 1 93 Wflm Inceptisols Tuffac. sandstone

II1H 0.28 0.03 2 0.28 0.30 0.26 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II1A 1.80 0.30 5 1.99 2.05 1.38 135 Lmrf Ultisols Tuffac. sandstone

II1A 2.55 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 1.22 0.36 4 1.24 1.60 0.79 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.15 0.04 8 0.16 0.19 0.09 135 Lmrf Ultisols Tuffac. sandstone

II1Ci 0.60 0.12 9 0.60 0.79 0.39 135 Lmrf Ultisols Tuffac. sandstone

II1F 2.29 0.26 4 2.30 2.60 1.98 135 Lmrf Ultisols Tuffac. sandstone

II1H 1.70 0.10 2 1.70 1.77 1.62 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II1A 0.43 1 93 Wflm Inceptisols Tuffac. sandstone

II1H 0.34 0.02 2 0.34 0.35 0.32 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II1A 2.11 0.10 5 2.11 2.26 2.01 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

183

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 2.14 1 93 Wflm Inceptisols Tuffac. sandstone

II1A 2.37 0.27 3 2.29 2.68 2.15 94 Swf Ultisols Tuffac. sandstone

II1A 2.01 0.10 6 2.00 2.16 1.92 126 Smf Ultisols Tuffac. sandstone Luquillo Exp. Forest

II1B 1.07 0.41 4 1.09 1.54 0.56 135 Lmrf Ultisols Tuffac. sandstone

II1B 1.44 0.08 3 1.44 1.52 1.36 94 Swf Ultisols Tuffac. sandstone

II1C 0.25 0.03 8 0.25 0.29 0.20 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.93 0.48 3 0.69 1.48 0.62 94 Swf Ultisols Tuffac. sandstone

II1C 0.32 0.09 6 0.29 0.49 0.25 126 Smf Ultisols Tuffac. sandstone Luquillo Exp. Forest

II1Ci 1.26 0.08 9 1.26 1.36 1.13 135 Lmrf Ultisols Tuffac. sandstone

II1Ci 1.54 0.18 5 1.48 1.76 1.36 126 Smf Ultisols Tuffac. sandstone Luquillo Exp. Forest

II1F 3.52 0.02 4 3.52 3.55 3.48 135 Lmrf Ultisols Tuffac. sandstone

II1H 1.62 0.02 2 1.62 1.63 1.60 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.79 0.22 5 0.78 1.13 0.58 135 Lmrf Ultisols Tuffac. sandstone

II1A 1.12 1 93 Wflm Inceptisols Tuffac. sandstone

II1A 0.94 0.30 3 0.77 1.29 0.76 94 Swf Ultisols Tuffac. sandstone

II1B 0.42 0.19 4 0.44 0.61 0.18 135 Lmrf Ultisols Tuffac. sandstone

II1B 1.14 0.37 3 1.04 1.55 0.84 94 Swf Ultisols Tuffac. sandstone

II1C 0.08 0.01 8 0.08 0.10 0.06 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.61 0.40 3 0.54 1.04 0.24 94 Swf Ultisols Tuffac. sandstone

II1Ci 0.46 0.06 9 0.46 0.55 0.34 135 Lmrf Ultisols Tuffac. sandstone

II1F 2.58 0.44 4 2.61 2.99 2.13 135 Lmrf Ultisols Tuffac. sandstone

II1H 0.73 0.03 2 0.73 0.75 0.70 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II1A 7.37 1.45 5 7.02 9.88 6.34 135 Lmrf Ultisols Tuffac. sandstone

II1A 10.82 1 93 Wflm Inceptisols Tuffac. sandstone

II1A 4.83 3.01 3 3.99 8.17 2.34 94 Swf Ultisols Tuffac. sandstone

II1B 5.15 1.33 4 5.39 6.40 3.42 135 Lmrf Ultisols Tuffac. sandstone

II1B 4.92 2.90 3 6.28 6.90 1.59 94 Swf Ultisols Tuffac. sandstone

II1C 1.94 0.23 8 1.89 2.27 1.54 135 Lmrf Ultisols Tuffac. sandstone

II1C 3.34 1.81 3 3.06 5.27 1.68 94 Swf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

184

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1Ci 4.76 0.56 9 4.78 5.83 3.92 135 Lmrf Ultisols Tuffac. sandstone

II1F 12.95 1.04 4 12.79 14.36 11.87 135 Lmrf Ultisols Tuffac. sandstone

II1H 7.83 0.32 2 7.83 8.06 7.61 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II1A 0.20 0.02 6 0.20 0.24 0.18 126 Smf Ultisols Tuffac. sandstone Luquillo Exp. Forest

II1C 0.01 0.01 6 0.01 0.03 0.01 126 Smf Ultisols Tuffac. sandstone Luquillo Exp. Forest

II1Ci 0.17 0.03 5 0.16 0.21 0.15 126 Smf Ultisols Tuffac. sandstone Luquillo Exp. Forest

Wood density (g/cc) II1C 0.34 0.04 5 0.34 0.40 0.30 126 Smf Ultisols Tuffac. sandstone Luquillo Exp. Forest

Ocotea moschata Wood density (g/cc) II1B 0.38 0.01 3 0.38 0.38 0.37 97 Wfs Inceptisols Tuffac. sandstone

Ocotea spathulata Aluminum (mg/g) II1A 0.58 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.05 1 93 Wflm Inceptisols Tuffac. sandstone

Ash (%) II1B 3.10 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.35 1 93 Wflm Inceptisols Tuffac. sandstone

C/N II1B 43 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 195 1 93 Wflm Inceptisols Tuffac. sandstone

Calcium (mg/g) II1A 6.43 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 2.12 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 1.41 1 93 Wflm Inceptisols Tuffac. sandstone

Iron (mg/g) II1A 1.88 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.04 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.03 1 93 Wflm Inceptisols Tuffac. sandstone

Magnesium (mg/g) II1B 0.58 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.26 1 93 Wflm Inceptisols Tuffac. sandstone

Manganese (mg/g) II1A 0.10 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.03 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.01 1 93 Wflm Inceptisols Tuffac. sandstone

Nitrogen (%) II1B 1.03 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.24 1 93 Wflm Inceptisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

185

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Phosphorus (mg/g) II1A 1.56 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 1.89 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.11 1 93 Wflm Inceptisols Tuffac. sandstone

Potassium (mg/g) II1A 18.65 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 11.29 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 1.49 1 93 Wflm Inceptisols Tuffac. sandstone

Odontosoria spp. Aluminum (mg/g) II6 1.52 1 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II6 1.60 1 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II6 0.79 1 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II6 1.49 1 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II6 0.20 1 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II6 1.04 1 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II6 0.49 1 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II6 9.72 1 135 Lmrf Ultisols Tuffac. sandstone

Olyra latifolia Aluminum (mg/g) II3 0.40 1 135 Lmrf Ultisols Tuffac. sandstone

C/N II3 25 1 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II3 0.77 1 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II3 35 1 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II3 0.13 1 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II3 1.03 1 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II3 0.27 1 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II3 1.01 1 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II3 0.48 1 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II3 9.39 1 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II3 0.24 1 135 Lmrf Ultisols Tuffac. sandstone

Ormosia krugii Ash (%) II1B 0.74 0.48 8 0.66 1.75 0.09 68 Swf Ultisols Tuffac. sandstone Class 3 LD
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

186

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1B 0.73 0.69 2 0.73 1.22 0.24 68 Swf Ultisols Tuffac. sandstone Class 3 ST

Calcium (mg/g) II1A 2.08 0.49 9 1.82 2.89 1.65 135 Lmrf Ultisols Tuffac. sandstone

II1B 2.77 1.55 8 3.01 4.87 0.84 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 0.19 0.12 2 0.19 0.27 0.11 68 Swf Ultisols Tuffac. sandstone Class 3 ST

II1B 1.74 0.46 9 1.85 2.38 0.84 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.29 0.12 12 0.27 0.52 0.07 135 Lmrf Ultisols Tuffac. sandstone

II1Ci 3.91 0.90 12 4.06 5.02 2.55 135 Lmrf Ultisols Tuffac. sandstone

II1F 1.75 0.52 3 1.59 2.34 1.34 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II1A 0.91 0.11 9 0.88 1.13 0.80 135 Lmrf Ultisols Tuffac. sandstone

II1B 0.43 0.17 9 0.41 0.65 0.18 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.06 0.03 12 0.07 0.09 0.02 135 Lmrf Ultisols Tuffac. sandstone

II1Ci 0.28 0.09 12 0.24 0.49 0.18 135 Lmrf Ultisols Tuffac. sandstone

II1F 0.77 0.09 3 0.72 0.87 0.71 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II1A 2.12 0.09 9 2.10 2.27 2.02 135 Lmrf Ultisols Tuffac. sandstone

II1B 0.40 0.20 8 0.38 0.72 0.15 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 0.58 0.52 2 0.58 0.94 0.21 68 Swf Ultisols Tuffac. sandstone Class 3 ST

II1B 0.95 0.23 9 0.88 1.25 0.60 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.21 0.02 12 0.21 0.24 0.18 135 Lmrf Ultisols Tuffac. sandstone

II1Ci 0.90 0.05 12 0.91 0.98 0.83 135 Lmrf Ultisols Tuffac. sandstone

II1F 1.27 0.11 3 1.21 1.39 1.20 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.76 0.06 9 0.76 0.85 0.65 135 Lmrf Ultisols Tuffac. sandstone

II1B 0.05 0.14 8 0.14 0.20 0.07 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 0.05 0.03 2 0.05 0.09 0.01 68 Swf Ultisols Tuffac. sandstone Class 3 ST

II1B 0.49 0.30 9 0.36 0.82 0.19 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.03 0.01 12 0.03 0.47 0.02 135 Lmrf Ultisols Tuffac. sandstone

II1Ci 0.48 0.18 12 0.18 0.20 0.13 135 Lmrf Ultisols Tuffac. sandstone

II1F 0.41 0.18 3 0.37 0.61 0.26 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II1A 4.45 0.59 9 4.26 5.78 3.78 135 Lmrf Ultisols Tuffac. sandstone

II1B 0.81 0.98 9 0.46 3.30 0.17 68 Swf Ultisols Tuffac. sandstone Class 3 LD
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

187

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1B 0.63 1 68 Swf Ultisols Tuffac. sandstone Class 3 ST

II1B 2.85 1.40 9 2.58 4.31 1.23 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.42 0.17 12 0.40 0.80 0.21 135 Lmrf Ultisols Tuffac. sandstone

II1Ci 1.75 0.55 12 1.54 2.68 1.03 135 Lmrf Ultisols Tuffac. sandstone

II1F 3.69 0.97 3 3.41 4.77 2.90 135 Lmrf Ultisols Tuffac. sandstone

Ouratea littoralis Aluminum (mg/kg) II1A 42 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 38 1 102 Smf Limestone Tuffac. sandstone Adult 

Ash (%) II1A 4.68 1 102 Sdf Mollisols Alluvial deposits Adult 

II1A 4.74 1 149 Sdf Mollisols Alluvial deposits Susua forest

Calcium (mg/g) II1A 11.82 1 102 Smf Limestone Tuffac. sandstone

II1A 12.07 1 149 Sdf Mollisols Alluvial deposits Susua forest

Cobalt (μg/g) II1A 0.75 1 102 Smf Limestone Tuffac. sandstone Adult 

Iron (mgkg) II1A 78 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 58 1 102 Smf Limestone Tuffac. sandstone Adult 

Magnesium (mg/g) II1A 3.09 1 102 Smf Limestone Tuffac. sandstone Adult 

II1A 2.66 1 149 Sdf Mollisols Alluvial deposits Susua forest

Manganese (mg/kg) II1A 48 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 52 1 102 Smf Limestone Tuffac. sandstone Adult 

Nickel (μg/g) II1A 34.15 1 102 Smf Limestone Tuffac. sandstone Adult 

Nitrogen (%) II1A 1.06 1 102 Smf Limestone Tuffac. sandstone Adult 

II1A 1.08 1 149 Sdf Mollisols Alluvial deposits Susua forest

Phosphorus (mg/g) II1A 0.54 1 102 Smf Limestone Tuffac. sandstone Adult 

 II1A 0.39 1 149 Sdf Mollisols Alluvial deposits Susua forest

Potassium (mg/g) II1A 6.22 1 102 Smf Limestone Tuffac. sandstone Adult 

II1A 5.00 1 149 Sdf Mollisols Alluvial deposits Susua forest

Zinc (μg/g) II1A 11.90 1 149 Sdf Mollisols Alluvial deposits Susua forest

Palicourea crocea Aluminum (mg/g) II2 0.32 0.10 2 0.32 0.39 0.25 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

188

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Calcium (mg/g) II2 5.38 1.64 2 5.38 6.54 4.23 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II2 0.08 0.06 2 0.08 0.12 0.04 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II2 3.29 1.52 2 3.29 4.37 2.22 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II2 0.06 0.04 2 0.06 0.08 0.03 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II2 1.03 0.21 2 1.03 1.18 0.88 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II2 0.71 0.16 2 0.71 0.82 0.60 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II2 6.31 26.46 2 46.31 65.02 27.59 135 Lmrf Ultisols Tuffac. sandstone

Palicourea riparia Aluminum (mg/g) II2A 6.76 1.73 12 6.63 11.28 4.51 135 Lmrf Ultisols Tuffac. sandstone

II2C 1.14 0.47 15 1.11 2.17 0.38 135 Lmrf Ultisols Tuffac. sandstone

II2E 1.35 0.64 2 1.35 1.81 0.90 135 Lmrf Ultisols Tuffac. sandstone

II2H 2.07 1 135 Lmrf Ultisols Tuffac. sandstone

II8A 15.09 3.66 2 15.09 17.68 2.50 146 Swf Ultisols Tuffac. sandstone

II8B 1.22 1.56 2 1.22 2.33 0.12 146 Swf Ultisols Tuffac. sandstone

Ash (%) II2A 9.13 1.32 12 8.97 11.22 7.36 94 Swf Ultisols Tuffac. sandstone

II2B 5.20 0.59 8 5.14 5.98 4.36 94 Swf Ultisols Tuffac. sandstone

II2C 3.96 1.90 10 4.22 6.76 1.38 94 Swf Ultisols Tuffac. sandstone

II2F 7.73 0.85 4 8.06 8.31 6.47 94 Swf Ultisols Tuffac. sandstone

II8A 10.41 0.57 2 10.41 10.81 10.01 146 Swf Ultisols Tuffac. sandstone

II8B 3.18 2.97 2 3.18 5.28 1.08 146 Swf Ultisols Tuffac. sandstone

C/N II8A 18 8 2 18 24 12 146 Swf Ultisols Tuffac. sandstone

II8B 75 8 2 75 81 69 146 Swf Ultisols Tuffac. sandstone

Calcium (mg/g) II2A 8.65 1.10 13 8.66 10.31 6.29 135 Lmrf Ultisols Tuffac. sandstone

II2B 11.02 1 135 Lmrf Ultisols Tuffac. sandstone

II2C 4.26 3.39 16 3.59 16.53 2.03 135 Lmrf Ultisols Tuffac. sandstone

II2E 1.21 0.04 2 1.21 1.23 1.18 135 Lmrf Ultisols Tuffac. sandstone

II2F 6.60 1 135 Lmrf Ultisols Tuffac. sandstone

II2H 4.73 1 135 Lmrf Ultisols Tuffac. sandstone

II8A 12.26 5.40 2 12.26 16.07 8.44 146 Swf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

189

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8B 3.61 4.49 2 3.61 6.78 0.43 146 Swf Ultisols Tuffac. sandstone

Carbon (%) II8A 51 3 2 51 53 49 146 Swf Ultisols Tuffac. sandstone

II8B 52 2 2 52 53 51 146 Swf Ultisols Tuffac. sandstone

Iron (mg/g) II2A 0.22 0.06 12 0.21 0.36 0.12 135 Lmrf Ultisols Tuffac. sandstone

II2C 0.09 0.04 15 0.09 0.20 0.04 135 Lmrf Ultisols Tuffac. sandstone

II2E 1.10 0.45 2 1.10 1.42 0.78 135 Lmrf Ultisols Tuffac. sandstone

II2H 0.19 1 135 Lmrf Ultisols Tuffac. sandstone

II8A 0.57 0.45 2 0.57 0.89 0.25 146 Swf Ultisols Tuffac. sandstone

II8B 0.03 0.03 2 0.03 0.05 0.01 146 Swf Ultisols Tuffac. sandstone

Magnesium (mg/g) II2A 4.21 0.67 13 4.17 5.36 2.87 135 Lmrf Ultisols Tuffac. sandstone

II2B 3.53 1 135 Lmrf Ultisols Tuffac. sandstone

II2C 1.00 0.40 16 0.91 1.98 0.52 135 Lmrf Ultisols Tuffac. sandstone

II2E 0.44 0.09 2 0.44 0.50 0.38 135 Lmrf Ultisols Tuffac. sandstone

II2F 3.86 1 135 Lmrf Ultisols Tuffac. sandstone

II2H 2.10 1 135 Lmrf Ultisols Tuffac. sandstone

II8A 3.48 0.83 2 3.48 4.07 2.89 146 Swf Ultisols Tuffac. sandstone

II8B 0.95 1.25 2 0.95 1.83 0.07 146 Swf Ultisols Tuffac. sandstone

Manganese (mg/g) II2A 0.15 0.07 12 0.15 0.28 0.07 135 Lmrf Ultisols Tuffac. sandstone

II2C 0.11 0.04 15 0.11 0.22 0.05 135 Lmrf Ultisols Tuffac. sandstone

II2E 0.05 0.05 2 0.05 0.09 0.02 135 Lmrf Ultisols Tuffac. sandstone

II2H 0.12 1 135 Lmrf Ultisols Tuffac. sandstone

II8A 0.20 0.17 2 0.20 0.32 0.09 146 Swf Ultisols Tuffac. sandstone

II8B 0.07 0.08 2 0.07 0.12 0.01 146 Swf Ultisols Tuffac. sandstone

Nitrogen (%) II2A 2.29 0.34 13 2.35 2.68 1.52 135 Lmrf Ultisols Tuffac. sandstone

II2A 2.68 0.30 12 2.68 3.08 2.12 94 Swf Ultisols Tuffac. sandstone

II2B 1.02 1 135 Lmrf Ultisols Tuffac. sandstone

II2B 1.15 0.20 8 1.13 1.37 0.94 94 Swf Ultisols Tuffac. sandstone

II2C 0.71 0.31 16 0.62 1.68 0.42 135 Lmrf Ultisols Tuffac. sandstone

II2C 0.96 0.39 10 0.97 1.67 0.33 94 Swf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

190

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II2E 0.68 0.21 2 0.68 0.82 0.53 135 Lmrf Ultisols Tuffac. sandstone

II2F 2.92 1 135 Lmrf Ultisols Tuffac. sandstone

II2F 2.35 0.94 4 1.86 3.43 1.75 94 Swf Ultisols Tuffac. sandstone

II2H 1.33 1 135 Lmrf Ultisols Tuffac. sandstone

II8A 3.33 1.77 2 3.33 4.58 2.08 146 Swf Ultisols Tuffac. sandstone

II8B 0.70 0.05 2 0.70 0.73 0.66 146 Swf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II2A 0.62 0.07 13 0.63 0.79 0.49 135 Lmrf Ultisols Tuffac. sandstone

II2A 0.85 0.22 12 0.84 1.21 0.49 94 Swf Ultisols Tuffac. sandstone

II2B 0.44 1 135 Lmrf Ultisols Tuffac. sandstone

II2B 0.55 0.15 8 0.55 0.76 0.32 94 Swf Ultisols Tuffac. sandstone

II2C 0.29 0.12 16 0.26 0.54 0.14 135 Lmrf Ultisols Tuffac. sandstone

II2C 0.48 0.26 10 0.46 0.91 0.17 94 Swf Ultisols Tuffac. sandstone

II2E 0.27 0.07 2 0.27 0.32 0.21 135 Lmrf Ultisols Tuffac. sandstone

II2F 1.73 1 116

II2F 1.43 0.50 4 1.41 2.03 0.87 94 Swf Ultisols Tuffac. sandstone

II2H 0.41 1 135 Lmrf Ultisols Tuffac. sandstone

II8A 1.04 0.55 2 1.04 1.43 0.65 146 Swf Ultisols Tuffac. sandstone

II8B 0.49 0.64 2 0.49 0.95 0.04 146 Swf Ultisols Tuffac. sandstone

Potassium (mg/g) II2A 13.78 1.96 13 14.17 16.90 9.51 135 Lmrf Ultisols Tuffac. sandstone

II2A 13.71 5.41 12 14.80 23.90 2.20 94 Swf Ultisols Tuffac. sandstone

II2B 4.32 1 135 Lmrf Ultisols Tuffac. sandstone

II2B 6.60 2.62 8 5.40 9.50 1.50 94 Swf Ultisols Tuffac. sandstone

II2C 7.05 0.90 16 7.31 8.05 5.17 135 Lmrf Ultisols Tuffac. sandstone

II2C 8.01 4.05 10 7.20 15.70 2.60 94 Swf Ultisols Tuffac. sandstone

II2E 4.97 0.62 2 4.97 5.41 4.54 135 Lmrf Ultisols Tuffac. sandstone

II2F 13.48 1 135 Lmrf Ultisols Tuffac. sandstone

II2F 15.10 1.18 4 14.70 16.80 14.10 94 Swf Ultisols Tuffac. sandstone

II2H 7.85 1 135 Lmrf Ultisols Tuffac. sandstone

II2C 8.72 8.37 2 8.72 14.63 2.80 146 Swf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

191

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8B 5.43 7.59 2 5.43 10.80 0.07 146 Swf Ultisols Tuffac. sandstone

Panicum maximum Aluminum (mg/g) II3 0.23 0.18 6 0.14 0.57 0.11 144 Mfs Histosol Alluvial deposits

Ash (%) II3 8.40 2.10 6 7.88 12.46 6.63 144 Mfs Histosol Alluvial deposits

C/N II3 56 16 6 50 86 45 144 Mfs Histosol Alluvial deposits

Calcium (mg/g) II3 3.63 1.23 6 4.27 4.58 1.56 144 Mfs Histosol Alluvial deposits

Carbon (%) II3 46 1 6 47 48 44 144 Mfs Histosol Alluvial deposits

Iron (mg/g) II3 0.24 0.27 6 0.12 0.77 0.04 144 Mfs Histosol Alluvial deposits

Magnesium (mg/g) II3 2.63 0.86 6 2.36 3.88 1.78 144 Mfs Histosol Alluvial deposits

Manganese (mg/g) II3 0.05 0.02 6 0.05 0.06 0.02 144 Mfs Histosol Alluvial deposits

Nitrogen (%) II3 0.88 0.18 6 0.95 1.03 0.54 144 Mfs Histosol Alluvial deposits

Phosphorus (mg/g) II3 1.63 0.18 6 1.71 1.81 1.39 144 Mfs Histosol Alluvial deposits

Potassium (mg/g) II3 10.55 2.43 6 11.66 13.02 6.72 144 Mfs Histosol Alluvial deposits

Pasture Aluminum (mg/g) II9A 4.21 0.51 4 4.42 4.56 3.45 153 Swf Ultisols Tuffac. sandstone

Ash (%) II9A 8.37 0.64 4 8.49 8.97 7.54 153 Swf Ultisols Tuffac. sandstone

C/N II3 26 3 24 26 32 20 110 Lmrf Ultisols Tuffac. sandstone Mameyes

II3 25 6 24 24 44 17 110 Wfs Ultisols Lava Sabana

II9A 101 5 4 101 108 97 153 Swf Ultisols Tuffac. sandstone

Calcium (mg/g) II9A 0.28 0.04 4 0.27 0.33 0.24 153 Swf Ultisols Tuffac. sandstone

Carbon (%) II3 45 3 24 45 49 36 110 Lmrf Ultisols Tuffac. sandstone Mameyes

II3 45 3 24 45 50 41 110 Wfs Ultisols Lava Sabana

II9A 49 0 4 48 49 48 153 Swf Ultisols Tuffac. sandstone

Iron (mg/g) II9A 6.26 0.91 4 6.50 7.07 4.96 153 Swf Ultisols Tuffac. sandstone

Magnesium (mg/g) II9A 0.37 0.05 4 0.38 0.41 0.31 153 Swf Ultisols Tuffac. sandstone

Manganese (mg/g) II9A 0.23 0.04 4 0.22 0.28 0.20 153 Swf Ultisols Tuffac. sandstone

Nitrogen (%) II3 1.76 0.25 24 1.50 2.21 1.34 110 Lmrf Ultisols Tuffac. sandstone Mameyes

II3 1.90 0.40 24 1.84 2.72 1.09 110 Wfs Ultisols Lava Sabana

II9A 0.48 0.02 4 0.49 0.50 0.45 153 Swf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

192

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Phosphorus (mg/g) II9A 0.20 0.01 4 0.19 0.22 0.19 153 Swf Ultisols Tuffac. sandstone

Potassium (mg/g) II9A 2.25 0.13 4 2.24 2.39 2.13 153 Swf Ultisols Tuffac. sandstone

Sodium (mg/g) II9A 1.41 0.03 4 1.41 1.44 1.38 153 Swf Ultisols Tuffac. sandstone

Paullinia pinnata C/N II4 53 48 2 53 86 19 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II4 47 0 2 47 47 47 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II4 1.53 1.39 2 1.53 2.51 0.54 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II4 0.18 0.04 2 0.18 0.21 0.16 135 Lmrf Ultisols Tuffac. sandstone

Paullinia spp. Aluminum (mg/g) II4 0.19 1 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II4 14.33 1 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II4 0.20 1 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II4 4.55 1 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II4 0.23 1 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II4 1.45 1 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II4 1.54 1 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II4 15.20 1 135 Lmrf Ultisols Tuffac. sandstone

Peperomia spp. Aluminum (mg/g) II3 0.27 0.18 5 0.27 0.56 0.11 135 Lmrf Ultisols Tuffac. sandstone

C/N II3 13 1 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II3 7.08 2.31 5 5.84 9.68 4.97 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II3 40 1 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II3 0.23 0.16 5 0.20 0.46 0.08 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II3 4.33 2.60 5 3.37 8.86 2.50 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II3 0.22 0.10 5 0.20 0.35 0.09 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II3 1.76 0.89 5 1.63 3.14 0.82 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II3 1.98 0.83 5 2.09 2.72 0.66 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II3 54.63 24.63 5 42.79 97.78 38.68 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II3 0.31 1 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

193

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Petitia domingensis Aluminum (mg/g) II1A 0.05 1 149 Sdf Mollisols Alluvial deposits Susua forest

Ash (%) II1A 4.76 1 149 Sdf Mollisols Alluvial deposits Susua forest

Calcium (mg/g) II1A 5.98 1 149 Sdf Mollisols Alluvial deposits Susua forest

Iron (mg/g) II1A 0.08 1 149 Sdf Mollisols Alluvial deposits Susua forest

Magnesium (mg/g) II1A 3.95 1 149 Sdf Mollisols Alluvial deposits Susua forest

Manganese (mg/g) II1A 0.05 1 149 Sdf Mollisols Alluvial deposits Susua forest

Nitrogen (%) II1A 1.35 1 149 Sdf Mollisols Alluvial deposits Susua forest

Phosphorus (mg/g) II1A 0.82 1 149 Sdf Mollisols Alluvial deposits Susua forest

Potassium (mg/g) II1A 6.16 1 149 Sdf Mollisols Alluvial deposits Susua forest

Zinc (μg/g) II1A 52.43 1 149 Sdf Mollisols Alluvial deposits Susua forest

Philodendron angustatum Aluminum (mg/g) II4 0.18 0.15 2 0.18 0.28 0.07 135 Lmrf Ultisols Tuffac. sandstone

C/N II4 44 1 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II4 5.89 2.57 2 5.89 7.71 4.07 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II4 42 1 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II4 0.97 1.03 2 0.97 1.70 0.24 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II4 7.31 7.96 2 7.31 12.94 1.68 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II4 1.60 0.84 2 1.60 2.20 1.01 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II4 0.84 0.16 2 0.84 0.95 0.73 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II4 0.67 0.16 2 0.67 0.78 0.56 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II4 8.82 10.33 2 8.82 16.13 1.51 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II4 0.14 1 135 Lmrf Ultisols Tuffac. sandstone

Philodendron scandens Aluminum (mg/g) II4 0.26 0.23 14 0.15 0.73 0.03 135 Lmrf Ultisols Tuffac. sandstone

C/N II4 34 20 12 26 83 17 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II4 7.09 1.89 14 6.61 12.42 5.11 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II4 43 2 12 43 46 40 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II4 0.30 0.37 14 0.16 1.25 0.03 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

194

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Magnesium (mg/g) II4 3.11 4.25 14 1.92 14.80 1.28 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II4 0.45 0.37 14 0.33 1.41 0.14 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II4 1.48 0.60 22 1.41 2.84 0.55 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II4 1.63 1.04 14 1.36 4.47 0.42 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II4 30.07 15.84 14 25.97 59.84 2.27 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II4 0.15 0.05 12 0.15 0.22 0.07 135 Lmrf Ultisols Tuffac. sandstone

Philodendron spp. Aluminum (mg/g) II4 0.82 0.68 16 0.81 2.56 0.08 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II4 7.04 2.25 16 6.76 10.36 3.36 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II4 1.02 0.98 16 0.81 3.04 0.02 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II4 1.91 0.75 16 2.01 3.34 0.81 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II4 0.47 0.21 16 0.39 0.83 0.18 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II4 1.18 0.40 15 1.05 1.85 0.37 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II4 1.49 0.79 16 1.44 2.86 0.26 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II4 34.19 12.77 16 38.14 52.46 8.91 135 Lmrf Ultisols Tuffac. sandstone

Phoradendron racemosum Calcium (mg/g) II3A 26.08 1 104 Sdf Mollisols Alluvial deposits

II3A 26.08 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Magnesium (mg/g) II3A 11.31 1 104 Sdf Mollisols Alluvial deposits

II3A 11.31 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Nitrogen (%) II3A 4.04 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Potassium (mg/g) II3A 33.75 1 104 Sdf Mollisols Alluvial deposits

II3A 33.75 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Phytolacca icosandra Aluminum (mg/g) II4 0.32 0.28 10 0.24 1.04 0.07 135 Lmrf Ultisols Tuffac. sandstone

II4A 2.29 0.42 12 2.39 2.79 1.41 141 Wflm Ultisols Tuffac. sandstone

II4C 0.57 0.40 1 0.38 1.31 0.25 141 Wflm Ultisols Tuffac. sandstone

II4E 1.26 0.70 9 1.07 2.91 0.68 141 Wflm Ultisols Tuffac. sandstone

II4E 0.83 0.37 10 1.74 2.35 1.20 141 Wflm Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

195

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II4F 0.07 1 141 Wflm Ultisols Tuffac. sandstone

Calcium (mg/g) II4 4.38 2.23 10 4.22 7.53 1.69 135 Lmrf Ultisols Tuffac. sandstone

II4A 6.52 2.01 12 6.44 9.86 3.24 141 Wflm Ultisols Tuffac. sandstone

II4C 3.32 1.40 11 3.10 5.66 1.38 141 Wflm Ultisols Tuffac. sandstone

II4E 4.21 1.63 9 4.19 7.47 2.13 141 Wflm Ultisols Tuffac. sandstone

II4E 8.89 3.18 10 9.84 12.80 4.91 141 Wflm Ultisols Tuffac. sandstone

II4F 3.26 1 141 Wflm Ultisols Tuffac. sandstone

Iron (mg/g) II4 0.22 0.33 10 0.11 1.15 0.07 135 Lmrf Ultisols Tuffac. sandstone

II4A 1.66 2.24 12 0.98 8.25 0.33 141 Wflm Ultisols Tuffac. sandstone

II4C 0.30 0.21 1 0.17 0.69 0.11 141 Wflm Ultisols Tuffac. sandstone

II4E 0.97 0.54 9 0.80 2.17 0.49 141 Wflm Ultisols Tuffac. sandstone

II4E 1.39 0.39 10 1.28 1.97 0.92 141 Wflm Ultisols Tuffac. sandstone

II4F 0.08 1 141 Wflm Ultisols Tuffac. sandstone

Magnesium (mg/g) II4 5.88 1.40 10 6.23 7.86 3.24 135 Lmrf Ultisols Tuffac. sandstone

II4A 8.01 2.13 12 8.12 11.24 3.83 141 Wflm Ultisols Tuffac. sandstone

II4C 4.68 2.77 11 3.75 10.80 1.39 141 Wflm Ultisols Tuffac. sandstone

II4E 1.77 0.37 9 1.73 2.44 1.36 141 Wflm Ultisols Tuffac. sandstone

II4E 3.19 1.23 10 2.91 6.06 1.70 141 Wflm Ultisols Tuffac. sandstone

II4F 4.39 1 141 Wflm Ultisols Tuffac. sandstone

Manganese (mg/g) II4 0.20 0.16 10 0.15 0.57 0.05 135 Lmrf Ultisols Tuffac. sandstone

II4A 1.59 0.86 12 1.84 2.55 0.22 141 Wflm Ultisols Tuffac. sandstone

II4C 0.19 0.13 11 0.17 0.51 0.06 141 Wflm Ultisols Tuffac. sandstone

II4E 0.21 0.08 9 0.18 0.39 0.15 141 Wflm Ultisols Tuffac. sandstone

II4E 0.26 0.06 10 0.25 0.38 0.21 141 Wflm Ultisols Tuffac. sandstone

II4F 0.79 1 141 Wflm Ultisols Tuffac. sandstone

Nitrogen (%) II4 1.70 0.68 10 1.55 1.59 0.68 135 Lmrf Ultisols Tuffac. sandstone

II4A 3.63 1.55 12 3.03 7.03 2.07 141 Wflm Ultisols Tuffac. sandstone

II4C 0.98 0.64 11 0.69 2.48 0.50 141 Wflm Ultisols Tuffac. sandstone

II4E 0.69 0.13 9 0.65 0.91 0.54 141 Wflm Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

196

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II4E 0.93 0.20 10 0.90 1.26 0.69 141 Wflm Ultisols Tuffac. sandstone

II4F 1.92 1 141 Wflm Ultisols Tuffac. sandstone

Phosphorus (mg/g) II4 0.81 0.58 10 0.71 2.05 0.19 135 Lmrf Ultisols Tuffac. sandstone

II4A 2.41 0.91 12 2.22 4.33 1.33 141 Wflm Ultisols Tuffac. sandstone

II4C 1.19 0.53 11 0.95 2.27 0.66 141 Wflm Ultisols Tuffac. sandstone

II4E 1.27 0.57 9 1.25 2.49 0.61 141 Wflm Ultisols Tuffac. sandstone

II4E 2.00 0.91 10 1.82 3.25 0.76 141 Wflm Ultisols Tuffac. sandstone

II4F 3.80 1 141 Wflm Ultisols Tuffac. sandstone

Potassium (mg/g) II4 58.12 19.88 10 60.75 89.46 26.24 135 Lmrf Ultisols Tuffac. sandstone

II4A 13.99 2.65 12 14.67 17.24 6.41 141 Wflm Ultisols Tuffac. sandstone

II4C 14.00 2.53 11 15.10 16.33 8.15 141 Wflm Ultisols Tuffac. sandstone

II4E 14.13 4.06 9 15.77 17.63 6.46 141 Wflm Ultisols Tuffac. sandstone

II4E 9.76 4.25 10 7.40 16.20 6.68 141 Wflm Ultisols Tuffac. sandstone

II4F 16.71 1 141 Wflm Ultisols Tuffac. sandstone

Phytolacca rivinoides Aluminum (mg/g) II4 0.14 0.07 3 0.14 0.21 0.06 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II4 1.71 0.70 3 2.04 2.17 0.91 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II4 0.13 0.08 3 0.16 0.20 0.04 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II4 3.82 1.68 3 3.43 5.57 2.36 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II4 0.09 0.04 3 0.09 0.14 0.05 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II4 1.25 1.04 3 0.92 2.42 0.41 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II4 0.83 0.86 3 0.47 1.82 0.21 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II4 36.57 17.43 3 41.82 50.76 17.12 135 Lmrf Ultisols Tuffac. sandstone

Pictetia aculeata Aluminum (mg/kg) II1A 40 21 14 41 82 12 151 Sdf Mollisols Alluvial deposits

II1A 60 1 102 Smf Limestone Tuffac. sandstone Old

Ash (%) II1A 10.82 1 102 Smf Limestone Tuffac. sandstone Old

II1A 7.57 1.67 11 8.03 8.61 2.68 151 Sdf Mollisols Alluvial deposits

C/N II1A 18 1 13 18 20 16 151 Sdf Mollisols Alluvial deposits
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

197

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Calcium (mg/g) II1A 20.53 1 102 Smf Limestone Tuffac. sandstone

II1A 20.86 4.07 14 22.21 24.07 9.26 151 Sdf Mollisols Alluvial deposits

II1Ci 34.71 1 104 Sdf Mollisols Alluvial deposits

II1Ci 34.70 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Carbon (%) II1A 52 1 13 52 53 51 151 Sdf Mollisols Alluvial deposits

Cobalt (μg/g) II1A 3.00 1 102 Smf Limestone Tuffac. sandstone Old

Iron (mg/kg) II1A 50 13 14 50 81 23 151 Sdf Mollisols Alluvial deposits

II1A 83 1 102 Smf Limestone Tuffac. sandstone Old

Magnesium (mg/g) II1A 2.55 1 104 Sdf Mollisols Alluvial deposits

II1A 2.82 1 102 Smf Limestone Tuffac. sandstone Old

II1A 2.55 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1A 2.12 0.44 14 2.14 3.02 0.92 151 Sdf Mollisols Alluvial deposits

II1Ci 1.04 1 104 Sdf Mollisols Alluvial deposits

II1Ci 1.04 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Manganese (mg/kg) II1A 42 11 14 41 66 16 151 Sdf Mollisols Alluvial deposits

II1A 11 1 102 Smf Limestone Tuffac. sandstone Old

Nickel (μg/g) II1A 14.45 1 102 Smf Limestone Tuffac. sandstone Old

Nitrogen (%) II1A 0.86 1 102 Smf Limestone Tuffac. sandstone Old

II1A 2.15 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1A 2.85 0.17 13 2.87 3.16 2.54 151 Sdf Mollisols Alluvial deposits

II1Ci 1.33 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Phosphorus (mg/g) II1A 0.82 1 104 Sdf Mollisols Alluvial deposits

II1A 0.40 1 102 Smf Limestone Tuffac. sandstone Old

II1A 0.82 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1A 0.69 0.14 14 0.71 0.81 0.26 151 Sdf Mollisols Alluvial deposits

II1Ci 0.12 1 104 Sdf Mollisols Alluvial deposits

II1Ci 0.12 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Potassium (mg/g) II1A 17.63 1 104 Sdf Mollisols Alluvial deposits

II1A 4.75 1 102 Smf Limestone Tuffac. sandstone Old
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

198

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 17.63 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1A 7.27 1.47 14 7.52 8.63 2.97 151 Sdf Mollisols Alluvial deposits

II1Ci 6.65 1 104 Sdf Mollisols Alluvial deposits

II1Ci 6.65 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Sulfur (%) II1A 0.27 0.02 13 0.27 0.30 0.22 151 Sdf Mollisols Alluvial deposits

Pilea inaequalis Aluminum (mg/g) II3 1.89 2.00 42 1.33 10.17 0.05 135 Lmrf Ultisols Tuffac. sandstone

C/N II3 21 3 20 20 26 17 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II3 29.65 10.24 42 28.53 60.43 1.24 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II3 36 1 20 36 39 34 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II3 2.23 2.89 42 1.10 14.02 0.20 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II3 7.79 2.41 42 7.35 12.92 0.84 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II3 0.24 0.12 42 0.23 0.66 0.02 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II3 1.70 0.35 53 1.75 2.34 0.48 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II3 1.18 0.34 42 1.16 1.98 0.27 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II3 34.91 16.71 42 31.90 108.39 4.54 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II3 0.51 0.21 20 0.47 1.13 0.22 135 Lmrf Ultisols Tuffac. sandstone

Pilea krugii Aluminum (mg/g) II3 1.49 0.01 2 1.49 1.50 1.48 135 Lmrf Ultisols Tuffac. sandstone

C/N II3 16 1 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II3 22.39 0.57 2 22.39 22.43 22.35 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II3 38 1 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II3 1.19 0.02 2 1.19 1.20 1.17 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II3 6.62 0.08 2 6.62 6.68 6.56 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II3 0.45 0.00 2 0.45 0.45 0.45 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II3 1.78 0.06 3 1.46 2.43 1.45 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II3 1.34 0.01 2 1.34 1.35 1.33 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II3 29.79 0.07 2 29.79 29.84 29.74 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II3 0.32 1 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

199

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Pimenta racemosa Ash (%) II2A 7.92 1.10 5 7.75 9.66 6.77 126 Ultisols Siltstone,sandstone Urban forest, RP

II2C 1.63 0.43 5 1.59 2.06 0.98 126 Ultisols Siltstone,sandstone Urban forest, RP

II2Ci 10.28 3.43 6 10.19 15.17 5.57 126 Ultisols Siltstone,sandstone Urban forest, RP

C/N II2A 59 5 5 59 64 52 126 Ultisols Siltstone,sandstone Urban forest, RP

II2C 425 201 5 339 777 287 126 Ultisols Siltstone,sandstone Urban forest, RP

II2Ci 102 26 6 90 136 76 126 Ultisols Siltstone,sandstone Urban forest, RP

Carbon (%) II2A 53 1 5 53 54 51 126 Ultisols Siltstone,sandstone Urban forest, RP

II2C 54 1 5 54 54 52 126 Ultisols Siltstone,sandstone Urban forest, RP

II2Ci 51 2 6 51 54 49 126 Ultisols Siltstone,sandstone Urban forest, RP

Nitrogen (%) II2A 0.90 0.09 5 0.88 1.04 0.83 126 Ultisols Siltstone,sandstone Urban forest, RP

II2C 0.14 0.05 5 0.16 0.19 0.07 126 Ultisols Siltstone,sandstone Urban forest, RP

II2Ci 0.53 0.11 6 0.56 0.65 0.39 126 Ultisols Siltstone,sandstone Urban forest, RP

Sulfur (%) II2A 0.16 0.02 5 0.16 0.20 0.15 126 Ultisols Siltstone,sandstone Urban forest, RP

II2C 0.03 0.01 5 0.03 0.05 0.02 126 Ultisols Siltstone,sandstone Urban forest, RP

II2Ci 0.14 0.07 6 0.12 0.23 0.07 126 Ultisols Siltstone,sandstone Urban forest, RP

Wood density (g/cc) II2C 0.67 0.07 5 0.71 0.72 0.58 126 Ultisols Siltstone,sandstone Urban forest, RP

Pine plantation Aluminum (mg/g) II9A 5.59 0.66 3 5.86 6.07 4.84 153 Swf Ultisols Tuffac. sandstone

Ash (%) II9A 7.12 0.16 3 7.15 7.27 6.95 153 Swf Ultisols Tuffac. sandstone

C/N II9A 70 1 3 69 70 69 153 Swf Ultisols Tuffac. sandstone

Calcium (mg/g) II8A 14.64 4.03 122 14.92 25.64 8.50 156 Swf Ultisols Tuffac. sandstone Other

II8A 5.44 1.44 131 5.21 16.80 4.12 156 Swf Ultisols Tuffac. sandstone Pine leaves

II8B 3.97 2.77 130 3.15 19.03 1.30 156 Swf Ultisols Tuffac. sandstone <2cm diameter

II8B 2.73 1.91 33 2.23 8.25 1.22 156 Swf Ultisols Tuffac. sandstone <2cm diameter

II8C 2.35 3.04 121 1.87 29.76 0.68 156 Swf Ultisols Tuffac. sandstone

II8D 6.84 2.35 128 6.43 13.89 3.40 156 Swf Ultisols Tuffac. sandstone

II9A 6.66 0.93 3 6.16 7.73 6.09 153 Swf Ultisols Tuffac. sandstone

II9A 14.20 2.48 129 14.18 21.64 7.99 156 Swf Ultisols Tuffac. sandstone Other
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

200

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II9A 3.96 0.66 120 3.97 5.04 0.95 156 Swf Ultisols Tuffac. sandstone Pine leaves

II9B 3.55 1.95 64 3.08 10.07 1.09 156 Swf Ultisols Tuffac. sandstone

II9C 4.00 3.58 50 2.58 15.18 0.25 156 Swf Ultisols Tuffac. sandstone

II9D 8.18 1.98 48 8.09 13.77 3.57 156 Swf Ultisols Tuffac. sandstone

Carbon (%) II9A 52 1 3 53 53 52 153 Swf Ultisols Tuffac. sandstone

Iron (mg/g) II9A 3.06 0.31 3 3.08 3.35 2.74 153 Swf Ultisols Tuffac. sandstone

Magnesium (mg/g) II8A 3.30 1.15 122 2.92 7.02 2.64 156 Swf Ultisols Tuffac. sandstone Other

II8A 1.51 0.29 131 1.44 3.42 1.20 156 Swf Ultisols Tuffac. sandstone Pine leaves

II8B 1.05 0.73 130 0.85 5.81 0.46 156 Swf Ultisols Tuffac. sandstone <2cm diameter

II8B 0.80 0.63 33 0.63 3.22 0.40 156 Swf Ultisols Tuffac. sandstone <2cm diameter

II8C 1.08 0.84 121 0.89 7.75 0.49 156 Swf Ultisols Tuffac. sandstone

II8D 1.98 0.52 128 4.88 4.24 1.50 156 Swf Ultisols Tuffac. sandstone

II9A 1.87 0.05 3 1.89 1.90 1.81 153 Swf Ultisols Tuffac. sandstone

II9A 3.56 0.81 129 3.52 5.95 1.03 156 Swf Ultisols Tuffac. sandstone Other

II9A 1.19 0.17 120 1.20 1.63 0.27 156 Swf Ultisols Tuffac. sandstone Pine leaves

II9B 0.91 0.54 64 0.73 2.75 0.27 156 Swf Ultisols Tuffac. sandstone

II9C 1.79 1.16 50 1.49 4.84 0.56 156 Swf Ultisols Tuffac. sandstone

II9D 2.15 0.46 48 2.10 3.61 1.40 156 Swf Ultisols Tuffac. sandstone

Manganese (mg/g) II9A 0.79 0.03 3 0.79 0.82 0.76 153 Swf Ultisols Tuffac. sandstone

Nitrogen (%) II8A 1.59 0.35 116 1.59 2.51 2.04 156 Swf Ultisols Tuffac. sandstone Other

II8A 0.58 0.13 124 0.56 1.37 0.59 156 Swf Ultisols Tuffac. sandstone Pine leaves

II8B 0.44 0.29 122 0.34 1.95 0.16 156 Swf Ultisols Tuffac. sandstone <2cm diameter

II8B 0.35 0.26 31 0.27 1.13 0.10 156 Swf Ultisols Tuffac. sandstone <2cm diameter

II8C 0.57 0.25 117 0.52 1.92 0.29 156 Swf Ultisols Tuffac. sandstone

II8D 1.14 0.26 122 1.10 2.03 0.97 156 Swf Ultisols Tuffac. sandstone

II9A 0.75 0.01 3 0.75 0.76 0.75 153 Swf Ultisols Tuffac. sandstone

II9A 1.63 0.31 115 1.58 2.47 0.99 156 Swf Ultisols Tuffac. sandstone Other

II9A 0.46 0.06 114 0.40 0.56 0.28 156 Swf Ultisols Tuffac. sandstone Pine leaves

II9B 0.44 0.32 55 0.33 1.64 0.08 156 Swf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

201

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II9C 0.89 0.34 48 0.87 1.61 0.27 156 Swf Ultisols Tuffac. sandstone

II9D 1.36 0.23 48 1.36 1.90 0.78 156 Swf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II8A 0.55 0.16 122 0.53 1.19 0.59 156 Swf Ultisols Tuffac. sandstone Other

II8A 0.23 0.09 131 0.22 1.12 0.19 156 Swf Ultisols Tuffac. sandstone Pine leaves

II8B 0.19 0.13 130 0.16 0.89 0.04 156 Swf Ultisols Tuffac. sandstone <2cm diameter

II8B 0.14 0.10 33 0.12 0.42 0.04 156 Swf Ultisols Tuffac. sandstone <2cm diameter

II8C 0.27 0.16 121 0.22 1.29 0.08 156 Swf Ultisols Tuffac. sandstone

II8D 0.52 0.12 128 0.51 0.96 0.45 156 Swf Ultisols Tuffac. sandstone

II9A 0.24 0.01 3 0.24 0.25 0.24 153 Swf Ultisols Tuffac. sandstone

II9A 0.59 0.17 129 0.60 1.33 0.24 156 Swf Ultisols Tuffac. sandstone Other

II9A 0.16 0.04 120 0.17 0.27 0.08 156 Swf Ultisols Tuffac. sandstone Pine leaves

II9B 0.20 0.24 64 0.15 1.66 0.04 156 Swf Ultisols Tuffac. sandstone

II9C 0.59 0.31 50 0.55 1.40 0.14 156 Swf Ultisols Tuffac. sandstone

II9D 0.80 0.27 48 0.72 1.89 0.39 156 Swf Ultisols Tuffac. sandstone

Potassium (mg/g) II8A 2.90 1.63 122 2.46 9.27 1.39 156 Swf Ultisols Tuffac. sandstone Other

II8A 1.09 0.49 131 0.96 5.15 0.72 156 Swf Ultisols Tuffac. sandstone Pine leaves

II8B 0.70 0.72 130 0.47 5.71 0.25 156 Swf Ultisols Tuffac. sandstone <2cm diameter

II8B 0.46 0.31 33 0.34 1.50 0.23 156 Swf Ultisols Tuffac. sandstone <2cm diameter

II8C 0.99 1.06 121 0.66 8.29 0.24 156 Swf Ultisols Tuffac. sandstone

II8D 1.79 1.92 128 1.16 10.21 0.73 156 Swf Ultisols Tuffac. sandstone

II9A 1.78 0.04 3 1.55 1.82 1.75 153 Swf Ultisols Tuffac. sandstone

II9A 5.40 2.00 129 5.20 10.75 1.88 156 Swf Ultisols Tuffac. sandstone Other

II9A 1.63 0.49 120 1.57 3.02 0.37 156 Swf Ultisols Tuffac. sandstone Pine leaves

II9B 0.81 0.93 64 0.55 6.21 0.11 156 Swf Ultisols Tuffac. sandstone

II9C 3.83 2.11 50 3.70 8.82 0.62 156 Swf Ultisols Tuffac. sandstone

II9D 3.21 1.88 48 2.80 13.88 1.76 156 Swf Ultisols Tuffac. sandstone

Sodium (mg/g) II9A 1.39 0.05 3 1.37 1.44 1.36 153 Swf Ultisols Tuffac. sandstone

Pinus caribaea Ash (%) II1A 2.93 1 101 Swf Ultisols Tuffac. sandstone Overstory
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

202

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8A 10.88 7.65 4 7.68 22.21 5.95 90 Swf Ultisols Tuffac. sandstone

II8A 3.19 0.39 7 3.30 3.74 2.63 90 Swf Ultisols Tuffac. sandstone Whole

II8A 8.24 0.31 2 8.24 8.46 8.02 90 Swf Ultisols Tuffac. sandstone Old

II8A 2.52 0.04 3 2.51 2.56 2.49 90 Swf Ultisols Tuffac. sandstone Recently 

II8A 7.70 5.03 6 4.96 15.31 3.51 90 Swf Ultisols Tuffac. sandstone Torn

II8B 3.92 5.82 8 1.89 18.31 1.64 90 Swf Ultisols Tuffac. sandstone

II8B 6.34 3.26 6 6.50 10.15 2.18 90 Swf Ultisols Tuffac. sandstone

II8C 0.71 0.36 3 0.71 0.96 0.45 90 Swf Ultisols Tuffac. sandstone Pine seeds

II8D 8.06 4.49 13 6.17 14.71 1.18 90 Swf Ultisols Tuffac. sandstone

II8D 10.17 4.03 4 11.24 13.56 5.71 90 Swf Ultisols Tuffac. sandstone

Calcium (mg/g) II1B 1.03 0.10 6 1.04 1.13 0.84 69 Swf Ultisols Tuffac. sandstone Fresh

II8A 4.89 0.63 4 4.67 5.80 4.41 90 Swf Ultisols Tuffac. sandstone

II8A 4.83 0.77 7 4.57 5.87 3.87 90 Swf Ultisols Tuffac. sandstone Whole

II8A 1.38 0.20 5 1.48 1.54 1.10 69 Swf Ultisols Tuffac. sandstone 6 months

II8A 1.38 0.18 5 1.48 1.55 1.13 69 Swf Ultisols Tuffac. sandstone 12 months

II8A 22.99 3.56 2 22.99 25.51 20.48 90 Swf Ultisols Tuffac. sandstone Old

II8A 5.11 0.95 3 4.63 6.20 4.50 90 Swf Ultisols Tuffac. sandstone Recently

II8A 5.68 0.84 6 5.95 6.50 4.64 90 Swf Ultisols Tuffac. sandstone Torn

II8B 3 .63 0.81 8 3.75 4.93 2.51 90 Swf Ultisols Tuffac. sandstone

II8B 2.10 0.49 5 2.17 2.5 1.29 69 Swf Ultisols Tuffac. sandstone 18 months

II8B 3.72 1.48 6 3.26 6.06 1.96 90 Swf Ultisols Tuffac. sandstone

II8C 1.74 1.38 3 1.30 3.29 0.63 90 Swf Ultisols Tuffac. sandstone Pine seeds

II8C 2.23 1.16 4 1.90 3.81 1.33 69 Swf Ultisols Tuffac. sandstone 24 months

II8D 1.81 0.23 4 1.90 1.96 1.47 69 Swf Ultisols Tuffac. sandstone 30 months

II8D 7.36 3.63 4 6.01 12.72 4.72 90 Swf Ultisols Tuffac. sandstone

lI8D 4.88 1.82 13 4.60 8.90 2.44 90 Swf Ultisols Tuffac. sandstone

Magnesium (mg/g) II1B 0.59 0.06 6 0.61 0.64 0.50 69 Swf Ultisols Tuffac. sandstone Fresh

II7B 0.48 0.08 5 0.51 0.53 0.33 69 Swf Ultisols Tuffac. sandstone 6 months

II7B 0.53 0.07 5 0.55 0.62 0.42 69 Swf Ultisols Tuffac. sandstone 12 months
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

203

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II7B 0.65 0.11 5 0.68 0.72 0.45 69 Swf Ultisols Tuffac. sandstone 18 months

II7B 0.75 0.29 4 0.64 1.17 0.55 69 Swf Ultisols Tuffac. sandstone 24 months

II7B 0.52 0.09 4 0.49 0.64 0.44 69 Swf Ultisols Tuffac. sandstone 30 months

II8A 1.67 0.13 4 1.69 1.81 1.50 90 Swf Ultisols Tuffac. sandstone

II8A 1.47 0.44 7 1.25 2.22 1.10 90 Swf Ultisols Tuffac. sandstone Whole

II8A 1.20 0.29 8 1.18 1.67 0.75 90 Swf Ultisols Tuffac. sandstone

II8A 5.85 1.83 2 5.85 7.15 4.56 90 Swf Ultisols Tuffac. sandstone Old

II8A 1.76 0.20 3 1.65 2.00 1.64 90 Swf Ultisols Tuffac. sandstone Recently

II8A 1.99 0.32 6 2.02 2.40 1.41 90 Swf Ultisols Tuffac. sandstone Torn

II8B 1.14 0.41 6 0.98 1.90 0.78 90 Swf Ultisols Tuffac. sandstone Loose litter

II8C 1.04 0.45 3 1.06 1.47 0.58 90 Swf Ultisols Tuffac. sandstone

II8D 1.85 0.74 13 1.83 3.58 0.80 90 Swf Ultisols Tuffac. sandstone

II8D 2.18 0.75 4 2.04 3.21 1.43 90 Swf Ultisols Tuffac. sandstone

Nitrogen (%) II1A 0.88 1 101 Swf Ultisols Tuffac. sandstone Overstory

II1B 0.13 0.02 6 0.13 0.15 0.11 69 Swf Ultisols Tuffac. sandstone Fresh

II7B 0.11 0.03 5 0.11 0.15 0.08 69 Swf Ultisols Tuffac. sandstone 6months

II7B 0.15 0.03 5 0.14 0.19 0.12 69 Swf Ultisols Tuffac. sandstone 12 months

II7B 0.22 0.05 5 0.22 0.29 0.16 69 Swf Ultisols Tuffac. sandstone 18 months

II7B 0.24 0.07 4 0.24 0.31 0.17 69 Swf Ultisols Tuffac. sandstone 24 months

II7B 0.22 0.03 4 0.22 0.24 0.19 69 Swf Ultisols Tuffac. sandstone 30 months

II8A 1.01 0.13 4 1.00 1.15 0.88 90 Swf Ultisols Tuffac. sandstone

II8A 0.47 0.12 7 0.49 0.65 0.32 90 Swf Ultisols Tuffac. sandstone Whole

II8A 1.32 0.12 2 1.32 1.40 1.23 90 Swf Ultisols Tuffac. sandstone Old

II8A 0.40 0.05 3 0.38 0.46 0.36 90 Swf Ultisols Tuffac. sandstone Recently 

II8A 0.95 0.23 6 0.97 1.28 0.60 90 Swf Ultisols Tuffac. sandstone Torn 

II8B 0.52 0.22 8 0.45 0.88 0.24 90 Swf Ultisols Tuffac. sandstone

II8B 0.60 0.19 6 0.64 0.85 0.28 90 Swf Ultisols Tuffac. sandstone

II8C 0.67 0.44 3 0.85 0.98 0.17 90 Swf Ultisols Tuffac. sandstone Pine seeds

II8D 0.91 0.41 13 1.06 1.51 0.23 90 Swf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

204

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8D 1.20 0.20 4 1.21 1.40 1.00 90 Swf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.78 1 101 Swf Ultisols Tuffac. sandstone Overstory

II1B 0.17 0.02 6 0.16 0.20 0.14 69 Swf Ultisols Tuffac. sandstone Fresh

II7B 0.11 0.03 5 0.09 0.17 0.09 69 Swf Ultisols Tuffac. sandstone 6months

II7B 0.10 0.02 5 0.10 0.12 0.09 69 Swf Ultisols Tuffac. sandstone 12 months

II7B 0.15 0.04 5 0.16 0.18 0.09 69 Swf Ultisols Tuffac. sandstone 18 months

II7B 0.13 0.03 4 0.13 0.16 0.09 69 Swf Ultisols Tuffac. sandstone 24 months

II7B 0.10 0.02 4 0.10 0.11 0.08 69 Swf Ultisols Tuffac. sandstone 30 months

II8A 0.49 0.07 4 0.47 0.58 0.43 90 Swf Ultisols Tuffac. sandstone

II8A 0.19 0.08 7 0.19 0.34 0.11 90 Swf Ultisols Tuffac. sandstone Whole

II8A 0.56 0.03 2 0.56 0.57 0.54 90 Swf Ultisols Tuffac. sandstone Old

II8A 0.24 0.04 3 0.23 0.28 0.21 90 Swf Ultisols Tuffac. sandstone Recently 

II8A 0.45 0.10 6 0.47 0.55 0.30 90 Swf Ultisols Tuffac. sandstone Torn 

II8B 0.21 0.10 8 0.18 0.42 0.11 90 Swf Ultisols Tuffac. sandstone

II8B 0.28 0.08 6 0.29 0.35 0.13 90 Swf Ultisols Tuffac. sandstone

II8C 0.29 0.20 3 0.24 0.51 0.11 90 Swf Ultisols Tuffac. sandstone Pine seeds

II8D 0.44 0.12 13 0.48 0.61 0.21 90 Swf Ultisols Tuffac. sandstone

II8D 0.58 0.07 4 0.58 0.66 0.51 90 Swf Ultisols Tuffac. sandstone

Potassium (mg/g) II1A 2.00 1 101 Swf Ultisols Tuffac. sandstone Overstory

II1B 0.73 0.09 6 0.76 0.85 0.59 69 Swf Ultisols Tuffac. sandstone Fresh

II7B 0.80 0.11 5 0.84 0.93 0.64 69 Swf Ultisols Tuffac. sandstone 6months

II7B 1.16 0.75 5 0.94 2.00 0.42 69 Swf Ultisols Tuffac. sandstone 12 months

II7B 0.68 0.32 5 0.54 1.04 0.34 69 Swf Ultisols Tuffac. sandstone 18 months

II7B 1.40 0.61 4 1.29 2.14 0.90 69 Swf Ultisols Tuffac. sandstone 24 months

II7B 0.37 0.3 4 0.23 0.81 0.20 69 Swf Ultisols Tuffac. sandstone 30 months

II8A 1.76 0.33 4 1.81 2.08 1.35 90 Swf Ultisols Tuffac. sandstone

II8A 1.96 0.53 7 1.87 3.06 1.71 90 Swf Ultisols Tuffac. sandstone Whole

II8A 2.38 0.06 2 2.38 2.42 2.34 90 Swf Ultisols Tuffac. sandstone Old

II8A 2.56 1.57 3 2.04 4.32 1.31 90 Swf Ultisols Tuffac. sandstone Recently 
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

205

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8A 1.42 0.60 6 1.35 2.22 0.78 90 Swf Ultisols Tuffaceoussandstone Torn 

II8B 1.81 0.47 8 1.94 2.41 1.20 90 Swf Ultisols Tuffac. sandstone

II8B 2.23 0.57 6 2.24 2.86 1.49 90 Swf Ultisols Tuffac. sandstone

II8C 1.67 1.11 3 1.16 2.94 0.90 90 Swf Ultisols Tuffac. sandstone Pine seeds

II8D 2.19 0.8 13 2.09 3.74 0.94 90 Swf Ultisols Tuffac. sandstone

II8D 2.24 0.54 4 2.10 2.97 1.78 90 Swf Ultisols Tuffac. sandstone

Wood density (g/cc) II1C 0.49 0.03 18 0.49 0.53 0.44 115

Pinus caribaea hondurensis Calcium (mg/g) II9A 3.25 0.45 22 3.17 4.21 2.25 139 Wfs Ultisols Tuffac. sandstone

II9A 11.03 2.29 20 10.79 16.45 8.54 139 Wfs Ultisols Tuffac. sandstone Other

II9B 2.52 0.79 5 2.31 3.81 1.66 139 Wfs Ultisols Tuffac. sandstone

II9C 1.65 1.01 6 1.37 3.54 0.81 139 Wfs Ultisols Tuffac. sandstone

II9D 4.64 1.45 5 4.62 6.05 2.35 139 Wfs Ultisols Tuffac. sandstone

Magnesium (mg/g) II9A 1.21 0.18 22 1.17 1.71 0.94 139 Wfs Ultisols Tuffac. sandstone

II9A 5.67 1.43 20 5.75 8.29 3.53 139 Wfs Ultisols Tuffac. sandstone Other

II9B 0.81 0.16 5 0.87 0.94 0.55 139 Wfs Ultisols Tuffac. sandstone

II9C 1.18 0.17 6 1.19 1.47 0.97 139 Wfs Ultisols Tuffac. sandstone

II9D 2.13 0.28 5 2.24 2.44 1.71 139 Wfs Ultisols Tuffac. sandstone

Nitrogen (%) II9A 0.37 0.06 3 0.37 0.52 0.24 139 Wfs Ultisols Tuffac. sandstone

II9A 1.75 0.34 21 1.74 2.42 1.05 139 Wfs Ultisols Tuffac. sandstone Other

II9B 0.32 0.18 10 0.26 0.66 0.14 139 Wfs Ultisols Tuffac. sandstone

II9C 0.57 0.18 11 0.56 0.91 0.28 139 Wfs Ultisols Tuffac. sandstone

II9D 1.24 0.61 9 1.16 2.80 0.65 139 Wfs Ultisols Tuffac. sandstone

Phosphorus (mg/g) II9A 0.18 0.06 22 0.18 0.30 0.04 139 Wfs Ultisols Tuffac. sandstone

II9A 0.86 0.18 20 0.86 1.28 0.43 139 Wfs Ultisols Tuffac. sandstone Other

II9B 0.18 0.10 5 0.15 0.29 0.08 139 Wfs Ultisols Tuffac. sandstone

II9C 0.48 0.30 6 0.46 0.85 0.08 139 Wfs Ultisols Tuffac. sandstone

II9D 0.74 0.11 5 0.76 0.87 0.57 139 Wfs Ultisols Tuffac. sandstone

Potassium (mg/g) II9A 1.84 0.38 22 1.82 2.74 0.97 139 Wfs Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

206

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II9A 7.51 2.23 20 7.99 12.50 2.81 139 Wfs Ultisols Tuffac. sandstone Other

II9B 1.46 0.53 5 1.14 2.10 1.02 139 Wfs Ultisols Tuffac. sandstone

II9C 1.74 0.92 6 1.87 2.79 0.51 139 Wfs Ultisols Tuffac. sandstone

II9D 3.20 0.66 5 2.97 3.93 2.46 139 Wfs Ultisols Tuffac. sandstone

Pinus elliottii Calcium (mg/g) II9A 2.99 0.36 22 2.98 3.61 2.39 139 Wfs Ultisols Tuffac. sandstone

II9A 9.28 4.16 22 8.37 19.51 3.22 139 Wfs Ultisols Tuffac. sandstone Other

II9B 3.08 1.56 6 2.68 5.72 1.68 139 Wfs Ultisols Tuffac. sandstone

II9C 1.13 0.87 5 0.65 2.56 0.52 139 Wfs Ultisols Tuffac. sandstone

II9D 4.48 1.92 6 4.00 8.25 2.78 139 Wfs Ultisols Tuffac. sandstone

Magnesium (mg/g) II9A 1.65 0.32 22 1.60 2.51 1.12 139 Wfs Ultisols Tuffac. sandstone

II9A 5.20 2.22 22 4.72 11.38 1.60 139 Wfs Ultisols Tuffac. sandstone Other

II9B 0.61 0.15 6 0.61 0.83 0.37 139 Wfs Ultisols Tuffac. sandstone

II9C 1.02 0.48 5 0.88 1.82 0.59 139 Wfs Ultisols Tuffac. sandstone

II9D 2.47 1.97 6 1.81 6.48 1.36 139 Wfs Ultisols Tuffac. sandstone

Nitrogen (%) II9A 0.38 0.13 22 0.36 0.90 0.28 139 Wfs Ultisols Tuffac. sandstone

II9A 1.34 0.31 22 1.37 2.02 0.78 139 Wfs Ultisols Tuffac. sandstone Other

II9B 0.30 0.10 11 0.32 0.49 0.12 139 Wfs Ultisols Tuffac. sandstone

II9C 1.15 1.04 10 0.84 3.43 0.12 139 Wfs Ultisols Tuffac. sandstone

II9D 0.83 0.20 11 0.83 1.24 0.57 139 Wfs Ultisols Tuffac. sandstone

Phosphorus (mg/g) II9A 0.22 0.05 22 0.22 0.31 0.14 139 Wfs Ultisols Tuffac. sandstone

II9A 0.52 0.16 22 0.54 0.78 0.18 139 Wfs Ultisols Tuffac. sandstone Other

II9B 0.14 0.08 6 0.15 0.24 0.02 139 Wfs Ultisols Tuffac. sandstone

II9C 0.37 0.16 5 0.34 0.61 0.21 139 Wfs Ultisols Tuffac. sandstone

II9D 0.49 0.16 6 0.49 0.66 0.27 139 Wfs Ultisols Tuffac. sandstone

Potassium (mg/g) II9A 1.94 0.37 22 1.90 2.57 1.09 139 Wfs Ultisols Tuffac. sandstone

II9A 5.21 2.14 22 4.75 9.56 1.15 139 Wfs Ultisols Tuffac. sandstone Other

II9B 1.01 0.28 6 1.02 1.44 0.62 139 Wfs Ultisols Tuffac. sandstone

II9C 1.81 0.77 5 1.47 2.91 1.09 139 Wfs Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

207

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II9D 2.28 0.43 6 2.27 2.76 1.64 139 Wfs Ultisols Tuffac. sandstone

Pinus massoniana Ash (%) II1B 1.27 0.52 2 1.27 1.63 1.70 68 Swf Ultisols Tuffac. sandstone Class 3-4 LD

II1B 1.04 0.23 2 1.04 1.20 0.87 68 Swf Ultisols Tuffac. sandstone Class 2 SD

II1B 1.22 0.72 6 0.97 2.65 0.74 68 Swf Ultisols Tuffac. sandstone Class 3 SD

Calcium (mg/g) II1B 2.15 0.07 2 2.15 2.20 2.10 68 Swf Ultisols Tuffac. sandstone Class 3 1 LD

II1B 2.83 0.66 2 2.83 3.29 2.36 68 Swf Ultisols Tuffac. sandstone Class 2 SD

II1B 1.98 0.72 6 2.05 3.19 1.10 68 Swf Ultisols Tuffac. sandstone Class 3 SD

Nitrogen (%) II1B 0.47 0.04 2 0.47 0.50 0.44 68 Swf Ultisols Tuffac. sandstone Class 3-4 LD

II1B 0.25 0.03 2 0.25 0.27 0.23 68 Swf Ultisols Tuffac. sandstone Class 2 SD

II1B 0.40 0.35 6 0.30 1.04 0.12 68 Swf Ultisols Tuffac. sandstone Class 3 SD

Phosphorus (mg/g) II1B 0.19 0.02 2 0.19 0.21 0.18 68 Swf Ultisols Tuffac. sandstone Class 3-4 LD

II1B 0.07 0.00 2 0.07 0.07 0.07 68 Swf Ultisols Tuffac. sandstone Class 2 SD

II1B 0.12 0.08 6 0.09 0.26 0.06 68 Swf Ultisols Tuffac. sandstone Class 3 SD

Potassium (mg/g) II1B 0.13 1 68 Swf Ultisols Tuffac. sandstone Class 3-4 LD

II1B 0.37 0.06 2 0.37 0.41 0.32 68 Swf Ultisols Tuffac. sandstone Class 2 SD

II1B 0.42 0.26 6 0.31 0.91 0.23 68 Swf Ultisols Tuffac. sandstone Class 3 SD

Pinus ocarpa Wood density (g/cc) II1C 0.41 0.04 3 0.39 0.46 0.38 115

Piper aduncum Aluminum (mg/g) II2A 0.71 1 129 Lmrf Ultisols Tuffac. sandstone

Ash (%) II2A 14.62 1 129 Lmrf Ultisols Tuffac. sandstone

C/N II2A 15 1 129 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II2A 13.95 1 129 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II2A 48 1 129 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II2A 0.61 1 129 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II2A 2.10 1 129 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II2A 0.09 1 129 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II2A 3.22 1 129 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

208

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Phosphorus (mg/g) II2A 0.91 1 129 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II2A 14.85 1 129 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II2A 0.50 1 129 Lmrf Ultisols Tuffac. sandstone

Piper glabrescens Aluminum (mg/g) II2A 0.19 0.12 14 0.16 0.55 0.05 135 Lmrf Ultisols Tuffac. sandstone

II2B 0.06 0.01 4 0.05 0.07 0.04 135 Lmrf Ultisols Tuffac. sandstone

II2C 0.06 0.03 14 0.05 0.13 0.03 135 Lmrf Ultisols Tuffac. sandstone

II2E 2.24 0.60 9 2.17 3.33 1.18 135 Lmrf Ultisols Tuffac. sandstone

II2H 0.24 0.10 8 0.25 0.37 0.11 135 Lmrf Ultisols Tuffac. sandstone

II8A 0.96 1.06 2 0.96 1.70 0.21 146 Swf Ultisols Tuffac. sandstone

II8B 0.13 0.16 2 0.13 0.24 0.02 146 Swf Ultisols Tuffac. sandstone

Ash (%) II8A 22.66 4.67 2 22.66 25.96 19.35 146 Swf Ultisols Tuffac. sandstone

II8B 6.65 2.17 2 6.65 8.18 5.11 146 Swf Ultisols Tuffac. sandstone

C/N II8A 13 4 2 13 16 10 146 Swf Ultisols Tuffac. sandstone

II8B 34 4 2 34 37 31 146 Swf Ultisols Tuffac. sandstone

Calcium (mg/g) II2A 10.60 1.15 14 10.35 13.10 9.08 135 Lmrf Ultisols Tuffac. sandstone

II2B 6.81 0.52 4 7.06 7.09 6.04 135 Lmrf Ultisols Tuffac. sandstone

II2C 3.35 0.84 14 3.23 4.76 1.97 135 Lmrf Ultisols Tuffac. sandstone

II2E 2.30 0.42 9 2.37 2.95 1.75 135 Lmrf Ultisols Tuffac. sandstone

II2H 8.15 1.53 8 8.51 10.09 5.45 135 Lmrf Ultisols Tuffac. sandstone

II8A 18.37 2.16 2 18.37 19.90 16.85 146 Swf Ultisols Tuffac. sandstone

II8B 6.29 1.05 2 6.29 7.03 5.54 146 Swf Ultisols Tuffac. sandstone

Carbon (%) II8A 43 1 2 43 44 42 146 Swf Ultisols Tuffac. sandstone

II8B 50 2 2 50 51 48 146 Swf Ultisols Tuffac. sandstone

Iron (mg/g) II2A 0.18 0.09 14 0.16 0.43 0.07 135 Lmrf Ultisols Tuffac. sandstone

II2B 0.08 0.02 4 0.09 0.10 0.06 135 Lmrf Ultisols Tuffac. sandstone

II2C 0.09 0.06 14 0.07 0.25 0.04 135 Lmrf Ultisols Tuffac. sandstone

II2E 2.35 0.93 9 2.24 4.43 1.32 135 Lmrf Ultisols Tuffac. sandstone

II2H 0.21 0.11 8 0.21 0.36 0.07 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

209

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8A 1.66 2.13 2 1.66 3.17 0.16 146 Swf Ultisols Tuffac. sandstone

II8B 0.13 0.02 2 0.13 0.14 0.11 146 Swf Ultisols Tuffac. sandstone

Magnesium (mg/g) II2A 3.55 0.95 14 3.48 6.29 2.27 135 Lmrf Ultisols Tuffac. sandstone

II2B 5.53 1.46 4 5.64 7.13 3.72 135 Lmrf Ultisols Tuffac. sandstone

II2C 3.86 1.32 14 3.52 7.53 1.80 135 Lmrf Ultisols Tuffac. sandstone

II2E 2.13 0.38 9 2.21 2.77 1.49 135 Lmrf Ultisols Tuffac. sandstone

II2H 5.67 0.74 8 5.52 6.57 4.68 135 Lmrf Ultisols Tuffac. sandstone

II8A 2.89 1.04 2 2.89 3.63 2.16 146 Swf Ultisols Tuffac. sandstone

II8B 3.75 1.88 2 3.75 5.08 2.42 146 Swf Ultisols Tuffac. sandstone

Manganese (mg/g) II2A 0.29 0.05 14 0.30 0.37 0.16 135 Lmrf Ultisols Tuffac. sandstone

II2B 0.16 0.03 4 0.17 0.19 0.12 135 Lmrf Ultisols Tuffac. sandstone

II2C 0.08 0.03 14 0.08 0.14 0.04 135 Lmrf Ultisols Tuffac. sandstone

II2E 0.21 0.07 9 0.20 0.30 0.11 135 Lmrf Ultisols Tuffac. sandstone

II2H 0.37 0.06 8 0.39 0.43 0.29 135 Lmrf Ultisols Tuffac. sandstone

II8A 0.33 0.17 2 0.328 0.45 0.21 146 Swf Ultisols Tuffac. sandstone

II8B 0.06 0.04 2 0.06 0.09 0.04 146 Swf Ultisols Tuffac. sandstone

Nitrogen (%) II2A 2.13 0.25 14 2.15 2.62 1.75 135 Lmrf Ultisols Tuffac. sandstone

II2B 1.06 0.09 4 1.02 1.19 1.00 135 Lmrf Ultisols Tuffac. sandstone

II2C 0.93 0.18 14 0.92 1.14 0.66 135 Lmrf Ultisols Tuffac. sandstone

II2E 1.29 0.18 9 1.25 1.61 1.06 135 Lmrf Ultisols Tuffac. sandstone

II2H 1.77 0.25 8 1.71 2.18 1.44 135 Lmrf Ultisols Tuffac. sandstone

II8A 3.47 1.04 2 3.47 4.20 2.73 146 Swf Ultisols Tuffac. sandstone

II8B 1.47 0.25 2 1.47 1.64 1.29 146 Swf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II2A 1.04 0.16 14 0.99 1.34 0.80 135 Lmrf Ultisols Tuffac. sandstone

II2B 0.79 0.15 4 0.78 0.98 0.61 135 Lmrf Ultisols Tuffac. sandstone

II2C 0.64 0.23 14 0.57 1.23 0.31 135 Lmrf Ultisols Tuffac. sandstone

II2E 0.68 0.08 9 0.66 0.83 0.59 135 Lmrf Ultisols Tuffac. sandstone

II2H 1.13 0.26 8 1.19 1.48 0.70 135 Lmrf Ultisols Tuffac. sandstone

II8A 0.89 0.08 2 0.89 0.95 0.84 146 Swf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

210

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8B 0.45 0.02 2 0.45 0.47 0.41 146 Swf Ultisols Tuffac. sandstone

Potassium (mg/g) II2A 24.21 6.26 14 26.29 33.86 11.68 135 Lmrf Ultisols Tuffac. sandstone

II2B 13.47 4.96 4 13.25 19.58 7.78 135 Lmrf Ultisols Tuffac. sandstone

II2C 15.83 8.60 14 12.25 34.90 6.27 135 Lmrf Ultisols Tuffac. sandstone

II2E 11.90 1.77 9 12.11 13.94 8.06 135 Lmrf Ultisols Tuffac. sandstone

II2H 26.54 7.27 8 27.67 36.17 15.13 135 Lmrf Ultisols Tuffac. sandstone

II8A 12.84 15.11 2 12.84 23.52 2.16 146 Swf Ultisols Tuffac. sandstone

II8B 6.68 7.60 2 6.68 12.05 1.30 146 Swf Ultisols Tuffac. sandstone

Piper hispidium Aluminum (mg/kg) II2A 137 85 11 130 350 46 135 Lmrf Ultisols Tuffac. sandstone

II2C 232 335 13 71 915 2 135 Lmrf Ultisols Tuffac. sandstone

II2E 1607 762 14 1331 3641 682 135 Lmrf Ultisols Tuffac. sandstone

II2H 495 457 3 382 997 105 135 Lmrf Ultisols Tuffac. sandstone

II8A 859 806 2 859 1429 290 146 Swf Ultisols Tuffac. sandstone

II8B 196 43 3 219 222 146 146 Swf Ultisols Tuffac. sandstone

Ash (%) II8A 16.14 1.16 2 16.14 16.96 15.32 146 Swf Ultisols Tuffac. sandstone

II8B 2.54 0.92 3 2.05 3.60 1.96 146 Swf Ultisols Tuffac. sandstone

C/N II8A 10 3 2 10 12 8 146 Swf Ultisols Tuffac. sandstone

II8B 65 1 3 65 66 65 146 Swf Ultisols Tuffac. sandstone

Calcium (mg/g) II2A 12.09 1.21 11 11.98 14.06 10.43 135 Lmrf Ultisols Tuffac. sandstone

II2C 4.65 1.39 13 4.11 6.56 1.54 135 Lmrf Ultisols Tuffac. sandstone

II2E 2.34 0.53 14 2.28 3.33 1.54 135 Lmrf Ultisols Tuffac. sandstone

II2H 10.03 0.20 3 9.93 10.26 9.89 135 Lmrf Ultisols Tuffac. sandstone

II8A 11.57 6.24 2 11.57 15.98 7.16 146 Swf Ultisols Tuffac. sandstone

II8B 2.87 0.50 3 2.66 3.43 2.52 146 Swf Ultisols Tuffac. sandstone

Carbon (%) II8A 49 2 2 49 50 48 146 Swf Ultisols Tuffac. sandstone

II8B 53 0 3 53 53 52 146 Swf Ultisols Tuffac. sandstone

Iron (mg/g) II2A 0.17 0.07 11 0.16 0.31 0.10 135 Lmrf Ultisols Tuffac. sandstone

II2C 0.52 0.96 13 0.10 2.74 0.03 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

211

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II2E 1.60 1.01 14 1.31 4.48 0.63 135 Lmrf Ultisols Tuffac. sandstone

II2H 0.39 0.37 3 0.30 0.80 0.07 135 Lmrf Ultisols Tuffac. sandstone

II8A 0.60 0.56 2 0.60 0.99 0.20 146 Swf Ultisols Tuffac. sandstone

II8B 0.09 0.01 3 0.08 0.09 0.08 146 Swf Ultisols Tuffac. sandstone

Magnesium (mg/g) II2A 4.14 0.58 11 4.20 5.13 3.35 135 Lmrf Ultisols Tuffac. sandstone

II2C 1.40 0.48 13 1.22 2.30 0.84 135 Lmrf Ultisols Tuffac. sandstone

II2E 1.37 0.58 14 1.20 3.12 0.84 135 Lmrf Ultisols Tuffac. sandstone

II2H 3.53 3.00 3 3.21 4.21 3.17 135 Lmrf Ultisols Tuffac. sandstone

II8A 2.34 1.80 2 2.34 3.62 1.07 146 Swf Ultisols Tuffac. sandstone

II8B 0.59 0.19 3 0.53 0.80 0.44 146 Swf Ultisols Tuffac. sandstone

Manganese (mg/g) II2A 0.14 0.08 11 0.14 0.24 0.03 135 Lmrf Ultisols Tuffac. sandstone

II2C 0.04 0.03 13 0.03 0.11 0.01 135 Lmrf Ultisols Tuffac. sandstone

II2E 0.24 0.06 14 0.27 0.32 0.13 135 Lmrf Ultisols Tuffac. sandstone

II2H 0.16 0.02 3 0.15 0.18 0.14 135 Lmrf Ultisols Tuffac. sandstone

II8A 0.09 0.06 2 0.09 0.13 0.05 146 Swf Ultisols Tuffac. sandstone

II8B 0.01 0.00 3 0.01 0.01 0.01 146 Swf Ultisols Tuffac. sandstone

Nitrogen (%) II2A 2.31 0.56 11 2.13 3.38 1.70 135 Lmrf Ultisols Tuffac. sandstone

II2C 0.80 0.34 13 0.64 1.54 0.50 135 Lmrf Ultisols Tuffac. sandstone

II2E 0.85 0.20 14 0.83 1.44 0.64 135 Lmrf Ultisols Tuffac. sandstone

II2H 1.52 0.39 3 1.57 1.88 1.10 135 Lmrf Ultisols Tuffac. sandstone

II8A 5.13 1.68 2 5.13 6.31 3.94 146 Swf Ultisols Tuffac. sandstone

II8B 0.80 0.01 3 0.80 0.81 0.80 146 Swf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II2A 1.63 0.84 11 1.20 3.54 0.90 135 Lmrf Ultisols Tuffac. sandstone

II2C 1.01 0.66 13 0.71 2.40 0.44 135 Lmrf Ultisols Tuffac. sandstone

II2E 0.64 0.11 14 0.63 0.91 0.44 135 Lmrf Ultisols Tuffac. sandstone

II2H 1.16 0.38 3 0.95 1.61 0.93 135 Lmrf Ultisols Tuffac. sandstone

II8A 1.18 0.11 2 1.81 1.26 1.10 146 Swf Ultisols Tuffac. sandstone

II8B 0.39 0.03 3 0.40 0.42 0.36 146 Swf Ultisols Tuffac. sandstone

Potassium (mg/g) II2A 26.82 7.64 11 22.16 37.93 18.74 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

212

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II2C 30.99 12.67 13 26.42 50.63 8.33 135 Lmrf Ultisols Tuffac. sandstone

II2E 12.84 2.73 14 11.51 18.31 8.33 135 Lmrf Ultisols Tuffac. sandstone

II2H 33.56 12.23 3 31.31 46.76 22.60 135 Lmrf Ultisols Tuffac. sandstone

II8A 11.25 14.00 2 11.25 21.15 1.35 146 Swf Ultisols Tuffac. sandstone

II8B 5.01 7.01 3 1.09 13.11 0.85 146 Swf Ultisols Tuffac. sandstone

Pisonia albida Calcium (mg/g) II4A 12.01 1 104 Sdf Mollisols Alluvial deposits New

II4A 16.03 5.24 7 17.29 22.26 3.03 104 Sdf Mollisols Alluvial deposits Mature

II4A 15.49 5.05 15 1.87 22.26 3.03 149 Sdf Mollisols Alluvial deposits Guanica Forest

II4Ci 11.69 1 104 Sdf Mollisols Alluvial deposits

II4Ci 11.69 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Magnesium (mg/g) II4A 9.68 1 104 Sdf Mollisols Alluvial deposits New

II4A 7.61 2.13 7 7.93 10.36 3.18 104 Sdf Mollisols Alluvial deposits Mature

II4A 7.88 2.11 15 8.27 10.36 3.18 149 Sdf Mollisols Alluvial deposits Guanica Forest

II4Ci 4.17 1 104 Sdf Mollisols Alluvial deposits

II4Ci 4.17 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Nitrogen (%) II4A 2.51 1 104 Sdf Mollisols Alluvial deposits New

II4A 1.89 0.26 7 1.75 2.31 1.64 104 Sdf Mollisols Alluvial deposits Mature

II4A 1.98 0.30 16 1.87 2.51 1.64 149 Sdf Mollisols Alluvial deposits Guanica Forest

II4Ci 0.71 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Phosphorus (mg/g) II4A 1.42 1 104 Sdf Mollisols Alluvial deposits New

II4A 0.95 0.30 7 0.83 1.57 0.68 104 Sdf Mollisols Alluvial deposits Mature

II4A 1.01 0.32 15 0.83 1.57 0.68 149 Sdf Mollisols Alluvial deposits Guanica Forest

II4Ci 0.16 1 104 Sdf Mollisols Alluvial deposits

II4Ci 0.16 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Potassium (mg/g) II4A 16.85 1 104 Sdf Mollisols Alluvial deposits New

II4A 13.64 4.70 7 11.83 22.29 8.04 104 Sdf Mollisols Alluvial deposits Mature

II4A 14.04 4.51 16 13.55 22.29 8.04 149 Sdf Mollisols Alluvial deposits Guanica Forest

II4Ci 6.23 1 104 Sdf Mollisols Alluvial deposits
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

213

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II4Ci 6.23 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Pisonia subcordata Aluminum (mg/kg) II4A 31 1 102 Smf Limestone Tuffac. sandstone Adult 

Ash (%) II4A 4.66 1 102 Smf Limestone Tuffac. sandstone Adult 

Calcium (mg/g) II4A 2.86 1 102 Smf Limestone Tuffac. sandstone Adult 

Cobalt (μg/g) II4A 0.30 1 102 Smf Limestone Tuffac. sandstone Adult 

Iron (mg/kg) II4A 45 1 102 Smf Limestone Tuffac. sandstone Adult 

Magnesium (mg/g) II4A 6.10 1 102 Smf Limestone Tuffac. sandstone Adult 

Manganese (mg/kg) II4A 48 1 102 Smf Limestone Tuffac. sandstone Adult 

Nickel (μg/g) II4A 69.75 1 102 Smf Limestone Tuffac. sandstone Adult 

Nitrogen (%) II4A 1.50 1 102 Smf Limestone Tuffac. sandstone Adult 

Phosphorus (mg/g) II4A 0.78 1 102 Smf Limestone Tuffac. sandstone Adult 

Potassium (mg/g) II4A 12.91 1 102 Smf Limestone Tuffac. sandstone

Pithecellobium dulce Wood density (g/cc) II1B 0.60 1 97 Wfs Inceptisols Tuffac. sandstone

Plumeria alba Ash (%) II2A 7.87 0.28 3 8.00 8.07 7.55 91 Sdf Mollisols Alluvial deposits InMature

II2A 9.33 0.99 3 9.34 10.31 8.33 91 Sdf Mollisols Alluvial deposits

II2A 10.37 1.76 4 10.41 11.97 8.71 91 Sdf Mollisols Alluvial deposits lst defoliation

II2A 8.65 0.03 2 8.65 8.67 8.63 91 Sdf Mollisols Alluvial deposits 2nd defoliation

II2A 8.40 0.50 4 8.37 8.94 7.92 91 Sdf Mollisols Alluvial deposits Mature

II2B 6.53 1 91 Sdf Mollisols Alluvial deposits

Calcium (mg/g) II2A 1.60 0.20 3 1.50 1.8 1.50 91 Sdf Mollisols Alluvial deposits InMature

II2A 25.50 9.90 3 30.60 31.80 14.00 91 Sdf Mollisols Alluvial deposits

II2A 33.30 11.40 4 34.40 43.60 21.00 91 Sdf Mollisols Alluvial deposits lst defoliation

II2A 18.80 5.30 2 18.80 22.50 15.00 91 Sdf Mollisols Alluvial deposits 2nd defoliation

II2A 22.60 2.80 4 22.40 26.20 19.40 91 Sdf Mollisols Alluvial deposits Mature

II2B 23.40 1 91 Sdf Mollisols Alluvial deposits

Nitrogen (%) II2A 3.73 0.43 3 3.76 4.15 3.29 91 Sdf Mollisols Alluvial deposits InMature
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

214

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II2A 1.27 0.41 3 1.50 1.51 0.79 91 Sdf Mollisols Alluvial deposits

II2A 1.21 0.11 4 1.24 1.31 1.06 91 Sdf Mollisols Alluvial deposits lst defoliation

II2A 0.75 0.01 2 0.75 0.75 0.74 91 Sdf Mollisols Alluvial deposits 2nd defoliation

II2A 1.80 0.35 4 1.94 2.05 1.28 91 Sdf Mollisols Alluvial deposits Mature

II2B 0.43 1 91 Sdf Mollisols Alluvial deposits

Phosphorus (mg/g) II2A 3.44 0.67 3 0.35 0.41 0.28 91 Sdf Mollisols Alluvial deposits InMature

II2A 0.86 0.07 3 0.89 0.90 0.78 91 Sdf Mollisols Alluvial deposits

II2A 1.32 0.33 4 1.42 1.57 0.84 91 Sdf Mollisols Alluvial deposits lst defoliation

II2A 1.81 0.12 2 1.81 1.90 1.72 91 Sdf Mollisols Alluvial deposits 2nd defoliation

II2B 0.28 1 91 Sdf Mollisols Alluvial deposits

II2A 0.27 0.33 4 0.12 0.75 0.08 91 Sdf Mollisols Alluvial deposits Mature

Potassium (mg/g) II2A 28.00 3.50 3 3.00 3.00 2.39 91 Sdf Mollisols Alluvial deposits InMature

II2A 4.70 3.40 3 6.40 7.00 0.80 91 Sdf Mollisols Alluvial deposits

II2A 2.60 2.30 4 2.10 5.40 0.70 91 Sdf Mollisols Alluvial deposits lst defoliation

II2A 0.60 0.00 2 0.60 0.60 0.60 91 Sdf Mollisols Alluvial deposits 2nd defoliation

II2A 14.20 5.80 4 16.70 17.90 5.60 91 Sdf Mollisols Alluvial deposits Mature

II2B 5.30 1 91 Sdf Mollisols Alluvial deposits

Plumeria obtusa Ash (%) II2A 6.10 0.26 2 6.10 6.28 5.91 91 Sdf Mollisols Alluvial deposits

II2B 7.67 1 91 Sdf Mollisols Alluvial deposits

Calcium (mg/g) II2A 12.7 3.00 2 12.7 14.80 10.50 91 Sdf Mollisols Alluvial deposits

II2B 17.7 1 91 Sdf Mollisols Alluvial deposits

Nitrogen (%) II2A 1.34 0.28 2 1.34 1.53 1.14 91 Sdf Mollisols Alluvial deposits

II2B 0.67 1 91 Sdf Mollisols Alluvial deposits

Phosphorus (mg/g) II2A 0.32 0.41 2 0.32 0.61 0.03 91 Sdf Mollisols Alluvial deposits

II2B 0.59 1 91 Sdf Mollisols Alluvial deposits

Potassium (mg/g) II2A 8.10 0.10 2 8.10 8.20 8.00 91 Sdf Mollisols Alluvial deposits

II2B 16.20 1 91 Sdf Mollisols Alluvial deposits
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

215

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Plumeria rubra (red) Ash (%) II2A 10.00 0.13 2 10.00 10.09 9.90 91 Sdf Mollisols Alluvial deposits Dorado

Calcium (mg/g) II2A 21.30 2.50 2 21.30 23.00 19.50 91 Sdf Mollisols Alluvial deposits Dorado

Nitrogen (%) II2A 2.28 0.17 2 2.28 2.40 2.16 91 Sdf Mollisols Alluvial deposits Dorado

Phosphorus (mg/g) II2A 0.30 0.01 2 0.30 0.31 0.30 91 Sdf Mollisols Alluvial deposits Dorado

Potassium (mg/g) II2A 26.30 0.40 2 26.30 26.50 26.00 91 Sdf Mollisols Alluvial deposits Dorado

Plumeria rubra (white) Ash (%) II2A 9.88 0.81 2 9.88 10.45 9.31 91 Sdf Mollisols Alluvial deposits Dorado

II2A 10.62 0.50 3 10.09 10.69 10.09 91 Sdf Mollisols Alluvial deposits Guanica Forest

Calcium (mg/g) II2A 7.50 2.80 2 7.50 9.50 5.50 91 Sdf Mollisols Alluvial deposits Dorado

II2A 18.80 8.10 3 15.90 28.00 12.50 91 Sdf Mollisols Alluvial deposits GuanicaForest

Nitrogen (%) II2A 2.69 0.44 2 2.69 3.00 2.38 91 Sdf Mollisols Alluvial deposits Dorado

II2A 2.67 0.14 3 2.60 2.84 2.58 91 Sdf Mollisols Alluvial deposits Guanica Forest

Phosphorus (mg/g) II2A 3.31 0.51 2 3.31 3.67 2.95 91 Sdf Mollisols Alluvial deposits Dorado

II2A 2.16 0.29 3 2.20 2.42 1.85 91 Sdf Mollisols Alluvial deposits Guanica Forest

Potassium (mg/g) II2A 35.80 3.20 2 35.80 38.00 33.50 91 Sdf Mollisols Alluvial deposits Dorado

II2A 29.80 3.00 3 29.50 32.90 27.00 91 Sdf Mollisols Alluvial deposits Guanica Forest

Polipodium chnoodes Aluminum (mg/g) II6 0.52 1 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II6 4.61 1 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II6 0.62 1 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II6 4.39 1 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II6 0.66 1 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II6 1.75 1 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II6 1.04 1 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II6 22.65 1 135 Lmrf Ultisols Tuffac. sandstone

Polybotrya cervina Aluminum (mg/g) II6 0.23 0.01 2 0.23 0.23 0.22 135 Lmrf Ultisols Tuffac. sandstone

II6 0.88 0.63 8 0.90 1.73 0.13 135 Lmrf Ultisols Tuffac. sandstone

C/N II6 34 12 3 39 44 20 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

216

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Calcium (mg/g) II6 3.02 0.58 2 3.02 3.43 2.61 135 Lmrf Ultisols Tuffac. sandstone

II6 5.60 3.48 8 6.98 9.22 0.65 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II6 45 4 3 44 49 42 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II6 0.18 0.07 2 0.18 0.23 0.14 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II6 0.37 0.24 8 0.31 0.84 0.08 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II6 2.84 0.05 2 2.84 2.88 2.81 135 Lmrf Ultisols Tuffac. sandstone

II6 4.76 1.90 8 5.13 7.61 1.48 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II6 0.16 0.09 2 0.16 0.23 0.10 135 Lmrf Ultisols Tuffac. sandstone

II6 0.18 0.11 8 0.20 0.32 0.03 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II6 1.33 0.18 2 1.33 1.45 1.20 135 Lmrf Ultisols Tuffac. sandstone

II6 1.80 0.59 10 2.08 2.48 1.04 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II6 1.08 0.69 2 1.08 1.57 0.59 135 Lmrf Ultisols Tuffac. sandstone

II6 1.30 0.54 8 1.27 2.01 0.61 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II6 19.33 0.12 2 19.33 19.42 19.25 135 Lmrf Ultisols Tuffac. sandstone

II6 33.47 15.80 8 39.77 52.00 10.43 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II6 0.50 0.25 3 0.42 0.78 0.30 135 Lmrf Ultisols Tuffac. sandstone

Polypodium crassifolium Aluminum (mg/g) II6 0.10 1 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II6 5.25 1 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II6 0.10 1 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II6 6.51 1 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II6 0.43 1 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II6 0.73 1 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II6 0.39 1 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II6 23.97 1 135 Lmrf Ultisols Tuffac. sandstone

Polypodium spp. Aluminum (mg/g) II6 1.33 1.54 2 1.33 2.42 0.24 135 Lmrf Ultisols Tuffac. sandstone

C/N II6 32 8 2 32 37 26 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II6 8.68 5.57 2 8.68 12.62 4.74 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

217

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Carbon (%) II6 44 1 2 44 45 43 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II6 0.74 0.74 2 0.74 1.26 0.21 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II6 3.50 1.85 2 3.50 4.81 2.19 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II6 0.74 0.06 2 0.74 0.78 0.69 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II6 1.44 0.22 3 1.47 1.64 1.21 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II6 1.95 0.29 2 1.95 2.15 1.74 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II6 24.24 5.74 2 24.24 28.30 20.18 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II6 0.29 0.09 2 0.29 0.35 0.22 135 Lmrf Ultisols Tuffac. sandstone

Pouteria multiflora Wood density (g/cc) II1C 0.78 0.02 3 0.79 0.80 0.76 118

Prescottia oligatha Aluminum (μg/g) II6A 3.35 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6A 1.08 1 155 Lmrf Ultisols Serpentinite Maricao

II6E 6.67 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6E 7.21 1 155 Lmrf Ultisols Serpentinite Maricao

II6F 0.73 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6F 1.18 1 155 Lmrf Ultisols Serpentinite Maricao

Calcium (mg/g) II6A 10.41 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6A 11.68 1 155 Lmrf Ultisols Serpentinite Maricao

II6E 8.40 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6E 6.71 1 155 Lmrf Ultisols Serpentinite Maricao

II6F 6.24 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6F 6.13 1 155 Lmrf Ultisols Serpentinite Maricao

Cobalt (μg/g) II6A 2.80 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6A 3.50 1 155 Lmrf Ultisols Serpentinite Maricao

II6E 4.20 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6E 11.20 1 155 Lmrf Ultisols Serpentinite Maricao

II6F 0.99 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6F 2.50 1 155 Lmrf Ultisols Serpentinite Maricao
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

218

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Copper (μg/g) II6A 32.20 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6A 15.90 1 155 Lmrf Ultisols Serpentinite Maricao

II6E 32.30 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6E 34.40 1 155 Lmrf Ultisols Serpentinite Maricao

II6F 19.60 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6F 15.20 1 155 Lmrf Ultisols Serpentinite Maricao

Chromium (μg/g) II6A 5.10 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6A 14.02 1 155 Lmrf Ultisols Serpentinite Maricao

II6E 6.00 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6E 760 1 155 Lmrf Ultisols Serpentinite Maricao

II6F 0.24 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6F 116 1 155 Lmrf Ultisols Serpentinite Maricao

Iron (μg/g) II6A 3.08 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6A 3.95 1 155 Lmrf Ultisols Serpentinite Maricao

II6E 4.74 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6E 27.77 1 155 Lmrf Ultisols Serpentinite Maricao

II6F 0.63 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6F 3.54 1 155 Lmrf Ultisols Serpentinite Maricao

Magnesium (mg/g) II6A 2.87 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6A 1.94 1 155 Lmrf Ultisols Serpentinite Maricao

II6E 2.16 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6E 1.75 1 155 Lmrf Ultisols Serpentinite Maricao

II6F 1.75 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6F 1.65 1 155 Lmrf Ultisols Serpentinite Maricao

Manganese (μg/g) II6A 292 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6A 133 1 155 Lmrf Ultisols Serpentinite Maricao

II6E 567 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6E 332 1 155 Lmrf Ultisols Serpentinite Maricao

II6F 102 1 155 Smf Ultisols Tuffac. sandstone Luquillo
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

219

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II6F 119 1 155 Lmrf Ultisols Serpentinite Maricao

Nickel (μg/g) II6A 9 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6A 24 1 155 Lmrf Ultisols Serpentinite Maricao

II6E 8 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6E 139 1 155 Lmrf Ultisols Serpentinite Maricao

II6F 3 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6F 25 1 155 Lmrf Ultisols Serpentinite Maricao

Nitrogen (%) II6A 1.80 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6E 1.60 1 155 Lmrf Ultisols Serpentinite Luquillo

II6E 1.80 1 155 Smf Ultisols Tuffac. sandstone Maricao

II6F 1.60 1 155 Lmrf Ultisols Serpentinite Luquillo

Phosphorus (mg/g) II6A 1.16 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6A 1.88 1 155 Lmrf Ultisols Serpentinite Maricao

II6E 0.96 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6E 95 1 155 Lmrf Ultisols Serpentinite Maricao

II6F 1.09 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6F 2.18 1 155 Lmrf Ultisols Serpentinite Maricao

potassium (mg/g) II6A 10.00 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6A 26.64 1 155 Lmrf Ultisols Serpentinite Maricao

II6E 6.65 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6E 5.57 1 155 Lmrf Ultisols Serpentinite Maricao

II6F 18.88 1 155 Smf Ultisols Tuffac. sandstone Luquillo

II6F 32.62 1 155 Lmrf Ultisols Serpentinite Maricao

Prestoea montana Aluminum (mg/g) II1A 1.01 1.42 6 0.22 3.48 0.04 93 Wflm Inceptisols Tuffac. sandstone

II1A 0.32 0.30 22 0.20 1.32 0.10 107 Rfs Inceptisols Tuffac. sandstone

II1A 0.16 0.06 108 0.15 0.37 0.03 147 Lmrf Ultisols Tuffac. sandstone

II1E 0.04 1 93 Wflm Inceptisols Tuffac. sandstone

Ash (%) II1A 7.55 3.18 6 7.69 11.81 2.50 93 Wflm Inceptisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

220

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 8.97 3.24 7 9.31 13.56 4.24 92 Smf Ultisols Tuffac. sandstone

II1A 7.92 1.29 17 7.75 10.72 4.46 107 Rfs Inceptisols Tuffac. sandstone

II1A 7.20 0.78 108 7.08 9.24 5.82 147 Lmrf Ultisols Tuffac. sandstone

II1B 2.18 0.56 15 2.18 3.54 1.27 68 Swf Ultisols Tuffac. sandstone Class 2 SD

II1B 2.99 1 68 Swf Ultisols Tuffac. sandstone Class 2 LD

II1B 2.04 0.60 27 1.99 3.22 1.09 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 2.81 0.59 8 2.97 3.51 1.72 68 Swf Ultisols Tuffac. sandstone Class 3 SD

II1B 3.12 0.79 12 3.07 4.34 2.12 68 Swf Ultisols Tuffac. sandstone Class 3/4 LD

II1B 6.19 3.60 3 7.51 8.94 2.12 68 Swf Ultisols Tuffac. sandstone Class 3/4 LD

II1B 2.82 1.00 7 2.41 4.77 1.84 68 Swf Ultisols Tuffac. sandstone Class 4/5 LD

II1B 3.48 0.76 2 0.76 4.02 2.94 68 Swf Ultisols Tuffac. sandstone Class 4/5 SD

II1B 3.30 0.08 2 3.20 3.26 3.14 68 Swf Ultisols Tuffac. sandstone Class 4/5 SD

II1B 2.41 0.29 5 2.33 2.74 2.05 69 Swf Ultisols Tuffac. sandstone Class 1 LD

II1B 1.28 0.69 6 1.30 2.41 0.54 69 Swf Ultisols Tuffac. sandstone Class 2 LD

II1B 2.93 1.94 18 2.21 8.14 0.87 69 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 2.28 0.65 4 2.14 3.14 1.71 69 Swf Ultisols Tuffac. sandstone Class 3 DS

II1B 2.21 0.62 2 2.21 2.65 1.77 69 Swf Ultisols Tuffac. sandstone Class 2/3 SD

II1B 2.05 0.07 2 2.05 2.10 2.00 69 Swf Ultisols Tuffac. sandstone Class 3/4 ST

II1E 2.10 1 93 Wflm Inceptisols Tuffac. sandstone

II1E 7.50 4.26 6 5.53 15.57 4.73 92 Smf Ultisols Tuffac. sandstone

II1E 7.64 2.75 9 7.17 14.57 5.51 92 Smf Ultisols Tuffac. sandstone

C/N II1A 43 27 6 33 97 24 93 Wflm Inceptisols Tuffac. sandstone

II1A 23 4 22 22 36 18 107 Rfs Inceptisols Tuffac. sandstone

II1A 29 2 108 30 33 24 147 Lmrf Ultisols Tuffac. sandstone

II1C 44 1 3 45 46 43 93 Wflm Inceptisols Tuffac. sandstone

II1E 318 1 93 Wflm Inceptisols Tuffac. sandstone

Calcium (mg/g) II1A 3.21 1.71 6 3.40 5.26 0.36 93 Wflm Inceptisols Tuffac. sandstone

II1A 4.80 0.95 22 4.82 7.11 3.37 107 Rfs Inceptisols Tuffac. sandstone

II1A 3.05 1 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

221

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 4.02 1.02 108 3.88 6.53 0.83 147 Lmrf Ultisols Tuffac. sandstone

II1B 4.00 1.20 15 3.50 6.20 2.70 68 Swf Ultisols Tuffac. sandstone Class 2 LD

II1B 3.10 1 68 Swf Ultisols Tuffac. sandstone Class 2 LD

II1B 4.30 1.30 27 4.10 7.70 2.20 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 3.20 0.80 8 3.10 4.20 1.80 68 Swf Ultisols Tuffac. sandstone Class 3 SD

II1B 8.70 4.10 12 8.50 18.00 4.50 68 Swf Ultisols Tuffac. sandstone Class 3/4 LD

II1B 9.10 2.90 3 9.00 12.00 6.20 68 Swf Ultisols Tuffac. sandstone Class 3/4 LD

II1B 5.30 1.70 7 4.70 8.00 3.30 68 Swf Ultisols Tuffac. sandstone Class 4/5 LD

II1B 6.80 0.40 2 6.80 7.00 6.50 68 Swf Ultisols Tuffac. sandstone Class 4/5 SD

II1B 5.50 1.40 2 5.50 6.50 4.50 68 Swf Ultisols Tuffac. sandstone Class 4/5 SD

II1B 2.44 0.28 5 2.45 2.87 2.17 69 Swf Ultisols Tuffac. sandstone Class 1 LD

II1B 3.74 2.03 6 3.83 6.92 1.49 69 Swf Ultisols Tuffac. sandstone Class 2 LD

II1B 4.89 2.37 18 4.59 10.70 1.70 69 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 3.85 0.69 4 3.84 4.70 3.00 69 Swf Ultisols Tuffac. sandstone Class 3 SD

