A Chronology of 

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By Craig M. Johnson

with contributions by

Stanley A. Ahler, Herbert Haas, and Georges Bonani

Smithsonian Institution

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Smithsonian Institution

Scholarly Press

smithsonian contributions to zoology • number 644

Catalog of Type Specimens 

of Recent Mammals:

Orders Didelphimorphia through 

Chiroptera (excluding Rodentia) in the 

National Museum of Natural History, 

Smithsonian Institution

Robert D. Fisher and Craig A. Ludwig
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smithsonian contributions to zoology • numb e r 6 4 4

Catalog of Type Specimens 

of Recent Mammals:

Orders Didelphimorphia through 

Chiroptera (excluding Rodentia) in the 

National Museum of Natural History, 

Smithsonian Institution

Robert D. Fisher and Craig A. Ludwig

WASHINGTON D.C.

2015
ABSTRACT

Fisher, Robert D. and Craig A. Ludwig, Catalog of Type Specimens of Recent Mammals: Orders Didelphimorphia through 

Chiroptera (excluding Rodentia) in the National Museum of Natural History, Smithsonian Institution. Smithsonian Contributions to Zoology, number 644, viii + 110 pages, 2015.—The type collection of Recent Mammals in the Division of 

Mammals, National Museum of Natural History, Smithsonian Institution, contains 821 specimens bearing names of 810 

species-group taxa of Didelphimorphia through Chiroptera, excluding Rodentia, as of June 2014. This catalog presents 

an annotated list of these holdings comprising 789 holotypes, 26 lectotypes, 11 syntypes (22 specimens), and 4 neotypes. 

Included are several specimens that should be in the collection but cannot be found or are now known to be in other collections. One hundred and twenty-eight of the names are new since the last type catalog covering these orders, Arthur J. 

Poole and Viola S. Schantz’s 1942 Catalog of the Type Specimens of Mammals in the United States National Museum, 

Incuding the Biological Surveys Collections (Bulletin of the United States National Museum, 178). Five specimens reported 

therein were subsequently sent to the National Museum’s Paleobiology Department collection and are mentioned only 

briefly in this work. They are Acratoenus? comes Miller, 1929, Paraconus serus Miller, 1929, Nesophontes hypomicrus 

Miller, 1929, Nesophontes paramicrus Miller, 1929, and Nesophontes zamicrus Miller, 1929. Orders and families are arranged systematically following D. E. Wilson and D. M. Reeder’s 2005 Mammal Species of the World: A Taxonomic and 

Geographic Reference,volume 1, third edition; within families, currently recognized genera are arranged alphabetically; 

within each currently recognized genus, species and subspecies accounts are arranged alphabetically by original published 

name. Information in each account includes original name and abbreviated citation thereto, current name if other than 

original, citation for first use of current name combination for the taxon (or new name combination if used herein for the 

first time), type designation, U.S. National Museum catalog number(s), preparation, age and sex, date of collection and 

collector, original collector number, type locality, and remarks as appropriate. Digital photographs of each specimen will 

serve as a condition report and will be attached to each electronic specimen record.

Cover images: Holotype of Sturnira angeli de la Torre, 1966 (USNM 361881). Left to right: dorsal, lateral, and ventral 

views of cranium. Photographs by D. F. Schmidt.

Published by SMITHSONIAN INSTITUTION SCHOLARLY PRESS

P.O. Box 37012, MRC 957

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Library of Congress Cataloging-in-Publication Data

National Museum of Natural History (U.S.)

Catalog of type specimens of recent mammals : orders Didelphimorphia through Chiroptera (excluding Rodentia) in 

the National Museum of Natural History, Smithsonian Institution / Robert D. Fisher and Craig A. Ludwig.

 pages cm. — (Smithsonian contributions to zoology ; number 644)

Includes bibliographical references and index.

1. Mammals—Type specimens—Catalogs and collections—Washington (D.C.) 2. National Museum of Natural 

History (U.S.)—Catalogs. I. Fisher, Robert D. II. Ludwig, Craig A. III. Title. 

QL708.2.N38 2015

599.074’753--dc23

2014040333

ISSN: 0081-0282 (print); 1943-6696 (online) 

Publication date (print): 24 March 2015

Ó The paper used in this publication meets the minimum requirements of the American National Standard for Permanence of Paper for Printed Library Materials Z39.48–1992. 
Contents

INTRODUCTION 1

Categories Of Name-Bearing Types Included 1

Research Approach 2

Classificatory Organization 2

Format of Accounts 2

Acknowledgments 3

ACCOUNTS OF NAME-BEARING TYPE SPECIMENS 3

Order Didelphimorphia Gill, 1872 3

 Family Didelphidae Gray, 1821 3

 Genus Caluromys J. A. Allen, 1900 3

 Genus Chacodelphys Voss, Gardner, and Jansa, 2004 3

 Genus Chironectes Illiger, 1811 3

 Genus Didelphis Linnaeus, 1758 4

 Genus Gracilinanus Gardner and Creighton, 1989 4

 Genus Marmosa Gray, 1821 4

 Genus Marmosops Matschie, 1916 5

 Genus Metachirus Burmeister, 1854 5

 Genus Monodelphis Burnett, 1830 6

 Genus Philander Brisson, 1762 6

 Genus Thylamys Gray, 1843 6

 Genus Tlacuatzin Voss and Jansa, 2003 6

Order Paucituberculata Ameghino, 1894 6

 Family Caenolestidae Trouessart, 1898 6

 Genus Lestoros Oehser, 1934 6

Order Dasyuromorphia Gill, 1872 6

 Family Dasyuridae Goldfuss, 1820 6

 Genus Pseudantechinus Tate, 1947 6

Order Diprotodontia Owen, 1866 7

 Family Phalangeridae Thomas, 1888 7

 Genus Ailurops Wagler, 1830 7

Order Macroscelidea Butler, 1956 7

 Family Macroscelididae Bonaparte, 1838 7

 Genus Elephantulus Thomas and Schwann, 1906 7

 Genus Petrodromus Peters, 1846 7
i v • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Order Hyracoidea Huxley, 1869 7

 Family Procaviidae Thomas, 1892 7

 Genus Dendrohyrax Gray, 1868 7

 Genus Heterohyrax Gray, 1868 7

Order Proboscidea Illiger, 1811 7

 Family Elephantidae Gray, 1821 7

 Genus Loxodonta Anonymous, 1827 7

Order Sirenia Illiger, 1811 8

 Family Trichechidae Gill, 1872 8

 Genus Trichechus Linnaeus, 1758 8

Order Cingulata Illiger, 1811 8

 Family Dasypodidae Gray, 1821 8

 Genus Cabassous McMurtrie, 1831 8

 Genus Dasypus Linnaeus, 1758 8

Order Pilosa Flower, 1883 8

 Family Bradypodidae Gray, 1821 8

 Genus Bradypus Linnaeus, 1758 8

 Family Megalonychidae Ameghino, 1889 8

 Genus Acratocnus Miller, 1929 8

 Genus Parocnus Miller, 1929 8

 Family Cyclopedidae Pocock, 1924 8

 Genus Cyclopes Gray, 1821 8

 Family Myrmecophagidae Gray, 1825 8

 Genus Myrmecophaga Linnaeus, 1758 8

Order Scandentia Wagner, 1855 9

 Family Tupaiidae Gray, 1825 9

 Genus Tupaia Raffles, 1821 9

Order Dermoptera Illiger, 1811 11

 Family Cynocephalidae Simpson, 1945 11

 Genus Galeopterus Thomas, 1908 11

Order Primates Linnaeus, 1758 12

 Family Lorisidae Gray, 1821 12

 Genus Nycticebus E. Geoffroy, 1812 12

 Family Galagidae Gray, 1825 13

 Genus Galago E. Geoffroy, 1796 13

 Family Tarsiidae Gray, 1825 13

 Genus Carlito Groves and Shekelle, 2010 13

 Genus Cephalopachus Swainson, 1835 13

 Genus Tarsius Storr, 1780 13

 Family Cebidae Bonaparte, 1831 14

 Genus Cebus Erxleben, 1777 14

 Genus Saguinas Hoffmannsegg, 1807 14

 Genus Sapajus Kerr, 1792 14

 Family Aotidae Elliot, 1913 14

 Genus Aotus Illiger, 1811 14

 Family Atelidae Gray, 1825 14

 Genus Alouatta Lacepede, 1799 14

 Genus Ateles E. Geoffroy, 1806 14

 Family Cercopithecidae Gray, 1821 15

 Genus Cercopithecus Linnaeus, 1758 15

 Genus Chlorocebus Gray, 1870 15

 Genus Colobus Illiger, 1811 16

 Genus Erythrocebus Trouessart, 1897 16
NUMBER 644 • v

 Genus Macaca Lacepede, 1799 16

 Genus Papio Erxleben, 1777 19

 Genus Presbytis Eschscholtz, 1821 19

 Genus Simias Miller, 1903 20

 Genus Trachypithecus Reichenbach, 1862 20

 Family Hylobatidae Gray, 1871 21

 Genus Hylobates Illiger, 1811 21

Order Lagomorpha Brandt, 1855 21

 Family Ochotonidae Thomas, 1897 21

 Genus Ochotona Link, 1795 21

 Family Leporidae G. Fischer, 1817 23

 Genus Brachylagus Miller, 1900 23

 Genus Lepus Linnaeus, 1758 23

 Genus Romerolagus Merriam, 1896 27

 Genus Sylvilagus Gray, 1867 27

Order Erinaceomorpha Gregory, 1910 32

 Family Erinaceidae G. Fischer, 1814 32

 Genus Atelerix Pomel, 1848 32

 Genus Echinosorex Blainville, 1838 32

 Genus Podogymnura Mearns, 1905 32

Order Soricomorpha Gregory, 1910 32

 Family Nesophontidae Anthony, 1916 32

 Genus Nesophontes Anthony, 1916 32

 Family Soricidae G. Fischer, 1814 32

 Genus Blarina Gray, 1838 32

 Genus Crocidura Wagler, 1832 33

 Genus Cryptotis Pomel, 1848 37

 Genus Megasorex Hibbard, 1950 40

 Genus Neomys Kaup, 1829 40

 Genus Notiosorex Coues, 1877 40

 Genus Sorex Linnaeus, 1758 40

 Genus Suncus Ehrenberg, 1832 49

 Genus Surdisorex Thomas, 1906 49

 Genus Sylvisorex Thomas, 1904 49

 Family Talpidae G. Fischer, 1814 49

 Genus Condylura Illiger, 1811 49

 Genus Dymecodon True, 1886 49

 Genus Euroscaptor Miller, 1940 50

 Genus Neurotrichus Gunther, 1880 50

 Genus Scalopus E. Geoffroy, 1803 50

 Genus Scapanus Pomel, 1848 50

Order Chiroptera Blumenbach, 1779 51

 Family Pteropodidae Gray, 1821 51

 Genus Acerodon Jourdan, 1837 51

 Genus Cynopterus F. Cuvier, 1824 51

 Genus Eonycteris Dobson, 1873 52

 Genus Harpyionycteris Thomas, 1896 52

 Genus Pteropus Brisson, 1762 52

 Family Rhinolophidae Gray, 1825 54

 Genus Rhinolophus Lacepede, 1799 54

 Family Hipposideridae Lydekker, 1891 55

 Genus Coelops Blyth, 1848 55

 Genus Hipposideros Gray, 1831 55
v i • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

 Family Megadermatidae H. Allen, 1864 56

 Genus Lavia Gray, 1838 56

 Genus Megaderma E. Geoffroy, 1810 56

 Family Rhinopomatidae Bonaparte, 1838 56

 Genus Rhinopoma E. Geoffroy, 1818 56

 Family Emballonuridae Gervais, 1855 56

 Genus Emballonura Temminck, 1838 56

 Genus Saccolaimus Temminck, 1838 57

 Genus Taphozous E. Geoffroy, 1818 57

 Family Phyllostomidae Gray, 1825 57

 Genus Anoura Gray, 1838 57

 Genus Ardops Miller, 1906 57

 Genus Artibeus Leach, 1821 57

 Genus Brachyphylla Gray, 1834 58

 Genus Carollia Gray, 1838 58

 Genus Centurio Gray, 1842 58

 Genus Chiroderma Peters, 1860 59

 Genus Choeroniscus Thomas, 1928 59

 Genus Dermanura Gervais, 1856 59

 Genus Ectophylla H. Allen, 1892 60

 Genus Erophylla Miller, 1906 60

 Genus Glossophaga E. Geoffroy, 1818 60

 Genus Leptonycteris Lydekker, 1891 61

 Genus Lichonycteris Thomas, 1895 61

 Genus Lonchophylla Thomas, 1903 61

 Genus Lophostoma d’Orbigny, 1836 61

 Genus Macrotus Gray, 1843 61

 Genus Micronycteris Gray, 1866 62

 Genus Mimon Gray, 1847 62

 Genus Monophyllus Leach, 1821 62

 Genus Phyllonycteris Gundlach, 1861 63

 Genus Platyrrhinus Saussure, 1860 63

 Genus Rhinophylla Peters, 1865 64

 Genus Sturnira Gray, 1842 64

 Genus Trachops Gray, 1847 64

 Genus Uroderma Peters, 1866 65

 Genus Vampyressa Thomas, 1900 65

 Genus Vampyrum Rafinesque, 1815 65

 Family Mormoopidae Saussure, 1860 65

 Genus Mormoops Leach, 1821 65

 Genus Pteronotus Gray, 1838 65

 Family Noctilionidae Gray, 1821 66

 Genus Noctilio Linnaeus, 1766 66

 Family Thyropteridae Miller, 1907 66

 Genus Thyroptera Spix, 1823 66

 Family Natalidae Gray, 1866 66

 Genus Chilonatalus Miller, 1898 66

 Genus Natalus Gray, 1838 66

 Family Molossidae Gervais, 1856 67

 Genus Chaerephon Dobson, 1874 67

 Genus Cheiromeles Horsfield, 1824 67

 Genus Eumops Miller, 1906 67

 Genus Molossus E. Geoffroy, 1805 68

 Genus Mormopterus Peters, 1865 68
NUMBER 644 • vii

 Genus Nyctinomops Miller, 1902 68

 Genus Promops Gervais, 1856 69

 Genus Tadarida Rafinesque, 1814 69

 Family Vespertilionidae Gray, 1821 70

 Genus Antrozous H. Allen, 1862 70

 Genus Corynorhinus H. Allen, 1865 70

 Genus Eptesicus Rafinesque, 1820 71

 Genus Histiotus Gervais, 1856 71

 Genus Hypsugo Kolenati, 1856 71

 Genus Kerivoula Gray, 1842 72

 Genus Laephotis Thomas, 1901 72

 Genus Lasiurus Gray, 1831 72

 Genus Miniopterus Bonaparte, 1837 73

 Genus Murina Gray, 1842 73

 Genus Myotis Kaup, 1829 73

 Genus Neoromicia Roberts, 1926 79

 Genus Nycticeius Rafinesque, 1819 80

 Genus Nyctophilus Leach, 1821 80

 Genus Parastrellus Hoofer, Van Den Bussche, 

and Horacek, 2006 80

 Genus Perimyotis Menu, 1984 81

 Genus Phoniscus Miller, 1905 81

 Genus Pipistrellus Kaup, 1829 81

 Genus Rhogeessa H. Allen, 1866 81

 Genus Scotophilus Leach, 1821 82

REFERENCES 83

INDEX OF TAXA 95

INTRODUCTION

The Division of Mammals, National Museum of Natural History (NMNH), Smithsonian Institution, formerly the United States National Museum (USNM), maintains the 

second-largest collection of mammal primary types in the world, numbering 3,202 specimens. Type specimens are “the international standards of reference that provide objectivity in zoological nomenclature” and as such “every institution in which name-bearing 

types are deposited should publish lists of name-bearing types in its possession or custody” 

(International Commission on Zoological Nomenclature, 1999:79). Although the official 

title of the museum was changed in 1969 (Yochelson, 1985) from United States National 

Museum to the National Museum of Natural History, the correct abbreviation for the museum’s cataloged holdings remains USNM. The abbreviation NMNH refers to the museum 

building and is not used as as prefix for catalog numbers, whereas USNM no longer refers 

to the museum itself and is only used in reference to museum catalog numbers.

This is the fifth type catalog of USNM mammals published to date, following Lyon 

and Osgood (1909; 1,415 specimens reported), Poole and Schantz (1942; 2,824 specimens reported), Fisher and Ludwig (2012; 843 specimens reported for the order Rodentia, suborders Sciuromorpha and Castorimorpha), and Fisher and Ludwig (2014; 945 

specimens reported for the order Rodentia, suborders Myomorpha, Anomaluromorpha, 

and Hystricomorpha). We continue to deviate from the two earlier catalogs and follow 

the precedent set by the NMNH Division of Fishes (Vari and Howe, 1991) and Division 

of Amphibians and Reptiles (Reynolds et al., 2007), as continued by Fisher and Ludwig 

(2012, 2014), in which the type collection is divided into smaller, more manageable taxonomic reports, rather than addressed in one monograph. The current work, the third of 

an anticipated four such publications, covers the orders Didelphimorphia through Chiroptera, excluding Rodentia, as ordered by Wilson and Reeder (2005).

Categories of Name-Bearing Types Included

Specimens included in this catalog are one of the following as defined by the International Code of Zoological Nomenclature (International Commission on Zoological 

Nomenclature, 1999; hereafter ICZN.

Holotype. The single specimen upon which a new nominal species-group taxon is 

based in the original publication (Article 73.1).

Catalog of Type Specimens of Recent Mammals:

Orders Didelphimorphia through Chiroptera (excluding 

Rodentia) in the National Museum of Natural History, 

Smithsonian Institution

Robert D. Fisher, U.S. Geological Survey, 

Patuxent Wildlife Research Center, National 

Museum of Natural History, Washington, D.C. 

20013-7012, USA; Craig A. Ludwig, Division 

of Mammals, Department of Vertebrate Zoology, 

National Museum of Natural History, Washington, D.C. 20013-7012, USA. Correspondence: 

fisherr@si.edu; ludwigc@si.edu. Manuscript received 2 July 2014; accepted 23 September 2014.
2 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Syntype. Any one of the multiple specimens on which a 

species-group name is equally based when no holotype was specified in the original publication and no subsequent designation 

of a lectotype has been published (Article 73.2). Syntypes are 

referred to as cotypes in Lyon and Osgood (1909) and Poole and 

Schantz (1942) as well as in other early literature.

Lectotype. One specimen selected from two or more specimens upon which a species-group name was originally based and 

designated in a publication subsequent to the original description to become the unique name bearer (Article 74.1).

Neotype. A specimen chosen as the name bearer of a speciesgroup taxon if no holotype, lectotype, syntype, or prior neotype 

is believed to be extant (Article 75.1).

Research Approach

In the early 1970s, accounts from a copy of Poole and 

Schantz (1942) were physically cut out and pasted onto 5 × 7 

index cards to create a master card file of types. Additional cards 

were created and added to the file as new taxa were described 

and their types installed in the collection. We checked the information on each card against the original description, the original skin label (when present), and the field catalog entry (when 

available). Information on each index card, often annotated by 

hand, thus served as the nucleus of each of the accounts. Other 

unpublished catalogs, such as the Museum Catalog, Merriam 

Collection Catalog, and X-Catalog, were consulted as needed.

Authors in Wilson and Reeder (2005) are followed for currently accepted names unless an online search of Zoological 

Record uncovered a more recent taxonomic arrangement or we 

became aware of an error of citation and attribution. In the taxonomic order presented herein (see the next section), this search 

was conducted beginning in December 2012 for the ealier accounts and running through March 2014 for the later accounts. 

Each specimen record in the electronic collection database was 

corrected as needed to reflect new information. The type publication citation was added to the electronic database if not already 

present. 

Digital images will be created for each specimen and will include dorsal, lateral, and ventral views of crania; dorsal and lateral views of mandibles; dorsal and ventral views of skins (lateral 

if possibly diagnostic); and both sides of all labels. All images 

will feature a metric scale bar. Additional views will be included 

if deemed appropriate. For lateral images, the traditional view of 

the left aspect will be presented except for skulls whose right side 

is more complete because of damage on the left. We will record 

specimen damage not captured in these images in the associated 

metadata when readily apparent but will not inspect specimens 

for minute damage. Images will be uploaded to the specimen 

database as they become available and can be accessed via the 

Internet (http://collections.mnh.si.edu/search/mammals/). Images 

will be created in the order in which accounts are presented. We 

anticipate it will take about six years for all images to be completed for specimens in this catalog. These images will not only 

serve as indicators of specimen condition, but will also allow 

researchers to verify certain characters of a type without having 

to visit the NMNH and handle the specimens, thereby providing 

a convenience to researchers and minimizing possible damage to 

irreplaceable specimens.

Classificatory Organization

Orders and families are sequentially arranged following 

Wilson and Reeder (2005). Within families, currently recognized 

genera are arranged alphabetically. Within each currently recognized genus, the specimen accounts are arranged alphabetically 

by original published name.

Format of Accounts

We follow the format set by Fisher and Ludwig (2014). Each 

account reads as follows:

• Original published name: Genus, species, and subspecies 

in bold followed by the author’s or authors’ names.

• Abbreviated type citation: Publication, volume, page, 

and date of publication. Dates are given as day, month 

(three-letter abbreviation), and year throughout.

• Currently recognized name: Genus, species, and subspecies, if different from the original published name, followed by the citation for the first use of current name 

combination for the taxon. Author names in parentheses 

indicate that the generic name used in the original description is different from the current generic one. The 

statement “new name combination” replaces the citation if the name is used for the first time herein. Current names are those used by the authors in Wilson and 

Reeder (2005) for the 17 orders included, unless more 

recent authors are cited.

• Type designation (holotype, lectotype, neotype, or syntype): USNM catalog number(s), preparation of specimen, age and sex, date collected and collector(s) name(s), 

and original number, which is usually the field number. 

Some type specimens were collected by personnel of the 

Smithsonian Venezuelan Project; the original numbers 

associated with these are preceeded by SVP.

• Type locality: Exact locality as given in the original description or lectotype designation, published restrictions, 

and supplementary data. Abbreviations are used for 

miles (mi), kilometers (km), feet (ft), and meters (m).

• Remarks: Additional information is provided as needed, 

but especially to explain types designated prior to becoming part of the USNM collection and reasons for considering certain specimens to be types when not specified as 

such in the original publication.

We make an effort to distinguish information that was included in the original description from that added later from 

other sources. Added information, such as amended or restricted 
NUMBER 644 • 3

localities, is given with citations. Brackets are used to identify 

supplementary data not contained in the original description 

and, when not from commonly used sources such as maps or 

online gazetteers, are attributed to a collector’s field catalog 

(FC), the Smithsonian Institution’s Museum Catalog (MC), the 

Merriam Collection Catalog (MCC), the skin label (SL), the XCatalog (XC), or a citation. Parentheses are placed around metric equivalents and current place names when this information 

was not included in the original publication.

Prior to January 1893, the USNM used a double-number 

cataloging system in which skins and fluid-preserved specimens 

were entered in one catalog and skeletal material in another. 

Many older specimens therefore have two numbers associated 

with them, and the proper museum record is indicated by a fraction such as 23651/31045. In this example, the numerator on 

a skin label indicates 23651 as the skin to which skull 31045 

belongs; on the associated skull label, 31045/23651, the numerator indicates 31045 as a skull to which the skin 23651 belongs. 

This cataloging system was abandoned at the end of 1892, after 

which a single, continuous numbering system was implemented 

for all specimen parts. The new system was initiated in January 1893, beginning with number 50001. The C. Hart Merriam 

private collection (MCC), donated to the USNM in 1911 and 

mentioned in many accounts, also used the double-numbering 

system.

Beginning in 1896 with the Biological Survey and continuing through 5 October 1951 with the Fish and Wildlife Service, 

specimens incidentally acquired through exchange or purchase, 

or received from or collected by persons other than permanent 

staff, were entered in a separate catalog, known as the X-Catalog, 

prior to being transferred to the Division of Mammals for formal 

cataloging. The X-Catalog thus served as an ersatz field catalog, and its entries frequently have data (referenced as XC) that 

were never entered into the museum catalog or onto skin labels. 

X-Catalog numbers are identified with an “-X” suffix.

Acknowledgments

We thank Robert P. Reynolds, USGS Patuxent Wildlife Research Center, NMNH, and Kristofer M. Helgen, Division of 

Mammals, NMNH, supervisors of Fisher and Ludwig, respectively, for support and encouragement to undertake this project. 

Funds to purchase the photographic setup, in order of significance, were provided by Carol E. Butler, assistant director for 

collections, NMNH; Don E. Wilson, chair, Department of Vertebrate Zoology, NMNH (since retired); and Robert P. Reynolds, 

to all of whom we are thankful. We are greatly indebted to David 

F. Schmidt, Division of Mammals, NMNH (since retired), with 

support from Linda K. Gordon, Division of Mammals, NMNH 

(since retired), who spent a considerable amount of time researching photographic equipment and methods, ordered the 

equipment, and set it up. Advice on photographic equipment, 

setup, and technique was graciously provided to David by Melvin J. Wachowiak and Elizabeth Webb, Museum Conservation 

Institute, Smithsonian Institution; Roy and Jason Larimer, Visionary Digital; and James L. Di Loreto, Office of Imaging and 

Photographic Services, NMNH. Renee Regan, a contractor 

funded by the Smithsonian Institution, Collection Information 

System, Information Resources Management pool, is currently 

doing photography for our earlier type catalog, and we hope 

to obtain additional funds to continue photography for the current publication. Alfred L. Gardner, USGS Patuxent Wildlife Research Center, NMNH, provided valuable insight to numerous 

taxonomic issues and guidance in interpreting various provisions 

of the International Code of Zoological Nomenclature. Alfred L. 

Gardner, Robert P. Reynolds, and Neal Woodman, all USGS 

Patuxent Wildlife Research Center, NMNH, read an earlier version of the manuscript and provided valuable comments.

ACCOUNTS OF NAME-BEARING 

TYPE SPECIMENS

Order Didelphimorphia Gill, 1872

Family Didelphidae Gray, 1821

Genus Caluromys J. A. Allen, 1900

Philander centralis Hollister

Proc. Biol. Soc. Wash. 27: 103, 11 May 1914a.

= Caluromys derbianus centralis (Hollister, 1914). See Goodwin 

(1946:285).

Holotype: USNM 12218/38522. Skin and skull. Adult female. 

Collected by W. Gabb, date unknown. Cataloged 19 Nov 

1874. No original number.

Type Locality: Talamanca, Costa Rica.

Genus Chacodelphys Voss, Gardner, 

and Jansa, 2004

Marmosa muscula Shamel

J. Wash. Acad. Sci. 20: 83, 4 Mar 1930a. Primary junior homonym of Didelphis (Marmosa) muscula Cabanas, 1846. 

Replaced with Marmosa formosa Shamel, J. Mammal. 11: 

311, 9 Aug 1930b.

= Chacodelphys formosa (Shamel, 1930). See Voss et al. (2004:2).

Holotype: USNM 236330. Skin and skull. Adult male. Collected 

9 Aug 1920 by A. Wetmore. Original number [SL: 1081].

Type Locality: [SL: Riacho Pilaga, 10 mi (16 km) NW of] kilometer 182, Formosa, Argentina.

Genus Chironectes Illiger, 1811

Chironectes panamensis Goldman

Smithson. Misc. Collect. 63(5): 1, 14 Mar 1914a.

= Chironectes minimus panamensis Goldman, 1914. See Cabrera (1958:44).
4 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Holotype: USNM 179164. Skin and skull. Old adult male. Collected 

23 Mar 1912 by E. A. Goldman. Original number 21562.

Type Locality: Cana, altitude 2,000 ft (610 m), [Darien], Panama.

Genus Didelphis Linnaeus, 1758

Didelphis marsupialis tabascensis J. A. Allen

Bull. Am. Mus. Nat. Hist. 14: 173, 15 Jun 1901a.

= Didelphis marsupialis caucae J. A. Allen, 1900. See A. L. Gardner (1973:26).

Holotype: USNM 100512. [Skin and skull]. [Lyon and Osgood 

(1908:1): Adult, SL: male]. Collected 7 Apr 1900 by E. W. 

Nelson and E. A. Goldman. Original number [SL: 14118].

Type Locality: Teapa, Tabasco, Mexico. 

Didelphis marsupialis texensis J. A. Allen

Bull. Am. Mus. Nat. Hist. 14: 172, 15 Jun 1901a.

= Didelphis virginiana californica Bennett, 1833. See A. L. Gardner (1973:30).

Holotype: USNM 33133/45137. Skin and skull. Adult male. 

Collected 13 Apr 1892 by F. B. Armstrong. Original number [SL: 12].

Type Locality: Brownsville, [Cameron County], Texas.

Remarks: Gardner (2005:6) used D. v. pigra for this taxon in 

error.

Didelphis richmondi J. A. Allen

Bull. Am. Mus. Nat. Hist. 14: 175, 15 Jun 1901a.

= Didelphis marsupialis caucae J. A. Allen, 1900. See A. L. Gardner (1973:26).

Holotype: USNM 33136/45140. Skin and skull. Adult female. 

Collected 10 Feb 1892, not 7 Feb as in original description, 

by C. W. Richmond. Original number [SL: 16].

Type Locality: Greytown (San Juan del Norte), [Rio San Juan], 

Nicaragua.

Didelphis yucatanensis J. A. Allen

Bull. Am. Mus. Nat. Hist. 14: 178, 15 Jun 1901a.

= Didelphis virginiana yucatanensis J. A. Allen 1901. See A. L. 

Gardner (1973:34).

Holotype: USNM 108299, not 100299 as in original description. [Skin and skull]. Adult male. Collected 1 Feb 1901, not 

29 Jan as in original description, by E. W. Nelson and E. A. 

Goldman. Original number [SL: 14468].

Type Locality: Chichenitza (Chichen Itza), Yucatan, [Mexico]. 

Didelphis yucatanensis cozumelae Merriam

Proc. Biol. Soc. Wash. 14: 101, 19 Jul 1901a.

= Didelphis virginiana yucatanensis J. A. Allen 1901. See A. L. 

Gardner (1973:34).

Holotype: USNM 108498. [Skin and skull]. Adult male. Collected 16 Apr 1901 by E. W. Nelson and E. A. Goldman. 

Original number 14700.

Type Locality: Cozumel Island, Yucatan (Quintana Roo), [SL: 

Mexico]. 

Genus Gracilinanus Gardner 

and Creighton, 1989

Gracilinanus perijae Hershkovitz

Fieldiana Zool. 70: 41, 30 Oct 1992.

= Gracilinanus marica (Thomas, 1898). See A. L. Gardner 

(2005:7).

Holotype: USNM 280881. Skin and skull. Adult male. Collected 

10 Mar 1943 by P. Hershkovitz. Original number 1826.

Type Locality: Las Marimondas, the site of a former coffee plantation, 1,450 m, eastern slope Serrania de Perija, Cordellera 

Oriental de los Andes, [Fonseca District], north and above 

Fonseca at the time in the department of Magdalena, now incorporated into the department of La Guajira, [SL: Colombia]. 

Genus Marmosa Gray, 1821

Marmosa demararae meridae Tate

Am. Mus. Novit. 493: 3, 26 Sep 1931.

See Voss and Jansa (2009:101) for use of Marmosa; meridae recognized as a valid subspecies of Micoureus demararae by 

Gardner and Creighton (2008:78).

Holotype: USNM 137510. Skin and skull. Old adult male. Collected 14 Jun 1904 by Briceño Gabaldon. No original number. [SL: 5393-X].

Type Locality: Cafetos de Merida, 1,630 m, Merida, [SL: 

Venezuela]. 

Marmosa isthmica Goldman

Smithson. Misc. Collect. 56(36): 1, 19 Feb 1912a.

See Rossi et al. (2010:42) for current use.

Holotype: USNM 170969. [Skin and skull]. Adult male. Collected 16 Feb 1911 by E. A. Goldman. Original number 

[FC: Nelson and Goldman] 20889. 

Type Locality: Rio Indio, near Gatun, Canal Zone (Colon), 

Panama. 

Marmosa murina mexicana Merriam

Proc. Biol. Soc. Wash. 11: 44, 16 Mar 1897a.

= Marmosa mexicana Merriam, 1897. See Bangs (1902:19).

Holotype: USNM 71526. [Skin and skull]. Immature male. Collected 28 Feb 1895 by E. W. Nelson and E. A. Goldman. 

Original number 7571.

Type Locality: Juquila, [FC: Altitude 5,000 ft (1,524 m)], Oaxaca, [FC: Mexico]. 

Marmosa ruatanica Goldman

Proc. Biol. Soc. Wash. 24: 237, 28 Nov 1911.

= Marmosa mexicana Merriam, 1897. See Rossi et al. (2010:24) 

for current use.

Holotype: USNM 7785/37700. Skin and skull. Adult, sex not 

given. Collected on unknown date by J. Akhurst. Cataloged 

4 Feb 1865.

Type Locality: Ruatan Island (Isla de Roatan). [Islas de la Bahia], 

off north coast of Honduras.
NUMBER 644 • 5

Remarks: The original description gave only the skin number 

USNM 7785; USNM 37700 is the skull that belongs to it. 

Marmosa xerophila Handley and Gordon

Vertebrate Ecology in the Northern Neotropics, p. 68, 4 Jan 1980.

Holotype: USNM 443819. Skin and skull. Adult male. Collected 

28 Jun 1986 by N. E. Peterson, F. P. Brown, Jr., and J. O. 

Matson. Original number SVP 23528.

Type Locality: La Isla, 15 m, near Cojoro, 37 km NNE Paraguaipoa, Department of Guajira (La Guajira), Colombia. 

Marmosa zeledoni Goldman

Proc. Biol. Soc. Wash. 24: 238, 28 Nov 1911.

See Rossi et al. (2010:37) for current use.

Holotype: USNM 12885/37133. Skin and skull. Adult male. 

Collected 1 Mar 1878 by J. C. Zeledon. Original number 

[SL: 119].

Type Locality: Navarro, [Miller and Kellogg (1955:9): near 

Orosi, 2,500-3,000 ft (762-914 m)], [Cartago], Costa Rica. 

Genus Marmosops Matschie, 1916

Marmosa caucae albiventris Tate

Bull. Am. Mus. Nat. Hist. 66: 182, 10 Aug 1933.

= Marmosops impavidus (Tschudi, 1845). See A. L. Gardner 

(2005:10).

Holotype: USNM 194378. Skin and skull. Adult male. Collected 

30 Apr 1915 by E. Heller. Original number 129.

Type Locality: Torontoy, near Cuzco (Cusco), altitude 8,000 ft 

(2,438 m) [FC: 8,500 ft (2,591 m)], [Cusco], Peru. 

Marmosa cracens Handley and Gordon

Vertebrate Ecology in the Northern Neotropics, p. 66, 4 Jan 

1980.

= Marmosops cracens (Handley and Gordon, 1980). See Gardner and Creighton (1989:4).

Holotype: USNM 418503. Skin and skull. Young adult male. 

Collected 11 Nov 1967 by N. E. Peterson, F. P. Brown, Jr., 

and J. O. Matson. Original number SVP 14837.

Type Locality: Near La Pastora, 11o12 ́N, 68o37 ́W, 150 m, 

14 km ENE Mirimire, Falcon, Venezuela. 

Marmosa invicta Goldman

Smithson. Misc. Collect. 60(2): 3, 20 Sep 1912b.

= Marmosops invictus (Goldman, 1912). See Gardner and 

Creighton (1989:4).

Holotype: USNM 178708. [Skin and skull]. Adult male. Collected 14 Mar 1912 by [SL: E. A. Goldman]. Original number 21517.

Type Locality: Cana, altitude 2,000 ft (610 m), [Darien], in the 

mountains of eastern Panama. 

Marmosa parvidens bishopi Pine

Mammalia 45: 63, 15 May 1981.

= Marmosops bishopi (Pine, 1981). See Voss et al. (2001:48).

Holotype: USNM 393535. Skin and skull. Adult female. Collected [SL: 5 Jul 1968] by R. H. Pine. Original number [SL: 

5176], not 5157 as given in original description.

Type Locality: 264 km N, by road, Xavantina, 12o51′S, 51o46 ́W, 

1,750 m, Serra do Roncador, Mato Grosso, Brazil. 

Marmosa parvidens pinheiroi Pine

Mammalia 45: 61, 15 May 1981.

= Marmosops pinheiroi (Pine, 1981). See Voss et al. (2001:49).

Holotype: USNM 461459. Skin and skull. Adult female. Collected 8 May 1969 by personnel of the Instituto Evandro 

Chagas. Original numbers T-391 and L5049.

Type Locality: Rio Amapari, Serra do Navio, 0o59 ́N, 52o03 ́W, 

Amapa, Brazil. 

Marmosa parvidens woodalli Pine

Mammalia 45: 62, 15 May 1981.

= Marmosops pinheiroi (Pine, 1981). See Voss et al. (2001:49).

Holotype: USNM 393532. Skin and skull. Adult female. Collected 14 Jun 1968 by R. H. Pine. Original number 5054; 

Belem Virus Laboratory number 20346.

Type Locality: “Nova Area Experimental,” Utinga, the wooded 

area surrounding the Belem waterworks, 1o27′S, 48o29′W, 

Belem, Para, Brazil. 

Marmosa purui G. S. Miller

Proc. Biol. Soc. Wash. 26: 31, 8 Feb 1913a.

= Marmosops noctivagus (Tschudi, 1844). See Voss and Jansa 

(2009:134).

Holotype: USNM 105543. [Skin in] alcohol, [skull removed]. 

Adult male. Collected Mar 1901 by J. B. Steere. No original 

number.

Type Locality: Hyutana-ham (Huitanaa), upper Purus River, 

[Amazonas], Brazil. 

Marmosops impavidus neblina A. L. Gardner

Advances in Neotropical Mammalogy. p. 414, 6 Feb 1990 [1989].

= Marmosops neblina A. L. Gardner, 1990. See Mustrangi and 

Patton (1997:22).

Holotype: USNM 560735. Skin and skull. Adult male. Collected 

5 Feb 1985 by A. L. Gardner. Original number 14436.

Type Locality: Camp VII, 00o50 ́40 ̋N, 65o58 ́10 ̋W, 1,800 m, 

Cerro de la Neblina, Territorio Federal Amazonas (Amazonas), Venezuela. 

Genus Metachirus Burmeister, 1854

Metachirus nudicaudatus dentaneus Goldman

Smithson. Misc. Collect. 56(36): 2, 19 Feb 1912a.

= Metachirus nudicaudatus colombianus J. A. Allen, 1900. See 

A. L. Gardner (2005:12).

Holotype: USNM 172732. [Skin and skull]. Adult male. Collected 12 Jan 1911 by E. A. Goldman. Original number [FC: 

Nelson and Goldman] 20837.

Type Locality: Gatun, Canal Zone (Colon), Panama. 
6 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Genus Monodelphis Burnett, 1830

Monodelphis kunsi Pine

Mammalia 39: 321, 20 Dec 1975.

Holotype: USNM 461348. Skin, skull, and skeleton. Adult male. 

Collected about Jul 1971 by E. Lotore. No original number.

Type Locality: La Granja, below 200 m, west bank of Rio Itonamas, 4 km north of Magdalena, Provincia Itenez, Departamento Beni (El Beni), Bolivia.

Remarks: Presented by M. L. Kuns, this specimen was originally 

prepared as a taxidermy mount by T. Rosas. The skeleton 

could not be found on 14 Mar 2013. 

P[eramys]. d[imidiata]. itatiayae Miranda-Ribeiro

Rev. Mus. Paul. 20: 421, 20 Jun 1936.

= Monodelphis sorex (Hensel, 1872). See Pine (1980:495).

Lectotype: USNM 461068 as designated by Avila Pires (1968:163). 

In alcohol [now a skin and skull]. [Avila Pires (1968:163): 

Adult] male. Collected in [SL: 1936] by Snr. Zikan. Original 

number [SL: 28. Museo National number 1311].

Type Locality: Campo Bello, E do Rio [SL: de Janeiro], Itatiaya 

(Itatiaia), [SL: Brazil].

Remarks: A type specimen was not designated by number in the 

original description. Avila Pires (1968:163) effectively designated a lectotype by listing this specimen as the type. (See 

ICZN, 1999, Recommendation 73F and Article 74.6). The 

specimen, originally in alcohol, had been made into a skin 

and skull by the time of Avila Pires’s catalog. It was cataloged as USNM 461068 on 5 Nov 1970. 

Peramys melanops Goldman

Smithson. Misc. Collect. 60(2): 2, 20 Sep 1912b.

= Monodelphis adusta (Thomas, 1897). See Handley (1966a:754).

Holotype: USNM 179609. [Skin, skull, and skeleton]. Adult 

male. Collected 23 May 1912 by E. A. Goldman. Original 

number 21737.

Type Locality: Cana, altitude [SL and FC: 2,800 ft (853 m)], not 

2,000 ft (610 m) as given in original description, [Darien], 

in the mountains of eastern Panama. 

Genus Philander Brisson, 1762

Metachirus fuscogriseus pallidus J. A. Allen

Bull. Am. Mus. Nat. Hist. 14: 215, 3 Jul 1901b.

= Philander opossum fuscogriseus (J. A. Allen, 1901). See Hershkovitz (1997:53).

Holotype: USNM 58158. [Skin and skull]. Adult male. Collected 

24 [SL and FC: Jan] 1894, not Jun as in original description, by 

E. W. Nelson and E. A. Goldman. Original number [SL: 5740].

Type Locality: Orizaba, [FC: altitude 4,200 ft (1,280 m)], Veracruz, Mexico. 

Genus Thylamys Gray, 1843

Marmosa tatei Handley

J. Wash. Acad. Sci. 46: 402, 13 Feb 1957.

= Thylamys tatei (Handley, 1957). See Reig et al. (1987:7).

Holotype: USNM 302915. Skin and skull, skinned from alcohol 

[SL: 8 Mar 1956]. Adult male. Collected Dec 1955 by J. M. 

de la Barrera. Original number 53/139.

Type Locality: Chasquitambo, 10o18′48 ̋S, 77o37′20 ̋W, 710 m, 

Ancachs (Ancash), Peru. 

Genus Tlacuatzin Voss and Jansa, 2003

Marmosa insularis Merriam

Proc. Biol. Soc. Wash. 12: 14, 27 Jan 1898a.

= Tlacuatzin canescens canescens (J. A. Allen, 1893). See A. L. 

Gardner (2005:18).

Holotype: USNM 89215. [Skin and skull]. Adult male. Collected 

16 May 1897 by E. W. Nelson and E. A. Goldman. Original 

number 11028.

Type Locality: Maria Madre Island, [FC: altitude 800 ft (244 m)], 

Tres Marias Islands, [Nayarit], Mexico. 

Marmosa oaxacae Merriam

Proc. Biol. Soc. Wash. 11: 43, 16 Mar 1897a.

= Tlacuatzin canescens canescens (J. A. Allen, 1893). See A. L. 

Gardner (2005:18).

Holotype: USNM 68240. [Skin and skull]. Adult female. Collected 14 Aug 1894 by E. W. Nelson and E. A. Goldman. 

Original number 6571.

Type Locality: City of Oaxaca, [FC: altitude 5,000 ft (1,524 m)], 

Oaxaca, Mexico. 

Order Paucituberculata Ameghino, 1894

Family Caenolestidae Trouessart, 1898

Genus Lestoros Oehser, 1934

Orolestes inca Thomas

Smithson. Misc. Collect. 68(4): 3, 10 Apr 1917.

= Lestoros inca (Thomas, 1917). See Oehser (1934:240).

Holotype: USNM 194401. [Skin, skull, and skeleton], [Poole 

and Schantz (1942:228): Adult] male. Collected [SL: 14 

May 1915 by E. Heller]. Original number 248.

Type Locality: Torontoy, [SL and FC: timberline], altitude 14,000 

ft (4,267 m), [Cusco], [SL: Peru]. 

Order Dasyuromorphia Gill, 1872

Family Dasyuridae Goldfuss, 1820

Genus Pseudantechinus Tate, 1947

Antechinus bilarni Johnson

Proc. Biol. Soc. Wash. 67: 77, 22 Mar 1954.

= Pseudantechinus bilarni (Johnson, 1954). See Kitchener and 

Caputi (1988:35).

Holotype: USNM 283970. Skin and skull. Adult female. Collected 

29 Oct 1948 by D. H. Johnson. Original number 5919.
NUMBER 644 • 7

Type Locality: Oenpelli (Gunbalanya), lat. 12o20 ́S, long. 

133o3 ́E, East Alligator River, Northern Territory, Australia. 

Order Diprotodontia Owen, 1866

Family Phalangeridae Thomas, 1888

Genus Ailurops Wagler, 1830

Phalanger furvus Miller and Hollister

Proc. Biol. Soc. Wash. 35: 115, 17 Oct 1922.

= Ailurops ursinus furvus (Miller and Hollister, 1922). See 

George (1987:509).

Holotype: USNM 219473. Skin and skull. Adult male. Collected 

5 Jan 1918 by H. C. Raven. Original number 3369.

Type Locality: Rano Rano (Ranorano), [FC: altitude about 

1,800 m], [Sulawesi Tengah], Middle Celebes (Sulawesi), 

[Indonesia]. 

Order Macroscelidea Butler, 1956

Family Macroscelididae Bonaparte, 1838

Genus Elephantulus

Thomas and Schwann, 1906

Elephantulus phaeus Heller

Smithson. Misc. Collect. 56(15): 8, 23 Dec 1910a.

= Elephantulus rufescens dundasi Dollman, 1910. See Corbet 

(1974:5).

Holotype: USNM 162074. [Skin and skull]. Adult male. Collected 26 Jun 1909 by J. A. Loring. Original number 6441.

Type Locality: Njoro O Solali, Sotik District, [Rift Valley], British East Africa (Kenya). 

Elephantulus rufescens mariakanae Heller

Smithson. Misc. Collect. 60(12): 10, 4 Nov 1912a.

= Elephantulus rufescens rufescens (Peters, 1878). See Corbet 

(1974:5).

Holotype: USNM 181821. [Skin and skull]. Adult female. Collected 31 Dec 1911 by E. Heller. Original number 5072.

Type Locality: Mariakani, [Coast Province], British East Africa 

(Kenya). 

Genus Petrodromus Peters, 1846

Petrodromus sultani sangi Heller

Smithson. Misc. Collect. 60(12): 12, 4 Nov 1912a.

= Petrodromus tetradactylus sultani Thomas, 1897. See Corbet 

(1974:2).

Holotype: USNM 181822. [Skin and skull]. Adult male. Collected 9 Nov 1911 by E. Heller. Original number 4698.

Type Locality: Summit of Mount Mbololo, altitude 4,000 ft 

(1,219 m), Taiti (Taita) Hills, [Coast Province], British East 

Africa (Kenya). 

Order Hyracoidea Huxley, 1869

Family Procaviidae Thomas, 1892

Genus Dendrohyrax Gray, 1868

Dendrohyrax validus True

Proc. U.S. Natl. Mus. 13: 228, 16 Sep 1890.

= Dendrohyrax arboreus (Hamilton Smith, 1827). See Shoshani 

(2005:87).

Lectotype: USNM 18986/34721 as designated by Lyon and Osgood (1909:26). Skin and skull. Adult male. Collected 17 

Jun 1888 by W. L. Abbott. Original number 8.

Type Locality: Mount Kilimanjaro, [Kilimanjaro], German East 

Africa (Tanzania).

Remarks: No type specimen was designated in the original description and True (1890:16) based the description on more 

than one specimen. Lyon and Osgood (1909:26) selected the 

above-listed specimen as the type, and effectively designated 

it as lectotype (ICZN, 1999: Article 74.5). 

Genus Heterohyrax Gray, 1868

Heterohyrax brucei albipes Hollister

Proc. Biol. Soc. Wash. 35: 135, 17 Oct 1922.

Holotype: USNM 181551. Skin and skull. Adult female. Collected 17 May 1911 by E. Heller. Original number [SL: 

1816].

Type Locality: [SL: Sotik, on the] Telek River, Loita Plains, [Rift 

Valley], Kenya Colony (Keyna). 

Order Proboscidea Illiger, 1811

Family Elephantidae Gray, 1821

Genus Loxodonta Anonymous, 1827

Elephas africanus rothschildi Lydekker

Proc. Zool. Soc. London 1: 399, 23 Apr 1907.

= Loxodonta africana (Blumenbach, 1797). See Roosevelt and 

Heller (1914:736).

Holotype: USNM 22247. [Cross-section of upper part of tusk]. 

Male. Inscribed on the tusk is: born in Africa, 1861; died at 

St. Thomas, [Ontario], Canada 15 Sep 1885. Date of collection, collector, and original number unknown.

Type Locality: Some part of the French Sudan (Mali), probably 

to the southward of Lake Chad.

Remarks: This locality is probably in Chad or Cameroon. 

Lydekker (1907:399) took for his type the statuette of 

“Jumbo” in the British Museum. This piece of tusk is from 

the original animal and therefore is part of the holotype. 

(See ICZN, 1999, Article 73.1.4.) In a letter from H. A. 

Ward to Spencer F. Baird, dated 23 Mar 1886, Ward stated 

that he had a few slices prepared from the part of one of 

the tusks of Jumbo which was buried in his head and he 
8 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

mailed a slice to Baird thinking it might be of interest to him 

(USNM accession record 17276). The piece was received 

26 Mar 1886 and cataloged 28 Sep 1886. Shoshani et al. 

(1986) chronicle the history and deposition of the remains 

of “Jumbo.” This specimen was not included in either Lyon 

and Osgood (1909) or Poole and Schantz (1942). 

Order Sirenia Illiger, 1811

Family Trichechidae Gill, 1872

Genus Trichechus Linnaeus, 1758

Manatus oweni Du Chaillu

Proc. Boston Soc. Nat. Hist. 7: 367, 1861.

= Trichechus senegalensis Link, 1795. See Gray (1865:1340).

Syntype: USNM 20907. [Skull and skeleton]. Age and sex unknown. Collected on unknown date by P. B. Du Chaillu. 

Cataloged [MC: 20 Apr 1883].

Type Locality: Central Africa. [Gray (1865:134): Mouth of the 

Gabon].

Remarks: Received from the British Museum (Natural History) 

on 19 Apr 1883. British Museum number 1388a. 

Order Cingulata Illiger, 1811

Family Dasypodidae Gray, 1821

Genus Cabassous McMurtrie, 1831

Tatoua (Ziphila) centralis G. S. Miller

Proc. Biol. Soc. Wash. 13: 4, 31 Jan 1899a.

= Cabassous centralis (G. S. Miller, 1899). See Palmer (1899:72).

Holotype: USNM 19464/35382. Skin and skull. Adult female. Collected 8 Jan 1891 by E. Wittkugel. Original number [SL: 8].

Type Locality: Chamelicon (Chamelecon), [Cortes], Honduras. 

Genus Dasypus Linnaeus, 1758

Tatu novemcinctum texanum Bailey

N. Am. Fauna 25: 52, 24 Oct 1905.

= Dasypus novemcinctus mexicanus Peters, 1864. See Hollister 

(1925:60).

Holotype: USNM 34352/46438. Skin and skull. Adult male. 

Collected 10 Jun 1892, not 1902 as in original description, 

by F. B. Armstrong. Original number 4.

Type Locality: Brownsville, [Cameron County], Texas. 

Order Pilosa Flower, 1883

Family Bradypodidae Gray, 1821

Genus Bradypus Linnaeus, 1758

Bradypus ignavus Goldman

Smithson. Misc. Collect. 60(22): 1, 28 Feb 1913.

= Bradypus variegatus ephippiger Philippi, 1870. See Hall 

(1981:280).

Holotype: USNM 179551. [Skin and skull]. Adult female. Collected 6 Apr 1912 by E. A. Goldman. Original number 21596.

Type Locality: Marraganti, [SL and FC: sea level], about 2 mi (3 

km) above Real de Santa Maria, near head of tidewater on 

the Rio Tuyra (Rio Tuira), [Darien], eastern Panama. 

Bradypus pygmaeus Anderson and Handley

Proc. Biol. Soc. Wash. 114: 17, 19 Apr 2001.

Holotype: USNM 579179. Skin and skull. Adult female. Collected 27 Mar 1991 by C. O. Handley, Jr. and E. P. Nelson. 

Original number EPN 166.

Type Locality: [SL: Northern Quarter], West Point, Isla Escudo 

de Veraguas, Bocas del Toro, Panama. 

Family Megalonychidae Ameghino 1889

Genus Acratocnus Miller, 1929

Acratocnus? comes G. S. Miller

Smithson. Misc. Collect. 81(9): 26, 30 Mar 1929.

Holotype: USNM 253178.

This specimen was transferred to the NMNH Paleobiology Department, date unknown. 

Genus Parocnus Miller, 1929

Parocnus serus G. S. Miller

Smithson. Misc. Collect. 81(9): 29, 30 Mar 1929.

Holotype: USNM 253228.

This specimen was transferred to the NMNH Paleobiology Department, date unknown.

Family Cyclopedidae Pocock, 1924

Genus Cyclopes Gray, 1821

Cyclopes mexicanus Hollister

Proc. Biol. Soc. Wash. 27: 210, 31 Oct 1914b.

= Cyclopes didactylus mexicanus Hollister, 1914. See Hall and 

Kelson (1952:316).

Holotype: USNM 11137/38534. Skin and skull. Adult, sex not 

given. Collected on unknown date by J. C. Spear. [MC: Cataloged 30 Dec 1872]. Original number [SL: 8].

Type Locality: Tehuantepec, Oaxaca, Mexico. 

Family Myrmecophagidae Gray, 1825

Genus Myrmecophaga Linnaeus, 1758

Myrmecophaga centralis Lyon

Proc. U.S. Natl. Mus. 31: 570, 14 Nov 1906a.

= Myrmecophaga tridactyla centralis Lyon, 1906. See Goldman 

(1920:64).

Holotype: USNM 15963. Skull only. Young adult, sex not given. 

Collected Jun 1876 by J. C. Zeledon. Original number 86.

Type Locality: Pacuare, Costa Rica. 
NUMBER 644 • 9

Order Scandentia Wagner, 1855

Family Tupaiidae Gray, 1825

Genus Tupaia Raffles, 1821

Tana cervicalis masae Lyon

Proc. U.S. Natl. Mus. 45: 148, 29 Nov 1913.

= Tupaia tana masae (Lyon, 1913). See Chasen (1940:7)

Holotype: USNM 121835. Skin and skull. Adult female. Collected 18 Feb 1903 by W. L. Abbott. Original number 2302.

Type Locality: Tana Masa (Pulau Tanahmasa), Batu Islands 

(Kepulauan Batu), [Sumatera Utara], west coast of Sumatra, 

[Indonesia]. 

Tana lingae Lyon

Proc. U.S. Natl. Mus. 45: 145, 29 Nov 1913.

= Tupaia tana lingae (Lyon, 1913). See Chasen (1940:7).

Holotype: USNM 101597. Skin and skull. Adult male. Collected 

16 Jul 1899 by W. L. Abbott. No original number.

Type Locality: On the peak of Linga Island (Pulau Lingga), altitude 2,000 ft (610 m), Rhio-Linga Archipelago (Kepulauan 

Lingga), [Kepulauan Riau, Sumatra, Indonesia]. 

Tana tana besara Lyon

Proc. U.S. Natl. Mus. 45: 141, 29 Nov 1913.

= Tupaia tana besara (Lyon, 1913). See Chasen (1940:6).

Holotype: USNM 142247. Skin and skull. Adult male. Collected 

17 Sep 1905 by W. L. Abbott. Original number 4458.

Type Locality: Opposite Pulo Jambu, below Tyan (Tayan), on 

north, or right, bank of Kapuas River (Sungai Kapuas), [Kalimantan Barat], western Borneo (Kalimantan), [Indonesia]. 

Tana tana tuancus Lyon

Proc. U.S. Natl. Mus. 45: 145, 29 Nov 1913.

= Tupaia tana speciosa (Wagner, 1841). See Helgen (2005:108).

Holotype: USNM 114412. Skin and skull. Adult male. Collected 

29 Jan 1902, not 1905 as in original description, by W. L. 

Abbott. Original number 1489.

Type Locality: Pulo (Pulau) Tuangku, Banjak Islands (Kepulauan 

Banyak), [Aceh], west coast of Sumatra, [Indonesia]. 

Tupaia anambae Lyon

Proc. U.S. Natl. Mus. 45: 89, 29 Nov 1913.

= Tupaia glis (Diard, 1820). See Chasen and Kloss (1928a:38). 

Subspecies of T. glis not recognized by Helgen (2005:106).

Holotype: USNM 101743. Skin and skull. Adult male. Collected 

23 Sep 1899 by W. L. Abbott. No original number.

Type Locality: Pulo Jimaja (Pulau Jemaja), Anamba Islands 

(Kepulauan Anambas), South China Sea, between Malay 

Peninsula and Borneo, [Kepulauan Riau, Indonesia]. 

