Checklist of North American 

Late Paleozoic Coral Species 

(Coelenterata, Anthozoa) 

By WILLIAM J. SANDO 

GEOLOGICAL SURVEY BULLETIN 

A bibliographic compilation through 1973 

of coral species described from the 

Mississippian, Pennsylvanian, and 

Permian rocks of North America 

1 3 8 7 

UNITED STATES GOVERNMENT PRINTING OFFICE, WASHINGTON 1974 
UNITED STATES DEPARTMENT OF THE INTERIOR 

ROGERS C. B. MORTON, Secretary 

GEOLOGICAL SURVEY 

V. E. McKelvey, Director 

Library of Congress Cataloging in Publication Data 

Sando, William Jasper. 

Checklist of North American late Paleozoic coral species (Coelenterata, 

Anthozoa) 

(Geological Survey bulletin; 1387) 

Includes bibliography. 

Supt. of Docs. no.: I 19.3:1387 

1. Anthozoa, Fossil. 2. Paleontology-Paleozoic. 3. PaletOntology-North 

America. I. Title. II. Series: United States. Geological Survey. Bulletin; 

1387. 

QE75.B9 no. 1387 [QE778] 557.3'08s [56,3.'.6] 74-23375 

For sale by the Superintendent of Documents, U.S. Government Printing Office 

Washington, D.C. 20402 -Price 70 cents (paper cover) 

Stock Number 2401-Q2576 
CONTENTS 

Pag.e · 

Abstract ------------------------------------------------------- 1 

Introduction ---------------------------------------------------- 1 

Species list ------------------------------ _______________ -------- 2 

References cited ------------------------------------------------ 26 

III 

CHECKLIST OF NORTH AMERICAN 

LATE PALEOZOIC CORAL SPECIES 

(COELENTERATA, ANTHOZOA) 

By WILLIAM J. SANDO 

ABSTRACT 

An objective list of coral species described from the Mississippian, Pennsylvanian, and Permian rocks of North Ame~ica includes 578 entries, of 

which four are now known to be from older rocks and four are probably 

erroneous references to older species. The list includes information on the 

original references for all species and the geologic age and geographic area 

from which the species have been described. 

INTRODUCTION 

This paper presents an objective list of formally named coral 

species that have been described from rocks of Mississippian, 

Pennsylvanian, and Permian age in North America through 1973. 

Comparative species references (cf., aff.) and unnamed species 

(sp.) are excluded. One nomen nudum (Texanophyllum skinneri) 

is included. The list does not include chaetetids, which I regard as 

hydrozoans. 

The list is arranged alphabetically by species name in the form 

in which this name was originally published, together with the 

author, original date of publication, and original generic assignment. Secondary references are indicated for species whose type 

localities are outside North America. All original and secondary 

references are included in the bibliography at the end of the 

paper. Subjective synonymy is omitted; objective synonyms are indicated only for replacernent names. Names proposed for varieties, 

subspecies, and formae are treated as species names. 

Age designations have been updated to indicate the latest interpretations. A few species originally described as late Paleozoic 

whose type specimens were later found to be from older rocks 

are included with appropriate explanation. The list also includes 

1 
2 NORTH AMERICAN LATE PALEOZOIC CORAL SPECIES 

a few corals that have been referred to pre-Mississippian species 

probably erroneously but without published verification of erroneous taxonomic assignment. 

Geographic designations are in standard political units for 

North America (Canada, Mexico, and the United States of 

America). Each geographic designation includes the original 

State or Province from which the species was described and other 

areas from which specimens assigned to the species have been described or illustrated. Geographic extensions of the usage of names 

are not documented in the bibliography; only references to original descriptions are given. Queried species assignments are included, but comparative species references (cf., aff.) are omitted. 

The nucleus for this compilation was the late Paleozoic species 

synonymy file in the U.S. Geological Survey, (U.S. National Museum) Washington, D.C. This file was begun by G. H. Girty and 

maintained continuous,ly until about 1957, after which the coverage became sporadic. Other compilations that were useful in 

searching for and checking species references are Bassler (1950), 

Branson (1948), Weller (1898), and Wilson (1974). 

I am indebted to A. K. Armstrong, E. W. Bamber, J. M. Cocke, 

C. L. Rowett, E. C. Wilson, and Geoffrey Cotton for checking preliminary versions of the list. Their expertise was valuable in assuring maximum accuracy and completeness for the compilation. 

I am also grateful to E. L. Yochelson and W. A. Oliver, Jr., for 

reviewing the final manuscript. 

Perfection is rarely, if ever, attained in any work of this kind. 

I would be grateful if any errors or omissions were called to 

my attention. 

SPECIES LIST 

[Miss.: Mississippian: Penn., Pennsylvanian; Perm., Permian] 

aages Jeffords 1947, Stereostylus 

Penn.: Iowa 

absitus Jeffords 1947, Stereostylus 

Penn. : Kansas 

acolumnatum Cocke 1970, Neokoninckophyllum 

Penn.: Kansas, Oklahoma 

aculeata Girty 1899, Syringopora 

Miss.: Alberta, Arizona, Colorado, Montana, New Mexico, Utah, 

Wyoming 

acus Moore and Jeffords 1941, Leonardophyllum 

Perm. : Texas 

acutus White and Whitfield, 1862, Zaphrentis 

Miss.: Iowa, Missouri 
SPECIES LIST 

adapertum Mo-ore and Jeffords, 1945, Lophophyllidium 

Penn. : Texas 

adelus Jeffords 1947, Stereostylus 

Penn. : Texas 

a.djunctiva White 1880a, Acervularia 

Miss.: Idaho 

adnatus Easton 1945c, Amplexus 

Miss. : Illinois 

agglomerata Grabau in Greene 1901a, Cera.topora 

Miss. : Indiana 

agglomerata Nelson 1962a, Syringopora 

Miss. : Alberta 

3 

aggregatum Simpson 1900, Schoenophyllum (equals Lithostrotion [Siphonodendron] genevievensis Easton 1957) 

Miss.: Kentucky (Easton, 1957) 

alani Cocke and Bowsher 1968, Sutherlandia 

Penn. : Oklahoma 

alaskaensis Armstrong 1970, Sciophyllmn 

Miss. : Alaska 

alaskaensis Rowett 1969, Durhamina 

Perm. : Alaska 

alleni Rowley 1901, AxophyllU'm? 

Penn. : Missouri 

americana Hayasaka 1936, Lithostrotionella 

Miss.: Alaska, Alberta, California, Illinois, Kentucky, Missouri (The 

California specimen is probably Permian) 

americana Weller, 1909, Beamnontia 

Miss. : Missouri 

americanus Weller 1909, Cladochonus 

Miss.: Arkansas, Missouri 

amesensis Bebout 1966, Stereostylus 

Penn.: Ohio 

a1noenum Moore and Jeffords 1945, Lophotichium 

Penn. : Oklahoma 

amplexa Rowley 1901, Aulopora 

Miss. : Missouri 

amsdenensis Branson and Greger 1918, Zaphrentis 

Miss.: Wyoming 

angustifolium Moore and Jeffords 1945, Lophophyllidiurn 

Penn.: Arkansas, Oklahoma 

anna Beede 1898, Aulopora.? 

Penn. : Kansas 

annae Jeffords 1947, Stereostylus 

Penn. : Kansas 

annectans Moore and Jeffords 1945, Stereoc,orypha 

Penn. : Texas 

aplatus Cummins 1891, Hadrophyllum 

Penn. : Texas 

araneosum Parks 1951, Faberophyllum 

Miss.: Utah 
4 NORTH AMERICAN LATE PALEOZOIC CORAL SPECIES 

arc.himediformis Rowley 1905, Amplexus 

Miss. : Missouri 

arcturusensis Stevens 1967, Kleopah·ina (Porfirievella) 

Perm.: Nevada 

arcuata Jeffords 1943, Caninia 

Miss. : New Mexico 

arcuata Weller, 1909, Cyathaxonia 

Miss.: Arkans-as, Missouri, Montana 

arcuatum Moore and Jeffords 1945, N eokoninckophyllum 

Penn.: Arizona, Texas 

arizelum Crickmay 1955, Korninckophyllum 

Miss. : Alberta 

a'rkansanum Girty 1910, Menophyllum 

Miss. : Arkansas, Oklahoma 

a.~arcum Jeffords 1947, Lophophyllidium 

Penn. : Oklahoma 

astraeiforme Warren 1927, Diphyphyllum 

Miss.: Alaska, Alberta 

austini Worthen 1890, Trachypo·ra 

Penn. : Kansas 

avonensis Bell 1929, Lophophyllum 

Miss.: Nova Scotia 

bailliei Nelson 1960, Lithostrotionella 

Miss. : Alberta 

ba.nJjense Warren 1927, Clisiophyllum? 

