A Chronology of 

Middle Missouri Plains 

Village Sites

By Craig M. Johnson

with contributions by

Stanley A. Ahler, Herbert Haas, and Georges Bonani

Smithsonian Institution

Scholarly Press

Smithsonian Institution

Scholarly Press

Catalog of Type Specimens 

of Recent Mammals:

Rodentia (Myomorpha, 

Anomaluromorpha, and 

Hystricomorpha) in the National 

Museum of Natural History, 

Smithsonian Institution

Robert D. Fisher and Craig A. Ludwig

smithsonian contributions to zoology • number 642
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smithsonian contributions to zoology • numb e r 6 4 2

Catalog of Type Specimens 

of Recent Mammals:

Rodentia (Myomorpha, 

Anomaluromorpha, and 

Hystricomorpha) in the National 

Museum of Natural History, 

Smithsonian Institution

Robert D. Fisher and Craig A. Ludwig

washington d.c.

2014
ABSTRACT

Fisher, Robert D. and Craig A. Ludwig, Catalog of Type Specimens of Recent Mammals: Rodentia (Myomorpha, Anomaluromorpha, and Hystricomorpha) in the National Museum of Natural History, Smithsonian Institution. Smithsonian 

Contributions to Zoology, number 642, vi + 125 pages, 2014.—The type collection of Recent mammals in the Division of 

Mammals, National Museum of Natural History, Smithsonian Institution, contains 945 specimens bearing names of 931 

species-group taxa of Rodentia (Myomorpha, Anomaluromorpha, and Hystricomorpha) as of August 2013. This catalog 

presents an annotated list of these holdings comprised of 905 holotypes, 16 lectotypes, 8 syntypes (48 specimens), and 2 

neotypes. In addition, we include 44 specimens that are part of syntype series that should be in the collection but cannot be 

found or are now known to be in other collections. One hundred and ten of the names are new since the last type catalog 

covering these suborders A lectotype for Mus peruvianus Peale, 1848, is newly designated herein. Nine specimens previously reported were subsequently sent to the vertebrate paleontology collection and are not included here. Suborders and 

families are ordered as in Carleton and Musser; within families, currently recognized genera are arranged alphabetically; 

within each currently recognized genus, accounts are arranged alphabetically by original published name. Information in 

each account includes original name and abbreviated citation thereto, current name if other than original, citation for first 

use of current name combination for the taxon (or new name combination if used herein for the first time), type designation, U.S. National Museum catalog number(s), preparation, age and sex, date of collection and collector, original collector 

number, type locality, and remarks as appropriate. Digital photographs of each specimen will serve as a condition report 

and will be attached to each electronic specimen record.

Cover images: Holotype of Evotomys ungava V. Bailey, 1897 (USNM 186492). Left to right: dorsal, lateral, and ventral 

views of cranium. Photographs by R. Regan.

Published by SMITHSONIAN INSTITUTION SCHOLARLY PRESS

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Copyright © 2014 Smithsonian Institution

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Library of Congress Cataloging-in-Publication Data

National Museum of Natural History (U.S.)

Catalog of type specimens of recent Mammals:Rodentia (Myomorpha, Anomaluromorpha, and 

Hystricomorpha in the National Museum of Natural History, Smithsonian Institution / Robert D. Fisher 

and Craig A. Ludwig.

 pages cm — (Smithsonian contributions to zoology ; number 642) 1. Rodents—Type specimens—

Catalogs and collections—Washington (D.C.) 2. National Museum of Natural History (U.S.)—Catalogs.

I. Fisher, Robert D. II. Ludwig, Craig A. III. Title.

QL737.R6N225 2014

599.35074’753--dc23

2013045347

ISSN: 0081-0282 (print); 1943-6696 (online)

The paper used in this publication meets the minimum requirements of the American National Standard for 

Permanence of Paper for Printed Library Materials Z39.48–1992.
Contents

INTRODUCTION 1

Categories of Name-Bearing Types Included 1

Research Approach 2

Classificatory Organization 2

Format of Accounts 2

Acknowledgments 3

ACCOUNTS OF NAME-BEARING TYPE SPECIMENS 3

Suborder Myomorpha Brandt, 1855 3

 Family Dipodidae G. Fischer, 1817 3

 Genus Allactaga F. Cuvier, 1837 3

 Genus Euchoreutes Sclater, 1891 3

 Genus Jaculus Erxleben, 1777 4

 Genus Napaeozapus Preble, 1899 4

 Genus Sicista Gray, 1827 4

 Genus Zapus Coues, 1875 5

 Family Spalacidae Gray, 1821 6

 Genus Eospalax G. M. Allen, 1938 6

 Genus Spalax Guldenstaedt, 1770 6

 Genus Tachyoryctes Ruppell, 1835 6

 Family Calomyscidae Vorontsov and Potapova, 1979 6

 Genus Calomyscus Thomas, 1905 6

 Family Nesomyidae Major, 1897 6

 Genus Cricetomys Waterhouse, 1840 6

 Genus Dendromus Smith, 1829 7

 Genus Saccostomus Peters, 1846 7

 Genus Steatomys Peters, 1846 7

 Family Cricetidae G. Fischer, 1817 8

 Genus Abrothrix Waterhouse, 1837 8

 Genus Akodon Meyen, 1833 8

 Genus Alticola Blanford, 1881 8

 Genus Arborimus Taylor, 1915 9

 Genus Baiomys True, 1894 9

 Genus Bibimys Massoia, 1979 10

 Genus Calomys Waterhouse, 1837 10
i v • smithsonian contributions to zoology

 Genus Chelemys Thomas, 1903 10

 Genus Dicrostonyx Gloger, 1841 10

 Genus Euneomys Coues, 1874 11

 Genus Geoxus Thomas, 1919 11

 Genus Habromys Hooper and Musser, 1964 11

 Genus Hodomys Merriam, 1894 11

 Genus Holochilus Brandt, 1835 12

 Genus Hyperacrius Miller, 1896 12

 Genus Ichthyomys Thomas, 1893 12

 Genus Isthmomys Hooper and Musser, 1964 12

 Genus Lasiopodomys Lataste, 1887 12

 Genus Lemmiscus Thomas, 1912 12

 Genus Lemmus Link, 1795 13

 Genus Lophiomys Milne-Edwards, 1867 14

 Genus Megadontomys Merriam, 1898 14

 Genus Melanomys Thomas, 1902 14

 Genus Microryzomys Thomas, 1917 14

 Genus Microtus Schrank, 1798 14

 Genus Myodes Pallas, 1811 23

 Genus Myopus Miller, 1910 25

 Genus Neacomys Thomas, 1900 26

 Genus Nectomys Peters, 1861 26

 Genus Nelsonia Merriam, 1897 26

 Genus Neodon Horsfield, 1841 26

 Genus Neofiber True, 1884 26

 Genus Neotoma Say and Ord, 1825 27

 Genus Neotomodon Merriam, 1898 34

 Genus Nephelomys Weksler, Percequillo, and Voss, 2006 34

 Genus Nyctomys Saussure, 1860 34

 Genus Ochrotomys Osgood, 1909 35

 Genus Oecomys Thomas, 1906 35

 Genus Oligoryzomys Bangs, 1900 35

 Genus Ondatra Link, 1795 35

 Genus Onychomys Baird, 1857 36

 Genus Oryzomys Baird, 1857 38

 Genus Osgoodomys Hooper and Musser, 1964 41

 Genus Ototylomys Merriam, 1901 41

 Genus Peromyscus Gloger, 1841 41

 Genus Phenacomys Merriam, 1889 52

 Genus Phodopus Miller, 1910 53

 Genus Phyllotis Waterhouse, 1837 53

 Genus Podomys Osgood, 1909 54

 Genus Reithrodon Waterhouse, 1837 54

 Genus Reithrodontomys Giglioli, 1874 54

 Genus Rheomys Thomas, 1906 59

 Genus Rhipidomys Tschudi, 1845 59

 Genus Scotinomys Thomas, 1913 59

 Genus Sigmodon Say and Ord, 1825 59

 Genus Sigmodontomys J. A. Allen, 1897 62

 Genus Synaptomys Baird, 1857 62

 Genus Thomasomys Coues, 1884 63

 Genus Transandinomys Weksler, Percequillo, and Voss, 2006 63

 Genus Tylomys Peters, 1866 64

 Genus Xenomys Merriam, 1892 64

 Genus Zygodontomys J. A. Allen, 1897 64
number 642 • v

 Family Muridae Illiger, 1811 64

 Genus Acomys Geoffroy, 1838 64

 Genus Aethomys Thomas, 1915 64

 Genus Apodemus Kaup, 1829 64

 Genus Apomys Mearns, 1905 65

 Genus Arvicanthis Lesson, 1842 66

 Genus Bandicota Gray, 1873 66

 Genus Batomys Thomas, 1895 66

 Genus Berylmys Ellerman, 1947 66

 Genus Brachiones Thomas, 1925 66

 Genus Bullimus Mearns, 1905 67

 Genus Bunomys Thomas, 1910 67

 Genus Chiropodomys Peters, 1869 67

 Genus Chrotomys Thomas, 1895 67

 Genus Crateromys Thomas, 1895 68

 Genus Dasymys Peters, 1875 68

 Genus Dephomys Thomas, 1926 68

 Genus Dipodillus Lataste, 1881 68

 Genus Echiothrix Gray, 1867 69

 Genus Eropeplus Miller and Hollister, 1921 69

 Genus Gerbilliscus Thomas, 1897 69

 Genus Gerbillus Desmarest, 1804 69

 Genus Grammomys Thomas, 1915 70

 Genus Hybomys Thomas, 1910 70

 Genus Hylomyscus Thomas, 1926 71

 Genus Lemniscomys Trouessart, 1881 71

 Genus Lenomys Thomas, 1898 71

 Genus Lenothrix Miller, 1903 71

 Genus Leopoldamys Ellerman, 1947 71

 Genus Limnomys Mearns, 1905 73

 Genus Lophuromys Peters, 1874 73

 Genus Mastomys Thomas, 1915 73

 Genus Maxomys Sody, 1936 74

 Genus Melasmothrix Miller and Hollister, 1921 77

 Genus Melomys Thomas, 1922 77

 Genus Meriones Illiger, 1811 77

 Genus Micromys Dehne, 1841 77

 Genus Mus Linnaeus, 1758 77

 Genus Mylomys Thomas, 1906 78

 Genus Myomyscus Shortridge, 1942 78

 Genus Nesokia Gray, 1842 79

 Genus Niviventer Marshall, 1976 79

 Genus Notomys Lesson, 1842 79

 Genus Oenomys Thomas, 1904 79

 Genus Otomys F. Cuvier, 1824 80

 Genus Palawanomys Musser and Newcomb, 1983 80

 Genus Paruromys Ellerman, 1954 80

 Genus Pelomys Peters, 1852 80

 Genus Pithecheirops Emmons, 1993 80

 Genus Praomys Thomas, 1892 80

 Genus Rattus G. Fischer, 1803 80

 Genus Sundamys Musser and Newcomb, 1983 88

 Genus Taeromys Sody, 1941 89

 Genus Tarsomys Mearns, 1905 89

 Genus Taterillus Thomas, 1910 89
v i • smithsonian contributions to zoology

 Genus Thallomys Thomas, 1920 89

 Genus Tokudaia Kuroda, 1943 90

 Genus Tryphomys Miller, 1910 90

 Genus Uranomys Dollman, 1909 90

 Genus Zelotomys Osgood, 1910 90

Suborder Anomaluromorpha Bugge, 1974 90

 Family Anomaluridae Gervais, 1849 90

 Genus Idiurus Matschie, 1894 90

 Family Pedetidae Gray, 1825 90

 Genus Pedetes Illiger, 1811 90

Suborder Hystricomorpha Brandt, 1855 90

 Family Bathyergidae Waterhouse, 1841 90

 Genus Heliophobius Peters, 1846 90

 Family Hystricidae G. Fischer, 1817 90

 Genus Atherurus F. Cuvier, 1829 90

 Genus Hystrix Linnaeus, 1758 91

 Genus Trichys Gunther, 1877 91

 Family Thryonomyidae Pocock, 1922 91

 Genus Thryonomys Fitzinger, 1867 91

 Family Erethizontidae Bonaparte, 1845 91

 Genus Coendou Lacepede, 1799 91

 Genus Erethizon F. Cuvier, 1823 91

 Family Chinchillidae Bennett, 1833 92

 Genus Lagostomus Brookes, 1828 92

 Family Caviidae G. Fischer, 1817 92

 Genus Hydrochoerus Brisson, 1762 92

 Family Dasyproctidae Bonaparte, 1838 92

 Genus Dasyprocta Illiger, 1811 92

 Family Cuniculidae Miller and Gidley, 1918 92

 Genus Cuniculus Brisson, 1762 92

 Family Ctenomyidae Lesson, 1842 92

 Genus Ctenomys Blainville, 1826 92

 Family Echimyidae Gray, 1825 93

 Genus Diplomys Thomas, 1916 93

 Genus Hoplomys J. A. Allen, 1908 93

 Genus Pattonomys Emmons, 2005 93

 Genus Proechimys J. A. Allen, 1899 93

 Family Myocastoridae Ameghino, 1904 93

 Genus Myocastor Kerr, 1792 93

 Family Capromyidae Smith, 1842 94

 Genus Geocapromys Chapman, 1901 94

 Genus Mysateles Lesson, 1842 94

 Genus Plagiodontia F. Cuvier, 1836 94

REFERENCES 95

INDEX OF TAXA 109
INTRODUCTION

The Division of Mammals, National Museum of Natural History (NMNH), Smithsonian Institution, formerly the United States National Museum (USNM), maintains 

the second largest collection of mammal primary types in the world, numbering 3,220 

specimens. Type specimens are “the international standards of reference that provide 

objectivity in zoological nomenclature,” and as such, “every institution in which namebearing types are deposited should publish lists of name-bearing types in its possession or 

custody” (International Commission on Zoological Nomenclature, 1999:79).

This is the fourth type catalog of USNM mammals published to date, following Lyon 

and Osgood (1909; 1,415 specimens reported), Poole and Schantz (1942; 2,824 specimens reported), and Fisher and Ludwig (2012; 843 specimens reported for the Order 

Rodentia, suborders Sciuromorpha and Castorimorpha). However, we continue to deviate from the two earlier catalogs and follow the precedent set by the NMNH Division of 

Fishes (Vari and Howe, 1991) and Division of Amphibians and Reptiles (Reynolds et al., 

2007), as continued by Fisher and Ludwig (2012), in which the type collection is divided 

into smaller, more manageable taxonomic reports, rather than addressed in one monograph. The current work, the second of an anticipated four such publications, completes 

the order Rodentia (suborders Myomorpha, Anomaluromorpha, and Hystricomorpha).

Categories of Name-Bearing Types Included

Specimens included in this catalog are one of the following as defined by the International Code of Zoological Nomenclature (International Commission on Zoological 

Nomenclature, 1999; hereafter ICZN).

Holotype. The single specimen upon which a new nominal species-group taxon is 

based in the original publication (Article 73.1).

Syntype. Any one of the multiple specimens on which a species-group name is 

equally based when no holotype was specified in the original publication and no subsequent designation of a lectotype has been published (Article 73.2). Syntypes are referred 

to as cotypes in Lyon and Osgood (1909) and Poole and Schantz (1942) as well as in 

other early literature.

Lectotype. One specimen selected from two or more specimens upon which a speciesgroup name is originally based and designated in a publication subsequent to the original 

description to become the unique name bearer (Article 74.1).

Catalog of Type Specimens of Recent Mammals:

Rodentia (Myomorpha, Anomaluromorpha, 

and Hystricomorpha) in the National Museum 

of Natural History, Smithsonian Institution

Robert D. Fisher, U.S. Geological Survey, 

Patuxent Wildlife Research Center, National Museum of Natural History, Smithsonian Institution, 

Washington, D.C. 20013-7012, USA; Craig A. 

Ludwig, Division of Mammals, Department of 

Vertebrate Zoology, National Museum of Natural History, Smithsonian Institution, Washington, D.C. 20013-7012, USA. Correspondence: 

fisherr@si.edu; ludwigc@si.edu. Manuscript received 26 March 2013; accepted 17 September 

2013.
2 • smithsonian contributions to zoology

Neotype. A specimen chosen as the name bearer of a species-group taxon if no holotype, lectotype, syntype, or prior neotype is believed to be extant (Article 75.1).

Research Approach

In the early 1970s, accounts from a copy of Poole and 

Schantz (1942) were physically cut out and pasted onto 5 × 7 

index cards to create a master card file of types. Additional cards 

were created and added to the file as new taxa were described 

and their types installed in the collection. We checked the information on each card against the original description, the original skin label (when present), and the field catalog entry (when 

available). Information on each index card, often annotated by 

hand, thus served as the nucleus of each of the accounts. Other 

unpublished catalogs, such as the Museum Catalog, Merriam 

Collection Catalog, and X-Catalog, were consulted as needed.

Authors in Wilson and Reeder (2005) are followed for currently accepted names unless an online search of Zoological Record uncovered a more recent taxonomic arrangement. In the 

taxonomic order presented herein (see the next section), this 

search was conducted through November 2011 for the beginning 

accounts and December 2012 for the later accounts. Each specimen record in the collection database was corrected as needed to 

reflect new information. The type publication citation was added 

to the electronic record if not already present. 

Digital images will be created for each specimen and will include dorsal, lateral, and ventral views of crania; dorsal and lateral views of mandibles; dorsal and ventral views of skins (lateral 

if possibly diagnostic); and both sides of all labels. All images 

will feature a metric scale bar. Additional views will be included 

if deemed appropriate. For lateral images, the traditional view 

of the left aspect will be presented, except for skulls whose right 

side is more complete because of damage on the left. We will 

record specimen damage not captured in these images in the associated metadata when readily apparent but will not inspect 

specimens for minute damage. Images will be uploaded to the 

specimen database as they become available and can be accessed 

via the Internet (http://collections.mnh.si.edu/search/mammals/). 

Images will be created in the order in which accounts are presented. We anticipate it will take about four years for all images 

to be completed for specimens in this catalog. These images will 

not only serve as indicators of specimen condition but also will 

allow researchers to verify certain characters of a type without 

having to visit the USNM and handle the specimens, thereby 

providing a convenience to researchers and minimizing possible 

damage to irreplaceable specimens.

Classificatory Organization

Suborders and families are sequentially arranged following 

Carleton and Musser (2005). Within families, currently recognized genera are arranged alphabetically. Within each currently 

recognized genus, the accounts are arranged alphabetically by 

original published name.

Format of Accounts

We follow the format set by Fisher and Ludwig (2012) with 

two exceptions. Type locality is now set apart, as a subheading, 

as is remarks, where appropriate.

Each account reads as follows:

• Original published name: genus, species, and subspecies 

in bold followed by the author’s or authors’ names.

• Abbreviated type citation: Publication, volume, page, 

and date of publication. Dates are given as day, threeletter abbreviation for month, and year throughout.

• Currently recognized name: Genus, species, and subspecies, if different from the original published name, followed by the citation for the first use of current name 

combination for the taxon. Author names in parentheses 

indicate the generic name used in the original description 

is different than the current one. The statement “new 

name combination” replaces the citation if the name is 

used for the first time herein. Unless more recent authors 

are cited, current usage follows the authors in Wilson 

and Reeder (2005): Myomorpha (Holden and Musser, 

2005; Musser and Carleton, 2005), Anomaluromorpha 

(Dieterlen, 2005), and Hystricomorpha (Woods and Kilpatrick, 2005). The aforementioned authors of the sections covering Myomorpha and Anomaluromorpha did 

not employ currently recognized subspecies and listed 

them as synonyms of unspecified rank of the genus and 

species to which they are currently assigned. We give a 

recent citation or two for their use where appropriate, 

primarily for North and Central American taxa.

• Type designation (holotype, lectotype, neotype, or syntype): USNM catalog number(s), preparation of specimen, age and sex, date collected and collector(s) name(s), 

and original number, which is usually the field number.

• Type locality: Exact locality as given in the original description or lectotype designation, published restrictions, 

and supplementary data. Abbreviations are used for 

miles (mi), kilometers (km), feet (ft), and meters (m).

• Remarks: As needed, but especially to explain types designated prior to becoming part of the USNM collection 

and reasons for considering certain specimens to be types 

when not specified as such in the original publication.

We have made every effort to distinguish information that 

was included in the original description from that added later 

from other sources. Added information, such as amended or restricted localities, is given with citations. Brackets are used to 

identify supplementary data not contained in the original description and, when not from commonly used sources such as 

maps or online gazetteers, are attributed to a collector’s Field 

Catalog (FC), the Smithsonian Institution’s Museum Catalog 

(MC), the Merriam Collection Catalog (MCC), the skin label 

(SL), the X-Catalog (XC), or a citation. Parentheses are placed 

around metric equivalents and current place-names when this 

information was not included in the original publication.
number 642 • 3

In the early years of the USNM a double-number cataloging 

system was used in which skins and alcoholics were entered in 

one catalog and skeletal material in another. Many older specimens therefore have two numbers associated with them, and 

the proper museum record is indicated by a fraction such as 

23651/31045. In this example, the numerator on a skin label 

indicates 23651 as the skin to which skull 31045 belongs; on 

the associated skull label, 31045/23651, the numerator indicates 31045 as a skull to which the skin 23651 belongs. This 

cataloging system was abandoned at the end of 1892, after 

which a single, continuous numbering system was implemented 

for all specimen parts. The new system was initiated in January 1893, beginning with number 50001. The C. Hart Merriam 

private collection (MCC), donated to the USNM in 1911 and 

mentioned in many accounts, also used the double-numbering 

system.

Finally, the X-Catalog requires explanation. Beginning with 

the Biological Survey and continuing with the Fish and Wildlife Service, specimens incidentally acquired through exchange 

or purchase or received from or collected by persons other than 

permanent staff were entered in this catalog prior to being transferred to the Division of Mammals for cataloging; X-Catalog 

numbers are identified with an “-X” suffix. The X-Catalog thus 

served as an ersatz field catalog, and its entries frequently have 

data (referenced as XC) that were never entered into the museum 

catalog or onto skin labels. The first entry in the X-Catalog was 

in 1896 and the last was on 5 October 1951.

Acknowledgments

We thank Robert P. Reynolds, U.S. Geological Survey 

(USGS) Patuxent Wildlife Research Center, NMNH, and 

Kristofer M. Helgen, Division of Mammals, NMNH, the supervisors of Fisher and Ludwig, respectively, for support and encouragement to undertake this project. Funds to purchase the 

photographic setup, in order of significance, were provided by 

Carol E. Butler, assistant director for collections, NMNH; Don 

E. Wilson, chair, Department of Vertebrate Zoology, NMNH 

(since retired); and Robert P. Reynolds, to whom we are thankful. We are greatly indebted to David F. Schmidt, Division of 

Mammals, NMNH (since retired), with support from Linda K. 

Gordon, Division of Mammals, NMNH (since retired), who 

spent a considerable amount of time researching photographic 

equipment and methods, ordered the equipment, and set up the 

photographic lab. Advice on photographic equipment, setup, 

and technique was graciously provided to David by Melvin J. 

Wachowiak and Elizabeth Webb, Museum Conservation Institute, Smithsonian Institution; Roy and Jason Larimer, Visionary 

Digital; and James L. Di Loreto, Office of Imaging and Photographic Services, NMNH. Renee Regan, a contractor working 

on funds provided by the Smithsonian Institution, Collection 

Information System, Information Resources Management pool 

funds, is currently doing photography for our earlier type catalog, and we hope to obtain additional funds to continue photography for the current publication. Alfred L. Gardner, USGS 

Patuxent Wildlife Research Center, NMNH, provided valuable 

insight to numerous taxonomic issues and guidance in interpreting various provisions of the International Code of Zoological 

Nomenclature. Michael D. Carleton, Division of Mammals, 

NMNH, and Robert P. Reynolds provided valuable comments 

that greatly improved the manuscript.

ACCOUNTS OF NAME-BEARING 

TYPE SPECIMENS

Order Rodentia Bowdich, 1821

Suborder Myomorpha Brandt, 1855

Family Dipodidae G. Fischer, 1817

Genus Allactaga F. Cuvier, 1837

Allactaga grisescens Hollister

Smithson. Misc. Collect. 60(14): 2, 29 Nov 1912.

= Allactaga sibirica (Forster, 1778). See Ognev (1948:185). Assigned to A. sibirica as synonym of unspecified rank by 

Holden and Musser (2005:877).

Holotype: USNM 175494. Skin and skull. Adult male. Collected 

28 Jul 1912 by N. Hollister. Original number 4395.

Type Locality: Chuisaya Steppe, 8 mi (13 km) south of KoshAgatch (Kosh-Agach), Siberia [Russia].

Allactaga mongolica longior Miller

Proc. Biol. Soc. Wash. 24: 54, 24 Feb 1911a.

= Allactaga sibirica (Forster, 1778). See Ognev (1948:183). 

Assigned to A. sibirica as synonym of unspecified rank 

by Holden and Musser (2005:877), but see Shenbrot 

(1993:105), who considered longior a valid subspecies of 

A. sibirica.

Holotype: USNM 155183. Skin and skull. Adult female. Collected 26 Jul 1909 by A. de C. Sowerby. Original number 

204.

Type Locality: 15 mi (24 km) northeast of Ching-ning-chow, altitude 6,200 ft (1,890 m), Kansu (Gansu), China.

Genus Euchoreutes Sclater, 1891

Euchoreutes naso alashanicus A. B. Howell

Proc. Biol. Soc. Wash. 41: 42, 16 Mar 1928.

Holden and Musser (2005:886) listed alashanicus as a synonym of unspecified rank; Corbet (1978:158), considered 

alashanicus a valid subspecies.

Holotype: USNM 240764. Skin, skull, and [trunk] skeleton. 

Adult female. Collected 26 Apr 1923 by F. R. Wulsin. Original number 1054.

Type Locality: Alashan Desert, Inner Mongolia, 100 mi (161 

km) north-northwest of Ningsia (Ningxia), Kansu (Nei 

Monggol), China.
4 • smithsonian contributions to zoology

Genus Jaculus Erxleben, 1777

Subspecies of Jaculus jaculus were not recognized by Holden 

and Musser (2005:884). We follow Corbet (1978:151–152) for 

the trinomial epithets.

Jaculus deserti fuscipes Ranck

Bull. U.S. Natl. Mus. 275: 226, 2 Oct 1968.

= Jaculus jaculus jaculus (Linnaeus, 1758). See Corbet 

(1978:152).

Holotype: USNM 322761. Skin and skull. Adult female. Collected 3 Dec 1961 by G. L. Ranck. Original number 909.

Type Locality: 7 km S El Gheddahia (Al Qaddahiyah), [Ranck 

(1968:68): 31°23′N, 15°12′E], Tripolitania Province, Libya.

Jaculus deserti rarus Ranck

Bull. U.S. Natl. Mus. 275: 228, 2 Oct 1968.

= Jaculus jaculus jaculus (Linnaeus, 1758). See Corbet 

(1978:151).

Holotype: USNM 319781. Skin and skull. Adult male. Collected 

26 Mar 1961 by H. W. Setzer. Original number 2987.

Type Locality: Ain Zueia (Ayn Zuwayyah), Gebel Uweinat, 

[Ranck (1968:68): 21°53′N, 24°50′E], Cyrenaica Province, 

Libya.

Jaculus deserti vastus Ranck

Bull. U.S. Natl. Mus. 275: 230, 2 Oct 1968.

= Jaculus jaculus jaculus (Linnaeus, 1758). See Corbet (1978: 

151).

Holotype: USNM 325837. Skin and skull. Adult male [SL: female]. Collected 29 Apr 1962 by G. L. Ranck. Original 

number 2116.

Type Locality: Wadi er Rueis (Wadi ar Ru’ays), Gebel el Herug 

el Asued, Tazerbo Oasis, [Ranck (1968:67): 26°52′N, 

18°31′E], Cyrenaica Province, Libya.

Jaculus jaculus arenaceous Ranck

Bull. U.S. Natl. Mus. 275: 234, 2 Oct 1968.

= Jaculus jaculus jaculus (Linnaeus, 1758). See Corbet 

(1978:152).

Holotype: USNM 322812. Skin and skull. Adult male. Collected 

9 Feb 1962 by G. L. Ranck. Original number 1624.

Type Locality: Edri (Idri), [Ranck (1968:68): 27°32′N, 13°12′E], 

Fezzan Province, Libya.

Jaculus jaculus collinsi Ranck

Bull. U.S. Natl. Mus. 275: 236, 2 Oct 1968.

= Jaculus jaculus jaculus (Linnaeus, 1758). See Corbet 

(1978:151).

Holotype: USNM 325797. Skin and skull. Adult male. Collected 

15 Apr 1962 by G. L. Ranck. Original number 2039.

Type Locality: Tazerbo Oasis (Tazirbu), [Ranck (1968:67): 

25°45′N, 21°09′E], Cyrenaica Province, Libya.

Jaculus jaculus cufrensis Ranck

Bull. U.S. Natl. Mus. 275: 238, 2 Oct 1968. 

= Jaculus jaculus jaculus (Linnaeus, 1758). See Corbet (1978:151).

Holotype: USNM 325785. Skin and skull. Adult female. Collected 2 Apr 1962 by G. L. Ranck. Original number 1898.

Type Locality: El Giof (Al Jawf), Cufra Oasis (Al Kufrah), [Ranck 

(1968:67): 24°11′N, 23°19′E], Cyrenaica Province, Libya.

Jaculus jaculus tripolitanicus Ranck

Bull. U.S. Natl. Mus. 275: 240, 2 Oct 1968.

= Jaculus jaculus jaculus (Linnaeus, 1758). See Corbet (1978:152).

Holotype: USNM 325773. Skin and skull. Adult male. Collected 

6 Mar 1962 by G. L. Ranck. Original number 1725.

Type Locality: 25 km N Gharian (Gharyan), [Ranck (1968:68): 

32°25′N, 13°03′E], Tripolitania Province, Libya.

Jaculus jaculus whitchurchi Ranck

Bull. U.S. Natl. Mus. 275: 242, 2 Oct 1968.

= Jaculus jaculus jaculus (Linnaeus, 1758). See Corbet 

(1978:151).

Holotype: USNM 325807. Skin and skull. Adult male. Collected 

18 Jun 1962 by G. L. Ranck [SL: and J. H. Shaw]. Original 

number GLR 2362.

Type Locality: 10 km S Agedabia (Ajdabiya), [Ranck (1968:67): 

30°42′N, 20°19′E], Cyrenaica Province, Libya.

Genus Napaeozapus Preble, 1899

Napaeozapus insignis frutectanus Jackson

Proc. Biol. Soc. Wash. 32: 9, 14 Feb 1919.

Holden and Musser (2005:892) listed frutectanus as a synonym of unspecified rank; Hall (1981:847) and Whitaker 

(1999a:666) recognized frutectanus as a valid subspecies.

Holotype: USNM 227349. Skin and skull. Adult male. Collected 

6 Sep 1917 by H. H. T. Jackson. Original number 896.

Type Locality: Crescent Lake, Oneida County, Wisconsin.

Zapus (Napaeozapus) insignis roanensis Preble

N. Am. Fauna 15: 35, 8 Aug 1899a.

= Napaeozapus insignis roanensis (Preble, 1899). See Miller 

(1900a:114). Assigned to N. insignis as synonym of unspecified rank by Holden and Musser (2005:893); Hall 

(1981:849) and Whitaker (1999a:666) recognized roanensis

as a valid subspecies.

Holotype: USNM 66283. [Skin and skull]. Adult male. Collected 

22 May 1894 by A. G. Wetherby. Original number [SL: 5].

Type Locality: Magnetic City, foot of Roan Mountain, [Mitchell 

County], North Carolina.

Genus Sicista Gray, 1827

Sicista napaea Hollister

Smithson. Misc. Collect. 60(14): 2, 29 Nov 1912.
number 642 • 5

Holotype: USNM 175195. Skin and skull. Adult male. Collected 

6 Aug 1912 by N. Hollister. Original number 4427.

Type Locality: Tapucha, Altai Mountains, Siberia, [Russia]. Restricted to 5 mi (8 km) south of Tapucha at 6,875 ft (2,096 m) 

by Hollister (1913a:521).

Sminthus flavus True

Proc. U.S. Natl. Mus. 17: 341, 15 Nov 1894a.

= Sicista concolor (Buchner, 1892). Assigned as synonym of unspecified rank by Holden and Musser (2005:892).

Holotype: USNM 20140/35503. Skin and skull. Adult male. 

Collected [SL and FC: 21] Jul 1891 by W. L. Abbott. No 

original number.

Type Locality: Central Kashmir, 11,000 ft (3,353 m), [Jammu 

and Kashmir], India.

Genus Zapus Coues, 1874

Holden and Musser (2005:892) did not recognize subspecies in the species of Zapus. We follow various authors for their 

current use.

Zapus hudsonius alascensis Merriam

Proc. Biol. Soc. Wash. 11: 223, 15 Jul 1897a.

Hall (1981:842) and Whitaker (1999b:667) recognized alascensis as a valid subspecies.

Holotype: USNM 73584. [Skin and skull]. Adult male. Collected 

5 Jul 1895 by C. P. Streator. Original number 4660.

Type Locality: Yakutat Bay, Alaska.

Zapus hudsonius campestris Preble

N. Am. Fauna 15: 20, 20 Aug 1899a.

King et al. (2006:4345) recognized campestris as a valid subspecies.

Holotype: USNM 65872. [Skin and skull]. Adult male. Collected 

12 Jun 1894 by B. H. Dutcher. Original number 600.

Type Locality: Bear Lodge Mountains, [FC: 5,100 ft (1,554 m), 

Crook County], Wyoming.

Zapus hudsonius preblei Krutzsch

Univ. Kans. Publ. Mus. Nat. Hist. 7: 452, 21 Apr. 1954.

King et al. (2006:4345) recognized preblei as a valid subspecies.

Holotype: USNM 73085. [Skin and skull]. Adult male. Collected 

23 Jul 1895 by E. A. Preble. Original number 435.

Type Locality: Loveland, [FC: 5,000 ft (1,524 m)], Larimer 

County, Colorado.

Zapus luteus Miller

Proc. Biol. Soc. Wash. 24: 253, 23 Dec 1911b.

= Zapus hudsonius luteus Miller, 1911. See Hafner et al. 

(1981:501). King et al. (2006:4345) considered luteus valid 

as a subspecies.

Holotype: USNM 133601. Skin and skull. Adult female. Collected 24 Jun 1904 by M. Surber. Original number 162.

Type Locality: Espanola, [SL and FC: altitude 5,000 ft (1,524 m)], 

[Rio Arriba County], New Mexico.

Zapus luteus australis V. Bailey

Proc. Biol. Soc. Wash. 26: 132, 21 May 1913.

= Zapus hudsonius luteus Miller, 1911. See Hafner et al. 

(1981:509). King et al. (2006:4345) considered luteus valid 

as a subspecies.

Holotype: USNM 160731. [Skin and skull]. Young adult female. 

Collected 23 Aug 1909 by E. A. Goldman. Original number 

[FC: Nelson and Goldman] 20402.

Type Locality: Socorro, [Socorro County], New Mexico.

Zapus major Preble

N. Am. Fauna 15: 24, 8 Aug 1899a.

= Zapus princeps oregonus Preble, 1899. See Krutzsch (1954: 

370). Hart et al. (2004:1) recognized oregonus as a valid 

subspecies.

Holotype: USNM 79983. [Skin and skull]. Adult female. Collected 4 Aug 1896 by C. H. Merriam and V. Bailey. Original 

number [FC: VB] 5720.

Type Locality: Warner Mountains, [FC: altitude 6,500 ft 

(1,981 m)], [Lake County], Oregon.

Zapus nevadensis Preble

N. Am. Fauna 15: 25, 8 Aug 1899a.

= Zapus princeps oregonus Preble, 1899. See Krutzsch (1954: 

370). Hart et al. (2004:1) recognized oregonus as a valid 

subspecies.

Holotype: USNM 94185. [Skin and skull]. Adult female. Collected 21 Jun 1898 by V. Bailey. Original number 6581.

Type Locality: [FC: Near Ruby P O, altitude 9,000 ft (2,743 m)], 

Ruby Mountains, [Elko County], Nevada.

Zapus pacificus Merriam

Proc. Biol. Soc. Wash. 11: 104, 26 Apr 1897b.

= Zapus princeps pacificus Merriam, 1897. See Krutzsch 

(1954:412). Hart et al. (2004:1) recognized pacificus as a 

valid subspecies.

Holotype: USNM 80445. [Skin and skull]. Young adult male. 

Collected 29 Aug 1896 by E. A. Preble. Original number 

1454.

Type Locality: Prospect, Rogue River Valley, [FC: Jackson 

County], Oregon.

Zapus princeps minor Preble

N. Am. Fauna 15: 23, 8 Aug 1899a.

Hart et al. (2004:1) recognized minor as a valid subspecies.

Holotype: USNM 73673. [Skin and skull]. Adult female. 

Collected 23 Jul 1895 by J. A. Loring. Original number 3123.

Type Locality: Wingard, near Carlton House, Saskatchewan, 

[SL: Canada].
6 • smithsonian contributions to zoology

Zapus princeps oregonus Preble

N. Am. Fauna 15: 24, 8 Aug 1899a.

Hart et al. (2004:1) recognized oregonus as a valid subspecies.

Holotype: USNM 78156. [Skin and skull]. Adult male. Collected 

29 May 1896 by E. A. Preble. Original number 959.

Type Locality: Elgin, Blue Mountains, [FC: Union County], 

Oregon.

Zapus tenellus Merriam

Proc. Biol. Soc. Wash. 11: 103, 26 Apr 1897b.

= Zapus hudsonius tenellus Merriam, 1897. See Hall (1934:377). 

Hall (1981:844) and Whitaker (1999b:667) recognized 

tenellus as a valid subspecies.

Holotype: USNM 66932. [Skin and skull]. Adult female. Collected 25 Aug 1894 by C. P. Streator. Original number 4196.

Type Locality: Kamloops, British Columbia, [Canada].

Zapus trinotatus montanus Merriam

Proc. Biol. Soc. Wash. 11: 104, 26 Apr 1897b.

Hall (1981:846) and Gannon (1999:670) recognized montanus

as a valid subspecies.

Holotype: USNM 79863. [Skin and skull]. Adult female. Collected 21 Aug 1896, not 19 Aug as in original description, 

by E. A. Preble. Original number 1388.

Type Locality: Crater Lake, [Lyon and Osgood (1909:43): 

Mount Mazama], [FC: Klamath County], Oregon.

Family Spalacidae Gray, 1821

Genus Eospalax G. M. Allen, 1938

Myotalpa cansus Lyon

Smithson. Misc. Collect. 50: 134, 9 Jul 1907a.

= Eospalax cansus (Lyon, 1907). See Zheng (1994:62). Specific 

validity upheld by Zhou and Zhou (2008:290).

Holotype: USNM 144022. Skin and skull. Adult female. Collected 7 May 1906 by W. W. Simpson. Original number 7.

Type Locality: Taocheo, Kan-su (Gansu), northwestern China.

Genus Spalax Guldenstaedt, 1770

Spalax berytensis Miller

Proc. Biol. Soc. Wash. 16: 162, 30 Nov 1903a.

= Spalax ehrenbergi Nehring, 1898. See Ellerman (1951:556).

Holotype: USNM 13009/36859. Skin and skull. Adult female. 

Collected Apr 1878 by W. T. Van Dyck. Original number 

[SL: 557].

Type Locality: Beyrout (Beirut), [Beyrouth Province], Syria 

(Lebanon).

Spalax dolbrogeae Miller

Proc. Biol. Soc. Wash. 16: 161, 30 Nov 1903a.

= Spalax leucodon Nordman, 1840. See Ellerman (1951:556).

Holotype: USNM 122109. Skin and skull. Adult male. Collected 

20 Mar 1903. Collector unknown [MC: Received from 

W. Schluter, Halle, Germany]. 

Type Locality: Malcocui, [SL: near Tulchea], Dobrudscha (Dobruja), [Tulcea Province], Rumania (Romania).

Genus Tachyoryctes Ruppell, 1835

Tachyoryctes rex Heller

Smithson. Misc. Collect. 56(9): 4, 22 Jul 1910a.

Holotype: USNM 163088. Skin and skull. Adult male. Collected 

7 Oct 1909 by J. A. Loring. Original number 7840.

Type Locality: Western slope of Mount Kenia (Kenya), altitude 

10,000 ft (3,048 m), [Kenya].

Family Calomyscidae Vorontsov and Potapova, 1979

Genus Calomyscus Thomas, 1905

Calomyscus bailwardi grandis Schlitter and Setzer

Proc. Biol. Soc. Wash. 86: 163, 31 May 1973.

= Calomyscus grandis Schlitter and Setzer, 1973. See Pavlinov et 

al. (1995:79).

Holotype: USNM 369548. Skin and skull. Adult female. Collected 28 Jun 1965 by R. G. Tuck Jr. Original number 1704.

Type Locality: 11 km ENE Fasham, Teheran (Tehran) Province, 

Iran.

Family Nesomyidae Major, 1897

Genus Cricetomys Waterhouse, 1840

Cricetomys gambianus enguvi Heller

Smithson. Misc. Collect. 59(16): 16, 5 Jul 1912.

= Cricetomys ansorgei Thomas, 1904. See Musser and Carleton 

(2005:932).

Holotype: USNM 181805. [Skin and skull]. Adult female. Collected 13 Nov 1911 by E. Heller. Original number 4741.

Type Locality: [FC: Summit of] Mount Umengo, altitude 5,000 ft 

(1,524 m), Taita Mountains, [Coast Province], British East 

Africa (Kenya).

Cricetomys gambianus osgoodi Heller

Smithson. Misc. Collect. 59(16): 16, 5 Jul 1912.

= Cricetomys ansorgei Thomas, 1904. See Musser and Carleton 

(2005:932).

Holotype: USNM 181806. [Skin and skull]. Aged male. Collected 20 Dec 1911 by E. Heller. Original number [SL: 

4926], not 7425 as in original description.

Type Locality: Mazeras, [Coast Province], British East Africa (Kenya).

Cricetomys gambianus raineyi Heller

Smithson. Misc. Collect. 59(16): 15, 5 Jul 1912.
number 642 • 7

= Cricetomys ansorgei Thomas, 1904. See Musser and Carleton 

(2005:932).

Holotype: USNM 181804. [Skin and skull]. Adult male. Collected 31 Aug 1911 by E. Heller. Original number 4187.

Type Locality: [SL: North Creek], Mount Gargues (Uaragess), 

altitude 6,000 ft (1,829 m), Mathews Range, [Rift Valley], 

British East Africa (Kenya).

Genus Dendromus Smith, 1829

Dendromus lineatus Heller

Smithson. Misc. Collect. 56(17): 4, 28 Feb 1911.

= Dendromus mystacalis (Heuglin, 1863). See Dieterlen 

(1971:103).

Holotype: USNM 164816. [Skin and skull]. Adult male. Collected 1 Feb 1910 by J. A. Loring. Original number 8921.

Type Locality: Rhino Camp, Lado [Enclave, Uganda].

Dendromus mesomelas percivali Heller

Smithson. Misc. Collect. 59(16): 5, 5 Jul 1912.

= Dendromus insignis (Thomas, 1903). See Thomas (1916:242).

Holotype: USNM 181791. [Skin and skull]. Adult female. Collected 26 Aug 1911 by A. B. Percival. Original number 

E. Heller 4100.

Type Locality: [SL: Summit of] Mount Gargues (Uaragess), altitude 7,000 ft (2,134 m) [SL: 7,100 ft (2,164 m)], Mathews 

Range, [Rift Valley], British East Africa (Kenya).

Dendromus spectabilis Heller

Smithson. Misc. Collect. 56(17): 3, 28 Feb 1911.

= Dendromus melanotis (Smith, 1834). See von Bohmann 

(1942:51).

Holotype: USNM 164815. [Skin and skull]. Adult female. Collected 18 Jan 1910 by J. A. Loring. Original number 8783.

Type Locality: Rhino Camp, Lado [Enclave, Uganda].

Dendromus whytei capitis Heller

Smithson. Misc. Collect. 59(16): 6, 5 Jul 1912.

= Dendromus mystacalis (Heuglin, 1863). See Musser and Carleton (1993:543).

Holotype: USNM 181792. [Skin and skull]. Adult female. Collected 8 Sep 1911 by E. Heller. Original number 4263.

Type Locality: [SL: Summit of] Mount Lololokui, altitude 6,000 

ft (1,829 m), Mathews Range, [Rift Valley], British East 

Africa (Kenya).

Dendromys nigrifrons True

Proc. U.S. Natl. Mus. 15: 462, 26 Oct 1892.

= Dendromus melanotis (Smith, 1834). See von Bohmann 

(1942:50).

Lectotype: USNM 19783/35263 as designated by Lyon and Osgood (1909:158). Alcoholic, skull removed. Female. Collected [Lyon and Osgood (1909:158): November] 1889(?) 

by W. L. Abbott. [MC: Skin cataloged 26 Sep 1891; skull 25 

Sep 1891]. Original number [MC: 10].

Type Locality: Mount Kilima-Njaro (Kilimanjaro), altitude 

[Lyon and Osgood (1909:158): 5,000 ft (1,524 m)], [Kilimanjaro], East Africa (Tanzania).

Remarks: As explained by Lyon and Osgood (1909:158) and Poole 

and Schantz (1942:454), a type was not designated by number in the original description, but the skull, USNM 35263, 

of one of Abbott’s five specimens was figured and marked 

“typical specimen.” This is the specimen that Lyon and Osgood (1909:158) regarded as “the type,” which constituted 

designation of a lectotype (ICZN, 1999: Article 74.5).

Genus Saccostomus Peters, 1846

Saccostomus isiolae Heller

Smithson. Misc. Collect. 59(16): 14, 5 Jul 1912.

= Saccostomus mearnsi Heller, 1910. See G. M. Allen and Lawrence (1936:102).

Holotype: USNM 181803. [Skin and skull]. Adult female. Collected 1 Jul 1911 by E. Heller. Original number 1908.

Type Locality: Isiola River, Northern Guaso Nyiro, British East 

Africa (Kenya).

Saccostomus mearnsi Heller

Smithson. Misc. Collect. 54(4): 3, 28 Feb 1910b.

Holotype: USNM 162882. [Skin and skull]. Adult male. Collected 28 Nov 1909 by E. A. Mearns. Original number 7292.

Type Locality: Changamwe, [Coast Province], British East Africa 

(Kenya).

Saccostomus umbriventer Miller

Smithson. Misc. Collect. 54(5): 1, 28 Feb 1910a.

= Saccostomus mearnsi Heller, 1910. See G. M. Allen and Lawrence (1936:102).

Holotype: USNM 162612. Skin and skull. Adult female. Collected 26 Jun 1909 by J. A. Loring [FC: and E. Heller]. 

Original number [FC: JAL 6447].

Type Locality: Njori Osolali (Sotik), [Rift Valley], British East 

Africa (Kenya).

Genus Steatomys Peters, 1846

Steatomys athi Heller

Smithson. Misc. Collect. 54(4): 3, 28 Feb 1910b.

= Steatomys parvus Rhoads, 1896. See Coetzee (1977:2).

Holotype: USNM 162883. [Skin and skull]. Adult male. Collected 22 Nov 1909 by J. A. Loring. Original number 8259.

Type Locality: Lukenia Hills, Athi Plains, British East Africa 

(Kenya).

Steatomys caurinus roseveari Swanepoel and Schlitter

Bull. Carnegie Mus. Nat. Hist. 6: 73, 7 Nov 1978.
8 • smithsonian contributions to zoology

= Steatomys caurinus Thomas, 1912. See Musser and Carleton 

(2005:943).

Holotype: USNM 467496. Skin and skull. Adult male. Collected 

22 Mar 1969 by L. W. Robbins. Original number 1106.

Type Locality: Yama, [SL and FC: 9°36′N, 6°18′W], Ivory Coast 

(Côte d’Ivoire).

Family Cricetidae G. Fischer, 1817

Genus Abrothrix Waterhouse, 1837

Akodon (Akodon) llanoi Pine

Mammalia 40: 63, 16 Jul 1976.

= Abrothrix olivaceus (Waterhouse, 1837). See Musser and Carleton (2005:1090).

Holotype: USNM 482144. Skin, skull, and skeleton. Adult male. 

Collected 2 May 1971 by J. P. Angle and D. Bridge. Original 

number 187396.

Type Locality: Head of Bahia Capitan Canepa, 54°51′S, 64°27′W, 

Isla de los Estados (Staten Island), Territorio Nacional de la 

Tierra del Fuego, Argentina.

Akodon (Akodon) markhami Pine

An. Inst. Patagon. 4: 423, 1973 (published Aug 1974).

= Abrothrix olivaceus markhami (Pine, 1974). See RodriguesSerrano et al. (2008:309) for current use.

Holotype: USNM 501221. Skin and skull. Adult male. Collected 

2 Apr 1971 by B. J. Markham. Original number 642.

Type Locality: 1.2 km NWW and 1.2 km NNW Puerto Eden, 

49°08′S, 74°27′W, 5–20 m, Isla Wellington, about 600 km 

NNW [SL: 420 km N, 250 km W] Punta Arenas, Magallanes 

[Province, Magallanes y de la Antártica Chilena], Chile.

Chraeomys inornatus Thomas

Smithson. Misc. Collect. 68(4): 2, 10 Apr 1917.

= Abrothrix jelskii (Thomas, 1894). See Musser and Carleton 

(2005:1089).

Holotype: USNM 194685. Skin and skull. [Poole and Schantz 

(1942:260): Adult] female. Collected [SL: 20 Jul 1915] by 

E. Heller. Original number 457.

Type Locality: [SL and FC: Occobamba Pass], altitude 13,000 ft 

(3,962 m), Ollantaytambo, Cuzco (Cusco), Peru.

Genus Akodon Meyen, 1833

Akodon surdus Thomas

Smithson. Misc. Collect. 68(4): 2, 10 Apr 1917.

Holotype: USNM 194663. [Skin and skull]. Adult male. Collected [SL: 28 Oct 1915] by E. Heller. Original number 835.

Type Locality: Huadquina, altitude 5,000 ft (1,524 m), Cuzco 

(Cusco), Peru.

Microxus torques Thomas

Smithson. Misc. Collect. 68(4): 3, 10 Apr 1917.

= Akodon torques (Thomas, 1917). See Thomas (1927:370).

Holotype: USNM 194607. [Skin and skull]. Adult male. Collected 28 May 1915 by E. Heller. Original number 276.

Type Locality: [SL and FC: Cedrobamba], Matchu (Machu) Picchu, altitude 10,000 ft (3,048 m) [SL and FC: 12,000 ft 

(3,658 m)], Cuzco (Cusco), Peru.

Genus Alticola Blanford, 1881

Alticola acmaeus Schwarz

Proc. Zool. Soc. London 108: 665, 26 Jan 1939a.

= Alticola albicaudus (True, 1894). See Rossolimo and Pavlinov 

(1992:160).

Holotype: USNM 198530. Skin and skull. Adult male. Collected 18 Oct 1913 by W. L. Abbott. Original number 

8282.

Type Locality: Mrobuk, [FC: altitude 14,500 ft (4,420 m)], Phyang Nullah, northwest of Leh, Lakak (Ladakh), [Jammu 

and Kashmir, India].

Alticola glacialis Miller

Proc. Biol. Soc. Wash. 26: 197, 23 Oct 1913a.

= Alticola argentatus (Severtzov, 1879). See Rossolimo (1989: 

113), who placed glacialis in A. argentatus; taxon overlooked by Musser and Carleton (2005).

Holotype: USNM 176071. Skin and skull. Adult female. 

Collected 23 Aug 1912 by W. L. Abbott. Original number 

7841.

Type Locality: Chogo Lungma Glacier, [FC: Basha Valley], altitude 11,000 ft (3,253 m), Baltistan [Region, Northern 

Areas, Pakistan].

Alticola roylei parvidens Schlitter and Setzer

Proc. Biol. Soc. Wash. 86: 165, 31 May 1973.

= Alticola argentatus (Severtzov, 1879). See Rossolimo and Pavlinov (1992:164).

Holotype: USNM 413551. Skin and skull. Adult male. Collected 

28 Sep 1966 by C. J. Phillips. Original number 744.

Type Locality: 20.5 mi (32.99 km) north Dir, 10,400 ft (3,170 m), 

Dir State (District), [Malakand Division, North-West Frontier], West Pakistan (Pakistan).

Arvicola albicauda True

Proc. U.S. Natl. Mus. 17: 12, 8 May 1894b.

= Alticola albicaudus (True, 1894). See Hinton (1926:314).

Holotype: USNM 20393/36816. Skin and skull. [Lyon and Osgood (1909:79): Adult] female. Collected 19 Dec 1891 by 

W. L. Abbott. No original number.

Type Locality: Braldu Valley, [SL: 11,000 ft (3,353 m)], Baltistan 

[Northern Areas, Pakistan].

Arvicola montosa True

Proc. U.S. Natl. Mus. 17: 11, 8 May 1894b.

= Alticola montosus (True, 1894). See Hinton (1926:315).
number 642 • 9

Holotype: USNM 20145/35508. Skin and skull. [Lyon and Osgood (1909:88): Half-grown] male. Collected 4 Oct 1891 

by W. L. Abbott. No original number.

Type Locality: Central Kashmir, altitude 11,000 ft (3,353 m), 

Jammu and Kashmir, [India].

Aschizomys lemminus Miller

Proc. Acad. Nat. Sci. Phila. 50: 369, 15 Oct 1898a.

= Alticola lemminus (Miller, 1898). See Ognev (1964:472).

Holotype: USNM 9911/37299. Skin, skull, [and dried body]. 

Adult [MC: male]. Collected by C. W. Baxter, date unknown. [MC: Skin cataloged Jan 1872; skull 29 Apr 1897]. 

Original number [SL: 423]. Miller (1898a:370) states that 

the skin was in alcohol for many years.

Type Locality: Kelsey Station, Plover Bay, Bering Strait, [Chukotskiy], Siberia (Russia).

Microtus acrophilus Miller

Proc. Acad. Nat. Sci. Phila. 51: 296, 9 Aug 1899a.

= Alticola stoliczkanus (Blanford, 1875). See Schwarz (1939a: 

663).

Holotype: USNM 62162. Skin and skull. Adult female. Collected 

25 Jul 1893 by W. L. Abbott. No original number.

Type Locality: On the Ladak (Ladakh) side of the Kara Korum 

Pass, altitude 17,000 ft (5,182 m), [Jammu and Kashmir, 

India].

Microtus cricetulus Miller

Proc. Acad. Nat. Sci. Phila. 51: 294, 9 Aug 1899a.

= Alticola stoliczkanus (Blanford, 1875). See Schwarz (1939a: 

663).

Holotype: USNM 84043. Skin and skull. Adult male. Collected 

11 Aug 1897 by W. L. Abbott. No original number.

Type Locality: Banks of the Tso Kyun, altitude 16,000 ft 

(4,877 m), [SL: Rupshu], Ladak (Ladakh) [District, Jammu 

and Kashmir, India].

Genus Arborimus Taylor, 1915

Phenacomys albipes Merriam

Proc. Biol. Soc. Wash. 14: 125, 19 Jul 1901a.

= Arborimus albipes (Merriam, 1901). See Johnson and Maser 

(1982:17).

Holotype: USNM 97236. Skin and skull. Adult male. Collected 

24 May 1899 by W. K. Fisher. Original number 821.

Type Locality: Redwoods near Arcata, Humboldt Bay, [Humboldt County], California.

Phenacomys longicaudus True

Proc. U.S. Natl. Mus. 13: 303, 15 Nov 1890.

= Arborimus longicaudus longicaudus (True, 1890). See Johnson (1968:27); trinomial recognized by Hall (1981:788) 

under Phenacomys and by Carey (1999:622) under 

Arborimus.

Holotype: USNM 19071/34808, not USNM 19071/25883 as in 

original description. Skin and skull. Collected [MC: Aug] 

1890 by A. Todd. No original number.

Type Locality: Marshfield, Coos County, Oregon.

Genus Baiomys True, 1894

Baiomys musculus grisescens Goldman

Proc. Biol. Soc. Wash. 45: 121, 30 Jul 1932a.

Subspecies recognized by Hall (1981:723).

Holotype: USNM 257083. [Skin and skull]. Adult female. Collected 6 Mar 1932 by C. F. Underwood. Original number 838.

Type Locality: Comayabuela (Comayagüela), altitude 3,100 

ft (945 m), just south of Tegucigalpa [Miller and Kellogg 

(1955:513): on Rio Grande opposite Tegucigalpa, Fancisco 

Morazin], Honduras.

Baiomys musculus handleyi Packard

Univ. Kans. Publ. Mus. Nat. Hist. 9: 399, 19 Dec 1958a.

Subspecies recognized by Hall (1981:723).

Holotype: USNM 275604. Skin and skull. Adult female. Collected 

24 Apr 1947 by C. O. Handley Jr. Original number 991.

Type Locality: [FC: Rio Blanco, 6.9 mi (11 km) W] Sacapulus, El 

Quiche (Quiché), Guatemala.

Peromyscus allex Osgood

Proc. Biol. Soc. Wash. 17: 76, 21 Mar 1904a.

= Baiomys taylori allex (Osgood, 1904). See Packard (1958b:17); 

subspecies recognized by Hall (1981:725) and Cameron 

(1999a:587).

Holotype: USNM 33429/45452. Skin and skull. Adult male [SL 

and FC: female]. Collected 7 Mar 1892 by E. W. Nelson. 

Original number [SL: 2029].

Type Locality: Colima, [FC: altitude 1,700 ft (518 m)], Colima, 

Mexico.

Peromyscus musculus nigrescens Osgood

Proc. Biol. Soc. Wash. 17: 76, 21 Mar 1904a.

= Baiomys musculus nigrescens (Osgood, 1904). See Miller 

(1912a:137); subspecies recognized by Hall (1981:724).

Holotype: USNM 76827. [Skin and skull]. Adult female. Collected 9 Dec 1895 by E. W. Nelson and E. A. Goldman. 

Original number [SL: 8719].

Type Locality: Valley of Comitan, [FC: altitude 5,700 ft 

(1,737 m)], Chiapas, Mexico.

Peromyscus taylori analogus Osgood

N. Am. Fauna 28: 256, 17 Apr 1909a.

= Baiomys taylori analogus (Osgood, 1909). See Miller 

(1912a:137); subspecies recognized by Hall (1981:725) and 

Cameron (1999a:587).

Holotype: USNM 120261. [Skin and skull]. Adult male. Collected 15 Jan 1903 by E. W. Nelson and E. A. Goldman. 

Original number [SL: 15764].

Type Locality: Zamora, Michoacan, [SL: Mexico].
1 0 • smithsonian contributions to zoology

Peromyscus taylori subater V. Bailey

N. Am. Fauna 25: 102, 24 Oct 1905.

= Baiomys taylori subater (V. Bailey, 1905). See Miller 

(1912a:137); subspecies recognized by Hall (1981:726) and 

Cameron (1999a:587).

Holotype: USNM 32616/44539. Skin and skull. Adult female. Collected 25 Feb 1892 by W. Lloyd. Original number 1122.

Type Locality: Bernard Creek, near [FC: 12 mi (19 km) west of] 

Columbia, Brazoria County, Texas.

Sitomys musculus Merriam

Proc. Biol. Soc. Wash. 7: 170, 29 Sep 1892a.

= Baiomys musculus musculus (Merriam, 1892). See Mearns 

(1907:381); subspecies recognized by Hall (1981:724).

Holotype: USNM 33437/45460. Skin and skull. Adult female 

[SL and FC: male]. Collected 9 Mar 1892 by E. W. Nelson. 

Original number 2055.

Type Locality: Near Colima City, [FC: altitude 1,700 ft (518 m)], 

[SL: Colima], Mexico.

Genus Bibimys Massoia, 1979

Akodon chacoensis Shamel

J. Wash. Acad. Sci. 21: 427, 19 Oct 1931.

= Bibimys chacoensis (Shamel, 1931). See Massoia (1980:280).

Holotype: USNM 236239. Skin and skull. Adult male. Collected 

20 Jul 1920 by A. Wetmore. Original number [SL: 1059].

Type Locality: [FC: Rio Quia], Las Palmas, Chaco, Argentina.

Genus Calomys Waterhouse, 1837

Hesperomys frida Thomas

Smithson. Misc. Collect. 68(4): 1, 10 Apr 1917.

= Calomys sorellus (Thomas, 1900). See Hershkovitz (1962:137).

Holotype: USNM 194779. [Skin and skull]. Female. Collected 

[SL and FC: 14 Apr 1915] by E. Heller. Original number 96.

Type Locality: [FC: Huarocondo River], Chospyoc, altitude 

10,000 ft (3,048 m), [Cusco], Peru.

Genus Chelemys Thomas, 1903

Notiomys vestitus alleni Osgood

Field Mus. Nat. Hist., Zool. Ser. 12: 124, 26 Oct 1925.

= Chelemys macronyx (Thomas, 1894). See Reig (1987:363).

Holotype: USNM 84227. [Skin and skull]. Adult male. Collected 

8 Feb 1897 by O. A. Peterson. Original number 372.

Type Locality: Upper Rio Chico, [SL: Cordilleros], Santa Cruz, 

southern Argentina.

Genus Dicrostonyx Gloger, 1841

Dicrostonyx groenlandicus clarus Handley

J. Wash. Acad. Sci. 43: 197, 24 Jun 1953.

Subspecies recognized by Hall (1981:835) and Nagy and Gower 

(1999:660).

Holotype: USNM 290952. Skin and skull. Old adult male. Collected 16 Jun 1949 by C. O. Handley Jr. Original number 

1285.

Type Locality: Near sea level at Cherie Bay, 5.4 mi (9 km) ENE 

of Mould Bay Station, Prince Patrick Island, District of 

Franklin, Northwest Territories, Canada, Lat. 76°19′N, 

Long. 119°02′W.

Dicrostonyx nelsoni Merriam

Proc. Wash. Acad. Sci. 2: 25, 14 Mar 1900a.

= Dicrostonyx nelsoni nelsoni (Merriam, 1900). See Honacki et 

al. (1982:483); subspecies recognized under D. groenlandicus by Hall (1981:836) and Nagy and Gower (1999:660).

Holotype: USNM 186500. Skin and skull, originally an alcoholic. Young adult female. Collected on an unknown date 

by E. W. Nelson. [MC: Received Aug 1879, cataloged 4 

May 1885]. Original number [SL: 448].

Type Locality: St. Michaels (Saint Michael), [Norton Sound], 

Alaska.

Remarks: This specimen, originally cataloged as USNM 14965, 

was exchanged to the C. H. Merriam collection and became 

5488/6163 thereof, the latter of which Merriam cited as the 

type. It returned to the USNM when the Merriam collection 

was donated and received its current museum number on 

20 Oct 1911.

Dicrostonyx richardsoni Merriam

Proc. Wash. Acad. Sci. 2: 26, 14 Mar 1900a.

Holotype: USNM 186501. [In alcohol, skull removed]. Adult 

male. Collected Jul 1859 by W. MacTavish. No original 

number.

Type Locality: Fort Churchill, [Manitoba, Canada].

Remarks: This specimen, originally cataloged as USNM 7756, 

was exchanged to the C. H. Merriam collection and became 

5489/6164, the latter of which Merriam cited as the type. 

It returned to the USNM when the Merriam collection was 

donated and received its current museum number on 20 Oct 

1911.

Dicrostonyx unalascensis Merriam

Proc. Wash. Acad. Sci. 2: 25, 14 Mar 1900a.

= Dicrostonyx unalascensis unalascensis Merriam, 1900. See Nelson (1929:145); subspecies recognized by Hall (1981:837) 

and Nagy and Gower (1999:660) under Dicrostonyx groenlandicus.

Holotype: USNM 99622. [Skull with some hair from pelt from 

owl pellets]. Adult. Collected 8 Jul 1899 by C. H. Merriam. 

No original number.

Type Locality: Unalaska [Island, Aleutian Islands], Alaska.

Dicrostonyx unalascensis peninsulae Handley

J. Wash. Acad. Sci. 43: 199, 24 Jun 1953.
number 642 • 1 1

= Dicrostonyx nelsoni peninsulae Handley, 1953. See Honacki 

et al. (1982:483); subspecies recognized by Hall (1981:836) 

and Nagy and Gower (1999:660) under Dicrostonyx groenlandicus.

Holotype: USNM 246377. Skin and skull. Old adult female. Collected 8 May 1925 by O. J. Murie. Original number 1993.

Type Locality: Near sea level at Urilla Bay, Unimak Island, [Aleutian Islands], Alaska.

Dicrostonyx unalascensis stevensoni Nelson

Proc. Biol. Soc. Wash. 42: 145, 30 Mar 1929.

Subspecies recognized by Hall (1981:837) under Dicrostonyx 

groenlandicus.

Holotype: USNM 235552. [Skin and skull]. Adult female. Collected 14 Dec 1920 by D. H. Stevenson. Original number 

[SL: 9, 22183-X].

Type Locality: Umnak Island, [Aleutian Islands], Alaska.

Genus Euneomys Coues, 1874

Euneomys noei Mann

Biologica (Santiago, Chile) 1: 95, Sep 1944.

= Euneomys mordax Thomas, 1912. See Pearson and Christie 

(1991:126).

Holotype: USNM 391817. [Skin, skull, and partial skeleton]. 

Subadult male. Collected 29 Feb 1944 by G. Mann. Original number 179.

Type Locality: Valle de la Junta [SL: La Junta (Lo Valdes)], Cajon 

del Rio Volcan, 2,400 m [SL: 2,500 m], [Chile].

Remarks: Published as G. Mann number 179; cataloged as 

USNM 391817 on 8 Dec 1967.

Euneomys petersoni J. A. Allen

Bull. Am. Mus. Nat. Hist. 19: 192, 9 May 1903.

Holotype: USNM 84198. [Skin and skull]. Adult female. Collected 

10 Feb 1897 by O. A. Peterson. Original number [SL: 390].

Type Locality: Upper Rio Chico de Santa Cruz, near the Cordilleras, Patagonia (Santa Cruz, Argentina).

Genus Geoxus Thomas, 1919

Oxymycterus microtus J. A. Allen

Bull. Am. Mus. Nat. Hist. 19: 189, 9 May 1903.

= Geoxus valdivianus (Philippi, 1858). See Musser and Carleton 

(2005:1116).

Holotype: USNM 84234. [Skin and skull]. Adult male. Collected 

7 Mar 1897 by O. A. Peterson. Original number [SL: 570].

Type Locality: Pacific slope of the Cordilleras near the head of 

the Rio Chico de Santa Cruz, [Santa Cruz, Argentina].

Genus Habromys Hooper and Musser, 1964

Peromyscus lepturus Merriam

Proc. Biol. Soc. Wash. 12: 118, 30 Apr 1898a.

= Habromys lepturus (Merriam, 1898). See Honacki et al. (1982: 

419).

Holotype: USNM 68612. [Skin and skull]. Adult male. Collected 

8 Jul 1894 by E. W. Nelson and E. A. Goldman. Original 

number 6381.

Type Locality: Mount Zempoaltepec, altitude 8,200 ft (2,499 m), 

Oaxaca, Mexico.

Peromyscus lophurus Osgood

Proc. Biol. Soc. Wash. 17: 72, 21 Mar 1904a.

= Habromys lophurus (Osgood, 1904). See Honacki et al. (1982:419).

Holotype: USNM 77219. [Skin and skull]. Adult male. Collected 

30 Dec 1895 by E. W. Nelson and E. A. Goldman. Original 

number [SL: 8956].

Type Locality: Todos Santos [FC: altitude 10,000 ft (3,048 m)], 

[Huehuetenango], Guatemala.

Peromyscus simulatus Osgood

Proc. Biol. Soc. Wash. 17: 72, 21 Mar 1904a.

= Habromys simulatus (Osgood, 1904). See Honacki et al. 

(1982:419).

Holotype: USNM 55028. [Skin and skull]. [Lyon and Osgood 

(1909:137): Adult] [FC: female]. Collected 12 Jul 1893 by 

E. W. Nelson and E. A. Goldman. Original number [SL: 5224].

Type Locality: [Poole and Schantz (1942:331): Near] Jico, altitude 6,000 ft (1,829 m), Veracruz, Mexico.

Genus Hodomys Merriam, 1894

Hodomys alleni guerrerensis Goldman

J. Wash. Acad. Sci. 28: 498, 15 Nov 1938a.

Subspecies recognized under Neotoma (Hodomys) alleni by Hall 

(1981:772).

Holotype: USNM 70574. Skin and skull. Adult male. Collected 

6 Jan 1895 by [E. W.] Nelson and [E. A.] Goldman. Original 

number 7321.

Type Locality: Acapulco, [SL and FC: near] sea level, Guerrero, 

Mexico.

Hodomys vetulus Merriam

Proc. Acad. Nat. Sci. Phila. 46: 236, 24 Sep 1894a.

= Hodomys alleni vetulus (Merriam, 1894). New name combination; subspecies recognized under Neotoma (Hodomys) 

alleni by Hall (1981:772).

Holotype: USNM 53656. [Skin and skull]. Adult male. Collected 

8 May 1893 by E. W. Nelson. Original number 4784.

Type Locality: Tehuacan, [FC: altitude 5,400 ft (1,646 m)], 

Puebla, Mexico.

Neotoma alleni Merriam

Proc. Biol. Soc. Wash. 7: 168, 29 Sep 1892a.

= Hodomys alleni alleni (Merriam, 1892). New name combination; subspecies recognized under Neotoma (Hodomys) alleni by Hall (1981:772).
1 2 • smithsonian contributions to zoology

Holotype: USNM 32709/44632. Skin and skull. Adult male. Collected 26 Jan 1892 by E. W. Nelson. Original number 1796.

Type Locality: Manzanillo, [FC: sea level up to 50 ft (15 m)], 

Colima, Mexico.

Genus Holochilus Brandt, 1835

Holochilus incarum Thomas

Proc. U.S. Natl. Mus. 58: 226, 10 Nov 1920a.

= Holochilus sciureus Wagner, 1842. See Massoia (1980:286). 

Holotype: USNM 194915. [Skin and skull]. Immature female. 

Collected 22 Dec 1914 by E. C. Erdis. Original number 22. 

Also E. Heller number 581.

Type Locality: Santa Ana, altitude 3,480 ft (1,061 m), Cuzco 

(Cusco), Peru.

Genus Hyperacrius Miller, 1896

Arvicola fertilis True

Proc. U.S. Natl. Mus. 17: 10, 8 May 1894b.

= Hyperacrius fertilis (True, 1894). See Hinton (1926:324).

Holotype: USNM 20147/35510. Skin and skull. [Lyon and Osgood (1909:85): Adult] female. Collected 30 Aug 1891 by 

W. L. Abbott. No original number.

Type Locality: Pir Panjal Range, altitude 8,500 ft (2,591 m), 

[Jammu and Kashmir], Kashmir (India).

Hyperacrius fertilis zygomaticus Phillips

J. Mammal. 50: 469, 22 Aug 1969.

= Hyperacrius fertilis (True, 1894).

Holotype: USNM 413574. Skin and skull. Adult female. Collected 7 Oct 1966 by C. J. Phillips. Original numbers 847; 

RT-B67290 [SL and FC: FF].

Type Locality: 6 mi (10 km) SW Utror, 8,900 ft (2,713 m), Swat 

[District, Malakand Division, North-West Frontier], West 

Pakistan (Pakistan).

Hyperacrius wynnei traubi Phillips

J. Mammal. 50: 463, 22 Aug 1969.

= Hyperacrius wynnei (Blanford, 1881).

Holotype: USNM 413577. Skin and skull. Adult male [SL and 

FC: both say female]. Collected 13 Oct 1966 by M. Iqbal. 

Original numbers 422; RT-B67331 [SL and FC: BB].

Type Locality: Yakh Tangai, 6,600 ft (2,012 m) [SL and FC: both 

say 6,900 ft (2,103 m)], Swat [District, Malakand Division, 

North-West Frontier], West Pakistan (Pakistan).

Microtus brachelix Miller

Proc. Acad. Nat. Sci. Phila. 51: 290, 9 Aug 1899a.

= Hyperacrius fertilis (True, 1894). See Hinton (1926:335).

Holotype: USNM 63445. Skin and skull. Young adult female. 

Collected 15 Nov 1895 by W. L. Abbott. No original number.

Type Locality: Nagmarg, altitude 9,000 ft (2,743 m), [Jammu 

and Kashmir], Kashmir (India).

Genus Ichthyomys Thomas, 1893

Ichthyomys pittieri Handley and Mondolfi

Acta Biol. Venezuelica 3: 417, 26 Nov 1963.

Holotype: USNM 342987. Skin and skull. Adult male. Collected 

20 Apr 1958 by B. Terzenbach and G. Medina. No original 

number.

Type Locality: Near the head of the Rio Limon, Parque Nacional 

de Rancho Grande, Estado Aragua, Venezuela.

Genus Isthmomys Hooper and Musser, 1964

Peromyscus pirrensis Goldman

Smithson. Misc. Collect. 60(2): 5, 20 Sep 1912a.

= Isthmomys pirrensis (Goldman, 1912). See Carleton (1980:124).

Holotype: USNM 178997. [Skin and skull]. Adult male. Collected 3 May 1912 by E. A. Goldman. Original number 

21675.

Type Locality: Near head of Rio Limon, altitude 4,500 ft 

(1,372 m), Mount Pirri, [Darien], eastern Panama.

Genus Lasiopodomys Lataste, 1887

Microtus pullus Miller

Proc. Biol. Soc. Wash. 24: 53, 24 Feb 1911a.

= Lasiopodomys mandarinus (Milne-Edwards, 1871). See 

Musser and Carleton (1993:516).

Holotype: USNM 155047. Skin and skull. Adult male. Collected 

11 Oct 1908 by A. de C. Sowerby. Original number 32.

Type Locality: Chiao Ch’eng Shan, altitude 7,000 ft (2,134 m), 

90 mi (145 km) west of Tai Yuan Fu, Shansi (Shanxi), China.

Microtus warringtoni Miller

Smithson. Misc. Collect. 60(28): 1, 31 Mar 1913b.

= Lasiopodomys brandtii Radde, 1861. See Musser and Carleton 

(1993:515).

Holotype: USNM 175861. Skin and skull. Adult male. Collected 

13 Aug 1912 by A. de C Sowerby. Original number 508.

Type Locality: Tabool, altitude 4,000 ft (1,219 m), Mongolia, 

100 mi (161 km) north of Kalgan, [Nei Monggol, China].

Genus Lemmiscus Thomas, 1912

Subspecies of Lemmiscus curtatus were not recognized by 

Musser and Carleton (2005:896). We follow Hall (1981:821–

822), who listed them under Lagurus curtatus, and Smith 

(1999:650).

Arvicola curtata Cope

Proc. Acad. Nat. Sci. Phila. 20: 2, presented at meeting of 21 

Jan 1868. Received prior to Jun 1868 by the Entomological 

Society of Philadelphia.

= Lemmiscus curtatus curtatus (Cope, 1868). See Goldman 

(1941a:70).
number 642 • 1 3

Holotype: USNM 10267/37419. [Fragments of skull and feet. 

Age and sex unknown]. Collected on unknown date by 

W. M. Gabb. [MC: Skin cataloged 15 Oct 1872; skull 5 

May 1898].

Type Locality: Pigeon springs, on the eastern boundary line of California east of Owens Valley, [Miller and Rehn (1901:127): 

Mount Magruder, Nevada, near boundary between Inyo 

County, California, and Esmeralda County, Nevada].

Remarks: Type not designated by number in original description. 

Coues (1877:215–216) borrowed the type from Cope and 

described its poor condition similar to that of 10267. The 

museum catalog entry for 10267 is marked “Copes type” 

and a Coues’s Monograph of North American Muridae

label associated with the specimen is marked “No. 10267, 

Type of Arvicola curtatus Cope.”

Arvicola (Chilotus) pallidus Merriam

Am. Nat. 22: 704, Aug 1888a.

= Lemmiscus curtatus pallidus (Merriam, 1888). See Goldman 

(1941a:70).

Holotype: USNM 186498. Skin and skull. Adult female. Collected 10 Sep 1887 by V. Bailey. Original number [SL: 226]. 

Merriam collection number 3852/4431.

Type Locality: Fort Buford, [Williams County, North] Dakota.

Remarks: Merriam designated the type as his collection number 

3852/4431. The Merriam collection was later donated to 

the USNM, and USNM 186498 was cataloged on 20 Oct 

1911.

Arvicola pauperrima Cooper

Am. Nat. 2: 535, Dec 1868.

= Lemmiscus curtatus pauperrimus (Cooper, 1868). See Goldman (1941a:70).

Holotype: USNM 10268/35071. [Skin and skull. Lyon and Osgood (1909:90): Adult. SL: Male]. Collected 9 Oct 1860 by 

J. G. Cooper. Original number 126.

Type Locality: Great Plain of Columbia, near Snake River, [SL: 

WT] (Washington).

Remarks: Type designated as Cooper’s number 126. Skin cataloged as USNM 10268 on 15 Oct 1872; skull 5 Mar 1891.

Lemmiscus curtatus levidensis Goldman

Proc. Biol. Soc. Wash. 54: 70, 31 Jul 1941a.

Holotype: USNM 148168. Skin and skull. Adult female. Collected 6 Aug 1906 by M. Cary. Original number 733.

Type Locality: Sand Hills, 5 mi (8 km) east of Canadian River [SL 

and FC: Creek] at west base of Medicine Bow Range, east 

of Weldon, North Park, altitude about 8,000 ft (2,438 m), 

Jackson County, Colorado.

Genus Lemmus Link, 1795

Subspecies of Lemmus trimucronatus were not recognized 

by Musser and Carleton (2005:988). We follow Hall (1981: 

828–830) and Batzli (1999a:654) for the trinomial epithets; 

however, they listed them under Lemmus sibiricus.

Lemmus alascensis Merriam

Proc. Wash. Acad. Sci. 2: 26, 14 Mar 1900a.

= Lemmus trimucronatus alascensis Merriam, 1900. See Anderson (1937:110).

Holotype: USNM 186499. In alcohol, [skull removed]. Adult female. Collected [MC: 11 Jun 1883 by J. Murdock. Original 

number 102]. Merriam collection number 5491/6166.

Type Locality: Point Barrow, Alaska.

Remarks: Published as Merriam collection number 5491/6166; 

cataloged as USNM 186499 on 20 Oct 1911.

Lemmus minusculus Osgood

N. Am. Fauna 24: 36, 23 Nov 1904b.

= Lemmus trimucronatus minusculus Osgood, 1904. See Davis 

(1944a:23).

Holotype: USNM 119612. [Skin and skull]. Adult male. Collected 1 Sep 1902 by W. H. Osgood and A. G. Maddren. 

Original number [FC: WHO 1903].

Type Locality: Kakhtul River, near its junction with the Malchatna (Mulchatna) River, Alaska.

Lemmus trimucronatus subarcticus Bee and Hall

Univ. Kans. Mus. Nat. Hist. Misc. Publ. 8: 109, 10 Mar 1956.

Holotype: USNM 303204. Skin and skull. Adult male. Collected 

24 Jul 1952 by J. W. Bee. Original number 520724-9. University of Kansas Museum of Natural History number 50876.

Type Locality: Lake Schroder, altitude 2,900 ft (884 m), Brooks 

Range, Alaska, 69°24′28′′[N], 145°09′50′′[W].

Remarks: Published as University of Kansas Museum of Natural 

History number 50876; cataloged as USNM 303204 on 26 

Jun 1956.

Lemmus yukonensis Merriam

Proc. Wash. Acad. Sci. 2: 27, 14 Mar 1900a.

= Lemmus trimucronatus yukonensis Merriam, 1900. See Bee 

and Hall (1956:113).

Holotype: USNM 98849. [Skin and skull]. [Lyon and Osgood 

(1909:77): Young] adult female. Collected 9 Aug 1899 by 

W. H. Osgood. Original number 769.

Type Locality: [FC: Near] Charlie Creek, Yukon River, Alaska.

Myodes nigripes True

Proc. U.S. Natl. Mus. 17: 2, 24 Apr 1894c. Preprint of Proc. U.S. 

Natl. Mus. 17: 242, 15 Nov 1894d.

= Lemmus trimucronatus nigripes (True, 1894). New name combination. Rausch (1953:127) relegated nigripes to subspecific status under L. sibiricus.

Holotype: USNM 59152. [Skin and skull]. [Lyon and Osgood 

(1909:76): Adult] male. Collected 18 Aug [probably 1883 

according to accession records] by C. H. Townsend. No 

original number.
1 4 • smithsonian contributions to zoology

Type Locality: St. George’s (Saint George) Island, [Pribilof Islands], Alaska.

Genus Lophiomys Milne-Edwards, 1867

Lophiomys thomasi Heller

Smithson. Misc. Collect. 59(16): 4, 5 Jul 1912.

= Lophiomys imhausi Milne-Edwards, 1867. See Hayman and 

Holt (1940:636).

Holotype: USNM 181789. [Skin and skull]. Adult male. Collected 1 Sep 1911 by E. Heller. Original number 4206.

Type Locality: [SL: North Creek], Mount Gargues (Uaragess), 

altitude 6,000 ft (1,829 m), Mathews Range, British East 

Africa (Kenya).

Genus Megadontomys Merriam, 1898

Peromyscus (Megadontomys) nelsoni Merriam

Proc. Biol. Soc. Wash. 12: 116, 30 Apr 1898a.

= Megadontomys nelsoni (Merriam, 1898). See Carleton (1989:37).

Holotype: USNM 55024. [Skin and skull]. Adult female. Collected 10 Jul 1893 by E. W. Nelson. Original number 5202.

Type Locality: Jico, altitude 6,000 ft (1,829 m), Veracruz, Mexico.

Peromyscus (Megadontomys) thomasi Merriam

Proc. Biol. Soc. Wash. 12: 116, 30 Apr 1898a.

= Megadontomys thomasi (Merriam, 1898). See Carleton (1980: 

129).

Holotype: USNM 70142. [Skin and skull]. Old male. Collected 

24 Dec 1894 by E. W. Nelson and E. A. Goldman. Original 

number 7250.

Type Locality: Mountains near Chilpancingo, altitude 9,700 ft 

(2,757 m), Guerrero, Mexico.

Genus Melanomys Thomas, 1902

Oryzomys idoneus Goldman

Smithson. Misc. Collect. 56(36): 5, 19 Feb 1912b.

= Melanomys caliginosus (Tomes, 1860). See Musser and Carleton (1993:708).

Holotype: USNM 171106. [Skin and skull]. Adult female. Collected 26 Mar 1911 by E. A. Goldman. Original number 

[FC: Nelson and Goldman] 21010.

Type Locality: Cerro Azul, altitude 2,500 ft (762 m), near the 

headwaters of the Chagres River, [Panama], Panama.

Genus Microryzomys Thomas, 1917

Oryzomys (Microryzomys) aurillus Thomas

Smithson. Misc. Collect. 68(4): 1, 10 Apr 1917.

= Microryzomys minutus (Tomes, 1860). See Carleton and 

Musser (1989:65).

Holotype: USNM 194795. [Skin and skull]. Female. Collected 

[SL: 15 May] 1915 by E. Heller. Original number 255.

Type Locality: Torontoy, [SL: altitude 14,000 ft (4,267 m)], 

[Cusco], Peru.

Genus Microtus Schrank, 1798

Arvicola (Mynomes) alticolus Merriam

N. Am. Fauna 3: 67, 11 Sep 1890a.

= Microtus longicaudus alticola (Merriam, 1890). See Goldman 

(1938b:491). Subspecies recognized by Hall (1981:807) and 

Smolen (1999a:632).

Holotype: USNM 17615/24551. Skin and skull. Adult female. 

Collected 31 Jul 1889 by C. H. Merriam and V. Bailey. 

Original number [FC: VB] 243.

Type Locality: Little Spring, San Francisco Mountain, altitude 

8,200 ft (2,500 m), [Coconino County], Arizona.

Arvicola apella LeConte

Proc. Acad. Nat. Sci. Phila. 6: 405, reported favorably for publication 25 Oct 1853. Receipt acknowledged 31 Dec 1853 by 

the Smithsonian Institution.

= Microtus pinetorum scalopsoides (Audubon and Bachman, 

1841). See V. Bailey (1900:64); subspecies recognized by 

Hall (1981:817) and Smolen (1999b:644).

Holotype: USNM 4714. [Skin with skull inside. Age, sex, and 

date collected unknown]. [MC: From J. E. LeConte’s collection. Cataloged 13 Apr 1861]. No original number.

Type Locality: Pennsylvania.

Remarks: Type not designated in original description. Coues 

(1877:223–224) refers to 4714 as the type of Arvicola 

apella LeConte. This specimen also bears a Coues label 

that is marked type of Arvicola apella LeConte. There is no 

reason to believe that LeConte had more than one specimen 

when he described Arvicola paella; therefore, USNM 4714 

is the holotype fixed by monotypy (ICZN, 1999: Article 

73.1.2).

Arvicola austerus LeConte

Proc. Acad. Nat. Sci. Phila. 6: 405, reported favorably for publication 25 Oct 1853. Receipt acknowledged 31 Dec 1853 by 

the Smithsonian Institution.

= Microtus ochrogaster ochrogaster (Wagner, 1842). See Hall 

and Kelson (1959:749); subspecies recognized by Hall 

(1981:813) and Stalling (1999:638).

Holotype: USNM 2249. [Skin only]. [MC: Collected by P. R. 

Hoy, date unknown. Cataloged 28 Feb 1857]. Original 

number [SL: 950].

Type Locality: [SL: Racine], [Racine County], Wisconsin.

Remarks: Type not designated by number in original description. 

It is entered in the museum catalog as the “type of Maj. 

LeConte.” Coues (1877:210, 214) refers to 2249 as the 

type of Arvicola austerus. There is no reason to believe that 

LeConte had more than one specimen when he described 

Arvicola paella; therefore, USNM 2249 is the holotype fixed 

by monotypy (ICZN, 1999: Article 73.1.2).
number 642 • 1 5

Arvicola austerus minor Merriam

Am. Nat. 22: 600, Jul 1888b.

= Microtus ochrogaster minor (Merriam, 1888). See Bole and 

Moulthrop (1942:160); subspecies recognized by Hall 

(1981:813) and Stalling (1999:638).

Holotype: USNM 186493. Skin and skull. Male. Collected 27 

Aug 1887 by [SL: V. Bailey]. Original number [SL: 210]. 

Merriam collection number 3827/4406.

Type Locality: Bottineau, [V. Bailey (1900:75): at base of] Turtle 

Mountain, [Bottineau County, North] Dakota.

Remarks: Published as Merriam collection number 3827/4406; 

cataloged as USNM 186493 on 19 Oct 1911.

Arvicola breweri Baird

Mammals Rep. Explor. Surv. 8(1): 525, 1857.

= Microtus breweri (Baird, 1857). See Miller (1896a:83).

Syntypes: Collected Jul 1856 by T. M. Brewer. No original 

numbers.

USNM 2828. [Lyon and Osgood (1909:82): Adult] male. 

Alcoholic.

USNM 2829. [Lyon and Osgood (1909:82): Adult] male. Alcoholic. Cannot be found.

USNM 2830/3213. [Lyon and Osgood (1909:82): Adult] female. 

Skin and skull.

USNM 2831/3214. [Lyon and Osgood (1909:82): Adult] male. 

Skin and skull.

USNM 2832/3215. Latter number incorrectly given as 3220 in 

original list. [Lyon and Osgood (1909:82): Immature] male. 

Skin and skull.

USNM 2833. Alcoholic. Currently number 4365 in the Museum of Comparative Zoology (Helgen and McFadden, 

2001:114).

Type Locality: Muskeeget (Muskeget) Island, [Nantucket County], 

Massachusetts.

Remarks: No type was designated in the original description. 

Specimens listed above appear in Baird’s (1857:526) list of 

specimens and are considered syntypes as there is no reason 

to believe any one in particular should be considered a holotype, nor is there a reason to designate a lectotype for a taxon 

that occurs on an island where no other Microtus occur.

Arvicola californica Peale

U.S. Explor. Exped. 8: 46, 1848.

= Microtus californicus californicus (Peale, 1848). See Miller 

(1912a:218); subspecies recognized by Hall (1981:801) and 

Heske and Lidicker (1999:627).

Holotype: USNM 10082/4156. [Skin and skull]. Collected [Kellogg (1918:9): 26 Oct 1841 by T. R. Peale]. No original 

number.

Type Locality: San Francisco Bay, California. Restricted by Kellogg (1918:9) to probably at San Francisquito Creek, near 

Palo Alto, [Santa Clara County], California.

Remarks: Type not designated by number in original description. Baird (1857:534) stated he had the original specimen, 

although he gave no museum number. The skin was cataloged in 1872, and the entry states “type of californicus.” 

One old skin label is marked Arvicola californica Peale, 

and the other, one of Coues’s Monographs of American 

Muridae labels, is marked “Type of Arvicola californica.” 

Coues (1877:173) lists 10082 as the type of “californicus.” 

There is no reason to believe that Peale had more than one 

specimen when he described Arvicola californica; therefore, 

USNM 10082/4156 is the holotype fixed by monotypy 

(ICZN, 1999: Article 73.1.2). V. Bailey (1900:35) believed 

that the skull to this specimen is either abnormal or came 

from a different specimen. See Kellogg (1918:9) for additional thoughts on this subject.

Arvicola (Pedomys) cinnamomea Baird

Mammals Rep. Explor. Surv. 8(1): 541, 1857.

= Microtus ochrogaster ochrogaster (Wagner, 1842). See Hall 

and Kelson (1959:749); subspecies recognized by Hall 

(1981:813) and Stalling (1999:638).

Holotype: USNM 591/1714. Skin [MC: misplaced before 1890 

and has not been seen since] and skull. [Lyon and Osgood 

(1909:83): Adult male]. Date collected unknown, [MC: 

cataloged 19 Mar 1855]. Collected by C. Cavileer. Original 

number [MC: 23].

Type Locality: Pembina, [Pembina County], Minnesota (North 

Dakota). V. Bailey (1900:74) stated he could not believe 

the specimen came from Pembina and was probably from 

somewhere within the range of Microtus ochrogaster 

ochrogaster.

Arvicola edax LeConte

Proc. Acad. Nat. Sci. Phila. 6: 405, reported favorably for publication 25 Oct 1853. Receipt acknowledged 31 Dec 1853 by 

the Smithsonian Institution.

= Microtus californicus californicus (Peale, 1848). See Kellogg 

(1918:5); subspecies recognized by Hall (1981:801) and 

Heske and Lidicker (1999:627).

Holotype: USNM 4721/37298. [Skin and skull]. Collected by 

[SL: Major] J. L. LeConte, date unknown. Skin cataloged 13 

Apr 1861; skull 27 Apr 1897. No original number.

Type Locality: [Baird (1857:532): South of San Francisco], [Kellogg (1918:5): probably Monterey], California.

Remarks: Type not designated by number in original description. In his list of specimens, Baird (1857:532) lists only one 

specimen from California and collected by LeConte and earlier states that his description is based on “the original of the 

species as described by Major LeConte.” Coues (1877:173) 

lists 4721 as the type of edax. There is no reason to believe 

that LeConte had more than one specimen when he described Arvicola edax; therefore, USNM 4721/37298 is the 

holotype fixed by monotypy (ICZN, 1999: Article 73.1.2).