Tupaia bunoae G. S. Miller

Proc. Wash. Acad. Sci. 2: 229, 20 Aug 1900a.

= Tupaia tana bunoae G. S. Miller, 1900. See Chasen (1940:7).

Holotype: USNM 101640. Skin and skull. Adult female. Collected 

5 Aug 1899 by [MC: W. L. Abbott]. No original number.

Type Locality: Pulo Bunoa (Pulau Benua), Tambelan Islands 

(Kepulauan Tambelan), [Kepulauan Riau, Indonesia]. 

Tupaia carimatae G. S. Miller

Proc. U.S. Natl. Mus 31: 61, 23 Jul 1906a.

= Tupaia splendidula carimatae G. S. Miller, 1906. See Medway 

(1961:271).

Holotype: USNM 125123. Skin and skull. Adult male. Collected 

2 Sep 1904 by W. L. Abbott. Original number 3716.

Type Locality: [SL: Telok Edar], Karimata Island (Pulau 

Karimata), [Kalimantan Barat, Borneo (Kalimantan), 

Indonesia]. 

Tupaia castanea G. S. Miller

Smithson. Misc. Collect. 45(1): 54, 6 Nov 1903a.

= Tupaia glis (Diard, 1820) See Chasen (1940:3). Subspecies of 

T. glis not recognized by Helgen (2006:106).

Holotype: USNM 115608. Skin and skull. Adult female. Collected 11 Aug 1902 by W. L. Abbott. Original number 1872.

Type Locality: [FC: Pasin Panjong], Pulo Bintang (Pulau Bintan), 

Rhio Archipelago (Kepulauan Riau), [Kepulauan Riau, Sumatra, Indonesia]. 

Tupaia cervicalis G. S. Miller

Smithson. Misc. Collect. 45(1): 59, 6 Nov 1903a.

= Tupaia tana cervicalis G. S. Miller, 1903. See Chasen (1940:7).

Holotype: USNM 121754. Skin and skull. Adult male. Collected 

14 Feb 1903 by W. L. Abbott. Original number 2294.

Type Locality: Tana Bala (Pulau Tanahbala), Batu Islands (Kepulauan Batu), [Sumatera Utara], Sumatra, [Indonesia]. 

Tupaia chrysogaster G. S. Miller

Smithson. Misc. Collect. 45(1): 58, 6 Nov 1903a.

Holotype: USNM 121572. Skin and skull. Adult female. Collected 21 Nov 1902 by W. L. Abbott. Original number 2078.

Type Locality: North Pagi Island (Pulau Pagai Utara), [Sumatera 

Barat], Sumatra, [Indonesia]. 

Tupaia chrysomalla G. S. Miller

Proc. Wash. Acad. Sci. 2: 232, 20 Aug 1900a.

= Tupaia glis (Diard, 1820). See Chasen and Kloss (1928a:38) 

who place chrysomalla in T. glis. Subspecies of T. glis not 

recognized by Helgen (2005:106).

Holotype: USNM 101710. Skin and skull. Adult female. Collected 24 Aug 1899 by [MC: W. L. Abbott]. No original 

number.

Type Locality: Pulo (Pulau) Siantan. Anambas Islands (Kepulauan Anambas), [Kepulauan Riau, Indonesia]. 

Tupaia cuyonis G. S. Miller

Proc. U.S. Natl. Mus. 38: 393, 19 Aug 1910.

= Tupaia moellendorffi cuyonis G. S. Miller, 1910. See Helgen 

(2005:107).
1 0 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Holotype: USNM 239182. Skin and skull. Adult male. Collected 

15 Jan 1903 by R. C. McGregor and A. Celestino. Original 

number Philippine Bureau of Science 26.

Type Locality: Island of Cuyo, [Palawan Province], [MC: Philippine Islands] (Philippines).

Remarks: The holotype was published as Philippine Bureau of 

Science number 26. It was cataloged as USNM 239182 on 

6 Feb 1923. 

Tupaia discolor Lyon

Proc. U.S. Natl. Mus. 31: 602, 18 Dec 1906b.

See Sargis et al. (2013:201) for current use.

Holotype: USNM 124703. Skin and skull. Adult female. Collected 24 May 1904 by W. L. Abbott. Original number 

3262.

Type Locality: Tanjong Rengsam, Island of Banka (Pulau 

Bangka), [Kepulauan Bangka Belitung], east of Sumatra, 

[Indonesia]. 

Tupaia ferruginea batamana Lyon

Proc. U.S. Natl. Mus. 31: 656, 16 Jan 1907a.

= Tupaia glis (Diard, 1820). See Lyon (1913:46). Subspecies of 

T. glis not recognized by Helgen (2005:106). 

Holotype: USNM 142151. Skin and skull. Adult female. Collected 15 Sep 1905 by C. B. Kloss. Original number 2.

Type Locality: Semimba Bay (Teluk Senimba), Batam Island 

(Pulau Batam), south of Malay Peninsula, [Kepulauan Riau, 

Indonesia]. 

Tupaia glis olivacea Kloss

J. Nat. Hist. Soc. Siam 3: 358, 31 Dec 1919.

= Tupaia belangeri olivacea Kloss, 1919. See Sargis et al. 

(2013:200) for current use.

Holotype: USNM 221562. Skin and skull. Adult male. Collected 

23 Oct 1916 by C. B. Kloss. Original number 2208.

Type Locality: Pak Bu, near Tachin (Samut Sakhon), [Samut 

Sakhon], Siam (Thailand).

Remarks: The holotype was published as C. B. Kloss number 

2208. It was cataloged as USNM 221562 on 2 Jul 1920. 

Tupaia gracilis edarata Lyon

Proc. U.S. Natl. Mus. 45: 118, 29 Nov 1913.

Holotype: USNM 153859. Skin and skull. Young adult female. 

Collected 5 Oct 1908 by W. L. Abbott. Original number 

6226.

Type Locality: Telok Edar, Karimata Island (Pulau Karimata), 

[Kalimantan Barat], off west coast of Borneo (Kalimantan), 

[Indonesia]. 

Tupaia inflata Lyon

Proc. U.S. Natl. Mus. 31: 600, 18 Dec 1906b.

= Tupaia gracilis inflata Lyon, 1906. See Lyon (1913:118).

Holotype: USNM 124709. Skin and skull. Adult male. Collected 

20 May 1904 by W. L. Abbott. Original number 3241.

Type Locality: Tanjong Rengsam, island of Banka (Pulau Bangka), 

[Kepulauan Bangka Belitung], east of Sumatra, [Indonesia]. 

Tupaia lacernata kohtauensis Shamel

J. Mammal. 11: 71, 11 Feb 1930c.

= Tupaia belangeri kohtauensis Shamel, 1930. See Sargis et al. 

(2013:200).

Holotype: USNM 253446. Skin and skull. Adult male. Collected 

19 Sep 1928 by H. M. Smith. Original number [SL: 2372].

Type Locality: Island of Koh Tau (Ko Tao), [Surat Thani, 

Thailand]. 

Tupaia longipes salatana Lyon

Proc. U.S. Natl. Mus. 45: 77, 29 Nov 1913.

= Tupaia salatana Lyon, 1913. See Sargis et al. (2013:201).

Holotype: USNM 151882. Skin and skull. [Poole and Schantz 

(1942:205): Adult, SL and FC: male]. Collected 11 Feb 

1908 by W. L. Abbott. Original number 5785.

Type Locality: 15 mi (24 km) from mouth of Pangkallahan 

River, [Kalimantan Selatan], eastern Borneo (Kalimantan), 

[Indonesia]. 

Tupaia natunae Lyon

Proc. Biol. Soc. Wash. 24: 168, 16 Jun 1911a.

= Tupaia splendidula natunae Lyon, 1911. See Medway (1961: 

271).

Holotype: USNM 104714. Skin and skull. Rather old adult 

female. Collected 27 Jun 1900 by W. L. Abbott. Original 

number 514.

Type Locality: Bunguran (Pulau Natuna Besar), Natuna Islands 

(Kepulauan Natuna Besar), [Kepulauan Riau, Indonesia]. 

Tupaia nicobarica surda G. S. Miller

Proc. U.S. Natl. Mus. 24: 774, 29 May 1902a.

Holotype: USNM 111757. Skin and skull. Adult male. Collected 

1 Mar 1901 by W. L. Abbott. Original number 899.

Type Locality: Little Nicobar Island, [Andaman and Nicobar Islands, India]. 

Tupaia pemangilis Lyon

Proc. Biol. Soc. Wash. 24: 168, 16 Jun 1911a.

= Tupaia glis (Diard, 1820). See Lyon (1913:48). Subspecies of 

T. glis not recognized by Helgen (2005:106).

 Holotype: USNM 112499. Skin and skull. Adult female. Collected 12 Jun 1901 by W. L. Abbott. Original number 1064.

Type Locality: Pulo Pemangil (Pulau Pemanggil), [Johor], off east 

coast of Malay Peninsula, [Malaysia]. 

Tupaia phaeura G. S. Miller

Proc. Acad. Nat. Sci. Phila. 54: 157, 11 Jun 1902b.

= Tupaia glis (Diard, 1820). See Chasen (1940:3). Subspecies of 

T. glis not recognized by Helgen (2005:106).

Holotype: USNM 113148. Skin and skull. Adult male. Collected 

4 Sep 1901 by W. L. Abbott. Original number 1275.
NUMBER 644 • 1 1

Type Locality: [FC: Sakana Bay], Sinkep Island (Pulau Singkep), 

[Lyon and Osgood (1909:253): Rhio Linga Archipelago 

(Kepulauan Lingga)], South China Sea, [Kepulauan Riau, 

Sumatra, Indonesia]. 

Tupaia picta fuscior Medway

J. Malay. Br. Roy. Asiat. Soc. 36 (part 3): 74, 1965.

Holotype: USNM 197198. Skin and skull. Adult male. Collected 

28 Jun 1913 by H. C. Raven. Original number [SL: 881].

Type Locality: Labuan Klambu (Labuhankelambu), east Kalimantan (Kalimantan Timur), Borneo (Kalimantan), 

[Indonesia]. 

Tupaia pulonis G. S. Miller

Smithson. Misc. Collect. 45(1): 56, 6 Nov 1903a.

= Tupaia glis (Diard, 1820). See Lyon (1913:47). Subspecies of 

T. glis not recognized by Helgen (2005:106).

 Holotype: USNM 112449. Skin and skull. Adult female. Collected 7 Jun 1901 by W. L. Abbott. Original number 1023.

Type Locality: Pulo Aor (Pulau Aur), [FC: South China Sea], off 

coast of Johore (Johor), [Malaysia]. 

Tupaia raviana Lyon

Proc. Biol. Soc. Wash. 24: 167, 16 Jun 1911a.

= Tupaia glis (Diard, 1820). See Chasen (1940:2). Subspecies of 

T. glis not recognized by Helgen (2005:106).

Holotype: USNM 104355. Skin and skull. Old adult male. Collected 18 Dec 1899 by W. L. Abbott. Orignial number 172.

Type Locality: Pulo Rawi (Ko Ra Wi), Butang Islands (Batang 

Group), off west coast of Malay Peninsula, [Satun, 

Thailand]. 

Tupaia riabus Lyon

Proc. U.S. Natl. Mus. 45: 88, 29 Nov 1913.

= Tupaia splendidulus riabus Lyon, 1913. See Helgen (2005:108).

Holotype: USNM 104881. Skin and skull. Adult female. Collected 

23 Aug 1900 by W. L. Abbott. Original number [SL: 636].

Type Locality: Pulo Riabu (Pulau Airabu), Anamba Islands (Kepulauan Anambas), [Kepulauan Riau, Indonesia]. 

Tupaia siaca Lyon

Proc. U.S. Natl. Mus. 34: 661, 14 Sep 1908.

= Tupaia ferruginea Raffles, 1821. See Sargis et al. (2013:201) 

for current use.

Holotype: USNM 144205. Skin and skull. Adult female. Collected 4 Nov 1906 by W. L. Abbott. Original number 4856.

Type Locality: Little Siak River (Sungai Siak Kecil), [Riau], eastern Sumatra, [Indonesia]. 

Tupaia sincepis Lyon

Proc. Biol. Soc. Wash. 24: 169, 16 Jun 1911a.

= Tupaia minor sincepis Lyon, 1911. See Lyon (1913:115).

Holotype: USNM 123105. Skin and skull. Adult male. Collected 

8 Aug 1903 by W. L. Abbott. Original number 2732.

Type Locality: Pulo (Pulau) Singkep, Rhio-Linga Archipelago, 

(Kepuluan Lingga), [Kepulauan Riau, Sumatra, Indonesia]. 

Tupaia sirhassenensis G. S. Miller

Proc. Wash. Acad. Sci. 3: 133, 26 Mar 1901a.

= Tupaia tana sirhassenensis G. S. Miller, 1901. See Chasen 

(1940:7).

Holotype: USNM 104712. Skin and skull. Adult male. Collected 

5 Jun 1900 by [MC: W. L. Abbott]. Original number 442.

Type Locality: Sirhassen Island (Pulau Serasan), Natunas (Kepulauan Natuna Selatan), [Kepulauan Riau, Indonesia]. 

Tupaia sordida G. S. Miller

Proc. Wash. Acad. Sci. 2: 231, 20 Aug 1900a.

= Tupaia glis (Diard, 1820). See Lyon (1913:48). Subspecies of 

T. glis not recognized by Helgen (2005:106).

Holotype: USNM 101747. Skin and skull. Adult male. Collected 

2 Oct 1899 by [MC: W. L. Abbott]. No original number.

Type Locality: Pulo (Pulau) Tioman, [Pahang, Malaysia]. 

Tupaia tephrura G. S. Miller

Smithson. Misc. Collect. 45(1): 57, 6 Nov 1903a.

= Tupaia glis (Diard, 1820). See Chasen (1940:3). Subspecies of 

T. glis not recognized by Helgen (2005:106).

Holotype: USNM 121752. Skin and skull. Adult female. Collected 12 Feb 1903 by W. L. Abbott. Original number 2276.

Type Locality: Tana Bala (Pulau Tanahbala), Batu Islands (Kepulauan Batu), [Sumatera Utara], Sumatra, [Indonesia]. 

Urogale cylindrura Mearns

Proc. U.S. Natl. Mus. 28: 435, 13 May 1905.

= Tupaia everetti Thomas, 1892. See Roberts et al. (2011:369).

Holotype: USNM 125287. Skin and skull. Adult male. Collected 

12 Jul 1904 by E. A. Mearns. Original number 5727.

Type Locality: Mount Apo, at the Bagobo village of Todaya, altitude 4,000 ft (1,219 m), [Davao del Sur], southern Mindanao, Philippine Islands (Philippines). 

Order Dermoptera Illiger, 1811

Family Cynocephalidae Simpson, 1945

Genus Galeopterus Thomas, 1908

Subspecies were not recognized by Stafford (2005:110).

Galeopithecus aoris G. S. Miller

Smithson. Misc. Collect. 45(1): 47, 6 Nov 1903a.

= Galeopterus variegatus (Audebert, 1799). See Chasen and 

Kloss (1929:20).

Holotype: USNM 112428. Skin and skull. Adult female. Collected 8 Jun 1901 by W. L. Abbott. Original number 1028.

Type Locality: Pulo Aor (Pulau Aur), off Coast of Johore (Johor), 

[Malaysia]. 
1 2 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Galeopithecus gracilis G. S. Miller

Smithson. Misc. Collect. 45(1): 49, 6 Nov 1903a.

= Galeopterus variegatus (Audebert, 1799). See Chasen (1935:24).

Holotype: USNM 104601. Skin and skull. Adult female. Collected 7 Jun 1900 by W. L. Abbott. Original number 461.

Type Locality: Sirhassen Island (Pulau Serasan), South Natunas 

(Kepulauan Natuna Selatan), [Kepulauan Riau, Indonesia]. 

Galeopithecus natunae G. S. Miller

Smithson. Misc. Collect. 45(1): 50, 6 Nov 1903a.

= Galeopterus variegatus (Audebert, 1799). See Chasen and 

Kloss (1928a:39).

Holotype: USNM 104602. Skin and skull. Adult female. Collected 16 Jul 1900 by W. L. Abbott. Original number 573.

Type Locality: Bunguran (Pulau Natuna Besar), North Natunas 

(Kepulauan Natuna Besar), [Kepulauan Riau, Indonesia]. 

Galeopithecus pumilus G. S. Miller

Smithson. Misc. Collect. 45(1): 46, 6 Nov 1903a.

= Galeopterus variegatus (Audebert, 1799). See Chasen and 

Kloss (1929:20).

Holotype: USNM 104448. Skin and skull. Adult male. Collected 

17 Dec 1899 by W. L. Abbott. Original number 165.

Type Locality: Pulo (Ko) Adang, Butang Islands (Batang Group), 

[Satun, Thailand]. 

Galeopithecus saturatus G. S. Miller

Smithson. Misc. Collect. 45(1): 51, 6 Nov 1903a.

= Galeopterus variegatus (Audebert, 1799). See Chasen (1940:20).

Holotype: USNM 121750. Skin and skull. Adult female. Collected 12 Feb 1903 by W. L. Abbott. Original number 2278.

Type Locality: Tana Bala (Pulau Tanahbala), Batu Islands (Kepulauan Batu), [Sumatera Utara], Sumatra, [Indonesia]. 

Galeopithecus tuancus G. S. Miller

Smithson. Misc. Collect. 45(1): 53, 6 Nov 1903a.

= Galeopterus variegatus (Audebert, 1799). See Chasen (1940:20).

Holotype: USNM 114375. Skin and skull. Adult female. Collected 22 Jan 1902 by W. L. Abbott. Original number 1454.

Type Locality: Pulo Tuanku (Pulau Tuangku), Banjak Islands 

(Kepulauan Banyak), [Aceh], Sumatra, [Indonesia]. 

Galeopterus abbotti Lyon

Proc. U.S. Natl. Mus. 40: 126, 25 Apr 1911b.

= Galeopterus variegatus (Audebert, 1799). See Chasen (1940:21).

Holotype: USNM 145577. Skin and skull. Adult female. Collected 

16 May 1907 by W. L. Abbott. Original number 5231.

Type Locality: Pulo (Pulau) Penebangan, [Kalimantan Barat], off 

west coast of Borneo (Kalimantan), [Indonesia]. 

Galeopterus borneanus Lyon

Proc. U.S. Natl. Mus. 40: 124, 25 Apr 1911b.

= Galeopterus variegatus (Audebert, 1799). See Chasen and 

Kloss (1929:18).

Holotype: USNM 151888. Skin, skull, [and skeleton]. Adult 

female. Collected 30 Jan 1908 by W. L. Abbott. Original 

number 5775.

Type Locality: Tjantung, [FC: upper river of that name (Sungai 

Cantung), 8 mi (13 km) from Klumpang Bay (Teluk Klumpang)], [Kalimantan Selatan], southeastern Borneo (Kalimantan), [Indonesia]. 

Galeopterus chombolis Lyon

Proc. U.S. Natl. Mus. 36: 486, 1 Jun 1909a.

= Galeopterus variegatus (Audebert, 1799). See Chasen and 

Kloss (1929:19).

Holotype: USNM 144375. Skin and skull. Adult female. Collected 3 Mar 1907 by W. L. Abbott. Original number 5091.

Type Locality: Pulo Jombol (Pulau Combol), Rhio-Linga Archipelago (Kepulauan Riau), [Kepulauan Riau, Sumatra, 

Indonesia]. 

Galeopterus lautensis Lyon

Proc. U.S. Natl. Mus. 40: 125, 25 Apr 1911b.

= Galeopterus variegatus (Audebert, 1799). See Chasen (1940:21).

Holotype: USNM 151886. Skin, skull, [and skeleton]. Adult female. Collected 23 Dec 1907 by [MC: W. L. Abbott]. Original number 5679.

Type Locality: [FC: Sungei (Sungai) Lima], Pulo (Pulau) Laut, 

[Kalimantan Selatan], off southeastern Borneo (Kalimantan), [Indonesia]. 

Order Primates Linnaeus, 1758

Family Lorisidae Gray, 1821

Genus Nycticebus E. Geoffroy, 1812

Nycticebus bancanus Lyon

Proc. U.S. Natl. Mus. 31: 536, 9 Nov 1906c.

See Munds et al. (2013:46) for current use.

Holotype: USNM 124907. Skin and skull. Adult female. Collected 24 Jun 1904 by W. L. Abbott. Original number 3432.

Type Locality: [FC: West side] Klabat Bay (Teluk Klabat), island 

of Banka (Pulau Bangka), [Kepulauan Bangka Belitung], 

east of Sumatra, [Indonesia]. 

Nycticebus borneanus Lyon

Proc. U.S. Natl. Mus. 31: 535, 9 Nov 1906c.

See Munds et al. (2013:46) for current use.

Holotype: USNM 142234. Skin and skull. Adult male. Collected 

15 Aug 1905 by W. L. Abbott. Original number 4322.

Type Locality: Along the Sakaiam River (Sungai Sekayam), a 

tributary of the Kapuas River (Sungai Kapuas), [Kalimantan 

Barat], western Borneo (Kalimantan), [Indonesia]. 

Nycticebus coucang brachycephalus Sody

Treubia 20: 145, 21 Dec 1949.
NUMBER 644 • 1 3

= Nycticebus coucang coucang (Boddaert, 1785). See Groves 

(1971:51).

Holotype: USNM 144328. [Skin and skull]. Adult female. Collected [SL: 15 Jan 1907 by W. L.] Abbott. Original number 

[SL: 4989].

Type Locality: Pulo Tebing Tinggi (Pulau Rantau), [Riau], east of 

Sumatra, [Indonesia].

Remarks: Sody (1949:145) did not give a specimen number for the 

type, but he gave all of the locality data given above that is not 

in brackets, and stated that this was the Nycticebus malaianus of Lyon (1908:669), in which Lyon also did not give a 

specimen number but referred a single specimen to Nycticebus malaianus and gave the locality as in the type description. 

USNM 144328 is the only specimen of Nycticebus collected 

by Abbott in the collection from this locality and therefore is 

the holotype by monotypy (ICZN, 1999: Article 73.1.2). 

Nycticebus coucang natunae Stone and Rehn

Proc. Acad. Nat. Sci. Phila. 54: 140, 4 Jun 1902.

= Nycticebus coucang coucang (Boddaert, 1785). See Groves 

(1971:51).

Holotype: USNM 104599. [Skin and skull]. [Lyon and Osgood 

(1909:282): Adult] male. Collected 28 Jul 1900 by W. L. 

Abbott. Original number [SL: 602].

Type Locality: Bunguran (Pulau Natuna Besar), Natuna Islands 

(Kepulauan Natuna Besar), [Kepulauan Riau, Indonesia]. 

Family Galagidae Gray, 1825

Genus Galago E. Geoffroy, 1796

Galago moholi cocos Heller

Smithson. Misc. Collect. 60(12): 1, 4 Nov 1912a.

= Galago cocos Heller, 1912. See Elliot (1913:257). See Butynski 

et al. (2006:21) for current use of cocos as a species.

Holotype: USNM 181810. [Skin and skull]. Adult male. Collected 16 Dec 1911 by E. Heller. Original number 4870.

Type Locality: Mazeras, [Coast Province], British East Africa 

(Kenya). 

Galago sotikae Hollister

Smithson. Misc. Collect. 72(2): 1, 22 Jan 1920.

= Galago senegalensis sotikae Hollister, 1920. See Schwarz 

(1931:55).

Holotype: USNM 142234. Skin and skull. Adult male. Collected 

22 May 1911 by E. Heller. Original number 1830.

Type Locality: Telek River, Sotik, [Rift Valley], British East Africa (Kenya). 

Family Tarsiidae Gray, 1825

Genus Carlito Groves and Shekelle, 2010

Tarsius fraterculus G. S. Miller

Proc. U.S. Natl. Mus. 38: 404, 19 Aug 1910.

= Carlito syrichta fraterculus (G. S. Miller, 1910). See Groves 

and Shekelle (2010:1074).

Holotype: USNM 239250. Skin, skull, and skeleton. Adult female. Collected 9 Apr 1906 by A. Celestino and M. Canton. 

Original number Philippine Bureau of Science 94.

Type Locality: Sevilla, [Bohol Province], Bohol [Island], 

[Philippines].

Remarks: The holotype was published as Philippine Bureau of 

Science number 94. It was cataloged as USNM 239250 on 

12 Feb 1923. 

Genus Cephalopachus Swainson, 1835

Tarsius borneanus Elliot

Bull. Am. Mus. Nat. Hist. 28: 153, 27 May 1910a.

= Cephalopachus bancanus borneanus (Elliot, 1910). See Groves 

and Shekelle (2010:1074).

Holotype: USNM 142242. [Skin and skull with carcass and 

brain in alcohol]. [Poole and Schantz (1942:252): Young 

adult] [SL: female]. Collected [SL: 10 Jul 1905 by W. L. Abbott. Original number 4259].

Type Locality: [FC: Batu Ampar], Landak River (Sungai 

Landak), [Kalimantan Barat], Borneo (Kalimantan), 

[Indonesia]. 

Tarsius saltator Elliot

Bull. Am. Mus. Nat. Hist. 28: 152, 27 May 1910a.

= Cephalopachus bancanus saltator (Elliot, 1910). See Groves 

and Shekelle (2010:1074).

Holotype: USNM 124992. [Skin and skull with carcass and 

brain in alcohol]. Adult female. Collected [SL: 7 Aug 1904 

by W. L. Abbott. Original number 3544].

Type Locality: [SL: Buding] [FC: Kampong], Billiton Island 

(Pulau Belitung), [Kepulauan Bangka Belitung, Sumatra, 

Indonesia]. 

Genus Tarsius Storr, 1780

Tarsius fuscus dentatus Miller and Hollister

Proc. Biol. Soc. Wash. 34: 103, 30 Jun 1921.

= Tarsius dentatus Miller and Hollister, 1921. See Groves 

(2005:127).

Holotype: USNM 218071. Skin and skull. Adult male. Collected 

15 Nov 1916 by H. C. Raven. Original number 2956.

Type Locality: Laboea Sore (Labuasore), north of Parigi, [Sulawesi Tengah], Celebes (Sulawesi), [Indonesia]. 

Tarsius pumilus Miller and Hollister

Proc. Biol. Soc. Wash. 34: 103, 30 Jun 1921.

Holotype: USNM 219454. Skin and skull. Adult female. Collected 31 Dec 1917 by H. C. Raven. Original number 

3366.

Type Locality: Rano Rano (Ranorano), [Sulawesi Tengah], Middle Celebes (Sulawesi), [Indonesia]. 
1 4 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Family Cebidae Bonaparte, 1831

Genus Cebus Erxleben, 1777

Cebus albifrons cesarae Hershkovitz

Proc. U.S. Natl. Mus. 98: 356, 10 May 1949.

See Ruiz-Garcia et al. (2010:1054) for current use.

Holotype: USNM 281588. [Skin and skull]. Old adult male. Collected 24 Aug 1942 by P. Hershkovitz. Original number 525.

Type Locality: Rio Guaimaral, a channel of the Rio Cesar, altitude 140 m, [SL: Valledupar District], Magdalena (El Cesar) 

Department, Colombia.

Remarks: Magdalena is still a Department, but the specimen was 

collected in El Cesar. 

Cebus albifrons pleei Hershkovitz

Proc. U.S. Natl. Mus. 98: 360, 10 May 1949.

See Ruiz-Garcia et al. (2010:1054) for current use.

Holotype: USNM 281624. Skin and skull. Male. Collected 29 

May 1943 by P. Hershkovitz. Original number 2029.

Type Locality: Swamplands of western bank of Rio Magdalena 

near village of Norosi at base of northern extremity of Cordillera Central, altitude approximately 50 m, Mompos, department of Bolivar, Colombia. 

Cebus capucinus limitaneus Hollister

Proc. Biol. Soc. Wash. 27: 105, 11 May 1914a.

= Cebus capucinus (Linnaeus, 1758). Subspecies not recognized 

by Groves (2005:137).

Holotype: USNM 16084/22845. Skin and skull. Adult male. 

Collected in 1887 by C. H. Townsend. No original number.

Type Locality: Segovia River (Rio Coco), eastern Honduras. Restricted by Hershkovitz (1949:347) to: Cabo Gracias a Dios 

at mouth of the Rio Segovia, eastern border between Honduras and Nicaragua. 

Genus Saguinas Hoffmannsegg, 1807

Midas elegantulus Slack

Proc. Acad. Nat. Sci. Phila. 13: 463, ordered printed 31 Dec 

1861. Receipt acknowledged 31 Mar 1862 by the Boston 

Society of Natural History.

= Saguinas labiatus labiatus E. Geoffroy, 1812. See Groves 

(2001:144).

Holotype: USNM 5132/37870. Skin [and skull]. Collected on unknown date by W. L. Herndon. [MC: Skin probably cataloged 

on 18 Nov 1861; skull on 12 Mar 1900]. No original number.

Type Locality: Amazon River and its tributaries, [Brazil?].

Remarks: The holotype was not designated by number in the 

original publication. Slack (1862:463) implied that he had 

only one specimen and that it belonged to the Smithsonian 

Institution. The specimen listed above is the only one in the 

collection bearing the data given in the original publication 

and is therefore the holotype by monotypy (ICZN 1999: 

Article 73.1.2). 

Genus Sapajus Kerr, 1792

Cebus margaritae Hollister

Proc. Biol. Soc. Wash. 27: 105, 11 May 1914a.

= Sapajus apaella margaritae (Hollister, 1914). New name combination. See Lynch Alfaro et al. (2012:272) for use of Sapajus.

Holotype: USNM 63219. Skin and skull. Old adult female. Collected 13 Jul 1895 by W. Robinson. Original number 520.

Type Locality: Margarita Island (Isla Margarita), [Nueva Esparta], Venezuela. 

Family Aotidae Elliot, 1913

Genus Aotus Illiger, 1811

Aotus zonalis Goldman

Smithson. Misc. Collect. 63(5): 6, 14 Mar 1914a.

See Defler and Bueno (2007:55) for current use.

Holotype: USNM 171231. Skin and skull. Adult female. Collected 29 Apr 1911 by E. A. Goldman. Original number 

[FC: Nelson and Goldman] 21101.

Type Locality: Gatun, altitude 100 ft (30 m), Canal Zone 

(Colon), Panama. 

Family Atelidae Gray, 1825

Genus Alouatta Lacepede, 1799

Alouatta palliata inconsonans Goldman

Smithson. Misc. Collect. 60(22): 17, 28 Feb 1913.

= Alouatta palliata (Gray, 1849). See Groves (2001:179).

Holotype: USNM 171068. [Skin and skull]. Adult male. Collected 23 Mar 1911 by E. A. Goldman. Original number 

[FC: Nelson and Goldman] 20995.

Type Locality: Cerro Azul, altitude 2,500 ft (762 m), near headwaters of the Chagres River, [Panama], Panama. 

Alouatta palliata mexicana Merriam

Proc. Biol. Soc. Wash. 15: 67, 22 Mar 1902a.

= Alouatta palliata (Gray, 1849). See Groves (2001:179).

Holotype: USNM 79398. [Skin and skull]. Adult male. Collected 

23 Apr 1896 by E. W. Nelson and E. A. Goldman. Original 

number 9551.

Type Locality: Minatitlan, [FC: altitude 100 ft (30 m)], Vera 

Cruz (Veracruz), Mexico. 

Genus Ateles E. Geoffroy, 1806

Ateles dariensis Goldman

Proc. Biol. Soc. Wash. 28: 101, 13 Apr 1915a.

= Ateles fuscipes rufiventris Sclater, 1872. See Groves (2001:188).

Holotype: USNM 179044. [Skin and skull]. Adult female. Collected 

29 Apr 1912 by E. A. Goldman. Original number 21664.

Type Locality: Near head of Rio Limon, altitude 5,200 ft (1,585 m), 

Mount Pirri (Cerro Pirre), [Darien], eastern Panama. 
NUMBER 644 • 1 5

Ateles geoffroyi panamensis Kellogg and Goldman

Proc. U.S. Natl. Mus. 96: 40, 2 Nov 1944.

= Ateles geoffroyi ornatus Gray, 1870. See Groves (2001:189).

Holotype: USNM 171489. Skin and skull. Adult female. Collected 8 Jun 1911 by E. A. Goldman. Original number [FC: 

Nelson and Goldman] 21165.

Type Locality: Cerro Bruja, about 15 mi (24 km) southeast of Portobello, altitude 2,000 ft (610 m), Province of Colon, Panama. 

Ateles geoffroyi yucatanensis Kellogg and Goldman

Proc. U.S. Natl. Mus. 96: 35, 2 Nov 1944.

Holotype: USNM 108531. Skin and skull. Adult male. Collected 

2 Apr 1901 by E. W. Nelson and E. A. Goldman. Original 

number 14652.

Type Locality: Puerto Morelos, altitude 100 ft (30 m), northeast 

coast of Quintana Roo, [Mexico]. 

Ateles tricolor Hollister

Proc. Biol. Soc. Wash. 27: 141, 10 Jul 1914c.

= Ateles geoffroyi vellerosus Gray, 1866. See Groves (2001:189).

Holotype: USNM 9427/8972. Skin and skull. Adult female. Collected 14 Jan 1869 by F. Sumichrast. Original number 42.

Type Locality: Santa Efigenia, [Kellogg and Goldman (1944:33): 

8 mi (13 km) north of Tapanatepec (San Pedro Tapanatepec)], Oaxaca, Mexico. 

Family Cercopithecidae Gray, 1821

Genus Cercopithecus Linnaeus, 1758

Lasiopyga albogularis kima Heller

Smithson. Misc. Collect. 61(17): 9, 21 Oct 1913a.

= Cercopithecus albogularis kibonotensis Lonnberg, 1908. See 

Dandelot (1974:20).

Holotype: USNM 182242. [Skin and skull]. Adult male. Collected 6 Nov 1911 by E. Heller. Original number 2555.

Type Locality: Mount Mbololo, Taita District, [Coast Province], 

British East Africa (Kenya). 

Lasiopyga albogularis maritima Heller

Smithson. Misc. Collect. 61(17): 8, 21 Oct 1913a.

= Cercopithecus albogularis kibonotensis Lonnberg, 1908. See 

Dandelot (1974:20).

Holotype: USNM 182272. [Skin and skull]. Adult female. Collected 17 Dec 1911 by E. Heller. Original number 2585.

Type Locality: Mazeras, [Coast Province], British East Africa (Kenya). 

Lasiopyga ascanius kaimosae Heller

Smithson. Misc. Collect. 61(17): 10, 21 Oct 1913a.

= Cercopithecus ascanius schmidti Matschie, 1892. See Dandelot 

(1974:22).

Holotype: USNM 182371. [Skin and skull]. Adult male. Collected 10 Feb 1912 by E. Heller. Original number 2692.

Type Locality: Upper Lukosa River, near the mission station of 

Kaimosi, [Rift Valley], British East Africa (Kenya). 

Lasiopyga leucampax [sic.] mauae Heller

Smithson. Misc. Collect. 61(17): 7, 21 Oct 1913a.

= Cercopithecus mitis stuhlmanni Matschie, 1893. See Dandelot 

(1974:19).

Holotype: USNM 173002. [Skin and skull]. Adult male. Collected 1 Nov 1910 by J. J. White. No original number.

Type Locality: Summit of Mau Escarpment, [SL: wagon road] 

between Londiana (Londiani) and Sirgoit, [Rift Valley], 

British East Africa (Kenya). 

Genus Chlorocebus Gray, 1870

Cercopithecus centralis luteus Elliot

Smithson. Misc. Collect. 56(7): 1, 11 Jun 1910b.

= Chlorocebus pygerythrus hilgerti (Neumann, 1902). See 

Groves (2001:202).

Holotype: USNM 163086. [Skin and skull]. [SL: Female]. Collected [SL: 12 Sep] [FC: 1909] by [SL: J. A. Loring]. Original 

number 7190.

Type Locality: Wambugu, [FC: altitude 5,300 ft (1,615 m)], 

southwest of Mount Kenia (Kenya), [Central Province], 

British East Africa (Kenya). 

Lasiopyga pygerythra arenaria Heller

Smithson. Misc. Collect. 61(17): 11, 21 Oct 1913a.

= Chlorocebus pygerythrus hilgerti (Neumann, 1902). See 

Groves (2001:202).

Holotype: USNM 182201. [Skin and skull]. Adult male. Collected 25 Jul 1911 by E. Heller. Original number 387.

Type Locality: Merille waterholes, Marsabit road, [North Eastern Province], British East Africa (Kenya). 

Lasiopyga pygerythra callida Hollister

Smithson. Misc. Collect. 59(3): 1, 2 Mar 1912a.

= Chlorocebus pygerythrus hilgerti (Neumann, 1902). See 

Groves (2001:202).

Holotype: USNM 162843. Skin, skull, [and appendicular skeleton]. Adult male. Collected 21 Jul 1909 by E. A. Mearns. 

Original number 6859.

Type Locality: South side of Lake Naivasha, [Rift Valley], British 

East Africa (Kenya). 

Lasiopyga pygerythra contigua Hollister

Smithson. Misc. Collect. 72(2): 2, 22 Jan 1920.

= Chlorocebus pygerythrus hilgerti (Neumann, 1902). See 

Groves (2001:202).

Holotype: USNM 163327. Skin and skull. Adult male. Collected 

30 Nov 1909 by E. A. Mearns. Original number 7321.

Type Locality: Changamwe, 6 mi (10 km) inland from Mombasa, [Coast Province], British East Africa (Kenya). 

Lasiopyga pygerythra tumbili Heller

Smithson. Misc. Collect. 61(17): 10, 21 Oct 1913a.

= Chlorocebus pygerythrus hilgerti (Neumann, 1902). See 

Groves (2001:202).
1 6 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Holotype: USNM 182229. [Skin and skull]. Adult male. Collected 1 Nov 1911 by E. Heller. Original number 2542.

Type Locality: Ndi, Taita District, [Coast Province], British East 

Africa (Kenya). 

Genus Colobus Illiger, 1811

Colobus abyssinicus percivali Heller

Smithson. Misc. Collect. 61(17): 6, 21 Oct 1913a.

= Colobus guereza percivali Heller, 1913. See Napier and Napier 

(1967:353).

Holotype: USNM 182138. [Skin and skull]. Adult male. Collected 22 Aug 1911 by E. Heller. Original number 2447.

Type Locality: [MC: North slope] Mount Uaragess (Warges), 

[MC: altitude 6,000 ft (1,829 m)], [Rift Valley], British East 

Africa (Kenya). 

Colobus abyssinicus roosevelti Heller

Smithson. Misc. Collect. 61(17): 5, 21 Oct 1913a.

= Colobus guereza matschiei Neumann, 1899. See Dandelot 

(1974:31).

Holotype: USNM 163261. [Skin and skull]. Adult male. Collected 6 Dec 1909 by T. Roosevelt. Original number 

E. Heller 543, not 513 as in original description.

Type Locality: Mau forest, near Enjoro (Njoro), [Rift Valley], 

British East Africa (Kenya). 

Colobus abyssinicus terrestris Heller

Smithson. Misc. Collect. 61(17): 7, 21 Oct 1913a.

= Colobus guereza occidentalis (Rochebrune, 1887). See Dandelot (1974:31).

Holotype: USNM 164756. [Skin, skull, and limb bones]. Adult 

female. Collected 20 Jan 1910 by K. Roosevelt. Original 

number E. Heller 623.

Type Locality: Rhino Camp, Lado Enclave, [Arua, Uganda]. 

Genus Erythrocebus Trouessart, 1897

Erythrocebus whitei Hollister

Smithson. Misc. Collect. 56(2): 11, 31 Mar 1910.

= Erythrocebus patas (Schreber, 1775). See Groves (2001:199).

Holotype: USNM 155440. Skin and skull. Adult male. Collected 

30 Sep 1908 by J. J. White. No original number.

Type Locality: Nzoia River, Guas Ngishu Plateau, [Western 

Province], British East Africa (Kenya). 

Genus Macaca Lacepede, 1799

Cynomolgus cagayanus Mearns

Proc. U.S. Natl. Mus. 28: 431, 13 May 1905.

= Macaca fascicularis fascicularis (Raffles, 1821). See Fooden 

(1991:22).

Holotype: USNM 125325. Skin and skull. Adult male. Collected 

25 Feb 1904 by E. A. Mearns. Original number 5771.

Type Locality: Cagayan Sulu Island, Sulu Sea, near Borneo, 

[Tawi Tawi Province], [SL: P. I.] (Philippines). 

Cynomolgus mindanensis Mearns

Proc. U.S. Natl. Mus. 28: 428, 13 May 1905.

= Macaca fascicularis philippensis (I. Geoffroy, 1843). See 

Groves (2001:228).

Holotype: USNM 123450. Skin and skull. Adult male. Collected 

26 Aug 1903 by E. A. Mearns. Original number 5620.

Type Locality: Pantar, altitude 1,900 ft (575 m), Mindanao, Philippine Islands (Philippines). 

Cynomolgus mindanensis apoensis Mearns

Proc. U.S. Natl. Mus. 28: 429, 13 May 1905.

= Macaca fascicularis philippensis (I. Geoffroy, 1843). See 

Groves (2001:228).

Holotype: USNM 125321. Skin and skull. Adult female. Collected 25 Jun 1904 by E. A. Mearns. Original number 5670.

Type Locality: Mount Apo, altitude 6,000 ft (1,829 m), southern 

Mindanao, Philippine Islands (Philippines). 

Cynomolgus suluensis Mearns

Proc. U.S. Natl. Mus. 28: 430, 13 May 1905.

= Macaca fascicularis fascicularis (Raffles, 1821). See Fooden 

(1995:72).

Holotype: USNM 125324. Skull without skin. Adult male. Collected 16 Nov 1903 by E. A. Mearns. Original number 

5750.

Type Locality: [SL and FC: Near the] foot of Crater Lake Mountain, on the Island of Sulu (Jolo), [Sulu Province], Philippine 

Islands (Philippines). 

Cynopithecus lembicus G. S. Miller

J. Mammal. 12: 413, 11 Nov 1931a.

= Macaca nigra (Desmarest, 1822). See Fooden (1969:71).

Holotype: USNM 217573. Skull only. Adult male. Collected 30 

May 1916 by H. C. Raven. Original number 2471.

Type Locality: Pulo (Pulau) Lembeh, [Sulawesi Utara], northeastern Celebes (Sulawesi), [Indonesia]. 

Macaca adusta G. S. Miller

Proc. U.S. Natl. Mus. 29: 559, 3 Feb 1906b.

= Macaca leonina (Blyth, 1863). See Groves (2001:223).

Holotype: USNM 124023. Skin and skull. Adult male. Collected 

22 Dec 1903 by W. L. Abbott. Original number 2929.

Type Locality: Champang, [FC: about 15 mi (24 km) north of 

Victoria Point], Tenasserim (Tanintharyi), [Burma]. 

Macaca broca G. S. Miller

Proc. U.S. Natl. Mus. 29: 558, 3 Feb 1906b.

= Macaca nemistrina Linnaeus, 1766. See Chasen (1940:65).

Holotype: USNM 19211/34930. Skin and skull. Adult male. 

Collected 21 Nov 1887 by C. F. Adams. Original number 

[MC: AA 111].
NUMBER 644 • 1 7

Type Locality: At Sapagaya River (Sungai Sapagaya), [Sabah], 

northeast Borneo, [Malaysia]. 

Macaca insulana G. S. Miller

Proc. U.S. Natl. Mus. 29: 560, 3 Feb 1906b.

= Macaca leonina (Blyth, 1863). See Groves (2001:223).

Holotype: USNM 104441. Skin and skull. Adult male. Collected 

1 Jan 1900 by W. L. Abbott. Original number 199.

Type Locality: Chance Island, Mergui Archipelago, [Tanintharyi, 

Burma]. 

Macaca irus atriceps Kloss

J. Nat. Hist. Soc. Siam 3: 347, 31 Dec 1919.

=Macaca fascicularis atriceps Kloss, 1919. See Napier and Napier (1967:349).

Holotype: USNM 236622. Skin and skull. Adult male. Collected 

30 Oct 1916 by C. B. Kloss. Original number 2283.

Type Locality: Koh Kram Island (Ko Khram), near Cape Liant, 

[Chon Buri], [SL: Inner Gulf of Siam], southeast Siam 

(Thailand).

Remarks: The holotype was published as CBK number 2283. It 

was cataloged as USNM 236622 on 17 Jul 1920. 

Macaca irus tua Kellogg

Proc. Biol. Soc. Wash. 57: 75, 31 Oct 1944.

= Macaca facicularis tua Kellogg, 1944. See Napier and Napier 

(1967:349).

Holotype: USNM 197663. Skin and skull. Adult male. Collected 

21 May 1913 by H. C. Raven. Original number 626.

Type Locality: Poelau Maratoea (Pulau Maratua), an island off 

the east coast of Borneo (Kalimantan), [Kalimantan Timur, 

Indonesia]. 

Macaca nemestrina indochinensis Kloss

J. Nat. Hist. Soc. Siam 3: 343, 31 Dec 1919.

= Macaca leonina (Blyth, 1863). See Groves (2001:223).

Holotype: USNM 236628. Skin and skull. Adult male. Collected 

12 Oct 1916 by C. B. Kloss. Original number 2148.

Type Locality: Lat Bua Kao (Lat Bua Khao), [Nakhon Ratchasima], east Siam (Thailand).

Remarks: The holotype was published as CBK number 2148. It 

was cataloged as USNM 236628 on 19 Jul 1920. 

Macacus fuscus G. S. Miller

Proc. U.S. Natl. Mus. 26: 476, 3 Feb 1903b.

= Macaca fascicularis fusca (G. S. Miller, 1903). See Fooden 

(1995:81).

Holotype: USNM 114164. Skin and skull. Old male. Collected 

20 Nov 1901 by W. L. Abbott. Original number 1348.

Type Locality: Simalur Island (Pulau Simeulue), [Aceh], northwestern Sumatra, [Indonesia]. 

Macacus pagensis G. S. Miller

Smithson. Misc. Collect. 45(1): 61, 6 Nov 1903a.

= Macaca pagensis (G. S. Miller, 1903). See G. S. Miller (1906b:557).

Holotype: USNM 121653. Skin and skull. Adult female. Collected 17 Nov 1902 by W. L. Abbott. Original number 2053.

Type Locality: South Pagi Island (Pulau Pagai Selatan), [Sumatera Barat], Sumatra, [Indonesia]. 

Macacus phaeura G. S. Miller

Smithson. Misc. Collect. 45(1): 63, 6 Nov 1903a.

= Macaca fascicularis fascicularis (Raffles, 1821). See Fooden 

(1995:72).

Holotype: USNM 121870. Skin and skull. Adult male. Collected 

20 Mar 1903 by W. L. Abbott. Original number 2399.

Type Locality: Siaba Bay (Teluk Siaba), Nias Island (Pulau Nias), 

[Sumatera Utara], Sumatra, [Indonesia]. 

Macacus pumilus G. S. Miller

Proc. Wash. Acad. Sci. 2: 241, 20 Aug 1900a.

= Macaca fascicularis fascicularis (Raffles, 1821). See Fooden 

(1995:71).

Holotype: USNM 101639. Skin and skull. Old adult male. Collected 6 Aug 1899 by [MC: W. L. Abbott]. No original number.

Type Locality: Pulo Bunoa (Pulau Benua), Tambelan Islands 

(Kepulauan Tambelan), [Kepulauan Riau, Indonesia]. 

Macacus rhesus villosus True

Proc. U.S. Natl. Mus. 17: 2, 8 May 1894a.

= Macaca mulatta (Zimmermann, 1780). See Pocock (1932:539).

Holotype: USNM 20120/35485. Skin and skull. Adult male. 

Collected 8 Sep 1891 by W. L. Abbott. Original number 

[MC: 4].

Type Locality: Lolab, altitude 7,500 ft (2,286 m), in the Vale of 

Kashmir [Jammu and Kashmir, India]. 

Macacus umbrosus G. S. Miller

Proc. U.S. Natl. Mus. 24: 789, 29 May 1902a.

= Macaca fascicularis umbrosa (G. S. Miller, 1902). See Fooden 

(1995:80).

Holotype: USNM 111795. Skin and skull. Adult male. Collected 

25 Feb 1901 by W. L. Abbott. Original number 888.

Type Locality: Little Nicobar Island, Nicobars, [Andaman and 

Nicobar Islands, India]. 

Pithecus agnatus Elliot

Proc. U.S. Natl. Mus. 38: 344, 6 Aug 1910c.

= Macaca fascicularis fascicularis (Raffles, 1821). See Fooden 

(1995:73).

Holotype: USNM 114409. [Skin and skull]. [Poole and Schantz 

(1942:241): Adult male]. Collected [SL: 26 Jan 1902 by 

W. L. Abbott. Original number 1471].

Type Locality: Pulo (Pulau) Tuangku, Banjak Islands (Kepulauan 

Banyak), [Aceh, Sumatra, Indonesia]. 

Pithecus baweanus Elliot

Proc. U.S. Natl. Mus. 38: 347, 6 Aug 1910c.
1 8 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

= Macaca fascicularis fascicularis (Raffles, 1821). See Fooden 

(1995:74).

Holotype: USNM 151829. [Skin and skull]. [Poole and Schantz 

(1942:241): Adult male]. Collected [SL: 24 Nov 1907 by 

W. L. Abbott. Original number 5565].

Type Locality: Bawean Island (Pulau Bawean), Java Sea, [Jawa 

Timur, Java, Indonesia]. 

Pithecus capitalis Elliot

Proc. U.S. Natl. Mus. 38: 350, 6 Aug 1910c.

= Macaca fascicularis fascicularis (Raffles, 1821). See Fooden 

(1995:75).

Holotype: USNM 83271. [Skin and skull]. [Poole and Schantz 

(1942:241): Adult] [SL: male]. Collected [SL: 7 Mar 1896 

by W. L. Abbott. Original number 105].

Type Locality: [MC: Prahmon], Trong (Trang), Lower Siam 

(Thailand). 

Pithecus carimatae Elliot

Proc. U.S. Natl. Mus. 38: 346, 6 Aug 1910c.

= Macaca fascicularis fascicularis (Raffles, 1821). See Medway 

(1965:84).

Holotype: USNM 125101. [Skin and skull]. [Poole and Schantz 

(1942:242): Adult] [SL: male]. Collected [SL: 24 Aug 1904 

by W. L. Abbott. Original number 3646].

Type Locality: Telok Pai, Karimata Islands (Kepulauan Karimata), 

[Kalimantan Barat, Borneo (Kalimantan), Indonesia]. 

Pithecus cupidus Elliot

Proc. U.S. Natl. Mus. 38: 348, 6 Aug 1910c.

= Macaca fascicularis fascicularis (Raffles, 1821). See Fooden 

(1995:74).

Holotype: USNM 151831. [Skin and skull]. [Poole and Schantz 

(1942:242): Adult] [SL: male]. Collected [SL: 8 Dec 1907 by 

W. L. Abbott. Original number 5584].

Type Locality: Pulo Mata Siri (Pulau Matasiri), 44 mi (71 km) 

from Pulo (Pulau) Laut, Java Sea, [Kalimantan Selatan, Borneo (Kalimantan), Indonesia]. 

Pithecus fuscus lasiae Lyon

Proc. U.S. Natl. Mus. 52: 453, 30 Dec 1916.

= Macaca fascicularis lasiae (Lyon, 1916). See Fooden (1995:81).

Holotype: USNM 114248. Skin and skull. Adult male. Collected 

5 Jan 1902 by W. L. Abbott. Original number 1398.

Type Locality: Pulo (Pulau) Lasia, [FC: 14 mi (23 km) SE of Simalur Island (Pulau Simeulue)], [Aceh, Sumatra, Indonesia]. 

Pithecus impudens Elliot

Proc. U.S. Natl. Mus. 38: 350, 6 Aug 1910c.

= Macaca fascicularis fascicularis (Raffles, 1821). See Napier 

(1981:13).

Holotype: USNM 115675. [Skin and skull]. [Poole and Schantz 

(1942:242): Adult] [SL: male]. Collected [SL: 24 Aug 1902 

by W. L. Abbott. Original number 1956].

Type Locality: Pulo (Pulau) Sugi, Rhio Archipelago (Kepulauan 

Riau), [Kepulauan Riau, Sumatra, Indonesia]. 

Pithecus lapsus Elliot

Proc. U.S. Natl. Mus. 38: 343, 6 Aug 1910c.

= Macaca fascicularis fascicularis (Raffles, 1821). See Fooden 

(1995:73).

Holotype: USNM 124863. [Skin and skull]. [Poole and Schantz 

(1942:243): Adult] [FC: male]. Collected [SL: 19 Jun 1904 

by W. L. Abbott. Original number 3418].

Type Locality: [FC: Tanjong Pamuja, north coast of] Island of 

Banka (Pulau Bangka), [Kepulauan Bangka Belitung], east 

of southern Sumatra, [Indonesia]. 

Pithecus lautensis Elliot

Proc. U.S. Natl. Mus. 38: 345, 6 Aug 1910c.

= Macaca fascicularis fascicularis (Raffles, 1821). See Fooden 

(1995:74).

Holotype: USNM 104854. [Skin and skull]. [Poole and Schantz 

(1942:243): Adult] [SL: male]. Collected [SL: 9 Aug 1900] 

by [MC: W. L. Abbott]. Original number [SL: 614].

Type Locality: Pulo (Pulau) Laut, Natuna Islands (Kepulauan 

Natuna Besar), [Kepulauan Riau, Indonesia]. 

Pithecus lingae Elliot

Proc. U.S. Natl. Mus. 38: 349, 6 Aug 1910c.

= Macaca fascicularis fascicularis (Raffles, 1821). See Fooden 

(1995:75).

Holotype: USNM 101603. [Skin and skull]. [Poole and Schantz 

(1942:243): Adult] [SL: male]. Collected [SL: 23 Jul 1899] 

by [MC: W. L. Abbott]. No original number.

Type Locality: Linga Island (Pulau Lingga), Rhio Archipelago 

(Kepulauan Lingga), [Kepulauan Riau, Sumatra, Indonesia]. 

Pithecus lingungensis Elliot

Proc. U.S. Natl. Mus. 38: 344, 6 Aug 1910c.

= Macaca fascicularis fascicularis (Raffles, 1821). See Fooden 

(1995:74).

Holotype: USNM 104853. [Skin and skull]. [Poole and Schantz 

(1942:243): Adult] [SL: male]. Collected [SL: 19 Jun 1900] 

by [MC: W. L. Abbott]. Original number [SL: 492].

Type Locality: Pulo (Pulau) Lingung, [SL and FC: near Bunguran 

(Pulau Natuna Besar)], Natuna Islands (Kepulauan Natuna 

Besar), [Kepulauan Riau, Indonesia]. 

Pithecus mandibularis Elliot

Proc. U.S. Natl. Mus. 38: 347, 6 Aug 1910c.

= Macaca fascicularis fascicularis (Raffles, 1821). See Medway 

(1965:84).

Holotype: USNM 142225. [Skin and skull]. [Poole and Schantz 

(1942:244): Adult] [SL: male]. Collected [SL: 18 Jun 1905 

by W. L. Abbott. Original number 4196].

Type Locality: Sungei Sama, near Pontianak, [Kalimantan Barat], 

Borneo (Kalimantan), [Indonesia]. 
NUMBER 644 • 1 9

Pithecus mansalaris Lyon

Proc. U.S. Natl. Mus. 52: 452, 30 Dec 1916.

= Macaca fascicularis fascicularis (Raffles, 1821). See Napier 

(1981:13).

Holotype: USNM 114560. Skin and skull. Adult male. Collected 

10 Mar 1902 by W. L. Abbott. Original number 1639.

Type Locality: Mansalar Island (Pulau Musala), [Sumatera 

Utara], [FC: west coast of Sumatra], [Indonesia]. 

Pithecus mindorus Hollister

Proc. U.S. Natl. Mus. 46: 328, 31 Dec 1913a.

= Macaca fascicularis philippensis (I. Geoffroy, 1843). See 

Fooden (1991:23).

Holotype: USNM 144674. Skin and skull. Adult male. Collected 

Nov 1906 by E. A. Mearns. Original number 6301.

Type Locality: Alag River, [SL and FC: Camp No. 2], [Mindoro 

Oriental Province], Mindoro, [SL and FC: P. I. (Philippines)]. 

Pithecus pullus A. B. Howell

Proc. Biol. Soc. Wash. 41: 41, 16 Mar 1928a.

= Macaca thibetana esau (Matschie, 1912). See Groves (2005:164).

Holotype: USNM 252157. Skin and skull. Young adult male. 

Collected on unknown date by F. T. Smith. Original number 

85. [MC: Cataloged 1 Jul 1927].

