U. S. Department of the Interior BLM-Alaska Open File Report 90 

Bureau of Land Management BLM/AK/ST-03/010+6700+025 

July 2003 

Alaska State Office 

222 West 7th Avenue 

Anchorage, Alaska 99513 

Toolik Lake Research Natural Area / ACEC 

Rare Plant Inventory, 2002 

Amy Breen Carroll, Carolyn Parker, Tim Craig 
Cover Photo 

Claytoniella bostockii (A. E. Pors.) Jurtz (photo by M. Tachibana). 

Authors 

Amy Breen Carroll works with the Institute of Arctic Biology as well as the Biology and Wildlife Departments at 

the University of Alaska, Fairbanks. Carolyn Parker is a research associate at the University of Alaska Museum 

Herbarium in Fairbanks. Tim Craig is a Wildlife Biologist in the BLM-Alaska Northern Field Office. 

The BLM Mission 

The Bureau of Land Management sustains the health, diversity and productivity of the public lands for the use 

and enjoyment of present and future generations. 

Disclaimer 

The mention of trade names or commercial products in this report does not constitute endorsement or recommendation for use by the federal government. 

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www.ak.blm.gov/affairs/sci_rpts.html. Related publications are also listed at http://juneau.ak.blm.gov. 
Toolik Lake Research Natural Area / ACEC
 

Rare Plant Inventory, 2002
 

Amy Breen Carroll, Carolyn Parker, Tim Craig 

BLM-Alaska Open File Report 90 

July 2003 

U. S. Department of the Interior
 

Bureau of Land Management
 

Abstract 

The Bureau of Land Management-Alaska Northern Field Office and the University 

of Alaska Museum Herbarium entered into an agreement to search for rare plants in 

the Toolik and Galbraith Lake Areas of Environmental Concern (ACEC) in the 

northern foothills of Alaska’s Brooks Range. The information gathered in this project 

will enable the Bureau of Land Management to address management issues 

concerning the conservation of rare plant species that occur on these lands. 

Claytoniella bostockii (A. E. Pors.) Jurtz (Portulaceae) [ = Montia bostockii (A. E. 

Pors. ) Welsh] was found during the inventory in the Toolik Lake ACEC. In addition, 

the inventory revealed that this plant is more common in the area than previously 

thought. C. bostockii was found in several vegetation complexes, but most frequently in the Moist and Dry Nonacidic Tundra complexes. 

Acknowledgements 

This work is the result of an agreement between the U.S. Department of the 

Interior, Bureau of Land Management, Northern Field Office and the University of 

Alaska Museum Herbarium. Tim Craig of the Bureau of Land Management’s 

Northern Field Office initiated and secured the funding for this project. 

We are grateful to Dave Murray, Curator Emeritus of the University of Alaska 

Museum Herbarium, for his assistance with the nomenclature of Claytoniella 

bostockii. We also wish to thank Marilyn and Skip Walker for sharing their Toolik 

Lake vegetation data and Anne-Lillian Schell and the Alaska Natural Heritage 

Program for sharing their line drawing of Claytoniella bostockii. Lastly, we thank 

Andrew Balser, Toolik Field Station GIS and Remote Sensing Manager, for providing 

maps of the Toolik Lake region and for assisting with the construction of inventory 

maps. 

i 

 

 

Table of Contents 

Abstract ............................................................................................................................................... i
 

Acknowledgments ............................................................................................................................... i
 

Introduction ........................................................................................................................................ 1
 

Methods ............................................................................................................................................. 1
 

Results and Conclusion ..................................................................................................................... 2
 

Literature Cited................................................................................................................................... 3
 

Figures 

Figure 1. Map of Alaska showing the location of the Toolik Lake Field Station ............................. 10
 

Figure 2. Toolik Lake RNA and General Land Status Map ............................................................ 11
 

Figure 3. Map of the 2002 Rare Plant Survey Area around Toolik Lake ....................................... 12
 

Figure 4. Line drawing of Claytoniella bostockii ............................................................................ 13
 

Figure 5. Aerial photograph map of Claytoniella bostockii occurences
 

within the 2002 Toolik Lake ACEC Survey Area ........................................................ 14
 