Arvicola (Pedomys) haydeni Baird

Mammals Rep. Explor. Surv. 8(1): 543, 1857.
1 6 • smithsonian contributions to zoology

= Microtus ochrogaster haydeni (Baird, 1857). See Osgood 

(1907a:48); subspecies recognized by Hall (1981:812) and 

Stalling (1999:638).

Holotype: USNM 699/1862. Skin and skull. [Lyon and Osgood 

(1909:86): Adult] male. Collected in 1854 by F. V. Hayden. 

Original number [MC: 74].

Type Locality: Fort Pierre, Nebraska (Stanley County, South 

Dakota).

Arvicola kennicottii Baird

Mammals Rep. Explor. Surv. 8(1): 547, 1857.

= Microtus pinetorum scalopsoides (Audubon and Bachman, 

1841). See V. Bailey (1900:64); subspecies recognized by 

Hall (1981:817) and Smolen (1999b:644).

Syntypes: The catalog entries are given below.

USNM 2834. Aux (Des) Plaines R., Ill. Alcohol. Sex and date 

collected unknown. Cataloged 31 May 1857. 

USNM 2835. Aux (Des) Plaines R., Ill. Alcohol. Sex and date 

collected unknown. Cataloged 31 May 1857.

USNM 2838. Aux (Des) Plaines R. Alcohol. Male. Date collected unknown. Cataloged 31 May 1857.

USNM 2839. Aux (Des) Plaines R. Alcohol. Male. Date collected unknown. Cataloged 31 May 1857.

USNM 2840. Aux (Des) Plaines R. Alcohol. Sex and date collected unknown. Cataloged 31 May 1857.

USNM 2920. West Northfield. Alcohol. Sex and date collected 

unknown. Cataloged 2 Jun 1857.

USNM 294. This number was incorrectly listed, as it belongs to 

a Sciurus from Maryland.

USNM 743/3233. West Northfield, Cook County. Female skin 

and skull. Collected spring 1855. Cataloged 1 Jun 1855.

USNM 2880. Anna, Ill. Male with skull inside. Collected 15 Apr. 

Cataloged 31 May 1857.

Type Locality: West Northfield, [Cook County], and Anna, 

[Union County], Illinois.

Remarks: Type not designated in original description, nor was 

the taxon included in type catalogs of Lyon and Osgood 

(1909) or Poole and Schantz (1942). Baird (1857:547) 

stated that this name applies to the specimens from Illinois, 

and in the table on page 548 he listed the above specimens, 

all collected by R. Kennicott. Only the skull belonging to 

USNM 743, number 3233, is currently in the collection.

Arvicola (Mynomes) longicaudus Merriam

Am. Nat. 22: 934, Oct 1888c.

= Microtus longicaudus longicaudus (Merriam, 1888). See Goldman (1938b:491); subspecies recognized by Hall (1981:808) 

and Smolen (1999a:652).

Holotype: USNM 186496. Skin and skull. Adult female. Collected 19 Jul 1888, not 1887 as in original description, by 

V. Bailey. Original number [SL: 105]. Merriam collection 

number 4509/5144.

Type Locality: Custer, in the Black Hills, [V. Bailey (1900:48): altitude 5,500 ft (1,676 m)], [Custer County, South] Dakota.

Remarks: Type published as Merriam collection number 

4509/5144; cataloged as USNM 186496 on 20 Oct 1911.

Arvicola longirostris Baird

Mammals Rep. Explor. Surv. 8(1): 530, 1857.

= Microtus montanus montanus (Peale, 1848). See Grinnell 

(1913:317); subspecies recognized by Sera and Early (2003:1).

Holotype: USNM 1268/2220. Mounted skin and [fragmentary] 

skull. [Lyon and Osgood (1909:87): Adult] female. Collected 

[MC: Sep 1855] by J. S. Newberry. No original number.

Type Locality: Upper Pit River, California.

Arvicola (Mynomes) macropus Merriam

N. Am. Fauna 5: 60, 30 Jul 1891a.

= Microtus richardsoni macropus (Merriam, 1891). See V. Bailey 

(1900:61); subspecies recognized by Hall (1981:820) and 

Ludwig (1999:646).

Holotype: USNM 23887/31291. Skin and skull. Adult female. 

Collected 16 Sep 1890 by C. H. Merriam and V. Bailey. 

Original number [FC: VB] 1803.

Type Locality: Pahsimeroi Mountains, [FC: head of Pahsimeroi 

River, altitude 9,700 ft (2,957 m) at or above timberline], 

[Custer County], Idaho.

Arvicola modesta Baird

Mammals Rep. Explor. Surv. 8(1): 535, 1857.

= Microtus pennsylvanicus modestus (Baird, 1857). See V. Bailey 

(1900:20); subspecies recognized by Hall (1981:795) and 

Tamarin (1999:642).

Holotype: USNM 594/1717. Skin and skull. Immature, sex unknown. Collected Aug–Sep 1853 by [F.] Kreutzfeldt. Original number 16.

Type Locality: Sawatch Pass [unattributed note with the skin 

label: Saguache Pass, Cochetopa Hills, south of Gunnison 

river], [Saguache County], Rocky Mountains, Colorado. 

Baird (1857:714) approximated the location of Sawatch 

Pass as “probably at the entrance of Sawatch Valley and 

near the Cochetopa Pass.”

Arvicola montana Peale

U.S. Explor. Exped. 8: 44, 1848.

= Microtus montanus montanus (Peale, 1848). See Miller (1912a: 

216); subspecies recognized by Sera and Early (2003:1).

Holotype: USNM number unknown.

Type Locality: Peale (1848:45) reported that “our specimen 

was obtained on the 4th of October, near the head waters 

of the Sacramento River, in California.” Miller and Rehn 

(1901:118) added near Mount Shasta, Siskiyou County.

Remarks: This specimen cannot be found and was listed as missing by Lyon and Osgood (1909) and Poole and Schantz 

(1942).

Arvicola (Mynomes) mordax Merriam

N. Am. Fauna 5: 61, 30 Jul 1891a.
number 642 • 1 7

= Microtus longicaudus longicaudus (Merriam, 1891). See Long 

(1965:653); subspecies recognized by Hall (1981:808) and 

Smolen (1999a:632).

Holotype: USNM 24231/31635. Skin and skull. Adult male. 

Collected 29 Sep 1890 by C. H. Merriam and V. Bailey. 

Original number [FC: VB 1903].

Type Locality: Sawtooth or Alturus Lake, [FC: altitude 7,200 ft 

(2,195 m)], east foot of Sawtooth Mountains, [Blaine 

County]. Idaho.

Arvicola (Mynomes) nanus Merriam

N. Am. Fauna 5: 63, 30 Jul 1891a.

= Microtus montanus nanus (Merriam, 1891). See Hall (1938:133); 

subspecies recognized by Sera and Early (2003:1).

Holotype: USNM 23853/31253. Skin and skull. Adult female, 

not male as in original description. Collected 16 Sep 1890 

by C. H. Merriam and V. Bailey. Original number [FC: VB 

1809].

Type Locality: [FC: Head of Pahsimeroi River, altitude 9,700 

ft (2,957 m)], Pahsimeroi Mountains, altitude 2,850 m or 

9,350 ft, [Custer County], Idaho.

Arvicola occidentalis Peale

U.S. Explor. Exped. 8: 45, 1848.

= Microtus townsendii townsendii (Bachman, 1839). See Hayman and Holt (1941:602). Subspecies recognized by Hall 

(1981:805) and Cornely (1999a:647).

Holotype: USNM 10083/4157. Skin and skull. Age and sex unknown. Collected by T. R. Peale, date unknown. Skin cataloged in 1872; skull in 1860.

Type Locality: Puget Sound, Oregon (Washington).

Remarks: Type not designated by number in original description. 

Although he gave no specimen number, Baird (1857:535) 

stated that the specimen he described is Peale’s original one. 

The 1872 entry in the museum catalog indicates “type of 

occidentalis” in the remarks column. The specimen bears a 

Coues’s Monograph of North American Muridae skin label 

on which is written “Type of Arvicola occidentalis Peale.” 

Coues (1877:173) listed 10083 as being the type of occidentalis. Therefore, the holotype is fixed by monotypy according to ICZN (1999: Articles 72.4.1.1 and 73.1.2).

Arvicola operarius Nelson

Proc. Biol. Soc. Wash. 8: 139, 28 Dec 1893.

= Microtus oeconomus operarius (Nelson, 1893). See Zimmermann (1942:187); subspecies recognized by Hall (1981:806) 

and Conroy (1999:639).

Holotype: USNM 14379/22225. Skin and skull. [Lyon and Osgood (1909:90): Adult] unsexed. Collected Nov 1879 by 

E. W. Nelson. Original number [SL: 122].

Type Locality: St. Michaels (Michael), Norton Sound, Alaska.

Arvicola phaeus Merriam

Proc. Biol. Soc. Wash. 7: 171, 29 Sep 1892a.

= Microtus mexicanus phaeus (Merriam, 1892). See V. Bailey 

(1900:54); subspecies retained by Hall (1981:815).

Holotype: USNM 33605/45640. Skin and skull. Adult female. Collected 21 Apr 1892 by E. W. Nelson. Original number 2516.

Type Locality: North slope Sierra Nevada de Colima, altitude 

[FC: about] 10,000 ft (3,048 m), Jalisco, Mexico.

Arvicola (Pitymys) pinetorum quasiater Coues

Proc. Acad. Nat. Sci. Phila. 26: 191, 15 Dec 1874.

= Microtus quasiater (Coues, 1874). See Miller (1896b:60). 

Holotype: USNM 3524. Skin. Age and sex unknown. Collected 

by R. Monte d’Oca, date unknown. Cataloged 18 Mar 

1859. No original number. There is no record of a skull, 

and the skin cannot be found.

Type Locality: Xalapa (Jalapa), [Veracruz], Mexico.

Arvicola riparia var. longipilis Baird

Mammals Rep. Explor. Surv. 8(1): 524, 1857.

= Microtus pennsylvanicus pennsylvanicus (Ord, 1815). See V. Bailey (1900:16); subspecies recognized by Hall (1981:795) and 

Tamarin (1999:642).

Syntypes: USNM 585/1708. Skin made from alcohol and skull. 

Female. West Northfield, [Cook County], Illinois. Collected 

by R. Kennicott, date unknown. Skin cataloged 19 Mar 

1855; skull 16 Mar 1855. The skin cannot be found.

USNM 867/1920. Skin and skull. Male. West Northfield, [Cook 

County], Illinois. Collected by R. Kennicott, date unknown. 

Cataloged 10 Oct 1855. Skin cannot be found.

USNM 2172. Skull only. Sex unknown. Racine, [Racine County], 

Wisconsin. Collected by Dr. Hoy, date unknown. Cataloged 

25 Feb 1856.

Type Locality: West Northfield, Illinois, and Racine, [Wisconsin].

Remarks: Type specimen not designated by number in original 

description, nor was a list of specimens presented. Baird 

(1857:524) stated that the name longipilis applies to those 

specimens from West Northfield, Illinois, and Racine, [Wisconsin]. The USNM collection currently contains three 

specimens that meet this stipulation; they were cataloged or 

labeled as riparius, thus eliminating other species of Microtus

that occur at these localities, and they were cataloged prior 

to 1857, so they would have been available to Baird. G. M. 

Allen (1931:260) considered these cotypes (syntypes), and we 

follow his lead. Catalog data are given above. A fourth specimen, USNM 745, is currently number 5292 in the Museum 

of Comparative Zoology (Helgen and McFadden, 2001:114). 

It is a Kennicott specimen from West Northfield, Illinois.

Arvicola scalopsoides Audubon and Bachman

Proc. Acad. Nat. Sci. Phila. 1: 97, read at stated meeting of 5 

Oct 1841. Receipt acknowledged 1 Dec 1841 by the Albany 

Institute.

= Microtus pinetorum scalopsoides (Audubon and Bachman, 

1841). See Batchelder (1896:187); subspecies recognized by 

Hall (1981:817) and Smolen (1999b:644).
1 8 • smithsonian contributions to zoology

Lectotype: USNM 10264 as designated by Coues (1877:224). 

[Skin with skull inside. Age and sex unknown]. [MC: Collected by/received from Major J. E. LeConte, date unknown. 

Cataloged 15 Oct 1872].

Type Locality: Long Island, [New York].

Remarks: Type not designated by number in original description. 

In the remarks column of the museum catalog “Type of scalopsoides” is written in the same hand as the original entry; 

the specimen bears a Coues’s Monograph of American Muridae skin label that states “10264 Type of A. ‘scalopsoides’ apud LeConte (= pinetorum) Long Island,” and Coues 

(1877:224) listed 10264 as the type of scalopsoides. Audubon and Bachman (1841:97) stated they had collected many 

specimens; therefore, Coues’s (1877:224) listing constituted 

designation of a lectotype (ICZN, 1999: Article 74.5).

Arvicola trowbridgii Baird

Mammals Rep. Explor. Surv. 8(1): 529, 1857.

= Microtus californicus californicus (Peale, 1848). See Kellogg 

(1918:5); subspecies recognized by Hall (1981:801) and 

Heske and Lidicker (1999:627).

Holotype: USNM 370/1284. Mounted skin [MC: from alcohol] 

and skull. Age and sex unknown. Collected on unknown 

date by W. P. Trowbridge. Cataloged 4 Jan 1855. No original number.

Type Locality: Monterey, [Monterey County], California.

Microtus abbreviatus Miller

Proc. Biol. Soc. Wash. 13: 13, 31 Jan 1899b.

= Microtus abbreviatus abbreviatus Miller, 1899. See Miller 

(1924:414); subspecies recognized by Hall (1981:819) and 

Cook and Klein (1999:624).

Holotype: USNM 15540/22429. Skin and skull. Young adult female. Collected 8 Sep 1885 by C. H. Townsend. No original 

number.

Type Locality: Hall Island, Bering Sea, [Alaska].

Microtus abbreviatus fisheri Merriam

Proc. Wash. Acad. Sci. 2: 23, 14 Mar 1900a.

Subspecies recognized by Hall (1981:819) and Cook and Klein 

(1999:624).

Holotype: USNM 97976. [Skin and skull]. Adult male. Collected 

15 Jul 1899 by A. K. Fisher. Original number 2189.

Type Locality: Saint Matthew Island, Bering Sea, [SL: Alaska].

Microtus amakensis Murie

J. Mammal. 11: 74, 11 Feb 1930.

= Microtus oeconomus amakensis Murie, 1930. See Hall and 

Cockrum (1952:309); subspecies recognized by Hall (1981: 

805) and Conroy (1999:639).

Holotype: USNM 246449. [Skin and skull]. Adult male. Collected 8 Jul 1925 by D. H. Stevenson. Original number 80.

Type Locality: Amak Island, [FC: 12 mi (19 km) off coast of 

Alaska Peninsula], [Aleutian Islands], Bering Sea, Alaska.

Microtus angusticeps V. Bailey

Proc. Biol. Soc. Wash. 12: 86, 30 Apr 1898a.

= Microtus longicaudus angusticeps V. Bailey, 1898. See Goldman (1938b:491); subspecies recognized by Hall (1981:807) 

and Smolen (1999a:632).

Holotype: USNM 17087/24008. Skin and skull. Adult male. 

Collected 16 Jun 1889 by T. S. Palmer. Original number 

151.

Type Locality: Crescent City, [Del Norte County], California.

Microtus aphorodemus Preble

N. Am. Fauna 22: 52, 31 Oct 1902a.

= Microtus pennsylvanicus aphorodemus Preble, 1902. See Anderson (1937:112); subspecies recognized by Hall (1981: 

792) and Tamarin (1999:642).

Holotype: USNM 106422. Skin and skull. Adult female. Collected 5 Aug 1900 by E. A. Preble. Original number 3208.

Type Locality: [Lyon and Osgood (1909:81): West shore Hudson 

Bay], barren grounds about 50 mi (80 km) south of Cape 

Eskimo, near mouth of Thlewiaza River, Keewatin [District, 

Nunavut, Canada].

Microtus arvalis duplicatus Miller

Cat. Mammals West. Eur., p. 686, 1912b.

Subspecies recognized by Gromov and Polyakov (1992:435).

Holotype: USNM 112213. Skin and skull. Adult male. Collected 

1901 by L. J. Boettcher. Original number 11. Holotype selected by von den Brink (1930:251).

Type Locality: Tenkitten, near Konigsberg (Kaliningrad), Germany (Kaliningradskaya, Russia).

Microtus bairdi Merriam

Proc. Biol. Soc. Wash. 11: 74, 21 Apr 1897c.

= Microtus oregoni bairdi Merriam, 1897. See Taylor (1920: 

180); subspecies recognized by Hall (1981:810) and Wolff 

(1999:640).

Holotype: USNM 79906. [Skin and skull]. Adult female. Collected 24 Aug 1896 by C. H. Merriam and V. Bailey. Original number [FC: VB] 5813.

Type Locality: Crater Lake, altitude about 7,800 ft (2,377 m), 

[SL and FC: north slope] Glacier Peak, [Klamath County], 

Oregon.

Microtus californicus constrictus V. Bailey

N. Am. Fauna 17: 36, 6 Jun 1900.

Subspecies recognized by Hall (1981:801) and Heske and 

Lidicker (1999:627).

Holotype: USNM 98347. [Skin and skull]. [Lyon and Osgood 

(1909:84): Adult] [FC: male]. Collected 6 Sep 1899 by 

V. Bailey. Original number 7174.

Type Locality: Cape Mendocino, [Humboldt County], California.

Microtus californicus vallicola V. Bailey

Proc. Biol. Soc. Wash. 12: 89, 30 Apr 1898a.
number 642 • 1 9

Subspecies recognized by Hall (1981:803) and Heske and 

Lidicker (1999:628).

Holotype: USNM 25393/32806. Skin and skull. Adult female. Collected 28 Dec 1890 by E. W. Nelson. Original number 149.

Type Locality: Lone Pine, [FC: upper edge of Alabamas], Inyo 

County, California. Restricted by A. B. Howell (1923a:266) 

to Lone Pine Creek, about 4,500 ft (1,372 m), Owens Valley, Inyo County, California.

Microtus canicaudus Miller

Proc. Biol. Soc. Wash. 11: 67, 21 Apr 1897a.

Holotype: USNM 75841. [Skin and skull]. Adult male. Collected 

1 Dec 1895 by B. J. Bretherton. Original number 2119, 

rather than 219 as in original description.

Type Locality: McCoy, Willamette Valley, [Polk County], Oregon.

Microtus dutcheri V. Bailey

Proc. Biol. Soc. Wash. 12: 85, 30 Apr 1898a.

= Microtus montanus dutcheri V. Bailey, 1898. See Grinnell 

(1913:317); subspecies recognized by Sera and Early (2003:1).

Holotype: USNM 29679/41848. Skin and skull. Adult male. Collected 10 Jul 1891 by B. H. Dutcher. Original number 69.

Type Locality: [FC: Head of] Big Cottonwood Meadows, near 

Mount Whitney, altitude 10,000 ft (3,048 m), [Tulare 

County], California.

Microtus elymocetes Osgood

Proc. Biol. Soc. Wash. 19: 71, 1 May 1906.

= Microtus oeconomus elymocetes Osgood, 1906. See Zimmermann (1942:188); subspecies recognized by Hall (1981:805) 

and Conroy (1999:639).

Holotype: USNM 137323. [Skin and skull]. Adult male. Collected 12 May 1905 by C. Sheldon. Original number 8. [SL: 

5448-X].

Type Locality: East side of Montague Island, Prince William 

Sound, [Valdez-Cordova], Alaska.

Microtus fortis uliginosus Jones and Johnson

Proc. Biol. Soc. Wash. 68: 193, 31 Dec 1955.

Subspecies recognized by Won and Smith (1999:26).

Holotype: USNM 298078. Skin and skull. Adult female. Collected 28 Aug 1952 by W. H. Lawrence. Original number 

B12098, rather than B-2098 as in original description.

Type Locality: Chip’o-ri, 38°08′N, 127°19′E, [South] Korea.

Microtus fulviventer Merriam

Proc. Biol. Soc. Wash. 12: 106, 30 Apr 1898b.

= Microtus mexicanus fulviventer Merriam, 1898. See Musser 

(1964:20). Subspecies recognized by Hall (1981:814).

Holotype: USNM 68250. [Skin and skull]. Adult male. Collected 

22 Aug 1894 by E. W. Nelson and E. A. Goldman. Original 

number 6601.

Type Locality: Cerro San Felipe, altitude 10,200 ft (3,109 m), 

Oaxaca, Mexico.

Microtus guatemalensis Merriam

Proc. Biol. Soc. Wash. 12: 108, 30 Apr 1898b.

Holotype: USNM 76777. [Skin and skull]. Adult male. Collected 

30 Dec 1895 by E. W. Nelson and E. A. Goldman. Original 

number 8960.

Type Locality: Todos Santos, altitude 10,000 ft (3,048 m), Huehuetenango, Guatemala.

Microtus innuitus Merriam

Proc. Wash. Acad. Sci. 2: 21, 14 Mar 1900a.

= Microtus oeconomus innuitus Merriam, 1900. See Zimmermann (1942:188); subspecies recognized by Hall (1981:805) 

and Conroy (1999:639).

Holotype: USNM 99373. Skull only from owl pellet. Adult, sex 

unknown. Collected 13 Jul 1899 by C. H. Merriam. No 

original number.

Type Locality: Northeast Cape, Saint Lawrence Island, Bering 

Sea, [SL: Alaska].

Microtus insularis V. Bailey

Proc. Biol. Soc. Wash. 12: 86, 30 Apr 1898a. 

Secondary junior homonym of Lemmus insularis Nilsson, 1844 

= Microtus agrestis (Linnaeus, 1761). Renamed Microtus 

nesophilus by V. Bailey (1898b:783).

= Microtus pennsylvanicus nesophilus V. Bailey, 1898. See Jones 

et al. (1986:4); subspecies recognized by Whitaker and 

Hamilton (1998:351).

Holotype: USNM 186494. Skin and skull. Immature male. Collected 6 Aug 1888 by B. H. Dutcher. No original number. 

Merriam collection number 4952/5706.

Type Locality: Great Gull Island, [Miller and Rehn (1901:118): 

off eastern extremity of Long Island, Suffolk County], New 

York.

Remarks: Published as Merriam collection number 4952/5706; 

cataloged as USNM 186494 on 20 Oct 1911.

Microtus kadiacensis Merriam

Proc. Biol. Soc. Wash. 11: 222, 15 Jul 1897a.

= Microtus oeconomus operarius (Nelson, 1893). See Paradiso 

and Manville (1961:78); subspecies recognized by Hall 

(1981:806) and Conroy( 1999:639).

Holotype: USNM 65827. Skin and skull. Adult female. Collected 

12 Sep 1893 by B. J. Bretherton. Original number 213.

Type Locality: Kodiak Island, Alaska.

Microtus longicaudus baileyi Goldman

J. Mammal. 19: 492, 14 Nov 1938b.

Subspecies recognized by Hall (1981:807) and Smolen (1999a: 

632).

Holotype: USNM 248924. Skin and skull. Adult male. Collected 

13 Jul 1929 by V. Bailey. Original number 10731.

Type Locality: Greenland Spring, head of Bright Angel Creek, 

altitude 8,000 ft (2,438 m), [SL and FC: Kaibab Forest], 

Grand Canyon National Park, Coconino County, Arizona.
2 0 • smithsonian contributions to zoology

Microtus ludovicianus V. Bailey

N. Am. Fauna 17: 74, 6 Jun 1900.

= Microtus ochrogaster ludovicianus V. Bailey, 1900. See Raun 

and Laughlin (1972:493); subspecies recognized by Hall 

(1981:812) and Stalling (1999:638).

Holotype: USNM 96624. Skin and skull. Adult male. Collected 

7 Apr 1899 by V. Bailey. Original number 6767.

Type Locality: Iowa [SL and FC: Station], Calcasieu Parish, 

Louisiana.

Microtus macfarlani Merriam

Proc. Wash. Acad. Sci. 2: 24, 14 Mar 1900a.

= Microtus oeconomus macfarlani Merriam, 1900. See Zimmermann (1942:187); subspecies recognized by Hall (1981:805) 

and Conroy (1999:639).

Holotype: USNM 9155/37347. Skin and skull. [Age and sex 

unknown]. Collected by R. MacFarlane [date unknown]. 

[MC: Skin cataloged Jul 1868; skull 31 Mar 1898]. Original number 3179.

Type Locality: Fort Anderson, [V. Bailey (1900:40): Anderson 

River], north of Great Bear Lake, Arctic America [Mackenzie District, Northwest Territories, Canada].

Microtus macrurus Merriam

Proc. Acad. Nat. Sci. Phila. 50: 353, 4 Oct 1898c.

= Microtus longicaudus macrurus Merriam, 1898. See Goldman 

(1938b:491); subspecies recognized by Hall (1981:809) and 

Smolen (1999a:632).

Holotype: USNM 66151. [Skin and skull]. Adult female. Collected 26 Jun 1894 by C. P. Streator. Original number 3975.

Type Locality: Lake Cushman, Olympic Mountains, [FC: Mason 

County], Washington.

Microtus mexicanus guadalupensis V. Bailey

Proc. Biol. Soc. Wash. 15: 118, 2 Jun 1902a.

Subspecies recognized by Hall (1981:814) as listed above and by 

Frey (1999a:635) under Microtus mogollonensis.

Holotype: USNM 109191. [Skin and skull]. Adult male. Collected 21 Aug 1901 by V. Bailey. Original number 7807.

Type Locality: [FC: Head of McKittrick Canyon], altitude 

7,800 ft (2,377 m), Guadalupe Mountains, [Culberson 

County], Texas.

Microtus mexicanus hualpaiensis Goldman

J. Mammal. 19: 493, 14 Nov 1938c.

Subspecies recognized by Hall (1981:814) as listed above and by 

Frey (1999a:635) under Microtus mogollonensis.

Holotype: USNM 244108. Skin and skull. Adult female. Collected 

1 Oct 1923 by E. A. Goldman. Original number 23554.

Type Locality: Hualpai (Hualapai) Peak, Hualpai (Hualapai) 

Mountains, altitude 8,400 ft (2,560 m), Mohave County, 

Arizona.

Microtus mexicanus madrensis Goldman

J. Mammal. 19: 493, 14 Nov 1938c.

Subspecies recognized by Hall (1981:814).

Holotype: USNM 98172. Skin and skull. Adult male. Collected 

2 Jul 1899 by [E. W.] Nelson and [E. A.] Goldman. Original 

number 13870.

Type Locality: Gavilan River, 5 mi (8 km) west of Colonia Garcia, about 60 mi (97 km) southwest of Casas Grandes, altitude 6,700 ft (2,042 m), Chihuahua, Mexico.

Microtus mexicanus subsimus Goldman

J. Mammal. 19: 494, 14 Nov 1938c.

Subspecies recognized by Hall (1981:815).

Holotype: USNM 116918. Skin and skull. Adult male. Collected 

3 May 1902 by [E. W.] Nelson and [E. A.] Goldman. Original number 15195.

Type Locality: Sierra Guadalupe, [FC: altitude about 9,500 ft 

(2,896 m)], southeastern Coahuila, Mexico.

Microtus miurus Osgood

N. Am. Fauna 21: 64, 26 Sep 1901a.

= Microtus miurus miurus Osgood, 1901. See Miller (1912a:220); 

subspecies recognized by Cole and Wilson (2010:75).

Holotype: USNM 107175. [Skin and skull]. Adult male. Collected 4 Sep 1900 by W. H. Osgood and E. Heller. Original 

number [FC: WHO] 1349.

Type Locality: Head of Bear Creek, in mountains near Hope 

City, Turnagain Arm, Cook Inlet, Alaska.

Microtus miurus oreas Osgood

Proc. Biol. Soc. Wash. 20: 61, 18 Apr 1907b.

Subspecies recognized by Cole and Wilson (2010:75).

Holotype: USNM 148596. [Skin and skull]. Adult male. Collected 8 Aug 1906 by C. Sheldon. Original number 47. [SL: 

6289-X].

Type Locality: Head of Toklat River, [SL: Mount McKinley], 

Alaskan Range, Alaska.

Microtus miurus paneaki Rausch

J. Wash. Acad. Sci. 40: 135, 21 Apr 1950.

= Microtus miurus muriei Nelson, 1931. See Hall and Cockrum (1952:311); subspecies recognized by Cole and Wilson 

(2010:75).

Holotype: USNM 290296. Skin and skull. Young adult male. 

Collected 2 Jun 1949 by R. Rausch. Original number 21.

Type Locality: Tolugak Lake, [SL: Anaktuvuk Pass], Brooks 

Range, Alaska. Lat. 68°24′N, Long. 152°10′W.

Microtus montanus arizonensis V. Bailey

Proc. Biol. Soc. Wash. 12: 88, 30 Apr 1898a.

Subspecies recognized by Sera and Early (2003:1).

Holotype: USNM 24778/32179. Skin and skull. Adult male. Collected 7 Nov 1890 by E. W. Nelson. Original number 153.

Type Locality: Springerville, [Apache County], Arizona.

Microtus montanus caryi V. Bailey

Proc. Biol. Soc. Wash. 30: 29, 21 Feb 1917.
number 642 • 2 1

= Microtus montanus nanus (Merriam, 1891). See Anderson 

(1954:493); subspecies recognized by Sera and Early (2003:1).

Holotype: USNM 168670. [Skin and skull]. Adult male. Collected 8 May 1910 by M. Cary. Original number 1912.

Type Locality: Milford, [FC: 5,500 ft (1,676 m)], Fremont 

County, Wyoming.

Microtus mordax bernardinus Merriam

Proc. Biol. Soc. Wash. 21: 145, 9 Jun 1908a.

= Microtus longicaudus bernardinus Merriam, 1908. See Goldman (1938b:492); subspecies recognized by Hall (1981:807) 

and Smolen (1999a:632).

Holotype: USNM 159632. [Skin and skull]. Adult male. Collected 21 Aug 1907 by V. Bailey. Original number 8749.

Type Locality: Dry Lake, at north base of San Gorgonio Peak, altitude 9,000 ft (2,743 m), San Bernardino Mountains, [San 

Bernardino County], California.

Microtus muriei Nelson

J. Mammal. 12: 311, 24 Aug 1931.

= Microtus miurus muriei Nelson, 1931. See Hall and Cockrum (1952:312); subspecies recognized by Cole and Wilson 

(2010:75).

Holotype: USNM 243482. [Skin and skull]. Adult female. Collected 

7 Mar 1923 by O. J. Murie. Original number [SL: 1737].

Type Locality: Kutuk River, tributary of Alatna River, Endicott 

Mountains, Alaska.

Microtus nanus canescens V. Bailey

Proc. Biol. Soc. Wash. 12: 87, 30 Apr 1898a.

= Microtus montanus canescens V. Bailey, 1898. See Hall (1938: 

133); subspecies recognized by Sera and Early (2003:1).

Holotype: USNM 90577. [Skin and skull]. Adult male. Collected 

12 Sep 1897 by J. A. Loring. Original number 4654.

Type Locality: Conconully, [Okanogan County], Washington.

Microtus nevadensis V. Bailey

Proc. Biol. Soc. Wash. 12: 86, 30 Apr 1898a.

= Microtus montanus nevadensis V. Bailey, 1898. See Hall 

(1935:423); subspecies recognized by Sera and Early (2003:1).

Holotype: USNM 26877/34291. Skin and skull. Adult male, not 

female as in original description. Collected 2 Mar 1891 by 

E. W. Nelson. Original number 577.

Type Locality: Ash Meadows, Nye County, Nevada.

Microtus nevadensis rivularis V. Bailey

Proc. Biol. Soc. Wash. 12: 87, 30 Apr 1898a.

= Microtus montanus rivularis V. Bailey, 1898. See V. Bailey 

(1900:29); subspecies recognized by Sera and Early (2003:1).

Holotype: USNM 186497. Skin and skull. Adult male. Collected 

6 Jan 1889 by V. Bailey. Original number 493. Merriam collection number 5172/5867.

Type Locality: Saint George, [Washington County], Utah.

Remarks: Published as Merriam collection number 5172/5867; 

cataloged as USNM 186497 on 20 Oct 1911.

Microtus oeconomus gilmorei Setzer

Proc. Biol. Soc. Wash. 65: 75, 25 Apr 1952.

= Microtus oeconomus operarius (Nelson, 1893). See Rausch 

(1953:130); subspecies recognized by Hall (1981:806) and 

Conroy (1999:639).

Holotype: USNM 293109. Skin and skull. Adult male. Collected 

4 Jul 1951 by H. W. Setzer. Original number 2396.

Type Locality: Point Lay, 163°04′W Long. and 69°46′N Lat., 

Alaska.

Microtus operarius endoecus Osgood

N. Am. Fauna 30: 23, 7 Oct 1909b.

= Microtus oeconomus macfarlani Merriam, 1900. See Anderson (1947:157); subspecies recognized by Hall (1981:805) 

and Conroy (1999:639).

Holotype: USNM 128327. [Skin and skull]. Adult male. Collected 21 Jun 1903 by W. H. Osgood. Original number [SL: 

2366].

Type Locality: Mouth of Charlie Creek, [FC: NE bank of] Yukon 

River, about 50 mi (80 km) above Circle, Alaska.

Microtus oregoni adocetus Merriam

Proc. Biol. Soc. Wash. 21: 145, 9 Jun 1908a.

Subspecies recognized by Hall (1981:810) and Wolff (1999: 

640).

Holotype: USNM 137995. [Skin and skull]. Adult male. Collected 30 Jul 1905 by A. S. Bunnell. Original number 135.

Type Locality: South Yolla Bolly Mountain, [Tehama County], 

California.

Microtus oregoni cantwelli Taylor

J. Mammal. 1: 180, 24 Aug 1920.

= Microtus oregoni oregoni (Bachman, 1839). See Dalquest 

(1948:357); subspecies recognized by Hall (1981:810) and 

Wolff (1999:640).

Holotype: USNM 232814. [Skin and skull]. Adult male. Collected 

13 Aug 1919 by G. G. Cantwell. Original number 1487.

Type Locality: Glacier Basin, altitude 5,935 ft (1,809 m), Mount 

Rainier, [Pierce County], Washington.

Microtus pennsylvanicus labradorius V. Bailey

Proc. Biol. Soc. Wash. 12: 88, 30 Apr 1898a.

Subspecies recognized by Hall (1981:794) and Tamarin (1999: 

642).

Holotype: USNM 186495. Skin and skull, originally in alcohol. Adult female. Collected 15 Nov 1882 by L. M. Turner. 

Original number [SL: 296]. Merriam collection number 

5481/6566.

Type Locality: Fort Chimo (Kuujjuaq), Ungava [District, Quebec, Canada].

Remarks: Published as MCC 5481/6566. Cataloged as USNM 

186495 on 20 Oct 1911.

Microtus pennsylvanicus wahema V. Bailey

J. Mammal. 1: 72, 2 Mar 1920.
2 2 • smithsonian contributions to zoology

= Microtus pennsylvanicus insperatus (J. A. Allen, 1894). See Anderson (1943:92); subspecies recognized by Hall (1981:793) 

and Tamarin (1999:642).

Holotype: USNM 212370. [Skin and skull]. Adult male. Collected 8 May 1916 by R. Kellogg. Original number 425.

Type Locality: Glendive, [FC: across Yellowstone River], [Dawson County], Montana.

Microtus pinetorum auricularis V. Bailey

Proc. Biol. Soc. Wash. 12: 90, 30 Apr 1898a.

Subspecies recognized by Hall (1981:816) and Smolen (1999b: 

644).

Holotype: USNM 34062/46139. Skin and skull. Adult male. 

Collected 26 May 1892 by V. Bailey. Original number 3649.

Type Locality: Washington, [Adams County], Mississippi.

Microtus pinetorum nemoralis V. Bailey

Proc. Biol. Soc. Wash. 12: 89, 30 Apr 1898a.

Subspecies recognized by Hall (1981:816) and Smolen (1999b: 

644).

Holotype: USNM 87246. [Skin and skull]. Adult female. Collected 7 Apr 1897 by J. A. Loring. Original number 3905.

Type Locality: Stilwell, Boston Mountains, [Adair County], Indian Territory (Oklahoma).

Microtus ravidulus Miller

Proc. Acad. Nat. Sci. Phila. 51: 284, 9 Aug 1899a.

= Microtus gregalis (Pallas, 1779). See Hayman and Holt (1941: 

615).

Holotype: USNM 62159. Skin and skull. Adult female. Collected 7 Nov 1893 by W. L. Abbott. No original number.

Type Locality: Okchi (Akchi), valley of the Aksai, altitude 

7,000 ft (2,134 m), eastern Turkestan (Xinjiang, China).

Microtus scirpensis V. Bailey

N. Am. Fauna 17: 38, 6 Jun 1900.

= Microtus californicus scirpensis V. Bailey, 1900. See Kellogg 

(1918:24); subspecies recognized by Hall (1981:803) and 

Heske and Lidicker (1999:628).

Holotype: USNM 25979/33385. Skin and skull. Adult female. 

Collected 26 Feb 1891 by V. Bailey. Original number 2520.

Type Locality: Amargosa River, near Nevada line, [FC: 10 mi 

(16 km) north of Borax Works], [Kellogg (1918:24): spring 

near Shoshone, altitude 1,560 ft (475 m)], Inyo County, 

California.

Microtus serpens Merriam

Proc. Biol. Soc. Wash. 11: 75, 21 Apr 1897c.

= Microtus oregoni serpens Merriam, 1897. See Taylor 

(1920:180); subspecies recognized by Hall (1981:810) and 

Wolff (1999:640).

Holotype: USNM 76303. [Skin and skull]. Adult male. Collected 

2 Dec 1895 by C. P. Streator. Original number 5068.

Type Locality: Agassiz, British Columbia, [Canada].

Microtus sitkensis Merriam

Proc. Biol. Soc. Wash. 11: 221, 15 Jul 1897a.

= Microtus oeconomus sitkensis Merriam, 1897. See Zimmermann (1942:187); subspecies recognized by Hall (1981:806) 

and Conroy (1999:639).

Holotype: USNM 73839. Skin and skull. [Lyon and Osgood 

(1909:92): Young] adult male. Collected 3 Aug 1895 by 

C. P. Streator. Original number 4745.

Type Locality: [FC: High watermark at edge of forest], Sitka, 

Alaska.

Microtus tshuktshorum Miller

Proc. Biol. Soc. Wash. 13: 11, 31 Jan 1899c.

= Microtus oeconomus tshuktshorum Miller, 1899. See Zimmermann (1942:184); subspecies recognized by Gromov and 

Erbajeva (1995:482).

Holotype: USNM 8419/37610. In alcohol, [skull removed]. 

Adult female. Collected on unknown date by Lt. [Lyon and 

Osgood (1909:92): John] Dawson. Received in 1866 [MC: 

skin cataloged 12 Jan 1866; skull 10 Nov 1898]. Original 

number [MC: 211].

Type Locality: Plover Bay, East Siberia (Bering Strait, Chukotskiy, Russia).

Microtus umbrosus Merriam

Proc. Biol. Soc. Wash. 12: 107, 30 Apr 1898b.

Holotype: USNM 68480. [Skin and skull]. Adult female. Collected 10 Jul 1894 by E. W. Nelson and E. A. Goldman. 

Original number 6412.

Type Locality: Mount Zempoaltepec, altitude 8,200 ft (2,499 m), 

Oaxaca, Mexico.

Microtus unalascensis Merriam

Proc. Biol. Soc. Wash. 11: 222, 15 Jul 1897a.

= Microtus oeconomus unalascensis Merriam, 1897. See 

Zimmermann (1942:187); subspecies recognized by Hall 

(1981:806) and Conroy (1999:639).

Holotype: USNM 30772/42672. Skin and skull. Immature female. Collected 13 Aug 1891 by C. H. Merriam. Original 

number Z.

Type Locality: Unalaska, Alaska.

Microtus unalascensis popofensis Merriam

Proc. Wash. Acad. Sci. 2: 22, 14 Mar 1900a.

= Microtus oeconomus popofensis Merriam, 1900. See Zimmermann (1942:187); subspecies recognized by Hall (1981:806) 

and Conroy (1999:639).

Holotype: USNM 97956. [Skin and skull]. Adult male. Collected 

16 Jul 1899 by W. E. Ritter. Original number 2200, [FC: A. 

K.] Fisher catalog.

Type Locality: Popof [Island], Shumagin Islands, Alaska.

Microtus yakutatensis Merriam

Proc. Wash. Acad. Sci. 2: 22, 14 Mar 1900a.
number 642 • 2 3

= Microtus oeconomus yakutatensis Merriam, 1900. See Zimmermann (1942:187); subspecies recognized by Hall (1981: 

806) and Conroy (1999:639).

Holotype: USNM 98005. [Skin and skull]. Adult male. Collected 

19 Jun 1899 by C. H. Merriam. Original number [FC: A. K. 

Fisher] 2101.

Type Locality: North shore Yakutat Bay, Alaska.

Pitymys parvulus A. H. Howell

Proc. Biol. Soc. Wash. 29: 83, 4 Apr 1916.

= Microtus pinetorum parvulus (A. H. Howell, 1916). See Hall 

(1981:817); subspecies recognized by Smolen (1999b:644).

Holotype: USNM 210485. Skin and skull. Adult female. Collected 15 Dec 1915 by R. T. Jackson. Original number [XC: 

42]. [SL: 13149-X].

Type Locality: Ocala, [SL and XC: Lynne Planting Station], 

[Marion County], Florida.

Pitymys pinetorum carbonarius Handley

J. Wash. Acad. Sci. 42: 152, 15 May 1952.

= Microtus pinetorum carbonarius (Handley, 1952). See Hall 

and Cockrum (1953:449); subspecies recognized by Hall 

(1981:816) and Smolen (1999b:644).

Holotype: USNM 191204. Skin and skull. Old adult female. 

Collected 14 Feb 1888 by J. B. Lewis. Original number 76. 

[SL: Merriam collection number 4157/4692].

Type Locality: Eubank, Pulaski County, Kentucky.

Pitymys pinetorum schmidti Jackson

Proc. Biol. Soc. Wash. 54: 201, 8 Dec 1941.

= Microtus pinetorum schmidti (Jackson, 1941). See Hall and 

Cockrum (1953:451); subspecies recognized by Hall (1981: 

816) and Smolen (1999b:644).

Holotype: USNM 248079. Skin and skull. Adult female. Collected 4 Sep 1927. Presented by F. J. W. Schmidt [who was 

probably the collector]. Original number 192. 24658-X.

Type Locality: Worden Township, Clark County, Wisconsin.

Genus Myodes Pallas, 1811

Clethrionomys gapperi arizonensis Cockrum and Fitch

Univ. Kans. Publ. Mus. Nat. Hist. 5: 291, 15 Nov 1952.

= Myodes gapperi arizonensis (Cockrum and Fitch, 1952). New 

name combination; subspecies recognized by Hall (1981:780) 

and Merritt (1999a:615) under Clethrionomys gapperi.

Holotype: USNM 158401. Skin and skull. Adult female. Collected 12 Sep 1908 by C. Birdseye. Original number 152.

Type Locality: Little Colorado River, altitude 8,300 ft (2,530 m), 

White Mountains, Apache County, Arizona.

Clethrionomys gapperi cascadensis Booth

Murrelet 26: 27, 10 Aug 1945.

= Myodes gapperi cascadensis (Booth, 1945). New name combination; subspecies recognized by Hall (1981:780) and Merritt (1999a:615) under Clethrionomys gapperi.

Holotype: USNM 242380. Skin and skull. Adult female. Collected 30 Oct 1921 by G. G. Cantwell. Original number 

2800.

Type Locality: [FC: Swank Creek], 2 mi (3.2 km) south of 

Blewett Pass, altitude 3,000 ft (914 m), Kittitas County, 

Washington.

Clethrionomys gapperi gauti Cockrum and Fitch

Univ. Kans. Publ. Mus. Nat. Hist. 5: 289, 15 Nov 1952.

= Myodes gapperi gauti (Cockrum and Fitch, 1952). New name 

combination; subspecies recognized by Hall (1981:781) and 

Merritt (1999a:615) under Clethrionomys gapperi.

Holotype: USNM 133515. Skin and skull. Adult male. Collected 

7 Aug 1904 by J. H. Gaut. Original number 3086.

Type Locality: Twinning, altitude 10,700 ft (3,261 m), Taos 

County, New Mexico.

Clethrionomys gapperi maurus Kellogg

Proc. Biol. Soc. Wash. 52: 37, 11 Mar 1939. 

= Myodes gapperi maurus (Kellogg, 1939). New name combination; subspecies recognized by Hall (1981:782) and Merritt 

(1999a:615) under Clethrionomys gapperi.

Holotype: USNM 267826. Skin and skull. Adult female. Collected 23 Jun 1938 by W. M. Perrygo and H. J. Cole. Original number 705.

Type Locality: 4.5 mi (7 km) southeast of Lynch, altitude 4,100 

ft (1,250 m), Black Mountains, Harlan County, Kentucky.

Clethrionomys gapperi solus Hall and Cockrum

Univ. Kans. Publ. Mus. Nat. Hist. 5: 304, 17 Nov 1952.

= Myodes gapperi solus (Hall and Cockrum, 1952). New name 

combination; subspecies recognized by Hall (1981:784) and 

Merritt (1999a:615) under Clethrionomys gapperi.

Holotype: USNM 74939. Skin and skull. Adult male. Collected 

22 Sep 1895 by C. P. Streator. Original number 4980.

Type Locality: Loring, Revillagigedo Island, Alaska.

Clethrionomys imaizumii Jameson

Pac. Sci. 15: 599, Oct 1961.

= Myodes imaizumii (Jameson, 1961). See Musser and Carleton 

(2005:1024).

Holotype: USNM 324906. Skin and skull. Adult male. Collected 

13 Feb 1959 by E. W. Jameson Jr. Original number 1083.

Type Locality: Nachi Falls, 300 m elevation, Wakayama-ken, 

Honshu, Japan.

Evotomys alascensis Miller

Proc. Acad. Nat. Sci. Phila. 50: 364, 15 Oct 1898b.

= Myodes rutilus dawsoni (Merriam, 1888). New name combination; subspecies recognized by Hall (1981:778) and Batzli 

(1999b:617) under Clethrionomys rutilus.

Holotype: USNM 14359/22226. Skin and skull. Adult male. Collected 26 Oct 1879 by E. W. Nelson. Original number 96.

Type Locality: Saint Michael’s, [Norton Sound], Alaska.
2 4 • smithsonian contributions to zoology

Evotomys californicus Merriam

N. Am. Fauna 4: 26, 8 Oct 1890b.

= Myodes californicus californicus (Merriam, 1890). New name 

combination; subspecies recognized by Hall (1981:784) and 

Alexander (1999:613) under Clethrionomys californicus.

Holotype: USNM 17011/23920. Skin and skull. Adult male. 

Collected 3 Jun 1889 by T. S. Palmer. Original number 110.

Type Locality: Eureka, Humboldt County, California.

Evotomys carolinensis Merriam

Am. J. Sci. (Ser. 3) 36: 460, Dec 1888d.

= Myodes gapperi carolinensis (Merriam, 1888). New name 

combination; subspecies recognized by Hall (1981:784) and 

Merritt (1999a:615) under Clethrionomys gapperi.

Holotype: USNM 186490. [In alcohol]. Adult female. Collected 

11 Aug 1887 by C. H. Merriam. No original number. Merriam collection number 3660.

Type Locality: [MCC: Summit] Roan Mountain, altitude 6,000 ft 

(1,828 m), [Mitchell County], North Carolina.

Remarks: Published as Merriam collection number 3660; cataloged as USNM 186490 on 19 Oct 1911.

Evotomys caurinus V. Bailey

Proc. Biol. Soc. Wash. 12: 21, 17 Jan 1898c.

= Myodes gapperi caurinus (V. Bailey, 1898). New name combination; subspecies recognized by Hall (1981:780) and Merritt (1999a:615) under Clethrionomys gapperi.

Holotype: USNM 89460. [Skin and skull]. Adult male. Collected 

18 Jul 1897 by E. A. Preble. Original number 2147.

Type Locality: Lund, east shore of Malaspina Inlet, [FC: 185 

mi (137 km) northwest of Vancouver], British Columbia, 

[Canada].

Evotomys galei Merriam

N. Am. Fauna 4: 23, 8 Oct 1890c.

= Myodes gapperi galei (Merriam, 1890). New name combination; subspecies recognized by Hall (1981:780) and Merritt 

(1999a:615) under Clethrionomys gapperi.

Holotype: USNM 186491. Skin and skull. Adult female. Collected 13 Jul 1889 by D. Gale. No original number. Merriam collection number 5685/6352.

Type Locality: Near Gold Hill, [V. Bailey (1897:126): Ward, 

about 6 mi (10 km) above Gold Hill], altitude 9,500 ft 

(2,896 m), Boulder County, Colorado.

Remarks: Published as Merriam collection number 5685/6352; 

cataloged as USNM 186491 on 19 Oct 1911.

Evotomys gapperi athabascae Preble

N. Am. Fauna 27: 178, 26 Oct 1908.

= Myodes gapperi athabascae (Preble, 1908). New name combination; subspecies recognized by Hall (1981:780) and Merritt (1999a:615) under Clethrionomys gapperi.

Holotype: USNM 109945. Skin and skull. Adult female, not 

male as in original description. Collected 27 Jun 1901 by 

E. A. and A. E. Preble. Original number [FC: EAP] 4235.

Type Locality: Fort Smith, Slave River, near the Athabaska 

(Alberta)-Mackenzie line, Mackenzie [District, Northwest 

Territories], Canada.

Evotomys gapperi brevicaudus Merriam

N. Am. Fauna 5: 119, 30 Jul 1891b.

= Myodes gapperi brevicaudus (Merriam, 1891). New name 

combination; subspecies recognized by Hall (1981:780) and 

Merritt (1999a:615) under Clethrionomys gapperi.

Holotype: USNM 186489. Skin and skull. Adult male. Collected 

21 Jul 1888 by V. Bailey. Original number 111. Merriam 

collection number 4507/5142.

Type Locality: [Miller and Rehn (1901:115): 3 mi (4.8 km) north 

of] Custer, [Miller and Rehn (1901:115): altitude about 

6,000 ft (1,828 m)], Black Hills, [Custer County], South 

Dakota.

Remarks: Published as Merriam collection number 4507/5142; 

cataloged as USNM 186489 on 19 Oct 1911.

Evotomys gapperi loringi V. Bailey

Proc. Biol. Soc. Wash. 11: 125, 13 May 1897.

= Myodes gapperi loringi (V. Bailey, 1897). New name combination; subspecies recognized by Hall (1981:782) and Merritt 

(1999a:615) under Clethrionomys gapperi.

Holotype: USNM 75795. [Skin and skull]. Adult male. Collected 

22 Nov 1895 by J. A. Loring. Original number 3438.

Type Locality: Portland, [Traill County], North Dakota.

Evotomys hercynicus suecicus Miller

Proc. Wash. Acad. Sci. 2: 101, 26 Jul 1900b.

= Myodes glareolus (Schreber, 1780). See Musser and Carleton 

(2005:1024).

Holotype: USNM 85046. Skin and skull. Adult female. Collected 6 Aug 1898 by J. A. Loring. Original number 5009.

Type Locality: [FC: Ultuna, 3 mi (4.8 km) south of] Upsala (Uppsala), [Uppsala Municipality, Uppsala], Sweden.

Evotomys idahoensis Merriam

N. Am. Fauna 5: 66, 30 Jul 1891a.

= Myodes gapperi idahoensis (Merriam, 1891). New name combination. Subspecies recognized by Hall (1981:781) and 

Merritt (1999a:615) under Clethrionomys gapperi.

Holotype: USNM 24283/31687. Skin and skull. Adult female. 

Collected 4 Oct 1890 by C. H. Merriam and V. Bailey. Original number [FC: VB] 1936.

Type Locality: Sawtooth or Alturas Lake, east foot of Sawtooth 

Mountains, [FC: 7,200 ft (1,295 m)], [Blaine County], Idaho.

Evotomys limitis V. Bailey

Proc. Biol. Soc. Wash. 26: 133, 21 May 1913.

= Myodes gapperi limitis (V. Bailey, 1913). New name combination; subspecies recognized by Hall (1981:781) and Merritt 

(1999a:615) under Clethrionomys gapperi.

Holotype: USNM 148335. [Skin and skull]. Adult male. Collected 27 Oct 1906 by V. Bailey. Original number 8572.
number 642 • 2 5

Type Locality: Willow Creek, a branch of the Gilita, altitude 

8,500 ft (2,591 m), Mogollon Mountains, [Catron County], 

New Mexico.

Evotomys mazama Merriam

Proc. Biol. Soc. Wash. 11: 71, 21 Apr 1897d.

= Myodes californicus mazama (Merriam, 1897). New name 

combination; subspecies recognized by Hall (1981:785) and 

Alexander (1999:613) under Clethrionomys californicus.

Holotype: USNM 79913. [Skin and skull]. Adult male. Collected 

15 Aug 1896 by C. H. Merriam and V. Bailey. Original 

number [FC: VB] 5765.

Type Locality: Crater Lake, [Miller and Rehn (1901:115): 

Mount Mazama], south end of Cascade Range, altitude 

7,000 ft, 2,130 m [FC: 7,000 ft–7,100 ft (2,164 m)], [Klamath County], Oregon.

Evotomys nivarius V. Bailey

Proc. Biol. Soc. Wash. 11: 136, 13 May 1897. 

= Myodes gapperi nivarius (V. Bailey, 1897). New name combination; subspecies recognized by Hall (1981:782) and Merritt (1999a:615) under Clethrionomys gapperi.

Holotype: USNM 66203. Skin and skull. Adult female. Collected 9 Jul 1894 by C. P. Streator. Original number 4025.

Type Locality: [FC: Lake Cushman], northwest slope of Mount 

Ellinor, altitude 4,000 ft, 1,220 m, Olympic Mountains, 

[FC: Mason County], Washington.

Evotomys norvegicus Miller

Proc. Wash. Acad. Sci. 2: 93, 26 Jul 1900b.

= Myodes glareolus (Schreber, 1780). See Musser and Carleton 

(2005:1024).

Holotype: USNM 84674. Skin and skull. Adult female. Collected 31 May 1898 by T. Stejneger. Original number 20.

Type Locality: Bergen, [Bergen Municipality, Hordaland], Norway.

Evotomys obscurus Merriam

Proc. Biol. Soc. Wash. 11: 72, 21 Apr 1897d.

= Myodes californicus obscurus (Merriam, 1897). New name 

combination; subspecies recognized by Hall (1981:785) and 

Alexander (1999:613) under Clethrionomys californicus.

Holotype: USNM 80413. [Skin and skull]. Adult male. Collected 

29 Aug 1896 by E. A. Preble. Original number 1455.

Type Locality: Prospect, Upper Rogue River Valley at west base 

of Mount Mazama, altitude about 2,600 ft or 800 m, [FC: 

Jackson County], Oregon.

Evotomys occidentalis Merriam

N. Am. Fauna 4: 25, 8 Oct 1890b.

= Myodes gapperi occidentalis (Merriam, 1890). New name 

combination; subspecies recognized by Hall (1981:785) and 

Merritt (1999a:615) under Clethrionomys gapperi.

Holotype: USNM 17447/24351. Skin and skull. Adult male. 

Collected 16 Aug 1889 by T. S. Palmer. Original number 

308.

Type Locality: Aberdeen, Chehalis (Grays Harbor) County, 

Washington.

Evotomys orca Merriam

Proc. Wash. Acad. Sci. 2: 24, 14 Mar 1900a.

= Myodes rutilus orca (Merriam, 1900). New name combination; subspecies recognized by Hall (1981:779) and Batzli 

(1999b:617) under Clethrionomys rutilus.

Holotype: USNM 98028. [Skin and skull]. Adult female. Collected 28 Jun 1899 by A. K. Fisher. Original number 2139.

Type Locality: Orca, Prince William Sound, Alaska.

Evotomys ungava V. Bailey

Proc. Biol. Soc. Wash. 11: 130, 13 May 1897.

= Myodes gapperi ungava (V. Bailey, 1897). New name combination; subspecies recognized by Hall (1981:784) and Merritt 

(1999a:615) under Clethrionomys gapperi.

Holotype: USNM 186492. Skin and skull. Adult male. Collected 

12 May 1883 by L. M. Turner. Original number 317. Merriam collection number 5471/6158.

Type Locality: Fort Chimo (Kuujjuaq), Ungava [District, Quebec, Canada].

Remarks: Originally cataloged as USNM 14209, this specimen 

was exchanged with the Merriam collection and recataloged 

on 19 Oct 1911 when that collection was donated to USNM.

Evotomys vasconiae Miller

Proc. Wash. Acad. Sci. 2: 96, 26 Jul 1900b.

= Myodes glareolus (Schreber, 1780). See Musser and Carleton 

(2005:1024).

Holotype: USNM 86994. Skin and skull. Adult male. Collected 

4 Jul 1899 by R. T. Young. Original number [SL: 625].

Type Locality: Montrejeau, [Haute-Garonne Department], 

Hautes Pyrenees (Midi-Pyrénées), France.

Evotomys wrangeli V. Bailey

Proc. Biol. Soc. Wash. 11: 120, 13 May 1897.

= Myodes gapperi wrangeli (V. Bailey, 1897). New name combination; subspecies recognized by Hall (1981:784) and Merritt (1999a:615) under Clethrionomys gapperi.

Holotype: USNM 74724. [Skin and skull]. Adult female. Collected 1 Sep 1895 by C. P. Streator. Original number 4835.

Type Locality: Wrangel (Wrangell), [SL: sea level], [FC: at high 

tide mark], Wrangel (Wrangell) Island, Alaska.

Genus Myopus Miller, 1910

Myopus morulus Hollister

Smithson. Misc. Collect. 60(14): 1, 29 Nov 1912.

= Myopus schisticolor (Lilljeborg, 1844). See Hayman and Holt 

(1941:561).

Holotype: USNM 175197. Skin and skull. Adult male. Collected 

6 Aug 1912 by N. Hollister. Original number 4437.

Type Locality: 5 mi (8 km) south of Tapucha, altitude 6,876 

ft (2,096 m) [SL: 6,725 ft (2,050 m)]. Tapucha is 125 mi 
2 6 • smithsonian contributions to zoology

(201 km) southwest of Biisk and 50 mi (80 km) northwest 

of Ongudai on the post road to Kosh-Agatch, Altai Mountains, [Altayskiy], Siberia (Russia).

Genus Neacomys Thomas, 1900

Neacomys pictus Goldman

Smithson. Misc. Collect. 60(2): 6, 20 Sep 1912a.

Holotype: USNM 178717. [Skin and skull]. Adult male. Collected 13 Mar 1912 by E. A. Goldman. Original number 

21512.

Type Locality: Cana (Santa Cruz de Cana), 1,800 ft (549 m), 

[Miller and Kellogg (1955:442): upper Rio Tuyru, Darien], 

Panama.

Genus Nectomys Peters, 1861

Nectomys squamipes pollens Hollister

Proc. Biol. Soc. Wash. 27: 104, 11 May 1914a.

Subspecies recognized by Ernest (1986:1).

Holotype: USNM 121399. Skin and skull. Adult male. Collected 

27 Jul 1902 by W. Foster. Original number 800.

Type Locality: Sapucay, [Paraguari], Paraguay.

Nectomys squamipes tarrensis Hershkovitz

Proc. U.S. Natl. Mus. 98: 51, 30 Jun 1948a.

= Nectomys rattus (Pelzein, 1883). See Musser and Carleton 

(2005:1133).

Holotype: USNM 279741. Skin and skull. Adult male. Collected 

8 Aug 1943, not 6 as in original description, by P. Hershkovitz. Original number 2319.

Type Locality: Rio Tarra, altitude 250 m, [SL: San Calixto], 

upper Rio Catatumbo, department of Norte de Santander, 

Colombia.

Nectomys squamipes vallensis Hershkovitz

Misc. Publ. Mus. Zool. Univ. Mich. 58: 61, 4 Jan 1944.

= Nectomys apicalis Peters, 1861. See Musser and Carleton 

(2005:1132).

Holotype: USNM 194911. Skin and skull. Adult male. Collected 

by E. C. Erdis. Original number 12. Number 577 of the Yale 

University–National Geographic Society’s Peruvian Expedition, 1914–1915.

Type Locality: Santa Ana, altitude 3,480 ft (1,061 m), Rio 

Urubamba Valley, department of Cuzco (Cusco), Peru.

Genus Nelsonia Merriam, 1897

Nelsonia goldmani Merriam

Proc. Biol. Soc. Wash. 16: 80, 29 May 1903a.

= Nelsonia goldmani goldmani Merriam, 1903. See Engstrom et 

al. (1992:880).

Holotype: USNM 125818. [Skin and skull]. Adult male. Collected 25 Feb 1903 by E. W. Nelson and E. A. Goldman. 

Original number 16021.

Type Locality: Mount Tancitaro, [FC: altitude about 9,000 ft 

(2,743 m)], Michoacan, Mexico.

Nelsonia neotomodon Merriam

Proc. Biol. Soc. Wash. 11: 278, 17 Dec 1897e.

Holotype: USNM 90891. [Skin and skull]. Adult male. Collected 

3 Sep 1897 by E. W. Nelson and E. A. Goldman. Original 

number 11625.

Type Locality: Mountains near Plateado, altitude 2,500 m or 

8,200 ft [FC: 8,500 ft (2,591 m)], Zacatecas, Mexico.

Genus Neodon Horsfield, 1841

Microtus pamirensis Miller

Proc. Acad. Nat. Sci. Phila. 51: 287, 9 Aug 1899a.

= Neodon juldaschi (Severtzov, 1879). See Hayman and Holt 

(1941:620).

Holotype: USNM 62161. Skin and skull. Adult male. Collected 

18 Jun 1894 by W. L. Abbott. No original number.

Type Locality: Tagdumbash Pamir, altitude 12,000 ft (3,658 m), 

[Xinjiang], Central Asia (China).

Genus Neofiber True, 1884

Neofiber alleni True

Science 4: 34, 11 Jul 1884.

= Neofiber alleni alleni True, 1884. See Miller (1924:424); 

subspecies recognized by Hall (1981:823) and Birkenholz 

(1999:651).

Holotype: USNM 1405/21948. [Originally in alcohol, now a 

skin, skull, skeleton, and viscera in alcohol]. [Lyon and Osgood (1909:80): Adult male]. Collected on unknown date 

by [Bangs (1898:183): F. Ulrich]. Received from W. Wittfeld 

24 Dec 1883; cataloged 17 Jan 1884. No original number.

Type Locality: Georgiana, [Bangs (1898:183): Merritt Island], 

[Brevard County], Florida.

Remarks: Type not designated by number in original description. Lyon and Osgood (1909:80) indicated that True said 

this was the original specimen, and according to museum 

records, it was the only Neofiber in the collection at the 

time of publication and was the only Neofiber in the series 

of nine specimens received from Wittfeld at that time. Holotype fixed by monotypy (ICZN, 1999: Article 73.1.2).

Neofiber alleni apalachicolae Schwartz

Occas. Pap. Mus. Zool. Univ. Mich. 547: 14, 29 Jul 1953.

Subspecies recognized by Hall (1981:823) and Birkenholz 

(1999:651).

Holotype: USNM 263704. Skin and skull. Adult male. Collected 

11 Apr 1938 by L. C. Goldman. Original number 569.

Type Locality: Apalachicola, east side of [FC: Apalachicola] 

River, Franklin County, Florida.

Neofiber alleni nigrescens A. H. Howell

J. Mammal. 1: 79, 2 Mar 1920a.
number 642 • 2 7

Subspecies recognized by Hall (1981:823) and Birkenholz 

(1999:651).

Holotype: USNM 228392. Skin and skull. Adult male. Collected 

2 Mar 1918 by A. H. Howell. Original number 2269.

Type Locality: Ritta [Island], south shore of Lake Okeechobee, 

[Palm Beach County], Florida.

Genus Neotoma Say and Ord, 1825

Neotoma albigula mearnsi Goldman

Proc. Biol. Soc. Wash. 28: 135, 29 Jun 1915a.

Subspecies recognized by Hall (1981:753) and Mares (1999:598).

Holotype: USNM 202981. [Skin and skull]. Adult male. Collected 21 Nov 1913 by E. A. Goldman. Original number 

22300.

Type Locality: Tinajas Altas, [SL and FC: east base of Gila 

Mountains, altitude about 1,400 ft (427 m)], near international boundary, [Yuma County], southwestern Arizona.

Neotoma albigula seri Townsend

Bull. Am. Mus. Nat. Hist. 31: 125, 14 Jun 1912.

Subspecies recognized by Hall (1981:753) and Mares (1999:598).

Holotype: USNM 198404. [Skin and skull]. Male, age not given. 

Collected [SL: 13] Apr 1911 by [SL: H. E. Anthony. Original number 157].

Type Locality: Tiburon Island, [Gulf of California, Sonora, 

Mexico].

Remarks: Townsend (1912:125) identified American Museum 

of Natural History (AMNH) 31940 as the type of his new 

subspecies; the specimen was recataloged as USNM 198404 

on 26 Jun 1914.

Neotoma albigula sheldoni Goldman

Proc. Biol. Soc. Wash. 28: 136, 29 Jun 1915a.

Subspecies recognized by Hall (1981:753) and Mares (1999:598).

Holotype: USNM 206812. [Skin and skull]. Adult male. Collected in 1915 by C. Sheldon. Original number [SL: 19. 

11956- X].

Type Locality: Papago Tanks, Pinacate Mountains, Sonora , 

Mexico.

Neotoma albigula warreni Merriam

Proc. Biol. Soc. Wash. 21: 143, 9 Jun 1908b.

= Neotoma leucodon warreni Merriam, 1908. New name combination; subspecies recognized by Hall (1981:754) and 

Mares (1999:598) under N. albigula.

Holotype: USNM 151051. [Skin and skull]. Adult male. Collected 28 Nov 1907 by M. Cary. Original number [SL: 

1271].

Type Locality: Guames Ranch, altitude 4,600 ft (1,402 m), [FC: 

Shell Rock Canyon], northwest corner of Baca County, 

Colorado.

Neotoma arizonae Merriam

Proc. Biol. Soc. Wash. 8: 110, 31 Jul 1893.

= Neotoma cinerea arizonae Merriam, 1893. See Goldman 

(1910:106); subspecies recognized by Hall (1981:768) and 

Grayson (1999:600).

Holotype: USNM 186482. Skin and skull. Adult female. Collected 21 May 1888 by J. Sullivan. Original number [MCC: 

14]. Merriam collection number 4249/4980.

Type Locality: Keam[s] Canyon, Apache (Navajo) County, Arizona.

Remarks: Published as Merriam collection number 4249/4980; 

cataloged as USNM 186482 on 26 Jun 1911.

Neotoma baileyi Merriam

Proc. Biol. Soc. Wash. 9: 123, 2 Jul 1894b.

= Neotoma floridana baileyi Merriam, 1894. See V. Bailey 

(1905:109); subspecies recognized by Hall (1981:748) and 

Post (1999:603).

Holotype: USNM 186483. Skin and skull. Adult female. Collected 16 Jun 1888 by V. Bailey. Original number 41. Merriam collection number 4311/5034.

Type Locality: [FC: Clarks Canyon near] Valentine, [Cherry 

County], Nebraska.

Remarks: Published as Merriam collection number 4311/5034; 

cataloged as USNM 186483 on 18 Oct 1911.

Neotoma bryanti Merriam

Am. Nat. 21: 191, Feb 1887a.

= Neotoma bryanti bryanti Merriam, 1887. See Patton et al. 

(2008:375) for current use.

Holotype: USNM 186481. Skin in alcohol, skull removed. Immature male. Collected 11 Jan 1885 by W. E. Bryant. 

Original number [MCC: 69a]. Merriam collection number 

2838/1833.

Type Locality: Cerros (Cedros) Island, altitude about 2,000 ft 

(610 m), lat. 28°12′N, off Lower California (Baja California), [Mexico].

Remarks: Published as Merriam collection number 2838/1833; 

cataloged as USNM 186481 on 18 Oct 1911.

Neotoma cinerea lucida Goldman

Proc. Biol. Soc. Wash. 30: 111, 23 May 1917.

Subspecies recognized by Hall (1981:769) and Grayson (1999: 

600).

Holotype: USNM 208968. [Skin and skull]. Adult female. Collected 1 Jul 1915 by L. J. Goldman. Original number 2282.

Type Locality: [SL and FC: North base] Charleston Peak, 

Charleston Mountains, [Clark County], Nevada.

Neotoma cumulator Mearns

Proc. U.S. Natl. Mus. 20: 3, 5 Mar 1897b. Preprint of Proc. U.S. 

Natl. Mus. 20: 503, 19 Jan 1898.

= Neotoma albigula venusta True, 1894. See Goldman 

(1910:34); subspecies recognized by Hall (1981:753) and 

Mares (1999:598).

Holotype: USNM 60348. Skin and skull. Adult male. Collected 

2 Apr 1894 by E. A. Mearns [and SL: F. X. Holzner]. Original number 3473.
2 8 • smithsonian contributions to zoology

Type Locality: Old Fort Yuma, San Diego (Imperial) County, 

California, [Miller (1924:379): on right bank of Colorado 

River opposite present town of Yuma, Arizona].

Neotoma desertorum Merriam

Proc. Biol. Soc. Wash. 9: 125, 2 Jul 1894b.

= Neotoma lepida lepida Thomas, 1893. See Goldman (1932b:61); 

subspecies recognized by Patton et al. (2008:386).

Holotype: USNM 25739/33139. Skin and skull. Adult male. 

Collected 31 Jan 1891 by T. S. Palmer. Original number 43.

Type Locality: Furnace Creek, Death Valley, [Inyo County], 

California.

Neotoma desertorum sola Merriam

Proc. Biol. Soc. Wash. 9: 126, 2 Jul 1894b.

= Neotoma bryanti intermedia Rhoads, 1894. See Patton et al. 

(2008:379).

Lectotype: USNM 31516 as designated by Jones and Fisher 

(1973:435). Skin only. Adult male. Collected 24 October 

1891 by E. W. Nelson. Original number 1369.

Type Locality: San Emigdio, Kern County, California.

Remarks: As explained by Jones and Fisher (1973:435), the holotype, USNM 31516/43381, of Merriam’s new subspecies 

was a composite, the skin an example of N. lepida gilva

and the skull of N. fucipes. They designated the skin as the 

lectotype.

Neotoma fallax Merriam

Proc. Biol. Soc. Wash. 9: 123, 2 Jul 1894b.

= Neotoma mexicana fallax Merriam, 1894. See Goldman 

(1910:56); subspecies recognized by Hall (1981:762) and 

Cornely (1999b:609).

Holotype: USNM 186484. Skin and skull. Adult male. Collected 

1 Nov 1889 by D. Gale. No original number. Merriam collection number 5678/6345.

Type Locality: Gold Hill, [MCC: 5,500 ft (1,676 m)], Boulder 

County, Colorado.

Remarks: Published as Merriam collection number 5678/6345; 

cataloged as USNM 186484 on 19 Oct 1911.

Neotoma ferruginea chamula Goldman

Proc. Biol. Soc. Wash. 22: 141, 25 Jun 1909.

= Neotoma mexicana chamula Goldman, 1909. See Hall 

(1955:329); subspecies recognized by Hall (1981:761) and 

Cornely (1999b:609).

Holotype: USNM 76061. [Skin and skull]. Adult female, not 

male as in original description. Collected 29 Sep 1895 by 

E. W. Nelson and E. A. Goldman. Original number 8489.

Type Locality: Mountains near San Cristobal, [SL and FC: 9,500 

ft (2,896 m)], not 8,400 ft as in original description, Chiapas, Mexico.

Neotoma ferruginea ochracea Goldman

Proc. Biol. Soc. Wash. 18: 30, 2 Feb 1905.

= Neotoma mexicana ochracea Goldman, 1905. See Hall (1955 

:330); subspecies recognized by Hall (1981:762) and 

Cornely (1999b:609).

Holotype: USNM 34142/46219. Skin and skull. Nearly adult 

male. Collected 21 May 1892 by E. W. Nelson. Original 

number 2653.

Type Locality: Atemajac, altitude about 4,500 ft (1,372 m), near 

Guadalajara, Jalisco, Mexico.

Neotoma ferruginea solitaria Goldman

Proc. Biol. Soc. Wash. 18: 31, 2 Feb 1905.

= Neotoma mexicana solitaria Goldman, 1905. See Hall (1955: 

331); subspecies recognized by Hall (1981:763) and Cornely 

(1999b:609).

Holotype: USNM 76908. [Skin and skull]. Adult male. Collected 

17 Dec 1895 by E. W. Nelson and E. A. Goldman. Original 

number 8813.

Type Locality: Nenton, altitude 3,500 ft (1,067 m), [Huehuetenango], Guatemala.

Neotoma floridana illinoensis A. H. Howell

Proc. Biol. Soc. Wash. 23: 28, 23 Mar 1910.

Subspecies recognized by Hall (1981:749) and Post (1999:603).

Holotype: USNM 167752. [Skin and skull]. Adult female. Collected 

12 Jan 1910 by J. Johnson. No original number. [SL: 7778-X].

Type Locality: Wolf Lake, [Union County], Illinois.

Neotoma fulviventer Merriam

Proc. Biol. Soc. Wash. 9: 121, 2 Jul 1894b.

= Neotoma mexicana torquata Ward, 1891. See Hall (1955:331); 

subspecies recognized by Hall (1981:763) and Cornely 

(1999b:609).

Holotype: USNM 50165. [Skin and skull]. Adult female. Collected 5 Nov 1892 by E. W. Nelson. Original number 3744.

Type Locality: Toluca Valley, [FC: 9,000 ft (2,743 m)], [Mexico], 

Mexico.

Neotoma fuscipes Baird

Mammals Rep. Explor. Surv. 8(1): 495, 1857.

= Neotoma fuscipes fuscipes Baird, 1857. See Miller (1912a:199); 

subspecies recognized by Hall (1981:765) and Cranford 

(1999:604). 

Holotype: USNM 22026. [Skeleton. Adult] male. Collected Feb 

1856 by E. Samuels. No original number.

Type Locality: Petaluma, [Sonoma County], California.

Remarks: Specimen originally cataloged 10 Apr 1857 as USNM 

2679 in alcohol; later prepared as a skeleton and assigned 

its current USNM number on 9 Mar 1885.

Neotoma fuscipes dispar Merriam

Proc. Biol. Soc. Wash. 9: 124, 2 Jul 1894b.

= Neotoma macrotis simplex True, 1894. New name combination; subspecies recognized by Hall (1981:767) and Cranford (1999:604) under N. fuscipes.
number 642 • 2 9

Holotype: USNM 25391/32804. Skin and skull. Adult male. Collected 25 Dec 1890 by [SL: V. Bailey]. Original number 2310.

Type Locality: Lone Pine, Owens Valley, [Inyo County], 

California.

Neotoma fuscipes streatori Merriam

Proc. Biol. Soc. Wash. 9: 124, 2 Jul 1894b.

= Neotoma macrotis streatori Merriam, 1894. New name combination; subspecies recognized by Hall (1981:767) and 

Cranford (1999:604) under N. fuscipes.

Holotype: USNM 64439. [Skin and skull]. Adult male. Collected 

4 Apr 1894 by C. P. Streator. Original number 3685.

Type Locality: Carbondale, Amador County, California.

Neotoma goldmani Merriam

Proc. Biol. Soc. Wash. 16: 48, 19 Mar 1903b.

Holotype: USNM 116894. [Skin and skull]. Young adult male. 

Collected 18 Apr 1902 by E. W. Nelson and E. A. Goldman. 

Original number 15101.

Type Locality: Saltillo, [FC: altitude 5,000 ft (1,524 m)], Coahuila, Mexico.

Neotoma insularis Townsend

Bull. Am. Mus. Nat. Hist. 31: 125, 14 Jun 1912.

See Patton et al. (2008:381) for current use.

Holotype: USNM 198405. [Skin and skull]. Age not given, male, 

not female as given in original description. Collected 11 Apr 

1911 by [SL: H. E. Anthony. Original number 136].

Type Locality: Angel de la Guardia Island, [Gulf of California], 

Lower California (Baja California), [Mexico].

Remarks: Published incorrectly as AMNH number 21922; 

should be AMNH 31922; cataloged as USNM 198405 on 

26 Jun 1914.

Neotoma intermedia angusticeps Merriam

Proc. Biol. Soc. Wash. 9: 127, 2 Jul 1894b.

= Neotoma albigula albigula Hartley, 1894. See Goldman 

(1910:31); subspecies recognized by Hall (1981:752) and 

Mares (1999:598).

Holotype: USNM 186485. Skin and skull. Adult male. Collected 

12 Apr 1886 by A. W. Anthony. Original number 62. Merriam collection number 2325/2833.

Type Locality: Southwest corner of Grant (Hidalgo) County, 5 

mi (8 km) from Mexican Boundary, New Mexico.

Remarks: Published as Merriam collection number 2325/2833; 

cataloged as USNM 186485 on 19 Oct 1911.

Neotoma intermedia devia Goldman

Proc. Biol. Soc. Wash. 40: 205, 2 Dec 1927.

= Neotoma devia Goldman, 1927. See Musser and Carleton 

(2005:1055) and Patton et al. (2008:388).

Holotype: USNM 226376. [Skin and skull]. Adult female. Collected 27 Jul 1917 by E. A. Goldman. Original number 

23152.

Type Locality: Tanner Tank, altitude 5,200 ft (1,585 m), Painted 

Desert, [Coconino County], Arizona.

Neotoma intermedia melanura Merriam

Proc. Biol. Soc. Wash. 9: 127, 2 Jul 1894b.

= Neotoma albigula melanura Merriam, 1894. See Goldman 

(1910:35); subspecies recognized by Hall (1981:752) and 

Mares (1999:598).

Holotype: USNM 17819/24756. Skin and skull. Young adult 

male. Collected 13 Nov 1889 by V. Bailey. Original number 

671.

Type Locality: Ortiz, Sonora, Mexico.

Neotoma intermedia notia Nelson and Goldman

Proc. Biol. Soc. Wash. 44: 108, 17 Oct 1931a.

= Neotoma bryanti bryanti Merriam, 1887. See Patton et al. 

(2008:375) for current use.

Holotype: USNM 146794. [Skin and skull]. Adult male. Collected 27 Jan 1906 by [E. W.] Nelson and [E. A.] Goldman. 

Original number 19017.

Type Locality: La Laguna, altitude 5,500 ft (1,676 m), Sierra 

de la Victoria, southern Lower California (Baja California 

Sur), Mexico.

Neotoma intermedia perpallida Goldman

Proc. Biol. Soc. Wash. 22: 139, 25 Jun 1909.

= Neotoma bryanti bryanti Merriam, 1887. See Patton et al. 

(2008:375) for current use.

Holotype: USNM 79061. [Skin and skull]. Young adult male. 

Collected 4 Aug 1895 by J. E. McLellan. Original number 

1449.

Type Locality: San Jose Island, [Gulf of California], off east coast 

of Lower California (Baja California Sur), Mexico.

Neotoma intermedia pretiosa Goldman

Proc. Biol. Soc. Wash. 22: 139, 25 Jun 1909.

= Neotoma bryanti bryanti Merriam, 1887. See Patton et al. 

(2008:375) for current use.

Holotype: USNM 146123. [Skin and skull]. Adult male. Collected 17 Nov 1905 by E. W. Nelson and E. A. Goldman. 

Original number 18589.

Type Locality: Matancita, also called Soledad, altitude 100 ft 

(30 m), [FC: 75 ft (23 m)], 50 mi (80 km) north of Magdalena Bay, Lower California (Baja California Sur), Mexico.

Neotoma intermedia ravida Nelson and Goldman

Proc. Biol. Soc. Wash. 44: 107, 17 Oct 1931a.

= Neotoma bryanti bryanti Merriam, 1887. See Patton et al. 

(2008:375) for current use.

Holotype: USNM 140692. [Skin and skull]. Adult male. Collected 7 Nov 1905 by [E. W.] Nelson and [E. A.] Goldman. 

Original number 18476.

Type Locality: Comondu, altitude 700 ft (213 m), southern 

Lower California (Baja California Sur), Mexico.
3 0 • smithsonian contributions to zoology

Neotoma intermedia vicina Goldman

Proc. Biol. Soc. Wash. 22: 140, 25 Jun 1909.

= Neotoma bryanti bryanti Merriam, 1887. See Patton et al. 

(2008:378) for current use.

Holotype: USNM 146803. [Skin and skull]. Adult male. Collected 9 Feb 1906 by E. W. Nelson and E. A. Goldman. 

Original number 19082.

Type Locality: Espiritu Santo Island, [FC: altitude 30 ft (9 m)], 

[Gulf of California], off coast of southern Lower California 

(Baja California Sur), Mexico.

Neotoma isthmica Goldman

Proc. Biol. Soc. Wash. 17: 80, 21 Mar 1904.

= Neotoma mexicana isthmica Goldman, 1904. See Hall (1955: 

330); subspecies recognized by Hall (1981:762) and Cornely 

(1999:609).

Holotype: USNM 73187. [Skin and skull]. Adult female. Collected 5 May 1895 by E. W. Nelson and E. A. Goldman. 

Original number 7843.

Type Locality: Huilotepec, 8 mi (13 km) south of Tehuantepec, 

altitude 100 ft (30 m), Oaxaca, Mexico.

Neotoma latifrons Merriam

Proc. Biol. Soc. Wash. 9: 121, 2 Jul 1894b.

= Neotoma leucodon latifrons Merriam, 1894. New name combination; subspecies recognized by Hall (1981:753) and 

Mares (1999:598) under N. albigula.

Holotype: USNM 50135. [Skin and skull]. Adult male. Collected 

8 Aug 1892 by E. W. Nelson. Original number 3058.

Type Locality: Querendaro, [FC: altitude 5,800 ft (1,768 m)], 

Michoacan, Mexico.

Neotoma lepida marshalli Goldman

J. Mammal. 20: 357, 14 Aug 1939.

Subspecies recognized by Hall (1981:757) and MacMillen (1999: 

606).

Holotype: USNM 263984. Skin and skull. Adult female. Collected 2 Jul 1938 by W. H. Marshall. Original number 216.

Type Locality: Carrington Island, Great Salt Lake, altitude about 

4,250 ft (1,295 m), [Tooele County], Utah.

Neotoma lepida monstrabilis Goldman

J. Mammal. 13: 62, 9 Feb 1932b.

Subspecies recognized by Hall (1981:757) and MacMillen (1999: 

606).

Holotype: USNM 243123. [Skin and skull]. Adult male. Collected 

10 Oct 1922 by E. A. Goldman. Original number 23379.

Type Locality: Ryan, altitude 6,000 ft (1,829 m), Kaibab National Forest, Coconino County, Arizona.

Neotoma leucodon Merriam

Proc. Biol. Soc. Wash. 9: 120, 2 Jul 1894b.

= Neotoma leucodon leucodon Merriam, 1894; subspecies recognized by Hall (1981:757) and Mares (1999:598) under 

N. albigula.

Holotype: USNM 50137. [Skin and skull]. Adult male. Collected 

14 Aug 1892 by E. W. Nelson. Original number 3076.

Type Locality: San Luis Potosi, [FC: altitude about 6,000 ft 

(1,829 m)], [SL and FC: San Luis Potosi], Mexico.

Neotoma leucodon zacatecae Goldman

Proc. Biol. Soc. Wash. 18: 30, 2 Feb 1905.

= Neotoma leucodon leucodon Merriam, 1894. See Hall and 

Genoways (1970:514), who placed zacatecae in the subspecies leucodon in the species albigula. Subspecies recognized 

by Hall (1981:757) and Mares (1999:598) under N. albigula.

Holotype: USNM 90957. [Skin and skull]. Adult female. Collected 4 Sep 1897 by E. W. Nelson and E. A Goldman. Original number 11641.

Type Locality: Plateado, altitude 7,600 ft (2,316 m), Zacatecas, 

Mexico.

Neotoma macrotis simplex True

Proc. U.S. Natl. Mus. 17: 2, 2 Jun 1894e. Preprint of Proc. U.S. 

Natl. Mus. 17: 354, 15 Nov 1894f.

Subspecies recognized by Hall (1981:767) and Cranford (1999: 

604) under N. fuscipes.

Holotype: USNM 3651/3597. Skin and skull. [Lyon and Osgood 

(1909:105): Adult male]. Collected 1857–1858 by J. Xantus. 

[MC: Original number 785/786. Cataloged 24 Mar 1859].

Type Locality: [Miller and Rehn (1901:105) Old] Fort Tejon, 

[Miller and Rehn (1901:105): in mountains south of Kern 

Lake, Kern County], [Miller (1924:390): Tehachapi Mountains], California.

Remarks: Original description gives measurements for skull 

number 3598.

N[eotoma] magister Baird

Mammals Rep. Explor. Surv. 8(1): 498, 1857.

Syntypes, as designated by Lyon and Osgood (1909:100):

USNM 12206. Partial left mandible.

USNM 12207. Partial right mandible.

USNM 12208. Partial right mandible.

USNM 12209. Partial right mandible.

USNM 12210. Partial left mandible. Noted as missing in 1940 

by Poole and Schantz (1942:292) and still cannot be found.

USNM 12211. Partial left mandible.

USNM 12212. Partial left mandible.

USNM 12213. Partial right mandible.

USNM 12214. Fragment of left premaxilla.

Collected by S. F. Baird, date unknown. Cataloged 24 Oct 1872.

Type Locality: A cave near Carlisle, Pennsylvania.

Remarks: The locality given by Baird (1857:498) is actually “in 

the Bone Caves of Pennsylvania,” and the accession record, 

1664, says “from Caves near Carlisle, Pa,” but the singular 

Cave was used in designating the syntypes and therefore becomes the type locality.

Neotoma martinensis Goldman

Proc. Biol. Soc. Wash. 18: 28, 2 Feb 1905.
number 642 • 3 1

= Neotoma bryanti martinensis Goldman, 1905. See Patton et al. 

(2008:380) for current use.

Holotype: USNM 81074. [Skin and skull]. Adult female. Collected 17 Jul 1896 by A. W. Anthony. Original number 39.

Type Locality: San Martin Island, Lower California (Baja California), Mexico.

Neotoma mexicana Baird

Proc. Acad. Nat. Sci. Phila. 7: 333, reported favorably for publication 24 Apr 1855a; received 15 Jun 1855 by the American 

Philosophical Society.

= Neotoma mexicana mexicana Baird, 1855; subspecies recognized by Hall (1981:762) and Cornely (1999b:609).

Holotype: USNM 289/1674. [Skin, cannot be found, and skull]. 

[Lyon and Osgood (1909:101): Adult], sex unknown. Collected by J. Potts, date unknown. [MC: Skin cataloged 6 Jul 

1854; skull 10 Mar 1855].

Type Locality: [Miller and Rehn (1901:106): Mountains] near 

Chihuahua [Baird (1857:492): City], [Chihuahua, Mexico].

Remarks: Type not designated by number in original description. 

The list of specimens in Baird (1857:492) shows that USNM 

289/1674 is the only one from the type locality collected 

by Potts, as reasoned by Lyon and Osgood (1909:101) and 

affirmed by Poole and Schantz (1942:293). As there is no 

reason to believe Baird had more than one specimen, the holotype is fixed by monotypy (ICZN, 1999: Articles 72.4.1.1 

and 73.1.2).

Neotoma mexicana bullata Merriam

Proc. Biol. Soc. Wash. 9: 122, 2 Jul 1894b.

Subspecies recognized by Hall (1981:761) and Cornely (1999b: 

609).

Holotype: USNM 16863/23774. Skin and skull. Adult male. 

Collected 1 Jun 1889 by V. Bailey. Original number 114.

Type Locality: Santa Catalina Mountains, [Pima County], Arizona.

Neotoma mexicana inopinata Goldman

J. Wash. Acad. Sci. 23: 471, 15 Oct 1933.

Subspecies recognized by Hall (1981:762) and Cornely (1999b: 

609).

Holotype: USNM 158395. Skin and skull. Adult male. Collected 

8 Oct 1908 by C. Birdseye. Original number 246. 

Type Locality: Chuska Mountains, altitude 8,800 ft (2,682 m), 

[Miller and Kellogg (1955:545): San Juan County], northwestern New Mexico.

Neotoma mexicana inornata Goldman

Proc. Biol. Soc. Wash. 51: 60, 18 Mar 1938d.

Subspecies recognized by Hall (1981:762) and Cornely (1999b: 

609).

Holotype: USNM 263386. Skin and skull. Adult male. Collected 22 

Sep 1937 by R. S. Sturgis. Original number 79. [SL: 28074-X].

Type Locality: Carmen Mountains, altitude 6,100 ft (1,859 m), 

Coahuila, Mexico.

Neotoma mexicana madrensis Goldman

Proc. Biol. Soc. Wash. 18: 31, 2 Feb 1905.

= Neotoma mexicana mexicana Baird, 1855. See Anderson 

(1972:363); subspecies recognized by Hall (1981:762) and 

Cornely (1999b:609).

Holotype: USNM 95244. [Skin and skull]. Adult female. Collected 26 Aug 1898 by E. W. Nelson and E. A. Goldman. 

Original number 12918.

Type Locality: Sierra Madre, near Guadalupe y Calvo, altitude 

7,000 ft (2,134 m), Chihuahua, Mexico.

Neotoma micropus Baird

Proc. Acad. Nat. Sci. Phila. 7: 333, reported favorably for publication 24 Apr 1855a; received 15 Jun 1855 by the American 

Philosophical Society.

= Neotoma micropus micropus Baird, 1855; subspecies recognized by Hall (1981:751) and Braun (1999:610).

Lectotype: USNM 554/1676 as selected by Lyon and Osgood 

(1909:102). [Skull only; skin cannot be found]. [Lyon and 

Osgood (1909:101): Adult] [MC: male]. Collected [MC: 

Mar 1853] by Lieut. [D. N.] Couch. Original number [Baird 

(1857:435): 86].

Type Locality: Charco Escondido, [Tamaulipas], Mexico.

Remarks: Type not designated by number in original description. 

Lyon and Osgood (1909:102) considered USNM 554/1676 

to be the type that constituted selection of lectotype (ICZN, 

1999: Article 74.5). The museum catalog for USNM 554 is 

marked “not found 1902,” lined through, and replaced by 

“In coll. 17 Nov 1916.” However, it was reported as missing by both Lyon and Osgood (1909:102) and Poole and 

Schantz (1942:294).

Neotoma micropus leucophaea Goldman

J. Wash. Acad. Sci. 23: 472, 15 Oct 1933.

Subspecies recognized by Hall (1981:751) and Braun (1999:610).

Holotype: USNM 251057. Skin and skull. Adult male. Collected 

6 May 1933 by W. P. Taylor. Original number [SL and XC: 

A204]. [SL: 27347-X].

Type Locality: White Sands, 10 mi (16 km) west of Point of 

Sands, White Sands National Monument, altitude 4,100 ft 

(1,250 m), Otero (Doña Ana) County, New Mexico.

Neotoma micropus littoralis Goldman

Proc. Biol. Soc. Wash. 18: 31, 2 Feb 1905.

Subspecies recognized by Hall (1981:751) and Braun (1999:610).

Holotype: USNM 92952. [Skin and skull]. Adult male. Collected 

10 Apr 1898 by E. A. Goldman. Original number [FC: Nelson and Goldman] 12281.

Type Locality: Alta Mira, altitude 100 ft (30 m) [FC: about 75 ft 

(23 m)], Tamaulipas, Mexico.