II1B 4.14 0.86 2 4.14 4.75 3.53 69 Swf Ultisols Tuffac. sandstone Class 2/3 SD

II1B 4.82 1.17 2 4.82 5.64 3.99 69 Swf Ultisols Tuffac. sandstone Class 3/4 ST

II1C 2.10 1.28 3 2.54 4.15 1.57 93 Wflm Inceptisols Tuffac. sandstone

II1C 4.12 1 135 Lmrf Ultisols Tuffac. sandstone

II1E 1.71 1 93 Wflm Inceptisols Tuffac. sandstone

Carbon (%) II1A 45 2 22 45 50 41 107 Rfs Inceptisols Tuffac. sandstone

II1A 46 1 108 46 47 45 147 Lmrf Ultisols Tuffac. sandstone

II1A 44 1 73 44 45 35 150 Swf Ultisols Tuffac. sandstone

Iron (mg/g) II1A 0.70 0.88 6 0.22 2.25 0.10 93 Wflm Inceptisols Tuffac. sandstone

II1A 0.21 0.11 22 0.17 0.50 0.09 107 Rfs Inceptisols Tuffac. sandstone

II1A 0.09 0.03 108 0.08 0.31 0.04 147 Lmrf Ultisols Tuffac. sandstone

II1E 0.15 1 93 Wflm Inceptisols Tuffac. sandstone

Magnesium (mg/g) II1A 2.06 1.39 6 1.89 3.68 0.64 93 Wflm Inceptisols Tuffac. sandstone

II1A 3.88 0.72 22 3.87 5.26 1.76 107 Rfs Inceptisols Tuffac. sandstone

II1A 3.17 1 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

222

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 2.48 0.51 108 2.47 3.48 0.23 147 Lmrf Ultisols Tuffac. sandstone

II1C 1.19 1.00 3 0.72 2.68 0.63 93 Wflm Inceptisols Tuffac. sandstone

II1C 2.48 1 135 Lmrf Ultisols Tuffac. sandstone

II1E 0.82 1 93 Wflm Inceptisols Tuffac. sandstone

Manganese (mg/g) II1A 0.34 0.24 6 0.32 0.68 0.02 93 Wflm Inceptisols Tuffac. sandstone

II1A 0.35 0.21 22 0.25 0.88 0.03 107 Rfs Inceptisols Tuffac. sandstone

II1A 0.40 0.11 108 0.40 0.64 0.14 147 Lmrf Ultisols Tuffac. sandstone

II1C 0.11 0.09 3 0.07 0.25 0.06 93 Wflm Inceptisols Tuffac. sandstone

II1E 0.09 1 93 Wflm Inceptisols Tuffac. sandstone

Nitrogen (%) II1A 1.20 0.45 6 1.13 1.76 0.46 93 Wflm Inceptisols Tuffac. sandstone

II1A 1.96 0.30 22 1.98 2.49 1.38 107 Rfs Inceptisols Tuffac. sandstone

II1A 1.25 1 135 Lmrf Ultisols Tuffac. sandstone

II1A 1.58 0.08 108 1.57 1.91 1.40 147 Lmrf Ultisols Tuffac. sandstone

II1A 2.04 0.30 73 2.00 2.84 1.53 150 Swf Ultisols Tuffac. sandstone

II1B 0.41 0.15 15 0.38 0.74 0.22 68 Swf Ultisols Tuffac. sandstone Class 2 LD

II1B 0.43 0.08 2 0.43 0.48 0.37 68 Swf Ultisols Tuffac. sandstone Class 2 SD

II1B 0.54 0.24 27 0.46 1.02 0.22 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 0.41 0.14 8 0.37 0.66 0.26 68 Swf Ultisols Tuffac. sandstone Class 3 SD

II1B 0.68 0.19 12 0.68 0.99 0.36 68 Swf Ultisols Tuffac. sandstone Class 3/4 LD

II1B 0.81 0.18 3 0.78 1.01 0.65 68 Swf Ultisols Tuffac. sandstone Class 3/4 LD

II1B 0.85 0.28 7 0.86 1.27 0.42 68 Swf Ultisols Tuffac. sandstone Class 4/5 LD

II1B 0.94 0.54 2 0.94 1.32 0.55 68 Swf Ultisols Tuffac. sandstone Class 4/5 SD

II1B 0.85 0.25 2 0.85 1.03 0.67 68 Swf Ultisols Tuffac. sandstone Class 4/5 SD

II1B 0.43 0.19 5 0.35 0.77 0.34 69 Swf Ultisols Tuffac. sandstone Class 1 LD

II1B 0.23 0.10 6 0.22 0.40 0.12 69 Swf Ultisols Tuffac. sandstone Class 2 LD

II1B 0.72 0.26 18 0.67 1.37 0.35 69 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 0.37 0.12 4 0.39 0.50 0.22 69 Swf Ultisols Tuffac. sandstone Class 3 SD

II1B 0.49 0.18 2 0.49 0.62 0.36 69 Swf Ultisols Tuffac. sandstone Class 2/3 SD

II1B 0.87 0.06 2 0.87 0.91 0.82 69 Swf Ultisols Tuffac. sandstone Class 3/4 ST
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

223

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1C 0.47 0.56 3 0.22 1.30 0.14 93 Wflm Inceptisols Tuffac. sandstone

II1C 1.10 1 135 Lmrf Ultisols Tuffac. sandstone

II1E 0.14 1 93 Wflm Inceptisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.63 0.34 6 0.61 1.03 0.26 93 Wflm Inceptisols Tuffac. sandstone

II1A 1.06 0.45 7 0.94 2.04 0.74 92 Smf Ultisols Tuffac. sandstone

II1A 1.03 0.28 22 1.05 1.59 0.47 107 Rfs Inceptisols Tuffac. sandstone

II1A 0.75 1 135 Lmrf Ultisols Tuffac. sandstone

II1A 0.74 0.12 108 0.74 1.06 0.24 147 Lmrf Ultisols Tuffac. sandstone

II1B 0.15 0.06 15 0.15 0.28 0.07 68 Swf Ultisols Tuffac. sandstone Class 2 LD

II1B 0.14 1 68 Swf Ultisols Tuffac. sandstone Class 2 SD

II1B 0.26 0.13 27 0.25 0.57 0.05 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 0.13 0.06 8 0.14 0.21 0.05 68 Swf Ultisols Tuffac. sandstone Class 3 SD

II1B 0.40 0.25 12 0.29 1.04 0.18 68 Swf Ultisols Tuffac. sandstone Class 3/4 LD

II1B 0.38 0.18 3 0.03 0.59 0.24 68 Swf Ultisols Tuffac. sandstone Class 3/4 LD

II1B 0.59 0.41 2 0.59 0.88 0.29 68 Swf Ultisols Tuffac. sandstone Class 4/5 SD

II1B 0.09 0.10 5 0.03 0.25 0.02 69 Swf Ultisols Tuffac. sandstone Class 2 LD

II1B 0.11 0.05 6 0.09 0.20 0.08 69 Swf Ultisols Tuffac. sandstone Class 2 LD

II1B 0.34 0.26 18 0.28 1.14 0.06 69 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 0.16 0.03 4 0.16 0.19 0.11 69 Swf Ultisols Tuffac. sandstone Class 3 SD

II1B 0.24 0.10 2 0.24 0.31 0.17 69 Swf Ultisols Tuffac. sandstone Class 2/3 SD

II1B 0.32 0.03 2 0.32 0.34 0.29 69 Swf Ultisols Tuffac. sandstone Class 3/4 ST

II1C 0.28 0.32 3 0.13 0.76 0.10 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.57 1 135 Lmrf Ultisols Tuffac. sandstone

II1E 0.12 1 93 Wflm Inceptisols Tuffac. sandstone

II1E 1.06 0.50 6 0.89 2.05 0.71 92 Smf Ultisols Tuffac. sandstone

II1E 1.11 0.27 9 1.02 1.79 0.88 92 Smf Ultisols Tuffac. sandstone

Il1B 0.52 0.19 7 0.58 0.80 0.26 68 Swf Ultisols Tuffac. sandstone Class 4/5 LD

Potassium (mg/g) II1A 7.71 5.30 6 10.29 12.05 0.90 93 Wflm Inceptisols Tuffac. sandstone

II1A 9.28 3.85 22 8.71 17.82 4.43 107 Rfs Inceptisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

224

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 1.47 1 135 Lmrf Ultisols Tuffac. sandstone

II1A 6.79 1.44 108 6.78 10.95 0.69 147 Lmrf Ultisols Tuffac. sandstone

II1B 2.85 0.43 5 2.76 3.46 2.37 68 Swf Ultisols Tuffac. sandstone Class 1 LD

II1B 0.52 1 68 Swf Ultisols Tuffac. sandstone Class 2 LD

II1B 3.10 4.90 18 1.52 18.60 0.22 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 0.27 1 68 Swf Ultisols Tuffac. sandstone Class 3 SD

II1B 0.35 1 68 Swf Ultisols Tuffac. sandstone Class 3/4 LD

II1B 1.41 0.38 2 1.41 1.68 1.14 68 Swf Ultisols Tuffac. sandstone Class 3/4 ST

II1B 2.85 0.43 5 2.76 3.46 2.37 69 Swf Ultisols Tuffac. sandstone Class 1 LD

II1B 0.52 0.25 6 0.53 0.96 0.25 69 Swf Ultisols Tuffac. sandstone Class 2 LD

II1B 3.10 4.90 18 1.52 18.6 0.22 69 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 2.73 1.74 4 3.21 4.24 0.27 69 Swf Ultisols Tuffac. sandstone Class 3 SD

II1B 4.35 1.18 2 4.35 5.18 3.51 69 Swf Ultisols Tuffac. sandstone Class 2/3 SD

II1B 1.41 0.38 2 1.41 1.68 1.14 69 Swf Ultisols Tuffac. sandstone Class 3/4 ST

II1C 5.58 3.70 3 4.77 10.76 2.03 93 Wflm Inceptisols Tuffac. sandstone

II1C 9.74 1 135 Lmrf Ultisols Tuffac. sandstone

II1E 4.60 1 93 Wflm Inceptisols Tuffac. sandstone

Sulfur (%) II1A 0.69 0.07 108 0.70 0.87 0.51 147 Lmrf Ultisols Tuffac. sandstone

II1A 0.69 0.16 54 0.67 1.19 0.45 150 Swf Ultisols Tuffac. sandstone

Prunus occidentalis Wood density (g/cc) II1C 0.78 0.04 5 0.80 0.82 0.72 117

Psychotria berteriana Aluminum (mg/g) II2A 0.89 0.19 25 0.85 1.39 0.64 135 Lmrf Ultisols Tuffac. sandstone

II2A 3.45 7.12 7 0.80 19.59 0.57 143 Wflm Inceptisols Tuffac. sandstone

II2A 1.03 1 93 Wflm Inceptisols Tuffac. sandstone

II2B 0.21 0.09 2 0.21 0.27 0.15 135 Lmrf Ultisols Tuffac. sandstone

II2B 0.82 0.34 2 0.82 1.07 0.58 93 Wflm Inceptisols Tuffac. sandstone

II2C 0.24 0.25 26 0.17 1.13 0.06 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

225

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II2C 0.92 1.59 6 0.28 4.16 0.15 143 Wflm Inceptisols Tuffac. sandstone <1cm

II2C 0.17 0.07 8 0.16 0.34 0.11 143 Wflm Inceptisols Tuffac. sandstone >1cm

II2E 1.24 0.70 13 1.07 2.73 0.27 135 Lmrf Ultisols Tuffac. sandstone

II2F 0.25 1 143 Wflm Inceptisols Tuffac. sandstone

II2H 0.64 0.19 6 0.68 0.87 0.36 135 Lmrf Ultisols Tuffac. sandstone

II8A 2.81 2.33 2 2.81 4.45 1.16 146 Swf Ultisols Tuffac. sandstone

II8B 0.28 0.12 2 0.28 0.36 0.19 146 Swf Ultisols Tuffac. sandstone

Ash (%) II2A 9.10 1.74 7 8.69 12.69 7.43 143 Wflm Inceptisols Tuffac. sandstone

II2A 10.40 1 93 Wflm Inceptisols Tuffac. sandstone

II2B 5.51 1 93 Wflm Inceptisols Tuffac. sandstone

II2C 4.14 0.63 6 4.07 5.26 3.37 143 Wflm Inceptisols Tuffac. sandstone <1cm

II2C 1.97 0.25 8 2.05 2.21 1.39 143 Wflm Inceptisols Tuffac. sandstone >1cm

II2F 5.06 1 143 Wflm Inceptisols Tuffac. sandstone

II8A 10.92 1.68 2 10.92 12.10 9.73 146 Swf Ultisols Tuffac. sandstone

II8B 2.49 0.62 2 2.49 2.92 2.05 146 Swf Ultisols Tuffac. sandstone

C/N II2A 21 5 7 19 30 16 143 Wflm Inceptisols Tuffac. sandstone

II2A 13 1 93 Wflm Inceptisols Tuffac. sandstone

II2B 51 1 93 Wflm Inceptisols Tuffac. sandstone

II2C 79 19 6 77 107 58 143 Wflm Inceptisols Tuffac. sandstone <1cm

II2C 133 26 8 140 157 91 143 Wflm Inceptisols Tuffac. sandstone >1cm

II2F 32 1 143 Wflm Inceptisols Tuffac. sandstone

II8A 14 4 2 14 17 11 146 Swf Ultisols Tuffac. sandstone

II8B 72 33 2 72 95 48 146 Swf Ultisols Tuffac. sandstone

Calcium (mg/g) II2A 8.64 1.17 25 8.68 10.70 6.49 135 Lmrf Ultisols Tuffac. sandstone

II2A 13.74 3.00 7 14.13 17.35 7.97 143 Wflm Inceptisols Tuffac. sandstone

II2A 12.23 1 93 Wflm Inceptisols Tuffac. sandstone Leaves

II2B 4.66 0.44 2 4.66 4.97 4.35 135 Lmrf Ultisols Tuffac. sandstone

II2B 9.93 2.33 2 9.93 11.57 8.28 93 Wflm Inceptisols Tuffac. sandstone

II2C 4.06 1.13 26 4.24 5.86 1.95 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

226

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II2C 5.89 1.31 5 5.82 7.52 3.98 143 Wflm Inceptisols Tuffac. sandstone <1cm

II2C 2.00 0.31 8 2.01 2.47 1.58 143 Wflm Inceptisols Tuffac. sandstone >1cm

II2E 3.17 0.76 13 2.89 4.75 2.29 135 Lmrf Ultisols Tuffac. sandstone

II2F 5.76 1 143 Wflm Inceptisols Tuffac. sandstone

II2H 8.17 1.40 6 7.89 10.83 7.03 135 Lmrf Ultisols Tuffac. sandstone

II8A 21.35 8.49 2 21.35 27.35 15.35 146 Swf Ultisols Tuffac. sandstone

II8B 4.78 2.35 2 4.78 6.45 3.12 146 Swf Ultisols Tuffac. sandstone

Carbon (%) II2A 46 2 7 47 47 42 143 Wflm Inceptisols Tuffac. sandstone

II2C 47 1 6 47 48 46 143 Wflm Inceptisols Tuffac. sandstone <1cm

II2C 48 0 8 48 49 48 143 Wflm Inceptisols Tuffac. sandstone >1cm

II2F 51 1 143 Wflm Inceptisols Tuffac. sandstone

II8A 50 5 2 50 54 47 146 Swf Ultisols Tuffac. sandstone

II8B 53 0 2 53 53 53 146 Swf Ultisols Tuffac. sandstone

Iron (mg/g) II2A 0.27 0.42 25 0.17 2.22 0.04 135 Lmrf Ultisols Tuffac. sandstone

II2A 0.39 0.66 7 0.16 1.89 0.11 143 Wflm Inceptisols Tuffac. sandstone

II2A 0.24 1 93 Wflm Inceptisols Tuffac. sandstone

II2B 0.07 0.02 2 0.07 0.08 0.05 135 Lmrf Ultisols Tuffac. sandstone

II2B 0.28 0.22 2 0.28 0.43 0.12 93 Wflm Inceptisols Tuffac. sandstone

II2C 0.15 0.29 26 0.07 1.29 0.01 135 Lmrf Ultisols Tuffac. sandstone

II2C 0.06 0.02 6 0.06 0.07 0.04 143 Wflm Inceptisols Tuffac. sandstone <1cm

II2C 0.09 0.10 8 0.04 0.29 0.02 143 Wflm Inceptisols Tuffac. sandstone >1cm

II2E 1.54 1.36 13 1.28 4.54 0.05 135 Lmrf Ultisols Tuffac. sandstone

II2F 0.03 1 143 Wflm Inceptisols Tuffac. sandstone

II2H 0.17 0.05 6 0.18 0.23 0.10 135 Lmrf Ultisols Tuffac. sandstone

II8A 1.01 0.52 2 1.01 1.37 0.64 146 Swf Ultisols Tuffac. sandstone

II8B 0.10 0.09 2 0.10 0.17 0.04 146 Swf Ultisols Tuffac. sandstone

Magnesium (mg/g) II2A 4.42 0.89 25 4.19 6.85 3.07 135 Lmrf Ultisols Tuffac. sandstone

II2A 5.58 1.16 7 5.36 7.57 3.81 143 Wflm Inceptisols Tuffac. sandstone

II2A 6.02 1 93 Wflm Inceptisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

227

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II2B 1.64 0.76 2 1.64 2.18 1.10 135 Lmrf Ultisols Tuffac. sandstone

II2B 4.30 1.38 2 4.30 5.28 3.33 93 Wflm Inceptisols Tuffac. sandstone

II2C 1.42 0.51 26 1.37 2.56 0.62 135 Lmrf Ultisols Tuffac. sandstone

II2C 2.22 0.82 6 2.31 3.14 0.94 143 Wflm Inceptisols Tuffac. sandstone <1cm

II2C 0.54 0.24 8 0.48 0.87 0.29 143 Wflm Inceptisols Tuffac. sandstone >1cm

II2E 0.92 0.65 13 0.80 3.03 0.51 135 Lmrf Ultisols Tuffac. sandstone

II2F 2.87 1 143 Wflm Inceptisols Tuffac. sandstone

II2H 3.97 1.28 6 3.72 2.72 0.43 135 Lmrf Ultisols Tuffac. sandstone

II8A 4.39 4.69 2 4.39 7.71 1.07 146 Swf Ultisols Tuffac. sandstone

II8B 0.70 0.08 2 0.70 0.76 0.64 146 Swf Ultisols Tuffac. sandstone

Manganese (mg/g) II2A 0.17 0.05 25 0.17 0.28 0.09 135 Lmrf Ultisols Tuffac. sandstone

II2A 0.50 0.22 7 0.47 0.79 0.23 143 Wflm Inceptisols Tuffac. sandstone

II2A 0.19 1 93 Wflm Inceptisols Tuffac. sandstone

II2B 0.23 0.13 2 0.23 0.33 0.14 135 Lmrf Ultisols Tuffac. sandstone

II2B 0.54 0.66 2 0.54 1.01 0.08 93 Wflm Inceptisols Tuffac. sandstone

II2C 0.13 0.06 26 0.13 0.27 0.04 135 Lmrf Ultisols Tuffac. sandstone

II2C 0.28 0.14 6 0.27 0.44 0.08 143 Wflm Inceptisols Tuffac. sandstone <1cm

II2C 0.05 0.03 8 0.04 0.09 0.02 143 Wflm Inceptisols Tuffac. sandstone >1cm

II2E 0.08 0.06 13 0.07 0.27 0.04 135 Lmrf Ultisols Tuffac. sandstone

II2F 0.17 1 143 Wflm Inceptisols Tuffac. sandstone

II2H 0.32 0.11 6 0.31 0.47 0.18 135 Lmrf Ultisols Tuffac. sandstone

II8A 0.16 0.09 2 0.16 0.23 0.10 146 Swf Ultisols Tuffac. sandstone

II8B 0.03 0.02 2 0.03 0.04 0.01 146 Swf Ultisols Tuffac. sandstone

Nitrogen (%) II2A 2.61 0.32 25 2.57 3.62 2.21 135 Lmrf Ultisols Tuffac. sandstone

II2A 2.35 0.51 7 2.46 3.05 1.42 143 Wflm Inceptisols Tuffac. sandstone

II2A 3.17 1 93 Wflm Inceptisols Tuffac. sandstone

II2B 0.65 0.07 2 0.65 0.70 0.60 135 Lmrf Ultisols Tuffac. sandstone

II2B 0.86 1 93 Wflm Inceptisols Tuffac. sandstone

II2C 0.57 0.14 26 0.56 0.93 0.35 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

228

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II2C 0.63 0.15 6 0.62 0.81 0.44 143 Wflm Inceptisols Tuffac. sandstone <1cm

II2C 0.38 0.09 8 0.35 0.54 0.30 143 Wflm Inceptisols Tuffac. sandstone >1cm

II2E 0.72 0.15 13 0.71 0.99 0.50 135 Lmrf Ultisols Tuffac. sandstone

II2F 1.61 1 143 Wflm Inceptisols Tuffac. sandstone

II2H 1.79 0.25 6 1.83 2.05 1.48 135 Lmrf Ultisols Tuffac. sandstone

II8A 3.96 1.60 2 3.96 5.09 2.83 146 Swf Ultisols Tuffac. sandstone

II8B 0.80 0.38 2 0.83 1.10 0.56 146 Swf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II2A 0.92 0.16 25 0.91 1.27 0.68 135 Lmrf Ultisols Tuffac. sandstone

II2A 0.82 0.20 7 0.81 1.19 0.57 143 Wflm Inceptisols Tuffac. sandstone

II2A 0.99 1 93 Wflm Inceptisols Tuffac. sandstone

II2B 0.37 0.01 2 0.37 0.38 0.36 135 Lmrf Ultisols Tuffac. sandstone

II2B 0.69 0.28 2 0.69 0.89 0.49 93 Wflm Inceptisols Tuffac. sandstone

II2C 0.32 0.06 26 0.32 0.40 0.21 135 Lmrf Ultisols Tuffac. sandstone

II2C 0.24 0.06 6 0.26 0.30 0.16 143 Wflm Inceptisols Tuffac. sandstone <1cm

II2C 0.13 0.02 8 0.13 0.17 0.10 143 Wflm Inceptisols Tuffac. sandstone >1cm

II2E 0.34 0.12 13 0.33 0.67 0.23 135 Lmrf Ultisols Tuffac. sandstone

II2F 0.57 1 143 Wflm Inceptisols Tuffac. sandstone

II2H 0.72 0.21 6 0.68 1.06 0.45 135 Lmrf Ultisols Tuffac. sandstone

II8A 0.98 0.10 2 0.94 1.06 0.91 146 Swf Ultisols Tuffac. sandstone

II8B 0.53 0.26 2 0.53 0.72 0.35 146 Swf Ultisols Tuffac. sandstone

Potassium (mg/g) II2A 28.63 9.19 25 27.70 47.97 17.89 135 Lmrf Ultisols Tuffac. sandstone

II2A 11.64 2.55 7 10.86 15.79 8.33 143 Wflm Inceptisols Tuffac. sandstone

II2A 17.31 1 93 Wflm Inceptisols Tuffac. sandstone

II2B 10.79 0.03 2 1.79 10.81 10.77 135 Lmrf Ultisols Tuffac. sandstone

II2B 10.40 4.61 2 10.40 13.66 7.14 93 Wflm Inceptisols Tuffac. sandstone

II2C 18.57 7.97 26 17.45 34.83 7.18 135 Lmrf Ultisols Tuffac. sandstone

II2C 6.44 1.69 6 5.72 8.64 4.96 143 Wflm Inceptisols Tuffac. sandstone <1cm

II2C 4.92 1.34 8 5.38 6.41 2.83 143 Wflm Inceptisols Tuffac. sandstone >1cm

II2E 12.23 4.61 13 11.40 23.94 7.93 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

229

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II2F 9.69 1 143 Wflm Inceptisols Tuffac. sandstone

II2H 23.95 5.19 6 23.83 31.53 17.53 135 Lmrf Ultisols Tuffac. sandstone

II8A 12.03 13.83 2 12.03 21.81 2.25 146 Swf Ultisols Tuffac. sandstone

II8B 3.67 1.91 2 3.67 5.02 2.32 146 Swf Ultisols Tuffac. sandstone

Psychotria maleolens Aluminum (mg/g) II2A 14.16 2.13 5 12.77 17.16 12.56 143 Wflm Inceptisols Tuffac. sandstone

II2C 3.52 1.18 6 3.40 5.34 2.05 143 Wflm Inceptisols Tuffac. sandstone <1cm

Ash (%) II2A 12.10 1.50 5 11.91 13.91 9.88 143 Wflm Inceptisols Tuffac. sandstone

II2C 3.36 0.79 6 3.14 4.53 2.51 143 Wflm Inceptisols Tuffac. sandstone <1cm

C/N II2A 34 4 5 35 40 30 143 Wflm Inceptisols Tuffac. sandstone

II2C 129 33 6 126 174 77 143 Wflm Inceptisols Tuffac. sandstone <1cm

Calcium (mg/g) II2A 11.78 0.55 5 11.63 12.58 11.13 143 Wflm Inceptisols Tuffac. sandstone

II2C 3.99 1.12 6 3.97 5.65 2.54 143 Wflm Inceptisols Tuffac. sandstone <1cm

Carbon (%) II2A 42 2 5 43 43 40 143 Wflm Inceptisols Tuffac. sandstone

II2C 47 0 6 47 47 46 143 Wflm Inceptisols Tuffac. sandstone <1cm

Iron (mg/g) II2A 0.30 0.34 5 0.15 0.91 0.14 143 Wflm Inceptisols Tuffac. sandstone

II2C 0.06 0.03 6 0.05 0.12 0.03 143 Wflm Inceptisols Tuffac. sandstone <1cm

Magnesium (mg/g) II2A 5.17 0.77 5 4.85 6.49 4.53 143 Wflm Inceptisols Tuffac. sandstone

II2C 1.49 0.54 6 1.47 2.12 0.86 143 Wflm Inceptisols Tuffac. sandstone <1cm

Manganese (mg/g) II2A 0.15 0.09 5 0.11 0.28 0.08 143 Wflm Inceptisols Tuffac. sandstone

II2C 0.05 0.40 6 0.03 0.11 0.02 143 Wflm Inceptisols Tuffac. sandstone <1cm

Nitrogen (%) II2A 1.24 0.11 5 1.24 1.37 1.08 143 Wflm Inceptisols Tuffac. sandstone

II2C 0.39 0.12 6 0.37 0.61 0.27 143 Wflm Inceptisols Tuffac. sandstone <1cm

Phosphorus (mg/g) II2A 0.44 0.05 5 0.47 0.48 0.35 143 Wflm Inceptisols Tuffac. sandstone

II2C 0.20 0.09 6 0.17 0.31 0.12 143 Wflm Inceptisols Tuffac. sandstone <1cm

Potassium (mg/g) II2A 4.80 0.66 5 4.54 5.95 4.33 143 Wflm Inceptisols Tuffac. sandstone

II2C 4.97 0.81 6 4.73 6.08 4.10 143 Wflm Inceptisols Tuffac. sandstone <1cm

Psychotria sp Ash (%) II2A 12.67 1.00 2 12.67 13.37 11.96 94 Swf Ultisols Tuffac. sandstone Tree diam. <3cm
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

230

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II2B 6.36 0.68 2 6.36 6.84 5.88 94 Swf Ultisols Tuffac. sandstone Tree diam. <3cm

II2C 4.63 1.94 2 4.63 6.00 3.25 94 Swf Ultisols Tuffac. sandstone Tree diam. <3cm

Nitrogen (%) II2A 2.36 0.04 2 2.36 2.39 2.33 94 Swf Ultisols Tuffac. sandstone Tree diam. <3cm

II2B 0.94 0.09 2 0.94 1.00 0.87 94 Swf Ultisols Tuffac. sandstone Tree diam. <3cm

II2C 0.83 0.30 2 0.83 1.04 0.62 94 Swf Ultisols Tuffac. sandstone Tree diam. <3cm

Phosphorus (mg/g) II2A 1.21 0.04 2 1.21 1.24 1.18 94 Swf Ultisols Tuffac. sandstone Tree diam. <3cm

II2B 0.72 0.06 2 0.72 0.76 0.67 94 Swf Ultisols Tuffac. sandstone Tree diam. <3cm

II2C 0.54 0.13 2 0.54 0.63 0.44 94 Swf Ultisols Tuffac. sandstone Tree diam. <3cm

Potassium (mg/g) II2A 20.79 3.49 2 20.79 23.25 18.32 94 Swf Ultisols Tuffac. sandstone Tree diam. <3cm

II2B 7.83 0.44 2 7.83 8.14 7.52 94 Swf Ultisols Tuffac. sandstone Tree diam. <3cm

II2C 6.88 0.67 2 6.88 7.35 6.40 94 Swf Ultisols Tuffac. sandstone Tree diam. <3cm

Pteris altissima C/N II6 15 1 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II6 40 1 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II6 2.63 1 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II6 0.51 1 135 Lmrf Ultisols Tuffac. sandstone

Pterocarpus macrocarpus Wood density (g/cc) II1B 0.52 0.04 6 0.52 0.45 0.53 97 Wfs Inceptisols Tuffac. sandstone

Pterocarpus officinalis Aluminum (mg/g) II1A 0.06 0.02 14 0.07 0.10 0.03 78 Mfs Mangle Swamp

II1A 0.05 0.03 12 0.05 0.11 0.03 83 Mfs Histosol Alluvial deposits

II1Ai 0.03 0.00 2 0.03 0.03 0.03 83 Mfs Histosol Alluvial deposits Rachis

II1Aiii 0.05 0.01 28 0.05 0.08 0.03 83 Mfs Histosol Alluvial deposits

II9A 0.55 2.18 49 0.09 11.61 0.03 84 Mfs Histosol Alluvial deposits Laguncularia 

II9A 0.35 1.05 87 0.10 6.03 0.02 84 Mfs Histosol Alluvial deposits Other

II9A 0.45 1.78 228 0.11 18.58 0.02 84 Mfs Histosol Alluvial deposits

II9B 0.19 0.59 74 0.06 3.42 0.01 84 Mfs Histosol Alluvial deposits

II9C 0.10 0.05 2 0.10 0.13 0.07 84 Mfs Histosol Alluvial deposits

II9D 1.89 6.51 69 0.24 40.26 0.05 84 Mfs Histosol Alluvial deposits
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

231

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Ash (%) II1A 7.67 0.86 38 7.68 9.87 6.23 76 Mfs Histosol Alluvial deposits

II1A 6.91 0.98 14 7.12 7.80 4.73 78 Mfs Histosol Alluvial deposits

II1B 6.44 1.03 30 6.58 8.25 4.62 76 Mfs Histosol Alluvial deposits

II1F 4.02 0.30 5 4.07 4.33 3.57 76 Mfs Histosol Alluvial deposits Seeds

II9A 13.30 3.36 49 12.84 25.56 8.89 84 Mfs Histosol Alluvial deposits

II9A 9.00 3.91 84 8.64 16.87 3.59 84 Mfs Histosol Alluvial deposits Other

II9A 6.04 3.04 211 5.43 34.79 2.80 84 Mfs Histosol Alluvial deposits

II9B 6.09 1.40 72 6.08 9.95 3.01 84 Mfs Histosol Alluvial deposits

II9C 4.18 0.94 2 4.18 4.84 3.51 84 Mfs Histosol Alluvial deposits

II9D 8.76 10.12 69 6.13 63.56 4.53 84 Mfs Histosol Alluvial deposits

II1A 0.48 0.06 9 0.47 0.56 0.41 83 Mfs Histosol Alluvial deposits

II1Ai 0.54 0.01 2 0.54 0.54 0.53 83 Mfs Histosol Alluvial deposits Rachis

II1Aiii 0.49 0.03 28 0.44 0.57 0.42 83 Mfs Histosol Alluvial deposits

C/N II1A 16 2 14 17 18 13 78 Mfs Entisols Swamps

II1A 35 3 9 35 40 31 83 Mfs Histosol Alluvial deposits

II1Ai 41 1 2 41 42 40 83 Mfs Histosol Alluvial deposits Rachis

II1Aiii 16 2 28 16 19 14 83 Mfs Histosol Alluvial deposits

Calcium (mg/g) II1A 21.63 3.01 38 21.29 28.97 15.44 76 Smf Inceptisols Swamps

II1A 20.13 5.10 14 22.43 25.47 9.52 78 Mfs Entisols Swamps

II1A 11.83 1.49 12 11.90 14.17 9.00 83 Mfs Histosol Alluvial deposits

II1Ai 15.66 0.28 2 15.66 15.86 15.46 83 Mfs Histosol Alluvial deposits Rachis

II1Aiii 9.149 1.74 28 9.099 12.13 5.65 83 Mfs Histosol Alluvial deposits

II1B 21.59 3.52 30 21.38 28.57 15.48 76 Smf Inceptisols Swamps

II1F 2.72 0.76 5 2.59 3.90 1.84 76 Smf Inceptisols Swamps Seeds

II9A 38.71 9.91 49 35.99 58.93 19.24 84 Mfs Histosol Alluvial deposits Laguncularia 

II9A 26.95 15.32 87 18.08 60.89 9.36 84 Mfs Histosol Alluvial deposits Other

II9A 15.23 3.54 228 14.62 33.64 8.71 84 Mfs Histosol Alluvial deposits

II9B 18.59 5.70 74 18.80 29.57 9.55 84 Mfs Histosol Alluvial deposits
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

232

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II9C 4.17 1.75 2 4.17 5.41 2.93 84 Mfs Histosol Alluvial deposits

II9D 15.40 5.38 69 14.36 32.74 5.92 84 Mfs Histosol Alluvial deposits

Carbon (%) II1A 43 0 14 43 44 43 78 Mfs Mangle Swamp

II1A 51 1 9 51 52 50 83 Mfs Histosol Alluvial deposits

II1Ai 52 0 2 52 52 52 83 Mfs Histosol Alluvial deposits Rachis

II1Aiii 54 1 28 54 54 53 83 Mfs Histosol Alluvial deposits

Iron (mg/g) II1A 0.05 0.02 14 0.05 0.09 0.02 78 Mfs Mangle Swamp

II1A 0.07 0.02 12 0.07 0.10 0.04 83 Mfs Histosol Alluvial deposits

II1Ai 0.04 0.01 2 0.04 0.04 0.04 83 Mfs Histosol Alluvial deposits Rachis

II1Aiii 0.09 0.04 28 0.07 0.21 0.05 83 Mfs Histosol Alluvial deposits

II9A 1.34 2.53 49 0.68 14.60 0.22 84 Mfs Histosol Alluvial deposits Laguncularia 

II9A 0.65 1.25 87 0.27 6.88 0.02 84 Mfs Histosol Alluvial deposits Other

II9A 0.55 1.91 228 0.15 20.45 0.04 84 Mfs Histosol Alluvial deposits

II9B 0.35 0.94 74 0.15 6.03 0.03 84 Mfs Histosol Alluvial deposits

II9C 0.25 0.34 2 0.25 0.49 0.01 84 Mfs Histosol Alluvial deposits

II9D 2.37 7.21 69 0.40 40.26 0.12 84 Mfs Histosol Alluvial deposits

Magnesium (mg/g) II1A 4.43 0.65 38 4.53 5.47 2.90 76 Mfs Histosol Alluvial deposits

II1A 5.85 2.08 14 6.13 8.02 2.53 78 Mfs Histosol Alluvial deposits

II1B 2.82 0.40 30 2.84 3.76 1.88 76 Mfs Histosol Alluvial deposits

II1G 2.18 0.23 5 2.20 2.49 1.94 76 Mfs Histosol Alluvial deposits Seeds

II9A 2.90 0.58 49 2.87 4.37 1.99 84 Mfs Histosol Alluvial deposits Laguncularia 

II9A 2.77 0.41 87 2.76 4.07 1.94 84 Mfs Histosol Alluvial deposits Other

II9A 3.59 0.56 228 3.53 6.12 2.14 84 Mfs Histosol Alluvial deposits

II9B 2.13 0.45 74 2.19 2.92 0.91 84 Mfs Histosol Alluvial deposits

II9C 1.94 0.09 2 1.94 2.01 1.88 84 Mfs Histosol Alluvial deposits

II9D 3.16 1.17 69 2.89 8.88 2.06 84 Mfs Histosol Alluvial deposits

II1A 1.95 0.47 12 2.00 2.71 1.31 83 Mfs Histosol Alluvial deposits

II1Ai 1.42 0.00 2 1.42 1.42 1.41 83 Mfs Histosol Alluvial deposits Rachis

II1Aiii 3.47 0.60 2 3.52 4.45 2.04 83 Mfs Histosol Alluvial deposits
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

233

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Manganese (mg/g) II1A 0.02 0.01 14 0.02 0.05 0.01 78 Mfs Mangle Swamp

II1A 0.06 0.02 12 0.06 0.09 0.03 83 Mfs Histosol Alluvial deposits

II1Ai 0.04 0.00 2 0.04 0.04 0.04 83 Mfs Histosol Alluvial deposits Rachis

II1Aiii 0.07 0.04 28 0.06 0.15 0.04 83 Mfs Histosol Alluvial deposits

II9A 0.13 0.06 49 0.11 0.35 0.07 84 Mfs Histosol Alluvial deposits Laguncularia 

II9A 0.24 0.16 87 0.22 0.78 0.04 84 Mfs Histosol Alluvial deposits Other

II9A 0.11 0.05 228 0.10 0.46 0.04 84 Mfs Histosol Alluvial deposits

II9B 0.09 0.05 74 0.07 0.25 0.02 84 Mfs Histosol Alluvial deposits

II9C 0.02 0.01 2 0.02 0.03 0.02 84 Mfs Histosol Alluvial deposits

II9D 0.17 0.14 69 0.13 0.85 0.06 84 Mfs Histosol Alluvial deposits

Nitrogen (%) II1A 1.77 0.16 38 1.75 2.21 1.40 76 Mfs Histosol Alluvial deposits

II1A 2.75 0.31 14 2.58 3.30 2.41 78 Mfs Mangle Swamp

II1A 1.45 0.14 9 1.46 1.65 1.25 83 Mfs Histosol Alluvial deposits

II1Ai 1.27 0.04 2 1.27 1.30 1.24 83 Mfs Histosol Alluvial deposits Rachis

II1Aiii 3.42 0.33 28 3.41 3.87 2.81 83 Mfs Histosol Alluvial deposits

II1B 1.08 0.13 30 1.08 1.33 0.79 76 Smf Inceptisols Swamps

II1G 2.80 0.08 5 2.82 2.88 2.67 76 Smf Inceptisols Swamps

Phosphorus (mg/g) II1A 0.55 0.13 38 0.54 0.80 0.31 76 Smf Inceptisols Swamps

II1A 0.91 0.15 14 0.86 1.23 0.71 78 Mfs Mangle Swamp

II1A 0.73 0.10 12 0.71 0.87 0.62 83 Mfs Histosol Alluvial deposits

II1Ai 0.48 0.02 2 0.48 0.49 0.46 83 Mfs Histosol Alluvial deposits Rachis

II1Aiii 1.11 0.16 28 1.07 1.50 0.86 83 Mfs Histosol Alluvial deposits

II1B 0.40 0.15 30 0.37 0.86 0.21 76 Smf Inceptisols Swamps

II1F 1.51 0.17 5 1.54 1.66 1.24 76 Smf Inceptisols Swamps Seeds

II9A 0.59 0.17 49 0.55 0.95 0.32 84 Mfs Histosol Alluvial deposits Laguncularia 

II9A 0.46 0.15 87 0.46 0.76 0.19 84 Mfs Histosol Alluvial deposits Other

II9A 0.39 0.14 228 0.37 0.82 0.19 84 Mfs Histosol Alluvial deposits

II9B 0.29 0.11 74 0.27 0.56 0.09 84 Mfs Histosol Alluvial deposits

II9C 1.15 0.00 2 1.15 1.15 1.14 84 Mfs Histosol Alluvial deposits
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

234

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II9D 0.94 0.21 69 0.94 1.47 0.54 84 Mfs Histosol Alluvial deposits

Potassium (mg/g) II1A 5.72 0.76 38 5.75 7.15 4.08 76 Smf Inceptisols Swamps

II1A 12.69 3.94 14 12.22 20.93 8.21 78 Mfs Mangle Swamp

II1A 10.66 0.91 12 10.66 12.29 8.86 83 Mfs Histosol Alluvial deposits

II1Ai 10.33 0.22 2 10.33 10.49 10.17 83 Mfs Histosol Alluvial deposits Rachis

II1Aiii 9.20 1.65 28 8.94 12.79 6.95 83 Mfs Histosol Alluvial deposits

II1B 5.11 1.08 30 5.09 6.66 3.16 76 Smf Inceptisols Swamps

II1G 12.42 1.15 5 12.33 13.89 11.10 76 Smf Inceptisols Swamps Seeds

II9A 5.32 1.38 49 5.25 9.41 3.63 84 Mfs Histosol Alluvial deposits Laguncularia 

II9A 4.09 1.53 87 3.81 8.07 1.63 84 Mfs Histosol Alluvial deposits Other

II9A 4.05 1.01 228 3.91 7.94 2.00 84 Mfs Histosol Alluvial deposits

II9B 3.33 1.33 74 3.08 7.70 1.01 84 Mfs Histosol Alluvial deposits

II9C 9.34 0.35 2 9.34 9.59 9.10 84 Mfs Histosol Alluvial deposits

II9D 4.35 1.63 69 3.88 9.17 2.21 84 Mfs Histosol Alluvial deposits

Sodium (mg/g) II1A 0.61 0.23 14 0.55 1.00 0.28 78 Mfs Mangle Swamp

II1A 1.51 0.65 12 1.36 2.79 0.92 83 Mfs Histosol Alluvial deposits

II1Ai 0.72 0.00 2 0.72 0.72 0.72 83 Mfs Histosol Alluvial deposits Rachis

II1Aiii 0.43 0.14 28 0.42 0.66 0.21 83 Mfs Histosol Alluvial deposits

Sulfur (mg/g) II1A 0.15 0.02 9 0.14 0.17 0.12 83 Mfs Histosol Alluvial deposits

II1Ai 0.13 0.00 2 0.13 0.13 0.13 83 Mfs Histosol Alluvial deposits Rachis

II1Aiii 0.41 0.05 28 0.41 0.52 0.31 83 Mfs Histosol Alluvial deposits

Wood density (g/cc) II1B 0.53 1 97 Wfs Inceptisols Tuffac. sandstone

Rhizophora mangle Aluminum (mg/kg) II1A 30 10 2 30 40 30 78 Mfs Mangle Swamp

II1A 10 10 12 10 30 10 81 Mfs Entisols Swamps Adult 

II1A 30 30 12 20 110 10 81 Mfs Entisols Swamps Dead 

II1A 13 4 12 12 23 8 81 Mfs Entisols Swamps Old

II1A 60 60 12 20 220 10 81 Mfs Entisols Swamps Yellow 

II1A 20 20 12 10 50 10 81 Mfs Entisols Swamps Young 
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

235

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 66 3 4 65 70 64 82 Mfs Entisols Swamps

II1B 146 3 2 146 148 144 82 Mfs Entisols Swamps Dead wood

II1B 31 3 3 32 34 28 87 Mfs Entisols Compound dunes

II1B 17 1 2 17 16 18 157 Mfs Entisols Compound dunes

II1C 50 10 4 50 70 40 82 Mfs Entisols Swamps Woody roots

II1E 1002 306 3 849 1354 802 80 Mfs Entisols Swamps

II1A 25 33 36 15 166 7 80 Mfs Entisols Swamps

II1C 48 10 4 48 57 38 80 Mfs Entisols Swamps

II1C 39 42 18 32 207 15 80 Mfs Entisols Swamps Young stems

II1E 108 112 4 91 241 11 80 Mfs Entisols Swamps

II1E 13 3 2 13 15 11 80 Mfs Entisols Swamps Roots & lenticels

Ash (%) II1A 14.67 2.99 9 14.22 19.64 11.44 72 Dfs Mangle Mangle Border

II1A 15.33 2.02 7 14.40 18.86 13.10 72 Dfs Mangle Mangle Canal

II1A 18.18 2.47 8 18.51 20.50 12.67 72 Dfs Mangle Mangle Hipersaline lagoon

II1A 15.05 0.64 5 15.18 15.91 14.28 72 Dfs Mangle Mangle Seedlings

II1A 1.59 0.31 9 1.49 2.24 1.25 72 Dfs Mangle Mangle Border

II1A 1.15 0.42 7 1.14 1.75 0.50 72 Dfs Mangle Mangle Canal

II1A 10.31 0.05 2 10.31 10.35 10.28 78 Mfs Mangle Swamp

II1A 8.55 0.58 36 8.47 9.63 7.44 80 Mfs Entisols Swamps

II1A 10.82 0.09 4 10.79 10.94 10.75 82 Mfs Entisols Swamps

II1A 17.20 0.80 3 85 Dfs Mangle Dune deposits

II1A 8.74 1 86 Mfs Entisols Swamps

II1A 13.14 1 86 Mfs Entisols Swamps Yellow

II1A 10.18 1 86 Mfs Entisols Swamps

II1Ai 13.18 2.12 5 13.77 16.68 11.67 75 Dfs Mangle Dune deposits

II1Aii 14.28 2.33 5 13.58 17.55 11.83 75 Dfs Mangle Dune deposits

II1B 10.44 0.02 2 10.44 10.45 10.42 82 Mfs Entisols Swamps Dead wood

II1B 3.81 1.65 3 3.99 5.36 2.07 87 Mfs Entisols Compound dunes

II1C 11.39 0.11 4 11.41 11.49 11.26 80 Mfs Entisols Swamps
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

236

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1C 11.17 2.22 18 10.82 14.72 6.83 80 Mfs Entisols Swamps Young stems

II1C 11.48 0.26 4 11.48 11.72 11.25 82 Mfs Entisols Swamps Woody roots

II1C 12.30 0.05 3 85 Dfs Mangle Dune deposits

II1C 10.60 1 86 Mfs Entisols Swamps

II1E 16.46 1.38 4 16.47 17.68 15.21 80 Mfs Entisols Swamps

II1E 6.07 0.12 2 6.07 6.15 5.98 80 Mfs Entisols Swamps Roots & lenticels

II1F 8.50 1 86 Mfs Entisols Swamps Mature

II1F 7.20 1 86 Mfs Entisols Swamps Inmature

II1G 16.50 1.10 3 85 Dfs Mangle Dune deposits

II6 9.30 0.70 3 85 Dfs Mangle Dune deposits Seedlings

II6 2.98 1 86 Mfs Entisols Swamps Inmature

II6 2.24 1 86 Mfs Entisols Swamps Mature

C/N II1A 26 0 2 26 26 25 78 Mfs Mangle Swamp

II1A 39 5 36 40 46 29 80 Mfs Entisols Swamps

II1A 40 5 12 41 46 32 81 Mfs Entisols Swamps Adult 

II1A 80 29 12 1 143 57 81 Mfs Entisols Swamps Dead 

II1A 39 4 12 40 43 32 81 Mfs Entisols Swamps Old

II1A 80 26 12 72 133 58 81 Mfs Entisols Swamps Yellow 

II1A 44 4 12 44 48 36 81 Mfs Entisols Swamps Young 

II1A 43 2 4 43 45 41 82 Mfs Entisols Swamps

II1B 229 0 2 229 229 229 82 Mfs Entisols Swamps Dead wood

II1B 208 38 2 208 235 181 87 Mfs Entisols Compound dunes

II1C 76 2 4 76 78 73 80 Mfs Entisols Swamps

II1C 79 19 18 88 104 47 80 Mfs Entisols Swamps Young stems

II1C 112 9 4 112 122 104 82 Mfs Entisols Swamps Woody roots

II1E 47 3 3 47 49 44 80 Mfs Entisols Swamps

II1E 76 5 4 75 83 73 80 Mfs Entisols Swamps

Calcium (mg/g) II1A 12.60 2.80 8 12.40 17.50 7.50 72 Dfs Mangle Mangle Hipersaline lagoon

II1A 11.50 4.20 7 11.40 17.50 5.00 72 Dfs Mangle Mangle Canal
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

237

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 15.90 3.10 9 14.90 22.40 12.50 72 Dfs Mangle Mangle Border

II1A 21.55 0.32 2 21.55 21.77 21.33 72 Dfs Mangle Mangle Yellow-fringe

II1A 17.81 1.67 2 17.81 18.99 16.63 72 Dfs Mangle Mangle Old-fringe

II1A 22.47 0.11 2 22.47 22.55 22.39 72 Dfs Mangle Mangle Senescent-fringe

II1A 16.12 0.23 2 16.12 16.29 15.96 72 Dfs Mangle Mangle Young-fringe

II1A 8.15 1.21 2 8.15 9.00 7.30 72 Dfs Mangle Mangle Old-salina

II1A 15.16 0.28 2 15.16 15.36 14.96 72 Dfs Mangle Mangle Senescent-salina

II1A 12.04 0.55 2 12.04 12.43 11.65 72 Dfs Mangle Mangle Yellow-salina

II1A 7.89 0.51 2 7.89 8.26 7.53 72 Dfs Mangle Mangle Young-salina

II1A 12.02 4.23 5 13.14 16.28 4.38 77 Mfs Swamp Swamp

II1A 18.33 0.20 2 18.33 18.47 18.19 78 Mfs Mangle Swamp

II1A 8.20 1.00 36 8.24 9.94 6.19 80 Mfs Entisols Swamps

II1A 8.66 1.54 12 8.95 10.87 5.90 81 Mfs Entisols Swamps Adult 

II1A 12.37 1.83 12 12.16 17.17 10.22 81 Mfs Entisols Swamps Dead

II1A 9.03 0.95 12 9.19 10.30 6.72 81 Mfs Entisols Swamps Old

II1A 12.35 1.42 12 12.60 14.67 9.81 81 Mfs Entisols Swamps Yellow

II1A 7.81 1.04 12 8.10 9.05 6.07 81 Mfs Entisols Swamps Young 

II1A 8.66 1.04 4 8.73 9.70 7.48 82 Mfs Entisols Swamps

II1B 29.31 0.62 2 29.31 29.75 28.87 82 Mfs Entisols Swamps Dead wood

II1B 7.14 7.15 3 3.39 15.39 2.64 87 Mfs Entisols Compound dunes

II1B 3.77 0.34 2 3.77 3.53 4.01 157 Mfs Entisols Compound dunes

II1C 24.82 0.62 4 24.94 25.37 24.03 80 Mfs Entisols Swamps

II1C 24.82 8.03 18 23.23 42.00 17.01 80 Mfs Entisols Swamps Young stems

II1C 27.07 2.38 4 26.04 30.61 25.51 82 Mfs Entisols Swamps Woody roots

II1E 2.32 0.08 3 2.28 2.40 2.27 80 Mfs Entisols Swamps

II1E 1.89 0.15 4 1.86 2.11 1.74 80 Mfs Entisols Swamps

II1E 3.14 0.04 2 3.14 3.17 3.12 80 Mfs Entisols Swamps Roots & lenticels

II9A 16.61 2.79 11 17.22 19.84 9.86 73 Dfs Mangle Dune deposits

II9B 25.47 3.54 11 24.42 32.37 20.70 73 Dfs Mangle Dune deposits
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

238

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II9C 3.78 1.33 11 3.55 5.89 1.70 73 Dfs Mangle Dune deposits

II9D 9.94 1.05 11 9.93 11.67 7.71 73 Dfs Mangle Dune deposits

Carbon (%) II1A 41 0 2 41 42 41 78 Mfs Mangle Swamp

II1A 47 3 36 48 50 38 80 Mfs Entisols Swamps

II1A 44 1 12 44 46 43 81 Mfs Entisols Swamps Adult 

II1A 44 1 12 44 46 42 81 Mfs Entisols Swamps Dead 

II1A 44 2 12 4 46 41 81 Mfs Entisols Swamps Old

II1A 44 2 12 44 46 40 81 Mfs Entisols Swamps Yellow 

II1A 44 1 12 44 45 43 81 Mfs Entisols Swamps Young 

II1A 50 0 4 50 51 50 82 Mfs Entisols Swamps

II1A 38 1 3 85 Dfs Mangle Dune deposits

II1B 48 0 2 48 48 48 82 Mfs Entisols Swamps Dead wood

II1C 45 0 4 45 45 45 80 Mfs Entisols Swamps

II1C 45 7 18 46 66 35 80 Mfs Entisols Swamps Young stems

II1C 49 0 4 49 49 49 82 Mfs Entisols Swamps Woody roots

II1C 40 1 3 85 Dfs Mangle Dune deposits

II1E 40 0 3 40 40 40 80 Mfs Entisols Swamps

II1E 40 1 4 40 41 39 80 Mfs Entisols Swamps

II1E 47 0 2 47 47 47 80 Mfs Entisols Swamps Roots & lenticels

II1G 38 0 3 85 Dfs Mangle Dune deposits

II6 43 1 3 85 Dfs Mangle Dune deposits Seedlings

Iron (mg/kg) II1A 50 0 2 50 50 50 78 Mfs Mangle Swamp

II1A 30 20 12 30 80 10 81 Mfs Entisols Swamps Adult 

II1A 40 10 12 40 50 20 81 Mfs Entisols Swamps Dead 

II1A 20 10 12 30 40 10 81 Mfs Entisols Swamps Old

II1A 30 10 12 30 50 10 81 Mfs Entisols Swamps Yellow 

II1A 20 10 11 20 30 10 81 Mfs Entisols Swamps Young 

II1A 40 0 4 40 40 30 82 Mfs Entisols Swamps

II1B 150 0 2 150 150 150 82 Mfs Entisols Swamps Dead wood
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

239

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1B 36 8 3 36 44 28 87 Mfs Entisols Compound dunes

II1B 15 1 2 15 14 16 157 Mfs Entisols Compound dunes

II1C 60 10 4 60 60 50 82 Mfs Entisols Swamps Woody roots

II1A 39 19 36 38 110 6 80 Mfs Entisols Swamps

II1C 62 20 4 63 83 40 80 Mfs Entisols Swamps

II1C 33 16 18 34 71 2 80 Mfs Entisols Swamps Young stems

II1E 3729 1627 3 3695 5372 2119 80 Mfs Entisols Swamps

II1E 155 32 4 151 192 127 80 Mfs Entisols Swamps

II1E 18 5 2 18 21 14 80 Mfs Entisols Swamps Roots & lenticels

Magnesium (mg/g) II1A 7.61 0.04 2 7.61 7.64 7.58 74 Dfs Mangle Dune deposits Senescent-fringe

II1A 6.70 0.15 2 6.70 6.81 6.59 74 Dfs Mangle Dune deposits Young-fringe

II1A 5.71 0.37 2 5.71 5.97 5.45 74 Dfs Mangle Dune deposits Old-salina

II1A 7.28 0.36 2 7.28 7.53 7.02 74 Dfs Mangle Dune deposits Senescent-salina

II1A 6.91 0.13 2 6.91 7.00 6.91 74 Dfs Mangle Dune deposits Yellow-salina

II1A 4.97 0.04 2 4.97 5.00 4.94 74 Dfs Mangle Dune deposits Mature-salina

II1A 1.02 0.21 4 0.94 1.38 0.78 72 Dfs Mangle Mangle Border

II1A 1.27 0.31 4 1.26 1.73 0.94 72 Dfs Mangle Mangle Canal

II1A 3.59 0.71 5 3.59 5.03 2.65 77 Mfs Swamp Swamp

II1A 4.54 0.08 2 4.54 4.60 4.48 78 Mfs Mangle Swamp

II1A 4.69 0.46 36 4.56 5.84 3.94 80 Mfs Entisols Swamps

II1A 4.48 0.84 12 4.21 6.41 3.61 81 Mfs Entisols Swamps Adult 

II1A 6.27 0.87 12 6.06 7.70 5.04 81 Mfs Entisols Swamps Dead 

II1A 4.51 0.78 12 4.48 5.66 3.35 81 Mfs Entisols Swamps Old

II1A 6.22 1.02 12 5.81 7.88 5.12 81 Mfs Entisols Swamps Yellow 

II1A 3.96 0.62 12 3.77 5.37 3.35 81 Mfs Entisols Swamps Young 

II1B 0.89 0.28 3 0.90 1.16 0.60 87 Mfs Entisols Compound dunes

II1B 0.69 0.00 2 0.69 0.69 0.69 157 Mfs Entisols Compound dunes

II1C 2.71 0.18 4 2.72 2.88 2.50 80 Mfs Entisols Swamps

II1C 2.56 0.53 18 2.63 3.97 1.49 80 Mfs Entisols Swamps Young stems
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

240

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1E 4.69 0.46 3 4.44 5.22 4.41 80 Mfs Entisols Swamps

II1E 3.76 0.52 4 3.82 4.31 3.10 80 Mfs Entisols Swamps

II1E 1.51 0.02 2 1.51 1.53 1.49 80 Mfs Entisols Swamps Roots & lenticels

II9A 6.13 1.32 11 6.53 7.40 2.83 73 Dfs Mangle Dune deposits

II9B 3.09 0.35 11 3.23 3.47 2.52 73 Dfs Mangle Dune deposits

II9C 1.78 0.46 11 1.73 2.60 1.10 73 Dfs Mangle Dune deposits

II9D 3.48 0.33 11 3.38 4.04 2.88 73 Dfs Mangle Dune deposits

II1A 4.83 1.04 4 4.83 5.85 3.80 82 Mfs Entisols Swamps

II1B 3.07 0.13 2 3.07 3.16 2.98 82 Mfs Entisols Swamps Dead wood

II1C 1.72 0.35 4 1.76 2.04 1.33 82 Mfs Entisols Swamps Woody roots

Manganese (mg/g) II1A 0.59 0.00 2 0.59 0.60 0.59 78 Mfs Mangle Swamp

II1A 0.06 0.03 12 0.06 0.11 0.02 81 Mfs Entisols Swamps Adult 

II1A 0.09 0.03 12 0.08 0.13 0.03 81 Mfs Entisols Swamps Dead 

II1A 0.06 0.03 12 0.06 0.10 0.03 81 Mfs Entisols Swamps Old

II1A 0.09 0.04 12 0.09 0.15 0.13 81 Mfs Entisols Swamps Yellow 

II1A 0.05 0.02 12 0.04 0.08 0.02 81 Mfs Entisols Swamps Young 

II1A 0.05 0.01 4 0.05 0.07 0.04 82 Mfs Entisols Swamps

II1B 0.02 0.00 2 0.02 0.02 0.02 82 Mfs Entisols Swamps Dead wood

II1B 0.05 0.04 3 0.03 0.10 0.02 87 Mfs Entisols Compound dunes

II1B 0.05 0.01 2 0.05 0.04 0.05 157 Mfs Entisols Compound dunes

II1C 0.04 0.00 4 0.05 0.05 0.04 82 Mfs Entisols Swamps Woody roots

II1A 0.06 0.03 36 0.06 0.12 0.02 80 Mfs Entisols Swamps

II1C 0.10 0.00 4 0.10 0.10 0.10 80 Mfs Entisols Swamps

II1C 0.06 0.03 18 0.05 0.14 0.02 80 Mfs Entisols Swamps Young stems

II1E 0.04 0.00 3 0.03 0.05 0.02 80 Mfs Entisols Swamps

II1E 0.01 0.02 4 0.01 0.01 0.01 80 Mfs Entisols Swamps

II1E 0.01 0.00 2 0.01 0.01 0.01 80 Mfs Entisols Swamps Roots & lenticels

Nitrogen (%) II1C 0.40 1 86 Mfs Entisols Swamps

II1A 1.07 0.31 3 0.96 1.55 0.73 72 Dfs Mangle Mangle Hipersaline lagoon
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

241

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 1.27 0.31 4 1.26 1.73 0.94 72 Dfs Mangle Mangle Canal

II1A 1.02 0.21 4 0.94 1.38 0.78 72 Dfs Mangle Mangle Border-control

II1A 1.12 0.23 5 1.08 1.51 0.89 75 Dfs Mangle Dune deposits Seedlings

II1A 1.21 0.16 5 1.23 1.41 0.97 77 Mfs Swamp Swamp

II1A 1.62 0.03 2 1.62 1.64 1.60 78 Mfs Mangle Swamp

II1A 1.23 0.22 36 1.21 1.65 0.85 80 Mfs Entisols Swamps

II1A 1.12 0.13 12 1.07 1.38 1.00 81 Mfs Entisols Swamps Adult 

II1A 0.60 0.15 12 0.64 0.74 0.32 81 Mfs Entisols Swamps Dead 

II1A 1.13 0.13 12 1.07 1.38 1.02 81 Mfs Entisols Swamps Old

II1A 0.59 0.13 12 0.61 0.73 0.34 81 Mfs Entisols Swamps Yellow 

II1A 1.02 0.11 12 0.99 1.21 0.91 81 Mfs Entisols Swamps Young 

II1A 1.17 0.06 4 1.17 1.23 1.12 82 Mfs Entisols Swamps

II1A 1.17 0.11 3 85 Dfs Mangle Dune deposits

II1A 0.97 1 86 Mfs Entisols Swamps

II1A 0.62 1 86 Mfs Entisols Swamps Yellow

II1A 1.07 1 86 Mfs Entisols Swamps

II1A 0.31 1 74 Dfs Mangle Dune deposits Yellow-fringe

II1A 0.55 1 74 Dfs Mangle Dune deposits Old-fringe

II1A 0.28 1 74 Dfs Mangle Dune deposits Senescent-fringe

II1A 0.84 1 74 Dfs Mangle Dune deposits Young-fringe

II1A 1.38 1 74 Dfs Mangle Dune deposits Old-salina

II1A 0.37 1 74 Dfs Mangle Dune deposits Senescent-salina

II1A 0.39 1 74 Dfs Mangle Dune deposits Yellow-salina

II1A 0.83 1 74 Dfs Mangle Dune deposits Mature-salina

II1Ai 1.42 0.07 5 1.42 1.52 1.36 75 Dfs Mangle Dune deposits

II1Aii 1.51 0.07 5 1.51 1.59 1.41 75 Dfs Mangle Dune deposits

II1B 0.21 0.00 2 0.21 0.21 0.21 82 Mfs Entisols Swamps Dead wood

II1B 0.22 0.03 2 0.22 0.24 0.19 87 Mfs Entisols Compound dunes

II1C 0.60 0.02 4 0.60 0.62 0.58 80 Mfs Entisols Swamps
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

242

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1C 0.62 0.27 18 0.52 1.40 0.38 80 Mfs Entisols Swamps Young stems

II1C 0.44 0.04 4 0.44 0.47 0.40 82 Mfs Entisols Swamps Woody roots

II1C 0.83 0.08 3 85 Dfs Mangle Dune deposits

II1E 0.86 0.05 3 0.86 0.90 0.81 80 Mfs Entisols Swamps

II1E 0.52 0.03 4 0.53 0.55 0.49 80 Mfs Entisols Swamps

II1E 0.46 0.00 2 0.46 0.46 0.46 80 Mfs Entisols Swamps Roots & lenticels

II1F 0.47 1 86 Mfs Entisols Swamps Mature

II1F 0.63 1 86 Mfs Entisols Swamps Inmature

II1G 1.27 0.09 3 85 Dfs Mangle Dune deposits

II6 0.86 0.07 3 85 Dfs Mangle Dune deposits Seedlings

II6 0.47 1 86 Mfs Entisols Swamps Inmature

II6 0.24 1 86 Mfs Entisols Swamps Mature

II9A 0.70 0.27 11 0.60 1.48 0.54 73 Dfs Mangle Dune deposits

II9B 0.72 0.08 11 0.68 0.88 0.65 73 Dfs Mangle Dune deposits

II9C 0.68 0.11 11 0.66 0.86 0.54 73 Dfs Mangle Dune deposits

II9D 0.58 0.12 11 0.57 0.78 0.41 73 Dfs Mangle Dune deposits

Phosphorus (mg/g) II1A 0.40 0.03 3 85 Dfs Mangle Dune deposits

II1C 0.20 0.04 3 85 Dfs Mangle Dune deposits

II1G 0.40 0.03 3 85 Dfs Mangle Dune deposits

II6 0.50 0.04 3 85 Dfs Mangle Dune deposits Seedlings

II1A 0.46 0.11 9 0.41 0.70 0.36 72 Dfs Mangle Mangle Border

II1A 0.65 0.35 7 0.48 1.42 0.36 72 Dfs Mangle Mangle Canal

II1A 0.41 0.11 5 0.36 0.60 0.33 72 Dfs Mangle Mangle Hipersaline lagoon

II1A 0.85 0.04 5 0.85 0.89 0.79 75 Dfs Mangle Dune deposits Seedlings

II1A 0.11 0.02 2 0.11 0.12 0.10 74 Dfs Mangle Dune deposits Yellow-fringe

II1A 0.38 0.00 2 0.38 0.38 0.38 74 Dfs Mangle Dune deposits Old-fringe

II1A 0.21 0.01 2 0.21 0.22 0.21 74 Dfs Mangle Dune deposits Senescen-fringe

II1A 0.41 0.02 2 0.41 0.42 0.40 74 Dfs Mangle Dune deposits Young m.-fringe

II1A 0.46 0.04 2 0.46 0.49 0.43 74 Dfs Mangle Dune deposits Old mature-salina
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

243

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 0.10 0.02 2 0.10 0.12 0.09 74 Dfs Mangle Dune deposits Senescent-salina

II1A 0.22 0.01 2 0.22 0.22 0.21 74 Dfs Mangle Dune deposits Yellow-salina

II1A 0.62 0.01 2 0.62 0.63 0.61 74 Dfs Mangle Dune deposits Young-salina

II1A 0.94 0.09 5 0.95 1.12 0.79 77 Mfs Swamp Swamp

II1A 0.77 0.00 2 0.77 0.77 0.77 78 Mfs Mangle Swamp

II1A 0.43 0.10 36 0.40 0.62 0.31 80 Mfs Entisols Swamps

II1A 0.46 0.08 12 0.44 0.60 0.37 81 Mfs Entisols Swamps Adult 

II1A 0.09 0.02 12 0.08 0.14 0.07 81 Mfs Entisols Swamps Dead 

II1A 0.40 0.08 12 0.38 0.56 0.32 81 Mfs Entisols Swamps Old

II1A 0.11 0.05 12 0.09 0.21 0.08 81 Mfs Entisols Swamps Yellow 

II1A 0.56 0.12 12 0.50 0.74 0.41 81 Mfs Entisols Swamps Young 

II1A 0.40 0.09 4 0.40 0.49 0.31 82 Mfs Entisols Swamps

II1A 0.68 1 86 Mfs Entisols Swamps

II1A 0.05 1 86 Mfs Entisols Swamps Yellow

II1A 0.37 1 86 Mfs Entisols Swamps

II1Ai 0.80 0.04 5 0.80 0.84 0.75 75 Dfs Mangle Dune deposits

II1Aii 0.83 0.05 5 0.85 0.88 0.77 75 Dfs Mangle Dune deposits

II1B 0.06 0.00 2 0.06 0.06 0.06 82 Mfs Entisols Swamps Dead wood

II1B 0.28 0.07 3 0.29 0.34 0.21 87 Mfs Entisols Compound dunes

II1B 0.19 0.03 2 0.19 0.17 0.21 157 Mfs Entisols Compound dunes

II1C 0.19 0.01 4 0.19 0.20 0.18 80 Mfs Entisols Swamps

II1C 0.31 0.15 18 0.20 0.62 0.17 80 Mfs Entisols Swamps Young stems

II1C 0.17 0.02 4 0.18 0.19 0.15 82 Mfs Entisols Swamps Woody roots

II1C 0.08 1 86 Mfs Entisols Swamps

II1E 0.30 0.03 3 0.29 0.33 0.27 80 Mfs Entisols Swamps

II1E 0.23 0.01 4 0.23 0.24 0.21 80 Mfs Entisols Swamps

II1E 0.12 0.00 2 0.12 0.12 0.11 80 Mfs Entisols Swamps Roots & lenticels

II1F 0.37 1 86 Mfs Entisols Swamps Mature

II1F 0.51 1 86 Mfs Entisols Swamps Inmature
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

244

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II6 0.61 1 86 Mfs Entisols Swamps iInmature

II6 0.50 1 86 Mfs Entisols Swamps Mature

II9A 0.29 0.15 11 0.28 0.70 0.12 73 Dfs Mangle Dune deposits

II9B 0.40 0.09 11 0.38 0.61 0.31 73 Dfs Mangle Dune deposits

II9C 5.63 0.74 11 0.64 0.96 0.37 73 Dfs Mangle Dune deposits

II9D 0.31 0.08 11 0.29 0.44 0.21 73 Dfs Mangle Dune deposits

Potassium (mg/g) II1A 2.90 0.20 3 85 Dfs Mangle Dune deposits

II1C 1.40 0.10 3 85 Dfs Mangle Dune deposits

II1G 2.40 0.10 3 85 Dfs Mangle Dune deposits

II6 2.60 0.30 3 85 Dfs Mangle Dune deposits Seedlings

II1A 2.10 4.00 7 2.00 2.80 1.60 72 Dfs Mangle Mangle Hipersaline lagoon

II1A 1.80 0.50 6 1.90 2.50 1.20 72 Dfs Mangle Mangle Canal

II1A 2.00 0.80 8 1.80 3.50 1.00 72 Dfs Mangle Mangle Border-control

II1A 6.61 2.93 5 7.35 9.83 5.94 75 Dfs Mangle Dune deposits Seedlings

II1A 8.85 1.03 2 8.85 9.58 8.12 74 Dfs Mangle Dune deposits Yellow-fringe

II1A 8.90 0.25 2 8.90 9.07 8.72 74 Dfs Mangle Dune deposits Old-fringe

II1A 5.56 0.13 2 5.56 5.65 5.48 74 Dfs Mangle Dune deposits Senescen-fringe

II1A 10.55 0.04 2 10.55 10.57 10.53 74 Dfs Mangle Dune deposits Young m.-fringe

II1A 6.41 0.42 2 6.41 6.70 6.11 74 Dfs Mangle Dune deposits Old mature-salina

II1A 6.56 0.05 2 6.56 6.60 6.56 74 Dfs Mangle Dune deposits Senescent-salina

II1A 6.29 0.15 2 6.29 6.40 6.19 74 Dfs Mangle Dune deposits Yellow-salina

II1A 8.57 0.17 2 8.57 8.69 8.45 74 Dfs Mangle Dune deposits Young m.-salina

II1A 9.67 0.61 2 9.67 10.10 9.24 74 Dfs Mangle Dune deposits Yellow-fringe

II1A 7.34 0.50 2 7.34 7.69 6.99 74 Dfs Mangle Dune deposits Old-fringe

II1A 10.56 3.89 5 11.38 14.63 3.91 77 Mfs Swamp Swamp

II1A 20.29 0.12 2 20.29 20.38 20.21 78 Mfs Mangle Swamp

II1A 6.03 0.35 36 6.07 6.52 5.32 80 Mfs Entisols Swamps

II1A 6.11 0.32 12 6.07 6.69 5.45 81 Mfs Entisols Swamps Adult 

II1A 5.64 1.18 12 5.64 7.66 3.96 81 Mfs Entisols Swamps Dead 
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

245

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 5.88 0.46 12 5.81 6.67 4.89 81 Mfs Entisols Swamps Old

II1A 5.98 1.81 12 6.48 7.90 2.75 81 Mfs Entisols Swamps Yellow 

II1A 5.79 0.18 12 5.79 6.13 5.55 81 Mfs Entisols Swamps Young 

II1A 5.08 0.30 4 5.05 5.48 4.76 82 Mfs Entisols Swamps

II1Ai 4.39 1.93 5 4.99 7.00 2.34 75 Dfs Mangle Dune deposits

II1Aii 6.29 1.49 5 6.00 8.73 4.99 75 Dfs Mangle Dune deposits

II1B 2.60 0.06 2 2.60 2.64 2.56 82 Mfs Entisols Swamps Dead wood

II1B 1.24 0.30 3 1.23 1.55 0.94 87 Mfs Entisols Compound dunes

II1B 0.75 0.46 2 0.75 0.42 1.07 157 Mfs Entisols Compound dunes

II1C 2.23 0.08 4 2.24 2.30 2.12 80 Mfs Entisols Swamps

II1C 2.80 0.83 18 2.39 4.26 1.96 80 Mfs Entisols Swamps Young stems

II1C 3.48 0.21 4 3.53 3.67 3.18 82 Mfs Entisols Swamps Woody roots

II1E 3.99 0.42 3 4.06 4.36 3.54 80 Mfs Entisols Swamps

II1E 4.64 0.37 4 4.71 5.02 4.13 80 Mfs Entisols Swamps

II1E 5.17 0.05 2 5.17 5.20 5.13 80 Mfs Entisols Swamps Roots & lenticels

II9A 5.78 2.00 11 5.06 9.13 3.08 73 Dfs Mangle Dune deposits

II9B 4.32 1.23 11 4.22 6.56 2.54 73 Dfs Mangle Dune deposits

II9C 5.63 0.74 11 5.56 6.68 4.55 73 Dfs Mangle Dune deposits

II9D 3.95 0.64 11 4.10 4.78 3.00 73 Dfs Mangle Dune deposits

Sodium (mg/g) II1A 9.84 0.23 2 9.84 10.00 9.84 74 Dfs Mangle Dune deposits Senescen-fringe

II1A 16.09 0.09 2 16.09 16.16 16.03 74 Dfs Mangle Dune deposits Young m.-fringe

II1A 22.98 1.44 2 22.98 24.00 21.97 74 Dfs Mangle Dune deposits Old mature-salina

II1A 34.47 0.27 2 34.47 34.66 34.28 74 Dfs Mangle Dune deposits Senescent-salina

II1A 40.85 0.65 2 40.85 41.31 40.38 74 Dfs Mangle Dune deposits Yellow-salina

II1A 26.88 0.47 2 26.88 27.22 26.55 74 Dfs Mangle Dune deposits Mature-salina

II1A 19.91 0.74 2 19.91 20.43 19.38 74 Dfs Mangle Dune deposits Yellow-fringe

II1A 14.61 0.53 2 14.61 14.99 14.24 74 Dfs Mangle Dune deposits Old-fringe

II1A 11.87 7.49 5 7.42 22.62 3.78 77 Mfs Swamp Swamp

II1A 14.00 0.27 2 14.00 14.19 13.81 78 Mfs Mangle Swamp
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

246

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 11.73 1.70 36 11.58 14.60 9.20 80 Mfs Entisols Swamps

II1A 12.26 1.22 12 12.17 14.75 10.65 81 Mfs Entisols Swamps Adult 

II1A 16.18 4.40 12 16.72 22.15 9.67 81 Mfs Entisols Swamps Dead 

II1A 14.73 4.18 12 14.60 20.35 10.06 81 Mfs Entisols Swamps Old

II1A 17.74 5.24 12 17.87 24.52 10.69 81 Mfs Entisols Swamps Yellow 

II1A 11.05 0.83 12 11.25 12.16 9.45 81 Mfs Entisols Swamps Young 

II1C 10.43 0.54 4 10.46 11.01 9.79 80 Mfs Entisols Swamps

II1C 11.01 1.21 18 10.94 15.10 9.26 80 Mfs Entisols Swamps Young stems

II1E 27.28 1.06 3 26.71 28.50 26.62 80 Mfs Entisols Swamps

II1E 48.24 5.96 4 49.31 53.73 40.62 80 Mfs Entisols Swamps

II1E 11.52 0.30 2 11.52 11.74 11.31 80 Mfs Entisols Swamps Roots & lenticels

II9A 16.25 3.79 11 17.05 20.73 6.72 73 Dfs Mangle Dune deposits

II9B 9.32 2.69 11 8.66 13.59 5.09 73 Dfs Mangle Dune deposits

II9C 9.77 2.08 11 9.16 14.65 7.33 73 Dfs Mangle Dune deposits

II9D 10.20 1.53 11 10.29 12.88 7.76 73 Dfs Mangle Dune deposits

Sulfur (mg/g) II1A 0.60 0.29 4 0.60 0.87 0.35 82 Mfs Entisols Swamps

II1B 0.11 0.01 2 0.11 0.12 0.10 82 Mfs Entisols Swamps Dead wood

II1C 0.14 0.02 4 0.14 0.16 0.12 82 Mfs Entisols Swamps Woody roots

Rourea surinamensis Aluminum (mg/g) II4 0.12 0.12 7 0.07 0.38 0.05 135 Lmrf Ultisols Tuffac. sandstone

C/N II4 41 5 2 41 44 37 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II4 5.18 0.85 7 5.44 6.15 3.93 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II4 45 3 2 45 47 43 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II4 0.11 0.12 7 0.05 0.37 0.03 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II4 2.07 0.56 7 2.28 2.67 1.18 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II4 0.60 0.48 7 0.46 1.59 0.09 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II4 0.93 0.22 9 0.93 1.27 0.62 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II4 0.69 0.26 7 0.69 1.12 0.36 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II4 6.92 2.34 7 7.13 10.93 3.97 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

247

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Sulfur (%) II4 0.31 0.04 2 0.31 0.34 0.29 135 Lmrf Ultisols Tuffac. sandstone

Roystonea borinquena Aluminum (mg/g) II1F 0.05 0.02 2 0.05 0.06 0.04 119

II1F 0.06 0.01 2 0.06 0.07 0.05 119 Fruits-no seeds

Ash (%) II1F 2.82 0.57 2 2.82 3.22 2.41 119

II1F 9.05 0.04 2 9.05 9.07 9.02 119 Fruits-no seeds

Calcium (mg/g) II1F 1.51 0.04 2 1.51 1.54 1.48 119

II1F 13.79 0.66 2 13.79 14.26 13.32 119 Fruits-no seeds

Iron (mg/g) II1F 0.47 0.02 2 0.47 0.48 0.46 119

II1F 0.04 0.01 2 0.04 0.05 0.04 119 Fruits-no seeds

Magnesium (mg/g) II1F 0.91 0.03 2 0.91 0.93 0.89 119

II1F 3.18 0.14 2 3.18 3.27 3.08 119 Fruits-no seeds

Manganese (mg/kg) II1F 15 10 2 15 15 14 119

II1F 21 1 2 21 21 20 119 Fruits-no seeds

Nitrogen (%) II1F 1.15 0.01 2 1.15 1.15 1.14 119

II1F 0.51 0.01 2 0.55 0.50 0.50 119 Fruits-no seeds

Phosphorus (mg/g) II1F 1.74 0.04 2 1.74 1.77 1.71 119

II1F 0.92 0.04 2 0.92 0.94 0.89 119 Fruits-no seeds

Potassium (mg/g) II1F 4.74 0.12 2 4.74 4.83 4.66 119

II1F 18.10 0.10 2 18.10 18.17 18.03 119 Fruits-no seeds

Sabicea hirsuta Aluminum (mg/g) II3 0.18 0.10 6 0.14 0.31 0.09 135 Lmrf Ultisols Tuffac. sandstone

C/N II3 41 15 2 41 52 30 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II3 3.75 1.38 6 3.27 6.44 2.64 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II3 45 0 2 45 45 45 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II3 0.20 0.15 6 0.15 0.45 0.06 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II3 2.14 0.71 6 2.10 3.05 1.30 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II3 0.41 0.21 6 0.36 0.78 0.16 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II3 1.28 0.50 8 1.05 2.04 0.74 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

248

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Phosphorus (mg/g) II3 0.81 0.24 6 0.69 1.17 0.61 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II3 23.80 10.17 6 22.92 42.45 11.88 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II3 0.13 0.07 2 0.13 0.18 0.08 135 Lmrf Ultisols Tuffac. sandstone

Sapindus saponaria Aluminum (mg/g) II1A 0.48 1 129 Lmrf Ultisols Tuffac. sandstone

Ash (%) II1A 3.69 1 129 Lmrf Ultisols Tuffac. sandstone

C/N II1A 15 1 129 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II1A 7.10 1 129 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II1A 55 1 129 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II1A 0.48 1 129 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II1A 0.77 1 129 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II1A 0.14 1 129 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II1A 3.62 1 129 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.70 1 129 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II1A 2.58 1 129 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II1A 0.51 1 129 Lmrf Ultisols Tuffac. sandstone

Sapium laurocerasus Aluminum (mg/g) II1A 0.27 0.44 15 0.09 1.62 0.01 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.29 0.42 15 0.06 1.12 0.00 135 Lmrf Ultisols Tuffac. sandstone

II1E 0.53 0.73 8 0.11 2.02 0.03 135 Lmrf Ultisols Tuffac. sandstone

II1H 0.27 0.01 2 0.27 0.28 0.27 135 Lmrf Ultisols Tuffac. sandstone

II8A 0.25 0.12 7 0.24 0.40 0.08 145 Wfs Ultisols Tuffac. sandstone

Ash (%) II1A 7.22 1 71 Lmrf Ultisols Tuffac. sandstone

II1B 2.56 1.13 31 2.43 5.58 1.05 68 Swf Ultisols Tuffac. sandstone Class 1 LD

II1B 1.80 0.81 8 1.66 3.62 1.02 68 Swf Ultisols Tuffac. sandstone Class 2 LD

II1B 1.73 0.72 8 1.75 2.95 0.71 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 1.66 0.20 2 1.66 1.80 1.52 68 Swf Ultisols Tuffac. sandstone Class 4 LD

II1B 2.52 1.11 3 3.02 3.29 1.25 68 Swf Ultisols Tuffac. sandstone Class 2 ST

II8A 11.90 1.12 7 12.01 13.25 10.31 145 Wfs Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

249

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

C/N II8A 40 2 7 40 42 37 145 Wfs Ultisols Tuffac. sandstone

Calcium (mg/g) II1A 7.57 4.08 15 7.69 13.98 1.69 135 Lmrf Ultisols Tuffac. sandstone

II1B 8.30 3.42 31 8.06 16.55 3.42 68 Swf Ultisols Tuffac. sandstone Class 1 LD

II1B 3.10 1.92 8 5.64 5.65 0.98 68 Swf Ultisols Tuffac. sandstone Class 2 LD

II1B 2.76 1.21 8 2.70 4.51 1.16 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 2.55 0.35 2 2.55 2.80 2.30 68 Swf Ultisols Tuffac. sandstone Class 4 LD

II1B 7.02 4.62 3 7.17 11.56 2.33 68 Swf Ultisols Tuffac. sandstone Class 2 ST

II1C 2.44 1.29 15 2.10 5.15 0.98 135 Lmrf Ultisols Tuffac. sandstone

II1E 6.02 4.27 8 6.45 11.99 1.11 135 Lmrf Ultisols Tuffac. sandstone

II1H 7.15 1.20 2 7.15 8.00 6.30 135 Lmrf Ultisols Tuffac. sandstone

II8A 16.73 1.67 7 16.03 19.89 14.86 145 Wfs Ultisols Tuffac. sandstone

Carbon (%) II1A 51 1 71 Lmrf Ultisols Tuffac. sandstone

II8A 47 1 7 46 48 45 145 Wfs Ultisols Tuffac. sandstone

Iron (mg/g) II1A 0.37 0.59 15 0.11 1.96 0.01 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.29 0.42 15 0.08 1.14 0.01 135 Lmrf Ultisols Tuffac. sandstone

II1E 0.56 0.76 8 0.27 2.31 0.05 135 Lmrf Ultisols Tuffac. sandstone

II1H 0.23 0.09 2 0.23 0.30 0.17 135 Lmrf Ultisols Tuffac. sandstone

II8A 0.25 0.15 7 0.24 0.45 0.10 145 Wfs Ultisols Tuffac. sandstone

Magnesium (mg/g) II1A 4.52 2.11 15 5.38 6.69 1.06 135 Lmrf Ultisols Tuffac. sandstone

II1C 1.64 1.01 15 1.31 4.12 0.94 135 Lmrf Ultisols Tuffac. sandstone

II1E 4.04 2.23 8 4.36 6.95 1.44 135 Lmrf Ultisols Tuffac. sandstone

II1H 4.66 1.52 2 4.66 5.73 3.59 135 Lmrf Ultisols Tuffac. sandstone

II8A 4.32 0.49 7 4.04 5.05 3.84 145 Wfs Ultisols Tuffac. sandstone

Manganese (mg/g) II1A 0.14 0.05 15 0.12 0.24 0.08 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.08 0.04 15 0.06 0.15 0.04 135 Lmrf Ultisols Tuffac. sandstone

II1E 0.19 0.11 8 0.19 0.38 0.07 135 Lmrf Ultisols Tuffac. sandstone

II1H 0.26 0.01 2 0.26 0.27 0.26 135 Lmrf Ultisols Tuffac. sandstone

II8A 0.21 0.04 7 0.21 0.28 0.15 145 Wfs Ultisols Tuffac. sandstone

Nitrogen (%) II1A 1.88 1 71 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

250

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 1.44 0.82 14 1.76 2.46 0.24 135 Lmrf Ultisols Tuffac. sandstone

II1B 0.22 0.07 31 0.20 0.46 0.12 68 Swf Ultisols Tuffac. sandstone Class 1 LD

II1B 0.19 0.08 8 0.17 0.31 0.10 68 Swf Ultisols Tuffac. sandstone Class 2 LD

II1B 0.33 0.25 8 0.21 0.79 0.14 68 Swf Ultisols Tuffac. sandstone Class 3 LD

II1B 0.66 0.01 2 0.66 0.67 0.65 68 Swf Ultisols Tuffaceoussandstone Class 4 LD

II1B 0.23 0.09 3 0.27 0.29 0.13 68 Swf Ultisols Tuffac. sandstone Class 2 ST

II1C 0.36 0.13 15 0.40 0.52 0.05 135 Lmrf Ultisols Tuffac. sandstone

II1E 1.02 0.62 8 1.06 1.68 0.28 135 Lmrf Ultisols Tuffac. sandstone

II1H 1.42 0.42 2 1.42 1.72 1.12 135 Lmrf Ultisols Tuffac. sandstone

II8A 1.17 0.05 7 1.15 1.24 1.11 145 Wfs Ultisols Tuffac. sandstone

Phosphorus (mg/g) II1A 1.13 0.53 15 1.47 1.69 0.28 135 Lmrf Ultisols Tuffac. sandstone

II1B 0.12 0.16 31 0.09 0.98 0.03 68 Swf Ultisols Tuffac. sandstone Class 1 LD

II1B 0.08 0.04 8 0.09 0.16 0.03 68 Swf Ultisols Tuffac. sandstone Class 2 LD

II1B 0.13 0.08 8 0.12 0.26 0.02 68 Swf Ultisols Tuffaceoussandstone Class 3 LD

II1B 0.27 0.01 2 0.27 0.28 0.27 68 Swf Ultisols Tuffac. sandstone Class 4 LD

II1B 0.09 0.04 3 0.10 0.12 0.05 68 Swf Ultisols Tuffac. sandstone Class 2 ST

II1C 0.47 0.18 1 0.48 0.84 0.25 135 Lmrf Ultisols Tuffac. sandstone

II1E 0.85 0.41 8 0.97 1.34 0.34 135 Lmrf Ultisols Tuffac. sandstone

II1H 1.20 0.57 2 1.20 1.61 0.80 135 Lmrf Ultisols Tuffac. sandstone

II8A 0.62 0.12 7 0.68 0.72 0.42 145 Wfs Ultisols Tuffac. sandstone

Potassium (mg/g) II1A 20.54 9.33 15 23.65 32.72 4.44 135 Lmrf Ultisols Tuffac. sandstone

II1B 0.76 0.22 8 1.56 1.12 0.47 68 Swf Ultisols Tuffac. sandstone Class 2 LD

II1B 0.86 0.72 8 0.66 2.38 0.23 68 Swf Ultisols Tuffaceoussandstone Class 3 LD

II1B 0.29 0.02 2 0.29 0.30 0.27 68 Swf Ultisols Tuffac. sandstone Class 4 LD

II1B 2.80 1.68 3 2.94 4.41 1.05 68 Swf Ultisols Tuffac. sandstone Class 2 ST

II1B 2.08 1.26 31 1.81 6.20 0.66 68 Swf Ultisols Tuffac. sandstone Class 1 LD

II1C 10.25 4.41 15 10.84 16.93 4.98 135 Lmrf Ultisols Tuffac. sandstone

II1E 15.75 11.46 8 13.57 29.91 3.84 135 Lmrf Ultisols Tuffac. sandstone

II1H 26.94 1.94 2 26.94 28.31 25.57 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

251

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8A 7.03 2.09 7 7.29 9.58 4.69 145 Wfs Ultisols Tuffac. sandstone

Sulfur (%) II1A 0.50 1 71 Lmrf Ultisols Tuffac. sandstone

II8A 0.52 0.08 7 0.57 0.60 0.38 145 Wfs Ultisols Tuffac. sandstone

Sauvagesia erecta Aluminum (mg/g) II3 0.19 1 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II3 3.42 1 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II3 0.16 1 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II3 1.33 1 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II3 0.74 1 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II3 0.70 1 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II3 0.33 1 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II3 7.83 1 135 Lmrf Ultisols Tuffac. sandstone

Schefflera morototoni Ash (%) II1A 7.90 0.47 6 7.85 8.45 7.40 126 Urban forest, RP

II1A 5.82 0.40 2 5.85 6.10 5.54 94 Swf Ultisols Tuffac. sandstone

II1B 2.11 1 68 Swf Ultisols Tuffac. sandstone Class 4 LD

II1B 2.49 0.85 2 2.49 3.09 1.89 94 Swf Ultisols Tuffac. sandstone

II1C 1.21 0.09 6 1.22 1.35 1.09 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 3.19 0.95 3 3.48 3.96 2.13 94 Swf Ultisols Tuffac. sandstone

II1Ci 8.50 0.76 5 8.38 9.54 7.61 126 Ultisols Siltstone,sandstone Urban forest, RP

II8A 5.56 1 71 Wfs Ultisols Tuffac. sandstone

C/N II1A 25 3 6 25 28 21 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 292 24 6 296 322 258 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 72 5 5 73 79 67 126 Ultisols Siltstone,sandstone Urban forest, RP

Calcium (mg/g) II1A 6.77 1.01 7 7.26 7.58 4.94 135 Lmrf Ultisols Tuffac. sandstone

II1B 4.30 1 68 Swf Ultisols Tuffac. sandstone Class 4 LD

II1B 22.71 5.74 5 22.73 29.99 17.08 135 Lmrf Ultisols Tuffac. sandstone

II1C 1.11 0.12 9 1.13 1.29 0.89 135 Lmrf Ultisols Tuffac. sandstone

II1Ci 9.13 0.97 8 9.20 10.46 7.17 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

252

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Carbon (%) II1A 51 1 6 51 52 51 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 52 0 6 52 52 52 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 48 0 5 48 49 48 126 Ultisols Siltstone,sandstone Urban forest, RP

II8A 56 1 71 Wfs Ultisols Tuffac. sandstone

Magnesium (mg/g) II1A 3.45 0.72 7 3.08 4.37 2.52 135 Lmrf Ultisols Tuffac. sandstone

II1B 4.87 0.93 5 4.82 5.90 3.84 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.76 0.08 9 0.73 0.89 0.66 135 Lmrf Ultisols Tuffac. sandstone

II1Ci 1.74 0.09 8 1.75 1.85 1.61 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II1A 2.08 0.25 6 2.01 2.38 1.82 126 Lmrf Ultisols Tuffac. sandstone Urban forest, RP

II1A 2.14 0.02 2 2.14 2.15 2.12 94 Swf Ultisols Tuffac. sandstone

II1A 1.32 0.42 7 1.51 1.66 1.69 135 Lmrf Ultisols Tuffac. sandstone

II1B 1.04 1 68 Swf Ultisols Tuffac. sandstone Class 4 LD

II1B 0.45 0.16 2 0.45 0.56 0.33 94 Swf Ultisols Tuffac. sandstone

II1B 0.68 0.06 5 0.66 0.78 0.63 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.18 0.01 6 0.18 0.20 0.16 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 0.43 0.11 3 0.40 0.56 0.34 94 Swf Ultisols Tuffac. sandstone

II1C 0.08 0.00 9 0.08 0.08 0.07 35 Lmrf Ultisols Tuffac. sandstone

II1Ci 0.67 0.04 5 0.66 0.71 0.62 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 0.35 0.04 8 0.35 0.39 0.28 135 Lmrf Ultisols Tuffac. sandstone

II8A 0.87 1 71 Wfs Ultisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.97 0.10 2 0.97 1.03 0.90 94 Swf Ultisols Tuffac. sandstone

II1A 0.83 0.24 7 0.78 1.15 0.56 135 Lmrf Ultisols Tuffac. sandstone

II1B 0.23 1 68 Swf Ultisols Tuffac. sandstone Class 4 LD

II1B 0.29 0.12 2 0.29 0.37 0.20 94 Swf Ultisols Tuffac. sandstone

II1B 0.46 0.09 5 0.45 0.56 0.34 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.31 0.09 3 0.27 0.41 0.25 94 Swf Ultisols Tuffac. sandstone

II1C 0.04 0.01 9 0.04 0.07 0.03 135 Lmrf Ultisols Tuffac. sandstone

II1Ci 0.15 0.03 8 0.16 0.19 0.12 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II1A 11.94 0.79 2 11.94 12.50 11.38 94 Swf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

253

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 7.31 1.47 7 6.99 9.43 5.07 135 Lmrf Ultisols Tuffac. sandstone

II1B 0.51 1 68 Swf Ultisols Tuffac. sandstone Class 4 LD

II1B 4.09 3.87 2 4.09 6.82 1.35 94 Swf Ultisols Tuffac. sandstone

II1B 3.92 0.89 5 4.19 4.74 2.61 135 Lmrf Ultisols Tuffac. sandstone

II1C 5.11 2.25 3 4.93 7.45 2.96 94 Swf Ultisols Tuffac. sandstone

II1C 0.71 0.15 9 0.67 0.91 0.49 135 Lmrf Ultisols Tuffac. sandstone

II1Ci 2.24 0.50 8 2.16 3.05 1.55 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II1A 0.35 0.04 6 0.33 0.43 0.31 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 0.01 0.01 4 0.01 0.02 0.01 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 0.07 0.01 5 0.07 0.83 0.05 126 Ultisols Siltstone,sandstone Urban forest, RP

II8A 0.32 1 71 Wfs Ultisols Tuffac. sandstone

Wood density (g/cc) II1C 0.16 0.03 5 0.17 0.20 0.13 126 Ultisols Siltstone,sandstone Urban forest, RP

Schizolobium parahybum Wood density (g/cc) II1C 0.29 0.03 5 0.28 0.33 0.26 97 Wfs Inceptisols Tuffac. sandstone

Schlegelia brachyantha Aluminum (mg/g) II1 0.02 1 135 Lmrf Ultisols Tuffac. sandstone

C/N II1 65 1 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II1 6.50 1 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II1 46 1 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II1 0.23 1 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II1 6.16 1 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II1 0.15 1 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II1 0.71 1 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II1 0.30 1 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II1 2.47 1 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II1 0.15 1 135 Lmrf Ultisols Tuffac. sandstone

Scleria canescens Aluminum (mg/g) II3 0.27 1 135 Lmrf Ultisols Tuffac. sandstone

C/N II3 20 1
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

254

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Calcium (mg/g) II3 0.96 1 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II3 35 1 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II3 0.06 1 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II3 0.73 1 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II3 0.74 1 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II3 1.10 0.93 2 1.10 1.75 0.44 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II3 0.93 1 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II3 10.10 1 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II3 0.35 1 135 Lmrf Ultisols Tuffac. sandstone

Scleria spp. Aluminum (mg/g) II3 0.39 0.29 30 0.34 1.81 0.11 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II3 1.85 0.48 30 1.80 2.80 1.06 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II3 0.20 0.21 30 0.16 1.21 0.06 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II3 1.22 0.20 30 1.21 1.70 0.72 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II3 0.74 0.44 30 0.63 2.16 0.10 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II3 0.78 0.26 30 0.72 1.35 0.44 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II3 0.59 0.27 30 0.52 1.26 0.20 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II3 11.18 6.16 30 8.71 24.96 4.00 135 Lmrf Ultisols Tuffac. sandstone

Secondary Forest Aluminum (mg/g) II1A 4.76 5.62 55 2.60 23.76 0.20 112 Lmrf Ultisols Tuffac. sandstone Bisley

II1A 4.94 5.37 51 2.67 20.63 0.20 112 Lmrf Ultisols Tuffac. sandstone Puente Roto

II8B 1.33 0.83 8 1.46 2.31 0.32 130 Mfs Ultisols Tuffac. sandstone <=2cm

II8B 0.69 0.71 9 0.46 2.37 0.06 130 Mfs Ultisols Tuffac. sandstone >2cm

II8D 11.11 3.25 10 11.07 16.25 6.43 124 Lmrf Ultisols Tuffac. sandstone Bisley

II9A 2.39 1.50 22 1.78 6.37 0.57 132 Wfs Mollisols Plutonic rocks

II1E 8.27 2.97 16 8.14 14.39 3.67 114 Wfs Ultisols Lava Lives <2mm

II1E 13.73 5.17 16 15.06 21.69 3.49 114 Wfs Ultisols Lava Dead roots <2mm

II1E 5.92 4.04 6 5.00 13.37 1.24 114 Wfs Ultisols Lava Live >2mm<5mm

II1E 5.70 2.59 5 5.32 9.26 2.88 114 Wfs Ultisols Lava Dead >2mm<5mm
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

255

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1E 3.44 1.60 6 3.20 5.79 1.59 114 Wfs Ultisols Lava >5mm

II1E 4.44 2.21 29 4.06 8.56 0.94 131 Wfs Mollisols Plutonic rocks <1cm

II1E 1.98 1.20 19 1.70 4.57 0.58 131 Wfs Mollisols Plutonic rocks >1cm<2cm

II1E 2.52 2.61 5 1.70 7.13 0.86 131 Wfs Mollisols Plutonic rocks >2cm

II3 4.60 3.04 3 3.54 10.99 1.77 142 Wflm Inceptisols Tuffac. sandstone

II3 5.44 2.88 17 5.45 11.66 1.81 142 Wflm Inceptisols Tuffac. sandstone

II6 6.77 4.47 17 6.48 13.92 0.57 142 Wflm Inceptisols Tuffac. sandstone

II8A 3.04 2.89 16 1.54 9.95 0.89 111 Rfs Inceptisols Tuffac. sandstone

II8A 1.74 1.30 12 1.49 4.02 0.37 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8A 7.69 4.09 32 7.13 15.29 1.82 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8A 1.70 1.03 11 1.54 4.31 0.47 124 Wflm Ultisols Tuffac. sandstone Dacroides-Adjuntas

II8A 3.59 2.92 17 3.16 9.61 0.29 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8A 4.93 4.46 9 4.16 13.03 0.56 130 Mfs Ultisols Tuffac. sandstone

II8A 9.15 6.15 35 7.46 23.53 2.21 132 Wfs Mollisols Plutonic rocks

II8A 7.21 5.21 69 6.23 26.62 1.17 136 Wfs Ultisols Tuffac. sandstone

II8A 2.49 1.41 66 2.21 7.23 0.18 140 Swf Ultisols Tuffac. sandstone Cubuy

II8A 0.68 0.25 6 0.71 0.96 0.35 140 Sdf Mollisols Alluvial deposits Guanica

II8A 2.99 3.15 144 2.08 19.54 0.10 140 Lmrf Ultisols Tuffac. sandstone Bisley

II8A 1.28 1.03 75 1.10 6.75 0.21 140 Wflm Inceptisols Tuffac. sandstone Palm Forest

II8A 2.53 1.53 20 2.22 5.02 0.36 140 Rfs Inceptisols Tuffac. sandstone Pico del Este

II8A 6.65 4.97 20 5.57 23.59 1.10 142 Wflm Inceptisols Tuffac. sandstone Dicotiledonea

II8A 4.73 2.95 21 3.54 9.99 0.72 142 Wflm Inceptisols Tuffac. sandstone

II8B 0.65 0.61 12 0.43 1.93 0.08 111 Rfs Inceptisols Tuffac. sandstone

II8B 2.14 1.32 13 1.82 5.85 0.85 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8B 4.15 3.40 8 2.26 9.13 1.41 124 Dacroides

II8B 6.63 6.20 48 5.03 27.80 0.08 132 Wfs Mollisols Plutonic rocks

II8B 1.53 1.70 51 1.04 8.17 0.18 136 Wfs Ultisols Tuffac. sandstone

II8B 1.00 0.71 65 0.71 2.76 0.06 140 Swf Ultisols Tuffac. sandstone Cubuy

II8B 0.31 0.20 11 0.27 0.64 0.07 140 Sdf Mollisols Alluvial deposits Guanica
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

256

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8B 1.25 1.65 139 0.75 13.05 0.04 140 Lmrf Ultisols Tuffac. sandstone Bisley

II8B 1.09 1.03 66 0.74 6.63 0.18 140 Wflm Inceptisols Tuffac. sandstone Palm Forest

II8B 1.28 0.94 17 1.11 3.12 0.19 140 Rfs Inceptisols Tuffac. sandstone Pico del Este

II8B 2.70 1.58 31 2.73 7.37 0.26 142 Wflm Inceptisols Tuffac. sandstone

II8C 3.19 3.22 16 2.06 14.14 0.98 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8C 2.03 0.69 7 2.06 3.26 1.01 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8C 2.26 1.69 8 1.89 5.59 0.44 130 Mfs Ultisols Tuffac. sandstone

II8C 9.46 12.64 21 3.12 40.30 0.42 132 Wfs Mollisols Plutonic rocks

II8C 0.33 1.64 2 4.33 5.49 3.17 142 Wflm Inceptisols Tuffac. sandstone

II8D 3.41 3.93 16 1.84 13.47 1.34 111 Rfs Inceptisols Tuffac. sandstone

II8D 19.92 3.01 9 20.75 23.81 14.23 124 Lmrf Ultisols Tuffac. sandstone Adjuntas

II8D 12.53 4.53 8 12.02 19.02 6.40 130 Mfs Ultisols Tuffac. sandstone

II8D 14.28 10.69 27 15.96 40.31 0.10 132 Wfs Mollisols Plutonic rocks

II8D 6.40 2.44 62 6.40 10.62 0.43 140 Swf Ultisols Tuffac. sandstone Cubuy

II8D 5.77 1.63 7 6.83 7.31 3.19 140 Sdf Mollisols Alluvial deposits Guanica

II8D 4.67 5.11 138 3.17 32.92 0.21 140 Lmrf Ultisols Tuffac. sandstone Bisley

II8D 4.30 3.38 67 3.32 17.06 0.29 140 Wflm Inceptisols Tuffac. sandstone Palm Forest

II8D 12.81 10.10 9 9.60 34.68 2.74 140 Rfs Inceptisols Tuffac. sandstone Pico del Este

II8D 8.29 3.84 8 8.32 13.11 2.47 142 Wflm Inceptisols Tuffac. sandstone Fern

II8D 14.63 6.79 42 13.78 27.17 4.18 142 Wflm Inceptisols Tuffac. sandstone

II9A 0.65 0.39 12 0.59 1.35 0.21 124

II9A 4.97 4.18 10 2.91 13.10 1.28 124

II9A 0.96 0.96 583 0.61 6.41 0.02 106 Lmrf Ultisols Tuffac. sandstone

II9A 1.45 1.25 138 1.08 6.59 0.13 130 Mfs Ultisols Tuffac. sandstone

II9A 1.25 0.13 3 1.27 1.37 1.11 153 Swf Ultisols Tuffac. sandstone

II9A 0.57 0.47 14 0.52 1.42 0.06 154 Wfs Ultisols Tuffac. sandstone

II9B 0.53 0.31 11 0.42 1.06 0.15 124

II9B 0.53 1.17 420 0.17 8.75 0.02 106 Lmrf Ultisols Tuffac. sandstone

II9B 0.97 1.44 96 0.43 9.05 0.02 130 Mfs Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

257

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II9B 2.79 2.68 29 1.97 8.92 0.11 132 Wfs Mollisols Plutonic rocks

II9B 0.13 0.14 14 0.09 0.46 0.04 154 Wfs Ultisols Tuffac. sandstone

II9C 2.17 1.50 15 2.11 5.75 0.42 124

II9C 0.68 0.77 463 0.45 8.41 0.01 106 Lmrf Ultisols Tuffac. sandstone

II9C 0.89 1.38 129 0.33 8.15 0.01 130 Mfs Ultisols Tuffac. sandstone

II9C 1.35 1.19 21 0.96 4.21 0.07 132 Wfs Mollisols Plutonic rocks

II9D 9.10 2.73 5 10.49 11.57 0.42 124 Dacroides 

II9D 1.23 1.63 464 0.82 20.70 0.04 106 Lmrf Ultisols Tuffac. sandstone

II9D 9.49 7.11 58 7.82 27.17 0.04 130 Mfs Ultisols Tuffac. sandstone

II9D 2.37 2.45 17 2.21 10.14 0.18 132 Wfs Mollisols Plutonic rocks

II9D 0.34 0.31 13 0.13 0.82 0.03 154 Wfs Ultisols Tuffac. sandstone

Ash (%) II1A 11.82 9.14 39 8.66 57.55 2.47 112 Lmrf Ultisols Tuffac. sandstone Bisley

II1A 11.42 8.11 39 9.11 43.26 5.03 112 Lmrf Ultisols Tuffac. sandstone Puente Roto

II8B 4.66 1.22 8 4.61 6.33 2.51 130 Mfs Ultisols Tuffac. sandstone <=2cm

II8B 3.88 1.37 9 3.51 6.24 2.40 130 Mfs Ultisols Tuffac. sandstone >2cm

II8D 24.98 8.34 10 25.10 36.43 14.12 124 Lmrf Ultisols Tuffac. sandstone Bisley

II9A 5.15 0.39 22 5.16 5.98 4.41 132 Wfs Mollisols Plutonic rocks

II1E 5.99 2.23 29 5.19 11.67 2.10 131 Wfs Mollisols Plutonic rocks <1cm

II1E 4.78 1.59 19 4.81 7.79 2.65 131 Wfs Mollisols Plutonic rocks >1cm<2cm

II1E 3.07 0.95 5 2.87 4.66 2.07 131 Wfs Mollisols Plutonic rocks >2cm

II3 12.86 4.23 10 14.29 19.20 6.35 142 Wflm Inceptisols Tuffac. sandstone

II3 17.39 4.99 9 16.54 24.69 10.69 142 Wflm Inceptisols Tuffac. sandstone

II6 9.49 2.96 14 8.83 15.44 3.11 142 Wflm Inceptisols Tuffac. sandstone

II8A 13.40 1.74 12 13.55 15.75 9.74 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8A 18.14 6.47 32 15.60 33.79 10.00 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8A 12.62 1.16 1 12.30 14.38 11.22 124 Dacroides-Adjuntas

II8A 13.33 3.12 17 13.42 19.79 8.90 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8A 13.84 5.90 9 12.88 25.99 8.05 130 Mfs Ultisols Tuffac. sandstone

