THOMAS BORGMEIER Bredin-A rchbo Id

Smithsonian Biological

^Survey of Dominica:

f The Phoridae of

Dominica (Diptera)

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY • 1969 NUMBER 23
SERIAL PUBLICATIONS OF THE SMITHSONIAN INSTITUTION

The emphasis upon publications as a means of diffusing knowledge was expressed

by the first Secretary of the Smithsonian Institution. In his formal plan for the Institution, Joseph Henry articulated a program that included the following statement:

"It is proposed to publish a series of reports, giving an account of the new discoveries

in science, and of the changes made from year to year in all branches of knowledge

not stricdy professional." This keynote of basic research has been adhered to over the

years in the issuance of thousands of titles in serial publications under the Smithsonian

imprint, commencing with Smithsonian Contributions to Knowledge in 1848 and continuing

with the following active series:

Smithsonian Annals of Flight

Smithsonian Contributions to Anthropology

Smithsonian Contributions to Astrophysics

Smithsonian Contributions to Botany

Smithsonian Contributions to the Earth Sciences

Smithsonian Contributions to Paleobiology

Smithsonian Contributions to Zoology

Smithsonian Studies in History and Technology

In these series, the Institution publishes original articles and monographs dealing

with the research and collections of its several museums and offices and of professional

colleagues at other institutions of learning. These papers report newly acquired facts,

synoptic interpretations of data, or original theory in specialized fields. Each publication is distributed by mailing lists to libraries, laboratories, institutes, and interested

specialists throughout the world. Individual copies may be obtained from the Smithsonian Institution Press as long as stocks are available.

S. DILLON RIPLEY

Secretary

Smithsonian Institution
SMITHSONIAN CONTRIBUTIONS TO

ZOOLOGY

NUMBER 23

Thomas Borgmeier

Smithsonian Biological

Survey of Dominica:

The Phoridae of

Dominica (Diptera)

ISSUED

NOV18 1969

SMITHSONIAN INSTITUTION PRESS

CITY OF WASHINGTON
ABSTRACT

Recent collecting by J. F. G. Clarke, O. S. Flint, Jr., R. J. Gagn6, P. Spangler, T. J.

Spilman, G. C. Steyskal, and W. W. Wirth for the Bredin-Archbold-Smithsonian

Biological Survey of Dominica resulted in a small but representative collection of

phorid flies, comprising 16 genera and 82 species, 43 of which are new. Of the

latter, 32 belong to the giant genus Megaselia. One new species from Costa Rica

has been added: Pachyneurella haplopyga.

Official publication date is handstamped in a limited number of initial copies and is recorded in

the Institution's annual report, Smithsonian Year.

UNITED STATES GOVERNMENT PRINTING OFFICE

WASHINGTON : 1969

For sale by the Superintendent of Documents, U.S. Government Printing Office

Washington, D.C. 20402 - Price 70 cents (paper cover)
Thomas Borgmeier Bredin-ArchboldSmithsonian Biological

Survey of Dominica:

The Phoridae of

Dominica (Diptera)

I wish to thank Dr. Willis W. Wirth for the opportunity to study this interesting material. My thanks are

also due to the "Conselho Nacional de Pesquisas," Rio

de Janeiro, for financial help in the form of a fellowship.

Abbreviations used in the text are as follows: USNM

(United States National Museum); MGZ (Museum

of Comparative Zoology); CTB (Collection of

Thomas Borgmeier); Academy of Natural Sciences

of Philadelphia (ANSP); su (supraantennal bristles).

Subfamily PHORINAE

Genus Dohrniphora Dahl

Dohrniphora cavifemur, new species

FIGURES 1, 2

FEMALE.—Length (combined) 2.5 mm. Head

width 0.65 mm. From pentagonal, about as broad as

long medially, width 0.36 mm, black-brown, yellowish

around bases of supraantennals, bristles of first and

second row placed in large circular yellowish spots.

Supraantennals divergent. First row straight, antials

more approximate than preocellars, a little farther

from each other than from anterolaterals, the latter

are removed from eye-margin. Second row slightly concave anteriorly, the preocellars farther from each other

than from mediolaterals. One strong lower postocular

bristle; one bristle on jowls; one weaker bristle on

cheeks. Third antennal joint small, suboval, dark

brown; arista long (0.70 mm), finely but distinctly

pubescent. Palpi yellow, moderately broad, length

without stem 0.18 mm, with 5 strong bristles near apex,

one of which is prolonged. Proboscis yellow, slightly

Reverend Thomas Borgmeier, OJF.M., Estrada Rio Grande

2116, Jacarepagud, G.B., Rio de Janeiro ZC-89, Brazil.

geniculate, basal part short, apical part (haustellum)

long (0.46 mm) and narrow (0.1 mm), nearly straight.

Thorax dark brown. Mesopleural hairs not perceptible in the micropreparation but probably present

above, as usually. Scutellum apparently with 4 unequal

bristles, the posterior are broken in type, the anterior

is 0.16 mm long.

Abdomen (Figure 1) dark brown, with 5 tergites.

Tergites 3-5 gradually narrowed, 5th elongate triangular. Pubescence scarce and short, more abundant

on segments 7-8; 7th segment on each side with a

chitinous plate. Cerci small.

Legs yellow, midcoxae fuscous, hind femur darkened apically. Foretibia 0.5 mm, with 4 to 5 dorsal

bristles. Midtibia with the usual basal pair of bristles

near base and one short anterior subapical bristled,

the dorsal hair-seam reaches the apical third. Hind

femur (Figure 2) about 3 times as long as broad, dorsally near base with a deep excavation. Hind tibia with

a complete dorsal hair-seam; the anterodorsal and

posterodorsal cilia fine.

Wing distinctly yellowish, anterior margin and apical margin of membrane infuscated. Length 2.15 mm,

width 0.98 mm. Costa 0.54 of wing length, ratio of

segments 15:4:2. Cilia short. Fork narrow. First vein

with the characteristic angle behind middle; 3rd vein

with a distinct bristle at base; 4th vein slightly concave;

5th nearly straight, a little upturned apically; 6th

slightly sinuate; 7th weak. Auxiliary margin with one

hair. Halteres dirty yellow.

MALE.—Similar to the female. Antenna brown,

small, short oval; arista 0.60 mm long. Scutellum with

2 bristles and 2 fine short hairs. Abdominal tergites 2

and 6 elongated, yellow on anterior half; 3-5 with a

broad yellow median spot anteriorly. Hypopygium

dark brown, at left with a moderate bristle. Anal tube

yellow, cylindrical, length 0.23 mm, apical hairs not

1
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

differentiated. Legs as in female, the basal excavation

of hind femora a little less pronounced, obtuse-angled

instead of rectangular; posterior face of hind femur

near base with two black obtuse erect spines very close

together. Wing length 1.70 mm, costa 0.53 of wing

length, ratio of segments 11:3:1.5.

MATERIAL EXAMINED.—Holotype female (USNM

70236, on slide), Dominica, Clarke Hall June 1964,

light trap (O. S. Flint). Allotype male (USNM, on

slide), 10-20 April 1964, same locality and collector.

DISCUSSION.—Dohrniphora cavifemur is a very

striking species, both sexes are easily recognized by the

deep dorsal excavation near base of the hind femur.

Dohrniphora divaricata (Aldrich)

FIGURE 3

Phora divaricata Aldrich, 1896, p. 437 [male, female; St.

Vincent, B.W.I.].—Brues, 1904, p. 349.

Dohrniphora divaricata (Aldrich).—Borgmeier, 1961, p. 111.

Dohrniphora venusta (Coquillett).—Malloch, 1912, p. 432

[part].

Dohrniphora diffusa Borgmeier, 1960, p. 281, fig. 38 [$ $ ;

Brazil]; 1961,p. Ill [synonymy].

Holotype male (USNM 7765) and 3 paratypes

(USNM, CTB), from St. Vincent, B.W.I. (J. M. Aidrich leg.).

MALE.—Length 1.8-2 mm. Frons pentagonal,

broader than long laterally (4:3), black-brown or

black, nearly opaque, without median line. First row

straight, the antials as far apart as the preocellars,

anterolaterals a little removed from eye. Second row

slightly concave, the bristles equidistant. One lower

postocular bristle, one bristle on jowls and one bristle

on cheeks. Third antennal joint normal, yellowish

brown; arista long, distinctly pubescent. Palpi reddish

yellow, small, with short bristles.

Thorax pale brown to brown, pleura brown above,

pale yellow below, sometimes entirely brown. Mesopleura hairy above. Scutellum with 4 bristles, the anterior hairlike and hardly one-half of the posterior.

Abdomen opaque, venter yellow, tergites black with

yellow apical seams, 2 and 6 elongated. Pubescence

scarce and short. Hypopygium large, dark brown, with

fine longish hairs; anal segment long and stout, nearly

parallel-sided, with short hairs.

Legs yellow, hind femur darkened at tip, midcoxa

brown. Foretibia with 4 or 5 dorsal bristles; foretarsus

very slender. Hair-seam of midtibia attaining apical

third. Hind femur broad, ventral edge sharpened, near

the greatest width on posterior face with a flat groove

and anterior to it with about 30-35 short black sensorial pins (Figure 3). Hind tibia without single

bristles except on apex.

Wing with grayish yellow turbidity, veins dark

brown. Costa about 0.52 of wing length, ratio of segments 25:7:3. Cilia short. Fork very narrow. Third

vein with a bristle at base; 4th weakly curved; 5th

straight, slightly upturned apically. Axillary margin

with one hair. Halteres yellow.

FEMALE.—Length 2-2.5 mm. Proboscis stout, nearly

straight. Abdomen black above, venter yellow; there

are 5 tergites, 3-5 gradually narrowed, all with yellow

apical seams. Hind femur normal. Wing as in male.

MATERIAL EXAMINED.—The holotype and 2 paratypes in USNM and one paratype male in my collection, from St. Vincent, B.W.I. (Aldrich). From

Dominica: Clarke Hall, 7 males and 2 females, taken

with Malaise trap and light trap, IV.64 (O. S. Flint)

and I.-III.65 (W. W. Wirth).

DISTRIBUTION.—Antilles, Central and South

America down to Argentine (see localities under

D. diffusa, in Borgmeier, 1960).

DISCUSSION.—Dohrniphora divaricata is very close

to D. perplexa Brues (1904, USA), but in the latter

there are not so many sensorial pins on hind femur of

male, the pleura are pale yellow and the anterior scutellars are about two-thirds of the posterior in length (see

Borgmeier, 1963a, p. 67, fig. 74).

NOTE.—D. divaricata has been confused for a long

time with D. venusta Coquillett (which is a synonym of

cornuta Bigot). Even Malloch (1912), who could

examine the types of divaricata and venusta, failed to

see that they are different, probably because he did

not examine the posterior face of the hind femur of

the male. As a matter of fact, cornuta has only 4 or 5

sensorial pins of unequal length on the hind femur,

and the female has only 4 tergal plates.

Dohrniphora dispar (Enderlein)

FIGURE 4

Phora dispar Enderlein, 1912, p. 20 [male; Brazil].

Dohrniphora dispar (Enderlein).—Schmitz, 1923, p. 55.—

Borgmeier, 1925b, p. 100; 1935a p. 431, figs. 2 [foreleg], 3

[head of male], 5 [hind femur], pi. 5: fig. [wing of type];

1960, p. 276, fig. 27 [3 9].

Holotype male and one paratype male (CTB),

Hammonia, S. Catarina, Brazil (Luederwaldt).
NUMBER 2 3

This is a very striking species, easily distinguished

by the following characters: antenna of male orange,

enlarged, about two-thirds eye-length. Pleura and all

coxae yellow; abdomen nicely cross-banded with yellow. Foretarsus of male slender, last joint fuscous,

slightly dilated and longer than preceding; hind femur

on posterior face near base with 50-60 blunt spinules

and one oval groove (Figure 4). Wing (see Borgmeier,

1935a, pi. 5: fig. 4): costa 0.51 segments 33:8:3, i.e.

1st section 3 times as long as 2+3. Female with 5 tergal

plates.

MATERIAL.—46 males and 14 females, Dominica:

Hodges River Mouth, Manets Gutter, South Chiltern,

Clarke Hall, Fond Figues River, I.-III. 1965, light trap

and Malaise trap (W. W. Wirth).

DISTRIBUTION.—Brazil, Argentine, Panama, Ecuador, Trinidad, Dominica W.I.

Dohrniphora cornuta (Bigot)

FIGURE 5

Phora cornuta Bigot, 1857, p. 827. [French edition: 1857,

p. 827]. Schmitz, in Lindner, 1938-1958 (1951), p. 241

[literature and synonyms].—Borgmeier, 1960, p. 277;

1963a, p. 65.

In my redescription of this species I failed to mention the short sparse hairs near dorsal border on the

posterior face of the hind femur; ventrobasally there

are only 4 to 5 black spinules two of which are longer

(Figure 5). The upper part of the pleura and the midcoxae are dark brown; there is a yellow cross-band

on 6th tergite of male. First costal section generally

3 times as long as 2 + 3. Female with 4 tergal plates.

MATERIAL EXAMINED.—Nine females and 23 males,

Dominica: Clarke Hall, Cabrit Swamp, Woodford

Hill, South Chiltern, Manets Gutter, I.-III. 1965, with

light trap and Malaise trap (W. W. Wirth).

Genus Coniceromyia Borgmeier

Coniceromyia Borgmeier, 1963b p. 450 (key).

Coniceromyia pilicoxa, new species

FIGURES 6, 7

MALE.—Length 2.6 mm. Head width 0.78 mm.

From flat, broader than long laterally (5:4), width

0.41 mm, black-brown, shining, finely shagreened, with

a fine median line and fine scattered hairs. Supraantennals weak, divergent. First row of frontal bristles

slightly convex, antials parallel, a little farther from

each other than from anterolaterals, somewhat removed from eye-margin. Second row slightly convex,

the bristles nearly equidistant. One strong lower postocular bristle directed downward; one weak bristle on

jowls; cheeks bare. Antennal grooves flat, not separated, reddish brown. Third antennal joint retortshaped, length 0.49 mm, reddish brown at base, apical

half blackish, the whole joint covered by numerous

whitish microtrichia; arista (broken in type) apical.

Palpi pale yellow, small, length 0.22 mm, with 4 short

bristles near apex.

Thorax shining, reddish yellow anteriorly, gradually

darkened behind, castaneous, pleura yellow; mesopleura bare. Scutellum black castaneous, with 2 long

posterior bristles and 2 shorter anterior bristles, the

later about two-fifths of the former in length.

Abdomen opaque, venter yellow, tergites 2-6 black,

2 and 6 slightly elongated, 3-5 subequal; 1st tergite

black, with the anterior and posterior borders yellow.

Pubescence scarce and short. Hypopygium large, globose, dark castaneous, ventral part brown.

Legs yellow, hind femur blackish at tip. Foretrochanter with a tuft of black curved hairs. Anterior face

of fore- and midcoxae and anterior face of midfemora

covered with dense long fine woolen sensorial hairs.

Foretibia (Figure 7) 0.78 mm, with 2 unequal dorsal

bristles on basal half and a series of short anterodorsal

setulae on apical half. Foretarsus scarcely longer than

tibia, relative length of tibia and tarsal joints 28:11:

7:6:5:4. Fore-metatarsus (Figure 7) thickened, as

broad as tibia, with a bare anteroventral furrow, apically without the slender process present in most species, instead with a fine curved bristle. Midtibia with

the usual basal pair of bristles and a short anterior

bristle near apex. Hind femur moderately broadened

at basal third, 3 times as long as broad, on posterior

face ventrobasally with a marginal strip of extremely

short black spinuli. Hind tibia with a long dorsal bristle

near base and one very short anterodorsal bristle near

apex.

Wing (Figure 6) grayish yellow, veins dark brown.

Length 2.78 mm, width 0.11. Costa not thickened,

about as thick as 3rd vein, index 0.53, ratio of segments 31:20. Cilia moderately short (about 0.1 mm),

closely set (there are about 9 between 1st and 3rd

veins). Third vein with a bristle at base; 4th concave,

deflected at tip; 5th but little curved before middle; 6th
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

slightly bisinuate; 7th weak. Axillary margin with one

hair. Halteres yellow.

FEMALE.—Length 2 mm. Head width 0.6 mm.

Width of frons 0.325 mm, lateral length 0.26 mm.

Third antennal joint piriform, length 0.23 mm, width

at base 0.117 mm, brown, darkened apically, covered

with very short whitish pubescence, without hairs.

Arista 0.42 mm, distinctly pubescent. Thorax darker

than in male, pleura reddish castaneous. Mesopleura

bare. Scutellars as in male. Tergites dull black, 2nd

elongated, 3-6 subequal, all with a distinct yellow

seam on hind margins. Foretibia 0.5 mm, with 2

bristles on basal half. Foretarsus one-fifth longer than

tibia; metatarsus slender, of normal structure, joints

2-4 slightly flattened, longer than broad, 2nd joint

about two-thirds of metatarsus in length. Coxate I-II

and midfemur without lanuginous vestiture. Wing

length 2.15 mm, width 0.88 mm. Costa 0.52 of wing

length, slightly thicker than 3rd vein; ratio of segments 21:16. Cilia 0.1 mm long, about 8 between 1st

and 3rd veins. Halteres yellow.

MATERIAL EXAMINED.—Holotype male (USNM

70237), Dominica, B.W.I., Trois Pitons, 4.V.64 (O. S.

Flint, Jr.); wing on slide. One paratype, female

(USNM), Dominica, W.I., Pont Casse, 1.7 m east,

12.111.65 (W. W. Wirth).

DISCUSSION.—Coniceromyia pilicoxa is a striking

species. The male is easily recognized by the lanuginous

vestiture of the fore- and midcoxae and midfemora.

In my key (1963) it runs to C. antricolor Schmitz

(Chile), from which it differs at once by the absence

of hairs on the 3rd antennal joint. The long costa and

the ratio of costal segments is distinctive for both

sexes; the only other species of the genus with the

costal index 0.5 are C. striativena Borgmeier (Mexico) and C. cubensis Brues ( $ ) .

The female is considerably smaller than the male,

but that occurs also in other species of the genus and

was first observed by Brues in C. cubensis.

Coniceromyia ciliatipes, new species

FIGURES 8-10

MALE.—Length 1.8 mm. Head width 0.65 mm.

Frons broader than long (4:3), width 0.36 mm, dark

castaneous, strongly shining, scarcely pubescent, with

a weak median line. Supraantennals short. First and

2nd row slightly convex, the bristles equidistant. One

lower postocular bristle; one bristle on jowls; cheeks

bare. Third antennal joint dark brown, retort-shaped,

length 0.45 mm. narrowed portion distinctly hairy;

arista one-half as long as 3rd antennal joint, distinctly

pubescent. Palpi pale yellow, narrow, length 0.16 mm,

with short bristles on apical 3rd.

Thorax shining, reddish yellow, darker and sometimes castaneous behind. Mesopleura bare. Scutellum

with 2 long posterior bristles; the anterior are hairlike and one-third to two-fifths of the posterior in

length.

Abdomen opaque, venter yellow; tergites black or

black brown, all with yellow apical seams; 2nd tergite

elongated, 3-6 subequal. Pubescence very scarce. Hypopygium large, globose, reddish brown, shining. Anal

segment not protruding.

Legs yellow, hind femur black at tip. Foretrochanter

at apex with a tuft of curved black hairs. Forefemur

(Figure 10) with long sensorial cilia on ventral border; similar but shorter and somewhat finer cilia are

present also on foretibia ventrally. Foretibia 0.49 mm,

with 2 dorsal bristles on 1st and 2nd third. Foretarsus

(Figure 8) 0.57 mm, broadened, metatarsus excavated

anteriorly forming a bare furrow, apically with a

slender fringed process, joints 2-5 somewhat longer

than broad, the width and length diminishing gradually. Midtibia with the usual bristles, the subapical

bristle as long as the anterodorsal one near base. Hind

femur maculated at tip, strongly broadened at basal

fourth, posterior face ventrobasally with a broad marginal band of extremely short sensorial pins. Hind tibia with one dorsal bristle at basal third and one short

anterodorsal subapically; 3 end-spurs as usual.

Wing (Figure 9) grayish yellow, veins dark brown,

costa darker than 1st and 3rd vein. Length 1.74 mm,

width 0.65 mm. Costa thickened, index 0.43, ratio of

segments 17:9. Cilia moderately long (0.1 mm). Third

vein with a bristle at base; 4th slightly concave, deflected apically; 5th and 6th but little curved, 5th with

a faint hook at base. Axillary margin with one hair.

Halteres pale yellow.

FEMALE.—Length 1.6 mm. Similar to the male.

Third antennal joint short oval, dark brown apically,

paler at base. Color of thorax and scutellars as in male.

Second tergite a little elongated, 3-6 subequal in

length. Cerci distinct, yellow. Forefemur and tibia not

ciliated on ventral border. Foretarsus longer than tibia

(22:19), metatarsus of normal structure. Hind femur

moderately broad, about 3 times as long as broad.
NUMBER 23

MATERIAL EXAMINED.—Holotype male (USNM

70238), Dominica, W.I., S. Chiltern Est, 20.11.65

(W. W. Wirth). Paratypes (USNM, CTB): 8 males

and 6 females (I.-III.65, W. W. Wirth), from the

following localities: S. Chiltern Est, Pont Gasse 1.5-

1.7 m east, Manets Gutter, Freshwater Lake, Clarke

Hall, Fond Figues River.

DISCUSSION.—In my key (1963b) C. ciliatipes runs

to C. latimana Malloch, but in the latter the thorax is

dark brown to black, the last tarsal joint of forelegs is

fuscous, the foremetatarsus ( $ ) is broader and the

forefemur and tibia lack the sensorial cilia on ventral

border; the anterior scutellars are also a little longer;

in the female the second costal segment seems to be

shorter in latimana. The females are difficult to distinguish, because sometimes the thorax of ciliatipes is

darkened.

Coniceromyia latimana (Malloch)

FIGURE 11

Conicera latimana Malloch, 1924, p. 73 [male; Puerto Rico].

Coniceromyia latimana (Malloch).—Schmitz, 1927, p. 66;

1929, p. 126.—Borgmeier, 1950, p. 298; 1963b, p. 455,

fig. 7.

Of this species Dr. Wirth collected on the Island of

Dominica at Clarke Hall, with a Malaise trap, 7 males

and 2 females, January-March 1965.

The male is easily recognized by the broad foremetatarsus (Figure 11) and the fuscous color of the

last tarsal joint of forelegs. The female differs from

C. ciliatipes by the dark color of the thorax and also by

the somewhat longer anterior scutellars which reach

one-half of the posterior, a character I failed to mention

in my redescription (1963b).

Genus Conicera Meigen

Conicera (Hypocerina) seticerca, new species

FIGURES 12-14

MALE.—Length 1.1 mm. Frons black, with two small

reclinate supraantennak very close together and 2 antials twice as far apart and inserted immediately above

the supraantennals; anterolateral bristles wanting.

Third antennal joint (Figure 14) black, conical,

length 0.14 mm, width 0.09 mm, shortly pubescent;

arista apical, twice as long as third antennal joint

(0.28 mm), relative length of arista! segments i-ra=

4:5:23; segments i-n and basal half of m thickened.

Palpi dark, slender, with about 6 short setulae, the apical one not differentiated.

Thorax black. Scutellum wkh 2 bristles and 2 minute hairs. Mesopleura bare.

Abdomen black. Hypopygium dark brown; pincers

of the epandrium very characteristic (Figures 12-13);

right forceps with the upper (proximal) process broad

and short, truncated at tip and with 4 short black

cone-shaped pins and several hairs; lower (distal)

process with numerous strong black teeth only partly

visible in dorsal view, and besides the hairs with about

7 long bristles near outer border; left forceps slightly

excavated apically, with numerous thick conelike

teeth, many hairs and 7 or 8 strong bristles on outer

border.

Legs dark brown; chaetotaxy as usual. Foretibia

0.27 mm, with a dorsal bristle above middle and a dorsal row of 8 short setulae on distal half; foretarsus

0.36 mm, ratio of tarsal joints 10:5:4:4:4, last joint

slightly thicker than preceding. Midtibia 0.29 mm,

dorsal bristle near base 0.15 mm. Hind femur 0.49 mm,

width 0.15 mm, ventrobasally with 10 fine hairs on

basal half. Hind tibia 0.39 mm, dorsal bristle on basal

third as long as ventral end-spur (0.15 mm).

Wing yellowish brown, veins dark brown. Length

1.09 mm, width 0.52 mm. Costa 0.44 of wing length,

ratio of segments 21:11. Cilia short, there are 10 pairs

from base to tip of first vein, and 3 single cilia on

2nd costal segment. Third vein thicker than costa,

with a bristle at base which is about as long as the

longer costal cilia. Mediastinalis obsolete. Fourth vein

concave, 5th slightly sinuous, 7th weak but complete.

Halteres dark.

MATERIAL EXAMINED.—Holotype male (USNM

70239, on slide) and one paratype (CTB; dissected,

on three slides), Dominica, W. I., Clarke Hall, light

trap, October 1964 (P. Spangler).

DISCUSSION.—This striking species seems to be

close to megalodus Schmitz (Brazil) but differs by the

shape and chaetotaxy of the hypopygial pincers (see

Schmitz, 1929, fig. 28 (6, 7, p. 118). Conicera {Hypocerina) neotropica Brues (1904; Grenada, W.I.), re-
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

cently redescribed by Borgmeier (1968), may also be

near, but has the 3rd antennal segment somewhat

longer and the arista relatively shorter; the structure

of the hypopygium is unknown.

Genus Gymnoptera Lioy

Gymnoptera Lioy, 1864, p. 79.—Schmitz, in Lindner, 1953,

p. 300.—Borgmeier, 1958, p. 307.

This genus is very similar to Conicera Meigen, but

can be easily recognized by the chaetotaxy of the hind

tibiae, which lack the dorsal bristle near base that are

always present in Conicera. The male has stilettoshaped appendages on the hind border of the hypopygium. The dimorphism in the venation of the wing is

remarkable; in the male the costa is shorter, and the

1st costal segment is longer than the 2nd; in the female

the costa is longer, and the 1st costal segment is shorter

than the 2nd. The larvae and pupae are very characteristic (see figures of Schmitz, in Lindner, 1953, p.

302).

Gymnoptera neotropica Borgmeier

Gymnoptera neotropica Borgmeier, 1958, p. 307, fig. la-d

[male; Rio de Janeiro].

FEMALE (undescribed).—Length 1.3-1.5 mm.

Third antennal joint dark brown, small, globose, pointed at tip, length 0.13 mm, width 0.11 mm; arista apical, long (0.58 mm in a large specimen), distinctly pubescent. Tergites brown, opaque, 2 and 6 a little elongated, 3-5 subequal in length; pubescence scarce,

short, nearly absent dorsally. Legs as in male. Wing

hyaline, heavy veins pale brown. Length 1.53 mm (in

female from Dominica 1.3 mm), width 0.72 mm

(0.62 mm). Costa 0.54 (0.53) of wing length, ratio

of costal segments 13:16 (11:14). Costal cilia short

and numerous, arranged in pairs until near extreme tip,

2nd segment with 10-11 pairs and one single cilia.

MATERIAL EXAMINED.—The type-series (holotype

and paratypes, males in CTB) from Rio de Janeiro;

many puparia and several males and females reared

from dead Calosoma beetle, 111.-IV. 1960 (T. Borgmeier). One female (on slide, USNM) from Dominica, W. I., Manets Gutter, 7 March 1965 (W. W.

Wirth), light trap.

Subfamily METOPININAE

Tribe METOPININI

Genus Megaselia Rondani

Key to the Species of Megaselia

1. Mesopleura hairy or bristled (Subgenus Aphiochaeta) 2

Mcsopleura bare (Subgenus Megaselia, sensu stricto) 14

2. Scutellum with 4 bristles (group I) 3

Scutellum with 2 bristles 5

3. Mesopleura with uniform hairs; 5th tergite of female and 5th-6th in male with an apical row

of bristles 1. M. (A.) setigera (Brues)

Mesopleura hairy and bristled 4

4. Tergites yellow-banded, in female of normal size 2. M. (A.) fasciiventris (Enderlein)

Tergites 1-3 black, 2-3 large (Figure 18), segments 4-10 (9 ) yellow or orange

3. M. {A.) lutticauda (Borgmeier)

5. Mesopleura with hairs and one or more bristles 6

Mesopleura with uniform hairs 12

6. Costa long (0.44 or more of wing) (group II) 7

Costa short; fork wanting (group III) 11

7. Mesopleura with one long bristle 8

Mesopleura with one or more moderate bristles 9

8. Costa 0.58-0.60 of wing length, 2 longer than 1 (Figure 23) . 4. M. (A.) pleurochaeta, new species

Costa 0.47 of wing length; 2 shorter than 1 (Figure 21) . . . .5. M. (A.) spiculata, new species

9. Two supraantennals; frons glossy, coarsely punctured . . . 6. M.(A.)punctifrons, new species

Four supraantennals 10
NUMBER 2 3

10. Thorax yellow; costal cilia very short and close 7. M.(A.) semiferruginea Bridarolli

Thorax dark; cilia long, widely placed (Figure 26) 8. M.(A.) parviseta, new species

11. Costa 0.34 of wing length (Figure 27) 9. M.(A.) destituta, new species

Costa 0.42 of wing length 10. M.(A.) species indeterminata

12. Costa long (group IV) 13

Costa short; 1 longer than 2 + 3 (Figure 31); fore-metatarsus thickened (Figure 33) (group V)

13. M.(A.) definita, new species

13. Costa 0.55 of wing length; 2 longer than 1; palpi of normal shape

11. M.(A.) gemella Borgmeier

Costa 0.48 of wing length; 1 longer than 2+ 3 (Figure 30); palpi of male swollen (Figures 28, 29)

12. M.(A.) pullidava, new species

14. Scutellum with 4 bristles (group VI) 15

Scutellum with 2 bristles 22

15. Fork missing; costa 0.41 of wing length (Figure 34) ... . 14. Af. (A/.) deprivata, new species

Fork present , 16

16. Hind tibial setulae biseriate 17

Hind tibial setulae uniseriate 18

17. Halteres yellow; costa 0.5 of wing length; 1 shorter than 2 + 3 . . . 15. M.(M.)pkta (Lehmann)

Halteres dark brown; costa 0.59 of wing length; 1 subequal to 2 . 16. M.(Af.) subpicta (Malloch)

18. Scutellars unequal 17. M.(M.) obscuripalpis, new species

18. &M.(M.) scalaris (Loew) in part

Scutellars equal or subequal 19

19. Supraantennals equal 18. 9 M.(M.) scalaris (Loew)

Supraantennals unequal 20

20. Costal segment: 2 longer than 1 (Figure 35) .. . 19. M.(M.) elegantula, new species

Costal segment: 2 shorter than 1 21

21. Costal cilia very short and close; female abdomen (fig. 39) orange or yellow; anal segment

long (Figure 38) 20. Af.(Af.) aurea (Aldrich)

Costal cilia long; tergites brown with yellow apical seams; anal segment short

21. M.(Af.) striativentrisBorgmeier

22. Costa long (0.44 or more of wing length) (group VII) 23

Costa short (group VIII) 54

23. Halteres yellow 24

Halteres dark 30

24. Two supraantennals; 6th tergite of female long and narrow; hypopygium with 4-5 bristles

laterally 22. M. (M.) seticauda (Malloch)

Four supraantennals 25

25. Supraantennals equal 20

Supraantennals unequal 28

26. Hind tibial setulae biseriate 15. Af. (M.)picta (Lehmann) p.p.

Hind tibial setulae uniseriate 27

27. Pleura yellow 18. d" Af. (A/.) scalaris (Loew) in part

Pleura brown 23. M. (M.) pleurofascia Borgmeier

28. Abdominal segments 3—5 (9) membranous (Figure 41); fourth sternite with 2 curved rows

of setulae (Figure 42) 24. Af. (Af.) hypochaeta, new species

Abdomen of female with 6 tergites 29

29. Sixth tergite (9) long, rounded behind (Figure 44). . . . 25. Af. (Af.) Uptacina, new species

Sixth tergite short, rounded anteriorly (Figure 48) . . . . 26. Af. (Af.) imitatrix, new species

30. Hind tibial setulae biseriate; upper postantennals two-thirds of frontal width apart; labellae

broad (Figure 50) 27. Af. (Af.) buccata, new species

Hind tibial setulae uniseriate 31

31. Frons glossy 32

Frons opaque or subshining . . . . 33

32. Fore-metatarsus (o* 9 ) thickened (Figure 52); coxae black . . 28. M.(M.)polita (Enderlein)

Fore-metatarsus slender; coxae yellow; hind femur brown, basal 3rd yellow

29. M.(M.)jemoralis (Endcrlein)

33. Costa shorter than one-half of wing (0.45-0.48) 34

Costa 0.5 or longer 40

34. Costal cilia long (Figure 56); black species 30. Af.(Af.) enderleini (Brues)

Costal cilia short 35

336-709 O—69 2
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

35. Labdlac (o") large, shield-shaped; costa 0.48 (Figure 58) . 31. M.(M.) labdlifera, new species

Labdlae not shield-shaped 36

36. Foretarsus (d

1

) thickened 37

Foretarsus not thickened 38

37. Fore-metatarsus strongly thickened (Figure 67) 32. M.(M.) metatarsalis new species

Fore-metatarsus slightly thickened (Figure 62) 33. M.(M.) tarsodes, new species

38. Third tergite of female constricted (Figure 69) 34. M.(M.) substricta, new species

Third tergite of female not constricted; costa 0.45 39

39. Hypopygium with 2 bristles laterally (fig. 71); antials vertically below laterals

35. M.(M.) diplochaeta, new species

Hypopygium without bristles; antials midway between supraantennals and laterals

36. Af.(Af.) horticola Borgmeier

40. Two supraantennals 41

Four supraantennals 43

41. Palpi of male slender, fusiform, with short setulae (Figure 77)

37. M.(M.) gracUipalpis, new.species

Palpi normal 42

42. Wing whitish, veins pale (Figure 79); anal tube long . . 38. M.(M.)pallidipennis, new species

Wing yellow, veins brown (Figure 81); anal tube short .. . 39. M.(M.)ferruginosa (Brues)

43. Male with a sternal pocket on 6th segment 44

Male without sternal pocket 45

44. Sternal pocket large (Figure 83); supraantennals subequal . 40. M.(M.) bursella, new species

Sternal pocket small (Figure 85); supraantennals unequal

41. M.(M.) burselloides, new species

45. Proboscis of female broad 46

Proboscis of female normal 47

46. Fifth segment ( 9 ) laterally with stiff hairs (Figure 87); sides of 6th tergite straight

42. M. (M.)fottirostris, new species

Fifth segment ( 9 ) without stiff hairs; sides of 6th tergite convex (Figure 91)

43. M. (M.) tumidirostris, new species

47. Veins rather straight; fork long 48

Veins normal 49

48. Tergites 3-5 ( 9 ) each laterally with a large stigma ... . 44. M. (M.) miristigma Borgmeier

Stigmata of normal size; terminal segments ( 9 ) with a chitinized hook-shaped structure . . .