Neotoma micropus planiceps Goldman

Proc. Biol. Soc. Wash. 18: 32, 2 Feb 1905.

Subspecies recognized by Hall (1981:751) and Braun (1999: 

610).
3 2 • smithsonian contributions to zoology

Holotype: USNM 82105. [Skin and skull]. Adult male. Collected 

16 Jan 1897 by E. W. Nelson and E. A. Goldman. Original 

number 10461.

Type Locality: Rio Verde, altitude [FC: about] 3,000 ft (914 m), 

San Luis Potosi, Mexico.

Neotoma montezumae Goldman

Proc. Biol. Soc. Wash. 18: 29, 2 Feb 1905.

= Neotoma leucodon leucodon Merriam, 1894. See Hall and 

Genoways (1970:515), who placed montezumae in the part 

of N. albigula that is now N. leucodon; subspecies recognized by Hall (1981:753) and Mares (1999:598) under 

N. albigula.

Holotype: USNM 81426. [Skin and skull]. Adult male. Collected 

17 Oct 1896 by E. W. Nelson and E. A. Goldman. Original 

number 10275.

Type Locality: Zimapan, altitude 7,500 ft (2,286 m) [FC: 6,200 ft 

(1,890 m)], Hidalgo, Mexico.

Neotoma navus Merriam

Proc. Biol. Soc. Wash. 16: 47, 19 Mar 1903b.

= Neotoma mexicana navus Merriam, 1903. See Hall (1955:330); 

subspecies recognized by Hall (1981:762) and Cornely 

(1999b:609).

Holotype: USNM 116895. [Skin and skull]. Adult female. Collected 26 Apr 1902 by E. W. Nelson and E. A. Goldman. 

Original number 15130.

Type Locality: Sierra Guadalupe, [FC: about 6,500 ft (1,981 m)], 

Coahuila, Mexico.

Neotoma nelsoni Goldman

Proc. Biol. Soc. Wash. 18: 29, 2 Feb 1905.

Holotype: USNM 54320. [Skin and skull]. Adult female. Collected 3 Jun 1893 by E. W. Nelson. Original number 4935.

Type Locality: Perote, altitude 7,800 ft (2,377 m), Veracruz, 

Mexico.

Neotoma nudicauda Goldman

Proc. Biol. Soc. Wash. 18: 28, 2 Feb 1905.

= Neotoma bryanti bryanti Merriam, 1887. See Patton et al. 

(2008:375).

Holotype: USNM 79073. [Skin and skull]. Nearly adult female. 

Collected 14 Oct 1895 by J. E. McLellan. Original number 

1517.

Type Locality: Carmen Island, [Gulf of California], Lower California (Baja California Sur), Mexico.

Neotoma occidentalis Baird

Proc. Acad. Nat. Sci. Phila. 7: 335, reported favorably for publication 24 Apr 1855b. Received 15 Jun 1855 by the American Philosophical Society.

= Neotoma cinerea occidentalis Baird, 1855. See Merriam 

(1891a:58); subspecies recognized by Hall (1981:769) and 

Grayson (1999:600).

Lectotype: USNM 572 as designated by Lyon and Osgood 

(1909:103). [Skin only, formerly mounted; no record of a 

skull]. [Lyon and Osgood (1909:102): Adult] [SL: male]. 

Collected [SL: 1854], [Lyon and Osgood (1909:102): probably Jun] by J. G. Cooper. Original number [SL: 16.]. [MC: 

Cataloged 14 Mar 1855].

Type Locality: Shoalwater (Willapa) Bay, [Pacific County, 

Washington].

Remarks: Type not designated by number in original description. Lyon and Osgood (1909:103) examined measurements 

given in Baird (1857:497) of the three specimens from the 

type locality and determined that USNM 572 was the 

only one whose measurements matched those given in the 

original description and considered it the type = lectotype 

(ICZN, 1999: Article 74.5).

Neotoma occidentalis fusca True

Proc. U.S. Natl. Mus. 17: 2, 27 Jun 1894e. Preprint of Proc. U.S. 

Natl. Mus. 17: 354, 15 Nov 1894f.

= Neotoma cinerea fusca True, 1894. See Trouessart (1897:544); 

subspecies recognized by Hall (1981:769) and Grayson 

(1999:600).

Holotype: USNM 3370. Skin only. [Age and sex unknown]. Collected by E. P. Vollum, [date unknown. No original number. 

MC: Cataloged in 1859, about 25 Feb].

Type Locality: Fort Umpqua, [Douglas County], Oregon.

Neotoma orizabae Merriam

Proc. Biol. Soc. Wash. 9: 122, 2 Jul 1894b.

= Neotoma mexicana torquata Ward, 1891. See Hall (1955:331); 

subspecies recognized by Hall (1981:763) and Cornely 

(1999:609).

Holotype: USNM 53653. [Skin and skull]. Adult male. Collected 

20 Apr 1893 by E. W. Nelson. Original number 4674.

Type Locality: Mount Orizaba, [FC: altitude 12,500 ft (3,810 m)], 

Puebla, Mexico.

Neotoma orolestes Merriam

Proc. Biol. Soc. Wash. 9: 128, 2 Jul 1894b.

= Neotoma cinerea orolestes Merriam, 1894. See Goldman 

(1910:104); subspecies recognized by Hall (1981:771) and 

Grayson (1999:600).

Holotype: USNM 35906/48215. Skin and skull. Adult male. Collected 13 Aug 1892 by J. A. Loring. Original number 482.

Type Locality: Saguache Valley, [FC: Tevebaugh’s Ranch], 20 mi 

(32 km) west of Saguache [FC: and 9 mi (14 km) south of 

Cochetopa Pass], [Saguache County], Colorado.

Neotoma palatina Goldman

Proc. Biol. Soc. Wash. 18: 27, 2 Feb 1905.

Holotype: USNM 90959. [Skin and skull]. Adult male. Collected 

12 Sep 1897 by E. W. Nelson and E. A. Goldman. Original 

number [SL and FC: 11710], not 7104 as given in original 

description.
number 642 • 3 3

Type Locality: Bolanos, altitude 2,800 ft (853 m), Jalisco, 

Mexico.

Neotoma parvidens Goldman

Proc. Biol. Soc. Wash. 17: 81, 21 Mar 1904.

= Neotoma mexicana parvidens Goldman, 1904. See Hall 

(1955:330); subspecies recognized by Hall (1981:762) and 

Cornely (1999:609).

Holotype: USNM 71586. [Skin and skull]. Adult female. Collected [SL and FC: 2 Mar 1895], not 27 Feb as in original 

description, by E. W. Nelson and E. A. Goldman. Original 

number 7587.

Type Locality: Juquila, altitude 5,000 ft (1,524 m), Oaxaca, 

Mexico.

Neotoma picta Goldman

Proc. Biol. Soc. Wash. 17: 79, 21 Mar 1904.

= Neotoma mexicana picta Goldman, 1904. See Hall (1955:331); 

subspecies recognized by Hall (1981:763) and Cornely 

(1999:609).

Holotype: USNM 70050. [Skin and skull]. Adult male. Collected 

20 Dec 1894 by E. W. Nelson and E. A. Goldman. Original 

number 7179.

Type Locality: Mountains near Chilpancingo, altitude 10,000 ft 

(3,048 m) [FC: 9,500 ft (2,896 m)], Guerrero, Mexico.

Neotoma pinetorum Merriam

Proc. Biol. Soc. Wash. 8: 111, 31 Jul 1893.

= Neotoma mexicana pinetorum Merriam, 1893. See Goldman 

(1910:58); subspecies recognized by Hall (1981:763) and 

Cornely (1999:609).

Holotype: USNM 17692/24628. Skin and skull. Adult female. 

Collected 16 Aug 1889 by V. Bailey. Original number 366.

Type Locality: San Francisco Mountain, [SL: 8,000 ft (2,438 m)], 

[Coconino County], Arizona.

Neotoma saxamans Osgood

N. Am. Fauna 19: 33, 6 Oct 1900.

= Neotoma cinerea occidentalis Baird, 1855. See Cowan and 

Guiguet (1956:195); subspecies recognized by Hall (1981: 

763) and Grayson (1999:600).

Holotype: USNM 98923. [Skin and skull]. Adult male. Collected 

19 Jun 1899 by W. H. Osgood. Original number 462.

Type Locality: Bennett City, head of Lake Bennett, British Columbia, [Canada].

Neotoma splendens True

Proc. U.S. Natl. Mus. 17: 1, 27 Jun 1894e. Preprint of Proc. U.S. 

Natl. Mus. 17: 353, 15 Nov 1894f.

= Neotoma fuscipes monochroura Rhoads, 1894. See Hooper 

(1938:217); subspecies recognized by Hall (1981:766) and 

Cranford (1999:604).

Holotype: USNM 19693. Skin, [no record of a skull]. [Lyon and 

Osgood (1909:105): Adult] male. Collected 25 Nov 1887, 

[collector unknown]. [Accession record 24265: purchased 

from C. K. Worthen]. [SL: Original number 211].

Type Locality: [Hooper (1938:217): Nicasio], Marin County, 

California.

Remarks: Although Hall (1981:766) placed splendens in N. f. 

fuscipes, his range map and list of marginal localities place 

it in monochroura.

Neotoma stephensi Goldman

Proc. Biol. Soc. Wash. 18: 32, 2 Feb 1905.

= Neotoma stephensi stephensi Goldman, 1905. See Goldman 

(1932b:66); subspecies recognized by Hall (1981:760) and 

Wallace and Jones (1999:611).

Holotype: USNM 117466. [Skin and skull]. Adult female. 

Collected 1 Jul 1902 by F. Stephens. Original number 

4192.

Type Locality: Hualpai (Hualapai) Mountains, altitude 6,300 ft 

(1,920 m), [Mohave County], Arizona.

Neotoma stephensi relicta Goldman

J. Mammal. 13: 66, 9 Feb 1932b.

Subspecies recognized by Hall (1981:760) and Wallace and Jones 

(1999:611).

Holotype: USNM 67780. [Skin and skull]. Adult female. 

Collected 22 Jul 1894 by A. K. Fisher. Original number 

1649.

Type Locality: Keams Canyon, [FC: 80 mi (129 km) north of 

Holbrook], Navajo County, Arizona.

Neotoma tenuicauda Merriam

Proc. Biol. Soc. Wash. 7: 169, 29 Sep 1892a.

= Neotoma mexicana tenuicauda Merriam, 1892. See Hooper 

(1955:22); subspecies recognized by Hall (1981:763) and 

Cornely (1999:609).

Holotype: USNM 33594/45629. Skin and skull. Adult [SL and 

FC: female], not male as in original description. Collected 

13 Apr 1892 by E. W. Nelson. Original number 2446.

Type Locality: North slope Sierra Nevada de Colima, altitude 

3,650 m or [FC: about] 12,000 ft, Jalisco, Mexico.

Neotoma tropicalis Goldman

Proc. Biol. Soc. Wash. 17: 81, 21 Mar 1904.

= Neotoma mexicana tropicalis Goldman, 1904. See Hall 

(1955:331); subspecies recognized by Hall (1981:763) and 

Cornely (1999:609).

Holotype: USNM 68593. [Skin and skull]. Adult male. Collected 

17 Jul 1894 by E. W. Nelson and E. A. Goldman. Original 

number 6468.

Type Locality: Totontepec, altitude 6,500 ft (1,981 m), Oaxaca, 

Mexico.

Neotoma venusta True

Proc. U.S. Natl. Mus. 17: 2, 27 Jun 1894e. Preprint of Proc. U.S. 

Natl. Mus. 17: 354, 15 Nov 1894f.
3 4 • smithsonian contributions to zoology

= Neotoma albigula venusta True, 1894. See Goldman (1910: 

33); subspecies recognized by Hall (1981:753) and Mares 

(1999:598).

Holotype: USNM 21696/36400. Skin and skull. [Lyon and Osgood (1909:106): Adult] male. Collected 30 Nov 1891 by 

F. Stephens. Original number [SL: 1800]. [MC: Purchased 

from C. K. Worthen].

Type Locality: Carrizo Creek, [Imperial County], California.

Remarks: The original number given above does not belong to 

Stevens; it is probably a Worthen collection number.

Teanopus phenax Merriam

Proc. Biol. Soc. Wash. 16: 81, 29 May 1903a.

= Neotoma phenax (Merriam, 1903). See Burt and Barkalow 

(1942:289).

Holotype: USNM 95841. [Skin and skull]. Adult female. Collected 4 Nov 1898 by E. A. Goldman. Original number 

13258.

Type Locality: Camoa, [FC: about 800 ft (244 m)], Rio Mayo, 

Sonora, [SL: Mexico].

Genus Neotomodon Merriam, 1898

Neotomodon alstoni Merriam

Proc. Biol. Soc. Wash. 12: 128, 30 Apr 1898d.

Holotype: USNM 50534. [Skin and skull]. Old male. Collected 

12 Oct 1892 by E. W. Nelson. Original number 3580.

Type Locality: Nahuatzin, altitude 8,500 ft (2,591 m), Michoacan, 

Mexico.

Neotomodon orizabae Merriam

Proc. Biol. Soc. Wash. 12: 129, 30 Apr 1898d.

= Neotomodon alstoni Merriam, 1898. See Davis (1944b:398).

Holotype: USNM 53486. [Skin and skull]. Old male. Collected 

26 Apr 1893 by E. W. Nelson. Original number 4747.

Type Locality: Mount Orizaba, altitude 9,500 ft (2,896 m), 

Puebla, Mexico.

Neotomodon perotensis Merriam

Proc. Biol. Soc. Wash. 12: 129, 30 Apr 1898d.

= Neotomodon alstoni Merriam, 1898. See Davis (1944b:398).

Holotype: USNM 54398. [Skin and skull]. Adult female. Collected 29 May 1893 by E. W. Nelson. Original number 

4897.

Type Locality: Cofre de Perote, altitude 9,500 ft (2,896 m), Veracruz, Mexico.

Genus Nephelomys Weksler, Percequillo, 

and Voss, 2006

Oryzomys pirrensis Goldman

Smithson. Misc. Collect. 60(22): 5, 28 Feb 1913.

= Nephelomys pirrensis (Goldman, 1913). See Weksler et al. 

(2006:18).

Holotype: USNM 178993. [Skin and skull]. Adult male. Collected 29 Apr 1912 by E. A. Goldman. Original number 

21662.

Type Locality: Near head of Rio Limon, mount Pirri, altitude 

4,500 ft (1,372 m), [Darien], eastern Panama.

Genus Nyctomys Saussure, 1860

Nyctomys sumichrasti costaricensis Goldman

J. Wash. Acad. Sci. 27: 422, 15 Oct 1937a.

Subspecies recognized by Hunt et al. (2004:1).

Holotype: USNM 250331. Skin and skull. Adult male. Collected 

12 Apr 1931 by C. F. Underwood. Original number [SL and 

XC: 272]. 26896-X.

Type Locality: San Geronimo de Pirris, hamlet on the main road 

to Pirris before reaching Jabillo, altitude about 100 ft (30 

m), about 2 mi (3.4 km) before the abrupt decent to the 

lowlands of Pozo Axul (Pozo Azul) and about 12 mi (19 

km) inland from Pirris, [Puntarenas], near the west coast of 

Costa Rica.

Nyctomys sumichrasti pallidulus Goldman

J. Wash. Acad. Sci. 27: 420, 15 Oct 1937a.

Subspecies recognized by Hunt et al. (2004:1).

Holotype: USNM 73302. Skin and skull. Adult male. Collected 

13 Jun 1895 by E. W. Nelson and E. A. Goldman. Original 

number 8079.

Type Locality: Santo Domingo, 8 mi (13 km) west of Lagunas on 

the Mexican Railroad, altitude 900 ft (274 m), Isthmus of 

Tehuantepec, Oaxaca, Mexico.

Nyctomys sumichrasti venustulus Goldman

Proc. Biol. Soc. Wash. 29: 155, 6 Sep 1916.

Subspecies recognized by Hunt et al. (2004:1).

Holotype: USNM 33149/45153. Skin and skull. Adult female. 

Collected 10 Feb 1892 by C. W. Richmond. Original 

number 17.

Type Locality: Greytown (San Juan del Norte), [Rio San Juan], 

Nicaragua.

Sitomys (Rhipidomys) decolorus True

Proc. U.S. Natl. Mus. 16: 689, 5 Feb 1894g.

= N[yctomys] s[umichrasti] decolorus (True, 1894). See Goldman (1916:155). Subspecies recognized by Hunt et al. 

(2004:1).

Holotype: USNM 21092/SL: 36129. Skin and skull. [Lyon and 

Osgood (1909:142): Young] adult female. Collected 17 Nov 

1890, not 11 Dec as in original description, by E. Wittkugel. 

Original number [SL: 9].

Type Locality: Rio de las Piedras, Cortez, Honduras.

Remarks: Type not designated as such by number in original description, but the single specimen is referred to by its USNM 

number and becomes the holotype as fixed by monotypy 

(ICZN, 1999: Article 73.1.2).
number 642 • 3 5

Genus Ochrotomys Osgood, 1909

Arvicola nuttalli Harlan

Mon. Am. J. Geol. Nat. Sci. 1: 446, Apr 1832.

= Ochrotomys nuttalli (Harlan, 1832). See Hooper (1958:23); subspecies not recognized by Feldhamer and Linzey (2008:26).

Neotype: USNM 95889 as designated by Packard (1969:401). 

Skin and skull. Adult female. [SL: Collected 6 Nov 1898 by 

A. K. Fisher. Original number 2004].

Type Locality: Lake Drummond, Dismal Swamp, Nansemond 

County (Suffolk City), Virginia, 30 mi (48 km) southwest of 

the original type locality at Norfolk, Virginia.

Peromyscus nuttalli flammeus Goldman

Proc. Biol. Soc. Wash. 54: 190, 8 Dec 1941b.

= Ochrotomys nuttalli (Harlan, 1832). See Hooper (1958:23); subspecies not recognized by Feldhamer and Linzey (2008:26).

Holotype: USNM 170591. Skin and skull. Adult female. Collected 16 Nov 1910 by W. G. Savage. Original number [XC: 

26]. 8221-X.

Type Locality: Delight, Pike County, southwestern Arkansas.

Peromyscus nuttalli lewisi A. H. Howell

J. Mammal. 20: 498, 14 Nov 1939a.

= Ochrotomys nuttalli (Harlan, 1832). See Hooper (1958:23); 

subspecies not recognized by Feldhamer and Linzey 

(2008:26).

Holotype: USNM 262415. Skin and skull. Adult female. Collected 30 Jan 1936 by J. B. Lewis. Original number 15. 

27823-X.

Type Locality: Amelia Court House, Amelia County, Virginia.

Genus Oecomys Thomas, 1906

Oryzomys frontalis Goldman

Smithson. Misc. Collect. 56(36): 6, 19 Feb 1912b.

= Oecomys trinitatis (J. A. Allen and Chapman, 1893). See 

Musser and Carleton (1993:716).

Holotype: USNM 171531. [Skin and skull]. Adult female. Collected 20 Jun 1911 by E. A. Goldman. Original Number 

21172.

Type Locality: Corozal, Canal Zone (Panama), Panama.

Genus Oligoryzomys Bangs, 1900

Oryzomys fulvescens lenis Goldman

Proc. Biol. Soc. Wash. 28: 130, 29 Jun 1915b.

= Oligoryzomys fulvescens lenis (Goldman, 1915). New name 

combination; subspecies recognized by Hall (1981:622) 

under Oryzomys fulvescens.

Holotype: USNM 125941. [Skin and skull]. Adult male. Collected 14 Feb 1903 by E. W. Nelson and E. A. Goldman. 

Original number 15498.

Type Locality: Los Reyes, Michoacan, Mexico.

Oryzomys fulvescens mayensis Goldman

N. Am. Fauna 43: 92, 23 Sep 1918a.

= Oligoryzomys fulvescens mayensis (Goldman, 1918). New 

name combination; subspecies recognized by Hall (1981: 

622) under Oryzomys fulvescens.

Holotype: USNM 107979. [Skin and skull]. Adult male. Collected 5 Jan 1901 by E. W. Nelson and E. A. Goldman. 

Original number 14405.

Type Locality: Apazote, near Yohaltum (Yohaltún), altitude 200 

ft (61 m), Campeche, Mexico.

Genus Ondatra Link, 1795

Fiber macrodon Merriam

Proc. Biol. Soc. Wash. 11: 143, 13 May 1897f.

= Ondatra zibethicus macrodon (Merriam, 1897). See Miller 

(1912a:230); subspecies recognized by Hall (1981:825) and 

Feldhamer (1999:653).

Holotype: USNM 75940. [Skin and skull]. Adult female. Collected 9 Oct 1895 by A. K. Fisher. Original number 1788.

Type Locality: Lake Drummond, Dismal Swamp, [Chesapeake 

City], Virginia.

Fiber spatulatus Osgood

N. Am. Fauna 19: 36, 6 Oct 1900.

= Ondatra zibethicus spatulatus (Osgood, 1900). See Miller 

(1912a:231); subspecies recognized by Hall (1981:827) and 

Feldhamer (1999:653).

Holotype: USNM 98567. [Skin and skull]. Young adult female. 

Collected 3 Jul 1899 by W. H. Osgood. Original number 

552.

Type Locality: Lake Marsh, Northwest Territories (Yukon), 

Canada.

Fiber zibethicus cinnamominus Hollister

Proc. Biol. Soc. Wash. 23: 125, 2 Sep 1910a.

= Ondatra zibethicus cinnamominus (Hollister, 1910). See Miller 

(1912a:232); subspecies recognized by Hall (1981:825) and 

Feldhamer (1999:653).

Holotype: USNM 186518. Skin and skull. Adult male. Collected 

14 Jan 1887 by A. B. Baker. No original number. Merriam 

collection number 3084/3724.

Type Locality: Wakeeney [MCC: Banner], Trego County, Kansas.

Remarks: Published as Merriam collection number 3084/3724; 

cataloged as USNM 186518 on 23 Oct 1911.

Fiber zibethicus hudsonius Preble

N. Am. Fauna 22: 53, 31 Oct 1902a.

= Ondatra zibethicus albus (Sabine, 1823). See Miller (1912a: 

231); subspecies recognized by Hall (1981:824) and Feldhamer (1999:653).

Holotype: USNM 106881. Skin and skull. Adult male. Collected 

9 Aug 1900 by A. E. Preble. Original number [FC: EAP 

3081].
3 6 • smithsonian contributions to zoology

Type Locality: [FC: Hudson Bay], Fort Churchill (Churchill), 

Keewatin (Manitoba), Canada.

Fiber zibethicus mergens Hollister

Proc. Biol. Soc. Wash. 23: 1, 2 Feb 1910b.

= Ondatra zibethicus mergens (Hollister, 1910). See Miller 

(1912a:231); subspecies recognized by Hall (1981:826) and 

Feldhamer (1999:653).

Holotype: USNM 156880. Skin and skull. Adult female. Collected 3 Apr 1908 by S. E. Piper. Original number 143.

Type Locality: [SL and FC: 5 mi (8 km) west] Fallon, [Churchill 

County], Nevada.

Fiber zibethicus ripensis V. Bailey

Proc. Biol. Soc. Wash. 15: 119, 2 Jun 1902a.

= Ondatra zibethicus ripensis (V. Bailey, 1902). See Miller 

(1912a:232); subspecies recognized by Hall (1981:827) and 

Feldhamer (1999:653).

Holotype: USNM 109012. [Skin, skull, and genitalia in alcohol]. 

Adult male. Collected 26 Jul 1901, not 25 Jul as in original 

description, by V. Bailey. Original number 7757.

Type Locality: Carlsbad (Eddy), Pecos River, [Eddy County], 

New Mexico.

Fiber zibethicus zalophus Hollister

Proc. Biol. Soc. Wash. 23: 1, 2 Feb 1910b.

= Ondatra zibethicus zalophus (Hollister, 1910). See Miller 

(1912a:231); subspecies recognized by Hall (1981:827) and 

Feldhamer (1999:653).

Holotype: USNM 131488. Skin and skull. [Age and sex not 

given]. Collected Oct 1903 by A. G. Maddren. Original 

number [XC: 21. 4285-X].

Type Locality: Becharof Lake, [Alaska Peninsula], Alaska.

Ondatra zibethica bernardi Goldman

Proc. Biol. Soc. Wash. 45: 93, 21 Jun 1932c.

= Ondatra zibethicus bernardi Goldman, 1932. See Miller 

and Kellogg (1955:618); subspecies recognized by Hall 

(1981:825) and Feldhamer (1999:653).

Holotype: USNM 250454. [Skin and skull]. Adult male. Collected 

12 Nov 1931 by V. Bailey. Original number A4372. 27010-X.

Type Locality: 4 mi (6km) south of Gadsden, Yuma County, 

Arizona.

Genus Onychomys Baird, 1857

Hesperomys (Onychomys) torridus Coues

Proc. Acad. Nat. Sci. Phila. 26: 183, 15 Dec 1874.

= Onychomys torridus torridus (Coues, 1874). See Hollister 

(1914b:456); subspecies recognized by Hall (1981:732) and 

Riddle (1999a:591).

Holotype: USNM 9886. [Skin without skull]. [Age and sex not 

given]. Collected [MC: 10 Jun 1867] by [SL: E. Palmer]. No 

original number.

 Type Locality: [SL: Camp Grant]. [Graham County], Arizona. 

Type locality restricted by Hoffmeister (1956:280) to along 

the San Pedro River, at Arivaipa Creek, 47 mi (76 km) 

northwest of the present Fort Grant, or 2 mi (3.2 km) south 

of Feldman, Pinal County, Arizona. 

Onychomys fuliginosus Merriam

N. Am. Fauna 3: 59, 11 Sep 1890a.

= Onychomys leucogaster fuliginosus Merriam, 1890. See Hollister (1913b:216); subspecies recognized by Hall (1981:728) 

and Riddle (1999b:590).

Holotype: USNM 17997/24908. Skin and skull. Adult female. 

Collected 27 Sep 1889 by C. H. Merriam and V. Bailey. 

Original number VB 547.

Type Locality: Black Tank lava beds, [FC: 25 mi (40 km) west], 

[SL: Painted Desert], [Miller and Kellogg (1955:516): 12 

mi (19 km) to 15 mi (24 km) north of Dead Man Flat], 

northeast of San Francisco Mountain, [Coconino County, 

Arizona].

Onychomys leucogaster albescens Merriam

Proc. Biol. Soc. Wash. 17: 124, 9 Jun 1904.

Subspecies recognized by Hall (1981:727) and Riddle (1999b: 

590).

Holotype: USNM 50040. [Skin and skull]. Adult female. Collected 12 Dec 1892 by C. P. Streator. Original number 2399.

Type Locality: Samalayuca, [FC: altitude 4,200 ft (1,280 m)], 

Chihuahua, Mexico.

Onychomys leucogaster aldousi Goldman

Proc. Biol. Soc. Wash. 55: 77, 25 Jun 1942a.

= Onychomys leucogaster brevicaudus Merriam, 1891. See Riddle and Choate (1986:248); subspecies recognized by Riddle 

(1999b:590).

Holotype: USNM 266502. Skin and skull. Adult male. Collected 

18 Apr 1941 by C. M. Aldous. Original number [SL: 133]. 

30165-X.

Type Locality: Desert Range Experiment Station, 50 mi (80 km) 

west of Milford, Millard (Beaver) County, Utah.

Onychomys leucogaster breviauritus Hollister

Proc. Biol. Soc. Wash. 26: 216, 20 Dec 1913b.

= Onychomys leucogaster arcticeps Rhoads, 1898. See Engstrom 

and Choate (1979:737); subspecies recognized by Riddle 

(1999b:590).

Holotype: USNM 19144/34872. Skin and skull. Adult female. Collected 16 Feb 1890 by J. C. Merrill. Original 

number 18.

Type Locality: Fort Reno, [Canadian County], Oklahoma.

Onychomys leucogaster brevicaudus Merriam

N. Am. Fauna 5: 52, 30 Jul 1891a.

Subspecies recognized by Hall (1981:727) and Riddle (1999b: 

590).
number 642 • 3 7

Holotype: USNM 23086/30532. Skin and skull. Adult male. 

Collected 15 Jul 1890 by V. Bailey and B. H. Dutcher. Original number [FC: VB 1442].

Type Locality: Blackfoot, [Bingham County], Idaho.

Onychomys leucogaster capitulatus Hollister

Proc. Biol. Soc. Wash. 26: 215, 20 Dec 1913b.

= Onychomys leucogaster fuliginosus Merriam, 1890. See Van 

Cura and Hoffmeister (1966:622); subspecies recognized by 

Hall (1981:728) and Riddle (1999b:590).

Holotype: USNM 202612. Skin and skull. Adult male. Collected 

26 Sep 1913 by E. A. Goldman. Original number 22234.

Type Locality: Lower end of Prospect Valley, altitude 4,500 ft 

(1,372 m), [Miller and Kellogg (1955:516): about 25 mi (40 

km) west-southwest of Supai], Hualpai (Hualapai) Indian 

Reservation, Grand Canyon, [Coconino County], Arizona.

Onychomys leucogaster melanophrys Merriam

N. Am. Fauna 2: 2, 30 Oct 1889a.

Subspecies recognized by Hall (1981:728) and Riddle (1999b:590).

Holotype: USNM 186477. Skin and skull. Adult male. Collected 

22 Dec 1888 by V. Bailey. Original number [SL: 458]. Merriam collection number 5199/5894.

Type Locality: Kanab, [Kane County], Utah.

Remarks: Published as Merriam collection number 5199/5894; 

assigned USNM 186477 on 18 Oct 1911.

Onychomys leucogaster utahensis Goldman

J. Mammal. 20: 354, 14 Aug 1939.

= Onychomys leucogaster brevicaudus Merriam, 1891. See Riddle and Choate (1986:248); subspecies recognized by Riddle 

(1999b:590).

Holotype: USNM 263955. Skin and skull. Adult female. Collected 22 Jun 1938 by W. H. Marshall. Original number 96.

Type Locality: South end of Stansbury Island, Great Salt Lake, 

altitude 4,250 ft (1,295 m), [Tooele County], Utah.

Onychomys longicaudus Merriam

N. Am. Fauna 2: 2, 30 Oct 1889a.

= Onychomys torridus longicaudus Merriam, 1889. See Merriam (1904:123); subspecies recognized by Hall (1981:731) 

and Riddle (1999a:591).

Holotype: USNM 186476. Skin and skull. Adult male. Collected 

4 Jan 1889 by V. Bailey. Original number [SL: 476]. Merriam collection number 5201/5896.

Type Locality: Saint George, [Washington County], Utah.

Remarks: Published as Merriam collection number 5201/5896; 

assigned USNM 186476 on 18 Oct 1911.

Onychomys longipes Merriam

N. Am. Fauna 2: 1, 30 Oct 1889a.

= Onychomys leucogaster longipes Merriam, 1889. See Hollister 

(1913b:216); subspecies recognized by Hall (1981:728) and 

Riddle (1999b:590).

Holotype: USNM 186478. Skin and skull. Adult female. Collected 11 Mar 1887 by W. Lloyd. Original number [MCC: 

3]. Merriam collection number 3207/3839.

Type Locality: Concho County, Texas.

Remarks: Published as Merriam collection number 3207/3839; 

cataloged as USNM 186478 on18 Oct 1911.

Onychomys melanophrys pallescens Merriam

N. Am. Fauna 3: 61, 11 Sep 1890a.

= Onychomys leucogaster pallescens Merriam, 1890. See J. A. 

Allen (1895a:225); subspecies recognized by Hall (1981: 

730) and Riddle (1999b:590).

Holotype: USNM 186479. Skin and skull. Adult male. Collected 

18 May 1888 by [MCC: J. Sullivan. Original number 11]. 

Merriam collection number 4252/4983.

Type Locality: Moki [Hoffmeister (1986:381): Hopi] Pueblos, 

[MCC: Keams Canyon], Apache (Navajo) County, Arizona.

Remarks: Published as Merriam collection number 4252/4983; 

cataloged as USNM 186479 on 18 Oct 1911.

Onychomys torridus arenicola Mearns

Proc. U.S. Natl. Mus. 19: 3, 25 May 1896a. Preprint of Proc. 

U.S. Natl. Mus. 19: 139, 21 Dec 1896b.

= Onychomys arenicola Mearns, 1896. See Hinesley (1979:119).

Holotype: USNM 20081/35483. Skin and skull. Adult male. 

Collected 29 Feb 1892 by E. A. Mearns and F. X. Holzner. 

Original number 1528.

Type Locality: Rio Grande, about 6 mi (10 km) above El Paso, 

[El Paso County], Texas.

Onychomys torridus canus Merriam

Proc. Biol. Soc. Wash. 17: 124, 9 Jun 1904.

= Onychomys arenicola Mearns, 1896. See Riddle (1999c:588).

Holotype: USNM 90843. [Skin and skull]. Adult female. Collected 23 Aug 1897 by E. W. Nelson and E. A. Goldman. 

Original number 11574.

Type Locality: San Juan Capistrano, [FC: altitude 3,700 ft 

(1,128 m)], Zacatecas, Mexico.

Onychomys torridus clarus Hollister

Proc. Biol. Soc. Wash. 26: 215, 20 Dec 1913b.

Subspecies recognized by Hall (1981:731) and Riddle (1999a:591).

Holotype: USNM 25453/32867. Skin and skull. Adult male. 

Collected 30 Dec 1890 by V. Bailey. Original number 2314.

Type Locality: Keeler, [Miller (1924:304): east shore of Owens 

Lake], [FC: about 25 ft (8 m) above lake], Inyo County, 

California.

Onychomys torridus perpallidus Mearns

Proc. U.S. Natl. Mus. 19: 43, 25 May 1896a. Preprint of Proc. 

U.S. Natl. Mus. 19: 140, 21 Dec 1896b.

= Onychomys torridus torridus (Coues, 1874). See Van Cura 

and Hoffmeister (1966:626); subspecies recognized by Hall 

(1981:732) and Riddle (1999a:591).
3 8 • smithsonian contributions to zoology

Holotype: USNM 60174. Skin and skull. Adult female. Collected 27 Mar 1894 by E. A. Mearns and F. X. Holzner. 

Original number 3301.

Type Locality: Left bank of the Colorado River, at Monument 

204, Mexican Boundary Line, [Miller and Rehn (1901:67): 

Yuma County, Arizona].

Onychomys torridus surrufus Hollister

Proc. U.S. Natl. Mus. 47: 472, 29 Oct 1914b.

= Onychomys arenicola Mearns, 1896. See Riddle (1999c:588).

Holotype: USNM 93839. Skin and skull. Adult female. Collected 

8 Jun 1898 by E. W. Nelson and E. A. Goldman. Original 

number 12484.

Type Locality: Miquihuana, [FC: altitude about 5,000 ft 

(1,524 m)], Tamaulipas, Mexico.

Onychomys torridus tularensis Merriam

Proc. Biol. Soc. Wash. 17: 123, 9 Jun 1904.

Subspecies recognized by Hall (1981:732) and Riddle (1999a:591).

Holotype: USNM 29711/41786. Skin and skull. Adult female. 

Collected 19 Jul 1891 by A. K. Fisher. Original number 792.

Type Locality: Bakersfield, Kern County, California.

Onychomys torridus yakiensis Merriam

Proc. Biol. Soc. Wash. 17: 124, 9 Jun 1904.

Subspecies recognized by Hall (1981:732) and Riddle (1999a:591).

Holotype: USNM 95855. [Skin and skull]. Adult female. Collected 28 Oct 1898 by E. A. Goldman. Original number 

[FC: Nelson and Goldman] 13158.

Type Locality: Camoa, [FC: altitude 800 ft (244 m)], Rio Mayo, 

southern Sonora, Mexico.

Genus Oryzomys Baird, 1857

Oryzomys albiventer Merriam

Proc. Wash. Acad. Sci. 3: 279, 26 Jul 1901b.

See Carleton and Arroyo-Cabrales (2009:117) for current use.

Holotype: USNM 82236. [Skin and skull]. Adult male. Collected 

6 Feb 1897 by E. W. Nelson and E. A. Goldman. Original 

number 10478.

Type Locality: Ameca, [FC: 4,000 ft (1,219 m)], Jalisco, Mexico.

Oryzomys alfaroi dariensis Goldman

Proc. Biol. Soc. Wash. 28: 128, 29 Jun 1915b.

Subspecies recognized by Hall (1981:616).

Holotype: USNM 178660. [Skin and skull]. Adult female. Collected 4 Mar 1912 by E. A. Goldman. Original number 21453.

Type Locality: Cana (Santa Cruz de Cana), altitude 2,000 ft 

(610 m), [Miller and Kellogg (1955:435): Upper Rio Tuyra], 

[Darien], Panama.

Oryzomys angusticeps Merriam

Proc. Wash. Acad. Sci. 3: 292, 26 Jul 1901b.

= Oryzomys rhabdops Merriam, 1901. See Musser and Carleton 

(1993:724).

Holotype: USNM 76816. [Skin and skull]. Adult male. Collected 

22 Jan 1896 by E. W. Nelson and E. A. Goldman. Original 

number 9190.

Type Locality: Volcan Santa Maria, altitude 9,000 ft (2,743 m), 

[Quezaltenango], Guatemala.

Oryzomys argentatus Spitzer and Lazell

J. Mammal. 59: 787, 24 Nov 1978.

= Oryzomys palustris natator Chapman, 1893. See Humphrey 

and Setzer (1989:565); subspecies recognized by Rose and 

Dueser (1999:555).

Holotype: USNM 514995. Skin, skull, and skeleton. Adult female. Collected 8 Feb 1973 by N. C. Spitzer. Original number NS 162.

Type Locality: Cudjoe Key, [SL: 2.3 mi (3.7 km) west of Kemp 

Channel Bridge], Monroe County, Florida.

Oryzomys chapmani caudatus Merriam

Proc. Wash. Acad. Sci. 3: 289, 26 Jul 1901b.

= Oryzomys chapmani Thomas, 1898. See Musser and Carleton 

(1993:721).

Holotype: USNM 68641. [Skin and skull]. Adult male. Collected 

31 Jul 1894 by E. W. Nelson and E. A. Goldman. Original 

number 6516.

Type Locality: Comaltepec, [FC: altitude 3,500 ft (1,067 m)], 

Oaxaca, Mexico.

Remarks: Hall (1981:615) listed this taxon as the monotypic 

Oryzomys caudatus.

Oryzomys chapmani dilutior Merriam

Proc. Wash. Acad. Sci. 3: 290, 26 Jul 1901b.

= Oryzomys chapmani dilutior Merriam, 1901. See Musser and 

Carleton (1993:721); subspecies recognized by Hall (1981: 

616) under O. alfaroi.

Holotype: USNM 93124. [Skin and skull]. Adult male. Collected 

10 Jan 1898 by E. W. Nelson and E. A. Goldman. Original 

number 12040.

Type Locality: Huauchinango, [FC: altitude 5,000 ft (1,524 m)], 

Puebla, Mexico.

Oryzomys chapmani saturatior Merriam

Proc. Wash. Acad. Sci. 3: 290, 26 Jul 1901b.

= Oryzomys saturatior saturatior Merriam, 1901. See Musser 

and Carleton (1993:725); subspecies recognized by Hall 

(1981:616) under O. alfaroi.

Holotype: USNM 76183. [Skin and skull]. Adult female. Collected 23 Oct 1895 by E. W. Nelson and E. A. Goldman. 

Original number 8574.

Type Locality: Tumbala, [FC: altitude 5,000 ft (1,524 m)], Chiapas, [SL: Mexico].

Oryzomys couesi regillus Goldman

Proc. Biol. Soc. Wash. 28: 129, 29 Jun 1915b.

Subspecies recognized by Hall (1981:612) under O. palustris.
number 642 • 3 9

Holotype: USNM 125945. [Skin and skull]. Adult male. Collected 17 Feb 1903 by E. W. Nelson and E. A. Goldman. 

Original number 15962.

Type Locality: Los Reyes, Michoacan, Mexico.

Oryzomys cozumelae Merriam

Proc. Biol. Soc. Wash. 14: 103, 19 Jul 1901c.

= Oryzomys couesi cozumelae Merriam, 1901. See Musser and 

Carleton (1993:721); subspecies recognized by Vega et al. 

(2007:210).

Holotype: USNM 108462. [Skin, skull, and baculum]. Adult 

male. Collected 8 Apr 1901 by E. W. Nelson and E. A. Goldman. Original number 14666.

Type Locality: Cozumel Island, Yucatan (Quintana Roo), [SL: 

Mexico].

Oryzomys crinitus Merriam

Proc. Wash. Acad. Sci. 3: 281, 26 Jul 1901b.

= Oryzomys couesi crinitus Merriam, 1901. See Goldman 

(1918a:36); subspecies recognized by Hall (1981:610) 

under O. palustris.

Holotype: USNM 50182. [Skin and skull]. Adult male. Collected 

30 Nov 1892 by E. W. Nelson and E. A. Goldman. Original 

number [FC: EWN] 3905.

Type Locality: Tlalpam, [FC: altitude 7,600 ft (2,316 m)], Federal District (Distrito Federal), Mexico.

Oryzomys crinitus aztecus Merriam

Proc. Wash. Acad. Sci. 3: 282, 26 Jul 1901b.

= Oryzomys couesi aztecus Merriam 1901. See Goldman 

(1918a:35); subspecies recognized by Hall (1981:609) 

under O. palustris.

Holotype: USNM 51173. [Skin and skull]. Adult male. Collected 

16 Jan 1893 by E. W. Nelson and E. A. Goldman. Original 

number [FC: EWN] 4290.

Type Locality: Yautepec, [FC: altitude 4,100 ft (1,250 m)], Morelos, Mexico.

Oryzomys gatunensis Goldman

Smithson. Misc. Collect. 56(36): 7, 19 Feb 1912b.

= Oryzomys couesi gatunensis Goldman, 1912. See Musser and 

Carleton (1993:721); subspecies recognized by Hall (1981: 

610) under O. palustris.

Holotype: USNM 171034. Skin and skull. Young male. Collected 

7 Mar 1911 by E. A. Goldman. Original number 20967.

Type Locality: Gatun, Canal Zone (Colon), Panama.

Oryzomys goldmani Merriam

Proc. Wash. Acad. Sci. 3: 288, 26 Jul 1901b.

= Oryzomys couesi couesi (Alston, 1877). See Goldman (1918a: 

30); subspecies recognized by Hall (1981:609) under 

O. palustris.

Holotype: USNM 78110. [Skin and skull]. Adult female. Collected 11 Apr 1896 by E. W. Nelson and E. A. Goldman. 

Original number 9511.

Type Locality: Coatzacoalcos, [FC: altitude 50 ft (15 m)], Veracruz, Mexico.

Oryzomys guerrerensis Goldman

Proc. Biol. Soc. Wash. 28: 127, 29 Jun 1915b.

= Oryzomys chapmani guerrerensis Goldman, 1915. See Musser 

and Carleton (1993:721); subspecies recognized by Hall 

(1981:616) under O. alfaroi.

Holotype: USNM 127517. Skin and skull. Adult male. Collected 

20 May 1903 by E. W. Nelson and E. A. Goldman. Original 

number 16454.

Type Locality: Omilteme, [Hall and Kelson (1959:562): 8,000 ft 

(2,438 m)], Guerrero, Mexico.

Oryzomys hylocetes Merriam

Proc. Wash. Acad. Sci. 3: 291, 26 Jul 1901b.

= Oryzomys saturatior hylocetes Merriam, 1901. See Musser 

and Carleton (1993:725); subspecies recognized by Hall 

(1981:616) under O. alfaroi.

Holotype: USNM 77605. [Skin and skull]. Old male. Collected 

14 Feb 1896 by E. W. Nelson and E. A. Goldman. Original 

number 9306.

Type Locality: Chicharras, [FC: altitude 3,400 ft (1,036 m)], 

Chiapas, Mexico.

Oryzomys jalapae rufinus Merriam

Proc. Wash. Acad. Sci. 3: 285, 26 Jul 1901b.

= Oryzomys couesi couesi (Alston, 1877). See Goldman (1918a: 

29); subspecies recognized by Hall (1981:609) under 

O. palustris.

Holotype: USNM 65499. [Skin and skull]. Adult female. Collected 27 Apr 1894 by E. W. Nelson and E. A. Goldman. 

Original number 6112.

Type Locality: Catemaco, [FC: altitude 1,000 ft (305 m)], Veracruz, Mexico.

Oryzomys melanotis colimensis Goldman

N. Am. Fauna 43: 51, 23 Sep 1918a.

Subspecies recognized by Hall (1981:614).

Holotype: USNM 33289/45317. Skin and skull. Adult female. 

Collected 2 Mar 1892 by E. W. Nelson. Original number 

1987.

Type Locality: Armeria, altitude about 100 ft (30 m), Colima, 

Mexico.

Oryzomys mexicanus peragrus Merriam

Proc. Wash. Acad. Sci. 3: 283, 26 Jul 1901b.

= Oryzomys couesi peragrus Merriam, 1901. See Goldman 

(1918a:39). Subspecies recognized by Hall (1981:611) 

under O. palustris.

Holotype: USNM 82119, not 82111 as in original description. 

[Skin and skull]. Young adult male. Collected 8 Jan 1897 by 

E. W. Nelson and E. A. Goldman. Original number 10398.

Type Locality: Rio Verde, [FC: altitude about 3,000 ft (914 m)], 

San Luis Potosi, Mexico.
4 0 • smithsonian contributions to zoology

Oryzomys natator floridanus Merriam

Proc. Wash. Acad. Sci. 3: 277, 26 Jul 1901b.

= Oryzomys palustris natator Chapman, 1893. See Humphrey 

and Setzer (1989:565), who considered coloratus, including 

floridanus, as a subspecies of O. palustris natator.

Holotype: USNM 71349. [Skin and skull]. Adult male. Collected 

29 Mar 1895 by J. A. Loring. Original number 2819.

Type Locality: Everglade [FC: Post Office] (Everglades City), 

Monroe (Collier) County, Florida.

Oryzomys nelsoni Merriam

Proc. Biol. Soc. Wash. 12: 15, 27 Jan 1898e.

Holotype: USNM 89200. [Skin and skull]. Adult male. Collected 

13 May 1897 by E. W. Nelson and E. A. Goldman. Original 

number 11022.

Type Locality: Maria Madre Island, [FC: altitude 800 ft (244 m)], 

Tres Marias Islands, [Nayarit], Mexico.

Oryzomys palatinus Merriam

Proc. Wash. Acad. Sci. 3: 290, 26 Jul 1901b.

= Oryzomys alfaroi palatinus Merriam, 1901. See Goldman 

(1918a:65); subspecies recognized by Hall (1981:616).

Holotype: USNM 99977. [Skin and skull]. Adult female. Collected 1 Apr 1900 by E. W. Nelson and E. A. Goldman. 

Original number 14080.

Type Locality: Teapa, Tabasco, Mexico.

Oryzomys palustris planirostris Hamilton

Proc. Biol. Soc. Wash. 68: 83, 3 Aug 1955.

= Oryzomys palustris natator Chapman, 1893. See Humphrey 

and Setzer (1989:565), who considered planirostris a junior 

synonym of O. palustris natator.

Holotype: USNM 514829. Skin and skull, but skull not received 

from Cornell. Adult female. Collected 11 Mar 1954 by W. J. 

Hamilton Jr. Original number 4344. Cornell University 

Mammal Collection number 8090.

Type Locality: 1 mi (1.6 km) west of third bridge that spans 

Matachla (Matlacha) Pass, Pine Island, Lee County, Florida.

Remarks: Published as Cornell University Mammal Collection 

number 8090; assigned USNM 514829 on 25 Aug 1977.

Oryzomys rhabdops Merriam

Proc. Wash. Acad. Sci. 3: 291, 26 Jul 1901b.

Holotype: USNM 76813. [Skin and skull]. Adult male. Collected 

15 Jan 1896 by E. [W.] Nelson and E. A. Goldman. Original 

number 9135.

Type Locality: Calel, altitude 10,000 ft (3,048 m), [Quetzaltenango], Guatemala.

Oryzomys richmondi Merriam

Proc. Wash. Acad. Sci. 3: 284, 26 Jul 1901b.

= Oryzomys couesi richmondi Merriam, 1901. See Goldman 

(1918a:32); subspecies recognized by Hall (1981:612) 

under O. palustris.

Holotype: USNM 36340/48705. Skin and skull. Adult male. Collected 21 Jun 1892 by C. W. Richmond. Original number 63.

Type Locality: Escondido River, 50 mi (80 km) above Bluefields, 

[Atlantico Sur], Nicaragua.

Oryzomys rostratus Merriam

Proc. Wash. Acad. Sci. 3: 293, 26 Jul 1901b.

= Oryzomys rostratus rostratus Merriam, 1901. See Goldman 

(1918a:53); subspecies recognized by Hall (1981:614) 

under O. melanotis.

Holotype: USNM 93112. [Skin and skull]. Old male. Collected 

5 Feb 1898 by E. W. Nelson and E. A. Goldman. Original 

number 12130.

Type Locality: [FC: Near] Metlaltoyuca, [FC: about 800 ft 

(244 m)], Puebla, Mexico.

Oryzomys rostratus megadon Merriam

Proc. Wash. Acad. Sci. 3: 294, 26 Jul 1901b.

Subspecies recognized by Hall (1981:614) under O. melanotis.

Holotype: USNM 99978. [Skin and skull]. Old male. Collected 

24 Mar 1900 by E. W. Nelson and E. A. Goldman. Original 

number 14062.

Type Locality: Teapa, Tabasco, Mexico.

Oryzomys rufus Merriam

Proc. Wash. Acad. Sci. 3: 287, 26 Jul 1901b.

= Oryzomys couesi mexicanus J. A. Allen, 1897. See Goldman 

(1918a:33); subspecies also recognized by Carleton and 

Arroyo-Cabrales (2009:118), with rufus in full synonymy.

Holotype: USNM 91404. [Skin and skull]. Old female. Collected 

20 Jun 1897 by E. W. Nelson and E. A. Goldman. Original 

number 11232.

Type Locality: Santiago, [FC: altitude 200 ft (61 m)], Tepec 

(Nayarit), Mexico.

Oryzomys teapensis Merriam

Proc. Wash. Acad. Sci. 3: 286, 26 Jul 1901b.

= Oryzomys couesi couesi (Alston, 1877). See Goldman (1918a: 

29); subspecies, including teapensis as a junior synonym, 

recognized by Hall (1981:609) under O. palustris.

Holotype: USNM 99973. [Skin and skull]. Young adult male. 

Collected 4 Apr 1900 by E. W. Nelson and E. A. Goldman. 

Original number 14100. 

Type Locality: Teapa, Tabasco, Mexico.

Oryzomys yucatanensis Merriam

Proc. Wash. Acad. Sci. 3: 294, 26 Jul 1901b.

= Oryzomys rostratus yucatanensis Merriam, 1901. See Goldman (1918a:55); subspecies recognized by Hall (1981:615) 

under O. melanotis.

Holotype: USNM 108139. [Skin and skull]. Young adult male. 

Collected 9 Feb 1901 by E. W. Nelson and E. A. Goldman. 

Original number 14527.

Type Locality: Chichenitza (Chichén Itzá), Yucatan, Mexico.
number 642 • 4 1

Oryzomys zygomaticus Merriam

Proc. Wash. Acad. Sci. 3: 285, 26 Jul 1901b.

= Oryzomys couesi zygomaticus Merriam, 1901. See Goldman (1918a:32); subspecies recognized by Hall (1981:612) 

under O. palustris.

Holotype: USNM 76794. [Skin and skull]. Adult male. Collected 

15 Dec 1895 by E. W. Nelson and E. A. Goldman. Original 

number 8798.

Type Locality: Nenton, [FC: 3,000 ft (914 m)], [Huehuetenango], 

Guatemala.

Genus Osgoodomys Hooper and Musser, 1964

Peromyscus banderanus vicinior Osgood

Proc. Biol. Soc. Wash. 17: 68, 21 Mar 1904a.

= Osgoodomys banderanus vicinior (Osgood, 1904). See Carleton (1980:123); subspecies recognized by Hall (1981:721) 

under Peromyscus banderanus.

Holotype: USNM 126503. [Skin and skull]. Adult male. Collected 23 Mar 1903 by E. W. Nelson and E. A. Goldman. 

Original number [SL: 16216].

Type Locality: La Salada, Michoacan, Mexico.

Genus Ototylomys Merriam, 1901

Ototylomys phyllotis Merriam

Proc. Wash. Acad. Sci. 3: 562, 29 Nov 1901d.

= Ototylomys phyllotis phyllotis Merriam, 1901. See Hayman and 

Holt (1941:374); subspecies recognized by Lawlor (1982:1).

Holotype: USNM 108099. [Skin and skull]. Old male. Collected 

17 Feb 1901 by E. W. Nelson and E. A. Goldman. Original 

number 14551.

Type Locality: Tunkas, Yucatan, [SL: Mexico].

Ototylomys phyllotis phaeus Merriam

Proc. Wash. Acad. Sci. 3: 563, 29 Nov 1901d.

= Ototylomys phyllotis phyllotis Merriam, 1901. See Lawlor 

(1969:36); subspecies recognized by Lawlor (1982:1).

Holotype: USNM 107940. [Skin and skull]. Adult female. Collected 28 Dec 1900 by E. W. Nelson and E. A. Goldman. 

Original number 14369.

Type Locality: Apazote, near Yohaltun, Campeche, [SL: Mexico].

Genus Peromyscus Gloger, 1841

Hesperomys (Vesperimus) affinis J. A. Allen

Proc. U.S. Natl. Mus. 14: 195, 24 Jul 1891.

= Peromyscus leucopus affinis (J. A. Allen, 1891). See Osgood 

(1909a:133); subspecies recognized by Hall (1981:685) and 

Lackey (1999:574).

Holotype: USNM 9382, not 7382 as in original description/8665. 

Skin and skull. Adult female. Collected 30 Oct 1868 by F. 

Sumichrast. Original number [SL: 23].

Type Locality: Barrio, Tehuantepec (Oaxaca), Mexico.

Hesperomys (Vesperimus) anthonyi Merriam

Proc. Biol. Soc. Wash. 4: 5, 15 Apr 1887b.

= Peromyscus eremicus anthonyi (Merriam, 1887). See Mearns 

(1907:438); subspecies recognized by Hall (1981:657) and 

Caire (1999:568).

Holotype: USNM 186480. Skin and skull. Adult male. Collected 

10 May 1886 by A. W. Anthony. Original number [MCC: 

87]. Merriam collection number 2333/2841.

Type Locality: Camp Apache, [Osgood (1909:249): Big Hachita 

(Hachit) Mountains], Grant (Hidalgo) County, about 

31°20′, extreme southwestern corner of the Territory and 

only about 4 mi (6 km) from the Mexican boundary, [Mearns 

(1907:438): near Monument No. 40], New Mexico.

Remarks: Published as Merriam collection number 2333/2841; 

assigned USNM 186480 on 18 Oct 1911.

[Hesperomys] articus [sic] Coues

Monogr. N. Am. Rodentia p. 61, 1877.

= Peromyscus maniculatus maniculatus (Wagner, 1845). See Miller 

(1912a:142); subspecies recognized by Hall (1981:679).

Holotype: USNM 3924. [Skin, skull inside]. [Age and sex not 

given]. Collected on unknown date by H. de Saussure. Original number [SL: 2]. [MC: Cataloged 14 May 1860].

Type Locality: Labrador, [Newfoundland, Canada].

Hesperomys austerus Baird

Proc. Acad. Nat. Sci. Phila. 7: 336, reported favorably for publication 24 Apr 1855b. Receipt acknowledged 15 Jun 1855 

by the American Philosophical Society.

= Peromyscus maniculatus austerus (Baird, 1855). See Osgood 

(1909a:63); subspecies recognized by Hall (1981:671).

Lectotype: USNM 364/1278, as designated by Poole and Schantz 

(1942:314). Skin, which cannot be found, and skull. [Lyon 

and Osgood (1909:125): Young adult], sex not given. Collected by G. Suckley [MC: probably 20 Jan 1854]. Original 

number [MC: 5].

Type Locality: [Osgood (1909a:63): Old] Fort Steilacoom, Puget 

Sound, [Pierce County, Washington]. Baird (1857:708) located Fort Steilacoom “near the head of Puget’s Sound, on 

the east side, Latitude 47°10′ Longitude 122°25′.”

Remarks: Poole and Schantz (1942:314, 315) gave a lengthy explanation for their selection of the lectotype that follows 

closely that given by Lyon and Osgood (1909:125); the latter, however, did not formally designate a lectotype. The 

skin, USNM 364, cannot be found and is marked in the 

museum catalog as not in the collection on 17 Nov 1916.

[Hesperomys] bairdii Coues

Monogr. N. Am. Rodentia p. 61, 1877.

= Peromyscus maniculatus maniculatus (Wagner, 1845). See Miller 

(1912a:142); subspecies recognized by Hall (1981:679).

Holotype: USNM 3925. [Skin, skull inside]. [Age and sex not 

given]. Collected on unknown date by H. de Saussure. No 

original number. [MC: Cataloged 14 May 1860].
4 2 • smithsonian contributions to zoology

Type Locality: Labrador, [Newfoundland, Canada].

Remarks: This specimen cannot be found. A note in the museum 

catalog indicates it has been missing at least since 1932.

Hesperomys boylii Baird

Proc. Acad. Nat. Sci. Phila. 7: 335, reported favorably for publication 24 Apr 1855b. Receipt acknowledged 15 Jun 1855 

by the American Philosophical Society.

= Peromyscus boylii boylii (Baird, 1855). See Mearns (1896a:3); subspecies recognized by Kalcounis-Ruppell and Spoon (2009:2).

Holotype: USNM 356/1270. [Mounted skin, skull, and tail vertebrae. Age and sex not given]. Collected by C. C. Boyle, 

[date unknown]. [MC: Skin cataloged 4 Jan 1855; skull 20 

Dec 1854].

Type Locality: Middle fork of the American River, [Osgood 

(1909a:142): near present town of Auburn, El Dorado 

County], California.

Remarks: Type not designated by number in original description. 

The type locality and collector are given, however, and Baird 

(1857:471) stated that “the specimen upon which this species was founded” was collected at the locality given above 

by Boyle. Baird (1857:471) lists three specimens, only one 

of which, USNM 356/1270, is from the type locality and 

was collected by Boyle.

Hesp[eromys] campestris LeConte

Proc. Acad. Nat. Sci. Phila. 6: 413, reported favorably for publication 25 Oct 1853. Receipt acknowledged 31 Dec 1853 by 

the Smithsonian Institution.

= Peromyscus leucopus noveboracensis (Fischer, 1829). See Osgood (1909a:117); subspecies recognized by Shipp-Pennock 

et al. (2005:810).

Lectotype: USNM 4726 as designated by Coues (1877:51, 62). 

Skin, [no record of a skull]. Young, [sex unknown]. Collected by J. E. LeConte, [date unknown. Cataloged 13 Apr 

1861].

Type Locality: New Jersey.

Remarks: Type not designated by number in original description. 

See Lyon and Osgood (1909:126) or Poole and Schantz 

(1942:315) for additional information about selection of 

the lectotype.

H[esperomys] cognatus LeConte

Proc. Acad. Nat. Sci. Phila. 7: 442, reported favorably for publication 25 Dec 1855. Receipt acknowledged, day and month 

unknown, 1856 by the Smithsonian Institution.

= Peromyscus gossypinus gossypinus (LeConte, 1853). See Osgood (1909a:136); subspecies recognized by Hall (1981:689) 

and Linzey (1999a:570).

Syntypes: 4708 and 4709 as designated by Coues (1877:51). 

[Skins with skulls inside. Age and sex not given]. [MC: Collected by J. E. LeConte], date unknown. [MC: Cataloged 13 

Apr 1861].

Type Locality: Georgia and South Carolina.

Hesperomys crinitus Merriam

N. Am. Fauna 5: 53, 30 Jul 1891a.

= Peromyscus crinitus crinitus (Merriam, 1891). See Hall and 

Hoffmeister (1942:56); subspecies recognized by Hall 

(1981:691) and Armstrong (1999a:567).

Holotype: USNM 24255/31659. Skin and skull. Adult male. 

Collected 10 Oct 1890 by C. H. Merriam and V. Bailey. 

Original number [FC: VB] 1945.

Type Locality: Shoshone Falls, Snake River, Idaho. Restricted to 

Shoshone Falls, north side of Snake River, Jerome County, 

Idaho, by Davis (1939:286).

Hesperomys eremicus Baird

Mammals Rep. Explor. Surv. 8(1): 479, 1857.

= Peromyscus eremicus eremicus (Baird, 1857). See J. A. Allen 

(1895a:226); subspecies recognized by Hall (1981:657) and 

Caire (1999:568).

Lectotype: USNM 2575/38003 as designated by Osgood 

(1909a:241). Skin in alcohol, [skull removed]. [Lyon and 

Osgood (1909:129): Female], age not given. Collected in 

1853, collector unknown. Received from G. H. Thomas.

Type Locality: [Poole and Schantz (1942:317): Old] Fort 

Yuma, [Imperial County], California, [Poole and Schantz 

(1942:317): on west side of Colorado River, opposite Yuma, 

Arizona]. [Baird (1857:709) located Fort Yuma “at mouth 

of Gila river, Altitude 355 ft (108 m). Latitude 32°32′ Longitude 114°36′”].

Hesperomys gambelii Baird

Mammals Rep. Explor. Surv. 8(1): 464, 1857.

= Peromyscus maniculatus gambelii (Baird, 1857). See Osgood 

(1909a:182); subspecies recognized by Hall (1981:673).

Syntypes: USNM 368/1282. Mounted skin with skull and [tail 

vertebrae]. Age and sex not given.

USNM 369/1283. Mounted skin, cannot be found, with skull 

and [tail vertebrae]. Age and sex not given. Collected on 

unknown date by W. P. Trowbridge. [MC: Skins cataloged 4 

Jan 1853; skulls 6 Jan 1855].

Type Locality: Monterey, [Monterey County], California.

Remarks: Syntypes designated by J. A. Allen (1893:190). The museum catalog indicates that both specimens were originally 

in alcohol and that USNM 369 was not found 15 Dec 1916.

H[esperomys] gracilis LeConte

Proc. Acad. Nat. Sci. Phila. 7: 442, reported favorably for publication 25 Dec 1855. Receipt acknowledged, day and month 

unknown, 1856 by the Smithsonian Institution.

= Peromyscus maniculatus gracilis (LeConte, 1855). See Osgood 

(1909a:42); subspecies recognized by Taylor and Hoffman 

(2010:404).

Holotype: USNM 10292/38002. [Skin and skull. Age and sex 

not given. Date collected and collector not given]. [MC: 

Skin cataloged 15 Oct 1872; skull 26 May 1902].

Type Locality: Michigan.
number 642 • 4 3

Remarks: Osgood (1909a:43) and Lyon and Osgood (1909:131) 

gave reasons for considering this specimen the type.

Hesperomys leucopus rufinus Merriam

N. Am. Fauna 3: 65, 11 Sep 1890a.

= Peromyscus maniculatus rufinus (Merriam, 1890). See Osgood 

(1909a:72); subspecies recognized by Hall (1981:682).

Holotype: USNM 17646/24582. Skin and skull. Adult female. 

Collected 22 Aug 1889 by C. H. Merriam and V. Bailey. 

Original number [FC: VB] 401.

Type Locality: San Francisco Mountain, altitude 9,000 ft 

(2,743 m), [Coconino County], Arizona.

Hesperomys megalotis Merriam

N. Am. Fauna 3: 64, 11 Sep 1890a.

= Peromyscus truei truei (Shufeldt, 1885). See Osgood (1909a: 

165); subspecies recognized by Hall (1981:704) and Yancey 

and Jones (1999:583).

Holotype: USNM 17943/24854. Skin and skull. Adult male. 

Collected 21 Sep 1889 by C. H. Merriam and [SL: V. Bailey]. Original number [FC: VB] 502.

Type Locality: Black Tank, Desert of the Little Colorado [FC: 

Painted Desert], [Coconino County], Arizona.

Hesperomys (Vesperimus) melanophrys Coues

Proc. Acad. Nat. Sci. Phila. 26: 181, 15 Dec 1874.

= Peromyscus melanophrys melanophrys (Coues, 1874). See J. A. 

Allen (1897:51); subspecies recognized by Hall (1981:708).

Holotype: USNM 10183/12453. [Skin and skull]. [Lyon and Osgood (1909:134): Adult], sex not given. Collected 11 Jul 

1871 by F. Sumichrast. Original number [SL: 2].

Type Locality: Santa Efigenia, [SL: Tehuantepec], [Oaxaca], [SL: 

Mexico].

[Hesperomys] parasiticus Baird

Mammals Rep. Explor. Surv. 8(1): 479, 1857.

= Peromyscus californicus californicus (Gambel, 1848). See Smith 

(1979:718); subspecies recognized by Merritt (1999b:566).

Lectotype: USNM 1156 as selected by Grinnell and Orr 

(1934:213). Skin [mounted]. Male, age not given. Collected 

Nov 1855 by J. C. Cooper. No original number.

Type Locality: Santa Clara, California. Type locality restricted 

to Santa Clara Valley, probably near Mountain View, Santa 

Clara County, California, by Grinnell and Orr (1934:213).

Hesp[eromys] sonoriensis LeConte

Proc. Acad. Nat. Sci. Phila. 6: 413, reported favorably for publication 25 Oct 1853. Receipt acknowledged 31 Dec 1853 by 

the Smithsonian Institution.

= Peromyscus maniculatus sonoriensis (LeConte, 1853). See Osgood (1909a:89); subspecies recognized by Hall (1981:683).

Holotype: USNM 146/1128. Skin and skull. [Lyon and Osgood 

(1909:138): Young], sex not given. Collected [MC: 24–28 

Sep 1851 by J. H. Clark]. No original number.

Type Locality: [MC: Santa Cruz], Sonora, [Mexico].

Remarks: Type not designated by number in original description. 

Baird (1857:475) stated “characterized as H. sonoriensis

by Major Leconte from specimen 146 of the Smithsonian 

collection.”

Hesperomys texana Woodhouse

Proc. Acad. Nat. Sci. Phila. 6: 242, reported favorably for publication 22 Feb 1853. Receipt acknowledged 13 Apr 1853 by 

the Smithsonian Institution.

= Peromyscus leucopus texanus (Woodhouse, 1853). See Osgood 

(1909a:127); subspecies recognized by Hall (1981:688) and 

Lackey (1999:574).

Holotype: USNM 2559/37026. [Skin in alcohol, fragments of 

skull]. Age and sex not given. Collected [Lyon and Osgood 

(1909:139): probably autumn of 1852] by S. W. Woodhouse. Original number not given.

Type Locality: On the Rio Grande near El Paso. [Osgood (1909a: 

127): Probably from the vicinity of Mason, Mason County, 

Texas].

Remarks: Type not designated by number in original description, 

but a single specimen is implied. Baird’s (1857:464) list of 

specimens shows USNM 2559 as the only specimen with 

data that match that given in the original description. See 

Osgood (1909a:128) for greater detail.

Hesperomys truei Shufeldt

Proc. U.S. Natl. Mus. 8: 405, 14 Sep 1885.

= Peromyscus truei truei (Shufeldt, 1885). See Thomas (1894: 

364); subspecies recognized by Hall (1981:704) and Yancey 

and Jones (1999:583).

Holotype: USNM 14954/35108. Skin, skull, and skeleton. Adult 

male. Collected 16 [SL and MC say 14] Mar 1885 by R. W. 

Shufeldt. Original number not given. 

Type Locality: Fort Wingate, [McKinley County], New Mexico.

Remarks: USNM number incorrectly given as 14904/35108 in 

original description.

Peromyscus allophylus Osgood

Proc. Biol. Soc. Wash. 17: 71, 21 Mar 1904a.

= Peromyscus gymnotis Thomas, 1894. See Musser (1971a:8).

Holotype: USNM 77657. [Skin and skull]. Adult female. Collected 21 Feb 1896 by E. W. Nelson and E. A. Goldman. 

Original number [SL: 9352].

Type Locality: Huehuetan, [FC: altitude 500 ft (152 m)], [Huehuetan Municipality], Chiapas, Mexico.

Peromyscus altilaneus Osgood

Proc. Biol. Soc. Wash. 17: 74, 21 Mar 1904a.

= Peromyscus mexicanus (Saussure, 1860). Subspecies undetermined; see Trujano-Alvarez and Alvarez-Castaneda (2010:112).

Holotype: USNM 76856. [Skin and skull]. Adult male. Collected 

30 Dec 1895 by E. W. Nelson and E. A. Goldman. Original 

number [SL: 8942].
4 4 • smithsonian contributions to zoology

Type Locality: Todos Santos, altitude 10,000 ft (3,048 m), [Huehuetenango], Guatemala.

Peromyscus amplus Osgood

Proc. Biol. Soc. Wash. 17: 62, 21 Mar 1904a.

= Peromyscus difficilis amplus Osgood, 1904. See Osgood 

(1909a:181); subspecies recognized by Fernandez et al. 

(2010:220).

Holotype: USNM 70158. [Skin and skull]. Adult female. Collected 12 Nov 1894 by E. W. Nelson and E. A. Goldman. 

Original number [SL: 7033].

Type Locality: Coixtlahuaca, [FC: altitude 7,000 ft (2,134 m)], 

Oaxaca, Mexico.

Peromyscus attwateri eremicoides Osgood

Proc. Biol. Soc. Wash. 17: 60, 21 Mar 1904a.

= Peromyscus pectoralis pectoralis Osgood, 1904. See Schmidly 

(1972:127); subspecies recognized by Hall (1981:694) and 

Schmidly (1999a:580).

Holotype: USNM 57729. [Skin and skull]. Adult male. Collected 

15 Dec 1893 by E. A. Goldman. Original number [SL: 235].

Type Locality: Mapimi, Durango, Mexico.

Peromyscus attwateri pectoralis Osgood

Proc. Biol. Soc. Wash. 17: 59, 21 Mar 1904a.

= Peromyscus pectoralis pectoralis Osgood, 1904. See V. Bailey 

(1906:57); subspecies recognized by Hall (1981:694) and 

Schmidly (1999a:580).

Holotype: USNM 81236. [Skin and skull]. Adult male. Collected 

30 Aug 1896 by E. W. Nelson and E. A. Goldman. Original 

number [SL: 10095].

Type Locality: Jalpan, [FC: altitude about 2,000 ft (610 m)], 

Queretaro, Mexico.

Peromyscus auritus Merriam

Proc. Biol. Soc. Wash. 12: 119, 30 Apr 1898a.

= Peromyscus megalops Merriam, 1898. See Huckaby (1980:11).

Holotype: USNM 68438. [Skin and skull]. Old female. Collected 

17 Sep 1894 by E. W. Nelson and E. A. Goldman. Original 

number 6795.

Type Locality: Mountains 15 mi (24 km) west of Oaxaca, [Osgood (1909a:214): altitude 9,300 ft (2,835 m)], Oaxaca, 

Mexico.

Peromyscus banderanus angelensis Osgood

Proc. Biol. Soc. Wash. 17: 69, 21 Mar 1904a.

= Peromyscus m[exicanus] angelensis Osgood, 1904. See Musser 

(1969:5); subspecies recognized by Trujano-Alvarez and 

Alvarez-Castaneda (2010:112).

Holotype: USNM 71442. [Skin and skull]. Adult female. Collected 13 Mar 1895 by E. W. Nelson and E. A. Goldman. 

Original number [SL: 7642].

Type Locality: Puerto Angel, [FC: altitude 100 ft–200 ft 

(30 m–61m)], Oaxaca, Mexico.

Peromyscus boylii ambiguus Alvarez

Univ. Kans. Publ., Mus. Nat. Hist. 14: 118, 29 Dec 1961.

= Peromyscus levipes ambiguus Alvarez, 1961. See CastroCampillo et al. (1999:1227); subspecies recognized by 

Alvarez-Castaneda and Gonzalez-Ruiz (2009:1).

Holotype: USNM 25692/33092. Skin and skull. Adult male. 

Collected 17 Feb 1891 by W. Lloyd. Original number 377.

Type Locality: Monterrey, [FC: altitude 2,000 ft (610 m)], Nuevo 

Leon, Mexico.

Remarks: Although Alvarez (1961:118) stated that the specimen 

was a skin and skull, he only gave the skull number, USNM 

33092; USNM 25692 is the skin that belongs to it.

Peromyscus boylei laceyi V. Bailey

N. Am. Fauna 25: 99, 24 Oct 1905.

= Peromyscus attwateri J. A. Allen, 1895. See Lee et al. (1972: 706).

Holotype: USNM 92746. [Skin and skull]. Adult male. Collected 

4 Dec 1897 by H. P. Attwater. No original number. 1372-X.

Type Locality: Turtle Creek, Kerr County, Texas.

Peromyscus boylii penicillatus Mearns

Proc. U.S. Natl. Mus. 19: 2, 25 May 1896a. Preprint of Proc. 

U.S. Natl. Mus. 19: 139, 21 Dec 1896b.

= Peromyscus nasutus penicillatus Mearns, 1896. New name 

combination. Musser and Carleton (2005:1075) assigned 

penicillatus to P. nasutus as a synonym of unspecified rank; 

subspecies recognized by Hall (1981:706) under P. difficilis.

Holotype: USNM 20034/35426. Skin and skull. Adult female. 

Collected 19 Feb 1892 by E. A. Mearns and F. X. Holzner. 

Original number 1463.

Type Locality: Foothills of the Franklin Mountains near El Paso, 

[El Paso County], Texas.

Peromyscus bullatus Osgood

Proc. Biol. Soc. Wash. 17: 63, 21 Mar 1904a.

Holotype: USNM 54405. [Skin and skull]. Adult female. Collected 3 Jun 1893 by E. W. Nelson and E. A. Goldman. 

Original number [SL and FC: EWN 4932].

Type Locality: Perote, [FC: altitude 7,800 ft (2,377 m)], Veracruz, Mexico.

Peromyscus canus Mearns

Proc. U.S. Natl. Mus. 18: 3, 25 Mar 1896c. Preprint of Proc. 

U.S. Natl. Mus. 18: 445, 23 May 1896d.

= Peromyscus leucopus texanus Woodhouse, 1853. See Osgood 

(1909a:127); subspecies recognized by Hall (1981:688) and 

Lackey (1999:574).

Holotype: USNM 21109/37096. Skin and skull. Adult female. Collected 13 Jan 1893 by E. A. Mearns. Original number 2208.

Type Locality: Fort Clark, Kinney County, Texas.

Peromyscus comptus Merriam

Proc. Biol. Soc. Wash. 12: 120, 30 Apr 1898a.

= Peromyscus megalops Merriam 1898. See Huckaby (1980:11).
number 642 • 4 5

Holotype: USNM 70191. [Skin and skull]. Adult male. Collected 

24 Dec 1894 by E. W. Nelson and E. A. Goldman. Original 

number 7266.

Type Locality: Mountains near Chilpancingo, [FC: altitude 

9,800 ft (2,987 m)], Guerrero, Mexico.

Peromyscus cozumelae Merriam

Proc. Biol. Soc. Wash. 14: 103, 19 Jul 1901c.

= Peromyscus leucopus cozumelae Merriam, 1901. See Osgood 

(1909a:135); subspecies recognized by Hall (1981:686) and 

Lackey (1999:574).

Holotype: USNM 108449. [Skin and skull]. Adult male. Collected 11 Apr 1901 by E. W. Nelson and E. A. Goldman. 

Original number 14686.

Type Locality: Cozumel Island, Yucatan (Quintana Roo), [SL: 

Mexico].

Peromyscus crinitus disparilis Goldman

Proc. Biol. Soc. Wash. 45: 90, 21 Jun 1932d.

Subspecies recognized by Hall (1981:691) and Armstrong 

(1999a:567).

Holotype: USNM 202989. Skin and skull. Adult male. Collected 

22 Nov 1913 by E. A. Goldman. Original number 22304.

Type Locality: [SL and FC: Near] Tinajas Altas, altitude 2,000 

ft (610 m) [SL and FC: 2,500 ft (762 m)], Gila Mountains, 

Yuma County, Arizona.

Peromyscus crinitus pergracilis Goldman

J. Mammal. 20: 356, 14 Aug 1939.

Subspecies recognized by Hall (1981:692) and Armstrong 

(1999a:567).

Holotype: USNM 264022. Skin and skull. [Age and] sex undetermined [FC: male]. Collected 22 Jun 1938 by W. H. Marshall. Original number 100.

Type Locality: South end of Stansbury Island, altitude 4,250 ft 

(1,295 m), Great Salt Lake, [Tooele County], Utah.

Peromyscus crinitus peridoneus Goldman

J. Mammal. 18: 92, 14 Feb 1937b.

= Peromyscus crinitus auripectus (J. A. Allen, 1893). See Hall 

and Hoffmeister (1942:64); subspecies recognized by Hall 

(1981:691) and Armstrong (1999a:567).

Holotype: USNM 202424. Skin and skull. Adult male. Collected 

19 Aug 1913 by E. A. Goldman. Original number 22145.

Type Locality: [SL and FC: Near] Bright Angel Trail, altitude 

4,800 ft (1,463 m), south side of Grand Canyon, Coconino 

County, Arizona.

Peromyscus eremicus arenarius Mearns

Proc. U.S. Natl. Mus. 19: 2, 25 May 1896a. Preprint of Proc. 

U.S. Natl. Mus. 19: 138, 21 Dec 1896b.

= Peromyscus eremicus eremicus (Baird, 1857). See Osgood 

(1909a:239); subspecies recognized by Hall (1981:658) and 

Caire (1999:568).

Holotype: USNM 20018/35413. Skin and skull. Adult male. 

Collected 25 Feb 1892 by E. A. Mearns and F. X. Holzner. 

Original number 1513.

Type Locality: Rio Grande, about 6 mi (10 km) from El Paso, [El 

Paso County], Texas.

Peromyscus eremicus avius Osgood

N. Am. Fauna 28: 247, 17 Apr 1909a.

Subspecies recognized by Hall (1981:657) and Caire (1999:568).

Holotype: USNM 147024. [Skin and skull]. Adult female. Collected 13 Feb 1906 by E. W. Nelson and E. A. Goldman. 

Original number [SL: 19125].

Type Locality: Ceralbo (Jacques Cousteau) Island, off east coast 

of southern Lower California (Baja California Sur), Mexico.

Peromyscus eremicus carmeni Townsend

Bull. Am. Mus. Nat. Hist. 31: 126, 14 Jun 1912.

= Peromyscus eva carmeni Townsend, 1912. See Lawlor (1971: 

17); subspecies recognized by Alvarez-Castaneda and CortesCalva (2003:1).

Holotype: USNM 198408. Skin and skull. Male, age not given. 

Collected [SL and MC: 3] Apr 1911, not 13 Apr as given in 

original description, by H. E. Anthony. Original number 109.

Type Locality: Carmen Island. [Gulf of California], Lower California (Baja California Sur), [SL: Mexico].

Remarks: Published as AMNH 31855; cataloged as USNM 

198408 on 26 Jun 1914.

Peromyscus eremicus insulicola Osgood

N. Am. Fauna 28: 246, 17 Apr 1909a.

Subspecies recognized by Hall (1981:658) and Caire (1999:568).

Holotype: USNM 147010. [Skin and skull]. Adult male. Collected 9 Feb 1906 by E. W. Nelson and E. A. Goldman. 

Original number [SL: 19077].

Type Locality: Espiritu Santo Island, [FC: altitude 30 ft (9 m)], 

off east coast of southern Lower California (Baja California 

Sur), Mexico.

Peromyscus eremicus papagensis Goldman

Proc. Biol. Soc. Wash. 30: 110, 23 May 1917.

Subspecies recognized by Hall (1981:659) and Caire (1999:568).

Holotype: USNM 210698. [Skin and skull]. Adult female. Collected [XC: 17] Feb 1915 by C. Sheldon. Original number 

[XC: 1]. [SL: 11962-X].

Type Locality: Pinacate Mountains, Sonora, Mexico.

Peromyscus eremicus phaeurus Osgood

Proc. Biol. Soc. Wash. 17: 75, 21 Mar 1904a.

Subspecies recognized by Hall (1981:660) and Caire (1999:568).

Holotype: USNM 50438. [Skin and skull]. Adult female. Collected 20 Aug 1892 by E. W. Nelson. Original number [SL: 

3227].

Type Locality: Hacienda La Parada, [FC: altitude about 6,000 ft 

(1,829 m)], San Luis Potosi, Mexico.
4 6 • smithsonian contributions to zoology

Peromyscus eremicus polypolius Osgood

N. Am. Fauna 28: 248, 17 Apr 1909a.

Subspecies recognized by Hall (1981:660) and Caire (1999:568).

Holotype: USNM 146074. [Skin and skull]. Adult male. Collected 30 Nov 1905 by E. W. Nelson and E. A. Goldman. 

Original number [SL: 18676].

Type Locality: Margarita Island, [FC: altitude 500 ft (152 m)], 

off west coast of southern Lower California (Baja California 

Sur), Mexico.

Peromyscus felipensis Merriam

Proc. Biol. Soc. Wash. 12: 122, 30 Apr 1898a.

= Peromyscus difficilis felipensis Merriam, 1898. See Osgood 

(1909a:182); subspecies recognized by Fernandez et al. 

(2010:220).

Holotype: USNM 68409. [Skin and skull]. Adult male. Collected 

22 Aug 1894 by E. W. Nelson and E. A. Goldman. Original 

number 6611.

Type Locality: Cerro San Felipe, altitude 10,200 ft (3,109 m), 

Oaxaca, Mexico.

Peromyscus goldmani Osgood

Proc. Biol. Soc. Wash. 17: 75, 21 Mar 1904a.

= Peromyscus merriami goldmani Osgood, 1904. See Hoffmeister and Diersing (1973:354); subspecies recognized by Hall 

(1981:661) and Armstrong (1999b:578).

Holotype: USNM 96340. [Skin and skull]. Adult female. Collected 19 Dec 1898 by [SL: E. W. Nelson and] E. A. Goldman. Original number [SL: Nelson and Goldman 13306].

Type Locality: Alamos, [FC: altitude about 1,200 ft (366 m)], 

Sonora, Mexico.

Peromyscus gossypinus restrictus A. H. Howell

J. Mammal. 20: 364, 14 Aug 1939b.

Subspecies recognized by Hall (1981:690) and Linzey (1999a: 

570).

Holotype: USNM 263539. Skin and skull. Adult female. Collected 

13 Mar 1938 by L. C. Goldman. Original number 479.

Type Locality: On Chadwick (Englewood) Beach, near Englewood, [SL and FC: Sarasota County], Florida.

Peromyscus gratus Merriam

Proc. Biol. Soc. Wash. 12: 123, 30 Apr 1898a.

= Peromyscus gratus gratus Merriam, 1898. See Modi and Lee 

(1984:722); subspecies recognized by Hall (1981:703), 

under P. truei, and Frey (1999b:571).

Holotype: USNM 50619. [Skin and skull]. [Lyon and Osgood 

(1909:131): Adult] male. Collected 30 Nov 1892 by E. W. 

Nelson. Original number 3927.

Type Locality: Tlalpam, [FC: altitude 7,600 ft (2,316 m)], Valley 

of Mexico [SL: Distrito Federal], [Mexico].

Peromyscus gratus gentilis Osgood

Proc. Biol. Soc. Wash. 17: 61, 21 Mar 1904a.

Subspecies recognized by Hall (1981:703), under P. truei, and 

Frey (1999b:571).

Holotype: USNM 78937. [Skin and skull]. Adult male. Collected 

27 Jun 1896 by E. W. Nelson and E. A. Goldman. Original 

number [SL: 9702].

Type Locality: Lagos, [FC: altitude 6,150 ft (1,875 m)], Jalisco, 

Mexico.

Peromyscus guardia Townsend

Bull. Am. Mus. Nat. Hist. 31: 126, 14 Jun 1912.

= Peromyscus guardia guardia Townsend, 1912. See Burt (1932: 

175); subspecies recognized by Mellink et al. (2002:107).

Holotype: USNM 198406. Skin and skull. Age not given, female. 

Collected 11 Apr 1911 by [SL: H. E. Anthony. Original 

number 135].

Type Locality: Angel de la Guardia Island, [Gulf of California], 

Lower California (Baja California), [SL: Mexico].

Remarks: Published as AMNH 31907; cataloged as USNM 

198406 on 26 Jun 1914.

Peromyscus guatemalensis Merriam

Proc. Biol. Soc. Wash. 12: 118, 30 Apr 1898a.

Holotype: USNM 76861. [Skin and skull]. Adult male. Collected 

31 Dec 1895 by E. W. Nelson and E. A. Goldman. Original 

number 8991.

Type Locality: Todos Santos, altitude 10,000 ft (3,048 m), [Huehuetenango], Guatemala.

Peromyscus hylaeus Osgood

Proc. Biol. Soc. Wash. 21: 141, 9 Jun 1908.

= Peromyscus keeni hylaeus Osgood, 1908. See Hogan et al. 

(1993:828); subspecies recognized by Hall (1981:677), 

under P. maniculatus, and Greenbaum (1999:572).

Holotype: USNM 127038. [Skin and skull]. [Lyon and Osgood 

(1909:132): Adult] [SL and FC: male]. Collected 15 May 

1903 by W. H. Osgood. Original number [SL: 2234].

Type Locality: Hollis, Kasaan Bay, Prince of Wales Island, Alaska.

Peromyscus hylocetes Merriam

Proc. Biol. Soc. Wash. 12: 124, 30 Apr 1898a.

Holotype: USNM 50423. [Skin and skull]. Adult female. Collected 27 Jul 1892 by E. W. Nelson and E. A. Goldman. 

Original number [FC: EWN] 2961.

Type Locality: Patzcuaro, altitude about 8,000 ft (2,438 m), Michoacan, Mexico.

Peromyscus leucocephalus A. H. Howell

J. Mammal. 1: 239, 4 Dec 1920b.

= Peromyscus polionotus leucocephalus A. H. Howell, 1920. See 

Sumner (1926:150); subspecies recognized by Mullen et al. 

(2009: Table S4).

Holotype: USNM 234358. Skin and skull. Adult female. Collected 26 Feb 1920 by A. H. Howell. Original number 

2392.
number 642 • 4 7

Type Locality: Santa Rosa Island, opposite Camp Walton, [Santa 

Rosa County], Florida.

Peromyscus leucopus aridulus Osgood

N. Am. Fauna 28: 122, 17 Apr 1909a.

Subspecies recognized by Hall (1981:685) and Lackey (1999: 

574).

Holotype: USNM 75704. [Skin and skull]. Adult female. Collected 12 Nov 1895 by J. A. Loring. Original number [SL: 

3378].

Type Locality: Fort Custer, [Big Horn County], Montana.

Peromyscus leucopus buxtoni Shipp-Pennock, Webster, and 

Freshwater

J. Mammal. 86: 809, 22 Aug 2005.

Holotype: USNM 569137. Skin and skull. Adult male. Collected 

15 Nov 1987 by D. Webster. Original number 1946. University of North Carolina, Wilmington number 1708.

Type Locality: Buxton Woods, Jeannette’s Sedge, Dare County, 

North Carolina.

Peromyscus leucopus easti Paradiso

Proc. Biol. Soc. Wash. 73: 21, 10 Aug 1960.

Subspecies recognized by Shipp-Pennock et al. (2005:810). 

Holotype: USNM 302764. Skin and skull. Adult male. Collected 

10 Apr 1956 by C. O. Handley Jr. Original number 3052.

Type Locality: [SL: Back Bay National Wildlife Refuge Headquarters], 6.8 mi (11 km) southeast of Pungo, Princess Anne 

County, Virginia.

Peromyscus leucopus incensus Goldman

Proc. Biol. Soc. Wash. 55: 157, 17 Oct 1942b.

Subspecies recognized by Hall (1981:686) and Lackey (1999: 

574).

Holotype: USNM 93069. [Skin and skull]. Adult female. Collected 28 Jan 1898 by E. A. Goldman. Original number [FC: 

Nelson and Goldman] 12107.

Type Locality: Metlaltoyuca, altitude 800 ft (244 m), Puebla, 

Mexico.

Peromyscus leucopus minnesotae Mearns

Proc. Biol. Soc. Wash. 14: 154, 9 Aug 1901.

= Peromyscus leucopus noveboracensis (Fischer, 1829). See Osgood (1909a:117); subspecies recognized by Hall (1981:688) 

and Shipp-Pennock et al. (2005:810). 

Holotype: USNM 82717. [Skin and skull]. Adult female. Collected 30 Nov 1890 by E. A. Mearns. Original number 

1181.

Type Locality: Fort Snelling, Hennepin County, Minnesota.

Peromyscus leucopus ochraceus Osgood

N. Am. Fauna 28: 124, 17 Apr 1909a.

Subspecies recognized by Hall (1981:688) and Lackey (1999: 

574).

Holotype: USNM 53301. [Skin and skull]. Adult male. Collected 

30 Apr 1893 by C. P. Streator. Original number [SL: 2674].

Type Locality: Winslow, [Navajo County], Arizona.

Peromyscus luteus Osgood

Proc. Biol. Soc. Wash. 18: 77, 21 Feb 1905.

= Peromyscus maniculatus luteus Osgood, 1905. See Osgood 

(1909a:77); subspecies recognized by Hall (1981:679).

Holotype: USNM 18658/25447. Skin and skull. Adult female. 

Collected 23 Apr 1890 by V. Bailey. Original number [SL: 

1079].

Type Locality: Kennedy, [Cherry County], Nebraska.

Peromyscus levipes Merriam

Proc. Biol. Soc. Wash. 12: 123, 30 Apr 1898a.

= Peromyscus levipes levipes Merriam, 1898. See CastroCampillo et al. (1999:1228); subspecies recognized by Alvarez-Castaneda and Gonzalez-Ruiz (2009:1).

 Holotype: USNM 53673. [Skin and skull]. Adult male. Collected 

12 May 1893 by E. W. Nelson. Original number 4799.

Type Locality: Mount Malinche, altitude 8,400 ft (2,560 m), 

Tlaxcala, Mexico.

Peromyscus madrensis Merriam

Proc. Biol. Soc. Wash. 12: 16, 27 Jan 1898e.

Holotype: USNM 89223. [Skin and skull]. Adult male. Collected 

18 May 1897 by E. W. Nelson and E. A. Goldman. Original 

number 11040.

Type Locality: Maria Madre Island, [FC: altitude 800 ft (244 m)], 

Tres Marias Islands, [Nayarit], Mexico.

Peromyscus maniculatus algidus Osgood

N. Am. Fauna 28: 56, 17 Apr 1909a.

= Peromyscus keeni algidus Osgood, 1909. See Hogan et al. 

(1993:828); subspecies recognized by Hall (1981:670), 

under P. maniculatus, and Greenbaum (1999:572).

Holotype: USNM 130013. [Skin and skull]. Adult male. Collected 

17 Sep 1903 by W. H. Osgood. Original number [SL: 2690].

Type Locality: Head of Lake Bennett, site of old Bennett City, 

British Columbia, [Canada].

Peromyscus maniculatus assimilis Nelson and Goldman

J. Mammal. 12: 305, 24 Aug 1931b.

Subspecies recognized by Hall (1981:671).

Holotype: USNM 77939. Skin and skull. Adult male. Collected 

24 Apr 1896 by A. W. Anthony. Original number [SL: 13].

Type Locality: Coronados Island, off west coast of Lower California (Baja California), Mexico.

Peromyscus maniculatus dorsalis Nelson and Goldman

J. Wash. Acad. Sci. 21: 535, 19 Dec 1931c.

Subspecies recognized by Hall (1981:673).