II8A 10.44 7.35 35 9.05 39.90 4.45 132 Wfs Mollisols Plutonic rocks
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

258

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8A 12.36 4.34 66 11.83 26.25 5.86 140 Swf Ultisols Tuffac. sandstone Cubuy

II8A 12.35 2.51 6 12.15 16.24 9.25 140 Sdf Mollisols Alluvial deposits Guanica

II8A 11.04 5.52 143 9.58 43.62 3.78 140 Lmrf Ultisols Tuffac. sandstone Bisley

II8A 5.73 3.21 72 5.12 15.39 0.51 140 Wflm Inceptisols Tuffac. sandstone Palm Forest

II8A 5.38 2.03 18 4.64 10.94 3.13 140 Rfs Inceptisols Tuffac. sandstone Pico del Este

II8A 12.68 5.55 19 11.99 23.01 4.52 142 Wflm Inceptisols Tuffac. sandstone Dicotiledonea

II8A 12.23 6.70 17 11.37 23.25 3.31 142 Wflm Inceptisols Tuffac. sandstone

II8A 5.75 4.25 16 3.19 14.46 2.55 111 Rfs Inceptisols Tuffac. sandstone

II8A 15.67 9.19 69 13.21 56.02 3.09 136 Wfs Ultisols Tuffac. sandstone

II8B 7.26 2.06 13 7.18 12.87 4.92 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8B 8.69 2.24 8 7.92 13.01 5.93 124 Dacroides

II8B 4.79 2.21 48 4.52 11.18 1.26 132 Wfs Mollisols Plutonic rocks

II8B 6.35 2.01 65 6.13 12.83 3.34 140 Swf Ultisols Tuffac. sandstone Cubuy

II8B 8.39 1.82 10 8.58 11.02 5.19 140 Sdf Mollisols Alluvial deposits Guanica

II8B 4.61 3.50 139 3.52 28.12 0.72 140 Lmrf Ultisols Tuffac. sandstone Bisley

II8B 3.60 2.09 65 3.11 11.86 0.64 140 Wflm Inceptisols Tuffac. sandstone Palm Forest

II8B 3.44 1.47 15 3.11 8.00 1.33 140 Rfs Inceptisols Tuffac. sandstone Pico del Este

II8B 6.72 3.12 29 6.83 17.38 1.94 142 Wflm Inceptisols Tuffac. sandstone

II8B 2.18 1.08 12 1.74 4.20 1.17 111 Rfs Inceptisols Tuffac. sandstone

II8B 4.52 3.07 50 3.87 19.85 0.78 136 Wfs Ultisols Tuffac. sandstone

II8C 6.16 3.66 12 4.97 15.20 2.14 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8C 3.77 0.99 7 3.96 5.16 2.34 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8C 4.55 2.61 8 4.07 9.30 2.03 130 Mfs Ultisols Tuffac. sandstone

II8C 4.55 2.42 13 4.24 9.43 0.15 132 Wfs Mollisols Plutonic rocks

II8D 30.00 6.12 9 29.32 38.67 21.74 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8D 24.14 7.22 8 23.27 35.18 13.41 130 Mfs Ultisols Tuffac. sandstone

II8D 20.35 15.84 27 21.86 51.81 1.77 132 Wfs Mollisols Plutonic rocks

II8D 26.65 8.91 62 25.35 57.08 11.49 140 Swf Ultisols Tuffac. sandstone Cubuy

II8D 28.99 3.34 7 30.64 32.09 23.10 140 Sdf Mollisols Alluvial deposits Guanica
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

259

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8D 12.38 8.45 137 10.46 59.23 2.45 140 Lmrf Ultisols Tuffac. sandstone Bisley

II8D 11.92 10.26 66 7.26 41.00 2.36 140 Wflm Inceptisols Tuffac. sandstone Palm Forest

II8D 21.63 12.94 8 23.74 42.75 5.26 140 Rfs Inceptisols Tuffac. sandstone Pico del Este

II8D 17.97 5.21 2 17.97 21.65 14.28 142 Wflm Inceptisols Tuffac. sandstone Fern

II8D 43.13 23.78 35 38.46 90.18 16.19 142 Wflm Inceptisols Tuffac. sandstone

II8D 5.94 6.04 16 4.26 21.09 1.03 111 Rfs Inceptisols Tuffac. sandstone

II9A 6.52 0.93 7 6.70 8.20 5.29 154 Wfs Ultisols Tuffac. sandstone

II9A 8.41 1.38 87 8.21 11.85 5.45 108 Wf-LM Ultisols Tuffac. sandstone Mameyes

II9A 7.80 4.35 95 6.96 46.08 2.62 108 Wf-LM Ultisols Tuffac. sandstone Sabana

II9A 7.15 1.33 20 6.84 9.55 5.30 113 Wfs Ultisols Lava

II9A 10.91 1.22 12 10.99 12.29 8.52 124

II9A 9.00 2.02 10 8.83 11.95 6.28 124

II9A 8.55 0.99 138 8.55 12.44 5.98 130 Mfs Ultisols Tuffac. sandstone

II9A 8.08 0.21 3 8.17 8.22 7.84 153 Swf Ultisols Tuffac. sandstone

II9B 5.08 0.94 7 4.99 6.53 3.88 154 Wfs Ultisols Tuffac. sandstone

II9B 4.55 1.94 59 4.38 8.94 0.54 108 Wf-LM Ultisols Tuffac. sandstone Mameyes

II9B 4.03 1.55 56 3.81 8.15 1.41 108 Wf-LM Ultisols Tuffac. sandstone Mameyes

II9B 3.78 1.96 16 3.06 9.42 1.76 113 Wfs Ultisols Lava

II9B 4.59 0.85 11 4.80 5.72 3.22 124

II9B 4.25 1.98 96 4.04 13.71 0.63 130 Mfs Ultisols Tuffac. sandstone

II9B 3.42 1.10 28 3.26 5.70 1.87 132 Wfs Mollisols Plutonic rocks

II9C 4.18 1.07 45 4.41 6.20 0.10 108 Wf-LM Ultisols Tuffac. sandstone Mameyes

II9C 4.78 1.28 49 4.85 7.89 2.44 108 Wf-LM Ultisols Tuffac. sandstone Sabana

II9C 4.51 0.81 18 4.24 5.91 3.12 113 Wfs Ultisols Lava

II9C 5.94 3.73 15 6.30 15.08 0.44 124

II9C 4.63 1.90 129 4.57 9.34 1.41 130 Mfs Ultisols Tuffac. sandstone

II9C 3.80 1.23 21 3.65 6.91 2.18 132 Wfs Mollisols Plutonic rocks

II9D 6.19 3.57 6 4.61 12.90 3.65 154 Wfs Ultisols Tuffac. sandstone

II9D 5.49 1.69 38 5.15 9.93 2.20 108 Wf-LM Ultisols Tuffac. sandstone Mameyes
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

260

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II9D 5.58 2.21 32 5.67 9.77 0.76 108 Wf-LM Ultisols Tuffac. sandstone Sabana

II9D 7.18 2.32 10 7.27 10.42 4.06 113 Wfs Ultisols Lava

II9D 21.90 6.77 5 23.60 28.22 0.44 124 Dacroides 

II9D 17.13 10.14 58 13.72 38.43 4.97 130 Mfs Ultisols Tuffac. sandstone

II9D 6.60 1.42 14 6.74 9.66 2.84 132 Wfs Mollisols Plutonic rocks

C/N II1A 25 8 55 25 47 12 112 Lmrf Ultisols Tuffac. sandstone Bisley

II1A 28 13 49 24 72 14 112 Lmrf Ultisols Tuffac. sandstone Puente Roto

II8B 41 13 8 36 67 28 130 Mfs Ultisols Tuffac. sandstone <=2cm

II8B 63 27 9 58 102 28 130 Mfs Ultisols Tuffac. sandstone >2cm

II8D 28 6 10 26 40 23 124 Lmrf Ultisols Tuffac. sandstone Bisley

II9A 27 3 22 28 32 22 132 Wfs Mollisols Plutonic rocks

II1E 39 14 29 35 95 22 131 Wfs Mollisols Plutonic rocks <1cm

II1E 45 22 19 37 114 28 131 Wfs Mollisols Plutonic rocks >1cm<2cm

II1E 35 2 5 34 37 32 131 Wfs Mollisols Plutonic rocks >2cm

II8A 36 3 16 36 41 32 111 Rfs Inceptisols Tuffac. sandstone

II8A 47 4 12 47 55 42 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8A 36 6 32 36 50 28 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8A 51 5 11 51 59 44 124 Dacroides-Adjuntas

II8A 43 8 17 40 55 34 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8A 22 2 9 22 26 19 130 Mfs Ultisols Tuffac. sandstone

II8A 20 3 33 20 25 14 132 Wfs Mollisols Plutonic rocks

II8B 68 14 12 66 99 53 111 Rfs Inceptisols Tuffac. sandstone

II8B 66 11 13 67 79 40 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8B 57 19 8 54 99 35 124 Dacroides

II8B 39 15 49 38 90 27 132 Wfs Mollisols Plutonic rocks

II8C 48 17 12 46 77 25 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8C 43 10 7 40 61 34 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8C 38 5 8 36 45 33 130 Mfs Ultisols Tuffac. sandstone

II8C 22 7 21 21 40 14 132 Wfs Mollisols Plutonic rocks
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

261

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8D 48 23 16 39 111 34 111 Rfs Inceptisols Tuffac. sandstone

II8D 24 3 9 24 28 20 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8D 15 2 8 15 18 13 130 Mfs Ultisols Tuffac. sandstone

II8D 22 11 27 17 53 13 132 Wfs Mollisols Plutonic rocks

II9A 33 9 130 31 97 21 106 Lmrf Ultisols Tuffac. sandstone

II9A 81 18 87 82 121 41 108 Wf-LM Ultisols Tuffac. sandstone Mameyes

II9A 53 15 95 49 94 34 108 Wf-LM Ultisols Tuffac. sandstone Sabana

II9A 68 8 12 70 75 46 124

II9A 56 10 10 57 75 41 124

II9A 25 5 138 25 62 19 130 Mfs Ultisols Tuffac. sandstone

II9A 61 1 3 61 62 59 153 Swf Ultisols Tuffac. sandstone

II9B 65 24 94 60 162 32 106 Lmrf Ultisols Tuffac. sandstone

II9B 50 17 63 48 93 8 108 Wf-LM Ultisols Tuffac. sandstone Mameyes

II9B 34 17 70 30 114 14 108 Wf-LM Ultisols Tuffac. sandstone Mameyes

II9B 80 11 11 80 98 65 124

II9B 47 29 96 41 219 22 130 Mfs Ultisols Tuffac. sandstone

II9B 42 9 30 44 60 26 132 Wfs Mollisols Plutonic rocks

II9C 38 9 100 37 77 18 106 Lmrf Ultisols Tuffac. sandstone

II9C 79 22 45 83 125 28 108 Wf-LM Ultisols Tuffac. sandstone Mameyes

II9C 74 17 49 71 115 38 108 Wf-LM Ultisols Tuffac. sandstone Sabana

II9C 40 20 15 33 100 26 124

II9C 30 10 129 27 62 16 130 Mfs Ultisols Tuffac. sandstone

II9C 25 5 21 23 39 18 132 Wfs Mollisols Plutonic rocks

II9D 43 9 85 42 72 47 106 Lmrf Ultisols Tuffac. sandstone

II9D 70 16 38 70 113 26 108 Wf-LM Ultisols Tuffac. sandstone Mameyes

II9D 54 22 39 51 138 17 108 Wf-LM Ultisols Tuffac. sandstone Sabana

II9D 41 4 5 43 44 15 124 Dacroides 

II9D 19 8 58 17 53 13 130 Mfs Ultisols Tuffac. sandstone

II9D 28 7 17 29 41 16 132 Wfs Mollisols Plutonic rocks
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

262

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Calcium (mg/g) II1A 14.63 4.55 55 14.59 28.09 4.15 112 Lmrf Ultisols Tuffac. sandstone Bisley

II1A 13.88 3.96 51 14.43 22.30 6.83 112 Lmrf Ultisols Tuffac. sandstone Puente Roto

II8B 9.39 1.48 8 9.69 10.74 6.09 130 Mfs Ultisols Tuffac. sandstone <=2cm

II8B 7.61 1.72 9 6.77 9.86 5.17 130 Mfs Ultisols Tuffac. sandstone >2cm

II8D 5.80 1.16 10 6.02 8.00 3.54 124 Lmrf Ultisols Tuffac. sandstone Bisley

II9A 6.58 0.99 22 6.43 8.48 5.00 132 Wfs Mollisols Plutonic rocks

II1E 7.60 1.70 16 7.87 10.46 3.83 114 Wfs Ultisols Lava Lives <2mm

II1E 9.77 2.54 16 10.03 14.63 6.79 114 Wfs Ultisols Lava Dead roots <2mm

II1E 5.99 4.27 6 5.06 12.38 1.35 114 Wfs Ultisols Lava Live >2mm<5mm

II1E 8.57 1.38 5 8.97 10.10 6.41 114 Wfs Ultisols Lava Dead >2mm<5mm

II1E 3.77 2.18 6 3.00 7.71 1.97 114 Wfs Ultisols Lava >5mm

II1E 4.18 1.54 29 4.15 7.13 1.68 131 Wfs Mollisols Plutonic rocks <1cm

II1E 5.51 2.14 19 6.30 9.11 2.12 131 Wfs Mollisols Plutonic rocks >1cm<2cm

II1E 3.54 1.23 5 2.92 5.00 2.38 131 Wfs Mollisols Plutonic rocks >2cm

II3 3.77 1.77 13 4.35 6.51 1.20 142 Wflm Inceptisols Tuffac. sandstone

II3 2.00 0.76 17 1.76 4.16 1.03 142 Wflm Inceptisols Tuffac. sandstone

II6 9.30 4.64 17 9.13 21.89 1.63 142 Wflm Inceptisols Tuffac. sandstone

II8A 4.43 0.81 16 4.74 5.32 2.54 111 Rfs Inceptisols Tuffac. sandstone

II8A 7.07 1.47 12 6.90 10.68 4.65 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8A 9.02 2.96 32 8.13 16.59 4.25 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8A 10.60 0.61 11 10.61 11.92 9.80 124 Dacroides-Adjuntas

II8A 11.76 1.77 17 11.31 14.88 9.27 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8A 14.31 2.56 9 14.26 18.19 10.48 130 Mfs Ultisols Tuffac. sandstone

II8A 8.42 1.78 35 8.08 11.21 4.43 132 Wfs Mollisols Plutonic rocks

II8A 5.64 2.17 69 5.32 14.83 1.70 136 Wfs Ultisols Tuffac. sandstone

II8A 11.27 3.91 66 10.59 22.53 6.00 140 Swf Ultisols Tuffac. sandstone Cubuy

II8A 38.37 10.42 6 36.89 56.29 26.28 140 Sdf Mollisols Alluvial deposits Guanica

II8A 8.18 3.03 144 7.24 17.90 3.53 140 Lmrf Ultisols Tuffac. sandstone Bisley

II8A 6.45 4.02 75 5.12 20.31 1.18 140 Swf Ultisols Tuffac. sandstone Palm Forest
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

263

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8A 4.52 1.67 20 4.17 8.74 2.51 140 Swf Ultisols Tuffac. sandstone Pico del Este

II8A 6.70 3.09 20 5.47 16.80 3.75 142 Wflm Inceptisols Tuffac. sandstone Dicotiledonea

II8A 3.57 1.64 21 3.03 8.73 1.79 142 Wflm Inceptisols Tuffac. sandstone

II8A 12.14 2.91 135 11.61 21.97 5.22 156 Swf Ultisols Tuffac. sandstone

II8B 3.60 1.72 12 3.16 7.47 1.97 111 Rfs Inceptisols Tuffac. sandstone

II8B 9.83 2.23 13 10.15 12.53 5.86 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8B 21.75 3.64 8 23.33 25.56 15.17 124 Dacroides

II8B 7.89 2.54 488 8.27 12.84 1.37 132 Wfs Mollisols Plutonic rocks

II8B 6.31 3.19 51 5.88 15.20 1.18 136 Wfs Ultisols Tuffac. sandstone

II8B 11.66 4.52 65 11.92 27.26 3.65 140 Swf Ultisols Tuffac. sandstone Cubuy

II8B 26.99 8.10 11 25.01 28.84 15.47 140 Sdf Mollisols Alluvial deposits Guanica

II8B 5.58 2.05 139 5.33 13.31 0.60 140 Lmrf Ultisols Tuffac. sandstone Bisley

II8B 6.206 4.68 66 4.912 30.07 1.69 140 Wflm Inceptisols Tuffac. sandstone Palm Forest

II8B 4.47 1.43 17 4.15 6.58 1.78 140 Rfs Inceptisols Tuffac. sandstone Pico del Este

II8B 6.05 3.57 31 5.28 17.63 1.62 142 Wflm Inceptisols Tuffac. sandstone

II8B 9.95 3.40 126 9.13 23.79 3.56 156 Swf Ultisols Tuffac. sandstone <2cm

II8B 8.06 4.10 23 8.46 15.98 2.15 156 Swf Ultisols Tuffac. sandstone >2cm

II8C 4.25 2.70 16 3.50 10.39 1.44 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8C 5.94 1.89 7 5.59 10.00 4.58 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8C 3.85 1.60 8 3.92 5.99 2.08 130 Mfs Ultisols Tuffac. sandstone

II8C 6.13 3.40 21 5.70 13.90 1.26 132 Wfs Mollisols Plutonic rocks

II8C 4.75 2.53 2 4.75 6.54 2.96 142 Wflm Inceptisols Tuffac. sandstone

II8C 6.14 3.95 86 5.58 32.23 1.04 156 Swf Ultisols Tuffac. sandstone

II8D 3.66 0.89 16 3.72 4.82 1.64 111 Rfs Inceptisols Tuffac. sandstone

II8D 11.05 1.82 9 11.72 12.98 7.23 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8D 14.78 1.67 8 15.14 16.94 12.21 130 Mfs Ultisols Tuffac. sandstone

II8D 6.52 1.94 27 6.70 9.55 1.68 132 Wfs Mollisols Plutonic rocks

II8D 10.56 3.95 62 10.22 21.04 3.97 140 Swf Ultisols Tuffac. sandstone Cubuy

II8D 72.05 10.33 7 75.08 84.78 54.27 140 Sdf Mollisols Alluvial deposits Guanica
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

264

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8D 6.83 2.89 138 6.09 20.24 1.26 140 Lmrf Ultisols Tuffac. sandstone Bisley

II8D 6.33 2.99 67 5.86 15.93 2.77 140 Wflm Inceptisols Tuffac. sandstone Palm Forest

II8D 2.52 0.63 9 2.32 3.55 1.71 140 Rfs Inceptisols Tuffac. sandstone Pico del Este

II8D 7.51 4.55 8 7.33 14.48 1.98 142 Wflm Inceptisols Tuffac. sandstone Fern

II8D 3.81 2.33 42 3.30 9.36 0.50 142 Wflm Inceptisols Tuffac. sandstone

II8D 10.04 2.89 124 10.17 21.48 4.01 156 Swf Ultisols Tuffac. sandstone

II9A 8.21 1.75 626 8.11 15.75 0.07 106 Lmrf Ultisols Tuffac. sandstone

II9A 5.99 0.81 12 5.68 8.06 5.12 124

II9A 10.91 3.52 10 10.50 17.62 6.08 124

II9A 16.35 3.17 138 16.31 26.47 3.06 130 Mfs Ultisols Tuffac. sandstone

II9A 10.09 0.48 3 10.04 10.59 9.64 153 Swf Ultisols Tuffac. sandstone

II9A 8.29 3.50 14 7.56 15.66 4.38 154 Wfs Ultisols Tuffac. sandstone

II9A 11.17 2.97 128 10.19 21.13 5.98 156 Swf Ultisols Tuffac. sandstone

II9B 5.97 2.54 462 5.61 20.16 1.17 106 Lmrf Ultisols Tuffac. sandstone

II9B 7.79 1.54 11 7.67 10.32 5.79 124

II9B 12.42 6.29 96 1.12 49.60 1.57 130 Mfs Ultisols Tuffac. sandstone

II9B 6.44 2.53 28 5.66 11.44 3.45 132 Wfs Mollisols Plutonic rocks

II9B 11.88 2.82 14 11.80 16.31 7.92 154 Wfs Ultisols Tuffac. sandstone

II9B 9.80 2.40 58 9.25 17.45 3.84 156 Swf Ultisols Tuffac. sandstone

II9C 4.33 2.68 63 3.75 24.05 0.42 106 Lmrf Ultisols Tuffac. sandstone

II9C 4.20 1.72 15 4.87 6.91 0.76 124

II9C 8.73 4.29 129 8.06 19.67 1.27 130 Mfs Ultisols Tuffac. sandstone

II9C 4.22 1.84 21 3.49 8.25 1.42 132 Wfs Mollisols Plutonic rocks

II9C 5.50 2.41 49 5.13 10.84 0.64 156 Swf Ultisols Tuffac. sandstone

II9D 8.47 4.24 505 7.88 36.74 1.32 106 Lmrf Ultisols Tuffac. sandstone

II9D 4.43 1.22 5 4.03 6.24 0.76 124 Dacroides 

II9D 15.87 3.62 58 15.84 25.62 7.82 130 Mfs Ultisols Tuffac. sandstone

II9D 9.08 2.36 15 9.62 11.95 4.33 132 Wfs Mollisols Plutonic rocks

II9D 7.49 4.68 13 5.14 16.27 1.82 154 Wfs Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

265

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II9D 9.67 2.80 49 9.75 19.89 2.87 156 Swf Ultisols Tuffac. sandstone

Carbon (%) II1A 44 6 55 46 51 16 112 Lmrf Ultisols Tuffac. sandstone Bisley

II1A 44 7 49 47 52 23 112 Lmrf Ultisols Tuffac. sandstone Puente Roto

II8D 35 4 10 35 40 29 124 Lmrf Ultisols Tuffac. sandstone Bisley

II9A 53 1 22 53 55 52 132 Wfs Mollisols Plutonic rocks

II1E 49 2 16 49 53 47 114 Wfs Ultisols Lava Lives <2mm

II1E 46 3 18 45 51 40 114 Wfs Ultisols Lava Dead <2mm

II1E 51 3 6 52 54 45 114 Wfs Ultisols Lava Live >2mm<5mm

II1E 51 2 5 51 53 49 114 Wfs Ultisols Lava Dead >2mm<5mm

II1E 52 2 6 52 53 49 114 Wfs Ultisols Lava >5mm

II1E 53 1 29 53 55 50 131 Wfs Mollisols Plutonic rocks <1cm

II1E 53 2 19 53 56 51 131 Wfs Mollisols Plutonic rocks >1cm<2cm

II1E 55 2 5 56 57 53 131 Wfs Mollisols Plutonic rocks >2cm

II3 46 4 13 46 51 38 142 Wflm Inceptisols Tuffac. sandstone

II3 42 3 13 40 46 38 142 Wflm Inceptisols Tuffac. sandstone

II6 47 3 15 47 53 43 142 Wflm Inceptisols Tuffac. sandstone

II8A 40 1 12 40 42 39 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8A 38 3 32 39 42 31 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8A 40 1 11 41 41 40 124 Dacroides-Adjuntas

II8A 40 1 17 40 42 37 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8A 50 5 33 52 55 33 132 Wfs Mollisols Plutonic rocks

II8A 45 5 69 45 55 24 136 Wfs Ultisols Tuffac. sandstone

II8A 51 5 20 51 58 42 142 Wflm Inceptisols Tuffac. sandstone Dicotiledonea

II8A 48 4 20 48 53 40 142 Wflm Inceptisols Tuffac. sandstone

II8B 43 1 13 43 44 40 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8B 42 1 8 43 44 40 124 Dacroides

II8B 52 2 49 52 55 49 132 Wfs Mollisols Plutonic rocks

II8B 51 3 50 50 55 43 136 Wfs Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

266

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8B 53 2 31 53 56 48 142 Wflm Inceptisols Tuffac. sandstone

II8C 44 2 13 44 46 39 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8C 45 0 7 44 45 44 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8C 50 6 21 51 55 29 132 Wfs Mollisols Plutonic rocks

II8C 55 1 2 55 56 54 142 Wflm Inceptisols Tuffac. sandstone

II8D 32 3 9 33 36 28 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8D 44 10 27 43 66 27 132 Wfs Mollisols Plutonic rocks

II8D 46 4 7 45 52 42 142 Wflm Inceptisols Tuffac. sandstone Fern

II8D 31 14 42 36 47 3 142 Wflm Inceptisols Tuffac. sandstone

II9A 48 1 130 48 51 40 106 Lmrf Ultisols Tuffac. sandstone

II9A 49 1 87 49 55 46 108 Wf-LM Ultisols Tuffac. sandstone Mameyes

II9A 49 1 95 49 51 45 108 Wf-LM Ultisols Tuffac. sandstone Sabana

II9A 51 1 20 51 53 49 113 Wfs Ultisols Lava

II9A 41 1 12 41 42 41 124

II9A 42 1 10 42 43 41 124

II9A 51 0 3 51 51 51 153 Swf Ultisols Tuffac. sandstone

II9B 50 2 94 49 56 46 106 Lmrf Ultisols Tuffac. sandstone

II9B 49 2 63 49 56 42 108 Wf-LM Ultisols Tuffac. sandstone Mameyes

II9B 49 2 70 49 53 45 108 Wf-LM Ultisols Tuffac. sandstone Mameyes

II9B 52 2 18 52 56 47 113 Wfs Ultisols Lava

II9B 44 0 11 44 45 44 124

II9B 52 1 30 52 56 50 132 Wfs Mollisols Plutonic rocks

II9C 49 2 100 49 52 44 106 Lmrf Ultisols Tuffac. sandstone

II9C 51 2 45 50 56 47 108 Wf-LM Ultisols Tuffac. sandstone Mameyes

II9C 50 2 49 50 54 42 108 Wf-LM Ultisols Tuffac. sandstone Sabana

II9C 53 1 19 53 55 51 113 Wfs Ultisols Lava

II9C 44 2 15 43 46 39 124

II9C 54 2 21 54 58 51 132 Wfs Mollisols Plutonic rocks

II9D 44 2 85 44 54 41 106 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

267

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II9D 48 2 38 49 52 44 108 Wf-LM Ultisols Tuffac. sandstone Mameyes

II9D 48 2 39 48 51 44 108 Wf-LM Ultisols Tuffac. sandstone Sabana

II9D 48 2 12 48 51 43 113 Wfs Ultisols Lava

II9D 36 3 5 35 41 2 124 Dacroides 

II9D 49 1 17 50 52 46 132 Wfs Mollisols Plutonic rocks

Iron (mg/g) II1A 5.97 7.72 55 2.88 31.92 0.14 112 Lmrf Ultisols Tuffac. sandstone Bisley

II1A 5.05 6.44 51 2.46 27.70 0.10 112 Lmrf Ultisols Tuffac. sandstone Puente Roto

II8B 0.99 0.84 8 0.61 2.47 0.38 130 Mfs Ultisols Tuffac. sandstone <=2cm

II8B 0.51 0.37 9 0.48 1.28 0.04 130 Mfs Ultisols Tuffac. sandstone >2cm

II8D 14.65 7.04 10 15.58 22.84 5.87 124 Lmrf Ultisols Tuffac. sandstone Bisley

II9A 0.13 0.04 22 0.12 0.26 0.09 132 Wfs Mollisols Plutonic rocks

II1E 8.57 3.42 16 8.10 15.52 3.08 114 Wfs Ultisols Lava Lives <2mm

II1E 17.62 7.80 16 17.56 32.12 4.68 114 Wfs Ultisols Lava Dead <2mm

II1E 7.22 5.92 6 5.48 18.55 1.46 114 Wfs Ultisols Lava Live >2mm<5mm

II1E 6.35 3.64 5 5.66 11.27 2.50 114 Wfs Ultisols Lava Dead >2mm<5mm

II1E 3.62 1.81 6 3.41 6.60 1.78 114 Wfs Ultisols Lava >5mm

II1E 1.27 0.69 29 1.02 2.66 0.38 131 Wfs Mollisols Plutonic rocks <1cm

II1E 1.04 0.86 19 0.87 2.78 0.19 131 Wfs Mollisols Plutonic rocks >1cm<2cm

II1E 0.99 0.68 5 0.64 1.97 0.36 131 Wfs Mollisols Plutonic rocks >2cm

II3 2.33 2.10 13 1.56 7.68 0.52 142 Wflm Inceptisols Tuffac. sandstone

II3 2.73 1.63 17 2.16 6.34 0.61 142 Wflm Inceptisols Tuffac. sandstone

II6 1.52 1.38 17 1.29 6.04 0.16 142 Wflm Inceptisols Tuffac. sandstone

II8A 0.90 0.81 16 0.50 2.62 0.20 111 Rfs Inceptisols Tuffac. sandstone

II8A 1.73 1.51 12 1.34 4.66 0.25 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8A 5.81 5.14 32 4.13 17.37 0.38 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8A 1.12 0.70 11 0.98 2.90 0.32 124 Dacroides-Adjuntas

II8A 2.37 2.25 17 1.80 7.06 0.15 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8A 6.01 6.30 9 4.50 19.01 0.50 130 Mfs Ultisols Tuffac. sandstone

II8A 3.13 3.11 35 1.23 15.45 0.23 132 Wfs Mollisols Plutonic rocks
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

268

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8A 5.82 4.85 69 4.84 26.77 0.74 136 Wfs Ultisols Tuffac. sandstone

II8A 2.89 1.88 66 2.54 8.94 0.15 140 Swf Ultisols Tuffac. sandstone Cubuy

II8A 0.60 0.20 6 0.62 0.80 0.32 140 Sdf Mollisols Alluvial deposits Guanica

II8A 2.72 3.93 144 1.56 30.68 0.22 140 Lmrf Ultisols Tuffac. sandstone Bisley

II8A 1.05 1.22 75 0.65 7.13 0.12 140 Wflm Inceptisols Tuffac. sandstone Palm Forest

II8A 1.52 1.46 20 1.34 5.95 0.17 140 Rfs Inceptisols Tuffac. sandstone Pico del Este

II8A 3.66 3.14 20 3.20 14.50 0.31 142 Wflm Inceptisols Tuffac. sandstone Dicotiledonea

II8A 2.86 2.11 21 2.28 6.93 0.30 142 Wflm Inceptisols Tuffac. sandstone

II8B 0.45 0.41 12 0.31 1.50 0.07 111 Rfs Inceptisols Tuffac. sandstone

II8B 2.11 1.77 13 1.85 6.98 0.57 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8B 1.31 0.90 8 0.93 2.86 0.20 124 Dacroides

II8B 0.43 0.52 48 0.22 2.24 0.07 132 Wfs Mollisols Plutonic rocks

II8B 1.23 1.46 51 0.66 7.14 0.09 136 Wfs Ultisols Tuffac. sandstone

II8B 0.94 0.79 65 0.68 3.66 0.08 140 Swf Ultisols Tuffac. sandstone Cubuy

II8B 0.29 0.17 11 0.25 0.52 0.08 140 Sdf Mollisols Alluvial deposits Guanica

II8B 1.17 2.08 139 0.53 18.07 0.05 140 Lmrf Ultisols Tuffac. sandstone Bisley

II8B 0.77 0.88 66 0.57 5.20 0.09 140 Wflm Inceptisols Tuffac. sandstone Palm Forest

II8B 1.33 1.72 17 0.62 6.90 0.11 140 Rfs Inceptisols Tuffac. sandstone Pico del Este

II8B 1.80 1.37 31 1.50 5.73 0.11 142 Wflm Inceptisols Tuffac. sandstone

II8C 4.57 4.04 16 2.72 15.41 1.06 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8C 1.27 0.43 7 1.46 1.68 0.77 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8C 2.80 2.50 8 2.20 8.06 0.36 130 Mfs Ultisols Tuffac. sandstone

II8C 2.24 3.45 21 1.01 14.78 0.11 132 Wfs Mollisols Plutonic rocks

II8C 3.17 1.66 2 3.17 4.41 1.92 142 Wflm Inceptisols Tuffac. sandstone

II8D 1.92 1.44 16 1.69 5.25 0.27 111 Rfs Inceptisols Tuffac. sandstone

II8D 18.67 5.07 9 20.15 26.84 12.63 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8D 14.67 6.26 8 14.85 24.05 4.82 130 Mfs Ultisols Tuffac. sandstone

II8D 5.95 5.54 27 5.44 16.67 0.01 132 Wfs Mollisols Plutonic rocks

II8D 9.23 4.10 62 9.61 16.07 0.23 140 Swf Ultisols Tuffac. sandstone Cubuy
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

269

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8D 6.10 1.70 7 6.26 8.12 3.24 140 Sdf Mollisols Alluvial deposits Guanica

II8D 4.61 6.19 138 2.68 46.97 0.15 140 Lmrf Ultisols Tuffac. sandstone Bisley

II8D 4.70 5.07 67 2.40 21.72 0.18 140 Wflm Inceptisols Tuffac. sandstone Palm Forest

II8D 14.36 18.33 9 8.01 50.59 1.43 140 Rfs Inceptisols Tuffac. sandstone Pico del Este

II8D 4.09 2.11 8 3.39 6.80 1.45 142 Wflm Inceptisols Tuffac. sandstone Fern

II8D 11.11 7.34 42 9.91 30.39 1.35 142 Wflm Inceptisols Tuffac. sandstone

II9A 1.31 2.74 583 0.40 29.61 0.01 106 Lmrf Ultisols Tuffac. sandstone

II9A 0.49 0.31 12 0.46 1.12 0.12 124

II9A 0.51 0.44 10 0.40 1.71 0.18 124

II9A 0.35 0.52 138 0.24 4.33 0.09 130 Mfs Ultisols Tuffac. sandstone

II9A 0.86 0.11 3 0.88 0.95 0.74 153 Swf Ultisols Tuffac. sandstone

II9A 0.28 0.20 14 0.31 0.60 0.03 154 Wfs Ultisols Tuffac. sandstone

II9B 0.38 1.27 420 0.13 18.75 0.01 106 Lmrf Ultisols Tuffac. sandstone

II9B 0.47 0.28 11 0.34 1.06 0.17 124

II9B 0.55 2.75 96 0.12 19.35 0.01 130 Mfs Ultisols Tuffac. sandstone

II9B 0.08 0.03 29 0.07 0.17 0.03 132 Wfs Mollisols Plutonic rocks

II9B 0.20 0.38 14 0.05 1.12 0.02 154 Wfs Ultisols Tuffac. sandstone

II9C 1.01 1.82 460 0.44 22.67 0.01 106 Lmrf Ultisols Tuffac. sandstone

II9C 3.32 2.39 15 3.56 7.92 0.24 124

II9C 0.37 0.78 129 0.20 7.48 0.01 130 Mfs Ultisols Tuffac. sandstone

II9C 0.33 0.24 21 0.30 1.13 0.04 132 Wfs Mollisols Plutonic rocks

II9D 1.23 3.72 464 0.37 43.77 0.01 106 Lmrf Ultisols Tuffac. sandstone

II9D 11.59 4.23 5 14.00 15.43 0.24 124 Dacroides 

II9D 9.43 8.96 58 6.76 27.44 0.02 130 Mfs Ultisols Tuffac. sandstone

II9D 0.53 0.57 17 0.39 2.22 0.10 132 Wfs Mollisols Plutonic rocks

II9D 0.27 0.21 13 0.19 0.60 0.05 154 Wfs Ultisols Tuffac. sandstone

Magnesium (mg/g) II8B 1.94 0.58 8 2.01 2.92 1.14 130 Mfs Ultisols Tuffac. sandstone <=2cm

II8B 2.21 1.21 9 2.47 3.49 0.47 130 Mfs Ultisols Tuffac. sandstone >2cm

II9A 2.29 0.33 22 2.26 3.01 1.77 132 Wfs Mollisols Plutonic rocks
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

270

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1E 1.95 0.54 16 1.81 3.00 1.29 114 Wfs Ultisols Lava Lives <2mm

II1E 17.71 0.03 16 1.62 2.28 1.27 114 Wfs Ultisols Lava Dead <2mm

II1E 1.49 0.31 6 1.39 2.05 1.22 114 Wfs Ultisols Lava Live >2mm<5mm

II1E 1.38 0.28 5 1.20 1.80 1.18 114 Wfs Ultisols Lava Dead >2mm<5mm

II1E 1.50 0.77 6 1.19 2.78 0.85 114 Wfs Ultisols Lava >5mm

II1E 1.15 0.31 29 1.15 2.00 0.67 131 Wfs Mollisols Plutonic rocks <1cm

II1E 1.24 0.39 19 1.19 1.99 0.65 131 Wfs Mollisols Plutonic rocks >1cm<2cm

II1E 0.96 0.36 5 0.77 1.38 0.65 131 Wfs Mollisols Plutonic rocks >2cm

II3 3.05 0.94 13 2.88 4.31 1.42 142 Wflm Inceptisols Tuffac. sandstone

II3 2.84 0.68 17 2.91 4.00 1.70 142 Wflm Inceptisols Tuffac. sandstone

II6 3.97 1.64 17 4.36 6.63 1.03 142 Wflm Inceptisols Tuffac. sandstone

II8A 3.68 0.86 9 3.70 5.25 2.38 130 Mfs Ultisols Tuffac. sandstone

II8A 2.36 0.49 35 2.41 3.51 1.27 132 Wfs Mollisols Plutonic rocks

II8A 1.72 0.67 69 1.59 3.33 0.63 136 Wfs Ultisols Tuffac. sandstone

II8A 2.62 0.48 66 2.48 3.74 1.81 140 Swf Ultisols Tuffac. sandstone Cubuy

II8A 2.03 0.61 6 2.06 2.79 1.07 140 Sdf Mollisols Alluvial deposits Guanica

II8A 1.91 0.56 144 1.93 3.32 0.67 140 Lmrf Ultisols Tuffac. sandstone Bisley

II8A 1.76 0.61 75 1.67 3.09 0.46 140 Wflm Inceptisols Tuffac. sandstone Palm Forest

II8A 1.69 0.68 20 1.41 4.02 1.18 140 Rfs Inceptisols Tuffac. sandstone Pico del Este

II8A 1.78 0.58 20 1.60 3.21 1.10 142 Wflm Inceptisols Tuffac. sandstone Dicotiledonea

II8A 1.32 0.61 21 1.20 3.43 0.50 142 Wflm Inceptisols Tuffac. sandstone

II8A 3.65 0.81 135 3.54 6.13 1.61 156 Swf Ultisols Tuffac. sandstone

II8B 1.70 0.62 48 1.68 3.11 0.37 132 Wfs Mollisols Plutonic rocks

II8B 1.12 0.47 51 1.00 3.05 0.46 136 Wfs Ultisols Tuffac. sandstone

II8B 2.00 0.46 65 1.96 3.35 1.21 140 Swf Ultisols Tuffac. sandstone Cubuy

II8B 0.87 0.36 11 0.85 1.85 0.53 140 Sdf Mollisols Alluvial deposits Guanica

II8B 1.16 0.68 139 1.10 6.21 0.23 140 Lmrf Ultisols Tuffac. sandstone Bisley

II8B 1.17 0.59 66 1.00 2.88 0.33 140 Wflm Inceptisols Tuffac. sandstone Palm Forest

II8B 0.76 0.14 17 0.75 1.03 0.51 140 Rfs Inceptisols Tuffac. sandstone Pico del Este
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

271

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8B 0.96 0.38 31 0.90 2.55 0.40 142 Wflm Inceptisols Tuffac. sandstone

II8B 2.41 0.89 126 2.25 7.82 0.33 156 Swf Ultisols Tuffac. sandstone <2cm

II8B 1.53 0.68 23 1.39 3.00 0.43 156 Swf Ultisols Tuffac. sandstone >2cm

II8C 1.31 0.48 8 1.25 2.03 0.74 130 Mfs Ultisols Tuffac. sandstone

II8C 2.14 1.43 21 1.92 7.81 1.22 132 Wfs Mollisols Plutonic rocks

II8C 1.58 0.44 2 1.58 1.88 1.27 142 Wflm Inceptisols Tuffac. sandstone

II8C 2.71 1.63 86 2.35 10.65 0.90 156 Swf Ultisols Tuffac. sandstone

II8D 4.22 0.93 8 4.17 5.38 2.76 130 Mfs Ultisols Tuffac. sandstone

II8D 1.65 0.41 27 1.67 2.39 0.94 132 Wfs Mollisols Plutonic rocks

II8D 2.86 0.89 62 2.63 4.48 1.43 140 Swf Ultisols Tuffac. sandstone Cubuy

II8D 2.59 0.37 7 2.72 2.96 1.83 140 Sdf Mollisols Alluvial deposits Guanica

II8D 1.78 0.60 138 1.69 3.65 0.58 140 Lmrf Ultisols Tuffac. sandstone Bisley

II8D 2.10 0.84 67 1.88 4.54 1.08 140 Wflm Inceptisols Tuffac. sandstone Palm Forest

II8D 1.04 0.31 9 0.95 1.71 0.70 140 Rfs Inceptisols Tuffac. sandstone Pico del Este

II8D 3.31 1.57 8 3.72 5.44 1.33 142 Wflm Inceptisols Tuffac. sandstone Fern

II8D 1.90 0.65 42 1.80 3.72 0.63 142 Wflm Inceptisols Tuffac. sandstone

II8D 4.87 1.79 125 4.70 10.26 1.47 156 Swf Ultisols Tuffac. sandstone

II9A 2.23 0.57 626 21.14 5.32 0.02 106 Lmrf Ultisols Tuffac. sandstone

II9A 1.36 0.26 12 1.27 1.96 1.16 124

II9A 2.41 0.43 10 2.32 3.08 1.76 124

II9A 3.23 0.77 138 3.18 5.38 1.22 130 Mfs Ultisols Tuffac. sandstone

II9A 2.34 0.05 3 2.37 2.38 2.28 153 Swf Ultisols Tuffac. sandstone

II9A 2.62 0.92 13 2.35 4.05 1.52 154 Wfs Ultisols Tuffac. sandstone

II9A 3.44 0.55 128 3.39 5.41 2.25 156 Swf Ultisols Tuffac. sandstone

II9B 1.38 0.65 462 1.27 6.67 0.37 106 Lmrf Ultisols Tuffac. sandstone

II9B 1.50 0.26 11 1.57 1.81 0.98 124

II9B 2.22 0.95 96 2.10 6.57 0.65 130 Mfs Ultisols Tuffac. sandstone

II9B 1.72 0.70 29 1.44 4.63 0.90 132 Wfs Mollisols Plutonic rocks

II9B 2.49 0.45 14 2.41 3.32 1.92 154 Wfs Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

272

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II9B 1.91 0.54 58 1.92 3.22 0.79 156 Swf Ultisols Tuffac. sandstone

II9C 1.65 0.82 463 1.48 7.84 0.12 106 Lmrf Ultisols Tuffac. sandstone

II9C 1.60 0.51 15 1.79 2.30 0.68 124

II9C 0.09 0.16 129 0.07 1.81 0.01 130 Mfs Ultisols Tuffac. sandstone

II9C 1.92 0.53 21 1.79 3.00 0.96 132 Wfs Mollisols Plutonic rocks

II9C 2.18 0.64 49 2.27 4.02 1.01 156 Swf Ultisols Tuffac. sandstone

II9D 2.87 1.15 505 2.73 7.46 0.46 106 Lmrf Ultisols Tuffac. sandstone

II9D 1.25 0.16 5 1.30 1.39 0.51 124 Dacroides 

II9D 4.74 1.04 58 4.66 7.23 2.53 130 Mfs Ultisols Tuffac. sandstone

II9D 4.26 1.44 17 4.63 6.84 1.82 132 Wfs Mollisols Plutonic rocks

II9D 3.01 1.76 13 2.37 6.97 1.08 154 Wfs Ultisols Tuffac. sandstone

II9D 3.03 0.87 49 2.91 6.13 1.29 156 Swf Ultisols Tuffac. sandstone

II1A 1.93 0.61 55 1.88 3.80 0.84 112 Lmrf Ultisols Tuffac. sandstone Bisley

II1A 2.19 0.77 51 2.17 4.27 0.98 112 Lmrf Ultisols Tuffac. sandstone Puente Roto

II8D 1.42 0.15 10 1.43 1.62 1.08 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8A 1.74 0.13 16 1.50 1.99 1.48 111 Rfs Inceptisols Tuffac. sandstone

II8A 1.51 0.07 12 1.50 1.65 1.41 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8A 1.61 0.40 32 1.56 2.63 0.98 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8A 1.10 0.06 11 1.08 1.22 1.00 124 Dacroides-Adjuntas

II8A 1.22 0.12 17 1.21 1.46 1.01 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8B 0.93 0.30 12 0.97 1.32 0.53 111 Rfs Inceptisols Tuffac. sandstone

II8B 1.24 0.20 13 1.25 1.51 0.83 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8B 1.11 0.25 8 1.04 1.63 0.83 124 Dacroides

II8C 1.03 0.49 16 0.84 1.99 0.46 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8C 0.84 0.14 7 0.81 1.11 0.70 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8D 1.90 0.31 16 1.94 2.30 1.27 111 Rfs Inceptisols Tuffac. sandstone

II8D 1.48 0.20 9 1.49 1.83 1.23 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II9C 2.82 1.22 129 2.68 5.74 0.83 130 Mfs Ultisols Tuffac. sandstone

Manganese (mg/g) II1A 0.63 0.64 55 0.36 2.41 0.05 112 Lmrf Ultisols Tuffac. sandstone Bisley
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

273

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 0.38 0.34 51 0.22 1.29 0.06 112 Lmrf Ultisols Tuffac. sandstone Puente Roto

II8B 0.17 0.11 8 0.15 0.30 0.03 130 Mfs Ultisols Tuffac. sandstone <=2cm

II8B 0.14 0.21 9 0.08 0.67 0.01 130 Mfs Ultisols Tuffac. sandstone >2cm

II8D 0.37 0.08 10 0.37 0.48 0.25 124 Lmrf Ultisols Tuffac. sandstone Bisley

II9A 0.72 0.18 22 0.65 1.04 0.43 132 Wfs Mollisols Plutonic rocks

II1E 0.37 0.15 16 0.37 0.76 0.14 114 Wfs Ultisols Lava Lives <2mm

II1E 0.69 0.30 16 0.67 1.29 0.18 114 Wfs Ultisols Lava Dead <2mm

II1E 0.20 0.07 6 0.19 0.33 0.12 114 Wfs Ultisols Lava Live >2mm<5mm

II1E 0.56 0.38 5 0.82 0.86 0.13 114 Wfs Ultisols Lava Dead >2mm<5mm

II1E 0.35 0.38 6 0.14 1.05 0.13 114 Wfs Ultisols Lava >5mm

II1E 0.23 0.15 29 0.23 0.61 0.03 131 Wfs Mollisols Plutonic rocks <1cm

II1E 0.20 0.09 19 0.21 0.45 0.02 131 Wfs Mollisols Plutonic rocks >1cm<2cm

II1E 0.17 0.06 15 0.14 0.27 0.11 131 Wfs Mollisols Plutonic rocks >2cm

II3 0.22 0.14 13 0.23 0.44 0.06 142 Wflm Inceptisols Tuffac. sandstone

II3 0.55 0.16 17 0.54 0.91 0.24 142 Wflm Inceptisols Tuffac. sandstone

II6 0.47 0.28 17 0.35 1.20 0.19 142 Wflm Inceptisols Tuffac. sandstone

II8A 0.28 0.21 16 0.17 0.69 0.08 111 Rfs Inceptisols Tuffac. sandstone

II8A 0.47 0.08 12 0.48 0.60 0.28 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8A 0.41 0.08 32 0.41 0.58 0.24 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8A 0.95 0.14 11 0.97 1.17 0.78 124 Dacroides-Adjuntas

II8A 1.07 0.21 17 1.10 1.60 0.73 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8A 0.29 0.19 9 0.29 0.65 0.10 130 Mfs Ultisols Tuffac. sandstone

II8A 1.19 0.61 35 1.05 3.49 0.50 132 Wfs Mollisols Plutonic rocks

II8A 0.66 0.41 69 0.64 1.83 0.04 136 Wfs Ultisols Tuffac. sandstone

II8A 0.40 0.27 66 0.39 1.44 0.07 140 Swf Ultisols Tuffac. sandstone Cubuy

II8A 0.06 0.01 6 0.06 0.08 0.04 140 Sdf Mollisols Alluvial deposits Guanica

II8A 0.31 0.15 144 0.27 1.00 0.09 140 Lmrf Ultisols Tuffac. sandstone Bisley

II8A 0.59 0.36 75 0.55 1.42 0.06 140 Wflm Inceptisols Tuffac. sandstone Palm Forest

II8A 0.14 0.10 20 0.12 0.36 0.03 140 Rfs Inceptisols Tuffac. sandstone Pico del Este
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

274

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8A 0.50 0.25 20 0.44 1.04 0.19 142 Wflm Inceptisols Tuffac. sandstone Dicotiledonea

II8A 0.33 0.17 21 0.26 0.87 0.11 142 Wflm Inceptisols Tuffac. sandstone

II8B 0.18 0.30 12 0.07 0.87 0.03 111 Rfs Inceptisols Tuffac. sandstone

II8B 0.31 0.08 13 0.30 0.46 0.20 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8B 1.43 1.00 8 0.93 3.72 0.82 124 Dacroides

II8B 1.05 1.42 48 0.73 9.82 0.09 132 Wfs Mollisols Plutonic rocks

II8B 0.38 0.28 51 0.36 1.27 0.02 136 Wfs Ultisols Tuffac. sandstone

II8B 0.26 0.22 65 0.20 0.99 0.06 140 Swf Ultisols Tuffac. sandstone Cubuy

II8B 0.03 0.01 11 0.03 0.07 0.02 140 Sdf Mollisols Alluvial deposits Guanica

II8B 0.16 0.10 139 0.13 0.69 0.03 140 Lmrf Ultisols Tuffac. sandstone Bisley

II8B 0.29 0.19 66 0.28 0.92 0.03 140 Wflm Inceptisols Tuffac. sandstone Palm Forest

II8B 0.09 0.06 17 0.07 0.19 0.02 140 Rfs Inceptisols Tuffac. sandstone Pico del Este

II8B 0.29 0.18 31 0.24 0.94 0.04 142 Wflm Inceptisols Tuffac. sandstone

II8C 0.18 0.12 15 0.12 0.44 0.07 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8C 0.37 0.18 7 0.33 0.73 0.21 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8C 0.10 0.07 8 0.11 0.22 0.03 130 Mfs Ultisols Tuffac. sandstone

II8C 0.77 0.67 21 0.41 1.95 0.14 132 Wfs Mollisols Plutonic rocks

II8C 0.25 0.15 2 0.25 0.35 0.14 142 Wflm Inceptisols Tuffac. sandstone

II8D 0.16 0.09 16 0.14 0.41 0.08 111 Rfs Inceptisols Tuffac. sandstone

II8D 1.56 0.35 9 1.52 2.05 1.10 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8D 0.65 0.28 8 0.69 1.09 0.28 130 Mfs Ultisols Tuffac. sandstone

II8D 1.20 0.55 27 1.35 2.59 0.09 132 Wfs Mollisols Plutonic rocks

II8D 0.54 0.35 62 0.48 2.07 0.11 140 Swf Ultisols Tuffac. sandstone Cubuy

II8D 0.18 0.04 7 0.17 0.26 0.14 140 Sdf Mollisols Alluvial deposits Guanica

II8D 0.36 0.21 138 0.33 1.44 0.02 140 Lmrf Ultisols Tuffac. sandstone Bisley

II8D 0.76 0.44 67 0.72 1.63 0.21 140 Wflm Inceptisols Tuffac. sandstone Palm Forest

II8D 0.12 0.06 9 0.12 0.24 0.04 140 Rfs Inceptisols Tuffac. sandstone Pico del Este

II8D 0.37 0.17 8 0.39 0.58 0.17 142 Wflm Inceptisols Tuffac. sandstone Fern

II8D 0.53 0.25 42 0.45 1.37 0.22 142 Wflm Inceptisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

275

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II9A 0.40 0.16 583 0.39 2.20 0.01 106 Lmrf Ultisols Tuffac. sandstone

II9A 0.50 0.08 12 0.50 0.64 0.40 124

II9A 0.39 0.12 10 0.39 0.57 0.23 124

II9A 0.14 0.05 138 0.13 0.29 0.02 130 Mfs Ultisols Tuffac. sandstone

II9A 1.07 0.05 3 1.05 1.13 1.03 153 Swf Ultisols Tuffac. sandstone

II9A 0.40 0.27 14 0.36 0.79 0.05 154 Wfs Ultisols Tuffac. sandstone

II9B 0.25 0.20 420 0.19 1.74 0.02 106 Lmrf Ultisols Tuffac. sandstone

II9B 0.34 0.06 11 0.34 0.45 0.24 124

II9B 0.13 0.15 96 0.09 0.08 0.01 130 Mfs Ultisols Tuffac. sandstone

II9B 0.80 0.79 29 0.63 4.40 0.22 132 Wfs Mollisols Plutonic rocks

II9B 0.24 0.26 14 0.07 0.74 0.03 154 Wfs Ultisols Tuffac. sandstone

II9C 0.21 0.18 463 0.17 2.17 0.02 106 Lmrf Ultisols Tuffac. sandstone

II9C 0.22 0.07 15 0.24 0.31 0.08 124

II9C 0.46 0.22 21 0.46 0.94 0.09 132 Wfs Mollisols Plutonic rocks

II9D 0.46 0.27 64 0.40 1.93 0.02 106 Lmrf Ultisols Tuffac. sandstone

II9D 0.23 0.07 5 0.20 0.34 0.07 124 Dacroides 

II9D 0.54 0.40 58 0.45 1.38 0.04 130 Mfs Ultisols Tuffac. sandstone

II9D 0.56 0.21 17 0.48 1.14 0.34 132 Wfs Mollisols Plutonic rocks

II9D 0.17 0.07 13 0.15 0.26 0.04 154 Wfs Ultisols Tuffac. sandstone

Nitrogen (%) II1A 1.91 0.58 55 1.88 3.82 0.95 112 Lmrf Ultisols Tuffac. sandstone Bisley

II1A 1.75 0.52 49 1.81 2.63 0.61 112 Lmrf Ultisols Tuffac. sandstone Puente Roto

II8B 1.15 0.29 8 1.25 1.58 0.67 130 Mfs Ultisols Tuffac. sandstone <=2cm

II8B 0.86 0.44 9 0.78 1.59 0.44 130 Mfs Ultisols Tuffac. sandstone >2cm

II8D 1.29 0.18 10 1.33 1.49 0.94 124 Lmrf Ultisols Tuffac. sandstone Bisley

II9A 1.96 0.21 22 1.93 2.37 1.67 132 Wfs Mollisols Plutonic rocks

II1E 1.87 0.48 16 1.89 2.67 0.64 114 Wfs Ultisols Lava Lives <2mm

II1E 1.84 0.50 18 1.98 2.48 0.59 114 Wfs Ultisols Lava Dead <2mm

II1E 0.96 0.30 6 0.96 1.32 0.45 114 Wfs Ultisols Lava Live >2mm<5mm

II1E 1.31 0.48 5 1.26 1.89 0.72 114 Wfs Ultisols Lava Dead >2mm<5mm
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

276

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1E 0.74 0.20 6 0.72 1.03 0.50 114 Wfs Ultisols Lava >5mm

II1E 1.49 0.39 29 1.51 2.30 0.57 131 Wfs Mollisols Plutonic rocks <1cm

II1E 1.34 0.41 19 1.40 1.93 0.49 131 Wfs Mollisols Plutonic rocks >1cm<2cm

II1E 1.59 0.16 5 1.65 1.79 1.41 131 Wfs Mollisols Plutonic rocks >2cm

II3 1.54 0.79 13 1.35 4.08 0.96 142 Wflm Inceptisols Tuffac. sandstone

II3 2.14 0.72 13 2.10 3.24 1.02 142 Wflm Inceptisols Tuffac. sandstone

II6 1.50 0.61 15 1.52 2.87 0.54 142 Wflm Inceptisols Tuffac. sandstone

II8A 1.82 0.11 9 1.83 1.96 1.65 130 Mfs Ultisols Tuffac. sandstone

II8A 2.59 0.32 33 2.58 3.27 2.06 132 Wfs Mollisols Plutonic rocks

II8A 1.21 0.10 16 1.19 1.36 1.09 111 Rfs Inceptisols Tuffac. sandstone

II8A 0.85 0.06 12 0.84 0.97 0.76 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8A 1.08 0.14 32 1.06 1.38 0.79 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8A 0.80 0.06 11 0.80 0.91 0.70 124 Dacroides-Adjuntas

II8A 0.96 0.14 17 1.02 1.14 0.75 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8A 1.51 0.37 69 1.52 2.31 0.57 136 Wfs Ultisols Tuffac. sandstone

II8A 1.08 0.37 20 1.00 2.53 0.70 142 Wflm Inceptisols Tuffac. sandstone Dicotiledonea

II8A 0.81 0.21 20 0.82 1.24 0.34 142 Wflm Inceptisols Tuffac. sandstone

II8A 1.01 0.20 124 0.99 2.22 0.66 156 Swf Ultisols Tuffac. sandstone

II8B 1.45 0.42 49 1.38 2.38 0.57 132 Wfs Mollisols Plutonic rocks

II8B 0.68 0.11 12 0.70 0.86 0.46 111 Rfs Inceptisols Tuffac. sandstone

II8B 0.67 0.13 13 0.64 1.00 0.56 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8B 0.79 0.20 8 0.79 1.16 0.44 124 Dacroides

II8B 0.78 0.20 50 0.76 1.29 0.29 136 Wfs Ultisols Tuffac. sandstone

II8B 0.72 0.20 31 0.70 1.20 0.34 142 Wflm Inceptisols Tuffac. sandstone

II8B 0.64 0.17 118 0.62 1.20 0.34 156 Swf Ultisols Tuffac. sandstone <2cm

II8B 0.52 0.18 21 0.53 0.85 0.21 156 Swf Ultisols Tuffac. sandstone >2cm

II8C 1.19 0.14 8 1.21 1.34 1.01 130 Mfs Ultisols Tuffac. sandstone

II8C 2.36 0.60 21 2.24 3.64 1.33 132 Wfs Mollisols Plutonic rocks

II8C 1.01 0.39 12 0.93 1.73 0.58 124 Lmrf Ultisols Tuffac. sandstone Bisley
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

277

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8C 1.07 0.21 7 1.11 1.31 0.74 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8C 0.86 0.04 2 0.86 0.88 0.83 142 Wflm Inceptisols Tuffac. sandstone

II8C 0.97 0.41 84 0.93 2.71 0.28 156 Swf Ultisols Tuffac. sandstone

II8D 2.34 0.18 8 2.28 2.61 2.16 130 Mfs Ultisols Tuffac. sandstone

II8D 2.20 0.67 27 2.01 3.52 1.18 132 Wfs Mollisols Plutonic rocks

II8D 1.01 0.27 16 1.04 1.32 0.41 111 Rfs Inceptisols Tuffac. sandstone

II8D 1.35 0.18 9 1.25 1.64 1.20 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8D 1.08 0.26 7 1.11 1.54 0.74 142 Wflm Inceptisols Tuffac. sandstone Fern

II8D 0.91 0.41 42 0.98 1.77 0.10 142 Wflm Inceptisols Tuffac. sandstone

II8D 1.10 0.20 117 1.10 1.51 0.56 156 Swf Ultisols Tuffac. sandstone

II9A 1.54 0.36 14 1.45 2.17 1.12 154 Wfs Ultisols Tuffac. sandstone

II9A 1.74 0.28 138 1.70 2.27 0.69 130 Mfs Ultisols Tuffac. sandstone

II9A 1.52 0.25 175 1.52 2.45 0.50 106 Lmrf Ultisols Tuffac. sandstone

II9A 1.65 0.35 87 1.68 2.42 0.84 108 Wf-LM Ultisols Tuffac. sandstone Mameyes

II9A 1.09 0.29 95 1.02 1.91 0.67 108 Wf-LM Ultisols Tuffac. sandstone Sabana

II9A 1.27 0.16 20 1.23 1.67 0.94 113 Wfs Ultisols Lava

II9A 0.62 0.10 12 0.60 0.92 0.54 124

II9A 0.78 0.14 10 0.75 1.04 0.55 124

II9A 0.84 0.02 3 0.84 0.86 0.83 153 Swf Ultisols Tuffac. sandstone

II9A 0.73 0.12 128 0.71 1.11 0.54 156 Swf Ultisols Tuffac. sandstone

II9B 0.85 0.20 14 0.90 1.24 0.06 154 Wfs Ultisols Tuffac. sandstone

II9B 1.12 0.38 96 1.10 1.97 0.21 130 Mfs Ultisols Tuffac. sandstone

II9B 1.32 0.33 30 1.19 2.06 0.87 132 Wfs Mollisols Plutonic rocks

II9B 0.92 0.31 129 0.87 1.72 0.30 106 Lmrf Ultisols Tuffac. sandstone

II9B 1.01 0.34 63 0.96 1.94 0.16 108 Wf-LM Ultisols Tuffac. sandstone Mameyes

II9B 0.69 0.33 70 0.63 2.31 0.28 108 Wf-LM Ultisols Tuffac. sandstone Mameyes

II9B 0.87 0.31 18 0.92 1.45 0.30 113 Wfs Ultisols Lava

II9B 0.56 0.08 11 0.56 0.68 0.45 124

II9B 0.60 0.16 58 0.60 1.03 0.27 156 Swf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

278

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II9C 1.58 0.44 129 1.62 2.67 0.73 130 Mfs Ultisols Tuffac. sandstone

II9C 2.24 0.36 21 2.25 2.90 1.38 132 Wfs Mollisols Plutonic rocks

II9C 1.38 0.35 138 1.34 2.90 0.65 106 Lmrf Ultisols Tuffac. sandstone

II9C 1.56 0.43 45 1.56 2.40 0.55 108 Wf-LM Ultisols Tuffac. sandstone Mameyes

II9C 1.49 0.32 49 1.44 2.27 0.78 108 Wf-LM Ultisols Tuffac. sandstone Sabana

II9C 1.38 0.32 19 1.34 2.36 1.00 113 Wfs Ultisols Lava

II9C 1.23 0.36 15 1.24 1.65 0.45 124

II9C 1.32 0.38 49 1.32 2.01 0.68 156 Swf Ultisols Tuffac. sandstone

II9D 1.06 0.60 13 0.73 2.14 0.45 154 Wfs Ultisols Tuffac. sandstone

II9D 2.15 0.49 58 2.18 3.27 0.82 130 Mfs Ultisols Tuffac. sandstone

II9D 1.86 0.54 17 1.70 3.13 1.13 132 Wfs Mollisols Plutonic rocks

II9D 1.26 0.39 122 1.15 2.47 0.67 106 Lmrf Ultisols Tuffac. sandstone

II9D 1.45 0.32 38 1.45 2.32 0.55 108 Wf-LM Ultisols Tuffac. sandstone Mameyes

II9D 1.13 0.45 39 1.10 2.83 0.34 108 Wf-LM Ultisols Tuffac. sandstone Sabana

II9D 1.21 0.44 12 1.08 2.30 0.81 113 Wfs Ultisols Lava

II9D 0.90 0.12 5 0.88 1.08 0.36 124 Dacroides 

II9D 1.14 0.33 48 1.10 2.00 0.49 156 Swf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.74 0.31 55 0.71 1.81 0.28 112 Lmrf Ultisols Tuffac. sandstone Bisley

II1A 0.69 0.28 51 0.74 1.19 0.17 112 Lmrf Ultisols Tuffac. sandstone Puente Roto

II8B 0.42 0.13 8 0.44 0.55 0.26 130 Mfs Ultisols Tuffac. sandstone <=2cm

II8B 0.32 0.16 9 0.28 0.57 0.15 130 Mfs Ultisols Tuffac. sandstone >2cm

II8D 0.53 0.11 10 0.53 0.66 0.32 124 Lmrf Ultisols Tuffac. sandstone Bisley

II9A 0.50 0.18 22 0.46 0.92 0.30 132 Wfs Mollisols Plutonic rocks

II1E 0.63 0.15 16 0.62 0.84 0.24 114 Wfs Ultisols Lava Lives <2mm

II1E 0.49 0.14 16 0.53 0.63 0.17 114 Wfs Ultisols Lava Dead roots <2mm

II1E 0.34 0.13 6 0.37 0.46 0.11 114 Wfs Ultisols Lava Live >2mm<5mm

II1E 0.32 0.05 5 0.31 0.39 0.25 114 Wfs Ultisols Lava Dead >2mm<5mm

II1E 0.27 0.11 6 0.24 0.48 0.17 114 Wfs Ultisols Lava >5mm

II1E 0.25 0.13 29 0.21 0.56 0.11 131 Wfs Mollisols Plutonic rocks <1cm
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

279

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1E 0.20 0.08 19 0.18 0.41 0.11 131 Wfs Mollisols Plutonic rocks >1cm<2cm

II1E 0.17 0.03 5 0.17 0.20 0.13 131 Wfs Mollisols Plutonic rocks >2cm

II3 0.71 0.44 13 0.62 2.16 0.42 142 Wflm Inceptisols Tuffac. sandstone

II3 0.80 0.19 17 0.76 1.25 0.54 142 Wflm Inceptisols Tuffac. sandstone

II6 0.66 0.23 17 0.60 1.05 0.30 142 Wflm Inceptisols Tuffac. sandstone

II8A 0.44 0.04 16 0.43 0.52 0.38 111 Rfs Inceptisols Tuffac. sandstone

II8A 0.20 0.05 12 0.19 0.34 0.13 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8A 0.31 0.08 32 0.30 0.46 0.14 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8A 0.17 0.03 11 0.16 0.22 0.12 124 Dacroides-Adjuntas

II8A 0.25 0.08 17 0.26 0.37 0.12 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8A 0.63 0.15 9 0.66 0.88 0.44 130 Mfs Ultisols Tuffac. sandstone

II8A 0.69 0.18 35 0.66 1.04 0.42 132 Wfs Mollisols Plutonic rocks

II8A 0.51 0.23 69 0.54 1.06 0.12 136 Wflm Inceptisols Tuffac. sandstone

II8A 0.57 0.11 66 0.57 0.84 0.35 140 Rfs Inceptisols Tuffac. sandstone Cubuy

II8A 0.40 0.09 6 0.42 0.50 0.23 140 Sdf Mollisols Alluvial deposits Guanica

II8A 0.49 0.18 144 0.45 1.11 0.14 140 Lmrf Ultisols Tuffac. sandstone Bisley

II8A 0.52 0.20 75 0.51 1.14 0.11 140 Wflm Inceptisols Tuffac. sandstone Palm Forest

II8A 0.35 0.12 20 0.34 0.64 0.21 140 Rfs Inceptisols Tuffac. sandstone Pico del Este

II8A 0.30 0.20 20 0.23 1.03 0.16 142 Wflm Inceptisols Tuffac. sandstone Dicotiledonea

II8A 0.29 0.09 21 0.28 0.48 0.16 142 Wflm Inceptisols Tuffac. sandstone

II8A 0.42 0.09 135 0.42 0.80 0.20 156 Swf Ultisols Tuffac. sandstone

II8B 0.25 0.12 12 0.23 0.49 0.13 111 Rfs Inceptisols Tuffac. sandstone

II8B 0.20 0.04 13 0.20 0.28 0.13 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8B 0.23 0.08 8 0.23 0.39 0.14 124 Dacroides

II8B 0.40 0.17 48 0.38 0.99 0.13 132 Wfs Mollisols Plutonic rocks

II8B 0.23 0.12 51 0.24 0.47 0.05 136 Wfs Ultisols Tuffac. sandstone

II8B 0.32 0.08 65 0.31 0.56 0.17 140 Swf Ultisols Tuffac. sandstone Cubuy

II8B 0.25 0.11 11 0.23 0.54 0.14 140 Sdf Mollisols Alluvial deposits Guanica

II8B 0.22 0.12 139 0.20 1.13 0.09 140 Lmrf Ultisols Tuffac. sandstone Bisley
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

280

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8B 0.33 0.14 66 0.28 0.84 0.15 140 Wflm Inceptisols Tuffac. sandstone Palm Forest

II8B 0.15 0.05 17 0.13 0.26 0.06 140 Rfs Inceptisols Tuffac. sandstone Pico del Este

II8B 0.20 0.10 31 0.19 0.69 0.08 142 Wflm Inceptisols Tuffac. sandstone

II8B 0.29 0.12 126 0.28 1.27 0.12 156 Swf Ultisols Tuffac. sandstone <2cm

II8B 0.23 0.11 23 0.20 0.51 0.09 156 Swf Ultisols Tuffac. sandstone >2cm

II8C 0.55 0.29 16 0.51 1.35 0.22 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8C 0.54 0.12 7 0.57 0.67 0.34 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8C 0.63 0.12 8 0.60 0.80 0.50 130 Mfs Ultisols Tuffac. sandstone

II8C 0.78 0.24 21 0.76 1.27 0.27 132 Wfs Mollisols Plutonic rocks

II8C 0.44 0.13 2 0.44 0.53 0.35 142 Wflm Inceptisols Tuffac. sandstone

II8C 0.51 0.27 86 0.45 1.35 0.06 156 Swf Ultisols Tuffac. sandstone

II8D 0.38 0.15 16 0.40 0.57 0.11 111 Rfs Inceptisols Tuffac. sandstone

II8D 0.60 0.15 9 0.58 0.90 0.41 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8D 0.95 0.16 8 1.00 1.16 0.65 130 Mfs Ultisols Tuffac. sandstone

II8D 0.78 0.32 27 0.83 1.24 0.21 132 Wfs Mollisols Plutonic rocks

II8D 0.62 0.14 62 0.63 0.92 0.32 140 Swf Ultisols Tuffac. sandstone Cubuy

II8D 0.47 0.03 7 0.47 0.52 0.41 140 Sdf Mollisols Alluvial deposits Guanica

II8D 0.46 0.18 138 0.45 1.24 0.14 140 Lmrf Ultisols Tuffac. sandstone Bisley

II8D 0.66 0.14 67 0.64 0.97 0.39 140 Wflm Inceptisols Tuffac. sandstone Palm Forest

II8D 0.34 0.11 9 0.32 0.53 0.19 140 Rfs Inceptisols Tuffac. sandstone Pico del Este

II8D 0.50 0.15 8 0.54 0.72 0.28 142 Wflm Inceptisols Tuffac. sandstone Fern

II8D 0.40 0.14 42 0.38 0.89 0.19 142 Wflm Inceptisols Tuffac. sandstone

II8D 0.57 0.13 125 0.57 1.05 0.28 156 Swf Ultisols Tuffac. sandstone

II9A 0.43 0.16 626 0.41 1.80 0.01 106 Lmrf Ultisols Tuffac. sandstone

II9A 0.15 0.06 12 0.13 0.32 0.11 124

II9A 0.25 0.05 10 0.24 0.35 0.17 124

II9A 0.65 0.17 138 0.63 1.14 0.28 130 Mfs Ultisols Tuffac. sandstone

II9A 0.28 0.01 3 0.28 0.29 0.27 153 Swf Ultisols Tuffac. sandstone

II9A 0.82 0.17 14 0.77 1.09 0.62 154 Wfs Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

281

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II9A 0.33 0.08 128 0.32 0.56 0.19 156 Swf Ultisols Tuffac. sandstone

II9B 0.27 0.12 462 0.25 1.36 0.09 106 Lmrf Ultisols Tuffac. sandstone

II9B 0.25 0.05 11 0.25 0.35 0.18 124

II9B 0.39 0.23 96 0.33 1.86 0.08 130 Mfs Ultisols Tuffac. sandstone

II9B 0.33 0.13 29 0.30 0.68 0.14 132 Wfs Mollisols Plutonic rocks

II9B 0.75 0.16 14 0.72 1.12 0.59 154 Wfs Ultisols Tuffac. sandstone

II9B 0.31 0.09 58 0.30 0.52 0.15 156 Swf Ultisols Tuffac. sandstone

II9C 0.82 0.26 463 0.81 3.90 0.14 106 Lmrf Ultisols Tuffac. sandstone

II9C 0.86 0.26 15 0.84 1.35 0.44 124

II9C 1.09 0.26 129 1.07 1.80 0.29 130 Mfs Ultisols Tuffac. sandstone

II9C 1.04 0.26 21 1.01 1.72 0.65 132 Wfs Mollisols Plutonic rocks

II9C 1.23 0.43 49 1.09 2.43 0.61 156 Swf Ultisols Tuffac. sandstone

II9D 0.50 0.26 505 0.43 2.00 0.11 106 Lmrf Ultisols Tuffac. sandstone

II9D 0.54 0.13 5 0.56 0.68 0.26 124 Dacroides 

II9D 1.17 0.41 58 1.20 2.79 0.22 130 Mfs Ultisols Tuffac. sandstone

II9D 0.72 0.34 17 0.69 1.70 0.37 132 Wfs Mollisols Plutonic rocks

II9D 0.81 0.40 13 0.74 1.36 0.32 154 Wfs Ultisols Tuffac. sandstone

II9D 0.91 0.54 49 0.77 3.88 0.18 156 Swf Ultisols Tuffac. sandstone

Potassium (mg/g) II1A 2.33 1.32 55 2.01 6.13 0.80 112 Lmrf Ultisols Tuffac. sandstone Bisley

II1A 2.17 1.33 51 1.82 6.32 0.43 112 Lmrf Ultisols Tuffac. sandstone Puente Roto

II8B 2.01 0.32 8 2.00 2.49 1.48 130 Mfs Ultisols Tuffac. sandstone <=2cm

II8B 2.18 1.12 9 2.23 3.94 0.86 130 Mfs Ultisols Tuffac. sandstone >2cm

II8D 1.86 0.38 10 1.97 2.36 1.01 124 Lmrf Ultisols Tuffac. sandstone Bisley

II9A 4.94 1.16 22 4.99 7.58 3.01 132 Wfs Mollisols Plutonic rocks

II1E 2.52 1.44 16 2.04 6.52 1.18 114 Wfs Ultisols Lava Lives <2mm

II1E 1.96 0.49 16 1.95 2.91 1.11 114 Wfs Ultisols Lava Dead roots <2mm

II1E 3.08 2.41 6 2.12 6.16 0.70 114 Wfs Ultisols Lava Live >2mm<5mm

II1E 1.56 0.42 5 1.29 2.07 1.20 114 Wfs Ultisols Lava Dead >2mm<5mm

II1E 2.36 3.38 6 1.10 9.22 0.53 114 Wfs Ultisols Lava >5mm
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

282

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1E 1.93 0.97 29 1.52 4.57 0.70 131 Wfs Mollisols Plutonic rocks <1cm

II1E 2.71 1.95 19 1.94 9.11 1.34 131 Wfs Mollisols Plutonic rocks >1cm<2cm

II1E 2.71 0.90 5 2.94 3.55 1.23 131 Wfs Mollisols Plutonic rocks >2cm

II3 10.17 7.24 13 7.37 32.71 5.32 142 Wflm Inceptisols Tuffac. sandstone

II3 15.00 4.31 17 14.98 24.73 7.35 142 Wflm Inceptisols Tuffac. sandstone

II6 9.11 2.94 17 9.41 14.51 3.96 142 Wflm Inceptisols Tuffac. sandstone

II8A 4.35 0.36 16 4.39 4.94 3.57 111 Rfs Inceptisols Tuffac. sandstone

II8A 1.55 0.82 12 1.39 4.08 0.87 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8A 1.41 0.56 32 1.28 4.10 0.78 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8A 1.71 0.22 11 1.66 2.25 1.39 124 Dacroides-Adjuntas

II8A 1.76 0.39 17 1.61 2.74 1.45 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8A 2.81 0.72 9 2.87 4.02 1.61 130 Mfs Ultisols Tuffac. sandstone

II8A 2.54 0.61 35 2.50 5.15 1.66 132 Wfs Mollisols Plutonic rocks

II8A 1.27 0.37 69 1.18 2.63 0.73 136 Wfs Ultisols Tuffac. sandstone

II8A 2.80 1.17 66 2.50 7.07 1.62 140 Swf Ultisols Tuffac. sandstone Cubuy

II8A 4.09 0.91 6 4.30 5.01 2.44 140 Swf Ultisols Tuffac. sandstone Guanica

II8A 2.64 1.56 144 2.27 8.40 0.67 140 Swf Ultisols Tuffac. sandstone Bisley

II8A 1.93 1.73 75 1.51 10.62 0.27 140 Wflm Inceptisols Tuffac. sandstone Palm Forest

II8A 2.77 1.79 20 2.64 8.57 1.00 140 Rfs Inceptisols Tuffac. sandstone Pico del Este

II8A 1.83 1.00 20 1.54 5.50 0.94 142 Wflm Inceptisols Tuffac. sandstone Dicotiledonea

II8A 1.11 0.33 21 1.09 1.85 0.58 142 Wflm Inceptisols Tuffac. sandstone

II8A 1.64 0.55 135 1.50 4.80 0.76 156 Swf Ultisols Tuffac. sandstone

II8B 2.37 0.73 12 2.25 3.78 1.49 111 Rfs Inceptisols Tuffac. sandstone

II8B 1.35 0.66 13 1.29 0.75 0.66 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8B 1.52 0.35 8 1.47 1.91 0.99 124 Dacroides 

II8B 2.26 0.82 48 2.22 4.60 0.75 132 Wfs Mollisols Plutonic rocks

II8B 1.16 0.49 51 1.08 2.79 0.12 136 Wfs Ultisols Tuffac. sandstone

II8B 2.34 0.63 65 2.21 4.78 1.39 140 Swf Ultisols Tuffac. sandstone Cubuy

II8B 3.42 1.08 11 3.45 5.30 1.92 140 Sdf Mollisols Alluvial deposits Guanica
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

283

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8B 1.61 1.35 139 1.26 10.24 0.36 140 Lmrf Ultisols Tuffac. sandstone Bisley

II8B 1.22 0.57 66 1.03 0.77 0.37 140 Wflm Inceptisols Tuffac. sandstone Palm Forest

II8B 1.38 0.83 17 1.37 3.40 0.48 140 Rfs Inceptisols Tuffac. sandstone Pico del Este

II8B 1.22 0.81 31 1.08 5.28 0.49 142 Wflm Inceptisols Tuffac. sandstone

II8B 1.02 0.64 126 0.86 0.88 0.42 156 Swf Ultisols Tuffac. sandstone <2cm

II8B 0.73 0.49 23 0.68 2.61 0.22 156 Swf Ultisols Tuffac. sandstone >2cm

II8C 2.61 1.67 16 2.22 6.91 0.92 124 Lmrf Ultisols Tuffac. sandstone Bisley

II8C 1.88 0.82 7 1.56 3.08 1.10 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8C 1.54 0.34 8 1.51 2.07 1.14 130 Mfs Ultisols Tuffac. sandstone

II8C 4.16 2.98 21 3.05 12.20 1.35 132 Wfs Mollisols Plutonic rocks

II8C 1.57 0.79 2 1.57 2.13 1.01 142 Wflm Inceptisols Tuffac. sandstone

II8C 2.65 2.43 86 1.81 11.98 0.45 156 Swf Ultisols Tuffac. sandstone

II8D 4.22 1.33 16 4.50 5.72 1.54 111 Rfs Inceptisols Tuffac. sandstone

II8D 1.60 0.33 9 1.52 2.18 1.13 124 Wflm Ultisols Tuffac. sandstone Adjuntas

II8D 2.29 0.51 8 2.22 3.40 1.68 130 Mfs Ultisols Tuffac. sandstone

II8D 2.33 1.24 27 2.03 6.44 1.21 132 Wfs Mollisols Plutonic rocks

II8D 2.45 0.77 62 2.36 5.35 1.43 140 Swf Ultisols Tuffac. sandstone Cubuy

II8D 4.37 0.29 7 4.40 4.61 3.81 140 Sdf Mollisols Alluvial deposits Guanica

II8D 2.60 2.51 138 1.88 21.44 0.65 140 Lmrf Ultisols Tuffac. sandstone Bisley

II8D 1.55 0.43 67 1.45 2.64 0.79 140 Wflm Inceptisols Tuffac. sandstone Palm Forest

II8D 1.57 0.55 9 1.51 2.72 0.86 140 Rfs Inceptisols Tuffac. sandstone Pico del Este

II8D 2.46 0.99 8 2.52 3.99 1.13 142 Wflm Inceptisols Tuffac. sandstone Fern

II8D 1.63 0.30 42 1.60 4.15 0.85 142 Wflm Inceptisols Tuffac. sandstone

II8D 1.68 1.10 125 1.41 9.88 0.67 156 Swf Ultisols Tuffac. sandstone

II9A 3.98 1.44 626 3.71 9.54 0.03 106 Lmrf Ultisols Tuffac. sandstone

II9A 1.96 0.38 12 1.86 3.10 1.64 124

II9A 3.27 0.90 10 3.30 4.72 1.81 124

II9A 6.41 1.89 138 6.31 11.77 2.59 130 Mfs Ultisols Tuffac. sandstone

II9A 8.76 0.20 3 8.76 8.97 8.56 153 Swf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

284

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II9A 6.37 1.85 14 5.83 9.16 4.56 154 Wfs Ultisols Tuffac. sandstone

II9A 4.92 1.41 128 4.74 7.95 1.88 156 Swf Ultisols Tuffac. sandstone

II9B 1.62 1.02 462 1.29 6.51 0.35 106 Lmrf Ultisols Tuffac. sandstone

II9B 3.83 0.67 11 3.97 4.90 2.84 124

II9B 2.80 1.87 96 2.41 13.06 0.60 130 Mfs Ultisols Tuffac. sandstone

II9B 2.76 0.81 29 2.40 4.54 1.49 132 Wfs Mollisols Plutonic rocks

II9B 6.25 1.54 14 5.96 9.56 4.53 154 Wfs Ultisols Tuffac. sandstone

II9B 2.00 1.59 58 1.46 9.29 0.43 156 Swf Ultisols Tuffac. sandstone

II9C 4.91 2.39 463 4.52 29.95 1.91 106 Lmrf Ultisols Tuffac. sandstone

II9C 4.49 1.43 15 4.59 8.10 2.07 124

II9C 9.48 5.30 129 7.71 31.24 1.79 130 Mfs Ultisols Tuffac. sandstone

II9C 6.08 1.73 21 5.76 11.19 4.08 132 Wfs Mollisols Plutonic rocks

II9C 8.17 2.21 49 8.09 12.68 0.78 156 Swf Ultisols Tuffac. sandstone

II9D 4.43 2.47 505 3.85 26.38 0.55 106 Lmrf Ultisols Tuffac. sandstone

II9D 2.67 0.47 5 2.64 3.37 1.43 124

II9D 5.30 2.31 58 4.92 12.90 2.05 130 Mfs Ultisols Tuffac. sandstone

II9D 6.52 2.39 16 6.26 11.65 2.97 132 Wfs Mollisols Plutonic rocks

II9D 8.48 3.08 13 8.08 14.59 2.99 154 Wfs Ultisols Tuffac. sandstone

II9D 6.18 3.71 49 5.05 16.48 1.70 156 Swf Ultisols Tuffac. sandstone

Sodium (mg/g) II9A 2.03 0.07 3 2.00 2.11 1.97 153 Swf Ultisols Tuffac. sandstone

Sulfur (%) II9A 0.29 0.04 22 0.30 0.39 0.23 132 Wfs Mollisols Plutonic rocks

II1E 0.34 0.08 16 0.36 0.45 0.09 114 Wfs Ultisols Lava Lives <2mm

II1E 0.29 0.09 18 0.29 0.39 0.08 114 Wfs Ultisols Lava Dead <2mm

II1E 0.22 0.13 6 0.20 0.44 0.06 114 Wfs Ultisols Lava Live >2mm<5mm

II1E 0.19 0.06 5 0.21 0.25 0.10 114 Wfs Ultisols Lava Dead >2mm<5mm

II1E 0.14 0.04 6 0.12 0.20 0.10 114 Wfs Ultisols Lava >5mm

II1E 0.29 0.16 29 0.24 0.72 0.09 131 Wfs Mollisols Plutonic rocks <1cm

II1E 0.22 0.11 19 0.21 0.46 0.06 131 Wfs Mollisols Plutonic rocks >1cm<2cm

II1E 0.23 0.16 5 0.13 0.43 0.11 131 Wfs Mollisols Plutonic rocks >2cm
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

285

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II3 0.39 0.14 13 0.36 0.69 0.20 142 Wflm Inceptisols Tuffac. sandstone

II3 0.57 0.16 13 0.54 0.82 0.25 142 Wflm Inceptisols Tuffac. sandstone

II6 0.36 0.15 15 0.34 0.72 0.18 142 Wflm Inceptisols Tuffac. sandstone

II8A 0.45 0.13 33 0.42 0.72 0.26 132 Wfs Mollisols Plutonic rocks

II8A 0.25 0.08 69 0.25 0.50 0.10 136 Wfs Ultisols Tuffac. sandstone

II8A 0.19 0.04 20 0.18 0.29 0.15 142 Wflm Inceptisols Tuffac. sandstone Dicotiledonea

II8A 0.17 0.05 20 0.17 0.27 0.08 142 Wflm Inceptisols Tuffac. sandstone

II8B 0.24 0.10 49 0.23 0.49 0.05 132 Wfs Mollisols Plutonic rocks

II8B 0.15 0.04 50 0.14 0.25 0.04 136 Wfs Ultisols Tuffac. sandstone

II8B 0.10 0.03 31 0.12 0.20 0.07 142 Wflm Inceptisols Tuffac. sandstone

II8C 0.40 0.22 21 0.37 0.91 0.13 132 Wfs Mollisols Plutonic rocks

II8C 0.13 0.00 2 0.13 0.13 0.13 142 Wflm Inceptisols Tuffac. sandstone

II8D 0.38 0.15 27 0.33 0.71 0.19 132 Wfs Mollisols Plutonic rocks

II8D 0.20 0.08 7 0.20 0.30 0.09 142 Wflm Inceptisols Tuffac. sandstone Fern

II8D 0.15 0.07 42 0.17 0.28 0.02 142 Wflm Inceptisols Tuffac. sandstone

II9A 0.31 0.05 130 0.31 0.41 0.15 106 Lmrf Ultisols Tuffac. sandstone

II9B 0.16 0.04 94 0.16 0.25 0.05 106 Lmrf Ultisols Tuffac. sandstone

II9B 0.16 0.04 30 0.16 0.25 0.09 132 Wfs Mollisols Plutonic rocks

II9C 0.21 0.04 100 0.21 0.38 0.13 106 Lmrf Ultisols Tuffac. sandstone

II9C 0.24 0.04 21 0.24 0.34 0.16 132 Wfs Mollisols Plutonic rocks

II9D 0.17 0.05 85 0.17 0.30 0.07 106 Lmrf Ultisols Tuffac. sandstone

II9D 0.21 0.08 17 0.19 0.40 0.09 132 Wfs Mollisols Plutonic rocks

Securidaca virgata Aluminum (mg/g) II4 0.15 0.13 6 0.13 0.39 0.03 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II4 1.40 0.52 6 1.30 2.21 0.92 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II4 0.20 0.11 6 0.17 0.39 0.09 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II4 0.91 0.39 6 0.90 1.44 0.43 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II4 0.08 0.04 6 0.09 0.14 0.02 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II4 1.01 0.30 6 0.91 1.41 0.75 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

286

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Phosphorus (mg/g) II4 0.64 0.24 6 0.58 1.06 0.42 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II4 8.79 1.55 6 9.19 10.28 6.05 135 Lmrf Ultisols Tuffac. sandstone

Selaginella krugii Aluminum (mg/g) II3 4.62 2.15 6 5.57 6.85 1.31 135 Lmrf Ultisols Tuffac. sandstone

C/N II3 31 3 4 29 35 29 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II3 2.49 1.09 6 2.37 3.83 0.88 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II3 43 2 4 43 45 41 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II3 4.14 2.17 6 4.56 6.48 0.84 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II3 1.76 0.37 6 1.74 2.22 1.31 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II3 0.54 0.18 6 0.60 0.70 0.29 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II3 1.37 0.15 9 1.42 1.55 1.14 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II3 0.55 0.11 6 0.58 0.67 0.38 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II3 9.73 3.24 6 9.15 15.32 6.82 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II3 0.46 0.11 4 0.44 0.61 0.35 135 Lmrf Ultisols Tuffac. sandstone

Slonea berteriana Aluminum (mg/g) II1A 0.09 0.04 2 0.09 0.12 0.05 67 Smf Ultisols Tuffac. sandstone

II1C 0.02 1 67 Smf Ultisols Tuffac. sandstone

II1H 0.16 1 135 Lmrf Ultisols Tuffac. sandstone

II7A 1.35 1.82 5 0.23 4.35 0.12 70 Lmrf Ultisols Tuffac. sandstone 1year

II7A 0.47 0.30 3 0.36 0.81 0.24 70 Lmrf Ultisols Tuffac. sandstone 2year

II7B 0.05 0.05 4 0.05 0.11 0.01 70 Lmrf Ultisols Tuffac. sandstone 1year

II7B 0.07 0.07 3 0.05 0.14 0.01 70 Lmrf Ultisols Tuffac. sandstone 2year

II7C 0.31 0.37 5 0.04 0.71 0.03 70 Lmrf Ultisols Tuffac. sandstone 1year

II7C 0.46 0.32 3 0.52 0.75 0.12 70 Lmrf Ultisols Tuffac. sandstone 2year

Ash (%) II1A 3.81 0.64 2 3.81 4.36 3.26 67 Smf Ultisols Tuffac. sandstone

II1C 0.75 1 67 Smf Ultisols Tuffac. sandstone

C/N II1A 49 21 2 49 70 30 67 Smf Ultisols Tuffac. sandstone

II1C 371 1 67 Smf Ultisols Tuffac. sandstone

Calcium (mg/g) II1A 7.78 0.36 2 7.78 8.27 7.39 67 Smf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

287

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 3.40 1.43 6 4.04 4.53 0.70 135 Lmrf Ultisols Tuffac. sandstone

II1B 6.86 3.69 5 6.06 1.31 3.11 135 Lmrf Ultisols Tuffac. sandstone

II1C 2.02 1 67 Smf Ultisols Tuffac. sandstone

II1C 2.26 0.63 8 2.46 2.92 1.14 135 Lmrf Ultisols Tuffac. sandstone

II1Ci 10.43 1.18 8 10.52 12.07 7.89 135 Lmrf Ultisols Tuffac. sandstone

II1F 4.83 1 135 Lmrf Ultisols Tuffac. sandstone

II1H 2.73 1 135 Lmrf Ultisols Tuffac. sandstone

II7A 12.85 0.46 5 12.94 13.50 12.30 70 Lmrf Ultisols Tuffac. sandstone 1year

II7A 6.72 3.27 3 7.33 9.64 3.19 70 Lmrf Ultisols Tuffac. sandstone 2year

II7B 10.05 4.10 4 10.56 14.01 5.10 70 Lmrf Ultisols Tuffac. sandstone 1year

II7B 5.39 3.40 3 5.33 8.82 2.02 70 Lmrf Ultisols Tuffac. sandstone 2year

II7C 16.19 9.47 5 21.69 25.37 5.11 70 Lmrf Ultisols Tuffac. sandstone 1year

II7C 3.24 0.76 3 3.05 4.08 2.60 70 Lmrf Ultisols Tuffac. sandstone 2year

Carbon (%) II1A 47 5 101 47 61 36 150 Swf Ultisols Tuffac. sandstone

Iron (mg/g) II1A 0.07 0.01 2 0.07 0.07 0.05 67 Smf Ultisols Tuffac. sandstone

II1C 0.02 1 67 Smf Ultisols Tuffac. sandstone

II1H 0.17 1 135 Lmrf Ultisols Tuffac. sandstone

II7A 1.99 2.84 5 0.27 6.44 0.12 70 Lmrf Ultisols Tuffac. sandstone 1year

II7A 0.49 0.41 3 0.28 0.96 0.23 70 Lmrf Ultisols Tuffac. sandstone 2year

II7B 0.08 0.06 4 0.08 0.15 0.01 70 Lmrf Ultisols Tuffac. sandstone 1year

II7B 0.07 0.04 3 0.06 0.12 0.03 70 Lmrf Ultisols Tuffac. sandstone 2year

II7C 0.50 0.61 5 0.09 1.18 0.03 70 Lmrf Ultisols Tuffac. sandstone 1year

II7C 0.49 0.43 3 0.40 0.95 0.12 70 Lmrf Ultisols Tuffac. sandstone 2year

Magnesium (mg/g) II1A 1.46 0.25 2 1.46 1.71 1.22 67 Smf Ultisols Tuffac. sandstone

II1A 1.72 0.67 6 1.94 2.40 0.47 135 Lmrf Ultisols Tuffac. sandstone

II1B 1.77 0.61 5 1.54 2.47 0.99 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.30 1 67 Smf Ultisols Tuffac. sandstone

II1C 0.30 0.02 8 0.30 0.33 0.27 135 Lmrf Ultisols Tuffac. sandstone

II1Ci 0.68 0.08 8 0.67 0.79 0.60 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

288

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1F 1.57 1 135 Lmrf Ultisols Tuffac. sandstone

II1H 1.61 1 135 Lmrf Ultisols Tuffac. sandstone

II7A 1.40 0.67 5 1.00 2.38 0.87 70 Lmrf Ultisols Tuffac. sandstone 1year

II7A 0.79 0.51 3 0.54 1.37 0.45 70 Lmrf Ultisols Tuffac. sandstone 2year

II7B 1.13 0.84 4 1.06 1.99 0.39 70 Lmrf Ultisols Tuffac. sandstone 1year

II7B 0.34 0.20 3 0.44 0.48 0.11 70 Lmrf Ultisols Tuffac. sandstone 2year

II7C 1.53 0.96 5 2.03 2.55 0.46 70 Lmrf Ultisols Tuffac. sandstone 1year

II7C 0.39 0.20 3 0.34 0.60 0.22 70 Lmrf Ultisols Tuffac. sandstone 2year

Manganese (mg/g) II1A 0.29 0.23 2 0.29 0.50 0.09 67 Smf Ultisols Tuffac. sandstone

II1C 0.04 1 67 Smf Ultisols Tuffac. sandstone

II1H 0.09 1 135 Lmrf Ultisols Tuffac. sandstone

II7A 0.24 0.18 5 0.14 0.47 0.10 70 Lmrf Ultisols Tuffac. sandstone 1year

II7A 0.17 0.09 3 0.20 0.25 0.07 70 Lmrf Ultisols Tuffac. sandstone 2year

II7B 0.13 0.10 4 0.13 0.23 0.05 70 Lmrf Ultisols Tuffac. sandstone 1year

II7B 0.09 0.11 3 0.03 0.22 0.02 70 Lmrf Ultisols Tuffac. sandstone 2year

II7C 0.45 0.40 5 0.57 0.97 0.05 70 Lmrf Ultisols Tuffac. sandstone 1year

II7C 0.11 0.10 3 0.07 0.22 0.04 70 Lmrf Ultisols Tuffac. sandstone 2year

Nitrogen (%) II1A 1.04 0.44 2 1.04 1.43 0.64 67 Smf Ultisols Tuffac. sandstone

II1A 1.22 0.39 5 1.40 1.52 0.55 135 Lmrf Ultisols Tuffac. sandstone

II1A 1.89 0.62 101 1.72 4.20 0.92 150 Swf Ultisols Tuffac. sandstone

II1B 0.93 0.33 5 0.93 1.45 0.57 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.13 1 67 Smf Ultisols Tuffac. sandstone

II1C 0.12 0.02 8 0.11 0.16 0.10 135 Lmrf Ultisols Tuffac. sandstone

II1Ci 0.49 0.06 8 0.48 0.63 0.46 135 Lmrf Ultisols Tuffac. sandstone

II1F 0.85 1 135 Lmrf Ultisols Tuffac. sandstone

II1H 1.59 1 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.56 0.10 2 0.56 0.65 0.47 67 Smf Ultisols Tuffac. sandstone

II1A 0.68 0.08 6 0.62 0.77 0.57 135 Lmrf Ultisols Tuffac. sandstone

II1B 0.62 0.16 5 0.61 0.81 0.38 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

289

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1C 0.08 1 67 Smf Ultisols Tuffac. sandstone

II1C 0.05 0.01 8 0.04 0.07 0.03 135 Lmrf Ultisols Tuffac. sandstone

II1Ci 0.16 0.02 8 0.17 0.19 0.13 135 Lmrf Ultisols Tuffac. sandstone

II1F 0.68 1 135 Lmrf Ultisols Tuffac. sandstone

II1H 0.84 1 135 Lmrf Ultisols Tuffac. sandstone

II7A 0.35 0.18 5 0.25 0.60 0.19 70 Lmrf Ultisols Tuffac. sandstone 1year

II7A 0.17 0.05 3 0.16 0.22 0.13 70 Lmrf Ultisols Tuffac. sandstone 2year

II7B 0.15 0.08 4 0.16 0.22 0.06 70 Lmrf Ultisols Tuffac. sandstone 1year

II7B 0.09 0.06 3 0.07 0.16 0.04 70 Lmrf Ultisols Tuffac. sandstone 2year

II7C 0.28 0.19 5 0.25 0.49 0.08 70 Lmrf Ultisols Tuffac. sandstone 1year

II7C 0.19 0.05 3 0.21 0.24 0.14 70 Lmrf Ultisols Tuffac. sandstone 2year

Potassium (mg/g) II1A 8.85 2.39 2 8.85 11.83 6.90 67 Smf Ultisols Tuffac. sandstone

II1A 7.57 1.76 6 6.97 10.87 6.32 135 Lmrf Ultisols Tuffac. sandstone

II1B 5.76 0.82 5 5.52 6.88 5.03 135 Lmrf Ultisols Tuffac. sandstone

II1C 2.19 1 67 Smf Ultisols Tuffac. sandstone

II1C 1.77 0.33 8 1.65 2.53 1.53 135 Lmrf Ultisols Tuffac. sandstone

II1Ci 5.35 0.37 8 5.30 6.07 4.86 135 Lmrf Ultisols Tuffac. sandstone

II1F 6.00 1 135 Lmrf Ultisols Tuffac. sandstone

II1H 13.17 1 135 Lmrf Ultisols Tuffac. sandstone

II7A 2.17 1.12 5 2.86 3.06 0.78 70 Lmrf Ultisols Tuffac. sandstone 1year

II7A 0.98 0.37 3 1.03 1.32 0.58 70 Lmrf Ultisols Tuffac. sandstone 2year

II7B 1.31 0.73 4 1.18 2.23 0.65 70 Lmrf Ultisols Tuffac. sandstone 1year

II7B 0.73 0.21 3 0.63 0.97 0.06 70 Lmrf Ultisols Tuffac. sandstone 2year

II7C 1.58 1.03 5 0.87 2.75 0.80 70 Lmrf Ultisols Tuffac. sandstone 1year

II7C 0.99 0.56 3 0.79 1.63 0.57 70 Lmrf Ultisols Tuffac. sandstone 2year

Sulfur (%) II1A 0.31 0.09 68 0.31 0.55 0.12 150 Swf Ultisols Tuffac. sandstone

Solanum torvum Aluminum (mg/g) II8A 1.55 1.33 2 1.55 2.49 0.61 146 Swf Ultisols Tuffac. sandstone

II8B 0.15 0.15 2 0.15 0.26 0.04 146 Swf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

290

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Ash (%) II8A 8.50 1.61 2 8.50 9.64 7.36 146 Swf Ultisols Tuffac. sandstone

II8B 3.34 1.82 2 3.34 4.63 2.05 146 Swf Ultisols Tuffac. sandstone

C/N II8A 15 1 2 15 16 14 146 Swf Ultisols Tuffac. sandstone

II8B 64 4 2 64 67 61 146 Swf Ultisols Tuffac. sandstone

Calcium (mg/g) II8A 12.91 5.29 2 12.91 16.65 9.17 146 Swf Ultisols Tuffac. sandstone

II8B 8.81 0.39 2 8.81 9.08 8.53 146 Swf Ultisols Tuffac. sandstone

Carbon (%) II8A 52 1 2 52 52 51 146 Swf Ultisols Tuffac. sandstone

II8B 52 1 2 52 53 51 146 Swf Ultisols Tuffac. sandstone

Iron (mg/g) II8A 1.14 0.78 2 1.14 1.70 0.59 146 Swf Ultisols Tuffac. sandstone

II8B 0.09 0.10 2 0.09 0.16 0.02 146 Swf Ultisols Tuffac. sandstone

Magnesium (mg/g) II8A 2.70 2.01 2 2.70 4.12 1.28 146 Swf Ultisols Tuffac. sandstone

II8B 2.07 2.14 2 2.07 0.58 0.56 146 Swf Ultisols Tuffac. sandstone

Manganese (mg/g) II8A 0.14 0.11 2 0.14 0.22 0.06 146 Swf Ultisols Tuffac. sandstone

II8B 0.03 0.02 2 0.03 0.05 0.02 146 Swf Ultisols Tuffac. sandstone

Nitrogen (%) II8A 3.52 0.43 2 3.52 3.82 3.21 146 Swf Ultisols Tuffac. sandstone

II8B 0.81 0.08 2 0.81 0.87 0.75 146 Swf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II8A 1.31 0.67 2 1.31 1.78 0.83 146 Swf Ultisols Tuffac. sandstone

II8B 0.58 0.13 2 0.58 0.67 0.48 146 Swf Ultisols Tuffac. sandstone

Potassium (mg/g) II8A 10.11 12.12 2 10.11 18.68 1.54 146 Swf Ultisols Tuffac. sandstone

II8B 5.58 5.46 2 0.58 9.44 1.71 146 Swf Ultisols Tuffac. sandstone

Spathodea campanulata Ash (%) II1A 6.70 0.50 5 6.94 7.20 6.10 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 2.68 0.58 5 2.83 3.45 2.04 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 11.08 1.88 6 11.03 13.18 8.53 126 Ultisols Siltstone,sandstone Urban forest, RP

C/N II1A 19 2 5 20 21 16 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 299 104 5 281 422 187 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 65 18 6 62 88 47 126 Ultisols Siltstone,sandstone Urban forest, RP

Carbon (%) II1A 51 1 5 51 52 51 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 50 0 5 51 51 50 126 Ultisols Siltstone,sandstone Urban forest, RP
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

291

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1Ci 46 1 6 46 47 45 126 Ultisols Siltstone,sandstone Urban forest, RP

Nitrogen (%) II1A 2.66 0.35 5 2.58 3.26 2.36 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 0.19 0.06 5 0.18 0.27 0.12 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 0.75 0.20 6 0.76 1.00 0.52 126 Ultisols Siltstone,sandstone Urban forest, RP

Sulfur (%) II1A 0.28 0.03 5 0.29 0.31 0.24 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 0.04 0.02 5 0.50 0.60 0.01 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 0.09 0.03 6 0.09 0.11 0.05 126 Ultisols Siltstone,sandstone Urban forest, RP

Wood density (g/cc) II1B 0.26 0.01 3 0.26 0.27 0.26 97 Wfs Inceptisols Tuffac. sandstone

II1C 0.23 0.06 5 0.22 0.32 0.18 126 Ultisols Siltstone,sandstone Urban forest, RP

Swietenia humilis Aluminum (mg/kg) II1A 62 20 6 50 94 45 103 Swf Ultisols Lava,tuff Mature-Coamo

II1A 168 29 3 160 206 136 103 Swf Ultisols Lava, tuff Mature-Guavate

Calcium (mg/g) II1A 18.73 4.17 6 19.10 25.69 12.93 103 Swf Ultisols Lava,tuff Mature-Coamo

II1A 16.97 4.02 3 16.04 21.93 12.32 103 Swf Ultisols Lava, tuff Mature-Guavate

Magnesium (mg/g) II1A 2.46 0.44 6 2.48 3.03 1.79 103 Swf Ultisols Lava,tuff Mature-Coamo

II1A 2.87 0.17 3 2.88 3.08 2.66 103 Swf Ultisols Lava, tuff Mature-Guavate

Manganese (mg/kg) II1A 14 6 6 15 22 5 103 Swf Ultisols Lava,tuff Mature-Coamo

II1A 12 8 3 10 23 5 103 Swf Ultisols Lava, tuff Mature-Guavate

Nitrogen (%) II1A 1.57 0.54 6 1.32 2.42 1.05 103 Swf Ultisols Lava,tuff Mature-Coamo

II1A 1.88 0.25 3 2.01 2.08 1.52 103 Swf Ultisols Lava, tuff Mature-Guavate

Phosphorus (mg/g) II1A 0.77 0.15 6 0.79 1.03 0.59 103 Swf Ultisols Lava,tuff Mature-Coamo

II1A 0.88 0.04 3 0.89 0.92 0.81 103 Swf Ultisols Lava, tuff Mature-Guavate

Potassium (mg/g) II1A 8.14 1.91 6 8.57 10.39 5.07 103 Swf Ultisols Lava,tuff Mature-Coamo

II1A 13.95 1.90 3 14.24 16.09 11.69 103 Swf Ultisols Lava, tuff Mature-Guavate

Swietenia macrophylla Aluminum (mg/kg) II1A 740 50 2 740 780 710 129 Lmrf Ultisols Tuffac. sandstone

II1A 124 45 5 123 213 64 103 Swf Ultisols Lava, tuff Mature-Guavate

II1A 42 11 6 37 61 30 103 Swf Ultisols Lava,tuff Mature-Coamo

Ash (%) II1A 7.41 1.17 6 7.78 8.76 5.66 94 Swf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

292

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 8.13 1 101 Lmwf Inceptisols Tuffac. sandstone Fresh 

II1A 5.64 1 101 SWf Ultisols Tuffac. sandstone Fresh 

II1A 6.87 0.02 2 6.87 6.88 6.85 129 Lmrf Ultisols Tuffac. sandstone

II1B 4.07 0.71 3 3.95 4.84 3.43 94 Swf Ultisols Tuffac. sandstone

II1C 3.80 0.87 6 3.90 4.72 2.42 94 Swf Ultisols Tuffac. sandstone

II8A 5.34 1.48 9 5.54 7.06 2.37 90 Swf Ultisols Tuffac. sandstone

II8B 4.32 0.77 4 4.43 5.08 3.36 90 Swf Ultisols Tuffac. sandstone

II8D 17.64 4.47 4 17.32 23.00 12.90 90 Swf Ultisols Tuffac. sandstone

C/N II1A 24 1 129 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II1A 10.15 4.41 5 12.54 14.10 3.53 104 Sdf Mollisols Alluvial deposits New