Type Locality: Kuatun, Fukien (Fujian), China. 

Pithecus sirhassenensis Elliot

Proc. U.S. Natl. Mus. 38: 345, 6 Aug 1910c.

= Macaca fascicularis fascicularis (Raffles, 1821). See Fooden 

(1995:74).

Holotype: USNM 104852. [Skin and skull]. [Poole and Schantz 

(1942:245): Adult] [SL: male]. Collected [SL: 8 Jun 1900 by 

W. L. Abbott. Original number 468].

Type Locality: Sirhassen Island (Pulau Serasan), Natuna Islands 

(Kepulauan Natuna Selatan), [Kepulauan Riau, Indonesia]. 

Pithecus vitis Elliot

Proc. U.S. Natl. Mus. 38: 346, 6 Aug 1910c.

= Macaca fascicularis aurea (E. Geoffroy, 1831). See Napier 

(1981:13).

Holotype: USNM 124176. [Skin and mandibles]. [Poole and 

Schantz (1942:246): Adult] [SL: male]. Collected [SL: 26 

Jan 1904 by W. L. Abbott. Original number 3076].

Type Locality: Domel Island (Letsok-aw Kyun), Mergui Archipelago, [Tanintharyi, Burma]. 

Genus Papio Erxleben, 1777

Papio anubis lestes Heller

Smithson. Misc. Collect. 61(19): 10, 8 Nov 1913b.

= Papio anubis (Lesson, 1827). Subspecies not recognized by 

Groves (2005:166).

Holotype: USNM 164633. [Skin and skull]. Adult male. Collected 19 Nov 1909 by J. A. Loring. Original number 8234.

Type Locality: Ulukenia (Lukenya) Hills, Athi Plains, British East 

Africa (Kenya). 

Papio anubis vigilis Heller

Smithson. Misc. Collect. 61(19): 11, 8 Nov 1913b.

= Papio anubis (Lesson, 1827). Subspecies not recognized by 

Groves (2005:166).

Holotype: USNM 182033. [Skin and skull]. Old male. Collected 

10 Jul 1911 by E. Heller. Original number 2337.

Type Locality: Lakiundu River, near its junction with the Northern Guaso Nyiro, British East Africa (Kenya). 

Genus Presbytis Eschscholtz, 1821

Presbytes batuanus G. S. Miller

Smithson. Misc. Collect. 45(1): 65, 6 Nov 1903a.

= Presbytis melalophos sumatrana (Muller and Schlegel, 1841). 

See Groves (2001:276).

Holotype: USNM 121810. Skin and skull. Adult male. Collected 

4 Mar 1903 by W. L. Abbott. Original number 2369.

Type Locality: Pulo Pinie (Pulau Pini), Batu Isalnds (Kepulauan 

Batu), [Sumatera Utara], Sumatra, [Indonesia]. 

Presbytes rhionis G. S. Miller

Smithson. Misc. Collect. 45(1): 64, 6 Nov 1903a.

= Presbytis siamensis rhionis (G. S. Miller, 1903). See Groves 

(2001:280).

Holotype: USNM 115665. Skin and skull. Adult female. Collected 15 Aug 1902 by W. L. Abbott. Original number 1888.

Type Locality: Telok Pemudong, Pulo Bintang (Pulau Bintan), 

Rhio Archipelago (Kepulauan Riau), [Kepulauan Riau, Sumatra, Indonesia]. 

Presbytis arwasca G. S. Miller

J. Mammal. 15: 126, 15 May 1934a.

= Presbytis chrysomelas cruciger (Thomas, 1892). See Groves 

(2001:278).

Holotype: USNM 83953. Skin and skull. Young adult male. Collected Aug 1894 by E. and C. Hose. No original number.

Type Locality: Miri, Sarawak, Borneo, [Malaysia]. 

Presbytis australis G. S. Miller

Smithson. Misc. Collect. 61(21): 28, 29 Dec 1913b.

= Presbytis femoralis femoralis (Martin, 1838). See Pocock 

(1934:901).

Holotype: USNM 112709. Skin and skull. Adult male. Collected 

31 Jul 1901 by W. L. Abbott. Original number 1196.

Type Locality: Jambu Luang, east coast of Johore (Johor), 

[Malaysia]. 

Presbytis cana G. S. Miller

Proc. U.S. Natl. Mus. 31: 275, 11 Sep 1906c.

= Presbytis siamensis cana G. S. Miller, 1906. See Groves 

(2001:280).
2 0 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Holotype: USNM 122915. Skin and skull. Adult male. Collected 

28 Jun 1903 by W. L. Abbott. Original number 2558.

Type Locality: Pulo (Pulau) Kundur, Rhio Archipelago (Kepulauan Riau), [Kepulauan Riau, Sumatra, Indonesia]. 

Presbytis canicrus G. S. Miller

Proc. Biol. Soc. Wash. 47: 15, 9 Feb 1934b.

= Presbytis hosei canicrus G. S. Miller, 1934. See Medway 

(1977:67).

Holotype: USNM 198282. Skin and skull. Young adult female. 

Collected 25 Nov 1913 by H. C. Raven. Original number 

1137.

Type Locality: [FC: Gunong Batu], on the Karangan River (Sungai Karangan), [Kalimantan Timur], Dutch Northeast Borneo (Kalimantan), [Indonesia]. 

Presbytis carimatae G. S. Miller

Proc. U.S. Natl. Mus. 31: 65, 23 Jul 1906a.

= Presbytis rubicunda carimatae G. S. Miller, 1906. See Medway 

(1965:82).

Holotype: USNM 125157. Skin, skull, [and brain in fluid]. Adult 

female. Collected [SL and FC: 2 Sep 1904], not 21 Aug as in 

original description, by W. L. Abbott. Original number [SL 

and FC: 3717], not 3627 as in original description.

Type Locality: Telok Edar, Karimata Island (Pulau Karimata), 

[Kalimantan Barat, Borneo (Kalimantan), Indonesia]. 

Presbytis catemana Lyon

Proc. U.S. Natl. Mus. 34: 672, 14 Sep 1908.

= Presbytis siamensis cana G. S. Miller, 1906. See Groves 

(2001:280).

Holotype: USNM 123149. Skin, skull [and brain in fluid]. Adult 

female. Collected [SL and FC: 28 Aug 1903], not 23 as in 

original description, by W. L. Abbott. Original number 2762.

Type Locality: Along the Kateman River (Sungai Kateman), 

[Riau], eastern Sumatra, [Indonesia]. 

Presbytis percura Lyon

Proc. U.S. Natl. Mus. 34: 671, 14 Sep 1908.

= Presbytis femoralis percura Lyon, 1908. See Wilson and Wilson (1977:220).

Holotype: USNM 144088. Skin and skull. Adult male. Collected 

26 Feb 1907 by W. L. Abbott. Original number 5083.

Type Locality: Kompei, eastern Sumatra, [Indonesia]. 

Pygathrix rubicunda rubida Lyon

Proc. U.S. Natl. Mus. 40: 139, 25 Apr 1911b.

= Presbytis rubicunda rubida (Lyon, 1911). See Medway (1965:81).

Holotype: USNM 153790. Skin, skull, skeleton, [and brain in 

fluid]. Adult female. Collected [SL and FC: 18 Jun 1908], 

not 17 as in original description, by W. L. Abbott. Original 

number 5979.

Type Locality: [FC: Tanjong] Batu Jurong, [Kalimantan Barat], 

southwestern Borneo, (Kalimantan), [Indonesia]. 

Genus Simias Miller, 1903

Simias concolor G. S. Miller

Smithson. Misc. Collect. 45(1): 67, 6 Nov 1903a.

Holotype: USNM 121659. Skin and skull. Adult male. Collected 

3 Dec 1902 by W. L. Abbott. Original number 2103.

Type Locality: [FC: Small island close to the south shore of Vekens Bay], South Pagi Island (Pulau Pagai Selatan), [Sumatera 

Barat], Sumatra, [Indonesia]. 

Genus Trachypithecus Reichenbach, 1862

Presbytis argenteus Kloss

J. Nat. Hist. Soc. Siam 3: 338, 31 Dec 1919.

= Trachypithecus phayrei crepusculus (Elliot, 1909). See Groves 

(2001:268).

Syntypes: USNM 236624. Skin and skull. Adult female. Collected 8 Oct 1916 by C. B. Kloss. Original number 2127. 

USNM 236625. Skin and skull. Adult male. Collected 11 

Oct 1916 by C. B. Kloss. Original number 2144.

Type Locality: Lat Bua Kao (Lat Bua Khao), [Nakon Ratchasima], East Siam (Thailand). [Pocock (1935:950): modified 

the locality to read = 30 mi (48 km) W. of Korat, some 100 

mi (161 km) N. E. of Bangkok in S. W. Siam, not in E. Siam].

Remarks: Kloss (1919:338) identified the two specimens listed 

above as types in his original description and they are therefore syntypes (ICZN: 1999, Article 73.2.1). Both specimens 

were published under their original numbers; they were cataloged and assigned their USNM numbers on 19 Jul 1920. 

Note: the type locality is in fact in eastern Thailand, contrary to Pocock’s assertion. 

Presbytis corvus G. S. Miller

Smithson. Misc. Collect. 61(21): 27, 29 Dec 1913b.

= Trachypithecus obscurus carbo (Thomas and Wroughton, 

1909). See Pocock (1935:947).

Holotype: USNM 123993. Skin and skull. Adult male. Collected 

10 Nov 1903 by W. L. Abbott. Original number 2852.

Type Locality: Pulo Terutau (Ko Tarutao), west coast of Malay 

Peninsula, [Satun, Thailand]. 

Presbytis cristata koratensis Kloss

J. Nat. Hist. Soc. Siam 3: 340, 31 Dec 1919.

= Trachypithecus germaini germaini (Milne-Edwards, 1876). See 

Groves (2001:265).

Holotype: USNM 236627. Skin and skull. Adult female. Collected [SL: 10] Oct 1916 by C. B. Kloss. Original number 

2136.

Type Locality: Lat Bua Kao (Lat Bua Khao), [Nakon Ratchasima], East Siam (Thailand). [Pocock (1935:950): modified 

the locality to read = 30 mi (48 km) W. of Korat, some 100 

mi (161 km) N. E. of Bangkok in S. W. Siam, not in E. Siam].

Remarks: The holotype was published as CBK number 2136; it 

was cataloged and assigned USNM 236627 on 19 Jul 1920. 
NUMBER 644 • 2 1

Note: the type locality is in fact in eastern Thailand, contrary to Pocock’s assertion. 

Presbytis vigilans G. S. Miller

Smithson. Misc. Collect. 61(21): 29, 29 Dec 1913b.

= Trachypithecus cristatus vigilans (G. S. Miller, 1913). See 

Groves (2001:265).

Holotype: USNM 104840. Skull only. Adult female. Collected 2 

Jun 1900 by W. L. Abbott. Original number 422.

Type Locality: Sirhassen Island (Pulau Serasan), South Natuna Islands 

(Kepulauan Natuna Selatan), [Kepulauan Riau, Indonesia]. 

Pygathrix flavicauda Elliot

Proc. U.S. Natl. Mus. 38: 352, 6 Aug 1910c.

= Trachypithecus obscurus flavicaudus (Elliot, 1910). See Pocock 

(1935:942).

Holotype: USNM 83259. [Skin and skull]. [Poole and Schantz 

(1942:249): Adult] [SL: female]. Collected [SL: 27 Mar 

1896 by W. L. Abbott]. No original number.

Type Locality: [MC: Prahmon], Trong (Trang), Lower Siam 

(Thailand). 

Pygathrix sanctorum Elliot

Proc. U.S. Natl. Mus. 38: 351, 6 Aug 1910c.

= Trachypithecus obscurus sanctorum (Elliot, 1910). See Groves 

(2001:268).

Holotype: USNM 104446. [Skin and skull]. [Poole and Schantz 

(1942:250): Adult] [SL: female]. Collected [SL: 16 Jan 1900, 

not Jun as in original description, by W. L. Abbott. Original 

number 235].

Type Locality: St. Matthew (Za Det Gyi) Island, Mergui Archipelago, [Tanintharyi, Burma]. 

Pygathrix ultima Elliot

Proc. U.S. Natl. Mus. 38: 351, 6 Aug 1910c.

= Trachypithecus cristatus cristatus (Raffles, 1821). See Groves 

(2001:265).

Holotype: USNM 83949. [Skin and skull]. [Poole and Schantz 

(1942:250): Adult] [SL: male]. Collected [SL: Jun 1895] by 

E. and C. Hose. No original number.

Type Locality: Mount (Bukit) Dulit, elevation 3,000 ft (914 m), 

[Sarawak], Borneo, [Malaysia]. 

Semnopithecus (Trachypithecus) (auratus) auratus ebenus

Brandon-Jones

Raffles Bull. Zool. 41: 15, 30 Jun 1995.

= Trachypithecus ebenus (Brandon-Jones, 1995). See Groves 

(2001:273).

Holotype: USNM 240489. Flat skin, skull, [and hand and feet 

bones]. Adult female. Collected in 1924 by F. R. Wulsin. No 

original number.

Type Locality: [SL: French Indochina], possibly from the vicinity 

of Lai Chau or, perhaps more probably, from the Fan Si Pan 

mountain chain at 22o30 ́N 103o50 ́E, Vietnam. 

Family Hylobatidae Gray, 1871

Genus Hylobates Illiger, 1811

Hylobates cinereus abbotti Kloss

Proc. Zool. Soc. London 1929: 119, 30 Apr 1929.

= Hylobates muelleri abbotti Kloss, 1929. See Medway (1977:74).

Holotype: USNM 142172. Skin, skull, [long bones, and brain in 

fluid]. Adult male. Collected 29 Jun 1905 by W. L. Abbott. 

Original number 4252.

Type Locality: Sungei Naja (Sungai Naya), Landak River (Sungai 

Landak), Pontianak, [Kalimantan Barat], southwest Borneo, (Kalimantan), [Indonesia]. 

Hylobates mulleri albibarbis Lyon

Proc. U.S. Natl. Mus. 40: 142, 25 Apr 1911b.

= Hylobates albibarbis Lyon, 1911. See Groves (2001:291).

Holotype: USNM 145327. Skin and skull. Adult male. Collected 

16 Aug 1907 by W. L. Abbott. Original number 5501.

Type Locality: Along the Matan River (Sungai Matan), a tributary of the Sempang (Sungai Simpang), [Kalimantan Barat], 

[SL: west Borneo (Kalimantan)], [Indonesia]. 

Symphalangus klossii G. S. Miller

Smithson. Misc. Collect. 45(1): 70, 6 Nov 1903a.

= Hylobates klossii (G. S. Miller, 1903). See Chasen and Kloss 

(1928b:808).

Holotype: USNM 121678. Skin and skull. Adult male. Collected 

13 Nov 1902 by W. L. Abbott. Original number 2032.

Type Locality: [FC: Sikakap Strait (Selat Sikakap)], South Pagi 

Island (Pulau Pagai Selatan), [Sumatera Barat], [FC: west] 

Sumatra, [Indonesia]. 

Order Lagomorpha Brandt, 1855

Family Ochotonidae Thomas, 1897

Genus Ochotona Link, 1795

Lagomys collaris Nelson

Proc. Biol. Soc. Wash. 8: 117, 21 Dec 1893.

= Ochotona collaris (Nelson, 1893). See Trouessart (1897:648).

Holotype: USNM 14384/36297. Skin and skull. Adult. Collected [SL: summer of] 1880 by E. W. Nelson. Original 

number 164.

Type Locality: Near the head of the Tanana River, about 200 mi 

(322 km) south of Fort Yukon, Alaska. 

Lagomys schisticeps Merriam

N. Am. Fauna 2: 11, 30 Oct 1889a.

= Ochotona princeps schisticeps (Merriam, 1889). See A. H. 

Miller (1936:175). 

Holotype: USNM 186517. Skin and skull. Adult male. Collected 

9 Jun 1888 by C. A. Allen. Original number [MCC: 265]. 

Merriam collection number 4725/5376.
2 2 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Type Locality: Donner, [Placer County], California.

Remarks: The holotype was published as the Merriam collection 

number; it was cataloged USNM 186517 on 23 Oct 1911. 

Ochotona (Pika) alpina argentata A. B. Howell

Proc. Biol. Soc. Wash. 41:116, 29 Jun 1928b.

= Ochotona argentata A. B. Howell, 1928. See Erbajeva 

(1997:505).

Holotype: USNM 240726. Skin, skull, and skeleton. Adult female. 

Collected 16 May 1923 by F. R. Wulsin. Original number 1059.

Type Locality: 15 mi (24 km) northwest of Ninghsia (Yinchuan), 

northern Kansu (Ningxia), China. 

Ochotona annectens G. S. Miller

Proc. Biol. Soc. Wash. 24: 54, 24 Feb 1911a.

= Ochotona cansus annectens G. S. Miller, 1911. See Smith 

(2008:278) for current use.

Holotype: USNM 155164. Skin and skull. Adult male. Collected 

27 Jul 1909 by A. de C. Sowerby. Original number 225.

Type Locality: 15 mi (24 km) northeast of Ching-Ning-Chow, 

altitude 6,200 ft (1,890 m), Kansu (Gansu), China. 

Ochotona cansus Lyon

Smithson. Misc. Collect. 50(14): 136, 9 Jul 1907b.

= Ochotona cansus cansus Lyon, 1907.

Holotype: USNM 144030. Skin and skull. Adult male. Collected 

8 Jun 1906 by W. W. Simpson. Original number 13.

Type Locality: Taocheo [Hoffmann and Smith (2005:186): 

(Lintan)], Kan-su (Gansu), China. 

Ochotona (Ochotona) erythrotis vulpina A. B. Howell

Proc. Biol. Soc. Wash. 41: 117, 29 Jun 1928b.

= Ochotona erythrotis (Buchner, 1890).

Holotype: USNM 240723. Skin and skull. Adult Male. Collected 

12 Aug [SL and FC: 1923], not 1927 as in original description, by F. R. Wulsin. Original number 1103.

Type Locality: 30 mi (48 km) west of Sining (Xining), Kansu 

(Qinghai), China. 

Ochotona fenisex brunnescens A. H. Howell

Proc. Biol. Soc. Wash. 32: 108, 20 May 1919.

= Ochotona princeps fenisex Osgood, 1913. See Hafner and 

Smith (2010:411) for current use.

Holotype: USNM 227259. Skin and skull. Subadult male. Collected 23 Aug 1917 by G. G. Cantwell. Original number 38. 

19568-X.

Type Locality: Keechelus, [Kittitas County], Washington. 

Ochotona fenisex fumosa A. H. Howell

Proc. Biol. Soc. Wash. 32: 109, 20 May 1919.

= Ochotona princeps fenisex Osgood, 1913. See Hafner and 

Smith (2010:411) for current use.

Holotype: USNM 91144. Skin and skull. Adult male. Collected 

4 Oct 1897 by J. A. Loring. Original number 4799.

Type Locality: Permilia (Pamelia) Lake, west base Mount Jefferson, [Linn County], Oregon. 

Ochotona levis Hollister

Proc. Biol. Soc. Wash. 25: 57, 13 Apr 1912b.

= Ochotona princeps princeps (Richardson, 1828). See Hoffmann and Smith (2005:191).

Holotype: USNM 12000/22241. Skin and skull. Adult female. 

Collected 24 Aug 1874 by E. Coues. Original number 4593.

Type Locality: Chief Mountain Lake (Waterton Lake), [Glacier 

County], Montana. 

Ochotona nigrescens Bailey

Proc. Biol. Soc. Wash. 26: 133, 21 May 1913.

= Ochotona princeps saxatilis Bangs, 1899. See Hoffmann and 

Smith (2005:192).

Holotype: USNM 147976. [Skin and skull]. Adult male. Collected 28 Aug 1906 by V. Bailey. Original number 8447.

Type Locality: Jemez Mountains, altitude [FC: about] 10,000 ft 

(3,048 m), [Sandoval County], New Mexico. Restricted by 

Bailey (1931:67) within the above locality to “on Goat Peak 

at the head of Santa Clara Creek.” 

Ochotona nitida Hollister

Smithson. Misc. Collect. 60(14): 4, 29 Nov 1912c.

= Ochotona alpina alpina (Pallas, 1773). See Hoffmann and 

Smith (2005:185).

Holotype: USNM 175390. Skin and skull. Adult female. Collected 8 Jul 1912 by N. Hollister. Original number 4257.

Type Locality: Tchegan-Burgazi Pass, Altai Mountains, near 

Mongolian border, south of Kosh-Agatch (Kosh-Agach), 

[Altayskiy], Siberia (Russia). 

Ochotona princeps brooksi A. H. Howell.

N. Am. Fauna 47: 30, 23 Sep 1924.

= Ochotona princeps fenisex Osgood, 1913. See Hafner and 

Smith (2010:411) for current use.

Holotype: USNM 69275. Skin and skull. Adult male. Collected 

24 Sep 1894 by C. P. Streator. Original number 4319.

Type Locality: Sicamous, [FC: altitude 1,200 ft (366 m)], British 

Columbia, [Canada]. 

Ochotona princeps lutescens A. H. Howell

Proc. Biol. Soc. Wash. 32: 105, 20 May 1919.

= Ochotona princeps princeps (Richardson, 1828). See Hoffmann and Smith (2005:191).

Holotype: USNM 108650. Skin and skull. Adult male. Collected 

26 Jul 1899 by G. F. Dippie. Original number 193. 2973-X.

Type Locality: Mount Inglesmaldie (Inglismaldie), near Banff, 

[Miller and Kellogg (1955:136): altitude about 8,000 ft 

(2,438 m)], Alberta, Canada. 

Ochotona princeps moorei M. C. Gardner

J. Wash. Acad. Sci. 40: 344, 15 Oct 1950.
NUMBER 644 • 2 3

= Ochotona princeps uinta Hollister, 1912. See Hafner and 

Smith (2010:412) for current use.

Holotype: USNM 248436. Skin and skull. Adult female. Collected 18 Jul 1928 by A. W. Moore. Original number 470. 

[SL: 24928-X].

Type Locality: Manti National Forest, 1 mi (1.6 km) northeast of 

Baldy Ranger Station, altitude 10,000 ft (3,048 m), Sanpete 

County, Utah. 

Ochotona saxatilis incana A. H. Howell

Proc. Biol. Soc. Wash. 32: 107, 20 May 1919.

= Ochotona princeps saxatilis Bangs, 1899. See Hoffmann and 

Smith (2005:195).

Holotype: USNM 128914. Skin and skull. Adult female. Collected 10 Aug 1903 by V. Bailey. Original number 8079.

Type Locality: Pecos Baldy, altitude 12,000 ft (3,658 m), [Santa 

Fe County], New Mexico. 

Ochotona schisticeps fuscipes A. H. Howell

Proc. Biol. Soc. Wash. 32: 110, 20 May 1919.

= Ochotona princeps schisticeps (Merriam, 1889). See Hafner 

and Smith (2010:411) for current use.

Holotype: USNM 158094. Skin and skull. Adult male. Collected 

7 Sep 1908 by W. H. Osgood. Original number 3475.

Type Locality: Brian Head, Parowan Mountains, [FC: Iron 

County], Utah. 

Ochotona schisticeps goldmani A. H. Howell

N. Am. Fauna 47: 40, 23 Sep 1924.

= Ochotona princeps princeps (Richardson, 1828). See Hoffmann and Smith (2005:191).

Holotype: USNM 236408. Skin and skull. Adult male. Collected 

18 Jun 1921 by L. J. Goldman. Original number 2754.

Type Locality: Echo Crater, Snake River Desert, 20 mi (32 km) 

southwest of Arco, [Butte County], Idaho. 

Ochotona schisticeps jewetti A. H. Howell

Proc. Biol. Soc. Wash. 32: 109, 20 May 1919.

= Ochotona princeps schisticeps (Merriam, 1889). See Hafner 

and Smith (2010:411) for current use.

Holotype: USNM 208352. Skin and skull. Adult male. Collected 

3 Sep 1915 by S. G. Jewett. Original number 2362.

Type Locality: Head of Pine Creek, near Cornucopia, south slope 

of Wallowa Mountains, Baker County, Oregon. 

Ochotona uinta Hollister

Proc. Biol. Soc. Wash. 25: 58, 13 Apr 1912b.

= Ochotona princeps uinta Hollister, 1912. See A. H. Howell 

(1924:19). See Hafner and Smith (2010:412) for current use.

Holotype: USNM 9750. Skin only. Adult. Collected 28 Sep 1870 

by F. V. Hayden. Original number 567.

Type Locality: Uinta Mountains, Utah. Restricted by A. H. Howell (1924:19) to “near head of east fork of Bear River, Uinta 

Mountains, Utah.” 

Ochotona uinta lemhi A. H. Howell

Proc. Biol. Soc. Wash. 32: 106, 20 May 1919.

= Ochotona princeps princeps (Richardson, 1828). See Hafner 

and Smith (2010:410) for current use.

Holotype: USNM 23543/30961. Skin and skull. Adult female. 

Collected 19 Aug 1890 by C. P. Streator. Original number 

174.

Type Locality: Lemhi [FC: Salmon River] Mountains, 10 mi 

(16 km) west of Junction, [FC: altitude about 9,000 ft 

(2,743 m)], [Lemhi County], Idaho. 

Ochotona uinta nevadensis A. H. Howell

Proc. Biol. Soc. Wash. 32: 107, 20 May 1919.

= Ochotona princeps princeps (Richardson, 1828). See Hafner 

and Smith (2010:410) for current use.

Holotype: USNM 94213. Skin and skull. Adult female. Collected 

21 Jun 1898 by V. Bailey. Original number 6580.

Type Locality: Ruby Mountains, altitude 10,500 ft (3,200 m), 

southwest of Ruby P. O., [Elko County], Nevada. 

Ochotona uinta ventorum A. H. Howell

Proc. Biol. Soc. Wash. 32: 106, 20 May 1919.

= Ochotona princeps princeps (Richardson, 1828). See Hafner 

and Smith (2010:410) for current use.

Holotype: USNM 176778. Skin and skull. Adult male. Collected 

19 Jul 1911 by H. E. Anthony. Original number 395.

Type Locality: [SL and FC: 4 mi (6 km) south of] Fremont Peak, 

[SL and FC: altitude 11,500 ft (3,505 m)], Wind River 

Mountains, [Fremont County], Wyoming. 

Family Leporidae G. Fischer, 1817

Genus Brachylagus Miller, 1900

Lepus idahoensis Merriam

N. Am. Fauna 5: 76, 30 Jul 1891.

= Brachylagus idahoensis (Merriam, 1891). See Lyon (1904: 

411).

Holotype: USNM 24045/31461. Skin and skull. Adult male. 

Collected 16 Sep 1890 by [SL: V. Bailey] and B. H. Dutcher. 

Original number [FC: VB] 1816.

Type Locality: Pahsimeroi Valley, [FC: near head Pahsimeroi 

River, altitude 7,000 ft (2,134 m)], [Davis (1939:363): near 

Goldburg, Custer County], Idaho. 

Genus Lepus Linnaeus, 1758

Lepus alleni tiburonensis Townsend

Bull. Am. Mus. Nat. Hist. 31: 120, 14 Jun 1912.

Holotype: USNM 198403. [Skin and skull]. [Poole and Schantz 

(1942:208): Adult] male. Collected 13 Apr [SL: 1911 by 

H. E. Anthony. Original number 160].

Type Locality: Tiburon Island, [Gulf of California, Sonora, 

Mexico].
2 4 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Remarks: The holotype was published as American Museum of 

Natural History number 31990. It was cataloged USNM 

198403 on 26 Jun 1914. 

Lepus americanus dalli Merriam

Proc. Wash. Acad. Sci. 2: 29, 14 Mar 1900.

Holotype: USNM 8996/7579. Skin and skull. Adult male. Collected [SL: 27 Jan 1867] by W. H. Dall. Original number 

[SL: 584].

Type Locality: Nulato River, Alaska.

Remarks: The holotype was designated by the skull number 

USNM 7579; USNM 8996 is the skin that belongs to it. 

Lepus americanus macfarlani Merriam

Proc. Wash. Acad. Sci. 2: 30, 14 Mar 1900.

= Lepus americanus dalli Merriam, 1900. See Youngman 

(1975:57).

Holotype: USNM 7111/14467. Skin [cannot be found] and skull. 

Adult male. Collected [MC: Mar 1863] by R. MacFarlane. 

Original number [MC: 319].

Type Locality: Fort Anderson, north of Great Bear Lake, [Miller 

and Rehn (1901:178): near mouth of Anderson River], 

Arctic America (Mackenzie District, Northwest Territories, 

Canada).

Remarks: The holotype was published as USNM 14467; USNM 

7111 is its corresponding skin, which cannot be found. 

Lepus arcticus canus Preble

N. Am. Fauna 22: 59, 31 Oct 1902.

= Lepus arcticus arcticus Ross, 1819. See Hoffmann and Smith 

(2005:196).

Holotype: USNM 106860. Skin and skull. Young adult male. 

Collected 17 Aug 1900 by E. A. Preble. Original number 

3347.

Type Locality: Barren grounds near Hubbart Point, about 75 mi 

(121 km) north of Fort Churchill, Keewatin (Manitoba), 

Canada. 

Lepus arcticus hubbardi Handley

Proc. Biol. Soc. Wash. 65: 199, 5 Nov 1952.

= Lepus arcticus monstrabilis Nelson, 1934. See Hoffmann and 

Smith (2005:196).

Holotype: USNM 290933. Skin and skull. Adult male. Collected 

11 Jun 1949 by C. O. Handley, Jr. Original number 1283.

Type Locality: At 350 ft (107 m) elevation, near Cherie Bay, 5 mi 

(8 km) [SL: 6 mi (10 km)] northeast of Mould Bay Station, 

Prince Patrick Island, District of Franklin, Northwest Territories, Canada, lat. 72o20 ́N, long. 119o08 ́W. 

Lepus arcticus monstrabilis Nelson

Proc. Biol. Soc. Wash. 47: 85, 8 Mar 1934.

Holotype: USNM 126169. [Skin and skull]. Adult male. Collected 24 Apr 1901, not 21 as in original description, by J. 

S. Warmbath. Original number [XC: 54]. 3923-X.

Type Locality: [SL: head of north arm] Buchanan Bay, Ellesmere 

Island, [Nunavut], northern Canada. 

Lepus arcticus porsildi Nelson

Proc. Biol. Soc. Wash. 47: 83, 8 Mar 1934.

= Lepus arcticus groenlandicus Rhoads, 1896. See Hoffmann 

and Smith (2005:196).

Holotype: USNM 248723. [Skin and skull]. Adult, sex not given. 

Collected 5 Sep 1926 by O. Hastrup, not by M. P. Porsild as 

given in original description. Original number [SL and XC: 

2]. 24637-X.

Type Locality: Near Julianehaab (Qaqortoq), lat. 61o20 ́N, not 

60o20 ́as in original description, Greenland. 

Lepus asellus G. S. Miller

Proc. Acad. Nat. Sci. Phila. 51: 380, 29 Sep 1899b.

= Lepus californicus texianus Waterhouse, 1848. See Hoffmann 

and Smith (2005:196).

Holotype: USNM 20895/36009. Skin and skull. Adult female. 

Collected 22 Oct 1891 by P. L. Jouy. Original number 270.

Type Locality: San Luis Potosi, [San Luis Potosi], Mexico. 

Lepus aurigineus Hollister

Proc. Biol. Soc. Wash. 25: 181, 24 Dec 1912d.

= Lepus tolai aurigineus Hollister, 1912. See G. M. Allen 

(1927:9).

Holotype: USNM 13761/38173. Skin and skull. Adult female. 

Collected 27 Dec 1880 by unknown collector. No original 

number.

Type Locality: Chiu Kinag (Jiujiang), northern Kiangsi (Jiangxi), 

China. 

Lepus bairdii Hayden

Am. Nat. 3: 115, May 1869.

= Lepus americanus bairdii Hayden, 1869. See J. A. Allen 

(1875:431).

Lectotype: USNM 4262/3801 as designated by Nelson (1909: 

109). Skin and skull. [Nelson (1909:109): Adult] male. Collected 2 Jun 1860 by F. V. Hayden. Original number 62.

Type Locality: Near Fremont Peak, summit of Wind River 

Mountains, [Fremont County], Wyoming.

Remarks: No type was designated in the original description, 

which was based on several specimens from the Wind River 

Mountains collected by F. V. Hayden in the first part of Jun 

1860. In his revision, Nelson (1909:109) called the abovelisted specimen the type, therefore designating a lectotype 

(ICZN, 1999: Article 74.5). 

Lepus brachyurus angustidens Hollister

Proc. Biol. Soc. Wash. 25: 183, 24 Dec 1912d.

Holotype: USNM 13871/38514. Skin and skull. Adult male. 

Collected 14 Dec 1882 by P. L. Jouy. No original number.

Type Locality: Tate Yama Mountain, Hondo Island (Honshu), 

Japan. 
NUMBER 644 • 2 5

Lepus californicus magdalenae Nelson

Proc. Biol. Soc. Wash. 20: 81, 22 Jul 1907.

= Lepus californicus xanti Thomas, 1898. See Hoffmann and 

Smith (2005:196).

Holotype: USNM 146168. [Skin and skull]. Adult male. Collected 26 Nov 1905 by E. W. Nelson and E. A. Goldman. 

Original number [SL: 18638].

Type Locality: Magdalena Island (Isla Santa Magdalena), [FC: 

20–50 ft (6–15 m)], Lower California (Baja California Sur), 

Mexico.

Remarks: Hoffmann and Smith (2005:196) used magdalenae as 

the subspecific name in error, as it is predated by xanti. This 

correction was noted on a working draft of the manuscript 

by R. D. Fisher and was agreed to by R. S. Hoffmann, but 

the correction did not make it into the final publication. 

Lepus campestris sierrae Merriam

Proc. Biol. Soc. Wash. 17: 132, 14 Jul 1904a.

= Lepus townsendii townsendii Bachman, 1839. See Orr 

(1940:37).

Holotype: USNM 67863. [Skin and skull]. [Lyon and Osgood 

(1909:130): Adult] female. Collected 9 Sep 1894 by F. Stephens. Original number 1889.

Type Locality: Hope Valley, altitude 7,800 ft (2,377 m), Alpine 

County, California. 

Lepus capensis abbotti Hollister

Proc. Biol. Soc. Wash. 31: 35, 16 May 1918a.

= Lepus capensis aegyptius Desmarest, 1822. See Hoffmann and 

Smith (2005:197).

Holotype: USNM 19014/34779. Skin and skull. Adult female. 

Collected 20 Sep 1888 by W. L. Abbott. No original number.

Type Locality: Plains east of Mount Kilimanjaro, British East Africa (Kenya). 

Lepus europaeus cinnamomeus Shamel

J. Mammal. 21: 77, 14 Feb 1940a.

= Lepus tolai cinnamomeus Shamel, 1940. See Hoffmann and 

Smith (2005:204).

Holotype: USNM 255006. Skin, skull, and skeleton. Adult male. 

Collected 13 Dec 1929 by D. C. Graham. Original number 

[SL: 321].

Type Locality: Suifu (Yibin), altitude 1,000 to 2,000 ft (305 to 

610 m), Szechwan (Sichuan), China. 

Lepus festinus Nelson

Proc. Biol. Soc. Wash. 17: 108, 18 May 1904.

= Lepus californicus texianus Waterhouse, 1848. See Hoffmann 

and Smith (2005:196).

Holotype: USNM 53490. [Skin and skull]. Adult male. Collected 

31 Mar 1893 by E. W. Nelson and E. A. Goldman. Original 

number [FC: EWN] 4522.

Type Locality: Irolo, [FC: altitude 7,600 ft (2,317 m)], Hidalgo, 

Mexico. 

Lepus gaillardi Mearns

Proc. U.S. Natl. Mus. 18: 560, 24 Jun 1896a.

= Lepus callotis gaillardi Mearns, 1896. See Anderson and 

Gaunt (1962:5).

Holotype: USNM 20525/35714. Skin and skull. Adult male. 

Collected 17 Jun 1892 by E. A. Mearns and F. X. Holzner. 

Original number [SL: 1885].

Type Locality: West fork of the Playas Valley, near monument 

Number 63, [FC: Plains between White Water and the San 

Luis Mountains], Mexican boundary line (Hidalgo County, 

New Mexico). 

Lepus grahami A. B. Howell

Proc. Biol. Soc. Wash. 41: 143, 15 Oct 1928c.

= Lepus oistolus Hodgson, 1840. See Smith (2008:290) for current use.

Holotype: USNM 239875. Skin and skull. Adult female. Collected late Jul or early Aug [FC: 31 Jul] 1923 by D. C. Graham. Original number 8.

Type Locality: Ulongkong, altitude about 10,000 ft (3,048 m), 

about 10 mi (16 km) south of Tatsienlu (Kangding), Szechwan (Sichuan), China. 

Lepus kakumegae Heller

Smithson. Misc. Collect. 59(16): 19, 5 Jul 1912b.

= Lepus microtis microtis Heuglin, 1865. See Hoffmann and 

Smith (2005:201).

Holotype: USNM 181809. [Skin and skull]. Adult female. Collected 12 Feb 1912 by E. Heller. Original number 5644.

Type Locality: Lukosa River, Kakumega (Kakamega) Forest, 

[Western Province], British East Africa (Kenya). 

Lepus klamathensis Merriam

N. Am. Fauna 16: 100, 28 Oct 1899.

= Lepus americanus washingtonii Baird, 1855. See Hoffmann 

and Smith (2005:195).

Holotype: USNM 92248. [Skin and skull]. Adult female. Collected 25 Jan 1898 by B. L. Cunningham. Original number 

86. [SL: 1191-X].

Type Locality: [Nelson (1909:107): Head of Wood River, near] 

Fort Klamath, [Klamath County], Oregon.

Remarks: Placement of washingtonii as a synonym of virginianus

by Hoffmann and Smith (2005:195) was an error caught 

by R. D. Fisher while he was reviewing a working copy of 

their manuscript. Further, washingtonii has priority over 

cascadensis, used by Hoffmann and Smith (2005:195), and 

is the current valid subspecific name. These corrections were 

agreed to by R. S. Hoffmann, but they did not make it into 

the published version of Hoffmann and Smith (2005:195). 

Lepus labradorius Miller

Proc. Biol. Soc. Wash. 13: 39, 29 May 1899c.

= Lepus arcticus bangsii Rhoads, 1896. See Hoffmann and Smith 

(2005:196).
2 6 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Syntypes: USNM 14149/37138. Skin [and skull]. [Lyon and Osgood (1909:29): Adult]. Collected 28 Sep 1882 by L. M. 

Turner. Original number 1180.

USNM 23132. Skull only. [Lyon and Osgood (1909:29): 

Adult]. Collected, date unknown, by L. M. Turner. Original 

number 2326. [MC: cataloged 12 Jan 1889].

Type Locality: Fort Chimo (Kuujjuaq), Ungava (Nunavik), [Quebec, Canada].

Remarks: These two specimens were designated as cotypes in 

the original description and are now syntypes (ICZN, 1999: 

Article 73.2.1); USNM 14149 was published as a skin only, 

to which skull number USNM 37138 belongs, and USNM 

23132 was erroneously published as 32132. 

Lepus merriami Mearns

Proc. U.S. Natl. Mus. 18: 2, 25 Mar 1896b. Preprint of Proc. 

U.S. Natl. Mus. 18: 444, 23 May 1896c.

= Lepus californicus melanotis Mearns, 1890. See Hoffmann 

and Smith (2005:196).

Holotype: USNM 83797. [Skin and skull]. Adult female. Collected 6 Apr 1893 by E. A. Mearns. Original number 2317.

Type Locality: Fort Clark, Kinney County, Texas.

Remarks: Published under the original number, 2317. Cataloged 

as USNM 83797 on 16 Sep 1897. 

Lepus merriami altamirae Nelson

Proc. Biol. Soc. Wash. 17: 109, 18 May 1904.

= Lepus californicus melanotis Mearns, 1890. See Hoffmann 

and Smith (2005:196).

Holotype: USNM 93691. [Skin and skull]. Adult male. Collected 

16 May 1898 by E. W. Nelson and E. A. Goldman. Original 

number 12365.

Type Locality: Alta Mira, [FC: altitude about 75 ft (23 m)], Tamaulipas, Mexico. 

Lepus othus Merriam

Proc. Wash. Acad. Sci. 2: 28, 14 Mar 1900.

= Lepus othus othus Merriam, 1900.

Holotype: USNM 15883. Skull only. Adult, sex unknown. Collected [MC: Feb 1877] by L. M. Turner. Original number 

[MC: 1418].

Type Locality: St. Michaels (St. Michael), [Norton Sound], 

Alaska. 

Lepus parnassius G. S. Miller

Proc. Biol. Soc. Wash. 16: 145, 12 Nov 1903c.

= Lepus europaeus parnassius Miller, 1903. See Ellerman 

(1951:436).

Holotype: USNM 122093. Skin and skull. Adult male. Collected 

10 Sep 1895. Collector unknown, received from W. Schluter 

of Halle, on the Saale, Germany.

Type Locality: Angorianni, north side of Lyakura (Parnassus) 

Mts., [Sterea Ellada], Greece. 

Lepus poadromus Merriam

Proc. Wash. Acad. Sci. 2: 29, 14 Mar 1900.

= Lepus othus othus Merriam, 1900. See Hoffmann and Smith 

(2005:202).

Holotype: USNM 98068. [Skin and skull]. [Lyon and Osgood 

(1909:30): Adult]. Collected 8 Jul 1899 by C. Palache. Original number A. K. Fisher 2207.

Type Locality: Stepovak Bay, Alaska Peninsula, [Alaska]. 

Lepus quercerus Hollister

Proc. Biol. Soc. Wash. 25: 182, 24 Dec 1912d.

= Lepus tibetanus stoliczkanus Blandford, 1875. See Hoffmann 

and Smith (2005:203).

Holotype: USNM 175446. Skin and skull. Adult male. Collected 

28 Jul 1912 by N. Hollister. Original number 4405.

Type Locality: Chuisaya Steppe, Altai Mountains, [Altayskiy], 

Siberia (Russia). 

Lepus raineyi Heller

Smithson. Misc. Collect. 59(16): 18, 5 Jul 1912b.

= Lepus microtis microtis Heuglin, 1865. See Hoffmann and 

Smith (2005:201).

Holotype: USNM 181808. [Skin and skull]. Adult female. Collected 21 Jul 1911 by P. J. Rainey. Original number Heller 

3061.

Type Locality: Longaya Water, 30 mi (48 km) south of Mount 

Marsabit, [Eastern Province], British East Africa (Kenya). 

Lepus saliens Osgood

N. Am. Fauna 19: 39, 6 Oct 1900.

= Lepus americanus dalli Merriam, 1900. See Youngman 

(1975:57).

Holotype: USNM 98956. [Skin and skull]. Adult male. Collected 

26 Jun 1899 by W. H. Osgood. Original number 504.

Type Locality: Caribou Crossing (Carcross), between Lake Bennett and Lake Tagish, Northwest Territory (Yukon), Canada. 

Lepus swinhoei sowerbyae Hollister

Proc. Biol. Soc. Wash. 25: 182, 24 Dec 1912d.

= Lepus tolai filchneri Matschie, 1907. See G. M. Allen (1927:8).

Holotype: USNM 172628. Skin and skull. Adult male. Collected 

5 Feb 1910 by A. de C. Sowerby. Original number 365.

Type Locality: Wu-tsai (Yancheng), altitude 6,000 ft (1,829 m), 

20 mi (32 km) west of Ning-wu-fu (Fenghuang), northern 

Shansi (Shanxi), China. 

Lepus texianus griseus Mearns

Proc. U.S. Natl. Mus. 18: 562, 24 Jun 1896a.

= Lepus californicus texianus Waterhouse, 1848. See Nelson 

(1909:142).

Holotype: USNM 21068/36108. Skin and skull. Adult female. 

Collected 22 Jun 1893 by Dr. [E. A.] Mearns. Original number 2353.
NUMBER 644 • 2 7

Type Locality: Fort Hancock, El Paso (Hudspeth) County, Texas.

Remarks: El Paso is still a valid county in Texas, but the type 

locality is in Hudspeth County. 

Lepus texianus wallawalla Merriam

Proc. Biol. Soc. Wash. 17: 137, 14 Jul 1904b.

= Lepus californicus deserticola Mearns, 1896. See Hoffmann 

and Smith (2005:196).

Holotype: USNM 23923/31328. Skin and skull. Adult female. 

Collected 18 Sep 1890 by C. P. Streator. Original number 271.

Type Locality: Touchet, Plains of Columbia, [Walla Walla 

County], Washington.

Remarks: The skull number was incorrectly given as 21328 in 

the original description. 

Lepus tularensis Merriam

Proc. Biol. Soc. Wash. 17: 136, 14 Jul 1904b.

= Lepus californicus californicus Gray, 1837. See Hoffmann and 

Smith (2005:196).

Holotype: USNM 126334. [Skin and skull]. Adult female. Collected 25 Oct 1900 by L. J. Goldman. Original number 

[SL:496].

Type Locality: Alila, in bottom of San Joaquin Valley, Tulare 

County, California. 

Lepus varronis G. S. Miller

Proc. Biol. Soc. Wash. 14: 97, 27 Jun 1901b.

= Lepus timidus varronis G. S. Miller. See G. S. Miller (1912a: 

528).

Holotype: USNM 105832. Skin and skull. Adult male. Collected 

5 Dec 1900 by E. H. Zollikofer. Original number 196.

Type Locality: Grisons, Heinzenberg, Canton of Graubunden, 

Switzerland. 

Lepus washingtonii Baird

Proc. Acad. Nat. Sci. Phila. 7: 333, reported favorably for publication 24 Apr 1855; receipt acknowledged 15 Jun 1855 by 

the American Philosophical Society.

= Lepus americanus washingtonii Baird, 1855. See J. A. Allen 

(1875:431).

Lectotype: USNM 280/1223 as designated by Lyon and Osgood (1909:31). Skin and skull. Adult, sex unknown. Collected 1 Apr 1854 by G. Suckley. Original number [MC: 

J. J. Stevens 7].

Type Locality: Steilacoom, Pierce County, Washington.

Remarks: Lyon and Osgood (1909:31) stated that for reasons 

given, and reiterated by Poole and Schantz (1942:215), this 

specimen must be regarded as the type, effectively designating it as the lectotype (ICZN, 1999: Article 74.5). Placement 

of washingtonii as a synonym of virginianus by Hoffmann 

and Smith (2005:195) was an error caught by R. D. Fisher 

while reviewing a working copy of their manuscript. Further, 

washingtonii has priority over cascadensis and is the current 

valid subspecific name. These corrections were agreed to 

by R. S. Hoffmann but they did not make it into the published version of Hoffmann and Smith (2005:195). The skin, 

USNM 280, also bears a Museum of Comparative Zoology 

label with the number 5500. In a letter in the type catalog 

file dated 22 Dec 1961 from C. O. Handley, Jr. to Barbara 

Lawrence, it was indicated that the skin was at the MCZ in 

May 1876; it was not at the USNM when Lyon and Osgood 

(1909:31) published their catalog but was present when 

Poole and Schantz (1942:215) published theirs. The circumstances surrounding the movement of the skin are unknown. 

Genus Romerolagus Merriam, 1896

Romerolagus nelsoni Merriam

Proc. Biol. Soc. Wash. 10: 173, 29 Dec 1896.

= Romerolagus diazi Ferrari-Perez, 1893. See Miller (1911b:229).

Holotype: USNM 57949. [Skin and skull]. Adult male. Collected 

6 Jan 1894 by E. W. Nelson and E. A. Goldman. Original 

number 5639.

Type Locality: [Miller and Rehn (1901:177): West slope of] 

Mount Popocatepetl, altitude 3,350 m or 11,000 ft, [SL: 

Mexico], Mexico. 

Genus Sylvilagus Gray, 1867

Lepus arizonae goldmani Nelson

Proc. Biol. Soc. Wash. 17: 107, 18 May 1904.

= Sylvilagus audubonii goldmani (Nelson, 1904). See Nelson 

(1909:225).

Holotype: USNM 96812. [Skin and skull]. Adult female, not 

male as in original description. Collected 20 Mar 1899 by 

E. A. Goldman. Original number [FC: Nelson and Goldman] 13588.

Type Locality: Culiacan, [FC: altitude about 175 ft (53 m)], 

Sinaloa, Mexico. 

Lepus arizonae major Mearns

Proc. U.S. Natl. Mus. 18: 557, 24 Jun 1896a.

= Sylvilagus audubonii arizonae (J. A. Allen, 1877). See Nelson 

(1909:222).

Holotype: USNM 18192/25090. Skin and skull. Adult male. Collected 23 Oct 1889 by L. Stejneger. Original number 3053.

Type Locality: Calabasas, [Santa Cruz County], Arizona. 

Lepus arizonae minor Mearns

Proc. U.S. Natl. Mus. 18: 557, 24 Jun 1896a.

= Sylvilagus audubonii minor (Mearns, 1896). See Nelson 

(1909:226).

Holotype: USNM 20104/37064. Skin and skull. Adult male. 

Collected 6 Feb 1892 by [E. A.] Mearns and [F. X.] Holzner. 

Original number 1418.

Type Locality: El Paso, [El Paso County], Texas. 
2 8 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Lepus audubonii Baird

Mammals Rep. Explor. Surv. 8(1): 608, 1857.

= Sylvilagus audubonii audubonii (Baird, 1857). See Nelson 

(1909:214).

Lectotype: USNM 1163/2045, as designated by Lyon and Osgood (1909:288). Skin and skull. Age and sex not given. 

Collected on unknown date by J. S. Newberry. [MC: Cataloged 31 Dec 1855]. Received from R. S. Williamson.

Type Locality: San Francisco, [San Francisco County], California.

Remarks: Lyon and Osgood (1909:288) stated that for reasons 

given, it seems reasonable to take this specimen as the type, 

effectively designating it as the lectotype (ICZN, 1999: Article 74.5). Neither skin nor skull was present at the time 

of Lyon and Osgood (1909:288). By the time Poole and 

Schantz (1942:216) published their type catalog, the skin 

was in the USNM and bore a Museum of Comparative Zoology label with number 5460 on it. The circumstances of 

its travel to the Museum of Comparative Zoology and its 

return are unknown. The skull was found in the general collection 21 Mar 1974 and installed in the type collection at 

that time. 

Lepus bachmani ubericolor G. S. Miller

Proc. Acad. Nat. Sci. Phila. 51: 383, 5 Oct 1899b.

= Sylvilagus bachmani ubericolor (G. S. Miller, 1899). See Lyon 

(1904:337).

Holotype: USNM 19964/35371. Skin and skull. Adult male. 

Collected 25 Feb 1890 by A. W. Anthony. Original number 

[SL: 1226].

Type Locality: Beaverton, [Washington County], Oregon. 

Lepus baileyi Merriam

Proc. Biol. Soc. Wash. 11: 148, 9 Jun 1897b.

= Sylvilagus audubonii baileyi (Merriam, 1897). See Nelson 

(1909:232).

Holotype: USNM 56016. [Skin and skull]. Adult female. Collected 17 Sep 1893 by C. H. Merriam and V. Bailey. Original number [FC: VB] 4372.

Type Locality: Spring Creek, east side of Bighorn Basin, [Big 

Horn County], Wyoming. 

Lepus brasiliensis var. gabbi J. A. Allen

Monogr. N. Am. Rodentia, p. 349, Aug 1877.

= Sylvilagus gabbi (J. A. Allen, 1877). See Lyon (1904:336). See 

Ruedas and Salazar-Bravo (2007:63) for current use.

Lectotype: 11371/37794 as designated by Nelson (1909:259). 

Skin and skull. [Lyon and Osgood (1909:33): Adult] male. 

Collected by [SL: J. C. Zeledon], date unknown. Skin cataloged [MC: between 28 May and 11 Dec 1873; the skull 20 

Oct 1889]. Received from W. M. Gabb.

Type Locality: Talamanca, Costa Rica. Restricted by Hershkovitz (1950:352) to Talamanca, Sipurio, Rio Sixaola, near the 

Caribbean Coast, [Limon], Costa Rica.

Remarks: Allen (1877:337) listed three specimens of his new 

variety. In his revision of The Rabbits of North America, 

Nelson (1909:259) designated USNM 11371/37794 as the 

type, effectively designating it the lectotype (ICZN, 1999: 

Article 74.5). Although Lyon and Osgood (1909:33) listed 

USNM 11371 and USNM 11372 as cotypes = syntypes, 

Nelson’s (1909:259) act takes precedent (ICZN, 1999: Article 73.2.2). 

Lepus floridanus caniclunis G. S. Miller

Proc. Acad. Nat. Sci. Phila. 51: 388, 5 Oct 1899b.

= Sylvilagus floridanus chapmani (J. A. Allen, 1899). See Lyon 

and Osgood (1909:32).

Holotype: USNM 63137. Skin and skull. Adult male. Collected 

27 Dec 1892 by E. A. Mearns. Original number 2172.

Type Locality: Fort Clark, [Kinney County], Texas. 

Lepus floridanus chiapensis Nelson

Proc. Biol. Soc. Wash. 17: 106, 18 May 1904.

= Sylvilagus floridanus aztecus (J. A. Allen, 1890). See Hoffmann 

and Smith (2005:209).

Holotype: USNM 75953. [Skin and skull]. Adult female. Collected 28 Sep 1895 by E. W. Nelson and E. A. Goldman. 

Original number 8483.

Type Locality: San Cristobal, [FC: 8,200 ft (2,499 m)], Chiapas, 

[SL: Mexico]. 

Lepus floridanus connectens Nelson

Proc. Biol. Soc. Wash. 17: 105, 18 May 1904.

= Sylvilagus floridanus connectens (Nelson, 1904). See Lyon and 

Osgood (1909:32).

Holotype: USNM 63660. [Skin and skull]. Adult male. Collected 

15 Feb 1894 by by E. W. Nelson and E. A. Goldman. Original number 5849.

Type Locality: Chichicaxtle, Veracruz, [SL: Mexico]. 

Lepus floridanus sanctidiegi G. S. Miller

Proc. Acad. Nat. Sci. Phila. 51: 386, 5 Oct 1899b.

= Sylvilagus audubonii arizonae (J. A. Allen, 1877). See Hoffmann and Smith (2005:208).

Holotype: USNM 60668. Skin and skull. Adult female. Collected 10 Jul 1894 by E. A. Mearns. Original number [SL: 

3783].

Type Locality: Monument No. 258, Pacific Ocean, Mexican 

Boundary Line, San Diego County, California. 

Lepus floridanus subcinctus G. S. Miller

Proc. Acad. Nat. Sci. Phila. 51: 384, 5 Oct 1899b.

= Sylvilagus floridanus orizabae (Merriam, 1893). See Hoffmann 

and Smith (2005:209).

Holotype: USNM 20435/35665. Skin and skull. Adult female. 

Collected 15 Jun 1892 by P. L. Jouy. Original number 415.

Type Locality: Hacienda El Molino, near Negrete, Michoacan, 

Mexico. 
NUMBER 644 • 2 9

Lepus floridanus yucatanicus G. S. Miller

Proc. Acad. Nat. Sci. Phila. 51: 388, 5 Oct 1899b.

= Sylvilagus floridanus yucatanicus (G. S. Miller, 1899). See Lyon 

(1904:336).

Holotype: USNM 11441/37772. Skin and skull. Adult female. 

Collected 22 Feb 1865 by A. Schott. Original number 207.

Type Locality: Merida, Yucatan, [Mexico]. 

Lepus graysoni J. A. Allen

Monogr. N. Am. Rodentia, p. 347, 1877.

= Sylvilagus graysoni (J. A. Allen, 1877). See Lyon (1904:336).

Holotype: USNM 8318. Skin with skull inside. [Lyon and Osgood (1909:33): Adult] female. Collected by A. J. Grayson, 

date unknown. [MC: Cataloged 15 Nov 1856].

Type Locality: Tres Marias Islands, [Nelson (1899:16): Undoubtedly from Maria Madre], [Nayarit, Mexico].

Remarks. J. A. Allen (1877:347) did not specifically designate 

the type by number but stated that the description was 

based mainly on the specimen from the Tres Marias Islands 

and referred a second specimen to his new taxon. In a table 

(J. A. Allen 1877:348) lists USNM 8318 as the specimen 

from the Tres Marias Islands. 

Lepus insonus Nelson

Proc. Biol. Soc. Wash. 17: 103, 18 May 1904.

= Sylvilagus insonus (Nelson, 1904). See Lyon and Osgood 

(1909:34).

Holotype: USNM 126878. [Skin and skull]. Adult female. Collected 20 May 1903 by E. W. Nelson and E. A. Goldman. 

Original number 16466.

Type Locality: Omilteme (Omiltemi), Guerrero, [SL: Mexico]. 