Holotype: USNM 80857. [Skin and skull]. Adult female. Collected 4 Aug 1896 by A. W. Anthony. Original number 96.
4 8 • smithsonian contributions to zoology

Type Locality: Natividad Island, off west coast of Lower California (Baja California), Mexico.

Peromyscus maniculatus elusus Nelson and Goldman

J. Wash. Acad. Sci. 21: 533, 19 Dec 1931c.

Subspecies recognized by Hall (1981:673).

Holotype: USNM 92049. [Skin and skull]. Adult male. Collected 

16 May 1897 by H. A. Gaylord. Original number [XC: 18]. 

1095-X.

Type Locality: Santa Barbara Island, [Channel Islands], off the west 

coast of southern California, [Ventura County, California].

Peromyscus maniculatus eremus Osgood

N. Am. Fauna 28: 47, 17 Apr 1909a.

Subspecies recognized by Hall (1981:673).

Holotype: USNM 150223. [Skin and skull]. Adult female. Collected 9 Aug 1907 by W. H. Osgood. Original number [SL: 

3197].

Type Locality: Pleasant Bay, Grindstone Island, Magdalen Islands, Quebec, [Canada].

Peromyscus maniculatus exterus Nelson and Goldman

J. Wash. Acad. Sci. 21: 532, 19 Dec 1931c.

Subspecies recognized by Hall (1981:673).

Holotype: USNM 92098. [Skin and skull]. Old adult female. 

Collected 23 May 1897 by H. A. Gaylord. Original number 

[XC: 80]. [SL: 1121-X].

Type Locality: San Nicolas Island, [Channel Islands], off the west 

coast of southern California, [Ventura County, California].

Peromyscus maniculatus gunnisoni Goldman

Proc. Biol. Soc. Wash. 50: 224, 28 Dec 1937c.

= Peromyscus maniculatus sonoriensis (LeConte, 1853). See Durrant (1952:307); subspecies recognized by Hall (1981:683).

Holotype: USNM 262845. Skin and skull. Adult female. Collected 1 Jun 1937 by A. M. Bailey and R. J. Niedrach. 

Original number not given. [SL: 28019-X]. [SL: Colorado 

Museum of Natural History number] 3010.

Type Locality: Gunnison Island, altitude about 4,300 ft (1,311 

m), Great Salt Lake, [Box Elder County], Utah.

Peromyscus maniculatus hollisteri Osgood

N. Am. Fauna 28: 62, 17 Apr 1909a.

Subspecies recognized by Hall (1981:677).

Holotype: USNM 130316. [Skin and skull]. Adult male. Collected 

23 Oct 1903 by N. Hollister. Original number [SL: 832].

Type Locality: Friday Harbor, San Juan Island, San Juan County, 

Washington.

Peromyscus maniculatus inclarus Goldman

J. Mammal. 20: 355, 14 Aug 1939.

Subspecies recognized by Hall (1981:677).

Holotype: USNM 264119. Skin and skull. Adult male. Collected 

7 Aug 1938 by W. H. Marshall. Original number 276.

Type Locality: Fremont Island, Great Salt Lake, altitude about 

4,250 ft (1,295 m), [Weber County], Utah.

Peromyscus maniculatus magdalenae Osgood

N. Am. Fauna 28: 101, 17 Apr 1909a.

Subspecies recognized by Hall (1981:679).

Holotype: USNM 146971. [Skin and skull]. Old male. Collected 

3 Dec 1906 by E. W. Nelson and E. A. Goldman. Original 

number [SL: 18749].

Type Locality: Magdalena Island, [FC: altitude 20 ft (6 m)], off 

west coast of southern Lower California (Baja California 

Sur), Mexico.

Peromyscus maniculatus margaritae Osgood

N. Am. Fauna 28: 95, 17 Apr 1909a.

Subspecies recognized by Hall (1981:679).

Holotype: USNM 146958. [Skin and skull]. Adult female. Collected 1 Dec 1905 by E. W. Nelson and E. A. Goldman. 

Original number [SL: 18726].

Type Locality: Margarita Island, [FC: altitude 20 ft (6 m)], off 

west coast of southern Lower California (Baja California 

Sur), Mexico.

Peromyscus maniculatus martinensis Nelson and Goldman

J. Wash. Acad. Sci. 21: 534, 19 Dec 1931c.

= Peromyscus maniculatus exiguus J. A. Allen, 1898. See Nelson 

and Goldman (1932:371); subspecies recognized by Hall 

(1981:673).

Holotype: USNM 138979. [Skin and skull]. Adult female. Collected 13 Aug 1905 by [E. W.] Nelson and [E. A.] Goldman. 

Original number 17726.

Type Locality: San Martin Island, [FC: 25 ft (8 m)], off west 

coast of Lower California (Baja California), Mexico.

Peromyscus maniculatus santacruzae Nelson and Goldman

J. Wash. Acad. Sci. 21: 532, 19 Dec 1931c.

Subspecies recognized by Hall (1981:682).

Holotype: USNM 35184/47449. Skin and skull. Old adult female. Collected 13 Jul 1892 by C. P. Streator. Original 

number 1923.

Type Locality: Santa Cruz Island, [Channel Islands], off the 

coast of southern California, [Santa Barbara County, 

California].

Remarks: Published as USNM 47449, the skull number to which 

skin number USNM 35184 belongs.

Peromyscus maniculatus streatori Nelson and Goldman

J. Wash. Acad. Sci. 21: 531, 19 Dec 1931c.

Subspecies recognized by Hall (1981:683).

Holotype: USNM 34631/46716. Skin and skull. Old adult male. 

Collected 24 Jun 1892 by C. P. Streator. Original number 

1861.

Type Locality: San Miguel Island, [Channel Islands], off the coast 

of southern California, [Santa Barbara County], California.
number 642 • 4 9

Peromyscus megalops Merriam

Proc. Biol. Soc. Wash. 12: 119, 30 Apr 1898a.

Holotype: USNM 71592. [Skin and skull]. Old male. Collected 

26 Mar 1895 by E. W. Nelson and E. A. Goldman. Original 

number 7733.

Type Locality: Mountains near Ozolotepec, [Huckaby (1980:11): 

at a ranch called Las Cieneguilla at 10,000 ft (3,048 m) in 

the Sierra Madre del Sur, near the village of Santa Maria 

Ozolotepec], Oaxaca, Mexico.

Peromyscus megalops melanurus Osgood

N. Am. Fauna 28:215, 17 Apr 1909a.

= Peromyscus melanurus Osgood, 1909. See Huckaby (1980:12).

Holotype: USNM 71385. [Skin and skull]. Adult male. Collected 

20 Mar 1895 by E. W. Nelson and E. A. Goldman. Original 

number [SL: 7703].

Type Locality: [Huckaby (1980:12): Below] Pluma, altitude 

4,600 ft (1,402 m) [FC: 3,000 ft (914 m)], Oaxaca, Mexico.

Peromyscus mekisturus Merriam

Proc. Biol. Soc. Wash. 12: 124, 30 Apr 1898a.

Holotype: USNM 64108. [Skin and skull]. Adult female. Collected 16 Mar 1894 by E. W. Nelson and E. A. Goldman. 

Original number 5951.

Type Locality: Chalchicomula, altitude 8,400 ft (2,560 m), 

Puebla, Mexico.

Peromyscus melanocarpus Osgood

Proc. Biol. Soc. Wash. 17: 73, 21 Mar 1904a.

Holotype: USNM 68610. [Skin and skull]. Young adult female. 

Collected 8 Jul 1894 by E. W. Nelson and E. A. Goldman. 

Original number [SL: 6379].

Type Locality: Mount Zempoaltepec, altitude 8,000 ft (2,438 m), 

Oaxaca, Mexico.

Peromyscus melanophrys consobrinus Osgood

Proc. Biol. Soc. Wash. 17: 66, 21 Mar 1904a.

Subspecies recognized by Hall (1981:708).

Holotype: USNM 79626. [Skin and skull]. Adult female. Collected 10 Jul 1896 by E. W. Nelson and E. A. Goldman. 

Original number [SL: 9818].

Type Locality: Berriozabal, [FC: altitude 6,600 ft (2,012 m)], 

Zacatecas, Mexico.

Peromyscus melanophrys zamorae Osgood

Proc. Biol. Soc. Wash. 17: 65, 21 Mar 1904a.

Subspecies recognized by Hall (1981:709).

Holotype: USNM 120288. [Skin and skull]. Adult female, not 

male as in original description. Collected 20 Jan 1903 by E. 

W. Nelson and E. A. Goldman. Original number [SL: 15783].

Type Locality: Zamora, Michoacan, Mexico.

Peromyscus melanotis zamelas Osgood

Proc. Biol. Soc. Wash. 17: 59, 21 Mar 1904a.

= Peromyscus melanotis J. A. Allen and Chapman, 1897. See 

Osgood (1909a:109).

Holotype: USNM 98197. [Skin and skull]. Adult female. Collected 24 Jul 1899, not 23 Jul as in original description, by E. 

W. Nelson and E. A. Goldman. Original number [SL: 13915].

Type Locality: [SL and FC: Near] Colonia Garcia, altitude 6,700 

ft (2,042 m), Chihuahua, Mexico.

Peromyscus merriami Mearns

Proc. U.S. Natl. Mus. 19: 2, 25 May 1896a. Preprint of Proc. 

U.S. Natl. Mus. 19: 138, 21 Dec 1896b.

= Peromyscus merriami merriami Mearns, 1896. See Hall and 

Kelson (1952:368); subspecies recognized by Hall (1981: 

661) and Armstrong (1999b:578).

Holotype: USNM 59234. Skin and skull. Adult female. Collected 

21 Jan 1894 by E. A. Mearns and F. X. Holzner. Original 

number 2815.

Type Locality: Sonoyta, on the Sonoyta River, Sonora, Mexico.

Peromyscus mexicanus orizabae Merriam

Proc. Biol. Soc. Wash. 12: 121, 30 Apr 1898a.

= Peromyscus mexicanus totontepecus Merriam, 1898. See 

Osgood (1909a:202); subspecies recognized by TrujanoAlvarez and Alvarez-Castaneda (2010:112).

Holotype: USNM 58197. [Skin and skull]. Adult male. Collected 

29 Jan 1894 by E. W. Nelson and E. A. Goldman. Original 

number 5787.

Type Locality: Orizaba, altitude 4,200 ft (1,280 m), Veracruz, 

Mexico.

Peromyscus mexicanus saxatilis Merriam

Proc. Biol. Soc. Wash. 12: 121, 30 Apr 1898a.

Subspecies recognized by Trujano-Alvarez and Alvarez-Castaneda (2010:112).

Holotype: USNM 77296. [Skin and skull]. Adult male. Collected 

19 Dec 1895 by E. W. Nelson and E. A. Goldman. Original 

number 8824.

Type Locality: Jacaltenango, [FC: altitude 5,400 ft (1,646 m)], 

Huehuetenango, Guatemala.

Peromyscus mexicanus teapensis Osgood

Proc. Biol. Soc. Wash. 17: 69, 21 Mar 1904a.

Subspecies recognized by Trujano-Alvarez and AlvarezCastaneda (2010:112).

Holotype: USNM 100022. [Skin and skull]. Adult female. Collected 25 Mar 1900 by E. W. Nelson and E. A. Goldman. 

Original number [SL: 14067].

Type Locality: Teapa, [Osgood (1909a:204): altitude 800 ft 

(244 m)], Tabasco, Mexico.

Peromyscus mexicanus totontepecus Merriam

Proc. Biol. Soc. Wash. 12: 120, 30 Apr 1898a.

Subspecies recognized by Trujano-Alvarez and AlvarezCastaneda (2010:112).
5 0 • smithsonian contributions to zoology

Holotype: USNM 68624. [Skin and skull]. Adult female. Collected 16 Jul 1894 by E. W. Nelson and E. A. Goldman. 

Original number 6465.

Type Locality: Totontepec, altitude 6,500 ft (1,981 m), Oaxaca, 

Mexico.

Peromyscus musculoides Merriam

Proc. Biol. Soc. Wash. 12: 124, 30 Apr 1898a.

= Peromyscus leucopus affinis (J. A. Allen, 1891). See Osgood 

(1909a:133); subspecies recognized by Hall (1981:685) and 

Lackey (1999:574).

Holotype: USNM 69661. [Skin and skull]. Old male. Collected 

14 Oct 1894 by E. W. Nelson and E. A. Goldman. Original 

number 6892.

Type Locality: Cuicatlan, altitude 1,800 ft (549 m), Oaxaca, 

Mexico.

Peromyscus oaxacensis Merriam

Proc. Biol. Soc. Wash. 12: 122, 30 Apr 1898a.

= Peromyscus aztecus oaxacensis Merriam, 1898. See Carleton (1979:205); subspecies recognized by Vazquez et al. 

(2001:1).

Holotype: USNM 68426. [Skin and skull]. Young adult male. 

Collected 1 Sep 1894 by E. W. Nelson and E. A. Goldman. 

Original number 6700.

Type Locality: Cerro San Felipe, altitude 10,000 ft (3,048 m) 

[FC: 9,000 ft (2,743 m)], Oaxaca, Mexico.

Peromyscus oreas rubidus Osgood

Proc. Biol. Soc. Wash. 14: 193, 12 Dec 1901b.

= Peromyscus maniculatus rubidus Osgood, 1901. See Osgood 

(1909a:65); subspecies recognized by Hall (1981:681).

Holotype: USNM 91650. [Skin and skull]. Young adult female. 

Collected 17 Nov 1897 by J. A. Loring. Original number 

4925.

Type Locality: Mendocino City, Mendocino County, California.

Peromyscus pectoralis laceianus V. Bailey

Proc. Biol. Soc. Wash. 19: 57, 1 May 1906.

Subspecies recognized by Hall (1981:693) and Schmidly (1999a: 

580).

Holotype: USNM 97063. [Skin and skull]. Adult male. Collected 

3 May 1899 by V. Bailey. Original number 6860.

Type Locality: Ranch of Howard Lacey, Turtle Creek, near Kerrville, [Kerr County], Texas.

Peromyscus polionotus decoloratus A. H. Howell

J. Mammal. 20: 363, 14 Aug 1939b.

Subspecies recognized by Hall (1981:668) and Holler (1999:582).

Holotype: USNM 245780. Skin and skull. [Poole and Schantz 

(1942:331): Adult] [SL: male]. Collected 25 May 1925 by 

A. H. Howell. Original number 2448.

Type Locality: Ponce Park, near Mosquito Inlet, [Volusia 

County], Florida.

Peromyscus polionotus griseobracatus Bowen

Bull. Fla. State Mus. 12: 16, 19 Apr 1968.

Subspecies recognized by Hall (1981:668) and Holler (1999:582).

Holotype: USNM 308957. [Skin and skull]. Adult female. Collected Apr 1955 by E. V. Komarek. Original number [SL: 

Ppa-28].

Type Locality: 20 mi (32 km) west of Fort Walton, about 5 mi 

(8 km) west of Navarre, Santa Rosa County, Florida.

Peromyscus polius Osgood

Proc. Biol. Soc. Wash. 17: 61, 21 Mar 1904a.

Holotype: USNM 98226. [Skin and skull]. Old female. Collected 

26 Jun 1899 by E. W. Nelson and E. A. Goldman. Original 

number [SL: 13846].

Type Locality: [SL and FC: Near] Colonia Garcia, [FC: altitude 

6,400 ft (1,951 m)], Chihuahua, Mexico.

Peromyscus prevostensis Osgood

N. Am. Fauna 21: 29, 26 Sep 1901c.

= Peromyscus keeni prevostensis Osgood, 1901. See Hogan et 

al. (1993:828); subspecies recognized by Hall (1981:681), 

under P. maniculatus, and Greenbaum (1999:572).

Holotype: USNM 100818. [Skin and skull]. Old adult female. 

Collected 5 Jul 1900 by W. H. Osgood and E. Heller. Original number [FC: WHO] 1135.

Type Locality: Prevost Island, [FC: north end of island on coast 

of Houston Stewert Channel], Queen Charlotte Islands, 

British Columbia, [Canada].

Peromyscus sitkensis Merriam

Proc. Biol. Soc. Wash. 11: 223, 15 Jul 1897a.

= Peromyscus keeni sitkensis Merriam, 1897. See Hogan et al. 

(1993:828); subspecies recognized by Greenbaum (1999: 

572).

Holotype: USNM 73809. [Skin and skull]. Adult male. Collected 

30 Jul 1895 by C. P. Streator. Original number 4720.

Type Locality: Sitka, Alaska.

Peromyscus sonoriensis blandus Osgood

Proc. Biol. Soc. Wash. 17: 56, 21 Mar 1904a.

= Peromyscus maniculatus blandus Osgood, 1904. See Osgood 

(1909a:84); subspecies recognized by Hall (1981:672).

Holotype: USNM 57635. [Skin and skull]. Adult female. Collected 27 Nov 1893 by E. A. Goldman. Original number 

[SL: 166].

Type Locality: Escalon, Chihuahua, Mexico.

Peromyscus sonoriensis fulvus Osgood

Proc. Biol. Soc. Wash. 17: 57, 21 Mar 1904a.

= Peromyscus maniculatus fulvus Osgood, 1904. See Osgood 

(1909a:86); subspecies recognized by Hall (1981:673).

Holotype: USNM 68655. [Skin and skull]. Adult male. Collected 

12 Jun 1894 by E. W. Nelson and E. A. Goldman. Original 

number [SL: 6277].
number 642 • 5 1

Type Locality: Oaxaca City, [FC: altitude 5,000 ft (1,524 m)], 

Oaxaca, Mexico.

Peromyscus spicilegus evides Osgood

Proc. Biol. Soc. Wash. 17: 64, 21 Mar 1904a.

= Peromyscus aztecus evides Osgood, 1904. See Carleton (1979: 

294); subspecies recognized by Vazquez et al. (2001:1).

Holotype: USNM 71426. [Skin and skull]. Adult male. Collected 

28 Feb 1895 by E. W. Nelson and E. A. Goldman. Original 

number [SL: 7572].

Type Locality: Juquila, [FC: altitude 5,000 ft (1,524 m)], Oaxaca, 

Mexico.

Peromyscus spicilegus simulus Osgood

Proc. Biol. Soc. Wash. 17: 64, 21 Mar 1904a.

= Peromyscus simulus Osgood, 1904. See Carleton (1977:41).

Holotype: USNM 88088. [Skin and skull]. Adult male. Collected 

18 Apr 1897 by E. W. Nelson and E. A. Goldman. Original 

number [SL: 10933].

Type Locality: San Blas, [FC: altitude 50 ft (15 m)], Tepic 

(Nayarit), Mexico.

Peromyscus stephani Townsend

Bull. Am. Mus. Nat. Hist. 31: 126, 14 Jun 1912.

Holotype: USNM 198407. Skin and skull. Age not given, male. 

Collected 14 Apr 1911 by [SL: H. E. Anthony]. Original 

number [SL: 162].

Type Locality: San Esteban Island, [Gulf of California], Lower 

California (Baja California), [SL: Mexico].

Remarks: Published as AMNH 31961; cataloged as USNM 

198407 on 26 Jun 1914.

Peromyscus stephensi Mearns

Proc. U.S. Natl. Mus. 19: 721, 30 Jul 1897a.

= Peromyscus crinitus stephensi Mearns, 1897. See Osgood 

(1909a:232); subspecies recognized by Hall (1981:692) and 

Armstrong (1999a:567).

Holotype: USNM 61026. Skin and skull. Adult female. Collected 9 May 1894 by E. A. Mearns. Original number [FC: 

Mearns and Holzner] 3512.

Type Locality: From the lowest water, on the wagon road, in 

a canyon, at the eastern base of the Coast Range Mountains, San Diego County, California. Grinnell (1933:171) 

restricted the locality to 3 mi (4.8 km) east of Mountain 

Spring, Imperial County, California.

Peromyscus tehuantepecus Merriam

Proc. Biol. Soc. Wash. 12: 122, 30 Apr 1898a.

= Peromyscus mexicanus mexicanus (Saussure, 1860). See Osgood (1909a:199); subspecies recognized by Hall (1981: 

710).

Holotype: USNM 75302. [Skin and skull]. Adult female, not 

male as in original description. Collected 23 May 1895 by 

E. W. Nelson and E. A. Goldman. Original number 7980.

Type Locality: [SL and FC: Near] Tehuantepec, [FC: altitude 

1,400 ft (427 m)], Oaxaca, Mexico.

Peromyscus texanus castaneus Osgood

Proc. Biol. Soc. Wash. 17: 58, 21 Mar 1904a.

= Peromyscus leucopus castaneus Osgood, 1904. See Osgood 

(1909a:133); subspecies recognized by Hall (1981:686) and 

Lackey (1999:574).

Holotype: USNM 107980. Skin, skull, [and baculum]. Adult 

male. Collected 19 Dec 1900 by E. W. Nelson and E. A. 

Goldman. Original number [SL: 14283].

Type Locality: Vicinity of [SL and FC: Near] Yohaltun, 

Campeche, Peninsula of Yucatan, Mexico.

Peromyscus texanus clementis Mearns

Proc. U.S. Natl. Mus. 18: 4, 25 Mar 1896c. Preprint of Proc. 

U.S. Natl. Mus. 18: 446, 23 May 1896d.

= Peromyscus maniculatus clementis Mearns, 1896. See Osgood 

(1909a:96); subspecies recognized by Hall (1981:673).

Holotype: USNM 61117. [Skin and skull]. Adult male. Collected 

27 Aug 1894 by E. A. Mearns. Original number 3819.

Type Locality: [Grinnell (1933:175): Pyramid Cove, southeast 

end] San Clemente Island, [Channel Islands, Los Angeles 

County], California.

Peromyscus texanus medius Mearns

Proc. U.S. Natl. Mus. 18: 4, 25 Mar 1896c. Preprint of Proc. 

U.S. Natl. Mus. 18: 446, 23 May 1896d.

= Peromyscus maniculatus gambelii (Baird, 1857). See Osgood 

(1909a:67); subspecies recognized by Hall (1981:673).

Holotype: USNM 61059. [Skin and skull]. Adult male. Collected 

4 Jun 1894 by E. A. Mearns. Original number 3623.

Type Locality: Nachoguero Valley, [Lyon and Osgood (1909: 

133): near monument number 238, Mexican boundary 

line], Lower California (Baja California), [Mexico].

Peromyscus texanus mesomelas Osgood

Proc. Biol. Soc. Wash. 17: 57, 21 Mar 1904a.

= Peromyscus leucopus mesomelas Osgood, 1904. See Osgood 

(1909a:132); subspecies recognized by Hall (1981:687) and 

Lackey (1999:574).

Holotype: USNM 58210. [Skin and skull]. Adult male. Collected 

20 Jan 1894 by E. W. Nelson and E. A. Goldman. Original 

number [SL: 5705].

Type Locality: Orizaba, [FC: altitude 4,000 ft (1,219 m)], Veracruz, Mexico.

Peromyscus tiburonensis Mearns

Proc. U.S. Natl. Mus. 19: 720, 30 Jul 1897a.

= Peromyscus eremicus tiburonensis Mearns, 1897. See Osgood 

(1909a:250); subspecies recognized by Hall (1981:660) and 

Caire (1999:568).

Holotype: USNM 63186. Skin and skull. Adult male. Collected 

25 Dec 1895 by J. W. Mitchell. Original number 1.
5 2 • smithsonian contributions to zoology

Type Locality: Tiburon Island, Gulf of California, [Sonora], 

Mexico.

Peromyscus tornillo Mearns

Proc. U.S. Natl. Mus. 18: 3, 3 Mar 1896c. Preprint of Proc. U.S. 

Natl. Mus. 18: 445, 23 May 1896d.

= Peromyscus leucopus tornillo Mearns, 1896. See Osgood 

(1909a:125); subspecies recognized by Hall (1981:688) and 

Lackey (1999:574).

Holotype: USNM 20225/35420. Skin and skull. Adult male. 

Collected 18 Feb 1892 by E. A. Mearns and F. X. Holzner. 

Original number 1458.

Type Locality: About 6 mi (10 km) above El Paso, on the Rio 

Grande, [El Paso County], Texas.

Peromyscus truei lagunae Osgood

N. Am. Fauna 28: 172, 17 Apr 1909a.

Subspecies recognized by Hall (1981:703) and Yancey and Jones 

(1999:583).

Holotype: USNM 147004. [Skin and skull]. Adult female. Collected 26 Jan 1906 by E. W. Nelson and E. A. Goldman. 

Original number [SL:19012].

Type Locality: La Laguna, [FC: 5,500 ft (1,676 m)], Laguna 

Mountains, Lower California (Baja California Sur), 

Mexico.

Peromyscus truei preblei V. Bailey

N. Am. Fauna 55: 188, 29 Aug 1936.

Subspecies recognized by Hall (1981:704) and Yancey and Jones 

(1999:583).

Holotype: USNM 78660. [Skin and] skull. Adult male. Collected 

28 Jun 1896 by E. A. Preble. Original number 1079.

Type Locality: Crooked River, 20 mi (32 km) southeast [Hoffmeister (1951:45): 12 mi (19 km) south and 6 mi (10 km) 

east] of Prineville, [Crook County], Oregon.

Peromyscus xenurus Osgood

Proc. Biol. Soc. Wash. 17: 67, 21 Mar 1904a.

= Peromyscus melanophrys xenurus Osgood, 1904. See Baker 

(1952:256); subspecies recognized by Hall (1981:709).

Holotype: USNM 94518. [Skin and skull]. Adult female. Collected 1 Jul 1898 by E. W. Nelson and E. A. Goldman. Original number [SL: 12677].

Type Locality: Durango, [FC: altitude 6,200 ft (1,890 m)], 

Durango, Mexico.

Peromyscus yucatanicus badius Osgood

Proc. Biol. Soc. Wash. 17: 70, 21 Mar 1904a.

Subspecies recognized by Young and Jones (1983:1).

Holotype: USNM 108016. [Skin and skull]. Adult female. Collected 28 Dec 1900 by E. W. Nelson and E. A. Goldman. 

Original number [SL: 14377].

Type Locality: Apazote, [FC: near Yohaltun], Campeche, 

Mexico.

Peromyscus zarhynchus Merriam

Proc. Biol. Soc. Wash. 12: 117, 30 Apr 1898a.

Holotype: USNM 76119. [Skin and skull]. Adult female. Collected [SL and FC: 26 Oct 1895], not 20 Oct as in original 

description, by E. W. Nelson and E. A. Goldman. Original 

number 8606.

Type Locality: Tumbala, [FC: altitude 5,000 ft (1,524 m)], 

Chiapas, Mexico.

Peromyscus zarhynchus cristobalensis Merriam

Proc. Biol. Soc. Wash. 12: 117, 30 Apr 1898a.

= Peromyscus zarhynchus Merriam, 1898. See Osgood (1909a: 

217).

Holotype: USNM 76109. [Skin and skull]. Adult female. Collected 2 Oct 1895 by E. W. Nelson and E. A. Goldman. 

Original number 8536.

Type Locality: San Cristobal, [FC: altitude 9,500 ft (2,896 m)], 

Chiapas, Mexico.

Peromyscus zelotes Osgood

Proc. Biol. Soc. Wash. 17: 67, 21 Mar 1904a.

= Peromyscus gratus gratus Osgood, 1904. See Modi and Lee 

(1984:722); subspecies recognized by Hall (1981:703), 

under P. truei, and Frey (1999b:571).

Holotype: USNM 50430. [Skin and skull]. Old female. Collected 8 Aug 1892 by E. W. Nelson. Original number [SL: 

3056].

Type Locality: Querendaro, [FC: altitude 5,800 ft (1,768 m)], 

Michoacan, Mexico.

Genus Phenacomys Merriam, 1889

Phenacomys celatus Merriam

N. Am. Fauna 2: 33, 30 Oct 1889b.

= Phenacomys ungava ungava Merriam, 1889. See Miller (1897b: 

84); subspecies recognized by McAllister and Hoffmann 

(1988:1), under P. intermedius, and George (1999:619).

Holotype: USNM 186486. Skin and skull, originally in alcohol. Adult male. Collected 10 Jun 1886 by N. A. Comeau. 

Original number not given. Merriam collection number 

2851/5988.

Type Locality: Godbout, P. Q. (Quebec), Canada.

Remarks: Type published as Merriam collection number 

2851/5988; cataloged as USNM 186486 on 19 Oct 1911.

Phenacomys intermedius celsus A. B. Howell

Proc. Biol. Soc. Wash. 36: 158, 1 May 1923b.

Subspecies recognized by Hall (1981:786) and Hallett (1999: 

618).

Holotype: USNM 109103. [Skin and skull]. Adult male. Collected 13 Aug 1901 by J. H. Gaut. Original number 250.

Type Locality: Muir Meadow, altitude 9,300 ft (2,835 m), Tuolumne Meadows, Yosemite National Park, [Tuolumne 

County], California.
number 642 • 5 3

Phenacomys intermedius levis A. B. Howell

Proc. Biol. Soc. Wash. 36: 157, 1 May 1923b.

Subspecies recognized by Hall (1981:786) and Hallett (1999: 618).

Holotype: USNM 72405. [Skin and skull]. Adult male. Collected 

23 May 1895 by A. H. Howell. Original number 13.

Type Locality: Saint Marys Lake, Teton (Glacier) County, 

Montana.

Phenacomys latimanus Merriam

N. Am. Fauna 2: 34, 30 Oct 1889b.

= Phenacomys ungava ungava Merriam, 1889. See A. B. 

Howell (1926:25); subspecies recognized by McAllister 

and Hoffmann (1988:1), under P. intermedius, and George 

(1999:619).

Holotype: USNM 186487. Skin and skull, [SL: skinned from alcohol]. Young adult male. Collected 4 Feb 1883 by L. M. 

Turner. Original number [SL: 300]. Merriam collection 

number 5484/6159.

Type Locality: Fort Chimo (Kuujjuaq), Ungava [District], Hudson Bay, [Quebec, Canada].

Remarks: Type published as Merriam collection number 

5484/6159; cataloged as USNM 186487 on 19 Oct 1911.

Phenacomys mackenzii Preble

Proc. Biol. Soc. Wash. 15: 182, 6 Aug 1902b.

= Phenacomys ungava mackenzii Preble, 1902. See Anderson 

(1942:58); subspecies recognized by McAllister and Hoffmann 

(1988:1), under P. intermedius, and George (1999: 619).

Holotype: USNM 110625. Skin and skull. Adult male. Collected 

29 Jun 1901 by E. A. Preble and A. E. Preble. Original number [FC: EAP] 4271.

Type Locality: Fort Smith, Slave River, near the AthabaskaMackenzie boundary line, [Mackenzie District, Northwest 

Territories], Canada.

Phenacomys orophilus Merriam

N. Am. Fauna 5: 65, 30 Jul 1891a.

= Phenacomys intermedius intermedius Merriam, 1889. See 

A. B. Howell (1926:15); subspecies recognized by Hall 

(1981:786) and Hallett (1999:618).

Holotype: USNM 23856/31256. Skin and skull. Adult female. 

Collected 28 Aug 1890 by C. H. Merriam and V. Bailey. 

Original number [FC: VB] 1710.

Type Locality: Salmon River Mountains [FC: = Lemhi Mountains], near head of Timber Creek, altitude 3,200 m or 

10,500 ft, [FC: west of Junction], Lemhi County, Idaho.

Phenacomys preblei Merriam

Proc. Biol. Soc. Wash. 11: 45, 16 Mar 1897g.

= Phenacomys intermedius intermedius Merriam, 1889. See A. 

B. Howell (1926:15); subspecies recognized by Hall (1981: 

786) and Hallett (1999:618).

Holotype: USNM 74513. [Skin and skull]. Adult male. Collected 

12 Aug 1895 by E. A. Preble. Original number 647.

Type Locality: Longs Peak, [FC: altitude about 9,000 ft 

(2,743 m), Larimer County], Colorado.

Phenacomys truei J. A. Allen

Bull. Am. Mus. Nat. Hist. 6: 331, 7 Nov 1894.

= Phenacomys intermedius intermedius Merriam, 1889. See 

A. B. Howell (1926:15); subspecies recognized by Hall 

(1981:786) and Hallett (1999:618).

Holotype: USNM 3056/36884. [Skin and skull]. Full grown but 

quite young, sex not given. Collected 10 Aug 1857 by W. A. 

Hammond. Original number [SL: 109].

Type Locality: Black Hills, [A. B. Howell (1926:19): now Laramie Mountains, Wyoming].

Remarks: Published under skin number USNM 3056 to which 

skull number USNM 36884 belongs.

Phenacomys ungava Merriam

N. Am. Fauna 2: 35, 30 Oct 1889b.

= Phenacomys ungava ungava Merriam, 1889. See A. B. 

Howell (1926:25); subspecies recognized by McAllister 

and Hoffmann (1988:1), under P. intermedius, and George 

(1999:619).

Holotype: USNM 186488. Skin and skull, from alcohol. Adult 

male. Collected spring 1884 by L. M. Turner. Original number [MCC: 525]. Merriam collection number 5468/6155.

Type Locality: Fort Chimo (Kuujjuaq), Ungava [District], Hudson Bay Territory, [Quebec, Canada].

Remarks: Type published as Merriam collection number 

5468/6155; cataloged as USNM 186488 on 19 Oct 1911.

Genus Phodopus Miller, 1910

Phodopus crepidatus Hollister

Smithson. Misc. Collect. 60(14): 3, 29 Nov 1912.

= Phodopus campbelli (Thomas, 1905). See Pavlinov and Rossolimo (1987:171).

Holotype: USNM 175480. Skin and skull. Adult male. Collected 

28 Jul 1912 by N. Hollister. Original number 4383.

Type Locality: Chuisaya Steppe, 8 mi (13 km) south of 

Kosh-Agatch, altitude 7,300 ft (2,225 m), Altai District 

(Altayskiy), Siberia (Russia).

Genus Phyllotis Waterhouse, 1837

Phyllotis arenarius chilensis Mann

Biologica (Santiago, Chile) 2: 84, Jul 1945.

= Phyllotis xanthopygus chilensis Mann, 1945. See Spotorno 

and Walker (1983:57); subspecies recognized by Steppan et 

al. (2007:811).

Holotype: USNM 391801. [Skin and skull]. Adult male. Collected 28 Mar 1945 by G. Mann. Original number 337.

Type Locality: Los alrededores de Parinacota, [Chile].

Remarks: Published as G. Mann number 337; cataloged as 

USNM 391801 on 8 Dec 1967.
5 4 • smithsonian contributions to zoology

Phyllotis wolffhuegeli Mann

Biologica (Santiago, Chile) 1: 108, Sep 1944.

= Phyllotis xanthopygus vaccarum Thomas, 1912. See Spotorno 

and Walker (1983:57); subspecies recognized by Steppan et 

al. (2007:811).

Holotype: USNM 391813. [Skin and cranium without mandibles]. Adult female. Collected 27 Feb 1944 by G. Mann. 

Original number 225.

Type Locality: Cajon del Rio Volcan, altitude 1,800 m, Lo Valdes, Bocatoma, Region Metropolitana, [Chile].

Remarks: Published as G. Mann number 225; cataloged as 

USNM 391813 on 8 Dec 1967.

Genus Podomys Osgood, 1909

Hesperomys macropus Merriam

N. Am. Fauna 4: 53, 8 Oct 1890d.

= Podomys floridanus (Chapman, 1889). See Carleton 

(1980:115) for generic usage and Osgood (1909a:227) for 

specific synonymy of macropus.

Holotype: USNM 16582/23513. Skin and skull. Adult male. Collected 5 May 1889 by M. M. Green. Original number 72.

Type Locality: [FC: Juno], Lake Worth, [Palm Beach County], 

Florida.

Genus Reithrodon Waterhouse, 1837

Reithrodon hatcheri J. A. Allen

Bull. Am. Mus. Nat. Hist. 19: 191, 9 May 1903.

= Reithrodon auritus pachycephalus (Philippi, 1900). See Osgood (1943:221); subspecies recognized by Pardinas and 

Galliari (2001:1).

Holotype: USNM 84210. [Skin and skull]. Adult male. Collected 

11 Mar 1897 by O. A. Peterson. Original number [SL: 600].

Type Locality: Pacific slope of the Cordilleras, head of the Rio 

Chico de Santa Cruz. Restricted to Perito Moreno National 

Park, about 48°S, 72°W, Santa Cruz Province, Argentina, by 

Pardinas and Galliari (2001:1).

Genus Reithrodontomys Giglioli, 1874

Reithrodon longicauda Baird

Mammals Rep. Explor. Surv. 8(1): 451, 1857.

= Reithrodontomys megalotis longicaudus Baird, 1857. See Grinnell (1913:303); subspecies recognized by Hall (1981:640) 

and Webster (1999:560).

Syntypes: As designated by Lyon and Osgood (1909:114).

USNM 1418/2282. Skin and skull. Female. Original number 

[MC: 231].

USNM 1419/2283. Skin and skull. [MC: Male]. Original number [MC: 232].

USNM 1583/2411. Skin and skull. [MC: Male]. Original number [MC: 239].

USNM 2581 through USNM 2591. In alcohol.

All collected by E. Samuels. Numbers 1418/2282 and 1419/2283 

cataloged 24 Apr 1856; 1583/2411 cataloged 24 Jun 1856; 

the alcoholics cataloged Apr 1857.

Type Locality: Petaluma, [Sonoma County], California.

Remarks: No type specimen designated in the original description. Baird (1857:452) listed the above 14 specimens plus 

one from San Francisco, California. J. A. Allen (1895b:129) 

and Miller and Rehn (1901:97) gave Petaluma as the type 

locality but did not select a type specimen. Lyon and Osgood (1909:114) designated the specimens from Petaluma 

as cotypes = syntypes and fixed the type locality at Petaluma. Numbers 2582, 2584, and 2588 are recorded in the 

museum catalog as “Distributed to C. E. Aiken.” Currently, 

the only syntype present in the collection is the skull of 

USNM 1418/2282. 

Reithrodon megalotis Baird

Mammals Rep. Explor. Surv. 8(1): 451, 1857.

= Reithrodontomys megalotis megalotis Baird, 1857. See A. H. 

Howell (1914:26); subspecies recognized by Hall (1981:640) 

and Webster (1999:560).

Lectotype: USNM 1039/2281 as designated by J. A. Allen 

(1895b:126). Mounted skin and skull. The skin cannot 

be found. Age and sex not given. Collected by C. B. [R.] 

Kennerly, date unknown. [MC: Skin cataloged 6 Dec 1855; 

skull 11 Apr 1856]. No original number.

Type Locality: Between Janos, Sonora, and San Luis Springs. 

Mearns (1907:460) gave the locality as Janos Plain, on 

wagon road between Janos, Chihuahua, [Mexico], and San 

Luis Springs, Monument No. 66, Grant (Hidalgo) County, 

New Mexico.

Remarks: Baird (1857:541) had two specimens from the above 

locality; J. A. Allen (1895b:126) stated the specimen given 

above should be considered the type, which action effectively designated a lectotype (ICZN, 1999: Article 74.5). A 

note in the museum catalog indicates that the skin was missing in Aug 1898.

Reithrodon montanus Baird

Proc. Acad. Nat. Sci. Phila. 7: 335, reported favorably for publication 24 Apr 1855b. Receipt acknowledged 20 Jul 1855 by 

the Smithsonian Institution.

= Reithrodontomys montanus montanus Baird, 1855. See Benson (1935:141); subspecies recognized by Hall (1981:636) 

and Dowler (1999:561).

Holotype: USNM 441/1306. [Mounted skin and skull. Age and 

sex not given]. Collected [J. A. Allen (1895b:24): 29 or 30 

Aug 1853 by F. Kreutzfeldt on Capt. E. G. Beckworth’s 

expedition from Westport, Missouri to the Pacific coast in 

1853–1854]. Original number [Baird (1857:450): 13].

Type Locality: In the vicinity of the Rocky Mountains, Lat. 38°. 

Baird (1857:450) subsequently repeated Rocky Mountains 

but gave the latitude as 39°. J. A. Allen (1895b:125) restricted the locality to upper part of the San Luis Valley, 
number 642 • 5 5

[Saguache County, Colorado]. Cary (1911:108) determined 

from the expedition itinerary that the type locality appears 

to have been on the east side of the San Luis Valley about 

one-third the distance between old Fort Massachusetts, now 

Fort Garland, and Cochetopa Pass, probably not far from 

the San Luis Lakes. He collected what he considered to be 

topotypes of R. montanus near Medano Creek on Medano 

Springs Ranch, 15 mi (24 km) northeast of Mosca and a 

short distance northeast of the San Luis Lakes.

Remarks: Type not designated by number in the original description, but the description was based on a single specimen; 

the above listed number is given in Baird (1857:13) and 

becomes the holotype as fixed by monotypy (ICZN, 1999: 

Article 73.1.2).

Reithrodontomys albescens Cary

Proc. Biol. Soc. Wash. 16: 53, 6 May 1903.

= Reithrodontomys montanus albescens Cary, 1903. See Benson 

(1935:141); subspecies recognized by Hall (1981:635) and 

Dowler (1999:561).

Holotype: USNM 116358. [Skin and skull]. Adult male. Collected 

31 Oct 1901 by M. Cary. Original number 411. [SL: 3431-X].

Type Locality: 18 mi (29 km) northwest of Kennedy, [Cherry 

County], Nebraska.

Reithrodontomys alleni A. H. Howell

N. Am. Fauna 36: 59, 5 Jun 1914.

= Reithrodontomys sumichrasti luteolus, A. H. Howell, 1914. See 

Hooper (1952:77); subspecies recognized by Hall (1981: 643).

Holotype: USNM 71563. [Skin and skull]. Adult male. Collected 

27 Mar 1895 by E. W. Nelson and E. A. Goldman. Original 

number 7749.

Type Locality: Mountains near Ozolotepec, altitude 10,000 ft 

(3,048 m), Oaxaca, Mexico.

Reithrodontomys amoles A. H. Howell

N. Am. Fauna 36: 40, 5 Jun 1914.

= Reithrodontomys megalotis amoles A. H. Howell, 1914. See 

Hooper (1952:64); subspecies recognized by Hall (1981: 

638) and Webster (1999:560).

Holotype: USNM 81234. [Skin and skull]. Adult female. Collected 20 Sep 1896 by E. W. Nelson and E. A. Goldman. 

Original number 10169.

Type Locality: Pinal de Amoles, [FC: altitude about 8,000 ft 

(2,438 m)], Queretaro, Mexico.

Reithrodontomys chrysopsis Merriam

Proc. Biol. Soc. Wash. 13: 152, 13 Jun 1900b.

= Reithrodontomys chrysopsis chrysopsis Merriam, 1900. 

See A. H. Howell (1914:66); subspecies recognized by 

Gonzalez-Ruiz et al. (2007:242).

Holotype: USNM 52031. [Skin and skull]. Adult male. Collected 

25 Feb 1893 by E. W. Nelson and E. A. Goldman. Original 

number [FC: EWN] 4405.

Type Locality: Mount Popocatepetl, [SL and FC: altitude 11,500 

ft (3,505 m)], [Mexico], Mexico.

Reithrodontomys chrysopsis tolucae Merriam

Proc. Wash. Acad. Sci. 3: 549, 29 Nov 1901e.

= Reithrodontomys chrysopsis chrysopsis Merriam, 1900. See 

Hooper (1952:86); subspecies recognized by Gonzalez-Ruiz 

et al. (2007:242).

Holotype: USNM 55895. [Skin and skull]. Adult female. Collected 10 Sep 1893 by E. W. Nelson and E. A. Goldman. 

Original number [FC: EWN] 5454.

Type Locality: [SL and FC: North slope] Volcan Toluca, altitude 

11,500 ft (3,505 m), State of Mexico, [Mexico].

Reithrodontomys colimae Merriam

Proc. Wash. Acad. Sci. 3: 551, 29 Nov 1901e.

= Reithrodontomys chrysopsis chrysopsis Merriam, 1900. See 

Hooper (1952:86); subspecies recognized by Gonzalez-Ruiz 

et al. (2007:242).

Holotype: USNM 33671/45706. Skin and skull. Adult male. 

Collected 13 Apr 1892, not 21 Apr as in original description, by E. W. Nelson and E. A. Goldman. Original number 

[FC: EWN] 2447.

Type Locality: Near timberline, Sierra Nevada de Colima, altitude [FC: about] 12,000 ft (3,658 m), Jalisco, Mexico.

Reithrodontomys colimae nerterus Merriam

Proc. Wash. Acad. Sci. 3: 551, 29 Nov 1901e.

= Reithrodontomys sumichrasti nerterus Merriam, 1901. See 

Hooper (1952:74); subspecies recognized by Hall (1981:643).

Holotype: USNM 33680/45715. Skin and skull. Adult female. 

Collected 21 Apr 1892 by E. W. Nelson and E. A. Goldman. 

Original number [FC: EWN] 2514.

Type Locality: Foothill region of Sierra Nevada de Colima, altitude [FC: about] 6,500 ft (1,981 m), Jalisco, Mexico.

Reithrodontomys costaricensis jalapae Merriam

Proc. Wash. Acad. Sci. 3: 552, 29 Nov 1901e.

= Reithrodontomys mexicanus mexicanus (Saussure, 1860). See 

A. H. Howell (1914:70); subspecies recognized by Hall 

(1981:650).

Holotype: USNM 108538. [Skin and skull]. Adult male. Collected 10 May 1901 by E. W. Nelson and E. A. Goldman. 

Original number 14712.

Type Locality: Jalapa, altitude 4,000 ft (1,219 m), Veracruz, Mexico.

Reithrodontomys difficilis Merriam

Proc. Wash. Acad. Sci. 3: 556, 29 Nov 1901e.

= Reithrodontomys fulvescens difficilis Merriam, 1901. See 

A. H. Howell (1914:50); subspecies recognized by Hall 

(1981:645) and Cameron (1999b:557).

Holotype: USNM 63735. [Skin and skull]. Adult male. Collected 

20 Feb 1894 by E. W. Nelson and E. A. Goldman. Original 

number 5868.
5 6 • smithsonian contributions to zoology

Type Locality: Orizaba, altitude 4,500 ft (1,372 m) [FC: 4,200 ft 

(1,280 m)], Veracruz, Mexico.

Reithrodontomys dorsalis Merriam

Proc. Wash. Acad. Sci. 3: 557, 29 Nov 1901e.

= Reithrodontomys sumichrasti dorsalis Merriam, 1901. See 

Hooper (1952:78); subspecies recognized by Hall (1981:643).

Holotype: USNM 77009. [Skin and skull]. Adult male. Collected 

14 Jan 1896 by E. W. Nelson and E. A. Goldman. Original 

number 9112.

Type Locality: Calel, [FC: 10,200 ft (3,109 m)], [Quetzaltenango], Guatemala.

Reithrodontomys dychei nebrascensis J. A. Allen

Bull. Am. Mus. Nat. Hist. 7: 122, 21 May 1895b.

= Reithrodontomys megalotis dychei J. A. Allen, 1895. See 

A. H. Howell (1914:30); subspecies recognized by Hall 

(1981:638) and Webster (1999:560).

Holotype: USNM 18624/25426. Skin and skull. Adult male. Collected 10 Apr 1890 by V. Bailey. Original number [SL: 1042].

Type Locality: Kennedy, [FC: probably 2,700 ft (823 m), Cherry 

County], Nebraska.

Reithrodontomys fulvescens chiapensis A. H. Howell

N. Am. Fauna 36: 53, 5 Jun 1914.

Subspecies recognized by Hall (1981:645) and Cameron (1999b: 

557).

Holotype: USNM 132865. [Skin and skull]. Adult male. Collected 2 May 1904 by [E. W.] Nelson and [E. A.] Goldman. 

Original number 16741.

Type Locality: Canjob, [FC: altitude 5,300 ft (1,615 m)], Chiapas, Mexico.

Reithrodontomys fulvescens mustelinus A. H. Howell

N. Am. Fauna 36: 54, 5 Jun 1914.

Subspecies recognized by Gonzalez-Ruiz et al. (2007:243).

Holotype: USNM 71549. [Skin and skull]. Adult female. Collected 18 Feb 1895 by [E. W.] Nelson and [E. A.] Goldman. 

Original number 7483.

Type Locality: Llano Grande, [FC: altitude 200 ft (61 m)], Oaxaca, Mexico.

Reithrodontomys fulvescens nelsoni A. H. Howell

N. Am. Fauna 36: 53, 5 Jun 1914.

Subspecies recognized by Hall (1981:647) and Cameron (1999b: 

557).

Holotype: USNM 33409/45432. Skin and skull. Adult female. 

Collected 9 Mar 1892 by E. W. Nelson. Original number 

2050.

Type Locality: Colima, [FC: altitude 1,700 ft (518 m)], Colima, 

Mexico.

Reithrodontomys goldmani Merriam

Proc. Wash. Acad. Sci. 3: 552, 29 Nov 1901e.

= Reithrodontomys mexicanus mexicanus (Saussure, 1860). See 

Hooper (1952:140); subspecies recognized by Hall (1981: 

650).

Holotype: USNM 93096. [Skin and skull]. Adult female. Collected 12 Feb 1898 by E. A. Goldman. Original number 

[FC: Nelson and Goldman] 12153.

Type Locality: [FC: Near] Metlaltoyuca, altitude [FC: about] 

800 ft (244 m), Puebla, [Mexico].

Reithrodontomys griseoflavus Merriam

Proc. Wash. Acad. Sci. 3: 553, 29 Nov 1901e.

= Reithrodontomys fulvescens tenuis J. A. Allen, 1899. See 

A. H. Howell (1914:45); subspecies recognized by Hall 

(1981:645) and Cameron (1999b:557).

Holotype: USNM 82196. [Skin and skull]. Adult male. Collected 

9 Feb 1897 by E. W. Nelson and E. A. Goldman. Original 

number 10501.

Type Locality: Ameca, altitude 4,000 ft (1,219 m), Jalisco, 

Mexico.

Reithrodontomys griseoflavus helvolus Merriam

Proc. Wash. Acad. Sci. 3: 554, 29 Nov 1901e.

= Reithrodontomys fulvescens helvolus Merriam, 1901. See 

A. H. Howell (1914:52); subspecies recognized by Hall 

(1981:647) and Cameron (1999b:557).

Holotype: USNM 68387. [Skin and skull]. Adult female. Collected 14 Aug 1894 by E. W. Nelson and E. A. Goldman. 

Original number 6576.

Type Locality: Oaxaca City, [FC: altitude 5,000 ft (1,524 m)], 

Oaxaca, Mexico.

Reithrodontomys griseus V. Bailey

N. Am. Fauna 25: 106, 24 Oct 1905.

= Reithrodontomys montanus griseus V. Bailey, 1905. See Benson (1935:141); subspecies recognized by Hall (1981:636) 

and Dowler (1999:561).

Holotype: USNM 87852. [Skin and skull]. Adult male. Collected 4 

Mar 1897 by H. P. Attwater. Original number 1068. 371-X.

Type Locality: San Antonio, [Bexar County], Texas.

Reithrodontomys hirsutus Merriam

Proc. Wash. Acad. Sci. 3: 553, 29 Nov 1901e.

Holotype: USNM 82200. [Skin and skull]. Old male. Collected 

15 Feb 1897 by E. W. Nelson and E. A. Goldman. Original 

number 10537.

Type Locality: Ameca, altitude 5,500 ft (1,676 m), Jalisco, Mexico.

Reithrodontomys humulis virginianus A. H. Howell

J. Mammal. 21: 346, 13 Aug 1940.

Subspecies recognized by Hall (1981:637) and Schmidly (1999b: 

558).

Holotype: USNM 263815. [Skin and skull]. Adult male. Collected 15 Feb 1938 by J. B. Lewis. Original number 158. 

28258-X.
number 642 • 5 7

Type Locality: Amelia [FC: Courthouse, Amelia County], 

Virginia.

Reithrodontomys klamathensis Merriam

N. Am. Fauna 16: 93, 28 Oct 1899.

= Reithrodontomys megalotis longicaudus (Baird, 1857). See 

A. H. Howell (1914:33); subspecies recognized by Hall 

(1981:640) and Webster (1999:560).

Holotype: USNM 95444. [Skin and skull]. Adult male. Collected 

18 Sep 1898 by W. H. Osgood. Original number 281.

Type Locality: Big Spring (Mayten), Shasta Valley, [Siskiyou 

County], California.

Reithrodontomys levipes Merriam

Proc. Wash. Acad. Sci. 3: 554, 29 Nov 1901e.

= Reithrodontomys hirsutus Merriam, 1901. See Hooper (1952: 

124). 

Holotype: USNM 88057. [Skin and skull]. Adult male. Collected 

30 Mar 1897 by E. W. Nelson and E. A. Goldman. Original 

number 10839.

Type Locality: San Sebastian, altitude 3,000 ft (914 m), Jalisco, 

Mexico.

Reithrodontomys levipes otus Merriam

Proc. Wash. Acad. Sci. 3: 555, 29 Nov 1901e.

= Reithrodontomys sumichrasti nerterus Merriam, 1901. See 

Hooper (1952:74); subspecies recognized by Hall (1981:643).

Holotype: USNM 33670/45705. Skin and skull. Adult female. 

Collected 11 Apr 1892 by E. W. Nelson and E. A. Goldman. 

Original number [FC: EWN] 2439.

Type Locality: Foothill region of Sierra Nevada de Colima, altitude [FC: about] 6,500 ft (1,981 m), Jalisco, Mexico.

Reithrodontomys levipes toltecus Merriam

Proc. Wash. Acad. Sci. 3: 555, 29 Nov 1901e.

= Reithrodontomys fulvescens toltecus Merriam, 1901. See A. H. 

Howell (1914:51); subspecies recognized by Gonzalez-Ruiz 

et al. (2007:242).

Holotype: USNM 50746. [Skin and skull]. Adult female. Collected 1 Dec 1892 by E. W. Nelson and E. A. Goldman. 

Original number [FC: EWN] 3935.

Type Locality: Tlalpam, [FC: altitude 7,600 ft (2,316 m)], Valley 

of Mexico, [Distrito Federal, Mexico].

Reithrodontomys megalotis caryi A. H. Howell

J. Mammal. 16: 143, 15 May 1935.

= Reithrodontomys megalotis aztecus J. A. Allen, 1893. See 

Jones and Mursaloglu (1961:19); subspecies recognized by 

Hall (1981:638) and Webster (1999:560).

Holotype: USNM 150714. Skin and skull. Adult female. Collected 4 Nov 1907 by M. Cary. Original number 1232.

Type Locality: Medano Ranch, 15 mi (24 km) northeast of 

Mosca, [FC: altitude 7,600 ft (2,316 m)], Alamosa County, 

Colorado.

Reithrodontomys megalotis deserti J. A. Allen

Bull. Am. Mus. Nat. Hist. 7: 127, 21 May 1895b.

= Reithrodontomys megalotis megalotis (Baird, 1857). See A. 

H. Howell (1914:26); subspecies recognized by Hall (1981: 

640) and Webster (1999:560).

Holotype: USNM 27720/39819. Skin and skull. Adult female. 

Collected 16 Mar 1891 by F. Stephens. Original number 

[SL: 149].

Type Locality: [FC: Lower end] Oasis Valley, Nye County, 

Nevada.

Reithrodontomys megalotis nigrescens A. H. Howell

N. Am. Fauna 36: 32, 5 Jun 1914.

= Reithrodontomys megalotis megalotis (Baird, 1857). See Hall 

(1946:499); subspecies recognized by Hall (1981:640) and 

Webster (1999:560).

Holotype: USNM 201616. [Skin and skull]. Adult male. Collected 9 Jun 1913 by L. E. Wyman. Original number 98.

Type Locality: Payette, [Payette County], Idaho.

Reithrodontomys megalotis obscurus Merriam

Proc. Wash. Acad. Sci. 3: 558, 29 Nov 1901e.

= Reithrodontomys zacatecae Merriam, 1901. See Musser and 

Carleton (1993:743).

Holotype: USNM 95277. [Skin and skull]. Adult male. Collected 

26 Aug 1898 by E. W. Nelson and E. A. Goldman. Original 

number 12900.

Type Locality: Sierra Madre, near Guadalupe y Calvo, [FC: altitude 7,000 ft (2,134 m)], Chihuahua, Mexico.

Reithrodontomys megalotis ravus Goldman

J. Mammal. 20: 355, 14 Aug 1939.

Subspecies recognized by Hall (1981:641) and Webster (1999:560).

Holotype: USNM 263963. Skin and skull. Adult male. Collected 

26 Jun 1938 by W. H. Marshall. Original number [SL: 161].

Type Locality: Sandbar at spring on lake shore, north end 

of Stansbury Island, Great Salt Lake, altitude 4,250 ft 

(1,295 m), [Tooele County], Utah.

Reithrodontomys megalotis zacatecae Merriam

Proc. Wash. Acad. Sci. 3: 557, 29 Nov 1901e.

= Reithrodontomys zacatecae Merriam, 1901. See Musser and 

Carleton (1993:743).

Holotype: USNM 91910. [Skin and skull]. Adult female. Collected 6 Dec 1897 by E. W. Nelson and E. A. Goldman. 

Original number 11859.

Type Locality: Valparaiso Mountains, [FC: altitude 8,200 ft 

(2,499 m)], Zacatecas, Mexico.

Reithrodontomys merriami J. A. Allen

Bull. Am. Mus. Nat. Hist. 7: 119, 21 May 1895b.

= Reithrodontomys humulis merriami J. A. Allen, 1895. See 

A. H. Howell (1914:21); subspecies recognized by Hall 

(1981:637) and Schmidly (1999b:558).
5 8 • smithsonian contributions to zoology

Holotype: USNM 32832/44853. Skin and skull. Adult male. 

Collected 15 Mar 1892 by W. Lloyd. Original number [SL: 

1162].

Type Locality: Austin Bayou, near [FC: 10 mi (16 km) southwest 

of] Alvin, Brazoria County, Texas.

Reithrodontomys mexicanus aurantius J. A. Allen

Bull. Am. Mus. Nat. Hist. 7: 137, 21 May 1895b.

= Reithrodontomys fulvescens aurantius J. A. Allen, 1895. See 

A. H. Howell (1914:48); subspecies recognized by Hall 

(1981:645) and Cameron (1999b:557).

Holotype: USNM 33963/45998. Skin and skull. Adult male. 

Collected 24 May 1892 by R. J. Thompson. Original number [SL: 174].

Type Locality: Lafayette, [Lafayette Parish], Louisiana.

Reithrodontomys mexicanus lucifrons A. H. Howell

Proc. Biol. Soc. Wash. 45: 125, 30 Jul 1932.

Subspecies recognized by Hall (1981:650).

Holotype: USNM 257086. Skin and skull. Adult female. Collected 

7 Feb 1932 by C. F. Underwood. Original number 709.

Type Locality: Cerro Cantoral, [Francisco Morazan], Honduras.

Reithrodontomys mexicanus minusculus A. H. Howell

Proc. Biol. Soc. Wash. 45: 125, 30 Jul 1932.

= Reithrodontomys mexicanus lucifrons A. H. Howell, 1932. See 

Hooper (1952:150); subspecies recognized by Hall (1981:650).

Holotype: USNM 257087. Skin and skull. Adult male. Collected 

27 Feb 1932 by C. F. Underwood. Original number 797.

Type Locality: Comayabuela (Comayaguela), just south of Tegucigalpa, [Francisco Morazan], Honduras.

Reithrodontomys microdon Merriam

Proc. Wash. Acad. Sci. 3: 548, 29 Nov 1901e.

= Reithrodontomys microdon microdon Merriam, 1901. See 

A. H. Howell (1914:80); subspecies recognized by Hall 

(1981:653).

Holotype: USNM 76923. [Skin and skull]. Adult female. Collected 31 Dec 1895 by E. W. Nelson and E. A. Goldman. 

Original number 8977.

Type Locality: Todos Santos, altitude 10,000 ft (3,048 m), [Huehuetenango], Guatemala.

Reithrodontomys microdon albilabris Merriam

Proc. Wash. Acad. Sci. 3: 549, 29 Nov 1901e.

Subspecies recognized by Hall (1981:653).

Holotype: USNM 68393. [Skin and skull]. Adult female. Collected 25 Aug 1894 by E. W. Nelson and E. A. Goldman. 

Original number 6629.

Type Locality: Cerro San Felipe, altitude 10,000 ft (3,048 m), 

Oaxaca, Mexico.

Reithrodontomys orizabae Merriam

Proc. Wash. Acad. Sci. 3: 550, 29 Nov 1901e.

= Reithrodontomys chrysopsis perotensis Merriam, 1901. See 

Hooper (1952:89); subspecies recognized by Hall (1981: 

642).

Holotype: USNM 53626. [Skin and skull]. Adult male. Collected 

24 Apr 1893 by E. W. Nelson and E. A. Goldman. Original 

number [FC: EWN] 4732.

Type Locality: Mount Orizaba, altitude 9,500 ft (2,896 m), 

Puebla, Mexico.

Reithrodontomys perotensis Merriam

Proc. Wash. Acad. Sci. 3: 550, 29 Nov 1901e.

= Reithrodontomys chrysopsis perotensis Merriam, 1901. See 

Davis (1944b:393); subspecies recognized by Hall (1981: 

642).

Holotype: USNM 54424. [Skin and skull]. Adult male. Collected 

31 May 1893 by E. W. Nelson and E. A. Goldman. Original 

number [FC: EWN] 4912.

Type Locality: Cofra (Cofre) de Perote, altitude 9,500 ft 

(2,896 m), Veracruz, Mexico.

Reithrodontomys rufescens luteolus A. H. Howell

N. Am. Fauna 36: 57, 5 Jun 1914.

= Reithrodontomys sumichrasti luteolus A. H. Howell, 1914. See 

Hooper (1952:77); subspecies recognized by Hall (1981:643).

Holotype: USNM 71558. [Skin and skull]. Adult female. Collected 28 Feb 1895 by E. W. Nelson and E. A. Goldman. 

Original number 7579.

Type Locality: Juquila, altitude 5,000 ft (1,524 m), Oaxaca, 

Mexico.

Reithrodontomys saturatus alticolus Merriam

Proc. Wash. Acad. Sci. 3: 556, 29 Nov 1901e.

= Reithrodontomys megalotis alticolus Merriam, 1901. See A. 

H. Howell (1914:37); subspecies recognized by Hall (1981: 

638) and Webster (1999:560).

Holotype: USNM 68392. [Skin and skull]. Old male. Collected 

24 Aug 1894 by E. W. Nelson and E. A. Goldman. Original 

number 6623.

Type Locality: Cerro San Felipe, near Oaxaca City, altitude 

10,000 ft (3,048 m), Oaxaca, Mexico.

Reithrodontomys saturatus cinereus Merriam

Proc. Wash. Acad. Sci. 3: 556, 29 Nov 1901e.

= Reithrodontomys megalotis saturatus J. A. Allen and Chapman, 1897. See Hooper (1952:55); subspecies recognized by 

Hall (1981:641) and Webster (1999:560).

Holotype: USNM 53623. [Skin and skull]. Adult male. Collected 

13 Apr 1893 by E. W. Nelson and E. A. Goldman. Original 

number [FC: EWN] 4659.

Type Locality: Chalchicomula, [FC: altitude 8,200 ft (2,499 m)], 

Puebla, Mexico.

Reithrodontomys tenuirostris Merriam

Proc. Wash. Acad. Sci. 3: 547, 29 Nov 1901e.
number 642 • 5 9

Holotype: USNM 76919. [Skin and skull]. Adult male. Collected 

29 Dec 1895 by E. W. Nelson and E. A. Goldman. Original 

number 8917.

Type Locality: Todos Santos, altitude 10,000 ft (3,048 m), [Huehuetenango], Guatemala.

Reithrodontomys tenuirostris aureus Merriam

Proc. Wash. Acad. Sci. 3: 548, 29 Nov 1901e.

= Reithrodontomys tenuirostris Merriam, 1901. See Hooper 

(1952:171).

Holotype: USNM 76939. [Skin and skull]. Adult female. Collected 15 Jan 1896 by E. W. Nelson and E. A. Goldman. 

Original number 9140.

Type Locality: Calel, altitude 10,200 ft (3,109 m), [Quetzaltenango], Guatemala.

Genus Rheomys Thomas, 1906

Rheomys raptor Goldman

Smithson. Misc. Collect. 60(2): 7, 20 Sep 1912a.

= Rheomys raptor raptor Goldman, 1912. See Voss (1988:357).

Holotype: USNM 179028. [Skin and skull]. Adult male. Collected 

28 Apr 1912 by E. A. Goldman. Original number 21658.

Type Locality: Near head of Rio Limon, altitude 4,500 ft 

(1,372 m), Mount Pirri, [Darien], eastern Panama.

Genus Rhipidomys Tschudi, 1845

Rhipidomys scandens Goldman

Smithson. Misc. Collect. 60(22): 8, 28 Feb 1913.

= Rhipidomys latimanus latimanus Tomes, 1860. See Tribe 

(1996:221).

Holotype: USNM 178987. [Skin and skull]. Old adult female. 

Collected 25 Apr 1912 by E. A. Goldman. Original number 

21640.

Type Locality: Near head of Rio Limon, altitude 5,000 ft 

(1,524 m) [SL and FC: 5,100 ft (1,554 m)], Mount Pirri, 

[Darien], eastern Panama.

Remarks: USNM 178937 was incorrectly given as the catalog 

number in the original description.

Rhipidomys wetzeli A. L. Gardner

In Redford and Eisenberg, eds., Advances in Neotropical Mammalogy, p. 417, 6 Feb 1990.

Holotype: USNM 560697. Skin and skull. Adult female. Collected 2 Dec 1984 by A. L. Gardner. Original number 14326.

Type Locality: Camp VII, 00°50′40′′N, 65°58′10′′W, 1,800 m 

[SL: 1,850 m], Cerro de la Neblina, Territorio Federal Amazonas, Venezuela.

Genus Scotinomys Thomas, 1913

Scotinomys teguina subnubilis Goldman

Proc. Biol. Soc. Wash. 48: 141, 22 Aug 1935.

= Scotinomys teguina teguina Alston, 1876. See Hooper (1972: 

15); subspecies recognized by Hall (1981:735).

Holotype: USNM 76353. Skin and skull. Adult male. Collected 

22 Aug 1895 by [E. W.] Nelson and [E. A.] Goldman. Original number 8371.

Type Locality: Ocuilapa, 10 mi (16 km) northwest of Ocozucuautla (Ocozocuautla) and about 25 mi (40 km) west of 

Tuxtla Gutierrez, altitude 3,500 ft (1,067 m), Chiapas, 

Mexico.

Genus Sigmodon Say and Ord, 1825

Sigmodon alleni V. Bailey

Proc. Biol. Soc. Wash. 15: 112, 2 Jun 1902b.

= Sigmodon alleni alleni V. Bailey, 1902. See Baker (1969:192); 

subspecies recognized by Hall (1981:743).

Holotype: USNM 88227. [Skin and skull]. Adult male. Collected 

15 Mar 1897 by E. W. Nelson and E. A. Goldman. Original 

number 10708.

Type Locality: [FC: Near] San Sebastian, [FC: altitude 3,800 ft 

(1,158 m)], Mascota, Jalisco, Mexico.

Sigmodon alticola V. Bailey

Proc. Biol. Soc. Wash. 15: 116, 2 Jun 1902b.

= Sigmodon leucotis alticola V. Bailey, 1902. See Baker (1969: 

222); subspecies recognized by Hall (1981:744).

Holotype: USNM 68231. [Skin and skull]. Adult male. Collected 

24 Aug 1894 by E. W. Nelson and E. A. Goldman. Original 

number 6624.

Type Locality: Cerro San Felipe, altitude 10,000 ft (3,048 m), 

Oaxaca, Mexico.

Sigmodon alticola amoles V. Bailey

Proc. Biol. Soc. Wash. 15: 116, 2 Jun 1902b.

= Sigmodon leucotis leucotis V. Bailey, 1902. See Baker (1969: 

220); subspecies recognized by Hall (1981:744).

Holotype: USNM 81430. [Skin and skull]. Adult male. Collected 

18 Sep 1896, not 1898 as given in original description, by 

E. W. Nelson and E. A. Goldman. Original number 10161.

Type Locality: Pinal de Amoles, altitude [FC: about] 7,000 ft 

(2,134 m), Queretaro, Mexico.

Sigmodon berlandieri Baird

Proc. Acad. Nat. Sci. Phila. 7: 333, reported favorably for publication 24 Apr 1855a. Receipt acknowledged 15 Jun 1855 

by the American Philosophical Society.

= Sigmodon hispidus berlandieri Baird, 1855. See Coues (1877: 

42); subspecies recognized by Hall (1981:736) and Cameron (1999c:594).

Lectotype: USNM 566/1687 as designated by Coues (1877:39). 

Skin, not found, and skull. [Lyon and Osgood (1909:109): 

Adult] female. Collected 1853 by D. N. Couch. No original 

number.

Type Locality: Rio Nasas (Nazas), Coahuila, Mexico.
6 0 • smithsonian contributions to zoology

Remarks: Baird (1855:333) indicated that berlandieri was based 

on specimens collected between San Antonio and El Paso 

by Mr. Clark and in northern Mexico by Lieut. Couch, 

but he did not select a type. In a subsequent publication, 

Baird (1857:506) listed several specimens, and in a table 

on page 505 he gave measurements for USNM 566. Bailey 

(1902b:107) stated that this specimen is generally considered the type. Lyon and Osgood (1909:109) and Poole and 

Schantz (1942:351) followed Bailey in listing it as the type. 

However, Coues (1877:39) had earlier, in his Table VIII listing measurements of Sigmodon, footnoted USNM 566 as 

the “type of S. ‘berlandieri’ Baird,” which action effectively 

fixed the lectotype (ICZN, 1999: Article 74.5). Notes in the 

museum catalog state that the specimen was originally in 

alcohol, skinned, and the body remained in alcohol and that 

neither were found 13 Feb 1903.

Sigmodon guerrerensis Nelson and Goldman

Proc. Biol. Soc. Wash. 46: 196, 26 Oct 1933.

= Sigmodon alleni vulcani J. A. Allen, 1906. See Baker (1969:194); 

subspecies recognized by Hall (1981:744).

Holotype: USNM 126936. Skin and skull. Adult female. Collected 21 May 1903 by [E. W.] Nelson and [E. A.] Goldman. 

Original number 16467.

Type Locality: Omilteme, altitude 8,000 ft (2,438 m), Guerrero, 

Mexico.

Sigmodon hispidus alfredi Goldman and Gardner

J. Mammal. 28: 57, 15 Feb 1947.

Subspecies recognized by Hall (1981:736) and Cameron 

(1999c:594).

Holotype: USNM 274918. Skin and skull. Adult male. Collected 

19 Mar 1946 by R. L. Landburg. Original number 1056. 

32296-X.

Type Locality: I. N. Pruitt Farm, near William’s Corner, 11 mi 

(18 km) north of Springfield, Baca County, Colorado.

Sigmodon hispidus confinis Goldman

Proc. Biol. Soc. Wash. 31: 21, 16 May 1918b.

Subspecies recognized by Hall (1981:737) and Cameron 

(1999c:594).

Holotype: USNM 204241. [Skin and skull]. Adult male. Collected 27 Jul 1914 by J. L. Peters. Original number 77.

Type Locality: Safford, altitude 2,900 ft (834 m), [Graham 

County], Arizona.

Sigmodon hispidus eremicus Mearns

Proc. U.S. Natl. Mus. 20: 4, 5 Mar 1897b. Preprint of Proc. U.S. 

Natl. Mus. 20: 504, 19 Jan 1898.

Subspecies recognized by Hall (1981:737) and Cameron 

(1999c:594).

Holotype: USNM 60319. Skin and skull. Adult male. Collected 

24 Mar 1894 by E. A. Mearns and F. X. Holzner. Original 

number 3367.

Type Locality: Cienega Well, 30 mi (48 km) south of Monument 

No. 204, Mexican boundary line, on the left bank of the 

Colorado River, Sonora, [Mexico].

Sigmodon hispidus floridanus A. H. Howell

Proc. Biol. Soc. Wash. 56: 73, 16 Jun 1943.

Subspecies recognized by Hall (1981:737) and Cameron (1999c: 

594).

Holotype: USNM 261264. Skin and skull. Adult female. Collected 1 Mar 1936 by A. H. Howell. Original number 

2813.

Type Locality: Canal Point, Palm Beach County, Florida.

Sigmodon hispidus jacksoni Goldman

Proc. Biol. Soc. Wash. 31: 22, 16 May 1918b.

= Sigmodon arizonae jacksoni Goldman, 1918. See Hall (1981: 

742); subspecies recognized by Davis and Young (1999: 

592).

Holotype: USNM 214121. [Skin and skull]. Subadult male. 

Collected 8 Jul 1916 by H. H. T. Jackson. Original number 

456.

Type Locality: 3 mi (5 km) north of Fort Whipple, altitude 5,000 

ft (1,524 m), near Prescott, [Yavapai County], Arizona.

Sigmodon hispidus komareki M. C. Gardner

Proc. Biol. Soc. Wash. 61: 97, 16 Jun 1948a.

Subspecies recognized by Hall (1981:739) and Cameron (1999c: 

594).

Holotype: USNM 207210. Skin and skull. Adult male. Collected 

9 Mar 1915 by L. J. Goldman. Original number 2128.

Type Locality: Woodville, altitude 616 ft (188 m), Jackson 

County, Alabama.

Sigmodon hispidus major V. Bailey

Proc. Biol. Soc. Wash. 15: 109, 2 Jun 1902b.

= Sigmodon arizonae major V. Bailey, 1902. See Hall (1981:742); 

subspecies recognized by Davis and Young (1999:592).

Holotype: USNM 96275. [Skin and skull]. Adult male. Collected 

20 Oct 1898 by E. W. Nelson and E. A. Goldman. Original 

number 13154.

Type Locality: Sierra de Choix, 50 mi (80 km) [Burt (1938:62): 

probably only 10 mi (16 km) or 15 mi (24 km)] northeast of 

Choix, [FC: altitude 4,500 ft–5,000 ft (1,373 m–1,524 m)], 

Sinaloa, Mexico.

Sigmodon hispidus microdon V. Bailey

Proc. Biol. Soc. Wash. 15: 111, 2 Jun 1902b.

= Sigmodon toltecus (Saussure, 1860). See Musser and Carleton 

(2005:1177).

Holotype: USNM 108467. [Skin and skull]. Adult male. Collected 13 Mar 1901 by E. W. Nelson and E. A. Goldman. 

Original number 14581.

Type Locality: Puerto Morelos, Yucatan (Quintana Roo), [FC: 

Mexico].
number 642 • 6 1

Sigmodon hispidus pallidus Mearns

Proc. U.S. Natl. Mus. 20: 4, 5 Mar 1897b. Preprint of Proc. U.S. 

Natl. Mus. 20: 504, 19 Jan 1898.

= Sigmodon hispidus berlandieri Baird, 1855. See V. Bailey 

(1902b:106); subspecies recognized by Hall (1981:736) and 

Cameron (1999c:594).

Holotype: USNM 20103/35464. Skin and skull. Adult male. 

Collected 19 Feb1892 by E. A. Mearns and F. X. Holzner. 

Original number 1461.

Type Locality: Left bank of the Rio Grande, about 6 mi (10 km) 

above El Paso and opposite the initial monument of the 

Mexican boundary, [El Paso County], Texas.

Sigmodon hispidus plenus Goldman

Proc. Biol. Soc. Wash. 41: 205, 18 Dec 1928.

= Sigmodon arizonae plenus Goldman, 1928. See Hall (1981: 

742); subspecies recognized by Davis and Young (1999:592).

Holotype: USNM 181086. [Skin and skull]. Adult male. Collected 

2 Feb 1913 by E. A. Goldman. Original number 21819.

Type Locality: Parker, altitude 350 ft (107 m), [La Paz County], 

Arizona.

Sigmodon hispidus saturatus V. Bailey

Proc. Biol. Soc. Wash. 15: 111, 2 Jun 1902b.

= Sigmodon toltecus (Saussure, 1860). See Musser and Carleton 

(2005:1177).

Holotype: USNM 99998. [Skin and skull]. Adult male. Collected 

5 Apr 1900 by E. W. Nelson and E. A. Goldman. Original 

number 14108.

Type Locality: Teapa, Tabasco, [SL: Mexico].

Sigmodon hispidus tonalensis V. Bailey

Proc. Biol. Soc. Wash. 15: 109, 2 Jun 1902b.

= Sigmodon mascotensis J. A. Allen, 1897. See Carleton et al. 

(1999:843), who allocated tonalensis to S. mascotensis and 

found no basis for subspecific recognition.

Holotype: USNM 75144. [Skin and skull]. Adult male. Collected 

9 Aug 1895 by E. W. Nelson and E. A. Goldman. Original 

number 8312.

Type Locality: Tonala, [FC: altitude 500 ft (152 m)], Chiapas, 

Mexico.

Sigmodon hispidus virginianus M. C. Gardner

Proc. Biol. Soc. Wash. 59: 137, 25 Oct 1946.

Subspecies recognized by Hall (1981:740) and Cameron (1999c: 

595).

Holotype: USNM 273535. Skin and skull. Adult female. Collected 30 Jan 1943 by J. B. Lewis. Original number 705. 

31384-X.

Type Locality: Triplet, altitude 160 ft (49 m), [SL: Seward Forest], Brunswick County, Virginia.

Sigmodon leucotis V. Bailey

Proc. Biol. Soc. Wash. 15: 115, 2 Jun 1902b.

= Sigmodon leucotis leucotis V. Bailey, 1902. See Baker (1969: 

220); subspecies recognized by Hall (1981:744).

Holotype: USNM 92001. [Skin and skull]. Adult female. Collected 2 Dec 1897 by E. W. Nelson and E. A. Goldman. 

Original number 11812.

Type Locality: Valparaiso Mountains, altitude 8,700 ft (2,652 m), 

Zacatecas, Mexico.

Sigmodon melanotis V. Bailey

Proc. Biol. Soc. Wash. 15: 114, 2 Jun 1902b.

= Sigmodon fulviventer melanotis V. Bailey, 1902. See Baker 

(1969:212); subspecies recognized by Hall (1981:742) and 

Shump (1999:593).

Holotype: USNM 50190. [Skin and skull]. Adult female. 

Collected 15 Jul 1892 by E. W. Nelson. Original number 

2834.

Type Locality: Patzcuaro, altitude 7,000 ft (2,134 m), Michoacan, Mexico.

Sigmodon minima Mearns

Proc. U.S. Natl. Mus. 17: 130, 19 Jul 1894.

= Sigmodon fulviventer minimus Mearns, 1894. See Baker and 

Greer (1962:123); subspecies recognized by Hall (1981:742) 

and Shump (1999:593).

Holotype: USNM 21187/37291. Skin [and skull]. Adult male. 

Collected 26 Apr 1892 by E. A. Mearns and F. X. Holzner. 

Original number [SL: 1704].

Type Locality: Upper Corner Monument [SL and FC: Number 

40], [Miller and Kellogg (1955:530): 1,500 m], [Hidalgo 

County], New Mexico, on the Mexican boundary line 100 

mi (161 km) west of the initial monument on the west bank 

of the Rio Grande.

Remarks: Published as skin number USNM 21187 to which 

skull number USNM 37291 belongs.

Sigmodon minimus goldmani V. Bailey

Proc. Biol. Soc. Wash. 26: 132, 21 May 1913.

= Sigmodon fulviventer goldmani V. Bailey, 1913. See Baker and 

Greer (1962:123); subspecies recognized by Hall (1981:742) 

and Shump (1999:593).

Holotype: USNM 167625. [Skin and skull]. Young adult female. 

Collected 28 Oct 1909 by E. A. Goldman. Original number 

[FC: Nelson and Goldman] 20695.

Type Locality: 7 mi (11 km) north of [SL and FC: Las] Palomas [FC: Hot Springs], altitude 4,200 ft (1,280 m), [Sierra 

County], New Mexico.

Sigmodon minimus woodi M. C. Gardner

J. Mammal. 29: 65, 13 Feb 1948b.

= Sigmodon fulviventer minimus Mearns, 1894. See Hall (1981: 

743); subspecies recognized by Shump (1999:593).

Holotype: USNM 266327. Skin and skull. Adult male. Collected 1 Dec 1940 by D. Wood Jr. Original number 12. 

30005-X.
6 2 • smithsonian contributions to zoology

Type Locality: East side of the Rio Grande [SL: River], 51 mi (82 

km) south of Albuquerque, near Bernardo, altitude 5,000 ft 

(1,524 m), Socorro County, New Mexico.

Sigmodon ochrognathus V. Bailey

Proc. Biol. Soc. Wash. 15: 115, 2 Jun 1902b.

Holotype: USNM 110333. [Skin and skull]. Adult female. Collected 13 Jun 1901 by V. Bailey. Original number 7681.

Type Locality: Chisos Mountains, altitude 8,000 ft (2,438 m), 

[Brewster County], Texas.

Sigmodon ochrognathus madrensis Goldman and Gardner

J. Mammal. 28: 58, 15 Feb 1947.

= Sigmodon ochrognathus V. Bailey, 1902. See Findley and Jones 

(1960:469).

Holotype: USNM 96268. Skin and skull. Adult male. Collected 

22 Sep 1898 by E. W. Nelson and E. A. Goldman. Original 

number 13051.

Type Locality: Foothills of Sierra Madre, about 30 mi (42 km) 

northwest of Parral, altitude about 6,200 ft (1,890 m), 

southern Chihuahua, Mexico.

Sigmodon planifrons Nelson and Goldman

Proc. Biol. Soc. Wash. 46: 197, 26 Oct 1933.

Holotype: USNM 71918. Skin and skull. Adult female. Collected 28 Feb 1895 by [E. W.] Nelson and [E. A.] Goldman. 

Original number 7569.

Type Locality: Juquila, altitude 5,000 ft (1,524 m), southwestern 

Oaxaca, Mexico.

Genus Sigmodontomys J. A. Allen, 1897

Nectomys alfari efficax Goldman

Smithson. Misc. Collect. 60(22): 7, 28 Feb 1913.

= Sigmodontomys alfari efficax Goldman, 1913. See Musser and 

Carleton (1993:748); subspecies recognized by Hall (1981: 

624) under Oryzomys alfari.

Holotype: USNM 178627. [Skin and skull]. Adult male. Collected 12 Mar 1912 by E. A. Goldman. Original number 

21498.

Type Locality: Cana, altitude 1,800 ft (549 m), [Darien], eastern 

Panama.

Genus Synaptomys Baird, 1857

Mictomys innuitus True

Proc. U.S. Natl. Mus. 17: 3, 24 Apr 1894c. Preprint of Proc. U.S. 

Natl. Mus. 17: 243, 15 Nov 1894d.

= Synaptomys borealis innuitus True, 1894. See A. B. Howell 

(1927a:28); subspecies recognized by Hall (1981:833) and 

West (1999:656).

Holotype: USNM 14838/24729. In alcohol, [skull removed]. 

Nursing female. Collected spring of 1884 by L. M. Turner. 

Original number [MC: 506].

Type Locality: Fort Chimo (Kuujjuaq), Ungava [District], Labrador (Quebec), [Canada].

Remarks: Published as USNM 14838, a specimen in alcohol; the 

skull was removed and cataloged as USNM 24729 on 16 

Nov 1899.

Synaptomys (Mictomys) bullatus Preble

Proc. Biol. Soc. Wash. 15: 181, 6 Aug 1902b.

= Synaptomys borealis borealis (Richardson, 1828). See A. B. 

Howell (1927a:22); subspecies recognized by Hall (1981: 

833) and West (1999:656).

Holotype: USNM 110632. Skin and skull. Adult male. Collected 

17 Jul 1901 by E. A. Preble. Original number 4511.

Type Locality: Trout Rock, [FC: Northern Arm], near [SL and 

FC: 25 mi (40 km) south of] Fort Rae, Great Slave Lake, 

Mackenzie [District, Northwest Territories], Canada.

Synaptomys cooperi Baird

Mammals Rep. Explor. Surv. 8(1): 558, 1857.

= Synaptomys cooperi cooperi Baird, 1857. See A. B. Howell 

(1927a:12); subspecies recognized by Hall (1981:831) and 

Linzey (1999b:658).

Holotype: USNM 1367/3230. Skin and skull. Received from 

W. Cooper of Hoboken, [New Jersey]. [MC: Skin cataloged 

26 Feb 1856; skull 8 Jun 1857].

Type Locality: No locality assigned in original description. Type 

locality fixed by Bole and Moulthrop (1942:146) at Jackson, Carroll County, New Hampshire.

Remarks: Type designated by skin number USNM 1367 by Baird 

(1857:556) to which skull number USNM 3230 belongs.

Synaptomys cooperi jesseni Long

Mammalia 51: 324, 20 Oct 1987.

Holotype: USNM 565670. Skin, skull, and skeleton. Adult female. Collected 28 Jul 1985 by C. A. Long and J. E. Long. 

Original number not given. University of Wisconsin–Stevens 

Point, Museum of Natural History number 6250.

Type Locality: [SL: Jessen’s Place], Swenson Road, north side of 

Washington Island, Door County, Wisconsin.

Remarks: Published as University of Wisconsin–Stevens Point, 

Museum of Natural History number 6250; cataloged as 

USNM 565670 on 10 May 1989.

Synaptomys cooperi kentucki Barbour

J. Mammal. 37: 414, 11 Sep 1956.

Subspecies recognized by Hall (1981:831) and Linzey (1999b: 

658).

Holotype: USNM 301891. Skin and skull. Adult male. Collected 

23 Feb 1953 by D. M. Linzey. Original number not given. 

University of Kentucky number 1964.

Type Locality: Sadieville, Scott County, Kentucky.

Synaptomys (Mictomys) dalli Merriam

Proc. Biol. Soc. Wash. 10: 62, 19 Mar 1896.
number 642 • 6 3

= Synaptomys borealis dalli Merriam, 1896. See A. B. Howell 

(1927a:24); subspecies recognized by Hall (1981:833) and 

West (1999:656).

Holotype: USNM 49373. Skull and skeleton. Adult male. Collected Feb 1867 by W. H. Dall. Original number not given.

Type Locality: Nulato, [A. B. Howell (1927a:24): Yukon River], 

Alaska.

Remarks: Published as USNM 10597, a specimen that was originally in alcohol. Sometime prior to Merriam’s description, 

the specimen was made into a skull and skeleton and was 

recataloged as USNM 49373 on 28 Apr 1894. The skin 

was reported missing by Lyon and Osgood (1909:77) and 

was probably discarded.

Synaptomys helaletes Merriam

Proc. Biol. Soc. Wash.10: 59, 19 Mar 1896.

= Synaptomys cooperi helaletes Merriam, 1896. See A. B. Howell 

(1927a:17); subspecies recognized by Hall (1981:831) and 

Rose (2011:60).

Holotype: USNM 75172. [Skin and skull]. Adult female. Collected 14 Oct 1895 by A. K. Fisher. Original number 1818.

Type Locality: [SL and FC: Lake Drummond], Dismal Swamp, 

[Chesapeake/Suffolk City], Virginia.

Synaptomys helaletes gossii Merriam

Proc. Biol. Soc. Wash. 10: 60, 19 Mar 1896.

= Synaptomys cooperi gossii Merriam, 1896. See Rhoads and 

Young (1897:307); subspecies recognized by Hall (1981: 

831) and Linzey (1999b:658).

Holotype: USNM 6915. Skull only. Old male. Collected 1866 

[MC: cataloged 4 May 1866] by B. F. Goss. Original number [MC: 1].

Type Locality: Neosho Falls, [Woodson County], Kansas.

Remarks: The name gossii has been attributed to Coues by 

some authors, but Coues (1877:235) simply listed the 

name as Arvicola (Synaptomys) gossii in the synonymy of 

Synaptomys cooperi as a manuscript name of Baird and 

provided no description. Merriam (1896:60) considered 

Coues’s usage a nomen nudum. See ICZN (1999: Articles 

11.6 and 12.1).

Synaptomys (Mictomys) sphagnicola Preble

Proc. Biol. Soc. Wash. 13: 43, 29 May 1899b.

= Synaptomys borealis sphagnicola Preble, 1899. See A. B. Howell 

(1927a:30); subspecies recognized by Hall (1981:834) and 

West (1999:656).

Holotype: USNM 96543. [Skin and skull]. Adult male. Collected 

29 Jun 1898 by E. A. Preble. Original number 2402.

Type Locality: [FC: Mouth of Dartmouth Brook], Fabyans, 

near base of Mount Washington, [Coos County], New 

Hampshire.

Synaptomys (Mictomys) truei Merriam

Proc. Biol. Soc. Wash. 10: 62, 19 Mar 1896.

= Synaptomys borealis truei Merriam, 1896. See Hall (1981: 

834); subspecies recognized by Hall (1981:834) and West 

(1999:656).

Holotype: USNM 3798/12101. Skin and skull. Young adult, 

sex not given. Collected 6 Aug 1859 by C. B. R. Kennerly. 

Original number [SL: 304].

Type Locality: Skagit Valley, [Skagit County], Washington. In the 

original description, Merriam (1896:62) added “probably 

in the mountains bordering Skagit Valley.”

Synaptomys (Mictomys) wrangeli Merriam

Proc. Biol. Soc. Wash. 10: 63, 19 Mar 1896.

= Synaptomys borealis truei Merriam, 1896. See Hall (1981: 

834); subspecies recognized by Hall (1981:834) and West 

(1999:656).

Holotype: USNM 74720. [Skin and skull]. Adult male. Collected 

6 Sep 1895 by C. P. Streator. Original number 4871.

Type Locality: Wrangell, [FC: Wrangell Island, Sea level], 

Alaska.

Genus Thomasomys Coues, 1884

Thomasomys daphne Thomas

Smithson. Misc. Collect. 68(4): 2, 10 Apr 1917.

Holotype: USNM 194902. [Skin and skull]. Female. Collected 

[SL: 29 Jul] 1915 by E. Heller. Original number 521.

Type Locality: [SL and FC: Tocopoqueu], Ocobamba Valley, altitude 9,100 ft (2,774 m), [Cusco], Peru.

Thomasomys gracilis Thomas

Smithson. Misc. Collect. 68(4): 2, 10 Apr 1917.

Holotype: USNM 194816. [Skin and skull]. Male. Collected [SL: 

8 Jun] 1915 by E. Heller. Original number 321.

Type Locality: [SL and FC: Runcaraccay Ruins], Matchu 

(Machu) Picchu, altitude 12,000 ft (3,658 m) [SL and FC: 

14,000 ft (4,267 m)], [Cusco], Peru.

Thomasomys notatus Thomas

Smithson. Misc. Collect. 68(4): 2, 10 Apr 1917.

Holotype: USNM 194548. [Skin and skull]. Male. Collected [SL: 

5 May] 1915 by E. Heller. Original number 173.

Type Locality: Torontoy, altitude 9,500 ft (2,894 m), [Cusco], 

Peru.

Genus Transandinomys Weksler, Percequillo, 

and Voss, 2006

Oryzomys bombycinus Goldman

Smithson. Misc. Collect. 56(36): 6, 19 Feb 1912b.

= Transandinomys bolivaris (J. A. Allen, 1901). See Weksler et 

al. (2006:25).

Holotype: USNM 171105. [Skin and skull]. Adult male. Collected 26 Mar 1911 by E. A. Goldman. Original number 

[FC: Nelson and Goldman] 21009.
6 4 • smithsonian contributions to zoology

Type Locality: Cerro Azul, altitude 2,500 ft (762 m), near headwaters of the Chagres River, [Panama], Panama.

