Paleocene Foraminifera of 

the Gulf Coastal Region 

of the United States 

and Adjacent Areas

By JOSEPH A. CUSHMAN

GEOLOGICAL SURVEY PROFESSIONAL PAPER 232

Descriptions and illustrations of smaller 

Foraminifera from the Gulf Coastal 

Region, Cuba, Central America, 

Haiti, and Trinidad

UNITED STATES GOVERNMENT PRINTING OFFICE, WASHINGTON : 1951
UNITED STATES DEPARTMENT OF THE INTERIOR

Oscar L. Chapman, Secretary

GEOLOGICAL SURVEY

W. E. Wrather, Director

For sale by the Superintendent of Documents, U. S. Government Printing Office 

Washington 25, D. C. - Price $1.75 (paper cover)
CONTENTS

Page Page

Abstract______--------------------_-----__----_-___ 1 Systematic descriptions—Continued

Introduction.___--_----___-________________-______- 1 Lagenidae _____________________-_____---------_ 13

List of loealities____-----------_-_-_-_-_-__--_-_-_-_ 1 Polymorphinidae-__-________--_-_------—— ____ 32

Systematic descriptions____-__-_-_-___-_-___.-____-_- 3 Nonionidae-___————————— ----- ———— ______ 37

Rhizamminidae_______-_-._--- —— -._ — ___ 3 Heterohelicidae___._ —.._- — _ — — _ — — — 37

Ammodiscidae-,—- — ---- — — _ — — ____ 4 Buliminidae————--_-- — — — — — — — 39

T ., ... . Ellipsoidimdae.-------------------------------- 45 Lituohdae__-__--------------_-------------_--- 4 ,, /..., ,_

_ , .., _ Rotaliidae____--_------_----_------------------ 47 Textulariidae—-— — __ — ___ — — — — — - 5 Amphisteginidae—— — — — — — — — — — — 56

Verneuilinidae— ______________ __ _ __________ 8 CM_idulinidae_____ _________ __._...._ — ... — _. 56

Valvulinidae__------------_-_-_____-___---__--- 9 Chilostomellidae— _____________________________ 58

Miliolidae-—---------------------------------- 10 Globigerinidae—------------------------------- 60

Ophthalmidiidae.-- _________________ ——________ 12 Globorotaliidae-_-_ ___--_ ____________________ 61

Trochamminidae- _______________________________ 12 Anomalinidae __________-______--_-__-------_--- 62

Placopsilinidae __--_-_-_-_--_---___-_-__-_-_____ 12 Index.________________________-_---_____-_---_---_ 69

ILLUSTRATIONS

PLATE 1. Rhizamminidae, Ammodiscidae, Lituolidae, Textulariidae_________________________________________ Following index

2. Textulariidae, Verneuilinidae, Valvulinidae, Miliolidae__________________________________________ d^

3. Miliolidae, Ophthalmidiidae, Trochamminidae, Placopsilinidae, Lagenidae__________________________ do

4-8. Lagenidae__________________________________________________________________________________ do

9. Lagenidae, Polymorphinidae, Buliminidae_____________________________________________________ do

10. Polymorphinidae-___________________________________________________________________________ do

11. Nonionidae, Heterohelicidae, Buliminidae---___________________________________________________ do

12. Buliminidae, Ellipsoidinidae-__________________________________________________________________ do

13. Lagenidae, Ellipsoidinidae, Rotaliidae__________________________________________________________ do

14-15. Rotaliidae._________________________________________________________________________________ do

16. Rotaliidae, Cassidulinidae, Chilostomellidae_____________________________________________________ do

17. Chilostomellidae, Globigerinidae, Globorotaliidae, Anomalinidae______-__-_____-_______-L-_______-_ do

18-19. Anomalinidae-______________________________________________________________________________ do20. Rotaliidae, Anomalinidae-____________________________________________________________________ do

21. Valvulinidae, Lagenidae, Anomalinidae___-________-_______________________________________-___- do

22. Anomalinidae-_________________________________-______________--____•______________-_______-- do

23. Rotaliidae, Anomalinidae-_____________-______________ v __________-_____-_____-________________ do

24. Rotaliidae, Cassidulinidae, Globorotaliidae_-_-_-_______________-_________________-_____.________ do

TABLES

Page 

TABLE 1. Distribution of Paleocene Rhizamminidae, Ammodiscidae, Lituolidae, Textulariidae, Verneuilinidae, Miliolida-?,

Ophthalmidiidae, Trochamminidae, and Placopsilinidae__ _______________________________________________ 12

2. Distribution of Paleocene Lagenidae__________________________________________________________________.._ 28

3. Distribution of Paleocene Polymorphinidae, Nonionidae, Heterohelicidae, Buliminidae, and EUipsoidinidae____. _ 44

4. Distribution of Paleocene Rotaliidae, Amphisteginidae, Cassidulinidae, Chilostomellidae, Globigerinidae, Globorotaliidae, and Anomalinidae--_________________________________________________________________________ 60

m

PALEOCENE FORAMINIFERA OF THE GULF COASTAL REGION OF THE UNITED STATES

AND ADJACENT AREAS

By JOSEPH A. CUSIIMAN l

ABSTRACT

This paper describes and figures the Foraminifera from the 

Paleocene of the Gulf Coastal Plain of the United States. The 

collections are largely from Texas, Arkansas, Mississippi, Alabama, and Tennessee. Records from related areas of Trinidad, 

Cuba, Haiti, and Central America are also included. Many of 

the species are excellent index fossils for study of the Paleocene. 

About 280 species and varieties are recorded and nearly all are 

illustrated.

INTRODUCTION

Some controversy exists as to what fossil material 

should be included in the Paleocene. In this report, 

well samples from Florida have been left out where the 

distribution of the species indicates latest Cretaceous 

rather than Paleocene age. Likewise, outcrop samples 

were left out where late Cretaceous species of Foraminifera, evidently redeposited, were associated with megafossils that definitely indicate Paleocene age. In the 

situation of well samples where the only fossils present 

are Foraminifera the age relations may also be obscured by the presence of redeposited fossils. To make 

the fauna more nearly complete, records from the West 

Indies region and Central America have been included.

Although many of the species of Foraminifera in the 

Paleocene are found also in the Upper Cretaceous and 

numerous others extend upward into the Wilcox group 

of the Eocene, many species remain that are excellent 

index fossils for the Paleocene. Many of these are 

very striking in their characters and should not be 

confused with species from other series or formations.

The work by Mrs. Helen JTeaime Plummer "Foraminifera of the Midway formation in Texas" (Texas 

Univ. Bull 2644, 1926 (1927), pp. 1-206, pis. 1-15, text 

figs. 1-13, table) is the pioneer work on this group. 

Later papers have been published on the Paleocene of 

other regions which have added much to the original 

fauna and undoubtedly many more species will be found 

as work progresses in different regions.

Many members of the U. S. Geological Survey have 

very kindly collected material for this work and a number of others have helped much. Their names will be 

found in the list of localities given here. Also, the 

collections of Dr. W. H. Deaderick, bequeathed to the

1 This report was received March 2, 1949. The author died on April 

16, 1949. Its editorial preparation for publication has involved no substantial changes from the text as left by the author.

U. S. National Museum, have added much to the faunas 

from Arkansas.

I wish to express my thanks to the many persons whohave collected material at my suggestion, to I ~. John, 

B. Reeside, Jr., for his help as chief of the Paleontology 

and Stratigraphy Branch, and to Miss Ruth Todd f OF 

her careful work in helping to prepare material, typing,, 

and in publishing several joint papers which have; 

added to the known faunas.

LIST OF LOCALITIES 2 

ALABAMA

PORTERS CREEK FORMATION, MATTHEWS LANDING MARL. 

MEMBER

1. Road cut on east slope to a branch in SW 14 sec12, T. 11 N., R. 9 E., Wilcox County, Air, F. S, 

MacNeil.

2. Road cut just south of Dixoii Creek, sec. 23, T. 13 

N., R. 6. E., Wilcox County, Ala, F. S. I lacNeiL

3. Foot of hill on south slope to Wolf Creek, probably 

in SE^NE^ sec. 20, T. 11 N., R. 12 E, Butler 

County, Ala. F. S. MacNeil.

4. On Alabama Highway 96, 10.2 miles northeast of 

Kimbrough, Wilcox County, Ala. C. G. Lalicker.

5. On Alabama Highway 96, 9.2 miles northeast of 

Kimbrough, Wilcox County, Ala. C. G. Lalicker.

6. North part of sec. 12, T. 12 N., R. 6 E., at Matthews 

Landing, 9 miles west and 2 miles north of Camden, Wilcox County, Ala. C. G. Laliclrer.

7. Naheola Landing on Tombigbee River, Choctaw 

County, Ala. T. W. Vaughan. U. S. G. S. 5647.

NAHEOLA FORMATION, COAL BLUFF MARL MEMBER

8. Creek bottom just west of the intersection of Caledonia, about 14 mile south of the center of sec. 

29, T. 11 N., R. 10 E., Wilcox County, Ala. F. S. 

MacNeil.

CLAYTON FORMATION, CHALYBEATE LIMESTONE MEMBER

9. Chalk overlying Ostrea pulaskensis zone, U. S. 

Highway 80, south of Sucarnoochee Creek, y2 

mile southwest of Livingston, Sumter County, 

Ala. L. W. Stephenson and W. H. Mor roe.

3 The localities are referred to by number in the individual descriptions 

and in the faunal tables.
PALEOCENE FORAMINHFERA OP GULF COASTAL REGION

CLAYTON FORMATION. ( HORIZON=BASE OF PORTERS CREEK 

CLAY OF WESTERN ALA.)

10. Glauconite bed 1 to iy2 feet thick between hard 

limestone of the Clayton formation and overlying 

fullers earth. Old fullers earth mine, sec. 23, 

T. 10 N., R. 25 E., just east of Pratts, Barbour 

County, Ala. F. S. MacNeil.

CLAYTON FORMATION, PINE BARREN MEMBER. (NEAR TOP)

11. 130 feet above water level on road leading down 

to ferry at Old Canton Landing, Alabama River, 

Wilcox County, Ala. L. W. Stephenson and 

W. H. Monroe.

12. Top of fullers earth zone, immediately underlying 

Turritella rock, SEfySEty sec. 18, T. 11 N., 

R. 14 E., Butler County, Ala. F. S. MacNeil.

TENNESSEE

TORTERS CREEK CLAY. ( PROBABLY IN TIPPAH SAND

LENTIL)

13. Cut on Southern Railway 200 feet east of milepost 

481, V/2 miles east of Middleton, Hardeman 

County, Tenn. U. S. G. S. 6495.

14. Cut on Southern Railway 260 feet east of milepost 

481, 11/2 miles east of Middleton, Hardeman 

County, Tenn. L. W. Stephenson.

MISSISSIPPI 

CLAYTON FORMATION, CHALYBEATE LIMESTONE MEMBER

15. Within 5 feet of top of Cretaceous, U. S. Highway 

45, 2 miles north of Scooba, Kemper County, 

Miss. F. F. Mellen.

16. South valley wall of Lynn Creek, sec. 22, T. 16 N., 

R. 15 E., Noxubee County, Miss. E. H. Rainwater and F. F. Mellen.

PORTERS CREEK CLAY

17. Field, 1,200 feet east of M. & O. Railroad, 3 miles 

north of Scooba, Kemper County, Miss. U. S. 

G. S. 6493.

ARKANSAS 

MIl>i\..Y FORMATION, LOWER PART

18. North bank of McNeil Creek,

sec. 10, T. 2 S., R. 15 W., at the north edge of 

Benton, Saline County, Ark. M. Gordon. Midway loc. 14.

19. North bank of McNeil Creek, about 600 feet west 

of Highway TO, NW^NE^NEVi sec. 10, T. 2 S., 

R. 15 W., Saline County, Ark. M. Gordon. Midway loc. 13.

19A. Ravine alongside lumber yard at Benton, Saline 

County, Ark. 100 feet north of the south line

of the SE14SW1/4SW1/4 sec. 11, T. 2 S., R. 15 W. 

Coll. first by R. A. Schmidt and J. A. Gardner. 

Additional material collected by M. Gordon and 

M. W. Ellis. Midway loc. 4.

20. West bank of Trace Creek, SEV4SY^V4 sec. 30, T. 2 

S., R, 15 W., Saline County, Ark CoU. first by 

R. A. Schmidt and J. A. Gardner. Additional 

material collected by M. Gordon and M. W. Ellis. 

Midway loc. 7.

21. Bed of McNeil Creek 50 to 75 yards upstream from 

Highway 70, SE cor. sec. 3, T. 2 S., R. 15 W., 

Saline County, Ark. M. Gordon and J. I. Tracey. 

Midway loc. 10.

22. Clay from just below limestone ledge in gully on 

the south side of U. S. Highway 67, 1.9 miles 

beyond bridge over Salt Creek, in the SEi^NWVi 

sec. 20, T. 2 S., R. 15 W., Saline County, Ark. 

R. P. Bryson, M. Gordon, M. W. Ellis, R. A. 

Schmidt, and J. A. Gardner. Midway loc. 5.

23. Upper bed on east bank of valley of Fourche Creek, 

about 1/2 mile north of Terry ton, 8.000 feet east of 

U. S. Highway 67 and 70, sec. 8, T. 1 S., R. 13 

W., Pulaski County, Ark. R. A . Schmidt and 

J. A. Gardner. Midway loc. 3.

MIDWAY FORMATION, UPPER FART

24. (Near base of upper part) Gully 400 to 500 feet

' south of NE cor. NW% sec. 16, T. 1 N., R. 12 W.,

600 to 700 feet up gully from Midway loc. 1,

southwest part of Little Rock, F ilaski County,

Ark. M. W. Ellis. Midway loc. 15.

25. (Base of upper part) about 1,000 feet south of 

Roosevelt Road viaduct, 200 feet east of railroad 

tracks in small gully heading east, just upstream 

from small abandoned bridge, NE^NW^ sec. 

16, T. 1 N., R. 12 W., Little Rock, Pulaski County, 

Ark. M. W. Ellis, R. A. Schmidt, and J. A. 

Gardner, Midway loc. 1.

MIDWAY FORMATION

26. Side of bank, north side of Highway 67, 8 miles 

southwest of Benton, Saline County, Ark. W. H. 

Deaderick. No. 324.

27. 200 yards southwest of Camp Marion on Highway 

67 between Pulaski-Saline County line and Little 

Rock, turn southeast and go through field and 

over suspension bridge to Bluffon Creek, Pulaski 

County, Ark. W. H. Deaderick. No. 342.

28. (At Paleozoic contact, Polk Creek shale) Natural 

erosion on Highway 67 about a mile south of 

Collegeville just beyond large sign, west side of 

road, Pulaski County, Ark., W. H. Deaderick. 

No. 344.

29. Roadside ditch, southeast side of road, Highway 

67, at Beirne Cross Road, 3.2 miles south of
RHIZAMMINIDAE

Gurdon railroad station, Clark County, Ark. 

W. H. Deaderick. No. 363.

"0. Eoadside ditch, northwest side of road, Highway 

67, at Beirne Cross Road, 3.2 miles south of Gurdon railroad station, Clark County, Ark. W. H. 

Deaderick. No. 387.

"1. Eoadside ditch, east side, Highway 67, 0.7 mile 

north of Curtis Cross Road, Clark County, Ark. 

W. H. Deaderick. No. 377.

WILLS POINT ( ?) FORMATION

?2. Deep roadside ditch, southeast side of road, Highway 67, 3.5 miles northeast of railroad station in 

Gurdon, Clark County, Ark. W. H. Deaderick. 

No. 364.

TEXAS 

WILLS POINT FORMATION

33. 1.9 miles north of Lockhart on road to Austin, 

Travis County, Tex. U. S. G. S. 10793.

34. Gully in jog in secondary road 4.6 miles west and 

a little north of Lockhart, Caldwell County, Tex. 

U. S. G. S. 10794.

35. Upper bed of Dry Brushy Creek, 6 miles south of 

Thrall, Williamson County, Tex. J. A. Gardner. 

U. S. G. S. 10796.

WILLS POINT FORMATION, MEXIA MEMBER

?6. About 2 miles north and slightly west of center of 

Mexia, Limestone County, Tex. C. G. Lalicker.

KINCAID FORMATION

37. On west or downthrow side of fault, 2 miles northwest of Lone Oak, Hunt County, Tex. L. W. 

Stephenson. U. S. G. S. 10797.

38. 3 + miles above bridge over Cedar Creek on AustinBed Rock road, Bastrop County, Tex. U. S. G. S. 

10786.

MIDWAY GROUP

39. Exposure in creek bed on northeast-southwest 

country road, 1.1 miles due east of Honest Ridge 

School, and 2.5 miles due north of the cross roads 

in Thelma, Limestone County, Tex. H. J. 

Plummer. Sta. 697.

40. Shallow ditch at road corner southeast of new 

Corsicana reservoir on road to Mildred, Navarro 

County, Tex. H. J. Plummer. Sta. 23.

41. Exposure in deep roadside ditch on steep hill 0.2 

mile east of road corner in north end of town of 

Commerce, on highway to Paris, Hunt County, 

Tex. H. J. Plummer. Sta. 3.

42. Clay pit, Mexia Brick Works, about 1 mile west of 

the town of Mexia, Limestone County, Tex. 

H. J. Plummer. Sta. 46.

43. Base of high bluff on west side of Colorado River 

between the Travis-Bastrop County line and the

mouth of Dry Creek, Bastrop County, Tex. 

H. J. Plummer. Stas. 67 and 379.

44. 4.6 miles north of Fentress, Caldwell County, Tex. 

L. W. Stephenson.

45. Branch on the F. Goericke farm, across fo^d 011 

Wilbarger Creek, iy2 miles south of Littig, 

Travis County, Tex. U. S. G. S. 10787.

46. 800 yards above bridge over Wilbarger Creek, near 

Travis County line, Tex. J. A. Gardner. U. S. 

G. S. 10788.

47. On road southwest corner of F. V. Hughes property, east Alvarado Survey, Van Zandt County, 

Tex.

48. On creek, west line W. W. Pitts property, east Alvarado Survey, Van Zandt County, Tex.

49. Dry Creek, 3% miles above bridge on first road east 

of Travis, Bastrop County, Tex. J. A. Gardner. 

U. S. G. S. 10795.

50. Butler salt dome, 1,000 feet southeast of Lalreport 

store, Freestone County, Tex. L. W. Steiohenson. U. S. G. S. 10789.

51. Right bank of Solomon's Creek, about % mile east 

of its junction with Wilbarger Creek, 5.8 miles 

by road south-southwest of railroad crossing 

south of Elgin, Bastrop County, Tex. H. J. 

Plummer. Sta. ll-T-3.

52. On south side of Foggyhead Creek in Smith'? pasture and about 0.15 mile west of the bridge on the 

Kerens-Round Prairie road, 3.8 miles by road 

south-southeast of the railroad station in Kerens, 

Navarro County, Tex. H. J. Plummer. Sta. 

174-T-6.

SYSTEMATIC DESCRIPTIONS

Family RHIZAMMINIDAE

Genus BATHYSIPHON M. Sars, 1872

Bathysiphon eocenicus Cushman and G. D. Hanna

Plate 1, figures 1, 2

Bathysiphon eocenica Cushman and G. D. Hanna, v California

Acacl. Sci. Proc., 4th ser., vol. 16, p. 210, pi. 13, fig? 2, 3,

1927. 

Cushman and McMasters, Jour. Paleontology, vol. 10, p. 508,

pi. 74, fig. 1,1936. 

Staesche and Hiltermann, Reichs. Bodenforschung Abhandl.,

n. ser., No. 201, pi. 37, figs. 4, 5,1940. 

Cushinan and Siegfus, San Diego Soc. Nat. History Trans.,

vol. 9. no. 34, p. 400, pi. 15, fig. 1, 1942. 

Kelley, Am. Assoc. Petroleum Geologists Bull., vol. 27, p. 11

(list), 1943.

Curran, idem, pp. 1378, 1381 (lists), 1943. 

Martin, Stanford Univ. Publ., Univ. Ser., Geol. Sci., vol. 3,

no. 3, p. 9 (list), 1943. 

Cushinan and Simonson, Jour. Paleontology, vol. 18, p. 193,

pi. 30, fig. 1, 1944. 

Cushman and Stone, Cushman Lab. Forarn. Research Special

Pub. 20, p. 2, pi. 1, fig. 1,1947.
PALEOCENE FORAMINHFERA OF GULF COASTAL REGION

Test elongate, compressed cylindrical; wall of fine 

white amorphous material, fairly smooth on the exterior; inferior tubular chamber distinct. Diameter 

of crushed specimens 0.62-0.75 mm.; diameter of tubular 

chamber averaging about 0.25 mm.

This species is widely recorded in the Eocene of California and also recorded from the Eocene of Peru and 

the lower Eocene of northwest Germany.

The Paleocene specimens are very similar to the types 

of this species and are unlike Cretaceous species that 

may be found redeposited in the Paleocene at some 

localities.

Midway group. Texas, Navarro County (40) ; Limestone 

County (42).

Family AMMODISCIDAE

Genus AMMODISCUS Reuss, 1861

Ammodiscus cf. A. cretaceus (Reuss) Cushman

Plate 1, figure 3

Specimens were found only at station 42 and may 

possibly be reworked from Upper Cretaceous deposits. 

One of the specimens is figured.

The forms figured by Mrs. Plummer as "Ammodiscus 

incertus (D'Orbigny)" (Texas Univ. Bull. 2644, p. 63, 

pi. 13, figs, la-d, 1927) are probably the same as the 

forms here referred to A. cf. A. cretaceus.

Family LITUOLIDAE

Genus HAPLOPHRAGMOIDES Cushman, 1910 

Haplophragmoides cf. H. excavata Cushman and Waters

Single specimens from two localities seem to belong 

in this species and may represent material reworked 

from the Upper Cretaceous. They are, however, more 

compressed than the types.

The somewhat distorted specimen figured by Mrs. 

Plummer as "Haplophragmoides cananensis (D'Orbigny)" (Texas Univ. Bull. 2644, p. 65, pi. 3, figs, la, 

b, 1927) may possibly be the same as those here referred 

to H. cf. H. excavata.

Midway formation. Arkansas, Clark County (31). 

Midway group. Texas, Limestone County (42).

Genus AMMOBACULITES Cushman, 1910

Ammobaculites expansus Plummer

Plate 1, figures 5-7

Ammobaculites expansus Plummer, Texas Univ. Bull. 3201, p.

65, pi. 5, figs. 4-6, 1933.

Test broadly elongate, very coarsely arenaceous and roughly 

finished, composed of angular quartz grains bound by a reddish 

and insoluble cement up to the apertural rim, which on many 

specimens is white; about the first 12 chambers arranged in a 

tight and very strongly compressed and slightly umbilicate coil 

of two convolutions; final two or three chambers arranged in 

rectilinear succession and also strongly compressed; sutures 

very obscure except between last two or three chambers of a fully

mature test where they are somewhat depressed; aperture a 

narrow elongate opening at the end of the final chamber. 

Length of holotype 0.58 mm.; breadth 0.40 mm.; thickness of 

final chamber 0.06 mm.

Like [A. midwayensis] this is a characteristic species of the 

uppermost upper Midway of Texas and often the only species 

present.—Plummer.

The type specimens came from the Paleocene, from 

north bank of Solomon's creek 6% miles south-southwest of Elgin, Bastrop County, Tex.

We have specimens of this species from the type 

locality, sent by Mrs. Plummer, but have found no specimens in our material.

Ammobaculites midwayensis Plummer 

Plate 1, figures 8-12

Ammobaculites midioayensis Plummer, Texos Univ. Bull. 3201,

p. 63, pi. 5, figs. 7-11,1933.

Test elongate, slender, very coarsely arenaceous and roughly 

finished, composed of angular quartz grains bound by a reddish 

insoluble cement up to the last chamber which is conspicuously 

whiter than the rest of the test; early 5 to 7 chambers arranged 

in a tight, compressed coil of a little more tl !?.n one convolution 

and with rounded periphery; succeeding chambers rectilinear, 

about as high as broad, and varying from uncompressed to 

considerably compressed; sutures visible ard very slightly depressed on some tests, but in general obscure or even hidden 

by the texture of the wall; aperture terminal and round or 

somewhat elongate, depending on the degree of compression 

due to forces of fossilization of 1 a somewl at plastic original 

test. Length of holotype 0.65 mm.; diameter of coil 0.20 mm.; 

width of rectilinear chambers 0.18 mm.—Plummer.

The types of this species are from the Paleocene, from 

north bank of Solomon's Creek near its junction with 

Wilbarger Creek, 6% miles south-southwest of Elgin, 

Bastrop County, Tex.

This seems to be a characteristic spe?ies of the upper 

50 to 75 feet of the Midway group in outcrops and makes 

a good index fossil for this part of the section in Texas.

Ammobaculites paleocenicus Cufhman

Plate 1, figures 13, 14 

Ammobaculites paleocenicus Cushman, Cushman Lab. Foram.

Research Contr., vol. 23, p. 77,1947. 

Ammobaculites midwayensis Cushman (no*: Plummer) idem,

vol. 16, p. 52, pi. 9, figs. 1, 2, 1940.

Test in the early stages close-coilec1 , planispiral, in 

the adult uncoiling and the chambers in a rectilinear 

series; chambers distinct, inflated, 6 to 9 in the coiled 

portion, only 2 or 3 in the uncoiled part, later ones circular in section; sutures distinct, depressed; wall finely 

arenaceous with occasional larger fragments, rather 

smoothly finished; aperture somewhat arcuate, narrow, 

elongate. Length up to 2.70 mm.; breadth 1.65 mm.; 

thickness 0.85 mm.

The types are from the Paleocene, U. S. Highway 80, 

S. of Sucarnoochee Creek, y2 mile SW. of Livingston, 

Sumter County, Ala.

Clay ton formation, Chalybeate limestone r1 ember. Alabama, 

Sumter County (9). Mississippi, Noxube^ County (16).
TEXTULARHDAE

Genus AMMOMARGINULINA Wiesner, 1931

Ammomarginulina sp.

Plate 1, figure 4

/.mmotnarginulina sp. Cushman and Renz, Cushman Lab. 

Foram. Research Contr., vol. 18, p. 3, pi. 1, fig. 1, 1942.

A single specimen figured here has an arenaceous 

test with the early portion close-coiled and the lastformed chambers uncoiled in a rectilinear series and 

much compressed. This slightly resembles Ammo- 

<" iculites expansus Plummer from the Paleocene of 

' T'exas but is not the same.

The present specimen is from the Paleocene 'Soldado 

r*ock formation, Soldado Kock, Trinidad, B. W. I.

Genus LITUOLA Lamarck, 1804

Lituola erecta Mellen and Gault

Plate 1, figure 17

erecta Mellen and Gault, Am. Midland Naturalist, vol. 

22, p. 472, text figs. 2, 4 (middle row), 1939.

The test is long, crosier-shaped, and slightly compressed in 

the early close-coiled portion. The latter portion is very abruptly uncoiled and erect in the adult forms. The numerous 

labyrinthic chambers have very irregular, rough interiors, the 

rnmber in the coil varying in the microspheric and meglospheric forms and in the young and adult stages. The sutures 

, re indistinct and but slightly depressed. The wall is arenaceous, well cemented, and rough on the exterior. The aperture 

i"? terminal and multiple or cribrate, and occupies all the 

rpertural face.

Maximum length 9 mm. ; width of coil 2.5 mm. ; thickness 2

•urn. — Mellen and Gault.

The types are from the Paleocene Clayton formation, 

outcrops behind the J. W. Martin home in the SE*4 

rec. 31, T. 19 K, B. 14 E., Oktibbeha County, Miss. 

'""he species is recorded from the Clayton formation in 

day, Oktibbeha, and Noxubee Counties of Mississippi.

"atuola erecta Mellen and Gault var. distincta Mellen and Gault

Plate 1, figure 18

Lituola erecta distincta Mellen and Gault, Am. Midland Naturalist, vol. 22, p. 473, text figs. 3, 4 (top row), 1939. 

The subspecies differs from L. erecta in the more evolute 

soiling, in the more depressed sutures, and in size, having a 

maximum length of 5 mm., a width of 2.5 mm., and a thickness 

)f 1 mm. The interiors of the chambers are smooth and the 

"•utures are distinct. — Mellen and Gault.

The types are from the Paleocene, from a 5- by 40- 

: nch sandy and calcareous lens in the Porters Creek 

"lay 5 or 10 feet above the Clayton formation, from

->, road cut <Hi the south valley wall of Lynn Creek, sec. 

22, T. 16 K ; E. 15 E., Noxubee County, Miss. It is 

"scorded from Clay County, Miss., and from the basal 

of the Midway formation north of Hope, Ark.

Family TEXTULARIIDAE

Genus SPIROPLECTAMMINA Cushman, 1927

Spiroplectammina rossae Plummer

Plate 1, figures 15, 16

Spiroplectammina rossae Plummer, Texas Univ. Bull. 3301, p. 

66, pi. 5, figs. 1-3, 1933.

Test very small, elongate, expanding gradually fro^ the 

somewhat bluntly pointed initial extremity, strongly compressed, thickest along the axis, white, composed of ve~y fine 

siliceous particles bound by an insoluble cement, thin shelled; 

early four or five chambers arranged in so minute a co;l that 

the average peripheral outline hardly suggests this stricture; 

later chambers typically biserial, narrow, slowly lengthening 

through early maturity and still more slowly through late maturity, sloping from the axis on each side at an angle of about 

45 degrees and gently curved downward along the outer edge; 

sutural limbations strongly elevated on each side of the center 

but not merged, and tapering rapidly toward the peripheral 

margin, which is very narrowly rounded and unflanged; aperture a rather high arch at the base of the septal face. Length 

of holotype, 0.32 mm.; width 0.19 mm.; thickness 0.09 mm.

This is another species that seems to be characteristic of the 

uppermost Midway beds and seems a definite guide in determining the Paleocene-Wilcox contact.—Plummer.

The types are from the Paleocene, from bank on 

Foggyhead Creek, 3.8 miles by road southeast of Kerens 

railroad station, and a few feet below the overlying 

Wilcox groups, Navarro County, Tex.

Spiroplectammina plummerae Cushman 

Plate 1, figures 19, 20

Spiroplectammina plummerae Cushman, Maryland Dept. Geol.,

Mines and Water Resources, Bull. 2, p. 226, pi. 16, fig. 2,

1948. 

Textularia carinata D'Orbigny, var. expansa Plumme~ (not

Spiroplectammina expansa LeRoy, 1941), Texas Univ. Bull.

2644, p. 67, pi. 3, fig. 3,1927.

Test somewhat elongate, much compressed, expanding rather rapidly from the acute initial end; periphery 

acute, somewhat lobulate, sometimes slightly car'nate; 

chambers numerous, early ones planispiral, later ones 

biserial, rather rapidly increasing in size and 1 °dght 

as added; sutures distinct, depressed, gently curved and 

somewhat oblique; wall finely arenaceous, smoothly finished ; aperture a low, rather broad opening at thQ- base 

of the inner margin of the last-formed chamber, which 

is slightly depressed. Length 0.40-0.65 mm.; breadth 

0.30-0.40 mm.; thickness 0.18-0.20 mm.

The types are from the Paleocene, upper pr.rt of 

Midway group, 2^4 miles southeast of Corsican?,, Navarro County, Tex. It has been recorded from the 

Eocene of well samples from Maryland.

Porters Creek formation, Matthews Landing marl member. 

Alabama, Butler County (3).

915259—31———2
6 PALEOC'ENE FORAMINHFERA OF GULF COASTAL REGION

Clayton formation, Chalybeate limestone member. Mississippi,

Noxubee County (16).

Midway formation, lower part. Arkansas, Saline County (20). 

Midway formation. Arkansas, Saline County (26). 

Midway group. Texas, Limestone County (42). 

Wills Point formation, Mexia member. Texas, Limestone

County (36).

Spiroplectammina wilcoxensis Cushman and Ponton 

Plate 1, figures 21-23

Spiroplectammina icilcoxensis Cushman and Ponton, Cushman

Lab. Foram. Research Contr., vol. 8, p. 51, pi. 7, fig. 1,

1932. 

Cushman and Garrett, idem, vol. 15, p. 78, pi. 13, figs. 1, 2,

1939. 

Toulmin, Jour. Paleontology, vol. 15, p. 571, pi. 78, fig. 1,

1941. 

Cushman, Am. Jour. Sci., vol. 242, p. 8, pi. 1, figs. 1, 2, 1944;

Cushman Lab. Foram. Research Contr., vol. 20, p. 19, pi.

3, figs. 26, 27; p. 29, pi. 5, fig. 1, 1944.

Test broad, compressed; periphery subacute; early 

chambers coiled; chambers distinct, low and broad in 

the early portion, gradually and rather regularly increasing in height as added, slightly overlapping, later 

ones slightly! inflated; sutures distinct, slightly depressed especially in the later portion, gently curved, 

making a very slight angle with the horizontal; wall 

arenaceous but rather smoothly finished; aperture a 

low curved opening at the base of the apertural face. 

Length of holotype 0.45 mm.; breadth 0.25 mm.; thickness 0.12 mm.

The types are from the Wilcox group of Eocene age, 

railroad cut, 1 mile north of Ozark, Ala.

Naheola formation, Coal Bluff marl member. Alabama, Wilcox

County (8). 

Clayton formation, Pine Barren member. Alabama, Butler

County (12).

Porters Creek clay. Tennessee, Hardeman County (13, 14). 

Midway formation, lower part. Arkansas, Saline County (22). 

Midway group. Texas, Bastrop County (43).

Spiroplectammina laevis (Roemer) Cushman var. cretosa Cushman

Plate 1, figure 24

Spiroplectammina laevis (Roemer) Cushman, var. cretosa Cushman, Cushman Lab. Foram. Research Contr., vol. 8, p. 87, 

pi. 11, fig. 3,1932.

Jennings, Bull. Am. Paleontology, vol. 23, no. 78, p. 12, pi. 1, 

fig. 2,1936.

Cushman, Cushman Lab. Foram. Research Contr., vol. 16, 

p. 52, pi. 9, fig. 3,1940.

Cushman and Todd, idem, vol. 18, p. 25, pi. 5, fig. 1, 1942.

Kline, Mississippi Geol. Survey Bull. 53, p. 13, pi. 1,- fig. 1, 

1943.

Cushman, Cushman Lab. Foram. Research Contr., vol. 20, 

p. 2, pi. 1, fig. 4; p. 84, 1944.

Cushman and Deaderick, Jour. Paleontology, vol. 18, p. 329, 

pi. 50, fig. 6,1944.

Cushman and Todd, Cushman Lab. Foram. Research Contr., 

vol. 22, p. 46, pi. 7, fig. 1,1946.

Cushman, U. S. Geol. Survey Prof. Paper 206, p. 27, pi. ? 

figs. 1-3, 1946.

Test tapering, usually somewhat longer than broad 

the greatest breadth toward the apertural end, peri 

phery subacute; apertural end only slightly rounded 

broad in end view, tapering rapidly to the subacute 

periphery; chambers with the early portion coiled, lateT 

biserial, distinct, the margin of the apertural face distinctly raised, giving a series of raised ridges at the 

suture lines and forming a raised zigzag line along the 

center of the test; wall finely arenaceous, stout, ncj 

usually collapsed; aperture a low opening on the innei 

margin, of the apertural face with the peripheral portion of the face extending forward so that the aperture 

itself is in a reentrant. Length up to 0.65 mm.; breadtr 

0.45 mm.; thickness 0.25 mm.

The types of this variety are from the upper part o: 

the Taylor marl (Upper Cretaceous), 5.1 miles froir 

Josephine along the highway to Nevada, Collin County 

Tex. 

Clayton formation, Chalybeate limestone member. Alabama

Sumter County (9).

Porters Creek clay. Mississippi, Kemper County (17). 

Porters Creek formation, Matthews Landing marl member

Alabama, Choctaw County (7).

Midway formation, lower part. Arkansas Saline County (20) 

Midway formation, upper part. Arkansas, Pulaski County (25) 

Midway formation. Arkansas, Saline County (26) ; Pulask'

County (27).

Spiroplectammina mexiaensis Lalicker 

Plate 1, figures 25, 26

Spiroplectammina mexiaensis Lalicker, C ishman Lab. Foram, 

Research Contr., vol. 11, p. 43, pi. 6, figs. 5, 6, 1935.

Test elongate, much compressed, sides parallel, planispiral 

portion broader £han remainder of test, periphery short, axial 

portion of test high, ridge-like, depressed area along each 

side near peripheral margin; chambers numerous, initial 4 or 

5 coiled, later biserial, low and broad, commonly depressed; 

sutures distinct, usually somewhat liml ?te because of depressed chambers, straight and oblique; wfll finely arenaceous, 

composed of very fine siliceous and calcareous fragments with 

considerable cement, rather smoothly finished; aperture in 

early biserial portion a low, narrow slit at the inner margin 

of the last-formed chamber, in a deep reentrant, in adult form 

the reentrant tends to close, causing the aperture to be distinctly above the inner margin of the chamber. Length of 

holotype 0.60 mm.; width 0.20 mm.; tl ickness 0.11 mm.— 

Lalicker.

The types are from the Paleocene Mexia member of 

the Wills Point formation, about 35 feet above top of 

the Tehuacana member of the Kincaid formation, from 

steep bank along eastern side of a tribitary flowing due 

north into Tehuacana Creek, about 2 miles in a direct 

line north and slightly west of the centergof the town 

of Mexia, about *4 niile west of railroad, Limestone 

County, Tex.

This is a rather common species in the Wills Point 

formation.
TEXTULARIIDAE

Spiroplectammina paleocenica Cushman 

Plate 1, figure 27

Spiroplectammina, paleocenica Cushman, Cushman Lab. Forarn. 

Research Contr., vol. 23,'p. 81, pi. 17, fig. 17, 1947.

Test very small, evenly tapering from the slightly 

rounded initial end to the greatest width at the apertural end, central portion thick and rounded, tapering 

rapidly to the subacute periphery, apertural end truncate; chambers of the early portion planispiral, later 

biserial, increasing slowly in height as added, not inflated; sutures fairly distinct, not depressed, nearly 

straight to slightly convex upward; wall finely arenaceous, smoothly finished; aperture a slight, low opening 

at the base of the inner margin of the last-formed 

chamber. Length 0.35-0.43 mm., breadth 0.22-0.25 mm., 

thickness 0.20-0.23 mm.

The types are from the Paleocene, road cut on east 

slope to a branch in SW % sec. 12, T. 11 N., R. 9 E., Wilcox County, Ala. (1).

Porters Creek clay. Mississippi, Kemper County (17). 

Porters Creek formation, Matthews Landing marl member.

Alabama, Wilcox County (1, 5). 

Midway group. Texas, Bastrop County (43).

Genus TEXTULARIA Defrance, 1824 

Textularia midwayana Lalicker

Plate 1, figures 28-30

Textularia midwayana Lalicker, Cushman Lab. Foram. Research Contr., vol. 11 p. 49, pi. 6, figs. 7-9, 1935. 

Test elongate, about twice as long as broad, early portion 

rapidly expanding, later portion with sides nearly parallel, 

almost circular in top view, periphery broadly rounded; chambers numerous, somewhat wider than high, except apertural 

chambers whose width and height are about equal, slightly inflated ; sutures distinct, depressed, straight, and in a horizontal 

position; wall rather coarsely arenaceous, composed of fine and 

coarse sand grains with considerable cement, somewhat roughened; aperture a small, comparatively high arched opening at 

the inner margin of the last-formed chamber, in a shallow 

reentrant. Length of holotype 0.73 mm.; greatest width 0.30 

mm.; thickness 0.24 mm.—Lalicker.

The types are from the Paleocene, uppermost part of 

the Midway group, 0.15 mile west of bridge on KerensRound Prairie road, 3.8 miles by road south-southeast 

of railroad station in Kerens, Navarro County, Tex., 

where it is frequent.

Midway formation, lower part. Arkansas, Saline County (20,

22). 

Midway formation. Arkansas, Saline County (26) ; Pulaski

County (27). 

Midway group. Texas, Bastrop County (43).

Textularia midwayana Lalicker var. pansa Cushman 

Plate 2, figure 1

Textularia midicayana Lalicker var. pansa Cushman, Cushman 

Lab. Foram. Research Contr., vol. 20, p. 30, pi. 5, fig. 2,1944.

Variety differing from the typical in the somewhat 

broader and more tapering form, the more compressed

test, and the flatter and only slightly convex apertural 

face.

The types of this variety are from the Paleocene Coal 

Bluff marl member of the Naheola formation, creek 

bottom just west of store at Caledonia, about V4 milesouth of center of sec. 29, T. 11 N., R. 10 E., Wilcox 

County, Ala. (8).

Naheola formation, Coal Bluff marl member. Alabama, Wilcox

County (8). 

Midway formation. Arkansas, Saline County (26).

Textularia plummerae Lalicker 

Plate 2, figures 2, 3

Textularia plummerae Lalicker, Cushman Lab. Foram. Research

Contr., vol. 11, p. 50, pi. 6, fig. 10, 1935. 

Glaessner, Moscow Univ. Problems of Paleontology, vol.

2-3, p. 364, 1937. 

Cushman, Cushman Lab. Foram. Research Contr., vol. 16,.

p. 52, pi. 9, fig. 4, 1940. 

Davis, Jour. Paleontology, vol. 15, p. 151, pi. 25, figs. 8, 9,.

1941.

Toulmin, idem, vol. 15, p. 572, pi. 78, fig. 3, 1941. 

Textularia eocenica Plumnier (not Giimbel), Texas Univ. BulL

2644, p. 67, pi. 3, fig. 2, 1927.

Test very elongate, tapering, somewhat compressed, periphery 

broadly rounded; chambers numerous, distinct, width and height 

almost equal, somewhat inflated; sutures distinct, moderately 

depressed, straight, nearly horizontal; wall coarsel^ arenaceous, composed of fine and coarse sand grains with a moderate 

amount of cement, roughly finished; aperture a rather low 

arched slit at the base of the last-formed chamber ir a slight 

reentrant in the septal face. Length of holotype 084 mm.; 

greatest width 0.27 mm.; thickness 0.21 mm.—Lalicker.

The types are from the Paleocene Mexia member of 

the Wills Point formation, about 35 feet above the 

Tehuacana member of the Kincaid formation, from a 

steep bank along the east side of a tributary flowing due= 

north into Tehuacana Creek, about 2 miles in a direct 

line north and slightly west of the center of the town 

of Mexia, about % mile west of railroad, Limestone 

County, Tex. 

Clayton formation, Chalybeate limestone member. Alabama,,

Sumter County (9).

Wills Point formation, Mexia member. Texas, LimestoneCounty (36). 

Midway group. Texas, Limestone County (42)

Textularia plummerae Lalicker var. arkansasana Cushman 

Plate 2, figures 4, 5

Textularia plummerae Lalicker var. arkansasana, Cushman,. 

Cushman Lab. Foram. Research Contr., vol. 23, p. 81, pi. 17,. 

figs. 18, 19, 1947. 

Textularia plummerae Lalicker var. Cushman and Told, idem,.

vol. 22, p. 46, pi. 7, fig. 2, 1946.

Variety differing from the typical in the greaterthickness and enlargement of the chambers at the apertural end and the much coarser arenaceous wall.

The types are from the Paleocene, side of bank, norths 

«ide of Highway 67, 8 miles southwest of Bentor, Saline 

County, Ark. (26). 

Midway formation, lower part. Arkansas Saline County (22).
8 PALEOCENE F6RAMINiIFERA OF GULF COASTAL'REGION

Midway formation, upper part. Arkansas, Pulaski County (25). 

Midway formation. Arkansas, Saline County (26).

Textularia portenta Cushman 

Plate 2, figure 6

Textularia portenta Cushman, Cushman Lab. Foram. Research 

Contr., vol. 23, p. 81, pi. 17, fig. 16,1947.

Test small, initial end subacute, increasing rapidly 

in width in the early stages, only slightly increasing in 

width in the adult, periphery subacute in the early 

stages, rounded in the adult; chambers fairly distinct, 

increasing rapidly in size and height as added, somewhat inflated in the adult; sutures distinct and slightly 

depressed in the adult portion, slightly oblique; wall 

very finely arenaceous, exterior very smooth; aperture 

semicircular, at the inner margin of the last-formed

•chamber. Length 0.45 to 0.60 mm., breadth 0.25 to 

0.32 mm., thickness 0.13 to 0.15 mm.

The types are from the Paleocene, middle part of 

the Midway group, foot of hill in south slope to Wolf 

€reek, probably in SE %, NE % sec. 20, T. 11 N., K. 12 

E., Butler County, Ala. (3).

This species somewhat resembles Textularia dibollensis Cushman and Applin but differs in the deeper 

sutures and very smooth exterior. It is very common 

tit the type locality but was not found in any of the 

other Paleocene material.

Family VERNEUILINIDAE

Genus GATTDRYIITA D'Orbigny, 1839

Gaudryina rudita Sandidge

Plate 2, figure 7

<Gaudryina rudita Sandidge, Am. Midland Naturalist, vol. 13, p.

342, pi. 31, figs. 19, 20, 1932. 

Cushman, Cushman Lab. Foram. Research Special Pub. 7,

p. 46, pi. 7, figs. 8-10, 1937. 

Cushman and Todd, Cushman Lab. Foram. Research Contr.,

vol. 19, p. 51, pi. 9, fig., 4, 1943. 

Cushman, idem, vol. 20, p. 84,1944. 

Cushman and Deaderick, Jour. Paleontology, vol. 18, p.

329, pi. 50, figs. 9, 10, 1944. 

Cushman, Cushman Lab. Foram. Research Special Pub. 7A,

p. 8, 1946; U. S. Geol. Survey Prof. Paper 206, p. 34, pi. 7,

figs. 23, 24; pi. 8, fig. 1, 1946. 

Textularia agglutlnans W. Berry (not D'Orbigny), in Berry and

Kelley, U. S. Nat. Mus. Proc., vol. 76, art. 19, p. 3, pi. 2,

fig. 1, 1929.

•Gaudryina rugosa Cushman (not D'Orbigny), Tennessee Div. 

Geology Bull. 41, p. 20, pi. 1, figs. 9, 10, 1931.

•Gaudryina minima Cushman (?) (not Egger), Jour. Paleontology, vol. 5, p. 301, pi. 34, figs. 5a, b, 1931.

The previous records for this species are all from the 

Upper Cretaceous. A few specimens from the basal 

part of the Midway group, 9 feet above water level, 

Old Canton Landing, Alabama River, Wilcox County, 

Ala., seem to be this species and are very perfect, showing no sign of having been reworked from older beds, 

although there is evidence of reworking in these beds.

The form recorded from the Paleocene of Mississippi 

by Kline as "Gaudryina rugrosa D'Orbigny" (Mississippi Geol. Survey Bull. 53, p. 14, pi. 7, fig. 11, 1943)

is probably this species.

Kincaid formation. Texas, Hunt County (37).

Gaudryina soldadoensis Cushman ard Renz 

Plate 2, figures 8, 9

Gaudryina soldadoensis Cushman and Ren-*, Cushman Lab.

Foram. Research Contr., vol. 18, p. 4, p1 , 1, fig. 1, 1942. 

Cushman, Cushman Lab. Foram. Reserrch Special Pub.

7A, p. 14, pi. 2, fig. 1; pi. 3, fig. 2, 1946. 

Gaudryina sp. Cushman, Cushman Lab. Foram. Research Contr.

vol. 16, p. 53, pi. 9, fig. 5, 1940. 

Gaudryina sp. Kline, Mississippi Geol. Survey Bull. 53, p. 15,

pi. 7, fig. 14, 1943.

Test somewhat longer than broad, slightly arcuate, 

early portion triangular and triserial, adult portion 

biserial, much compressed, the broader faces slightly 

concave, narrower one deeply so except in the final 

chamber in the adult; chambers distinct except in the 

early portion; sutures in the adult strongly depressed; 

wall finely arenaceous, the exterior slightly roughened; 

aperture a rounded opening near the inner margin of 

the last-formed chamber. Length of h^otype 1.3 mm.; 

breadth 0.65 mm.; thickness 0.35 mm.

The types are from the Paleocene Sol dado Rock formation, Soldado Rock, Trinidad, B. W. I. It also 

occurs in the Chalybeate limestone member of the Clayton formation of Alabama (9) and in the Clay ton formation of Mississippi as recorded by Fline.

Gaudryina sp.

A few specimens occurring in the Paleocene appear 

to be reworked Cretaceous species and without further 

evidence are not included here. Some of the other records such as "Graudryina rugosa D'Orbigny" (Kline, 

Mississippi Geol. Survey Bull. 53, p. 14, pi. 7, fig. 11, 

1943) from the Porters Creek clay may slso be reworked 

Cretaceous material.

Genus CLAVULINOIDES Cushman, 1936 

Clavulinoides midwayensis Cushman

Plate 2, figures 10-16

Clavulinoides midwayensis Cushman, Cushman Lab. Foram. Research Special Pub. 6, p. 21, pi. 3, figs. 9, 15, 1936; Special 

Pub. 7, p. 126, pi. 18, figs. 8, 9,1937; Cushman Lab. Foram. 

Research Contr., vol. 16, p. 53, pi. 9, fig. 6, 1940.

Kline, Mississippi Geol. Survey Bull. 53, p. 13, pi. 1, fig. 2, 

1943.

Applin and Jordan, Jour. Paleontology, vol. 19, p. 131 

(list), 1945.

Cushman and Todd, Cushman Lab. Foram. Research Contr., 

vol. 22, p. 46, pi. 7, figs. 3-6,1946.

Cushman, Cushman Lab. Foram. Research Special Pub.

7A, p. 34, 1946.

Clavulina angularls Plummer (not D'Orbigny), Texas Univ. 

Bull. 2644, p. 70, pi. 3, figs. 4, 5,1927.

Test in the microspheric form very rapidly increasing in diameter toward the apertural end, test in the
VALVULINIDAE 9

megalospheric form with the sides parallel in the adult, 

and in the last-formed chambers with the diameter 

decreasing, triangular throughout, or in the megalospheric form in the later portion rounded, sides concave 

in the megalospheric form in the adult becoming convex; chambers distinct, not inflated except in the last 

chambers of the megalospheric form; sutures distinct, 

very slightly depressed in the earlier portions and in 

the microspheric form, but in the megalospheric form 

becoming deeply depressed in the adult; wall coarsely 

arenaceous, often rather roughly finished; aperture in 

the megalospheric form rounded, in the microspheric 

form with somewhat irregular lobes projecting toward 

the angles of the test. Length up to 1.25 mm.; diameter 0.30-0.60 mm.

The types are from the Paleocene, from a shallow 

ditch at road corner, SE. of New Corsicana Reservoir, 

on road to Mildred, Navarro County, Tex. (401).

Clayton formation, Chalybeate limestone member. Alabama,

Sumter County (9).

Porters Creek clay. Mississippi, Kemper County (17). 

Midway formation, upper part. Arkansas, Pulaski County

(24,25). 

Midway formation. Arkansas, Saline County (26) ; Pulaski

County (27) ; Clark County (30). 

Midway group. Texas, Navarro County (40) ; Limestone

County (42) ; Bastrop County (43) ; Caldwell County (44) ;

Van Zandt County (48). 

Wills Point formation. Texas, Travis County (33) ; Caldwell

County (34). 

Mexia member. Texas. Limestone County (36).

Family VALVULINIDAE 

Genus VALVULAMMINA Cushman, 1933

Valvulammina nassauensis Applin and Jordan var. cubana 

Cushman and Bermudez

Plate 21, figure 16

Valvulammina nassauensis Applin and Jordan var. cubana Cushman and Bermudez, Cushman Lab. Foram. Research 

Contr., vol. 24, p. 68, pi. 11, fig. 1, 1948.

Variety differing from the typical form in the larger 

size, more tapering form and more pointed initial end. 

Length of holotype 1.17 mm.; breadth 0.87 mm.

The types are from the Paleocene Madruga formation, under highway bridge on Central San Antonio, 

Madruga, Habana Province, Cuba.

Genus MARSSONELLA Cushman, 1933 

Marssonella oxycona (Reuss) Cushman

Plate 2, figure 21

Gaudryina oxycona, Keuss, Akad. Wiss. Wien, Math-naturwiss. 

Kl., Sitzungsber., vol. 40, p. 229, pi. 12, fig. 3, 1860; idem, 

vol. 46, pt. 1, p. 33,1862 (1863).

Marssonella oxycona Cushman, Cushman Lab. Foram. Research 

Contr., vol. 9, p. 36, pi. 4, fig. 13,1933.

A few poorly preserved specimens were found in the 

basal part of the Midway group, 9 feet above present 

water level, Old Canton Landing, Alabama River,

Wilcox County, Ala. These have the appearance of 

being reworked from the Cretaceous. The spe°ies is 

also recorded by Kline (Mississippi Geol. Survey Bull. 

53, p. 16, pi. 7, fig. 10,1943) "from the uppermost Clayton chalk from a test hole (M65A) on the Dr. T. D. 

Houston property (NW*4, NE%, NW^ sec. 10, T. 

16 S., R. 3 E.), above a road cut ^ mile west of Prairie 

Creek," Clay County, Miss. The single specincen recorded is also a fragmentary one and may probably 

also be a reworked specimen.

Marssonella cf. M. indentata (Cushman and Jarvis) Cushman 

Rare specimens from the Paleocene of Cuba, under 

highway bridge 011 Central San Antonio, Madruga, 

Habana Province, Cuba, seem very close to this Cretaceous species. It is recorded by Van den Bole1 from 

the Paleocene of Guatemala and British Honduras 

(Thesis Univ. Utrecht, Amsterdam, p. 122, pi. 18, figs. 

2a, b, 1946). Specimens are not entirely typical.

Genus DOROTHIA Flummer, 1931 

Dorothia alabamensis Cushman

Plate 2, figure 17

Dorothia alabamensis Cushman, Cushman Lab. Foram. Research Cont-r., vol. 16, p. 53, pi. 9, fig. 7, 1940.

Applin and Jordan, Jour. Paleontology, vol. 19, p. 131 

(list), 1945.

Cushman, Cushman Lab. Foram. Research Special Pub. 

8A, p. 29, pi. 2, fig. 21, 1947.

Test tapering throughout, broadest at the apertural 

end, usually distinctly twisted, little if at all compressed, earliest whorl with 4 or 5 chambers later 

triserial, and in the adult biserial; chambers distinct, 

somewhat inflated, increasing rather regularly in size 

as added, slightly overlapping; sutures distinct, depressed, more strongly so in the adult portion; wall 

finely arenaceous, smoothly finished but not polished; 

aperture a low, broad opening at the base of the lastformed chamber at the inner margin. Length up to 

1.15 mm.; breadth 0.60 mm.; thickness 0.30 mm.

The types are from the Paleocene, U. S. Highway 80, 

South of Sucarnoochee Creek, y2 mile SW. of Livingston, Sumter County, Ala. (9).

Clayton formation, Chalybeate limestone member. Alabama,

Sumter County (9). 

Midway formation. Arkansas, Clark County (30).

Genus MARTINOTTIELLA Cushman, 1933

Martinottiella paleocenica Cushman

Plate 2, figure 19

Martinottiella, paleocenica, Cushman, Cu,shman Lab. Foram. Research Contr., vol. 23, p. 82, pi. 18, fig. 1, 1947.

Test minute, early portion irregularly triangular in 

section, later portion with a short, irregular biserial 

stage followed by a uniserial stage, rounded in section; 

chambers of the early portion indistinct, those of the 

later biserial and uniserial stages very few in number,
10 PALEOCENE FOKAMINIIFERA OF GULF COASTAL REGION

Jiot increasing in diameter as added, very slightly inflated ; sutures of the early portion indistinct, later ones

•distinct and slightly depressed; wall finely arenaceous, 

smoothly finished; aperture terminal, rounded, with a 

slight neck. Length 0.30-0.40 mm., diameter 0.12-0.15 

mm.

The types are from the Paleocene Naheola formation, 

road cut on east slope to a branch in SW14 sec-12, T. 11, 

N., E. 9 E., Wilcox County, Ala. This small but distinct species seems to be found only in the Naheola 

formation.

Porters Creek formation, Matthews Landing marl member. Alabama, Wilcox County (5, 6).

Genus SCHENCKIELLA Thalmann, 1942

Schenckiella alabamensis Cushman

Plate 2, figure 18

Listerella laevis Cushman (not Finlay), Cushman Lab. Foram. 

Research Contr., vol. 16, p. 54, pi. 9, fig. 8,1940.

ScliencJciella alabamensis Cushman, Cushman Lab. Forani. Research Special Pub. SA, p. 51, pi. 8, fig. 10,1947.

Test elongate, slender, of rather uniform diameter 

throughout, triserial portion rounded; chambers mostly 

rather indistinct, later ones becoming more inflated, 6 or 

7 in the uniserial portion in the adult; sutures indistinct 

except in the last portion, where they are often distinctly 

depressed; wall finely arenaceous, smoothly finished; 

aperture terminal, rounded, small, often with a slight 

lip. Length up to 1.20 mm.; diameter 0.35 mm.

The types are from the Paleocene, U. S. Highway 80, 

S. of Sucarnoochee Creek, y2 mile S.W. of Livingston, 

Sumter County, Ala.

Clayton formation, Chalybeate limestone member. Alabama, 

Sumter County (9). Mississippi, Noxubee County (16).

Genus TRITAXILINA Cushman, 1911

Tritaxilina cubensis Cushman and Bermudez

Plate 2, figure 20

Tritaxilina cubetisis Cushinan and Bermudez, Cushman Lab. 

Foram. Research Contr., vol. 12, pi. 10, figs. 24, 25, 1936; 

vol. 13, p. 7,1937.

Cushman, Cushman Lab. Foram. Research Special Pub. 8, 

p. 156, pi. 18, figs. 4, 5, 1937; Cushman Lab. Foram. Research Contr., vol. 16, p. 54, pi. 9, fig. 9, 1940. »

Test elongate, fusiform, about twice as long as broad, 

greatest breadth at or a little above the middle, apertural end rounded, initial end pointed; chambers fairly 

distinct with deep excavation at the base at either side, 

middle of chambers extending downward; sutures fairly 

distinct, depressed; wall coarsely arenaceous but rather 

smoothly finished; aperture small, narrow, at right 

angles to the inner margin, in a distinct depression. 

Length up to 1.50 mm.; diameter 0.75 mm.

The types are from the Eocene, lower beds of "El 

Husillo" Quarry, Puentes Grandes, Havana, Cuba.

•Clayton formation, Chalybeate limestone member. Alabama, 

Sumter County (9).

Family MILIOLIDAE

Genus ftTIINftTIELOCTILINA D'Ortrigny, 1826

Quinqueloculina naheolensis Cushman

Plate 2, figure 22

Quinqueloculina naheolensis Cushman, Cushman Lab. Foram. 

Research Contr., vol. 23, p. 82, pi. 17, fig. 14, 1947.

Test very small, about twice as long as broad, periphery rounded, base broadly rounded, apertural end 

truncate, sides only slightly convex; chambers slightly 

inflated, increasing very little in breadth as added; sutures fairly distinct, very slightly depressed; wall 

smooth; aperture terminal, circular, without a neck, 

with a distinct tooth, broad at the inner end and occasionally slightly bifid. Length 0.30-0.35 mm., breadth 

0.15-0.18 mm., thickness 0.12-0.15 mm.

The types are from the Paleocene Matthews Landing 

marl member of the Porters Creek formation, Alabama 

Highway 96, 9.2 miles NE. of Kimbrough, Wilcox 

County, Ala. (5). It also occurs at other localities in 

Alabama and seems to be a distinctive fossil for this 

formation. 

Porters Creek formation, Matthews Landing marl member.

Alabama, Wilcox County (1, 2, 5, 6).

Quinqueloculina pulcherrima CusHman 

Plate 2, figure 23

Quinqueloculina pulcherrima Cushman, Cuslman Lab. Foram. 

Research Contr., vol. 20, p. 30, pi. 5, fig. 5, 1944.

Test very small, less than twice as long as broad, periphery rounded or subangular; chambers inflated, basal 

end little if at all projecting, apertural end without a 

neck, obliquely truncate, sutures depressed; wall ornamented with numerous fine, but distinct, longitudinal 

costae, the channels between with slight transverse 

ridges; aperture elliptical, with a sligH thickening of 

the lip, and a simple elongate tooth on tH inner margin. 

Length of holotype 0.28 mm., breadth 0.16 mm.

The types are from the Paleocene Coal Bluff marl 

member of the Naheola formation, cre^k bottom, just 

west of store at Caledonia, about y± mile south of center 

of sec. 29, T. 11 N., K. 10 E., Wilcox County, Ala. (8).

Porters Creek clay. Tennessee, Hardeman County (13).

Porters Creek formation, Matthews Landing marl member. 

Alabama, Butler County (3).

Naheola formation, Coal Bluff marl mernbe". Alabama, Wilcox County (8).

Quinqueloculina alabamensis Cushman 

Plate 2, figure 24

Quinqueloculina alatamensis Cushman, Cusl man Lab. Foram. 

Research Contr., vol. 20, p. 30, pi. 5, fig. 4, 1944.

Test small, about 2,y2 times as long as broad, periphery angled; chambers with the basal end protruding, 

apertural end with a short cylindrical neck, chambers 

triangular in transverse section with the peripheral 

angle with a thickened ridge; sutures slightly if at all
MILIOLIDtAE 11

Repressed; wall smooth; aperture at the end of the 

short neck, with a very slight thickening and a small, 

s :mple tooth on the inner border. Length of holotype 

(,47 mm.; breadth 0.20 mm.

The types are from the Paleocene Coal Bluff marl 

riember of the Naheola formation, creek bottom, just 

vest of store at Caledonia, about y± mile south of center 

(f sec. 29, T. 11 N., R. 10 E., Wilcox County, Ala. (8).

This species somewhat resembles Q. mauricens's 

Howe, from the Cook Mountain formation of Louisiana, but is smaller, more elongate, the keels sharper, 

s> nd the chambers more angled.

loiters Creek formation, Matthews Landing marl member. 

Alabama, Butler County (3).

1'aheola formation, Coal Bluff marl member. Alabama, Wilcox County (8).

Quinqueloculina plummerae Cushman and Todd 

Plate 2, figures 25, 26

<~ uinqueloculina plummerae Cushman and Todd, Cushman Lab.

Foram. Research Contr., vol. 18, p. 25, pi. 5, fig. 2, 1942.

Cushman, idem, vol. 20, p. 30, pi. 5, fig. 3, 1944/ 

<~ uinqueloculina ferussacii Plummer (not D'Orbigny), Texas

Univ. Bull. 2644, p. 161, pi. 12, fig. 10, 1927.

Test small, nearly as broad as long, in end view about 

twice as broad as thick, periphery truncate with 

rounded, carinate edges; chambers distinct, the sides 

fattened or slightly concave toward the periphery; 

sutures distinct but little if at all depressed; wall 

smooth; aperture terminal, without a neck, but with 

i slightly thickened lip and a short blunt tooth. 

Length 0.20 to 0.25 mm.; breadth 0.15 mm.; thickness 

0.08 mm.

The types are from the Paleocene Naheola formation, 

o-reensand bed, upper fossiliferous horizon, Naheola 

Landing, Tombigbee Kiver, Ala. (7).

Porters Creek clay. Tennessee, Hardeman County (13, 14).

Forters Creek formation, Matthews Landing marl member. 

Alabama, Wilcox County (4) ; Choctaw County (7).

1'aheola formation. Coal Bluff marl member. Alabama, Wilcox County (8).

Tills Point formation, Mexia member. Texas, Limestone 

County (36).

• Quinqueloculina plummerae Cushman and Todd var. flectata 

Cushman

Plate 2, figure 27

^tuinqueloculina, plummerae Cushman and Todd var. flectata 

Cushman, Cushman Lab. Foram. Research Contr., vol. 

23, p. 82, pi. 17, fig. 15,1947.

Variety differing from the typical form in the lastformed chambers, which have the periphery in a tort ious line and in some specimens with a depressed area 

at the edge.

The types of this variety are from the Paleocene, road 

<^t on east slope to a branch in SW % sec. 12, T. 11 N., 

T, 9 E., Wilcox County, Ala. (1). ' It occurs com- 

. lonly at the type locality and several others in Wilcox 

bounty, but does not seem to occur with the typical

form of the species. This variety should make a good 

index fossil for the Porters Creek formation.

Porters Creek formation, Matthews Landing marl member. Alabama, Wilcox County (1, 2, 5, 6).

Genus SPIROLOCULINA D'Orbigny, 1826

Spiroloculina alabamensis Cushman

Plate 3, figure 1

Spiroloculina alabamensis Cushman, Cushman Lab. Foram. 

Research Contr., vol. 20, p. 31, pi. 5, fig. 8, 1944.

Test small, 2 to 2,y2 times as long as broad, slightly 

concave in the early portion, periphery flat, sharply 

angled at the margins; chambers few, rapidly increasingly in size as added, of equal width throughout, 

strongly projecting at the base, extending into a prominent neck at the apertural end; sutures distinct, depressed ; wall glistening but roughened with irregular, 

short, longitudinal, incised lines, with occasionally a 

longitudinal costa on the peripheral face; aperture 

quadrangular, with a short, simple tooth on the inner 

margin. Length 0.35 to 0.40 mm., breadth 0.15 to 0.25 

mm., thickness 0.07 to 0.08 mm.

The types are from the Paleocene Coal Bluff marl 

member of the Naheola formation, creek bottom, just 

west of store at Caledonia, about *4 mile south of center 

of sec. 29, T. 11 N., E. 10 E., Wilcox County, Ala, (8).

Porters Creek formation, Matthews Landing marl member.

Alabama, Butler County (3). 

Naheola formation, Coal Bluff marl member. Alabama, Vilcox

County (8).

Genus TBJLOCULINA D'Orbigny, 1826

Triloculina alabamensis Cushman

Plate 3, figure 2

Triloculina alabamensis Cushman, Cushman Lab. Foran. Research Contr., vol. 20, p. 32, pi. 5, fig. 11, 1944.

Triloculina inflata Terquem (not D'Orbigny), Soc. g6ol. France 

Mem., 3rd ser., vol. 2, p. 165, pi. 17 (25), figs. 4-6,1882.

Test small, about twice as long as broad, periphery 

broadly rounded; chambers inflated, the base extending back in a broad curve, apertural end obliquely 

truncate without a neck, broadest near the base^ tapering gradually to the apertural end; sutures distinct, depressed; wall smooth; aperture semicircular, with a 

distinct lip and a simple or slightly bifid tooth on the 

inner margin. Length of holotype 0.33 mm., breadth 

0.18 mm.

The types are from the Paleocene Coal Bluff 

marl member of the Naheola formation, creek bottom, 

just west of store at Caledonia, about % mile south of 

center of sec. 29, T. 11 N., R. 10 E., Wilcox Connty, 

Ala. (8).

