RONALD W. HODG Nearctic Walshiidae

Notes and New Taxa

(Lepidoptera: Gelechioidea)

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY-1969 NUMBER 18

SMITHSONIAN CONTRIBUTIONS TO

ZOOLOGY

NUMBER 18

Ronald w. Hodges Nearctic Walshiidae

Notes and New Taxa

(Lepidoptera:

Gelechioidea)

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ABSTRACT

Knowledge concerning the Nearitic members of the Walshiidae is made current by

presenting new information concerning distribution and food plants; describing 14

species (Periploca hostiata, hortatrix, P. opinatrix, P. devia, P. dipapha, P. labes, Aeaea

placatrix, Sorhagenia baucidis, S. pexa, Siskiwitia alticolans, Perimede eremos,

P. parilis, P. circitor, P. maniola) and one genus (Siskiwitia) as new; presenting new

keys to separate species of Periploca, Sorhagenia, and Perimede; and presenting a

list of the known species.

When I discussed the members of various genera of

Walshiidae (Hodges, 1961, 1962a, 1962b, 1962c, and

1964), many of the species were known from few localities. During the past few years I have collected and

received for study examples of several of the known

species as well as 14 new species and one new genus.

This additional information necessitates rewriting some

of the keys to accommodate the new species. To judge

from the number of new species in some genera (Periploca Braun, Aeaea Chambers, Sorhagenia Spuler, and

Perimede Chambers), it appears as though many new

ones remain to be discovered. The small size and relatively uniform maculation make discrimination of

of many species impossible on the basis of external

characters. Also, the rearing of a second species of

Periploca from cedar-apple galls indicates that host

plant information may not be adequate for species

determination in this genus. For these reasons genitalic

preparations were made for all specimens of Periploca

(with the exception of the series of facula Hodges and

orichalcella (Clemens)), the females of Aeaea species,

and all specimens of Perimede (except erransella

Chambers).

Recently, I have had the opportunity to collect

rather extensively in several areas. For the sake of

brevity in citing localities I use an abbreviated form

(indicated in parentheses) for the following: Arizona:

Fort Valley, 7,350 feet, 7/2 miles NW Flagstaff, Coconino County (Fort Valley) ; Hart Prairie, 8,500

feet, 10 miles NNW Flagstaff, Coconino County (Hart

Ronald W. Hodges, Systematic Entomology Laboratory

{United States Department of Agriculture), Smithsonian Institution, Washington, D.C. 20560.

Prairie); Hochderffer Hill, 8,500 feet, 12/2 miles

NNW Flagstaff, Coconino County (Hochderffer Hill) ;

4 miles ESE Pine, 5,400 feet, Gila County (Pine) ;

Vail Lake Road, 6,500 feet, 9/2 miles SE Flagstaff,

Coconino County (Vail Lake); West Fork [Oak

Creek], 6,500 feet, 16 miles SW Flagstaff, Coconino

County (West Fork). Arkansas: Devil's Den State

Park, Washington County (Devil's Den); Hartford,

Sebastian County (Hartford). Florida: Lake Placid,

Archbold Biological Station, Highlands County (Archbold) ; Fisheating Creek, Palmdale, Glades County

(Palmdale); Parker Islands, Highlands County

(Parker Islands). Maryland: Plummers Island [in

Potomac River about 5 miles upstream from Washington, D.C], Montgomery County (Plummers Island).

South Dakota: Hardy Work Center, T3N, R1E, S30

[about 20 miles SW of Lead on U.S. route 85], Lawrence County (Hardy). Wyoming: 6 miles NW Newcastle, Weston County (Newcastle). In addition

Murray O. Glenn of Henry, Illinois, and Charles P.

Kimball of West Barnstable, Massachusetts, have furnished me with specimens collected by themselves and

by others. The citations [MOG] and [USNM] are used

for the collections of Mr. Murray O. Glenn and the

United States National Museum, respectively.

I wish to thank Dr. Harold Grant (deceased) of the

Academy of Natural Sciences, Philadelphia, and

Messrs. John D. Bradley and Allan Watson of the

British Museum (Natural History) for allowing me to

study specimens in their care. The photographs of

maculation were taken by Victor Krantz, staff photographer, Smithsonian Institution.

1
o SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Genus Periploca Braun

Key to North American Species Based on Genitalia

1. Males

 2

Females

 1 9

2. Aedeagus concave dorsally orichalcella (Clemens)

Aedeagus convex or sinuous dorsally 3

3. Valvae asymmetrical &&**• Hodges

Valvae symmetrical ^

4. A long, cylindrical extension from inner surface of valva lobes, new species

Valva unarmed 5

5. Vinculum smooth margined or produced posteriorly 6

Vinculum emarginate posteriorly 13

6. Vinculum produced medially /«'« Hodges

Vinculum smooth margined medially 7

7. Valva triangular ceanothiella (Cosens)

Valva not triangular 8

8. Aedeagus armed externally 9

Aedeagus unarmed 11

9. Costal margin of valva deeply excavated between two-thirds and apex, apex of valva narrowly

acute hostiata, new species

Costal margin of valva sinuous or rounded, apex of valva broadly acute 10

10. Costal margin of valva sinuous, 1 or 2 stout setae from costal margin . opinatrix, new species

Costal margin of valva rounded; setae on valva generally uniform in size, none extremely

stout dipapha, new species

11. Apex of aedeagus upturned hortatrix, new species

Apex of aedeagus straight 12

12. Costal margin of valva strongly indented before apex, apex narrowly acute

gleditschiaeella (Chambers)

Costal margin of valva even, apex broadly acute atrata Hodges

13. Aedeagus unarmed jacula Hodges

Aedeagus armed 14

14. Aedeagus armed dorsally funebris Hodges

Aedeagus armed laterally 15

15. Aedeagus armed on right side 16

Aedeagus armed on left side 18

16. Armature of aedeagus a long and a short projection nigra Hodges

Armature of aedeagus several, short spines 17

17. Aedeagus with 5 to 6 readily visible projections, apex sharply constricted and turned to the

left devia, new species

Aedeagus with numerous, very small projections, apex slightly twisted to the right

cata Hodges

18. Aedeagus with a long projection preceded by a few short ones laeta Hodges

Aedeagus with several, short, asymmetrical projections mimula Hodges

19. Ductus bursae not heavily sclerotized before ostium bursae 20

Ductus bursae heavily sclerotized before ostium bursae 26

20. Posterior margin of 6th sternite invaginated medially orichalcella (Clemens)

Posterior margin of 6th sternite even 21

21. Ductus bursae nearly straight, relatively broad ceanothiella (Cosens)

Ductus bursae twisted several times, slender 22

22. A finger-like imagination on anterior portion of lamella antevaginalis .- 23

No finger-like invagination on anterior portion of lamella antevaginalis 24

23. Width of signum more than one-half basal width of corpus bursae; diameter of invagination

on genital plate approximately equal to diameter of ductus bursae

opinatrix, new species

Width of signum less than one-third basal width of corpus bursae; diameter of invagination

on genital plate less than one-half diameter of ductus bursae ... . hortatrix, new species
NUMBER 18

24. Length of apophyses postenores more than 5 times maximum width of 9th abdominal segment

gleditschiaeella (Chambers)

Length of apophyses postenores less than 2 times maximum width of 9th abdominal segment. 25

25. Ostium bursae on anterior margin of 7th segment, lamella postvaginalis a broadly curving

script "x," signum less than one-fourth width of corpus bursae atrata Hodges

Ostium bursae at one-third on 7th sternite, lamella postvaginalis a very narrow, open

ellipse (open anteriorly), signum approximately one-third width of corpus bursae ... .

labes, new species

26. Posterior margin of 8th abdominal segment with numerous setae 27

Posterior margin of 8th abdominal segment with few setae 28

27. Heavily sclerotized portion of ductus bursae hookshaped facula Hodges

Heavily sclerotized portion of ductus bursae angulate slightly beyond middle . funtbris Hodges

28. Anterior margin of 7th sternite simple, straight cata Hodges

Anterior margin of 7 th sternite modified with lobes, in vagina tions, or productions .. . 29

29. Anterior margin of 7th sternite produced medially laeta Hodges

Anterior margin of 7th sternite emarginate at center or at least less anterad of immediately

adjacent margin 30

30. Anterior margin of 7th sternite emarginate medially devia, new species

Anterior margin of 7th sternite slightly convex anteriorly, flanked by a pair of more anterior

lobes 31

31. A transverse row of setae on dorsal surface of 9th abdominal segment usually present. ... .

nigra Hodges

Dorsal surface of 9th abdominal segment usually lacking a row of setae . . . mimula Hodges

Periploca orichalcella (Clemens), new combination

Elachista ?orichalcella Clemens, 1864, p. 430.