Figure 6. Vegetation map showing Claytoniella bostockii occurences
 

within the 2002 Toolik Lake ACEC Survey Area ........................................................ 15
 

Tables 

Table 1. List of possible rare plants within the Toolik Lake ACEC ................................................... 4
 

Table 2. Definitions of rare species ranks used by the Alaska Natural Heritage Program............... 5
 

Table 3. One mile2 sections surveyed for rare plants in the Toolik Lake ACEC in 2002 .................. 6
 

Table 4. Claytoniella bostockii occurences within the 2002 Toolik Lake ACEC Survey Area .......... 7
 

ii 

 

 

 

Introduction 

Nearly 140,000 acres managed by the Bureau of 

Land Management in the vicinity of Toolik and 

Galbraith Lakes, located in the northern foothills of 

Alaska’s Brooks Range, were identified as Areas of 

Critical Environmental Concern in 1991. These 

lands include the Toolik Lake Research Natural Area 

(RNA) and the Galbraith Lake Outstanding Natural 

Area (Figs. 1 and 2). The Toolik Field Station, 

which is administered by the University of Alaska 

Fairbanks, is located in the Toolik Lake RNA. 

Currently, 88 different Long-term Ecological 

Research (LTER) sites are permitted within the 

Toolik RNA. These projects and other future 

development may affect the native vegetation within 

these Areas of Critical Environmental Concern 

(ACEC). 

The Dalton Highway Utility Corridor 

Management Plan (USDI 1989) and a subsequent 

report (Lipkin and Parker 1995) indicate two rare 

plants, Claytoniella bostockii (A. E. Pors.) Jurtz. 

(Portulacaceae) [= Montia bostockii (A. E. Pors.) 

Welsh] (Fig. 4) and Erigeron muirii Gray 

(Asteraceae), occur in the Toolik Lake ACEC. In 

addition, the management plan indicates a high 

probability of the occurrence of rare plants in the 

Galbraith Lake ACEC. This document directs the 

Bureau of Land Management to “inventory ACECs 

to delineate crucial habitat for Montia bostockii..” 

and to monitor crucial plant habitats and populations 

in the Galbraith Lake ACEC. 

To this end, the U. S. Department of the Interior, 

Bureau of Land Management, Northern Field Office 

and the University of Alaska Museum Herbarium 

entered into an agreement to search at least 3,000 

acres/year for rare plants; in the vicinity of Toolik 

and Galbraith Lakes. In addition to those plants 

previously encountered in the vicinity of Toolik 

Lake (Claytoniella bostockii and Erigeron muirii), 

we searched for other rare plants that we suspected 

to occur as well, within the survey area (Lipkin 

2000, Carolyn Parker, personal communication; 

Tables 1 and 2). The inventory began in 2002 and 

will occur over the next 5 years, as funding is 

available. This project will enable the Bureau of 

Land Management to address management issues 

concerning the conservation of rare plant species that 

occur on these lands. This report outlines the work 

accomplished during the 2002 field season. 

Methods 

In July and August 2002, we inventoried lands in 

the vicinity of the Toolik Field Station, including all 

high use research areas and the lake perimeter. To 

accomplish this, we surveyed a 3 x 3 mile area 

(5,760 acres or 23.31 km2) with Toolik Lake at its 

center. These dimensions were selected to coincide 

with section lines that provided a clear boundary for 

the survey area (Fig. 3). We further divided the 

study area into square mile sections and then 

searched each of these sections for rare plants. We 

walked line transects that were no more than 400 m 

apart within each of these sections and oriented 

transects either north to south or east to west. Search 

areas were recorded on 1 mile2

 aerial photographs of 

the area and for each rare population encountered we 

recorded: 

1. the approximate location on the map using a 

GeoExplorer 3 GPS unit (Trimble Navigation Limited, 

Sunnyvale, CA) for populations covering more than 

5000 m2

 and 

2. the exact location via a GPS point for populations 

covering less than or equal to 5000 m2

. 