Oryzomys medius Robinson and Lyon

Proc. U.S. Natl. Mus. 24: 142, 3 Oct 1901.

= Transandinomys talamancae (J. A. Allen, 1891). See Weksler 

et al. (2006:25).

Holotype: USNM 105405. Skin and skull. Young adult male. 

Collected 8 Aug 1900 by [SL: W. Robinson]. Original number [SL: 1655].

Type Locality: San Julian, 8 mi (13 km) east of La Guaira, [Vargas], Venezuela.

Oryzomys talamancae J. A. Allen

Proc. U.S. Natl. Mus. 14: 193, 24 Jul 1891.

= Transandinomys talamancae (J. A. Allen, 1891). See Weksler 

et al. (2006:25).

Holotype: USNM 12222/22742. Skin and skull. Adult male. 

Collected, date unknown, by W. M. Gabb, [MC: cataloged 

19 Nov 1874]. Original number [SL: 99].

Type Locality: Talamanca, [Limon, Costa Rica].

Genus Tylomys Peters, 1866

Tylomys bullaris Merriam

Proc. Wash. Acad. Sci. 3: 561, 29 Nov 1901d.

Holotype: USNM 76058. [Skin and skull]. Immature male. Collected 7 Sep 1895 by E. W. Nelson and E. A. Goldman. 

Original number 8406.

Type Locality: Tuxtla, [FC: altitude 2,600 ft (798 m)], Chiapas, 

Mexico.

Tylomys tumbalensis Merriam

Proc. Wash. Acad. Sci. 3: 560, 29 Nov 1901d.

Holotype: USNM 76059. [Skin and skull]. Young adult male. 

Collected 23 Oct 1895 by E. W. Nelson and E. A. Goldman. 

Original number 8568.

Type Locality: Tumbala, [FC: altitude 5,000 ft (1,524 m)], 

Chiapas, Mexico.

Genus Xenomys Merriam, 1892

Xenomys nelsoni Merriam

Proc. Biol. Soc. Wash. 7: 161, 29 Sep 1892b.

Holotype: USNM 33281/45286. Skin and skull. Adult male. 

Collected 21 Mar 1892 by E. W. Nelson. Original number 

2288.

Type Locality: Hacienda Magdalena, [FC: altitude about 1,500 

ft (457 m)], Colima, Mexico.

Genus Zygodontomys J. A. Allen, 1897

Zygodontomys cherriei ventriosus Goldman

Smithson. Misc. Collect. 56(36): 8, 19 Feb 1912b.

= Zygodontomys brevicauda cherriei (J. A. Allen, 1895). See 

Voss (1991:59).

Holotype: USNM 171098. [Skin and skull]. Adult male. Collected 12 Mar 1911 by E. A. Goldman. Original number 

[FC: Nelson and Goldman] 20975.

Type Locality: Tabernilla, Canal Zone (Colon), Panama.

Family Muridae Illiger, 1811

Genus Acomys Geoffroy, 1838

Acomys hystrella Heller

Smithson. Misc. Collect. 56(17): 13, 28 Feb 1911.

= Acomys cineraceus Hueglin, 1877. See Musser and Carleton 

(2005:1195).

Holotype: USNM 164821. [Skin and skull]. Adult female. Collected 4 Feb 1910 by J. A. Loring. Original number 8929.

Type Locality: Nimule, [Al-lstiwa’lyah, Southern Region], Uganda 

(South Sudan).

Acomys ignites montanus Heller

Smithson. Misc. Collect. 63(7): 12, 24 Jun 1914.

= Acomys kempi Heller, 1914. See Musser and Carleton (2005: 

1197).

Holotype: USNM 182901. [Skin only]. Adult female. Collected 

26 Feb 1911 by A. B. Percival. Original number 309.

Type Locality: North slope of Mount Marsabit, altitude 4,600 ft 

(1,402 m), [Eastern Province], British East Africa (Kenya).

Genus Aethomys Thomas, 1915

Epimys kaiseri centralis Heller

Smithson. Misc. Collect. 63(7): 10, 24 Jun 1914.

= Aethomys hindei (Thomas, 1902). See Musser and Carleton 

(1993:568).

Holotype: USNM 165035. [Skin and skull]. Adult male. 

Collected 11 Jan 1910 by J. A. Loring. Original number 

8633.

Type Locality: Rhino Camp, Lado Enclave, [Arua], Uganda.

Epimys kaiseri turneri Heller

Smithson. Misc. Collect. 63(7): 8, 24 Jun 1914.

= Aethomys kaiseri (Noack, 1887). See Thomas (1926:177).

Holotype: USNM 183395. [Skin and skull]. Adult female. Collected [SL: 19 Jan 1912] by H. J. A. Turner. Original number 

[SL: E. Heller] 5121.

Type Locality: Kisumu, [Kisumu District, Nyanza], British East 

Africa (Kenya).

Genus Apodemus Kaup, 1829

Apodemus agrarius chejuensis Johnson and Jones

Proc. Biol. Soc. Wash. 68: 171, 31 Dec 1955.

= Apodemus agrarius (Pallas, 1771).
number 642 • 6 5

Holotype: USNM 299204. Skin and skull. Adult male. Collected 

7 Sep 1954 by G. W. Byers. Original number J. K. Jones Jr. 

1641.

Type Locality: 10 mi (16 km) northeast of Mosulp’o, 420 m, 

Cheju Do, Quelpart Island, [Jeju], [SL: South] Korea.

Apodemus agrarius pallescens Johnson and Jones

Proc. Biol. Soc. Wash. 68: 169, 31 Dec 1955.

= Apodemus agrarius (Pallas, 1771).

Holotype: USNM 299161. Skin and skull. Adult female. Collected 26 Sep 1954 by J. K. Jones Jr. Original number 1713.

Type Locality: 8 mi (13 km) southwest of Kunsan, 10 m, [SL: 

South] Korea.

Apodemus microtis Miller

Proc. Biol. Soc. Wash. 25: 60, 13 Apr 1912c.

= Apodemus uralensis (Pallas, 1811). See Musser and Carleton 

(1993:574).

Holotype: USNM 155471. Skin and skull. Young adult male. Collected 11 Nov 1909 by W. Ruckbeil. Original number 12.

Type Locality: Vicinity of Dzharkent (Zharkent), [Almaty], Russian Turkestan (Kazakhstan).

Apodemus nigritalus Hollister

Smithson. Misc. Collect. 60(24): 1, 13 Mar 1913c.

= Apodemus peninsulae (Thomas, 1907). See Jones (1956:346).

Holotype: USNM 175164. Skin and skull. Adult male. Collected 

6 Aug 1912 by N. Hollister. Original number 4438.

Type Locality: Tepucha [SL: Dapucha], 125 mi (201 km) southeast of Biisk and 50 mi (80 km) northwest of Ongudai, Altai 

Mountains, [Altayskiy], Siberia (Russia).

Apodemus peninsulae sowerbyi Jones

Univ. Kans. Publ., Mus. Nat. Hist. 9: 343, 15 Aug 1956.

= Apodemus peninsulae (Thomas, 1907).

Holotype: USNM 175523. Skin and skull. Adult female. Collected 

23 May 1912 by A. de C. Sowerby. Original number 456.

Type Locality: [SL: Mountains] 30 mi (48km) west of Kuei-huacheng, 7,000 ft (2,134 m), Northern Shansi (Nei Monggol), 

[SL: Northern] China.

Apodemus praetor Miller

Proc. Biol. Soc. Wash. 27: 89, 11 May 1914.

= Apodemus peninsulae (Thomas, 1907). See Jones (1956:346).

Holotype: USNM 197792. Skin and skull. Adult male. Collected 

6 Jul 1913 by A. de C. Sowerby. Original number 625.

Type Locality: Sungaree River, 60 mi (97 km) southwest of Kirin 

(Jilin), [SL: 500 ft (152 m)], Kirin (Jilin) Province, Manchuria (China).

Mus arianus griseus True

Proc. U.S. Natl. Mus. 17: 8, 8 May 1894b. Junior primary homonym of Mus sylvaticus var. griseus Palumbo, 1868. Renamed Apodemus flavicollis rusiges by Miller (1913c:81).

= Apodemus rusiges Miller, 1913. See Musser and Carleton 

(1993:573).

Syntypes: USNM 20139/35502. Skin and skull. [Lyon and 

Osgood (1909:149): Adult] female. Central Kashmir, altitude 8,500 ft (2,591 m), [Jammu and Kashmir], Kashmir 

(India). Collected 8 Oct 1891 by W. L. Abbott. No original 

number.

USNM 20144/35507. Skin and skull. [Lyon and Osgood 

(1909:149): Young adult] male. Pir Panjal Pass, 8,500 ft 

(2,591 m), [Jammu and Kashmir], Kashmir (India). Collected 31 Aug 1891 by W. L. Abbott. No original number.

USNM 20151/35514. Skin and skull. Immature male. Mountains of central Kashmir, altitude 10,000 ft (3,048 m). 

[Jammu and Kashmir], Kashmir (India). Collected 13 Sep 

1891 by W. L. Abbott. No original number.

Type Locality: The three localities listed above comprise the type 

locality (ICZN, 1999: Article 73.2.3).

Remarks: Holotype not designated in original description, which 

was based on these three specimens that True (1894:8) listed 

by number.

Genus Apomys Mearns, 1905

Apomys bardus Miller

Proc. U.S. Natl. Mus. 38: 402, 19 Aug 1910b.

= Apomys insignis Mearns, 1905. See Musser (1982:19).

Holotype: USNM 144582. Skin and skull. Adult male. Collected 

28 May 1906 by E. A. Mearns. Original number 6166.

Type Locality: Summit of Mount Bliss, altitude 5,750 ft 

(1,753 m), Mindanao, [Philippines].

Apomys hylocœtes Mearns

Proc. U.S. Natl. Mus. 28: 456, 13 May 1905.

= Apomys hylocoetes Mearns, 1905. See Poole and Schantz 

(1942:441).

Holotype: USNM 125426. Skin and skull. Adult female. Collected 2 Jul 1904 by E. A. Mearns. Original number 5696.

Type Locality: Mount Apo, altitude 6,000 ft (1,829 m), southern 

Mindanao, Philippine Islands (Philippines).

Remarks: Mearns (1905:456) spelled his new name with a 

ligature, Apomys hylocœtes, which constitutes an incorrect original spelling that must be amended (ICZN, 1999: 

Article 32.5), as Poole and Schantz (1942:441) did in reporting Apomys hylocoetes. Apomys hylocetes as used in 

Musser (1982:3 and throughout) and Musser and Carleton 

(2005:1282) is an incorrect subsequent spelling (ICZN, 

1999: Article 33.3).

Apomys insignis Mearns

Proc. U.S. Natl. Mus. 28: 459, 13 May 1905.

Holotype: USNM 125230. Skin and skull. Adult female. Collected 8 Jul 1904 by E. A. Mearns. Original number 5711.

Type Locality: Mount Apo, altitude 6,000 ft (1,829 m), southern 

Mindanao, Philippine Islands (Philippines).
6 6 • smithsonian contributions to zoology

Apomys major Miller

Proc. U.S. Natl. Mus. 38: 402, 19 Aug 1910b.

= Apomys datae (Meyer, 1899). See Musser (1982:9).

Holotype: USNM 151513. Skin and skull. Adult female. Collected 2 Aug 1907 by E. A. Mearns. Original number 6531.

Type Locality: Haights-in-the-Oaks, altitude 7,000 ft (2,134 m), 

Benguet [SL: Province], Luzon, [SL: Philippine Islands] 

(Philippines).

Apomys microdon Hollister

Proc. U.S. Natl. Mus. 46: 327, 31 Dec 1913d.

Holotype: USNM 155145. Skin and skull. Adult male. Collected 

8 May 1909 by D. B. Mackie. Original number 6.

Type Locality: Biga, Catanduanes Island, [Catanduanes Province], [SL: Philippine Islands] (Philippines).

Apomys musculus Miller

Proc. U.S. Natl. Mus. 38: 403, 19 Aug 1910b.

Holotype: USNM 145770. Skin and skull. Adult female. Collected 6 May 1907 by E. A. Mearns. Original number 6409.

Type Locality: Camp John Hay, Baguio, altitude 5,000 ft 

(1,524 m), Benguet [FC: Province], Luzon, [SL: Philippine 

Islands] (Philippines).

Apomys petraeus Mearns

Proc. U.S. Natl. Mus. 28: 458, 13 May 1905.

= Apomys hylocoetes Mearns, 1905. See Musser (1982:20).

Holotype: USNM 125245. Skin and skull. Adult female. Collected 30 Jun 1904 by E. A. Mearns. Original number 

5690.

Type Locality: Mount Apo, altitude 7,600 ft (2,316 m), southern 

Mindanao, Philippine Islands (Philippines).

Apomys sacobianus Johnson

Proc. Biol. Soc. Wash. 75: 318, 31 Dec 1962.

Holotype: USNM 304352. Skin and skull. Adult male. Collected 

16 Aug 1956 by D. H. Johnson. Original number 8555.

Type Locality: [SL and FC: Water point on] Sacobia River, Clark 

Air Base, Pampanga Province, Luzon, Philippine Islands 

(Philippines).

Genus Arvicanthis Lesson, 1842

Arvicanthis abyssinicus centrosus Hollister

Smithson. Misc. Collect. 66(10): 1, 26 Oct 1916.

= Arvicanthis niloticus (E. Geoffroy, 1803). See Misonne (1974:18).

Holotype: USNM 165167. Skin and skull. Adult male. Collected 

21 Jan 1910 by J. A. Loring. Original number 8817.

Type Locality: Rhino Camp, Lado [Enclave, Arua, Uganda].

Arvicanthis abyssinicus virescens Heller

Smithson. Misc. Collect. 63(7): 11, 24 Jun 1914.

= Arvicanthis nairobae J. A. Allen, 1909. See Musser and Carleton (1993:577).

Holotype: USNM 183922. [Skin and skull]. Adult male. Collected 15 Nov 1911 by E. Heller. Original number 4775.

Type Locality: Voi, [SL: Sagalla, base], [Coast Province], British 

East Africa (Kenya).

Arvicanthis jebelae Heller

Smithson. Misc. Collect. 56(17): 9, 28 Feb 1911.

= Arvicanthis niloticus (E. Geoffroy, 1803). See Misonne (1974:18).

Holotype: USNM 164826. [Skin and skull. Male]. Collected 16 

Jan 1910 by J. A. Loring. Original number 8762.

Type Locality: Rhino Camp, Lado Enclave, [Arua, Uganda].

Genus Bandicota Gray, 1873

Bandicota siamensis Kloss

J. Nat. Hist. Soc. Siam 3: 382, 31 Dec 1919.

= Bandicota indica (Bechstein, 1800). See Ellerman (1949:94).

Holotype: USNM 221559. Skin and skull. Adult male. Collected 

23 Oct [SL: 1916, not 1216 as in original description] by 

C. B. Kloss. Original number 2218.

Type Locality: [SL: Pak Bu], Tachin (Samut Sakhon), [Samut 

Sakhon], Central Siam (Thailand).

Remarks: Kloss used his field number 2218 in the original description; cataloged as USNM 221559 on 2 Jul 1920.

Genus Batomys Thomas, 1895

Batomys dentatus Miller

Proc. U.S. Natl. Mus. 38: 400, 19 Aug 1910b.

Holotype: USNM 151506. Skin and skull. Adult male. Collected 

31 Jul 1907 by E. A. Mearns. Original number 6484.

Type Locality: Haights-in-the-Oaks, altitude 7,000 ft (2,134 m), 

Benguet [SL: Province], Luzon, [SL: Philippine Islands] 

(Philippines).

Genus Berylmys Ellerman, 1947

Mus ferrocanus Miller

Proc. Biol. Soc. Wash. 13: 140, 21 Apr 1900c.

= Berylmys bowersi (Anderson, 1897). See Musser and Newcomb (1983:357).

Holotype: USNM 86737. Skin and skull. Adult female. Collected 

15 Jan 1899 by [SL: W. L. Abbott]. No original number.

Type Locality: [SL: Khow Nok Ram (Khao Nok Ra)], altitude 

3,000 ft (914 m), Trong (Trang), Siam (Thailand).

Genus Brachiones Thomas, 1925

Gerbillus arenicolor Miller

Proc. Biol. Soc. Wash. 13: 163, 31 Oct 1900d.

= Brachiones przewalskii (Buchner, 1899). See Corbet (1978:128).

Holotype: USNM 62153, not 62143 as in original description. 

Skin and skull. Adult male. Collected 9 Feb 1894 by W. L. 

Abbott. No original number.
number 642 • 6 7

Type Locality: In jungle on Yarkand River, east of Maralbashi, 

[Xinjiang], Eastern Turkestan (China).

Genus Bullimus Mearns, 1905

Bullimus bagobus Mearns

Proc. U.S. Natl. Mus. 28: 450, 13 May 1905.

Holotype: USNM 125248. Skin and skull. Adult female. Collected 13 Jul 1904 by E. A. Mearns. Original number 5719.

Type Locality: Todaya, a Bagobo village at 4,000 ft (1,219 m), 

Mount Apo, southern Mindanao, [Davao del Sur Province], 

Philippine Islands (Philippines).

Rattus bagobus barkeri Johnson

J. Wash. Acad. Sci. 36: 318, 17 Sep 1946.

= Bullimus bagobus Mearns, 1905. See Musser and Heaney 

(1992:115).

Holotype: USNM 278141. Skin and skull. Young adult male. 

Collected 7 Mar 1945 by J. E. Barker. Prepared by D. H. 

Johnson. Original number [SL: DHJ] 276.

Type Locality: Village of Mercedes, 5 mi (8 km) west of Guiuan, 

altitude about 200 ft (61 m), southeastern Samar Island, 

[Eastern Samar Province], Philippine Islands (Philippines).

Genus Bunomys Thomas, 1910

Mus andrewsi J. A. Allen

Bull. Am. Mus. Nat. Hist. 30: 336, 21 Dec 1911.

= Bunomys andrewsi (J. A. Allen, 1911). See Musser (1981a:117).

Holotype: USNM 175899. [Skin and skull]. Adult male. Collected 13 Dec 1909 by R. C. Andrews. Original number 

[SL: 25].

Type Locality: [SL: Labuan Balanda], Buton Island (Pulau Buton), 

[Sulawesi Tenggara], Celebes (Sulawesi), [Indonesia].

Rattus adspersus Miller and Hollister

Proc. Biol. Soc. Wash. 34: 71, 31 Mar 1921a.

= Bunomys andrewsi (J. A. Allen, 1911). See Musser (1981a:117).

Holotype: USNM 219602. Skin and skull. Adult male. Collected 

22 Jan 1918 by H. C. Raven. Original number 3427.

Type Locality: Pinedapa, [Sulawesi Tengah], Middle Celebes 

(Sulawesi), [Indonesia].

Rattus nigellus Miller and Hollister

Proc. Biol. Soc. Wash. 34: 72, 31 Mar 1921a.

= Bunomys chrysocomus (Hoffmann, 1887). See Musser (1981a: 

115).

Holotype: USNM 218140. Skin and skull. Adult male. Collected 

8 Nov 1916 by H. C. Raven. Original number 2936.

Type Locality: Bumbaroedjaba, near Toboli, [Sulawesi Tengah], 

Middle Celebes (Sulawesi), [Indonesia].

Rattus penitus Miller and Hollister

Proc. Biol. Soc. Wash. 34: 72, 31 Mar 1921a.

= Bunomys penitus (Miller and Hollister, 1921). See Musser 

(1981a:117).

Holotype: USNM 218686. Skin and skull. Adult male. Collected 

21 Jan 1917 by H. C. Raven. Original number 3109.

Type Locality: Goenoeng Lehio, southwest of Lake Lindoe 

(Rano Lindu), altitude 6,000 ft (1,829 m), [Sulawesi Tengah], Middle Celebes (Sulawesi), [Indonesia].

Rattus rallus Miller and Hollister

Proc. Biol. Soc. Wash. 34: 73, 31 Mar 1921a.

= Bunomys chrysocomus (Hoffmann, 1887). See Musser (1981a: 

117).

Holotype: USNM 219595. Skin and skull. Adult female. Collected 7 Sep 1917 by H. C. Raven. Original number 3233.

Type Locality: Gimpoe (Gimpu), [Sulawesi Tengah], Middle Celebes (Sulawesi), [Indonesia].

Rattus sericatus Miller and Hollister

Proc. Biol. Soc. Wash. 34: 73, 31 Mar 1921a.

= Bunomys penitus (Miller and Hollister, 1921). See Musser 

(1981a:117).

Holotype: USNM 219627. Skin and skull. Adult male. Collected 

19 Dec 1917 by H. C. Raven. Original number 3340.

Type Locality: Rano Rano, east of Lake Lindoe (Rano Lindu) 

and north of Lake Poso (Danau Poso), altitude 6,000 ft 

(1,829 m), [Sulawesi Tengah], Middle Celebes (Sulawesi), 

[Indonesia].

Genus Chiropodomys Peters, 1869

Chiropodomys muroides Medway

J. Malay. Branch R. Asiat. Soc. 36: 133, Dec 1965.

Holotype: USNM 301101. Skin and skull. Adult female. Collected 10 Aug 1953 by D. H. Johnson, [SL: says R. Traub. 

Original number B-20622].

Type Locality: [SL: Tenompak], Bunda Tuhan, Kinabalu. [SL: 

North Borneo (Sabah)], [Borneo, Malaysia].

Chiropodomys niadis Miller

Smithson. Misc. Collect. 45: 40, 6 Nov 1903b.

= Chiropodomys gliroides (Blyth, 1856). See Musser (1979:408).

Holotype: USNM 121867. Skin and skull. Adult female. Collected 30 Mar 1903 by W. L. Abbott. Original number 

2413.

Type Locality: Lafau, Nias Island (Pulau Nias), [Sumatera 

Utara], Sumatra, [Indonesia].

Genus Chrotomys Thomas, 1895

Chrotomys gonzalesi Rickart and Heaney

Proc. Biol. Soc. Wash. 104: 389, 18 Jun 1991.

Holotype: USNM 458952. Skull with carcass [including skin] in 

fluid. Adult male. Collected 28 Apr 1988 by L. R. Heaney. 

Original number 4176.
6 8 • smithsonian contributions to zoology

Type Locality: Western slope of Mount Isarog, 4 km N, 21 km 

E Naga, elevation 1,350 m, Camarines Sur Province, Luzon 

Island, Philippines, 13°40′N, 123°22′E.

Chrotomys whiteheadi mindorensis Kellogg

Proc. Biol. Soc. Wash. 58: 123, 20 Sep 1945a.

= Chrotomys mindorensis Kellogg, 1945. See Musser et al. 

(1982:514).

Holotype: USNM 277639. Skin and skull. Adult male. Collected 

7 May 1945 by R. M. Roecker. Original number 54.

Type Locality: 3 mi (5 km) south-southeast of San Jose (Central), 

altitude 200 ft (61 m), Mindoro Island, Philippine Islands 

(Philippines).

Genus Crateromys Thomas, 1895

Crateromys paulus Musser and Gordon

J. Mammal. 62: 515, 20 Aug 1981.

Holotype: USNM 522023. Skin and skull. Adult male. Collected 

4 Apr 1953 by P. Soriano and presented by E. Gelena. No 

original number.

Type Locality: Ilin Island, Mindoro Occidental Province, [Mindoro], Philippines.

Genus Dasymys Peters, 1875

Dasymys helukus Heller

Smithson. Misc. Collect. 54(4): 2, 28 Feb 1910b.

= Dasymys incomtus (Sundevall, 1847). See Hayman and Holt 

(1941:122).

Holotype: USNM 162889. [Skin and skull]. Adult male. Collected 19 Nov 1909 by E. Heller. Original number 1239.

Type Locality: Sirgoit [FC: Lake], Guas Ngishu plateau, British 

East Africa (Kenya).

Dasymys helukus nigridius Hollister

Smithson. Misc. Collect. 66(10): 2, 26 Oct 1916.

= Dasymys incomtus (Sundevall, 1847). See Hayman and Holt 

(1941:122).

Holotype: USNM 162465, Skin and skull. Adult female. 

Collected 20 Aug 1909 by J. A. Loring. Original number 

7054.

Type Locality: Naivasha Station, [Rift Valley], British East Africa 

(Kenya).

Remarks: For some unknown reason, this taxon was published 

as J. A. Loring number 7054 even though it was cataloged 

as USNM 162465 on 30 Dec 1909.

Dasymys orthos Heller

Smithson. Misc. Collect. 56(17): 13, 28 Feb 1911.

= Dasymys incomtus (Sundevall, 1847). See Misonne (1974:19).

Holotype: USNM 164824. [Skin and skull]. Adult male. Collected [SL and FC: 6], not 1 as in original description, Jan 

1910 by J. A. Loring. Original number 8601.

Type Locality: Butiaba, Albert Nyanza (Lake Albert), [Bulisa], 

Uganda.

Dasymys rufulus Miller

Proc. Wash. Acad. Sci. 2: 639, 28 Dec 1900e.

Holotype: USNM 83844. Skin and skull. Adult male. Collected 

30 Mar 1897 by [SL: R. P. Currie. Original number 19].

Type Locality: Mount Coffee, [Montserrado], Liberia, Africa.

Dasymys savannus Heller

Smithson. Misc. Collect. 56(17): 14, 28 Feb 1911.

= Dasymys incomtus (Sundevall, 1847). See Hayman and Holt 

(1941:122).

Holotype: USNM 164471. Skin and skull. Adult female. Collected 28 Oct 1909 by J. A. Loring. Original number 8182.

Type Locality: Fort Hall, [Central Province], British East Africa 

(Kenya).

Genus Dephomys Thomas, 1926

Mus defua Miller

Proc. Wash. Acad. Sci. 2: 635, 28 Dec 1900e.

= Dephomys defua (Miller, 1900). See Thomas (1926:177).

Holotype: USNM 83837. Skin and skull. Adult male. Collected 

[SL and FC: 3], not 13 as in original description, May 1897 

by [SL: R. P. Currie]. Original number 53.

Type Locality: Mount Coffee, [Lyon and Osgood (1909:147): 

altitude about 400 or 500 ft (122–152 m)], [Montserrado], 

Liberia, Africa.

Genus Dipodillus Lataste, 1881

See Abiadh et al. (2010) and Ndiaye et al. (2012), who make 

a strong case for Dipodillus being recognized as a subgenus of 

Gerbillus.

Dipodillus maghrebi Schlitter and Setzer

Proc. Biol. Soc. Wash. 84: 387, 29 Feb 1972.

Holotype: USNM 472714. Skin and skull. Adult male. Collected 

8 May 1970 by R. E. Vaden. Original number 6019.

Type Locality: 15 km WSW Taounate, 34°29′N, [SL and FC: 

4°], not 40° as in original description, 48′W, Fes Province, 

Morocco.

Dipodillus zakariai Cockrum, Vaughan, and Vaughan

Mammalia 40: 320, 30 Oct 1976.

Holotype: USNM 498997. Skin and skull. Adult male. Collected 

21 [SL: May], not Mai as in original description, 1975 by T. 

and P. Vaughan. Original number 3270.

Type Locality: 0.5 mi (0.8 km) E Kellabine, Kerkennah Islands, approximately 42 km airline E Sfax, [Safaqis Province], Tunisia.

Gerbillus campestris brunnescens Ranck

Bull. U.S. Natl. Mus. 275: 133, 2 Oct 1968.
number 642 • 6 9

= Dipodillus campestris (Loche, 1867). See Pavlinov et al. (1990: 

182).

Holotype: USNM 302180. Skin and skull. Adult male. Collected 

9 Nov 1955 by H. W. Setzer. Original number 2724.

Type Locality: 5 km SE Derna (Darnah), [Ranck (1968:67): 

32°44′N, 22°40′E], Cyrenaica. Libya.

Genus Echiothrix Gray, 1867

Echiothrix brevicula Miller and Hollister

Proc. Biol. Soc. Wash. 34: 67, 31 Mar 1921a.

= Echiothrix centrosa Miller and Hollister, 1921. See Musser 

and Carleton (2005:1321).

Holotype: USNM 219744. Skin and skull. Adult male. Collected 

29 Jan 1919 by H. C. Raven. Original number 3467.

Type Locality: Pinedapa, about 5 mi (8 km) inland from Gulf of 

Tomini, near Mapane, [Sulawesi Tengah], Middle Celebes 

(Sulawesi), [Indonesia].

Echiothrix centrosa Miller and Hollister

Proc. Biol. Soc. Wash. 34: 67, 31 Mar 1921a.

Holotype: USNM 218706. Skin and skull. Adult male. Collected 

9 Jan 1917 by H. C. Raven. Original number 3077.

Type Locality: Winatoe, between Koelawi and Gimpoe (Gimpu), 

[Sulawesi Tengah], Middle Celebes (Sulawesi), [Indonesia].

Genus Eropeplus Miller and Hollister, 1921

Eropeplus canus Miller and Hollister

Proc. Biol. Soc. Wash. 34: 95, 30 Jun 1921b.

Holotype: USNM 218707. Skin and skull. Female. Collected 12 

Jan 1917 by H. C. Raven. Original number 3079.

Type Locality: Goenoeng Lehio, altitude 6,000 ft (1,829 m), 

southwest from Lake Lindoe (Rano Lindu), [Sulawesi Tengah], Middle Celebes (Sulawesi). [Indonesia].

Genus Gerbilliscus Thomas, 1897

Gerbillus (Gerbillurus) setzeri Schlitter

Bull. South. Calif. Acad. Sci. 72: 13, 7 Jun 1973.

= Gerbilliscus setzeri (Schlitter, 1973). See Granjon et al. (2012:15).

Holotype: USNM 342253. Skin and skull. Young adult female. 

Collected 22 Nov 1963 by A. C. Riser. Original number 359.

Type Locality: Gobabeb, [Schmidt et al. (2008:34): 23°33′S, 

15°02′E], 1 mi (1.6 km) E Namib Desert Research Station, [FC: along Kuiseb River, 68 km ESE of Walvis Bay], 

[Erongo], South West Africa (Namibia).

Tatera nigricauda percivali Heller

Smithson. Misc. Collect. 63(7): 8, 24 Jun 1914.

= Gerbilliscus nigricaudus (Peters, 1878). See Musser and Carleton (2005:1220).

Holotype: USNM 183945. [Skin and skull]. Adult female. Collected 

[SL: 9 Sep 1911] by A. B. Percival. Original number 792.

Type Locality: Lorian Swamp, altitude 700 ft (213 m) [FC: 750 ft 

(229 m)], [North Eastern Province], British East Africa (Kenya).

Tatera pothae Heller

Smithson. Misc. Collect. 56(9): 2, 22 Jul 1910a.

= Gerbilliscus robustus (Cretzchmar, 1826). See Musser and 

Carleton (2005:1221).

Holotype: USNM 161716. Skin and skull. Adult female. Collected 30 Apr 1909 by J. A. Loring. Original number 6013.

Type Locality: Potha, Kapiti Plains, [Eastern Province], British 

East Africa (Kenya).

Tatera pringlei Hubbard

Zool. Afr. 5: 237, 1970.

= Gerbilliscus inclusus (Thomas and Wroughton, 1908). See 

Musser and Carleton (2005:1219).

Holotype: USNM 381474. Skin and skull. Adult male. Collected 

17 Jul 1965 by [SL: C. A. Hubbard]. Original number 

T2710.

Type Locality: Ubweri Village, Muheza, in northeast Tanzania.

Remarks: Published as T2710; cataloged as USNM 381474 on 

30 Jan 1967.

Tatera varia Heller

Smithson. Misc. Collect. 56(9): 1, 22 Jul 1910a.

= Gerbilliscus boehmi (Noack, 1887). See Musser and Carleton 

(2005:1218).

Holotype: USNM 162249. [Skin and skull]. Adult male. Collected 15 Jun 1909 by J. A. Loring. Original number 6338.

Type Locality: Loletai Plains, Southern Guaso Nyiro River, Sotik 

District, [Rift Valley], British East Africa (Kenya).

Genus Gerbillus Desmarest, 1804

Gerbillus andersoni blanci Cockrum, Vaughan, and Vaughan

Mammalia 40: 470, 20 Mar 1977.

= Gerbillus andersoni de Winton, 1902.

Holotype: USNM 498998. Skin and skull. Adult male. Collected 

[SL: 4 Apr], not 8 Apr as in original description, 1975 by 

T. and P. Vaughan. Original number 3171.

Type Locality: 2 km NE Bordj Cedria, 23 km SE Tunis, about 

36°42′N, 10°25′E, [Tunis], [SL: Tunisia].

Gerbillus aureus nalutensis Ranck

Bull. U.S. Natl. Mus. 275: 90, 2 Oct 1968.

= Gerbillus latastei Thomas and Trouessart, 1903. See Cockrum 

(1977:78).

Holotype: USNM 321816. Skin and skull. Adult male. Collected 

12 Nov 1961 by H. W. Setzer. Original number 3113.

Type Locality: 40 km ENE Nalut, [Ranck (1968:68): 32°01′N, 

11°22′E], Tripolitania Province, Libya.

Gerbillus cheesmani aquilus Schlitter and Setzer

Proc. Biol. Soc. Wash. 86: 167, 31 May 1973.
7 0 • smithsonian contributions to zoology

= Gerbillus aquilus Schlitter and Setzer, 1973. See Lay and 

Nadler (1975:437).

Holotype: USNM 354623. Skin and skull. Adult male. Collected 

8 Dec 1964 by R. G. Tuck Jr. Original number 1222.

Type Locality: 60 km W Kerman, Kerman Province, Iran.

Gerbillus cheesmani subsolanus Schlitter and Setzer

Proc. Biol. Soc. Wash. 86: 168, 31 May 1973.

= Gerbillus aquilus Schlitter and Setzer, 1973. See Lay and 

Nadler (1975:437).

Holotype: USNM 369335. Skin and skull. Adult female. Collected 3 Feb 1965 by R. G. Tuck Jr. Original number 1283.

Type Locality: 56 km E Nok Kundi, Kalat Division, [Balochistan], Pakistan.

Gerbillus eatoni inflatus Ranck

Bull. U.S. Natl. Mus. 275: 97, 2 Oct 1968.

= Gerbillus andersoni de Winton, 1902. See Cockrum et al. 

(1976:469).

Holotype: USNM 325527. Skin and skull. Adult male. Collected 

1 Jun 1962 by G. L. Ranck. Original number 2203.

Type Locality: 10 km SW Fort Capuzzo (Musa’id), [Ranck 

(1968:67): 31°31′N, 24°59′E], Cyrenaica Province, Libya.

Gerbillus eatoni versicolor Ranck

Bull. U.S. Natl. Mus. 275: 98, 2 Oct 1968.

= Gerbillus andersoni de Winton, 1902. See Cockrum et al. 

(1976:469).

Holotype: USNM 325298. Skin and skull. Adult male. Collected 

15 Jun 1962 by G. L. Ranck and J. H. Shaw. Original number [FC: GLR] 2341.

Type Locality: 2 km N Coefia (Al Kuwayfiyah), [Ranck (1968: 

67): 32°14′N, 20°11′E], Cyrenaica Province, Libya.

Gerbillus gerbillus aeruginosus Ranck

Bull. U.S. Natl. Mus. 275: 103, 2 Oct 1968.

= Gerbillus gerbillus (Oliver, 1801).

Holotype: USNM 325146. Skin and skull. Adult male. Collected 

3 Apr 1962 by G. L. Ranck. Original number 1920.

Type Locality: El Giof (Al Jawf), Cufra (Al Kufrah) Oasis, [Ranck 

(1968:67): 24°11′N, 23°19′E], Cyrenaica Province, Libya.

Gerbillus gerbillus discolor Ranck

Bull. U.S. Natl. Mus. 275: 106, 2 Oct 1968.

= Gerbillus gerbillus (Oliver, 1801).

Holotype: USNM 322461. Skin and skull. Adult male. Collected 

18 Dec 1961 by G. L. Ranck. Original number 1116.

Type Locality: Ghat, [Schmidt et al. (2008:34): 24°58′N, 

10°11′E], Fezzan (Ghat) Province, Libya.

Gerbillus gerbillus psammophilous Ranck

Bull. U.S. Natl. Mus. 275: 112, 2 Oct 1968.

= Gerbillus gerbillus (Oliver, 1801).

Holotype: USNM 325078. Skin and skull. Adult male. Collected 

16 Mar 1962 by G. L. Ranck. Original number 1790.

Type Locality: Gialo (Jalu) Oasis, [Ranck (1968:67): 29°15′N, 

21°14′E], Cyrenaica Province, Libya.

Gerbillus pyramidum aureus Setzer

Proc. Biol. Soc. Wash. 69: 179, 31 Dec 1956a.

= Gerbillus latastei Thomas and Trouessart, 1903. See Cockrum 

(1977:78).

Holotype: USNM 302078. Skin and skull. Adult male. Collected 

22 Oct 1955 by H. W. Setzer. Original number 2624.

Type Locality: 12 km W Zliten, [Ranck (1968:68): 32°29′N, 

14°26′E], Tripolitania Province, Libya.

Gerbillus pyramidum favillus Setzer

Proc. Biol. Soc. Wash. 69: 180, 31 Dec 1956a.

= Gerbillus latastei Thomas and Trouessart, 1903. See Cockrum 

(1977:78).

Holotype: USNM 302088. Skin and skull. Adult male. Collected 

24 Oct 1955 by H. W. Setzer. Original number 2637.

Type Locality: 2 km E Sirte (Surt), [Ranck (1968:68): 31°12′N, 

16°38′E], Tripolitania Province, Libya.

Gerbillus pyramidum hamadensis Ranck

Bull. U.S. Natl. Mus. 275: 118, 2 Oct 1968.

= Gerbillus tarabuli Thomas, 1902. See Lay (1983:348).

Holotype: USNM 321827. Skin and skull. Adult male. Collected 

14 Nov 1961 by G. L. Ranck. Original number 886.

Type Locality: 5 km E Derg (Dirj), [Ranck (1968:68): 30°12′N, 

10°29′E], Tripolitania Province, Libya.

Gerbillus pyramidum tibesti Setzer and Ranck

Proc. Biol. Soc. Wash. 84: 55, 30 Jun 1971.

= Gerbillus tarabuli Thomas, 1902. See Granjon et al. (1999:305).

Holotype: USNM 319721. Skin and skull. Adult male. Collected 

10 Apr 1961 by H. W. Setzer. Original number 3024.

Type Locality: Zouar, [Schmidt et al. (2008:34): 20°27′N, 

16°32′E], Tibesti Mountains, [Tibesti], Republic of Chad.

Genus Grammomys Thomas, 1915

Thamnomys dolichurus littoralis Heller

Smithson. Misc. Collect. 59(16): 10, 5 Jul 1912.

= Grammomys dolichurus (Smuts, 1832). See Hutterer and Dieterlen (1984:8).

Holotype: USNM 181799. [Skin and skull]. Adult male. Collected 22 Dec 1911 by E. Heller. Original number 4949.

Type Locality: Mazeras, [Coast Province], British East Africa (Kenya).

Genus Hybomys Thomas, 1910

Arvicanthis planifrons Miller

Proc. Wash. Acad. Sci. 2: 641, 28 Dec 1900e.

= Hybomys planifrons (Miller, 1900). See Carleton and Robbins 

(1985:956).

Holotype: USNM 83814. Skin and skull. Adult male. Collected 

26 Apr 1897 by [SL: R. P. Currie. Original number 46].
number 642 • 7 1

Type Locality: Mount Coffee. [Miller (1900e:631): altitude 400 

to 500 ft (122 to 142 m)], [Montserrado], Liberia, Africa.

Genus Hylomyscus Thomas, 1926

Epimys alleni kaimosae Heller

Smithson. Misc. Collect. 59(16): 7, 5 Jul 1912.

= Hylomyscus kaimosae (Heller, 1912). See Fadda et al. (2001: 

585), who listed H. kaimosae as valid without explanation; 

specific status corroborated by Nicolas et al. (2006:789).

Holotype: USNM 181794. [Skin and skull]. Adult female. Collected 1 Feb 1912 by E. Heller. Original number 5527.

Type Locality: Kaimosi, Kakumega (Kakamega) forest, [Rift 

Valley/Western Province], British East Africa (Kenya).

Epimys endorobae Heller

Smithson. Misc. Collect. 56(9): 3, 22 Jul 1910a.

= Hylomyscus endorobae (Heller, 1910). See Carleton et al. 

(2006:311).

Holotype: USNM 162888. [Skin and skull]. Adult male. Collected 27 Nov 1909 by E. Heller. Original number 1261.

Type Locality: Western edge of the Mau Forest, 25 mi (40 km) 

[SL and FC: 30 mi (48 km)] north of Edoma (Eldama) Ravine, [SL and FC: Guasa Ngishu Plateau], altitude 8,600 ft 

(2,621 m), [Rift Valley, Kenya].

Genus Lemniscomys Trouessart, 1881

Arvicanthis pulchellus micropus Heller

Smithson. Misc. Collect. 56(17): 9, 28 Feb 1911.

= Lemniscomys striatus massaicus (Pagenstecher, 1885). See 

Hollister (1919:139); subspecies recognized by Castiglia 

and Oguge (2008:57).

Holotype: USNM 164825. [Skin and skull]. Adult female. 

Collected 24 Jan 1910 by J. A. Loring. Original number 

8861.

Type Locality: Rhino Camp, Lado [Enclave, Arua, Uganda].

Lemniscomys dorsalis mearnsi Heller

Smithson. Misc. Collect. 63(7): 12, 24 Jun 1914.

= Lemniscomys rosalia (Thomas, 1904). See Musser and Carleton (1993:602).

Holotype: USNM 163616. [Skin and skull]. Adult female. 

Collected 11 Sep 1909 by J. A. Loring. Original number 

7152.

Type Locality: Fort Hall, altitude 6,200 ft (1,890 m), [Central 

Province], British East Africa (Kenya).

Lemniscomys hoogstraali Dieterlen

Bonn. Zool. Beitr. 42: 11, 11 Apr 1991.

Holotype: USNM 342078. Skin and skull. Adult male. Collected 

23 Jan 1961 by H. Hoogstraal. Original number 13391.

Type Locality: Paloich, 12 mi (19 km) N Niayok, [SL: near Nile], 

approx. 10°26′N, 32°33′E, Upper Nile Province, Sudan 

(South Sudan).

Lemniscomys pulchellus spermophilus Heller

Smithson. Misc. Collect. 59(16): 11, 5 Jul 1912.

= Lemniscomys striatus massaicus (Pagenstecher, 1885). See 

Hollister (1919:139); subspecies recognized by Castiglia 

and Oguge (2008:57).

Holotype: USNM 181800. [Skin and skull]. Adult female. Collected 26 Aug 1911 by E. Heller. Original number 4103.

Type Locality: [SL: Summit], Mount Gargues (Uaragess), Altitude 7,000 ft (2,134 m), Mathews Range, [Rift Valley], British East Africa (Kenya).

Genus Lenomys Thomas, 1898

Lenomys longicaudus Miller and Hollister

Proc. Biol. Soc. Wash. 34: 95, 30 Jun 1921b.

= Lenomys meyeri (Jentink, 1879). See Laurie and Hill (1954:97).

Holotype: USNM 219712. Skin and skull. Adult female. Collected 1 Sep 1917 by H. C. Raven. Original number 3203.

Type Locality: Gimpoe (Gimpu), [Sulawesi Tengah], Middle Celebes (Sulawesi), [Indonesia].

Genus Lenothrix Miller, 1903

Lenothrix canus Miller

Proc. U.S. Natl. Mus. 26: 466, 3 Feb 1903c.

Holotype: USNM 114386. Skin and skull. Adult male. Collected 

27 Jan 1902 by W. L. Abbott. Original number [SL: 1477].

Type Locality: Pulo Tuanku (Pulau Tuangku), [SL: Banjak Islands (Kepulauan Banyak)], [Aceh, Sumatra, Indonesia].

Genus Leopoldamys Ellerman, 1947

Epimys luta Miller

Smithson. Misc. Collect. 61(21): 18, 29 Dec 1913d.

= Leopoldamys sabanus (Thomas, 1887). See Musser (1981b: 

257).

Holotype: USNM 151917. Skin and skull. Adult female. Collected 24 Dec 1907 by W. L. Abbott. Original number 5682.

Type Locality: [FC: Sungei Lima], Pulo (Pulau) Laut, [Kalimantan Selatan], Dutch southeast Borneo (Kalimantan), 

[Indonesia].

Epimys nasutus Lyon

Proc. U.S. Natl. Mus. 40: 104, 25 Apr 1911.

= Leopoldamys sabanus (Thomas, 1887). See Musser (1981b: 

256).

Holotype: USNM 145519. Skin and skull. Adult male. Collected 

21 May 1907 by W. L. Abbott. Original number 5270.

Type Locality: Pulo (Pulau) Panebangan, [Kalimantan Barat], off 

west coast of Borneo (Kalimantan), [Indonesia].

Epimys stentor Miller

Smithson. Misc. Collect. 61(21): 19, 29 Dec 1913d.

= Leopoldamys sabanus (Thomas, 1887). See Musser (1981b: 

257).
7 2 • smithsonian contributions to zoology

Holotype: USNM 124044. Skin and skull. Adult female. Collected 2 Jan 1904 by W. L. Abbott. Original number 2983.

Type Locality: James Island (Pulo Buda), Mergui Archipelago, 

[Tanintharyi, Burma].

Epimys vociferans clarae Miller

Smithson. Misc. Collect. 61(21): 20, 29 Dec 1913d.

= Leopoldamys sabanus (Thomas, 1887). See Musser (1981b: 

257).

Holotype: USNM 124115. Skin and skull. Adult male. Collected 

10 Jan 1904 by W. L. Abbott. Original number 3019.

Type Locality: Clara Island, Mergui Archipelago, [Tanintharyi, 

Burma].

Epimys vociferans insularum Miller

Smithson. Misc. Collect. 61(21): 19, 29 Dec 1913d.

= Leopoldamys sabanus (Thomas, 1887). See Musser (1981b: 

257).

Holotype: USNM 104167. Skin and skull. Adult female. Collected 27 Feb 1900 by W. L. Abbott. Original number 329.

Type Locality: Domel Island (Letsok-aw Kyun), Mergui Archipelago, [Tanintharyi, Burma].

Mus balae Miller

Smithson. Misc. Collect. 45: 33, 6 Nov 1903b.

= Leopoldamys sabanus (Thomas, 1887). See Musser (1981b: 

257).

Holotype: USNM 121781. Skin and skull. Adult female. Collected 12 Feb 1903 by W. L. Abbott. Original number 2274.

Type Locality: Tana Bala (Pulau Tanabala), Batu Islands (Kepulauan Batu), [Sumatera Utara], Sumatra, [Indonesia].

Mus fremens Miller

Proc. Acad. Nat. Sci. Phila. 54: 154, 11 Jun 1902a.

= Leopoldamys sabanus (Thomas, 1887). See Musser (1981b: 

257).

Holotype: USNM 113087. Skin and skull. Adult male. Collected 

4 Sep 1901 by [SL: W. L. Abbott]. Original number 1273.

Type Locality: [FC: Sakana Bay], Sinkep Island (Pulau Singkep), 

South China Sea [Poole and Schantz: (1942:463): RhioLinga (Lingga) Archipelago], [Kepulauan Riau, Sumatra, 

Indonesia].

Mus lucas Miller

Smithson. Misc. Collect. 45: 30, 6 Nov 1903b.

= Leopoldamys sabanus (Thomas, 1887). See Musser (1981b: 

257).

Holotype: USNM 104190. Skin and skull. Adult female. Collected 20 Jan 1900 by W. L. Abbott. Original number 253.

Type Locality: St. Luke (Za Det Nge) Island, Mergui Archipelago, [Tanintharyi, Burma].

Mus masae Miller

Smithson. Misc. Collect. 45: 32, 6 Nov 1903b.

= Leopoldamys sabanus (Thomas, 1887). See Musser (1981b: 

257).

Holotype: USNM 121822. Skin and skull. Adult female. Collected 21 Feb 1903 by W. L. Abbott. Original number 2327.

Type Locality: Tana masa (Pulau Tanahmasa), Batu Islands 

(Kepulauan Batu), [Sumatera Utara, Sumatra, Indonesia].

Mus matthaeus Miller

Smithson. Misc. Collect. 45: 29, 6 Nov 1903b.

= Leopoldamys sabanus (Thomas, 1887). See Musser (1981b: 

257).

Holotype: USNM 104159. Skin and skull. Adult male. Collected 

18 Jan 1900 by W. L. Abbott. Original number 243.

Type Locality: St. Matthew (Za Det Gyi) Island, Mergui Archipelago, [Tanintharyi, Burma].

Mus soccatus Miller

Smithson. Misc. Collect. 45: 30, 6 Nov 1903b.

= Leopoldamys siporanus (Thomas, 1895). See Musser (1981b: 

257).

Holotype: USNM 121549. Skin and skull. Adult male. Collected 

29 Dec 1902 by W. L. Abbott. Original number 2183.

Type Locality: North Pagi Island (Pulau Pagai Utara), [Mentawai 

Islands, Sumatera Barat], Sumatra, [Indonesia].

Mus strepitans Miller

Proc. Wash. Acad. Sci. 2: 207, 20 Aug 1900f.

= Leopoldamys sabanus (Thomas, 1887). See Musser (1981b: 

257).

Holotype: USNM 101697. Skin and skull. Adult female. Collected 10 Sep 1899 by [W. L. Abbott]. No original number.

Type Locality: Pulo (Pulau) Siantan, Anambas Islands (Kepulauan Anambas), [Kepulauan Riau, Indonesia].

Mus stridens Miller

Smithson. Misc. Collect. 45: 28, 6 Nov 1903b.

= Leopoldamys sabanus (Thomas, 1887). See Musser (1981b: 

257).

Holotype: USNM 104992. Skin and skull. Adult male. Collected 

10 Oct 1900 by W. L. Abbott. Original number 702.

Type Locality: Tioman Island (Pulau Tioman), off southeast 

coast of Malay Peninsula, [Pahang, Malaysia].

Mus stridulus Miller

Smithson. Misc. Collect. 45: 29, 6 Nov 1903b.

= Leopoldamys sabanus (Thomas, 1887). See Musser (1981b: 

257).

Holotype: USNM 104196. Skin and skull. Adult female. Collected 12 Mar 1900 by W. L. Abbott. Original number 350.

Type Locality: Bentinck Island, Mergui Archipelago, [Tanintharyi, Burma].

Mus vociferans Miller

Proc. Biol. Soc. Wash. 13: 138, 21 Apr 1900c.
number 642 • 7 3

= Leopoldamys sabanus (Thomas, 1887). See Musser (1981b: 

257).

Holotype: USNM 86736. Skin and skull. Adult male. Collected 

21 Feb 1899 by [W. L. Abbott]. No original number.

Type Locality: [SL: Khow Sai Dow (Khao Soi Dao)], altitude 

1,000 ft (305 m), in the mountains of Trong (Trang), Lower 

Siam (Thailand).

Mus vociferans lancavensis Miller

Proc. Biol. Soc. Wash. 13: 188, 21 Dec 1900g.

= Leopoldamys sabanus (Thomas, 1887). See Musser (1981b: 

257).

Holotype: USNM 104173. Skin and skull. Adult female. Collected 6 Dec 1899 by [W. L. Abbott]. Original number 122.

Type Locality: Pulo Lankawi (Pulau Langkawi), [Kedah, 

Malaysia].

Rattus fremens mansalaris Lyon

Proc. U.S. Natl. Mus. 52: 450, 30 Dec 1916a.

= Leopoldamys sabanus (Thomas, 1887). See Musser (1981b: 

257).

Holotype: USNM 114583. Skin and skull. Adult male. Collected 

4 Mar 1902 by W. L. Abbott. Original number 1591.

Type Locality: Pulo Mansalar (Pulau Musala), off west coast of 

Sumatra, [Sumatera Utara, Indonesia].

Rattus fremens tuancus Lyon

Proc. U.S. Natl. Mus. 52: 451, 30 Dec 1916a.

= Leopoldamys sabanus (Thomas, 1887). See Musser (1981b: 

257).

Holotype: USNM 114402. Skin and skull. Adult male. Collected 

26 Jan 1902 by W. L. Abbott. Original number 1470.

Type Locality: Pulo Tuanku (Pulau Tuangku), Banjak Islands 

(Kepulauan Banyak), [Aceh, Sumatra, Indonesia].

Rattus vociferans tapanulius Lyon

Proc. Biol. Soc. Wash. 29: 209, 22 Sep 1916b.

= Leopoldamys sabanus (Thomas, 1887). See Musser (1981b: 

257).

Holotype: USNM 114453. Skin and skull. Adult male. Collected 

16 Feb 1902 by W. L. Abbott. Original number 1538.

Type Locality: Tapanuli Bay (Teluk Tapanuli), west coast of Sumatra, [Sumatera Utara, Indonesia].

Genus Limnomys Mearns, 1905

Limnomys mearnsi Hollister

Proc. U.S. Natl. Mus. 46: 324, 31 Dec 1913d.

= Limnomys sibuanus Mearns, 1905. See Musser (1977a:11).

Holotype: USNM 144622. Skin and skull. Adult female. Collected 7 Jun 1905 by E. A. Mearns. Original number 6190.

Type Locality: Summit of Grand Malindang Peak (Mount Malingdang), altitude 9,000 ft (2,743 m), [Misamis Occidental 

Province], Mindanao, [Philippines].

Limnomys sibuanus Mearns

Proc. U.S. Natl. Mus. 28: 452, 13 May 1905.

Holotype: USNM 125228. Skin and skull. Adult female. Collected 30 Jun 1904 by E. A. Mearns. Original number 5688.

Type Locality: Mount Apo, altitude 6,600 ft (2,012 m), [Musser 

and Carleton (2005:1352): Davao City Province], southern 

Mindanao, Philippine Islands (Philippines).

Genus Lophuromys Peters, 1874

Lophuromys aquilus margarettae Heller

Smithson. Misc. Collect. 59(16): 7, 5 Jul 1912.

= Lophuromys aquilus (True, 1892).

Holotype: USNM 181793. Skin and skull. Adult male. Collected 

27 Aug 1911 by E. Heller. Original number 4126.

Type Locality: [SL: North Creek]. Mount Garguess (Uaragess), 

altitude 6,000 ft (1,829 m), Mathews Range, [Rift Valley], 

British East Africa (Kenya).

Lophuromys nudicaudus Heller

Smithson. Misc. Collect. 56(17): 11, 28 Feb 1911.

Holotype: USNM 125436. [Skin and skull]. Adult male. Collected 4 Aug 1903 by G. L. Bates. Original number 26.

Type Locality: Efulen (Efoulan), Bula Country, [Sud], Kamerun 

(Cameroon).

Lophuromys pyrrhus Heller

Smithson. Misc. Collect. 56(17): 10, 28 Feb 1911.

= Lophuromys ansorgei (de Winton, 1896). See Hollister (1919: 

112).

Holotype: USNM 164823. [Skin and skull]. Adult male. Collected 24 Jan 1910 by J. A. Loring. Original number 8853.

Type Locality: Rhino Camp, Lado Enclave, [Arua, Uganda].

Mus aquilus True

Proc. U.S. Natl. Mus. 15: 460, 26 Oct 1892.

= Lophuromys aquilus (True, 1892). See Thomas (1897:795).

Holotype: USNM 18997/34723. Skin and skull. Adult male. 

Collected 11 Apr 1888 by [MC: W. L. Abbott]. No original 

number.

Type Locality: Mount Kilima-Njaro (Kilimanjaro), altitude 

8,000 ft (2,438 m), [Kilimanjaro, Tanzania].

Genus Mastomys Thomas, 1915

Epimys concha [sic] ismailiae Heller

Smithson. Misc. Collect. 63(7): 9, 24 Jun 1914.

= Mastomys natalensis (Smith, 1834). See Musser and Carleton 

(2005:1365).

Holotype: USNM 165108. [Skin and skull]. Adult male. 

Collected 23 Feb 1910 by J. A. Loring. Original number 

9056.

Type Locality: Gondokoro, [Central Equatoria], Uganda (South 

Sudan).
7 4 • smithsonian contributions to zoology

Epimys coucha durumae Heller

Smithson. Misc. Collect. 59(16): 9, 5 Jul 1912.

= Mastomys natalensis (Smith, 1834). See Musser and Carleton 

(2005:1365).

Holotype: USNM 181796. [Skin and skull]. Adult male. Collected 23 Dec 1911 by E. Heller. Original number 5002.

Type Locality: Mazeras, [Coast Province], British East Africa 

(Kenya).

Epimys coucha neumani Heller

Smithson. Misc. Collect. 59(16): 8, 5 Jul 1912.

= Mastomys natalensis (Smith, 1834). See Musser and Carleton 

(2005:1365).

Holotype: USNM 181795. [Skin and skull]. [SL: Male]. Collected 26 Sep 1911 by E. Heller. Original number 4372.

Type Locality: Neuman’s Boma, Northern Guaso Nyiro (Ewaso 

Ng’iro) River, [Rift Valley], British East Africa (Kenya).

Epimys panya Heller

Smithson. Misc. Collect. 56(9): 2, 22 Jul 1910a.

= Mastomys natalensis (Smith, 1834). See Musser and Carleton 

(2005:1365).

Holotype: USNM 161886. [Skin and skull]. Adult male. Collected 23 May 1909 by J. A. Loring. Original Number 6220.

Type Locality: Juja Farm, Athi Plains, [Central Province], British 

East Africa (Kenya).

Mus tana True

Proc. U.S. Natl. Mus. 16: 602, 25 Oct 1893.

= Mastomys natalensis (Smith, 1834). See Musser and Carleton 

(2005:1365).

Holotype: USNM 21004/36055. Skin, skull, [and body in alcohol]. [Lyon and Osgood (1909:155): Adult] female. Collected Nov 1892 by W. A. Chanler and L. von Hohnel. No 

original number.

Type Locality: Along the Tana River, between the coast and 

Hameye, a point about 300 mi (483 km) from its mouth, 

[Coast Province], East Africa (Kenya).

Rattus coucha tinctus Hollister

Smithson. Misc. Collect. 68(10): 1, 16 Jan 1918.

= Mastomys natalensis (Smith, 1834). See Musser and Carleton 

(2005:1365).

Holotype: USNM 183294. Skin and skull. Adult male. Collected 

24 Jan 1912 by E. Heller. Original number 5188.

Type Locality: Kaimosi, Kavirondo, [Western Province], British 

East Africa (Kenya).

Genus Maxomys Sody, 1936

Epimys batus Miller

Proc. Biol. Soc. Wash. 24: 27, 24 Feb 1911c.

= Maxomys whiteheadi (Thomas, 1894). See Musser et al. 

(1979:593).

Holotype: USNM 121792. Skin and skull. Adult female. Collected 4 Mar 1903 by W. L. Abbott. Original number 2368.

Type Locality: Pulo Pinie (Pulau Pini), Batu Islands (Kepulauan 

Batu), [Sumatera Utara], Sumatra, [Indonesia].

Epimys perflavus Lyon

Proc. U.S. Natl. Mus. 40: 108, 25 Apr 1911.

= Maxomys surifer (Miller, 1900). See Musser et al. (1979:593).

Holotype: USNM 151918. Skin and skull. Adult male. Collected 

25 Dec 1907 by W. L. Abbott. Original number 5684.

Type Locality: [FC: Sungei Lima], Pulo (Pulau) Laut, [Kalimantan Selatan], off southeast coast of Borneo (Kalimantan), 

[Indonesia].

Epimys saturatus Lyon

Proc. U.S. Natl. Mus. 40: 109, 25 Apr 1911.

= Maxomys surifer (Miller, 1900). See Musser et al. (1979:593).

Holotype: USNM 145523. Skin and skull. Adult male. Collected 

17 May 1907 by W. L. Abbott. Original number 5236.

Type Locality: Pulo (Pulau) Panebangan, [Kalimantan Barat], off 

west coast of Borneo (Kalimantan), [Indonesia].

Epimys ubecus Lyon

Proc. U.S. Natl. Mus. 40: 109, 25 Apr 1911.

= Maxomys surifer (Miller, 1900). See Musser et al. (1979:593).

Holotype: USNM 151931. Skin and skull. Adult male. Collected 

3 Jan 1908 by W. L. Abbott. Original number 5727.

Type Locality: Pulo (Pulau) Sebuku, [Kalimantan Selatan], off 

southeast coast of Borneo (Kalimantan), [Indonesia].

Mus anambae Miller

Proc. Wash. Acad. Sci. 2: 205, 20 Aug 1900f.

= Maxomys surifer (Miller, 1900). See Musser et al. (1979:593).

Holotype: USNM 101737. Skin and skull. Adult female. Collected 21 Sep 1899 by [SL: W. L. Abbott]. No original 

number.

Type Locality: Pulo Jimaja (Pulau Jemaja), Anambas Islands 

(Kepulauan Anambas), [Kepulauan Riau, Indonesia].

Mus asper Miller

Proc. Biol. Soc. Wash. 13: 145, 21 Apr 1900c.

= Maxomys whiteheadi (Thomas, 1894). See Musser et al. (1979: 

593).

Holotype: USNM 86767. Skin and skull. Adult female. Collected 

2 Feb 1899 by [SL: W. L. Abbott]. No original number.

Type Locality: [SL: Khow Sai Dow (Khao Soi Dao)], altitude 

about 1,000 ft (305 m), Trong (Trang), Siam (Thailand).

Mus batamanus Lyon

Proc. U.S. Natl. Mus. 31: 654, 16 Jan 1907b.

= Maxomys whiteheadi (Thomas, 1894). See Musser et al. (1979: 

593).

Holotype: USNM 143232. Skin and skull. Adult male. Collected 

30 Mar 1906 by C. B. Kloss. Original number 75.
number 642 • 7 5

Type Locality: Senimba Bay (Teluk Senimba), Batam Island 

(Palau Batam), off southern end of Malay Peninsula, [Kepulauan Riau, Sumatra, Indonesia].

Mus bentincanus Miller

Smithson. Misc. Collect. 45: 38, 6 Nov 1903b.

= Maxomys surifer (Miller, 1900). See Musser et al. (1979:593).

Holotype: USNM 104269. Skin and skull. Adult female. Collected 11 Mar 1900 by W. L. Abbott. Original number 348.

Type Locality: Bentinck Island, Mergui Archipelago, [Tanintharyi, Burma].

Mus carimatae Miller

Proc. U.S. Natl. Mus. 31: 59, 23 Jul 1906a.

= Maxomys surifer (Miller, 1900). See Musser et al. (1979:593).

Holotype: USNM 125079. Skin and skull. Adult male. Collected 

20 Aug 1904 by W. L. Abbott. Original number 3612.

Type Locality: Telok Pai, Karimata Island (Pulau Karimata), [Kalimantan Barat, Borneo (Kalimantan), Indonesia].

Mus casensis Miller

Smithson. Misc. Collect. 45: 38, 6 Nov 1903b.

= Maxomys surifer (Miller, 1900). See Musser et al. (1979:593).

Holotype: USNM 104249. Skin and skull. Adult male. Collected 

28 Dec 1899 by W. L. Abbott. Original number 188.

Type Locality: Chance Island, Mergui Archipelago, [Tanintharyi, 

Burma].

Mus catellifer Miller

Proc. U.S. Natl. Mus. 26: 464, 3 Feb 1903c.

= Maxomys surifer (Miller, 1900). See Musser et al. (1979:593).

Holotype: USNM 114590. Skin and skull. Adult female. Collected 3 Mar 1902 by W. L. Abbott. Original number 1587.

Type Locality: Pulo Mansalar (Pulau Musala), off Tapanuli Bay 

(Teluk Tapanuli), [Sumatera Utara], Sumatra, [Indonesia].

Mus domelicus Miller

Smithson. Misc. Collect. 45: 39, 6 Nov 1903b.

= Maxomys surifer (Miller, 1900). See Musser et al. (1979:593).

Holotype: USNM 104257. Skin and skull. Adult female. Collected 24 Feb 1900 by W. L. Abbott. Original number 320.

Type Locality: Domel Island (Letsok-aw Kyun), Mergui Archipelago, [Tanintharyi, Burma].

Mus lingensis Miller

Proc. Wash. Acad. Sci. 2: 206, 20 Aug 1900f.

= Maxomys rajah (Thomas, 1894). See Musser et al. (1979:593).

Holotype: USNM 101614. Skin and skull. Adult male. Collected 

15 Jul 1899 by [SL: W. L. Abbott]. No original number.

Type Locality: [SL: Peak], Linga Island (Pulau Lingga), [SL: altitude 

1,000 ft (305 m)], [Kepulauan Riau, Sumatra, Indonesia].

Mus luteolus Miller

Smithson. Misc. Collect. 45: 36, 6 Nov 1903b.

= Maxomys surifer (Miller, 1900). See Musser et al. (1979:593).

Holotype: USNM 104276. Skin and skull. Adult female. Collected 15 Jan 1900 by W. L. Abbott. Original number 226.

Type Locality: St. Matthew (Za Det Gyi) Island, Mergui Archipelago, [Tanintharyi, Burma].

Mus mandus Lyon

Proc. U.S. Natl. Mus. 34: 644, 14 Sep 1908.

= Maxomys whiteheadi (Thomas, 1894). See Musser et al. (1979: 

593).

Holotype: USNM 144225. Skin and skull. Adult male. Collected 

23 Nov 1906 by W. L. Abbott. Original number 4898.

Type Locality: Sungei (Sungai) Mandau, tributary to the Siak 

River (Sungai Siak), eastern Sumatra, [Riau, Indonesia].

Mus pagensis Miller

Smithson. Misc. Collect. 45: 39, 6 Nov 1903b.

= Maxomys pagensis (Miller, 1903). See Musser et al. (1979:593).

Holotype: USNM 121629. Skin and skull. Adult male. Collected 

23 Dec 1902 by W. L. Abbott. Original number 2153.

Type Locality: [FC: Sikakep Straight], South Pagi Island (Pulau 

Pagai Selatan), [Sumatera Barat], Sumatra, [Indonesia].

Mus pellax Miller

Proc. Biol. Soc. Wash. 13: 147, 21 Apr 1900c.

= Maxomys rajah (Thomas, 1894). See Musser et al. (1979:593).

Holotype: USNM 86755. Skin and skull. Adult female. Collected 

5 Feb 1899 by [SL: W. L. Abbott]. No original number.

Type Locality: [SL: Khow Sai Dow (Khao Soi Dao)], altitude about 

1,000 ft (305 m), Trong (Trang), Lower Siam (Thailand).

Mus serutus Miller

Proc. U.S. Natl. Mus. 31: 59, 23 Jul 1906a.

= Maxomys surifer (Miller, 1900). See Musser et al. (1979:593).

Holotype: USNM 125032. Skin and skull. Adult male. Collected 

17 Aug 1904 by W. L. Abbott. Original number 3590.

Type Locality: Pulo (Pulau) Serutu, Karimata Islands (Kepulauan Karimata), [Kalimantan Barat, Borneo (Kalimantan), 

Indonesia].

Mus surifer Miller

Proc. Biol. Soc. Wash. 13: 148, 21 Apr 1900c.

= Maxomys surifer (Miller, 1900). See Musser et al. (1979:593).

Holotype: USNM 86746. Skin and skull. Adult male. Collected 

14 Jan 1899 by [SL: W. L. Abbott]. No original number.

Type Locality: [SL: Khow Nok Ram (Khao Nok Ra)], altitude 

about 3,000 ft (914 m), in the mountains of Trong (Trang), 

Lower Siam (Thailand).

Mus surifer butangensis Miller

Proc. Biol. Soc. Wash. 13: 190, 21 Dec 1900g.

= Maxomys surifer (Miller, 1900). See Musser et al. (1979:593).

Holotype: USNM 104309. Skin and skull. Adult male. Collected 

16 Dec 1899 by [SL: W. L. Abbott]. Original number 157.
7 6 • smithsonian contributions to zoology

Type Locality: Pulo (Ko) Adang, Butang Islands (Batong Group), 

[Satun, Thailand].

Mus surifer flavidulus Miller

Proc. Biol. Soc. Wash. 13: 189, 21 Dec 1900g.

= Maxomys surifer (Miller, 1900). See Musser et al. (1979:593).

Holotype: USNM 104330. Skin and skull. Adult female. Collected 4 Dec 1899 by [SL: W. L. Abbott]. Original number 

109.

Type Locality: Pulo Lankawi (Pulau Langkawi), [Kedah, Malaysia].

Mus umbridorsum Miller

Smithson. Misc. Collect. 45: 37, 6 Nov 1903b.

= Maxomys surifer (Miller, 1900). See Musser et al. (1979:593).

Holotype: USNM 104227. Skin and skull. Adult male. Collected 

24 Jan 1900 by W. L. Abbott. Original number 269.

Type Locality: Loughborough (Lord Loughborough) Island, 

Mergui Archipelago, [Tanintharyi, Burma].

Rattus hellwaldii cereus Miller and Hollister

Proc. Biol. Soc. Wash. 34: 74, 31 Mar 1921a.

= Maxomys hellwaldii (Jentink, 1878). See Musser et al. (1979: 

593).

Holotype: USNM 200232. Skin and skull. Adult male. Collected 

30 Nov 1914 by H. C. Raven. Original number 1846.

Type Locality: Toli Toli (Tolitoli), [Sulawesi Tengah], northwestern Celebes (Sulawesi), [Indonesia].

Rattus hellwaldii localis Miller and Hollister

Proc. Biol. Soc. Wash. 34: 74, 31 Mar 1921a.

= Maxomys hellwaldii (Jentink, 1878). See Musser et al. (1979: 

593).

Holotype: USNM 218120. Skin and skull. Adult male. Collected 

1 Dec 1916 by H. C. Raven. Original number 2987.

Type Locality: Laboea Sore (Labuasore), north of Parigi, [Sulawesi Tengah], Celebes (Sulawesi), [Indonesia].

Rattus lingensis antucus Lyon

Proc. U.S. Natl. Mus. 52: 449, 30 Dec 1916a.

= Maxomys surifer (Miller, 1900). See Musser et al. (1979:593).

Holotype: USNM 114390. Skin and skull. Adult female. Collected 28 Jan 1902 by W. L. Abbott. Original number 1484.

Type Locality: Pulo Tuanku (Pulau Tuangku), Banjak Islands 

(Kepulauan Banyak), [Aceh, Sumatra, Indonesia].

Rattus lingensis banacus Lyon

Proc. U.S. Natl. Mus. 52: 449, 30 Dec 1916a.

= Maxomys surifer (Miller, 1900). See Musser et al. (1979:593).

Holotype: USNM 114294. Skin and skull. Adult male. Collected 

19 Jan 1902 by W. L. Abbott. Original number 1434.

Type Locality: Pulo Bankaru (Pulau Bangkaru), Banjak Islands 

(Kepulauan Banyak), [Aceh, Sumatra, Indonesia].

Rattus lingensis mabalus Lyon

Proc. U.S. Natl. Mus. 52: 449, 30 Dec 1916a.

= Maxomys surifer (Miller, 1900). See Musser et al. (1979: 

593).

Holotype: USNM 121825. Skin and skull. Adult female. Collected 19 Feb 1903 by W. L. Abbott. Original number 2310.

Type Locality: Tana Masa (Pulau Tanamasa), Batu Islands (Kepulauan Batu), [Sumatera Utara, Sumatra, Indonesia].

Rattus lingensis pinacus Lyon

Proc. U.S. Natl. Mus. 52: 450, 30 Dec 1916a.

= Maxomys surifer (Miller, 1900). See Musser et al. (1979:593).

Holotype: USNM 121846. Skin and skull. Adult female. Collected 2 Mar 1903 by W. L. Abbott. Original number 2349.

Type Locality: Pulo Pinie (Pulau Pini), Batu Islands (Kepulauan 

Batu), [Sumatera Utara, Sumatra, Indonesia].

Rattus melanurus Shamel

J. Mammal. 21: 76, 15 Feb 1940.

= Maxomys whiteheadi (Thomas, 1894). See Musser et al. (1979: 

593).

Holotype: USNM 197371. Skin and skull. Adult male. Collected 

12 Sep 1913 by H. C. Raven. Original number [SL: 1025].

Type Locality: Pulo (Pulau) Miang Besar, just out from Sangkulirang Bay (Teluk Sangkulirang), [Kalimantan Timur], Borneo (Kalimantan), [Indonesia].

Rattus musschenbroekii tetricus Miller and Hollister

Proc. Biol. Soc. Wash. 34: 68, 31 Mar 1921a.

= Maxomys musschenbroekii Jentink, 1878. See Musser et al. 

(1979:593).

Holotype: USNM 219613. Skin and skull. Adult male. Collected 

27 Aug 1917 by H. C. Raven. Original number 3184.

Type Locality: Gimpoe (Gimpu), southwest from Lake Lindoe 

(Rano Lindu), [Sulawesi Tengah], Middle Celebes (Sulawesi), 

[Indonesia].

Rattus perasper Shamel

J. Mammal. 21: 76, 15 Feb 1940.

= Maxomys ochraceiventer (Thomas, 1894). See Musser et al. 

(1979:593).

Holotype: USNM 198255. Skin and skull. Adult male. Collected 

5 Dec 1913 by H. C. Raven. Original number [SL: 1210].

Type Locality: Gunong Batu, [SL: North bank, Sungai Karangan, Sangkulirang Bay (Teluk Sangkulirang)], [Kalimantan 

Timur], Borneo (Kalimantan), [Indonesia].

Rattus rajah koratis Kloss

J. Nat. Hist. Soc. Siam 3: 376, 31 Dec 1919.

= Maxomys surifer (Miller, 1900). See Musser et al. (1979: 

593).

Holotype: USNM 221509. Skin and skull. Adult male. Collected 

18 Oct 1916 by C. B. Kloss. Original number 2187.

Type Locality: Lat Bua Kao (Lat Bua Khao), [Nakhon Ratchasima], E. Siam (Thailand).

Remarks: Published as CBK number 2187; cataloged as USNM 

221509 on 1 Jul 1920.
number 642 • 7 7

Rattus rajah kramis Kloss

J. Nat. Hist. Soc. Siam 3: 377, 31 Dec 1919.

= Maxomys surifer (Miller, 1900). See Musser et al. (1979:593).

Holotype: USNM 221489. Skin and skull. Adult male. Collected 

30 Oct 1916 by C. B. Kloss. Original number 2277.

Type Locality: Koh Kram (Ko Khram), Inner Gulf of Siam, 

[Chon Buri, Thailand].

Remarks: Published as CBK number 2277; cataloged as USNM 

221489 on 1 Jul 1920.

Genus Melasmothrix Miller and Hollister, 1921

Melasmothrix naso Miller and Hollister

Proc. Biol. Soc. Wash. 34: 94, 30 Jun 1921b.

Holotype: USNM 219752. Skin and skull. Old male. Collected 2 

Jan 1918 by H. C. Raven. Original number 3368.

Type Locality: Rano Rano (Ranorano), [Sulawesi Tengah], Middle Celebes (Sulawesi), [Indonesia].

Genus Melomys Thomas, 1922

Melomys cervinipes albiventer Kellogg

Proc. Biol. Soc. Wash. 58: 69, 7 May 1945b.

= Melomys burtoni (Ramsay, 1887). See Mahoney and Richardson (1988:161).

Holotype: USNM 237782. Skin and skull. Adult male. Collected 

20 Oct 1920 by C. M. Hoy. Original number 861.

Type Locality: Either Brooks Creek or Douglas River, 100 

mi (161 km) south of Port Darwin, Northern Territory, 

Australia.

Genus Meriones Illiger, 1811

Meriones caudatus amplus Ranck

Bull. U.S. Natl. Mus. 275: 165, 2 Oct 1968.

= Meriones libycus Lichtenstein, 1823. See Lay and Nadler 

(1969:45).

Holotype: USNM 322681. Skin and skull. Adult male. Collected 

8 Jan 1962 by G. L. Ranck. Original number 1245.

Type Locality: El Gatrun (Al Qatrun), [Ranck (1968:68): 

24°57′N, 14°39′E], Fezzan Province, Libya.

Meriones caudatus luridus Ranck

Bull. U.S. Natl. Mus. 275: 173, 2 Oct 1968.

= Meriones libycus Lichtenstein, 1823. See Lay and Nadler 

(1969:45).

Holotype: USNM 325597. Skin and skull. Adult male. Collected 

29 May 1962 by G. L. Ranck. Original number 2187.

Type Locality: Bahr el (at) Tubat, 21 km ESE Giarabub (Al Jaghbub), [Ranck (1968:68): 29°36′N, 24°53′E], Cyrenaica 

Province, Libya.

Meriones erythrourus farsi Schlitter and Setzer

Proc. Biol. Soc. Wash. 86: 170, 31 May 1973.

= Meriones libycus Lichtenstein, 1823. See Corbet (1978:127).

Holotype: USNM 329186. Skin and skull. Adult male. Collected 

27 Mar 1963 by G. L. Ranck. Original number 3911.

Type Locality: 3 km N Bariz, 50 km N Lar, [SL: Laristan], Fars 

(Hormozgan) Province, Iran.

Meriones libycus auratus Ranck

Bull. U.S. Natl. Mus. 275: 183, 2 Oct 1968.

= Meriones shawi (Duvernoy, 1842). See Lay and Nadler 

(1969:45).

Holotype: USNM 302233. Skin and skull. Adult male. Collected 

29 Oct 1955 by H. W. Setzer. Original number 2672.

Type Locality: Gheminez (Qaminis), [Ranck (1968:67): 31°29′N, 

20°E], Cyrenaica Province, Libya.

Meriones shawi azizi Setzer

Proc. Biol. Soc. Wash. 69: 205, 31 Dec 1956b.

= Meriones shawi (Duvernoy, 1842).

Holotype: USNM 302242. Skin and skull. Adult male. Collected 

9 Nov 1955 by H. W. Setzer. Original number 2717.

Type Locality: 5 km SE Derna (Darnah), [Ranck (1968:67): 

32°44′N, 22°40′E], Cyrenaica, Libya.

Genus Micromys Dehne, 1841

Micromys minutus hertigi Johnson and Jones

Proc. Biol. Soc. Wash. 68: 167, 31 Dec 1955.

= Micromys minutus (Pallas, 1771).

Holotype: USNM 299104. Skin and skull. Adult female. Collected 11 Sep 1954 by W. D. Thomas. Original number J. K. 

Jones Jr. 1675.

Type Locality: 2 mi (3 km) southeast Mosulp’o, altitude 3 m, 

Quelpart Island, Cheju Do, Korea.

Genus Mus Linnaeus, 1758

Leggada naivashae Heller

Smithson. Misc. Collect. 54(4): 2, 28 Feb 1910b.

= Mus triton (Thomas, 1909). See Hollister (1919:95).

Holotype: USNM 162885. [Skin and skull]. Adult male. Collected 18 Oct 1909 by E. Heller. Original number 1186.

Type Locality: Eastern edge of Naivasha Plains at the [SL: western] base of the Aberdare Mountains, [SL and FC: altitude 

9,100 ft (2,774 m)], [Central Province], British East Africa 

(Kenya).

Mus bellus enclavae Heller

Smithson. Misc. Collect. 56(17): 8, 28 Feb 1911.

= Mus musculoides Temminck, 1853. See Musser and Carleton 

(1993:625).

Holotype: USNM 164818. [Skin and skull]. Adult male. Collected 10 Jan 1910 by J. A. Loring. Original number 8613.

Type Locality: Rhino Camp, Lado Enclave, [Arua], [SL: Uganda].

Mus bellus gondokorae Heller

Smithson. Misc. Collect. 56(17): 8, 28 Feb 1911.
7 8 • smithsonian contributions to zoology

= Mus musculoides Temminck, 1853. See Musser and Carleton 

(1993:625).

Holotype: USNM 164820. [Skin and skull]. Adult male. Collected 25 Feb 1910 by J. A. Loring. Original number 9089.

Type Locality: Gondokoro, [Central Equatoria], Uganda (South 

Sudan).

Mus bellus petilus Hollister

Smithson. Misc. Collect. 66(10): 3, 26 Oct 1916.

= Mus musculoides Temminck, 1853. See Musser and Carleton 

(1993:625).

Holotype: USNM 162397. Skin and skull. Adult male. Collected 

28 Jun 1909 by J. A. Loring. Original number 6450.

Type Locality: Southern Guaso Nyiro (Ewaso Ng’iro) River, British East Africa (Kenya).

Mus commissarius Mearns

Proc. U.S. Natl. Mus. 28: 449, 13 May 1905.

= Mus musculus castaneus Waterhouse, 1843. See Schwarz and 

Schwarz (1943:63).

Holotype: USNM 125213. Skin and skull. Adult female. Collected 19 Jul 1904 by E. A. Mearns. Original number 5734.

Type Locality: Military commissary building at Davao, [Davao 

City Province], southern Mindanao, Philippine Islands 

(Philippines).

Mus gratus soricoides Heller

Smithson. Misc. Collect. 63(7): 10, 24 Jun 1914.

= Mus musculoides Temminck, 1853. See Musser and Carleton 

(1993:625).

Holotype: USNM 183544. [Skin and skull]. Adult male. Collected 8 Nov 1911 by E. Heller. Original number 4675.

Type Locality: [SL: Summit] Mount Mbololo, Taita Hills, [Coast 

Province], British East Africa (Kenya).

Mus gratus sungarae Heller

Smithson. Misc. Collect. 56(17): 7, 28 Feb 1911.

= Mus musculoides Temminck, 1853. See Musser and Carleton 

(1993:625).

Holotype: USNM 163487. [Skin and skull]. Old male. Collected 

20 Sep 1909 by J. A. Loring. Original number 7425.

Type Locality: West Kenia Forest Station (Mount Kenya), altitude 7,500 ft (2,286 m), British East Africa (Kenya).

Mus musculoides emesi Heller

Smithson. Misc. Collect. 56(17): 5, 28 Feb 1911.

= Mus mahomet Rhoads, 1896. See Musser and Carleton (1993: 

624).

Holotype: USNM 164819. [Skin and skull]. Adult male. Collected 27 Dec 1909 by J. A. Loring. Original number 8497.

Type Locality: Kabula Muliro (Kabulamuliro), [Mityana], 

Uganda, Africa.

Mus peruvianus Peale

U.S. Explor. Exped. 8: 51, 1848.

= Mus musculus domesticus Schwarz and Schwarz, 1943. See 

Musser and Carleton (2005:1399).

Lectotype: USNM 4955, as herein designated. Alcoholic, skull 

removed. Unsexed adult. Date collected and collector not 

given; cataloged 30 May 1861.

Type Locality: Near Callao, [Callao], Peru.

Remarks: Type not designated by number in original description. Lyon and Osgood (1909:289) and Poole and Schantz 

(1942:574) presumed the type to be the above specimen but 

listed it as missing. This specimen was found in the collection on 14 Feb 1994. It bears a U.S. Exploring Expedition 

label and is the only known specimen of Mus among those 

mentioned by Peale (1848:51) to be extant. We designate 

USNM 4955 as lectotype in order to finally attach the name 

peruvianus to an identifiable specimen.

Mus setzeri Petter

Mammalia 42: 377, 31 Dec 1978.

Holotype: USNM 428742. [Skin and skull]. Male. Collected 21 

May 1967 by M. S. [SL and FC: S. W.] Goussard. Original 

number [SL: 633].

Type Locality: 82 km a l’ouest de Mohembo, Botswana, pres de 

la frontier de Namibie (Namibia).

Mus tenellus acholi Heller

Smithson. Misc. Collect. 56(17): 6, 28 Feb 1911.

= Mus sorella (Thomas, 1909). See Verheyen (1965:204).

Holotype: USNM 164817. [Skin and skull]. Adult male. Collected 13 Jan 1910 by J. A. Loring. Original number 8671.

Type Locality: Rhino Camp, Lado Enclave, [Arua, Uganda], Africa.

Mus wamae Heller

Smithson. Misc. Collect. 56(17): 5, 28 Feb 1911.

= Mus sorella (Thomas, 1909). See Verheyen (1965:204).

Holotype: USNM 161777. [Skin and skull]. Adult female. Collected 6 May 1909 by J. A. Loring. Original number 6061.

Type Locality: Kapiti Plains, [Eastern Province], British East Africa (Kenya).

Genus Mylomys Thomas, 1906

Pelomys roosevelti Heller

Smithson. Misc. Collect. 54(4): 1, 28 Feb 1910b.

= Mylomys dybowskii (Pousargues, 1893). See Misonne (1974:22).

Holotype: USNM 162881. [Skin and skull]. Adult male. Collected 16 Nov 1909 by E. Heller. Original number 1231.

Type Locality: Nzoia River, Guas Ngishu plateau, [Western 

Province], British East Africa (Kenya).

Genus Myomyscus Shortridge, 1942

Epimys niveiventris ulae Heller

Smithson. Misc. Collect. 56(9): 3, 22 Jul 1910c.

= Myomyscus brockmani (Thomas, 1908). See Musser and 

Carleton (2005:1414).
number 642 • 7 9

Holotype: USNM 162887. [Skin and skull]. Adult female. Collected 19 Nov 1909 by J. A. Loring. Original number 8221.

Type Locality: Ulu Kenia (Lukenya) Hills, [SL: Athi Plains], 

[Eastern Province], British East Africa (Kenya).

Genus Nesokia Gray, 1842

Nesokia indica chitralensis Schlitter and Setzer

Proc. Biol. Soc. Wash. 86: 171, 31 May 1973.

= Nesokia indica (Gray, 1830). 

Holotype: USNM 354362. Skin and skull. Adult male. Collected 

18 Aug 1964 by R. H. Pine. Original number 2496 [SL and 

FC: B67939A].

Type Locality: 4 mi (6 km) N Chitral, altitude approximately 

4,800 ft (1,463 m), Chitral State (District), [Malakand Division, North-West Frontier, Khyber Pakhtunkhwa], Pakistan.

Genus Niviventer Marshall, 1976

Epimys barussanus Miller

Proc. Biol. Soc. Wash. 24: 26, 24 Feb 1911c.

= Niviventer cremoriventer (Miller, 1900). See Musser (1981b:237).

Holotype: USNM 141208. Skin and skull. Adult male. Collected 

10 Mar 1905 by W. L. Abbott. Original number 4016.

Type Locality: Mouth of Mojeia River, Nias Island (Pulau Nias), 

[Sumatera Utara], Sumatra, [Indonesia].

Epimys gracilis Miller

Smithson. Misc. Collect. 61(21): 21, 29 Dec 1913d.

= Niviventer fulvescens (Gray, 1847). See Musser (1981b:237).

Holotype: USNM 101520. In alcohol, [skull removed]. Adult female. Collected on unknown date by L. Fea. Cataloged 15 

Dec 1899. No original number.

Type Locality: Summit of Mount Mooleyit (Mulayit Taung), 

northern Tenasserim (Kayin), [Burma].

Epimys lepidus Miller

Smithson. Misc. Collect. 61(21): 20, 29 Dec 1913d.

= Niviventer fulvescens (Gray, 1847). See Musser (1981b:237).

Holotype: USNM 104127. Skin and skull. Adult female. Collected 19 Feb 1900 by W. L. Abbott. Original number 315.

Type Locality: Bok Pyin (Bokpyin), southern Tenasserim (Tanintharyi), [Burma].

Epimys mengurus Miller

Proc. Biol. Soc. Wash. 24: 27, 24 Feb 1911c.

= Niviventer cremoriventer (Miller, 1900). See Musser (1981b:237).

Holotype: USNM 125021. Skin and skull. Adult male. Collected 

14 Aug 1904 by W. L. Abbott. Original number 3581.

Type Locality: Bukit Menguru, Billiton Island (Pulau Belitung), 

[Kepulauan Bangka Belitung], Sumatra, [Indonesia].

Epimys solus Miller

Smithson. Misc. Collect. 61(22): 21, 29 Dec 1913d.

= Niviventer cremoriventer (Miller, 1900). See Musser (1981b:237).

Holotype: USNM 123944. Skin and skull. Adult male. Collected 

16 Nov 1903 by W. L. Abbott. Original number 2902.

Type Locality: Pulo Terutau (Ko Tarutao), west coast of Malay 

Peninsula, [Satun, Thailand].

Epimys spatulatus Lyon

Proc. U.S. Natl. Mus. 40: 111, 25 Apr 1911.

= Niviventer cremoriventer (Miller, 1900). See Musser (1981b:237).

Holotype: USNM 145499. Skin and skull. Adult male. Collected 

10 May 1907 by W. L. Abbott. Original number 5214.

Type Locality: Pulo Lamukotan (Pulau Lemukutan), [Kalimantan 

Barat], off west coast of Borneo (Kalimantan), [Indonesia].

Mus cremoriventer Miller

Proc. Biol. Soc. Wash. 13: 144, 21 Apr 1900c.

= Niviventer cremoriventer (Miller, 1900). See Musser (1981b:237).

Holotype: USNM 86770. Skin and skull. Adult male. Collected 

16 Jan 1899 by [SL: W. L. Abbott]. No original number.

Type Locality: [SL: Khow Nok Ram (Khao Nak Ra)], altitude 

about 3,000 ft (914 m), mountains of Trong (Trang), Lower 

Siam (Thailand).

Mus flaviventer Miller

Proc. Wash. Acad. Sci. 2: 204, 20 Aug 1900f.

= Niviventer cremoriventer (Miller, 1900). See Musser (1981b:237).

Holotype: USNM 101739. Skin and skull. Adult male. Collected 

18 Sep 1899 by [MC: W. L. Abbott]. No original number.

Type Locality: Pulo Jimaja (Pulau Jemaja), Anambas Islands 

(Kepulauan Anambas), [Kepulauan Riau, Indonesia].

Mus gilbiventer Miller

Smithson. Misc. Collect. 45: 35, 6 Nov 1903c.

= Niviventer cremoriventer (Miller, 1900). See Musser (1981b:237).

Holotype: USNM 104153. Skin and skull. Adult male. Collected 

2 Feb 1900 by W. L. Abbott. Original number 295.

Type Locality: Sullivan (Lampi) Island, Mergui Archipelago, 

[Tanintharyi, Burma].

Genus Notomys Lesson, 1842

Notomys carpentarius Johnson

Proc. Biol. Soc. Wash. 72: 186, 4 Nov 1959.

= Notomys aquilo Thomas, 1921. See Ride (1970:227).

Holotype: USNM 284353. Skin and skull. Adult female. Collected 8 Jun 1948 by D. H. Johnson. Original number 

5525.

Type Locality: Umbakumba, 13°15′S, 136°45′E, Port Langdon, 

northeastern corner of Groote Eylandt, Northern Territory, 

Australia.

Genus Oenomys Thomas, 1904

Oenomys hypoxanthus vallicola Heller

Smithson. Misc. Collect. 63(7): 11, 24 Jun 1914.

= Oenomys hypoxanthus (Pucheran, 1855).
8 0 • smithsonian contributions to zoology

Holotype: USNM 162614. [Skin and skull]. Adult female. Collected 15 Jul 1909 by J. A. Loring. Original number 6640.

Type Locality: Lake Naivasha, [Rift Valley], British East Africa 

(Kenya).

Genus Otomys F. Cuvier, 1824

Otomys orestes dollmani Heller

Smithson. Misc. Collect. 59(16): 5, 5 Jul 1912.

= Otomys dollmani Heller, 1912. See Hollister (1919:147).

Holotype: USNM 181790. [Skin and skull]. Adult male. Collected 27 Aug 1911 by E. Heller. Original number 4125.

Type Locality: [SL: North Creek], Mount Gargues (Uaragess), altitude 7,000 ft (2,134 m), [SL: 6,000 ft (1,829 m)], Mathews 

Range, [Rift Valley], British East Africa (Kenya).