Elachista {?)concolorella Chambers, 1875b, p. 55. [New synonymy.]

Periploca purpuriella Braun, 1919, p. 261. [New synonymy.]

I have examined the type of this species and find it

to be conspecific with concolorella and purpuriella. The

type is in the Academy of Natural Sciences,

Philadelphia.

Continued collecting has turned up specimens from

several areas. I judge that orichalcella is not an uncommon species but that its size contributes to the

adults being overlooked. I have taken a moderate

series in northwestern Arkansas and relatively few

elsewhere; however, the records (Hodges, 1962b, p.

86 and here) indicate that it occurs across the southern

portion of the United States from Florida to California and north to Arkansas and Virginia.

RECORDS.—Florida: Archbold, 1 $, 16-22 May

1964; Palmdale, 1 $, 7-10 May 1964. Illinois: Putnam County, 1 $, 23 July 1963. Arkansas: Devil's

Den, 37 & $, 24 May-20 July 1966; Hartford, 2 $ $ ,

9 June 1966.

Periploca ceanothiella (Cosens)

Periploca ceanthiella (Cosens).—Hodges, 1962b, p. 86.

RECORDS.—South Dakota: Hardy, 1 3 , 11 July

1965. Arizona: Fort Valley, 3 t S , 2 9 9, 21 June6 July 1961; Hart Prairie, 1 * , 7 July 1961.

The South Dakota record represents a considerable

northcentral extension in the known range of

ceanothiella.

Periploca gleditschiaeella (Chambers)

Periploca gleditschiaeella (Chambers).—Hodges, 1962b, p.

88.

A small number of specimens was taken at light

immediately adjacent to Gleditsia triacanthos L.

(honeylocust) in northwest Arkansas.

RECORDS.—Arkansas: Devil's Den, 4 $ 3,12June -

22 July 1966. Illinois: Putnam County, 19, 7 June

1967.

Periploca hostiata, new species

FIGURE 7

A small, shining, gray-black species. Head, thorax,

and forewing shining gray black; tongue yellow white;

shaft of antenna yellow white overlaid with black;

hind wing yellow white; cilia slightly gray. Legs pale

yellow gray basally, gray black distally, apices of segments pale. Alar expanse: 8.3 mm. Male genitalia:

as in Figure 7 (RWH slide 4459). Costal margin of

valva incurved at two-thirds length, apex sharply

acute; apex of aedeagus with a short, dorsal flange, a

series of small triangular projections ventrolaterally at

three-fourths, others dorsally beyond one-half; pos-
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

terior margin of vinculum truncate. Female genitalia:

no specimens available.

FOOD PLANT.—Unknown.

HOLOTYPE.—Male, 2.5 mi W Ft. Simcoe, Yakima

County, Washington, 31 July 1962, J. F. G. Clarke

(RWH slide 4459). USNM Type 69791.

Periploca hostiata is nearest gleditschiaeella in characters of the valva and vinculum. It may easily be

separated from the latter, however, by having grayblack scales and an armed aedeagus.

Periploca hortatrix, new species

FIGURES 1, 10, 28, 29

A small, dark, shining gray-black species. Head,

basal half of antenna, forewing, and legs shining gray

black; distal half of antenna, hind wing, coxae, and

apices of tibiae and tarsal segments pale yellowish

white. Alar expanse: <$ 9 mm. 9 6.5-7.0 mm. Male

genitalia: as in Figure 10 (RWH slide 4414). Valva

with acute apex, costal margin moderately swollen

medially, then slightly depressed at four-fifths; aedeagus unarmed, apex abruptly upturned; vinculum

smooth margined posteriorly. Female genitalia: as in

Figures 28 and 29 (RWH slides 4096, 4097). Posterior

margin of eighth abdominal segment with few setae;

ostium bursae near anterior margin of seventh sternite,

surrounded by a lightly sclerotized plate, a medial,

fingerlike invagination near anterior margin of plate;

ductus bursae twisted five or six times; signa paired,

each an invaginated unit, appearing broadly elliptical

in outline, anterior half of ellipse heavily sclerotized,

posterior half lightly sclerotized.

FOOD PLANT.—Unknown.

HOLOTYPE.—Female, Devil's Den State Park, Washington County, Arkansas, 20 July 1966, R. W. Hodges

(RWH slide 4097). USNM Type 69779.

PARATYPES.—Same locality as for type, 1 $, 25 May

1966 (RWH slide 4414) [USNM]: 1 9, 10 June 1966

(RWH slide 4096) [University of Arkansas]. 2 H ,

Putnam County, Illinois, 4, 7 June 1966 and 1967,

M. O. Glenn (MOG preparations 10468, 10768)

[MOG].

This species seems to be near atrata and opinatrix

but differs from the former in that the male of hortatrix has the apex of the valva acute and lacks a triangular projection on the saccular margin. From the

latter the males of hortatrix differ in having the aedeagus unarmed. The female of hortatrix differs from that

of opinatrix in having few setae (uniserial) at the

posterior margin of the eighth abdominal segment and

in having the fingerlike invagination of the lamella

antevaginalis slender, nearly twice as long as the opening. In opinatrix the setae are numerous and biserial.

and the fingerlike process is broad and blunt.

Periploca opinatrix, new species

FIGURES 8, 9, 27

A small, dark, gray-black species. Head, thorax, and

forewings shining gray black. Dorsal surface of second

segment of labial palpus, base of tongue, and hind wing

pale yellowish white. Legs yellowish white overlaid

with gray black, apex of fore- and midtibiae yellowish

white, hind leg mainly yellowish white, dusted with

dark gray black, apices of tarsal segments paler than

remainder of segments. Alar expanse: 9-10 mm. Male

genitalia: as in Figures 8 and 9 (RWH slide 3953).

Costal margin of valva strongly arched with a strong

seta at two-thirds length and another at one-third, apex

of valva acute; aedeagus sinuous, armed dorsally and

on right side at three-fourths with four conical projections (dorsal pair larger than lateral pair); basal margin of vinculum emarginate. Female genitalia: as in

Figure 27 (RWH slide 3960). Ostium bursae near anterior margin of seventh sternite, opening nearly onefourth total length of segment, surrounded by a lightly

sclerotized region; ductus bursae coiled five or six

times; signa paired, near base of corpus bursae, each

heavily sclerotized, shaped as half of an ellipse; anterior

margin of seventh sternite produced medially to accommodate opening of stout fingerlike pouch; setae on

posterior margin of eighth abdominal segment numerous, multiserial.

FOOD PLANT.—Unknown.

HOLOTYPE.—Male, 6 mi NW Newcastle, Wyoming,

13 July 1965, R. W. Hodges (RWH slide 3953).

USNM Type 69780.

PARATYPES.—Same locality as for holotype, 23 &,

6 9 9, 13-20 July 1965 (RWH slides 3950, 51, 54-57,

59, 60) [USNM].

Males of opinatrix may be separated from those of

other species of Periploca by having the aedeagus armed

with four conical projections at three-fourths, the anterior margin of the vinculum emarginate, and the apex

of the valvae acute. Females of opinatrix are nearest

those of hortatrix but may be separated by having the

fingerlike pouch at the base of the seventh sternite

large and stout; the opening is approximately one-half

the width of the ostium bursae.
NUMBER 18

Variation is expressed mainly by the amount of dark

gray-black scales on the hind legs. As specimens become worn, the dark gray-black scales are lost, so the

forewings appear very light.

Periploca atrata Hodges

Periploca atrata Hodges, 1962b, p. 90.

RECORDS.—Arizona: Hochderffer Hill, 1 $, 16 July

1961; Madera Canyon, Bog Springs Gamp Ground,

Santa Cruz County, 1 8, 1 9, 10-26 July 1964; Southwestern Research Station, Chiricahua Mountains, Cochise County, 5,400 ft, 1 S, 31 July-2 August 1964;

Vail Lake, 1 S , 11 July 1961.

Periploca mimula Hodges

Periploca mimula Hodges, 1962b, p. 92.

Females of mimula and nigra are nearly inseparable

on maculation or characters of the terminalia. A possible point of difference is as indicated in the key to

species. Some females of nigra have a transverse, dorsal

row of setae (6-8) on the ninth abdominal segment,

whereas this area has no setae in most examined specimens of mimula. Features of the genital plate and the

ductus bursae are somewhat variable and so far have

been useless to distinguish between the two.