In addition, a rough estimate of the population 

size and associated vegetation complex was 

recorded, following the classification established by 

Walker (2000) and Walker et al (In prep). The 

inventory began July 11 and concluded July 21, 

2002. We also conducted a brief search on August 

29 for Erigeron muirii on the rocky southeast-facing 

slope of “Jade Mountain,” a 3,057 ft peak southwest 

of the Toolik Field Station. After the inventory was 

complete, we mapped the occurrences of all rare 

Toolik Lake Rare Plant Inventory, 2002 1 
 

 

 

 

 

 

 

 

plants encountered with ArcView GIS 3.2 

(Environmental Sciences Research Institute, Inc., 

Redlands, CA). Field maps were deposited at the 

University of Alaska Museum Herbarium. 

Results and Conclusion 

We found one rare plant species during the 2002 

inventory, Claytoniella bostockii (Table 4, Fig. 5). 

This species was first documented by T. Jorgenson at 

Toolik Lake in 1978 and four more specimens have 

been collected over the years at Toolik Lake, the last 

by D. A. Walker in July 1988, all were deposited at 

the University of Alaska Museum Herbarium 

(Lipkin and Parker 1995). These previous 

specimens of Claytoniella bostockii were collected 

from the north shore of Toolik Lake “on the 

peninsula that juts toward the center of the lake” 

(Lipkin and Parker 1995). 

Claytoniella bostockii is endemic to eastern 

Beringia including several disjunct populations at 

Toolik Lake and in the Wrangell-St. Elias 

Mountains, Tetlin Hills, Nutzotin Mountains and 

eastern Yukon-Tanana Uplands. The state rank for 

this species is S3 (Lipkin 2000). However, Lipkin 

and Parker (1995) indicate that “although not yet 

documented from more than 20 locations, it has been 

found at an increasing number of sites and almost 

certainly will be found at additional sites.” 

Our inventory revealed Claytoniella bostockii is 

more common in the Toolik Lake ACEC than 

previously thought. Not only is the species well 

established north of Toolik Lake, it also occurs to the 

west and south of the lake (Fig. 5) where we located 

populations adjacent to boardwalk trails and research 

sites. We do not know if these populations have 

increased or decreased since they were first 

discovered in 1978. We found this species was 

easiest to locate during its most conspicuous stage - 

flowering. Future inventories should search for 

Claytoniella bostockii at the height of flowering 

(mid-July in the Toolik Lake region). 

The vegetation of the Toolik Lake area is well 

documented (Walker et al. 1994; Walker and Walker 

1996; Walker 2000; Walker et al. In prep). As a 

result, we are able to describe the habitat where 

Claytoniella bostockii was most prolific (Table 4 and 

Fig. 6). Previously, this habitat was described as 

“wet meadows on ridge tops, alpine slopes and by 

lake shores, as well as frost boils, and wet ridge crest 

gravels” (Lipkin and Parker 1995). We found 

Claytoniella bostockii occurred most frequently 

within the Moist and Dry Nonacidic Tundra 

Complexes. These vegetation complexes are 

dominated by Carex membranacea and by 

Astragalus umbellatus and Dryas integrifolia, 

respectively (Walker 2000; Walker et al In prep). 

However, Claytoniella bostockii also occurs within 

several other vegetation complexes to a lesser extent 

including Rich Fen, Snowbed and Moist and Dry 

Acidic Tundra. Therefore, future inventories for 

Claytoniella bostockii should broaden their search 

efforts to include a range of vegetation complexes. 

2 Toolik Lake Rare Plant Inventory, 2002 
 

 

 

 

Literature Cited 

Lipkin, R. 2000. Alaska Botanical Biodiversity: 

AKNHP Vascular Plant Tracking List. 

http://www.uaa.alaska.edu/enri/aknhp_web/ 

biodiversity/botanical/vascular_species_concern/ 

species_table/listall.html (April 4, 2000). 

Lipkin, R. and C. Parker. 1995. Rare vascular 

plants of the BLM Dalton Highway Corridor. 

Alaska Natural Heritage Program, University of
 

Alaska Anchorage and University of Alaska
 

Museum Fairbanks. 79 pp.
 