Miss.: Alberta 

banffense Warren 1927, Lithostrotion 

Miss.: Alaska, Alberta 

basaltiforme Haughton in M'Clintock 1857, Lithostrotion 

Perm.: Northwest Territories (Arctic Islands) 

basaltiforme Owen 1852, Lithostrotion 

Miss.: Alabama, Iowa, Kentucky 

bassoi Nelson 1962a, Syringopora 

Miss.: Alberta, British Columbia 

beecheri Grabau 1899, Monilopora 

Miss.: Arkansas, Indiana 

belc,heri Harker 1960, Caninia 

Perm.: Northwest Territories (Arctic Islands) 

bennetti Beede 1898, Cladochonus 

Penn. : Kansas 

berthiaumi Merriam 1942, Lithostrotion (Lithostrotionella) 

Perm.: Oregon 

bicarinatus Easton 1951, Triplophyllites ( Triplophyllites) 

Miss. : Indiana 

bicostatus Miller 1891, Amplexus 

Miss.: Missouri 

bifidus Weller 1909, Palaeacis 

Miss.: Missouri, New Mexico 

bilateralis Easton 1945b, Caninia 

Miss.: Montana 
SPECIES LIST 

billingsi Dawson 1868, Cyathophyllum 

Miss. : Nova Scotia 

birdi Armstrong 1970, Lithostrotionella 

Miss.: Alaska 

blairi Miller 1891, Amplexus 

Miss.: Indiana, Iowa, Missouri 

blairi Miller, 1892, J."Wicrocyclus 

Miss.: Missouri, New Mexico 

blandum Moore and Jeffords 1945, Lophophyllidium 

Penn. : Oklahoma 

bollibokkensis Coogan 1960, Stereostylus 

Perm. : California 

bordeni Greene 1900, Cyathaxonia 

Miss. : Indiana 

borealis Stuckenberg 1895, Lithostrotion (Ghernys:hev and Stepa.nov, 1916) 

Perm.: Northwest Territories (Arctic Islands) 

bourbonense Cocke 1970, Dibunophyllum 

Penn. : Kansas 

branneri Miller and Gurley 1893, Mic.helinia 

Penn. : Illinois 

brevifolius Jeffords 194 7, Lophamplexus 

Penn. : Colorado, Kansas, Oklahoma 

brevis Weller 1909, Amplexus 

Miss.: Missouri 

brokawi Wilson and Langenheim 1962, Lithostrotionella 

Perm.: Nevada 

brucei Gocke 1966, Dibunophyll-um 

Penn. : Oklahoma 

brushensis Bebout 1966, Stereostylus 

Penn.: Ohio 

burlingi Sutherland 1958, Pseudoza.phrentoides? 

Miss.: British Columbia 

burlingtonensis Keyes 1942, Aulopora (equals A. gra,cilis Keyes, 1894) 

Miss.: Iowa, Missouri 

buttensis Stevens 1967, HeTitschioides (Heritschioides) 

Perm.: Nevada 

calcariformis Hall 1882, Zaphrentis 

Miss.: Alabama, Alberta, Indiana, Kentucky 

calceola White and Whitfield 1862, Lophophyllum 

Miss.: Arizona, Iowa, Missouri, New Mexico 

californicus Tischle,r 1956, Triplophyllites ( Triplophyllites) 

Miss.: California 

californiense Langenheim and Tischler, 1960, Enygmophyllum 

Miss.: California 

californiense Meek 1864, Lithostrotion? 

Perm.: California 

callosum Moore and Jeffords 1945, Barytichisma 

Penn.: Arkansas, Oklahoma, Texas 

ca,lyculus Miller 1891, Zaphrentis 

Miss.: Missouri 

5 
6 NORTH AMERICAN LATE PAL·EOZOIC CORAL SPECIES 

canadens1's Gastelnau 1843, Axinura 

(Type sped men is probably a drift specimen derived from Devonian 

(Ehlers in Lang, Smith, and Thomas, 1940). As commonly used it 

equals Acrocyathus floriforrnis d'Orbigny 1849 (Easton, 1973)) 

Miss.: Alabama, Illinois, Kentucky, Missouri, Tennessee, Virginia 

caneyana Morgan 1924, Pachypora 

Penn.: Oklahoma 

cannonensis Winchell 1869, Zaphrentis 

Miss.?: Tenness·ee 

captiosis Moore and Jeffords 1945, Lophamplexus 

Penn.: Oklahoma 

capulifoormis Rowley 1900, Zaphrentis 

Miss. : Mis•souri 

carbonaria Mather 1915, Pachypora 

Penn.: Arkansas, Oklahoma 

carbonaria White 1862, Striatopora 

Miss. : Iowa, Missouri 

carinata Girty, 1910, Palaeacis 

Miss.: Arkans·as 

carinatus Worthen 1890, Zaphrentis 

Miss.: Indiana, Iowa, Kentucky 

ca,scadense Warren 1927, Lophophyllum? 

Miss. : Alberta 

cassa Sutherland 1958, Amplexi-Zaphrentis 

Miss.: British Columbia 

cassedayi Milne-Eldwards 1860, Zaphrentis 

Miss.: Indiana, Kentucky, Ohio 

ca,stelnaui Hayasaka 1936, Lithostrotionella 

Miss.: Alabama, California, Georgia, Illinois, Missouri, Virginia, West 

Virginia 

casteri Armstrong 1958, Zaphriphyllum 

Miss. : New Mexico 

cavernosa Milier 1891, Pa1aeacis 

Miss. : Indi•ana 

cavum Easton 1944, Pseudocryptophyllum 

Miss.: Missouri 

centralis Milne-Edwards and Haime 1851, Zaphrentis 

Miss.: Illinois, Iowa, Kentucky, Missouri 

centrocylindrica Driscoll and Newton 1969, Syringopora. 

Penn.: Montana 

chesterensis Worthen 1890, ZaphrenUs 

Miss.: Illinois, Missouri 

clwuteauensis Miller 1891, Zaphrentis 

Miss.: Georgia, Indiana, Missouri 

circinatus Easton and Gutschick 1953, Lithostrotionella 

Miss. : Arizona 

C'irc,ularis Easton 1958, Triplophyllites (H omalophyllites) 

Miss. : New Mexico, Sonora 

clathrum Cocke 1970, Dibunophyllum 

Penn. : Kansas 
SPECIES LIST 

clavigerum Easton 1962, Bradyphyllurn 

Miss. and Penn.?: Montana 

clifjordana Milne-Edwards and Haime 1851, Zaphrentis 

Miss.: Alabama, Georgia, Illinois, Iowa, Kentucky, Missouri, Ohio 

clinatus Greene 1904b, Zaphrentis 

Miss.: Indiana, Kentucky, Missouri, New Mexico 

coaptum Moore and Jeffords 1945, Lophophyllidium 

Penn. : Arkansas 

columbicum Smith 1935, Waagenophyllum 

Perm.: British Columbia 

columnirotata Langenheim and Tischler 1960, Rylstonia 

Miss. : Arizona, California 

compac,ta Nelson 1962a, Syringopora 

Mis.s. : Alberta 

complexum Jeffords 1942, Lophophyllidium 

Penn.: Oklahoma 

compressa Easton 1943h, Palaeosmilia 

Miss. : Tennessee 

compressa Greene 1899a, Cyathaxonia (equals C. venusta Greene) 

Miss. : Indiana 

com,pressa Milne-Edwards 1860, Zaphrentis 

Miss.: Alabama, Georgia, Indiana, Kentucky, Tennessee, Utah 

compressum Jeffords 1942, Lophophyllidiurn 

Penn.: Texas 

compressum Meek and Worthen 1860, Sphenopoterium 

Miss. : Illinois 

confertum Jeffords 1942, Lophophyllidiwm 

Penn. : Oklahoma 

confluens Easton 1958, Lithostro,tionella 

Miss.: Arizona, New Mexico, Sonora 

conica Easton 1944, Palaeacis 

Miss. : Missouri 

conicum Castelnau 1843, Cyathophyllum 

Miss. : Illinois 

c,oniforme Jeffords 1947, Lophophyllidium 

Penn.: Kansas, Missouri, Oklahoma 

connorsensis Easton 1960, Lithostrotion [Diphyphyllum] 