In 1962 the type number was stated incorrectly. It

should read "Cornell University type number 3823."

RECORDS.—Arizona: Vail Lake, 13,1 1 July 1961.

Arkansas: Devil's Den, 3 SS, 19-21 May, 13 July

1966. New Mexico: Gran Quivira National Monument, Socorro County, 6,600 ft, 1 S, 1-3 July 1964.

Washington: 2 mi W Mazama, Okanogan County,

1 9 , 29 July 1962. Wyoming: Newcastle, 2 $ $ , 1 9,

28 June 18-20 July 1965.

Periploca cata Hodges

Periploca cata Hodges, 1962b, p. 93.

The holotype of cata is a female rather than a male

as stated on page 94 (Hodges, 1962b).

RECORDS.—Arkansas: Devil's Den, 10 $ $, 19

May-27 June 1966. Iowa: Shenandoah, 5 $ 3,3 9 9,

8 June 1946, reared from cedar rust gall.

Periploca devia, new species

FIGURES 11,30-32

A small, dark, gray-black species. Head, thorax, and

forewing shining gray black. Dorsal surface of second

segment of labial palpus, base of tongue, frons, and

coxae shining yellowish white; scape of antenna and

base of shaft dark, remainder yellowish white. Hind

wing yellowish white, becoming slightly darker distally;

ventral surface much lighter than dorsal surface. Foreleg yellowish white heavily overlaid with gray black.

Tibia and tarsus of midleg dark, apices of segments

pale yellowish white. Hind leg gray brown. Alar expanse: 8-10 mm. Male genitalia: as in Figure 11

(RWH slide 4371). Apex of aedeagus recurved, armed

with a row of 6-8 conical projections on right side from

approximately one-half to three-fourths; costal

margin of valva strongly convex, apex blunt; posterior

margin of vinculum emarginate medially. Female

genitalia: as in Figures 30-32 (RWH slide 4365).

Ostium bursae at approximately one-third distance

from anterior margin of seventh sternite; genital plate

developed as a narrow, sclerotized band around ostium

bursae; anterior margin of seventh sternite slightly invaginated medially, a pair of shallow lobes immediately

laterad of this invagination; ductus bursae heavily

sclerotized just before ostium bursae, preceded by a

short, less heavily sclerotized section, the remainder

lightly sclerotized, looped five or six times; signa a pair

of linear invaginations near base of corpus bursae,

inner margin of each with a pair of inwardly curving

hooks.

FOOD PLANT.—Unknown.

HOLOTYPE.—Male, Fort Valley, 7,350 ft, 7/ 2 mi

NW Flagstaff, Coconino County, Arizona, 22 June

1961, Ronald W. Hodges (RWH slide 4371). USNM

Type 69781.

PARATYPES.—Same locality as for holotype, 1 $,

19 June 1961 (RWH slide 4372) [USNM]; 1 9, Hart

Prairie, 8,500 ft, 10 mi NNW Flagstaff, Coconino

County, Arizona, 14 July 1961, Ronald W. Hodges

(RWH slide 4374) [USNM]; 2 9 9, Hochderffer Hill,

8,500 ft, 12 J/2 mi NNW Flagstaff, Coconino County,

Arizona, 16 July and 8 August 1961, Ronald W.

Hodges (RWH slides 4365, 4366) [Cornell University,

USNM].

The male genitalia of devia are nearest those of cata

in general configuration but differ in having the

aedeagus armed with a relatively small number (5-6)

of easily visible projections, the apex of the aedeagus

sharply constricted and curved to the left and extending but a short distance beyond the posterior margin of

the vinculum. The female genitalia are nearest those

of laeta, differing in having the anterior margin of the
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

seventh sternite incurved medially and the basal,

sclerotized portion of the ductus bursae heavily

sclerotized immediately before the ostium bursae, then

less heavily sclerotized.

Periploca nigra Hodges

Periploca nigra Hodges, 1962b, p. 94.

RECORDS.—Arkansas: Devil's Den, 11 $$,299 ,

20 May- 5 June 1966.

Periploca fessa Hodges

Periploca fessa Hodges, 1962b, p. 95.

RECORDS.—Arkansas: Devil's Den, 5 $ $, 20

May-3 June and 22 July 1966. Wyoming: Newcastle, 1 4,13 July 1965.

Periploca facula Hodges

Periploca facula Hodges, 1962b, p. 96.

RECORDS.—Arizona: Fort Valley, 2 $ $, 22 June

1961. Wyoming: Newcastle, 4 $ $, 7 9 9, reared

from cedar apple galls on Juniperus species, collected

23 July, issued 3-18 August 1965; 14,1 3 July 1965.

Periploca dipapha, new species

FIGURES 12, 13

A small, shining, yellowish-brown and pale yellow

moth. All surfaces shining yellow at certain angles of

light incidence, shining yellowish brown overlying pale

yellowish white scales on outer surface of second and

third segments of labial palpus, dorsal surface of scape

(apex pale) and base of shaft of antenna, thorax, forewing, and femur, tibia, and tarsus of foreleg. Alar expense: 8.5 mm. Male genitalia: as in Figures 12 and 13

(RWH slide 1060). Aedeagus with three prominant

conical projections on dorsal surface between twothirds and three-fourths length, apex blunt with a dorsal, conical projection just before end; costal margin

of valva nearly evenly convex, apex broadly acute; posterior margin of vinculum evenly rounded. Female

genitalia: no specimens available.

FOOD PLANT.—Unknown.

HOLOTYPE.—Male, Madera Canyon, 4,880 ft, Santa

Rita Mtns., Arizona, July 27, 1959, R. W. Hodges

(RWH slide 1060). Cornell University type number

4268.

The male genitalia of dipapha may be separated

from those of funebris by having the costal margin of

the valva broadly rounded (sinuous in funebris) and

only three preapical, conical projections on the dorsal

surface of the aedeagus (funebris has a series of large

projections and many small, triangular ridges in the

same location).

The unique specimen of dipapha is not fresh, so the

description of maculation will probably be modified as

others turn up. Dark scales are likely to be more abundant than is indicated in the description.

Periploca labes, new species

FIGURES 14, 33

A small, pale yellow species. Nearly all scaled surfaces shining, pale yellowish white, posterior surface of

scape of antenna, tegula, thorax, and forewing dusted

with light brown scales. Femur, tibia, and tarsus of

foreleg brown, apex of tibia pale yellow, apices of

tarsal segments lighter than remainder of segment.

Tibia of midleg with a few yellow-brown scales before

apex, ventral surface of tarsus yellow brown. Alar expense: 7 mm. Male genitalia: as in Figure 14 (RWH

slide 1163). Aedeagus unarmed, sinuous, tapering to

apex; costal margin of valva rounded, apices, rounded,

a long, curved, medial projection rising from inner

surface extending beyond apex; part of vinculum

heavily sclerotized as a narrow band articulating with

tegumen, remainder lightly sclerotized. Female genitalia: as in Figure 33 (RWH slide 4377). A single,

sparse row of setae on eighth segment; ostium bursae

circular, surrounded by an incomplete (open anteriorly), heavily sclerotized ring, located slightly posterad

of anterior margin of seventh sternite; ductus bursae

membranous with approximately seven loops; corpus

bursae lightly sclerotized; a pair of crescent-shaped

signa at basal third of corpus bursae.

FOOD PLANT.—Unknown.

HOLOTYPE.—Female, Vail Lake Road, 6,500 ft, 9l/2

mi SE Flagstaff, Coconino County, Arizona, 11 July

1961, Ronald W. Hodges (RWH slide 4377). USNM

69782.

PARATYPE.—Male, Madera Canyon, 4,880 ft, Santa

Rita Mtns., Arizona, July 17, 1959, R. W. Hodges

(RWH sUde 1163) [Cornell University].

I have associated the sexes of labes because of the

similarity in maculation. Inasmuch as no other Periploca species is so pale in a fresh condition, this character seems to be diagnostic. The pale yellowish color
NUMBER 1 8

of labes immediately separates it from other species of

Periploca. Possession of the long projection from the

inner surface of the valva will immediately separate

males of labes from those of other species of Periploca.

The presence of crescent-shaped signa, membranous

ductus bursae, ostium bursae posterad of anterior margin of seventh sternite, seventh sternite basically simple, and a single row of setae on the eighth segment

will, in combination, characterize females of labes.

Genus Synploca Hodges

Synploca gumia Hodges

Synploca gumia Hodges, 1964, p. 295.