USDI. 1989. Utility Corridor Proposed Resource
 

Management Plan and Final Environmental Impact
 

Statement. Bureau of Land Management, Northern
 

Field Office, Fairbanks, Alaska.
 

Walker, D. A. and M. D. Walker. 1996. Terrain and 

vegetation of the Imnavait Creek Research Site. In:
 

Reynolds, J. F. and J. D. Tenhunen (eds.). 

Landscape function: implications for ecosystem
 

response to disturbance, a case study in arctic tundra.
 

Springer-Verlag Ecological Studies 120:73-108.
 

Walker, D. A. 2000. The Hierarchical Geobotanical
 

Atlas of the Toolik Lake/Kuparuk River region in 

Northern Alaska. 

http://www.geobotany.uaf.edu/arcticgeobot/
 

tlrveg.html
 

(December 16, 2000)
 

Walker, D. A., M. D. Walker, M. K. Raynolds, A. W. 

Balser, H. A. Maier. In prep. Terrain and vegetation 

of the Toolik Lake Research Site. In: Hobbie, J. 

(ed.). Long-term ecological research in the Arctic. 

Oxford University Press, Inc. 

Walker, M. D., D. A. Walker, and N. A. Auerbach. 

1994. Plant communities of a tussock tundra 

landscape in the Brooks Range Foothills, Alaska. 

Journal of Vegetation Science 5:843-866. 

Toolik Lake Rare Plant Inventory, 2002 3 
TABLE 1. List of rare plants that potentially could occur with in the Toolik Lake ACEC. Global and state species 

ranks are defined in Table 2. 

Scientific Name Global Rank State Rank 

Aster pygmaeus G3 S1S2 

Beckwithia glacialis ssp. alaskensis [=Ranunculus glacialis 

ssp. 1] G4T2 S2 

Claytonia arctica G3 S1 

Draba micropetala G4 S1S2 

Erigeron muirii G2 S2 

Montia bostockii G3 S3 

Oxytropis arctica var. barnebyana G4T2 S2 

Pedicularis hirsuta G5? S1 

Potentilla stipularis G5 S1 

4 Toolik Lake Rare Plant Inventory, 2002 
 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

TABLE 2. Definitions of rare species ranks used by the Alaska Natural Heritage Program (Lipkin 2002). 

Species Global Rankings

G1: Critically imperiled globally 

G2: Imperiled globally 

G3: Rare or uncommon globally 

G4: Apparently secure globally, but cause long-term concern 

G5: Demonstrated secure globally 

G?: Unranked 

G#G#: Global rank of species uncertain, best described as a range between 

the two ranks 

G#Q: Taxonomically questionable 

G#T#: Global rank of species and global rank of the described variety or 

subspecies of the species 

GU: Unrankable 

GH: Historical occurrence 

GX: Extinct 

HYB: Hybrid 

Species State Rankings

S1: Critically imperiled in state 

S2: Imperiled in state 

S3: Rare or uncommon in state 

S4: Apparently secure in state, but with cause for long-term concern 

S5: Demonstrably secure in state 

S#S#: State rank of species uncertain, best described as a range between 

the two ranks 

S?: Unranked 

SU: Unrankable 

SA: Accidental 

SR: Reported from the state, but not yet verified 

SRF: Reported falsely 

SP: Potential to occur in the state 

HYB: Hybrid 

SSYN: Synonym 

Qualifiers

B: Breeding status 

N: Non-breeding status 

?: Inexact 

Q: Questionable taxonomy 

Toolik Lake Rare Plant Inventory, 2002 5 
 

TABLE 3. One mile2 sections surveyed for rare plants in the Toolik Lake ACEC in 2002. 