Perm.: Nevada 

conoideum Moore and Jeffords 1945, Lophophyllidium 

Penn.: Arkansas, Texas 

contortus Easton 1962, Fasciculiarnplexus 

Penn.?: Montana 

cooperi Ross and Ross 1962, N eokoninckophyllum 

Penn. : Texas 

coralloides Sowerby 1814, Amplexus (Milne-Edwards and Haime, 1851) 

Miss.: Illinois, New Mexico 

cordillerensis Easton 1958, Cyathaxonia 

Miss.: Sonora 

cordillerensis Easton, 1960, Lonsdaleia 

Perm.: California, Nevada 

7 
8 NORTH AMERICAN LATE PALEOZOIC CORAL SPECIES 

co'tniculum Miller 1891, Amplexus 

Miss.: Georgia, Missouri, Sonora 

cornucopiae Michelin in Gervais 1840, Caninia (Sloss, 1945) 

Miss.: Montana 

corrugatus Mather 1915, Amplexus 

Penn.: Arizona, Arkansas, Oklahoma, Texas 

crassa McCoy 1844, Jania (Weller, 1909) 

Miss. : Missouri 

crassiseptatum Moore and Jeffords 1945, Hapsiphyllum 

Penn.: Oklahoma 

crassum Moore and Jeffords 1945, Barytichisma 

Penn. : Texas 

cumberlandica Troost 1840, Calamopora 

Miss. : Tennesse'e 

cuneata Snider 1915, Palaeacis 

Miss.: Arkansas, Oklahoma 

cuneatum Meek and Worthen 1860, Sphenopoterium 

Miss.: Indiana 

cuneiformis Milne-Edwards 1860, Palaeacis 

Miss. : Alabama, Indiana 

curva Shimer 1926, Aulopora 

Miss. : Alberta 

cylindraceus Worthen 1890, Zaphrenf'is 

Miss.: Illinois, Missouri 

cylindrica Girty 1908, Lindst1·oemia 

Perm. : Texas 

cylindrica Scouler in McCoy 1844, Siphonophyllia (Marcou, 1858) 

Miss.: New Mexico 

cylincrricmn Dobrolyubova and Kabakovich 1948, Axophyllum (Cocke, 1970) 

Penn. : Kansas 

cylindricum Girty 1915, Axophyllum 

Penn. : Missouri 

cymba Seebach 1866, Palaeacis 

Miss.: Iowa 

cyndon Rafinesque and Clifford 1820, Turbinolia 

Miss.: Kentucky, Pennsylvania (Pennsylvania record is probably Pennsylvanian) 

cystoides Grabau in Greene 1901a, Romingeria 

Miss. : Indiana 

cystosum Gocke 1970, Neokoninckophyllum 

Penn. : Kansas 

dalii Milne-Edwards and Haime 1851, Zaphrentis 

Miss.: Alberta, Georgia, Illinois, Indiana, Iowa, Missouri 

dawsoni Lambe 1899a, Cyathophyllum 

Miss.: Nova Scotia 

deciensis Ross and Ross 1962, N eokmtinckophyllum 

Perm. : Texas 

declinis Miller 1891, Zaphrentis 

Miss. : Indiana 

delicata Ross and Ross 1963, Amplexocarinia 

Penn. and Perm. : Texas 
SPECIES LIST 

delicatum Bassler, 1937, Hadrophyllum 

Miss.: Kentucky 

deltense Rowett 1969, Auloclisia 

Perm.: Alaska 

deminu..tivum Easton 1944, Metriophyllum 

Miss. : Missouri 

densa Carruthers 1908, Zaphrentis (Davis, 1956) 

Miss.: Utah 

densum Moore and Jeffords 1945, Lophotichium 

Penn.: Oklahoma 

depressum Meek and Worthen 1866, Spherzopoter£um 

Miss. : Illinois 

dibolium Cocke 1970, Dibunophyllum 

Penn. : Kansas, Oklahoma 

dilatata Easton 1960, Lithostrotiorn [Lithostrotiorzella] 

Perm.: Nevada 

dilatatus Easton 1945c, Amplexus 

Miss.: Illinois, Utah 

dingmanae Nelson 1962a, Syringopora 

Miss. : Alberta 

disseptum Sutherland 1954, Zaph'riphyllurn 

Miss.: Northwest Territories 

distinctum Jeffords 1942, LophophyllidiU/m 

Penn.: Kansas 

distinctum Moore and Jeffords 1941, Leonardophyllum 

Perm.: Texas 

distorta Worthen ·in Worthen and Meek 1875, Cyathaxonia 

Penn.: Illinois, Missouri, Pennsylvania 

divergens White 1877, Favosites 

Miss.: Arizona, Iowa 

drummondi Nelson 1962a, Syringopora 

Miss. : Alberta 

dumosa Hyde 1953, Cladochonus 

Miss.: Ohio 

dunbari Moore and Jeffords 1941, Lophophyllidium 

Perm.: Kansas 

dunbari Ross and Ross 1962, N eokonirzckophyllum 

Penn. : Texas 

duncanae Hoare 1964, Ac,aciapora 

Perm.: Nevada 

durhami Langenheim and Tischler 1960, Meniscophyllum 

Miss. : California 

dutroi Armstrong 1972a, Lithostrotion (Siphonodendron) 

Miss. : Alaska 

dux Sutherland 1954, Kakwiphyllum 

Miss. : British Columbia, Nevada 

eastoni Langenheim and Tischler 1960, Ekvasophyllum 

Miss. : California 

eastoni Moore and Jeffords 1945, Lophophyllidium 

Penn.: Arkansas 

9 
10 NORTH AMERICAN LATE PALEOZOIC CORAL SPECIES 

elegente Cocke 1970, Dibunophyllum 

Penn.: Kansas 

eliasi Moo,re and Jeffords 1941, Lophamplexus 

Perm.: Kansas, Oklahoma 

elkoensis Hoa:re 1964, Caninia 

Perm.: Nevada 

elliptica White, 1862, Zaphrentis 

Miss.: Arizona, Illinois, Iowa, Kentucky, Missouri 

elongatum Jeffords 1942, LophophyUidiurn 

Penn.: Oklahoma 

elpasoensis Armstrong 1962, Koninckophyllum 

Penn.: Texas (A. K. Armstrong, oral commun., 1974) 

elyensis Wilson and Langenheim 19~6~2, Eastonoides 

Perm.: Nevada 

enclinotabulatum Sutherland 1958, Ekvasophyllurn 

Miss.: British Columbia 

enniskilleni Milne~ Edwards and Haime 1851, Zaphrentis (Sutherland, 1958) 

Miss.: Northwest Territories 

enorme Meek and Worthen 1860, Sphenopoterium 

Miss and Devonian.: Indiana, Iowa, Missouri, New Mexico 

enormis Easton 1945b, Caninia 

Miss.: Montana 

eugeneae White 1884, Michelinia 

Penn.: Arkansas, Illinois, Indiana, Kansas, Missouri, Oklahoma 

eumetrium Crickmay 1955, Allotropiophyllum 

Miss. : Alberta 

erecavatum Easton 1944, Clinophyllum 

Miss. : Missouri 

erecavatum Girty 1899, Menophyllum 

Miss.: Montana, Utah, Wyoming 

erecavatum Ulrich 1917, Menophyllum 

Miss. : Kentucky 

erecentric,a Meek 1873, Zaphrentis 

Miss.: Montana, Nevada, Utah 

erecentricus Moore and Jeffords 1945, Zaphrentoides 

Penn. : Texas 

ereigua Miller 1891, Zaphrentis 

Miss.: Illinois, Kentucky, Missouri 

ereigum Jeffords 1948, Dibunophyllum 

Penn. : Kansas 

exile Moore and Jeffords 1945, Lophophyllidium 

Penn. : Oklahoma 

exilimura Mather 1915, Michelinia 

Penn. : Oklahoma 

expansa White 1880b, Michilinia 

Miss.: Alberta, Arizona, Iowa, Missouri, Utah 

expansum Jeffords 1942, Lophophyllidium 

Penn.: Kansas, Missouri 

expansum White 1876a, Lophophyllum 

Miss.: Iowa 
SPECIES LIST 11 

expansus Easton 1945c, Amplexus 

Miss.: Oklahoma, Utah 

extumidum Moore and Jeffords 1945, Lophophyllidium 

Penn. : Texas 

fiaccida Easton 1943b, Caninia 

Miss. : Alabama 

fiexuosum Warren 1927, Lithostrotion (equals Diphyphyllum sinuosum 

Kelly) 