Lepus margaritae G. S. Miller

Proc. Biol. Soc. Wash. 12: 97, 30 Apr 1898a.

= Sylvilagus floridanus margaritae (G. S. Miller, 1898). See 

Hershkovitz (1950:347).

Holotype: USNM 63217. [Skin and skull]. Adult male. Collected 

1 Jul 1895 by W. Robinson. Original number 369.

Type Locality: Margarita Island (Isla de Margarita), [Nueva Esparta], Venezuela. 

Lepus orizabae Merriam

Proc. Biol. Soc. Wash. 8: 143, 29 Dec 1893.

= Sylvilagus floridanus orizabae (Merriam, 1893). See Nelson 

(1909:183).

Holotype: USNM 53318. [Skin and skull]. Adult female. Collected 24 Apr 1893 by E. W. Nelson. Original number 4730.

Type Locality: Mount Orizaba, altitude about 9,500 ft (2,896 m), 

Puebla, Mexico. 

Lepus pinetis robustus Bailey

N. Am. Fauna 25: 159, 24 Oct 1905.

= Sylvilagus floridanus robustus (Bailey, 1905). See Hall and Kelson (1951:56). See Nalls et al. (2012:6) for current use.

Holotype: USNM 18262/25165. Skin and skull. Adult female. 

Collected 6 Jan 1890 by V. Bailey. Original number 873.

Type Locality: [FC: Finleys Ranch], altitude 6,000 ft (1,829 m), 

[SL: Fort Davis], Davis Mountains, [Jeff Davis County], Texas. 

Lepus simplicicanus G. S. Miller

Proc. Biol. Soc. Wash. 15: 81, 25 Apr 1902c.

= Sylvilagus floridanus chapmani (J. A. Allen, 1899). See Lyon 

and Osgood (1909:37).

Holotype: USNM 21805/36508. Skin and skull. Adult female. 

Collected 19 Oct 1891 by F. B. Armstrong. Original number 

[SL: 1402].

Type Locality: Brownsville, [Cameron County], Texas. 

Lepus sylvaticus var. arizonae J. A. Allen

Monogr. N. Am. Rodentia, p. 332, 1877.

= Sylvilagus audubonii arizonae (J. A. Allen, 1877). See Nelson 

(1909:222).

Holotype: USNM 8439. Skin of [Lyon and Osgood (1909:31): 

adult] male. Collected 8 Sep 1865 by E. Coues. Original 

number 1563.

Type Locality: Beales Springs, 50 mi (80 km) west of Fort Whipple, [Mohave County], Arizona.

Remarks: J. A. Allen (1877:332) did not explicitly designate a type 

by number, but stated that the taxon was based primarily on 

a specimen collected by Coues on 8 Sep 1865 at the locality 

given above. In the list of specimens, (J. A. Allen, 1877:340), 

number 8439 is the only specimen that matches those criteria. 

Lepus sylvaticus holzneri Mearns

Proc. U.S. Natl. Mus. 18: 554, 24 Jun 1896a.

= Sylvilagus floridanus holzneri (Mearns, 1896). See Lyon 

(1904:336).

Holotype: USNM 58937. [Skin and skull]. Adult female. Collected 29 Aug 1893 by F. X. Holzner. Original number 989.

Type Locality: [FC: Tanner’s Canyon at the saw mill], in the 

Douglas Spruce zone near the summit of the Huachuca 

Mountains, [Cochise County], southern Arizona. 

Lepus sylvaticus rigidus Mearns

Proc. U.S. Natl. Mus. 18: 555, 24 Jun 1896a.

= Sylvilagus floridanus holzneri (Mearns, 1896). See Nelson 

(1909:178).

Holotype: USNM 20336/35537. Skin and skull. Adult male. 

Collected 21 Apr 1892 by [E. A.] Mearns and [F. X.] Holzner. Original number 1680.

Type Locality: [MC: Near Carrizalillo Spring], near Monument 

No. 31, Carrizalillo Mountains, Mexican Boundary Line, 

[Luna County, New Mexico]. 

Lepus trowbridgii Baird

Proc. Acad. Nat. Sci. Phila. 7: 333, reported favorably for publication 24 Apr 1855; receipt acknowledged 15 Jun 1855 by 

the American Philosophical Society.
3 0 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

= Sylvilagus bachmani bachmani (Waterhouse, 1839). See Nelson (1909:247).

Lectotype: USNM 310/1235 as designated by Lyon and Osgood 

(1909:37). Skin and skull. Age and sex not given. Collected 

in 1853 by W. P. Trowbridge. No original number.

Type Locality: Monterey County, California.

Remarks: A type was not designated in the original description. 

Lyon and Osgood (1909:37) give their reasons for selecting 

this specimen as the type = lectotype (ICZN, 1999: Article 

74.5). Although Lyon and Osgood’s (1909:37) selection 

also establishes the type locality as given above, it should 

be noted the USNM skull catalog reads “Monterey? Cal. 

Perhaps San Diego” and the skin label reads “Presidio or 

Monterey, near San Francisco.” Further, the skin label also 

adds Museum of Comparative Zoology number 1458, perhaps explaining why Lyon and Osgood (1909:37) reported 

the skin as lost. The skin was at the USNM at the time Poole 

and Schantz (1942:223) published their type catalog. 

Lepus truei J. A. Allen

Bull. Am. Mus. Nat. Hist. 3: 192, 10 Dec 1890.

= Sylvilagus gabbi truei (J. A. Allen, 1890). See Ruedas and 

Salazar-Bravo (2007:68) for current use.

Holotype: USNM 6357/34878. Skin and skull. Adult, sex not 

given. Collected by C. Sartorius, date unknown. Original 

number e. [MC: Skin cataloged Apr 1863; skull 27 Nov 

1890].

Type Locality: Mirador, [Veracruz], Mexico.

Remarks: In a cataloging mixup, the skull was erroneously cataloged as USNM 25953 and was published as such. It is correctly presented here as USNM 34878. 

Lepus veraecrucis pacificus Nelson

Proc. Biol. Soc. Wash. 17: 104, 18 May 1904.

= Sylvilagus cunicularius cunicularius (Waterhouse, 1848). See 

Hoffmann and Smith (2005:209).

Holotype: USNM 70622. [Skin and skull]. Adult male. Collected 

9 Jan 1895 by E. W. Nelson and E. A. Goldman. Original 

number 7340.

Type Locality: Acapulco, [FC: near sea level], Guerrero, [SL: 

Mexico]. 

Sylvilagus aquaticus littoralis Nelson

N. Am. Fauna 29: 273, 31 Aug 1909.

Holotype: USNM 33848/45883. Skin and skull. Adult female. Collected 4 May 1892 by V. Bailey. Original number [SL: 3581].

Type Locality: Houma, [Terrebonne Parish], Louisiana. 

Sylvilagus audubonii cedrophilus Nelson

Proc. Biol. Soc. Wash. 20: 83, 22 Jul 1907.

= Sylvilagus audubonii warreni Nelson, 1907. See Hoffmeister 

and Lee (1963:514).

Holotype: USNM 148287. [Skin and skull]. Adult female. Collected 6 Nov 1906 by V. Bailey. Original number [SL: 8595].

Type Locality: Cactus Flat, [FC: altitude 5,200 ft (1,585 m)], 20 

mi (32 km) north of Cliff, [Grant County], New Mexico. 

Sylvilagus audubonii neomexicanus Nelson

Proc. Biol. Soc. Wash. 20: 83, 22 Jul 1907.

= Sylvilagus audubonii baileyi (Merriam, 1897). See Hoffmann 

and Smith (2005:208).

Holotype: USNM 118477. [Skin and skull]. Adult male. Collected 23 Sep 1902 by J. H. Gaut. Original number [SL: 

506].

Type Locality: [FC: Fifteen mi (24 km) south of] Fort Sumner, 

[De Baca County], New Mexico. 

Sylvilagus audubonii vallicola Nelson

Proc. Biol. Soc. Wash. 20: 82, 22 Jul 1907.

= Sylvilagus audubonii audubonii (Baird, 1857). See Hoffmann 

and Smith (2005:208).

Holotype: USNM 31257/43122. Skin and skull. Adult female. 

Collected 22 Oct 1891 by E. W. Nelson. Original number 

[SL: 1353].

Type Locality: San Emigdio Ranch, Kern County, California. 

Sylvilagus audubonii warreni Nelson

Proc. Biol. Soc. Wash. 20: 83, 22 Jul 1907.

Holotype: USNM 148632. [Skin and skull]. Adult female. Collected 4 Jan 1907 by C. H. Smith. Original number [XC: 

23]. [SL: 6312-X].

Type Locality: [FC: Sagebrush south of] Coventry, [Montrose 

County], Colorado. 

Sylvilagus bachmani exiguus Nelson

Proc. Biol. Soc. Wash. 20: 84, 22 Jul 1907.

Holotype: USNM 139607. [Skin and skull]. Adult male. Collected 19 Sep 1905 by E. W. Nelson and E. A. Goldman. 

Original number [SL: 18153].

Type Locality: Yubay, [SL: 30 mi (48 km) southeast of Calamahue], [FC: altitude 200 ft (61 m)], central Lower California 

(Baja California), Mexico. 

Sylvilagus brasiliensis sanctaemartae Hershkovitz

Proc. U.S. Natl. Mus. 100: 353, 26 May 1950.

Holotype: USNM 279993. Skin and skull. Adult female. Collected 12 Mar 1942 by P. Hershkovitz. Original number 

232.

Type Locality: Colonia Agricola de Caracolicito, altitude 335 

m, Rio Ariguani, southern slope of the Sierra Nevada Santa 

Marta, department of Magdalena (Cesar/Magdalena), 

Colombia. 

Sylvilagus cognatus Nelson

Proc. Biol. Soc. Wash. 20: 82, 22 Jul 1907.

Holotype: USNM 136569. [Skin and skull]. Adult, sex not given. 

Collected Feb 1905 by A. Rea. No original number [SL: 

5331-X].
NUMBER 644 • 3 1

Type Locality: Near summit of Manzano Mountains, altitude 

10,000 ft (3,048 m), New Mexico. Restricted by Frey et 

al. (1997:331) to: the vicinity of Rea Ranch, 1.9 km N and 

13.4 km W of Tajique, T6N, R5E, NE 1⁄4 of NW 1⁄4 Sec 

9, N34o45′05.39 ̋W106o25′18.04 ̋ on the northeast side of 

Bosque Peak at 2,880 m elevation, Manzano Mountains, 

Torrence County, New Mexico. 

Sylvilagus floridanus ammophilus A. H. Howell

J. Mammal. 20: 365, 14 Aug 1939.

= Sylvilagus floridanus floridanus (J. A. Allen, 1890). See Hoffmann and Smith (2005:209).

Holotype: USNM 70313. Skin and skull. Adult male. Collected 

22 Jan 1895 by J. A. Loring. Original number 2672.

Type Locality: Oak Lodge, on peninsula opposite Micco, [FC: 

Brevard County], Florida. 

Sylvilagus floridanus hitchensi Mearns

Proc. U.S. Natl. Mus. 39: 227, 9 Jan 1911.

= Sylvilagus floridanus mallurus (Thomas, 1898). See Hoffmann 

and Smith (2005:209).

Holotype: USNM 155577. [Skin and skull]. Adult female. Collected 

13 May 1910 by E. A. Mearns. Original number [SL: 7419].

Type Locality: Smiths Island (Smith Island), Northampton 

County, Virginia. 

Sylvilagus floridanus hondurensis Goldman

Proc. Biol. Soc. Wash. 45: 122, 30 Jul 1932.

Holotype: USNM 257062. [Skin and skull]. Adult male. Collected 

3 Dec 1931 by C. F. Underwood. Original number 628.

Type Locality: Monte Redondo, altitude about 5,100 ft (1,555 m), 

about 30 mi (48 km) northwest of Tegucigalpa, Honduras. 

Sylvilagus floridanus macrocorpus Diersing and Wilson

Smithson. Contrib. Zool. 297: 8, 8 Jan 1980.

Holotype: USNM 509011. Skin and skull. Adult female. Collected 20 Mar 1975 by C. Jones. Original number 5251.

Type Locality: Estanzuela, 1,372 m, Nayarit, Mexico. 

Sylvilagus floridanus restrictus Nelson

Proc. Biol. Soc. Wash. 20: 82, 22 Jul 1907.

= Sylvilagus floridanus orizabae (Merriam, 1893). See Hoffmann 

and Smith (2005:209).

Holotype: USNM 33687/45722. Skin and skull. Adult male. Collected 25 Apr 1892 by E. W. Nelson. Original number 2576.

Type Locality: Zapotlan (Ciudad Guzman), [FC: at north base of 

Sierra Nevada, about 5,000 ft (1,524 m)], Jalisco, Mexico.

Remarks: Published under the skin number 33687, to which 

skull number 45722 belongs. 

Sylvilagus floridanus similis Nelson

Proc. Biol. Soc. Wash. 20: 82, 22 Jul 1907.

= Sylvilagus floridanus mearnsii (J. A. Allen, 1894). See Hoffmann and Smith (2005:209).

Holotype: USNM 69517. [Skin and skull]. Adult male. Collected 

10 Nov 1894 by C. P. Streator. Original number 4442.

Type Locality: Valentine, [Cherry County], Nebraska.

Remarks: Hoffmann and Smith (2005:209) incorrectly listed alacer as the subspecific name for this taxon. This error was 

caught by R. D. Fisher while reviewing the Hoffmann and 

Smith manuscript. R. S. Hoffmann agreed that that mearnsii 

should be the valid subspecies, but the correction did not 

make it into the final publication of Hoffmann and Smith 

(2005:209). 

Sylvilagus gabbi messorius Goldman

Smithson. Misc. Collect. 60(2): 13, 20 Sep 1912b.

= Sylvilagus gabbi gabbi (J. A. Allen, 1877). See Ruedas and 

Salazar-Bravo (2007:68) for current use.

Holotype: USNM 179569. [Skin and skull]. Adult male. Collected 23 May 1912 by E. A. Goldman. Original number 

21736.

Type Locality: Cana, altitude 1,800 ft (549 m), [Darien], eastern 

mountains of Panama. 

Sylvilagus graysoni badistes Diersing and Wilson

Smithson. Contrib. Zool. 297: 15, 8 Jan 1980.

= Sylvilagus graysoni (J. A. Allen, 1877). See Hoffmann and 

Smith (2005:210).

Holotype: USNM 512550. Skin and skull. Adult male. Collected 24 Mar 1976 by D. E. Wilson. Original number 

3730.

Type Locality: San Juanito Island, Tres Marias Islands, Nayarit, 

Mexico. 

Sylvilagus mansuetus Nelson

Proc. Biol. Soc. Wash. 20: 83, 22 Jul 1907.

Holotype: USNM 79041. [Skin and skull]. Adult [SL: female], 

not male as in original description. Collected 2 Aug 1895 by 

J. E. McLellan. Original number [SL: 1444].

Type Locality: San Jose Island, Gulf of California, [Baja California Sur], Mexico. 

Sylvilagus obscurus Chapman, Cramer, Dippenaar, and Robinson

Proc. Biol. Soc. Wash. 105: 858, 18 Dec 1992.

Holotype: USNM 567187. Skin, skull, [and one femur]. Adult 

female. Collected 8 Apr 1974 by J. A. Chapman. Original 

number DS-58.

Type Locality: Dolly Sods Scenic Area, Grant County, West 

Virginia. 

Sylvilagus palustris hefneri Lazell

J. Mammal. 65: 26, 28 Feb 1984.

Holotype: USNM 555687. Skin, skull, and skeleton. Adult female. Collected 4 Feb 1980 by J. Lazell. Original number 

[SL: Z-08259].

Type Locality: S. Lower Sugarloaf Key, Monroe County, 

Florida. 
3 2 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Order Erinaceomorpha Gregory, 1910

Family Erinaceidae G. Fischer, 1814

Genus Atelerix Pomel, 1848

Erinaceus sotikae Heller

Smithson. Misc. Collect. 56(15): 1, 23 Dec 1910a.

= Atelerix albiventris (Wagner, 1841). See Corbet (1988:149).

Holotype: USNM 162112. [Skin and skull]. Adult male. Collected 28 Jun 1909 by J. A. Loring. Original number 6451.

Type Locality: Guaso Nyiro, Sotik District, [Rift Valley], British 

East Africa (Kenya). 

Genus Echinosorex Blainville, 1838

Gymnura gymnura minor Lyon

Proc. U.S. Natl. Mus. 36: 453, 27 May 1909b.

= Echinosorex gymnura gymnura (Raffles, 1822). See Corbet 

(1988:128).

Holotype: USNM 86783. Skin and skull. Adult male. Collected 

12 Jan 1899 by W. L. Abbott. No original number.

Type Locality: Khow Nok Ram (Khao Nok Ra), altitude 2,000 ft 

(610 m), Trong (Trang), Lower Siam (Thailand). 

Genus Podogymnura Mearns, 1905

Podogymnura truei Mearns

Proc. U.S. Natl. Mus. 28: 437, 13 May 1905.

Holotype: USNM 125286. In alcohol, skull removed. Adult 

female. Collected 25 Jun 1904 by E. A. Mearns. Original 

number 5667.

Type Locality: Mount Apo, at 6,000 ft (1,829 m), [Davao del 

Sur], southern Mindanao, Philippine Islands (Philippines). 

Order Soricomorpha Gregory, 1910

Family Nesophontidae Anthony, 1916

Genus Nesophontes Anthony, 1916

Nesophontes hypomicrus G. S. Miller

Smithson. Misc. Collect. 81(9): 4, 20 Mar 1929.

Holotype: USNM 253077.

This specimen was transferred to the NMNH Paleobiology Department, date unknown. 

Nesophontes paramicrus G. S. Miller

Smithson. Misc. Collect. 81(9): 3, 20 Mar 1929.

Holotype: USNM 253063.

This specimen was transferred to the NMNH Paleobiology Department, date unknown. 

Nesophontes zamicrus G. S. Miller

Smithson. Misc. Collect. 81(9): 3, 20 Mar 1929.

Holotype: USNM 252090.

This specimen was transferred to the NMNH Paleobiology Department, date unknown. 

Family Soricidae G. Fischer, 1814

Genus Blarina Gray, 1838

Blarina angusticeps Baird

Mammals Rep. Explor. Surv. 8(1): 47, 1857.

= Blarina brevicauda talpoides (Gapper, 1830). See Hall and Kelson (1959:55). See Webster et al. (2011:37) for current use.

Holotype: USNM 1318/2120, not 2520 as in original desctiption. Skin and skull made from alcohol. Age and sex not 

given. Collected on unknown date by Z. Thompson. Cataloged [MC: Feb 1856]. Original number [SL: 203].

Type Locality: Burlington, [Chittenden County], Vermont. 

Blarina brevicauda knoxjonesi Webster

Contrib. Mammal. p. 52, 5 Sep 1996.

Not listed by Hutterer (2005:269). See Webster et al. (2011:36) 

for current use.

Holotype: USNM 567349. Skin and skull. Adult male. Collected 

16 Nov 1982 by W. D. Webster. Original number 1294. Formerly University of North Carolina, Wilmington 486.

Type Locality: [SL: Dow Road], Carolina Beach, New Hanover 

County, North Carolina, 34o01 ́32 ̋N, 77o54 ́36 ̋W. 

Blarina brevicauda shermani Hamilton

Proc. Biol. Soc. Wash. 68: 37, 20 May 1955.

= Blarina shermani Hamilton, 1955. See Benedict et al. (2006:1) 

for current use.

Holotype: USNM 514828. Skin and skull. Adult female. Collected 13 Feb 1954 by W. J. Hamilton, Jr. Original number 

4280. Formerly Cornell University number 8026.

Type Locality: Two mi (3.2 km) North Fort Myers, Lee County, 

Florida. 

Blarina carolinensis peninsulae Merriam

N. Am. Fauna 10: 14, 31 Dec 1895a.

See Benedict et al. (2006:1) for current use.

Holotype: USNM 70874. [Skin and skull]. Adult male. Collected 

2 Mar 1895 by J. A. Loring. Original number 2777.

Type Locality: [FC: Fork of] Miami River, [FC: 5 mi (8 km) West 

of Miami], Dade (Miami-Dade) County, Florida. 

Blarina telmalestes Merriam

N. Am. Fauna 10: 15, 31 Dec 1895a.

= Blarina brevicauda talpoides (Gapper, 1830). See Webster et al. 

(2011:37) for current use.

Holotype: USNM 71823. [Skin and skull]. Adult female. Collected 6 Jun 1895 by A. K. Fisher. Original number 1775.

Type Locality: Lake Drummond, Dismal Swamp, [Chesapeake/

Suffolk City], Virginia. 
NUMBER 644 • 3 3

Sorex carolinensis Bachman

J. Acad. Nat. Sci. Phila. 7: 336, 1837.

= Blarina carolinensis carolinensis (Bachman, 1837). See Baird 

(1857:45).

Neotype: USNM 574157 as designated by Handley and Varn 

(1994:396). Skin and skull. Adult male. Collected 27 Jul 

1989 by C. O. Handley, Jr. and M. Varn. Original number 

COH 15236.

Type Locality: Beside Awendaw Creek, 3.2 km E Awendaw 

Post Office, Charleston County, South Carolina. 

Genus Crocidura Wagler, 1832

Crocidura agadiri Vesmanis and Vesmanis

Zool. Abhand. Staat. Mus. Tierk. 36: 67, 30 Apr 1980.

= Crocidura tarfayaensis Vesmanis and Vesmanis, 1980. See 

Hutterer (1987:530).

Holotype: USNM 470588. S, B (skin and skull). Female, age not 

given. Leg. (Collected) 20 Dec 1969 by M. G. Hearst. Original number 567.

Type Locality: 15 km No. (NE) Tiznit, Agadir Provinz (Province), Marakko (Morocco), 29o49′N, 9o39′W. 

Crocidura alchemillae Heller

African Game Trails p. 480, 1910b.

= Crocidura fumosa Thomas, 1904. See Dippenaar and Meester 

(1989:31).

Holotype: USNM 163087. [Skin and skull]. Adult male. Collected 17 Oct 1909 by E. Heller. Original number 1177.

Type Locality: Summit of Aberdare Range, altitude 10,500 ft 

(3,200 m), [Central Province], British East Africa (Kenya). 

Crocidura aleksandrisi Vesmanis

Bonn. Zool. Beitr. 28: 3, May 1977.

Holotype: USNM 325010. [Skin and skull]. Female, age not 

given. Collected 12 Jun 1962 by G. L. Ranck and J. H. 

Shaw. Original number JHS 213.

Type Locality: 5 km W Tocra, 32o31 ́N, 20o34 ́E, Cyrenaica, [SL: 

Libya]. 

Crocidura alpina Heller

Smithson. Misc. Collect. 56(9): 5, 22 Jul 1910c.

= Crocidura allex Osgood, 1910. See Hollister (1918b:68).

Holotype: USNM 163089. [Skin and skull]. Adult female. Collected 4 Oct 1909 by J. A. Loring. Original number 7728.

Type Locality: West slope of Mount Kenia (Kenya), altitude 

10,000 ft (3,048 m), British East Africa (Kenya). 

Crocidura andamanensis G. S. Miller

Proc. U.S. Natl. Mus. 24: 777, 29 May 1902a.

Holotype: USNM 111825. Skin and skull. Adult male. Collected 

16 Jan 1901 by W. L. Abbott. Original number 851.

Type Locality: MacPherson Strait, South Andaman Island, [Andaman and Nicobar Islands, India]. 

Crocidura beatus G. S. Miller

Proc. U.S. Natl. Mus. 38: 392, 19 Aug 1910.

Holotype: USNM 144647. Skin, skull, [and skeleton]. Adult 

male. Collected 28 May 1906 by E. A. Mearns. Original 

number 6173.

Type Locality: Summit of Mount Bliss, altitude 5,750 ft (1,753 m), 

Mindanao, [SL: P. I. (Philippines)]. 

Crocidura bolivari tamrinensis Vesmanis and Vesmanis

Zool. Abhand. Staat. Mus. Tierk. 36: 41, 30 Apr 1980.

= Crocidura viaria (I. Geoffroy, 1834). See Hutterer (1984:209).

Holotype: USNM 476091. S, B (skin and skull). Male, age not 

given. Leg. (Collected) 11 Apr 1970 by L. W. Robbins. Original number 3007.

Type Locality: 10 km N Tamri, 30o21′N, 9o49′W (Schmidt et 

al. 2008:34: 30o46 ́N, 9o50 ́W), Agadir Provinz (Province), 

Marakko (Morocco). 

Crocidura caudata G. S. Miller

Proc. Biol. Soc. Wash. 14: 42, 25 Apr 1901c.

= Crocidura sicula sicula G. S. Miller, 1901. See Vogel et al. 

(1989:246).

Holotype: USNM 103302. In alcohol, [skull removed]. Young 

adult female. Collected 21 Jun 1900 by D. Coolidge. Original number 1365.

Type Locality: Palermo, Sicily (Sicilia), [Italy]. 

Crocidura chaouianensis Vesmanis and Vesmanis

Zool. Abhand. Staat. Mus. Tierk. 36: 64, 30 Apr 1980.

= Crocidura russula yebalensis Cabrera, 1913. See Hutterer 

(1987:529).

Holotype: USNM 476081. S, B (skin and skull). Female, age 

not given. Leg. (Collected) 23 Apr 1970 by L. W. Robbins. 

Original number 3152.

Type Locality: 3 km N Settat, 33o02′N, 7o46′W (Schmidt et al. 

2008:34; 33o02′N, 07o37′W), Settat Provinz (Province), 

Marokko (Morocco). 

Crocidura daphnia Hollister

Smithson. Misc. Collect. 66(8): 1, 23 May 1916a.

= Crocidura olivieri nyansae Neumann, 1900. See Hutterer 

(2005:244).

Holotype: USNM 164898. Skin and skull. Adult female. Collected 19 Feb 1910 by J. A. Loring. Original number 9009.

Type Locality: Gondokoro, [Central Equatoria], Uganda (South 

Sudan). 

Crocidura elongata Miller and Hollister

Proc. Biol. Soc. Wash. 34: 101, 30 Jun 1921.

Holotype: USNM 217534. Skin and skull. Adult male. Collected 

1 Aug 1916 by H. C. Raven. Original number 2790.

Type Locality: Temboan, southwest from Tondano Lake, 

[Sulawesi Utara], northeastern Celebes (Sulawesi), 

[Indonesia]. 
3 4 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Crocidura essaouiranensis Vesmanis and Vesmanis

Zool. Abhand. Staat. Mus. Tierk. 36: 61, 30 Apr 1980.

= Crocidura whitakeri de Winton, 1897. See Hutterer (1987:530).

Holotype: USNM 485839. S, B (skin and skull). Male, age not 

given. Leg. (Collected) 2 Mar 1972 by H. L. Norman. Original number 555.

Type Locality: 5 km no. (NE) Essaouira, 31o31′N, 9o46′W, Safi 

Provinz (Province), Marokko (Morocco). 

Crocidura gouliminensis Vesmanis and Vesmanis

Zool. Abhand. Staat. Mus. Tierk. 36: 52, 30 Apr 1980.

= Crocidura tarfayaensis Vesmanis and Vesmanis, 1980. See 

Hutterer (1987:530).

Holotype: USNM 482864. S, B (skin and skull), Female, age not 

given. Leg. (Collected) 24 Jun 1971 by R. E. Vaden. Original number 6706.

Type Locality: 28 km SW Goulimine, 28o46′N, 10o14′W, Agadir 

Provinz (Province), Marokko (Morocco). 

Crocidura grandis G. S. Miller

Proc. U.S. Natl. Mus. 38: 393, 19 Aug 1910.

Holotype: USNM 144648. Skin and skull. Adult male. Collected 

3 Jun 1906 by E. A. Mearns. Original number 6181.

Type Locality: [FC: McMurray Flats], Grand Malindang Mountain, altitude 6,100 ft (1,859 m), [Misamis Occidental Province], Mindanao, [SL: P. I. (Philippines)]. 

Crocidura grisea A. B. Howell

Proc. Biol. Soc. Wash. 39: 137, 27 Dec 1926.

= Crocidura attenuata Milne-Edwards, 1872. See G. M. Allen 

(1938:122).

Holotype: USNM 238229. Skin and skull. Adult female. Collected 

23 Nov 1921 by A. de C. Sowerby. Original number 1076.

Type Locality: 75 mi (121 km) southwest of Yeng-ping-fu (Nanping), Fukien (Fujian), China. 

Crocidura grisescens A. B. Howell

J. Mammal. 9: 60, 9 Feb 1928d.

= Crocidura attenuata Milne-Edwards, 1872. See Jiang and 

Hoffmann (2001:1068).

Holotype: USNM 252187. Skin and skull. Adult female. Collected on unknown date by F. T. Smith. [MC: Cataloged 1 

Jul 1927]. Original number 89.

Type Locality: Kuatun (Guadun?), Fukien (Fujian), China. 

Crocidura halconus G. S. Miller

Proc. U.S. Natl. Mus. 38: 391, 19 Aug 1910.

= Crocidura grayi Dobson, 1890. See Heaney et al. (1987:36).

Holotype: USNM 144652. Skin and skull. Young adult female. 

Collected 18 Nov 1906 by E. A. Mearns. Original number 

6316.

Type Locality: [FC: Posey Flats], Spur of main ridge of Mount 

Halcon, altitude 6,300 ft (1,920 m), [Oriental Mindoro], 

Mindoro, [SL: P. I. (Philippines)]. 

Crocidura heljanensis safii Vesmanis and Vesmanis

Zool. Abhand. Staat. Mus. Tierk. 36: 49, 30 Apr 1980.

= Crocidura russula yebalensis Cabrera, 1913. See Hutterer 

(1987:529).

Holotype: USNM 476078. S, B (skin and skull). Male, age not 

given. Leg. (Collected) 16 Apr 1970 by M. G. Hearst. Original number 1442.

Type Locality: 13 km S Safi, 32o11′N, 9o15′W, Safi Provinz 

(Province), Marokko (Morocco). 

Crocidura hildegardeae altae Heller

Smithson. Misc. Collect. 60(12): 9, 4 Nov 1912a.

= Crocidura hildegardeae Thomas, 1904. Subspecies not recognized by Hutterer (2005:234).

Holotype: USNM 181819. Skin and skull. Adult male. Collected 

31 Aug 1911 by E. Heller. Original number 4190.

Type Locality: [SL: North Creek], altitude 6,000 ft (1,829 m), 

Mount Garguez (Warges), Mathews Range, [Rift Valley], 

British East Africa (Kenya). 

Crocidura hildegardeae procera Heller

Smithson. Misc. Collect. 60(12): 10, 4 Nov 1912a

= Crocidura hildegardeae Thomas, 1904. Subspecies not recognized by Hutterer (2005:234).

Holotype: USNM 181820. [Skin and skull]. Adult female. Collected 9 Sep 1911 by E. Heller. Original number 4274.

Type Locality: Summit of Mount Lololokui altitude 6,000 ft 

(1,829 m), northern Guaso Nyiro district, British East Africa (Kenya). 

Crocidura horsfieldi kurodai Jameson and Jones

Proc. Biol. Soc. Wash. 90: 461, 17 Oct 1977.

= Crocidura rapax kurodai Jameson and Jones, 1977. See Hutterer (2005:247).

Holotype: USNM 358115. Skin and skull. Adult female. Collected Jan 1969 by G. S. Jones. Original number T1451.

Type Locality: Linkou, [SL: 25o05′N, 121o23′E], Taipei Hsien, 

Taiwan. 

Crocidura lea Miller and Hollister

Proc. Biol. Soc. Wash. 34: 102, 30 Jun 1921.

Holotype: USNM 217553. Skin and skull. Adult male. Collected 

3 Aug 1916 by H. C. Raven. Original number 2837.

Type Locality: Temboan, [Sulawesi Utara], northeastern Celebes 

(Sulawesi), [Indonesia]. 

Crocidura lepidura Lyon

Proc. U.S. Natl. Mus. 34: 662, 14 Sep 1908.

Holotype: USNM 123140. Skin and skull. Adult female. Collected 1 Sep 1903 by W. L. Abbott. Original number 2772.

Type Locality: Kateman River, [Riau], eastern Sumatra, [Indonesia]. 

Crocidura levicula Miller and Hollister

Proc. Biol. Soc. Wash. 34: 103, 30 Jun 1921.
NUMBER 644 • 3 5

Holotype: USNM 219450. Skin and skull. Adult female. Collected 13 Feb 1918 by H. C. Raven. Original number 3521.

Type Locality: Pinedapa, [Sulawesi Tengah], Middle Celebes (Sulawesi), [Indonesia]. 

Crocidura lignicolor G. S. Miller

Proc. Wash. Acad. Sci. 2: 39, 30 Mar 1900b.

= Crocidura gmelini (Pallas, 1811). See Hoffmann (1996:161).

Holotype: USNM 62180. Skin [and skull]. Adult female. Collected 10 Jan 1894 by [MC: W. L. Abbott]. No original 

number.

Type Locality: Jungles east of Maralbashi near the Yarkand 

River, eastern Turkestan (Xinjiang, China). 

Crocidura littoralis Heller

Smithson. Misc. Collect. 56(15): 5, 23 Dec 1910a.

Holotype: USNM 164642. [Skin and skull]. Adult male. Collected 6 Jan 1910 by J. A. Loring. Original number 8598.

Type Locality: Butiaba, east shore of Albert Nyanza, [Bulisa], 

Uganda. 

Crocidura longipes Hutterer and Happold

Bonn. Zool. Monogr. 18: 53, 1983.

Holotype: USNM 402309. Skin and skull. Adult male. Collected 

2 Jun 1967, not Jul as in original description, by J. C. Geest. 

Original number [SL: 3534].

Type Locality: Dada, 11o34′N, 4o29′E, [SL and FC: Northern 

Region (Kebbi)], western Nigeria. 

Crocidura lutrella Heller

Smithson. Misc. Collect. 56(15): 4, 23 Dec 1910a.

= Crocidura parvipes Osgood, 1910. See Hutterer (1986:31).

Holotype: USNM 164640. [Skin and skull]. Adult male. Collected 14 Jan 1910 by J. A. Loring. Original number 8729.

Type Locality: Rhino Camp, Lado Enclave, [Arua, Uganda]. 

Crocidura lutreola Heller

Smithson. Misc. Collect. 60(12): 8, 4 Nov 1912a.

= Crocidura hildegardeae Thomas, 1904. See Hollister 

(1918b:64).

Holotype: USNM 181818. [Skin and skull]. Adult female. Collected 6 Nov 1911 by E. Heller. Original number 4635.

Type Locality: [SL: Summit] Mount Mbololo, Taita Hills, altitude 5,000 ft (1,524 m), [Coast Province], British East Africa (Kenya). 

Crocidura maanjae Heller

Smithson. Misc. Collect. 56(15): 4, 23 Dec 1910a.

= Crocidura hildegardeae Thomas, 1904. See Cabrera 

(1925:152).

Holotype: USNM 164639. [Skin and skull]. Adult male. Collected 27 Dec 1909 by J. A. Loring. Original number 8502.

Type Locality: Kabula Muliro (Kabulamuliro), [Mityana], [SL: 

Uganda]. 

Crocidura mesatanensis Vesmanis and Vesmanis

Zool. Abhand. Staat. Mus. Tierk. 36: 69, 30 Apr 1980.

= Crocidura whitakeri de Winton, 1897. See Hutterer (1987:530).

Holotype: USNM 476088. S, B (skin and skull). Female, age not 

given. Leg. (Collected) 30 Apr 1970 by M. G. Hearst. Original number 1582.

Type Locality: 10 km S Oued Zem, 32o46′N, 6o35′W, Khouribga 

Provinz (Province), Marakko (Morocco). 

Crocidura mimula G. S. Miller

Proc. Biol. Soc. Wash. 14: 95, 27 Jun 1901d.

See Bannikova et al. (2006:118) for current use.

Holotype: USNM 105801. Skin and skull. Adult female. Collected 1 Dec 1900 by E. H. Zollikofer. Original number 192.

Type Locality: Zuberwangen, St. (Sankt) Gallen, Switzerland. 

Crocidura mindorus G. S. Miller

Proc. U.S. Natl. Mus. 38: 392, 19 Aug 1910.

Holotype: USNM 144654. Skin, skull, [and body in alcohol]. 

Young adult male. Collected 19 Nov 1906 by E. A. Mearns. 

Original number 6321.

Type Locality: [FC: Posey Flats], summit main ridge of Mount 

Halcon, altitude 6,300 ft (1,920 m), [Oriental Mindoro], 

Mindoro, [Philippines]. 

Crocidura mutesae Heller

Smithson. Misc. Collect. 56(15): 3, 23 Dec 1910a.

Holotype: USNM 164636, not 174636 as in original description. [Skin and skull]. Adult female. Collected 22 Dec 1909 

by J. A. Loring. Original number 8430.

Type Locality: Kampala, [Kampala], Uganda. 

Crocidura nicobarica G. S. Miller

Proc. U.S. Natl. Mus. 24: 776, 29 May 1902a.

Holotype: USNM 111788. In alcohol, [skull removed]. Adult 

female. Collected 15 Mar 1901 by W. L. Abbott. Original 

number 931.

Type Locality: Great Nicobar Island, [Andaman and Nicobar 

Islands, India]. 

Crocidura nigripes Miller and Hollister

Proc. Biol. Soc. Wash. 34: 101, 30 Jun 1921.

= Crocidura nigripes nigripes Miller and Hollister, 1921.

Holotype: USNM 217545. Skin and skull. Adult male. Collected 

4 Aug 1916 by H. C. Raven. Original number 2866.

Type Locality: Temboan, southwest from Tondano Lake, [Sulawesi Utara], northeastern Celebes (Sulawesi), [Indonesia]. 

Crocidura nigripes lipara Miller and Hollister

Proc. Biol. Soc. Wash. 34: 101, 30 Jun 1921.

Holotype: USNM 219444. Skin and skull. Adult male. Collected 

2 Sep 1917 by H. C. Raven. Original number 3207.

Type Locality: Gimpoe (Gimpu), [Sulawesi Tengah], Middle Celebes (Sulawesi), [Indonesia]. 
3 6 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Crocidura nilotica Heller

Smithson. Misc. Collect. 56(15): 3, 23 Dec 1910a.

= Crocidura nigrofusca Matschie, 1895. See Hutterer (1993:91).

Holotype: USNM 164638. [Skin and skull]. Adult female. Collected 17 Jan 1910 by J. A. Loring. Original number 8768.

Type Locality: Rhino Camp, Lado Enclave, [Arua, Uganda]. 

Crocidura parvipes nisa Hollister

Smithson. Misc. Collect. 66(8): 2, 23 May 1916a.

= Crocidura parvipes Osgood, 1910. See Hutterer (2005:245).

Holotype: USNM 182440. Skin and skull. Adult female. Collected 20 Jan 1912 by E. Heller. Original number 5126.

Type Locality: Kibabe, Kisumu, [Nyanza], British East Africa 

(Kenya). 

Crocidura pergrisea G. S. Miller

Proc. Biol. Soc. Wash. 26: 113, 3 May 1913c.

Holotype: USNM 175918. Skin and skull. Adult female. Collected 16 Oct 1912 by W. L. Abbott. Original number 8036.

Type Locality: Skoro Loomba, [FC: valley to Shigar], altitude 

9,500 ft (2,896 m), Shigar, Baltistan (Gilgit-Baltistan), 

[Pakistan]. 

Crocidura planiceps Heller

Smithson. Misc. Collect. 56(15): 5, 23 Dec 1910a.

Holotype: USNM 164641. [Skin and skull]. Adult male. Collected 23 Jan 1910 by J. A. Loring. Original number 8837.

Type Locality: Rhino Camp, Lado Enclave, [Arua, Uganda]. 

Crocidura pullata G. S. Miller

Proc. Biol. Soc. Wash. 24: 241, 28 Nov 1911c.

Holotype: USNM 173213. Skin and skull. Adult male. Collected 

9 Oct 1910 by W. L. Abbott. Original number 7421.

Type Locality: [FC: Saagam], Kotihar [FC: Valley], altitude 7,000 

ft (2,134 m), [Jammu and Kashmir, India]. 

Crocidura raineyi Heller

Smithson. Misc. Collect. 60(12): 7, 4 Nov 1912a.

Holotype: USNM 181817. [Skin and skull]. Adult male. Collected Sep 1911 by E. Heller. Original number 4195.

Type Locality: [SL: North Creek], Mount Garguez (Warges), 

Mathews Range, altitude 6,000 ft (1,829 m), [Rift Valley], 

British East Africa (Kenya). 

Crocidura rhoditis Miller and Hollister

Proc. Biol. Soc. Wash. 34: 102, 30 Jun 1921.

Holotype: USNM 217550. Skin and skull. Adult male. Collected 

3 Aug 1916 by H. C. Raven. Original number 2834.

Type Locality: Temboan, [Sulawesi Utara], northeastern Celebes 

(Sulawesi), [Indonesia]. 

Crocidura russula hosletti Jameson and Jones

Proc. Biol. Soc. Wash. 90: 465, 17 Oct 1977.

= Crocidura shantungensis quelpartis Kuroda, 1934. See Jiang 

and Hoffmann (2001:1072).

Holotype: USNM 398640. Skin and skull. Adult male. Collected 

11 Oct 1963 by E. W. Jameson, Jr. Original number 1150.

Type Locality: 10 mi (16 km) W Taichung, Taichung Hsien, 

Taiwan. 

Crocidura shantungensis G. S. Miller

Proc. Biol. Soc. Wash. 14: 158, 9 Aug 1901e.

= Crocidura shantungensis shantungensis G. S. Miller, 1901.

Holotype: USNM 86151. Skin and skull. Adult, sex not given. 

Collected Jun 1898 by P. D. Bergen. No original number.

Type Locality: Chimeh, Shantung (Shandong), northern China. 

Crocidura sicula G. S. Miller

Proc. Biol. Soc. Wash. 14: 41, 25 Apr 1901c.

= Crocidura sicula sicula G. S. Miller, 1901.

Holotype: USNM 103301. Skin and skull. Adult male. Collected 

20 Jun 1900 by D. Coolidge. Original number 1332.

Type Locality: Palermo, Sicily (Sicilia), [Italy]. 

Crocidura simiolus Hollister

Smithson. Misc. Collect. 66(8): 3, 23 May 1916a.

= Crocidura zaphiri Dollman, 1915. See Osgood (1936:224).

Holotype: USNM 197959. Skin and skull. Adult female. Collected 25 Sep 1913 by H. J. A. Turner. Original number 3.

Type Locality: Kisumu, [Nyanza], British East Africa (Kenya). 

Crocidura suahelae Heller

Smithson. Misc. Collect. 60(12): 6, 4 Nov 1912a.

= Crocidura viaria (E. Geoffroy, 1834). See Hutterer (1984:209).

Holotype: USNM 181815. [Skin and skull]. Adult male. Collected 17 Dec 1911 by E. Heller. Original number 4875.

Type Locality: Mazeras, [Coast Province], British East Africa 

(Kenya). 

Crocidura sururae Heller

Smithson. Misc. Collect. 56(15): 2, 23 Dec 1910a.

= Crocidura olivieri sururae Heller, 1910. See Hutterer 

(2005:244).

Holotype: USNM 164637. [Skin and skull]. Adult male. Collected 18 Jan 1910 by J. A. Loring. Original number 8777.

Type Locality: Rhino Camp, Lado Enclave, [Arua, Uganda]. 

Crocidura tarfayaensis Vesmanis and Vesmanis

Zool. Abhand. Staat. Mus. Tierk. 36: 47, 30 Apr 1980.

Holotype: USNM 470590. S, B (skin and skull). Female, age not 

given. Leg. (Collected) 5 Dec 1969 by L. W. Robbins. Original number 2088.

Type Locality: 8 km S Tarfaya, 27o50′N, 12o30′W (Schmidt et al. 

2008:34: 27o53 ́N, 12o52 ́W), Tarfaya Provinz (Province), 

Morokko (Morocco). 

Crocidura tiznitensis Vesmanis and Vesmanis

Zool. Abhand. Staat. Mus. Tierk. 36: 54, 30 Apr 1980.

= Crocidura tarfayaensis Vesmanis and Vesmanis, 1980. See 

Hutterer (1987:530).
NUMBER 644 • 3 7

Holotype: USNM 470589. S, B (skin and skull). Male, age not 

given. Leg. (Collected) 20 X [SL and FC: Dec] 1969 by L. W. 

Robbins. Original number 2241.

Type Locality: 15 km no. (NE) Tiznit, 29o49′N, 9o39′W, Agadir 

Provinz (Province), Marokko (Morocco). 

Crocidura turba lakiundae Heller

Smithson. Misc. Collect. 60(12): 6, 4 Nov 1912a.

= Crocidura nigrofusca Matschie, 1895. See Hutterer (1993:91).

Holotype: USNM 181816. [Skin and skull]. Adult female. Collected 12 Jul 1911 by E. Heller. Original number 1986.

Type Locality: Lakiundu River, near its junction with the Northern Guaso Nyiro River, British East Africa (Kenya). 

Crocidura zaianensis Vesmanis and Vesmanis

Zool. Abhand. Staat. Mus. Tierk. 36: 62, 30 Apr 1980.

= Crocidura whitakeri de Winton, 1897. See Hutterer (1987:530).

Holotype: USNM 476071. S, B (skin and skull). Female, age not 

given. Leg. (Collected) 27 Apr 1970 by R. E. Vaden. Original number 5982.

Type Locality: 5 km N Ben-Slimane, 33o39′N, 7o08′W, Settat 

Provinz (Province), Marokko (Morocco). 

Heliosorex roosevelti Heller

Smithson. Misc. Collect. 56(15): 6, 23 Dec 1910a.

= Crocidura roosevelti (Heller, 1910). See Hollister (1918b:68).

Holotype: USNM 164643. [Skin and skull]. Adult male, not female as in original description. Collected 21 Jan 1910 by 

J. A. Loring. Original number 8812.

Type Locality: Rhino Camp, Lado Enclave, [Arua, Uganda]. 

Myosorex muricauda G. S. Miller

Proc. Wash. Acad. Sci. 2: 645, 28 Dec 1900c.

= Crocidura muricauda (G. S. Miller, 1900). See Hollister 

(1916b:669).

Holotype: USNM 83809. In alcohol, [skull removed]. Adult 

male. Collected 5 Apr 1897 by [SL: R. P. Currie]. Original 

number 32.

Type Locality: Mount Coffee, [Montserrado], Liberia, Africa. 

Genus Cryptotis Pomel, 1848

Blarina alticola Merriam

N. Am. Fauna 10: 27, 31 Dec 1895a.

= Cryptotis alticola (Merriam, 1895). See G. S. Miller (1912b:27).

Holotype: USNM 52047. [Skin and skull]. Adult male. Collected 

25 Feb 1893 by E. W. Nelson. Original number 4396.

Type Locality: Mount Popocatepetl, 11,500 ft (3,505 m), [SL: 

Mexico], Mexico. 

Blarina berlandieri Baird

Mammals Rep. Explor. Surv., 8(1): 53, 1857.

= Cryptotis parvus berlandieri (Baird, 1857). See Davis (1941:413).

Lectotype: USNM 2159 as designated by Lyon and Osgood 

(1909:327). In alcohol, [Poole and Schantz (1942:175): skull 

removed Apr 1936]. Age and sex not given. [MC: Prepared 

by J. L. Berlandier] and [Lyon and Osgood (1909:237): 

probably collected by him]. Date of collection not given. 

Received from D. N. Couch. [MC: Cataloged 14 Feb 1857].

Type Locality: Vicinity of Matamoras (Matamoros), [Tamaulipas], Mexico.

Remarks: No type was designated in the original description. 

Baird (1857:55) listed four specimens. Lyon and Osgood 

designated USNM 2159 as the type = lectotype (ICZN, 

1999: Article 74.5) because it was figured. 

Blarina exilipes Baird

Mammals Rep. Explor. Surv., 8(1): 51, 1857.

= Cryptotis parvus parvus (Say, 1823). See Merriam (1895a:17).

Lectotype: USNM 2157 as designated by Lyon and Osgood 

(1909:327). In alcohol, [Poole and Schantz (1942:175): 

skull removed Apr 1936], but remains in alcohol. Age and 

sex not given. Collected by B. L. C. Wailes, date unknown. 

[MC: Cataloged 14 Feb 1857].

Type Locality: Washington, [Adams County], Mississippi.

Remarks: No type was designated in the original description. 

Baird (1857:53) listed eight specimens. Lyon and Osgood 

designated USNM 2157 as the type = lectotype (ICZN, 

1999; Article 74.5) because it was figured. 

B[larina]. eximius Baird

Mammals Rep. Explor. Surv., 8(1): 52, 1857.

= Cryptotis parvus parvus (Say, 1823). See Merriam (1895a:17).

Syntypes: USNM 639/1790. Skin [cannot be found] and skull. 

Male, age not given. Obtained from G. Engleman, date 

unknown. [MC: Cataloged 19 Apr 1855. Original number 42].

USNM 2166/3109. Skin and skull. Age and sex not 

given. Obtained from R. Kennicott, date unknown. [MC: 

Skin cataloged 17 Feb 1858; skull Feb or Mar 1857]. No 

original number.

Type Locality: Saint Louis, [Saint Louis City], Missouri and 

DeKalb County, Illinois.

Remarks: Baird (1857:52 and 53) based this taxon soley on the 

two specimens listed above. They are therefore syntypes 

(ICZN, 1999: Article 73.2) and the type locality encompasses both localities (ICZN, 1999: Article 73.2.3). 

Blarina floridana Merriam

N. Am. Fauna 10: 19, 31 Dec 1895a.

= Cryptotis parvus floridanus (Merriam, 1895). See Harper 

(1927:270).

Holotype: USNM 16510/23937. [In alcohol, skull removed]. 

Age and sex not given. Collected 22 Apr 1889 by M. M. 

Green. Original number 44.

Type Locality: Chester Shoal, 11 mi (18 km) north of Cape Canaveral, Brevard County, Florida. 

Blarina fossor Merriam

N. Am. Fauna 10: 28, 31 Dec 1895a.
3 8 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

= Cryptotis goldmani machetes (Merriam, 1895). See Hutterer 

(2005:271).

Holotype: USNM 68545. [Skin and skull]. Adult female. Collected 10 Jul 1894 by E. W. Nelson and E. A. Goldman. 

Original number 6419.

Type Locality: Mount Zempoaltepec, altitude 10,500 ft (3,200 m) 

[FC: 8,200 ft (2,499 m)], Oaxaca, Mexico. 

Blarina magna Merriam

N. Am. Fauna 10: 28, 31 Dec 1895a.

= Cryptotis magnus (Merriam, 1895). See G. S. Miller (1912b:28).

Holotype: USNM 68575. [Skin and skull]. Old male. Collected 

24 Jul 1894 by E. W. Nelson and E. A. Goldman. Original 

number 6493.

Type Locality: Totontepec, altitude 6,800 ft (2,073 m), Oaxaca, 

[SL: Mexico]. 

Blarina mayensis Merriam

Proc. Wash. Acad. Sci. 3: 559, 29 Nov 1901b.

= Cryptotis mayensis (Merriam, 1901). See G. S. Miller 

(1912b:26).

Holotype: USNM 108087. [Skin and skull]. Adult female. Collected 5 Feb 1901 by E. W. Nelson and E. A. Goldman. 

Original number 14495.

Type Locality: Maya ruin at Chichenitza (Chichen Itza), Yucatan, [SL: Mexico]. 

Blarina (Soriciscus) mexicana Coues

Bull. U.S. Geol. and Geogr. Surv. Terr. 3: 652, 15 May 1877.

= Cryptotis mexicanus (Coues, 1877). See G. S. Miller 

(1911d:221).

Holotype: USNM 3525/4438. Skin and skull. Age and sex not given. 

Collected by R. Montes de Oca, date not given. [MC: Skin cataloged 18 Mar 1859; skull Sep 1861]. No original number.

Type Locality: Xalapa, [Veracruz], Mexico. 

Blarina mexicana goldmani Merriam

N. Am. Fauna 10: 25, 31 Dec 1895a.

= Cryptotis goldmani goldmani (Merriam, 1895). See Choate 

(1970:247).

Holotype: USNM 70244. [Skin and skull]. Young adult male. 

Collected 23 Dec 1894 by E. W. Nelson and E. A. Goldman. 

Original number 7231.

Type Locality: Mountains near Chilpancingo, altitude 10,000 ft 

(3,048 m) [FC: 9,600 ft (2,926 m)], Guerrero, Mexico. 

Blarina mexicana machetes Merriam

N. Am. Fauna 10: 26, 31 Dec 1895a.

= Cryptotis goldmani machetes (Merriam, 1895). See Hutterer 

(2005:271).

Holotype: USNM 71456. [Skin and skull]. Adult female. Collected 26 Mar 1895 by E. W. Nelson and E. A. Goldman. 

Original number 7723.

Type Locality: Mountains near Ozolotepec, altitude 10,000 ft 

(3,048 m), Oaxaca, [SL: Mexico]. 

Blarina mexicana peregrina Merriam

N. Am. Fauna 10: 24, 31 Dec 1895a.

= Cryptotis peregrinus (Merriam, 1895). See Woodman and 

Timm (1999:1).

Holotype: USNM 68317. [Skin and skull]. Adult male. Collected 

12 Sep 1894 by E. W. Nelson and E. A. Goldman. Original 

number 6748.

Type Locality: Mountains 15 mi (24 km) west of Oaxaca, altitude 9,500 ft (2,896 m), [SL: Oaxaca], Mexico. 

Blarina nelsoni Merriam

N. Am. Fauna 10: 26, 31 Dec 1895a.

= Cryptotis nelsoni (Merriam, 1895). See G. S. Miller (1912b:27).

Holotype: USNM 65437. [Skin and skull]. Adult [SL: female]. 

Collected 13 May 1894 by E. W. Nelson and E. A. Goldman. Original number 6253.

Type Locality: Volcano of Tuxtla (Volcan Tuxtla), altitude 4,800 

ft (1,463 m), Veracruz, Mexico. 

Blarina obscura Merriam

N. Am. Fauna 10: 23, 31 Dec 1895a.

= Cryptotis obscurus (Merriam, 1895). See G. S. Miller 

(1912b:26).

Holotype: USNM 55634. [Skin and skull]. Young adult female. Collected 27 Aug 1893 by E. W. Nelson. Original number 5377.

Type Locality: Tulancingo, altitude 8,500 ft (2,591 m), Hidalgo, 

Mexico. 

Blarina soricina Merriam

N. Am. Fauna 10: 22, 31 Dec 1895a.

= Cryptotis parvus soricinus (Merriam, 1895). See Choate 

(1970:267).

Holotype: USNM 50762. [Skin and skull]. Adult male. Collected 

5 Dec 1892 by E. W. Nelson. Original number 3989.

Type Locality: Tlalpam, Valley of Mexico, [Miller and Rehn 

(1901:247): 10 mi (16 km) south of the city of Mexico], 

altitude 7,600 ft (2,317 m), [SL and FC: DF (Distrito Federal)], [Mexico]. 

Cryptotis frontalis G. S. Miller

Proc. Biol. Soc. Wash. 24: 222, 31 Oct 1911d.

= Cryptotis goldmani machetes (Merriam, 1895). See Hutterer 

(2005:271).

Holotype: USNM 123429. In alcohol, [skull removed]. Adult 

female. Collected [Poole and Schantz (1942:176): about 

1870] by F. Sumichrast. No original number.

Type Locality: Near Tehuantepec City, [Oaxaca], Mexico. 

Cryptotis goodwini Jackson

Proc. Biol. Soc. Wash. 46: 81, 27 Apr 1933.

Holotype: USNM 77074. Skin and skull. Adult male. Collected 

13 Jan 1896 by E. W. Nelson and E. A. Goldman. Original 

number 9073.

Type Locality: Calel, altitude 10,200 ft (3,109 m), [Quezaltenango], Guatemala. 
NUMBER 644 • 3 9

Cryptotis gracilis G. S. Miller

Proc. Biol. Soc. Wash. 24: 221, 31 Oct 1911d.

Holotype: USNM 12236/38471. Skin and skull. Adult, sex not 

given. Collected on unknown date by W. M. Gabb. [MC: 

Skin cataloged 19 Nov 1874; skull between 21 Mar and 31 

May 1911]. No original number.

Type Locality: Head of Lari River, near base of Pico Blanco, altitude about 6,000 ft (1,829 m), Talamanca, [Puntarenas], 

Costa Rica. 

Cryptotis griseoventris Jackson

Proc. Biol. Soc. Wash. 46: 80, 27 Apr 1933.

Holotype: USNM 75894. Skin and skull. Adult male. Collected 

4 Oct 1895 by E. W. Nelson and E. A. Goldman. Original 

number 8545.

Type Locality: San Cristobal, altitude 9,500 ft (2,896 m), Chiapas, Mexico. 