Meridian Township Range Section Date Surveyed 

Umiat T9S R11E 19 Jul-20-02 

Umiat T9S R11E 20 Jul-19-02 

Umiat T9S R11E 21 Jul-18-02 

Umiat T9S R11E 30 North: Jul-11-02 

South: Jul-16-02 

Umiat T9S R11E 29 North: Jul-19-02 

South: Jul-13-02 

Umiat T9S R11E 28 West: Jul-15-02 

East: Jul-18-02 

Umiat T9S R11E 31 North: Jul-16-02 and Aug-29-02 

South: Jul-20-02 

Umiat T9S R11E 32 North: Jul-13-02 

South: Jul-21-02 

Umiat T9S R11E 33 Jul-17-02 

6 Toolik Lake Rare Plant Inventory, 2002 
 

TABLE 4. Claytoniella bostockii occurrences within the 2002 Toolik Lake ACEC Survey Area. Vegetation 

complexes follow the nomenclature of Walker (2000) and Walker et al. (In prep). “Primary vegetation 

complex” is defined as the most common vegetation type within a polygon, while “Other Vegetation Complex” 

refers to any additional vegetation type overlapped by a polygon. 

ID SHAPE DATE COLLECTOR # OF ABUNDANCE STAGE PRIMARY VEGETATION COMPLEX OTHER VEGETATION COMPLEX 

INDIVIDUALS AREA (m2

) 

1 POLYGON Jul-20-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 79358 MOIST NONACIDIC TUNDRA RICH FEN, DRY ACIDIC TUNDRA 

2 POLYGON Jul-20-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 5757 DRY ACIDIC TUNDRA MOIST NONACIDIC TUNDRA 

3 POLYGON Jul-20-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 1406 SNOWBED RICH FEN 

4 POLYGON Jul-20-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 16905 MOIST NONACIDIC TUNDRA RICH FEN, DRY ACIDIC TUNDRA 

5 POLYGON Jul-20-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 5015 MOIST NONACIDIC TUNDRA 

6 POLYGON Jul-20-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 6957 DRY ACIDIC TUNDRA MOIST NONACIDIC TUNDRA 

7 POLYGON Jul-20-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 3083 DRY ACIDIC TUNDRA 

8 POLYGON Jul-20-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 6528 RICH FEN MOIST ACIDIC TUNDRA 

9 POLYGON Jul-20-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 284588 MOIST NONACIDIC TUNDRA MOIST ACIDIC TUNDRA, SHRUB 

TUNDRA, SNOWBED 

10 POLYGON Jul-20-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 27383 MOIST ACIDIC TUNDRA SNOWBED 

11 POLYGON Jul-20-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 1495 MOIST NONACIDIC TUNDRA 

12 POLYGON Jul-20-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 5483 SHRUB TUNDRA MOIST NONACIDIC TUNDRA 

13 POLYGON Jul-19-02 AMY BREEN 

CARROLL 

100+ FREQUENT FLOWER 920195 MOIST NONACIDIC TUNDRA DRY NONACIDIC TUNDRA, DRY ACIDIC 

TUNDRA, MOIST ACIDIC TUNDRA, 

SNOWBED, RICH FEN, RIPARIAN 

SHRUNLAND 

14 POLYGON Jul-19-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 1591 MOIST NONACIDIC TUNDRA DRY NONACIDIC TUNDRA 

15 POLYGON Jul-19-02 AMY BREEN 

CARROLL 

100+ FREQUENT FLOWER 294105 MOIST NONACIDIC TUNDRA SNOWBED 

16 POLYGON Jul-19-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 2984 MOIST NONACIDIC TUNDRA SNOWBED 

17 POLYGON Jul-19-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 1534 MOIST NONACIDIC TUNDRA 

18 POLYGON Jul-19-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 4033 MOIST NONACIDIC TUNDRA 

19 POLYGON Jul-19-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 27774 MOIST NONACIDIC TUNDRA DRY ACIDIC TUNDRA 

20 POLYGON Jul-19-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 4846 MOIST NONACIDIC TUNDRA 

21 POLYGON Jul-19-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 1207 MOIST NONACIDIC TUNDRA 

22 POLYGON Jul-19-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 17947 MOIST NONACIDIC TUNDRA 

23 POLYGON Jul-19-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 10888 MOIST NONACIDIC TUNDRA 

24 POLYGON Jul-19-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 7084 MOIST NONACIDIC TUNDRA 

25 POLYGON Jul-19-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 2452 MOIST NONACIDIC TUNDRA 

Toolik Lake Rare Plant Inventory, 2002 7 
ID SHAPE DATE COLLECTOR NUMBER OF ABUNDANCE STAGE PRIMARY VEGETATION OTHER VEGETATION COMPLEX 