Miss. : Alberta 

fioriformis Hayasaka 1936, Lithostrotionella 

Miss. : Alaska 

floriformis d'Orbigny 1849, Acrocyathus 

Miss. : Indiana 

fragilis Mather 1915, Cladochonus 

Penn.: Oklahoma, Texas 

f'i·agilis White and St. John 1867, Amplexus 

Miss.: Arkansas, Iowa, Missouri 

ftatateeta McCutcheon and Wilson 1961, Ptolemaia 

Perm.: Nevada 

gabbi Meek 1864, Clisiophyllum 

Perm.: California 

garnettense Cocke 1970, Geyerophyllum 

Penn.: Kansas 

gasperense Butts 1922, Campophyllum 

Miss.: Alabama, Georgia, Kentucky, Tennes•see, Virginia 

genevievensis Easton 1957, Lithostrotion [Siphonodendron] 

Miss.: Alberta, British Columbia, Kentucky 

genic,ulatus Worthen 1890, Amplexus 

Miss.: Illinois, Kentucky, Utah 

geometrica Girty 1899, Aulopora 

Miss.: Alberta, Montana, Wyoming 

gibsoni White 1884, Zaphrentis 

Penn. : Arkansas, Colorado, Indiana, Oklahoma, Texas 

girtyi Armstrong 1970, Faberophyllum 

Miss. : Alaska 

girtyi Cocke 1970, Geyerophyllum 

Penn. : Kansas 

girtyi Haack 1914, Cyathaxonia 

Perm.: Coahuila 

girtyi Hayasaka 1936, Litho•strotionella 

Miss.: California, Illinois, Montana, Utah 

girtyi Jeffords 1942, Lophophyllidium 

Penn. : Oklahoma 

girtyi Moore and Jeffords 1941, H eritschia 

Perm. : Kansas 

glabrum Keyes 1894, Cyathophyllum 

Mis·s.: Iowa, Missouri 

glans White 1862, Zaphrentis 

Miss.: Iowa, Missouri 

goldcreekensis Hoare 1964, Caninia 

Perm.: Nevada 
12 NORTH AMERICAN LATE PALEOZOIC CORAL SPECIES 

gorbyi Miller 1891, Leptopora 

Miss.: Illinois, Iowa, Missouri, New Mexico 

grabaui Greene 1901a, Enallophyllum 

Miss.: Indiana 

· gracile Moore and Jeffords 1945, N eokoninckophyllum 

Penn. : Texas 

gracilis Keyes 1894, A ulopora 

Miss.: Iowa, Missouri 

gracUium Hoare 1964, Bradyphyllum 

Perm.: Nevada 

grandis Heritsch 1939, Petalaxis (Harker, 1960) 

Perm.: Northwest Territories (Arctic Islands) 

greenei Rowley in Greene 1901b, Blothrophyllum 

Miss. : Indiana 

greeni Rowley 1901, Coleophyllum? 

Miss.: Missouri · 

hadrum Jeffords 1947, Lophophyllidium 

Penn. : Kansas, Oklahoma 

hagei Sutherland 1954, Liardiphyllum 

Miss.: Northwest Territories 

haimeanum Ulrich 1917, Menophyllum 

Miss. : Kentucky 

hanseni Wilson and Langenheim 1962, Caninia 

Perm.: Nevada 

hansoni Coeke and Haynes 1973, Dibunophyllum 

Penn.: Oklahoma 

harkeri Nelson 1962b, Michelinia 

Penn.: Yukon Territory 

harkeri Sutherland 1958, Ekvasophyllum? 

Miss.: British Columbia 

harmodites Milne-Edwards and Haime 1851, Lithostrotion 

(Type specimen is a Silurian coral from Lake Huron region. As commonly used, it equals Lithostrotion [Siphonodendron] genevievensis 

Easton 1957) 

Miss.: Idaho, Kentucky, Tennessee 

harveyi White 1862, Syringopora 

Mis.s. : Alberta, Arizona, Iowa, Missouri 

heckeli Cocke 1970, Neokoninckophyllum 

Penn. : Kansas 

hemisphaerica Hayasaka 1936, Lithostrotionella 

Miss.: Idaho, Illinois, Kentucky, Mis·souri, Tennessee, Utah, Virginia, 

West Virginia 

hespere Grove 1935, Triplophyllum 

Miss. : New Mexico 

hessensis Ross and Ross 1962, Dibunophyllum 

Perm.: Texas 

hians Easton 1944, Rotiphyllum 

Miss.: Missouri, Montana 

killae Wilson and Langenheim 1962, Heritschioides 

Perm.: Nevada 
SPECIES LIST 

hudsoni Wilson and Langenheim 1962, Orionastrea 

Perm.: Nevada 

hypatiae Wilson 1963, Multithecopora 

Penn.: Nevada 

hystric.osum Cocke 1970, Dibunophyllum 

Penn. : Kansas 

ida Winchell 1865, Zaphrentis 

Miss.: Indiana, Ohio, Tennessee 

idonium Moore and J effo.rds 1945, Lophovphyllidium 

Penn. : Oklahoma, Texas 

ignekensis Armstrong 1972a, Lithostrotion (Siphonodendron) 

Miss.: Alaska 

ignotum Moore and Jeffords 1945, Lophophyllidium 

Penn.: Oklahoma 

illmoisensis Worthen 1890, Zaphrentis 

Miss. : Illinois, Iowa, Missouri 

illipahensis Easton 1960, Lonsdaleia 

Perm.: Nevada 

immota Moore and Jeffords 1945, Striatopora 

Penn. : Arkansas 

improcerum Moore and Jeffords 1945, Lophotichium 

Penn.: Oklahoma 

imulum Moore and Jeffords 1945, Empodesnw 

Penn. : Texas 

inauditum Moore and Jeffords 1945, Dibunophyllum? 

Penn. : Arizona, Oklahoma 

inclinatum Parks 1951, Ekvasophyllum 

Miss.: Utah 

inconferta Lonsdale 1845, Stylastrea (Salter, 1855) 

Perm.: Northwest Territories (Arctic Islands) 

inc,onstans Easton and Gutschick 1953, Lithostrotion (Diphyphyllum) 

Miss.: Arizona, Utah 

incrassatum Eas:ton and Gutschick 1953, Caninophyllum 

Mis,s.: Arizona, California 

indianensis Beede in Cumings and Beede 1906, Michelinia 

Miss.: Indiana 

indifferens Sutherland 1958, Amplexi-Zaphrentis 

Miss. : Northwest Territories 

infundibulum Worthen in Worthen and Meek 1875, Axophyllum 

Penn. : illinois, Mis,souri 

invagindtum Easton 1958, Cystelasma 

Miss. : Sonora 

iovaensis Worthen 1890, Anisophyllum? 