Cryptotis guerrerensis Jackson

Proc. Biol. Soc. Wash. 46: 80, 27 Apr 1933.

= Cryptotis goldmani goldmani (Merriam, 1895). See Choate 

(1970:247).

Holotype: USNM 126895. Skin and skull. Adult female. Collected 17 May 1903 by E. W. Nelson and E. A. Goldman. 

Original number 16429.

Type Locality: Omilteme (Omiltemi), altitude about 8,000 ft 

(2,438 m), Guerrero, Mexico. 

Cryptotis jacksoni Goodwin

Am. Mus. Novit. 1267: 1, 10 Dec 1944.

= Cryptotis gracilis G. S. Miller, 1911. See Choate (1970:281).

Holotype: USNM 116649. Adult female. Skin and skull. Collected 

27 Feb 1902 by M. Cary. Original number [SL: 540. 3549-X].

Type Locality: [XC: Estancia Jimenez], Volcan Irazu, [SL: Costa 

Rica]. 

Cryptotis lacertosus Woodman

J. Mammal. 91: 576, 16 Jun 2010.

Holotype: USNM 569443. Skin and skull. Adult female. Collected 30 Jul 2005 by W. Bulmer, R. P. Eckerlin, and J. O. 

Matson. Original number JOM 7124.

Type Locality: 5 km SW San Mateo Ixtatan, 3,110 m, [FC: 

15o48.604′N, 91o30.326′W], Huehuetenango, Guatemala. 

Cryptotis mam Woodman

J. Mammal. 91: 573, 16 Jun 2010.

Holotype: USNM 77053. Skin and skull. Adult male. Collected 

27 Dec 1895 by E. W. Nelson and E. A. Goldman. Original 

number 8908.

Type Locality: Above Todos Santos Cuchumatan, approximately 

10,000 ft (3,048 m), approximately 15o36′N, 91o37′W, 

Huehuetenango, Guatemala. 

Cryptotis merus Goldman

Smithson. Misc. Collect. 60(2): 17, 20 Sep 1912b.

Holotype: USNM 178976. [Skin and skull]. Adult male. Collected 

2 May 1912 by E. A. Goldman. Original number 21669.

Type Locality: Near head of Rio Limon, Mount Pirri, altitude 

4,500 ft (1,372 m), [Darien], eastern Panama. 

Cryptotis nigrescens merriami Choate

Univ. Kans. Publ., Mus. Nat. Hist. 19: 277, 30 Dec 1970.

= Cryptotis merriami Choate, 1970. See Woodman and Timm 

(1993:14).

Holotype: USNM 77050. Skin and skull. Adult female. Collected 21 Dec 1895 by E. W. Nelson and E. A. Goldman. 

Original number 8846.

Type Locality: Jacaltenango, 5,400 ft (1,646 m), Huehuetenango, 

Guatemala. 

Cryptotis oreoryctes Woodman

Zool. J. Linn. Soc. 163: 1280, 25 Nov 2011.

Holotype: USNM 569877. Skin, skull, and partial skeleton. 

Adult female. Collected 4 i (Jan) 2007 by N. Woodman. 

Original number J. O. Matson 7288.

Type Locality: Chelemha Cloud Forrest Reserve, [SL: Yaliju 

Mountain], 2,090 m.a.m.s.l. (m), about 15o23 ́N, 90o04 ́W, 

Alta Verapaz, Guatemala. 

Cryptotis pergracilis macer G. S. Miller

Proc. Biol. Soc. Wash. 24: 223, 31 Oct 1911d.

= Cryptotis parvus berlandieri (Baird, 1857). See Choate 

(1970:260).

Holotype: USNM 15565/38494. In alcohol, [skull removed]. 

Adult female. Collected on unknown date by A. Duges. 

[MC: Skin cataloged 19 Oct 1886; skull 30 Aug 1911]. 

Original number [MC: 103].

 Type Locality: Near Guanajuato city, [Guanajuato], Mexico. 

Cryptotis pergracilis nayaritensis Jackson

Proc. Biol. Soc. Wash. 46:79, 27 Apr 1933.

= Cryptotis parvus berlandieri (Baird, 1857). See Choate 

(1970:260).

Holotype: USNM 88015. Skin and skull. Adult male. Collected 

13 Apr 1897 by E. W. Nelson and E. A. Goldman. Original 

number 10902.

Type Locality: Tepic, altitude 3,000 ft (914 m), Nayarit, Mexico. 

Cryptotis pergracilis pueblensis Jackson

Proc. Biol. Soc. Wash. 46:79, 27 Apr 1933.

= Cryptotis parvus pueblensis Jackson, 1933. See Choate 

(1970:264).

Holotype: USNM 92720. Skin and skull. Adult male. Collected 

6 Jan 1898 by E. A. Goldman. Original number Nelson and 

Goldman 12014.

Type Locality: Huachinango, altitude 5,000 ft (1,524 m), Puebla, 

Mexico. 

Cryptotis tamensis Woodman

Proc. Biol. Soc. Wash. 115: 254, 2 Jul 2002.
4 0 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Holotype: USNM 418567. Skin, skull, and body in fluid. Adult 

female. Collected 8 Mar 1968 by the Smithsonian Venezuelan Project team led by N. E. Peterson. Original number 

SVP 21863.

Type Locality: Near Paramo de Tama, 7o27′N, 72o26′W, 2,415m, 

35 km S, 22 km W of San Cristobal, reported as 41 km 

SW San Cristobal and 12 km SSE Las Delicias by Handley 

(1976:64), Buena Vista, State of Tachira, Venezuela. 

Cryptotis zeteki Setzer

J. Wash. Acad. Sci. 40: 299, 15 Sep 1950.

= Cryptotis nigrescens (J. A. Allen, 1895). See Choate (1970:279).

Holotype: USNM 290466. In alcohol, skull removed. Adult female. Collected Apr 1949 by E. Graetz. No original number.

Type Locality: Cerro Punta, 6,500 ft (1,829 m), lat. 8o42′N, 

long. 82o48′W, Chiriqui Province, Republic of Panama. 

Sorex cinereus Bachman

J. Acad. Nat. Sci. Phila. 7: 373, 1837.

= Cryptotis parvus parvus (Say, 1823). See G. S. Miller 

(1912b:24).

Syntype: USNM 94/1771. Skin and skull. Age and sex not given. 

Collected by W. Wesner, date unknown. [MC: Skin cataloged 26 Apr 1852; skull 18 Apr 1855]. No original number.

Type Locality: Goose Creek, about 22 mi (35 km) from Charleston. Restricted by Handley and Varn (1994:379) to Goose 

Creek, about 22 mi (35 km) NNW by road of Charleston, 

Berkeley County, South Carolina.

Remarks: Bachman (1837) based his description on six specimens from the type locality. According to Handley and Varn 

(1994:397) three of the six specimens are known to exist; 

two are at the Academy of Natural Sciences of Philadelphia 

(Academy of Natural Sciences of Drexel University), although Koopman (1976:3) did not list them; and the one 

given above, which was received in exchange with the Academy. [MC: The skin, USNM 94, was transferred to the University of Michigan 1 Apr 1859], but Handley could not find 

it there on 31 May 1954 (Handley and Varn, 1994:397).

Genus Megasorex Hibbard, 1950

Notiosorex gigas Merriam

Proc. Biol. Soc. Wash. 11: 227, 15 Jul 1897c.

= Megasorex gigas (Merriam, 1897). See Hibbard (1950:127).

Holotype: USNM 88012. [Skin and skull]. Adult female. Collected 15 Mar 1897 by E. W. Nelson and E. A. Goldman. 

Original number 10706.

Type Locality: Mountains at Milpillas, [FC: altitude 3,800 ft 

(1,158 m)], near San Sebastian, Jalisco, Mexico. 

Genus Neomys Kaup, 1829

Neomys fodiens minor G. S. Miller

Proc. Biol. Soc. Wash. 14: 45, 25 Apr 1901c.

= Neomys fodiens (Pennant, 1771). Subspecies not recognized by 

Hutterer (2005:279).

Holotype: USNM 101311. Skin and skull. Adult male. Collected 

8 Jul 1899 by R. T. Young. Original number 641.

Type Locality: Montrejeau, in the foothills of Pyrenees, Haut 

Pyrenees (Haute-Garonne Department), [Mid-Pyrenees], 

France. 

Genus Notiosorex Coues, 1877

Sorex (Notiosorex) crawfordi Coues

Bull. U.S. Geol. and Geogr. Surv. Terr. 3: 646, 15 May 1877.

= Notiosorex crawfordi (Coues, 1877). See Merriam (1895a: 

32).

Holotype: USNM 2653/4437. In alcohol, skull removed. Age 

and sex not given. Collected on unknown date by S. W. 

Crawford. [MC: Alcoholic cataloged 28 Apr 1857; skull 

Sep 1861]. No original number.

Type Locality: Fort Bliss or vicinity, [Miller and Rehn (1901:244): 

about 2 mi (3.2 km) above El Paso, El Paso County], New 

Mexico (Texas). 

Sorex (Notiosorex) evotis Coues

Bull. U.S. Geol. and Geogr. Surv. Terr. 3: 652, 15 May 1877.

= Notiosorex evotis (Coues, 1877). See Carraway and Timm 

(2000:310).

Holotype: USNM 9066. [Skull only]. Age and sex not given. 

Collected [SL: Feb 1868] by F. Bischoff. 

Type Locality: [SL: Rio] Mazatlan, [Sinaloa], Mexico. 

Genus Sorex Linnaeus, 1758

Atophyrax bendirii Merriam

Trans. Linn. Soc. New York 2: 217, 28 Aug 1884.

= Sorex bendirii bendirii (Merriam, 1884). See Dobson (1890:fig. 

17).

Holotype: USNM 186442. In alcohol [currently a skin, skull, 

and partial skeleton]. Adult male. Collected 1 Aug 1882 by 

C. E. Bendire. No original number. [SL: MCC 496/1200].

Type Locality: About 1 mi (1.6 km) from Williamson’s River, 

some 18 mi (29 km) southeast of Fort Klamath, [Klamath 

County], Oregon.

Remarks: Type not designated by number in original description, 

but it was expressly stated that the description was based on 

a single specimen. The specimen listed above is the only one 

in the collection matching the data given in the original description and therefore is the holotype by monotypy (ICZN, 

1999: Article 73.1.2). Originally part of the Merriam Collection, this specimen was cataloged and assigned USNM 

186442 on 13 Oct 1911. 

Microsorex hoyi intervectus Jackson

Proc. Biol. Soc. Wash. 38: 125, 13 Nov 1925a.

= Sorex hoyi hoyi Baird, 1857. See Diersing (1980:91).
NUMBER 644 • 4 1

Holotype: USNM 226979. Skin and skull. Adult female. Collected 

17 Aug 1917 by H. H. T. Jackson. Original number 820.

Type Locality: Lakewood, Oconto County, Wisconsin. 

Microsorex hoyi montanus Brown

Proc. Biol. Soc. Wash. 79: 50, 23 May 1966.

= Sorex hoyi montanus (Brown, 1966). See Diersing (1980:99).

Holotype: USNM 399498. Skin and skull. Adult male. Collected 

30 Sep 1963 by L. N. Brown. Original number [SL: 1449].

Type Locality: Edge of Trail’s Divide Pond, [SL: 10,000 ft (3,048 

m)], 1/4 mi (0.4 km) S of University of Wyoming Summer 

Science Camp on State Highway 130, [Long (1972:190): 6 

mi (10 km) W] Centennial, [SL: Medicine Bow Mountains], 

Albany County, Wyoming.

Remarks: Published as University of Wyoming Museum of Zoology specimen number K-307. 

Microsorex hoyi washingtoni Jackson

Proc. Biol. Soc. Wash. 38: 125, 13 Nov 1925a.

= Sorex hoyi hoyi Baird, 1857, See Diersing (1980:91).

Holotype: USNM 91007. Skin and skull. Adult female. Collected 

26 Sep 1897 by V. Bailey. Original number 6293.

Type Locality: Loon Lake, Stevens County, Washington. 

Microsorex winnemana Preble

Proc. Biol. Soc. Wash. 23: 101, 24 Jun 1910.

= Sorex hoyi winnemana (Preble, 1910). See Diersing (1980:98).

Holotype: USNM 126320. Skin and skull. Young adult female. 

Collected 25 Apr 1903 by E. A. Preble. No original number. 

[SL: 3929-X].

Type Locality: Bank of Potomac River near Stubblefield Falls, 

[SL: 4 mi (6 km) below Great Falls], Fairfax County, 

Virginia. 

Neosorex albibarbis Cope

Proc. Acad. Nat. Sci. Phila. 14: 188, ordered published 29 Apr 

1862. Receipt acknowledged 27 May 1862 by the Smithsonian Institution.

= Sorex albibarbis albibarbis (Cope, 1862). See Hope et al. 

(2014:730) for current use.

Syntype: USNM 11239/38743. In alcohol, skull removed. Adult 

female. Collected Sep 1859 by E. D. Cope. No original 

number.

Type Locality: Profile Lake, Franconia Mountains, [Grafton 

County], New Hampshire.

Remarks: No type was designated in original description. Cope 

(1862) had two specimens, both from the locality given 

above. The whereabouts of the second specimen is unknown. The alcoholic part of this specimen was destroyed 

and its skeleton preserved on 6 Sep 1956, so it is now a skull 

and skeleton. 

Neosorex navigator Baird

Mammals Rep. Explor. Surv. 8(1): 11, 1857.

= Sorex navigator navigator (Baird, 1857). See Hope et al. 

(2014:730) for current use.

Holotype: USNM 629/1780. Skin and skull. Age and sex not 

given. Collected [note on SL: 31 Aug 1853] by J. G. Cooper. 

Original number [MC: 37].

Type Locality: Fort Vancouver, [SL: Columbia River], WT. Restricted by Jackson (1928:186) to near head of Yakima 

River, Cascade Mountains, [Kittitas County], Washington.

Remarks: A note on the skin label states, “according to Dr. Cooper found in lake near summit of Cascade Mountains.” 

Sorex alascensis shumaginensis Merriam

Proc. Wash. Acad. Sci. 2: 18, 14 Mar 1900.

= Sorex monticola shumaginensis Merriam, 1900. See Hennings 

and Hoffmann (1977:14).

Holotype: USNM 97993. [Skin and skull]. Adult male. Collected 

[SL: 17] Jul 1899 by De Alton Saunders. Original number 

[FC: A. K.] Fisher 2210.

Type Locality: Popof Island, Shumagin Islands, Alaska. 

Sorex (Microsorex) alnorum Preble

N. Am. Fauna 22: 72, 31 Oct 1902.

= Sorex hoyi alnorum Preble, 1902. See Hutterer (2005:289).

Holotype: USNM 107014. Skin and skull. Adult female. Collected 27 Jun 1900 by E. A. and A. E. Preble. Original number [FC: EAP] 2662.

Type Locality: [FC: Lower end] Robinson Portage, [Anderson 

(1947:22): Upper Hays River about 35 mi (56 km) southwest of Oxford Lake at about lat. 54o30 ́N, long. 96oW], 

Keewatin (Manitoba), Canada. 

Sorex alpinus hercynicus G. S. Miller

Ann. Mag. Nat. Hist. (series 8) 3: 417, May 1909.

= Sorex alpinus Schinz, 1837. Subspecies of S. alpinus not recognized by Hutterer (2005:253).

Holotype: USNM 112928. Skin and skull. Adult male. Collected 

10, not 18 as in original description, Oct 1901 by F. L. J. 

Boettcher. Original number 265.

Type Locality: Mauseklippe, Bodethal (Bodetal), [FC: 630 m], 

Harz Mountains, [Sachsen-Anhalt], Germany. 

Sorex amoenus Merriam

N. Am. Fauna 10: 69, 31 Dec 1895b.

= Sorex vagrans vagrans Baird, 1857. See Findley (1955:52).

Holotype: USNM 29784/41863. Skin and skull. Adult male. 

Collected 22 Jul 1891 by E. W. Nelson. Original number 

1129.

Type Locality: Mammoth Pass, altitude 10,000 ft (3,048 m), head 

of Owens River, east slope of the Sierra Nevada, California. 

Restricted by Howell (1923:266) to immediate vicinity of 

Mammoth, 8,000 ft (2,438 m), Mono County, California. 

Sorex araneus alticola G. S. Miller

Proc. Biol. Soc. Wash. 14: 43, 25 Apr 1901c.
4 2 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

= Sorex araneus Linnaeus, 1758. Subspecies of S. araneus not 

recognized by Hutterer (2005:283).

Holotype: USNM 85930. Skin and skull. Adult male. Collected 

17 Oct 1898 by J. A. Loring. Original number 5731, not 

5782 as in original description.

Type Locality: Near Meiringen, altitude 2,100 m, [Bern], Switzerland. 

Sorex araneus bergensis G. S. Miller

Ann. Mag. Nat. Hist. (series 8) 3: 416, May 1909.

= Sorex araneus Linnaeus, 1758. Subspecies of S. araneus not 

recognized by Hutterer (2005:283).

Holotype: USNM 84664. Skin and skull. Adult female. Collected 

10 Jun 1898 by T. Stejneger. Original number 31.

Type Locality: [FC: Ystaas, about 1,000 ft (305 m)], Gravin 

(Granvin), Hardanger, [Hordaland], Norway. 

Sorex araneus euronotus G. S. Miller

Proc. Biol. Soc. Wash. 14: 44, 25 Apr 1901c.

= Sorex coronatus Millet, 1828. See Meylan and Hausser 

(1978:120) who placed euronotus in coronotus as a valid 

subspecies. Hutterer (2005:287) did not recognize subspecies of S. coronatus.

Holotype: USNM 101321. Skin and skull. Adult male. Collected 

8 Jul 1899 by R. T. Young. Original number 642.

Type Locality: Montrejeau, in foothills of Pyrenees, Haute Pyrenees (Hautes-Pyrenees), [Midi-Pyrenees], France. 

Sorex arcticus laricorum Jackson

Proc. Biol. Soc. Wash. 38: 127, 13 Nov 1925b.

Holotype: USNM 186837. Skin and skull. Adult male. Collected 

19 Feb 1886 by V. Bailey. Original number 75. Old (MCC) 

number 2019/2552.

Type Locality: Elk River, Sherburne County, Minnesota. 

Sorex bairdi Merriam

N. Am. Fauna 10: 77, 31 Dec 1895b.

= Sorex bairdi bairdi Merriam, 1895.

Holotype: USNM 17414/24318. Skin and skull. Adult female. 

Collected 2 Aug 1889 by T. S. Palmer. Original number 270.

Type Locality: Astoria, [Clatsop County], Oregon. 

Sorex (Atophyrax) bendirii albiventer Merriam

N. Am. Fauna 10: 97, 31 Dec 1895b.

= Sorex bendirii albiventer Merriam, 1895. See Jackson 

(1926:58).

Holotype: USNM 66198. [Skin and skull]. Adult male. Collected 

7 Jul 1894 by C. P. Streator. Original number 4021.

Type Locality: Lake Cushman, Olympic Mountains, [Mason 

County], Washington. 

Sorex (Atophyrax) bendirii palmeri Merriam

N. Am. Fauna 10: 97, 31 Dec 1895b.

= Sorex bendirii palmeri Merriam, 1895. See Jackson (1926:58).

Holotype: USNM 17338/24263. Skin and skull. Old female. 

Collected 29 Jul 1889 by T. S. Palmer. Original number 256.

Type Locality: Astoria, [Clatsop County], Oregon. 

Sorex californicus Merriam

N. Am. Fauna 10: 80, 31 Dec 1895b.

= Sorex ornatus ornatus Merriam, 1895. See Hutterer (2005:294).

Holotype: USNM 32578/44426. Skin and skull. Adult male. Collected 15 Feb 1892 by C. P. Streator. Original number 1583.

Type Locality: Walnut Creek, Contra Costa County, California. 

Sorex californicus juncensis Nelson and Goldman

Proc. Biol. Soc. Wash. 22: 27, 10 Mar 1909.

= Sorex ornatus juncensis Nelson and Goldman, 1909. See Junge 

and Hoffmann (1981:34).

Holotype: USNM 139594. [Skin and skull]. Young adult, sex 

unknown. Collected 1 Sep 1905 by E. W. Nelson and E. A. 

Goldman. Original number 17916.

Type Locality: Socorro, [FC: 10 ft (3 m)], 15 mi (24 km) south 

of San Quintin, Lower California (Baja California), Mexico. 

Sorex dispar blitchi Schwartz

J. Elisha Mitchell Sci. Soc. 72: 26, 1 May 1956.

Holotype: USNM 301993. [Skin and skull]. Male, age not given. 

Collected 30 Oct 1955 by E. L. Blitch and A. Schwartz. 

Original number AS 3525.

Type Locality: 2 mi (3.2 km) NE Wagon Road Gap, elevation 

4,525 ft (1,379 m), Haywood County, North Carolina. 

Sorex dobsoni Merriam

N. Am. Fauna 5: 33, 30 Jul 1891.

= Sorex vagrans vagrans Baird, 1857. See Findley (1955:52).

Holotype: USNM 24274/31678. Skin and skull. Adult female. 

Collected 3 Oct 1890 by C. H. Merriam and V. Bailey. Original number [FC: VB] 1929.

Type Locality: Sawtooth or Alturas Lake, [G. S. Miller (1912b: 

14): east base of] Sawtooth Mountains, altitude about 

2,200 m (7,200 ft), [Blaine County], Idaho. 

Sorex durangae Jackson

Proc. Biol. Soc. Wash. 38: 127, 13 Nov 1925b.

= Sorex monticola monticola Merriam, 1890. See Hennings and 

Hoffmann (1977:15).

Holotype: USNM 94540. Skin and skull. Adult male. Collected 

19 Jul 1898 by E. W. Nelson and E. A. Goldman. Original 

number 12774.

Type Locality: El Salto, [FC: altitude 8,400 ft (2,560 m)], Durango, Mexico. 

Sorex emarginatus Jackson

Proc. Biol. Soc. Wash. 38: 129, 13 Nov 1925b.

Holotype: USNM 90847. Skin and skull. Young adult female. 

Collected 17 Sep 1897 by E. W. Nelson and E. A. Goldman. 

Original number 11765.
NUMBER 644 • 4 3

Type Locality: Sierra Madre near Bolanos, altitude 7,600 ft 

(2,317 m), Jalisco, Mexico. 

Sorex (Microsorex) eximius Osgood

N. Am. Fauna 21: 71, 26 Sep 1901a.

= Sorex hoyi eximius Osgood, 1901. See Diersing (1980:89).

Holotype: USNM 107126. [Skin and skull]. Adult female. Collected 14 Sep 1900 by W. H. Osgood and E. Heller. Original 

number [FC: WHO] 1395.

Type Locality: Tyonek, Cook Inlet, Alaska. 

Sorex fimbripes Bachman

J. Acad. Nat. Sci. Phila. 7: 391, 1837.

= Sorex cinereus cinereus Kerr, 1792. See Jackson (1928:40).

Holotype: USNM 84556. [In alcohol, skull removed]. Age and 

sex not given. Collected on unknown date by W. R. Johnson. No original number. [MC: Cataloged 29 Apr 1898].

Type Locality: On the high table-land on a branch of Drury’s 

(Drury) Run, a tributary of the west bank (branch?) of the 

Susquehanna River, [Clinton County, Pennsylvania].

Remarks: A type specimen was not designated by number in 

the original description. However, the taxon was based on 

a single specimen and for many reasons Lyon and Osgood 

(1909:243) and Poole and Schantz (1942:185) considered 

USNM 84556 to be that specimen; holotype fixed by monotypy (ICZN, 1999: Article 73.1.3). According to Coues 

(1877:641), Baird examined Bachman’s type preserved in 

the Philadelphia Academy. How it ended up in the USNM is 

unknown. Koopman (1976) makes no mention of it being in 

the Academy of Natural Sciences of Philadelphia (Academy 

of Natural Sciences of Drexel University). 

Sorex fisheri Merriam

N. Am. Fauna 10: 86, 31 Dec 1895b.

= Sorex longirostris fisheri Merriam, 1895. See Jackson (1928:87).

Holotype: USNM 75166. [Skin and skull]. Adult male. Collected 

11 Oct 1895 by A. K. Fisher. Original number 1800.

Type Locality: Lake Drummond, Dismal Swamp, [Chesapeake/

Suffolk City], Virginia. 

Sorex fontinalis Hollister

Proc. U.S. Natl. Mus. 40: 378, 17 Apr 1911a.

= Sorex cinereus fontinalis Hollister, 1911. See Poole (1937:96).

Holotype: USNM 85439. Skin and skull. Old adult female. Collected 6 Nov 1898 by G. S. Miller, Jr. No original number.

Type Locality: Cold Springs Swamp, near Beltsville, [Prince 

Georges County], Maryland. 

Sorex fumeus umbrosus Jackson

Proc. Biol. Soc. Wash. 30: 149, 27 Jul 1917.

Holotype: USNM 150065. Skin and skull. Adult male. Collected 

29 Jul 1907 by W. H. Osgood. Original number 3140.

Type Locality: James River, Antigonish County, Nova Scotia, 

[Canada]. 

Sorex glacialis Merriam

Proc. Wash. Acad. Sci. 2: 16, 14 Mar 1900.

= Sorex monticola alascensis Merriam, 1895. See Alexander 

(1996:241).

Holotype: USNM 97709. [Skin and skull]. Adult male. Collected 

12 Jun 1899 by A. K. Fisher. Original number 2056.

Type Locality: Point Gustavus, on east side of entrance to Glacier 

Bay, Alaska. 

Sorex godmani Merriam

Proc. Biol. Soc. Wash. 11: 229, 15 Jul 1897c.

= Sorex salvini salvini Merriam, 1897. See Woodman at al. 

(2012:214) for current use.

Holotype: USNM 77044. Skin and skull. Adult female. Collected 28 Jan 1896 by E. W. Nelson and E. A. Goldman. 

Original number 9239.

Type Locality: Volcano (Volcan) Santa Maria, altitude 9,000 ft 

(2,743 m), Quezaltenango, Guatemala. 

Sorex haydeni Baird

Mammals Rep. Explor. Surv. 8(1): 29, 1857.

Lectotype: USNM 1685 as designated by Lyon and Osgood 

(1909:244). In alcohol, skull not removed. Male, age not 

given. Collected by F. V. Hayden. No original number.

Type Locality: Fort Union, [Lyon and Osgood (1909:244): 

near Fort Buford], [Williams County], Nebraska (North 

Dakota).

Remarks: A type was not designated by number in original description. Lyon and Osgood (1909:244) selected the specimen given above as the type = lectotype (ICZN, 1999: 

Article 74.5). 

Sorex hoyi Baird

Mammals Rep. Explor. Surv. 8(1): 32, 1857.

= Sorex hoyi hoyi Baird, 1857. See Diersing (1980:90).

Lectotype: USNM 632/1783 as designated by Jackson (1928:203). 

Skin [and skull]. [Lyon and Osgood (1909:244): Old] male. 

[SL: Collected by P. R. Hoy], date unknown. [MC: Cataloged 19 Apr 1855].

Type Locality: Racine, [Racine County], Wisconsin.

Remarks: Type not designated by number in original description. 

Baird had two specimens and Lyon and Osgood (1909:244) 

considered one of them, USNM 1688, as the type. However, Preble (1910:203) determined USNM 1688 to be a 

specimen of Sorex cinereus and regarded the skull on which 

Baird based his description, USNM 1783, as the type. In his 

revision, Jackson (1928:203) formally designated the skin 

and skull as lectotype. 

Sorex idahoensis Merriam

N. Am. Fauna 5: 32, 30 Jul 1891.

= Sorex cinereus cinereus Kerr, 1792. Unassigned to subspecies 

by Hutterer (2005:286), but see Jackson (1928:41) who assigned it to S. c. cinereus, as did Hall (1981:28).
4 4 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Holotype: USNM 23527/30945. Skin and skull. Adult female. 

Collected 26 Aug 1890 by C. H. Merriam and V. Bailey. 

Original number [FC: VB] 1674.

Type Locality: Timber Creek, altitude about 2,500 m (8,200 ft), 

[FC: 10 mi (16 km) west of Junction], Salmon River (Lemhi) 

Mountains, [Lemhi County], Idaho. 

Sorex lagunae Nelson and Goldman

Proc. Biol. Soc. Wash. 22: 27, 10 Mar 1909.

= Sorex ornatus lagunae Nelson and Goldman, 1909. See Jackson (1928:169).

Holotype: USNM 147119. [Skin and skull]. Adult female. Collected 29 Jan 1906 by E. W. Nelson and E. A. Goldman. 

Original number 19036.

Type Locality: La Laguna, altitude 5,500 ft (1,676 m), Sierra 

Laguna (Sierra de la Laguna), Lower California (Baja California Sur), Mexico. 

Sorex leucogenys Osgood

Proc. Biol. Soc. Wash. 22: 52, 17 Apr 1909.

= Sorex merriami Dobson, 1890. See Benson and Bond 

(1939:348) who considered leucogenys a subspecies of 

S. merriami.

Holotype: USNM 157952. [Skin and skull]. Adult female. Collected 12 Aug 1908 by W. H. Osgood. Original number [SL: 

3318].

Type Locality: Mouth of the canyon of Beaver River, about 3 mi 

(5 km) east of Beaver [FC: and 1 mi (1.6 km) east of Fort 

Cameron, Beaver River], Beaver County, Utah. 

Sorex longicauda elassodon Osgood

N. Am. Fauna 21: 35, 26 Sep 1901b.

= Sorex monticola elassodon Osgood, 1901. See Hennings and 

Hoffmann (1977:16).

Holotype: USNM 100597. [Skin and skull]. Young adult male. 

Collected 13 Jun 1900 by W. H. Osgood and E. Heller. 

Original number [FC: WHO] 1030.

Type Locality: Cumshewa Inlet, [Poole and Schantz (1942:87): 

near old Indian Village of Clew], Moresby Island, Queen 

Charlotte Islands, British Columbia, [Canada]. 

Sorex longicauda prevostensis Osgood

N. Am. Fauna 21: 35, 26 Sep 1901b.

= Sorex monticola prevostensis Osgood, 1901. See Hennings and 

Hoffmann (1977:16).

Holotype: USNM 100618. [Skin and skull]. Adult male. Collected 3 Jul 1900 by W. H. Osgood and E. Heller. Original 

number [FC: WHO] 1089.

Type Locality: [FC: North end of] Prevost Island, [FC: on coast 

of Houston Stewart Channel], Queen Charlotte Islands, 

British Columbia, [Canada]. 

Sorex macrodon Merriam

N. Am. Fauna 10: 82, 31 Dec 1895b.

Holotype: USNM 58272. [Skin and skull]. Adult male. Collected 

26 Jan 1894 by E. W. Nelson and E. A. Goldman. Original 

number 5759.

Type Locality: Orizaba, altitude 4,200 ft (1,280 m), Vera Cruz 

(Veracruz), Mexico. 

Sorex macropygmaeus G. S. Miller

Proc. Biol. Soc. Wash. 14: 158, 9 Aug 1901e.

= Sorex caecutiens Laxmann, 1785. See Hoffmann (1987:107).

Holotype: USNM 84021. Skin and skull. Adult male. Collected 

23 Sep 1897 by Mrs. L. Stejneger. Original number LS 

8019.

Type Locality: Petropaulski (Petropavlovsk Kamchatskiy), Kamchatka (Kamchatskiy Kray), [Russia]. 

Sorex merriami Dobson

Monogr. Insectivora, pt. 3, fasc. 1, pl. 23, fig. 6, and explanation, 

May 1890.

Holotype: USNM 186441. [In alcohol, skull removed]. Adult female. Collected [SL: 26 Dec 1884 by C. E. Bendire]. Original number [MC and MCC: 635].

Type Locality: [Merriam (1895b:88): On the Little Bighorn 

River, about 1.5 mi (2.4 km) above] Fort Custer, [Big Horn 

County], Montana.

Remarks: Published as MCC 1001 to which skull number 4861 

belonged. Cataloged as USNM 186441 on 13 Oct 1911. 

Sorex milleri Jackson

Proc. Biol. Soc. Wash. 60: 131, 9 Oct 1947.

Holotype: USNM 274950. Skin, skull, and dry body. Adult female? Collected 1 Nov 1940 by F. W. Miller. Original number 19. Biological Survey miscellaneous 32327-X.

Type Locality: Madera Camp, altitude 8,000 ft (2,438 m), Carmen Mountains, Coahuila, Mexico.

Remarks: [SL: Donated by the Dallas Museum of Natural 

History]. 

Sorex montereyensis Merriam

N. Am. Fauna 10: 79, 31 Dec 1895b.

= Sorex trowbridgii montereyensis Merriam. See Jackson 

(1922:264).

Holotype: USNM 32000/44810. Skin and skull. Adult male. 

Collected 1 Oct 1891 by V. Bailey. Original number 3336.

Type Locality: Monterey, [Monterey County], California. 

Sorex monticolus Merriam

N. Am. Fauna 3: 43, 11 Sep 1890a.

= Sorex monticola monticola Merriam.

Holotype: USNM 17599/24535. Skin and skull. Adult male. 

Collected 28 Aug 1889 by C. H. Merriam and V. Bailey. 

Original number [SL: VB] 406.

Type Locality: [FC: Near Spring Creek], altitude 3,500 m = 

11,500 ft, San Francisco Mountain, [Coconino County], 

Arizona. 
NUMBER 644 • 4 5

Sorex navigator alaskanus Merriam

Proc. Wash. Acad. Sci. 2: 18, 14 Mar 1900.

= Sorex alaskanus Merriam. See Jackson (1926:58).

Holotype: USNM 97713. [Skin and skull]. Adult male. Collected 

12 Jun 1899 by A. K. Fisher. Original number 2058.

Type Locality: Point Gustavus, Glacier Bay, Alaska. 

Sorex nevadensis Merriam

N. Am. Fauna 10: 71, 31 Dec 1895b.

= Sorex vagrans vagrans Baird, 1857. See Findley (1955:52).

Holotype: USNM 24891/32302. Skin and skull. Adult male. 

Collected 20 Nov 1890, not 24 as in original description, by 

V. Bailey. Original number 2150.

Type Locality: Reese River, [Miller and Kellogg (1955:20): at 

about 6,000 ft (1,829 m)], [FC: at a line between Lander 

and Nye Counties], Nevada. 

Sorex obscurus alascensis Merriam

N. Am. Fauna 10: 71, 31 Dec 1895b.

= Sorex monticola alascensis Merriam, 1895. See Hennings and 

Hoffmann (1977:15).

Holotype: USNM 73539. [Skin and skull]. Young adult female. 

Collected 10 Jul 1895 by C. P. Streator. Original number 

[SL: 4676].

Type Locality: Yakutat Bay, Alaska. 

Sorex obscurus isolatus Jackson

J. Wash. Acad. Sci. 12: 263, 4 Jun 1922.

= Sorex monticola isolatus Jackson, 1922. See Hennings and 

Hoffmann (1977:16).

Holotype: USNM 177719. Skin and skull. Adult male. Collected 

21 May 1911 by F. A. Wetmore. Original number 517.

Type Locality: Mouth of Millstone Creek, Nanaimo, Vancouver 

Island, British Columbia, [Canada]. 

Sorex obscurus longicauda Merriam

N. Am. Fauna 10: 74, 31 Dec 1895b.

= Sorex monticola longicaudus Merriam, 1895. See Hennings 

and Hoffmann (1977:16).

Holotype: USNM 74711. [Skin and skull]. Young adult male. Collected 9 Sep 1895 by C. P. Streator. Original number 4891.

Type Locality: Wrangel (Wrangell), [FC: Wrangel (Wrangell) Island], southeast Alaska. 

Sorex obscurus neomexicanus Bailey

Proc. Biol. Soc. Wash. 26: 133, 21 May 1913.

= Sorex neomexicanus Bailey, 1913. See Alexander (1996:21).

Holotype: USNM 100440. [Skin and skull]. Old adult male. Collected 29 May 1900 by V. Bailey. Original number 7383.

Type Locality: Cloudcroft, altitude 9,000 ft (2,743 m) in the Sacramento Mountains, [Otero County], New Mexico. 

Sorex obscurus parvidens Jackson

J. Mammal. 2: 161, 19 Aug 1921.

= Sorex ornatus parvidens Jackson, 1921. See Woodman 

(2012:837).

Holotype: USNM 56561. Skin and skull. Adult male. Collected 

3 Oct 1893 by J. E. McLellan. Original number 242.

Type Locality: Spring known as Thurman’s Camp, altitude about 

7,500 ft (2,286 m) [FC: 9,000 ft (2,743 m)], western side 

of San Bernardino Peak, San Bernardino Mountains, [San 

Bernardino County], California. 

Sorex obscurus permiliensis Jackson

Proc. Biol. Soc. Wash. 31: 128, 29 Nov 1918.

= Sorex bairdi permiliensis Jackson, 1918. See Carraway 

(1990:40).

Holotype: USNM 91048. Skin and skull. Adult male. Collected 

2 Oct 1897 by J. A. Loring. Original number 4756.

Type Locality: Permilia (Pamelia) Lake, west base of Mount Jefferson, Cascade Range, [Linn County], Oregon. 

Sorex obscurus ventralis Merriam

N. Am. Fauna 10: 75, 31 Dec 1895b.

= Sorex ventralis Merriam, 1895. See Elliot (1903:148).

Holotype: USNM 68342. [Skin and skull]. Adult male. Collected 

26 Aug 1894 by E. W. Nelson and E. A. Goldman. Original 

number 6636.

Type Locality: Cerro San Felipe, altitude 10,000 ft (3,048 m), 

Oaxaca, Mexico. 

Sorex oreopolus Merriam

Proc. Biol. Soc. Wash. 7: 173, 29 Sep 1892.

Holotype: USNM 33663/45698. Skin and skull. Adult male. 

Collected 22 Apr 1892 by E. W. Nelson. Original number 

2517.

Type Locality: [Miller (1912b:17): North Slope] Sierra Nevada 

de Colima, altitude [SL: about] 10,000 ft (3,048 m), Jalisco, 

Mexico. 

Sorex orizabae Merriam

N. Am. Fauna 10: 71, 31 Dec 1895b.

Holotype: USNM 53633. [Skin and skull]. Adult female. Collected 24 Apr 1893 by E. W. Nelson. Original number 4733.

Type Locality: [Jackson (1928:113): West slope] Mount Orizaba, altitude 9,500 ft (2,896 m), Puebla, Mexico. 

Sorex ornatus Merriam

N. Am. Fauna 10: 79, 31 Dec 1895b.

= Sorex ornatus ornatus Merriam, 1895.

Holotype: USNM 31333/43198 Skin and skull. Adult male. Collected 19 Oct 1891 by E. W. Nelson. Original number 1328.

Type Locality: Head of San Emigdio Canyon, Mount Pinos, 

[Kern County], California. 

Sorex pachyurus Baird

Mammals Rep. Explor. Surv. 8(1), 1857.

= Sorex arcticus arcticus Kerr, 1792. See Hutterer (2005:284).
4 6 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Lectotype: USNM 1674/38820 as designated by Lyon and Osgood (1909:247). Originally in alcohol, the skull and skeleton were removed and cataloged as USNM 38820 on 13 

Sep 1923. The skin was apparently destroyed. Age and sex 

not given. Obtained from C. Cavileer. No original number.

Type Locality: Pembina, [Pembina County], North Dakota, not 

Minnesota as in original description.

Remarks: A type was not designated by number in original description. Lyon and Osgood (1909:247) considered USNM 

1674 to be the type = lectotype (ICZN, 1999: Article 74.5). 

Sorex pacificus Coues

Bull. U.S. Geogr. Surv. Terr. 3: 650, 15 May 1877.

= Sorex pacificus pacificus Coues, 1877. See Carraway (1990:54).

Holotype: USNM 3266. [Skin with fragment of skull inside]. 

Age and sex not given. [MC: Received from E. P. Vollum. 

Cataloged Mar 1858].

Type Locality: Fort Umpqua, [Miller and Rehn (1901:242): 

mouth of Umpqua River, Douglas County], Oregon. 

Sorex pacificus cascadensis Carraway

Special Publs. Mus. Texas Tech Univ. 32: 55, 7 Jul 1990.

Holotype: USNM 204479. Skin and skull. Adult female. Collected 5 Jul 1914 by V. Bailey. Original number 9650.

Type Locality: McKenzie Bridge, Lane County, Oregon. 

Sorex palustris hydrobadistes Jackson

J. Mammal. 7: 57, 15 Feb 1926.

Holotype: USNM 229061. Skin and skull. Adult male. Collected 

23 Jul 1918 by H. H. T. Jackson. Original number 1211.

Type Locality: Withee, Clark County, Wisconsin. 

Sorex palustris turneri Johnson

Proc. Biol. Soc. Wash. 64: 110, 24 Aug 1951.

= Sorex albibarbis turneri Johnson, 1951. See Hope et al. 

(2014:730) for current use.

Holotype: USNM 282879. Skin and skull. Adult male. Collected 

by R. Traub and D. Koelle; prepared by D. H. Johnson. 

Original number DHJ 5216.

Type Locality: Fort Chimo (Kuujjuaq), on eastern bank of Koksoak River, lat. 58o8 ́N, long. 68o15 ́W, Ungava District, 

Quebec, Canada. 

Sorex personatus arcticus Merriam

Proc. Wash. Acad. Sci. 2: 17, 14 Mar 1900. Primary junior homonym of Sorex arcticus Kerr, 1792. Replaced with Sorex cinereus hollisteri by Jackson (1925c:55).

Holotype: USNM 99305. [Skin and skull]. Adult female. Collected 14 Sep 1899 by W. H. Osgood. Original number 910.

Type Locality: Saint Michaels (Saint Michael), Alaska. 

Sorex personatus streatori Merriam 

N. Am. Fauna 10: 62, 31 Dec 1895b.

= Sorex cinereus streatori Merriam, 1895. See Jackson 

(1925c:56).

Holotype: USNM 73537. [Skin and skull]. Adult male. Collected 

9 Jul 1895 by C. P. Streator. Original numbr 4674.

Type Locality: Yakutat, about lat. 59o35 ́, Alaska. 

Sorex planiceps G. S. Miller

Proc. Biol. Soc. Wash. 24: 242, 28 Nov 1911c.

Holotype: USNM 173915. Skin and skull. Adult male. Collected 

30 May 1911 by W. L. Abbott. Original number 7714.

Type Locality: [FC: Kiar Nye], Dachin, Altitude 9,000 ft (2,743 

m), Khistwar, Kashmir (Jammu and Kashmir), [India]. 

Sorex preblei Jackson

J. Wash. Acad. Sci. 12: 263, 4 Jun 1922.

Holotype: USNM 208032. Skin and skull. Adult male. Collected 

3 Jul 1915 by E. A. Preble. Original number 5972.

Type Locality: Jordan Valley, altitude 4,200 ft (1,280 m), Malheur County, Oregon. 

Sorex pribilofensis Merriam

N. Am. Fauna 10: 87, 31 Dec 1895b.

Holotype: USNM 30911. In alcohol. Adult female. Collected 29 

Jul 1891 by C. H. Merriam. No original number.

Type Locality: Saint Paul Island, Pribilof Islands, Bering Sea, 

Alaska. 

Sorex roboratus Hollister

Smithson. Misc. Collect. 60(24): 2, 13 Mar 1913b.

Holotype: USNM 175436. Skin and skull. Adult male. Collected 

7 Aug 1912 by N. Hollister. Original number 4451.

Type Locality: Dapucha, Altai Mountains, [Altayskiy], Siberia 

(Russia). 

Sorex salvini Merriam

Proc. Biol. Soc. Wash. 112: 229, 15 Jul 1897c.

= Sorex salvini salvini Merriam, 1897. See Woodman et al. 

(2012:214) for current use.

Holotype: USNM 77035. [Skin and skull]. Adult female. Collected 12 Jan 1896 by E. W. Nelson and E. A. Goldman. 

Original number 9057.

Type Locality: Calel, altitude 10,200 ft (3,109 m), Tontonicapan 

(Quetzaltenango), Guatemala. 

Sorex saussurei Merriam

Proc. Biol. Soc. Wash. 7: 173, 29 Sep 1892.

See Carraway (2007:73) for current use.

Holotype: USNM 33667/45702. Skin and skull. Adult female. 

Collected 23 Apr 1892 by E. W. Nelson. Original number 

2538.

Type Locality: [Miller (1912b:19): North slope] Sierra Nevada 

de Colima, altitude [FC: about] 8,000 ft (2,438 m), Jalisco, 

Mexico. 

Sorex saussurei caudatus Merriam

N. Am. Fauna 10: 84, 31 Dec 1895b. Primary junior homonym of Sorex caudatus Hodgson, 1849 and Horsfield, 
NUMBER 644 • 4 7

1851. Replaced with Sorex saussurei mutablis by Merriam 

(1898b:782).

= Sorex veraepacis mutabilis Merriam, 1898. See Jackson 

(1925b:130).

Holotype: USNM 69600. [Skin and skull]. Young adult female. 

Collected 21 Oct 1894 by E. W. Nelson [SL and FC: and 

E. A. Goldman]. Original number 6963.

Type Locality: Reyes, [Poole and Schantz (1942:193): near Cuicatlan], altitude 10,200 ft (3,109 m), Oaxaca, Mexico. 

Sorex saussurei cristobalensis Jackson

Proc. Biol. Soc. Wash. 38: 129, 13 Nov 1925b.

= Sorex salvini cristobalensis Jackson, 1925. See Woodman et al. 

(2012:214) for current use.

Holotype: USNM 75883. Skin and skull. Adult female. Collected 19 Sep 1895 by E. W. Nelson and E. A. Goldman. 

Original number 8429.

Type Locality: San Cristobal, altitude 8,400 ft (2,560 m), Chiapas, Mexico. 

Sorex saussurei oaxacae Jackson

Proc. Biol. Soc. Wash. 38: 128, 13 Nov 1925b.

= Sorex salvini oaxacae Jackson, 1925. See Woodman et al. 

(2012:214) for current use.

Holotype: USNM 71467. Skin and skull. Adult female. Collected 27 Mar 1895 by E. W. Nelson and E. A. Goldman. 

Original number 7748.

Type Locality: Mountains near Ozolotepec, altitude 10,000 ft 

(3,048 m), Oaxaca, Mexico. 

Sorex saussurei veraecrucis Jackson

Proc. Biol. Soc. Wash. 38: 128, 13 Nov 1925b.

= Sorex salvini veraecrucis Jackson, 1925. See Woodman et al. 

(2012:214) for current use.

Holotype: USNM 55106. Skin and skull. Adult female. Collected 14 Jul 1893 by E. W. Nelson. Original number 5235.

Type Locality: Xico, altitude 6,000 ft (1,829 m), Vera Cruz 

(Veracruz), Mexico. 

Sorex sclateri Merriam

Proc. Biol. Soc. Wash. 11: 228, 15 Jul 1897c.

Holotype: USNM 75872. [Skin and skull]. Adult female. Collected 23 Oct 1895 by E. W. Nelson and E. A. Goldman. 

Original number 8567.

Type Locality: Tumbala, altitude 5,000 ft (1,524 m), [Tumbala 

Municipality], Chiapas, Mexico. 

Sorex shastensis Merriam

N. Am. Fauna 16: 87, 28 Oct 1899.

=Sorex vagrans vagrans Baird, 1857. See Findley (1955:52).

Holotype: USNM 95450. [Skin and skull]. [Lyon and Osgood 

(1909:248): Adult] [SL: female]. Collected 26 Sep 1898 by 

W. H. Osgood. Original number 317.

Type Locality: Wagon Camp, Mount Shasta, altitude 5,700 ft 

(1,737 m), [Siskiyou County, California]. 

Sorex sonomae tenelliodus Carraway

Special Publs. Mus. Texas Tech Univ. 32: 33, 7 Jul 1990.

Holotype: USNM 565663. Skin and skull. Adult female. Collected 

13 May 1983 by R. G. Anthony. Original number X-560. 

Oregon State University Fish and Wildlife number 8251.

Type Locality: 4 mi (6.4 km) S, 10.5 mi (17 km) E Blue River, 

East Fork Creek, T17S, R6E, sec. 7, Lane County, Oregon.

Remarks: Transferred to the NMNH on 2 May 1989. 

Sorex sphagnicola Coues

Bull. U.S. Geol. and Geogr. Surv. Terr. 3: 650, 15 May 1877.

= Sorex arcticus arcticus Kerr, 1792. See Jackson (1928:69).

Holotype: USNM 6361. Skin only. Age and sex not given. Collected on unknown date by W. L. Hardisty. No original 

number. [MC: Cataloged in Apr 1863].

Type Locality: Vicinity of Fort Liard, [Dehcho Region, Northwest Territories, Canada]. 

Sorex stizodon Merriam

N. Am. Fauna 10: 98, 31 Dec 1895b.

Holotype: USNM 75885. [Skin and skull]. Adult female. Collected 25 Sep 1895 by E. W. Nelson and E. A. Goldman. 

Original number 8473.

Type Locality: San Cristobal, [FC: altitude 9,000 ft (2,743 m)], 

Chiapas, Mexico. 

Sorex suckleyi Baird

Mammals Rep. Explor. Surv. 8(1): 18, 1857.

= Sorex vagrans vagrans Baird, 1857. See Merriam (1895b:67).

Lectotype: USNM 1677 as designated by Lyon and Osgood 

(1909:250). In alcohol. Age and sex not given. Collected by 

G. Suckley in [MC: 1856. Cataloged on 23 Oct 1856]. No 

original number.

Type Locality: Fort Steilacoom, Puget Sound, [Pierce County], 

Washington.

Remarks: Of the seven specimens mentioned in the original description, Lyon and Osgood (1909:250) stated that USNM 

1677 is here taken as the type = lectotype (ICZN, 1999: 

Article 74.5). 

Sorex tenellus Merriam

N. Am. Fauna 10: 81, 31 Dec 1895b.

Holotype: USNM 25083/32495. Skin and skull. Adult, sex unknown. Collected 22 Dec 1890 by E. W. Nelson. Original 

number 131.

Type Locality: Near summit of Alabama Hills, Lone Pine, Owens 

Valley, [Inyo County], California. Restricted by A. B. Howell 

(1923:266) to Lone Pine Creek, about 4,500 ft (1,372 m), 

Owens Valley, Inyo County, California. 

Sorex tenellus lyelli Merriam

Proc. Biol. Soc. Wash. 15: 75, 22 Mar 1902b.

= Sorex lyelli Merriam, 1902. See Jackson (1928:57).

Holotype: USNM 109530. [Skin and skull]. Young adult male. Collected 29 Aug 1901 by W. K. Fisher. Original number 2275.
4 8 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Type Locality: Mount Lyell, [Grinnell (1933:79): near head of 

Lyell Fork of Tuolumne River, at about 10,000 ft (3,048 m) 

altitude], [Tuolumne County], California. 

Sorex tenellus myops Merriam

Proc. Biol. Soc. Wash. 15: 76, 22 Mar 1902b.

= Sorex tenellus Merriam, 1895. See Hall and Kelson (1959:36).

Holotype: USNM 29559/41634. Skin and skull. Adult female. Collected 13 Jul 1891 by E. W. Nelson. Original number 1101.

Type Locality: [FC: Pipers Creek (Cottonwood Creek), east 

slope near main peak of] White Mountains, [FC: 9488 ft 

(2,892 m)], Mono County, California. Restricted by Howell 

(1923:266) to Cottonwood Creek, about 9,550 ft (2,911 m), 

on east slope of the White Mountains, Inyo (Mono) County, 

California. 

Sorex tenellus nanus Merriam

N. Am. Fauna 10: 81, 31 Dec 1895b.

= Sorex nanus Merriam, 1895. See Jackson (1928:174).

Holotype: USNM 73773. [Skin and skull]. [Poole and Schantz 

(1942:196): Young] adult female. Collected 3 Aug 1895 by 

E. A. Preble. Original number [SL: 567].

Type Locality: Estes Park, [FC: Larimer County], Colorado. 

Sorex thompsoni Baird

Mammals Rep. Explor. Surv. 8(1): 34, 1857.

= Sorex hoyi thompsoni Baird, 1857. See Diersing (1980:94).

Lectotype: USNM 1686/38838 as designated by Lyon and Osgood (1909:250). In alcohol, [skull removed]. Age and sex 

not given. Obtained from Z. Thompson. No original number. 

Date collected unknown. [MC: Cataloged 23 Oct 1856].

Type Locality: Burlington, [Chittenden County], Vermont.

Remarks: A type was not designated by number in original description. From the three specimens mentioned, Lyon and 

Osgood (1909:250) condidered USNM 1686 to be the type 

= lectotype (ICZN, 1999: Article 74.5). The skull, still in 

the specimen in 1909, has been removed and was assigned 

USNM 38838 on 13 Jun 1923. 

Sorex trigonirostris Jackson

J. Wash. Acad. Sci. 12: 264, 4 Jun 1922.

= Sorex vagrans vagrans Baird, 1857. See Hennings and Hoffmann (1977:8).

Holotype: USNM 203608. Skin and skull. Adult female. Collected 

5 May 1914 by L. J. Goldman. Original number 1308.

Type Locality: Ashland, altitude 1,975 ft (602 m), Jackson 

County, Oregon. 

Sorex trowbridgii Baird

Mammals Rep. Explor. Surv. 8(1): 13, 1857.

= Sorex trowbridgii trowbridgii Baird, 1857.

Lectotype: USNM 813/3088 as designated by Poole and Schantz 

(1942:196). Skin and skull. Male. Collected on unknown 

date by J. Wayne. No original number. [MC: Skin cataloged 

on 28 Jul 1855, skull on Jan 1857].

Type Locality: Astoria, [Miller and Rehn (1901:239): mouth of 

Columbia River, Clatsop County], Oregon.

Remarks: A type was not designated by number in original description. Baird (1857:15) listed four specimens by number and Poole and Schantz (1942:196) selected USNM 

813/3088 as the lectotype. 

Sorex trowbridgii destructioni Scheffer and Dalquest

J. Mammal. 23: 334, 14 Aug 1942.

Holotype: USNM 271634. Skin and skull. Adult female. Collected 22 Apr 1941 by V. B. Scheffer. Original number 1151.

Type Locality: Destruction Island, Jefferson County, Washington. 

Sorex trowbridgii humboldtensis Jackson

J. Wash. Acad. Sci. 12: 264, 4 Jun 1922.

Holotype: USNM 97271. Skin and skull. Adult male. Collected 

11 Jun 1899 by W. K. Fisher. Original number 914. 

Type Locality: [SL: Redwood] Carson’s Camp, Mad River, Humboldt Bay, Humboldt County, California. 

Sorex tundrensis Merriam

Proc. Wash. Acad. Sci. 2: 16, 14 Mar 1900.

Holotype: USNM 99286. [Skin and skull]. [Lyon and Osgood 

(1909:250): Adult]. Sex not given. Collected 13 Sep 1899 

by W. H. Osgood. Original number 902.

Type Locality: Saint Michaels (Saint Michael), Alaska. 

Sorex vagrans Baird

Mammals Rep. Explor. Surv. 8(1): 15, 1857.

= Sorex vagrans vagrans Baird, 1857.

Lectotype: USNM 1675 as designated by Lyon and Osgood 

(1909:251). In alcohol. [Lyon and Osgood (1909:251): 

Adult] male. Received from J. G. Cooper, date unknown. 

No original number. [MC: Cataloged 23 Oct 1856].

Type Locality: Shoalwater (Willapa) Bay, [Pacific County], W. T. 

(Washington).

Remarks: A type was not designated in original description. 

Of the seven specimens mentioned, Lyon and Osgood 

(1909:251) selected USNM 1675 as the type = lectotype 

(ICZN, 1999: Article 74.5). 

Sorex vagrans obscuroides Findley

Univ. Kans. Publ., Mus. Nat. Hist. 9: 58, 10 Dec 1955.

= Sorex monticola obscurus Merriam, 1895. See Hennings and 

Hoffmann (1977:15).

Holotype: USNM 30064/42074. Skin and skull. First year female. Collected 9 Aug 1891 by F. Stephens. Original number 811.

Type Locality: Bishop Creek, 6600 ft (2,012 m), [SL and FC: 

Sierra Nevada Mountains], Inyo County, California. 

Sorex vagrans similis Merriam

N. Am. Fauna 5: 34, 30 Jul 1891. Primary junior homonym of 

Sorex similis Hensel, 1855. Renamed Sorex obscurus by 

Merriam (1895b:72).
NUMBER 644 • 4 9

= Sorex monticola obscurus Merriam, 1895. See Hennings and 

Hoffmann (1977:17).

Holotype: USNM 23525/30943. Skin and skull. [Lyon and Osgood (1909:249): Adult] female. Collected 26 Aug 1890 by 

[SL: V. Bailey] and B. H. Dutcher. Original number [FC: VB] 

1670.