45. M. (M.) hamaticauda, new species

49. Last joint of midtarsus (cf) twice as long as preceding (Figure 96)

46. M. (M.)paula, new species

Last joint of midtarsus of normal length 50

50. Fork large; 4th vein starting behind fork; antials vertically below laterals

47. M. (M.)flampleura (Malloch)

Fork of medium size; antials inside of laterals 51

51. Arista short 52

Arista long 53

52. Hind femur black at tip; 5th tergite (c?) yellow; hairs of ventral fringe of hind femur short

and close (Figure 100) 48. M. (M.) crinellifemur, new species

Hind femur not maculated; 5th tergite (o") brown; hairs of hind femur longer, widely placed

(Figure 106) 49. M. (M.) subaristalis, new species

53. Fifth tergite (9 ) excavated behind, 6th transverse (Figure 110)

50. M. (M.) totaflava, new species

Fifth tergite (9 ) straight behind; 6th oval, longer than broad (Figure 112)

51. M. (M.) alajuelensis (Malloch)

54. Fork wanting; costa 0.42 (Figure 115) 52. M. (M.) mutilata, new species

Fork present 55

55. Halteres yellow; 3rd vein with a bristle at base 53. M. (A/.) brevicostalis (Wood)

Halteres dark; 3rd vein without a bristle at base (Figure 117)

54. M. (M.) crinipyga, new species
NUMBER 2 3

1. Megaselia (Aphiochaeta) setigera (Braes)

Aphiochaeta aurea var. setigera Brues, 1919a, p. 505 [9 ;

Grenada].

Megaselia (Aphiochaeta) setigera (Brues).—Borgmeier,

1962b, pp. 301 [key], 317, figs. 1-3, 14 [Grenada; Costa

Rica; Panama; Dominica].

Holotype female (MCZ), paratypes females (MCZ,

CTB), Grenada, Brues collector.

MALE (undescribed).—Length 1.30-2.00 mm. Frons

broader than long laterally (4:3), yellow to brown,

subshining, pubescent. Lower supraantenals weak,

one-half to two-thirds of upper, the latter about onefourth of frontal width apart. Antials a little below

level of upper postantennals, slightly farther from them

than from laterals which are a little above level of

upper supraantennals. One lower postocular bristle;

one bristle on jowls; cheeks with a few hairs. Third antennal joint brown, of normal size. Palpi yellow, well

bristled.

Thorax yellow. Mesopleura with a few short hairs.

Scutellum with 4 equal bristles.

Abdomen opaque, venter yellow; tergites black, second yellow at base and with bristles laterally; 5th and

6th with bristles on hind margin, the lateral ones longer.

Hypopygium brown, not large, on both sides with

bristly hairs. Anal segment long, yellow, end-hairs differentiated.

Legs yellow, hind femur black at tip. Foretarsus very

slender. Hair-seam of midtibia nearly complete. Hind

femur on basal hah* of ventral edge with 6-7 setuliform

hairs. Posterodorsal cilia of hind tibia short, the basal

ones weak.

Wing grayish yellow, veins brown. Costa 0.56 of

wing length, exceeding a little tip of 3rd vein, ratio of

segments 14:12:4. Cilia very short and close. Fourth

vein starting at fork, 5th and 6th sinuous. Axillary

margin with 5-6 hairs. Halteres brown.

FEMALE.—Length 1.5-2.8 mm. Wing (Borgmeier,

1962b fig. 14); costa exceeding slightly tip of 3rd vein

(Figure 14 is not correct in this respect). Tergites

as in his Figure 1 (Borgmeier, 1962b); 3rd as long as

2nd, with the hind margin straight; 4th rudimental,

variable in size, semicircular; 5th transverse, with 8

long bristles on hind margin.

MATERIAL EXAMINED.—Besides the specimens listed

in a previous paper (1962) I have seen from Dominica

18 dried specimens (6 $ , 12 9 ) and 79 preserved in

alcohol (34 & 45 9 ) from: Clarke Hall, S. Chiltern

Est., Manets Gutter, Fond Figues River (Wirth 1965,

I-III; Steyskal, May 1966).

DISCUSSION.—A very striking species of group I,

easily recognized by the long bristles on 5th tergite of

female, and on hind margin of 5th and 6th tergite in

male.

2. Megaselia (Aphiochaeta) fasciiventris

(Enderlein)

FIGURES 15-17

Aphiochaeta punctifemur var. fasciiventris Enderlein, 1912,

p. 33 [ 9 ; Hammonia=Corupa, S. Catarina, Brazil].

Megaselia (Aphiochaeta) fasciiventris (Enderlein).—Borgmeier, 1935a, p. 462, pi. 5: fig. 5 [wing; type]; 1962b, p.

302 [key].

Aphiochaeta caribbaea Brues, 1919a, p. 504 [9 ; Grenada,

W.I.].

Megaselia (Aphiochaeta) caribbaea (Brues).—Borgmeier,

1962b, p. 313, fig. 15 [type]; 1967c, p. 404 [synonymy].

MALE (undescribed).—Length 1.6-2.0 mm. Frons

broader than long (4:3), black, subshining, sparsely

pubescent, with a distinct median line. Supraantennal

bristles (su) subequal, the upper one-third or more

of frontal width apart, the lower one-eighth. Antials

on level of lower su, about midway between upper su

and eye; anterolaterals slightly above level of upper

su near eye-margin. Second row slightly convex, the

bristles equidistant. One long lower postocular; jowls

with 2 bristles; cheeks with 5-6 weak bristles. Third

antennal joint globose, dark brown; arista long, distinctly pubescent. Palpi brownish yellow, flat, with 5

strong bristles.

Thorax yellow to dark brown, subshining. Pubescence short. Mesopleura hairy, with 1-3 moderate

bristles or prolonged hairs. Scutellum with 4 long bristles, the anterior sometimes a trifle weaker.

Abdomen opaque; venter yellow; tergites subequal,

black with yellow apical bands. Pubescence distinct,

bristle-like laterally, longer on hind margin of sixth.

Hypopygium (Figures 16, 17) small, black, with several hairs on both sides; ventral plate broadly rounded.

Anal segment yellow, of median length, apical hairs

not differentiated.

Legs brownish yellow. Foretibia with a dorsal row of

black spinuli. Foretarsus slender, metatarsus a little

longer than 2 + 3. Hair-seam of midtibia reaching

apical fifth, posterodorsal setulae distinct, some anterodorsal setulae on basal half. Hind femur broad

(ratio 25:65), on basal half of ventral edge with about

11 stiff short setulae.

Wing (Figure 15, $ from Nova Teutonia) brownish, veins brown. Length 1.76 mm, width 0.75 mm.
10 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Costa 0.53 of wing length (0.56 in male from Dominica), ratio of segments 14:12:4 (Dominica: 14:

13:4). Cilia moderately long (0.1 mm), about 9 pairs

between 1st vein and tip of costa. Mediastinalis present.

One distinct bristle at base of 3rd vein. Fork acute;

4th vein slightly concave, starting at fork with a weak

S-curvature; 5th bent before middle; 6th sinuous.

Axillary margin with 4 hairs. Halteres yellow or brown.

FEMALE.—Length 2.0-2.3 mm. Similar to male.

Costa 0.55 of wing length in lectotype, 0.56 in type of

A. caribbaea; ratio of segments 16:15:5. Tergites somewhat shining.

MATERIAL EXAMINED.—Besides the type from

Corupa, S. Catarina, Brazil, I have seen 30 males and

14 females from Nova Teutonia, and 7 males and

3 females from Petropolis, R.J., Brazil. From Dominica: 2 males in alcohol, Clarke Hall, March 1965

(Wirth).

DISCUSSION.—M. (A.) fasciiventris belongs to

group I. There are 4 long scutellars of equal length in

both sexes; the species is wrongly placed in my key

(1962b) among those with unequal scutellars.

VARIATION.—The color of the thorax and halteres

is variable. In the 2 males from Dominica the costa is

rather long (0.56 of wing length), the 4th vein is more

curved, the 5th is less arcuate; the halteres are dark;

the hypopygium is identical to males from Nova

Teutonia.

3. Megaselia (Aphiochaeta) luteicauda (Borgmeier)

FIGURES 18-20

Aphiochaeta luteicauda Borgmeier, 1925b, p. 145 [9 ;

Petr6polis].

Megaselia (Aphiochaeta) luteicauda (Borgmeier).—Borgmeier, 1962b, p. 301 [in key].

Lectotype female (CTB) and one paratype female

(CTB), from Petropolis, R. J.; Borgmeier and Ronchi,

collectors.

MALE (undescribed).—Length 1.6-2.0 mm. Frons

about as broad as long medially, black to blackish

brown, subshining, scarcely pubescent, with a fine

median line. Su subequal, the lower somewhat shorter

(5:4 or 6:5), the upper about one-third of frontal

width apart, the lower one-sixth. Antials on level of

lower su, near anterior corner of frons, a little removed

from eye-margin; anterolaterals slightly above level of

upper su, near eye-margin. Second row slightly convex,

the bristles equidistant. One strong lower postocular

bristle; jowls-cheeks with an ascending row of 8 moderate bristles. Third antennal joint globose, brown;

arista long, distinctly pubescent. Palpi flattened, yellowish, with 5 long bristles. Proboscis short, diskshaped, circular when seen from below.

Thorax dark castaneous, subshining, Pubescence

short. Mesopleura with a few fine hairs above and 1-3

bristly hairs behind. Scutellum with 4 long bristles.

Abdomen opaque, black, grayish pollinose; 2nd tergite elongated, 3-6 subequal, sparsely pubescent, some

bristly hairs laterally and on hind margin of 6th. Hypopygium (Figure 20) small, black brown, at left with

a row of 5 bristly hairs near lower margin. Anal segment long (0.2 mm), yellow, with short hairs, apical

hairs not differentiated.

Lews yellowish brown to castaneous. Foretibia with

a dorsal row of black spinuli. Foretarsus slender, metatarsus about as long as 2-4. Midtibial setulae biseriate

on basal half, hair-seam reaching apical sixth. Hind

femur broad, ventral hairs on basal half short, delicate,

Posterodorsal setulae on hind tibiae strong below middle, weaker above.

Wing (Figure 19, <$ from Dominica) yellowish

brown, veins brown. Costa 0.56 of wing length, ratio

of segments 14: 14:3. Cilia long (0.13 mm), there are

6 pairs between ra and r3. Mediastinalis obsolete. Third

vein with a bristle at base. Fork not large, acute; 4th

vein starting at fork with a very faint hook (sometimes

missing), curved near base, nearly straight at middle;

veins 5-6 sinuous. Axillary margin with 3 hairs.

Halteres yellow, peduncle reddish brown.

FEMALE.—Length 2.0-3.5 mm. Upper su sometimes

more than one-third of frontal width apart, antials

sometimes nearly vertically below anterolaterals. Third

antennal joint small. Proboscis short, narrow. Abdomen

(Figure 18) opaque, black on segments 1-3, venter

enclosed, the rest yellow or orange, 1st tergite short;

2nd and 3rd large trapezoid, form a little variable, with

short sparse pubescence, marginal hairs on 2nd (sometimes also on 3rd) stiff, bristle-like. Segments 3-10

membranous, retractile, often nearly entirely withdrawn under 3rd tergite; 3-6 indistinctly segmented,

practically bare, 5th very short, 7-10 abbreviated. Cerci

indistinct. Costa 0.55-0.57 of wing length, segments

1 and 2 subequal, 1st sometimes a little longer (23:20),

there are 7-8 pairs of cilia between ra and r3.

MATERIAL EXAMINED.—Besides the types from

Petropolis, Rio de Janeiro, I have seen both sexes from

Nova Teutonia, Brazil (Plaumann) and Costa Rica

(Higuito, San Mateo; Suiza de Turrialba; P. Schild

leg.) ; numerous females from Curitiba, (Wiltuschnig,
NUMBER 2 3 11

XII. 1925) ; one female from Loreto, Misiones, Argentina (Ogloblin 11.X36), on dead opilionid. From

Dominica I have seen 12 males and 8 females from the

following localities (Wirth I.—Ill, 1965) : d'Leau

Gommier, Manets Gutter, Sylvania, Layou R. mouth,

Pont Casse, Fond Figures R. and Clarke Hall.

DISCUSSION.—This striking species belongs to group

I. The female is easily recognized by the color and

structure of the abdomen, the male by the long anal

tube. The species comes close to M. pyteryacantha

Borgmeier which has the scutellars unequal and the

female abdomen different.

4. Megaselia (Aphiochaeta) pleurochaeta,

new species

FIGURE 23

MALE.—Length 1.2-1.7 mm. Frons subquadrate,

slightly broader than long (9:8), yellow, subshining,

scarcely hairy, medium furrow distinct. Four supraantennals, the lower very weak and hardly one-half of

the upper which are about one-third of frontal width

apart. Antials a little below level of upper supraantennals, a little farther from them than from eyemargin and from anterolaterals; the latter are slightly

higher placed. Preocellars as far from each other as

upper supraantennals. One lower postocular bristle;

jowls and cheeks with a series of fine hairs. Third

antennal joint brownish yellow, of normal size; arista

one-third longer than width of frons, distinctly

pubescent. Palpi yellow, not large, with 6 short bristles.

Thorax yellow, subshining; pleura pale. Scutellum

with 2 bristles and 2 small hairs. Mesopleura with a

few hairs above and a strong bristle behind which

is about as long as the scutellars.

Abdomen: venter pale yellow; tergites dark brown,

a little paler brown at middle and on anterior half of

2nd tergite; the latter with 2 bristles laterally. Pubescence scarce and short, longer on hind margin of

6th tergite. Venter on hind margins of segments 3-6

with 2-4 bristle-like hairs. Hypopygium dark brown,

of medium size, dull, with hairs below and on each

side with 2 conspicuous bristles behind; dorsum bare.

Anal tube long, yellow, with short hairs, end-hairs

strong.

Legs yellow, hind femur blackish at tip. Foretarsus

slender. Hind femur moderately broad, with a distinct fringe of longish hairs on ventral edge of basal

half. Hind tibia with about 10 distinct posterodorsal

setulae.

Wing grayish yellow, veins brown. Length 1.2-1.5

mm. Costa in male 0.58 of wing length, ratio of segments 20:24:5. Cilia short. Fourth vein concave, very

faintly recurved at base; 5th and 6th slightly sinuous.

Anal margin broadly convex. Axillary margin with 2

short hairs. Halteres dark brown.

FEMALE.—Similar to male. Tergites brown; 2nd

elongated, reaching sides, 3-6 gradually narrowed, 6th

with a median notch anteriorly, 2nd with 2 bristles

laterally; pubescence very scarce. Terminal segments

fleshy. Venter with hairs as in male. Wing (Figure

23) : costa up to 0.6 of wing length, ratio of segments 22:27:4.

MATERIAL EXAMINED.—Holotype male (USNM

70240), Dominica, Pont Casse, 1.7 miles east, 12.111.65

(W. W. Wirth). Paratypes (USNM, CTB) : 5 males

and 2 females, partly in alcohol, same data as type; 1

female, Pont Casse, 11.IV.1966 (R. J. Gagne).

DISCUSSION.—M. pleurochaeta seems to be close

to M. macrohaeta Malloch (1912; Puerto Rico), but

in the latter the female has 4 scutellars, the halteres are

yellow, and the first costal division is longer than the

second.

5. Megaselia (Aphiochaeta) spiculata, new species

FIGURES 21, 22

MALE.—Length 1.56 mm. Head width 0.52 mm.

Frons black, slightly broader than long laterally (6:5),

width 0.36 mm, median length 0.30 mm, distinctly

pubescent, with a deep median furrow. Supraantennals subequal, the lower about three-fourths of upper;

the upper one-fifth of frontal width apart, the lower

close together. Antials farther from su than from

anterolaterals, the latter in line with upper su, the antials slightly lower. Second row nearly straight, the preocellars farther from mediolaterals than from each

other. Upper and lower postoculars strong. Cheeks

with some setulae. Third antennal segment black, globose, of normal size; arista 0.39 mm, distinctly pubescent. Palpi yellow, of normal shape, with 5 strong

bristles on apical 3rd and some setulae.

Thorax black-brown. Pubescence short, bristle-like

behind. Two dorsocentrals. Mesopleura with about 15

short hairs and one long bristle (0.176 mm), about as

long as scutellars. Scutellum with 2 bristles and 2

minute hairs.

Abdomen black brown, venter dark; tergites subequal, first tergite yellow, 2-6 with a broad median

yellow spot anteriorly, forming nearly a yellow longi-
12

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

tudinal strip. Pubescence conspicuous, longer on hind

margin of 6th. Hypopygium (Figure 22) small, dark

brown, with numerous bristles or bristly hairs. Anal

segment yellow, of medium length (0.13 mm), hairy,

end-hairs conspicuous but not much differentiated.

Legs yellow, hind femur with a large apical macula.

Foretibia 0.33 mm; tarsus slender, 0.46 mm, metatarsus longer than 2 + 3. Midtibia 0.44 mm, with distinct

anterodorsal and posterodorsal setulae; seam nearly

complete; end-spur four-fifths of metatarsus. Hind

femur broad (ratio 10:25), ventrobasal hairs short,

inconspicuous. Hind tibia 0.59 mm, with 8 strong

moderately short setulae.

Wing (Figure 21) distinctly grayish yellow, veins

pale brown, 4-7 evanescent at tip. Length 1.37 mm,

width 0.59 mm. Costa 0.47 of wing length, ratio of

segments 10:8:4; cilia moderately long (0.1 mm),

about 6 pairs between 1st vein and apex of costa. Mediastinalis obsolete. Fork acute. Third vein with a distinct bristle at base. Fourth starting at fork with a

faint hook, curved at basal fourth, remainder nearly

straight; 5th feebly recurved at base, broadly bent

before middle; 6th sinuous. Axillary margin with 3

hairs. Halteres black.

MATERIAL EXAMINED.—Holotype, on slide (USNM

70241), Dominica, Manets Gutter, 7 Mar. 1965 (W.

W. Wirth).

DISCUSSION.—M. (A.) spiculata, new species, is

very close to M. (A.) parviseta, new species. For differences, see page 13.

6. Megaselia (Aphiochaeta) punctifrons,

new species

FIGURE 24

FEMALE.—Length with the terminal segments partly

protruding 1.75 mm. Head width 0.48 mm. From

convex, subquadrate, width 0.26 mm, deep black, distinctly shining, with scattered coarse piligerous punctures and a fine median furrow. Two thin supraantennals close together (about one-seventh of frontal width

apart and more approximate than the preocellars).

Antials reclinate, placed on anterior border, closer to

supraantennals than to anterolaterals which are distinctly higher. Second row slightly concave. Eyes large.

One strong bristle on lower corner of eye; cheeks with

4 hairs. Third antennal joint orange, somewhat oval,

of normal size. Arista brown, one-fifth longer than

width of frons, distinctly pubescent. Palpi yellow,

partly withdrawn into the mouth opening, with about

5 bristles (the apical is the longest). Proboscis short.

Thorax black or brownish black, distinctly shining,

pleura glossy. Mesopleura with hairs and one moderately long bristle behind (length 0.169 mm). Scutellum with 2 bristles (length 0.195 mm) and 2 minute

hairs.

Abdomen black, venter enclosed. Tergites reaching

sides, 2-5 subequal, glossy laterally, somewhat pruinose

dorsally, 6th entirely glossy. Pubescence scarce, short.

End-segments fleshy, 7th with a narrow tergal plate

and a circle of delicate hairs. Cerci small.

Legs black brown, forelegs paler. Foretarsus slender.

Foretibia with a dorsal row of short setulae. Posterodorsal setulae on midtibia distinct. Hind femur broad,

no ventral fringe on basal half. Hind tibia with about

7 distinct setulae, the upper short, those on lower half

strong and widely placed.

Wing (Figure 24) grayish brown, veins dark brown.

Length 1.55 mm, width 0.62 mm. Costa 0.53 of wing

length, ratio of segments 15:10:4. Cilia short, closely

placed. First vein nearly straight; 4th gently curved

on basal 3rd, base obsolete, apex faintly deflected; 6th

slightly sinuous. Axilliary margin with 2 hairs. Halteres

black.

MATERIAL EXAMINED.—Holotype female (USNM

70242), Dominica, S. Chiltern Est., 2 Feb. 1965 (W.

W. Wirth).

DISCUSSION.—M. (A.) punctifrons, new species,

belongs to group II. It is a nice strongly shining species

and is easily recognized by the two supraantennals and

the coarse frontal punctures.

7. Megaselia (Aphiochaeta) semiferruginea

Bridarolli

Megaselia (Aphiochaeta) semiferruginea Bridarolli, 1940,

p. 494 ($9; Argentina).—Borgmeier, 1962b, p. 303

[key].

Holotype male and paratype female (CTB), from

Carlos Paz, Sierras de Cordoba, Argentina; Bridarolli

collector.

MATERIAL EXAMINED.—Three females from Dominica, in alcohol, Clarke Hall, March 1965 (W. W.

Wirth).

This species belongs to group II (costa long, mesopleura hairy and with some setulae or prolonged hairs).

It may be confused with M. gemella Borgmeier, but the

latter lacks the mesopleural setulae, and the 2nd
NUMBER 2 3 13

costal segment is longer than the 1st; in M. semiferruginea the first and second costal segments are of equal

length and the foretarsus is rather stout in male.

8. Megaselia (Aphiochaeta) parviseta, new species

FIGURES 25-26

MALE.—Length 1.40 mm. Head width 0.52 mm.

Frons black-brown, broader than long laterally (4:3),

width 0.26 mm, sparsely pubescent, with a distinct

median line. Su equal or nearly so, the upper about

one-third of frontal width apart, the lower one-fifth.

Antials close to anterolaterals, on a slightly lower level

than upper su, anterolaterals almost in line with

upper su. One upper and one lower postocular bristle;

one bristle on jowls; cheeks with some weaker bristles.

Third antennal joint black, small, globose, width 0.09

mm; arista 0.39 mm. Palpi yellow, with 5 long bristles.

Thorax black-brown. Pubescence short, bristle-like

behind. Mesopleura with 4 short hairs above and 2

small bristles (length 0.09 mm). Scutellum with 2

bristles and 2 hairs.

Abdomen black. Tergites subequal. Pubescence

scarce, longer hairs on hind comers of tergites and on

hind margin of 6th. Hypopygium (Figure 26) blackbrown, with bristly hairs. Anal segment short, 0.09 mm,

yellow, end-hairs not differentiated.

Legs yellow, hind femur not darkened at tip. Foretibia 0.33 mm; tarsus slender, 0.49 mm, metatarsus as

long as 2 + 3. Midtibia 0.41 mm, cilia biseriate, seam

reaching apical fourth. Hind femur about 3-times as

long as broad, on basal half of ventral edge with 8

moderately long curved hairs. Hind tibia 0.54 mm,

with 10 distinct short setulae.

Wing (Figure 25) grayish yellow, veins pale brown,

4-7 evanescent at tip. Length 1.37 mm, width 0.57 mm.

Costa 0.52 of wing length, ratio of segments 24:20:7.

Cilia moderately long (0.11 mm), 6 pairs between 1st

vein and apex of costa. Mediastinalis obsolete. Fork

acute. Third vein with a fine bristle at base; 4th

starting at fork with a faint hook, feebly curved near

base, remainder nearly straight; 5th a little recurved at

base, remainder feebly sinuous; 6th slightly sinuate

Axillary margin with 3 hairs. Halteres yellow.

MATERIAL EXAMINED.—Holotype on two slides

(USNM 70243), Dominica, Pont Casse, 12 Feb. 1965

(Wirth).

DISCUSSION.—M. (A.) parviseta, new species, is

extremely close to M. (A.) spiculata, new species, but

in the latter the upper su are more approximated, the

mesopleural bristle is long, the hind femur is black at

tip, the anal segment is longer, and the costa is shorter;

the color of the halteres is also different.

REMARK.—The type-specimen was badly damaged

by accident during laboratory work, after the description and drawings had been made.

9. Megaselia (Aphiochaeta) destituta, new species

FIGURE 27

FEMALE.—Length 1.00 mm. Head width 0.31 mm.

Frons as broad as long medially (0.18 mm), lateral

length 0.11 mm, black or black-brown, pubsecent, with

a distinct median line. Four supraantennals of equal

length, the upper about one-fourth of frontal width

apart, the lower one-eighth. Bristles of 1st row forming

an evenly convex line with upper su, antials a little

farther from su than from anterolaterals, as far from

su as the latter from each other. Second row slightly

convex, the bristles equidistant. Upper postocular

distinct; lower strong. Jowls + cheeks with 3 bristles.

Third antennal joint black, of normal size, globose,

width 0.065 mm; arista 0.27 mm, shortly pubescent.

Palpi yellow, slender, with 4 long bristles on apical

half.

Thorax black or black-brown, Mesopleura with a

few short hairs and one very short bristle (0.03 mm).

Scutellum with 2 bristles and 2 minute hairs.

Abdomen black-brown, venter grayish yellow. Tergites 2-6 subequal, gradually narrowed, 6th transverse,

with a gland opening basally. Pubescence short, scarce;

venter on hind margins of 5th and 6th segments with

a transverse row of hairs placed in dark spots. Terminal

segments short. Cerci small.

Legs yellow, hind legs brownish yellow. Foretibia

0.20 mm. Fore-metatarsus slightly thickened, one-half

of tibia in length, about as long as joints 2 + 3. Midtibia

0.24 mm, hair-seam ending at middle; spur two-thirds

of metatarsus. Hind femur slender, length 0.39 mm,

width 0.10 mm, ventrobasally with 4 short widely

placed fine hairs. Hind tibia 0.35 mm, with 12 very

delicate short posterodorsal cilia.

Wing (Figure 27) slightly grayish, veins pale brown,

4-7 paler, evanescent at tip. Length 0.93 mm, width
14

0.42 mm. Costa 0.34 of wing length, slightly thickened,

especially on distal half, ratio of segments 3:2, with 8

marginal cilia, the longest 0.065 mm; there are 3 cilia

between tip of 1st vein and apex of costa; besides the

marginal bristles there is a dorsal row of 13 closely

set short hairs. Mediastinalis incomplete. Fork wanting.

Third vein without a bristle at base, apex upcurved and

thickened; 4th vein evenly arcuate, obsolete at base;

5-6 feebly sinuous, 7th indistinct. Axillary margin with

2 hairs. Halteres black.

MATERIAL EXAMINED.—Holotype female (USNM

70244) on slide, Dominica, Manets Gutter, 7 March

1965, light trap (Wirth).

DISCUSSION.—M. (A.) destituta, new species, is extremely close to M. tumidula Borgmeier (1962b,

Brazil) but the latter is larger, the costa is somewhat

longer, the 3rd vein is not distinctly upcurved at apex

and less thickened, and there are 4 cilia between r1-]"3

(see Borgmeier, 1962b, fig. 33); 6th more distinctly

sinuate, the upper su are also farther apart from each

other.

10. Megaselia (Aphiochaeta) species indetenninata

There is one male before me, labeled: "Dominica,

Manets Gutter, 5 March 1965, W. W. Wirth" and

"Megaselia (Aph.) lanceata Borg. det. W. W. Wirth."

This specimen has both wings badly damaged at base;

I put them on slide. The costa is less thickened than

in M. (A.) lanceata Borgmeier (Mexico); wing

length 0.91 mm, costa 0.41 of wing length. This male

probably belongs to a new species which is close to

M. (A.) tumidula Borgmeier (1962b, p. 338, figs. 33-

34; Nova Teutonia, Brazil). According to Dr. Wirth,

there is a 2nd male in the National Museum of Natural

History from the same locality (collected 7 March

1965, W. W. Wirth) which may be in better condition.

MATERIAL EXAMINED.—One female (USNM, on

slide), from Dominica, Pont Casse, April 1964, O. S.

Flint, at light, probably belongs to this species. Wing

length 1.24 mm, costa 0.42 of wing length.

I refrain from naming and describing this species

until additional material is available, preferably

preserved in alcohol.

In M.(A.) destituta, new species, described above,

which is also near, the costa is shorter (0.34 of wing

length).

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

11. Megaselia (Aphiochaeta) gemella Borgmeier

Megaselia (Aphiochaeta) gemella Borgmeier, 1962b,

pp. 304 [key], 349, fig. 43 [ <J 9].

Holotype male (CTB), paratypes one male 3 females (CTB, USNM), from Petropolis, R.J., Brazil,

and Nova Teutonia, S. Catarina, Brazil.

MATERIAL EXAMINED.—From Dominica, 8 males

and 24 females (pinned, one on slide) and 3 males and

30 females (in alcohol), from the following localities:

Clarke Hall, S. Chiltern Est, d'Leau Gommier, Tareau

Cliffs, Antrim 1000', Fond Figures R. (Collectors:

W. W. Wirth, Jan.-Mar. 1965; O. S. Flint, 1964; R. J.

Gagne, 1966).

The species belongs to group IV. The mesopleura

are hairy. Supraantennals unequal. The male antenna

is oval and somewhat enlarged; the palpal bristles are

short. Hypopygium on each side with 3-4 weak bristles; anal segment very long (0.30 mm), end-hairs

differentiated. Costa 0.55; 2 longer than 1. Halteres

brown.

12. Megaselia (Aphiochaeta) pulliclava, new species

FIGURES 28-30

MALE.—Length 1.55 mm. Frons as broad as long

medially (0.31 mm), dark brown, slightly shining,

sparsely pubescent, with a fine median line. Su subequal, the upper one-half of frontal width apart, the

lower one-eighth. Antials slightly below level of upper

su, vertically below anterolaterals which are distinctly

higher. Second row convex, the bristles equidistant.

Two strong bristles near lower corner of eye. Cheeks

with 2 hairs. Third antennal joint brown, of normal

size, globose; arista 0.39 mm, distinctly pubescent.

Palpi (Figures 28, 29) orange, rather long (0.23 mm),

thickened, in dorsal view of normal shape, slightly depressed above; ventrally on distal half with a large

oval groove which bears a few minute hairs; sides on

basal half with a weak depression; there are 3 short

black spinulae: 2 near apex, the apical being a little

longer, and one on middle of lateral border.

Thorax yellowish brown, subshining. Pubescence

short, not noticeably longer behind. Mesopleura with

a patch of short hairs above. Scutellum with 2 bristles

and 2 minute hairs.

Abdomen opaque, dark brown, venter yellow; tergites with a fine yellow apical seam; tergite 2 slightly

elongated, 3-6 subequal. Pubescence short and scarce,
NUMBER 2 3 15

hind margin of 6th tergite with longer hairs. Hypopygium small, dark brown, on both sides with some

bristly hairs. Anal segment rather long (0.15 mm), pale

yellow, end-hairs not longer than the other hairs.

Legs yellow, hind femur black at tip. Foretibia 0.36

mm; foretarsus slender, metatarsus as long as joints

2-4. Midtibia 0.45 mm, hair-seam reaching apical

fourth, spur nearly as long as metatarsus. Hind femur

about 3 times as long as broad; basal half of ventral

edge with 9 short curved hairs or setulae. Hind tibia

0.65 mm, with 6 distinct posterodorsal setulae on distal

two-thirds and some shorter ones on basal third.

Wing (Figure 30) brownish yellow, veins brown.

Length 1.50 mm, width 0.67 mm. Costa 0.48, ratio of

segments 13:7:4. Cilia moderately short (0.09 mm),

6 pairs between tip of first vein and apex of costa.

Mediastinalis obsolete. Fork acute. Third vein with a

bristle at base; 4th weakly and evenly curved, starting

behind fork without an S-curvature; 5th but little

bent; 6th slightly bisinuate. Anal angle well marked.

Axillary margin with 3 hairs. Halteres dark brown,

stem yellow.

MATERIAL EXAMINED.—Holotype male, pinned specimen (USNM 70245), Dominica, Clarke Hall 11-20

Jan. 1965, light trap (W. W. Wirth).

DISCUSSION.—M. (A.) pulliclava, new species, is

extremely close to M. sublutea Malloch (1912;

U.S.A.), recently redescribed by Borgmeier (1964 p.

344). In M. sublutea the palpi are pale yellow, clubshaped and apparently shorter than in the present

species; furthermore, the halteres of M. sublutea are decidedly pale yellow and the 4th vein is recurved at base.

REMARK.—The ventral groove on distal half of

palpi described above may be due to desiccation; as a

matter of fact, the excavation is present in both palpi

of the type-specimen.