This is not the same as D'Orbigny's species from the 

Recent and Late Tertiary of Europe. It is apparently 

identical with the form figured by Terquem from the 

middle Eocene of the Paris Basin.
12 PALEOCENE FORAMINHFERA OF GULF COASTAL REGION

Triloculina natchitochensis Howe 

Plate 3, figure 3

Triloculina, natchitochensis Howe, Louisiana Dept. Cons. Geol.

Bull. 14, p. 38, pL 3, figs. 3-5, 1939.

Cushman, Cushman Lab. Foram. Research Contr., vol. 20, 

p. 32, pi. 5, fig. 7, 1944.

Test small, elongate, only the last two chambers showing 

definitely, and these added at angles of approximately 180 

degrees; wall white, thick, longitudinally coarsely costate; aperture simple, subcircular, at the end of a long neck.—Howe.

Length 0.40 to 0.50 mm., breadth 0.15 to 0.20 mm.

Our specimens are slightly smaller than the type but 

seem to have the same general characters. The types 

are from the Eocene Cook Mountain formation, of Louisiana. The only specimens in our Paleocene collections 

referable to it are from Wilcox County, Ala. (8).

Triloculina deca Cushman 

Plate 3, figures 4, 5

Triloculina natchitochensis Howe var. deca Cushman, Cushman 

Lab. Foram. Research Contr., vol. 20, p. 32, pi. 5, figs. 9, 10, 

1944.

Test small, in front view broadly oval, periphery 

strongly convex, in end view roughly triangular; chambers distinct, broad, strongly curved, the basal end 

broadly curved, apertural end broad and not projecting 

into a definite neck; sutures distinct, slightly depressed; 

wall ornamented with obliquely longitudinal depressions and ridges; aperture nearly circular, with a 

slightly thickened lip and a small, simple tooth at the 

inner side of the opening. Length 0.35 to 0.50 mm., 

breadth 0.20 to 0.25 mm., thickness 0.15 to 0.18 mm.

The types are from the Paleocene Coal Bluff marl 

member of Naheola formation, creek bottom, just west 

of store at Caledonia, about 14 mile south of center of 

sec. 29, T. 11 N., E. 10 E., Wilcox County, Ala. (8).

This species seems to be distinct from the form with 

which it was originally described as a variety.

Porters creek formation, Matthews Landing marl member.

Alabama, Wilcox County (1). 

Naheola formation, Coal Bluff marl member. Alabama, Wilcox

County (8).

"Triloculina laevigata Bornemann" 

Plate 3, figures 6, 7

Triloculina laevigata Plummer, Texas Univ. Bull. 2644, p. 161,

pi. 12, fig. 11, 1927.

Cushman and Todd, Cushman Lab. Foram. Research 

Contr., vol. 18, p. 26, pi. 5, fig. 3,1942.

Under this name Mrs. Plummer figured a single 

specimen found in the Texas material from locality 40. 

I have found specimens questionably referable to this 

form in our material at only one locality, 7.

Family OPHTHALMIDIIDAE

Genus CORWTTSPIRA Schultze, 1854

Cornuspira cf. C. byramensis Crshman

Plate 3, figures 8, 9

Specimens from the Paleocene of Alabama are very 

close to G. byramensis, described frcm the Oligocene 

and recorded from the Wilcox Eocene of Alabama.

Mrs. Plummer recorded and figured a single specimen from the Paleocene of Texas as "Comu&pira carinata (Costa)" (Texas Univ. Bull. 2644, p. 160, pi. 12, 

fig. 9,1927). It has a carinate margin and may be different from the form recorded here as G. cf. G. lyrainensis.

Porters Creek formation, Matthews Landing marl member. 

Alabama, Butler County (3).

Family TROCHAMMIWID^ E

Genus TROCHAMMINA Parker and Tones, 1859

Trochammina cf. T. texana Cushman and Waters

Plate 3, figure 10

Specimens from two localities ar-> comparable to 

T. texana and show no sign of reworking. In the Upper 

Cretaceous the species is restricted to beds of the 

Navarro group.

Midway group. Texas, Navarro County (40) ; Limestone County 

(42).

Family PLACOFSILINIDAE

Genus ADHAEREWTIA Plummer, 1938

Adhaerentia midwayensis Plummer

Plate 3, figures 11-13

Adhaerentia midwayensis Plummer, Am. Midland Naturalist,

vol. 19, p. 242, text figs, la-e, 1938.

Cushman, Cushman Lab. Foram. Research Contr., vol. 16, 

p. 54, pi. 9, figs. 10, 11, 1940.

Test elongate, sides nearly parallel, attached in the 

early stages; chambers fairly distinct in the later portion, early 4 or 5 chambers compactly biserial, later 

loosely biserial and last few chambers in the adult 

uniserial, somewhat labyrinthic; sutures rather indistinct, somewhat depressed in all but the earliest portion ; wall composed of various materials with considerable cement, somewhat roughened to fairly smooth on 

the exterior; aperture in the early stages single and 

rounded to slightly lunar toward the apex of the test on 

the apertural face, later becoming terminal and in the 

last-formed chambers in the adult multiple, consisting 

of two or three irregular openings. Maximum length 

about 5 mm., average thickness about 0.8 mm.

The types are from the Paleocene Clayton formation, 

outcrop on a branch of Dixon Creek, bed of stream, on 

State Highway 96 about 1 mile south of junction with 

Highway 28, sec. 1, T. 13 N., R. 6 E., Wilcox County, 

Ala.
TABLE 1.—Distribution of Pateocene Rhizamminidae, Ammodiscidae, Lituolidae, Textulariidae, Verneuilinidae, Miliolidae, Ophthalmidiidae, Trochamminidae, and Placopsilinidae in the Gulf Coastal region 

[Prepared by J. B. Reeside, Jr. Numbers refer to locality list on pages 1-3. K<= localities reported by V. H. One, P=localities reported by H. J. PlummerandC. a. Lalicker, C=Cuba and Haiti, G=Guatemala and Honduras, S=»Soldado Eock, Trinidad, B. W. I.]

Rhizamminidae Flttthifniphftn eocf-nicwg

Ammodiscidae 

Ammodiscus cf. A. cretaceut

Lituolidae 

HaplophTdfftnoidfs cf H esc&wotd

Aimnobaculites expansun

•naXf.fiffn^f9ui

AmmofflOToinulinQ so Jjituote erects .

Textulariidae

wilcoxensis .

l&evis var. cretosa

mexjaensis .

pafcocfttico

Textttterfa midwaydiui

midwtttuiM var "oajwa

«flW*W»

Verneoiiinidae Gbtt&fta* fiotfte-. #ois(iKfrHstitif _ -

Valvaltaidae Vabmlammtoa ntumuensto var. «uta«a_

ffif M. fa<tentflia

J)«ro*M<ioJ«6flMwnsi« ____ . ..........

^^^^^^^abam^n^ —————

TritaxUms cvbensis

MtUolidse

mSib&tRGIUiS

Spinloculmadtiteniensis

TrSocwMfW ttfafofttncnsis

Oplitbalmidudae

Troohamminidae

PtecopsUinWae

Alabama

1

i

X

X

X

X

2

X

X

3

X

X

X

X

X

X

4

X

5

,

X

X

X

X

6

Ji.

X

X

7

X

X

X

8

X

X

X

X

X

X

X,

9

X

X

X

X

X

X

X

X

10 11 12

X

...„

Term.

13

X

X

X

14

X

X

X

Miss.

15 16

X

X

X

17

X

X

X

K

X

X

X

X

X

Arkansas

18 19 20

-

X

X

X

21 22

X

X

X

f23

u

24

X

25

X

X

X

2A

X

X

X 

X

X

X

27

X

X

X

28

i

29 30

X

X

,

31

X

32

Texas

33

X

34

X

35 36

X

X

X

X

37

X

38 39 40

X

X

X

41 42

X

X

X

X

X

'•X

43

X

X

X

44

X

45 46 47 48

X

49 50 51 52

.

P

X

X

X

X

X

X

T.

o

X

Jt"

G

.*

---•

8

X

X

-^

915259—51 (Face p. 12)

LAGENIDAE 13

Clayton formation, Chalybeate limestone member. Alabama,

Sumter County (9). Mississippi, Noxubee County (16). 

Porters Creek clay. Tennessee, Hardeman County (14).

Family LAGENIDAE

Genus ROBUIUS Montfort, 1808

Robulus midwayensis (Plummer)

Plate 3, figures 14-17

Cristellaria midwayensis Plummer, Texas Univ. Bull. 2644, p.

95, pi. 13, fig. 5, 1927. 

Lenticulina midwayensis Plummer, idem. Bull. 3201, pp. 54,

61, 64 (lists), 1933. 

Lenticulina midwayana Israelsky, Sixth Pacific Sci. Congress,

Proc., p. 573, pi. 2, figs. 7, 8, 1939. 

Robulus midwayensis Cushnian, Cushnian Lab. Foram. Research

Contr., vol. 16, p. 54, pi. 9, fig. 12, 1940. 

Toulmin, Jour. Paleontology, vol. 15, p. 579, pi. 78, fig. 23,

text fig. 2G, 1941. 

Cushnian and Todd, Cushnian Lab. Foram. Research Contr.,

vol. 18, p. 26, pi. 5, figs. 4, 5, 1942. 

Kline, Mississippi Geol. Survey Bull. 53, p. 17, pi. 1, fig. 3,

1943. 

Cushnian, Cushmaa Lab. Forarn. Research Contr., vol. 20,

p. 33, pi. 5, fig. 15, 1944. 

Cooper, Jour. Paleontology, vol. 18, p. 351, pi. 55, figs. 22,

23, 1944. 

Cushrnan and Todd, Cushnian Lab. Foram. Research Contr.,

vol. 22, p. 47, pi. 7, fig. 7,1946.

Test large, circular, very closely coiled, full bodied, though 

somewhat compressed; periphery distinctly angular but not 

flanged in its typical form; chambers 10-12 in adult form, 

smooth, narrow, gently curved, radiate from a conspicuous 

central boss and tapering somewhat toward the peripheral margin; aperture at apex of broad septal face. Diameter up to 

1.5 mm., usually less.—Plummer.

The types are from the Paleoeene, in base of high 

bluff on west side of Colorado Kiver between the TravisBastrop county line and the mouth of Dry Creek. 

(Bastrop Quadrangle), Tex.

This species is widely distributed in the Paleoeene 

and shows a considerable amount of variation in the 

number of chambers and the relative amount of elevation of the sutures and umbones.

Porters Creek clay. Tennessee, Hardernan County (13, 14).

Mississippi, Kemper County (17). 

Porters Creek formation, Matthews Landing marl member.

Alabama, Butler County (3) ; Wilcox County (5, 6) ; Choctaw

County (7).

Naheola formation, Coal Bluff marl member. Alabama, Wilcox County (8).

Clayton formation. Alabama, Barbour County (10). 

Chalybeate limestone member. Alabama, Surnter County (9). 

Pine Barren member, Alabama, Butler County (12). 

Midway formation, lower part. Arkansas, Saline County (18,

19, 19A, 20, 21). 

Midway formation, upper part. Arkansas, Pulaski County (24,

25). 

Midway formation. Arkansas, Saline County (26) ; Pulaski

County (28) ; Cla.rk County (30, 31). 

Midway group. -Texas, Limestone County (39, 42) ; Navarro

County (40) ; Hunt County (41) ; Bastrop County (43, 49) ;

Caldwell County (44) ; Travis County (45, 46).

Willis Point formation. Texas, Travis County (33) ; Caldwell

County (34) ; Williamson County (35). 

Kincaid formation. Texas, Hunt County (37) ; Bastrop Countj

. (38).

Robulus midwayensis (Plummer) var. carinatus (Plummer) 

Cushman

« Plate 3, figures 18-20

Cristellaria midwayensis Plummer, var. carinata Plummer, 

Texas Univ. Bull. 2644, p. 97, text fig. 5, 1927.

Robulus midivayensis (Plummer) var. carinatus (Plummer) 

Cushman, Cushman Lab. Foram. Research Contr., vol. 16, 

p. 55, pi. 9, fig. 13,1940.

Variety differing from the typical form in having a 

well developed peripheral flange.

The types of the variety are from the Paleoeene, basal 

part of Midway group, gully in field west of north - 

south stretch of road about 3 miles in a straight linQ< 

northwest of Campbell or 4 miles by road from that 

town to Neyland, Hunt County, Tex.

There seem to be all gradations between this variet7 

and the typical form in the development of the peripheral flange. It seems to be characteristic of the lower 

part of the Paleoeene of Texas and Alabama.

Midway formation. Arkansas, Clark County (30, 31). 

Midway group. Texas, Hunt County (41) ; Van Zandt County

(47, 48) ; Freestone County (50).

Wills Point formation. Texas, Caldwell County (34). 

Kincaid formation. Texas, Hunt County (37).

Robulus pseudo-mamilligerus (Plummer) Cushman 

Plate 4, figures 1-5

Cristellaria pseudo-mamilligera Plummer, Texas Univ. Bull.

2644, p. 98, pi. 7, fig. 11,1927. 

Robulus pseudo-mamilligerus Cushman, Cushman Lab. Foram.

Research Contr., vol. 16, p. 55, pi. 9, fig. 16, 1940. 

Kline, Mississippi Geol. Survey Bull. 53, p. 19, pi. 1, fig. 12,

1943.

Cushnian and Todd, Cushman Lab. Foram. Research Contr., 

vol. 22, p. 47, pi. 7, fig. 10,1946.

Test very slightly elongate, strongly compressed; periphery 

bounded by a rather thick keel that shows on some specimens a 

slight lobation; chambers 9-11 in final convolution, distinctly 

curved; sutures marked by conspicuous tapering elevatiors 

curving outward from an irregularly developed central boss or 

from a group of protuberances; aperture radiate, protruding. 

Diameter up to 1.6 mm.—Plummer.

The types are from the Paleoeene, exposure along a 

small branch about % mile NW. of Tehuacana and 0.2 

mile N. of the Tehuaeana-Waeo road on the first road 

turning north, Limestone County, Tex.

The species is most common in the glauconitic middle 

portion of the Midway group of Texas and is recorded 

from the Paleoeene of Arkansas and Alabama and from 

the uppermost part of the Porters Creek clay of 

Mississippi.
14 PALEOCENE FORAMINIFERA OF GULF COASTAL REGION

Clayton formation, Chalybeate limestone member. Alabama,

Sumter County (9).

Midway formation, upper part. Arkansas, Pulaski County (25). 

Midway formation. Arkansas, Clark County (31). 

Midway group. Texas, Limestone County (39) ; Caldwell

County (44). 

Wills Point formation. Texas, Travis County (33) ; Caldwell

County (34).

• 

Robulus turbinatus (Plummer) Cushman

Plate 4, figures 6-9

Cristcttaria turbinata Plummer, Texas Univ. Bull. 2644, p. 93,

pi. 7, fig. 4, 1927. 

lobultis turbinatus Cushman, Cushman Lab. Forani. Research

Contr., vol. 16, p. 55, pi. 9, fig. 17, 1940. 

Kline, Mississippi Geol. Survey Bull. 53, p. 20, pi. 1, fig.

7, 1943. 

Cushman and Todd, Cushman Lab. Foram. Research Contr.,

vol. 22, p. 47, pi. 7, fig. 11, 1946.

Test circular, considerably compressed; peripheral margin 

sharp and extended into a fragile, white flange that is typically 

ragged; chambers 8 in final convolution, narrow, smooth ; sutures 

strongly elevated and of about equal width from the large 

unibonal area to the periphery, very strongly curved; aperture 

at apex of narrow septal face. Diameter up to 0.6 mm.— 

Plummer.

The types are from the Paleocene, clay pit, Mexia 

Brick Works, about 1 mile west of the town of Mexia, 

Limestone County, Tex.

The species occurs in the Paleocene of Texas, Arkansas, Alabama, and Mississippi.

Porters Creek formation, Matthews Landing marl member.

Alabama, Butler County (3). 

Clayton formation, Chalybeate limestone member. Alabama,

Sumter County (9). Mississippi, Noxubee County (16). 

Midway formation, upper part. Arkansas, Pulaski County (24,

25). Midway formation. Arkansas, Saline County (26) ;

Pulaski County (27) ; Clark County (30, 31). 

Midway group. Texas, Navarro County (40) ; Limestone

County (42) ; Caldwell County (44). 

Wills Point formation. Texas, Travis County (33) ; Caldwell

County (34). 

Mexia member. Texas, Limestone County (36).

Robulus degolyeri (Plummer) Bandy 

Plate 3, figures 21, 22

Cristellaria degolyeri Plummer, Texas Univ. Bull. 2644, p. 97,

pi. 7, fig. 7,1927. 

Robulus degolyeri Bandy, Jour. Paleontology, vol. IS, p. 368, pi.

60, fig. 5, 1944. 

Cushman and Todd, Cushman Lab. Foram. Research Contr.,

vol. 22, p. 47, pi. 7, fig. 13, 1946. 

Lenticulvtia degolyeri Kline, Mississippi Geol. Survey Bull. 53,

p. 21, pi. 1, fig. 8, 1943.

Test somewhat longer than broad, moderately compressed; 

peripheral margin very sharp and bounded by a ragged flange; 

chambers 7 to 9, gently curved, smooth; sutures marked by 

strong elevations of clear shell matter tapering outward from 

a conspicuous umbonal boss; aperture at apex of an elongate 

septal face. Length up to 0.8 mm.; usually less.—Plummer.

The types are from the Paleocene, clay pit, Mexia 

Brick Works, about 1 mile west of the town of Mexia, 

Limestone County, Tex.

The species has been recorded from the Paleocene of 

Texas, Mississippi, and Arkansas, and from the middle 

Eocene of Oregon.

Midway formation, upper part. Arkansas, Pulaski Connty (25). 

Midway formation. Arkansas, Pulaski County (27) ; Clark

County (29, 30). 

Midway group, Texas, Caldwell County (44) ; Van Zandt

County (47, 48).

Robulus pseudo-eostatus (Plummer) Cushmar 

Plate 4, figures 10-12

Gristellaria pseudo-costata Plummer, Texas Univ. Bull. 2644, p.

98, pi. 7, fig. 9, 1927.

Robulus pseudo-eostatus Cushman, Cushman Lab. Fo^am. Research Contr., vol. 16, p. 55, pi. 9, fig. 18, 1940. 

Kline, Mississippi Geol. Survey Bull. 53, p. 18, p\ 1, fig. 5, 

1943.

Test much compressed, not completely involute, changing from 

the loosely coiled form to linear development in specimens developed beyond maturity; periphery bound by a distinct, thin, 

transparent flange; chambers 7-8 in last whorl, ornamented by 

two to four irregularly developed thin costae that follow roughly 

the direction of coiling; sutures marked by thin, high, uneven 

ridges; aperture protruding from a strongly inflated septal face. 

Diameter up to .7 mm.—Plummer.

The types are from the Paleocene, gully in fi eld west 

of north-south stretch of road about 3 miles in a straight 

line northwest of Campbell or 4 miles by road f ~om that 

town on highway to Neyland, Hunt County, Tex.

This highly ornate species is fairly common at a number of localities in the Texas area but very r-f.re elsewhere.

Clayton formation, Chalybeate limestone member. Mississippi,

Noxubee County (16).

Porters Creek clay. Mississippi, Kemper County (17). 

Midway formation. Arkansas, Clark County (29, 31). 

Midway group, Texas, Hunt County. (41) ; Travis County (45,

46) ; Van Zandt County (47) ; Freestone County (50). 

Kincaid formation. Texas, Hunt County (37).

Robulus pseudo-eostatus (Plummer) Cushman var. comis 

Cushman, new name

Plate 4, figures 13, 14

Robulus pseudo-eostatus (Plummer) Cushman var. inornat'iis 

Kline (not D'Orbigny), Mississippi Geol. Survey Bull. 53, 

p. IS, pi. 1, fig. 6,1943.

Test much compressed, almost completely involute; periphery 

with a distinct, thin, transparent flange which is usually broken ; 

chambers about 7 in final whorl, smooth; sutures marked by thin, 

high ridges which vary from smooth to crenulated; septal face 

less strongly inflated than in species; aperture less distinctly 

protuberant. Average diameter about 0.5 mm.—Kline.

The types are from the Ostrea pulaskensis bed of the 

Paleocene Clayton formation, test hole M 93, 3.8 to 11.0 

feet below surface, iy2 miles northwest of Montpelier, 

Clay County, Miss.

The variety is recorded as characteristic of tiH Clayton formation but also occurs in the Porters Creek clay 

of Mississippi.

Clayton formation, Chalybeate limestone member. Mississippi, 

Noxubee County (16).
LAGENIDAE 15

Robulus alabamensis Cushman 

Plate 4, figures 15, 16

Robulus alabamensis Cushman, Cushman Lab. Foram. Research

Contr., vol. 20, p. 33, pi. 5, fig. 13,1944.

Cushman and Todd, idem, vol. 22, p. 47, pi. 7, fig. 8, 1946. 

Robulus sp. Cushman, idem, vol. 16, p. 55, pi. 9, fig. 23, 1940.

Test small for the genus, umbonate, close coiled, 

periphery within the area of each adult chamber slightly flattened, sharply angled but not carinate; chambers 

numerous, 8 to 10 in the final coil, distinct, of uniform 

shape, increasing gradually and regularly in size as 

added, the apertural angle translucent; sutures distinct, 

nearly tangential, very slightly curved, not depressed; 

wall smooth, polished; aperture radiate, with a slight 

opening at the upper end of the ventral face. Diameter 

of holotype, 0.70 mm., thickness 0.40 mm.

The types are from the Paleocene Coal Bluff marl 

member of the Naheola formation, creek bottom, just 

west of store at Caledonia, about 14 mile south of center 

of sec. 29, T. 11 N., R. 10 E., Wilcox County, Ala. (8).

This species seems related to R. pondi Cushman, of 

the Upper Cretaceous, but differs in the fewer chambers, the non-nodose periphery, and the more umbonate 

center.

Naheola formation, Coal Bluff marl member. Alabama, Wilcox 

County (8).

Clayton formation, Chalybeate limestone member. Alabama, 

Sumter County (9).

Midway formation, upper part. Arkansas, Pulaski County (25).

Midway formation. Arkansas, Clark County (29-31).

Midway group. Texas, Hunt County (41) ; Caldwell County 

(44) ; Travis County (46) ; Van Zandt County (47) ; Freestone County (50).

Robulus wilcoxensis Cushman and Ponton 

Plate 4, figure 17

Robulus wilcoxensis Cushman and Ponton, Cushman Lab.

Foram. Research Contr., vol. 8, p. 52, pi. 7, fig. 3, 1932. 

Toulmin, Jour. Paleontology, vol. 15, p. 579, pi. 78, figs. 24,

25; text fig. 2H, 1941. 

Cushman and Todd, Cushman Lab. Foram. Research Contr.,

vol. 18, p. 27, pi. 5, fig. 7,1942. 

Kline, Mississippi Geol. Survey Bull. 53, p. 20, pi. 7, fig. 27,

1943.

Test compressed, close-coiled except in the oldest portion where one or two chambers may become uncoiled, 

periphery in the earlier portion with a narrow blunt 

keel that becomes obsolescent and rounded in the last 

chambers; chambers numerous, 9 or 10 in the lastformed coil of the adult, later ones slightly inflated and 

uncoiling, early ones of uniform shape, gradually increasing in size as added; sutures distinct, rather 

strongly curved, in the early portion limbate and raised, 

then becoming flush with the surface and in the adult 

slightly depressed; wall smooth except for the early 

raised sutures; aperture radiate, terminal in the adult, 

in the earlier chambers at the outer peripheral angle.

Length 1.00-1.15 mm., breadth 0.75-0.80 mm., thickness 

0.25-0.30 mm.

The types are from Eocene of Wilcox age from railroad cut, 1 mile north of Ozark, Ala.

Porters Creek formation, Matthews Landing marl member.

Alabama, Choctaw County (7). 

Midway group. Texas, Limestone County (39).

Robulus wilcoxensis Cushman and Ponton var. dissentius 

Cushman and Todd

Plate 4, figure 18

Robulus wilcoxensis Cushman and Ponton var. dissentia Cushman and Todd, Cushman Lab. Foram. Research Contr., 

vol. 22, p. 48, pi. 7, fig. 12,1946.

Variety differing from the typical form in the deeply 

incised sutures, more umbonate center, which is frequently ornamented with small bead-like elevations, and 

the more inflated chambers.

The types of the variety are from the Paleocene, 

about 1,000 feet south of Roosevelt Road viaduct, 200 

feet east of railroad tracks, in small gully heading 

east, just upstream from small abandoned bridge, 

NE14NW1/4 sec. 16, T. 1 K, R. 12 W., Little Rock, Ark. 

(25).

This seems to be a rare variety, common at the type 

locality, but rare elsewhere.

Porters Creek formation, Matthews Landing marl member.

Alabama, Butler County (3). 

Midway formation, upper part. Arkansas, Pulaski County (25).

Robulus arkansasanus Cushman and Todd 

Plate 4, figure 19

Robulus arkansasana Cushman and Todd, Cushman Lab. Foram. 

Research Contr., vol. 22, p. 48, pi. 7, fig. 14, 1946.

Test close coiled, strongly umbonate; periphery acute 

but not keeled, very slightly lobulate; chambers distinct, very slightly inflated, seven or eight in the adult 

coil, increasing very gradually and uniformly in size 

as added; sutures distinct, strongly limbate, curved, 

raised, broadest near the umbo where they tend to fuse; 

wall smooth except for the raised sutures; aperture at 

the peripheral angle of the last-formed chamber, 

slightly projecting, with a distinct median slit extending slightly down the apertural face. Diameter 1.25 

to 1.50 mm., thickness 0.65 to 0.75 mm.

The types are from the Paleocene, about 1,000 feet 

south of Roosevelt Road viaduct, 200 feet east of railroad tracks, in small gully heading east, just upstream 

from small abandoned bridge, NE^NW^ sec. 16, T. 

1 N., R. 12 W., Little Rock, Ark. (25).

Midway formation, upper part. Arkansas, Pulaski County (25). 

Midway formation. Arkansas, Pulaski County (27) ; Clark

County (30) ; Navarro County (40) ; Caldwell County (44) ;

Van Zandt County (47). 

Kincaid formation. Texas, Hunt County (37).
16 PALEOC'ENE FORAMINIFERA OF GULF COASTAL REGION

Robulus cf. R. rosettus (Giimbel) Cushman 

Plate 4, figures 20, 21

Robulina rosetta Giimbel, K. bayer. Akad. Wiss., Math.-phys.

Abt., Abh., Kl. 2, vol. 10, p. 642, pi. 1, fig. 73, 1870. 

Robulus rosetta Cushman, Cushman Lab. For am. Research

Contr., vol. 16, p. 55, pi. 9, fig. 24,1940. 

Cushman and Renz, idem, vol. 18, p. 5, pi. 1, fig. 4, 1942. 

Kline, Mississippi Geol. Survey Bull. 53, p. 19, pi. 1, fig. 9, 

1943.

Specimens compared with R. rosettus, described by 

Giimbel from the lower Eocene of Bavaria, are from 

the Paleocene of Alabama, Trinidad, Mississippi, and 

Texas. They have few, five or six, chambers in the adult 

whorl, distinctly inflated, periphery slightly lobulate 

with a distinct keel; sutures depressed and strongly 

curved; aperture in the adult with a slight neck. 

Clayton formation, Chalybeate limestone member. Mississippi,

Noxubee County (16).

Midway formation. Arkansas, Clark County (31). 

Wills Point formation. Texas, Williamson County (35).

Robulus insulsus Cushman

Plate 5, figures 1-3

Robulus insulsus Cushman, Cushman Lab. Foram. Research 

Contr., vol. 23, p. 83, pi. 18, figs. 2, 3, 1947.

Cristellaria orbicularis Plummer (not D'Orbigny), Texas Univ. 

Bull. 2644, p. 92, pi. 7, fig. 1, 1927.

Test small, close coiled, very thick; periphery acute 

and slightly keeled; chambers six or seven in number, 

not inflated, increasing very gradually in size as added; 

sutures distinct, very strongly curved, sometimes slightly raised, usually flush with the surface; wall smooth, 

polished, somewhat transparent; aperture radiate, 

slightly projecting. Diameter 0.30 to 0.40 mm., thickness 0.20 to 0.25 mm.

The types are from the Paleocene, 4.6 miles north of 

Feiitress, Caldwell County, Tex. (44).

This species was first figured from the Paleocene by 

Mrs. Plummer as cited above. It occurs in Texas and 

Alabama in the Paleocene and is very constant in its 

size and general characters. It differs from R. orbicularis (D'Orbigny) in the much smaller size and fewer 

chambers.

Porters Creek formation, Matthews Landing marl member.

Alabama, Butler County (3). 

Midway group. Texas, Caldwell County (44).

Robulus piluliferus Cushman 

Plate 5, figure 4

Robulus piluliferus Cushrnan, Cushman Lab. Foram. Research 

Contr., vol. 23, p. 83, pi. 18, fig. 4,1947.

Test of medium size, close-coiled, apertural face 

slightly concave with raised borders, periphery acute, 

very slightly keeled; chambers ten to twelve in the adult 

coil, not inflated, of uniform shape and increasing very 

evenly and gradually in size as added; sutures distinct,

curved, raised, the earlier ones often ending in a rounded 

boss near the umbo and broken into a series of bead-like 

projections toward the inner portion; wall smooth except for the raised sutures; aperture radiate, at the 

peripheral margin of the last-formed chamber. I}iameter of holotype 1.50 mm., thickness 0.65 mm.

The types are from the Paleocene, upper bed at Dry 

Brushy Creek, 6 miles south of Thrall, TVilliamson 

County, Tex. (35).

The species differs from R. midwayensis (Plummer) 

in the generally smaller size and the beaded natures.

Midway formation. Arkansas, Clark County (30).

Midway Group. Texas, Caldwell County (44).

Wills Point formation. Texas, Williamson County (35).

Robulus sp. 

Plate 5, figures 9, 10

Robulus sp. Cushman and Todd, Cushman Lab. Foram. Research Contr., vol. 22, p. 48, pi. 7, figs. 15,16,1916.

Rare specimens of a peculiar form with numerous 

chambers, tending to become slightly uncoiled in the 

adult, are inadequate for specific description.

Midway formation, upper part. Arkansas, Pulaski County (25).

Genus FLANULARIA Defrance, 1824

Planularia toddae Cushman

Plate 5, figures 5-8

Planularia toddae Cushman, Cushman Lab. Foram. Research 

Contr., vol. 20, p. 34, pi. 5, fig. 12,1944.

Astacolus jugleri Toulmin (not Reuss), Jour. Paleontology, vol. 

15, p. 580, pi. 78, figs. 27, 28,1941.

Planularia sp. Cushman and Todd, Cushman Lab. Foram. Research Contr., vol. 18, p. 27, pi. 5, fig. 8,1942.

Test small, compressed, very slightly umbonate, 

periphery subacute; chambers distinct, later ones tending to become more elongate, increasing very gradually 

in size as added; sutures distinct, very slightly curved, 

somewhat thickened, not depressed; wall smooth, polished; aperture radiate, at the peripheral angle. 

Length of holotype 0.45 mm., breadth 0.27 mm., thickness 0.15 mm.

The types are from the Paleocene Coal Pluff marl 

member of the Naheola formation, creek bottom, just 

west of store at Caledonia, about y4= mile south of center 

of sec. 29, T. 11 N., R 10 E., Wilcox County, Ala. (8).

This species differs from "Cristellaria jugleri Reuss" 

in its much smaller size, much less elongate form, slight 

increase in length of the chambers, and less curved ventral face.

Porters Creek clay. Tennessee, Hardeman County (13,14).

Porters Creek formation, Matthews Landing marl member. Alabama, Wilcox County (1, 2, 5).

Naheola formation, Coal Bluff marl member. Alabrma, Wilcox 

County (8).

Clayton formation, Pine Barren member. Alabama, Wilcox 

County (11) ; Butler County (12).
LAGENIDAE 17

Genus MARGINUIINA D'Orbigny, 1826

Marginulina tuberculata (Plummer) Cushman and Bermudez

Plate 5, figures 11-13

Cristellaria subaculeata Cushman var. tuberculata Plummer,

Texas Univ. Bull. 2644, p. 101, pi. 7, fig. 2; pi. 14, fig. 1,1927. 

Hemicristellaria subaculeata (Gushman) var. tuberculata

Kline, Mississippi Geol. Survey Bull. 53, p. 22, pi. 1, fig. 11,

1943. 

Margimilina tuberculata Cushman and Bermudez, Cushman

Lab. Foram. Research Contr., vol. 24, p. 69, pi. 11, fig. 5,

1948.

Test elongate, somewhat compressed; periphery rounded on 

early chambers and very bluntly angular on later chambers of 

mature forms; chambers numerous, smooth, first six or seven 

piano-spiral followed by a linear succession of short, compact 

chambers; sutures marked by rows of distinct beadlike tubercles best developed on the coiled portion of the test and giving 

place to more ridgelike elevations between later chambers 

or even to depressions in extreme maturity; aperture protruding, radiate, peripheral. Length up to 1.4 mm.; average 1 mm.— 

Plummer.

The types are from the Paleocene Mexia member of 

the Wills Point formation, clay pit of Mexia Brick 

Works, about 1 mile west of the town of Mexia, Limestone County, Tex.

The species is largely confined to the upper part of 

the Midway group of Texas, but rare specimens were 

found in the Naheola formation of Alabama, and it is 

also recorded from the Porters Creek clay of Mississippi and the Madruga formation of Cuba.

Porters Creek formation, Matthews Landing marl member.

Alabama, Wilcox County (5). 

Wills Point formation. Texas, Travis County (33) ; Caldwell

County (34).

Mexia member. Texas, Limestone County (36). 

Midway group. Texas, Limestone County (39, 42) ; Caldwell

County (44).

Marginulina earlandi (Plummer) Cushman 

Plate 5, figures 14-16

Cristellaria earlandi Plummer, Texas Univ. Bull. 2644, p. 103,

pi. 7, fig. 10, 1927. 

Marginulina earlandi Cushman, Cushman Lab. Foram. Research

Contr., vol. 16, p. 56, pi. 9, figs. 14, 15, 1940. 

Kline, Mississippi Geol. Survey Bull. 53, p. 22, pi. 1, fig. 10,

1943.

Test very elongate, much compressed; peripheral margin narrowly rounded; chambers numerous, first six or seven closely 

coiled about a conspicuous and only slightly protruding boss, 

later ones in an erect series; sutures oblique in both the coiled 

and uncoiled portion of test, strongly elevated in rather even 

development in the coiled area but more greatly thickened on 

each side of the linear series; aperture marginal. Length probably up to about 3 mm.—Plummer.

The types are from the Paleocene, clay pit, Mexia 

Brick Works, about 1 mile west of the town of Mexia, 

Limestone County, Tex.

The species is not common in our material but rare 

specimens occur in the Paleocene of Alabama and it is 

also recorded from Mississippi.

Clay ton formation. Alabama, Barbour County (10). Chalybeate limestone member. Alabama, Sumter County (9). 

Midway group. Texas, Limestone County (39, 42).

Marginulina toulmini Cushman

Plate 5, figure 17 

Marginulina toulmini Cushman, Cushman Lab. Foram. Research

Contr., vol. 20, p. 34, pi. 5, fig. 16, 1944.

Vaginulinopsis brantlyi Toulmin (not Hemicristellaria T>rantlyi 

Garrett), Jour. Paleontology, vol. 15, p. 583, pi. 79, figs. 

8-10,1941.

Marffinulina sp. Cushman, Cushman Lab. Foram. Research 

Contr., vol. 20, p. 21, pi. 4, fig. 4,1944.

Test elongate, compressed, the early portion close 

coiled, later adult portion uncoiled; periphery in the 

adult becoming nearly straight; chambers distinct, 6 

to 8 in the coiled portion, rapidly increasing in size, 

the adult uncoiled portion consisting of 3 or 4 chambers increasing gradually in height but not in width; 

sutures distinct, slightly curved, raised, with a row of 

small tubercles of clear shell material increasing in 

number in the uncoiled portion; wall mostly sniooth 

except for the sutural beading, the last one or two 

chambers in the adult occasionally with a few, small, 

slightly elongate tubercles; aperture radiate, at the 

outer peripheral angle, in the adult with a slight neck. 

Length of holotype 1.00 mm., breadth 0.37 mm., thickness 0.25 mm.

The types are from the Paleocene Coal Bluff marl 

member of the Naheola formation, creek bottom, just 

west of store at Caledonia, about % mile south of center 

of sec. 29, T. 11 N., R. 10 E., Wilcox County, Ala. (8). 

It was abundant at the type locality but not found elsewhere in our material.

Marginulina eximia Neugeboren

Plate 5, figures 18,19 

Marginulina eximia Neugeboren, Ver. Mitth. Siebenburg. Ver.

Nat., Jahrg. 2, p. 129, pi. 4, fig. 17,1851. 

Cushman and Ponton, Cushman Lab. Foram. Research

Contr., vol. 8, p. 54, pi. 7, fig. 8, 1932. 

Cushman, idem, vol. 15, p. 56, pi. 9, figs. 27, 28, 1939. 

Beck, Jour. Paleontology, vol. 17, p. 597, pi. 104, figs. 15, 16,

1943. 

Kelley, Am. Assoc. Petroleum Geologists Bull., vol. 27, p. 8

(list), 1943. 

Martin, Stanford Univ. Publ., Univ. Ser., Geol. Sci., vol.

3, no. 3, p. 11 (list), 1943. 

Cushman, Cushman Lab. Foram. Research Contr., vol. 20,

p. 35, pi. 5, fig. 14, 1944. 

Cushman and Todd, idem, vol. 22, p. 48, pi. 7, fig. 17, 1946.

The types of this species are from the Miocene of 

Lapugy, Hungary. Specimens that appear to belong 

to that species occur in the American Paleocene and 

younger portions of the Eocene. They show a considerable amount of variation, as do most species of the 

genus. 

Naheola formation, Coal Bluff marl member. Alabama. Wilcox

County (8). 

Midway formation, upper part. Arkansas, Pulaski County (24,

25). 

Midway group. Texas, Bastrop County (43).
18 PALEOOENE FORAMINHFERA OF GULF COASTAL REGION

Marginulina cf. M. scitula Bornemann 

Plate 5, figures 20, 21

Rare specimens from a few localities have been compared with Bornemann's species, but not enough specimens have been available to make the identification 

certain. Two of these are figured for reference.

Porters Creek formation, Matthews Landing marl member.

Alabama, Choctaw County (7). 

Clayton formation, Chalybeate limestone member. Alabama,

Sumter County (9).

Midway formation. Arkansas, Clark County (30) 

Midway group. Texas, Caldwell County (44).

Marginulina cf. M. subrecta Franke 

Plate 5, figure 22

Very rare specimens from the Paleocene of Arkansas 

and Texas may be compared with M. subrecta, described 

from the Paleocene of Germany.

Midway formation, upper part. Arkansas, Pulaski County

(25). 

Midway group. Texas, Caldwell County (44).

Marginulina cf. M. dubia Neugeboren 

Plate 5, figure 23

Rare specimens from a very few localities in Arkansas, Texas, and Alabama somewhat resemble this 

species, described from the Miocene of Lapugy, Hungary, but are probably not the same. Specimens are 

too few to warrant a specific determination. 

Clayton formation, Pine Barren member. Alabama, Butler

County (12).

Midway formation, upper part. Arkansas, Pulaski County

(25).

Midway group. Texas, Caldwell County (44). 

Wills Point formation, Mexia member. Texas, Limestone

County (36).

Marginulina cf. M. havanensis Cushman and Bermudez 

Plate 5, figure 24

Rare specimens from the Paleocene of Tennessee resemble this species described from the Eocene of Cuba. 

They also are somewhat like specimens from the American upper Eocene referred to M. tennis Bornemann. 

Porters Creek clay. Tennessee, Hardeman County (13, 14).

Marginulina cf. M. glabra D'Orbigny 

Plate 5, figures 25-27

Specimens from the upper part of the Paleocene, 

Midway group of Texas resemble M. glabra but show 

a considerable amount of variation. The species has 

been recorded from the Paleocene of Texas by Plummer 

(Texas Univ. Bull. 2644, p. 104, pi. 6, figs. 3a-g, 192T) 

and as "Vaginulina gldbra, (D'Orbigny)" from the 

Paleocene Porters Creek clay of Mississippi by Kline 

(Mississippi Geol. Survey Bull. 53, p. 32, pi. 3, fig. 4, 

1943).

Wills Point formation. Texas, Caldwell County (34). 

Midway group. Texas, Limestone County (39) ; Bastrop 

County (43) ; Caldwell County (44).

Marginulina cf. M. hamata (Franke) Cushman and Todd 

Plate 5, figures 28, 29

Marginulina cf. M. hamata Cushman and Todd, C ishman Lab. 

Foram. Research Contr., vol. 22, p. 49, pi. 7, figs. 19, 20,. 

1946.

A few specimens in the Paleocene material from 

Arkansas are similar in form to this species, described 

as "Cristellaria hamata" by Franke from the Paleocene 

of Denmark. Our specimens are much smaller than 

the European ones. 

Midway formation, upper part. Arkansas, Pulaski County

(25).

Marginulina distincta Cushman and Bermvdez 

Plate 21, figure 6

Marginulina distincta Cushman and Bermudez, Cushman Lab. 

Foram. Research Contr., vol. 24, p. 69, pi. 11, fig. 2, 1948.

Test elongate, early portion close coiled, becoming 

uncoiled in the adult portion, axis slightly curved, early 

portion somewhat compressed, adult portion circular 

in transverse section; chambers distinct, early ones not 

inflated, adult ones distinctly inflated, increasing in size 

rapidly as added, the last three making up r, large proportion of the test; sutures distinct, earlier ones slightly 

raised, later ones depressed; wall ornamented with numerous fine longitudinal costae, those of the later portion independent of the sutures; aperture radiate, at the 

peripheral angle, somewhat projecting. Length of 

holotype 0.80 mm.; diameter 0.30 mm.

The types are from the Paleocene Madruga formation, highway bridge on Central San Antonio, Madruga, Habana Province, Cuba.

This species differs from M. havanensis Cushman and 

Bermudez in the shorter and stouter form, more 

strongly oblique sutures, and the costate ornamentation.

Marginulina sp. A 

Plate 5, figures 30, 31

The figured specimen is similar to that figured by 

Mrs. Plummer from the Paleocene of Texas as ^Marginulina regularis D'Orbigny" (Texas Univ. Bull. 2644, 

p. 107, pi. 5, fig. 7, 1927). It is not the same as 

D'Orbigny's Miocene species but not enough specimens 

are available to warrant a specific identification.

Midway group. Texas, Caldwell County (44).

Marginulina sp. B 

Plate 5, figures 32, 33

Specimens from the Paleocene of Arkansas are too 

rare to give the full characters for specific determination but are figured for future reference.

Midway formation, upper part. Arkansas, Pulaski County (25). 

Midway formation. Arkansas, Saline County (26).
LAGENIDAE 19

Genus DENTAIINA D'Orbigny, 1826

Dentalina pseudo-obliquestriata (Plummer) Cushman

Plate 6, figure 1-3

Nodosaria pseudo-obliquestriata Plummer, Texas Univ. Bull.

2644, p. 87, pi. 4, fig. 18,1927.

Dentalina pseudo-obliquestriata, Cushrnau, Cushman Lab.

Foram. Research Contr., vol. 16, p. 58, pi. 10, fig. 18, 1940.

Cushman and Todd, idem, vol. 18, p. 28, pi. 5, fig. 10, 1942.

Martin, Stanford Univ. Publ., Univ. Ser., Geol. Sci., vol. 3,

no. 3, p. 10 (list), 1943. 

Kline, Mississippi Geol. Survey Bull. 53, p. 26, pi. 2, fig. 14,

1943. 

Cushman and Todd, Cushman Lab. Foram. Research Contr.,

vol. 22, p. 50, pi. 8, fig. 18,1946.

Test long, slender, arcuate, tapering toward the aboral extremity; chambers numerous, strongly inflated, ornamented by 

coarse costae that follow the length of the test somewhat 

obliquely; sutures strongly constricted; aperture protruding, 

round, somewhat eccentric. Length up to 2 mm.—Plummer.

The types are from the Paleocene, gully in field west 

of north-south stretch of road about 3 miles in a straight 

line northwest of Campbell or 4 miles by road from 

that town on highway to Neyland, Hunt County, Tex.

The species occurs in Texas, Arkansas, and Alabama 

and the characters seem to be fairly constant for species 

in this genus. It seems to be most-common in the lower 

part of the Paleocene.

Clayton formation, Chalybeate limestone member. Alabama, 

Sumter County (9).

Pine Barren member. Alabama, Butler County (12).

Porters Creek clay. Tennessee, Hardeman County (13).

Porters Creek formation, Matthews Landing marl member. Alabama, Choctaw County (7).

Midway formation, lower part. Arkansas, Saline County (21).

Midway formation, upper part. Arkansas, Pujaski County (25).

Midway formation. Arkansas, Clark County (29)

Midway group. Texas, Limestone County (39) ; Hunt County 

(41) ; Bastrop County (43) ; Caldwell County (44) ; Travis 

County (45) ; Van Zandt County (47).

Kincaid formation. Texas, Bastrop County (38).

Dentalina gardnerae (Plummer) Cushman 

Plate 6, figures 4-7

Marginulina gardnerae Plummer, Texas Univ. Bull. 2644, p. 106,

pi. 5, fig. 11,1927.

Dentalina (?) gardnerae Cushman, Cushman Lab. Foram. Research Contr., vol. 16, p. 57, pi. 10, figs. 10-12, 1940. 

Toulrnin, Jour. Paleontology, vol. 15, p. 585, pi. 79, fig. 15,

1941. 

Cushman and Todd, Cushman Lab. Foram. Research Contr.,

vol. 18, p. 29, pi. 5, figs. 11,12, 1942. 

Kline, Mississippi Geol. Survey Bull. 53, p. 26, pi. 2, fig. 14,

1943. 

Cushman and Todd, Cushman Lab. Foram. Research Contr.,

vol. 22, p. 50, pi. 8, fig. 18, 1946. 

Cushman and Bermudez, idem, vol. 24, p. 69, pi. 11, fig. 3,

1948.

Test elongate, straight to slightly arcuate, somewhat stout, 

tapering bluntly toward the initial extremity; early chambers 

very slightly compressed, later ones round in transverse section, 

compact, subcylindrical, narrow; sutures evident as dark bands 

or faint lines, constricted only between the last two or three

chambers, early sutures oblique or displaying even a suggestion 

of coiling, later ones transverse; aperture eccentric, protruding, 

radiate. Length up to 1.5 mm.; average 1 mm.—Plumrner.

The types are from the Paleocene, from fossiliferou^ 

clays, 2^o miles northwest of Kidgeway around the bend 

of a small creek that flows northward into South Sulphur River, Hopkins County, Tex.

This is a widely distributed species in the Paleocene 

and should make a good index fossil.

Clayton formation, Chalybeate limestone member. Alabama, 

Sumter County (9).

Porters Creek clay. Tennessee, Hardeman County (13).

Porters Creek formation, Matthews Landing marl member. Alabama, Choctaw County (7).

Midway formation, lower part. Arkansas, Saline County (20).

Midway formation, upper part. Arkansas, Pulaski County (25).

Midway formation. Arkansas, Saline County (26) ; Pulasl i 

County (27, 28) ; Clark County (29, 31).

Midway group. Texas, Hunt County (41) ; Bastrop County 

(43) ; Travis County (45, 46) ; Van Zandt County (47) ; Freestone County (50).

Kincaid formation. Texas, Hunt County (37) ; Bastrop County 

(38).

Dentalina colei Cushman and Dusenbury 

Plate 6, figures 8-10

Dentalina colei Cushman and Dusenbury, Cushman Lab. Foram.

Research Contr., vol. 10, p. 54, pi. 7, figs. 10-12, 1934. 

Parr, Royal Soc. West Australia Jour., vol. 24, p. 76, pi. 1,

fig. 8,1937-38. 

Toulmin, Jour. Paleontology, vol. 15, p. 584, pi. 79, fig. 12,.

1941.

Beck, idem, vol. 17, p. 598, pi. 105, fig. 18,1943. 

Curran, Am. Assoc. Petroleum Geologists Bull., vol. 27, pri.

1378, 1381 (lists), 1943. 

Martin, Stanford Univ. Publ., Univ. Ser., Geol. Sci., vol. 3,

no. 3, p. 10 (list), 1943. 

Cushman, Cushman Lab. Foram. Research Contr., vol. 2\

p. 37, pi. 5, figs. 25-28, 1944.

Cushman and Todd, idem, vol. 22, p. 49, pi. 8, fig. 2, 194<5. 

Vaffinulina legmnen (Linn6) var. elegans Cole (not D'Orbigny),

Bull. Am. Paleontology, vol. 14, no. 51, p. 21, pi. 3, figs.

10, 11, 1927. ' 

Plummer, Texas Univ. Bull. 2644, p. 110, pi. 6, fig. 1, 1927.

Test elongate, tapering, with the greatest width ner.r 

the apertural end, slightly curved; chambers distinct, 

earliest ones slightly compressed, later ones circular in 

transverse section, increasing very gradually in size 

and height as added, little if at all inflated; sutures flu^h 

with the surface, slightly oblique; wall smooth; aperture radiate, toward the inner side of the apertural 

face. Length 1.00 to 2.50 mm., diameter 0.12 to 0.2T 

mm.

The types are from the Eocene Poway conglomerate, 

Murray Canyon, LaJolla Quadrangle, San Diego 

County, Calif.

Some specimens tend toward the form of D. gardnerae (Plummer), but the test is more slender and more 

curved than in that species. It has a wide distribution.

Porters Creek clay. Mississippi, Kemper County (17). 

Porters Creek formation. Matthews Landing marl member. 

Alabama, Butler County (3).
20 PALEOCENE FORAMINHFERA OF GULF COASTAL REGION

Naheola formation, Coal Bluff marl member. Alabama, Wilcox

County (8). 

Clayton formation, Pine Barren member. Alabama, Wilcox

County (11). Chalybeate limestone member. Mississippi,

Kemper County (16).

Midway formation, lower part. Arkansas, Saline County (21). 

Midway formation, upper part. Arkansas, Pulaski County (25). 

Midway formation. Arkansas, Saline County (26) ; Pulaski

County (27) ; Clark County (29, 31). 

Midway group. Texas, Limestone County (39, 42) ; Hunt

County (41) ; Caldwell County (44) ; Van Zandt County (47) ;

Freestone County (50). 

Wills Point formation. Texas, Caldwell County (34). Mexia

member. Texas, Limestone County (36). 

Kincaid formation. Texas, Hunt County (37).

Dentalina plummerae Cushman 

Plate 6, figures 11-15

Dentalina plummerae Cushman, Cushman Lab. Foram. Research Contr., vol. 16, p. 57, pi. 10, figs. 7-9,19?, 1940.

Kline, Mississippi Geol. Survey Bull. 53, p. 25, pi. 2, fig. 6, 

1943.

Cushman and Todd, Cushman Lab. Foram. Research Contr., 

vol. 22, p. 50, pi. 8, fig. 3,1946.

Test slender, elongate, slightly curved, tapering, 

greatest breadth at the last-formed chamber, initial end 

rounded in the megalospheric form, pointed in microspheric ; chambers distinct, rapidly increasing in size as 

added, the later ones much inflated, subspherical; 

sutures of the later portion very strongly depressed, 

limbate; wall smooth; aperture with a distinct, rather 

elongate, tapering neck, radiate. Length 1.30 to 2.00 

mm., diameter 0.20 to 0.32 mm.

The types are from the Paleocene, U. S. Highway 80, 

south of Sucarnoochee Creek, y2 mile southwest of Livingston, Sumter County, Ala.

Most of the specimens in our series are from the Paleocene of Texas but it occurs at a few stations in other 

areas. It has been rather rare at all localities. Specimens referred to as "Nodosaria soluta (Reuss)" (Plummer, Texas Univ. Bull. 2644, p. 78, pi. 4, fig. 10, 1927) 

may belong here.

Porters Creek clay. Mississippi, Kemper County (17).

Porters Creek formation, Matthews Landing marl member. Alabama, Butler County (3).

Clayton formation? Chalybeate limestone member. Alabama, 

Sumter County (9).

Midway formation, upper part. Arkansas, Pulaski County (25).

Midway formation. Arkansas, Saline County (26) ; Pulaski 

County (27) ; Clark County (29, 30).

Midway group. Texas, Hunt County (41) ; Limestone County 

(42) ; Caldwell County (44).

Wills Point formation. Texas, Travis County (33).

Dentalina insulsa Cushman 

Plate 6, figures 16,17

Dentalina insulsa Cushman, Cushman Lab. Foram. Research 

Contr., vol. 23, p. 84, pi. 18, figs. 6, 7,1947.

Marginulina costata Plummer (not Batsch), Texas Univ. Bull. 

2644, p. 107, pi. 5, figs. 8a-c, 1927.

Dentalina sp. B Cushman and Todd, Cushman Lab. Foram. Research Contr., vol. 22, p. 51, pi. 8, figs. 15,16,1946.

Test slightly curved, the megalospheric form with 

very little increase in diameter from the prolo^ulum, .the 

microspheric form more tapering and compressed, initial end often with a distinct spine; chambers inflated 

in the adult portion, increasing very gradually in size 

as added and becoming more distinct; sutures distinct 

except in the early stages of the microspheric form 

where they are obscured by the costae of the wall, in the 

later portion depressed; wall ornamented wi*:h numerous, distinct, longitudinal costae, often slightly oblique 

to the longitudinal axis especially in the microspheric 

form; aperture radiate, usually at one side of the axis 

of the test, only slightly projecting. Length 1.00 to 1.40 

mm., diameter 0.20 to 0.30 mm.

The types are from the Paleocene, about 1,000 feet 

south of Roosevelt Road viaduct, 200 feet east of railroad tracks, in small gully heading east, just upstream 

from small abandoned bridge, NE^NWi/i se-,. 16, T. 1 

N., R. 12 W., Little Rock, Ark. (25).

This species differs from D. pseitdo-nasuta Cushman 

and Todd in the much shorter apertural necK broader 

and generally larger test, the costae somewhat oblique, 

and the less distinct chambers in the early portion.

Porters Creek clay. Tennessee, Hardeman County (13). 

Midway formation, upper part. Arkansas, Pulaski County

(25). 

Midway formation. Arkansas, Saline County (fft) ; Clark

County (30).

Midway group. Texas, Limestone County (39, 42). 

Wills Point formation. Texas, Travis County (33).

Dentalina naneolensis Cushman and Todd 

Plate 6, figure 18

Dentalina delicatula Cushman var. naheolensis Cushman and

Todd, Cushman Lab. Foram. Research Contr., vol. 18, p. 30,

pi. 5, figs. 14, 15, 1942. 

Dentalina naheolensis Cushman, idem. vol. 20, p. 35, pi. 5, fig.

17, 1944. Cushman and Todd, idem, vol. 22, F. 50, pi. 8,

fig. 8,1946.

Test small, slightly curved, initial end with a slight 

spine, apertural end with the distinct neck at the margin; chambers few, usually 4 or 5 in the adult, somewnat inflated, of nearly uniform size and shap?-; sutures 

distinct, slightly depressed and somewhat oblioue; wall 

ornamented with numerous fine longitudinal, slightly 

oblique costae; aperture radiate, terminal w; th a distinct, slightly tapering neck. Length of holotype 0.65 

mm., diameter 0.10 mm.

The types are from the Paleocene Naheola formation, 

greensand bed, upper fossiliferous horizon, Naheola 

Landing, Tombigbee River, Ala. (7).

Most of the records are from the Naheola formation, 

with a very few elsewhere. It should make a good index fossil for the Paleocene.

Porters Creek clay. Tennessee, Hardeman County (13, 14). 

Porters Creek formation. Matthews Landing mar? member.

Alabama, Wilcox County (1, 2, 4-6) ; Butler County (3) ;

Choctaw County (7).
LAGENIDAE 21

Naheola formation. Coal Bluff marl member. Alabama, Wilcox County (8). 

Clayton formation, Pine Barren member. Alabama, Wilcox

County (11); Butler County (12). 

Midway formation, lower part. Arkansas, Saline County

(21,22).

Midway formation, upper part. Arkansas, Pulaski County (25). 

Midway formation. Arkansas, Pulaski County (27). 

Kincaid formation. Texas, Hunt County (37) ; Bastrop

County (38).

Dentalina pseudo-nasuta Cushman and Todd 

Plate 6, figures 19, 20

Dentalina, pseudo-nasuta, Cushman and Todcl, Cushman Lab.

Foram. Research Contr., vol. 22, p. 51, pi. 8, figs. 13, 14,

1946. 

Dentalina cf. D. pungens Cushman (not Reuss), idem, vol. 15, p.

57, pi. 10, fig. 9, 1939.

Test small, elongate, slender; initial end with a single 

long spine, apertural end produced; chambers distinct, 

inflated, increasing very gradually in width as added, 

becoming more inflated in the adult portion; sutures 

distinct, somewhat depressed, slightly oblique; wall 

ornamented with numerous low costae, often slightly 

oblique on the earlier chambers; aperture terminal, 

slightly on the concave side of the test, radiate, projecting and tapering to a pointed end. Length 0.65 to 1.05 

mm., breadth 0.10 to 0.16 mm.

The types are from the Paleocene, about 1,000 feet 

south of Roosevelt Road viaduct,. 200 feet east of railroad tracks in small gully heading east, just upstream 

from small abandoned bridge, NE 14 NW *4 sec. 16, 

T. 1 N., R. 12 W., Little Rock, Ark. (25).

Similar specimens were recorded as Dentalina cf. 

D. pungens Reuss from the Eocene of cores from the 

western Atlantic.

Dentalina alabamensis Cushman 

Plate 6, figures 21-24

Dentalina, alabamensis Cushman, Cushman Lab. Foram. Research Contr., vol. 20, p. 35, pi. 5, figs. 18-21,1944.

Test slender, initial end pointed, sometimes with a 

slight spine, gently tapering, one side nearly straight or 

slightly concave, the other slightly convex; chambers 

few, distinct, the last ones in the adult slightly inflated, 

earlier ones small and slightly coiled; sutures distinct, 

earlier ones strongly oblique and slightly curved, becoming slightly separated; sutures of the early portion 

little if at all depressed, gradually becoming deeply depressed in the adult; wall ornamented with numerous 

longitudinal costae, as many as 20 in the adult chambers; aperture radiate, terminal, with a prominent, 

slightly tapering neck. Length of holotype, 1.17 mm., 

diameter 0.20 mm.

The types are from the Paleocene Coal Bluff marl 

member of the Naheola formation, creek bottom, just 

west of store at Caledonia, about 14 mile south of center 

of sec. 29, T. 11 N., R. 10 E., Wilcox County, Ala. (8). 

The species has occurred only at the type locality.

Dentalina inepta Cushman

Plate 6, figure 25 

Dentalina inepta Cushman, Cushman Lab. Foram. Research

Contr., vol. 23, p. 83, pi. 18, fig. 5, 1947.

Dentalina sp. A Cushman and Todd, Cushman Lab. Foram. 

Research Contr., vol. 22, p. 51, pi. 8, fig. 10.

Test elongate, slender, slightly curved, later portion 

slightly compressed, basal end broadly rounded; 

chambers of nearly uniform width, distinctly inflated, 

increasing very slightly in size as added; sutures distinct, depressed, slightly oblique; wall smooth; aperture at the inner margin of the last-formed chamber, 

radiate, slightly projecting. Length 0.60 to 1.00 mm.; 

breadth 0.16 to 0.25 mm.

The types are from the Paleocene, about 1,000 feet 

south of Roosevelt Road viaduct, 200 feet east of railroad tracks, in small gully heading east, just upstream 

from small abandoned bridge, NE 14 NW 14 sec. 16, 

T. 1 N., R. 12 W., Little Rock, Ark. (25).

The species differs from Z>. colei Cushman and 

Dusenbury in the less curved and more slender test, 

which is slightly compressed, and the nearly uniform 

size of the chambers.

The species occurs at several localities in Arkansas. 

Texas, and Alabama.

Porters Creek formation, Matthews Landing marl member.

Alabama, Butler County (3).

Midway formation, upper part. Arkansas, Pulaski County (25). 

Midway group. Texas, Caldwell County (44).

Dentalina aculeata D'Orbigny

Plate 6, figure 26 

Dentalina aculeata D'Orbigny, Soc. g6ol. France Mem., ser. 1,

vol. 4, p. 13, pi. 1, figs. 2, 3,1940. 

Cushman and Todd, Cushman Lab. Foram. Researcl

Contr., vol. 22, p. 51, pi. 8, fig. 9,1946. 

A few specimens from the Paleocene of Texas anc1 

Arkansas resemble this species, described from the 

Cretaceous of France and recorded from the Upper 

Cretaceous of America.

Clayton formation, Chalybeate limestone member. Mississippi,

Noxubee County (16).

Midway formation, upper part. Arkansas, Pulaski County (25). 

Midway group. Texas, Limestone County (39); Caldwer

County (44). 

Wills Point formation, Mexia member. Texas, Limestone

County (36).

Dentalina wilcoxensis Cushman 

Plate 6, figures 27-29

Dentalina wilcoxensis Cushman, Am. Jour. Sci., vol. 242, p. 8, 

pi. 1, figs. 5, 6,1944; Cushman Lab. Foram. Research Contr., 

vol. 20, p. 22, pi. 4, fig. 10; p. 36, pi. 5, figs. 22-24, 1944.

Dentalina, sp. Cushman and Todd, idem, vol. 18, p. 30, pi. 5 

fig. 20, 1942.

Test small, elongate, of nearly uniform diameter 

throughout, circular or slightly compressed in transverse section, rounded at the initial end; chambers few, 

slightly if at all inflated, increasing only slightly in 

height as added; sutures distinct but not depressed,
22 PALEOC'ENE FOKAMINHFERA OF GULF COASTAL REGION

strongly oblique; wall smooth; aperture terminal, 

rounded, with the border finely toothed. Length 0.50 

mm., diameter 0.10 to 0.12 mm.

The types of this species are from the Eocene Bashi 

marl member of the Hatchetigbee formation, of the 

Wilcox group of Alabama. Specimens that seem referable to this species occur rarely in the Paleocene.

Midway formation, upper part. Arkansas, Pulaski County (25).

Dentalina eocenica Cushman 

Plate 6, figures 30-33

Dentalina eocenica Oushman, Cushman Lab. Foram. Research 

Contr., vol. 20, p. 36, pi. 6, fig. 1, 1944.

Cushman and Todd, idem, vol. 22, p. 50, pi. 8, figs. 4, 5,1946. 

Nodosaria pauperata Plummer (not D'Orbigny), Texas Univ.

Bull. 2644, p. 79, pi. 4, fig. 11, 1927. 

Nodosaria cf. N. pauperata Cushman and Todd, Cushman Lab.

Foram. Research Contr., vol. 18, p. 29, pi. 5, figs. 17, 18,

1942.

Test small, slender, slightly curved, increasing very 

slightly in diameter; chambers distinct, but only 

slightly inflated in the later portion, increasing gradually and rather regularly in size as added, the lastformed one in the adult nearly twice as high as broad; 

sutures distinct, depressed slightly in the later portion, 

earlier ones slightly oblique, later ones nearly at right 

angles to the long axis of the test; wall smooth; aperture 

radiate, terminal, nearly in the center of the terminal 

face. Length 0.75 to 0.85 mm., diameter 0.12 to 

0.14 mm.

The types are from the Paleocene Coal Bluff marl 

member of the Naheola formation, creek bottom, just 

west of store at Caledonia, about y± niile south of center 

of sec. 29, T. 11 N., R. 10 E., Wilcox County, Ala.

It is possible that the incomplete form figured by 

Cooper (Jour. Paleontology, vol. 18, p. 347, pi. 54, fig. 

21, 1944) as "Dentalina cf. D. cooperensis Cushman," 

from the Paleocene of Illinois, may belong here.

Porters Creek formation, Matthews Landing marl member.

Alabama, Wilcox County (1, 2, 4, 5) ; Butler County (3). 

Naheola formation, Coal Bluff marl member. Alabama, Wilcox

County (8).

Clayton formation, Pine Barren member. Alabama, Butler 

County (12). 

Chalybeate limestone member. Mississippi, Noxubee

County (16). 

Midway formation, upper part. Arkansas, Pulaski County (24,

25).

Midway formation. Arkansas, Saline County (26). 

Wills Point formation. Texas, Caldwell County (34). 

Mexia member. Texas, Limestone County (36).

Dentalina cf. D. mucronata Neugefooren 

Plate 6, figures 34, 35

Specimens similar to those figured from the Paleocene of Arkansas (Cushman and Todd, Cushman Lab. 

Foram. Research Contr., vol. 22, p. 51, pi. 8, fig. 12, 

1 946) and from the Naheola formation of Alabama

(idem, vol. 16, p. 57, pi. 10, fig. 27, 1940) occur at a 

number of other stations in Alabama.

Porters Creek formation, Matthews Landing marl member.

Alabama, Butler County (3) ; Wilcox County (6) ; Choctaw

County (7). 

Clayton formation, Pine Barren member. Alabama, Butler

County (12). 

Midway formation, upper part. Arkansas, Pulaski County

(25).

Dentalina nasuta Cushman

Plate 6, figures 37, 38

Dentalina nasuta Cushman, Cushman Lab. Forarr, Research

Contr., vol. 15, p. 57, pi. 10, figs. 10, 11, 1939. 

Cushman and Todd, idem, vol. 22, p. 50, pi. 8, figs'. 6, 7, 1946.

Test elongate, slender, slightly curved, gradually 

tapering from the subacute initial end to the greatest 

breadth at the last-formed chamber, apertural end produced into a long, tapering cone; chambers distinct, 

somewhat inflated toward the apertural end, increasing 

rather rapidly in size and height in the adult portion, 

slightly overlapping; sutures distinct, depressed in the 

later portion, somewhat oblique; wall smooth; aperture radiate, at the end of a long, conical, tapering projection of the apertural end of the last-formed chamber. 

Length up to 1.75 mm., diameter 0.15 to 0.25 mm.

The types are from the Eocene of a core taken off the 

Atlantic coast of North America.

Midway formation, upper part. Arkansas, Pulaski County (25). 

Midway group. Texas, Bastrop County (43).

Dentalina cf. D. pauperata D'OrMgny 

Plate 6, figure 36

Rare specimens from the Paleocene of Alabama and 

Texas seem closely related to D. pauperata but not 

identical. Not enough specimens are available for a 

full specific determination.

Porters Creek formation, Matthews Landing marl member.

Alabama, Choctaw County (7).

Clayton formation. Alabama, Barbour County (10). 

Chalybeate limestone member. Alabama, Sumter Crunty (9). 

Wills Point formation. Texas, Caldwell County (34). 

Midway group. Texas, Limestone County (39) ; Travis County

(45).

Dentalina sp. A

Plate 6, figure 39

This form occurs as very rare specimens s.t several 

localities in Texas and Alabama. Not enough specimens are available to warrant a specific identification. 