Genus Palawanomys Musser 

and Newcomb, 1983

Palawanomys furvus Musser and Newcomb

Bull. Am. Mus. Nat. Hist. 174: 335, 29 Jun 1983.

Holotype: USNM 478113. Skin and skull. Adult female. Collected 15 Apr 1962 by D. S. Rabor. Original number 1438.

Type Locality: Mount Mantalingajan, 4,500 ft (1,372 m), 

Brook’s Point Municipality, Palawan Province, Palawan Island, [Philippines].

Genus Paruromys Ellerman, 1954

Rattus dominator camurus Miller and Hollister

Proc. Biol. Soc. Wash. 34: 96, 30 Jun 1921b.

= Paruromys dominator (Thomas, 1921). See Musser and Newcomb (1983:404).

Holotype: USNM 219566. Skin and skull. Adult male. Collected 

15 Jan 1918 by H. C. Raven. Original number 3384.

Type Locality: Pinedapa, [Sulawesi Tengah], Middle Celebes (Sulawesi), [Indonesia].

Genus Pelomys Peters, 1852

Pelomys fallax iridescens Heller

Smithson. Misc. Collect. 59(16): 12, 5 Jul 1912.

= Pelomys fallax Peters, 1852.

Holotype: USNM 181801. [Skin and skull]. Adult male. Collected 5 Nov 1911 by E. Heller. Original number 4620.

Type Locality: [SL: Summit], Mount Mbololo, altitude 5,000 ft 

(1,524 m), Taita Mountains (Hills), [Coast Province, Kenya].

Genus Pithecheirops Emmons, 1993

Pithecheirops otion Emmons

Proc. Biol. Soc. Wash. 106: 753, 17 Dec 1993.

Holotype: USNM 574500. Skin, skull, and body, including left 

feet and organs, in fluid. Juvenile male. Collected 21 Sep 

1991 by L. H. Emmons. Original number [SL: 673], not 763 

as in original description.

Type Locality: About 1 km NE of Danum Valley Field Centre, 

by road, and about 500 m NW of road on a study plot trail, 

4°58′N, 117°48′E, elevation about 150 m, approximately 

75 km W Lahad Datu, State of Sabah, [Borneo], Malaysia.

Genus Praomys Thomas, 1892

Epimys taitae Heller

Smithson. Misc. Collect. 59(16): 9, 5 Jul 1912.

= Praomys taitae (Heller, 1912). See G. M. Allen and Lawrence 

(1936:95) for name combination and Carleton and Stanley 

(2012:444) for current use as a valid species.

Holotype: USNM 181797. [Skin and skull]. Adult male. Collected 5 Nov 1911 by E. Heller. Original number 4611.

Type Locality: [SL: Summit] Mount Mbololo, altitude 5,000 ft 

(1,524 m), Taita Mountains (Hills), [Coast Province], British East Africa (Kenya).

Mus peromyscus Heller

Smithson. Misc. Collect. 52: 472, 13 Nov 1909a.

= Praomys jacksoni (de Winton, 1897). See G. M. Allen (1939: 

410).

Holotype: USNM 161905. Skin and skull. Adult male. Collected 

9 Jun 1909 by E. Heller. Original number 1011.

Type Locality: Njoro O Nyiro, Sotik, [Rift Valley], British East 

Africa (Kenya).

Mus tullbergi rostratus Miller

Proc. Wash. Acad. Sci. 2: 637, 28 Dec 1900e.

= Praomys rostratus (Miller, 1900). See Van der Straten and Verheyen (1981:263).

Holotype: USNM 83836. Skin and skull. Adult male. Collected 

7 May 1897 by [SL: R. P. Currie]. Original number 60.

Type Locality: Mount Coffee, [Miller (1900e:631): altitude 

400 ft–500 ft (122–152 m)], [Montserrado], Liberia.

Genus Rattus G. Fischer, 1803

Epimys basilanus Hollister

Proc. U.S. Natl. Mus. 46: 322, 31 Dec 1913d.

= Rattus exulans (Peale, 1848). See Ellerman (1949:65).

Holotype: USNM 144635. Skin and skull. Adult male. Collected 

Feb 1906 by E. A. Mearns. Original number 6039.

Type Locality: Basilan Island, [Basilan Province, Philippines].

Epimys benguetensis Hollister

Proc. U.S. Natl. Mus. 46: 323, 31 Dec 1913d.

= Rattus tanezumi Temminck, 1844. See Musser and Carleton 

(1993:660).

Lectotype: USNM 145790 as designated by Musser (1977b:6). 

Skin only. Adult female. Collected 1 May 1907 by E. A. 

Mearns. Original number 6405.
number 642 • 8 1

Type Locality: Camp John Hay, Baguio, altitude 5,000 ft 

(1,524 m), Benguet Province, Luzon, [SL: P. I. (Philippines)].

Remarks: Musser (1977b:6) pointed out that the skull with this 

number is a Rattus nitidus and designated the skin as the 

lectotype.

Epimys calcis Hollister

Proc. Biol. Soc. Wash. 24: 89, 15 May 1911.

= Rattus exulans (Peale, 1848). See Ellerman (1949:65).

Holotype: USNM 145771. Skin and skull. Young adult male. 

Collected 8 May 1907 by E. A. Mearns. Original number 

6412.

Type Locality: Baguio, Limestone Hills, near Lime Kiln, at 5,000 

ft (1,524 m), Benguet Province, Luzon, P.I. (Philippines).

Epimys coloratus Hollister

Proc. U.S. Natl. Mus. 46: 317, 31 Dec 1913d.

= Rattus tanezumi Temminck, 1844. See Musser and Carleton 

(1993:660).

Holotype: USNM 144571. Skin and skull. Adult female. Collected 21 Feb 1906 by E. A. Mearns. Original number 6044.

Type Locality: Musser’s Plantation, 3 mi (5 km) [FC: 4 mi 

(6 km)] west of Isabela, [Basilan Province], Basilan Island, 

[Philippines].

Epimys fulmineus Miller

Smithson. Misc. Collect. 61(21): 9, 29 Dec 1913d.

= Rattus tiomanicus (Miller, 1900). See Musser and Carleton 

(2005:1492).

Holotype: USNM 112349. Skin and skull. Adult female. Collected 27 May 1901 by W. L. Abbott. Original number 950.

Type Locality: St. (Santa) Barbe Island (Pulau Pejantan), South 

China Sea, [Kepulauan Riau, Indonesia].

Epimys gala Miller

Proc. U.S. Natl. Mus. 38: 398, 19 Aug 1910b.

= Rattus everetti (Gunther, 1879). See Ellerman (1949:65).

Holotype: USNM 144633. Skin and skull. Adult male. Collected 

30 Nov 1906 by E. A. Mearns. Original number 6334.

Type Locality: Alag River, [Mindoro Oriental Province], Mindoro, [SL: P. I. (Philippines)].

Epimys leucophaeatus Hollister

Proc. U.S. Natl. Mus. 46: 320, 31 Dec 1913d.

= Rattus exulans (Peale, 1848). See Ellerman (1949:65).

Holotype: USNM 155144. Skin and skull. Adult female. Collected 27 May 1909 by D. B. Mackie. Original number 5.

Type Locality: [SL: Albay], Bagamanoc, [Catanduanes Province], 

Catanduanes Island, [Philippines].

Epimys maerens Miller

Proc. Biol. Soc. Wash. 24: 26, 24 Feb 1911c.

= Rattus tiomanicus (Miller, 1900). See Musser and Califia 

(1982:24).

Holotype: USNM 141193. [Skin and skull]. Adult female. Collected 11 Mar 1905 by W. L. Abbott. Original number 4062.

Type Locality: Mouth of Mojeia River, Nias Island, [Sumatera 

Utara]. Sumatra, [Indonesia].

Epimys mara Miller

Smithson. Misc. Collect. 61(21): 10, 29 Dec 1913d.

= Rattus tiomanicus (Miller, 1900). See Musser and Califia 

(1982:15).

Holotype: USNM 196751. Skin and skull. Adult female. Collected 28 Aug 1912 by H. C. Raven. Original number 187.

Type Locality: Maratua Island (Pulau Maratua), [Kalimantan 

Timur], Dutch Southeast Borneo (Kalimantan), [Indonesia].

Epimys mayonicus Hollister

Proc. U.S. Natl. Mus. 46: 319, 31 Dec 1913d.

= Rattus exulans (Peale, 1848). See Ellerman (1949:65).

Holotype: USNM 144600. Skin and skull. Adult male. Collected 

5 Jun 1907 by E. A. Mearns. Original number 6438.

Type Locality: Mount Mayon, altitude 4,000 ft (1,219 m), Albay 

Province, Luzon, [Philippines].

Epimys neglectus ducis Lyon

Proc. U.S. Natl. Mus. 40: 99, 25 Apr 1911.

= Rattus tiomanicus (Miller, 1900). See Medway (1965:121).

Holotype: USNM 145511. Skin and skull. Adult male. Collected 

4 May 1907 by W. L. Abbott. Original number 5174.

Type Locality: Pulo (Pulau) Datu, [Kalimantan Barat], off west 

coast of Borneo (Kalimantan), [Indonesia].

Epimys neglectus lamucotanus Lyon

Proc. U.S. Natl. Mus. 40: 100, 25 Apr 1911.

= Rattus tiomanicus (Miller, 1900). See Medway (1965:122).

Holotype: USNM 145497. Skin and skull. Adult female. Collected 10 May 1907 by W. L. Abbott. Original number 

5224.

Type Locality: Pulo Lamukotan (Pulau Lemukutan), [Kalimantan 

Barat], off west coast of Borneo (Kalimantan), [Indonesia].

Epimys norvegicus socer Miller

Proc. Biol. Soc. Wash. 27: 90, 11 May 1914.

= Rattus norvegicus (Berkenhout, 1769). See G. M. Allen (1926:8).

Holotype: USNM 144020. Skin and skull. Adult male. Collected 

30 Jan 1905 by W. W. Simpson. Original number 5.

Type Locality: Taocheo (Lintan), Kansu (Gansu), China.

Epimys ornatulus Hollister

Proc. U.S. Natl. Mus. 46: 322, 31 Dec 1913d.

= Rattus exulans (Peale, 1848). See Ellerman (1949:65).

Holotype: USNM 239245. Skin and skull. Adult female. Collected 8 Feb 1903 by R. C. McGregor and A. Celestino. 

Original number Philippine Bureau of Science 37.

Type Locality: Cagayancillo, Cagayan Island, [Palawan Province, Philippines].
8 2 • smithsonian contributions to zoology

Remarks: Published as Philippine Bureau of Science 37; cataloged as USNM 239245 on 12 Feb 1923.

Epimys pannellus Miller

Smithson. Misc. Collect. 61(21): 8, 29 Dec 1913d.

= Rattus tanezumi Temminck, 1844. See Musser and Carleton 

(1993:660).

Holotype: USNM 104121. Skin and skull. Adult male. Collected 

19 Dec 1899 by W. L. Abbott. Original number 175.

Type Locality: Pulo Rawi (Ko Ra Wi), Butang Islands (Batong 

Group), [Satun, Thailand].

Epimys querceti Hollister

Proc. Biol. Soc. Wash. 24: 90, 15 May 1911.

= Rattus exulans (Peale, 1848). See Ellerman (1949:65).

Holotype: USNM 145833. Skin and skull. Adult male. Collected 

1 Aug 1907 by E. A. Mearns. Original number 6491.

Type Locality: Haights-in-the-Oaks, altitude 7,000 ft (2,134 m), 

Benguet Province, P. I. (Philippines).

Epimys rattus dentatus Miller

Smithson. Misc. Collect. 61(21): 14, 29 Dec 1913d.

= Rattus tanezumi Temminck, 1844. See Musser and Carleton 

(1993:660).

Holotype: USNM 111929. Skin and skull. Adult male. Collected 

13 Dec 1900 by W. L. Abbott. Original number 783.

Type Locality: Hastings Island, Mergui Archipelago, [Tanintharyi, Burma].

Epimys rattus exsul Miller

Smithson. Misc. Collect. 61(21): 15, 29 Dec 1913d.

= Rattus tanezumi Temminck, 1844. See Musser and Carleton 

(1993:660).

Holotype: USNM 124046. Skin and skull. Adult male. Collected 

31 Dec 1903 by W. L. Abbott. Original number 2958.

Type Locality: James Island (Pulo Buda), Mergui Archipelago, 

[Tanintharyi, Burma].

Epimys rattus fortunatus Miller

Smithson. Misc. Collect. 61(21): 15, 29 Dec 1913d.

= Rattus tanezumi Temminck, 1844. See Musser and Carleton 

(1993:660).

Holotype: USNM 104148. Skin and skull. Adult male. Collected 

29 Dec 1899 by W. L. Abbott. Original number 191.

Type Locality: Chance Island, Mergui Archipelago, [Tanintharyi, 

Burma].

Epimys rattus insulanus Miller

Smithson. Misc. Collect. 61(21): 14, 29 Dec 1913d.

= Rattus tanezumi Temminck, 1844. See Musser and Carleton 

(1993:660).

Holotype: USNM 104147. Skin and skull. Adult male. Collected 

7 Mar 1900 by W. L. Abbott. Original number 334.

Type Locality: Helfer (Hel-fer) Island, Mergui Archipelago, 

[Tanintharyi, Burma].

Epimys rattus pauper Miller

Smithson. Misc. Collect. 61(21): 13, 29 Dec 1913d.

= Rattus tiomanicus (Miller, 1900). See Musser and Califia (1982:24).

Holotype: USNM 104828. Skin and skull. Adult male. Collected 

4 Jun 1900 by W. L. Abbott. Original number 433.

Type Locality: Sirhassen Island (Pulau Serasan), South Natuna 

Islands (Kepulauan Natuna Selatan), [Kepulauan Riau, 

Indonesia].

Epimys rattus shigarus Miller

Proc. Biol. Soc. Wash. 26: 198, 23 Oct 1913a.

= Rattus pyctoris (Hodgson, 1845). See Musser and Carleton 

(2005:1482).

Holotype: USNM 176132. Skin and skull. Adult male. Collected 

4 Oct 1912 by [SL: W. L. Abbott]. Original number 8010.

Type Locality: [FC: Mango, on a hillside about 1,300 ft (396 m) 

above bottom], Shigar Valley, altitude 9,000 ft (2,743 m), 

Baltistan (Gilgit-Baltistan), [Northern Areas, Pakistan].

Epimys rattus turbidus Miller

Smithson. Misc. Collect. 61(21): 12, 29 Dec 1913d.

= Rattus tanezumi Temminck, 1844. See Musser and Carleton 

(1993:660).

Holotype: USNM 196746. Skin and skull. Young adult female. 

Collected 21 Jun 1912 by H. C. Raven. Original number 81.

Type Locality: Tanggarung (Tenggarong), south bank of Mahakam River (Sungai Mahakam), [Kalimantan Timur], 

Dutch Southeast Borneo (Kalimantan), [Indonesia].

Epimys rattus viclana Miller

Smithson. Misc. Collect. 61(21): 13, 29 Dec 1913d.

= Rattus tiomanicus (Miller, 1900). See Musser and Califia (1982:24).

Holotype: USNM 123861. Skin and skull. Adult female, not 

male as given in original description. Collected 3 Nov 1903 

by W. L. Abbott. Original number 2800.

Type Locality: Pulo Lankawi (Pulau Langkawi), off west coast of 

Malay Peninsula, [Kedah, Malaysia].

Epimys roa Miller

Smithson. Misc. Collect. 61(21): 10, 29 Dec 1913d.

= Rattus tiomanicus (Miller, 1900). See Musser and Califia (1982:24).

Holotype: USNM 112444. Skin and skull. Adult female. Collected 8 Jun 1901 by W. L. Abbott. Original number 1025.

Type Locality: Pulo Aor (Pulau Aur), off east coast of Johore 

(Johor), [Malaysia].

Epimys robiginosus Hollister

Proc. U.S. Natl. Mus. 46: 318, 31 Dec 1913d.

= Rattus tanezumi Temminck, 1844. See Musser and Carleton 

(1993:660).

Holotype: USNM 239426. Skin and skull. Adult male. Collected 

7 Feb 1903 by R. C. McGregor and A. Celestino. Original 

number Philippine Bureau of Science 33.

Type Locality: Cagayancillo, Cagayan Island, [Palawan Province, Philippines].
number 642 • 8 3

Remarks: Published as Philippine Bureau of Science 33; cataloged as USNM 239426 on 12 Feb 1923.

Epimys tingius Miller

Smithson. Misc. Collect. 61(21): 9, 29 Dec 1913d.

= Rattus tiomanicus (Miller, 1900). See Musser and Califia (1982:24).

Holotype: USNM 112723. Skin and skull. Adult male. Collected 

5 Aug 1901 by W. L. Abbott. Original number 1201.

Type Locality: Pulo (Pulau) Tinggi, off east coast of Johore 

(Johor), [Malaysia].

Epimys tua Miller

Smithson. Misc. Collect. 61(21): 12, 29 Dec 1913d.

= Rattus tiomanicus (Miller, 1900). See Musser and Califia (1982:15).

Holotype: USNM 196752. Skin and skull. Adult female. Collected 28 Aug 1912 by H. C. Raven. Original number 188.

Type Locality: Maratua Island (Pulau Maratua), [Kalimantan 

Timur], Dutch Southeast Borneo (Kalimantan), [Indonesia].

Epimys tyrannus Miller

Proc. U.S. Natl. Mus. 38: 397, 19 Aug 1910b.

= Rattus everetti (Gunther, 1879). See Musser and Carleton 

(1993:651).

Holotype: USNM 239247. Skin and skull. Adult male. Collected 

15 May 1902 by R. C. McGregor and A. Celestino. Original 

number Philippine Bureau of Science 8.

Type Locality: Ticao [Island, Masbate Province, Philippines].

Remarks: Published as Philippine Bureau of Science 8; cataloged 

as USNM 239247 on 12 Feb 1923.

Epimys vigoratus Hollister

Proc. U.S. Natl. Mus. 46: 321, 31 Dec 1913d.

= Rattus exulans (Peale, 1848). See Ellerman (1949:65).

Holotype: USNM 144637. Skin and skull. Adult male. Collected 

15 Nov 1906 by E. A. Mearns. Original number 6306.

Type Locality: Mount Halcon, spur of main ridge at 4,500 ft (1,372 

m), Mindoro, [Oriental Mindoro Province, Philippines].

Limnomys picinus Hollister

Proc. U.S. Natl. Mus. 46: 325, 31 Dec 1913d.

= Rattus mindorensis (Thomas, 1898). See Musser (1977a:9).

Lectotype: USNM 144605, as selected by Musser (1977a:9). 

Skin only. Adult female. Collected 16 Nov 1906 by E. A. 

Mearns. Original number 6311.

Type Locality: Spur of main ridge of Mount Halcon, altitude 

4,500 ft (1,372 m), Mindoro, [Oriental Mindoro Province, 

Philippines].

Remarks: According to Musser (1977a:9), the skull originally 

associated with Hollister’s type specimen is a Limnomys 

mearnsi and belongs to USNM 144620. Musser (1977a:9) 

selected the skin to be the lectotype.

Mus albigularis Mearns

Proc. U.S. Natl. Mus. 28: 440, 13 May 1905.

= Rattus everetti (Gunther, 1879). See Ellerman (1949:65).

Holotype: USNM 125258. Skin and skull. Adult male. Collected 

3 Jul 1904 by E. A. Mearns. Original number 5699.

Type Locality: Mount Apo, 7,600 ft (2,317 m), southern Mindanao, Philippine Islands (Philippines).

Mus atratus Miller

Proc. U.S. Natl. Mus. 24: 767, 29 May 1902b. 

Primary junior homonym of Mus atratus Philippi, 1900. Renamed Mus atridorsum by Miller (1903d:50).

= Rattus rattus (Linnaeus, 1758). See Musser and Newcomb 

(1983:481).

Holotype: USNM 111868. Skin and skull. Adult female. Collected 7 Jan 1901 by W. L. Abbott. Original number 818.

Type Locality: Barren Island, Andamans [Andaman and Nicobar 

Islands, India].

Mus bullatus Lyon

Proc. U.S. Natl. Mus. 34: 646, 14 Sep 1908.

= Rattus annandalei (Bonhote, 1903). See Chasen (1940:161).

Holotype: USNM 143447. Skin and skull. Adult male. Collected 

16 Mar 1906 by W. L. Abbott. Original number 4723.

Type Locality: Pulo (Pulau) Rupat, off east coast of Sumatra, 

[Riau, Indonesia].

Mus burrescens Miller

Proc. U.S. Natl. Mus. 24: 771, 29 May 1902b.

= Rattus burrus (Miller, 1902). See Musser and Heaney (1985:21).

Holotype: USNM 111789. Skin and skull. Adult female. Collected 12 Mar 1901 by W. L. Abbott. Original number 926.

Type Locality: Great Nicobar Island, [Andaman and Nicobar 

Islands, India].

Mus burrulus Miller

Proc. U.S. Natl. Mus. 24: 770, 29 May 1902b.

= Rattus andamanensis (Blyth, 1860). See Musser and Carleton 

(2005:1464).

Holotype: USNM 111817. Skin and skull. Adult male. Collected 

25 Jan 1901 by W. L. Abbott. Original number 865. 

Type Locality: Car Nicobar, [Andaman and Nicobar Islands, 

India].

Mus burrus Miller

Proc. U.S. Natl. Mus. 24: 768, 29 May 1902b.

= Rattus burrus (Miller, 1902). See Hayman and Holt (1941:184).

Holotype: USNM 111811. Skin and skull. Adult female. Collected 5 Feb 1901 by W. L. Abbott. Original number 881.

Type Locality: Trinkut (Trinkat) Island, Nicobars [Andaman and 

Nicobar Islands, India].

Mus buruensis J. A. Allen

Bull. Am. Mus. Nat. Hist. 30: 336, 21 Dec 1911.

= Rattus exulans (Peale, 1848). See Ellerman (1949:64).

Holotype: USNM 175900. [Skin and skull]. Adult male. Collected 11 Dec 1909 by R. C. Andrews. Original number [SL: 

24]. AMNH number 31296.
8 4 • smithsonian contributions to zoology

Type Locality: Bouru Island, (Pulau Buru), Molucca Islands, [SL: 

Dutch East Indes], [Maluku, Indonesia].

Remarks: Published as AMNH 31296; cataloged as USNM 

175900 on 13 Mar 1913.

Mus clabatus Lyon

Proc. U.S. Natl. Mus. 31: 596, 18 Dec 1906.

= Rattus exulans (Peale, 1848). See Ellerman (1949:64).

Holotype: USNM 124888. Skin and skull. Old adult female. Collected 25 Jun 1904 by W. L. Abbott. Original number 3439.

Type Locality: Klabat Bay (Teluk Klabat), island of Banka (Pulau 

Bangka), east of Sumatra, [Kepulauan Bangka Belitung, 

Indonesia].

Mus enganus Miller

Proc. U.S. Natl. Mus. 30: 821, 4 Jun 1906b.

= Rattus enganus (Miller, 1906). See Lyon (1916a:460).

Holotype: USNM 140976. Skin and skull. Adult male. Collected 

4 Dec 1904 by W. L. Abbott. Original number 3823.

Type Locality: Engano Island (Pulau Enggano), [Bengkulu, Sumatra, Indonesia].

Mus exulans Peale

U.S. Explor. Exped. 8: 47, 1848.

= Rattus exulans (Peale, 1848). See Tate (1935:146).

Lectotype: USNM 3730, as designated by Lyon and Osgood 

(1909:148). Mounted skin. Age and sex unknown. Collected on unknown date by the U.S. Exploring Expedition, 

1838–1842. Cataloged 20 Dec 1859. No original number.

Type Locality: Tahiti, [MC: Society Islands].

Remarks: Type not designated by number in original description, which mentioned specimens from several islands. This 

specimen was mounted on a stand, the bottom of which 

bears the following information: “3730 Mus exulans, Peale 

(Type) (Mus penicillatus, Gould) Tahiti Rat Society Ids. T. 

R. Peale.” Lyon and Osgood (1909:148) considered this 

specimen to be the “type,” an identification that effectively 

served to designate the lectotype (ICZN, 1999: Article 

74.5). Poole and Schantz (1942:462) confirmed the designation of USNM 3730 as the lectotype.

Mus flebilis Miller

Proc. U.S. Natl. Mus. 24: 762, 29 May 1902b.

= Rattus andamanensis (Blyth, 1860). See Musser and Carleton 

(2005:1464).

Holotype: USNM 111841. Skin and skull. Adult female. Collected 10 Jan 1901 by W. L. Abbott. Original number 827.

Type Locality: Henry Lawrence Island, Andamans [Andaman 

and Nicobar Islands, India].

Mus julianus Miller

Smithson. Misc. Collect. 45: 34, 6 Nov 1903b.

= Rattus tiomanicus (Miller, 1900). See Musser and Califia 

(1982:24).

Holotype: USNM 112393. Skin and skull. Adult female. Collected 2 Jun 1901 by W. L. Abbott. Original number 987.

Type Locality: St. Julian Island (Kemudi), South China Sea, 

[Kepulauan Riau, Indonesia].

Mus kelleri Mearns

Proc. U.S. Natl. Mus. 28: 444, 13 May 1905.

= Rattus tanezumi Temminck, 1844. See Musser and Carleton 

(1993:660).

Holotype: USNM 125278. Skin and skull. Adult female. Collected 20 Jul 1904 by E. A. Mearns. Original number 5738.

Type Locality: [SL: Commissary building at] Davao, [Davao 

City Province], southern Mindanao, Philippine Islands 

(Philippines).

Mus lugens Miller

Smithson. Misc. Collect. 45: 33, 6 Nov 1903b.

= Rattus lugens (Miller, 1903). See Musser and Heaney (1985:21).

Holotype: USNM 121533. Skin and skull. Adult female. Collected 15 Nov 1902 by W. L. Abbott. Original number 2046.

Type Locality: North Pagi Island (Pulau Pagai Utara), [Sumatera 

Barat], Sumatra, [Indonesia].

Mus magnirostris Mearns

Proc. U.S. Natl. Mus. 28: 441, 13 May 1905.

= Rattus norvegicus (Berkenhout, 1769). See Musser (1977b:12).

Holotype: USNM 125212. Skin and skull. Adult female. Collected 15 Jan 1904 by E. A. Mearns. Original number 5639.

Type Locality: Old Spanish hospital, Zamboanga, [Zamboanga 

Province], western Mindanao, Philippine Islands (Philippines).

Mus mindanensis Mearns

Proc. U.S. Natl. Mus. 28: 442, 13 May 1905.

= Rattus tanezumi Temminck, 1844. See Musser and Carleton 

(1993:660).

Holotype: USNM 125274. Skin and skull. Adult male. Collected 

9 Jul 1904 by E. A. Mearns. Original number 5719.

Type Locality: Todaya, altitude 4,000 ft (1,219 m), Mount Apo, 

[Davao del Sur Province], southern Mindanao, [FC: Philippine Islands (Philippines)].

Mus obscurus Miller

Proc. Wash. Acad. Sci. 2: 213, 20 Aug 1900f. 

Primary junior homonym of Mus obscurus Waterhouse, 1837. 

Renamed Mus pullus by Miller (1901a:178).

= Rattus exulans (Peale, 1848). See Ellerman (1949:64).

Holotype: USNM 101764. Skin and skull. Adult male. Collected 

1 Oct 1899 by [MC: W. L. Abbott]. No original number.

Type Locality: Tioman Island (Pulau Tioman), [Pahang, Malaysia].

Mus pannosus Miller

Proc. Biol. Soc. Wash. 13: 190, 21 Dec 1900g.

= Rattus tanezumi Temminck, 1844. See Musser and Carleton 

(1993:660).
number 642 • 8 5

Holotype: USNM 104110. [Skin and skull]. Adult male. Collected 14 Dec 1899 by [MC: W. L. Abbott]. Original 

number 146.

Type Locality: Pulo (Ko) Adang, Butang Islands (Batong Group), 

[Satun, Thailand].

Mus pantarensis Mearns

Proc. U.S. Natl. Mus. 28: 448, 13 May 1905.

= Rattus exulans (Peale, 1848). See Ellerman (1949:65).

Holotype: USNM 123294. Skin and skull. Adult female. Collected 4 Sep 1903 by E. A. Mearns. Original number 5622.

Type Locality: Pantar, altitude 1,907 ft (581 m), Mindanao, Philippine Islands (Philippines).

Mus pulliventer Miller

Proc. U.S. Natl. Mus. 24: 765, 29 May 1902b.

= Rattus tanezumi Temminck, 1844. See Musser and Carleton 

(1993:660).

Holotype: USNM 111790. Skin and skull. Adult female. Collected 12 Mar 1901 by W. L. Abbott. Original number 927.

Type Locality: Great Nicobar Island, Nicobars [Andaman and 

Nicobar Islands, India].

Mus siantanicus Miller

Proc. Wash. Acad. Sci. 2: 210, 20 Aug 1900f.

= Rattus tiomanicus (Miller, 1900). See Musser and Califia 

(1982:24).

Holotype: USNM 101705. Skin and skull. Adult male. Collected 

11 Sep 1899 by [MC: W. L. Abbott]. No original number.

Type Locality: Pulo (Pulau) Siantan, Anamba Islands (Kepulauan 

Anambas), [Kepulauan Riau, Indonesia].

Mus simalurensis Miller

Proc. U.S. Natl. Mus. 26: 458, 3 Feb 1903c.

= Rattus simalurensis (Miller, 1903). See Hayman and Holt 

(1941:181).

Holotype: USNM 114216. Skin and skull. Adult female. Collected 14 Dec 1901 by W. L. Abbott. Original number 1372.

Type Locality: Simalur Island (Pulau Simeulue), [Aceh, Sumatra, 

Indonesia].

Mus stoicus Miller

Proc. U.S. Natl. Mus. 24: 759, 29 May 1902b.

= Rattus stoicus (Miller, 1902). See Poole and Schantz (1942:481).

Holotype: USNM 111834. Skin and skull. Adult male. Collected 

9 Jan 1901 by W. L. Abbott. Original number 820.

Type Locality: Henry Lawrence Island, Andamans [Andaman 

and Nicobar Islands, India].

Mus surdus Miller

Proc. U.S. Natl. Mus. 26: 460, 3 Feb 1903c.

= Rattus exulans (Peale, 1848). See Ellerman (1949:64).

Holotype: USNM 114184. Skin and skull. Adult male. Collected 

11 Dec 1901 by W. L. Abbott. Original number 1359.

Type Locality: Simalur Island (Pulau Simeulue), [Aceh, Sumatra, 

Indonesia].

Mus taciturnus Miller

Proc. U.S. Natl. Mus. 24: 762, 29 May 1902b.

= Rattus stoicus (Miller, 1902). See Musser and Newcomb 

(1983:481).

Holotype: USNM 111828. Skin and skull. Adult male. Collected 

16 Jan 1901 by W. L. Abbott. Original number 854.

Type Locality: [FC: MacPherson Strait], South Andaman Island, 

Andamans [Andaman and Nicobar Islands, India].

Mus tagulayensis Mearns

Proc. U.S. Natl. Mus. 28: 439, 13 May 1905.

= Rattus everetti (Gunther, 1879). See Ellerman (1949:65).

Holotype: USNM 125264. Skin and skull. Adult male. Collected 

15 Jul 1904 by E. A. Mearns. Original number 5732.

Type Locality: At sea level at Tagulaya, on the Gulf of Davao, 

foot of Mount Apo, [Davao del Sur Province], southern 

Mindanao, Philippine Islands (Philippines).

Mus tambelanicus Miller

Proc. Wash. Acad. Sci. 2: 212, 20 Aug 1900f.

= Rattus tiomanicus (Miller, 1900). See Musser and Califia 

(1982:24).

Holotype: USNM 101665. Skin and skull. Adult male. Collected 

10 Aug 1899 by [MC: W. L. Abbott]. No original number.

Type Locality: Big Tambelan Island (Pulau Tambelan Besar), 

[Kepulauan Tambelan, Kepulauan Riau, Indonesia].

Mus tiomanicus Miller

Proc. Wash. Acad. Sci. 2: 209, 20 Aug 1900f.

= Rattus tiomanicus (Miller, 1900). See Medway (1965:118).

Holotype: USNM 101763. Skin and skull. Adult male. Collected 

4 Oct 1899 by [MC: W. L. Abbott]. No original number.

Type Locality: Tioman Island (Pulau Tioman), [MC: off Malay 

Peninsula], [Pahang, Malaysia].

Mus todayensis Mearns

Proc. U.S. Natl. Mus. 28: 445, 13 May 1905.

= Rattus exulans (Peale, 1848). See Ellerman (1949:64).

Holotype: USNM 125224. Skin and skull. Adult female. Collected 11 Jul 1904 by E. A. Mearns. Original number 5722.

Type Locality: Todaya, altitude 4,000 ft (1,219 m), on Mount 

Apo, [Davao del Sur Province], southern Mindanao, Philippine Islands (Philippines).

Mus vitiensis Peale

U.S. Explor. Exped. 8: 49, 1848.

= Rattus exulans (Peale, 1848). See Tate (1935:146).

Holotype: USNM 3731. Skin only. Sex and age not given. Collected on unknown date by U.S. Exploring Expedition, 

1838–1842. Cataloged 20 Dec 1859. No original number.

Type Locality: Feejee (Fiji) Islands.
8 6 • smithsonian contributions to zoology

Remarks: Type not designated by number in original description, which referenced only a single specimen. USNM 3731 

is the only specimen collected by the Exploring Expedition 

from Fiji and becomes the holotype as fixed by monotypy 

(ICZN, 1999: Article 73.1.2). The specimen was reported 

as missing by Lyon and Osgood (1909:289). Poole and 

Schantz (1942:486) reported that the specimen was originally mounted and on display prior to Jun 1910 when it was 

made into a study skin.

Mus vulcani Mearns

Proc. U.S. Natl. Mus. 28: 446, 13 May 1905.

= Rattus exulans (Peale, 1848). See Ellerman (1949:64).

Holotype: USNM 125216. Skin and skull. Adult female. Collected 26 Jun 1904 by E. A. Mearns. Original number 5674.

Type Locality: Mount Apo, 7,600 ft (2,317 m), southern Mindanao, Philippine Islands (Philippines).

Mus vulcani apicis Mearns

Proc. U.S. Natl. Mus. 28: 447, 13 May 1905.

= Rattus exulans (Peale, 1848). See Ellerman (1949:64).

Holotype: USNM 125229. Skin, skull, [and carcass in alcohol]. 

Adult female. Collected 6 Jun 1904 by E. A. Mearns. Original number 5709.

Type Locality: [SL: West peak], summit of Mount Apo, altitude 

9,700 ft (2,957 m), southern Mindanao, Philippine Islands 

(Philippines).

Mus zamboangae Mearns

Proc. U.S. Natl. Mus. 28: 443, 13 May 1905.

= Rattus tanezumi Temminck, 1844. See Musser and Carleton 

(1993:660).

Holotype: USNM 125279. Skin and skull. Adult male. Collected 

20 Jan 1904 by E. A. Mearns. Original number 5753.

Type Locality: Zamboanga, [Zamboanga Province], western 

Mindanao, Philippine Islands (Philippines).

Rattus concolor solatus Kellogg

Proc. Biol. Soc. Wash. 58: 65, 7 May 1945c.

= Rattus exulans (Peale, 1848). See Laurie and Hill (1954:106).

Holotype: USNM 277317. Skin and skull. Adult male. Collected 23 Oct 1944 by J. F. C. [SL: Cassel] and R. M. R. [SL: 

Roecker]. Original number 4.

Type Locality: [MC: 1 mi (1.6 km) N of Wama], Morotai Island 

(Palau Morotai), Gilolo Group, Moluccas (Maluku Utara), 

Netherlands East Indies (Indonesia).

Rattus facetus Miller and Hollister

Proc. Biol. Soc. Wash. 34: 96, 30 Jun 1921b.

= Rattus marmosurus Thomas, 1921. See Musser (1973:24).

Holotype: USNM 218677. Skin and skull. Adult female. Collected 15 Jan 1917 by H. C. Raven. Original number 3082.

Type Locality: [FC: Sungai Soepoi], Goenoeng Lehio, southwest 

of Lake Lindoe (Rano Lindu), altitude 6,000 ft (1,829 m) 

[FC: between 6,000–7,166 ft (1,829–2,184m)], Middle Celebes (Sulawesi), [Sulawesi Tengah, Indonesia].

Rattus hoffmanni linduensis Miller and Hollister

Proc. Biol. Soc. Wash. 34: 70, 31 Mar 1921a.

= Rattus hoffmanni (Matschie, 1901).

Holotype: USNM 218700. Skin and skull. Adult female. Collected 28 Mar 1917 by H. C. Raven. Original number 3141.

Type Locality: Tomado, Lake Lindoe (Rano Lindu), Middle Celebes (Sulawesi), [Sulawesi Tengah, Indonesia].

Rattus hoffmanni subditivus Miller and Hollister

Proc. Biol. Soc. Wash. 34: 70, 31 Mar 1921a.

= Rattus nitidus (Hodgson, 1845). See Musser (1971b:4).

Holotype: USNM 219691. Skin and skull. Adult female. Collected 18 Sep 1917 by H. C. Raven. Original number 3270.

Type Locality: Toware, Bada, Middle Celebes (Sulawesi), 

[Indonesia].

Rattus humiliatus insolatus A. B. Howell

Proc. Biol. Soc. Wash. 40: 44, 5 Mar 1927b.

= Rattus norvegicus (Berkenhout, 1769). See Musser and Newcomb (1985:16).

Holotype: USNM 172569. Skin and skull. Adult female. Collected 

12 Jan 1909 by A. de C. Sowerby. Original number 413.

Type Locality: 12 mi (19 km) south of Yenanfu (Yah’an), altitude 

4,000 ft (1,219 m), Shensi (Shaanxi), China.

Rattus humiliatus sowerbyi A. B. Howell

Proc. Biol. Soc. Wash. 41: 42, 16 Mar 1928.

= Rattus norvegicus (Berkenhout, 1769). See Ellerman (1951:589).

Holotype: USNM 199620. Skin and skull. Young adult male. 

Collected 15 Oct 1914 by A. de C. Sowerby. Original 

number 735.

Type Locality: Near Imienpo (Yimianpo), altitude 500 ft (152 

m), north Kirin (Heilongjiang), Manchuria (China).

Rattus mollicomus Miller and Hollister

Proc. Biol. Soc. Wash. 34: 71, 31 Mar 1921a.

= Rattus hoffmanni (Matschie, 1901). See Musser (1971b:17).

Holotype: USNM 217752. Skin and skull. Adult male. Collected 

10 Apr 1916 by H. C. Raven. Original number 2433.

Type Locality: Goenoeng Kalabat (Gunung Klabat), altitude 

6,500 ft (1,981 m), [Sulawesi Utara], northeastern Celebes 

(Sulawesi), [Indonesia].

Rattus morotaiensis Kellogg

Proc. Biol. Soc. Wash. 58: 66, 7 May 1945c.

Holotype: USNM 277312. Skin and skull. Adult male. Collected 26 Oct 1944 by J. F. C. [SL: Cassel] and R. M. R. [SL: 

Roecker]. Original number 9.

Type Locality: [MC: 1 mi (1.6 km) N of Wama], Morotai Island 

(Palau Morotai), Gilolo Group, Moluccas (Maluku Utara), 

Netherlands East Indies (Indonesia).
number 642 • 8 7

Rattus palelae Miller and Hollister

Proc. Biol. Soc. Wash. 34: 69, 31 Mar 1921a.

= Rattus tanezumi Temminck, 1844. See Musser and Carleton 

(1993:660).

Holotype: USNM 200063. Skin and skull. Adult female. Collected 2 Aug 1914 by H. C. Raven. Original number 1619.

Type Locality: Pulo Paleleh (Pulau Panjang), [Sulawesi Tengah], 

north coast of Celebes (Sulawesi), [Indonesia].

Rattus rattus ambersoni Schwarz and Schwarz

An. Esc. Nac. Cienc. Biol. Mex. 14: 130, 15 Mar 1967.

= Rattus tiomanicus (Miller, 1900). See Musser and Califia 

(1982:25).

Holotype: USNM 197537. Skin and skull. Adult male. Collected 

28 May 1913, not 26 May as in original description, by H., 

not R. as in original description, C. Raven. Original number 

685.

Type Locality: Sangka Laki Island (Pulau Sanga Laki), Maratua 

Group, [Kalimantan Timur], east coast of Borneo (Kalimantan), [Indonesia].

Rattus rattus exiguus A. B. Howell

Proc. Biol. Soc. Wash. 40: 43, 5 Mar 1927b.

= Rattus losea (Swinhoe, 1871). See G. M. Allen (1940:1005).

Holotype: USNM 238185. Skin and skull. Adult female. Collected 1 Dec 1921 by A. de C. Sowerby. Original number 

1139.

Type Locality: 70 mi (113 km) southwest of Yenpingfu (Nanping), altitude 500 ft (152 m), Fukien (Fujian), China.

Rattus rattus koratensis Kloss

J. Nat. Hist. Soc. Siam 3: 379, 31 Dec 1919.

= Rattus andamanensis (Blyth, 1860). See Musser and Carleton 

(2005:1464).

Holotype: USNM 221512. Skin and skull. Adult female. Collected 19 Oct 1916 by C. B. Kloss. Original number 2196.

Type Locality: Lat Bua Kao (Sathani Lat Bua Khao), [Nakhon 

Ratchasima], East Siam (Thailand).

Remarks: Published as CBK 2196; cataloged as USNM 221512 

on 1 Jul 1920.

Rattus rattus kramensis Kloss

J. Nat. Hist. Soc. Siam 3: 379, 31 Dec 1919.

= Rattus tanezumi Temminck, 1844. See Musser and Carleton 

(1993:660).

Holotype: USNM 221521. Skin and skull. Adult female. Collected 30 Oct 1916 by C. B. Kloss. Original number 2281.

Type Locality: Koh Kram (Ko Khram), Inner Gulf of Siam, 

[Chon Buri, Thailand].

Remarks: Published as CBK 2281; cataloged as USNM 221521 

on 2 Jul 1920.

Rattus rattus lanensis Kloss

J. Nat. Hist. Soc. Siam 3: 378, 31 Dec 1919.

= Rattus tanezumi Temminck, 1844. See Musser and Carleton 

(1993:660).

Holotype: USNM 221523. Skin and skull. Adult female. Collected 29 Oct 1916 by C. B. Kloss. Original number 2261.

Type Locality: Koh (Ko) Lan, Inner Gulf of Siam, [Chon Buri, 

Thailand].

Remarks: Published as CBK 2261; cataloged as USNM 221523 

on 2 Jul 1920.

Rattus rattus mansorius Johnson

Proc. Biol. Soc. Wash. 75: 317, 31 Dec 1962.

= Rattus tanezumi Temminck, 1844. See Musser and Carleton 

(1993:660).

Holotype: USNM 302027. Skin and skull. Adult female. Collected 1 Sep 1955 by W. B. Jackson. Original number 31.

Type Locality: Kolonia, [SL: across from “Circle”], Ponape [Island], Caroline Islands.

Rattus rattus mesanis Kloss

J. Nat. Hist. Soc. Siam 3: 379, 31 Dec 1919.

= Rattus tanezumi Temminck, 1844. See Musser and Carleton 

(1993:660).

Holotype: USNM 221535. Skin and skull. Adult male. Collected 

2 Nov 1916 by C. B. Kloss. Original number 2320.

Type Locality: Koh Mesan Island (Ko Samae San), near Cape 

Liant (Laem Samae San), [Chon Buri], S. E. Siam (Thailand).

Remarks: Published as CBK 2320; cataloged as USNM 221535 

on 2 Jul 1920.

Rattus rattus piperis Schwarz and Schwarz

An. Esc. Nac. Cienc. Biol. Mex. 14: 132, 15 Mar 1967.

= Rattus tiomanicus (Miller, 1900). See Musser and Califia 

(1982:25).

Holotype: USNM 151961. Skin and skull. Adult female, not 

male as in original description. Collected 11 Dec 1907 by 

W. L. Abbott. Original number 5608.

Type Locality: Mata Siri Island (Pulau Matasiri), [Kalimantan Selatan], south of Borneo (Kalimantan), Java Sea, [Indonesia].

Rattus rattus umbriventer Kellogg

Proc. Biol. Soc. Wash. 58: 121, 20 Sep 1945a.

= Rattus argentiventer (Robinson and Kloss, 1916). See Musser 

(1973:12).

Holotype: USNM 277675. Skin and skull. Adult male. Collected 

31 May 1945 by A. L. Gordon. Original number 69.

Type Locality: 1 mi (1.6 km) SW Progreso, altitude about 20 ft (6 

m), Mindoro Island, [Mindoro Occidental Province], Philippine Islands (Philippines).

Rattus raveni Miller and Hollister

Proc. Biol. Soc. Wash. 34: 68, 31 Mar 1921a.

= Rattus exulans (Peale, 1848). See Ellerman (1949:64).

Holotype: USNM 199976. Skin and skull. Adult male. Collected 

16 Dec 1914 by H. C. Raven. Original number 1963.
8 8 • smithsonian contributions to zoology

Type Locality: Toli Toli (Tolitoli), [Sulawesi Tengah], North Celebes (Sulawesi), [Indonesia].

Rattus raveni eurous Miller and Hollister

Proc. Biol. Soc. Wash. 34: 69, 31 Mar 1921a.

= Rattus exulans (Peale, 1848). See Laurie and Hill (1954:106).

Holotype: USNM 199927. Skin and skull. Adult male. Collected 

15 Oct 1914 by H. C. Raven. Original number 1724.

Type Locality: Molengkapoti, Kwandang, [Gorontalo], North 

Celebes (Sulawesi), [Indonesia].

Rattus simalurensis babi Lyon

Proc. U.S. Natl. Mus. 52: 447, 30 Dec 1916a.

= Rattus simalurensis (Miller, 1903).

Holotype: USNM 114280. Skull only. Adult male. Collected 13 

Jan 1902 by W. L. Abbott. No original number.

Type Locality: Pulo (Pulau) Babi, [MC: 17 mi (27 km) SE Simalur Island (Pulau Simeulue)], [Aceh, Sumatra, Indonesia].

Rattus simalurensis lasiae Lyon

Proc. U.S. Natl. Mus. 52: 446, 30 Dec 1916a.

= Rattus simalurensis (Miller, 1903).

Holotype: USNM 114254. Skin and skull. Adult female. Collected 6 Jan 1902 by W. L. Abbott. Original number 1400.

Type Locality: Pulo (Pulau) Lasia, [FC: 14 mi (23 km) SE of Simalur Island (Pulau Simeulue)], [Aceh, Sumatra, Indonesia].

Genus Sundamys Musser 

and Newcomb, 1983

Epimys borneanus Miller

Smithson. Misc. Collect. 61(21): 15, 29 Dec 1913d.

= Sundamys muelleri (Jentink, 1879). See Musser and Newcomb 

(1983:401).

Holotype: USNM 196749. Skin and skull. Adult female. Collected 12 Aug 1912 by H. C. Raven. Original number 157.

Type Locality: Telok Karang, Tigau (Teluk Karangtigau), [Kalimantan Timur], Dutch Southeast Borneo (Kalimantan), 

[Indonesia].

Epimys crassus Lyon

Proc. U.S. Natl. Mus. 40: 103, 25 Apr 1911.

= Sundamys muelleri (Jentink, 1879). See Musser and Newcomb 

(1983:401).

Holotype: USNM 145471. Skin and skull. Adult male. Collected 

8 May 1907 by W. L. Abbott. Original number 5190.

Type Locality: Pulo Lamukoton (Pulau Lemukutan), [Kalimantan 

Barat], off west coast of Borneo (Kalimantan), [Indonesia].

Epimys pollens Miller

Smithson. Misc. Collect. 61(21): 17, 29 Dec 1913d.

= Sundamys muelleri (Jentink, 1879). See Musser and Newcomb 

(1983:401).

Holotype: USNM 124691. Skin and skull. Adult male. Collected 

22 May 1904 by W. L. Abbott. Original number 3250.

Type Locality: Tanjong Rengsam, Banka (Pulau Bangka), [Kepulauan Bangka Belitung, Sumatra, Indonesia].

Epimys potens Miller

Smithson. Misc. Collect. 61(21): 17, 29 Dec 1913d.

= Sundamys muelleri (Jentink, 1879). See Musser and Newcomb 

(1983:401).

Holotype: USNM 114384. Skin and skull. Adult female. Collected 5 Feb 1902 by W. L. Abbott. Original number 1516.

Type Locality: [FC: Ujong Tambaga, east side of] Pulo (Pulau) 

Tuangku, Banjak Islands (Kepulauan Banyak), [Aceh, Sumatra, Indonesia].

Epimys sebucus Lyon

Proc. U.S. Natl. Mus. 40: 102, 25 Apr 1911.

= Sundamys muelleri (Jentink, 1879). See Musser and Newcomb 

(1983:401).

Holotype: USNM 151964. Skin and skull. Adult male. Collected 

2 Jan 1908 by W. L. Abbott. Original number 5717.

Type Locality: [FC: North end of] Pulo (Pulau) Sebuku, [Kalimantan Selatan], off southeastern coast of Borneo (Kalimantan), [Indonesia].

Epimys valens Miller

Smithson. Misc. Collect. 61(21): 17, 29 Dec 1913d.

= Sundamys muelleri (Jentink, 1879). See Musser and Newcomb 

(1983:401).

Holotype: USNM 114285. Skin and skull. Adult female. Collected 19 Jan 1902 by W. L. Abbott. Original number 1430.

Type Locality: Pulo (Pulau) Bangkaru, Banjak Islands (Kepulauan Banyak), [Aceh, Sumatra, Indonesia].

Epimys victor Miller

Smithson. Misc. Collect. 61(21): 16, 29 Dec 1913d.

= Sundamys muelleri (Jentink, 1879). See Musser and Newcomb 

(1983:401).

Holotype: USNM 115422. Skin and skull. Adult female. Collected 

25 May 1902 by W. L. Abbott. Original number 1706.

Type Locality: Near [FC: Palawan, about 35 mi (56 km) from] 

mouth of Rumpin River, Pahang, [Malaysia].

Mus chombolis Lyon

Proc. U.S. Natl. Mus. 36: 484, 1 Jun 1909.

= Sundamys muelleri (Jentink, 1879). See Musser and Newcomb 

(1983:401).

Holotype: USNM 144393. Skin and skull. Adult female. Collected 10 Mar 1907 by W. L. Abbott. Original number 5100.

Type Locality: Pulo Jombol (Pulau Combol), Rhio Linga Archipelago (Kepulauan Riau), [Kepulauan Riau, Sumatra, 

Indonesia].

Mus domitor Miller

Proc. U.S. Natl. Mus. 26: 461, 3 Feb 1903c.

= Sundamys muelleri (Jentink, 1879). See Musser and Newcomb 

(1983:401).
number 642 • 8 9

Holotype: USNM 114621. Skin and skull. Adult female. Collected 4 Mar 1902 by W. L. Abbott. Original number 1592.

Type Locality: Pulo Mansalar (Pulau Musala), at entrance to Tapanuli Bay, [Sumatera Utara], Sumatra, [Indonesia].

Mus firmus Miller

Proc. Acad. Nat. Sci. Phila. 54: 155, 11 Jun 1902a.

= Sundamys muelleri (Jentink, 1879). See Musser and Newcomb 

(1983:401).

Holotype: USNM 113038. Skin and skull. Adult female. Collected 25 Aug 1901 by W. L. Abbott. Original number 1215.

Type Locality: [FC: Mantuda Bay, W coast of] Linga Island 

(Pulau Lingga), [Kepulauan Lingga, Kepulauan Riau, Sumatra, Indonesia].

Mus integer Miller

Proc. Wash. Acad. Sci. 3: 119, 26 Mar 1901b.

= Sundamys muelleri (Jentink, 1879). See Musser and Newcomb 

(1983:401).

Holotype: USNM 104837. Skin and skull. Adult male. Collected 

7 Jun 1900 by [SL: W. L. Abbott. Original number 455].

Type Locality: Sirhassen Island (Pulau Serasan), South Natunas 

(Kepulauan Natuna Selatan), [Kepulauan Riau, Indonesia].

Mus validus Miller

Proc. Biol. Soc. Wash. 13: 141, 21 Apr 1900c.

= Sundamys muelleri (Jentink, 1879). See Musser and Newcomb 

(1983:401).

Holotype: USNM 86741. Skin and skull. Adult male. Collected 

18 Feb 1899 by [SL: W. L. Abbott]. No original number.

Type Locality: [SL: Khow Sai Dow (Khao Soi Dao)], altitude 

about 1,000 ft (305 m), mountains of Trong (Trang), Lower 

Siam (Thailand).

Rattus balmasus Lyon

Proc. U.S. Natl. Mus. 52: 447, 30 Dec 1916a.

= Sundamys muelleri (Jentink, 1879). See Musser and Newcomb 

(1983:401).

Holotype: USNM 121765. Skin and skull. Adult female. Collected 13 Feb 1903 by W. L. Abbott. Original number 

2286.

Type Locality: Tana Bala (Palau Tanabala), Batu Islands (Kepulauan Batu), [Sumatera Utara, Sumatra, Indonesia].

Rattus pinatus Lyon

Proc. U.S. Natl. Mus. 52: 448, 30 Dec 1916a.

= Sundamys muelleri (Jentink, 1879). See Musser and Newcomb 

(1983:401).

Holotype: USNM 121778. Skin and skull. Adult male. Collected 

4 Mar 1903 by W. L. Abbott. Original number 2367.

Type Locality: Pulo Pinie (Palau Pini), Batu Islands (Kepulauan 

Batu), [Sumatera Utara, Sumatra, Indonesia].

Rattus virtus Lyon

Proc. Biol. Soc. Wash. 29: 210, 22 Sep 1916b.

= Sundamys muelleri (Jentink, 1879). See Musser and Newcomb 

(1983:401).

Holotype: USNM 144223. Skin and skull. Adult male. Collected 

10 Dec 1906 by W. L. Abbott. Original number 4944.

Type Locality: Siak River (Sungai Siak), near mouth of Gasip 

River (Sungai Gasip), [Riau], eastern Sumatra, [Indonesia].

Genus Taeromys Sody, 1941

Rattus hamatus Miller and Hollister

Proc. Biol. Soc. Wash. 34: 97, 30 Jun 1921b.

= Taeromys hamatus (Miller and Hollister, 1921). See Musser 

and Newcomb (1983:487).

Holotype: USNM 218680. Skin and skull. Adult male. Collected 

16 Jan 1917 by H. C. Raven. Original number 3095.

Type Locality: [FC: Sungai Saepoi], Goenoeng Lehio, [FC: altitude between 6,166 and 7,155 ft (1,879 and 2,184 m)], 

[Sulawesi Tengah], Middle Celebes (Sulawesi), [Indonesia].

Rattus punicans Miller and Hollister

Proc. Biol. Soc. Wash. 34: 98, 30 Jun 1921b.

= Taeromys punicans (Miller and Hollister, 1921). See Musser 

and Newcomb (1983:487).

Holotype: USNM 219625. Skin and skull. Adult female. Collected 7 Feb 1918 by H. C. Raven. Original number 3501.

Type Locality: Pinedapa, [Sulawesi Tengah], Middle Celebes (Sulawesi), [Indonesia].

Genus Tarsomys Mearns, 1905

Tarsomys apoensis Mearns

Proc. U.S. Natl. Mus. 28: 453, 13 May 1905.

Holotype: USNM 125280. In alcohol, skull removed. Adult 

male. Collected 5 Jul 1904 by E. A. Mearns. Original number 5706.

Type Locality: Mount Apo, altitude 6,700 ft (2,057 m), southern 

Mindanao, Philippine Islands (Philippines).

Genus Taterillus Thomas, 1910

Taterillus arenarius Robbins

Proc. Biol. Soc. Wash. 87: 397, 31 Dec 1974.

Holotype: USNM 401919. Skin, skull, [and cleared and stained 

baculum]. Adult male. Collected 9 Apr 1967 by C. B. Robbins. Original number 799.

Type Locality: Tiguent, [SL and FC: 17°16′N, 16°01′W], Trarza 

Region, Mauritania.

Genus Thallomys Thomas, 1920

Thamnomys loringi Heller

Smithson. Misc. Collect. 52: 471, 13 Nov 1909a.

= Thallomys loringi (Heller, 1909). See Thomas (1920b:141).

Holotype: USNM 161904. Skin and skull. Adult female. Collected 17 Jul 1909 by J. A. Loring. Original number 6684.
9 0 • smithsonian contributions to zoology

Type Locality: Lake Naivasha, [Rift Valley], British East Africa 

(Kenya).

Genus Tokudaia Kuroda, 1943

Tokudamys osimensis muenninki Johnson

Proc. Biol. Soc. Wash. 59: 170, 23 Dec 1946.

= Tokudaia muenninki (Johnson, 1946). See Musser and Carleton (1993:670).

Holotype: USNM 278757. Skin and skull. Adult female. Collected 24 Sep 1945 by D. H. Johnson and O. A. Muennink. 

Original number DHJ 479. [SL: NAMRU-2 No. 686].

Type Locality: Hentona, western coast of northern Okinawa Island, Riu Kiu (Ryukyu) Islands, [Okinawa, Japan].

Genus Tryphomys Miller, 1910

Tryphomys adustus Miller

Proc. U.S. Natl. Mus. 38: 399, 19 Aug 1910b.

Holotype: USNM 151511. Skin and skull. Old female. Collected 

26 Jul 1907 by E. A. Mearns. Original number 6457.

Type Locality: Haights-in-the-Oaks, [FC: 7,000 ft (2,134 m)], 

Benguet [SL: Province], Luzon [FC: Island], [SL: Philippine 

Islands (Philippines)].

Genus Uranomys Dollman, 1909

Uranomys ugandae Heller

Smithson. Misc. Collect. 56(17): 12, 28 Feb 1911.

= Uranomys ruddi Dollman, 1909. See Misonne (1974:31).

Holotype: USNM 164822. [Skin and skull]. Adult male. Collected 2 Jan 1910 by J. A. Loring. Original number 8570.

Type Locality: Kikonda, [Kibale], Uganda.

Genus Zelotomys Osgood, 1910

Zelotomys hildegardeae vinaceus Heller

Smithson. Misc. Collect. 59(16): 10, 5 Jul 1912.

= Zelotomys hildegardeae (Thomas, 1902).

Holotype: USNM 181798. [Skin and skull]. Adult female. Collected 4 Nov 1911 by E. Heller. Original number 4578.

Type Locality: [SL: Entonei], Ndi, Mount Mbololo, altitude 

3,000 ft (914 m) [SL: 3,800 ft (1,158 m)], Taita Mountains 

(Hills), [Coast Province], British East Africa (Kenya).

Suborder Anomaluromorpha Bugge, 1974

Family Anomaluridae Gervais, 1849

Genus Idiurus, Matschie, 1894

Idiurus macrotis Miller

Proc. Biol. Soc. Wash. 12: 73, 24 Mar 1898c.

Holotype: USNM 83625. [Skin and skull]. Young adult male. 

Collected 18 Jun 1895 by G. L. Bates. Original number 4.

Type Locality: Efulen (Efoulan), Cameroon District, [Sud], West 

Africa (Cameroon).

Family Pedetidae Gray, 1825

Genus Pedetes Illiger, 1811

Pedetes cafer dentatus Miller

Proc. Biol. Soc. Wash. 40: 113, 26 Sep 1927a.

= Pedetes surdaster (Thomas, 1902). See Dieterlen (2005:1535).

Holotype: USNM 251879. Skin and skull. Adult male. Collected 

8 Jul 1926 by A. Loveridge. No original number.

Type Locality: Near Dodoma, Tanganyika Territory, Africa 

(Tanzania).

Pedetes surdaster currax Hollister

Smithson. Misc. Collect. 68(10): 3, 16 Jan 1918.

= Pedetes surdaster (Thomas, 1902).

Holotype: USNM 181762. Skin and skull. Adult male. Collected 

7 May 1911 by E. Heller. Original number 1802.

Type Locality: Kabalolot Hill, Sotik, [Rift Valley], British East 

Africa (Kenya).

Pedetes surdaster larvalis Hollister

Smithson. Misc. Collect. 68(10): 2, 16 Jan 1918.

= Pedetes surdaster (Thomas, 1902).

Holotype: USNM 163304. Skin and skull. Adult female. Collected 21 Nov 1909 by J. A. Loring. Original number 8250.

Type Locality: Ulukenia Hills, Athi Plains, British East Africa 

(Kenya).

Suborder Hystricomorpha Brandt, 1855

Family Bathyergidae Waterhouse, 1841

Genus Heliophobius Peters, 1846

Georychus kapiti Heller

Smithson. Misc. Collect. 52: 469, 24 Sep 1909b.

= Heliophobius argenteocinereus kapiti (Heller, 1909). See Hayman and Holt (1940:85).

Holotype: USNM 161708. [Skin and skull]. Adult female. Collected 3 May 1909 by J. A. Loring. Original number 6027.

Type Locality: Potha, Kapiti Plains, [Eastern Province], British 

East Africa (Kenya).

Family Hystricidae G. Fischer, 1817

Genus Atherurus F. Cuvier, 1829

Atherura zygomatica Miller

Smithson. Misc. Collect. 45: 42, 6 Nov 1903b.

= Atherurus macrourus (Linnaeus, 1758). See Chasen (1940:189). 

Holotype: USNM 112429. Skin and skull. Adult female. Collected 6 Jun 1909 by W. L. Abbott. Original number 1009.
number 642 • 9 1

Type Locality: Pulo Aor (Pulau Aur), off coast of Johore (Johor), 

[Malaysia].

Atherurus terutaus Lyon

Proc. U.S. Natl. Mus. 32: 587, 29 Jun 1907c.

= Atherurus macrourus (Linnaeus, 1758). See Chasen (1940:189). 

Holotype: USNM 123971. Skin and skull. Adult male. Collected 

10 Apr 1904 by W. L. Abbott. Original number 3223.

Type Locality: Pulo Terutau (Ko Tarutao), about 15 geographical miles (24 km) west of the Malay Peninsula, where the 

one hundredth meridian east of Greenwich cuts the west 

coast of the Malay Peninsula [Satun, Thailand].

Genus Hystrix Linnaeus, 1758

Thecurus major Schwarz

J. Mammal. 20: 246, 14 May 1939b.

= Hystrix crassipinus (Gunther, 1877). See Chasen (1940:188).

Holotype: USNM 196777. Skin and skull. Adult male. Collected 

8 Jul 1912 by H. C. Raven. Original number 118.

Type Locality: Near Samarinda, left bank of lower Mahakam 

River, [Kalimantan Timur], East Borneo (Kalimantan), 

[Indonesia].

Thecurus sumatrae Lyon

Proc. U.S. Natl. Mus. 32: 583, 29 Jun 1907c.

= Hystrix sumatrae (Lyon, 1907). See Van Weers (1978:25).

Holotype: USNM 143432. Skin and skull. Adult male. Collected 

17 Jan 1906 by W. L. Abbott. Original number 4637.

Type Locality: Aru Bay (Teluk Aru), [Sumatera Utara], east coast 

of Sumatra, [Indonesia].

Genus Trichys Gunther, 1877

Trichys macrotis Miller

Proc. U.S. Natl. Mus. 26: 469, 3 Feb 1903c.

= Trichys fasciculata (Shaw, 1801). See Van Weers (1976:19).

Holotype: USNM 144488. Skin and skull. Adult female. Collected 20 Feb 1902 by W. L. Abbott. Original number 1555.

Type Locality: Tapanuli Bay (Teluk Tapanuli), [Sumatera Utara], 

northwestern Sumatra, [Indonesia].

Family Thryonomyidae Pocock, 1922

Genus Thryonomys Fitzinger, 1867

Thryonomys gregorianus pusillus Heller

Smithson. Misc. Collect. 59(16): 17, 5 Jul 1912.

= Thryonomys gregorianus gregorianus (Thomas, 1894). See 

Misonne (1974:7).

Holotype: USNM 181807. [Skin and skull]. Adult female. Collected 3 Nov 1911 by E. Heller. Original number 4566, not 

4853 as given in original description.

Type Locality: Ndi, Mt. Mbololo, [Coast Province], British East 

Africa (Kenya).

Family Erethizontidae Bonaparte, 1845

Genus Coendou Lacepede, 1799

Coendou koopmani Handley and Pine

Mammalia 56: 238, 8 Jul 1992.

= Coendou nycthemera (Olfers, 1818). See Woods and Kilpatrick 

(2005:1547).

Holotype: USNM 519689. Skin and skull. Adult, sex unknown. 

Collected 31 Oct 1975 by R. Lainson. Original number [SL: 

M3089].

Type Locality: Belem, Para, Brazil.

Sphiggure spinosa F. Cuvier

Mem. Mus. Hist. Nat. 9: 433, Nov 1823.

= Coendou spinosus (Cuvier, 1823). See Voss (2011:15) for current use of Coendou and specific validity of spinosus.

Neotype: USNM 115122, as selected by Voss (2011:28). Skin 

and skull. Young adult male. Collected 14 Oct [SL: 1901] 

by W. Foster. Original number [SL: 536]. 

Type Locality: Sapucay (Sapucai), 25°40′S, 57°38′W, Paraguari, 

Paraguay.

Genus Erethizon F. Cuvier, 1823

Erethizon epixanthum doani H. H. Bailey

Bailey Mus. Libr. Nat. Hist. Bull. 12: 1, 15 Jan 1937.

= Erethizon dorsatum dorsatum (Linnaeus, 1758). See Hall 

(1981:851).

Holotype: USNM 347446. Skin and skull. Adult female. Collected 11 Apr 1935 by E. Doane. Original number [SL: 24]. 

Bailey Collection of Natural History number 718.

Type Locality: Piniware River, 15 mi (24 km) NW Red Bay, Labrador, [Newfoundland, Canada].

Remarks: Published as BCNH number 718; cataloged as USNM 

347446 on 4 Nov 1969.

Erethizon epixanthus couesi Mearns

Proc. U.S. Natl. Mus. 19: 723, 30 Jul 1897a.

= Erethizon dorsatum couesi Mearns, 1897. See Hall (1946: 

649).

Holotype: USNM 7814/6501. Skin and skull [with both femurs, 

left scapula, and humerus]. Nearly adult. No sex given. Collected on unknown date by E. Coues. Cataloged 19 Feb 

1865. No original number.

Type Locality: Fort Whipple, [Yavapai County], Arizona.

Erethizon epixanthus myops Merriam

Proc. Wash. Acad. Sci. 2: 27, 14 Mar 1900a.

= Erethizon dorsatum myops Merriam, 1900. See Anderson and 

Rand (1943:293).

Holotype: USNM 59140. [Skin and skull]. Old female. Collected 

Sep 1893 by C. H. Townsend. No original number.

Type Locality: Portage Bay, Alaska Peninsula, [Alaska].
9 2 • smithsonian contributions to zoology

Family Chinchillidae Bennett, 1833

Genus Lagostomus Brookes, 1828

Lagostomus maximus petilidens Hollister

Proc. Biol. Soc. Wash. 27: 58, 20 Mar 1914c.

Holotype: USNM 172801. Skull only. [Poole and Schantz (1942: 

256): male]. Collected 1910 by A. Hrdlicka. No original 

number.

Type Locality: [MC: Los Posos], 8 mi (13 km) north of Carmen 

de Patagones, [Province of Buenos Aires], Argentina.

Family Caviidae G. Fischer, 1817

Genus Hydrochoerus Brisson, 1762

Hydrochoerus hydrochaeris notialis Hollister

Proc. Biol. Soc. Wash. 27: 58, 20 Mar 1914c.

= Hydrochoerus hydrochaeris (Linnaeus, 1766).

Holotype: USNM 154186. Skin, skull, and skeleton, skeleton 

not found, [and brain in alcohol]. Adult male. [National 

Zoological Park records: received 7 Dec 1904 from J. N. 

Ruffin, U.S. Consul at Asuncion, died 6 Jan 1909]. National 

Zoological Park number 1555.

Type Locality: Paraguay.

Hydrochoerus isthmius Goldman

Smithson. Misc. Collect. 60(2): 11, 20 Sep 1912a.

Holotype: USNM 179703. [Skin and skull]. Adult male. Collected 

4 Apr 1912 by E. A. Goldman. Original number 21591.

Type Locality: Marraganti, near the head of tidewater on the Rio 

Tuyra, [SL: sea level], [Darien], eastern Panama.

Family Dasyproctidae Bonaparte, 1838

Genus Dasyprocta Illiger, 1811

Dasyprocta punctata bellula Kellogg

Proc. Biol. Soc. Wash. 59: 59, 11 Mar 1946.

Holotype: USNM 277152. Skin, skull, [and penis in alcohol]. 

Adult male. Collected 11 May 1944 by J. P. E. Morrison. 

Original number 29.

Type Locality: Isla San Jose, Archipelago de las Perlas, Golfo de 

Panama, [Panama], Panama.

Dasyprocta punctata chiapensis Goldman

Smithson. Misc. Collect. 60(22): 13, 28 Feb 1913.

Holotype: USNM 77997. [Skin and skull]. Adult female. Collected 26 Feb 1896 by E. W. Nelson and E. A. Goldman. 

Original number 9430.

Type Locality: Huehuetan, [SL: altitude 300 ft (91 m)], southern 

Chiapas, Mexico.

Dasyprocta punctata dariensis Goldman

Smithson. Misc. Collect. 60(22): 11, 28 Feb 1913.

Holotype: USNM 179056. [Skin and skull]. Adult female. Collected 

24 Apr 1912 by E. A. Goldman. Original number 21637.

Type Locality: Near head of Rio Limon, altitude 5,200 ft (1,585 m), 

Mount Pirri, [Darien], eastern Panama.

Dasyprocta punctata richmondi Goldman

Proc. Biol. Soc. Wash. 30: 114, 23 May 1917.

Holotype: USNM 51333. [Skin and skull]. Adult female. Collected 

16 Nov 1892 by C. W. Richmond. Original number 156.

Type Locality: Escondido River, 50 mi (80 km) above Bluefields, 

[Atlantico Sur], Nicaragua.

Dasyprocta punctata underwoodi Goldman

J. Wash. Acad. Sci. 21: 481, 19 Nov 1931.

Holotype: USNM 256459. [Skin and skull]. Adult male. Collected 

1 Jun 1931 by C. F. Underwood. Original number [MC: 478].

Type Locality: San Geronimo (San Jeronimo), Pirris, [Puntarenas], western Costa Rica.

Dasyprocta punctata yucatanica Goldman

Smithson. Misc. Collect. 60(22): 12, 28 Feb 1913.

Holotype: USNM 108293. [Skin, skull, and baculum]. Adult 

male. Collected 22 Dec 1900 by E. W. Nelson and E. A. 

Goldman. Original number 14347.

Type Locality: Apazote, near Yohaltun, Campeche, Mexico.

Family Cuniculidae Miller And Gidley, 1918

Genus Cuniculus Brisson, 1762

Agouti paca nelsoni Goldman

Smithson. Misc. Collect. 60(22): 9, 28 Feb 1913.

= Cuniculus paca nelsoni (Goldman, 1913). See Hollister (1913e:79).

Holotype: USNM 65952. [Skin and skull]. Adult male. Collected 

30 Apr 1894 by E. W. Nelson and E. A. Goldman. Original 

number 6135.

Type Locality: Catemaco, [FC: altitude 1,000 ft (305 m)], southern Veracruz, Mexico.

Family Ctenomyidae Lesson, 1842

Genus Ctenomys Blainville, 1826

Ctenomys robustus J. A. Allen

Bull. Am. Mus. Nat. Hist. 19: 185, 9 May 1903.

Primary junior homonym of Ctenomys robustus Phillippi, 1896; 

replaced with Ctenomys osgoodi by J. A. Allen (1905:191).

= Ctenomys magellanicus osgoodi J. A. Allen, 1905. See Osgood 

(1943:120). 

Holotype: USNM 84149. [Skin and skull]. Adult male. Collected 

20 Feb 1897 by O. A. Peterson. Original number [SL: 485].

Type Locality: Rio Chico de Santa Cruz, near the Cordilleras, 

[Santa Cruz, Argentina].

Remarks: According to Lyon and Osgood (1909:42), “Type designated as No. 84194, an error for 84149. No. 84194 is a 
number 642 • 9 3

specimen of C. sericeus. The measurements of No. 84149 

agree with those given by J. A. Allen as belonging to the 

Type, and the specimen itself bears Allen’s type label.”

Ctenomys sericeus J. A. Allen

Bull. Am. Mus. Nat. Hist. 19: 187, 9 May 1903.

Holotype: USNM 84189. [Skin and skull]. Adult male. Collected 

5 Feb 1897 by O. A. Peterson. Original number [SL: 331].

Type Locality: Cordilleras, upper Rio Chico de Santa Cruz, 

[Santa Cruz], Patagonia (Argentina).

Remarks: According to Lyon and Osgood (1909:42), “Type designated as No. 84191, which is an error for No. 84189. 

The measurements of the skin and skull ascribed by Allen 

to the type are those of 84189 and do not agree with those 

of 84191, a smaller individual. To indicate further that No. 

84189 is the type, Allen tied to the specimen the standard 

red type label of the American Museum of Natural History, 

with the italicized words crossed out and the words U. S. 

Nat. added as well as the name Ctenomys sericeus Allen.”

Family Echimyidae Gray, 1825

Genus Diplomys Thomas, 1916

Isothrix darlingi Goldman

Smithson. Misc. Collect. 60(2): 12, 20 Sep 1912a.

= Diplomys labilis (Bangs, 1901). See Handley (1966:787).

Holotype: USNM 179577. [Skin and skull]. Young female. Collected 

11 May 1912 by E. A. Goldman. Original number 21739.

Type Locality: Marraganti, [SL: sea level], near Real de Santa 

Maria, on the Rio Tuyra, [Darien], eastern Panama.

Genus Hoplomys J. A. Allen, 1908

Hoplomys geothalsi Goldman

Smithson. Misc. Collect. 56(36): 10, 19 Feb 1912b.

= Hoplomys gymnurus (Thomas, 1897). See Handley (1959:8).

Holotype: USNM 170972. [Skin and skull]. Young female. Collected 16 Feb 1911 by E. A. Goldman. Original number 20888.

Type Locality: Rio Indio, near Gatun, Canal Zone (Colon), Panama.

Hoplomys gymnurus wetmorei Handley

Smithson. Misc. Collect. 139(4): 9, 3 Jul 1959.

= Hoplomys gymnurus (Thomas, 1897).

Holotype: USNM 307057. Skin and skull. Adult male. Collected 

1 Mar 1958 by A. Wetmore. Original number 1479.

Type Locality: Escudo de Veraguas, Prov. Bocas del Toro, Panama.

Remarks: Hoplomys gymnurus was considered monotypic by 

Woods and Kilpatrick (2005:1583), who overlooked the 

name wetmorei.

Genus Pattonomys Emmons, 2005

Loncheres flavidus Hollister

Proc. Biol. Soc. Wash. 27: 143, 10 Jul 1914d.

= Pattonomys flavidus (Hollister, 1914). See Emmons (2005:282).

Holotype: USNM 63218. Skin and skull. Adult male. Collected 

9 Jul 1895 by W. Robinson. Original number 479.

Type Locality: El Valle, Margarita Island (Isla de Margarita), 

[Nueva Esparta], Venezuela.

Genus Proechimys J. A. Allen, 1899

Proechimys guairae Thomas

Proc. Biol. Soc. Wash. 14: 27, 2 Apr 1901.

Holotype: USNM 102731. Skin and skull. [FC: Adult] male. 

Collected 8 Jul 1900 by Messrs. [M. W.] Lyon [Jr.] and 

[W.] Robinson. Original number 81.

Type Locality: La Guaira, [Vargas], Venezuela.

Proechimys guyannensis magdalenae Hershkovitz

Proc. U.S. Natl. Mus. 97: 136, 6 Jan 1948b.

= Proechimys magdalenae Hershkovitz, 1948. See Emmons and 

Feer (1990:274).

Holotype: USNM 280170. Skin and skull. Adult male. Collected 

20 Jun 1943 by P. Hershkovitz. Original number 2098.

Type Locality: Rio San Pedro, a small stream in the northern 

foothills of the Cordillera Central, above the village of 

Norosi, altitude 178 m, Bolivar, Colombia.

Proechimys rubellus Hollister

Proc. Biol. Soc. Wash. 27: 57, 20 Mar 1914c.

= Proechimys semispinosus rubellus Hollister, 1914. See Goldman (1920:120).

Holotype: USNM 12901/38587. Skin and skull. Adult “[male].” 

Collected [on unknown date] by J. C. Zeledon. Skin cataloged 23 Jun 1878. Original number [MC: 112].

Type Locality: [MC: Pacuare], Angostura Valley, Costa Rica.

Proechimys semispinosus ignotus Kellogg

Proc. Biol. Soc. Wash. 59: 61, 11 Mar 1946.

Holotype: USNM 277160. Skin and skull. Adult male. Collected 

8 May 1944 by J. P. E. Morrison. Original number 28.

Type Locality: Isla San Jose, Archipelago de las Perlas, Golfo de 

Panama, [Panama], Panama.

Proechimys steerei Goldman

Proc. Biol. Soc. Wash. 24: 238, 28 Nov 1911.

Holotype: USNM 105535. [Skin and skull]. Adult male. Collected [FC: 15 Feb] 1901 by J. B. Steere. Original number 

[MC: 77].

Type Locality: [FC: Hyutanahan], Rio Purus, a southern tributary of the Amazon, [Amazonas], in northwestern Brazil.

Family Myocastoridae Ameghino, 1904

Genus Myocastor Kerr, 1792

Myocastor coypus santacruzae Hollister

Proc. Biol. Soc. Wash. 27: 57, 20 Mar 1914c.
9 4 • smithsonian contributions to zoology

Holotype: USNM 96513. Skin and skull. Adult “[male].” Collected [Poole and Schantz (1942:490): Jul to Sep 1896] by 

W. Frakes. Original number not given. [SL: 1977-X].

Type Locality: Near Los Palmares, north bank of the Rio Salado, Santa Fe, not Santa Cruz as in original description, 

Argentina.

Family Capromyidae Smith, 1842

Genus Geocapromys Chapman, 1901

Capromys brachyurus thoracatus True

Proc. U.S. Natl. Mus. 11: 469, 3 Sep 1889.

= Geocapromys thoracatus (True, 1889). See Morgan (1985:29).

Lectotype: USNM 15897/22691, as designated by Lyon and Osgood (1909:41). Skin, skull, [and femurs]. [Lyon and Osgood (1909:41): Adult] [SL: male]. Collected [SL: 6 Mar 

1887 by C. H. Townsend]. No original number.

Type Locality: Little Swan Island, [SL: Caribbean Sea], at the 

entrance of the Gulf of Honduras, [Swan Islands].

Remarks: Type not designated by number in the original description, but according to Lyon and Osgood (1909:41), “of the 

two original specimens Dr. True has personally selected the 

above as the type and placed a red label upon it.” This statement by Lyon and Osgood effectively designated the lectotype (ICZN, 1999: Article 74.5).

Genus Mysateles Lesson, 1842

Capromys prehensilis gundlachi Chapman

Bull. Am. Mus. Nat. Hist. 14: 317, 12 Nov 1901.

= Mysateles prehensilis gundlachi (Chapman, 1901). See Kratochvil et al. (1978:17).

Holotype: USNM 103905. Skin and skull. [Lyon and Osgood 

(1909:41): Adult] male. Collected 4 Jul 1900 by [Lyon 

and Osgood (1909:41): W.] Palmer and [Lyon and Osgood 

(1909:41): J. H.] Riley. Original number 505.

Type Locality: Nueva Gerona, Isle of Pines, [Isla de la Juventud], 

[SL: Cuba].

Genus Plagiodontia F. Cuvier, 1836

Plagiodontia hylaeum Miller

Proc. U.S. Natl. Mus. 72(16): 4, 30 Sep 1927b.

= Plagiodontia aedium hylaeum Miller, 1927. See Anderson 

(1965:97).

Holotype: USNM 239887. Skin, skull, and skeleton. Young 

adult male. Collected 23 Nov 1923 by W. L. Abbott. Original number [SL: B].

Type Locality: Guarabo, 10 mi (16 km) east of Jovero, Samana 

Province, Dominican Republic.
References

Abiadh, A., M. Chetoui, T. Lamine-Cheniti, E. Capanna, and P. Colangelo. 2010. Molecular Phylogenetics of the 

Genus Gerbillus (Rodentia, Gerbillinae): Implications for Systematics, Taxonomy, and Chromosomal Evolution. Molecular Phylogenetics and Evolution, 56:513–518. http://dx.doi.org/10.1016/j.ympev.2010.04.018.

Alexander, L. F. 1999. “Western Red-Backed Vole, Clethrionomys californicus.” In The Smithsonian Book of 

North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 612–613. Washington, D.C.: Smithsonian 

Institution Press.

Allen, G. M. 1926. Rats (Genus Rattus) from the Asiatic Expeditions. American Museum Novitates, 217:1–16.

———. 1931. Type Specimens of Mammals in the Museum of Comparative Zoology. Bulletin of the Museum of 

Comparative Zoology, 71:229–289.

———. 1939. A Checklist of African Mammals. Bulletin of the Museum of Comparative Zoology at Harvard 

College, 83:1–763.

———. 1940. “The Mammals of China and Mongolia.” In Natural History of Central Asia, Volume 11, pt. 2, ed. 

W. Granger, pp. 621–1350. New York: American Museum of Natural History.

Allen, G. M., and B. L. Lawrence. 1936. Scientific Results of an Expedition to Rainforest Regions in Eastern 

Africa. III. Mammals. Bulletin of the Museum of Comparative Zoology at Harvard College, 79:31–126.

Allen, J. A. 1891. Descriptions of Two Supposed New Species of Mice from Costa Rica and Mexico, with Remarks 

on Hesperomys melanophrys of Coues. Proceedings of the United States National Museum, 14:193–196. 

http://dx.doi.org/10.5479/si.00963801.850.193.

———. 1893. On a Collection of Mammals from the San Pedro Martir Region of Lower California, with Notes 

on Other Species, Particularly of the Genus Sitomys. Bulletin of the American Museum of Natural History, 

5:188–202.

———. 1894. Descriptions of Ten New North American Mammals, and Remarks on Others. Bulletin of the 

American Museum of Natural History, 6:317–332.

———. 1895a. On a Collection of Mammals from Arizona and Mexico, Made by Mr. W. W. Price, with Field 

Notes by the Collector. Bulletin of the American Museum of Natural History, 7:193–258.

———. 1895b. On the Species of the Genus Reithrodontomys. Bulletin of the American Museum of Natural History, 7:107–143.

———. 1897. Further Notes on Mammals Collected in Mexico by Dr. Audley C. Buller, with Descriptions of New 

Species. Bulletin of the American Museum of Natural History, 9:47–58.

———. 1903. Descriptions of New Rodents from Southern Patagonia, with a Note on the Genus Euneomys

Coues, and an Addendum to Article IV, on Siberian Mammals. Bulletin of the American Museum of Natural 

History, 19:185–196.

———. 1905. “Mammalia of Southern Patagonia.” In Reports of the Princeton University Expedition to Patagonia, 1896–1899, pt. 1, ed. W. B. Scott III, pp. 1–210. Princeton, N.J.: Princeton University Press.

———. 1911. Mammals Collected in the Dutch East Indies by Mr. Roy C. Andrews on the Cruise of the ‘Albatross’ in 1909. Bulletin of the American Museum of Natural History, 30:335–339.

Alvarez, T. 1961. Taxonomic Status of Some Mice of the Peromyscus boylii Group in Eastern Mexico, with Description of a New Subspecies. University of Kansas Publications, Museum of Natural History, 14:111–120.

Alvarez-Castaneda, S. T., and P. Cortes-Calva. 2003. Peromyscus eva. Mammalian Species, 738:1–3. http://dx.doi

.org/10.1644/738.

Alvarez-Castaneda, S. T., and N. Gonzalez-Ruiz. 2009. Peromyscus levipes (Rodentia: Cricetidae). Mammalian 

Species, 824:1–6. http://dx.doi.org/10.1644/824.1.

Anderson, R. M. 1937. “Mammals and Birds of the Western Arctic District, Northwest Territories, Canada.” In 

Canada’s Western Northland, pp. 97–122. Ottawa, Can.:J. O. Patenaude, I. S. O., King’s Printer.

———. 1942. Canadian Voles of the Genus Phenacomys with Description of Two New Canadian Subspecies. 

Canadian Field-Naturalist, 56:56–60.
9 6 • smithsonian contributions to zoology

———. 1943. A Prior Name Revived for the Bean Mouse. Canadian Field-Naturalist, 57:92.

———.1947 [1946]. Catalog of Canadian Recent Mammals. Bulletin of the National Museum of Canada, 102:1–238.

Anderson, R. M., and A. L. Rand. 1943. Variation in the Porcupine (Genus Erethizon) in Canada. Canadian Journal of Research, Section D, 21:292–309. 

http://dx.doi.org/10.1139/cjr43d-024.

Anderson, S. 1954. Subspeciation in the Meadow Mouse, Microtus montanus, 

in Wyoming and Colorado. University of Kansas Publications, Museum of 

Natural History, 7:489–506.

———. 1965. Conspecificity of Plagiodontia aedium and P. hylaeum (Rodentia). 

Proceedings of the Biological Society of Washington, 78:95–98.

———. 1972. Mammals of Chihuahua, Taxonomy and Distribution. Bulletin of 

the American Museum of Natural History, 148:151–410.

Armstrong, D. M. 1999a. “Canyon Mouse, Peromyscus crinitus.” In The Smithsonian Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 

566–567. Washington, D.C.: Smithsonian Institution Press.

———. 1999b. “Mesquite Mouse, Peromyscus merriami.” In The Smithsonian 

Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 577–

578. Washington, D.C.: Smithsonian Institution Press.

Audubon, J. J., and J. Bachman. 1841. Descriptions of New Species of Quadrupeds 

Inhabiting North America. Proceedings of the Academy of Natural Sciences 

of Philadelphia, 1:91–103.

Bailey, H. H. 1937. Two New North American Mammals. The Bailey Museum and 

Library of Natural History Bulletin, 12:1–4.

Bailey, V. 1897. Revision of the American Voles of the Genus Evotomys. Proceedings of the Biological Society of Washington, 11:113–138.

———. 1898a. Description of Eleven New Species and Subspecies of Voles. Proceedings of the Biological Society of Washington, 12:85–90.

———. 1898b. A New Name for Microtus insularis Bailey. Science, n.s., 8:782–

783. http://dx.doi.org/10.1126/science.8.205.782-a.

———. 1898c. A New Species of Evotomys from British Columbia. Proceedings of 

the Biological Society of Washington, 12:21–22.

———. 1900. Revision of the American Voles of the Genus Microtus. North American Fauna, 17:1–88.

———. 1902a. Seven New Mammals from Western Texas. Proceedings of the Biological Society of Washington, 15:117–120.

———. 1902b. Synopsis of the North American Species of Sigmodon. Proceedings 

of the Biological Society of Washington, 15:101–116.

———. 1905. Biological Survey of Texas. North American Fauna, 25:1–222.

———. 1906. A New White-Footed Mouse from Texas. Proceedings of the Biological Society of Washington, 19:57–58.

———. 1913. Ten New Mammals from New Mexico. Proceedings of the Biological Society of Washington, 26:129–134.

———. 1917. A New Subspecies of Meadow Mouse from Wyoming. Proceedings 

of the Biological Society of Washington, 30:29–30.

———. 1920. Identity of the Bean Mouse of Lewis and Clark. Journal of Mammalogy, 1:70–72.

———. 1936. The Mammals and Life Zones of Oregon. North American Fauna, 

55:1–416.

Baird, S. F. 1855a. Characteristics of Some New Species of Mammalia, Collected by 

the U. S. and Mexican Boundary Survey, Major W. H. Emory, U. S. A. Commissioner. Proceedings of the Academy of Natural Sciences of Philadelphia, 

7:331–333.

———. 1855b. Characteristics of Some New Species of North American Mammalia, Collected Chiefly in Connection with the U. S. Surveys of a Railroad 

Route to the Pacific. Proceedings of the Academy of Natural Sciences of 

Philadelphia, 7:333–336.

———. 1857. “Mammals.” In Reports of Explorations and Surveys to Ascertain 

the Most Practicable and Economical Route for a Railroad from the Mississippi River to the Pacific Ocean. Part 1. General Report upon the Zoology 

of the Several Pacific Railroad Routes, Volume 8, bk. 1.Washington, D.C.: 

Beverly Tucker.

Baker, R. H. 1952. Geographic Range of Peromyscus melanophrys, with Descriptions of New Subspecies. University of Kansas Publications, Museum of 

Natural History, 5:251–258.

———. 1969. Cotton Rats of the Sigmodon fulviventer Group. Miscellaneous Publications, Museum of Natural History, University of Kansas, 51:177–232.

Baker, R. H., and J. K. Greer. 1962. Mammals of the Mexican State of Durango. 

Publications of the Museum, Michigan State University, Biological Series,

2:25–154.

Bangs, O. 1898. The Land Mammals of Peninsular Florida and the Coast Region of Georgia. Proceedings of the Boston Society of Natural History, 

28:157–285.

Barbour, R. W. 1956. Synaptomys cooperi in Kentucky, with Description of a 

New Subspecies. Journal of Mammalogy, 37:413–416. http://dx.doi.org/10

.2307/1376742.

Batchelder, C. F. 1896. Some Facts in Regard to the Distribution of Certain Mammals in New England and Northern New York. Proceedings of the Boston 

Society of Natural History, 27:185–193.

Batzli, G. O. 1999a. “Brown Lemming, Lemmus sibiricus.” In The Smithsonian 

Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 653–

654. Washington, D.C.: Smithsonian Institution Press.

———. 1999b. “Northern Red-Backed Vole, Clethrionomys rutilus.” In The Smithsonian Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 

616–617. Washington, D.C.: Smithsonian Institution Press.

Bee, J. W., and E. R. Hall. 1956. Mammals of Northern Alaska. Miscellaneous 

Publications, Museum of Natural History, University of Kansas, 8:1–309.

Benson, S. B. 1935. The Status of Reithrodontomys montanus (Baird). Journal of 

Mammalogy, 16:139–142. http://dx.doi.org/10.2307/1374363. 

Birkenholz, D. E. 1999. “Round-Tailed Muskrat, Neofiber alleni.” In The Smithsonian Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 

650–651. Washington, D.C.: Smithsonian Institution Press.

Bole, B. P., Jr., and P. N. Moulthrop. 1942. The Ohio Recent Mammal Collection 

in the Cleveland Museum of Natural History. Scientific Publications of the 

Cleveland Museum of Natural History, 5:84–181.

Booth, E. S. 1945. A New Red-Backed Vole from Washington State. Murrelet, 

26:27–28.

Bowen, W. W. 1968. Variation and Evolution of Gulf Coast Populations of Beach 

Mice, Peromyscus polionotus. Bulletin of the Florida State Museum, 12:1–91.

Braun, J. K. 1999. “Southern Plains Woodrat, Neotoma micropus.” In The Smithsonian Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 

609–610. Washington, D.C.: Smithsonian Institution Press.

Burt, W. H. 1932. Descriptions of Heretofore Unknown Mammals from Islands 

in the Gulf of California, Mexico. Transactions of the San Diego Society of 

Natural History, 7:161–182.