13. Megaselia (Aphiochaeta) definita, new species

FIGURES 31-33

MALE.—Length 1.00-1.30 mm. Head width (paratype) 0.39 mm. Frons quadrate, width 0.20 mm, black

brown, sparsely pubescent, with a distinct median line.

Supraantennals equal, the upper about one-fourth of

frontal width apart, the lower more approximate. Antials in line with upper su, about midway between

them and anterolaterals which are higher near eyemargin. Second row nearly straight, the preocellars

farther from mediolaterals than from each other. Up336-709 O—69 3

per postocular bristle strong. Three bristles near lower

corner of eyes. Third antennal joint black-brown, globose, of normal size; arista 0.30 mm, distinctly pubescent. Palpi slender, yellowish, well bristled.

Thorax black-brown. Pubescence short, longer behind. Mesopleura with about 13 hairs (in holotype),

two of which are longer and bristle-like (length 0.05

mm). Scutellum with 2 bristles and 2 minute hairs.

Abdomen dark brown, venter enclosed. Tergites subequal in length. Hypopygium (Figure 32, from paratype) castaneous, higher than long, right side rounded

behind, with a conspicuous bristle below and an ascending row of 5 setulae; 3 hairs near posterior border. Anal segment brownish yellow, of medium length,

hairy, tergite on each side near apex with a long

bristle directed downward; end-hairs not differentiated.

Legs yellowish brown to dark brown. Foretibia in

holotype 0.19 mm (in paratype 0.26 mm); foretarsus

(Figure 33) 0.37 mm (0.27 mm); metatarsus as thick

as tibia at apex, one-half as long as tibia; joints 2-5

gradually attenuated. Midtibia 0.26 mm (0.32 mm);

end-spur two-thirds of metatarsus. Hind femur 3 times

as long as broad, with about 7 curved setuliform hairs.

Hind tibia 0.36 mm (0.45 mm), with 12 short posterodorsal setulae.

Wing (Figure 31, drawn from holotype) slightly

grayish yellow, veins brown. Length 0.81 mm (1.08

mm), width 0.37 mm (0.46 mm). Costa 0.42 (0.41)

of wing length; Ratio of segments 20:8:5 (25:10:7).

Cilia short (0.05-0.06 mm), there are 4 between rx

and apex of costa. Mediastinalis obsolete. Third vein

with a small bristle at base. Fork narrow, acute, posterior arm slightly curved. Fourth vein starting at fork

(base obsolete in holotype), evenly concave; 5th slightly recurved at base; 6th feebly sinuous; 7th weak.

Axillary margin with 2 hairs. Halteres dark.

FEMALE.—Similar to male. Abdomen brown. Tergites subequal in length, 6th transverse, with a yellowish gland-opening at base medially. The thickening of

the fore-metatarsus is not so conspicuous as in male.

In 3 specimens there are only 3 costal cilia between ri

and tip of costa.

MATERIAL EXAMINED.—Holotype male, on slide

(USNM 70246), Dominica, Cabrit Swamp, 13 June

1964, at light (O. S. Flint). Paratypes (all on slides)

(USNM, CTB) : 1 female, Cabrit Swamp, 18 June

1964 at light, (O. S. Flint); 2 females, Clarke Hall,

21-29 April 1964 (O. S. Flint); 1 male and 2 females,

light trap, 21-31 March 1965 (W. W. Wirth).
16

DISCUSSION.—This small species belongs to group

V and is close to M. coalescens Borgmeier, 1962b

(Brazil), but differs by the structure and chaetotaxy of

the male hypopygium and the dilated foretarsus in

male.

14. Megaselia (Megaselia) deprivata, new species

FIGURE 34

MALE.—Length 1 mm. Frons broader than long

medially (5:4), anterior width 0.25 mm, black-brown,

slightly shining, distinctly pubescent, with a fine

median furrow. Four supraantennals apparently subequal (the lower partly broken), the upper nearly onefourth of frontal width apart, a little farther than the

preocellars. Antials near anterior corner of frons, directed vertically toward median line, almost vertically below anterolaterals, twice as far from upper

supraantennals than from anterolaterals. Third antennal joint dark brown, of normal size and shape; arista

as long as width of frons, strongly hairy. Palpi yellow,

not large, the bristles of median length or short.

Thorax subshining, dark brown, pleura paler below;

mesopleura bare. Scutellum with 4 bristles, the posterior somewhat shorter (10:13) and close together, invading disk and nearly in line with the anterior

bristles.

Abdomen black, dull. Tergites 2-6 gradually a little

reduced in length. Pubescence scarce and short, somewhat longer laterally and on hind margin of 6th tergite.

Hypopygium black, small, higher than long, with some

hairs below; anal tube short, yellowish, end-hairs

inconspicuous.

Legs yellowish coxae enclosed, hind legs darker.

Foretibia 0.26 mm. Foretarsus short, a little longer than

tibia, metatarsus as long as joints 2+3. Hind femur

with 3 longish hairs on basal 4th of ventral edge.

Posterodorsal cilia short, closely set.

Wing (Figure 34) distinctly brownish, veins dark

brown. Length 1.1 mm, width 0.52 mm. Costa 0.41 of

wing length, ratio of segments 17:13. Fork wanting.

Costal cilia short (0.08 mm), about 10 pairs between

humeral crossvein and apex of costa. Mediastinalis

present. Third vein somewhat upcurved on distal third,

slightly thickened apically; 4th slightly concave, a little

deflected at extreme apex; 5th but little curved; 6th

sinuous. Axillary margin with 2 hairs. Halteres black.

MATERIAL EXAMINED.—Holotype male (USNM

70247), Dominica, Freshwater Lake, 21 January 1965

fW. W. Wirth).

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

DISCUSSION.—M. (M.) deprivata, new species,

belongs to group VI. It is a very peculiar species, easily

recognizable by the length and position of the four

scutellars, the short costa, and the absence of a fork.

15. Megaselia (Megaselia) picta (Lehmann)

Phora picta Lehmann, 1822, p. 43.

This cosmopolitan species was recently redescribed

by Borgmeier (1966). The number of scutellars is

variable (2-4). The specimens examined from

Dominica have only 2 scutellars. See literature, synonyms and main characters in Borgmeier, 1966, p. 10;

1967b, p. 96.

MATERIAL EXAMINED.—Eight females (4 pinned, 4

in alcohol), and 4 males (2 pinned, 2 in alcohol),

Dominica: S. Chiltern Est.; Clarke Hall; Fond Figues

R. (Wirth, I.-III. 1965).

16. Megaselia (Megaselia) subpicta (Malloch)

Aphiochaeta subpicta Malloch, 1912, p. 452, pi. 35: fig. 3

[ 9 ; Biscayne Bay, Fla.].

Megaselia (M.) subpicta (Malloch).—Borgmeier, 1962b, pp.

306 [key], 361, fig. 56 [ $ 9 ; U.S.A.; Brazil]; 1966, pp. 2

[key], 8, fig. 1 [wing of type].

Megaselia (M.) malacophaga Borgmeier, 1958 p. 372, figs.

71 [abdomen of female], 76 [hypopygium], 79 [wing], [$ 9 ;

Rio de Janeiro, Petropolis, Curitiba, Nova Teutonia].

Holotype female (USNM 14843), Biscayne Bay, Fla.

Seen 1961.

MALE AND FEMALE.—Length 1.8-3.5 mm. Frons

quadrate, yellow or brown. Supraantennals equal or

nearly so, the upper one-half of frontal width apart.

Antials vertically below laterals. Third antennal joint

small, reddish yellow. Arista long. Palpi yellow, with

moderate bristles.

Thorax yellow. Mesopleura bare. Scutellum with 4

equal bristles.

Abdomen opaque, venter yellow. Tergites black, 2nd

yellow anteriorly, 3-5 with yellow median spots, in

male sometimes 5th entirely yellow. Second tergite laterally with bristles ( $ 9 ), 6th in male with apical

bristles. Hypopygium (see Borgmeier, 1958, fig. 76)

small, brown, with 1 or 2 bristles laterally below. Anal

segment long, sheath-shaped, yellow, end-hairs differentiated. Tergites of female as in Borgmeier's Figure 71 (1958).
NUMBER 2 3 17

Legs yellow, hind femur maculated at tip, with 5-6

strong hairs on basal half of ventral edge. Hind tibial

setulate biseriate, the posterodorsal ones strong.

Wing grayish yellow, veins brown. Costa longer

than one-half of wing (0.54—0.60), First segment equal

or subequal to 2nd. Cilia very short and close. Fourth

vein starting at fork, sometimes slightly deflected at extreme tip; 5th starting vertically below apex of first

vein; veins 5-7 but little bent. Axillary margin with 5

hairs. Halteres dark.

MATERIAL EXAMINED.—Besides the holotype 9 from

Florida and numerous specimens from Brazil, I have

seen 28 males and 24 females from the following localities of Dominica: South Chiltern Est., Sylvania,

Manets Gutter, Springfield Est., and Clarke Hall (W.

W. Wirth collector, January-March, 1965).

DISCUSSIONS.—I am now convinced that M. {M.)

malacophaga Borgmeier is a synonym of M. (M.)

subpicta. The extreme tip of fourth vein seems to be

variable; sometimes the apical deflexion is distinct, as

in the specimens from Brazil; sometimes it is obsolete or wanting, as in the type and the Dominican

specimens.

17. Megasetia (Megaselia) obscuripalpis,

new species

FIGURES 36, 37

FEMALE.—Length 1.70 mm, with the terminal segments extended 1.80 mm. Frons very little longer

medially than broad anteriorly (11:10), width 0.26

mm, yellow, subshining, sparsely pubescent, with a distinct median line. Su unequal, the lower hair-like and

about one-half in length, the upper one-fourth of

frontal width apart. Antials in line with upper su, farther from anterolaterals than from su, anterolaterals

higher placed, near eye-margin. Second row slightly

convex, the bristles equidistant. Upper postocular long.

Two divergent bristles near lower corner of eye; cheeks

with 2 setulae. Third antennal joint not large, globose,

yellowish brown, darkened at tip; arista 0.39 mm, distinctly pubescent. Palpi brown, of normal shape, with

6 short bristles on apical half. Proboscis short.

Thorax subshining, yellow, pleura paler. Pubescence

short, bristle-like behind between 2 dorsocentrals.

Mesopleura bare. Four scutellars, the posterior long,

the anterior thin and one-half in length of the posterior.

(9:18).

Abdomen black-brown, 6th tergite with a yellow

triangular mark at middle extending from base to

apex. Tergites 2-6 subequal in length, 2 reaching sides,

3-6 gradually narrowed, 3-5 transverse, 6th (Figure

37) quadratic, as long as broad. Pubescence extremely

scarce, a few short hairs on 2nd tergite laterally, and

longer bristle-like hairs on hind margin of 6th tergite;

7th tergite with a narrow tergal plate; 8th segment

black, chitinized, glossy. Cerci small.

Legs yellow, hind femur black at tip. Foretibia 0.36

mm; foretarsus slightly longer than tibia (0.41 mm),

metatarsus a little longer than 2+3 . Midtibia 0.52 mm,

seam reaching apical fourth; spur four-fifths of metatarsus. Hind femur 3 times as long as broad; the hairs

on basal half of ventral edge short and inconspicuous.

Hind tibia 0.57 mm, with 11 fine posterodorsal setulae.

Wing (Figure 36) distinctly grayish yellow, veins

dark brown. Length 1.70 mm, width 0.72 mm. Costa

0.51 segment 16:11:3. Cilia short (0.05 mm) and

close, there are 11 pairs between tip of 1st vein and apex

of costa. Mediastinalis obsolete. Fork narrow, acute.

Third vein without a bristle at base; 4th starting at

fork, slightly concave; 5th but little curved, nearly

straight on distal half; 6th sinuous; 7th weak; 4-7

evanescent at apex. Axillary margin with 4 hairs.

Halteres yellow.

MATERIAL EXAMINED.—Holotype female (pinned,

USNM 70248), Dominica, Fond Figues R. 400 ft,

29 April 1966. (R. J. Gagne).

DISCUSSION.—M. obscuripalpis belongs to group

VI. In my key (1962 p. 307) the species runs to couplet 55, but it must be noted that the 2 species mentioned there are wrongly placed: punctifemur is a

Physoptera, and M. subflava belongs to group VII.

There are several undescribed neotropical species

which have the anterior scutellars one-half as long as

the posterior. M. obscuripalpis may be separated from

its allies by the brown palpi, the length of costa and the

ratio of costal segments. The form of the 6th tergite

may also be distinctive.

18. Megaselia (Megaselia) scalar is (Loew)

Phora scalaris Loew, 1866, p. 53.—Borgmeier, 1966, p. 36,

figs. 23, 24 [female abdomen]; 1967b, p. 101.

This widely distributed species was recently redescribed by Borgmeier (1966). The supraantennals are

of equal length and widely separated. The antials are

nearly vertically below the laterals. The female is easily
18

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

recognized by the structure of the 6th abdominal tergite

which is saddle-like. The hypopygium has a single

bristle on each side below; the anal segment is large

and the apical hairs are strong. The female has 4

subequal scutellars (group VI); the male generally

only two (group VII).

MATERIAL EXAMINED.—From Dominica: 2 females

in alcohol, Clarke Hall, March 1965 (Wirth). 1 female,

Clarke Hall, May 1966 (Steyskal); 1 male and 1 female, Clarke Hall, 11-20 Jan. 1965 (Wirth); 1 male,

Clarke Hall, 26-28 Feb. 1965 (Wirth); 1 male, Hodges

R. mouth, reared from Calathea, 22 Feb. 1965

(Wirth).

19. Megaselia (Megaselia) elegantula, new species

FIGURE 35

FEMALE.—Length, with the abdominal segments

fully extended, 2.00 mm. Head width, 0.45 mm. Frons

as broad as long medially (0.23 mm), slightly narrower anteriorly (0.20 mm), reddish yellow, finely

pubescent, with a distinct median line. Lower su rudimental, about one-third in length of the upper which

are about one-fourth of anterior frontal width apart.

Antials reclinate, slightly divergent, much approximate

to upper su, a trifle lower, twice as far from eye-margin

than from su, about midway between su and lx which

are slightly above level of su, near eye-margin. Second

row nearly straight, preocellars farther from each other

than from mediolaterals. Upper postocular well developed; lower postocular strong; no bristle on jowls;

cheeks with 2 weak bristles. Third antennal joint of

normal size, globose, width 0.09 mm, brownish; arista

0.39 mm, distinctly pubescent. Palpi pale yellow, slender, with short bristles on apical half.

Thorax reddish yellow, pleura pale. Pubescence

short, longer behind, Scutellum with 4 bristles of equal

length.

Abdomen: venter pale yellow; tergites subequal in

length, dark brown, with short hairs on hind margins;

2nd tergite with a group of short hairs laterally, no

bristles. Venter with a few hairs on hind margins of

sternites 4-7; 7th segment with a narrow elongate

tergal plate. Cerci distinct.

Legs pale yellow, hind femur maculated at tip. Foretibia 0.28 mm. Foretarsus slender, 0.39 mm. Midtibia

0.36 mm, hair-seam reaching apical fourth. Hind

femur 3 times as long as broad, ventrobasal hairs inconspicuous. Hind tibia 0.48 mm, with a complete

dorsal hair-seam and 6 strong posterodorsal setulae,

the longest below middle nearly as long as width of

tibia.

Wing (Figure 35) grayish yellow, veins brown.

Length 1.30 mm, width 0.52 mm. Costa 0.54 of wing

length, ratio of segments 19:22:7. Cilia moderately

short (0.091 mm), there are about 8 pairs between tip

of 1st vein and apex of costa. Mediastinalis distinct.

Fork acute. Third vein with a conspicuous bristle at

base; 4th starting a little behind fork, distinctly arcuate

on basal half, less so on distal half; 5th slightly sinuate;

6th nearly straight; veins 4-7 obsolete at apex. Anal

margin convex, the angle not pronounced. Axillary

margin with 2 hairs. Halteres dark brown.

Holotype female, on slide (USNM 70249), Dominica, Clarke Hall 1-10 Feb. 1965, light trap (Wirth).

DISCUSSION.—M. (M.) elegantula, new species, belongs to group VI; it is easily recognized by the position of the antials, 4 scutellars, venation and strong

posterodorsal setulae of t3.

20. Megaselia (Megaselia) aurea (Aldrich)

FIGURES 38-40

Phora aurea Aldrich, 1896, p. 477 [$ 9; St. Vincent,

B.W.I.].—Coquillett, 1900 p. 253.

Aphiochaeta aurea (Aldrich).—Brues, 1904, p. 365, pi. VII:

fig. 35.—Malloch, 1912, p. 467.

Megaselia (Af.) aurea (Aldrich).—Borgmeier, 1962b, p. 294,

306 [key]; 1966, pp. 2, 15.

Aphiochaeta rubriventris Borgmeier, 1925b, p. 156, pi. v:

fig. 22.

Holotype female and 4 paratypes females (USNM),

one paratype female (CTB), 1 paratype male (MCZ),

St. Vincent, B.W.I.

Additional notes to my redescription (1966) are as

follows.

MALE.—Length 1.3-2.4 mm. Su unequal. Hypopygium (Figure 38) of medium size, dark brown,

on each side with numerous short hairs and one moderate bristle below. Ventral plate large, broadly

rounded. Anal segment long, yellow, with short hairs

and 2 pairs of longer ones near dorsal edge of tergite,

which are about as long as the apical hairs. Hind

femur (Figure 40) slender, more than 3 times as long

as broad, on basal half of ventral edge with 6 conspicuous curved hairs, the longest is about one-half of
NUMBER 2 3 19

femural width. Costa (male from Dominica) 0.65,

ratio of segments 21:19:4.

FEMALE.—Length 1.5-3 mm. The structure of the

abdominal tergites can best be observed in pregnant

individuals preserved in alcohol (Figure 39). Second

tergite moderately elongated, reaching sides, well

chitinized; 3rd shorter than 2nd, weakly chitinized,

the outlines ill defined, at each side with a transverse

oval whitish area which is shining and often surrounded

by a dark area; 4th and 5th rudimental, nearly membranous but recognizable by the very short pubescence,

4th slightly transverse, 5th subquadrate; 6th well

chitinized, rectangular, longer than broad. The color

of the abdominal segments 1-6 is highly variable (the

terminal segments 7-10 are always black) :

In a paratype, tergite 1 is reddish brown; 2 reddish

brown at middle, black laterally; 3 black-brown; 4-5

black with a deep red tinge; 6 deep black; sides of

abdomen deep red above, black below; venter yellow.

In specimens from Grenada, W.I., tergites 1-5 are

orange, 6 is black; sometimes also 2-3 are black laterally; in one specimen 1-4 are orange, 5-6 black.

In specimens from Dominica, W.I., tergites 1-3 are

brown, 2 sometimes yellow anteriorly; 4 is orange; 5

dark brown, sometimes orange; 6 black, venter orange,

often black laterally.

In specimens from Brazil (Rio de Janeiro; Belem,

Para), Bolivia, Costa Rica, and United States (Florida), tergites 1-6 are orange; in 2 specimens from

Costa Rica 6 is black at sides, yellow at middle.

Costa in paratype 0.62 of wing length, ratio of segment 27:22:5; in specimens from Dominica the first

section is generally equal to 2 + 3 or shorter; in specimens from Rio de Janeiro, Bolivia, Costa Rica, Florida, 1 is longer than 2 + 3.

MATERIAL EXAMINED.—Besides the material listed

in a previous paper (1966) I have seen 6 pinned males

and 25 females, and 50 males and 45 females in alcohol, from the following localities of Dominica:

Clarke Hall, Manets Gutter, Fond Figues River, S.

Chiltern Est. and Pont Casse, January-March 1965

(W. W. Wirth).

DISCUSSION.—M.{M.) aurea comes close to M.

rubronigra Borgmeier (Costa Rica), but the latter

is much darker and the costa exceeds the tip of 3rd

vein. The female of M. aurea can be easily recognized

by the peculiar structure of the 3rd tergite and the color

of the abdomen.

21. Megaselia (Megaselia) striativentris Borgmeier

Megaselia {Megaselia) striativentris Borgmeier, 1962b, pp.

306 [key], 367 [ $ $ ; Nova Teutonia; Curitiba, Parana,

Petropolis, R. J., Costa Rica].

Holotype male (CTB), Nova Teutonia, Brazil.

Paratypes males and females (CTB, USNM).

Frons quadrate or subquadrate, dark brown. Su

unequal. Antials lower than upper su, removed from

eye-margin, sometimes midway between su and lx the

latter nearly in line with upper su. Third antennal

joint orange, darkened at tip.

Thorax yellow. Mesopleura bare. Scutellum with

4 equal bristles.

Abdomen subshining, venter yellow. Tergites black

with yellow apical bands; 2nd with bristles laterally

( S 9 ); 6th tergite in male with apical bristles. Hypopygium small, brown, hairy. Anal segment moderately

short, as long as epandrium, yellow, with short hairs.

Legs yellow, hind femur black at tip. Foretarsus

slender. Hair-seam of midtibia nearly complete. Hairs

on ventral edge of hind femur of moderate length.

Hind tibia with 6-8 setulae, the upper weak, the remainder strong. Wing slightly brownish. Costa 0.56

(holotype), ratio of segments 16:15:5. Cilia long (0.12

mm). Fourth vein starting at fork, 5-6 sinuous. Haltere

brown [not yellow; sometimes ochreous brown].

MATERIAL EXAMINED.—More than 150 specimens

from Brazil and some from Costa Rica. From Dominica

I have seen 10 pinned specimens and 150 specimens

preserved in alcohol (59) , from the following localities: Clarke Hall, S. Chiltern Est., and Manets Gutter

(Jan.-March 1965, W. W. Wirth).

DISCUSSION.—M. (M.) striativentris can be confused with M. fimbriata Borgmeier (1962b, Brazil)

which is very similar but has some anterodorsal setulae

on hind tibia and the antials are nearly vertically below

the anterolaterals.

22. Megaselia (Megaselia) seticauda (Malloch)

Aphiochaeta seticauda Malloch, 1914, p. 32 [& 9 ; Irazu,

Costa Rica].

Megaselia (M.) seticauda (Malloch).—Borgmeier, 1962b, p.

309 [key], 413, figs. 74 [hypopygium], 93 [wing $ ]; 1966,

pp. 3 [key], 44.

Megaselia (M.) disparisecta Borgmeier, 1962b, p. 414, figs.

86 [abdomen, 9], 92 [wing, 9]; 1966, p. 45 [syn.]; 1967c,

p. 405 [syn.].

This species, originally described from Costa Rica, is

widely distributed. I have seen the types and other
20

specimens from Costa Rica; furthermore from Brazil

(Petropolis, Nova Teutonia), Ecuador, Mexico

(Nogales), United States (Texas) and one female

from Dominica, Clarke Hall, 11-20 January 1965,

light trap (Wirth).

The species was redescribed by the present writer

in 1962 (b) and 1966. There are only two supraantennals in both sexes, close together; the antials are close

to su and nearly parallel. The hypopygium has on each

side behind a series of 4 or 5 strong moderately long

bristles; anal segment long. Sixth tergite in female long

and narrow. Costa 0.52 in male, up to 0.58 in female;

first costal section equal to 2nd, or 2nd longer than 1st.

Cilia very short. Halteres yellow or brown.

DISCUSSION.—The female is easily recognized by the

long 6th tergite; the male by the bristles of the hypopygium. The length of the 2nd costal segment is variable and caused the synonym M. disparisecta.

23. Megaselia (Megaselia) pleurofascia Borgmeier

Megaselia (Megaselia) pleurofascia Borgmeier, 1962b, p. 418

[ j {; Nova Teutonia].

One pinned male from Dominica, Clarke Hall, 11-

20 March 1968 (Wirth) seems to be referable to this

species, originally described from southern Brazil.

24. Megaselia (Megaselia) hypochaeta, new species

FIGURES 41-43

FEMALE.—Length 2.4 mm. Head width 0.65 mm.

Frons as broad as long laterally (0.28 mm), slightly

longer medially (0.32 mm), yellow, sparsely pubescent,

with a fine median line. Four supraantennals, the lower

rudimental, placed nearly vertically below the upper

which are close together, occupying about one-fourth

of frontal width. Antials a little lower than upper

supraantennals, about as far from them as from eyemargin; anterolaterals distinctly higher, near eyemargin. Second row slightly convex anteriorly, the

bristles equidistant; the preocellars farther apart from

each other than upper supraantennals. Ocular cilia

distinct; upper postocular bristle strong; one lower

postocular bristle, and one on jowls; cheeks narrow,

with 2 moderate bristles. Third antennal joint reddish

yellow, small globose, diameter 0.1 mm; arista distinctly pubescent. Palpi of normal shape, with short

bristles.

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Thorax yellow, with 2 dorsocentrals; pubescence

short, longer and bristle-like in front of scutellum.

Mesopleura bare. Scutellum with 2 bristles and 2

minute hairs.

Abdomen (Figures 41-42) reddish yellow, tergites

1-2, 6 and terminal segments blackish brown; 2nd tergite yellow on anterior half, with some short bristles

laterally. Segments 2-5 mostly membranous, bare; 3rd

segment with an ill-defined, weakly chitinized small

dark plate. Posterior border of 5th segment blackish;

6th tergite trapezoid. Terminal segments tubiform,

fleshy. Venter yellow, except 6th segment and a subapical band of 5th which are blackish. The 4th ventrite has posteriorly on each side a curved row of about

20 closely set black setulae (Figure 42) ; in front of

them there are some sparse bristly-hairs; 6th ventrite

with sparse short hairs inserted in chitinous spots; 7th

segment with a narrow striplike tergite dilated anteriorly; a similar narrow sternite is present, slightly

dilated posteriorly. Cerci indistinct.

Legs yellow; hind femur maculated at tip. Foretibia

0.44 mm; foretarsus 0.52 mm, all joints longer than

broad; fore-metatarsus about as long as joints 2-4 combined. Midlegs missing in type. Hind femur 0.91 mm,

width 0.28, with about 9 longish setuliform hairs.

Hind tibia 0.85 mm, with 9 posterodorsal setulae, the

3 on median 3rd are stronger and nearly as long as

width of tibia.

Wing (Figure 43) grayish yellow, veins pale brown.

Length 2.08 mm, width 0.78 mm. Costa 0.53 of wing

length, ratio of costal segments 20: 15 :4. Mediastinalis present. Costal cilia short, more or less 10 pairs

between tip of 1st vein and apex of costa. Fork acute,

narrow. Fourth vein slightly concave, starting at fork,

basal S-curvature indistinct; 5th arcuate before middle,

remainder nearly straight; 6th feebly sinuous; 7th weak

but distinct. Axillary margin with 4 hairs. Halteres

yellow.

MATERIAL EXAMINED.—Holotype female, on slide

(USNM 70250), Dominica, Clarke Hall, 21-31 Jan.

1965, W. W. Wirth, light trap.

DISCUSSION.—A striking species, easily recognized

by the structure of the abdomen and especially by the

two curved rows of setulae on fourth ventrite. M. aurea

is similar in general appearance, but has four scutellars, longer costa, and the structure of the abdomen

is quite different.
NUMBER 2 3

25. Megaselia (Megaselia) leptacina, new species

FIGURES 44-46

FEMALE.—Length 1.40-1.50 mm. Head width 0.42

mm. Frons dark brown, as broad as long medially

(0.20 mm), lateral length 0.15 mm, sparsely pubescent,

with a fine median line. Supraantennals unequal, the

lower thin and a little more than one-half of upper

(5:9), the upper about one-fourth of frontal width

apart, the lower one-eighth. Antials close to lower corner of frons, slightly below level of upper su, anterolaterals a trifle above level of su, near eye-margin.

Second row slightly convex, the bristles equidistant.

One upper postocular, and one lower; jowls-cheeks

with a series of about 4 hairs. Third antennal joint

black-brown, of normal size, globose, width 0.09 mm;

arista as long as head width, finely pubescent. Palpi

yellow, slender, of normal length, with 5 long bristles

on apical half.

Thorax reddish yellow, pleura castaneous but epipleura pale yellow. Pubescence short. Mesopleura bare.

Scutellum with 2 bristles and 2 minute hairs.

Abdomen (Figures 44) dark brown, venter grayish

brown. Tergites 2-5 subequal in length, all reaching

sides; 6th elongated, rounded behind, with a distinct

gland-opening at base. Pubescence short and scarce,

more conspicuous near lateral corner of 2nd and on

hind margin of 5th tergite. Venter sparsely hairy.

Terminal segments fleshy, tubiform; 7th segment with

a pen-shaped tergal plate, 7th sternite reduced to a

narrow median strip; 8th withdrawn; cerci indistinct.

Legs pale yellow, coxae enclosed, hind femur

brownish yellow on distal half. Foretibia 0.26 mm;

foretarsus slender, 0.36 mm, metatarsus longer than

joints 2 + 3. Midtibia 0.36 mm; hair-seam reaching

apical fourth; spur two-thirds of metatarsus. Hind

femur (Figure 46) slender, length 0.52 mm, width 0.13

mm, basal half of ventral edge with 5-6 conspicuous

curved hairs, the longest one-half of width of femur

in length. Hind tibia 0.46 mm, with 15 fine short

posterodorsal cilia.

Wing (Figure 45) grayish yellow, veins pale

brown. Length 1.19 mm, width 0.52 mm. Costa 0.56 of

wing length, ratio of segments 21:18:6. Cilia short

(0.052 mm) and close, there are 10 pairs between tip

of 1st vein and apex of costa. Mediastinalis weak. Fork

acute. Third vein without a bristle at base; 4th arcuate;

5th curved before middle, remainder nearly straight;

21

6th sinuous. Anal margin broadly convex; axillary

margin with 2 hairs. Halteres yellow.

MATERIAL EXAMINED.—Holotype female, on slide

(USNM 70251), Dominica, S. Chiltern Est., 20 Feb.

1965 (W. W. Wirth). Paratypes females (USNM,

CTB) 4 females on slide, same locality and date; one

pinned female, Clarke Hall, 11-20 Mar. 1965 (W. W.

Wirth).

DISCUSSION.—M. (M.) leptacina, new species, is

close to M. tubulifera Borgmeier (1962b, Brazil), but

the latter has the sixth tergite transverse.

26. Megaselia (Megaselia) imitatrix, new species

FIGURES 47-48

FEMALE.—Length 1.70 mm, extended in alcohol

2.00-2.50 mm. Frons subquadrate, broader than long

laterally (4:3), yellow, sometimes infuscated above,

shortly pubescent, with a distinct median line. Upper

postantennals one-fifth or less of frontal width apart,

more approximate than the preocellars. Lower postantennals very weak, hardly one-half of upper in

length. Antials about midway between anterolaterals

and upper postantennals, nearly in line with latter;

anterolaterals a little higher near eye-margin. Second

row slightly convex. Eyes large. Jowls with 2 divergent

bristles. Cheeks narrow, with 4 hairs. Third antennal

joint globular, width 0.13 mm, dusky yellow; arista

0.5 mm distinctly pubescent. Palpi pale yellow, of normal shape, well bristled.

Thorax yellow. Mesopleura bare. Scutellum with 2

bristles and 2 fine hairs.

Abdomen (Figure 48) venter yellow, laterally

slightly darkened on segments 3-5, the darkened areas

separated by yellow intervals. Tergites black, 1-2 yellow on anterior half; 3-5 gradually narrowed, subequal

in length, with a yellow median spot anteriorly not

reaching posterior border; 6th tergite shorter and

broader than 5th, nearly semicircular anteriorly, with

a transverse median yellow spot; 7th segment with a

black trapezoid tergite. Pubescence on dorsum very

scarce and short, longer on hind margin of 6th and

7th tergites; venter distinctly hairy on segments 3-6;

7th segment with a small sternal plate provided with

2 bristles. Cerci small.

Legs yellow, hind femur black at tip, on basal half

of ventral edge with curved hairs, the basal ones short,

the remaining setiform. Hind tibia with 6 strong

widely placed posterodorsal setulae.
22

Wing (Figure 47) yellowish, veins brown. Length

1.30 mm to 1.50 mm. Costa about 0.5, ratio of segments 24:17:7. Cilia short (0.06 mm). Mediastinalis

present. Fork large, acute; 3rd vein without a bristle

at base; 4th starting behind fork, evenly curved; 5th

but little curved; 6th weakly sinuous. Axillary margin

with 4 hairs. Halteres yellow, penduncle brown.

MALE.—Length 1.60 mm, in alcohol up to 2.00 mm.

Similar to female. Tergites black, 1-2 and 6 yellow on

anterior half, 2-4 with a yellow median spot not reaching posterior border, 5th sometimes nearly entirely yellow. Pubescence scarce and short, 6th with bristly

hairs on hind margin. Hypopygiura small, blackbrown, with hairs, at left with a single conspicuous

bristle. Anal segment long (0.23 mm), pale yellow,

end-hairs differentiated but not so strong as in M. {M.)

scalaris. Foretarsus slender, longer than tibia (4.3).

Costa nearly one-half wing length, first segment generally a little longer than 2 + 3.

Holotype (in alcohol; USNM 70252), Dominica,

Cabrit Swamp, 23 Feb. 1965, light trap (Wirth). Paratypes 9 females and 15 males, from the following localities: Cabrit Swamp, 23 Feb. 1965 [1 female]; Clarke

Hall, 21-31 Jan. 1965 (Wirth); Puerto Rico, Barrio

Rio Grande, El Verde, 1963-1966, sticky traps, G. E.

Drewery.

DISCUSSION.—M.(M.) imitatrix, new species, is

very similar in appearance to M. (M.) scalaris (Loew),

but the female of the latter has four scutellars and 9

and $ have four subequal postantennals; it differs

also by the position of the antials and the structure of

the abdominal tergites 5-6.