It somewhat resembles D. inepta Cushman tut is not 

compressed and the chambers are nearly spherical.

Porters Creek formation, Matthews Landing mar1 member.

Alabama, Wilcox County (5). 

Clayton formation, Pine Barren member. Alabama, Butler

County (12).

Kincaid formation. Texas, Bastrop County (38). 

Midway group. Texas, Travis County (45).
LAGENIDAE 23

Dentalina sp. B 

Plate 6, figure 41

Dentalina sp. Cushman, Cushman Lab. Foram. Research Contr., 

vol. 16, p. 57, pi. 10, fig. 13,1940.

Specimens from two localities in the Paleocene of 

Texas are similar to those found in the Paleocene of 

Alabama but are very rare and not specifically identifiable. The species nearest to this is probably D. alternata (Jones), from the Upper Cretaceous, and these 

may possibly be reworked specimens.

Clayton formation, Chalybeate limestone member. Alabama,

Sumter County (9). 

Wills Point formation, Mexia member. Texas, Limestone

County (36). 

Midway group. Texas, Limestone County (39) ; Caldwell

County (44).

Dentalina sp. C 

Plate 6, figure 40

Dentalina sp. C Cushman and Todd, Cushman Lab. Foram. 

Research Contr., vol. 22, 52, pi. 8, fig. 17, 1946.

A few very slightly compressed specimens, similar to 

that figured, are insufficient to warrant description.

Midway formation, upper part. Arkansas, Pulaski County 

(25).

Genus NODOSARIA Lamarck, 1812

Nodosaria affinis Reuss

Plate 7, figures 3-6

Nodosaria, affinis Rouss, Versteinerungen bohm. Kreideformation, pt. 1, p. 26, pi. 13, fig. 16, 1845; Palaeontographica,

vol. 20, pt. 2, 1872-75, p. 83, pi. 2 (20), fig. 12, 1874. 

Perner, Forani. Ceskeho Cenomami, p. 57, pi. 6, figs. 10,

14a, b, 1892. 

Franke, Greifswald Univ., Geol.-paleont. Inst, Abh., vol. 6,

p. 37, pi. 3, fig. 25, 1925. 

W. Berry and Kelley, U. S. Nat. Mus. Proc., vol. 76, art. 19,

p. 6, pi. 1, fig. 8, 1929. 

Cushman, Tennessee Div. Geology Bull. 41, p. 30, pi. 3,

figs. 16-20, 1931; Jour. Paleontology, vol.-5, p. 305, pi.

35, figs. 3-5 (not fig. 2), 1931; Cushman Lab. Foram.

Research Contr., vol. 7, p. 38, pi. 5, fig. 4, 1931. 

Cushman and Jarvis, U. S. Nat. Mus. Proc., vol. 80, art. 14,

p. 34, pi. 10, fig. 13, 1932. 

Cushman, Geol. Soc. America Bull., vol. 47, p. 417, 1936;

Cushman Lab. Foram. Research Contr., vol. 16, p. 59, pi.

10, figs. 30-33; p. 86, pi. 15, figs. 8-23, 1940; vol. 18, p. 30,

pi. 5, figs. 27-29; p. 58, 1942.

Cushman and Renz, idem, vol. 18, p. 6, pi. 1, figs. 8-10, 1942. 

Cushman and Todd, idem, vol. 19, p. 57, pi. 10, fig. 11, 1943. 

Cushman, idem, vol. 20, p. 7, pi. 2, fig. 1; p. 37, pi. 6, figs.

2-5; p. 87, pi. 13, fig. 18, 1944. 

Cushman and Deaderick, Jour. Paleontology, vol. 18, p. 333-

pi. 51, figs. 19-21, 1944. 

Cushman and Todd, Cushman Lab. Foram. Research Contr.,

vol. 22, p. 52, pi. 8, figs. 21-24, 1946. 

Cushman, U. S. Geol. Survey Prof. Paper 206, p. 70, pi. 25,

figs. 8-23, 1946.

Cushman and Renz, Cushman Lab. Foram. Research Special Pub. 18, p. 30, pi. 5, fig. 1, 1946.

Nodosaria vertebralis Plummer (not Batsch), Texas Univ. Bull. 

2644, p. 88, pi. 5, fig. 10, 1927.

Nodosaria proximo, W. Berry and Kelley (not Silvestri), U. S. 

Nat. Mus. Proc., vol. 76, art. 19, p. 7, pi. 1, fig. 13, 1929.

Nodosaria cf. N. vertebralis (Batsch) Cooper, Jour. Paleontology, vol. 18, p. 350, pi. 55, figs. 14,15, 1944.

Nodosaria cf. N. sippei Reuss, Cooper, idem, p. 350, pi. 55, figs. 

18,19,1944.

Test elongate, of differing shape in the microspheric 

and megalospheric forms; the former with many chambers and tapering, the greatest width near the apertural 

end; the latter with the chambers of nearly uniform 

diameter throughout; chambers distinct, inflated, especially toward the apertural end, initial end usually 

with a stout spine; sutures distinct, depressed, often 

somewhat limbate; wall ornamented by numerous 

(usually 13 to 15) longitudinal costae, continuous over 

the adjacent chambers, usually sharp and plate-like; 

aperture radiate, terminal, with a slight projection of 

the apertural face. Length up to 2.00 mm. or more, 

diameter normally about 0.30 mm., but in extreme 

megalospheric forms may be as much as 0.75 mm.

This species apparently has a very long range and 

wide distribution. It occurs throughout most of the 

American Upper Cretaceous and is common in the 

Paleocene and later formations.

Porters Creek formation, Matthews Landing marl member.

Alabama, Wilcox County (2, 4) ; Butler County (3) ; Choctaw

County (7).

Naheola formation. Coal Bluff marl member. Alabama, Wilcox County (8). 

Clayton formation, Chalybeate limestone member. Alabama,

Sumter County (9). Mississippi, Noxubee County (16). 

Pine Barren member. Alabama, Butler County (12). 

Midway formation, lower part. Arkansas, Saline County (18,

20). 

Midway formation, upper part. Arkansas, Pulaski County

(25). 

Midway formation. Arkansas, Saline County (26) ; Pulaski

County (27). 

Midway group. Texas, Hunt County (41); Bastrop County

(43, 49) ; Caldwell County (44); Van Zandt County (47, 48) ;

Freestone County (50). 

Wills Point formation. Texas, Travis County (33) ; Williamson

County (35).

Nodosaria latejugata Giimbel 

Plate 7, figures 1, 2

Nodosaria latejugata Gumbel, K. bayer. Akad. Wiss., Math.-phys.

Abt, Abh., Kl. 2, vol. 10, p. 619, pi. 1, fig. 32,1870. 

Kline, Mississippi Geol. Survey Bull. 53, p. 27, pi. 2, fig. 10,

1943. 

(?) Cooper, Jour. Paleontology, vol. 18, p. 348, pi. 55, figs.

24, 25, 1944. 

Cushman, Cushman Lab. Foram. Research Contr., vol. 20,

p. 37, pi. 6, figs. 6-8, 1944. 

Cushman and Todd, idem, vol. 22, p. 52, pi. 8, figs. 19, 20,

1946.

Nodosaria affinis Plummer (not Reuss), Texas Univ. Bull. 2644, 

p. 89, pi. 14, figs. 2a-d, 1927.

There are many records in the literature for this 

species but many of them are without figures and only
24 PALEOCENE FORAMINIFERA OF GULF COASTAL REGION1

those that relate to this particular fauna are given here. 

It differs from Nodosaria affinis Eeuss mainly in the 

greater inflation of the chambers and consequent 

strongly depressed sutures. Some specimens are rather 

difficult to allocate and the two species are evidently 

closely related. N. latejugata is apparently not found 

in the Upper Cretaceous.

Porters Creek formation, Matthews Landing marl member. Alabama, Butler County (3).

Clayton formation, Chalybeate limestone member. Mississippi, 

Noxubee County (16).

Midway formation, lower part. Arkansas, Saline County (21).

Midway formation, upper part. Arkansas, Pulaski County (25).

Midway formation. Arkansas, Saline County (26) ; Pulaski 

County (28) ; Clark County (29, 31).

Midway group. Texas, Limestone County (39, 42) ; Hunt 

County (41) ; Travis County (45, 46) ; Van Zandt County (47, 

48) ; Bastrop County (49) ; Freestone County (50).

Wills Point formation. Texas, Caldwell County (34) ; Williamson County (35).

Kincaid formation. Texas, Hunt County (37) ; Bastrop County 

(38).

Nodosaria macneili Cushman

Plate 7, figure 7

Nodosaria macneili Cushman, Cushman Lab, Foram. Research

Contr., vol. 20, p. 37, pi. 6, fig. 9,1944. 

Nodosaria paupercula Toulmin (not Reuss), Jour. Paleontology,

vol. 15, p. 588, pi. 79, figs. 29, 30,1941.

Test slightly tapering, nearly straight, of rather few 

chambers, basal end rounded; chambers inflated; later 

ones nearly spherical and more separated; sutures of 

the early portion little if at all depressed, gradually becoming deeply depressed in the adult; wall ornamented 

with numerous longitudinal costae, as many as 20 in the 

adult chambers; aperture radiate, terminal, with a 

prominent, slightly tapering neck. Length of holotype 

1.17 mm., diameter 0.20 mm.

The types are from the Paleocene Coal Bluff marl 

member of the Naheola formation, creek bottom, just 

west of store at Caledonia, about *4 mile south of center 

of sec. 29, T. 11 N., B. 10 E., Wilcox County, Ala. (8).

The species is evidently limited in its distribution, occurring in our material only in the Naheola formation 

of Alabama and the Clayton formation of Tennessee.

Naheola formation, Coal Bluff marl member. Alabama, Wilcox

County (8). 

Porters Creek clay. Tennessee, Hardeman County (13, 14).

Nodosaria of. N. amphioxys Reuss 

Plate 7, figure 8

Nodosaria cf. N. amphioays Reuss, Cushman and Todd, Cushman Lab. Foram. Research Contr., vol. 22, p. 52, pi. 8, fig. 25, 

1946. 

Nodosaria oligotoma Plumrner (not Reuss), Texa*s Univ. Bull.

2644, p. 87, pi. 4, fig. 14, 1927. 

Cushman, Cushman Lab. Foram. Research Contr., vol. 16,

p. 59, pi. 10, figs. 25, 26,1940.

Kline, Mississippi Geol. Survey Bull. 53, p. 27, pi. 2, fig. 16, 

1943.

Specimens from the American Paleocere have been 

referred to this species with some question. The types 

are from the Upper Cretaceous of Saxony r,nd it occurs 

in the upper portion of the Taylor marl and in the 

Navarro group of the southeastern United States. The 

references above are to Paleocene occurrences in this 

same area : Arkansas, Texas, Alabama, and Mississippi. 

They all seem to come under one species and are more 

like ^V. amphioxys than ^V. oligotoma.

Porters Creek formation, Matthews Landing marl member.

Alabama, Butler County (3). 

Clayton formation, Chalybeate limestone member. Mississippi,

Noxubee County (16). 

Midway formation, upper part. Arkansas, Pulaski County

(24, 25).

Midway group. Texas, Caldwell County (44). 

Wills Point formation. Texas, Caldwell County (34).

Nodosaria? cf. N. longiseata D'Orbigny 

Plate 7, figures 9, 10

sfiria Jonyiscata Plummer, Texas Univ. Bull. 2644, p. 82,

pi. 4, fig. 17, 1927. 

Cushman, Cushman Lab. Foram. Research Contr., vol. 16,

p. 59, pi. 10, fig. 28, 1940.

Cushman and Todd, idem, vol. 22. p. 52, pi. 8, %. 26, 1944. 

Nodosarina (Nodosaria) longiscata Kline, Mississippi Geol. 

Survey Bull. 53, p. 28, pi. 2, fig. 8, 1943.

Very rare specimens from the Paleocene of Texas, 

Arkansas, Mississippi, and Alabama are referred to this 

species with considerable question.

Porters Creek formation, Matthews Landing marl member.

Alabama, Choctaw County (7). 

Clayton formation, Pine Barren member. Alabama, Butler

County (12). 

Midway formation, upper part. Arkansas, Pulaski County

(25). 

Midway group. Texas, Limestone County (39) ; Van Zandt

County (47). 

Wills Point formation, Mexia member. Texas, Limestone

County (36).

Genus CHRYSALOGONIUM Schubert, 1907 

Chrysalogonium grant! (Plummer) Toulmin 

Plate 13, figure 11

Nodosaria granti Plummer, Texas Univ. Bull. 264^ p. 83, pi. 5,

figs. Qa-d, 1927. 

Dentalina granti Plummer, idem, Bull. 3101, p. 149, pi. 11, figs.

8, 9, 1931. 

Jennings, Bull. Am. Paleontology, vol. 23, no. 78, p. 176,

pi. 29, fig. 6, 1936. 

Ellipsonodosariaf granti Cushman, Cushman Lab. Foram. Research Contr., vol. 12, p. 51, pi. 9, figs. 3-5, 1936. 

Cushman, idem, vol. 16, p. 69, pi. 12, fig. 3, 1940. 

Cushman and Todd, idem, vol. 19, p. 67, pi. 11. fig. 30, 1943. 

Cushman, U. S. Geol. Survey Prof. Paper 2C6, p. 136, pi.

56, figs. 24-26, 1946. 

Clirysalogonium granti Toulmin, Jour. Paleontology, vol. 15, p.

589, pi. 79, figs. 34, 35, 1941. 

Kline, Mississippi Geol. Survey Bull. 53, p. 29, pi. 2, fig. 9,

1943.

Cooper, Jour. Paleontology, vol. 18, p. 346, p1 . 54, figs. 26, 

27, 1944.
LAGENIDAE 25

Test very elongate, slightly arcuate, slightly tapering, initial end with a stout spine; chambers numerous, 

later ones becoming slightly inflated, sometimes but not 

always pyriform in shape, increasing in relative length 

as added; sutures transverse, indistinct, sometimes unobservable in the early part of the test, later becoming 

progressively more constricted; wall smooth, thick, 

opaque; aperture terminal, a bluntly pointed cribrate 

cone. Length up to 3 mm. or more, diameter up to 

0.25 mm., usually less.

This species has been recorded frequently from the 

Upper Cretaceous and lower Eocene under various 

generic names. The test is very easily broken and 

usually only fragments are found. Recently Mrs. 

Plummer sent me a specimen showing the aperture, 

which is characteristic of Chrysalogonivm and the species is here included under that genus. It is common in 

our Paleocene material.

Clay ton formation, Chalybeate limestone member. Alabama,

Sumter County (9). Mississippi, Noxubee County (16). 

Midway formation. Arkansas, Saline County (26) ; Pulaski

County (27) ; Clark County (30, 31). 

Midway group. Texas, Hunt County (41) ; Caldwell County

(44) ; Travis County (45, 46) ; Van Zandt County (47) ;

Freestone County (50). 

Wills Point formation. Texas, Travis County (33) ; Caldwell

County (34). 

Kincaid formation. Texas, Hunt County (37).

Chrysalogonium arkansasanum Cushman and Todd 

Plate 7, figures 11,12

Chrysalogonium arkansasanum Cushman and Todd, Cushman 

Lab. Foram. Research Contr., vol. 22, p. 52, pi. 9, figs. 1, 2, 

1946.

Chrysalogonium cf. C. texanum Cushman, idem, vol. 16, p. 60, 

pi. 10, fig. 14, 1940.

Test very elongate, nearly straight to slightly curved, 

very slightly tapering, widest toward the apertural end, 

initial end with a short spine; chambers distinct, 

numerous, slightly inflated, averaging about twice as 

long as broad, increasing very slightly in relative length 

as added; sutures distinct, very slightly depressed; 

wall smooth; aperture terminal, cribrate, raised in a 

blunt cone. Length up to 3.75 mm., diameter up to 

0.25 mm.

The types are from the Paleocene, about 1,000 feet 

south of Roosevelt Road viaduct, 200 feet east of railroad tracks, in small gully heading east, just upstream 

from small abandoned bridge, NE%NW^4 sec. 16, T. 

1 N., R. 12 W., Little Rock, Ark. (25).

This species is very close to G. granti (Plummer) 

but differs in having more evenly inflated and nonpyriform chambers.

Porters Creek formation, Matthews Landing marl member.

Alabama, Butler County (3). 

Clayton formation, Chalybeate limestone member. Alabama,

Sumter County (9). 

Midway formation, upper part. Arkansas, Pulaski County (25).

Chrysalogonium eocenicum Cushman and Todd 

Plate 7, figures 13-15

Chrysalogonium eocenicum Cushman and Todd, Cushman Lab. 

Foram. Research Contr., vol. 22, p. 53, pi. 9, figs. 3-5, 1946.

Test elongate, slender, distinctly curved, tapering, 

widest toward the apertural end, initial end subacute; 

chambers distinct, numerous, slightly inflated, slightly 

longer than broad in the adult, increasing little in relative length as added; sutures distinct, depressed and 

slightly oblique in the earlier portion; wall smooth; 

aperture terminal, cribrate, little if at all projecting 

above the contour of the chamber. Length up to 1.25 

mm., breadth up to 0.15 mm.

The types are from the Paleocene, about 1,000 feet 

south of Roosevelt Road viaduct, 200 feet east of railroad tracks, in small gully heading east, just upstream 

from small abandoned bridge, NE*4NW*4 sec. 16, T. 

1 N., R. 12 W., Little Rock, Ark. (25).

This species is very constant in its characters and 

occurs at a number of localities in the Paleocene of 

Arkansas, Texas, and Alabama.

Porters Creek formation, Matthews Landing marl member.

Alabama, Wilcox County (2) ; Butler County (3). 

Clayton formation. Alabama, Barbour County (10). 

Midway formation, upper part. Arkansas, Pulaski County

(25).

Midway formation. Arkansas, Clark County (30). 

Midway group. Texas, Caldwell County (44).

Genus PSEUDOGLANDTTLINA Cushman, 1929 

Fseudoglandulina manifesta (Reuss) Cushman 

Plate 7, figures 16, 17

Glandulina manifesta Reuss, Haidinger's Naturwiss. Abh., vol.

4, pt. 1, p. 22, pi. 1, fig. 4,1851. 

Franke, Preuss. geol. Landesanstalt Abh., n. ser., vol. Ill,

p. 52, pi. 4, fig. 28, 1928. 

Cushman and Church, California Acad. Sci. Proc., 4th ser.

vol. 18, p. 511, pi. 39, fig. 10,1929.

Nodosaria manifesta Cushman, Am. Assoc. Petroleum Geologists Bull., vol. 10, p. 594, pi. 18, fig. 8,1926. 

Sandige, Jour. Paleontology, vol. 6, p. 278, pi. 42, fig. 8,1932. 

Pseudoglandulina manifesta Cushman, Cushman Lab. Foram.

Research Contr., vol. 16, p. 60, pi. 11, fig. 1, 1940. 

Toulmin, Jour. Paleontology, vol. 15, p. 590, pi. 79, fig. 32,

1941.

Cushman and Todd, Cushman Lab. Foram. Research Contr., 

vol. 18, p. 31, pi. 6, figs. 1, 2, 1942; vol 19, p. 58, pi. 10, 

fig. 15, 1943. 

Kline, Mississippi Geol. Survey Bull. 53, p. 30, pi. 2, fig. 12,

1943. 

Cushman and Todd, Cushman Lab. Foram. Research Contr.,

vol. 22, p. 53, pi. 9, figs. 6-9,1946. 

Cushman, U. S. Geol. Survey Prof. Paper 206, p. 76, pi. 27,

figs. 20-26, 1946. 

Nodosaria larva Carsey, Texas Univ. Bull. 2612, p. 31, pi. 2,

fig. 2, 1926. 

Nodosaria radicula Plummer (Not Linne"), idem. Bull. 2644, p.

77, pi. 4, fig. 9, 1927.

Nodosaria Jiumilis Cushman (not Roemer), Tennessee Div. 

Geology Bull. 41, p. 32, pi. 4, fig. 5, 1931.
26 PALEOCENE FOBAMINIFERA OF GULF COASTAL REGION

Test uniserial; the microspheric form rapidly tapering, initial end pointed, often with a small spine; the 

megalospheric form with a large proloculum forming 

a rounded base and the test much more nearly cylindrical; chambers in the early stages strongly overlapping with little or no inflation, in the adult stage much 

inflated; sutures in the early stages flush with the surface, in the adult depressed; wall smooth and polished; 

aperture terminal, radiate. Length 0.70 to 0.90 mm., 

diameter 0.25 to 0.35 mm.

The types are from the Upper Cretaceous of Lemberg. The species occurs in the American Upper 

Cretaceous, most abundantly in beds of Navarro and 

Taylor age. The Paleocene specimens seem identical 

with the Cretaceous ones but show some variation, 

especially in the rounded or acute base, evidently dependent on whether microspheric or megalospheric. 

Porters Creek formation, Matthews Landing marl member. Alabama, Choctaw County (7). 

Clayton formation, Chalybeate limestone member. Alabama,

Sumter County (9). 

Midway formation, upper part. Arkansas, Pulaski County

(25). 

Midway formation. Arkansas, Saline County (26) ; Pulaski

County (27). 

Midway group. Texas, Navarro County (40) ; Caldwell County

(44). 

Wills Point formation. Texas, Caldwell County (34).

Pseudoglandulina cf. P. caudigera (Schwager) Cushman 

Plate 7, figures 18, 19

Pseudoglandulina cf. P. caudigera- Schwager, Cushman* Lab.

Foram. Research Contr., vol. 16, p. 60, pi. 11, figs. 2, 3,

1940. 

Kline, Mississippi Geol. Survey Bull. 53, p. 29, pi. 2, fig. 17,

1943.

Specimens from the Paleocene of Alabama, Mississippi, and Texas have been referred with some question 

to Schwager's species, the types of which are from the 

lower Eocene of northern Africa. They are shorter 

and more tapering than typical specimens of P. manifesto, (Reuss) Cushman but otherwise are much like the 

microspheric forms of that species. The specimens referred to "Nodosaria (Glandulina) laemgata D'Orbigiiy var. occidentalis Cushman" by Mrs. Plummer 

(Texas Univ. Bull. 2644, p. 75, pi. 4, fig. 8, 1927) may 

belong to Pseudoglandulina caudigera.

Porters Creek clay. Mississippi, Kemper County (17). 

Porters Creek formation, Matthews Landing marl member.

Alabama, Wilcox County (2, 4, 5) ; Butler County (3). 

Midway formation. Arkansas, Pulaski County (27). 

Midway group. Texas, Bastrop County (43) ; Caldwell County

(44).

Pseudoglandulina pygmaea (Reuss) Cushman 

Plate 7, figures 20-22

Glandulina pygmaea Reuss, Haidinger's Naturwiss. Abh., vol.

4, pt. 1, p. 6, pi. 1, fig. 3, 1851.

Pseudoglandulina, pygmaea Cushman, Cushman Lab. Foram. 

Research Contr., vol. 16, p. 60, pi. 11, fig. 4, 1940.

Cushman and Todd, idem, vol. 22, p. 54, pi. 9, figs. 10, 11,

1946. 

Cushman, U. S. Geol. Survey Prof. Paper 206, p. 76, pi. 27,

figs. 27, 28, 1946. 

Nodosaria (Glandulina) comata Plummer (not Bitsch), Texas

Univ. Bull. 2644, p. 76, pi. 4, fig. 7, 1927.

Pseudoglandulina comata Kline (not Batsch), Mississippi Geol. 

Survey Bull. 53, p. 29, pi. 2, fig. 13, 1943.

Test short and stout, only a little longer than broad, 

circular in transverse section, initial ard pointed; 

chambers few, much overlapping, the final chamber 

making up more than half the test; sutures indistinct; 

wall ornamented by numerous longitudinal costae; 

aperture terminal, radiate. Length 0.30 to 0.50 mm., 

diameter 0.25 to 0.40 mm.

The types are from the Upper Cretaceous of Lemberg. It occurs in the American Cretaceous in beds 

of Navarro and Taylor age and similar specimens are 

widely distributed in the American Paleocene.

Clayton formation, Chalybeate limestone member. Alabama, 

Sumter County (9). Mississippi, Noxubee County (16).

Pine Barren member. Alabama, Butler County (12).

Midway formation, upper part. Arkansas, Pulaski County 

(24,25).

Midway formation. Arkansas, Saline County (26) ; Pulaski 

County (27) ; Clark County (29-31).

Midway group. Texas, Hunt County (41) ; Bastrop County 

(43) ; Travis County (45) ; Van Zandt County (47) ; Freestone County (50).

Kincaid formation. Texas, Bastrop County (38).

Pseudoglandulina madrugaensis Cushman and Bermudez

Plate 21, figure 4

Pseudoglandulina madrugaensis Cushman and Bermudez, Cushman Lab. Foram. Research Contr., vol. 24, p. 70, pi. 11, 

fig. 7, 1948.

Test elongate, circular in transverse section, rectilinear; chambers distinct, inflated, increasing very 

gradually in size as added; sutures distinct, depressed; 

wall ornamented with very fine longitudinal costae; 

aperture terminal, radiate. Length of holotype 1.05 

mm.; diameter 0.40 mm.

The types are from the Paleocene Madruga formation, under highway bridge on Central Sni Antonio, 

Madruga, Habana Province, Cuba.

The species differs from P. manifesta (Keuss) in the 

slightly higher chambers, deeper sutures, and finely 

costate surface.

Genus IINGKTLINA D'Orbigny, 1826

Lingulina naheolensis Cushman

Plate 7, figure 23

Lingulina naheolensis Cushman, Cushman Lab. Foram. Research Contr., vol. 23, p. 18,1947.

Lingulina minuta Cushman (not Franke), idem, vol. 20, p. 38, 

pi. 6, fig. 10, 1944.

Test minute, about twice as long as t^oad, compressed, tapering from the acute initial end to the 

greatest breadth at the middle of the last-formed chamber, thence narrowing to the bluntly angled, apertural
LAGENIDAE 27

end; chambers few, increasing rapidly in size as added, 

strongly overlapping; sutures distinct, not depressed, 

at right angles to the long axis of the test; wall smooth; 

aperture terminal, radiate, not projecting. Length of 

holotype 0.35 mm., breadth 0.15 mm., thickness 0.12 mm. 

The types are from the Paleocene Coal Bluff marl 

member of the Naheola formation, creek bottom, just 

west of store at Caledonia, about ^4 mile south of center 

of sec. 29, T. 11 N., E. 10 E., Wilcox County, Ala. (8).

Porters Creek formation, Matthews Landing marl member.

Alabama, Butler County (3). 

Naheola formation, Coal Bluff marl member. Alabama, Wilcox

County (8).

lingulina cf. 1. wilcoxensis Cushman and Ponton

Plate 7, figure 24

Lingulina cf. L. wilcoxensis Cushman and Todd, Cushman Lab. 

Foram. Research Contr., vol. 22, p. 54, pi. 9, fig. 12, 1946.

The only record for this form in the Paleocene is 

from Arkansas (25) ; it is represented by a single, incomplete specimen.

Genus SARACENARIA Defrance, 1824

Saracenaria trigonata (Plummet) Cushman

Plate 7, figures 25, 26

Cristellaria trigonata Plummer, Texas Univ. Bull. 2644, p. 101,

pi. 7, fig. 3,1927.

Saracenaria trigonata Cushman, Cushman Lab. Foram. Research CoQtr., vol. 16, p. 61, pi. 9, fig. 31, 1940. 

Kline, Mississippi Geol. Survey Bull. 53, p. 31, pi. 3, fig. 2,

1943.

Test elongate, triangular in cross section, tapering toward the 

oral end; peripheral margin carinate; early chambers closely 

coiled, later ones sharply carinate on each side of the septal 

face; sutures as dark lines; aperture at the apex of a long, broad, 

septal face on mature specimens. Length up to 0.7 mm.— 

Plummer.

The types are from the Paleocene, clay pit, Mexia 

Brick Works, about 1 mile west of the town of Mexia, 

Limestone County, Tex.

The species is recorded by Kline from the Paleocene 

of Clay County, Mississippi.

Porters Creek formation, Matthews Landing marl member. Alabama, Wilcox County (5).

Clay to Q formation, .Chalybeate limestone member. Alabama, 

Sumter County (9).

Saracenaria sublatifrons (Plummer) Kline 

Plate 7, figure 27

Cristellaria siiblatifrons Plummer, Texas Univ. Bull. 2644, p. 100,

pi. 7, fig. 6,1927. 

Saracenaria cf. sublatifrons Kline, Mississippi Geol. Survey Bull.

53, p. 31, pi. 3, fig. 1,1943.

Test elongate, smooth, tapering at both ends; peripheral margin bluntly angular; chambers few, passing from very slightly 

spiral to linear oblique in rapidly lengthening series, later chambers bluntly triangular but not keeled; sutures strongly oblique, 

smooth, distinct; aperture at apex of a long, narrow, slightly 

inflated septal face marked by a faint longitudinal furrow. 

Length up to 0.5 mm.—Plummer.

The types are from the Paleocene, clay pit, Mexir, 

Brick Works, about 1 mile west of the town of Mexia, 

Limestone County, Tex.

Wills Point formation, Mexia member. Texas, Limestone

County (36). 

Midway group. Texas, Caldwell County (44).

Saracenaria midwayensis Kline 

Plate 7, figures 28-30

Saracenaria midwayensis Kline, Mississippi Geol. Survey Bull.

53, p. 30, pi. 3, fig. 3,1943.

Test nearly triangular, periphery acute, apertural face elliptical ; closely coiled; chambers few, 4 to 5 usually visible, increasing rapidly in size; sutures distinct, depressed slightly or not 

at all, slightly curved; wall smooth, glossy; aperture peripheral, 

radiate. Average length 0.5 mm.—Kline.

The types came from the Paleocene Porters Creek 

clay, iy2 miles northwest of Montpelier, Clay County, 

Miss. Kline also reports that this species is widely distributed in Mississippi but is confined to the Porters 

Creek clay. No specimens were found in our collections

Genus VAGINTJLINA D'OrMgny, 1826

Vaginulina gracilis Plummer

Plate 8, figures 1-3

Vaginulina gracilis Plummer, Texas Univ. Bull. 2644, p. Ill, pi.

6, fig. 5,1927. 

Cushman, Cushman Lab. Foram. Research Contr., vol. 16

p. 61, pi. 9, fig. 27, 1940. 

Kline, Mississippi Geol. Survey Bull. 53, p. 32, pi. 3, fig. 7,

1943. 

Cooper, Jour. Paleontology, vol. 18, p. 351, pi. 55, figs. 5, 6

1944.

Test long, slender, slightly arcuate, gradually tapering toward the initial end, compressed; chambers numerous, short, 

smooth, compact, except the primordial chamber which is bulbous in megalospheric forms; sutures oblique on early portior 

of test to less oblique above, expressed outwardly by distinct 

and narrow ridges that extend around the apertural margin; 

wall moderately strong; aperture marginal, protruding, radiate. 

Length up to 2.5 mm.—Plummer.

The types are from the Paleocene, gully in field west 

of north-south stretch of road about 3 miles in a straight 

line northwest of Campbell or 4 miles by road from that 

town on highway to Neyland, Hunt County, -Tex.

This species is very common in the Paleocene of 

America and, together with V. midwayana Fox and 

Koss, makes an excellent index fossil for the Paleocene,

Clayton formation, Pine Barren member. Alabama, Butler

County (12). 

Chalybeate limestone member. Mississippi, Noxubee

County (16).

Porters Creek clay. Mississippi, Kemper County (17). 

Midway formation. Arkansas, Saline County (26) ; Pulaski

County (27) ; Clark County (29, 31). 

Midway group. Texas, Hunt County (41) ; Bastrop County

(43, 49) ; Travis County (45, 46) ; Van Zandt County (47) ;

Freestone County (50). 

Kincaid formation. Texas, Hunt County (37) ; Bastrop County

(38).
28 PALEOC'ENE FORAMINHFERA OF GULF COASTAL REGION

Vaginulina midwayana Fox and Ross 

Plate 8, figures 4-9

Vaginulina midwayana Fox and Ross, Jour. Paleontology, vol.

16, p. 669, 1942. 

Vaffinulina rotusta Plummer (not Chapman), Texas Univ. Bull.

2644, p. 112, pi. 6, fig. 4; pi. 13, fig. 3,1927. 

Cushman, Cushman Lab. Foram. Research Contr., vol. 16,

p. 61, pi. 10, figs. 1-4,1940.

Cushman and Renz, idem, vol. 18, p. 6, pi. 2, figs. 1, 2, 1942. 

Cushman and Todd, idem, vol. 18, p. 32, pi. 5, figs. 21, 22,

1942. 

Kline, Mississippi Geol. Survey Bull. 53, p. 34, pi. 3, fig. 8,

1943. 

Cooper, Jour. Paleontology, vol. 18, p. 351, pi. 55, figs. 16,

17, 1944.

Applin and Jordan, idem, vol. 19, p. 132 (list), 1945. 

Cushman and Todd, Cushman Lab. Foram. Research Contr.,

vol. 22, p. 54, pi. 9, figs. 15-19, 1946.

Test elongate, moderately broad, stout, somewhat compressed, 

tapering bluntly in megalospheric forms but very acutely in 

microspheric forms; chambers smooth, few, first two or three 

very slightly twisted followed by the usual straight linear 

series; sutures oblique, conspicuously marked by sharp, high 

ridges that on most specimens encircle the test, though a slight 

amount of discontinuity is frequently evident; aperture on extreme margin. Length up to 1.4 mm.—Plummer.

The types are from the Paleocene, clay pit, Mexia 

Brick Works, about 1 mile west of the town of Mexia, 

Limestone County, Tex.

This is an excellent index fossil for the Paleocene. 

Some specimens are difficult to distinguish from V. 

gracilis Plummer, but V. midwayanifl. has a greater 

amount of ornamentation and the costae extend out on 

the peripheral margins and become spinose. The general form is also shorter and broader.

Porters Creek formation, Matthews Landing marl member.

Alabama, Choctaw County (7). 

Clayton formation, Chalybeate limestone member. Alabama,

Sumter County (9).

Midway formation, lower part. Arkansas, Saline County (20). 

Midway formation, upper part. Arkansas, Pulaski County

(25). 

Midway formation. Arkansas, Saline County (26) ; Pulaski

County (27, 28) ; Clark County (30, 31). 

Midway group. Texas, Limestone County (39, 42) ; Navarro

County (40) ; Bastrop County (43) ; Caldwell County (44). 

Wills Point formation. Texas, Travis County (33) ; Caldwell

County (34). 

Mexia member. Texas, Limestone County (36).

Vaginulina longiforma (Plummer) Cushman 

Plate 8, figures 10-15

Gristellaria longiforma Plummer, Texas Univ. Bull. 2644, p.

102, pi. 13, fig. 4, 1927. 

Hemicristellaria longiforma Plummer, idem, Bull. 3201, pp. 54,

61, 62, 64 (lists), 1933.

VayinuUna longiforma Cushman, Cushman Lab. Foram. Research Contr., vol. 20, p. 38, pi. 6, figs. 11-13, 1944. 

Cushman and Todd, idem, vol. 22, p. 54, pi. 9, figs. 13, 14,

1946.

Cushman and Bermudez, idem, vol. 24, p. 70, pi. 11, fig. 4, 

1948.

Test elongate, stout, broad, compressed; few ear'y chambers 

more or less coiled, later chambers erect and somewhat oblique; 

sutures very strongly limbate on each side but appr.rent OD the 

margins as lines or less distinct ridges; aperture marginal, 

protruding, radiate. Length up to 1.7 mm.—Plummer.

The types are from the Paleocene, exposure along an 

east-west road about three-quarters of a mil0- north of 

New Hope between two creeks, Freestone Comity, Tex.

This species is very distinct from the preceding ones 

and should also make a good index fossil for the upper 

portion of the Paleocene.

Porters Creek formation, Matthews Landing marl member.

Alabama, Wilcox County (2, 4) ; Butler County (3). 

Naheola formation, Coal Bluff marl member. Alabama, Wilcox

County (8).

Midway formation, lower part. Arkansas, Saline County (21). 

Midway formation, upper part. Arkansas, Pulaski County

(25). 

Midway formation. Arkansas, Saline County (23) ; Pulaski

County (27). 

Wills Point formation. Texas, Travis County (33) ; Caldwell

County (34).

Vaginulina plumoides Plummer 

Plate 8, figures 16-18

Vaginulina plumoides Plummer, Texas Univ. Bull. 2644, p. 113,

pi. 6, fig 6, 1927. 

Cushman, Cushman Lab. Foram. Research Co^tr., vol 16,

p. 61, pi. 9, fig. 26,1940.

Cushman and Renz, idem, vol. 18, p. 6, pi. 1, fig. 6,1942. 

Kline, Mississippi Geol. Survey Bull. 53, p. 33, pi. 3, fig. 6,

1943. 

Cushman and Todd, Cushman Lab. Foram. Research Contr.,

vol. 22, p. 55, pi. 9, fig. 20,1944.

Test very thin, wing shaped, acuminate posteriorly and anteriorly, spreading rapidly upward; chambers very oblique and 

somewhat curved, ornamented by very fine delicate striae parallel to the direction of growth; aperture protruding. Length 

up to .8 mm.—Plummer.

The types are from the Paleocene, shallow ditch at 

road corner southeast of new Corsicana rerervoir on 

the road to Mildred, Navarro County, Tex.

This species, although not as common as smie of the 

others, is very distinctive and makes an excellent marker 

for the Paleocene. Typical specimens also o°.cur in the 

Paleocene of Trinidad.

Porters Creek formation, Matthews Landing mr.rl member. 

Alabama, Butler County (3)

Midway formation, upper part. Arkansas, Pulrski County 

(25).

Midway formation. Arkansas, Clark County (29).

Midway group. Texas, Bastrop County (43) ; Calc'well County 

(44) ; Travis County (45) ; Van Zandt County (47) ; Freestone County (50).

Wills Point formation. Texas, Caldwell County (34).

Vaginulina semilaevis Cushman and Bermudez 

Plate 21, figure 5

Vaginulina semilaevis Cushman and Bermudez, Crshman Lab. 

Foram. Research Contr., vol. 24, p. 70, pi. 11, f g. 6, 1948.

Test elongate, somewhat compressed, slender, increasing in width to the last-formed chamber, initial end


LAGENIDAE 29

acute with a slight spine, periphery slightly rounded; 

chambers of the early portion rather indistinct, later 

ones distinct and very slightly inflated, increasing 

rather evenly in size as added; sutures of the early portion indistinct, later ones distinct and slightly depressed, strongly oblique; wall of the early portion 

nearly smooth, later portion with longitudinal costae 

independent of the sutures; aperture terminal, radiate. 

Length of holotype 2.30 mm.; breadth 0.30 mm. The 

types are from the Paleocene Madruga formation, San 

Juan y Martinez, Pinar del Rio Province, Cuba.

This species differs from V. plumoides Plummer in 

the more slender, elongate form, higher chambers, and 

smooth early portion.

Vaginulina sp. 

Plate 8, figure 19

Vaginulina sp. A Cushman and Todd, Cushman Lab. Foram. 

Research Contr., vol. 22, p. 55, pi. 8, fig. 11, 1946.

Rare specimens of a compressed, faintly costate 

form may be new but are too few to warrant description.

Midway formation, upper part. Arkansas, Pulaski County (25).

Genus PALMULA Lea, 1833

Falmula tmdensis (Hantken) Cushman

Plate 7, figures 31, 32

Flabellina ludensis Hantken, K. Ungar. geol. Anstalt Mitth.,

vol. 4, p. 44, pi. 4, fig. 17,1875 (1881). 

Frondicularia ludensis Plummer, Texas Univ. Bull. 2644, p.

116, pi. 5, fig. 5, 1927.

Palmula cf. P. budensis Cushman, Cushman Lab. Foram. Research Contr., vol. 16, p. 62, pi. 10, figs, 5, 6,1940. 

Cushman and Todd, idem, vol. 18, p. 33, pi. 6, fig. 3, 1942. 

Kline, Mississippi Geol. Survey Bull. 53, p. 34, pi. 3, fig. 9,

1943.

Cushman and Todd, Cushman Lab. Foram. Research Contr., 

vol. 22, p. 55, pi. 9, figs. 21, 22, 1946.

Test strongly compressed, elongate oval; chambers 

distinct, earlier ones loosely coiled, later extending back 

011 both sides toward the base, narrow, increasing only 

slightly in size as added; sutures distinct, slightly depressed, very slightly curved; wall smooth; aperture 

terminal, radiate. Length 0.75 to 1.40 mm.

The types are from the upper Eocene of Hungary. 

Specimens referred to this species have been recorded 

from the Paleocene of Texas, Arkansas, Mississippi, 

and Alabama, but seem to be very rare.

The figures of "Frondicularia oldhami Plummer" 

(Texas Univ. Bull. 2644, p. 117, text figs. 12a-c, 1927) 

are similar to specimens of Palmula budensis, but no 

specimens are available.

Porters Creek formation, Matthews Landing marl member.

Alabama, Choctaw County (7). . . 

Clayton formation, Chalybeate limestone member. Alabama,

Sumter County (9). 

Midway formation, upper part. Arkansas, Pulaski County (25).

Midway formation. Arkansas, Saline County (26). 

Wills Point formation. Texas, Caldwell County (34).

Palmula delicatissima (Plummer) Cushman 

Plate 7, figures 33-35

Frondicularia delicatissima Plummer, Texas Univ. Bull. 2644, p.

120, pi. 5, fig. 4,1927. 

Palmula delicatissima Cushman, Cushman Lab. Foram. Research

Contr., vol. 16, p. 61, pi. 9, figs. 28, 29, 1940. 

Kline, Mississippi Geol. Survey Bull. 53, p. 35, pi. 3, fig. 12, 

1943.

Test very thin, broadly subovate, tapering rapidly toward the 

apertural extremity and bluntly rounded posteriorly; peripheral 

edges narrowly quadrate; early chambers Flabelline and irregularly coiled, later series typically sagittate; sutures delicate, 

thin, raised ridges from which branch a few wavy elevations 

especially near the apertural extremities of the sutures; shell 

wall coarsely punctate; aperture protruding. Length up to .95 

mm., average .5 mm.—Plummer.

The types are from the Paleocene, clay in section of 

Tehuacana Creek, 200 feet east of the Mexia-Wortham 

road bridge, Limestone County, Tex.

This species is apparently characteristic of the upper 

portion of the Paleocene of Texas and Alabama, and is 

recorded from the Porters Creek clay of Mississippi.

Clayton formation, Chalybeate limestone member. Alabama,

Sumter County (9). 

Wills Point formation, Mexia member. Texas, Limestone

County (36). 

Midway group. Texas, Limestone County (42).

Palmula rugosa (D'Orbigny) Cushman 

Plate 7, figures 36, 37

Flabellina rugosa D'Orbigny, Soc. geol. France Mem., 1st ser.,

vol. 4, p. 23, pi. 2, figs. 4, 5, 7, 1840. 

Reuss (in Geinitz), Grundriss der Verstein., p. 658, pi. 24,

fig. 23, 1845-46. 

D'Orbigny, Prodome de pa!6ontologie stratigraphique universelle des animaux mollusques et rayonnes, vol. 2, p.

281, No. 1379, 1850. 

Reuss, Akad. Wiss. Wien, Math.-naturwiss. Kl., Denkschr.,

vol. 7, pt. 1, p. 67,1854; idem, Sitzungsber., vol. 40, p. 215,

1860; idem, vol. 52, Abt. 1, p. 453,1865. 

Karrer, K. K. geol. Reichsanst. Jahrb., vol. 20, p. 176, 1870. 

Marsson, Naturw. Ver. Neu-Vorpommern u. Riigen Mitt.,

Jahrg. 10, p. 140, 1878. 

Goes, K. svenska vetensk. akad. Handl., vol. 15, pt. 4, No. 2,

pi. 2, fig. 4, 1889. 

Beissel, Preuss. geol. Landesanstalt Abb.., new ser., vol. 3,

p. 47, pi. 9, figs. 20-24; pi. 16, figs. 30, 31, 1891. 

Egger, K. bayer. Akad. Wiss., Math.-naturh. Abt., Abb.., Kl.

2, vol. 21, p. 108, pi. 10, figs. 5, 6; pi. 13, figs. 1, 2, 1899;

Naturwiss. Ver. Passau Ber., p. 30, pi. 1, fig. 8,1907. 

Cushman, Cushman Lab. Foram. Research Contr., vol. 3,

p. 189, 1927. 

Sandidge, Jour. Paleontology, vol. 6, p. 279, pi. 42, fig. 22,

1982. 

Brotzen, Zeitschr. Deutschen Palastina-Vereins, Jahrg.,

p. 45, 1934. 

Cushman, Cushman Lab. Foram. Research Contr., vol. 11, p.

83, pi. 13, figs. 1-6,1935.

915259—51-
30 PALEOCENE FGRAMIN&FERA OF GULF COASTAL REGION

Frondicularia rugosa Plummer, Texas Univ. Bull. 2644, p. 118,

pi. 5, fig. 1, 1927. 

Palmula rugosa Cushman, Foraminifera, Ed. 3, Key, pi. 20, fig.

8, 1940; Cushman Lab. Foram Research Contr., vol. 16,

p. 62, pi. 9, fig. 30, 1940; idem, vol. 20, p. 8, pi. 2, fig. 6,

1944. 

Kline, Mississippi Geol. Survey Bull. 53, p. 36, pi. 3, fig.

11, 1943. 

Cushman and Deaderick, Jour. Paleontology, vol. 18, p.

335, pi. 52, fig. 7, 1944. 

Flabellina, interpunctata Von der Marck, Naturh. Ver. preuss.

Rheinland Verh., vol. 15, p. 53, pi. 1, fig. 5, 1858. 

Reuss, Akad. Wiss. Wien, Math.-naturwiss. Kl., Sitzungsber.,

vol. 40, p. 216, pi. 9, fig. 1, 1860. 

Heron-Alien and Barland, Royal Micr. Soc. Jour., p. 422,

pi. 8, fig. 5, 1910. 

Franke, Naturh. Ver. preuss. Rheinlande u. Westfalens

Verh., 69 Jahrg., vol. 59, 1912, p. 277 (1913). 

Chapman, W. Australia Geol. Survey Bull. 72, p. 34, pi.

10, fig. 91, 1917. 

Franke, Greifswald Univ., Geol.-paleont. Inst., Abh., vol.

6, p. 64, pi. 5, fig. 13, 1925; Preuss. geol. Landesanstalt

Abh., new ser., vol. Ill, p. 92, pi. 8, fig. 17, 1928. 

Cushman, Cushman Lab. Foram. Research Contr., vol. 6,

p. 30, pi. 4, figs. 16, 17, 1930; Jour. Paleontology, vol. 5,

p. 307, pi. 35, fig. 9,1931. 

Plummer, Texas Univ. Bull. 3101, p. 163, pi. 12, figs. 1-3,

1931.

Cushman, Jour. Paleontology, vol. 6, p. 336, 1932. 

Sandidge, Am. Midland Naturalist, vol. 13, p. 194, pi. 19,

figs. 12-14, 1932; Jour. Paleontology, vol. 6, p. 279, pi.

42, fig. 21, 1932. 

Frondicularia, projecta Carsey, Texas Univ. Bull. 2612, p. 41, pi.

6, fig. 5, 1926. 

Flabellina projecta Plummer, idem, Bull. 3101, p. 165, pi. 12,

figs. 5-8, 1931.

Frondioularia 'baudoiiiniana Cushman (not D'Orbigny), Cushman Lab. Foram. Research Contr., vol. 2, pt. 1, p. 21, pi. 

3, fig. 5. 1926; Jour. Paleontology, vol. 1, p. 155, pi. 24, fig. 

13, 1927.

Test sagittate to rhomboid or even broadly elliptical 

in outline, much compressed, sides nearly flat, periphery 

truncate; early chambers coiled, especially in the microspheric form, later chambers chevron-shaped, narrow 

and of uniform width, extending back 011 both sifles, 

often at first nearly enclosing the earlier coiled chambers; sutures raised and sharp, later chambers with a 

loop at the apical end; wall between the raised sutures 

with a series of small, raised papillae; apertural end 

projecting, with a slight neck. Length up to 1.20 mm., 

breadth 1.00 mm.

The types of this species are from the Upper Cretaceous Craie blanche of the Paris Basin. It is a very 

common species in the Upper Cretaceous of America. 

It occurs especially in the lower portion of the Paleocene of Texas, Alabama, and Mississippi. Some of the 

specimens seem worn and may have been reworked from 

Upper Cretaceous beds.

Clayton formation, Chalybeate limestone member. Mississippi,

Noxubee County (16). 

Midway formation. Arkansas, Clark County (31).

Palmula primitiva Cushman var. paleocenica C'lshman

Plate 7, figure 38

Palmula primitiva Cushman var. paleocenica Cushman, Cushman Lab. Foram. Research Contr., vol. 24, p. 43, pi. 8, fig. 7, 

1948.

Palmula cf. P. primitiva Cushman, Cushman Lab Foram. Research Contr., vol. 16, p. 62, pi. 9, fig. 33, 1940. 

Kline, Mississippi Geol. Survey Bull. 53, p. 35, pi. 3, fig. 10, 

1943.

Variety differing from the typical form in the shorter 

broader shape and the thicker wall.

The types are from the Paleocene Ostrea pulaskensis 

bed, old abandoned roadway south of Sucarnoochee 

Creek, about %0 mile upstream* from crossing of U. S. 

Highway 80, % mile southwest of Livings! on, Sumter 

County, Ala. (9).

This is probably the same as the form figured by 

Kline from the Paleocene of Mississippi, where it is 

recorded from the Porters Creek clay and the Clayton 

formation.

Genus FRONDICTTIABIA Defrance, 1824

Frondicularia midwayensis Cushman

Plate 9, figures 4, 5

Frondicularia midwayensis Cushman, Cushman Lab. Forarn.

Research Contr., vol. 16, p. 63, pi. 10, figs. 34, 35, 1940. 

Frondicularia sp. Cushman, idem, vol. 20, p. 39, pi. 6, fig. 14,1944.

Test large, much compressed, irregularly rhomboid 

in front view, with the greatest breadth toward its 

apertural end, initial end with a slight spine; chambers 

numerous, distinct, of uniform shape, increasing very 

gradually in height as added, proloculum globular; 

sutures distinct, slightly raised above the general surface; wall of the proloculum with a few longitudinal 

costae, the surface in the adult finely but distinctly 

papillate; aperture terminal, radiate, slightly projecting. Length 1.75 to 2.50 mm., breadth 0.50 to 0.65 mm.

The types are from the Paleocene, U. S. Highway 80, 

south of Sucarnoochee Creek, i/2 mile south vest of Livingston, Sumter County, Ala. (9).

Porters Creek formation, Matthews Landing marl member.

Alabama, Butler County (3). 

Clayton formation, Chalybeate limestone member. Alabama,

Sumter County (9). 

Midway group. Texas, Bastrop County (43).

Frondicularia naheolensis Cushman and T^dd 

Plate 8, figures 24, 25

Frondicularia naheolensis Cushman and Todd, CTshman Lab.

Foram. Research Contr., vol. 18, p. 33, pi. 6, figs. 5, 6,1942;

vol. 22, p. 55, pi. 9, fig. 23,1946.

Frondicularia arcMaciana D'Orbigny var. strigillata Plummer

(not Bagg), Texas Univ. Bull. 2644, p. 114, pi. 5, fig 2, 1927.

Frondicularia cf. F. frankei Kline, Mississippi Geol. Survey Bull.

53, p. 37, pi. 3, fig. 14, 1943.

Test elongate, compressed, the periphery truncate, 

flattened or slightly concave, the sides angled, some-
LAGENIDAE 31

times almost keeled, the sides nearly parallel for most of 

their length, initial end with a short stout spine, apertural end tapering to a blunt point; chambers few, 

strongly overlapping at the sides; sutures distinct, 

somewhat limbate, little if at all raised, strongly oblique ; wall ornamented with numerous very fine longitudinal costae, independent of the sutures, the proloculum in the megalospheric form with a strong central 

costa; aperture terminal, radiate. Length of holotype 

0.87 mm. including the spine, breadth 0.25 mm., thickness 0.13 mm.

The types are from the Paleocene Naheola formation, 

greensand bed, upper fossiliferous horizon, Naheola 

Landing, Tombigbee River, Ala.

Specimens of this species are mostly from the upper 

part of the Paleocene but seem to be well distributed in 

the area covered by our collections. It should make a 

good index fossil for this part of the section.

Porters Creek formation, Matthews Landing marl member.

Alabama, Wilcox County (4) ; Choctaw County (7). 

Midway formation, upper part. Arkansas, Pulaski County (25). 

Midway formation. Arkansas, Saline County (26). 

Midway group. Texas, Limestone County (39) ; Caldwell

County (44). 

Wills Point formation. Texas, Caldwell County (34).

Frondicularia cf. F. goldfussi Reuss 

Plate 8, figures 20-23

Frondicularia goldfussi Reuss Plummer, Texas Univ. Bull. 2644,

p. 115, pi. 5, fig. 3,1927. 

Cushman, Cushman Lab. Foram. Research Contr., vol. 16,

p. 62, pi. 10, fig. 29.1940.

Nodosarina (Frondicularia) goldfussi Kline, Mississippi Geol. 

Survey Bull. 53, p. 28, pi. 2, fig. 7,1943.

Rare specimens from the Paleocene of Texas and Alabama may be compared with F. goldfussi with some 

question. They represent only the early stages and no 

adult specimens were found to confirm the identification. The specimen figured by Mrs. Plummer and 

copied here shows a slightly greater development.

The species is recorded by Kline as fairly abundant 

from three localities in the Porters Creek clay in Clay 

County, Miss.

Midway formation. Arkansas, Clark County (30). 

Wills Point formation, Mexia member. Texas, Limestone 

County (36).

Frondicularia cf. F. frankei Cushman 

Plate 9, figures 1-3

A few specimens from the Paleocene of Alabama 

have been recorded and figured under this name (Cushman, Cushman Lab. Foram. Research Contr., vol. 16, 

p. 63, pi. 10, figs. 36, 37, 1940). They may possibly be 

reworked specimens from Upper Cretaceous beds. 

Similar specimens are recorded and figured from the 

Paleocene of Mississippi (Kline, Mississippi Geol. Survey Bull. 53, p. 37, pi. 3, fig. 14,1943).

Clayton formation, Chalybeate limestone member. Alabama,

Sumter County (9).

Midway formation. Arkansas, Clark County (30). 

Frondicularia sp. A 

Plate 9, figure 6

Frondicularia sp A. Cushman and Todd, Cushman Lab. Foram. 

Research Contr., vol. 22, p. 55, pi. 8, fig. 11,1946.

The peculiar species figured occurs at a few localities 

in the Paleocene of Arkansas and Alabama, but no 

adult specimens were found to give the full specific 

characters. 

Clayton formation, Pine Barren member. Alabama, Butler

County (12). 

Midway formation, upper part. Arkansas, Pulaski County

(25). 

Midway formation. Arkansas, Saline County (26).

Genus IAGENA Walker and Jacob, 1798

Lagena cf. L. laevis (Montagu) Williamson

Plate 9, figures 8, 9

Lagena cf. L. laevis (Montagu) Williamson, Cushman and Todd,

Cushman Lab. Foram Research Contr., vol. 18, p. 34, pi.

6, fig. 4, 1942. 

Kline, Mississippi Geol. Survey Bull. 53, p. 38, pi. 4, fig. 4, 1943. ' 

Cushman and Todd, Cushman Lab. Foram. Research Contr.,

vol. 22, p. 56, pi. 9, fig 26,1946.

Specimens similar to those figured are rather common in some of the Paleocene material. It is recorded 

from the Paleocene of Arkansas, the Naheola formation of Alabama, and the Porters Creek clay and Clayton formation of Mississippi.

Porters Creek clay. Mississippi, Kemper County (17).

Porters Creek formation, Matthews Landing marl member. 

Alabama, Wilcox County (1, 2, 4) ; Butler County (3) ; Choctaw County (7).

Midway formation, lower part. Arkansas, Saline County (20).

Midway formation, upper part. Arkansas, Pulaski County 

(25).

Midway formation. Arkansas, Saline County (26).

Lagena sulcata (Walker and Jacob) Parker and Jones var.

semiinterrupta W. Berry

Plate 9, figures 10,11

Lagena 'sulcata (Walker and Jacob) Parker and Jones var.

semiinterrupta W. Berry, in Berry and Kelley, U. S Nat.

Mus. Proc., vol. 76, art. 19, p. 5, pi. 3* fig. 19, 1929. 

Cushman, Tennessee Div. Geology Bull. 41, p. 37, pi. 5, figs.

9-11, 1931 

Jennings, Bull. Am. Paleontology, vol. 23, no. 78, p. 23, pi. 2,

fig. 23, 1936. 

Berquist, Mississippi Geol. Survey Bull. 49, p. 52, pi. 5, figs.

21, 22, 1942.

Kline, idem, Bull. 53, p. 39, pi. 7, figs. 15, 16, 1943. 

Cushman, U. S. Geol. Survey Prof. Paper 206, p. 95, pi. 39,

figs. 18-21, 1946.

Specimens from several stations may be referred to 

this variety with some question. They are very small 

but have the general ornamentation. The species is 

known mainly in Upper Cretaceous beds of Navarro 

age but is recorded from the Eocene Jackson group and
32 PALEOC'ENE FORAMINUFERA OP GULF COASTAL REGION

the Paleocene Porters Creek clay and Clayton formation of Mississippi.

Clayton formation. Alabama, Barbour County (10). 

Wills Point formation. Texas, Caldwell County (34).

Mexia member. Texas, Limestone County (36). 

Midway group. Texas, Caldwell County (44).

Lagena cf. L. acuticosta Reuss 

Plate 9, figure 12

Very rare specimens from the Paleocene are not entirely typical of L. acuticosta but are compared with 

that species until more specimens are available.

Naheola formation, Coal Bluff marl member. Alabama, Wilcox

County (8). 

Clayton formation, Chalybeate limestone member. Alabama,

Sumter County (9).

Pine Barren member. Alabama, Butler County (12). 

Midway formation, lower part. Arkansas, Saline County (20,

21).

Midway formation. Arkansas, Pulaski County (27). 

Midway group. Texas, Caldwell County (44). 

Wills Point formation. Texas, Caldwell County (34).

Lagena cf. L. costata (Williamson) Reuss 

Plate 9, figure' 13

Lagena sp. A Cushman and Todd, Cushman Lab. Foram. Research Contr., vol. 22, p. 56, pi. 9, fig. 27, 1946.

Very rare specimens from a few Paleocene localities may be 

compared with L. costata with some question. They are rather 

finely costate and have a slender neck when well preserved. 

Porters Creek clay. Tennessee, Hardeman County (13). 

Porters Creek formation, Matthews Landing marl member.

Alabama, Butler County (3). 

Clayton formation, Pine Barren member. Alabama, Butler

County (12).

Midway formation, upper part. Arkansas, Pulaski County (25). 

Midway group. Texas, Caldwell County (44).

"Lagena cf. L. primigera H. B. Brady" 

Plate 9, figure 14

This species is recorded by Kline (Mississippi Geol. 

Survey Bull. 53, p. 38, pi. 7, fig. 21, 1943) from the 

uppermost part of the Clayton formation of Clay 

County, Miss. A single specimen was all that was 

found. Kline's figure is copied on our plate.

Family POLYMORPHINIDAE

"V Genus GTTTTTTLINA D'Orbigny, 1839 

Guttulina problema D'Orbigny 

Plate 9, figures 15-18

Specimens referred to this widely ranging species 

occur at a number of stations but show a very considerable amount of variation. It is also recorded from 

Mississippi as fairly common in the Clayton formation 

and extremely rare in the Porters Creek clay (Kline, 

Mississippi Geol. Survey Bull. 53, p. 39, pi. 4, fig. 2, 

1943).

Porters Creek clay. Tennessee, Hardeman County (13, 14). 

Porters Creek formation, Matthews Landing^ marl member.

Alabama, Wilcox County (2, 4, 6) ; Butler County (3) ;

Choctaw County (7). 

Naheola formation. Coal Bluff marl member. Alabama, Wilcox

County (8). 

Clayton formation. Alabama, Barbour County (10).

Pine Barren member. Alabama, Wilcox County (11). 

Midway formation, lower part. Arkansas, Saline County (18,

19A, 20, 21).

Midway formation, upper part. Arkansas, PulaskJ County (25). 

Midway formation. Arkansas, Saline County (26) ; Pulaski

County (27); Clark County (31).

Midway group. Texas, Van Zandt County (47).

Guttulina hantkeni Cushman and Ozava 

Plate 9, figures 19-21

Guttulina hantkeni Cushman and Ozawa, U. S. Nat. Mus. Proc.,

vol. 77, art. 6, p. 33, pi. 5, figs. 4-6,1980. 

Cushman and Dusenbury, Cushman Lab. Foram. Research

Contr., vol. 10, p. 60, pi. 8, fig. 5,1934. 

Cushman and Todd, idem, vol. 18, p. 34, pi. 6, figs. 11, 12,

1942. 

Curran, Am. Assoc. Petroleum Geologists Bull., vol. 27, p.

1381 (list), 1943. 

Cushman and Todd, Cushman Lab. Foram. Research Contr.,

vol. 22, p. 56, pi. 10, fig. 2, 1946. 

Polymorphina acuta Hantken (not D'Orbigny), Jahrb. K. Ungar.

geol. Anstalt Mitth., vol. 4, p. 60, pi. 8, fig. 4 (acuminata on

explanation of plate), 1875 (1881).

Test oval, botryoidal, more or less rounded at the 

base, acute at the apertural end, greatest breadth above 

the middle; chambers ovate, only slightly embracing, 

arranged in a contraclockwise, quinqueloculine series, 

each succeeding chamber removed farther from the 

base; sutures distinct, strongly depressed; wall smooth; 

aperture radiate, projecting. Length 0.60 to 1.20 

mm., breadth 0.35 to 0.75 mm., thickness 0.30 to 0.60 

mm.

The types are from the Eocene of Hungary, and the 

species has a wide range in the Eocene. Numerous 

specimens occur in our Paleocene material and show a 

considerable amount of variation.

Porters Creek clay. Tennessee, Hardeman County (13). 

Porters Creek formation, Matthews Landing marl member.

Alabama, Butler County (3) ; Wilcox County (6) ; Choctaw

County (7). 

Clayton formation, Pine Barren member. Alal ama, Wilcox

County (11). 

Midway formation, upper part. Arkansas, Pulaski County

(24,25). 

Midway formation. Arkansas, Saline County (26) ; Pulaski

County (27) ; Clark County (29). 

Midway group. Texas, Caldwell County (44).

Guttulina wilcoxensis Cushman and Ponton 

Plate 9, figure 22

Guttulina, wilooxensis Cushman and Ponton, Cushman Lab.

Foram. Research Contr., vol. 8, p. 60, pi. 8, figs. 1, 2,

1932. 

Cushman and Garrett, idem, vol. 15, p. 80, pi. 14, figs. 8, 9,

1989. 

Cushman, Am. Jour. Sci., vol. 242, p. 9, pi. 3, fig. 8, 1944;

Cushman Lab. Foram Research Contr., vol. 20, p. 23;

p. 39, pi. 6, fig. 18,1944.
POLYMORPHINIDAE 33

Test elongate, fusiform, composed of comparatively 

few chambers, distinct, inflated, elongate, usually two 

or three times as long as wide, each chamber added in 

the adult only reaching to about one-half or two-thirds 

of the distance to the base of the preceding chamber, 

sutures distinct, depressed; wall smooth, finely perforate ; aperture radiate, terminal. Length 0.60 to 0.75 

mm., diameter 0.20 to 0.25 mm.

The types are from the Eocene of Wilcox age from 

railroad cut, 1 mile north of Ozark, Ala.

The species has been recorded from a number of 

localities of Wilcox age and also from the Coal Bluff 

member of the Naheola formation of the Paleocene. 

In the Paleocene it is largely confined to the Naheola 

formation, with one occurrence in the Clay ton formation.

Porters Creek clay. Tennessee, Hardeman County (14). 

Porters Creek formation, Matthews Landing inarl member.

Alabama, Wilcox County (1, 2, 4). 

Naheola formation, Coal Bluff marl member. Alabama, Wilcox

County (8). 

Clayton formation, Pine Barren member. Alabama, Wilcox

County (11).

Genus GLOBULINA D'Orbigny, 1839

Globulina gibba D'OrMgny

Plate 9, figures 26-28

(For earlier references, see U. S. Geol. Survey Prof. Paper 181, 

p. 25,1935.)

So many forms have been placed under this specific 

name that it is difficult to try to give a complete synonymy. In the Paleocene material specimens seemingly 

referable to this species have occurred at numerous 

localities. These show a considerable degree of variation. Fistulose forms occur but are rather rare.

Porters Creek clay. Tennessee, Hardeman County (13, 14).

Mississippi, Kemper County (17). 

Porters Creek formation, Matthews Landing marl member.

Alabama, Wilcox County (1, 2, 4-6) ; Butler County (3) ;

Choctaw County (7).

Naheola formation, Coal Bluff marl member. Alabama, Wilcox County (8). 

Clayton formation, Pine Barren member, Alabama, Butler

County (12). 

Chalybeate limestone member. Mississippi, Noxubee

County (16). 

Midway formation, lower part. Arkansas, Saline County (19A,

20-22); Pulaski County (23). 

Midway formation, upper part. Arkansas, Pulaski County (24,

25). 

Midway formation. Arkansas, Saline County (26) ; Pulaski

County (27); Clark County (30). 

Midway group. Texas, Hunt County (41) ; Bastrop County (43,

49) ; Caldwell County (44) ; Travis County (45) ; Van Zandt

County (47).

Wills Point formation. Texas, Williamson County (35). 

Kincaid formation. Texas, Hunt County (37).

Globulina rotundata (Bornemann) Cushman and Ozawa 

Plate 9, figures 29-33

(For earlier references, see U. S. Geol. Survey Prof. Paper 181,

p. 27, 1935.) 

Parr, Royal Soc. West Australia Jour., vol. 24, p. 80, pi. 2,

fig. 2,1937-8.

Bergquist, Mississippi Geol. Survey Bull. 49, p. 56, 1942. 

Crespin, Commonwealth of Australia, Min. Res. Survey,

Bull. 9 (Pal. Ser. No. 4), p. 80 (list), 1943. 

Cushman, Cushman Lab. Foram. Research Contr., vol. 20,

p. 40, pi. 6, figs. 20-22,1944. 

Cushman and Todd, idem, vol. 21, p. 14, 1945; vol. 22, p. 56,

pi. 10, fig. 4, 1946. 

Cushman, Cushman Lab. Forain. Research Special Pub. 16,

p. 19, pi. 4, figs. 14, 15, 1946.