Miss.: Iowa 

irregularis Cocke and Bowsher 1968, S1dherlandia 

Penn.: Kansas, Oklahoma 

jagoensis Armstrong 1972b, Corwenia 

Penn. : Alaska 

jasperensis Kelly 1942, Lithostrotionella 

Miss.: Alberta, Nevada 

13 
14 NORTH AMERICAN LATE PALEOZOIC' CORAL SPE,CIES 

jewetti Cocke 1970, Geyerophyllum 

Penn. : Kansas 

juddi Thomson 1893, Campophyllum (Sloss, 1945) 

Miss. : Montana 

kansasense Miller and Gurley 1893, Campophyllum 

Penn. : Missouri 

kansasense Moore and Jeffords 1941, Malonophyllum 

Perm.: Kansas 

katoi Rowett 1969, Syringopora 

Perm. : Alaska 

kingi Jeffords 1955, Palaeacis 

Penn. : Texas 

kingi Ross and Ross 1962, Leonardophyllum 

Penn. : Texas 

klawockensis Armstrong 1970, Diphyphyllum 

Miss. : Alaska · 

knoxi Sutherland 1954, Canadiphyllum 

Miss.: British Columbia 

konincki Milne-Edwards and Haime 1851, Clisiophyllum (Armstrong, 1962) 

Penn.: Texas (A. K. Armstrong, oral commun., 1974) 

kunthi Stuckenberg 1888, Petalaxis (Harker, 1960) 

Perm.: Northwest Territories (Arctic Islands) 

lambarti Harker and McLaren 1950, Sciophyllu..m 

Miss. : Alaska, Yukon Territory 

lambii Ben 1929, Dibunophyllum 

Miss. : Nova Scotia 

lanc,eolatus Worthen 1890, Zaphrentis 

Miss.: Alabama, Illinois, Indiana, Kentucky, Tennessee, Utah 

lanesvillense Miller 1891, Cystelasma 

Miss. : Indiana 

languidum Parks 1951, Faberophyllum 

Miss.: Utah 

lanosum Jeffords 1947, Lophophyllidium 

Penn. : Texas -

latebrosa Moore and Jeffords 1945, Michelinia 

Penn.: Arkansas, Oklahoma, Texas 

leathamense Parks 1951, Faberophyllum 

Miss.: Utah 

lenis Jeffords 1947, Stereostylus 

Penn.: Kansas, Missouri 

lepidus Moore and Jeffords 1945, Pseudozaphrentoides 

Penn. : Texas 

leptosphragma Armstrong 1962, Michelinia 

Miss.: Arizona, New Mexico 

linnensis Cocke 1970, Caninia 

Penn. : Kansas 

lisburnensis Armstrong 1972a, Lithostrotion (Siphonodendron) 

Miss.: Alaska 

lochmanae Armstrong 1962, Lithostrotionella 

Miss. : Arizona, New Mexico 
longi Rowley 1901, Aulopora 

Miss. : Missouri 

SPECIES LIST 

lutarius Jeffords 1947, Lophamplexus? 

Penn. : Oklahoma 

macounii Lambe 1899a, Litho,strotion 

Miss.: Alaska, British Columbia 

macrostoma Romer 1883, Monilopora 

Miss.: Kentucky 

magnificum Jeffords 1942, Lophophyllidium 

Penn. : Oklahoma 

magnussoni Nelson 1962a, Syringopora 

Miss. : Alberta 

major Easton 1943a, Lonsdaleia 

Miss. : Arkansas 

mamillaris Castelnau 1843, Astraea 

15 

Miss.: Alabama, California, Illinois, Indiana, Iowa, Kentucky, Michigan, Missouri, Tennessee, Virginia (California occurrence is p.robably Permian) 

mancus Winchell 1865, Favosites? 

Miss.: Indiana 

mccutcheonae Wilson and Langenheim 1962, Syringopora 

Perm.: Nevada 

mcfarlani Ehlers and Stumm 1953, Palastraea 

Miss.: Kentucky 

mclareni Sutherland 1958, Lithostrotion [Lithostrotionella] [Thysanophyllum] 

Miss.: Alaska, British Columbia 

nwekana Girty 1910, Michelinia 

Miss.: Arkans.as, British Golumb~a, Oklahoma 

merriamorum Langenheim and. McCutcheon 1959, Bayhaium 

Perm. : California 

metum Moore and Jeffords 1945, Lophophyllidium 

Penn. : Arkansas, Oklahoma 

micra Kelly 1942, Lithostrotionella 

Miss. : Alberta 

microstylum White, 1880b, Lithostrotion 

Miss. : Alberta, Iowa, Missouri 

1nilichus Jeffords 1947, Stereostylus 

Penn. : Kansas 

minas Dawson 1868, Zaphrentis 

Miss. : Nova Scotia 

minatoi Rowett 1969, Sinopora 

Perm. : Alaska 

minnewankensis Shimer 1926, Triplophyllum 

Miss. : Alberta 

minor Easton 1943a, Lonsdaleia 

Miss. : Arkansas 

m·inor Weller 1909, Cyathaxonia 

Miss.: Arkansas, Missouri, Oklahoma, Texas, Utah 

minutum Jeffords 1942, Lophophyllidium 

Penn.: Oklahoma 
16 NORTH AMERICAN LATE PALEOZOIC ~CORAL, SPECIES 

minutum Simpson 1900, Meniscophyllum 

Miss. : Missouri 

mirabile Moore and Jeffords 1941, Sochkineophyllum 

Perm.: Kansas 

missouriensis Fraunfelter 1965, Dibunophyllmn 

Penn. : Missouri 

mokomokensis Easton 1960, Lithostrotion [Lithostrotionella] 

Penn. or Perm.: Nevada 

monroense Beede in Cumings and Beede 1906, Syringopora 

Miss.: Indiana 

rnontanensis Easton 1962, Caninia 

Miss. : Montana 

moorei Jeffords 1948, Dibunophyllum 

Penn.: Colorado, Oklahoma, Texas 

moorei Wells 1944, Striatopora 

Penn.: Texas 

moormanensis Stevens 1967, Heritschioides (EoheritscMoides) 

Perm.: Nevada 

morrowense Rowett and Sutherland 1964, Leonardophyllum 

Penn. : Oklahoma 

mulleri Wilson and Langeheim 1962, Sciophyllum 

Perm.: Nevada 

multattenuata McChesney 1860, Syringopora 

Penn.: Illino~s, Kansas, Missouri, Nebraska, Nevada, Oklahoma, Utah 

multiconum Parks 1951, Turbophyllum 

Miss.: Utah 

multilamellata Hall 1852, Zaphrentis 

Miss.: Utah 

multiradiata Hayasaka 1936, Lithostrotionella 

Miss.: Idaho, U tab 

tnundulum Jeffords 1942, Lophophyllidiurn 

Penn. : Oklahoma 

murale Jeffords 1942, Lophophyllidium 

Penn. : Kansas 

1nutabile Kelly 1942, Diphyphyllum 

Miss. : Alberta 

nasorakensis Armstrong 1972a, Diphyphyllum 

Miss. : Alaska 

nauvooense Bassler 1937, Hadrophyllum 

Miss. : Illinois 

nevadense Meek 1877, Cyathophyllum (Campophyllum?) 

Miss.: Utah 

nevadensis Hoare 1964, Cladochonus 

Perm.: Nevada 

nevadaensis Stevens 1967, Kleopatrina (Porfirievella) 

Perm.: Nevada 

newelli Jeffords 1942, Lophophyllidium 

Penn. : Oklahoma 

niakensis Armstrong 1972a, Lithostrotionella 

Miss.: Alaska 
SPECIES LIST 

nitellus Moore and Jeffords 1945, Pseudo.zaphrentoides 

Penn.: Arizona, Oklahoma 

northropi Armstrong 1962, A mplexizaphrentis 

Miss.: Arizona, New Mexico 

nunnensis Armstrong 1962, H apsiphyllum 

Miss. : New Mexico 

obtusum Meek and Worthen 1860, Sphenopoterium 

Miss. : Illinois, Iowa, Missouri 

occidentale Easton 1958, Rotiphyllum 

Miss. : Sonora 

occidentalis Meek 1877, Syringopora 

Penn. or Perm.: Utah 

occidentalis Merriam 1942, Litlwstrotion (Lithostrotionella) 

Perm. : Oregon 

occidentalis Sando 1963, Aulina 

Miss.: Arizona 

occ;ultum Parks 1951, Faberophyllum 

Miss.: Utah 

ochocoensis Merriam 1942, Waagenophyllum 

Perm.: Oregon 

oculinum Sando 1963, Lithostrotion (Siphonodendron) 

Miss. : Arizona 

ohioensis Bebout 1966, Stereostylus 

Penn.: Ohio 

ohioensis Stumm 1969, Amplexizaphrentis 

Miss.: Ohio 

oklahomense Cocke 1966, Dibunophyllum 

Penn.: Oklahoma 

oklahomense Rowett and Sutherland 1964, Koninckophyllum 

Penn. : Oklahoma 

oklahomensis Snide·r 1915, Pachypora 

Penn. : Arkansas, Oklahoma 

oppelti Greene 1900, Zaphrentis 

Miss.: Indiana 

ordinatus Ross and Ross 1962, Pseudozaphrentoides 

Penn. : Texas 

oregonensis Merriam 1942, Dibunophyllum 

Miss. : Oregon 

oregonensis Merriam 1942, L-ithostrotion (Siphonodendron) 

Miss.: Oregon 

01rientale Thomson 1880, Thysanophyllum (Armstrong, 1970b) 

Miss.: Alaska 

O'V·ale Bassler 1937, Hadrophyllum 

Miss.: Alabama, Georgia, Kentucky 

ovibos Salter 1855, Zaphrentis 

Perm.: Northwest Territories (Arctic Islands) 

packardi Merriam 1942, Lithostrotion (Lithostrotion) 

Miss.: Oregon 

palmatus Easton 1944, Triplophyllites 

Miss.: Illinois 

17 
18 NORTH AMERICAN LATE PALEOZOIC CORAL SPECIES 

panamintense L~angenheim and Tischler 1960, Clisiophyllum? 