27. Megaselia (Megaselia) buccata, new species

FIGURES 49-51

MALE.—Length (somewhat extended in balsam)

2.00 mm. Head width 0.57 mm. Frons distinctly higher than broad, width 0.23 mm, lateral length 0.28 mm,

median length 0.33 mm, black-brown, distinctly pubescent, with a distinct median line. Su equal, the upper

two-thirds of frontal width apart, the lower about onefourth. Antials close to upper su and near eye-margin,

vertically below anterolaterals, the latter distinctly

higher, twice as far from antials as antials from upper

su. Second row slightly concave anteriorly, nearly

straight, the preocellars farther from each other than

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

from mediolaterals. One long lower postocular bristle;

jowls-cheeks with an ascending row of 6 bristles, the

upper 3 slightly weaker. Third antennal joint dark

brown, globose, of normal size, width 0.10 mm. Palpi

pale yellow, of normal shape, with 5 bristles of moderate length. Proboscis of peculiar shape; labrum

small, triangular; labellae (Figure 50) yellow, broad,

with a few fine hairs, apparently without pseudotracheae.

Thorax brownish yellow, pleura pale. Pubescence

short, longer behind. Mesopleura bare. Scutellum

with 2 bristles and 2 hairs.

Abdomen: venter yellow; tergites dark brown; 2

and 6 a little elongated, 3-5 subequal in length. Pubescence short, some bristly hairs on 2nd laterally and

on hind margin of 6th. Hypopygium (Figure 57) dark

brown, small, with short hairs; anal segment pale yellow, long, cylindrical, slightly curved downwards, endhairs not longer than hairs on tergite.

Legs yellow, hind femur with a black macula at

apex. Foretibia 0.36 mm, with a dorsal row of short

spinuli; foretarsus slender, 0.49 mm, metatarsus a

little longer than joints 2 + 3. Midtibia 0.52 mm, hairseam complete, 4 distinct anterodorsal and 4 postercdorsal setulae on upper two-thirds; spur four-fifths of

metatarsus. Hind femur 3 times as long as broad, width

0.26 mm, length 0.75 mm, ventrobasally with a row

of 5 conspicuous curved hairs not reaching middle.

Hind tibia 0.65 mm, setulae biseriate: 6 strong posterodorsal which are as long as width of tibia, and 4

anterodorsal which are weaker.

Wing (Figure 49) infuscated, veins dark brown.

Length 1.48 mm, width 0.62 mm. Costa 0.52, ratio of

segments 24:19:11. Cilia short (0.065 mm), close,

about 10 pairs between tip of first vein and apex of

costa. Fork long and narrow, acute. Mediastinalis obsolete; 3rd vein with a small bristle at base; 4th concave starting at fork with a slight S-curvature, 5th but

little curved, 6th very feebly sinuous. Axillary margin with 5 hairs. Halteres dark brown.

MATERIAL EXAMINED.—Holotype male (on slide;

USNM 70253), Dominica, Clarke Hall, 8-10 Jan.

1965, Malaise trap (Wirth).

DISCUSSION.—M.(M.) buccata, new species, is a

very striking species of group VII. It runs to M. anterospinosa in my key (1926b), but is completely different

by the peculiar labellae, the elongate frons, the frontal

chaetotaxy, and the venation.
NUMBER 2 3 23

28. Megaselia (Megaselia) polita (Enderlein)

FIGURES 52-53

Aphiochaeta polita Enderlein, 1912, p. 30 [9 ; Hammonia].—

Malloch, 1914, p. 29 [Costa Rica].

Megaselia (Megaselia) polita (Enderlein).—Borgmeier,

1935a , p . 449 , pi. 5 : fig. 7 [win g of holotype , $9];

1962b , p . 30 7 [key].

Holotype female (CTB), Hammonia, S. Catarina,

Brazil (Luederwaldt).

Frons quadratic, black, glossy. Supraantennals subequal. Antials and anterolaterals nearly in line with

upper su. Thorax black, shining. Mesopleura bare.

Scutellum with 2 bristles. Abdomen black. Hypopygium

glossy, castaneous. Anal segment of medium length,

yellow, end-hairs not differentiated. Legs black. Foremetatarsus (Figure 52) somewhat thickened in both

sexes. Hind tibial setulae strong. Hairs on ventral edge

of hind femur short. Wing (Figure 43, drawn from

holotype) distinctly brownish; length 1.56 mm, width

0.67 mm; costa 0.56, cilia moderately short (0.09

mm), closely set, about 11-12 pairs between 1st vein

and apex, ratio of segments 25:24:10. Third vein

without bristle at base; 4th starting at fork or behind

it, slightly deflected at tip. Anal margin broadly convex. Axillary margin with 2 hairs. Halteres black.

MATERIAL EXAMINED.—Besides the type, I have seen

numerous specimens from Nova Teutonia, S. Catarina,

some specimens from Petropolis, Rio de Janeiro; one

from Peru (Tingo Maria); two from Costa Rica (Suiza

de Turrialba). From Dominica: 1 female, pinned. S.

Chiltern Est. (Wirth); 2 females and 1 male in alcohol, Clarke Hall, March 1965 (Wirth).

DISCUSSION.—M.(M.) polita can be confused with

M. littoralis Malloch (Costa Rica), but the latter has

slender foretarsi, the base of wing is yellow, the cilia are

widely placed (see Borgmeier, 1962b, fig. 66), and the

forelegs are yellow. Also close to M. anthracina Borgmeier (Brazil), which has a shorter costa, unequal su,

and bristly hairs on ventral edge of hind femur.

29. Megaselia (Megaselia) femoralis (Enderlein)

FIGURES 54-55

Aphiochaeta femoralis Enderlein, 1912, p. 30 [9 ; Hammonia].—Malloch, 1914, p. 29 [ 9 ; Costa Rica].

Megaselia (Megaselia) femoralis (Enderlein).—Borgmeier,

1932, p. 51 [9 ; Costa Rica]; 1935a, p. 450, pi. 5: fig. 13

[type]; 1966, pp. 4 [key], 56 [8 9 ; Texas, U.S.].

Lectotype female and 3 paratypes females (CTB),

from Hammonia, S. Catarina, Brazil (Luederwaldt).

336-709 O—69 4

The female types were redescribed by Borgmeier

(1935a) and the male was first described in 1966. The

hypopygium (Figure 54) is small, black, and shows

several fine short hairs on both sides. The left ventral

plate (Figure 55) is pointed at tip, and the right is

broadly rounded. The anal segment is of medium size,

yellow; the apical hairs are not differentiated.

MATERIAL EXAMINED.—Besides the types, I have

seen numerous specimens from Brazil (Nova Teutonia,

Curitiba, Petropolis) and some from Costa Rica (Suiza

de Turrialba, Collector P. Schild). From Dominica, 36

females and 18 males are before me, mostly pinned,

some in alcohol and 4 on slides, Clarke Hall, Jan.-Feb.

1965 (Wirth).

DISCUSSION.—M. (M.) femoralis belongs to group

VII. It is easily recognized by the striking coloration

of the hind femur, the glossy froms and by venational

characters.

30. Megaselia (Megaselia) enderleini (Brues)

FIGURES 56, 57

Aphiochaeta enderleini Brues, 1912, p. 135 [nomen novum]

Megaselia {Megaselia) enderleini (Brues).—Borgmeier,

1935a, p. 447, fig. 16.

Aphiochaeta minuta Enderlin, 1912, p. 36 [$ ° ; Hammonia,

Brazil, not Aldrich].

Lectotype male (CTB) and 1 paratype male (CTB),

Hammonia, Brazil.

MALE.—Length 1.3-1.6 mm. Head width 0.42 mm.

Frons broader than long (4:3), black, subopaque,

scarcely pubescent, with a distinct median furrow.

Lower su variable in length, one-half or three-fourths

of upper, nearly vertically below upper which are about

one-fifth of frontal width apart. Anterolaterals nearly

in line with antials and upper su; antials farther from

su than from laterals. Third antennal joint black, globose. Palpi black-brown, well bristled.

Thorax black. Mesopleura bare. Scutellum with 2

bristles and 2 minute hairs.

Abdomen black, opaque. Tergites subequal, scarcely

and shortly pubescent, longer bristle-like hairs on hind

margin of 6th. Hypopygium black-brown, with hairs

and on each side with a single bristle. Anal segment

very short, black-brown, end-hairs not differentiated.

Legs black to black-brown. Foretarsus slender,

Metartarsus longer than 2 + 3. Hind femur rather

broad (ratio 8:21, 9:25, 10:25, the latter in specimens

from Dominica), ventral hairs on basal half short,

inconspicuous. Hind tibia with fine cilia.
24

Wing (Figures 56-57) nearly hyaline, somewhat

brownish in specimens from Dominica; heavy veins

brown, light veins pale (distinct in specimens from

Dominica). Length (lectotype) 1.40 mm, width 0.59

mm. Costa 0.44-0.45 (lectotype) 0.46 (Nova Teutonia), 0.45 (Petropolis), 0.46 (Dominica). Ratio of

segments 19:8:5 (lectotype); 10:6:3 (Nova Teutonia), 12:7:4 (Petropolis). Cilia long (0.13 mm),

there are 4 or 5 pairs between tip of 1st vein and apex

of costa. Third vein without bristle at base; 4th slightly

concave, 5th slightly recurved at base, 6th weakly sinuous; anal margin concave, axillary margin with 5-6

hairs. Halteres black.

FEMALE.—Similar to the male. Tergites subequal.

Pubescence scarce. There are 5-6 costal cilia between

rx and r8.

MATERIAL EXAMINED.—Type and 1 paratype, Hammonia, Brazil; many specimens from Nova Teutonia,

Brazil, and Petropolis, Brazil; a few from Costa Rica

and Mexico, DF.; 35 specimens, mostly in alcohol,

from Dominica (Jan.-Mar. 1956) Clarke Hall, S.

Chiltern Est., PontCasse (Wirth) ; 1, Freshwater Lake,

rV.66(R.J.Gagne).

DISCUSSION.—M. (M.) enderleini is a striking

species; form, color, and venation of wing are characteristic. Comes close to M. atricolor Borgmeier

(Brazil), but the latter has four equal supraantennals.

31. Megaselia (MegaseUa) labellifera, new species

FIGURES 58-60

MALE.—Length 1.40 mm. Head width 0.46 mm.

Frons black, slightly broader than long laterally (6:5),

width 0.23 mm, pubescent, with a distinct median

line. Su unequal, the lower thin and about one-half

of the upper, the upper nearly one-fourth of frontal

width apart, the lower one-ninth. Antials in line with

upper su, farther from them than from anterolaterals

which are a little higher near eye-margin. One upper

and one lower postocular bristle. Cheeks with 2 or 3

setulae. Third antennal joint black, of normal size, globose; arista 0.39 mm, finely pubescent. Palpi brown, of

normal shape, with bristles of moderate length. Proboscis short, dark brown; labellae large, shield-shaped,

each labella excavated ventrally.

Thorax black-brown. Pubescence short, longer behind. Mesopleura bare. Scutellum with 2 bristles and 2

minute hairs.

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Abdomen black-brown, venter dark. Tergites subequal in length, with scarce short pubescence, hairs on

hind margin of 6th tergite bristle-like. Venter sparsely

hairy. Hypopygium (Figure 59) small, dark brown,

with some fine bristly hairs; ventral plate large, hairy.

Anal segment short, yellowish brown, end-hairs hardly

longer than the other hairs. Legs castaneous. Foretibia

0.28 mm; fore tarsus slender; metatarsus as long as

2 + 3. Midtibia 0.33 mm. Hind femur (Figure 60)

nearly 3 times as long as broad, on basal half of ventral edge with about 7 longish curved hairs. Hind tibia

0.46 mm, with 10 short posterodorsal setulae.

Wing (Figure 58) distinctly yellowish brown, veins

brown. Length 1.17 mm, width 0.52 mm. Costa 0.48

of wing length, ratio of segments 18:14:7. Cilia short

(0.078 mm), there are 6 pairs between tip of 1st vein

and apex of costa. Mediastinalis distinct. Fork acute.

Third vein without a bristle at base; 4th vein curved

near base, remainder nearly straight; 5th and 6th

slightly sinuous; 7th weak. Axillary margin with 2

hairs. Halteres black.

Holotype, on slide (USNM 70254), Dominica, S.

Chiltern Est. 20 Feb. 1965 (Wirth).

DISCUSSION.—M. (M.) labellifera, new species,

comes close to M. labellata Borgmeier (1962b, Brazil),

but the latter has the labellae larger and of red color,

the supraantennals are equal, the antials are vertically

below ll5 and the venation is different.

32. MegaseUa (MegaseUa) metatarsalts, new species

FIGURES 64-67

MALE.—Length 1.00 mm. Head width 0.39 mm.

Frons broader than long medially (3:2), width 0.20

mm, yellow, with sparse fine hairs and a fine median

line. Lower su reduced to hairs, upper about one-fourth

of frontal width apart. Antials close to anterolaterals,

both practically in line with upper su. Second row

nearly straight, preocellars a little farther from mediolaterals than from each other. Upper postocular developed; 2 unequal bristles near lower corner of eye.

Third antennal segment of normal size, globose, width

0.078 mm, brownish; arista 0.31 mm, distinctly

pubescent. Palpi yellow, slender, with 6 long bristles on

apical half.

Thorax yellow. Pubescence short. Mesopleura bare.

Scutellum with 2 bristles and 2 short hairs.

Abdomen: venter yellow; tergites pale brown, subequal in length, with sparse hairs, those on hind margin
NUMBER 2 3 25

of 6th not conspicuously longer. Hypopygium (Figure

66) brown, asymmetrical, sides apparently connected

beneath anal segment; each side with several bristly

hairs, lobe of left side broadly rounded; lobe of right

side narrow, a short tooth-like process and a short digitiform process (dotted line in Figure 66) are

probably part of the connecting bridge. Anal segment

short, yellow, end-hairs short.

Legs entirely yellow, coxae enclosed. Foretibia 0.19

mm; fore-metatarsus (Figure 67) strongly dilated,

broader than tibia, with a broad longitudinal furrow

on ventral side, length 0.14 mm, width 0.052 mm;

joints 2-5 small, combined 0.16 mm. Midtibia 0.29

mm, hair-seam reaching apical fourth; end-spur twothirds of metatarsus. Hind femur 3 times as long as

broad, on basal half of ventral edge with 5 long curved

hairs (Figure 65). Hind tibia 0.39 mm, with 6 fine

short posterodorsal cilia beginning at basal fourth.

Wing (Figure 64) grayish yellow, veins pale brown.

Length 0.98 mm, width 0.39 mm. Costa 0.48 of wing

length, ratio of segments 15:11:5. Cilia short (0.05

mm), about 8 pairs between tip of 1st vein and apex of

costa. Mediastinalis missing. Fork acute. Third vein

without a bristle at base; 4th weakly arcuate, especialy on basal half; 5th and 6th feebly sinuous; 7th

weak. Anal margin very broadly convex; axillary margin with 2 hairs. Halteres dark.

Holotype on slide (USNM 70255), Dominica,

Clarke Hall, 1-10 Feb. 1965, light trap (Wirth).

DISCUSSION.—In my key (1962b) M. (M.) metatarsalis, new species, runs to M. flavipes, but the latter

is larger, has tarsal joints 1-3 slightly dilated and the

costa longer.

33. Megaselia (Megaselia) tarsodes, new species

FIGURES 61-63

MALE.—Length 1.00 mm. Head width 0.39 mm.

Frons broader than long laterally (5:4), width 0.19

mm, lateral length 0.15 mm, median length 0.16 mm,

dark brown, sparsely pubescent, with a distinct median

line. Lower supraantennals thin and a little more than

one-half of upper, which are one-fourth of frontal

width apart (the lower one-fifth). Antials in line with

upper su, nearly vertically below anterolaterals which

are a little higher. Second row convex, the preocellars

farther from each other than the upper supraantennals. Upper postocular bristles distinct. One lower

postocular, and one bristle on jowls; cheeks narrow,

with one setula. Third antennal segment of normal

size and shape, dark brown; arista 0.32 mm, distinctly

pubescent. Palpi (Figure 63) slender, length 0.09 mm,

yellowish brown, with 5 long bristles on apical half.

Thorax reddish yellow. Pubescence short, 2 dorsocentrals. Mesopleura bare. Scutellum with 2 bristles

and 2 minute hairs.

Abdomen brown, venter enclosed. Tergites subequal

in length. Pubescence short, scarce; 6th segment behind with a circle of bristly hairs. Hypopygium brown,

on each side with a few weak bristles or bristly hairs.

Anal segment very short, hairy, end-hairs not differentiated.

Legs yellow, hind femur maculated at tip. Foretibia

0.23 mm, width at tip 0.04 mm. Foretarsus (Figure 62)

longer than tibia, length 0.33 mm, joints 1-4 thickened,

gradually attenuated, all joints longer than broad;

metatarsus nearly as broad as end of tibia (0.03 mm).

Midtibia 0.30 mm, hairseam ending below middle,

posterodorsal cilia distinct; end-spur as long as metatarsus. Hind femur 0.45 mm, about 3 times as long

as broad, with about 7 conspicuous curved hairs on

basal half of ventral edge. Hind tibia 0.42 mm, with

10 fine posterodorsal setulae.

Wing (Figure 61) slightly yellowish, veins brown.

Length 1.00 mm, width 0.47 mm. Costa 0.47 of wing

length, ratio of segments 24:15:8; cilia short (0.06

mm), about 7 pairs between tip of 1st vein and apex

of costa. Mediastinalis distinct. Posterior arm of fork

curved. Fourth vein starting behind fork, evenly

arcuate; 5th and 6th veins slightly sinuous; 7th weak.

Anal angle not pronounced; axillary margin with 2

hairs. Halteres dark.

Holotype, (USNM 70256, on slide), Dominica,

21-29 Apr. 1964, light trap (O. S. Flint).

DISCUSSION.—M. (M.) tarsodes, new species, is a

small species close to M. communis Borgmeier, recognizable by the dilated foretarsus.

34. Megaselia (Megaselia) substricta, new species

FIGURES 68, 69

FEMALE.—Length 1.2 mm. Head width 0.41 mm.

Frons broader than long laterally (4:3), width 0.20

mm, median length 0.22 mm, brownish yellow, with

sparse pubescence and a fine median line. Lower

supraantennals rudimental, close together; upper

supraantennals one-fourth of frontal width apart.

Antials nearly in line with upper supraantennals,
26

twice as far from them as from eye-margin. Anterolaterals a trifle higher, close to antials. Second row

convex anteriorly, the preocellars farther apart than

upper supraantennals. Upper postocular bristle distinct; ocular cilia weak. One lower postocular bristle,

and one on jowls; cheeks with one moderate bristle.

Third antennal joint brown, globose, length about 0.09

mm. Arista 0.44 mm, distinctly pubescent. Palpi

slender, whitish yellow, with about 5 bristles on apical

third. Labrum piriform, labellae slender.

Thorax yellow, with 2 dorsocentrals. Pubescence

short, somewhat longer before scutellum. Mesopleura

bare. Scutellum with 2 bristles and 2 minute hairs.

Abdomen yellow, venter enclosed; tergites (Figure

69) brown, 3-6 modified. Third tergite as long as 2nd,

constricted at middle, the sides excavate; 4th short,

5th and 6th trapezoid; 4th and 5th with an anterior

seam (indicated by dotted lines) which is normally

covered by the hind border of the preceding tergite;

6th with a gland opening at base. Pubescence short

and very scarce. 7th tergite subtriangular, concave

anteriorly. Terminal segments fleshy, short.

Legs entirely yellow, hind femur not maculated.

Foretibia 0.28 mm; foretarsus slender, length 0.36

mm; metatarsus slightly longer than joints 2-3 combined (11:10). Midtibia 0.37 mm; end-spur nearly as

long as metatarsus (14:17). Hind femur 0.52 mm,

width 0.18 mm, ventrobasally with some short hairs

and about 5 fine longish hairs on basal half. Hind

tibia 0.48 mm, with about 12 short posterodorsal cilia.

Wing (Figure 68) slightly grayish, veins pale. Length

1.23 mm, width 0.52 mm. Costa 0.47 of wing length,

ratio of segments 18:13:6. Cilia moderately short

(0.078 mm), there are about 7 pairs between tip of

1st vein and apex of costa. Mediastinalis missing. Fork

acute. Third vein with a conspicuous bristle at base;

4th vein obsolete at base, slightly concave; 5th but

little curved; 6th feebly sinuous; 7th weak. Axillary

margin with 2 hairs. Halteres dark.

MALE.—Length 1.17 mm. Costa 0.5 of wing length.

There are 6 pairs of cilia between tip of 1st vein and

apex of costa. Hypopygium pale brown, with fine hairs;

anal segment short (0.052 mm), dirty yellow, with

short hairs, apical hairs not differentiated.

MATERIAL EXAMINED.—Holotype female, (USNM

70257, on slide), Dominica, Clarke Hall, 1-10 May

1964, light trap (O. S. Flint). Paratypes (USNM,

CTB): one female on slide, 21-29 April 1964, same

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

locality and collector; 4 females and 1 male in alcohol, Clarke Hall, 11-20 Mar. 1965 (W. W. Wirth).

DISCUSSION.—M. (M.) substricta, new species,

comes very close to M. mendax Borgmeier (1962b,

Brazil), but the latter has a longer costa (see Figure

70, wing of paratype) and the structure of the third

and fourth tergite is different.

35. Megaselia (Megaselia) diplochaeta, new species

FIGURES 71-73

MALE.—Length 1.00 mm. Head width 0.33 mm.

Frons broader than long laterally (4:3), width 0.20

mm, black-brown, shining, pubescent, with a distinct

median furrow. Four supraantennals subequal (4:3),

the upper about one-fourth of frontal width apart, the

lower one-eighth; the upper are more approximate

than the preocellars. Antials in line with upper su,

vertically below anterolaterals which are higher, near

eye-margin. Second row slightly convex, preocellars

farther from each other than from mediolaterals.

Upper postocular bristle developed; 2 bristles near

lower corner of eye. Third antennal joint black, length

0.08 mm; arista long (0.39 mm), distinctly pubescent.

Palpi yellow, well bristled.

Thorax dark brown. Mesopleura bare. Scutellum

with 2 bristles and 2 minute hairs.

Abdomen dark brown. Tergites subequal. Pubescence extremely scarce and short. Hypopygium (Figure 71) dark brown, slightly produced and pale behind; there are 3 small hairs close together near apex;

sides each with 2 conspicuous bristles, above them at

right 2 setulae, lower border at right with a small

notch. Left ventral plate (visible in Figure 71) white,

slender, sinuous, ciliate. Anal segment of medium size,

brown, hairy, on each side with a long hair directed

downward, end-hairs short.

Legs yellow to brownish yellow. Measurements

(holotype): foretibia 0.23 mm, midtibia 0.28 mm,

hind tibia 0.39 mm. Foretarsus slender, longer than

tibia (4:3). Hair-seam of midtibia reaching apical

3rd; end-spur two-thirds of metatarsus. Hind femur

(Figure 73) about 3 times as long as broad, with 7

curved longish hairs on basal half of ventral edge.

Posterodorsal cilia of hind tibia delicate.

Wing (Figure 72) brownish yellow, veins brown.

Length 1.00 mm, width 0.39 mm. Costa 0.44 of wing

length, ratio of segments 15:10:5. Cilia short (0.05

mm), about 6 between tip of 1st vein and apex of
NUMBER 2 3 27

costa. Mediastinalis obsolete. Fork acute. Fourth vein

weakly arcuate, slightly recurved at extreme base and

apex; 5th feebly bent before middle, remainder nearly

straight; 6th slightly sinuate; 7th weak, approximate

to anal margin which is broadly convex. Axillary margin with one long and one short hair. Halteres dark.

FEMALE.—Similar to male. Tergites dark brown,

subequal, 6th trapezoid, with a gland opening at base

medially. Cerci distinct.

MATERIAL EXAMINED.—Holotype male (on slide;

USNM 70258), Dominica, Manets Gutter, 7 Mar.

1965, (W. W. Wirth). Paratypes (USNM, CTB): 3

males on slides, 2 males and 1 female in alcohol, same

locality, date and collector.

DISCUSSION.—M.{M.) diplochaeta, new species,

belongs to group VII. The male is easily recognized

by the conspicuous two bristles on each side of the hypopygium; the female may be separated from its nearest allies by the fourth vein which is slightly but distinctly deflected at tip.

36. Megaselia (Megaselia) horticola Borgmeier

FIGURE 74

Megaselia (Megaselia) horticola Borgmeier, 1962b, pp. 311

[key], 442 [ $ 9 ; Rio de Janeiro, Brazil].

Holotype male (CTB), paratypes 6 males and 36 females, Rio de Janeiro, Brazil (CTB, USNM).

One female (pinned specimen) from Dominica,

Layou R. Mouth, 20 Jan. 1965 (Wirth) agrees perfectly with the types. The wing of a paratype female

is represented in Figure 74; length of this wing 1.17

mm, costa 0.45, ratio of segments 9:5:3. Female of

Dominica: wing length 1.04 mm, costa 0.44, ratio of

segments 8:5:2.5. The costa is slightly thickened on

distal half in both sexes.

DISCUSSION.—M.{M.) horticola is close to M. obscura Brues (1904, St. Vincent, B.W.I.), which also

has the costa thickened on distal half, but in the latter

the costa is longer (0.5) and the fifth and sixth veins

are more distinctly convergent at middle (see Borgmeier, 1962b, fig. 98, p. 443).

37. Megaselia (Megaselia) gracilipalpis, new species

FIGURES 75-77

MALE.—Length 1.00 mm. Head width (paratype)

0.59 mm. Frons nearly twice as broad as long laterally,

width 0.30 mm, yellowish brown, subshining, pubescent, with a distinct median line. Only 2 developed su

are present, occupying about one-fourth of frontal

width; lower su rudimental, reduced to hairs which

are about one-third of the upper in length. Antials in

line with upper su, farther from them than from eyemargin ; anterolaterals a little higher, near eye-margin.

Second row straight, the preocellars farther from

mediolaterals than from each other. Upper postocular

developed; lower postocular strong; jowls and cheeks

with a series of closely set setulae. Third antennal

joint somewhat enlarged, compressed, lenticular,

densely and shortly pubescent, length 0.19 mm, pale

yellow; arista relatively short, 0.40 mm, finely pubescent. Palpi (Figure 77) yellow, slender, fusiform,

length 0.24 mm, with very short setulae and fine hairs,

apical setula slightly differentiated.

Thorax subshining, reddish yellow to brownish,

pleura (in alcohol) pale yellow. Pubescence short,

longer behind. Mesopleura bare. Scutellum with 2

bristles and 2 minute hairs.

Abdomen dark brown, venter yellow; 2nd tergite

yellow on basal half. Tergites 2-6 subequal in length.

Pubescence scarce, short, 2nd tergite with a few moderate bristles laterally; 6th sternite with a group of

short hairs behind. Hypopygium (Figure 76) dark

brown, not large, on both sides with several long

bristles and setulae, right side behind above with a

small notch, rounded below; ventral plate large. Anal

segment of good size, whitish or yellow, end-hairs

hardly longer than the other hairs.

Legs entirely yellow, coxae enclosed, hind femur not

maculated at tip. Foretibia 0.39 mm, midtibia 0.44 mm,

hind tibia 0.63 mm. Hair-seam of midtibia reaching

apical 3rd; cilia distinct; end-spur nearly as long as

metatarsus. Hind femur 3 times as long as broad, on

basal half of ventral edge with about 8 delicate curved

hairs. Posterodorsal cilia of hind tibia weak.

Wing (Figure 75) brownish yellow, veins dark

brown. Length (paratype) 1.69 mm, width 0.72 mm.

Costa 0.5 of wing length, ratio of segments 16:9:4.

Cilia moderately short (0.09 mm), about 7 pairs

between tip of 1st vein and apex of costa. Mediastinalis

incomplete. Fork acute, posterior arm curved. Third

vein without a bristle at base; 4th starting behind

fork, evenly curved, slightly recurved at base and apex;

5th weakly bisinuous; 6th distinctly sinuate. Axillary

margin with 2 hairs. Halteres pale brown.

MATERIAL EXAMINED.—Holotype male (USNM

70259; mounted); Dominica, Clarke Hall 11-20
28

Jan. 1965 (W. W. Wirth). Paratypes (USNM, CTB)

2 males, one in alcohol and the others on 2 slides,

same locality, date and collector.

DISCUSSION.—Megaselia (Megaselia) gracilipalpis,

new species, is a very striking species, easily recognized

by the slender palpi and the structure and chaetotaxy

of the hypopygium; the venation is also characteristic.

38. Megaselia (Megaselia) pallidipennis,

new species

FIGURES 78-80

MALE.—Length 1.3 mm. Head width 0.44 mm.

Frons (Figure 78) subquadrate, slightly broader than

long laterally (6:5), width 0.23 mm, black-brown, with

sparse hairs and a distinct median line. Two supraantennals close together. First row convex, antials

reclinate, nearly on level of su, slightly farther from

anterolaterals than from su. Second row slightly concave, the preocellars more approximate than the

antials. There is a weak supranumerary bristle on

lower corner of frons, the exact length of which could

not be ascertained. Upper postocular bristle distinct;

ocular cilia weak. One moderate lower postocular; one

weak bristle on jowls; cheeks with one setula above.

Third antennal joint globose, yellowish brown; arista

0.44 mm, distinctly pubescent. Palpi yellow, not large,

with moderate bristles.

Thorax black-brown, with 2 dorsocentrals. Mesopleura bare. Scutellum with 2 bristles and 2 minute

hairs.

Abdomen black-brown, venter enclosed. Relative

length of abdominal tergites 12:10:8:8:12. Pubescence

scarce, and short, longer and bristle-like on hind

border of 6th tergite. Second tergite with some bristles

laterally. Hypopygium black-brown, at right with a

single moderate bristle behind. Anal segment white,

rather long (0.19 mm), with longish hairs, the endhairs hardly longer.

Legs brownish yellow, hind legs darker. Foretibia

0.26 mm; foretarsus slender, 0.65 mm, metatarsus

slightly longer than 2+3(9:8). Midtibia 0.35 mm, the

dorsal hair-seam reaching apical fifth; end-spur threefourths of metatarsus. Hind femur (Figure 80) 3 times

as long as broad, length 0.52 mm, with about 12 curved

hairs on basal half of ventral edge. Hind tibia 0.46 mm,

the posterodorsal cilia short.

Wing (Figure 79) grayish white, heavy veins extremely pale, veins 4-6 almost not perceptible, 7th

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

nearly extinguished. Length 1.17 mm, width 0.52 mm.

Costa 0.5 ratio of segments 32:22:7. Cilia short (0.04

mm) about 8 pairs between 1st vein and tip of costa.

Mediastinalis obsolete, 4th vein evenly curved; 5th

weakly arcuate before middle; 6th slightly sinuous.

Axillary margin with 3 hairs. Halteres dark brown.

Holotype (on slide, USNM 70260), Dominica,

Clarke Hall, 21-29 Apr. 1964, light trap (O. S. Flint).

Paratype (on slide, CTB), 1-10 May 1964, same

locality and collector.

DISCUSSION.—M. (M.) pallidipennis, new species,

belongs to group VII. It is recognizable by the very

pale wings and the white anal segment.

39. Megaselia (Megaselia) ferruginosa (Brues)

FIGURE 81

Aphiochaeta ferruginosa Brues, 1912, p. 136 [nomen novum].

Megaselia (Megaselia) ferruginosa (Brues).—Borgmeier,

1935a, p. 451 [9 ; type]; 1962b, p. 309 [key].

Aphiochaeta ferruginea Enderlein, 1912, p. 37 [9 ; Hammonia; not Brunetti 1912]

Lectotype female (CTB), Hammonia, S. Catarina,

Brazil.

MALE (undescribed).—Length 1.2-1.3 mm. Frons

about as broad as long medially, width 0.22 mm,

black-brown, subshining, scarcely pubescent, with a

distinct median line. Two weak su close together, more

aproximate than preocellars, the lower rudimental, reduced to fine hairs. Antials in line with su, a little

farther from them than from laterals which are slightly

higher near eye-margin. Second row slightly convex,

the bristles equidistant. Cheeks with fine hairs. Third

antennal joint globose, ferruginous or orange; arista

distinctly pubescent. Palpi yellow, with short bristles.

Thorax dark castaneous, subshining. Pubescence

short. Mesopleura bare. Scutellum with 2 bristles and

2 hairs.

Abdomen opaque, black-brown; tergites 2-6 subequal, nearly bare. Hypopygium yellowish brown, with

fine hairs; anal segment short, yellow, end-hairs not

differentiated.

Legs yellow, coxae enclosed, hind femur and tibia

sometimes brown. Foretarsus slender, metatarsus as

long as 2-4. Hair-seam of midtibia reaching apical

third. Hind femur slender, 4 times as long as broad;

ventrobasal hairs short, not reaching middle. Posterodorsal cilia of hind tibia short and delicate.

Wing (Figure 81; male from Nova Teutonia) grayish yellow, veins brown. Length 1.19 mm, width 0.54
NUMBER 2 3 29

mm. Costa 0.5 of wing length, ratio of segments 2:12:5

(in male from Dominica 11:6:2). Cilia short (0.03

mm), about 9 pairs between rt and r3. Third vein

without a bristle at base. Fork not large, posterior arm

arcuate. Fourth vein evenly concave, starting at fork;

5th and 6th sinuous. Axillary margin with 2 hairs.

Halteres dark brown.

FEMALE.—Length 1.5-1.8 mm. Similar to male.

Tergites 2-6 subequal, scarcely pubescent, 2nd with 2

bristly hairs laterally; 6th tergite trapezoid, in cne

specimen semicircular. Wing length 1.56 mm to 1.82

mm. Costa 0.53 (type) or 0.54 (Nova Teutonia), ratio

of segments in type 15:10:4 (in specimen from Nova

Teutonia 15:8:3); in 2 specimens from Nova Teutonia

fork large, costa 0.54, ratio 14:8:5, and 4th starting

well beyond fork.