II1A 11.73 2.60 6 12.20 15.37 8.55 104 Sdf Mollisols Alluvial deposits Young 

II1A 11.60 5.79 15 11.59 24.37 1.85 104 Sdf Mollisols Alluvial deposits

II1A 15.60 3.14 5 14.52 20.81 10.88 103 Swf Ultisols Lava, tuff Mature-Guavate

II1A 17.22 2.26 6 17.42 21.98 13.75 103 Swf Ultisols Lava,tuff Mature-Coamo

II1A 20.18 1.73 2 20.18 21.40 18.96 129 Lmrf Ultisols Tuffac. sandstone

II8A 15.02 4.93 9 16.68 20.19 5.85 90 Swf Ultisols Tuffac. sandstone

II8B 12.71 0.47 4 12.52 13.24 12.36 90 Swf Ultisols Tuffac. sandstone

II8D 17.24 5.64 4 19.84 21.11 10.76 90 Swf Ultisols Tuffac. sandstone

II9A 14.07 3.20 22 13.97 18.35 6.90 139 Wfs Ultisols Tuffac. sandstone

II9A 9.03 2.59 22 9.37 15.42 4.59 139 Wfs Ultisols Tuffac. sandstone Other

II9B 5.76 2.38 7 5.00 10.59 3.03 139 Wfs Ultisols Tuffac. sandstone

II9C 4.43 2.23 6 5.12 6.78 1.52 139 Wfs Ultisols Tuffac. sandstone

II9D 8.40 2.56 5 7.65 12.82 6.41 139 Wfs Ultisols Tuffac. sandstone

Carbon (%) II1A 53 0 2 53 53 52 129 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II1A 1.41 0.14 2 1.41 1.51 1.31 129 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II1A 1.26 0.36 5 1.45 1.62 0.79 104 Sdf Mollisols Alluvial deposits New

II1A 1.45 0.47 6 1.35 2.22 0.84 104 Sdf Mollisols Alluvial deposits Young 

II1A 1.32 0.25 17 1.28 2.00 0.88 104 Sdf Mollisols Alluvial deposits

II1A 2.77 0.48 5 2.81 3.63 2.03 103 Swf Ultisols Lava, tuff Mature-Guavate
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

293

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 2.40 0.80 6 2.34 3.89 1.18 103 Swf Ultisols Lava,tuff Mature-Coamo

II1A 2.02 0.17 2 2.02 2.14 1.90 129 Lmrf Ultisols Tuffac. sandstone

II8A 2.55 0.56 9 2.63 3.20 1.38 90 Swf Ultisols Tuffac. sandstone

II8B 2.70 0.25 4 2.72 2.94 2.44 90 Swf Ultisols Tuffac. sandstone

II8D 3.00 0.42 4 3.22 3.26 2.52 90 Swf Ultisols Tuffac. sandstone

II9A 3.63 0.86 22 3.43 5.70 2.27 139 Wfs Ultisols Tuffac. sandstone

II9A 3.97 1.48 22 4.16 7.29 2.02 139 Wfs Ultisols Tuffac. sandstone Other

II9B 1.64 0.82 7 1.69 3.06 0.52 139 Wfs Ultisols Tuffac. sandstone

II9C 2.48 0.83 6 2.49 3.61 1.31 139 Wfs Ultisols Tuffac. sandstone

II9D 2.72 0.80 5 2.58 3.89 1.80 139 Wfs Ultisols Tuffac. sandstone

Manganese (mg/g) II1A 0.06 0.00 2 0.06 0.06 0.06 129 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II1A 1.90 0.15 6 1.96 2.05 1.74 94 Swf Ultisols Tuffac. sandstone

II1A 1.66 1 101 Lmwf Inceptisols Tuffac. sandstone Fresh 

II1A 1.86 1 101 SWf Ultisols Tuffac. sandstone Fresh 

II1A 1.02 0.16 5 0.99 1.20 0.77 104 Sdf Mollisols Alluvial deposits New

II1A 1.25 0.09 6 1.28 1.33 1.01 104 Sdf Mollisols Alluvial deposits Young 

II1A 1.27 0.83 14 0.94 3.34 0.41 104 Sdf Mollisols Alluvial deposits

II1A 1.62 0.14 5 1.55 1.83 1.44 103 Swf Ultisols Lava, tuff Mature-Guavate

II1A 1.46 0.16 6 1.38 1.77 1.29 103 Swf Ultisols Lava,tuff Mature-Coamo

II1A 2.23 0.01 2 2.23 2.23 2.22 129 Lmrf Ultisols Tuffac. sandstone

II1B 0.48 0.11 3 0.46 0.59 0.38 94 Swf Ultisols Tuffac. sandstone

II1C 0.58 0.23 6 0.58 0.96 0.27 94 Swf Ultisols Tuffac. sandstone

II8A 1.15 0.21 9 1.22 1.47 0.81 90 Swf Ultisols Tuffac. sandstone

II8B 0.70 0.21 4 0.69 0.92 0.51 90 Swf Ultisols Tuffac. sandstone

II8D 1.47 0.08 4 1.45 1.57 1.40 90 Swf Ultisols Tuffac. sandstone

II9A 0.97 0.24 22 0.98 1.56 0.63 139 Wfs Ultisols Tuffac. sandstone

II9A 1.55 0.41 22 1.44 2.35 0.93 139 Wfs Ultisols Tuffac. sandstone Other

II9B 0.37 0.17 12 0.37 0.64 0.15 139 Wfs Ultisols Tuffac. sandstone

II9C 0.72 0.17 11 0.70 0.97 0.53 139 Wfs Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

294

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II9D 1.17 0.18 9 1.43 1.82 1.24 139 Wfs Ultisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.58 0.18 5 0.60 0.84 0.33 104 Sdf Mollisols Alluvial deposits New

II1A 0.73 0.19 6 0.78 0.95 0.48 104 Sdf Mollisols Alluvial deposits Young 

II1A 0.81 0.93 17 0.48 3.63 0.16 104 Sdf Mollisols Alluvial deposits

II1A 0.73 0.18 5 0.71 1.14 0.54 103 Swf Ultisols Lava, tuff Mature-Guavate

II1A 0.69 0.12 6 0.70 0.86 0.54 103 Swf Ultisols Lava,tuff Mature-Coamo

II1A 1.10 0.24 6 1.03 1.39 0.84 94 Swf Ultisols Tuffac. sandstone

II1A 0.88 1 101 Lmwf Inceptisols Tuffac. sandstone Fresh 

II1A 0.88 1 101 SWf Ultisols Tuffac. sandstone Fresh 

II1A 0.55 0.03 2 0.55 0.57 0.53 129 Lmrf Ultisols Tuffac. sandstone

II1B 0.56 0.04 3 0.57 0.59 0.51 94 Swf Ultisols Tuffac. sandstone

II1C 0.94 0.71 6 0.72 2.12 0.23 94 Swf Ultisols Tuffac. sandstone

II8A 0.31 0.07 9 0.28 0.44 0.24 90 Swf Ultisols Tuffac. sandstone

II8B 0.20 0.05 4 0.20 0.25 0.15 90 Swf Ultisols Tuffac. sandstone

II8D 0.44 0.06 4 0.46 0.49 0.38 90 Swf Ultisols Tuffac. sandstone

II9A 0.39 0.15 22 0.37 0.71 0.12 139 Wfs Ultisols Tuffac. sandstone

II9A 0.66 0.40 22 0.56 2.10 0.26 139 Wfs Ultisols Tuffac. sandstone Other

II9B 0.22 0.22 7 0.14 0.64 0.03 139 Wfs Ultisols Tuffac. sandstone

II9C 0.57 0.27 6 0.59 0.99 0.27 139 Wfs Ultisols Tuffac. sandstone

II9D 1.28 0.80 5 1.06 2.68 0.71 139 Wfs Ultisols Tuffac. sandstone

Potassium (mg/g) II1A 10.23 4.08 6 9.60 15.52 6.26 94 Swf Ultisols Tuffac. sandstone

II1A 6.00 1 101 Lmwf Inceptisols Tuffac. sandstone Fresh 

II1A 8.00 1 101 SWf Ultisols Tuffac. sandstone Fresh 

II1A 9.84 2.67 5 8.08 13.63 7.32 104 Sdf Mollisols Alluvial deposits New

II1A 14.59 5.04 6 17.52 18.81 7.23 104 Sdf Mollisols Alluvial deposits Young 

II1A 9.33 4.09 17 7.57 18.72 2.27 104 Sdf Mollisols Alluvial deposits

II1A 14.04 3.71 5 12.87 22.70 10.37 103 Swf Ultisols Lava, tuff Mature-Guavate

II1A 7.98 1.28 6 7.64 10.55 6.74 103 Swf Ultisols Lava,tuff Mature-Coamo

II1A 5.01 0.54 2 5.01 5.39 4.63 129 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

295

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1B 7.60 0.39 3 7.51 8.03 7.26 94 Swf Ultisols Tuffac. sandstone

II1C 6.34 2.81 6 5.10 10.30 3.00 94 Swf Ultisols Tuffac. sandstone

II8A 2.61 1.36 9 2.15 5.08 1.20 90 Swf Ultisols Tuffac. sandstone

II8B 1.95 0.94 4 1.81 2.95 1.08 90 Swf Ultisols Tuffac. sandstone

II8D 2.43 1.35 4 2.32 3.83 1.14 90 Swf Ultisols Tuffac. sandstone

II9A 5.40 1.31 22 5.26 7.98 3.01 139 Wfs Ultisols Tuffac. sandstone

II9A 5.01 2.09 22 4.09 10.78 2.76 139 Wfs Ultisols Tuffac. sandstone Other

II9B 2.44 1.85 7 2.22 5.48 0.45 139 Wfs Ultisols Tuffac. sandstone

II9C 4.47 1.16 6 4.58 5.97 3.14 139 Wfs Ultisols Tuffac. sandstone

II9D 4.40 1.89 5 3.56 6.99 2.64 139 Wfs Ultisols Tuffac. sandstone

Sulfur (%) II1A 0.39 0.02 2 0.39 0.40 0.38 129 Lmrf Ultisols Tuffac. sandstone

Swietenia macrophylla Aluminum (mg/kg) II1A 166 119 3 93 323 85 103 Swf Ultisols Lava, tuff Guavate c.-shady 

xmahagoni II1A 90 41 10 89 185 20 103 Swf Ultisols Lava, tuff Guavate fert.-shady

II1A 137 36 8 123 208 94 103 Swf Ultisols Lava, tuff Guavate fert.-sun

II1A 307 140 4 260 551 176 103 Lmwf Inceptisols Tuffac. sandstone Harvey 

II1A 161 66 5 189 227 60 103 SWf Ultisols Tuffac. sandstone Rio Chiquito

II1A 102 15 3 104 118 82 103 Smf Alfisols Sandstone,sillstone Vieques c.-sun

II1A 129 24 5 129 160 92 103 Swf Ultisols Lava, tuff Guavate c.-sun

Calcium (mg/g) II1A 13.89 3.91 3 15.61 17.90 8.55 103 Swf Ultisols Lava, tuff Guavate c.-shady 

II1A 16.88 5.01 10 16.87 27.21 10.30 103 Swf Ultisols Lava, tuff Guavate fert.-shady

II1A 13.29 2.76 8 13.15 17.80 9.14 103 Swf Ultisols Lava, tuff Guavate fert.-sun

II1A 19.19 3.58 4 19.13 24.91 12.54 103 Lmwf Inceptisols Tuffac. sandstone Harvey 

II1A 10.10 3.67 5 8.25 16.08 6.37 103 SWf Ultisols Tuffac. sandstone Rio Chiquito

II1A 8.21 2.73 3 7.51 11.57 5.53 103 Smf Alfisols Sandstone,sillstone Vieques c.-sun

II1A 13.19 2.93 5 13.43 18.06 9.59 103 Swf Ultisols Lava, tuff Guavate c.-sun

II1C 12.99 1 103 SWf Ultisols Tuffac. sandstone Rio Chiquito

Magnesium (mg/g) II1A 1.92 0.43 3 1.86 2.50 1.40 103 Swf Ultisols Lava, tuff Guavate c.-shady 

II1A 1.51 0.36 10 1.44 2.33 1.15 103 Swf Ultisols Lava, tuff Guavate fert.-shady
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

296

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 2.2 0.50 8 1.96 3.25 1.76 103 Swf Ultisols Lava, tuff Guavate fert.-sun

II1A 2.42 0.35 4 2.41 2.93 1.95 103 Lmwf Inceptisols Tuffac. sandstone Harvey 

II1A 2.65 0.69 5 2.46 3.59 1.79 103 SWf Ultisols Tuffac. sandstone Rio Chiquito

II1A 1.43 0.14 3 1.46 1.59 1.25 103 Smf Alfisols Sandstone,sillstone Vieques c.-sun

II1A 1.71 0.26 5 1.59 2.09 1.42 103 Swf Ultisols Lava, tuff Guavate c.-sun

II1C 0.53 1 103 SWf Ultisols Tuffac. sandstone Rio Chiquito

Manganese (mg/kg) II1A 23 6 3 27 27 14 103 Swf Ultisols Lava, tuff Guavate c.-shady 

II1A 34 12 10 34 57 15 103 Swf Ultisols Lava, tuff Guavate fert.-shady

II1A 20 7 8 18 34 11 103 Swf Ultisols Lava, tuff Guavate fert.-sun

II1A 38 16 4 36 63 17 103 Lmwf Inceptisols Tuffac. sandstone Harvey 

II1A 21 6 5 22 28 13 103 SWf Ultisols Tuffac. sandstone Rio Chiquito

II1A 20 7 3 18 29 14 103 Smf Alfisols Sandstone,sillstone Vieques c.-sun

II1A 41 12 5 37 56 26 103 Swf Ultisols Lava, tuff Guavate c.-sun

Nitrogen (%) II1A 1.66 0.33 3 1.85 1.90 1.22 103 Swf Ultisols Lava, tuff Guavate c.-shady 

II1A 1.77 0.21 10 1.79 2.18 1.40 103 Swf Ultisols Lava, tuff Guavate fert.-shady

II1A 1.73 0.15 8 1.71 1.92 1.46 103 Swf Ultisols Lava, tuff Guavate fert.-sun

II1A 1.72 0.06 4 1.74 1.78 1.61 103 Lmwf Inceptisols Tuffac. sandstone Harvey 

II1A 1.75 0.29 5 1.84 2.04 1.22 103 SWf Ultisols Tuffac. sandstone Rio Chiquito

II1A 1.25 0.10 3 1.30 1.34 1.12 103 Smf Alfisols Sandstone,sillstone Vieques c.-sun

II1A 1.64 0.16 5 1.67 1.83 1.38 103 Swf Ultisols Lava, tuff Guavate c.-sun

II1C 0.28 1 103 SWf Ultisols Tuffac. sandstone Rio Chiquito

Phosphorus (mg/g) II1A 0.78 0.08 3 0.74 0.88 0.72 103 Swf Ultisols Lava, tuff Guavate c.-shady 

II1A 0.73 0.08 10 0.75 0.82 0.56 103 Swf Ultisols Lava, tuff Guavate fert.-shady

II1A 0.88 0.20 8 0.81 1.20 0.66 103 Swf Ultisols Lava, tuff Guavate fert.-sun

II1A 0.69 0.09 4 0.66 0.83 0.59 103 Lmwf Inceptisols Tuffac. sandstone Harvey 

II1A 0.70 0.25 5 0.59 1.19 0.55 103 SWf Ultisols Tuffac. sandstone Rio Chiquito

II1A 0.78 0.14 3 0.72 0.96 0.66 103 Smf Alfisols Sandstone,sillstone Vieques c.-sun

II1A 0.66 0.04 5 0.67 0.70 0.61 103 Swf Ultisols Lava, tuff Guavate c.-sun

II1C 0.16 1 103 SWf Ultisols Tuffac. sandstone Rio Chiquito
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

297

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Potassium (mg/g) II1A 14.53 2.52 3 15.80 16.83 11.20 103 Swf Ultisols Lava, tuff Guavate c.-shady 

II1A 10.49 1.66 10 9.95 12.86 8.42 103 Swf Ultisols Lava, tuff Guavate fert.-shady

II1A 16.4 2.58 8 16.41 21.11 12.91 103 Swf Ultisols Lava, tuff Guavate fert.-sun

II1A 13.45 4.67 4 12.18 21.36 8.68 103 Lmwf Inceptisols Tuffac. sandstone Harvey 

II1A 8.78 2.06 5 9.43 11.61 5.87 103 SWf Ultisols Tuffac. sandstone Rio Chiquito

II1A 6.65 1.37 3 6.03 8.50 5.46 103 Smf Alfisols Sandstone,sillstone Vieques c.-sun

II1A 10.36 2.12 5 10.18 13.03 7.79 103 Swf Ultisols Lava, tuff Guavate c.-sun

II1C 5.56 1 103 SWf Ultisols Tuffac. sandstone Rio Chiquito

Swietenia mahagoni Aluminum (mg/kg) II1A 71 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 20 2 2 20 18 22 102 Smf Limestone Tuffac. sandstone Young 

II1A 57 11 4 51 67 43 103 Swf Ultisols Lava,tuff Coamo

II1A 60 1 103 Smf Alfisols Sandstone,sillstone Young-Vieques

Ash (%) II1A 6.51 1 149 Sdf Mollisols Alluvial deposits Susua forest

Calcium (mg/g) II1A 4.31 1.09 2 4.31 3.54 5.08 102 Smf Limestone Tuffac. sandstone Young 

II1A 10.86 0.84 4 10.59 12.19 10.00 103 Swf Ultisols Lava,tuff Coamo

II1A 8.16 1 103 Smf Alfisols Sandstone,sillstone Young-Vieques

II1A 10.37 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1Ci 18.86 7.26 4 18.66 27.54 10.47 103 Lmwf Inceptisols Tuffac. sandstone Harvey

II1Ci 23.91 5.15 10 22.14 33.79 15.50 103 SWf Ultisols Tuffac. sandstone Rio Chiquito

Cobalt (μg/g) II1A 0.33 0.04 2 0.33 0.35 0.30 102 Smf Limestone Tuffac. sandstone Young 

Iron (mg/kg) II1A 101 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 35 2 2 35 37 33 102 Smf Limestone Tuffac. sandstone Young 

Magnesium (mg/g) II1A 2.64 0.14 2 2.64 2.74 2.54 102 Smf Limestone Tuffac. sandstone Young 

II1A 1.10 0.22 4 1.08 1.37 0.85 103 Swf Ultisols Lava,tuff Coamo

II1Aia 1.20 1 103 Smf Alfisols Sandstone,sillstone Young-Vieques

II1A 4.07 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1Ci 0.73 0.40 4 0.74 1.14 0.33 103 Lmwf Inceptisols Tuffac. sandstone Harvey

II1Ci 0.54 0.36 10 0.45 1.89 0.28 103 SWf Ultisols Tuffac. sandstone Rio Chiquito
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

298

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Manganese (mg/kg) II1A 18 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 20 2 2 20 19 22 102 Smf Limestone Tuffac. sandstone Young 

II1A 9 5 4 6 16 5 103 Swf Ultisols Lava,tuff Coamo

II1Aia 15 1 103 Smf Alfisols Sandstone,sillstone Young-Vieques

Nickel (μg/g) II1A 4.10 0.42 2 4.10 4.40 3.80 102 Smf Limestone Tuffac. sandstone Young 

Nitrogen (%) II1A 1.17 0.07 2 1.17 1.23 1.12 102 Smf Limestone Tuffac. sandstone Young 

II1A 1.32 0.19 4 1.33 1.51 1.10 103 Swf Ultisols Lava,tuff Coamo

II1Aia 1.26 1 103 Smf Alfisols Sandstone,sillstone Young-Vieques

II1A 1.10 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1Ci 0.59 0.14 4 0.67 0.70 0.40 103 Lmwf Inceptisols Tuffac. sandstone Harvey

II1Ci 0.31 0.09 10 0.28 0.50 0.20 103 SWf Ultisols Tuffac. sandstone Rio Chiquito

Phosphorus (mg/g) II1A 0.68 0.01 2 0.68 0.69 0.67 102 Smf Limestone Tuffac. sandstone Young 

II1A 0.69 0.03 4 0.69 0.74 0.64 103 Swf Ultisols Lava,tuff Coamo

II1Aia 0.83 1 103 Smf Alfisols Sandstone,sillstone Young-Vieques

II1A 0.43 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1Ci 0.35 0.19 4 0.40 0.51 0.01 103 Lmwf Inceptisols Tuffac. sandstone Harvey

II1Ci 0.13 0.04 10 0.13 0.20 0.06 103 SWf Ultisols Tuffac. sandstone Rio Chiquito

Potassium (mg/g) II1A 8.00 1.35 2 8.00 8.96 7.05 102 Smf Limestone Tuffac. sandstone Young 

II1A 7.20 0.48 4 7.27 7.97 6.49 103 Swf Ultisols Lava,tuff Coamo

II1Aia 8.46 1 103 Smf Alfisols Sandstone,sillstone Young-Vieques

II1A 5.53 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1Ci 14.03 4.72 4 14.03 19.01 9.10 103 Lmwf Inceptisols Tuffac. sandstone Harvey

II1Ci 6.82 1.06 10 7.17 8.22 5.02 103 SWf Ultisols Tuffac. sandstone Rio Chiquito

Zinc (μg/g) II1A 16.25 1 149 Sdf Mollisols Alluvial deposits Susua forest

Calcium (mg/g) II1A 11.35 4.91 51 12.07 24.37 1.85 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 27.16 7.64 4 27.27 33.93 20.15 149 Sdf Mollisols Alluvial deposits Guanica Forest

Magnesium (mg/g) II1A 1.34 0.33 55 1.30 2.22 0.79 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 0.21 0.04 4 0.21 0.25 0.17 149 Sdf Mollisols Alluvial deposits Guanica Forest

Nitrogen (%) II1A 1.22 0.59 55 1.19 3.34 0.41 149 Sdf Mollisols Alluvial deposits Guanica Forest
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

299

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1Ci 0.27 0.03 4 0.27 0.31 0.24 149 Sdf Mollisols Alluvial deposits Guanica Forest

Phosphorus (mg/g) II1A 0.75 0.72 54 0.53 3.63 0.16 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 0.06 0.04 4 0.06 0.10 0.02 149 Sdf Mollisols Alluvial deposits Guanica Forest

Potassium (mg/g) II1A 10.57 4.57 55 8.10 18.81 2.27 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 7.22 1.86 4 7.23 9.02 5.39 149 Sdf Mollisols Alluvial deposits Guanica Forest

Swietenia sp Aluminum (mg/kg) II1A 26 17 16 20 70 10 103 Wfs Ultisols Lava Mature f.-Sabana

II1A 101 23 7 110 135 63 103 Wfs Ultisols Lava Old-Sabana

II1A 16 4 2 14 21 13 103 Wfs Ultisols Lava Young-Sabana

II1Aiib 151 24 3 161 175 120 103 Wfs Ultisols Lava Mature f.-Sabana

Calcium (mg/g) II1A 12.51 2.27 16 11.70 16.19 9.69 103 Wfs Ultisols Lava Mature f.-Sabana

II1A 16.97 3.26 7 18.39 20.55 11.65 103 Wfs Ultisols Lava Old-Sabana

II1A 4.32 0.50 2 4.30 4.80 3.87 103 Wfs Ultisols Lava Young-Sabana

II1Aiib 12.88 3.10 2 12.88 15.58 10.16 103 Wfs Ultisols Lava Mature f.-Sabana

Magnesium (mg/g) II1A 3.66 0.92 16 3.72 5.38 2.46 103 Wfs Ultisols Lava Mature f.-Sabana

II1A 3.62 0.95 7 3.16 5.47 2.49 103 Wfs Ultisols Lava Old-Sabana

II1A 2.52 0.64 2 2.50 3.12 1.95 103 Wfs Ultisols Lava Young-Sabana

II1Aiib 3.86 0.40 3 3.80 4.50 3.45 103 Wfs Ultisols Lava Mature f.-Sabana

Manganese (mg/kg) II1A 36 7 16 36 47 25 103 Wfs Ultisols Lava Mature f.-Sabana

II1A 43 15 7 42 75 27 103 Wfs Ultisols Lava Old-Sabana

II1A 14 4 2 15 18 10 103 Wfs Ultisols Lava Young-Sabana

II1Aiib 50 13 3 50 61 37 103 Wfs Ultisols Lava Mature f.-Sabana

Nitrogen (%) II1A 2.09 0.17 16 2.03 2.43 1.91 103 Wfs Ultisols Lava Mature f.-Sabana

II1A 1.77 0.28 7 1.81 2.34 1.44 103 Wfs Ultisols Lava Old-Sabana

II1A 1.77 0.22 2 1.77 1.98 1.57 103 Wfs Ultisols Lava Young-Sabana

II1Aiib 1.67 0.04 3 1.67 1.71 1.62 103 Wfs Ultisols Lava Mature f.-Sabana

Phosphorus (mg/g) II1A 0.73 0.10 16 0.74 0.85 0.54 103 Wfs Ultisols Lava Mature f.-Sabana

II1A 0.55 0.09 7 0.54 0.75 0.45 103 Wfs Ultisols Lava Old-Sabana

II1A 1.21 0.20 2 1.21 1.40 1.02 103 Wfs Ultisols Lava Young-Sabana
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

300

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1Aiib 0.68 0.07 3 0.66 0.77 0.60 103 Wfs Ultisols Lava Mature f.-Sabana

Potassium (mg/g) II1A 11.08 1.77 16 10.55 13.92 8.81 103 Wfs Ultisols Lava Mature f.-Sabana

II1A 8.80 1.96 7 8.31 12.30 5.77 103 Wfs Ultisols Lava Old-Sabana

II1A 14.35 1.66 2 14.37 15.81 12.86 103 Wfs Ultisols Lava Young-Sabana

II1Aiib 9.49 1.37 2 9.51 10.72 8.21 103 Wfs Ultisols Lava Mature f.-Sabana

Syzygium jambos Aluminum (mg/g) II1A 0.07 0.04 6 0.07 0.12 0.04 129 Lmrf Ultisols Tuffac. sandstone

II1A 0.19 0.02 2 0.19 0.21 0.18 131 Wfs Mollisols Plutonic rocks

II1B 0.15 0.05 2 0.15 0.18 0.11 131 Wfs Mollisols Plutonic rocks

II1C 0.12 0.00 2 0.12 0.13 0.12 131 Wfs Mollisols Plutonic rocks

II1E 0.61 0.19 3 0.52 0.83 0.49 131 Wfs Mollisols Plutonic rocks

II6 1.30 0.44 25 1.27 2.26 0.62 129 Lmrf Ultisols Tuffac. sandstone Decomp.

Ash (%) II1A 5.16 0.82 6 5.15 6.07 3.96 126 Ultisols Siltstone,sandstone Urban forest, RP

II1A 4.00 0.23 6 3.95 4.27 3.69 129 Lmrf Ultisols Tuffac. sandstone

II1A 5.59 1.05 8 4.83 6.82 4.02 94 Swf Ultisols Tuffac. sandstone Understory

II1A 4.15 0.25 2 4.15 4.33 3.97 131 Wfs Mollisols Plutonic rocks

II1B 4.71 0.75 6 4.22 5.79 3.92 94 Swf Ultisols Tuffac. sandstone Understory

II1B 2.30 0.81 2 2.30 2.87 1.72 101 Wfs Ultisols Lava

II1B 3.10 0.45 2 3.10 3.41 2.78 131 Wfs Mollisols Plutonic rocks

II1C 1.55 0.38 6 1.44 2.24 1.21 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 3.25 0.71 8 3.93 4.42 2.06 94 Swf Ultisols Tuffac. sandstone Understory

II1C 1.57 0.52 2 1.57 1.93 1.20 131 Wfs Mollisols Plutonic rocks

II1Ci 5.45 0.82 5 5.22 6.76 4.53 126 Ultisols Siltstone,sandstone Urban forest, RP

II1E 2.80 0.32 3 2.63 3.16 2.60 131 Wfs Mollisols Plutonic rocks

II6 7.74 1.15 25 7.67 9.69 5.35 129 Lmrf Ultisols Tuffac. sandstone Decomp.

C/N II1A 39 3 6 41 42 34 126 Ultisols Siltstone,sandstone Urban forest, RP

II1A 37 1 6 37 38 35 129 Lmrf Ultisols Tuffac. sandstone

II1A 36 2 2 36 37 34 131 Wfs Mollisols Plutonic rocks

II1B 79 9 2 79 85 72 131 Wfs Mollisols Plutonic rocks
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

301

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1C 406 193 6 306 655 235 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 126 1 2 126 127 126 131 Wfs Mollisols Plutonic rocks

II1Ci 90 22 5 86 118 59 126 Ultisols Siltstone,sandstone Urban forest, RP

II1E 60 25 3 47 89 45 131 Wfs Mollisols Plutonic rocks

II6 30 4 25 29 37 23 129 Lmrf Ultisols Tuffac. sandstone Decomp.

Calcium (mg/g) II1A 6.65 0.51 6 6.79 7.19 5.89 129 Lmrf Ultisols Tuffac. sandstone

II1A 6.33 0.75 2 6.33 6.86 5.81 131 Wfs Mollisols Plutonic rocks

II1B 3.30 1.42 2 3.30 4.31 2.29 101 Wfs Ultisols Lava

II1B 5.69 1.49 2 5.69 6.71 4.66 131 Wfs Mollisols Plutonic rocks

II1C 1.61 0.04 2 1.61 1.58 0.64 131 Wfs Mollisols Plutonic rocks

II1E 2.51 0.32 3 2.66 2.74 2.14 131 Wfs Mollisols Plutonic rocks

II6 11.17 1.67 25 11.81 14.80 7.86 129 Lmrf Ultisols Tuffac. sandstone Decomp.

Carbon (%) II1A 54 0 6 54 54 53 126 Ultisols Siltstone,sandstone Urban forest, RP

II1A 53 0 6 53 54 53 129 Lmrf Ultisols Tuffac. sandstone

II1A 55 0 2 55 55 54 131 Wfs Mollisols Plutonic rocks

II1B 52 0 2 52 52 51 131 Wfs Mollisols Plutonic rocks

II1C 52 0 6 52 52 51 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 52 0 2 52 52 52 131 Wfs Mollisols Plutonic rocks

II1Ci 52 0 5 52 53 52 126 Ultisols Siltstone,sandstone Urban forest, RP

II1E 53 1 3 53 53 52 131 Wfs Mollisols Plutonic rocks

II6 52 1 25 52 54 50 129 Lmrf Ultisols Tuffac. sandstone Decomp.

Iron (mg/g) II1A 0.05 0.01 6 0.05 0.08 0.04 129 Lmrf Ultisols Tuffac. sandstone

II1A 0.08 0.03 2 0.08 0.10 0.06 131 Wfs Mollisols Plutonic rocks

II1B 0.11 0.00 2 0.11 0.11 0.11 131 Wfs Mollisols Plutonic rocks

II1C 0.10 0.04 2 0.10 0.12 0.07 131 Wfs Mollisols Plutonic rocks

II1E 0.67 0.10 3 0.64 0.78 0.59 131 Wfs Mollisols Plutonic rocks

II6 1.64 0.88 25 1.40 4.35 0.50 129 Lmrf Ultisols Tuffac. sandstone Decomp.

Magnesium (mg/g) II1A 3.05 0.49 6 3.16 3.51 2.24 129 Lmrf Ultisols Tuffac. sandstone

II1A 3.74 0.29 2 3.74 3.94 3.53 131 Wfs Mollisols Plutonic rocks
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

302

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1B 2.43 0.61 2 2.43 2.86 2.01 131 Wfs Mollisols Plutonic rocks

II1C 0.95 0.11 2 0.95 1.03 0.87 131 Wfs Mollisols Plutonic rocks

II1E 1.70 0.20 3 1.78 1.85 1.47 131 Wfs Mollisols Plutonic rocks

II6 1.95 0.53 25 1.84 3.30 1.35 129 Lmrf Ultisols Tuffac. sandstone Decomp.

Manganese (mg/kg) II1A 22 14 6 16 42 9 129 Lmrf Ultisols Tuffac. sandstone

II6 103 99 25 73 550 34 129 Lmrf Ultisols Tuffac. sandstone Decomp.

II1A 8 2 2 8 9 6 131 Wfs Mollisols Plutonic rocks

II1B 26 13 2 26 35 16 131 Wfs Mollisols Plutonic rocks

II1C 6 1 131 Wfs Mollisols Plutonic rocks

II1E 24 3 2 24 26 22 131 Wfs Mollisols Plutonic rocks

Nitrogen (%) II1A 1.37 0.12 6 1.32 1.58 1.29 126 Ultisols Siltstone,sandstone Urban forest, RP

II1A 1.44 0.05 6 1.45 1.49 1.38 129 Lmrf Ultisols Tuffac. sandstone

II1A 1.07 0.16 8 1.06 1.30 0.85 94 Swf Ultisols Tuffac. sandstone Understory

II1A 1.53 0.10 2 1.53 1.60 1.46 131 Wfs Mollisols Plutonic rocks

II1B 0.61 0.10 6 0.58 0.82 0.51 94 Swf Ultisols Tuffac. sandstone Understory

II1B 0.53 0.12 2 0.53 0.61 0.44 101 Wfs Ultisols Lava

II1B 0.66 0.07 2 0.66 0.71 0.61 131 Wfs Mollisols Plutonic rocks

II1C 0.15 0.06 6 0.17 0.22 0.08 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 0.48 0.17 8 0.51 0.71 0.25 94 Swf Ultisols Tuffac. sandstone Understory

II1C 0.41 0.00 2 0.41 0.41 0.41 131 Wfs Mollisols Plutonic rocks

II1Ci 0.62 0.18 5 0.61 0.90 0.44 126 Ultisols Siltstone,sandstone Urban forest, RP

II1E 0.96 0.33 3 1.13 1.17 0.58 131 Wfs Mollisols Plutonic rocks

II6 1.81 0.30 25 1.77 2.38 1.36 129 Lmrf Ultisols Tuffac. sandstone Decomp.

Phosphorus (mg/g) II1A 0.73 0.05 6 0.74 0.77 0.62 129 Lmrf Ultisols Tuffac. sandstone

II1A 0.66 0.11 8 0.59 0.83 0.55 94 Swf Ultisols Tuffac. sandstone Understory

II1A 0.51 0.02 2 0.51 0.52 0.49 131 Wfs Mollisols Plutonic rocks

II1B 0.45 0.19 6 0.38 0.83 0.35 94 Swf Ultisols Tuffac. sandstone Understory

II1B 0.34 0.01 2 0.34 0.40 0.28 101 Wfs Ultisols Lava

II1B 0.29 0.06 2 0.29 0.33 0.25 131 Wfs Mollisols Plutonic rocks
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

303

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1C 0.47 0.25 8 0.38 0.87 0.18 94 Swf Ultisols Tuffac. sandstone Understory

II1C 0.20 0.01 2 0.20 0.20 0.19 131 Wfs Mollisols Plutonic rocks

II1E 0.23 0.01 3 0.23 0.24 0.22 131 Wfs Mollisols Plutonic rocks

II6 0.38 0.04 25 0.37 0.46 0.33 129 Lmrf Ultisols Tuffac. sandstone Decomp.

Potassium (mg/g) II1A 8.06 0.25 6 8.03 8.44 7.75 129 Lmrf Ultisols Tuffac. sandstone

II1A 5.00 1.20 8 5.30 6.90 3.20 94 Swf Ultisols Tuffac. sandstone Understory

II1A 3.98 0.18 2 3.98 4.11 3.86 131 Wfs Mollisols Plutonic rocks

II1B 3.40 1.40 6 4.10 4.80 1.30 94 Swf Ultisols Tuffac. sandstone Understory

II1B 4.30 0.33 2 4.30 4.54 4.07 101 Wfs Ultisols Lava Understory

II1B 3.34 0.10 2 3.34 3.41 2.27 131 Wfs Mollisols Plutonic rocks

II1C 3.60 1.40 8 3.30 5.50 1.00 94 Swf Ultisols Tuffac. sandstone

II1C 2.66 1.20 2 2.66 3.50 1.81 131 Wfs Mollisols Plutonic rocks

II1E 2.02 0.15 3 2.03 2.16 1.86 131 Wfs Mollisols Plutonic rocks

II6 0.68 0.21 25 0.64 1.19 0.31 129 Lmrf Ultisols Tuffac. sandstone Decomp.

Sulfur (%) II1A 0.14 0.01 6 0.14 0.15 0.11 126 Ultisols Siltstone,sandstone Urban forest, RP

II1A 0.20 0.01 6 0.20 0.22 0.18 129 Lmrf Ultisols Tuffac. sandstone

II1A 0.36 0.02 2 0.36 0.38 0.48 131 Wfs Mollisols Plutonic rocks

II1B 0.27 0.02 2 0.27 0.28 0.25 131 Wfs Mollisols Plutonic rocks

II1C 0.04 0.02 6 0.04 0.07 0.03 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 0.14 0.02 2 0.14 0.15 0.12 131 Wfs Mollisols Plutonic rocks

II1Ci 0.25 0.05 5 0.25 0.32 0.19 126 Ultisols Siltstone,sandstone Urban forest, RP

II1E 0.44 0.14 3 0.49 0.55 0.28 131 Wfs Mollisols Plutonic rocks

II6 0.15 0.02 25 0.15 0.19 0.11 129 Lmrf Ultisols Tuffac. sandstone Decomp.

Wood density (g/cc) II1C 0.56 0.05 5 0.55 0.62 0.51 126 Ultisols Siltstone,sandstone Urban forest, RP

Tabebuia aemantha Aluminum (mg/kg) II1A 17 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 10 1 102 Smf Limestone Tuffac. sandstone Adult 

Ash (%) II1A 5.79 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 3.56 1 102 Smf Limestone Tuffac. sandstone Adult 
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

304

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Calcium (mg/g) II1A 4.15 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 2.41 1 102 Smf Limestone Tuffac. sandstone Adult 

Cobalt (μg/g) II1A 0.70 1 102 Smf Limestone Tuffac. sandstone Adult 

Iron (mg/kg) II1A 66 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 16 1 102 Smf Limestone Tuffac. sandstone Adult 

Magnesium (mg/g) II1A 2.57 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 1.45 1 102 Smf Limestone Tuffac. sandstone Adult 

Manganese (mg/kg) II1A 18 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 8 1 102 Smf Limestone Tuffac. sandstone Adult 

Nickel (μg/g) II1A 40.65 1 102 Smf Limestone Tuffac. sandstone Adult 

Nitrogen (%) II1A 1.30 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 0.83 1 102 Smf Limestone Tuffac. sandstone Adult 

Phosphorus (mg/g) II1A 0.60 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 0.58 1 102 Smf Limestone Tuffac. sandstone Adult 

Potassium (mg/g) II1A 7.88 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 15.36 1 102 Smf Limestone Tuffac. sandstone

Zinc (μg/g) II1A 11.42 1 149 Sdf Mollisols Alluvial deposits Susua forest

Tababuia heterophylla Aluminum (μg/g) II1A 5.68 4.31 2 5.58 9.65 1.90 58 Lmrf Ultisols Serpentinite

Ash (%) II1A 4.82 0.60 3 4.52 5.51 4.42 101 Swf Ultisols Tuffac. sandstone

II1A 4.88 1.09 7 4.65 6.52 3.20 94 Swf Ultisols Tuffac. sandstone

II1B 4.23 1 94 Swf Ultisols Tuffac. sandstone

II1C 3.15 2.15 5 2.21 5.77 1.22 94 Swf Ultisols Tuffac. sandstone Fresh

II1A 5.06 0.39 5 4.91 5.67 4.66 126 Ultisols Siltstone,sandstone Urban forest, RP

II1A 3.37 0.88 51 3.15 6.87 2.23 138 Wfs Ultisols Lava

II1C 1.01 0.22 5 0.95 1.35 0.80 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 4.44 0.38 6 4.33 5.02 4.09 126 Ultisols Siltstone,sandstone Urban forest, RP

C/N II1A 32 2 5 31 35 28 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 368 112 5 346 556 264 126 Ultisols Siltstone,sandstone Urban forest, RP
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

305

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1Ci 128 29 6 133 166 87 126 Ultisols Siltstone,sandstone Urban forest, RP

Calcium (mg/g) II1A 5.32 0.25 2 5.34 5.56 5.05 58 Lmrf Ultisols Serpentinite

Carbon (%) II1A 53 0 5 53 53 52 126 Ultisols Siltstone,sandstone Urban forest, RP

II1A 50 1 51 50 51 48 126 Ultisols Siltstone,sandstone

II1C 55 0 5 55 56 55 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 54 0 6 55 55 54 126 Ultisols Siltstone,sandstone Urban forest, RP

Cobalt (μg/g) II1A 0.93 0.42 2 0.90 1.35 0.57 58 Lmrf Ultisols Serpentinite

Copper (μg/g) II1A 7.09 3.55 2 6.99 10.69 3.69 58 Lmrf Ultisols Serpentinite

Chromium (μg/g) II1A 108 80 2 109 178 35 58 Lmrf Ultisols Serpentinite

Iron (μg/g) II1A 551 586 2 524 1111 43 58 Lmrf Ultisols Serpentinite

Magnesium (mg/g) II1A 2.56 0.34 2 2.57 2.87 2.24 58 Lmrf Ultisols Serpentinite

Manganese (μg/g) II1A 43 26 2 43 66 20 58 Lmrf Ultisols Serpentinite

Niquel (μg/g) II1A 136 18 2 135 155 118 58 Lmrf Ultisols Serpentinite

Nitrogen (%) II1A 1.20 0.27 3 1.09 1.51 1.01 101 Swf Ultisols Tuffac. sandstone

II1A 1.60 0.62 7 1.65 2.28 0.70 94 Swf Ultisols Tuffac. sandstone

II1B 0.67 1 94 Swf Ultisols Tuffac. sandstone

II1C 0.65 0.40 5 0.49 1.09 0.28 94 Swf Ultisols Tuffac. sandstone Fresh

II1A 1.66 0.13 5 1.67 1.85 1.50 126 Ultisols Siltstone,sandstone Urban forest, RP

II1A 1.45 18.00 51 1.45 1.89 1.10 138 Wfs Ultisols Lava

II1A 1.12 0.24 2 1.12 1.33 0.89 58 Lmrf Ultisols Serpentinite

II1C 0.16 0.04 5 0.16 0.21 0.10 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 0.45 0.11 6 0.41 0.62 0.33 126 Ultisols Siltstone,sandstone Urban forest, RP

Phosphorus (mg/g) II1A 0.80 0.22 3 0.81 1.01 0.57 101 Swf Ultisols Tuffac. sandstone

II1A 1.31 0.55 7 1.15 2.12 0.55 94 Swf Ultisols Tuffac. sandstone

II1B 0.86 1 94 Swf Ultisols Tuffac. sandstone

II1C 0.37 0.16 5 0.34 0.63 0.22 94 Swf Ultisols Tuffac. sandstone Fresh

II1A 13.87 2.47 2 14.01 16.05 11.42 58 Lmrf Ultisols Serpentinite

Potassium (mg/g) II1A 7.67 2.08 3 7.00 10.00 6.00 101 Swf Ultisols Tuffac. sandstone

II1A 13.20 5.00 7 13.10 20.50 4.80 94 Swf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

306

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1B 10.84 1 94 Swf Ultisols Tuffac. sandstone

II1C 7.70 8.80 5 3.60 23.40 3.10 94 Swf Ultisols Tuffac. sandstone

II1A 11.33 3.73 2 11.32 14.61 8.08 58 Lmrf Ultisols Serpentinite

Sulfur (%) II1A 0.18 0.04 5 0.17 0.26 0.16 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 0.04 0.02 5 0.04 0.07 0.01 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 0.05 0.01 6 0.05 0.07 0.04 126 Ultisols Siltstone,sandstone Urban forest, RP

Wood density (g/cc) II1C 0.49 0.02 20 0.49 0.53 0.46 115

II1C 0.51 0.01 5 0.50 0.53 0.50 126 Ultisols Siltstone,sandstone Urban forest, RP

Tabebuia rigida Aluminum (mg/g) II1A 0.17 1 93 Wflm Inceptisols Tuffac. sandstone

II1A 0.62 0.30 39 0.59 1.29 0.17 107 Rfs Inceptisols Tuffac. sandstone

II1B 0.08 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.06 1 93 Wflm Inceptisols Tuffac. sandstone

Ash (%) II1A 4.47 1 93 Wflm Inceptisols Tuffac. sandstone

II1A 3.60 0.60 26 3.63 4.74 2.38 107 Rfs Inceptisols Tuffac. sandstone

II1B 2.22 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.62 1 93 Wflm Inceptisols Tuffac. sandstone

C/N II1A 38 1 93 Wflm Inceptisols Tuffac. sandstone

II1A 34 5 39 34 46 23 107 Rfs Inceptisols Tuffac. sandstone

II1B 146 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 460 1 93 Wflm Inceptisols Tuffac. sandstone

Calcium (mg/g) II1A 2.92 1 93 Wflm Inceptisols Tuffac. sandstone

II1A 2.23 1.09 39 1.89 6.91 1.15 107 Rfs Inceptisols Tuffac. sandstone

II1B 2.31 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.64 1 93 Wflm Inceptisols Tuffac. sandstone

Carbon (%) II1A 46 2 39 47 49 42 107 Rfs Inceptisols Tuffac. sandstone

Iron (mg/g) II1A 0.11 1 93 Wflm Inceptisols Tuffac. sandstone

II1A 0.59 0.38 39 0.52 2.34 0.17 107 Rfs Inceptisols Tuffac. sandstone

II1B 0.07 1 93 Wflm Inceptisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

307

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1C 0.06 1 93 Wflm Inceptisols Tuffac. sandstone

Magnesium (mg/g) II1A 3.09 1 93 Wflm Inceptisols Tuffac. sandstone

II1A 2.23 0.43 39 2.24 3.09 1.32 107 Wflm Inceptisols Tuffac. sandstone

II1B 1.50 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.41 1 93 Wflm Inceptisols Tuffac. sandstone

Manganese (mg/g) II1A 0.12 1 93 Wflm Inceptisols Tuffac. sandstone

II1A 0.03 0.03 39 0.03 0.19 0.02 107 Rfs Inceptisols Tuffac. sandstone

II1B 0.42 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.05 1 93 Wflm Inceptisols Tuffac. sandstone

Nitrogen (%) II1A 1.15 1 93 Wflm Inceptisols Tuffac. sandstone

II1A 1.41 0.22 39 1.40 1.94 0.94 107 Rfs Inceptisols Tuffac. sandstone

II1B 0.31 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.10 1 93 Wflm Inceptisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.67 1 93 Wflm Inceptisols Tuffac. sandstone

II1A 0.73 0.23 39 0.71 1.41 0.40 107 Rfs Inceptisols Tuffac. sandstone

II1B 0.50 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.14 1 93 Wflm Inceptisols Tuffac. sandstone

Potassium (mg/g) II1A 8.74 1 93 Wflm Inceptisols Tuffac. sandstone

II1A 7.05 3.92 39 5.36 20.52 3.04 107 Rfs Inceptisols Tuffac. sandstone

II1B 3.21 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 1.04 1 93 Wflm Inceptisols Tuffac. sandstone

Wood density (g/cc) II1B 0.56 0.05 23 0.56 0.67 0.46 97 Wfs Inceptisols Tuffac. sandstone

Tabebuia spp. Calcium (mg/g) II1A 20.26 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 19.28 0.15 2 19.28 19.39 19.18 149 Sdf Mollisols Alluvial deposits Guanica Forest

Magnesium (mg/g) II1A 15.56 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 7.00 0.25 2 7.00 7.18 6.83 149 Sdf Mollisols Alluvial deposits Guanica Forest

Nitrogen (%) II1A 1.75 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 1.39 0.41 4 1.39 1.76 1.03 149 Sdf Mollisols Alluvial deposits Guanica Forest
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

308

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Phosphorus (mg/g) II1Ci 0.86 0.54 4 0.88 1.33 0.34 149 Sdf Mollisols Alluvial deposits Guanica Forest

Potassium (mg/g) II1A 17.67 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 20.08 0.60 2 20.08 20.50 19.66 149 Sdf Mollisols Alluvial deposits Guanica Forest

Tabonuco forest Aluminum (mg/g) II9A 1.01 0.40 4 0.98 1.52 0.55 153 Swf Ultisols Tuffac. sandstone

Ash (%) II9A 6.97 0.59 4 7.08 7.48 6.22 153 Swf Ultisols Tuffac. sandstone

C/N II9A 33 1 4 33 33 31 153 Swf Ultisols Tuffac. sandstone

Calcium (mg/g) II9A 9.14 1.30 4 9.41 10.37 7.35 153 Swf Ultisols Tuffac. sandstone

Carbon (%) II9A 51 1 4 51 53 50 153 Swf Ultisols Tuffac. sandstone

Iron (mg/g) II9A 0.23 0.12 4 0.18 0.41 0.16 153 Swf Ultisols Tuffac. sandstone

Magnesium (mg/g) II9A 2.98 0.36 4 3.07 3.30 2.47 153 Swf Ultisols Tuffac. sandstone

Manganese (mg/g) II9A 0.28 0.07 4 0.27 0.38 0.21 153 Swf Ultisols Tuffac. sandstone

Nitrogen (%) II9A 1.58 0.06 4 1.57 1.66 1.52 153 Swf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II9A 0.57 0.04 4 0.57 0.61 0.53 153 Swf Ultisols Tuffac. sandstone

Potassium (mg/g) II9A 7.41 0.59 4 7.39 8.11 6.75 153 Swf Ultisols Tuffac. sandstone

Sodium (mg/g) II9A 2.65 0.29 4 2.57 3.06 2.40 153 Swf Ultisols Tuffac. sandstone

Tabonuco Sec. Forest Aluminum (mg/g) II8A 5.26 4.76 11 3.13 14.59 1.25 133 Wfs Ultisols Tuffac. sandstone

II8B 1.53 1.30 9 1.18 4.37 0.39 133 Wfs Ultisols Tuffac. sandstone

II8C 1.92 1.55 11 1.60 6.13 0.68 133 Wfs Ultisols Tuffac. sandstone

II8D 20.90 6.20 9 22.39 27.84 11.07 133 Wfs Ultisols Tuffac. sandstone

Ash (%) II8A 11.78 4.36 11 9.85 20.17 8.14 133 Wfs Ultisols Tuffac. sandstone

II8B 4.43 2.29 9 3.54 8.56 2.70 133 Wfs Ultisols Tuffac. sandstone

II8C 3.44 2.07 11 2.78 9.20 2.07 133 Wfs Ultisols Tuffac. sandstone

II8D 25.85 5.66 8 28.42 30.93 14.48 133 Wfs Ultisols Tuffac. sandstone

C/N II8A 39 6 11 39 48 31 133 Wfs Ultisols Tuffac. sandstone

II8B 71 13 9 74 86 53 133 Wfs Ultisols Tuffac. sandstone

II8C 59 5 11 57 66 50 133 Wfs Ultisols Tuffac. sandstone

II8D 25 2 9 25 28 22 133 Wfs Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

309

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Calcium (mg/g) II8A 11.11 2.33 11 10.86 16.34 8.20 133 Wfs Ultisols Tuffac. sandstone

II8B 9.36 1.71 9 9.46 11.59 6.56 133 Wfs Ultisols Tuffac. sandstone

II8C 4.02 0.89 11 4.07 5.49 2.33 133 Wfs Ultisols Tuffac. sandstone

II8D 9.00 1.05 9 8.65 10.70 7.70 133 Wfs Ultisols Tuffac. sandstone

Carbon (%) II8A 49 3 11 50 52 43 133 Wfs Ultisols Tuffac. sandstone

II8B 51 1 9 51 53 48 133 Wfs Ultisols Tuffac. sandstone

II8C 54 3 11 55 56 48 133 Wfs Ultisols Tuffac. sandstone

II8D 40 4 9 40 47 36 133 Wfs Ultisols Tuffac. sandstone

Iron (mg/g) II8A 3.96 4.40 11 2.24 12.57 0.67 133 Wfs Ultisols Tuffac. sandstone

II8B 1.07 1.15 9 0.58 3.41 0.21 133 Wfs Ultisols Tuffac. sandstone

II8C 1.53 1.49 11 1.15 5.47 0.29 133 Wfs Ultisols Tuffac. sandstone

II8D 22.02 8.92 9 26.46 32.46 8.96 133 Wfs Ultisols Tuffac. sandstone

Magnesium (mg/g) II8A 1.99 0.39 11 2.06 2.49 1.48 133 Wfs Ultisols Tuffac. sandstone

II8B 1.47 0.32 9 1.52 2.00 1.06 133 Wfs Ultisols Tuffac. sandstone

II8C 1.39 0.29 11 1.35 1.81 0.86 133 Wfs Ultisols Tuffac. sandstone

II8D 1.97 0.27 9 1.98 2.48 1.47 133 Wfs Ultisols Tuffac. sandstone

Manganese (mg/g) II8A 1.00 0.20 11 1.06 1.34 0.75 133 Wfs Ultisols Tuffac. sandstone

II8B 0.49 0.15 9 0.48 0.71 0.32 133 Wfs Ultisols Tuffac. sandstone

II8C 0.16 0.07 11 0.14 0.32 0.09 133 Wfs Ultisols Tuffac. sandstone

II8D 1.47 0.27 9 1.46 1.91 1.00 133 Wfs Ultisols Tuffac. sandstone

Nitrogen (%) II8A 1.26 0.14 11 1.27 1.41 1.04 133 Wfs Ultisols Tuffac. sandstone

II8B 0.74 0.14 9 0.70 0.95 0.58 133 Wfs Ultisols Tuffac. sandstone

II8C 0.93 0.11 11 0.93 1.10 0.78 133 Wfs Ultisols Tuffac. sandstone

II8D 1.65 0.21 9 1.64 2.04 1.41 133 Wfs Ultisols Tuffac. sandstone

Phosphorus (mg/g) II8A 0.33 0.08 11 0.29 0.46 0.22 133 Wfs Ultisols Tuffac. sandstone

II8B 0.22 0.06 9 0.20 0.33 0.14 133 Wfs Ultisols Tuffac. sandstone

II8C 0.46 0.16 11 0.48 0.67 0.19 133 Wfs Ultisols Tuffac. sandstone

II8D 0.52 0.09 9 0.53 0.69 0.39 133 Wfs Ultisols Tuffac. sandstone

Potassium (mg/g) II8A 1.62 0.49 11 1.37 2.60 1.13 133 Wfs Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

310

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8B 1.17 0.44 9 1.03 1.98 0.72 133 Wfs Ultisols Tuffac. sandstone

II8C 2.90 1.09 11 3.35 4.13 0.62 133 Wfs Ultisols Tuffac. sandstone

II8D 1.30 0.22 9 1.24 1.80 1.08 133 Wfs Ultisols Tuffac. sandstone

Tectaria trifoliata Aluminum (mg/g) II6 1.05 1.16 9 0.75 4.01 0.11 135 Lmrf Ultisols Tuffac. sandstone

C/N II6 24 7 7 23 38 16 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II6 7.16 1.80 9 6.57 11.19 5.35 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II6 42 2 7 43 44 40 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II6 1.04 1.09 9 0.67 3.63 0.22 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II6 4.69 1.95 9 3.73 9.15 2.95 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II6 0.14 0.05 9 0.12 0.24 0.09 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II6 1.87 0.36 11 1.89 2.44 1.16 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II6 0.85 0.24 9 0.91 1.24 0.39 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II6 22.79 11.62 9 26.11 35.37 3.54 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II6 0.79 0.33 7 0.73 1.36 0.42 135 Lmrf Ultisols Tuffac. sandstone

Telypteris deltoidea C/N II6 30 1 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II6 39 1 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II6 1.29 1 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II6 0.41 1 135 Lmrf Ultisols Tuffac. sandstone

Terebraria recinosa Aluminum (μg/g) II1A 30 1 58 Lmrf Ultisols Serpentinite

II1A 113 1 58 Lmrf Ultisols Serpentinite

Calcium (mg/g) II1A 7.64 1 58 Lmrf Ultisols Serpentinite

II1A 10.15 1 58 Lmrf Ultisols Serpentinite

Cobalt (μg/g) II1A 0.79 1 58 Lmrf Ultisols Serpentinite

II1A 1.44 1 58 Lmrf Ultisols Serpentinite

Copper (μg/g) II1A 7 1 58 Lmrf Ultisols Serpentinite

II1A 14 1 58 Lmrf Ultisols Serpentinite
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

311

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Chromium (μg/g) II1A 2 1 58 Lmrf Ultisols Serpentinite

II1A 3 1 58 Lmrf Ultisols Serpentinite

Iron (μg/g) II1A 35 1 58 Lmrf Ultisols Serpentinite

II1A 135 1 58 Lmrf Ultisols Serpentinite

Magnesium (mg/g) II1A 35.05 1 58 Lmrf Ultisols Serpentinite

II1A 1.74 1 58 Lmrf Ultisols Serpentinite

Manganese (μg/g) II1A 18 1 58 Lmrf Ultisols Serpentinite

II1A 76 1 58 Lmrf Ultisols Serpentinite

Niquel (μg/g) II1A 54 1 58 Lmrf Ultisols Serpentinite

II1A 62 1 58 Lmrf Ultisols Serpentinite

Nitrogen (%) II1A 0.80 1 58 Lmrf Ultisols Serpentinite

II1A 0.90 1 58 Lmrf Ultisols Serpentinite

Phosphorus (mg/g) II1A 9.18 1 58 Lmrf Ultisols Serpentinite

II1A 10.62 1 58 Lmrf Ultisols Serpentinite

Potassium (mg/g) II1A 7.38 1 58 Lmrf Ultisols Serpentinite

II1A 17.42 1 58 Lmrf Ultisols Serpentinite

Terminalia ivorensis Ash (%) II8A 8.28 1 90 Swf Ultisols Tuffac. sandstone New

II8A 9.64 1 90 Swf Ultisols Tuffac. sandstone Old

II8A 9.97 2.08 6 9.99 13.08 7.43 90 Swf Ultisols Tuffac. sandstone

II8B 4.50 0.44 3 4.54 4.91 4.04 90 Swf Ultisols Tuffac. sandstone

II8C 9.59 1 90 Swf Ultisols Tuffac. sandstone Seeds

II8D 19.59 7.28 3 15.47 27.99 15.31 90 Swf Ultisols Tuffac. sandstone

Calcium (mg/g) II8A 19.95 1 90 Swf Ultisols Tuffac. sandstone New

II8A 18.98 1 90 Swf Ultisols Tuffac. sandstone Old

II8A 22.31 6.64 6 23.15 29.07 14.45 90 Swf Ultisols Tuffac. sandstone

II8B 11.15 2.93 3 9.94 14.49 9.02 90 Swf Ultisols Tuffac. sandstone

II8C 9.75 1 90 Swf Ultisols Tuffac. sandstone Seeds

II8D 12.05 3.94 3 10.19 16.58 9.38 90 Swf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

312

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II9A 13.30 2.78 22 13.86 17.79 7.84 139 Wfs Ultisols Tuffac. sandstone

II9A 11.59 2.53 22 11.07 17.93 8.45 139 Wfs Ultisols Tuffac. sandstone Other

II9B 8.13 1.36 5 7.96 10.16 6.40 139 Wfs Ultisols Tuffac. sandstone

II9C 5.05 1.23 6 4.90 7.30 3.59 139 Wfs Ultisols Tuffac. sandstone

II9D 7.75 1.29 6 7.76 9.55 5.66 139 Wfs Ultisols Tuffac. sandstone

Magnesium (mg/g) II8A 4.20 1 90 Swf Ultisols Tuffac. sandstone New

II8A 3.80 1 90 Swf Ultisols Tuffac. sandstone Old

II8A 4.39 0.49 6 4.34 5.14 3.66 90 Swf Ultisols Tuffac. sandstone

II8B 2.31 0.20 3 2.37 2.47 2.09 90 Swf Ultisols Tuffac. sandstone

II8C 2.53 1 90 Swf Ultisols Tuffac. sandstone Seeds

II8D 2.87 0.44 3 2.72 3.36 2.52 90 Swf Ultisols Tuffac. sandstone

II9A 4.68 1.01 22 4.42 6.99 3.40 139 Wfs Ultisols Tuffac. sandstone

II9A 3.31 1.13 22 2.87 6.49 2.08 139 Wfs Ultisols Tuffac. sandstone Other

II9B 2.43 0.71 5 2.05 3.50 1.81 139 Wfs Ultisols Tuffac. sandstone

II9C 2.55 0.76 6 2.45 3.63 1.72 139 Wfs Ultisols Tuffac. sandstone

II9D 2.79 0.23 6 2.74 3.21 2.58 139 Wfs Ultisols Tuffac. sandstone

Nitrogen (%) II8A 1.13 1 90 Swf Ultisols Tuffac. sandstone New

II8A 1.16 1 90 Swf Ultisols Tuffac. sandstone Old

II8A 1.13 0.17 6 1.16 1.34 0.90 90 Swf Ultisols Tuffac. sandstone

II8B 0.64 0.04 3 0.65 0.68 0.60 90 Swf Ultisols Tuffac. sandstone

II8C 1.17 1 90 Swf Ultisols Tuffac. sandstone Seeds

II8D 1.21 0.04 3 1.22 1.25 1.17 90 Swf Ultisols Tuffac. sandstone

II9A 1.17 0.30 22 1.04 1.97 0.78 139 Wfs Ultisols Tuffac. sandstone

II9A 1.14 0.28 23 0.98 1.62 0.72 139 Wfs Ultisols Tuffac. sandstone Other

II9B 0.48 0.09 10 0.50 0.60 0.32 139 Wfs Ultisols Tuffac. sandstone

II9C 1.52 0.47 11 1.67 2.06 0.74 139 Wfs Ultisols Tuffac. sandstone

II9D 1.28 0.31 11 1.28 1.58 0.50 139 Wfs Ultisols Tuffac. sandstone

Phosphorus (mg/g) II8A 0.19 1 90 Swf Ultisols Tuffac. sandstone New

II8A 0.42 1 90 Swf Ultisols Tuffac. sandstone Old
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

313

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II8A 0.35 0.12 6 0.36 0.50 0.19 90 Swf Ultisols Tuffac. sandstone

II8B 0.22 0.05 3 0.23 0.27 0.16 90 Swf Ultisols Tuffac. sandstone

II8C 0.60 1 90 Swf Ultisols Tuffac. sandstone Seeds

II8D 0.51 0.01 3 0.50 0.52 0.50 90 Swf Ultisols Tuffac. sandstone

II9A 0.48 0.18 22 0.42 0.99 0.18 139 Wfs Ultisols Tuffac. sandstone

II9A 0.44 0.15 22 0.41 0.72 0.17 139 Wfs Ultisols Tuffac. sandstone Other

II9B 0.22 0.04 5 0.23 0.27 0.16 139 Wfs Ultisols Tuffac. sandstone

II9C 0.79 0.30 6 0.78 1.19 0.48 139 Wfs Ultisols Tuffac. sandstone

II9D 0.71 0.13 6 0.69 0.88 0.56 139 Wfs Ultisols Tuffac. sandstone

Potassium (mg/g) II8A 1.74 1 90 Swf Ultisols Tuffac. sandstone New

II8A 1.82 1 90 Swf Ultisols Tuffac. sandstone Old

II8A 3.76 1.83 6 2.95 6.63 1.74 90 Swf Ultisols Tuffac. sandstone

II8B 2.07 0.98 3 1.98 3.09 1.14 90 Swf Ultisols Tuffac. sandstone

II8C 2.41 1 90 Swf Ultisols Tuffac. sandstone Seeds

II8D 2.65 1.28 3 2.66 3.92 1.37 90 Swf Ultisols Tuffac. sandstone

II9A 5.07 1.40 22 4.74 8.39 3.20 139 Wfs Ultisols Tuffac. sandstone

II9A 5.20 1.44 22 4.87 8.29 3.31 139 Wfs Ultisols Tuffac. sandstone Other

II9B 3.60 0.87 5 3.50 4.85 2.57 139 Wfs Ultisols Tuffac. sandstone

II9C 5.90 1.32 6 5.65 8.03 4.02 139 Wfs Ultisols Tuffac. sandstone

II9D 4.60 1.48 6 4.98 5.95 2.16 139 Wfs Ultisols Tuffac. sandstone

Ternstroemia stahlii Aluminum (mg/kg) II1A 51 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 56 1 102 Smf Limestone Tuffac. sandstone Old

II1A 26 1 102 Smf Limestone Tuffac. sandstone Young

Ash (%) II1A 5.85 1 149 Sdf Mollisols Alluvial deposits Susua forest

Calcium (mg/g) II1A 8.69 1 102 Smf Limestone Tuffac. sandstone Old

II1A 2.72 1 102 Smf Limestone Tuffac. sandstone Young

II1A 9.71 1 149 Sdf Mollisols Alluvial deposits Susua forest

Cobalt (μg/g) II1A 0.80 1 102 Smf Limestone Tuffac. sandstone Old
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

314

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1A 0.50 1 102 Smf Limestone Tuffac. sandstone Young

Iron (mg/kg) II1A 87 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 53 1 102 Smf Limestone Tuffac. sandstone Old

II1A 24 1 102 Smf Limestone Tuffac. sandstone Young

Magnesium (mg/g) II1A 2.67 1 102 Smf Limestone Tuffac. sandstone Old

II1A 1.78 1 102 Smf Limestone Tuffac. sandstone Young

II1A 4.67 1 149 Sdf Mollisols Alluvial deposits Susua forest

Manganese (mg/kg) II1A 22 1 149 Sdf Mollisols Alluvial deposits Susua forest

II1A 13 1 102 Smf Limestone Tuffac. sandstone Old

II1A 11 1 102 Smf Limestone Tuffac. sandstone Young

Nickel (μg/g) II1A 11 1 102 Smf Limestone Tuffac. sandstone Old

II1A 18 1 102 Smf Limestone Tuffac. sandstone Young

Nitrogen (%) II1A 0.74 1 102 Smf Limestone Tuffac. sandstone Old

II1A 1.04 1 102 Smf Limestone Tuffac. sandstone Young

II1A 0.83 1 149 Sdf Mollisols Alluvial deposits Susua forest

Phosphorus (mg/g) II1A 0.32 1 102 Smf Limestone Tuffac. sandstone Old

II1A 0.81 1 102 Smf Limestone Tuffac. sandstone Young

II1A 0.34 1 149 Sdf Mollisols Alluvial deposits Susua forest

Potassium (mg/g) II1A 7.62 1 102 Smf Limestone Tuffac. sandstone Old

II1A 12.29 1 102 Smf Limestone Tuffac. sandstone Young

II1A 7.03 1 149 Sdf Mollisols Alluvial deposits Susua forest

Zinc (μg/g) II1A 5 1 149 Sdf Mollisols Alluvial deposits Susua forest

Tetragastris balsamifera Calcium (mg/g) II1A 5.67 1.45 3 6.11 6.86 4.05 135 Lmrf Ultisols Tuffac. sandstone

II1B 5.54 2.92 3 6.56 7.82 2.83 135 Lmrf Ultisols Tuffac. sandstone

II1C 5.94 2.72 3 7.05 7.90 2.83 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II1A 1.66 0.14 3 1.67 1.79 1.52 135 Lmrf Ultisols Tuffac. sandstone

II1B 1.01 0.22 3 1.06 1.20 0.77 135 Lmrf Ultisols Tuffac. sandstone

II1C 1.10 0.17 3 1.14 1.24 0.92 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

315

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Nitrogen (%) II1A 1.40 0.04 3 1.40 1.44 1.37 135 Lmrf Ultisols Tuffac. sandstone

II1B 0.48 0.12 3 0.54 0.56 0.35 135 Lmrf Ultisols Tuffac. sandstone

 II1C 0.50 0.11 3 0.56 0.57 0.37 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.67 0.00 3 0.67 0.67 0.67 135 Lmrf Ultisols Tuffac. sandstone

II1B 0.33 0.11 3 0.34 0.44 0.22 135 Lmrf Ultisols Tuffac. sandstone

II1C 0.34 0.08 3 0.37 0.41 0.25 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II1A 7.20 0.50 3 7.06 7.75 6.78 135 Lmrf Ultisols Tuffac. sandstone

II1B 4.53 1.26 3 4.33 5.88 3.38 135 Lmrf Ultisols Tuffac. sandstone

II1C 5.07 0.74 3 4.91 5.89 4.43 135 Lmrf Ultisols Tuffac. sandstone

Thespesia grandiflora Ash (%) II1A 9.34 0.48 6 9.23 10.00 8.84 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 1.77 0.62 6 1.46 2.58 1.26 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 9.32 2.17 5 8.31 11.76 7.02 126 Ultisols Siltstone,sandstone Urban forest, RP

C/N II1A 24 1 6 24 26 23 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 212 92 6 200 321 102 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 58 13 5 52 72 43 126 Ultisols Siltstone,sandstone Urban forest, RP

Carbon (%) II1A 53 0 6 53 54 53 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 55 0 6 55 55 54 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 53 2 5 52 56 51 126 Ultisols Siltstone,sandstone Urban forest, RP

Nitrogen (%) II1A 2.21 0.08 6 2.22 2.28 2.07 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 0.31 0.15 6 0.28 0.54 0.17 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 0.95 0.21 5 1.02 1.20 0.71 126 Ultisols Siltstone,sandstone Urban forest, RP

Sulfur (%) II1A 0.27 0.02 6 0.27 0.30 0.24 126 Ultisols Siltstone,sandstone Urban forest, RP

II1C 0.03 0.01 6 0.03 0.05 0.02 126 Ultisols Siltstone,sandstone Urban forest, RP

II1Ci 0.11 0.04 5 0.12 0.15 0.08 126 Ultisols Siltstone,sandstone Urban forest, RP

Wood density (g/cc) II1C 0.49 0.00 5 0.51 0.52 0.43 126 Ultisols Siltstone,sandstone Urban forest, RP

Thelypteris deltoidea Aluminum (mg/g) II6 0.97 0.81 38 0.71 3.16 0.04 135 Lmrf Ultisols Tuffac. sandstone

C/N II6 33 4 20 33 38 20 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

316

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Calcium (mg/g) II6 3.76 0.87 38 3.75 5.66 1.90 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II6 41 1 20 41 43 38 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II6 0.81 0.83 38 0.47 3.30 0.05 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II6 3.80 0.60 38 3.74 5.82 2.85 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II6 0.30 0.50 8 0.18 3.18 0.06 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II6 1.21 0.21 51 1.22 1.97 0.71 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II6 0.57 0.18 38 0.53 1.15 0.30 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II6 6.62 2.40 37 6.54 14.15 2.66 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II6 0.47 0.14 20 0.44 0.78 0.29 135 Lmrf Ultisols Tuffac. sandstone

Thouinia striata Calcium (mg/g) II1A 3.12 1 104 Sdf Mollisols Alluvial deposits New

II1A 8.07 4.08 7 7.17 13.63 0.94 104 Sdf Mollisols Alluvial deposits Mature

II1A 7.41 4.16 15 6.79 13.63 0.94 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 15.48 1 104 Sdf Mollisols Alluvial deposits

II1Ci 15.48 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Magnesium (mg/g) II1A 1.11 1 104 Sdf Mollisols Alluvial deposits New

II1A 1.22 0.41 7 1.31 1.75 0.43 104 Sdf Mollisols Alluvial deposits Mature

II1A 1.20 0.38 15 1.28 1.75 0.43 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 2.36 1 104 Sdf Mollisols Alluvial deposits

II1Ci 2.36 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Nitrogen (%) II1A 1.88 1 104 Sdf Mollisols Alluvial deposits New

II1A 1.86 0.17 6 1.86 2.12 1.65 104 Sdf Mollisols Alluvial deposits Mature

II1A 1.85 0.15 16 1.85 2.12 1.65 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 0.75 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Phosphorus (mg/g) II1A 0.87 1 104 Sdf Mollisols Alluvial deposits New

II1A 0.80 0.10 7 0.79 1.01 0.67 104 Sdf Mollisols Alluvial deposits Mature

II1A 0.81 0.09 16 0.80 1.01 0.67 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 0.39 1 104 Sdf Mollisols Alluvial deposits

II1Ci 0.39 1 149 Sdf Mollisols Alluvial deposits Guanica Forest
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

317

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Potassium (mg/g) II1A 11.88 1 104 Sdf Mollisols Alluvial deposits New

II1A 10.34 0.75 7 10.19 12.16 9.35 104 Sdf Mollisols Alluvial deposits Mature

II1A 10.53 0.88 16 10.21 12.16 9.35 149 Sdf Mollisols Alluvial deposits Guanica Forest

II1Ci 9.50 1 104 Sdf Mollisols Alluvial deposits

II1Ci 9.50 1 149 Sdf Mollisols Alluvial deposits Guanica Forest

Torralbasia cuneifolia Aluminum (mg/g) II1B 0.11 1 93 Wflm Inceptisols Tuffac. sandstone

Ash (%) II1B 2.76 1 93 Wflm Inceptisols Tuffac. sandstone

C/N II1B 106 1 93 Wflm Inceptisols Tuffac. sandstone

Calcium (mg/g) II1B 8.65 1 93 Wflm Inceptisols Tuffac. sandstone

Iron (mg/g) II1B 0.12 1 93 Wflm Inceptisols Tuffac. sandstone

Magnesium (mg/g) II1B 0.95 1 93 Wflm Inceptisols Tuffac. sandstone

Manganese (mg/g) II1B 0.01 1 93 Wflm Inceptisols Tuffac. sandstone

Nitrogen (%) II1B 0.42 1 93 Wflm Inceptisols Tuffac. sandstone

Phosphorus (mg/g) II1B 0.53 1 93 Wflm Inceptisols Tuffac. sandstone

Potassium (mg/g) II1B 2.90 1 93 Wflm Inceptisols Tuffac. sandstone

Trema micranthum Ash (%) II2A 9.37 1 94 Swf Ultisols Tuffac. sandstone

II2B 3.48 1 94 Swf Ultisols Tuffac. sandstone

II2C 2.12 1 94 Swf Ultisols Tuffac. sandstone

Nitrogen (%) II2A 3.63 1 94 Swf Ultisols Tuffac. sandstone

II2B 0.72 1 94 Swf Ultisols Tuffac. sandstone

II2C 0.51 1 94 Swf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II2A 2.14 1 94 Swf Ultisols Tuffac. sandstone

II2B 0.70 1 94 Swf Ultisols Tuffac. sandstone

II2C 0.37 1 94 Swf Ultisols Tuffac. sandstone

Potassium (mg/g) II2A 17.93 1 94 Swf Ultisols Tuffac. sandstone

II2B 3.35 1 94 Swf Ultisols Tuffac. sandstone

II2C 3.50 1 94 Swf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

318

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Trichilia palida Ash (%) II1A 9.71 1 71 Swf Ultisols Tuffac. sandstone

Carbon (%) II1A 52 1 71 Swf Ultisols Tuffac. sandstone

Nitrogen (%) II1A 1.42 1 71 Swf Ultisols Tuffac. sandstone

Sulfur (%) II1A 0.29 1 71 Swf Ultisols Tuffac. sandstone

Trichipteris borinquena Aluminum (mg/g) II6 0.88 0.99 4 0.84 1.82 0.03 135 Lmrf Ultisols Tuffac. sandstone

C/N II6 29 7 13 28 38 17 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II6 4.45 2.53 4 3.49 8.10 2.72 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II6 44 2 13 43 47 42 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II6 1.15 0.82 4 1.16 1.88 0.42 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II6 6.07 1.62 4 6.12 7.55 4.48 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II6 0.12 0.01 4 0.12 0.13 0.11 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II6 1.63 0.42 14 1.55 2.48 1.19 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II6 0.69 0.36 4 0.60 1.15 0.40 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II6 10.42 9.27 4 7.94 22.63 3.17 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II6 0.39 0.18 13 0.33 0.90 0.22 135 Lmrf Ultisols Tuffac. sandstone

Trichipteris procera Aluminum (mg/g) II6 2.59 1 135 Lmrf Ultisols Tuffac. sandstone

C/N II6 37 1 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II6 2.65 1 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II6 41 1 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II6 0.09 1 135 Lmrf Ultisols Tuffac. sandstone

Magnesium (mg/g) II6 2.71 1 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II6 0.13 1 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II6 1.11 1 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II6 0.83 1 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II6 16.48 1 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II6 0.27 1 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

319

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Trichostigma octandrum C/N II4 46 1 135 Lmrf Ultisols Tuffac. sandstone

Carbon (%) II4 45 1 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II4 0.96 1 135 Lmrf Ultisols Tuffac. sandstone

Sulfur (%) II4 0.15 1 135 Lmrf Ultisols Tuffac. sandstone

Turpinia occidentalis Ash (%) II1A 5.41 0.40 2 5.41 5.69 5.12 101 Swf Ultisols Tuffac. sandstone

II1B 2.30 0.13 2 2.30 2.39 2.21 101 Swf Ultisols Tuffac. sandstone

Calcium (mg/g) II1A 11.35 1 101 Swf Ultisols Tuffac. sandstone

II1B 7.05 2.59 2 7.05 8.89 5.22 101 Swf Ultisols Tuffac. sandstone

Nitrogen (%) II1A 1.46 0.10 2 1.46 1.53 1.39 101 Swf Ultisols Tuffac. sandstone

II1B 0.29 0.11 2 0.29 0.36 0.21 101 Swf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II1A 0.85 0.12 2 0.85 0.93 0.76 101 Swf Ultisols Tuffac. sandstone

II1B 0.25 0.13 2 0.25 0.34 0.16 101 Swf Ultisols Tuffac. sandstone

Potassium (mg/g) II1A 13.34 0.90 2 13.34 13.98 12.70 101 Swf Ultisols Tuffac. sandstone

II1B 6.75 0.45 2 6.75 7.07 6.44 101 Swf Ultisols Tuffac. sandstone

Urera baccifera Aluminum (mg/g) II2 0.21 1 135 Lmrf Ultisols Tuffac. sandstone

II2A 0.07 0.01 4 0.06 0.09 0.06 135 Lmrf Ultisols Tuffac. sandstone

II2C 0.05 0.03 5 0.04 0.08 0.02 135 Lmrf Ultisols Tuffac. sandstone

II2H 0.09 0.00 2 0.09 0.09 0.08 135 Lmrf Ultisols Tuffac. sandstone

Calcium (mg/g) II2 19.81 1 135 Lmrf Ultisols Tuffac. sandstone

II2A 37.30 12.30 4 38.00 51.25 21.952 135 Lmrf Ultisols Tuffac. sandstone

II2C 8.36 2.42 5 8.13 11.02 4.70 135 Lmrf Ultisols Tuffac. sandstone

II2H 15.11 2.21 2 15.11 16.67 13.55 135 Lmrf Ultisols Tuffac. sandstone

Iron (mg/g) II2 0.22 1 135 Lmrf Ultisols Tuffac. sandstone

II2A 0.08 0.01 4 0.08 0.10 0.06 135 Lmrf Ultisols Tuffac. sandstone

II2C 0.02 0.01 5 0.02 0.04 0.01 135 Lmrf Ultisols Tuffac. sandstone

II2H 0.13 0.09 2 0.13 0.19 0.06 135 Lmrf Ultisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

320

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Magnesium (mg/g) II2 6.49 1 135 Lmrf Ultisols Tuffac. sandstone

II2A 10.79 4.90 4 9.47 17.52 6.68 135 Lmrf Ultisols Tuffac. sandstone

II2C 5.50 1.05 5 5.77 6.66 3.82 135 Lmrf Ultisols Tuffac. sandstone

II2H 7.24 1.01 2 7.24 7.95 6.52 135 Lmrf Ultisols Tuffac. sandstone

Manganese (mg/g) II2 0.18 1 135 Lmrf Ultisols Tuffac. sandstone

II2A 0.30 0.22 4 0.23 0.61 0.13 135 Lmrf Ultisols Tuffac. sandstone

II2C 0.13 0.03 5 0.13 0.15 0.09 135 Lmrf Ultisols Tuffac. sandstone

II2H 0.17 0.04 2 0.17 0.20 0.15 135 Lmrf Ultisols Tuffac. sandstone

Nitrogen (%) II2 1.27 1 135 Lmrf Ultisols Tuffac. sandstone

II2A 2.69 0.51 4 2.82 3.14 1.96 135 Lmrf Ultisols Tuffac. sandstone

II2C 0.62 0.05 5 0.60 0.71 0.57 135 Lmrf Ultisols Tuffac. sandstone

II2H 1.36 0.01 2 1.36 1.36 1.35 135 Lmrf Ultisols Tuffac. sandstone

Phosphorus (mg/g) II2 4.34 1 135 Lmrf Ultisols Tuffac. sandstone

II2A 2.90 0.71 4 2.892 3.73 2.07 135 Lmrf Ultisols Tuffac. sandstone

II2C 1.12 0.56 5 0.80 2.07 0.77 135 Lmrf Ultisols Tuffac. sandstone

II2H 1.43 0.23 2 1.43 1.59 1.27 135 Lmrf Ultisols Tuffac. sandstone

Potassium (mg/g) II2 50.59 1 135 Lmrf Ultisols Tuffac. sandstone

II2A 41.05 9.15 4 44.73 47.29 27.45 135 Lmrf Ultisols Tuffac. sandstone

II2C 31.00 2.58 5 31.75 33.52 26.87 135 Lmrf Ultisols Tuffac. sandstone

II2H 32.82 0.70 2 32.82 33.31 32.33 135 Lmrf Ultisols Tuffac. sandstone

Xylosma schwaneckeanum Aluminum (mg/g) II1A 9.34 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 3.42 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 2.70 1 93 Wflm Inceptisols Tuffac. sandstone

Ash (%) II1B 4.90 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 1.26 1 93 Wflm Inceptisols Tuffac. sandstone

C/N II1B 69 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 692 1 93 Wflm Inceptisols Tuffac. sandstone

Calcium (mg/g) II1A 18.37 1 93 Wflm Inceptisols Tuffac. sandstone
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

321

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

II1B 5.12 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 1.67 1 93 Wflm Inceptisols Tuffac. sandstone

Iron (mg/kg) II1A 1530 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 70 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 4 1 93 Wflm Inceptisols Tuffac. sandstone

Magnesium (mg/g) II1A 0.22 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 1.12 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.37 1 93 Wflm Inceptisols Tuffac. sandstone

Manganese (mg/g) II1A 0.49 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.21 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.17 1 93 Wflm Inceptisols Tuffac. sandstone

Nitrogen (%) II1B 0.63 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.07 1 93 Wflm Inceptisols Tuffac. sandstone

Phosphorus (mg/g) II1A 1.62 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 0.76 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 0.05 1 93 Wflm Inceptisols Tuffac. sandstone

Potassium (mg/g) II1A 11.32 1 93 Wflm Inceptisols Tuffac. sandstone

II1B 11.50 1 93 Wflm Inceptisols Tuffac. sandstone

II1C 2.05 1 93 Wflm Inceptisols Tuffac. sandstone

Young Secondary forest Calcium (mg/g) II8A 9.76 2.11 10 9.05 12.91 7.54 98 Lmrf Ultisols Tuffac. sandstone Mameyes Basin

II8B 9.48 2.75 10 8.74 15.32 6.15 98 Lmrf Ultisols Tuffac. sandstone Mameyes Basin

II8D 9.37 2.36 10 8.47 14.63 6.76 98 Lmrf Ultisols Tuffac. sandstone Mameyes Basin

Magnesium (mg/g) II8A 3.13 0.63 10 2.88 4.30 2.31 98 Lmrf Ultisols Tuffac. sandstone Mameyes Basin

II8B 2.32 0.77 10 2.25 3.59 1.36 98 Lmrf Ultisols Tuffac. sandstone Mameyes Basin

II8D 3.55 0.96 10 3.26 5.46 2.53 98 Lmrf Ultisols Tuffac. sandstone Mameyes Basin

Phosphorus (mg/g) II8A 0.62 0.12 10 0.59 0.83 0.48 98 Lmrf Ultisols Tuffac. sandstone Mameyes Basin

II8B 0.43 0.05 10 0.45 0.50 0.35 98 Lmrf Ultisols Tuffac. sandstone Mameyes Basin

II8D 0.74 0.13 10 0.69 0.96 0.61 98 Lmrf Ultisols Tuffac. sandstone Mameyes Basin
Table 8. Element concentration of plant parts by species. Tissue or plant part codes are in Table 2. Source code are in appendix 1. Life zone 

codes are Table 1.

322

Species Element Tissue Mean Sd n Median Max Min Source Life Soil order Geology and/or Notes Species

zone parent material #

Potassium (mg/g) II8A 3.55 1.04 10 3.20 5.57 2.34 98 Lmrf Ultisols Tuffac. sandstone Mameyes Basin

II8B 2.86 0.78 10 2.98 3.81 1.10 98 Lmrf Ultisols Tuffac. sandstone Mameyes Basin

II8D 2.90 0.76 10 2.53 4.08 1.98 98 Lmrf Ultisols Tuffac. sandstone Mameyes Basin

Zanthoxylum flavum Wood density (g/cc) II1C 0.78 0.08 4 0.70 0.50 0.71 115

Zanthoxylum martinicense Wood density (g/cc) II1B 0.43 0.03 3 0.44 0.45 0.40 97 Wfs Inceptisols Tuffac. sandstone
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

333

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Alluvial deposits Mollisols Sdf Calcium (mg/g) 0--5 4.55 1.60 2 4.55 5.68 3.42 1 Disturb surface

0--5 5.44 0.88 8 5.70 6.24 3.60 1 Disturb plot

0--5 5.12 1.37 14 5.10 7.25 2.58 1 Undisturb plot

5--15 5.16 0.88 8 4.91 6.95 4.20 1 Disturb plot

5--15 5.18 1.17 14 4.99 7.13 2.88 1 Undisturb plot

5--15 4.50 1 1 Undisturb deep

5--15 3.84 1.11 2 3.84 4.63 3.05 1 Undisturb surface

ECEC (cmol/kg-1) 0--5 29.32 12.43 2 29.32 38.12 8.00 1 Disturb surface

0--5 36.58 6.61 8 38.08 42.66 24.47 1 Disturb plot

0--5 35.13 11.36 14 32.42 53.00 16.38 1 Undisturb plot

5--15 34.48 7.17 8 33.61 47.99 27.01 1 Disturb plot

5--15 35.41 9.38 14 36.12 50.20 17.86 1 Undistwb plot

5--15 25.56 8.63 2 25.56 31.67 19.46 1 Undisturb surface

Loss on ignition (%) 0--5 71.21 10.23 20 73.07 84.86 46.12 1 Disturb plot

0--5 67.03 10.38 5 63.69 81.04 5.00 1 Disturb surface

0--5 64.62 13.15 38 63.15 87.30 42.33 1 Undisturb plot

5--15 75.93 9.85 20 78.56 87.69 57.46 1 Disturb plot

5--15 74.43 5.79 5 73.02 81.16 9.85 1 Disturb deep

5--15 69.24 13.34 40 69.07 93.18 32.17 1 Undisturb plot

5--15 69.20 7.89 4 70.03 77.76 58.95 1 Undisturb deep

5--15 53.58 11.94 5 51.21 73.38 41.13 1 Undisturb surface

Magnesium (mg/g) 0--5 0.96 0.26 8 0.99 1.34 0.63 1 Disturb plot

0--5 0.66 0.47 2 0.66 0.99 0.33 1 Disturb surface

0--5 1.00 0.54 14 0.76 2.03 0.37 1 Undisturb plot

5--15 0.89 0.37 8 0.76 1.46 0.53 1 Disturb plot

5--15 1.00 0.44 14 1.04 2.05 0.39 1 Undisturb plot
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

334

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

5--15 0.81 1 1 Undisturb deep

5--15 0.67 0.31 2 0.67 0.89 0.45 1 Undisturb surface

Nitrogen (%) 0--5 1.01 0.26 8 0.99 1.40 0.67 1 Disturb plot

0--5 1.14 0.31 5 1.26 1.43 0.17 1 Disturb surface

0--5 1.15 0.30 14 1.23 1.62 0.60 1 Undisturb plot

5--15 0.81 0.26 8 0.74 1.20 0.55 1 Disturb plot

5--15 0.94 0.17 5 0.96 1.09 0.26 1 Distwb deep

5--15 1.00 0.29 14 1.01 1.39 0.47 1 Undisturb plot

5--15 0.99 0.40 2 0.99 1.27 0.71 1 Undisturb deep

5--15 1.43 0.41 4 1.59 1.71 0.82 1 Undisturb surface

Organic matter (%) 0--5 19.63 7.58 20 19.68 36.79 8.56 1 Disturb plot

0--5 23.63 9.33 38 23.89 41.33 8.66 1 Undisturb plot

5--15 16.13 7.41 20 13.93 30.66 7.57 1 Disturb plot

5--15 15.51 4.15 5 17.24 18.42 8.36 1 Disturb deep

5--15 22.11 5.91 5 22.78 28.40 13.88 1 Disturb surface

5--15 19.64 8.05 39 19.80 37.65 6.71 1 Undisturb plot

5--15 18.34 6.40 4 17.86 26.48 11.16 1 Undisturb deep

5--15 29.98 8.75 5 32.24 38.09 15.02 1 Undisturb surface

pH (H2O) 0--5 7.66 0.18 8 7.65 7.95 7.39 1 Disturb plot

0--5 7.65 0.17 33 7.66 7.99 7.28 1 Undisturb plot

5--15 7.68 0.16 16 7.70 7.87 7.37 1 Disturb plot

5--15 7.70 0.17 33 7.69 7.97 7.27 1 Undisturb plot

Phosphorus (mg/kg) 0--5 535 141 20 509 902 363 1 Disturb plot

0--5 590 130 38 600 1010 370 1 Undisturb plot

5--15 490 140 20 480 930 330 1 Disturb plot

5--15 460 80 5 460 560 140 1 Distwb deep
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

335

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

5--15 590 150 5 650 730 80 1 Disturb surface

5--15 550 120 40 540 850 340 1 Undisturb plot

5--15 610 100 4 650 670 460 1 Undisturb deep

5--15 600 70 5 620 690 510 1 Undisturb surface

Potassium (mg/g) 0--5 0.53 0.17 8 0.48 0.77 0.32 1 Disturb plot

0--5 0.45 0.21 14 0.39 0.93 0.12 1 Undisturb plot

5--15 0.50 0.18 8 0.54 0.74 0.28 1 Disturb plot

5--15 0.42 0.21 2 0.42 0.56 0.27 1 Disturb surface

5--15 0.47 0.19 14 0.46 0.88 0.01 1 Undisturb plot

5--15 0.42 1 1 Undisturb deep

5--15 0.31 0.17 2 0.31 0.43 0.19 1 Undisturb surface

Alluvial deposits Mollisols Sdf Loss on ignition (%) 0-5 21.52 1 2 Ewell site-grassy

0-5 21.18 1.55 4 21.27 22.88 19.29 2 Ewell site-quad 1-4

0-5 12.83 1 2 North forest

0-5 25.93 1 2 South forest

0-5 22.47 1 2 Eastern forest

0-5 34.59 1 2 West forest

5--10 20.46 1 2 Ewell site-grassy

5--15 19.13 1.18 4 19.02 20.41 18.06 2 Ewell site-quad 1-4

5--15 18.80 1 2 North forest

5--15 23.04 1 2 South forest

5--15 21.63 1 2 Eastern forest

5--10 29.76 1 2 West forest

Organic matter (%) 0-5 9.32 1 2 Ewell site-grassy

0-5 10.07 1.05 4 10.29 10.98 8.74 2 Ewell site-quad 1-4

0-5 11.34 1 2 North forest
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

336

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0-5 13.36 1 2 South forest

0-5 11.54 1 2 Eastern forest

0-5 21.25 1 2 West forest

5--10 8.57 1 2 Ewell site-grassy

5--10 8.72 0.89 4 8.58 9.91 7.81 2 Ewell site-quad 1-4

5--10 8.78 1 2 North forest

5--10 11.75 1 2 South forest

5--10 10.33 1 2 Eastern forest

5--10 17.39 1 2 West forest

pH (H2O) 0-5 7.54 1 2 Ewell site-grassy

0-5 7.55 0.11 4 7.56 7.67 7.41 2 Ewell site-quad 1-4

0-5 7.66 1 2 South forest

0-5 7.75 1 2 Eastern forest

0-5 7.67 1 2 West forest

5--10 7.44 1 2 Ewell site-grassy

5--10 7.58 0.10 4 7.58 7.68 7.48 2 Ewell site-quad 1-4

5--10 7.61 1 2 South forest

5--10 7.74 1 2 Eastern forest

5--10 7.68 1 2 West forest

Phosphorus (mg/kg) 0-5 11 1 2 Ewell site-grassy

0-5 11 1 4 10 12 10 2 Ewell site-quad 1-4

0-5 12 1 2 North forest

0-5 16 1 2 South forest

0-5 12 1 2 Eastern forest

0-5 19 1 2 West forest

5--10 10 1 2 Ewell site-grassy
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

337

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

5--10 9 1 4 9 10 8 2 Ewell site-quad 1-4

5--10 12 1 2 North forest

5--10 11 1 2 South forest

5--10 10 1 2 Eastern forest

5--10 18 1 2 West forest

Phosphorus-total (mg/g) 0-5 0.73 1 2 Ewell site-grassy

0-5 0.63 0.08 4 0.62 0.73 0.54 2 Ewell site-quad 1-4

0-5 0.93 1 2 North forest

0-5 0.79 1 2 South forest

0-5 0.64 1 2 Eastern forest

0-5 0.84 1 2 West forest

5--10 0.65 1 2 Ewell site-grassy

5--10 0.58 0.15 4 0.56 0.77 0.41 2 Ewell site-quad 1-4

5--10 0.66 1 2 North forest

5--10 0.68 1 2 South forest

5--10 0.60 1 2 Eastern forest

5--10 0.80 1 2 West forest

Alluvial deposits Mollisols Sdf Bulk density (g/cc) 0--4 0.86 0.07 4 0.84 0.95 0.81 3 Soil pit

6--16 0.88 0.14 7 0.80 1.03 0.72 3 Soil pit

18--63 1.21 0.13 6 1.23 1.35 0.99 3 Soil pit

Clay (%) 0-4 34 10 4 34 47 23 3 Soil pit

6--16 50 4 7 52 56 46 3 Soil pit

18--63 30 12 6 28 48 19 3 Soil pit

Loss on ignition (%) 0-4 3.12 1.33 4 3.04 4.81 1.59 3 Soil pit

6--16 6.98 1.94 7 6.98 9.70 4.40 3 Soil pit

18--63 1.41 1.69 6 0.69 4.61 0.24 3 Soil pit
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

338

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Moisture (%) 0-4 20.85 2.64 4 20.11 24.55 18.65 3 Soilpit

6--16 20.80 4.43 7 22.27 25.27 12.13 3 Soil pit

18--63 12.33 5.32 6 11.64 21.77 6.97 3 Soil pit

Nitrogen (%) 0-4 0.24 0.10 4 0.21 0.39 0.17 3 Soil pit

6--16 0.45 0.11 7 0.43 0.63 0.32 3 Soil pit

18--63 0.07 0.04 6 0.05 0.12 0.02 3 Soil pit

pH (H2O) 0-4 7.97 0.10 4 7.96 8.08 7.87 3 Soil pit

6--16 7.88 0.16 7 7.82 8.23 7.75 3 Soil pit

18--63 8.11 0.21 6 8.15 8.42 7.84 3 Soil pit

Phosphorus-total (mg/g) 0-4 0.28 0.16 4 0.24 0.51 0.14 3 Soil pit

6--16 0.36 0.06 7 0.38 0.42 0.24 3 Soil pit

18--63 0.17 0.07 6 0.16 0.30 0.08 3 Soil pit

Sand (%) 0-4 25 6 4 23 34 19 3 Soil pit

6--16 18 3 7 17 24 14 3 Soil pit

18--63 15 4 6 14 21 10 3 Soil pit

Silt (%) 0-4 41 9 4 37 55 35 3 Soil pit

6--16 32 5 7 34 37 23 3 Soil pit

18--63 55 16 6 58 71 31 3 Soil pit

Tuffaceous sandstone Inceptisols Lmwf Bulk density (g/cc) 0--24 0.57 0.09 11 0.59 0.70 0.32 4 Landslide-scarse veg.

0--24 0.52 0.10 11 0.49 0.69 0.32 4 Landslide-with veg.

0--21 0.62 0.32 10 0.53 1.51 0.39 4 Landslide-without veg.