INDIVIDUALS COMPLEX AREA (m2

) 

26 POLYGON Jul-19-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 5442 DRY ACIDIC TUNDRA 

27 POLYGON Jul-18-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 3873 RIPARIAN SHRUBLAND MOIST ACIDIC TUNDRA 

28 POLYGON Jul-11-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 11073 DRY ACIDIC TUNDRA 

29 POLYGON Jul-11-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 1276 DRY ACIDIC TUNDRA 

30 POLYGON Jul-11-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 61581 DRY ACIDIC TUNDRA SNOWBED 

31 POLYGON Jul-16-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 21204 DRY ACIDIC TUNDRA 

32 POLYGON Jul-16-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 7904 DRY ACIDIC TUNDRA 

33 POLYGON Jul-16-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 14476 DRY ACIDIC TUNDRA MOIST NONACIDIC TUNDRA 

34 POLYGON Jul-16-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 3500 DRY ACIDIC TUNDRA SNOWBED 

35 POLYGON Jul-11-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 516282 MOIST NONACIDIC TUNDRA SNOWBED, DRY ACIDIC TUNDRA 

36 POLYGON Jul-11-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 1599 DRY ACIDIC TUNDRA 

37 POLYGON Jul-16-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 5970 MOIST NONACIDIC TUNDRA DRY ACIDIC TUNDRA, MOIST NONACIDIC 

TUNDRA 

38 POLYGON Jul-16-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 5674 DRY ACIDIC TUNDRA MOIST NONACIDIC TUNDRA, MOIST 

ACIDIC TUNDRA 

39 POLYGON Jul-16-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 3738 MOIST NONACIDIC TUNDRA SNOWBED 

40 POLYGON Jul-11-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 7983 MOIST NONACIDIC TUNDRA DRY NONACIDIC TUNDRA 

41 POLYGON Jul-11-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 20170 MOIST ACIDIC TUNDRA MOIST NONACIDIC TUNDRA, DRY 

NONACIDIC TUNDRA, RICH FEN 

42 POLYGON Jul-11-02 AMY BREEN 

CARROLL 

100+ FREQUENT FLOWER 21682 MOIST ACIDIC TUNDRA 

43 POLYGON Jul-19-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 219714 MOIST NONACIDIC TUNDRA SNOWBED, MOIST ACIDIC TUNDRA, DRY 

NONACIDIC TUNDRA, DRY ACIDIC 

TUNDRA 

44 POLYGON Jul-19-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 2324 MOIST NONACIDIC TUNDRA 

45 POLYGON Jul-19-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 1261 MOIST NONACIDIC TUNDRA 

46 POLYGON Jul-21-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 106750 MOIST NONACIDIC TUNDRA DRY ACIDIC TUNDRA, RIPARIAN 

SHRUBLAND, MOIST ACIDIC TUNDRA 

47 POLYGON Jul-21-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 2917 DRY ACIDIC TUNDRA 

48 POLYGON Jul-21-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 7463 DRY ACIDIC TUNDRA MOIST ACIDIC TUNDRA 

49 POLYGON Jul-21-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 5498 MOIST ACIDIC TUNDRA DRY ACIDIC TUNDRA 

50 POLYGON Jul-13-02 AMY BREEN 

CARROLL 

100+ FREQUENT FLOWER 41776 DRY ACIDIC TUNDRA MOIST NONACIDIC TUNDRA, SHRUB 

TUNDRA 

51 POLYGON Jul-13-02 AMY BREEN 

CARROLL 

100+ FREQUENT FLOWER 41758 DRY ACIDIC TUNDRA MOIST NONACIDIC TUNDRA, SNOWBED 

52 POLYGON Jul-21-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 41814 MOIST ACIDIC TUNDRA DRY ACIDIC TUNDRA 

8 Toolik Lake Rare Plant Inventory, 2002 
ID SHAPE DATE COLLECTOR NUMBER OF ABUNDANCE STAGE PRIMARY VEGETATION OTHER VEGETATION COMPLEX 

INDIVIDUALS COMPLEX AREA (m2

) 