MATERIAL EXAMINED.—Besides the type from Brazil,

I have seen 2 females from Petropolis, Rio de Janeiro,

Brazil; 5 females and 5 males from Nova Teutonia,

Brazil; 2 males and 3 females from Higuito, San Mateo,

Costa Rica (P. Schild); 2 males from Dominica: Pont

Casse, 12 Feb. 1965, and Clarke Hall, 11-20 Jan. 1965

(Wirth).

DISCUSSION.—M. (M.) ferruginosa belongs to

group VII. It is easily recognized by the reddish color

of the antennae, the weak su, the pale hypopygium and

by the venation.

VARIATION.—The two females with large fork are

rather puzzling, but I think they are individual

variants.

40. Megaselia (Megasetia) bursella, new species

FIGURES 82-83

MALE.—Length 1.60 mm. Head width 0.52 mm.

Frons brownish black, broader than long laterally

(4:3), width 0.26 mm, sparsely pubescent, with a distinct median line. Four su, the lower weaker and about

three-fourths of the upper in length, the latter are more

approximate than the preocellars and are about onefourth of frontal width apart. Antials slightly below

level of upper su, farther from them than from eyemargin. Upper and lower postocular bristles strong;

one bristle on jowls; cheeks with 3 setulae. Third antennal joint brown, slightly enlarged, length 0.13 mm;

arista 0.39 mm, distinctly pubescent. Palpi brown,

slender, the bristles short or of moderate length.

Thorax reddish brown. Pubescence short, behind

longer and bristle-like. Mesopleura bare. Scutellum

with 2 bristles and 2 minute hairs.

Abdomen dark brown, venter yellow. Tergites subequal. Pubescence short, hind margin of 6th tergite

with bristly hairs. Hypopygium (Figure 83) small,

dark brown, sides connected beneath anal segment,

connecting bridge at middle angulate (in profile) and

bearing at tip 2 curved bristles directed downwards;

each lateral lobe of the epandrium with several weak

bristles or bristly hairs. Sixth sternite with a deep median pocket flanked on each side by an angulate lobe

which bears numerous short marginal hairs. Sternites

4-5 with a transverse apical row of hairs. Anal segment

of median length, pale yellow, end-hairs differentiated.

Legs yellow, hind femur maculated at extreme tip.

Foretibia 0.33 mm. Foretarsus slender, metatarsus

nearly as long as joints 2—4 combined. Midtibia 0.39

mm, hair-seam ending subapically. Hind femur 3 times

as long as broad, on basal half of ventral edge with 15

short closely set curved hairs. Hind tibia 0.52 mm, with

9 fine posterodorsal setulae.

Wing (Figure 82) grayish yellow, veins dark brown.

Length 1.35 mm, width 0.62 mm. Costa 0.51 of wing

length, ratio of segments 13:9:3. Cilia short (0.065

mm), close, there are about 10 pairs between Ti and

tip of costa. Mediastinalis present. Fork acute. Third

vein without a bristle at base; 4th evenly arcuate, 5th

very weakly sinuous, 6th sinuate. Axillary margin with

2 hairs. Halteres dark brown.

Holotype, on slide (USNM 70261), Dominica,

Clarke Hall, 1-10 Feb. 1965, light trap (Wirth).

DISCUSSION.—M. bursata Borgmeier (1966, Arizona) has a similar pocket on 6th sternite, but has four

scutellars and the hypopygium is quite different. This

new species is most closely related to M. (M.) burselloides, new species, but differing by the structure of

the sternal pocket and the hypopygium.

REMARK.—The drawing of the hypopygium was

made after removing the internal copulatory organs.

41. Megaselia (Megaselia) burselloides, new species

FIGURES 84-85

MALE.—Length 1.65 mm. Head width 0.52 mm.

Frons black or black-brown, broader than long laterally (4:3), width 0.54 mm, sparsely pubescent, median

line distinct. Lower su weak, about one-half in length

of upper which are one-fourth of frontal width apart.

Antials a little below level of upper su, twice as far

from them as from eye-margin, anterolaterals only

slightly higher, near eye-margin. Second row nearly
30

straight, the bristles equidistant. Two divergent

bristles near lower corner of eye; cheeks with 2 setulae.

Third antennal joint dark brown, globose; arista 0.39

mm, distinctly pubescent. Palpi slender, brownish yellew, with bristles of medium length.

Thorax black-brown. Pubescence short, longer

behind, Mesopleura bare. Scutellum with 2 bristles and

2 minute hairs.

Abdomen dark brown, venter yellow. Tergites subequal. Pubescence scarce and short, longer on hind

corners, bristle-like on hind margin of 6th. Hypopygium (Figure 85) dark brown, sides of epandrium connected beneath anal segment, connecting bridge produced into a rounded point, each lateral Jobe of epandrium with several bristly hairs. Left ventral plate

large, broadly rounded; right ventral plate triangular.

Sixth sternite with a small pocket or pouch, flanked

on each side by a short rounded process which bears

2 setulae; below the pocket are 2 or 3 hairs on each

side. Anal segment whitish yellow, of medium size,

end-hairs differentiated.

Legs yellow, hind femur maculated at extreme tip.

Foretibia 0.28 mm; foretarsus slender, metatarsus

longer than 2 + 3. Midtibia 0.41 mm; seam nearly

complete. Hind femur on basal half of ventral edge

with 9 longish curved hairs. Hind tibia 0.57, with 8

fine posterodorsal setulae.

Wing (Figure 84) grayish yellow, veins dark brown.

Length 1.40 mm, width 0.62 mm. Costa 0.53 of wing

length, ratio of segments 25:18:6. Cilia moderately

short (0.091 mm), there are 8 marginals between tip

of 1st vein and apex of costa. Mediastinalis present.

Fork acute. Third vein without a bristle at base; 4th

starting with a faint hook, concave; 5th slightly bisinuate, 6th sinuous. Axillary margin with 2 hairs. Halteres

dark brown.

Holotype, on slide (USNM 70262), Dominica,

Clarke Hall, 10 Jan. 1965, Malaise trap (Wirth).

DISCUSSION.—M. (M.) burselloid.es, new species,

is very close to M. (M.) bursella, new species, but the

sternal pocket is different and the fifth vein is more

curved on distal half.

REMARK.—The drawing of the hypopygium was

made after removing the internal copulatory organs.

42. Megaselia (Megaselia) fortirostris, new species

FIGURES 86-88

FEMALE.—Length 1.30-1.40 mm. Head width 0.40

mm. Frons brown, broader than long laterally (4:3),

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

width (holotype) 0.20 mm, scarcely pubescent, with a

fine median line. Supraantennals unequal, the lower

rudimental, very thin and more or less one-half in

length of the upper which are one-fourth of frontal

width apart, the lower are slightly more approximate.

Antials about in line with upper su, close to anterolaterals which are a little higher. Second row convex,

the bristles equidistant. Two divergent bristles near

lower comer of eye; cheeks with 3 hairs. Third antennal joint dark brown, of normal size, globose, width

0.08 mm, arista 0.35 mm, distinctly pubescent. Palpi

yellowish brown, slender, with 6 long bristles on apical

half. Epistoma not produced. Proboscis (Figure 88)

dark brown, length 0.26 mm, labrum voluminous,

pear-shaped, length 0.18 mm, width 0.15 mm. Labellae

narrow.

Thorax reddish yellow or yellowish brown. Pubescence short, longer behind. Mesopleura bare. Scutellum with 2 bristles and 2 minute hairs.

Abdomen dark brown, venter grayish yellow. Tergites subequal, gradually diminishing a little in length,

6th with a gland-opening at base. Pubescence on

tergites scarce and short. Venter distinctly hairy; 5th

segment (Figure 87) laterally above with several stiff

bristly hairs (length 0.065 mm); 6th tergite trapezoid,

the sides straight. Terminal segments fleshy, cerci

small.

Legs pale yellow, hind femur blackened at tip. Foretibia 0.26 mm (in paratype 0.28 mm); tarsus slender,

0.33 mm (0.36 mm) ; metatarsus a little longer than

2 + 3. Midtibia 0.28 mm (0.31 mm), hair-seam reaching apical third, cilia distinct, spur five-sixths of metatarsus. Hind femur 3 times as long as broad, on basal

half of ventral edge with 6 longish curved hairs. Hind

tibia 0.41 mm (0.49 mm), with about 10 short fine

posterodorsal setulae.

Wing (Figure 86) grayish yellow, veins pale brown,

4-7 evanescent at apex. Length 1.10 mm (in paratype

1.22 mm), width 0.52 mm (0.59 mm). Costa 0.53

of wing length, ratio of segments 24:16:5. Cilia short

(0.078 mm), 7 pairs between rx and apex of costa.

Mediastinalis distinct. Fork small, acute. Third vein

without a bristle at base; 4th concave, especially on apical half; 5th sinuous, 6th weakly bisinuate, 7th weak.

Axillary margin with 2 hairs. Halteres dark brown.

Holotype (on slide, USNM 70263) and 1 paratype

(on 2 slides, CTB), Dominica, Manets Gutter, 7 Mar.

1965 (Wirth).

DISCUSSION.—Easily recognized by the four stiff

bristly hairs laterally on fifth segment. Extremely close
NUMBER 2 3 31

to M. (M.) tumidirostris, new species, which lacks the

stiff hairs on fifth segment. M. crassirostris is a black

species, with the epistoma produced.

43. Megaselia (Megaselia) tumidirostris,

new species

FIGURES 89-91

Among material received from the United States

National Museum I found one female on slide from

Dominica, which belongs to a new species that is extremely close to M.(M.)fortirostris, new species, described above. A detailed description would be for the

greatest part a repetition of that of M. (M.)fortirostris,

new species. The main differences are: (1) The

structure of the labrum (Figure 90) is somewhat

different, being less voluminous. (2) The sixth tergite

(Figure 91) has the lateral borders slightly curved

instead of straight. (3) The fifth segment lacks the

stiff hairs present on the pleural area in M. fortirostris.

(4) The costa is shorter (index 0.5); length of wing

(Figure 89) 1.17 mm; ratio of costal segments 20:13:6;

therefore, the second segment is relatively shorter; the

costal cilia are shorter and more closely set (there are

nine pairs between tip of first vein and apex of costa);

the fourth vein is less arcuate on distal half, and the

fifth is more distinctly sinuous.

MATERIAL EXAMINED.—Holotype on slide (USNM

70264), Dominica, Clarke Hall, June 1964, light trap

(O. S. Flint).

44. Megaselia (Megaselia) miristigma Borgmeier

FIGURE 97

Megaselia (Megaselia) miristigma Borgmeier, 1962b, pp. 308

[key], 390, fig. 69 [abdomen, 2 ; Nova Teutonia;

Petr6polis; San Jose, Costa Rica].

Holotype female (CTB), Nova Teutonia, Brazil; 4

paratypes (GTB, USNM), Nova Teutonia, Petropolis,

Brazil; S. Jose, Costa Rica).

MALE (undescribed).—Length 1.4-1.8 mm. Frons

as broad as long laterally, slightly longer medially, yellow or brownish, densely pubescent, opaque, with a

distinct median line. Su weak, the lower thin and a

little more than one-half of upper, the latter one-fourth

of frontal width apart. Antials not much inclined toward median line, slightly above level of upper su, a

little farther from laterals than from su; laterals higher

on each side in oblique line with antials and upper su.

336-7 0» O—69 5

Second row straight, the bristles equidistant. One

strong lower postocular; jowls with 3 weak bristles;

cheeks with 3 short hairs. Third antennal joint small,

globose, brown; arista one-third longer than frontal,

width, shortly pubescent. Palpi clavate, yellow, with 6

bristles. Proboscis brown, labrum small, labellae large,

semicircular, without pseudotracheae.

Thorax yellow. Mesopleura bare. Scutellum with 2

bristles and 2 minute hairs.

Abdomen opaque, venter yellow. Tergites 2-6 subequal, brown, sometimes paler medially and on basal

half of 2nd. Pubescence scarce, short, not conspicuously

longer on hind margin of 6th. Hypopygium of medium

size, brown, with fine hairs; anal segment of medium

length, yellow, end-hairs not differentiated.

Legs yellow, hind femur blackened at tip. Foretarsus

slender, metatarsus slightly longer than 2-4. Hairseam on midtibia complete, setulae biseriate, the anterodorsal weak. Hind femur nearly 3 times as long as

broad, the ventral hairs on basal half short. Hind tibia

with 6 strong setulae, sometimes 2 additional weaker

ones are present.

Wing narrow (Figure 97), yellowish, veins pale

brown. Length 1.4-1.8 mm. Costa 0.53, ratio of segments 14:12:5. Cilia very short and close. Mediastinalis wanting. Fork long, narrow, acute. Third vein

straight, with a minute hair at base; 4th-7th but little

curved, 4th starting in front of fork, sometimes obsolete

basally, slightly deflected at extreme tip. Anal margin

broadly convex; axillary margin with 2 short hairs.

Halteres dark brown.

FEMALE.—See original description. Tergites 3-5 with

a large stigma laterally (see Borgmeier, 1962, fig. 69).

MATERIAL EXAMINED.—The types (see above); 1

female in alcohol, Dominica, Fond Figues River

9-12 Mar. 1965 (Wirth); 1 female and 6 males, all on

slides, Puerto Rico, El Verde Exp. St., Barrio Rio

Grande, 1963-1966, G. E. Drewery (No. 26).

DISCUSSION.—The female is easily recognized by the

large stigmata; the male by venational characters.

45. Megaselia (Megaselia) hamaticauda,

new species

FIGURES 92, 93

MALE.—Length 1.25-1.37 mm. Frons yellow, subopaque, about as broad as long laterally (10:9), width

0.26 mm, a little longer medially (0.3 mm), with

scattered hairs and a fine median furrow. There are
32 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

4 supraantennals, the upper are shorter than the antials and are about one-fourth of frontal width apart;

they are more approximate than the preocellars; the

lower supraantennals are very weak and about onehalf of the upper in length. First row distinctly convex,

the antials are one-half of frontal width apart. Second

row slightly concave anteriorly, the bristles equidistant.

Eyes large. One strong lower postocular bristle; cheeks

narrow, with about 6 hairs or weak bristles. Third antennal joint small, orange; arista about one-third

longer than frontal width, shortly pubescent. Palpi yellow, about one-half as broad as long, length 0.13 mm,

with moderately long bristles. Proboscis orange, with

very broad labellae.

Thorax yellow, subshining; pleura pale, mesopleura

bare. Scutellum with 2 bristles and 2 minute hairs.

Postscutellum brown.

Abdomen opaque, venter yellow. Tergites 2-5 subequal in length, 6th elongated; 1st brownish, 2 and 5

yellow, 3, 4, and 6 with a black or brown spot laterally, not reaching middle; in the paratype the tergites

are a little darker and have distinct yellow seams on

hind margins. Pubescence scarce, short, almost limited

to hind margins. Hypopygium dark brown, subshining,

with a few short hairs. Anal tube yellow, of medium

length, as long as dorsum of hypopygium, end-hairs

moderately strong (wanting in paratype).

Legs yellow, hind femur darkened at tip. Foretarsus

slender. Dorsal hair-seam of midtibia complete, cilia

biseriate on upper half. Hind femur broad, ventral

fringe on basal half wanting (the hairs are not longer

than the pubescence). Posterodorsal setulae of hind

tibia about 7, strong and about as long as width of

tibia, except the basal ones.

Wing (Figure 92) grayish yellow, veins dirty yellow.

Length (paratype) 1.37 mm, width 0.5 mm. Costa

0.56 of wing length, ratio of segments 13:8:7. Cilia

short. Humeral cross-vein indistinct, mediastinalis

wanting. Base of 3rd vein with a short hair. Fork long.

Fourth vein starting behind fork, obsolete and slightly

curved basally, remainder nearly straight except at

apex which is somewhat deflected; 5th but little

curved; 6th slightly sinuous. Anal margin broadly convex. Axillary margin with 2 hairs. Halteres dark brown.

FEMALE.—Length 2.25 mm. Similar to male. Third

antennal joint small, orange. Proboscis short, labellae

normal. One lower postocular bristle; jowls with one

bristle; cheeks with 8 hairs. Thorax yellow, postscutellum brown. Tergites 1-5 subequal, 6th elongated; 1,

3, and 4 dark brown, 2 and 5 yellow, somewhat brownish at middle; 6th nearly black, with 2 faint yellow

longitudinal stripes; 6th with a broad yellow seam,

seams on 4th and 5th narrow. Pubescence nearly absent. Seventh segment black, tergite small; intersegmental membrane black, surrounding the base of a

peculiar chitinous structure which comprises segments

8-10 (Figure 93); this structure is black-brown, glossy,

striolate in side view and distinctly concave above, terminating behind on each side in an obtuse erect hook

which nearly touch each other; at base above there

are several fine hairs, one of which is longer. Hind

femur as in male. Wing length 2 mm. Costa 0.6 of

wing length.

MATERIAL EXAMINED.—Holotype male (USNM

70265) and paratype female (USNM), Dominica,

Fond Figues River, 3 Feb. 1965, rain forest (W. W.

Wirth); one paratype male (CTB), same locality,

9-12 Mar. 1965 (W. W. Wirth); one paratype male,

1.5 m east Pont Casse, 11 Apr. 1966 (R. J. Gagn6);

one paratype male (in alcohol), Malaise trap, 12-14

Oct. 1964 (P. J. Spangler).

DISCUSSION.—M. (M.) hamaticauda, new species,

belongs to group VII. The venation is very similar to

M. orthoneura Borgmeier (1961; Brazil), but in the

latter the female terminal segments are fleshy and the

male has a small hypopygium and a distinct ventral

fringe on hind femur; the color of the tergites is also

different.

Three species are known from the Philippines, in

which the female is provided with curious chitinous

structures on the seventh and eighth abdominal segments: M. (M.) appendiculata Brues, M. (M.) bihamulata Brues and M. (M.) carinata Borgmeier (see

Borgmeier, 1967b, pp. 151—155).

46. Megaselia (Megaselia) paula, new species

FIGURES 95,96

MALE.—Length 0.90-1.00 mm. Head width (paratype, on slide) 0.31 mm. Frons brownish, grayish pruinose, opaque, broader than long laterally (3:2),

width 0.19 mm, pubescent, with a fine median line.

Lower su rudimental, hair-like; upper not quite onefourth of frontal width apart (4:14). Antials on level

of upper su, slightly farther from them than from eye

and from anterolaterals, the latter a little higher. Second row feebly convex, the bristles equidistant. Upper

and lower postocular long. No bristle on jowls; cheeks
NUMBER 23 33

with a series of 5 hairs. Third antennal joint yellow

or brownish yellow, globose, width 0.065 mm; arista

0.20 mm, distinctly pubescent. Palpi yellow, of normal size, slightly angulate on upper border near base,

with 5 long bristles on apical half. Proboscis small.

Thorax subshining, yellow to brownish yellow. Pubescence short, not noticeably longer behind. Mesopleura bare. Scutellum with 2 bristles and 2 very minute hairs.

Abdomen opaque, venter yellow, tergites dark

brown, subequal in length, 6th yellow on anterior half.

Pubescence scarce, mostly short, some longer hairs on

hind margin on 6th tergite. Hypopygium of medium

size, dark brown, dull, with longish hairs. Anal segment of medium length, yellow, with long hairs, endhairs not differentiated.

Legs entirely yellow. Foretibia (in paratype) 0.19

mm; foretarsus 0.28 mm, metatarsus about as long as

2 + 3. Midtibia 0.26 mm; seam reaching apical third;

cilia fine; end-spur three-fourths of metatarsus; last

joint of mid tarsus (Figure 96) a little thickened, about

twice as long as preceding. Hind femur 3 times as

long as broad, on ventral edge near base with about 4

longish curved hairs. Hind tibia 0.32 mm with 10 very

fine cilia; last joint of hind tarsus slightly longer than

preceding.

Wing (Figure 95) slightly grayish yellow, veins

brown (yellow in paratype on slide). Length 1.04 mm

in holotype, 0.85 mm in mounted paratype, width of

latter 0.39 mm. Costa 0.5 of wing length, ratio of segments 14:10:5. Cilia short (0.05 mm), widely placed,

there are 6 pairs between r^ and apex of costa. Mediastinalis weak. Fork large, anterior arm steep, posterior

arm curved. Third vein without a bristle at base; 4th

starting behind fork, evenly arcuate; 5th and 6th

sinuous. Anal margin broadly convex. Axillary margin

with 2 hairs. Halteres dark brown.

FEMALE.—Length 1.3 mm. Similar to male. Tergites 2-6 subequal, gradually narrowed, scarcely pubescent, hairs on hind margin of 6th not noticeably

longer; 8th segment dark. Tarsi of normal length.

Wing length 1.17 mm.

MATERIAL EXAMINED.—Holotype male (pinned,

USNM 70266), Dominica, Fond Figues River, 20

Jan. 1965 (W. W. Wirth); allotype female (on slide,

USNM), and 2 males paratypes: one pinned

(USNM), another on slide (CTB), Fond Figues

River, 3Feb. 1965 (Wirth).

DISCUSSION.—M. (M.) paula, new species, belongs

to the group of species whose males have the last midtarsal joint twice as long as the preceding: M. lutea

Meigen (Europe, U.S.A.), M. scutellaris Wood (Europe), M. {M.) serotina Borgmeier (Brazil), M. luteoides Schmitz (Formosa), M. tetrachaeta Beyer

(New Guinea), and M. postorta Borgmeier (Australia). M. paula, new species, differs from M. serotina

by smaller size, the ratio of costal segments (in M.

serotina 1 is longer than 2 + 3), and its less curved

fifth vein (see Figure 94.).

47. Megaselia (Megaselia) flavopleura (Malloch)

FIGURES 98, 99

Aphiochaeta flavopleura Malloch, 1914, p. 31 [9 ; Costa

Rica].

Megaselia (Megaselia) flavopleura (Malloch).—Borgmeier,

1962b, pp. 309, 429, fig. 90 [3 ; paratype].

Holotype female (ANSP), one paratype male (not

female, CTB), from Costa Rica.

MALE.—Length 1.20-1.40 mm. Frons somewhat

broader than long medially (5:4), brown, rather shining, with sparse pubescence and a distinct median line.

Lower su weak, about one-half of upper which are

one-fourth of frontal width apart (the lower oneeighth). Antials in line with upper su, near eyemargin, vertically below anterolaterals. Second row

convex, the bristles equidistant. Two long divergent

bristles near lower corner of eye; cheeks with a few

hairs. Third antennal joint brown, of normal size;

arista one-third longer than width of frons. Palpi narrow, yellow, with long bristles.

Thorax yellow, pleura pale. Mesopleura bare. Scutellum with 2 bristles and 2 minute hairs.

Abdomen: venter yellow; tergites brown, 2-6 subequal in length, sparsely hairy, hairs on hind border

of 6th bristle-like. Venter sparsely hairy. Hypopygium

(Figure 98) of medium size, pale brown, with a few

short hairs laterally. Anal segment short, pale yellow,

with short hairs and one longer hair on each side directed downwards, as long as apical hairs.

Legs yellow. Foretarsus slender, metatarsus longer

than 2 + 3. Hind femur on basal half with several

longish delicate curved hairs. Hind tibia with about

13 fine short cilia.

Wing (see Borgmeier, 1962b p. 399, fig. 20, drawn

from paratype) slightly yellowish, veins dirty yellow.

Length (paratype) 1.4 mm. Costa 0.56-0.59 length
34

of wing; segments 23:23:9. Cilia moderately short

(0.1 mm), 8 or 9 pairs between 1st vein and apex

of costa. Third vein without a bristle at base. Fork

large, anterior arm rather steep. Fourth starting behind fork, 5 and 6 sinuous. Axillary margin with 2

hairs. Halteres dark brown.

FEMALE.—Length 1.3-1.5 mm. Similar to male.

Tergites pale brown, sparsely hairy. Tergites 5 and 6

(Figure 99) moderately convex laterally, hind corners conspicuous; 6th transverse. Costa up to 0.6 length

of wing, 2nd segment a little longer than first

(10:11:5).

MATERIAL EXAMINED.—Paratype male, Costa Rica

(holotype female seen in Philadelphia in 1961); 10

males and 10 females (one pinned, one on slide, the

rest in alcohol) from Dominica, Manets Gutter,

15 Mar. 1965 (Wirth), and Fond Figues River,

27 Feb. 1965(Wirth).

DISCUSSION.—Easily separated from M. (M.) alajuelensis by the position of the antials, the long costa,

the large fork, the fourth vein starting behind fork,

and the structure of tergites 5 and 6.

48. Megaselia (Megaselia) crinellifemur,

new species

FIGURES 100-104

MALE.—Length 1.40-1.60 mm. Head width about

0.50 mm. Frons slightly broader than long laterally

(5:4), width 0.26 mm, yellow or brownish yellow,

infuscated above, subopaque, pubescent, with a distinct

median line. Su unequal, the lower thin and about

one-half in length of the upper which are one-fourth

of frontal width apart. Antials nearly in line with upper su (slightly lower), about midway between su and

anterolaterals, the latter not much higher, near eyemargin. Second row slightly convex, the preocellars

slightly farther from 12 than from each other, but not

so much approximate as upper su. Upper postoculars

well developed. Two divergent strong bristles near

lower eye-corner; cheeks with 3 weaker bristles. Third

antennal joint slightly enlarged, globose, yellow, infuscated at tip, sometimes brownish. Arista short (0.32

mm), distinctly pubescent. Palpi yellow, of normal size,

the bristles moderately long. Proboscis small, yellow.

Thorax subshining, yellow or brownish yellow.

Pubescence short. Mesopleura bare. Scutellum with 2

bristles and 2 hairs.

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Abdomen: venter yellow; tergites black-brown, 5th

yellow, also 2nd along anterior border yellow and

sometimes a fine median yellow stripe on 2-4; 2nd and

6th slightly elongate, 3-5 subequal. Pubescence extremely scarce, short, 2nd with 3 and 4 moderate bristles laterally, and 6th with bristly hairs on hind margin.

Venter with conspicuous apical rows of bristly hairs

on segments 3-6. Hypopygium (Figure 103) dark

brown, with short hairs laterally, one of which is longer; left ventral plate (Figure 104) broadly rounded,

whitish yellow; right process triangular. Anal segment

long (0.19 mm), sheat-shaped, whitish yellow, slightly

infuscated on basal half, with fine hairs, end-hairs differentiated.

Legs yellow, hind femur blackened at tip. Foretibia

0.30 mm, with a dorsal row of fine cilia; foretarsus slender, 0.36 mm, metatarsus a little longer than 2 + 3.

Midtibia 0.41 mm, seam reaching apical fourth, cilia

fine, spur three-fourths of metatarsus. Hind femur

(Figure 100) 3 times as long as broad, on basal half

of ventral edge with a fringe of about 12-14 short

stiff hairs, the basal 9-10 are closely set and straight.

Hind tibia 0.52 mm, with about 9 short setulae, the

basal ones very weak.

Wing (Figure 101) grayish yellow, veins dark brown.

Length 1.40 mm, width 0.59 mm. Costa 0.53 length

of wing, ratio of segments 24:21:6. Cilia short, 0.065

mm, there are 9 pairs between tip of 1st vein and apex

of costa. Fork narrow, acute. Mediastinalis incomplete;

3rd vein without a bristle at base; 4th evenly arcuate,

starting at fork with a very faint hook; 5th but little

curved before middle; 6th feebly sinuous. Axillary

margin with 3 hairs. Halteres brown.

FEMALE.—Similar to male. Length 1.80-2.00 mm.

Tergites (Figure 102) black, 6th with a median yellow

spot anteriorly; venter yellow, lateral areas darkbanded; the tergites are subequal in length and gradually narrowed; 6th subquadrate, slightly broader than

long. Pubescence very scarce and short; 2nd with some

moderate bristles laterally; venter with a transverse

apical row of bristly hairs on segments 3-6; 7th tergite

long and narrow. Fringe on hind femur as in male.

MATERIAL EXAMINED.—Holotype male (pinned;

USNM 70267), Dominica, Fond Figues R., 30 Jan.

1965 (Wirth). Paratypes (USNM CTB), pinned: 4

males 3 females, from the following localities: S. Chiltern Est. 2 Feb. 1965 (Wirth); Clarke Hall, 11-20

Feb. 1965 (Wirth), June 1966 (G. Steyskal); in al-
NUMBER 2 3 35

cohol: 18 males 4 females, Clark Hall, 1-10 Feb. 1965

(Wirth).

DISCUSSION.—M. (M.) crinellifemur, new species,

is very close to M. subflava Malloch (Puerto Rico), but

differs by the close fringe on hind femur and by venational characters. Dr. Wirth was kind enough to compare one paratype male with the holotype male of M.

subflava (USNM 14857). The latter is represented

by a good series from Puerto Rico and will be treated

in a forthcoming paper. Also close to M.(M.) subaristalis, new species, which is smaller, the ventral plate

is triangular, the hind femur is not maculated at tip,

and the 1st vein is more straight near apex.

49. Megaselia (Megaselia) subaristalis, new species

FIGURES 105-109

MALE.—Length 1.2-1.3 mm. Head width 0.45 mm.

Frons subquadrate, brown, median furrow distinct.

Lower supraantennals very weak and about one-half

of upper, the latter more approximate than the preocellars. Antials inclined toward median line, about

midway between upper supraantennals and anterolaterals, which are slightly higher placed. Third antennal

joint (Figure 105) brownish, somewhat enlarged, oval,

about one-half eye-length; arista dorsal, hardly twice

as long as 3rd antennal joint, pubescent. Palpi short,

yellow, the bristles moderately long. Proboscis short.

Thorax yellowish brown (in alcohol reddish yellow),

pleura pale. Mesopleura bare. Scutellum with 2

bristles.

Abdomen: ventor yellow. Tergites brown, subequal

in length. Pubescence scarce, short, longer on hind

margin of 6th tergite. Venter with a transverse row

of bristle-like hairs on hind margin of 6th segment.

Hypopygium (Figure 108) dark brown, higher than

long, on both sides with hairs and one weak bristle

on lower border. Both sides of dorsal part connected

below anal tube. Right ventral plate (Figure 107)

small, whitish, triangular; left ventral plate large,

rounded triangular (Figure 107), generally curved

and spoon-like in side view (Figure 108). Anal tube

long, yellow, with short hairs; end-hairs strong.

Legs yellow, hind femur (Figure 109) infuscated

at tip, moderately broad, with a distinct fringe of about

8 longish hairs on ventral edge of basal half. Foretarsus

slender. Hind tibial setulae delicate but distinct, weaker

on upper half.

Wing (Figure 106) brownish yellow, veins brown.

Length (paratype) 1.13 mm, width 0.5 mm. Costa 0.55

of wing length, ratio of segments 10-9-2. Cilia short

(0.06-0.08 mm). Fourth vein evenly concave, faintly

deflected at extreme tip; 5th and 6th slightly sinuous;

7th weak. Axillary margin with 3 hairs. Halteres dark

brown.

FEMALE.—Length 1.5 mm. Similar to male. Third

antennal joint not enlarged; arista about two and onehalf length of 3rd antennal joint. Tergites dark brown,

2nd a little elongated, 3-5 subsequal, 6th small, with

a whitish median notch anteriorly. Pubescence scarce,

2nd tergite with some hairs laterally. Terminal segments fleshy, 7th with a narrow dorsal plate. Fringe on

hind femur as in male.

MATERIAL EXAMINED.—Holotype male (USNM

70268), pinned (from alcohol), Dominica, Point Lolo,

0.5 miles west, 25 Jan. 1965, at light (W. W. Wirth).

Paratypes: 1 female (USNM),pinned (fromalcohol),

and 13 males (in alcohol), same locality, date and

collector.

DISCUSSION.—M. (M.) subaristalis, new species,

belongs to group VII. In my key to the neotropical

species (1962b) it runs to M. tumidicornis, but the

latter has a shorter costa (0.45). The species may be

recognized by the enlarged 3rd antennal joint in male,

the rather short arista, and the structure of the hypopygium, especially the left ventral plate. Close to M.

crinellifemur (see p. 35).

50. Megaselia (Megaselia) totaftava, new species

FIGURES 110-111

FEMALE.—Length 1.3-1.5 mm, fully extended 1.9

mm. Head width 0.44 mm. Frons yellow, quadratic,

width 0.20 mm, pubescent, with a distinct median

line. Su unequal, the lower thin and less than one-half

in length, the upper about one-fourth of frontal width

apart, the lower one-eighth. Antials nearly in line with

upper su, twice as far from them than from eye-margin, close to anterolaterals which are slightly higher

near eye-margin. Second row slightly convex, the bristles equidistant. One upper and one lower postocular

bristle; one bristle on jowls; cheeks with 2 setulae.

Third antennal joint yellowish brown, globose, of normal size, width 0.09 mm; arista 0.39 mm, finely pubescent. Palpi whitish, with moderate bristles.

Thorax reddish yellow, pleura pale. Pubescence

short, longer behind. Mesopleura bare. Scutellum with

2 bristles and 2 minute hairs.
36

Abdomen ochreous yellow, venter pale yellow. Tergites 2-6 subequal, gradually narrowed, 5th (Figure

110) on hind margin with a small excavation at

middle, 6th tranverse, sides weakly convex, with a

gland opening at base. Pubescence scarce and short,

some short bristly hairs on 2nd tergite laterally.

Terminal segments tubiform, whitish; 7th with a

trapezoid tergal plate, sternite a linear strip dilated

apically; 8th segment behind with 2 very small tergal

plates close together. Cerci small.