RECORDS.—Arizona: Fort Valley, 1 3, 25 July 1961;

Pine, H, 5 September 1-961.

Genus Walshia Clemens

Walshia miscecolorella (Chambers)

Walshia miscecolorella (Chambers).—Hodges, 1961, p. 69.

This species becomes moderately common at light

over a relatively long period of time. The work of

George Manglitz (in litt.) at Lincoln, Nebraska, indicates that the species can be a minor economic pest—

in this instance of sweet clover.

When series of specimens are available, several general statements can be made about populations: The

California specimens are large and pale, usually the

basal dark brown patch contrasts sharply with the rest

of the wing which is pale yellow, and the hind wings

often are pale yellow. The northern Arizona population is represented by large specimens (average: 16.4

mm, range: 11.8-18.6 mm, 70 specimens measured)

with relatively uniform markings; the yellow-brown

area beyond the basal, dark brown patch is moderately

dark and becomes suffused yellow-brown in a short

distance. A series of specimens from northwest Arkansas is distinctly small (average: 11.6 mm, range: 9.4-

15.0 mm, 31 specimens measured) and relatively dark,

and another series (reared) from Lincoln, Nebraska,

is moderate sized (average: 13.5 mm, range: 11.0—

15.4 mm, 16 specimens measured) and relatively dark

colored.

This species shows more evidence of variation than

the other Walshia species, but the difference may be a

result of more material being collected than for the

others.

333-341 O—

RECORDS.—Arizona: Fort Valley, 38 3 3, 3 9 9,

20 June-1 August 1961; Hart Prairie, 85 3 3 , 7 9 9 ,

23 June-22 August 1961, Hochderffer Hill, 2 3 3,

9 July 1961; Vail Lake, 2 9 9, 11 July 1961; West

Fork, 8 3 3, 4-15 July 1961. Arkansas: Devil's Den,

32 3 3 , 6 9 9 , 20 May-22 July 1966. Florida: Archbold, 1 3,1-7 May 1964. Nebraska: Lincoln, 7 3 3,

9 9 9, issued 12-17 March 1967. Nevada: Kernshaw

Ruan State Park near Caliente, 1 3, 21 June 1963.

Walshia exemplata Hodges

Walshia exemplata Hodges, 1961, p. 73.

RECORDS.—Arkansas: Devil's Den, 13, 6 July 1966.

Walshia particornella (Busck)

Walshiaparticornella (Busck).—Hodges, 1961,p.73.

RECORDS.—Arkansas: Devil's Den, 6 9 9 , 20 May13 July 1966.

Of the six specimens taken in Arkansas, one was

extremely fresh and has patches of raised scales on the

forewings. It appears as though particornella does fit

better in the genus Walshia with this character being

present. These particular scales must be extremely

deciduous to be lost while specimens still appear fresh.

Genus Aeaea Chambers

Aeaea venifica Hodges

FIGURE 36

Aeaea venifica Hodges, 1964, p. 301.

The first recognized females of venifica were taken

by Mr. Glenn. Subsequently, a large series of males

and females was collected in northwestern Arkansas.

As seems to be true for many of the small species of

Microlepidoptera, venifica can be relatively common,

but it is not collected by most workers.

The following description of the female genitalia is

given to supplement the original statement concerning

the species. Female genitalia: as in Figure 36 (RWH

slide 4415). Ostium bursae at or slightly beyond middle

of seventh sternite; genital plate with a low ridge

from anterior margin to ostium bursae flanked by a

shallow, depressed, elliptical, sclerotized region, latter

with a series of concentric, semicircular lines laterally;

ductus bursae narrow at base then expanding at level

of anterior margin of seventh stemite, coiled once;
8 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

corpus bursae nearly membranous, walls with fine

spicules, signa a pair of inwardly pointing thorns.

The series of concentric ridges on the genital plate

of venifica ally it with A. duke do Hodges; however,

the genitalia of the former have a median ridge on the

genital plate, the ostium bursae on the seventh sternite

(not the margin), and the posterior margin of the

seventh sternite even (not emarginate).

RECORDS.—Arkansas: Devil's Den, 62 $ $,

19 9 9, 24 May-10 July 1961. Illinois: Putnam

County, 2 4 4 , 2 9 9 , 30 May-5 June 1962. Maryland: Plummers Island, 3 4 4,599 , 20-28 July

1962.

Aeaea ostryaeella Chambers

Aeaea ostryaeella Chambers.—Hodges, 1964, p. 302.

RECORDS.—Arkansas: Devil's Den, 13 4 4, 4 9 9 ,

4 June- 6 July 1966. Maryland, Plummers Island,

1 4 , 20 July 1962.

Aeaea quadricustatella Chambers

Aeaea quadricustatella Chambers.—Hodges, 1964, p. 303.

RECORDS.—Arkansas: Devil's Den, 56 4 4,11 9 9,

5 June-22 July 1966. Florida: Archbold, 31 $ $,

26 9 9, 1 May-8 June 1964; Parker Island, 1 9 ,

26-29 May 1964.

Aeaea placatrix, new species

FIGURE 35

A small, dark gray-black species. Maculation as for

A. juvantis Hodges. Alar expanse: 7 mm. Male genitalia: no specimens available. Female genitalia: as in

Figure 35 (RWH slide 4141). Genital plate relatively

complicated, a pair of flanges running from anterior

to posterior margins of seventh sternite, commencing

near lateral margin, terminating medially in a pair

of pointed, free-standing ridges; ostium bursae in anterior section of seventh abdominal segment, opening

to exterior near or at end of segment; ductus bursae

broadest at ostium bursae, tapering to slightly beyond

half length, then gradually broadening to corpus

bursae, basal one-fourth moderately sclerotized; corpus

bursae nearly membranous, wall with fine spicules;

signa a pair of inwardly pointing spines.

FOOD PLANT.—Unknown.

HOLOTYPE.—Female, Devil's Den State Park, Washington County, Arkansas, 20 June 1966, R. W. Hodges

(RWH slide 4141). USNM Type 69783.

Females of placatrix are closest to those of A. victor

Hodges but may be separated by having the pair of

flanges starting near the lateral margins of the seventh

abdominal segment rather than near the middle.

Genus Chrysopeleia Chambers

Chrysopeleia purpuriella Chambers

Chrysopeleia purpuriella Chambers.—Hodges, 1964, p. 315.

Males of purpuriella can be identified by examination of the abdomen without resorting to genitalic

preparation. The modified scales of the eighth abdominal sternite are easily visible.

RECORDS.—Arkansas: Devil's Den, 10 $ 4,10 $ $ ,

1-19 July 1966. Maryland: Plummers Island, 2 9 9,

20 July 1962.

Genus Stilbosis Clemens

Examination of male genitalia of additional Stilbosis species has shown that Aeaea and Stilbosis are

very close if not the same. The character of the saccular

margin of the valva being heavily sclerotized is good

for some but not all species of Stilbosis. An extremely

obvious difference is the nature of the scales: Stilbosis

has metallic scales that are shades of yellow and brown;

Aeaea has scales that are dull gray-black and white.

Differences in the nature of the scales are usually not

important enough to separate genera; however, until

I have had the opportunity to study the numerous

Neotropical representatives of Stilbosis, I prefer to

maintain the two genera as distinct

Stilbosis lonchocarpella Busck

FIGURES 2,15, 34

Stilbosis lonchocarpella Busck, 1934, p. 157.

This species has been taken commonly by Mrs.

Spencer Kemp at Key Largo, Florida. Busck cites

Lonchocarpus sericeus as the host of the larva; however, this plant is not known to occur in Florida, so

another legume probably serves as host there.