Bulk density (g/cc) 0--22 1.09 0.23 6 0.99 1.43 0.88 5 Rio Icaco

0-23 1.22 0.11 6 1.19 1.39 1.10 5 Lanslide

0-23 0.89 0.14 12 0.88 1.11 0.71 5 Toro Willey top

0-23 0.78 0.17 28 0.83 1.05 0.38 5 Toro-186

Tuffaceous sandstone Ultisols Lmrf 0--23 0.97 0.13 23 0.97 1.22 0.71 5 Rio Grande-186
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

339

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Tuffaceous sandstone Inceptisols Lmrf 0-24 0.72 0.13 12 0.71 0.99 0.54 5 930-Mt. Briton

0-24 1.10 0.11 6 1.13 1.24 0.96 5 CS-2

0--24 1.07 0.17 6 1.09 1.28 0.77 5 Yunque road

0--25 1.05 0.37 6 1.08 1.63 0.56 5 West Fork top

0--25 1.01 0.20 6 0.98 1.37 0.83 5 West Fork nemoca

0--25 0.84 0.20 6 0.86 1.07 0.57 5 West Fork bottom

0--25 1.07 0.12 6 1.07 1.24 0.92 5 West Fork steep

Tuffaceous sandstone Ultisols Lmrf 0--25 1.11 0.28 6 1.05 1.44 0.82 5 CV-l

Tuffaceous sandstone Inceptisols Lmrf 0--25 0.78 0.18 12 0.75 1.02 0.54 5 Dwarf-l

0--25 0.94 0.27 6 0.94 1.24 0.61 5 Yunque road

Clay (%) 0--22 27 9 6 30 36 15 5 Rio Icaco

0--23 29 13 6 34 43 11 5 Lanslide

0--23 35 15 12 31 59 14 5 Toro Willey top

0--23 64 20 29 74 85 18 5 Toro-186

Tuffaceous sandstone Ultisols Lmrf 0--23 64 18 24 67 84 11 5 Rio Grande-186

Tuffaceous sandstone Inceptisols Lmrf 0--24 35 17 12 31 61 12 5 930-Mt. Britton

0--24 26 7 6 25 35 17 5 CS-2

0--24 28 15 6 25 47 10 5 Yunque road

0--25 27 10 6 25 45 15 5 West Fork top

0--25 41 13 6 43 57 24 5 West Fork nemoca

0--25 53 18 6 54 81 31 5 West Fork bottom

Tuffaceous sandstone Inceptisols Lmrf 0--25 31 12 6 28 48 18 5 West Fork steep

Tuffaceous sandstone Ultisols Lmrf 0--25 22 11 6 20 37 9 5 CV-l

Tuffaceous sandstone Inceptisols Lmrf 0--25 40 16 12 46 60 15 5 Dwarf-l

Lmrf 0--25 28 13 6 32 40 10 5 Yunque bath

Lmrf Organic matter (%) 0--22 3.03 1.34 6 2.55 4.74 1.50 5 Rio Icaco
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

340

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Lmrf 0--23 4.02 1.29 6 3.74 5.66 2.58 5 Lanslide

Lmrf 0--23 8.53 4.54 12 7.04 15.73 2.52 5 Toro Willey top

0--23 11.90 7.26 29 9.69 28.16 1.69 5 Toro-186

Tuffaceous sandstone Ultisols Lmrf 0--23 6.82 2.91 23 7.00 14.58 3.13 5 Rio Grande-186

Tuffaceous sandstone Inceptisols Lmrf 0--24 10.29 4.36 12 11.38 16.16 2.83 5 930-Mt. Britton

0--24 4.34 2.37 6 3.77 7.26 1.73 5 CS-2

0--24 8.74 3.56 6 7.92 14.76 5.04 5 Yunque road

0--25 12.50 8.46 6 10.25 23.08 3.03 5 West Fork top

0--25 12.28 4.81 6 13.02 17.66 4.66 5 West Fork nemoca

0--25 12.37 7.92 6 9.10 23.33 4.66 5 West Fork bottom

0--25 7.46 3.68 6 6.87 12.58 2.27 5 West Fork steep

Tuffaceous sandstone Ultisols Lmrf 0--25 5.07 2.49 6 4.99 9.54 2.56 5 CV-l

Tuffaceous sandstone Inceptisols Lmrf 0--25 12.82 7.90 12 12.86 27.67 2.16 5 Dwarf-l

0--25 10.02 5.86 6 9.52 18.21 3.60 5 Yunque bath

Sand (%) 0-22 57 4 6 57 63 51 5 Rio Icaco

0-23 58 15 6 54 78 44 5 Lanslide

0-23 40 16 12 47 57 11 5 Toro Willey top

0-23 15 19 29 6 65 1 5 Toro-186

Tuffaceous sandstone Ultisols Lmrf 0--23 15 15 24 13 75 1 5 Rio Grande-186

Tuffaceous sandstone Inceptisols Lmrf 0-24 32 19 12 27 70 7 5 930-Mt. Britton

0-24 53 7 6 56 61 40 5 CS-2

0--24 53 23 6 52 79 26 5 Yunque road

0--25 50 17 6 54 67 18 5 West Fork top

0--25 26 6 6 27 33 17 5 West Fork nemoca

0--25 13 9 6 11 26 2 5 West Fork bottom

0--25 43 13 6 40 59 28 5 West Fork steep
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

341

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Tuffaceous sandstone Ultisols Lmrf 0--25 58 10 6 61 72 46 5 CV-l

Tuffaceous sandstone Inceptisols Lmrf 0--25 24 13 12 22 46 10 5 Dwarf-l

0--25 55 16 6 47 82 42 5 Yunque bath

Silt (%) 0-22 16 9 6 13 30 7 5 Rio Icaco

0-23 13 2 6 13 18 11 5 Lanslide

0-23 27 12 12 31 44 5 5 Toro Willey top

0-23 22 7 29 20 43 13 5 Toro-186

Tuffaceous sandstone Ultisols Lmrf 0--23 21 10 22 19 58 12 5 Rio Grande-186

Tuffaceous sandstone Inceptisols Lmrf 0-24 34 12 12 32 52 19 5 930-Mt. Britton

0-24 21 6 6 20 30 15 5 CS-2

0--24 20 8 6 23 27 7 5 Yunque road

0--25 23 7 6 22 37 17 5 West Fork top

0--25 33 11 6 32 46 23 5 West Fork nemoca

0--25 34 10 6 36 43 16 5 West Fork bottom

0--25 26 6 6 24 39 23 5 West Fork steep

Tuffaceous sandstone Ultisols Lmrf 0--25 19 3 6 18 22 16 5 CV-l

Tuffaceous sandstone Inceptisols Lmrf 0--25 35 5 12 34 46 29 5 Dwarf-l

0--25 17 5 6 19 22 8 5 Yunque bath

Tuffaceous sandstone Ultisols Smf Aluminum (cmol/kg-l) 0-15 4.41 1.36 9 4.60 5.80 1.60 6 Cubuy pine

0-15 1.74 1.11 9 1.80 3.40 0.10 6 Cubuy secondary forest

0-15 4.01 1.12 9 4.20 5.30 1.70 6 El Verde

0-15 1.83 0.71 9 1.90 3.10 0.40 6 Guzmán pine

0-15 0.70 0.64 9 0.60 2.00 0.10 6 Guzmán secondary forest

0-15 0.46 0.28 9 0.30 1.00 0.20 6 Harvey-mahagony

15-30 3.10 1.44 9 3.00 4.80 0.60 6 Cubuy secondary forest

15-30 3.81 1.10 9 4.40 5.20 1.90 6 El Verde
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

342

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

15-30 2.37 0.69 9 2.40 3.30 1.30 6 Guzmán pine

15-30 1.29 0.54 9 1.20 2.30 0.70 6 Guzmán secondary forest

15-30 1.41 0.78 9 1.50 2.70 0.30 6 Harvey-mahagony

15-30 3.08 0.51 9 3.00 4.10 2.50 6 Cubuy pine

Aluminum-total (%) 0-15 8.57 2.85 9 9.46 9.98 1.02 6 Guzmán pine

0-15 9.99 1.00 9 9.93 11.19 8.53 6 Harvey-mahagony

0-15 10.81 1.15 9 11.22 12.19 9.33 6 El Verde

0-15 13.04 2.96 9 14.38 17.39 8.64 6 Guzmán secondary forest

0-15 6.66 1.08 9 6.75 8.59 5.18 6 Cubuy secondary forest

0-15 8.78 1.04 9 9.12 10.13 7.08 6 Cubuy pine

15-30 10.56 1.08 9 10.42 12.74 9.21 6 Guzmán pine

15-30 11.25 1.45 9 11.53 13.06 8.47 6 Harvey-mahagony

15-30 12.14 1.86 9 11.97 14.92 9.37 6 El Verde

15-30 13.61 3.55 9 14.37 18.02 7.53 6 Guzmán secondary forest

15-30 6.81 0.80 9 7.04 7.72 5.59 6 Cubuy secondary forest

15-30 9.19 1.06 9 9.32 10.87 7.77 6 Cubuy pine

Bulk density (g/cc) 0-15 0.90 0.09 9 0.92 1.03 0.78 6 Guzmán pine

0-15 0.89 0.07 9 0.92 0.98 0.78 6 Harvey-mahagony

0-15 0.71 0.08 9 0.68 0.88 0.61 6 El Verde

0-15 0.85 0.12 9 0.91 0.96 0.59 6 Guzmán secondary forest

0-15 0.79 0.10 9 0.84 0.88 0.61 6 Cubuy secondary forest

0-15 0.76 0.08 9 0.76 0.89 0.65 6 Cubuy pine

15-30 0.93 0.09 9 0.94 1.04 0.74 6 Guzmán pine

15-30 0.91 0.06 9 0.92 1.00 0.81 6 Harvey-mahagony

15-30 0.88 0.06 9 0.88 0.94 0.76 6 El Verde

15-30 0.84 0.10 9 0.86 0.99 0.66 6 Guzmán secondary forest
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

343

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

15-30 0.90 0.05 9 0.88 0.98 0.85 6 Cubuy secondary forest

15-30 1.05 0.06 9 1.04 1.14 0.94 6 Cubuy pine

Calcium (mg/kg) 0-15 470 115 9 457 670 325 6 Guzmán pine

0-15 877 184 9 838 1227 608 6 Harvey-mahagony

0-15 389 108 9 422 554 227 6 El Verde

0-15 1023 289 9 964 1546 637 6 Guzmán secondary forest

0-15 810 577 9 584 2080 325 6 Cubuy secondary forest

0-15 300 110 9 268 487 138 6 Cubuy pine

15-30 354 79 9 333 472 233 6 Guzmán pine

15-30 615 257 9 555 1126 349 6 Harvey-mahagony

15-30 218 102 9 171 455 131 6 El Verde

15-30 756 443 9 765 1355 153 6 Guzmán secondary forest

15-30 389 380 9 278 1302 100 6 Cubuy secondary forest

15-30 284 103 9 256 464 166 6 Cubuy pine

Calcium-total (mg/kg) 0-15 1369 511 9 1148 2419 719 6 Guzmán pine

0-15 3750 642 9 3812 4594 2728 6 Harvey-mahagony

0-15 2116 540 9 1946 3180 1631 6 El Verde

0-15 4844 1500 9 3808 7065 3550 6 Guzmán secondary forest

0-15 4086 5222 9 1793 16933 1026 6 Cubuy secondary forest

0-15 962 467 9 968 1809 292 6 Cubuy pine

15-30 984 299 9 870 1457 649 6 Guzmán pine

15-30 2555 488 9 2532 3263 1917 6 Harvey-mahagony

15-30 1355 459 9 1359 2033 736 6 El Verde

15-30 4233 2406 9 3800 8429 1708 6 Guzmán secondary forest

15-30 1544 1513 9 1046 5316 501 6 Cubuy secondary forest

15-30 793 275 9 718 1172 440 6 Cubuy pine
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

344

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Clay (%) 0-15 43 8 9 47 52 29 6 Guzmán pine

0-15 28 13 9 26 50 14 6 Harvey-mahagony

0-15 37 20 9 29 78 19 6 El Verde

0-15 32 8 9 31 43 23 6 Guzmán secondary forest

0-15 44 20 9 36 83 24 6 Cubuy secondary forest

0-15 59 18 9 58 81 35 6 Cubuy pine

15-30 52 13 9 52 74 34 6 Guzmán pine

15-30 43 10 9 40 60 32 6 Harvey-mahagony

15-30 57 20 9 46 89 34 6 El Verde

15-30 38 8 9 41 48 25 6 Guzmán secondary forest

15-30 76 15 9 81 88 41 6 Cubuy secondary forest

15-30 68 19 9 68 89 42 6 Cubuy pine

ECEC (cmol/kg-1) 0-15 6.96 0.86 9 6.86 8.16 5.83 6 Guzmán pine

0-15 11.16 2.35 9 10.33 14.21 7.56 6 Harvey-mahagony

0-15 10.43 0.42 9 10.36 11.11 9.76 6 El Verde

0-15 25.73 9.30 9 28.00 39.01 13.12 6 Guzmán secondary forest

0-15 8.71 2.48 9 8.57 12.32 5.52 6 Cubuy secondary forest

0-15 7.54 0.70 9 7.33 8.72 6.41 6 Cubuy pine

15-30 6.04 0.83 9 6.13 6.93 4.27 6 Guzmán pine

15-30 9.01 1.48 9 8.52 11.16 7.29 6 Harvey-mahagony

15-30 7.56 0.97 9 7.85 8.74 5.86 6 El Verde

15-30 24.76 13.97 9 31.60 40.47 8.16 6 Guzmán secondary forest

15-30 9.30 9.66 9 5.80 34.79 4.37 6 Cubuy secondary forest

15-30 5.69 0.62 9 5.37 6.59 5.04 6 Cubuy pine

Iron (mg/kg) 0-15 415 84 9 411 605 346 6 Guzmán pine

0-15 195 41 9 204 246 125 6 Harvey-mahagony
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

345

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0-15 1520 504 9 1533 2330 841 6 El Verde

0-15 289 117 9 291 486 125 6 Guzmán secondary forest

0-15 2428 1216 9 1926 4218 474 6 Cubuy secondary forest

0-15 1492 615 9 1410 2791 851 6 Cubuy pine

15-30 165 57 9 162 285 79 6 Guzmán pine

15-30 117 41 9 106 200 62 6 Harvey-mahagony

15-30 675 434 9 579 1669 234 6 El Verde

15-30 228 184 9 119 514 70 6 Guzmán secondary forest

15-30 1400 853 9 1254 3165 480 6 Cubuy secondary forest

15-30 435 174 9 392 852 230 6 Cubuy pine

Iron-total (%) 0-15 9.15 0.63 9 9.19 10.52 8.47 6 Guzmán pine

0-15 8.17 0.57 9 8.17 8.95 7.33 6 Harvey-mahagony

0-15 9.26 1.48 9 9.59 11.00 7.20 6 El Verde

0-15 7.90 1.02 9 7.77 9.77 6.68 6 Guzmán secondary forest

0-15 6.79 0.92 9 7.10 8.16 5.08 6 Cubuy secondary forest

0-15 8.32 2.99 9 7.62 15.90 6.11 6 Cubuy pine

15-30 10.16 0.66 9 10.06 11.02 9.31 6 Guzmán pine

15-30 8.80 0.56 9 9.00 9.54 8.06 6 Harvey-mahagony

15-30 10.72 1.29 9 10.55 12.75 8.65 6 El Verde

15-30 8.67 2.03 9 7.96 12.03 6.43 6 Guzmán secondary forest

15-30 7.71 0.69 9 8.00 8.50 6.64 6 Cubuy secondary forest

15-30 7.69 0.66 9 7.75 8.69 6.62 6 Cubuy pine

Magnesium (mg/kg) 0-15 284 75 9 287 424 176 6 Guzmán pine

0-15 479 204 9 408 760 198 6 Harvey-mahagony

0-15 334 66 9 359 432 224 6 El Verde

0-15 1781 1143 9 2289 3434 281 6 Guzmán secondary forest
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

346

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0-15 226 159 9 159 515 62 6 Cubuy secondary forest

0-15 94 88 9 54 301 28 6 Cubuy pine

15-30 197 57 9 210 265 97 6 Guzmán pine

15-30 350 129 9 341 655 212 6 Harvey-mahagony

15-30 194 37 9 189 250 141 6 El Verde

15-30 2084 1444 9 2763 3633 225 6 Guzmán secondary forest

15-30 100 59 9 72 201 25 6 Cubuy secondary forest

15-30 62 37 9 44 115 27 6 Cubuy pine

Magnesium-total (mg/kg) 0-15 1033 129 9 1027 1301 823 6 Guzmán pine

0-15 2297 629 9 2169 3682 1662 6 Harvey-mahagony

0-15 948 152 9 957 1135 694 6 El Verde

0-15 14093 8300 9 13305 27746 2065 6 Guzmán secondary forest

0-15 1665 1534 9 1122 5373 635 6 Cubuy secondary forest

0-15 601 193 9 532 1094 464 6 Cubuy pine

15-30 991 93 9 979 1201 914 6 Guzmán pine

15-30 2876 1838 9 2089 7097 1542 6 Harvey-mahagony

15-30 780 160 9 766 982 536 6 El Verde

15-30 17617 11930 9 15079 35029 1656 6 Guzmán secondary forest

15-30 886 453 9 859 1900 464 6 Cubuy secondary forest

15-30 463 60 9 448 586 394 6 Cubuy pine

Manganese (mg/kg) 0-15 412 166 9 386 750 242 6 Guzmán pine

0-15 269 99 9 238 492 179 6 Harvey-mahagony

0-15 114 58 9 117 222 51 6 El Verde

0-15 223 148 9 140 462 96 6 Guzmán secondary forest

0-15 106 46 9 118 180 27 6 Cubuy secondary forest

0-15 177 83 9 209 283 60 6 Cubuy pine
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

347

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

15-30 154 128 9 112 387 33 6 Guzmán pine

15-30 270 95 9 299 445 162 6 Harvey-mahagony

15-30 60 38 9 39 108 16 6 El Verde

15-30 149 176 9 56 462 14 6 Guzmán secondary forest

15-30 50 55 9 22 170 5 6 Cubuy secondary forest

15-30 118 81 9 100 274 28 6 Cubuy pine

Manganese-total (mg/kg) 0-15 3842 1615 9 3441 6962 1904 6 Guzmán pine

0-15 2685 412 9 2590 3443 2130 6 Harvey-mahagony

0-15 593 280 9 486 1124 259 6 El Verde

0-15 2207 627 9 2119 3212 1230 6 Guzmán secondary forest

0-15 454 253 9 426 873 104 6 Cubuy secondary forest

0-15 766 399 9 722 1646 260 6 Cubuy pine

15-30 2717 1866 9 2573 6221 595 6 Guzmán pine

15-30 2969 464 9 2810 6486 1647 6 Harvey-mahagony

15-30 589 285 9 518 1124 261 6 El Verde

15-30 1703 881 9 1601 3384 487 6 Guzmán secondary forest

15-30 389 267 9 421 938 92 6 Cubuy secondary forest

15-30 678 451 9 602 1557 231 6 Cubuy pine

Nitrogen (%) 0-15 0.23 0.02 9 0.22 0.26 0.20 6 Guzmán pine

0-15 0.28 0.04 9 0.30 0.32 0.22 6 Harvey-mahagony

0-15 0.42 0.02 9 0.42 0.46 0.37 6 El Verde

0-15 0.23 0.11 9 0.19 0.40 0.07 6 Guzmán secondary forest

0-15 0.32 0.04 9 0.32 0.37 0.24 6 Cubuy secondary forest

0-15 0.29 0.07 9 0.30 0.40 0.14 6 Cubuy pine

15-30 0.14 0.04 9 0.14 0.22 0.09 6 Guzmán pine

15-30 0.14 0.04 9 0.14 0.19 0.08 6 Harvey-mahagony
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

348

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

15-30 0.26 0.03 9 0.25 0.30 0.22 6 El Verde

15-30 0.11 0.11 9 0.05 0.31 0.02 6 Guzmán secondary forest

15-30 0.20 0.03 9 0.18 0.25 0.16 6 Cubuy secondary forest

15-30 0.21 0.02 9 0.22 0.24 0.18 6 Cubuy pine

Organic matter (%) 0-15 3.77 0.83 9 3.80 4.64 1.72 6 Guzmán pine

0-15 5.21 0.72 9 5.08 6.14 3.76 6 Harvey-mahagony

0-15 8.43 0.49 9 8.44 9.05 7.36 6 El Verde

0-15 4.61 1.85 9 4.45 7.43 1.69 6 Guzmán secondary forest

0-15 6.76 0.81 9 6.91 7.94 5.27 6 Cubuy secondary forest

0-15 6.16 1.15 9 6.04 8.70 4.51 6 Cubuy pine

15-30 2.25 0.51 9 2.25 3.28 1.54 6 Guzmán pine

15-30 2.59 0.68 9 2.51 3.45 1.63 6 Harvey-mahagony

15-30 4.91 0.72 9 4.78 6.24 3.90 6 El Verde

15-30 1.98 1.66 9 1.13 5.40 0.56 6 Guzmán secondary forest

15-30 4.09 0.70 9 3.79 5.60 3.55 6 Cubuy secondary forest

15-30 3.72 0.57 9 3.64 4.57 2.80 6 Cubuy pine

pH (H2O) 0-15 4.42 0.13 9 4.42 4.68 4.24 6 Guzmán pine

0-15 4.56 0.20 9 4.53 4.89 4.32 6 Harvey-mahagony

0-15 4.40 0.12 9 4.36 4.63 4.28 6 El Verde

0-15 5.32 0.24 9 5.34 5.60 5.02 6 Guzmán secondary forest

0-15 4.79 0.25 9 4.74 5.38 4.59 6 Cubuy secondary forest

0-15 4.49 0.30 9 4.50 5.02 3.98 6 Cubuy pine

15-30 4.48 0.12 9 4.53 4.61 4.24 6 Guzmán pine

15-30 4.47 0.13 9 4.49 4.63 4.31 6 Harvey-mahagony

15-30 4.51 0.12 9 4.49 4.74 4.37 6 El Verde

15-30 5.35 0.32 9 5.36 5.96 4.93 6 Guzmán secondary forest
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

349

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

15-30 4.69 0.21 9 4.65 5.18 4.49 6 Cubuy secondary forest

15-30 4.75 0.19 9 4.84 4.92 4.35 6 Cubuy pine

pH (KCl) 0-15 4.11 0.19 9 4.05 4.58 3.89 6 Guzmán pine

0-15 4.54 0.19 9 4.60 4.79 4.22 6 Harvey-mahagony

0-15 3.93 0.12 9 3.89 4.19 3.77 6 El Verde

0-15 4.73 0.39 9 4.72 5.58 4.29 6 Guzmán secondary forest

0-15 4.37 0.25 9 4.34 4.92 4.06 6 Cubuy secondary forest

0-15 4.23 0.20 9 4.17 4.67 3.94 6 Cubuy pine

15-30 4.07 0.10 9 4.05 4.22 3.91 6 Guzmán pine

15-30 4.17 0.22 9 4.06 4.52 3.97 6 Harvey-mahagony

15-30 4.00 0.10 9 3.95 4.16 3.90 6 El Verde

15-30 4.49 0.30 9 4.56 5.06 4.15 6 Guzmán secondary forest

15-30 4.16 0.44 9 3.96 4.94 3.82 6 Cubuy secondary forest

15-30 4.36 0.21 9 4.37 4.70 3.93 6 Cubuy pine

Phosphorus (mg/kg) 0-15 5 2 9 6 7 2 6 Guzmán pine

0-15 11 2 9 11 14 8 6 Harvey-mahagony

0-15 18 3 9 18 21 12 6 El Verde

0-15 6 5 9 5 16 1 6 Guzmán secondary forest

0-15 16 5 9 16 25 6 6 Cubuy secondary forest

0-15 20 5 9 21 28 13 6 Cubuy pine

15-30 2 1 9 2 5 1 6 Guzmán pine

15-30 5 1 9 5 7 3 6 Harvey-mahagony

15-30 6 2 9 7 9 4 6 El Verde

15-30 4 4 9 1 11 1 6 Guzmán secondary forest

15-30 5 3 9 4 10 2 6 Cubuy secondary forest

15-30 9 3 9 10 13 5 6 Cubuy pine
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

350

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Phosphorus-total (mg/kg) 0-15 377 49 9 362 453 316 6 Guzmán pine

0-15 229 76 9 212 416 135 6 Harvey-mahagony

0-15 351 59 9 359 443 250 6 El Verde

0-15 257 142 9 264 422 68 6 Guzmán secondary forest

0-15 255 86 9 186 386 155 6 Cubuy secondary forest

0-15 263 40 9 276 316 194 6 Cubuy pine

15-30 301 37 9 308 349 223 6 Guzmán pine

15-30 172 48 9 158 256 116 6 Harvey-mahagony

15-30 273 61 9 271 359 183 6 El Verde

15-30 187 149 9 141 418 36 6 Guzmán secondary forest

15-30 134 68 9 131 285 40 6 Cubuy secondary forest

15-30 189 60 9 180 279 108 6 Cubuy pine

Potassium (mg/kg) 0-15 107 43 9 98 184 63 6 Guzmán pine

0-15 756 141 9 770 1007 578 6 Harvey-mahagony

0-15 281 36 9 286 328 201 6 El Verde

0-15 651 211 9 715 855 246 6 Guzmán secondary forest

0-15 383 572 9 205 1905 121 6 Cubuy secondary forest

0-15 139 47 9 116 243 102 6 Cubuy pine

15-30 56 28 9 43 113 32 6 Guzmán pine

15-30 489 112 9 490 662 321 6 Harvey-mahagony

15-30 145 20 9 141 176 114 6 El Verde

15-30 404 169 9 430 638 134 6 Guzmán secondary forest

15-30 92 36 9 74 163 64 6 Cubuy secondary forest

15-30 115 109 9 80 404 60 6 Cubuy pine

Potassium-total (%) 0-15 0.58 0.06 9 0.56 0.72 0.53 6 Guzmán pine

0-15 0.36 0.08 9 0.42 0.45 0.27 6 Harvey-mahagony
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

351

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0-15 0.54 0.13 9 0.55 0.73 0.34 6 El Verde

0-15 0.78 0.08 9 0.80 0.90 0.66 6 Guzmán secondary forest

0-15 0.45 0.13 9 0.43 0.77 0.35 6 Cubuy secondary forest

0-15 0.68 0.24 9 0.63 1.30 0.50 6 Cubuy pine

15-30 0.57 0.05 9 0.57 0.66 0.49 6 Guzmán pine

15-30 0.34 0.07 9 0.31 0.44 0.27 6 Harvey-mahagony

15-30 0.57 0.12 9 0.56 0.72 0.39 6 El Verde

15-30 0.77 0.09 9 0.75 0.92 0.67 6 Guzmán secondary forest

15-30 0.41 0.08 9 0.38 0.57 0.32 6 Cubuy secondary forest

15-30 0.62 0.07 9 0.64 0.68 0.48 6 Cubuy pine

Sand (%) 0-15 25 8 9 22 36 16 6 Guzmán pine

0-15 36 15 9 43 52 16 6 Harvey-mahagony

0-15 36 18 9 44 54 5 6 El Verde

0-15 27 9 9 28 41 16 6 Guzmán secondary forest

0-15 23 15 9 22 42 2 6 Cubuy secondary forest

0-15 18 12 9 16 34 1 6 Cubuy pine

15-30 19 12 9 21 35 2 6 Guzmán pine

15-30 24 7 9 27 31 9 6 Harvey-mahagony

15-30 21 14 9 24 38 1 6 El Verde

15-30 22 7 9 24 32 14 6 Guzmán secondary forest

15-30 6 8 9 3 27 1 6 Cubuy secondary forest

15-30 14 10 9 11 31 4 6 Cubuy pine

Silt (%) 0-15 33 2 9 32 38 31 6 Guzmán pine

0-15 36 7 9 34 55 32 6 Harvey-mahagony

0-15 27 8 9 27 44 17 6 El Verde

0-15 41 6 9 40 55 34 6 Guzmán secondary forest
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

352

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0-15 33 8 9 35 43 15 6 Cubuy secondary forest

0-15 23 8 9 26 32 9 6 Cubuy pine

15-30 29 3 9 28 34 25 6 Guzmán pine

15-30 33 4 9 33 38 27 6 Harvey-mahagony

15-30 22 8 9 23 37 10 6 El Verde

15-30 40 3 9 41 44 34 6 Guzmán secondary forest

15-30 19 7 9 17 32 11 6 Cubuy secondary forest

15-30 19 10 9 21 30 6 6 Cubuy pine

Sodium (mg/kg) 0-15 31 5 9 30 41 23 6 Guzmán pine

0-15 89 9 9 90 105 73 6 Harvey-mahagony

0-15 96 16 9 97 116 75 6 El Verde

0-15 138 40 9 136 192 58 6 Guzmán secondary forest

0-15 60 25 9 69 88 15 6 Cubuy secondary forest

0-15 25 8 9 24 38 13 6 Cubuy pine

15-30 28 7 9 25 37 19 6 Guzmán pine

15-30 80 22 9 87 109 40 6 Harvey-mahagony

15-30 70 13 9 74 85 52 6 El Verde

15-30 155 68 9 170 245 54 6 Guzmán secondary forest

15-30 56 23 9 55 90 12 6 Cubuy secondary forest

Tuffaceous sandstone Ultisols Smf 15-30 16 6 9 15 22 10 6 Cubuy Pine

Tuffaceous sandstone Inceptisols Wflm Manganese (mg/kg) 0-10 21 15 4 17 42 5 7 Toro Trail-forest

0-10 36 29 8 37 76 1 7 Toro Trail-landslide

Tuffaceous sandstone Inceptisols Wflm 0-10 11 9 2 11 17 5 7 Rio Cubuy-landslide

Tuffaceous sandstone Ultisols Wflm 0-10 28 1 7 Quebrada La Coca-forest

0-10 59 19 2 59 73 46 7 Quebrada La Coca-landslide

0-10 36 1 7 Quebrada Sonadora-forest
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

353

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0-10 111 6 2 111 115 106 7 Queb.Sonadora-landslide

0-10 16 1 7 Rio Cubuy-forest

0-10 10 1 7 Rio Espiritu Santo-forest

Tuffaceous sandstone Ultisols Wflm 0-10 97 9 2 97 103 90 7 Rio Esp. Santo-landslide

Tuffaceous sandstone Inceptisols Wflm Phosphorus (mg/kg) 0-10 22 10 4 19 36 15 7 Toro Trail-forest

0-10 8 6 8 8 17 1 7 Toro Trail-landslide

0-10 5 5 2 5 8 1 7 Rio Cubuy-landslide

Tuffaceous sandstone Ultisolls Wflm 0-10 110 1 7 Rio Espiritu Santo-forest

0-10 1 0 2 1 1 1 7 Rio Esp. Santo-landslide

0-10 20 1 7 Quebrada La Coca-forest

0-10 6 1 2 6 7 6 7 Quebrada La Coca-landslide

0-10 26 1 7 Quebrada Sonadora-forest

0-10 2 2 2 2 4 1 7 Quebrada Sonadora-landslide

0-10 24 1 7 Rio Cubuy-forest

Tuffaceous sandstone Inceptisols Wflm Potassium (mg/kg) 0-10 113 44 4 108 164 42 7 Toro Trail-forest

0-10 48 29 8 40 87 16 7 Toro Trail-landslide

Tuffaceous sandstone Ultisols Wflm 0-10 99 1 7 Quebrada La Coca-forest

0-10 62 10 2 62 69 54 7 Quebrada La Coca-landslide

0-10 115 1 7 Queb.Sonadora-forest

0-10 43 34 2 43 67 19 7 Quebrada Sonadora-landslide

0-10 128 1 7 Rio Cubuy-forest

0-10 44 31 2 44 66 23 7 Rio Cubuy-landslide

0-10 155 1 7 Rio Espiritu Santo-forest

0-10 42 9 2 42 48 36 7 Rio Esp. Santo-landslide

Tuffaceous sandstone Ultisols Smf Clay (%) 0--30 51 10 3 53 60 38 8 Barrancas, Barranquitas

Tuffaceous sandstone Inceptisols Swf 0--30 47 1 8 Buena Vista, Las Marias
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

354

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Lava, tuff Ultisols Smf 0--30 40 9 2 40 46 33 8 Carite Forest

0--30 37 1 8 Cercadilla, Cayey

0--30 41 10 2 41 48 34 8 Cerro El Gato, Cayey

Limestone Ultisols Swf 0--30 44 16 4 44 60 30 8 Cerro Gordo, Moca

Tuffaceous sandstone Ultisols Swf 0--30 19 1 8 Cubuy, Mab

Plutonic rocks Inceptisols Smf 0--30 15 1 8 Espino, SanLorenzo

Lava, tuff Ultisols Swf 0--30 34 1 8 Guavate Forest

Alluvial deposits Inceptisols Smf 0--30 17 1 8 Guayanes, Yabucoa

Tuffaceous sandstone Ultisols Swf 0--30 59 4 3 60 62 55 8 Guzmán Abajo, Mab

Serpentinite Ultisols Swf 0--30 52 1 8 Indiera Arriba, Maricao

Tuffaceous sandstone Ultisols Swf 0--30 54 4 2 54 57 50 8 Las Marias

Limestone Entisols Swf 0--30 65 1 8 Mamey, Moca

Plutonic rocks Inceptisols Swf 0--30 16 2 3 16 18 15 8 Paso Palmas, Utuado

0--30 27 8 2 27 32 21 8 Piedras Blancas, Yabucoa

Plutonic rocks Ultisols Swf 0--30 22 1 8 Saltos Caguana, Utuado

30--100 64 4 2 64 67 60 8 Barrancas, Barranquitas

Tuffaceous sandstone Inceptisols Swf 30--100 37 1 8 Buena Vista, Las Marias

Lava, tuff Ultisols Smf 30--100 44 9 2 54 60 48 8 Carite Forest

30--100 55 1 8 Cercadilla, Cayey

30--100 52 6 2 52 56 48 8 Cerro El Gato, Cayey

Limestone Ultisols Swf 30--100 51 12 4 56 58 34 8 Cerro Gordo, Moca

Tuffaceous sandstone Ultisols Swf 30--100 36 1 8 Cubuy, Mab

Plutonic rocks Inceptisols Smf 30--100 12 1 8 Espino, San Lorenzo

Lava, tuff Ultisols Swf 30--100 60 1 8 Guavate Forest

Alluvial deposits Inceptisols Smf 30--100 25 1 8 Guayanes, Yabucoa

Tuffaceous sandstone Ultisols Swf 30--100 70 5 3 72 74 64 8 Guzmán Abajo, Mab
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

355

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Serpentinite Ultisols Swf 30--100 57 1 8 Indiera Arriba, Maricao

Tuffaceous sandstone Ultisols Swf 30--100 52 2 2 52 54 50 8 Las Marias

Limestone Entisols Swf 30--100 65 1 8 Mamey, Moca

Plutonic rocks Inceptisols Swf 30--100 16 4 3 16 21 12 8 Paso Palmas, Utuado

30--100 23 1 2 23 23 22 8 Piedras Blancas, Yabucoa 

Plutonic rocks Ultisols Swf 30--100 22 1 8 Saltos Caguana, Utuado

Tuffaceous sandstone Ultisols Smf Loss on ignition (%) 0--30 13.59 3.43 3 15.28 15.85 9.64 8 Barrancas, Barranquitas

Tuffaceous sandstone Inceptisols Swf 0--30 9.57 1 8 Buena Vista, Las Marias

Lava, tuff Ultisols Smf 0--30 13.86 3.44 2 13.86 16.29 11.43 8 Carite Forest

Tuffaceous sandstone Ultisols Swf 0--30 9.32 1 8 Cercadilla, Cayey

0--30 12.14 0.90 2 12.14 12.77 11.50 8 Cerro El Gato, Cayey

Limestone Ultisols Swf 0--30 12.24 2.31 4 11.73 15.25 10.24 8 Cerro Gordo, Moca

Tuffaceous sandstone Ultisols Swf 0--30 16.26 1 8 Cubuy, Mab

Plutonic rocks Inceptisols Smf 0--30 6.11 1 8 Espino, San Lorenzo

Lava, tuff Ultisols Swf 0--30 11.15 1 8 Guavate Forest

Alluvial deposits Inceptisols Smf 0--30 3.78 1 8 Guayanes, Yabucoa

Tuffaceous sandstone Ultisols Swf 0--30 15.45 1.06 3 15.39 16.54 14.42 8 Guzmán Abajo, Mab

Serpentinite Ultisols Swf 0--30 12.80 1 8 Indiera Arriba, Maricao

Tuffaceous sandstone Ultisols Swf 0--30 11.71 2.62 2 11.71 13.56 9.86 8 Las Marias

Limestone Entisols Swf 0--30 8.98 1 8 Mamey, Moca

Plutonic rocks Inceptisols Swf 0--30 5.02 0.32 3 4.99 5.35 4.72 8 Paso Palmas, Utuado

0--30 7.71 0.16 2 7.71 7.82 7.59 8 Piedras Blancas, Yabucoa

Plutonic rocks Ultisols Swf 0--30 6.15 1 8 Saltos Caguana, Utuado

Tuffaceous sandstone Ultisols Smf 30--100 11.09 2.84 2 11.09 13.10 9.08 8 Barrancas, Barranquitas

Tuffaceous sandstone Inceptisols Swf 30--100 11.22 1 8 Buena Vista, Las Marias

Lava, tuff Ultisols Smf 30--100 11.77 1.49 2 11.77 12.82 10.71 8 Carite Forest
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

356

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Tuffaceous sandstone Ultisols Swf 30--100 8.54 1 8 Cercadilla, Cayey

Tuffaceous sandstone Ultisols Swf 30--100 8.15 0.41 2 8.15 8.44 7.86 8 Cerro El Gato, Cayey

Limestone Ultisols Swf 30--100 12.46 1.27 4 12.79 13.52 10.74 8 Cerro Gordo, Moca

Tuffaceous sandstone Ultisols Swf 30--100 9.81 1 8 Cubuy, Mab

Plutonic rocks Inceptisols Smf 30--100 4.47 1 8 Espino, San Lorenzo

Lava, tuff Ultisols Swf 30--100 12.87 1 8 Guavate Forest

Alluvial deposits Inceptisols Smf 30--100 3.18 1 8 Guayanes, Yabucoa

Tuffaceous sandstone Ultisols Swf 30--100 14.88 1.18 3 14.81 16.09 13.73 8 Guzmán Abajo, Mab

Serpentinite Ultisols Swf 30--100 12.60 1 8 Indiera Arriba, Maricao

Tuffaceous sandstone Ultisols Swf 30--100 10.31 0.63 2 10.31 10.75 9.86 8 Las Marias

Limestone Entisols Swf 30--100 9.38 1 8 Mamey, Moca

Plutonic rocks Inceptisols Swf 30--100 4.94 0.73 3 4.93 5.67 4.21 8 Paso Palmas, Utuado

Plutonic rocks Inceptisols Smf 30--100 5.78 0.62 2 5.78 6.22 5.34 8 Piedras Blancas, Yabucoa 

Plutonic rocks Ultisols Swf 30--100 7.00 1 8 Saltos Caguana, Utuado

Tuffaceous sandstone Ultisols Smf Organic matter (%) 0--30 3.90 1.13 3 3.52 5.17 3.01 8 Barrancas, Barranquitas

30--100 1.70 0.09 2 1.70 1.76 1.63 8 Barrancas, Barranquitas

Tuffaceous sandstone Inceptisols Swf 0--30 2.14 1 8 Buena Vista, Las Marias

30--100 0.50 1 8 Buena Vista, Las Marias

Lava, tuff Ultisols Smf 0--30 3.59 1.51 2 3.59 4.66 2.52 8 Carite Forest

30--100 2.11 0.30 2 2.11 2.32 1.89 8 Carite Forest

Tuffaceous sandstone Ultisols Swf 0--30 2.77 1 8 Cercadilla, Cayey

30--100 1.25 1 8 Cercadilla, Cayey

0--30 4.42 0.70 2 4.42 4.91 3.92 8 Cerro El Gato, Cayey

30--100 1.39 0.18 2 1.39 1.52 1.26 8 Cerro El Gato, Cayey

Limestone Ultisols Swf 0--30 2.47 0.48 4 2.49 3.02 1.88 8 Cerro Gordo, Moca

30--100 0.76 0.42 4 0.76 1.26 0.25 8 Cerro Gordo, Moca
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

357

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Tuffaceous sandstone Ultisols Swf 0--30 6.41 1 8 Cubuy, Mab

30--100 1.38 1 8 Cubuy, Mab

Plutonic rocks Inceptisols Smf 0--30 2.64 1 8 Espino, San Lorenzo

30--100 1.13 1 8 Espino, San Lorenzo

Lava, tuff Ultisols Swf 0--30 2.78 1 8 Guavate Forest

30--100 1.14 1 8 Guavate Forest

Alluvial deposits Inceptisols Smf 0--30 1.27 1 8 Guayanes, Yabucoa

30--100 0.63 1 8 Guayanes, Yabucoa

Tuffaceous sandstone Ultisols Swf 0--30 2.68 0.59 3 2.88 3.15 2.02 8 Guzmán Abajo, Mab

30--100 1.05 0.14 3 1.13 1.13 0.88 8 Guzmán Abajo, Mab

Serpentinite Ultisols Swf 0--30 3.77 1 8 Indiera Arriba, Maricao

30--100 1.38 1 8 Indiera Arriba, Maricao

Tuffaceous sandstone Ultisols Swf 0--30 3.15 0.35 2 3.15 3.39 2.90 8 Las Marias

30--100 1.51 0.18 2 1.51 1.64 1.38 8 Las Marias

Limestone Entisols Swf 0--30 1.51 1 8 Mamey, Moca

30--100 0.63 1 8 Mamey, Moca

Plutonic rocks Inceptisols Swf 0--30 1.51 0.58 3 1.64 2.02 0.88 8 Paso Palmas, Utuado

30--100 0.88 0.66 3 1.26 1.26 0.12 8 Paso Palmas, Utuado

0--30 1.89 0.89 2 1.89 2.52 1.26 8 Piedras Blancas, Yabucoa 

30--100 0.88 0.71 2 0.88 1.38 0.38 8 Piedras Blancas, Yabucoa

Plutonic rocks Ultisols Swf 0--30 1.70 1 8 Saltos Caguana, Utuado

Plutonic rocks Ultisols Swf 30--100 1.01 1 8 Saltos Caguana, Utuado

Tuffaceous sandstone Ultisols Smf pH (H2O) 0--30 4.44 0.17 3 4.52 4.56 4.24 8 Barrancas, Barranquitas

30--100 4.40 0.16 2 4.40 4.51 4.28 8 Barrancas, Barranquitas

Tuffaceous sandstone Inceptisols Swf 0--30 4.61 1 8 Buena Vista, Las Marias

30--100 4.74 1 8 Buena Vista, Las Marias
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

358

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Lava, tuff Ultisols Smf 0--30 4.52 0.20 2 4.52 4.66 4.38 8 Carite Forest

30--100 4.52 0.04 2 4.52 4.55 4.49 8 Carite Forest

Tuffaceous sandstone Ultisols Swf 0--30 4.37 1 8 Cercadilla, Cayey

30--100 4.08 1 8 Cercadilla, Cayey

0--30 4.23 0.03 2 4.23 4.25 4.21 8 Cerro El Gato, Cayey

30--100 4.19 0.23 2 4.19 4.35 4.02 8 Cerro El Gato, Cayey

Limestone Ultisols Swf 0--30 4.46 0.07 4 4.44 4.55 4.39 8 Cerro Gordo, Moca

30--100 4.37 0.12 4 4.37 4.52 4.23 8 Cerro Gordo, Moca

Tuffaceous sandstone Ultisols Swf 0--30 5.20 1 8 Cubuy, Mab

30--100 5.64 1 8 Cubuy, Mab

Plutonic rocks Inceptisols Smf 0--30 5.06 1 8 Espino, San Lorenzo

30--100 5.03 1 8 Espino, San Lorenzo

Lava, tuff Ultisols Swf 0--30 4.94 1 8 Guavate Forest

30--100 4.73 1 8 Guavate Forest

Alluvial deposits Inceptisols Smf 0--30 5.03 1 8 Guayanes, Yabucoa

30--100 4.88 1 8 Guayanes, Yabucoa

Tuffaceous sandstone Ultisols Swf 0--30 4.64 0.07 3 4.60 4.72 4.59 8 Guzmán Abajo, Mab

30--100 4.75 0.08 3 4.78 4.82 4.66 8 Guzmán Abajo, Mab

Serpentinite Ultisols Swf 0--30 4.44 1 8 Indiera Arriba, Maricao

30--100 4.36 1 8 Indiera Arriba, Maricao

Tuffaceous sandstone Ultisols Swf 0--30 4.34 0.05 2 4.34 4.37 4.30 8 Las Marias

30--100 4.30 0.09 2 4.30 4.36 4.23 8 Las Marias

Limestone Entisols Swf 0--30 4.30 1 8 Mamey, Moca

30--100 4.27 1 8 Mamey, Moca

Plutonic rocks Inceptisols Swf 0--30 5.14 0.34 3 4.96 5.53 4.94 8 Paso Palmas, Utuado

30--100 5.27 0.23 3 5.18 5.53 5.10 8 Paso Palmas, Utuado
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

359

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--30 4.52 0.01 2 4.52 4.52 4.51 8 Piedras Blancas, Yabucoa

30--100 4.62 0.14 2 4.62 4.72 4.52 8 Piedras Blancas, Yabucoa

Plutonic rocks Ultisols Swf 0--30 5.30 1 8 Saltos Caguana, Utuado

30--100 5.19 1 8 Saltos Caguana, Utuado

Tuffaceous sandstone Ultisols Smf Phosphorus-total (mg/g) 0--30 0.46 0.03 3 0.46 0.49 0.42 8 Barrancas, Barranquitas

30--100 0.50 0.13 2 0.50 0.59 0.41 8 Barrancas, Barranquitas

Tuffaceous sandstone Inceptisols Swf 0--30 0.29 1 8 Buena Vista, Las Marias

30--100 0.27 1 8 Buena Vista, Las Marias

Lava, tuff Ultisols Smf 0--30 0.34 0.14 2 0.34 0.44 0.24 8 Carite Forest

30--100 0.31 0.08 2 0.31 0.37 0.25 8 Carite Forest

Tuffaceous sandstone Ultisols Swf 0--30 0.39 1 8 Cercadilla, Cayey

30--100 0.35 1 8 Cercadilla, Cayey

0--30 0.34 0.00 2 0.34 0.34 0.34 8 Cerro El Gato, Cayey

30--100 0.28 0.06 2 0.28 0.31 0.24 8 Cerro El Gato, Cayey

Limestone Ultisols Swf 0--30 0.19 0.06 4 0.19 0.24 0.13 8 Cerro Gordo, Moca

30--100 0.20 0.09 4 0.20 0.29 0.10 8 Cerro Gordo, Moca

Tuffaceous sandstone Ultisols Swf 0--30 0.45 1 8 Cubuy, Mab

30--100 0.30 1 8 Cubuy, Mab

Plutonic rocks Inceptisols Smf 0--30 0.26 1 8 Espino, San Lorenzo

30--100 0.23 1 8 Espino, San Lorenzo

Lava, tuff Ultisols Swf 0--30 0.21 1 8 Guavate Forest

30--100 0.13 1 8 Guavate Forest

Alluvial deposits Inceptisols Smf 0--30 0.09 1 8 Guayanes, Yabucoa

30--100 0.06 1 8 Guayanes, Yabucoa

Tuffaceous sandstone Ultisols Swf 0--30 0.20 0.07 3 0.24 0.24 0.12 8 Guzmán Abajo, Mab

30--100 0.22 0.02 3 0.22 0.23 0.19 8 Guzmán Abajo, Mab
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

360

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Serpentinite Ultisols Swf 0--30 0.44 1 8 Indiera Arriba, Maricao

30--100 0.35 1 8 Indiera Arriba, Maricao

Tuffaceous sandstone Ultisols Swf 0--30 0.55 0.09 2 0.55 0.62 0.49 8 Las Marias

30--100 0.49 0.09 2 0.49 0.55 0.42 8 Las Marias

Limestone Entisols Swf 0--30 0.15 1 8 Mamey, Moca

30--100 0.09 1 8 Mamey, Moca

Plutonic rocks Inceptisols Swf 0--30 0.33 0.20 3 0.25 0.56 0.19 8 Paso Palmas, Utuado

30--100 0.26 0.12 3 0.22 0.40 0.17 8 Paso Palmas, Utuado

0--30 0.11 0.01 2 0.11 0.12 0.10 8 Piedras Blancas, Yabucoa

30--100 0.09 0.00 2 0.09 0.10 0.09 8 Piedras Blancas, Yabucoa

Plutonic rocks Ultisols Swf 0--30 0.36 1 8 Saltos Caguana, Utuado

30--100 0.29 1 8 Saltos Caguana, Utuado

Tuffaceous sandstone Ultisols Smf Sand (%) 0--30 16 6 3 18 21 9 8 Barrancas, Barranquitas

30--100 13 0 2 13 13 13 8 Barrancas, Barranquitas

Tuffaceous sandstone Inceptisols Swf 0--30 18 1 8 Buena Vista, Las Marias

30--100 21 1 8 Buena Vista, Las Marias

Lava, tuff Ultisols Smf 0--30 20 4 2 20 23 17 8 Carite Forest

30--100 20 3 2 20 22 18 8 Carite Forest

Tuffaceous sandstone Ultisols Swf 0--30 17 1 8 Cercadilla, Cayey

30--100 13 1 8 Cercadilla, Cayey

0--30 21 4 2 21 23 18 8 Cerro El Gato, Cayey

30--100 18 3 2 18 21 16 8 Cerro El Gato, Cayey

Limestone Ultisols Swf 0--30 13 7 4 11 23 8 8 Cerro Gordo, Moca

30--100 16 7 4 14 26 9 8 Cerro Gordo, Moca

Tuffaceous sandstone Ultisols Swf 0--30 37 1 8 Cubuy, Mab

30--100 23 1 8 Cubuy, Mab
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

361

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Plutonic rocks Inceptisols Smf 0--30 63 1 8 Espino, San Lorenzo

30--100 66 1 8 Espino, San Lorenzo

Lava, tuff Ultisols Swf 0--30 37 1 8 Guavate Forest

30--100 8 1 8 Guavate Forest

Alluvial deposits Inceptisols Smf 0--30 58 1 8 Guayanes, Yabucoa

30--100 56 1 8 Guayanes, Yabucoa

Tuffaceous sandstone Ultisols Swf 0--30 12 3 3 11 14 9 8 Guzmán Abajo, Mab

30--100 4 1 3 4 6 3 8 Guzmán Abajo, Mab

Serpentinite Ultisols Swf 0--30 14 1 8 Indiera Arriba, Maricao

30--100 10 1 8 Indiera Arriba, Maricao

Tuffaceous sandstone Ultisols Swf 0--30 12 1 2 12 12 11 8 Las Marias

30--100 15 2 2 15 16 13 8 Las Marias

Limestone Entisols Swf 0--30 11 1 8 Mamey, Moca

30--100 11 1 8 Mamey, Moca

Plutonic rocks Inceptisols Swf 0--30 59 3 3 57 62 56 8 Paso Palmas, Utuado

30--100 61 6 3 64 65 55 8 Paso Palmas, Utuado

0--30 53 7 2 53 58 48 8 Piedras Blancas, Yabucoa 

30--100 55 5 2 55 59 52 8 Piedras Blancas, Yabucoa

Plutonic rocks Ultisols Swf 0--30 43 1 8 Saltos Caguana, Utuado

30--100 44 1 8 Saltos Caguana, Utuado

Tuffaceous sandstone Ultisols Smf Silt (%) 0--30 33 5 3 31 38 30 8 Barrancas, Barranquitas

30--100 24 4 2 24 27 21 8 Barrancas, Barranquitas

Tuffaceous sandstone Inceptisols Swf 0--30 34 1 8 Buena Vista, Las Marias

30--100 42 1 8 Buena Vista, Las Marias

Lava, tuff Ultisols Smf 0--30 40 5 2 40 44 37 8 Carite Forest

30--100 36 1 2 36 37 36 8 Carite Forest
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

362

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Tuffaceous sandstone Ultisols Swf 0--30 46 1 8 Cercadilla, Cayey

30--100 33 1 8 Cercadilla, Cayey

0--30 38 6 2 38 43 34 8 Cerro El Gato, Cayey

30--100 30 3 2 30 32 28 8 Cerro El Gato, Cayey

Limestone Ultisols Swf 0--30 42 13 4 41 59 30 8 Cerro Gordo, Moca

30--100 34 5 4 32 41 29 8 Cerro Gordo, Moca

Tuffaceous sandstone Ultisols Swf 0--30 44 1 8 Cubuy, Mab

30--100 41 1 8 Cubuy, Mab

Plutonic rocks Inceptisols Smf 0--30 22 1 8 Espino, San Lorenzo

30--100 23 1 8 Espino, San Lorenzo

Lava, tuff Ultisols Swf 0--30 29 1 8 Guavate Forest

30--100 32 1 8 Guavate Forest

Alluvial deposits Inceptisols Smf 0--30 25 1 8 Guayanes, Yabucoa

30--100 20 1 8 Guayanes, Yabucoa

Tuffaceous sandstone Ultisols Swf 0--30 29 1 3 29 31 28 8 Guzmán Abajo, Mab

30--100 26 4 3 24 31 23 8 Guzmán Abajo, Mab

Serpentinite Ultisols Swf 0--30 34 1 8 Indiera Arriba, Maricao

30--100 32 1 8 Indiera Arriba, Maricao

Tuffaceous sandstone Ultisols Swf 0--30 35 4 2 35 37 32 8 Las Marias

30--100 33 0 2 33 33 33 8 Las Marias

Limestone Entisols Swf 0--30 25 1 8 Mamey, Moca

30--100 24 1 8 Mamey, Moca

Plutonic rocks Inceptisols Swf 0--30 25 4 3 25 29 22 8 Paso Palmas, Utuado

30--100 23 3 3 24 25 20 8 Paso Palmas, Utuado

0--30 20 0 2 20 20 20 8 Piedras Blancas, Yabucoa 

30--100 22 4 2 22 25 19 8 Piedras Blancas, Yabucoa
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

363

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Plutonic rocks Ultisols Swf 0--30 35 1 8 Saltos Caguana, Utuado

30--100 34 1 8 Saltos Caguana, Utuado

Tuffaceous sandstone Ultisols Lmrf Aluminum (cmol/kg-l) 0--10 3.12 1.82 4 2.74 5.46 1.53 9 Top

0--10 2.43 0.73 9 2.30 3.36 1.33 9 Ridge

0--10 1.96 1.42 3 2.64 2.91 0.33 9 Side

0--10 2.86 1.32 4 2.92 4.15 1.47 9 Slope

0--10 1.69 0.86 9 1.76 2.83 0.21 9 Valley

10--20 2.69 0.86 4 2.55 3.85 1.81 9 Top

10--20 2.36 0.86 9 2.16 3.77 1.19 9 Ridge

10--20 2.12 1.71 3 2.85 3.34 0.16 9 Side

10--20 2.56 0.54 4 2.61 3.05 1.97 9 Slope

10--20 1.92 1.04 9 1.83 3.33 0.22 9 Valley

20--30 2.41 0.73 4 2.92 4.15 1.47 9 Top

20--30 2.25 1.00 9 1.83 3.77 1.15 9 Ridge

20--30 2.24 1.64 3 2.85 3.34 0.16 9 Side

20--30 2.47 0.46 4 2.92 4.15 1.47 9 Slope

20--30 1.69 1.02 9 1.83 3.15 0.24 9 Valley

Calcium (mg/g) 0--10 0.21 0.20 4 0.22 0.38 0.04 9 Top

0--10 0.14 0.06 9 0.15 0.24 0.05 9 Ridge

0--10 0.29 0.13 3 0.26 0.43 0.18 9 Side

0--10 0.20 0.19 4 0.17 0.43 0.04 9 Slope

0--10 0.45 0.24 9 0.43 0.86 0.13 9 Valley

10--20 0.09 0.10 4 0.07 0.23 0.01 9 Top

10--20 0.08 0.08 9 0.04 0.23 0.03 9 Ridge

10--20 0.11 0.08 3 0.07 0.21 0.07 9 Side

10--20 0.08 0.08 4 0.08 0.17 0.01 9 Slope
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

364

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--20 0.30 0.26 9 0.16 0.71 0.06 9 Valley

20--30 0.12 0.17 4 0.05 0.38 0.01 9 Top

20--30 0.05 0.05 9 0.04 0.16 0.01 9 Ridge

20--30 0.07 0.06 3 0.04 0.13 0.03 9 Side

20--30 0.09 0.09 4 0.06 0.21 0.01 9 Slope

20--30 0.34 0.30 9 0.20 0.83 0.05 9 Valley

Effective cation exchange (cmol/kg-l) 0--10 3.12 1.82 4 2.74 5.46 1.53 9 Top

0--10 2.43 0.73 9 2.30 3.36 1.33 9 Ridge

0--10 1.96 1.42 3 2.64 2.91 0.33 9 Side

0--10 2.86 1.32 4 2.92 4.15 1.47 9 Slope

0--10 1.69 0.86 9 1.76 2.83 0.21 9 Valley

10--20 2.69 0.86 4 2.55 3.85 1.81 9 Top

10--20 2.36 0.86 9 2.16 3.77 1.19 9 Ridge

10--20 2.12 1.71 3 2.85 3.34 0.16 9 Side

10--20 4.38 0.84 4 4.17 5.50 3.65 9 Slope

10--20 5.03 1.09 9 5.36 6.14 3.21 9 Valley

20--30 4.93 3.92 4 4.20 11.30 1.85 9 Top

20--30 3.96 2.90 9 3.05 10.31 1.60 9 Ridge

20--30 3.79 2.84 3 4.04 6.23 0.74 9 Side

20--30 4.41 2.34 4 4.35 8.61 1.91 9 Slope

20--30 5.04 3.20 9 4.41 10.10 1.39 9 Valley

Iron (mg/g) 0--10 1.67 1.44 4 1.50 3.35 0.32 9 Top

0--10 1.57 0.73 9 1.60 2.66 0.48 9 Ridge

0--10 2.45 0.16 3 2.51 2.58 2.27 9 Side

0--10 3.15 3.15 4 2.09 7.70 0.74 9 Slope

0--10 0.81 0.25 9 0.80 1.39 0.58 9 Valley
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

365

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--20 0.83 0.65 4 0.77 1.58 0.22 9 Top

10--20 0.57 0.46 9 0.43 1.70 0.23 9 Ridge

10--20 0.88 0.64 3 0.53 1.62 0.48 9 Side

10--20 2.73 3.20 4 1.71 7.23 0.28 9 Slope

10--20 0.41 0.17 9 0.37 0.79 0.21 9 Valley

20--30 0.54 0.39 4 0.56 0.89 0.14 9 Top

20--30 0.32 0.28 9 0.22 0.97 0.08 9 Ridge

20--30 0.29 0.16 3 0.37 0.40 0.11 9 Side

20--30 0.84 0.67 4 0.82 1.50 0.22 9 Slope

20--30 0.31 0.15 9 0.32 0.64 0.10 9 Valley

Magnesium (mg/g) 0--10 0.13 0.13 4 0.08 0.31 0.03 9 Top

0--10 0.12 0.05 9 0.11 0.21 0.04 9 Ridge

0--10 0.19 0.11 3 0.22 0.28 0.07 9 Side

0--10 0.17 0.10 4 0.17 0.26 0.07 9 Slope

0--10 0.19 0.06 9 0.21 0.31 0.11 9 Valley

10--20 0.15 0.16 3 0.09 0.38 0.04 9 Top

10--20 0.09 0.03 9 0.09 0.13 0.05 9 Ridge

10--20 0.12 0.11 3 0.10 0.24 0.03 9 Side

10--20 0.13 0.12 4 0.10 0.29 0.03 9 Slope

10--20 0.14 0.04 9 0.15 0.23 0.09 9 Valley

20--30 0.19 0.26 4 0.09 0.57 0.03 9 Top

20--30 0.14 0.18 9 0.10 0.63 0.04 9 Ridge

20--30 0.08 0.08 3 0.07 0.17 0.02 9 Side

20--30 0.15 0.16 4 0.10 0.36 0.03 9 Slope

20--30 0.15 0.05 9 0.15 0.22 0.09 9 Valley

Manganese (mg/kg) 0--10 23 21 4 21 48 3 9 Top
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

366

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--10 25 13 9 23 46 9 9 Ridge

0--10 15 5 3 18 19 10 9 Side

0--10 17 13 4 14 36 6 9 Slope

0--10 27 16 9 23 61 11 9 Valley

10--20 23 25 3 44 45 3 9 Top

10--20 21 21 9 14 58 1 9 Ridge

10--20 9 6 3 8 15 2 9 Side

10--20 14 12 4 15 25 2 9 Slope

10--20 23 14 9 21 57 9 9 Valley

20--30 35 39 4 35 71 1 9 Top

20--30 17 16 9 18 48 1 9 Ridge

20--30 25 26 3 21 53 2 9 Side

20--30 20 20 4 19 40 1 9 Slope

20--30 23 6 9 20 34 17 9 Valley

Organic matter (%) 0--10 5.17 1.30 4 5.11 6.81 3.66 9 Top

0--10 5.47 0.37 9 5.55 5.94 4.79 9 Ridge

0--10 6.10 0.21 3 6.08 6.32 5.90 9 Side

0--10 4.68 1.20 4 4.83 5.90 3.15 9 Slope

0--10 4.71 1.21 9 4.47 7.54 3.19 9 Valley

10--20 3.36 0.65 4 3.44 3.97 2.60 9 Top

10--20 3.61 0.69 9 3.74 4.53 2.54 9 Ridge

10--20 3.93 0.17 3 3.91 4.11 3.78 9 Side

10--20 3.24 0.38 4 3.29 3.56 2.81 9 Slope

10--20 3.44 0.58 9 3.62 4.15 2.54 9 Valley

20--30 2.91 0.56 4 3.01 3.46 2.16 9 Top

20--30 2.65 0.84 9 2.60 4.28 1.43 9 Ridge
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

367

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

20--30 2.93 0.69 3 2.72 3.70 2.36 9 Side

20--30 5.74 3.31 4 5.89 8.72 2.45 9 Slope

20--30 2.89 0.68 9 2.73 3.97 1.85 9 Valley

pH (H2O) 0--10 4.66 0.32 4 4.65 5.04 4.31 9 Top

0--10 4.63 0.21 9 4.59 5.00 4.29 9 Ridge

0--10 4.85 0.69 3 4.51 5.64 4.40 9 Side

0--10 4.67 0.23 4 4.67 4.93 4.41 9 Slope

0--10 4.79 0.26 9 4.75 5.39 4.50 9 Valley

10--20 4.86 0.23 4 4.90 5.09 4.55 9 Top

10--20 4.78 0.22 4 4.82 5.05 4.39 9 Ridge

10--20 4.89 0.55 3 4.65 5.52 4.51 9 Side

10--20 4.85 0.24 4 4.91 5.09 4.51 9 Slope

10--20 4.95 0.25 9 4.94 5.39 4.54 9 Valley

20--30 4.92 0.22 4 4.96 5.13 4.61 9 Top

20--30 4.88 0.26 9 4.88 5.16 4.32 9 Ridge

20--30 5.00 0.42 3 4.95 5.44 4.61 9 Side

20--30 4.87 0.20 4 4.88 5.10 4.63 9 Slope

20--30 5.04 0.24 9 4.99 5.39 4.73 9 Valley

Phosphorus (mg/kg) 0--10 7 3 4 6 11 5 9 Top

0--10 10 2 9 9 13 7 9 Ridge

0--10 14 3 3 12 18 12 9 Side

0--10 10 6 4 9 18 4 9 Slope

0--10 12 5 9 10 21 6 9 Valley

10--20 4 2 4 4 6 2 9 Top

10--20 4 1 9 4 6 2 9 Ridge

10--20 4 1 3 4 5 4 9 Side
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

368

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--20 6 4 4 5 12 3 9 Slope

10--20 7 3 9 7 12 2 9 Valley

20--30 3 1 4 3 3 2 9 Top

20--30 3 1 9 3 6 2 9 Ridge

20--30 3 1 3 3 4 3 9 Side

20--30 3 0 4 3 4 3 9 Slope

20--30 4 2 9 4 8 2 9 Valley

Potassium (mg/g) 0--10 0.10 0.02 4 0.10 0.12 0.07 9 Top

0--10 0.18 0.25 9 0.09 0.85 0.07 9 Ridge

0--10 0.12 0.02 3 0.12 0.15 0.10 9 Side

0--10 0.10 0.01 4 0.10 0.11 0.08 9 Slope

0--10 0.13 0.08 9 0.10 0.32 0.04 9 Valley

10--20 0.05 0.01 4 0.05 0.05 0.03 9 Top

10--20 0.05 0.01 9 0.05 0.07 0.03 9 Ridge

10--20 0.05 0.02 3 0.05 0.06 0.04 9 Side

10--20 0.06 0.03 4 0.06 0.09 0.04 9 Slope

10--20 0.08 0.06 9 0.06 0.23 0.02 9 Valley

20--30 0.04 0.02 4 0.04 0.06 0.03 9 Top

20--30 0.03 0.01 9 0.03 0.06 0.01 9 Ridge

20--30 0.12 0.10 3 0.10 0.23 0.04 9 Side

20--30 0.04 0.02 4 0.05 0.06 0.02 9 Slope

20--30 0.08 0.07 9 0.06 0.24 0.03 9 Valley

Sodium (mg/kg) 0--10 78 31 4 74 119 45 9 Top

0--10 56 15 9 55 80 31 9 Ridge

0--10 69 3 3 69 71 66 9 Side

0--10 56 14 4 54 74 42 9 Slope
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

369

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--10 56 18 9 52 87 30 9 Valley

10--20 51 23 4 52 74 28 9 Top

10--20 46 22 9 41 78 1 9 Ridge

10--20 56 18 3 46 76 45 9 Side

10--20 40 10 4 41 49 29 9 Slope

10--20 50 16 9 52 76 25 9 Valley

20--30 51 33 4 53 86 12 9 Top

20--30 42 23 9 34 88 16 9 Ridge

20--30 47 12 3 42 61 39 9 Side

20--30 40 15 4 42 54 24 9 Slope

20--30 47 18 9 42 82 19 9 Valley

Tuffaceous sandstone Ultisols Swf Aluminum (cmol/kg-l) 0--35 1.12 0.73 10 1.11 2.45 0.31 10

25--60 0.88 0.73 10 0.84 2.23 0.01 10

50--85 0.90 0.95 7 0.53 2.81 0.01 10

85--100 0.56 0.34 4 0.58 0.95 0.11 10

100--120 0.72 0.48 4 0.99 1.00 0.16 10

Aluminum-total (%) 0--35 12.68 2.40 10 12.22 16.87 10.20 10

25--60 13.99 3.13 10 13.71 20.16 9.89 10

50--85 19.92 2.68 7 20.16 22.40 9.89 10

85--100 19.34 4.08 4 20.18 22.82 14.15 10

100--120 19.31 3.13 4 19.91 22.43 14.98 10

Carbon/Nitrogen 0--35 15 2 10 16 19 12 10

25--60 20 15 10 16 62 13 10

50--85 13 2 7 13 62 10 10

85--100 13 2 4 12 16 11 10

100--120 15 8 4 13 26 9 10
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

370

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Calcium (mg/g) 0--35 1.08 0.66 10 1.16 2.14 0.12 10

25--60 0.82 0.57 10 0.97 1.57 0.10 10

50--85 0.83 0.50 7 0.78 1.57 0.10 10

85--100 0.94 0.54 4 1.00 1.53 0.22 10

100--120 1.38 0.17 4 1.32 1.63 1.25 10

Calcium-total (%) 0--35 0.43 0.17 10 0.43 0.76 0.14 10

25--60 0.39 0.16 10 0.37 0.76 0.15 10

50--85 0.31 0.31 7 0.17 0.98 0.09 10

85--100 0.39 0.32 4 0.28 0.86 0.15 10

100--120 0.27 0.05 4 0.26 0.34 0.22 10

ECEC (cmol/kg-1) 0--35 11.91 3.75 10 12.10 19.25 5.70 10

25--60 9.70 3.82 10 10.50 14.18 3.91 10

50--85 10.20 3.06 7 10.46 14.18 3.91 10

85--100 9.12 3.52 4 9.42 12.83 4.81 10

100--120 11.61 2.80 4 10.99 15.14 9.33 10

Iron (mg/g) 0--35 1.21 0.71 10 1.16 2.48 0.42 10

25--60 0.63 0.45 10 0.51 1.34 0.06 10

50--85 0.30 0.28 7 0.23 1.34 0.03 10

85--100 0.18 0.25 4 0.07 0.55 0.02 10

100--120 0.11 0.09 4 0.07 0.25 0.05 10

Iron-total (%) 0--35 5.40 1.08 10 5.38 6.65 3.95 10

25--60 6.09 1.23 10 6.66 7.43 4.05 10

50--85 8.41 1.41 7 8.45 10.72 4.05 10

85--100 8.45 1.19 4 8.62 9.67 6.88 10

100--120 8.22 1.94 4 8.14 10.64 5.97 10

Magnesium-total (%) 0--35 1.13 0.26 10 1.07 1.54 0.78 10
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

371

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

25--60 1.31 0.44 10 1.10 2.34 0.97 10

50--85 1.19 0.50 7 1.05 2.34 0.86 10

85--100 1.60 0.54 4 1.55 2.18 1.13 10

100--120 1.27 0.25 4 1.22 1.58 0.97 10

Manganese (mg/g) 0--35 0.08 0.05 10 0.06 0.18 0.03 10

25--60 0.10 0.09 10 0.08 0.30 0.02 10

50--85 0.06 0.04 7 0.07 0.30 0.01 10

85--100 0.04 0.03 4 0.04 0.07 0.01 10

100--120 0.05 0.02 4 0.05 0.07 0.03 10

Manganese-total (%) 0--35 0.07 0.03 10 0.05 0.15 0.04 10

25--60 0.11 0.07 10 0.12 0.28 0.03 10

50--85 0.10 0.05 7 0.12 0.28 0.03 10

85--100 0.13 0.05 4 0.13 0.18 0.06 10

100--120 0.16 0.07 4 0.16 0.23 0.07 10

Nitrogen (%) 0--35 0.40 0.12 10 0.40 0.63 0.23 10

25--60 0.19 0.11 10 0.18 0.38 0.02 10

50--85 0.08 0.03 7 0.08 0.38 0.02 10

85--100 0.04 0.02 4 0.04 0.06 0.02 10

100--120 0.03 0.01 4 0.03 0.04 0.01 10

Organic matter (%) 0--35 10.51 3.78 10 10.38 16.76 5.32 10

25--60 5.30 3.07 10 5.07 10.50 0.44 10

50--85 1.54 1.02 7 1.79 10.50 0.06 10

85--100 0.93 0.57 4 0.81 1.68 0.43 10

100--120 0.56 0.27 4 0.50 0.94 0.31 10

pH (H2O) 0--35 4.76 0.37 10 4.78 5.32 4.05 10

25--60 4.80 0.44 10 4.85 5.37 4.07 10
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

372

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

50--85 4.87 0.50 7 4.86 5.61 4.07 10

85--100 5.02 0.98 4 4.88 6.12 4.18 10

100--120 4.97 0.88 4 5.01 5.80 4.04 10

Phosphorus (mg/kg) 0--35 31 16 10 29 60 13 10

25--60 12 15 10 7 52 1 10

50--85 2 1 7 2 52 1 10

85--100 1 1 4 1 3 1 10

100--120 1 1 4 1 2 1 10

Phosphorus-total (%) 0--35 0.04 0.02 10 0.04 0.06 0.01 10

25--60 0.03 0.01 10 0.02 0.05 0.01 10

50--85 0.02 0.01 7 0.02 0.05 0.01 10

85--100 0.02 0.01 4 0.02 0.03 0.01 10

100--120 0.02 0.01 4 0.01 0.03 0.01 10

Potassium (mg/g) 0--35 0.24 0.08 10 0.27 0.33 0.10 10

25--60 0.15 0.06 10 0.16 0.24 0.06 10

50--85 0.12 0.06 7 0.14 0.24 0.04 10

85--100 0.11 0.05 4 0.10 0.17 0.05 10

100--120 0.10 0.06 4 0.09 0.17 0.05 10

Sodium (mg/g) 0--35 0.15 0.04 10 0.15 0.21 0.07 10

25--60 0.14 0.07 10 0.13 0.25 0.05 10

50--85 0.18 0.09 7 0.14 0.30 0.05 10

85--100 0.19 0.11 4 0.17 0.33 0.10 10

100--120 0.17 0.06 4 0.17 0.23 0.12 10

Serpentinite Oxisols Smf Bulk density (g/cc) 0--18 1.25 1 11 Nipe

Limestone Entisols Smf 0--18 1.52 1 11 St. Lucie

Limestone Oxisols Smf 0--18 1.35 1 11 Coto
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

373

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Limestone Ultisols Smf 0--18 1.29 1 11 Almirante

Sandstone, sillstone Entisols Smf 0--18 1.36 1 11 Corcega

Alluvial deposits Alfisols Smf 0--18 1.58 1 11 Islote

Alluvial deposits Ultisols Smf 0--18 1.76 1 11 Guayabo

Blanket deposits Ultisols Smf 0--18 1.05 1 11 Espinosa

Alluvial deposits Mollisols Smf 0--18 1.35 1 11 Toa

Blanket deposits Ultisols Smf 0--18 1.63 1 11 Sabana Seca

Alluvial deposils Entisols Smf 0--18 1.44 1 11 Cataño

Sandstone, sillstone Entisols Smf 0--18 1.28 1 11 Piñones

Alluvial deposits Mollisols Smf 0--18 1.02 1 11 Colinas

Sandstone, sillstone Inceptisols Smf 0--18 1.24 1 11 Malaya

Plutonic rocks Ultisols Smf 0--18 1.41 1 11 Jayuya

Tuffaceous sandstone Entisols Swf 0--18 1.11 1 11 Fortuna

Lava, tuff Alfisols Smf 0--18 1.18 1 11 Cayagua

Alluvial deposits Alfisols Smf 0--18 1.96 1 11 Dune

Alluvial deposits Mollisols Smf 0--18 0.97 1 11 Ensenada

Blanket deposits Inceptisols Smf 0--18 1.13 1 11 Jaucas

Limestone Vertisols Sdf 0--18 1.35 1 11 Fraternidad

Tuffaceous sandstone Mollisols Smf 0--18 1.26 1 11 Estación

Tuffaceous sandstone Inceptisols Smf 0--18 1.28 1 11 Jacana

Tuffaceous sandstone Ultisols Smf 0--18 1.16 1 11 Cabo Rojo

Tuffaceous sandstone Inceptisols Smf 0--18 1.23 1 11 Lajas

Tuffaceous sandstone Inceptisols Smf 0--18 1.45 1 11 Caguas

Alluvial deposits Entisols Smf 0--18 1.29 1 11 San Germán

Siltstone, sandstone Ultisols Sdf 0--18 1.16 1 11 Alonso

Lava, tuff Ultisols Smf 0--18 1.43 1 11 Ciales
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

374

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Alluvial deposits Alfisols Sdf 0--18 1.68 1 11 Machete

Lava, tuff Inceptisols Sdf 0--18 1.45 1 11 Vives

Tuffaceous, sandstone Alfisols Smf 0--18 1.39 1 11 Via

Serpentinite Oxisols Smf Clay (%) 0--18 27 1 11 Nipe

Limestone Entisols Smf 0--18 6 1 11 St. Lucie

Limestone Oxisols Smf 0--18 25 1 11 Coto

Limestone Ultisols Smf 0--18 40 1 11 Almirante

Sandstone, sillstone Entisols Smf 0--18 27 1 11 Córcega

Alluvial deposits Alfisols Smf 0--18 6 1 11 Islote

Alluvial deposits Ultisols Smf 0--18 21 1 11 Guayabo

Blanket deposits Ultisols Smf 0--18 15 1 11 Espinosa

Alluvial deposits Mollisols Smf 0--18 25 1 11 Toa

Blanket deposits Ultisols Smf 0--18 12 1 11 Sabana Seca

Alluvial deposits Entisols Smf 0--18 5 1 11 Cataño

Sandstone, sillstone Entisols Smf 0--18 14 1 11 Piñones

Alluvial deposits Mollisols Smf 0--18 50 1 11 Colinas

Sandstone, sillstone Inceptisols Smf 0--18 67 1 11 Malaya

Plutonic rocks Ultisols Smf 0--18 20 1 11 Jayuya

Tuffaceous sandstone Entisols Swf 0--18 20 1 11 Fortuna

Lava, tuff Alfisols Smf 0--18 23 1 11 Cayagua

Alluvial deposits Alfisols Smf 0--18 3 1 11 Dune

Alluvial deposits Mollisols Smf 0--18 10 1 11 Ensenada

Blanket deposits Inceptisols Smf 0--18 12 1 11 Jaucas

Limestone Vertisols Sdf 0--18 29 1 11 Fraternidad

Tuffaceous sandstone Mollisols Smf 0--18 24 1 11 Estación

Tuffaceous sandstone Inceptisols Smf 0--18 43 1 11 Jacana
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

375

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Tuffaceous sandstone Ultisols Smf 0--18 17 1 11 Cabo Rojo

Tuffaceous sandstone Inceptisols Smf 0--18 42 1 11 Lajas

Tuffaceous sandstone Inceptisols Smf 0--18 39 1 11 Caguas

Alluvial deposits Entisols Smf 0--18 24 1 11 San Germán

Siltstone, sandstone Ultisols Sdf 0--18 24 1 11 Alonso

Lava, tuff Ultisols Smf 0--18 26 1 11 Ciales

Alluvial deposits Alfisols Sdf 0--18 26 1 11 Machete

Lava, tuff Inceptisols Sdf 0--18 36 1 11 Vives

Tuffaceous, sandstone Alfisols Smf 0--18 21 1 11 Via

Serpentinite Oxisols Smf Organic matter (%) 0--18 4.72 1 11 Nipe

Limestone Entisols Smf 0--18 0.60 1 11 St. Lucie

Limestone Oxisols Smf 0--18 3.59 1 11 Coto

Limestone Ultisols Smf 0--18 4.91 1 11 Almirante

Sandstone, sillstone Entisols Smf 0--18 3.33 1 11 Córcega

Alluvial deposits Alfisols Smf 0--18 1.41 1 11 Islote

Alluvial deposits Ultisols Smf 0--18 1.79 1 11 Guayabo

Blanket deposits Ultisols Smf 0--18 3.06 1 11 Espinosa

Alluvial deposits Mollisols Smf 0--18 2.36 1 11 Toa

Blanket deposits Ultisols Smf 0--18 2.30 1 11 Sabana Seca

Alluvial deposits Entisols Smf 0--18 2.66 1 11 Cataño

Sandstone, sillstone Entisols Smf 0--18 3.54 1 11 Piñones

Alluvial deposits Mollisols Smf 0--18 8.09 1 11 Colinas

Sandstone, sillstone Inceptisols Smf 0--18 3.65 1 11 Malaya

Plutonic rocks Ultisols Smf 0--18 0.76 1 11 Jayuya

Tuffaceous sandstone Entisols Swf 0--18 3.34 1 11 Fortuna

Lava, tuff Alfisols Smf 0--18 2.91 1 11 Cayagua
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

376

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Alluvial deposits Alfisols Smf 0--18 0.35 1 11 Dune

Alluvial deposits Mollisols Smf 0--18 2.10 1 11 Ensenada

Blanket deposits Inceptisols Smf 0--18 2.61 1 11 Jaucas

Limestone Vertisols Sdf 0--18 4.40 1 11 Fraternidad

Tuffaceous sandstone Mollisols Smf 0--18 3.05 1 11 Estación

Tuffaceous sandstone Inceptisols Smf 0--18 4.67 1 11 Jacana

Tuffaceous sandstone Ultisols Smf 0--18 7.51 1 11 Cabo Rojo

Tuffaceous sandstone Inceptisols Smf 0--18 4.60 1 11 Lajas

Tuffaceous sandstone Inceptisols Smf 0--18 3.39 1 11 Caguas

Alluvial deposits Entisols Smf 0--18 4.61 1 11 San Germán

Siltstone, sandstone Ultisols Sdf 0--18 6.42 1 11 Alonso

Lava, tuff Ultisols Smf 0--18 2.55 1 11 Ciales

Alluvial deposits Alfisols Sdf 0--18 1.31 1 11 Machete

Lava, tuff Inceptisols Sdf 0--18 1.81 1 11 Vives

Tuffaceous, sandstone Alfisols Smf 0--18 3.09 1 11 Via

Serpentinite Oxisols Smf pH (H2O) 0--18 5.43 1 11 Nipe

Limestone Entisols Smf 0--18 5.65 1 11 St. Lucie

Limestone Oxisols Smf 0--18 5.57 1 11 Coto

Limestone Ultisols Smf 0--18 5.81 1 11 Almirante

Sandstone, sillstone Entisols Smf 0--18 7.70 1 11 Córcega

Alluvial deposits Alfisols Smf 0--18 7.87 1 11 Islote

Alluvial deposits Ultisols Smf 0--18 5.15 1 11 Guayabo

Blanket deposits Ultisols Smf 0--18 7.67 1 11 Espinosa

Alluvial deposits Mollisols Smf 0--18 5.64 1 11 Toa

Blanket deposits Ultisols Smf 0--18 6.80 1 11 Sabana Seca

Alluvial deposits Entisols Smf 0--18 8.26 1 11 Cataño
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

377

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Sandstone, sillstone Entisols Smf 0--18 8.14 1 11 Piñones

Alluvial deposits Mollisols Smf 0--18 7.26 1 11 Colinas

Sandstone, sillstone Inceptisols Smf 0--18 4.35 1 11 Malaya

Plutonic rocks Ultisols Smf 0--18 6.03 1 11 Jayuya

Tuffaceous sandstone Entisols Swf 0--18 6.87 1 11 Fortuna

Lava, tuff Alfisols Smf 0--18 4.73 1 11 Cayagua

Alluvial deposits Alfisols Smf 0--18 8.86 1 11 Dune

Alluvial deposits Mollisols Smf 0--18 8.23 1 11 Ensenada

Blanket deposits Inceptisols Smf 0--18 8.09 1 11 Jaucas

Limestone Vertisols Sdf 0--18 7.64 1 11 Fraternidad

Tuffaceous sandstone Mollisols Smf 0--18 7.28 1 11 Estación

Tuffaceous sandstone Inceptisols Smf 0--18 6.16 1 11 Jacana

Tuffaceous sandstone Ultisols Smf 0--18 5.82 1 11 Cabo Rojo

Tuffaceous sandstone Inceptisols Smf 0--18 7.21 1 11 Lajas

0--18 6.28 1 11 Caguas

Alluvial deposits Entisols Smf 0--18 6.80 1 11 San Germán

Siltstone, sandstone Ultisols Sdf 0--18 7.22 1 11 Alonso

Lava, tuff Ultisols Smf 0--18 4.99 1 11 Ciales

Alluvial deposits Alfisols Sdf 0--18 5.72 1 11 Machete

Lava, tuff Inceptisols Sdf 0--18 7.29 1 11 Vives

Tuffaceous, sandstone Alfisols Smf 0--18 5.08 1 11 Via

Serpentinite Oxisols Smf Sand (%) 0--18 36 1 11 Nipe

Limestone Entisols Smf 0--18 91 1 11 St. Lucie

Limestone Oxisols Smf 0--18 49 1 11 Coto

Limestone Ultisols Smf 0--18 32 1 11 Almirante

Sandstone, sillstone Entisols Smf 0--18 41 1 11 Córcega
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

378

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Alluvial deposits Alfisols Smf 0--18 83 1 11 Islote

Alluvial deposits Ultisols Smf 0--18 69 1 11 Guayabo

Blanket deposits Ultisols Smf 0--18 66 1 11 Espinosa

Alluvial deposits Mollisols Smf 0--18 22 1 11 Toa

Blanket deposits Ultisols Smf 0--18 74 1 11 Sabana Seca

Alluvial deposits Entisols Smf 0--18 86 1 11 Cataño

Sandstone, sillstone Entisols Smf 0--18 71 1 11 Piñones

Alluvial deposits Mollisols Smf 0--18 18 1 11 Colinas

Sandstone, sillstone Inceptisols Smf 0--18 10 1 11 Malaya

Plutonic rocks Ultisols Smf 0--18 53 1 11 Jayuya

Tuffaceous sandstone Entisols Swf 0--18 22 1 11 Fortuna

Lava, tuff Alfisols Smf 0--18 57 1 11 Cayagua

Alluvial deposits Alfisols Smf 0--18 97 1 11 Dune

Alluvial deposits Mollisols Smf 0--18 80 1 11 Ensenada

Blanket deposits Inceptisols Smf 0--18 73 1 11 Jaucas

Limestone Vertisols Sdf 0--18 42 1 11 Fraternidad

Tuffaceous sandstone Mollisols Smf 0--18 36 1 11 Estación

Tuffaceous sandstone Inceptisols Smf 0--18 25 1 11 Jacana

Tuffaceous sandstone Ultisols Smf 0--18 51 1 11 Cabo Rojo

Tuffaceous sandstone Inceptisols Smf 0--18 24 1 11 Lajas

Tuffaceous sandstone Inceptisols Smf 0--18 29 1 11 Caguas

Alluvial deposits Entisols Smf 0--18 39 1 11 San Germán

Siltstone, sandstone Ultisols Sdf 0--18 34 1 11 Alonso

Lava, tuff Ultisols Smf 0--18 52 1 11 Ciales

Alluvial deposits Alfisols Sdf 0--18 41 1 11 Machete

Lava, tuff Inceptisols Sdf 0--18 34 1 11 Vives
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

379

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Tuffaceous, sandstone Alfisols Smf 0--18 43 1 11 Via

Serpentinite Oxisols Smf Silt (%) 0--18 37 1 11 Nipe

Limestone Entisols Smf 0--18 3 1 11 St. Lucie

Limestone Oxisols Smf 0--18 26 1 11 Coto

Limestone Ultisols Smf 0--18 28 1 11 Almirante

Sandstone, sillstone Entisols Smf 0--18 31 1 11 Córcega

Alluvial deposits Alfisols Smf 0--18 11 1 11 Islote

Alluvial deposits Ultisols Smf 0--18 10 1 11 Guayabo

Blanket deposits Ultisols Smf 0--18 19 1 11 Espinosa

Alluvial deposits Mollisols Smf 0--18 53 1 11 Toa

Blanket deposits Ultisols Smf 0--18 15 1 11 Sabana Seca

Alluvial deposits Entisols Smf 0--18 9 1 11 Cataño

Sandstone, sillstone Entisols Smf 0--18 15 1 11 Piñones

Alluvial deposits Mollisols Smf 0--18 32 1 11 Colinas

Sandstone, sillstone Inceptisols Smf 0--18 23 1 11 Malaya

Plutonic rocks Ultisols Smf 0--18 28 1 11 Jayuya

Tuffaceous sandstone Entisols Swf 0--18 58 1 11 Fortuna

Lava, tuff Alfisols Smf 0--18 20 1 11 Cayagua

Alluvial deposits Alfisols Smf 0--18 0 1 11 Dune

Alluvial deposits Mollisols Smf 0--18 11 1 11 Ensenada

Blanket deposits Inceptisols Smf 0--18 15 1 11 Jaucas

Limestone Vertisols Sdf 0--18 29 1 11 Fraternidad

Tuffaceous sandstone Mollisols Smf 0--18 40 1 11 Estación

Tuffaceous sandstone Inceptisols Smf 0--18 32 1 11 Jacana

Tuffaceous sandstone Ultisols Smf 0--18 32 1 11 Cabo Rojo

Tuffaceous sandstone Inceptisols Smf 0--18 34 1 11 Lajas
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

380

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Tuffaceous sandstone Inceptisols Smf 0--18 33 1 11 Caguas

Alluvial deposits Entisols Smf 0--18 37 1 11 San Germán

Siltstone, sandstone Ultisols Sdf 0--18 42 1 11 Alonso

Lava, tuff Ultisols Smf 0--18 23 1 11 Ciales

Alluvial deposits Alfisols Sdf 0--18 33 1 11 Machete

Lava, tuff Inceptisols Sdf 0--18 30 1 11 Vives

Tuffaceous, sandstone Alfisols Smf 0--18 35 1 11 Via

Tuffaceous, sandstone Ultisols Swf Aluminum (cmol/kg-1) 0--10 0.30 0.21 48 0.25 0.82 0.02 12 Landslide

Calcium (mg/g) 0--10 0.83 0.45 48 0.79 1.51 0.05 12 Landslide

ECEC (cmol/kg-1) 0--10 9.08 3.76 48 9.82 14.85 1.30 12 Landslide

Iron (mg/g) 0--10 0.18 0.12 48 0.14 0.65 0.06 12 Landslide

Magnesium (mg/g) 0--10 0.48 0.23 48 0.50 0.86 0.02 12 Landslide

Manganese (mg/g) 0--10 0.21 0.11 48 0.20 0.52 0.04 12 Landslide

Phosphorus (mg/kg) 0--10 9 8 48 6 39 1 12 Landslide

Potassium (mg/g) 0--10 0.13 0.09 48 0.10 0.43 0.03 12 Landslide

Sodium (mg/g) 0--10 0.08 0.03 48 0.08 0.15 0.04 12 Landslide

Swamps Inceptisols Smf Bulk density (g/cc) 0--7.6 1.33 0.32 2 1.33 1.56 1.11 13 Humacao

Swamps Ultisols Smf 0--7.6 0.96 1 13 Cienaga Alta

Tuffaceous sandstone Ultisols Swf 0--7.6 1.12 1 13 El Verde

Tuffaceous sandstone Ultisols Lmwf 0--7.6 1.21 1 13 Villalba

0--7.6 0.93 1 13 Lares

Sandstone, sillstone Ultisols Swf 0--7.6 1.29 1 13 San Sebastián

Plutonic rocks Ultisols Smf 0--7.6 0.69 1 13 Jayuya

Limestone Mollisols Smf 0--7.6 0.70 1 13 Rio Abajo

Swamps Inceptisols Smf Clay (%) 0--7.6 29 12 2 29 37 20 13 Humacao

Swamps Ultisols Lmrf 0--7.6 35 1 13 Cienaga Alta
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

381

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Tuffaceous sandstone Ultisols Swf 0--7.6 50 1 13 El verde

Tuffaceous sandstone Ultisols Lmwf 0--7.6 42 19 2 42 56 29 13 Villalba

0--7.6 47 1 13 Lares

Sandstone, sillstone Ultisols Swf 0--7.6 29 1 13 San Sebastián

Plutonic rocks Ultisols Smf 0--7.6 68 1 13 Jayuya

Limestone Mollisols Smf 0--7.6 60 1 13 Rio Abajo

Swamps Inceptisols Smf Loss on ignition (%) 0--7.6 8.07 2.88 2 8.07 10.10 6.03 13 Humacao

Swamps Ultisols Lmrf 0--7.6 18.54 1 13 Cienaga Alta

Tuffaceous sandstone Ultisols Swf 0--7.6 12.58 1 13 El Verde

Tuffaceous sandstone Ultisols Lmwf 0--7.6 10.15 4.50 2 10.15 13.33 6.96 13 Villalba

0--7.6 7.36 1 13 Lares

Sandstone, sillstone Ultisols Swf 0--7.6 11.57 1 13 San Sebastián

Plutonic rocks Ultisols Smf 0--7.6 18.87 1 13 Jayuya

Limestone Mollisols Smf 0--7.6 12.69 1 13 Rio Abajo

Swamps Inceptisols Smf Organic matter (%) 0--7.6 5.08 2.05 2 5.08 6.53 3.63 13 Humacao

Swamps Ultisols Lmrf 0--7.6 7.39 1 13 Cienaga Alta

Tuffaceous sandstone Ultisols Swf 0--7.6 6.33 1 13 El Verde

Tuffaceous sandstone Ultisols Lmwf 0--7.6 6.17 2.40 2 6.17 7.86 4.47 13 Villalba

Tuffaceous sandstone Ultisols Lmwf 0--7.6 4.50 1 13 Lares

Sandstone, sillstone Ultisols Swf 0--7.6 7.40 1 13 San Sebastián

Plutonic rocks Ultisols Smf 0--7.6 11.09 1 13 Jayuya

Limestone Mollisols Smf 0--7.6 6.04 1 13 Rio Abajo

Swamps Inceptisols Smf pH (H2O) 0--7.6 5.50 0.14 2 5.50 5.60 5.40 13 Humacao

Swamps Ultisols Lmrf 0--7.6 5.14 1 13 Cienaga Alta

Tuffaceous sandstone Ultisols Swf 0--7.6 5.16 1 13 El Verde

Tuffaceous sandstone Ultisols Lmwf 0--7.6 7.00 0.75 2 7.00 7.53 6.47 13 Villalba
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

382

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Tuffaceous sandstone Ultisols Lmwf 0--7.6 4.40 1 13 Lares

Sandstone, sillstone Ultisols Swf 0--7.6 6.60 1 13 San Sebastián

Plutonic rocks Ultisols Smf 0--7.6 4.30 1 13 Jayuya

Limestone Mollisols Smf 0--7.6 5.07 1 13 Rio Abajo

Swamps Inceptisols Smf Phosphorus (mg/kg) 0--7.6 9 6 2 9 13 4 13 Humacao

Swamps Ultisols Lmrf 0--7.6 4 1 13 Cienaga Alta

Tuffaceous sandstone Ultisols Swf 0--7.6 2 1 13 El Verde

Tuffaceoussandstone Ultisols Lmwf 0--7.6 28 17 2 28 41 16 13 Villalba

0--7.6 7 1 13 Lares

Sandstone, sillstone Ultisols Swf 0--7.6 8 1 13 San Sebastián

Plutonic rocks Ultisols Smf 0--7.6 4 1 13 Jayuya

Limestone Mollisols Smf 0--7.6 5 1 13 Rio Abajo

Swamps Inceptisols Smf Phosphorus-total (%) 0--7.6 0.03 0.01 2 0.03 0.04 0.03 13 Humacao

Swamps Ultisols Lmrf 0--7.6 0.07 1 13 Cienaga Alta

Tuffaceous sandstone Ultisols Swf 0--7.6 0.04 1 13 El Verde

Tuffaceous sandstone Ultisols Lmwf 0--7.6 0.07 0.00 2 0.07 0.08 0.07 13 Villalba

Tuffaceous sandstone Ultisols Lmwf 0--7.6 0.03 1 13 Lares

Sandstone, sillstone Ultisols Swf 0--7.6 0.03 1 13 San Sebastián

Plutonic rocks Ultisols Smf 0--7.6 0.09 1 13 Jayuya

Limestone Mollisols Smf 0--7.6 0.05 1 13 Rio Abajo

Swamps Inceptisols Smf Sand (%) 0--7.6 41 4 2 41 44 38 13 Humacao

Swamps Ultisols Lmrf 0--7.6 41 1 13 Cienaga Alta

Tuffaceous sandstone Ultisols Swf 0--7.6 26 1 13 El Verde

Tuffaceous sandstone Ultisols Lmwf 0--7.6 35 22 2 35 51 20 13 Villalba

Tuffaceous sandstone Ultisols Lmwf 0--7.6 27 1 13 Lares

Sandstone, sillstone Ultisols Swf 0--7.6 27 1 13 San Sebastián
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

383

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Plutonic rocks Ultisols Smf 0--7.6 12 1 13 Jayuya

Limestone Mollisols Smf 0--7.6 14 1 13 Rio Abajo

Swamps Inceptisols Smf Silt (%) 0--7.6 30 8 2 30 36 25 13 Humacao

Swamps Ultisols Lmrf 0--7.6 25 1 13 Cienaga Alta

Tuffaceous sandstone Ultisols Swf 0--7.6 25 1 13 El Verde

Tuffaceous sandstone Ultisols Lmwf 0--7.6 22 3 2 22 24 20 13 Villalba

Tuffaceous sandstone Ultisols Lmwf 0--7.6 26 1 13 Lares

Sandstone, sillstone Ultisols Swf 0--7.6 44 1 13 San Sebastián

Plutonic rocks Ultisols Smf 0--7.6 20 1 13 Jayuya

Limestone Mollisols Smf 0--7.6 26 1 13 Rlo Abajo

Limestone Ultisols Smf Aluminum (cmol/kg-1) 0-12 0.21 0.04 3 0.23 0.24 0.16 14 Guajataca Forest

Blanket deposits Alfisols Smf 0-30 0.12 1 14 Cambalache Forest

Alluvial deposits Vertisols Sdf 0-30 0.06 1 14 Fort Allen, Juana Díaz

Limestone Ultisols Smf 0-30 0.08 0.05 8 0.07 0.14 0.04 14 Guajataca Forest

Alluvial deposits Mollisols Sdf 0-30 0.11 1 14 Guánica Forest

Swamps Mollisols Sdf 0-30 0.13 0.02 2 0.13 0.14 0.11 14 Guayanilla forest

Limestone Ultisols Swf 0-30 0.17 1 14 Rio Abajo forest

Tuffaceous sandstone Ultisols Smf 0-30 0.11 0.01 3 0.11 0.12 0.10 14 Susua Forest

Sandstone, sillstone Alfisols Smf 0-30 0.09 1 14 Vieques

Limestone Ultisols Smf Calcium (mg/g) 0-12 6.56 3.32 3 5.09 10.36 4.23 14 Guajataca Forest

Blanket deposits Alfisols Smf 0-30 2.50 1 14 Cambalache Forest

Alluvial deposits Vertisols Sdf 0-30 4.16 1 14 Fort Allen, Juana Díaz

Limestone Ultisols Smf 0-30 4.88 1.63 8 4.92 6.97 2.62 14 Guajataca Forest

Alluvial deposits Mollisols Sdf 0-30 5.14 1 14 Guánica Forest

Swamps Mollisols Sdf 0-30 9.18 0.72 2 9.18 9.69 8.67 14 Guayanilla Forest
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

384

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Limestone Ultisols Swf 0-30 10.99 1 14 Rio Abajo Forest

Tuffaceous sandstone Ultisols Smf 0-30 1.27 0.29 3 1.43 1.44 0.94 14 Susua Forest

Sandstone, sillstone Alfisols Smf 0-30 1.18 1 14 Vieques

Limestone Ultisols Smf Clay (%) 0-12 18 12 3 18 31 7 14 Guajataca Forest

Blanket deposits Alfisols Smf 0-30 23 1 14 Cambalache Forest

Alluvial deposits Vertisols Sdf 0-30 40 1 14 Fon Allen, Juana Díaz

Limestone Ultisols Smf 0-30 38 20 8 38 73 8 14 Guajataca Forest

Alluvial deposits Mollisols Sdf 0-30 5 1 14 Guánica Forest

Swamps Mollisols Sdf 0-30 34 39 2 34 61 7 14 Guayanilla Forest

Lava, tuff Ultisols Swf 0-30 18 1 14 Rio Abajo Forest

Tuffaceous sandstone Ultisols Smf 0-30 53 19 3 62 66 31 14 Susua Forest

Sandstone, sillstone Alfisols Smf 0-30 23 1 14 Vieques

Limestone Ultisols Smf ECEC (cmol/kg-1) 0-12 38.49 18.68 3 30.39 59.85 25.24 14 Guajataca Forest

Blanket deposits Alfisols Smf 0-30 14.98 1 14 Cambalache Forest

Alluvial deposits Vertisols Sdf 0-30 38.44 1 14 Fort Allen, Juana Díaz

Limestone Ultisols Smf 0-30 32.26 13.58 8 39.67 40.51 16.59 14 Guajataca Forest

Alluvial deposits Mollisols Sdf 0-30 28.95 1 14 Guánica Forest

Swamps Mollisols Sdf 0-30 60.89 7.11 2 60.89 65.91 55.86 14 Guayanilla Forest

Limestone Ultisols Swf 0-30 58.16 1 14 Rio Abajo Forest

Tuffaceous sandstone Ultisols Smf 0-30 54.60 20.53 3 63.19 69.44 31.17 14 Susua Forest

Sandstone, sillstone Alfisols Smf 0-30 9.29 1 14 Vieques

Limestone Ultisols Smf Iron (mg/g) 0-12 0.10 0.12 3 0.06 0.24 0.01 14 Guajataca Forest

Blanketdeposits Alfisols Smf 0-30 0.16 1 14 Cambalache Forest

Alluvial deposits Vertisols Sdf 0-30 0.01 1 14 Fort Allen, Juana Díaz

Limestone Ultisols Smf 0-30 0.14 0.12 8 0.09 0.39 0.03 14 Guajataca Forest

Alluvial deposits Mollisols Sdf 0-30 0.02 1 14 Guánica Forest
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

385

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Swamps Mollisols Sdf 0-30 0.05 0.06 2 0.05 0.09 0.01 14 Guayanilla Forest

Limestone Ultisols Swf 0-30 0.04 1 14 Rio Abajo Forest

Tuffaceous sandstone Ultisols Smf 0-30 0.16 0.01 3 0.16 0.17 0.15 14 Susua Forest

Sandstone, sillstone Alfisols Smf 0-30 0.20 1 14 Vieques

Limestone Ultisols Smf Magnesium (mg/g) 0-12 0.53 0.21 3 0.45 0.77 0.36 14 Guajataca Forest

Blanket deposits Alfisols Smf 0-30 0.23 1 14 Cambalache Forest

Alluvial deposits Vertisols Sdf 0-30 1.71 1 14 Fort Allen, Juana Díaz

Limestone Ultisols Smf 0-30 0.30 0.17 8 0.33 0.54 0.12 14 Guajataca Forest

Alluvial deposits Mollisols Sdf 0-30 0.25 1 14 Guánica Forest

Swamps Mollisols Sdf 0-30 1.59 1.32 2 1.59 2.52 0.65 14 Guayanilla Forest

Limestone Ultisols Swf 0-30 0.24 1 14 Rio Abajo Forest

Tuffaceous sandstone Ultisols Smf 0-30 5.71 2.56 3 6.64 7.68 2.81 14 Susua Forest

Sandstone, sillstone Alfisols Smf 0-30 0.28 1 14 Vieques

Limestone Ultisols Smf Manganese (mg/g) 0-12 0.21 0.11 3 0.22 0.31 0.09 14 Guajataca Forest

Blanket deposits Alfisols Smf 0-30 0.35 1 14 Cambalache Forest

Alluvial deposits Vertisols Sdf 0-30 0.04 1 14 Fort Allen, Juana Díaz

Limestone Ultisols Smf 0-30 0.09 0.05 8 0.09 0.17 0.01 14 Guajataca Forest

Limestone Ultisols Smf 0-30 0.05 1 14 Guajataca Forest

Swamps Mollisols Sdf 0-30 0.07 0.00 2 0.07 0.07 0.07 14 Guayanilla Forest

Limestone Ultisols Swf 0-30 0.03 1 14 Rio Abajo Forest

Tuffaceous sandstone Ultisols Smf 0-30 0.13 0.03 3 0.15 0.15 0.10 14 Susua Forest

Sandstone, sillstone Alfisols Smf 0-30 0.10 1 14 Vieques

Limestone Ultisols Smf Organic matter (%) 0-12 14.76 5.43 3 12.14 21.01 11.14 14 Guajataca Forest

Blanket deposits Alfisols Smf 0-30 5.01 1 14 Cambalache Forest

Alluvial deposits Vertisols Sdf 0-30 3.14 1 14 Fort Allen, Juana Díaz

Limestone Ultisols Smf 0-30 6.62 1.40 8 6.43 8.74 5.00 14 Guajataca Forest
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

386

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Limestone Ultisols Smf 0-30 5.82 1 14 Guajataca Forest

Swamps Mollisols Sdf 0-30 5.70 0.93 2 5.70 6.36 5.04 14 Guayanilla Forest

Limestone Ultisols Swf 0-30 16.73 1 14 Rio Abajo Forest

Tuffaceous sandstone Ultisols Smf 0-30 5.10 1.11 3 5.19 6.17 3.95 14 Susua Forest

Sandstone, sillstone Alfisols Smf 0-30 2.10 1 14 Vieques

Limestone Ultisols Smf pH (H2O) 0-12 6.23 0.72 3 6.61 6.69 5.40 14 Guajataca Forest

Blanket deposits Alfisols Smf 0-30 6.13 1 14 Cambalache Forest

Alluvial deposits Vertisols Sdf 0-30 8.05 1 14 Fon Allen, Juana Díaz

Limestone Ultisols Smf 0-30 6.57 0.72 8 6.59 7.80 5.40 14 Guajataca Forest

Alluvial deposits Mollisols Sdf 0-30 7.86 1 14 Guánica Forest

Swamps Mollisols Sdf 0-30 6.89 1.06 2 6.89 7.64 6.14 14 Guayanilla Forest

Lava, tuff Ultisols Swf 0-30 7.02 1 14 Rio Abajo Forest

Tuffaceous sandstone Ultisols Smf 0-30 6.20 0.05 3 6.22 6.23 6.14 14 Susua Forest

Sandstone, sillstone Alfisols Smf 0-30 5.89 1 14 Vieques

Limestone Ultisols Smf pH (KCl) 0-12 5.58 0.81 3 6.00 6.09 4.65 14 Guajataca Forest

Blanket deposits Alfisols Smf 0-30 5.71 1 14 Cambalache Forest

Alluvial deposits Vertisols Sdf 0-30 6.99 1 14 Fort Allen, Juana Díaz

Limestone Ultisols Smf 0-30 5.93 0.71 8 5.93 7.13 4.79 14 Guajataca Forest

Alluvial deposits Mollisols Sdf 0-30 7.36 1 14 Guánica Forest

Swamps Mollisols Sdf 0-30 6.10 1.16 2 6.10 6.92 5.28 14 Guayanilla Forest

Lava, tuff Ultisols Swf 0-30 6.35 1 14 Rio Abajo Forest

Tuffaceous sandstone Ultisols Smf 0-30 5.25 0.15 3 5.3 5.37 5.09 14 Susua Forest

Sandstone, sillstone Alfisols Smf 0-30 4.69 1 14 Vieques

Limestone Ultisols Smf Phosphorus (mg/kg) 0-12 48 6 3 47 54 43 14 Guajataca Forest

Blanket deposits Alfisols Smf 0-30 38 1 14 Cambalache Forest

Alluvial deposits Vertisols Sdf 0-30 63 1 14 Fort Allen, Juana Díaz
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

387

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Limestone Ultisols Smf 0-30 48 12 8 49 71 29 14 Guajataca Forest

Alluvial deposits Mollisols Sdf 0-30 48 1 14 Guánica Forest

Swamps Mollisols Sdf 0-30 58 3 2 58 60 56 14 Guayanilla Forest

Limestone Ultisols Swf 0-30 95 1 14 Rio Abajo Forest

Tuffaceous sandstone Ultisols Smf 0-30 21 5 3 21 26 16 14 Susua Forest

Sandstone, sillstone Alfisols Smf 0-30 39 1 14 Vieques

Limestone Ultisols Smf Potassium (mg/g) 0-12 0.15 0.06 3 0.14 0.21 0.10 14 Guajataca Forest

Blanket deposits Alfisols Smf 0-30 0.08 1 14 Cambalache Forest

Alluvial deposits Vertisols Sdf 0-30 0.42 1 14 Fort Allen, Juana Díaz

Limestone Ultisols Smf 0-30 0.14 0.08 8 0.12 0.32 0.09 14 Guajataca Forest

Alluvial deposits Mollisols Sdf 0-30 0.27 1 14 Guánica Forest

Swamps Mollisols Sdf 0-30 0.46 0.18 2 0.46 0.59 0.33 14 Guayanilla forest

Limestone Ultisols Swf 0-30 0.21 1 14 Rio Abajo Forest

Tuffaceous sandstone Ultisols Smf 0-30 0.12 0.04 3 0.11 0.16 0.09 14 Susua Forest

Sandstone, sillstone Alfisols Smf 0-30 0.15 1 14 Vieques

Limestone Ultisols Smf Sand (%) 0-12 43 2 3 43 45 42 14 Guajataca Forest

Blanket deposits Alfisols Smf 0-30 46 1 14 Cambalache Forest

Alluvial deposits Vertisols Sdf 0-30 22 1 14 Fon Allen, Juana Díaz

Limestone Ultisols Smf 0-30 24 11 8 25 35 3 14 Guajataca Forest

Alluvial deposits Mollisols Sdf 0-30 27 1 14 Guánica Forest

Swamps Mollisols Sdf 0-30 19 8 2 19 25 14 14 Guayanilla Forest

Lava, tuff Ultisols Swf 0-30 59 1 14 Rio Abajo Forest

Tuffaceous sandstone Ultisols Smf 0-30 24 22 3 13 49 9 14 Susua Forest

Sandstone, sillstone Alfisols Smf 0-30 61 1 14 Vieques

Limestone Ultisols Smf Silt (%) 0-12 38 14 3 40 51 24 14 Guajataca Forest

Blanket deposits Alfisols Smf 0-30 30 1 14 Cambalache Forest
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

388

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Alluvial deposits Vertisols Sdf 0-30 38 1 14 Fon Allen, Juana Díaz

Limestone Ultisols Smf 0-30 39 17 8 39 77 23 14 Guajataca Forest

Alluvial deposits Mollisols Sdf 0-30 68 1 14 Guánica Forest

Swamps Mollisols Sdf 0-30 47 31 2 47 68 25 14 Guayanilla Forest

Lava, tuff Ultisols Swf 0-30 23 1 14 Rio Abajo Forest

Tuffaceous sandstone Ultisols Smf 0-30 23 3 3 25 25 20 14 Susua Forest

Sandstone, sillstone Alfisols Smf 0-30 17 1 14 Vieques

Limestone Ultisols Smf Sodium (mg/g) 0-12 0.17 0.04 3 0.16 0.21 0.14 14 Guajataca Forest

Blanket deposits Alfisols Smf 0-30 0.06 1 14 Cambalache Forest

Alluvial deposits Vertisols Sdf 0-30 0.52 1 14 Fort Allen, Juana Díaz

Limestone Ultisols Smf 0-30 0.11 0.05 8 0.12 0.17 0.05 14 Guajataca Forest

Limestone Ultisols Smf 0-30 0.08 1 14 Guajataca Forest

Swamps Mollisols Sdf 0-30 0.12 0.06 2 0.12 0.16 0.07 14 Guayanilla Forest

Limestone Ultisols Swf 0-30 0.11 1 14 Rio Abajo Forest

Tuffaceous sandstone Ultisols Smf 0-30 0.10 0.02 3 0.11 0.12 0.08 14 Susua Forest

Sandstone, sillstone Alfisols Smf 0-30 0.13 1 14 Vieques

Tuffaceous sandstone Ultisols Lmrf Aluminum (cmol/kg-l) 0-10 1.35 0.49 3 1.12 1.91 1.01 15 TR-l

0-15 0.51 0.49 2 0.51 0.85 0.16 15 Young Secondary Forest

15-30 1.68 2.29 2 1.68 3.30 0.06 15 Young Secondary Forest

30-35 1.85 1.32 3 1.82 3.18 0.55 15 TRl

30-50 1.09 1.39 2 1.09 2.07 0.10 15 Young Secondary Forest

50-75 2.18 0.98 3 2.12 3.18 1.23 15 TR-l

Bulk density (g/cc) 0-10 0.73 0.15 3 0.65 0.90 0.63 15 TR-l

0-15 0.64 0.05 2 0.64 0.67 0.60 15 Young Secondary Forest

15-30 0.56 0.21 2 0.56 0.70 0.41 15 Young Secondary Forest

30-35 0.91 0.25 3 0.81 1.20 0.73 15 TRl
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

389

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

30-50 0.84 0.27 2 0.84 1.03 0.65 15 Young Secondary Forest

50-75 1.01 0.00 3 1.03 1.04 0.96 15 TR-I

Calcium (mg/g) 0-10 0.16 0.04 3 0.17 0.19 0.11 15 TR-l

0-15 0.92 1.00 2 0.92 1.63 0.22 15 Young Secondary Forest

15-30 1.52 0.27 2 1.52 1.71 1.33 15 Young Secondary Forest

30-35 0.06 0.03 3 0.06 0.09 0.02 15 TRl

30-50 0.85 0.83 2 0.85 1.44 0.26 15 Young Secondary Forest

50-75 0.11 0.10 3 0.07 0.23 0.03 15 TR-l

Clay (%) 0-10 34 9 3 36 42 25 15 TR-l

0-15 38 1 2 38 38 37 15 Young Secondary Forest

15-30 41 23 2 41 57 25 15 Young Secondary Forest

30-35 25 1 3 25 26 24 15 TRl

30-50 45 25 2 45 63 27 15 Young Secondary Forest

50-75 27 7 3 25 34 21 15 TR-l

ECEC (cmol/kg-1) 0-10 3.02 0.31 3 3.12 3.27 2.68 15 TR-l

0-15 12.26 10.15 2 12.26 19.44 5.08 15 Young Secondary Forest

15-30 19.38 4.96 2 19.38 22.88 15.87 15 Young Secondary Forest

30-35 2.70 0.92 3 2.83 3.54 1.72 15 TRl

30-50 12.31 9.41 2 12.31 18.96 5.65 15 Young Secondary Forest

50-75 3.52 0.48 3 3.65 3.93 2.99 15 TR-l

Iron (mg/g) 0-10 1.23 0.11 3 1.29 1.29 1.10 15 TR-l

0-15 1.40 0.45 2 1.40 1.71 1.08 15 Young Secondary Forest

15-30 0.19 0.14 2 0.19 0.30 0.09 15 Young Secondary Forest

30-35 0.44 0.47 3 0.19 0.98 0.15 15 TRl

30-50 0.14 0.12 2 0.14 0.23 0.06 15 Young Secondary Forest

50-75 0.11 0.05 3 0.09 0.17 0.07 15 TR-l
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

390

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Magnesium (mg/g) 0-10 0.07 0.03 3 0.06 0.11 0.05 15 TR-l

0-15 0.58 0.65 2 0.58 1.04 0.12 15 Young Secondary Forest

15-30 1.11 0.20 2 1.11 1.25 0.96 15 Young Secondary Forest

30-35 0.05 0.03 3 0.05 0.07 0.01 15 TRl

30-50 0.74 0.79 2 0.74 1.30 0.18 15 Young Secondary Forest

50-75 0.08 0.07 3 0.05 0.16 0.02 15 TR-l

Manganese (mg/kg) 0-10 12 7 3 10 20 6 15 TR-l

0-15 196 49 2 196 230 161 15 Young Secondary Forest

15-30 45 35 2 45 70 20 15 Young Secondary Forest

30-35 6 8 3 2 15 2 15 TRl

30-50 41 20 2 41 20 27 15 Young Secondary Forest

50-75 10 7 3 7 17 4 15 TR-l

Organic matter (%) 0-10 3.14 0.12 3 3.08 3.28 3.06 15 TR-l

0-15 8.21 0.03 2 8.21 8.23 8.19 15 Young Secondary Forest

15-30 1.60 1.10 2 1.60 2.37 0.82 15 Young Secondary Forest

30-35 1.59 1.29 3 1.18 3.04 0.55 15 TRl

30-50 1.29 0.60 2 1.29 1.71 0.86 15 Young Secondary Forest

50-75 0.48 0.13 3 0.48 0.61 0.36 15 TR-l

pH (H2O) 0-10 4.80 0.36 3 4.90 5.10 4.40 15 TR-l

0-15 5.30 0.42 2 5.30 5.60 5.00 15 Young Secondary Forest

15-30 5.50 0.28 2 5.50 5.70 5.30 15 Young Secondary Forest

30-35 4.83 0.21 3 4.90 5.00 4.60 15 TRl

30-50 5.70 0.00 2 5.70 5.70 5.70 15 Young Secondary Forest

50-75 5.00 0.10 3 5.00 5.10 4.90 15 TR-l

pH (KCl) 0-10 3.83 0.15 3 3.80 4.00 3.70 15 TR-l

0-15 4.45 0.35 2 4.45 4.70 4.20 15 Young Secondary Forest
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

391

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

15-30 4.20 0.57 2 4.20 4.60 3.80 15 Young Secondary Forest

30-35 3.87 0.21 3 3.80 4.10 3.70 15 TRl

30-50 4.15 0.49 2 4.15 4.50 3.80 15 Young Secondary Forest

50-75 3.87 0.06 3 3.90 3.90 3.80 15 TR-l

Phosphorus (mg/kg) 0-10 10 2 3 10 12 8 15 TR-l

0-15 36 1 2 36 37 35 15 Young Secondary Forest

15-30 9 4 2 9 12 6 15 Young Secondary Forest

30-35 4 3 3 2 7 2 15 TRl

30-50 8 2 2 8 10 7 15 Young Secondary Forest

50-75 1 1 3 2 2 0 15 TR-l

Potassium (mg/kg) 0-10 53 11 3 56 62 40 15 TR-l

0-15 688 231 2 688 851 524 15 Young Secondary Forest

15-30 166 114 2 166 247 85 15 Young Secondary Forest

30-35 22 25 3 10 50 5 15 TRl

30-50 98 6 2 98 102 94 15 Young Secondary Forest

50-75 7 2 3 7 10 5 15 TR-l

Sand (%) 0-10 54 7 3 51 62 49 15 TR-l

0-15 16 3 2 16 18 14 15 Young Secondary Forest

15-30 10 3 2 10 12 8 15 Young Secondary Forest

30-35 45 6 3 46 51 39 15 TRl

30-50 13 9 2 13 19 6 15 Young Secondary Forest

50-75 41 7 3 41 47 34 15 TR-l

Silt (%) 0-10 12 2 3 13 13 9 15 TR-l

0-15 49 20 2 49 63 35 15 Young Secondary Forest

15-30 49 20 2 49 63 35 15 Young secondary forest

30-35 30 7 3 29 37 23 15 TRl
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