Legs entirely yellow, hind femur not darkened at

tip. Foretibia 0.28 mm; tarsus slender, 0.37 mm.

Midtibia 0.39 mm, hair-seam reaching apical fourth;

9 distinct posterodorsal cilia; end-spur three-fourths of

metatarsus. Hind femur nearly 3 times as long as

broad, ventrobasally with 5 longish curved hairs. Hind

tibia 0.50 mm, with 12 fine posterodorsal cilia.

Wing (Figure 111) grayish yellow, veins pale

brown. Length 1.24 mm, width 0.52 mm. Costa 0.52

of wing length, ratio of segments 21:17:6. Cilia short

(0.05 mm) and close, there are about 10 pairs between tip of 1st vein and apex of costa. Fork acute.

Mediastinalis obsolete. Third vein without a bristle at

base; 4th concave on basal fourth, remainder nearly

straight; 5th and 6th slightly sinuous. Anal margin

convex, axillary margin with 2 hairs. Halteres

brownish.

MATERIAL EXAMINED.—Holotype female, on slide

(USNM 70269), S. Chiltern Est. 20 Feb. 1965

(Wirth); paratypes in alcohol (USNM, CTB): 65

females, S. Chiltern Est., Clarke Hall and Pont Casse,

I-III 1965 (Wirth); 1 pinned female, Clarke Hall,

11-20 Jan. 1965.

DISCUSSION.—M. (M.) totaflava, new species,

comes close to M.(M.) alajuelensis but differs by the

structure of tergites 5-7, the shorter costal cilia and

other characters.

51. Megaselia (Megaselia) alajuelensis (Malloch)

FIGURES 112-114

Aphiochaeta alajuelensis Malloch, 1914, p. 32 [<$ 9 j Costa

Rica].

Megaselia (Megaselia) alajuelensis (Malloch).—Borgmeier,

1962b, pp. 309,410, fig. 91 [Costa Rica; Brazil].

Holotype male and paratype female, Alajuela, Costa

Rica (ANSP); 1 paratype male (CTB).

MALE.—Length 1.00-1.80 mm. Frons subquadrate,

as broad as long medially, somewhat shorter laterally,

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

yellow or brown, pubescent, with a distinct median

line. Lower su weak, about one-half in length of the

upper which are one-fourth of frontal width apart.

Antials very close to anterolaterals, inside of them and

slightly lower; 2nd row slightly convex, the bristles

equidistant. Third antennal joint brown, globose, of

normal size; arista one-third longer than width of

frons. Palpi yellow, with long bristles.

Thorax yellow. Mesopleura bare. Scutellum with

2 bristles and 2 hairs.

Abdomen: venter yellow. Tergites brown, 6th sometimes yellow. Pubescence short, hairs on hind-margin

of 6th bristle-like. Venter with transverse apical rows

of conspicuous bristly hairs on segments 3-6. Hypopygium (Figure 113) pale brown, small, hairy, hairs

short or of moderate length. Anal segment short, yellow, with short hairs, end-hairs not differentiated.

Legs entirely yellow, hind femur not darkened at

tip. Foretarsus slender. Hind femur ventrobasally with

about 7 longish curved hairs. Hind tibia with 12 fine

setulae, those below middle somewhat stronger.

Wing (see Borgmeier, 1962b, fig. 91), grayish, yellow, veins dirty yellow. Costa 0.52 of wing length, ratio

of segments (in paratype) 10:8:3. Cilia moderately

short (0.07-0.08 mm), there are 6 pairs between 1st

vein and tip of costa; 3rd vein without a bristle at

base; 4th vein starting at fork; 5th and 6th sinuous.

Axillary margin with 3 hairs. Anal margin broadly

convex. Halteres brown.

FEMALE.—Length 1.10-2.00 mm. Similar to male.

Proboscis (Figure 114) narrow, yellow. Tergites (Figure 112) pale brown, subsequal in length; hind corners of 5th broadly rounded; lateral borders of 6th

convex. Pubescence conspicuous on lateral borders of

tergites; 2nd with bristly hairs. Venter sparsely hairy,

hairs shorter than in male. Costa 0.52-0.53 of wing

length; there are 7-8 cilia between 1st vein and apex

of costa.

MATERIAL EXAMINED.—Besides the types from

Costa Rica and numerous specimens from Brazil, I

saw about 300 specimens from Dominica: 25 pinned

specimens, 48 on slides, and more than 200 in alcohol,

from the following localities: Clarke Hall, Pont Casse,

Manets Gutter, Fond Figures R., d'Leau Gommier,

S. Chiltern Est. (W. W. Wirth; O. S. Flint; T. J. Spilman; 1964-1965).

DISCUSSION.—M. (M.) alajuelensis comes close to

M. (A/.) flavopleura Malloch, but the latter has a

longer costa, the ratio of length of segments is different,
NUMBER 23 37

the fork is larger, and the fourth vein starts behind

fork; the female of M. (M.) alajuelensis can be recognized by the form of the sixth tergite which is distinctive; it is best seen in alcohol material.

VARIATION.—The variation in body-length is remarkable.

52. Megaselia (Megaselia) mutilata, new species

FIGURE 115

FEMALE.—Length 1.00 mm. Head width 0.32 mm.

Frons black brown, as broad as long medially (0.19

mm), lateral length about 0.11 mm, finely pubescent,

with a distinct median line. Four su of equal length,

the upper two-fifths of frontal width apart (6:15), the

lower one-seventh. Antials approximate to eye-margin,

a little above level of upper su, nearly vertically below

anterolaterals which are distinctly higher near eyemargin. Second row nearly straight, the bristles equidistant. Upper postocular bristles strong. A pair of

divergent bristles near lower corner of eye, cheeks

with 4 setulae. Third antennal joint black-brown, of

normal size, width 0.078 mm; arista 0.26 mm, distinctly pubescent. Palpi yellow, of normal shape, with

5-6 bristles on apical half.

Thorax black-brown. Pubescence short, longer behind. Mesopleura bare. Scutellum with 2 bristles and

2 minute hairs.

Abdomen dark brown, venter yellow. Tergites 2-6

subequal in length, gradually narrowed. Pubescence

scarce, short; venter on hind margins of segments 4-6

with a transverse row of hairs inserted in dark spots.

Terminal segments short, fleshy; cerci distinct.

Legs yellow, hind legs more or less brownish yellow.

Foretibia 0.22 mm. Foretarsus slender (0.29 mm),

metatarsus about as long as joints 2 + 3. Midtibia 0.27

mm, hair-seam reaching apical third; end-spur about

two-thirds of metatarsus. Hind femur a little more

than 3 times as long as broad, ventrobasal hairs rudimental. Hind tibia 0.39 mm, posterodorsal cilia inconspicuous, very fine.

Wing (Figure 115) grayish yellow, veins brown.

Length 1.01 mm, width 0.44 mm. Costa 0.42 or 0.43

of wing length, ratio of segments 22:21, with about

13 marginal cilia, length 0.065 mm (there are 5 between rx and apex of costa). Fork and mediastinalis

missing. Third vein with a conspicuous bristle at base,

nearly as long as longest costal cilia; 4th vein weakly

arcuate; 5th but little curved before middle; 6th

slightly sinuous; 7th weak. Axillary margin with 2

hairs. Halteres dark brown.

MATERIAL EXAMINED.—Holotype female, on slide

(USNM 70270), Dominica, Manets Gutter, 7 Mar.

1965, light trap (Wirth).

DISCUSSION.—Among the neotropical species of

group VIII easily recognized by the unforked 3rd vein.

53. Megaselia (Megaselia) brevicostalis (Wood)

Phora brevicostalis Wood, 1910, pp. 243, 246 [$ Q ; England].

Aphiochaeta brevicostalis (Wood).—Wood, 1912, p. 171.—

Braes, 1915a, p. 113.—Lundbeck, 1922, p. 401, fig. 119.

Megaselia (Megaselia) brevicostalis (Wood).—Borgmeier,

1966, pp. 6 [key], 106, figs. 89-90 [$ 9 ; U.S.; South

America].

Aphiochaeta fuscopedunculata Malloch, 1912, p. 498 [Kaslo,

B.C.].—Borgmeier, 1966, p. 107 [synonymy].

Megaselia (Megaselia) frequens Borgmeier, 1962b, p. 312,

448 [Brazil]; 1966, p. 107 [synonymy].

This European species, recently recorded by Borgmeier from North and South America, is represented

by two males: 1 pinned specimen from Dominica,

Layou, 29 May 1966 (G. Steyskal); and 1 male in

alcohol, Dominica, Clarke Hall, Feb. 1965, light trap

(W. W. Wirth). The species has a small bristle at base

of 3rd vein.

54. Megaselia (Megaselia) crinipyga, new species

FIGURES 116-117

MALE.—Length 1.00 mm. Head width 0.31 mm.

Frons slightly broader than long laterally (6:5), about

as broad as long medially (0.21 mm), black-brown,

pubescent, with a distinct median line. Supraantennals

of equal length, the upper about one-third of frontal

width apart, the lower one-sixth. Antials on anterior

border, farther from supraantennals than from eyemargin ; anterolaterals higher, near eye-margin. Second

row weakly convex, the bristles equidistant. Upper

postocular bristle developed. Two bristles near lower

corner of eye. Third antennal joint black, globose,

width 0.8 mm; arista 0.27 mm, distinctly pubescent.

Palpi yellow, well bristled.

Thorax black-brown. Pubescence short, longer behind. Mesopleura bare. Scutellum with 2 bristles and 2

minute hairs.

Abdomen black-brown, venter dark. Tergites subequal in length. Pubescence scarce and short, hardly

longer on hind margin of 6th tergite. Hypopygium
38 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

(Figure 116) small, dark brown, globose, on both sides

with numerous hairs and 1 or 2 bristles. Anal segment

yellowish brown, very short, end-hairs delicate, inconspicuous, directed downward.

Legs castaneous to yellowish brown. Foretibia 0.23

mm, midtibia 0.29 mm, hind tibia 0.41 mm. Foretarsus

slender, 0.33 mm, metatarsus as long as joints 2+3 .

Hind femur about 3 times as long as broad, length

0.44 mm, ventrobasal hairs short and delicate, not

reaching middle. Posterodorsal cilia of hind tibia short

and fine.

Wing (Figure 117) grayish yellow, veins pale brown,

4-7 evanescent apically. Length 0.91 mm, width 0.41

mm. Costa 0.42 length of wing, ratio of segments

13:7:5. Cilia short (0.05 mm) about 5 between tip

of 1st vein and apex of costa. Mediastinalis missing.

Fork narrow and acute. Third vein without a bristle

at base; 4th slightly arcuate on basal half, remainder

straight; 5th but little curved, very weakly recurved at

base; 6th slightly sinuous; 7th weak. Anal margin

broadly convex. Axillary margin with 2 hairs. Halteres

dark.

FEMALE.—Similar to the male. Tergites of subequal

length, 2-6 gradually narrow, 7-10 fleshy. Cerci distinct Wing length 0.91 mm, width 0.39 mm. Costa

0.42, segments 13:6:5.

MATERIAL EXAMINED.—Holotype male (on slide,

USNM 70271), Dominica, Cabrit Swamp, 18 June

1964, at light (O. S. Flint). Paratypes (USNM, CTB

all on slides): 5 males and 2 females, same locality,

date and collector.

DISCUSSION.—M. (M.) crinipyga, new species,

comes close to M. brevicostalis Wood, but in the latter

the halteres are yellow, and the third vein has a bristle

at base.

Genus Woodiphora Schmitz

Woodiphora (Woodiphora) magnipalpis (Aldrich)

FIGURES 118-120

Phora magnipalpis Aldrich, 1896, p. 438 [$, 9 ; St. Vincent

W.I.].

Aphiochaeta magnipalpis (Aldrich).—Brues, 1904, p. 365,

pi. vi: fig. 36 [wing].

Woodiphora magnipalpis (Aldrich).—Schmitz, 1926, p. 75.

Borgmeier, 1932, p. 50 [Costa Rica; Brazil]; 1963a, p. 136,

fig. 155 [forefemur].

Aphiochaeta pilosa Malloch, 1914, p. 33, pi. 1: fig. 4 [? .

Costa Rica].—Borgmeier, 1963a, p. 138 [synonymy].

Aphiochaeta velutinipes Brues, 1924, p. 158 [$ ; U.S.].—

Borgmeier, 1963a, p. 138 [synonymy].

Aphiochaeta opaciventris Brues, 1924, p. 159 [& ; Grenada,

W.I.].—Borgmeier, 1967b, p. 263 [synonymy].

Holotype male (USNM 7759), paratypes males and

females (USNM, MCZ, CTB), all from St. Vincent,

B.W.I. (Aldrich).

MALE.—Palpi (Figure 119) enlarged, with microscopic hairs. Forefemur (Figure 120) with a characteristic black stria on median third of ventral furrow.

Foretarsus slender. Fore-metatarsus, with an anteroventral series of short and some longer sensorial hairs.

Wing clear. Costa 0.54-0.56 of wing length; ratio of

segments 26:22:8 (paratype), 25:22:8 or 22:20:8

(males from Dominica), 26:19:7 (holotype male of A.

opaciventris), 22:20:8 (syntype male of A. opaciventris). Halteres variable: yellow (types) or yellowish

brown or dark brown (specimens from Grenada and

Dominica).

FEMALE.—Palpus slender (Figure 118),-with short

bristles and fine hairs. Forefemur without stria. Foretarsus somewhat stout. Wing as in male. Costal segments 24:22:9. Halteres yellow or dark.

MATERIAL EXAMINED.—Besides the types listed

above, I have seen from Dominica: 25 pinned males

and 18 females, and numerous specimens of both sexes

in alcohol, from the following localities: Macoucheri,

Grande Savane, Woodford Hill, Pagua Bay, Hodges

River Mouth, Fond Figues River, Cabrit Swamp,

Clarke Hall, all collected by W. W. Wirth (Jan. to

Mar. 1965).

DISTRIBUTION.—Antilles, North America, Central

America, Brazil.

DISCUSSION.—The male is easily recognized by the

black stria on median third of forefemur (Figure 120).

SYNONYMY.—As I pointed out in a previous paper

(1963a, p. 138), Aphiochaeta pilosa Malloch (1914)

and Aphiochaeta velutinipes Brues (1924) are synonyms of W. magnipalpis. To these must be added

Aphiochaeta opaciventris Brues (1924), based on a

single male from Grenada, W. I. According to Brues

(1924, p. 161), "This species is at once distinguished

from magnipalpis Aldrich from the same island [the

type-locality of W. magnipalpis is St. Vincent!] by (1)

the much more slender legs, (2) opaque velvety black

abdomen, (3) normally curved fifth vein and (4) by

the complete absence of any cilia on the middle tibiae.

(5) The palpi although similarly enlarged in the male,

are more slender than in magnipalpis."
NUMBER 23

In February 1965, during a visit to the Harvard

Museum, I had the opportunity to examine the holotype male of A. opaciventris (the wing on slide is before me) which I compared carefully with a paratype

male of W. magnipalpis in my collection. I reached

the following conclusions with regard to the differences

appointed by Brues: (1) the legs are not more slender

in A. opaciventris; (2) the abdomen is opaque velvety

black also in W. magnipalpis; (3) the fifth vein is less

curved before middle in A. opaciventris; (4) the midlegs are missing in the holotype of A. opaciventris, but

in a syntype the cilia are present on mid tibia; (5) the

palpi of male are not more slender in A. opaciventris.

The only real differences discovered are (a) the ratio of costal segments; in the type of A. opaciventris

the first section is exactly as long as the other two together [not longer, as stated by Brues]; (b) the fifth

vein is less curved; (c) the halteres are dark. That the

differences of (a) and (b) are due to variation, is

suggested by the fact that they are not present in a

syntype of A. opaciventris taken by Brues together

with the type. That the color of the halteres is variable,

seems to be evident from the presence of the black

stria on forefemur in all males examined and which I

consider as an extraordinary character of high taxonomic value. I conclude from this that A. opaciventris

must be a synonym of W. magnipalpis.

REMARK.—For the remaining species of this genus

see my improved key to the world species (Borgmeier,

1967b, p. 264).

Genus Johowia Silva Figueroa

Johowia parvicornis, new species

FIGURES 121-123

MALE.—Length 1.5-1.9 mm. Head width 0.65 mm.

Frons longer than broad (4:3), width above 0.26 mm,

width below 0.23 mm, a little depressed anteriorly at

middle, black, subopaque, with a distinct median line.

Four equal supraantennals, between and above them

there are some proclinate bristly hairs which are not

always conspicuous; the upper supraantennals are approximately one-half of frontal width apart, the lower

one-third. Antials near eye, on a lower level than lower

supraantennals, vertically below anterolaterals, the

latter are much higher, about at middle between antials

and mediolaterals. Second row straight or slightly convex anteriorly, the preocellars a little farther from each

39

other than from mediolaterals. Eyes large, approximate

below, where they are about one-half of frontal width

apart. One strong lower postocular bristle directed

downward; one weak bristle on jowls; cheeks very

narrow, with 8 weak bristles. Third antennal joint

(Figure 123) dark brown, very small, globose, width

0.09 mm; arista dorsal, long (0.40 mm), finely pubescent. Palpi (Figure 122) yellow, small, clavate, length

0.15 mm, at apex with 5 strong bristles and a smaller

one. Proboscis short, labrum very small, labella with 8

pseudotracheae.

Thorax shining, dark castaneous, pleura paler,

sternopleura yellow. Mesopleura with short uniform

hairs. Scutellum with 2 bristles and 2 short hairs.

Abdomen dull, venter yellow, tergites black-brown,

with grayish pruinosity, 2-5 subequal in length, 6th

a little longer, all with yellow seams on hind margins.

Pubescence scarce and short, not noticeably longer on

6th tergite, 2nd with bristly hairs laterally. Venter

hairy. Hypopygium small, dark brown, both sides of

dorsal part connected beneath anal tube, with short

hairs 2 or 3 of which behind are bristle-like; ventral

plate small, triangular. Anal segment short, somewhat

depressed, dark brown, with short hairs.

Legs slender, yellow, coxae enclosed, hind legs dark

brown except for the coxae and the basal 3rd of femora

which are yellow. Foretibia 0.52 mm; tarsus thin, 0.65

mm, metatarsus longer than joints 2 + 3. Midtibia with

a complete dorsal hair-seam and 7 distinct posterodorsal setulae. Hind femur 3 times as long as broad,

hairs on ventral edge short and inconspicuous. Hind

tibiae with a dorsal hair-seam and 10 distinct posterodorsal setulae increasing gradually in length.

Wing (Figure 121) brownish, veins dark brown.

Length 1.9 mm, width 0.65 mm. Costa 0.57 of wing

length, ratio of segments 17:15:5. Cilia short and very

close. Fork long, acute. Fourth vein with a slight Scurvature at base, concave, deflected at extreme tip;

5th but little curved; 6th slightly sinuous; 7th weak.

Axillary margin with 4 hairs. Halteres yellow with a

brownish tinge.

FEMALE.—Length 1.8 mm. Similar to the male. The

extranumerary supraantennals are more conspicuous.

Third antennal joint a little smaller. Tergites subequal.

Terminal segments fleshy, 7th and 8th with a narrow

chitinous tergite; segments 9-10 tubiform, cerci not

differentiated.

Holotype male (USNM 70272), Dominica, Pont

Casse, 1.7 m east, 10.111.65 (W. W. Wirth). Paratypes
40

(USNM, CTB) 4 males and 1 female (W. W. Wirth),

from the following localities: Pont Casse 10.111.65;

S. Chiltern Est. 20.11.65; Clarke Hall 18.1.65; Manets

Gutter 7. amd 10.111.65. One of the paratypes

(crushed; in alcohol) was dissected for the study of

details; another was badly damaged by accident.

DISCUSSION.—/. parvicornis, new species, is very

close to / . pilipleura Borgmeier (1960; Rio de

Janeiro), but in the latter the mesopleural hairs are

not of uniform length, the costa is longer and extends

beyond tip of third vein, the ratio of costal segments is

different, the sixth tergite has some strong bristles laterally on hind margin, the hypopygium is bristled, and

the anal segment is yellow and compressed.

Genus Apocephalus Coquillett

Apocephalus (Mesophora) insulanus, new species

FIGURES 124-127

MALE.—Length 1.6-1.7 mm. Head width 0.65 mm.

Frons yellow or fuscous yellow, nearly twice as broad

as long medially (12:7), width 0.31 mm. Supraantennals, anterolaterals and mediolaterals wanting. Antials

one-sixth of frontal width apart, a little more approximate than the preocellars, the latter nearly one-third

of frontal width apart. Upper postocular bristle weak,

lower strong; jowls and cheeks bare. Third antenna!

joint (Figure 127) enormous, band-shaped on distal

half, yellow, infuscated on apical half, length 0.44

mm; arista subapical, length 0.20 mm. Palpi (Figure

125) yellow, small and narrow, pointed at apex, length

0.18 mm, with short setulae.

Thorax reddish yellow. Mesopleura bare. Scutellum

with 2 bristles and 2 minute hairs.

Abdomen dull, nearly bare. Venter yellow. Tergites

2 and 6 elongated. Color of tergites somewhat variable,

1, 2, and 6 yellow anteriorly, black posteriorly; 3-5

black, with yellow median spots; all with fine yellow

apical seams. Hypopygium (Figure 126) black-brown,

sides produced behind and apparently connected beneath anal segment, with short hair; apex of left side

whitish; ventral part flat. Anal segment brown, slender, curved, length 0.23 mm, end-hairs differentiated.

Legs yellow, hind femur darkened at tip. Foretarsus

slender, longer than tibia (9:7), fore-metatarsus about

as long as joints 2-4 combined. Midtarsus normal, endspur nearly as long as midmetatarsus (10:11). Hind

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

femur about 3 times as long as broad. Hind tibia with

7 posterodorsal setulae.

Wing (Figure 124) slightly grayish yellow, veins

dark brown. Length 1.56 mm, width 0.65 mm. Costa

0.5 of wing length, ratio of segments 30:18:5. Cilia

short and close, about 8 pairs between 1st and 3rd

veins. Fork small. Fourth vein evenly concave; 5th

nearly straight; 6th but slightly sinuous; 7th weak.

Axillary margin with 3 hairs. Halteres dark brown,

stem yellow.

FEMALE.—Length 1.8 mm. Similar to the male.

Mediolaterals present, 2nd row convex, the preocellars

farther from mediolaterals than from each other. Third

antennal joint conical, twice as long as broad basally,

not reaching vertical border; arista apical, about twothirds of antenna in length. Palpi yellow, longer than

in male (0.23 mm), not pointed at tip, narrow, about

3 times as long as broad, with extremely short hairs,

practically bare. Tergites 2 and 6 elongated, all reddish

yellow with black transverse bands apically. Ovipositor

black, glossy, lanceolate.

MATERIAL EXAMINED.—Holotype male (USNM

70273), Dominica, S. Chiltern Est. 2 Feb. 1965 (W.

W. Wirth). Paratypes (USNM, CTB): one male,

Clarke Hall, 16 Feb. 1965 (Wirth); one male, Fond

Figues River, 21-31 Mar. 1965 (Wirth); one female,

Pont Casse, 29 Jan. 1965 (Wirth); one male (CTB),

Trinidad, B.W.I., Arima Valley, Feb. 1964 (Wygodzinsky & Rozen).

DISCUSSION.—A. (M.) insulanus, new species, may

be confused with A. (M.) antennatus Malloch (1912;

U.S.), but in the latter the halteres are yellow and the

hypopygium (Figure 129) is bristled; the costa is also

a little shorter and the wing is broader (Figure 128).

Figures 128 and 129 were drawn from a male from

Gatlinburg, Tennessee.

Apocephalus (Apocephalus) aztecae Borgmeier

Apocephalus {Apocephalus) aztecae Borgmeier, 1961, p. 56,

figs. 78, 86 [ovipositor and wing; 9 ; Panama].

Holotype female (CTB, mounted) and 4 paratypes

(in alcohol), from Barro Colorado Island, Canal Zone,

16.V.56 (Rettenmeyer), taken with Azteca species.

MALE (undescribed).—Length 1 mm. Similar to

the female. Frons as broad as long medially, width 0.26

mm, dark brown, grayish pruinose, opaque, with a

distinct median line and 2 rudimental supraantennals

close together. First row strongly convex, the antials

parallel and about one-third of frontal width apart.
NUMBER 2 3 41

Second row distinctly convex, the bristles equidistant.

Third antennal joint oval, length 0.14 mm, yellow

basally, darkened apically; arista subapical, short.

Palpi pale yellow, small, with short bristles. Posterior

scutellars slightly shorter than the anterior, the tips of

the 4 scutellars are in line. Abdomen dull, venter yellow. Tergite 2 elongated, 3-5 short, 6th a little longer;

tergites 1-5 velvety dark brown, with fine yellow seams

on hind margins, 6th yellow on basal half, dark behind.

Hypopygium dark brown, somewhat grayish, with

short hairs. Anal segment slender, yellow, apical hairs

differentiated. Legs yellow, hind femur with anteroventral bristles on apical half. Wing as in female.

Halteres yellow.

FEMALE.—In the female from Dominica and in one

female of the type-series 2 microscopic hairs are present

below the rudimental supraantennals. The 2nd row

is distinctly convex anteriorly (not slightly, as stated in

the original description). Scutellars as in male described above. Sternite of the basal capsule of the

ovipositor with a transverse row of 4 hairs (not bristles) ; apical hairs are absent on the sternite, but the

apex is pointed and slightly curved downwards, forming a short tooth.

MATERIAL EXAMINED.—One male allotype (USNM)

Dominica, Clarke Hall, 21-31 Jan. 1965 (W. W.

Wirth); 1 female in alcohol (USNM), Clarke Hall,

1-10 Feb. 1965 (Wirth). One female and 1 male from

Clarke Hall were destroyed by Psocids. I reexamined

the type-series in my collection.

DISCUSSION.—A. (A.) aztecae is close to A. angustinervis Borgmeier (1961, Brazil), but differs by the

yellow halteres, the shorter costa and the structure of

the ovipositor.

Genus Apodicrania Borgmeier

Apodicrania Borgmeier, 1933, p. 384; 1963b, p. 479 [key].

TYPE-SPECIES.—Syneura termitophila Borgmeier,

1923.

Apodicrania termitophila (Borgmeier)

Syneura termitophila Borgmeier, 1923b, p. 57, fig. 6 [S 9 ;

Petr6polis, Rio de Janeiro]; 1925b, p. 233, pi. xi; fig. 53.

Apodicrania termitophila (Borgmeier).—Borgmeier, 1933,

p. 384; 1963b, p. 479.

Two males and 1 female (in alcohol), Dominica,

Clarke Hall, 1-10 Feb. 1963, light trap (W. W. Wirth).

In my collection also from Central America and

from Brazil.

Genus Pseudacteon Coquillett

Pseudacteon antiguensis (Malloch)

LITERATURE AND SYNONYMS.—See Borgmeier,

1963a, p. 200, figs. 196 (ovipositor), 197 (hypopygium).

Holotypemale (USNM 14896) and paratypes males

(USNM, CTB), Antigua, B.W.I.

This is a very striking species which is a parasite

of Solenopsis geminata and S. saevissima. The female

is easily recognized by the large horse-shoe shaped

ovipositor, and the male by the digitiform processes

of the hypopygium. See the redescription and figures in

Borgmeier (1963a).

MATERIAL EXAMINED (according to Wirth).—18

males and 24 females, of which I have seen 3 males and

5 females, Dominica, Clarke Hall, Malaise trap, 11-

31 Jan. 1965 (W. W. Wirth).

DISTRIBUTION.—Antilles, Mexico, Brazil. Probably

occurring also in the southern United States.

Pseudacteon dorymyrmecis Borgmeier

Pseudacteon doTymyrmeeis Borgmeier, 1925a, p. 45 [9 ; Rio

de Janeiro].

Holotype female and 2 paratypes (CTB), Rio de

Janeiro, 3.III.1925 (Borgmeier), with Dorymyrmex

pyramicus.

FEMALE.—Frons with 2 supraantennals and 2:4:4:4

bristles. Antials inclined toward median line. Inner

bristles of 2nd row not reduced in size. Third antennal

joint oval, dark brown; arista short, subapical, practically bare. Palpi small, pale yellow, with short setulae,

the apical hardly differentiated. Scutellum with 4

bristles, the anterior weak. Second tergite elongated,

3-5 short, subequal; 6th tergite represented only by

a rudimental plate on each side which bears 2 hairs.

Sixth sternite near base with a transverse row of 6

erect hairs, the median pair being close together and

bristlelike, the laterals short. Ovipositor black, glossy,

lanceolate, depressed and pointed, flat above; 8th

sternite forming a small tooth basally which is perceptible in side-view. Legs yellow. Midtibia with a dorsal hair-seam which terminates below middle. Hind

femur with some anteroventral setulae on distal half.
42 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Hind tibia with a complete dorsal hair-seam. Hind

metatarsus about as long as joints 2-3 combined. Wing

clear. Costa 0.36 ratio of segments 2:1. Cilia short,

about 13 pairs. Seven vein obsolete. Axillary margin

with 3 hairs. Halteres whitish yellow.

MALE (undescribed).—Similar to the female. Hypopygium without digitiform appendages. Anal segment long, whitish yellow, end-hairs conspicuous.

MATERIAL EXAMINED.—Two males and 5 females

(USNM, CTB), Dominica, Clarke Hall, Fond Figues

River, I.-III.65 (W. W. Wirth), with light trap and

Malaise trap.

DISCUSSION.—P. dorymyrmecis is close to the

European P. formicarum Verrall, which lacks the dorsal hair-seam on midtibia. Also close to P. curriei Malloch (British Columbia), but in the latter the costal

cilia are longer, the median bristles on sixth sternite are

widely placed and the ovipositor is transversely convex

above.

Pseudacteon arcuatus, new species

FIGURE 133

FEMALE.—Length 0.91 mm. Head width 0.33 mm.

Frons black, with 2 proclinate supraantennals and

4:4:4 bristles. Antials near lower corner of frons, inclined toward median line; anterolaterals higher, near

eye-margin. Intermediary bristles wanting. Third

antennal joint dark brown, length 0.19 mm, conical,

distinctly pubescent; arista wanting. Palpi small, yellow, with a few short hairs and one longer hair at apex.

Thorax black-brown. Two dorsocentrals. Mesopleura bare. Scutellum with 2 bristles.

Abdomen dark brown; tergites 2 and 6 elongated,

3rd and 4th short, 5th a little longer. Sixth sternite

with a median pair of long hairs and laterally with 2

or 3 shorter hairs. Ovipositor (Figure 133) heavily

chitinized, black and brown; 7th segment with an

elongate chitinous stripe at each side; lateral lobes of

8th segment oval from above, excavated dorsally;

medial apical piece arc-shaped.

Legs brownish yellow, anterior legs paler. Foretibia

0.19 mm; foretarsus 0.32 mm, metatarsus not as long as

joints 2 + 3 combined. Midtibia 0.22 mm; midtarsus

0.33 mm. Hind femur slender, more than 3 times as

long as broad, ventrobasally with some moderately

long hairs. Hind tibia 0.26 mm, very slender near

base, with a dorsal hair-seam and some delicate posterodorsal cilia.

Wing grayish yellow, veins pale brown. Length 0.93

mm, width 0.46 mm. Costa 0.37 of wing length, ratio

of segments 14:7. Coastal cilia short, 14 pairs in all,

of which 4 pairs on 2nd segment. Fourth vein slightly

concave near base, remainder nearly straight; veins

6 and 7 evanescent apically. Axillary margin with 2

hairs. Halteres brownish.

MATERIAL EXAMINED.—Holotype female (USNM

70274, on slide), Dominica, Clarke Hall, light trap,

Nov. 1964 (T. J. Spilman).

DISCUSSION.—The general structure of the ovipositor is similar to P. pradei Borgmeier from Brazil,

but the shape of the lateral lobes and of the median

apical piece are quite different (see Borgmeier, 1925b,

fig. 48, p. 245).

Genus Metopina Macquart

Metopina recurvata, new species

FIGURES 130-132

FEMALE.—Length 1.0-1.3 mm. Frons dark brown,

shining, sparsely pubescent, with a distinct median line.

Four porrect subequal supraantennals forming a trapezium. Antials inclined toward median line, about midway between upper supraantennals and laterals, the

points of insertion in an oblique straight line. Preocellars weak, nearly in line with anterior ocellus, as

far apart as inner borders of posterior ocelli. Vertical

row with 6 bristles as usual. Third antennal joint

brown, normal; arista apical, with long pubescence.

Palpi yellowish brown, with 5 long bristles.

Thorax brown, pleura yellowish below. Scutellum

with 2 bristles and 2 hairs.

Abdomen dull, dark brown venter enclosed. Tergites

1-4 (Figure 130) transverse, 5th semicircular with a

large gland opening; 6th segment fused together with

5th as usual, membranous. Tergites nearly bare; venter

on segments 3-6, sides of 5-6 and dorsum of 6 with

numerous short hairs inserted in black chitinous rings;

7th segment with a narrow dorsal plate; apical hairs

short.

Legs yellow. Foretarsus longer than tibia. Hind

metatarsus broad, with 4 or 5 transverse combs.

Wing (Figure 131) brownish, veins dark brown,

membrane infuscated beyond costa and along 2 basal

thirds of 6th vein. Length 1.04 mm, width at distal

third 0.45 mm. Costa 0.53 of wing length, ratio of segments 7:12. Cilia short and close. Fourth vein straight;
NUMBER 2 3 43

5th strongly bent near base, the prolongation of the

basal part would meet the costa at a right angle; 6th

with the usual sudden bend behind middle, apical part

gently arcuate. Halteres dark brown.

MALE.—Length 0.8 mm. Similar to the female.