Type Locality: [Poole and Schantz (1942:197): Near] Timber 

Creek, altitude about 2,500 m (8,300 ft), Salmon River 

(Lemhi) Mountains, 10 mi (16 km) west of Junction, [Lemhi 

County], Idaho. 

Sorex vancouverensis Merriam

N. Am. Fauna 10: 70, 31 Dec 1895b.

= Sorex vagrans vagrans Baird, 1857. See George and Smith 

(1991:88).

Holotype: USNM 71913. [Skin and skull]. Adult male. Collected 

10 May 1895 by C. P. Streator. Original number 4592.

Type Locality: Goldstream, Vancouver Island, British Columbia, 

[Canada]. 

Sorex veraepacis chiapensis Jackson

Proc. Biol. Soc. Wash. 38: 129, 13 Nov 1925b.

Holotype: USNM 75877. Skin and skull. Adult female. Collected 1 Oct 1895 by E. W. Nelson and E. A. Goldman. 

Original number 8528.

Type Locality: San Cristobal, altitude 9,500 ft (2,896 m), Chiapas, Mexico. 

Sorex yaquinae Jackson

Proc. Biol. Soc. Wash. 31: 127, 29 Nov 1918.

= Sorex pacificus pacificus Coues, 1877. See Carraway (1990:54).

Holotype: USNM 73051. Skin and skull. Adult female. Collected 

18 Jul 1895 by B. J. Bretherton. Original number 2031.

Type Locality: Yaquina Bay, [Lincoln County], Oregon. 

Genus Suncus Ehrenberg, 1832

Pachyura occultidens Hollister

Proc. U.S. Natl. Mus. 46: 303, 31 Dec 1913a.

= Suncus murinus (Linnaeus, 1766). See Corbet and Hill 

(1992:36).

Holotype: USNM 175761. In alcohol, skull removed. Young 

adult male. Collected Aug [MC: Sep] 1912 by [MC: C. 

Fox]. No original number.

Type Locality: Ilo-ilo (Iloilo), Iloilo Province, Panay, [Philippines]. 

Genus Surdisorex Thomas, 1906

Surdisorex polulus Hollister

Smithson. Misc. Collect. 66(1): 1, 10 Feb 1916c.

Holotype: USNM 163992. Skin and skull. Adult male. Collected 

30 Sep 1909 by J. A. Loring. Original number 7562.

Type Locality: West side Mount Kenia (Kenya), altitude 10,700 

ft (3,261 m), British East Africa (Kenya). 

Genus Sylvisorex Thomas, 1904

Pachyura lixa aequatoria Heller

Smithson. Misc. Collect. 60(12): 4, 4 Nov 1912a.

= Sylvisorex aequatorius (Heller, 1912). New name combination. 

See Hutterer (2005:257) who raised aequatorius to specific 

status and Dubey et al. (2008:953) for the use of Sylvisorex.

Holotype: USNM 181814. [Skin and skull]. Adult male. Collected 19 Nov 1911 by E. Heller. Original number 4815.

Type Locality: Summit of Mount Sagalla, altitude 4,000 ft 

(1,219 m), Taita Hills, [Coast Province], British East Africa 

(Kenya). 

Sylvisorex gemmeus Heller

Smithson. Misc. Collect. 56(15):7, 23 Dec 1910d.

= Sylvisorex megalura gemmeus Heller, 1910. See Heim de Balsac and Meester (1977:7) who reduced gemmeus to subspecific status and Dubey et al. (2008:960) for current use of 

Sylvisorex.

Holotype: USNM 164644. [Skin and skull]. Adult male. Collected 14 Jan 1910 by J. A. Loring. Original number 8730.

Type Locality: Rhino Camp, Lado Enclave, [Arua, Uganda]. 

Sylvisorex konganensis Ray and Hutterer

Ecotropica 1: 93, 1996 [1995].

Holotype: USNM 580655. In alcohol, skull removed. Young 

adult female. Collected 8 Apr 1994 by J. C. Ray. Original 

number 1736.

Type Locality: Near Kongana Camp, 02o47 ́N, 16o25 ́E, DzangaSangha Forest Reserve, SW Central African Republic. 

Family Talpidae G. Fischer, 1814

Genus Condylura Illiger, 1811

Condylura cristata parva Paradiso

Proc. Biol. Soc. Wash. 72: 103, 24 Jul 1959.

Holotype: USNM 293291. Skin and skull. Adult male. Collected 

2 Dec 1939 by L. Llewellyn. Original number [SL: 565].

Type Locality: [SL: Spoon Mountain], 5 mi (8 km) NW of Stewart, Patrick County, Virginia.

Remarks: Hutterer (2005:301) placed C. c. parva as a junior 

synonym of C. c. nigra following Hartman (1999:67), who 

placed nigra as a synonym of parva, despite nigra having 

priority. However, we believe Hartman (1999:67) erred in 

doing so and meant to place nigra as a synonym of cristata,

as he correctly listed the range of cristata to include the type 

locality, Nova Scotia, of nigra. In our opinion, Peterson and 

Yates (1980:1) correctly placed nigra as a junior synonym of

cristata as did van Zyll de Jong (1983:180). 

Genus Dymecodon True, 1886

Dymecodon pilirostris True

Proc. U.S. Natl. Mus. 9: 97, 2 Sep 1886a.
5 0 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Holotype: USNM 15291/22139. In alcohol, [skull removed]. 

[Lyon and Osgood (1909:234): Immature] [MC: male]. 

[MC: Collected on unknown date by E. S. Morse. No original number. Received from the Boston Society of Natural 

History 19 Feb 1878. Cataloged 21 Dec 1885].

Type Locality: Mouth of Yeddo (Tokyo) Bay, [MC: Yenosima 

(Enoshima)], [Honshu], Japan. 

Genus Euroscaptor Miller, 1940

Euroscaptor grandis G. S. Miller

J. Mammal. 21: 444, 14 Nov 1940a.

Holotype: USNM 255370. Skin and skull. Adult female. Collected Jan 1930 by D. C. Graham. Original number 370.

Type Locality: Mount Omei (Emei Shan), altitude 5,000 ft (1,524 

m), Szechuan (Sichuan), China. 

Talpa parvidens G. S. Miller

J. Mammal. 21: 203, 14 May 1940b.

= Euroscaptor parvidens (G. S. Miller, 1940). See G. S. Miller 

(1940a:444).

Holotype: USNM 258342. In alcohol, skull removed. Adult 

male, not female as given in the original description (G. S. 

Miller, 1940a:444). Collected 13 Jun 1933 by E. Poilane. 

No original number.

Type Locality: Agricultural station of Blao (Bao Loc), [Note 

in type file: altitude 800 m], Delegation de Djynrinh, near 

upper Donai river, Annam (Lam Dong), French Indo-China 

(Vietnam). 

Genus Neurotrichus Gunther, 1880

Neurotrichus gibbsi major Merriam

N. Am. Fauna 16: 88, 28 Oct 1899.

= Neurotrichus gibbsii gibbsii (Baird, 1857). See G. S. Miller 

(1924:16).

Holotype: USNM 65321. Skin and skull. Adult male. Collected 

18 May 1894 by C. P. Streator. Original number 3789.

Type Locality: Carberry Ranch, altitude 4,100 ft (1,250 m), [FC: 

4,050 ft (1,234 m)], between Mounts Shasta and Lassen, 

[FC: 50 mi (80 km) NE of Redding on stage road to Alturus 

(Alturas)], Shasta County, California. 

Urotrichus gibbsii Baird

Mammals Rep. Explor. Surv. 8(1): 76, 1857.

= Neurotrichus gibbsii gibbsii (Baird, 1857). See Gunther 

(1880:pl. 42).

Holotype: USNM 662/1843. Skin and skull. Immature of unknown sex. Collected [MC: 15] Jul 1854 by G. Gibbs. Original number 15.

Type Locality: White River, Cascade Mountains, W. T. (Washington). Restricted by Dalquest and Burgner (1941:12) 

to Naches Pass, 4,500 ft (1,372 m), Pierce County, 

Washington. 

Genus Scalopus E. Geoffroy, 1803

Scalopus aquaticus caryi Jackson

Proc. Biol. Soc. Wash. 27: 20, 2 Feb 1914a.

Holotype: USNM 116799. Skin and skull. Young adult male. 

Collected 18 Sep 1901 by M. Cary. Original number 307. 

[SL: 3640-X].

Type Locality: Neligh, [Antelope County], Nebraska. 

Scalopus aquaticus howelli Jackson

Proc. Biol. Soc. Wash. 27: 19, 2 Feb 1914a.

Holotype: USNM 177931. Skin and skull. Adult male. Collected 4 Jan 1912 by L. S. Golsan. No original number. 

[SL: 8877-X].

Type Locality: Autaugaville, [Autauga County], Alabama. 

Scalopus aquaticus machrinoides Jackson

Proc. Biol. Soc. Wash. 27: 19, 2 Feb 1914a.

Holotype: USNM 169717. Skin and skull. Adult male. Collected 

1 Jun 1910 by W. E. Berg. Original number [XC: 16]. [SL: 

8139-X].

Type Locality: Manhattan, [Riley County], Kansas. 

Scalopus aquaticus pulcher Jackson

Proc. Biol. Soc. Wash. 27: 20, 2 Feb 1914a.

= Scalopus aquaticus aereus (Bangs, 1896). See Yates and 

Schmidly (1977:25).

Holotype: USNM 170698. Skin and skull. Adult male. Collected 

20 Jan 1911 by W. G. Savage. Original number [XC: 58]. 

[SL: 8316-X].

Type Locality: Delight, [Pike County], Arkansas. 

Scalopus inflatus Jackson

Proc. Biol. Soc. Wash. 27: 21, 2 Feb 1914a.

= Scalopus aquaticus inflatus Jackson, 1914. See Yates and 

Schmidly (1977:28).

Holotype: USNM 52709. Skin and skull, Young adult, sex unknown. Collected 1893 by F. B. Armstrong. Original number [SL: 1].

Type Locality: 45 mi (72 km) from Brownsville, Texas [in] Tamaulipas, Mexico. 

Genus Scapanus Pomel, 1848

Scalops aeneus Cassin

Proc. Acad. Nat. Sci. Phila. 6: 299, reported favorably for publication 22 Feb 1853. Receipt acknowledged 13 Apr 1853 by 

the Smithsonian Institution.

= Scapanus townsendii townsendii (Bachman, 1839). See True 

(1896:51).

Holotype: USNM 3725. Skin only. Adult, [sex unknown]. Collected on unknown date by the U.S. Exploring Expedition. 

No original number. [MC: Cataloged on 20 Dec 1859].

Type Locality: Oregon.
NUMBER 644 • 5 1

Remarks: As stated by Lyon and Osgood (1909:233), the original description was based on a single specimen in the collection of the Exploring Expedition. The above listed specimen 

has a U.S. Exploring Expedition label and is undoubtedly 

this “single specimen.” 

Scapanus alpinus Merriam

Proc. Biol. Soc. Wash. 11: 102, 26 Apr 1897d.

= Scapanus latimanus dilatus True, 1894. See Palmer (1937:291).

Holotype: USNM 79967. [Skin and skull]. Adult male. Collected 

18 Aug 1896 by V. Bailey. Original number 5784.

Type Locality: Crater Lake, altitude about 7,000 ft (2,120 m), 

Mount Mazama, [Klamath County], Oregon. 

Scapanus dilatus True

Proc. U.S. Natl. Mus. 17: 2, 24 Apr 1894b. Preprint of Proc. U.S. 

Natl. Mus. 17: 242, 15 Nov 1894c.

= Scapanus latimanus dilatus True, 1894. See Jackson (1915:72).

Holotype: USNM 186628. [Skull and] skeleton. [MCC: Adult]. 

Collected in 1883 by C. E. Bendire. No original number. 

Merriam collection number 1286.

Type Locality: Fort Klamath, [Klamath County], Oregon.

Remarks: Published as Merriam collection number 1286. Cataloged USNM 186628 on 3 Nov 1911. 

Scapanus latimanus sericatus Jackson

Proc. Biol. Soc. Wash. 27: 55, 20 Mar 1914b.

Holotype: USNM 109548. Skin and skull. Adult female. Collected 20 Aug 1901 by W. K. Fisher. Original number 2227.

Type Locality: Yosemite, [G. S. Miller (1924:12): Yosemite Valley], Mariposa County, California. 

Scapanus orarius True

Proc. U.S. Natl. Mus. 19: 52, 21 Dec 1896.

= Scapanus orarius orarius True, 1896.

Holotype: USNM 1381/37434. Skin and skull. Adult female. Collected 30 Aug 1855 by J. G. Cooper. No original number.

Type Locality: Shoalwater (Willapa) Bay, [Pacific County], 

Washington.

Remarks: Published as USNM 1381, to which 37434 belongs. 

Scapanus orarius schefferi Jackson

N. Am. Fauna 38: 63, 30 Sep 1915.

Holotype: USNM 204997. Skin and skull. Adult male. Collected 

8 Aug 1914 by T. H. Scheffer. No original number. [SL: 

11231-X].

Type Locality: Walla Walla, Walla Walla County, Washington. 

Scapanus truei Merriam

Proc. Biol. Soc. Wash. 11: 102, 26 Apr 1897d.

= Scapanus latimanus dilatus True, 1894. See Jackson (1915:72).

Holotype: USNM 79290. [Skin and skull]. Adult, sex not given. Collected 15 June 1896 by C. P. Streator. Original number 5289.

Type Locality: Lake City, Modoc County, California. 

Order Chiroptera Blumenbach, 1779

Family Pteropodidae Gray, 1821

Genus Acerodon Jourdan, 1837

Pteropus arquatus Miller and Hollister

Proc. Biol. Soc. Wash. 34: 100, 30 Jun 1921.

 = Acerodon celebensis (Peters, 1867). See Musser et al. 

(1982:319).

Holotype: USNM 218612. Skin and skull. Adult male. Collected 

5 Jan 1917 by H. C. Raven. Original number 3067.

Type Locality: Koelawi (Kulawi), [Sulawesi Tengah], Middle Celebes (Sulawesi), [Indonesia]. 

Genus Cynopterus F. Cuvier, 1824

Cynopterus angulatus G. S. Miller

Proc. Acad. Nat. Sci. Phila. 50: 316, 25 Jul 1898b.

= Cynopterus sphinx angulatus G. S. Miller, 1898. See Hill and 

Thonglongya (1972:175).

Holotype: USNM 83569. In alcohol, [skull removed]. Adult male. 

Collected in 1896 by W. L. Abbott. No original number.

Type Locality: [MC: Lay Song Hong], Trong (Trang), Lower 

Siam (Thailand). 

Cynopterus babi Lyon

Proc. U.S. Natl. Mus. 52: 438, 30 Dec 1916.

= Cynopterus sphinx babi Lyon, 1916. See Kitchener and Maharadatunkamsi (1991:308).

Holotype: USNM 114269. Skin and skull. [Poole and Schantz 

(1942:137): Adult] [MC: female]. Collected 8 Jan 1902 by 

W. L. Abbott. Original number 1411.

Type Locality: Pulo (Pulau) Babi, [FC: 17 mi (27 km) SE of Simalur Island (Pulau Simeulue)], [Aceh, Sumatra, Indonesia]. 

Cynopterus major G. S. Miller

Proc. Biol. Soc. Wash. 19: 62, 1 May 1906d.

= Cynopterus titthaecheilus major G. S. Miller. See Hill 

(1983:121).

Holotype: USNM 141236. Skin and skull. Adult male. Collected 

10 Mar 1905 by W. L. Abbott. Original number 4021.

Type Locality: Mojeia River, Nias Island (Pulau Nias), [Sumatera 

Utara], west Sumatra, [Indonesia]. 

Cynopterus minutus G. S. Miller

Proc. Biol. Soc. Wash. 19: 63, 1 May 1906d.

Holotype: USNM 141240. Skin and skull. Adult male. Collected 

11 Mar 1905 by W. L. Abbott. Original number 4043.

Type Locality: [FC: Mojeia River], Nias Island (Pulau Nias), [Sumatera Utara], west Sumatra, [Indonesia]. 

Cynopterus pagensis G. S. Miller

Proc. Biol. Soc. Wash. 19: 62, 1 May 1906d.
5 2 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

= Cynopterus sphinx pagensis G. S. Miller, 1906. See Hill 

(1983:118).

Holotype: USNM 121581. Skin and skull. Adult female. Collected 12 Nov 1902 by W. L. Abbott. Original number 2028.

Type Locality: North Pagi Island (Pulau Pagai Utara), [Sumatera 

Barat], west Sumatra, [Indonesia]. 

Cynopterus princeps G. S. Miller

Proc. Biol. Soc. Wash. 19: 61, 1 May 1906d.

= Cynopterus horsfieldii princeps G. S. Miller, 1906. See Corbett 

and Hill (1992:72).

Holotype: USNM 141235. Skin and skull. Adult female. Collected 10 Mar 1905 by W. L. Abbott. Original number 4020.

Type Locality: Mojeia River, Nias Island (Pulau Nias), [Sumatera 

Utara], west Sumatra, [Indonesia]. 

Cynopterus sphinx serasani Paradiso

Proc. Biol. Soc. Wash. 84: 296, 4 Nov 1971.

Holotype: USNM 104744. Skin and skull. Adult male. Collected 

10 Jun 1900 by W. L. Abbott. Original number [SL: 472].

Type Locality: Serasan Island (Pulau Serasan), Natuna Islands 

(Kepulauan Natuna Selatan), [Kepulauan Riau], Indonesia. 

Niadias [sic.] minor Lyon

Proc. U.S. Natl. Mus. 34: 665, 14 Sep 1908. Junior secondary 

homonym of Cynopterus (Cynonycteris) minor Trouessart, 

1878. Replaced with Cynopterus horsfieldii lyoni by Andersen (1912:827).

= Cynopterus horsfieldii harpax Thomas and Wroughton, 1909. 

See Koopman (1994:33).

Holotype: USNM 144264. In alcohol, [skull removed]. Nearly 

adult male. Collected 14 Dec 1906 by W. L. Abbott. Original number 4951.

Type Locality: Confluence of the Gasip (Sungai Gasip) and Siak 

(Sungai Siak) Rivers [FC: Siak River (Sungai Siak) near 

mouth of Gasip River (Sungai Gasip), [Riau], eastern Sumatra, [Indonesia].

Remarks: Niadias is an incorrect subsequent spelling of Niadius

G. S. Miller, 1906. 

Genus Eonycteris Dobson, 1873

Eonycteris robusta G. S. Miller

Proc. Biol. Soc. Wash. 26: 73, 22 Mar 1913d.

Holotype: USNM 175849. In alcohol, [skull removed]. Collected 

5 Jul 1908 by P. Bartsch. No original number.

Type Locality: In cave at Montalban, [Rizal], Luzon, Philippine 

Islands (Philippines). 

Genus Harpyionycteris Thomas, 1896

Harpyionycteris celebensis Miller and Hollister

Proc. Biol. Soc. Wash. 34: 99, 30 Jun 1921.

Holotype: USNM 219349. Skin and skull. Adult female. Collected 23 Aug 1917 by H. C. Raven. Original number 3176.

Type Locality: Gimpoe (Gimpu), [Sulawesi Tengah], Middle Celebes (Sulawesi), [Indonesia]. 

Genus Pteropus Brisson, 1762

Pteropus aldabrensis True

Proc. U.S. Natl. Mus. 16: 1, 14 Jul 1893a. Preprint of Proc. U.S. 

Natl. Mus. 16: 533, 21 Oct 1893b.

Syntypes: USNM 20984/36053 and 20985/36054. Skins and 

skulls. Adult males. Collected 26 Sep and 5 Oct 1892 respectively by W. L. Abbott. No original numbers.

Type Locality: Aldabra Island, [SL: northwest of Madagascar], 

[Seychelles].

Remarks: Type not designated in original description but the two 

specimens given above were listed by number and therefore 

are syntypes (ICZN, 1999: Article 73.2). 

Pteropus balutus Hollister

Proc. Biol. Soc. Wash. 26: 111, 3 May 1913c.

= Pteropus pumilus G. S. Miller, 1911. See Klingener and Creighton (1984:395).

Holotype: USNM 144760. Skin and skull. Adult female. Collected 23 Jan 1906 by E. A. Mearns. Original number 6021.

Type Locality: Balut Isalnd, Serangani Group, [Davao del Sur Province], south of Mindanao, Philippine Islands (Philippines). 

Pteropus baveanus G. S. Miller

Proc. Biol. Soc. Wash. 19: 63, 1 May 1906d.

= Pteropus alecto aterrimus Matchie, 1899. See Corbet and Hill

(1992:64).

Holotype: USNM 125482. Skin and skull. Adult male. Collected 

19 Jul 1904 by W. Grasshoff. Original number 16.

Type Locality: Bawean Island (Pulau Bawean), Java Sea. [Jawa 

Timur, Java, Indonesia]. 

Pteropus cagayanus Mearns

Proc. U.S. Natl. Mus. 28: 433, 13 May 1905.

= Pteropus hypomelanus cagayanus Mearns, 1905. See Andersen 

(1912:121).

Holotype: USNM 125289. Skin and skull. Adult male. Collected 

25 Feb 1904 by E. A. Mearns. Original number 5755.

Type Locality: Cagayan Sulu Island, near west side of the Sulu 

Sea, [Tawi Tawi Province], Philippine Islands (Philippines). 

Pteropus coxi Helgen, Helgen, and Wilson

Am. Mus. Novit. 3646: 14, 25 Jun 2009.

Holotype: USNM 3791/3953. Skin and skull; the skull is now 

lost. Probably male. Collected by the U.S. Exploring Expedition of 1838–1842. No original number.

Type Locality: Samoan Islands. 

Pteropus enganus G. S. Miller

Proc. U.S. Natl. Mus. 30: 822, 4 Jun 1906e.

= Pteropus hypomelanus enganus G. S. Miller, 1906. See Andersen (1908:364).
NUMBER 644 • 5 3

Holotype: USNM 140966. Skin and skull. Adult male. Collected 

4 Nov 1904 by W. L. Abbott. Original number 3774.

Type Locality: Pulo (Pulau) Dua, Engano (Pulau Enggano), 

[Bengkulu, Sumatra, Indonesia]. 

Pteropus faunulus G. S. Miller

Proc. U.S. Natl. Mus. 24: 785, 29 May 1902a.

Holotype: USNM 111730. Skin and skull. Adult male. Collected 

23 Jan 1901 by W. L. Abbott. Original number 864.

Type Locality: Car Nicobar, [Andaman and Nicobar Islands, 

India]. 

Pteropus geminorum G. S. Miller

Smithson. Misc. Collect. 45: 60, 6 Nov 1903a.

= Pteropus hypomelanus geminorum G. S. Miller, 1903. See Andersen (1908:363).

Holotype: USNM 104464. Skin and skull. Adult female. Collected 28 Jan 1900 by W. L. Abbott. Original number 283.

Type Locality: South Twin Island, Mergui Archipelago, [Tanintharyi, Burma]. 

Pteropus lanensis Mearns

Proc. U.S. Natl. Mus. 28: 432, 13 May 1905.

= Pteropus vampyrus lanensis Mearns, 1905. See Andersen 

(1908:368).

Holotype: USNM 123291. Skin and skull. Adult male. Collected 

7 Sep 1903 by E. A. Mearns. Original number 5626.

Type Locality: [FC: Camp] Pantar, near Lake Lanao, altitude 

1,907 ft (581m), Mindanao, [Lanao del Sur Province], Philippine Islands (Philippines). 

Pteropus lanigera H. Allen

Proc Am. Philos. Soc. 28: 70, 10 May 1890a.

= Pteropus pelagicus insularis Jacquinot and Pucheran, 1853. 

See Buden et al. (2013:113) for current use.

Lectotype: USNM 19066/37815 as designated by Helgen 

(2001:99). Skin and skull. Age and sex not given. Collector and date of collection unknown. Original number [SL: 

Wards Natural Science Establishment 4397].

Type Locality: Somoa Islands. Restricted to Caroline Islands by 

Andersen (1912:298) and probably Chuuk Lagoon Islands, 

[Federated States of Micronesia], by Buden et al. (2013:128).

Remarks: This specimen was purchased from Wards in Aug 

1890 and the skin was cataloged on 21 Aug 1890; the skull 

on 14 Nov 1899. Long considered the type by Lyon and Osgood (1909:257) and Poole and Schantz (1942:141), Helgen (2001:99) formally made it the lectotype. 

Pteropus lepidus G. S. Miller

Proc. Wash. Acad. Sci. 2: 237, 20 Aug 1900a.

= Pteropus hypomelanus lepidus G. S. Miller, 1900. See Andersen (1908:361).

Holotype: USNM 101670. Skin and skull. Adult female. Collected 15 Aug 1899 by [MC: W. L. Abbott]. No original 

number.

Type Locality: Kaju Ara or Saddle Island (Pulau Mendarik), 

Tambelan Islands (Kepulauan Tambelan), [Kepulauan Riau, 

Indonesia]. 

Pteropus mearnsi Hollister

Proc. Biol. Soc. Wash. 26: 112, 3 May 1913c.

= Pteropus speciosus Andersen, 1908. See Heaney et al. 

(1987:39).

Holotype: USNM 144739. Skin and skull. Adult male. Collected 

26 Jan 1906 by E. A. Mearns. Original number 6024.

Type Locality: Isabella (Isabela), Basilan Island, [Basilan Province], Philippine Islands (Philippines). 

Pteropus niadicus G. S. Miller

Proc. Biol. Soc. Wash. 19: 64, 1 May 1906d.

= Pteropus melanotus niadicus G. S. Miller, 1906. See Koopman 

(1994:24).

Holotype: USNM 141233. Skin and skull. Adult male. Collected 

5 Mar 1905 by W. L. Abbott. Original number 3981.

Type Locality: Teliwaa, Nias Island (Pulau Nias), [Sumatera 

Utara], Sumatra, [Indonesia]. 

Pteropus pumilus G. S. Miller

Proc. U.S. Natl. Mus. 38: 394, 19 Aug 1910.

Holotype: USNM 144758. Skin and skull. Adult male. Collected 

21 Jan 1906 by [MC: E. A. Mearns]. Original number 6019.

Type Locality: Palmas Island (Pulau Miangas), southeast of Mindanao, [Sulawesi Utara, Indonesia]. 

Pteropus samoensis Peale

U.S. Explor. Exped. 8: 20, 1848.

= Pteropus samoensis samoensis Peale, 1848.

Lectotype: USNM 8597/37860 as designated by Helgen et al. 

(2009:22). Skin and skull. Age and sex not given. Collected 

on unknown date by the U.S. Exploring Expedition. [MC: 

Skin cataloged on 30 Jul 1866; the skull on 21 Feb 1900]. 

No original number.

Type Locality: Tutuila Island, [American Samoa].

Remarks: Three specimens had long been considered syntypes 

of this taxon, (see Lyon and Osgood, 1909:258; Poole and 

Schantz, 1942:142), until Helgen et al. (2009:22) designated 

USNM 8597/37860 as the lectotype. 

Pteropus vociferus Peale

U.S. Explor. Exped. 8: 19, 1848.

= Pteropus hypomelanus? Temminck, 1853. See Andersen 

(1912:120) who discussed vociferus under P. hypomelanus 

tomesi but condidered it to be undeterminable. Simmons 

(2005:338) stated that vociferus Peale, 1848 may be an 

older name for P. hypomelanus Temminck, 1853.

Holotype: USNM 3961. Skin only. [SL: Adult] male. Collected 

on unknown date by the U.S. Exploring Expedition. [MC: 

Cataloged 31 May 1860. Original number 7].

Type Locality: Island of Mangsi, straits of Balabac, [Palawan 

Province, Philippines].
5 4 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Remarks: A type was not designated by number in the original description, but Peale (1848:19) mentions only a single 

specimen from the type locality. USNM 3961 is unquestionably it and is therefore a holotype fixed by monotypy 

(ICZN, 1999: Article 73.1.2). 

Family Rhinolophidae Gray, 1825

Genus Rhinolophus Lacepede, 1799

Rhinolophus affinis nesites Andersen

Proc. Zool. Soc. London 1905 2: 104, 17 Oct 1905a.

Holotype: USNM 104753. In alcohol, [skull removed]. Adult female. Collected 24 Jul 1900 by W. L. Abbott. No original 

number.

Type Locality: Bunguran Island (Pulau Natuna Besar), N. Natunas 

(Kepulauan Natuna Besar), [Kapulauan Riau, Indonesia]. 

Rhinolophus circe Andersen

Proc. U.S. Natl. Mus. 29: 657, 7 Mar 1906a.

= Rhinolophus acuminatus circe Andersen, 1906. See Chasen 

(1940:39).

Holotype: USNM 141343. In alcohol, [skull removed]. Adult 

male. Collected [FC: 15 Mar 1905] by W. L. Abbott. Original number 4094.

Type Locality: [FC: Mojeia River], [Pulau] Nias, [Sumatera 

Utara, Sumatra, Indonesia]. 

Rhinolophus coelophyllus shameli Tate

Am. Mus. Novit. 1219: 3, 17 Feb 1943.

= Rhinolophus shameli Tate, 1943. See Hill and Thonglongya 

(1972:183).

Holotype: USNM 267255. Skin and skull. Male, age not given. 

Collected [SL: 12 May 1937] by H. G. Deignan. Original 

number [SL: 157].

Type Locality: Ko Chang Island, Gulf of Siam, [Trat, Thailand]. 

Rhinolophus hirsutus Andersen

Ann. Mag. Nat. Hist. (series 7) 16: 289, Sep 1905b.

= Rhinolophus macrotis hirsutus Andersen, 1905. See Tate 

(1943:2).

Holotype: USNM 105487. In alcohol, [skull removed]. Adult female. Collected [MC: Dec 1887] by J. B. Steere. No original 

number.

Type Locality: Guimaras Island, [Guimaras Province], Philippine 

Islands (Philippines). 

Rhinolophus inops Andersen

Ann. Mag. Nat. Hist. (series 7) 16: 284, Sep 1905c.

Holotype: USNM 125314. In alcohol, [skull removed]. Adult 

male. Collected 8 Jul 1904 by E. A. Mearns. Original number 5713.

Type Locality: Jodaya (Todaya), altitude 4,000 ft (1,219 m), 

Mount Apo, [Davao del Sur], Mindanao, [Philippines]. 

Rhinolophus keniensis Hollister

Smithson. Misc. Collect. 66(1): 2, 10 Feb 1916c.

= Rhinolophus clivosus keniensis Hollister, 1916. See Koopman 

(1975:384).

Holotype: USNM 166352. In alcohol, skull removed. Adult 

male. Collected 27 Aug 1909 by E. Heller. Original number 

1154.

Type Locality: [FC: Base camp], West side of Mount Kenia 

(Kenya), altitude 7,000 ft (2,134 m), British East Africa 

(Kenya). 

Rhinolophus mehelyi tuneti Deleuil and Labbe

Bull. Soc. Sci. Nat. Tunis. 8: 3, 1955.

Neotype: USNM 498999 as designated by Cockrum (1977:686). 

Skin and skull. Male, age not given. Collected 4 May 1975 

by T. and P. Vaughan. Original number TPV 3244.

Type Locality: [SL: At bat cave] 4 km NE El Haouaria, [Nabeul], 

Tunisia.

Remarks: A type was not designated in the original publication 

and the type series was lost. Cockrum (1977:686) incorrectly gave the specimen number as USNM 498998 in the 

neotype designation. 

Rhinolophus minutus G. S. Miller

Proc. Wash. Acad. Sci. 2: 235, 20 Aug 1900a. Primary junior 

homonym of Vespertilio minutus Montagu, 1808 applied 

to the British race of Rhinolophus hipposideros. See Andersen (1905b:129). Renamed Rhinolophus minutillus by G. S. 

Miller (1906f:41).

= Rhinolophus pusillus minutillus G. S. Miller, 1906. See Hill 

and Yoshiyuki (1980:186).

Holotype: USNM 101715. In alcohol, [skull removed]. Adult 

male. Collected Sep 1899 by W. L. Abbott. No original 

number.

Type Locality: Pulo (Pulau) Siantan, Anambas Islands (Kepulauan Anambas), [Kepulauan Riau, Indonesia]. 

Rhinolophus nereis Andersen

Proc. Zool. Soc. London 1905 2: 90, 17 Oct 1905a.

Holotype: USNM 101714. In alcohol, [skull removed]. Adult 

female. Collected Sep 1899 by W. L. Abbott. No original 

number.

Type Locality: Pulo (Pulau) Siantan, Anambas Group (Kepulauan Anambas), [Kepulauan Riau, Indonesia]. 

Rhinolophus solitarius Andersen

Ann. Mag. Nat. Hist. (series 7) 16: 250, Aug 1905d.

= Rhinolophus trifoliatus solitarius Andersen, 1905. See Chasen 

(1940:41).

Holotype: USNM 124767. In alcohol, [skull removed]. Adult 

male. Collected 18 Jun 1904 by W. L. Abbott. Original 

number [SL: 3415].

Type Locality: Tanjong Pamuja, Banka (Pulau Bangka), [Kepulauan Bangka Belitung, Sumatra, Indonesia]. 
NUMBER 644 • 5 5

Rhinolophus spadix G. S. Miller

Proc. Wash. Acad. Sci. 3: 136, 26 Mar 1901a.

= Rhinolophus borneensis spadix G. S. Miller, 1901. See Chasen 

(1940:37).

Holotype: USNM 104752. In alcohol, [skull removed]. Adult female. Collected [MC: 5] Jun 1900 by [MC: W. L. Abbott]. 

No original number.

Type Locality: Sirhassen Island (Pulau Serasan), South Natunas (Kepulauan Natuna Selatan), [Kepulauan Riau, 

Indonesia]. 

Rhinolophus trifoliatus niasensis Andersen

Proc. U.S. Natl. Mus. 29: 658, 7 Mar 1906a.

Holotype: USNM 141350. In alcohol, originally in formalin, 

[skull removed]. Adult male. Collected 15 Mar 1905 by 

W. L. Abbott. Original number 4088.

Type Locality: [FC: Mojeia River], [Pulau] Nias, [Sumatera 

Utara, Sumatra, Indonesia]. 

Rhinolophus virgo Andersen

Proc. Zool. Soc. London 1905 2: 88, 17 Oct 1905a.

Holotype: USNM 101966. In alcohol, [skull removed]. Adult 

female. Collected 1899–1900 by L. M. McCormick. No 

original number.

Type Locality: [FC: Cave near Pasacao], Luzon, Province 

of S. Camarines (Camarines Sur), Philippine Islands 

(Philippines). 

Family Hipposideridae Lydekker, 1891

Genus Coelops Blyth, 1848

Chilophylla hirsuta G. S. Miller

Proc. U.S. Natl. Mus. 38: 395, 19 Aug 1910.

= Coelops robinsoni hirsutus (G. S. Miller, 1910). See Corbet 

and Hill (1992:116).

Holotype: USNM 144821. In alcohol, skin only. Adult female. 

Collected 2 Dec 1906 by E. A. Mearns. Original number 

6337.

Type Locality: Alag River, opposite mouth of Egbert River, [Oriental Mindoro Province], Mindoro, [Philippines].

Remarks: G. S. Miller (1910:395) noted that the entire specimen 

was supposed to be in alcohol, but that he could find only 

the skin; that remains the case today. 

Coelops inflata G. S. Miller

Proc. Biol. Soc. Wash. 41: 85, 16 Mar 1928.

= Coelops frithii inflatus G. S. Miller, 1928. See Simmons 

(2005:367).

Holotype: USNM 238991. In alcohol, [skull removed]. Adult 

male. Collected 7 Apr 1922 by A. de C. Sowerby. Original 

number 1277.

Type Locality: Near Yen-ping-fu (Nanping), altitude 2,000 ft 

(610 m), Fukien (Fujian), China. 

Genus Hipposideros Gray, 1831

Hipposideros barbensis G. S. Miller

Proc. Wash. Acad. Sci. 2: 233, 20 Aug 1900a.

= Hipposideros larvatus barbensis G. S. Miller, 1900. See Chasen 

(1940:46).

Holotype: USNM 101625. Skin and skull. Adult male. Collected 

1 Aug 1899 by [MC: W. L. Abbott]. No original number.

Type Locality: Ste. Barbe Island (Pulau Pejantan), [SL: China 

Sea], [Kepulauan Riau, Indonesia]. 

Hipposideros bicolor gilberti Johnson

Proc. Biol. Soc. Wash. 72: 183, 4 Nov 1959.

= Hipposideros ater gilberti Johnson, 1959. See Hill (1963:33).

Holotype: USNM 284170. Skin and skull. Adult male. Collected 

1 Nov 1948 by D. H. Johnson. Original number 5936.

Type Locality: Oenpelli (Gunbalanya), East Alligator River, lat. 

12o21′S, long. 133o04′E, Northern Territory, Australia. 

Hipposideros gentilis toala Shamel

J. Mammal. 21: 352. 13 Aug 1940b.

= Hipposideros ater saevus Andersen, 1918. See Hill (1963:30).

Holotype: USNM 219390. In alcohol, [skull removed]. Adult 

female. Collected 16 Sep 1917 by H. C. Raven. Original 

number [SL: 3255].

Type Locality: [MC: Bada], Toeare (Tuare), [Sulawesi Tengah], 

Celebes (Sulawesi), [Indonesia]. 

Hipposideros insolens Lyon

Proc. U.S. Natl. Mus. 40: 129, 25 Apr 1911b.

= Hipposideros galeritus insolens Lyon, 1911. See Jenkins and 

Hill (1981:286).

Holotype: USNM 154389. In alcohol, [skull removed]. Adult 

male. Collected 31 Dec 1908 by W. L. Abbott. Original 

number 6274.

Type Locality: Near Lowatsi on the upper Pasir River (Sungai 

Pasir), [Kalimantan Timur], southeastern Borneo (Kalimantan), [Indonesia]. 

Hipposideros nicobarulae G. S. Miller

Proc. U.S. Natl. Mus. 24: 781, 29 May 1902a.

See Douangboubpha et al. (2011:73) for current use.

Holotype: USNM 111874. In alcohol, [skull removed]. Adult 

male. Collected 2 Mar 1901 by W. L. Abbott. No original 

number.

Type Locality: Little Nicobar, [FC: Nicobar Islands], [Andaman 

and Nicobar Islands, India]. 

Hipposideros pelingensis Shamel

J. Mammal. 21: 353, 13 Aug 1940b.

Holotype: USNM 219433. In alcohol, [skull removed]. Adult 

male, not female as given in original description. Collected 

May 1918 by H. C. Raven. Original numbers [FC: 3571 

and 8 P].
5 6 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Type Locality: [FC: Near Liang], Peling Island (Pulau Peling), 

[Sulawesi Tengah], Celebes (Sulawesi), [Indonesia]. 

Family Megadermatidae H. Allen, 1864

Genus Lavia Gray, 1838

Lavia rex G. S. Miller

Proc. Biol. Soc. Wash. 18: 227, 9 Dec 1905a.

= Lavia frons rex G. S. Miller, 1905. See Koopman (1975:383).

Holotype: USNM 18993/38197. In alcohol, skull removed. 

Adult male. Collected in 1889 by W. L. Abbott. No original 

number.

Type Locality: Taveta, [Coast Province], German (British) East 

Africa (Kenya). 

Genus Megaderma E. Geoffroy, 1810

Megaderma carimatae G. S. Miller

Proc. U.S. Natl. Mus. 31: 63, 23 Jul 1906a.

= Megaderma spasma carimatae G. S. Miller, 1906. See Chasen 

(1940:34).

Holotype: USNM 125185. In alcohol, [skull removed]. Adult 

female. Collected 31 Aug 1904 by W. L. Abbott. Original 

number 3709.

Type Locality: Tanjong Karimata Tua, Karimata Island (Pulau 

Karimata), [Kalimantan Barat, Borneo (Kalimantan), 

Indonesia]. 

Megaderma lasiae Lyon

Proc. U.S. Natl. Mus. 52: 439, 30 Dec 1916.

= Megaderma spasma lasiae Lyon, 1916. See Chasen (1940:34).

Holotype: USNM 114249. Skin and skull. [Poole and Schantz 

(1942:119): Adult] [SL: male]. Collected 6 Jan 1902 by 

W. L. Abbott. Original number 1401.

Type Locality: Pulo (Pulau) Lasia, [FC: 14 mi (23 km) SE of Simalur Island (Pulau Simeulue)], [Aceh, Sumatra, Indonesia]. 

Megaderma niasense Lyon

Proc. U.S. Natl. Mus. 52: 440, 30 Dec 1916.

= Megaderma spasma niasense Lyon, 1916. See Chasen (1940:35).

Holotype: USNM 141305. In alcohol, [skull removed]. Adult 

male. Collected 13 Mar 1905, not 15 as in original description, by W. L. Abbott. Original number 4066.

Type Locality: Pulo (Pulau) Nias, Mojeia River, [Sumatera Utara, 

Sumatra, Indonesia]. 

Megaderma siumatis Lyon

Proc. U.S. Natl. Mus. 52: 439, 30 Dec 1916.

= Megaderma spasma siumatis Lyon, 1916. See Chasen 

(1940:35).

Holotype: USNM 114227. Skin and skull. Adult female. Collected 27 Dec 1901 by W. L. Abbott. Original number 1390.

Type Locality: Pulo (Pulau) Siumat, [Aceh, Sumatra, Indonesia]. 

Megaderma spasma celebensis Shamel

J. Mammal. 21: 352, 13 Aug 1940b.

Holotype: USNM 216945. Skin and skull. Adult male. Collected 

26 Feb 1916 by H. C. Raven. Original number [SL: 2404].

Type Locality: Likoepang (Likupang), [Sulawesi Utara], Celebes 

(Sulawesi), [Indonesia]. 

Family Rhinopomatidae Bonaparte, 1838

Genus Rhinopoma E. Geoffroy, 1818

Rhinopoma microphyllum harrisoni Schlitter and DeBlase

Mammalia 38: 658, 30 Jun 1975.

See Akmali et al. (2011:288) for current use.

Holotype: USNM 350124. Skin and skull. Adult male. Obtained 

18 Nov 1963 and prepared by H. W. Setzer. Original number 3094.

Type Locality: 10 km SE Kazerun, 29o34 ́N, 51o46 ́E, Fars Province, Iran. 

Family Emballonuridae Gervais, 1855

Genus Emballonura Temminck, 1838

Emballonura anambensis G. S. Miller

Proc. Wash. Acad. Sci 2: 236, 20 Aug 1900a.

= Emballonura alecto anambensis G. S. Miller, 1900. See Corbet 

and Hill (1992:84).

Holotype: USNM 101716. In alcohol, [skull removed]. Adult female. Collected 26 Aug 1899 by [W. L. Abbott]. No original 

number.

Type Locality: Pulo Mobur (Pulau Mubur), Anamba Islands 

(Kepulauan Anambas), [Kepulauan Riau, Indonesia]. 

Emballonura peninsularis G. S. Miller

Proc. Acad. Nat. Sci. Phila. 50: 323, 25 Jul 1898b.

= Emballonura monticola Temminck, 1838. See Tate and Archbold (1939:8).

Holotype: USNM 83575. In alcohol, [skull removed]. Adult 

male. Collected Nov 1896 by W. L. Abbott. No original 

number.

Type Locality: [MC: Lay Song Hong], Trong (Trang), Lower 

Siam (Thailand). 

Emballonura pusilla Lyon

Proc. U.S. Natl. Mus. 40: 132, 25 Apr 1911b.

= Emballonura monticola Temminck, 1838. See Tate and Archbold (1939:8).

Holotype: USNM 153940. In alcohol, skull removed. Adult 

female. Collected 29 Sep 1908 by W. L. Abbott. Original 

number 6222, not 6223 as in original publication.

Type Locality: Mankol, Kendawangan River (Sungai Kendawangan), [Kalimantan Barat], southwestern Borneo (Kalimantan), [Indonesia]. 
NUMBER 644 • 5 7

Emballonura sulcata G. S. Miller

Proc. Biol. Soc. Wash. 24: 161, 16 Jun 1911e.

= Emballonura semicaudata sulcata G. S. Miller, 1911. See Kuroda (1938:91).

Holotype: USNM 151568. Skin and skull. Adult, sex not given. 

Collected 16 Feb 1900 by H. F. Moore. No originl number.

Type Locality: Uola Island (Weno), Truk Group, [Chuuk], Caroline Islands, [Federated States of Micronesia]. 

Vespertilio semicaudatus Peale

U.S. Explor. Exped. 8: 23, 1848.

= Emballonura semicaudata semicaudata (Peale, 1848). See 

Wagner (1855:698).

Holotype: USNM 3727. [Mummy without skull. Sex and age 

unknown]. Collected on unknown date by the U.S. Exploring Expedition. [MC: Cataloged on 20 Dec 1859].

Type Locality: Samoan Islands.

Remarks: The type was not designated by number in the original description, but the description was based on a single 

specimen collected by the U.S. Exploring Expedition. This is 

the only specimen from the Expedition known to be extant 

and undoubtedly the holotype by monotypy (ICZN, 1999: 

Article 73.1.2). 

Genus Saccolaimus Temminck, 1838

Taphonycteris capito Hollister

Proc. U.S. Natl. Mus. 46: 308, 31 Dec 1913a.

= Saccolaimus saccolaimus pluto (G. S. Miller, 1910). See Simmons (2005:382).

Holotype: USNM 155149. Skin and skull. Adult female. Collected in 1909 by D. B. Mackie. Original number [SL: 1].

Type Locality: Pandan, [Catanduanes Province], Catanduanes 

Island, [Philippines]. 

Taphozous pluto G. S. Miller

Proc. U.S. Natl. Mus. 18: 396, 19 Aug 1910.

= Saccolaimus saccolaimus pluto (G. S. Miller, 1910). See Simmons (2005:382).

Holotype: USNM 144812. Skin and skull. Adult female. Collected 

28 Mar 1906 by E. A. Mearns. Original number 6054.

Type Locality: Mercedes, 9 mi (14 km) east of Zamboanga, 

[Zamboanga Province], Mindanao, [Philippines]. 

Genus Taphozous E. Geoffroy, 1818

Taphozous cavaticus Hollister

Proc. Biol. Soc. Wash. 26: 157, 30 Jun 1913d.

= Taphozous melanopogon cavaticus Hollister, 1913. See Chasen 

(1940:33).

Holotype: USNM 141092. Skin and skull. Adult female. Collected 15 Jan 1905 by W. L. Abbott. Original number 3944.

Type Locality: Moeara Dua, near Emma-haven (Teluk Bayur), 

south of Padang, [Sumatera Barat], West Sumatra, [Indonesia]. 

Taphozous solifer Hollister

Proc. Biol. Soc. Wash. 26: 157, 30 Jun 1913d.

= Taphozous melanopogon philippinensis Waterhouse, 1845. 

See Simmons (2005:384).

Holotype: USNM 113010. In alcohol. Adult male. Collected 28 

Mar 1901 by M. L. Robb. No original number.

Type Locality: [MC: Near] Peking (Beijing), [Beijing], China. 

Family Phyllostomidae Gray, 1825

Genus Anoura Gray, 1838

Anoura cultrata Handley

Proc. U.S. Natl. Mus. 112: 463, 6 Oct 1960.

Holotype: USNM 309396. Skin and skull. Adult female. Collected 7 Feb 1959 by C. O. Handley, Jr. and B. R. Feinstein. 

Original number [SL: COHJr.] 4747.

Type Locality: Tacarcuna Village [SL: Camp], altitude 3,200 ft 

(975m), Rio Pucro, Darien, Panama. 

Anoura latidens Handley

Proc. Biol. Soc. Wash. 97: 513, 17 Aug 1984.

Holotype: USNM 370119. Skin and skull. Adult female. Collected 18 Aug 1965 by A. L. and M. D. Tuttle. Original 

number SVP 629.

Type Locality: Pico Avila, [Handley (1976:79): 10o33′N, 

66o52′W], 2,150 m, 5 km NNE Caracas = “Hotel Humbolt (Humboldt), 9.4 km N Caracas”, Distrito Federal, 

Venezuela. 

Genus Ardops Miller, 1906

Ardops annectens G. S. Miller

Proc. Biol. Soc. Wash. 26: 33, 8 Feb 1913a.

 = Ardops nichollsi annectens G. S. Miller, 1913. See Jones and 

Schwartz (1967:10).

Holotype: USNM 113502. In alcohol, [skull removed]. Adult 

female. Collected 1 Feb 1902 by H. S. Branch. No original 

number.

Type Locality: Island of Guadeloupe, Lesser Antilles. 

Stenoderma luciae G. S. Miller

Proc. Acad. Nat. Sci. Phila. 54: 407, 12 Sep 1902d.

= Ardops nichollsi luciae (G. S. Miller, 1902). See Jones and 

Schwartz (1967:9).

Holotype: USNM 110921. In alcohol, [skull removed]. Adult 

female. Collected 4 Feb 1901 by H. S. Branch. No original 

number.

Type Locality: Santa (St.) Lucia Island, West Indies. 

Genus Artibeus Leach, 1821

Artibeus amplus Handley

Fieldiana Zool. n.s. 39: 164, 31 Dec 1987.
5 8 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Holotype: USNM 440932. Skin and skull. Adult female. Collected 15 Apr 1968 by N. E. Peterson, F. P. Brown, Jr., and 

J. O. Matson. Original number SVP 22086.

Type Locality: Kasmera, 270 m, 21 km SW Machiques, across 

the Rio Yasa from the Kasmera Biological Station, eastern 

foothills of the Sierra de Perija, [Handley (1976:74): 9o59′N, 

72o43′W], Estado Zulia, Venezuela. 

Artibeus hirsutus Andersen

Ann. Mag. Nat. Hist. (series 7) 18: 420, Dec 1906b.

Holotype: USNM 126449. Skin [and skull]. Adult male. Collected 16 Mar 1903 by [E. W.] Nelson and [E. A.] Goldman. 

Original number 16168.

Type Locality: La Salada, Michoacan, Mexico. 

Artibeus intermedius koopmani Wilson

Bull. Am. Mus. Nat. Hist. 206: 230, 12 Sep 1991.

= Artibeus lituratus koopmani Wilson, 1991. See Simmons 

(2005:419).

Holotype: USNM 512378. Skin and skull. Adult male. Collected 

18 Mar 1976 by D. E. Wilson. Original number 3678.

Type Locality: Maria Cleofas, Tres Marias Islands, Nayarit, Mexico. 

Artibeus jamaicensis praeceps Andersen

Ann. Mag. Nat. Hist. (series 7) 18: 421, Dec 1906b.

= Artibeus jamaicensis jamaicensis Leach, 1821. See Koopman 

(1968:6).

Holotype: USNM 113503. In alcohol, [skull removed]. Adult 

male. Collected [MC: 22 Jan 1902] by H. S. Branch. No 

original number.

Type Locality: Guadeloupe, W. I. (West Indies). 

Artibeus jamaicensis triomylus Handley

Ann. Inst. Biol. Univ. Mex. 36: 299, 20 Jun 1966b.

Holotype: USNM 126554. Skin and skull. Adult female. Collected 17 Apr 1903 by E. W. Nelson and E. A. Goldman. 

Original number 16302.

Type Locality: Papayo (El Papayo), Guerrero, Mexico. 

Genus Brachyphylla Gray, 1834

Brachyphylla pumila G. S. Miller

Proc. Biol. Soc. Wash. 31: 39, 16 May 1918.

= Brachyphylla nana G. S. Miller, 1902. See Jones and Carter 

(1976:30) who recognized pumila as a valid subspecies of 

nana.

Holotype: USNM 218354. Skull only. Age and sex not given. 

Collected [MC: Fall 1917] by W. L. Abbott. No original 

number.

Type Locality: Cave known as “Trou de Bon Dieu,” near Port de 

Paix, Haiti, Greater Antilles. 

Brachyphylla minor G. S. Miller

Proc. Biol. Soc. Wash. 26: 32, 8 Feb 1913a.

= Brachyphylla cavernarum minor G. S. Miller, 1913. See Koopman (1968:5).

Holotype: USNM 101528. In alcohol, [skull removed]. Adult 

female. Collected 14 Jun [MC: 24 Sep] 1899 by P. McDonough. No original number.

Type Locality: Coles Cave, St. Thomas Parish, Barbados, Lesser 

Antilles.

Remarks: We have been unable to resolve the difference between 

the dates given in the original description and the Museum 

Catalog for date collected. 

Brachyphylla nana G. S. Miller

Proc. Acad. Nat. Sci. Phila. 54: 409, 12 Sep 1902d.

Holotype: USNM 103828. Skull only. [Lyon and Osgood 

(1909:266): Adult], sex not given. Collected 10 Mar 1900 

by W. Palmer and J. H. Riley. Original number 108.

Type Locality: El Guama, [Pinar del Rio], Cuba, Greater Antilles. 

Genus Carollia Gray, 1838

Carollia castanea H. Allen

Proc. Am. Philos. Soc. 28: 19, 25 Feb 1890b.

Holotype: USNM 12914/36834. In alcohol, skull removed. [Lyon 

and Osgood (1909:265): Adult] male. Collected [Lyon and 

Osgood (1909:265): probably 1876] by J. C. Zeledon. 

Type Locality: [Pine (1972:18): Angostura], [San Jose], Costa Rica.

Remarks: The holotype was published as USNM 12914, to 

which USNM 36834 belongs. The skin was cataloged 23 on 

Jun 1878; the skull 11 Nov 1893. 

Hemiderma subrufum Hahn

Proc. Biol. Soc. Wash. 18: 247, 9 Dec 1905.

= Carollia subrufa (Hahn, 1905). See G. S. Miller (1924:54).

Holotype: USNM 75127. [Skin and skull.] Adult male. Collected 

29 Jul 1895 by E. W. Nelson and E. A. Goldman. Original 

number 8235.

Type Locality: Santa Ifigenia (Efigenia), [FC: altitude 1,200 ft 

(366 m)], [Miller and Kellogg (1955:74): 8 mi (13 km) 

northwest of Tepantepec near west coast of Oaxaca], Oaxaca, Mexico. 

Hemiderma tricolor G. S. Miller

Proc. Acad. Nat. Sci. Phila. 54: 408, 12 Sep 1902d.

= Carollia perspicillata (Linnaeus, 1758). See Pine (1972:50).

Holotype: USNM 114005. Skin and skull. Adult female. Collected 5 Dec 1901 by W. Foster. Original number 589.

Type Locality: Cave at Sapucay, [Paraguari], Paraguay. 

Genus Centurio Gray, 1842

Centurio mcmurtrii H. Allen

Proc. Acad. Nat. Sci. Phila. 13: 360, ordered published 26 Nov 

1861; receipt acknowledged 31 Dec 1861 by the Boston Society of Natural History.
NUMBER 644 • 5 9

= Centurio senex senex Gray, 1842. See Rehn (1901:297).

Holotype: USNM 8289/37806. [In alcohol, skull removed]. 

[Rehn (1901:300): Adult male]. Collected by [MC: C. Sartorius], date unknown. Skin cataloged between Jul and Oct 

1865; skull 14 Nov 1899. No original number.

Type Locality: Mirador, [Veracruz], Mexico.

Remarks: The type was not designated by number in the original 

description, but the description was based on a single specimen in the Smithsonian collection. Rehn (1901:300) determined that specimen to be USNM 8289/37806. 

Centurio senex greenhalli Paradiso

Mammalia 31: 601, Dec 1968. 

Holotype: USNM 346847. In alcohol with skull removed. Adult 

female. Collected 8 Nov 1961 by A. Greenhall. Original 

number [SL: 61-2928].

Type Locality: Port of Spain, St. George County (Parish), Trinidad, [Trinidad and Tobago]. 

Genus Chiroderma Peters, 1860

Chiroderma gorgasi Handley

Proc. U.S. Natl. Mus. 112: 464, 6 Oct 1960.

= Chiroderma trinitatum gorgasi Handley, 1960. See BarrigaBonilla (1965:245) for first use of current name combination and Gardner (2008a:324) for current use.

Holotype: USNM 309903. Skin, skull, [and skeleton]. Adult 

male. Collected 6 Mar 1959 by C. O. Handley, Jr, and B. R. 

Feinstein. Original number [SL: COH, Jr.] 5436.

Type Locality: Tacarcuna Village [SL: Camp], 3,200 ft (975 m), 

Rio Pucro, Darien, Panama. 

Chiroderma isthmicum G. S. Miller

Proc. U.S. Natl. Mus. 42: 25, 6 Mar 1912c.

= Chiroderma villosum jesupi J. A. Allen. See Handley (1960: 

466).

Holotype: USNM 173834. In alcohol, [skull removed]. Adult female. Collected May 1911 by A. Busck. No original number.

Type Locality: Cabima (La Cabima), [Panama], Panama. 

Chiroderma salvini scopaeum Handley

Ann. Inst. Biol. Univ. Mex. 36: 297, 20 Jun 1966b.