RECORDS.—Florida: Key Largo, Monroe County,

28 4 4 , 14 9 9,1 6 December-20 April.
NUMBER 18

Genus Sorhagenia Spuler

Key to Nearctic Species Based on Genitalia

1. Males 2

Females 5

2. Costal lobe of valva longer than saccular lobe 3

Costal lobe of valva shorter than saccular lobe' 4

3. Costal lobe of valva broadly rounded at apex; saccular lobe relatively evenly sclerotized, more

heavily sclerotized than costal lobe baucidis, new species

Costal lobe of valva acute at apex, more heavily sclerotized than saccular lobe nimbosa (Braun)

4. Medial lobe of valva relatively constant in width, slightly expanded at apex; costal lobe slightly

expanded before apex, then tapering to sharply pointed apex pexa, new species

Medial lobe of valva expanded distally, more than twice medial width; costal lobe tapering

evenly to narrowly rounded apex daedala Hodges

5. Signa absent pexa, new species

Signa present 6

6. Pregenital plate rectangular nimbosa (Braun)

Pregenital plate triangular 7

7. Ductus bursae slender, width nearly uniform; sclerotized, anterior arms of pregenital plate

narrow, posterior margin excavated daedala Hodges

Ductus bursae moderate, anterior portion nearly twice as wide as posterior, sclerotized portion;

sclerotized, anterior portion of pregenital plate broad, posterior margin truncate

baucidis, new species

Sorhagenia baucidis, new species

FIGURES 3, 16-17,37-38

A small, dark gray species. Head and thorax dark

gray, scales tipped with pale gray to off-white, thorax

darker than head; apex of third segment of labial

palpus off-white; apex of antenna with five half-segments pale gray, each pale half-segment separated by

one and one-half dark gray segments. Scales of forewing much as for thorax, dorsal half from just beyond

base to one-half pale gray, a few completely dark

gray scales at base and some on middle third near

costa; five patches of raised scales, a costal and

dorsal pair at one-fourth, a costal and dorsal pair at

one-half, and a medial one at end of cell, first pair

and outer one unicolorous dark gray, middle pair

lighter and tipped with pale gray; a row of three dark

gray (almost black) spots on outer margin; cilia gray,

scales of costal portion tipped with pale gray, dorsal

ones unicolorous. Hind wing unicolorus gray. Legs

gray to dark gray, scales tipped with pale gray; foreleg with apex and middle of tibia and apices of tarsal

segments 1, 2, 4, 5 pale gray; midlegs absent on type;

tibia of hind leg with pale oblique streaks subbasally,

medially, and apically, apices of tarsal segments offwhite; legs darker distally than basally. Alar expanse:

10.0-10.5 mm. Male genitalia: as in Figures 16 and

17 (MOG slide 25). Costal lobe of valva expanded

for distal two-thirds, apex broadly rounded, medial

lobe short with sinuous ventral margin; costal lobe

heavily sclerotized, margins nearly parallel, apex

rounded; juxta roughly "T" shaped. Eighth abdominal

sternite moderately heavily sclerotized. Female genitalia: as in Figures 37 and 38 (RWH slide 4529). Pregenital plate triangular, apex truncate; basal portion

of ductus bursae sclerotized, distal half with transverse

ridges, spiculate; corpus bursae spherical, two signa

present. Apophyses anteriores and posteriores subequal

in length.

FOOD PLANT.—Unknown.

HOLOTYPE.—Male, Putnam County, Illinois, June

17, 1965, M. O. Glenn (MOG slide 25). USNM Type

69784.

PARATYPES.—Same locality as for holotype, 2 $ & ,

2 9 9 , 25 June-2 July 1966 and 1967 (MOG preparation 20468, RWH slides 4528, 4529) [MOG,

USNM].

Sorhagenia baucidis is similar in maculation to both

nimbosa and daedala but differs in having a patch of

raised scales on the costal portion of the forewing at

one-fourth; the latter two species lack this patch.

Sorhagenia pexa, new species

FIGURES 4, 18, 20, 39

A small, light brown moth. Maculation as in Figure

4. Scales of head and inner suface of labial palpus pale

orange, a row of brown (tipped with pale orange)
10

scales bordering eye from base of antenna to tongue,

occiput slightly darker than frons; outer surface of second segment of labial palpus mottled brown, apex grayish yellow and pale orange, basal two-thirds of third

segment grayish yellow and pale orange, apical third

mottled brown; antenna mainly brown to four-fifths

then pale orange, apex mottled brown and pale

orange, ventral surface of scape pale orange. Thorax

pale orange with tegula, apex of mesothorax , and sides

of mesothorax brown. Forewing with 4 patches of

raised scales, 1 between fold and dorsal margin at onefourth, 1 just beyond one-half, 1 between fold and costa

at one-half, and 1 at end of cell, base of wing and

several streaks on costal half brownish black, an

oblique pale orange band from costa to tornus, beyond

this a series of 5 brown-black dots on costal margin and

2 on outer margin, cilia gray brown with pale orange

streak from end of oblique band at tornus. Hind wing

gray. Abdomen mottled brown and brownish orange

dorsally, brownish orange ventrally. Foreleg brown,

apex of coxa and tibia pale yellow, base of first and

second tarsal segments mottled brown, third segment

brown, remainder pale orange. Midleg with yellowwhite coxa; femur gray brown, apex brown; tibia

mottled light brown, grayish orange, and orange white,

apices of scales pale; base and apex of 1st, 2d, and 4th,

base of 3d, and all of 5th tarsal segments orange white,

remainder brown. Hind leg with yellowish white coxa

and femur, upper margin of femur gray brown; tibia

with oblique, brown band from two-fifths to four-fifths,

a brown streak near base and one just before apex, remainder of scales grayish orange tipped with orange

white, spurs mottled gray brown and orange white;

base and apex of first 4 tarsal segments and all of 5th

orange white, remainder brown. Alar expanse: 12-14

mm. Male genitalia: as in Figures 18 and 20 (RWH

slide 4055). Costal lobe of valva slender, slightly expanded before apex, medial lobe uniformly slender,

slightly enlarged at apex; saccular lobe with a broad

base then spatulate, inner surface densely covered with

long setae; uncus minutely trilobate; eighth sternite

sclerotized as horseshoe. Female genitalia: as in Figure

39 (RWM slide 4056). Ostium bursae at three-fourths,

projecting from seventh sternite; basal third of ductus

bursae heavity sclerotized, smooth walled, distal twothirds with granulose walls, corpus bursae lightly

sclerotized, walls with numerous spicules, signa absent,

but spicules more closely set in one region.

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

FOOD PLANT.—Unknown.

HOLOTYPE.—Male, Devil's Den State Park, Washington County, Arkansas, 20 June 1966, R. W. Hodges

(RWH slide 4055). USNM Type 69785.

PARATYPES.—Same locality as for holotype, 2 9 9 ,

9, 13 July 1966 (RWH slide 4056) [University of Arkansas, USNM]. 3 9 9 , Putnam County, Illinois, 6-^10

July 1963 and 1964, M. O. Glenn (MOG slide 24,

RWH slide 3166) [MOG, USNM].

Sorhagenia pexa may easily be separated from the

other species of Sorhagenia by the light brown color of

the forewings; gray to gray-black is the predominant

color of the latter. Genitalic differences are as given in

the key.

Genus Siskiwitia, new genus

TYPE-SPECIES.—Siskiwitia alticolans Hodges, new

species.

Head: smooth scaled, labial palpus porrect, third

segment three-fourths length of second segment, apex

acute; tongue short; maxillary palpus short; antenna

about three-fifths length of forewing, simple, pecten

absent. Forewing: broadly lanceolate, apex pointed;

11 veins present, Cu2 absent; M2 short stalked with

R^ + Rs + Mx; R4, R5, and Ma stalked, Mx to outer

margin, R6 to costa. Hindwing: linear; 8 veins present;

Ma and RB stalked. Male genitalia: valva with a moderately heavily sclerotized, slender lobe from costal

margin at two-thirds length of valva; articulating

mechanism between aedeagus and bases of valvae

short; vinculum apparently absent, valvae articulating

medially with one another; uncus heavily sclerotized,

rod shaped. Female genitalia: not known.

Siskiwitia is nearest Nepotula Hodges on characters of the male genitalia but differs in having a welldeveloped uncus. The venation differs from that of

Nepotula in that M2 is stalked with R4 + R5 + Mx in the

forewing and that R8 and Mi are stalked in the hind

wing. Raised scales are absent on the forewings of Siskiwitia, present on those of Nepotula.

Siskiwitia alticolans, new species

FIGURES 5, 19

A minute, shining black (forewing with a white bar)

species. Head: shining metallic silver and yellow silver

at certain angles of light incidence, otherwise nearly
NUMBER 18 11

black; tongue pale yellow. Thorax and forewing black, margin swollen, saccular margin nearly straight; aedeshining dark bronze at some angles of light incidence,

individual scales with nonlustrous margins, a transverse, white band at two-thirds length of wing. Hind

wing shining dark gray. Legs: (not all present) shining

dark gray to black, apex of hind tibia and all of tibial

spurs white. Alar expanse: 6 mm. Male genitalia: as in

Figure 19 (RWH slide 4423). Valva somewhat triangular, a moderately heavy sclerotized, slender lobe

from costal margin at two-thirds length of valva, costal

agus concave dorsally; uncus stout, pointed, heavily

sclerotized. Female genitalia: no specimens available.

FOOD PLANT.—Unknown.