Test ellipsoidal or ovoid to cylindrical, rounded at 

the base, slightly produced at the apertural end; chambers rounded, almost as long as broad, arranged in a 

nearly triserial series, each succeeding chamber removed 

much farther from the base, rarely becoming almost 

uniserial in the last chamber; sutures only slightly 

depressed, generally distinct; wall smooth, thick, often 

with fistulose tubes; aperture radiate. Length 0.45 to 

0.90 mm., breadth 0.30 to 0.60 mm., thickness 0.25 to 

0.55 mm.

According to the records this species appears to have 

a long range and wide distribution. Specimens referable to it occurred at a number of localities in our 

Paleocene material and show a considerable amount of 

variation. Occasional specimens show a fistulose 

development.

Porters Creek clay. Tennessee, Hardeman County (14). 

Porters Creek formation, Matthews Landing marl member.

Alabama, Wilcox County (2, 6) ; Butler County (3). 

Naheola formation, Coal Bluff marl member. Alabama, Wilcox

County (8).

Midway formation, upper part. Arkansas, Pulaski County (25). 

Midway formation. Arkansas, Saline County (26) ; Pulaski

County (28) ; Clark County (29). 

Midway group. Texas, Hunt County (41) ; Caldwell County

(44). 

Eincaid formation. Texas, Hunt County (37).

Globulina cf. G. minuta (Roemer) Reuss 

Plate 9, figure 34

Very rare specimens from the Paleocene of Arkansas ' 

and Texas are compared with G. minuta, as not enough 

specimens are available to make the specific identification certain.

Midway formation, upper part. Arkansas, Pulaski County (25). 

Midway group. Texas, Caldwell County (44).

Genus PYRULINA ITOrbigny, 1839

Pyruliha cf. P. eylihdroides (Eoemer) Cushman and Ozawa 

Plate 9, figures 23, 24

Pyrulina cylindroid.es (Roemer) Kline, Mississippi Geol. Survey Bull. 53, p. 40, pi. 7, fig. 5,1943.

Cushman, Cushman Lab. Foram. Research Contr., vol. 2(1, 

p. 40, pi. 6; fig. 23, 1944.

Cushman and Todd, idem, vol. 22, p. 57, pi. 10, fig. 6, 1944.
34 PALEOCENE FORAMINIFEEA OF GULF COASTAL REGION

This species is recorded with some question as to 

its presence in the Paleocene of Mississippi, Alabama, 

and Arkansas because specimens here compared with P. 

cylindroides are very rare and not entirely typical. 

Very rare specimens of a similar form were also found 

in the Paleocene of Texas.

Porters Creek clay. Tennessee, Hardeman County (14).

Porters Creek formation, Matthews Landing marl member. Alabama, Wilcox County (1, 2).

Naheola formation, Coal Bluff marl member. Alabama, Wilcox 

County (8).

Midway formation, lower part. Arkansas, Saline County (20).

Midway formation, upper part. Arkansas, Pulaski County (25).

Midway formation. Arkansas, Pulaski County (27).

Midway group. Texas, Caldwell County (34).

Pyrulina extensa (Gushmau) Cushman and Ozawa 

Plate 9, figure 25

Polymorphina extensa Cushman, U. S. Nat. Mus. Bull. 104, pt.

4, p. 156, pi. 41, figs. 7, 8,1923. 

Pyrulina extensa Cushman and Ozawa, idem, Proc., vol. 77, art.

6, p. 53, pi. 12, fig. 5, 1930.

The only Paleocene record for this species is from 

Arkansas (Cushman and Todd, Cushman Lab. Foram. 

Research Contr., vol. 22, p. 57, pi. 10, fig. 7,1946). One 

other specimen from Arkansas is the only additional 

material found in the numerous samples examined. 

Midway formation, upper part. Arkansas, Pulaski County

(24,25).

Genus PSETTDOPOLYMORPHINA Cushman and Ozawa, 1928

Pseudopolymorphina wilcoxensis Cushman and Ponton

Plate 10, figure 1

Pseudopolymorphina wilcoxensis Cushman and Ponton, Cushman Lab. Foram. Kesearch Contr., vol. 8, p. 61, pi. 8, figs. 

5, 6,1932.

Cushman and Garrett, idem, vol. 15, p. 81, pi. 14, fig. 11,1939.

Cushman, idem, vol. 20, p. 23, pi. 4, fig. 19; p. 41, pi. 7, fig 2, 

1944.

Test somewhat compressed, only slightly longer than 

broad, periphery broadly rounded, apertural end 

slightly produced, early chambers irregularly spiral, 

later ones becoming biserial; chambers distinct, very 

slightly if at all depressed in the early stages, slightly 

so in the adult; wall thick, opaque, ornamented with 

definite, short, slightly raised, somewhat elongate 

papillae arranged generally lengthwise of the test but 

irregularly placed; aperture fairly large, radiate, terminal, slightly projecting. Length 0.60 to 0.75 mm., 

breadth 0.55 to 0.60 mm., thickness 0.30 to 0.35 mm.

The types are from the Eocene Wilcox group, from 

railroad cut, 1 mile north of Ozark, Ala.

Most of the records for this species are from beds of 

the Wilcox group but it occurs in the Paleocene, in the 

Coal Bluff marl member of the Naheola formation of 

Alabama (8).

Pseudopolymorphina sp. A

PseudopolymorpMna sp. A Cushman and Todd, Cushman Lab. 

Foram. Kesearch Contr., vol. 22, p. 57, pi. 10, fig. 8, 1946.

The only specimen found in the Paleocene is from 

Arkansas and is noted here for the record until more 

material can be found and a specific identification established. 

Midway formation, upper part. Arkansas, Pulaskf County (25).

Genus SIGMOMORPHINA Cushman and Ozawa, 1928

Sigmomorphina semitecta (Reuss) Cushman and Ozawa var.

terquemiana (Fornasini) Cushman and 02 aw a

Plate 10, figures 2, 3

Polymorphina amygdaloides Reuss var. terquemiana Fornasini, 

Accad. sci. 1st. Bologna Mem., 5th ser., vol. 9, 1900-1902, 

p. 72, fig. 25 (in text), 1902. 

Sifftnomorphina semitecta (Reuss) Cushman and Ozawa var.

terqttemiana (Fornasini) Cushman and Ozawa, U. S.

Nat. Mus. Proc., vol. 77, art. 6, p. 129, pi. 33, figs. 4, 5;

pi. 34, figs. 2, 3; pi. 35, fig. 1; 1980. 

Cushman, Tennessee Div. Geology Bull. 41, p. 42, pi. 6,

figs. 12a-c, 1931; U. S. Geol. Survey Prof. Paper 181, p.

28, 1935. 

Howe, Louisiana Dept. Cons. Geol. Bull. 14, p. 55, pi. 7,

figs. 11,12, 1939. 

Cushman and Todd, Cushman Lab. Foram. Research Contr.,

vol. 18, p. 35, pi. 6, figs. 16,17, 1942.

Cushman and Applin, idem, vol. 19, p. 37, pi. 7, fig. 23, 1943. 

Cushman, idem, vol. 20, p. 41, pi. 7, fig. 3, 1944. 

Cushman and Todd, idem, vol. 21, p. 90, pi. 14, fig. 11,1945;

vol. 22, p. 57, 1946. 

Cushman, U. S. Geol. Survey Prof. Paper 206, p. 98, pi. 41.

fig. 13, 1946.

Test compressed, oval to elongate, lanceolate, tapering toward the aperture; chambers elongate, arranged 

in a clockwise sigmoid series, all extending down to 

the base, but not involute; sutures scarcely depressed, 

distinct; wall smooth; aperture radiate. Length 0.30 

to 0.55 mm., breadth 0.20 to 0.35 mm., thickness 0.10 to 

0.20 mm.

Specimens referable to this variety are fairly common in the Paleocene. From the records the variety 

has a wide range in the Upper Cretaceous and Eocene.

Porters Creek clay. Tennessee, Hardeman County (13, 14). 

Porters Creek, formation, Matthews Landing nnrl member.

Alabama, Wilcox County (2, 5, 6) ; Butler County (3) ;

Choctaw County (7). 

Naheola formation. Coal Bluff marl member. Alabama, Wilcox

County (8). 

Clayton formation, Pine Barren member. Alabr.ma, Wilcox

County (11) ; Butler County (12). 

Midway formation, lower part. Arkansas, Saline County (19,

20, 21). 

Midway formation, upper part. Arkansas, Pulaski County (25).

Sigmomorphina wilcoxensis Cushman and Ponton 

Plate 10, figure 4

SiffmomorpMna wilcoxensis Cushman and Ponton, Cushman 

Lab. Foram. Research Contr., vol. 8, p. 61, pi. 8, fig. 7, 

1932.
POLYMORPHINIDAE 35

Toulmin, Jour. Paleontology, vol. 15, p. 595, pi. 80, fig.

15,1941. 

Cushman and Todd, Cushman Lab. Foram. Research Contr.,

vol. 18, p. 35, pi. 6, fig. 15, 1942. 

Cushman, idem, vol. 20, p. 41, pi. 7, fig. 4, 1944. 

Cushman and Todd, idem, vol. 22, p. 57, pi. 10, fig. 9, 1946.

Test elongate, in the type specimen about twice as 

long as broad, much compressed, periphery rounded; 

chambers in the early portion in an elongate spiral, later 

ones becoming somewhat sigmoid, successive chambers 

in the adult being removed rapidly from the base, inflated; sutures distinct, later ones slightly depressed, 

earlier ones flush with the surface; wall smooth, finely 

perforated; aperture comparatively large, terminal, 

radiate. Length 0.90 mm., breadth 0.45 mm., thickness 

0.20 mm.

The types are from the Eocene Wilcox group from 

railroad cut, 1 mile north of Ozark, Ala.

Specimens that seem referable to this species occurred 

at a number of localities in the Paleocene of Alabama, 

Tennessee, and Arkansas.

Porters Creek clay. Tennessee, Hardeman County (13,14). 

Porters Creek formation, Matthews Landing marl member.

Alabama, Wilcox County (1, 4) ; Butler County (3) ; Choctaw

County (7). 

Naheola formation. Coal Bluff marl member. Alabama,

Wilcox County (8). 

Midway formation, upper part. Arkansas, Pulaski County (25).

Sigmomorphiua cf. S. williamsoni (Terquem) Cushman and Ozawa 

Plate 10, figure 5

Sifftnomorphina cf. williamsoni (Terquem) Cushman, Cushman Lab. Foram. Research Contr., vol. 20, p. 41, pi. 7, 

fig. 5, 1944.

Specimens resembling /S. willimnsoni are recorded 

from the Paleocene of Alabama, and a single specimen 

occurred in a sample from Tennessee, making the only 

Paleocene records for this form.

Naheola formation, Coal Bluff marl member. Alabama, Wilcox

County (8). 

Porters Creek clay. Tennessee, Hardeman County (13).

Sigmomorphiua soldadoensis Cushman and Renz 

Plate 10, figure 6

Sigmomorphina soldadoensis Cushman and Renz, Cushman 

Lab. Foram. Research Contr. vol. 18, p. 7, pi. 2, fig. 5, 1942.

Test about twice as long as broad, the greatest 

breadth above the middle, strongly compressed, two 

biserial chambers in the adult making up more than 

half the test, earlier ones sigmoid; chambers of the last 

part slightly inflated; sutures distinct, but only slightly 

depressed in the adult portion; wall smooth; aperture 

terminal, radiate. Length of holotype 0.60 mm., 

breadOi 0.30 mm., thickness 0.15 mm.

The types are from the Paleocene Soldado Rock formation, Soldado Rock, Trinidad, B. W. I. It was not 

found in any of our other material.

Genus POLYMORPHINA D'Orbigny, 1826

Polymorphina cushmani Plummer

Plate 10, figures 8-12

Polymorphina cushmani Plummer, Texas Univ. Bull. 2644, p.

125, pi. 6, fig. 9 ; pi. 15, fig. 1,1927. 

Cushman and Ozawa, U. S. Nat. Mus. Proc., vol. 77, art. 6,

p. 117, pi. 30, fig. 8, 1930. 

Kline, Mississippi Geol. Survey Bull. 53, p. 41, pi. 4, fig. 8,

1943. 

Cushman and Todd, Cushman Lab. Foram. Research Contr.,

vol. 22, p. 57, pi. 10, fig. 10, 1946.

Test broadly ovoid, strongly compressed; peripheral margin 

narrowly rounded and somewhat lobate; sutures marked by 

faint depressions toward the margins but marked by irregularly disposed and broken elevations down the central axis on 

each side of the test; aperture extended, radiate. Length up 

to 1 mm.—Plummer.

The types are from the Paleocene, about 5^ miles 

clue south and very slightly west of Littig where the 

440-foot contour cuts a northeast-southwest road 

(Bastrop quadrangle), Bastrop County, Tex.

Typical specimens occur in the Paleocene of Texas, 

Arkansas and Mississippi. 

Pojrters Creek formation, Matthews Landing marl member.

Alabama, Butler County (3). 

Midway formation, upper part. Arkansas, Pulaski County

(25).

Midway formation. Arkansas, Saline County (26). 

Midway group. Texas, Bastrop County (43) ; Caldwell County

(44) ; Van Zandt County (47, 48).

Polymorphina frondea (Cushman) Cushman 

Plate 10, figure 13

Bolivina frondea Cushman, U. S. Geol. Survey Prof. Paper 

129-F, p. 126, pi. 29, fig. 3, 1922; Prof. Paper 133, p. 20, 

1923.

Polymorphina frondea Cushman, Cushman Lab. Foram. Research Contr., vol. 5, p. 41, 1929.

This species has been recorded by Kline (Mississippi 

Geol. Survey Bull. 53, p. 41, pi. 4, fig. 7, 1943) from 

the Porters Creek clay and Clay ton formation of Mississippi. In the Chalybeate limestone member of th«s 

Clay ton formation of Alabama (9), a single specimen 

recorded as Polymorphina sp. (Cushman, Cushmain 

Lab. Foram. Research Contr., vol. 16, p. 64, pi. 11, fig. 

8, 1940) seems to be the same as that recorded from 

Mississippi.

Polymorphina sp.

Plate 10, figure 7

Rare specimens from the Paleocene of Alabama evidently represent the early stages of a smooth form related to P. cushmani Plummer, but no adult specimens 

were found. They seem identical with the form figured by Kline (Mississippi Geol. Survey Bull. 53, p. 41, 

pi. 4, fig. 6, 1943) from the Clayton formation of 

Mississippi as "Sigmomorphina sp." 

Clayton. formation, Pine Barren member. Alabama, Butler

County (12).
36 PALEOCENE FORAMINIFERA OF GUI/F COASTAL REGION

Genus POIYMOEPHINEILA Cushman and Hanzawa, 1936 

Polymorphinella cf. P. elongata Toulmin

Plate 10, figures 14-16

Rare specimens from the Paleocene of Texas and 

Alabama seem related to P. elongata, described from 

the Eocene Wilcox group of Alabama.

Porters Creek formation, Matthews Landing marl member.

Alabama, Butler County (3). 

Wills Point formation. Texas, Caldwell County (34).

Polymorphinella sp. 

Plate 10, figure 17

Polymorphinella sp. Cushman, Cushman Lab. Foram. Research 

Contr., vol. 20, p. 41, pi. 7, fig. 6, 1944.

The only record for this species is from the Paleocene 

Coal Bluff marl member of the Naheola formation (8).

Genus BUIIOPORA ftuenstedt, 1856

Bullopora laevis (Sollas) Wickenden

Plate 10, figure 18

Webbina laevis Sollas, Geol. Mag., dec. 2, vol. 4, p. 103, pi. 6,

figs. 1-3,1877. 

Vitreiveblina laevis Chapman, idem, dec. 3, vol. 8, p. 53, pi. 2,

fig. 4, 1892; Annals and Mag. Nat. History, ser. 6, vol.

18, p. 332, text fig. 3, 1896; Royal Micr. Soc. Jour., p. 585,

pi. 12, fig. 12, 1896. 

Bagg, U. S. Geol. Survey Bull. 88, p. 36, pi. 2, figs. 4a, b,

1898. 

. Chapman, Annals and Mag. Nat. History, ser. 7, vol. 3,

p. 314, 1899. 

Weller, New Jersey Geol. Survey, Paleontology, vol. 4, p. 205,

pi. 1, figs. 40, 41, 1907. 

Chapman, Western Australia Geol. Survey Bull. 72, p. 37,

pi. 11, fig. 101,1917.

Plummer, Texas Univ. Bull. 2644, p. 128, pi. 8, fig. 3, 1927. 

Bullopora laems (Sollas) Wickenden, Jour. Paleontology, vol.

6, p. 206, pi. 29, figs. 6-8, 1932. 

Cushman, Cushman Lab. Foram. Research Special Pub. 5,

pi. 22, fig. 24, 1933; idem, Contr., vol. 16, p. 64, pi. 11, fig.

12, 1940. 

Tappan, Jour. Paleontology, vol. 14, p. 115, pi. 18, fig. 6,

1940; idem, vol. 17, p. 507, pi. 81, figs. 11, 12, 1943. 

Cushman and Todd, Cushman Lab. Foram. Research Contr.,

vol. 19, p. 63, pi. 11, fig. 8, 1943. 

Kline, Mississippi Geol. Survey Bull. 53, p. 43, pi. 4, fig. 13,

1943. 

Lozo, Am. Assoc. Petroleum Geologists Bull., vol. 27, p. 1066

(list), 1943; Am. Midland Naturalist, vol. 31, p. 560,

pi. 3, fig. 2, 1944. 

Cushman, Cushman Lab. Foram. Research Contr., vol. 20,

p. 9, pi. 2, fig. 33, 1944; U. S. Geol. Survey Prof. Paper

206, p. 98, pi. 42, figs. 1-4, 1946.

Test attached, consisting of a CrlobuUna-like stage 

followed by a series of globular, oval, fusiform, or 

irregularly shaped chambers, generally in a single series joined by stolon-like connections, or branching into 

two or more series, wall smooth.

The globular shape of the chambers is retained even 

where the test is attached to an elongate object such as 

a Nodosaria. The species evidently has a fairly long 

range in the Upper Cretaceous and Tertiary.

Porters Creek formation, Matthews Landing marl member.

Alabama, Butler County (3). 

Clayton formation, Chalybeate limestone member. Alabama,

Sumter County (9). Mississippi, Noxubee Courty (16). 

Midway group. Texas, Caldwell County (44).

"Bullopora laevis (Sollas) var. hispida Kliue"

Bullopora laevis (Sollas) var. hispida Kline (not B. chapmani 

(Plummer) var. hispida Kline), Mississippi Geol. Survey 

Bull. 53, p. 43, pi. 4, fig. 16,1943.

This hispid variety described from the Porters Creek 

clay 2,1/2 miles north of Pheba, Miss., needs a new name 

as the name hispida has already been used.

Bullopora chapmani (Plummer) Cushman 

Plate 10, figures 19-22

Vitrewebbma chapmani Plummer, Texas Univ. Bull. 2644, p.

128, pi. 8, fig. 2,1927. 

Bullopora chapmani Cushman, Cushman Lab. Foranv Research

Contr., vol. 16, p. 64, pi. 11, figs. 10,11, 1940. 

Kline, Mississippi Geol. Survey Bull. 53, p. 42, pi. 4, fig. 11,

1943. 

Cushman and Todd, Cushman Lab. Foram. Research

Contr., vol. 22, p. 58, pi. 10, figs. 13, 14, 1946 

Cushman and Bermudez, idem, vol. 24, p. 71, pi. 11, fig. 8,

1948.

Test adherent, composed of strongly inflated, perfectly smooth, 

elongate chambers joined by narrow slender tubes, and edged by 

a slight flange that is so thin and merges so well irto the shell 

on which it is adherent, that it is hardly evident unless the test 

has been broken away from its support. Averag? length of 

single chamber .7 nim.; average width .3 mm.—Plummer.

The types are from the Paleocene, Alamo Brick Co. 

clay pit southeast of San Antonio on north £*ide of the 

road leading southeast from the city and abont 14 mile 

from the area broken up into city blocks, Bexr.r County, 

Tex.

Rare specimens with elongate chambers oc~ur in the 

Paleocene material. It is not as common as B. laevis.

Clayton formation, Chalybeate limestone member. Alabama,

Sumter County (9).

Midway formation, upper part. Arkansas, Pulaski C?unty (25> 

Midway group. Texas, Freestone County (50).

Bullopora chapmani (Plummer) Cushman var. hispida, Kline 

Plate 10, figure 23

Bullopora chapmani (Plummer) var. hispida Kline, Mississippi 

Geol. Survey Bull. 53, p. 43, pi. 4, fig. 12,1943.

Variety differs from the typical form of the species in having 

a finely hispid surface. The variety resembles the species very 

closely in size and shape, but may readily be identified by its hispid test. No intermediate forms between species and variety 

have been noted.—Kline.

The variety was described from the Porters Creek 

clay, 21/0 miles north of Pheba, Miss.

Genus RAMULIN A Rupert Jones, 1875

Ramulina cf. K. aculeata (D'OrMgny) Wright

Plate 10, figures 24-26

Rare specimens occur in our Paleocene material. JR. 

aculeata has been recorded from widely separated
NONIONIDAE 37

areas and from both the Cretaceous and Tertiary. It 

has been previously recorded from the Paleocene (Gushman, Cushman. Lab. Forani. Research Contr., vol. 16, 

p. 64, pi. 11, figs. 13, 14, 1940 and Cushman and Todd, 

idem, vol. 18, p. 35, pi. 6, fig. 24, 1942; vol. 22, p. 58, pi. 

10, fig. 12, 1946). A smooth specimen evidently belonging to this genus was figured by Mrs. Plummer 

from the Paleocene of Texas (Texas Univ. Bull. 2644, 

p. 127, pi. 8, fig. 7, 1927). The early stages were not 

found at any of the localities.

Porters Creek formation, Matthews Landing marl member.

Alabama, Butler County (3) ; Choctaw County (7). 

Clayton formation, Chalybeate limestone member. Alabama,

Sumter County (9). Mississippi, Noxubee County (16). 

Midway formation, upper part. Arkansas, Pulaski County (25). 

Midway formation. Arkansas, Pulaski County (27). Caldwell County (44). 

Wills Point formation. Texas, Travis County (33).

Family NONIONIDAE

Genus NONIONELLA Cushman, 1926

Nonionella insecta (Sclrwager) Cushman and Ponton

Plate 11, figure 1 

Anomalina insecta Schwager, Palaeontographica, vol. 30, Pal.

Theil, p. 128, pi. 28 (5), figs. 1, 2, 1883. 

Nonionella insecta Cushman and Ponton, Cushman Lab. Forana.

Research Contr., vol. 8, p. 65, pi. 8, figs. 13, 14, 1932. 

Glaessner, Problems of Paleontology, Moscow Univ., vols.

2-3, p. 368, 1937. 

Cushman, U. S. Geol. Survey Prof. Paper 191, p. 29, pi. 8,

fig. 1, 1939. 

Toulmin, Jour. Paleontology, vol. 15, p. 597, pi. 80, fig. 22,

1941. '

Cushman, Cushman Lab. Forana. Research Contr., vol. 20, 

p. 42, 1944.

The types of this species are from the middle Eocene 

of northern Africa. It has been recorded from the 

Eocene Wilcox group of Alabama, in the Tuscahoma 

sand, the Hatchetigbee formation, and the Salt Mountain limestone. Bare specimens assigned to this species 

occurred in the Paleocene material from Alabama and 

Arkansas.

Porters Creek formation, Matthews Landing marl member.

Alabama, Butler County (3). 

Naheola formation, Coal Bluff marl member. Alabama, Wilcox

County (8). 

Midway formation, lower part. Arkansas, Saline County (21).

Nonionella soldadoensis Cushman and Renz

Plate 11, figures 2-4 

Non.ion.eUn soldadoensis Cushman and Renz, Cushman Lab.

Forana. Research Contr., vol. 18, p. 7, pi. 2, fig. 7, 1942. 

Nonionina turgida Plummer (not Willianason) (part), Texas 

Univ. Bull. 2644, p. 159, pi. 12, fig. 6,1927.

Test somewhat longer than broad, strongly compressed, periphery rounded, ventral side with the basal 

portion of the umbilical area covered, in peripheral 

view with the sides nearly parallel; chambers distinct, 

slightly inflated on the ventral side, all visible on the 

dorsal side, on the ventral side with the last-formed 

chamber extended into a lobe over the umbilical region;

915259—51———4

sutures distinct, slightly depressed; wall smooth; aperture extending from the periphery ventrally under the 

lobe of the last-formed chamber. Length of holotype 

0.40 mm., breadth 0.25 mm., thickness 0.08 mm.

The types are from the Paleocene Soldado Rock formation, Soldaclo Rock, Trinidad, B. W. I.

Midway formation, lower part. Arkansas, Pulaski County (23). 

Midway formation. Arkansas, Saline County (26) ; Clark 

County (29).

Nonionella sp. 

Plate 11, figures 5, 6

Nonionina turgida Plummer (not Williamson) (part), Texas 

Univ. Bull. 2644, p. 159, pi. 12, fig. 7,1927.

A single specimen from the Paleocene of Alabama 

here figured seems very close to the figure given by Mrs. 

Plummer of a specimen from the Paleocene of Texas. 

More specimens are needed to give this a specific identification.

Porters Creek formation, Matthews Landing marl member. Alabama, Butler County (3).

Family HETEROHELICIDAE 

Genus BOLIVINOPSIS Yakovlev, 1891

Bolivinopsis sp.

Two very small specimens, evidently the young stages 

but definitely belonging in this genus, were found in the, 

Paleocene Naheola formation on Alabama Highway 96, 

9.2 miles northeast of Kimbrough, Wilcox County, Ala. 

(5).

Genus GtJMBELINA Egger, 1899

Giimbelina midwayensis Cushman

Plate 11, figures 7, 8

Giimbelina midwayensis Cushman, Cushman Lab. Foram. Research Contr., vol. 16, p. 65, pi. 11, fig. 15, 1940. 

Cushman and Todd, idem, vol. 22, p. 58, pi. 10, fig. 15, 1946

Test small, compressed, usually twice as long as broad, 

rapidly tapering, with the greatest breadth formed by 

the last pair of chambers, periphery rounded throughout, lobulate; chambers with breadth and height about 

equal, slightly overlapping, inflated, increasing rapidly 

in height as added; sutures distinct, depressed, very 

slightly curved; wall finely spinose; aperture high, 

arched, with distinct lateral flanges. Length 0.18 to 

0.22 mm., breadth 0.10 to 0.12 mm., thickness 0.15 mm.

The types are from the Paleocene, U. S. Highway 80, 

south of Sucarnoochee Creek, y2 mile southwest of Livingston, Sumter County, Ala.

This seems to be an index fossil for the Paleocene, occurring in our material from Alabama, Arkansas, anc1 

Texas.

Porters Creek formation, Matthews Landing marl member. Alabama, Wilcox County (5).

Clayton formation, Chalybeate limestone member. Alabama, 

Sumter County (9). 

Pine Barren member. Alabama, Butler County (12).
38 PALEOCENE FORAMIMtFERA OF GULF COASTAL REGION

Midway formation, lower part. Arkansas, Saline County (20). 

Midway formation, upper part. Arkansas, Pulaski County (24,

25).

Midway formation. Arkansas, Clark County (30). 

Midway group. Texas, Caldwell County (44). 

Wills Point formation. Texas, Caldwell County (34).

Gihubelina trinitatensis Cushman and Renz

Plate 11, figure 9

CriimbeUna trinitatensis Cushman and Renz, Cushman Lab. 

Forarn. Research Contr., vol. 18, p. 8, pi. 2, fig. 8, 1942.

Test slightly longer than broad, moderately compressed, rapidly tapering, greatest breadth formed by 

the last-formed pair of chambers, periphery rounded, 

lobulate; chambers with breadth and height about equal, 

the last-formed pair in the adult usually much larger 

than the remainder of the test; sutures distinct, depressed, straight, nearly at right angles to the elongate 

axis; wall smooth or slightly hispid; aperture high, 

arched. Length of holotype 0.27 mm., breadth 0.20 

mm., thickness 0.15 mm.

The types are from the Paleocene Soldado Rock formation, Soldado Rock, Trinidad, B. W. I. It has not 

been recorded elsewhere.

Giimbelina morsel Zline 

Plate 11, figure 10

Gumbelina morsei Kline, Mississippi Geol. Survey Bull. 53, p. 

44, pi. 7, fig. 12, 1943.

Test small, about twice as long as broad, regularly tapering, 

with greatest breadth at apertural end, periphery rounded and 

lobulate; chambers with breadth greater than height, inflated, 

increasing rapidly in size, especially in breadth; sutures distinct, depressed; wall finely but distinctly spinose; aperture 

high, arched, with distinct lateral flanges.

This minute species resembles Giimbelina midwayensis Cushman, from the Midway of Alabama, but may readily be distinguished from that species by the proportionately greater 

breadth of the chambers.

The species is common in the Porters Creek and transitional 

beds, but is not known from typical Clayton beds.—Kline.

The types are from the Paleocene, test hole M 49, 2% 

miles north of Pheba, Clay County, Miss.

This species seems very similar to G. trinitatensis 

Cusliman and Renz.

Genus RECTOGUMBELINA Cushman, 1932

Rectogiimbelina alabamensis Cushman

Plate 11, figure 11

Rectogiimbelina alabamensis Cushman, Cushman Lab. Foram.

Research Contr., vol. 16, p. 65, pi. 11, fig. 16, 1940. 

Cooper, Jour. Paleontology, vol. 18, p. 351, pi. 55, figs. 3, 4, 

1944.

Test minute, elongate, early portion definitely biserial, 6rumbelina-lik&, later portion uniserial; chambers of the early portion not much inflated, later ones 

distinctly so; sutures not depressed in the early portion,

later much depressed; wall very finely hispid throughout ; aperture small, nearly terminal, with a very slightly raised edge above the upper margin. Length 0.17 

to 0.22 mm., diameter 0.06 to 0.08 mm.

The types are from the Paleocene, U. S. Highway 80, 

south of Sucarnoochee Creek, % mile southwest of Livingston, Sumter County, Ala. (9). The only other 

record is that of Cooper, from the Paleocene Porters 

Creek clay of Illinois.

Genus PLANOGLOBULINA Cushman, 1927 

"Planoglobulina acervulinoides (Egger) Cushman"

This species is recorded by Kline (Mississippi Geol. 

Survey Bull. 53, p. 45, pi. 7, fig. 1,1943) from the Paleocene Clayton formation of Mississippi. As it is represented by a single specimen and is typically a Cretaceous species it may have been redeposited from Upper 

Cretaceous beds.

Genus EOUVIGERINA Cushman, 1926

Eouvigerina excavata Cushman

Plate 11, figure 12

Eouvigerina excavata Cushman, Cushman Lab. Foram. Research Contr., vol. 16, p. 66, pi. 11, fig. 18, 1940. 

Cushman and Todd, idem, vol. 18, p. 35, pi. 6, figs. 20, 21.

1942. 

Kline, Mississippi Geol. Survey Bull. 53, p. 45, pi. 4, fig. 19,

1943. 

Cushman, Am. Jour. Sci., vol. 242, p. 10, pi 1, fig 18, 1944:

Cushman Lab. Foram. Research Contr., vol. 20, p. 42, pi. 7,

fig. 7, 1944. 

Cushman and Todd, idem, vol. 22, p. 58, pi. 10, fig. 16, 1946.

Test small, mostly biserial, in the adult quadrangular 

in end view, tapering, greatest breadth formed by the 

last two chambers, initial end rounded; chambers very 

distinct, the broader faces deeply excavated, the angles 

of the chamber raised into narrow plate-like projections; sutures distinct, strongly raised; wall smooth, 

finely perforate; aperture terminal, rounded, with a 

distinct neck and lip. Length 0.18 to 0.25 mm., diameter 

0.08 to 0.10 mm.

The types are from the Paleocene, U. S. Highway 80, 

south of Sucarnoochee Creek, i/2 mile southwest of Livingston, Sumter County, Ala.

Except for a single record from the Bashi marl member of the Hatchetigbee formation of tr<s Wilcox 

group (Eocene), this is recorded only from the Paleocene, where it occurs at a number of localities in Alabama, Arkansas, Mississippi, and Texas.

Porters Creek clay. Tennessee, Hardeman County (14). 

Porters Creek formation, Matthews Landing marl member. Alabama, Wilcox County (5, 6) ; Choctaw County (7). 

Naheola formation, Coal Bluff marl member. Alabama, Wilcox

County (8).

Clayton formation, Chalybeate limestone member. Alabama, 

Sumter County (9).

Pine Barren member. Wilcox County (11) ; Butler County (12).
BULIMINIDAE

Midway formation, lower part. Arkansas, Saline County (20-

22). 

Midway formation, upper part. Arkansas, Pulaski County (24.

25).

Midway formation. Arkansas, Clark County (30). 

Midway group, Texas, Bastrop County (43); Caldwell County

(44).

Wills Point formation. Texas, Caldwell County (34). 

Mexia member. Texas, Limestone County (36).

Genus FSEUDOUVIGERINA Cushman, 1927

Fseudouvigerina naheolensis Cushman and Todd

Plate 11, figures 13-15

Pseudoiivigerina naheolensis Cushman and Todd, Cushman Lab. 

Foram. Research Contr., vol. 18, p. 36, pi. 6, figs. 18, 19,

1942. 

Kline, Mississippi Geol. Survey Bull. 53, p. 46, pi. 7, fig. 2,

1943. 

Cushman, Am. Jour. Sci., vol. 242, p. 11, pi. 1, figs. 15, 16,

1944.

Cushman and Todd, Cushman Lab. Foram. Research Contr., 

vol. 22, p. 59, pi. 10, fig. 17, 1946.

Test small, broadly fusiform in front view, in end 

view triangular with the sides flattened or somewhat 

concave, the periphery angled and slightly carinate, or 

slightly truncate; chambers distinct, very slightly inflated in the last whorl; sutures distinct, slightly depressed in the later portion; wall distinctly perforate, 

generally smooth; aperture circular, at the end of a 

short but distinct cylindrical neck with a slight phialine 

lip. Length 0.30 mm., breadth 0.15 mm.

The types are from the Paleocene Naheola formation, 

greensand bed, upper fossiliferous horizon, Naheola 

Landing, Tombigbee River, Ala.

Except for a single record from the Eocene Bashi 

marl member of the Hatchetigbee formation of the Wilcox group of Alabama, the species seems limited to the 

Paleocene.

Porters Creek clay. Tennessee, Hardeman County (13, 14).

Porters Creek formation, Matthews Landing marl member. 

Alabama, Butler County (3) ; Wilcox County (4, 5) ; Choctaw County (7).

Clayton formation, Pine Barren member. Alabama, Wilcox 

County (11).

Midway formation, lower part. Arkansas, Saline County (20- 

22).

Midway. formation, upper part. Arkansas, Pulaski County 

(25).

Midway formation. Arkansas, Saline County (26).

Genus SIFHOGENERINOIDES Cushman, 1927

Siphogenerinoides eleganta (Flummer) Cushman

Plate 11, figures 16-19

Siphogenerina, eleganta, Plummer, Texas Univ. Bull. 2644, p. 126,

pi. 8, fig. 1,1927.

Bifarina eleganta Plummer, idem, Bull. 3201, pp. 54, 62, 1933. 

Siphogenerinoides eleganta, Cushman, Cushman Lab. Foram. 

Research Contr., vol. 16, p. 66, pi. 11, fig. 17,1940.

Cushman and Renz, idem, vol. 18, p. 8,1942.

Kline, Mississippi Geol. Survey Bull. 53, p. 46, pi. 4, fig. 18,

1943.

Cooper, Jour. Paleontology, vol. 18, p. 351, pi. 54, fig. 14,1944. 

Cushman, Cushman Lab. Foram. Research Contr., vol. 20,

p. 42, pi. 7, fig. 8, 1944.

Cushman and Todd, idem, vol. 22, p. 59, pi. 10, fig. 18, 1946. 

Cushman and Bermudez, idem, vol. 24, p. 71, pi. 11, fig. 9,

1948.

Test elongate; early chambers biserial merging into a succession of alternately oblique chambers that very rarely reach a 

Nodosarian development; very earliest portion of test marked 

by indistinct and irregularly developed longitudinal striations 

and spinulose projections that disappear rapidly upward; mature chambers very smooth and coarsely punctate; sutures 

sharply depressed; aperture terminal, elliptical, bounded by a 

short, flaring rim and connected to earlier apertures by an inner 

tube. Length up to 0.9 mm.; average 0.5 mm.—Plummer.

The types are from the Paleocene, clay pit, Mexia 

Brick Works, about 1 mile west of the town of Mexia, 

Limestone County, Tex.

This seems to be an excellent index fossil for the 

Paleocene. It has occurred at numerous stations in our 

Paleocene material from Alabama, Mississippi, Texas, 

and Arkansas. It has been recorded by Kline from the 

Porters Creek clay and the Clayton formation of Mississippi and by Cooper from the Porters Creek clay of 

Illinois. The species also occurs in the Paleocene Soldado rock formation of Trinidad.

Naheola formation, Coal Bluff marl member. Alabama, Wilcox County (8). 

Clayton formation, Chalybeate limestone member. Mississippi,

Noxubee County (16).

Porters Creek clay. Mississippi, Kemper County (17). 

Midway formation, lower part. Arkansas, Saline County (19A,

21). 

Midway formation, upper part. Arkansas, Pulaski County (24,

25). 

Midway formation. Arkansas, Pulaski County (27) ; Clark

County (29, 30). 

Midway group. Texas, Limestone County (39, 42) ; Hunt

County (41) ; Caldwell County (44) ; Van Zandt County (47). 

Wills Point formation. Texas, Travis County (33) ; Caldwell

County (34).

Mexia member. Texas, Limestone County (36).

Family BULIMINIDAE

Genus BULIMINELLA Cushman, 1911

Buliminella elegantissima (D'Orbigny) Cushman

Plate 11, figure 20

(For references, see U. S. Geol. Survey Prof. Paper 210-D, p. 67, 

pi. 17, figs. 106-12, 1947.)

Numerous specimens from the Paleocene of Arkansas 

and Alabama give the first records for this species earlier than the Wilcox group (Eocene). The Paleocene 

specimens are very small but otherwise seem to be typical and are common at all localities.

Clayton formation, Pine Barren member. Alabama, Wilcox

County (11) ; Butler County (12). 

Midway formation, lower part. Arkansas, Saline County

(18,19).
40 PALEOC'ENE FORAMINIFERA OF GULF COASTAL REGION

Genus ROBERTINA D'Orbigny, 1846

Robertina wilcoxensis Cushman and Ponton

Plate 11, figure 21

Robert ina wilcoxensis Cushman and Ponton, Cushman Lab.

Foram. Research Contr., vol. 8, p. 66, pi. 8, fig. 19, 1932. 

Cushman and Parker, idem, vol. 12, p. 96, pi. 16, fig. 13, 1936. 

Cushman and Garrett, idem, vol. 15, p. 82, pi. 14, fig. 16,1939. 

Cushman and Todd, idem, vol. 18, p. 36, pi. 6, fig. 22, 23,1942. 

Cushman, Am. Jour. Sci., vol. 242, p. 11, pi. 1, fig. 17, 1944;

Cushman Lab. Foram. Research Contr., vol. 20, p. 42, pi. 7,

fig. 9, 1944. 

Cushman and Parker, U. S. Geol. Survey Prof. Paper 210-D,

p. 71, pi. IS, fig. 1, 1947.

Test elongate, about twice as long as broad, fusiform, 

greatest breadth slightly above the middle; chambers 

in an elongate spiral, those in each whorl alternating, 

very slightly inflated; sutures distinct, not depressed; 

wall smooth, finely perforate; aperture on the ventral 

side formed by an elongate narrow opening deeply 

entering the apertural face. Length 0.35 mm., breadth 

0.18 mm., thickness 0.15 mm.

The types are from the Eocene Wilcox group from 

railroad cut, 1 mile north of Ozark, Ala.

Occasional specimens from the Paleocene Naheola 

formation of Alabama are probably to be placed under 

this species, recorded previously only from the Eocene 

of Alabama. These make the oldest records for this 

genus.

Porters Creek formation, Matthews Landing marl member. 

Alabama, Wilcox County (5, 6) ; Choctaw County (7).

Naheola formation. Coal Bluff marl member. Alabama, Wilcox County (/S).

Genus BULIMINA D'Orbigny, 1826

Bulimina cacumenata Cushman and Parker

Plate 11, figures 22, 23

Bulimina cacumenata Cushman and Parker, Cushman Lab.

Foram. Research Contr., vol. 12, p. 40, pi. 7, fig. 3, 1936. 

Cushman, idem, vol. 16, p. 67, pi. 11, fig. 20, 1940. 

Cushman and Todd, idem, vol. 18, p. 37, 1942. 

Kline, Mississippi Geol. Survey Bull. 53, p. 47, pi. 7, fig. 8,

1943. 

Cushman and Todd, Cushman Lab. Foram. Research Contr.,

vol. 22, p. 59, pi. 10, fig. 20, 1946. 

Cushman and Parker, U. S. Geol. Survey Prof. Paper 210-D,

p. 92, pi. 21, fig. 27, 1947.

Test small, somewhat fusiform, greatest width 

slightly above the middle, gradually tapering to a long, 

subacute point; chambers numerous, 6 to 7 whorls, those 

of the last whorl somewhat inflated; arranged in a 

slightly twisted series, those of adjacent series meeting 

in a zigzag line; sutures distinct in the upper part, 

obscure in the lower part of the test, very slightly depressed ; wall, except for the last whorl and occasionally 

for the next to the last, covered with irregular, low, 

closely set costae, last whorl smooth, coarsely perforate;

aperture loop-shaped, with a slight lip. Length 0.20 to 

0.23 mm., diameter 0.10 to 0.11 mm. 

• The types are from the Paleocene, 3 miles above 

bridge over Cedar Creek on Austin-Bed Rock road, 

Bastrop County, Tex.

The species is an excellent index fossil for the Paleocene as it is usually rather common. It occurs in our 

Paleocene material from Alabama, Texas, and Arkansas, and has been recorded by Kline from the Paleocene 

of Mississippi.

Porters Creek formation, Matthews Landing marl member. Alabama, Wilcox County (1, 2, 4-6) ; Butler County (3) ; Choctaw County (7).

Clayton formation. Alabama, Barbour County (10).

Pine Barren member. Alabama, Wilcox Courty (11) ; Butler County (12).

Midway formation, lower part. Arkansas, Saline County (19- 

22); Pulaski County (23).

Midway formation, upper part. Arkansas, PulasM County (24, 

25).

Midway formation. Arkansas, Saline County (26) ; Clark 

County (29, 30).

Midway group. Texas, Bastrop County (43) ; Ca.ldwell County 

(44).

Wills Point formation. Texas, Caldwell County (34).

Bulimina arkadelphiana Cushman and Parker var. midwayensis 

Cushman and Parker

Plate 11, figures 25, 26

Bulimina, arJcadelpMana Cushman and Parker var. midwayensis 

Cushman and Parker, Cushman Lab. Foram. Research 

Contr., vol. 12, p. 42, pi. 7, figs. 9, 10, 1936. 

Kline, Mississippi Geol. Survey Bull. 53, p. 47, pi. 7, fig. 9,

1943. 

Cushman and Parker, U. S. Geol. Survey Prof. Paper 210-1',

p. 92, pi. 21, figs. 24, 25, 1947.

Bulimina aculeata Plummer (not D'Orbigny), Texas Univ. Bull. 

2644, p. 73, pi. 4, fig. 3, 1927.

Test small, about 1% times as long as brord, tapering, 

usually with a well-defined basal spine; chambers distinct, undercut at base, giving a "collared" effect, about 

5 whorls, last-formed chambers inflated, sutures distinct, 

depressed; wall of all but the last-formed whorl, covered with sharp, fine spines, usually extending from the 

lower edges of the chambers, last-formed wl orl smooth, 

finely perforate; aperture loop-shaped with a well-defined lip. Length 0.26 to 0.38 mm., diameter 0.16 to 

0.25 mm.

The types are from the Paleocene, road cnt south of 

Reservoir 3^ miles southeast of Corsicana, Tex.

Specimens are not common but occurred at several 

stations in Arkansas and Texas. It has be^n recorded 

by Kline from the Porters Creek clay of Mississippi.

Midway formation, upper part. Arkansas, Pulaski County (24). 

Midway formation. Arkansas, Clark County (30). 

Midway group. Texas, Limestone County (39). 

Wills Point formation, Mexia member. Texas', Limestone 

County (36).
BULIMESTIDAE 41

Bulimina kugleri Cushman and Renz 

Plate 11, figure 24

Bulimina Jcugleri Oushman and Renz, Cushman Lab. Foram. Research Contr., vol. 18, p. 9, pi. 2, fig. 9, 1942. 

Cushman and Parker, U. S. Geol. Survey Prof. Paper 210- 

D, p. 93, pi. 30, fig. 13, 1947.

Test elongate, fusiform, about twice as long as broad, 

greatest breadth at about the middle; chambers distinct, 

slightly inflated, elongate; sutures distinct, very slightly depressed; 1 wall smooth; aperture a high, arched, 

slightly curved opening at the base of the inner margin 

of the last-formed chamber. Length of holotype 0.50 

mm., diameter 0.23 mm.

The types are from the Paleocene Soldado Rock formation, Soldado Rock, Trinidad, B. W. I. It has not 

been recorded elsewhere.

Submenus DESINOBULIMINA Cushman and Parker, 1940 

Bulimina (Desiuobulimina) quadrata Plummer

Plate 11, figures 27-30

Bnlimina (Ellipsobulimina) quadrata Plummer, Texas Univ.

Bull. 2644, p. 72, pi. 4, figs. 4, 5, 1927. 

Bulimina quadrata Jennings, Bull. Am. Paleontology, vol. 23, no.

78, p. 30, pi. 3, fig. 19, 1936. 

Albritton and Phleger, Jour. Paleontology, vol. 11, p. 352,

1937. 

Bulimina (Desinobulimina) quadrata Cushman, Cushman Lab.

Foram. Research Contr., vol. 16, p. 67, pi. 11, fig. 21, 1940. 

Kline, Mississippi Geol. Survey Bull. 53, p. 48, pi. 4, fig. 20,

1943. 

Applin and Jordan, Jour. Paleontology, vol. 19, p. 131 (list),

1945. 

Cushman and Todd, Cushman Lab. Foram. Research Contr.,

vol. 22, p. 60, pi. 10, fig. 19, 1946. 

Cushman and Parker, U. S. Geol. Survey Prof. Paper 210-D,

p. 128, pi. 29, figs. 18, 19, 1947.

Test of megalospheric form (Form A) almost cylindrical, 

stout, increasing in diameter only very slightly from the broad 

blunt initial end toward the broadly rounded oral extremity; 

microspheric form (Form B) pointed aborally through a succession of small chambers that follow the proloculum to the 

later mature chambers that comprise a test identical in shape 

with that of the much more frequent megalospheric form; chambers smooth, very little inflated, broad, and short; sutures as 

sharp lines in early part of test and faintly depressed above; 

wall thin; aperture a large vertical slit on the inner side of the 

last chamber and connected with all previous apertures by an 

inner tube that traverses the entire length of the shell. Length 

up to 0.65 mm. in megalospheric form, average 0.5 mm.; up 

to 0.8 mm. in microspheric form.—Plummer.

The types are from the Paleocene, clay pit, Mexia 

Brick Works, about 1 mile west of the town of Mexia, 

Limestone County, Tex.

The species is often very common in some of the 

Paleocene samples examined. It is recorded from the 

Upper Cretaceous but these probably are to be included 

under B. kickapooensis Cole.

Clayton formation, Chalybeate limestone member. Alabama, 

Sumter County (9).

Midway formation, upper part. Arkansas, Pulaski County (25). 

Midway formation. Arkansas, Saline County (26) ; Clark

County (30). 

Midway group. Texas, Limestone County (39, 42) ; Caldwell

County (44).

Wills Point formation. Texas, Caldwell County (34). 

Mexia member. Texas, Limestone County (36).

Genus ENTOSOLENIA Ehrenberg, 1848 

Entosolenia crumenata Cushman

Plate 12, figures 1, 2 ,

Entosolenia crumenata Cushman, Cushman Lab. Foram. Research Contr., vol. 11, p. 31, pi. 4, fig. 9, 1935.

Cushman and McGlamery, U. S. Geol. Survey Prof. Paper 

189-D, p. 109, pi. 26, fig. 11, 1938.

Cushman, Cushman Lab. Foram. Research Contr., vol. 15, 

p. 66, pi. 11, figs. 4, 5, 1939.

Cushman and McGlamery, U. S. Geol. Survey Prof. Paper 

197-B, p. 70, pi. 5, figs. 16-18, 1942.

Cushman and Todd, Cushman Lab. Foram. Research Contr., 

vol. 18, p. 37, pi. 6, fig. 28, 1942.

Cushman, idem, vol. 20, p. 43, pi. 7, fig. 12,1944.

Cushman and Ellisor, Jour. Paleontology, vol. 19, p. 563, 

pi. 76, fig. 3, 1945.

Test single-chambered, longer than broad, compressed; periphery truncate, with a single, broad keel 

in the median line and lesser ones at the angles, these 

forming a broadly elliptical or oval area in side view 

in the middle of each face, the sides continuing into the 

very distinct, compressed, tubular neck; wall clear; 

aperture terminal, narrowly elliptical. Length 0.35 to 

0.40 mm., breadth 0.18 to 0.20 mm., thickness 0.10 to 

0.12 mm.

The types are from the Oligocene of Mississippi and 

it has been recorded from the Oligocene of Texas and 

from Eocene cores from the Atlantic. Specimens show 

considerable variation and more than one species may 

have been included under this name.

Porters Creek formation, Matthews Landing marl member. 

Alabama, Wilcox County (2, 5) ; Butler County (3) ; Choctaw 

County (7).

Naheola formation. Coal Bluff marl member. Alabama, Wilcox County (8).

Entosolenia cf. E. laevigata (Reuss) Cushman and McGlamery

Plate 12, figures 3, 4

Rare specimens from the Paleocene of Alabama are 

compared to E. laevigata with some question.

Porters Creek formation, Matthews Landing marl member. 

Alabama, Wilcox County (5) ; Choctaw County (7).

Naheola formation. Coal Bluff marl member. Alabama, Wilcox County (8).

Entosolenia cf. E. orbignyana (Segueuza) Cushman 

Plate 12, figure 5

Specimens from the Porters Creek clay of Mississippi 

have been referred to as "Lagena orbignyaaia (Seguenza)" by Kline (Mississippi Geol. Survey Bull. 53, 

p. 38, pi. 4, fig. 5, 1943). Kline's figure is reproduced 

on our plate.
42 PALEOCENE FORAMINIFERA OF GULF COASTAL REGION

Entosolenia morsei Kline 

Plate 12, figure 6

Entosolenia morsei Kline, Mississippi Geol. Survey Bull. 53, p. 48,

pi. 4, fig. 17,1943.

Test ovate to globular, greatest width usually a little below 

center, with short apical spine; surface smooth; aperture round, 

small, at end of short but distinct neck. Length of holotype 

0.46 mm.; breadth 0.34 mm.—Kline.

The types are from the Paleocene Porters Creek clay, 

from shallow test hole, 2*4 miles north of Pheba, Clay 

County, Miss. It is also recorded from the Clayton 

formation of Mississippi. It was found in our material at only two localities. The type figure is copied 

on our plate.

Porters Creek clay. Mississippi, Kemper County (17). 

Midway group. Texas, Caldwell County (44).

Entosolenia cf. E. apiculata (Reuss) Cushman 

Plate 9, figure 7

Laaena apiculata Keuss, Akad. Wiss. Wien, Math.-naturwiss. 

Kl., Sitzungsber., vol. 46, p. 318, pi. 1, figs. 4-8, 10, 11, 

1862.

Plummer, Texas Univ. Bull. 2644, p. 75, pi. 4, fig. 6,1927. 

Cushman and Todd, Cushman Lab. Foram. Research Contr.,

vol. 22, p. 56, pi. 9, fig. 25, 1946.

Entosolenia. cf. E. apioulata Cushman, Cushman Lab. Foram. Research Contr., vol. 20, p. 43, pi. 7, fig. 10, 1944.

Bare specimens from a few localities in the Paleocene 

may be compared questionably with E. a/pwuidta. 

Some specimens are circular in section and others are 

compressed. The form is close to-E. morsei Kline but 

does not have a definite neck as in that species.

Porters Creek formation, Matthews Landing marl member.

Alabama, Wilcox County (2) ; Butler County (3). 

Naheola formation, Coal Bluff marl member. Alabama, Wilcox

County (8). 

Midway formation, upper part. Arkansas, Pulaski County

(25).

Entosolenia sp. 

Plate 12, figure 7

Very rare specimens of the smooth form here figured 

occurred at but two localities in the Paleocene of Alabama. More specimens are needed to give a definite 

specific determination and it is here figured for future 

reference.

Porters Creek formation, Matthews Landing marl member. 

Alabama, Wilcox County (2) ; Butler County (3).

Genus VIRGULINA D'OrMgny, 1826

Virgulina wilcoxensis Cushman and Ponton

Plate 12, figures 8, 9

Virgulina wilcoxensis Cushman and Ponton, Cushman Lab.

Foram. Research Contr., vol. 8, p. 67, pi. 8, fig. 22, 1932. 

Cushman, Cushman Lab. Foram. Research Special Pub. 9, p.

67, pi. 8, fig. 22,1937. 

Cushman and Garrett, Cushman Lab. Foram. Research

Contr., vol. 15, p. 82, pi. 14, figs. 19-21, 1939. 

Cushman, idem, vol. 16, p. 67, pi. 11, fig. 19, 1940.

Kline, Mississippi Geol. Survey Bull. 53, p. 4?, pi. 6, fig. 24,

1943. 

Cushman, Am. Jour. Sci., vol. 242, p. 11, pi. 1, figs. 19, 20,

1944; Cushman Lab. Foram. Research Contr., vol. 20, p.

24, pi. 4, fig. 24; p, 43, 1944.

Test elongate, fusiform, somewhat compressed, 

about 2V4 times as long as broad, early portion irregularly spiral, adult irregularly biserial, periphery 

rounded; chambers distinct, very slightly inflated; sutures distinct, very slightly depressed; v^all smooth, 

distinctly perforate; aperture a broad, comma-shaped 

opening at the base of the apertural face in the median 

line. Length 0.50 mm., breadth 0.15 mm., thickness 

0.10 mm.

The types are from the Eocene Wilcox group, from 

railroad cut, 1 mile north of Ozark, Ala.

Specimens apparently identical with the species from 

the Wilcox strata occur at numerous localities in the 

Paleocene of Alabama, Texas, and Arkansas, often very 

abundantly. It is also recorded by Kline from the 

Porters Creek clay and the Clayton formation of Mississippi.

Naheola formation, Coal Bluff marl member. Alabama, Wilcox County (8). 

Clayton formation. Alabama, Barbour County (30).

Chalybeate limestone member. Alabama, S'Tmter County

(9).

Pine Barren member. Alabama, Wilcox Courty (11) ; Butler County (,12).

Porters Creek clay. Tennessee, Hardeman Count^ (13). 

Midway formation, lower part. Arkansas, Saline County

(18, 19) ; Pulaski County (23).

Midway formation, upper part. Arkansas, Pulask' County (24). 

Midway formation. Arkansas, Saline County (2<*). 

Midway group, Texas, Hunt County (41).

Virgulina naheolensis Cushman 

Plate 12, figure 10

Virgulina naheolensis Cushman, Cushman Lab. Foram. Research

Contr., vol. 20, p. 78, 1944. 

Virgulina alabamensis Cushman (not Cushman and McGlamery), idem, vol. 20, p. 43, pi. 7, fig. 13,1944.

Test about 3 times as long as broad, fusiform, somewhat compressed, periphery rounded, very slightly lobulate, little if at all twisting in the early stages, adult 

with 8 to 10 biserial chambers; chambers distinct, very 

slightly if at all inflated, biserial, much higher than 

broad; sutures distinct, very strongly oMique, very 

slightly curved, little if at all depressed; v^all smooth, 

finely perforate; aperture comma-shaped, broadest at 

the outer end. Length 0.42 to 0.60 mm., breadth 0.12 

to 0.15 mm., thickness 0.07 to 0.08 mm.

The types are from the Paleocene Coal Bluff marl 

member of the Naheola formation, creek bottom, just 

west of store at Caledonia, about V4 mile south of center 

of sec. 29, T. 11 N., R 10 E., Wilcox County, Ala. (8).

This species seems to be very rare in thQ. Paleocene 

material examined but is small and might easily be overlooked.
BULIMINIDAE

Genus BOLIVINA D'Orbigny, 1839

Bolivina midwayensis Cushman

Plate 12, figures 11, 12

Bolivina midwayensis Cushman, Cushman Lab. Foram. Research

Special Pub. 6, p. 50, pi. 7, fig. 12,1936; Special Pub. 9, p.

45, pi. 6, figs. 11-13, 1937; idem, Contr., vol. 16, p. 67,

pi. 11, fig. 22, 1940. 

Kline, Mississippi Geol. Survey Bull. 53, p. 49, pi. 4, fig. 14,

1943. 

Cushinan, Am. Jour. Sci., vol. 242, p. 11, pi. 1, figs. 22, 23,

1944; Cushinan Lab. Foram. Research Contr., vol. 20, p.

44, pi. 7, fig. 14, 1944.

Test elongate, very slightly tapering, much compressed, periphery rounded, biserial throughout; chambers distinct, slightly inflated, low and broad, very 

slightly overlapping, of rather uniform shape throughout; sutures distinct, slightly depressed, very strongly 

oblique, forming an angle of at least 45° with the horizontal, and slightly curved; wall smooth, very finely 

perforate; aperture an oval opening, tending very 

slightly to be somewhat removed from the inner margin 

of the last-formed chamber. Length up to 0.85 mm., 

breadth 0.15 to 0.18 mm., thickness 0.08 to 0.10 mm.

The types are from the Paleocene, from a shallow 

ditch at road corner southeast of New Corsicana Reservoir on road to Mildred, Navarro County, Tex.

This species occurs at numerous localities in Alabama, 

Mississippi, and Texas. It seems to have a considerable 

vertical range within the Paleocene and should make 

a good index fossil for the Paleocene. It has, however, 

been recorded with some question from the Bashi marl 

member of Hatchetigbee formation of Wilcox age of 

Alabama.

Porters Creek clay. Mississippi, Kemper County (17).

Porters Creek formation. Matthews Landing marl member.

Alabama, Wilcox County (5). 

Naheola formation, Coal Bluff marl member. Alabama, Wilcox

County (8). 

Clayton formation, Chalybeate limestone member. Mississippi,

Noxubee County (16).

Midway formation. Arkansas, Clark County (30). 

Midway group. Texas, Caldwell County (44). 

Wills Point formation, Mexia member. Texas, Limestone

County (36).

Bolivina crenulata Cushman 

Plate 12, figures 13, 14

Bolivina crenulata Cushman, Cushman Lab. Foram. Research 

Special Pub. 6, p. 50, pi. 7, fig. 13, 1936; Special Pub. 9, p. 

53, pi. 6, figs. 33, 34, 1937; idem, Contr., vol. 20, p. 44, pi. 7, 

figs. 15, 16, 1944.

Test about twice as long as broad, tapering, greatest 

breadth near the apertural end, periphery subacute, 

slightly lobulate; chambers numerous, rather indistinct, 

later ones slightly inflated, about twice as broad as high; 

sutures rather indistinct, sigmoid, slightly oblique; wall 

finely perforate, ornamented by distinct, longitudinal 

ridges, and the base of the chambers crenulate, forming

a pattern of irregularly rounded depressions. Lengtt 

0.40 mm., breadth 0.20 mm., thickness 0.15 mm.

The types are from the upper Eocene Kleinzellner 

Tegel, clay pit, near Budapest, Hungary.

Rare specimens seemingly identical with this species 

occurred in the Paleocene Coal Bluff marl member of 

the Naheola formation, creek bottom, just west of storo 

at Caledonia, about }4 mil6 south of center of sec. 29, 

T. 11 N., R. 10 E., Wilcox County, Ala. (8). 

Bolivina budensis (Hantken) Cushman 

Plate 12, figure 15

Bolivima budensis (Hantken) Cushman, Cushman Lab. Foram,

Research Special Pub. 9, p. 47, pi. 6, figs. 21-23, 1937. 

Cushman, Cushman Lab. Foram. Research Contr., vol. 20. 

p. 44, pi. 7, fig. 17, 1944.

A few specimens from the Paleocene of Alabama 

seem very close to this species, which elsewhere is recorded only from the upper Eocene of Europe. Our 

specimens may be a new species, but not enough specimens are available to warrant a full description.

Naheola formation, Coal Bluff marl member. Alabama, Wilcox 

County (8).

Bolivina sp. A 

Plate 12, figure 16

The figured specimen from the Paleocene Naheola 

formation of Alabama has very oblique sutures, smooth 

surface, and peculiar early chambers that seem almost 

as though partially divided. No other specimens were 

found.

Porters Creek formation, Matthews Landing marl member. Alr.- 

barna, Choctaw County (7).

Bolivina sp. B 

Plate 12, figure 17

The figured specimen, from the Paleocene of Alabama, has a broad test, rounded periphery, nearly hor> 

zontal sutures, and the early portion with a rough surface. More material is needed to warrant a specific 

determination.

Porters Creek formation, Matthews Landing marl member 

Alabama, Choctaw County (7).

Genus LOXOSTOMTTM: Ehrenberg, 1854

Loxostomum applinae (Flummer) Nuttall

Plate 12, figure 18

Bolivina applini Plummer, Texas Univ. Bull. 2644, p. 69, pi. 4,

fig. 1, 1927. 

Loxostomum applinae Nuttall, Jour. Paleontology, vol. 4, p. 28',

pi. 24, figs. 4, 5, 1930.

Plummer, Texas Univ. Bull. 3201, p. 54 (list), 1933. 

Cushman, Cushman Lab. Foram. Research Special Pub. 9,

p. 173, pi. 20, fig. 20,1937; idem, Contr., vol. 16, p. 68, pi. 11,

fig. 23,1940. 

Kline, Mississippi Geol. Survey Bull. 53, p. 50, pi. 5, fig. 7,

1943. 

Cushman and Todd, Cushman Lab. Foram. Research Cont~,,

vol. 22, p. 60, pi. 10, fig. 22,1946.
44 PALEOCENE FOEAMINIFERA OF GULF COASTAL, REGION

Cushman and Bermudez, idem, vol. 24, p. 71, pi. 11, fig. 10, 

1948.

Test very elongate, about five times as long as broad, 

slender, only slightly compressed, very gradually tapering, the adult portion with the sides nearly parallel; 

chambers numerous, fairly distinct, later ones slightly 

inflated, increasing rather regularly in size as added, 

earlier ones distinctly biserial, later ones tending to become somewhat uniserial, the basal border in the adult 

extending backward in definite lobes with straight 

reentrants between; sutures distinct, in the early portion somewhat limbate, later depressed and crenulate, 

very slightly oblique; wall distinctly perforate, earlier 

portion sometimes with slight traces of longitudinal 

grooves or costae, later portion smooth except for the 

crenulations along the sutures; aperture in the adult an 

elongate, oval opening, with a slight lip, tending to become terminal. Length up to 1.00 mm., diameter 0.18 

to 0.20 mm.

The types are from the Paleocene, clay pit of Mexia 

Brick Works, about 1 mile west of the town of Mexia, 

Limestone County, Tex.

Although there are one or two records from other 

regions and formations, this seems to be a characteristic species of the Paleocene of Alabama, Arkansas, Mississippi, and Texas.

Porters Creek clay. Mississippi, Kemper County (17).

Porters Creek formation, Matthews Landing marl member. Alabama, Butler County (3).

Clayton formation, Chalybeate limestone member. Alabama, 

Sumter County (9). Mississippi, Noxubee County (16). 

Pine Barren member. Alabama, Wilcox County (11) ; 

Butler County (12).

Midway formation, lower part. Arkansas, Saline County (19A).

Midway formation, upper part. Arkansas, Pulaski County (24, 

25).

Midway formation. Arkansas, Saline County (26) ; Pulaski 

County (27) ; Clark County (30, 31).

Midway group. Texas, Limestone County (39, 42). Caldwell 

County (44).

Wills Point formation, Mexia member. Texas, Limestone 

County (36).

Loxostomum plummerae Cushman 

Plate 12, figures 19, 20

Loxostoma plummerae Cushman, Cushrnan Lab. Foram. Research Special Pub. 6, p. 59, pi. 8, fig. 13, 1936; Special 

Pub. 9, p. 174, pi. 20, fig. 21, 1937.

Cushman and Todd, idem, Contr., vol. 22, p. 60, pi. 10, fig. 

21, 1946.

Test small, elongate, 3% to 4 times as long as broad, 

very slightly compressed, the early portion biserial, 

later strongly tending to become uniserial; chambers 

distinct, somewhat inflated throughout, more strongly 

so in the adult; sutures distinct, depressed, nearly at 

right angles to the periphery; wall distinctly perforate, smooth; aperture in the adult terminal, elongate,

elliptical, with a slight lip. Length 0.40 mm., breadth 

0.10 to 0.12 mm.

The types are from the Paleocene, Dry Creek, % 

mile above bridge on first road east of Travis-Bastrop 

County line, Bastrop County, Tex. (49).

This species is much less common than L. applinae, 

being recorded only from the Paleocene of Texas and 

Arkansas.

Midway formation, upper part. Arkansas, Pulaski County (24,

25).

Midway group. Texas, Bastrop County (49). 

Wills Point formation. Texas, Caldwell County (34).

Loxostomum deadericki Cushman 

Plate 12, figures 21-23

Loxostomum deadericM Cushman, Cushman Lab. Foram. Research Contr., vol. 23, p. 85, pi. 18, figs. 8-10, 1947.

Test elongate, slender, little if at all compressed, early 

portion biserial and somewhat twisted, later chambers 

becoming uniserial; chambers distinct, slightly inflated, 

increasing very gradually in size as added; sutures distinct, slightly depressed, earlier ones strongly oblique, 

becoming less so, and in the last-formed portion at right 

angles to the elongate axis; wall smooth, but distinctly 

perforate; aperture in the adult terminal, narrow. 

Length 0.60 to 0.90 mm., breadth 0.17 to 0.20 mm; thickness 0.15 to 0.18 mm.

The types are from the Paleocene, north side of Highway 67, 8 miles southwest of Benton, Saline County, 

Ark. (26). The species is very common at the type 

locality.

It differs from L. applinae (Plummer) in the less 

tapering form and only slight compression, and is without the basal indentations.

Loxostomum deadericki Cushman var. exilis Cushman

Plate 12, figures 24-26

Loxostomum deadericki Cushman var. exilis Cushman, Cushman Lab. Foram. Research Contr., vol. 23, p. 85, pi. 18, figs. 

11-13, 1947.

Variety differing from the typical form in tl*°. smaller 

size and more slender form, deeper sutures and more 

inflated chambers, and the more tapering earl;? portion. 