Holotype: USNM 338711. Skin and skull. Adult female. Collected 16 Aug 1960 by A. L. Gardner. Original number 

1565.

Type Locality: Pueblo Juarez, Colima, Mexico.

Remarks: Previously University of Arizona, Department of Biology number 7952. Received in exchange from E. L. Cockrum. Cataloged USNM 338711 on 26 Aug 1964. 

Genus Choeroniscus Thomas, 1928

Choeroniscus periosus Handley

Proc. Biol. Soc. Wash. 79: 84, 23 May 1966c.

Holotype: USNM 344918. In alcohol, skull removed. Adult female. Collected 1 Feb 1963 by W. A. Thornton. Original 

number 592.

Type Locality: Rio Raposo, near sea level, 27 km south of Buenaventura, Departamento de Valle [del Cauca], Colombia. 

Genus Dermanura Gervais, 1856

We follow Hoofer et al. (2008) and Solari et al. (2009) in 

recognizing Dermanura at the generic level.

Artibeus aztecus Andersen

Ann. Mag. Nat. Hist. (series 7) 18: 422, Dec 1906b.

= Dermanura azteca (Andersen, 1906). See Owen (1987:64) for 

name combination and Solari et al. (2009:284) for current use.

Holotype: USNM 52050. Skin [and skull]. Adult male. Collected 

12 Feb 1893 by E. W. Nelson. Original number 4332.

Type Locality: Tetela del Volcan, [FC: altitude 8,000 ft (2,438 m)], 

Morelos, Mexico. 

Artibeus gnomus Handley

Fieldiana Zool. n.s. 39: 167, 31 Dec 1987.

= Dermanura gnoma (Handley, 1987). See Hoofer et al. 

(2008:10).

Holotype: USNM 387534. Skin and skull. Adult female. Collected 14 Jun 1966 by A. L. and M. D. Tuttle. Original number SVP 9298.

Type Locality: El Manaco, [Handley (1976:69): 150 m, 6o17 ́N, 

61o19 ́W], KM 74, 59 km SE El Dorado, Bolivar, Venezuela. 

Artibeus incomitatus Kalko and Handley

Z. Saugetierkd. 59: 260, Oct 1994.

= Dermanura watsoni (Thomas, 1901). See Solari et al. (2009: 

284).

Holotype: USNM 579125. Skin and skull. Adult male. Collected 

19 Mar 1991 by C. O. Handley, Jr. Original number COH 

17002.

Type Locality: A meter or so above sea level near West Point, Isla 

Escudo de Veraguas, Bocas del Toro, Panama. 

Dermanura phaeotis G. S. Miller

Proc. Acad. Nat. Sci. Phila. 54: 405, 12 Sep 1902d.

= Dermanura phaeotis phaeotis G. S. Miller, 1902. See Hoofer et 

al. (2008:10) for current use.

Holotype: USNM 108176. Skin and skull. Adult female. Collected 10 Feb 1901 by E. W. Nelson and E. A. Goldman. 

Original number 14537.

Type Locality: Chichen Itza, Yucatan, [SL: Mexico]. 

Dermanura rava G. S. Miller

Proc. Acad. Nat. Sci. Phila. 54: 404, 12 Sep 1902d.

See Hoofer et al. (2008:10) for current use.

Holotype: USNM 113338. Skin and skull. Adult male. Collected 

10 Aug 1900 by G. Flemming. Original number 12.
6 0 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Type Locality: San Javier, [SL: altitude 60 ft (18 m)], northern 

Ecuador. 

Stenoderma tolteca Saussure

Rev. Mag. Zool. series 2, 12: 427, Oct 1860.

= Dermanura tolteca tolteca (Saussure, 1860). See Owen 

(1987:64) for name combination and Solari et al. (2009:284) 

for current use.

Neotype: USNM 6981/38954 as designated by Davis (1969:26). 

In alcohol with skull removed. Adult male. Collected on unknown date by [MC: C. Sartorius]. Original number [SL: 

105]. [MC: Skin cataloged on 13 Apr 1863; skull on 27 

Nov 1931].

Type Locality: Mirador, Veracruz, Mexico. 

Genus Ectophylla H. Allen, 1892

Ectophylla alba H. Allen

Proc. U.S. Natl. Mus. 15: 442, 26 Oct 1892.

Holotype: USNM 15950. Skin, without skull, in alcohol. [A. L. 

Gardner’s notation on card in the type file: penis proves 

specimen to be a male]. Age indeterminate. Collected [MC: 

1 Jul 1887 by C. H. Townsend. Original number 313].

Type Locality: Vicinity of Segovia River (Rio Coco), [Gracias a 

Dios], eastern Honduras.

Remarks: According to Miller and Kellogg (1955:77), the locality given above is equal to Comarca de El Cabo, northeastern Nicaragua. 

Genus Erophylla Miller, 1906

Erophylla planifrons mariguanensis Shamel

J. Wash. Acad. Sci. 21: 252, 4 Jun 1931a.

= Erophylla sezekorni mariguanensis Shamel. 1931. See Hall 

(1981:171).

Holotype: USNM 255593. In alcohol, [skull removed]. Adult 

male. Collected 20 Jul 1930 by P. Bartsch. Original number 

[MC: 169].

Type Locality: Abrahams Hill, Mariguana (Mayaguana) Island, 

[Bahamas]. 

Phyllonycteris bombifrons G. S. Miller

Proc. Biol. Soc. Wash. 13: 36, 29 May 1899d.

= Erophylla bombifrons bombifrons (G. S. Miller, 1899). See G. 

S. Miller (1906g:84).

Holotype: USNM 86274. In alcohol, [skull removed]. Adult male. 

Collected 18 Jan 1899 by P. Beckwith. No original number.

Type Locality: In a limestone cave near Bayamon, Province of 

San Juan, Puerto Rico. 

Phyllonycteris planifrons G. S. Miller

Proc. Biol. Soc. Wash. 13: 34, 29 May 1899d.

= Erophylla sezekorni planifrons (G. S. Miller, 1899d). See G. M. 

Allen (1917:167).

Holotype: USNM 62517. In alcohol, [skull removed]. Adult male. 

Collected 18 Mar 1886 by J. E. Benedict. No original number.

Type Locality: Nassau, New Providence [Island], Bahamas.

Remarks: Originally USNM 15453, this specimen was recataloged USNM 62517 on 13 Jul 1895. 

Genus Glossophaga E. Geoffroy, 1818

Glossophaga elongata G. S. Miller

Proc. Biol. Soc. Wash. 13: 124, 6 Apr 1900d.

= Glossophaga longirostris elongata G. S. Miller, 1900. See 

Koopman (1958:437).

Holotype: USNM 101871. Skin and skull from specimen in 

formalin. Adult female. Collected 4 Dec 1899 by [SL: L. J. 

Guthrie]. No original number.

Type Locality: Willemstad, Curacao, [Netherlands Antilles],West 

Indies.

Remarks: According to the museum catalog, this specimen was 

made into a skin and skull on 27 Feb 1900. 

Glossophaga longirostris campestris Webster and Handley

Occas. Pap., Mus. Texas Tech Univ. 100: 8, 18 Mar 1986.

Holotype: USNM 389033. Skin and skull. Adult female. Collected 

8 Apr 1967 by N. Peterson, D. Peacock, and R. Peacock, 

Smithsonian Venezuelan Project. Original number SVP 12679.

Type Locality: Hato San Jose, 300 m, [Handley (1976:73): 

6o49′N, 63o29′W], 20 km W Paragua, = 146 km S, 7 km E 

Ciudad Bolivar, Bolivar, Venezuela. 

Glossophaga mutica Merriam

Proc. Biol. Soc. Wash. 12: 18, 27 Jan 1898a.

= Glossophaga soricina mutica Merriam, 1898. See G. S. Miller 

(1913e:420).

Holotype: USNM 89271. [Skin and skull]. Adult male. Collected 

8 May 1897 by E. W. Nelson and E. A. Goldman. Original 

number 10976.

Type Locality: Maria Madre Island, [FC: altitude 50 ft (15 m)], 

Tres Marias Islands, [Nayarit], Mexico. 

Glossophaga rostrata G. S. Miller

Proc. Biol. Soc. Wash. 26: 32, 8 Feb 1913a.

= Glossophaga longirostris rostrata G. S. Miller, 1913. See G. S. 

Miller (1913e:423).

Holotype: USNM 111500. Skin and skull. Adult male. Collected 

25 Aug 1900 by P. Gellineau. Original number 29.

Type Locality: Westerhall Estate, Grenada, Lesser Antilles. 

Glossophaga soricina microtis G. S. Miller

Proc. U.S. Natl. Mus. 46: 419, 31 Dec 1913e.

= Glossophaga soricina soricina (Pallas, 1766). See Alvarez et 

al. (1991:1).

Holotype: USNM 115061. In alcohol, [skull removed]. Adult female. Collected Jun 1901 by W. Foster. Original number 66.

Type Locality: Sapucay (Sapucai), [Paraguari], Paraguay. 
NUMBER 644 • 6 1

Glossophaga villosa H. Allen

Proc. U.S. Natl. Mus. 18: 779, 27 Oct 1896. Primary junior 

homonym of Glossophaga villosa Rengger, 1830. Replaced

with Glossophaga truei by H. Allen (1897:153).

= Glossophaga soricina soricina (Pallas, 1766). See G. S. Miller 

(1913e:418).

Holotype: USNM 9523/37388. Skin in alcohol, skull removed. 

Adult, sex unknown. Collector and date of collection unknown. The skin was cataloged between 12 Feb and 17 Mar 

1870; the skull 21 Apr 1898.

Type Locality: La Guayra (Guaira), [Vargas], Venezuela.

Remarks: Published as USNM 9523 to which skull number 

37388 belongs. H. Allen referred to his two specimens as 

9522 and 9523 in error. They are actually 9523 and 9524, 

with 9524 being a specimen without any locality data. The 

type is designated by number 9522, an error for 9523. 

The type locality is questionable as it is given as Guyana, 

Venezuela in the original museum catalog. A copy of this 

catalog, and no doubt the one H. Allen referred to for the 

locality data, gives it as Guyara, Venezuela. See Lyon and 

Osgood (1909:264) or Poole and Schantz (1942:131) for 

more lengthy discussion. 

Genus Leptonycteris Lydekker, 1891

Leptonycteris curasoae G. S. Miller

Proc. Biol. Soc. Wash. 13: 126, 6 Apr 1900d.

Holotype: USNM 101851. In alcohol, [skull removed]. Adult 

male. Collected [MC: Nov-Dec 1899 by L. J. Guthrie]. No 

original number.

Type Locality: [Note with specimen: 1 mi (1.6 km) NW of Willemstad], Curacao, [Netherlands Antilles], West Indies. 

Genus Lichonycteris Thomas, 1895

Lichonycteris degener G. S. Miller

J. Mammal. 12: 411, 11 Nov 1931b.

See Griffiths and Gardner (2008:242) for current use.

Holotype: USNM 239520. Skin and skull. Adult female. Collected 22 Jul 1923 by C. R. Aschemeier. Original number 

[SL: 1437].

Type Locality: Para (Belem), [Para], Brazil. 

Genus Lonchophylla Thomas, 1903

Lonchophylla cadenai Woodman and Timm

Proc. Biol. Soc. Wash. 119: 462, 22 Dec 2006.

Holotype: USNM 483359. Skin and skull. Adult male. Collected 18 Feb 1967 by A. A. Arata and M. E. Thomas. 

Original number 858.

Type Locality: 29 km SE Buenaventura, 75 m elevation, 

east bank of Rio Zabaletas, across from village of Zabaletas, 3o44′N, 76o57′W, Valle del Cauca Department, 

Colombia. 

Lonchophylla concava Goldman

Smithson. Misc. Collect. 63(5): 2, 14 Mar 1914a.

See Albuja and Gardner (2005:446) for current use.

Holotype: USNM 179621. Skin and skull. Adult male. Collected 

20 May 1912 by E. A. Goldman. Original number 21701.

Type Locality: Cana, altitude 2,000 ft (610 m), [Darien], eastern 

Panama. 

Lonchophylla fornicata Woodman

Proc. Biol. Soc. Wash. 120: 343, 17 Oct 2007.

Holotype: USNM 483360. Skin and skull. Adult male. Collected 

18 Nov 1966 by M. E. Thomas. [MC: and A. A. Arata]. 

Original number 914.

Type Locality: 29 km SE Buenaventura, 75 m elevation, east 

bank of Rio Zabaletas, across from village of Zabaletas, 

3o44′N, 76o57′W, Valle del Cauca Department, Colombia. 

Lonchophylla robusta G. S. Miller

Proc. U.S. Natl. Mus. 42: 23, 6 Mar 1912c.

Holotype: USNM 173854. In alcohol, [skull removed]. Adult 

male. Collected 14 Apr 1911 by A. Busck. No original 

number.

Type Locality: In cave on Chilibrillo River, [Panama], Panama. 

Genus Lophostoma d’Orbigny, 1836

Lophostoma kalkoae Velazco and Gardner

J. Mammal. 93: 606, 30 Apr 2012.

Holotype: USNM 582249. In alcohol, skull removed. Adult 

male. Collected 11 Oct 1998 by E. K. V. Kalko. Original 

number 118.

Type Locality: Pipeline Road, near the former Limbo Hunt Club, 

9o9′50 ̋N, 79o45′10 ̋W, Soberania National Park, Colon 

Province, Panama. 

Lophostoma venezuelae Robinson and Lyon

Proc. U.S. Natl. Mus. 24: 154, 3 Oct 1901.

= Lophostoma brasiliense Peters, 1866. See Simmons (2005:406).

Holotype: USNM 102919. In alcohol, [skull removed]. Adult 

female. Collected [SL: 4 Aug 1900 by M. W. Lyon, Jr. and 

W. Robinson. Original number 199].

Type Locality: Macuto, [Lyon and Osgood (1909:267): 8 mi (13 

km) east of La Guaira], [Vargas], Venezuela. 

Genus Macrotus Gray, 1843

Macrotus californicus Baird

Proc. Acad. Nat. Sci. Phila. 10: 116, ordered to be published 25 

May 1858.

Holotype: USNM 2347. In alcohol. Age and sex not given. Collected on unknown date by G. H. Thomas. No original 

number. [MC: Cataloged 27 Mar 1857].

Type Locality: [Miller and Rehn (1901:278): Old] Fort Yuma, 

[Miller (1924:46): Imperial County], California, [Miller 
6 2 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

(1924:46): on right bank of Colorado River, opposite present town of Yuma, Arizona].

Remarks: The holotype was not designated by number in the 

original description, which was based on a single specimen 

from Fort Yuma, California. Measurements given in the 

description match those of USNM 2347 as given in Baird 

(1859) and is therefore the holotype fixed by monotypy 

(ICZN, 1999: Article 37.1.2). This specimen cannot be 

found. The last record of it is H. Allen (1894:39) in which 

he lists the specimen by number and refers to it as the type. 

Macrotus waterhousii compressus Rehn

Proc. Acad. Nat. Sci. Phila. 56: 434, 30 Jun 1904.

Holotype: USNM 122484. [Skin and skull]. Adult female. Collected 

6 Jul 1903 by S. H. Derickson. Original number [SL: 90].

Type Locality: Eleuthera [SL: Island], Bahamas.

Remarks: Received from J. H. Riley, original number 1108. 

Macrotus waterhousii heberfolium Shamel

J. Wash. Acad. Sci. 21: 252, 4 Jun 1931a.

= Macrotus waterhousii waterhousii Gray, 1843. See Anderson 

and Nelson (1965:14).

Holotype: USNM 255651. In alcohol, [skull removed]. Adult 

male. Collected 23 Jul 1930 by P. Bartsch. No original 

number.

Type Locality: Kingston, Providencialis (Providenciales) Island, 

[Caicos Islands, Turks and Caicos Islands]. 

Macrotus waterhousii jamaicensis Rehn

Proc. Acad. Nat. Sci. Phila. 56: 432, 27 Jun 1904.

Holotype: USNM 8553/37543. Skin and skull. Adult female. 

Collected on unknown date by W. T. March. No original 

number. [MC: Skin cataloged on 3 Jul 1866; the skull on 6 

Oct 1898].

Type Locality: Spanishtown (Spanish Town), [St. Catherine], 

Jamaica. 

Genus Micronycteris Gray, 1866

Micronycteris megalotis mexicanus G. S. Miller

Proc. Acad. Nat. Sci. Phila. 50: 329, 2 Aug 1898c.

= Micronycteris megalotis (Gray, 1842). See Porter et al. 

(2007:1211) who assigned mexicana to megalotis at unspecified rank.

Holotype: USNM 52105. In alcohol, [skull removed]. Adult female. Collected 4 Apr 1892 by E. W. Nelson. Original number 2389.

Type Locality: Plantinar, [FC: altitude about 4,000 ft (1,219 m)], 

Jalisco, Mexico. 

Micronycteris microtis G. S. Miller

Proc. Acad. Nat. Sci. Phila. 50: 328, 12 Jul 1898c.

Holotype: USNM 16366/23364. Skin and skull [SL: originally in 

alcohol]. Adult male. Collected on unknown date by L. F. H. 

Birt. No original number. [MC: Cataloged 2 Feb 1889 as an 

alcoholic; skull cataloged 16 Apr 1889].

Type Locality: Greytown (San Juan del Norte), [Rio San Juan], 

Nicaragua. 

Micronycteris sanborni Simmons

Am. Mus. Novit. 3158: 6, 14 Feb 1996.

Holotype: USNM 555702. Skin, skull, [and photographs]. Adult 

female. Collected 26 Mar 1978 by A. L. Gardner. Original 

number 13745.

Type Locality: Sitio Luanda, Itaitera, 4 km S of Crato, Ceara, 

Brazil. 

Genus Mimon Gray, 1847

Mimon cozumelae Goldman

Proc. Biol. Soc. Wash. 27: 75, 11 May 1914b.

Holotype: USNM 203191. Skin and skull. Adult, sex unknown. 

Collected on unknown date by G. F. Gaumer. No original 

number. [MC: Cataloged 5 May 1914]. [SL: 11030-X].

Type Locality: Cozumel Island, off the east coast of Yucatan 

(Quintana Roo), Mexico.

Remarks: Originally numbered 6232 in the L. L. Dyche collection. 

Mimon crenulatum keenani Handley

Proc. U.S. Natl. Mus. 112: 460, 6 Oct 1960.

Holotype: USNM 311951. Skin and skull. Adult male. Collected 

25 Aug 1959 by C. M. Keenan. Original number 4127.

Type Locality: [SL: Civilian housing area], Fort Gulick, Panama 

Canal Zone (Colon), [Panama]. 

Genus Monophyllus Leach, 1821

Monophyllus clinedaphus G. S. Miller

Proc. Wash. Acad. Sci. 2: 36, 30 Mar 1900e.

= Monophyllus redmani clinedaphus G. S. Miller, 1900. See 

Schwartz and Jones (1967:6).

Holotype: USNM 5120/37405. In alcohol, [skull removed]. 

Adult male. No additional data given. [MC: Fluid-preserved 

specimen cataloged 30 Oct 1861; the skull 2 May 1898].

Type Locality: No locality given. Schwartz and Jones (1967:6) 

restricted the type locality to the vicinity of Baracoa, Oriente 

Province (Guantanamo), Cuba. 

Monophyllus cubanus G. S. Miller

Proc. Acad. Nat. Sci. Phila. 54: 410, 11 Sep 1902d.

= Monophyllus redmani clinedaphus G. S. Miller, 1900. See 

Schwartz and Jones (1967:6).

Holotype: USNM 113674. Skin and skull. Adult male. Collected 

6 Feb 1902 by W. Palmer. Original number 645.

Type Locality: Baracoa, [Guantanamo], eastern Cuba. 

Monophyllus cubanus ferreus G. S. Miller

Proc. Biol. Soc. Wash. 31: 40, 16 May 1918.
NUMBER 644 • 6 3

= Monophyllus redmani clinedaphus G. S. Miller, 1900. See 

Schwartz and Jones (1967:6).

Holotype: USNM 219151. Skin and skull. Adult male. Collected 

7 Dec 1917 by W. L. Abbott. Original number [SL: A].

Type Locality: In a cave 8 mi (13 km) W. S. W. Jeremie, [Schwartz 

and Jones (1967:6): Department du Sud], Haiti. 

Monophyllus luciae G. S. Miller

Proc. Acad. Nat. Sci. Phila. 54: 411, 11 Sep 1902d.

= Monophyllus plethodon luciae G. S. Miller, 1902. See Schwartz 

and Jones (1967:13).

Holotype: USNM 106095. In alcohol, [skull removed]. Adult 

male. Collected 4 Feb 1901 by H. S. Branch. No original 

number.

Type Locality: Santa (St.) Lucia Island. 

Monophyllus plethodon G. S. Miller

Proc. Wash. Acad. Sci. 2: 35, 30 Mar 1900e.

= Monophyllus plethodon plethodon G. S. Miller, 1900. See 

Schwartz and Jones (1967:12).

Holotype: USNM 101530. In alcohol, [skull removed]. Adult 

male. Collected 1 Aug 1899 by P. McDonough. No original 

number.

Type Locality: St. Michaels (Saint Michael) Parish, Barbados. 

Monophyllus portoricensis G. S. Miller

Proc. Wash. Acad. Sci. 2: 34, 30 Mar 1900e.

= Monophyllus redmani portoricensis G. S. Miller, 1900. See See 

Schwartz and Jones (1967:11).

Holotype: USNM 86261. In alcohol, [skull removed]. Adult 

male. Collected 18 Jan 1899 by P. Beckwith. No original 

number.

Type Locality: Cave near Bayamon, [San Juan], Puerto Rico. 

Genus Phyllonycteris Gundlach, 1861

Phyllonycteris obtusa G. S. Miller

Smithson. Misc. Collect. 81(9): 10, 30 Mar 1929.

= Phyllonycteris poeyi obtusa G. S. Miller, 1929. See Jones and 

Carter (1976:30).

Holotype: USNM 253095. Skull only. Age and sex unknown. 

Collected Mar 1925 by G. S. Miller, Jr. No original number.

Type Locality: In the “crooked cave” near the Atalaye plantation, St. Michel, Haiti. 

Reithronycteris aphylla G. S. Miller

Proc. Acad. Nat. Sci. Phila. 50: 334, 2 Aug 1898c.

= Phyllonycteris (Reithronycteris) aphylla (G. S. Miller, 1898). 

See Koopman (1952:257).

Holotype: USNM 255514. In alcohol, [skull removed]. Adult 

male. Collector and date of collection unknown. [MC: Cataloged as found in collection without a number on 16 Jul 

1930].

Type Locality: Jamaica.

Remarks: Described as Number 9, Museum of the Institute of 

Jamaica. 

Genus Platyrrhinus Saussure, 1860

Platyrrhinus fusciventris Velazco, Gardner, and Patterson

Zool. J. Linn. Soc. 159: 803, Jul 2010.

Holotype: USNM 560806. Skin and skull. Adult female. Collected 

23 Feb 1985 by A. L. Gardner. Original number 14605.

Type Locality: Base camp, Cerro Neblina, 00o49′50 ̋N, 

66o09′40 ̋W, 140 m, Amazonas, Venezuela. 

Platyrrhinus nitelinea Velazco and Gardner

Proc. Biol. Soc. Wash. 122: 268, 14 Oct 2009.

Holotype: USNM 513461. Skin and skull. Adult female. Collected 9 Aug 1976 by A. L. Gardner. Original number 

13227.

Type Locality: 1 km SW Puente de Moromoro, 3,480 ft 

(1,060 m), 03o44′S, 79o44′W, El Oro Province, Ecuador. 

Vampyrops aquilus Handley and Ferris

Proc. Biol. Soc. Wash. 84: 521, 29 Feb 1972.

= Platyrrhinus aquilus (Handley and Ferris, 1972). See Velazco 

and Gardner (2009:253).

Holotype: USNM 338025. Skin and skull. Lactating adult female. Collected 19 Feb 1964 by C. O. Handley, Jr. Original 

number 12655.

Type Locality: Head of Rio Pucro, 4,100 ft (1,250 m), Cerro 

Mali, Darien, Panama. 

Vampyrops aurarius Handley and Ferris

Proc. Biol. Soc. Wash. 84: 522, 29 Feb 1972.

= Platyrrhinus aurarius (Handley and Ferris, 1972). See Koopman (1993:190).

Holotype: USNM 387163. Skin and skull. Adult male. Collected 

18 May 1966 by M. D. Tuttle and A. L. Tuttle. Original 

number SVP 8329.

Type Locality: KM 125, 1,000 m, 85 km SSE El Dorado, [Handley (1976:74): 5o59′N, 61o26′W], Bolivar, Venezuela. 

Vampyrops fumosus G. S. Miller

Proc. Acad. Nat. Sci. Phila. 54: 405, 11 Sep 1902d.

= Platyrrhinus infuscus (Peters, 1880). See Koopman (1993: 

191).

Holotype: USNM 105530. Skin and skull. Adult female. Collected 24 Mar 1901 by J. B. Steere. Original number 83, not 

24 as given in original description.

Type Locality: [SL: Hyutanaham (Huitanaa), Upper] Purus 

River, [Amazonas], Brazil. 

Vampyrops latus Handley and Ferris

Proc. Biol. Soc. Wash. 84: 519, 29 Feb 1972.

= Platyrrhinus brachycephalus brachycephalus (Rouk and 

Carter, 1972). See Gardner (2008b:322) for current use.
6 4 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Holotype: USNM 364408. Skin, skull, [and partial skeleton]. 

Adult male. Collected 12 Aug 1964 by A. L. Tuttle. Original 

number 1789.

Type Locality: San Juan, 900 ft (274 m), Province of Oxapampa, 

Department of Pasco, Peru. 

Vampyrops latus saccharus Handley and Ferris

Proc. Biol. Soc. Wash. 84: 521, 29 Feb 1972.

= Platyrrhinus brachycephalus saccharus (Handley and Ferris, 

1972). See Gardner (2008b:333) for current use.

Holotype: USNM 408411. Skin and skull. Adult male. Collected 

20 Jul 1967 by N. E. Peterson. Original number SVP 14362.

Type Locality: Manacal, 300 m, 5 km S and 25 km E Carupano, 

[Handley (1976:76): 10o37′N, 61o09′W], Sucre, Venezuela. 

Genus Rhinophylla Peters, 1865

Rhinophylla alethina Handley

Proc. Biol. Soc. Wash. 79: 86, 23 May 1966c.

Holotype: USNM 324988. Skin and skull. Adult male. Collected 

13 Jul 1962 by W. A. Thornton. Original number 172.

Type Locality: [SL: Virology Field Station], at the Rio Raposo, 

near sea level, 27 km south of Buenaventura, Departamento 

de Valle [del Cauca], Colombia. 

Genus Sturnira Gray, 1842

Sturnira angeli de la Torre

Proc. Biol. Soc. Wash. 79: 271, 1 Dec 1966.

See Velazco and Patterson (2013:694) for current use.

Holotype: USNM 361881. Skin and skull. Adult male. Collected 

21 Feb 1962 by R. F. Klinikowski. No original number.

Type Locality: 6 mi (10 km) NE Roseau, 1,000 ft (305 m), St. 

Paul Parish, Dominica, Windward Islands, West Indies. 

Remarks: Originally numbered 5354 in the Albert Schwartz 

collection. 

Sturnira aratathomasi Peterson and Tamsitt

Life Sci. Occas. Pap., Royal Ont. Mus. 12: 1, 15 Nov 1968.

Holotype: USNM 395158. Skin, skull, [and partial skeleton]. 

Adult male. Collected 1 Dec 1966 by M. Thomas [SL: ArataThomas]. Original number University del Valle-Tulane 635.

Type Locality: 2 km south of Pance, 1,650 m, approximately 

20 km southwest of Cali, Dept. (Departamento) Valle [del 

Cauca], Colombia.

Remarks: Published as University del Valle-Tulane number 635. 

Cataolged USNM 395158 some time between 2 Jul and 10 

Sep 1968. 

Sturnira lilium bogotensis Shamel

Proc. Biol. Soc. Wash. 40: 129, 26 Sep 1927.

= Sturnira bogotensis Shamel, 1927. See Handley (1976:25).

Holotype: USNM 251989. Skin and skull. Adult female. Collected [SL: 25 Nov 1925] by [MC: Niceforo Maria]. No 

original number.

Type Locality: Bogota, Estacion La Uribe, [Cundinamarca], 

Colombia. 

Sturnira lilium parvidens Goldman

Proc. Biol. Soc. Wash. 30: 116, 23 May 1917.

= Sturnira parvidens Goldman, 1917. See Iudica (2000:197) for 

first use of name combination and Velazco and Patterson 

(2013:694) for current use.

Holotype: USNM 126555. [Skin and skull]. Adult female. Collected 17 Apr 1903 by E. W. Nelson and E. A. Goldman. 

Original number 16313.

Type Locality: Papayo (El Papayo), about 25 mi (40 km) northwest of Acapulco, Guerrero, Mexico. 

Sturnira paulsoni de la Torre and Schwartz

Proc. Biol. Soc. Wash. 79: 301, 1 Dec 1966.

See Velazco and Patterson (2013:694) for current use.

Holotype: USNM 361882. Skin and skull. Adult female. 

Collected 30 Dec 1961 by D. R. Paulson. No original 

number.

Type Locality: Lowrt, 1,000 ft (305 m), St. Andrew Parish, St. 

Vincent, British Windward Islands, Lesser Antilles, [Saint 

Vincent and the Grenadines].

Remarks: Originally numbered 5333 of the Albert Schwartz 

collection. 

Sturnira thomasi de la Torre and Schwartz

Proc. Biol. Soc. Wash. 79: 299, 1 Dec 1966.

Holotype: USNM 361883. Skin and skull. Adult male. Collected 

26 Jan 1963 by R. Thomas. No original number.

Type Locality: Sofaia, 1,200 ft (366 m), Guadeloupe, French 

Leeward Islands, Lesser Antilles.

Remarks: Originally numbered 5413 of the Albert Schwartz collection. 

Sturnirops mordax Goodwin

Am. Mus. Novit. 976: 1, 4 May 1938.

= Sturnira mordax (Goodwin, 1938). See Davis et al. (1964:381).

Holotype: USNM 250310. Skin and skull. Adult male. Collected 

14 Jul 1931 by C. F. Underwood. Original number [SL:543. 

26881-X].

Type Locality: El Sauce Peralta, a farm on the Atlantic Railroad, 

less than halfway from San Jose to Limon, probably about 

1,000 ft (305 m) elevation, [Cartago, Costa Rica]. 

Genus Trachops Gray, 1847

Trachops coffini Goldman

Proc. Biol. Soc. Wash. 38: 23, 12 Mar 1925.

= Trachops cirrhosus coffini Goldman, 1925. See Felten (1956: 

369).

Holotype: USNM 244266. Skin, skull, and nearly complete skeleton. Adult female. Collected 30 Jun 1923 by H. Malleis. 

Original number 388.

Type Locality: Guyo, Peten, Guatemala. 
NUMBER 644 • 6 5

Genus Uroderma Peters, 1866

Uroderma convexum Lyon

Proc. Biol. Soc. Wash. 15: 83, 25 Apr 1902.

See Mantilla-Meluk (2014:17) for current use.

Holotype: USNM 111722. In alcohol, [skull removed]. Young 

adult female. Collected 28 May 1901 by J. W. Humphreys. 

No original number.

Type Locality: Colon, [Colon], Colombia (Panama). 

Remarks: We know this Colon to be in Panama rather than Colombia because the title of the type publication refers to a 

new bat from the Isthmus of Panama.

Genus Vampyressa Thomas, 1900

Vampyressa elisabethae Tavares, Gardner, Ramirez-Chaves, 

and Velazco

Am. Mus. Novit. 3813: 19, 2 Oct 2014.

Holotype: USNM 319283. Skin and skull. Adult male. Collected 

13 Sep 1961 by V. J. Tipton. Original number 8120, not 

8006 as given in original description.

Type Locality: Rancho Mojica, upper Changena River, ±20 mi 

(32.2 km) SSW of Changuinola, 9o6′N, 82o34′W, 1,463 m, 

Provincia Bocas del Toro, Panama.

Vampyressa minuta G. S. Miller

Proc. U.S. Natl. Mus. 42: 25, 6 Mar 1912c.

= Vampyressa thyone Thomas, 1909. See Lim et al. (2003:25).

Holotype: USNM 173832. In alcohol, [skull removed]. Immature female. Collected May 1911 by A. Busck. No original 

number.

Type Locality: Cabima (La Cabima), [Panama], Panama. 

Genus Vampyrum Rafinesque, 1815

Vampyrus spectrum nelsoni Goldman

Proc. Biol. Soc. Wash. 30: 115, 23 May 1917.

= Vampyrum spectrum (Linnaeus, 1758). See G. S. Miller 

(1924:49). Considered monotypic by Simmons (2005:412).

Holotype: USNM 78127. In alcohol, skull removed. Adult male. 

Collected 14 May 1896 by E. W. Nelson. Original number 

[FC: Nelson and Goldman] 9579.

Type Locality: Coatzacoalcos, [FC: 75 ft (23 m)], Vera Cruz 

(Veracruz), Mexico. 

Family Mormoopidae Saussure, 1860

Genus Mormoops Leach, 1821

Mormoops intermedia G. S. Miller

Proc. Biol. Soc. Wash. 13: 160, 31 Oct 1900f.

= Mormoops megalophylla intermedia G. S. Miller, 1900. See 

Rehn (1902:170).

Holotype: USNM 102174. In alcohol. Adult female. Collected 

29 Apr 1900 by [MC: L. J. Guthrie]. No original number.

Type Locality: Cave at Hatto, on the north coast of Curacao, 

[Netherlands Antilles], West Indies. 

Remarks: USNM accession number 36524 contains a note from 

Guthrie about his bat collection in which he stated “. . . the 

majority were collected in the large cave on the North coast; 

but there are bats in the lot from every corner of the island . . .” A careful reading of the museum catalog reveals 

that this specimen came from a cave on the north coast, but 

not necessarily the cave at Hatto. 

Mormoops megalophylla senicula Rehn

Proc. Acad. Nat. Sci. Phila. 54: 169, 11 Jun 1902.

= Mormoops megalophylla megalophylla (Peters, 1864). See 

Smith (1972:115).

Holotype: USNM 84801. Skin and skull. Adult female. Collected 

3 Dec 1897 by E. A. Mearns. Original number [SL: 4273].

Type Locality: Fort Clark, Kinney County, Texas. 

Mormoops tumidiceps G. S. Miller

Proc. Acad. Nat. Sci. Phila. 54: 403, 11 Sep 1902d.

= Mormoops megalophylla tumidiceps G. S. Miller, 1902. See 

Goodwin and Greenhall (1961:225).

Holotype: USNM 186443. In alcohol, [skull removed]. Adult 

male. Collected in 1889 by W. Robinson. No original 

number.

Type Locality: Point Gourde Caves, Trinidad, [Trinidad and Tobago].

Remarks: Published as Merriam Collection number 5581. Cataloged USNM 186443 on 13 Oct 1911. 

Genus Pteronotus Gray, 1838

Chilonycteris mexicana G. S. Miller

Proc. Acad. Nat. Sci. Phila. 54: 401, 11 Sep 1902d.

= Pteronotus parnellii mexicanus (G. S. Miller, 1902). See Hall 

and Dalquest (1963:217).

Holotype: USNM 89277. Skin and skull. Adult male. Collected 

9 Jun 1897 by E. W. Nelson and E. A. Goldman. Original 

number 11132.

Type Locality: San Blas, [FC: altitude 75 ft (23 m)], Tepic 

(Nayarit), Mexico. 

Chilonycteris parnellii gonavensis Koopman

J. Mammal. 36: 110, 25 Feb 1955.

= Pteronotus parnellii gonavensis (Koopman, 1955). See Smith 

(1972:68).

Holotype: USNM 295009. Skull lacking incisors, mandible, 

and ear region. Age and sex not given. Collected [MC: Mar 

1929 by A. J. Poole. Original number 1A].

Type Locality: Cave near En Café, La Gonave Island (Ile de la 

Gonave), [Ouest], Republic of Haiti. 

Chilonycteris portoricensis G. S. Miller

Proc. Acad. Nat. Sci. Phila. 54: 400, 3 Sep 1902d.

= Pteronotus parnellii portoricensis (G. S. Miller, 1902). See 

Smith (1972:68).
6 6 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Holotype: USNM 102358. Skin and skull. Adult female. Collected 19 Mar 1900 by L. Stejneger and C. W. Richmond. 

Original number [SL: G].

Type Locality: Cave near Pueblo Viejo, Puerto Rico, W. I. (West 

Indies). 

Family Noctilionidae Gray, 1821

Genus Noctilio Linnaeus, 1766

Noctilio leporinus mexicanus Goldman

Proc. Biol. Soc. Wash. 28: 136, 29 Jun 1915b.

= Noctilio leporinus mastivus Vahl, 1797. See Davis (1973: 

869).

Holotype: USNM 126672. [Skin and skull]. Adult male. Collected 17 Apr 1903 by E. W. Nelson and E. A. Goldman. 

Original number 16318.

Type Locality: Papayo, Guerrero, Mexico. 

Family Thyropteridae Miller, 1907

Genus Thyroptera Spix, 1823

Thyroptera discifera abdita Wilson

Proc. Biol. Soc. Wash. 89: 307, 12 Oct 1976.

Holotype: USNM 51538. In alcohol, skull removed. Adult female. Collected 25 Jul 1892 by J. E. Mitchell. Original number [SL: 2].

Type Locality: 50 mi (80 km) E [SL: from] Bluefields, Escondido 

River, [Atlantico Sur], Nicaragua. 

Thyroptera discifera major G. S. Miller

J. Mammal. 12: 411, 11 Nov 1931b.

= Thyroptera discifera discifera (Lichtenstein and Peters, 1855). 

See Wilson (1976:306).

Holotype: USNM 105419. Skin and skull. Adult female. Collected 17 Jul 1900 by W. Robinson and M. W. Lyon, Jr. 

Original number [FC: 122].

Type Locality: San Julian, [Vargas], Venezuela.

Remarks: Originally W. Robinson collection number 1511. 

Family Natalidae Gray, 1866

Genus Chilonatalus Miller, 1898

Chilonatalus macer G. S. Miller

Proc. Biol. Soc. Wash. 27: 225, 29 Dec 1914a.

See Tejedor (2011:26) for current use.

Holotype: USNM 113724. In alcohol, [skull removed]. Adult female. Collected 6 Feb 1902 by W. Palmer. Original number 

699.

Type Locality: Baracoa, [Silva-Taboada (1979:220): in a cave 

identified as la Cueva de la Majana], [Tejedor (2011:26): 

Guantanamo], Cuba. 

Chilonatalus tumidifrons G. S. Miller

Proc. Biol. Soc. Wash. 16: 119, 30 Sep 1903d.

Holotype: USNM 122024. In alcohol, [skull removed]. Adult 

male. Collected 12 Jul 1903 by J. H. Riley. Original number 

157.

Type Locality: In a cave near Sandy Point, southwest corner 

of Watlings Island (San Salvador Island), Bahama Islands, 

(Bahamas). 

Natalus (Chilonatalus) brevimanus G. S. Miller

Proc. Acad. Nat. Sci. Phila. 50: 328, 12 Jul 1898c.

= Chilonatalus micropus (Dobson, 1880). See Tejedor (2011:30) 

who placed brevamanus in what he considered a monotypic 

micropus.

Holotype: USNM 15835. In alcohol. Adult male. Collector unknown. Presented by C. B. Cory and cataloged [MC: 25 

May 1887].

Type Locality: Old Providence Island (Isla de Providencia), Caribbean Sea, [San Andres y Providencia, Colombia]. 

Genus Natalus Gray, 1838

Natalus dominicensis Shamel

Proc. Biol. Soc. Wash. 41: 67, 16 Mar 1928.

= Natalus stramineus Gray, 1838. See Goodwin (1959:5) who 

placed dominicensis in stramineus as a valid subspecies and 

Tejedor (2011:73) who considered stramineus to be monotypic.

Holotype: USNM 113605. In alcohol, [skull removed]. Adult 

male. Collected May–Jul [SL: 18 Jul] 1901 by H. S. Branch. 

No original number.

Type Locality: Island of Dominica, [MC: West Indies]. 

Natalus major G. S. Miller

Proc. Acad. Nat. Sci. Phila. 54: 398, 3 Sep 1902d.

Holotype: USNM 101395. In alcohol, [skull removed]. Adult male. 

Collected [Poole and Schantz (1942): 1869–1871], by W. M. 

Gabb. No original number. [MC: Received from the Academy 

of Natural Sciences of Philadelphia (Academy of Natural Sciences of Drexel University); cataloged 3 Nov 1899].

Type Locality: [MC: From cave] near Savaneta (Sabaneta), San 

(Santo) Domingo, [Santiago, Dominican Republic], West 

Indies. 

Natalus mexicanus G. S. Miller

Proc. Acad. Nat. Sci. Phila. 54: 399, 3 Sep 1902d.

See Tejedor (2011:60) for current use.

Holotype: USNM 96496. In alcohol, [skull removed]. Adult female. Collected Jul or Aug 1897 by J. F. Abbott. No original 

number. [SL: 1953-X].

Type Locality: Santa Anita, Lower California (Baja California 

Sur), Mexico. 

Natalus stramineus natalensis Goodwin

Am. Mus. Novit. 1977: 5, 22 Dec 1959.
NUMBER 644 • 6 7

= Natalus macrourus (Gervais, 1856). See Tejedor (2011:39) 

who placed natalensis in espiritosantensis and Garbino and 

Tejedor (2013:237) who determined that espiritosantensis 

was a junior synonym of macrourus.

Holotype: USNM 242830. Skin and skull [in alcohol, skull removed]. Adult female. Collected Apr 1921 by E. C. Green. 

No original number. [SL: 22838-X].

Type Locality: Natal, Rio Grande do Norte, Brazil. 

Natalus tumidirostris G. S. Miller

Proc. Biol. Soc. Wash. 13: 160, 31 Oct 1900f.

See Tejedor (2011:79) who considered tumidirostris to be monotypic.

Holotype: USNM 102106. In alcohol, [skull removed]. Adult 

male. Collected 1 May 1900 by [MC: L. J. Guthrie]. No 

original number.

Type Locality: In a cave at Hatto, [MC: a country estate], north 

side of Curacao, [Netherlands Antilles], West Indies. 

Family Molossidae Gervais, 1856

Genus Chaerephon Dobson, 1874

Chaerephon luzonus Hollister

Proc. U.S. Natl. Mus. 46: 312, 31 Dec 1913a.

= Chaerephon plicatus luzonus Hollister, 1913. See Hill 

(1961:52) who placed luzonus as a subspecies of Tadarida 

plicata and Freeman (1981:152) who placed it in 

Chaerephon.

Holotype: USNM 144881. In alcohol, skull removed. Adult male 

[A. L. Gardner in litt.: but is obviously a female]. Collected 

on unknown date by E. A. Mearns. No original number. 

[MC: Cataloged 13 Nov 1907].

Type Locality: Cagayan, Cagayan Province, Northern Luzon, 

[Philippines]. 

Chaerephon pumilus naivashae Hollister

Smithson. Misc. Collect. 66(1): 4, 10 Feb 1916c.

= Chaerephon pumilus (Cretzschmar, 1830–1831). Subspecies 

not recognized by Simmons (2005:435).

Holotype: USNM 166658. In alcohol, skull removed. [Poole and 

Schantz (1942:120): Adult] male. Collected 7 Aug 1909 by 

J. A. Loring. Original number 6955.

Type Locality: Naivasha Station, [Rift Valley], British East Africa 

(Kenya). 

Nyctinomus jobensis G. S. Miller

Proc. Biol. Soc. Wash. 15: 246, 16 Dec 1902e.

= Chaerephon jobensis (G. S. Miller, 1902). See G. S. Miller 

(1907:245).

Holotype: USNM 18545/38035. Skin and skull [in alcohol, skull 

removed]. Adult male [Lyon and Osgood (1909:280): female]. Collected on unknown date by O. Beccari. Original 

number [MC: 33. Cataloged 15 Oct 1902].

Type Locality: Ansus, Island of Jobie (Yapen), northwestern New 

Guinea, [Papua, Indonesia]. 

Nyctinomus pusillus G. S. Miller

Proc. Biol. Soc. Wash. 15: 245, 16 Dec 1902e.

= Chaerephon pusillus (G. S. Miller, 1902). See G. S. Miller 

(1907:245) who placed pusillus in Chaerephon and Goodman and Ratrimomanarivo (2007:398) for current use.

Holotype: USNM 20097, not 20091 as in original description/37852. In alcohol, [skull removed]. Adult female. 

Collected on unknown date by W. L. Abbott. No original 

number. [MC: Alcoholic cataloged 30 Jun 1893; skull 12 

Feb 1900].

Type Locality: Aldabra Island, [Aldabra Atoll, Seychelles], Indian Ocean. 

Genus Cheiromeles Horsfield, 1824

Cheiromeles parvidens Miller and Hollister

Proc. Biol. Soc. Wash. 43: 100, 30 Jun 1921.

Holotype: USNM 219350. Skin and skull. Adult female. Collected 20 Feb 1918 by H. C. Raven. Original number 

3547.

Type Locality: Pinedapa, [Sulawesi Tengah], Middle Celebes (Sulawesi), [Indonesia]. 

Genus Eumops Miller, 1906

Eumops amazonicus Handley

Proc. Biol. Soc. Wash. 68: 177, 31 Dec 1955a.

= Eumops hansae Sanborn, 1932. See Gardner et al. (1970:727).

Holotype: USNM 123827. Skin and skull. Young adult, sex not 

given. Collected in 1899 by “Bicego.” No original number.

Type Locality: Manaos (Manaus), Amazonas, Brazil.

Remarks: Museu Paulista number 365a. 

Eumops hansae Sanborn

J. Mammal. 13: 356, 2 Nov 1932.

Holotype: USNM 200993. [Skin and skull]. Adult male. Collected in 1901 by H. Ehrhardt. No original number.

Type Locality: Colonia Hansa, near Joinville, Santa Catherina 

(Catarina), Brazil.

Remarks: Museu Paulista number 1339. 

Molossus californicus Merriam

N. Am. Fauna 4: 31, 8 Oct 1890b.

= Eumops perotis californicus (Merriam, 1890). See Sanborn 

(1932:351).

Holotype: USNM 186448. [In alcohol]. Adult female. Collected 

14 Dec 1889 by E. C. Thurber. No original number. Merriam collection number 5736.

Type Locality: Alhambra, Los Angeles County, California.

Remarks: Published as Merriam collection number 5736. Cataloged as USNM 186448 on 13 Oct 1911. 
6 8 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Nyctinomus orthotis H. Allen

Proc. Am. Philos. Soc. 26: 4, 3 Dec 1889a. Preprint of Proc. Am. 

Philos. Soc. 26: 561, 18 Dec 1889b.

= Eumops glaucinus (Wagner, 1843). See Sanborn (1932:353). 

Considered monotypic by Eger (2008:412).

Holotype: USNM 9397/37447. Skin [in alcohol, skull removed]. 

Age and sex not given. Collected probably Aug 1868 by 

W. T. March. No original number.

Type Locality: Spanishtown (Spanish Town), [St. Catherine, 

Jamaica].

Remarks: Published as USNM 9397 to which skull number 

37447 belongs. 

Genus Molossus E. Geoffroy, 1805

Molossus debilis G. S. Miller

Proc. U.S. Natl. Mus. 46: 90, 23 Aug 1913f.

= Molossus molossus debilis G. S. Miller, 1913. See Jones and 

Phillips (1970:143).

Holotype: USNM 110935. In alcohol, [skull removed]. Adult 

female. Collected in 1901 by W. H. Alexander. No original 

number.

Type Locality: St. Kitts, [Saint Kitts and Nevis], Lesser Antilles. 

Molossus fortis G. S. Miller

Proc. U.S. Natl. Mus. 46: 89, 23 Aug 1913f.

= Molossus molossus fortis G. S. Miller, 1913. See Varona 

(1974:42).

Holotype: USNM 102319. In alcohol, [skull removed]. Adult 

male. Collected 5 Mar 1900 by L. Stejneger and C. W. Richmond. No original number.

Type Locality: Luquillo, Porto (Puerto) Rico. 

Molossus nigricans G. S. Miller

Proc. Acad. Nat. Sci. Phila. 54: 395, 3 Sep 1902d.

= Molossus rufus E. Geoffroy, 1805. See Dolan (1989:44). Considered monotypic by Eger (2008:425).

Holotype: USNM 90941. Skin and skull. Adult male. Collected 

2 Aug 1897 by E. W. Nelson and E. A. Goldman. Original 

number 11433.

Type Locality: Acaponeta, [FC: altitude 150 ft (46 m)], Tepic 

(Nayarit), Mexico. 

Molossus pretiosus G. S. Miller

Proc. Acad. Nat. Sci. Phila. 54: 396, 3 Sep 1902d.

Holotype: USNM 102761. Skin and skull. Adult male. Collected 

13 Jul 1900 by W. Robinson and M. W. Lyon, Jr. Original 

number 106.

Type Locality: La Guaira, [Vargas], Venezuela. 

Molossus pygmaeus G. S. Miller

Proc. Biol. Soc. Wash. 13: 162, 31 Oct 1900f.

= Molossus molossus pygmaeus G. S. Miller, 1900. See Simmons 

(2005:441).

Holotype: USNM 102104. In alcohol, [skull removed]. Adult 

female. Collected 16 Jan 1900 by [MC: L. J. Guthrie]. No 

original number.

Type Locality: In an attic near Willemstad, Curacao, [Netherlands Antilles], West Indies. 

Genus Mormopterus Peters, 1865

Mormopterus phrudus Handley

Proc. Biol. Soc. Wash. 69: 197, 31 Dec 1956.

Holotype: USNM 194449. Skin and skull. Adult male. Collected 

26 Jun 1915 by E. Heller. Original number 361.

Type Locality: San Miguel Bridge, altitude 6,000 ft (1,829 m), 

Urubamba River, Machu Picchu, Cuzco (Cusco), Peru. 

Tadarida loriae cobourgiana Johnson

Proc. Biol. Soc. Wash. 72: 185, 4 Nov 1959.

= Mormopterus loriae cobourgianus (Johnson, 1959). See Flannery (1995:479).

Holotype: USNM 284243. Skin and skull. Adult female. Collected 25 Sep 1948 by D. H. Johnson. Original number 

5846.

Type Locality: Black Rock Point on north shore of Van Diemen 

Gulf, 15 mi (24 km) southeast of Cape Don lighthouse, Cobourg Peninsula, Northern Territory, Australia, lat. 11o26′S, 

long. 131o56′E. 

Genus Nyctinomops Miller, 1902

Dysopes aurispinosus Peale

U.S. Explor. Exped. 8: 21, 1848.

= Nyctinomops aurispinosus (Peale, 1848). See G. S. Miller 

(1902d:393).

Holotype: USNM 3726. [Skin only in alcohol]. [Lyon and Osgood (1909:279): Adult male]. Collected by the U.S. Exploring Expedition on 18 Nov 1838-1842. No original 

number.

Type Locality: About 100 mi (161 km) south of Cape St. Roque 

(Cabo Sao Roque), Brazil.

Remarks: The holotype was not designated by number in the 

original description. However, Peale (1848:21) speaks of a 

single specimen that flew aboard the U.S. ship Peacock. The 

specimen listed above was first cataloged 20 Dec 1859 as 

USNM 3726 and then a second time on 31 Oct 1861 when it 

was assigned USNM 5476. In both instances the entry reads 

Molossus aurispinosus Brazil, U.S. Exploring Expedition. 

There is no question this is the specimen Peale (1848:21) 

described and is the holotype by monotypy (ICZN, 1999: 

Article 73.1.2). 

Nyctinomops yucatanicus G. S. Miller

Proc. Acad. Nat. Sci. Phila. 54: 393, 3 Sep 1902d.

= Nyctinomops laticaudatus yucatanicus G. S. Miller, 1902. See 

Freeman (1981:163).
NUMBER 644 • 6 9

Holotype: USNM 108166. Skin and skull. Adult female. Collected 9 Feb 1901 by E. W. Nelson and E. A. Goldman. 

Original number 14521.

Type Locality: Chichen Itza, Yucatan, [SL: Mexico]. 

Nyctinomus europs H. Allen

Proc. Am. Philos. Soc. 26: 1, 3 Dec 1889a. Preprint of Proc. Am. 

Philos. Soc. 26: 558, 18 Dec 1889b.

= Nyctinomops laticaudatus europs (H. Allen, 1889). See AvilaFlores et al. (2002:1).

Lectotype: USNM 101498 as designated by Shamel (1931b:14). 

In alcohol, [skull removed]. Adult female. Collected on unknown date by Prof. [MC: C. F.] Harte [MC: Hartt]. No 

original number. [MC: Cataloged 5 Dec 1899].

Type Locality: Brazil. Restricted by Cabrera (1958:121) to Corumba, Mato Grosso do Sul, Brazil. 

Nyctinomus femorosaccus Merriam

N. Am. Fauna 2: 23, 30 Oct 1889b.

= Nyctinomops femorosaccus (Merriam, 1889). See G. S. Miller 

(1902d:393).

Holotype: USNM 186447. In alcohol. Adult male. Collected 27 

Mar 1885 by F. Stephens. Original number [MCC: 252]. 

Merriam Collection number 2276.

Type Locality: Agua Caliente (Palm Springs), [MCC: west end] 

Colorado Desert, [Riverside County], California.

Remarks: Published as Merriam Collection number 2276. Cataloged USNM 186447 on 13 Oct 1911. 

Nyctinomus macrotis nevadensis H. Allen

Bull. U.S. Natl. Mus. 43: 171, 14 Mar 1894.

= Nyctinomops macrotis (Gray, 1840). See G. S. Miller 

(1902d:393). N. macrotis was considered to be monotypic 

by Simmons (2005:447).

Lectotype: USNM 15178/36569 as designated by J. A. Allen 

(1894:326, footnote). In alcohol, skull removed. Immature 

female. Collected [MC: 11 Aug 1885] by J. Mullen. No 

original number.

Type Locality: California and Nevada, but restricted by selection 

of the lectotype to California.

Remarks: The original description was based on two specimens. J. A. Allen (1894:326, footnote) selected USNM 

15178/36569 to be the type = lectotype (ICZN, 1999: Article 74.5). 

Genus Promops Gervais, 1856

Promops pamana G. S. Miller

Proc. Biol. Soc. Wash. 26: 33, 8 Feb 1913a.

= Promops nasutus (Spix, 1823). See Goodwin and Greenhall 

(1962:11) who placed pamana in nasutus. Eger (2008:436) 

considered P. nasutus to be monotypic.

Holotype: USNM 105528. Skin only [in alcohol]. Adult male. 

Collected 22 Mar 1901 by J. B. Steere. No original number.

Type Locality: Hyutanaham (Huitanaa), Upper Purus River, 

[Amazonas], Brazil. 

Genus Tadarida Rafinesque, 1814

Nycticea cynocephala LeConte

In McMurtrie, The Animal Kingdom 1: 432, 1831.

= Tadarida brasiliensis cynocephala (LeConte, 1831). See 

Schwartz (1955:108).

Holotype: USNM 4742. [Skin with skull inside]. Age and sex unknown. Collected on unknown date by [MC: Maj. LeConte]. 

No original number. [MC: Cataloged 13 Apr 1861].

Type Locality: Unknown. Long presumed to be the LeConte 

plantation (Woodmanston), near Riceboro, Liberty County, 

Georgia.

Remarks: A type was not designated by number in the original description. USNM 4742 has long been considered the 

type, Lyon and Osgood (1909:279) and Poole and Schantz 

(1942:123). It is a LeConte specimen, bears two old labels 

that have cynocephalus written on them, and is entered in 

the museum catalog as such. 

Nyctinomus antillularum G. S. Miller

Proc. Acad. Nat. Sci. Phila. 54: 398, 3 Sep 1902d.

= Tadarida brasiliensis antillularum (G. S. Miller, 1902). See 

Schwartz (1955:108).

Holotype: USNM 113188. In alcohol, [skull removed]. Adult 

male. Collected 5 Aug 1901 by H. S. Branch [MC: C. E. 

Ashcraft, Jr.]. No original number.

Type Locality: Roseau, [Saint George], Dominica, West Indies. 

Nyctinomus mohavensis Merriam

N. Am. Fauna 2: 25, 30 Oct 1889c.

= Tadarida brasiliensis mexicana (Saussure, 1860). See G. S. 

Miller (1924:86) who placed mohavensis in mexicana and 

Schwartz (1955:108).

Holotype: USNM 186446. In alcohol. Adult male. Collected 8 

Mar 1889 by V. Bailey. Original number [MC: 643]. Merriam collection number 5418.