HOLOTYPE.—Male, Arizona, Santa Cruz County,

Madera Canyon, 5,100 ft, Bog Spring Camp Ground,-

Santa Rita Mts., July 10-26, 1964, D. R. Davis (RWH

slide 4423, wing slide 111). USNM Type 69786.

Siskiwitia alticolans looks like a small Periploca that

has a white, transverse band on the forewing.

Genus Perimede Chambers

Key to the Species

1. Males 2

Females 8

2. Spines on glands associated with genitalia symmetrical 3

Spines on glands associated with genitalia asymmetrical 6

3. Glandular spines shorter than valvae 4

Glandular spines equal to or longer than valvae 5

4. Apex of valva drawn to point ventrally, spine curved at one-half to two thirds

circitor, new species

Apex of valva rounded; spine nearly straight, slightly curved at one-third . parilis, new species

5. Glandular spines sinuate, apex of valva rounded latris Hodges

Glandular spines straight, apex of valva drawn to point ventrally ... . erransella Chambers

6. Right glandular spine coiled battis Hodges

Right glandular spine not coiled 7

7. Apices of uncus blunt; valva narrow, saccular margin with an expanded shoulder near base. .

ricina Hodges

Apices of uncus acute; valva broader, saccular margin without a distinct shoulder near base. .

falcata Braun

8. Seventh sternite symmetrical 10

Seventh sternite asymmetrical 9

9. Ostium bursae at middle of 7th sternite ricina Hodges

Ostium bursae at apex of 7th sternite latris Hodges

10. Ostium bursae extending beyond posterior margin of 7th tergite, on raised prominence.

falcata Braun

Ostium bursae not extending beyond posterior margin of 7th tergite, on raised prominence or

not 11

11. Lamella antevaginalis narrowly triangular, extending beyond anterior margin of 7th sternite. •

parilis, new species

Lamella antevaginalis variously shaped, not extending beyond anterior margin of 7th sternite. 12

12. Ostium bursae open to ventral surface, lamella antevaginalis a sclerotized rim 13

Ostium bursae not immediately open to ventral surface, lamella antevaginalis extending beyond

ostium bursae ventrally making a flange 15

13. Ostium bursae laterally compressed; lamella antevaginalis with a narrow, medial slit, extending

to posterior margin of 7th sternite circitor, new species

Ostium bursae broad 14

14. Lamella antevaginalis broadest at center, ostium bursae nearly at posterior margin of seventh

sternite, no sclerotized band posterad of opening maniola, new species

Lamella antevaginalis evenly narrow, a sclerotized band between ostium bursae and posterior

margin of 7th sternite nemos, new species

15. A pair of invaginations containing scales anterior to ostium bursae battis Hodges

Lacking such invaginations erransella Chambers
12

Perimede erransella Chambers

Perimede erransella Chambers.—Hodges 1962c, p.147.

RECORDS.—Arkansas: Devil's Den, 8 S $, 3 2 2,

29 May-21 July 1966. Florida: Archbold, 1 5,1- 7

May 1964; Pensacola, 2 £ 5, 1 5, 24 July, 1 December 1961, 22 February 1962. North Carolina: Durham,

3 3 5,599, 4 December 1946-13 January 1947,

ex Carya tomentosa and Quercus coccinea.

Perimede eremos, new species

FIGURES 40, 42

A small, dark, gray-black species. Head shining offwhite; outer surface of second segment of labial palpus (except apex), a series of scales along dorsal

and anterior margins of eyes, and antenna dark

gray. Thorax with dark gray tegula, pro- and mesonota

medium gray, metanotum pale gray. Forewing mottled

dark gray; with four black spots, one on fold at twofifths length of wing, one at middle of wing, one at

end of cell, and one at apex of wing, first two followed

by gray-white scales, last two preceded by gray-white

scales; a white spot on costal margin and one on dorsal

margin of wing at three-fourths length of wing.- Hind

wing slightly paler than forewing. Proleg: coxa mottled

pale and medium gray, appearing light; femur and

tibia dark gray, apex of epiphysis and tibia off-white;

a dark gray saddle on first, second, and fourth tarsal

segments, all of third segment dark gray, remaining

elements off-white. Midleg: mainly off-white to white;

coxa and femur speckled with medium gray; basal

three-fifths of tibia dark gray dorsally, a few medium

gray scales on distal two-fifths and on spurs, tarsus

with a few scattered gray scales. Hind leg: coxa and

femur white with a few dark gray—almost black—

scales, these more numerous at base and apex of femur;

tibia medium gray, a few white scales at base of middle

tibial spurs, apex and spurs white; tarsus darker than

tibia, base and apex of first four and all of fifth segment white. Abdomen: tergum medium gray, apices

of segments slightly paler, apex of sixth tergite offwhite; sternum mottled off-white and gray, mainly

off-white. Alar expanse: 7-8 mm. Male genitalia:

no specimens available. Female genitalia: as in

Figures 40 and 42 (RWH slide 4086). Ostium bursae from middle to near caudal margin of seventh

sternite, irregularly round tosubquadrate; lamella antevaginalis a narrow, heavily sclerotized band; ductus

bursae slender, lightly sclerotized; corpus bursae memSMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

branous, signa a pair of inwardly pointing spurs, bases

on wall of corpus bursae lightly and evenly sclerotized;

anterolateral margins of seventh sternite lightly

sclerotized; extending around sides as a pair of flaps.

FOOD PLANT.—Unknown.

HOLOTYPE.—Female, Devil's Den State Park, Washington County, Arkansas, 27 June 1966, R. W. Hodges

(RWH slide 4086). USNM Type 69788. Paratype:

One 9, same locality as for holotype, 17 June 1966

[USNM].

Consistent characters to separate eremos from other

species of Perimede are the symmetrical female genitalia with the ostium bursae being roughly quadrilateral and the signa being simple, stout projections.

Perimede ricina Hodges

Perimede ricina Hodges, 1962c, p. 150.

RECORDS.—Falls Church, Virginia, 19, 9 July 1962.

Perimede parilis, new species

FIGURES 6, 21-23,25,41

A small, dark, gray-black moth. Maculation: as in,

Figure 6. General color pattern is much as for eremos.

Head usually shining pale to medium gray; inner surface of labial palpus shining orange white, outer surface dark gray; base of tongue medium to dark gray;

several half-segments on dorsal surface of antenna pale

gray. Forewing with a yellowish white patch in costal

scales just before apex; a few whitish scales on apical

one-fifth of wing; and an incomplete "V" of whitish

scales connecting white spots at three-fourths length of

wing, point of "V" toward apex. Proleg: coxa gray,

apex yellowish white; femur dark gray, base and apex

white; tibia dark gray, area opposite epiphysis and

apex white; tarsus dark gray, base and apex of first

two segments, apex of fourth, and all of fifth segment

white, ventral surface yellowish white to white. Midleg: coxa yellowish white with a few pale gray scales;

femur medium gray, base yellowish white; tibia dark

gray basally, yellowish white distally; spurs and distal

half with scattered pale gray scales; tarsus mainly yellowish white, a few dark gray scales near apex of first

segment, a dark gray saddle on middle of second, on

distal two-thirds of third, and basal two-thirds of fourth

segment. Hind leg: coxa yellowish white mottled with

pale gray; femur yellowish white, dark gray (almost

black at base and apex); tibia gray, apex, a transverse

streak at one-half, and spurs yellowish white, spurs
NUMBER 18 13

with dark gray scales just before apex; tarsus dark gray,

base and apex of first four segments and all of fifth

segment yellowish white. Alar expanse: 8-12 mm. Male

genitalia: as in Figures 21-23 and 25 (RWH slides

4057, 4087). Valva angled at middle, distal half with

relatively parallel costal and saccular margins, slightly

expanded at apex, apex rounded; aedeagus expanded

for basal two-fifths then sharply constricted, tapering

gradually to acute apex, curved downward; saccus

broad; lobes of uncus slender, parallel sided, apices

slightly bent at one-third; lightly sclerotized lobe with

setae at apex. Female genitalia: as in Figure 41 (RWH

slide 4090). Ostium bursae extending from middle

nearly to posterior margin of seventh sternite, opening

lightbulb-shaped; lamella antevaginalis narrowly triangular; ductus bursae short, relatively broad, expanding gradually to corpus bursae; corpus bursae lightly

sclerotized, walls with numerous, fine lines; signa a

pair of inwardly projecting spines arising from an elliptical, sclerotized patch on wall of corpus bursae.

FOOD PLANT.—Unknown.