392

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

30-50 43 16 2 43 54 31 15 Young secondary forest

50-75 33 1 3 32 34 32 15 TR-l

Sodium (mg/kg) 0-10 30 10 3 40 40 20 15 TR-l

0-15 130 30 2 130 150 110 15 Young secondary forest

15-30 110 10 2 110 120 100 15 Young secondary forest

30-35 30 10 3 30 40 30 15 TRl

30-50 130 20 2 130 140 110 15 Young secondary forest

50-75 40 20 3 30 60 30 15 TR-l

Serpentinite Ultisols Lmrf Clay (%) 0-10 24 6 6 24 31 16 16 Forest Keeper/H

0-10 20 5 6 20 27 14 16 Mototrola Tower

10--20 27 11 6 24 46 15 16 Forest Keeper/H

10--20 19 8 6 18 31 10 16 Mototrola Tower

20--30 28 9 6 31 39 14 16 Forest Keeper/H

20--30 27 6 6 28 32 16 16 Mototrola Tower

30-40 31 9 6 29 48 21 16 Forest Keeper/H

30-40 27 5 6 27 34 19 16 Mototrola Tower

Organic matter (%) 0-10 7.33 0.50 6 7.30 8.10 6.68 16 Forest Keeper/H

0-10 8.88 1.77 6 8.87 11.25 6.07 16 Mototrola Tower

10--20 6.12 1.40 6 6.08 8.01 4.72 16 Forest Keeper/H

10--20 6.31 0.69 6 6.01 7.19 5.55 16 Mototrola Tower

20--30 4.63 1.14 6 4.82 6.01 2.96 16 Forest Keeper/H

20--30 4.71 0.90 6 4.78 5.97 3.20 16 Mototrola Tower

30-40 3.82 0.57 6 3.96 4.46 2.93 16 Forest Keeper/H

30-40 3.99 0.32 6 3.95 4.57 3.62 16 Mototrola Tower

Sand (%) 0-10 59 7 6 60 67 50 16 Forest Keeper/H

0-10 59 7 6 60 67 50 16 Mototrola Tower
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

393

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--20 64 11 6 67 78 50 16 Forest Keeper~I

10--20 64 11 6 67 78 50 16 Mototrola Tower

20--30 54 8 6 54 69 47 16 Forest Keeper/H

20--30 54 8 6 54 69 47 16 Mototrola Tower

30-40 56 7 6 56 67 45 16 Mototrola Tower

30--40 56 7 6 56 67 45 16 Forest Keeper/H

Silt (%) 0-10 29 3 6 28 34 26 16 Forest Keeper/H

0-10 21 4 6 21 26 17 16 Mototrola Tower

10--20 28 8 6 26 41 19 16 Forest Keeper/H

10--20 17 5 6 17 25 12 16 Mototrola Tower

20--30 24 6 6 23 35 18 16 Forest Keeper/H

20--30 19 3 6 19 25 15 16 Mototrola Tower

30-40 19 3 6 20 23 14 16 Forest Keeper/H

30-40 17 3 6 17 21 14 16 Mototrola Tower

Serpentinite Ultisols Lmrf Aluminum (cmol/kg-l) 0--10 0.13 0.05 22 0.12 0.25 0.06 17 Plantation

0--10 0.12 0.04 3 0.13 0.15 0.07 17 Control plot

10--20 0.11 0.05 18 0.10 0.20 0.02 17 Plantation

10--20 0.10 0.05 3 0.08 0.16 0.07 17 Control plot

20--30 0.11 0.04 21 0.11 0.18 0.04 17 Plantation

20--30 0.10 0.04 4 0.08 0.16 0.06 17 Control plot

30--40 0.11 0.05 18 0.11 0.23 0.03 17 Plantation

30--40 0.10 0.05 3 0.08 0.15 0.06 17 Control plot

Calcium (cmol/kg-l) 0--10 7.80 2.14 22 7.89 13.03 3.84 17 Plantation

0--10 5.82 1.53 3 5.06 7.58 4.82 17 Control plot

10--20 7.89 1.84 18 7.74 10.67 3.98 17 Plantation

10--20 5.72 1.65 3 5.32 7.54 4.31 17 Control plot
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

394

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

20--30 7.26 2.47 21 7.43 13.67 3.72 17 Plantation

20--30 5.39 1.01 4 5.05 6.87 4.59 17 Control plot

30--40 6.87 2.18 18 7.15 10.66 3.46 17 Plantation

30--40 4.79 1.50 3 4.09 6.51 3.77 17 Control plot

Calcium (mg/g) 0--10 1.56 0.43 22 1.58 2.61 0.77 17 Plantation

0--10 1.16 0.31 3 1.01 1.52 0.96 17 Control plot

10--20 1.58 0.37 18 1.55 2.13 0.80 17 Plantation

10--20 1.15 0.33 3 1.06 1.51 0.86 17 Control plot

20--30 1.45 0.49 21 1.49 2.73 0.74 17 Plantation

20--30 1.08 0.20 4 1.01 1.37 0.92 17 Control plot

30--40 1.37 0.44 18 1.43 2.13 0.69 17 Plantation

30--40 0.96 0.30 3 0.82 1.30 0.75 17 Control plot

ECEC (cmol/kg-1) 0--10 9.40 2.51 22 9.65 14.30 4.80 17 Plantation

0--10 6.99 2.11 3 5.80 9.42 5.74 17 Control plot

10--20 9.39 2.21 18 9.38 12.85 4.78 17 Plantation

10--20 6.64 2.03 3 6.08 8.90 4.95 17 Control plot

20--30 8.47 2.70 21 8.73 14.76 4.33 17 Plantation

20--30 6.16 1.30 4 5.71 8.07 5.15 17 Control plot

30--40 8.07 2.45 18 8.49 11.66 4.18 17 Plantation

30--40 5.28 1.44 3 4.64 6.93 4.28 17 Control plot

Iron (mg/kg) 0--10 25 20 22 17 71 6 17 Plantation

0--10 12 11 3 6 25 5 17 Control plot

10--20 28 22 18 15 67 5 17 Plantation

10--20 8 6 3 6 15 4 17 Control plot

20--30 19 16 21 12 55 2 17 Plantation

20--30 6 2 4 5 9 4 17 Control plot
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

395

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

30--40 17 12 18 13 50 4 17 Plantation

30--40 6 4 3 4 10 3 17 Control plot

Potassium (cmol/kg-l) 0--10 0.12 0.03 22 0.12 0.18 0.06 17 Plantation

0--10 0.14 0.03 3 0.15 0.16 0.10 17 Control plot

10--20 0.10 0.03 18 0.11 0.14 0.05 17 Plantation

10--20 0.09 0.03 3 0.09 0.12 0.07 17 Control plot

20--30 0.08 0.02 21 0.08 0.13 0.05 17 Plantation

20--30 0.08 0.02 4 0.07 0.11 0.07 17 Control plot

30--40 0.08 0.02 18 0.07 0.14 0.04 17 Plantation

30--40 0.07 0.02 3 0.06 0.09 0.05 17 Control plot

Potassium (mg/kg) 0--10 46 12 22 47 68 25 17 Plantation

0--10 53 13 3 58 63 38 17 Control plot

10--20 39 9 18 42 55 21 17 Plantation

10--20 36 9 3 34 46 28 17 Control plot

20--30 31 8 21 29 51 18 17 Plantation

20--30 30 9 4 26 43 26 17 Control plot

30--40 30 9 18 27 53 17 17 Plantation

30--40 25 7 3 22 33 20 17 Control plot

Loss on ignition (%) 0--10 4.63 0.87 18 4.81 5.88 3.13 17 Plantation

0--10 3.95 1.25 3 3.46 5.38 3.02 17 Control plot

10--20 4.39 0.83 18 4.40 5.98 3.27 17 Plantation

10--20 3.36 1.01 3 3.44 4.33 2.32 17 Control plot

20--30 3.89 0.78 18 3.80 5.43 2.70 17 Plantation

20--30 2.96 0.60 3 2.94 3.57 2.38 17 Control plot

30--40 3.64 0.74 18 3.52 5.24 2.47 17 Plantation

30--40 2.81 0.74 3 2.75 3.58 2.10 17 Control plot
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

396

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Magnesium (cmol/kg-l) 0--10 1.15 0.47 22 0.99 2.01 0.54 17 Plantation

0--10 0.83 0.51 3 0.59 1.42 0.49 17 Control plot

10--20 1.07 0.45 18 0.89 1.91 0.56 17 Plantation

10--20 0.65 0.28 3 0.54 0.97 0.44 17 Control plot

20--30 0.84 0.38 21 0.76 1.80 0.36 17 Plantation

20--30 0.53 0.19 4 0.47 0.81 0.37 17 Control plot

30--40 0.82 0.31 18 0.80 1.27 0.44 17 Plantation

30--40 0.27 0.17 3 0.34 0.40 0.08 17 Control plot

Magnesium (mg/g) 0--10 0.14 0.06 22 0.12 0.24 0.06 17 Plantation

0--10 0.10 0.06 3 0.07 0.17 0.06 17 Control plot

10--20 0.13 0.05 18 0.11 0.23 0.07 17 Plantation

10--20 0.08 0.03 3 0.07 0.12 0.05 17 Control plot

20--30 0.10 0.05 21 0.09 0.22 0.04 17 Plantation

20--30 0.06 0.02 4 0.06 0.10 0.05 17 Control plot

30--40 0.10 0.04 18 0.10 0.15 0.05 17 Plantation

30--40 0.03 0.02 3 0.04 0.05 0.01 17 Control plot

Manganese (mg/kg) 0--10 14 6 22 13 28 8 17 Plantation

0--10 11 5 3 10 16 7 17 Control plot

10--20 13 6 18 12 29 6 17 Plantation

10--20 7 2 3 7 9 6 17 Control plot

20--30 10 4 21 10 19 4 17 Plantation

20--30 6 2 4 5 9 4 17 Control plot

30--40 10 4 18 10 17 5 17 Plantation

30--40 5 2 3 4 7 4 17 Control plot

Nitrogen (%) 0--10 0.09 0.02 22 0.10 0.13 0.05 17 Plantation

0--10 0.08 0.03 3 0.07 0.12 0.06 17 Control plot
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

397

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--20 0.10 0.04 18 0.09 0.20 0.05 17 Plantation

10--20 0.07 0.02 3 0.08 0.08 0.04 17 Control plot

20--30 0.08 0.08 21 0.06 0.40 0.02 17 Plantation

20--30 0.05 0.02 4 0.06 0.06 0.03 17 Control plot

30--40 0.05 0.02 18 0.05 0.10 0.03 17 Plantation

30--40 0.04 0.02 3 0.04 0.05 0.02 17 Control plot

Sodium (cmol/kg-l) 0--10 0.21 0.15 22 0.16 0.72 0.07 17 Plantation

0--10 0.08 0.05 3 0.11 0.11 0.03 17 Control plot

10--20 0.22 0.16 18 0.17 0.69 0.08 17 Plantation

10--20 0.08 0.04 3 0.06 0.12 0.05 17 Control plot

20--30 0.19 0.14 21 0.15 0.53 0.05 17 Plantation

20--30 0.07 0.04 4 0.06 0.12 0.04 17 Control plot

30--40 0.20 0.14 18 0.15 0.62 0.06 17 Plantation

30--40 0.05 0.04 3 0.03 0.10 0.03 17 Control plot

Sodium (mg/g) 0--10 0.05 0.03 22 0.04 0.17 0.02 17 Plantation

0--10 0.02 0.01 3 0.03 0.03 0.01 17 Control plot

10--20 0.05 0.04 18 0.04 0.16 0.02 17 Plantation

10--20 0.02 0.01 3 0.01 0.03 0.01 17 Control plot

20--30 0.04 0.03 21 0.03 0.12 0.01 17 Plantation

20--30 0.02 0.01 4 0.01 0.03 0.01 17 Control plot

30--40 0.05 0.03 18 0.03 0.14 0.01 17 Plantation

30--40 0.01 0.01 3 0.01 0.02 0.01 17 Control plot

Phosphorus (mg/kg) 0--10 7 2 22 7 10 4 17 Plantation

0--10 5 3 3 5 8 3 17 Control plot

10--20 7 2 18 6 10 4 17 Plantation

10--20 4 1 3 4 5 3 17 Control plot
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

398

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

20--30 5 2 21 5 9 2 17 Plantation

20--30 3 1 4 3 4 2 17 Control plot

30--40 0 2 18 4 9 3 17 Plantation

30--40 2 1 3 2 3 2 17 Control plot

pH (H2O) 0--10 7.60 0.35 18 7.63 8.09 6.72 17 Plantation

0--10 7.80 0.40 3 7.86 8.16 7.37 17 Control plot

10--20 7.57 0.40 18 7.71 8.06 6.67 17 Plantation

10--20 7.89 0.29 3 7.96 8.14 7.57 17 Control plot

20--30 7.64 0.38 18 7.71 8.16 6.80 17 Plantation

20--30 7.85 0.23 3 7.82 8.10 7.64 17 Control plot

30--40 7.76 0.33 18 7.82 8.15 6.77 17 Plantation

30--40 8.01 0.28 3 8.15 8.19 7.68 17 Control plot

pH (KCl) 0--10 6.73 0.42 18 6.68 7.36 6.00 17 Plantation

0--10 6.99 0.29 3 7.00 7.28 6.70 17 Control plot

10--20 6.72 0.54 18 6.84 7.48 5.76 17 Plantation

10--20 7.07 0.34 3 7.15 7.36 6.69 17 Control plot

20--30 6.84 0.52 18 6.87 7.68 5.99 17 Plantation

20--30 7.22 0.21 3 7.25 7.41 6.99 17 Control plot

30--40 6.93 0.47 18 7.00 7.72 5.62 17 Plantation

30--40 7.35 0.38 3 7.40 7.70 6.95 17 Control plot

Alluvial deposits Inceptisols Dfs Aluminum (cmol/kg-l) 0--10 0.39 0.13 4 0.40 0.52 0.24 18 Control

0--10 0.33 0.12 19 0.30 0.65 0.15 18 Plantation

Carbon (%) 0--10 3.76 0.50 4 3.76 4.36 3.15 18 Control

0--10 3.68 0.52 19 3.85 4.60 2.37 18 Plantation

Calcium (cmol/kg-l) 0--10 40.24 7.51 4 40.96 48.51 30.52 18 Control

0--10 38.17 6.84 19 36.44 50.96 27.07 18 Plantation
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

399

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Calcium (mg/g) 0--10 8.05 1.50 4 8.19 9.70 6.11 18 Control

0--10 7.64 1.37 19 7.29 10.19 5.42 18 Plantation

ECEC (cmol/kg-1) 0--10 51.46 5.82 4 52.48 56.45 44.43 18 Control

0--10 49.97 6.15 19 49.14 62.61 40.43 18 Plantation

Iron (mg/kg) 0--10 13 8 4 14 21 4 18 Control

0--10 13 7 19 12 34 4 18 Plantation

Potassium (cmol/kg-l) 0--10 1.28 0.30 4 1.28 1.60 0.96 18 Control

0--10 1.04 0.18 19 0.98 1.44 0.76 18 Plantation

Potassium (mg/g) 0--10 0.49 0.12 4 0.49 0.61 0.37 18 Control

0--10 0.39 0.07 19 0.37 0.55 0.29 18 Plantation

Magnesium (cmol/kg-l) 0--10 8.58 3.16 4 8.90 11.61 4.92 18 Control

0--10 9.46 2.27 19 10.11 12.37 4.20 18 Plantation

Magnesium (mg/g) 0--10 1.03 0.38 4 1.07 1.39 0.59 18 Control

0--10 1.14 0.27 19 1.21 1.48 0.50 18 Plantation

Manganese (mg/kg) 0--10 28 7 4 29 34 19 18 Control

0--10 35 10 19 37 48 17 18 Plantation

Nitrogen (%) 0--10 0.40 0.05 4 0.40 0.45 0.33 18 Control

0--10 0.41 0.06 19 0.43 0.52 0.28 18 Plantation

Sodium (cmol/kg-l) 0--10 0.97 0.15 4 0.99 1.13 0.78 18 Control

0--10 0.98 0.15 19 0.97 1.34 0.70 18 Plantation

Sodium (mg/g) 0--10 0.22 0.03 4 0.23 0.26 0.18 18 Control

0--10 0.23 0.03 1 0.22 0.31 0.16 18 Plantation

Phosphorus (mg/kg) 0--10 17 9 4 16 29 8 18 Control

0--10 12 3 19 12 21 8 18 Plantation

Sulfur (%) 0--10 0.04 0.01 4 0.05 0.05 0.03 18 Control

Alluvial deposits Inceptisols Dfs 0--10 0.04 0.01 19 0.05 0.06 0.02 18 Plantation
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

400

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Alluvial deposits Histosol Mfs Aluminum (cmol/kg-l) 0--10 0.12 0.05 17 0.12 0.18 0.04 19 Plantation

0--10 0.10 0.06 3 0.12 0.15 0.04 19 Control

10--20 0.08 0.04 20 0.08 0.15 0.03 19 Plantation

10--20 0.07 0.03 4 0.08 0.10 0.03 19 Control

20--30 0.08 0.04 17 0.08 0.15 0.02 19 Plantation

20--30 0.10 0.04 3 0.09 0.14 0.06 19 Control

30--40 0.04 0.03 20 0.03 0.12 0.01 19 Plantation

30--40 0.05 0.01 3 0.04 0.06 0.04 19 Control

Carbon (%) 0--10 1.93 0.35 17 1.84 2.81 1.39 19 Plantation

0--10 2.20 0.22 3 2.30 2.35 1.95 19 Control

10--20 1.56 0.32 20 1.62 2.12 0.69 19 Plantation

10--20 1.63 0.39 4 1.62 2.03 1.24 19 Control

20--30 1.27 0.41 17 1.26 1.83 0.40 19 Plantation

20--30 1.63 0.96 3 1.56 2.62 0.71 19 Control

30--40 1.33 0.79 20 1.19 2.96 0.19 19 Plantation

30--40 1.59 0.85 3 1.21 2.56 1.00 19 Control

Calcium (cmol/kg-l) 0--10 6.81 1.27 17 6.76 10.32 4.51 19 Plantation

0--10 7.72 2.11 3 7.00 10.09 6.06 19 Control

10--20 6.41 1.31 20 6.79 8.63 4.01 19 Plantation

10--20 8.23 1.43 4 8.35 9.85 6.36 19 Control

20--30 5.73 1.57 17 5.80 8.68 3.37 19 Plantation

20--30 6.73 3.13 3 6.68 9.88 3.63 19 Control

30--40 4.79 1.55 20 4.57 8.53 2.56 19 Plantation

30--40 6.19 3.26 3 5.30 9.80 3.46 19 Control

Calcium (mg/g) 0--10 1.36 0.26 17 1.35 2.06 0.90 19 Plantation

0--10 1.54 0.42 3 1.40 2.02 1.21 19 Control
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

401

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--20 1.28 0.26 20 1.36 1.73 0.80 19 Plantation

10--20 1.65 0.29 4 1.67 1.97 1.27 19 Control

20--30 1.15 0.31 7 1.16 1.74 0.67 19 Plantation

20--30 1.35 0.63 3 1.34 1.98 0.73 19 Control

30--40 0.96 0.31 20 0.91 1.71 0.51 19 Plantation

30--40 1.24 0.62 3 1.06 1.96 0.69 19 Control

ECEC (cmol/kg-1) 0--10 9.31 1.58 17 9.34 12.48 6.04 19 Plantation

0--10 9.42 2.86 3 8.38 12.66 7.23 19 Control

10--20 8.46 1.99 20 8.88 11.63 5.13 19 Plantation

10--20 10.23 2.05 4 10.73 12.14 7.32 19 Control

20--30 7.38 2.05 17 7.05 11.45 4.09 19 Plantation

20--30 8.32 3.90 3 8.47 12.14 4.34 19 Control

30--40 6.10 1.95 20 6.09 10.49 3.20 19 Plantation

30--40 7.25 3.68 3 6.33 11.30 4.13 19 Control

Iron (mg/kg) 0--10 28 18 17 23 73 8 19 Plantation

0--10 25 27 3 11 56 9 19 Control

10--20 23 15 20 19 61 5 19 Plantation

10--20 20 22 4 32 50 8 19 Control

20--30 20 15 17 17 52 1 19 Plantation

20--30 17 13 3 18 30 3 19 Control

30--40 15 13 20 12 42 1 19 Plantation

30--40 9 8 3 5 18 3 19 Control

Potassium (cmol/kg-l) 0--10 0.34 0.07 17 0.32 0.57 0.28 19 Plantation

0--10 0.31 0.03 3 0.29 0.35 0.28 19 Control

10--20 0.27 0.03 20 0.26 0.35 0.23 19 Plantation

10--20 0.28 0.02 4 0.29 0.29 0.25 19 Control
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

402

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

20--30 0.24 0.02 17 0.23 0.29 0.21 19 Plantation

20--30 0.26 0.04 3 0.25 0.30 0.22 19 Control

30--40 0.22 0.02 20 0.23 0.25 0.20 19 Plantation

30--40 0.24 0.02 3 0.24 0.25 0.22 19 Control

Potassium (mg/g) 0--10 0.13 0.03 17 0.12 0.22 0.11 19 Plantation

0--10 0.12 0.01 3 0.11 0.13 0.11 19 Control

10--20 0.10 0.01 20 0.10 0.13 0.09 19 Plantation

10--20 0.11 0.01 4 0.11 0.11 0.09 19 Control

20--30 0.09 0.01 17 0.09 0.11 0.08 19 Plantation

20--30 0.10 0.02 3 0.10 0.11 0.08 19 Control

30--40 0.08 0.01 20 0.09 0.10 0.07 19 Plantation

30--40 0.09 0.01 3 0.09 0.10 0.08 19 Control

Loss on ignition (%) 0--10 4.67 0.82 17 4.80 6.04 3.01 19 Plantation

0--10 4.16 1.82 3 3.48 6.22 2.77 19 Control

10--20 4.01 0.87 20 4.25 5.26 2.31 19 Plantation

10--20 4.36 0.65 4 4.56 4.88 3.42 19 Control

20--30 3.46 0.93 17 3.31 5.76 2.10 19 Plantation

20--30 3.17 1.60 3 2.94 4.87 1.71 19 Control

30--40 2.66 0.64 20 2.55 3.79 1.63 19 Plantation

30--40 2.47 1.26 3 2.12 3.86 1.42 19 Control

Magnesium (cmol/kg-l) 0--10 1.68 0.43 17 1.67 2.69 1.05 19 Plantation

0--10 1.18 0.69 3 0.92 1.96 0.66 19 Control

10--20 1.35 0.61 20 1.22 2.49 0.64 19 Plantation

10--20 1.50 0.61 4 1.79 1.83 0.58 19 Control

20--30 0.98 0.40 17 0.92 1.72 0.38 19 Plantation

20--30 1.11 0.71 3 1.21 1.76 0.35 19 Control
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

403

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

30--40 0.72 0.35 20 0.66 1.37 0.22 19 Plantation

30--40 0.66 0.40 3 0.59 1.09 0.31 19 Control

Magnesium (mg/g) 0--10 0.22 0.05 17 0.20 0.32 0.13 19 Plantation

0--10 0.14 0.08 3 0.11 0.24 0.08 19 Control

10--20 0.16 0.07 20 0.15 0.30 0.08 19 Plantation

10--20 0.18 0.07 4 0.22 0.22 0.07 19 Control

20--30 0.12 0.05 17 0.11 0.21 0.05 19 Plantation

20--30 0.13 0.09 3 0.15 0.21 0.04 19 Control

30--40 0.09 0.04 20 0.08 0.17 0.03 19 Plantation

30--40 0.08 0.05 3 0.07 0.13 0.04 19 Control

Manganese (mg/kg) 0--10 22 6 17 22 35 13 19 Plantation

0--10 20 9 3 16 31 13 19 Control

10--20 15 4 20 15 23 9 19 Plantation

10--20 19 8 4 21 26 10 19 Control

20--30 11 4 17 11 20 4 19 Plantation

20--30 12 7 3 15 17 4 19 Control

30--40 9 4 20 9 17 3 19 Plantation

30--40 8 4 3 7 13 5 19 Control

Nitrogen (%) 0--10 0.15 0.03 17 0.15 0.19 0.06 19 Plantation

0--10 0.11 0.08 3 0.09 0.19 0.05 19 Control

10--20 0.11 0.04 20 0.12 0.02 0.04 19 Plantation

10--20 0.12 0.03 4 0.12 0.16 0.08 19 Control

20--30 0.09 0.05 17 0.08 0.17 0.02 19 Plantation

20--30 0.07 0.07 3 0.06 0.14 0.01 19 Control

30--40 0.05 0.03 20 0.04 0.11 0.01 19 Plantation

30--40 0.06 0.07 2 0.06 0.11 0.01 19 Control
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

404

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Sodium (cmol/kg-l) 0--10 0.37 0.25 17 0.29 1.00 0.10 19 Plantation

0--10 0.11 0.01 3 0.11 0.12 0.11 19 Control

10--20 0.36 0.25 20 0.28 1.20 0.09 19 Plantation

10--20 0.15 0.04 4 0.15 0.18 0.10 19 Control

20--30 0.35 0.21 17 0.28 0.92 0.10 19 Plantation

20--30 0.13 0.06 3 0.11 0.20 0.09 19 Control

30--40 0.33 0.20 20 0.24 0.87 0.08 19 Plantation

30--40 0.12 0.02 3 0.12 0.14 0.10 19 Control

Sodium (mg/g) 0--10 0.08 0.06 17 0.07 0.23 0.02 19 Plantation

0--10 0.03 0.00 3 0.03 0.03 0.03 19 Control

10--20 0.08 0.06 20 0.06 0.28 0.02 19 Plantation

10--20 0.03 0.01 4 0.04 0.04 0.02 19 Control

20--30 0.08 0.05 17 0.07 0.21 0.02 19 Plantation

20--30 0.03 0.01 3 0.03 0.05 0.02 19 Control

30--40 0.08 0.05 20 0.06 0.20 0.02 19 Plantation

30--40 0.03 0.01 3 0.03 0.03 0.02 19 Control

Phosphorus (mg/kg) 0--10 8 2 17 8 12 5 19 Plantation

0--10 9 4 3 9 12 5 19 Control

10--20 6 1 20 6 10 4 19 Plantation

10--20 8 2 4 8 9 5 19 Control

20--30 4 1 17 4 6 2 19 Plantation

20--30 5 3 3 5 9 2 19 Control

30--40 3 1 20 3 5 2 19 Plantation

30--40 3 2 3 2 6 2 19 Control

pH (H2O) 0--10 7.65 0.42 17 7.83 8.20 6.81 19 Plantation

0--10 7.75 0.68 3 8.09 8.20 6.97 19 Control
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

405

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--20 7.69 0.53 20 7.82 8.21 6.11 19 Plantation

10--20 7.48 0.61 4 7.45 8.08 6.93 19 Control

20--30 7.66 0.45 17 7.86 8.16 6.72 19 Plantation

20--30 7.90 0.32 3 7.92 8.20 7.57 19 Control

30--40 7.88 0.48 20 8.01 8.24 6.31 19 Plantation

30--40 8.10 0.21 3 8.19 8.26 7.86 19 Control

pH (KCl) 0--10 6.98 0.45 17 7.09 7.42 5.57 19 Plantation

0--10 6.97 0.84 3 7.40 7.51 6.00 19 Control

10--20 6.98 0.55 20 7.21 7.52 5.50 19 Plantation

10--20 6.55 0.93 4 6.49 7.49 5.75 19 Control

20--30 6.90 0.51 17 6.95 7.62 5.77 19 Plantation

20--30 7.16 0.48 3 7.18 7.63 6.68 19 Control

30--40 7.19 0.53 20 7.34 7.72 5.72 19 Plantation

30--40 7.41 0.19 3 7.35 7.62 7.26 19 Control

Sulfur (mg/kg) 0--10 170 60 17 170 320 100 19 Plantation

0--10 150 110 3 120 270 60 19 Control

10--20 140 60 17 110 320 70 19 Plantation

10--20 130 70 4 140 200 40 19 Control

20--30 160 80 17 110 360 70 19 Plantation

20--30 140 60 3 110 200 100 19 Control

30--40 140 50 18 140 250 60 19 Plantation

30--40 70 30 3 70 90 40 19 Control

Alluvial deposits Mollisols Sdf C/N 0--10 17 5 35 18 28 18 20 Guanica Forest

Carbon (%) 0--10 11.47 4.49 35 12.27 23.00 4.87 20 Guanica Forest

Nitrogen (%) 0--10 0.68 0.28 35 0.59 1.16 0.40 20 Guanica Forest

Sulfur (%) 0--10 0.03 0.02 35 0.02 0.09 0.01 20 Guanica Forest
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

406

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Tuffaceous sandstone Ultisols Swf C/N 0--10 15 2 18 15 20 12 21 El Verde

0--10 11 1 61 11 15 8 21 Mameyes

0--10 13 2 71 13 18 10 21 Sabana

Carbon (%) 0-10 9.93 4.28 18 8.87 19.50 5.06 21 El Verde

0--10 4.98 1.37 61 4.95 8.49 2.35 21 Mameyes

0--10 5.55 1.78 71 5.17 14.21 2.78 21 Sabana

Nitrogen (%) 0--10 0.63 0.18 18 0.57 0.99 0.39 21 El Verde

0--10 0.46 0.09 61 0.46 0.68 0.29 21 Mameyes

0--10 0.42 0.10 71 0.47 0.87 0.24 21 Sabana

Sulfur (%) 0--10 0.09 0.02 18 0.09 0.13 0.06 21 El Verde

0--10 0.05 0.01 61 0.05 0.08 0.03 21 Mameyes

0--10 0.07 0.01 71 0.07 0.10 0.05 21 Sabana

Tuffaceous sandstone Ultisols Lmrf Aluminum (mg/kg) 0--10 90 44 3 80 138 52 22 Hot water extraction

0--10 357 77 3 360 432 279 22 Hot water extraction

10--20 26 156 3 65 304 11 22 Hot water extraction

10--40 260 110 5 220 425 141 22 Hot water extraction

20--40 6 8 4 3 17 1 22 Hot water extraction

Calcium (mg/kg) 0--10 2 2 3 3 3 22 Hot water extraction

0--10 91 85 3 88 178 8 22 Ammonium acetate extraction

0--10 8 3 3 8 11 6 22 Hot water extraction

0--10 189 76 3 193 262 111 22 Ammonium acetate extraction

10--20 3 2 2 3 4 1 22 Hot water extraction

10--20 30 21 3 38 45 5 22 Ammonium acetate extraction

10--40 5 4 5 4 13 2 22 Hot water extraction

10--40 84 37 5 67 149 60 22 Ammonium acetate extraction

20--40 1 0 4 1 1 1 22 Hot water extraction
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

407

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

20--40 11 10 5 10 28 1 22 Ammonium acetate extraction

Iron (mg/kg) 0--10 83 53 3 81 136 31 22 Hot water extraction

0--10 112 61 3 101 178 57 22 Hot water extraction

10--20 133 165 3 74 319 5 22 Hot water extraction

10--40 50 18 5 52 66 20 22 Hot water extraction

20--40 7 8 4 3 19 1 22 Hot water extraction

Magnesium (mg/kg) 0--10 3 1 3 3 3 2 22 Hot water extraction

0--10 68 50 3 44 125 36 22 Ammonium acetate extraction

0--10 9 4 3 8 13 5 22 Hot water extraction

0--10 108 60 3 74 177 73 22 Ammonium acetate extraction

10--20 2 2 3 1 5 1 22 Hot water extraction

10--20 30 8 3 33 37 21 22 Ammonium acetate extraction

10--40 4 2 5 4 6 1 22 Hot water extraction

10--40 64 44 5 46 142 38 22 Ammonium acetate extraction

20--40 0.40 0.25 6 0.38 0.86 0.16 22 Hot water extraction

20--40 11 6 6 9 22 6 22 Ammonium acetate extraction

Phosphorus (mg/kg) 0--10 0.19 0.16 3 0.11 0.38 0.08 22 Hot water extraction

0--10 3.87 1.44 3 3.96 5.25 2.38 22 Hot water extraction

10--20 0.36 0.38 3 0.27 0.79 0.04 22 Hot water extraction

10--40 2.90 1.30 5 3.47 4.08 0.92 22 Hot water extraction

20--40 0.09 0.07 2 0.09 0.14 0.04 22 Hot water extraction

Potassium (mg/kg) 0--10 6 3 3 6 9 3 22 Hot water extraction

0--10 17 6 3 19 22 10 22 Hot water extraction

10--20 6 6 3 4 13 1 22 Hot water extraction

10--40 12 6 5 16 17 5 22 Hot water extraction

20--40 0.73 0.45 6 0.70 1.35 0.22 22 Hot water extraction
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

408

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--10 50 15 3 46 67 38 22 Ammonium acetate extraction

0--10 65 18 3 70 79 45 22 Ammonium acetate extraction

10--20 25 3 3 27 27 21 22 Ammonium acetate extraction

10--40 42 10 5 45 54 30 22 Ammonium acetate extraction

20--40 15 8 6 11 29 8 22 Ammonium acetate extraction

Sodium (mg/kg) 0--10 21 3 3 20 24 19 22 Ammonium acetate extraction

0--10 49 19 3 49 69 30 22 Ammonium acetate extraction

10--20 14 1 3 14 16 14 22 Ammonium acetate extraction

10--40 35 10 5 34 44 19 22 Ammonium acetate extraction

20--40 8 2 6 8 10 4 22 Ammonium acetate extraction

Swamp Mangle Mfs Aluminum (cmol/kg-l) 0--20 0.41 0.28 18 0.32 0.94 0.15 23

20--30 0.25 1 23

Calcium (cmol/kg-l) 0--20 51.52 11.38 18 57.11 67.22 29.64 23

20--30 61.34 1 23

Calcium (mg/g) 0--20 10.30 2.28 18 11.42 13.44 5.93 23

20--30 12.27 1 23

Carbon (%) 0--20 28.32 4.94 16 30.64 33.39 18.89 23

Iron (mg/g) 0--20 0.59 0.65 18 0.08 1.61 0.03 23

20--30 0.01 1 23

Loss on ignition (%) 0--20 62.60 5.87 18 63.68 72.08 50.56 23

20--30 78.31 1 23

Magnesium (mg/g) 0--20 4.68 2.46 18 2.81 8.30 2.31 23

20--30 2.02 1 23

Magnesium (cmol/kg-l) 0--20 38.97 20.50 18 23.43 69.16 19.24 23

20--30 16.81 1 23

Manganese (mg/g) 0--20 0.07 0.04 18 0.07 0.16 0.02 23
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

409

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

20--30 0.07 1 23

Nitrogen (%) 0--20 1.56 0.35 16 1.64 1.95 1.10 23

Organic matter (%) 0--20 28.35 5.72 18 30.61 37.73 17.15 23

20--30 34.86 0.69 2 34.86 35.35 34.37 23

pH (H2O) 0--20 6.30 1.12 9 7.06 7.43 4.93 23

20--30 7.14 0.00 1 23

pH (KCl) 0--20 6.15 1.04 9 6.83 7.12 4.87 23

20--30 6.76 0.00 1 6.76 6.76 6.76 23

Phosphorus (mg/kg) 0--20 45 16 18 44 63 26 23

20--30 23 1 23

Potassium (cmol/kg-l) 0--20 2.27 1.82 18 1.53 5.89 0.32 23

20--30 0.29 1 23

Potassium (mg/g) 0--20 0.89 0.71 18 0.60 2.30 0.12 23

20--30 0.11 1 23

Sodium (cmol/kg-l) 0--20 75.83 62.29 18 36.78 169.50 9.41 23

20--30 8.06 1 23

Sodium (mg/g) 0--20 17.44 14.33 18 8.46 38.99 2.16 23

20--30 1.85 1 23

Swamps Entisols Mfs Aluminum (mg/g) 0--30 0.11 0.04 26 0.10 0.20 0.03 24

100-150 0.09 0.02 26 0.09 0.13 0.05 24

Aluminum-total (mg/g) 0--30 16.71 2.80 26 16.36 23.07 12.44 24

100-150 13.55 2.09 26 13.67 18.25 9.17 24

Calcium (mg/g) 0--30 5.97 1.29 6 6.30 7.53 2.01 24

100-150 6.44 0.82 26 6.38 7.71 4.70 24

Calcium-total (mg/g) 0--30 5.70 0.63 26 5.77 7.01 4.06 24

100-150 5.48 0.34 26 5.50 5.96 4.82 24
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

410

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Carbon (%) 0--30 25.31 6.61 26 27.05 32.16 7.15 24

100-150 26.76 4.62 26 28.62 32.65 14.06 24

Iron-total (mg/g) 0--30 17.77 2.29 26 17.58 21.15 12.01 24

100-150 17.08 2.81 26 17.60 22.06 11.30 24

Magnesium (mg/g) 0--30 10.52 2.33 26 10.97 13.19 3.24 24

100-150 12.83 1.64 26 13.01 15.64 10.06 24

Magnesium-total (mg/g) 0--30 11.80 1.51 26 1.63 14.01 8.09 24

100-150 12.95 0.84 26 12.79 14.30 11.24 24

Manganese (mg/kg) 0--30 130 5 26 120 250 7 24

100-150 160 5 26 160 280 10 24

Manganese-total (mg/g) 0--30 0.08 0.03 26 0.07 0.17 0.05 24

100-150 0.07 0.01 26 0.07 0.10 0.05 24

Nitrogen (%) 0--30 0.95 0.27 26 1.05 1.19 0.28 24

100-150 0.83 0.15 26 0.87 0.97 0.37 24

Phosphorus (mg/g) 0--30 0.05 0.01 26 0.05 0.07 0.02 24

100-150 0.05 0.01 26 0.05 0.06 0.03 24

Phosphorus-total (mg/g) 0--30 0.20 0.05 26 0.20 0.29 0.11 24

100-150 0.13 0.03 26 0.12 0.18 0.08 24

Potassium (mg/g) 0--30 3.62 0.92 26 3.74 4.85 0.97 24

100-150 4.21 0.85 26 4.02 5.45 2.85 24

Potassium-total (mg/g) 0--30 3.64 0.32 26 3.68 4.06 2.82 24

100-150 3.75 0.27 26 3.80 4.25 3.02 24

Sulfur (%) 0--30 6.77 2.19 26 7.38 9.69 1.60 24

 100-150 7.65 1.55 26 8.36 9.46 3.96 24

Swamp Histosol Mfs Aluminum (mg/kg) 0--20 9 11 10 3 32 6 25 Low salinity

0--20 130 110 10 120 320 20 25 Intermediates alinity
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

411

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--20 75 126 10 30 3240 11 25 High salinity

Calcium (mg/g) 0--20 1.00 0.28 10 1.05 1.28 0.52 25 Low salinity

0--20 0.60 0.16 10 0.62 0.85 0.39 25 Intermediates alinity

0--20 0.93 0.70 10 0.69 2.22 0.34 25 High salinity

Carbon (%) 0--20 20.39 6.22 10 21.97 28.06 10.12 25 Low salinity

0--20 17.82 5.53 10 15.78 27.10 11.19 25 Intermediates alinity

0--20 18.94 1.23 10 18.94 20.31 16.94 25 High salinity

Iron (mg/g) 0--20 0.05 0.06 10 0.02 0.19 0.01 25 Low salinity

0--20 0.05 0.04 10 0.04 0.11 0.01 25 Intermediates alinity

0--20 0.02 0.02 10 0.01 0.09 0.01 25 High salinity

Loss on ignition (%) 0--20 44.32 9.80 10 47.56 55.91 28.39 25 Low salinity

0--20 39.78 9.31 10 35.84 55.35 28.99 25 Intermediates alinity

0--20 40.14 2.03 10 40.27 42.86 37.80 25 High salinity

Magnesium (mg/g) 0--20 0.74 0.21 10 0.75 1.02 0.42 25 Low salinity

0--20 0.67 0.03 10 0.69 0.77 0.54 25 Intermediates alinity

0--20 0.89 0.53 10 0.81 1.82 0.33 25 High salinity

Manganese (mg/kg) 0--20 14 14 10 8 39 2 25 Low salinity

0--20 10 7 10 8 22 4 25 Intermediates alinity

0--20 20 21 10 14 58 1 25 High salinity

Nitrogen (%) 0--20 0.63 0.15 10 0.70 0.79 0.37 25 Low salinity

0--20 0.57 0.12 10 0.65 0.69 0.40 25 Intermediates alinity

0--20 1.36 0.87 10 1.08 3.00 0.58 25 High salinity

pH (H2O) 0--20 5.95 0.21 10 6.05 6.14 5.69 25 Low salinity

0--20 5.66 0.36 10 5.48 6.10 5.25 25 Intermediates alinity

0--20 5.21 0.47 10 5.34 5.75 4.38 25 High salinity

pH (KCl) 0--20 5.55 0.15 10 5.60 5.78 5.37 25 Low salinity
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

412

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--20 5.29 0.33 10 5.11 5.77 4.94 25 Intermediates alinity

0--20 4.82 0.43 10 4.89 5.32 4.08 25 High salinity

Phosphorus (mg/kg) 0--20 4 3 10 3 11 2 25 Low salinity

0--20 3 2 10 3 6 2 25 Intermediates alinity

0--20 2 10 10 2 3 2 25 High salinity

Potassium (mg/g) 0--20 0.17 0.08 10 0.15 0.30 0.10 25 Low salinity

0--20 0.20 0.03 10 0.20 0.26 0.15 25 Intermediates alinity

0--20 0.22 0.08 10 0.21 0.34 0.11 25 High salinity

Sulfur (%) 0--20 1.38 0.44 10 1.51 1.90 0.68 25 Low salinity

0--20 1.09 0.39 10 1.02 1.71 0.57 25 Intermediates alinity

0--20 1.03 0.19 10 1.06 1.29 0.74 25 High salinity

Swamps Entisols Mfs Aluminum (mg/g) 0--20 0.16 0.11 12 0.13 0.34 0.03 26 Hot water extraction

Calcium (mg/g) 0--20 3.48 1.07 12 3.68 5.02 1.72 26 Hot water extraction

0--20 4.01 0.70 12 4.28 4.73 2.61 26 Ammonium Acetate extraction

Carbon (%) 0--20 21.52 6.09 12 23.46 28.70 9.43 26

Iron (mg/g) 0--20 0.07 0.06 12 0.04 0.20 0.01 26 Hot water extraction

Loss on ignition (%) 0--20 61.38 14.40 12 66.03 74.90 33.39 26

Magnesium (mg/g) 0--20 8.54 2.30 12 8.86 11.41 4.14 26 Hot water extraction

0--20 9.72 2.31 12 10.54 11.90 5.06 26 Ammonium acetate extraction

Manganese (mg/g) 0--20 0.02 0.01 12 0.02 0.06 0.01 26 Hot water extraction

0--20 0.02 0.01 12 0.01 0.04 0.01 26 Ammonium acetate extraction

Nitrogen (%) 0--20 0.92 0.27 12 0.97 1.27 0.36 26

pH (H2O) 0--20 4.16 0.31 12 4.18 4.59 3.76 26

pH (KCl) 0--20 4.06 0.32 12 4.07 4.50 3.66 26

Phosphorus (mg/kg) 0--20 8 5 12 7 22 2 26 Hot water extraction

0--20 257 140 12 213 572 82 26 Ammonium acetate extraction
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

413

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Potassium (mg/g) 0--20 1.65 0.74 12 1.77 2.52 0.11 26 Hot water extraction

0--20 0.05 0.01 12 0.05 0.06 0.03 26 Ammonium acetate extraction

Tuffaceous sandstone Ultisols Lmrf Aluminum (mg/g) 0--10 0.15 0.09 36 0.14 0.38 0.02 27 Gap 1993

0--10 0.20 0.14 17 0.17 0.46 0.02 27 Control 1993

0--10 0.34 0.12 36 0.34 0.59 0.13 27 Gap 1994

0--10 0.30 0.16 17 0.31 0.57 0.08 27 Control 1994

0--10 0.32 0.13 34 0.34 0.70 0.10 27 Gap 1995

0--10 0.28 0.15 17 0.29 0.57 0.02 27 Control 1995

0--10 0.32 0.13 36 0.31 0.68 0.06 27 Gap 1996

0--10 0.29 0.16 17 0.31 0.55 0.06 27 Control 1996

0--10 0.32 0.11 44 0.32 0.53 0.14 27 Gap 1997

0--10 0.15 0.08 9 0.12 0.25 0.03 27 Control 1997

0--10 0.29 0.11 30 0.31 0.52 0.13 27 Gap 1998

0--10 0.27 0.14 14 0.27 0.56 0.07 27 Control 1998

0--10 0.37 0.11 30 0.39 0.55 0.11 27 Gap 1999

0--10 0.27 0.14 14 0.30 0.51 0.04 27 Control 1999

10--35 0.18 0.11 30 0.16 0.51 0.001 27 Gap 1993

10--35 0.18 0.12 14 0.18 0.35 0.02 27 Control 1993

10--35 0.37 0.14 30 0.36 0.68 0.11 27 Gap 1994

10--35 0.27 0.14 14 0.30 0.48 0.05 27 Control 1994

10--35 0.37 0.16 32 0.35 0.66 0.04 27 Gap 1995

10--35 0.28 0.17 14 0.26 0.73 0.05 27 Control 1995

10--35 0.35 0.14 30 0.34 0.72 0.13 27 Gap 1996

10--35 0.32 0.18 13 0.31 0.81 0.02 27 Control 1996

10--35 0.32 0.11 37 0.33 0.58 0.16 27 Gap 1997

10--35 0.15 0.09 7 0.17 0.27 0.05 27 Control 1997
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

414

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--35 0.34 0.13 36 0.33 0.64 0.10 27 Gap 1998

10--35 0.24 0.14 16 0.25 0.45 0.05 27 Control 1998

10--35 0.35 0.15 36 0.34 0.62 0.06 27 Gap 1999

10--35 0.24 0.11 16 0.26 0.38 0.06 27 Control 1999

Calcium (mg/g) 0--10 0.13 0.10 36 0.10 0.40 0.02 27 Gap 1993

0--10 0.14 0.09 17 0.11 0.33 0.03 27 Control 1993

0--10 0.28 0.16 36 0.24 0.70 0.04 27 Gap 1994

0--10 0.26 0.17 17 0.17 0.64 0.12 27 Control 1994

0--10 0.30 0.20 34 0.26 0.73 0.03 27 Gap 1995

0--10 0.34 0.19 17 0.31 0.66 0.06 27 Control 1995

0--10 0.28 0.19 36 0.27 0.83 0.02 27 Gap 1996

0--10 0.33 0.17 17 0.31 0.62 0.06 27 Control 1996

0--10 0.17 0.11 44 0.16 0.47 0.01 27 Gap 1997

0--10 0.30 0.19 9 0.28 0.70 0.06 27 Control 1997

0--10 0.29 0.16 30 0.26 0.78 0.03 27 Gap 1998

0--10 0.35 0.15 14 0.31 0.64 0.13 27 Control 1998

0--10 0.34 0.23 30 0.26 1.21 0.09 27 Gap 1999

0--10 0.36 0.28 14 0.23 1.08 0.09 27 Control 1999

10--35 0.04 0.02 30 0.04 0.11 0.01 27 Gap 1993

10--35 0.05 0.04 14 0.03 0.13 0.01 27 Control 1993

10--35 0.10 0.06 30 0.10 0.25 0.01 27 Gap 1994

10--35 0.10 0.09 14 0.06 0.31 0.03 27 Control 1994

10--35 0.12 0.12 32 0.07 0.44 0.01 27 Gap 1995

10--35 0.13 0.11 14 0.09 0.39 0.01 27 Control 1995

10--35 0.12 0.09 30 0.09 0.34 0.01 27 Gap 1996

10--35 0.12 0.10 14 0.08 0.32 0.01 27 Control 1996
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

415

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--35 0.06 0.06 37 0.04 0.25 0.01 27 Gap 1997

10--35 0.16 0.20 7 0.13 0.34 0.02 27 Control 1997

10--35 0.11 0.08 36 0.10 0.32 0.01 27 Gap 1998

10--35 0.15 0.13 16 0.10 0.50 0.03 27 Control 1998

10--35 0.17 0.14 36 0.11 0.59 0.02 27 Gap 1999

10--35 0.16 0.17 16 0.08 0.63 0.04 27 Control 1999

Magnesium (mg/g) 0--10 0.07 0.03 36 0.07 0.15 0.03 27 Gap 1993

0--10 0.06 0.03 17 0.05 0.11 0.03 27 Control 1993

0--10 0.16 0.08 36 0.13 0.41 0.06 27 Gap 1994

0--10 0.12 0.06 17 0.10 0.23 0.05 27 Control 1994

0--10 0.16 0.07 34 0.16 0.33 0.05 27 Gap 1995

0--10 0.15 0.08 17 0.15 0.30 0.03 27 Control 1995

0--10 0.16 0.06 36 0.16 0.28 0.05 27 Gap 1996

0--10 0.14 0.08 17 0.12 0.30 0.06 27 Control 1996

0--10 0.11 0.06 44 0.10 0.32 0.03 27 Gap 1997

0--10 0.14 0.05 9 0.16 0.22 0.07 27 Control 1997

0--10 0.17 0.07 30 0.16 0.32 0.05 27 Gap 1998

0--10 0.15 0.07 14 0.13 0.27 0.06 27 Control 1998

0--10 0.20 0.11 30 0.17 0.59 0.06 27 Gap 1999

0--10 0.14 0.08 14 0.11 0.30 0.05 27 Control 1999

10--35 0.05 0.02 30 0.05 0.11 0.02 27 Gap 1993

10--35 0.04 0.03 14 0.03 0.12 0.01 27 Control 1993

10--35 0.10 0.05 30 0.08 0.22 0.03 27 Gap 1994

10--35 0.08 0.06 14 0.05 0.18 0.01 27 Control 1994

10--35 0.10 0.06 32 0.08 0.22 0.02 27 Gap 1995

10--35 0.09 0.07 14 0.08 0.22 0.01 27 Control 1995
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

416

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--35 0.10 0.06 30 0.79 0.23 0.02 27 Gap 1996

10--35 0.08 0.07 14 0.06 0.25 0.01 27 Control 1996

10--35 0.06 0.04 37 0.05 0.17 0.01 27 Gap 1997

10--35 0.12 0.08 7 0.12 0.24 0.03 27 Control 1997

10--35 0.10 0.06 36 0.07 0.21 0.02 27 Gap 1998

10--35 0.09 0.07 16 0.06 0.20 0.01 27 Control 1998

10--35 0.12 0.07 36 94.00 0.27 0.03 27 Gap 1999

10--35 0.09 0.08 16 0.06 0.27 0.02 27 Control 1999

Manganese (mg/kg) 0--10 16 17 36 9 81 3 27 Gap 1993

0--10 10 16 17 5 73 3 27 Control 1993

0--10 17 9 36 14 52 5 27 Gap 1994

0--10 14 9 17 11 33 3 27 Control 1994

0--10 22 24 34 18 119 1 27 Gap 1995

0--10 13 8 17 14 25 3 27 Control 1995

0--10 27 21 36 20 95 3 27 Gap 1996

0--10 15 7 17 16 31 4 27 Control 1996

0--10 11 8 44 10 35 1 27 Gap 1997

0--10 16 2 9 16 20 14 27 Control 1997

0--10 31 34 30 20 147 3 27 Gap 1998

0--10 16 6 14 16 28 8 27 Control 1998

0--10 19 14 30 17 69 3 27 Gap 1999

0--10 11 7 14 10 23 2 27 Control 1999

10--35 21 40 30 8 188 1 27 Gap 1993

10--35 7 14 14 3 55 1 27 Control 1993

10--35 14 15 30 12 78 1 27 Gap 1994

10--35 12 12 14 10 39 1 27 Control 1994
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

417

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--35 20 33 32 10 139 1 27 Gap 1995

10--35 8 8 14 6 22 1 27 Control 1995

10--35 26 33 27 13 121 1 27 Gap 1996

10--35 12 13 12 7 46 1 27 Control 1996

10--35 9 13 37 4 71 1 27 Gap 1997

10--35 15 8 7 13 26 6 27 Control 1997

10--35 29 50 36 12 185 1 27 Gap 1998

10--35 11 9 16 7 27 1 27 Control 1998

10--35 18 15 33 13 80 1 27 Gap 1999

10--35 7 9 16 4 33 1 27 Control 1999

pH (H2O) 0--10 4.48 0.20 36 4.52 4.92 4.04 27 Gap 1994

0--10 4.47 0.18 17 4.42 4.81 4.21 27 Control 1994

0--10 4.44 0.27 34 4.37 5.18 4.05 27 Gap 1995

0--10 4.46 0.24 17 4.44 4.91 4.11 27 Control 1995

0--10 4.57 0.27 36 4.53 5.21 4.13 27 Gap 1996

0--10 4.69 0.40 17 4.74 5.43 4.17 27 Control 1996

0--10 4.54 0.35 44 4.42 5.23 3.86 27 Gap 1997

0--10 5.14 0.17 9 5.14 5.36 4.77 27 Control 1997

0--10 5.92 7.30 30 4.56 4.45 4.05 27 Gap 1998

0--10 4.64 0.32 14 4.71 5.06 4.11 27 Control 1998

0--10 4.88 0.29 30 4.88 5.30 3.97 27 Gap 1999

0--10 4.86 0.25 14 4.85 5.24 4.58 27 Control 1999

10--35 4.55 0.14 30 4.50 4.75 4.20 27 Gap 1994

10--35 4.49 0.20 14 4.45 4.79 4.17 27 Control 1994

10--35 4.44 0.25 32 4.39 5.00 3.85 27 Gap 1995

10--35 4.92 0.19 14 4.52 4.79 4.22 27 Control 1995
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

418

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--35 4.51 0.27 30 4.47 5.05 4.50 27 Gap 1996

10--35 4.69 0.26 14 4.64 5.29 4.28 27 Control 1996

10--35 4.58 0.32 37 4.51 5.38 4.14 27 Gap 1997

10--35 5.07 0.26 7 5.05 5.46 4.66 27 Control 1997

10--35 4.55 0.24 36 4.52 5.00 4.13 27 Gap 1998

10--35 4.66 0.30 16 4.64 5.12 4.21 27 Control 1998

10--35 4.99 0.26 36 4.97 5.53 4.01 27 Gap 1999

10--35 4.97 0.32 16 5.04 5.41 4.18 27 Control 1999

pH (KCl) 0--10 3.91 0.15 36 3.89 4.44 3.68 27 Gap 1996

0--10 3.91 0.27 17 4.01 4.32 3.42 27 Control 1996

0--10 3.92 0.13 44 3.94 4.15 3.58 27 Gap 1997

0--10 4.13 0.11 9 4.19 4.29 3.97 27 Control 1997

0--10 3.97 0.12 30 3.99 4.18 3.70 27 Gap 1998

0--10 4.02 0.23 14 4.07 4.38 3.58 27 Control 1998

0--10 3.92 0.11 30 3.91 4.20 3.64 27 Gap 1999

0--10 3.93 0.23 14 3.99 4.24 3.49 27 Control 1999

10--35 3.93 0.11 30 3.93 4.11 3.70 27 Gap 1996

10--35 3.97 0.23 14 3.93 4.57 3.65 27 Control 1996

10--35 3.95 0.08 37 3.94 4.11 3.76 27 Gap 1997

10--35 4.18 0.15 7 4.12 4.42 4.03 27 Control 1997

10--35 4.02 0.09 36 4.01 4.17 3.85 27 Gap 1998

10--35 4.08 0.24 16 4.03 4.54 3.78 27 Control 1998

10--35 3.95 0.09 36 3.93 4.12 3.75 27 Gap 1999

10--35 4.02 0.33 16 3.96 4.91 3.40 27 Control 1999

Potassium (mg/kg) 0--10 25 16 36 21 65 2 27 Gap 1993

0--10 36 19 16 36 74 8 27 Control 1993
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

419

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--10 54 22 36 50 154 24 27 Gap 1994

0--10 81 46 17 67 202 22 27 Control 1994

0--10 62 38 34 50 158 14 27 Gap 1995

0--10 77 54 17 62 237 27 27 Control 1995

0--10 50 29 36 45 135 13 27 Gap 1996

0--10 60 23 17 59 107 27 27 Control 1996

0--10 31 14 43 29 65 3 27 Gap 1997

0--10 78 35 9 68 152 39 27 Control 1997

0--10 53 17 30 54 85 18 27 Gap 1998

0--10 75 38 14 58 174 43 27 Control 1998

0--10 326 112 30 325 517 91 27 Gap 1999

0--10 188 82 14 165 328 97 27 Control 1999

10--35 130 11 30 9 48 1 27 Gap 1993

10--35 19 5 9 18 27 11 27 Control 1993

10--35 28 16 30 24 72 9 27 Gap 1994

10--35 33 15 14 33 72 13 27 Control 1994

10--35 30 20 32 24 91 7 27 Gap 1995

10--35 26 15 14 19 58 15 27 Control 1995

10--35 19 15 26 17 72 1 27 Gap 1996

10--35 27 20 14 20 77 6 27 Control 1996

10--35 16 8 35 14 44 6 27 Gap 1997

10--35 37 17 7 29 58 15 27 Control 1997

10--35 26 11 35 26 56 9 27 Gap 1998

10--35 31 19 16 22 73 13 27 Control 1998

10--35 245 91 36 262 508 71 27 Gap 1999

10--35 128 61 16 108 281 75 27 Control 1999
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

420

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Tuffaceous sandstone Ultisols Lmrf Iron (mg/g) 0--10 0.80 0.47 133 0.76 2.22 0.12 28

0--10 1.24 1.07 63 0.73 4.02 0.13 28 Control

Loss on ignition (%) 0--10 18.14 2.72 133 17.94 31.58 12.82 28

0--10 18.76 3.06 63 19.48 24.62 9.04 28 Control

Manganese (mg/kg) 0--10 72 80 133 43 445 3 28

0--10 98 124 63 48 702 2 28 Control

Organic matter (%) 0--10 5.62 1.84 133 5.69 10.09 2.15 28

0--10 6.36 2.43 63 6.24 13.12 1.25 28 Control

Phosphorus (mg/kg) 0--10 9 5 133 9 33 1 28

0--10 13 11 63 10 46 1 28 Control

Potassium (mg/g) 0--10 0.07 0.03 133 0.06 0.19 0.02 28

0--10 0.09 0.06 63 0.08 0.33 0.01 28 Control

Tuffaceous sandstone Ultisols Swf Aluminum (cmol/kg-l) 0--10 1.55 3.35 80 0.25 13.99 0.10 29

10--35 2.55 4.22 85 0.63 15.63 0.04 29

C/N 0--10 15 1 80 15 18 12 29

10--35 18 4 85 17 31 14 29

Calcium (cmol/kg-l) 0--10 22.19 15.72 80 24.12 51.28 0.43 29

10--35 21.11 16.44 85 20.33 52.86 0.13 29

Calcium (mg/g) 0--10 4.44 3.14 80 4.82 10.26 0.09 29

10--35 4.22 3.29 85 4.07 10.57 0.03 29

Carbon (%) 0--10 3.84 0.75 80 3.73 5.64 2.05 29

10--35 1.81 0.43 85 1.84 2.84 0.79 29

ECEC (cmol/kg-1) 0--10 35.05 18.40 80 39.78 69.74 8.31 29

10--35 36.41 18.42 85 41.03 67.31 6.41 29

Iron (mg/g) 0--10 0.25 0.34 80 0.08 1.39 0.01 29

10--35 0.24 0.34 85 0.07 1.30 0.02 29
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

421

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Loss on ignition (%) 0--10 15.50 2.96 80 16.19 19.35 9.88 29

10--35 12.61 2.94 85 13.89 16.07 5.76 29

Magnesium (cmol/kg-1) 0--10 10.90 5.88 80 12.78 19.96 0.50 29

10--35 12.29 7.33 85 13.72 27.37 0.33 29

Magnesium (mg/g) 0--10 1.31 0.71 80 1.53 2.39 0.06 29

10--35 1.48 0.88 85 1.65 3.28 0.04 29

Manganese (mg/g) 0--10 0.12 0.05 80 0.12 0.30 0.05 29

10--35 0.84 0.04 85 0.09 0.16 0.01 29

Nitrogen (%) 0--10 0.26 0.05 80 0.26 0.36 0.13 29

10--35 0.11 0.04 85 0.11 0.20 0.03 29

pH (H2O) 0--10 5.70 0.64 80 5.81 6.72 4.03 29

10--35 5.49 0.60 85 5.53 6.49 4.21 29

pH (KCl) 0--10 4.55 0.49 80 4.63 5.29 3.38 29

10--35 4.09 0.32 85 4.05 4.76 3.51 29

Phosphorus (mg/kg) 0--10 11 5 80 10 25 5 29

10--35 6 3 85 5 15 2 29

Potassium (mg/g) 0--10 0.14 0.08 80 0.12 0.32 0.05 29

10--35 0.08 0.07 85 0.04 0.26 0.01 29

Sodium (cmol/kg-l) 0--10 0.41 0.17 80 0.42 0.94 0.11 29

10--35 0.46 0.26 85 0.47 1.17 0.08 29

Sodium (mg/g) 0--10 0.09 0.04 80 0.10 0.22 0.02 29

10--35 0.11 0.06 85 0.11 0.27 0.02 29

Compound dunes Ultisols Mfs Potassium (cmol/kg-l) 0--10 0.37 0.07 4 0.40 0.42 0.26 30 Ensenada Brei

10--40 0.45 0.50 11 0.25 2.04 0.12 30 Ensenada Brei

40--50 0.46 0.15 4 0.44 0.65 0.30 30 Ensenada Brei

Alluvial deposits Alfisol Mfs 0--10 0.31 0.09 5 0.36 0.38 0.20 30 Cerro Gordo
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

422

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--50 0.13 0.04 5 0.13 0.17 0.06 30 Cerro Gordo

70--80 0.19 0.01 2 0.19 0.19 0.18 30 Cerro Gordo

Compound dunes Ultisols Mfs Calcium (cmol/kg-l) 0--10 2.49 0.90 4 2.93 2.98 1.14 30 Ensenada Brei

10--40 1.73 1.27 11 1.36 4.29 0.52 30 Ensenada Brei

40--50 2.20 1.37 4 1.97 4.05 0.81 30 Ensenada Brei

Alluvial deposits Alfisol Mfs 0--10 2.57 1.34 5 1.82 4.07 1.24 30 Cerro Gordo

10--50 1.48 0.91 5 1.11 2.85 0.67 30 Cerro Gordo

70--80 4.64 0.49 2 4.64 4.99 4.29 30 Cerro Gordo

Compound dunes Ultisols Mfs Calcium (mg/g) 0--10 0.50 0.18 4 0.59 0.60 0.23 30 Ensenada Brei

10--40 0.35 0.25 11 0.27 0.86 0.10 30 Ensenada Brei

40--50 0.44 0.27 4 0.39 0.81 0.16 30 Ensenada Brei

Alluvial deposits Alfisol Mfs 0--10 0.51 0.27 5 0.36 0.81 0.25 30 Cerro Gordo

10--50 0.30 0.18 5 0.22 0.57 0.13 30 Cerro Gordo

70--80 0.93 0.10 2 0.93 1.00 0.86 30 Cerro Gordo

Compound dunes Ultisols Mfs Iron (mg/g) 0--10 0.06 0.00 4 0.06 0.06 0.05 30 Ensenada Brei

10--40 0.06 0.07 11 0.03 0.28 0.02 30 Ensenada Brei

40--50 0.03 0.01 4 0.03 0.04 0.02 30 Ensenada Brei

Alluvial deposits Alfisol Mfs 0--10 0.07 0.02 5 0.06 0.09 0.05 30 Cerro Gordo

10--50 0.04 0.02 5 0.03 0.06 0.02 30 Cerro Gordo

70--80 0.03 0.00 2 0.03 0.03 0.03 30 Cerro Gordo

Compound dunes Ultisols Mfs Magnesium (cmol/kg-l) 0--10 1.57 0.55 4 1.40 2.36 1.11 30 Ensenada Brei

10--40 1.32 1.07 11 0.78 4.05 0.53 30 Ensenada Brei

40--50 2.68 1.80 4 2.40 5.13 0.81 30 Ensenada Brei

Alluvial deposits Alfisol Mfs 0--10 2.75 1.33 5 2.08 4.63 1.83 30 Cerro Gordo

10--50 1.70 1.98 5 0.74 5.19 0.58 30 Cerro Gordo

70--80 4.28 0.11 2 4.28 4.35 4.20 30 Cerro Gordo
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

423

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Compound dunes Ultisols Mfs Magnesium (mg/g) 0--10 0.19 0.07 4 0.17 0.28 0.13 30 Ensenada Brei

10--40 0.16 0.13 11 0.09 0.49 0.06 30 Ensenada Brei

40--50 0.32 0.22 4 0.29 0.62 0.10 30 Ensenada Brei

Alluvial deposits Alfisol Mfs 0--10 0.33 0.16 5 0.25 0.56 0.17 30 Cerro Gordo

10--50 0.20 0.24 5 0.09 0.62 0.07 30 Cerro Gordo

70--80 0.51 0.01 2 0.51 0.52 0.50 30 Cerro Gordo

Compound dunes Ultisols Mfs Manganese (mg/kg) 0--10 62 16 4 69 73 38 30 Ensenada Brei

10--40 23 6 11 22 34 16 30 Ensenada Brei

40--50 20 6 4 21 26 13 30 Ensenada Brei

Alluvial deposits Alfisol Mfs 0--10 52 12 5 47 68 40 30 Cerro Gordo

10--50 21 9 5 20 30 11 30 Cerro Gordo

70--80 25 0 2 25 25 25 30 Cerro Gordo

Compound dunes Ultisols Mfs pH (H2O) 0--10 5.90 0.40 4 5.80 6.40 5.60 30 Ensenada Brei

10--40 6.00 1.00 11 5.90 6.80 5.20 30 Ensenada Brei

40--50 6.00 0.50 4 5.80 6.70 5.50 30 Ensenada Brei

Alluvial deposits Alfisol Mfs 0--10 6.20 0.60 5 6.00 7.20 5.60 30 Cerro Gordo

10--50 5.80 0.20 5 5.70 6.10 5.50 30 Cerro Gordo

70--80 6.30 0.00 2 6.30 6.30 6.30 30 Cerro Gordo

Compound dunes Ultisols Mfs Phosphorus (mg/kg) 0--10 12 2 4 13 14 9 30 Ensenada Brei

10--40 9 12 11 6 5 4 30 Ensenada Brei

40--50 3 1 4 3 3 2 30 Ensenada Brei

Alluvial deposits Alfisol Mfs 0--10 7 1 5 8 8 6 30 Cerro Gordo

10--50 3 1 5 3 5 1 30 Cerro Gordo

70--80 2 0 2 2 2 2 30 Cerro Gordo

Compound dunes Ultisols Mfs Potassium (mg/g) 0--10 0.14 0.03 4 0.15 0.16 0.10 30 Ensenada Brei

10--40 0.17 0.21 11 0.10 0.78 0.05 30 Ensenada Brei
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

424

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

40--50 0.18 0.06 4 0.17 0.25 0.12 30 Ensenada Brei

Alluvial deposits Alfisol Mfs 0--10 0.12 0.03 5 0.14 0.17 0.08 30 Cerro Gordo

10--50 0.05 0.02 5 0.05 0.07 0.02 30 Cerro Gordo

70--80 0.07 0.00 2 0.07 0.07 0.07 30 Cerro Gordo

Tuffaceous sandstone Ultisols Lmrf Aluminum (cmol/kg-l) 0--20 0.29 1 31

20--100 9.21 4.62 2 9.21 12.48 5.94 31

100-150 11.08 2.07 3 12.03 12.50 8.71 31

200-400 11.14 2.90 2 11.14 13.19 9.09 31

400-500 15.08 0.50 3 15.31 15.42 14.50 31

500-680 4.77 4.99 4 2.96 11.94 1.23 31

Calcium (cmol/kg-l) 0--20 7.05 1 31

20--100 1.28 0.58 2 1.28 1.70 0.87 31

100-150 0.59 0.15 3 0.64 0.72 0.42 31

200-400 0.49 0.17 2 0.49 0.60 0.37 31

400-500 0.95 0.17 3 1.01 1.08 0.76 31

500-680 7.40 7.15 4 7.21 14.23 0.97 31

Calcium (mg/g) 0--20 1.41 1 31

20--100 0.26 0.12 2 0.26 0.34 0.17 31

100-150 0.12 0.03 3 0.13 0.14 0.08 31

200-400 0.10 0.03 2 0.10 0.12 0.07 31

400-500 0.19 0.03 3 0.20 0.22 0.15 31

500-680 1.48 1.43 4 1.44 2.85 0.19 31

Carbon (%) 0--20 3.58 1 31

20--100 0.32 0.05 2 0.32 0.35 0.28 31

100-150 0.38 0.17 3 0.34 0.57 0.23 31

200-400 0.18 0.05 2 0.18 0.22 0.15 31
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

425

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

400-500 0.08 0.02 3 0.07 0.10 0.07 31

500-680 0.07 0.03 4 0.08 0.09 0.04 31

ECEC (cmol/kg-1) 0--20 10.72 1 31

20--100 11.43 3.71 2 11.43 14.05 8.81 31

100-150 12.37 2.42 3 13.26 14.21 9.63 31

200-400 12.60 3.68 2 12.60 15.20 9.99 31

400-500 18.13 0.41 3 18.01 18.59 17.79 31

500-680 18.78 7.43 4 17.64 8.34 4.65 31

Iron (mg/kg) 0--20 340 1 31

20--100 70 40 2 70 110 40 31

100-150 80 20 3 70 100 70 31

200-400 80 20 2 80 90 60 31

400-500 80 10 3 60 70 60 31

500-680 40 10 4 40 50 30 31

Loss on ignition (%) 0--20 18.33 1 31

20--100 13.26 0.32 2 13.26 13.49 13.03 31

100-150 14.22 0.7 3 14.34 14.86 13.47 31

200-400 13.30 0.86 2 13.30 13.90 12.69 31

400-500 12.13 0.19 3 12.07 12.34 11.99 31

500-680 11.09 0.99 4 11.36 11.90 9.72 31

Magnesium (cmol/kg-1) 0--20 2.66 1 31

20--100 0.67 0.23 2 0.67 0.83 0.50 31

100-150 0.52 0.26 3 0.43 0.82 0.32 31

200-400 0.73 0.60 2 0.73 1.16 0.31 31

400-500 1.78 0.21 3 1.88 1.93 1.53 31

500-680 4.89 3.55 4 4.61 8.50 1.83 31
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

426

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Magnesium (mg/g) 0--20 0.32 1 31

20--100 0.08 0.03 2 0.08 0.10 0.06 31

100-150 0.06 0.03 3 0.05 0.10 0.04 31

200-400 0.09 0.07 2 0.09 0.14 0.04 31

400-500 0.21 0.03 3 0.23 0.23 0.18 31

Magnesium (mg/g) 500-680 0.59 0.43 4 0.55 1.02 0.22 31

0--20 3.44 1 31

20--100 2.24 1.33 2 2.24 3.18 1.30 31

100-150 2.42 2.14 3 1.68 4.83 0.75 31

200-400 6.78 0.54 2 6.78 7.16 6.39 31

400-500 4.45 1.68 3 3.49 6.39 3.48 31

500-680 1.36 1.75 4 0.58 3.97 0.31 31

Nitrogen (%) 0--20 0.23 1 31

20--100 0.03 0.01 2 0.03 0.04 0.02 31

100-150 0.03 0.02 3 0.02 0.05 0.02 31

200-400 0.02 0.00 2 0.02 0.02 0.01 31

400-500 0.01 0.00 3 0.01 0.02 0.01 31

pH (H2O) 0--20 4.73 1 31

20--100 4.80 0.15 2 4.78 4.88 4.67 31

100-150 4.88 0.14 3 4.96 4.97 4.72 31

200-400 4.54 0.46 2 4.54 4.86 4.21 31

400-500 4.98 0.03 3 4.99 5.00 4.95 31

500-680 5.25 0.43 4 5.29 5.67 4.76 31

pH (KCl) 0--20 4.47 1 31

20--100 3.81 0.11 2 3.81 3.89 3.73 31

100-150 3.72 0.04 3 3.74 3.74 3.67 31
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

427

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

200-400 3.74 0.08 2 3.74 3.80 3.68 31

400-500 3.64 0.04 3 3.64 3.68 3.60 31

500-680 3.72 0.08 4 3.72 3.80 3.64 31

Potassium (mg/g) 0--20 0.18 1 31

20--100 0.05 0.03 2 0.05 0.07 0.03 31

100-150 0.02 0.00 3 0.02 0.02 0.02 31

200-400 0.03 0.01 2 0.03 0.04 0.02 31

400-500 0.06 0.01 3 0.06 0.06 0.05 31

500-680 0.12 0.04 4 0.13 0.14 0.06 31

 Potassium (cmol/kg-l) 0--20 0.47 1 31

20--100 0.14 0.08 2 0.14 0.20 0.08 31

100-150 0.05 0.00 3 0.05 0.05 0.05 31

200-400 0.08 0.03 2 0.08 0.10 0.06 31

400-500 0.15 0.02 3 0.15 0.16 0.13 31

500-680 0.31 0.09 4 0.35 0.36 0.17 31

Sodium (cmol/kg-l) 0--20 0.26 1 31

20--100 0.13 0.02 2 0.13 0.14 0.12 31

100-150 0.12 0.02 3 0.12 0.14 0.10 31

200-400 0.16 0.01 2 0.16 0.17 0.16 31

400-500 0.17 0.00 3 0.18 0.18 0.17 31

500-680 0.27 0.07 4 0.29 0.33 0.17 31

Sodium (mg/g) 0--20 0.06 1 31

20--100 0.03 0.00 2 0.03 0.03 0.03 31

100-150 0.03 0.00 3 0.03 0.03 0.02 31

200-400 0.04 0.00 2 0.04 0.04 0.04 31

400-500 0.04 0.00 3 0.04 0.04 0.04 31
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

428

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

500-680 0.06 0.02 4 0.07 0.08 0.04 31

Sulfur (%) 0--20 0.06 1 31

20--100 0.06 0.01 2 0.06 0.06 0.05 31

100-150 0.05 0.01 3 0.06 0.06 0.04 31

200-400 0.04 0.03 2 0.04 0.05 0.02 31

400-500 0.01 0.00 3 0.01 0.01 0.01 31

Alluvial deposits Oxisols Mfs Carbon (%) 1500 10.29 2.60 399 9.92 23.30 5.11 32 Lagoon sediments

C/N 1500 12 2 399 12 16 4 32 Lagoon sediments

Nitrogen (%) 1500 0.90 0.31 399 0.87 4.40 0.47 32 Lagoon sediments

Alluvial deposits Oxisols Mfs Aluminum (cmol/kg-l) 0--15 0.17 0.05 12 0.18 0.23 0.09 33 Quebradillas

0--15 0.05 0.05 3 0.04 0.11 0.01 33 Isabela

Bulk density(g/cc) 0--15 0.92 0.20 12 0.93 1.23 0.57 33 Quebradillas

Camuy formation Alfisol Mfs 0--15 1.09 0.58 3 1.22 1.59 0.45 33 Isabela

Alluvial deposits Oxisols Mfs C/N 0--15 91 41 12 84 148 42 33 Quebradillas

Camuy formation Alfisol Mfs 0--15 299 392 3 129 747 21 33 Isabela

Alluvial deposits Oxisols Mfs Calcium (cmol/kg-l) 0--15 25.06 9.30 12 26.58 36.75 13.05 33 Quebradillas

Camuy formation Alfisol Mfs 0--15 23.60 23.84 3 19.74 49.13 1.93 33 Isabela

Alluvial deposits Oxisols Mfs Calcium (mg/g) 0--15 5.01 1.86 12 5.32 7.35 2.61 33 Quebradillas

Camuy formation Alfisol Mfs 0--15 4.72 4.77 3 3.95 9.83 0.39 33 Isabela

Alluvial deposits Oxisols Mfs Carbon (%) 0--15 38.51 2.78 12 38.05 42.18 34.45 33 Quebradillas

Camuy formation Alfisol Mfs 0--15 38.37 8.69 3 41.37 45.17 28.58 33 Isabela

Alluvial deposits Oxisols Mfs ECEC (cmol/kg-1) 0--15 28.15 10.06 12 30.178 40.95 14.66 33 Quebradillas

Camuy formation Alfisol Mfs 0--15 30.11 32.32 3 21.77 65.79 2.77 33 Isabela

Alluvial deposits Oxisols Mfs Iron (mg/g) 0--15 0.01 0.05 12 0.01 0.03 0.01 33 Quebradillas

Camuy formation Alfisol Mfs 0--15 0.05 0.06 3 0.01 0.12 0.01 33 Isabela

Alluvial deposits Oxisols Mfs Magnesium (cmol/kg-1) 0--15 2.36 1.03 12 2.39 3.95 0.95 33 Quebradillas
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

429

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Camuy formation Alfisol Mfs 0--15 5.86 8.31 3 1.54 15.44 0.59 33 Isabela

Alluvial deposits Oxisols Mfs Magnesium (mg/g) 0--15 0.28 0.12 12 0.29 0.47 0.11 33 Quebradillas

Camuy formation Alfisol Mfs 0--15 0.70 1.00 3 0.19 1.85 0.07 33 Isabela

Alluvial deposits Oxisols Mfs Manganese (mg/g) 0--15 0.03 0.01 12 0.03 0.04 0.01 33 Quebradillas

Camuy formation Alfisol Mfs 0--15 0.02 0.00 3 0.02 0.02 0.02 33 Isabela

Alluvial deposits Oxisols Mfs Nirogen (%) 0--15 0.50 0.20 12 0.46 0.82 0.28 33 Quebradillas

Camuy formation Alfisol Mfs 0--15 0.58 0.68 3 0.32 1.35 0.06 33 Isabela

Alluvial deposits Oxisols Mfs Organic matter (%) 0--15 7.88 3.55 12 7.01 14.23 3.34 33 Quebradillas

Camuy formation Alfisol Mfs 0--15 9.34 10.34 3 5.93 20.96 1.14 33 Isabela

Alluvial deposits Oxisols Mfs pH (H2O) 0--15 7.69 0.19 12 7.70 7.89 7.16 33 Quebradillas

Camuy formation Alfisol Mfs 0--15 7.23 0.91 3 7.70 7.81 6.19 33 Isabela

Alluvial deposits Oxisols Mfs pH (KCl) 0--15 7.14 0.22 12 7.18 7.37 6.51 33 Quebradillas

Camuy formation Alfisol Mfs 0--15 6.61 1.07 3 7.19 7.26 5.38 33 Isabela

Alluvial deposits Oxisols Mfs Phosphorus (mg/g) 0--15 0.05 0.03 12 0.04 0.10 0.02 33 Quebradillas

Camuy formation Alfisol Mfs 0--15 0.03 0.01 3 0.20 0.04 0.02 33 Isabela

Alluvial deposits Oxisols Mfs Potassium (mg/g) 0--15 0.16 0.07 12 0.16 0.31 0.07 33 Quebradillas

Camuy formation Alfisol Mfs 0--15 0.11 0.03 3 0.12 0.13 0.08 33 Isabela

Alluvial deposits Oxisols Mfs Potassium (cmol/kg-l) 0--15 0.42 0.19 12 0.41 0.81 0.19 33 Quebradillas

Camuy formation Alfisol Mfs 0--15 0.30 0.07 3 0.33 0.35 0.22 33 Isabela

Alluvial deposits Oxisols Mfs Sodium (cmol/kg-l) 0--15 0.14 0.09 12 0.12 0.33 0.04 33 Quebradillas

Camuy formation Alfisol Mfs 0--15 0.30 0.40 3 0.13 0.76 0.03 33 Isabela

Alluvial deposits Oxisols Mfs Sodium (mg/g) 0--15 0.03 0.02 12 0.03 0.08 0.01 33 Quebradillas

Camuy formation Alfisol Mfs 0--15 0.07 0.09 3 0.03 0.17 0.01 33 Isabela

Tuffaceous sandstone Inceptisols Rfs Aluminum (cmol/kg-l) 0--10 5.75 2.20 20 5.46 10.10 2.82 34 Ridge

0--10 6.21 2.30 19 7.10 9.46 1.16 34 Riparian

0--10 5.18 1.25 19 4.84 7.57 3.20 34 Slope
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

430

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--10 5.33 1.53 20 5.04 9.44 2.93 34 Slope-ridge

0--10 5.15 1.72 21 5.24 7.90 2.02 34 Slope-riparian

0--10 3.91 1.50 20 3.37 6.59 1.82 34 Streambank

10--25 4.27 1.16 20 3.88 7.26 2.94 34 Ridge

10--25 5.24 2.12 20 5.80 8.44 0.64 34 Riparian

10--25 4.41 1.15 21 4.32 6.11 2.21 34 Slope

10--25 4.08 0.85 19 4.22 6.54 2.72 34 Slope-ridge

10--25 4.32 1.66 20 3.87 7.57 2.59 34 Slope-riparian

10--25 4.57 1.54 20 4.54 7.33 1.93 34 Streambank

25--50 2.68 0.44 19 2.65 3.59 1.94 34 Ridge

25--50 5.54 2.03 20 4.90 9.40 0.64 34 Riparian

25--50 2.42 0.53 18 2.42 3.72 1.68 34 Slope

25--50 2.91 0.52 21 2.88 3.87 2.20 34 Slope-ridge

25--50 4.32 1.81 19 3.79 8.68 2.46 34 Streambank

Calcium (cmol/kg-l) 0--10 0.57 0.63 20 0.38 2.83 0.15 34 Ridge

0--10 1.30 1.13 19 1.04 4.07 0.18 34 Riparian

0--10 0.27 0.24 19 0.15 0.74 0.07 34 Slope

0--10 0.43 0.20 20 0.42 0.86 0.16 34 Slope-ridge

0--10 0.50 0.45 21 0.27 1.72 0.10 34 Slope-riparian

0--10 1.39 1.12 20 0.83 2.85 0.24 34 Streambank

10--25 0.11 0.06 20 0.15 0.23 0.04 34 Ridge

10--25 1.14 1.88 20 0.54 8.57 0.04 34 Riparian

10--25 0.09 0.05 21 0.08 0.22 0.02 34 Slope

10--25 0.13 0.05 19 0.12 0.24 0.06 34 Slope-ridge

10--25 0.20 0.20 20 0.11 0.73 0.04 34 Slope-riparian

10--25 0.80 0.77 20 0.36 2.33 0.07 34 Streambank
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

431

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

25--50 0.03 0.01 19 0.03 0.05 0.01 34 Ridge

25--50 0.86 1.41 20 0.46 6.47 0.05 34 Riparian

25--50 0.03 0.03 18 0.02 0.12 0.01 34 Slope

25--50 0.03 0.01 21 0.03 0.05 0.01 34 Slope-ridge

25--50 0.49 0.57 19 0.23 2.05 0.04 34 Streambank

Calcium (mg/kg) 0--10 114 126 20 77 566 31 34 Ridge

0--10 260 225 19 209 814 36 34 Riparian

0--10 54 48 19 31 148 14 34 Slope

0--10 86 41 20 83 171 31 34 Slope-ridge

0--10 100 89 21 53 344 21 34 Slope-riparian

0--10 279 225 20 165 570 47 34 Streambank

10--25 23 11 20 21 47 8 34 Ridge

10--25 227 376 20 108 1713 8 34 Riparian

10--25 18 10 21 16 44 4 34 Slope

10--25 25 10 19 24 48 12 34 Slope-ridge

10--25 40 39 20 23 146 8 34 Slope-riparian

10--25 159 154 20 72 465 15 34 Streambank

25--50 5 2 19 5 10 2 34 Ridge

25--50 173 281 20 91 1294 10 34 Riparian

25--50 5 5 18 5 24 1 34 Slope

25--50 6 2 21 6 11 3 34 Slope-ridge

25--50 97 113 19 45 409 9 34 Streambank

Carbon (%) 0--10 5.08 2.07 20 4.37 11.10 3.03 34 Ridge

0--10 7.28 2.71 19 6.35 13.25 3.34 34 Riparian

0--10 4.68 1.57 19 4.33 7.70 2.10 34 Slope

0--10 4.26 1.07 20 4.10 6.83 2.75 34 Slope-ridge
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

432

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--10 6.01 1.88 21 5.41 9.81 3.36 34 Slope-riparian

0--10 2.61 1.24 20 2.60 5.26 0.97 34 Streambank

10--25 2.77 1.08 20 2.64 5.28 1.12 34 Ridge

10--25 6.31 2.38 20 6.16 10.25 2.42 34 Riparian

10--25 3.50 1.25 21 3.47 5.70 1.46 34 Slope

10--25 2.72 1.00 19 2.58 4.69 1.50 34 Slope-ridge

10--25 4.92 1.79 20 4.39 7.68 2.82 34 Slope-riparian

10--25 2.62 1.28 20 2.19 5.23 0.98 34 Streambank

25--50 1.17 0.35 19 1.12 1.88 0.65 34 Ridge

25--50 5.23 2.72 20 4.36 13.10 1.40 34 Riparian

25--50 1.35 0.61 18 1.22 2.97 0.71 34 Slope

25--50 1.07 0.20 21 1.03 1.61 0.71 34 Slope-ridge

25--50 1.91 1.27 19 1.37 4.92 0.81 34 Streambank

Effective cation exchange capacity 0--10 7.17 2.80 20 6.48 14.62 3.08 34 Ridge

{Ca+K+Mg(cmol/kg-1) 0--10 8.63 1.83 19 8.12 11.89 5.15 34 Riparian

0--10 6.03 1.44 19 5.49 8.83 3.74 34 Slope

0--10 6.45 1.84 20 6.14 11.55 3.99 34 Slope-ridge

0--10 6.26 1.95 21 5.88 9.99 2.58 34 Slope-riparian

0--10 6.17 1.72 20 6.45 9.58 3.63 34 Streambank

10--25 4.77 1.27 20 4.26 8.03 3.27 34 Ridge

10--25 7.11 1.43 20 7.38 10.19 4.40 34 Riparian

10--25 4.87 1.21 21 4.95 6.75 2.44 34 Slope

10--25 4.54 0.94 19 4.55 7.33 3.13 34 Slope-ridge

10--25 4.90 1.81 20 4.31 8.37 3.01 34 Slope-riparian

10--25 6.01 1.22 20 5.73 8.39 3.96 34 Streambank

25--50 2.84 0.47 19 2.86 3.81 2.02 34 Ridge
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

433

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

25--50 7.09 1.82 20 7.30 10.67 4.35 34 Riparian

25--50 2.58 0.59 18 2.56 3.95 1.85 34 Slope

25--50 3.06 0.52 21 3.06 4.08 2.26 34 Slope-ridge

25--50 5.32 2.01 19 4.75 9.43 3.06 34 Streambank

Iron (mg/g) 0--10 2.01 0.63 20 1.72 3.20 1.43 34 Ridge

0--10 1.01 0.47 19 1.05 1.94 0.49 34 Riparian

0--10 1.89 0.72 19 1.74 3.37 0.88 34 Slope

0--10 1.88 0.47 20 2.02 2.74 1.00 34 Slope-ridge

0--10 1.56 0.42 21 1.62 2.50 0.91 34 Slope-riparian

0--10 0.73 0.35 20 0.59 1.59 0.33 34 Streambank

10--25 1.49 0.49 20 1.47 2.50 0.69 34 Ridge

10--25 0.79 0.35 20 0.69 1.60 0.25 34 Riparian

10--25 1.51 0.43 21 1.58 2.37 0.78 34 Slope

10--25 1.20 0.43 19 1.17 2.29 0.12 34 Slope-ridge

10--25 1.18 0.29 20 1.22 1.62 0.50 34 Slope-riparian

10--25 0.66 0.18 20 0.59 1.04 0.46 34 Streambank

25--50 0.56 0.30 19 0.44 1.37 0.24 34 Ridge

25--50 0.68 0.38 20 0.52 1.40 0.22 34 Riparian

25--50 0.72 0.35 18 0.59 0.13 0.18 34 Slope

25--50 0.49 0.19 21 0.46 1.24 0.30 34 Slope-ridge

25--50 0.68 0.32 19 0.56 1.57 0.36 34 Streambank

Loss on ignition (%) 0--10 13.42 4.60 20 13.13 25.56 7.86 34 Ridge

0--10 21.38 3.89 19 21.55 29.58 12.57 34 Riparian

0--10 12.14 2.30 19 12.28 15.99 6.80 34 Slope

0--10 12.28 2.66 20 12.21 18.71 8.06 34 Slope-ridge

0--10 13.49 4.11 21 12.06 25.90 8.49 34 Slope-riparian
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

434

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--10 12.62 4.03 20 13.51 19.44 6.79 34 Streambank

10--25 8.97 2.15 20 8.73 13.88 6.16 34 Ridge

10--25 20.03 3.09 20 20.35 25.26 13.08 34 Riparian

10--25 9.63 1.99 21 10.12 12.66 5.70 34 Slope

10--25 8.47 2.54 19 8.65 14.02 4.51 34 Slope-ridge

10--25 11.94 4.53 20 10.17 21.35 7.23 34 Slope-riparian

10--25 13.55 3.46 20 14.85 18.41 7.93 34 Streambank

25--50 7.09 1.71 19 6.35 10.53 4.75 34 Ridge

25--50 18.60 7.47 20 17.99 47.76 9.67 34 Riparian

25--50 6.41 1.32 18 6.45 8.55 4.18 34 Slope

25--50 6.60 1.27 21 6.19 8.81 4.51 34 Slope-ridge

25--50 11.64 3.85 19 11.96 17.45 1.46 34 Streambank

Magnesium (cmol/kg-l) 0--10 0.49 0.23 20 0.44 1.12 0.24 34 Ridge

0--10 0.71 0.30 19 0.74 1.40 0.24 34 Riparian

0--10 0.29 0.10 19 0.25 0.47 0.15 34 Slope

0--10 0.34 0.18 20 0.29 0.75 0.14 34 Slope-ridge

0--10 0.33 0.24 21 0.21 1.05 0.08 34 Slope-riparian

0--10 0.59 0.29 20 0.66 0.93 0.19 34 Streambank

10--25 0.15 0.07 20 0.13 0.29 0.03 34 Ridge

10--25 0.44 0.24 20 0.47 0.90 0.11 34 Riparian

10--25 0.15 0.07 21 0.14 0.35 0.07 34 Slope

10--25 0.12 0.06 19 0.11 0.28 0.06 34 Slope-ridge

10--25 0.19 0.14 20 0.15 0.50 0.04 34 Slope-riparian

10--25 0.39 0.23 20 0.40 0.81 0.07 34 Streambank

25--50 0.05 0.02 19 0.05 0.09 0.03 34 Ridge

25--50 0.42 0.32 20 0.37 1.13 0.04 34 Riparian
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

435

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

25--50 0.05 0.02 18 0.05 0.11 0.02 34 Slope

25--50 0.04 0.02 21 0.03 0.09 0.01 34 Slope-ridge

25--50 0.24 0.18 19 0.19 0.60 0.04 34 Streambank

Magnesium (mg/kg) 0--10 59 27 20 52 135 28 34 Ridge

0--10 85 36 19 89 168 28 34 Riparian

0--10 34 12 19 31 56 18 34 Slope

0--10 40 21 20 35 90 16 34 Slope-ridge

0--10 40 29 21 26 126 9 34 Slope-riparian

0--10 71 34 20 79 111 23 34 Streambank

10--25 18 9 20 16 35 4 34 Ridge

10--25 53 29 20 57 108 13 34 Riparian

10--25 18 9 21 17 42 8 34 Slope

10--25 14 7 19 13 33 7 34 Slope-ridge

10--25 23 17 20 17 60 5 34 Slope-riparian

10--25 47 27 20 48 97 9 34 Streambank

25--50 6 2 19 6 11 4 34 Ridge

25--50 50 38 20 44 136 5 34 Riparian

25--50 6 3 18 6 13 2 34 Slope

25--50 5 3 21 4 11 1 34 Slope-ridge

25--50 29 22 19 23 72 5 34 Streambank

Manganese (mg/kg) 0--10 5 4 20 4 15 2 34 Ridge

0--10 41 46 19 25 195 4 34 Riparian

0--10 4 2 19 3 8 2 34 Slope

0--10 6 3 20 5 13 2 34 Slope-ridge

0--10 6 5 21 4 22 1 34 Slope-riparian

0--10 80 41 20 90 131 15 34 Streambank
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

436

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--25 2 1 20 2 3 1 34 Ridge

10--25 39 49 20 12 138 2 34 Riparian

10--25 2 1 21 2 4 1 34 Slope

10--25 3 1 19 2 6 1 34 Slope-ridge

10--25 4 3 20 3 11 0.4 34 Slope-riparian

10--25 54 34 20 53 121 5 34 Streambank

25--50 0.75 0.40 19 0.75 1.51 0.04 34 Ridge

25--50 29 36 20 19 138 1 34 Riparian

25--50 1.6 1.6 17 1.0 6.1 0.3 34 Slope

25--50 1.6 1.1 17 1.2 4.2 0.3 34 Slope-ridge

25--50 35 26 19 35 83 3 34 Streambank

Nitrogen (%) 0--10 0.22 0.08 20 0.22 0.40 0.03 34 Ridge

0--10 0.33 0.09 19 0.30 0.55 0.22 34 Riparian

0--10 0.23 0.08 19 0.23 0.45 0.10 34 Slope

0--10 0.21 0.05 20 0.21 0.33 0.14 34 Slope-ridge

0--10 0.25 0.09 21 0.23 0.44 0.13 34 Slope-riparian

0--10 0.13 0.07 20 0.13 0.27 0.05 34 Streambank

10--25 0.14 0.05 20 0.13 0.23 0.07 34 Ridge

10--25 0.28 0.07 20 0.27 0.43 0.16 34 Riparian

10--25 0.17 0.06 21 0.17 0.29 0.08 34 Slope

10--25 0.14 0.05 19 0.14 0.23 0.08 34 Slope-ridge

10--25 0.20 0.08 20 0.17 0.33 0.11 34 Slope-riparian

10--25 0.15 0.07 20 0.15 0.27 0.06 34 Streambank

25--50 0.07 0.02 19 0.06 0.13 0.03 34 Ridge

25--50 0.19 0.08 20 0.18 0.39 0.06 34 Riparian

25--50 0.06 0.02 18 0.06 0.12 0.04 34 Slope
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

437

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

25--50 0.06 0.01 21 0.06 0.08 0.04 34 Slope-ridge

25--50 0.10 0.06 19 0.08 0.22 0.04 34 Streambank

pH (H2O) 0--10 4.28 0.14 20 4.29 4.47 3.94 34 Ridge

0--10 4.63 0.31 19 4.69 5.23 4.15 34 Riparian

0--10 4.32 0.23 19 4.36 4.64 3.78 34 Slope

0--10 4.31 0.16 20 4.28 4.72 4.05 34 Slope-ridge

0--10 4.55 0.24 21 4.58 4.98 4.15 34 Slope-riparian

0--10 4.89 0.21 20 4.96 5.17 4.41 34 Streambank

10--25 4.42 0.16 20 4.42 4.62 4.10 34 Ridge

10--25 4.77 0.49 20 4.80 6.51 4.31 34 Riparian

10--25 4.38 0.15 21 4.41 4.62 4.04 34 Slope

10--25 4.43 0.11 19 4.42 4.66 4.24 34 Slope-ridge

10--25 4.62 0.21 20 4.60 5.08 4.20 34 Slope-riparian

10--25 4.82 0.19 20 4.90 5.10 4.49 34 Streambank

25--50 4.70 0.15 19 4.72 4.98 4.26 34 Ridge

25--50 4.79 0.28 20 4.25 5.55 4.37 34 Riparian

25--50 4.60 0.24 18 4.25 5.08 4.33 34 Slope

25--50 4.52 0.24 21 4.57 4.86 4.12 34 Slope-ridge

25--50 4.75 0.21 19 4.75 5.10 4.38 34 Streambank

pH (KCl) 0--10 3.55 0.13 20 3.58 3.73 3.18 34 Ridge

0--10 3.87 0.19 19 3.91 4.28 3.56 34 Riparian

0--10 3.66 0.16 19 3.71 3.88 3.39 34 Slope

0--10 3.66 0.13 20 3.68 3.89 3.43 34 Slope-ridge

0--10 3.78 0.17 21 3.81 4.00 3.52 34 Slope-riparian

0--10 3.94 0.14 20 3.96 4.23 3.67 34 Streambank

10--25 3.82 0.12 20 3.79 4.09 3.58 34 Ridge
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

438

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--25 4.04 0.45 20 3.96 5.67 3.71 34 Riparian

10--25 3.73 0.11 21 3.73 3.94 3.53 34 Slope

10--25 3.81 0.08 19 3.80 3.96 3.62 34 Slope-ridge

10--25 3.89 0.13 20 3.92 4.04 3.64 34 Slope-riparian

10--25 3.92 0.09 20 3.91 4.04 3.75 34 Streambank

25--50 4.01 0.06 19 4.02 4.09 3.87 34 Ridge

25--50 3.95 0.21 20 3.93 4.64 3.52 34 Riparian

25--50 4.00 0.09 18 4.03 4.13 3.78 34 Slope

25--50 4.01 0.05 21 4.01 4.10 3.93 34 Slope-ridge

25--50 3.95 0.09 19 3.96 4.10 3.78 34 Streambank

Phosphorus (mg/kg) 0--10 7 2 20 7 12 4 34 Ridge

0--10 14 4 19 14 24 10 34 Riparian

0--10 8 3 19 7 15 4 34 Slope

0--10 6 1 20 6 10 4 34 Slope-ridge

0--10 15 3 21 14 20 10 34 Slope-riparian

0--10 6 2 20 6 10 3 34 Streambank

10--25 3.5 2.4 20 3.2 9.5 0.5 34 Ridge

10--25 12 4 20 11 21 6 34 Riparian

10--25 6 3 21 6 12 2 34 Slope

10--25 3 1 19 3 5 1 34 Slope-ridge

10--25 11 3 20 11 17 7 34 Slope-riparian

10--25 6 2 20 6 10 4 34 Streambank

25--50 1 1 11 1 3 1 34 Ridge

25--50 8 3 20 8 17 1 34 Riparian

25--50 1.4 1.3 17 1.0 6.0 0.3 34 Slope

25--50 1 0 9 1 1 1 34 Slope-ridge
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

439

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

25--50 5 1 19 5 7 1 34 Streambank

Potassium (cmol/kg-l) 0--10 0.16 0.04 20 0.15 0.23 0.11 34 Ridge

0--10 0.21 0.06 19 0.19 0.32 0.10 34 Riparian

0--10 0.14 0.02 19 0.14 0.17 0.11 34 Slope

0--10 0.15 0.03 20 0.15 0.24 0.10 34 Slope-ridge

0--10 0.16 0.05 21 0.16 0.28 0.09 34 Slope-riparian

0--10 0.15 0.06 20 0.15 0.26 0.07 34 Streambank

10--25 0.10 0.03 20 0.10 0.16 0.03 34 Ridge

10--25 0.15 0.04 20 0.14 0.25 0.09 34 Riparian

10--25 0.11 0.03 21 0.10 0.18 0.04 34 Slope

10--25 0.10 0.03 19 0.10 0.16 0.04 34 Slope-ridge

10--25 0.10 0.04 20 0.09 0.22 0.04 34 Slope-riparian

10--25 0.13 0.06 20 0.14 0.24 0.04 34 Streambank

25--50 0.04 0.01 17 0.03 0.07 0.01 34 Ridge

25--50 0.11 0.06 20 0.11 0.27 0.03 34 Riparian

25--50 0.04 0.03 18 0.03 0.12 0.02 34 Slope

25--50 0.03 0.01 20 0.03 0.05 0.01 34 Slope-ridge

25--50 0.11 0.10 19 0.09 0.45 0.04 34 Streambank

Potassium (mg/g) 0--10 0.06 0.01 20 0.059 0.09 0.04 34 Ridge

0--10 0.08 0.02 19 0.07 0.12 0.04 34 Riparian

0--10 0.05 0.07 19 0.05 0.06 0.04 34 Slope

0--10 0.06 0.01 20 0.06 0.09 0.04 34 Slope-ridge

0--10 0.06 0.02 21 0.06 0.10 0.03 34 Slope-riparian

0--10 0.06 0.02 2 0.06 0.10 0.03 34 Streambank

10--25 0.04 0.01 20 0.04 0.06 0.01 34 Ridge

10--25 0.06 0.02 20 0.05 0.09 0.03 34 Riparian
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

440

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--25 0.04 0.01 21 0.04 0.07 0.02 34 Slope

10--25 0.04 0.01 19 0.04 0.06 0.02 34 Slope-ridge

10--25 0.04 0.02 20 0.03 0.09 0.02 34 Slope-riparian

10--25 0.05 0.02 20 0.05 0.09 0.02 34 Streambank

25--50 0.01 0.01 17 0.01 0.03 0.01 34 Ridge

25--50 0.04 0.02 20 0.04 0.10 0.01 34 Riparian

25--50 0.02 0.01 18 0.01 0.05 0.01 34 Slope

25--50 0.012 0.004 20 0.012 0.019 0.004 34 Slope-ridge

25--50 0.04 0.04 19 0.03 0.17 0.01 34 Streambank

Sodium (cmol/kg-l) 0--10 0.20 0.06 20 0.19 0.34 0.11 34 Ridge

0--10 0.20 0.07 19 0.18 0.36 0.10 34 Riparian

0--10 0.16 0.05 19 0.15 0.25 0.07 34 Slope

0--10 0.21 0.06 20 0.22 0.28 0.12 34 Slope-ridge

0--10 0.13 0.08 21 0.10 0.27 0.03 34 Slope-riparian

0--10 0.14 0.07 20 0.14 0.27 0.03 34 Streambank

10--25 0.14 0.07 20 0.10 0.28 0.06 34 Ridge

10--25 0.14 0.05 20 0.14 0.22 0.04 34 Riparian

10--25 0.11 0.04 21 0.12 0.17 0.04 34 Slope

10--25 0.12 0.04 19 0.11 0.20 0.06 34 Slope-ridge

10--25 0.09 0.05 20 0.08 0.19 0.01 34 Slope-riparian

10--25 0.13 0.08 20 0.12 0.29 0.02 34 Streambank

25--50 0.05 0.02 19 0.05 0.09 0.03 34 Ridge

25--50 0.16 0.07 20 0.16 0.40 0.08 34 Riparian

25--50 0.05 0.03 18 0.04 0.09 0.01 34 Slope

25--50 0.06 0.02 21 0.06 0.09 0.02 34 Slope-ridge

25--50 0.16 0.15 19 0.13 0.69 0.04 34 Streambank
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

441

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Sodium (mg/kg) 0--10 45 13 20 44 77 26 34 Ridge

0--10 45 15 19 42 83 22 34 Riparian

0--10 37 12 19 33 57 16 34 Slope

0--10 48 13 20 50 65 27 34 Slope-ridge

0--10 29 18 21 23 62 7 34 Slope-riparian

0--10 32 16 20 32 61 7 34 Streambank

10--25 32 16 20 24 65 14 34 Ridge

10--25 33 11 20 33 50 9 34 Riparian

10--25 26 8 21 28 39 9 34 Slope

10--25 27 9 19 26 45 13 34 Slope-ridge

10--25 20 11 20 18 44 30 34 Slope-riparian

10--25 30 18 20 27 66 5 34 Streambank

25--50 12 4 19 11 21 6 34 Ridge

25--50 36 17 20 36 92 18 34 Riparian

25--50 10 6 18 10 20 2 34 Slope

25--50 13 4 21 14 20 5 34 Slope-ridge

25--50 37 34 19 30 158 9 34 Streambank

Sulfur (%) 0--10 0.40 0.01 20 0.04 0.07 0.02 34 Ridge

0--10 0.07 0.02 19 0.06 0.12 0.04 34 Riparian

0--10 0.04 0.01 19 0.04 0.61 0.02 34 Slope

0--10 0.04 0.01 20 0.04 0.06 0.02 34 Slope-ridge

0--10 0.05 0.02 21 0.04 0.08 0.03 34 Slope-riparian

0--10 0.03 0.01 20 0.02 0.06 0.01 34 Streambank

10--25 0.03 0.01 20 0.02 0.04 0.02 34 Ridge

10--25 0.06 0.02 20 0.06 0.10 0.04 34 Riparian

10--25 0.03 0.01 21 0.03 0.05 0.01 34 Slope
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

442

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--25 0.03 0.01 19 0.02 0.04 0.02 34 Slope-ridge

10--25 0.04 0.02 20 0.03 0.07 0.02 34 Slope-riparian

10--25 0.03 0.01 20 0.02 0.06 0.01 34 Streambank

25--50 0.02 0.01 19 0.01 0.02 0.01 34 Ridge

25--50 0.05 0.02 20 0.04 0.09 0.02 34 Riparian

25--50 0.02 0.01 18 0.02 0.03 0.01 34 Slope

25--50 0.02 0.00 21 0.01 0.02 0.01 34 Slope-ridge

25--50 0.02 0.01 19 0.02 0.05 0.01 34 Streambank

Tuffaceous sandstone Ultisols Lmrf Organic matter (%) 0--12 2.20 1 35

65--95 1.41 0.23 4 1.43 1.65 1.12 35

105--145 1.89 0.20 4 0.87 1.12 0.60 35

155-195 0.54 0.06 4 0.52 0.61 0.47 35

205-225 0.52 0.19 3 0.47 0.73 0.37 35

Serpentinite Ultisols Lmrf Aluminum (cmol/kg-l) 0--60 0.13 0.12 12 0.10 0.49 0.03 36

Aluminum-total (mg/g) 0--60 112.59 23.17 12 157.79 209.86 79.01 36

Calcium (cmol/kg-l) 0--60 4.13 2.50 12 3.65 8.70 0.43 36

Calcium (mg/g) 0--60 0.83 0.50 12 0.73 1.74 0.09 36

Calcium-total (mg/g) 0--60 118.68 38.35 12 110.92 206.23 75.97 36

Cobalt-total (mg/g) 0--60 0.53 0.08 12 0.38 0.67 0.43 36

Copper-total (mg/g) 0--60 0.02 0.01 12 0.02 0.04 0.01 36

Cromium-total (mg/g) 0--60 6.68 1.17 12 6.86 8.33 5.10 36

ECEC (cmol/kg-1) 0--60 36.98 9.88 12 36.16 54.41 24.86 36

Iron (mg/g) 0--60 0.31 0.17 12 0.25 0.76 0.12 36

Iron-total (mg/g) 0--60 317.14 54.03 12 321.68 409.82 240.03 36

Magnesium (cmol/kg-l) 0--60 32.16 9.61 12 31.36 49.66 18.93 36

Magnesium (mg/g) 0--60 3.78 1.13 12 3.69 5.84 2.23 36
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