Mis,s. : California 

pandatus Jeffords 1947 Stereostylus 

Penn. : Kans.a_s 

parva Greene 1900, Cyathaxonia 

Miss. : Indiana 

parvum Cocke 1969, Dibunophyllum 

Penn.: Kansas 

parvum Stevens 1967, Heritschioides (Eoheritschioides) 

Perm.: Nevada 

patulum Cocke 1970, Geyerophyllum 

Penn. : Kansas 

paucicinctus Easton and Gutschick 1953, Triplophyllites (Homalophyllites) 

Miss.: Arizona, Utah 

paucitabulata Moore and Jeffords 1945, Multithecopora 

Penn. : Texas 

pedunculatum Bassler 1937, Hadrophyllum 

Miss.: Kentucky, Tennessee 

pelaeus Jeffords 1947, Stereostylus 

Penn.: Kansas, Oklahoma 

pellaensis Worthen 1890, Zaphrentis 

Miss.: Iowa, Kentucky 

penchiensis Yoh 1927, Multithecopora (Nelson, 1962b) 

Penn.: Yukon Territory 

pennsylvanica Shimer 1926, Syringopora 

Miss. : Alberta 

pennsylvanicum Shimer 1926, Lithostrotion 

Miss. : Alaska, Alberta 

peratrovichensis Armstrong 19·70, Lithostrotionella 

Miss. : Alaska 

perelegans Billings 1859, Syringopora (Davis, 1956) 

(Type specimen i's Devonian) 

Miss.: Utah 

permiana Girty 1908, Lindstroemia. 

Perm. : Texas 

perplexum Cocke 1969, Dibunophyllum 

Penn. : Kans;as 

perplexum Cocke 1970, Neokoninckophyllum 

Penn. : Kansas 

persimilis Easton and Gutschick 1953, Triplophyllites ( Triplophyllites) 

Miss.: Arizona 

perversus Jeffords, 1947, Stereostylus 

Penn.: Kansas, Oklahoma 

petilum Cocke 1970, Neakoninckophyllum 

Penn. : Kansas 

petoczi Rowett 1969, Caninia 

Perm. : Alaska 

phainus Jeffords 1947, Stereostylus 

Penn. : Kansas 
S·PECIES LIST 

phractus J·effords 1947, Lophamplexus 

Penn.: Kansas 

pictoense Billings in Dawson 1868, Lithostrot·ion 

Miss. : N orva Scotia 

pilata Sutherland 1958, A mplexi-Zaphrentis 

Miss.: British Oo~umbi.a 

pinnatus Easton 1944, H apsiphyllum (H oma1ophyllites) 

Miss.: Missouri 

pisgahense Parks 1951, Faberophyllum 

Miss.: Utah 

placenta White 1880b, Michilinia? 

Miss.: Missouri, Montana, Wyoming 

plummeri Jeffords 1947, Lophophyllid·ium 

Penn. : Texas 

plummeri Wells 1944, Striatopora 

Penn. : Texas 

princeps Easton 1960, Lithostrotion [Thysanophyllum] 

Perm.: Nevada, Utah 

princetonensis Ulrich 1917, M enophyllum 

Miss.: Georgia, Kentucky 

princ.etonensis Ulrich 1905, Michelinia 

Mis·s.: Kentucky 

problematicum Goldfuss 1826, Pleurodictyum (WincheH, 1870) 

(Type s.pec,imen is Devonian) 

Miss.: Ohio 

procera Rowley 1901, Leptopora 

M.iss. : lllinoi•s, Missouri 

p'l'ofunda Milne-Edwards and Haime 1851, Cyathaxonia 

19 

Penn.: Alberta, Arkansas, Colorado, Illinois, Iowa, Kansas, Kentucky, 

Michigan, Nebraska, Ohio, Oklahoma, Pennsylvania, Texas, West Virginia (Alberta reference is Mi:ssissippian) 

prolifera McChesney 1860, Cyathaxonia 

Penn.: Illinois, Indiana, Iowa, Mis.souri, Ohio, Okl·ahoma, Texas, Utah 

proliferum Hall 1858, Lithostrotion 

Miss.: Alabama, Georgia, Illinois, Kentucky, Michigan, Tennessee, Utah, 

Virginia 

prona Milne-Edwards 1860, Zaphrentis 

Mists. : Indiana 

prosseri Beede 1898, Aulopora 

Penn.: Colorado, Kansas, Oklahoma 

proteus Sutherland 1958, Ekvasophyllum 

Miss.: Northwest Territories 

quinqueseptatum Ulrich 1905, Cystelasma 

Miss.: Alabama, Georgia, Kentucky, Virginia 

radiatum Jeffords 1942, Lophophyllidium 

Penn. : Kansas 

radicosa Ulrich 1917, Zaphrentis 

Miss.: Georgia, Kentucky 

radicosurn Girty 1912, Lophophyllum 

Penn.: Oklahoma, Texas 
20 NORTH AMERICAN LATE PAL,EOZOIC CORAL SPECIES 

radicula Rowley 1900, Zaphrentis 

Miss. : Missouri 

radigerus Rowley 1901, Amplexus 

Miss. : Missouri 

rainbowewis Rowett 1969, Timania 

Perm. : Alaska 

r-amosa Rowley 1901, Leptopora 

Mi,ss.: Illinois, Missouri 

ramulosa Goldfuss 1826, Syringopora (Lambe, 1899b) 

Miss. : Alberta 

'l'andolphensis Fraunfelter 1970, Dibunophyllum 

Penn.: Missouri 

readi Merriam 1942, Campophyllum? 

Miss. : Oregon 

referta Moore and Jeffords 1945, Mic,helinia 

Penn. : Oklahoma, Texas 

reiseri Armstrong 1973, Lithostrotion 

Miss.: Alaska 

religiosa Moore and Jeffords 1945, Striatopora 

Penn. : Texas 

repletum Moore and Jeffords 1945, Barytichisma 

Penn.: Texas 

reticulata Goldfuss 1826, Syringopora (Lambe, 1899h) 

Mis,s. ( ?) : British Columbia 

retusum Moore and Jeffords 1945, Hapsiphyllum 

Penn. : Texas 

reversa Worthen 1890, Zaphrentis 

Miss.: Illinois, Missouri, Utah 

rockfordensis Miller and Gurley 1897, Amplexus? 

Miss.: Illinois, Indiana 

romingeri Bassler 1937, Hadrophyllum 

Mi1ss.: Kentucky 

'I'Ossi Minato and Kato 1965, Y abeiphyllum 

Penn. or Perm. : Texas 

r'ltdis White and St. John 1867, Axophyllum 

Penn. : Kans13;s, Illinois, Indiana, Iowa, Missouri 

'l"udyi Nelson 1962.a, Syringopora 

Miss. : Alberta 

rugosum Hall 1852, Faviphyllum 

Miss.: Utah 

rugosum Ulrich 1905, Cystelasma 

Miss. : Indiana, Kentucky 

rugosus Weller 1909, Amplexus 

Miss.: Arkansas, Missouri 

sana Moore and Jeffords 1945, Paracaninia? 