HOLOTYPE.—Male, Devil's Den State Park, Washington County, Arkansas, 10 July 1966, R. W. Hodges

(RWH slide 4057). USNM Type 69787.

PARATYPES.—4 $ S, 5 9 2, same locality as for holotype, 21 June-14 July 1966 (RWH slides 4087, 4088,

4090, 4435-7, 4440-2) [University of Arkansas,

USNM]. 3 $$, 4 $2, Putnam County, Illinois, 7

June-14 August 1961 to 1967, M. O. Glenn (MOG

slides 18-20, 19168, 19368, 20068; RWH slide 4532)

[MOG, USNM].

Some variation occurs in the amount of light or dark

scales on the head, with individuals appearing very

light to dark gray. Fresh specimens generally have the

vertex darker than worn ones. Flown specimens take

on a brownish cast as the overlying dark gray scales

on the wings are worn away.

The combination of characters, glandular spines

symmetrical, apices of valva rounded, and lobes of

uncus linear, will separate males of parilis from those

of other species of Perimede. In turn, the females may

be distinguished by the lamella antevaginalis being

narrow and extending from the middle of the seventh

sternite to beyond the anterior margin of this segment.

Perimede circitor, new species

FIGURES 24, 26, 43, 45

A small, dark gray species. Head: occiput medium

gray, irons and lower part of vertex pale gray; inner

surface of labial palpus light orange, outer surface

dark gray (scale bases pale), third segment about

three-fourths length of second; base of tongue mottled

pale gray and orange white; thorax and wings as for

eremos. Foreleg: coxa medium gray, apex yellowish

white; femur slightly darker gray, apex narrowly pale

gray; tibia dark gray, a white streak opposite epiphysis,

apex white; tarsus nearly black, base and apex of first,

apex of second and fourth, and all of fifth segment

white. Midleg: coxa yellowish white; femur yellowish

white basally becoming pale gray distally; tibia mottled

yellowish white and pale gray, dark gray basally, spurs

yellowish white with a dark gray ring just before apex;

tarsus mainly yellowish white, second through fourth

segments with dark gray saddles. Hind leg: Coxa mottled yellowish white and medium gray; femur yellowish white, heavily dusted with dark gray at base and

apex; tibia mottled pale and dark gray (scale bases

pale), a transverse, pale gray streak at base of first pair

of tibial spurs and at apex pale gray to off-white,

spurs off-white, ringed with gray before apices; tarsus

dark gray, base and apex of first, apices of second, third,

and all of fifth segment off-white. Alar expanse: 7-10

mm. Male genitalia: as in Figures 24 and 26 (RWH

slide 4085). Valva relatively linear, tapering gradually

from base to two-thirds length, apex drawn to a point

on ventral margin; apices of uncus very slender on distal half, apices rounded; aedeagus slender, apex

slightly downturned. Glands associated with genitalia

bilobate, ventral one with series of setae on outer onethird of inner surface; spines slender, tapering fairly

evenly from base to apex, curved near middle. Seventh

sternite with heavily sclerotized margin; eighth sternite

composed of three sclerites in shape of tulip, lateral

sclerites relatively slender, medial one "T" shaped,

cross of "T" broad, base slender; eighth tergite shaped

like a low margined "U," sclerite narrowest medially.

Female genitalia: as in Figures 43 and 45 (RWH slide

4089). Ostium bursae narrow, at posterior margin of

seventh sternite; lamella antevaginalis with a narrow

medial slit; ductus bursae very narrow from ostium

bursae; expanding quickly to corpus bursae; corpus

bursae with a slight pouch near base, walls with fine

creases; signa two, slender, inwardly directed pyramids, lightly sclerotized base elliptical with a tearshaped border from origin of pyramid.

FOOD PLANT.—Unknown.

HOLOTYPE.—Female, Devil's Den State Park, Wash-
14

ington County, Arkansas, 18 July 1966, R.W. Hodges

(RWH slide 4089). USNM Type 69790.

PARATYPES.—1 $ , 3 9 9, same locality as for holotype, 3 June-22 July 1966 (RWH slides 4085, 4438,

4439, 4443) [University of Arkansas, USNM]. 3'8 8 ,

Putnam County, Illinois, 15 July, 2 August 1966 and

1967, M. O. Glenn (MOG preparation 20168, RWH

slides 4530,4531) [MOG, USNM].

The shape of the valva is much like that of erransella; however, the apices of the uncus being rounded

rather than acute, the glandular spine being curved

and arising from a broad lobe rather than being

straight and arising from a goosenecked apex of a lobe

will easily separate males of the two species. Females

of circitor may be separated from those of other species

of Perimede by the lamella antevaginalis having a narrow, medial slit and by the corpus bursae having a

lateral, basal expansion.

Perimede maniola, new species

FIGURES 44, 46

A small, dark gray-brown species. Head: base of

tongue and frons shining, pale gray, vertex and occiput

becoming medium to dark gray; outer surface of labial

palpus dark gray brown, base of third segment pale,

inner surface pale orange to orange white; antenna

gray, some scales of shaft pale. Thorax: dark gray,

metanotum pale gray. Forewing: dark gray brown,

scale bases pale; four indistinct solid dark spots, followed and preceded by off-white scales as for eremos,

apical dark spot set off distally by pale scale bases; a

faint gray-white costal patch at three-fourths length

of wing. Hindwing: pale to medium gray. Foreleg:

coxa pale yellow white and medium gray; femur medium gray; tibia dark gray, apex and a small patch at

apex of epiphysis off-white; tarsus dark gray, base

and apex of first two, base of third, apex of fourth, and

all of fifth segment off-white. Midleg: (worn on

all specimens) appearing yellowish white with gray

and dark gray scales. Hind leg: coxa mottled yellowish white, medium gray, and dark gray brown, femur

pale yellowish white, base and apex dark gray brown;

tibia dark gray, off-white at apex and at base of first

pair of spurs, tibial spurs off-white, dark gray just before apex; tarsus dark gray, base and apex of first four

and all of fifth segment pale yellowish white. Alar

expanse: 9.0-10.5 mm. Male genitalia: no specimens

available. Female genitalia: as in Figures 44 and 46

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

(MOG slide 22). Ostium bursae just before posterior

margin of seventh sternite, rounded lamella antevaginalis roughly diamond shaped, lateral edges extended

posteriorly; ductus bursae expanding rapidly to fuse

with corpus bursae, no separation between the two;

corpus bursae lightly sclerotized, walls with numerous,

fine lines; signa two inwardly projecting cones.

FOOD PLANT.—Unknown.

HOLOTYPE.—Female, Putnam County, Illinois,

August 19, 1965, M. O. Glenn (MOG slide 22).

USNM Type 69789.

PARATYPES.—2 9 9, same locality as for holotype,

July 11 and August 17, 1965 (MOG slides 21, 23)

[MOG, USNM].

This is an evenly colored species which, as it becomes worn, turns pale. The female genitalia of

maniola are nearest those of eremos but may easily be

separated as indicated in the key.

Perimede battis Hodges

Perimede battis Hodges, 1962c, p. 153.

RECORDS.—Arizona: Pine, 1 $ , 1 9,1, 5 September

1961.

Genus Ithome Chambers

Ithome concolorella (Chambers)

Ithome concolorella (Chambers).—Hodges, 1962a, p. 85.

RECORDS.—Arkansas: Devil's Den, 174 $ $,

124 9 9, 19 May-23 July 1966.

This species was very common over most of the time

that I collected in Arkansas. Late one afternoon I observed several adults as they were resting and flying

around the inflorescences of Schrankia nuttallii (DC)

Standley. I judge that the larvae feed on the developing flowers and fruits of this plant.

Ithome curvipunctella (Walsingham), new

combination

Anybia curvipunctella Walsingham, 1892, p. 538.

Eriphia quinquepunctata Forbes, 1931, p. 361. [New

synonymy.]

Ithome quinquepunctata (Forbes).—Hodges, 1962a, p. 88.

I have dissected and examined the genitalia of the

female holotype of curvipunctella in the British

Museum (NH) and so am able to recognize the given

synonymy.
NUMBER 18 15

lthome edax Hodges

Ithome edax Hodges, 1962a, p. 88.

When this species was described, a question was

present as to the plant species associated with the

common name ebony. Subsequently, it has been suggested that what was meant is Texas ebony (Pithecollobium flexicaule (Bentham) Coulter).