443

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Magnesium-total (mg/g) 0--60 26.49 9.23 12 26.60 42.72 13.39 36

Manganese (mg/g) 0--60 0.22 0.11 12 0.20 0.40 0.06 36

Manganese-total (mg/g) 0--60 7.75 1.43 12 7.57 10.22 5.67 36

Nickel-total (mg/g) 0--60 2.44 0.54 12 2.45 3.53 1.51 36

Nitrogen (%) 0--60 0.44 0.24 12 0.35 0.98 0.25 36

Organic matter (%) 0--60 10.65 5.66 12 9.21 24.49 5.00 36

pH (H2O) 0--60 6.10 0.37 12 6.13 6.55 5.17 36

pH (KCl) 0--60 5.41 0.20 11 5.46 5.70 5.09 36

Phosphorus (mg/kg) 0--60 6 4 12 4 13 2 36

Posphorus-total (mg/g) 0--60 0.57 0.16 12 0.55 0.88 0.38 36

Potassium (cmol/kg-l) 0--60 0.31 0.13 12 0.30 0.59 0.16 36

Potassium (mg/g) 0--60 0.12 0.05 12 0.12 0.23 0.06 36

Potassium-total (mg/g) 0--60 1.38 0.37 12 1.34 2.20 0.93 36

Sodium (cmol/kg-l) 0--60 0.24 0.08 12 0.25 0.39 0.13 36

Sodium (mg/g) 0--60 0.05 0.02 12 0.06 0.09 0.03 36

Tuffaceous sandstone Inceptisols Wflm Calcium (mg/g) 0--15 2.65 0.67 5 2.86 3.27 1.91 37

15--30 1.67 0.39 5 1.51 2.19 1.21 37

Iron (mg/g) 0--15 0.42 0.13 5 0.40 0.58 0.29 37

15--30 0.42 0.16 5 0.33 0.65 0.27 37

Loss on ignition (%) 0--15 14.31 1.93 5 15.15 15.64 10.91 37

15--30 12.08 1.42 5 11.76 14.50 10.72 37

Magnesium (mg/g) 0--15 0.43 0.07 5 0.46 0.50 0.35 37

15--30 0.32 0.11 5 0.29 0.50 0.24 37

Manganese (mg/g) 0--15 0.08 0.05 5 0.10 0.13 0.03 37

15--30 0.07 0.03 5 0.07 0.12 0.03 37

Nitrogen (%) 0--15 0.19 0.05 5 0.20 0.23 0.10 37
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

444

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

15--30 0.12 0.02 5 0.11 0.16 0.10 37

Organic matter (%) 0--15 4.51 1.32 5 5.05 5.37 2.20 37

15--30 2.71 0.40 5 2.75 3.10 2.17 37

pH (H2O) 0--15 5.50 0.41 5 5.75 5.86 4.91 37

15--30 5.32 0.17 5 5.29 5.49 5.10 37

pH (KCl) 0--15 4.43 0.53 5 4.70 4.83 3.53 37

15--30 4.09 0.26 5 4.24 4.29 3.66 37

Phosphorus (mg/g) 0--15 0.02 0.00 5 0.02 0.02 0.01 37

15--30 0.02 0.00 5 0.02 0.02 0.01 37

Potassium (mg/g) 0--15 0.10 0.03 5 0.09 0.14 0.08 37

15--30 0.13 0.04 5 0.13 0.17 0.08 37

Tuffaceous sandstone Ultisols Lmrf Aluminum (cmol/kg-l) 0--15 1.89 1.24 9 1.38 3.64 0.35 38 Dieffenbachia seguine

0--15 8.17 6.62 17 6.94 19.47 0.19 38 Ichnanthus pallens

Aluminum (cmol/kg-l) 0--15 1.59 1.82 11 0.79 6.17 0.34 38 Pilea inaequalis

Base saturation (cmol/kg-l) 0--15 22.00 19.00 9 16.00 53.00 3.00 38 Dieffenbachia seguine

0--15 57.48 34.22 17 59.29 93.44 1.71 38 Ichnanthus pallens

0--15 13.05 14.47 11 6.97 48.81 3.09 38 Pilea inaequalis

C/N 0--15 18.00 2.00 10 18.00 21.00 16.00 38 Dieffenbachia seguine

0--15 23.00 5.00 17 22.00 37.00 17.00 38 Ichnanthus pallens

0--15 18.00 3.00 11 17.00 22.00 14.00 38 Pilea inaequalis

Calcium (cmol/kg-l) 0--15 4.66 2.00 10 4.83 7.30 1.34 38 Dieffenbachia seguine

0--15 2.23 2.11 17 1.77 6.62 0.24 38 Ichnanthus pallens

0--15 5.85 1.47 11 5.94 8.22 2.97 38 Pilea inaequalis

Calcium (mg/g) 0--15 0.93 0.40 10 0.97 1.46 0.27 38 Dieffenbachia seguine

0--15 0.45 0.42 17 0.35 1.32 0.05 38 Ichnanthus pallens

0--15 1.17 0.29 11 1.19 1.64 0.59 38 Pilea inaequalis
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

445

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Carbon (%) 0--15 3.98 0.76 10 3.94 5.63 2.89 38 Dieffenbachia seguine

0--15 4.34 1.49 17 4.22 7.30 2.69 38 Ichnanthus pallens

0--15 3.69 0.70 11 3.82 4.98 2.49 38 Pilea inaequalis

Effective cation exchange capacity 0--15 9.52 2.08 10 9.57 12.75 6.81 38 Dieffenbachia seguine

{Ca+K+Mg(cmol/kg-1) 0--15 12.69 4.56 17 11.35 22.34 6.68 38 Ichnanthus pallens

0--15 11.72 1.73 11 11.89 15.30 9.18 38 Pilea inaequalis

Iron (mg/g) 0--15 0.52 0.24 10 0.46 0.99 0.28 38 Dieffenbachia seguine

0--15 1.69 1.00 17 1.84 3.30 0.25 38 Ichnanthus pallens

0--15 0.46 0.19 11 0.40 0.94 0.26 38 Pilea inaequalis

Loss on ignition (%) 0--15 17.56 2.34 10 18.72 19.50 12.31 38 Dieffenbachia seguine

0--15 17.53 3.08 17 17.76 23.36 12.78 38 Ichnanthus pallens

0--15 16.49 1.91 11 16.12 19.75 12.92 38 Pilea inaequalis

Magnesium (cmol/kg-l) 0--15 2.75 0.97 10 2.92 4.25 1.41 38 Dieffenbachia seguine

0--15 1.85 1.15 17 1.72 3.81 0.54 38 Ichnanthus pallens

0--15 3.74 1.47 11 3.36 7.06 2.20 38 Pilea inaequalis

Magnesium (mg/g) 0--15 0.33 0.12 10 0.35 0.51 0.17 38 Dieffenbachia seguine

0--15 0.22 0.14 17 0.21 0.46 0.07 38 Ichnanthus pallens

0--15 0.48 0.18 11 0.40 0.85 0.26 38 Pilea inaequalis

Manganese (mg/kg) 0--15 59 32 10 54 110 14 38 Dieffenbachia seguine

0--15 30 34 17 9 120 3 38 Ichnanthus pallens

0--15 132 40 11 119 202 88 38 Pilea inaequalis

Nitrogen (%) 0--15 0.22 0.05 10 0.22 0.33 0.17 38 Dieffenbachia seguine

0--15 0.20 0.09 17 0.21 0.34 0.07 38 Ichnanthus pallens

0--15 0.21 0.04 11 0.23 0.26 0.14 38 Pilea inaequalis

pH (H2O) 0--15 4.91 0.16 10 4.91 5.20 4.69 38 Dieffenbachia seguine

0--15 4.56 0.36 17 4.40 5.39 4.07 38 Ichnanthus pallens
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

446

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--15 5.05 0.21 11 5.00 5.40 4.72 38 Pilea inaequalis

pH (KCl) 0--15 4.13 0.18 10 4.13 4.44 3.90 38 Dieffenbachia seguine

0--15 3.71 0.42 17 3.65 4.54 3.11 38 Ichnanthus pallens

0--15 4.21 0.18 11 4.24 4.42 3.88 38 Pilea inaequalis

Phosphorus (mg/kg) 0--15 7 2 10 7 10 4 38 Dieffenbachia seguine

0--15 12 5 17 12 24 5 38 Ichnanthus pallens

0--15 7 2 11 7 11 5 38 Pilea inaequalis

Potassium (cmol/kg-l) 0--15 0.20 0.08 10 0.22 0.32 0.10 38 Dieffenbachia seguine

0--15 0.21 0.06 17 0.19 0.31 0.10 38 Ichnanthus pallens

0--15 0.34 0.15 11 0.27 0.74 0.20 38 Pilea inaequalis

Potassium (mg/g) 0--15 0.08 0.03 10 0.08 0.12 0.04 38 Dieffenbachia seguine

0--15 0.08 0.02 17 0.07 0.12 0.04 38 Ichnanthus pallens

0--15 0.13 0.06 11 0.10 0.28 0.08 38 Pilea inaequalis

Sodium (cmol/kg-l) 0--15 0.21 0.05 10 0.21 0.28 0.14 38 Dieffenbachia seguine

0--15 0.23 0.08 17 0.20 0.46 0.12 38 Ichnanthus pallens

0--15 0.21 0.05 11 0.20 0.34 0.16 38 Pilea inaequalis

Sodium (mg/g) 0--15 0.05 0.01 10 0.05 0.07 0.03 38 Dieffenbachia seguine

0--15 0.05 0.02 17 0.05 0.11 0.03 38 Ichnanthus pallens

0--15 0.05 0.01 11 0.05 0.08 0.04 38 Pilea inaequalis

Tuffaceous sandstone Ultisols Swf Clay (%) 0--30 38 15 4 40 52 21 39 Caracoles-El Verde

Tuffaceous sandstone Ultisols Wf-LM 0--30 13 4 4 13 17 8 39 Toro Negro

Lava, tuff Ultisols Swf 0--30 33 16 6 30 55 14 39 Guavate

Limestone Mollisols Smf 0--30 21 29 6 9 77 1 39 Rio Abajo

Limestone Ultisols Smf 0--30 5 4 4 4 9 1 39 Guajataca

Tuffaceous sandstone Inceptisols Wfs 0--30 35 23 4 38 55 11 39 Sabana

Tuffaceous sandstone Ultisols Wfs 0--30 51 1 39 Arboretum
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

447

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Tuffaceous sandstone Ultisols Swf pH (H2O) 0--30 4.65 0.39 4 4.55 5.20 4.30 39 Caracoles-El Verde

Tuffaceous sandstone Ultisols Wf-LM 0--30 5.08 0.05 4 5.10 5.10 5.00 39 Toro Negro

Lava, tuff Ultisols Swf 0--30 4.77 0.14 6 4.75 5.00 4.60 39 Guavate

Limestone Mollisols Smf 0--30 6.02 1.03 6 6.20 7.30 4.80 39 Rio Abajo

Limestone Ultisols Smf 0--30 7.10 0.45 4 7.15 7.60 6.50 39 Guajataca

Tuffaceous sandstone Inceptisols Wfs 0--30 5.28 0.55 4 5.15 6.00 4.80 39 Sabana

Tuffaceous sandstone Ultisols Wfs 0--30 5.16 1 39 Arboretum

Tuffaceous sandstone Ultisols Swf Sand (%) 0--30 34 16 4 4 55 20 39 Caracoles-El Verde

Tuffaceous sandstone Ultisols Wf-LM 0--30 53 8 4 53 60 45 39 Toro Negro

Lava, tuff Ultisols Swf 0--30 23 15 6 15 44 11 39 Guavate

Limestone Mollisols Smf 0--30 42 23 6 44 66 7 39 Rio Abajo

Limestone Ultisols Smf 0--30 33 9 4 32 44 25 39 Guajataca

Tuffaceous sandstone Inceptisols Wfs 0--30 16 9 4 17 25 6 39 Sabana

Tuffaceous sandstone Ultisols Wfs 0--30 16 1 39 Arboretum

Tuffaceous sandstone Ultisols Swf Silt (%) 0--30 27 4 4 28 31 24 39 Caracoles-El Verde

Tuffaceous sandstone Ultisols Wf-LM 0--30 34 4 4 35 38 29 39 Toro Negro

Lava, tuff Ultisols Swf 0--30 44 10 6 41 59 34 39 Guavate

Limestone Mollisols Smf 0--30 37 13 6 39 54 16 39 Rio Abajo

Limestone Ultisols Smf 0--30 62 5 4 63 67 55 39 Guajataca

Tuffaceous sandstone Inceptisols Wfs 0--30 49 14 4 48 64 35 39 Sabana

Tuffaceous sandstone Ultisols Wfs 0--30 33 1 39 Arboretum

Limestone Mollisols Smf Calcium (mg/kg) 0--30 4891 2861 10 3988 9119 1372 40 Rio Abajo

Blanket deposits Alfisols Smf 0--30 1598 910 12 1476 3216 173 40 Cambalache forest

Lava, tuff Ultisols Swf 0--30 5528 1729 2 5528 6750 4305 40 Cayey

0--30 2425 1260 2 2425 3316 1534 40 Guavate forest

Plutonic rocks Inceptisols Mfs 0--30 1343 1 40 Patillas
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

448

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Tuffaceous sandstone Ultisols Wfs 0--30 6600 804 4 6504 7634 5760 40 Caribbean National Forest

Limestone Mollisols Smf Effective cation exchange capacity 0--30 29 13 10 24 48 9 40 Rio Abajo

Blanket deposits Alfisols Smf {Ca+K+Mg(cmol/kg-1) 0--30 11 4 12 10 19 4 40 Cambalache forest

Lava, tuff Ultisols Swf 0--30 38 12 2 38 46 30 40 Cayey

0--30 18 5 2 18 22 14 40 Guavate forest

Plutonic rocks Inceptisols Mfs 0--30 9 1 40 Patillas

Tuffaceous sandstone Ultisols Wfs 0--30 45 4 4 45 49 40 40 Caribbean National Forest

Limestone Mollisols Smf Magnesium (mg/kg) 0--30 273 109 10 252 486 161 40 Rio Abajo

Blanket deposits Alfisols Smf 0--30 119 49 12 112 204 46 40 Cambalache forest

Lava, tuff Ultisols Swf 0--30 1156 344 2 1156 1399 912 40 Cayey

0--30 548 59 2 548 589 506 40 Guavate forest

Plutonic rocks Inceptisols Mfs 0--30 204 1 40 Patillas

Tuffaceous sandstone Ultisols Wfs 0--30 1267 131 4 1232 1446 1158 40 Caribbean National Forest

Limestone Mollisols Smf Potassium (mg/kg) 0--30 260 54 10 268 371 172 40 Rio Abajo

Blanket deposits Alfisols Smf 0--30 226 52 12 229 292 141 40 Cambalache forest

Lava, tuff Ultisols Swf 0--30 258 40 2 258 286 229 40 Cayey

0--30 303 73 2 203 254 151 40 Guavate forest

Plutonic rocks Inceptisols Mfs 0--30 131 1 40 Patillas

Tuffaceous sandstone Ultisols Wfs 0--30 279 17 4 279 299 259 40 Caribbean National Forest

Limestone Mollisols Smf Sodium (mg/kg) 0--30 35 18 10 29 82 21 40 Rio Abajo

Blanket deposits Alfisols Smf 0--30 16 8 12 13 37 8 40 Cambalache forest

Lava, tuff Ultisols Swf 0--30 77 7 2 77 82 73 40 Cayey

0--30 57 28 2 57 77 38 40 Guavate forest

Plutonic rocks Inceptisols Mfs 0--30 56 1 40 Patillas

Tuffaceous sandstone Ultisols Wfs 0--30 60 37 4 65 100 11 40 Caribbean National Forest

Tuffaceous sandstone Ultisols Lmrf Aluminum (cmol/kg-l) 0--10 6.98 1.29 11 6.64 8.74 4.94 41
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

449

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--20 6.10 0.95 11 6.02 8.11 4.36 41

20--30 5.64 0.77 11 5.75 6.88 3.87 41

Calcium (cmol/kg-l) 0--10 1.21 0.97 11 0.89 3.74 0.39 41

10--20 0.51 0.37 11 0.33 1.30 0.16 41

20--30 0.26 0.17 11 0.24 0.60 0.05 41

Calcium (mg/g) 0--10 0.24 0.19 11 0.18 0.75 0.08 41

10--20 0.10 0.07 11 0.07 0.26 0.03 41

20--30 0.05 0.03 11 0.05 0.12 0.01 41

Effective cation exchange capacity 0--10 10.77 1.24 11 10.63 12.93 8.64 41

{Ca+K+Mg(cmol/kg-1) 10--20 8.36 0.99 11 8.25 10.10 6.53 41

20--30 7.41 1.17 11 7.23 9.70 5.23 41

Iron (mg/g) 0--10 2.11 0.40 11 2.00 2.86 1.54 41

10--20 1.78 0.49 11 1.59 2.75 1.34 41

20--30 1.63 0.39 11 1.52 2.60 1.16 41

Magnesium (cmol/kg-l) 0--10 1.75 0.29 11 1.65 2.26 1.37 41

10--20 1.13 0.20 11 1.07 1.62 0.87 41

20--30 0.87 0.30 11 0.72 1.53 0.57 41

Magnesium (mg/g) 0--10 0.21 0.03 1 0.20 0.27 0.16 41

10--20 0.14 0.02 11 0.13 0.19 0.10 41

20--30 0.10 0.04 11 0.09 0.18 0.07 41

Manganese (mg/kg) 0--10 4 2 11 3 8 2 41

10--20 3 1 11 2 5 1 41

20--30 2 1 11 2 50 1 41

Organic matter (%) 0--10 10.19 3.21 11 9.40 17.20 7.23 41

10--20 6.90 2.40 11 6.37 11.00 2.75 41

20--30 12.36 2.59 10 12.38 15.77 6.37 41
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

450

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

pH (H2O) 0--10 3.96 0.12 11 3.97 4.18 3.72 41

10--20 4.02 0.16 11 4.05 4.18 3.64 41

20--30 4.12 0.14 11 4.16 4.29 3.89 41

pH (KCl) 0--10 3.47 0.11 11 3.48 3.62 3.31 41

10--20 3.55 0.11 11 3.55 3.72 3.36 41

20--30 3.60 0.09 11 3.60 3.77 3.43 41

Phosphorus (mg/kg) 0--10 9 2 11 10 11 6 41

10--20 6 2 11 5 8 4 41

20--30 4 1 11 4 6 2 41

Potassium (cmol/kg-l) 0--10 0.17 0.10 11 0.13 0.36 0.07 41

10--20 0.12 0.08 11 0.09 0.27 0.04 41

20--30 0.10 0.08 11 0.06 0.23 0.02 41

Potassium (mg/g) 0--10 0.07 0.04 11 0.05 0.14 0.03 41

10--20 0.05 0.03 11 0.04 0.10 0.02 41

20--30 0.04 0.03 11 0.02 0.09 0.01 41

Sodium (cmol/kg-l) 0--10 0.66 0.11 11 0.65 0.87 0.46 41

10--20 0.90 0.08 11 0.50 0.59 0.35 41

20--30 0.54 0.51 11 0.38 2.04 0.25 41

Sodium (mg/g) 0--10 0.15 0.03 11 0.15 0.20 0.11 41

10--20 0.11 0.02 11 0.11 0.14 0.08 41

Tuffaceous sandstone Ultisols Lmrf 20--30 0.12 0.12 11 0.09 0.47 0.06 41

Tuffaceous sandstone Inceptisols Smf Clay (%) 0--10 36 13 8 40 47 13 42

10--20 32 12 8 36 43 13 42

20--30 28 12 8 28 45 5 42

30--40 26 12 8 31 39 6 42

Organic matter (%) 0--10 5.45 2.30 8 5.41 9.32 2.74 42
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

451

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--20 3.39 2.07 8 2.82 6.98 1.30 42

20--30 1.99 1.11 8 1.66 4.32 0.85 42

30--40 1.71 0.98 8 1.68 3.39 0.69 42

pH (H2O) 0--10 7.16 1.01 8 6.94 8.68 5.75 42

10--20 7.47 1.05 8 7.16 9.07 6.15 42

20--30 7.83 0.74 8 7.50 8.96 6.93 42

30--40 7.61 0.71 8 7.57 8.68 6.89 42

Sand (%) 0--10 33 25 8 19 75 15 42

10--20 37 24 8 26 76 13 42

20--30 42 25 8 40 89 12 42

30--40 47 29 8 38 91 15 42

Silt (%) 0--10 32 13 8 36 46 12 42

10--20 32 13 8 34 47 11 42

20--30 25 15 8 25 43 4 42

30--40 27 16 8 30 47 3 42

Alluvial deposits Inceptisols Mfs Bulk density (g/cc) 0--8 1.40 1 43 Yabucoa

0--19 0.97 1 43 Yabucoa

36--43 1.54 1 43 Yabucoa

Limestone Mollisols Smf 0--8 0.66 0.18 2 0.66 0.79 0.53 43 Rio Abajo

15--23 1.07 1 43 Rio Abajo

Alluvial deposits Inceptisols Mfs Calcium-total (mg/g) 0--8 1.70 1 43 Yabucoa

0--19 0.78 1 43 Yabucoa

Alluvial deposits Inceptisols Mfs 36--43 1.03 1 43 Yabucoa

Limestone Mollisols Smf 0--8 9.28 2.01 2 9.28 10.70 7.86 43 Rio Abajo

15--23 25.06 1 43 Rio Abajo

Alluvial deposits Inceptisols Mfs Clay (%) 0--8 7 1 43 Yabucoa
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

452

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--19 29 1 43 Yabucoa

36--43 10 1 43 Yabucoa

Limestone Mollisols Smf 0--8 27 20 2 27 41 13 43 Rio Abajo

15--23 11 1 43 Rio Abajo

Alluvial deposits Inceptisols Mfs Magnesium-total (mg/g) 0--8 1.34 1 43 Yabucoa

0--19 1.70 1 43 Yabucoa

36--43 2.24 1 43 Yabucoa

Limestone Mollisols Smf 0--8 9.27 1.02 2 9.27 9.99 8.55 43 Rio Abajo

15--23 6.96 1 43 Rio Abajo

Alluvial deposits Inceptisols Mfs Nitrogen (%) 0--8 0.16 1 43 Yabucoa

0--19 0.11 1 43 Yabucoa

36--43 0.06 1 43 Yabucoa

Limestone Mollisols Smf 0--8 0.30 0.11 2 0.30 0.38 0.22 43 Rio Abajo

15--23 0.33 1 43 Rio Abajo

Alluvial deposits Inceptisols Mfs Organic matter (%) 0--8 2.29 1 43 Yabucoa

0--19 1.70 1 43 Yabucoa

Alluvial deposits Inceptisols Mfs 36--43 0.65 1 43 Yabucoa

Limestone Mollisols Smf 0--8 4.16 2.52 2 4.16 5.94 2.37 43 Rio Abajo

15--23 4.91 1 43 Rio Abajo

Alluvial deposits Inceptisols Mfs Phosphorus-total (mg/g) 0--8 0.45 1 43 Yabucoa

0--19 0.33 1 43 Yabucoa

36--43 0.32 1 43 Yabucoa

Limestone Mollisols Smf 0--8 0.62 0.29 2 0.62 0.82 0.41 43 Rio Abajo

15--23 1.12 1 43 Rio Abajo

Alluvial deposits Inceptisols Mfs Potassium-total (mg/g) 0--8 1.48 1 43 Yabucoa

0--19 1.61 1 43 Yabucoa
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

453

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

36--43 1.27 1 43 Yabucoa

Limestone Mollisols Smf 0--8 2.80 0.25 2 2.80 2.98 2.63 43 Rio Abajo

15--23 3.72 1 43 Rio Abajo

Alluvial deposits Inceptisols Mfs Sand (%) 0--8 72 1 43 Yabucoa

0--19 41 1 43 Yabucoa

Alluvial deposits Inceptisols Mfs 36--43 66 1 43 Yabucoa

Limestone Mollisols Smf 0--8 27 23 2 27 43 11 43 Rio Abajo

Limestone Mollisols Smf 15--23 53 1 43 Rio Abajo

Alluvial deposits Inceptisols Mfs Silt (%) 0--8 21 1 43 Yabucoa

0--19 30 1 43 Yabucoa

Alluvial deposits Inceptisols Mfs 36--43 24 1 43 Yabucoa

Limestone Mollisols Smf 0--8 46 3 2 46 48 44 43 Rio Abajo

15--23 36 1 43 Rio Abajo

Alluvial deposits Inceptisols Mfs Sodium-total (mg/g) 0--8 0.47 1 43 Yabucoa

0--19 0.33 1 43 Yabucoa

36--43 0.32 1 43 Yabucoa

Limestone Mollisols Smf 0--8 0.78 0.01 2 0.78 0.79 0.77 43 Rio Abajo

15--23 0.99 1 43 Rio Abajo

Lava Ultisols Wfs Calcium (%) 0--6 0.06 0.01 4 0.06 0.08 0.05 44

5--20 0.05 0.01 5 0.05 0.06 0.05 44

12--39 0.06 0.01 3 0.06 0.07 0.05 44

28--58 0.06 0.01 4 0.07 0.07 0.05 44

Organic matter (%) 0--6 7.83 1.76 4 8.43 9.18 5.29 44

5--20 2.41 1.04 5 2.01 4.01 1.51 44

12--39 0.80 0.07 3 0.76 0.88 0.75 44

28--58 0.32 0.13 4 0.38 0.38 0.12 44
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

454

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

pH (H2O) 0--6 4.22 0.28 4 4.16 4.61 3.95 44

5--20 4.05 0.10 5 4.01 4.16 3.92 44

12--39 4.11 0.09 3 4.13 4.18 4.01 44

28--58 4.03 0.11 4 4.01 4.17 3.91 44

Phosphorus (%) 0--6 0.02 0.00 4 0.02 0.02 0.01 44

5--20 0.01 0.01 5 0.01 0.02 0.01 44

12--39 0.01 0.00 3 0.01 0.01 0.01 44

28--58 0.02 0.00 4 0.02 0.02 0.02 44

Potassium (%) 0--6 0.07 0.02 4 0.07 0.09 0.06 44

5--20 0.08 0.02 5 0.07 0.10 0.06 44

12--39 0.04 0.03 3 0.06 0.06 0.01 44

28--58 0.07 0.03 4 0.06 0.10 0.04 44

Tuffaceous sandstone Ultisols Wfs Aluminum (cmol/kg-1) 0--10 6.20 0.76 4 6.25 6.90 5.40 45 Khaya nyassica

0--10 5.60 0.10 3 5.60 5.70 5.50 45 Pinus elliottii

0--10 5.70 0.56 3 5.60 6.30 5.20 45 Terminalia ivorensis

0--10 3.73 2.48 3 3.00 6.50 1.70 45 Anthocephalus cadamba

0--10 5.43 0.66 4 5.65 5.90 4.50 45 Swietenia macrophylla

0--10 3.37 1.54 3 4.10 4.40 1.60 45 Hibiscus elatus

0--10 4.20 1.73 3 3.80 6.10 2.70 45 Eucalyptus saligna

0--10 4.40 1.40 3 3.80 6.00 3.40 45 Pinus caribaea var. hondurensis

0--10 2.67 0.38 3 2.50 3.10 2.40 45 Hernandia sonora

0--10 15.85 24.81 4 4.40 53.00 1.60 45 Eucalyptus pathentinervis

Aluminum-total (mg/g) 0--10 46.05 1.79 4 45.39 48.69 44.71 45 Khaya nyassica

0--10 62.85 6.89 4 65.22 68.24 52.74 45 Pinus elliottii

0--10 57.85 6.89 4 55.49 67.71 52.51 45 Terminalia ivorensis

0--10 39.09 8.68 4 35.20 52.06 33.90 45 Anthocephalus cadamba
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

455

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--10 65.38 8.32 4 68.35 71.36 53.46 45 Swietenia macrophylla

0--10 91.26 8.51 4 93.93 98.01 79.16 45 Hibiscus elatus

0--10 97.62 8.10 4 97.67 104.94 90.19 45 Eucalyptus saligna

0--10 59.72 27.91 4 55.28 95.98 32.50 45 Pinus caribaea var. hondurensis

0--10 32.60 5.25 4 33.82 36.98 25.77 45 Hernandia sonora

0--10 35.72 2.95 4 36.70 38.07 31.40 45 Eucalyptus pathentinervis

Bulk density (g/cc) 0--10 0.60 0.11 4 0.56 0.77 0.53 45 Khaya nyassica

0--10 0.73 0.08 4 0.73 0.81 0.63 45 Pinus elliottii

0--10 0.45 0.04 4 0.45 0.51 0.41 45 Terminalia ivorensis

0--10 0.56 0.08 4 0.57 0.63 0.46 45 Anthocephalus cadamba

0--10 0.57 0.08 5 0.56 0.66 0.48 45 Swietenia macrophylla

0--10 0.69 0.12 4 0.70 0.82 0.54 45 Hibiscus elatus

0--10 0.54 0.06 4 0.54 0.61 0.47 45 Eucalyptus saligna

0--10 0.81 0.10 4 0.83 0.91 0.69 45 Pinus caribaea var. hondurensis

0--10 0.50 0.12 4 0.52 0.62 0.33 45 Hernandia sonora

0--10 0.72 0.14 5 0.74 0.88 0.50 45 Eucalyptus pathentinervis

Calcium (mg/kg) 0--10 299 55 4 323 331 217 45 Khaya nyassica

0--10 214 51 3 205 268 168 45 Pinus elliottii

0--10 236 38 3 227 278 204 45 Terminalia ivorensis

0--10 362 279 3 323 658 105 45 Anthocephalus cadamba

0--10 300 116 4 266 467 202 45 Swietenia macrophylla

0--10 274 151 3 192 448 182 45 Hibiscus elatus

0--10 550 245 3 502 816 333 45 Eucalyptus saligna

0--10 237 26 3 250 253 207 45 Pinus caribaea var. hondurensis

0--10 239 48 3 214 294 208 45 Hernandia sonora

0--10 203 121 4 160 379 114 45 Eucalyptus pathentinervis
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

456

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Calcium-total (mg/g) 0--10 0.26 0.06 4 0.26 0.33 0.18 45 Khaya nyassica

0--10 0.20 0.15 4 0.20 0.39 0.03 45 Pinus elliottii

0--10 0.19 0.03 4 0.21 0.21 0.15 45 Terminalia ivorensis

0--10 0.49 0.17 4 0.45 0.71 0.33 45 Anthocephalus cadamba

0--10 0.52 0.32 4 0.45 0.95 0.25 45 Swietenia macrophylla

0--10 0.44 0.25 4 0.54 0.61 0.07 45 Hibiscus elatus

0--10 1.17 0.38 4 1.09 0.85 0.63 45 Eucalyptus saligna

0--10 0.49 0.04 4 0.50 0.53 0.43 45 Pinus caribaea var. hondurensis

0--10 0.63 0.11 4 0.60 0.77 0.55 45 Hernandia sonora

0--10 0.81 0.37 4 0.87 1.14 0.37 45 Eucalyptus pathentinervis

Clay (%) 0--10 32 1 45 Khaya nyassica

0--10 18 1 45 Pinus elliottii

0--10 34 1 45 Terminalia ivorensis

0--10 34 1 45 Anthocephalus cadamba

0--10 26 1 45 Swietenia macrophylla

0--10 22 1 45 Hibiscus elatus

0--10 25 1 45 Eucalyptus saligna

0--10 27 1 45 Pinus caribaea var. hondurensis

0--10 25 1 45 Hernandia sonora

0--10 24 1 45 Eucalyptus pathentinervis

Iron (mg/g) 0--10 4.93 1.57 4 4.79 6.75 3.39 45 Khaya nyassica

0--10 2.93 0.91 3 2.49 3.98 2.32 45 Pinus elliottii

0--10 2.56 0.58 3 2.50 3.17 2.01 45 Terminalia ivorensis

0--10 1.94 0.51 3 2.05 2.39 1.39 45 Anthocephalus cadamba

0--10 3.95 0.43 4 3.90 4.45 3.56 45 Swietenia macrophylla

0--10 2.83 0.74 3 3.15 3.36 1.98 45 Hibiscus elatus
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

457

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--10 1.78 0.38 3 1.76 2.16 1.41 45 Eucalyptus saligna

0--10 1.84 0.86 3 1.40 2.82 1.29 45 Pinus caribaea var. hondurensis

0--10 8.61 1.24 3 8.40 9.94 7.48 45 Hernandia sonora

0--10 2.52 0.74 4 2.23 3.62 2.00 45 Eucalyptus pathentinervis

Iron-total (mg/g) 0--10 64.29 3.91 4 63.49 69.73 60.46 45 Khaya nyassica

0--10 77.14 2.73 4 77.92 79.40 73.30 45 Pinus elliottii

0--10 74.73 5.30 4 76.45 78.99 67.04 45 Terminalia ivorensis

0--10 55.24 4.66 4 53.07 62.21 52.61 45 Anthocephalus cadamba

0--10 60.85 4.67 4 60.11 66.49 56.67 45 Swietenia macrophylla

0--10 62.30 5.12 4 63.77 66.71 54.96 45 Hibiscus elatus

0--10 64.60 8.02 4 68.03 69.63 52.72 45 Eucalyptus saligna

0--10 79.38 8.75 4 77.84 90.79 71.05 45 Pinus caribaea var. hondurensis

0--10 56.08 2.20 4 56.24 58.57 53.28 45 Hernandia sonora

0--10 64.42 3.06 4 64.78 67.60 60.53 45 Eucalyptus pathentinervis

Magnesium (mg/kg) 0--10 183 26 4 192 204 195 45 Khaya nyassica

0--10 24 51 3 99 182 90 45 Pinus elliottii

0--10 209 15 3 202 226 199 45 Terminalia ivorensis

0--10 262 81 3 286 328 172 45 Anthocephalus cadamba

0--10 177 65 4 149 274 137 45 Swietenia macrophylla

0--10 211 69 3 175 291 168 45 Hibiscus elatus

0--10 415 212 3 489 580 175 45 Eucalyptus saligna

0--10 220 49 3 231 263 167 45 Pinus caribaea var. hondurensis

0--10 277 77 3 234 366 230 45 Hernandia sonora

0--10 226 87 4 193 354 163 45 Eucalyptus pathentinervis

Magnesium-total (mg/g) 0--10 0.89 0.05 4 0.90 0.94 0.83 45 Khaya nyassica

0--10 0.85 0.10 4 0.87 0.96 0.72 45 Pinus elliottii
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

458

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--10 1.34 0.24 4 1.24 1.69 1.18 45 Terminalia ivorensis

0--10 0.87 0.45 4 0.68 1.55 0.59 45 Anthocephalus cadamba

0--10 0.86 0.18 4 0.85 1.05 0.68 45 Swietenia macrophylla

0--10 0.94 0.02 4 0.94 0.97 0.93 45 Hibiscus elatus

0--10 2.01 1.24 4 1.49 3.85 1.23 45 Eucalyptus saligna

0--10 0.75 0.32 4 0.74 1.08 0.45 45 Pinus caribaea var. hondurensis

0--10 0.83 0.09 4 0.84 0.94 0.72 45 Hernandia sonora

0--10 0.77 0.17 4 0.78 0.92 0.60 45 Eucalyptus pathentinervis

Manganese (mg/kg) 0--10 26 9 4 22 39 20 45 Khaya nyassica

0--10 16 3 3 16 19 13 45 Pinus elliottii

0--10 115 21 3 111 138 96 45 Terminalia ivorensis

0--10 83 18 3 76 103 70 45 Anthocephalus cadamba

0--10 38 12 4 32 56 31 45 Swietenia macrophylla

0--10 23 10 3 24 32 13 45 Hibiscus elatus

0--10 74 24 3 60 102 60 45 Eucalyptus saligna

0--10 13 7 3 11 21 8 45 Pinus caribaea var. hondurensis

0--10 8 3 3 8 11 5 45 Hernandia sonora

0--10 17 7 4 1 22 7 45 Eucalyptus pathentinervis

Manganese-total (mg/g) 0--10 0.04 0.02 4 0.03 0.07 0.01 45 Khaya nyassica

0--10 0.07 0.01 4 0.08 0.08 0.07 45 Pinus elliottii

0--10 0.41 0.13 4 0.35 0.61 0.33 45 Terminalia ivorensis

0--10 0.25 0.10 4 0.25 0.35 0.17 45 Anthocephalus cadamba

0--10 0.21 0.08 4 0.18 0.34 0.15 45 Swietenia macrophylla

0--10 0.08 0.01 4 0.08 0.09 0.07 45 Hibiscus elatus

0--10 0.28 0.09 4 0.26 0.39 0.21 45 Eucalyptus saligna

0--10 0.07 0.03 4 0.06 0.13 0.05 45 Pinus caribaea var. hondurensis
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

459

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--10 0.05 0.02 4 0.06 0.07 0.03 45 Hernandia sonora

0--10 0.25 0.25 4 0.16 0.62 0.09 45 Eucalyptus pathentinervis

Nitrogen (%) 0--10 0.42 0.07 3 0.42 0.48 0.35 45 Khaya nyassica

0--10 0.26 0.04 3 0.27 3.00 0.22 45 Pinus elliottii

0--10 0.51 0.02 3 0.52 0.52 0.48 45 Terminalia ivorensis

0--10 0.43 0.03 3 0.44 0.45 0.40 45 Anthocephalus cadamba

0--10 0.50 0.03 3 0.49 0.53 0.47 45 Swietenia macrophylla

0--10 0.37 0.05 3 0.37 0.42 0.33 45 Hibiscus elatus

0--10 0.38 0.06 3 0.35 0.45 0.33 45 Eucalyptus saligna

0--10 0.23 0.02 3 0.24 0.24 0.21 45 Pinus caribaea var. hondurensis

0--10 0.47 0.04 3 0.05 0.51 0.43 45 Hernandia sonora

0--10 0.42 0.02 3 0.42 0.43 0.40 45 Eucalyptus pathentinervis

Organic matter (%) 0--10 8.93 1.04 3 8.90 9.98 7.90 45 Khaya nyassica

0--10 6.45 1.12 3 6.38 7.60 5.36 45 Pinus elliottii

0--10 9.99 1.06 3 10.52 10.68 8.77 45 Terminalia ivorensis

0--10 8.70 0.09 3 8.73 8.78 8.60 45 Anthocephalus cadamba

0--10 8.84 0.87 3 8.76 9.75 8.02 45 Swietenia macrophylla

0--10 8.44 0.95 3 8.76 9.19 7.37 45 Hibiscus elatus

0--10 9.80 2.10 3 10.89 11.14 7.38 45 Eucalyptus saligna

0--10 5.61 0.42 3 5.37 6.10 5.36 45 Pinus caribaea var. hondurensis

0--10 9.83 0.16 3 9.88 9.96 9.65 45 Hernandia sonora

0--10 9.69 1.39 3 9.86 10.98 8.22 45 Eucalyptus pathentinervis

pH (H2O) 0--10 4.52 0.07 3 4.55 4.56 4.44 45 Khaya nyassica

0--10 4.64 0.05 3 4.63 4.69 4.60 45 Pinus elliottii

0--10 4.64 0.03 3 4.63 4.67 4.61 45 Terminalia ivorensis

0--10 4.66 0.11 3 4.63 4.78 4.57 45 Anthocephalus cadamba
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

460

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--10 4.42 0.11 3 4.42 4.52 4.31 45 Swietenia macrophylla

0--10 4.73 0.19 3 4.64 4.95 4.61 45 Hibiscus elatus

0--10 4.75 0.06 3 4.75 4.80 4.69 45 Eucalyptus saligna

0--10 4.74 0.05 3 4.75 4.78 4.69 45 Pinus caribaea var. hondurensis

0--10 4.83 0.03 3 4.83 4.85 4.80 45 Hernandia sonora

0--10 4.83 0.27 3 4.78 5.12 4.59 45 Eucalyptus pathentinervis

Phosphorus (mg/kg) 0--10 37 5 4 40 40 29 45 Khaya nyassica

0--10 21 8 3 24 26 12 45 Pinus elliottii

0--10 31 3 3 32 33 28 45 Terminalia ivorensis

0--10 25 6 3 28 29 19 45 Anthocephalus cadamba

0--10 28 1 4 28 30 27 45 Swietenia macrophylla

0--10 25 12 3 31 33 12 45 Hibiscus elatus

0--10 26 8 3 30 30 17 45 Eucalyptus saligna

0--10 11 7 3 14 15 3 45 Pinus caribaea var. hondurensis

0--10 36 6 3 38 40 29 45 Hernandia sonora

0--10 14 2 4 15 16 12 45 Eucalyptus pathentinervis

Phosphorus-total (mg/g) 0--10 0.31 0.03 4 0.30 0.35 0.28 45 Khaya nyassica

0--10 0.32 0.03 4 0.33 0.36 0.29 45 Pinus elliottii

0--10 0.39 0.04 4 0.38 0.45 0.37 45 Terminalia ivorensis

0--10 0.51 0.06 4 0.49 0.59 0.46 45 Anthocephalus cadamba

0--10 0.60 0.10 4 0.61 0.69 0.49 45 Swietenia macrophylla

0--10 0.54 0.04 4 0.55 0.58 0.49 45 Hibiscus elatus

0--10 0.59 0.07 4 0.58 0.67 0.52 45 Eucalyptus saligna

0--10 0.51 0.21 4 0.41 0.82 0.39 45 Pinus caribaea var. hondurensis

0--10 0.39 0.02 4 0.38 0.41 0.37 45 Hernandia sonora

0--10 0.35 0.02 4 0.35 0.37 0.33 45 Eucalyptus pathentinervis
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

461

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Potassium (mg/kg) 0--10 277 23 4 288 290 243 45 Khaya nyassica

0--10 208 45 3 188 259 176 45 Pinus elliottii

0--10 269 19 3 261 291 255 45 Terminalia ivorensis

0--10 254 53 3 267 300 196 45 Anthocephalus cadamba

0--10 269 38 4 256 325 240 45 Swietenia macrophylla

0--10 242 46 3 220 295 210 45 Hibiscus elatus

0--10 225 65 3 250 274 152 45 Eucalyptus saligna

0--10 158 47 3 152 208 115 45 Pinus caribaea var. hondurensis

0--10 270 35 3 290 290 230 45 Hernandia sonora

0--10 186 20 4 194 200 157 45 Eucalyptus pathentinervis

Potassium-total (mg/g) 0--10 2.10 0.05 4 2.11 2.14 2.03 45 Khaya nyassica

0--10 2.39 0.12 4 2.42 2.51 2.23 45 Pinus elliottii

0--10 2.35 0.11 4 2.31 2.51 2.28 45 Terminalia ivorensis

0--10 1.45 0.15 4 1.39 1.67 1.37 45 Anthocephalus cadamba

0--10 1.93 0.03 4 1.94 1.95 1.89 45 Swietenia macrophylla

0--10 2.46 0.09 4 2.47 2.56 2.35 45 Hibiscus elatus

0--10 2.64 0.04 4 2.63 2.69 2.59 45 Eucalyptus saligna

0--10 2.54 0.73 4 2.39 3.47 1.93 45 Pinus caribaea var. hondurensis

0--10 1.44 0.07 4 1.46 1.50 1.33 45 Hernandia sonora

0--10 1.63 0.08 4 1.61 1.75 1.57 45 Eucalyptus pathentinervis

Sand (%) 0--10 15 1 45 Khaya nyassica

0--10 12 1 45 Pinus elliottii

0--10 21 1 45 Terminalia ivorensis

0--10 11 1 45 Anthocephalus cadamba

0--10 19 1 45 Swietenia macrophylla

0--10 10 1 45 Hibiscus elatus
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

462

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--10 23 1 45 Eucalyptus saligna

0--10 5 1 45 Pinus caribaea var. hondurensis

0--10 29 1 45 Hernandia sonora

0--10 13 1 45 Eucalyptus pathentinervis

Silt (%) 0--10 53 1 45 Khaya nyassica

0--10 70 1 45 Pinus elliottii

0--10 45 1 45 Terminalia ivorensis

0--10 55 1 45 Anthocephalus cadamba

0--10 55 1 45 Swietenia macrophylla

0--10 68 1 45 Hibiscus elatus

0--10 52 1 45 Eucalyptus saligna

0--10 68 1 45 Pinus caribaea var. hondurensis

0--10 46 1 45 Hernandia sonora

0--10 63 1 45 Eucalyptus pathentinervis

Sodium (mg/kg) 0--10 55 14 4 56 71 37 45 Khaya nyassica

0--10 51 4 3 49 56 48 45 Pinus elliottii

0--10 63 6 3 63 69 58 45 Terminalia ivorensis

0--10 75 28 3 64 107 55 45 Anthocephalus cadamba

0--10 83 25 4 91 104 47 45 Swietenia macrophylla

0--10 57 12 3 59 67 44 45 Hibiscus elatus

0--10 134 52 3 156 171 74 45 Eucalyptus saligna

0--10 8 13 3 93 96 73 45 Pinus caribaea var. hondurensis

0--10 128 43 3 152 154 78 45 Hernandia sonora

0--10 67 26 4 67 94 42 45 Eucalyptus pathentinervis

Tuffaceous sandstone Ultisols Mfs Aluminum saturation (%) 0--15 7.10 14.93 28 2.33 75.27 0.57 46

15--30 21.77 25.86 25 8.43 80.12 0.42 46
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

463

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Aluminum (cmol/kg-l) 0--15 1.00 1.37 28 0.57 7.03 0.18 46

15--30 2.12 2.39 25 0.75 7.08 0.11 46

Bulk density (g/cc) 0--15 0.52 0.14 28 0.55 0.73 0.10 46

15--30 0.75 0.19 25 0.80 0.97 0.27 46

Calcium (cmol/kg-l) 0--15 16.24 9.52 28 14.18 32.92 0.45 46

15--30 5.98 4.27 25 4.29 14.80 0.07 46

Calcium (mg/g) 0--15 3.25 1.90 28 2.84 6.58 0.09 46

15--30 1.20 0.85 25 0.86 2.96 0.01 46

Effective cation exchange capacity 0--15 26.07 13.28 28 21.62 52.78 9.34 46

{Ca+K+Mg(cmol/kg-1) 15--30 13.39 5.16 25 10.85 26.42 8.38 46

Iron (mg/g) 0--15 0.20 0.12 28 0.19 0.41 0.04 46

15--30 0.16 0.08 25 0.14 0.34 0.04 46

Loss on ignition (%) 0--15 25.87 5.41 28 24.85 40.27 16.82 46

15--30 17.80 1.82 25 17.59 21.59 14.79 46

Magnesium (cmol/kg-l) 0--15 8.02 4.27 28 6.83 17.28 1.58 46

15--30 4.62 2.45 25 4.29 10.26 1.38 46

Magnesium (mg/g) 0--15 0.96 0.51 28 0.82 2.07 0.19 46

15--30 0.56 0.30 25 0.52 1.23 0.17 46

Manganese (mg/g) 0--15 0.28 0.14 28 0.28 0.63 0.05 46

15--30 0.12 0.13 25 0.10 0.64 0.01 46

Nitrogen (%) 0--15 0.58 0.28 28 0.51 1.30 0.15 46

15--30 0.19 0.08 25 0.18 0.33 0.04 46

Organic matter (%) 0--15 10.56 4.94 28 9.76 23.72 2.53 46

15--30 3.16 1.48 25 2.99 6.83 0.77 46

pH (H2O) 0--15 5.42 0.56 28 5.35 6.47 4.30 46

15--30 5.14 0.47 25 5.05 5.97 4.45 46
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

464

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

pH (KCl) 0--15 4.74 0.53 28 4.74 5.82 3.48 46

15--30 4.29 0.47 25 4.98 4.98 3.59 46

Phosphorus (mg/kg) 0--15 15 7 28 14 37 7 46

15--30 5 2 25 5 11 2 46

Potassium (cmol/kg-l) 0--15 0.37 0.18 28 0.33 0.84 0.08 46

15--30 0.25 0.29 25 0.13 1.30 0.05 46

Potassium (mg/g) 0--15 0.14 0.07 28 0.13 0.32 0.03 46

15--30 0.09 0.11 25 0.05 0.50 0.02 46

Sodium (cmol/kg-l) 0--15 0.45 0.26 28 0.38 0.98 0.08 46

15--30 0.42 0.26 25 0.41 0.96 0.07 46

Sodium (mg/g) 0--15 0.10 0.06 28 0.09 0.23 0.02 46

15--30 0.10 0.06 25 0.09 0.22 0.02 46

Plutonic rocks Mollisols Wfs Aluminum (cmol/kg-l) 0--15 14.86 1.97 18 14.64 19.42 12.09 47

15--30 14.37 1.41 21 14.05 16.61 10.81 47

Bulk density (g/cc) 0--15 0.74 0.11 18 0.74 0.93 0.52 47

15--30 1.12 0.16 18 1.14 1.42 0.75 47

C/N 0--15 12 1 18 12 13 11 47

15--30 12 0 21 12 13 11 47

Calcium (cmol/kg-l) 0--15 0.55 0.31 18 0.50 1.65 0.18 47

15--30 0.20 0.10 21 0.17 0.50 0.10 47

Calcium (mg/g) 0--15 0.11 0.06 18 0.10 0.33 0.04 47

15--30 0.04 0.02 21 0.03 0.10 0.02 47

Carbon (%) 0--15 4.18 0.81 18 4.22 5.85 2.81 47

15--30 2.18 0.67 21 2.00 3.42 1.41 47

Effective cation exchange capacity 0--15 16.41 1.90 18 16.04 20.68 13.59 47

{Ca+K+Mg(cmol/kg-1) 15--30 15.09 1.39 21 14.68 17.18 11.46 47
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

465

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Iron (mg/g) 0--15 0.73 0.30 18 0.63 1.59 0.40 47

15--30 0.51 0.20 21 0.40 0.89 0.28 47

Loss on ignition (%) 0--15 15.58 3.20 18 16.74 21.10 10.30 47

15--30 14.54 9.27 21 12.97 53.09 8.28 47

Magnesium (cmol/kg-l) 0--15 0.60 0.17 18 0.59 1.01 0.33 47

15--30 0.27 0.07 21 0.25 0.43 0.18 47

Magnesium (mg/g) 0--15 0.07 0.02 18 0.07 0.12 0.04 47

15--30 0.03 0.01 21 0.30 0.05 0.02 47

Manganese (mg/g) 0--15 0.12 0.07 18 0.11 0.25 0.02 47

15--30 0.06 0.05 21 0.54 0.19 0.01 47

Nitrogen (%) 0--15 0.34 0.06 18 0.33 0.48 0.24 47

15--30 0.18 0.05 21 0.17 0.28 0.12 47

pH (H2O) 0--15 4.02 0.25 18 4.00 4.46 3.45 47

15--30 4.23 0.21 21 4.22 4.58 3.66 47

pH (KCl) 0--15 3.45 0.18 18 3.49 3.67 3.13 47

15--30 3.59 0.09 21 3.60 3.80 3.44 47

Phosphorus (mg/kg) 0--15 28 4 18 29 34 21 47

15--30 12 4 21 11 19 8 47

Potassium (cmol/kg-l) 0--15 0.29 0.06 18 0.27 0.42 0.20 47

15--30 0.16 0.05 21 0.16 0.25 0.10 47

Potassium (mg/g) 0--15 0.11 0.02 18 0.10 0.16 0.07 47

15--30 0.06 0.02 21 0.06 0.09 0.04 47

Sodium (cmol/kg-l) 0--15 0.12 0.03 18 0.11 0.18 0.07 47

15--30 0.09 0.04 21 0.08 0.24 0.06 47

Sodium (mg/kg) 0--15 27 7 18 26 4 15 47

15--30 21 9 21 18 54 14 47
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

466

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Tuffaceous sandstone Ultisols Wfs Aluminum (cmol/kg-l) 0--15 0.40 0.35 69 0.22 1.33 0.07 48 Mahogany plantation-fumigated

0--15 0.63 0.57 50 0.40 1.81 0.02 48 Mahogany plantation

0--15 2.18 1.14 24 2.43 4.03 0.25 48 Pine plantation-fumigated

0--15 2.69 1.29 37 3.09 4.56 0.09 48 Pine plantation

0--15 0.94 0.57 27 0.82 2.21 0.19 48 Guzman secondary forest-fumigated

0--15 1.25 0.67 34 1.15 2.79 0.09 48 Guzman secondary forest

0--15 1.73 0.85 69 1.66 3.84 0.32 48 Tabonuco secondary forest-fumigated

0--15 2.52 1.09 54 2.47 4.86 0.20 48 Tabonuco secondary forest

0--15 0.16 0.04 23 0.16 0.27 0.07 48 Toa Baja

0--15 0.06 0.02 21 0.06 0.10 0.02 48 Toa Baja-fumigated

Calcium (cmol/kg-l) 0--15 6.42 4.56 69 5.74 38.59 2.01 48 Mahogany plantation-fumigated

0--15 7.71 87.28 50 5.58 40.96 2.29 48 Mahogany plantation

0--15 2.92 0.87 24 3.04 4.95 1.63 48 Pine plantation-fumigated

0--15 2.86 1.07 37 2.69 5.62 1.01 48 Pine plantation

0--15 3.67 1.61 27 3.48 8.85 1.47 48 Guzman secondary forest-fumigated

0--15 5.00 6.62 34 3.62 41.16 1.39 48 Guzman secondary forest

0--15 2.65 1.33 69 2.59 6.38 0.55 48 Tabonuco secondary forest-fumigated

0--15 3.06 1.35 54 3.09 7.05 0.74 48 Tabonuco secondary forest

0--15 7.54 1.71 23 7.45 12.06 4.92 48 Toa Baja

0--15 7.18 1.59 21 7.56 9.89 4.58 48 Toa Baja-fumigated

Calcium (mg/g) 0--15 1.28 0.91 69 1.15 7.72 0.40 48 Mahogany plantation-fumigated

0--15 1.54 1.46 50 1.12 8.19 0.46 48 Mahogany plantation

0--15 0.59 0.17 24 0.61 0.99 0.33 48 Pine plantation-fumigated

0--15 0.57 0.21 37 0.54 1.12 0.20 48 Pine plantation

0--15 0.73 0.32 27 0.70 1.77 0.29 48 Guzman secondary forest-fumigated

0--15 1.00 1.33 34 0.72 8.23 0.28 48 Guzman secondary forest
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

467

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--15 0.53 0.27 69 0.52 1.28 0.11 48 Tabonuco secondary forest-fumigated

0--15 0.61 0.27 54 0.62 1.41 0.15 48 Tabonuco secondary forest

0--15 1.51 0.34 23 1.49 2.41 0.98 48 Toa Baja

0--15 1.44 0.32 21 1.51 1.98 0.92 48 Toa Baja-fumigated

0--15 1.95 0.26 23 1.93 2.48 1.45 48 Toa Baja

Effective cation exchange capacity 0--15 12.12 5.09 69 11.41 45.55 5.39 48 Mahogany plantation-fumigated

{Ca+K+Mg(cmol/kg-1) 0--15 13.70 7.34 50 11.71 46.68 6.63 48 Mahogany plantation

0--15 8.01 1.14 24 7.84 10.29 6.34 48 Pine plantation-fumigated

0--15 8.20 1.34 37 8.25 10.97 5.53 48 Pine plantation

0--15 8.95 2.57 27 8.44 17.11 5.83 48 Guzman secondary forest-fumigated

0--15 10.49 6.81 34 8.78 46.88 6.61 48 Guzman secondary forest

0--15 6.83 1.82 69 6.90 10.90 3.22 48 Tabonuco secondary forest-fumigated

0--15 8.04 1.52 54 8.09 11.29 4.15 48 Tabonuco secondary forest

0--15 10.27 2.16 23 10.28 16.18 6.57 48 Toa Baja

Iron (mg/g) 0--15 0.74 0.50 69 0.54 2.01 0.14 48 Mahogany plantation-fumigated

0--15 0.68 0.42 50 0.52 1.70 0.19 48 Mahogany plantation

0--15 0.49 0.37 24 0.40 1.53 0.11 48 Pine plantation-fumigated

0--15 0.54 0.36 37 0.43 1.62 0.12 48 Pine plantation

0--15 0.63 0.31 27 0.54 1.44 0.27 48 Guzman secondary forest-fumigated

0--15 0.72 0.33 34 0.75 1.55 0.28 48 Guzman secondary forest

0--15 2.70 1.54 69 2.46 8.04 0.85 48 Tabonuco secondary forest-fumigated

0--15 2.77 1.65 54 2.77 7.77 0.23 48 Tabonuco secondary forest

0--15 4.41 1.13 23 4.34 6.90 2.53 48 Toa Baja

Magnesium (cmol/kg-l) 0--15 5.09 1.19 69 5.31 6.83 1.37 48 Mahogany plantation-fumigated

0--15 5.18 0.90 50 5.15 6.73 3.09 48 Mahogany plantation

0--15 2.78 0.65 24 2.58 4.24 1.63 48 Pine plantation-fumigated
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

468

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--15 2.52 0.60 37 2.38 3.93 1.55 48 Pine plantation

0--15 4.03 1.12 27 3.92 7.22 2.42 48 Guzman secondary forest-fumigated

0--15 3.94 0.92 34 3.83 6.47 2.43 48 Guzman secondary forest

0--15 2.20 0.76 69 2.05 4.15 1.00 48 Tabonuco secondary forest-fumigated

0--15 2.26 0.74 54 2.19 1.08 0.87 48 Tabonuco secondary forest

0--15 1.59 0.65 23 1.35 2.81 0.77 48 Toa Baja

0--15 1.29 0.55 21 1.12 2.65 0.55 48 Toa Baja-fumigated

Magnesium (mg/g) 0--15 0.61 0.14 69 0.64 0.82 0.16 48 Mahogany plantation-fumigated

0--15 0.62 0.11 50 0.62 0.81 0.37 48 Mahogany plantation

0--15 0.33 77.00 24 0.31 0.51 0.20 48 Pine plantation-fumigated

0--15 0.30 0.07 37 0.29 0.47 0.19 48 Pine plantation

0--15 0.48 0.14 27 0.47 0.87 0.29 48 Guzman secondary forest-fumigated

0--15 0.47 0.11 34 0.46 0.78 0.29 48 Guzman secondary forest

0--15 0.26 0.09 69 0.25 0.50 0.12 48 Tabonuco secondary forest-fumigated

0--15 0.27 0.09 54 0.26 0.59 0.13 48 Tabonuco secondary forest

0--15 0.19 0.08 23 0.16 0.34 0.09 48 Toa Baja

0--15 0.16 0.07 21 0.13 0.32 0.07 48 Toa Baja-fumigated

Manganese (mg/kg) 0--15 17 5 23 15 31 12 48 Toa Baja

0--15 58 66 69 38 419 5 48 Mahogany plantation-fumigated

0--15 42 27 50 34 120 8 48 Mahogany plantation

0--15 130 101 24 86 344 20 48 Pine plantation-fumigated

0--15 67 43 37 72 195 14 48 Pine plantation

0--15 87 37 27 95 155 19 48 Guzman secondary forest-fumigated

0--15 61 26 34 53 123 12 48 Guzman secondary forest

0--15 111 90 69 62 445 19 48 Tabonuco secondary forest-fumigated

0--15 92 67 54 56 286 15 48 Tabonuco secondary forest
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

469

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--15 6 1 21 6 9 4 48 Toa Baja-fumigated

Nitrogen (%) 0--15 0.14 0.03 23 0.13 0.19 0.08 48 Toa Baja

pH (H2O) 0--15 7.21 0.42 23 7.32 7.72 5.96 48 Toa Baja

pH (KCl) 0--15 6.58 0.48 23 6.81 7.11 5.30 48 Toa Baja

Phosphorus (mg/kg) 0--15 9 2 23 8 13 7 48 Toa Baja

0--15 9 3 69 9 20 5 48 Mahogany plantation-fumigated

0--15 5 2 50 5 12 1 48 Mahogany plantation

0--15 5 3 24 4 12 1 48 Pine plantation-fumigated

0--15 4 2 37 4 9 1 48 Pine plantation

0--15 9 4 27 7 17 3 48 Guzman secondary forest-fumigated

0--15 6 3 34 7 11 2 48 Guzman secondary forest

0--15 15 5 69 14 32 7 48 Tabonuco secondary forest-fumigated

0--15 7 2 54 7 15 3 48 Tabonuco secondary forest

0--15 12 3 21 12 17 8 48 Toa Baja-fumigated

Potassium (cmol/kg-l) 0--15 0.57 0.23 23 0.52 1.16 0.32 48 Toa Baja

0--15 0.21 0.09 69 0.19 0.53 0.10 48 Mahogany plantation-fumigated

0--15 0.17 0.08 50 0.16 0.37 0.07 48 Mahogany plantation

0--15 0.13 0.00 24 0.13 0.22 0.05 48 Pine plantation-fumigated

0--15 0.12 0.08 37 0.09 0.31 0.04 48 Pine plantation

0--15 0.32 0.17 27 0.29 0.84 0.14 48 Guzman secondary forest-fumigated

0--15 0.29 0.15 34 0.27 0.79 0.10 48 Guzman secondary forest

0--15 0.25 0.07 69 0.24 0.44 0.14 48 Tabonuco secondary forest-fumigated

0--15 0.20 0.06 54 0.19 0.37 0.10 48 Tabonuco secondary forest

0--15 0.21 0.06 21 0.20 0.37 0.11 48 Toa Baja-fumigated

Potassium (mg/g) 0--15 0.08 0.04 69 0.08 0.21 0.04 48 Mahogany plantation-fumigated

0--15 0.07 0.03 50 0.06 0.14 0.03 48 Mahogany plantation
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

470

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--15 0.05 0.02 24 0.05 0.09 0.02 48 Pine plantation-fumigated

0--15 0.05 0.03 37 0.04 0.12 0.01 48 Pine plantation

0--15 0.12 0.07 27 0.11 0.33 0.05 48 Guzman secondary forest-fumigated

0--15 0.12 0.06 34 0.10 0.31 0.04 48 Guzman secondary forest

0--15 0.10 0.03 69 0.09 0.17 0.05 48 Tabonuco secondary forest-fumigated

0--15 0.08 0.02 54 0.08 0.15 0.04 48 Tabonuco secondary forest

0--15 0.22 0.09 23 0.20 0.44 0.12 48 Toa Baja

0--15 0.08 0.03 21 0.08 0.14 0.04 48 Toa Baja-fumigated

Sodium (cmol/kg-l) 0--15 0.42 0.17 23 0.39 0.85 0.18 48 Toa Baja

Sodium (mg/g) 0--15 0.10 0.04 23 0.09 0.20 0.04 48 Toa Baja

Tuffaceous sandstone Ultisols Lmrf Aluminum (cmol/kg-l) 0--10 3.80 0.44 6 3.68 4.45 3.36 49 Cloud forest-dry soil

0--10 2.35 0.53 6 2.41 2.92 1.67 49 Colorado forest-dry forest

0--10 6.62 1.38 6 6.37 8.70 4.84 49 Tabonuco forest-dry forest

0--10 0.51 0.28 5 0.40 0.94 0.28 49 Cloud forest-wet soil

0--10 1.12 0.32 6 1.32 1.34 0.62 49 Colorado forest-wet soil

0--10 2.50 0.54 6 2.65 3.18 1.72 49 Tabaonuco forest-wet soil

10--35 3.36 0.40 5 3.34 3.88 2.84 49 Cloud forest-dry soil

10--35 2.15 0.51 5 2.46 2.57 1.47 49 Colorado forest-dry forest

10--35 0.75 1.08 5 4.84 5.95 3.42 49 Tabonuco forest-dry forest

10--35 0.72 0.12 5 0.72 0.90 0.60 49 Cloud forest-wet soil

10--35 1.02 0.29 5 0.92 1.46 0.78 49 Colorado forest-wet soil

10--35 1.71 0.71 5 1.79 2.57 0.85 49 Tabaonuco forest-wet soil

Calcium (mg/kg) 0--10 365 87 6 380 528 293 49 Cloud forest-dry soil

0--10 272 114 6 257 411 139 49 Colorado forest-dry forest

0--10 99 29 6 101 130 62 49 Tabonuco forest-dry forest

0--10 90 41 6 82 165 63 49 Cloud forest-wet soil
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

471

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--10 150 76 6 148 234 73 49 Colorado forest-wet soil

0--10 28 12 6 26 470 14 49 Tabaonuco forest-wet soil

10--35 116 27 5 131 140 85 49 Cloud forest-dry soil

10--35 91 45 5 73 170 61 49 Colorado forest-dry forest

10--35 14 5 5 13 22 10 49 Tabonuco forest-dry forest

10--35 55 22 5 44 81 36 49 Cloud forest-wet soil

10--35 44 35 5 29 103 16 49 Colorado forest-wet soil

10--35 7 5 5 5 13 3 49 Tabaonuco forest-wet soil

Iron (mg/g) 0--10 0.41 0.24 6 2.38 2.86 2.20 49 Cloud forest-dry soil

0--10 1.77 0.21 6 1.77 2.15 1.50 49 Colorado forest-dry forest

0--10 3.35 2.59 6 3.01 7.21 1.39 49 Tabonuco forest-dry forest

0--10 2.53 0.61 6 2.27 3.75 2.19 49 Cloud forest-wet soil

0--10 1.51 0.34 6 1.50 1.88 0.93 49 Colorado forest-wet soil

0--10 2.30 1.09 6 1.97 3.70 1.31 49 Tabaonuco forest-wet soil

10--35 2.36 0.15 5 2.29 2.60 2.26 49 Cloud forest-dry soil

10--35 0.13 0.10 5 1.23 1.36 1.12 49 Colorado forest-dry forest

10--35 0.99 0.52 4 0.87 1.70 0.54 49 Tabonuco forest-dry forest

10--35 2.69 0.56 5 2.52 3.51 2.05 49 Cloud forest-wet soil

10--35 0.84 0.09 5 0.82 0.97 0.71 49 Colorado forest-wet soil

10--35 1.32 0.75 5 1.17 2.45 0.52 49 Tabaonuco forest-wet soil

Loss on ignition (%) 0--10 35.36 2.08 6 35.09 37.83 33.37 49 Cloud forest-dry soil

0--10 13.95 1.36 6 13.87 15.63 12.16 49 Colorado forest-dry forest

0--10 21.54 2.95 6 21.89 26.37 18.09 49 Tabonuco forest-dry forest

10--35 24.65 3.60 5 24.53 28.29 19.60 49 Cloud forest-dry soil

10--35 10.98 1.40 5 11.28 12.38 8.95 49 Colorado forest-dry forest

10--35 14.79 1.30 5 14.24 16.30 13.63 49 Tabonuco forest-dry forest
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

472

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Magnesium (mg/g) 0--10 0.15 0.05 6 0.14 0.21 0.12 49 Cloud forest-dry soil

0--10 0.11 0.02 6 0.11 0.13 0.08 49 Colorado forest-dry forest

0--10 0.15 0.02 6 0.15 0.18 0.11 49 Tabonuco forest-dry forest

0--10 0.04 0.02 6 0.03 0.06 0.03 49 Cloud forest-wet soil

0--10 0.06 0.01 6 0.06 0.07 0.03 49 Colorado forest-wet soil

0--10 0.05 0.02 6 0.05 0.07 0.03 49 Tabaonuco forest-wet soil

10--35 0.06 0.02 5 0.07 0.08 0.04 49 Cloud forest-dry soil

10--35 0.08 0.02 5 0.07 0.12 0.06 49 Colorado forest-dry forest

10--35 0.06 0.02 5 0.05 0.10 0.05 49 Tabonuco forest-dry forest

10--35 0.03 0.01 5 0.03 0.04 0.02 49 Cloud forest-wet soil

10--35 0.04 0.02 5 0.04 0.06 0.02 49 Colorado forest-wet soil

10--35 0.03 0.01 5 0.02 0.04 0.02 49 Tabaonuco forest-wet soil

Manganese (mg/kg) 0--10 11 2 6 11 14 8 49 Cloud forest-dry soil

0--10 20 12 6 14 35 11 49 Colorado forest-dry forest

0--10 12 5 6 14 16 5 49 Tabonuco forest-dry forest

0--10 0.97 0.47 6 0.98 1.61 0.24 49 Cloud forest-wet soil

0--10 10 10 6 6 26 1 49 Colorado forest-wet soil

0--10 3 2 6 3 5 2 49 Tabaonuco forest-wet soil

10--35 9 9 5 6 25 4 49 Cloud forest-dry soil

10--35 15 12 5 10 37 8 49 Colorado forest-dry forest

10--35 2 1 4 2 3 1 49 Tabonuco forest-dry forest

10--35 4.09 6.77 4 0.99 14.21 0.16 49 Cloud forest-wet soil

10--35 6 5 5 4 15 3 49 Colorado forest-wet soil

10--35 0.79 0.60 5 0.69 1.69 0.08 49 Tabaonuco forest-wet soil

Nitrogen (%) 0--10 0.60 0.05 6 0.60 0.65 0.54 49 Cloud forest-dry soil

0--10 0.19 0.04 6 0.20 0.25 0.15 49 Colorado forest-dry forest
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

473

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--10 0.32 0.06 6 0.30 0.41 0.25 49 Tabonuco forest-dry forest

10--35 0.33 0.08 5 0.33 0.41 0.21 49 Cloud forest-dry soil

10--35 0.13 0.01 5 0.13 0.15 0.11 49 Colorado forest-dry forest

10--35 0.14 0.01 5 0.14 0.16 0.13 49 Tabonuco forest-dry forest

Organic matter (%) 0--10 24.58 1.05 6 24.64 25.84 22.97 49 Cloud forest-dry soil

0--10 7.80 1.29 6 7.75 9.63 6.19 49 Colorado forest-dry forest

0--10 9.85 2.39 6 10.17 13.73 6.87 49 Tabonuco forest-dry forest

10--35 15.91 4.05 5 16.39 19.61 10.07 49 Cloud forest-dry soil

10--35 4.27 0.72 5 4.56 4.76 3.01 49 Colorado forest-dry forest

10--35 3.59 0.58 5 3.37 4.41 3.05 49 Tabonuco forest-dry forest

pH (H2O) 0--10 4.89 0.05 6 4.88 4.96 4.85 49 Cloud forest-dry soil

0--10 4.81 0.11 6 4.86 4.89 4.63 49 Colorado forest-dry forest

0--10 4.25 0.14 6 4.22 4.44 4.11 49 Tabonuco forest-dry forest

0--10 5.19 0.09 6 5.22 5.29 5.03 49 Cloud forest-wet soil

0--10 5.14 0.13 6 5.08 5.37 5.02 49 Colorado forest-wet soil

0--10 5.12 0.06 6 5.12 5.19 5.03 49 Tabaonuco forest-wet soil

10--35 5.07 0.05 5 5.04 5.13 5.03 49 Cloud forest-dry soil

10--35 4.74 0.09 5 4.72 4.84 4.62 49 Colorado forest-dry forest

10--35 4.36 0.26 5 4.43 4.62 4.07 49 Tabonuco forest-dry forest

10--35 5.20 0.15 5 5.15 5.47 5.10 49 Cloud forest-wet soil

10--35 5.30 0.22 5 5.18 5.66 5.13 49 Colorado forest-wet soil

10--35 5.17 0.09 5 5.18 5.29 5.05 49 Tabaonuco forest-wet soil

pH (KCl) 0--10 4.09 0.06 6 4.06 4.16 4.03 49 Cloud forest-dry soil

0--10 3.95 0.07 6 3.96 4.05 3.83 49 Colorado forest-dry forest

0--10 3.55 0.16 6 3.53 3.76 3.32 49 Tabonuco forest-dry forest

10--35 4.09 0.03 5 4.11 4.12 4.05 49 Cloud forest-dry soil
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

474

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--35 4.02 0.09 5 4.03 4.12 3.90 49 Colorado forest-dry forest

10--35 3.72 0.15 5 3.79 3.85 3.50 49 Tabonuco forest-dry forest

Phosphorus (mg/kg) 0--10 13 2 6 13 15 10 49 Cloud forest-dry soil

0--10 9 1 6 9 10 7 49 Colorado forest-dry forest

0--10 14 5 6 16 18 6 49 Tabonuco forest-dry forest

10--35 9 2 5 10 12 6 49 Cloud forest-dry soil

10--35 4 1 5 4 5 3 49 Colorado forest-dry forest

10--35 3 1 4 3 4 1 49 Tabonuco forest-dry forest

0--10 10 2 6 9 12 8 49 Cloud forest-wet soil

0--10 4 1 6 4 5 3 49 Colorado forest-wet soil

0--10 7 2 6 7 9 5 49 Tabaonuco forest-wet soil

10--35 9 2 5 9 11 5 49 Cloud forest-wet soil

10--35 3 1 5 3 3 2 49 Colorado forest-wet soil

10--35 5 1 5 6 6 3 49 Tabaonuco forest-wet soil

Potassium (mg/kg) 0--10 114 19 6 122 130 78 49 Cloud forest-dry soil

0--10 90 25 6 90 121 50 49 Colorado forest-dry forest

0--10 94 25 6 88 143 72 49 Tabonuco forest-dry forest

10--35 70 10 5 72 85 59 49 Cloud forest-dry soil

10--35 59 17 5 67 76 38 49 Colorado forest-dry forest

10--35 34 7 4 33 42 27 49 Tabonuco forest-dry forest

0--10 40 9 6 41 52 26 49 Cloud forest-wet soil

0--10 47 10 6 50 58 31 49 Colorado forest-wet soil

0--10 48 27 6 59 73 7 49 Tabaonuco forest-wet soil

10--35 35 12 5 31 48 22 49 Cloud forest-wet soil

10--35 29 15 5 27 52 13 49 Colorado forest-wet soil

10--35 11 8 5 13 20 1 49 Tabaonuco forest-wet soil
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

475

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Sodium (mg/g) 0--10 0.10 0.02 6 0.10 0.12 0.08 49 Cloud forest-dry soil

0--10 0.05 0.01 6 0.05 0.07 0.04 49 Colorado forest-dry forest

0--10 0.10 0.01 6 0.12 0.13 0.11 49 Tabonuco forest-dry forest

10--35 0.08 0.01 5 0.08 0.09 0.06 49 Cloud forest-dry soil

10--35 0.04 0.01 5 0.04 0.05 0.02 49 Colorado forest-dry forest

10--35 0.07 0.01 5 0.07 0.09 0.06 49 Tabonuco forest-dry forest

Tuffaceous sandstone Ultisols Lmrf Carbon (%) 0--10 7.20 2.95 16 6.36 12.18 3.57 50 Dacroydes excelsa

0--10 7.09 4.82 21 5.54 23.03 3.05 50 Guarea guidonia

10--20 3.10 0.58 19 3.18 3.94 2.03 50 Dacroydes excelsa

10--20 4.99 3.54 25 3.85 15.92 2.74 50 Guarea guidonia

Nitrogen (%) 0--10 0.47 0.14 16 0.44 0.77 0.28 50 Dacroydes excelsa

0--10 0.47 0.16 21 0.41 0.90 0.28 50 Guarea guidonia

10--20 0.25 0.04 19 0.25 0.34 0.17 50 Dacroydes excelsa

10--20 0.34 0.12 25 0.31 0.70 0.24 50 Guarea guidonia

Sulfur (%) 0--10 0.07 0.01 16 0.06 0.09 0.05 50 Dacroydes excelsa

0--10 0.07 0.02 21 0.07 0.12 0.04 50 Guarea guidonia

10--20 0.04 0.01 19 0.04 0.06 0.04 50 Dacroydes excelsa

10--20 0.06 0.02 25 0.06 0.10 0.03 50 Guarea guidonia

Plutonic rocks Inceptisols Mfs Aluminum-total (mg/g) 0--15 34.87 4.29 16 35.52 42.71 28.66 51 Secondary forest-tobacco

0--15 47.88 7.05 15 46.75 62.99 38.97 51 Secondary forest-coffee

15--30 37.59 6.80 17 35.27 55.63 29.99 51 Secondary forest-tobacco

15--30 49.82 9.80 20 47.33 66.79 36.13 51 Secondary forest-coffee

Calcium-total (mg/g) 0--15 4.24 1.16 16 4.38 5.84 1.50 51 Secondary forest-tobacco

0--15 5.38 1.72 15 5.76 7.49 2.43 51 Secondary forest-coffee

15--30 3.51 1.22 17 3.57 5.79 1.43 51 Secondary forest-tobacco

15--30 4.43 1.60 20 5.21 6.04 2.05 51 Secondary forest-coffee
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

476

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Carbon (%) 0--15 1.60 0.38 16 1.65 2.43 1.08 51 Secondary forest-tobacco

0--15 1.71 0.47 15 1.65 2.61 1.02 51 Secondary forest-coffee

15--30 0.89 0.32 17 0.87 1.63 0.48 51 Secondary forest-tobacco

15--30 0.75 0.39 18 0.61 1.57 0.32 51 Secondary forest-coffee

Iron-total (mg/g) 0--15 43.79 2.70 16 43.84 49.87 39.46 51 Secondary forest-tobacco

0--15 60.85 8.43 15 59.79 74.24 46.26 51 Secondary forest-coffee

15--30 45.49 4.77 17 44.31 59.25 39.97 51 Secondary forest-tobacco

15--30 60.89 10.83 20 62.18 80.23 45.15 51 Secondary forest-coffee

Loss on ignition (%) 0--15 91.66 1.47 14 91.47 93.81 89.91 51 Secondary forest-tobacco

0--15 91.85 1.86 15 91.86 94.38 88.57 51 Secondary forest-coffee

15--30 94.18 1.62 15 93.60 96.87 91.83 51 Secondary forest-tobacco

15--30 95.66 1.76 18 95.88 98.18 91.87 51 Secondary forest-coffee

Magnesium-total (mg/g) 0--15 8.59 1.74 16 8.48 12.07 5.60 51 Secondary forest-tobacco

0--15 14.14 3.28 15 14.54 18.14 8.71 51 Secondary forest-coffee

15--30 8.47 1.87 17 8.10 12.45 5.68 51 Secondary forest-tobacco

15--30 14.65 4.20 20 16.21 20.56 7.39 51 Secondary forest-coffee

Manganese-total (mg/g) 0--15 1.18 0.13 16 1.20 1.36 0.90 51 Secondary forest-tobacco

0--15 1.18 0.18 15 1.15 1.41 0.76 51 Secondary forest-coffee

15--30 1.18 0.14 17 1.20 1.41 0.97 51 Secondary forest-tobacco

15--30 1.12 0.14 20 1.11 1.36 0.75 51 Secondary forest-coffee

Nitrogen (%) 0--15 0.14 0.03 16 0.15 0.20 0.10 51 Secondary forest-tobacco

0--15 0.15 0.04 15 0.15 0.23 0.10 51 Secondary forest-coffee

15--30 0.09 0.03 17 0.09 0.14 0.05 51 Secondary forest-tobacco

15--30 0.07 0.03 18 0.06 0.14 0.04 51 Secondary forest-coffee

pH (H2O) 0--15 6.39 0.36 14 6.39 6.91 5.48 51 Secondary forest-tobacco

0--15 6.26 0.27 15 6.25 6.67 5.91 51 Secondary forest-coffee
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

477

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

15--30 6.07 0.48 15 6.12 6.90 5.33 51 Secondary forest-tobacco

15--30 6.04 0.29 18 6.08 6.48 5.52 51 Secondary forest-coffee

pH (KCl) 0--15 4.80 0.40 14 4.80 5.39 3.95 51 Secondary forest-tobacco

0--15 4.61 0.34 15 4.62 5.06 4.00 51 Secondary forest-coffee

15--30 4.33 0.56 15 4.39 5.60 3.54 51 Secondary forest-tobacco

15--30 4.11 0.30 18 4.01 4.74 3.78 51 Secondary forest-coffee

Phosphorus-total (mg/g) 0--15 0.35 0.10 16 0.32 0.58 0.22 51 Secondary forest-tobacco

0--15 0.28 0.06 15 0.27 0.38 0.18 51 Secondary forest-coffee

15--30 0.30 0.07 17 0.28 0.44 0.17 51 Secondary forest-tobacco

15--30 0.21 0.05 20 0.20 0.31 0.14 51 Secondary forest-coffee

Potassium-total (mg/g) 0--15 1.71 0.28 16 1.64 2.56 1.32 51 Secondary forest-tobacco

0--15 1.85 0.20 15 1.85 2.28 1.52 51 Secondary forest-coffee

15--30 1.54 0.19 17 1.47 1.99 1.28 51 Secondary forest-tobacco

15--30 1.68 0.18 20 1.69 1.99 1.35 51 Secondary forest-coffee

Sulfur (%) 0--15 0.02 0.01 16 0.02 0.03 0.01 51 Secondary forest-tobacco

0--15 0.02 0.01 15 0.02 0.03 0.01 51 Secondary forest-coffee

15--30 0.01 0.00 17 0.01 0.02 0.01 51 Secondary forest-tobacco

15--30 0.01 0.01 18 0.01 0.02 0.01 51 Secondary forest-coffee

Tuffaceous sandstone Ultisols Smf C/N 0--10 13 2 28 13 17 11 52

Carbon (%) 0--10 5.43 1.71 29 5.19 9.51 2.97 52

Loss on ignition (%) 0--10 18.57 4.12 28 17.95 30.35 13.61 52

Nitrogen (%) 0--10 0.41 0.10 29 0.40 0.63 0.25 52

pH (H2O) 0--10 5.45 0.76 24 5.34 7.12 4.48 52

pH (KCl) 0--10 4.75 0.80 24 4.63 6.70 3.80 52

Sulfur (%) 0--10 0.06 0.01 29 0.06 0.08 0.04 52

Tuffaceous sandstone Ultisols Smf Aluminum-total (mg/g) 0--10 24.16 11.60 23 24.75 45.52 6.57 53
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

478

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Calcium-total (mg/g) 0--10 1.41 1.36 49 0.89 6.35 0.29 53

Carbon (%) 0--10 1.86 0.67 26 2.08 2.79 0.34 53

Iron (mg/g) 0--10 0.08 0.03 4 0.07 0.10 0.05 53

Iron-total (mg/g) 0--10 27.85 12.13 23 33.92 47.21 8.87 53

Loss on ignition (%) 0--10 13.05 4.79 48 13.51 21.95 2.27 53

Magnesium-total (mg/g) 0--10 2.95 2.06 49 2.46 8.74 0.34 53

Manganes (mg/g) 0--10 0.02 0.00 4 0.02 0.02 0.02 53

Manganese-total (mg/g) 0--10 0.95 0.61 49 1.00 2.40 0.13 53

Nitrogen (%) 0--10 0.19 0.07 26 0.21 0.28 0.03 53

pH (H2O) 0--10 4.54 0.58 18 4.46 6.38 3.91 53

pH (KCl) 0--10 3.75 0.61 18 3.59 5.82 3.28 53

Phosphorus (mg/g) 0--10 1.49 0.33 4 1.50 1.79 1.16 53

Phosphorus-total (mg/g) 0--10 0.88 0.79 49 0.60 4.32 0.26 53

Potassium (mg/g) 0--10 0.30 0.07 4 0.30 0.36 0.24 53

Potassium-total (mg/g) 0--10 1.89 0.92 49 1.65 5.21 0.79 53

Tuffaceous sandstone Ultisols Mfs Aluminum (cmol/kg-l) 0-15 3.01 2.06 22 2.40 8.28 0.60 54

Aluminum-total (mg/g) 0-15 46.63 10.72 22 47.01 62.25 28.22 54

C/N 0-15 14 3 22 14 24 11 54

Calcium (cmol/kg-l) 0-15 4.80 2.35 2 4.73 10.03 1.19 54

Calcium (mg/g) 0-15 0.96 0.47 22 0.95 2.01 0.24 54

Calcium-total (mg/g) 0-15 1.13 0.65 22 1.04 2.66 0.27 54

Carbon (%) 0-15 9.60 6.02 22 8.28 27.42 1.42 54

ECEC (cmol/kg-1) 0-15 10.70 2.72 22 10.33 17.31 6.56 54

Iron (mg/g) 0-15 0.89 0.59 22 0.70 2.89 0.07 54

Iron-total (mg/g) 0-15 52.52 15.61 22 54.05 90.67 27.62 54

Loss on ignition (%) 0-15 30.68 12.75 22 29.64 55.82 8.53 54
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

479

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Magnesium (cmol/kg-l) 0-15 1.93 0.89 22 1.75 4.48 0.56 54

Magnesium (mg/g) 0-15 0.23 0.11 22 0.21 0.54 0.07 54

Magnesium-total (mg/g) 0-15 4.89 2.32 22 4.62 9.39 0.56 54

Manganese (mg/g) 0-15 0.13 0.09 22 0.09 0.41 0.05 54

Manganese-total (mg/g) 0-15 1.06 0.65 22 0.98 2.95 0.17 54

Nitrogen (%) 0-15 0.66 0.36 22 0.60 1.68 0.09 54

pH (H2O) 0-15 4.80 0.37 21 4.81 5.70 4.32 54

pH (KCl) 0-15 3.72 0.26 21 3.64 4.26 3.28 54

Phosphorus (mg/g) 0-15 0.04 0.02 22 0.03 0.08 0.01 54

Phosphorus-total (mg/g) 0-15 0.78 0.34 22 0.75 1.39 0.22 54

Potassium-total (mg/g) 0-15 1.60 0.39 22 1.54 2.61 1.00 54

Potassium (cmol/kg-l) 0-15 0.52 0.20 22 0.48 1.07 0.23 54

Potassium (mg/g) 0-15 0.20 0.08 22 0.18 0.41 0.09 54

Sodium (cmol/kg-l) 0-15 0.44 0.06 22 0.44 0.57 0.37 54

Sodium (mg/g) 0-15 0.10 0.01 22 0.10 0.13 0.09 54

Sulfur (%) 0-15 0.09 0.04 22 0.09 0.20 0.01 54

Tuffaceous sandstone Ultisols Mfs Aluminum (meq/100g) 0--15 11.54 13.47 11 5.00 34.79 0.73 55

Dfs Bulk density (g/cc) 0--15 0.27 0.06 11 0.28 0.34 0.12 55

Dfs 15--30 0.57 0.21 2 0.57 0.71 0.42 55

Dfs C/N 0--15 9 1 11 9 10 8 55

Dfs Calcium (cmol/kg-l) 0--15 49.11 1.82 14 39.35 90.05 27.36 55

Dfs 15--30 58.02 4.62 2 58.02 61.29 54.76 55

Dfs Calcium (mg/g) 0--15 9.82 4.37 14 7.87 18.01 5.47 55

Dfs 15--30 11.61 0.92 2 11.61 12.26 1.95 55

Dfs Carbon (%) 0--15 25.28 7.95 11 23.60 41.28 17.39 55

Dfs Effective cation exchange capacity 0--15 69.89 27.54 14 57.65 119.49 41.11 55
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

480

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Dfs {Ca+K+Mg(cmol/kg-1) 15--30 70.46 6.10 2 70.49 74.81 66.17 55

Dfs Iron (mg/g) 0--15 0.28 0.36 14 0.12 1.16 0.01 55

Dfs 15--30 0.62 0.41 2 0.62 0.91 0.34 55

Dfs Loss on ignition (%) 0--15 47.32 11.38 14 42.80 73.20 35.43 55

Dfs 15--30 28.74 4.09 2 28.74 31.63 25.85 55

Dfs Magnesium (cmol/kg-l) 0--15 4.08 0.77 14 3.85 5.88 2.81 55

Dfs 15--30 4.78 1.67 2 4.78 5.96 3.60 55

Dfs Magnesium (mg/g) 0--15 0.49 0.09 14 0.46 0.71 0.34 55

Dfs 15--30 0.57 0.20 2 0.57 0.72 0.43 55

Dfs Manganese (mg/kg) 0--15 9 4 14 10 15 2 55

Dfs 15--30 6 4 2 6 8 3 55

Dfs Nitrogen (%) 0--15 2.65 0.91 14 2.22 4.40 1.74 55

Dfs 15--30 1.28 0.13 2 1.28 1.37 1.19 55

Dfs pH (H2O) 0--15 4.34 0.81 11 4.27 5.94 3.41 55

Dfs 15--30 3.89 0.30 2 3.89 4.10 3.68 55

Dfs pH (KCl) 0--15 4.15 0.74 11 4.20 5.57 3.21 55

Dfs 15--30 3.78 0.25 2 3.78 3.95 3.60 55

Dfs Phosphorus (mg/g) 0--15 1.87 2.10 14 0.74 5.81 0.20 55

Dfs 15--30 4.76 0.44 2 4.76 5.08 4.45 55

Dfs Potassium (cmol/kg-l) 0--15 2.79 2.30 14 2.07 7.77 0.86 55

Dfs 15--30 5.75 1.22 2 5.75 6.62 4.89 55

Dfs Potassium (mg/g) 0--15 1.06 0.88 14 0.79 2.95 0.33 55

Dfs 15--30 2.19 0.46 2 2.19 2.51 1.86 55

Dfs Sodium (cmol/kg-l) 0--15 4.84 2.04 14 5.14 8.16 1.54 55

Dfs 15--30 1.94 1.93 2 1.94 3.30 0.57 55

Dfs Sodium (mg/g) 0--15 1.11 0.47 14 1.18 1.88 0.36 55
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

481

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Dfs 15--30 0.45 0.44 2 0.45 0.76 0.13 55

Dfs Sulfur (%) 0--15 0.57 0.19 11 0.54 0.91 0.32 55

Tuffaceous sandstone Ultisols Wfs Aluminum (cmol/kg-l) 0-9 1.01 0.43 5 0.70 1.55 0.69 56 Coffee plantation

0-9 1.02 0.22 5 0.95 1.33 0.81 56 Pasture

C/N 0-9 12 1 5 11 13 10 56 Coffee plantation

0-9 13 2 5 12 16 12 56 Pasture

Calcium (cmol/kg-l) 0-9 14.98 2.91 5 14.38 18.23 11.97 56 Coffee plantation

0-9 7.70 2.96 5 9.51 9.95 3.09 56 Pasture

Calcium (mg/g) 0-9 3.00 0.58 5 2.88 3.65 2.39 56 Coffee plantation

0-9 1.54 0.59 5 1.90 1.99 0.62 56 Pasture

Carbon (%) 0-9 7.78 1.93 5 7.61 9.79 5.69 56 Coffee plantation

0-9 6.09 2.68 5.58 10.66 3.71 56 Pasture

Effective cation exchange capacity 0-9 25.00 3.07 5 23.33 28.60 22.44 56 Coffee plantation

{Ca+K+Mg(cmol/kg-1) 0-9 12.82 3.06 5 14.40 15.97 7.30 56 Pasture

Iron (mg/g) 0-9 0.16 0.04 5 0.17 0.21 0.11 56 Coffee plantation

0-9 0.32 0.08 5 0.28 0.45 0.26 56 Pasture

Loss on ignition (%) 0-9 25.17 3.07 5 26.63 27.76 20.92 56 Coffee plantation

0-9 18.23 4.97 5 18.74 25.78 12.50 56 Pasture

Magnesium (cmol/kg-l) 0-9 7.75 0.90 5 7.92 8.52 6.35 56 Coffee plantation

0-9 2.96 0.80 5 3.04 3.75 1.76 56 Pasture

Magnesium (mg/g) 0-9 0.93 0.11 5 0.95 1.02 0.76 56 Coffee plantation

0-9 0.36 0.10 5 0.36 0.45 0.21 56 Pasture

Manganese (mg/g) 0-9 0.21 0.04 5 0.20 0.26 0.17 56 Coffee plantation

0-9 0.10 0.03 5 0.09 0.14 0.07 56 Pasture

Nitrogen (%) 0-9 0.67 0.12 5 0.71 0.78 0.50 56 Coffee plantation

0-9 0.45 0.13 5 0.43 0.66 0.31 56 Pasture
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

482

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

pH (H2O) 0-9 5.85 0.24 5 5.93 6.04 5.45 56 Coffee plantation

0-9 5.47 0.19 5 5.41 5.80 5.35 56 Pasture

pH (KCl) 0-9 4.87 0.26 5 4.98 5.09 4.44 56 Coffee plantation

0-9 4.48 0.22 5 4.45 4.83 4.22 56 Pasture

Phosphorus (mg/kg) 0-9 21 3 5 23 25 17 56 Coffee plantation

0-9 16 3 5 14 22 14 56 Pasture

Potassium (cmol/kg-l) 0-9 0.69 0.09 5 0.70 0.79 0.56 56 Coffee plantation

0-9 0.68 0.08 5 0.66 0.80 0.60 56 Pasture

Potassium (mg/g) 0-9 0.26 0.03 5 0.27 0.30 0.21 56 Coffee plantation

0-9 0.26 0.03 5 0.25 0.30 0.23 56 Pasture

Sodium (cmol/kg-l) 0-9 0.57 0.22 5 0.50 0.96 0.39 56 Coffee plantation

0-9 0.45 0.02 5 0.45 0.48 0.42 56 Pasture

Sodium (mg/g) 0-9 0.13 0.05 5 0.11 0.22 0.09 56 Coffee plantation

0-9 0.15 0.01 5 0.10 0.11 0.10 56 Pasture

Sulfur (%) 0-9 0.15 0.03 5 0.15 0.18 0.09 56 Coffee plantation

0-9 0.10 0.03 5 0.10 0.15 0.07 56 Pasture

Alluvial deposits Inceptisols Dfs Aluminum-total (mg/g) 0--15 19.90 5.59 15 21.20 28.12 7.30 57

15--30 21.72 4.99 15 23.43 28.17 13.69 57

Bulk density (g/cc) 0--15 0.81 0.35 15 0.75 1.34 0.13 57

15--30 0.89 0.42 15 0.88 1.54 0.07 57

C/N 0--15 28 9 15 31 43 12 57

15--30 45 26 15 38 120 15 57

Calcium-total (mg/g) 0--15 48.66 20.54 15 50.11 84.47 18.57 57

15--30 51.44 29.21 15 52.32 116.27 9.53 57

Carbon (%) 0--15 4.72 3.58 15 3.84 17.24 2.87 57

15--30 6.01 4.79 15 4.12 19.68 2.36 57
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

483

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Iron-total (mg/g) 0--15 25.82 8.06 15 28.89 37.25 11.80 57

15--30 27.01 8.59 15 32.29 36.81 13.28 57

Loss on ignition (%) 0--15 18.45 11.60 15 15.51 52.78 8.76 57

15--30 19.29 11.53 15 14.78 41.96 7.91 57

Magnesium-total (mg/g) 0--15 14.44 5.50 15 12.88 31.57 7.67 57

15--30 13.17 2.32 15 13.34 16.45 8.08 57

Manganese-total (mg/g) 0--15 1.09 0.33 15 1.08 1.64 0.34 57

15--30 1.00 0.43 15 1.15 1.71 0.40 57

Nitrogen (%) 0--15 0.19 0.17 15 0.13 0.76 0.08 57

15--30 0.16 0.13 15 0.09 0.47 0.05 57

pH (H2O) 0--15 7.77 0.24 15 7.78 8.32 7.35 57

15--30 7.87 0.34 15 7.86 8.53 7.29 57

pH (KCl) 0--15 7.50 0.19 15 7.47 8.02 7.28 57

15--30 7.37 0.32 15 7.48 7.79 6.49 57

Phosphorus-total (mg/g) 0--15 1.79 2.34 15 0.73 7.22 0.47 57

15--30 0.97 0.92 15 0.58 3.86 0.48 57

Potassium-total (mg/g) 0--15 6.28 1.45 15 6.68 7.96 2.87 57

15--30 7.08 1.21 15 7.17 9.11 4.61 57

Sulfur (%) 0--15 0.32 0.43 15 0.12 1.70 0.03 57

15--30 0.50 0.59 15 0.11 1.61 0.02 57

Serpentinite Ultisols Lmrf Aluminum (cmol/kg-l) 0--5 0.08 0.04 20 0.07 0.15 0.03 58 Baseball park

0--5 0.14 0.04 20 0.14 0.22 0.06 58 Farmlands

0--5 0.17 0.08 20 0.15 0.34 0.01 58 Charcoal house

0--5 0.19 0.10 31 0.18 0.59 0.04 58 Houses

0--5 0.16 0.07 9 0.16 0.25 0.04 58 Mature forest

5--10 0.07 0.04 24 0.07 0.14 0.07 58 Baseball park
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

484

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

5--10 0.12 0.04 24 0.13 0.19 0.03 58 Farmlands

5--10 0.13 0.06 24 0.12 0.29 0.04 58 Charcoal house

5--10 0.14 0.05 36 0.14 0.24 0.03 58 Houses

5--10 0.11 0.06 12 0.12 0.21 0.02 58 Mature forest

Bulk density (g/cc) 0--5 0.44 0.04 20 0.42 0.55 0.38 58 Baseball park

0--5 0.56 0.11 20 0.56 0.77 0.40 58 Farmlands

0--5 0.39 0.10 20 0.40 0.55 0.18 58 Charcoal house

0--5 0.52 0.22 31 0.56 0.95 0.15 58 Houses

0--5 0.23 0.08 9 0.21 0.38 0.17 58 Mature forest

5--10 0.71 0.11 24 0.7 0.92 0.49 58 Baseball park

5--10 0.69 0.13 24 0.72 0.99 0.48 58 Farmlands

5--10 0.44 0.14 24 0.39 0.73 0.26 58 Charcoal house

5--10 0.52 0.30 36 0.58 1.15 0.10 58 Houses

5--10 0.27 0.11 12 0.32 0.38 0.08 58 Mature forest

C/N 0--5 7 0 20 7 8 7 58 Baseball park

0--5 7 1 20 7 8 6 58 Farmlands

0--5 8 1 20 8 10 6 58 Charcoal house

0--5 8 1 31 8 9 7 58 Houses

0--5 8 1 9 8 9 7 58 Mature forest

5--10 7 0 24 7 8 6 58 Baseball park

5--10 7 0 24 7 7 6 58 Farmlands

5--10 7 1 24 7 10 6 58 Charcoal house

5--10 7 1 36 7 9 6 58 Houses

5--10 7 0 12 7 8 7 58 Mature forest

Calcium (cmol/kg-l) 0--5 47.21 15.49 20 52.54 67.13 18.82 58 Baseball park

0--5 34.65 8.44 20 37.40 50.74 17.06 58 Farmlands
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

485

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--5 47.04 9.38 20 46.96 74.78 29.84 58 Charcoal house

0--5 45.40 16.04 31 42.04 71.05 15.91 58 Houses

0--5 68.69 8.31 9 67.99 80.75 58.56 58 Mature forest

5--10 49.61 15.27 24 51.34 71.35 21.74 58 Baseball park

5--10 33.13 8.64 24 35.30 49.27 11.98 58 Farmlands

5--10 44.86 9.22 24 44.65 65.96 29.81 58 Charcoal house

5--10 43.09 16.48 36 36.47 70.17 15.34 58 Houses

5--10 66.46 8.07 12 68.21 79.88 54.67 58 Mature forest

Calcium (mg/g) 0--5 9.44 3.10 20 10.51 13.43 3.76 58 Baseball park

0--5 6.93 1.69 20 7.48 10.15 3.41 58 Farmlands

0--5 9.41 1.88 20 9.39 14.96 5.97 58 Charcoal house

0--5 9.08 3.21 31 8.41 14.21 3.18 58 Houses

0--5 13.74 1.66 9 13.60 16.15 11.71 58 Mature forest

5--10 9.92 3.05 24 10.27 14.27 4.35 58 Baseball park

5--10 6.62 1.73 24 7.06 9.85 2.40 58 Farmlands

5--10 8.97 1.84 24 8.93 13.19 5.96 58 Charcoal house

5--10 8.62 3.30 36 7.29 14.03 3.07 58 Houses

5--10 13.29 1.65 12 13.64 15.98 10.94 58 Mature forest

Carbon (%) 0--5 4.15 0.72 20 4.31 5.64 2.88 58 Baseball park

0--5 4.05 1.11 20 3.78 7.04 2.78 58 Farmlands

0--5 7.27 2.16 20 6.79 11.65 4.65 58 Charcoal house

0--5 7.52 3.98 31 5.73 17.35 2.42 58 Houses

0--5 11.67 4.36 9 9.75 20.46 7.86 58 Mature forest

5--10 3.62 0.63 24 3.73 4.76 2.46 58 Baseball park

5--10 3.33 1.08 24 3.28 6.74 1.73 58 Farmlands

5--10 6.20 2.09 24 5.35 11.32 3.86 58 Charcoal house
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

486

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

5--10 6.33 3.43 36 4.89 15.63 1.64 58 Houses

5--10 8.42 1.91 12 8.56 11.68 5.89 58 Mature forest

Effective cation exchange capacity 0--5 55.19 15.59 20 61.72 73.74 26.79 58 Baseball park

{Ca+K+Mg(cmol/kg-1) 0--5 40.43 8.72 20 42.80 58.27 21.54 58 Farmlands

0--5 52.87 11.16 20 51.91 85.88 32.69 58 Charcoal house

0--5 51.32 17.63 31 46.27 78.10 18.83 58 Houses

0--5 76.57 8.36 9 77.20 90.03 65.76 58 Mature forest

5--10 56.77 15.15 24 59.32 77.64 28.45 58 Baseball park

5--10 38.28 9.15 24 39.95 55.55 15.19 58 Farmlands

5--10 50.18 11.24 24 48.23 74.96 33.07 58 Charcoal house

5--10 48.24 18.15 36 40.22 78.04 18.15 58 Houses

5--10 73.18 8.04 12 74.45 87.26 62.81 58 Mature forest

Iron (mg/g) 0--5 0.51 1.57 20 0.01 5.70 0.01 58 Baseball park

0--5 6.95 2.77 20 6.64 12.62 3.12 58 Farmlands

0--5 4.55 5.31 20 0.02 13.08 0.01 58 Charcoal house

0--5 0.01 0.01 31 0.01 0.03 0.01 58 Houses

0--5 0.01 0.01 9 0.01 0.02 0.01 58 Mature forest

5--10 1.04 2.08 24 0.01 5.87 0.01 58 Baseball park

5--10 6.37 2.17 24 6.02 11.38 3.20 58 Farmlands

5--10 3.12 4.22 24 0.02 10.56 0.01 58 Charcoal house

5--10 0.01 0.01 36 0.01 0.03 0.01 58 Houses

5--10 0.01 0.00 12 0.01 0.02 0.01 58 Mature forest

Loss on ignition (%) 0--5 25.57 1.63 20 22.67 25.47 19.45 58 Baseball park

0--5 17.23 3.69 20 16.28 26.64 11.86 58 Farmlands

0--5 26.98 6.73 20 25.67 42.76 19.39 58 Charcoal house

0--5 27.65 12.68 31 21.19 53.67 9.84 58 Houses
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

487

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--5 40.57 9.84 9 38.19 60.53 31.66 58 Mature forest

5--10 21.15 1.3 24 21.54 23.31 18.73 58 Baseball park

5--10 15.13 3.21 24 14.50 24.02 8.74 58 Farmlands

5--10 23.83 5.90 24 22.68 39.77 17.28 58 Charcoal house

5--10 24.95 12.59 36 18.56 53.47 7.82 58 Houses

5--10 33.46 5.59 12 32.94 42.41 26.57 58 Mature forest

Magnesium (cmol/kg-l) 0--5 6.76 0.99 20 6.61 8.46 5.35 58 Baseball park

0--5 4.40 1.35 20 4.39 8.38 1.85 58 Farmlands

0--5 4.54 2.52 20 3.86 12.85 1.98 58 Charcoal house

0--5 4.40 2.04 31 3.81 9.20 1.77 58 Houses

0--5 6.65 1.28 9 6.60 7.90 4.70 58 Mature forest

5--10 6.08 0.89 24 6.13 7.67 4.76 58 Baseball park

5--10 3.86 1.28 24 3.73 7.05 1.49 58 Farmlands

5--10 4.10 2.40 24 3.32 10.81 2.07 58 Charcoal house

5--10 3.80 1.91 36 3.05 9.50 1.61 58 Houses

5--10 5.60 0.97 12 5.87 6.87 3.98 58 Mature forest

Magnesium (mg/g) 0--5 0.81 0.12 22 0.79 1.02 0.64 58 Baseball park

0--5 0.53 0.16 20 0.53 1.01 0.22 58 Farmlands

0--5 0.55 0.30 20 0.46 1.54 0.24 58 Charcoal house

0--5 0.53 0.25 31 0.46 1.10 0.21 58 Houses

0--5 0.80 0.15 9 0.79 0.95 0.56 58 Mature forest

5--10 0.73 0.11 24 0.74 0.92 0.57 58 Baseball park

5--10 0.46 0.15 24 0.45 0.85 0.18 58 Farmlands

5--10 0.49 0.29 24 0.40 1.30 0.25 58 Charcoal house

5--10 0.46 0.23 36 0.37 1.14 0.19 58 Houses

5--10 0.67 0.12 12 0.70 0.82 0.48 58 Mature forest
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

488

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Manganese (mg/g) 0--5 0.12 0.05 20 0.11 0.30 0.08 58 Baseball park

0--5 0.06 0.02 20 0.07 0.08 0.01 58 Farmlands

0--5 0.08 0.02 20 0.07 0.13 0.04 58 Charcoal house

0--5 0.07 0.04 31 0.08 0.14 0.01 58 Houses

0--5 0.10 0.05 9 0.11 0.16 0.02 58 Mature forest

5--10 0.10 0.03 24 0.09 0.20 0.07 58 Baseball park

5--10 0.05 0.02 24 0.06 0.07 0.01 58 Farmlands

5--10 0.07 0.02 24 0.07 0.10 0.03 58 Charcoal house

5--10 0.07 0.03 36 0.06 0.14 0.01 58 Houses

5--10 0.06 0.04 12 0.04 0.12 0.02 58 Mature forest

Nitrogen (%) 0--5 0.58 0.09 20 0.61 0.72 0.40 58 Baseball park

0--5 0.53 0.08 20 0.56 0.66 0.37 58 Baseball park

0--5 0.58 0.20 20 0.53 1.16 0.36 58 Farmlands

0--5 0.55 0.18 20 0.51 1.09 0.34 58 Farmlands

0--5 0.96 0.28 20 0.88 1.64 0.57 58 Charcoal house

0--5 0.86 0.29 20 0.83 1.47 0.20 58 Charcoal house

0--5 0.99 0.51 31 0.81 1.95 0.28 58 Houses

0--5 1.49 0.43 9 1.33 2.21 0.97 58 Mature forest

5--10 0.52 0.09 24 0.53 0.66 0.35 58 Baseball park

5--10 0.47 0.08 24 0.48 0.60 0.32 58 Baseball park

5--10 0.50 0.17 24 0.49 1.05 0.27 58 Farmlands

5--10 0.47 0.16 24 0.47 0.99 0.26 58 Farmlands

5--10 0.84 0.25 24 0.80 1.34 0.51 58 Charcoal house

5--10 0.89 0.51 36 0.69 2.09 0.18 58 Houses

5--10 1.18 0.26 12 1.22 1.58 0.80 58 Mature forest

Organic matter (%) 0--5 8.98 1.57 20 9.31 12.19 6.22 58 Baseball park
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