Length 0.40 to 0.60 mm., breadth 0.10 to 0.12 mm., thickness 0.08 to 0.10 mm.

The types are from the Paleocene Clayton formation, 

top of fuller's earth zone, immediately underlying 

Turritella rock, SE 14 SE y± sec. 18, T. 11 N., R. 14 E., 

Butler County, Ala. (12).

Genus ANGULOGEEJNA Cushman, 1927

Angulogerina wilcoxensis (Cushman and Ponton) Cushman and

Garrett

Plate 12, figure 27

Pseudouvigerina, wilcoxensis Cushman and Ponton, Cushman 

Lab. Forani. Research Contr., vol. 8, p. 66, pi. 8, fi#. 18, 1932.

* W)
ELUPSOIDINIDAE 45

Angulogerina wilooxensis Cushman and Garrett, idem, vol. 15,

p. 84, pi. 14, figs. 24, 25,1939. 

Toulmin, Jour. Paleontology, vol. 15, p. 599, pi. 80, flg. 30,

1941. 

Cushman, Cushman Lab. Foram. Research Contr., vol. 20,

p. 44, pi. 7, flg. 18, 1944.

Test small, elongate, about twice as long as broad, 

sides in the adult nearly parallel, triangular in transverse section, the angles with two distinct ribs and a 

deep channel between; chambers fairly distinct, 

strongly curved, not depressed; wall coarsely perforate; 

aperture in the adult terminal, with a short neck and 

slight lip. Length 0.30 mm., diameter 0.15 mm.

The types are from the Eocene Wilcox group, from 

railroad cut, 1 mile north of Ozark, Ala.

This species is rare in the Paleocene, specimens occurring in the Coal Bluff marl member of the Naheola formation in Alabama and in the Clay ton formation of 

Tennessee.

Naheola formation, Coal Bluff marl member. Alabama, Wilcox County (8). 

Porters Creek clay. Tennessee, Hardeman County (13,14).

Angulogerina virginiana Cushman 

Plate 12, figure 28

Angulogerina virginiana Cushman, Cushman Lab. Foram. Research Contr., vol. 20, p. 25, pi. 4, fig. 23; p. 44, pi. 7, fig. 19, 

1944.

Test minute, elongate, tapering, triangular in section, 

the sides concave and the angles slightly rounded, triserial except for the last 2 or 3 chambers in the adult, 

which are irregularly uniserial; chambers of the early 

portion indistinct, in the adult somewhat separated and 

more inflated; sutures indistinct except in the very last 

portion in the adult where they are somewhat depressed; 

wall ornamented with fine longitudinal costae continuous over most of the test; aperture in the adult terminal, with a short neck and slight lip. Length 0.30 to 

0.40 mm., diameter 0.10 to 0.12 mm.

The types are from the Eocene Aquia formation, 19- 

22-foot sample, below second rock layer, Marlboro 

Point, Stafford County, Va.

The only Paleocene occurrence of specimens referable 

to this species is from the Coal Bluff marl member of 

the Naheola formation, creek bottom, just west of store 

at Caledonia, about 14 mile south of center of sec. 29, T. 

11 N., E. 10 E., Wilcox County, Ala. (8).

Genus TRIFARINA Cushman, 1923

Trifarina herberti Cushman and Renz

Plate 12, figures 29, 30

Trifarina, herberti Cushman and Renz, Cushman Lab. Foram. 

Research Contr., vol. 18, p. 9, pi. 2, figs. 11, 12, 1942.

Test elongate, about twice as long as broad, triangular 

in transverse section, sides slightly concave, angles 

rounded, rapidly tapering in the microspheric form, 

more fusiform in the megalospheric; chambers few,

increasing rapidly in height as added; sutures distinct, 

depressed, strongly curved upward in the middle of each 

lateral face; wall smooth; aperture terminal, rounded, 

at the end of a short, tubular neck. Length 0.50 to 

0.55 mm., diameter 0.25 mm.

The types are from the Paleocene Soldado Bock formation, Soldaclo Eock, Trinidad, B. W. I. It has not 

been recorded elsewhere.

Family ELLIPSOIDINIDAE

Genus PLEUROSTOMELIA Reuss, 1860

Pleurostomella paleocenica Cushman

Plate 12, figures 31-33

Pleurostomella paleocenica Cushman, Cushman Lab. Foram.

Research Contr., vol. 23, p. 86, pi. 18, figs. 14,15, 1947. 

Pleurostomella alternans Plummer (not Schwager), Texas Univ.

Bull. 2644, p. 69, pi. 4, fig. 2, 1927. 

Pleurostomella cf. P. ftrevis var. alternans Kline (not

Schwager), Mississippi Geol. Survey Bull. 53, p. 50, pi. 6,

flg. 23, 1943.

Test very small, slender, taj>ering from the slightly 

rounded base to the greatest breadth formed by the last 

pair of chambers, nearly circular in transverse section; 

chambers few, distinctly inflated in the later portion, 

nearly as wide as high, increasing rapidly in size in the 

early portion, only slightly in the later portion; sutures 

distinct, later ones strongly depressed; wall smooth; 

aperture on the upper part of the inner face of the 

last-formed chamber, with a deeper reentrant toward 

the base of the opening, without definite tooth-like projections. Length 0.38 to 0.45 mm., breadth 0.10 to 0.12 

mm., thickness 0.10 to 0.12 mm.

The types are from the Paleocene, 4.6 miles north of 

Fentress, Caldwell County, Tex.

This species has been referred to P. alternans 

Schwager, a name that has been used for specimens 

from Cretaceous to Recent. A comparison with topotypes from the Pliocene of the Pacific shows that the 

Paleocene species differs in the much smaller size, fewer 

and shorter chambers, and smaller and more rounded 

aperture.

Kline has recorded it from the Porters Creek clay of 

Mississippi and specimens occur in the Paleocene collections from Texas and Arkansas.

Midway formation. Arkansas, Clark County (30). Midway

group. Texas, Caldwell County (44). 

Wills Point formation, Mexia member. Texas, Limestone

County (36).

Genus NODOSARELLA Rzehak, 1895

Nodosarella attenuata (Plummer) Cushman

Plate 12, figures 34-37

Ellvpsopleurostomella attenuata Plummer, Texas Univ. Bull. 

2644, p. 131, pi. 8, fig. 6,1927. *

Nodosarella attenuata Cushman, Cushman Lab. Foram. Research Contr., vol. 16, p. 68, pi. 11, fig. 30,1940. 

Test long, tapering, straight or slightly uneven; chambers

inflated, smooth, first two or three Textularian followed by a

Bifarine series that merges into true Nodosarian succession;
46 PALEOCENE FORAMINIFERA OF GULF COASTAL REGION

sutures faint lines in early part of test and faint constrictions 

above; successive apertures connected by an internal column, 

and final aperture a somewhat curved terminal fissure one side 

of which arches slightly over the other. Length up to 0.8 mm.— 

Plummer.

The types are from the Paleocene, clay pit, Mexia 

Brick Works, about 1 mile west of the town of Mexia, 

Limestone County, Tex.

Clayton formation, Chalybeate limestone member. Alabama,

Sumter County (9).

Midway formation. Arkansas, Saline County (26). 

Midway group. Texas, Limestone County (42).

Nodosarella paleocenica Cushman and Todd 

Plate 12, figure 38

Nodosarella paleocenica Cushman and Todd, Cushman Lab. 

Foram. Research Contr., vol. 22, p. 60, pi. 10, fig. 23, 1946.

Test small, elongate, often slightly curved, tapering 

from the very narrow initial end to the greatest width 

formed by the last one or two chambers, rounded in section ; chamber§ distinct, later ones inflated, increasing 

rather rapidly in size and height as added, in the adult 

slightly longer than broad; sutures distinct, in the later 

portion depressed; wall smooth; aperture terminal, narrow, elongate, with a slightly raised lip at one side. 

Length up to 1.00 mm., breadth 0.20 to 0.23 mm.

The types are from the Paleocene, about 1,000 feet 

south of Roosevelt Road viaduct, 200 feet east of railroad tracks, in small gully heading east, just upstream 

from small abandoned bridge, NEi/4 NW*4 sec. 16, T. I 

N., R. 12 W., Little Rock, Ark. (25).

The only other specimen is from the Matthews Landing marl member of the Porters Creek formation on 

Alabama Highway 96, 9.2 miles northeast of Kimbrough, Wilcox County, Ala. (5).

Genus ELLIPSONODOSARIA A. Silvestri, 1900 

Ellipsonodosaria plummerae Cushman 

Plate 13, figures 1, 2 »

Nodosaria sagrinensis Plummer (not Bagg), Texas Univ. Bull.

2644, p. 85, pi. 4, fig. 16,1927.

Ellipsonodosaria plummerae Cushman, Cushman Lab. Foram. 

Research Contr., vol. 16, p. 69, pi. 12, figs. 4, 5, 1940.

Kline, Mississippi Geol. Survey Bull. 53, p. 51, pi. 5, fig. 8,

1943. 

Cooper, Jour. Paleontology, vol. 18, p. 352, pi. 54, figs. 18,19,

1944. 

Cushman and Todd, Cushman Lab. Foram. Research Contr.,

vol. 22, p. 61, pi. 10, fig. 24, 1946.

Ellipsonodosaria sagrinensis Cooper (not Bagg), Jour. Paleontology, vol. 18, p. 352, pi. 54, fig. 22,1944.

Test elongate, very slightly tapering, chambers in a 

straight linear series, circular in transverse section, initial end with one or more short spines; chambers distinct, later o*s becoming pyriform with the greatest 

breadth toward the base, which is somewhat excavated, 

increasing in size very gradually as added; sutures 

deeply excavated in the later portion; wall ornamented 

byrlow, longitudinal costae, broken into irregular short

spines, and limited largely to the upper part of the 

chamber, ending often in short spines at th°, ridge near 

the base of the chamber; aperture terminal, rounded, 

with a distinct tooth at one side, and with a definite neck 

and slight lip. Length 0.80 to 1.00 mm., diameter 0.10 

to 0.14 mm.

The types are from the Paleocene, from U. S. Highway 80, south of Sucarnoochee Creek, y2 mile southwest 

of Livingston, Sumter County, Ala.

This is a very common species in the Paleocene and 

should make a good index fossil. In addition to the 

localities given below it has been recorded by Kline 

from the Paleocene of Mississippi and by Cooper from 

the Paleocene of Illinois. 

Porters Creek formation, Matthews Landing irarl member.

Alabama, Butler County (3). 

Clayton formation, Chalybeate limestone member. Alabama,

Sumter County (9). Mississippi, Noxubee Comty (16).

Pine Barren member. Alabama, Butler County (12). 

Midway formation, upper part. Arkansas, Pulaski County

(24, 25). 

Midway formation. Arkansas, Saline County (26) ; Pulaski

County (27) ; Clark County (30, 31). 

Midway group. Texas, Limestone County (39, 42) ; Navarro

County (40) ; Bastrop County (43) ; Caldwell County (44) ;

Travis County (45) ; Freestone County (50). 

Wills Point formation. Texas, Travis County (33) ; Caldwell

County (34).

Mexia member. Texas, Limestone County (3?). 

Kincaid formation. Texas, Hunt County (37).

Ellipsonodosaria paleocenica Cushman and Todd 

Plate 13, figures 3-5

Ellipsonodosaria paleocenic-a Cushman and Todd, C tshman Lab.

Foram. Research Contr., vol. 22, p. 61, pi. 10, fig. 26, 1946. 

Ellipsonodosaria sp. Cushman, idem, vol. 16, p. 70, pi. 12, figs.

1, 2, 1940.

Test elongate, slender, straight, tapering very gradually from the very narrow initial end to the greatest 

breadth near the apertual end; chambers distinct, inflated, numerous as many as 14 in the adult, increasing 

very gradually in size as added, strongly inflated, subspherical ; sutures distinct, strongly depressed throughout ; wall nearly smooth or slightly hispid in the earliest 

portion, in the last two or three chambers with a trace 

of very short spines; aperture terminal with a short 

neck and distinct lip, with a short tooth at one side of 

the opening. Length 1.50 to 1.90 mm., breadth 0.20 to 

0.25 mm.

The types are from the Paleocene, about 1,000 feet 

south of Roosevelt Road viaduct, 200 feet e<r,st of railroad tracks, in small gully heading east, jus4:, upstream 

from small abandoned bridge, NE% NW1^ sec. 16, T. 

1 N., R. 12 W., Little Rock, Ark.

This should make a good index fossil for the Paleocene. It may be distinguished from E. plummerae by 

its more rounded chambers and smoother surface.

Porters Creek clay. Mississippi, Kemper County (17). 

Porters Creek formation, Matthews Landing marl member. 

Alabama, Wilcox County (5).
ROTALIIDAE 47

Clayton formation, Chalybeate limestone member. Alabama, 

Sumter County (9). Mississippi, Noxubee County (16).

Midway formation, lower part. Arkansas, Saline County (20).

Midway formation, lower part. Arkansas, Pulaski County (25).

Midway formation. Arkansas, Saline County (26). Midway, 

group. • Texas, Limestone County (39) ; Navarro County (40) ; 

Travis County (46) ; Bastrop County (49).

Wills Point formation. Texas, Williamson County (35).

Ellipsonodosaria midwayensis Cushman and Todd 

Plate 13, figures 6-10

Ellipsonodosaria midwayensis Cushman and Todd, Cushman Lab. 

Foram. Research Contr., vol. 22, p. 61, pi. 10, fig. 25, 1946. 

Nodosaria spinulosa Plummer (not Nautilus spinulosus Montagu), Texas Univ. Bull. 2644, p. 84, pi. 4, fig. 19,1927. 

Ellipsonodosaria spinulosa (Plummer) Cooper, Jour. Paleontology, vol. 18, p. 352, pi. 54, fig. 20, 1944.

Ellipsonodosaria alexanderi Cushman (not Cushman, 1936), 

Cushman Lab. Foram. Research Contr., vol. 16, p. 69, 

pi. 11, figs. 27-29, 1940.

Kline (part), Mississippi Geol. Survey Bull. 53, pi. 5, fig. 

1 (not fig. 2), 1943.

Test elongate, slender, arcuate, gradually tapering 

from the acute initial end to the greatest width at the 

last-formed chamber, initial end with one or more distinct, short spines; chambers distinct, pyriform in 

shape, greatest width near the base, sides tapering but 

convex; sutures distinct, strongly depressed; wall ornamented with numerous, platelike longitudinal costae 

ending in two series of spinose projections near the base 

of the chambers, in the earlier portion often with the 

costae continuous over several chambers; aperture terminal, with a very short neck and distinct lip. Length 

2.00 to 2.60 mm., breadth 0.25 to 0.30 mm.

The types are from the Paleocene, about 1,000 feet 

south of Eoosevelt Road viaduct, 200 feet east of railroad tracks, in small gully heading east, just upstream 

from small abandoned bridge, NE1^ NW14 sec. 16, T. 

1 N., E. 12 W., Little Rock, Ark. (25).

This species should make a good index fossil for the 

Paleocene. It occurs mostly in material from Texas 

but there are occasional specimens from Arkansas and 

a questionable one from Alabama. One of Kline's figures is of this species and gives a record from the Paleocene of Mississippi. Cooper's specimens from Illinois should apparently be placed here.

Clayton formation. Alabama, Barbour County (10). 

Midway formation, upper part. Arkansas, Pulaski County

(24, 25). 

Midway formation. Arkansas, Pulaski County (27) ; Clark

County (29). 

Midway group. Texas, Limestone County (39, 42) ; Navarro

County (40) ; Hunt County (41) ; Van Zandt County (47,

48) ; Freestone County (50).

Ellipsonodosaria alexanderi Cushman

Plate 13, figure 12

Ellipsonodosaria alexanderi Cushman, Cushman Lab. Foram.

Research Contr., vol. 12, p. 52, pi. 9, figs. 6-9, 1936. 

Kline (part), Mississippi Geol. Survey Bull. 53, p. 51, pi. 

5, fig. 2 (not fig. 1), 1943.

Cushman, U. S. Geol. Survey Prof. Paper 206, p. 135, p1, 

56, figs. 12-15, 1946.

Test elongate, straight or slightly curved, microspheric form increasing rather rapidly in diameter from 

the small proloculum, the megalospheric form with tli e 

proloculum having nearly as great a diameter as the 

last-formed chambers; chambers distinct, inflated, increasing rather gradually in length, the adult chambers 

about twice as long as broa'd; sutures distinct, strong]^ 

depressed; wall ornamented with short backward!^ 

pointing spines, in the early stages of the microspher; c 

form with a single ring of spines slightly below tre 

middle of the chamber, in the adult with numerous 

spines rather irregularly scattered over the surface; 

aperture a semicircular opening with a single tooth; 

with a distinct neck and slightly raised lip. Length up 

to 2.00 mm., diameter 0.20 mm.

The types are from the Upper Cretaceous Taylor 

marl in a road cut 14.4 miles south of Paris and 0.9 

mile north of Lake City, Delta County, Tex.

This is typically an Upper Cretaceous species. Very 

rare specimens from a single Paleocene locality in Texas 

may be referred here with some question. One of th-3 

specimens figured by Kline from the Paleocene of Mississippi seems to belong here. 

Midway group. Texas, Caldwell County (44).

Family BOTALIIDAE

Genus SFIBILLINA Ehreuberg, 1843

Spirillina selseyeusis Heron-Alien and Earlaud

Plate 13, figures 13, 14

Spirillina selseyensis Heron-Alien and Earland, Royal Micr. Soc.

Jour., 1909, p. 440, pi. 18, figs. 6, 7. 

Cushman and Garrett, Cushman Lab. Foram. Research

Contr., vol. 15, p. 84, pi. 14, figs. 28, 29,1939. 

Cushman and Todd, idem, vol. 18, p. 38, pi. 7, fig. 2, 1942 

Cushman, -idem, vol. 20, p. 45, pi. 7, fig. 20,1944.

Test close coiled, dorsal side somewhat concave, ventral side flattened or slightly concave near the cente^; 

spiral suture distinct; wall of the dorsal side with a 

series of rectangular depressions extending from the 

peripheral thickening to the inner margin of uniform 

size, ventral side with slightly oblique linear depressions with raised areas between; aperture at the end of 

the tubular chamber. Diameter 0.25 to 0.30 mm.

The types of this species are from the Eocene of S4- 

sey Bill, England. It has been recorded in America 

from the Eocene Wilcox group of Woods Bluff, Alabama, and from the Paleocene Naheola formation, of 

Alabama.

Naheola formation. Coal Bluff marl member. Alabama, Vilcox County (8).

Porters Creek clay. Tennessee, Hardeman County (13, 14). 

Porters Creek formation. Matthews Landing marl menil ^r.

Alabama, Choc taw County (7).
48 PALEOCENE FORAMINIFERA OF GULF COASTAL REGION

Spirillina cf. S. vivipara Ehrenberg 

Plate 13, figure 15

Spirillina cf. vivipara Ehrenberg, Cushman and Todd, Cushman 

Lab. Foram. Research Contr., vol. 18, p. 38, pi. 7, fig. 1, 1943. 

Cushman, idem, vol. 20, p. 44, pi. 7, fig. 21, 1944.

Single specimens have been recorded and figured as 

noted above.

Porters Creek formation, Matthews Landing marl member.

Alabama, Wilcox County (5) ; Choctaw County (7). 

Naheola formation. Coal Bluff marl member. Alabama, Wilcox County (8).

Spirillina sp.

Plate 13, figures 16, 17

Spirillina sp. Cushman and Renz, Cushman Lab. Foram. Research Contr., vol. 18, p. 10, pi. 2, figs. 14, 15,1942.

Specimens from the Paleocene of Soldado Rock, Trinidad, B. W. I., evidently belong to this genus or to Conicospirittina. They are not preserved well enough for 

a specific description.

Genus PATELLINA Williamson, 1858

Patellina sp. 

Plate 13, figure 18

Patellina sp. Cushman, Cushman Lab. Foram. Research Contr., 

vol. 20, p. 45, pi. 7, fig. 22, 1944.

Very rare specimens from the Paleocene Coal Bluff 

marl member of the Naheola formation of Wilcox 

County, Ala., and from Tennessee seem to belong to this 

genus, but more specimens are needed to give an adequate description.

Naheola formation, Coal Bluff marl member. Alabama, Wilcos

County (8). 

Porters Creek clay. Tennessee, Hardeman County (13).

Genus PATELLINOIDES Heron-Alien and Earland, 1932 

Patellinoides? sp. 

Plate 13, figure 19

Patellinoides? sp. Cushman and Todd, Cushman Lab. Foram. Research Contr., vol. 18, p. 38, pi. 7, fig. 3, 1942.

A single specimen from the Paleocene at the type 

locality of the Naheola formation, Naheola Landing, 

on Tombigbee River, Choctaw County, Ala. (7), is referred with some question to this genus. The adult 

chambers make up about half a coil. The sutures on the 

dorsal side are raised and plate-like. It apparently 

does not belong to any described species, but more specimens are needed for a full specific description.

Genus DISCORBIS Lamarck, 1804

Discorbis midwayensis Cushman

Plate 13, figure 20

Discorbis midwayensis Cushman, Cushman Lab. Foram. Research Contr., vol 16, p. 70, pi. 12, fig. 6, 1940. 

Kline, Mississippi Geol. Survey Bull. 53, p. 52, pi. 5, figs. 

9,10, 1943.

Test trochoid, plano-convex, dorsal side somewhat 

convex, ventral side flattened or even slightly concave,

periphery subacute but not keeled, ventral side umbilicate; chambers normally 7 in the last-forni°,d whorl, 

fairly distinct, of uniform shape, increasing gradually 

in size as added, slightly inflated on the dorsal side; 

sutures dfstinctly curved on both dorsal and ventral 

sides, slightly depressed; wall distinctly papillate on 

both dorsal and ventral sides, the center of the dorsal 

side sometimes slightly umbonate and smooth; aperture 

on the ventral side, in the umbilical region, low and 

elongate, with a distinct, overhanging lip. Diameter 

0.65 to 0.80 mm., thickness 0.18 to 0.25 mm.

The types are from the Paleocene, from an old abandoned roadway, south of Sucarnoochee Cre?k, about 

y10 mile upstream from crossing of U. S. Highway 80, 

1/2 mile southwest of Livingston, Sumter County, 

Ala. (9).

Clayton formation, Pine Barren member. Alabama, Butler

County (12). . 

Midway formation. Arkansas, Saline County (26) ; Pulaski

County (28) ; Clark County (31).

Discorbis midwayensis Cushman var. soldadoensis Cushman and

Renz

Plate 13, figure 21

Discorbis midwayensis Cushman var. soldadoensis Cushman 

and Renz, Cushman Lab. Foram. Research Contr., vol. 18, 

p. 10, pi. 3, fig. 4,1942.

Variety differing from the typical form in th°! thicker, 

more convex test, fewer chambers, deeper umbilical region, narrower and more triangular and more inflated 

final chamber.

The types of the variety are from the Paleocene Soldado Rock formation, Soldado Rock, Trinidad, B. W. I. 

It has not been recorded elsewhere.

Discorbis midwayensis Cushman var. trinitatensis CurTiman and

Renz 

Plate 13, figure 22

Discorbis midwayensis Cushman var. trinitatensis Cushman 

and Renz, Cushman Lab. Foram. Research Contr., vol. 18, 

p. 10, pi. 3, fig. 5,1942. 

Cushman, idem, vol. 20, p. 45, pi. 7, fig. 23, 1944.

Variety differing from the typical form in the ornamentation of the test, consisting of thickened and raised 

sutures on the dorsal side and deeper sutures on the ventral side.

The types of the variety are from the Paleocene Soldado Rock formation, Soldado Rock, Trinidad, B. W. I., 

and it has not been recorded elsewhere.

Discorbis infrequent Plummer 

Plate 13, figure 23

Disoorbis infrequens Plummer, Texas Univ. Bull. 2644, p. 138,

pi. 9, figs, la-c, 1927.

Test very small, round, trochoid, the dorsal face beir-? broadly 

conical and the ventral face almost fiat; peripheral margin 

acute, very faintly lobate; chambers 3 or 4 in final whorl, narrow, strongly curved, very smooth; aperture a narrow arched
ROTALIIDAE 49

opening along the edge of the final chamber between the periphery 

and the umbilicus. Diameter up to 0.3 mm.—Plummer.

The types are from the Paleocene, exposure in side of 

steep west-facing hill on Richland-Streetman road U/2 

miles southwest of Richland where roads branch off 

both northeast and southwest with a short offset, Navarro County, Tex.

The species is recorded as exceedingly rare. No 

specimens have been found in our material.

Genus LAMARCKINA Berthelin, 1881

Lamarckina naheolensis Cushman and Todd

Plate 14, figures 4-6

Lamarckina naheolensis Cushman and Todd, Cushman Lab. 

Foram. Research Contr., vol. 18, p. 39, pi. 7, figs. 5-7, 1942. 

Cushman, idem, vol. 20, p. 45, pi. 7, fig 25, 1944.

Test small, longer than broad, dorsal side flattened or 

even slightly concave in the earlier portion, ventral side 

strongly convex, deeply umbilicate, periphery acute, 

even slightly keeled in the early stages; chambers distinct, about 8 in the adult whorl, increasing rapidly in 

size as added; sutures distinct on the dorsal side, slightly 

limbate as each chamber tends to slightly overlap the 

previous one, gently curved, on the ventral side indistinct; wall smooth, distinctly perforate on the dorsal 

side, smooth and polished on the ventral side; aperture 

opening into the umbilical cavity, with a large, slightly 

convex lip extending into and partially covering the 

umbilicus. Length of holotype 0.25 mm., breadth 0.20 

mm., thickness 0.13 mm.

The types are from the Paleocene Naheola formation, 

greensand bed, upper fossiliferous horizon, Naheola 

Landing, Tombigbee River, Ala.

Porters Creek formation, Matthews Landing marl member. 

Alabama, Wilcox County (2, 4-6) ; Choctaw County (7).

Naheola formation. Coal Bluff marl member. Alabama, Wilcox County (8).

Lamarckina limbata Cushman and Todd 

Plate 14, figures 1-3

Lamarckina limbata Cushman and Todd, Cushman Lab. Foram. 

Research Contr., vol. 18, p. 39, pi. 7, figs. 8-10, 1942.

Test about as broad as long, biconvex, periphery acute 

and slightly carinate; chambers distinct, 5 or 6 in the 

adult whorl, increasing rapidly in size as added; sutures on the dorsal side strongly limbate and raised, 

only slightly curved toward the periphery, indistinct 

on the ventral side, especially on the earlier chambers, 

smooth and polished on the ventral side; aperture 

opening into the umbilical cavity, without a definite 

lip. Length of holotype 0.32 mm., breadth 0.28 mm., 

thickness 0.18 mm.

The types are from the Paleocene Naheola formation, greensand bed, upper fossiliferous horizon, Naheola Landing, Tombigbee River, Ala.

Porters Creek formation, Matthews Landing marl membe". 

Alabama, Butler County (3), Choctaw County (7).

Lamarckina rugulosa Plummer 

Plate 13, figure 24

Lamarckina rugulosa Plummer, Ms, in Cushman, Cushman Lab. 

Foram. Research Contr., vol. 2, pt. 1, p. 8, pi. 3, figs. 6a-c, 

1926; Univ. Texas Bull. 2644, p. 140, pi. 9, figs. 3a-c, 1927.

Test very broadly elliptical to almost round in outline, molerately compressed in average development to nearly globular 

in extreme old age; convolutions not over 1%; chambers 5-6 

in the last-formed whorl, enlarging rapidly, very smooth and 

glistening on the ventral face, highly granular on the dorsal 

face; dorsal sutures slightly depressed between last 2 or 3 

chambers, but commonly obliterated by the granulations, or 

rarely marked by faint ridges of smooth shell material; ventral 

sutures faintly depressed; umbilicus deeply excavated; aperture a low arch on the umbilical edge of the final chamber und^r 

a narrow and delicately fringed flap on well-preserved tes^s. 

Maximum length up to 0.50 mm.; maximum breadth 0.40 mm.; 

thickness 0.30 mm.—Plummer in Cushman.

The types are from the Paleocene, road cut sou^h 

of city reservoir, 3% miles southeast of Corsicana, 

Tex. It has also been recorded from the Clayton formation, from a bluff on the south side of Owl Creek, 

2% miles northeast of Ripley, Tippah County, Miss.

Wills Point formation. Texas, Caldwell County (34).

Mexia member. Texas, Limestone County (36). 

Midway group. Texas, Bastrop County (43) ; Caldwell Cour+y 

(44).

Lamarckina paleocenica Cushman 

Plate 14, figure 7

Lamarckina paleocenica Cushman, Cushman Lab. Foram. Research Contr., vol. 24, p. 44, pi. 8, fig. 8, 1948.

Test small, dorsal side moderately convex, ventral 

side flattened or slightly concave, slightly umbilicate, 

periphery subacute; chambers distinct on the dorsal 

side, rather indistinct on the ventral side, 6 or 7 in 

the adult whorl, increasing rapidly in size as added; 

sutures distinct on the dorsal side, slightly thicker ed 

and somewhat raised in the earlier portion, the last one 

or two often depressed, rather indistinct on the ventral 

side; wall smooth except for the raised sutures of the 

dorsal side; aperture ventral, beneath an extended lobe 

of the last-formed chamber. Length 0.32 to 0.37 mm., 

breadth 0.27 to 0.30 mm., thickness 0.15 to 0.18 mm.

The types are from the Paleocene Clayton formation, 

cut on Southern Railway, 200 feet east of mile post 

481, 1% miles east of Middleton, Hardeman County, 

Tenn.

The species is common at the type locality and another locality a short distance away and at one Alabama 

locality. It differs from L. ripleyensis Cushman in 

he more compressed and smaller test, fewer chambers, 

and less raised sutures.

Porters Creek clay, Tennessee, Hardeman County (13, 14). 

Porters Creek formation, Matthews Landing marl member. 

Alabama, Wilcox County (1).
50 PALEOCENE FORAMINilFERA OF GULF COASTAL REGION

Genus VALVUIINERIA Cushman, 1926

Valvulineria allomorphinoides (Reuss) Cushman

Plate 14, figures 8, 9

Valvulina allomorphinoides Reuss, Akad. Wiss. Wien, Math.-

naturwiss. Kl., Sitzungsber., vol. 40, p. 223, pi. 11, figs.

6a-c, 1860. 

Discorbina allomorphinoides Franke, Greifswald Univ. Geol.-

palaeont. Inst, Abh., vol. 6, p. 91, pi. 8, figs, lla, b, 1925,

Preuss. geol. Landesanstalt Abh., new ser., vol. Ill, p. 189,

pi. 18, figs. 7a, b, 1928. 

DiscorMs allomorphinoides Cushman, Am. Assoc. Petroleum

Geologists Bull., vol. 10, p. 606, pi. 20, figs. 18, 19; pi. 21,

fig. 5, 1926.

Plummer, Texas Univ. Bull. 2644, p. 139, pi. 9, fig. 2, 1927. 

Valvulineria allomorpMnoides Cushman, Cushman Lab. Foram.

Research Contr., vol. 7, p. 43, pi. 6, figs. 2a-c, 1931; Tennessee Div. Geology Bull. 41, p. 53, pi. 9, figs. 6a-c, 1931; U. S.

Nat. Mus. Proc., vol. 80, art. 14, p. 46, pi. 13, figs. 17a-c,

1932. 

Brotzen, Sveriges geol. Undersb'kning, ser. C, No. 396, p.

153, pi. 11, figs, la-c, text fig. 56, 1936. 

Morozova, Soc. naturalistes Moscou Bull., n. ser., vol. 47,

p. 78, pi. 2, figs. 21, 22, 1939. 

Cushman, Cushman Lab. Foram. Research Contr., vol. 16, p.

70, pi. 12, fig. 9, 1940. 

Cushman and Hedberg, idem, vol. 17, p. 96, pi. 23, figs.

9a-c, 1941. 

Kline, Mississippi Geol. Survey Bull. 53, p. 52, pi. 5, figs. 11,

12, 1943.

Applin and Jordan, Jour. Paleontology, vol. 19, p. 132 (list), 

1945.

Cushman and Todd, Cushman Lab. Foram. Research Contr., 

vol. 22, p. 62, pi. 11, fig. 1,1946.

Cushman, U. S. Geol. Survey Prof. Paper 206, p. 138, pi. 57, 

figs. 6, 7, 1946.

Cushman and Renz, Cushman Lab. Foram. Research Special

Pub. 18, p. 44, pi. 17, figs. 13, 14, 1946.

Schijfsma, Med. Geol. Stichting, ser. C-V, No. 7, p. 89,1946. 

Cushman and Renz, Cushman Lab. Foram. Research Contr.,

vol. 23, p. 48, 1946.

Test trochoid, biconvex, slightly longer than broad, 

oval, periphery rounded; chambers distinct, very 

slightly inflated, 4 to 6 or T in the adult whorl, usually 

5; sutures distinct, dorsally slightly curved, not depressed ; wall smooth, very finely perforate; aperture a 

low opening, on the ventral side, beneath an overhanging, plate like lip. Length 0.25 to 0.40 mm., breadth 

0.20 to 0.35 mm., thickness 0.13 to 0.20 mm.

This species was described from the Upper Cretaceous of Europe and has a wide distribution in the 

Cretaceous and lower Eocene.

Porters Creek formation, Matthews Landing marl member.

Alabama, Wilcox County (2); Butler County (3). 

Clayton formation, Chalybeate limestone member. Alabama,

Sumter County (9).

Midway formation, upper part. Arkansas, Pulaski County (25). 

Midway formation. Arkansas, Saline County (26) ; Pulaski 

County (27) ; Clark County (30).

Midway group. Texas, Limestone County (39, 42) ; Caldwell 

County (44).

Wills Point formation, Mexia member. Texas, Limestone

County (36). 

Kincaid formation. Texas, Hunt County (37).

Valvulineria wilcoxensis Cushman and Ponton 

Plate 14, figures 10-13

Valvulineria loilcoxemis Cushman and Ponton, Cusl Tuan Lab.

Foram. Research Contr., vol. 8, p. 70, pi. 9, fig. 6, 1932. 

Cushman and Garrett, idem, vol. 15, p. 85, pi. 15 figs. 1, 2,

1939. 

Cushman, Am. Jour. Sci., vol. 242, p. 13, pi. 1, fiTO. 36, 37,

1944; Cushman Lab. Foram. Research Contr., vo1 . 20, p. 26,

pi. 4, fig. 26, 1944.

Test nearly circular in side view, periphery slightly 

lobulate, nearly bilaterally symmetrical, umbilical region slightly depressed on the ventral side; chambers 

very distinct, 6 or 7 in the last-formed whorl, inflated; 

sutures distinct, slightly curved, strongly linbate on 

the ventral side, less so on the dorsal side in the early 

stages, sutures of later portion depressed, not limbate; 

wall smooth, distinctly perforate; aperture a low opening, extending from near the periphery along the ventral side beneath the somewhat flattened lip of the ventral margin of the chamber. Diameter of t^pe 0.75 

mm., thickness 0.30 mm.

The types of this species are from the Eocene Tuscahoma sand, railroad cut, 1 mile north of Ozark, Ala. 

The species has not previously been recorded from the 

Paleocene, but rather typical specimens occur at a number of localities.

Porters Creek clay. Tennessee, Hardeman County (IP). 

Midway formation, lower part. Arkansas, Saline County (21). 

Midway formation. Arkansas, Saline County (26). 

Midway group, Texas, Bastrop County (43) ; Travi* County 

(45).

Valvulineria extensa Cushman and Bermudez 

Plate 23, figures 6-8

Valvulineria extensa Cushman and Bermudez, Cushr>an Lab. 

Foram. Research Contr., vol. 24, p. 72, pi. 12, figs. 7-9, 1948.

Test trochoid, the dorsal side very slightly convex, 

ventral side increasing rapidly in thickness in the adult, 

periphery subacute to slightly rounded;'chambers distinct, numerous, 10 or more in the adult whorl, increasing very gradually in size as addecl, with a ventral lobe 

over the umbilical area in the early stages becoming 

high and extending outward in the later port: on; sutures distinct, slightly depressed, curved; wall smooth, 

distinctly perforate; aperture extending from the 

periphery along the ventral margin of the last-formed 

chamber. Length 0.45-0.50 mm.; breadth 0.35-0.40 

mm.; thickness 0.28-0.32 mm.

The typ.es are from the Paleocene Madruga formation, under highway bridge, Central San J ntonio, 

Madruga, Habana Province, Cuba.

This species differs from V. herricki (Hadley) in the 

larger number of chambers, more concave periphery, 

and more open umblical region.
ROTALIIDAE 51

Valvulineria madrugaensis Cushman and Bermudez 

Plate 23, figure 5

Valvulineria madrugaensis Cushman and Bermudez, Cushman 

Lab. Foram. Research Contr., vol. 24, p. 73, pi. 12, fig. 10, 

1948.

Test trochoid, with the dorsal side much less convex 

than the ventral, periphery rounded, becoming more 

acute in the later portion; chambers distinct in the 

later portion, slightly inflated, numerous, earlier ones 

indistinct, not inflated, increasing very gradually in 

size as added, with a narrow, ventral lobe over the umbilical area; sutures distinct in the later portion and 

slightly depressed, earlier ones indistinct; wall smooth; 

aperture extending from the periphery along the ventral margin of the last-formed chamber. Length of 

holotype 0.65 mm.; breadth 0.50 mm.; thickness 0.35 mm.

The types are from the Paleocene Madruga formation, under highway bridge, Central San Antonio, Madruga, Habana Province, Cuba.

This species differs from V. eaetensa Cushman and 

Bermudez in the somewhat larger size, tendency for the 

last-formed chambers to not reach the periphery on the 

dorsal side, and the less projecting ventral lobe.

Valvulineria insueta Cushman and Bermudez 

Plate 23, figures 2-4

Valvulineria insueta Cushman and Bermudez, Cushman Lab. 

Foram. Research Contr., vol. 24, p. 85, pi. 14, figs. 7-9, 1948.

Test trochoid, biconvex, dorsal side nearly flat, ventral side strongly convex, slightly umbilicate, periphery broadly rounded; chambers few, 4 or 5 in the 

adult whorl, inflated, increasing very gradually in size 

as added; sutures distinct, depressed, curved; wall 

smooth; aperture at the ventral margin of the lastformed chamber with a distinct lip, partially covering 

the umbilicus. Length of holotype 0.55 mm.; breadth 

0.47 mm.; thickness 0.32 mm.

The types are from the Paleocene Madruga formation, under highway bridge on Central San Antonio, 

Madruga, Habana Province, Cuba.

This species resembles V. advena Cushman and Siegfus, but differs in the slightly larger number of chambers, the last-formed one making up much less of the 

surface, and the much less prominent lip.

Genus GYROIDINA D'Orbigny, 1826

Gyroidina subangulata (Flummer) Cushman

Plate 14, figures 14, 15

Rotalia soldanii (D'Orbigny var. subangulata Plummer, Texas

Univ. Bull. 2644, p. 154, pi. 12, fig. 1, 1927. 

Gyroidina subangulata Cushman, Cushman Lab. Foram. Research Contr., vol. 16, p. 71, pi. 12, fig. 7, 1940. 

Cushman and Renz, idem, vol. 18, p. 11, pi. 2, fig. 18, 1942. 

Kline, Mississippi Geol..Survey Bull. 53, p. 53, pi. 5, figs. 13-

15, 1943.

Applin and Jordan, Jour. Paleontology, vol. 19, p. 132 (list), 

1945.

Cushman and Todd, Cushman Lab. Foram. Research Contr., 

vol. 22, p. 62, pi. 11, figs. 2-4, 1946.

Test almost plano-convex, the dorsal side being flat or faintly 

convex, the ventral side very strongly convex, composed of about 

two convolutions; peripheral margin bluntly angular; chambers 

8-9 in final whorl; sutures slightly depressed between the last 

two or three chambers on both sides and around the small umbilical excavation, otherwise plain or faintly elevated, moderately oblique dorsally and radiate ventrally; shell wall very 

finely punctate, very smooth, glistening; aperture a long narrow 

slit at the base of a broad septal face extending from a point 

below the periphery almost into the umbilicus. Diameter up to 

0.4mm.; usually less.—Plummer.

The types are from the Paleocene, shallow ditch at 

road corner southeast of new Corsicana reservoir on 

the road to Mildred, Navarre County, Tex.

Porters Creek formation, Matthews Landing marl member.

Alabama, Butler County (3). 

Clayton formation, Chalybeate limestone member. Alabama,

Sumter County (9).

Pine Barren member. Alabama, Butler County (12). 

Midway formation, lower part. Arkansas, Saline County (19,

19A, 20, 22). 

Midway formation, upper part. Arkansas, Pulaski County (24,

25). 

Midway formation. Arkansas, Saline County (26) ; Pulaski

County (27) ; Clark County (30). 

Midway group. Texas, Limestone County (39, 42) ; Navarro

County (40); Bastrop County (43,49) ; Caldwell County (44) ;

Travis County (46). 

Wills Point formation. Texas, Travis County (33) ; Caldwell

County (34). 

Mexia member. Texas, Limestone County (36).

Gyroidina ae^uilateralis (Plummer) Cushman 

Plate 14, figures 16, 17

Rotalia aequilateralis Plummer, Texas Univ. Bull. 2644, p. 155, 

pi. 12, fig. 3, 1927.

Gyroidina aequilateralis Cushman, Cushman Lab. Foram. Research Contr., vol. 20, p. 45, pi. 7, fig. 24,1944.

Gyroidina subangulata Cushman and Todd (not Plummer), idem, 

vol. 18, p. 40, pi. 7, figs. 11,12, 1942.

Test almost equally biconvex, composed of about two and onehalf convolutions that increase very slowly in width; peripheral 

margin narrowly rounded, faintly lobate in the last-formed 

portion of the test; chambers compact, numerous, about 10 in 

final whorl; dorsal sutures distinct, narrow, tapering bands 

without elevation, strongly curved with a slight angulation, but 

not oblique, depressed gently between last two or three chambers 

only; ventral sutures elevated most markedly around the small 

umbilicus and tapering radially toward the margin, dark on tests 

filled with mineral matter; aperture a very narrow slit at base 

of septal face and bearing a very narrow extended lip. Diameter up to 0.4 mm.; usually less.—Plummer.

The types are from the Paleocene, shallow ditch at 

road corner southeast of the new Corsicana reservoir, or 

road to Mildred, Navarro County, Tex.

Porters Creek formation, Matthews Landing marl member. 

Alabama, Wilcox County (1, 2, 4-6) ; Butler County (3); 

Choctaw County (7).

Naheola formation. Coal Bluff marl member. Alabama, Wilcox County (8).

Clayton formation, Pine Barren member. Alabama, Butler 

County (12).
52 PALEOCENE FORAMIMIFERA OF GULF COASTAL REGION

Midway formation, lower part. Arkansas, Saline County (22). 

Midway group. Texas, Bastrop County (49). 

Kincaid formation. Texas, Hunt County (37) ; Bastrop County 

(38).

Gyroidina madrugaensis Cushman and Bermudez 

Plate 24, figures 4-6

Gyroidina madrugaensis Cushman and Bermudez, Cushman Lab. 

Foram. Research Contr., vol. 24, p. 85, pi. 14, figs. 10-12, 

1948.

Test small, about as broad as long, thick, ventral side 

slightly umbilicate, dorsal side depressed in the center, 

periphery broadly rounded; chambers distinct, somewhat inflated, about six in the final whorl, increasing 

very gradually in size as added, on the dorsal side with 

a lobular extension, partially filling the central depression; sutures distinct, slightly depressed, ventrally 

somewhat sinuate, dorsally slightly curved; wall 

smooth; aperture a low opening at the ventral margin 

of the last-formed chamber extending from just below 

the periphery nearly to the umbilical area. Length 

0.30-0.35 mm.; breadth 0.25-0.28 mm.; thickness 0.25 

mm.

The types are from the Paleocene Madruga formation, under highway bridge 011 Central San Antonio, 

Madruga, Habana Province, Cuba.

This species differs from Gr. subangulata (Plummer) 

in the fewer chambers to the w^horl and very depressed 

dorsal side with an extension of the chambers over this 

area.

Genus EPONIDES Montfort, 1808

Eponides elevatus (Plummer) Cushman and Renz 

Plate 14, figures 18, 19

Tmncatulina elevata Plummer, Texas Univ. Bull. 2644, p. 142,

pi. 11, fig. 1, 1927.

Eponides ele-vata Cushman and Renz, Cushman Lab. Foram. Research Contr., vol. 18, p. 11, pi. 3, fig. 1,1942.

Test subconical, the dorsal face being highly elevated and the 

ventral face gently convex; periphery bluntly angular; chambers usually 7 in final convolution; dorsal sutures moderately 

oblique, almost straight, strongly elevated; ventral sutures less 

elevated, radiates from a distinct, thick ridge of shell matter 

surrounding a small excavated umbilicus; shell wall coarsely 

punctate; aperture a small, arched slit near the periphery. 

Diameter up to 0.35 mm.—Plummer.

The types are from the Paleocene, exposure along a 

small branch about % mile northwest of Tehuacana, 

and 0.2 of a mile north of Tehuacana-Waco road at first 

road turning north, Limestone County, Tex.

Porters Creek clay. Tennessee, Hardeman County (13. 14). 

Midway formation, lower part. Arkansas, Saline County (21). 

Midway group. Texas, Hunt County (41) ; Bastrop County (43,

49) ; Travis County (46). 

Kincaid formation. Texas, Hunt County (37) ; Bastrop

County (38).

Eponides lotus (Schwager) Cushman and Ponton 

Plate 14, figure 21

Pulvinulina lotus Schwager, Palaeontographica, vol. 30, Pal. 

Theil, p. 132, pi. 28 (5), fig. 9, 1883.

Eponides lotus Cushman and Ponton, Cushman Lab. Foram. Research Contr., vol. 8, p. 71, pi. 9, fig. S, 1932.

Glaessner, Problems of Paleontology, Moscow Univ., vols. 

2-3, p. 379, pi. 3, fig. 26, 1937.

Bermudez, Soc. cubana hist. nat. Mem., vol. 12, p. 7, 1938.

Cushman and Garrett, Cushman Lab. Forsrn. Research 

Contr., vol. 15, p. 85, pi. 15, figs. 3-6,1939.

Israelsky, Sixth Pacific Sci. Congress, Proc., p. 578, pi. 5, 

figs. 1-4; pi. 6, fig. 1, 1939.

Cushman and Todd, Cushman Lab. Foram. Research Contr., 

vol. 18, p. 40, pi. 7, figs. 13, 14, 1942.

Thalmann, Stanford Univ. Publ., Univ. Ser., Geol. Sci., vol. 

3, no. 1, p. 13 (list), 1942.

Curran, Am. Assoc. Petroleum Geologists Bull., vol. 27, p. 

1379 (list), 1943.

Cushman, Am. Jour. Sci., vol. 242, p. 13, pi. 2, fiV?s. 5, 6, 1944; 

Cushman Lab. Foram. Research Contr., vol. 20, p. 26; p. 

46, pi. 7, fig. 26, 1944.

It seems very difficult to distinguish between this 

species and E. elevatiis. In this material the specimens 

with the sutures generally flush with the surface or 

slightly depressed and with a generally smooth test 

have been placed in E. lotus.

Porters Creek formation, Matthews Landing in^rl member.

Alabama, Wilcox County (1, 2, 6) ; Choctaw County (7). 

Naheola formation Coal Bluff marl member. Alabama, Wilcox

County (8). 

Clayton formation, Pine Barren member. Alabama, Wilcox

County (11) ; Butler County (12). 

Midway formation, lower part. Arkansas, Saline County (18,

19).

Midway formation. Arkansas, Pulaski County (2°). 

Midway group. Texas, Hunt County (41) ; Travis County (45).

Eponides plummerae Cushman 

Plate 14, figures 20, 22

Eponides plummerae Cushman, Cushman Lab. Foram. Research

Contr., vol. 24, p. 44, pi. 8, fig. 9, 1948. 

Truncatulina tenera Plummer (not H. B. Brady), Texas Univ.

Bull. 2644, p. 146, pi. 9, fig. 5, 1927. 

Eponides cf. E. tenera Kline, Mississippi Geol. Survey Bull. 53, p.

53, pi. 5, figs. 16-18, 1943. 

EponMes cf. E. liaidingerii Cushman and Todd (not D'Orbigny),

Cushman Lab. Foram. Research Contr., vol. 22, p. 62, pi. 11,

figs. 5, 6, 1946. 

Eponides sp. Cushman, idem, vol. 16, p. 71, pi. 12, fig. 8, 1940.

Test small, biconvex, periphery subacute; chambers 

distinct but not inflated, usually six in the adult whorl, 

increasing very uniformly in size as added; sutures distinct but not depressed except very slightly on the ventral side, 011 the dorsal side somewhat oblique, on the 

ventral side nearly radial, very slightly curbed; wall 

smooth, polished; aperture elongate, narrow, 011 the 

ventral side of the last-formed chamber running from 

just below the periphery almost to the umbilical area, 

often with a very slight overhanging lip. Diameter 

0.32 to 0.37 mm., thickness 0.22 to 0.25 mm.

The types' are from the Paleocene, exposure in creek 

bed on northeast-southwest country road, 1.1 mi. due 

east of Honest Ridge School, and 2.5 mi. dua, north of 

the cross roads in Thelma, Limestone County, Tex. 

(39).
ROTALIIDAE' 53

This species differs from E. tenera (H. B. Brady) in 

its more convex dorsal side and more oblique sutures 

on the dorsal side.

Clayton formation, Chalyeate limestone member. Alabama,

Sumter County (9). 

Midway formation, upper part. Arkansas, Pulaski County (24,

25). 

Midway formation. Arkansas, Pulaski County (27) ; Clark

County (30). 

Midway group. Texas, Limestone County (39) ; Navarro

County (40) ; Caldwell County (44). 

Wills Point formation. Texas, Travis County (33).

Eponides vaubelleui (Van den Bold) Cushman and Bermudez 

Plate 20, figures 11-13; plate 23, figure 10

fdbicides varibelleni Van den Bold, Thesis Univ. Utrecht, Amsterdam, p. 125, pi. 18, figs. 8a-c, 1946.

Eponides vanbelleni Cushman and Bermudez, Cushman Lab. 

Foram. Research Contr., vol. 24, p. 71, pi. 12, figs. 1-3, 

1948.

Test trochoid, unequally biconvex, rotaliform. Dorsal side 

only slightly elevated, ventral side strongly convex. Chambers 

numerous in the last formed whorl (at least 14), but hardly 

visible, owing to the ornamentation; sutures very indistinct 

except dorsally in the few last-formed chambers. There they 

are slightly depressed. Ventral side strongly ornamented with 

numerous fiat papillae, vermicular in the umbilical region, diminishing in size and becoming nodular towards the periphery. 

Dorsal side less strongly ornamented in the same way. Aperture a short arched slit at the periphery and the ventral side 

and probably extending on the dorsal side. But as this could 

not be seen very well owing to the rather poor state of preservation of the tests, the reference to Cibitides is questionable.— 

Van den Bold.

Diameter 0.65 mm., height 0.40 mm.

This species was described from the lower Eocene of 

Guatemala and British Honduras. Specimens apparently identical occurred in the Paleocene Madruga formation, under highway bridge on Central San Antonio, 

Madruga, Habana Province, Cuba, and indicate that 

from the position of the aperture the species belongs in 

Eponides.

Eponides gratiosus Cushman and Bermudez 

Plate 23, figure 9

Eponides gratiosus Cushman and Bermudez, Cushman Lab. 

Foram. Research Contr., vol. 24, p. 71, pi. 11, fig. 14, 1948.

Test trochoid, unequally biconvex, the ventral side 

very strongly convex, dorsal side much less so, periphery 

acute; chambers fairly distinct on the ventral side, those 

of the last-formed whorl on the dorsal side fairly distinct, earlier whorls obscured by the surface ornamentation; sutures mostly indistinct, ventrally slightly 

curved, dorsally strongly oblique; wall of the ventral 

side somewhat roughened but not definitely papillate, 

dorsal side finely but distinctly papillate except over the 

last few chambers, which are fairly smooth; aperture 

ventral, a low elongate opening at the margin of the 

last-formed chamber. Diameter of holotype 0.63 mm.'; 

thickness 0.38 mm.

The types are from the Paleocene Madruga formation, under highway bridge on Central San Antonio, 

Madruga, Habana Province, Cuba.

This species differs from E. vanbelleni (Van den 

Bold) in the acute periphery, smoother ventral side, and 

nearly straight ventral sides in edge view. 

Genus PARRELLA Finlay, 1939 

Parrella expansa Toulmin 

Plate 16, figures 11, 12

Parrella expansa Toulmin, Jour. Paleontology, vol. 15, p. 604,

text figs. 3, 4F, 4G, 1941. 

Kline, Mississippi Geol. Survey Bull. 53, p. 54, pi. 5, figs. 19,

20, 1943. 

Truncatulina culler Plummer (not Parker and Jones), Texas

Univ. Bull. 2644, p. 147, pi. 10, fig. 1; pi. 15, fig. 2, 1927. 

PulvinuUnella culter (Parker and Jones) var. mexicana Cushman (not Cole), Cushman Lab. Foram. Research Contr., 

vol. 16, p. 72, pi. 12, fig. 12,1940.

Pulvinulinella culter (Parker and Jones) var. midwayana Cushman and Todd, idem, vol. 22, p. 63, pi. 11, fig. 12,1946. 

Test trochiform, close coiled, subcircular in outline, ventral 

side strongly convex, dorsal side nearly flat with earliest coils 

gently elevated, periphery sharply acute with a broad thin flange; 

all chambers visible on dorsal side, with flange of all coils forming a limbate whorl suture, only chambers in final convolution 

visible ventrally, eight or nine chambers in the final whorl; 

dorsal sutures broad, .oblique, slightly curved, flush or slightly 

elevated, ventral sutures limbate between early chambers of the 

final whorl, generally depressed between the last two or three 

chambers, radiating from a small umbilical filing and curving 

backward near the junction with the peripheral flange; wall 

smooth, perforate; aperture consisting of two portions, a narrow, elliptical opening extending from the base of the final chamber near the periphery obliquely across the apertural face, at 

an angle of 45 degrees or more to the peripheral plane, and a 

very narrow slit at base of the septal face extending ventrally 

toward the umbilical area from the dorsal end of the oblique 

opening. Average diameter 0.5 mm.—Touunin.

The types are from the Paleocene Wills Point formation, abandoned clay pit in west edge of Mexia, Tex.

Porters Creek clay. Mississippi, Kemper County (17). 

Midway formation, upper part. Arkansas, Pulaski County (25). 

Midway formation. Arkansas, Pulaski County (27) ; Clark

County (30). 

Midway group. Texas, Limestone County (39, 42) ; Navarre

County (40) ; Caldwell County (44). 

Wills Point formation. Texas, Williamson County (35). 

Mexia member. Texas, Limestone County (36).

Parrella macneili Cushman 

Plate 16, figures 13, 14

Parrella macneili Cushman, Cushman Lab. Foram. Researcl 

Contr., vol. 20, p. 47, pi. 7, figs. 30, 31, 1944.

Test small, close coiled, trochoid, consisting of about 

4 chambers in the adult whorl; periphery acute, slightly 

keeled and serrate; chambers fairly distinct, increasing 

rapidly in size as added, the last-formed one strongly 

inflated on the ventral side, making up nearly half the 

surface of the test; sutures indistinct, tangential, little 

if at all depressed; wall slightly papillate in the umbonal region of the ventral side, on the dorsal sid? 

papillate along the sutures; aperture fairly large, ven-
54 PALEOCENE FORAMINIFERA OF GULF COASTAL REGION

tral, with a rounded, somewhat triangular lip curved on 

the outer side making a rounded indentation into the 

ventral face. Diameter 0.35 to 0.45 mm., thickness 

0.10 mm.

The types are from the Paleocene Coal Bluff marl 

member of the Naheola formation, creek bottom, just 

west of store at Caledonia, about y± mile south of center • 

of sec. 29, T. 11 N., R. 10 E., Wilcox County, Ala. (8). 

It is known only from the type locality.

Genus COLEITES Plummer, 1934

Coleites reticulosus (Plummer) Plummer

Plate 15, figures 1-5

Pulmnulina reticulosa Plummer, Texas Univ. Bull. 2644, p. 152,

pi. 12, fig. 5, 1927. 

Coleites reticulosus Plummer, Am. Midland Naturalist, vol. 15,

p. 606, pi. 24, figs. 5-9, 1934. 

Cushman and Garrett, Cushman Lab. Foram. Research

Contr., vol. 15, p. 87, pi. 15, figs. 14-20, 1939. 

Cushman, idem, vol. 16, p. 71, pi. 12, fig. 20, 1940. 

Cushman and Bermudez, idem, vol. 24, p. 81, pL 13, figs. 1-4, 

1948.

Test broadly elongate, strongly compressed, flat or gently 

convex on the dorsal side, more convex on the ventral side, 

coarsely and irregularly reticulate on both sides except over the 

septal face; periphery very sharp and bounded by a ragged 

flange; early chambers tightly coiled through two or more convolutions, very narrow, curved, gradually lengthening, 7 in the 

final whorl; fully mature chambers short, broad, and rapidly 

becoming almost rectilinear; foramina connected by a stout tube, 

which in the final chamber extends to the aperture; early sutures 

completely obscured by the coarse reticulation of the test but 

clearly visible when test is mounted in liquid, strongly curved 

dorsally but somewhat less curved ventrally, later sutures very 

slightly constricted and not quite parallel; aperture of very 

young tests an elliptical opening on the ventral side of the 

peripheral angle above the base of the septal face, but with 

growth of the test the opening becomes more narrow and longer 

and develops a faint blunt tooth on its ventral edge. Length 

of most mature test observed 0.85 mm.; breadth 0.52 mm.; thickness 0.21 mm.—Plummer, 1934.

Porters Creek formation, Matthews Landing marl member. Alabama, Butler County (3). 

Midway formation. Arkansas, Clark County (30).

Coleites pasionensis Cushman and Bermudez 

Plate 24, figure 7

Coleites pasionensis Cushman and Bermudez, Cushman Lab. 

Foram. Research Contr., vol. 24, p. 84, pi. 14, fig. 6,1948.

Test only slightly longer than broad, thick, the dorsal 

side strongly convex, ventral side less convex, the adult 

portion tending to uncoil and much more compressed, 

periphery subacute, in the adult with a raised and thickened border on the dorsal side; chambers obscured by 

the surface ornamentation except the last two or three; 

sutures obscure, except the last two or three which are 

strongly raised and thickened on the dorsal side, only 

slightly so on the ventral side, only slightly curved; 

wall ornamented with very coarse reticulations on the 

early coiled portion, becoming much less coarse in the 

later uncoiled portion; aperture an elongate slit on the

ventral margin of the last-formed chamber. Length 

of holotype 1.12 mm.; breadth 0.85 mm.; thickness 

0.38 mm.

The types are from the lower Eocene, Rio Pasion, 

Peten Province, Guatemala.

This species differs from G. reticulosus (Plummer) 

in the very coarse reticulations of the early portion, the 

raised and thickened sutures of the dorsal side, and the 

more irregular periphery.

Coleites guatemalensis Cushman and Bern udez 

Plate 24, figures 8-10

Coleites guatemalensis Cushman and Bermudez, Cushman Lab. 

Foram. Research Contr., vol. 24, p. 84, pi. 14, f gs. 3-5, 1948.

Test strongly compressed, early portion trochoid and 

slightly thicker than the later portion, adult portion uncoiling but somewhat curved, periphery subacute, the 

early portion irregularly dentate, later slightly lobulate; chambers of the early trochoid portion rather indistinct, in the later uncoiled portion mucl more distinct; sutures distinct except in the early portion, 

strongly curved, depressed on the ventral side and 

slightly raised and limbate on the dorsal side; wall of 

the early coiled portion finely reticulate, tiH uncoiled 

portion nearly smooth; aperture in the adult elongate 

in the middle of the terminal border, sligl tly on the 

ventral side. Length of holotype 1.00 mm.; breadth 

0.70 mm.; thickness 0.10 mm.

The types are from the lower Eocene, B.io Pasion, 

Peten Province, Guatemala.

This species differs from C. dbuillotensis Cushman 

and Bermudez in the broader form, smoother test, and 

slightly raised, limbate sutures on the dorsal side.

Coleites abuillotensis Cushman and Bermudez 

Plate 24, figures 11, 12

Coleites abuillotensis Cushman and Bermudez, Cushman Lab. 

Foram. Research Contr., vol. 24, p. 83, pi. 14, figs. 1, 2, 1948.

Test elongate, strongly compressed, nearly flat, 3 

times as long as broad, greatest breadth toward the central portion, slightly convex ventrally, peripheral margin subacute; chambers numerous, 14 to 15 comprising 

the entire test, early portion involute, formed by 7 to 8 

chambers, each connected with the preceding by the 

apertural openings; sutures slightly depressed, strongly 

curved on both sides of the test; wall distinctly reticulate; aperture a longitudinal slit at the middle of 

the peripheral edge of the last-formed chamber. 

Length of holotype 1.11 mm.; breadth 0.55 mm.; 

thickness 0.12 mm.

The types are from the lower Eocene, Abuillot Kiver, 

Central Plain, Haiti, and the species is known only from 

the type locality.

This species differs from O. reticulosus (Plummer) in 

the less convex ventral side, more compressed form, and 

less acute peripheral margin.
ROTALIIDAE' 55

Genus ROTALIA Lamarck, 1804

Rotalia havanensis Cushman and Bermudez

Plate 15, figure 11

Rotalia havanensis Cushman and Bermudez Cushman Lab. 

Foram. Research Contr., vol. 23, p. 24, pi. 5, fig, 5, 1947.

Test rather small, strongly and nearly evenly biconvex, periphery rounded; chambers fairly distinct, very 

slightly if at all inflated, 9 or 10 in the adult whorl, increasing very gradually in size as added; sutures fairly 

distinct, very slightly curved but nearly radial, little 

if at all depressed; wall fairly smooth, except near the 

aperture on the ventral side where it is distinctly papillate, on the dorsal side with a series of fine linear papillae near the periphery and on the ventral side with a 

series of short oblique costae on each chamber at the 

basal peripheral angle, umbonal boss strongly developed ; aperture a low opening just below the periphery 

on the ventral side of the last-formed chamber. 

Length of holotype 0.60 mm., breadth 0.50 mm., thickness 0.30 mm.

The types are from the Paleocene, water well, 200 

meters north of Cerro Avenue on Rancho Boyeroa 

road, Habana Province, Cuba, and the species is frequent in samples of Paleocene age from the vicinity of 

Havana.

Rotalia madmgaeusis Cushman and Bermudez 

Plate 15, figure 12

Rotalia madrugaensis Cushman and Bermudez, Cushman Lab. 

Foram. Research Contr., vol. 23, p. 24, pi. 5, fig. 4, 1947.

Test of medium size, strongly biconvex but with the 

central areas of both sides slightly depressed, periphery 

broadly rounded; chambers strongly inflated in the 

later portion, about 10 in the adult whorl, increasing in 

size rather rapidly as added in the last portion, earlier 

ones indistinct; sutures indistinct except in the last 

whorl, where they are radial on both sides and become 

increasingly depressed; wall strongly ornamented with 

longitudinal costae confined to each chamber, those of 

the earlier portion of the final whorl broken up into 

very short ones, almost papillate, ventral side with 

several large papillae in the umbilical area; aperture a 

low opening on the ventral edge of the last-formed 

chamber, toward the periphery. Length of holotype 

0.85 mm., breadth 0.65 mm., thickness 0.37 mm.

The types are from the Paleocene, under bridge in the 

highway, Central San Antonio, Madruga, Habana Province, Cuba, and the species occurs in numerous samples 

of Paleocene age from the vicinity of Madruga.

Genus EPISTOMINA Terquem, 1883

Epistomina eoceuica Cushman and M. A. Hauna

Plate 15, figures 13, 14

Epistomina eocenica Cushman and M. A. Hanna, San Diego 

Soc. Nat. History Trans., vol. 5, no. 4, p. 53, pi. 5, figs. 4, 

5, 1927.

Cushman and Schenck, California Univ., Dept. Geol. Sci.,

Bull., vol. 17, p. 313, pi. 44, fig. 9, 1928. 

Weinzierl and Applin, Jour. Paleontology, vol. 3, p. 407,1929. 

Cushman and McMasters, idem, vol. 10, p. 515, pi. 76, fig. 5,

1936. 

Cushman and Frizzell, Cushman Lab. Foram. Research

Contr., vol. 19, p. 87, pi. 15, figs. 9, 10, 1943. 

Martin, Stanford Univ. Publ., Univ. Ser., Geol. Sci., vol. 3,

no. 3, p. 10 (list), 1943. 

Cushman and Simonson, Jour. Paleontology, vol. 18, p. 202,

pi. 34, fig. 6, 1944.

Pulvinulina partschiana Plummer (not D'Orbigny), Texas Univ. 

Bull. 2644, p. 153, pi. 11, fig. 5, 1927.

Test trochoid, biconvex, periphery subacute, ventral side more 

convex than the dorsal, composed of about three coils, the adult 

coil having seven chambers, which are distinct due to the darker 

appearance of the chamber walls between the sutures which 

are raised and very white and confluent with the raised border; 

aperture a narrow slit near the inner margin of the chamber on 

the ventral side and a supplementary one consisting of a longnarrow slit, parallel to and just below the periphery on the 

ventral side; sutures on the ventral side coalescing in an umbilical thickening. Diameter 0.50 mm.—Cushman and Hanna. 

Wills Point formation, Mexia member. Texas, Limestone

County (36). 

Midway group. Texas, Limestone County (39, 42) ; Navarre

County (40).

Genus SIPHONINA Reuss, 1850

Siphouiua prima Flummer

Plate 15, figures 7-9

Siphomna prima Plummer, Texas Univ. Bull. 2644, p. 148, pi.

12, fig. 4, 1927. 

Cushman, U. S. Nat. Mus. Proc., vol. 72, art. 20, p. 2, pi. 2,

fig. 4, 1927.

Plummer, Texas Univ. Bull. 3201, pp. 54, 62 (lists), 1933. 

Jennings, Bull. Am. Paleontology, vol. 23, no. 78, p. 33, pi,

4, fig. 3, 1936. 

Cushman, Cushman Lab. Foram. Research Contr., vol. 16, p.

71, pi. 12, fig. 10, 1940. 

Cushman and Todd, idem, vol. 18, p. 40, pi. 7, figs. 16, 17,

1942. 

Kline, Mississippi Geol. Survey Bull. 53, p. 55, pi. 5, figs.

21, 22, 1943. 

Cooper, Jour. Paleontology, vol. 18, p. 353, pi. 55, figs. 7-S,

1944. 

- Cushman and Todd, Cushman Lab. Foram. Research Contr.,

vol. 22, p. 62, pi. 11, figs. 7, 8', 1946. 

Cushman, U. S. Geol. Survey Prof. Paper 206, p. 143, pL

59, figs. 3-5, 1946.