———. 1938. Faunal Relationships on Geographic Distributions of Mammals in 

Sonora, Mexico. Miscellaneous Publications, Museum of Zoology, University of Michigan, 39:1–77.

Burt, W. H., and F. S. Barkalow Jr. 1942. A Comparative Study of the Bacula of 

Wood Rats (Subfamily Neotominae). Journal of Mammalogy, 42:287–297. 

http://dx.doi.org/10.2307/1374996.

Caire, W. 1999. “Canyon Mouse, Peromyscus eremicus.” In The Smithsonian Book 

of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 567–568. 

Washington, D.C.: Smithsonian Institution Press.

Cameron, G. N. 1999a. “Northern Pigmy Mouse, Baiomys taylori.” In The Smithsonian Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 

586–587. Washington, D.C.: Smithsonian Institution Press.

———. 1999b. “Fulvous Harvest Mouse, Reithrodontomys fulvescens.” In The 

Smithsonian Book of North American Mammals, ed. D. E. Wilson and 

S. Ruff, pp. 556–557. Washington, D.C.: Smithsonian Institution Press.

———. 1999c. “Hispid Cotton Rat, Sigmodon hispidus.” In The Smithsonian 

Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 593–

595. Washington, D.C.: Smithsonian Institution Press.

Carey, A. B. 1999. “Red Tree Vole, Arborimus longicaudus.” In The Smithsonian Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 

620–622. Washington, D.C.: Smithsonian Institution Press.

Carleton, M. D. 1977. Interrelationships of Populations of the Peromyscus boylii

Species Group (Rodentia: Muridae) in Western Mexico. Occasional Papers of 

the Museum of Zoology, University of Michigan, 675:1–47.

———. 1979. Taxonomic Status and Relationships of Peromyscus boylii from El 

Salvador. Journal of Mammalogy, 60:280–296. http://dx.doi.org/10.2307

/1379799.

———. 1980. Phylogenetic Relationships in Neotomine–Peromyscine Rodents 

(Muroidea) and a Reappraisal of the Dichotomy within New World Cricetinae. Miscellaneous Publications, Museum of Zoology, University of Michigan, 157:1–146.

———. 1989. “Systematics and Evolution.” In Advances in the Study of Peromyscus (Rodentia), ed. G. L. Kirkland Jr. and J. N. Layne, pp. 7–141. Lubbock: 

Texas Tech University Press.

Carleton, M. D., and J. Arroyo-Cabrales. 2009. Review of the Oryzomys couesi

complex (Rodentia: Cricetidae: Sigmodontinae) in Western Mexico. Bulletin 
number 642 • 9 7

of the American Museum of Natural History, 331:94–127. http://dx.doi.org

/10.1206/582-3.1.

Carleton, M. D., R. D. Fisher, and A. L. Gardner. 1999. Identification and Distribution of Cotton Rats, Genus Sigmodon (Muridae: Sigmodontinae), of Nayarit, 

Mexico. Proceedings of the Biological Society of Washington, 112:813–856.

Carleton, M. D., and G. G. Musser. 1989. Systematic Studies of Oryzomyine Rodents (Muridae, Sigmodontinae): A Synopsis of Microryzomys. Bulletin of 

the American Museum of Natural History, 191:1–83.

———. 2005. “Order Rodentia.” In Mammal Species of the World: A Taxonomic 

and Geographic Reference, Volume 2, 3rd ed., ed. D. E. Wilson and D. M. 

Reeder, pp. 745–752. Baltimore, Md.: Johns Hopkins University Press.

Carleton, M. D., J. C. K. Peterhans, and W. T. Stanley. 2006. Review of the Hylomyscus denniae Group (Rodentia: Muridae) in Eastern Africa, with Comments on the Generic Allocation of Epimys endorobae Heller. Proceedings of 

the Biological Society of Washington, 119:293–325.

Carleton, M. D., and C. B. Robbins. 1985. On the Status and Affinities of Hybomys 

planifrons (Miller, 1900) (Rodentia: Muridae). Proceedings of the Biological 

Society of Washington, 98:956–1003.

Carleton, M. D., and W. T. Stanley. 2012. Species Limits within the Praomys delectorum Group (Rodentia: Muridae: Murinae) of East Africa: A Morphometric Reassessment and Biogeographical Implications. Zoological Journal 

of the Linnean Society, 165:420–469. http://dx.doi.org/10.1111/j.1096-3642

.2012.00817.x.

Cary, M. 1903. A New Reithrodontomys from Western Nebraska. Proceedings of 

the Biological Society of Washington, 16:53–54.

———. 1911. A Biological Survey of Colorado. North American Fauna, 33:1–256.

Castiglia, R., and N. Oguge. 2008. Karyotype Divergence between Two Subspecies of the Typical Striped Grass Mouse Lemniscomys striatus (Linnaeus, 

1758) (Rodentia, Muridae). Mammalia, 72:57–60. http://dx.doi.org/10.1515

/MAMM.2008.001.

Castro-Campillo, A. H., H. R. Roberts, D. J. Schmidly, and R. D. Bradley. 1999. 

Systematic Status of Peromyscus boylii ambiguus Based on Morphologic 

and Molecular Data. Journal of Mammalogy, 80:1214–1231. http://dx.doi

.org/10.2307/1383172.

Chapman, F. M. 1901. A Revision of the Genus Capromys. Bulletin of the American Museum of Natural History, 14:313–323.

Chasen, F. N. 1940. A Handlist of Malaysian Mammals: A Systematic List of the 

Mammals of the Malay Peninsula, Sumatra, Borneo, and Java, Including the 

Adjacent Small Islands. Bulletin of the Raffles Museum, 15:1–209.

Cockrum, E. L. 1977. Status of the Hairy Footed Gerbil, Gerbillus latastei

Thomas and Trouessart. Mammalia, 41:75–80. http://dx.doi.org/10.1515

/mamm.1977.41.1.75.

Cockrum, E. L., and K. L. Fitch. 1952. Geographic Variation in Red-Backed Mice 

(Genus Clethrionomys) of the Southern Rocky Mountain Region. University 

of Kansas Publications, Museum of Natural History, 5:281–292.

Cockrum, E. L., T. C. Vaughan, and P. J. Vaughan. 1976. A Review of North African Short-Tailed Gerbils (Dipodillus) with Description of a New Taxon from 

Tunisia. Mammalia, 40:313–326. http://dx.doi.org/10.1515/mamm.1976.40

.2.313.

———. 1977. Gerbillus andersoni de Winton, a Species New to Tunisia. Mammalia, 40:467–473. http://dx.doi.org/10.1515/mamm.1976.40.3.467.

Coetzee, C. G. 1977. “Genus Steatomys.” In The Mammals of Africa: An Identification Manual, ed. J. Meester and H. W. Setzer, pt. 6.8, pp. 1–4. Washington, 

D.C.: Smithsonian Institution Press.

Cole, F. R., and D. E. Wilson. 2010. Microtus miurus (Rodentia: Cricetidae). Mammalian Species, 42 (855):75–89. http://dx.doi.org/10.1644/855.1.

Conroy, J. B. 1999. “Tundra Vole, Microtus oeconomus.” In The Smithsonian 

Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 638–

639. Washington, D.C.: Smithsonian Institution Press.

Cook, J. A., and D. R. Klein. 1999. “Insular Vole, Microtus abbreviatus.” In The 

Smithsonian Book of North American Mammals, ed. D. E. Wilson and 

S. Ruff, pp. 623–624. Washington, D.C.: Smithsonian Institution Press.

Cooper, J. G. 1868. The Fauna of Montana Territory. American Naturalist, 2:528–

538. http://dx.doi.org/10.1086/270303.

Cope, E. D. 1868. Untitled. Proceedings of the Academy of Natural Sciences of 

Philadelphia, 20:2.

Corbet, G. B. 1978. The Mammals of the Palaearctic Region: A Taxonomic Review.

London: British Museum (Natural History).

Cornely, J. E. 1999a. “Townsend’s Vole, Microtus townsendii.” In The Smithsonian Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 

646–647. Washington, D.C.: Smithsonian Institution Press.

———. 1999b. “Mexican Woodrat, Neotoma mexicana.” In The Smithsonian 

Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 608–

609. Washington, D.C.: Smithsonian Institution Press.

Coues, E. 1874. Synopsis of the Muridae of North America. Proceedings of the 

Academy of Natural Sciences of Philadelphia, 26:173–196.

———. 1877. “Muridae.” In Monographs of North American Rodentia, United 

States Geological Survey of the Territories, ed. E. Coues and J. A. Allen, pp. 

1–264. Washington, D.C.: Government Printing Office.

Cowan, I. McT., and C. J. Guiguet. 1956. The Mammals of British Columbia. British Columbia Provincial Museum Handbook, 11:1–413.

Cranford, J. A. 1999. “Dusky-Footed Woodrat, Neotoma fuscipes.” In The Smithsonian Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 

603–604. Washington, D.C.: Smithsonian Institution Press.

Cuvier, F. 1823 [1822].. Examen des especies du genre porc-epic, et formation des 

genres du sous-genres Acanthion, Erethizon, Sinethere, et Sphiggure. Memoires du Museum d’Historie Naturelle, 9:413–437.

Dalquest, W. W. 1948. Mammals of Washington. University of Kansas Publications, Museum of Natural History, 2:1–144.

Davis, R., and P. J. Young. 1999. “Arizona Cotton Rat, Sigmodon arizonae.” In 

The Smithsonian Book of North American Mammals, ed. D. E. Wilson and 

S. Ruff, pp. 591–592. Washington, D.C.: Smithsonian Institution Press.

Davis, W. B. 1939. The Recent Mammals of Idaho. Caldwell, Idaho: Caxton 

Printers.

———. 1944a. Geographic Variation in Brown Lemmings (Genus Lemmus). Murrelet, 25:19–25. http://dx.doi.org/10.2307/3533754.

———. 1944b. Notes on Mexican Mammals. Journal of Mammalogy, 25:370–

403. http://dx.doi.org/10.2307/1374900. 

Dieterlen, F. 1971. Beitrage sur Systematik, Okologie, und Biologie der Gattung 

Dendromus (Dendromurinae, Cricetidae, Rodentia), insbesondere ihrer 

zentralafrikanischen Formen. Saugetierkundliche Mitterlungen, 19:97–132.

———. 1991. Lemniscomys hoogstraali, an New Murid Species from Sudan. Bonner Zoologische Beitrage, 42:11–15.

———. 2005. “Family Pedetidae.” In Mammal Species of the World: A Taxonomic 

and Geographic Reference, Volume 2, 3rd ed., ed. D. E. Wilson and D. M. 

Reeder, p. 1535. Baltimore, Md.: Johns Hopkins University Press.

Dowler, R. C. 1999. “Plains Harvest Mouse, Reithrodontomys montanus.” In 

The Smithsonian Book of North American Mammals, ed. D. E. Wilson and 

S. Ruff, pp. 560–561. Washington, D.C.: Smithsonian Institution Press.

Durrant, S. D. 1952. Mammals of Utah, Taxonomy and Distribution. University of 

Kansas Publications, Museum of Natural History, 6:1–549.

Ellerman, J. R. 1949. The Families and Genera of Living Rodents. Volume 3, pt. 1. 

London: British Museum (Natural History). 

———. 1951. “Order Rodentia.” In Checklist of Palaearctic and Indian Mammals, 

1758 to 1946, ed. J. R. Ellerman and T. C. S. Morrison-Scott, pp. 456–712. 

London: Trustees of the British Museum.

Emmons, L. H. 1993. A New Genus and Species of Rat from Borneo (Rodentia: 

Muridae). Proceedings of the Biological Society of Washington, 106:752–

761.

———. 2005. “A Revision of the Genera of Arboreal Echimyidae (Rodentia: Echimyidae, Echimyinae) with Description of Two New Genera.” In Mammal 

Diversification: From Chromosomes to Phylogeography; a Celebration of the 

Career of James L. Patton, ed. E. A. Lacey and P. Myers. University of California Publications in Zoology, 133:247–306.

Emmons, L. H., and F. Feer. 1990. Neotropical Rainforest Mammals: A Field 

Guide. Chicago: University of Chicago Press.

Engstrom, M. D., and J. R. Choate. 1979. Systematics of the Northern Grasshopper Mouse (Onychomys leucogaster) on the Central Great Plains. Journal of 

Mammalogy, 60:723–739. http://dx.doi.org/10.2307/1380188.

Engstrom, M. D., O. Sanchez-Herrera, and G. Urbano-Vidales. 1992. Distribution, 

Geographic Variation, and Systematic Relationships within Nelsonia (Rodentia: Sigmodontinae). Proceedings of the Biological Society of Washington, 

105:867–881.

Ernest, K. A. 1986. Nectomys squamipes. Mammalian Species, 265:1–5. http://dx

.doi.org/10.2307/3503779.

Fadda, C., M. Corti, and E. Verheyen. 2001. Molecular Phylogeny of Myomys/

Stenocephalemys Complex and Its Relationships with Related African Genera. Biochemical Systematics and Ecology, 29:585–596. http://dx.doi.org/10

.1016/S0305-1978(00)00087-9.

Feldhamer, G. A. 1999. “Muskrat, Ondatra zibethicus.” In The Smithsonian Book 

of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 652–653. 

Washington, D.C.: Smithsonian Institution Press.
9 8 • smithsonian contributions to zoology

Feldhamer, G. A., and D. W. Linzey. 2008. “The Golden Mouse: Taxonomy and 

Natural History.” In The Golden Mouse, Ecology and Conservation, ed. 

G. W. Barrett and G. A. Feldhamer, pp. 21–35. New York: Springer.

Fernandez, J. A., F. Garcia-Campusano, and M. S. Hafner. 2010. Peromyscus difficilis (Rodentia: Cricetidae). Mammalian Species, 42:220–229. http://dx.doi

.org/10.1644/867.1.

Findley, J. S., and C. J. Jones. 1960. Geographic Variation in the Yellow-Nosed 

Cotton Rat. Journal of Mammalogy, 46:462–469. http://dx.doi.org/10.2307

/1377534.

Fisher, R. D., and C. A. Ludwig. 2012. Catalog of Type Specimens of Recent Mammals: Rodentia (Sciuromorpha and Castorimorpha) in the National Museum 

of Natural History, Smithsonian Institution. Smithsonian Contributions to 

Zoology, No. 640. Washington, D.C.: Smithsonian Institution Scholarly 

Press. http://dx.doi.org/10.5479/si.00810282.640.1. 

Frey, J. K. 1999a. “Mogollon Vole, Microtus mogollonensis.” In The Smithsonian Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 

634–635. Washington, D.C.: Smithsonian Institution Press.

———. 1999b. “Osgood’s Mouse, Peromyscus gratus.” In The Smithsonian Book 

of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 570–571. 

Washington, D.C.: Smithsonian Institution Press.

Gannon, W. L. 1999. “Pacific Jumping Mouse, Zapus trinotatus.” In The Smithsonian Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 

669–670. Washington, D.C.: Smithsonian Institution Press.

Gardner, A. L. 1990. “Two New Mammals from Southern Venezuela and Comments on the Affinities of the Highland Fauna of Cerro de la Neblina.” In 

Advances in Neotropical Mammalogy, ed. K. H. Redford and J. F. Eisenberg, 

pp. 411–424. Gainesville, Fla.: Sandhill Crane Press.

Gardner, M. C. 1946. A New Cotton Rat from Virginia. Proceedings of the Biological Society of Washington, 59:137–138.

———. 1948a. An Undescribed Eastern Cotton Rat. Proceedings of the Biological 

Society of Washington, 61:97–98.

———. 1948b. An Undescribed Cotton Rat from New Mexico. Journal of Mammalogy, 29:65. http://dx.doi.org/10.2307/1375282.

George, S. B. 1999. “Eastern Heather Vole, Phenacomys ungava.” In The Smithsonian Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 

618–619. Washington, D.C.: Smithsonian Institution Press.

Goldman, E. A. 1904. Descriptions of Five New Mammals from Mexico. Proceedings of the Biological Society of Washington, 17:79–82.

———. 1905. Twelve New Wood Rats of the Genus Neotoma. Proceedings of the 

Biological Society of Washington, 18:27–34.

———. 1909. Five New Woodrats of the Genus Neotoma from Mexico. Proceedings of the Biological Society of Washington, 22:139–142.

———. 1910. Revision of the Wood Rats of the Genus Neotoma. North American 

Fauna, 31:1–107. http://dx.doi.org/10.3996/nafa.31.0001.

———. 1911. Three New Mammals from Central and South America. Proceedings 

of the Biological Society of Washington, 24:237–240.

———. 1912a. New Mammals from Eastern Panama. Smithsonian Miscellaneous 

Collections, 60(2):1–18.

———. 1912b. Descriptions of Twelve New Species and Subspecies of Mammals 

from Panama. Smithsonian Miscellaneous Collections, 56(36):1–11.

———. 1913. Descriptions of New Mammals from Panama and Mexico. Smithsonian Miscellaneous Collections, 60(22):1–20.

———. 1915a. Five New Mammals from Mexico and Arizona. Proceedings of the 

Biological Society of Washington, 28:133–138.

———. 1915b. Five New Rice Rats of the Genus Oryzomys from Middle America. 

Proceedings of the Biological Society of Washington, 28:127–130.

———. 1916. A New Vesper Rat from Nicaragua. Proceedings of the Biological 

Society of Washington, 29:155–156.

———. 1917. New Mammals from North and Middle America. Proceedings of the 

Biological Society of Washington, 30:107–116.

———. 1918a. The Rice Rats of North America (Genus Oryzomys). North American Fauna, 43:1–100. http://dx.doi.org/10.3996/nafa.43.0001.

———. 1918b. Five New Mammals from Arizona and Colorado. Proceedings of 

the Biological Society of Washington, 31:21–26.

———. 1920. Mammals of Panama. Smithsonian Miscellaneous Collections, 

69(5):1–309.

———. 1927. A New Wood Rat from Arizona. Proceedings of the Biological Society of Washington, 40:205–206.

———. 1928. Three New Rodents from Western Arizona. Proceedings of the Biological Society of Washington, 41:203–206.

———. 1931. A New Agouti from Costa Rica. Journal of the Washington Academy of Sciences, 21:481.

———. 1932a. Two New Mammals from Honduras. Proceedings of the Biological 

Society of Washington, 45:121–124.

———. 1932b. Review of Wood Rats of Neotoma lepida Group. Journal of Mammalogy, 13:59–67. http://dx.doi.org/10.2307/1374173.

———. 1932c. A New Muskrat from Arizona. Proceedings of the Biological Society of Washington, 45:93–94.

———. 1932d. Two New Rodents from Arizona. Proceedings of the Biological 

Society of Washington, 45:89–92.

———. 1933. New Mammals from Arizona, New Mexico, and Colorado. Journal 

of the Washington Academy of Sciences, 23:463–473.

———. 1935. A New Brown Mouse of the Genus Scotinomys from Mexico. Proceedings of the Biological Society of Washington, 48:141–142.

———. 1937a. New Rodents from Middle America. Journal of the Washington 

Academy of Sciences, 27:418–423.

———. 1937b. A New Canyon Mouse of the Genus Peromyscus from Arizona. 

Journal of Mammalogy, 18:92–93. http://dx.doi.org/10.2307/1374319.

———. 1937c. Four New Mammals from Utah. Proceedings of the Biological Society of Washington, 50:221–226.

———. 1938a. A New Woodrat of the Genus Hodomys. Journal of the Washington Academy of Sciences, 28:498–499.

———. 1938b. Notes on the Voles of the Microtus longicaudus Group. Journal of 

Mammalogy, 19:491–492. http://dx.doi.org/10.2307/1374240.

———. 1938c. Three New Races of Microtus mexicanus. Journal of Mammalogy, 

19:493–495. http://dx.doi.org/10.2307/1374241.

———. 1938d. Six New Rodents from Coahuila and Texas and Notes on the Status 

of Several Described Forms. Proceedings of the Biological Society of Washington, 51:55–62.

———. 1939. Nine New Mammals from Islands in the Great Salt Lake, Utah. 

Journal of Mammalogy, 20:351–357. http://dx.doi.org/10.2307/1374264.

———. 1941a. Remarks on Voles of the Genus Lemmiscus, with One Described as 

New. Proceedings of the Biological Society of Washington, 54:69–72.

———. 1941b. A New Western Subspecies of Golden Mouse. Proceedings of the 

Biological Society of Washington, 54:189–192.

———. 1942a. Three New Rodents from Southern Utah. Proceedings of the Biological Society of Washington, 55:75–78.

———. 1942b. A New White-Footed Mouse from Mexico. Proceedings of the Biological Society of Washington, 55:157–158.

Goldman, E. A., and M. C. Gardner. 1947. Two New Cotton Rats. Journal of 

Mammalogy, 28:57–59. http://dx.doi.org/10.2307/1375495.

Gonzalez-Ruiz, N., J. Ramirez-Pulido, and H. H. Genoways. 2007. Review of the 

Harvest Mice (Genus Reithrodontomys) in the Mexican State of Mexico. 

Western North American Naturalist, 67:238–250. http://dx.doi.org/10.3398

/1527-0904(2007)67[238:ROTHMG]2.0.CO;2. 

Granjon, L., A. Bonnet, W. Hamdine, and V. Volobouev. 1999. Reevaluation of the 

Taxonomic Status of North African Gerbils Usually Referred to as Gerbillus pyramidum (Gerbillinae, Rodentia): Chromosomal and Biometrical Data. 

Zeitschrift für Saugetierkunde, 64:298–307.

Granjon, L., P. Colangelo, C. Tatard, M. Colyn, G. Dobigny, and V. Nicolas. 2012. 

Intrageneric Relationships within Gerbilliscus (Rodentia, Muridae, Gerbillinae), with Characterization of an Additional West African Species. Zootaxa, 

3325:1–25.

Grayson, D. K. 1999. “Bushy-Tailed Woodrat, Neotoma cinerea.” In The Smithsonian Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 

598–600. Washington, D.C.: Smithsonian Institution Press.

Greenbaum, I. F. 1999. “Northwestern Deer Mouse, Peromyscus keeni.” In The 

Smithsonian Book of North American Mammals, ed. D. E. Wilson and 

S. Ruff, pp. 571–572. Washington, D.C.: Smithsonian Institution Press.

Grinnell, J. 1913. A Distributional List of the Mammals of California. Proceedings 

of the California Academy of Sciences, 4th ser., 3:265–390.

———. 1933. Review of the Recent Mammal Fauna of California. University of 

California Publications in Zoology, 40:71–234.

Grinnell, J., and R. T. Orr. 1934. Systematic Review of the californicus Group of the 

Rodent Genus Peromyscus. Journal of Mammalogy, 15:210–220. http://dx

.doi.org/10.2307/1373853.

Gromov, I. M., and M. A. Erbajeva. 1995. The Mammals of Russia and Adjacent 

Territories: Lagomorphs and Rodents. [In Russian.] St. Petersburg: Russian 

Academy of Sciences, Zoological Institut. 

Gromov, I. M., and I. Ya. Polyakov. 1992. Fauna of the USSR: Voles (Microtinae).

Volume 3, bk. 8. New Delhi: Pauls Press.

Hafner, D. J., K. E. Peterson, and T. L. Yates. 1981. Evolutionary Relationships of 

Jumping Mice (Genus Zapus) of the Southwestern United States. Journal of 

Mammalogy, 62:501–512. http://dx.doi.org/10.2307/1380398.
number 642 • 9 9

Hall, E. R. 1934. Mammals Collected by T. T. and E. B. McCabe in the Brown 

Lake Region of British Columbia. University of California Publications in 

Zoology, 40:363–386.

———. 1935. Nevadan Races of the Microtus montanus Group of Meadow Mice. 

University of California Publications in Zoology, 40:417–428.

———. 1938. Notes on the Meadow Mice Microtus montanus and M. nanus with 

Description of a New Subspecies from Colorado. Proceedings of the Biological Society of Washington, 51:131–134.

———. 1946. The Mammals of Nevada. Berkeley: University of California Press.

———. 1955. A New Subspecies of Wood Rat from Nayarit, Mexico with New 

Name-Combinations for the Neotoma mexicana Group. Journal of the 

Washington Academy of Sciences, 45:328–332.

———. 1981. The Mammals of North America. Volume 2. 2nd ed. New York: 

John Wiley.

Hall, E. R., and E. L. Cockrum. 1952. Comments on the Taxonomy and Geographic Distribution of North American Microtines. University of Kansas 

Publications, Museum of Natural History, 5:293–312.

———. 1953. A Synopsis of North American Microtine Rodents. University of 

Kansas Publications, Museum of Natural History, 5:373–498.

Hall, E. R., and H. H. Genoways. 1970. Taxonomy of the Neotoma albigula Group 

of Woodrats in Central Mexico. Journal of Mammalogy, 51:504–516. http://

dx.doi.org/10.2307/1378390. 

Hall, E. R., and D. F. Hoffmeister. 1942. Geographic Variation in the Canyon 

Mouse, Peromyscus crinitus. Journal of Mammalogy, 23:51–65. http://

dx.doi.org/10.2307/1374855.

Hall, E. R., and K. R. Kelson. 1952. Comments on the Taxonomy and Geographic 

Distribution of Some North American Rodents. University of Kansas Publications, Museum of Natural History, 5:343–371.

———. 1959. The Mammals of North America. Volume 2. New York: Ronald Press.

Hallett, J. G. 1999. “Western Heather Vole, Phenacomys intermedius.” In The 

Smithsonian Book of North American Mammals, ed. D. E. Wilson and 

S. Ruff, pp. 617–618. Washington, D.C.: Smithsonian Institution Press.

Hamilton, W. J., Jr. 1955. Two New Rice Rats (Genus Oryzomys) from Florida. 

Proceedings of the Biological Society of Washington, 68:83–86.

Handley, C. O., Jr. 1952. A New Pine Mouse (Pitymys pinetorum carbonarius) 

from the Southern Appalachian Mountains. Journal of the Washington Academy of Sciences, 42:152–153.

———.1953. Three New Lemmings (Dicrostonyx) from Arctic America. Journal of 

the Washington Academy of Sciences, 43:197–200.

———. 1959. A Review of the Genus Hoplomys (Thick-Spined Rats), with Description of a New Form from Isla de Escudo de Veraguas, Panama. Smithsonian Miscellaneous Collections, 139(4):1–10.

———. 1966. “Checklist of the Mammals of Panama.” In Ectoparasites of Panama, ed. R. L. Tipton and V. J. Tipton, pp. 753–795. Chicago: Field Museum 

of Natural History.

Handley, C. O., Jr., and E. Mondolfi. 1963. A New Species of Fish-Eating Rat, 

Icthyomys, from Venezuela. Acta Biologica Venezuelica, 3:417–419.

Handley, C. O., Jr., and R. H. Pine. 1992. A New Species of Prehensile-Tailed 

Porcupine, Genus Coendou Lacepede, from Brazil. Mammalia, 56:237–244.

Harlan, R. 1832. Description of a New Species of Quadruped of the Genus Arvicola of Lacepede, or Hypudoes, of Illiger. Monthly American Journal of 

Geology and Natural Science, 1:446–447.

Hart, E. B., M. C. Belk, E. Jordan, and M. W. Gonzales. 2004. Zapus princeps. 

Mammalian Species, 749:1–7. http://dx.doi.org/10.1644/749. 

Hayman, R. W., and G. W. C. Holt. 1940. “A List of Named Forms (1757–1936).” 

In The Families and Genera of Living Rodents. Volume 1, Rodents Other 

Than Muridae, ed. J. R. Ellerman, pp. 85–653. London: British Museum 

(Natural History).

———. 1941. “A List of Named Forms (1758–1936).” In The Families and Genera of Living Rodents. Volume 2, Family Muridae, ed. J. R. Ellerman, pp. 

76–641. London: British Museum (Natural History).

Helgen, K. M., and T. L. McFadden. 2001. Type Specimens of Recent Mammals in 

the Museum of Comparative Zoology. Bulletin of the Museum of Comparative Zoology, 157:1–181.

Heller, E. 1909a. Two New Rodents from British East Africa. Smithsonian Miscellaneous Collections, 52:471–472.

———. 1909b. A New Rodent of the Genus Georychus. Smithsonian Miscellaneous Collections, 52:469–470.

———. 1910a. Descriptions of Seven New Species of East African Mammals. 

Smithsonian Miscellaneous Collections, 56(9):1–5.

———. 1910b. Five New Rodents from British East Africa. Smithsonian Miscellaneous Collections, 54(4):1–4.

———. 1910c. Descriptions of Seven New Species of East African Mammals. 

Smithsonian Miscellaneous Collections, 56(9):1–5.

———. 1911. New Species of Rodents and Carnivores from Equatorial Africa. 

Smithsonian Miscellaneous Collections, 56(17):1–16.

———. 1912. New Rodents from British East Africa. Smithsonian Miscellaneous 

Collections, 59(16):1–20.

———. 1914. New Subspecies of Mammals from Equatorial Africa. Smithsonian 

Miscellaneous Collections, 63(7):1–12.

Hershkovitz, P. 1944. A Systematic Review of the Neotropical Water Rats of the 

Genus Nectomys (Cricetinae). Miscellaneous Publications, Museum of Zoology, University of Michigan, 58:9–88.

———. 1948a. Mammals of Northern Colombia. Preliminary Report No. 3: Water Rats (Genus Nectomys), with Supplemental Notes on Related Forms. 

Proceedings of the United States National Museum, 98:49–56. http://dx.doi

.org/10.5479/si.00963801.3221.49.

———. 1948b. Mammals of Northern Colombia. Preliminary Report No. 2: 

Spiny Rats (Echimyidae), with Supplemental Notes on Related Forms. Proceedings of the United States National Museum, 97:125–140. http://dx.doi

.org/10.5479/si.00963801.3214.125.

———. 1962. Evolution of Neotropical Cricetine Rodents (Muridae) with Special 

Reference to the Phyllotine Group. Fieldiana Zoology, 46:1–524.

Heske, E. J., and W. Z. Lidicker Jr. 1999. “California Vole, Microtus californicus.” In The Smithsonian Book of North American Mammals, ed. D. E. 

Wilson and S. Ruff, pp. 626–628. Washington, D.C.: Smithsonian Institution Press.

Hinesley, L. L. 1979. Systematics and Distribution of Two Chromosome Forms in 

the Southern Grasshopper Mouse, Genus Onychomys. Journal of Mammalogy, 69:117–128. http://dx.doi.org/10.2307/1379763.

Hinton, M. A. C. 1926. Monograph of the Voles and Lemmings (Microtinae), Living and Extinct. Volume 1. London: British Museum (Natural History).

Hoffmeister, D. F. 1951. A Taxonomic and Evolutionary Study of the Pinon Mouse, 

Peromyscus truei. Illinois Biological Monographs, 21:1–104.

———. 1956. Mammals of the Graham (Pinaleno) Mountains, Arizona. American 

Midland Naturalist, 55:257–288. http://dx.doi.org/10.2307/2422591.

———. 1986. Mammals of Arizona. Tucson: University of Arizona Press.

Hoffmeister, D. F., and V. E. Diersing. 1973. The Taxonomic Status of Peromyscus 

merriami goldmani Osgood, 1904. Southwestern Naturalist, 18:354–357. 

http://dx.doi.org/10.2307/3669759.

Hogan, K. M., M. C. Hedin, H. S. Koh, S. K. Davis, and I. F. Greenbaum. 

1993. Systematic and Taxonomic Implications of Karyotypic, Electrophoretic, and Mitochondrial-DNA Variation in Peromyscus from the Pacific 

Northwest. Journal of Mammalogy, 74:819–831. http://dx.doi.org/10.2307

/1382420.

Holden, M. E., and G. G. Musser. 2005. “Family Dipodidae.” In Mammal Species 

of the World: A Taxonomic and Geographic Reference, Volume 2, 3rd ed., 

ed. D. E. Wilson and D. M. Reeder, pp. 871–893. Baltimore, Md.: Johns 

Hopkins University Press.

Holler, N. R. 1999. “Oldfield Mouse, Peromyscus polionotus.” In The Smithsonian Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 

580–582. Washington, D.C.: Smithsonian Institution Press.

Hollister, N. 1910a. A New Muskrat from the Great Plains. Proceedings of the 

Biological Society of Washington, 23:125–126.

———1910b. Descriptions of Two New Muskrats. Proceedings of the Biological 

Society of Washington, 23:1–2.

———. 1911. Two New Species of Epimys from Luzon. Proceedings of the Biological Society of Washington, 24:89–90.

———. 1912. New Mammals from the Highlands of Siberia. Smithsonian Miscellaneous Collections, 60(14):1–6.

———. 1913a. Mammals Collected by the Smithsonian-Harvard Expedition to the 

Altai Mountains, 1912. Proceedings of the United States National Museum, 

45:507–532. http://dx.doi.org/10.5479/si.00963801.45-1990.507.

———. 1913b. Three New Species of Grasshopper Mice. Proceedings of the Biological Society of Washington, 26:215–216.

———. 1913c. Two New Mammals from the Siberian Altai. Smithsonian Miscellaneous Collections, 60(24):1–3.

———. 1913d. A Review of the Philippine Land Mammals in the United States 

National Museum. Proceedings of the United States National Museum, 

46:299–341. http://dx.doi.org/10.5479/si.00963801.46-2028.299.

———. 1913e. The Type Species of Cuniculus Brisson. Proceedings of the Biological Society of Washington, 26:79.

———. 1914a. Four New Mammals from Tropical America. Proceedings of the 

Biological Society of Washington, 27:103–106.
100 • smithsonian contributions to zoology

———. 1914b. A Systematic Account of the Grasshopper Mice. Proceedings of 

the United States National Museum, 47:427–489. http://dx.doi.org/10.5479/

si.00963801.47-2057.427.

———. 1914c. Four New Neotropical Rodents. Proceedings of the Biological Society of Washington, 27:57–60.

———. 1914d. Descriptions of Four New Mammals from Tropical America. Proceedings of the Biological Society of Washington, 27:141–144.

———. 1916. Three New Murine Rodents from Africa. Smithsonian Miscellaneous 

Collections, 66(10):1–3.

———. 1918. New Rodents from British East Africa. Smithsonian Miscellaneous 

Collections, 68(10):1–3.

———. 1919. East African Mammals in the United States National Museum Part 

II. Rodentia, Lagomorpha, and Tubulidentata. Bulletin of the United States 

National Museum, 99:1–184. http://dx.doi.org/10.5479/si.03629236.99.i.

Honacki, J. H., K. E. Kinman, and J. W. Koeppl, eds. 1982. Mammal Species of 

the World: A Taxonomic and Geographic Reference. Lawrence, Kans.: Allen 

Press.

Hooper, E. T. 1938. Geographic Variation in Woodrats of the Species Neotoma 

fuscipes. University of California Publications in Zoology, 42:213–246.

———. 1952. A Systematic Review of the Harvest Mice (Genus Reithrodontomys) 

of Latin America. Miscellaneous Publications, Museum of Zoology, University of Michigan, 77:1–256.

———. 1955. Mammals of Western Mexico. Occasional Papers of the Museum of 

Zoology, University of Michigan, 565:1–26.

———. 1958. The Male Phallus in Mice of the Genus Peromyscus. Miscellaneous 

Publications, Museum of Zoology, University of Michigan, 105:1–24.

———. 1972. A Synopsis of the Rodent Genus Scotinomys. Occasional Papers of 

the Museum of Zoology, University of Michigan, 665:1–32.

Howell, A. B. 1923a. Corrected Type Localities of Some California Mammals. 

Journal of Mammalogy, 4:266.

———. 1923b. Descriptions of Two New Rodents of the Genus Phenacomys. Proceedings of the Biological Society of Washington, 36:157–158.

———. 1926. Voles of the Genus Phenacomys. North American Fauna, 48:1–66.

———. 1927a. Revision of the American Lemming Mice. North American Fauna, 

50:1–37.

———. 1927b. Two New Chinese Rats. Proceedings of the Biological Society of 

Washington, 40:43–46.

———. 1928. New Chinese Mammals. Proceedings of the Biological Society of 

Washington, 41:41–44.

Howell, A. H. 1910. Notes on Mammals of the Middle Mississippi Valley, with 

Description of a New Woodrat. Proceedings of the Biological Society of 

Washington, 23:23–34.

———. 1914. Revision of the American Harvest Mice (Genus Reithrodontomys). 

North American Fauna, 36:1–97.

———. 1916. Description of a New Pine Mouse from Florida. Proceedings of the 

Biological Society of Washington, 29:83–84.

———. 1920a. Description of a New Race of the Florida Water-Rat (Neofiber alleni). Journal of Mammalogy, 1:79–80. http://dx.doi.org/10.2307/1373747.

———. 1920b. Description of a New Species of Beach Mouse from Florida. Journal of Mammalogy, 1:237–240. http://dx.doi.org/10.2307/1373248.

———. 1932. Descriptions of Two New Harvest Mice from Honduras. Proceedings of the Biological Society of Washington, 45:125–126.

———. 1935. The Harvest Mice of the San Luis Valley, Colorado. Journal of Mammalogy, 16:143–144. http://dx.doi.org/10.2307/1374364.

———. 1939a. Description of a New Subspecies of the Golden Mouse. Journal of 

Mammalogy, 20:498. http://dx.doi.org/10.2307/1374599. 

———. 1939b. Descriptions of Five New Mammals from Florida. Journal of Mammalogy, 20:363–365. http://dx.doi.org/10.2307/1374266.

———. 1940. A New Race of the Harvest Mouse (Reithrodontomys) from Virginia. Journal of Mammalogy, 21:346. http://dx.doi.org/10.2307/1374764.

———. 1943. Two New Cotton Rats from Florida. Proceedings of the Biological 

Society of Washington, 56:73–76.

Hubbard, C. A. 1970. A New Species of Tatera from Tanzania with a Description of Its Life History and Habits Studied in Captivity. Zoologica Africana, 

5:237–247.

Huckaby, D. C. 1980. Species Limits in the Peromyscus mexicanus Group (Mammalia: Rodentia: Muroidea). Contributions in Science, Natural History Museum of Los Angeles County, 326:1–24.

Humphrey, S. R., and H. W. Setzer. 1989. Geographic Variation and Taxonomic 

Revision of Rice Rats (Oryzomys palustris and O. argentatus) of the United 

States. Journal of Mammalogy, 70:557–570. http://dx.doi.org/10.2307

/1381427.

Hunt, J. L., J. E. Morris, and T. L. Best. 2004. Nyctomys sumichrasti. Mammalian 

Species, 754:1–6. http://dx.doi.org/10.1644/754.

Hutterer, R., and F. Dieterlen. 1984. Zwei Neue Arten der Gattung Grammomys

aus Athiopien und Kenia (Mammalia: Muridae). Stuttgarter Beitrage zur 

Naturkunde, Series A (Biologie), 374:1–18.

International Commission on Zoological Nomenclature. 1999. International Code 

of Zoological Nomenclature. London: International Trust for Zoological 

Nomenclature.

Jackson, H. H. T. 1919. The Wisconsin Napaeozapus. Proceedings of the Biological Society of Washington, 32:9–10.

———. 1941. A New Pine Mouse, Genus Pitymys, from Wisconsin. Proceedings of 

the Biological Society of Washington, 54:201–202.

Jameson, E. W., Jr. 1961. Relationships of the Red-Backed Voles of Japan. Pacific 

Science, 15:594–604.

Johnson, D. H. 1945. The Spiny Rats of the Riu Kiu Islands. Proceedings of the 

Biological Society of Washington, 59:169–170.

———. 1946. A New Philippine Rat Allied to “Bullimus” bagobus Mearns. Journal of the Washington Academy of Sciences, 36:317–320.

———. 1959. Four New Mammals from the Northern Territory of Australia. Proceedings of the Biological Society of Washington, 72:183–188.

———. 1962. Two New Murine Rodents. Proceedings of the Biological Society of 

Washington, 75:317–319.

Johnson, D. H., and J. K. Jones Jr. 1955. Three New Rodents of the Genera Micromys and Apodemus from Korea. Proceedings of the Biological Society of 

Washington, 68:167–174.

Johnson, M. L. 1968. Application of Blood Protein Electrophoretic Studies to Problems in Mammalian Taxonomy. Systematic Zoology, 17:23–30. http://dx.doi

.org/10.2307/2412392.

Johnson, M. L., and C. Maser. 1982. Genetic Relationships of Phenacomys albipes. 

Northwest Science, 56:17–19.

Jones, C., and R. D. Fisher. 1973. Comments on the Type-Specimen of Neotoma 

desertorum sola Merriam 1894 (Mammalia: Rodentia). Proceedings of the 

Biological Society of Washington, 86:435–438.

Jones, J. K., Jr. 1956. Comments on the Taxonomic Status of Apodemus peninsulae, with Description of a New Subspecies from North China. University of 

Kansas Publications, Museum of Natural History, 9:337–346.

Jones, J. K., Jr., D. C. Carter, H. H. Genoways, R. S. Hoffmann, D. W. Rice, and 

C. Jones. 1986. Revised Checklist of North American Mammals North of 

Mexico. Occasional Papers, The Museum of Texas Tech University, 107:1–22.

Jones, J. K., Jr., and D. H. Johnson. 1955. A New Reed Vole, Genus Microtus, 

from Central Korea. Proceedings of the Biological Society of Washington, 

68:193–196.

Jones, J. K., Jr., and B. Mursaloglu. 1961. Geographic Variation in the Harvest 

Mouse, Reithrodontomys megalotis, on the Central Great Plains and in Adjacent Regions. University of Kansas Publications, Museum of Natural History, 14:9–27.

Kalcounis-Ruppell, M. C., and T. R. Spoon. 2009. Peromyscus boylii (Rodentia: 

Cricetidae). Mammalian Species, 838:1–14. http://dx.doi.org/10.1644/838.1.

Kellogg, R. 1918. Revision of the Microtus californicus Group of Meadow Mice. 

University of California Publications in Zoology, 21:1–42.

———. 1939. A New Red-Backed Mouse from Kentucky and Virginia. Proceedings of the Biological Society of Washington, 52:37–40.

———. 1945a. Two New Philippine Rodents. Proceedings of the Biological Society 

of Washington, 58:121–124.

———1945b. A New Australian Naked-Tailed Rat (Melomys). Proceedings of the 

Biological Society of Washington, 58:69–72.

———. 1945c. Two Rats from Morotai Island. Proceedings of the Biological Society of Washington, 58:65–68.

———. 1946. Three New Mammals from the Pearl Islands, Panama. Proceedings 

of the Biological Society of Washington, 59:57–62.

King, T. L., J. F. Switzer, C. L. Morrison, M. S. Eackles, C. C. Young, B. A. Lubinski, and P. A. Cryon. 2006. Comprehensive Genetic Analyses Reveal Evolutionary Distinction of a Mouse (Zapus hudsonius preblei) Proposed for 

Delisting from the US Endangered Species Act. Molecular Ecology, 15:4331–

4359. http://dx.doi.org/10.1111/j.1365-294X.2006.03080.x. 

Kloss, C. B. 1919. On Mammals Collected in Siam. Journal of the Natural History 

Society of Siam, 3:333–407.

Kratochvil, J. I., L. Rodrigues, and V. Barus. 1978. Capromyinae (Rodentia) of 

Cuba. Acta Scientiarum Naturalium, Academiae Scientiarum Bohemoslovacae (Brno), n.s., 12(11):1–60.

Krutzsch, P. H. 1954. North American Jumping Mice (Genus Zapus). University of 

Kansas Publications, Museum of Natural History, 7:349–472.
number 642 • 101

Lackey, J. A. 1999. “White-Footed Mouse, Peromyscus leucopus.” In The Smithsonian Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 

572–574. Washington, D.C.: Smithsonian Institution Press.

Laurie, E. M. O., and J. E. Hill. 1954. List of Land Mammals of New Guinea, Celebes, 

and Adjacent Islands 1758–1952. London: British Museum (Natural History).

Lawlor, T. E. 1969. A Systematic Study of the Rodent Genus Ototylomys. Journal 

of Mammalogy, 50:28–42. http://dx.doi.org/10.2307/1378627.

———. 1971. Distribution and Relationships of Six Species of Peromyscus in Baja 

California and Sonora, Mexico. Occasional Papers of the Museum of Zoology, University of Michigan, 661:1–22.

———. 1982. Ototylomys phyllotis. Mammalian Species, 181:1–3.

Lay, D. M. 1983. Taxonomy of the Genus Gerbillus (Rodentia: Gerbillinae) with 

Comments on the Application of Generic and Subgeneric Names and an Annotated List of Species. Zeitschrift für Saugetierkunde, 48:329–354.

Lay, D. M., and C. F. Nadler. 1969. Hybridization in the Rodent Genus Meriones. I. 

Breeding and Cytological Analysis of Meriones shawi (♀) X Meriones libycus

(♂) Hybrids. Cytogenetics, 8:35–50.

———. 1975. A Study of Gerbillus (Rodentia: Muridae) East of the Euphrates River. 

Mammalia, 39:423–445. http://dx.doi.org/10.1515/mamm.1975.39.3.423.

LeConte, J. 1853. Descriptions of Three New Species of American Arvicola, with 

Remarks upon Some Other American Rodents. Proceedings of the Academy 

of Natural Sciences of Philadelphia, 6:404–415.

———. 1855. Description of Two New Species of Hesperomys. Proceedings of the 

Academy of Natural Sciences of Philadelphia, 7:442.

Lee, M. R., D. J. Schmidly, and C. C. Huheey. 1972. Chromosomal Variation in 

Certain Populations of Peromyscus boylii and Its Systematic Implications. 

Journal of Mammalogy, 53:697–707. http://dx.doi.org/10.2307/1379208.

Linzey, A. V. 1999a. “Cotton Mouse, Peromyscus gossypinus.” In The Smithsonian Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 

569–570. Washington, D.C.: Smithsonian Institution Press.

———. 1999b. “Southern Bog Lemming, Synaptomys cooperi.” In The Smithsonian Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 

657–658. Washington, D.C.: Smithsonian Institution Press.

Long, C. A. 1965. The Mammals of Wyoming. University of Kansas Publications, 

Museum of Natural History, 14:493–758.

———. 1987. A New Subspecies of Southern Bog Lemming from Two Lake Michigan Isles (Mammalia: Rodentia). Mammalia, 51:324–326.

Ludwig, D. R. 1999. “Water Vole, Microtus richardsoni.” In The Smithsonian 

Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 644–

646. Washington, D.C.: Smithsonian Institution Press.

Lyon, M. W., Jr. 1906. Mammals of Banka, Mendanau, and Billiton Islands, between Sumatra and Borneo. Proceedings of the United States National Museum, 31:575–612. http://dx.doi.org/10.5479/si.00963801.31-1498.575.

———. 1907a. Notes on a Small Collection of Mammals from the Province of KanSu, China. Smithsonian Miscellaneous Collections, 50:133–138.

———. 1907b. Mammals of Batam Island, Rhio Archipelago. Proceedings of the 

United States National Museum, 31:653–657. http://dx.doi.org/10.5479/

si.00963801.31-1502.653.

———. 1907c. Notes on the Porcupines of the Malay Peninsula and Archipelago. 

Proceedings of the United States National Museum, 32:575–594. http://

dx.doi.org/10.5479/si.00963801.32-1552.575.

———. 1908. Mammals Collected in Eastern Sumatra By Dr. W. L. Abbott During 

1903, 1906, and 1907, with Descriptions of New Species and Subspecies. 

Proceedings of the United States National Museum, 34:619–679. http://

dx.doi.org/10.5479/si.00963801.34-1626.619.

———. 1909. Additional Notes on Mammals of the Rhio-Linga Archipelago, 

with Descriptions of New Species and a Revised List. Proceedings of the 

United States National Museum, 36:479–491. http://dx.doi.org/10.5479/si

.00963801.36-1684.479.

———. 1911. Mammals Collected By Dr. W. L. Abbott on Borneo and Some of the 

Small Adjacent Islands. Proceedings of the United States National Museum, 

40:53–146. http://dx.doi.org/10.5479/si.00963801.40-1809.53.

———. 1916a. Mammals Collected By Dr. W. L. Abbott on the Chain of Islands 

Lying off the Western Coast of Sumatra, with Descriptions of Twenty-Eight 

New Species and Subspecies. Proceedings of the United States National Museum, 52:437–462. http://dx.doi.org/10.5479/si.00963801.52-2188.437.

———. 1916b. Two New Mammals from Sumatra. Proceedings of the Biological 

Society of Washington, 29:209–212. 

Lyon, M. W., Jr., and W. H. Osgood. 1909. Catalogue of the Type-Specimens of 

Mammals in the United States National Museum, Including the Biological 

Survey Collection. Bulletin of the United States National Museum, 62:1–325. 

http://dx.doi.org/10.5479/si.03629236.62.i.

MacMillen, R. E. 1999. “Desert Woodrat, Neotoma lepida.” In The Smithsonian Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 

604–606. Washington, D.C.: Smithsonian Institution Press.

Mahoney, J. A., and B. J. Richardson. 1988. “Muridae.” In Zoological Catalogue 

of Australia. Volume 5, Mammalia, ed. D. W. Walton, pp. 154–192. Canberra: Australian Government Publishing Service.

Mann, G. 1944. Dos nuevas especies de Roedores. Biologica (Santiago, Chile), 1:95–113.

———. 1945. Mamiferos de Tarapaca. Biologica (Santiago, Chile), 2:23–98.

Mares, M. A. 1999. “White-Throated Woodrat, Neotoma albigula.” In The Smithsonian Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 

596–598. Washington, D.C.: Smithsonian Institution Press.

Massoia, E. 1980. El estado sistematico de cuatro especies de Cricetidos Sudamericanos y comentarios sabre otras especies congenericas (Mammalia-Rodentia). 

Ameghiniana, 17:280–287.

McAllister, J. A., and R. S. Hoffmann. 1988. Phenacomys intermedius. Mammalian 

Species, 205:1–8. http://dx.doi.org/10.2307/3504242.

Mearns, E. A. 1894. Description of a New Species of Cotton Rat (Sigmodon minima) from New Mexico. Proceedings of the United States National Museum, 

17:129–130. http://dx.doi.org/10.5479/si.00963801.994.129.

———. 1896a. Preliminary Diagnoses of New Mammals from the Mexican Border 

of the United States. Proceedings of the United States National Museum, 

19:1–4. Preprints.

———. 1896b. Preliminary Diagnoses of New Mammals from the Mexican Border 

of the United States. Proceedings of the United States National Museum, 

19:137–140. http://dx.doi.org/10.5479/si.00963801.1103.137. 

———. 1896c. Preliminary Diagnoses of New Mammals from the Mexican Border 

of the United States. Proceedings of the United States National Museum, 

18:1–4. Preprints.

———. 1896d. Preliminary Diagnoses of New Mammals from the Mexican Border 

of the United States. Proceedings of the United States National Museum, 

18:443–447. http://dx.doi.org/10.5479/si.00963801.1075.443.

———. 1897a. Descriptions of Six New Mammals from North America. Proceedings of the United States National Museum, 19:719–724. http://dx.doi

.org/10.5479/si.00963801.1121.719.

———. 1897b. Preliminary Diagnoses of New Mammals of the Genera Sciurus, 

Castor, Neotoma, and Sigmodon, from the Mexican Border of the United 

States. Proceedings of the United States National Museum, 20:1–4. Preprints.

———. 1898. Preliminary Diagnoses of New Mammals of the Genera Sciurus, 

Castor, Neotoma, and Sigmodon, from the Mexican Border of the United 

States. Proceedings of the United States National Museum, 20:501–505. 

http://dx.doi.org/10.5479/si.00963801.1132.501.

———. 1901. On the Mainland Forms of the Eastern Deermouse, Peromyscus 

leucopus (Rafinesque). Proceedings of the Biological Society of Washington, 

14:153–155.

———. 1905. Descriptions of New Genera and Species of Mammals from the Philippine Islands. Proceedings of the United States National Museum, 28:425–

460. http://dx.doi.org/10.5479/si.00963801.1402.425.

———. 1907. Mammals of the Mexican Boundary of the United States, Part 1: 

Families Didelphidae to Muridae. Bulletin of the United States National Museum, 56:1–530.

Medway, L. 1965. Mammals of Borneo; Field Keys and an Annotated Checklist. 

Journal of the Malayan Branch of the Royal Asiatic Society, 36:1–193.

Mellink, E., G. Ceballos, and J. Luevano. 2002. Population Demise and Extinction 

Threat of the Angel de la Guardia Deer Mouse (Peromyscus guardia). Biological Conservation, 108:107–111. http://dx.doi.org/10.1016/S0006-3207

(02)00095-2.

Merriam, C. H. 1887a. Description of a New Species of Wood-Rat from Cerros 

Island, Off Lower California. American Naturalist, 21:191–193.

———. 1887b. Description of a New Mouse from New Mexico, Hesperomys (Vesperimus) anthonyi sp. nov. Proceedings of the Biological Society of Washington, 

4:5–8.

———. 1888a. Description of a New Species of Field-Mouse (Arvicola pallidus) 

from Dakota. American Naturalist, 22:702–705. http://dx.doi.org/10.1086

/274761.

———. 1888b. Description of a New Prairie Meadow Mouse (Arvicola austerus 

minor) from Dakota and Minnesota. American Naturalist, 22:598–601. 

http://dx.doi.org/10.1086/274743.

———. 1888c. Description of a New Species of Meadow Mouse from the Black 

Hills of Dakota. American Naturalist, 22:934–935.

———. 1888d. Remarks on the Fauna of the Great Smoky Mountains; with Description of a New Species of Redbacked Mouse (Evotomys carolinensis). 

American Journal of Science, 3rd ser., 36:458–460.
102 • smithsonian contributions to zoology

———. 1889a. Descriptions of Two New Species and One New Subspecies of 

Grasshopper Mouse, with a Diagnosis of the Genus Onychomys, and a Synopsis of the Species and Subspecies. North American Fauna, 2:1–5. http://dx

.doi.org/10.3996/nafa.2.0001.

———. 1889b. Description of a New Genus (Phenacomys) and Four New Species 

of Arvicolinae. North American Fauna, 2:27–35. http://dx.doi.org/10.3996

/nafa.2.0009.

———. 1890a. Results of a Biological Survey of the San Francisco Mountain Region and Desert of the Little Colorado, Arizona. North American Fauna, 

3:1–136. http://dx.doi.org/10.3996/nafa.3.0001.

———. 1890b. Descriptions of Two New Species of Evotomys from the Pacific 

Coast Region of the United States. North American Fauna, 4:25–26. http://

dx.doi.org/10.3996/nafa.4.0004.

———. 1890c. Description of a New Evotomys from Colorado. North American 

Fauna, 4:23–24. http://dx.doi.org/10.3996/nafa.4.0003.

———. 1890d. Description of a New Species of Hesperomys from Southern Florida. 

North American Fauna, 4:53–54. http://dx.doi.org/10.3996/nafa.4.0011.

———. 1891a. Results of a Biological Reconnoissance of South-Central Idaho. 

North American Fauna, 5:1–108.

———. 1891b. Description of a New Evotomys from the Black Hills of South 

Dakota. North American Fauna, 5:119–133. http://dx.doi.org/10.3996/nafa

.5.0006.

———. 1892a. Descriptions of Nine New Mammals Collected by E. W. Nelson 

in the States of Colima and Jalisco, Mexico. Proceedings of the Biological 

Society of Washington, 7:164–174.

———. 1892b. Description of a New Genus and Species of Murine Rodent (Xenomys nelsoni) from the State of Colima, Western Mexico. Proceedings of the 

Biological Society of Washington, 7:159–163.

———. 1893. Two New Wood Rats from the Plateau Region of Arizona (Neotoma pinetorum and N. arizonae), with Remarks on the Validity of the Genus Teonoma of Gray. Proceedings of the Biological Society of Washington, 

8:109–112.

———. 1894a. A New Subfamily of Murine Rodents—The Neotominae—With Description of a New Genus and Species and a Synopsis of the Known Forms. 

Proceedings of the Academy of Natural Sciences of Philadelphia, 46:225–252.

———. 1894b. Abstract of a Study of the American Wood Rats, with Descriptions 

of Fourteen New Species and Subspecies of the Genus Neotoma. Proceedings 

of the Biological Society of Washington, 9:117–128.

———. 1896. Revision of the Lemmings of the Genus Synaptomys, with Descriptions of New Species. Proceedings of the Biological Society of Washington, 

10:55–64.

———. 1897a. Descriptions of Five New Rodents from the Coast Region of 

Alaska. Proceedings of the Biological Society of Washington, 11:221–223.

———. 1897b. Three New Jumping Mice (Zapus) from the Northwest. Proceedings of the Biological Society of Washington, 11:103–104.

———. 1897c. The Voles of the Subgenus Chilotus, with Descriptions of New Species. Proceedings of the Biological Society of Washington, 11:73–75.

———. 1897d. Descriptions of Two New Red Backed Mice (Evotomys) from Oregon. Proceedings of the Biological Society of Washington, 11:71–72.

———. 1897e. Nelsonia neotomodon, a New Genus and Species of Murine Rodent from Mexico. Proceedings of the Biological Society of Washington, 

11:277–279.

———. 1897f. Description of a New Muskrat from the Great Dismal Swamp, Virginia. Proceedings of the Biological Society of Washington, 11:143.

———. 1897g. Phenacomys preblei, a New Vole from the Mountains of Colorado. 

Proceedings of the Biological Society of Washington, 11:45.

———. 1898a. Descriptions of Twenty New Species and a New Subgenus of Peromyscus from Mexico and Guatemala. Proceedings of the Biological Society 

of Washington, 12:115–125.

———. 1898b. Descriptions of Two New Subgenera and Three New Species of 

Microtus from Mexico and Guatemala. Proceedings of the Biological Society 

of Washington, 12:105–108.

———. 1898c. Descriptions of Three New Rodents from the Olympic Mountains, 

Washington. Proceedings of the Academy of Natural Sciences of Philadelphia, 50:352–353.

———. 1898d. A New Genus (Neotomodon) and Three New Species of Murine 

Rodents from the Mountains of Southern Mexico. Proceedings of the Biological Society of Washington, 12:127–129.

———. 1898e. Mammals of the Tres Marias Islands, off Western Mexico. Proceedings of the Biological Society of Washington, 12:13–19.

———. 1899. Results of a Biological Survey of Mount Shasta California. North 

American Fauna, 16:1–179.

———. 1900a. Papers from the Harriman Alaska Expedition. I. Descriptions of 

Twenty-Six New Mammals from Alaska and British North America. Proceedings of the Washington Academy of Sciences, 2:13–30.

———. 1900b. Description of a New Harvest Mouse (Reithrodontomys) from 

Mexico. Proceedings of the Biological Society of Washington, 13:152.

———. 1901a. Two New Rodents from Northwestern California. Proceedings of 

the Biological Society of Washington, 14:125–126.

———. 1901b. Synopsis of the Rice Rats (Genus Oryzomys) of the United States 

and Mexico. Proceedings of the Washington Academy of Sciences, 3:273–

295.

———. 1901c. Six New Mammals from Cozumel Island, Yucatan. Proceedings of 

the Biological Society of Washington, 14:99–104.

———. 1901d. Seven New Mammals from Mexico, Including a New Genus of 

Rodents. Proceedings of the Washington Academy of Sciences, 3:559–563.

———. 1901e. Descriptions of 23 New Harvest Mice (Genus Reithrodontomys). 

Proceedings of the Washington Academy of Sciences, 3:547–558.

———. 1903a. Four New Mammals, Including a New Genus (Teanopus), from 

Mexico. Proceedings of the Biological Society of Washington, 16:79–82.

———. 1903b. Two New Wood Rats (Genus Neotoma) from State of Coahuila, 

Mexico. Proceedings of the Biological Society of Washington, 16:47–48.

———. 1904. Four New Grasshopper Mice, Genus Onychomys. Proceedings of 

the Biological Society of Washington, 17:123–126.

———. 1908a. Four New Rodents from California. Proceedings of the Biological 

Society of Washington, 21:145–148.

———. 1908b. Three New Rodents from Colorado. Proceedings of the Biological 

Society of Washington, 21:143–144.

Merritt, J. F. 1999a. “Southern Red-Backed Vole, Clethrionomys gapperi.” In 

The Smithsonian Book of North American Mammals, ed. D. E. Wilson and 

S. Ruff, pp. 613–615. Washington, D.C.: Smithsonian Institution Press.

———. 1999b. “California Mouse, Peromyscus californicus.” In The Smithsonian Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 

565–566. Washington, D.C.: Smithsonian Institution Press.

Miller, G. S., Jr. 1896a. The Beach Mouse of Muskeget Island. Proceedings of the 

Boston Society of Natural History, 27:75–87.

———. 1896b. Genera and Subgenera of Voles and Lemmings. North American 

Fauna, 12:1–78. http://dx.doi.org/10.3996/nafa.12.0001.

———. 1897a. Description of a New Vole from Oregon. Proceedings of the Biological Society of Washington, 11:67–68.

———. 1897b. Synopsis of the Voles of the Genus Phenacomys. Proceedings of the 

Biological Society of Washington, 11:77–87.

———. 1898a. Description of a New Genus and Species of Microtine Rodent from 

Siberia. Proceedings of the Academy of Natural Sciences of Philadelphia, 

50:368–371.

———. 1898b. Notes on the Arctic Red-Backed Mice. Proceedings of the Academy 

of Natural Sciences of Philadelphia, 50:358–367.

———. 1898c. Description of a New Rodent of the Genus Idiurus. Proceedings of 

the Biological Society of Washington, 12:73–76.

———. 1899a. The Voles Collected by Dr. W. L. Abbott in Central Asia. Proceedings of the Academy of Natural Sciences of Philadelphia, 51:281–298.

———. 1899b. A New Vole from Hall Island, Bering Sea. Proceedings of the Biological Society of Washington, 13:13–14.

———. 1899c. Description of a New Vole from Eastern Siberia. Proceedings of the 

Biological Society of Washington, 13:11–12.

———. 1900a. Key to the Land Mammals of Northeastern North America. Bulletin of the New York State Museum, 8:61–160.

———. 1900b. Preliminary Revision of the European Redbacked Mice. Proceedings of the Washington Academy of Sciences, 2:83–109.

———. 1900c. Seven New Rats Collected by Dr. W. L. Abbott in Siam. Proceedings 

of the Biological Society of Washington, 13:137–150.

———. 1900d. A New Gerbille from Eastern Turkestan. Proceedings of the Biological Society of Washington, 13:163–164.

———. 1900e. A Collection of Small Mammals from Mount Coffee, Liberia. Proceedings of the Washington Academy of Sciences, 2:631–649.

———. 1900f. Mammals Collected By Dr. W. L. Abbott on Islands in the North 

China Sea. Proceedings of the Washington Academy of Sciences, 2:203–246.

———. 1900g. Mammals Collected by Dr. W. L. Abbott on Pulo Lankawi and 

the Butang Islands. Proceedings of the Biological Society of Washington, 

13:187–193.

———. 1901a. A New Name for Mus obscurus Miller. Proceedings of the Biological Society of Washington, 14:178.

———. 1901b. Mammals Collected by Dr. W. L. Abbott on the Natuna Islands. 

Proceedings of the Washington Academy of Sciences, 3:111–138.
number 642 • 103

———. 1902a. Mammals Collected By Dr. W. L. Abbott in the Region of the Indragiri River, Sumatra. Proceeding of the Academy of Natural Sciences of 

Philadelphia, 54:143–159.

———. 1902b. The Mammals of the Andaman and Nicobar Islands. Proceedings of the United States National Museum, 24:751–795. http://dx.doi.

org/10.5479/si.00963801.24-1269.751.

———. 1903a. Descriptions of Two New Mole Rats. Proceedings of the Biological 

Society of Washington, 16:161–164.

———. 1903b. Seventy New Malayan Mammals. Smithsonian Miscellaneous Collections, 45:1–70.

———. 1903c. Mammals Collected by Dr. W. L. Abbott on the Coast and Islands 

of Northwest Sumatra. Proceedings of the United States National Museum, 

26:437–484. http://dx.doi.org/10.5479/si.00963801.26-1317.437.

———. 1903d. A New Name for Mus atratus Miller. Proceedings of the Biological 

Society of Washington, 16:50.

———. 1906a. Mammals Collected By Dr. W. L. Abbott in the Karimata Islands, 

Dutch East Indies. Proceedings of the United States National Museum, 

31:55–66. http://dx.doi.org/10.5479/si.00963801.31-1481.55.

———. 1906b. The Mammals of Engano Island, West Sumatra. Proceedings of 

the United States National Museum, 30:819–825. http://dx.doi.org/10.5479

/si.00963801.1472.819.

———. 1910a. A New Rodent of the Genus Saccostomus from British East Africa. 

Smithsonian Miscellaneous Collections, 54(5):1–2.

———. 1910b. Descriptions of Two New Genera and Sixteen New Species of Mammals from the Philippine Islands. Proceedings of the United States National 

Museum, 38:391–404. http://dx.doi.org/10.5479/si.00963801.38-1757.391.

———. 1911a. Four New Chinese Mammals. Proceedings of the Biological Society 

of Washington, 24:53–56.

———. 1911b. A New Jumping Mouse from New Mexico. Proceedings of the 

Biological Society of Washington, 24:253–254.

———. 1911c. Descriptions of Six New Mammals from the Malay Archipelago. 

Proceedings of the Biological Society of Washington, 24:25–28.

———. 1912a. List of North American Land Mammals in the United States National Museum. Bulletin of the United States National Museum, 79:1–455.

———. 1912b. Catalogue of the Mammals of Western Europe (Europe Exclusive 

of Russia) in the Collection of the British Museum. London: British Museum 

(Natural History).

———. 1912c. Two New Murine Rodents from Turkestan. Proceedings of the Biological Society of Washington, 25:59–60.

———. 1913a. Two New Murine Rodents from Baltistan. Proceedings of the Biological Society of Washington, 26:197–198.

———. 1913b. A New Vole from Mongolia. Smithsonian Miscellaneous Collections, 60(28):1–2.

———. 1913c. Some Overlooked Names of Sicilian Mammals. Proceedings of the 

Biological Society of Washington, 26:80–81.

———. 1913d. Fifty-One New Malayan Mammals. Smithsonian Miscellaneous 

Collections, 61(21):1–30.

———. 1914. Two New Murine Rodents from Eastern Asia. Proceedings of the 

Biological Society of Washington, 27:89–92.

———. 1924. List of North American Recent Mammals, 1923. Bulletin of the 

United States National Museum, 128:1–673. http://dx.doi.org/10.5479/

si.03629236.128.i.

———. 1927a. A New Pedetes from Tanganyika Territory. Proceedings of the Biological Society of Washington, 40:113–114.

———. 1927b. The Rodents of the Genus Plagiodontia. Proceedings of the United 

States National Museum, 72(16):1–8.

Miller, G. S., Jr. and N. Hollister. 1921a. Descriptions of Sixteen New Murine Rodents 

from Celebes. Proceedings of the Biological Society of Washington, 34:67–76.

———. 1921b. Twenty New Mammals Collected by H. C. Raven in Celebes. Proceedings of the Biological Society of Washington, 34:93–104.

Miller, G. S., Jr., and R. Kellogg. 1955. List of North American Recent Mammals. 

Bulletin of the United States National Museum, 205:1–954. http://dx.doi

.org/10.5479/si.03629236.205.

Miller, G. S., Jr., and J. A. G. Rehn. 1901. Systematic Results of the Study of North 

American Land Mammals to the Close of the Year 1900. Proceedings of the 

Boston Society of Natural History, 30:1–352.

Misonne, X. 1974. “Order Rodentia, Part 6.” In The Mammals of Africa: An Identification Manual, ed. J. Meester and H. W. Setzer, pp 1–39. Washington, 

D.C.: Smithsonian Institution Press.

Modi, W. S., and M. R. Lee. 1984. Systematic Implications of Chromosomal Banding Analyses of Populations of Peromyscus truei (Rodentia: Muridae). Proceedings of the Biological Society of Washington, 97:716–723.

Morgan, G. S. 1985. Taxonomic Status and Relationships of the Swan Island Hutia, Geocapromys thoracatus (Mammalia: Rodentia: Capromyidae), and the 

Zoogeography of the Swan Islands Vertebrate Fauna. Proceedings of the Biological Society of Washington, 98:29–46.

Mullen, L. M., S. N. Vignieri, J. A. Gore, and H. E. Hoekstra. 2009. Adaptive 

Basis of Geographic Variation: Genetic, Phenotypic, and Environmental Differences Among Beach Mouse Populations. Proceedings of the Royal Society 

B, Biological Sciences, 276:3809–3818.

Murie, O. J. 1930. A New Alaskan Microtus. Journal of Mammalogy, 11:74–75. 

http://dx.doi.org/10.2307/1373790.

Musser, G. G. 1964. Notes on Geographic Distribution, Habitat, and Taxonomy 

of Some Mexican Mammals. Occasional Papers of the Museum of Zoology, 

University of Michigan, 636:1–22.

———. 1969. Notes on Peromyscus (Muridae) of Mexico and Central America. 

American Museum Novitates, 2357:1–23.

———. 1971a. Peromyscus allophylus Osgood: A Synonym of Peromyscus gymnotis Thomas (Rodentia, Muridae). American Museum Novitates, 2453:1–10.

———. 1971b. Results of the Archbold Expeditions. No. 94. Taxonomic Status 

of Rattus tatei and Rattus frosti, Two Taxa of Murid Rodents Known from 

Middle Celebes. American Museum Novitates, 2454:1–19.

———. 1973. Zoogeographical Significance of the Ricefield Rat, Rattus argentiventer, on Celebes and New Guinea and the Identity of Rattus pesticulus. 

American Museum Novitates, 2511:1–30.

———. 1977a. Results of the Archbold Expeditions. No. 100. Notes on the Philippine Rat, Limnomys, and the Identity of Limnomys picinus, a Composite. 

American Museum Novitates, 2636:1–14.

———. 1977b. Epimys benguetensis, a Composite, and One Zoogeographic View 

of Rat and Mouse Faunas in the Philippines and Celebes. American Museum 

Novitates, 2624:1–15.

———. 1979. Results of the Archbold Expeditions. No. 102. The Species of Chiropodomys, Arboreal Mice of Indochina and the Malay Archipelago. Bulletin of the American Museum of Natural History, 162:377–445.

———. 1981a. The Giant Rat of Flores and Its Relatives East of Borneo and Bali. 

Bulletin of the American Museum of Natural History, 169:67–176.

———. 1981b. Result of the Archbold Expeditions. No. 105. Notes on Systematics of Indo-Malayan Murid Rodents, and Descriptions of New Genera and 

Species from Ceylon, Sulawesi, and the Philippines. Bulletin of the American 

Museum of Natural History, 168:225–334.

———. 1982. Results of the Archbold Expeditions. No. 108. The Definition of 

Apomys, a Native Rat of the Philippine Islands. American Museum Novitates, 2746:1–43.

Musser, G. G., and D. Califia. 1982. Results of the Archbold Expeditions. No. 106. 

Identities of Rats from Pulau Maratua and Other Islands off East Borneo. 

American Museum Novitates, 2726:1–30.

Musser, G. G., and M. D. Carleton. 1993. “Family Muridae.” In Mammal Species 

of the World: A Taxonomic and Geographic Reference, 2nd ed., ed. D. E. 

Wilson and D. M. Reeder, pp. 501–755. Washington, D.C.: Smithsonian Institution Press.

———. 2005. “Superfamily Muroidea.” In Mammal Species of the World: A Taxonomic and Geographic Reference, Volume 2, 3rd ed., ed. D. E. Wilson and 

D. M. Reeder, pp. 894–1531. Baltimore, Md.: Johns Hopkins University Press.

Musser, G. G., and L. K. Gordon. 1981. A New Species of Crateromys (Muridae) from the Philippines. Journal of Mammalogy, 62:513–525. http://dx.doi

.org/10.2307/1380399.

Musser, G. G., L. K. Gordon, and H. Sommer. 1982. Species Limits in the Philippine Murid, Chrotomys. Journal of Mammalogy, 63:514–521. http://dx.doi

.org/10.2307/1380457.

Musser, G. G., and L. R. Heaney. 1985. Philippine Rattus: A New Species from the 

Sulu Archipelago. American Museum Novitates, 2818:1–32.

———. 1992. Philippine Rodents: Definition of Tarsomys and Limnomys Plus a 

Preliminary Assessment of Phylogenetic Patterns among Native Philippine 

Murines (Murinae, Muridae), Bulletin of the American Museum of Natural 

History, 211:1–138.

Musser, G. G., J. T. Marshall Jr., and Boeadi. 1979. Definition and Contents of 

the Sundaic Genus Maxomys (Rodentia, Muridae). Journal of Mammalogy, 

60:592–606. http://dx.doi.org/10.2307/1380101.

Musser, G. G., and C. Newcomb. 1983. Malaysian Murids and the Giant Rat of Sumatra. Bulletin of the American Museum of Natural History, 174:327–598.

———. 1985. Definitions of Indochinese Rattus losea and a New Species from 

Vietnam. American Museum Novitates, 2814:1–32.

Nagy, T. R., and B. A. Gower. 1999. “Northern Collared Lemming, Dicrostonyx 

groenlandicus.” In The Smithsonian Book of North American Mammals, ed. 
104 • smithsonian contributions to zoology

D. E. Wilson and S. Ruff, pp. 659–660. Washington, D.C.: Smithsonian Institution Press.

Ndiaye, A., K. Bâ, V. Aniskin, T. Benazzou, P. Chevret, A. Konečný, M. Sembène, 

C. Tatard, G. J. Kergoat, and L. Granjon. 2012. Evolutionary Systematics 

and Biogeography of Endemic Gerbils (Rodentia, Muridae) from Morocco: 

An Integrative Approach. Zoologica Scripta, 41:11–28. http://dx.doi.org/10

.1111/j.1463-6409.2011.00501.x.

Nelson, E. W. 1893. Description of a New Species of Arvicola, of the Mynomes

Group, from Alaska. Proceedings of the Biological Society of Washington, 

8:139–142.

———.1929. Description of a New Lemming from Alaska. Proceedings of the Biological Society of Washington, 42:143–146.

———. 1931. A New Vole of the Subgenus Stenocranius from Alaska. Journal of 

Mammalogy, 12:310–312. http://dx.doi.org/10.2307/1373885.

Nelson, E. W., and E. A. Goldman. 1931a. Two New Woodrats from Lower California. Proceedings of the Biological Society of Washington, 44:107–110.

———. 1931b. New Carnivores and Rodents from Mexico. Journal of Mammalogy, 12:302–306. http://dx.doi.org/10.2307/1373882.

———. 1931c. Six New White-Footed Mice (Peromyscus maniculatus Group) 

from Islands off the Pacific Coast. Journal of the Washington Academy of 

Sciences, 21:530–535.

———. 1932. The Name of the San Martin Island Mouse—A Correction. Journal 

of Mammalogy, 13:370–371.

———. 1933. Three New Rodents from Southern Mexico. Proceedings of the Biological Society of Washington, 46:195–198.

Nicolas, V., S. Querouil, E. Verheyen, W. Verheyen, J. F. Mboumba, M. Dillem, 

and M. Colyn. 2006. Mitochondrial Phylogeny of African Wood Mice, Genus Hylomyscus (Rodentia, Muridae): Implications for Their Taxonomy and 

Biogeography. Molecular Phylogenetics and Evolution, 38:779–793. http://

dx.doi.org/10.1016/j.ympev.2005.11.024.

Ognev, S. I. 1948. Zveri SSSR i prilezhaschashchikh stran: Gryzuny (prodolzhenie). (Zveri vostochnoi Evropy i severonoi Azii) [Mammals of the USSR and 

adjacent countries: Rodents (continued). (Mammals of eastern Europe and 

northern Asia)]. Akademiya Nauk SSSR, 6:1–559.

———. 1964. Mammals of the USSR and Adjacent Countries: Rodents (continued. (Mammals of Eastern Europe and Northern Asia) [A translation og S. I. 

Ognev, 1950, Zveri SSSR i prilezhashchikh stran: Gryzuny (prodolzhenie), 

(Zverivostochnoi Evropy I severnoi Azli]. Israel Program for Scientific Translations, Jerusalem, 7L1—626.

Osgood, W. H. 1900. Results of a Biological Reconnoissance of the Yukon River 

Region. North American Fauna, 19:1–45.

———. 1901a. Natural History of the Cook Inlet Region, Alaska. North American 

Fauna, 21:51–87.

———. 1901b. A New White-Footed Mouse from California. Proceedings of the 

Biological Society of Washington, 14:193–194.

———. 1901c. Natural History of the Queen Charlotte Islands, British Columbia. 

North American Fauna, 21:1–50. http://dx.doi.org/10.3996/nafa.21.0001.

———.1904a. Thirty New Mice of the Genus Peromyscus from Mexico and Guatemala. Proceedings of the Biological Society of Washington, 17:55–77.

———. 1904b. A Biological Reconnaissance of the Base of the Alaska Peninsula. 

North American Fauna, 24:1–86. http://dx.doi.org/10.3996/nafa.24.0001.

———. 1905. A New Name for the Peromyscus nebracensis of Certain Authors. 

Proceedings of the Biological Society of Washington, 18:77.

———. 1906. A New Vole from Montague Island, Alaska. Proceedings of the Biological Society of Washington, 19:71–72.

———. 1907a. Some Unrecognized and Misapplied Names of American Mammals. Proceedings of the Biological Society of Washington, 20:43–52.

———. 1907b. A Collection of Mammals from the Region of Mount McKinley, 

Alaska. Proceedings of the Biological Society of Washington, 20:59–64.

———. 1908. A New White-Footed Mouse from Alaska. Proceedings of the Biological Society of Washington, 21:141–142.

———. 1909a. Revision of the Mice of the American Genus Peromyscus. North 

American Fauna, 28:1–285. http://dx.doi.org/10.3996/nafa.28.0001.

———. 1909b. Biological Investigations in Alaska and Yukon Territory. North 

American Fauna, 30:1–96. http://dx.doi.org/10.3996/nafa.30.0001.

———. 1925. The Long-Clawed South American Rodents of the Genus Notiomys. 

Field Museum of Natural History, Zoological Series, 12:113–125.

———. 1943. The Mammals of Chile. Field Museum of Natural History, Zoological Series, 30:1–268.

Packard, R. L. 1958a. New Subspecies of the Rodent Baiomys from Central 

America. University of Kansas Publications, Museum of Natural History, 

9:397–404.

———. 1958b. The Taxonomic Status of Peromyscus allex Osgood. Proceedings of 

the Biological Society of Washington, 71:17–20.

———. 1969. Taxonomic Review of the Golden Mouse, Ochrotomys nuttalli. Miscellaneous Publications, Museum of Natural History, University of Kansas, 

51:373–406.

Paradiso, J. L. 1960. A New White-Footed Mouse (Peromyscus leucopus) from 

Southeastern Virginia. Proceedings of the Biological Society of Washington, 

73:21–24.

Paradiso, J. L., and R. H. Manville. 1961. Taxonomic Notes on the Tundra Vole 

(Microtus oeconomus) in Alaska. Proceedings of the Biological Society of 

Washington, 74:77–92.

Pardinas, U. F. J., and C. A. Galliari. 2001. Reithrodon auritus. Mammalian Species, 

664:1–8. http://dx.doi.org/10.1644/1545-1410(2001)664<0001:RA>2.0.CO;2.

Patton, J. L., D. G. Huckaby, and S. T. Alvarez-Castaneda. 2008 [2007] . The 

Evolutionary History and a Systematic Revision of Woodrats of the Neotoma 

lepida Group. University of California Publications in Zoology, 135:1–411.

Pavlinov, I. Ya, Yu. A. Dubrovsky, O. L. Rossolimo, and E. G. Potapova. 1990. 

Gerbils of the World. [In Russian.] Moscow: Nauka.

Pavlinov, I. Ya., and O. L. Rossolimo. 1987. Sistematica Miekopitayushchikh USSR 

[Systematics of the mammals of the USSR]. Moscow: Moscow University 

Press.

Pavlinov, I. Ya., E. L. Yakhontov, and A. K. Agadzhanyan. 1995. Mammals of Eurasia. I. Rodentia. Taxonomic and Geographic Guide. Archives of the Zoological Museum, Moscow State University, 32:1–236.

Peale, T. R. 1848. United States Exploring Expedition during the Years 1838, 1839, 

1840, 1841, 1842 under the Command of Charles Wilkes, U.S.N. Volume 8, 

Mammalia and Ornithology. Philadelphia: C. Sherman.

Pearson, O. P., and M. I. Christie. 1991. Sympatric Species of Euneomys (Rodentia, 

Cricetidae). Studies on Neotropical Fauna and Environment, 26:121–127.

Petter, F. 1978. Une Souris Nouvelle Du Sud De L’Afrique: Mus setzeri sp. nov. 

Mammalia, 42:377–379.

Phillips, C. J. 1969. Review of Central Asian Voles of the Genus Hyperacrius, with 

Comments on Zoogeography, Ecology, and Ectoparasites. Journal of Mammalogy, 50:457–474. http://dx.doi.org/10.2307/1378773.

Pine, R. H. 1973 [1974]. Una nueve Especies de Akodon (Mammalia: Rodentia: 

Muridae) de la Isla Wellington, Magallanes, Chile. Anales del Instituto de la 

Patagonia, 4:423–426.

———. 1976. A New Species of Akodon (Mammalia: Rodentia: Muridae: Cricetinae) from Isla De Los Estados, Argentina. Mammalia, 40:63–68. http://

dx.doi.org/10.1515/mamm.1976.40.1.63.

Poole, A. J., and V. S. Schantz. 1942. Catalog of the Type Specimens of Mammals 

in the United States National Museum, Including the Biological Surveys Collection. Bulletin of the United States National Museum, 178:1–705. http://

dx.doi.org/10.5479/si.03629236.178.1.

Post, D. 1999. “Eastern Woodrat, Neotoma floridana.” In The Smithsonian Book 

of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 601–603. 

Washington, D.C.: Smithsonian Institution Press.

Preble, E. A. 1899a. Revision of the Jumping Mice of the Genus Zapus. North 

American Fauna, 15:1–42.

———. 1899b. Description of a New Lemming Mouse from the White Mountains, New Hampshire. Proceedings of the Biological Society of Washington, 

13:43–45.

———. 1902a. A Biological Investigation of the Hudson Bay Region. North American Fauna, 22:1–140.

———. 1902b. Descriptions of New Species of Synaptomys and Phenacomys from 

Mackenzie, Canada. Proceedings of the Biological Society of Washington, 

15:181–182.

———. 1908. Biological Investigations of the Athabaska-Mackenzie Region. 

North American Fauna, 27:1–574.

Ranck, G. L. 1968. Rodents of Libya: Taxonomy, Ecology, and Zoogeographical 

Relationships. Bulletin of the United States National Museum, 275:1–264. 

http://dx.doi.org/10.5479/si.03629236.275.1.

Raun, G. G., and H. E. Laughlin. 1972. Sub-Recent Vertebrate Remains from a 

Site in Southern Texas with Comments on Microtus (Pedomys) ludovicianus. 

Southwestern Naturalist, 16:436–439. http://dx.doi.org/10.2307/3670079.

Rausch, R. 1950. Notes on Microtine Rodents from the Brooks Range, Arctic 

Alaska. Journal of the Washington Academy of Sciences, 40:133–136.

———. 1953. On the Status of Some Arctic Mammals. Journal of the Arctic Institute of North America, 6:91–148.

Reig, O. A. 1987. An Assessment of the Systematics and Evolution of the Akodontini, with the Description of New Fossil Species of Akodon (Cricetidae: Sigmodontinae). Fieldiana: Zoology, n.s., 39:347–399.
number 642 • 105

Reynolds, R. P., S. W. Gotte, and C. H. Ernst. 2007. Catalog of Type Specimens 

of Recent Crocodilia and Testudines in the National Museum of Natural 

History, Smithsonian Institution. Smithsonian Contributions to Zoology, 

No. 626. Washington, D.C.: Smithsonian Institution Scholarly Press. http://

dx.doi.org/10.5479/si.00810282.626.

Rhoads, S. N., and R. T. Young. 1897. Notes on a Collection of Small Mammals 

from Northeastern North Carolina. Proceedings of the Academy of Natural 

Sciences of Philadelphia, 49:303–312.

Rickart, E. A., and L. R. Heaney. 1991. A New Species of Chrotomys (Rodentia: 

Muridae) from Luzon Island, Philippines. Proceedings of the Biological Society of Washington, 104:387–398.

Riddle, B. R. 1999a. “Southern Grasshopper Mouse, Onychomys torridus.” In 

The Smithsonian Book of North American Mammals, ed. D. E. Wilson and 

S. Ruff, pp. 590–591. Washington, D.C.: Smithsonian Institution Press.

———. 1999b. “Northern Grasshopper Mouse. Onychomys leucogaster.” In 

The Smithsonian Book of North American Mammals, ed. D. E. Wilson and 

S. Ruff, pp. 588–590. Washington, D.C.: Smithsonian Institution Press.

———. 1999c. “Mearns’ Grasshopper Mouse, Onychomys arenicola.” In The 

Smithsonian Book of North American Mammals, ed. D. E. Wilson and 

S. Ruff, p. 588. Washington, D.C.: Smithsonian Institution Press.

Riddle, B. R., and J. R. Choate. 1986. Systematics and Biogeography of Onychomys leucogaster in Western North America. Journal of Mammalogy, 67:233–

255. http://dx.doi.org/10.2307/1380877.

Ride, W. D. L. 1970. A Guide to the Native Mammals of Australia. Melbourne: 

Oxford University Press.

Robbins, C. B. 1974. Comments on the Taxonomy of the West African Taterillus

(Rodentia: Cricetidae) with the Description of a New Species. Proceedings of 

the Biological Society of Washington, 87:395–404.

Robinson, W., and M. W. Lyon Jr. 1901. An Annotated List of Mammals Collected 

in the Vicinity of La Guaira, Venezuela. Proceedings of the United States National Museum, 24:135–162. http://dx.doi.org/10.5479/si.00963801.1246

.135.

Rodrigues-Serrano, E., C. E. Hernandez, and R. E. Palma. 2008. A New Record 

and an Evaluation of the Phylogenetic Relationships of Abrothrix olivaceus 

markhami (Rodentia: Sigmodontinae). Mammalian Biology, 73:309–317. 

http://dx.doi.org/10.1016/j.mambio.2007.10.003.

Rose, R. K. 2011. Range Extension for the Dismal Swamp Southern Bog Lemming, 

Synaptomys cooperi helaletes, in Virginia. Banisteria, 38:60–64.

Rose, R. K., and R. D. Dueser. 1999. “Marsh Rice Rat, Oryzomys palustris.” In 

The Smithsonian Book of North American Mammals, ed. D. E. Wilson and 

S. Ruff, pp. 554–555. Washington, D.C.: Smithsonian Institution Press.

Rossolimo, O. L. 1989. Revision of Royle’s High-Mountain Vole Alticola (A.)

argentatus (Mammalia: Cricetidae). [In Russian.] Zoologiecheskii Zhurnal, 

68(8):104–114.

Rossolimo, O. L., and I. Pavlinov. 1992. “Species and Subspecies of Alticola s. str. 

(Rodentia: Arvicolidae).” In Prague Studies in Mammalogy, ed. J. Horacek 

and V. Vohralik, pp. 149–176. Prague: Charles University Press.

Schlitter, D. A. 1973. A New Species of Gerbil from South West Africa with Remarks on Gerbillus tytonis Bauer and Niethammer, 1959 (Rodentia: Gerbillinae). Bulletin of the Southern California Academy of Science, 72:13–18.

Schlitter, D. A., and H. W. Setzer. 1972. A New Species of Short-Tailed Gerbil (Dipodillus) from Morocco (Mammalia: Cricetidae: Gerbillinae). Proceedings of 

the Biological Society of Washington, 84:385–392.

———. 1973. New Rodents (Mammalia: Cricetidae, Muridae) from Iran and Pakistan. Proceedings of the Biological Society of Washington, 86:163–174.

Schmidly, D. J. 1972. Geographic Variation in the White-Ankled Mouse, Peromyscus pectoralis. The Southwestern Naturalist, 17:113–138. http://dx.doi

.org/10.2307/3670366.

———. 1999a. “White-Ankled Mouse, Peromyscus pectoralis.” In The Smithsonian Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 

579–580. Washington, D.C.: Smithsonian Institution Press.

———. 1999b. “Eastern Harvest Mouse, Reithrodontomys humulis.” In The 

Smithsonian Book of North American Mammals, ed. D. E. Wilson and 

S. Ruff, pp. 557–558. Washington, D.C.: Smithsonian Institution Press.

Schmidt, D. F., C. A. Ludwig, and M. D. Carleton. 2008. The Smithsonian Institution African Mammal Project (1961–1972): An Annotated Gazetteer 

of Collecting Localities and Summary of Its Taxonomic and Geographic 

Scope. Smithsonian Contributions To Zoology, No. 628. Washington, 

D.C.: Smithsonian Institution Scholarly Press. http://dx.doi.org/10.5479/si.

00810282.628.

Schwartz, A. A. 1953. Systematic Study of the Water Rat (Neofiber alleni). Occasional Papers, Museum of Zoology, University of Michigan, 547:1–27.

Schwarz, E. 1939a. On Mountain Voles of the Genus Alticola Blanford: A Taxonomic and Genetic Analysis. Proceedings of the Zoological Society of London, 108:663–668.

———. 1939b. A New Porcupine (Thecurus) from Borneo. Journal of Mammalogy, 20:246–248. http://dx.doi.org/10.2307/1374385.

Schwarz, E., and H. K. Schwarz. 1943. The Wild and Commensal Stocks of the 

House Mouse, Mus musculus Linnaeus. Journal of Mammalogy, 24:59–72. 

http://dx.doi.org/10.2307/1374781.

———. 1967. A Monograph of the Rattus rattus Group. Anales de la Escuela 

Nacional Ciencias Biologicas Mexico, 14:79–178.

Sera, W. E., and C. N. Early. 2003. Microtus montanus. Mammalian Species, 

716:1–10. http://dx.doi.org/10.1644/716.

Setzer, H. W. 1952. A New Subspecies of Microtus oeconomus from Alaska. Proceedings of the Biological Society of Washington, 65:75–76.

———. 1956a. Two New Gerbils from Libya. Proceedings of the Biological Society 

of Washington, 69:179–182.

———. 1956b. A New Jird from Libya. Proceedings of the Biological Society of 

Washington, 69:205–206.

Setzer, H. W., and G. L. Ranck. 1971. A New Gerbil (Genus Gerbillus) from the 

Chad. Proceedings of the Biological Society of Washington, 84:55–58.

Shamel, H. H. 1931. Akodon chacoensis, a New Cricetine Rodent from Argentina. 

Journal of the Washington Academy of Sciences, 21:427–429.

———. 1940. Three New Mammals from Asia. Journal of Mammalogy, 21:76–78.

Shenbrot, G. I. 1993 [1991]. Revision of Subspecies Systematics of Jerboas of the 

Genus Allactaga of the USSR Fauna. [In Russian with English abstract.] 

Trudy Zoologicheskovo Instituta, Academiya Nauk SSSR, 243:81–109. 

Shipp-Pennock, M. A., W. D. Webster, and D. W. Freshwater. 2005. Systematics 

of the White-Footed Mouse (Peromyscus leucopus) in the Mid-Atlantic Region. Journal of Mammalogy, 86:803–813. http://dx.doi.org/10.1644/1545

-1542(2005)086[0803:SOTWMP]2.0.CO;2.

Shufeldt, R. W. 1885. Description of Hesperomys truei, a New Species Belonging to 

the Subfamily Murinae. Proceedings of the United States National Museum, 

8:403–408. http://dx.doi.org/10.5479/si.00963801.8-529.403.