489

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--5 8.75 2.39 20 8.18 15.23 6.01 58 Farmlands

0--5 15.71 4.68 20 14.69 25.19 10.05 58 Charcoal house

0--5 16.26 8.60 31 12.39 37.52 5.24 58 Houses

0--5 25.24 9.43 9 21.08 44.23 17.00 58 Mature forest

5--10 7.82 1.37 24 8.07 10.3 5.31 58 Baseball park

5--10 7.15 2.33 24 7.10 14.57 3.75 58 Farmlands

5--10 13.40 4.52 24 11.58 24.47 8.34 58 Charcoal house

5--10 13.69 7.42 36 10.58 33.79 3.55 58 Houses

5--10 18.20 4.14 12 18.51 25.26 12.74 58 Mature forest

pH (H2O) 0--5 7.36 0.32 20 7.40 7.74 6.53 58 Baseball park

0--5 7.77 0.09 20 7.76 7.92 7.61 58 Farmlands

0--5 7.77 0.14 20 7.79 7.91 7.36 58 Charcoal house

0--5 7.81 0.17 31 7.81 8.08 7.48 58 Houses

0--5 7.55 0.12 9 7.56 7.74 7.36 58 Mature forest

5--10 7.4 0.33 24 7.51 7.76 6.70 58 Baseball park

5--10 7.85 0.09 24 7.82 8.04 7.69 58 Farmlands

5--10 7.84 0.09 24 7.87 7.98 7.67 58 Charcoal house

5--10 7.89 0.13 36 7.87 8.06 7.58 58 Houses

5--10 7.65 0.15 12 7.68 7.87 7.38 58 Mature forest

pH (KCl) 0--5 6.62 0.38 20 6.69 7.03 5.57 58 Baseball park

0--5 7.24 0.13 20 7.29 7.41 7.02 58 Farmlands

0--5 7.21 0.13 20 7.25 7.35 6.76 58 Charcoal house

0--5 7.28 0.13 31 7.28 7.53 7.02 58 Houses

0--5 7.00 0.13 9 7.01 7.21 6.85 58 Mature forest

5--10 6.64 0.37 24 6.81 7.01 5.89 58 Baseball park

5--10 7.23 0.14 24 7.28 7.40 6.97 58 Farmlands
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

490

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

5--10 7.25 0.08 24 7.26 7.36 7.03 58 Charcoal house

5--10 7.30 0.13 36 7.31 7.55 7.01 58 Houses

5--10 7.06 0.19 12 7.09 7.29 6.80 58 Mature forest

Phosphorus (mg/kg) 0--5 7 1 20 7 10 5 58 Baseball park

0--5 14 2 20 14 18 10 58 Farmlands

0--5 15 4 20 15 29 9 58 Charcoal house

0--5 21 5 31 20 33 11 58 Houses

0--5 17 7 9 19 28 6 58 Mature forest

5--10 6 1 24 6 8 4 58 Baseball park

5--10 13 3 24 13 18 7 58 Farmlands

5--10 13 3 24 12 21 8 58 Charcoal house

5--10 21 7 36 20 37 8 58 Houses

5--10 12 4 12 12 18 6 58 Mature forest

Potassium (cmol/kg-l) 0--5 0.91 0.21 20 0.88 1.38 0.43 58 Baseball park

0--5 1.05 0.29 20 1.07 1.54 0.15 58 Farmlands

0--5 0.90 0.18 20 0.85 1.45 0.71 58 Charcoal house

0--5 1.05 0.28 31 0.94 1.65 0.46 58 Houses

0--5 0.84 0.05 9 0.87 0.89 0.76 58 Mature forest

5--10 0.77 0.19 2 0.78 1.07 0.27 58 Baseball park

5--10 0.97 0.19 24 0.94 1.41 0.71 58 Farmlands

5--10 0.83 0.15 24 0.81 1.37 0.58 58 Charcoal house

5--10 0.97 0.21 36 0.91 1.56 0.70 58 Houses

5--10 0.80 0.09 12 0.82 0.93 0.64 58 Mature forest

Potassium (mg/g) 0--5 0.35 0.08 20 0.33 0.53 0.16 58 Baseball park

0--5 0.40 0.11 20 0.41 0.59 0.06 58 Farmlands

0--5 0.34 0.07 20 0.33 0.55 0.27 58 Charcoal house
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

491

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--5 0.40 0.11 31 0.36 0.63 0.18 58 Houses

0--5 0.32 0.02 9 0.33 0.34 0.29 58 Mature forest

5--10 0.29 0.07 24 0.30 0.41 0.10 58 Baseball park

5--10 0.37 0.07 24 0.36 0.54 0.27 58 Farmlands

5--10 0.32 0.06 24 0.31 0.52 0.22 58 Charcoal house

5--10 0.37 0.08 36 0.35 0.59 0.27 58 Houses

5--10 0.30 0.03 12 0.31 0.35 0.24 58 Mature forest

Sodium (cmol/kg-l) 0--5 0.23 0.05 20 0.25 0.32 0.15 58 Baseball park

0--5 0.20 0.09 20 0.17 0.38 0.08 58 Farmlands

0--5 0.22 0.07 20 0.20 0.36 0.13 58 Charcoal house

0--5 0.28 0.13 31 0.25 0.59 0.08 58 Houses

0--5 0.22 0.04 9 0.21 0.27 0.15 58 Mature forest

5--10 0.24 0.07 24 0.23 0.40 0.14 58 Baseball park

5--10 0.20 0.12 24 0.16 0.50 0.08 58 Farmlands

5--10 0.25 0.14 24 0.20 0.62 0.12 58 Charcoal house

5--10 0.24 0.10 36 0.22 0.55 0.08 58 Houses

5--10 0.22 0.06 12 0.21 0.36 0.15 58 Mature forest

Sodium (mg/kg) 0--5 54 12 20 57 74 34 58 Baseball park

0--5 45 20 20 39 88 19 58 Farmlands

0--5 51 16 20 46 83 31 58 Charcoal house

0--5 64 29 31 58 135 17 58 Houses

0--5 50 9 9 49 62 35 58 Mature forest

5--10 55 16 24 53 92 33 58 Baseball park

5--10 47 28 24 38 115 19 58 Farmlands

5--10 57 32 24 45 142 27 58 Charcoal house

5--10 56 24 36 50 125 20 58 Houses
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

492

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

5--10 50 14 12 48 82 34 58 Mature forest

Serpentinite Ultisols Lmrf Aluminum (cmol/kg-l) 0--15 0.17 0.10 10 0.16 0.37 0.04 59

Bulk density (g/cc) 0--15 0.84 0.10 10 0.83 1.04 0.71 59

Calcium (cmol/kg-l) 0--15 4.22 2.53 10 3.57 8.33 1.58 59

Calcium (mg/g) 0--15 0.84 0.51 10 0.71 1.67 0.32 59

Clay (%) 0--15 44 5 10 45 49 31 59

ECEC (cmol/kg-1) 0--15 19.47 13.11 10 14.91 36.51 4.40 59

Iron (mg/g) 0--15 0.24 0.07 10 0.23 0.37 0.13 59

Magnesium (cmol/kg-l) 0--15 14.57 12.48 10 10.71 33.90 2.22 59

Magnesium (mg/g) 0--15 1.75 1.50 10 1.29 4.07 0.27 59

Manganese (mg/g) 0--15 0.07 0.03 10 0.07 0.13 0.05 59

Nitrogen (%) 0--15 0.29 0.08 10 0.27 0.41 0.16 59

Organic matter (%) 0--15 7.80 1.48 10 0.67 9.99 5.69 59

pH (H2O) 0--15 5.30 0.40 10 5.40 5.90 4.80 59

pH (KCl) 0--15 5.90 5.00 10 6.00 6.60 5.30 59

Phosphorus (mg/kg) 0--15 9 2 10 10 12 6 59

Potassium (mg/g) 0--15 0.14 0.04 10 0.14 0.20 0.08 59

Potassium (cmol/kg-l) 0--15 0.36 0.10 10 0.36 0.52 0.21 59

Potassium (mg/g) 0--15 0.14 0.04 10 0.14 0.20 0.08 59

Sand (%) 0--15 24 6 10 23 40 17 59

Silt (%) 0--15 32 4 10 34 36 26 59

Sodium (cmol/kg-l) 0--15 0.15 0.06 10 0.17 0.27 0.08 59

Sodium (mg/g) 0--15 0.04 0.01 10 0.04 0.06 0.02 59

Alluvial deposits Mollisols Sdf Aluminum (cmol/kg-l) 0--10 0.39 0.46 241 0.18 2.19 0.01 60

Aluminum-total (mg/g) 0--10 19.34 13.61 391 15.10 58.17 1.18 60

Calcium (cmol/kg-l) 0--10 46.59 17.85 241 47.23 97.06 15.78 60
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

493

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Calcium (mg/g) 0--10 9.32 3.57 241 9.45 19.41 3.16 60

Calcium-total (mg/g) 0--10 143.81 97.56 391 114.23 346.58 1.91 60

Carbon (%) 0--10 19.60 8.59 343 16.46 46.07 8.45 60

ECEC (cmol/kg-1) 0--10 55.60 20.64 241 56.91 117.38 23.10 60

Iron (mg/g) 0--10 0.02 0.02 239 0.01 0.13 0.01 60

Iron-total (mg/g) 0--10 20.81 14.39 391 16.07 58.79 1.53 60

Loss on ignition (%) 0--10 32.79 16.22 415 30.50 80.02 11.47 60

Magnesium (cmol/kg-l) 0--10 6.60 3.13 241 6.17 19.72 1.70 60

Magnesium (mg/g) 0--10 0.79 0.38 241 0.74 2.37 0.20 60

Magnesium-total (mg/g) 0--10 4.47 1.87 391 4.42 14.06 0.21 60

Manganese (mg/g) 0--10 0.05 0.04 239 0.04 0.57 0.01 60

Manganese-total (mg/g) 0--10 0.41 0.26 391 0.32 1.14 0.04 60

Nitrogen (%) 0--10 1.30 0.65 343 1.25 3.14 0.36 60

pH (H2O) 0--10 7.87 0.21 412 7.93 8.25 7.04 60

pH (KCl) 0--10 7.26 0.20 212 7.29 7.72 6.30 60

Phosphorus (mg/g) 0--10 0.01 0.01 239 0.01 0.14 0.00 60

Phosphorus-total (mg/g) 0--10 0.65 0.25 391 0.65 1.42 0.03 60

Potassium (cmol/kg-l) 0--10 1.58 1.24 239 1.38 18.93 0.28 60

Potassium (mg/g) 0--10 0.60 0.47 239 0.53 7.19 0.11 60

Potassium-total (mg/g) 0--10 8.89 2.58 391 9.28 13.89 0.76 60

Sodium (cmol/kg-l) 0--10 0.46 0.28 237 0.42 1.24 0.01 60

Sodium (mg/g) 0--10 0.11 0.07 237 0.10 0.29 0.01 60

Sulfur (%) 0--10 0.17 0.10 343 0.16 0.45 0.02 60

Tuffaceous sandstone Ultisols Wfs Aluminum saturation (%) 0--10 61.69 14.46 3 58.11 77.60 49.35 61 Bisley

0--10 49.46 18.38 5 51.63 66.23 20.02 61 Dwarf forest

0--10 48.41 23.87 6 43.68 82.52 24.17 61 El Verde
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

494

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--10 29.99 27.76 11 21.12 100.00 6.70 61 Montane palm forest

0--10 44.81 18.83 15 40.49 77.64 21.78 61 Palo Colorado forest

0--10 31.45 25.82 15 15.77 85.34 5.25 61 Tabonuco forest

10--20 56.62 17.99 3 49.52 77.08 43.26 61 Bisley

10--20 63.32 9.43 6 61.55 77.38 52.21 61 Dwarf forest

10--20 61.31 28.33 6 67.93 92.74 24.30 61 El Verde

10--20 41.43 30.92 8 38.23 10.00 4.01 61 Montane palm forest

10--20 5.47 21.32 16 58.26 86.87 10.16 61 Palo Colorado forest

10--20 30.52 26.43 17 21.51 82.76 3.64 61 Tabonuco forest

20--30 57.48 16.54 3 54.67 73.92 40.84 61 Bisley

20--30 67.14 9.58 5 65.46 80.15 57.93 61 Dwarf forest

20--30 57.11 31.53 6 66.44 84.77 16.09 61 El Verde

20--30 40.45 30.12 7 34.44 100.00 6.58 61 Montane palm forest

20--30 58.11 19.68 15 53.97 85.52 23.67 61 Palo Colorado forest

20--30 31.55 29.84 15 17.45 85.58 3.10 61 Tabonuco forest

30--40 64.48 14.61 2 64.48 74.81 54.15 61 Bisley

30--40 70.60 7.89 6 69.60 82.90 62.72 61 Dwarf forest

30--40 65.53 36.04 5 88.02 89.64 6.46 61 El Verde

30--40 40.74 31.82 7 6.20 8.81 2.63 61 Montane palm forest

30--40 53.69 26.20 15 52.89 90.88 7.37 61 Palo Colorado forest

30--40 23.12 24.26 16 10.87 66.41 2.53 61 Tabonuco forest

Aluminum (cmol/kg-l) 0--10 2.12 0.67 3 1.75 2.90 1.72 61 Bisley

0--10 5.01 1.95 5 6.20 6.59 2.02 61 Dwarf forest

0--10 4.40 3.23 6 3.75 9.34 1.06 61 El Verde

0--10 1.60 0.89 11 1.56 3.12 0.72 61 Montane palm forest

0--10 2.08 1.14 15 1.57 4.46 1.00 61 Palo Colorado forest
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

495

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--10 3.28 3.02 15 2.01 12.80 0.77 61 Tabonuco forest

10--20 1.55 0.68 3 1.21 2.34 1.11 61 Bisley

10--20 4.23 1.27 6 4.72 5.65 2.65 61 Dwarf forest

10--20 4.86 3.92 6 4.35 11.14 0.93 61 El Verde

10--20 1.76 0.85 8 1.83 2.80 0.36 61 Montane palm forest

10--20 1.88 1.14 16 1.53 4.08 0.20 61 Palo Colorado forest

10--20 3.76 3.73 17 2.67 11.55 0.40 61 Tabonuco forest

20--30 1.49 0.37 3 1.36 1.91 1.20 61 Bisley

20--30 4.09 1.21 5 3.87 6.14 2.89 61 Dwarf forest

20--30 4.65 4.16 6 4.03 11.33 0.58 61 El Verde

20--30 1.82 0.85 7 1.77 2.81 0.49 61 Montane palm forest

20--30 1.95 1.36 15 1.49 4.38 0.43 61 Palo Colorado forest

20--30 2.83 2.38 15 2.21 8.89 0.36 61 Tabonuco forest

30--40 1.93 1.15 2 1.93 2.74 1.11 61 Bisley

30--40 3.86 0.66 6 3.83 5.01 3.08 61 Dwarf forest

30--40 5.33 4.88 5 5.79 12.67 0.19 61 El Verde

30--40 1.84 1.07 5 0.25 0.47 0.18 61 Montane palm forest

30--40 1.55 1.24 15 1.38 4.01 0.05 61 Palo Colorado forest

30--40 2.34 2.21 16 1.79 8.59 0.29 61 Tabonuco forest

Calcium (cmol/kg-l) 0--10 0.50 0.30 3 0.44 0.89 0.33 61 Bisley

0--10 2.51 1.80 5 2.38 5.20 0.89 61 Dwarf forest

0--10 1.69 0.84 6 1.71 2.87 0.39 61 El Verde

0--10 3.43 2.25 10 2.68 7.48 1.37 61 Montane palm forest

0--10 1.29 0.75 15 1.44 2.50 0.27 61 Palo Colorado forest

0--10 7.62 7.68 15 6.64 27.10 0.26 61 Tabonuco forest

10--20 0.46 0.26 3 0.35 0.76 0.27 61 Bisley
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

496

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--20 0.70 0.27 6 0.70 1.11 0.29 61 Dwarf forest

10--20 0.74 0.67 6 0.60 1.80 0.05 61 El Verde

10--20 2.59 1.98 7 2.20 6.36 0.89 61 Montane palm forest

10--20 0.52 0.34 16 0.48 1.30 0.08 61 Palo Colorado forest

10--20 8.02 7.43 17 7.39 23.14 0.40 61 Tabonuco forest

20--30 0.44 0.31 3 0.34 0.78 0.19 61 Bisley

20--30 0.57 0.23 5 0.58 0.85 0.23 61 Dwarf forest

20--30 0.85 0.69 6 0.66 1.77 0.18 61 El Verde

20--30 2.90 1.60 6 2.53 4.84 0.76 61 Montane palm forest

20--30 0.45 0.36 15 0.35 1.28 0.05 61 Palo Colorado forest

20--30 7.98 7.40 15 7.59 24.83 0.30 61 Tabonuco forest

30--40 0.45 0.21 2 0.45 0.59 0.30 61 Bisley

30--40 0.40 0.18 6 0.45 0.55 0.12 61 Dwarf forest

30--40 0.42 0.51 5 0.19 1.30 0.09 61 El Verde

30--40 2.76 1.90 6 2.50 6.22 0.49 61 Montane palm forest

30--40 0.24 0.18 15 0.20 0.70 0.01 61 Palo Colorado forest

30--40 8.38 7.32 17 7.47 25.04 0.69 61 Tabonuco forest

Calcium (mg/g) 0--10 0.11 0.06 3 0.09 0.18 0.07 61 Bisley

0--10 0.50 0.36 5 0.48 1.04 0.18 61 Dwarf forest

0--10 0.34 0.17 6 0.34 0.57 0.08 61 El Verde

0--10 0.69 0.45 10 0.54 1.50 0.27 61 Montane palm forest

0--10 0.26 0.15 15 0.29 0.50 0.06 61 Palo Colorado forest

0--10 1.53 1.54 15 1.33 5.42 0.05 61 Tabonuco forest

10--20 0.09 0.05 3 0.07 0.15 0.05 61 Bisley

10--20 0.14 0.05 6 0.14 0.22 0.06 61 Dwarf forest

10--20 0.15 0.13 6 0.12 0.36 0.01 61 El Verde
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

497

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--20 0.52 0.40 7 0.44 1.27 0.18 61 Montane palm forest

10--20 0.10 0.07 16 0.09 0.26 0.02 61 Palo Colorado forest

10--20 1.60 1.49 17 1.48 4.63 0.08 61 Tabonuco forest

20--30 0.09 0.06 3 0.07 0.16 0.04 61 Bisley

20--30 0.11 0.05 5 0.12 0.17 0.05 61 Dwarf forest

20--30 0.17 0.14 6 0.13 0.35 0.04 61 El Verde

20--30 0.58 0.32 6 0.51 0.97 0.15 61 Montane palm forest

20--30 0.09 0.07 15 0.07 0.26 0.01 61 Palo Colorado forest

20--30 1.60 1.48 15 1.52 4.97 0.06 61 Tabonuco forest

30--40 0.09 0.04 2 0.09 0.12 0.06 61 Bisley

30--40 0.08 0.04 6 0.09 0.11 0.02 61 Dwarf forest

30--40 0.08 0.10 5 0.04 0.26 0.02 61 El Verde

30--40 0.55 0.38 6 0.50 1.24 0.10 61 Montane palm forest

30--40 0.05 0.04 15 0.04 0.14 0.01 61 Palo Colorado forest

30--40 1.68 1.65 17 1.49 5.01 0.14 61 Tabonuco forest

Effective cation exchange capacity 0--10 3.41 0.37 3 3.49 3.74 3.01 61 Bisley

{Ca+K+Mg(cmol/kg-1) 0--10 10.16 1.54 5 9.82 12.76 8.76 61 Dwarf forest

0--10 8.14 2.76 6 9.09 10.92 4.38 61 El Verde

0--10 7.33 3.14 11 7.27 12.78 1.56 61 Montane palm forest

0--10 4.75 1.85 15 4.29 9.41 2.52 61 Palo Colorado forest

0--10 16.37 14.15 15 13.55 54.02 3.02 61 Tabonuco forest

10--20 2.69 0.41 3 2.80 3.04 2.24 61 Bisley

10--20 6.60 1.45 6 6.99 8.05 4.68 61 Dwarf forest

10--20 6.83 3.12 6 6.20 12.01 3.83 61 El Verde

10--20 5.57 2.32 8 5.34 8.99 1.95 61 Montane palm forest

10--20 3.29 1.34 16 3.05 5.35 1.41 61 Palo Colorado forest
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

498

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--20 16.81 15.13 17 11.18 51.24 3.94 61 Tabonuco forest

20--30 2.63 0.29 3 2.58 2.94 2.36 61 Bisley

20--30 6.09 1.47 5 5.62 8.36 4.83 61 Dwarf forest

20--30 6.85 3.80 6 5.69 13.77 3.61 61 El Verde

20--30 5.84 2.82 7 6.56 9.77 1.77 61 Montane palm forest

20--30 3.14 1.62 15 2.53 6.93 1.24 61 Palo Colorado forest

20--30 15.02 12.74 15 13.22 50.94 3.16 61 Tabonuco forest

30--40 2.86 1.14 2 2.86 3.66 2.05 61 Bisley

30--40 5.55 1.16 6 6.01 6.82 3.72 61 Dwarf forest

30--40 6.76 4.58 5 6.58 14.13 2.91 61 El Verde

30--40 5.61 2.19 7 1.99 3.37 0.20 61 Montane palm forest

30--40 2.42 1.31 15 2.25 4.64 0.68 61 Palo Colorado forest

30--40 15.13 12.26 17 10.72 5.37 1.41 61 Tabonuco forest

Iron (mg/g) 0--10 2.16 0.31 5 2.33 2.47 1.73 61 Dwarf forest

0--10 1.68 0.37 5 1.73 2.09 1.14 61 El Verde

0--10 0.57 0.31 8 0.45 0.99 0.25 61 Montane palm forest

0--10 1.24 0.63 10 1.00 2.48 0.48 61 Palo Colorado forest

0--10 1.55 1.04 11 1.67 3.39 0.30 61 Tabonuco forest

10--20 1.62 0.48 6 1.64 2.11 0.76 61 Dwarf forest

10--20 1.37 0.67 5 1.47 2.09 0.33 61 El Verde

10--20 0.45 0.33 5 0.40 0.98 0.13 61 Montane palm forest

10--20 0.83 0.54 11 0.87 1.94 0.13 61 Palo Colorado forest

10--20 1.47 1.28 13 1.38 4.19 0.30 61 Tabonuco forest

20--30 1.37 0.91 5 1.77 2.26 0.08 61 Dwarf forest

20--30 0.82 0.53 5 0.66 1.69 0.27 61 El Verde

20--30 0.40 0.34 5 0.31 0.99 0.13 61 Montane palm forest
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

499

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

20--30 0.79 0.59 11 0.75 2.24 0.13 61 Palo Colorado forest

20--30 1.74 1.21 10 1.60 3.69 0.28 61 Tabonuco forest

30--40 0.70 0.69 6 0.46 1.63 0.08 61 Dwarf forest

30--40 0.43 0.47 5 0.27 1.24 0.07 61 El Verde

30--40 0.42 0.46 5 0.22 1.23 0.13 61 Montane palm forest

30--40 0.39 0.35 10 0.19 0.93 0.05 61 Palo Colorado forest

30--40 1.66 1.17 12 1.52 3.89 0.29 61 Tabonuco forest

Loss on ignition (%) 0--10 20.23 4.21 28 20.35 34.62 14.35 61 Bisley

0--10 25.61 7.47 17 23.92 38.74 15.76 61 Dwarf forest

0--10 22.24 2.88 33 22.13 30.71 15.31 61 El Verde

0--10 20.22 4.85 15 18.60 28.60 14.76 61 Montane palm forest

0--10 10.40 4.18 19 9.80 19.74 6.11 61 Palo Colorado forest

0--10 20.97 6.78 22 19.06 37.59 12.89 61 Tabonuco forest

0--20 22.05 3.27 7 23.65 24.83 16.80 61 Jimenez, Caribbean National Forest

0--20 14.06 3.68 7 13.45 19.18 10.19 61 Road 191

10--20 18.08 2.56 30 18.32 22.03 12.89 61 Bisley

10--20 23.21 7.96 17 25.52 34.09 11.98 61 Dwarf forest

10--20 20.16 1.90 32 20.10 24.32 15.47 61 El Verde

10--20 17.66 3.38 13 17.40 26.28 11.90 61 Montane palm forest

10--20 9.87 4.95 21 8.30 19.10 3.81 61 Palo Colorado forest

10--20 19.42 6.41 23 18.63 35.48 10.33 61 Tabonuco forest

20--30 16.80 2.66 28 16.33 21.12 10.84 61 Bisley

20--30 20.12 7.43 17 20.20 31.82 10.44 61 Dwarf forest

20--30 18.99 1.59 32 19.15 22.13 13.50 61 El Verde

20--30 17.10 4.18 13 16.39 26.74 9.41 61 Montane palm forest

20--30 9.20 3.82 22 7.90 17.00 3.52 61 Palo Colorado forest
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

500

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

20--30 23.40 22.60 20 19.53 116.41 9.13 61 Tabonuco forest

20--40 19.33 3.09 6 20.22 22.61 15.34 61 Jimenez, Caribbean National Forest

20--40 12.91 3.70 6 12.53 17.68 8.78 61 Road 191

30--40 17.57 2.58 28 17.76 24.18 10.66 61 Bisley

30--40 17.45 6.09 17 16.67 28.46 0.57 61 Dwarf forest

30--40 18.75 1.73 26 19.09 23.22 14.92 61 El Verde

30--40 16.89 4.42 14 16.32 28.48 9.45 61 Montane palm forest

30--40 8.58 3.27 21 7.92 16.43 3.19 61 Palo Colorado forest

30--40 17.09 5.94 20 17.73 26.85 6.22 61 Tabonuco forest

40--50 17.17 2.23 7 16.92 21.04 13.93 61 El Verde

40--50 17.70 2.43 4 18.81 19.96 14.41 61 Jimenez, Caribbean National Forest

40--50 10.99 1.92 5 11.89 12.73 8.65 61 Road 191

Magnesium (cmol/kg-l) 0--10 0.45 0.15 3 0.49 0.58 0.28 61 Bisley

0--10 1.34 0.25 5 1.18 1.61 1.14 61 Dwarf forest

0--10 1.47 0.94 6 1.35 2.63 0.21 61 El Verde

0--10 1.87 0.68 10 2.08 2.88 0.61 61 Montane palm forest

0--10 0.87 0.57 15 0.67 1.87 0.26 61 Palo Colorado forest

0--10 4.58 5.09 15 2.97 20.04 0.35 61 Tabonuco forest

10--20 0.44 0.14 3 0.45 0.58 0.30 61 Bisley

10--20 0.77 0.21 6 0.74 1.14 0.49 61 Dwarf forest

10--20 0.84 0.38 6 0.86 1.27 0.17 61 El Verde

10--20 1.13 0.44 7 1.20 1.53 0.32 61 Montane palm forest

10--20 0.46 0.35 16 0.36 1.27 0.13 61 Palo Colorado forest

10--20 4.13 4.45 17 2.18 14.91 0.41 61 Tabonuco forest

20--30 0.44 0.23 3 0.38 0.69 0.24 61 Bisley

20--30 0.72 0.26 5 0.77 1.02 0.31 61 Dwarf forest
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

501

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

20--30 1.03 0.26 6 0.93 1.49 0.76 61 El Verde

20--30 14.14 0.42 6 1.26 1.55 0.34 61 Montane palm forest

20--30 0.39 0.27 15 0.25 1.01 0.12 61 Palo Colorado forest

20--30 3.40 3.75 15 2.23 14.65 0.30 61 Tabonuco forest

30--40 0.24 0.15 2 0.24 0.34 0.13 61 Bisley

30--40 0.60 0.27 6 0.65 0.87 0.20 61 Dwarf forest

30--40 0.77 0.11 5 0.83 0.86 0.63 61 El Verde

30--40 1.03 0.42 6 1.07 1.49 0.35 61 Montane palm forest

30--40 0.35 0.22 15 0.31 0.97 0.09 61 Palo Colorado forest

30--40 3.63 3.49 17 2.51 13.72 0.52 61 Tabonuco forest

Magnesium (mg/g) 0--10 0.05 0.02 3 0.06 0.07 0.03 61 Bisley

0--10 0.16 0.03 5 0.14 0.19 0.14 61 Dwarf forest

0--10 0.18 0.11 6 0.16 0.32 0.03 61 El Verde

0--10 0.22 0.08 10 0.25 0.35 0.07 61 Montane palm forest

0--10 0.10 0.07 15 0.08 0.22 0.03 61 Palo Colorado forest

0--10 0.55 0.61 15 0.36 2.41 0.04 61 Tabonuco forest

10--20 0.05 0.02 3 0.05 0.07 0.04 61 Bisley

10--20 0.09 0.03 6 0.09 0.14 0.06 61 Dwarf forest

10--20 0.10 0.05 6 0.10 0.15 0.02 61 El Verde

10--20 0.14 0.05 7 0.14 0.18 0.04 61 Montane palm forest

10--20 0.06 0.04 16 0.04 0.15 0.02 61 Palo Colorado forest

10--20 0.50 0.54 17 0.26 1.79 0.05 61 Tabonuco forest

20--30 0.05 0.04 3 0.05 0.08 0.03 61 Bisley

20--30 0.09 0.03 5 0.09 0.12 0.04 61 Dwarf forest

20--30 0.12 0.03 6 0.11 0.18 0.09 61 El Verde

20--30 0.14 0.05 6 0.15 0.19 0.04 61 Montane palm forest
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

502

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

20--30 0.05 0.03 15 0.03 0.12 0.01 61 Palo Colorado forest

20--30 0.41 0.45 15 0.27 1.76 0.04 61 Tabonuco forest

30--40 0.03 0.02 2 0.03 0.04 0.02 61 Bisley

30--40 0.07 0.03 6 0.08 0.11 0.03 61 Dwarf forest

30--40 0.09 0.01 5 0.10 0.10 0.08 61 El Verde

30--40 0.12 0.05 6 0.13 0.18 0.04 61 Montane palm forest

30--40 0.04 0.03 15 0.04 0.12 0.01 61 Palo Colorado forest

30--40 0.44 0.42 17 0.30 1.65 0.06 61 Tabonuco forest

Manganese (mg/g) 0--10 0.02 0.01 5 0.01 0.02 0.01 61 Dwarf forest

0--10 0.08 0.08 5 0.05 0.20 0.01 61 El Verde

0--10 0.32 0.15 8 0.25 0.52 0.19 61 Montane palm forest

0--10 0.03 0.02 10 0.03 0.05 0.01 61 Palo Colorado forest

0--10 0.23 0.22 11 0.13 0.63 0.02 61 Tabonuco forest

10--20 0.01 0.00 6 0.01 0.01 0.01 61 Dwarf forest

10--20 0.07 0.10 5 0.02 0.23 0.01 61 El Verde

10--20 0.23 0.07 5 0.19 0.30 0.15 61 Montane palm forest

10--20 0.01 0.01 11 0.01 0.03 0.01 61 Palo Colorado forest

10--20 0.24 0.27 13 0.11 0.83 0.01 61 Tabonuco forest

20--30 0.05 0.11 5 0.01 0.25 0.01 61 Dwarf forest

20--30 0.09 0.11 5 0.04 0.27 0.01 61 El Verde

20--30 0.18 0.06 5 0.21 0.23 0.09 61 Montane palm forest

20--30 0.01 0.01 11 0.01 0.03 0.01 61 Palo Colorado forest

20--30 0.32 0.23 10 0.32 0.64 0.06 61 Tabonuco forest

30--40 0.10 0.14 6 0.01 0.30 0.01 61 Dwarf forest

30--40 0.09 0.12 5 0.05 0.30 0.01 61 El Verde

30--40 0.15 0.07 5 0.14 0.21 0.05 61 Montane palm forest
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

503

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

30--40 0.01 0.01 10 0.01 0.03 0.01 61 Palo Colorado forest

30--40 0.27 0.15 12 0.29 0.48 0.05 61 Tabonuco forest

Nitrogen (%) 0--10 0.51 0.17 5 0.58 0.69 0.30 61 Dwarf forest

0--10 0.23 0.09 6 0.23 0.37 0.14 61 Montane palm forest

0--10 0.18 0.09 5 0.15 0.29 0.09 61 Palo Colorado forest

0--10 0.37 0.16 5 0.34 0.62 0.22 61 Tabonuco forest

10--20 0.37 0.18 6 0.41 0.54 0.15 61 Dwarf forest

10--20 0.15 0.10 5 0.13 0.33 0.06 61 Montane palm forest

10--20 0.13 0.09 5 0.08 0.25 0.05 61 Palo Colorado forest

10--20 0.30 0.08 6 0.31 0.41 0.20 61 Tabonuco forest

20--30 0.26 0.20 5 0.19 0.53 0.08 61 Dwarf forest

20--30 0.15 0.11 5 0.12 0.33 0.06 61 Montane palm forest

20--30 0.13 0.09 6 0.11 0.23 0.05 61 Palo Colorado forest

20--30 0.29 0.09 5 0.32 0.40 0.20 61 Tabonuco forest

30--40 0.14 0.16 6 0.07 0.42 0.02 61 Dwarf forest

30--40 0.14 0.11 5 0.10 0.33 0.06 61 Montane palm forest

30--40 0.07 0.05 6 0.04 0.14 0.02 61 Palo Colorado forest

30--40 0.24 0.11 6 0.23 0.36 0.09 61 Tabonuco forest

Organic matter (%) 0--10 6.33 2.44 79 5.90 17.57 1.86 61 Bisley

0--10 17.70 8.64 27 19.95 33.51 3.46 61 Dwarf forest

0--10 7.59 2.11 79 7.27 15.46 4.10 61 El Verde

0--10 6.36 3.65 24 5.59 15.32 2.17 61 Montane palm forest

0--10 6.68 3.52 25 5.53 13.65 2.19 61 Palo Colorado forest

0--10 9.21 5.94 27 6.54 29.03 3.71 61 Tabonuco forest

0--20 5.48 2.01 13 6.36 7.54 1.56 61 Jimenez, Caribbean National Forest

0--20 6.90 2.88 12 5.49 12.68 4.09 61 Road 191
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

504

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--20 4.46 1.67 78 4.44 14.63 0.73 61 Bisley

10--20 15.15 8.83 27 16.15 31.65 0.32 61 Dwarf forest

10--20 5.44 1.61 82 5.12 10.35 2.50 61 El Verde

10--20 4.27 3.17 22 3.41 11.50 0.62 61 Montane palm forest

10--20 5.12 3.19 25 4.22 11.85 1.07 61 Palo Colorado forest

10--20 7.68 4.67 27 7.14 17.76 2.59 61 Tabonuco forest

20--30 3.41 1.02 73 3.65 5.11 1.00 61 Bisley

20--30 12.51 8.53 27 11.79 26.42 0.47 61 Dwarf forest

20--30 4.36 1.43 77 4.03 7.82 1.52 61 El Verde

20--30 3.54 2.63 23 3.01 11.26 0.25 61 Montane palm forest

20--30 4.49 3.27 27 3.19 12.85 1.11 61 Palo Colorado forest

20--30 6.50 3.23 24 6.05 14.06 1.75 61 Tabonuco forest

20--40 3.46 1.34 13 3.96 5.14 0.93 61 Jimenez, Caribbean National Forest

20--40 4.58 2.75 11 3.73 10.53 1.87 61 Road 191

30--40 2.54 1.06 76 2.53 5.28 0.72 61 Bisley

30--40 8.63 7.14 27 5.80 22.94 0.28 61 Dwarf forest

30--40 3.51 1.66 63 3.26 10.63 0.74 61 El Verde

30--40 3.10 2.61 23 2.63 12.02 0.56 61 Montane palm forest

30--40 3.05 2.66 25 1.82 8.55 0.66 61 Palo Colorado forest

30--40 5.26 3.27 230 4.43 11.91 1.40 61 Tabonuco forest

40--50 2.66 0.71 10 2.57 4.04 1.47 61 El Verde

40--50 2.59 1.26 10 2.51 5.39 0.67 61 Jimenez, Caribbean National Forest

40--50 2.82 1.42 11 2.50 5.88 1.20 61 Road 191

pH (H2O) 0--10 4.85 0.29 83 4.80 5.88 3.92 61 Bisley

0--10 4.65 0.16 27 4.67 5.00 4.31 61 Dwarf forest

0--10 4.92 0.25 75 4.92 5.52 4.31 61 El Verde
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

505

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--10 5.05 0.20 21 5.08 5.41 4.62 61 Montane palm forest

0--10 4.60 0.24 25 4.67 4.94 3.93 61 Palo Colorado forest

0--10 4.94 0.25 26 4.90 5.85 4.61 61 Tabonuco forest

0--20 4.80 0.13 13 4.78 5.05 4.65 61 Jimenez, Caribbean National Forest

0--20 4.67 0.24 12 4.67 5.04 4.28 61 Road 191

10--20 4.90 0.26 83 4.88 5.96 4.29 61 Bisley

10--20 4.72 0.21 27 4.72 5.25 4.28 61 Dwarf forest

10--20 4.97 0.19 79 5.00 5.44 4.48 61 El Verde

10--20 4.98 0.23 20 5.00 5.37 4.60 61 Montane palm forest

10--20 4.67 0.25 25 4.72 5.05 4.15 61 Palo Colorado forest

10--20 4.90 0.30 27 4.83 5.53 4.41 61 Tabonuco forest

20--30 4.93 0.22 75 4.92 5.45 4.37 61 Bisley

20--30 4.79 0.18 27 4.78 5.11 4.43 61 Dwarf forest

20--30 5.00 0.21 73 5.02 5.62 4.47 61 El Verde

20--30 5.03 0.17 20 5.04 5.35 4.74 61 Montane palm forest

20--30 4.72 0.31 27 4.70 5.23 4.03 61 Palo Colorado forest

20--30 4.98 0.20 24 4.99 5.33 4.52 61 Tabonuco forest

20--40 4.93 0.15 12 4.95 5.12 4.63 61 Jimenez, Caribbean National Forest

20--40 5.13 1.46 12 4.72 9.71 4.30 61 Road 191

30--40 5.01 0.23 79 5.01 5.48 4.39 61 Bisley

30--40 4.77 0.22 27 4.79 5.22 4.26 61 Dwarf forest

30--40 5.05 0.21 63 5.02 5.78 4.65 61 El Verde

30--40 4.99 0.20 22 4.97 5.43 4.62 61 Montane palm forest

30--40 4.81 0.36 25 4.84 5.68 4.17 61 Palo Colorado forest

30--40 5.10 0.30 23 5.14 4.66 4.49 61 Tabonuco forest

40--50 4.69 0.26 9 4.68 5.14 4.28 61 El Verde
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

506

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

40--50 5.04 0.21 10 5.05 5.36 4.71 61 Jimenez, Caribbean National Forest

40--50 4.85 0.19 11 4.89 5.11 4.59 61 Road 191

pH (KCl) 0--10 4.04 0.27 83 3.98 5.20 3.15 61 Bisley

0--10 3.85 0.12 27 3.86 4.21 3.59 61 Dwarf forest

0--10 4.07 0.24 75 4.04 4.73 3.52 61 El Verde

0--10 0.10 0.22 21 4.15 4.55 3.64 61 Montane palm forest

0--10 3.89 0.19 25 3.95 4.18 3.42 61 Palo Colorado forest

0--10 3.91 0.21 26 3.93 4.48 3.43 61 Tabonuco forest

0--20 3.89 0.13 13 3.95 4.06 3.69 61 Jimenez, Caribbean National Forest

0--20 4.01 0.13 12 4.05 4.16 3.83 61 Road 191

10--20 4.08 0.23 83 4.05 5.19 3.64 61 Bisley

10--20 3.89 0.08 27 3.90 4.06 3.74 61 Dwarf forest

10--20 4.07 0.20 79 4.04 4.47 3.58 61 El Verde

10--20 4.02 0.17 20 4.01 4.42 3.74 61 Montane palm forest

10--20 4.02 0.22 25 4.03 4.63 3.57 61 Palo Colorado forest

10--20 3.90 0.24 27 3.91 4.31 3.48 61 Tabonuco forest

20--30 4.10 0.20 75 4.08 4.62 3.62 61 Bisley

20--30 3.92 0.09 27 3.93 4.07 3.77 61 Dwarf forest

20--30 4.06 0.20 73 4.04 4.67 3.62 61 El Verde

20--30 4.07 0.16 20 4.09 4.40 3.76 61 Montane palm forest

20--30 4.05 0.22 27 4.06 4.47 3.55 61 Palo Colorado forest

20--30 3.92 0.19 24 3.91 4.28 3.51 61 Tabonuco forest

20--40 4.21 0.80 12 4.05 6.68 3.65 61 Jimenez, Caribbean National Forest

20--40 4.05 0.13 12 4.02 4.26 3.88 61 Road 191

30--40 4.12 0.23 79 4.10 4.67 3.61 61 Bisley

30--40 3.91 0.12 27 3.91 4.26 3.61 61 Dwarf forest
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

507

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

30--40 4.08 0.23 63 4.02 4.99 3.68 61 El Verde

30--40 4.07 0.18 22 4.09 4.54 3.78 61 Montane palm forest

30--40 4.22 0.44 25 4.11 5.38 3.62 61 Palo Colorado forest

30--40 3.96 0.20 23 3.91 4.37 3.52 61 Tabonuco forest

40--50 3.93 0.11 9 3.91 4.08 3.82 61 El Verde

40--50 4.09 0.27 10 4.18 4.49 3.65 61 Jimenez, Caribbean National Forest

40--50 4.11 0.22 11 4.04 4.63 3.94 61 Road 191

Phosphorus (mg/kg) 0--10 13 4 3 12 18 10 61 Bisley

0--10 25 12 5 29 40 9 61 Dwarf forest

0--10 22 6 6 21 32 15 61 El Verde

0--10 12 4 8 12 19 7 61 Montane palm forest

0--10 16 14 10 11 44 2 61 Palo Colorado forest

0--10 25 6 11 26 34 13 61 Tabonuco forest

10--20 7 1 3 6 8 6 61 Bisley

10--20 18 15 6 19 37 2 61 Dwarf forest

10--20 9 5 6 9 18 4 61 El Verde

10--20 8 6 5 4 18 4 61 Montane palm forest

10--20 10 8 9 7 25 1 61 Palo Colorado forest

10--20 23 9 13 25 38 10 61 Tabonuco forest

20--30 4 1 3 4 4 3 61 Bisley

20--30 17 17 4 15 36 1 61 Dwarf forest

20--30 6 4 6 5 12 2 61 El Verde

20--30 5 5 5 5 16 3 61 Montane palm forest

20--30 14 9 7 11 24 2 61 Palo Colorado forest

20--30 21 9 10 19 35 10 61 Tabonuco forest

30--40 2 1 2 2 2 1 61 Bisley
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

508

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

30--40 33 16 2 33 44 21 61 Dwarf forest

30--40 2 1 4 2 3 1 61 El Verde

30--40 6 5 5 6 14 2 61 Montane palm forest

30--40 6 3 5 6 9 2 61 Palo Colorado forest

30--40 18 8 12 17 31 5 61 Tabonuco forest

Potassium (cmol/kg-l) 0--10 0.48 0.05 5 0.50 0.53 0.40 61 Dwarf forest

0--10 0.43 0.08 5 0.41 0.53 0.32 61 El Verde

0--10 0.53 0.25 8 0.61 0.75 0.15 61 Montane palm forest

0--10 0.36 0.18 10 0.37 0.71 0.13 61 Palo Colorado forest

0--10 0.49 0.28 11 0.38 1.02 0.19 61 Tabonuco forest

10--20 0.29 0.12 6 0.28 0.42 0.16 61 Dwarf forest

10--20 0.23 0.15 5 0.21 0.48 0.09 61 El Verde

10--20 0.33 0.11 5 0.30 0.47 0.21 61 Montane palm forest

10--20 0.26 0.15 11 0.26 0.49 0.07 61 Palo Colorado forest

10--20 0.35 0.26 13 0.24 0.86 0.07 61 Tabonuco forest

20--30 0.19 0.10 5 0.14 0.30 0.08 61 Dwarf forest

20--30 0.16 0.08 5 0.12 0.30 0.10 61 El Verde

20--30 0.27 0.09 5 0.25 0.42 0.20 61 Montane palm forest

20--30 0.24 0.15 11 0.26 0.48 0.06 61 Palo Colorado forest

20--30 0.37 0.24 10 0.34 0.75 0.07 61 Tabonuco forest

30--40 0.16 0.08 6 0.16 0.26 0.07 61 Dwarf forest

30--40 0.10 0.09 5 0.07 0.25 0.04 61 El Verde

30--40 0.29 0.13 6 0.39 0.52 0.20 61 Montane palm forest

30--40 0.15 0.12 10 0.12 0.37 0.04 61 Palo Colorado forest

30--40 0.38 0.16 12 0.36 0.67 0.05 61 Tabonuco forest

Potassium (mg/g) 0--10 0.18 0.02 5 0.19 0.20 0.15 61 Dwarf forest
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

509

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--10 0.16 0.03 5 0.16 0.20 0.12 61 El Verde

0--10 0.20 0.09 8 0.23 0.28 0.06 61 Montane palm forest

0--10 0.14 0.07 10 0.14 0.27 0.05 61 Palo Colorado forest

0--10 0.19 0.11 11 0.15 0.39 0.07 61 Tabonuco forest

10--20 0.11 0.04 6 0.11 0.16 0.06 61 Dwarf forest

10--20 0.09 0.06 5 0.08 0.18 0.04 61 El Verde

10--20 0.12 0.04 5 0.11 0.18 0.08 61 Montane palm forest

10--20 0.10 0.06 11 0.10 0.19 0.03 61 Palo Colorado forest

10--20 0.13 0.10 13 0.09 0.33 0.03 61 Tabonuco forest

20--30 0.07 0.04 5 0.06 0.12 0.03 61 Dwarf forest

20--30 0.06 0.03 5 0.05 0.12 0.04 61 El Verde

20--30 0.10 0.03 5 0.10 0.16 0.08 61 Montane palm forest

20--30 0.09 0.06 11 0.10 0.18 0.02 61 Palo Colorado forest

20--30 0.14 0.09 10 0.13 0.29 0.03 61 Tabonuco forest

30--40 0.06 0.03 6 0.06 0.10 0.03 61 Dwarf forest

30--40 0.04 0.03 5 0.03 0.09 0.02 61 El Verde

30--40 0.11 0.05 5 0.09 0.18 0.07 61 Montane palm forest

30--40 0.06 0.05 10 0.04 0.14 0.02 61 Palo Colorado forest

30--40 0.15 0.06 12 0.14 0.26 0.02 61 Tabonuco forest

Sodium (cmol/kg-l) 0--10 0.28 0.06 3 0.30 0.33 0.22 61 Bisley

0--10 0.83 0.13 5 0.78 1.03 0.70 61 Dwarf forest

0--10 0.34 0.23 4 0.43 0.49 0.01 61 El Verde

0--10 0.59 0.16 10 0.60 0.84 0.30 61 Montane palm forest

0--10 0.28 0.13 15 0.26 0.66 0.13 61 Palo Colorado forest

0--10 0.67 0.36 12 0.67 1.35 0.14 61 Tabonuco forest

10--20 0.24 0.11 3 0.25 0.34 0.13 61 Bisley
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

510

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--20 0.60 0.15 6 0.64 0.74 0.41 61 Dwarf forest

10--20 0.30 0.21 4 0.38 0.44 0.01 61 El Verde

10--20 0.40 0.13 7 0.39 0.57 0.16 61 Montane palm forest

10--20 0.25 0.16 16 0.19 0.66 0.10 61 Palo Colorado forest

10--20 0.78 0.69 14 0.57 2.44 0.12 61 Tabonuco forest

20--30 0.26 0.01 3 0.27 0.27 0.25 61 Bisley

20--30 0.52 0.15 5 0.61 0.65 0.29 61 Dwarf forest

20--30 0.39 0.07 3 0.35 0.47 0.34 61 El Verde

20--30 0.43 0.12 6 0.44 0.57 0.22 61 Montane palm forest

20--30 0.20 0.09 15 0.18 0.42 0.09 61 Palo Colorado forest

20--30 0.65 0.59 13 0.54 2.46 0.13 61 Tabonuco forest

30--40 0.26 0.06 2 0.26 0.30 0.21 61 Bisley

30--40 0.53 0.16 6 0.59 0.65 0.24 61 Dwarf forest

30--40 0.36 0.01 2 0.36 0.36 0.35 61 El Verde

30--40 0.38 0.11 6 1.07 1.49 0.35 61 Montane palm forest

30--40 0.17 0.10 15 0.16 0.36 0.08 61 Palo Colorado forest

30--40 0.73 0.69 15 0.58 2.97 0.10 61 Tabonuco forest

Sodium (mg/g) 0--10 0.07 0.01 3 0.07 0.08 0.05 61 Bisley

0--10 0.19 0.03 5 0.18 0.24 0.16 61 Dwarf forest

0--10 0.10 0.01 3 0.11 0.11 0.09 61 El Verde

0--10 0.14 0.04 10 0.14 0.19 0.07 61 Montane palm forest

0--10 0.06 0.03 15 0.06 0.15 0.03 61 Palo Colorado forest

0--10 0.15 0.08 12 0.15 0.31 0.03 61 Tabonuco forest

10--20 0.06 0.02 3 0.06 0.08 0.03 61 Bisley

10--20 0.14 0.03 6 0.15 0.17 0.09 61 Dwarf forest

10--20 0.09 0.01 3 0.10 0.10 0.08 61 El Verde
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

511

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--20 0.09 0.03 7 0.09 0.13 0.04 61 Montane palm forest

10--20 0.06 0.04 16 0.04 0.15 0.02 61 Palo Colorado forest

10--20 0.18 0.16 14 0.13 0.56 0.03 61 Tabonuco forest

20--30 0.06 0.00 3 0.06 0.03 0.06 61 Bisley

20--30 0.12 0.04 5 0.14 0.15 0.07 61 Dwarf forest

20--30 0.09 0.02 3 0.08 0.11 0.08 61 El Verde

20--30 0.10 0.26 6 0.10 0.13 0.05 61 Montane palm forest

20--30 0.05 0.02 15 0.04 0.10 0.02 61 Palo Colorado forest

20--30 0.15 0.14 13 0.01 0.57 0.03 61 Tabonuco forest

30--40 0.06 0.02 2 0.06 0.07 0.05 61 Bisley

30--40 0.12 0.04 6 0.14 0.15 0.06 61 Dwarf forest

30--40 0.08 0.00 2 0.08 0.08 0.08 61 El Verde

30--40 0.09 0.02 6 0.09 0.12 0.05 61 Montane palm forest

30--40 0.04 0.02 15 0.04 0.08 0.02 61 Palo Colorado forest

30--40 0.17 0.16 15 0.13 0.68 0.02 61 Tabonuco forest

Alluvial deposits Mollisols Sdf Aluminum (cmol/kg-l) 0--15 0.09 0.11 114 0.05 0.51 0.01 62

Bulk density (g/cc) 0--15 0.63 0.11 80 0.66 0.81 0.25 62

Calcium (cmol/kg-l) 0--15 52.84 13.29 114 53.65 86.39 14.79 62

Calcium (mg/g) 0--15 10.57 2.66 114 10.73 17.28 2.96 62

ECEC (cmol/kg-1) 0--15 61.83 14.64 114 60.72 99.84 19.85 62

Iron (mg/kg) 0--15 7 4 114 6 20 2 62

Potassium (cmol/kg-l) 0--15 2.86 3.63 114 0.77 13.39 0.31 62

Potassium (mg/g) 0--15 0.32 0.15 114 0.29 0.65 0.12 62

Loss on ignition (%) 0--15 25.15 6.60 114 23.70 42.25 14.17 62

Magnessium (cmol/kg-1) 0--15 4.99 1.97 114 4.38 11.30 2.25 62

Magnesium (mg/g) 0--15 0.60 0.24 114 0.53 1.36 0.27 62
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

512

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Manganese (mg/g) 0--15 0.09 0.07 114 0.05 0.28 0.01 62

Sodium (cmol/kg-l) 0--15 1.05 1.45 114 0.22 5.22 0.02 62

Sodium (mg/g) 0--15 0.24 0.33 114 0.05 1.20 0.01 62

Nitrogen (%) 0--15 0.66 0.21 114 0.64 1.31 0.30 62

Organic matter (%) 0--15 11.93 4.97 114 10.25 23.76 4.81 62

Phosphorus (mg/kg) 0--15 8 6 114 6 27 3 62

pH (H2O) 0--15 7.74 0.27 114 7.81 8.18 7.01 62

pH (KCl) 0--15 7.03 0.28 114 7.08 7.48 6.04 62

Alluvial deposits Histosol Mfs Aluminum (cmol/kg-l) 0--10 0.11 0.07 3 0.09 0.18 0.05 63 Casuarina equisetifolia plantation

0--10 0.09 0.10 3 0.05 0.20 0.02 63 Eucalyptus robusta plantation

0--10 0.16 0.20 3 0.05 0.39 0.04 63 Leucaena leucocephala plantation

10--25 0.05 0.03 3 0.04 0.08 0.03 63 Casuarina equisetifolia plantation

10--25 0.10 0.03 3 0.09 0.13 0.07 63 Eucalyptus robusta plantation

10--25 0.13 0.13 3 0.06 0.28 0.05 63 Leucaena leucocephala plantation

C/N 0--10 13 3 3 12 16 11 63 Casuarina equisetifolia plantation

0--10 18 9 3 16 28 11 63 Eucalyptus robusta plantation

0--10 20 10 3 20 29 10 63 Leucaena leucocephala plantation

10--25 18 6 3 21 22 11 63 Casuarina equisetifolia plantation

10--25 31 1 3 42 42 10 63 Eucalyptus robusta plantation

10--25 57 42 3 70 90 10 63 Leucaena leucocephala plantation

Calcium (cmol/kg-l) 0--10 6.94 0.77 3 7.23 7.53 6.07 63 Casuarina equisetifolia plantation

0--10 6.64 1.83 3 6.94 8.30 4.68 63 Eucalyptus robusta plantation

0--10 5.59 1.55 3 5.43 7.22 4.13 63 Leucaena leucocephala plantation

10--25 5.63 0.59 3 5.35 6.31 5.24 63 Casuarina equisetifolia plantation

10--25 6.15 1.98 3 5.30 8.41 4.73 63 Eucalyptus robusta plantation

10--25 4.71 2.06 3 3.97 7.03 3.12 63 Leucaena leucocephala plantation
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

513

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Calcium (mg/g) 0--10 1.39 0.15 3 1.45 1.51 1.22 63 Casuarina equisetifolia plantation

0--10 1.33 0.37 3 1.39 1.66 0.94 63 Eucalyptus robusta plantation

0--10 1.12 0.31 3 1.09 1.44 0.83 63 Leucaena leucocephala plantation

10--25 1.13 0.12 3 1.07 1.26 1.05 63 Casuarina equisetifolia plantation

10--25 1.23 0.40 3 1.06 1.68 0.95 63 Eucalyptus robusta plantation

10--25 0.94 0.41 3 0.79 1.41 0.62 63 Leucaena leucocephala plantation

Carbon (%) 0--10 1.77 0.45 3 1.64 2.27 1.41 63 Casuarina equisetifolia plantation

0--10 1.84 0.14 3 1.86 1.97 1.70 63 Eucalyptus robusta plantation

0--10 2.28 0.31 3 2.22 2.61 2.00 63 Leucaena leucocephala plantation

10--25 1.38 0.59 3 1.26 2.02 0.85 63 Casuarina equisetifolia plantation

10--25 1.73 0.33 3 1.67 2.08 1.43 63 Eucalyptus robusta plantation

10--25 2.19 0.48 3 2.11 2.71 1.75 63 Leucaena leucocephala plantation

Effective cation exchange capacity 0--10 9.60 0.74 3 9.96 10.08 8.75 63 Casuarina equisetifolia plantation

{Ca+K+Mg(cmol/kg-1) 0--10 8.48 2.25 3 8.74 10.59 6.12 63 Eucalyptus robusta plantation

0--10 8.20 3.19 3 7.09 11.79 5.71 63 Leucaena leucocephala plantation

10--25 7.50 0.55 3 7.45 8.07 6.98 63 Casuarina equisetifolia plantation

10--25 7.44 2.29 3 6.28 10.08 5.97 63 Eucalyptus robusta plantation

10--25 6.84 3.79 3 4.91 11.20 4.40 63 Leucaena leucocephala plantation

Iron (mg/kg) 0--10 17 3 3 17 20 14 63 Casuarina equisetifolia plantation

0--10 21 6 3 24 25 14 63 Eucalyptus robusta plantation

0--10 28 38 3 7 72 5 63 Leucaena leucocephala plantation

10--25 14 12 3 9 28 6 63 Casuarina equisetifolia plantation

10--25 9 6 3 7 16 8 63 Eucalyptus robusta plantation

10--25 32 52 3 2 92 1 63 Leucaena leucocephala plantation

Loss on ignition (%) 0--10 8.78 1.62 3 7.98 10.64 7.71 63 Casuarina equisetifolia plantation

0--10 4.92 0.46 3 5.05 5.31 4.41 63 Eucalyptus robusta plantation
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

514

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--10 6.71 4.39 3 4.46 11.76 3.90 63 Leucaena leucocephala plantation

10--25 5.70 1.64 3 5.28 7.51 4.32 63 Casuarina equisetifolia plantation

10--25 4.52 1.53 3 5.24 5.55 2.76 63 Eucalyptus robusta plantation

10--25 5.56 4.64 3 2.91 10.92 2.85 63 Leucaena leucocephala plantation

Magnesium (cmol/kg-l) 0--10 1.88 0.24 3 1.97 2.06 1.60 63 Casuarina equisetifolia plantation

0--10 1.34 0.35 3 1.43 1.63 0.95 63 Eucalyptus robusta plantation

0--10 1.89 1.25 3 1.20 3.34 1.14 63 Leucaena leucocephala plantation

10--25 1.34 0.4 3 1.30 1.75 0.96 63 Casuarina equisetifolia plantation

10--25 0.80 0.18 3 0.75 1.00 0.64 63 Eucalyptus robusta plantation

10--25 1.46 1.3 3 0.84 2.96 0.59 63 Leucaena leucocephala plantation

Magnesium (mg/g) 0--10 0.23 0.03 3 0.24 0.25 0.19 63 Casuarina equisetifolia plantation

0--10 0.16 0.04 3 0.17 0.20 0.11 63 Eucalyptus robusta plantation

0--10 0.23 0.15 3 0.15 0.40 0.14 63 Leucaena leucocephala plantation

10--25 0.16 0.05 3 0.16 0.21 0.12 63 Casuarina equisetifolia plantation

10--25 0.10 0.02 3 0.09 0.12 0.08 63 Eucalyptus robusta plantation

10--25 0.18 0.16 3 0.10 0.36 0.07 63 Leucaena leucocephala plantation

Manganese (mg/kg) 0--10 5 1 3 5 6 4 63 Casuarina equisetifolia plantation

0--10 9 3 3 7 13 7 63 Eucalyptus robusta plantation

0--10 33 44 3 11 84 4 63 Leucaena leucocephala plantation

10--25 3 1 3 3 4 2 63 Casuarina equisetifolia plantation

10--25 3 1 3 3 3 2 63 Eucalyptus robusta plantation

10--25 2 2 3 1 4 1 63 Leucaena leucocephala plantation

Nitrogen (%) 0--10 0.14 0.01 3 0.14 0.14 0.13 63 Casuarina equisetifolia plantation

0--10 0.11 0.04 3 0.12 0.15 0.07 63 Eucalyptus robusta plantation

0--10 0.13 0.06 3 0.11 0.20 0.09 63 Leucaena leucocephala plantation

10--25 0.08 0.02 3 0.08 0.09 0.06 63 Casuarina equisetifolia plantation
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

515

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--25 0.08 0.06 3 0.05 0.15 0.04 63 Eucalyptus robusta plantation

10--25 0.08 0.09 3 0.03 0.18 0.03 63 Leucaena leucocephala plantation

pH (H2O) 0--10 7.64 0.11 3 7.66 7.73 7.52 63 Casuarina equisetifolia plantation

0--10 7.68 0.08 3 7.67 7.76 7.60 63 Eucalyptus robusta plantation

0--10 7.49 0.80 3 7.85 8.05 6.58 63 Leucaena leucocephala plantation

10--25 7.66 0.40 3 7.60 7.99 7.22 63 Casuarina equisetifolia plantation

10--25 7.82 0.36 3 7.66 8.23 7.57 63 Eucalyptus robusta plantation

10--25 7.55 1.06 3 8.06 8.26 6.34 63 Leucaena leucocephala plantation

pH (KCl) 0--10 6.89 0.40 3 7.02 7.21 6.45 63 Casuarina equisetifolia plantation

0--10 6.90 0.46 3 6.77 7.41 6.51 63 Eucalyptus robusta plantation

0--10 6.65 1.09 3 6.95 7.55 5.44 63 Leucaena leucocephala plantation

10--25 7.34 0.23 3 7.43 7.50 7.08 63 Casuarina equisetifolia plantation

10--25 7.26 0.46 3 7.19 7.75 6.84 63 Eucalyptus robusta plantation

10--25 6.91 1.26 3 7.51 7.76 5.46 63 Leucaena leucocephala plantation

Phosphorus (mg/kg) 0--10 5 0 3 5 5 5 63 Casuarina equisetifolia plantation

0--10 6 1 3 6 7 5 63 Eucalyptus robusta plantation

0--10 6 2 3 6 7 4 63 Leucaena leucocephala plantation

10--25 4 1 3 4 5 3 63 Casuarina equisetifolia plantation

10--25 4 1 3 5 5 3 63 Eucalyptus robusta plantation

10--25 4 3 3 3 8 2 63 Leucaena leucocephala plantation

Potassium (cmol/kg-l) 0--10 0.03 0.04 3 0.28 0.32 0.24 63 Casuarina equisetifolia plantation

0--10 0.22 0.01 3 0.22 0.22 0.21 63 Eucalyptus robusta plantation

0--10 0.31 0.02 3 0.32 0.33 0.29 63 Leucaena leucocephala plantation

10--25 0.26 0.03 3 0.27 0.28 0.22 63 Casuarina equisetifolia plantation

10--25 0.20 0.01 3 0.20 0.21 0.19 63 Eucalyptus robusta plantation

10--25 0.25 0.07 3 0.24 0.32 0.19 63 Leucaena leucocephala plantation
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

516

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Potassium (mg/g) 0--10 0.11 0.02 3 0.11 0.12 0.09 63 Casuarina equisetifolia plantation

0--10 0.08 0.00 3 0.08 0.08 0.08 63 Eucalyptus robusta plantation

0--10 0.19 0.01 3 0.12 0.12 0.11 63 Leucaena leucocephala plantation

10--25 0.10 0.01 3 0.10 0.11 0.08 63 Casuarina equisetifolia plantation

10--25 0.08 0.00 3 0.08 0.08 0.07 63 Eucalyptus robusta plantation

10--25 0.10 0.03 3 0.09 0.12 0.07 63 Leucaena leucocephala plantation

Sodium (cmol/kg-l) 0--10 0.39 0.15 3 0.38 0.55 0.25 63 Casuarina equisetifolia plantation

0--10 0.20 0.06 3 0.23 0.24 0.13 63 Eucalyptus robusta plantation

0--10 0.24 0.25 3 0.12 0.52 0.07 63 Leucaena leucocephala plantation

10--25 0.22 0.12 3 0.15 0.36 0.14 63 Casuarina equisetifolia plantation

10--25 0.20 0.16 3 0.17 0.38 0.06 63 Eucalyptus robusta plantation

10--25 0.29 0.35 3 0.11 0.69 0.06 63 Leucaena leucocephala plantation

Sodium (mg/kg) 0--10 90 34 3 87 125 58 63 Casuarina equisetifolia plantation

0--10 46 14 3 53 55 30 63 Eucalyptus robusta plantation

0--10 55 56 3 28 119 17 63 Leucaena leucocephala plantation

10--25 50 29 3 34 84 33 63 Casuarina equisetifolia plantation

10--25 47 37 3 40 87 15 63 Eucalyptus robusta plantation

10--25 66 80 3 26 158 13 63 Leucaena leucocephala plantation

Sulfur (%) 0--10 0.03 0.00 3 0.03 0.03 0.03 63 Casuarina equisetifolia plantation

0--10 0.04 0.02 3 0.03 0.06 0.03 63 Eucalyptus robusta plantation

0--10 0.04 0.00 3 0.04 0.04 0.03 63 Leucaena leucocephala plantation

10--25 0.02 0.01 3 0.03 0.03 0.02 63 Casuarina equisetifolia plantation

10--25 0.03 0.01 3 0.03 0.03 0.02 63 Eucalyptus robusta plantation

10--25 0.03 0.00 3 0.03 0.03 0.03 63 Leucaena leucocephala plantation

Alluvial deposits Histosol Mfs Carbon (%) 0--10 1.82 0.20 13 1.73 2.05 1.68 64 C. equisetifolia + E. robusta

0--10 1.75 0.02 3 1.74 1.77 1.73 64 C. equisetifolia + L. leucaena
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

517

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--10 1.91 0.37 3 2.07 2.17 1.48 64 Casuarina equisetifolia

0--10 1.75 0.41 3 1.67 2.19 1.39 64 Eucalyptus robusta

0--10 2.24 0.59 3 2.29 2.81 1.63 64 Leucaena leucocephala

10--20 1.87 0.01 2 1.87 1.87 1.86 64 C. equisetifolia + E. robusta

10--20 1.41 0.12 3 1.45 1.51 1.28 64 C. equisetifolia + L. leucaena

10--20 1.44 0.26 4 1.35 1.82 1.23 64 Casuarina equisetifolia

10--20 1.53 0.16 3 1.61 1.63 1.35 64 Eucalyptus robusta

10--20 1.50 0.74 3 1.70 2.12 0.68 64 Leucaena leucocephala

20--30 1.16 0.37 2 1.16 1.42 0.90 64 C. equisetifolia + E. robusta

20--30 0.91 0.32 3 0.81 1.26 0.65 64 C. equisetifolia + L. leucaena

20--30 1.36 0.41 3 1.18 1.83 1.07 64 Casuarina equisetifolia

20--30 1.60 0.21 3 1.72 1.72 1.35 64 Eucalyptus robusta

20--30 1.17 0.68 3 1.41 1.69 0.40 64 Leucaena leucocephala

30--40 0.80 0.18 2 0.80 0.92 0.67 64 C. equisetifolia + E. robusta

30--40 1.00 0.64 4 1.01 1.56 0.43 64 C. equisetifolia + L. leucaena

30--40 1.61 0.54 3 1.61 2.16 1.08 64 Casuarina equisetifolia

30--40 2.01 0.66 4 2.25 2.50 1.05 64 Eucalyptus robusta

30--40 1.08 1.27 4 0.59 2.96 0.19 64 Leucaena leucocephala

Nitrogen (%) 0--10 0.13 0.05 13 0.14 0.21 0.05 64 C. equisetifolia + E. robusta

0--10 0.14 0.03 13 0.14 0.19 0.08 64 C. equisetifolia + L. leucaena

0--10 0.11 0.03 15 0.11 0.18 0.06 64 Casuarina equisetifolia

0--10 0.11 0.03 13 0.10 0.18 0.06 64 Eucalyptus robusta

0--10 0.11 0.04 14 0.12 0.19 0.06 64 Leucaena leucocephala

10--20 0.13 0.02 11 0.12 0.19 0.11 64 C. equisetifolia + E. robusta

10--20 0.12 0.04 13 0.13 0.20 0.07 64 C. equisetifolia + L. leucaena

10--20 0.10 0.03 13 0.10 0.16 0.05 64 Casuarina equisetifolia
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

518

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--20 0.09 0.04 12 0.08 0.17 0.04 64 Eucalyptus robusta

10--20 0.09 0.03 12 0.09 0.15 0.06 64 Leucaena leucocephala

20--30 0.10 0.03 9 0.10 0.16 0.05 64 C. equisetifolia + E. robusta

20--30 0.11 0.11 10 0.07 0.40 0.04 64 C. equisetifolia + L. leucaena

20--30 0.06 0.03 10 0.06 0.14 0.02 64 Casuarina equisetifolia

20--30 0.06 0.02 10 0.06 0.10 0.02 64 Eucalyptus robusta

20--30 0.07 0.03 9 0.06 0.12 0.03 64 Leucaena leucocephala

30--40 0.08 0.02 8 0.07 0.11 0.04 64 C. equisetifolia + E. robusta

30--40 0.05 0.02 10 0.04 0.08 0.03 64 C. equisetifolia + L. leucaena

30--40 0.04 0.02 9 0.04 0.08 0.01 64 Casuarina equisetifolia

30--40 0.04 0.03 10 0.03 0.08 0.01 64 Eucalyptus robusta

30--40 0.04 0.02 10 0.04 0.07 0.02 64 Leucaena leucocephala

Sulfur (mg/kg) 0--10 250 60 3 220 320 210 64 C. equisetifolia + E. robusta

0--10 230 40 3 250 250 190 64 C. equisetifolia + L. leucaena

0--10 100 3 3 200 200 100 64 Casuarina equisetifolia

0--10 120 10 3 120 130 110 64 Eucalyptus robusta

0--10 130 30 3 130 150 100 64 Leucaena leucocephala

10--20 100 0 2 100 100 100 64 C. equisetifolia + E. robusta

10--20 100 40 3 150 180 100 64 C. equisetifolia + L. leucaena

10--20 90 20 3 90 110 70 64 Casuarina equisetifolia

10--20 130 30 3 140 150 90 64 Eucalyptus robusta

10--20 90 1 64 Leucaena leucocephala

20--30 300 100 2 300 400 200 64 C. equisetifolia + E. robusta

20--30 200 50 3 220 230 140 64 C. equisetifolia + L. leucaena

20--30 100 40 3 90 150 70 64 Casuarina equisetifolia

20--30 100 10 3 100 110 90 64 Eucalyptus robusta
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

519

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

20--30 120 40 3 110 170 90 64 Leucaena leucocephala

30--40 200 100 2 200 300 200 64 C. equisetifolia + E. robusta

30--40 150 30 4 140 200 120 64 C. equisetifolia + L. leucaena

30--40 120 10 2 120 130 110 64 Casuarina equisetifolia

30--40 120 20 4 120 140 90 64 Eucalyptus robusta

30--40 100 50 3 90 150 60 64 Leucaena leucocephala

Aluminum (cmol/kg-l) 0--10 0.10 0.05 13 0.10 0.17 0.01 64 C. equisetifolia + E. robusta

0--10 0.07 0.05 13 0.08 0.14 0.01 64 C. equisetifolia + L. leucaena

0--10 0.14 0.08 15 0.15 0.26 0.02 64 Casuarina equisetifolia

0--10 0.10 0.05 13 0.08 0.18 0.04 64 Eucalyptus robusta

0--10 0.10 0.05 14 0.09 0.18 0.01 64 Leucaena leucocephala

10--20 0.08 0.06 11 0.09 0.18 0.01 64 C. equisetifolia + E. robusta

10--20 0.05 0.03 13 0.05 0.10 0.01 64 C. equisetifolia + L. leucaena

10--20 0.11 0.09 13 0.10 0.24 0.01 64 Casuarina equisetifolia

10--20 0.09 0.06 12 0.09 0.17 0.01 64 Eucalyptus robusta

10--20 0.11 0.06 12 0.10 0.20 0.02 64 Leucaena leucocephala

20--30 0.05 0.04 9 0.03 0.12 0.01 64 C. equisetifolia + E. robusta

20--30 0.06 0.04 10 0.06 0.12 0.01 64 C. equisetifolia + L. leucaena

20--30 0.08 0.06 10 0.07 0.18 0.01 64 Casuarina equisetifolia

20--30 0.07 0.04 10 0.08 0.14 0.02 64 Eucalyptus robusta

20--30 0.07 0.06 9 0.06 0.16 0.01 64 Leucaena leucocephala

30--40 0.06 0.04 8 0.06 0.12 0.01 64 C. equisetifolia + E. robusta

30--40 0.06 0.06 10 0.04 0.18 0.01 64 C. equisetifolia + L. leucaena

30--40 0.07 0.07 9 0.03 0.23 0.01 64 Casuarina equisetifolia

30--40 0.05 0.04 10 0.05 0.13 0.02 64 Eucalyptus robusta

30--40 0.05 0.03 10 0.04 0.09 0.01 64 Leucaena leucocephala
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

520

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

C/N 0--10 11 1 3 11 12 11 64 C. equisetifolia + E. robusta

0--10 12 1 3 12 13 1 64 C. equisetifolia + L. leucaena

0--10 13 1 3 14 14 12 64 Casuarina equisetifolia

0--10 17 4 3 16 22 14 64 Eucalyptus robusta

0--10 15 7 3 12 23 11 64 Leucaena leucocephala

10--20 15 1 2 15 16 15 64 C. equisetifolia + E. robusta

10--20 11 1 3 11 13 10 64 C. equisetifolia + L. leucaena

10--20 17 7 4 15 28 12 64 Casuarina equisetifolia

10--20 24 12 3 23 36 12 64 Eucalyptus robusta

10--20 17 11 3 11 30 10 64 Leucaena leucocephala

20--30 9 1 2 9 10 9 64 C. equisetifolia + E. robusta

20--30 10 1 3 10 11 9 64 C. equisetifolia + L. leucaena

20--30 21 11 3 23 31 9 64 Casuarina equisetifolia

20--30 57 31 3 71 78 21 64 Eucalyptus robusta

20--30 17 9 3 12 27 11 64 Leucaena leucocephala

30--40 9 1 2 9 10 9 64 C. equisetifolia + E. robusta

30--40 17 3 4 18 20 13 64 C. equisetifolia + L. leucaena

30--40 91 29 3 107 108 57 64 Casuarina equisetifolia

30--40 140 122 4 116 307 21 64 Eucalyptus robusta

30--40 42 65 4 10 140 8 64 Leucaena leucocephala

Calcium (cmol/kg-l) 0--10 8.48 2.27 13 8.43 13.19 5.63 64 C. equisetifolia + E. robusta

0--10 9.42 2.92 13 8.92 15.19 6.59 64 C. equisetifolia + L. leucaena

0--10 10.23 3.43 15 9.39 15.64 5.84 64 Casuarina equisetifolia

0--10 8.81 2.55 13 8.45 13.38 4.51 64 Eucalyptus robusta

0--10 8.70 4.76 14 7.21 19.66 3.84 64 Leucaena leucocephala

10--20 8.82 1.48 11 8.82 11.24 6.34 64 C. equisetifolia + E. robusta
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

521

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--20 9.34 3.11 13 7.91 15.48 6.46 64 C. equisetifolia + L. leucaena

10--20 9.51 4.01 13 7.67 16.32 4.01 64 Casuarina equisetifolia

10--20 8.12 2.69 12 7.87 12.53 4.06 64 Eucalyptus robusta

10--20 8.92 4.91 12 7.99 19.58 3.98 64 Leucaena leucocephala

20--30 7.81 1.52 9 8.17 9.82 5.31 64 C. equisetifolia + E. robusta

20--30 6.89 1.33 10 6.62 9.18 4.38 64 C. equisetifolia + L. leucaena

20--30 7.23 3.57 10 6.31 12.73 3.72 64 Casuarina equisetifolia

20--30 7.34 3.22 10 6.67 13.67 3.37 64 Eucalyptus robusta

20--30 6.85 3.14 9 6.54 14.04 3.73 64 Leucaena leucocephala

30--40 7.24 2.15 8 7.33 10.04 4.19 64 C. equisetifolia + E. robusta

30--40 6.27 1.63 10 6.32 9.09 4.06 64 C. equisetifolia + L. leucaena

30--40 6.97 3.78 9 5.19 12.45 3.57 64 Casuarina equisetifolia

30--40 5.90 2.84 10 5.38 10.66 2.56 64 Eucalyptus robusta

30--40 5.90 2.77 10 5.60 12.26 3.39 64 Leucaena leucocephala

Calcium (mg/g) 0--10 1.70 0.45 13 1.69 2.64 1.13 64 C. equisetifolia + E. robusta

0--10 1.88 0.59 13 1.78 3.04 1.32 64 C. equisetifolia + L. leucaena

0--10 2.05 0.69 15 1.88 3.13 1.17 64 Casuarina equisetifolia

0--10 1.76 0.51 13 1.69 2.68 0.90 64 Eucalyptus robusta

0--10 1.74 0.95 14 1.44 3.93 0.77 64 Leucaena leucocephala

10--20 1.76 0.30 11 1.76 2.25 1.27 64 C. equisetifolia + E. robusta

10--20 1.87 0.62 13 1.58 3.10 1.29 64 C. equisetifolia + L. leucaena

10--20 1.90 0.80 13 1.54 3.27 0.80 64 Casuarina equisetifolia

10--20 1.62 0.54 12 1.57 2.51 0.81 64 Eucalyptus robusta

10--20 1.78 0.98 12 1.60 3.92 0.80 64 Leucaena leucocephala

20--30 1.56 0.30 9 1.63 1.96 1.06 64 C. equisetifolia + E. robusta

20--30 1.38 0.27 10 1.32 1.84 0.88 64 C. equisetifolia + L. leucaena
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

522

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

20--30 1.45 0.71 10 1.26 2.55 0.74 64 Casuarina equisetifolia

20--30 1.47 0.65 10 1.33 2.73 0.67 64 Eucalyptus robusta

20--30 1.37 0.63 9 1.31 2.81 0.75 64 Leucaena leucocephala

30--40 1.45 0.43 8 1.47 2.01 0.84 64 C. equisetifolia + E. robusta

30--40 1.25 0.33 10 1.26 1.82 0.81 64 C. equisetifolia + L. leucaena

30--40 1.39 0.76 9 1.04 2.49 0.71 64 Casuarina equisetifolia

30--40 1.18 0.57 10 1.08 2.13 0.51 64 Eucalyptus robusta

30--40 1.18 0.55 10 1.12 2.45 0.68 64 Leucaena leucocephala

Effective cation exchange capacity 0--10 10.73 2.87 13 10.55 16.32 7.37 64 C. equisetifolia + E. robusta

{Ca+K+Mg(cmol/kg-1) 0--10 12.20 3.20 13 11.22 18.29 8.41 64 C. equisetifolia + L. leucaena

0--10 12.35 3.75 15 11.55 18.49 7.11 64 Casuarina equisetifolia

0--10 10.60 2.91 13 9.57 16.98 6.04 64 Eucalyptus robusta

0--10 10.85 5.65 14 9.70 23.74 4.80 64 Leucaena leucocephala

10--20 10.85 2.00 11 11.01 14.70 7.77 64 C. equisetifolia + E. robusta

10--20 11.64 3.61 13 10.32 19.00 7.67 64 C. equisetifolia + L. leucaena

10--20 11.38 4.49 13 9.66 19.20 5.13 64 Casuarina equisetifolia

10--20 9.64 3.18 12 8.95 15.39 5.20 64 Eucalyptus robusta

10--20 11.01 5.84 12 10.04 23.95 4.78 64 Leucaena leucocephala

20--30 9.42 1.72 9 9.73 11.65 7.05 64 C. equisetifolia + E. robusta

20--30 8.47 1.26 10 8.50 10.98 6.61 64 C. equisetifolia + L. leucaena

20--30 8.54 4.02 10 7.55 14.97 4.33 64 Casuarina equisetifolia

20--30 8.35 3.25 10 7.67 14.76 4.09 64 Eucalyptus robusta

20--30 8.41 3.77 9 8.06 16.45 4.42 64 Leucaena leucocephala

30--40 8.67 2.34 8 8.75 11.48 5.76 64 C. equisetifolia + E. robusta

30--40 7.87 1.78 10 7.99 10.57 4.91 64 C. equisetifolia + L. leucaena

30--40 8.24 4.18 9 6.33 14.45 4.32 64 Casuarina equisetifolia
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

523

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

30--40 6.84 2.98 10 6.55 11.66 3.20 64 Eucalyptus robusta

30--40 7.12 3.28 10 6.93 14.23 3.96 64 Leucaena leucocephala

Iron (mg/kg) 0--10 30 23 13 23 73 9 64 C. equisetifolia + E. robusta

0--10 29 16 12 28 61 4 64 C. equisetifolia + L. leucaena

0--10 23 13 15 22 53 9 64 Casuarina equisetifolia

0--10 12 5 13 9 22 5 64 Eucalyptus robusta

0--10 19 19 14 9 59 6 64 Leucaena leucocephala

10--20 23 18 11 15 67 7 64 C. equisetifolia + E. robusta

10--20 29 19 12 29 72 10 64 C. equisetifolia + L. leucaena

10--20 21 15 13 14 59 7 64 Casuarina equisetifolia

10--20 11 6 12 11 28 5 64 Eucalyptus robusta

10--20 20 18 12 10 56 5 64 Leucaena leucocephala

20--30 23 14 9 17 52 6 64 C. equisetifolia + E. robusta

20--30 21 11 10 16 39 8 64 C. equisetifolia + L. leucaena

20--30 11 10 10 8 32 2 64 Casuarina equisetifolia

20--30 8 3 10 8 13 1 64 Eucalyptus robusta

20--30 15 16 9 7 55 5 64 Leucaena leucocephala

30--40 20 12 8 20 41 4 64 C. equisetifolia + E. robusta

30--40 17 9 10 14 33 5 64 C. equisetifolia + L. leucaena

30--40 7 4 9 5 14 3 64 Casuarina equisetifolia

30--40 6 4 10 6 10 1 64 Eucalyptus robusta

30--40 13 9 10 11 26 4 64 Leucaena leucocephala

Loss on ignition (%) 0--10 6.31 1.88 12 5.86 10.20 3.33 64 C. equisetifolia + E. robusta

0--10 5.63 0.94 13 5.16 6.86 4.47 64 C. equisetifolia + L. leucaena

0--10 5.11 1.05 14 5.23 7.02 3.72 64 Casuarina equisetifolia

0--10 4.64 1.01 13 4.40 6.88 3.01 64 Eucalyptus robusta
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

524

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--10 4.81 1.50 13 4.84 8.26 3.13 64 Leucaena leucocephala

10--20 5.72 1.61 11 5.29 9.60 4.32 64 C. equisetifolia + E. robusta

10--20 5.25 1.10 13 4.70 7.72 4.15 64 C. equisetifolia + L. leucaena

10--20 4.59 1.24 13 4.29 7.18 2.31 64 Casuarina equisetifolia

10--20 4.04 1.23 12 3.62 6.05 2.34 64 Eucalyptus robusta

10--20 5.32 4.18 12 4.14 18.31 2.86 64 Leucaena leucocephala

20--30 5.11 1.62 9 4.44 7.90 3.78 64 C. equisetifolia + E. robusta

20--30 4.13 1.14 9 3.82 6.91 3.16 64 C. equisetifolia + L. leucaena

20--30 3.28 0.61 9 3.30 4.29 2.48 64 Casuarina equisetifolia

20--30 3.38 1.00 9 3.08 5.01 2.10 64 Eucalyptus robusta

20--30 4.13 1.53 9 3.93 7.67 2.90 64 Leucaena leucocephala

30--40 4.07 0.85 8 3.71 5.24 3.27 64 C. equisetifolia + E. robusta

30--40 3.35 0.60 10 3.48 4.35 2.29 64 C. equisetifolia + L. leucaena

30--40 2.62 0.75 9 2.51 3.56 1.63 64 Casuarina equisetifolia

30--40 2.69 0.61 10 2.50 3.69 1.78 64 Eucalyptus robusta

30--40 2.80 0.43 10 2.83 3.44 2.18 64 Leucaena leucocephala

Magnesium (cmol/kg-l) 0--10 1.48 0.59 13 1.45 2.61 0.54 64 C. equisetifolia + E. robusta

0--10 1.81 0.41 13 1.88 2.35 0.99 64 C. equisetifolia + L. leucaena

0--10 1.30 0.31 15 1.32 2.05 0.77 64 Casuarina equisetifolia

0--10 1.10 0.26 13 1.01 1.75 0.70 64 Eucalyptus robusta

0--10 1.32 0.58 14 1.22 2.37 0.69 64 Leucaena leucocephala

10--20 1.30 0.47 11 1.07 2.12 0.78 64 C. equisetifolia + E. robusta

10--20 1.56 0.66 13 1.30 2.75 0.81 64 C. equisetifolia + L. leucaena

10--20 1.12 0.36 13 1.13 1.71 0.63 64 Casuarina equisetifolia

10--20 0.82 0.23 12 0.73 1.38 0.58 64 Eucalyptus robusta

10--20 1.15 0.70 12 0.95 2.14 0.11 64 Leucaena leucocephala
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

525

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

20--30 1.07 0.20 9 1.09 1.36 0.77 64 C. equisetifolia + E. robusta

20--30 1.02 0.29 10 0.97 1.51 0.62 64 C. equisetifolia + L. leucaena

20--30 0.75 0.36 10 0.67 1.55 0.36 64 Casuarina equisetifolia

20--30 0.62 0.14 10 0.62 0.85 0.38 64 Eucalyptus robusta

20--30 0.93 0.48 9 0.76 1.80 0.48 64 Leucaena leucocephala

30--40 0.89 0.23 8 0.87 1.21 0.51 64 C. equisetifolia + E. robusta

30--40 0.95 0.25 10 0.98 1.26 0.52 64 C. equisetifolia + L. leucaena

30--40 0.69 0.23 9 0.74 1.00 0.34 64 Casuarina equisetifolia

30--40 0.53 0.19 10 0.51 0.84 0.30 64 Eucalyptus robusta

30--40 0.69 0.36 10 0.61 1.22 0.22 64 Leucaena leucocephala

Magnesium (mg/g) 0--10 0.18 0.07 13 0.17 0.31 0.06 64 C. equisetifolia + E. robusta

0--10 0.22 0.05 13 0.23 0.28 0.12 64 C. equisetifolia + L. leucaena

0--10 0.16 0.04 15 0.16 0.25 0.09 64 Casuarina equisetifolia

0--10 0.13 0.03 13 0.12 0.21 0.08 64 Eucalyptus robusta

0--10 0.16 0.07 1 0.15 0.28 0.08 64 Leucaena leucocephala

10--20 0.16 0.06 11 0.13 0.25 0.09 64 C. equisetifolia + E. robusta

10--20 0.19 0.08 13 0.16 0.33 0.10 64 C. equisetifolia + L. leucaena

10--20 0.13 0.04 13 0.14 0.21 0.08 64 Casuarina equisetifolia

10--20 0.10 0.03 12 0.09 0.17 0.07 64 Eucalyptus robusta

10--20 0.14 0.08 12 0.11 0.26 0.01 64 Leucaena leucocephala

20--30 0.13 0.02 9 0.13 0.16 0.09 64 C. equisetifolia + E. robusta

20--30 0.12 0.04 10 0.12 0.18 0.07 64 C. equisetifolia + L. leucaena

20--30 0.09 0.04 10 0.08 0.19 0.04 64 Casuarina equisetifolia

20--30 0.07 0.02 10 0.07 0.10 0.05 64 Eucalyptus robusta

20--30 0.11 0.06 9 0.09 0.22 0.06 64 Leucaena leucocephala

30--40 0.11 0.03 8 0.11 0.15 0.06 64 C. equisetifolia + E. robusta
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

526

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

30--40 0.11 0.03 10 0.12 0.15 0.06 64 C. equisetifolia + L. leucaena

30--40 0.08 0.03 9 0.09 0.12 0.04 64 Casuarina equisetifolia

30--40 0.06 0.02 10 0.06 0.10 0.04 64 Eucalyptus robusta

30--40 0.08 0.04 10 0.07 0.15 0.03 64 Leucaena leucocephala

Manganese (mg/kg) 0--10 16 9 13 16 31 3 64 C. equisetifolia + E. robusta

0--10 14 8 13 14 25 2 64 C. equisetifolia + L. leucaena

0--10 18 11 15 14 38 6 64 Casuarina equisetifolia

0--10 13 6 13 13 23 5 64 Eucalyptus robusta

0--10 13 0 14 11 26 4 64 Leucaena leucocephala

10--20 14 8 11 13 29 3 64 C. equisetifolia + E. robusta

10--20 11 7 13 14 20 1 64 C. equisetifolia + L. leucaena

10--20 14 7 13 16 25 5 64 Casuarina equisetifolia

10--20 11 4 12 11 18 5 64 Eucalyptus robusta

10--20 12 8 12 12 31 3 64 Leucaena leucocephala

20--30 9 7 9 11 20 3 64 C. equisetifolia + E. robusta

20--30 10 5 10 10 18 3 64 C. equisetifolia + L. leucaena

20--30 7 10 10 8 11 3 64 Casuarina equisetifolia

20--30 7 3 10 8 11 3 64 Eucalyptus robusta

20--30 7 5 9 6 19 3 64 Leucaena leucocephala

30--40 10 6 8 9 17 3 64 C. equisetifolia + E. robusta

30--40 10 4 10 11 17 4 64 C. equisetifolia + L. leucaena

30--40 7 2 9 6 11 4 64 Casuarina equisetifolia

30--40 6 3 10 6 11 3 64 Eucalyptus robusta

30--40 7 3 10 7 10 3 64 Leucaena leucocephala

pH (H2O) 0--10 7.56 0.43 12 7.71 8.04 6.81 64 C. equisetifolia + E. robusta

0--10 7.67 0.22 13 7.71 8.03 7.23 64 C. equisetifolia + L. leucaena
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

527

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

0--10 7.79 0.20 14 7.74 8.19 7.56 64 Casuarina equisetifolia

0--10 7.84 0.26 13 7.85 8.14 7.25 64 Eucalyptus robusta

0--10 7.79 0.33 13 7.92 8.20 7.03 64 Leucaena leucocephala

10--20 7.67 0.35 11 7.78 8.06 6.95 64 C. equisetifolia + E. robusta

10--20 7.61 0.37 13 7.62 8.10 6.65 64 C. equisetifolia + L. leucaena

10--20 7.84 0.25 13 7.88 8.17 7.21 64 Casuarina equisetifolia

10--20 7.93 0.11 12 7.96 8.09 7.75 64 Eucalyptus robusta

10--20 7.81 0.45 12 7.98 8.21 6.63 64 Leucaena leucocephala

20--30 7.67 0.32 9 7.82 7.91 6.96 64 C. equisetifolia + E. robusta

20--30 7.73 0.25 9 7.82 8.01 7.25 64 C. equisetifolia + L. leucaena

20--30 7.84 0.29 9 7.91 8.16 7.26 64 Casuarina equisetifolia

20--30 7.97 0.15 9 7.97 8.19 7.71 64 Eucalyptus robusta

20--30 7.96 0.27 9 8.03 8.18 7.31 64 Leucaena leucocephala

30--40 7.71 0.42 8 7.81 8.06 6.71 64 C. equisetifolia + E. robusta

30--40 7.88 0.26 10 7.95 8.16 7.25 64 C. equisetifolia + L. leucaena

30--40 8.04 0.15 9 8.05 8.24 7.76 64 Casuarina equisetifolia

30--40 8.02 0.13 10 8.06 8.24 7.76 64 Eucalyptus robusta

30--40 7.99 0.15 10 8.02 8.18 7.66 64 Leucaena leucocephala

pH (KCl) 0--10 6.74 0.57 9 7.04 7.23 5.81 64 C. equisetifolia + E. robusta

0--10 6.81 0.31 10 6.81 7.27 6.23 64 C. equisetifolia + L. leucaena

0--10 6.98 0.26 11 6.95 7.37 6.57 64 Casuarina equisetifolia

0--10 7.14 0.13 10 7.15 7.34 6.86 64 Eucalyptus robusta

0--10 7.07 0.57 10 7.30 7.63 6.04 64 Leucaena leucocephala

10--20 6.85 0.46 9 6.97 7.33 6.03 64 C. equisetifolia + E. robusta

10--20 6.70 0.60 10 6.70 7.34 5.43 64 C. equisetifolia + L. leucaena

10--20 7.06 0.35 10 7.05 7.40 6.19 64 Casuarina equisetifolia
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

528

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--20 7.22 0.21 9 7.16 7.60 6.94 64 Eucalyptus robusta

10--20 7.08 0.57 9 7.20 7.82 5.82 64 Leucaena leucocephala

20--30 6.97 0.21 7 7.03 7.18 6.56 64 C. equisetifolia + E. robusta

20--30 6.95 0.45 6 7.09 7.33 6.21 64 C. equisetifolia + L. leucaena

20--30 7.28 0.34 6 7.26 7.73 6.87 64 Casuarina equisetifolia

20--30 7.28 0.28 6 7.36 7.49 6.72 64 Eucalyptus robusta

20--30 7.28 0.66 6 7.45 7.83 5.99 64 Leucaena leucocephala

30--40 6.98 0.24 6 7.00 7.34 6.61 64 C. equisetifolia + E. robusta

30--40 6.97 0.41 6 6.92 7.45 6.43 64 C. equisetifolia + L. leucaena

30--40 7.35 0.29 6 7.37 7.72 6.99 64 Casuarina equisetifolia

30--40 7.35 0.10 6 7.37 7.49 7.21 64 Eucalyptus robusta

30--40 7.31 0.36 6 7.71 7.73 6.69 64 Leucaena leucocephala

Phosphorus (mg/kg) 0--10 8 2 13 8 11 5 64 C. equisetifolia + E. robusta

0--10 7 2 13 7 9 2 64 C. equisetifolia + L. leucaena

0--10 8 2 15 8 16 6 64 Casuarina equisetifolia

0--10 7 2 13 7 12 3 64 Eucalyptus robusta

0--10 7 2 14 8 12 4 64 Leucaena leucocephala

10--20 7 1 11 7 9 6 64 C. equisetifolia + E. robusta

10--20 6 2 13 6 9 2 64 C. equisetifolia + L. leucaena

10--20 7 2 13 7 12 4 64 Casuarina equisetifolia

10--20 6 2 12 6 10 4 64 Eucalyptus robusta

10--20 7 2 12 7 10 4 64 Leucaena leucocephala

20--30 6 2 9 6 9 4 64 C. equisetifolia + E. robusta

20--30 4 1 10 4 6 3 64 C. equisetifolia + L. leucaena

20--30 4 1 10 4 6 2 64 Casuarina equisetifolia

20--30 4 2 10 4 7 2 64 Eucalyptus robusta
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

529

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

20--30 4 2 9 4 8 3 64 Leucaena leucocephala

30--40 5 2 8 4 9 3 64 C. equisetifolia + E. robusta

30--40 4 0 10 4 4 3 64 C. equisetifolia + L. leucaena

30--40 3 1 9 3 5 2 64 Casuarina equisetifolia

30--40 4 2 10 3 7 2 64 Eucalyptus robusta

30--40 3 1 10 3 5 2 64 Leucaena leucocephala

Potassium (cmol/kg-l) 0--10 0.17 0.10 13 0.13 0.37 0.06 64 C. equisetifolia + E. robusta

0--10 0.17 0.11 13 0.14 0.39 0.03 64 C. equisetifolia + L. leucaena

0--10 0.17 0.08 15 0.13 0.35 0.10 64 Casuarina equisetifolia

0--10 0.15 0.08 13 0.12 0.30 0.05 64 Eucalyptus robusta

0--10 0.16 0.10 14 0.15 0.35 0.07 64 Leucaena leucocephala

10--20 0.14 0.07 11 0.12 0.28 0.06 64 C. equisetifolia + E. robusta

10--20 0.13 0.10 13 0.09 0.34 0.03 64 C. equisetifolia + L. leucaena

10--20 0.15 0.07 13 0.13 0.27 0.07 64 Casuarina equisetifolia

10--20 0.14 0.07 12 0.11 0.25 0.05 64 Eucalyptus robusta

10--20 0.14 0.08 12 0.12 0.28 0.05 64 Leucaena leucocephala

20--30 0.12 0.08 9 0.08 0.28 0.04 64 C. equisetifolia + E. robusta

20--30 0.11 0.08 10 0.07 0.24 0.04 64 C. equisetifolia + L. leucaena

20--30 0.11 0.08 10 0.08 0.23 0.04 64 Casuarina equisetifolia

20--30 0.12 0.07 10 0.10 0.24 0.05 64 Eucalyptus robusta

20--30 0.12 0.09 9 0.08 0.24 0.04 64 Leucaena leucocephala

30--40 0.11 0.07 8 0.10 0.24 0.03 64 C. equisetifolia + E. robusta

30--40 0.13 0.09 10 0.08 0.23 0.03 64 C. equisetifolia + L. leucaena

30--40 0.11 0.07 9 0.08 0.22 0.04 64 Casuarina equisetifolia

30--40 0.12 0.08 10 0.09 0.24 0.05 64 Eucalyptus robusta

30--40 0.12 0.09 10 0.07 0.24 0.03 64 Leucaena leucocephala
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

530

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Potassium (mg/g) 0--10 0.07 0.04 13 0.05 0.14 0.03 64 C. equisetifolia + E. robusta

0--10 0.07 0.04 13 0.06 0.15 0.01 64 C. equisetifolia + L. leucaena

0--10 0.07 0.03 15 0.05 0.14 0.04 64 Casuarina equisetifolia

0--10 0.06 0.03 13 0.05 0.12 0.02 64 Eucalyptus robusta

0--10 0.06 0.04 14 0.06 0.13 0.03 64 Leucaena leucocephala

10--20 0.05 0.03 11 0.05 0.11 0.02 64 C. equisetifolia + E. robusta

10--20 0.05 0.04 13 0.04 0.13 0.01 64 C. equisetifolia + L. leucaena

10--20 0.06 0.03 13 0.05 0.10 0.03 64 Casuarina equisetifolia

10--20 0.05 0.03 12 0.04 0.10 0.02 64 Eucalyptus robusta

10--20 0.05 0.03 12 0.05 0.11 0.02 64 Leucaena leucocephala

20--30 0.04 0.03 9 0.03 0.11 0.02 64 C. equisetifolia + E. robusta

20--30 0.04 0.03 10 0.03 0.09 0.01 64 C. equisetifolia + L. leucaena

20--30 0.04 0.03 10 0.03 0.09 0.02 64 Casuarina equisetifolia

20--30 0.05 0.03 10 0.04 0.09 0.02 64 Eucalyptus robusta

20--30 0.05 0.03 9 0.03 0.09 0.01 64 Leucaena leucocephala

30--40 0.04 0.03 8 0.04 0.09 0.01 64 C. equisetifolia + E. robusta

30--40 0.05 0.03 10 0.03 0.09 0.01 64 C. equisetifolia + L. leucaena

30--40 0.04 0.03 9 0.03 0.08 0.01 64 Casuarina equisetifolia

30--40 0.05 0.03 10 0.03 0.09 0.02 64 Eucalyptus robusta

30--40 0.05 0.03 10 0.03 0.09 0.01 64 Leucaena leucocephala

Sodium (cmol/kg-l) 0--10 0.49 0.44 13 0.38 1.44 0.09 64 C. equisetifolia + E. robusta

0--10 0.72 0.48 13 0.60 1.61 0.17 64 C. equisetifolia + L. leucaena

0--10 0.52 0.44 15 0.29 1.34 0.10 64 Casuarina equisetifolia

0--10 0.44 0.43 13 0.24 1.49 0.11 64 Eucalyptus robusta

0--10 0.56 0.68 14 0.22 2.28 0.07 64 Leucaena leucocephala

10--20 0.52 0.35 11 0.38 1.11 0.11 64 C. equisetifolia + E. robusta
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

531

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

10--20 0.57 0.36 13 0.53 1.20 0.17 64 C. equisetifolia + L. leucaena

10--20 0.49 0.47 13 0.22 1.40 0.12 64 Casuarina equisetifolia

10--20 0.47 0.43 12 0.32 1.22 0.09 64 Eucalyptus robusta

10--20 0.69 0.72 12 0.34 2.17 0.08 64 Leucaena leucocephala

20--30 0.39 0.24 9 0.29 0.78 0.09 64 C. equisetifolia + E. robusta

20--30 0.39 0.24 10 0.38 0.92 0.15 64 C. equisetifolia + L. leucaena

20--30 0.36 0.42 10 0.23 1.47 0.05 64 Casuarina equisetifolia

20--30 0.20 0.12 10 0.15 0.36 0.08 64 Eucalyptus robusta

20--30 0.43 0.49 9 0.23 1.61 0.08 64 Leucaena leucocephala

30--40 0.36 0.27 8 0.27 0.97 0.12 64 C. equisetifolia + E. robusta

30--40 0.46 0.22 10 0.48 0.87 0.17 64 C. equisetifolia+ L. leucaena

30--40 0.40 0.49 9 0.19 1.67 0.12 64 Casuarina equisetifolia

30--40 0.23 0.13 10 0.22 0.47 0.06 64 Eucalyptus robusta

30--40 0.35 0.36 10 0.21 1.20 0.08 64 Leucaena leucocephala

Sodium (mg/g) 0--10 0.11 0.10 13 0.09 0.33 0.02 64 C. equisetifolia + E. robusta

0--10 0.17 0.11 13 0.14 0.37 0.04 64 C. equisetifolia+ L. leucaena

0--10 0.12 0.10 15 0.07 0.31 0.02 64 Casuarina equisetifolia

0--10 0.10 0.10 13 0.06 0.34 0.03 64 Eucalyptus robusta

0--10 0.13 0.16 14 0.05 0.53 0.02 64 Leucaena leucocephala

10--20 0.12 0.08 11 0.09 0.25 0.03 64 C. equisetifolia + E. robusta

10--20 0.13 0.08 13 0.12 0.28 0.04 64 C. equisetifolia+ L. leucaena

10--20 0.11 0.11 13 0.05 0.32 0.03 64 Casuarina equisetifolia

10--20 0.11 0.10 12 0.07 0.28 0.02 64 Eucalyptus robusta

10--20 0.16 0.17 12 0.08 0.50 0.02 64 Leucaena leucocephala

20--30 0.09 0.06 9 0.07 0.18 0.02 64 C. equisetifolia + E. robusta

20--30 0.09 0.06 10 0.09 0.21 0.04 64 C. equisetifolia+ L. leucaena
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

532

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

20--30 0.08 0.10 10 0.05 0.34 0.01 64 Casuarina equisetifolia

20--30 0.05 0.03 10 0.04 0.83 0.02 64 Eucalyptus robusta

20--30 0.10 0.11 9 0.05 0.37 0.02 64 Leucaena leucocephala

30--40 0.08 0.06 8 0.06 0.22 0.03 64 C. equisetifolia + E. robusta

30--40 0.11 0.05 10 0.11 0.20 0.04 64 C. equisetifolia + L. leucaena

30--40 0.09 0.11 9 0.04 0.38 0.03 64 Casuarina equisetifolia

30--40 0.05 0.03 10 0.05 0.11 0.01 64 Eucalyptus robusta

30--40 0.08 0.08 10 0.05 0.28 0.02 64 Leucaena leucocephala

Tuffaceous sandstone Entisols Mfs Calcium (mg/g) 0--30 4.47 1.27 2 4.47 5.36 3.57 65 Canovanas

0--30 5.86 0.37 2 5.86 6.12 5.60 65 Carolina

30--76 4.67 2.16 2 4.67 6.19 3.14 65 Canovanas

30--76 5.39 0.63 2 5.39 5.84 4.95 65 Carolina

Magnesium (mg/g) 0--30 0.42 0.05 2 0.42 0.46 0.39 65 Canovanas

0--30 0.28 0.03 2 0.28 0.30 0.26 65 Carolina

30--76 0.45 0.21 2 0.45 0.60 0.30 65 Canovanas

30--76 0.31 0.00 2 0.31 0.32 0.31 65 Carolina

Plutonic rocks Inceptisols Mfs Aluminum (cmol/kg-l) 0--15 9.50 4.23 12 8.94 16.12 4.63 66

15--30 9.60 3.93 12 9.24 15.07 5.02 66

Bulk density (g/cc) 0--15 0.65 0.08 2 0.67 0.79 0.49 66

15--30 1.02 0.12 12 1.07 1.17 0.76 66

Calcium (cmol/kg-l) 0--15 1.12 0.92 12 0.76 3.36 0.36 66

15--30 0.26 0.26 12 0.16 1.02 0.07 66

Calcium (mg/g) 0--15 0.22 0.18 12 0.15 0.67 0.07 66

15--30 0.05 0.05 12 0.03 0.20 0.01 66

Effective cation exchange capacity 0--15 11.98 3.83 12 1.52 17.42 7.16 66

{Ca+K+Mg(cmol/kg-1) 15--30 10.60 3.78 12 10.34 15.96 6.25 66
Table 9. Element concentration in soils and other soil characteristics by depth, geology, soil order, and life zone. Life zone codes are in Table 

1. 

533

total

Geology and Soil Life Depth

parent material order zone Element (cm) Mean Sd n Median Max Min Source Site/notes 

Iron (mg/g) 0--15 2.04 0.56 12 1.98 3.06 1.24 66

15--30 1.44 0.51 12 1.44 2.42 0.60 66

Magnesium (mg/g) 0--15 0.12 0.04 12 0.12 0.22 0.08 66

15--30 0.07 0.03 12 0.06 0.13 0.04 66

Magnessium (cmol/kg-1) 0--15 1.00 0.33 12 1.00 1.88 0.67 66

15--30 0.55 0.22 2 0.49 1.13 0.30 66

Manganese (mg/kg) 0--15 14 13 12 11 42 1 66

15--30 6 8 12 4 30 0 66

Nitrogen (%) 0--15 0.24 0.04 12 0.24 0.31 0.16 66

15--30 0.11 0.03 12 0.12 0.15 0.07 66

Organic matter (%) 0--15 5.84 0.84 12 5.70 7.66 4.78 66

15--30 2.60 0.42 12 2.60 3.16 1.73 66

pH (H2O) 0--15 4.09 0.23 12 4.13 4.51 3.79 66

15--30 4.12 0.19 12 4.08 4.39 3.78 66

pH (KCl) 0--15 3.60 0.20 12 3.59 3.91 3.25 66

15--30 3.63 0.15 12 3.65 3.82 3.37 66

Phosphorus (mg/kg) 0--15 14 4 12 15 22 9 66

15--30 5 1 11 5 7 2 66

Potassium (cmol/kg-l) 0--15 0.24 0.15 12 0.22 0.50 0.05 66

15--30 0.13 0.06 8 0.14 0.20 0.07 66

Potassium (mg/g) 0--15 0.09 0.06 12 0.09 0.19 0.02 66

15--30 0.05 0.02 8 0.05 0.08 0.03 66

Sodium (cmol/kg-l) 0--15 0.13 0.05 12 0.11 0.23 0.09 66

15--30 0.08 0.03 12 0.07 0.12 0.04 66

Sodium (mg/kg) 0--15 30 11 12 26 53 20 66

15--30 18 6 12 16 29 6 66