List of Nearctic Walshiidae

(Synonyms in italics)

Periploca Braun, 1919

orichalcella (Clemens), 1864

concolorella (Chambers), 1875b

purpuriellaBTaun, 1919

ceanothiella (Cosens), 1908

gleditschiaeella (Chambers), 1876

hostiata, new species

hortatrix, new species

opinatrix, new species

atrata Hodges, 1962b

mimula Hodges, 1962b

laeta Hodges, 1962b

cata Hodges, 1962b

devia, new species

nigra Hodges, 1962b

fessa Hodges, 1962b

gulosa Hodges, 1962b

facula Hodges, 1962b

funebris Hodges, 1962b

dipapha, new species

iabes, new species

Synploca Hodges, 1964

gumia Hodges, 1964

Neoploca Hodges, 1964

corusca Hodges, 1964

Walshia Clemens, 1964

particornella (Busck), 1909

amorphella Clemens, 1864

miscecolorella (Chambers), 1875b

miscecalonella (Chambers), 1875a

dispar Hodges, 1961

similis Hodges, 1961

exemplata Hodges, 1961

Nepotula Hodges, 1964

secura Hodges, 1964

Aeaea Chambers, 1874b

Amaurogramma Braun, 1919

juvantis Hodges, 1964

dulcedo Hodges, 1964

venifica Hodges, 1964

venatrix Hodges, 1964

ostryaeella Chambers, 1874b

quadricustatella Chambers, 1880

stipator Hodges, 1964

risor Hodges, 1964

victor Hodges, 1964

placatrix, new species

sagana Hodges, 1964

rhynchosiae Hodges, 1964

ex tens a (Braun), 1919

Stilbosis Clemens, 1860

tesquella Clemens, 1860

quinquicristatelta (Chambers), 1881

nubila Hodges, 1964

lonchocarpella Busck, 1934

Chrysopeleia Chambers, 1874b

purpuriella Chambers, 1874b

Sorhagenia Spuler, 1910

Cystioecetes Braun, 1915

nimbosa (Braun), 1915

nubila Hodges, 1964

baucidis, new species

pexa, new species

Siskiwitia, new genus

alticolans, new species

Perimede, Chambers, 1874a

erransella Chambers, 1874a

falcata (Braun), 1919

eremos, new species

ricina Hodges, 1962c

parilis, new species

circitor, new species

maniola, new species

latris Hodges, 1962c

battis Hodges, 1962c

Ithome Chambers, 1875c

concolorella (Chambers), 1875b

unimaculella Chambers, 1875c

curvipunctella (Walsingham), 1892

quinquepunctata (Forbes), 1931

edax Hodges, 1962a

lassula Hodges, 1962a

ferax Hodges, 1962a

Obithome Hodges, 1964

punctiferella (Busck), 1906

Literature Cited

Braun, Annette F.

1915. New Genera and Species of Tineina. Canadian

Entomologist, 47:188-197.

1919. Notes on Cosmopterygidae, with Descriptions of

New Genera and Species (Microlepidoptera). Entomological News, 30:260-264.

Busck, August

1906. Tineid Moths from Southern Texas, with Description of New Species. Proceedings of the U.S. National Museum, 30( 1465):721-736.

1909. Notes on Microlepidoptera, with Descriptions of

New North American Species. Proceedings of the

Entomological Society of Washington, 11: 87-103.

1934. Microlepidoptera of Cuba. Entomological Americana, 13 (newseries) (4): 151-203.
16 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Chambers, V. T.

1874a. Micro-Lepidoptera. Canadian Entomologist, 6:

49-52.

1874b. Micro-Lepidoptera. Canadian Entomologist, 6:

72-77.

1875a. Tineina from Texas. Canadian Entomolgist, 7:

30-35.

1875b. Tineina from Texas. Canadian Entomologist, 7:

51-56.

1875c. Tineina from Texas. Canadian Entomologist, 7:

92-95.

1876. Tineina. Canadian Entomologist, 8:135-138.

1880. Descriptions of Some New Tineina, with Notes on

a Few Old Species. Journal of the Cincinnati Society of Natural History, 2:179-194.

1881. New Species of Tineina. Journal of the Cincinnati

Society of Natural History, 3:289-296.

Clemens, Brackenridge

1860. Contributions to American Lepidopterology—No.

4. Proceedings of the Academy of Natural Sciences

of Philadelphia, 1860:156-174.

1864. North American Microlepidoptera. Proceedings of

the Entomological Society of Philadelphia, 2:415-

430.

Cosens, A.

1908. A New Lepidopterous Gall-Producer. Canadian Entomologist, 40:107-108.

Forbes, Wm. T. M.

1931. Supplementary Report of the Heterocera or Moths

of Porto Rico. Journal of the Department of Agriculture of Porto Rico, 4(4): 339-394.

Hodges, Ronald W.

1961. A Review of the Genus Walshia Clemens with Descriptions of New Species (Lepidoptera: Gelechioidea). Bulletin of the Brooklyn Entomological

Society, 56:66-80.

1962a. The Genus Ithome in North America North of

Mexico (Walshiidae). Journal of the Lepidopterists'

Society, 15:81-90.

1962b. A Review of the Genus Periploca with Descriptions

of Nine New Species (Lepidoptera: Gelechioidea).

Pan-Pacific Entomologist, 38:83-97.

1962c. The Genus Perimede Chambers in North America

North of Mexico. Proceedings of the Entomological

Society of Washington, 64:145-154.

1964. A Review of the North American Moths of the

Family Walshiidae (Lepidoptera: Gelechioidea).

Proceedings of the U.S. National Museum, 115

(3485):289-330.

Spuler, Arnold

1910. Die Schmetterlinge Euro pas. Volume 2. Stuttgart.

Walsingham, Lord [deGrey, Thomas]

1892. On the Micro-Lepidoptera of the West Indies. Proceedings of the Zoological Society of London, 1891:

492-549.
NUMBER 18 17

FIGURES 1-3.—1, Periploca hortatrix, new species, Devil's

Den State Park, Arkansas; 2, Stilbosis lonchocarpella Busck,

Key Largo, Florida; 3, Sorhagenia baucidis, new species, Putnam County, Illinois.
18 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

FIGURES 4-6.—4, Sorhagenia pexa, new species, Devil's Den

State Park, Arkansas; 5, Siskiwitia alticolans, new species,

Madera Canyon, Santa Rita Mountains, Arizona; 6, Perimede

parilis, new species, Devil's Den State Park, Arkansas.
NUMBER 18 19

FIGURES 7-10.—Male genitalia: 7, Periploca hostiata, new

species; 8-9, P. opinatrix, new species; 10, P. hortatrix, new

species.
20 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

FIGURES 11-14.—Male genitalia: 11, Periploca devia, new

species; 12-13, P. dipapha, new species; 14, P. labes, new

species.
NUMBER 18 21

FIGURES 15-17.—Male genitalia: 15, Stilbosis lonchocarpella

Busck; 16—17, Sorhagenia baucidis, new species (16, eighth

stemite).
22 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

FIGURES 18-20.—Male genitalia: 18, 20, Sorhagenia pexa,

new species (20, eighth sternite); 19, Siskiwitia alticolans,

new species.
NUMBER 18 23

FIGURES 21-23.—Male genitalia: Perimede parilis, new

species, 21, genitalia with accessory glands removed; 22

genitalia with accessory glands in place; 23, accessory gland.
24 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

FIGURES 24-26.—Male genitalia: 24, 26, Perimede circitor,

new species (26, eighth abdominal segment); 25, Perimede

parilis, new species, eighth abdominal segment.
NUMBER 18 25

FIGURES 27-29.—Female genitalia: 27, Periploca opinatrix,

new species; 28-29, P. hortatrix, new species (29, ostium

bursae and bursa copulatrix).
26 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

FIGURES 30-33.—Female genitalia: 30-32, Periploca devia,

new species (31, genital plate, 32, corpus bursae); 33, P. labes,

new species.
NUMBER 18 27

FIGURES 34—36.—Female genitalia: 34, Stilbosis lonchocarpella Busck; 35, Aeaea placatrix, new species; 36, A. venifica

Hodges.
28 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

FIGURES 37-39.—Female genitalia: 37-38, Sorhagenia

baucidis, new species (37, genital plate and ductus bursae);

39, S. pexa, new species.
NUMBER 18 29

FIGURES 40-43.—Female genitalia: 40, 42, Perimede eremos,

new species; 41, P. parilis, new species; 43, P. circitor, new

species.
30

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

FIGURES 44—46.—Female genitalia: 44, 46, Perimede maniola,

new species (46, eighth abdominal segment); 45, P. circitor,

new species, eighth abdominal segment and bursa copulatrix.

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