Penn.: Texas 

sauridens White 1875, Lophophyllum 

Penn. :Colorado, New Mexico 

schucherti Heritsch 1936b, Palaeosmilia 

Perm.: Texas 
BPECIES LIST 

scopulosa Moore and Jeffords 1945, Michelinia 

Penn. : Arizona, Arkansas, Oklahoma 

sedaliense White 1880b, Chonophyllum 

Miss.: Iowa, Missouri 

semimuralis Sando 1963, Aulina 

Miss. : Arizona 

seminolensis Gocke and Bowsher 1968, Sutherlandia 

Penn. : Oklahoma 

septarugosum Moore and Jeffords 1941, Duplophyllum 

Perm.: Texas 

septata Greene 1901a, Cystelasma 

Miss.: Indiana 

shawi Wilson and Langenheim 1962, Cladochonus 

Perm.: Nevada 

shimeri Crickmay 1955, Lonsdaleia 

Mis.s. : Alberta, Arizona 

simplex Hayasaka 1936, Lithostrotionella 

Miss.: Utah, Wyoming 

simplex Moore and Jeffords 1945, Neokoninckophyllum 

Penn.: Oklahoma 

simulans Moore and Jeffords 1941, Timorphyllum 

Perm.: Texas 

sinuosum Kelly 1942, Diphyphyllum 

Miss.: Alaska, Alberta 

skinneri Heritsch 1937, Texanophyllum (nomen nudum) 

Perm.: Texas 

skinneri Ross and Ros·s 1962, Lophophyllid·ium 

Perm.: Texas 

solidum Ross and Ross 1962, Lophophyllidium 

Penn. and Perm.: Texas 

sonoraensis Armstrong 1962, Amplexizaphrentis 

Miss.: Arizona, New Mexico 

sonorense Easton 1958, Caninophyllum 

Miss. : Sonora 

spanius Jeffords 1947, Lophamplexus 

Penn. : Oklahoma 

spatiosus Moore and Jeffords 1945, Pseudozaphrentoides 

Penn.: Texas 

spergenensis Worthen 1890, Zaphrentis 

Miss. : Illinois, Indiana, Iowa, Missouri 

spinosum Jeffords 1947, Lophophyllidium 

Penn.: Texas 

spinulata Girty 1908, Cladopora 

Perm.: Coahuila, Texas 

spinulifera Hall 1858, Zaphrentis 

Miss.: Illinois, Iowa 

spinulosa Milne-Edwards and Haime 1851, Zaphrentis 

21 

Miss.: Alabama, Arkansas, J}eorgia, Iowa, Michigan Missouri, Montana, Oklahoma, Virginia 
22 NORTH AMERICAN: LATE PALEOZOIC CORAL SPECIE'S 

spissata Moore and Jeffords 1945, Michelinia 

Penn. : Arkansas, Oklahoma 

spissata Moore and Jeffords 1945, Stereocorypha 

Penn. : Oklahoma 

stansburyii Hall 1852, Zaphrentis 

Mis.s. : Idaho, New Mexico, Utah 

stelcki Nelson 1960, LithostTotionella 

Miss.: Alberta, British Columbia 

stokesi Milne-Edwards and Haime 1851, Lithostrotion 

(Originally described as Carboniferous from Manitoba. 

Type s.pecimen is a drift specimen derived from Devonian (Easton and 

Oliver, 1973)) 

striatissimus Easton 1946, Cladochonus 

Miss.: Missouri 

striatum Rowe<tt 1969, Cryptophyllum 

Penn. : Alaska 

striatus Easton 1944, Cladoc.honus (equals C. striatissimus Easton, 1946) 

Miss.: Missouri 

strimplei Cocke and Molinary 1973, N eokoninckophyllum 

Penn. : Oklahoma 

stuarti Nelson 1962b, Favosites? 

Perm.: Yukon Territory 

sturgeoni Bebout 1966, Stereostylus 

Penn.: Ohio 

subcaespitosum Meek 1873, Cyathophyllu.m 

Miss. : Idaho, Nevada 

subcrassus Ea;ston and Gutschick 1953, Triplophyllites (Homalophyllites) 

Miss.: Arizona, Utah 

subcylindrica Mather 1915, Michelinia 

Penn. : Oklahoma 

sublaeve Meek 1864, Lithostrotion 

Perm. : California 

subramosa Ulrich 1905, Michelinia 

Miss.: Georgia, Kentucky 

subramosum Easton 1958, Pleurodic,tyum 

Miss. : Sonora 

succinctus Armstrong 1970, Lithosf1·otion (Siphonodendron) 

Miss. : Alaska 

summitensis Rowetrt 1969, H eritschioides 

Perm. : Alaska 

surcularia Girty 1899, SyringopoTa 

Miss.: Alberta, Arizona, Colorado, Iowa, Montana, New Mexico, Utah, 

Wyoming 

sutherlandi Armstrong 1962, Vesiculophyllum 

Miss. : New Mexico 

sutherlandi Rowett 1969, DuThamina 

Perm. : Alaska 

tabularia Hoare 1964, Cornwallia 

Perm.: Nevada 

tabulata Hayasaka 1936, Lithostrotionella 

Miss.: Ldaho 
SIPEGIE.S LIST 23 

tabulatum Beede in Cumings and Beede 1906, Cystelasma 

Miss.: Indiana 

talonatum Easton 1943a, Koninckophyllum 

Miss.: Arkansas 

tantilla Miller 1891, Zaphrentis 

Miss.: Colorado, Illinois, Iowa, Missouri 

taylori Sutherland 1958, A mplexi-Zaphrentis 

Miss.: British Columbia, Northwest Territories 

tenella Miller 1891, Zaphrentis 

Miss.: Missouri, Sonora 

tennesseense Miller and Gurley 1895, Hadrophyllwm 

Miss. : Alabama, Tennessee 

tennesseensis Easton 1943b, Caninia 

Miss.: Tennessee 

tenuicula Moore and Jeffords 1945, Michelinia 

Penn. : Oklahoma 

teres Girty 1899, Clisiophyllum 

Miss. : Arizona, New Mexico, Wyoming 

tergidus Ross and Ross 1962, Stereostylus 

Perm.: Texas 

testata Moore and Jeffords 1945, Palaeacis 

Penn. : Texas 

texanum Heritsch 1936a, Waagenophyllurn 

Perm.: Texas 

texanum Moore and Jeffords 1945, Rodophyllurm 

Penn. : Texas 

texanum Okulitch and Albritton 1937, Malonophyl.lurn 

Perm.: Texas 

tcxanum Shumard 1859, Campophyllurn? 

Perm.: Texas 

texasensis Moore and Jeffords 1945, Cladochonus 

Penn. : Oklahoma, Texas 

torquium Owen 1852, Campophyllurn 

Penn.: Alberta, Arizona, Arkansas, Colorado, Illinois, Indiana, Iowa, 

Kansas, Missouri, Nebra.ska, Nevada, Oklahoma 

trac,hyporoides Wells 1944, Str·iatopora 

Penn. : Texas 

trigonalis Easton 1945a, Kinkaidia 

Miss.: Arkansas, Illinois 

trojana Easton 1960, Caninia 

Penn.: Nevada 

tschernyschewi Minato 1960, lpciphyllum 

Perm.: Northwest Territorie•s (Arctic Islands) 

tubifera Easton 1958, Syringopora 

Miss. : Sonora 

tubifera Hayasaka 1936 Lithostrotionella 

Mis•s.: Montana 

tubulata Girty 1908, Cladopora? 

Perm. : Texas 

tumida Easton 1962, Longiclava 

Miss. : Montana 
24 NORTH AMERICAN LATE PALEOZOIC CORAL SPECIES 

tumidum Moore and Jeffords 1945, Hapsiphyllum 

Penn. : Texas 

tumulus Salter 1855, Clisiophyllum 

Perm.: Northwest Territories (Arctic Islands) 

turbinata Billings 1859, "Favosites" (Allen and Leste.r, 1954) 

(Type specimen is Devonian) 

Miss.: Georgia 

turbineum Parks 1951, Ekvasophyllum 

Miss.: Utah 

tushanense Chi 1931, Koninckophyllum (Cocke, 1970) 