53 POLYGON Jul-21-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 10132 RIPARIAN SHRUBLAND MOIST NONACIDIC TUNDRA, RICH FEN, 

SHRUB TUNDRA 

54 POLYGON Jul-21-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 1145 DRY ACIDIC TUNDRA MOIST NONACIDIC TUNDRA 

55 POLYGON Jul-21-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 2151 RIPARIAN SHRUBLAND SHRUB TUNDRA 

56 POLYGON Jul-13-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 52858 MOIST ACIDIC TUNDRA MOIST NONACIDIC TUNDRA 

57 POLYGON Jul-21-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 4803 MOIST ACIDIC TUNDRA SHRUB TUNDRA, MOIST NONACIDIC 

TUNDRA 

58 POLYGON Jul-21-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 3863 DRY ACIDIC TUNDRA RIPARIAN SHRUBLAND 

59 POLYGON Jul-21-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 4430 MOIST NONACIDIC TUNDRA RICH FEN 

60 POINT Jul-16-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 600 MOIST NONACIDIC TUNDRA 

61 POINT Jul-16-02 AMY BREEN 

CARROLL 

>50 COMMON FLOWER 300 DRY ACIDIC TUNDRA 

62 POINT Jul-16-02 AMY BREEN 

CARROLL 

>50 COMMON FLOWER 150 DRY ACIDIC TUNDRA 

63 POINT Jul-16-02 AMY BREEN 

CARROLL 

<50 COMMON FLOWER 500 MOIST NONACIDIC TUNDRA 

64 POINT Jul-16-02 AMY BREEN 

CARROLL 

<50 COMMON FLOWER 150 SNOWBED 

65 POINT Jul-13-02 AMY BREEN 

CARROLL 

<100 FREQUENT FLOWER 200 MOIST NONACIDIC TUNDRA 

66 POINT Jul-13-02 AMY BREEN 

CARROLL 

100+ COMMON FLOWER 5000 MOIST NONACIDIC TUNDRA 

67 POINT Jul-21-02 AMY BREEN 

CARROLL 

>10 COMMON FLOWER 50 MOIST NONACIDIC TUNDRA 

Toolik Lake Rare Plant Inventory, 2002 9 
FIGURE 1. Map of Alaska showing the location of the Toolik Field Station. Map courtesy of Andrew Balser, Toolik Field Station GIS and Remote 

Sensing Manager. 

10 Toolik Lake Rare Plant Inventory, 2002
 
FIGURE 2. Toolik Lake Research Natural Area and General Land Status. Map courtesy of Andrew Balser, Toolik Lake Field Station GIS and 

Remote Sensing Manager 

Toolik Lake Rare Plant Inventory, 2002 11 
FIGURE 3. Map of the 2002 Rare Plant Survey Area around Toolik Lake. Each square represents a one mile2

 section within the Umiat 

Meridian’s T9S and R11E. Numbers in the center of each square identify the section. 

12 Toolik Lake Rare Plant Inventory, 2002
 
FIGURE 4. Line drawing of Claytoniella bostockii (A. E. Pors.) Jurtz. Synonyms for this species include: Claytonia bostockii A. E. Pors., Montia 

bostockii (A. E. Pors.) Welsh, and Montiastrum bostockii (A. E. Pors.) O. Nilsson. This species was placed in the Claytoniella genus due to its 

pollen and seed morphology and its leaf arrangement (Boris Yurtzev, personal communication). Drawing reproduced with permission of the 

illustrator, Anne-Lillian Schell and the Alaska Natural Heritage Program. 

Toolik Lake Rare Plant Inventory, 2002 13 
FIGURE 5. Aerial photograph map of Claytoniella bostockii occurrences within the 2002 Toolik Lake ACEC Survey Area (aerial photograph, 

August 1982). 

14 Toolik Lake Rare Plant Inventory, 2002 
FIGURE 6. Vegetation map showing Claytoniella bostockii occurrences within the 2002 Toolik Lake ACEC Survey Area. Vegetation complex 

nomenclature follows the classification established by Walker (2000) and Walker et al (In Prep). 

Toolik Lake Rare Plant Inventory, 2002 15