Shump, K. A., Jr. 1999. “Tawny-Bellied Cotton Rat, Sigmodon fulviventer.” In 

The Smithsonian Book of North American Mammals, ed. D. E. Wilson and 

S. Ruff, pp. 592–593. Washington, D.C.: Smithsonian Institution Press.

Smith, H. D. 1999. “Sagebrush Vole, Lemmiscus curtatus.” In The Smithsonian 

Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 649–

650. Washington, D.C.: Smithsonian Institution Press.

Smith, M. F. 1979. Geographic Variation in Genic and Morphological Characters in Peromyscus californicus. Journal of Mammalogy, 60:705–722. http://

dx.doi.org/10.2307/1380187.

Smolen, M. J. 1999a. “Long-Tailed Vole, Microtus longicaudus.” In The Smithsonian Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 

630–632. Washington, D.C.: Smithsonian Institution Press.

———. 1999b. “Woodland Vole, Microtus pinetorum.” In The Smithsonian Book 

of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 642–644. 

Washington, D.C.: Smithsonian Institution Press.

Spitzer, N. C., and J. D. Lazell, Jr. 1978. A New Rice Rat (Genus Oryzomys) from 

Florida’s Lower Keys. Journal of Mammalogy, 59:787–792. http://dx.doi.org

/10.2307/1380143.

Spotorno, A. E., and L. I. Walker. 1983. Analisis el Electroforetico y Biometrico 

de dos Especies de Phyllotis En Chile Central. Revista Chilena de Historia 

Natural, 56:51–59.

Stalling, D. T. 1999. “Prairie Vole, Microtus ochrogaster.” In The Smithsonian 

Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 637–

638. Washington, D.C.: Smithsonian Institution Press.

Steppan, S. J., O. Ramirez, J. Banbury, D. Huchon, V. Pacheco, L. I. Walker, and 

A. E. Spotorno. 2007. A Molecular Reappraisal of the Leaf-Eared Mice Phyllotis and Their Relatives. University of California Publications in Zoology, 

134:799–826.

Sumner, F. B. 1926. An Analysis of Geographic Variation in Mice of the Peromyscus polionotus Group from Florida and Alabama. Journal of Mammalogy, 

7:149–184. http://dx.doi.org/10.2307/1373508.

Swanepoel, P., and D. A. Schlitter. 1978. Taxonomic Review of the Fat Mice (Genus 

Steatomys) of West Africa (Mammalia: Rodentia). Bulletin of the Carnegie 

Museum of Natural History, 6:53–76.

Tamarin, R. H. 1999. “Meadow Vole, Microtus pennsylvanicus.” In The Smithsonian Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 

641–642. Washington, D.C.: Smithsonian Institution Press. 

Tate, G. H. H. 1935. Rodents of the Genera Rattus and Mus from the Pacific 

Islands. Bulletin of the American Museum of Natural History, 68:145–178.
106 • smithsonian contributions to zoology

Taylor, W. P. 1920. A New Meadow Vole from the Cascade Mountains of Washington. Journal of Mammalogy, 1:180–182.

Taylor, Z. S., and S. M. G. Hoffman. 2010. Mitochondrial DNA Genetic Structure 

Transcends Natural Boundaries in Great Lakes Populations of Woodland 

Deer Mice (Peromyscus maniculatus gracilis). Canadian Journal of Zoology, 

88:404–415. http://dx.doi.org/10.1139/Z10-010.

Thomas, O. 1894. Descriptions of Some New Neotropical Muridae. Annals and 

Magazine of Natural History, 6th ser., 14:346–360.

———. 1897. On the Mammals of Nyasaland: Fourth Notice. Proceedings of 

the Zoological Society of London, 64:788–798. http://dx.doi.org/10.1111

/j.1096-3642.1896.tb03079.x.

———. 1901. A New Spiny Rat from La Guaira, Venezuela. Proceedings of the 

Biological Society of Washington, 14:27–28.

———. 1916. Three New African Mice of the Genus Dendromus. Annals and 

Magazine of Natural History, 8th ser., 18:241–243.

———. 1917. Preliminary Diagnoses of New Mammals Obtained by the YaleNational Geographic Society Peruvian Expedition. Smithsonian Miscellaneous Collections, 68(4):1–3.

———. 1920a. Report on the Mammalia Collected by Mr. Edmund Heller during 

the Peruvian Expedition of 1915 under the Auspices of Yale University and 

the National Geographic Society. Proceedings of the United States National 

Museum, 58:217–249. http://dx.doi.org/10.5479/si.00963801.58-2333.217.

———. 1920b. The Generic Positions of “Mus” nigricauda, Thos., and woosnami, 

Schwann. Annals and Magazine of Natural History, 9th ser., 5:140–142.

———. 1926. The Generic Position of Certain African Muridae, Hitherto Referred 

to Aethomys and Praomys. Annals and Magazine of Natural History, 9th 

ser., 17:174–179.

———. 1927. The Godman-Thomas Expedition to Peru. V. On Mammals Collected by M. R. W. Hendee in the Province of San Martin, N. Peru, Mostly at 

Yurac Yacu. Annales and Magazine of Natural History, 9th ser., 19:361–375.

Townsend, C. H. 1912. Mammals Collected by the ‘Albatross’ Expedition in Lower 

California in 1911, with Descriptions of New Species. Bulletin of the American Museum of Natural History, 31:117–130.

Tribe, C. J. 1996. The Neotropical Rodent Genus Rhipidomys (Cricetidae: 

Sigmodontinae)—A Taxonomic Revision. Ph.D. diss., University College, 

London.

Trouessart, E. L. 1897. Catalogus Mammalium tam Viventium quam Fossilium. 

Berlin: R. Friedlander.

True, F. W. 1884. A Muskrat with a Round Tail. Science, 4:34. http://dx.doi.org

/10.1126/science.ns-4.75.34.

———. 1889. On the Mammals Collected in Eastern Honduras in 1887 by Mr. 

Charles H. Townsend, with a Description of a New Subspecies of Capromys

from Little Swan Island. Proceedings of the United States National Museum, 

11:469–472. http://dx.doi.org/10.5479/si.00963801.744.469.

———. 1890. Description of a New Species of Mouse, Phenacomys longicaudus, 

from Oregon. Proceedings of the United States National Museum, 13:303–

304. http://dx.doi.org/10.5479/si.00963801.826.303.

———. 1892. An Annotated Catalog of the Mammals Collected by Dr. W. L. Abbott in the Kilima-Njaro Region, East Africa. Proceedings of the United States 

National Museum, 15:445–480. http://dx.doi.org/10.5479/si.00963801.15

-915.445.

———. 1893. Notes on a Small Collection of Mammals from the Tana River, East 

Africa, with Descriptions of New Species. Proceedings of the United States 

National Museum, 16:601–603. http://dx.doi.org/10.5479/si.00963801.954

.601.

———. 1894a. On the Rodents of the Genus Sminthus in Kashmir. Proceedings of 

the United States National Museum, 17:341–343. http://dx.doi.org/10.5479

/si.00963801.1004.351.

———. 1894b. Notes on Mammals of Baltistan and the Vale of Kashmir, Presented 

to the National Museum by Dr. W. L. Abbott. Proceedings of the United States 

National Museum, 17:1–16. http://dx.doi.org/10.5479/si.00963801.976.

———. 1894c. Diagnosis of New North American Mammals. Proceedings of the 

United States National Museum, 17:1–3. Preprints. 

———. 1894d. Diagnosis of New North American Mammals. Proceedings of the 

United States National Museum, 17:241–243. http://dx.doi.org/10.5479/si

.00963801.999.241.

———. 1894e. Diagnoses of Some Undescribed Wood Rats (Genus Neotoma) in 

the National Museum. Proceedings of the United States National Museum, 

17:1–3. Preprints.

———. 1894f. Diagnoses of Some Undescribed Wood Rats (Genus Neotoma) in 

the National Museum. Proceedings of the United States National Museum, 

17:353–355. http://dx.doi.org/10.5479/si.00963801.1006.353.

———. 1894g. Description of a New Species of Mouse (Sitomys decolorus) 

from Central America. Proceedings of the United States National Museum, 

16:689–690. http://dx.doi.org/10.5479/si.00963801.963.689.

Trujano-Alvarez, A. L., and S. T. Alvarez-Castaneda. 2010. Peromyscus mexicanus (Rodentia: Cricetidae). Mammalian Species, 42:111–118. http://dx.doi

.org/10.1644/858.1.

Van Cura, N. J., and D. F. Hoffmeister. 1966. A Taxonomic Review of the Grasshopper Mice, Onychomys, in Arizona. Journal of Mammalogy, 47:613–630. 

http://dx.doi.org/10.2307/1377892.

Van der Straten, E., and W. N. Verheyen. 1981. Etude biometrique du genre Praomys en Cote d’Ivoire. Bonner Zoologische Beitrage, 32:249–264.

Van Weers, D. J. 1976. Notes on Southeast Asian Porcupines (Hystricidae, Rodentia) I. On the Taxonomy of the Genus Trichys Gunther, 1877. Beaufortia, 

25:15–31.

———. 1978. Notes on Southeast Asian Porcupines (Hystricidae, Rodentia) III. 

On the Taxonomy of the Subgenus Thecurus Lyon, 1907 (Genus Hystrix

Linnaeus, 1758). Beaufortia, 28:17–33.

Vari, R. P., and J. C. Howe. 1991. Catalog of the Type Specimens of Recent Fishes 

in the National Museum of Natural History, Smithsonian Institution. 1: 

Characiformes (Teleostei: Ostariophysi). Smithsonian Contributions to Zoology, No. 517. Washington, D.C.: Smithsonian Institution Scholarly Press. 

http://dx.doi.org/10.5479/si.00810282.517.

Vazquez, L. B., G. N. Cameron, and R. A. Medellin. 2001. Peromyscus aztecus. 

Mammalian Species, 649:1–4.

Vega, R. E., E. Vazquez-Dominguez, A. Mejia-Puente, and A. D. Cuaron. 2007. 

Unexpected High Levels of Genetic Variability and Population Structure of 

an Island Endemic Rodent (Oryzomys couesi cozumelae). Biological Conservation, 137:210–222. http://dx.doi.org/10.1016/j.biocon.2007.02.007.

Verheyen, W. N. 1965. Contribution a l’Etude, Systematique de Mus sorella

(Thomas, 1909). Revue de Zoologie et de Botanique Africanes, 71:194–212.

von Bohmann, L. 1942. Die Gattung Dendromus A. Smith. Versuch einer 

naturlichen Gruppierung (Ergebnisse der Ostafrika-Reise 1937 UthmollerBohmann. VIII). Zoologischer Anzeiger, 193:33–53.

von den Brink, F. H. 1930. Zur Nomenclatur Zweier Europalacher Microtus-Forman. Zeitschrift für Saugetierkunde, 4:251.

Voss, R. S. 1988. Systematics and Ecology of Ichthyomine Rodents (Muroidea): 

Patterns of Morphological Evolution in a Small Adaptive Radiation. Bulletin 

of the American Museum of Natural History, 188:259–493.

———. 1991. An Introduction to the Neotropical Muroid Rodent Genus Zygodontomys. Bulletin of the American Museum of Natural History, 210:1–113.

———. 2011. Revisionary Notes on Neotropical Porcupines (Rodentia: Erethizontidae) 3. An Annotated Checklist of the Species of Coendou Lacepede, 1799. 

American Museum Novitates, 3720:1–36.

Wallace, A. M., and C. Jones. 1999. “Stephen’s Woodrat, Neotoma stephensi.” In 

The Smithsonian Book of North American Mammals, ed. D. E. Wilson and 

S. Ruff, p. 611. Washington, D.C.: Smithsonian Institution Press.

Webster, W. D. 1999. “Western Harvest Mouse, Reithrodontomys megalotis.” In 

The Smithsonian Book of North American Mammals, ed. D. E. Wilson and 

S. Ruff, pp. 558–560. Washington, D.C.: Smithsonian Institution Press.

Weksler, M., A. R. Percequillo, and R. S. Voss. 2006. Ten New Genera of Oryzomine Rodents (Cricetidae: Sigmodontinae). American Museum Novitates, 

3537:1–29.

West, S. D. 1999. “Northern Bog Lemming, Synaptomys borealis.” In The Smithsonian Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 

655–656. Washington, D.C.: Smithsonian Institution Press.

Whitaker, J. O., Jr. 1999a. “Woodland Jumping Mouse, Napaeozapus insignis.” 

In The Smithsonian Book of North American Mammals, ed. D. E. Wilson 

and S. Ruff, pp. 665–666. Washington, D.C.: Smithsonian Institution Press.

———. 1999b. “Meadow Jumping Mouse, Zapus hudsonius.” In The Smithsonian Book of North American Mammals, ed. D. E. Wilson and S. Ruff, pp. 

666–667. Washington, D.C.: Smithsonian Institution Press.

Whitaker, J. O., Jr., and W. J. Hamilton, Jr. 1998. Mammals of the Eastern United 

States. 3rd ed. Ithaca, N.Y.: Cornell University Press.

Wilson, D. E., and D. M. Reeder, eds. 2005. Mammal Species of the World: A 

Taxonomic and Geographic Reference. Volume 2. 3rd ed. Baltimore, Md.: 

Johns Hopkins University Press.

Wolff, J. O. 1999. “Creeping Vole, Microtus oregoni.” In The Smithsonian Book of 

North American Mammals, ed. D. E. Wilson and S. Ruff, p. 640. Washington, D.C.: Smithsonian Institution Press.

Won, C., and K. G. Smith. 1999. History and Current Status of Mammals of the 

Korean Peninsula. Mammal Review, 29:3–33. http://dx.doi.org/10.1046/

j.1365-2907.1999.00034.x.
number 642 • 107

Woodhouse, S. W. 1853. Description of a New Species of Mouse, of the Genus 

Hesperomys, Waterhouse. Proceedings of the Academy of Natural Sciences 

of Philadelphia, 6:242.

Woods, C. A., and C. W. Kilpatrick. 2005. “Infraorder Hystrichognathi.” In Mammal Species of the World: A Taxonomic and Geographic Reference, Volume 

2, 3rd ed., ed. D. E. Wilson and D. M. Reeder, pp. 1538–1600. Baltimore, 

Md.: Johns Hopkins University Press.

Yancey, F. D., and C. Jones. 1999. “Pinyon Mouse, Peromyscus truei.” In The 

Smithsonian Book of North American Mammals, ed. D. E. Wilson and 

S. Ruff, pp. 582–583. Washington, D.C.: Smithsonian Institution Press.

Young, C. J., and J. K. Jones, Jr. 1983. Peromyscus yucatanicus. Mammalian Species, 196:1–3. http://dx.doi.org/10.2307/3503978.

Zheng, S. 1994. “Classification and Evolution of the Siphneidae.” In Rodent and 

Lagomorph Families of Asian Origin and Diversification, ed. Y. Tomida, 

C. K. Li, and T. Setoguchi. National Science Museum Monographs, 8:57–76.

Zhou, C., and K. Zhou. 2008. The Validity of Different Zokor Species and the 

Genus Eospalax Inferred from Mitochondrial Gene Sequences. Integrative 

Zoology, 3:290–298.

Zimmermann, K. 1942. Zur Kenntnis von Microtus oeconomus (Pallas). Archive 

fur Naturgeschichte, 11:174–197.

Index of Taxa

abbreviatus

Microtus, 18

Abrothrix

jelskii, 8

markhami, 8

olivaceus, 8

abyssinicus

Arvicanthis, 66

acholi

Mus, 78

acmaeus

Alticola, 8

Acomys

cineraceus, 64

hystrella, 64

ignites, 64

kempi, 64

montanus, 64

acrophilus

Microtus, 9

adocetus

Microtus, 21

adspersus

Rattus, 67

adustus

Tryphomys, 90

aedium

Plagiodontia, 94

aeruginosus

Gerbillus, 70

Aethomys

hindei, 64

kaiseri, 64

affinis

Hesperomys, 41

Peromyscus, 41, 50

Agouti

nelsoni, 92

paca, 92

agrarius

Apodemus, 64, 65
110 • smithsonian contributions to zoology

Akodon

chacoensis, 10

llanoi, 8

markhami, 8

surdus, 8

torques, 8

alascensis

Evotomys, 23

Lemmus, 13

Zapus, 5

alashanicus

Euchoreutes, 3

albescens

Onychomys, 36

Reithrodontomys, 55

albicauda

Arvicola, 8

albicaudus

Alticola, 8

albigula

Neotoma, 27, 29, 34

albigularis

Mus, 83

albilabris

Reithrodontomys, 58

albipes

Arborimus, 9

Phenacomys, 9

albiventer

Melomys, 77

Oryzomys, 38

albus

Ondatra, 35

aldousi

Onychomys, 36

alfari

Nectomys, 62

Sigmodontomys, 62

alfaroi

Oryzomys, 38, 40

alfredi

Sigmodon, 60

algidus

Peromyscus, 47

Allactaga

grisescens, 3

longior, 3

mongolica, 3

sibirica, 3

alleni

Epimys, 71

Hodomys, 11

Neofiber, 26

Neotoma, 11

Notiomys, 10

Reithrodontomys, 55

Sigmodon, 59, 60

allex

Baiomys, 9

Peromyscus, 9

allophylus

Peromyscus, 43

alstoni

Neotomodon, 34

alticola

Microtus, 14

Sigmodon, 59

Alticola

acmaeus, 8

albicaudus, 8

argentatus, 8

glacialis, 8

lemminus, 9

montosus, 8

parvidens, 8

roylei, 8

stoliczkanus, 9

alticolus

Arvicola, 14

Reithrodontomys, 58

altilaneus

Peromyscus, 43

amakensis

Microtus, 18

ambersoni

Rattus, 87

ambiguus

Peromyscus, 44

amoles

Reithrodontomys, 55

Sigmodon, 59

amplus

Meriones, 77

Peromyscus, 44

analogus

Baiomys, 9

Peromyscus, 9

anambae

Mus, 74

andamanensis

Rattus, 83, 84, 87

andersoni

Gerbillus, 69, 70

andrewsi

Bunomys, 67

Mus, 67

angelensis

Peromyscus, 44

angusticeps

Microtus, 18

Neotoma, 29

Oryzomys, 38

annandalei

Rattus, 83

ANOMALURIDAE, 90

ANOMALUROMORPHA, 90

ansorgei

Cricetomys, 6, 7

Lophuromys, 73

anthonyi

Hesperomys, 41

Peromyscus, 41

antucus

Rattus, 76

apalachicolae

Neofiber, 26

apella

Arvicola, 14

aphorodemus

Microtus, 18

apicalis

Nectomys, 26

apicis

Mus, 86

Apodemus

agrarius, 64, 65

chejuensis, 64

microtis, 65

nigritalus, 65

pallescens, 65

peninsulae, 65

praetor, 65

rusiges, 65

sowerbyi, 65

uralensis, 65

apoensis

Tarsomys, 89

Apomys

bardus, 65

datae, 66

hylocoetes, 65, 66

insignis, 65

major, 66

microdon, 66

musculus, 66

petraeus, 66

sacobianus, 66

aquilo

Notomys, 79

aquilus

Gerbillus, 69, 70

Lophuromys, 73

Mus, 73

Arborimus

albipes, 9

longicaudus, 9

arcticeps

Onychomys, 36

arenaceous

Jaculus, 4

arenarius

Peromyscus, 45

Phyllotis, 53

Taterillus, 89

arenicola

Onychomys, 37, 38

arenicolor

Gerbillus, 66

argentatus

Alticola, 8

Oryzomys, 38

argenteocinereus

Heliophobius, 90

argentiventer

Rattus, 87

arianus

Mus, 65

aridulus

Peromyscus, 47

arizonae

Neotoma, 27

Sigmodon, 60, 61

arizonensis

Clethrionomys, 23

Microtus, 20

Myodes, 23

articus [sic]

Hesperomys, 41

arvalis

Microtus, 18

Arvicanthis

abyssinicus, 66

centrosus, 66

jebelae, 66

micropus, 71

nairobae, 66

niloticus, 66

planifrons, 70

pulchellus, 71

virescens, 66

Arvicola

albicauda, 8

alticolus, 14

apella, 14

austerus, 14, 15

breweri, 15

californica, 15

cinnamomea, 15

curtata, 12

edax, 15

fertilis, 12

haydeni, 15

kennicottii, 16

longicaudus, 16

longipilis, 17

longirostris, 16

macropus, 16

minor, 15

modesta, 16

montana, 16

montosa, 8

mordax, 16

nanus, 17

nuttalli, 35

occidentalis, 17

operarius, 17

pallidus, 13

pauperrima, 13

phaeus, 17

pinetorum, 17

quasiater, 17

riparia, 17

scalopsoides, 17

trowbridgii, 18

Aschizomys

lemminus, 9

asper

Mus, 74

assimilis

Peromyscus, 47

athabascae

Evotomys, 24

Myodes, 24

Atherura

zygomatica, 90
number 642

• 111

Atherurus

macrourus, 90, 91

terutaus, 91

athi

Steatomys, 7

atratus

Mus, 83

attwateri

Peromyscus, 44

aurantius

Reithrodontomys, 58

auratus

Meriones, 77

aureus

Gerbillus, 69, 70

Reithrodontomys, 59

auricularis

Microtus, 22

aurillus

Oryzomys, 14

auripectus

Peromyscus, 45

auritus

Peromyscus, 44

Reithrodon, 54

austerus

Arvicola, 14, 15

Hesperomys, 41

Peromyscus, 41

australis

Zapus, 5

avius

Peromyscus, 45

azizi

Meriones, 77

aztecus

Oryzomys, 39

Peromyscus, 50, 51

Reithrodontomys, 57

babi

Rattus, 88

badius

Peromyscus, 52

bagobus

Bullimus, 67

Rattus, 67

baileyi

Microtus, 19

Neotoma, 27

bailwardi

Calomyscus, 6

Baiomys

allex, 9

analogus, 9

grisescens, 9

handleyi, 9

musculus, 9, 10

nigrescens, 9

subater, 10

taylori, 9, 10

bairdi

Microtus, 18

bairdii

Hesperomys, 41

balae

Mus, 72

balmasus

Rattus, 89

banacus

Rattus, 76

banderanus

Osgoodomys, 41

Peromyscus, 41, 44

Bandicota

indica, 66

siamensis, 66

bardus

Apomys, 65

barkeri

Rattus, 67

barussanus

Epimys, 79

basilanus

Epimys, 80

batamanus

Mus, 74

BATHYERGIDAE, 90

Batomys

dentatus, 66

batus

Epimys, 74

bellula

Dasyprocta, 92

bellus

Mus, 77

benguetensis

Epimys, 80

bentincanus

Mus, 75

berlandieri

Sigmodon, 59, 61

bernardi

Ondatra, 36

bernardinus

Microtus, 21

Berylmys

bowersi, 66

berytensis

Spalax, 6

Bibimys

chacoensis, 10

blanci

Gerbillus, 69

blandus

Peromyscus, 50

boehmi

Gerbilliscus, 69

bolivaris

Transandinomys, 63

bombycinus

Oryzomys, 63

borealis

Synaptomys, 62, 63

borneanus

Epimys, 88

bowersi

Berylmys, 66

boylei

Peromyscus, 44

boylii

Hesperomys, 42

Peromyscus, 42, 44

brachelix

Microtus, 12

Brachiones

przewalskii, 66

brachyurus

Capromys, 94

brandtii

Lasiopodomys, 12

breviauritus

Onychomys, 36

brevicauda

Zygodontomys, 64

brevicaudus

Evotomys, 24

Myodes, 24

Onychomys, 36, 37

brevicula

Echiothrix, 69

breweri

Arvicola, 15

Microtus, 15

brockmani

Myomyscus, 78

brunnescens

Gerbillus, 68

bryanti

Neotoma, 27, 28, 29, 30, 31, 

32

bullaris

Tylomys, 64

bullata

Neotoma, 31

bullatus

Mus, 83

Peromyscus, 44

Synaptomys, 62

Bullimus

bagobus, 67

Bunomys

andrewsi, 67

chrysocomus, 67

penitus, 67

burrescens

Mus, 83

burrulus

Mus, 83

burrus

Mus, 83

Rattus, 83

burtoni

Melomys, 77

buruensis

Mus, 83

butangensis

Mus, 75

buxtoni

Peromyscus, 47

cafer

Pedetes, 90

calcis

Epimys, 81

californica

Arvicola, 15

californicus

Evotomys, 24

Microtus, 15, 18, 22

Myodes, 24, 25

Peromyscus, 43

caliginosus

Melanomys, 14

Calomys

sorellus, 10

CALOMYSCIDAE, 6

Calomyscus

bailwardi, 6

grandis, 6

campbelli

Phodopus, 53

campestris

Dipodillus, 69

Gerbillus, 68

Hesperomys, 42

Zapus, 5

camurus

Rattus, 80

canescens

Microtus, 21

canicaudus

Microtus, 19

cansus

Eospalax, 6

Myotalpa, 6

cantwelli

Microtus, 21

canus

Eropeplus, 69

Lenothrix, 71

Onychomys, 37

Peromyscus, 44

capitis

Dendromus, 7

capitulatus

Onychomys, 37

CAPROMYIDAE, 94

Capromys

brachyurus, 94

gundlachi, 94

prehensilis, 94

thoracatus, 94

carbonarius

Microtus, 23

Pitymys, 23

carimatae

Mus, 75

carmeni

Peromyscus, 45

carolinensis

Evotomys, 24

Myodes, 24

carpentarius

Notomys, 79
112 • smithsonian contributions to zoology

caryi

Microtus, 20

Reithrodontomys, 57

cascadensis

Clethrionomys, 23

Myodes, 23

casensis

Mus, 75

castaneus

Mus, 78

Peromyscus, 51

catellifer

Mus, 75

caudatus

Meriones, 77

Oryzomys, 38

caurinus

Evotomys, 24

Myodes, 24

Steatomys, 7, 8

CAVIIDAE, 92

celatus

Phenacomys, 52

celsus

Phenacomys, 52

centralis

Epimys, 64

centrosa

Echiothrix, 69

centrosus

Arvicanthis, 66

cereus

Rattus, 76

cervinipes

Melomys, 77

chacoensis

Akodon, 10

Bibimys, 10

chamula

Neotoma, 28

chapmani

Oryzomys, 38, 39

cheesmani

Gerbillus, 69, 70

chejuensis

Apodemus, 64

Chelemys

macronyx, 10

cherriei

Zygodontomys, 64

chiapensis

Dasyprocta, 92

Reithrodontomys, 56

chilensis

Phyllotis, 53

CHINCHILLIDAE, 92

Chiropodomys

gliroides, 67

muroides, 67

niadis, 67

chitralensis

Nesokia, 79

chombolis

Mus, 88

Chraeomys

inornatus, 8

Chrotomys

gonzalesi, 67

mindorensis, 68

whiteheadi, 68

chrysocomus

Bunomys, 67

chrysopsis

Reithrodontomys, 55, 58

cineraceus

Acomys, 64

cinerea

Neotoma, 27, 32, 33

cinereus

Reithrodontomys, 58

cinnamomea

Arvicola, 15

cinnamominus

Fiber, 35

Ondatra, 35

clabatus

Mus, 84

clarae

Epimys, 72

clarus

Dicrostonyx, 10

Onychomys, 37

clementis

Peromyscus, 51

Clethrionomys

arizonensis, 23

cascadensis, 23

gapperi, 23

gauti, 23

imaizumii, 23

maurus, 23

solus, 23

Coendou

koopmani, 91

nycthemera, 91

spinosus, 91

cognatus

Hesperomys, 42

colimae

Reithrodontomys, 55

colimensis

Oryzomys, 39

collinsi

Jaculus, 4

coloratus

Epimys, 81

commissarius

Mus, 78

comptus

Peromyscus, 44

concha [sic]

Epimys, 73

concolor

Rattus, 86

Sicista, 5

confinis

Sigmodon, 60

consobrinus

Peromyscus, 49

constrictus

Microtus, 18

cooperi

Synaptomys, 62, 63

costaricensis

Nyctomys, 34

Reithrodontomys, 55

coucha

Epimys, 74

Rattus, 74

couesi

Erethizon, 91

Oryzomys, 38, 39, 40, 41

coypus

Myocastor, 93

cozumelae

Oryzomys, 39

Peromyscus, 45

crassipinus

Hystrix, 91

crassus

Epimys, 88

Crateromys

paulus, 68

cremoriventer

Mus, 79

Niviventer, 79

crepidatus

Phodopus, 53

CRICETIDAE, 8

Cricetomys

ansorgei, 6, 7

enguvi, 6

gambianus, 6

osgoodi, 6

raineyi, 6

cricetulus

Microtus, 9

crinitus

Hesperomys, 42

Oryzomys, 39

Peromyscus, 42, 45, 51

cristobalensis

Peromyscus, 52

CTENOMYIDAE, 92

Ctenomys

magellanicus, 92

osgoodi, 92

robustus, 92

sericeus, 93

cufrensis

Jaculus, 4

cumulator

Neotoma, 27

CUNICULIDAE, 92

Cuniculus

nelsoni, 92

paca, 92

currax

Pedetes, 90

curtata

Arvicola, 12

curtatus

Lemmiscus, 12, 13

dalli

Synaptomys, 62, 63

daphne

Thomasomys, 63

dariensis

Dasyprocta, 92

Oryzomys, 38

darlingi

Isothrix, 93

Dasymys

helukus, 68

incomtus, 68

nigridius, 68

orthos, 68

rufulus, 68

savannus, 68

Dasyprocta

bellula, 92

chiapensis, 92

dariensis, 92

punctata, 92

richmondi, 92

underwoodi, 92

yucatanica, 92

DASYPROCTIDAE, 92

datae

Apomys, 66

dawsoni

Myodes, 23

decoloratus

Peromyscus, 50

decolorus

Nyctomys, 34

Sitomys, 34

defua

Dephomys, 68

Mus, 68

Dendromus

capitis, 7

insignis, 7

lineatus, 7

melanotis, 7

mesomelas, 7

mystacalis, 7

percivali, 7

spectabilis, 7

whytei, 7

Dendromys

nigrifrons, 7

dentatus

Batomys, 66

Epimys, 82

Pedetes, 90

Dephomys

defua, 68

deserti

Jaculus, 4

Reithrodontomys, 57

desertorum

Neotoma, 28
number 642

• 113

devia

Neotoma, 29

Dicrostonyx

clarus, 10

groenlandicus, 10

nelsoni, 10, 11

peninsulae, 10, 11

richardsoni, 10

stevensoni, 11

unalascensis, 10, 11

difficilis

Peromyscus, 44, 46

Reithrodontomys, 55

dilutior

Oryzomys, 38

Diplomys

labilis, 93

DIPODIDAE, 3

Dipodillus

campestris, 69

maghrebi, 68

zakariai, 68

discolor

Gerbillus, 70

dispar

Neotoma, 28

disparilis

Peromyscus, 45

doani

Erethizon, 91

dolbrogeae

Spalax, 6

dolichurus

Grammomys, 70

Thamnomys, 70

dollmani

Otomys, 80

domelicus

Mus, 75

domesticus

Mus, 78

dominator

Paruromys, 80

Rattus, 80

domitor

Mus, 88

dorsalis

Lemniscomys, 71

Peromyscus, 47

Reithrodontomys, 56

dorsatum

Erethizon, 91

ducis

Epimys, 81

duplicatus

Microtus, 18

durumae

Epimys, 74

dutcheri

Microtus, 19

dybowskii

Mylomys, 78

dychei

Reithrodontomys, 56

easti

Peromyscus, 47

eatoni

Gerbillus, 70

ECHIMYIDAE, 93

Echiothrix

brevicula, 69

centrosa, 69

edax

Arvicola, 15

efficax

Nectomys, 62

Sigmodontomys, 62

ehrenbergi

Spalax, 6

elusus

Peromyscus, 48

elymocetes

Microtus, 19

emesi

Mus, 78

enclavae

Mus, 77

endoecus

Microtus, 21

endorobae

Epimys, 71

Hylomyscus, 71

enganus

Mus, 84

Rattus, 84

enguvi

Cricetomys, 6

Eospalax

cansus, 6

Epimys

alleni, 71

barussanus, 79

basilanus, 80

batus, 74

benguetensis, 80

borneanus, 88

calcis, 81

centralis, 64

clarae, 72

coloratus, 81

concha [sic], 73

coucha, 74

crassus, 88

dentatus, 82

ducis, 81

durumae, 74

endorobae, 71

exsul, 82

fortunatus, 82

fulmineus, 81

gala, 81

gracilis, 79

insulanus, 82

insularum, 72

ismailiae, 73

kaimosae, 71

kaiseri, 64

lamucotanus, 81

lepidus, 79

leucophaeatus, 81

luta, 71

maerens, 81

mara, 81

mayonicus, 81

mengurus, 79

nasutus, 71

neglectus, 81

neumani, 74

niveiventris, 78

norvegicus, 81

ornatulus, 81

pannellus, 82

panya, 74

pauper, 82

perflavus, 74

pollens, 88

potens, 88

querceti, 82

rattus, 82

roa, 82

robiginosus, 82

saturatus, 74

sebucus, 88

shigarus, 82

socer, 81

solus, 79

spatulatus, 79

stentor, 71

taitae, 80

tingius, 83

tua, 83

turbidus, 82

turneri, 64

tyrannus, 83

ubecus, 74

ulae, 78

valens, 88

viclana, 82

victor, 88

vigoratus, 83

vociferans, 72

epixanthum

Erethizon, 91

epixanthus

Erethizon, 91

eremicoides

Peromyscus, 44

eremicus

Hesperomys, 42

Peromyscus, 41, 42, 45, 46, 

51

Sigmodon, 60

eremus

Peromyscus, 48

Erethizon

couesi, 91

doani, 91

dorsatum, 91

epixanthum, 91

epixanthus, 91

myops, 91

ERETHIZONTIDAE, 91

Eropeplus

canus, 69

erythrourus

Meriones, 77

Euchoreutes

alashanicus, 3

naso, 3

Euneomys

mordax, 11

noei, 11

petersoni, 11

eurous

Rattus, 88

eva

Peromyscus, 45

everetti

Rattus, 81, 83, 85

evides

Peromyscus, 51

Evotomys

alascensis, 23

athabascae, 24

brevicaudus, 24

californicus, 24

carolinensis, 24

caurinus, 24

galei, 24

gapperi, 24

hercynicus, 24

idahoensis, 24

limitis, 24

loringi, 24

mazama, 25

nivarius, 25

norvegicus, 25

obscurus, 25

occidentalis, 25

orca, 25

suecicus, 24

ungava, 25

vasconiae, 25

wrangeli, 25

exiguus

Peromyscus, 48

Rattus, 87

exsul

Epimys, 82

exterus

Peromyscus, 48

exulans

Mus, 84

Rattus, 80, 81, 82, 83, 84, 

85, 86, 87, 88

facetus

Rattus, 86

fallax

Neotoma, 28

Pelomys, 80

farsi

Meriones, 77

fasciculata

Trichys, 91
114 • smithsonian contributions to zoology

favillus

Gerbillus, 70

felipensis

Peromyscus, 46

ferrocanus

Mus, 66

ferruginea

Neotoma, 28

fertilis

Arvicola, 12

Hyperacrius, 12

Fiber

cinnamominus, 35

hudsonius, 35

macrodon, 35

mergens, 36

ripensis, 36

spatulatus, 35

zalophus, 36

zibethicus, 35, 36

firmus

Mus, 89

fisheri

Microtus, 18

flammeus

Peromyscus, 35

flavidulus

Mus, 76

flavidus

Loncheres, 93

Pattonomys, 93

flaviventer

Mus, 79

flavus

Sminthus, 5

flebilis

Mus, 84

floridana

Neotoma, 27, 28

floridanus

Oryzomys, 40

Podomys, 54

Sigmodon, 60

fortis

Microtus, 19

fortunatus

Epimys, 82

fremens

Mus, 72

Rattus, 73

frida

Hesperomys, 10

frontalis

Oryzomys, 35

frutectanus

Napaeozapus, 4

fuliginosus

Onychomys, 36, 37

fulmineus

Epimys, 81

fulvescens

Niviventer, 79

Oligoryzomys, 35

Oryzomys, 35

Reithrodontomys, 55, 56, 57, 

58

fulviventer

Microtus, 19

Neotoma, 28

Sigmodon, 61

fulvus

Peromyscus, 50

furvus

Palawanomys, 80

fusca

Neotoma, 32

fuscipes

Jaculus, 4

Neotoma, 28, 29, 33

gala

Epimys, 81

galei

Evotomys, 24

Myodes, 24

gambelii

Hesperomys, 42

Peromyscus, 42, 51

gambianus

Cricetomys, 6

gapperi

Clethrionomys, 23

Evotomys, 24

Myodes, 23, 24, 25

gatunensis

Oryzomys, 39

gauti

Clethrionomys, 23

Myodes, 23

gentilis

Peromyscus, 46

Geocapromys

thoracatus, 94

Georychus

kapiti, 90

geothalsi

Hoplomys, 93

Geoxus

valdivianus, 11

Gerbilliscus

boehmi, 69

inclusus, 69

nigricaudus, 69

robustus, 69

setzeri, 69

gerbillus

Gerbillus, 70

Gerbillus

aeruginosus, 70

andersoni, 69, 70

aquilus, 69, 70

arenicolor, 66

aureus, 69, 70

blanci, 69

brunnescens, 68

campestris, 68

cheesmani, 69, 70

discolor, 70

eatoni, 70

favillus, 70

gerbillus, 70

hamadensis, 70

inflatus, 70

latastei, 69, 70

nalutensis, 69

psammophilous, 70

pyramidum, 70

setzeri, 69

subsolanus, 70

tarabuli, 70

tibesti, 70

versicolor, 70

gilbiventer

Mus, 79

gilmorei

Microtus, 21

glacialis

Alticola, 8

glareolus

Myodes, 24, 25

gliroides

Chiropodomys, 67

goldmani

Nelsonia, 26

Neotoma, 29

Oryzomys, 39

Peromyscus, 46

Reithrodontomys, 56

Sigmodon, 61

gondokorae

Mus, 77

gonzalesi

Chrotomys, 67

gossii

Synaptomys, 63

gossypinus

Peromyscus, 42, 46

gracilis

Epimys, 79

Hesperomys, 42

Peromyscus, 42

Thomasomys, 63

Grammomys

dolichurus, 70

grandis

Calomyscus, 6

gratus

Mus, 78

Peromyscus, 46, 52

gregalis

Microtus, 22

gregorianus

Thryonomys, 91

griseobracatus

Peromyscus, 50

griseoflavus

Reithrodontomys, 56

grisescens

Allactaga, 3

Baiomys, 9

griseus

Mus, 65

Reithrodontomys, 56

groenlandicus

Dicrostonyx, 10

guadalupensis

Microtus, 20

guairae

Proechimys, 93

guardia

Peromyscus, 46

guatemalensis

Microtus, 19

Peromyscus, 46

guerrerensis

Hodomys, 11

Oryzomys, 39

Sigmodon, 60

gundlachi

Capromys, 94

Mysateles, 94

gunnisoni

Peromyscus, 48

guyannensis

Proechimys, 93

gymnotis

Peromyscus, 43

gymnurus

Hoplomys, 93

Habromys

lepturus, 11

lophurus, 11

simulatus, 11

hamadensis

Gerbillus, 70

hamatus

Rattus, 89

Taeromys, 89

handleyi

Baiomys, 9

hatcheri

Reithrodon, 54

haydeni

Arvicola, 15

Microtus, 16

helaletes

Synaptomys, 63

Heliophobius

argenteocinereus, 90

kapiti, 90

hellwaldii

Maxomys, 76

Rattus, 76

helukus

Dasymys, 68

helvolus

Reithrodontomys, 56

hercynicus

Evotomys, 24

hertigi

Micromys, 77

Hesperomys

affinis, 41

anthonyi, 41

articus [sic], 41

austerus, 41

bairdii, 41
number 642

• 115

boylii, 42

campestris, 42

cognatus, 42

crinitus, 42

eremicus, 42

frida, 10

gambelii, 42

gracilis, 42

leucopus, 43

macropus, 54

megalotis, 43

melanophrys, 43

parasiticus, 43

rufinus, 43

sonoriensis, 43

texana, 43

torridus, 36

truei, 43

hildegardeae

Zelotomys, 90

hindei

Aethomys, 64

hirsutus

Reithrodontomys, 56, 57

hispidus

Sigmodon, 59, 60, 61

Hodomys

alleni, 11

guerrerensis, 11

vetulus, 11

hoffmanni

Rattus, 86

hollisteri

Peromyscus, 48

Holochilus

incarum, 12

sciureus, 12

hoogstraali

Lemniscomys, 71

Hoplomys

geothalsi, 93

gymnurus, 93

wetmorei, 93

hualpaiensis

Microtus, 20

hudsonius

Fiber, 35

Zapus, 5, 6

humiliatus

Rattus, 86

humulis

Reithrodontomys, 56, 57

Hybomys

planifrons, 70

hydrochaeris

Hydrochoerus, 92

Hydrochoerus

hydrochaeris, 92

isthmius, 92

notialis, 92

hylaeum

Plagiodontia, 94

hylaeus

Peromyscus, 46

hylocetes

Oryzomys, 39

Peromyscus, 46

hylocoetes

Apomys, 65, 66

Hylomyscus

endorobae, 71

kaimosae, 71

Hyperacrius

fertilis, 12

traubi, 12

wynnei, 12

zygomaticus, 12

hypoxanthus

Oenomys, 79

hystrella

Acomys, 64

HYSTRICIDAE, 90

HYSTRICOMORPHA, 90

Hystrix

crassipinus, 91

sumatrae, 91

Ichthyomys

pittieri, 12

idahoensis

Evotomys, 24

Myodes, 24

Idiurus

macrotis, 90

idoneus

Oryzomys, 14

ignites

Acomys, 64

ignotus

Proechimys, 93

illinoensis

Neotoma, 28

imaizumii

Clethrionomys, 23

Myodes, 23

imhausi

Lophiomys, 14

incarum

Holochilus, 12

incensus

Peromyscus, 47

inclarus

Peromyscus, 48

inclusus

Gerbilliscus, 69

incomtus

Dasymys, 68

indica

Bandicota, 66

Nesokia, 79

inflatus

Gerbillus, 70

innuitus

Microtus, 19

Mictomys, 62

Synaptomys, 62

inopinata

Neotoma, 31

inornata

Neotoma, 31

inornatus

Chraeomys, 8

insignis

Apomys, 65

Dendromus, 7

Napaeozapus, 4

Zapus, 4

insolatus

Rattus, 86

insperatus

Microtus, 22

insulanus

Epimys, 82

insularis

Microtus, 19

Neotoma, 29

insularum

Epimys, 72

insulicola

Peromyscus, 45

integer

Mus, 89

intermedia

Neotoma, 28, 29, 30

intermedius

Phenacomys, 52, 53

iridescens

Pelomys, 80

isiolae

Saccostomus, 7

ismailiae

Epimys, 73

Isothrix

darlingi, 93

isthmica

Neotoma, 30

isthmius

Hydrochoerus, 92

Isthmomys

pirrensis, 12

jacksoni

Praomys, 80

Sigmodon, 60

jaculus

Jaculus, 4

Jaculus

arenaceous, 4

collinsi, 4

cufrensis, 4

deserti, 4

fuscipes, 4

jaculus, 4

rarus, 4

tripolitanicus, 4

vastus, 4

whitchurchi, 4

jalapae

Oryzomys, 39

Reithrodontomys, 55

jebelae

Arvicanthis, 66

jelskii

Abrothrix, 8

jesseni

Synaptomys, 62

juldaschi

Neodon, 26

julianus

Mus, 84

kadiacensis

Microtus, 19

kaimosae

Epimys, 71

Hylomyscus, 71

kaiseri

Aethomys, 64

Epimys, 64

kapiti

Georychus, 90

Heliophobius, 90

keeni

Peromyscus, 46, 47, 50

kelleri

Mus, 84

kempi

Acomys, 64

kennicottii

Arvicola, 16

kentucki

Synaptomys, 62

klamathensis

Reithrodontomys, 57

komareki

Sigmodon, 60

koopmani

Coendou, 91

koratensis

Rattus, 87

koratis

Rattus, 76

kramensis

Rattus, 87

kramis

Rattus, 77

labilis

Diplomys, 93

labradorius

Microtus, 21

laceianus

Peromyscus, 50

laceyi

Peromyscus, 44

Lagostomus

maximus, 92

petilidens, 92

lagunae

Peromyscus, 52

lamucotanus

Epimys, 81

lancavensis

Mus, 73

lanensis

Rattus, 87
116 • smithsonian contributions to zoology

larvalis

Pedetes, 90

lasiae

Rattus, 88

Lasiopodomys

brandtii, 12

mandarinus, 12

latastei

Gerbillus, 69, 70

latifrons

Neotoma, 30

latimanus

Phenacomys, 53

Rhipidomys, 59

Leggada

naivashae, 77

lemminus

Alticola, 9

Aschizomys, 9

Lemmiscus

curtatus, 12, 13

levidensis, 13

pallidus, 13

pauperrimus, 13

Lemmus

alascensis, 13

minusculus, 13

nigripes, 13

subarcticus, 13

trimucronatus, 13

yukonensis, 13

Lemniscomys

dorsalis, 71

hoogstraali, 71

massaicus, 71

mearnsi, 71

pulchellus, 71

rosalia, 71

spermophilus, 71

striatus, 71

lenis

Oligoryzomys, 35

Oryzomys, 35

Lenomys

longicaudus, 71

meyeri, 71

Lenothrix

canus, 71

Leopoldamys

sabanus, 71, 72, 73

siporanus, 72

lepida

Neotoma, 28, 30

lepidus

Epimys, 79

lepturus

Habromys, 11

Peromyscus, 11

leucocephalus

Peromyscus, 46

leucodon

Neotoma, 27, 30, 32

Spalax, 6

leucogaster

Onychomys, 36, 37

leucophaea

Neotoma, 31

leucophaeatus

Epimys, 81

leucopus

Hesperomys, 43

Peromyscus, 41, 42, 43, 44, 

45, 47, 50, 51, 52

leucotis

Sigmodon, 59, 61

levidensis

Lemmiscus, 13

levipes

Peromyscus, 44, 47

Reithrodontomys, 57

levis

Phenacomys, 53

lewisi

Peromyscus, 35

libycus

Meriones, 77

limitis

Evotomys, 24

Myodes, 24

Limnomys

mearnsi, 73

picinus, 83

sibuanus, 73

linduensis

Rattus, 86

lineatus

Dendromus, 7

lingensis

Mus, 75

Rattus, 76

littoralis

Neotoma, 31

Thamnomys, 70

llanoi

Akodon, 8

localis

Rattus, 76

Loncheres

flavidus, 93

longicauda

Reithrodon, 54

longicaudus

Arborimus, 9

Arvicola, 16

Lenomys, 71

Microtus, 14, 16, 17, 18, 19, 

20, 21

Onychomys, 37

Phenacomys, 9

Reithrodontomys, 54, 57

longior

Allactaga, 3

longipes

Onychomys, 37

longipilis

Arvicola, 17

longirostris

Arvicola, 16

Lophiomys

imhausi, 14

thomasi, 14

Lophuromys

ansorgei, 73

aquilus, 73

margarettae, 73

nudicaudus, 73

pyrrhus, 73

lophurus

Habromys, 11

Peromyscus, 11

loringi

Evotomys, 24

Myodes, 24

Thallomys, 89

Thamnomys, 89

losea

Rattus, 87

lucas

Mus, 72

lucida

Neotoma, 27

lucifrons

Reithrodontomys, 58

ludovicianus

Microtus, 20

lugens

Mus, 84

Rattus, 84

luridus

Meriones, 77

luta

Epimys, 71

luteolus

Mus, 75

Reithrodontomys, 55, 58

luteus

Peromyscus, 47

Zapus, 5

mabalus

Rattus, 76

macfarlani

Microtus, 20, 21

mackenzii

Phenacomys, 53

macrodon

Fiber, 35

Ondatra, 35

macronyx

Chelemys, 10

macropus

Arvicola, 16

Hesperomys, 54

Microtus, 16

macrotis

Idiurus, 90

Neotoma, 28, 29, 30

Trichys, 91

macrourus

Atherurus, 90, 91

macrurus

Microtus, 20

madrensis

Microtus, 20

Neotoma, 31

Peromyscus, 47

Sigmodon, 62

maerens

Epimys, 81

magdalenae

Peromyscus, 48

Proechimys, 93

magellanicus

Ctenomys, 92

maghrebi

Dipodillus, 68

magister

Neotoma, 30

magnirostris

Mus, 84

mahomet

Mus, 78

major

Apomys, 66

Sigmodon, 60

Thecurus, 91

Zapus, 5

mandarinus

Lasiopodomys, 12

mandus

Mus, 75

maniculatus

Peromyscus, 41, 42, 43, 47, 

48, 50, 51

mansalaris

Rattus, 73

mansorius

Rattus, 87

mara

Epimys, 81

margarettae

Lophuromys, 73

margaritae

Peromyscus, 48

markhami

Abrothrix, 8

Akodon, 8

marmosurus

Rattus, 86

marshalli

Neotoma, 30

martinensis

Neotoma, 30, 31

Peromyscus, 48

masae

Mus, 72

mascotensis

Sigmodon, 61

massaicus

Lemniscomys, 71

Mastomys

natalensis, 73, 74

matthaeus

Mus, 72

maurus

Clethrionomys, 23

Myodes, 23

maximus

Lagostomus, 92

Maxomys

hellwaldii, 76

musschenbroekii, 76

ochraceiventer, 76
number 642 • 117

pagensis, 75

rajah, 75

surifer, 74, 75, 76, 77

whiteheadi, 74, 75, 76

mayensis

Oligoryzomys, 35

Oryzomys, 35

mayonicus

Epimys, 81

mazama

Evotomys, 25

Myodes, 25

mearnsi

Lemniscomys, 71

Limnomys, 73

Neotoma, 27

Saccostomus, 7

medius

Oryzomys, 64

Peromyscus, 51

megadon

Oryzomys, 40

Megadontomys

nelsoni, 14

thomasi, 14

megalops

Peromyscus, 44, 49

megalotis

Hesperomys, 43

Reithrodon, 54

Reithrodontomys, 54, 55, 56, 

57, 58

mekisturus

Peromyscus, 49

melanocarpus

Peromyscus, 49

Melanomys

caliginosus, 14

melanophrys

Hesperomys, 43

Onychomys, 37

Peromyscus, 43, 49, 52

melanotis

Dendromus, 7

Oryzomys, 39

Peromyscus, 49

Sigmodon, 61

melanura

Neotoma, 29

melanurus

Peromyscus, 49

Rattus, 76

Melasmothrix

naso, 77

Melomys

albiventer, 77

burtoni, 77

cervinipes, 77

mengurus

Epimys, 79

mergens

Fiber, 36

Ondatra, 36

Meriones

amplus, 77

auratus, 77

azizi, 77

caudatus, 77

erythrourus, 77

farsi, 77

libycus, 77

luridus, 77

shawi, 77

merriami

Peromyscus, 46, 49

Reithrodontomys, 57

mesanis

Rattus, 87

mesomelas

Dendromus, 7

Peromyscus, 51

mexicana

Neotoma, 28, 30, 31, 32, 33

mexicanus

Microtus, 17, 19, 20

Oryzomys, 39, 40

Peromyscus, 43, 44, 49, 51

Reithrodontomys, 55, 56, 58

meyeri

Lenomys, 71

microdon

Apomys, 66

Reithrodontomys, 58

Sigmodon, 60

Micromys

hertigi, 77

minutus, 77

micropus

Arvicanthis, 71

Neotoma, 31

Microryzomys

minutus, 14

microtis

Apodemus, 65

microtus

Oxymycterus, 11

Microtus

abbreviatus, 18

acrophilus, 9

adocetus, 21

alticola, 14

amakensis, 18

angusticeps, 18

aphorodemus, 18

arizonensis, 20

arvalis, 18

auricularis, 22

baileyi, 19

bairdi, 18

bernardinus, 21

brachelix, 12

breweri, 15

californicus, 15, 18, 22

canescens, 21

canicaudus, 19

cantwelli, 21

carbonarius, 23

caryi, 20

constrictus, 18

cricetulus, 9

duplicatus, 18

dutcheri, 19

elymocetes, 19

endoecus, 21

fisheri, 18

fortis, 19

fulviventer, 19

gilmorei, 21

gregalis, 22

guadalupensis, 20

guatemalensis, 19

haydeni, 16

hualpaiensis, 20

innuitus, 19

insperatus, 22

insularis, 19

kadiacensis, 19

labradorius, 21

longicaudus, 14, 16, 17, 18, 

19, 20, 21

ludovicianus, 20

macfarlani, 20, 21

macropus, 16

macrurus, 20

madrensis, 20

mexicanus, 17, 19, 20

minor, 15

miurus, 20, 21

modestus, 16

montanus, 16, 17, 19, 20, 21

mordax, 21

muriei, 20, 21

nanus, 17, 21

nemoralis, 22

nesophilus, 19

nevadensis, 21

ochrogaster, 14, 15, 16, 20

oeconomus, 17, 18, 19, 20, 

21, 22, 23

operarius, 17, 19, 21

oreas, 20

oregoni, 18, 21, 22

pamirensis, 26

paneaki, 20

parvulus, 23

pennsylvanicus, 16, 17, 18, 

19, 21, 22

phaeus, 17

pinetorum, 14, 16, 17, 22, 23

popofensis, 22

pullus, 12

quasiater, 17

ravidulus, 22

richardsoni, 16

rivularis, 21

scalopsoides, 14, 16, 17

schmidti, 23

scirpensis, 22

serpens, 22

sitkensis, 22

subsimus, 20

townsendii, 17

tshuktshorum, 22

uliginosus, 19

umbrosus, 22

unalascensis, 22

vallicola, 18

wahema, 21

warringtoni, 12

yakutatensis, 22, 23

Microxus

torques, 8

Mictomys

innuitus, 62

mindanensis

Mus, 84

mindorensis

Chrotomys, 68

Rattus, 83

minima

Sigmodon, 61

minimus

Sigmodon, 61

minnesotae

Peromyscus, 47

minor

Arvicola, 15

Microtus, 15

Zapus, 5

minusculus

Lemmus, 13

Reithrodontomys, 58

minutus

Micromys, 77

Microryzomys, 14

miurus

Microtus, 20, 21

modesta

Arvicola, 16

modestus

Microtus, 16

mollicomus

Rattus, 86

mongolica

Allactaga, 3

monochroura

Neotoma, 33

monstrabilis

Neotoma, 30

montana

Arvicola, 16

montanus

Acomys, 64

Microtus, 16, 17, 19, 20, 21

Reithrodon, 54

Reithrodontomys, 54, 55, 56

Zapus, 6

montezumae

Neotoma, 32

montosa

Arvicola, 8

montosus

Alticola, 8

mordax

Arvicola, 16

Euneomys, 11

Microtus, 21

morotaiensis

Rattus, 86

morulus

Myopus, 25

muelleri

Sundamys, 88, 89
118 • smithsonian contributions to zoology

muenninki

Tokudaia, 90

Tokudamys, 90

MURIDAE, 64

muriei

Microtus, 20, 21

muroides

Chiropodomys, 67

Mus

acholi, 78

albigularis, 83

anambae, 74

andrewsi, 67

apicis, 86

aquilus, 73

arianus, 65

asper, 74

atratus, 83

balae, 72

batamanus, 74

bellus, 77

bentincanus, 75

bullatus, 83

burrescens, 83

burrulus, 83

burrus, 83

buruensis, 83

butangensis, 75

carimatae, 75

casensis, 75

castaneus, 78

catellifer, 75

chombolis, 88

clabatus, 84

commissarius, 78

cremoriventer, 79

defua, 68

domelicus, 75

domesticus, 78

domitor, 88

emesi, 78

enclavae,77 

enganus, 84

exulans, 84

ferrocanus, 66

firmus, 89

flavidulus, 76

flaviventer, 79

flebilis, 84

fremens, 72

gilbiventer, 79

gondokorae, 77

gratus, 78

griseus, 65

integer, 89

julianus, 84

kelleri, 84

lancavensis, 73

lingensis, 75

lucas, 72

lugens, 84

luteolus, 75

magnirostris, 84

mahomet, 78

mandus, 75

masae, 72

matthaeus, 72

mindanensis, 84

musculoides, 77, 78

musculus, 78

obscurus, 84

pagensis, 75

pannosus, 84

pantarensis, 85

pellax, 75

peromyscus, 80

peruvianus, 78

petilus, 78

pulliventer, 85

rostratus, 80

serutus, 75

setzeri, 78

siantanicus, 85

simalurensis, 85

soccatus, 72

sorella, 78

soricoides, 78

stoicus, 85

strepitans, 72

stridens, 72

stridulus, 72

sungarae, 78

surdus, 85

surifer, 75

taciturnus, 85

tagulayensis, 85

tambelanicus, 85

tana, 74

tenellus,78 

tiomanicus, 85

todayensis, 85

triton, 77

tullbergi, 80

umbridorsum, 76

validus, 89

vitiensis, 85

vociferans, 72, 73

vulcani, 86

wamae, 78

zamboangae, 86

musculoides

Mus, 77, 78

Peromyscus, 50

musculus

Apomys, 66

Baiomys, 9, 10

Mus, 78

Peromyscus, 9

Sitomys, 10

musschenbroekii

Maxomys, 76

Rattus, 76

mustelinus

Reithrodontomys, 56

Mylomys

dybowskii, 78

Myocastor

coypus, 93

santacruzae, 93

MYOCASTORIDAE, 93

Myodes

arizonensis, 23

athabascae, 24

brevicaudus, 24

californicus, 24, 25

carolinensis, 24

cascadensis, 23

caurinus, 24

dawsoni, 23

galei, 24

gapperi, 23, 24, 25

gauti, 23

glareolus, 24, 25

idahoensis, 24

imaizumii, 23

limitis, 24

loringi, 24

maurus, 23

mazama, 25

nigripes, 13

nivarius, 25

obscurus, 25

occidentalis, 25

orca, 25

rutilus, 23, 25

solus, 23

ungava, 25

wrangeli, 25

MYOMORPHA, 3

Myomyscus

brockmani, 78

myops

Erethizon, 91

Myopus

morulus, 25

schisticolor, 25

Myotalpa

cansus, 6

Mysateles

gundlachi, 94

prehensilis, 94

mystacalis

Dendromus, 7

nairobae

Arvicanthis, 66

naivashae

Leggada, 77

nalutensis

Gerbillus, 69

nanus

Arvicola, 17

Microtus, 17, 21

napaea

Sicista, 4

Napaeozapus

frutectanus, 4

insignis, 4

roanensis, 4

naso

Euchoreutes, 3

Melasmothrix, 77

nasutus

Epimys, 71

Peromyscus, 44

natalensis

Mastomys, 73, 74

natator

Oryzomys, 38, 40

navus

Neotoma, 32

Neacomys

pictus, 26

nebrascensis

Reithrodontomys, 56

Nectomys

alfari, 62

apicalis, 26

efficax, 62

pollens, 26

rattus, 26

squamipes, 26

tarrensis, 26

vallensis, 26

neglectus

Epimys, 81

nelsoni

Agouti, 92

Cuniculus, 92

Dicrostonyx, 10, 11

Megadontomys, 14

Neotoma, 32

Oryzomys, 40

Peromyscus, 14

Reithrodontomys, 56

Xenomys, 64

Nelsonia

goldmani, 26

neotomodon, 26

nemoralis

Microtus, 22

Neodon

juldaschi, 26

Neofiber

alleni, 26

apalachicolae, 26

nigrescens, 26

Neotoma

albigula, 27, 29, 34

alleni, 11

angusticeps, 29

arizonae, 27

baileyi, 27

bryanti, 27, 28, 29, 30, 31, 

32

bullata, 31

chamula, 28

cinerea, 27, 32, 33

cumulator, 27

desertorum, 28

devia, 29

dispar, 28

fallax, 28

ferruginea, 28

floridana, 27, 28

fulviventer, 28

fusca, 32

fuscipes, 28, 29, 33

goldmani, 29

illinoensis, 28
number 642 • 119

inopinata, 31

inornata, 31

insularis, 29

intermedia, 28, 29, 30

isthmica, 30

latifrons, 30

lepida, 28, 30

leucodon, 27, 30, 32

leucophaea, 31

littoralis, 31

lucida, 27

macrotis, 28, 29, 30

madrensis, 31

magister, 30

marshalli, 30

martinensis, 30, 31

mearnsi, 27

melanura, 29

mexicana, 28, 30, 31, 32, 33

micropus, 31

monochroura, 33

monstrabilis, 30

montezumae, 32

navus, 32

nelsoni, 32

notia, 29

nudicauda, 32

occidentalis, 32, 33

ochracea, 28

orizabae, 32

orolestes, 32

palatina, 32

parvidens, 33

perpallida, 29

phenax, 34

picta, 33

pinetorum, 33

planiceps, 31

pretiosa, 29

ravida, 29

relicta, 33

saxamans, 33

seri, 27

sheldoni, 27

simplex, 28, 30

sola, 28

solitaria, 28

splendens, 33

stephensi, 33

streatori, 29

tenuicauda, 33

torquata, 28, 32

tropicalis, 33

venusta, 27, 33, 34

vicina, 30

warreni, 27

zacatecae, 30

neotomodon

Nelsonia, 26

Neotomodon

alstoni, 34

orizabae, 34

perotensis, 34

Nephelomys

pirrensis, 34

nerterus

Reithrodontomys, 55, 57

Nesokia

chitralensis, 79

indica, 79

NESOMYIDAE, 6

nesophilus

Microtus, 19

neumani

Epimys, 74

nevadensis

Microtus, 21

Zapus, 5

niadis

Chiropodomys, 67

nigellus

Rattus, 67

nigrescens

Baiomys, 9

Neofiber, 26

Peromyscus, 9

Reithrodontomys, 57

nigricauda

Tatera, 69

nigricaudus

Gerbilliscus, 69

nigridius

Dasymys, 68

nigrifrons

Dendromys, 7

nigripes

Lemmus, 13

Myodes, 13

nigritalus

Apodemus, 65

niloticus

Arvicanthis, 66

nitidus

Rattus, 86

nivarius

Evotomys, 25

Myodes, 25

niveiventris

Epimys, 78

Niviventer

cremoriventer, 79

fulvescens, 79

noei

Euneomys, 11

norvegicus

Epimys, 81

Evotomys, 25

Rattus, 81, 84, 86

notatus

Thomasomys, 63

notia

Neotoma, 29

notialis

Hydrochoerus, 92

Notiomys

alleni, 10

vestitus, 10

Notomys

aquilo, 79

carpentarius, 79

noveboracensis

Peromyscus, 42, 47

nudicauda

Neotoma, 32

nudicaudus

Lophuromys, 73

nuttalli

Arvicola, 35

Ochrotomys, 35

Peromyscus, 35

nycthemera

Coendou, 91

Nyctomys

costaricensis, 34

decolorus, 34

pallidulus, 34

sumichrasti, 34

venustulus, 34

oaxacensis

Peromyscus, 50

obscurus

Evotomys, 25

Mus, 84

Myodes, 25

Reithrodontomys, 57

occidentalis

Arvicola, 17

Evotomys, 25

Myodes, 25

Neotoma, 32, 33

ochracea

Neotoma, 28

ochraceiventer

Maxomys, 76

ochraceus

Peromyscus, 47

ochrogaster

Microtus, 14, 15, 16, 20

ochrognathus

Sigmodon, 62

Ochrotomys

nuttalli, 35

Oecomys

trinitatis, 35

oeconomus

Microtus, 17, 18, 19, 20, 21, 

22, 23

Oenomys

hypoxanthus, 79

vallicola, 79

Oligoryzomys

fulvescens, 35

lenis, 35

mayensis, 35

olivaceus

Abrothrix, 8

Ondatra

albus, 35

bernardi, 36

cinnamominus, 35

macrodon, 35

mergens, 36

ripensis, 36

spatulatus, 35

zalophus, 36

zibethica, 36

zibethicus, 35, 36

Onychomys

albescens, 36

aldousi, 36

arcticeps, 36

arenicola, 37, 38

breviauritus, 36

brevicaudus, 36, 37

canus, 37

capitulatus, 37

clarus, 37

fuliginosus, 36, 37

leucogaster, 36, 37

longicaudus, 37

longipes, 37

melanophrys, 37

pallescens, 37

perpallidus, 37

surrufus, 38

torridus, 36, 37, 38

tularensis, 38

utahensis, 37

yakiensis, 38

operarius

Arvicola, 17

Microtus, 17, 19, 21

orca

Evotomys, 25

Myodes, 25

oreas

Microtus, 20

Peromyscus, 50

oregoni

Microtus, 18, 21, 22

oregonus

Zapus, 5, 6

orestes

Otomys, 80

orizabae

Neotoma, 32

Neotomodon, 34

Peromyscus, 49

Reithrodontomys, 58

ornatulus

Epimys, 81

orolestes

Neotoma, 32

orophilus

Phenacomys, 53

orthos

Dasymys, 68

Oryzomys

albiventer, 38

alfaroi, 38, 40

angusticeps, 38

argentatus, 38

aurillus, 14

aztecus, 39

bombycinus, 63

caudatus, 38

chapmani, 38, 39

colimensis, 39

couesi, 38, 39, 40, 41
120 • smithsonian contributions to zoology

Oryzomys (continued)

cozumelae, 39

crinitus, 39

dariensis, 38

dilutior, 38

floridanus, 40

frontalis, 35

fulvescens, 35

gatunensis, 39

goldmani, 39

guerrerensis, 39

hylocetes, 39

idoneus, 14

jalapae, 39

lenis, 35

mayensis, 35

medius, 64

megadon, 40

melanotis, 39

mexicanus, 39, 40

natator, 38, 40

nelsoni, 40

palatinus, 40

palustris, 38, 40

peragrus, 39

pirrensis, 34

planirostris, 40

regillus, 38

rhabdops, 38, 40

richmondi, 40

rostratus, 40

rufinus, 39

rufus, 40

saturatior, 38, 39

talamancae, 64

teapensis, 40

yucatanensis, 40

zygomaticus, 41

osgoodi

Cricetomys, 6

Ctenomys, 92

Osgoodomys

banderanus, 41

vicinior, 41

osimensis

Tokudamys, 90

otion

Pithecheirops, 80

Otomys

dollmani, 80

orestes, 80

Ototylomys

phaeus, 41

phyllotis, 41

otus

Reithrodontomys, 57

Oxymycterus

microtus, 11

paca

Agouti, 92

Cuniculus, 92

pachycephalus

Reithrodon, 54

pacificus

Zapus, 5

pagensis

Maxomys, 75

Mus, 75

palatina

Neotoma, 32

palatinus

Oryzomys, 40

Palawanomys

furvus, 80

palelae

Rattus, 87

pallescens

Apodemus, 65

Onychomys, 37

pallidulus

Nyctomys, 34

pallidus

Arvicola, 13

Lemmiscus, 13

Sigmodon, 61

palustris

Oryzomys, 38, 40

pamirensis

Microtus, 26

paneaki

Microtus, 20

pannellus

Epimys, 82

pannosus

Mus, 84

pantarensis

Mus, 85

panya

Epimys,74 

papagensis

Peromyscus, 45

parasiticus

Hesperomys, 43

Paruromys

dominator, 80

parvidens

Alticola, 8

Neotoma, 33

parvulus

Microtus, 23

Pitymys, 23

parvus

Steatomys, 7

Pattonomys

flavidus, 93

paulus

Crateromys, 68

pauper

Epimys, 82

pauperrima

Arvicola, 13

pauperrimus

Lemmiscus, 13

pectoralis

Peromyscus, 44, 50

Pedetes

cafer, 90

currax, 90

dentatus, 90

larvalis, 90

surdaster, 90

PEDETIDAE, 90

pellax

Mus, 75

Pelomys

fallax, 80

iridescens, 80

roosevelti, 78

penicillatus

Peromyscus, 44

peninsulae

Apodemus, 65

Dicrostonyx, 10, 11

penitus

Bunomys, 67

Rattus, 67

pennsylvanicus

Microtus, 16, 17, 18, 19, 21, 

22

peragrus

Oryzomys, 39

perasper

Rattus, 76

percivali

Dendromus, 7

Tatera, 69

perflavus

Epimys, 74

pergracilis

Peromyscus, 45

peridoneus

Peromyscus, 45

peromyscus

Mus, 80

Peromyscus

affinis, 41, 50

algidus, 47

allex, 9

allophylus, 43

altilaneus, 43

ambiguus, 44

amplus, 44

analogus, 9

angelensis, 44

anthonyi, 41

arenarius, 45

aridulus, 47

assimilis, 47

attwateri, 44

auripectus, 45

auritus, 44

austerus, 41

avius, 45

aztecus, 50, 51

badius, 52

banderanus, 41, 44

blandus, 50

boylei, 44

boylii, 42, 44

bullatus, 44

buxtoni, 47

californicus, 43

canus, 44

carmeni, 45

castaneus, 51

clementis, 51

comptus, 44

consobrinus, 49

cozumelae, 45

crinitus, 42, 45, 51

cristobalensis, 52

decoloratus, 50

difficilis, 44, 46

disparilis, 45

dorsalis, 47

easti, 47

elusus, 48

eremicoides, 44

eremicus, 41, 42, 45, 46, 51

eremus, 48

eva, 45

evides, 51

exiguus, 48

exterus, 48

felipensis, 46

flammeus, 35

fulvus, 50

gambelii, 42, 51

gentilis, 46

goldmani, 46

gossypinus, 42, 46

gracilis, 42

gratus, 46, 52

griseobracatus, 50

guardia, 46

guatemalensis, 46

gunnisoni, 48

gymnotis, 43

hollisteri, 48

hylaeus, 46

hylocetes, 46

incensus, 47

inclarus, 48

insulicola, 45

keeni, 46, 47, 50

laceianus, 50

laceyi, 44

lagunae, 52

lepturus, 11

leucocephalus, 46

leucopus, 41, 42, 43, 44, 45, 

47, 50, 51, 52

levipes, 44, 47

lewisi, 35

lophurus, 11

luteus, 47

madrensis, 47

magdalenae, 48

maniculatus, 41, 42, 43, 47, 

48, 50, 51

margaritae, 48

martinensis, 48

medius, 51

megalops, 44, 49

mekisturus, 49
number 642

• 121

melanocarpus, 49

melanophrys, 43, 49, 52

melanotis, 49

melanurus, 49

merriami, 46, 49

mesomelas, 51

mexicanus, 43, 44, 49, 51

minnesotae, 47

musculoides, 50

musculus, 9

nasutus, 44

nelsoni, 14

nigrescens, 9

noveboracensis, 42, 47

nuttalli, 35

oaxacensis, 50

ochraceus, 47

oreas, 50

orizabae, 49

papagensis, 45

pectoralis, 44, 50

penicillatus, 44

pergracilis, 45

peridoneus, 45

phaeurus, 45

pirrensis, 12

polionotus, 46, 50

polius, 50

polypolius, 46

preblei, 52

prevostensis, 50

restrictus, 46

rubidus, 50

rufinus, 43

santacruzae, 48

saxatilis, 49

simulatus, 11

simulus, 51

sitkensis, 50

sonoriensis, 43, 48, 50

spicilegus, 51

stephani, 51

stephensi, 51

streatori, 48

subater, 10

taylori, 9, 10

teapensis, 49

tehuantepecus, 51

texanus, 43, 44, 51

thomasi, 14

tiburonensis, 51

tornillo, 52

totontepecus, 49

truei, 43, 52

vicinior, 41

xenurus, 52

yucatanicus, 52

zamelas, 49

zamorae, 49

zarhynchus, 52

zelotes, 52

perotensis

Neotomodon, 34

Reithrodontomys, 58

perpallida

Neotoma, 29

perpallidus

Onychomys, 37

peruvianus

Mus, 78

petersoni

Euneomys, 11

petilidens

Lagostomus, 92

petilus

Mus, 78

petraeus

Apomys, 66

phaeurus

Peromyscus, 45

phaeus

Arvicola, 17

Microtus, 17

Ototylomys, 41

Phenacomys

albipes, 9

celatus, 52

celsus, 52

intermedius, 52, 53

latimanus, 53

levis, 53

longicaudus, 9

mackenzii, 53

orophilus, 53

preblei, 53

truei, 53

ungava, 52, 53

phenax

Neotoma, 34

Teanopus, 34

Phodopus

campbelli, 53

crepidatus, 53

phyllotis

Ototylomys, 41

Phyllotis

arenarius, 53

chilensis, 53

vaccarum, 54

wolffhuegeli, 54

xanthopygus, 53, 54

picinus

Limnomys, 83

picta

Neotoma, 33

pictus

Neacomys, 26

pinacus

Rattus, 76

pinatus

Rattus, 89

pinetorum

Arvicola, 17

Microtus, 14, 16, 17, 22, 23

Neotoma, 33

Pitymys, 23

piperis

Rattus, 87

pirrensis

Isthmomys, 12

Nephelomys, 34

Oryzomys, 34

Peromyscus, 12

Pithecheirops

otion, 80

pittieri

Ichthyomys, 12

Pitymys

carbonarius, 23

parvulus, 23

pinetorum, 23

schmidti, 23

Plagiodontia

aedium, 94

hylaeum, 94

planiceps

Neotoma, 31

planifrons

Arvicanthis, 70

Hybomys, 70

Sigmodon, 62

planirostris

Oryzomys, 40

plenus

Sigmodon, 61

Podomys

floridanus, 54

polionotus

Peromyscus, 46, 50

polius

Peromyscus, 50

pollens

Epimys, 88

Nectomys, 26

polypolius

Peromyscus, 46

popofensis

Microtus, 22

potens

Epimys, 88

pothae

Tatera, 69

praetor

Apodemus, 65

Praomys

jacksoni, 80

rostratus, 80

taitae, 80

preblei

Peromyscus, 52

Phenacomys, 53

Zapus, 5

prehensilis

Capromys, 94

Mysateles, 94

pretiosa

Neotoma, 29

prevostensis

Peromyscus, 50

princeps

Zapus, 5, 6

pringlei

Tatera, 69

Proechimys

guairae, 93

guyannensis, 93

ignotus, 93

magdalenae, 93

rubellus, 93

semispinosus, 93

steerei, 93

przewalskii

Brachiones, 66

psammophilous

Gerbillus, 70

pulchellus

Arvicanthis, 71

Lemniscomys, 71

pulliventer

Mus, 85

pullus

Microtus, 12

punctata

Dasyprocta, 92

punicans

Rattus, 89

Taeromys, 89

pusillus

Thryonomys, 91

pyctoris

Rattus, 82

pyramidum

Gerbillus, 70

pyrrhus

Lophuromys, 73

quasiater

Arvicola, 17

Microtus, 17

querceti

Epimys, 82

raineyi

Cricetomys, 6

rajah

Maxomys, 75

Rattus, 76, 77

rallus

Rattus, 67

raptor

Rheomys, 59

rarus

Jaculus, 4

rattus

Epimys, 82

Nectomys, 26

Rattus, 83, 87

Rattus

adspersus, 67

ambersoni, 87

andamanensis, 83, 84, 87

annandalei, 83

antucus, 76

argentiventer, 87

babi, 88
122 • smithsonian contributions to zoology

Rattus (continued)

bagobus, 67

balmasus, 89

banacus, 76

barkeri, 67

burrus, 83

camurus, 80

cereus, 76

concolor, 86

coucha, 74

dominator,80 

enganus, 84

eurous, 88

everetti, 81, 83, 85

exiguus, 87

exulans, 80, 81, 82, 83, 84, 

85, 86, 87, 88

facetus, 86

fremens, 73

hamatus, 89

hellwaldii, 76

hoffmanni, 86

humiliatus, 86

insolatus, 86

koratensis, 87

koratis, 76

kramensis, 87

kramis, 77

lanensis, 87

lasiae, 88

linduensis, 86

lingensis, 76

localis, 76

losea, 87

lugens, 84

mabalus, 76

mansalaris, 73

mansorius, 87

marmosurus, 86

melanurus, 76

mesanis, 87

mindorensis, 83

mollicomus, 86

morotaiensis, 86

musschenbroekii, 76

nigellus, 67

nitidus, 86

norvegicus, 81, 84, 86

palelae, 87

penitus, 67

perasper, 76

pinacus, 76

pinatus, 89

piperis, 87

punicans, 89

pyctoris, 82

rajah, 76, 77

rallus, 67

rattus, 83, 87

raveni, 87, 88

sericatus, 67

simalurensis, 85, 88

solatus, 86

sowerbyi, 86

stoicus, 85

subditivus, 86

tanezumi, 80, 81, 82, 84, 85, 

86, 87

tapanulius, 73

tetricus, 76

tinctus, 74

tiomanicus, 81, 82, 83, 84, 

85, 87

tuancus, 73

umbriventer, 87

virtus, 89

vociferans, 73

raveni

Rattus, 87, 88

ravida

Neotoma, 29

ravidulus

Microtus, 22

ravus

Reithrodontomys, 57

regillus

Oryzomys, 38

Reithrodon

auritus, 54

hatcheri, 54

longicauda, 54

megalotis, 54

montanus, 54

pachycephalus, 54

Reithrodontomys

albescens, 55

albilabris, 58

alleni, 55

alticolus, 58

amoles, 55

aurantius, 58

aureus, 59

aztecus, 57

caryi, 57

chiapensis, 56

chrysopsis, 55, 58

cinereus, 58

colimae, 55

costaricensis, 55

deserti, 57

difficilis, 55

dorsalis, 56

dychei, 56

fulvescens, 55, 56, 57, 58

goldmani, 56

griseoflavus, 56

griseus, 56

helvolus, 56

hirsutus, 56, 57

humulis, 56, 57

jalapae, 55

klamathensis, 57

levipes, 57

longicaudus, 54, 57

lucifrons, 58

luteolus, 55, 58

megalotis, 54, 55, 56, 57, 58

merriami, 57

mexicanus, 55, 56, 58

microdon, 58

minusculus, 58

montanus, 54, 55, 56

mustelinus, 56

nebrascensis, 56

nelsoni, 56

nerterus, 55, 57

nigrescens, 57

obscurus, 57

orizabae, 58

otus, 57

perotensis, 58

ravus, 57

rufescens, 58

saturatus, 58

sumichrasti, 55, 56, 57, 58

tenuirostris, 58, 59

tenuis, 56

toltecus, 57

tolucae, 55

virginianus, 56

zacatecae, 57

relicta

Neotoma, 33

restrictus

Peromyscus, 46

rex

Tachyoryctes, 6

rhabdops

Oryzomys, 38, 40

Rheomys

raptor, 59

Rhipidomys

latimanus, 59

scandens, 59

wetzeli, 59

richardsoni

Dicrostonyx, 10

Microtus, 16

richmondi

Dasyprocta, 92

Oryzomys, 40

riparia

Arvicola, 17

ripensis

Fiber, 36

Ondatra, 36

rivularis

Microtus, 21

roa

Epimys, 82

roanensis

Napaeozapus, 4

Zapus, 4

robiginosus

Epimys, 82

robustus

Ctenomys, 92

Gerbilliscus, 69

roosevelti

Pelomys, 78

rosalia

Lemniscomys, 71

roseveari

Steatomys, 7

rostratus

Mus, 80

Oryzomys, 40

Praomys, 80

roylei

Alticola, 8

rubellus

Proechimys, 93

rubidus

Peromyscus, 50

ruddi

Uranomys, 90

rufescens

Reithrodontomys, 58

rufinus

Hesperomys, 43

Oryzomys, 39

Peromyscus, 43

rufulus

Dasymys, 68

rufus

Oryzomys, 40

rusiges

Apodemus, 65

rutilus

Myodes, 23, 25

sabanus

Leopoldamys, 71, 72, 73

Saccostomus

isiolae, 7

mearnsi, 7

umbriventer, 7

sacobianus

Apomys, 66

santacruzae

Myocastor, 93

Peromyscus, 48

saturatior

Oryzomys, 38, 39

saturatus

Epimys, 74

Reithrodontomys, 58

Sigmodon, 61

savannus

Dasymys, 68

saxamans

Neotoma, 33

saxatilis

Peromyscus, 49

scalopsoides

Arvicola, 17

Microtus, 14, 16, 17

scandens

Rhipidomys, 59

schisticolor

Myopus, 25

schmidti

Microtus, 23

Pitymys, 23

scirpensis

Microtus, 22

sciureus

Holochilus, 12
number 642

• 123

Scotinomys

subnubilis, 59

teguina, 59

sebucus

Epimys, 88

semispinosus

Proechimys, 93

seri

Neotoma, 27

sericatus

Rattus, 67

sericeus

Ctenomys, 93

serpens

Microtus, 22

serutus

Mus, 75

setzeri

Gerbilliscus, 69

Gerbillus, 69

Mus, 78

shawi

Meriones, 77

sheldoni

Neotoma, 27

shigarus

Epimys, 82

siamensis

Bandicota, 66

siantanicus

Mus, 85

sibirica

Allactaga, 3

sibuanus

Limnomys, 73

Sicista

concolor, 5

napaea, 4

Sigmodon

alfredi, 60

alleni, 59, 60

alticola, 59

amoles, 59

arizonae, 60, 61

berlandieri, 59, 61

confinis, 60

eremicus, 60

floridanus, 60

fulviventer, 61

goldmani, 61

guerrerensis, 60

hispidus, 59, 60, 61

jacksoni, 60

komareki, 60

leucotis, 59, 61

madrensis, 62

major, 60

mascotensis, 61

melanotis, 61

microdon, 60

minima, 61

minimus, 61

ochrognathus, 62

pallidus, 61

planifrons, 62

plenus, 61

saturatus, 61

toltecus, 60, 61

tonalensis, 61

virginianus, 61

vulcani, 60

woodi, 61

Sigmodontomys

alfari, 62

efficax, 62

simalurensis

Mus, 85

Rattus, 85, 88

simplex

Neotoma, 28, 30

simulatus

Habromys, 11

Peromyscus, 11

simulus

Peromyscus, 51

siporanus

Leopoldamys, 72

sitkensis

Microtus, 22

Peromyscus, 50

Sitomys

decolorus, 34

musculus, 10

Sminthus

flavus, 5

soccatus

Mus, 72

socer

Epimys, 81

sola

Neotoma, 28

solatus

Rattus, 86

solitaria

Neotoma, 28

solus

Clethrionomys, 23

Epimys, 79

Myodes, 23

sonoriensis

Hesperomys, 43

Peromyscus, 43, 48, 50

sorella

Mus, 78

sorellus

Calomys, 10

soricoides

Mus, 78

sowerbyi

Apodemus, 65

Rattus, 86

SPALACIDAE, 6

Spalax

berytensis, 6

dolbrogeae, 6

ehrenbergi, 6

leucodon, 6

spatulatus

Epimys, 79

Fiber, 35

Ondatra, 35

spectabilis

Dendromus, 7

spermophilus

Lemniscomys, 71

sphagnicola

Synaptomys, 63

Sphiggure

spinosa, 91

spicilegus

Peromyscus, 51

spinosa

Sphiggure, 91

spinosus

Coendou, 91

splendens

Neotoma, 33

squamipes

Nectomys, 26

Steatomys

athi, 7

caurinus, 7, 8

parvus, 7

roseveari, 7

steerei

Proechimys, 93

stentor

Epimys, 71

stephani

Peromyscus, 51

stephensi

Neotoma, 33

Peromyscus, 51

stevensoni

Dicrostonyx, 11

stoicus

Mus, 85

Rattus, 85

stoliczkanus

Alticola, 9

streatori

Neotoma, 29

Peromyscus, 48

strepitans

Mus, 72

striatus

Lemniscomys, 71

stridens

Mus, 72

stridulus

Mus, 72

subarcticus

Lemmus, 13

subater

Baiomys, 10

Peromyscus, 10

subditivus

Rattus, 86

subnubilis

Scotinomys, 59

subsimus

Microtus, 20

subsolanus

Gerbillus, 70

suecicus

Evotomys, 24

sumatrae

Hystrix, 91

Thecurus, 91

sumichrasti

Nyctomys, 34

Reithrodontomys, 55, 56, 57, 

58

Sundamys

muelleri, 88, 89

sungarae

Mus, 78

surdaster

Pedetes, 90

surdus

Akodon, 8

Mus, 85

surifer

Maxomys, 74, 75, 76, 77

Mus, 75

surrufus

Onychomys, 38

Synaptomys

borealis, 62, 63

bullatus, 62

cooperi, 62, 63

dalli, 62, 63

gossii, 63

helaletes, 63

innuitus, 62

jesseni, 62

kentucki, 62

sphagnicola, 63

truei, 63

wrangeli, 63

Tachyoryctes

rex, 6

taciturnus

Mus, 85

Taeromys

hamatus, 89

punicans, 89

tagulayensis

Mus, 85

taitae

Epimys, 80

Praomys, 80

talamancae

Oryzomys, 64

Transandinomys, 64

tambelanicus

Mus, 85

tana

Mus, 74

tanezumi

Rattus, 80, 81, 82, 84, 85, 

86, 87

tapanulius

Rattus, 73

tarabuli

Gerbillus, 70

tarrensis

Nectomys, 26
124 • smithsonian contributions to zoology

Tarsomys

apoensis, 89

Tatera

nigricauda, 69

percivali, 69

pothae, 69

pringlei, 69

varia, 69

Taterillus

arenarius, 89

taylori

Baiomys, 9, 10

Peromyscus, 9, 10

Teanopus

phenax, 34

teapensis

Oryzomys, 40

Peromyscus, 49

teguina

Scotinomys, 59

tehuantepecus

Peromyscus, 51

tenellus

Mus, 78

Zapus, 6

tenuicauda

Neotoma, 33

tenuirostris

Reithrodontomys, 58, 59

tenuis

Reithrodontomys, 56

terutaus

Atherurus, 91

tetricus

Rattus, 76

texana

Hesperomys, 43

texanus

Peromyscus, 43, 44, 51

Thallomys

loringi, 89

Thamnomys

dolichurus, 70

littoralis, 70

loringi, 89

Thecurus

major, 91

sumatrae, 91

thomasi

Lophiomys, 14

Megadontomys, 14

Peromyscus, 14

Thomasomys

daphne, 63

gracilis, 63

notatus, 63

thoracatus

Capromys, 94

Geocapromys, 94

THRYONOMYIDAE, 91

Thryonomys

gregorianus, 91

pusillus, 91

tibesti

Gerbillus, 70

tiburonensis

Peromyscus, 51

tinctus

Rattus, 74

tingius

Epimys, 83

tiomanicus

Mus, 85

Rattus, 81, 82, 83, 84, 85, 87

todayensis

Mus, 85

Tokudaia

muenninki, 90

Tokudamys

muenninki, 90

osimensis, 90

toltecus

Reithrodontomys, 57

Sigmodon, 60, 61

tolucae

Reithrodontomys, 55

tonalensis

Sigmodon, 61

tornillo

Peromyscus, 52

torquata

Neotoma, 28, 32

torques

Akodon, 8

Microxus, 8

torridus

Hesperomys, 36

Onychomys, 36, 37, 38

totontepecus

Peromyscus, 49

townsendii

Microtus, 17

Transandinomys

bolivaris, 63

talamancae, 64

traubi

Hyperacrius, 12

Trichys

fasciculata, 91

macrotis, 91

trimucronatus

Lemmus, 13

trinitatis

Oecomys, 35

trinotatus

Zapus, 6

tripolitanicus

Jaculus, 4

triton

Mus, 77

tropicalis

Neotoma, 33

trowbridgii

Arvicola, 18

truei

Hesperomys, 43

Peromyscus, 43, 52

Phenacomys, 53

Synaptomys, 63

Tryphomys

adustus, 90

tshuktshorum

Microtus, 22

tua

Epimys, 83

tuancus

Rattus, 73

tularensis

Onychomys, 38

tullbergi

Mus, 80

tumbalensis

Tylomys, 64

turbidus

Epimys, 82

turneri

Epimys, 64

Tylomys

bullaris, 64

tumbalensis, 64

tyrannus

Epimys, 83

ubecus

Epimys, 74

ugandae

Uranomys, 90

ulae

Epimys, 78

uliginosus

Microtus, 19

umbridorsum

Mus, 76

umbriventer

Rattus, 87

Saccostomus, 7

umbrosus

Microtus, 22

unalascensis

Dicrostonyx, 10, 11

Microtus, 22

underwoodi

Dasyprocta, 92

ungava

Evotomys, 25

Myodes, 25

Phenacomys, 52, 53

uralensis

Apodemus, 65

Uranomys

ruddi, 90

ugandae, 90

utahensis

Onychomys, 37

vaccarum

Phyllotis, 54

valdivianus

Geoxus, 11

valens

Epimys, 88

validus

Mus, 89

vallensis

Nectomys, 26

vallicola

Microtus, 18

Oenomys, 79

varia

Tatera, 69

vasconiae

Evotomys, 25

vastus

Jaculus, 4

ventriosus

Zygodontomys, 64

venusta

Neotoma, 27, 33, 34

venustulus

Nyctomys, 34

versicolor

Gerbillus, 70

vestitus

Notiomys, 10

vetulus

Hodomys, 11

vicina

Neotoma, 30

vicinior

Osgoodomys, 41

Peromyscus, 41

viclana

Epimys, 82

victor

Epimys, 88

vigoratus

Epimys, 83

vinaceus

Zelotomys, 90

virescens

Arvicanthis, 66

virginianus

Reithrodontomys, 56

Sigmodon, 61

virtus

Rattus, 89

vitiensis

Mus, 85

vociferans

Epimys, 72

Mus, 72, 73

Rattus, 73

vulcani

Mus, 86

Sigmodon, 60

wahema

Microtus, 21

wamae

Mus, 78

warreni

Neotoma, 27

warringtoni

Microtus, 12

wetmorei

Hoplomys, 93

wetzeli

Rhipidomys, 59
number 642

• 125

whitchurchi

Jaculus, 4

whiteheadi

Chrotomys, 68

Maxomys, 74, 75, 76

whytei

Dendromus, 7

wolffhuegeli

Phyllotis, 54

woodi

Sigmodon, 61

wrangeli

Evotomys, 25

Myodes, 25

Synaptomys, 63

wynnei

Hyperacrius, 12

xanthopygus

Phyllotis, 53, 54

Xenomys

nelsoni, 64

xenurus

Peromyscus, 52

yakiensis

Onychomys, 38

yakutatensis

Microtus, 22, 23

yucatanensis

Oryzomys, 40

yucatanica

Dasyprocta, 92

yucatanicus

Peromyscus, 52

yukonensis

Lemmus, 13

zacatecae

Neotoma, 30

Reithrodontomys, 57

zakariai

Dipodillus, 68

zalophus

Fiber, 36

Ondatra, 36

zamboangae

Mus, 86

zamelas

Peromyscus, 49

zamorae

Peromyscus, 49

Zapus

alascensis, 5

australis, 5

campestris, 5

hudsonius, 5, 6

insignis, 4

luteus, 5

major, 5

minor, 5

montanus, 6

nevadensis, 5

oregonus, 5, 6

pacificus, 5

preblei, 5

princeps, 5, 6

roanensis, 4

tenellus, 6

trinotatus, 6

zarhynchus

Peromyscus, 52

zelotes

Peromyscus, 52

Zelotomys

hildegardeae, 90

vinaceus, 90

zibethica

Ondatra, 36

zibethicus

Fiber, 35, 36

Ondatra, 35, 36

Zygodontomys

brevicauda, 64

cherriei, 64

ventriosus, 64

zygomatica

Atherura, 90

zygomaticus

Hyperacrius, 12

Oryzomys, 41
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