Penn.: Kansas, Oklahoma 

typa WincheH 1863, Leptopora 

Miss. : Illinois, Iowa, Missouri, New Mexico 

typicum Sando 1961, Ankhelasma 

Miss.: Utah 

uddeni Ross and Ross 1963, Dibunophyllum 

Penn. : Texas 

ulius Jeffords 1947, Lophamplexus 

Penn.: Kansas, Missouri 

ulrichanum Girty 1903, Menophyllum 

Miss. : Arizona, Colorado 

ulrichi Worthen 1890, Zaph·rentis 

Miss. : Kentucky 

umbonata Seebach 1866, Palaeacis 

Miss.: Iowa 

vagus Jeffords 1947, Lophamplexus 

Penn. : Kansas, Oklahoma 

'L•aleriae Newell 1935, Dibunophyllum 

Penn.: Kansas, Oklahoma 

vallum Hoare 1966, Bayhaimn 

Perm.: Nevada 

vallum Rosis and Ros~s 1962, Lophophyllidium 

Penn. : Texas 

valmeyerensis Weller 1909, Favosites 

Miss.: Missouri, Ohio 

variabile Cocke 1970, N eokoninckophyllum 

Penn. : Kansas 

variabilis Easton 1943a, Canhwst~·otion 

Miss.: Arkansas, Illinois 

'/Jarsoviensis Worthen 1890, Zaphrentis 

Miss.: Illinois, Iowa 

venosum Armstrong 1970, Diphyphyllwm 

Miss. : Alaska 

venusta Greene 1904b, Cyathaxonia 

Miss. : Indiana 

venusta Moore and Jeffords 1945, Acaciapora 

Penn. : Texas 

vermicularis Rowley 1901, Amplexus 

Miss. : Missouri 

verneuilanurn Milne-Edwards and Haime 1851, Baryphyllum 

Miss. : Tennessee 
SPECIES LIST 

verneuilanum Milne-Edwards and Haime 1851, 1-'·rochophyllum 

Miss.: Kentucky 

verticillata Goldfuss 1826, Syringopora (Davis, 1956) 

(Type s.pecimen is Silurian) 

Miss.: Utah 

verticillatum Barbour 1911, Craterophyllum 

Penn.: N ehraska 

veryi Greene 1904a, Lithodrumus 

Miss.: Kentucky 

vescum Moore and Jeffords 1945, Lophotich1'um 

Penn.: Arkansas, Oklahoma 

vesicularis Hayasaka 1936, Lithostrotionella 

Miss.: Alaska 

vesiculosum Easton 1958, Rotiphyllum 

Miss. : Sonora 

vidriensis Ross and Ross 1962, Lophophyllidium 

Perm. : Texas 

virginica Butts 1941, Sy·ringopo·ra 

Miss.: Alberta, British Columbia, Georgia, Virginia 

wahooensis Armstrong 1972b, Lithostrotionella 

Penn. : Alaska 

walcotti Moore and Jeffords 1945, Palaeacis 

Penn. : Texas 

wannensis Newell 1935, Zaphrentis 

Penn.: Oklahoma 

ward~· Howell 1945, Gymnophyllum 

Penn. : Oklahoma 

warreni Nelson 1960, Lithostrotion 

Miss. : Alaska, Alberta 

washburni Meuiam 1942, Waagenophyllu.tn 

Perm.: Oregon 

welleri Grove 1935, Triplophyllum 

Miss.: Missouri 

westii Beede 1898, Amplexus 

Penn. : Kansas, Missouri 

wewokanum Jeffords 1947, Lophophyllidium 

Penn. : Oklahoma 

whitfieldi White 187 4, Favosites (equals F. divergens White) 

Miss. : Arizona 

'lDhitneyi Meek 1877, Lithostrotion 

Miss.: Alberta, Utah 

whitneyi White 1875, Lithostrotion 

Penn. ( ?) : Nevada 

williamsi Armstrong 1970, Faberophyllurn 

Miss.: Alaska 

'tdlliamsi Greene 1899b, Michelin·ia 

Miss. : Indiana 

'Wilsonae Sutherland 1958, Paracaninia? 

Miss.: Northwest Territories 

'toilsoni Minato and Kato 1965, Kleopatrina (Kleopatrina) 

Perm.: Nevada 

25 
26 NORTH AMERICAN LATE PALEOZOIC CORAL SPECIES 

1.oimani Heritsch 1939, Roemeripora (Harker, 1960) 

Perm.: Northwest Territories (Arctic Islands) 

'Winchelli Rowley 1900, Cyathaxonia 

Miss.: Miss•ouri 

winchelli White 1879, Leptopora 

(Originally described as from Carboniferous of Utah. 

Type s.pecimen is a coral of Ordovician or Silurian age (Sando, 1965) ) 

woodi Wilson and Langenheim 1962, H eritschioides 

Perm.: Nevada 

wortheni Weller 1909, Zaphrentis (equals Triplophyllum welleri Grove) 

Miss.: Iowa, Kentucky,· Missouri 

yandelli Milne-Edwards and Ha:ime 1851, Amplexus 

Devonian and Miss. ( ?) : Mis,souri 

zaphrentiformis Greene 1901a, Bordenia 

Miss.: Indiana, Kentucky 

zaphrentiformis White 1876b, Amplexus 

Penn. : Colorado, Utah 

REFERENCES CITED 

Allen, A. T., Jr., and Lester, J. G., 1954, Cont:ributions to the paleontology 

of northwe.st Georgia: Georgia Geol. Survey Bull. 62, 166 p., 42 pls. 

Armstrong, A. K., 1958, The Mississippian of west-c:entral New Mexico: New 

Mexico Bur. Mines and Mineral R.e:sourees Mem. 5, 32 p., 6 pls., 21 figs. 

--- 1962, Stratigraphy and paleontology of the Missis·sip·pian System in 

s1outhwes,tern New Mexico and a.djacent s~outheaste,rn Arizona: New 

Mexico Bur. Mines and Mineral Resources Mem. 8, 99 p., 12 pls., 41 

figs .. 

--- 1970, Mississippian rugose cor,als, Peratrovich Formation, west coast, 

Prince of Wales Island, southeastern Alaska: U.S. Geo:l. Survey Prof. 

Pape:r 534, 44 p., 13 pls., 30 figs. 

--- 19·72a, Biostratigraphy of Missi·s.sippia.n lithostrotionoid corals, Lis~ 

burne Group, arctic Alaska: U.S. Geol. Survey Prof. Paper 743-A, 28 

p., 9 p1s., 25 figs. 

--- 1972b, Pennsylvanian carbonates, paleo·e·colo:gy, and rugo·S·e colonial 

corals, north flank, eastern Brooks Range, arctic Alaska: U.S. Geol. 

Survey Prof. Paper 747, 21 p., 8 pls., 16 figs. 

---· 1973, Lithostrotion reiseri n. sp., a cerioid coral from Meramec-age 

beds, Lisburne Group, arctic Alaska: U.S. Geol. Survey Jour. Res,earch, 

v. 1, no. 2, p. 137-145, 10 figs. 

Barbour, E. H., 1911, A new Carboniferous coral, Craterophyllum verticillatum: Nebraska Geol. Survey, v. 4, pt. 3, p. 38-49, 4 pis. 

Bassler, R. S., 1937, The Paleozoic rugose coral family Paleocyclidae: Jour. 

Paleontology, v. 11, no. 3, p. 189-201, pls. 30-32. 

--- 1950, Faunal lisrts and descriptions of Paleozoic corals: Geol. Soc. 

America Mem. 44, 315 p., 20 pls. 

Bebout, D. G., 196.6, Upp.er Penns•ylvanian Conemaugh corals from Ohio: 

Kansas Univ. Paleont. Contr., Paper 6, 7 p., 3 pls., 2 fi·gs. 

Beede, J. W., 1898, New corals from the Kansas Carboniferous: Kansas 

Univ. Quart., v. 7, no. 1, ser. A, p. 16-18. 
REFERENCES CITED 27 

Bell, W. A., 1929, Horton-Windsor district, Nova Scotia: Canada Geol 

Survey Mem. 155, 268 p., 36 pls., 14 figs. 

Billings, Elkanah, 1859, On the fossil corals of the Devonian rocks of Canada 

west: Canadian Jour. Indus.try, Science, and Art, neiW ser., v. 4, p. 

97-140, 29 figs. 

Branson, C. C., 1948, Bibliographic index of Permian invertebrates: Geol. 

Soc. America Mem. 26, 1049 p. 

Branson, E. B., and Greger, D. K., 19·18, Amsden Formation of the east 

slope of the Wind River Mountains of Wyoming and its fauna: Geol. 

Soc. America Bull., v. 29, no. 6, p. 309-326, pls. 18, 19. 

Butts, Charles, 1917, Descriptions and correlation of the Mississippian formations of we·stern Kentucky: Frankfort, Ky., Kentucky Geol. Survey, 

119 p. 

--- 1922, The Mississippian s.eries of eastern Kentucky; a regional interpretation of the stratigraphic relations of the Subcarboniferous 

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28 NORTH AMERICAN LATE PALEOZOIC' CORAL SPEGIE1S 

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30 NORTH AMERICAN LATE PALEOZOIC CORAL SPECIES 

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34 NORTH AME:RICAN LATE PALEOZOIC CORAL· SPECIES 

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