Test small, nearly circular, about equally biconvex 

but much compressed; periphery angled, sharply acute 

and delicately serrate, very slightly lobate; chambers 

usually 5 in the last-formed volution, very slightly 

inflated on the ventral side; sutures distinct, obliquely 

curved, marked by the serrate edges of the chambers 

of the dorsal side, not depressed, on the ventral side 

more nearly radial, very slightly curved, somewhat depressed ; wall smooth, distinctly and somewhat coarsely 

perforate; aperture a small, narrowly elliptical opering on the ventral side close to the periphery, the elorgate axis in the axis of coiling, without a definite necK 

Diameter up to 0.25 mm., thickness 0.12 mm.
56 PALEO'CENE FORAMINIFERA OF GULF COASTAL REGION

The types are from the Paleocene, clay pit, Mexia 

Brick Works, about 1 mile west of the town of Mexia, 

Limestone County, Tex.

Porters Creek formation, Matthews Landing marl member. 

Alabama, Butler County (3) ; Wilcox County (5, 6) ; Choctaw County (7).

Clayton formation, Chalybeate limestone member. Alabama, 

Sumter County (9). Mississippi, Noxubee County (16).

Midway formation, lower part. Arkansas, Saline County (19, 

19A-21) ; Pulaski County (23)*

Midway formation, upper part. Arkansas, Pulaski County (24, 

25).

Midway formation. Arkansas, Saline County (26) ; Pulaski 

County (27) ; Clark County (30).

Midway group. Texas, Caldwell County (44).

Kincaid formation. Texas, Hunt County (37) ; Bastrop County (38).

Siphonina wilcoxensis Cushman

Plate 15, figure 10

Siplwnina ^l>ilcoxensis Cushman, U. S. Nat. Mus. Proc., vol. 72,

art. 20, p. 3, pi. 2, figs. 1-3, 1927. 

Cushman and Ponton, Cushman Lab. Foram. Research

Contr., vol. 8, p. 70, pi. 9, fig. 7,1932. 

Cushman and Garrett, idem, vol. 15, p. 86, pi. 15, figs. 7-9,

1939. 

Israelsky, Sixth Pacific Sci. Congress Proc., p. 578, pi. 7,

fig. 3, 1939. 

Woulmin, Jour. Paleontology, vol. 15, p. 605, pi. 81, figs.

15, 16, 1941. 

Cushman, Am. Jour. Sci., vol. 242, p. 14, pi. 2, figs. 3, 4,

1944; Cushman Lab. Foram. Research Contr., vol. 20,

p. 27; p. 46, pi. 7, fig. 27, 1944.

Test small, nearly circular, biconvex but slightly 

more inflated on the ventral side, compressed, periphery angled, sharply acute and delicately serrate, 

slightly lobulate; chambers usually six in the lastformed volution, slightly inflated on the ventral side; 

sutures on the dorsal side somewhat indistinct, strongly 

oblique, slightly curved, somewhat marked by the serrate edges of the chambers, not depressed, on the ventral 

side very nearly radial, distinctly depressed; wall 

smooth, distinctly and coarsely perforate; aperture a 

small, elliptical opening on the ventral side close to 

the periphery, with a distinct lip but with the neck only 

slightly developed or wanting. Diameter up'to 0.30 

mm., thickness 0.16 mm.

The types are from the Eocene, Nanafalia formation, 

upper portion of Nanaf alia Bluff, Tombigbee River, 

Ala.

Naheola formation, Coal Bluff marl member. Alabama, Wilcox County (8).

Clayton formation, Pine Barren member. Alabama, Butler 

County (12).

Midway formation. Arkansas, Pulaski County (28).

Genus CANCRIS Montfort, 1808

Cancris mauryae Cushman and Renz

Plate 15, figure 6

Cancris mauryae Cushman and Renz, Cushman Lab. Foram.

Research Contr., vol. 18, p. 11, pi. 2, fig. 17, 1942. 

Cushman and Todd, idem, vol. 18, p. 86, pi. 22, fig. 8, 1942.

Test nearly equally biconvex, periphery subacute and 

slightly keeled; chambers distinct, about 10 in the final 

whorl, increasing rather rapidly in size as added, the 

last 2 or 3 making up more than half of the test, slightly 

inflated; sutures distinct, slightly limbate and raised, 

especially on the dorsal side; wall smooth; aperture a 

low slit below the ventral extension of the last-formed 

chambers. Length of holotype 0.70 mm., breadth 0.50 

mm., thickness 0.35 mm.

The types are from the Paleocene Soldado Rock formation, Soldado Rock, Trinidad, B. W. I., and the 

species has not been recorded elsewhere.

Family AMPHTSTEGINIDAE

Genus ASTERIGERINA D'Orbigny, 1839

Asterigerina primaria Plummer

Plate 15, figure 15

Asterigerina primaria Plummer, Texas Univ. Bull. 2644, p. 157,

pi. 12, fig. 8, 1927. 

Toulmin, Jour. Paleontology, vol. 15, p. 606, p1 . 81, fig. 22,

1941. 

Cushman and Todcl, Cushman Lab. Foram. Research Contr.,

vol. 18, p. 41, pi. 7, fig. 18, 1942. 

Cushman, idem, vol. 20, p. 46, pi. 7, fig. 28,1944.

Test round, obtusely conical, the dorsal side being distinctly 

elevated and the ventral side almost fiat; peripheral margin 

sharply keeled, lobate; number of whorls about two in welldeveloped tests; chambers 5-6 in final convolution, smooth, 

conspicuously punctate; dorsal sutures moderately curved, 

faintly elevated, produced peripherally to form t^e marginal 

keel, rather strongly elevated in the early portion of the spire; 

ventral sutures strongly curved but partly masked by the supplementary chambers formed by the umbilical lotes over the 

successive apertures of the last whorl; aperture a strongly 

arched slit directed toward the irregularly pustulate umbilicus. 

Diameter up to 0.3 mm.—Plummer.

The types are from the Paleocene, exposure in base 

of high bluff on west side of Colorado River between 

the Travis-Bastrop county line and the mouth of Dry 

Creek (Bastrop quadrangle), Bastrop County, Tex.

Porters Creek formation, Matthews Landing mrrl member. 

Alabama, Wilcox County (2) ; Choctaw County (7).

Naheola formation. Coal Bluff marl member. Alabama, Wilcox County (8).

Midway group. Texas, Bastrop County (43).

Family CASSIDULINIDAE

Genus CERATOBULIMINA Toula, 1915

Ceratobulimina perplexa (Plummer) Cushman an-* Harris

Plate 16, figures 1-5

Rotalia perplexa Plummer, Texas Univ. Bull. 2644, j\ 156, pi. 12,

fig. 2, 1927.

Ceratobulimina perplexa Cushman and Harris, Cushman Lab.

Foram. Research Contr., vol. 3, p. 173, pi. 29, fig. 2, 1927.

Plummer, Texas Univ. Bull. 3201, pp. 54, 62 (lists), 1933;

(part), Am. Midland Naturalist, vol. 17, p. 4PO, text figs.

1-4, 1936.

Glaessner, Moscow Univ. Studies in Micropaleontology, vol. 

1, pp. 20, 23, pi. 1, figs. 2, 3; pi. 2, fig. 25, 1937.
CAISISIDTJLINIDAE, 57

Cushman and Todd, Cushman Lab. Foram. Research Contr., 

vol. 18, p. 41, pi. 7, figs. 23, 24,1942.

Cushman, idem, vol. 22, p. 108, pi. 17, figs. 3-5, 1946. 

Test oval, about equally biconvex, considerably compressed; 

peripheral margin broadly rounded, somewhat lobate; chambers smooth, glistening, finely punctate, gently curving, 6 to the 

final whorl; dorsal sutures marked by thick, smooth or very 

slightly elevated, tapering bands that become distinctly angular at their broadest points; ventral sutures depressed, radiate 

from a sunken umbilicus; aperture a conspicuous round opening at the base of the septal face and protected by an arched 

flap that is directed inte the umbilicus. Length up to 0.5 mm.; 

average 0.35 mm.—Plummer.

The types are from the Paleocene, shallow ditch at 

road corner southeast of new Corsicana reservoir on the 

road to Mildred, Navarro County, Tex.

Porters Creek formation, Matthews Landing marl member.

Alabama, Wilcox County (5) ; Choctaw County (7). 

Wills Point formation. Texas, Travis County (33) ; Caldwell County (34).

Mexia member. Texas, Limestone County (36). 

Midway group. Texas, Bastrop County (43).

Genus ALABAMINA Toulmin, 1941

Alabamina wilcoxensis Toulmin

Plate 16, figures 6, 7

Alabamina imlcoseensls Toulmin, Jour. Paleontology, vol. 15, p.

603, pi. 81, figs. 10-14, text figs. 4A-C, 1941. 

Cushman, Cushman Lab. Foram. Research Contr., vol. 24,

p. 14, pi. 2, figs. 18-20, 1948.

Pulvinulina exigiia H. B. Brady var. obtma Plummer (not Burrows and Holland), Texas Univ. Bull. 2644, p. 151, pi. 11, 

fig. 2, 1927.

Pulvmulinella exigua (H. B. Brady) var. obtusa Cushman and 

Ponton (not Burrows and Holland), Cushman Lab. Foram. 

Research Contr., vol. 8, p. 71, pi. 9, fig. 9, 1932. 

PulvinulineUa obtusa (Burrows and Holland) Cushman and

Garrett, idem, vol. 15, p. 87, pi. 15, figs. 12, 13, 1939. 

Cushman and Renz, idem, vol. 18, p. 11, pi. 2, fig. 16, 1942. 

Cushman and Todd, idem, vol. 18, p. 42, pi. 7, figs. 19, 20,

1942.

Cushman, Am. Jour. Sci. vol. 242, p. 14, pi. 2, figs. 7, 8, 1944; 

Cushman Lab. Foram. Research Contr., vol. 20, p. 27, pi. 

4, fig. 32; p. 46, pi. 7, fig. 29,1944. 

Applin and Jordan, Jour. Paleontology, vol. 19, p. 132 (list),

1945.

Cushman and Todd, Ctishman Lab. Foram. Research Contr., 

vol. 21, p. 101, pi. 16, figs. 7, 8, 1945; vol. 22, p. 63, pi. 11, 

figs. 9,10, 1946.

Test troehiform, close coiled, subcircular in outline, unequally 

biconvex or planoconvex, ventral side strongly convex, periphery 

bluntly acute; five of six chambers in the final whorl; wall 

smooth, finely perforate; sutures flush with the surface, on the 

dorsal side almost straight, oblique to the periphery, on the 

ventral side radial; aperture a long narrow opening on the 

ventral side at the base of the septal face, with a supplementary 

indentation extending peripherally from the true aperture, 

formed by a fold in the wall of the test and not opening into the 

interior of the chamber. Diameter up to 0.42 mm.; height up 

to 0.21 mm.—Toulmin.

The types are from the Eocene Wilcox group, railroad cut one mile north of Ozark, Ala.

Paratypes of this species have been compared with 

topotypes of Burrows and Holland's variety and the 

two seem distinct.

Porters Creek clay. Tennessee, Hardeman County (13, 14).

Mississippi, Kemper County (17). 

Porters Greek formation, Matthews Landing marl member.

Alabama, Wilcox County (1, 2, 4-6) ; Butler County (3) ;

Choctaw County (7).

Naheola formation. Coal Bluff marl member. Alabama, Wilcox County (8). 

Clayton formation, Pine Barren member. Alabama, Butler

County (12). 

Midway formation, lower part. Arkansas, Saline County (19A,

22). 

Midway formation, upper part. Arkansas, Pulaski County (24,

25). 

Midway formation. Arkansas, Saline County (26) ; Pulaski

County (27) ; Clark County (29-31). 

Midway group. Texas, Limestone County (39) ; Navarro

County (40) ; Hunt County (41) ; Bastrop County (43) ; Caldwell County (44). 

Wills Point formation. Texas, Travis County (33) ; Caldwell

County (34).

Mexia member. Texas, Limestone County (36). 

Kincaid formation. Texas, Hunt County (37).

Alabamina wilcoxensis Toulmin var. limbata (Plummer) Gushmar 

Plate 16, figure 8

Pulvinulwa exigua, H. B. Brady var. limbata, Plummer, Texas

Univ. Bull. 2644, p. 152, pi. 11, figs. 4a-c, 1927. 

From the type of this species this varietal form differs in having strong sutural elevations on the dorsal side and slight elevations on the ventral side. Diameter up to 0.5 mm.— 

Plummer.

The types of the variety are from the Paleocene, deep 

roadside ditch on steep hill 0.2 of a mile east of road 

corner in north end of town of Commerce on highway?" 

to Paris, Hunt County, Tex. (41).

This variety is evidently related to Alabamina wilcoxensis Toulmin. It is known only from the typ?- 

locality.

Genus PSETTDOPARRELLA Cushman and Ten Dam, 1948

Pseudoparrella cf. P. exigua (H. B. Brady) Cushman

Plate 16, figures 9, 10

Very rare specimens in the Paleocene material are 

different from Alabmnina wtlGoxensis Toulmin and 

may be compared with P. eoeigua with considerable quertion, but not enough well preserved specimens are available to give the full characters. Specimens from the 

Paleocene of Texas have been referred to this species 

by Plummer (Texas Univ. Bull. 2644, p. 150, pi. 11, fig. 

3,1927) and by Kline from the Paleocene of Mississippi 

(Mississippi Geol. Survey Bull. 53, p. 55, pi. 5, figs. 23, 

24,1943).

Porters Creek clay. Tennessee, Hardeman County (13, 14). 

Midway group. Texas, Van Zandt County (47).
58 PALEOCENE FORAMINIFERA OF GULF COASTAL REGION

Pseudoparrella madrugaensis Gushman and Bermudez 

Plate 24, figures 1-3

Pseudoparrella madr.ugaensis Cushman and Bermudez, Gushman Lab. Foram. Research Contr., vol. 24, p. 73, pi. 12, figs. 

11-13, 1948.

Test trochoid, nearly equally biconvex, ventral side 

slightly more convex, periphery acute, slightly keeled; 

chambers distinct, very slightly inflated, about 5 in the 

adult whorl, increasing rather evenly in size as added; 

sutures distinct, dorsally strongly curved, ventrally 

nearly radial; wall smooth, distinct perforate; aperture at the ventral margin of the last-formed chamber 

and extending outward parallel to the periphery, narrow and with a distinct lip. Diameter 0.35-0.40 mm.; 

thickness 0.15-0.18 mm.

The types are from the Paleocene Madruga formation, under highway bridge, Central San Antonio, Habana Province, Cuba.

This species differs from P. obtusa (Burrows and 

Holland) in the definite keel, curved dorsal sutures, and 

less extended aperture.

Genus EPISTOMINOIDES Plummer, 1934 

Epistominoides midwayensis Plummer

Plate 16, figures 15, 16

Epistominoides midivayen-sis Plummer, Am. Midland Naturalist, vol. 15, p. 605, pi. 24, fig. 4, 1934.

Cushman and Todd, Cushman Lab. Foram. Research Contr., 

vol. 18, p. 42, pi. 7, figs. 21, 22, 1942.

Brotzen, Sveriges geol. undersokning, ser. C, no. 451, p. 

36, text fig. 11, 1942.

Cushman, Cushman Lab. Foram. Research Contr., vol. 20, 

p. 47, pi. 8, fig. 1, 1944.

Test elongate, somewhat more than half as broad as long, 

strongly bilaterally unsymmetrical, umbilicate; periphery 

sharply carinate but not distinctly flanged; dorsal face gently 

convex, ventral face somewhat more convex; chambers 7-8 

in the mature whorl, increasing successively at the rate of 

about 1.2, subdivided internally by a thin partition that extends 

from the short, slit-like aperture into each chamber across its 

ventro-anterior angle making by its junction with the ventral 

wall a supplementary ventral suture so close to the anterior 

edge of the chamber and so nearly parallel to it as to be 

somewhat obscure; septal face triangular, width about threequarters the height and marked externally by the junction of 

the inner partition from the anterior end of the apertural slit 

part way across the septal face and parallel to its ventral edge; 

ventral sutures nearly straight and marked by a slight thickening of the ventral wall between each true suture and the 

parallel supplementary suture to form blunt ridges that radiate 

from a small umbilical depression; dorsal sutures narrowly 

elevated, radiating from a broad central boss, and sharply curved 

backward near the periphery; aperture a short elongate slit located on the peripheral angle at the end of the septal face and 

bounded by protruding ventral and dorsal lips. Length of 

holotype 0.55 mm.; breadth 0.32 mm.; thickness 0.27 mm.— 

Plummer.

The types are from the Paleocene, expoure in base 

of high bluff on west side of Colorado River between 

the Travis-Bastrop county line and the mouth of Dry 

Creek (Bastrop quadrangle), Bastrop County, Tex.

This species differs from E. wilcowensis (Cushman 

and Ponton) in its broader chambers, and umbilicate, 

strongly asymmetrical, and more compressed form.

Porters Creek clay. Tennessee, Hardeman County (13).

Porters Creek formation. Matthews Landing mr.rl member. 

Alabama, Butler County (3) ; Wilcox County (4) ; Choctaw 

County (7).

Naheola formation. Coal Bluff marl member. Alabama, Wilcox County (8).

Midway group. Texas, Bastrop County (43).

Epistominoides wilcoxensis (Gushman and Ponton) Plummer 

Plate 16, figures 17, 18

Saracenaria imlcoxensis Cushman and Ponton, Cushman Lab.

Foram. Research Contr., vol. 8, p. 54, pi. 7, fig. 9, 1932. 

Epistominoides wilcoxensis Plummer, Am. Midland Naturalist,

vol. 15, p. 604, pi. 24, figs. 1-3, 1934. 

Glaessner, Moscow Univ. Studies in Micropaleo^tology, vol.

1, p. 21, pi. 2, fig. 22, 1937. 

Cushman, Foraminifera, 3d Ed., Key, pi. 48, figs. 15,16,1940.

Test elongate, sharply trihedral, with septal face nearly as 

broad as ventral and dorsal faces, very nearly bilaterally symmetrical, involute but showing very slightly more of the coil on 

the dorsal side of the consistently left-handed spire; chambers 

6 or 7 to the whorl, increasing successively in length at the 

rate of about 1.4, each one subdivided very unequally by a thin 

inner wall or partition that extends from the dorsrl lip of the 

aperture obliquely across the ventro-anterior angle cf the chamber and is attached to the septal face, to the ventral wall, and 

to the septum on the ventral side of the foramen; sutures flush 

to elevated, radiating from a boss on each side of the test, almost 

straight across both ventral and dorsal faces but curved sharply 

backward near the periphery; umbilical area filled with shell 

matter; aperture a narrow and elongate slit located in the 

peripheral angle of the final chamber at the end of the septal 

face and bounded by protruding ventral and dorsal lips.

—Plummer.

The types are from the Eocene Wilcox group in railroad cut 1 mile north of Ozark, Ala.

Length 0.50 mm., breadth 0.30 mm., thickness 0.25 

mm.

Family CHILOSTOMElllDAE

Genus ALLOMORPHINA Reuss, 1850

Allomorphina paleocenica Cushman

Plate 16, figures 19-22

Allomorphina paleocenica Cushman, Cushman Lab. Foram.

Research Contr., vol. 24, p. 45, pi. 8, fig. 10, 194^. 

Allomorphina trigona Plummer (not Reuss), Texas Univ. Bull.

2644, p. 129, pi. 8, fig. 5, 1927. 

Kline, Mississippi Geol. Survey Bull. 53, p. 56, jl. 6, flgs. 1,

2, 1943.

Cushman and Todd, Cushman Lab. Foram. Research Contr., 

vol. 22, p. 63, pi. 11, figs. 11,15, 1946.

Test semielliptical in outline, trochoid, one side nearly 

straight, the other strongly curved, periphery?- broadly 

rounded; chambers distinct, slightly inflated, three in a 

whorl in the adult, increasing very rapidly in size as 

added; sutures distinct, slightly depressed; wall 

smooth; aperture an elongate opening on the ventral 

side at the base of the last-formed chamber, with a
CHILOSTOMELLIDAE 59

distinct, overhanging lip. Length 0.35 to 0.45 mm., 

breadth 0.25 to 0.30 mm., thickness 0.22 to 0.27 mm.

The types are from the Paleocene, 4.6 miles north of 

Fentress, Caldwell County, Tex. (44).

This species differs from A. trigona Reuss in the much 

broader form and the wide aperture. In 1936 Jennings 

proposed a new name, A. halli (Bull. Am. Paleontology, 

vol. 23, no. 78, p. 34, pi. 4, fig. 5,1936), referring to Mrs. 

Plummer's figures. His figures from the Hornerstown 

marl of Ntew Jersey are quite different from those of 

Mrs. Plummer and of later authors given above. The 

chambers are much more rounded and even in size in 

his figures. Therefore the name paleocenica would 

seem to represent the species from Texas, Arkansas, 

Mississippi, and Alabama.

In the original description of A. paleocenica, an error 

in the use of the type figures was made. Figure 10& 

is not the ventral view of the holotype but instead is a 

figure copied from Cushman and Todd (Cushman Lab. 

Foram. Research Contr., vol. 22, p. 63, pi. 11, fig. 11, 

1946) of a specimen from the Paleocene of Arkansas. 

On the present plate 16 this error is corrected and figures 

21a and & are the dorsal and ventral views of the holotype of A. paleocenica; the ventral view, figure 21&, 

being first published here.

Porters Creek formation, Matthews Landing marl member.

Alabama, Wilcox County (2, 4, 5) ; Butler County (3). 

Midway formation, upper part. Arkansas, Pulaski County (25). 

Midway formation. Arkansas, Clark County (30). 

Midway group. Texas, Caldwell County (44).

Allomorphina subtriangularis (Kline) Cushman

Plate 17, figures 1, 2 

Chilostomella subtriangularis Kline, Mississippi Geol. Survey

Bull. 53, p. 56, pi. 6, fig. 3,1943.

Test subtriangular, about one and one-half times as long as 

broad, greatest width a little above middle, ends bluntly 

rounded; wall smooth, finely punctate; aperture narrow, 

nearly straight. Average length 0.75 mm.—Kline.

The types are from the Paleocene Porters Creek clay, 

iy2 miles northwest of Montpelier, Clay County, Miss.

The dorsal side of specimens shows the chambers to 

be arranged in a triserial manner and the species seems 

to belong in Allomorphina.

Midway formation. Arkansas, Clark County (30).

Midway group. Texas, Caldwell County (44).

Wills Point formation. Texas, Caldwell County (34).

Allomorphina globulosa Plummer 

Plate 17, figure 3

Allomorphina, globulosa Plummer, Texas Univ. Bull. 2644, p. 130, 

pi. 8, figs. 4a, b, 1927.

Test bluntly ellipsoidal, last three chambers only partly embracing on superior face thus revealing the inner whorl; chambers very smooth and thin shelled; sutures not depressed; aperture a narrow slit at base of final chamber and protected by a 

conspicuous, somewhat flaring lip. Length of only specimen 

.35 mm.

From A. trigona Reuss this new species differs in showing no 

angulation of outline and in the greater overlap of its chambers,

so that the last two almost complete the final whorl. As only 

one specimen has been found in the Midway material no statement can be made regarding its distribution either geographically or stratigraphically.—Plummer.

As this is based on a single specimen it is difficult to 

determine the full characters. The type specimen is 

from the Paleocene, gully close to the short northwestsoutheast road about 2% miles S. 25° E. of Littig (Bastrop quadrangle), Bastrop County, Tex.

Genus CHILOSTOMELLOIDES Cushman, 1926

CMlostomelloides eocenica Cushman

Plate 17, figures 4, 5

CMlostomelloides eocenioa Cushman, Cushman Lab. Foram. Research Contr., vol. 1, pt. 4, p. 78, pi. 11, fig. 20,1926.

Plummer, Texas Univ. Bull. 2644, p. 129, pi. 8, fig. 8, 1927.

Cushman, Cushman Lab. Foram. Research Contr., vol. 16, p. 

72, pi. 12, fig. 11,1940.

Kline, Mississippi Geol. Survey Bull. 53, p. 57, pi. 6, fig. 8, 

1943.

Test elongate, about 2% times as long as broad, ends 

rounded, sides gently convex, tapering toward either 

end; wall smooth, very finely punctate; aperture semicircular, standing out at an angle from the contour of 

the test, with a distinct, slightly thickened lip. Length 

0.5 mm.

The types are from the Paleocene of the Mexia Oil 

Field, Texas.

Clayton formation, Chalybeate limestone member. Alabama,

Sumter County (9). 

Midway formation. Arkansas, Clark County (30).

*

Genus PTTLLENIA Parker and Jones, 1862

Pullenia quinqueloba (Reuss) Reuss var. angusta Cushman and

Todd

Plate 17, figure 6

Pullenia quinqueloba (Reuss) var. angiista Cushman and Todd,

Cushman Lab. Foram. Research Contr., vol. 19, p. 10, pi. 2,

figs. 3, 4, 1943; idem, vol. 22, p. 63, 1946. 

Cushman, idem, Special Pub. 16, p. 37, pi. 7, fig. 11, 1946. 

Pullenia qidnqueloba Plummer (not Reuss), Texas Univ. Bull.

2644, p. 136, pi. 8, fig. 12, 1927. 

Cole, Bull. Am. Paleontology, vol. 14, no. 51, p. 32, pi. 5, fig.

15, 1927. 

Cushman, Cushman Lab. Foram. Research Contr., vol. 15,

p. 73, pi, 12, fig. 17, 1939; idem, vol. 16, p. 72, pi. 12, figs.

13, 14, 1940. 

Toulmin, Jour. Paleontology, vol. 15, p. 607, pi. 81, fig. 24,

1941. 

Cushman and Todd, Cushman Lab. Foram. Research Contr.,

vol. 18, p. 42, pi. 7, fig. 15. 1942. 

Kline, Mississippi Geol. Survey Bull. 53, p. 57, pi. 6, figs. 4f

7, 1943. 

Applin and Jordan, Jour. Paleontology, vol. 19, p. 132 (list),

1945.

Variety differing from the typical form in the smaller 

size and somewhat narrower test. Length 0.25/ to 0.5C 

mm., breadth 0.20 to 0.40 mm., thickness 0.15 to 0.30 mm.

The types of the variety are from the Paleocene, clay 

pit of Mexia Brick Works about 1 mile west of Mexia> 

Limestone County, Tex.
60 PALEOCENE FORAMINIFERA OF GULF COASTAL REGION

Porters Creek clay. Mississippi, Kemper County (17). 

Porters Creek formation, Matthews Landing marl member.

Alabama, Butler County (3); Choctaw County (7). 

Clayton formation, Chalybeate limestone member. Alabama,

Sumter County (9). Mississippi, Noxubee County (16). 

Midway formation, upper part. Arkansas, Pulaski County (24,

25).

Midway formation. Arkansas, Pulaski County (27). 

Midway group. Texas, Limestone County (39) ; Caldwell

County (44). 

Wills Point formation, Mexia member. Texas, Limestone

County (36).

Family GLOBIGERINIDAE

Genus GLOBIGERINA D'OrMgny, 1826

Globigerina pseudo-bulloides piummer

Plate 17, figures 7, 8

GloMgerina pseudo-bulloides Plummer, Texas Univ. Bull. 2644,

p. 133, pi. 8, fig. 9, 1927.

Nuttall, Jour. Paleontology, vol. 4, p. 290,1930. 

Plummer, Texas Univ. Bull. 3201, pp. 54, 62 (lists), 1933. 

Glaessner, Moscow Univ. Problems of Paleontology, vol. 2-3,

p. 382, pi. 4, fig. 31, 1937. 

Cushman, Cushman Lab. Foram. Research Contr., vol. 16, p.

72, pi. 12, fig. 16, 1940.

Cushman and Todd, idem, vol. 18, p. 43, pi. 8, figs. 3, 4, 1942. 

Kline, Mississippi Geol. Survey Bull. 53, p. 58, pi. 6, figs.

9-11, 1943. 

Cushman, Cushman Lab. Foram. Research Contr., vol. 20,

p. 48, pi. 8, fig. 3, 1944. 

Applin and Jordan, Jour. Paleontology, vol. 19, p. 131 (list),

1945. 

Cushman and Todd, Cushman Lab. Foram. Research Contr.,

vol. 22, p. 64, 1946.

Test rotaliform, very obtusely trochoid to plane dorsally, composed of about two and one-half convolutions, of which the last 

consists most generally of 5 (rarely 6) highly ventricose chambers increasing rapidly in size; periphery broadly rounded and 

lobate; shell wall thin and distinctly punctate but finely reticulate ; superior face bearing a spire of small chambers only very 

slightly elevated, if at all, above the circumambient chambers of 

the final whorl; interior face less convex and with a very distinct, though not large, umbilical depression; aperture a single 

moderately large, lunate opening on the last chamber extending 

from the margin to the umbilicus and edged with a narrow, 

delicate, flaring lip. Diameter up to 0.4 mm.—Plummer.

The types are from the Paleocene, shallow ditch at 

road comer southeast of new Corsicana reservoir on the 

road to Mildred, Navarro County, Tex.

Porters Creek clay. Tennessee, Hardeman County (14). Mississippi, Kemper County (17).

Porters Creek formation, Matthews Landing marl member. 

Alabama, Wilcox County (1, 2, 4-6) ; Butler County (3) ; 

Choctaw County (7).

Naheola' formation. Coal Bluff marl member. Alabama, Wilcox County (8). 

Clayton formation. Alabama, Barbour County (10).

Chalybeate limestone member. Alabama, Sumter County

(9). Mississippi, Noxubee County (16). 

Pine Barren member. Alabama, Wilcox County (11) ; Butler County (12). 

Midway formation, lower part. Arkansas, Saline County (18,

. 19A-22) ; Pulaski County (23).

Midway formation, upper part. Arkansas, Pulaski County 

(24, 25).

Midway formation. Arkansas, Saline County (26) ; Pulaski

County (27) ; Clark County (29, 30). 

Midway group. Texas, Limestone County (39, 42) ; Navarro

County (40) ; Hunt County (41) ; Bastrop Courty (43, 49) ;

Caldwell County (44) ; Travis County (45) ; Freestone County

(50). 

Wills Point formation. Texas, Travis County (33) ; Caldwell

County (34) ; Williamson County (35).

Mexia member. Texas, Limestone County (36). 

Kincaid formation. Texas, Hunt County (37) ; Bastrop County

(38).

GloMgerina compressa Plummer

Plate 17, figure 9

GloMgerina compressa Plummer, Texas Univ. Bull. 2644, p. 135,

pi. 8, fig. 11,1927. 

Jennings, Bull. Am. Paleontology, vol. 23, no. 78, p. 193, pi.

31, fig. 8, 1936. 

Glaessner, Moscow Univ. Problems of Paleontology, vol. 2-3,

p. 382, pi. 4, fig. 32, 1937. 

Toulmin, Jour. Paleontology, vol. 15, p. 607, pi. 82, figs. 1, 2,

1941. 

Cushman and Todd, Cushman Lab. Foram. Research Contr.,

vol. 18, p. 44, pi. 8, figs. 5, 6, 1942. 

Kline, Mississippi Geol. Survey Bull. 53, p. 5S, pi. 6, figs.

5, 6, 1943. 

Cooper, Jour. Paleontology, vol. 18, p. 353, pi. 54, figs. 8, 9,

1944. 

Cushman, Cushman Lab. Foram. Research Contr., vol. 20,

p. 47, pi. 8, fig. 2, 1944.

Test small, rotaliform, closely coiled, somewhat compressed, 

equally biconvex; peripheral margin bluntly angular, lobate; 

chambers increasing gradually, 5 in last-formed whorl, moderately inflated, overlapping on dorsal face; sutures distinctly depressed and strongly curved on the dorsal side; shell wall thin, 

smooth, finely punctate; aperture a single moderately arched 

slit protected by a definite flaring flap at base of septal face and 

extending into the small but distinct umbilical depression. 

Diameter up to 0.4 mm.; average 0.3 mm.—Plummer.

The types are from the Paleocene, shallow ditch at 

road corner southeast of new Corsicana reservoir on the 

road to Mildred, Navarro County, Tex.

Porters Creek formation, Matthews Landing marl member.

Alabama, Wilcox County (4) ; Choctaw County (7). 

Naheola formation, Coal Bluff marl member. Alabama, Wilcox

County (8). 

Midway formation. Arkansas, Olark County (30).

Globigerina triloculinoides Plummer 

Plate 17, figures 10, 11

Glolifferina triloculinoides Plummer, Texas Univ. Bull. 2644,

p. 134, pi. 8, fig. 10, 1927. 

Jennings, Bull. Am. Paleontology, vol. 23, no. 7°, p. 193, pi.

31, fig. 10, 1936. 

Glaessner, Moscow Univ. Problems of Paleontology, vol.

2-3, p. 382, pi. 4, fig. 33, 1937. 

Cushman, Cushman Lab. Foram. Research CoTitr., vol. 16,

p. 72, pi. 12, fig. 15, 1940. 

Toulmin, Jour. Paleontology, vol. 15, p. 607, r1 . 82, fig. 3,

1941. 

Cushman and Todd, Cushman Lab. Foram. Research Contr.,

vol. 18, p. 43, pi. 8, figs. 1, 2,1942. 

Thalmann, Stanford Univ. Publ., Univ. Ser., Geol. Sci., vol.

3, no. 1, p. 13 (list), 1942. 

Martin, idem, vol. 3, no. 3, p. 10 (list), 1943.
TABLE 4.- 

[Prepared by J. B. Reeside, Jr.

-Distribution of Paieocene Rotaiiidae, Amphisteginidae, Cassidulinidae, Chilostomellidae, Globigerinidae, Globorotaliidae, and Anomalinidae in the Gulf Coastal region 

Numbers refer to locality list on pages 1-3. K=»localities reported by V. H. Kline, P=localities reported by H. J. Plummer and C. Q. Lalicker, C=Cuba and Haiti, G=Guatemala and Honduras, S=»goldado Rock, Trinidad, B. W. I.]

Rotaiiidae 

Spirittina selseyensis ..... ———————

PoteUinasp— ... _____ ——————

Eponides elevatus- ___ ..... ———— ..

ParreBa expansa _____ —— . ————

macneUL...... ....... ................

Odettes reticules-us .... — ———————

abuiUotensis .. ____ .... ..........

Epistomina eocenica ....... . ... .. ———

Stphonina prima ....... ..... —————

Amphisteginidae 

Asterigerinapriinarfa.. ...... ... .... ...

Cassidulinidae 

CeratobvMmina perplexa ................

Alabamina wttcoxensis— , ........ .......

wttcoxenitis var. limbata... ___ ——— PseudopaTrella cf. P. exifWr ...........

madrugaensis ............ _ ... — ...

wOcoxensis. ... ......... .... ... . ......

ChUostomellidae 

AUomoTphina paleocenica ..... ...... ... .

svbtriangularis-.. ...... .... . .........

globutosa ............. .... ... ..... . ....

Chilostomettoides eocenica..,.. .... ..... .

Pullenla guinqueloba var. angusta .......

Globlgerinidae 

Glooiperina pseudo-btdloides ...... ... ....

eompresta.. ... ..... ..... .... .........

trOocutfaoUea............. ...........

Globorotaliidae 

OloborotalM wilcoxensis var. acute- ...... 

creueata var. aegw ........... . ... .... .

SL ®, fflwm&ronocea ____ .... __ ..

a&etri ................. ... ........ v _

AnomalinidaG 

Anomattnauinbonifera ..................

midwitsensis...... .... .... ... ___ ...

•rittdwagewig var. trockoMea ..........

acuta..... ..... ...... . ................

fiwSo&ato.— ... ___ ...............

cleinenttaM.. ........................

clementmw var. assimilis ............

weUert~ " _

martineMnsii .........................

cvbam — ,._... __ ...... ______

mairugMniis..-.. .... .... ...... ... ...

sp. A......... ......... ..............

sp. JB, ———— ... _ .... __ .........

Boldia vemdersluiti _________ ,..

dbfaides proecMrsorttt* .................

oBeai.................................

newtmMe... .........................

howelli

blanpiedi.. ... .. .......................

brwni....... .........................

mirtflcus ..... . ............ . ..........

Alabama

1

X

X

X

X

X

....

X

X

2

X

X

X

X

X

X

X

X

X 

X

X

X

X

X

3

X

X

X

X

X

X

X

X

X

*

X

X

X

X

4

X

X

X

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X

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X

X 

X

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5

X

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•It

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*x~

X

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6

X

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"x"

7

X 

X

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TT

X

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X

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'X

8

X 

X

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X 

X

X

X

X

2£

X

X

9

X

X

X

X

X

X

X

X

X

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"x"

10

"X

X

X

X

11

X

X

X

X

12

X

X

X

X

X

X

X

X

Tenn.

13

X

X

X

X

X

X

X

X

....

X

14

X

X

X

X

X

X

X

Miss.

15

....

16

X

X 

X

X

X

17

X

X

X 

X

X

X

X

X

K

X

X

—

X

Arkansas

18

X

X

X

X

19

X

X

X

X

X

X

X

X

20

X

X

X

X

X

21

X

....

X

X

X

X

22

X

X

— -

X

x

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X

at"

sT

x

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23

....

X

X

X

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24

X

X

X

X

X

X

X

X

X

X

X

25

X

X

X

X

X

X

X

X

X

X

X

X

X

X

26

X

X

X

X

X

X

X

"x"

X

X

x

27

X

X

X

X

X

X

X

X

X 

X 

X

X

X

28

X

X

X

....

....

....

29

X

X

X

30

X

X

X

X

X

X

X

X

X

X

X

X

X

X 

X

X

X

31

X

X

X

32

TM

....

Texas

33

X

X

X

X

X

X

X

X

X

34

X

X

X

X

X

X

X

X

X

35

X

-—

X

....

....

36

X

X

X

X

X

X

X

X

X

X

X

X

X

X

37

X

X

X

X

X

X

X

X

X

38

X

X

X

....

X

....

X

X

39

X

X

X

X

X

X

X

X

X

X

X

X

40

X

X

X

X

X

X

X

X

X

X

41

....

X

X

X 

X

X

X

X

X

X

X

X

X

X

42

X

X

X

X

....

X

X

X

'

43

X

X

X

X

X

X

X

X

X

....

X

X

X

44

X

X

X

X

X

X

X

X

X

X

X

X 

X

X

X

X

45

X

X

X

....

X

X

46

X

X

X

X

47

X

X

X

48

X

49

X

X

X

X

X

X

X

50

X

, ,

51

1

52 P

X

X

X

X

X

C

X

X

X

X

X

X

X

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x

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8

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X

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X

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915269—SI (Face p. 60)

GLOBIGERINIDAE 61

Beck, Jour. Paleontology, vol. 17, p. 609, pi. 108, figs. 2, 3,

1943. 

Kelley, Am. Assoc. Petroleum Geologists Bull., vol. 27, p.

11 (list), 1943.

Curran, idem, pp. 1378, 1381 (lists), 1943. 

Kline, Mississippi Geol. Survey Bull. 53, p. 59, pi. 6, figs. 12,

13, 1943. 

Cushman, Cushman Lab. Foram. Research Contr., vol. 20, p.

48, pi. 8, fig. 4, 1944. 

Cooper, Jour. Paleontology, vol. 18, p. 353, pi. 54, figs. 12,

13, 1944.

Applin and Jordan, idem, vol. 19, p. 132 (list), 1945. 

Cushman and Todd, Cushman Lab. Foram. Research

Contr., vol. 22, p. 64, 1946.

Test spiral, trochoid, composed of about 2 convolutions, the 

last of which is composed of 3% very rapidly increasing and 

highly globose chambers; periphery very broadly rounded and 

distinctly lobate; shell surface strongly reticulate; superior 

face rounded with a very low spire of neatly coiled tiny chambers of the preceding whorl; inferior face rounded with a very 

shallow umbilical depression; aperture a small arched slit on 

the last chamber and edged with a more or less prominent, 

delicately notched flap that extends from a point near the 

periphery to the umbilical depression. Greatest diameter up 

to 0.35 mm.; usually less.—Plummer.

The types are from the Paleocene, shallow ditch at 

road corner southeast of new Corsicana Reservoir on 

the road to Mildred, Navarro County, Tex.

Porters Creek formation, Matthews Landing marl member. 

Alabama, Wilcox County (2, 4, 5) ; Butler County (3) ; 

Choctaw County (7).

Naheola formation. Coal Bluff marl member. Alabama, Wilcox County (8). 

Clayton formation. Alabama, Barbour County (10).

Chalybeate limestone member. Alabama, Sumter County

(9). Mississippi, Noxubee County (16). 

Pine Barren member. Alabama, Butler County (12). 

Porters Creek clay. Mississippi, Kemper County (17). 

Midway formation, lower part. Arkansas, Saline County (19A,

22). 

Midway formation, upper part. Arkansas, Pulaski County (24,

25).

Midway formation. Arkansas, Clark County (30). 

Midway group. Texas, Limestone County (39, ^2) ; Navarro 

County (40) ; Hunt County (41) ; Caldwell County (44); 

Bastrop County (49).

Wills Point formation. Texas, Travis County (33) ; Caldwell County (34).

Mexia member. Texas, Limestone County (36). 

Kincaid formation. Texas, Hunt County (37).

Family GLOBOROTALILDAE 

Genus G10BOB.OTALIA Cushman, 1927

Globorotalia wilcoxensis Cushman and Ponton var. acuta Toulmin 

Plate 17, figures 12, 13

Globorotalia wilcoxensis Cushman and Ponton var. acuta Toulmin, Jour. Paleontology, vol. 15, p. 608, pi. 82, figs. 6-8. 

Cushman and Renz, Cushman Lab. Foram. Research Contr.,

vol. 18, p. 12, pi. 3, fig. 2, 1942.

Cushman, Am. Jour. Sci., vol. 242, p. 15, pi. 2, flgs. 16, 17, 

1944; Cushman Lab. Forani. Research Contr., vol. 20, p. 

48, pi. 8, fig. 5, 1944.

Test trochiform, plano-convex, dorsal side flat, ventral side 

strongly convex, deeply umbilicate, periphery strongly lobate, 

915259—51———5

acute, and bounded by a thick flange; chambers distinct, about 

4% in the last whorl, increasing regularly in size as added; 

sutures distinct, on the dorsal side slightly curved, limbate, 

slightly if at all depressed, on the ventral side radiate, depressed; 

wall roughened with minute, low spinose processes, especially 

along the peripheral border ; aperture an arched opening on the 

ventral side of the final chamber extending from the peripheral 

flange to the umbilicus. Length 0.46 mm.; width 0.37 mm.; 

thickness 0.24 mm.—Toulmin.

The types are from the Eocene Salt Mountain limestone, from Richmond Branch, half a mile north of Salt 

Mountain, Clarke County, Ala.

Porters Creek formation, Matthews Landing marl member.

Alabama, Wilcox County (2, 4-6). 

Naheola formation, Coal Bluff marl member. Alabama, Wilcox

County (8). 

Wills Point formation, Mexia member, Texas, Limestone

County (36). 

Midway group. Texas, Hunt County (41).

Globorotalia crassata (Cushman) Cole var. aequa Cushman and

Renz

Plate 17, figure 14

Globorotalia crassata (Cushman) Cole var. aequa Cushman anc1 

Renz, Cushman Lab. Foram. Research Contr., vol. 18 

p. 12, pi. 3, fig. 3, 1942. 

Cushman and Todd, idem, vol. 18, p. 44, pi. 8, figs. 7-9, 1942.

Variety differing from the typical form in the much 

smoother surface and the broader and more arcuate 

chambers, especially the later ones.

The types of the variety are from the Paleocene 

Soldado formation, Soldado Rock, Trinidad, B. W. I.

Porters Creek formation, Matthews Landing marl member. 

Alabama, Choctaw County (7).

Globorotalia of. G. memibranacea (Ehrenberg) White 

Plate 17, figure 15

This species was recorded from the type locality (7) 

of the Naheola formation of Alabama by a single specimen and that not entirely typical (Cushman and Todd, 

Cushman Lab. Foram. Research Contr., vol. 18, p. 45, 

pi. 8, fig. 10,1942). No additional specimens have been 

found that could be assigned to this species.

Globorotalia albeari Cushman and Bermudez 

Plate 24, figures 13-15

Globorotalia albeari Cushman and Bermudez, Cushman Lab. 

Foram. Research Contr., vol 25, p. 33, pi. 6, flgs. 13-15, 1949.

Test very small for the genus, strongly biconvex, dorsal side showing all the coils and ventral side only the 

last-formed whorl, periphery somewhat rounded; 

chambers not very distinct, 9 or 10 in the last-formed 

whorl, only slightly inflated ventrally, increasing very 

gradually in size as added; sutures fairly distinct but 

only slightly depressed, except in the last whorl on tl °, 

ventral side, rather strongly curved on the dorsal side; 

wall slightly spinose, coarsely perforate; aperture an 

elongate opening on the ventral side of the last-formed 

chamber extending from nearly the inner end to tl ^
62 PALEOCENE FORAMINIFERA OF GULF COASTAL REGION

periphery and with a distinct thin lip. Diameter 0.30 

to 0.32 mm., thickness 0.20 mm.

The types are from the Paleocene JVladruga formation, under highway bridge on Central San Antonio, 

Madruga, Habana Province, Cuba.

This species cliffers from G. praemenardii Cushman 

and Stainforth in its smaller size, more numerous chambers, more strongly biconvex test, and less umbilicate 

ventral side.

Family ANOMALINIDAE

Genus ANOMALINA D'Orbigny, 1826

Anomalina umbonifera (Schwager) Cushman and Ponton

Plate 17, figure 16

Discorbina umbonifera Schwager, Palaeontographica, vol 30,

Pal. Teil, p. 126, pi. 27 (4), fig. 14,1883. 

Anomalina umbonifera Cushman and Panton, Cushman Lab.

Foram. Research Contr., vol. 8, p. 72, pi. 9, fig. 11, 1932. 

Cushman, Am. Jour. Sci., vol. 242, p. 15, pi. 2, figs. 18, 19,

1932; Cushman Lab. Foram. Research Coutr., vol. 20,

p. 27, pi. 4, fig. 30; p. 49, pi. 8, fig. 6, 1944.

Test strongly compressed, ventral side completely involute, dorsal side slightly evolute, periphery slightly 

rounded; chambers distinct, slightly inflated, 7 or 8 

chambers making up the adult whorl, increasing gradually and rather uniformly in size as added; sutures 

distinct, slightly limbate, gently curved; wall smooth, 

finely perforate; aperture at the base of the last-formed 

chamber, near the periphery. Diameter up to 0.40 mm., 

thickness 0.15 mm.

The types of this species are from the middle Eocene 

of northern Africa. It is recorded from the Paleocene 

and Eocene (Wilcox group) of the United States.

Naheola formation, Coal Bluff marl member. Alabama, Wilcox 

County (8).

Anomalina midwayensis (Flummer) Cushman 

Plate 17, figures 17-19

Truncatulina midwayensis Plummer, Texas Univ. Bull. 2644,

p. 141, pi. 9, fig. 7; pi. 15, fig. 3,1927.

Anomalina midwayensis Cushman, Cushman Lab. Foram. Research Contr., vol. 16, p. 73, pi. 12, fig. 18, 1940.

Kline, Mississippi Geol. Survey Bull. 53, p. 60, pi. 6, figs. 17, 

18, 1943.

Cooper, Jour. Paleontology, vol. 18, p. 354, pi. 54, figs. 15-17, 

19-44.

Cushman and Todd, Cushman Lab. Foram. Research Contr., 

vol. 22, p. 64, pi. 11, figs. 18, 19, 1946.

Test almost equally biconvex, moderately compressed; peripheral margin rounded; convolutions about 2, the final one 

being strongly embracing; chambers usually 9 in final whorl, 

conspicuously punctate, gradually increasing, moderately curving; sutures broadly elevated on both sides, tapering toward 

the margin, and curved; aperture a slit at base of septal face 

under a narrow lip that extends to the umbilicus. Diameter 

up to 0.5 mm.; usually less.—Plummer.

The types are from the Paleocene, shallow ditch at 

road corner southeast of new Corsicana reservoir on the 

road to Mildred, Navarro County, Tex.

Clayton formation, Chalybeate limestone member. Alabama,

Sumter County (9).

Porters Creek clay. Mississippi, Kemper County (17). 

Midway formation, upper part. Arkansas, Pulrski County

(24, 25). 

Midway formation. Arkansas, Pulaski County (27) ; Clark

County (29-31). 

Midway group. Texas, Limestone County (39, 42) ; Navarro

County (40) ; Hunt County (41) ; Caldwell County (44) ;

Travis County (46) ; Van Zandt County (47) ; Bastrop

County (49). 

Wills Point formation. Texas, Travis County (33) ; Caldwell

County (34).

Mexia member. Texas, Limestone County (36). 

Kincaid formation. Texas, Hunt County (37) ; Bastrop County

(38).

Anomalina midwayensis (Flummer) Cushman var. trochoidea

(Plummer) Kline 

Plate 18, figures 1, 2

Truncatulina midicayensis Plummer var. trochoidea Plummer,

Texas Univ. Bull. 2644, p. 142, pi. 9, fig. 8, 1927. 

Anomalina midwayensis (Plummer) Cushman var. trochoidea

Kline, Mississippi Geol. Survey Bull. 53, p. 61, pi. 6, figs.

15, 16, 1943.

From the type this variety is distinguished by the more 

strongly trochoid dorsal coiling of the convolutions Diameter 

up to 0.6 mm.; usually less.—Plummer.

The types are from the Paleocene, exposure along a 

small branch about % mile northwest of Tehuacana, 

and 0.2 mile north of the Tehuacana-Waco road on the 

first road turning north, Limestone County, Tex.

Midway formation. Arkansas, Saline County (26) ; Pulaski

County (27) ; Clark County (30). 

Midway group. Texas, Caldwell County (44).

Anomalina acuta Flummer 

Plate 18, figures 3-6

Anomalina ammonoides (Reuss) var. acuta Plummer, Texas

Univ. Bull. 2644, p. 149, pi. 10, fig. 2,1927.

Anomalina acuta Glaessner, Moscow Univ. Problems of Paleontology, vol. 2-3, p. 386, pi. 5, fig. 40, 1937. 

Toulmin, Jour. Paleontology, vol. 15, p. 608, pi. 82, figs.

9, 10, 1941. 

Cushman and Renz, Cushman Lab. Foram Reserrch Contr.,

vol. 18, p. 12, pi. 3, fig. 6, 1942. 

Kline, Mississippi Geol. Survey Bull. 53, p. 59, pi. 5, figs. 3,

4, 1943. 

Cooper, Jour. Paleontology, vol. 18, p. 353, pi. 51, figs. 3-5,

1944. 

Cushman and Todd, Cushman Lab. Foram. Research Contr.,

vol. 22, p. 64, pi. 11, figs. 13, 14, 1946.

Test involute, much compressed, almost equally biconvex but 

slightly more flattened above; peripheral margin subacute; 

chambers numerous, about 13-15 to the final convolution, narrow, and slightly curving; sutures marked by more or less distinct limbations, which on the ventral face terminate along the 

inner edge of the convolution in a series of fine bead's that surround a thick spiral or irregular filling of translucent shell 

material in the umbilical recess, and on the dorsal face merge 

at the center into a more or less prominently developed elevated 

boss; shell wall distinctly but not coarsely punctate; aperture 

an arched opening over the peripheral margin and extending
ANOMALINIDAE 63

toward the umbilicus. Diameter up to 0.4 mm.; average 0.25 

mm.—Plummer.

The types are from the Paleocene, shallow ditch at 

road corner southeast of new Corsicana reservoir on 

the road to Mildred, Navarro County, Tex.

Porters Creek formation, Matthews Landing marl member.

Alabama, Wilcox County (2) ; Butler County (3). 

Clayton formation, Chalybeate limestone member. Mississippi,

Noxubee County (16). 

Midway formation, lower part. Arkansas, Saline County (20,

22) ; Pulaski County (23). 

Midway formation, upper part. Arkansas, Pulaski County (24,

25). 

Midway formation. Arkansas, Saline County (26) ; Pulaski

County 27; Clark County (30). 

Midway group. Texas, Hunt County (41) ; Bastrop County

(43, 49) ; Caldwell County (44) ; Travis County (45). 

Wills Point formation. Texas, Travis County (33).

Mexia member. Texas, Limestone County (36). 

Kincaid formation. Texas, Hunt County (37) ; Bastrop County

(38).

Anomalina basilobata Cushman and Renz 

Plate IS, figure 7

Anomalina basilobata Cushman and Renz, Cushman Lab. Foram. 

Research Contr., vol. 18, p. 12, pi. 3, fig. 7, 1942.

Test nearly equally biconvex, periphery rounded, central region of each side somewhat depressed, ventral 

side umbilicate; chambers about 9 in the adult whorl, 

increasing gradually and rather regularly in size as 

added, very slightly if at all inflated, at the base on 

both sides with a projection toward the center; sutures distinct, gently curved; wall smooth; aperture 

at the base of the last-formed chamber at the median 

line. Diameter of holotype 0.40 mm., thickness 0.10 

mm.

The types are from the Paleocene Soldado Rock formation, Soldado Rock, Trinidad, B. W. I. It has not 

been recorded elsewhere.

Anomalina clementiana (D'Orbigny) Franke 

Plate 18, figures 8-11

Rosalind clementiana, D'Orbigny, Soc. geol. France Mem., 1st

ser., vol. 4, p. 37, pi. 3, figs. 23-25,1840. 

Anomalitia clementiiui Franke, Greifswald Univ., Geol.-

palaeont. Inst., Abh., vol. 6, p. 85, pi. 7, figs. 12a-c, 1925;

Preuss. geol. Landesanstalt Abh., new ser., vol. Ill, p.

179, pi. 16, figs. 9a-c, 1928. 

Cushman, Tennessee Div. Geology Bull. 41, p. 61, pi. 13, figs.

la-c, 1931; Cushman Lab. Foram. Research Contr., vol.

7, p. 46, pi. 6, figs. lOa-c, 1931. 

Jennings, Bull. Am. Paleontology, vol. 23, no. 78, p. 38, pi.

5, figs. 2a, b, 1936. 

Cushman, Cushman Lab. Foram. Research Contr., vol. 16, p.

29, pi. 5, figs. 7, 8, 1940. 

Cushman and Goudkoff, idem, vol. 20, p. 63, pi. 10, fig. 12,

1944. 

Cushman and Todd, idem, vol. 22, p. 64, pi. 11, figs. 16, 17,

1946.

Cushman, U. S. Geol. Survey Prof. Paper 206, p. 155, pi. 63, 

figs. 12, 13, 1946.

Test somewhat tending toward planispiral in the 

adult, trochoidi in the young, variable, compressed, 

periphery rounded; chambers distinct, 7 to 9 in the lastformed whorl; sutures on the dorsal side curved, limbate 

and strongly raised except in the last few chambers of 

the adult, which are smooth, in many specimens slightly 

depressed; sutures on the ventral side nearly radial, 

depressed, the ends of the chambers raised between the 

sutures; wall smooth except for the ornamentation already noted in the earlier portions; aperture peripheral 

and extending onto the dorsal side. Diameter 0.30 to 

0.40 mm., thickness 0.10 to 0.15 mm.

Porters Creek clay. Mississippi, Keniper County (17). 

Midway formation, upper part. Arkansas, Pulaski County (25). 

Midway formation. Arkansas, Pulaski County (27). 

Midway group. Texas, Caldwell County (44). 

Wills Point formation, Mexia member. Texas, Limestone County 

(36).

Anomalina clementiana (D'Orbigny) Franke var. assimilis 

Cushman and Bermudez

Plate 21, figures 10-12

Anomalina clementiana (D'Orbigny) var. assimilis Cushman 

and Bermudez, Cushman Lab. Foram. Research Contr., vol. 

24, p. 74, pi. 12, fig. 17, 1948.

Variety differing from the typical form in the slightly 

narrower chambers and slightly thicker test.

The types are from the Paleocene Madruga formation, under highway bridge on Central San Antonio, 

Madruga, Habana Province, Cuba.

The variety (A. clementiana assimitis) is very closely 

related to the species which is from the Upper 

Cretaceous.

Anomalina welleri (Plummer) Plummer 

Plate 18, figure 12

Truncatulina welleri Plummer, Texas Univ. Bull. 2644, p. 143,

pi. 9, fig. 6, 1927. 

Anomalina welleri Plummer, idem, Bull. 3201, pp. 54, 62 (lists),

1933. 

Nonionella welleri Kline, Mississippi Geol. Survey Bull. 53, p. 44,

pi. 4, fig. 21,1943.

Test small, equally biconvex, considerably compressed; chambers 10-11 in final whorl, strongly punctate, narrow, curved increasing gradually in size; sutures distinct, narrow tapering 

slightly elevated, and curved in a broad gentle swing, those on 

the ventral side .being joined in a low ridge about the small 

umbilical depression; aperture a low arch very close to the 

periphery and narrowing toward the umbilicus under a narrow 

lip. Diameter up to 0.35 mm.; usually about 0.25 mm.— 

Plummer.

The types are from the Paleocene, shallow ditch at 

road corner southeast of new Corsicana reservoir on the 

road to Mildred, Navarro County, Tex.

Midway group. Texas, Travis County (46).
64 PALEOCENE FORAMINIFERA OF GULF COASTAL REGION

Anomalina martinezensis Cushman and Bermudez 

Plate 21, figures 7-9

Anomalina martinezensis Cushman and Bermudez, Cushman 

Lab. Foram. Research Contr., vol. 24, p. 74, pi. 12, figs 14-16, 

1948.

Test trochoid, biconvex, both dorsal and ventral sides 

depressed in the median portion, periphery broadly 

rounded, tending to become slightly angular in the 

adult; chambers distinct, somewhat inflated, about 8 in 

the adult whorl, increasing very gradually in size as 

added; sutures distinct, limbate and slightly raised, 

dorsally oblique and somewhat curved, ventrally nearly 

radial; wall very coarsely perforate; aperture at the 

peripheral margin extending onto the ventral side at 

the base of the last-formed chamber, with a distinct lip. 

Diameter 0.65-0.80 mm.; thickness 0.32-0.40 mm.

The types are from the Paleocene, San Juan y Martinez, Pinar del Bio Province, Cuba.

This species resembles A. midwayetisis (Plummer) 

but differs in the more coarsely perforated wall, larger 

size, and much rougher surface. It also resembles 

Cibicides vulgar is (Plummer).

Anomalina cubana Cushman and Bermudez 

Plate 21, figures 13-15

Anomalina cubana Cushman and Bermudez, Cushman Lab. 

Foram. Research Contr., vol. 24, p. 86, pi. 15, figs. 7-9, 1948.

Test unequally biconvex, d'orsal side nearly flattened 

in the central area or slightly convex, ventral side more 

strongly convex, with a distinct umbilical depression, 

periphery broadly rounded; chambers fairly distinct, 

slightly inflated on the dorsal side, strongly so on the 

ventral side, 5 or 6 in the adult whorl, increasing rather 

rapidly but uniformly in size as added; sutures fairly 

distinct and slightly depressed on the dorsal side, 

strongly depressed ventrally, sinuous; wall smooth, distinctly perforate; aperture at the base of the final chamber extending from near the periphery slightly onto the 

ventral side, without a definite lip. Length of holotype 

0.85 mm.; breadth 0.70 mm.; thickness 0.40 mm.

The types are from the Paleocene Madruga formation, under highway bridge on Central San Antonio, 

Madruga, Habana Province, Cuba.

This species differs from A. tnadrugaensis Cushman 

and Bermudez in the smoother wall, more sinuous sutures, and the largely ventral and lipless aperture.

Anomalina madrugaensis Cushman and Bermudez 

Plate 22, figures 1-3

Anomalina maarugaensis Cushman and Bermudez, Cushman 

Lab. Foram. Research Contr., vol. 24, p. 86, pi. 15, figs. 4-6, 

1948.

Test of medium size, nearly bilaterally symmetrical, 

dorsal and ventral sides both depressed in the middle 

area, periphery rounded; chambers distinct, somewhat 

inflated, about 6 in the adult whorl, of uniform shape,

very gradually increasing in size as added; sutures distinct, depressed, very slightly curved; wall smooth, 

coarsely perforate; aperture at the base of the peripheral margin of the last-formed chamber, extending 

slightly onto the ventral side, with a slight lip. Length 

of holotype 0.70 mm.; breadth 0.57 mm.; thickness 0.30 

mm.

The types are from the Paleocene Madruga formation, under highway bridge on Central Sail Antonio, 

Madruga, Habana Province, Cuba.

This species resembles some of the forms that have 

been referred to A. grosserugosa (Giimbel) but differs 

from the typical form of that species in the bilaterally 

symmetrical test and fewer chambers.

Anomalina praespissiformis Cushman and Bermudez 

Plate 22, figures 7-9

Anomalina praespissiformis Cushman and Bermudez, Cushman 

Lab. Foram. Research Contr., vol. 24, p. 86, pi. 15, figs. 1-3, 

1948.

Test fairly small, strongly compressed, about equally 

biconvex, periphery rounded, ventral side umbilicate, 

dorsal side slightly depressed in the earlier portion; 

chambers distinct, 10-12 in the adult whorl, of uniform 

shape, increasing very gradually in size as added, very 

slightly inflated; sutures distinct, very slightly depressed, curved; wall smooth; aperture at the base of 

the peripheral margin of the last-formed ch amber extending over onto the ventral side, with a very slight lip. 

Length of holotype 0.47 mm.; breadth 0.40 rr m.; thickness 0.15 mm.

The types are from the Paleocene Madruga formation, under highway bridge on Central San Antonio, 

Madruga, Habana Province, Cuba.

This species strongly resembles A. alazanensis Nuttall 

var. spissiformis Cushman and Stainforth from the 

Oligocene of Trinidad but differs in the siraller size, 

fewer chambers, and more strongly curved sutures.

Anomalina sp. A 

Plate 18, figure 13

Anomalina sp. Cushman and Renz, Cushman Lab. Foram. Research Contr., vol. 18, p. 13, pi. 3, fig. 8,1942.

The figured specimen is one of a series of a rather variable 

form which, from the material, is difficult to describe. It does 

not seem identical with any of the species described from the 

Midway group. It is like some of the Tertiary forirs that have 

been referred to Anomalina grossemgosa (Giimbel).—Cushman 

and Itenz.

The specimens are from the Paleocene Soldado Rock 

formation, Soldado Rock, Trinidad, B. W. I.

Anomalina sp. B 

Plate 18, figures 14, 15

Anomalina sp. Cushman and Todd, Cushman Lab. Foram. Research Contr., vol. 18, p. 45, pi. 8, figs. 11, 12, If 42.

A compressed species with a very coarsely perforate wall 

occurs in the Naheola material, but it is difficult to identify it
ANOMALESTIDAE 65

with any of the species described from this part of the Eocene.— 

Cushman and Todd.

The specimens are from the Naheola formation, 

Naheola Landing, Tombigbee River, Ala. (7). 

Genus BOLDIA Van Bellen, 1946 

Boldia vandersluisi Van den Bold 

Plate 23, figure 1

Boldia vandersluisi Van den Bold, Thesis Univ. Utrecht, p. 124,

pi. 18, fig. 6,1946.

Van Bellen, Cushman Lab. Foram. Research Gontr., vol. 22, 

p. 122, text fig. 1, 1946.

Test large, slightly biconcave, ventral side with a heavy keel. 

Periphery broadly truncate. About 6 chambers in the last 

formed whorl visible dorsally. Sutures very indistinct. Aperture ventral. Diam.: 1.00; T: 0.60.—Van Bellen.

The types are from the lower Eocene or Paleocene of 

Guatemala and questionable specimens have been recorded from British Honduras.

Boldia carinata Cushman and Bermudez 

Plate 22, figures 4-6

Boldia carinata Cushman and Bermudez, Cushman Lab. Foram. 

Research Contr., vol. 24, p. 75, pi. 11, figs. 11-13, 1948.

Test trochoid in the early stages, becoming planispiral 

in the adult, periphery in the adult truncate with a 

median depression, the dorsal angle carinate, both dorsal and ventral sides concave in the adult, ventral side 

somewhat umbilicate; chambers fairly distinct, inflated 

veiitrally, very slightly so dorsally, usually 8 chambers 

in the adult whorl; sutures indistinct dorsally, ventfally 

distinct and depressed, somewhat sinuous; wall coarsely 

perforate, slightly roughened, not papillate, dorsal side 

with a slight stellate appearance owing to the lip above 

the aperture; aperture extending from the periphery 

along the dorsal margin of the last-formed chamber, 

with a slight lip. Diameter 0.50-0.60 mm.; thickness 

0.38-0.43 mm.

The types are from the Paleocene Madruga formation, under highway bridge, Central San Antonio, 

Madruga, Habana Province, Cuba.

This species differs from B. vandersluisi Van den Bold 

in the smaller size, and more definitely keeled and more 

concave periphery.

The original description of the genus gives the aperture as ventral but in this and the following two species 

it is largely peripheral and extending over onto what 

is regarded as the dorsal side. There is a question as 

to which is the dorsal side, and in these three species 

from Cuba the dorsal side is opposite to what it is 

stated to be in the other known species of the genus. 

To determine the dorsal and ventral sides either young 

specimens or sections should be studied.

Boldia madrugaensis Cushman and Bermndez 

Plate 22, figures 10-12

Boldio nmdrugaensis Cushman and Bermudez, Cushman Lab. 

Foram. Research Contr., vol. 24, p. 75, pi. 12, figs. 4-6, 1948.

Test trochoid in the young, nearly planispiral in the 

adult, periphery in the adult becoming somewhat truncate in the last portion, earlier portion rounded, both 

dorsal and ventral sides somewhat concave, ventral side 

somewhat umbilicate; chambers fairly distinct, 7 or 8 in 

the adult whorl, distinctly inflated on the ventral side; 

sutures distinct and depressed veiitrally, slightly curved, 

nearly radial, dorsally indistinct; wall coarsely perforate, smooth on the ventral side, dorsal side with numerous, irregular raised portions radiating out toward 

the periphery; aperture peripheral and extending over 

onto the dorsal side as a narrow slit at the inner margin 

of the last-formed chamber, with a distinct lip. Diameter 0.45-0.65 mm.; thickness 0.30-0.45 mm.

The types are from the Paleocene Madruga formation, under highway bridge on Central San Antonio, 

Madruga, Habana Province, Cuba.

This species differs from B. vandersluisi Van den 

Bold in the much smaller size, less truncate border, umbilicate ventral side, and irregularly papillate dorsal 

side.

Boldia cnbensis Cushman and Bermudez 

Plate 22, figures 13, 14

Boldia cubensis Cushman and Bermudez, Cushman Lab. Foram. 

Research Contr., vol. 24, p. 74, pi. 11, figs. 15, 16, 1948.