Anal tube hardly protruding. Hind femur without a

sensorial organ on posterior face. Wing (Figure 132)

length 0.78 mm, width at apical third 0.37 mm. Costa

0.5 of wing length, thickened from middle of 1st section to near apex, gradually attenuated beyond tip of

1 st vein; ratio of segments 2:3.

MATERIAL EXAMINED.—Holotype female, on slide

(USNM 70275), (Dominica, Clarke Hall, 11, 20 Mar.

1965 (W. W. Wirth). Paratypes (USNM, CTB) 9

females and 11 males (3 & 6 $ in alcohol), Dominica:

South Chiltern Est., Fond Figues River, Cabrit Swamp,

Pont Casse, Manets Gutter, d'Leau Gommier, Trafalgar Falls, Freshwater Lake, I.-III.1965 (W. W.

Wirth).

DISCUSSION.—M. recurvata, new species, is close to

M. fenyesi Malloch redescribed by Borgmeier (1963a,

p. 212), but in the latter the fifth vein is not so strongly

recurved basally and the basal part is not so steep; the

apical part of 6th vein is straight; the ratio of costal

segments is 4:5 in the female; the costa is not thickened

in the male; the structure of female tergites is completely different.

Genus Dahliphora Schmitz

Dahliphora dispar Borgmeier

FIGURES 134-136

Dahliphora dispar Borgmeier, 1961, pp. 107, figs. 147 [antenna

of male], 149 [heavy veins of wing].

Hototype and Type (<J) (CTB), from Petropolis,

Rio de Janeiro, Brazil, X.I950 (Borgmeier); 2$

paratypes (CTB) from Amapa, Serra do Navio, Brazil

(Bicellicoll.).

MALE.—Length about 0.8 mm. Frons jet-black, lacking the supraantennals, the anterolaterals and the

mediolaterals; 2 preocellars present; vertex with 6

bristles. Third antennal joint (see Borgmeier, 1961,

fig. 147) black, onion-shaped, width 0.78 mm; arista

differentiated, thick, length 0.19 mm, with long

pubescence. Palpi small, narrow, black brown, with

short bristles. Thorax black-brown, mesopleura bare.

Scutellum with 2 subequal bristles. Abdomen black,

dull, tergites subequal. Hypopyguim small, dark brown,

with longish hairs. Anal segment short, apical hairs

differentiated. Legs black-brown. Foretibia 0.16 mm;

foretarsus 0.19 mm, metatarsus as long as joints 2 + 3.

Midtibia 0.16 mm, end-spur about two-thirds of metatarsus. Hind femur 3 times as long as broad. Hind tibia

0.228 mm, without hair-seam (contrary to my statement in 1961) but with a dorsal row of fine cilia. Wing

(Figure 135) slightly brownish, heavy veins brown,

light veins pale brown. Length 0.677 mm, width 0.343

mm. Costa 0.5 of wing length, ratio of segments 17:15.

Cilia short, about 15 pairs. Third vein gently curved;

4th very little concave; 5th strongly curved near base,

distal two-thirds nearly straight; 6th slightly bisinuate;

7th very close to anal margin. Axillary margin without

hairs. Halteres black.

FEMALE (undescribed).—Similar to the male. Frons

with 2 reclinate supraantennals, 2 preocellars, and 6

vertical bristles; all bristles weak. Third antennal joint

(Figure 136) normal, black, onion-shaped, width 0.07

mm; arista 3-segmented, apical, length 0.22 mm, with

long pubescence. Tergites subequal, black. Wing 0.7

mm, width 0.378 mm. Costa 0.5 of wing length, ratio

of segments 17:19. Third vein strongly bent at apical

4th; 5th bent at basal 3rd (in male at basal 4th).

MATERIAL EXAMINED.—Two males (USNM, CTB)

and 1 female (allotype USNM; paratype 9, CTB)

Dominica: South Chiltem Est. 2.II.65; Clarke Hall

18.1.65; Fond Figues River, 3.II.65 (W. W. Wirth).

DISCUSSION.—D. dispar differs from D. crenaticornis by the structure of the antennae and the number

of scutellars. The sexual dimorphism in the venation

is remarkable.

Genus Pachyneurella Braes

Pachyneurella Brues, 1904, pp. 382, pi. ix: fig. 58 [wing], 59

[hypopygium].

Puliciphora Dahl [in part].—Brues, 1906, p. 14.—Borgmeier

and Schmitz, 1923, p. 140 [note].—Schmitz, 1929, pp. 25,

157,159.—Borgmeier, 1960, p. 314.

Subgenus Termitometoecus Borgmeier and Schmitz, 1923,

p. 140 [type-species: Puliciphora (Termitometoecus) bicolor Borgmeier and Schmitz].

TYPE-SPECIES.—Phora venata Aldrich, 1896.

MALE.—Frons broader than long, with 4 porrect supraantennals and 2:4:6 bristles. Third antennal point

globose or oval; arista apical. Antennal foveae coalescent. Mesopleura bare. Costa with a slight swelling

before tip of 1st vein; 1st costal division shorter than

2nd; 3rd vein unforked. Costal cilia hair-like. Mediastinalis present; veins 4-7 reaching margin. Axillary
44

margin without hairs. Legs slender. Hind tibia without

dorsal hair-seam and without posterodorsal cilia.

Hypopygium asymmetrical, with bristles. Anal tube

protruding.

FEMALE.—Degenerate, wingless. Tegument strongly

shining. Frons with 4 proclinate supraantennals and

0:0:4 bristles. Three ocelli. Third antennal joint globose, pointed; arista apical. Thorax short. Prothoracic

bristle wanting. Hind corners without a papilla-like

elevation. Abdomen strongly convex; 5th tergite at base

with a gland-opening and a small flap covering it; 6th

tergite very small.

DISCUSSION.—This genus which was for a long time

considered as a synonym of Puliciphora, must be reinstated, as is evident from a comparative study of three

species represented by males. The male differs from

Puliciphora by the bristly hypopygium and by the

slight swelling on first costal segment. The female has

only four vertical bristles; the bristles of the first and

second rows are wanting. The tegument is strongly

shining and there is no papilla for the insertion of the

alar bristle on the hind corners of the thorax.

Four species are known: Pachyneurella venata (Aldrich) ($9), P. bicolor Borgmeier and Schmitz

( $ 9 ), P. nitida (Malloch) [new combination] ( 9 ),

and P. haplopyga, new species ( $ ) (p. 46).

Pachyneurella venata (Aldrich)

FIGURES 140,141

Phora venata Aldrich, 1896, p. 436 [$ ; St. Vincent, W. I.].

Pachyneurella venata (Aldrich).—Brues, 1904 p. 382, pi.

Dt: figs. 58 [wing], 59 [hypopygium].

Puliciphora venata (Aldrich).—Brues, 1906, p. 15; 1919a,

p. 506; 1923, p. 22 [$ 9, Grenada; not Bolivia].—Borgmeier, 1960, p. 336 [in part; not 9, not fig. 97 = bicolor].

Lectotype male, herewith designated (USNM) and

1 paratype male (MCZ), from St. Vincent, B.W.I.

(Aldrich). Probably 1 "type" also in British Museum,

London (teste Brues, 1904, p. 382).

MALE.—Length 1.0-1.3 mm. Frons broader than

long laterally (3:2), black-brown, opaque, sparsely

pubescent, without median furrow. There are 4 equal

porrect supraantennals and 2:4:6 bristles; anterolaterals wanting; Antials inclined toward median line,

close to mediolaterals; 2nd row concave anteriorly.

Jowls-cheeks with an ascending row of 5 bristles. Third

antennal joint dark brown, globose, of normal size;

arista apical, distinctly pubescent. Palpi yellowish

brown to dark brown, narrow, with short bristles.

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Thorax dark brown, opaque. Two dorsocentrals.

Mesopleura bare. Scutellum with 2 bristles.

Abdomen black-brown to black, dull. Second tergite

a little elongated, tergites 3-6 subequal in length, 6th

tergite followed by a membranous yellow apical band

which partly covers the hypopygium. Pubescence short,

6th segment laterally and ventrally with hairs placed in

chitinous spots. Hypopygium (Figures 141-142) castaneous, large, asymmetrical, longer than high, at left

behind with a long black process which is somewhat

flattened, and at right with a pale process; at left above

there is a single bristle near the base of the anal tube;

and at right there are apically about 6 bristly hairs.

Anal segment large, depressed, the tergites laterally on

each side with 4 strong bristles; apical hairs of ventrite

inconspicuous.

Legs yellowish brown, forecoxae paler. Foretarsus

slender, metatarsus slightly concave on anterior border,

but not suddenly constricted as in Pachyneurella bicolor. End-spur on mid- and hind tibia short. Hind

tibia without hair-seam and without cilia. Hind metatarsus with 5 transverse combs of hairs.

Wing (Figure 140) brownish yellow, veins dark

brown. Length 1.37 mm, width 0.67 mm. Costa 0.54 of

wing length, ratio of segments 2:3; 1st section at distal

3rd with a small swelling. Costal cilia very short and

close. Fourth vein nearly straight; 5th but little curved before middle, remainder nearly straight; 6th

slightly sinuous. Axillary margin without hairs.

Halteres dark brown.

FEMALE (undescribed).—Length 1.2 mm. Frons

transversely convex, pale brown to dark brown, shining, pubescent, with 4 supraantennals and 4 bristles on

vertical border. Three ocelli. Eyes somewhat reduced,

about twice as long as 3rd antennal joint; the latter is

brown, globose, pointed; arista apical, distinctly pubescent. Palpi brown, with long bristles.

Thorax short, about twice as broad as long, shining,

reddish yellow or yellowish brown, lighter than head.

Prothoracic bristle wanting. Hind corners without alar

papilla. Hind border with 6 bristles, the 2 inner ones

close together, the sublateral close to the lateral, the

latter placed a little more cephalad, the sublateral in

line with the inner bristles. Abdomen strongly convex,

shining. Venter fuscous; the 6 tergites deep black

castaneous; the structure of the tergites more or less

as in Pachyneurella bicolor, 6th very small; 5th with

a small gland opening basally, with a small raisable flap
NUMBER 2 3 45

covering it. Pubescence short, distinct, yellowish; hairs

on membranous parts of segments 4-6 partially placed

in dark chitinous spots. Cerci distinct. Legs yellow.

MATERIAL EXAMINED.—Besides the types (USNM,

MCZ), I have seen 23 males and 2 females from

Dominica: Clarke Hall, Manets Gutter, Pont Casse,

Sylvania, Trafalgar Falls, d'Leau Gommier, Fond

Figues R., South Chiltern Est, Macoucheri, I.—III.

1965 (W. W. Wirth).

DISCUSSION.—I confused Pachyneurella venata for

a long time with P. bicolor Borgmeier and Schmitz

(1923; Brazil) owing to the fact that the venation of

the males is very similar, both having the small swelling of the costa in the first section; the comparison

of the male genitalia (Figures 137-138 with 141-142)

shows that bicolor is not a synonym of P. venata as I

wrongly stated in a previous paper (1960). The female

I described in 1960 as venata, is actually bicolor. The

first true venata females I have seen are the two specimens mentioned above from Dominica; the abdomen

is uniformly dark brown and not cross-banded. The

females were mentioned by Brues (1919a, p. 506) but

not described, from Grenada, W. I.; they must have

been lost because they are not in the Brues collection;

the females mentioned from Bolivia were probably

bicolor.

A single male (on slide) from San Jos6, Costa Rica,

in the author's collection (H. Schmidt leg.) shows that

there is a third species (see Pachyneurella haplopyga,

new species, below) with the characteristic small swelling of the costa in the first costal section; the hypopygium (Figure 139) is quite different from either venata

or bicolor.

Pachyneurella haplopyga, new species

FIGURE 139

MALE.—Length 1.3 mm. Frons black-brown. Anterolateral bristles wanting. Jowls-cheeks with an ascending row of bristles. Third antennal joint dark brown,

of normal size (diameter 0.078 mm). Arista long, distinctly pubescent. Palpi slender, brown, with moderate

bristles.

Thorax dark brown. Two dorsocentrals. Mesopleura bare. Scutellum with 2 bristles.

Abdomen dark brown. Tergites subequal in length,

6th apically with a membranous seam covering base

of hypopygium. The latter is relatively small, but essentially of the same structure as in Pachyneurella

venata and P. bicolor. The most important distinctive

character is the moderate lanceolate process at the left

side, which seems to be homologous to the black process

present in P. venata. Hind border of epandrium at left

with a row of 5 or 6 bristles; hind corner at right

rounded apically and with bristles; the ventral process

at right is also present but hardly discernible in the

preparation of the unique specimen.

Legs brown. Foretibia 0.25 mm; foretarsus slender,

fore-metatarsus apparently not concave on anterior

border. Midtibia 0.32 mm, hind tibia 0.40 mm. Endspurs II and III short.

Wing grayish yellow, veins pale brown. Length 1.32

mm, width 0.59 mm; first costal segment with a slight

swelling behind middle. Venation very similar to P.

venata; 4th vein slightly upturned at extreme base

(straight in P. venata). Halteres dark.

MATERIAL EXAMINED.—Holotype male, (CTB, on

slide), Costa Rica, San Jose, H. Schmidt, 1940.

DISCUSSION.—Differs from P. venata by details of

the hypopygium, especially by the lanceolate process

at left side of the epandrium, and by the fourth vein.

Genus Homalophora Borgmeier

Homalophora Borgmeier, 1923a, p. 178, fig. 14.

TYPE-SPECIES.—Homalophora reichenspergeri

Borgmeier, 1923a ( 9 ).

The male of this genus is recognizable by the long

sensorial hairs near base of forefemur and the long

chitinous clasper at right of hypopygium.

Five species are known: H. attae Borgmeier ( S 2 ;

Brazil) ; H. epichaeta Borgmeier (2 ; Panama) ; H.

metae Seevers ( 9 ; Colombia); H. reichenspergeri

Borgmeier ( 9 ; Brazil) ; H. sensibilis, new species ( & ;

Dominica, W.I.).

Homalophora sensibilis, new species

FIGURES 143-146

MALE.—Length 1-1.2 mm. Frons broader than long,

a little produced medially, dark brown, scarcely shining, sparsely pubescent, with 4 equal porrect supraantennals and 2:4:6 bristles; anterolaterals wanting;

antials inclined toward median line, about midway

between upper supraantennals and mediolaterals;
46 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

second row concave anteriorly. Third antennal joint

normal, dark brown; arista apical, longer than eye,

distinctly pubescent. Palpi of normal shape, with moderate bristles.

Thorax dark brown, subopaque. Mesopleura bare.

Scutellum with 2 bristles and 2 minute hairs.

Abdomen dark brown, dull. Tergites 2-5 subequal,

6th elongated. Pubescence scarce and short. Hypopygium (Figures 145, 146) dark brown, large, asymmetrical; dorsal part at right reduced to a small

triangular piece close to anal tube, provided with 3

short hairs; dorsal part at left large, rounded behind;

ventral part at right with a long slender forceps curved

and spoon-shaped apically, medially in profile (from

left) with 2 short fingerlike processes and between

them 2 blunt black teeth; anal segment long, brown,

with short hairs.

Legs brown, forecoxae yellowish. Forefemora with a

dark ventral stria on basal half and about 7 fine long

sensorial hairs (Figure 143). Foretarsus slender.

Apical spur of midtibia short. Hind femur ventrobasally at extreme base of posterior face with a short

narrow black stria which at great magnification shows

2 blunt elevations (Figure 144); ventral edge with 3

or 4 short hairs near base. Hind tibia without hairseam but with numerous fine cilia. Hind metatarsus

with 5 transverse combs of hairs.

Wing grayish yellow, veins dark brown. Length 1.0

mm, width 0.5 mm. Costa 0.51 of wing length, ratio

of segments 14:23. Cilia very short. Veins 4-7 approximately as in Pachyneurella venata. Halteres dark

brown.

MATERIAL EXAMINED.—Holotype male, pinned

(USNM 70276), Dominica, Macoucherie, 15 Mar.

1965 (W. W. Wirth). Paratypes: 2 males, 1 pinned

and the other in alcohol (USNM), Manets Gutter,

7 Mar. 1965 (Wirth); 1 male, on slide (USNM),

Clarke Hall, Junae 1964 (O. S. Flint); 1 male, pinned

(CTB), Clarke Hall, 1-10 Feb. 1965 (Wirth); 1 male,

on slide (CTB), Cabrit Swamp, 23 Feb. 1965 (Wirth).

DISCUSSION.—H. sensibilis, new species, is very close

to H. attae Borgmeier (1958, p. 402, figs. 104-106) but

differs by the right clasper of the hypopygium which

is more slender and slightly dilated and spoon-shaped

at apex. The venation is also slightly different: in

H. sensibilis, new species, the fourth vein is practically

straight, in H. attae it is slightly upturned at extreme

apex.

Genus Chonocephalus Wandolleck

Chonocephalus Wandolleck.—Borgmeier, 1963, p. 247; 1967a,

p. 201.

TYPE-SPECIES.—Chonocephalus dorsalis Wandolleck (Bismarck Archipelago).

Mr. Charles N. Colyer was kind enough to call my

attention to the fact that in the checklist of Chonocephalus I published in 1967 (a, p. 202) four species described by Paulian from Madagascar (1958, Memoirs

Institute Scientifique Madagascar, (E) vol. 9, pp. 10-

14, figs, la-d and 2a-m) are missing; these are: C.

dimakae Paulian, C. hibisci Paulian, C. madagascariensis Paulian, and C. vadoni Paulian.

Chonocephalus furcatus Borgmeier

FIGURES 147, 148

Chonocephalus furcatus Borgmeier, 1967a, p. 206, figs. 27,

30 [=figs. 147 and 148 in the present paper].

Original description:

MALE.—Length about 1.2 mm. Head width 0.30 mm.

Frons dark brown. External bristle on upper border of antennal grooves 0.10 mm long. Third antennal joint pointed,

about as broad as long. Palpi of normal shape and size, apical

hair slightly differentiated.

Thorax castaneous; humeri, pteropleura and stemopleura

paler. Scutellum with 4 bristles, the anterior a little shorter.

Abdomen opaque. Tergites 2-5 subequal in length, 5th

elongated, all with a fine yellowish seam behind. Hypopygium

[Figure 148] high and short, brown, slightly shining. Epandrium at left side of anal opening with a slender process (p)

and farther below with a long single bristle; at both sides

of the anal opening there is a series of hairs which are curved

apically. Hypandrium at left with a long slender fork [Figure

148J and at right with a lobulate process provided apically

with two fingerlike structures close together [Figure 147] and

which became visible only after removing the inner copulatory organs.

Legs slender, yellowish brown, mid tibia whitish yellow

on distal half. Fore tibia 0.20 mm, fore metatarsus 0.10 mm.

Mid tibia 0.26 mm, mid metatarsus 0.13 mm. Hind tibia 0.36

mm; hind tarsus dilated as usual; hind metatarsus 0.14 mm.

End-spurs of mid and hind tibia very short.

Wing brownish, clouded more or less as in punctifascia

Borgmeier (1935[b]) but with less contrast [see remarks p. 48,

under C. punctifascia]. Length 1.20 mm, width 0.53 mm. Costa

about 0.6 of wing length, exceeding the point of contact of

3rd vein with the costa; apical swelling of 3rd vein slightly

diverging from anterior border of wing. Fourth vein slightly

concave; 5th straight except at apex which is slightly deflected ; 6th slightly sinuous. Halteres dark brown.

Holotype male, mounted on minuten pin (USNM [70440]),

from Dominica, South Chiltern Est., 20 Feb. 1965 (W. W.
NUMBER 2 3 47

Wirth). One paratype, dissected, on slide (CTB), Dominica,

Trafalgar Falls, 6 March 1965 (W. W. Wirth).

DISCUSSION.—Chonocephalus furcatus is easily recognized

by the slender fork on the left side of the hypopygium.

Chonocephalus dominicanus Borgmeier

FIGURE 149

Chonocephalus dominicanus Borgmeier, 1967a, p. 206, fig. 19

[=fig. 149 in the present paper].

Original description:

MALE.—Length 0.91 mm; of a mounted specimen 0.8 mm.

Head width about 0.23 mm. Frons dark brown, opaque;

structure and chaetotaxy as usual. Ocellar bristles 0.065. mm.

Third antennal joint dark brown, pointed, about as long as

broad (0.065 mm); arista 0.19 mm. Palpi brown, of normal

shape and size.

Thorax brown, opaque. Tergites 2-5 subequal in length,

6th elongated; apical seams faint, yellow. Hypopygium [Figure 149] globose, pale brown, slightly shining. Epandrium with

numerous hairs disposed as in [Figure 149]; posterior border

not well perceptible in micro-preparation. Hypandrium mostly

concealed by the internal copulatory organs (i).

Legs brownish yellow, mid and hind tibiae paler on distal

half. Fore tibia 0.156 mm, mid tibia 0.247 mm, hind tibia

0.325 mm. All tarsi longer than respective tibiae. End-spurs

of mid and hind tibia rudimental.

Wing yellowish brown, faintly shaded. Length 0.94 mm,

width 0.44 mm. Costa 0.56 of wing length. Fourth vein

weakly concave; 5th nearly straight, slightly deflected near

apex; 6th slightly sinuous. Halteres dark brown.

Holotype male (on slide USNM [70441], Dominica, Clarke

Hall, 11-20 May 1964 (O. S. Flint). One paratype male (on

slide, CTB), ibidem, June 1964 (O. S. Flint), light trap. One

male from Clarke Hall, Jan. 1965 (W. W. Wirth) was lost

during laboratory work.

DISCUSSION.—Chonocephalus dominicanus comes close to

punctifascia Borgmeier (1936[b]), but the latter has a longer

costa (up to 0.66 of wing length).

Chonocephalus jamaicensis Brues

Chonocephalus jamaicensis Brues, 1915b, p. 102 [9 ; Jamaica].—Borgmeier, 1963a, p. 249.

MATERIAL EXAMINED.—Numerous females in alcohol, collected on decaying fruit, Dominica, Clarke

Hall, 29 Feb. 1965 (W. W. Wirth).

DISTRIBUTION.—Antilles, Mexico, U.S. (Louisiana),

Surinam.

The female was recently redescribed by Borgmeier

(1963a). It is characterized by the hyaline dots on

hind margins of tergites. The male is unknown.

Chonocephalus mexicanus Silvestri

Chonocephalus mexicanus Silvestri, 1911, p. 172, figs. 1, 2

[ 9 ; Cordoba, Mex.].—Borgmeier, 1963a, p. 250.

Chonocephalus buccatus Malloch, 1912, p. 510, pi. 40: fig. 5

( 9 ; Guatemala).

One female, (USNM on slide), Dominica, Clarke

Hall, 1-30 June, 1964 (O. S. Flint). Hind margins of

abdominal tergites without hyaline dots. The holotype

was collected by Silvestri and is probably in his collection at Portici.

Chonocephalus punctifascia Borgmeier

Chonocephalus punctifascia Borgmeier, 1935b, p. 257, figs.

1-13, pi. 1: fig. 1 [wing; $ 9 ; Rio de Janeiro]; 1967a,

p. 210.

There are 4 males on slides, which agree perfectly

with the males from Rio de Janeiro; Dominica, Clarke

Hall, May to June 1964 (O. S. Flint).

In a previous paper (1963a, p. 210) I noted the

following:

Remark: The micro-preparation of the wing of Chonocephalus punctifascia from which the photo was made which was

published by Borgmeier (1935b, pi. 1: fig. 1), is in my collection. It does not show such a strong contrast between the

darker areas and the lighter ones as shown in the mentioned

figure. The contrast is probably due to the fact that a

panchromatic film was used by the photographer, J.

Barbosa.

It is possible that C. punctifascia is a synonym of C.

jamaicensis, as I surmised in a previous paper (1963a).

Males associated with females from the type-locality

are necessary to decide this question.

Tribe Beckerinini

Genus Beckerina Malloch

Beckerina Malloch, 1910, pp. 17, 90.—Borgmeier, 1963a,

p. 128.—Schmitz, in Lindner, 1956, p. 376.

TYPE-SPECIES.—Phora umbrimargo Becker

(Europe).

When I described Beckerina lucifrons (1925b), I

failed to make a comparison with B. fuscohalterata

Enderlein, because Schmitz, who had seen the type of

the latter, stated wrongly that it had only two scutellar

bristles. When I redescribed the type later (1935a), I

did not realize that the differences from B. lucifrons

Borgmeier (1925b) are not satisfactory: the ratio of

frontal width/length given for B. fuscohalterata (5:3)
48 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

is practically the same as that given for B. lucifrons

(4:2.5), and the venation is also practically identical.

Therefore, I consider lucifrons as a synonym of B.

fuscolaterata, which seems to be the only species of the

genus with hairy mesopleura.

B. nigricornis Borgmeier (1925b) is a color variant

of B. chelifera Borgmeier, as is shown by a re-examination of the types in my collection. Both are synonyms of

B. neotropica Brues (1919a; Grenada), of which I

have seen the holotype (MCZ) ; a paratype is in my

collection. Brues, in his description, does not mention

the dorsal hair-seam on the hind tibia; he refers only

to the absence of bristles and setulae. The differences

indicated by Beyer in his key (1966, p. 171) for the

costal sections of B. chelifera ("I deutlich kuerzer als

II") and nigricornis ("I deutlich kuerzer als 11 +

III") are proved to be nonexistent by the photos of

the wings I published in 1925 (pi. in: figs. 12-13).

As I pointed out in a previous paper (1966, p. 124),

B. relicta Borgmeier (1963a) is a synonym of B. aliena

Malloch (1919) which has four scutellars, not two as

wrongly stated by Malloch.

All species of the genus Beckerina can be divided

into three groups, on account of the absence or presence of one or more hair-seams on the hind tibia: (a)

hind tibia simple; (b) hind tibia with one hair-seam;

(c) hind tibia with several hair seams.

Beyer in his description of B. burmicola does not

refer to the condition of the mesopleura, the length of

costa, and the ratio of costal segments. In the following

key I supposed that the mesopleura are bare, and that

the costal index is approximately 0.5 of wing length.

Key to the World Species of the Genus Beckerina

1. Hind tibia simple 2

Hind tibia with 1 or more hair-scams 6

2. Frons with 4 + 4:4:4 bristles; costa 0.68 (U.S.) (cf) B. orphnephiloides Malloch

Frons with 2 + 4:4:4 bristles; costa shorter 3

3. Halteres black (Europe; Alaska) (cf 9 ) B. umbrimargo Becker

Halteres yellow 4

4. Yellow species (U.S.) (cf 9 ) B. luteola Malloch

Dark species 5

5. Costa 0.47; wing length 1.8 mm (U.S.) (cf 9 ) B. similata Malloch

Costa 0.5 (?); wing length 3 mm (Burma) (cf9) B. burmicola Beyer

6. Hind tibia with 1 hair-seam 7

Hind tibia with more than 1 hair-seam 12

7. Halteres dark (Antilles; Central and South America) (cf 9 ) B. neotropica Brues

Halteres yellow 8

8. Frons with 4 + 4:4:4 bristles 9

Frons with 2 + 4:4:4 bristles 10

9. Foretarsus (cf) dilated (Brazil) (cf 9 ) B. irregtdaris Borgmeier

Foretarsus (cf) slender (Costa Rica) (cf 9 ) B. costaricana Borgmeier

10. First costal section one and one-half as long as 2 (9)5 . aliena Malloch

First costal section one and one-fourth as long as 2 or subequal 11

11. Hypopygium laterally with 2 finger-like processes behind (Dominica, W.I.)

(cf 9 ) B. dactylata, new species

Hypopygium with 1 short process (Brazil) (cf 9 ) B. luteihalterata Borgmeier

12. Mesopleura hairy above (Brazil) (cf9 )B. Juscohalterata Enderlein

Mesopleura bare 13

13. Costa 0.52; the 3 lateral bristles of frons equidistant (New Zealand) . (cf) B. polysticha Schmitz

Costa shorter; the 3 lateral bristles on frons not equidistant 14

14. Costa 0.42; sections 22:11:4 (Brazil) (cf 9 ) B. nudipleura Borgmeier

Corta 0.47; sections 25:15:7 (Dominica, W.I.) (cf) B. dominie ana, new species
NUMBER 2 3 49

Beckerina dactylata, new species

FIGURES 150, 151

MALE.—Length 1.6 mm. Head width 0.65 mm.

Frons broader than long laterally (3:2), black, anterior margin yellowish, shining, scarcely pubescent. Two

divergent postantennals close together. First row

slightly convex, the antials twice as far apart as the

preocellars. Second row straight. Eyes large. Three

bristles on inferior corner of eyes. Cheeks narrow, with

4 short hairs. Third antennal joint globular, of normal

size, reddish brown, darkened at apex. Arista a little

longer than frontal width, distinctly pubescent. Palpi

pale yellow, or normal size, with strong but moderately

long bristles.

Thorax reddish yellow. Mesopleura bare. Scutellum

with 4 long bristles nearly in line, the inner ones slightly

shorter as usual.

Abdomen: venter yellow; tergites black, opaque,

with narrow yellow hind apical margins; tergites 2 and

6 elongate, 3-5 subequal. Pubescence extremely scarce

and short. Hypopygium large, yellow, subshining, on

each side with 2 finger-shaped processes, the posterior

one (forceps) with short spinuli on inner side near

apex (Figure 151). Anal segment papilliform.

Legs yellow, hind femur darkened at tip. Foretarsus

slender. Hind femur without fringe on ventral edge.

Mid- and hind tibia with a complete dorsal row of

palisade-like hairs, but without posterodorsal cilia.

Wing (Figure 150) grayish yellow, veins brown.

Length (holotype) 1.45 mm, width 0.62 mm. Costa

0.46 of wing length, ratio of segments 13:7:4. Costal

cilia short. Venation similar to B. luteihalterata Borgmeier (1925b, pi. 3 fig. 11), but the costa is not so

conspicuously thickened on distal half, and the 5th

vein is nearly straight. Veins 4-7 evanescent apically.

Mediastinalis present. Axillary margin with 3 bristles.

Halteres yellow.

FEMALE.—Length 1.8 mm. Similar to the male.

Tergites subequal, gradually narrowed, brown anteriorly, behind with transverse black band and yellow

seams. Cerci distinct. Wing length 1.95 mm. Costa

about 0.5 of wing length, ratio of segments 18:11:4.

MATERIAL EXAMINED.—Holotype male (pinned,

abdomen on card; USNM 70277), Dominica, South

Chiltern Est., 2 Feb. 1965 (W. W. Wirth). Paratypes:

1 female (pinned; USNM), South Chiltern Est. 20

Feb. 1965 (W. W. Wirth); 1 female (on slide;

USNM), Clarke Hall, 21-29 Apr. 1964 (O. S. Flint);

1 male (pest-damaged; CTB; wing on slide), Manets

Gutter, 2 Mar. 1965 (W. W. Wirth); 1 female (CTB),

Fond Figues River, 12 Apr. 1966 (R. J. Gagn6).

DISCUSSION.—B. dactylata, new species, comes close

to B. luteihalterata Borgmeier from Brazil, but the latter has only one finger-like process on each side of the

hypopygium and the process is shorter; the venation is

also different (fifth vein distinctly curved; costa more

thickened on apical half).

Beckerina dominicana, new species

FIGURE 152

MALE.—Length 1.6-1.8 mm. Frons distinctly broader than long laterally (3:2), width 0.31 mm, honey

yellow, shining, sparsely hairy, with a few crossed

hairs anteriorly, without median line. Postantennals

strong, close together. First row nearly straight, the

antials about two-fifths of frontal width apart, a little

more than the preocellars; anterolaterals close to anterior corner of frons, a little removed from eye-margin. Second row slightly convex, the bristles equidistant. One strong lower postocular bristle; 2 strong

bristles on jowls; cheeks with a few short hairs. Third

antennal joint round, brown; arista short (0.39 mm),

pubescent. Palpi pale yellow, flat, the bristles strong

but of moderate length.

Thorax subshining, yellowish brown or brownish

yellow above, pleura pale yellow. Two dorsocentrals.

Scutellum with 4 subequal bristles.

Abdomen dull, venter yellow; tergites velutinous

black-brown, 2-5 subequal, 6th a litle longer, all with

pale brown or yellowish apical bands. Pubescence

short, scarce dorsally. Hypopygium small, brown, at

left with a small rounded hairy lobe; a digitiform

process is not perceptible; ventral plate rounded. Anal

segment yellow, short, depressed, with short hairs.

Legs yellow, hind femur blackish near tip. Foretarsus slender, longer than tibia (4:3). Mid tibia with

several irregular straight or oblique hair-seams, some

of which ending at middle, 1 reaching apex. Hind

femur 3 times as long as broad. Hind tibia with a complete anterior hair-seam and several irregular ones on

dorsal face, between both there is a bare strip with a

single row of short hairs; no setulae.

Wing (Figure 152) grayish yellow, veins dark brown.

Length (paratype) 1.48 mm (1.69 in holotype), width

0.59 mm. Costa 0.47 of wing length, ratio of segments

25:15:7. Cilia moderately long (0.11 mm), widely

placed; there are 5 pairs between 1st and 3rd veins.
50 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Mediastinalis wanting. Fork long, acute. Fourth vein

weakly concave; 5th but little curved; 6th slightly

sinuous. Axillary margin with 3 hairs. Halteres dark

brown.

Holotype male (USNM 70278), paratype male

(CTB; mostly destroyed by psocids; wing on slide),

Dominica, South Chiltern Est., 2 Feb. 1965 (W. W.

Wirth).

DISCUSSION.—B. dominicana, new species, is close

to B. nudipleura, which has a shorter costa and a

longer first costal section.

Literature Cited

Aldrich, J. M.

1896. Dolichopodidae and Phoridae. In Williston, On

the Diptera of St. Vincent (West Indies). Transactions of the Entomological Society of London

(1896): 309-345, 435-438, plate 12:figures 108-

119.