Test in the early stages trochoid, in the adult becoming nearly planispiral, periphery in the adult becoming 

truncate, earlier portion with a narrowly rounded border, both dorsal and ventral sides somewhat concave; 

chambers fairly distinct, about 6 in the adult whorl, 

slightly inflated, increasing very gradually in size as 

added; sutures rather indistinct, curved, little if at all 

depressed; wall smooth except for the umbilical area of 

the dorsal side, which has a stellate thickening; aperture a low opening at the peripheral margin, extending 

over 011 the dorsal side, with a slight lip. Diameter of 

holotype 0.70 mm.; thickness 0.45 mm.

The types are from the Paleocene Madruga formation, under highway bridge on Central San Antonio, 

Madruga, Habana Province, Cuba.

The species differs from B. vandersluisi Van den Bold 

in the smaller size, less truncate periphery, and the 

more curved sutures.

Genus CIBICIDES Montfort 1808

Cibicides praecursorius (Schwager) Cushman and Ponton 

Plate 19, figures 1-6

Discorbina praecursoria Schwager, Palaeontographica, vol. 30,

Pal. Theil, p. 125, pi. 24 (4), fig. 12; pi. 29 (6), fig. 16,1883.

Cibicides praecursorius Cushman and Ponton, Oushman Lab.

Foram. Research Contr., vol. 8, p. 72, pi. 9, fig. 14, 1932.

Cushman and Garrett, idem, vol. 15, p. 88, 1939.

Toulmin, Jour. Paleontology, vol. 15, p. 610, pi. 82, figs. 19-

21, 1941.

Cushman and Renz, Cushman Lab. Foram. Research Contr., 

vol. 18, p. 13, pi. 3, fig. 9,1942.
66 PALEOCENE FORAMINIFERA OF GULF COASTAL REGION

Cushman and Todd, idem, vol. 18, p. 45, pi. 8, figs. 17-20,

1942. 

Kline, Mississippi Geol. Survey Bull. 53, p. 62, pi. 5, figs. 5,

6, 1943. 

Cushman, Cushman Lab. Foram. Research Contr., vol. 20, p.

28, pi. 4, fig. 28; p. 49, pi. 8, figs. 17-20, 1944; Am. Jour.

Sci., vol. 242, p. 18, pi. 2, figs. 23, 24,1944. 

Oushman and Todd, Cushman Lab. Foram. Research Contr.,

vol. 22, p. 65, pi. 11, figs. 20, 21, 1946.

Test with the dorsal side flat or slightly concave, ventral side convex, periphery subacute; chambers distinct, 

slightly inflated on the ventral side, 6 to 8 in the adult 

whorl, increasing gradually in size as added; sutures 

distinct, curved, slightly limbate; wall smooth, coarsely 

perforate; aperture extending from the peripheral margin over onto the dorsal side beneath the somewhat 

extended lip of the last-formed chamber. Length up 

to 0.50 mm., breadth 0.40 mm., thickness 0.12 mm.

Porters Creek formation, Matthews Landing marl member. Alabama, Wilcox County (1, 2, 4) ; Choctaw County (7).

Naheola formation. Coal Bluff marl member. Alabama, Wilcox 

County (8).

Clay ton formation. Alabama, Barbour County (10).

Pine Barren member. Alabama, Wilcox County (11).

Porters Creek clay. Tennessee, Hardeman County (13, 14).

Midway formation, lower part. Arkansas, Saline County (18, 

19A, 20-22) ; Pulaski County (23).

Midway formation, upper part. Arkansas, Pulaski County (24, 

25).

Midway group. Texas, Van Zandt County (47).

Cibicides alleni (Plummer) Plummer 

Plate 18, figures 16, 17

Truncatulina alleni Plummer, Texas Univ. Bull. 2644, p. 144, pi.

10, fig. 4, 1927. 

Cibicides alleni Plummer, idem, Bull. 3201, pp. 54, 61 (lists),

1933. 

Cushman, Cushman Lab. Foram. Research Contr., vol. 16,

p. 73, pi. 12, fig. 19, 1940. 

Kline, Mississippi Geol. Survey Bull. 53, p. 61, pi. 6, figs. 21,

22, 1943. 

Cooper, Jour. Paleontology, vol. 18, p. 354, pi. 54, figs. 24, 25,

1944.

Test almost equally biconvex, the ventral side of most specimens being the more rounded; periphery subacute and bordered 

by a band of clear shell material, faintly lobate in its latest 

development; chambers 10-11 in last convolution, very coarsely 

punctate, previous whorls concealed by strong elevations of 

shell matter that follow the base of the chambers on the dorsal 

face; sutures on dorsal side marked by conspicuous elevations of 

transparent shell matter that taper and curve gently toward the 

periphery; sutures of the ventral face very slightly elevated 

and curving outward from the large smooth umbilical boss; 

aperture a large arched opening over the periphery and extending 

farther downward on the ventral side. Diameter up to 0.7 

mm.; average about 0.4 mm.—Plummer.

The types are from the Paleocene, shallow ditch at 

road corner southeast of new Corsicana reservoir on the 

road to Mildred, Navarro County, Tex.

Porters Creek formation, Matthews Landing marl member. Alabama, Wilcox County (2, 4-6).

Clayton formation, Chalybeate limestone member. Alabama, 

Sumter County (9) 

Pine Barren member. Alabama. Butler County (12).

Porters Creek clay. Mississippi, Kemper County (17). 

Midway formation. Arkansas, Saline County ("">) ; Pulaski

County (27) ; Clark County (30). 

Midway group, Texas, Limestone County (39) ; Navarro County

(40) ; Hunt County (41) ; Bastrop County (43) ; Travis

County (45).

Cibicides vulgaris (Plummer) Cushman

Plate 19, figures 7-11

Truncatulina vulgaris Plummer, Texas Univ. Bull. 2644, p. 145,

pi. 10, fig. 3, 1927. 

Cibicides vulgaris Cushman, Cushman Lab. Foram. Research

Contr., vol. 16, p. 73, pi. 12, fig. 21, 1940. 

Kline, Mississippi Geol. Survey Bull. 53, p. 63, pi. 6, figs. 19,

20, 1943. 

Cooper, Jour. Paleontology, vol. 18, p. 354, pi. 54, figs. 28,

29, 1944. 

Cushman and Todd, Cushman Lab. Foram. Research Contr.,

vol. 22, p. 65, pi. 11, figs. 22, 23, 1946.

Test almost equally biconvex, the ventral face I ^ing slightly 

the more elevated; peripheral margin broadly rounded, frequently somewhat lobate; chambers 7-9 in last wl orl, last two 

or three distinctly turgid; sutures marked by strong elevations 

of clear shell material curving gently toward t*ie periphery 

from a very high ridge of irregularly disposed mass of shell 

matter that follows the inner edge of the whorl an^ produces a 

more or less well-developed spiral on both faces; sh-ill wall more 

coarsely punctate than any other species in the fauna; aperture 

a long, arched slit extending from the periphery toward the 

umbilicus under a narrow lip. Diameter up to 0.6 mm.— 

Plummer.

The types are from the Paleocene, road cut near top 

of hill on Corsicana-Navarro Koad just south of junction with the Mildred Koad, Navarro County, Tex.

Clayton formation, Chalybeate limestone member. Alabama,

Sumter County (9).

Porters Creek clay. Mississippi, Kemper County (17). 

Midway formation, lower part. Arkansas, Saline C?unty (19A). 

Midway formation, upper part. Arkansas, Pulaski County (25). 

Wills Point formation. Texas, Travis County (SB) ; Caldwell

County (34). 

Midway group. Texas, Limestone County (39) ; Bastrop County

(43) ; Caldwell County (44) ; Van Zandt County (48).

Cibicides newmanae (Plummer) Cushman and Todd 

Plate 19, figures 12-14

Discorbis neiomanae Plummer, Texas Univ. Bull. 2644, p. 138,

pi. 9, fig. 4, 1927. 

Cibicides newmanae Cushman and Todd, Cushman Lab. Foram.

Research Contr., vol. 18, p. 46, pi. 8, fig. 16, 19 i2. 

Test slightly oval, dorsal face being somewhat convex and 

the ventral face fiat or slightly concave; peripl <iral margin 

sharp and thin; chambers smooth, distinctly punctate, moderately curved, increasing rather rapidly in size, usually 8 in final 

whorl; sutures on both sides curved and marked by tapering 

dark Lines; aperture a distinct slit extending from a point near 

the margin into the small excavated umbilical depression under 

an apertural lip. Greatest diameter up to 0.4 mm.; average 0.3 

mm.—Plummer.

The types are from the Paleocene, ditch along ElginAustin road 1.4 miles northeast of Littig close to the 

county line (Bastrop quadrangle), Bastrop County, 

Tex. "
ANOMALESFIDAE 67

Porters Creek formation, Matthews Landing marl member. Alabama, Wilcox County (1, 2, 4) ; Butler County (3) ; Choctaw 

County (7).

Porters Creek clay. Tennessee, Hardeman County (14).

Midway formation, lower part. Arkansas, Saline County (18).

Midway group. Texas, Hunt County (41).

Cibicides howelli Toulmin 

Plate 19, figures 15-17

Cibicides hoioelli Toulmin, Jour. Paleontology, vol. 15, p. 609,

pi. 82, figs. 16-18, 1941. 

Cushman and Renz, Cushman Lab. Foram. Research Contr.,

vol. 18, p. 13, pi. 3, fig. 10, 1942.

Cushman, Am. Jour. Sci., vol. 242, p. 18, pi. 2, figs. 21, 22, 

1944; Cushman Lab. Foram. Research Contr., vol. 20, 

p. 28, pi. 4, fig. 29; p. 50, pi. 8, fig. 9,1944.

Cibicides cf. C. pseudoungerianus Cushman and Garrett, Cushman Lab. Foram. Research Contr., vol. 15, p. 88, pi. 15, 

figs. 25, 26, 1939.

Test plano-convex, subcircular in outline, earlier 

whorls on dorsal surface obscured by growth of shell 

material, ventral surface very convex, almost conical, 

capped by an elevated, low, rounded, smooth umbonal 

boss of clear to translucent shell material, periphery 

narrowly angled in the early portion of the last whorl, 

becoming bluntly angled or rounded in the later portion; chambers 9 or 10 in number in the final whorl, 

increasing gradually in size as added; dorsal sutures 

moderately curved, ventral sutures more gently curved 

than the dorsal, early sutures on dorsal and ventral sides 

flush with the surface, later sutures slightly depressed; 

wall coarsely perforate; aperture a slit-like opening at 

the base of the final chamber, arching across the periphery and extending onto the dorsal side. Length up 

to 0.55 mm.; width up to 0.42 mm.; thickness up to 

0.24 mm.

The types are from the Eocene Salt Mountain limestone, from Richmond Branch, y2 mile north of Salt 

Mountain, Clarke County, Ala.

Porters Creek clay. Mississippi, Kemper County (17).

Porters Creek formation, Matthews Landing marl member. 

Alabama, Butler County (3) ; Wilcox County (5).

Naheola formation, Coal Bluff marl member. Alabama, Wilcox County (8).

Clay ton formation. Alabama, Barbour County (10). 

Pine Barren member. Alabama, Wilcox County (11).

'Midway formation, lower part. Arkansas, Saline County (22).

Midway formation, upper part. Arkansas, Pulaski County (24).

Midway formation. Arkansas, Saline County (26).

Midway group. Texas, Hunt County (41).

Cibicides blanpiedi Toulmin 

Plate 20, figures 1-5

Cibicides blanpiedi Toulmin, Jour. Paleontology, vol. 15, p. 609,

pi. 82, figs. 11-13, 1941. 

Cushman and Todd, Cushman Lab. Foram. Research Contr.,

vol. 18, p. 46, pi. 8, figs. 13-15, 1942.

Cushman, Am. Jour. Sci., vol. 242, p.' 18, pi. 2, fig. 20, 1944; 

Cushman Lab. Foram. Research Contr., vol. 20, p. 50, pi. 8, 

fig. 10, 1944.

Test subcircular, plano-convex, non-lobate, with acute peripheral angle; dorsal side flat, somewhat evolute, the final whorl

of chambers partially embracing the preceding whorl; ventral 

side very convex, subconical, involute; chambers distinct, eigfct 

or nine in the final whorl, on the dorsal side curved and unirflated, on the ventral side curved and strongly produced vehtrally, with the ventral end of each chamber extending beyond tha t 

of the preceding; sutures on the dorsal side distinct, flush witv 

the surface, moderately curved, somewhat limbate in the earl" 

portion of the test but becoming non-limbate in the latest portion, sutures on the ventral side distinct, flush with the surface, 

somewhat sigmoid; wall smooth, very finely perforate, polished ; 

aperture a slit on the dorsal side at the inner margin of the lart 

two or three chambers, arching across the periphery. Diameter up to 0.31 mm.; height up to 0.22 mm.—Toulmin.

The types are from the Eocene Salt Mountain limestone, in bed of Richmond Branch, Clarke County, Ala.

Porters Creek formation, Matthews Landing marl membe".

Alabama, Wilcox County (5) ; Choctaw County (7). 

Naheola formation. Coal Bluff marl member. Alabama, WJlcox County (8).

Cibicides browni Kline 

Plate 20, figures 6-8

Cibicides browni Kline, Mississippi Geol. Survey Bull. 53, p. 6^, 

pi. 7, figs. 18-20, 1943.

Test plano-convex, ventral side convex, dorsal side flat, frequently showing signs of attachment, periphery subacute brt 

not keeled; chambers usually 5 in the last-formed whorl, di^- 

tinct, increasing rapidly in size, inflated on ventral side; sutures distinctly curved on dorsal and ventral sides, depressed on 

ventral side; aperture near periphery, small.—Kline.

The types are from the Paleocene Clayton formatkxn, 

test hole M 49, surface to 8.7 feet below, 2% miles north 

of Pheba, Clay County, Miss.

Naheola formation, Coal Bluff marl member. Alabama, Wilcox County (8).

Porters Creek clay. Tennessee, Hardeman County (14).

Midway formation, lower part. Arkansas, Saline County (18, 

22).

Midway group. Texas, Hunt County (41).

Cibicides cf. C. williamsoni Garrett 

Plate 20, figure 9

Cibicides williamsoni Garrett, Jour. Paleontology, vol. 15, p. If 6,

pi. 26, fig. 15, 1941. 

Cibicides cf. C. williamsoni Cushman and Renz, Cushman Lab.

Foram. Research Contr., vol. 18, p. 14, pi. 3, fig. 12,1942.

Specimens from the Paleocene Soldado Rock formation, Soldado Rock, Trinidad, are smaller than autotypes with which they have been compared, but see^n 

to be related. The typical form is recorded from the 

middle Eocene of southern Alabama and Mississippi.

Cibicides cf. C. semiplectus (Schwager) Cushman and Ponton 

Plate 20, figure 10

Pulvinulina semiplecta Schwager, Palaeontographica, vol. 30,

Pal. Theil, p. 130, pi. 27 (4), fig. 16,1883.

Cibicides cf. C. semiplectus Cushman and Renz, idem, vol. 18, 

p. 14, pi. 3, fig. 11, 1942.

The figured specimen is from the Paleocene Soldado 

Rock formation, Soldado Rock, Trinidad, B. W. I. T/ *> 

specimen is not entirely typical as the last-formed
68 PALEOCENE FORAMINIFERA OF GULF COASTAL REGION

chamber has not reached the full development that 

sometimes occurs in this species.

This form is not the same as that recorded as G. semiplectus from the Eocene Wilcox group of Alabama and 

later found to be Amrnocibicides pontoni Earland, an 

arenaceous species belonging in the Trochamminidae.

Cibicides miriflcus Cushman and Bermudez 

Plate 20,'figures 14-16

Cibicides miriflcus Cushman and Bermudez, Cushman Lab. 

Forani. Research Contr., vol. 24, p. 88, pi. 16, figs. 1-3, 1948.

Test compressed, nearly equally biconvex, ventral 

side slightly more convex than the dorsal side, periphery angled, subacute, ventral side with a small but 

distinct umbonal boss, dorsal side with the central portion depressed in a close spiral; chambers distinct, 11 

to 13 in the adult whorl, of uniform size, very gradually 

increasing in size as added, little if at all inflated; 

sutures distinct, not depressed except the last few on 

the ventral side, slightly curved; wall smooth, finely 

perforate; aperture at the base of the last-formed 

chamber, a small but distinct raised opening at the 

periphery, then extending over onto the dorsal side, with 

a very slight lip near the periphery. Length of holotype 0.52 mm.; breadth 0.47 mm.; thickness 0.23 mm.

The types are from the Paleocene Madruga formation, San Juan y Martinez, Pinar del Rio Province, 

Cuba.

This species differs from G. madrugaetisis Cushman 

and Bermudez in the more compressed test, sharper 

peripheral angle, larger number of chambers, and larger 

proportion of the dorsal side taken up by the lastformed whorl.

Cibicides reprimatus Cushman and Bermudez 

Plate 20, figures 17, 18

Cibicides reprimatus Cushman and Bermudez, Cushman Lab. 

Foram. Research Contr., vol. 24, p. 87, pi. 15, figs. 13, 14, 

1948.

Test plano-convex, dorsal side flattened, ventral side 

strongly convex, periphery subacute, ventral margins 

slightly concave, ventral side with a slight umbilical 

boss; chambers fairly distinct, especially in the later

portion, increasing very gradually in size as added, on 

the dorsal side with a distinct oral depression in the 

middle of the inner margin; sutures fairly distinct, 

very slightly depressed in the later portion of the ventral side and all of the dorsal side, ventrally curved, 

slightly sinuous, dorsally irregular with distinct lobular projections; wall smooth except for the depressions 

of the dorsal side; aperture at the peripheral margin 

extending onto the dorsal side, with a slight lip at the 

peripheral portion. Length of holotype 0.77 mm.; 

breadth 0.70 mm.; thickness 0.40 mm.

The types are from the Paleocene Madruga formation, under highway bridge on Central San Antonio, 

Madruga, Habana Province, Cuba.

This species is peculiar in its dorsal side, differing 

from G. madt^ugaensis Cushman and Bermudez in the 

more plano-convex form, very small umbilical boss, 

and the indentations of the dorsal side.

Cibicides madrugaensis Cushman and Bermudez 

Plate 21, figures 1-3

Cibicides madrugaetisis Cushman and Bermudez, Cufhman Lab. 

Foram. Research Contr., vol. 24, p. 87, pi. 15, figs. 10-12, 

1948.

Test rather small, nearly equally biconvex, ventral 

side slightly more convex than the dorsal, periphery 

angled, but slightly rounded, ventral side distinctly umbonate; chambers fairly distinct, little if at all inflated, 

6 to 8 in the final whorl, of rather uniform size and 

shape; sutures distinct, strongly curved, little if at all 

depressed, distinctly limbate on the ventral side; wall 

smooth, finely perforate; aperture narrow, at the basal 

margin of the last-formed chamber, extending very 

slightly on the ventral side but nearly the length of the 

chamber on the dorsal side. Length of hole type 0.40 

mm.; breadth 0.37 mm.; thickness 0.25 mm.

The types are from the Paleocene Madruga formation, under highway bridge on Central San Antonio, 

Madruga, Habana Province, Cuba.

This species differs from C. alleni (Plummer) in the 

smaller size, more umbonate ventral side and smooth 

dorsal side.
INDEX

[Italic numbers indicate descriptions

A Page

Abstract-—_———————————————————_--___-..__.________ 1 

abuillotensis, Colettes...——.„_.—_________________ 54, 60 (tab.), pi. 24 

acervulinoides, Planoglobulina..........._„_____._.__._______ 38, 44 (tab.)

Acknowledgments for aid—.____________________________ 1 

aculeata, Bulimina......„_.___________________________ 40

Dentalina.........———————..„_.._.______.__ 91, 28 (tab.), pi. 6

Ramulina——............____.______________ 36, 44 (tab.), pi. 10

acuta, Anomalina—............................................. 60 (tab.), 69, pi. 18

Anomalina ammonoides........__-____. -______________:._ 62

Globorotalia wilcoxensts______________________ 60 (tab.), 61, pi. 17

Polymorphina..._________'________________________ 32

acuticosta, Lagena............._-__.__.____________ 28 (tab.), 32, pi. 9

Adhaerentia midwayensis..........-——_——„_.__—______ H, 12 (tab.), pi. 3

aequa, Globorotalia crassata..—.—__-____._________ 60 (tab.), 61, pi. 17 

aequilateralis, Oyroidina......—-___-____._________ 51, 60 (tab.), pi. 14

Rotalia......... _______________'___________________ 51

affinis, Nodosaria.—.——————_—..._.__________ %S, 28 (lab.), pi. 7 

agglutinans, Textularia.———...„____-________________. __ 8 

Alabama, localities..————————————.——.—____________ 1-2

alabamensls, Dentalina-......._—__—__._—_________ 81, 28 (tab.), pi. 6

Dorofhia......———— -_______-____________-__ 9, 12 (tab.), pi. 2

Quingueloculina...............__-_.______________ 10, 12 (tab.), pi. 2

Rectogumbelina___________________________ 58, 44 (tab.), pi. 11 

Robulus.............__-_......__...._____...._.... 15, 28 (tab.), pi. 4

Schenckiella.. __-____________________-___ 10, 12 (tab.), pi. 2 

Spiroloculina.....————.................................... 11, 12 (tab.), pi. 3

Triloculina-....-_______-__..._..._____.____. 11, 12 (tab.), pi. 3

Virgulina........______.__-____._________________ 42

Alabamina wilcoxensis————————— ___________ 67, 60 (tab.), pi. 16

ivilcoxensis limbata—..................____________ 57, 60 (tab.), pi. 16

alazanensis spissiformis, Anomalina.............-........_„_.._.__... 64

albeari, Globorotalia.............................................. 60 (tab.), 61, pi. 24

alexanderi, Ellipsonodosaria...................................... 44 (tab.), 47, pi. 13

atteni, Cibicides............................................... 60 (tab.), 68, 68, pi. 18

Truncatidina...................—...—.....—....................—..—. 66

Allomorphina globulosa.......................................... 69, 60 (tab.), pi. 17

hatti-.............——.———.——-..—........................... 59

pakocenica.................................................. 58, 60 (tab.), pi. 16

subtriangularis........————.—————...__-_.____ 69, 60 (tab.), pi. 17

trigona.—............................._______________________ 58

allomorphinoides, Discorbina.................................................. 50

Discorbis....................—..—......———..—.............. ____ 50

Valvulina................................................................. 50

Valvulineria.......................................______ 50, 60 (tab.), pi. 14

alterans, Pleurostometta...———————————.--__„________.__.___ 45

Pleurostometta brevis...................................................... 45

atternata, Dentalina——..... _„_________________________ 23

Ammobaculites expansus.......................................... 4, 12 (tab.), pi. 1

midwayensis...———————.——-———___—_____ 4, 12 (tab.), pi. 1

paleocenicus.—..............—..________________ 4, 12 (tab.), pi. 1

Ammodiscidae————————————————__--_.—___________ 4,12 (tab.) 

Ammodiscus cretaceus.—...——._——-—__-_.________ 4, 12 (.tab.), pi. 1 

Ammomarginulina sp-—„—_—_—_._______________ 5, 12 (tab.), pi. 1 

ammonoides acuta, Anomalina................................................ 62

amphioxys, Nodosaria...................—...............____ 94, 28 (tab.), pi. 7

Amphisteginidae.—.————————————„__._—___________ 56, 60 (tab.) 

amygdaloides terquemiana, Polymorphina...................................... 34

angularis, Clavulina........................................................... 8

Angulogenna virginiana......................................... 44 (tab.), 45, pi. 12

wilcoxensis............................__________. 44, 44 (tab.), 45, pi. 12

angusta, Pullenia quinqueloba........—......................... 59, 60 (tab.), pi. 17

Anomalina acuta—.............___.__.____________ 60 (tab.), 6%, pi. 18

alazanensis spissiformis-...........—........._____________ 64

ammonoides acuta...................................____________ 62

basilobata.................................................... 60 (tab.), 63, pi. 18

clementiana..................—.....—..-................... 60 (tab.), 63, pi. 18

assimilis......................________________ 60 (tab.), 63, pi. 21

cubana............ ._______-___.____________ 60 (tab.), 64, pi. 21

grosserugosa.—........................................................... 64

insecta.....................................................________ 37

Anomalina acuta—Continued Page 

madrugaensis................................................ 60 (tab.), 64, pi. 22

martinezensis................................................ 60 (tab.), 64, pi. 21

midwayensis............................................. 60 (tab.), 68, 64, pi. 17

trochoidea-—...........................—..—..—— 60 (tab.), 68, pi. 18

praespissiformis...................................—........ 60 (tab.), 64, pi. 22

umbonifera..—-'—-.—.................................... 60 (tab.), 62, pi. 17

wetteri...—..................——..————.—.————— 60 (tab.), 63, pi, 18

sp. A—.———_-____-_____-_______-____ 60 (tab.), 64, pi. 18 

sp. B_.,_._....._._-_________-____-______ 60 (tab.), 64, pi. 18

Anomalinidae—.....____„____________—.-———————— 60 (tab.), 62

apiculata, Entosolenia........................................... 4%, 44 (tab.), pi. 12

Lagena...................._._-.———————————————————————— 42

applinae, Loxostomum.-............................————— 4$, 44 (tab.1, pi. 12

applini, Bolivina..—.—.............——..—————————-————— 43

archiaciana, strigillata, Frondicularia.........................——————— 30

arkadelphiana midwayensis, Bulimina............................ $, 44 (tab.), pi. 11

Arkansas, localities—————————————————————————————————— 2-3 

arkansasana, Textularia plummerae..........———.—————. 7, 12 (tab.), pi. 2

arkansasanum, Chrysalogonium........—.....-...—...——.—.. US, 28 (tab.), pi. 7

arkansasanus, Robulus——————.............—......———— 16, 28 (tab.), pi. 4

assimilis, Anomalina clementiana—.........———————— 60 (tab.), 63, pi. 21

Astacolusjugleri........................................——.....—....——. 16

Asterigerina primaria—.—————..————.._—————— 56, 60 (tab.), pi. Ifi

attenuata, Nodosarella.....——.......—...............———... 44 (tab.), 45, pi. 12

Ellipsopleurostomella...................———.———.———-— ————— 4ff

basilobata, Anomalina.....————...——.________————. 60 (tab.), 65, pi. If

Bathysiphon eocenicus.....——....——..——...—————— 3, 12 (tab.), pi. 1

baudouiniana, Frondicularia.......................—...——...—.............. 3P

blanpiedi, Cibicides........————.....—....——_———-— 60 (tab.), 87, pi. 20

Bifarina eleganta......—.—.——————————————————— 3f

carinata——....——....——...........—...—......——.. 60 (tab.) 65, pi. 22

Boldia carinata..___...________.__._.——————————— 60 (tab.), 65, pi. 22 

cubensis madrugaensis.——...————.——————— 60 (tab.), 65, pi. 22

vanderslutsi——————-..———.....——————— 60 (tab.), 65, pi. 23

BolMna applini...................————-——.————————— 43

budensis........—————....————————...—————— 43, 44 (tab.), pi. 12

crenulata————...... ... ... ... ... ... ...——...—...—— 4$, 44 (tab.), pi. 12

frondea............——......——————.—————..—.———————— 35

midwayensis....—.._________..„.————.——————— 43, 44 (tab.), pi. 12

sp. A..—————————————————— 4$, 44 (tab.), pi. 12

sp. B_—._____...___-__-_____—-____—- 4S, 44 (tab.), pi. 12 

Bolivinopsis sp._———_—————.————————————————————— 27, 44 (tab.) 

brantlyi, Hemicristellaria............———.—.-——..-—.—--..——-—-—-- 17

Vaginulinopsis—..........—.—————.————————.——-—--— 17

brevis alternans, Pleurostomella....—.——.————..—.—————..———...... 45

browni, Cibicides..———————-———————————.———— 60 (tab.), 67, pi. 20

budensis, Bolivina.-....————..——————...............—— 43, 44 (tab.), pi. 12

Fhbellina..—..——————.——....__—-——.—————— 29

Frondicularia.——————————————— ———————————— 2 1)

Palmula--..————.———————————-——.—- 28 (tab.), 39, pi. 7

Bulimina aculeata... ————— ——————— ———————————————————— 40 

arkadelphiana midwayensis.....—.——————..——— 40, 44 (tab.), pi. 11

cacumenata—————————.————————— 40, 44 (tab.), pi. 11

(Desinobulimina) quadrata—____———-————————— 41, 44 (tab.), pi. 11 

(Jjyiipsobulimina) quadrata....—————————————————————————— 41

kickapooensis.........——-.———.———————————--——— 41

kugleri....____________-____________-__ 41, 44 (tab.), pi. 11

BulimineUa elegantissima———........—————.—————.... 39, 44 (tab.), pi. 11

Buliminidae... ___-_______————————————-————-—— 39; 44 (tab.) 

BuUopora chapmani.———..———————....—————— 36, 44 (tab.), pi. 10

chapmani hispida——-..................................... 36,44 (tab.), pi. 10

laeais...______-____________-_________ 36, 44 (tab.), pi. 10 

hispida———————————————————— 36, 44 (tab)

byramensis, Cornuspira... ——————————————————————— If, 12 (tab.), pi. 3

C

cacumenata, Bulimina- ______—-——————————————— 40, 44 (tab.), pi. 15 

Cancris mauryae.———........—————...._—._————— 66, 60 (tab.), pi. 11

69
70 INDEX

Page

carinata, Boldta___________________-_________ 60 (tab.), 65, pi. 22 

Cristellaria midwayensis--...--- — --—-..- — - — —......_-._--_--..__ 13

carinata expansa, Textularia.... ___.___________.________ 5

carinatus, Robulus midwayensis.___—.._._______.__ IS, 28 (tab.), pi. 3 

Cassidulinidae_________________________________ 56, 60 (tab.)

caudigera, Pseudoglandulina.-. ______.__._._______ 86, 28 (tab.), pi. 7

Ceratobulimina perplexa. ____________________._ S6. 60 (tab.), pi. 16 

Chalybeate limestone member, localities_____ _____..________ 1,2 

chapmani, Bullopora.... _______________________ 36, 44 (tab.), pi. 10

hispida, Buliopora........ ....____.... ..... __________ __S6, 44 (tab.), pi. 10

Vitrewebbina______-_..___.._-_____-.________._____________ 36

Chilostomella subtriangularis..........__________.____________ 59

Chilostomellidae____________..—_-_____......____ 58, 60 (tab.)

Chilostomelloides eocenica... ___________________ 59, 60 ^ab.), pi. 17

Chrysalogoniumarkansasanum.. __________._.______ 85, 28 (tab.), pi. 7 

eocenicum..——...................______________ 85, 28 (tab.), pi. 7

granti..-.-.--............................................... 84, 28 (tab.), pi. 13

texanum......__....___..._____________._.____.___ 25

CibieidesaUeni............................................... 60 (tab.), 66, 68, pi. 18

blanpiedi..-.-.............................................. 60 (tab.), 67, pi. 20

browni— ___.....____.._____....____..._.. 60 (tab.), 67, pi. 20

hotoeUi...................................................... 60 (tab.), 67, pi. 19

madrugaensis....-.-.--..-._._._.._._____........ 60 (tab.), 68, pi. 21

mirificus—................................................. 60 (tab.), 68, pi. 20

newmanae. __..................___........._........ 60 (tab.), 66, pi. 19

praecursorius................................................ 60 (tab.), 65, pi. 19

pseudoungerianus....................................._ ___....__._. 67

reprimatus-..-........................................._.. 60 (tab.), 68, pi. 20

semiplectvs....... _________.___......___..... 60 (tab.), 67, pi. 20

vanbelleni................................................................. 63

vulgaris..........................................._..__.. 60 (.tab.), 64, 66, pi. 19

wiUiamsonL................................................. 60 (tab.), 67, pi. 20

Clavulina angularis............................._-._..__..--..._...____ 8

Clavulinoides midwayensis........................... _.___..___. 8,12 (tab.), pi. 2

Clayton formation, localities...._...___...._.._._......__.._.. 1-2

dementiana, Anomalina...................... ................... 60 (tab.), 63, pi. 18

assimilis, Anomalina....................................._ 60 (tab.), 63, pi. 21

Rosalina............................ ......__.____...._____.__ 63

Coal Bluff marl member, localities_..__.._.__......_.__..... 1

cold, Dentalina................................................... 19, 28 (tab.), pi. 6

Colettes abuillotensis.----................. ............_._.... 54, 60 (tab.), pi. 24

guatemalensis................................ ._____ ._ 54, 60 (tab.), pi. 24

pasionensis—................................................ 54, 60 (tab.), pi. 24

reticulosus....................................____...._ 64, 60 (tab.), pi. 15

comata, Nodosaria (Olandulina)............................................... 26

Pseudoglandulina......................................................... 26

comis, Robuhispseudo-costatus—.................................. 14, 28 (tab.), pi. 4

compressa, Globigerina................................ ...____ 60, 60 (tab.), pi. 17

Cornuspirabyramensis.-----.--............................. _____ 1$, 12 (tab.), pi. 3

costata, Lagena................................................... 28 (tab.), 38, pi. 9

Marginulina.............................................................. 20

crassata aequa, Oloborotalia........................... ........._ 60 (tab.), 61, pi. 17

crenulata, Bolimna........ _______________._.._._.__._ 43, 44 (tab.), pi. 12

cretaceus, Ammodiscus............................................ 4, 12 (tab.), pi. 1

cretosa, Spiroplectammina laems.......................... ______.. e, 12 (tab.), pi. 1

Cristellaria degolyeri........................................................... 14

earlandi............. ........._........._...._.._.._______......._. 17

hamata.......................................___.._. ..................... 18

longiforma ...................................____________________ 28

midwayensis.............................................................. 13

carinata............................................................... 13

orbtcularis pseudo-costata—........................... ___________________ 14

pseudo-mamilligera................. ________________________________..... 13

subaculeata tuberculata........................... ........___._.._______. 17

siiblatifrons..............................._..._.._._... ________________. 27

trigonata.............................._.____._.._..____________________. 27

turbinata..................................... ______________________ 14

crumenata, Entosolenia.......................................... 41, 44 (tab.), pi. 12

cubana, Anomalina.............................................. 60 (tab.), 64, pi. 21

I'alvulammina nassauensis.............................._.___ 9, 12 (.tab.), pi. 21

cubensis, Boldia....._.______._____.__..........__ 60 (tab.), 65, pi. 22

3>..ari.iHa................................................... w, 12 (tab.), pi. 2

cutter mexicana, PuMnulineUa................................_.___._____ 53

midwayana, Pulvinulinella—.............................................. 53

Truncatulina............,__...._____...___...._....___.... 53

cushmani, Polymorphina........................................ 35, 44 (tab.), pi. 10

cylindroides, Pyrvlina............................................. SS, 44 (tab.), pi. 9

D 

deadericki, Loxostomum.......................................... 44, 44 (tab.), pi. 12

deadericki exllis, Loxostomum.................................... 44, 44 (tab.), pi. 12

deca THtocu/ina.................................................. li, 12 (tab.), pi. 3

Page 

degolyeri, Cristellaria.......................................................... 14

Lenticulina........................ — .-..-..—.-...--.....-..- — -.-...-... 14

Robulru...................................................... 14, 28 (tab.), pi. 3

delicatissima, Frondicularia................................................... 29

Palmula........................——......................... 28 (tab.), 89, pi. 7

delicatula naheolensis, Dentalina........-......—— ....-....................... 20

Dentalina aculeata............................... ...___-.__--.- 81, 28 (tab.), pi. 6

alabamensis-----------...................—............... 81, 28 (tab.), pi. 6

alternata.-.---.-.-.. —.......-.-.—...-.......—.-........................ 23

cofci——— —— —— — — —— —— ————— — ————— 19, 28 (tab.), pi. 6

delicatula naheolensis.-...-.......—....................................... 20

eocenica............_____ —— ______-_____--_-. 88, 28 (tab.), pi. 6

gardnerae..................................................... 19, 28 (tab.), pi. 6

granti..-..-. — --. —— ————————————— —— — ——— 24

inepta...................—...........-..... —— ............. 81, 28 (tab.), pi. 6

insulta....--........ — .....—.. —........—...... —........ 80, 28 (tab.), pi. 6

mucronata—-.............................................. 88, 28 (tab.), pi. 6

naheolensit..............................-...— --.---........ 80, 28 (tab.), pi. 6

nasuta......m........_---.--.--..-.-.-_-..-....._...__ 88, 28 (tab.), pi. 6

pauperata......................................... ——...... 88, 28 (tab.), pi. 6

plummerae................................................... 80, 28 (tab.), pi. 6

pseudo-nasut a............. — —................-„.——.—— 81, 28 (tab.), pi. 6

pseudo-obliquestriata........—.............--....-....-....... 19, 28 (tab.), pi. 6

fungens......._ — -.---- — — — — .-— — - — -.. -... — —— — — - 21

wilcoxensis..--.----.-----.---—---------------------- 81, 28 (tab.), pi. 6

sp. A.---... — ———— ————— —— —— ——— _.... 21, 88, 28 (tab.), pi. 6

sp. B...-——————— — —— —— ————— 20, 83, 28 (tab.), pi. 6

sp. C.___......——————— ————————————— 83, 28 (tab.), pi. 6

sp..-...-. — — ——— ——— ———— —— ——— ———————— ——.-— 21

Desinobulimina-----....- — _——— — — __ ——.——— — —. ——....-- 41

Discorbina allomorphinoides......--...--.......-- ———————————. — —— 50

praecursoria............................................................... 65

umbonifera...... .__._..___.._--—-_-__---_-—_-.—..---..--- 62

Discorbis allomorphinoides—.—.- — — ..--..... —.....--..............—--...... 50

infrequent.......'........................... —— ............. 48, 60 (tab.), pi. 13

midwayensis——............................................. 48, 60 (tab.), pi. 13

soldadoensis.---—.. —————————————————— 48, 60 (tab.), pi. 13

trinitatensis—........................................... 48, 60 (tab.), pi. 13

newmanae.............___.____-.-._-_-__........ —.— ——-. 66

dissentius, Robulus wilcoxensis.-.................................. 15, 28 (tab.), pi. 4

distincta, Lituolaerecta.. ___.......„.— —.—.. ——— ————————- 5, pi. 1

Marginulina..... ————— —— —— —— —— ————————— 18, 28 (tab.), pi. 21

Dorothia alabamensis.--. ____.___-...-._____._____ 9, 12 (tab.), pi. 2 

dubia, Marginulina. —— ————————————— — — —— — 18, 28 (tab.), pi. 5

E

earlandi, Cristellaria..--- ____.._-.--_.--___-_._---_.___-__..-------.--.. 17

Marginulina_________.---._-_.... —... — _----_.. 17, 28 (tab.), pi. 5

elegans, Vaginulina legumen................ ....-....—..-..--......-........ 19

eleganta, Bifarina.................................. ....-. —... —— — --..-.. 39

Siphogenerina—-.... ....... ——__..._ —. — --..-—-_.——. ———.. 39

Siphogenerinoides.....—.. ____.._._-.-.____.... 55,44 (tab.), pi. 11

elegantissima, Buliminella.....______._____-______ 3ft, 44 (tab.), pi. 11

elevata, Truncatulina—-....--.... ..--._ ——.--_----..-. — ...... —— — —... 52

elevatus, Eponides................. — ......__.-.-__.____ 52, 60 (tab.), pi. 14

Ellipsobulimina........................—.—...._. — ---...——.. ——....... 41

Ellipsoidinidae............... ____________________........44 (tab.), 45

Ellipsonodosaria alexanderi....................................... 44 (tab.), 47, pi. 13

granti... ...______________——- — — — -—— —_._---...---. 24

midwayensis.............. ___ — _--._-.__--._-_--__.... 44 (tab.), 47, pi. 13

paleocenica................-._._.--__--- — __.--.-.__ 44 (tab.), 46, pi. 13

plummerae................... — .—.......................... 44 (tab.), 46, pi. 13

sagrinensis..-... ..--..----_. ——....--..---..... — ——._———. —...... 46

spinulosa..........................— .......-............................. 47

sp..—————— —— ——————— —— ——— — .- — —— —— —— —— — ..- 46

Ettipsopleurostomella aftenuata.........-.... —.......— ...........—.....---— 45

elongata, Polymorphi-iiella.—......... — ...-...-...______ 36, 44 (tab.), pi. 10

Entosolenia apiculata.................. .______________.. 4$, 44 (tab.), pi. 12

crumenata................................___._______ 41, 44 (tab.), pi. 12

laevigata—.................................................. 41, 44 (tab.), pi. 12

morsei——...._.._....__ — ---..-_...-_..___.. J$, 44 (tab.), pi. 12

orbignyana................................. — ............... 41, 44 (tab.), pi. 12

sp_————--——————————————— —————— 4S, 44 (tab.), pi. 12

eocenica, Chilostomelloides........................................ 59, 60 (tab.), pi. 17

Dentalina....................———. ———————————_ 88, 28 (tab.), pi. 6

Epistomina—... ___.________..._____.___ 55. 60 (tab.), pi. 15 

Textularia................................................................ 7

eocenicum, Chrysalogonium....................................... 8". 28 (tab.), pi. 7

eocenicus, Bathysiphon—.......................................... S, 12 (tab.), pi. 1

Eouvigerina excavata............................................. 38 44 (tab.), pi. 11

Epistomina eocenicu............................................. 55, 60 (tab.), pi. 15
INDEX 71

Page 

Epiitominoides midwayensis...... ......_._________._._ 58, 60 (tab.), pi. 16

wilcoxensis................__.___._.__.._____._..... 58, 60 (tab.), pi. 16

Eponides elevatus______....___.____........__........_ 58, 60 (tab.), pi. 14

gratiosus.—.... .____....______...._____________ 63, 60 (tab.), pi. 23

haidingeri——-.— ...--_.-_..---___-_..--________.________________...._ 52

lotus.......-._-_-:----_-_-_---__---_____.___________________ 52, 60 (tab.), pi. 14

plummerae-.--. __._.__.__..._________...___ SS, 60 (tab.), pi. 14

tenera.-... ___....__......__._.._....__________________.. 52

vanbelleni.. _______.______.________________. 53, 60 (tab.), pis. 20, 23

sp_... —....... — --.___.-„--______._______.___.__.___..____________.___ 52

erecta, Lituola........ __............_._......_______________... 5, pi. 1

erecta distincta, Lituola..... _____..__.___________________________________ 5, pi. 1

excavata, Eouvigerina. ....._______._........._._-..._______ 38, 44 (tab.), pi. 11

Haplophragmoides........___________.______________ ..4, 12 (tab.)

exigua, Pseudoparrella............. .____._-_____-___-.__.... 57, 60 (tab.), pi. 16

exigualimbata, Pulvinulina............. ___-._...-..._...----._....___ 57

Pulvinulinella........... ____________________________________ 57

exigua obtusa, Pulvinulina—....... ____.___...___,________ _____________ 57

Pulvinulinella............. .._....___._.____.._____._._______... 57

exilis, Loxostomumdeadericki.-----............ .-...-..___._ 44, 44 (tab.), pi. 12

eximia, Marginulina........................... _______________ 17, 28 (tab.), pi. 5

expansa, Parrella................. ______....___.. ...__S3, 60 (tab.), pi. 16

Textularia carinata...-----------... ....--.....---.-...—--...__.-.__.. 5

expansus, Ammobaculites_--.._------_-___.....______.._.__ 4, 12 (tab.), pi. 1

extensa, Polymorphina..._._._..___.____._______...______ 34

Pyrulina............._....... _______.____.________. 84. 44 (tab.), pi. 9

Valvulineria............. ......______..___......__ 50, 60 (tab.), pi. 23

F 

ferussacii, Ouinqueloculina. .._.............___----...-....--..------..... 11

FlabeUina buden&is—.......... _........________-_-._____--.-.____-__--.._ 29

interpunctata_....-------_-_.....___............................... 30

projecta.......... ...._.___...._....._._._--..._._.---....__._-..-- 30

rugosa... ................———.......—..................__......... 29

flectata, Quinqueloculina plummerae............. ..........__._ 11, 12 (tab.), pi. 2

frankei, Frondicularia.............. ._____....___.._.. 28 (tab.), 30, 31, pi. 9

frondea, Bolieina.............___....___....____......____. 35

Polymorphina............................. ...__.._____ 85, 44 (tab.), pi. 10

Frondicularia archiaciana strigillata..____.___.-..--.....-..___..-.__. 30

baudouiniana................________..---......-----_....-_-._. 30

budensis.-.............___..____......_.._..__.__..._.___ 29

delicatissima....................... ..___...__....___.....___ 29

frankei-.--................................................28 (tab.), 30, 31, pi. 9

goldfussi—.................................-. —— ........... 28 (tab.), 31, pi. 8

interpunctata..... ........__.........__.........._.-.__----._._ 30

midwayensis—....—..... _________....__...___ 28 (tab.), 30, pi. 9

naheolensis....... ...._._______.__-__....._.. 28 (tab.), 30, pi. 8

oldhami....................____________....___________ 29

projecta........ ._..__.__________.________._____ 30

rugosa......... ........__....__.............._.__--.-.-_.._-_..... 30

sp. A........._..„...--...--......-.--....--..-.......-.... 28 (tab.), SI, pi. 9

O

gardnerae, Dentalina-... _.__........_.__................. 18, 28 (tab.), pi. 6

Marginulina-..-..- ..._._......__._........._-------._----_.-- 19

Gaudryina minima.... .....__......__........._..........__....... 8

oxycona....__........___.._..........__.............__... 9

Tudtta....................................----..-....... -.-.- 8, 12 (tab.), pi., 2

rugosa.

soldadoensis.-.. _____.___.______._____._.____ 8, 12 (tab.), pi. 2

sp——____._--_-.____..-._.___.__..„..____________..__.____._._ 8, 12 (tab.)

gibba, Globulina. ..... — ......................................... S3, 44 (tab.), pi. 9

glabra, Marginulina------.............. .________________________ 18, 28 (tab.), pi. 5

Glandulina manifesta. ..............__-_._.______._._________._......... 25

pygmaea.-..........___.........._...._..._.___._....._......_.. 26

Globigerina compressa.............................-- — ........... 60, 60 (tab.), pi. 17

pseudo-bulloides...-...._..._.__.......___......_______ 60, 60 (tab.), pi. 17

triloculinoides--........ ________________.____.__... 60, 60 (tab.), pi. 17

Globigerinidae_....-__-____----.-___..........________________________ 60, 60 (tab.)

Globorotaliaaibeari—....... ____________________ 60 (tab.), 61, pi. 24

crassata aequa.............._.......___.._______._._ 60 (tab.), 61, pi. 17

membranacea... __...._.....................___._._. 60 (tab.), 61, pi. 17

wilcoxensis acuta—........... .....-.-.-.-...-........_......- 60 (tab.), 61, pi. 17

Globorotaliidae...----.-_-_-------.___--__-____________..............*. 60 (tab.), 61

Globulina gibba.... .___.._______________________._ 33, 44 (tab.), pi. 9

minuta.......... .....____________...______.... S3, 44 (tab.), pi. 9

rotundata-... ....—....-.-.................................. 33,44 (tab.), pi. 9

globulosa, AUomorphina. ._..__....._.......___.____.___...._ 59, 60 (tab.), pi. 17

goldfussi, Frondicularia............. .___.__.__..__.... 28 (tab.), SI, pi. 8.

Nodosarina.....___......_....___.._._..._____.......... 31

gracilis, Vaginulina...............................__._______ 27, 28 (tab.), pi. 8

Page 

granti, Chrysalogonium.........................—............ ... 24, 28 (tab.), pi. 13

Dentalina............... _____....___.-___-__-___._________ 24

Ellipsonodosaria...-----..------- __-——.----.-„--..._-.„-.-.——— 24

Nodosaria—-___.___.._.__..-.----.---.-._--__.-.__....... 24

gratiosus, Eponides................................ ..._____. S3, 60 (tab.), pi. 23

grosserugosa, Anomalina.--..—......................... ___________.__ 64

guatemalensis, Colettes__....._..__...___._._----._______. 54,60 (tab.), pi. 2.

Gumbelina midwayensis.-.-----................. ...________. 37, 44 (tab.), pi. 11

morsei.________..______.___________...__ 38, 44 (tab.), pi. 11 

trinitatensig..............................--..... — ........... 38, 44 (tab.), pi. 11

Guttulinahantkeni--...——.... .............._...-- — — .-___ 32,44 (tab.), pi..

problema..............................................—.... SS, 44 (tab.), pi. £

wilcoxensis.............................. ._._ —....___1 32, 44 (tab.), pi. t

Gyroidina aequilateralis.... ...___.______.....______ 51, 60 (tab.), pi. 14

madrugaensis..... _........._____---_..---_--____---- 52, 60 (tab.), pi. 24

subangulata....-..... ........_____.._........._... 51, 60 (tab.), pi. 14

H 

haidingeri, Eponides_.__.__....__———.......-.-__. — ....————— 52

halli, AUomorphina............................................... ...--------- 59

hamata, Cristellaria......--....-...-....———......-.--_..--.--„. ——— — 18

Maiginulina.....................__...........__........ 18, 28 (tab.), pi. 5

hantkeni, Guttulina............................................... 32, 44 (tab.), pi. 9

Haplophragmoides excavata...-............----.-... ...—_.----._-..-..- 4, 12 (tab.)

havanensis, Marginullna..--...-...-.-....— - — ................... 18, 28 (tab.), pi. 5

Rotalia-.——................................................ 56, 60 (tab.), pi. 16

Hemicristellaria brantlyi................................--...-...........—... 17

longiforma................................-........ ....—..-- — ————— 2P

subaculeata tuberculata.................... ... ——— —..... ———— —— -. 17

herberti, Trifarina...................................... —— ...... 44 (tab.), 45, pi. 12

herricki, Valvulineria....................................... __..._—-—— 50

Heterohellcidae.___........___....... — -—..-- — .„. — .--—.-. 37, 44 (tab.)

hispida, Bullopora chapmani-. _..._...--......„.,--..__-- 86, 44 (tab.), pi. 10

Bulloporalaevis .................................................... 36, 44 (tab.)

howelli, Cibicides—............................................. 60 (tab.), 67, pi. 19

humilis, Nodosaria.............................. ..... — -_-_-.. ———— ——... 25

indentata, Marssonella.. --------- ... .... ...... ... ....... .... .. — ..... — 9, 12 (tab.)

inepta, Dentalina—.-.. ........... ... ... . _ -...—-—... _ SI, 22, 28 (tab.), pi. 6

inflata, Trilocuhna..- ______ — _ ———— . — -.-- —— —————— — — 11

infrequens, Discorbis- ... .... ....... ...... ... ........... .... . ..... 48, 60 (tab.), pi. 13

inornatus, Robulus pseudo-costatus ..... —— ——— -------- — .-..- —————— .. 14

insecta, Anomalina.- ... . ... ........ . ——— . ———— - — — -_._ ——————— -- 3?

insueta, Valvulineria—.----- — ..... ... - ...... ................ 61, 60 (tab.), pi. Vinsulsa, Dentalina.— ... .............. ...... ....................... 10, 28 (tab.), pi. 6

insulsus, Robulus- ____ - ___ ... . ______ - _ ---- __ ----- 16, 28 (tab.), pi. 5

interpunctata, Flabettina. . . . ...... .. ... .. ....... ..... .... .......... ...... ..... 3.

Frondicularia... ... --......- ... .. .......... .. .... .. .... ...... ... — .. — .. 30

jugleri, Astacolus

K 

kickapooensis, Bulimina —.. ------------ — ----..---.---.--....- — -- ——— -- 41

Kincaid formation, localities— ------------- — ------ — - —— ...----..-.- ——— 3

kugleri, Bulimina _ - ____________ - __ .-...- _ ..___-- 41, 44 (tab.), pi. 11

laeitigata, Entosolenia—..............................-........ — . 41, 44 (tab.), pi. n

Nodosaria (Glandulina)..-....... _..------.-___--_--._-----...---.---— ?°

Triloculina-.-............................................... IS, 12 (tab.), pi. 3

laevis, Bullopora—............................................. 36, 44 (tab.), pi. IT

Lagena.. ______-..-„„-- — .--.____...___-___ 28 vtab.), 31, pi. 9 

ListereUa-..——..—-—-- — - — --- — --—-——— —— .———- 10 

Vitrewebbina.- ___...... —.. —— ----------- —.-...-...- — ————... 36

Webbina.—.-.—..—...-.-—— — ———- — - — — — ———-- 36

laevis cretosa, Spiroplectammina................. ——.....--.-..— 6, 12 (tab.), pi. 1

laeois hispida, Bullopora.—.......__....—.--...——--___ — -——— 36, 44 (tab.)

Lagena acuiicosta...................:..........-.........—...— 28 (tab.), 32, pi. 9

apiculata__..._..........---.----.-..-..--_-----.------------—...... 4-2

costata——.... ___--_-._-_. — .-.———.--------- —------ 28 (tab.), 32, pi. 9

laevis......................................................— 28 (tab.), SI, pi. 9

orbignyana......................................-.--.----........---..---. 41

i primigera—....... .....................——— — ..-—... 28 (tab.), 32, pi. 9

sulcata semiinterrupta........................................ 28 (tab.), 31, pi. 9

sp. A..............................................................——. 32

Lagenidae.—--.. .....--..-..._ — ....._...--.,.---..-..___... 13, 28 (tab.)
72 INDEX

Page 

Lamarckinalimbata........-___.....____________.... 49, 60 (tab.), pi. 14

naheolensis..............-__.____________.___ 49, 60 (tab.), pi. 14

paleocenica.................................................. 49, 60 (tab.), pi. 14

ripleyensis..—............................__..___.........__.___. 49

rugulosa........................................_____._. 49, 60 (tab.), pi. 13

larva, Nodosaria................................................____...... 25

latejugata, Nodosaria.............................................. %S, 28 (tab.), pi. 7

legumen elegans, Vaginulina...................................____-_.-.. 19

Lenticulina degolyeri....-..................................................... 14

midwayana............................................. ._._.--.-__..-. 13

midwayensis......................................................._._.. 13

limbata, Alabaminawilcoxensis.................................. 67, 60 (tab.), pi. 16

Lamarckina................................................. 49, 60 (tab.), pi. 14

PuMnulinaexigua.—..................................................... 57

Pulvinulinella exigua...................................................... 57

Llngulina minuta............................................................. 26

naheolensis......................---......—...........___ 96, 28 (tab.), pi. 7

wilcoxensis................................................... 27, 28 (tab.), pi. 7

Listerella laevis...... ._______......___._______............._ 10

Lituola erecta.---............................................................ 5, pi. 1

erecta distincta........................................................... 6, pi. 1

Lituolidae—-——-—- — -— — ---——--..„____..-......—. — .- 4, 12 (tab.)

Localities, Alabama...---.---.------......._-....._...............---..._.... 1

Arkansas____________...______________.__._._.. 2-3 

Mississippi-_____.....__.....____..____..__-_------- 2

Tennessee._________-..._________________ — -•-_.... 2

Texas.—.__....__— _—— —— ——— —— .__....................... 3

longiforma, Cristellaria........................................................ 28

Hemicristellaria........____....____________.________. 28

Vaginulina................................................... 98, 28 (tab.), pi. 8

longiscata, Nodosaria.............................................. 84, 28 (tab.), pi. 7

Nodosarina................................................................ 24

lotus, Epmides.................................................. 68, 60 (tab.), pi. 14

Pulrinulina____._.._............__...-_._....--__------- 52

Loxostomum applinae........................................... 48, 44 (.tab.), pi. 12

deadericki................................................... 44, 44 (tab.), pi. 12

exilis.................................................... 44, 44 (tab.), pi. 12

plummerae.................................................. 44, 44 (tab.), pi. 12

M

macneili, Nodosaria............................................... %4, 28 (tab.)

ParreUa..................................................... £3, 60 (tab.),

madrugaensis, Anomalina....................................... 60 (tab.), 64,

Boldia....................................................... 60 (tab.), 66,

Cibicides—-.-............................................. 60 (tab.), 68,

Gyroidina.................................................... 68, 60 (tab.),

Pseudoglandulina............................................ %6, 28 (tab.),

PseudoparreUa.............................................. 68, 60 (tab.),

Rotalia...................................................... 65, 60 (tab.),

Valvulineria................................................. 51, 60 (tab.),

manifesto, Glandulina_.-....-._.-_.........___-------__-____.

Nodosaria.................................................................

Pseudoglandulina......................................... S5, 26, 28 (tab.)

Marginulina costata...........................................................

distincta..................................................... 18, 28 (tab.),

dubia-—.-................................................. 18, 28 (tab.)

earlandi...................................................... 17, 28 (tab.)

eximia—..................................................... 17, 28 (tab.)

gardnerae...........................................................

glabra........................................................ 18, 28 (tab.)

hamata....................................................... 18, 28 (tab)

havanensis.................................................... 18, 28 (tab.)

regularis..................................................................

scitula......................................................... 18, 28 (tab.)

subrecta...................................................... 18, 28 (tab.)

toulmini...................................................... 17, 28 (tab.)

tubercvlata................................................... 17, 28 (tab.)

sp. A—-.. — -.—————————————————— — ——— 19, 28 (tab.)

sp.B ......................................................... 18, 28 (tab.)

,pl.7 

pi. 16 

pi. 22 

pi. 22 

pi. 21 

pi. 24 

pi. 21 

pi. 24 

pi. 15 

pi. 23 

25 

25

,pl.7 

20

pi. 21 

, pi. 5 

, pi. 5 

, pi. 5 

19

.Pl. 5 

, pi. 5 

, pi. 5 

18

, pi. 5 

, pi. 5 

, pi. 5 

, pi. 5 

,pl. 5 

, pi. 5 

17 

Marssonella indentata.................................................... 9, 12 (tab.)

oxycona—..................................................... 9, 12 (tab.), pi. 2

martinezensis, Anomalina................................. __..---60 (tab.), 84, pi. 21

Martinottiettapaleocenica.—........................................ 9, 12 (tab.), pi. 2

Mathews Landing marl member, localities.....__.___-...--......-.-...... 1

mauryae, Cancris................................................ 56, 60 (tab.), pi. 15

membranacea, Oloborotalia....................................... 60 (tab.), 61, pi. 17

Mexia member, locality-____-._-__..._----------__...___..--- 3

mexiaensis, Spiroplectammina..................................... 6, 12 (tab.), pi. 1

mexicana, Pulvinulinella culler................................................ 53

Midway formation, localities_............__..___._........._.-.- 2-3

Midway group, localities...---..-------.-..._--_.-..-..---........._...... 3

spPage 

midwayana, Lenticulina............................................ ... —..— 13

pansa, Textularia________...____________-__ 7, 12 (tab.), pi. 2 

Pulvinulinella cutter....................................................... 53

Textularia..................................................... 7, 12 (tab.), pi. 1

Vaginulina................................................... 28, 28 (tab.), pi. 8

midwayensis, Adhaerentia......................................... IS, 12 (tab.), pi. 3

Ammobaculites.......................................—...... 4, 12 (tab.), pi. 1

Anomalina.............................................. 60 (tab.), 68, 64, pi. 17

Bolivina-.................................................... 4$, 44 (tab.), pi. 12

Bulimina arkadelphiana..................................... 40, 44 (tab.), pi. 11

davulinoides.................................................. 8, 12 (tab.), pi. 2

Cristellaria................................................................ 13

Discorbis.................................................... 48, 60 (tab.), pi. 13

Ellipsonodosaria............................................. 44 (tab.), 47, pi. 13

Epistominoides.............................................. 68, 60 (tab.), pi. 16

Frondicularia................................................. 28 (tab.), SO, pi. 9

Oumbelina.................................................. ST, 44 (tab.), pi. 11

Lenticulina...................*........................................... 13

Robulus...................................................... IS, 28 (tab.), pi.3

Saracenaria.—............................................... 87, 28 (tab.), pi. 7

Truncatulina........................————. ————— — ———— —— —— 62

midwayensis carinata, Cristellaria............. ————————————-——--—— 13

carinatus, Robulus............................................ IS, 28 (tab.), pi. 3

midwayensis soldadoensis, Discorbis.............................. 4&, 60 (tab.), pi. 13

midwayensis trinitatensis, Discorbis.............................. 4$, 60 (tab.), pi. 13

midwayensis trochoidea, Anomalina.............................. 60 (tab.), 62, pi. 18

Truncatulina................................................. ——— —.. 62

Mfliolidae———..————————————————— 10,12 (tab.)

minima, Gaudryina-.......................................................... 8

minuta, Globulina................................................ SS, 44 (tab.), pi. 9

Lingulina................................................................. 26

mirificus, Cibicides—————————..........——-------- 60 (tab.), 68, pi. 20

Mississippi, localities..----------- ——— -.... ————— —. — —— -----——— 2

morsel, Entosolenia...................—..........——.... — ... 4%, 44 (tab.), pi. 12

Oumbelina—.........—————..—.—————... — .. SS, 44 (tab.), pi. 11

mucronata, Dentalina.-..............____..______.__._ S9, 28 (tab.), pi. 6

N 

Naheola formation, localities——.--.-—----.--..-.-.--.-.-.--.-------._..._- 1

naheolensis, Dentalina............ — —........—................. SO, 28 (tab.), pi. 6

Dentalina delicatula....................................................... 20

Frondicularia...... —— ——— —— —————————————— 28 (tab.), SO, pi. 8

Lamarckina...........................—— — ——— _ ——— _ 4£, 60 (tab.), pi. 14

Lingulina.....................................——......_.. SB. 28 (tab.), pi. 7

Pseudouvigerina............................................. SS, 44 (tab.), pi. 11

Ouingueloculina.............................................. 10, 12 (tab.), pi. 2

Virgulina———. — ———.———————.————. 4$, 44 (tab.), pi. 12

nassauensis cubana, Valvulammina............................... S, 12 (tab.), pi. 21

nasuta, Dentalina.........———-—-—————————_.. $9, 28 (tab.), pi. 6

natchitochensis, Triloculina........—........—.—............... 18, 12 (tab.), pi. 3

Nautilus spinulosus........................................................... 47

neivmanae, Discorbis.......................................................... 66

Cibicides.......................—............................ 6C (tab.), 66, pi. 19

Nodosarella attenuata...................-........................ 44 (tab.), 45, pi. 12

paleocenica.................... ____..—...__.--_-_._.. 44 (tab.), 46, pi. 12

Nodosaria affinis.......... ————......................... — —— S3, 28 (tab.), pi. 7

amphioxys .................................———............ $4, 28 (tab.), pi. 7

granti.............................................. —— ...——.—........ 24

humilis.............'.—................-..-.-.....--..--......---.-----... 25

larvat..............................------------ —— — - — - — ----------- 25

latejugata..............................—.................... S3, 28 (tab.), pi. 7

longiscata........————————............................... 84, 28 (tab.), pi. 7

macneili... — . —— ... ————..............—................. $4, 28 (tab.), pi. 7

manifesto. _.........___......___-------—----------—— — ——... 25

oligotoma................................................................. 24

pauperata................................................................. 22

paupercula................................................................ 24

proxima............ — ———— ————— ———— —— — ————— ——— — — ——— — 23

pseudo-obliquestriata. 

radic ula.--..........

sagrinensis..........

spimdosa...........

vertebralis...........

zippei_............

__ ——— _- —— ——— — —— —— — - ——— ———--- 19

— — ——— —— ————— ——— ——— — —— — 25

— —————— ———— ———— ——— ————— — — — 46

————————— —————— ——————— — —— 47

———— — ——— ——— ——————t — — — ——— 23

——— ——— — — — ——— ——— — ———— — ——— 23

Nodosaria (Glandulina) comata................................................ 26

Nodosaria (Glandulina) laevigata....---.----.-----............................ 26

Nodosarina goldfussi.......................................................... 31

longiscata................ ....— — ---._ — - — — --—___ — -------- —.-. 24

Nonionella insecta........_____.__-__.._____ —._ 37, 44 (tab.), pi. 11

soldadoensis.... ...................................—-...... S7, 44 (tab.), pi. 11

welleri................................................— .....————. 63

sp—...... — __._--- —--_- — - —— —— - — ——------ 37, 44 (tab.), pi. 11
INDEX

Page

Nonionidae...-._____________________.___________ 37, 44 (tab.) 

Nonionina turgida..... .....____.._____...._____._.._...... 37

O

obtusa, PuMnulina exigua. 

Pulvinulinetta.........

exigua............

oldhami, Frondicularia.-... 

oligotoma, Nodosaria......

Ophthalmidiidae--—.— ... 

orbicularis, CristeUaria....

.............................................. 57

——— .—.- — __.____._........__...... 57

——— ————————— — ——— —— ——— ——— 57

.._———— ———— . ——— ————— — —————— 29

.__.___._____...._.-._..__.--__-.__.. 24

......—.—....„ — ......-.-.....-..-..... 12

—— — ————— ——— ——— ___ ——— .————— 16 

orbignyana, Entosolenia..........................___......_ 41, 44 (tab.), pi. 12

Lagena...........................................-..—..............._... 41

oxycona, Oaudryina.................. —— ._ __.__.—_—------_.___-----.- 9

Marssonella..-................................................ 9, 12 (tab.), pi. 2

paleocenica, Allomorphina....................................... 58, 60 (tab.), pi. 16

Ellipsonodosaria.......................... ___________ 44 (tab.), 46, pi. 13

Lamarckina......................._____.____.____ 49, 60 (tab.), pi. 14

Martinottiella.................................................. 9, 12 (tab.), pi. 2

Nodosarella.—.............................................. 44 (tab.), 46, pi. 12

Palmula primitiva............................................ 28 (tab.), 30, pi. 7

Pleurostomella............................................... 44 (tab.), 45, pi. 12

Spiroplectammina........................... .....___..._ 7, 12 (tab.), pi. 1

paleocenicus, Ammobaculites-...................................... 4, 12 (tab.), pi. 1

Palmula budensiS-............................ __.______. 28 (tab.), 29, pi. 7

delicatissima................................................. 28 (tab.), 89, pi. 7

primitiva.........__-.._.._._._.__...._....._________. 30

paleocenica..-..... _______._______________ 28 (tab.), 30, pi. 7

rugosa........................................................ 28 (tab.), 29, pi. 7

pansa, Textularia midwayana. .................................... 7, 12 (tab.), pi. 2

Parrella expansa.......,._._______._______.____..... S3, 60 (tab.), pi. 16

macneili—.................................................. 53, 60 (tab.), pi. 16

partschiana, Pulvinulina.......___.________.........._....__ 55

pasionensis, Colettes............................................. 54, 60 (tab.), pi. 24

Patellina sp..................................................... 48, 60 (tab.), pi. 13

Patellinoidessp.................................................. 48, 60 (tab.), pi. 13

pauperata, Dentalina............................................. S%, 28 (tab.), pi. 6

Nodosaria................................................................ 22

paupercula, Nodosaria...............................................___. 24

perplexa, Ceratobulimina......................________._.. 56, 60 (tab.), pi. 16

Rotalia................................................................... 56

piluUferus, Robulus............................................... 18, 28 (tab.), pi. 5

Pine Barren member, localities.__...._.______-------___..__ 2

Placopsilinidae--..________—______________.____ 12,12 (tab.) 

PlanoglobuJina acervulinoides........................................... 38, 44 (tab.)

Planularia toddae...............^................................ 16, 28 (tab.), pi. 5

sp. — ———————— ——— —— ——— ——————— ——— ——— ————— 16

Pleurostomella alternans........................................____.......... 45

brevis alternans............................................................ 45

paleocenica....... ..__...______...______....__. 44 (tab.), 45, pi. 12

plummerae, Dentalina............................................ 20, 28 (tab.), pi. 6

Ellipsonodosaria.--.......................................... 44 (tab.), 46, pi. 13

Eponides.................................................... 58, 60 (tab.), pi. 14

Loxostomum................................................ 44, 44 (tab.), pi. 12

Ouinqueloculina.............................................. 11, 12 (tab.), pi. 2

Spiroplectammina............................................. 5, 12 (tab.), pi. 1

Textularia..................................................... 7, 12 (tab.), pi. 2

plummerae arkansasana, Textularia.-..-.-..-....................... 7, 12 (tab.), pi. 2

flectata, Quinqueloculina...................................... 11, 12 (tab.), pi. 2

plumoides, Vaginulina........................................ $8, 28 (tab.), 29, pi. 8

Polymorphina acuta........................................................... 32

amygdaloides terquemiana................................................. 34

cushmani—................................................. 35, 44 (tab.), pi. 10

extensa.........................................____...._.._..___... 34

frondea...................................................... 35, 44 (tab.), pi. 10

sp______.—_——.__——. ———————————————— 35, 44 (tab.), pi. 10

Polymorphinella elongata........................................ SB, 44 (tab.), pi. 10

sp........................................................... 36, 44 (tab.), pi. 10

Polymorphinidae..—_____.__ '................................... 32, 44 (tab.)

Textularia..--—.......................................... — - 8, 12 (tab.), pi. 2

Porters Creek formation, localities___.___...________--..-.. 1, 2

praecursoria, Discorbina... _____._______._...._..___--.-... 65

praecursorius, Cibicides.......................................... 60 (tab.), 65, pi. 19

praespissiformis, Anomalina..................................... 60 (tab.), 64, pi. 22

prima, Siphonina..... ....____._._____._______._._ 56, 60 (tab.), pi. 15

primaria, Asterigerina..............................______.. 56, 60 (tab.), pi. 15

primigera, Lagena.. __________.____________.__ 28 (tab.),&?, pi. 9 

primitiva, Palmula............................................................ 30

paleocenica, Palmula.......................................... 28 (tab.), SO, pi. 7

problema, Outtulina.... __...__......__.__.__________ 38, 44 (tab.), pi. 9

Page 

projecta, Flabellina....................................................—.... 30

Frondicularia.....................................—...................... 30

proxima, Nodosaria.................................................-.......... 23

pseudo-bulloides, Olobigerina..................................... 60, 60 (tab.), pi. 17

pseudo-costata, CristeUaria..................................................... 14

pseudo-costatus, Robulus.......................................... 14, 28 (tab.), pi. 4

comis, Robulus............................................... 14, 28 (.tab.), pi. 4

inornatus, Robulus..............——.—-....——....—......——--.-—. 14

Pseudoglandulina caudigera....................................... 26, 28 (tab.), pi. 7

comata...........................................—..........——————— 26

madmgaensis................................................ 86, 28 (tab.), pi. 21

manifesta.—.............................................. SB, 26, 28 (tab.), pi. 7

pygmaea........................................—........... 26, 28 (tab.), pi. 7

pseudo-mamilligera, CristeUaria..-.---........................................ 13

pseudo-mamilligerus, Robulus..................................... 13, 28 (tab.), pi. 4

pseudo-nasuta, Dentalina......................................... SI, 28 (tab.), pi. 6

pseudo-obliquestriata, Dentalina.....—.—........................ 19, 28 (tab.), pi. 6

Nodosaria........................................................—....... 19

Pseudoparrella exigua......................_.-... ................. 67, 60 (tab.), pi. 16

madrugaensis................................................ 58, 60 (tab.), pi. 24

Pseudopolymorphina wilcoxensis................................. 54,44 (tab.), pi. 10

sp. A.........——...—.— -————.-———.———— 34, 44 (tab.)

pseudoungerianus, abicides.................................... ————————. 67

Pseudouvigerina naheolensis...................................... 39, 44 (tab.), pi. 11

wilcoxensis-.----.--.-—--..-...-----------------------------------........ 44

puleherrima, Quinqueloculina..................................... 10, 12 (tab.), pi. 2

Pl.llenia quinqueloba.......................................................... 59

quinqueloba angusta.......................................... 59, 60 (tab.), pi. 17

PuMnulina exigua limbata .................................................... 57

exigua obtusa........................................................-..... 57

lotus-.-..... — ———.....-----------------------------------—------ 52

partschiana...............

reticulosa..................

semiplecta................

Pulvinulinella culler mexicana. 

cutter midwayana..........

exigua obtusa...............

obtusa....— ——— —...

——— — — — — .———— -- — — — . 55

— —— — ——— — — — — —— — — ——— 54

. ————— ————— —— ———•—— —— —— 67

—— ————— ——— ———— ————— —— — — 53

—— ——— ——— ——— ——— ————— — ——— . 53

—— ——————————— —— ——— ————— 57

. ——————— ——— ———— ———— ——————— 57

pungens, Dentdlina........................................................... 21

pygmaea, Olandulina.......................................................... 26

Pseudoglandulina............................................. 36, 28 (tab.), pi. 7

Pyrulina cylindroides.............................—.............. 38,44 (tab.), pi. 9

extewsa —.———————— —————— ————— 84, 44 (tab.), pi. 9

Q 

quadrata, Bulimina (Desinobulimina)............. ........——... 4!, 44 (tab.), pi. 11

Bulimina (FJlipsobulimina)............................................... 41

quinqueloba, Pullenia................ ———————— — .. —. — ——————— 59

quinqueloba angusta, Pullenia.................................... b9, 60 (tab.), pi. 17

Quinqueloculina ferussadi..................................................... 11

alabamensis--.---...........—............................... 10, 12 (tab.), pi. 2

naheolensis- —-..—--——-- — — -— ———— — — ——- 10, 12 (tab.), pi. 2

plummerae——............................................... 11, 12 (tab.), pi. 2

flectata—...————....—...........................— 11, 12 (tab.), pi. 2

pulcherrima——.——————————— —— — — ———— 10, 12 (tab.), pi. 2

R 

radicula, Nodosaria............................................................ 25

Ramuhna aculeata.........................................'...... 36, 44 (tab.), pi. 10

Rectogumbelina alabamensis ..................................... 38, 44 ^tab.), pi. 11

regularis, Marginulina......................................................... 18

rugulosa, Lamarckina..-......................................... 49, 60 (tab.), pi. 13

reprimatus, Cibicides...........—............... ——............ 60 (tab.), 68, pi. 20

reticulosa, PuMnulina........................................................ 54

reticulosus, Colettes.....................——..................... 54, 60 (tab.), pi. 15

Rhizamminidae._________....___.__ —— —_.___.—— 3,12 (tab.)

ripleyensis, Lamarckina....................................................... 49

Robertina wilcoxensis...................—.————............. 40, 44 (tab.), pi. 11

Robulina rosetta.......................——..—....—.........——.——..... 16

Robulus alabamensis.............................................. IB, 28 (tab.), pi. 4

a rkansasan us................................................. Iff, 28 (tab.), pi. 4

degolyeri...—— ————————————————————... 14, 28 (tab.), pi. 3

insulsus......................——........................._ 16, 28 (tab.), pi. 5

midwayensis.................................................. IS, 28 (tab.), pi. 3

carinatus...............................................— IS, 28 (tab.), pi. 3

piluliferut... — —————.—.———.——...————— 16, 28 (tab.), pi. 5

pseudo-costatus............................................... 14, 28 (tab.), pi. 4

comis------.. — — ._—- ————-__--.--._. 14, 28 (tab.), pi. 4

inornatus............................................................. 14

pseudo-mammiligerus........................ ___-...__- IS, 28 (tab.), pi. 4

rosettus——————— ——. ————. —— ————- 16, 28 (tab.), pi. 4

turbinatus.---—-.— —————-.—.—.........———..... 14, 28 (tab.), pi. 4
74 INDEX

Page 

Robulus alabamensis—Continued

wilcoxensis.................................................... 15, 28 (tab.), pi. 4

dissentius—.............................................. IB, 28 (tab.), pi. 4

sp...----------------------------------------------------- 16, 28 (tab.), pi. 5

robusta, Vaginulina.-.---..................................................... 28

Rosalina clementiana.......................................................... 63

rosetta, Robulina........... _____——____..__--._„__..-._—___.__... 16

rosettus, Robulus...............____.._______.________ 16, 28 (tab.), pi. 4

rossae, Spiroplectammina.......................................... B, 12 (tab.), pi. 1

Rotalia aeguilateralis-......................................................... 51

havanensis................................................... 55, 60 (tab.), pi. 15

madrugaensis__-— — -—----——_-_- — — - — __........... 55, 60 (tab.), pi. 15

perplexa.................................................................. 56

soldanii subangulata.- .................................................... 51

Rotalitdae—__.—--- — .-- — ---..——-.. — ..-.. — ....„ — ...... 47, 60 (tab.)

rotundata, Globulina.............................................. SS, 44 (tab.), pi. 9

rudita, Gaudryina..— --.. — —.—-.—. ............................. 8, 12 (tab.), pi. 2

rugosa, Flabellina............................................................. 29

Frondicularia.-----------------...^................................_... 30

Gaudryina..........................................___............... 8

Palmula..................................................... 28 (tab.), %9, pi. 7

rugulosa, Lamarckina............................................ 49, 60 (tab.), pi. 13

sagrinensis, Ellipsonodosaria....— .... ...... ........ . .... ...... .........

Nodosaria ..... ---... ... .. ... ..................... ... ...... ..........

Saracenaria midwayensis ..... ............ ... ......... ... ... ....... 87, 28 (tab.)

sublatifrons ..... — __.. ___ _ . _______ . ___ ___ ._ _ $7, 28 (tab.)

trigonata ........ ... ......................... ... .. ...... ....... 27, 28 (tab.)

Schenckiella alabamensis.. ... ..... ........ ___ ...... ___ .. _ 1C, 12 (tab.)

scitula, Marginulina........................................... ... 18, 28 (tab.)

selseymsis, Spirittina..... ... ... ...... .......... .. ... ... ..... .... 47, 60 (tab.),

semiinterrupta, Lafcna sulcata. ........ .... .... ..... ... ........... 28 (tab.), SI

semilaevfs, Vaginulina ...... . ...... ... ........... .... ......... ... $8, 28 (tab.),

semiplecta, Pulvinulina. ................ .... ....... .. ... ....... ......... .... ..

semiplectus, Cibicides.. .......................................... 60 (tab.), 67,

semitecta terquemiana, Sigmomorphina .... . ..... . .... . .... ....... S4, 44 (tab.),

Sigmomorphina semitecta terquemiana .... .. ... .. ... . ... .......... S4, 44 (tab.),

soldadoensis ......... ... .. —— — _____ --------- _ ....... SB, 44 (tab.),

wilcoxensis... ...... ... ... ..... .... ... . ... ..... ....... ....... S4, 44 (tab.),

williamsoni.... .......... ... ... ... . ____ ..... _ ...... _ _. SB, 44 (tab.),

sp.

46 

46

,pl.7 

,pl.7 

,pl.7 

58

, pi. 2 

, pi. 5 

pi. 13 

, Pl-9 

pi. 21 

67

pi. 20 

pi. 10 

pi. 10 

pi. 10 

pi. 10 

pi. 10 

35 

Siphogenerina eleganta-------...-............................................. 39

Siphogenerinoides eleganta...—._.-_ —..———.......__._... S9, 44 (tab.), pi. 11

Siphonina prima................................................ 55, 60 (tab.), pi. 15

wilcoxensis.................................................. 56, 60 (tab.), pi. 15

soldadoensis, Discorbis midwayensh.............................. 48, 60 (tab.), pi. 13

Gaudryina..................................................... 8, 12 (tab.), pi. 2

NonioneOa.................................................. 37, 44 (tab.), pi. 11

Siamomorphina............................................. SB, 44 (tab.), pi. 10

soldanii subangulata, Rotalia.................................................. 51

spinulosa, Ellipsonodosaria.................................................... 47

Nodosaria..........------_-..._..____________........._... 47

spinulosus, Nautilus.......................................................... 47

Spirittina selseyensis............................................. 47, 60 (tab.), pi. 13

mvipara.....--.----..---.-................................ 48, 60 (tab.), pi. 13

sp........................................................... 48, 60 (tab.), pi. 13

Spiroloculina alabamensis.-....................................... 11, 12 (tab.), pi. 3

Spiroplectammina laevis cretosa.................................... 6, 12 (tab.), pi. 1

mexiaensis..................................................... 6, 12 (tab.), pi. 1

paleoeenica.................................................... 7, 12 (tab.), pi. 1

plummerae.............................. .___...._____ 5,12 (tab.), pi. 1

rossae...............-......................................... 6, 12 (tab.), pi. 1

wilcoxensis-................................................... 6, 12 (tab.), pi. 1

spissiformis, Anomalinaalazanensis........................................... 64

strigillata, Frondiculariaarchfaciana.......................... ....__...__.. 30

subaculeata, Cristellaria tuberculata.......................... ...___....._.. 17

Hemicristellaria........................................................... 17

subangulata, Rotalia soldanii.................................................. 51

Gyroidina.--.—............................................ 51, 60 (tab.), pi. 14

sublatifrons, CristeUaria..-.................................................... 27

Saracenaria................................................... 27, 28 (tab.), pi. 7

subrecta, Marginulina............................................. 18, 28 (tab.), pi. 5

subtriangvlaris, AllomorpMna................................... 69, 60 (tab.), pi. 17

ChilostomeUa............................................................. 59

sulcata semiinterrupta, Lagena...........................____'. 28 (tab.), SI, pi. 9

T

tenera, Eponides...-.

Truncatuttna....

Tennessee, localities.

Page 

terquemiana, Polymorphina amygdaloides...................................... 34

Sigmomorphina semitecta.-... . —._-.-—..——_„ ——.— $4, 44 (tab.), pi. 10

texana, Trochammina..... _______ __...—___----------_ —.. 1%, 12 (tab.), pi. 3

texanum, Chrysalogonium.............................. —————.... —. —... 25

Texas, localities-—____.....__.__.——.—— —.—-——_.____ 3

Textularia agglutinans.....-------------- ...— —— ——...——————————-- 8

carinata eipansa........ _______—.__._.———————— ———............. 5

eocenita.............. — — ——— ———— — —. —.... —— — — — ___-_... 7

midwayana...-_._. — -_——————. —.. —— ———— —.. 7, 12 (tab.), pi. 1

pansa.... ....._..... — ....__..._______.......___ 7, 12 (tab.), pi. 2

plummerae............. ...__________..___....— — _ 7, 12 (tab.), pi. 2

arkansasana............................................... 7, 12 (tab.), pi. 2

portenta.------........___------___..______----- 8, 12 (tab.), pi. 2

Textulariidae--——- — __-------.-———-—-—---.----—_ — .. 5, 12 (tab.)

Tippah sand lentil locality___„._ ——— ————— ——— ———....__._. 2

toddae, Planularia.,................ .._____.______.___ 16, 28 (tab.), pi. B

toulmini, Marginulina.............. .______._________ 17, 28 (tab.), pi. 5

Trifarina herberti................................................ 44 (tab.), 45, pi. 12

trigona, AllomorpMna.... ..--....—........................................... 58

trigonata, Cristellaria..--.--.--.-- —_ —_. ———— — — _. ——... —.. —— ——— 27 

Saracenaria-..-.............................................. 17/28 vtab.), pi. 7

Trilocvlina alabamensis........................................... 11, 12 (tab.), pi. 3

deca.-...-................................................... 18, 12 (tab.), pi. 3

inHata...................................--.--.--........-.......___.. 11

laevigata.. ._.._..—____________________-.__.. 1&, 12 Uab.), pi. 3

natchitochensis.................... _______________.-. n, 12 (tab.), pi. 3

triloculinoides, Globigerina....................................... 6L, 60 (tab.), pi. 17

trinitatensis, Discorbis midwayensis.............................. 4$, 60 itab.), pi. 13

Gumbelina.--............................................... SS, 44 (tab.), pi. 11

Tritaxilina cubensis........................................ ..—.. 10, 12 (tab.), pi. 2

Trochammina texana............................................. 1%, 12 itab.), pi. 3

Trochamminidae_.__________...__..—..______...__ 12,12 (tab.) 

trochoidea, Anomalina midwayensis............... ———————.. 60 (tab.), 6%, pi. 18

Truncatulina midioayensis................................................ 62

Truncatulina alleni......................................................__. 66

cuUer..................................................................... 53

elevata........

midwayensis..

trochoidea.

tenera.........

vulgaris..... ._......._._.._

welleri........ .______....

tuberculata, Cristellaria subaculeata. 

Hemicristellaria subaculeata....

..................................... 52

.. —— .. —......————— —— -.__-. 62

—.. — . —— — —— —— ——— . —— ——— 62

— — ..—————- —— — - —— —— - 52

—— .. —— . ——....._ —— .......... 66

. — ..... — ......................... 63

.— .„-.. ——— - ——— .... —...... 17

.——.. —— — —— - — —— - —— -.-_ 17 

Marginulina.................................................. 17, 28 (tab.), pi. 5

turbinata, CristeUaria................................... ———._._..— 14

turbinatus, Robulus.............................................. 14, 28 (tab.), pi. 4

turgida, Nonionina...........--..-................. ...— — — ——..————— 37

U 

umbonifera, Anomalina....---................................... 60 (tab.), 62, pi. 17

Discorbina..................-...-......................................... 62

Vaginulina gracilis........—.... ______.____...._...... 27, 28 (tab.), pi. 8

legumen elegans.. .................................———————.——.... 19

longiforma.................................................... 28, 28 (tab.), pi. 8

midwayana................................................... SS, 28 (tab.), pi. 8

plumoides.—............................................. 2«, 28 (tab.), 29, pi. 8

robusta.................................................................... 28

semilaevis................................................... ^-28 (tab.), pi. 21

sp. A.....--- —.-...—.............. ..——............ 28 (tab.), 29, pi. 8

Vaginuhnopsis brantlyi.... ____..-.-————————.—————....._..-.. 17

Valvulammlna nassauensis cubana................................ 9, 12 (tab.), pi. 21

Valvulinaallomorphinoides—.................................................. 50

Valvulineria aUomorphinoides.................................... BO, 60 (tab.), pi. 14

extema...................................................... 5t". 60 (tab.), pi. 23

herrkki................................................................... 50

insueta.. —................................................. B.\ 60 (tab.), pi. 23

madrugaensis.............................................. .. 5,', 60 (tab.), pi. 23

wilcoxensis-............ _________..—..———.——. SO, 60 (tab.), pi. 14

V&tvviMdaie............................................................ 9,12 (tab.)

vanbeUeni, Cibicides.. ......................................._____.___ 53

Eponides.—............................................ 6S, 60 (tab.), pis. 20, 23

vandersluisi, Boldia.............................................. 60 (tab.), 65, pi. 23

Verneuilinidae_..._____.____-___-_—.-———__... 8,12 (tab.) 

eertebralis, Nodosaria—........................................................ 23

virginiana, Angulogerina......................................... 44 (tab.), 45, pi. 12

Virgulina alabamensis......................................_———.———„—— 42

naheolensis. ......................_______________. 4$, 44 (tab.), pi. 12

wilcoxensis.................................................. 4."!, 44 (tab.), pi. 12
INDEX 75

Vitrewebbina chapmani. 

laevis.-----.-.......

Page 

-___.___-__-......._.-.-.--__...-....---_-.. 36

._- — --..-.._-....... —....................... 36

sivipara, Spirillina________._-_______._______.. 48, 60 (tab.), pi. 13 

vulgoris, Cibicides............................................ 60 (tab.), 64, 66, pi. 19

Truncatulina............................................................. 66

W 

Webbina laevis................................................................. 36

welleri, Anomalina.............................................. 60 (tab.), S3, pi. 18

Nc.nionella................................................................ 63

Truncatulina............................................................. 63

wilccxtnsis, Alabamina.......................................... 57, 60(tab.), pi. 16

Angulogerina............... — ........................... 44, 44 (tab.), 45, pi. 12

Cibicides..................................................... 60 (tab.), 67, pi. 20

Dentalina.................................................... HI, 28 (tab.), pi. 6

Epistominoides....... __.-___________....__.. 58, 60 (tab.), pi. 16

GuttuMna—.................................................. 38, 44 (tab.), pi. 9

Lingulina.....-............................................... $7, 28 (tab.), pi. 7

Pseudopdymorphina........................................ 34, 44 (tab.), pi. 10

- Page 

wilcoxensis, Alabamina—Continued

Pseudouvigerina—......................................-..... .__————— 44

Pobertina- ................................................. 40, 44 (tab.), pi. 11

Bobulus—.. — .—--...—— —— .—. — —..—— — — — 15, 28 (tab.), pi. 4

Sorocewaria..---------------------------- —— ............................. 58

Sigmomorphina..--........................._-.--__--.------ S4, 44 (tab.), pi. 10

Siphonina.................................................. SB, 60 (tab.), pi. 15

SpiTOplectammina................................ ............ 6, 12 (tab.), pi. 1

Valvulineria................................................. 50, 60 (tab.), pi. 14

Virgvlina—................................................. 4®, 44 (tab.), pi. 12

wilcoxensis acuta, OloboTotaUa..................------------------ 60 (tab.), 61, pi. 17

wilcoxensis dissentius, Robulus. _ ..-.-----_---__'--_----- — —.- 15, 28 (tab.), pi. 4

wilcoxensis limbata, Alabamina....... ........................... 57, 60 (tab.), pi. 16

williamsoni, Cibicides........ ......... ...........-......... — --. 60 (tab.), 67, 20

SiqmomoTphina..................- ............_______--. 35, 44 (tab.), pi. 10

Wills Point formation, localities--.-----.---------------.----------------— 3

zippei, Nodosaria-...,..........................................-------------- 23

o

PLATES 1-24
PLATE 1

FIGURES 1, 2. Bathysiphon eocenicus Cushman and G. D. Hanna (p. 3). X 27. 1. Front view. 2. Apertural view.

3. Ammodiscus cf. A. cretaceus (Reuss) Cushman (p. 4). X 54.

4. Ammomarginulina pp. (p. 5). (After Cushman and Renz.) X 54. a, Side view; b, peripheral view. 

5-7. Ammobaculites expansus Plummer (p. 4). (After Plummer.) X 50. 5. Holotype. 6. Paratype. a, side view;

b, apertural view. 7. Holotype by transmitted light. 

8-12. Ammobaculites midwayensis Plummer (p. 4). (After Plummer.) X 50. 8, 9, Holotype, 8, from exterior; 9, by

transmitted light. 10-12. Paratypes. a, Side view; b, apertural view. 

13, 14. Ammobaculites paleocenicus Cushman (p. 4). X 23. 13. Holotype. 14. Paratype. a, Side view; 6, apertural

view.

15, 16. Spiroplectammina rossae Plummer (p. 5). (After Plummer.) X 100. 15. Holotype. a, Front view; b, apertural 

view. 16. Paratype, by transmitted light.

17. Lituola erecta Mellen and Gault (p. 5). (After Mellen and Gault.) X 55. a, Longitudinal section; b, transverse 

section.

18. Lituola erecta Mellen and Gault var. distincta Mellen and Gault (p. 5). (After Mellen and Gault.) X 45. a,

Longitudinal section; b, transverse section.

19, 20. Spiroplectammina plummerae Cushman (p. 5). 19. X 54. 20. Holotype. (After Plummer.) X 45. 

21-23. Spiroplectammina wilcoxensis Cushman and Ponton (p. 6). 21. X 60. 22, 23. X 54.

24. Spiroplectammina laevis (Roemer) Cushman var. cretosa Cushman (p. 6). X 54. 

25, 26. Spiroplectammina mexiaensis Lalicker (p. 6). (After Lalicker.) X 68. 25. Holotype. a. Front vHw; b, side

view; c, apertural view. 26. Paratype.

27. Spiroplectammina paleocenica Cushman (p. 7). X 55. Holotype. a, Front view; b, apertural view. 

28-30. Textularia midwayana Lalicker (p. 7). (After Lalicker.) X 68. 28. Holotype. a. Front view; b, side view; 

c, apertural view. 29, 30. Paratypes.
GEOLOGICAL SURVEY PUOFESSIONAL PAPER 232 PLATE 1

RHIZAMMINIDAE, AMMODISCTDAE, LITUOLIDAE, TEXTULARTIDAE
PLATE 2 "

FIGURE 1. Textularia midwayana Lalicker var. pansa Cushman (p. 7). X 60. Holotype. a. Front view; b, apertural view.

2, 3. Textularia plummerae Lalicker (p. 7). 2. (After Lalicker.) X 63. Holotype. a, Front view; b, side view; c,

apertural view. 3. X 54. 

4, 5. Textularia plummerae Lalicker var. arkansasana Cushman (p. 7). X 40. 4. Holotype. 5. Paratype.

6. Textularia portenta Cushman (p. 8). X 48. Holotype. a, Front view; fe, apertural view.

7. Gaudryina rudita Sandidge (p. 8). X 27. 

8, 9. Gaudryina soldadoensis Cushman and Renz (p. 8). 8. X 38. 9. (After Cushman and Renz.) X 63. Holotype.

a, Side view; b, apertural view.

10-16. Clavulinoides midwayensis Cushman (p. 8). 11. Holotype. a, Front view; b, apertural view. 10, 16. Microspheric 

forms. 11-15. Megalospheric forms. 10, 11. X 27. 12. X 40. 13-16. X 15.

17. Dorothia alabamensis Cushman (p. 9). X 30. Holotype.

18. Schenckiella alabamensis Cushman (p. 10). X 30. Holotype.

19. Martinottiella paleocenica Cushman (p. 9). X 48. Holotype. a, Front view; b, apertural view.

20. Tritaxilina cubensis Cushman and Bermudez (p. 10). X 48.

21. Marssonella oxycona (Reuss) Cushman (p. 9). (After Kline.) X 33.

22. Quinqueloculina naheolensis Cushman (p. 10). X 60. Holotype. a, b, Opposite sides; c, apertural view.

23. Quinqueloculina pulcherrima Cushman (p. 10). X 60. Holotype. a, Front view; b, apertural view.

24. Quinqueloculina alabamensis Cushman (p. 10). X 60. Holotype. a, Front view; b, apertural view. 

25, 26. Quinqueloculina plummerae Cushman and Todd (p. 11). 25. X 110. Holotype. 26. X 60. a, a. Front views;

b, b, apertural views. 

27. Quinqueloculina plummerae Cushman and Todd var. flectata Cushman (p. 11). X 48. Holotype.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 282 PLATE 2

TEXTULARTTDAE, VERNEUTLTM IDAE, V ALVULINIDAE, MTLTOLIDAE
PLATE 3

FIGURE 1. Spiroloculina alabamensis Cushman (p. 11). X 58. Holotype. a, Front view; b, apertural view.

2. Triloculina alabamensis Cushman (p. 11). X 60. Holotype. a, Front view; 6, apertural view.

3. Triloculina natchitochensis Howe (p. 12). X 60.

4, 5. Triloculina deca Cushman (p. 12). X 60. 4. Holotype. 5. Paratype. a, a, Front views; b, b, apertural views. 

6,7. "Triloculina laevigata Bornemann" (p. 12). 6. (After Plummer.) X 45. a, Front view; 6, apertural view. 7. X 120. 

8, 9. Cornuspira cf. C. byramensis Cushman (p. 12). 8. X 54. 9. (After Plummer.) X 47.

10. Trochammina cf. T. texana Cushman and Waters (p. 12). X 54. 

11-13. Adhaerentia midwayensis Plummer (p. 12). 11. (After Plummer.) X 15. Holotvpe. a, Front view; b, apertural

view. 12, 13. X 30. 

14-17. Robulus midwayensis (Plummer) Cole and Gillespie (p. 13). 14, 15. (After Plummer.) 14. X 30. Holotype.

a, Side view; b, peripheral view. 15. X 27. 16. X 37. X 46. 

18-20. Robulus midwayensis (Plummer) Cole and Gillespie var. carinatus (Plummer) Cushman (p. 13). 18. (After Plummer.)

X 23. Holotype. a, Side view; b, peripheral view. 19, 20. X 27.

21, 22. Robulus degolyeri (Plummer) Bandy (p. 14). 21. (After Plummer.) X 40. Holotype. a, Side view; b, peripheral 

view. 22. X 27.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 232 PLATE 3

MILIOUDAE, OPHTIIALMIDIIDAE, THOCHAMMINIDAE, PLACOPSILINIDAE, LAGENIDAE
PLATE 4

FIGVRES 1-5. Robulus pseudo-mamilligerus (Phimnicr) Cushmaii (p. 13). 1, 2. (After Plummer.) X 24. 1. Holotvpe. 2.

Paratype. 3-5. X 27.

6-9. Robulus turbinatus (Plummer) Cushman (p. 14). 6. (After Plummer.) X 46. Holotype. a, Side view; b, peripheral view. 7. X 30. 8, 9. X 56. 

10-12. Robulus pseudo-costatus (Plummer) Cushman (p. 14). 10. (After Plummer.) X 46. Holotype. a, Side view;

6, peripheral view. 11. X 43. 12. (After Kline.) X 33. 

13, 14. Robulus pseudo-costatus (Plummer) Cushman var. romis Cushman, new name (p. 14). 13. X 54. 14. (After

Kline.) X 33. Holotype. 

15, 16. Robulus alabamensis Cushman (p. 15). 15. X 46. Holotype. a, Side view; b, peripheral view. 16. X 40.

17. Robulus wilcorensis Cushman and Ponton (p. 15). X 75.

18. Robulus wilcoxensis Cushman and Ponton var. dissentius Cushman and Todd (p. 15). X 54. Holotype.

19. Robulus arkansasanus Cushman and Todd (p. 15). X 27. Holotype. a, Side view; b, peripheral view. 

20, 21. Robulus cf. R. rosettus (Giimbel) Cushman (p. 16). 20. X 30. 21. (After Cushman and Renz.) X 53. o, Side 

view; b, apertural view.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 232 PLATE 4

LAGRNIDAE
PLATE 5

FIGURES 1-3. Robulus insulsus Cushman (p. 16). 1, 2. X 56. 3. X 50. 1. Paratype. 2. Holotype. 3. (After Plummer.)

a, Side view; b, peripheral view.

4. Robulus piluliferus Cushman (p. 16). X 28. Holotype.

5-8. Planularia toddae Cushman (p. 16). 5. X 60. Hototype. a, Side view; b, peripheral view. 6-8. X 54. 

9,10. Robulus sp. (p. 16). 9. X 40. 10. X 27. 

11-13. Marginuhna tuberculata (Plummer) Cushman and Bermudez (p. 17). 11,12. X 54. 13. (After Plummer.) X 27.

Holotype. 

14-16. MarginuUna earlandt (Plummer) Cushman (p. 17). 14. (After Plummer.) X 27. Holotype. 15. Microspheric

young. X 30. 16. Megalospheric adult. X 30.

17. MarginuUna tout-mini Cushman (p. 17). 17. X 50. Holotype. a, Side view; b, end view. 

18, 19. MarginuUna eximia Neugeboren (p. 17). 18. X 40. 19. X 60. 

20, 21. MarginuUna cf. M. scitula Bornemann (p. 18). X 35.

22. MarginuUna cf. M. subrecta Franke (p. 18). X 55.

23. MarginuUna cf. M. dubia Neugeboren (p. 18). X 55.

24. MarginuUna cf. M. havanensis Cushman and Bermudez (p. 18). X 27. 

25-27. MarginuUna cf. M. glabra D'Orbigny (p. 18). X 55.

28, 29. MarginuUna cf. M. hamata (Franke)" Cushman and Todd (p. 18). X 40. 

30, 31. MarginuUna sp. A (p. 18). 31. X 54. 32. (After Plummer.) X 40. 

32, 33. MarginuUna sp. B (p. 18). X 40.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 232 PLATE 5

LAGENTDAE
	PLATE 6

FIGURES 1-3. Dentalina pscudo-obliquestriata (Plummer) Cushman (p. 19). 1. X 55. 2. X 15. 3. (After Plunnner.) X 24.

	Holotype.

4-7. Dentalina gardnerae (Plummer) Cushman (p. 19). 4-6. (After Plummer.) X 46. 7. X 27.

8-10. Dentalina colei Cushman and Duscnbury (p. 19). 8,9. X 60. 10. X 27.

11-15. Dentalina plummerae Cushman (p. 20). 11-14. X 40. 15. X 27. 13. Holotype. 11,12,14. Paratypes.

16, 17. Dentalina insulsa Cushman (p. 20). 16. X 30. Holotype. 17. X 45. Paratype.

18. Dentalina naheolensis Cushman and Todd (p. 20). X 55.

19, 20. Dentalina pseudo-nasuta Cushman and Todd (p. 21). X 40. 19. Holotype. 20. Paratype.

21-24. Dentalina alabamensis Cushman (p. 21). X 60. 21. Plolotype. 22-24. Paratypes.

25. Dentalina inepta Cushman (p. 21). X 45. Holotype.

26. Dentalina aruleata D'Orbigny (p. 21). X 40.

27-29. Dentalina wilcoxensis Cushman (p. 21). X 60.

30-33. Dentalina eocenica Cushman (p. 22). X 55. 33. Holotype.

34, 35. Dentalina cf. D. mucronata Neugeboren (p. 22). 34. X 55. 35. X 40.

36. Dentalina cf. D. pauperata d'Orbigny (p. 22). X 40.

37, 38. Dentalina nasuta Cushman (p. 22). X 55.

39. Dentalina sp. A (p. 22). X 55.

40. Dentalina sp. C (p. 23). X 40.

41. Dentalina sp. B (p. 23). X 40.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 232 PLATE 6

LA(JKMI)\K
PLATE 7

6. X 30. 

a, Side view; b, apertural view.

X 55.

27. 11. Holotype. 12. Paratype. 

13. Holotype. 14, 15. Paratypes.

X 40.

FIGURES 1, 2. Nodosaria latejugata Giimbel (p. 23). X 15.

3-6. Nodosaria affinis Reuss (p. 23). 3. X 15. 4, 5. X 27.

7. Nodosaria macrieili Cushman (p. 24). X 50. Holotype.

8. Nodosaria cf. N. amphiozys Reuss (p. 24). X 40.

9, 10. Nodosaria? cf. N. longiscata D'Orbigny (p. 24). 9. X 40. 10.

11, 12. Chrysalogonium arkansasanum Cushman and Todd (p. 25). X

13-15. Chrysalogonium eocenicum Cushman and Todd (p. 25). X 40.

16, 17. Pseudoglandulina manifesto (Reuss) Cushman (p. 25). X 40.

18, 19. Pseudoglandulina cf. P. caudigera (Schwager) Cushman (p. 26).

20—22. Pseudoglandulina pygmaea (Reuss) Cushman (p. 26). X 40.

23. Lingulina naheolensis Cushman (p. 26). X 60. Holotype. a, Front view; b, apertural view.

24. Lingulina cf. L. wilcoxensis Cushman and Ponton (p. 27). X 15.

25,26. Saracenaria trigonata (Piummer) Cushman (p. 27). 25. (After Plummer.) X 50. Holotype. a, Oblique side 

	view; b, front view. 26. (After Kline.) X 33.

27. Saracenaria sublatifrons (Plummer) Kline (p. 27). (After Plummer.) X 45. Holotype. a, Side view; b, front view.

28-30. Saracenaria midwayensis Kline (p. 27). (After Kline.) X 33. 28, Front view; 29, rear view; 30, side view.

31, 32. Palmula budensis (Hantken) Cushman (p. 27). X 40.

33-35. Palmula delicatissima (Plummer) Cushman (p. 29). 33. (After Plummer.) X 50. Holotype. 34,35. X 40.

36,37. Palmula rugosa (D'Orbigny) Cushaman (p. 29). 36. (After Plummer.) X 45. 37. X 40.

38. Palmula primitiva Cushman var. paleocenica Cushman (p. 30). X 40. Holotype.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 232 PLATE 7

LAGENIDAK
PLATE 8

Fir.VREs 1-3. Vaginulina gracilis Plummer (p. 27). 1. X 30. 2, 3. (After Plummer.) X 24.

4-9. Vaginulina midwayana Fox and Ross (p. 27). 4-7. X 30. 8, 9. (After Plummer.) X 24 

10-15. Vaginulina longiforma (Plummer) Cushmau (p. 28). 10, 14, 15. X 27. 11-13. X 50. 

16-18. Vaginulina plumoides Plummer (p. 28). 16. (After Ciishman and Renz.) X 50. a. Side view; b, peripheral view.

17, 18. X 40.

19. Vaginulina sp. (p. 29). X 40.

20-23. Frondicularia cf. F. goldfussi Reuss (p. 31). 20. (After Plummer.) X 25. 21, 22. X 55. 23. X 40. 

24, 25. Frondicularia naheolensis Cushman and Todd (p. 30). 24. X 75. Holotype. a. Front view; b, side view; c, 

apertural view. 25. X 40.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 232 PLATE 8

LACENTDAK
	PLATE 9

FIGURES 1-3. Frondicularia cf. F. frankei Cushman (p. 31). 1. X 30. 2. X 40. 3. (After Kline.) X 33.

4, 5. Frondicularia midwayensis Cushman (p. 30). X 40. 1. Holotype. 5. Paratype.

6. Frondicularia sp. A (p. 31). X 40.

7. Entosolenia cf. E. apiculata (Reuss) Cushman (p. 42). X 40.

8, 9. Lagena cf. L. laevis (Montagu) Williamson (p. 31). 8. X 40. 9. X 95.

10, 11. Lagena sulcata (Walker and Jacob) Parker and Jones var. semiinterrupta W. Berry (p. 31). (After Kline.) X 60.

12. Lagena cf. L. acuticosta Reuss (p. 32). X 40.

13. Lagena cf. L. costata (Williamson) Reuss (p. 32). X 40.

14. "Lagena cf. L. primigera H. B. Brady" (p. 3 2). (After Kline.) X 60.

15-18. Guttulina problema d'Orbigny (p. 32)". 15. X 100. 16, 18. X 40. 17. X 55.

19-21. Guttulina hantkeni Cushman and Ozawa (p. 32). 19. X 95. 20. X 55. 21. X 40.

22. Guttulina wilcoxensis Cushman and Ponton (p. 32). X 50.

23, 24. Pyrulina cf. P. cylindroides (Roemer) Cushman and Ozawa (p. 33). 23. X 50. 24. X 40.

25. Pyrulina extensa (Cushman) Cushman and Ozawa (p. 34). X 40.

26-28. Globulina gibba D'Orbigny (p. 33). 26. X 95. 27. X 50. 28. X 40.

29-33. Globulina rotundata (Bornemann) Cushman and Ozawa (p. 33). 29, 31-33. X 50. Fistulose forms. 30. X 55.

34. dlobulina cf. G minuta (Roemer) Reuss (p. 33). X 40.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 232 PLATE 9

LAGENTDAE, POLYMORPHTNIDAE, BULIMTNIDAE
PLATE 10

FIGURE 1. Pseudopolymorphina wilcoxensis Cushman and Ponton (p. 34). X 50.

2, 3. Sigmomorphina semitecta (Reuss) Cushman and Ozawa var. terquemiana (Fornasini) Cushman and Ozawa (p. 34). 

2. X 110. 3. X 40.

4. Sigmomorphina wilcoxensis Cushman and Ponton (p. 34). X 40.

5. Sigmomorphina cf. S. williamsoni (Terquem) Cushman and Ozawa (p. 34). X 60.

6. Sigmomorphina soldadoensis Cushman and Renz (p. 35). (After Cushman and Renz.) X 55. Holotype. a, Front 

view; b, apertural view.

7. Polymorphina sp. (p. 35). (After Kline.) X 33. 

8-12. Polymorphina cushmani Plummer (p. 35). 8. (After Plummer.) X 24. Holotype. 9. (After Kline.) X 33.

10. X 15. 11, 12. X 55. Young stages.

13. Polymorphina frondea (Cushman) Cushman (p. 35). (After Kline.) X 33. 

14-16. Polymorphinella cf. P. elongata Toulmin (p. 36). X 55.

17. Polymorphinella sp. (p. 36). X 60.

18. Bullopora laevis (Sollas) Wickenden (p. 36). X 30.

19-22. Bullopora chapmani (Plummer) Cushman (p. 36). 19. X 40. 20, 21. (After Plummer.) X 24. 20. Holotype. 

21. Paratype, detached chamber. 22. X 22.

23. Bullopora chapmani (Plummer) Cushman var. hispida Kline (p. 36). (After Kline.) X 33. Holotvpe. 

24-26. Ramulina cf. R. aculeata (D'Orbigny) Wright (p. 36). 24, 25. X 35. 26. X 40.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 232 PLATE 10

POL YMOR PIIIM D A E
PLATE 11

FIGURE 1. Nonionella insecta (Schwager) Cushman and Ponton (p. 37). X 90.

2-4. Nonionella soldadoensis Cushman and Renz (p. 37). 2. (After Cushman and Renz.) X 55. Holotype. a, 6, Opposite

sides; c, peripheral view. 3. (After Plummer, "Nonionina turgida".) X 45. a, b, Opposite sides; r, peripheral view.

4. X 90.

5, 6. Nonionella sp. (p. 37). 5. X 55. 6. (After Plummer.) X 100. a, Side view; b, apertural view. 

7, 8. Giimbelina midwayensis Cushman (p. 37). 7. X 155. Holotype. 8. X 100.

9. Giimbelina trinitatensis Cushman and Renz (p. 38). (After Cushman and Renz.) X 50. Holotype. a, Front view; 

b, side view.

10. Giimbelina morsei Kline (p. 38). (After Kline.) X 60. Holotype.

11. Rectogumbelina alabamensis Cushman (p. 38). X 120. Holotype.

12. Eouvigerina excavata Cushman (p. 38). X 120. Holotype.

13-15. Pseudouvigerina naheolensis Cushman and Todd (p. 39). 13. X 120. Holotype. 14. X 120. Paratype. 15. X 55. 

16-19. Siphogenerinoides eleganta (Plummer) Cushman (p. 39). 16-18. (After Plummer.) X 45. 16. Holotype. 17, 18. 

Paratypes. 19. X 100.

20. Buliminella elegantissima (D'Orbigny) Cushman (p. 39). X 90.

21. Robertina wilcoxensis Cushman and Ponton (p. 40). X 60. 

22,23. Bulimina cacumenata Cushman and Parker (p. 40). 22. X 125. 23. X 100. a, Front view; b, rear view: r, apertural

view. 

24. Bulimina kugleri Cushman and Renz (p. 41). (After Cushman and Renz.) X 55. Holotype. a, Front view; b, rear

view; c, apertural view. 

25, 26. Bulimina arkadelphiana Cushman and Parker var. midwayensis Cushman and Parker (p. 40). (After Cushman and

Parker.) X 80. 25. Paratvpe. 26. Holotype. a, a, Front views; b, b, rear views; c, c, apertural views.

27-30. Bulimina (Desinobulimina) quddrata Plummer (p. 41). 27, 28. (After Plummer.) X 50. 29, 30. X 60. 27, 29. Microspheric forms. 28, 30. Megalospheric forms.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 232 PLATE 11

NONTONTDAE, HETEROHELTCTDAE, BULTMTNTDAE
PLATE 12

FIGURES 1, 2. Entosolenia crumenata Cushman (p. 41). 1. X 120. 2. X 60.

3, 4. Entosolenia cf. E. laevigata (Reuss) Cushman and McGlamery (p. 41). 3. X 115. 4. X 60.

5. Entosolenia cf. E. orbignyana (Seguenza) Cushman (p. 41). (After Kline.) X 45. a, Front view; b, apcrlural view.

6. Entosolenia morsei Kline (p. 42). (After Kline.) X 33. Holotype.

7. Entosolenia sp. (p. 42). X 90.

8, 9. Virgulina wilcoxensis Cushman and Ponton (p. 42). X 90.

10. Virgulina naheolensis Cushman (p. 42). X 60. Holotype. a, Front view; b, apertural view.

11, 12. Bolivina midwayensis Cushman (p. 43). X 85. 11. Holotype. 12. Paratype. a, a, Front views; b, b, apertural

	views. 

13, 14. Bolivina crenulata Cushman (p. 43). X 60.

15. Bolivina budensis (Hantken) Cushman (p. 43). X 60.

16. Bolivina sp. A (p. 43). X 150.

17. Bolivina sp. B (p. 43). X 150.

18. Loxostomum applinae (Plummer) Nuttall (p. 43). X 85. a, Front view; b, side view; c, apertural view.

19, 20. Loxostomum plummerae Cushman (p. 44). 19. X 115. Holotype. a, Front view; b, apertural view. 20. X 40.

21-23. Loxostomum deadericki Cushman (p. 44). X 60. 21. Holotype. 22, 23. Paratypes. a, Front view; b, apertural

	view. 

24-26. Loxostomum deadericki Cushman var. exilis Cushman (p. 44). X 60. 24. Holotype. 25, 26. Paratypes.

27. Angulogerina wilcoxensis (Cushman and Ponton) Cushman and Garrett (p. 44). X 60.

28. Angulogerina virginiana Cushman (p. 45). X 60.

29, 30. Trifarina herberti Cushman and Renz (p. 45). (After Cushman and Ilenz.) X 53. 29. Holotype. a, Side view;

b, apertural view. 30. Paratype.

31-33. Pleurostomella paleocenica Cushman (p. 45). 31, 32. X 55. 31. Holotype. 32. Paratype. 33. X 90.

34-37. Nodosarella attenuata (Plummer) Cushman (p. 45). 34-36. (After Plummer.) X 50. 34. Holotype. 35, 36.

	Paratypes. a, Side view; b, apertural view. 37. X 95. 

38. Nodosarella paleocenica Cushman and Todd (p. 46). X 30. Holotype
GEOLOGICAL SURVEY PROFESSIONAL PAPER 232 PLATE 12

HUUMIIMIDAK, KLUPSOiniMIMK
PLATE 13

X 54. 

.5. X 40. 

Holotype. 7-9. (After

Holotype.

Plurnmer.) X 24.

X 60. 

X 60. a, b, Opposite sides. 14. X 90.

16a, b, Opposite sides; c, peripheral view. 17a. Side

FIGURES 1, 2. Ellipsonodosaria plummerae Cushman (p. 46). 1. (After Plummer.) X 50. 2. 

3-5. Ellipsonodosaria paleocenica Cushman and Todd (p. 46). 3. X 40. 4. X 55. 

6-10. Ellipsonodosaria midwayensis Cushman and Todd (p. 47). 6. X 24. 

10. (After Kline.) X 33.

11. Chrysalogoniurn granti (Plumrner) Toulmin (p. 24). X 40.

12. Ellipsonodosaria alexanderi Cushman (p. 47). (After Kline.) 

13, 14. Spirillina selseyensis Heron-Alien and Earland (p. 47). 13.

15. Spinllina cf. S. vivipara Ehrenberg (p. 48). X 60. 

16, 17. Spirillina sp. (p. 48). (After Cushman and Renz.) X 55. 

view; b, peripheral view.

18. Patellina sp. (p. 48). X 60.

19. Patellinoidesf sp. (p. 48). X 150. a, Dorsal view; b, ventral view.

20. Discorbis midwayensis Cushman (p. 48). X 40. Holotype. a, Dot sal view;

21. Discorbis midwayensis Cushman var. soldadoensis Cushman and Renz (p. 48). 

Holotype. a, Dorsal view; b, ventral view; r, peripheral view.

22. Discorbis midwayensis Cushman var. trinitatensis Cushman and Renz (p. 48). 

Holotype. a, Dorsal view; b, ventral view; c, peripheral view.

23. Discorbis infrequens Plummer (p. 48). (After Plummer.) X 50. Holotype. 

peripheral view.

24. Lamarckina rugulosa Plummer (p. 49). (After Plummer.) X 50. Holotype. a, Dorsal view; b, ventral view; 

c, peripheral view.

b, ventral view. 

(After Cushman and Renz.) X 55.

(After Cushman and Renz.) X 55. 

a, Dorsal view; b, ventral view; c,
GEOLOGICAL SURVEY PROFESSIONAL PAPER 232 PLATE 13

LAGENTDAE, ELLIPSOTDTNTDAE, ROTALTTDAE
PLATE 14

FIGURES 1-3. Lamarckina iimbata Cushman and Todd (p. 49). X 115. 1. Holotype. Dorsal view. 2. Paratypc. Periphera

view. 3. Paratype. Ventral view.

4-6. Lamarckina naheofensis Cushman and Todd (p. 49). X 150. 4. Holotype. Dorsal view. 5. Paratype. Peripheral view. 6. Paratype. Ventral view.

7. Lamarckina paleocenica Cushinan (p. 49). X 90. Holotype. a, Dorsal view: b, ventral view; r, peripheral view. 

8, 9. Valvulineria allomorphinoides (Reuss) Cushman (p. 50). 8. X 40. a. Dorsal view; b, ventral view. 9. X 55. 

10-13. Valvulineria wilcoxensis Cushman and Ponton (p. 50). X 55. 

14, 15. Gyroidina subangulata (Plummer) Cushman (p. 51). 14. (After Plummer.) X 50. Holotype. a. Dorsal view; b,

ventral view; c, peripheral view. 15. X 40. a. Dorsal view; b, ventral view. 

16, 17. Gyroidina aequilateraiis (Plummer) Cushman (p. 51). 16. (After Plummer.) X 50. Holotype. a, Dorsal view;

b, ventral view; c, peripheral view. 17. X 60. a, Dorsal view; b, ventral view. 

18, 19. Eponides elevatus (Plummer) Cushinan and Renz (p. 52). 18. (After Cushman and Renz.) X 55. 19. (After

Plummer.) X 50. o, a. Dorsal views; b, b, ventral views; c, c, peripheral views. 

20, 22. Eponides plummerae Cushman (p. 52). 20. (After Plummer.) X 50. a. Dorsal view; b, ventral view; c, peripheral

view. 22. Holotype. a, Dorsal view, X 45; b, ventral view, X 55. 

21. Eponides lotus (Schwager) Cushman and Ponton (p. 52). X 60. a. Dorsal view; b, ventral view.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 232 PLATK 14

ROTALTTDAE
PLATE 15

1. X 50. Holotype. 

4. Early coiled stage.

a, Dorsal view; b, 

5. Young specimen

FIGURES 1-5. Coleites reticulosus (Plummer) Plummer (p. 54). (After Plummer.) 

ventral view. 2-5. X 54. 2. Dorsal view. 3. Peripheral view, 

with final chamber broken showing internal tube. 

6. Cancris mauryae Cushman and Renz (p. 56). (After Cushman and Kenz.) X 54.

b, ventral view; c, peripheral view.

7-9. Siphonina prima Plummer (p. 55). 7. (After Plummer.) X 50. Holotype. a, Dorsal view; b, ventral view; c, peripheral view. 8, 9. X 55. 8. Dorsal view. 9. Ventral view.

10. Siphonina wilcoxensis Cushman (p. 56). X 60. Dorsal view.

11. Rotalia havnnensis Cushman and Bermudez (p. 55). (After Cushman and Bermudez.) X 100. Holotype. 

view; b, ventral view: c, peripheral view.

12. Rotalia madntgaensis Cushman and Bermudez (p. 55). (After Cushman and Bermudez.) X 70.

a, Dorsal view; b, ventral view; c, peripheral view. 

13, 14. Epistomina eocenica Cushman and M. A. Hanna (p. 55). 13. (After Plummer.) X 50. a. Dorsal view; b, ventral

view; c, peripheral view. 14. X 27. Dorsal view. 

15. Asterigerina primaria Plummer (p. 56). (After Plummer.) X 50. a, Dorsal view; b, ventral view; c, peripheral view.

Holotype. a, Dorsal view;

a, Dorsal 

Holotype.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 232 PLATE 15

ROTALTTDAE
PLATE 16

FIGURES 1-5. Ceratobidimina perplexa (Plummer) Cushman and Harris (p. 56). 1-3. (After Plummer.) X 50. 1. Dorsal view.

2. Peripheral view. 3. Ventral view. 4, 5. X 110. 4. Dorsal view. 5. Ventral view. 

6,7. Alabamina vnlcoxensis Toulmin (p. 57). 6. (After Plummer.) X 50. 7. (After Cushman and Renz.) X 54. a, a,

Dorsal views; 6, 6, ventral views; c, c, peripheral views. 

8. Alabamina wilcoxensis Toulmin var. limbata (Plummer) Cushman (p. 57). (After Plummer.) X 100. Holotype

a. Dorsal view; b, ventral view; c, peripheral view. 

9, 10. Pseudoparrella cf. P. exigua (H. B. Brady) Cushman (p. 57). 9. (After Kline.) X 33. a, Dorsal view; 6, ventral

view. 10o, Dorsal view, X 54; 106, ventral view, X 45. 

11, 12. Parrella expansa Toulmin (p. 53). 11. (After Plummer.) X 50. a, Dorsal view; b, ventral view; c, peripheral

view. 12. X 52. a, Dorsal view; b, ventral view. 

13, 14. Parrella macneili Cushman (p. 53). X 60. 13. Holotype. 14. Paratype. a, a, Dorsal views; 6, 6, ventral views;

c, peripheral view. 

15, 16. Epistominoides midway ensis Plummer (p. 58). 15. (After Plummer.) X 73. Holotype. 16. X 60. a, a, Dorsal

views; b, b, ventral views; c, peripheral view. 

17, 18. Epistominoides wilcoxensis (Cushman and Ponton) Plummer (p. 58). (After Plummer.) X 73. 17. Specimen

showing interior aperture. 18o, Dorsal view; b, ventral view; c, peripheral view.

19-22. Allomorphina paleocenica Cushman (p. 58). 19. (After Kline.) X 33. a, Dorsal view; b, ventral view. 20. (After 

Plummer.) X 50. a, Dorsal view; 6, ventral view. 21. X 60. Holotype. a, Dorsal view; 6, ventral view. 

22. X 40.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 232 PLATE 16

ROTALTTDAE, CASSIDULTNIDAE, CHFLOSTOM ELLID AE
PLATE 17

FIGURES 1, 2. Allomorphina subtriangularis (Kline) Cushman (p. 59). 1. (After Kline.) X 33. Holotype. 2. X 54.

3. Allomorphina globulosa Plummer (p. 59). (After Plummer.) X 50. Holotype. a, Dorsal view; 6, ventral view. 

4,5. Chilostomelloides eocenica Cushman (p. 59). 4. (After Plummer.) X 50. a, Side view; b, ventral view. 5. (After

Kline.) X 30. Ventral view. 

6. Pullenia quinqueloba (Reuss) Reuss var. angusta Cushman and Todd (p. 59). X 75. Holotype. a, Side view;

b, apertmal view. 

7, 8. Globigerina pseudo-bulloides Plummer (p. 60). 7. (After Plummer.) X 50. Holotype. 8. X 40. a, a, Dorsal

views; b, b, ventral views; c, peripheral view. 

9. Globigerina compressa Plummer (p. 60) (After Plummer.) X 50. Holotype. a, Dorsal view; b, ventral view; c,

peripheral view. 

10, 11. Globigerina triloculinoides Plummer (p. 61). 10. (After Plummer.) X 50. Holotype. 11. X 40. a, a, Dorsal

views; b, b, ventral views.

12,13. Globorotalia wilcoxensis Cushman and Ponton var. acuta Toulmin (p. 61). 12. X 60. a, Dorsal view; b, peripheral 

view. 13. (After Cushman and Renz.) X 54 a, Dorsal view; b, ventral view; c, peripheral view.

14. Globorotalia crassata (Cushman) Cole var. aequa Cushman and Renz (p. 61). (After Cushman and Renz.) X 54. 

Holotype. a, Dorsal view; b, ventral view; c. peripheral view.

15. Globorotalia cf. C. membranacea (Ehrenberg) White (p. 61). X 160. a, Dorsal view; b, ventral view.

16. Anomalina umbonifera (Schwager) Cushman and Ponton (p. 62). X 60. a, Dorsal view; b, ventral view. 

17-19. Anomalina midwayensis (Plummer) Cushman (p. 62). 17, 18. X 54. 17. Dorsal view. 18. Ventral view. 

19. (After Plummer.) X 50. Holotype. a, Dorsal view; b, ventral view; c, peripheral view.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 232 PLATE 17

CHILOSTOMELLIDAE, GLOBIGERINTDAE, ,GLOBOROTALIIDAE, ANOMALINIDAE
PLATE 18

FIGURES 1, 2. Anomalina midwayensis (Plummer) Cushman var. trochoidea (Plummer) Kline (p. 62). 1. (After Plummer.)

X 50. Holotype. a, Dorsal view; 6, ventral view; c, peripheral view. 2. X 54. Dorsal view:

3-6. Anomalina acuta Plummer (p. 62). 3. (After Plummer.) X 50. Holotype. a, Dorsal view; 6, ventral view; 

c, peripheral view. 4. (After Cushman and Renz.) X 54. a, Dorsal view; 6, ventral view; c, peripheral view. 

5, 6. X 54. 5. Ventral view. 6. Dorsal view. 

7. Anomalina basilobata Cushman and Renz (p. 63). (After Cushman and Renz.) X 54. Holotype. a, Dorsal

view; b, ventral view; c, peripheral view.

8-11. Anomalina dementiana (D'Orbigny) Franke (p. 63). 8, 9. X 40. 10, 11. X 54. 8, 11. Ventral views. 9, 10, 

Dorsal views.

12. Anomalina welleri (Plummer) Plummei (p. 63). (After Plummer.) X 50. Holotype. a, Dorsal view; b, ventral 

view; c, peripheral view.

13. Anomalina sp. A (p. 64). (After Cushman and Renz.) X54. a, Dorsal view; 6, ventral view; c, peripheral view. 

14,15. Anomalina sp. B (p. 64). (After Cushman and Todd.) X 112. 15. Dorsal view. 14, Ventral view. 

16, 17. Cibicides alleni (Plummer) Plummer (p. 66). 16. (After Plummer.) X 50. Holotype. a, Dorsal view; 6, ventral 

view; c, peripheral view. 17. X 40. a, Dorsal view; 6, ventral view.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 232 PLATE 18

ANOMALIMDAE
PLATE 19

FICTRES 1-6. Cibicides praecursorius (Schwager) Cushrnan and Ponton (p. 65). 1, 6. X 60. a, a, Dorsal views; b, b, ventral views.

2-5. X 112. 2, 5. Dorsal views. 3, 4. Ventral views. 

7-11. Cibicides vulgaris (Plurrmer) Cushrnan (p. 66). 7. (After Plummer.) X 50. Holotype. a, Dorsal view; b, ventral

view; c, peripheral view. 8-10. X 40. 11. X 54. 8, 9, 11. Dorsal views. 10. Ventral view. 

12-14. Cibicides newmanae (Plumrrer) Cushinan and Todd (p. 66). 12. (After Plummer.) X 50. Holotype. o, Dorsal

view; 6, ventral view; c, peripheral view. 13, 14. X 90.

15-17. Cibicides howelli Toulmin (p. 67). X 60. 15. (After Toulmin.) Holotype. a, Dorsal view; b, ventral view; c, 

peripheral view. 16. Dorsal view. 17. Ventral view.
PROFESSIONAL PAPER 232 PLATE 16

ANOMALINTDAE
PLATE 20

FIGURES 1-5. Cibicides blanpiedi Toulmin (p. 67). 1. (After Toulmin.) X 75. Holotype. 2. X 60. o, o, Dorsal views; b, b,

ventral views; e, peripheral view. 3-5. X 120. 3. Dorsal view. 4. Peripheral view. 5. Ventral view. 

6-8. Cibicides browni Kline (p. 67). (After Kline.) X 33. 6. Dorsal view. 7. Ventral view. 8. Peripheral view.

9. Cibicides cf. C. williamsoni Garrett (p. 67). (After Cushman and Renz.) X 54. o, Dorsal view; 6, ventral view; c, 

peripheral view.

10. Cibicides cf. C. semiplectus (Schwager) Cushman and Ponton (p. 67). (After Cushman and Renz.) X 54. o,

Dorsal view; b, ventral view; c, peripheral view. 

11-13. Eponides vanbelleni (Van den Bold) Cushman and Bermudez (p. 53). 11. (After Van den Bold.) X 25. Holotype.

o, Dorsal view; b, ventral view; c, peripheral view. 12, 13. X 45. 12. Ventral view. 13. Dorsal view. 

14-16. Cibicides mirificus Cushman and Bermudez (p. 68). (After Cushman and Bermudez.) X 80. 14. Holotype,

dorsal view. 15, 16. Paratypes. 15. Ventral view. 16. Peripheral view. 

17, 18. Cibicides reprimatus Cushman and Bermudez (p. 68). (After Cushman and Bermudez.) X 50. 17. Holotype,

dorsal view. 18. Paratype. a, Ventral view; b, peripheral view.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 232 PLATE 20

ROTALTTDAE, ANOMALT MIME
PLATE 21

FicrREs 1-3. Cibicides madrugaensis Cushman and Bermudez (p. 68). (After Cushman and Bermudez.) X 100. 1. Holotype,

dorsal view. 2, 3. Paratypes. 2. Ventral view. 3. Peripheral view.

4. Pseudoglandulina madrugaensis Cushman and Bermudez (p. 26). (After Cushman and Bermudez.) 

type.

5. Vaginulina semilaevis Cushman and Bermudez (p. 28). (After Cushman and Bermudez.) X 22. Holotype.

6. Marginulina distincta Cushman and Bermudez (p. 18). (After Cushman and Bermudez.) X 50. Holotype. 

7-9. Anomalina martinezensis Cushman and Bermudez (p. 64). (After Cushman and Bermudez.)

dorsal view. 8, 9. Paratypes. 8. Ventral view. 9. Peripheral view.

10-12. Anomalina clementiana (D'Orbigny) Franke var. assimilis Cushman and Bermudez (p. 63). 

Bermudez.) X 70. 10. Holotype, dorsal view. 11, 12. Paratypes. 11. Ventral view 

13-15. Anomalina cubana Cushman and Bermudez (p. 64). (After Cushman and Bermudez.)

dorsal view. 14, 15. Paratypes. 14. Ventral view. 15. Peripheral view.

16. Valvulammina nassauensis Applin and Jordan var. cubana Cushman and Bermudez (p. 9). 

Bermudez.) X 30. Holotype. a, Dorsal view; 6, ventral view.

X 37. HoloX 50. 7. Holotype,

(After Cushman and 

12. Peripheral view. 

X 55. 13. Holotype,

(After Cushman and
GEOLOGICAL SURVEY PROFESSIONAL PAPER 233 PLATE 21

VALVULTNTDAE, LAGENTDAE, ANOMALTNTDAE
PLATE 22

FIGURES 1-3. Anomalina madrugaensis Cushiwan and Bermudez (p. 64). (After Cushman and Bermtidez.) X 65. 1. Holotype,

ventral view. 2, 3, Paratypes. 2. Dorsal view. 3. Peripheral view. 

4-6. Boldia carinata Cushman and' Bermtidez (p. 65). (After Cushman and Bermudez.) X 70. 4. Holotype, ventral

view. 5, 6. Paratypes. 5. Dorsal view. 6. Peripheral view.

7-9. Anomalina praespissiformis Cushman and Bermudez (p. 64). (After Cushman and Bermudez.) X 95. 7. Holotype, dorsal view. 8, 9. Paratypes. 8. Ventral view. 9. Peripheral view. 

10-12. Boldia madrugaensis Cushman and Bermtidez (p. 65). (After Cushman and Bermtidez.) X 45. 10. Holotype,

dorsal view. 11, 12. Paratypes. 11. Ventral view. 12. Peripheral view.

13, 14. Boldia cubensis Cushman and Bermtidez (p. 65). (After Cushman and Bermtidez.) X 50. 13. Holotype. 14. 

Paratype. a, a, Dorsal views; b, b, ventral views; c, c, peripheral views.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 232 PLATE 22

ANOMALTNIDAE
PLATE 23

FIGURE 1. Boldia vandersluisi Van den Bold (p. 65). (After Van den Bold.) X 27. a, Dorsal view; b, ventral view; c, peripheral

view. 

2-4. Valvulineria insueta Cushman and Bermudez (p. 51). (After Cushman and Bermudez.) X 75. 2. Holotype, dorsal

view. 3, 4. Paratypes. 3. Ventral view. 4. Peripheral view. 

5. Valvulineria madrugaensis Cushman and Bermudez (p. 51). (After Cushman and Bermudez.) X 50. Holotype.

a. Dorsal view; 6, ventral view; c, peripheral view.

6-8. Valvulineria extensa Cushman and Bermudez (p. 50). (After Cushman and Bermudez.) X 70. 6. Holotype, ventral 

view. 7, 8. Paratypes. 7. Peripheral view. 8. Dorsal view.

9. Eponides gratiosus Cushman and Bermudez (p. 53). (After Cushman and Bermudez.) X 50. Holotype. a. Dorsal 

view; 6, ventral view; c, peripheral view.

10. Eponides vanbelleni (Van den Bold) Cushman and Bermudez (p. 53). (After Cushman and Bernnidez.) X 50. a, 

Dorsal view; 6, ventral view; c, peripheral view.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 232 PLATE 23

ROTALITDAE, ANOMALTNIDAE
PLATE 24

FIGURES 1-3. Pseudoparretta madrugaensis Cushman and Bermudez (p. 58). (After Cushman and Bermudez.) X 115. 1, 3.

Paratypes. 1. Dorsal view. 3. Ventral view. 2. Holotype, peripheral view. 

4-6. Gyroidina madrugaensis Cushman and Bermudez (p. 52). (After Cushman and Bermudez.) X 130. 4. Holotype,

dorsal view. 5, 6, Paratypes. 5. Peripheral view. 6. Ventral view. 

7. Coleites pasionensis Cushman and Bermudez (p. 54). (After Cushman and Bermudez.) X 50. Holotype. a,

Dorsal view; b, ventral view; c, end view, showing aperture. 

8-10. Coleites guatemalensis Cushman and Bermudez (p. 54). (After Cushman and Bermuez.) X 50. 8. Holotype,

dorsal view. 9, 10. Paratypes. 9. Ventral view. 10. Apertural view. 

11, 12. Coleites abuillotensis Cushman and Bermudez (p. 54). (After Cushman and Bermudez.) X 56. 11. Holotype.

a, Dorsal view; b, ventral view. 12. Paratype, apertural view.

13-15. Globorotalia albeari Cushman and Bermudez'(p. 61). (After Cushman and Bermudez.) .X 110. 13. Holotype, 

dorsal view. 14, 15. Paratypes. 14. Ventral view. 15. Peripheral view.
GEOLOGICAL, SURVEY PROFESSIONAL PAPER 232 PLATE 24

ROTALITDAE, CASSIDULINIDAE, GLOBOROTALTIDAE