Beyer, E.

1966. Neue und Wenig Bekannte Phoriden, Zumeist aus

dem Bishop Museum, Honolulu. Pacific Insects,

8:165-217, 20 figures.

Bigot, J. M. F.

1857. Dipteros. Pages 328-349. In Sagra, Historia fisica

politico y natural de la Isla de Cuba, 7:371 pages.

Paris "1856." [French edition: 1857:783-829.]

Borgmeier, T.

1923a. Neue Phoridengattungen aus Brasilien. Zeitschrift

des Deutschen Vereins fuer Wissenschaft und

Kunst,3 (1922): 155-181, 14 figures.

1923b. Novos Phorideos brasileiros. Boletim do Museu

Nacional, Rio de Janeiro, 1:51-59, 6 figures.

1925a. Eine neue Myrmecophile Phoride aus Brasilien.

Societas Entomologica, Stuttgart, 40:45.

1925b. Novos subsidios para o conhecimento da familia

Phoridae. Archivos do Museu Nacional, Rio de

Janeiro, 25:85-281, 64 figures, 17 plates.

1932. Sobre alguns Phorideos neotropicaes do Museu de

Hamburgo. Revista de Entomologia, 2:43-52, 1

figure.

1933. Apodicrania, um novo genero da familia Phoridae.

Revista de Entomologia, 3:384.

1935a. Die Typen der von Prof. G. Enderlein im Jahre

1912 Beschriebenen Suedamerikanischen Phoriden. Revista de Entomologia 5:427-470, 17 figures, plates 5-6.

1935b. Sobre o cyclo evolutivo de Chonocephalus Wandolleck e uma nova especie de Melaloncha Brues,

endoparasita de abelhas. Archivos do Instituto

de Biologia Vegetal, 2:255-265, 14 figures, 1 plate.

1950. Revision der Gattung Coniceromyia Borgmeier,

nebst Beschreibung von sechs neuen Artcn. Revista de Entomologia, 21:281-299, 18 figures, 4

plates.

1958. Neue Beitraege zur Kenntnis der Neotropischen

Phoriden. Studia Entomologia 1:305-406, 121

figures.

1960. Gefluegelte und Ungefluegelte Phoriden aus der

Neotropischen Region, nebst Beschreibung von

sieben neuen Gattungen. Studia Entomologia,

3:257-374, 124 figures.

1961. Weitere Beitraege zur Kenntnis der Neotropischen

Phoriden, nebst Beschreibung einiger DohrniphoraArten aus der Indo-Australischen Region. Studia

Entomologia, 4:1-112, 181 figures.

1962a. Some New North American species of the Dipterous Family Phoridae. Rivista Brasileira de Biologia,

22:65-82, 17 figures.

1962b. Versuch einer Uebersicht der Neotropischen Megaselia-Arten, sowie neue oder Wenig Bekannte

Phoriden Verschiedener Gattungen. Studia Entomologia, 5:289-488, 132 figures.

1963a. Revision of the North American Phorid Flies, Part

I. Studia Entomologia, 6:1-256, 232 figures.

1963b. New or Little Known Coniceromyia, and Some

Other Neotropical and Paleotropical Phoridae.

Studia Entomologia, 6:449-480, 74 figures.

1964. Revision of the North American Phorid Flies,

Part II. Studia Entomologia, 7:257-416, 194

figures.

1966. Revision of the North American Phorid Flies,

Part III. Studia Entomologia, 8(1965): 1-160,

111 figures.

1967a. New or Little Known Phorid Flies, with the description of Two New Genera. Anais da Academia Brasileira de Ciencias, 39:195-210, 30

figures.

1967b. Studies on Indo-Australian Phorid Flies, Based

Mainly on Material of the Museum of Comparative Zoology and the United States National Museum, Part II. Studia Entomologia, 10:81-276,

177 figures.

1967c. Miscellaneous Studies on Phorid Flies, II. Revista

Brasileira de Biologia, 27:395-405, 22 figures.

1968. Miscellaneous Studies on Phorid Flies, III. Revista

Brasileira de Biologia, 28:167-176, 5 figures.

Borgmeier, T., and H. Schmitz

1923. Beitrag zur Kenntnis der Phoriden Brasiliens. Zeitschrift des Deutschen Vereins fuer Wissenschaft

und Kunst, 3(1922): 127-148, 2 figures, 1 plate.

Bridarolli, A.

1940. Cinco nuevos Phoridae argentinos del g6nero Megaselia. Revista de Entomologia, 11:487-495.

Brues, C. T.

1904. A Monograph of the North American Phoridae.

Transactions of the American Entomological Society, 29(1903) :331-404, plates V-IX.

1906. Family Phoridae. In Wytsman, Genera Insectorum, fasc. 44:1-22, 2 color plates.

1912. Synonymical Notes on Phoridae. Psyche, 19:

135-136.

1915a. A Synonymic Catalogue of the Dipterous Family

Phoridae. Bulletin of the Wisconsin Natural History Society, 12(1914):85-152.
NUMBER 2 3 51

1915b. A New Wingless Phorid Fly from Jamaica. Psyche,

22: 102-104.

1919a. The Phoridae of Grenada. Bulletin of the Museum of Comparative Zoology, 62:492-506, 2 figures.

1919b. Phoridae. Pages 431-433, in Johnson, A Revised

List of the Diptera of Jamaica. Bulletin of the

American Museum of Natural History, 41:421-

449.

1923. A New Genus of Myrmecophilous Phoridae, With

Notes on Some Related Forms. Psyche, 30:18-22

1 figure.

1924. New and Unrecorded Species of the Family Phoridae. Psyche, 31:155-161.

Coquillett,

1900. Report on a Collection of Dipterous Insects from

Puerto Rico. Proceedings of the U.S. National

Museum, 22:249-270.

Lehmann, J. G. C.

1822. Observationes Zoologicae Praesertim in Faunam

Hamburgensem. Pugillus primus. Indicem Scholarum Publice Privatimque in Hamburgensium

Gymnasio Academico. 55 pages. Hamburg,

Germany.

Enderlein, G.

1912. Die Phoridenfauna Sued-Brasiliens. Stettiner

Etomologische Zeitung, 73:16-45, 2 figures.

Lioy, P.

1864. I ditteri distributi secondo un nuovo metodo di

classificazione naturale (conclusion). Atti Istituto Veneto Scienze, Lettere ed Arti, serie III,

10:59-84.

Loew,

1866. Diptera Americae septentrionalis indigena, Centuria septima. Berlin Entomoligische Zeitschrift,

(1866):1-54.

Lundbeck, W.

1922. Diptera Danica, Part VI: Pipunculidae, Phoridae.

Copenhagen, 447 pages, 131 figures (Phoridae pp.

69-447).

Malloch, J. R.

1910. Scottish Phoridae, with Tables of All the British

Species and Notes of Localities. Annals of Scottish

Natural History, 1910:15-21, 87-92.

1912. The Insects of the Dipterous Family Phordiae in

the United States National Museum. Proceedings

of the United States National Museum, 43:411-

529, 7 plates.

1914. Costa Rican Diptera, Paper I: A partial Report

on the Borboridae, Phoridae and Agromyzidae.

Transactions of the American Entomological Society, 41:8-36, 1 plate (Phoridae pp. 24-35).

1924. A New Species of Conicera from Porto Rico.

Proceedings of the Entomological Society of Washington, 26:73-74.

Schmitz, H.

1923. Typenstudien an Phoriden. Jaarb. Nat. Gen.

Limb. 1920-1923; 49-59.

1926. Neue Gattungen und Arten europaeischer Phoriden. Encycl. Ent., series B II, Diptera 2(1925):

73-85, 2 figures.

1927. Revision der Phoridengattungen, mit Beschreibung

neuer Gattungen und Arten. Natuurhistorisch

Maandblad, 16:59-68.

1929. Revision der Phoriden. Berlin und Bonn: F.

Duemmler. 211 pages, 49 figures, 2 plates.

1938-1958. Family Phoridae. In Lindner, Die Fliegen

der Palaearktischen Region. 512 pages, 285 figures,

10 plates. Fortgesetzt von E. Beyer.

Silvestri, F.

1911. Con tribute alia conoscenza dei mirmecofili del

Messico. Bollentino del Laboratorio di Zoologia,

Portici 5:172-195, 17 figures (Phoridae pp.

172-178, figs. 1-5).

Wood, J. H.

1910. On the British Species of Phora. The Entomologist's Monthly Magazine, 46, 149-154, 195-202,

243-249.

1912. On the British Species of Phora. The Entomologist's Monthly Magazine, 48:94-99, 166-181, 3

figures, 1 plate.
52

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

FIGURES 1-11.—Dohrniphora cavifemur, new species: 1, abdomen of female; 2, hind femur of

female. D. divaricata, 3, male, base of hind femur. D. dispar, 4, male, base of hind femur.

D. cornuta: 5, male, base of hind femur. Coniceromyia pilicoxa, new species: 6, wing of male;

7, male, foretibia and metatarsus. C. ciliatipes, new species: 8, foretarsus of male; 9, wing of

male paratype; 10, forefemur of male. C. latimana: 11, foretarsus of male. [Drawn by T.

Borgmeier.J
NUMBER 23 53

18

FIGURES 12-20.—Conicera (Hypocerina) seticerca, new species: 12, left forceps; 13, right

forceps; 14, antenna of male. Megaselia (Aphiochaeta) fasciiventris: 15, wing of male from

Nova Teutonia; 16-17, hypopygium, lateral. M. (A.) luteicauda: 18, abdomen of female; 19,

wing of male; 20, hypopygium. [Drawn by T. Borgmeier ]
54 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

FIGURES 21—33.—Megaselia (Aphiochaeta) spiculata, new species: 21, wing of holotype; 22,

hypopygium. M. (A.) pleurochaeta, new species: 23, wing. M. (A.) punctifrons, new species:

24, wing. M. (A.) parviseta, new species: 25, wing; 26, hypopygium. M. (A.) destituta, new

species: 27, wing. M. (A.) pulliclava, new species: 28, palpus, dorsal; 29, palpus, lateral; 30,

wing. M. (A.) definita, new species: 31, wing of male; 32, hypopygium; 33, foretibia and

tarsus. [Drawn by T. Borgmeier.]
NUMBER 23 55

37

FIGURES 34-43.—Megaselia (Megaselia) depnvata, new species: 34, wing. M. (M.) elegantula,

new species: 35, wing. M. (M.) obscuripalpis, new species: 36, wing; 37, tergites 5-6. Af. (M.)

aurea: 38, hypopygium; 39, abdomen of female; 40, hind femur of male. M. (M.) hypochaeta,

new species: 41, abdomen, dorsal; 42, abdomen, ventral; 43, wing. [Drawn by T. Borgmeier.]
56

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

53

FIGURES 44—55.—Megaselia (Megaseiia) leptacina, new species: 44, abdomen; 45, wing; 46,

hind femur. M. (Af.) imitatrix, new species: 47, wing; 48, abdomen. M. (M.) buccata, new

species: 49, wing; 50, proboscis; 51, hypopygium. M. (M.) polita: 52, foretibia and tarsus of

type; 53, wing of female holotype. M. (A/.) femoralis: 54, hypopygium; 55, hypandrium with

ventral plates. [Drawn by T. Borgmeier.]
NUMBER 23 57

FIGURES 56—67.—Megaselia (Megaselia) enderleini: 56, wing of holotype male; 57, wing of

male from Dominica. M. (M.) labellifera, new species: 58, wing; 59, hypopygium; 60, hind

femur. M. (A/.) tarsodes, new species: 61, wing of holotype; 62, fore tibia and tarsus; 63,

palpus. M. (M.) metatarsalis, new species: 64, wing; 65, hind femur; 66, hypopygium; 67,

foretibia and tarsus. [Drawn by T. Borgmeier.]
58

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

75

11

FIGURES 68—77.—Megaselia (Megaselia) substricta, new species: 68, wing; 69, abdomen. M.

(A/.) mendax: 70, wing of paratype. M. (M.) diplochaeta, new species: 71, hypopygium; 72,

wing of holotype; 73, hind femur. M. (M.) horticola: 74, wing of female paratype. M. (M.)

gracilipalpis, new species: 75, wing; 76, hypopygium; 77, palpus. [Drawn by T. Borgmeier.]
NUMBER 2 3 59

FIGURES 78-85.—Megaselia (Megaselia) pallidipennis, new species: 78, frons; 79, wing; 80,

hind femur. M. (Af.) ferruginosa: 81, wing of male from Nova Teutonia, Brazil. M. (Af.)

bursella, new species: 82, wing; 83, hypopygium and sixth abdominal segment, lateral. M. (Af.)

burselloides, new species: 84, wing; 85, hypopygium and sixth abdominal segment, lateral.

[Drawn by T. Borgmeier.]
60

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

FIGURES 86-97.—Megaselia (Megaselia) fortirostris, new species: 86, wing; 87, abdominal segments 4-10, lateral; 88, proboscis of female. M. (M.) tumidirostris, new species: 89, wing;

90, proboscis of female; 91, tergites 5-6. M. (M.) hamaticauda, new species: 92, wing of male;

93, terminal segments of female abdomen, lateral. M. (M.) serotina: 94, wing of holotype male.

M. (Af.) paula, new species: 95, wing of holotype male; 96, midtarsus of male. M. (M.) miristigma: 97, wing of male from Porto Rico. [Drawn by T. Borgmeier.]
NUMBER 2 3 61

c

\ I

FIGURES 98-109.—Megaselia (Megaselia) flavopleura: 98, hypopygium; 99, tergites 5 and 6.

M. (M.) crindlifemur, new species: 100, hind femur; 101, wing of male; 102, abdomen of female;

103, hypopygium; 104, hypandrium and ventral plates. M. (M.) subaristalis, new species: 105,

antenna of male; 106, wing of male; 107, hypandrium and ventral plates; 108, hypopygium;

109, hind of femur of male. [Drawn by T. Borgmeier.]
62

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

117

114

FIOURES 110-120.—Megaselia {Megaselia) totaflava, new species: 110, abdomen of female; 111,

wing. M. (Af.) alajuelensis: 112, abdomen of female; 113, hypopygium; 114, proboscis of

female. Af. (Af.) mutilata, new species: 115, wing of female holotype. M. (Af.) crimpyga,

new species: 116, hypopygium; 117, wing of male. Woodiphora (Woodiphora) magnipalpis:

118, palpus of female; 119, palpus of male; 120, forefemur of male. [Drawn by T. Borgmeier.]
NUMBER 2 3 63

121

122

124

125

FIGURES 121-129.—Johowia parvicornis, new species: 121, wing of male; 122, palpus of male;

123, antenna of male. Apocephalus (Mesophora) insulanus, new species: 124, wing of male;

125, palpus of male; 126, hypopygium; 127, antenna of male. A. (M.) antennatus: 128, wing of

male; 129, hypopygium. [Drawn by T. Borgmeier.]
64

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

140

FIGURES 130-142.—Metopina recurvata, new species: 130, abdomen of female; 131, wing of

female; 132, wing of male. Pseudacteon arcuatus, new species: 133, ovipositor, dorsal. Dahliphora dispar: 134, wing of female; 135, wing of male; 136, antenna of female. Pachyneurella

bicolor: 137, hypopygium from left; 138, hypopygium from right. P. haplopyga, new species:

139, hypopygium, lateral. P. venata: 140, wing of male; 141, hypopygium, dorsal; 142, hypopygium, lateral. [Drawn by T. Borgmeier.]
NUMBER 2 3 65

FIGURES 143-152.—Homalophora sensibUis, new species: forefemur of male; 144, base of hind

femur; 145, hypopygium from right: 146, hypopygium from left. Chonocephalus furcatus: 147,

fingerlike structures of hypandrium; 148, hypopygium. C. dominicanus: 149, hypopygium

[i-internal copulatory organ]. Beckerina dactylata, new species: 150, wing of male; 151, hypopygium. B. dominie ana, new species: 152, wing of male. [Drawn by T. Borgmeier]
INDEX

[Names of new species in italics, page numbers of principal accounts in boldface.]

alajuelensis, Megaselia, 8, 34, 36, 62 (figs.)

aliena, Beckerina, 48

angustinervis, Apocephalus, 41

antennatus, Apocephalus, 40

antcrospinosa, Megaselia, 22

anthracina, Megaselia, 23

antiguensis, Pseudacteon, 41

Aphiochaeta, 7

caribbea, 9

fuscopedunculata, 37

opaciventris, 38

pilosa, 38

rubriventris, 18

velutinipes, 38

Apocephalus, 40

angustinervis, 41

antennatus, 40

aztecae, 40

instdanus, 40, 63 (figs.)

Apodicrania, 41

termitophia, 41

appendiculata, Megaselia, 32

arcuatus, Pseudacteon, 42, 64 (fig.)

atricolor, Coniceromyia, 4

atricolor, Megaselia, 24

attae, Homalophora, 45

aurea, Megaselia 7, 18, 20, 55 (figs.)

Azteca, 40

aztecae, Apocephalus, 40

Beckerina, 47, 48 (key)

Beckerina aliena, 48

burmicola, 48

chelifera, 48

costaricana, 48

dactylata, 48, 49, 65 (figs.)

dominicana, 48, 49, 65 (fig.)

fuscohalterata, 48

irregularis, 48

lucifrons, 47

luteihalterata, 48

luteola, 48

neotropica, 48

nigricornis, 48

nudipleura, 48

orphnephiloides, 48

polysticha, 48

relicta, 48

similata, 48

umbrimargo, 48

Beckerinini, 47

bicolor, Pachyneurella, 43, 44, 45

bihamulata, Megaselia, 32

brevicostalis, Megaselia, 8, 37, 38

buccata, Megaselia, 7, 22, 56 (figs.)

burmicola, Beckerina, 48

bursella, Megaselia, 8, 29, 59 (figs.)

burselloides, Megaselia, 8, 29, 30, 59 (figs.)

Calosoma, 6

caribbea, Aphiochaeta, 9

carinata, Megaselia, 32

cavifemuT, Dohrniphora, 1, 52 (figs.)

chelifera, Beckerina, 48

Chonocephalus, 46

dimakae, 46

dominicanus, 47, 65 (figs.)

dorsalis, 46

furcatus, 46, 65 (figs.)

hibisci, 46

jamaicensis, 47

madagascariensis, 46

mexicanus, 47

punctifascia, 46, 47

vadoni, 46

cUiatipes, Coniceromyia, 4, 52 (figs.)

coalescens, Megaselia, 16

communis, Megaselia, 25

Conicera, 5

megalodus, 5

neotropica, 5

seticerca, 5, 53 (figs.)

Coniceromyia, 3

atricolor, 4

ciliatipes, 4, 52 (figs.)

cubensis, 4

latimana, 5, 52 (fig.)

pilicoxa, 3, 52 (figs.)

striativena, 4

cornuta, Dohrniphora, 3, 52 (fig.)

costaricana, Beckerina, 48

crassirostris, Megaselia, 31

crenaticornis, Dahliphora, 43

crinellifemur, Megaselia, 8, 34, 61 (figs.)

crinipyga, Megaselia, 8, 37, 62 (figs.)

cubensis, Coniceromyia, 4

curriei, Pseudacteon, 42

dactylata, Beckerina, 48, 49, 65 (figs.)

Dahliphora, 43

crenaticornis, 43

dispar, 43, 64 (figs.)

definita, Megaselia, 7, 15, 54 (figs.)

66
NUMBE

R 2 3

67

depnvata, Mcgaselia, 7, 16, 55 (fig.)

destituta, Megaselia, 7, 13, 54 (fig.)

diffusa, Dohrniphora, 2

dimakae, Chonocephalus, 46

diplochaeta, Megaselia, 8, 26, 58 (figs.)

dispar, Dahliphora, 43, 64 (figs.)

Dohrniphora, 2, 52 (fig.)

disparisecta, Megaselia, 19

divaricata, Dohrniphora, 2, 52 (fig.)

Dohrniphora, 1

cavifemur, 1, 52 (figs.)

cornuta, 3, 52 (fig.)

diffusa, 2

dispar, 2, 52 (fig.)

divaricata, 2, 52 (fig.)

perplexa, 2

venusta, 2

dominicana, Beckerina, 48, 49, 65 (fig.)

dominicanus, Chonocephalus, 47, 65 (fig.)

dorsalis, Chonocephalus, 46

dorymyrmecis, Pseudacteon, 41

Dorymyrmex, 41

elegantula, Megaselia, 7, 18, 55 (fig.)

enderleini, Megaselia, 7, 23, 57 (figs.)

epichaeta, Homalophora, 45

fasciiventris, Megaselia, 6, 9, 53 (figs.)

femoralis, Megaselia, 7, 23, 56 (figs.)

fenyesi, Metopina, 43

ferruginea, Megaselia, 28

ferruginosa, Megaselia, 8, 28, 59 (fig)

fimbriata, Megaselia, 19

flavipes, Megaselia, 25

flavopleura, Megaselia, 8, 33, 36, 61 (figs.)

formicarum, Pseudacteon, 42

fortirostris, Megaselia, 8, 30, 60 (figs.)

frequens, Megaselia, 37

furcatus, Chonocephalus, 46, 65 (figs.)

fuscohalterata, Beckerina, 48

fuscopedunculata, Aphiochaeta, 37

gemella, Megaselia, 7, 12, 14

gracilipalpis, Megaselia, 8, 27, 58 (figs.)

Gymnoptera, 6

neotropica, 6

hamaticauda, Megaselia, 8, 31, 60 (figs.)

haplopyga, Pachyneurella, 44, 45, 64 (fig.)

hibisci, Chonocephalus, 46

Homalophora, 45

attae, 45

epichaeta, 45

reichenspergeri, 45

scnsibilis, 45, 65 (figs.)

horticola, Megaselia, 8, 27

Hypocerina, 5

hypochaeta, Megaselia, 7, 20, 55 (figs.)

imitatrix, Megaselia, 7, 21, 56 (figs.)

insulanus, Apocephalus, 40, 63 (figs.)

irregularis, Beckerina, 48

jamaicensis, Chonocephalus, 47

Johowia, 39

parvicornis, 39, 63 (figs.)

pilipleura, 40

labellata, Megaselia, 24

labellifera, Megaselia, 8, 24, 57 (figs.)

lanceata, Megaselia, 14

latimana, Coniceromyia, 5, 52 (fig.)

leptacina, Megaselia, 7, 21, 56 (figs.)

littoralis, Megaselia, 23

lucifrons, Beckerina, 47

lutea, Megaselia, 33

luteicauda, Megaselia, 6, 10, 53 (figs.)

luteihalterata, Beckerina, 48

luteoides, Megaselia, 33

luteola, Beckerina, 48

macrochaeta, Megaselia, 11

madagascariensis, Chonocephalus, 46

magnipalpis, Woodiphora, 38,62 (figs.)

malacophaga, Megaselia, 16

megalodus, Conicera, 5

Megaselia, 7 (key)

alajuelensis, 8, 34, 36, 62 (figs.)

anterospinosa, 22

anthrancina, 23

appendiculata, 32

atricolor, 24

aurea, 7, 18, 20, 55 (figs.)

bihamulata, 32

brevicostalis, 8, 37, 38

buccata, 7, 22, 56 (figs.)

bursella, 8, 29, 59 (figs.)

burselloides, 8, 29, 30, 59 (figs.)

carinata, 32

coalescens, 16

communis, 25

crassirostris, 31

crinellifemur, 8, 34, 61 (figs.)

crinipyga, 8, 37, 62 (figs.)

definite, 7, 15, 54 (figs.)

deprivata, 7, 16, 55 (fig.)

destituta, 7, 13, 54 (fig.)

diplochaeta, 8, 26, 58 (figs.)

disparisecta, 19

elegantula, 7, 18, 55 (fig.)

enderleini, 7, 23, 57 (figs.)

fasciiventris, 6, 9, 53 (figs.)

femoralis, 7, 23, 56 (figs.)

ferruginea, 28

ferruginosa, 8, 28, 59 (fig.)

fimbriata, 19

flavipes, 25

flavopleura, 8, 33, 36, 61 (figs.)

fortirostris, 8, 30, 60 (figs.)

frequens, 37

gemella, 7, 12, 14

gracilipalpis, 8, 27, 58 (figs.)

hamaticauda, 8, 31, 60 (figs.)

horticola, 8, 27

hypochaeta, 7, 20, 55 (figs.)
6

8 INDE

X

Megasclia—Continued

imitatrix, 7, 21, 56 (figs.)

labellata, 24

labellifera, 8, 24, 57 (figs.)

lanccata, 14

Uptacina, 7, 21, 56 (figs.)

littoralis, 23

lutea, 33

luteicauda, 6, 10, 53 (figs.)

lutcoides, 33

macrochacta, 11

malacophaga, 16

mendax, 26

rnetatarsalts, 8, 24, 57 (figs.)

minuta, 23

miristigma, 8, 31, 60 (fig.)

mutilate, 8, 37, 62 (fig.)

obscura, 27

obscuripalpis, 7, 17, 55 (figs.)

orthoneura, 32

pallidipennis, 8, 28, 59 (figs.)

parvheta, 7, 12, 13, 54 (figs.)

Paula, 8, 32, 60 (figs.)

picta, 7, 16

pleurochaeta, 6, 11, 54 (fig.)

plcurofascia, 7, 20

polita, 7, 23, 56 (figs.)

postorta, 33

pteryacantha, 11

pulliclava, 7, 14, 54 (figs.)

punctifrons, 6, 12, 54 (fig.)

rubronigra, 19

scalaris, 7, 17, 22

scutellaris, 33

scmifcrruginca, 7, 12

scrotina, 33

scticauda, 7, 19

sctigera, 6, 9

spiculata, 6, 11, 13, 54 (figs.)

striativentris, 7, 19

subaristalts, 8, 35, 61 (figs.)

subflava, 17, 35

sublutea, 15

subpicta, 7, 16

substricta, 8, 25, 58 (figs.)

tarsodes, 8, 25, 57 (figs.)

tetrachaeta, 33

totaflava, 8, 35, 62 (figs.)

nibulifera, 21

tumidicornis, 35

tumidirostris, 8, 30, 31, 60 (figs.)

tumidula, 14

mendax, Mcgaselia, 26

Mcsophora, 40

mctac, Homalophora, 45

metatarsalis, Mcgaselia, 8, 24, 57 (figs.)

Metopina, 42

recurvata, 42, 64 (figs.)

Metopininae, 7

Metopinini, 7

mcxicanus, Chonoccphalus, 47

minuta, Megaselia, 23

miristigma, Megaselia, 8, 31, 60 (fig.)

mutilata, Megaselia, 8, 37, 62 (fig.)

ncotropica, Beckerina, 48

Conicera, 5

Gymnoptera, 6

nigricornis, Beckerina, 48

nitida, Pachyneurella, 44

nudipleura, Beckerina, 48

obscura, Megaselia, 27

obscuripalpis, Megaselia, 7, 17, 55 (figs.)

opaciventris, Aphiochacta, 38

orphnephiloides, Beckerina, 48

orthoneura, Megaselia, 32

Pachyneurella, 43

bicolor, 43, 44, 45

haplopyga, 44, 45, 64 (fig.)

nitida, 44

pallidipennis, Megaselia, 8, 28, 59 (figs.)

parvxeornis, Johowia, 39, 63 (figs.)

parviseta, Megaselia, 7, 12, 13, 54 (figs.)

paula, Megaselia, 8, 32, 60 (figs.)

perplcxa, Dohrniphora, 2

Phorinac, 1

Physoptera punctifemur, 17

picta, Megaselia, 7, 16

pilicoxa, Coniceromyia, 3, 52 (figs.)

pilipleura, Johowia, 40

pilosa, Aphiochaeta, 38

pleurochaeta, Megaselia, 6, 11, 54 (fig.)

pleurofascia, Megaselia, 7, 20

polita, Megaselia, 7, 23, 56 (figs.)

polysticha, Beckerina, 48

postorta, Megaselia, 33

pradei, Pseudacteon, 42

Pseudacteon, 41

antiguensis, 41

arcuatus, 42, 64 (fig.)

curriei, 42

dorymyrmecis, 41

formicarum, 42

pradei, 42

pteryacantha, Megaselia, 11

Puliciphora, 43

pulliclava, Megaselia, 7, 14, 54 (figs.)

punctifascia, Chonoccphalus, 46, 47

punctifemur, Physoptera, 17

punctifrons, Megaselia, 6, 12, 54 (fig.)

recurvata, Metopina, 42, 64 (figs.)

reichenspergeri, Homalophora, 45

relicta, Beckerina, 48

rubriventris, Aphiochaeta, 18

rubronigra, Megaselia, 19

scalaris, Megaselia, 7, 17, 22

scutellaris, Megaselia, 33

semiferruginea, Megasclia, 7, 12

sensibilis, Homalophora, 45, 65 (figs.)

scrotina, Megaselia, 33

seticauda, Megasclia, 7, 19
NUMBER 2 3

seticerca, Conicera, 5, 53 (figs.)

setigcra, Mcgasdia, 6, 9

similata, Beckerina, 48

Solcnopsis, 41

spiculata, Megaselia, 6, 11, 13, 54 (figs.)

striativena, Coniccromyia, 4

striativcntris, Megaselia, 7, 19

subaristalis, Megaselia, 8, 35, 61 (figs.)

subflava, Megaselia, 17, 35

sublutea, Megaselia, 15

subpicta, Megaselia, 7, 16

substricta, Megaselia, 8, 25, 58 (figs.)

tarsodes, Megaselia, 8, 25, 57 (figs.)

Termitometoecus, 43

69

termitophila, Apodicrania, 41

tetrachaeta, Megaselia, 33

totaflava, Megaselia, 8, 35, 62 (figs.)

tubulifera, Megaselia, 21

tumidicornis, Megaselia, 35

tumidirostris, Megaselia, 8, 30, 31, 60 (figs.)

tumidula, Megaselia, 14

umbrimargo, Beckerina, 48

vadoni, Chonocephalus, 46

velutinipes, Aphiochaeta, 38

venata, Pachyneurella, 43, 44, 64 (figs.)

venusta, Dohrniphora, 2

Woodiphora, 38

magnipalpis, 38, 62 (figs.)

U^. SOVERNMENT f»«NTIN« OFFICE: 1988 O S38-709

Publication in Smithsonian Contributions to Zoology

Manuscripts for serial publications are accepted by the Smithsonian Institution Press, subject

to substantive review, only through departments of the various Smithsonian museums. NonSmithsonian authors should address inquiries to the appropriate department. If submission is

invited, the following format requirements of the Press will govern the preparation of copy.

(An instruction sheet for the preparation of illustrations is available from the Press on request.)

Copy must be typewritten, double-spaced, on one side of standard white bond paper, with

1/4" top and left margins, submitted in ribbon copy with a carbon or duplicate, and accompanied

by the original artwork. Duplicate copies of all material, including illustrations, should be retained

by the author. There may be several paragraphs to a page, but each page should begin with a new

paragraph. Number consecutively all pages, including title page, abstract, text, literature cited,

legends, and tables. The minimum length is 30 pages of typescript and illustrations.

The title should be complete and clear for easy indexing by abstracting services. Taxonomic

titles will carry a final line indicating the higher categories to which the taxon is referable:

"(Hymenoptera: Sphecidae)." Include an abstract as an introductory part of the text. Identify

the author on the first page of text with an unnumbered footnote that includes his professional

mailing address. A table of contents is optional. An index, if required, may be supplied by the

author when he returns page proof.

Two headings are used: (1) text heads (boldface in print) for major sections and chapters

and (2) paragraph sideheads (caps and small caps in print) for subdivisions. Further headings

may be worked out with the editor.

In taxonomic keys, number only the first item of each couplet; if there is only one couplet,

omit the number. For easy reference, number also the taxa and their corresponding headings

throughout the text; do not incorporate page references in the key.

In synonymy, use the short form (taxon, author, date, page) with a full reference at the end

of the paper under "Literature Cited." Begin each taxon at the left margin with subsequent lines

intended about three spaces. Within a taxon, use a period-dash (.—) to separate each reference.

Enclose with square brackets any annotation in or at the end of the taxon. For references within

the text, use the author-date system: "(Jones, 1910)" or "Jones (1910)." If the reference is

expanded, abbreviate the data: "Jones (1910, p. 122, pi. 20: fig. 1)."

Simple tabulations in the text (e.g., columns of data) may carry headings or not, but they

should not contain rules. Formal tables must be submitted as pages separate from the text, and

each table, no matter how large, should be pasted up as a single sheet of copy.

For measurements and weights, use the metric system instead of (or in addition to) the

English system.

Illustrations (line drawings, maps, photographs, shaded drawings) can be intermixed

throughout the printed text. They will be termed Figures and should be numbered consecutively;

however, if a group of figures is treated as a single figure, the individual components should be

indicated by lowercase italic letters on the illustration, in the legend, and in text references:

"Figure 9&." If illustrations (usually tone photographs) are printed separately from the text as

full pages on a different stock of paper, they will be termed Plates, and individual components

should be lettered (Plate 96) but may be numbered (Plate 9: figure 2). Never combine the

numbering system of text illustrations with that of plate illustrations. Submit all legends on pages

separate from the text and not attached to the artwork.

In the bibliography (usually called "Literature Cited"), spell out book; journal, and article

titles, using initial caps with all words except minor terms such as "and, of, the." (For capitalization of titles in foreign languages, follow the national practice of each language.) Underscore

(for italics) book and journal titles. Use the colon-parentheses system for volume, number, and

page citations: "10(2) :5-9." Spell out such words as "figures" and "plates" (or "pages" when

used alone).

For free copies of his own paper, a Smithsonian author should indicate his requirements on

"Form 36" (submitted to the Press with the manuscript). A non-Smithsonian author will receive

50 free copies; order forms for quantities above this amount with instructions for payment will be

supplied when page proof is forwarded.