Type Locality: [FC: At a pond 3 mi (5 km) south of] Fort Mohave, [Mohave County], Arizona.

Remarks: Published as Merriam collection number 5418. Cataloged as USNM 186446 on 13 Oct 1911. 

Tadarida constanzae Shamel

Proc. U.S. Natl. Mus. 78(19): 10, 6 May 1931b.

= Tadarida brasiliensis constanzae Shamel, 1931. See Schwartz 

(1955:108)

Holotype: USNM 217205. In alcohol, [skull removed]. Adult male. 

Collected 26 Sep 1916 by W. L. Abbott. No original number.

Type Locality: Constanza, Dominican Republic. 

Tadarida intermedia Shamel

Proc. U.S. Natl. Mus. 78(19): 7, 6 May 1931b.
7 0 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

= Tadarida brasiliensis intermedia Shamel, 1931. See Schwartz 

(1955:108).

Holotype: USNM 78488. In alcohol, [skull removed]. Adult 

male. Collected 11 Dec 1895 by E. W. Nelson and E. A. 

Goldman. Original number [SL: 8773].

Type Locality: Valley of Comitan, [FC: altitude 5,700 ft 

(1,737 m)], Chiapas, Mexico. 

Family Vespertilionidae Gray, 1821

Genus Antrozous H. Allen, 1862

Antrozous minor G. S. Miller

Proc. Acad. Nat. Sci. Phila. 54: 389, 3 Sep 1902d.

=Antrozous pallidus minor G. S. Miller, 1902. See Goldman 

(1951:356).

Holotype: USNM 79096. Skin and skull. Adult, sex not given. Collected 20 Sep 1895 by J. E. McLellan. No original number.

Type Locality: Comondu, Lower California (Baja California 

Sur), Mexico. 

Antrozous pallidus cantwelli Bailey

N. Am. Fauna 55: 391, 29 Aug 1936.

= Antrozous pallidus pallidus (LeConte, 1855). See Martin and 

Schmidly (1982:28).

Holotype: USNM 232362. Skin and skull. Adult female. Collected 28 May 1919, not 1918 as in original description, by 

G. G. Cantwell. Original number 1211.

Type Locality: Rogersburg, [MC: mouth, Grande Ronde River, 

700 ft (213 m)], Asotin County, Washington. 

Antrozous pallidus obscurus Baker

Southwest. Nat. 12: 329, 31 Oct 1967.

Holotype: USNM 395263. Skin and skull. Young adult male. 

Collected 31 Jul 1966 by P. Dalby and F. Welling; prepared 

by R. H. Baker. Original number 5338.

Type Locality: Acuna Ranch, 800 m, approximately 40 km north 

of Gonzales in the Sierra de Tamaulipas, Tamaulipas, Mexico.

Remarks: Published as 11362, the Museum, Michigan State University. Cataloged USNM 395263 on 10 Dec 1968. 

Antrozous pallidus pacificus Merriam

Proc. Biol. Soc. Wash. 11: 180, 1 Jul 1897e.

Holotype: USNM 29815. In alcohol. [Lyon and Osgood 

(1909:278): Adult female, not male as in original description]. Collected 28 Jun 1891 by C. H. Merriam. Original 

number 208.

Type Locality: Old Fort Tejon, Canada del las Uvas (Grapevine 

Canyon), [Tehachapi Mountains, Kern County], California. 

V[espertilio]. pallidus LeConte

Proc. Acad. Nat. Sci. Phila. 7: 437, reported favorably for publication 25 Dec 1855. Receipt acknowledged in 1856 by the 

Smithsonian Institution.

= Antrozous pallidus pallidus (LeConte, 1855). See H. Allen 

(1864:68).

Holotype: USNM 152/1134. Skin and skull. Age and sex unknown. Collected in 1851 by J. H. Clark. No original number.

Type Locality: [SL: El Paso, El Paso County, Texas].

Remarks: A type was not designated by number in the original 

description. H. Allen (1864:69) considered USNM 152/1134 

the type and Baird (1859:5) stated that the one described by 

LeConte was taken at El Paso. There is no indication that 

LeConte had more than the one specimen, so this is the holotype fixed by monotypy (ICZN, 1999: Article 73.1.2). The 

skull of this specimen, USNM 1134, was reported lost by 

both Lyon and Osgood (1909:278) and Poole and Schantz 

(1942:145); it was found and reunited with the skin on 7 

Mar 1980. The skin of this specimen was cataloged a second 

time as USNM 5467. It now bears both numbers and is on 

the electronic database as USNM 152. LeConte (1855:437) 

gave California as habitat; Baird (1859:5) clearly stated that 

LeConte’s specimen came from El Paso. 

Genus Corynorhinus H. Allen, 1865

Corynorhinus macrotis pallescens G. S. Miller

N. Am. Fauna 13: 52, 16 Oct 1897a.

= Corynorhinus townsendii pallescens G. S. Miller, 1897. See 

Handley (1955b:147).

Holotype: USNM 66534, not 65534 as in original description. 

Skin [and skull]. Adult female. Collected 3 Aug 1894 by 

A. K. Fisher. Original number 1715.

Type Locality: Keam (Keams) Canyon, [FC: 80 mi (129 km) 

north of Holbrook], Navajo County, Arizona. 

Corynorhinus megalotis mexicanus G. M. Allen

Bull. Mus. Comp. Zool. 60: 347, 20 Apr 1916.

= Corynorhinus mexicanus G. M. Allen, 1916. See Handley 

(1955b:148).

Holotype: USNM 98285. Skin and skull. Adult female. Collected 25 Aug 1899 by E. W. Nelson and E. A. Goldman. 

Original number [SL: 13955].

Type Locality: Near Pacheco, Chihuahua, Mexico. 

Corynorhinus townsendii australis Handley

J. Wash. Acad. Sci. 45: 147, 27 May 1955b.

Holotype: USNM 297265. In alcohol, skull removed. Adult female. Collected 20 Dec 1952 by A. Malaga-Alba. Original 

number 1053.

Type Locality: [MC: Boca Mine], 2 mi (3 km) W Jacala, 5,500 ft 

(1,676 m), Hidalgo, Mexico. 

Corynorhinus townsendii ingens Handley

J. Wash. Acad. Sci. 45: 148, 27 May 1955b.

Holotype: USNM 296767. Skin and skull. Adult male. Collected 

4 Dec 1950 by J. A. Sealander. No original number. [SL: University of Arkansas, Zoology Department number M50-14].
NUMBER 644 • 7 1

Type Locality: Hewlitt Cave, 12 mi (19 km) W Fayetteville, 

Washington County, Arkansas. 

Corynorhinus townsendii virginianus Handley

J. Wash. Acad. Sci. 45: 148, 27 May 1955b.

Holotype: USNM 269163. Skin and skull. Adult male. Collected 

12 Nov 1939 by W. J. Stephenson. No original number.

Type Locality: Schoolhouse Cave, 4.4 mi (7 km) NE Riverton, 

[SL: 22 mi (35 km) SE Petersburg], 2,205 ft (672 m), Pendleton County, West Virginia. 

Genus Eptesicus Rafinesque, 1820

Eptesicus hispaniolae G. S. Miller

Proc. Biol. Soc. Wash. 31: 39, 16 May 1918.

= Eptesicus fuscus hispaniolae G. S. Miller, 1918. See Shamel 

(1945:108).

Holotype: USNM 217207. In alcohol, [skull removed]. Adult 

male. Collected 22 Sep 1916 by W. L. Abbott. No original 

number.

Type Locality: Constanza, [SL: 5,000 ft (1,524 m)], Santo Domingo (Dominican Republic). 

Eptesicus lynni Shamel

Proc. Biol. Soc. Wash. 58: 107, 18 Jul 1945.

= Eptesicus fuscus lynni Shamel, 1945. See Simmons (2005:454).

Holotype: USNM 258488. In alcohol, skull removed. Adult female. Collected 11 Jul 1932 by W. G. Lynn. Original number B25.

Type Locality: A cave [MC: 3 mi (5 km)] east of Montego Bay, 

[MC: a few hundred yards off the main road which encircles 

the island], [Saint James], Jamaica. 

Eptesicus pallidus Young

Proc. Acad. Nat. Sci. Phila. 60: 408, 14 Oct 1908.

= Eptesicus fuscus pallidus Young, 1908. See G. S. Miller 

(1912b:62).

Holotype: USNM 142526. [Skin and skull]. Adult female. Collected 22 Jul 1903 by R. T. Young. Original number [SL: 

1315].

Type Locality: Boulder, [Boulder County], Colorado. 

Eptesicus serotinus pallens G. S. Miller

Proc. Biol. Soc. Wash. 24: 53, 24 Feb 1911a.

Holotype: USNM 155156. Skin and skull. Adult male. Collected 

4 Aug 1909 by A. de C. Sowerby. Original number 246.

Type Locality: Ch’eng-yuan-hsien (Chengyang Xiang?), [FC: 

4,150 ft (1,265 m)], 70 mi (113 km) west of Ch’ing-yang-fu 

(Qingyang), Kansu (Ningxia), China. 

Eptesicus wetmorei Jackson

Proc. Biol. Soc. Wash. 29: 37, 24 Feb 1916.

= Eptesicus fuscus wetmorei Jackson, 1916. See Shamel 

(1945:108).

Holotype: USNM 179142. In alcohol, skull removed. Young 

adult male. Collected 29 May 1912 by A. Wetmore. Original number 900.

Type Locality: Maricao, altitude 1,375 ft (419 m), Puerto Rico. 

S[cotophilus]. miradorensis H. Allen

Proc. Acad. Nat. Sci. Phila. 18: 287, ordered published 28 Aug 

1866.

= Eptesicus fuscus miradorensis (H. Allen, 1866). See G. S. 

Miller (1912b:62).

Holotype: USNM 5411; ANSP 1842. In alcohol. Age and sex 

not given. Collected on unknown date by Doctor Sartorius. 

No original number.

Type Locality: Mirador, [Veracruz], Mexico.

Remarks: A type was not designated by number in the original 

description. Allen had only one specimen from the above locality and collected by Sartorius, and he said it was in the 

Smithsonian. USNM 5411 is the only specimen in the collection that matches the data given by Allen and therefore is the 

holotype by monotypy (ICZN, 1999; Article 73. 1. 2). This 

specimen has been missing from the Smithsonian collection at 

least since Mar 1902. R. D. Fisher confirmed its presence in 

the ANSP collection on 19 Jun 1996 bearing ANSP number 

1842, even though it is not listed by Koopman (1976). 

Vespertilio fuscus bahamensis G. S. Miller

N. Am. Fauna 13: 101, 16 Oct 1897a.

= Eptesicus fuscus bahamensis (G. S. Miller, 1897). See Shamel 

(1945:108).

Holotype: USNM 76537. In alcohol. Adult male. Collected 

Spring 1894 by C. J. Maynard. No original number.

Type Locality: Nassau, New Providence [Island], Bahamas. 

Genus Histiotus Gervais, 1856

Histiotus humboldti Handley

Proc. Biol. Soc. Wash. 109: 2, 16 Apr 1996.

Holotype: USNM 370968. Skin, skull, and [partial] skeleton. 

Adult lactating female. Collected 24 Jul 1965 by C. O. 

Handley, Jr. Original number SVP 92.

Type Locality: Los Venados, 4 km NNW Caracas [SL: 5 mi (8 

km) N of Caracas], 10o32′N, 66o54′W, 1,498 m, Distrito 

Federal, Venezuela. 

Genus Hypsugo Kolenati, 1856

Pipistrellus curtatus G. S. Miller

Proc. Biol. Soc. Wash. 24: 25, 24 Feb 1911f.

= Hypsugo macrotis (Temminck, 1840). See Simmons (2005:491).

Holotype: USNM 141019. In alcohol, [skull removed]. Adult 

female. Collected 1 Dec 1904 by W. L. Abbott. Original 

number 3785.

Type Locality: Engano Island (Pulau Enggano), [Bengkulu], Sumatra, [Indonesia]. 
7 2 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Genus Kerivoula Gray, 1842

Kerivoula bombifrons Lyon

Proc. U.S. Natl. Mus. 40: 134, 25 Apr 1911b.

= Kerivoula pellucida (Waterhouse, 1845). See Hill (1966:539).

Holotype: USNM 145616. In alcohol, skull removed. Adult 

male. Collected 15 Aug 1907 by W. L. Abbott. Original 

number 5497.

Type Locality: Along Matan River (Sungai Matan), [Kalimantan 

Barat], Borneo (Kalimantan), [Indonesia]. 

Kerivoula depressa G. S. Miller

Proc. Biol. Soc. Wash. 19: 64, 1 May 1906d.

= Kerivoula hardwickii (Horsfield, 1824). See Hill (1966:543) 

who placed depressa as a subspecies of hardwickii. Simmons (2005:526) considered hardwickii to be monotypic.

Holotype: USNM 18533/38194. In alcohol, [skull removed]. 

Adult female. Collected on unknown date in 1890 by 

L. Fea. Original number [MC: 29. Skin cataloged 2 May 

1890; skull 4 Feb 1904].

Type Locality: Biapo, Carin (Karin) Hills, northeast Tounghoo 

(Toungoo), [Kayin ?], southern Burma, India. 

Kerivoula engana G. S. Miller

Proc. U.S. Natl. Mus. 30: 825, 4 Jun 1906e.

= Kerivoula hardwickii (Horsfield, 1824). See Hill (1966:543) 

who placed engana as a subspecies of hardwickii. Simmons 

(2005:526) considered hardwickii to be monotypic.

Holotype: USNM 141020. In alcohol, [skull removed]. Adult 

male. Collected 3 Nov 1904 by W. L. Abbott. Original number 3766.

Type Locality: Pulo (Pulau) Dua, Engano (Pulau Enggano), 

[Bengkulu, Sumatra, Indonesia]. 

Kerivoula minuta G. S. Miller

Proc. Acad. Nat. Sci. Phila. 50: 321, 12 Jul 1898b.

Holotype: USNM 83547. In alcohol, [skull removed]. Adult 

male. Collected Sep 1896 by W. L. Abbott. No original 

number.

Type Locality: [MC: Lay Song Hong], Trong (Trang), Siam 

(Thailand). 

Genus Laephotis Thomas, 1901

Laephotis botswanae Setzer

Proc. Biol. Soc. Wash. 84: 260, 4 Nov 1971.

Holotype: USNM 425349. Skin and skull. Adult female. Collected 24 May 1967 by S. W. Goussard. Original number 

691.

Type Locality: 50 mi (80 km) W, 12 mi (19 km) S Shakawe, 

[Schmidt et al. (2008:35): 18o32′S, 21o05′E], Botswana. 

Laephotis namibensis Setzer

Proc. Biol. Soc. Wash. 84: 259, 4 Nov 1971.

Holotype: USNM 342152. Skin and skull. Adult female. Collected 22 Nov 1963 by R. E. Cole. Original number 270.

Type Locality: Namib Research Station, Gobabeb, [Schmidt et al. 

(2008:35): 23o33′S, 15o22′E], South West Africa (Namibia). 

Genus Lasiurus Gray, 1831

Atalapha semota H. Allen

Proc. U.S. Natl. Mus. 13: 173, 9 Sep 1890c.

= Lasiurus cinereus semotus (H. Allen, 1890). See Sanborn and 

Crespo (1957:12).

Lectotype: USNM 15631/38983 as designated by Lyon and Osgood (1909:276). In alcohol, [skull removed]. Adult female. 

Collected on unknown date by V. Knudsen. No original 

number. [MC: Cataloged 4 Jan 1887].

Type Locality: Sandwich (Hawaiian) Islands.

Remarks: Published as USNM 15631, to which USNM 38983 

belongs. 

Atalapha teliotis H. Allen

Proc. Am. Philos. Soc. 29: 5, 10 Apr 1891.

= Lasiurus blossevillii teliotis (H. Allen, 1891). See G. S. Miller 

(1897a:110).

Holotype: USNM 84555. In alcohol, skull removed and lost. 

Age, sex, collector, and date collected unknown. [MC: Cataloged 27 Apr 1898].

Type Locality: Unknown. Lyon and Osgood (1909:277) stated 

probably California.

Remarks: A type was not designated by number in the original description. However, Allen based his description on a 

single specimen that was sent to him by J. G. Cooper of 

the California Academy of Sciences (Lyon and Osgood, 

1909:277). USNM 84555 bears an original label marked 

Atalapha teliotis, Cal. Acad. N. S. It is without doubt that 

single specimen and therefore is the holotype by monotypy 

(ICZN, 1999: Article 73.1.2). 

Dasypterus floridanus G. S. Miller

Proc. Acad. Nat. Sci. Phila. 54: 392, 3 Sep 1902d.

= Lasiurus intermedius floridanus (G. S. Miller, 1902). See Hall 

and Jones (1961:84).

Holotype: USNM 111379. Skin and skull. Adult female. Collected 28 Mar 1901 by E. A. Mearns. Original number 

5183.

Type Locality: Lake Kissimmee, [Osceola County], Florida. 

Lasiurus borealis ornatus Hall

Univ. Kans. Publ., Mus. Nat. Hist. 5: 226, 15 Dec 1951.

= Lasiurus blossevillii teliotis (H. Allen, 1891). See Baker et al. 

(1988:9).

Holotype: USNM 8492/37578. Skin and skull. Age and sex not 

given. Obtained from F. Sumichrast. No original number.

Type Locality: Penuela, [SL: near Cordova (Cordoba)], Veracruz, 

[Mexico]. 
NUMBER 644 • 7 3

Lasiurus castaneus Handley

Proc. U.S. Natl. Mus. 112: 468, 6 Oct 1960.

Holotype: USNM 310263. Skin, skull, [and skeleton]. Adult female. Collected 6 Mar 1959 by C. O. Handley, Jr. and B. R. 

Feinstein. Original number [SL: COH, Jr.] 5445.

Type Locality: Tacarcuna Village [SL: Camp], 3,200 ft (975 m), 

Rio Pucro, Darien, Panama. 

Lasiurus degelidus G. S. Miller

J. Mammal. 12: 410, 11 Nov 1931c.

Holotype: USNM 96188, not 96187 as in original description. 

In alcohol, [skull removed]. Adult female. Collector and 

date not given. Received from the Institute of Jamaica in 

1898. [SL: 1908-X].

Type Locality: Sutton’s (Suttons), District of Vere, [Clarendon], 

Jamaica. 

Lasiurus intermedius H. Allen

Proc. Acad. Nat. Sci. Phila. 14: 246, ordered published 29 Apr 

1862.

= Lasiurus intermedius intermedius H. Allen, 1862.

Lectotype: USNM 6136 as designated by Lyon and Osgood 

(1909:292). [In alcohol]. Age, sex, and collector not given. 

[MC: Received from L. B. Couch. Cataloged 7 Feb 1863].

Type Locality: Matamoros, Tamaulipas, [Mexico].

Remarks: Lyon and Osgood (1909:292), reiterated by Poole and 

Schantz (1942:571), gave their reasons for selecting USNM 

6136 as the type = lectotype. A note written in the Museum 

Catalog indicates that the specimen could not be found on 5 

Mar 1902; its whereabouts is still unknown. 

Lasiurus minor G. S. Miller

J. Mammal. 12: 410, 11 Nov 1931c.

Holotype: USNM 256505. Skull with complete set of molariform teeth, but lacking canines, incisors, and mandible. 

Age and sex not given. Collected 6–7 Mar 1931 by H. W. 

Krieger. No original number.

Type Locality: From owl pellets in the Voute l’Eglise, a cave situated near the Jacmel road a few kilometers north of Trouin, 

altitude 1,350 ft (411 m), [Ouest], Haiti. 

Myotis quebecensis Yourans

Le Natural. Can. 57: 65, Mar 1930.

= Lasiurus borealis (Muller, 1776). See Maheux (1930:185-186).

Holotype: USNM 255564. Skin with skull inside. Adult male. 

Collected in 1927 by E. Yourans. No original number.

Type Locality: Anse-a-Wolfe (Anse au Foulon), Quebec, Canada. 

Genus Miniopterus Bonaparte, 1837

Miniopterus natalensis arenarius Heller

Smithson. Misc. Collect. 60(12): 2, 4 Nov 1912a.

Holotype: USNM 181811. [Skin and skull]. Adult female. Collected 4 Oct 1911 by E. Heller. Original number 4413.

Type Locality: Guaso Nyuki, Northern Guaso Nyiro (Ewaso 

Ng’iro) River, British East Africa (Kenya). 

Miniopterus paululus Hollister

Proc. U.S. Natl. Mus. 46: 311, 31 Dec 1913a.

Holotype: USNM 105493. In alcohol, skull removed. Adult, sex 

not given. Collected Dec 1887 by J. B. Steere. No original 

number.

Type Locality: Guimaras Island, [Guimaras Province], [MC: 

Philippine Islands] (Philippines). 

Genus Murina Gray, 1842

Murina huttoni fuscus Sowerby

J. Mammal. 3: 46, 8 Feb 1922.

= Murina fusca Sowerby, 1922. See Honacki et al. (1982:183).

Holotype: USNM 199672. Skin and skull. [Poole and Schantz 

(1942:150): Adult] female. Collected 28 Sep 1914 by A. de 

C. Sowerby. Original number 702.

Type Locality: Forest of North Kirin, [SL: 500 ft (152 m)], 

Imienpo (Yimianpo) area, [SL: near Ninguta (Ning’an)], 

Manchuria [Heilongjiang, China]. 

Genus Myotis Kaup, 1829

Myotis abbotti Lyon

Proc. U.S. Natl. Mus. 52: 441, 30 Dec 1916.

= Myotis hasseltii abbotti Lyon, 1916. See Hill (1983:160).

Holotype: USNM 121611. In alcohol, skull removed. Adult female. Collected 3 Jan 1903 by W. L. Abbott. Original number 2191.

Type Locality: North Pagi Island (Pulau Pagai Utara), [Sumatera 

Barat, Sumatra, Indonesia]. 

Myotis adversus continentis Shamel

J. Mammal. 23: 323, 14 Aug 1942a.

= Myotis hasseltii continentis Shamel, 1942. See Hill (1983:163).

Holotype: USNM 251718. Skin and skull. Adult female. Collected 8 Aug 1926 by H. M. Smith. Original number [SL: 

1274].

Type Locality: Bangkok (Krung Thep), [Krung Thep], C. Siam 

(Thailand). 

Myotis altifrons Hollister

Smithson. Misc. Collect. 56(26): 3, 5 Dec 1911b.

= Myotis volans longicrus (True, 1886). See Miller and Allen 

(1928:140).

Holotype: USNM 174133. Skin and skull. Adult male. Collected 

5 Sep 1911 by J. H. Riley. Original number 2317.

Type Locality: Henry House, altitude 3,350 ft (1,021 m), Alberta, [Canada]. 

Myotis austroriparius mumfordi Rice

Q. J. Fla. Acad. Sci. 13: 67, 17 May 1955.
7 4 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

= Myotis austroriparius (Rhoads, 1897). Considered monotypic 

by LaVal (1970:50).

Holotype: USNM 293800. Skin and skull. Adult male. Collected 

11 Jan 1949 by R. Mumford. Original number [SL: 9].

Type Locality: Bronson Cave, Spring Mill State Park, 3 mi (5 km) 

east of Mitchell, Lawrence County, Indiana. 

Myotis baileyi Hollister

Proc. Biol. Soc. Wash. 22: 44, 10 Mar 1909.

= Myotis occultus Hollister, 1909. See Miller and Allen (1928:97).

Holotype: USNM 125787. [Skin and skull]. Adult female in alcohol [FC: made into skin 7 Nov 1910]. Collected 14 Sep 

1902 by N. Hollister. Original number 195.

Type Locality: Base of White Mountains, altitude 7,500 ft 

(2,286 m), near Ruidoso, [Lincoln County], New Mexico. 

Myotis californicus caurinus G. S. Miller

N. Am. Fauna 13: 72, 16 Oct 1897a.

Holotype: USNM 72219. In alcohol, [skull removed]. Adult 

male. Collected in 1895 by J. H. Keen. No original number.

Type Locality: [SL: Massett], [Graham Island], Queen Charlotte 

Islands, British Columbia, [Canada]. 

Myotis californicus pallidus Stephens 

Proc. Biol. Soc. Wash. 13: 153, 13 Jun 1900. Junior secondary 

homonym of Kerivoula pallida Blyth, 1863. Replaced with 

Myotis californicus stephensi by Dalquest (1946:67).

Holotype: USNM 99829. [Skin and skull]. [Poole and Schantz 

(1942:151): Adult] male. Collected 1 Apr 1895 by F. Stephens. Original number 2498.

Type Locality: Vallecito, [SL: altitude 1,600 ft (488 m)], San 

Diego County, California. 

Myotis capitaneus Nelson and Goldman

Proc. Biol. Soc. Wash. 22: 28, 10 Mar 1909.

= Myotis volans volans (H. Allen, 1866). See Miller and Allen 

(1928:139).

Holotype: USNM 146046. [Skin and skull]. Adult male. Collected 12 Nov 1905 by E. W. Nelson and E. A. Goldman. 

Original number 18526.

Type Locality: San Jorge, altitude 100 ft (30 m) [FC: 50 ft 

(15 m)], 30 mi (48 km) southwest of Comondu, Lower California (Baja California Sur), Mexico. 

Myotis carimatae G. S. Miller

Proc. U.S. Natl. Mus. 31: 62, 23 Jul 1906a.

= Myotis adversus carimatae G. S. Miller, 1906. See Hill 

(1983:162).

Holotype: USNM 125154. In alcohol, [skull removed]. Adult 

female. Collected 28 Aug 1904 by W. L. Abbott. Original 

number 3673.

Type Locality: Telok (Teluk) Edar, Karimata Island (Pulau Karimata), [Kalimantan Barat, Borneo (Kalimantan), Indonesia]. 

Myotis deignani Shamel

J. Mammal. 23: 324, 14 Aug 1942a.

= Myotis horsfieldii deignani Shamel, 1942. See Hill (1983:157).

Holotype: USNM 267264. Skin and skull. Adult male. Collected 

20 Oct 1936 by H. G. Deignan. Original number [SL: 97].

Type Locality: Chiengmai (Chiang Mai), [SL: altitude 1,000 ft 

(305 m)], [Chiang Mai], Siam (Thailand). 

Myotis diminutus Moratelli and Wilson

Mamm. Biol. 76: 609, Sep 2011.

Holotype: USNM 528569. Skin and skull. Sub-adult male. Collected 11 Feb 1979 by D. E. Wilson. Original number DEW 

4724.

Type Locality: Rio Palenque Science Center, elevation about 

150 m, 0o35′11 ̋S, 79o21′53 ̋W, 47 km S (by road) from 

Santo Domingo, Los Rios, Ecuador. 

Myotis dominicensis G. S. Miller

Proc. Biol. Soc. Wash. 15: 243, 16 Dec 1902f.

Holotype: USNM 113564. In alcohol, [skull removed]. Adult 

male. Collected 20 Jul 1901 by H. S. Branch. No original 

number.

Type Locality: Dominica, West Indies. 

Myotis findleyi Bogan

J. Mammal. 59: 524, 21 Aug 1978.

Holotype: USNM 512417. Skin and skull. Adult male. Collected 

14 Mar 1976, collector unknown. Original number C. B. 

Robbins 5756.

Type Locality: Isla Maria Magdalena, Islas Tres Marias, Nayarit, 

[Mexico]. 

Myotis flavus Shamel

J. Mammal. 25: 191, 25 May 1944.

= Myotis formosus watasei (Kishida, 1924). See Simmons 

(2005:507).

Holotype: USNM 239908. In alcohol, [skull removed]. Adult female. Collected Jul 1923 by Y. Horikawa. Original number 

[SL: 6].

Type Locality: Enri, Formosa (Taiwan). 

Myotis grisescens A. H. Howell

Proc. Biol. Soc. Wash. 22: 46, 10 Mar 1909.

Holotype: USNM 157517. Skin and skull. Adult male. Collected 

31 Aug 1908 by A. H. Howell. Original number 1233.

Type Locality: Nickajack Cave, near Shellmound, Marion 

County, Tennessee. 

Myotis handleyi Moratelli, Gardner, De Oliveira, and Wilson

Am. Mus. Novit. 3780: 11, 16 Sep 2013.

Holotype: USNM 370932. Skin and skull. Adult male. Collected 

19 Aug 1965 by the Smithsonian Venezuelan Project team 

[SL: M. D. Tuttle]. Original number SVP 753.
NUMBER 644 • 7 5

Type Locality: Pico Avila, [SL: Hotel Humboldt], elevation 2,092 

m, 5 km northeast [SL: 9.4 km N] of Caracas, about 10.33o 

(10o33′) N, 66.52o (66o52′) W, Distrito Federal, Venezuela. 

Myotis hirsutus A. B. Howell

Proc. Biol. Soc. Wash. 39: 139, 27 Dec 1926.

= Myotis fimbriatus (Peters, 1871). See Corbet and Hill (1992: 

125).

Holotype: USNM 238863. Skin and skull. Adult female. Collected 7 Apr 1922 by A. de C. Sowerby. Original number 

1358.

Type Locality: Near Yenpingfu (Nanping), altitude 2,000 ft 

(610 m), Fukien (Fujian), [SL: SE] China. 

Myotis longicrus amotus G. S. Miller

Proc. Biol. Soc. Wash. 27: 212, 31 Oct 1914b.

= Myotis volans amotus G. S. Miller, 1914. See Miller and Allen 

(1928:145).

Holotype: USNM 54437. Skin and skull. Adult female. Collected 

27 May 1893 by E. W. Nelson. Original number 4873.

Type Locality: Cofre de Perote, altitude 12,500 ft (3,810 m), 

Vera Cruz (Veracruz), Mexico. 

Myotis longicrus interior G. S. Miller

Proc. Biol. Soc. Wash. 27: 211, 31 Oct 1914b.

= Myotis volans interior G. S. Miller, 1914. See Miller and Allen 

(1928:142).

Holotype: USNM 133426. Skin and skull. Adult male. Collected 

23 Jul 1904 by V. Bailey. Original number 8182.

Type Locality: 5 mi (8 km) south of Twining, altitude 11,300 

ft (3,444 m) [FC: 11,400 ft (3,475 m)], Taos County, New 

Mexico. 

Myotis lucifugus alascensis G. S. Miller

N. Am. Fauna 13: 63, 16 Oct 1897a.

Holotype: USNM 77416. In alcohol, [skull removed]. Adult 

female. Collected 5 Aug 1895 by C. P. Streator. Original 

number 4754.

Type Locality: Sitka, Alaska. 

Myotis lucifugus phasma Miller and Allen

Bull. U.S. Natl. Mus. 144: 53, 25 May 1928.

= Myotis yumanensis yumanensis (H. Allen, 1864). See Harris 

and Findley (1962:194).

Holotype: USNM 148159. Skin and skull. Adult female. Collected 28 Aug 1906 by M. Cary. Original number 792.

Type Locality: [Armstrong (1972:59): Little] Snake River, south 

of Sunny Peak, Routt (Moffat) County, Colorado. 

Myotis micronyx Nelson and Goldman

Proc. Biol. Soc. Wash. 22: 28, 10 Mar 1909.

= Myotis evotis micronyx Nelson and Goldman, 1909. See Manning (1993:43).

Holotype: USNM 146044. [Skin and skull]. Adult male. Collected 8 Nov 1905 by E. W. Nelson and E. A. Goldman. 

Original number 18490.

Type Locality: Comondu, altitude 700 ft (213 m), Lower California (Baja California Sur), Mexico. 

Myotis nesopolus G. S. Miller

Proc. Biol. Soc. Wash. 13: 123, 6 Apr 1900d.

Holotype: USNM 101849. Skin and skull from specimen in alcohol [SL: 8 Feb 1900]. Adult male. Collected 4 Nov 1899 

by [SL: L. B. Smith] not L. J. Guthrie as given in original 

description. No original number.

Type Locality: [SL: Punda], near Willemstad, Curacao, [Netherlands Antilles], West Indies. 

Myotis niasensis Lyon

Proc. U.S. Natl. Mus. 52: 441, 30 Dec 1916.

= Myotis muricola niasensis Lyon, 1916. See Hill (1983:153).

Holotype: USNM 121876. In alcohol, skull removed. Age and 

sex not given. Collected 20 Mar 1903 by W. L. Abbott. 

Original number 2402.

Type Locality: Siaba Bay, Nias Island (Pulau Nias), [Sumatera 

Utara], off west coast of Sumatra, [Indonesia]. 

Myotis nigricans extremus Miller and Allen

Bull. U.S. Natl. Mus. 144: 181, 25 May 1928.

Holotype: USNM 77670. Skin and skull. Adult female. Collected 1 Mar 1896 by E. W. Nelson and E. A. Goldman. 

Original number 9455.

Type Locality: Huehuetan, altitude 300 ft (91 m), [Huehuetan 

Municipality], Chiapas, Mexico. 

Myotis occultus Hollister

Proc. Biol. Soc. Wash. 22: 43, 10 Mar 1909.

Holotype: USNM 137098. Skin and skull. Adult male. Collected 

14 May 1905 by N. Hollister. Original number 2237.

Type Locality: West side of Colorado River [SL: bottom], 10 

mi (16 km) above Needles, [San Bernardino County], 

California. 

Myotis pernox Hollister

Smithson. Misc. Collect. 56(26): 4, 5 Dec 1911b.

= Myotis lucifugus pernox Hollister, 1911. See Crowe (1943:395).

Holotype: USNM 174134. Skin and skull. Adult male. Collected 

6 Sep 1911 by J. H. Riley. Original number 2318.

Type Locality: Henry House, Alberta, [Canada]. 

Myotis petax Hollister

Smithson. Misc. Collect. 60(14): 6, 29 Nov 1912c.

See Matveev et al. (2005:33) for current use.

Holotype: USNM 175189. Skin and skull. Adult female. Collected 30 Jul, not Jun as in original description, 1912 by 

N. Hollister. Original number 4412.
7 6 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Type Locality: Kosh-Agatch (Kosh-Agach), Chuisaya Steppe, altitude 7,300 ft (2,225 m), Altai District (Altayskiy), Siberia 

(Russia). 

Myotis petax loukashkini Shamel

Proc. Biol. Soc. Wash. 55: 103, 13 Aug 1942b.

= Myotis daubentonii loukashkini Shamel, 1942. See Kruskop 

(2002:94).

Holotype: USNM 270561. Skin and skull. Adult female. Collected 

25 Jun 1937 by A. S. Loukashkin. Original number 487.

Type Locality: Wutalienchieh, Third Lake, Heilungkiang (Heilongjiang) Province, North Manchuria (China). 

Myotis siligorensis thaianus Shamel

J. Mammal. 23: 323, 14 Aug 1942a.

Holotype: USNM 260612. Skin and skull, jaws missing. Adult 

male. Collected 22 Jan 1936 by H. G. Deignan. Original 

number [SL: 51].

Type Locality: Chiengmai (Chiang Mai), [SL: 1,000 ft (305 m)], 

[Chiang Mai], [FC: northern] Siam (Thailand). 

Myotis simus riparius Handley

Proc. U.S. Natl. Mus. 112: 466, 6 Oct 1960.

= Myotis riparius Handley, 1960. See LaVal (1973a: 32).

Holotype: USNM 310255. Skin and skull. Adult female. Collected 9 Feb 1959 by C. O. Handley, Jr. and B. R. Feinstein. 

Original number [FC: COH, Jr.] 4843.

Type Locality: Tacarcuna Village [SL: Camp], 3,200 ft (975 m), 

Rio Pucro, Darien, Panama. 

Myotis sowerbyi A. B. Howell

Proc. Biol. Soc. Wash. 39: 138, 27 Dec 1926.

= Myotis siligorensis sowerbyi A. B. Howell, 1926. See Simmons 

(2005:516).

Holotype: USNM 238869. Skin and skull. Adult female. Collected 

7 Apr 1922 by A. de C. Sowerby. Original number 1366.

Type Locality: Near Yenpingfu (Nanping), altitude 2,000 ft 

(610 m), not 3,000 ft as in original description, Fukien (Fujian), China. 

Myotis thysanodes G. S. Miller

N. Am. Fauna 13: 80, 16 Oct 1897a.

Holotype: USNM 29827/38894. In alcohol, [skull removed]. 

Adult female. Collected 5 Jul 1891 by T. S. Palmer. Original 

number 235.

Type Locality: Old Fort Tejon, [Tehachapi Mountains, Kern 

County], California. 

Myotis winnemana Nelson

Proc. Biol. Soc. Wash. 26: 183, 8 Aug 1913.

= Myotis leibii (Audubon and Bachman, 1842). See Glass and 

Baker (1965:205).

Holotype: USNM 150275. [Skin and skull]. Adult male. Collected 

31 Aug 1907 by A. K. Fisher. Original number [SL: 2432].

Type Locality: Plummers Island, in Potomac River 10 mi (16 km) 

above Washington, [Montgomery County], Maryland. 

Myotis yumanensis lutosus Miller and Allen

Bull. U.S. Natl. Mus. 144: 72, 25 May 1928.

Holotype: USNM 50783. Skin and skull. Adult female. Collected 

17 Jul 1892 by E. W. Nelson. Original number [SL: 2860].

Type Locality: Patzcuaro, [FC: altitude 7,000 ft (2,134 m)], Michoacan, Mexico. 

Myotis yumanensis saturatus G. S. Miller

N. Am. Fauna 13: 68, 16 Oct 1897a.

Holotype: USNM 17399/24306. Skin and skull. Adult male. Collected 13 Sep 1889 by T. S. Palmer. Original number 392.

Type Locality: Hamilton, [Skagit County], Washington. 

Pipistrellus cinnamomeus G. S. Miller

Proc. Acad. Nat. Sci. Phila. 54: 390, 3 Sep 1902d.

= Myotis fortidens fortidens Miller and Allen, 1928. See Hall 

and Dalquest (1950:586).

Holotype: USNM 100231. Skin and skull. Adult female. Collected 4 May 1900 by E. W. Nelson and E. A. Goldman. 

Original number 14136.

Type Locality: Montecristo, Tabasco, Mexico. 

Vespertilio affinis H. Allen

Smithson. Misc. Collect. 7(1): 53, Jun 1864.

= Myotis lucifugus lucifugus (LeConte, 1831). See G. S. Miller 

(1897a:20).

Holotype: USNM 5342. In alcohol, [skull removed and cannot 

be found]. [Lyon and Osgood (1909:270): Adult female]. 

Presented by Dr. Shumard. Date of collection, collector, and 

original number not given [MC: Cataloged 30 Oct 1861].

Type Locality: Fort Smith, [Sebastian County], Arkansas.

Remarks: The description was based on a single specimen, 

USNM 5342, listed by number. 

V[espertilio]. agilis H. Allen

Proc. Acad. Nat. Sci. Phila. 18: 282, ordered published 28 Aug 

1866.

= Myotis californicus mexicanus (Saussure, 1860). See Miller 

and Allen (1928:159).

Holotype: ANSP 1874. In alcohol, [skull removed]. [Koopman 

(1976:5): Adult female]. Collected by Dr. Sartorius. Date 

unknown.

Type Locality: Mirador, [Veracruz], Mexico.

Remarks: H. Allen (1866:282) stated that the single specimen 

was in the Museum of the Smithsonian Institution but did 

not give a number. Both Lyon and Osgood (1909:290) and 

Poole and Schantz (1942:572) stated that nothing is known 

of the specimen. According to Koopman (1976:5), and verified by R. D. Fisher on 19 Jun 1996, it is in the collection of 

the Academy of Natural Sciences of Philadelphia (Academy 

of Natural Sciences of Drexel University). 
NUMBER 644 • 7 7

Vespertilio ciliolabrum Merriam

Proc. Biol. Soc. Wash. 4: 2, 17 Dec 1886.

= Myotis ciliolabrum (Merriam, 1886). See van Zyll de Jong 

(1984:2525).

Holotype: USNM 186444. [In alcohol, skull removed]. Adult female. Collected Aug 1885 by A. B. Baker. No original number. Merriam collection number 2797.

Type Locality: Near Banner, Trego County, Kansas.

Remarks: Published as Merriam collection number 2797. Cataloged USNM 186444 on 13 Oct 1911. 

Vespertilio evotis H. Allen

Smithson. Misc. Collect. 7(1): 48, Jun 1864.

= Myotis evotis evotis (H. Allen, 1864). See G. S. Miller 

(1897a:77).

Lectotype: USNM 5389 as designated by Lyon and Osgood 

(1909:290). In alcohol. Age and sex not given. Collected 

by A. S. Taylor, date unknown. No original number. [MC: 

Cataloged 31 Oct 1861].

Type Locality: Monterey, [Monterey County], California.

Remarks: A type was not designated by number in the original description. Lyon and Osgood (1909:290) stated that 

USNM 5389 should be considered the type = lectotype 

(ICZN, 1999: Article 74.5). This specimen cannot be found. 

V[espertilio]. exiguous H. Allen

Proc. Acad. Nat. Sci. Phila. 18: 281, ordered published 28 Aug 

1866.

= Myotis nigricans nigricans (Schinz, 1821). See Miller and Allen 

(1928:178).

Holotype: USNM 5373, ANSP 5626. In alcohol. [MC: Female], 

age not given. Collector unknown. [MC: Received from Dr. 

Hayer. Cataloged 30 Oct 1861].

Type Locality: Aspinwall (Colon), N. G. (Colon, Panama).

Remarks: The museum catalog states that this specimen was not 

found 5 Mar 1902. According to Koopman (1976:6), and 

verified by R. D. Fisher on 19 Jun 1996, it is in the collection 

of the Academy of Natural Sciences of Philadelphia (Academy of Natural Sciences of Drexel University) as ANSP 

5626 and also bears the ANSP number 1813. 

V[espertilio]. exilis H. Allen

Proc. Acad. Nat. Sci. Phila. 18: 283, ordered published 28 Aug 

1866.

= Myotis californicus californicus (Audubon and Bachman, 

1842). See G. S. Miller (1897a:69).

Holotype: USNM 5402, ANSP 1841. In alcohol. Male, age not 

given. Collected by J. Xantus, date unknown. Original number [MC: 3895. Cataloged 31 Oct 1861].

Type Locality: Cape (Cabo) St. (San) Lucas, [Baja California Sur, 

Mexico].

Remarks: This specimen, listed by number (USNM 5402) in the 

original description, was originally USNM 4977, cataloged 

on 31 May 1861. It is also a syntype of Vespertilio oregonensis H. Allen, 1864. Overlooked by both Lyon and Osgood (1909) and Poole and Schantz (1942), this specimen is 

currently ANSP number 1841. See Koopman (1976:5) and 

verified by R. D. Fisher on 19 Jun 1996. 

Vespertilio gryphus var. septentrionalis Trouessart

Cat. Mamm. Viv. Foss. Vol. 1: 131, 1897.

= Myotis septentrionalis (Trouessart, 1897). See van Zyll de Jong 

(1979:993).

Lectotype: USNM 8188/38663 as designated by Miller and Allen 

(1928:106). In alcohol, skull removed. Adult female. Collected on unknown date by Dr. [MC: J. B.] Gilpin. No original number. [MC: Skin cataloged in 1865; skull 26 Feb 1914].

Type Locality: Halifax, Nova Scotia, [Canada]. 

Vespertilio longicrus True

Sci. 8: 588, 24 Dec 1886b.

= Myotis volans longicrus (True, 1886). See Miller and Allen 

(1928:140).

Holotype: USNM 15623/22480. [In alcohol, skull removed]. 

Young adult female. Collected on unknown date by D. S. 

Jordan. No original number. [MC: Cataloged 16 Dec 1886].

Type Locality: Vicinity of Puget Sound, [Washington].

Remarks: A type was not designated by number in the original description, but a single specimen is implied. It is undoubtedly USNM 15623/22480 and type is written in the 

“Remarks” column of the museum catalog. The holotype is 

fixed by monotypy (ICZN, 1999: Article 73.1.2). 

V[espertilio]. lucifugus LeConte

In McMurtrie, The Animal Kingdom 1: 431, 1831.

= Myotis lucifugus lucifugus (LeConte, 1831). See G. S. Miller 

(1897a:59).

Syntype: USNM 4741. Skin only. Age and sex not given. Collected on unknown date by J. E. LeConte. No original number. [MC: Cataloged 13 Apr 1861].

Type Locality: Not given. [MC: United States]. [Miller and Rehn 

(1901:256): Georgia; probably the LeConte plantation 

(Woodmanston) near Riceboro, Liberty County].

Remarks: A note in the USNM skin catalog says “now (Mar 

1927) in the British Museum Number 7.1.1.534.” Miller 

and Allen (1928:44) stated that “this specimen may well be 

one of the specimens on which LeConte based his original 

account of the animal.” 

Vespertilio macropus H. Allen

Proc. Acad. Nat. Sci. Phila. 18: 288, ordered published 28 Aug 

1866.

= Myotis yumanensis yumanensis (H. Allen, 1864). See G. S. 

Miller (1897a:66).

Holotype: USNM 84549. [Skin only]. Mature [SL: male]. Collected [SL: 1 Nov 1866; but in a note to C. H. Merriam, 

Coues says 30 Oct 1865] by E. Coues. No original number. 

Dr. Coues’s private collection.
7 8 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Type Locality: Near Fort Majaor (Mohave), [note from Coues 

to Merriam dated 30 Jan 1898 states “Piute Spring, about 

25 mi (40 km) W of Fort Mojave”], New Mexico (Mohave 

County, Arizona).

Remarks: A type was not designated by number in the original description, which was based on a single specimen in 

Coues’s private collection. USNM 84549, bearing a Coues 

private collection label, is marked V. macropus Nov. Type, 

and is the only one in the collection that matches the data 

given by Allen. It is without question the specimen upon 

which the taxon was based and therefore is the holotype 

by monotypy (ICZN, 1999: Article 73.1.2). It was found in 

the collection unnumbered and cataloged USNM 84549 on 

21 Apr 1898. The discrepancy between the date collected 

on the skin label and the one given by Coues is difficult to 

understand, but we believe Coues to be correct, as H. Allen 

(1866:288) stated he was reporting on specimens collected 

by Coues in 1864 and 1865 and the date the description 

was ordered published, 28 Aug 1866, was before the date 

collected given on the skin label, 1 Nov 1866. 

Vespertilio melanorhinus Merriam

N. Am. Fauna 3: 46, 11 Sep 1890a.

= Myotis melanorhinus (Merriam, 1890). See Simmons (2005: 

511).

Holotype: USNM 18684. In alcohol. [SL: Made into a skin and 

skull 7 Nov 1910]. Adult male. Collected 4 Aug 1889 by C. H. 

Merriam [SL: and V. Bailey]. Original number [FC: V. B.] 275.

Type Locality: Little Spring, north base of San Francisco Mountain, [Miller and Kellogg (1955:96): 8,250 ft (2,515 m)], 

[Coconino County], Arizona. 

V[espertilio]. mundus H. Allen

Proc. Acad. Nat. Sci. Phila. 18: 280, ordered published 28 Aug 

1866.

= Myotis nigricans nigricans (Schinz, 1821). See LaVal (1973a:7).

Holotype: USNM 5547, ANSP 1812. In alcohol. Young female. 

No collector, original number, or date collected given. [MC: 

Cataloged 6 Feb 1862].

Type Locality: Maracaibo, [Zulia], Venezuela.

Remarks: This specimen is currently ANSP 1812, collection of 

the Academy of Natural Sciences of Philadelphia (Academy of Natural Sciences of Drexel University) (Koopman, 

1976:6), as verified by R. D. Fisher on 19 Jun 1996. 

Vespertilio nitidus H. Allen

Proc. Acad. Nat. Sci. Phila. 14: 247, ordered published 29 Apr 

1862.

= Myotis californicus californicus (Audubon and Bachman, 

1842). See G. S.Miller (1912b:56).

Lectotype: USNM 1981 as designated by G. S. Miller (1897a:69). 

[Skin]. Age and sex not given. Collected by W. Hutton, date 

unknown. No original number. [MC: Cataloged 13 Dec 

1856 and then later, 31 Oct 1861 as USNM 5437].

Type Locality: Monterey, [Monterey County], California.

Remarks: A type was not designated in the original description. 

G. S. Miller (1897a:69), as first revisor, selected Monterey 

as the type locality and refered Allen’s specimens from Fort 

Steilacoom to his new taxon, Myotis californicus caurinus.

H. Allen (1862:247) had only one specimen from Monterey, 

USNM 1981. Although there currently is a skull in the collection that bears USNM 1981, it is in error. Number 1981 

in the bone catalog is a Putorius vison from Carlisle, Pennsylvania and the skull in question is a Parastrellus hesperus (A. L. Gardner, pers. com. to R. D. Fisher). There is no 

reason to believe that a skull was ever cataloged for this 

specimen. 

Vespertilio nitidus henshawii H. Allen

Bull. U.S. Natl. Mus. 43: 103, 14 Mar 1894.

= Myotis melanorhinus (Merriam, 1890). See Simmons (2005: 

511).

Syntype: USNM 12450. [In alcohol]. Gravid female. Collected in 

1872 [MC: Jul 1874] by H. W. Henshaw. Original number 

[MC: 1237].

Syntype: USNM 102426. [In alcohol, skull removed]. Gravid female. Collected in 1872 [MC: Jul 1874] by H. W. Henshaw. 

Original number [MC: 1237].

Type Locality: Near Fort Wingate, [McKinley County], New 

Mexico.

Remarks: Both specimens were originally cataloged as USNM 

12450. Later, on 12 Jun 1900, one was recataloged as 

USNM 102426. 

Vespertilio obscurus H. Allen

Proc. Acad. Nat. Sci. Phila. 18: 281, ordered published 28 Aug 

1866.

= Myotis yumanensis yumanensis (H. Allen, 1864). See Goldman 

(1914c:102).

Holotype: USNM 8223/38357. In alcohol, [skull removed]. 

Young male. Collected on unknown date by J. Xantus. No 

original number. [MC: Skin cataloged between 10 Jul and 

27 Oct 1865; skull 6 Feb 1909].

Type Locality: Lower (Baja) California, [Mexico].

Remarks: Published as USNM 8223, to which USNM 38357 

belongs. 

Vespertilio oregonensis H. Allen

Smithson. Misc. Collect. 7(1): 61, Jun 1864.

= Myotis californicus californicus (Audubon and Bachman, 

1842). See G. S. Miller (1897a:69).

Syntype: USNM 5402, ANSP 1841. In alcohol. Male. Collected 

by J. Xantus, date unknown. Original number [MC: 3895. 

Originally cataloged as USNM 4977 on 31 May 1861; recataloged as USNM 5402 on 31 Oct 1861].

Syntype: USNM 5405. In alcohol. [Lyon and Osgood (1909:273): 

Female]. Collected in [MC: 1855] by G. H. Thomas. No 

original number. [MC: Cataloged 31 Oct 1861].
NUMBER 644 • 7 9

Syntype: USNM 5537. In alcohol. Male. Collected by G. H. 

Thomas [MC: probably in 1855. Cataloged 6 Feb 1862]. 

No original number.

Syntype: USNM 5512/37335. Skin and skull. No data were originally associated with this specimen. It is labeled Vespertilio 

oregonensis Major LeConte. [MC: Skin originally cataloged 

as USNM 4740 on 13 Apr 1861. Recataloged as USNM 

5512 on 1 Nov 1861. Skull cataloged on 2 Mar 1898]. 

Type Locality: The type locality comprises all of the localities 

published for the syntypes (ICZN, 1999: Article 73.2.3). 

They are USNM 5402: Cape Saint (Cabo San) Lucas, [Baja 

California Sur, Mexico]; USNM 5405 and 5537: Fort 

Yuma, Arizona, ([Imperial County], California); and USNM 

5512/37335: no published locality [MC: United States].

Remarks: See G. S. Miller (1897a:33) for the application of Vespertilio oregonensis to these specimens. USNM 5402 is also 

the holotype of Vespertilio exilis H. Allen and is in the collection of the Academy of Natural Sciences of Philadelphia 

(Academy of Natural Sciences of Drexel University) as ANSP 

1841 (Koopman, 1976:5), as verified by R. D. Fisher on 19 

Jun 1996. USNM 5537 cannot be found. USNM 5512 was 

published under that number, to which skull 37335 belongs. 

Vespertilio ruber E. Geoffroy

Ann. Mus. Hist. Nat. Paris 8: 204, 1806.

= Myotis ruber (E. Geoffroy, 1806). See Thomas (1902:493).

Neotype: USNM 115097 as designated by LaVal (1973a:45). 

Skin and skull, mandible missing. Adult male. Collected 22 

May 1901 by W. Foster. Original number [SL: 347].

Type Locality: Sapucay (Sapucai), elevation 300 m, [Paraguari], 

Paraguay. 

Vespertilio subulatus keenii Merriam

Am. Nat. 29: 860, 1 Sep 1895c.

= Myotis keenii (Merriam, 1895). See Miller and Allen (1928: 

104).

Holotype: USNM 72922. [In alcohol, skull removed]. Adult female. Collected summer of 1894 by J. H. Keen. No original 

number.

Type Locality: Massett (Masset), Graham Island, Queen Charlotte Islands, B. C. (British Columbia), [Canada]. 

Vespertilio tenuidorsalis H. Allen

Proc. Acad. Nat. Sci. Phila. 18: 283, ordered published 28 Aug 

1866.

= Myotis californicus californicus (Audubon and Bachman, 

1842). See G. S. Miller (1897a:69).

Holotype: USNM 5533, ANSP 1839. In alcohol. Female. Collected by J. Xantus. No other data provided.

Type Locality: Cape St. (Cabo San) Lucas, Lower California 

(Baja California Sur), [Mexico].

Remarks: The specimen number given by Allen must be incorrect as USNM 5533 is cataloged as a V. nitidus from Puget 

Sound collected by C. B. Kennerley. This specimen is currently in the collection of the Academy of Natural Sciences 

of Philadelphia (Academy of Natural Sciences of Drexel 

University), ANSP 1839, and bears the locality information 

given here under Type Locality (Koopman, 1976:5), as verified by R. D. Fisher on 19 Jun 1996. 

Vespertilio volans H. Allen

Proc. Acad. Nat. Sci. Phila. 18: 282, ordered published 28 Aug 

1866.

= Myotis volans volans (H. Allen, 1866). See Goldman 

(1914c:102).

Holotype: USNM 5398/37326, ANSP 1834. In alcohol, [skull 

removed]. Age and sex not given. Collected on unknown 

date by J. Xantus. No original number. [MC: Skin cataloged 

31 Oct 1861; skull 1 Mar 1898].

Type Locality: Cape (Cabo) San Lucas, Lower California (Baja 

California Sur), [Mexico].

Remarks: Published as USNM 5398 to which USNM 37326 belongs. The fluid specimen is in the collection of the Academy of Natural Sciences of Drexel University as ANSP 1834 

(Koopman, 1976:5), as verified by R. D. Fisher on 19 Jun 

1996. 

Vespertilio yumanensis H. Allen

Smithson. Misc. Collect. 7(1): 58, Jun 1864.

= Myotis yumanensis yumanensis (H. Allen, 1864). See G. S. 

Miller (1897a:66).

Syntype: USNM 5367. In alcohol. Female. Collected by G. H. 

Thomas in 1855. No additional data given.

Syntype: USNM 6019. In alcohol. Male. Collected by G. H. 

Thomas in 1855. No additional data given.

Syntype: USNM 6020. In alcohol. Female. Collected by G. H. 

Thomas in 1855. No additional data given.

Syntype: USNM 6021. In alcohol. Female. Collected by G. H. 

Thomas in 1855. No additional data given.

Type Locality: Fort Yuma, [Imperial County], California.

Remarks: None of these specimens can be found. Remarks in 

the Museum Catalog for USNM 5367 and 6021 state “not 

found 5 Mar 1902.” For USNM 6020 the remark is “distributed to Royal Mus. Copenhagen, 6 Aug 1866.” H. Allen 

(1866:285) stated that his type series of this taxon was not 

available to him for his 1866 work as they were mislaid during the fire at the Smithsonian Institution in Jan 1865 and 

have not since been found. 

Genus Neoromicia Roberts, 1926

Eptesicus ugandae Hollister

Smithson. Misc. Collect. 66(1): 3, 10 Feb 1916c.

= Neoromicia somalicus ugandae (Hollister, 1916). See Simmons 

(2005:495).

Holotype: USNM 166520. In alcohol, skull removed. Adult female. Collected 15 Feb 1910 by J. A. Loring. Original number 9022.
8 0 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Type Locality: Ledgus, [Central Equatoria], Uganda (South 

Sudan). 

Pipistrellus helios Heller

Smithson. Misc. Collect. 60(12): 3, 4 Nov 1912a.

= Neoromicia helios (Heller, 1912). See Simmons (2005:494).

Holotype: USNM 181813. [Skin and skull]. Adult male. Collected 22 Jul 1911 by E. Heller. Original number 3065.

Type Locality: Merelle Water, 30 mi (48 km) south of Mt. Marsabit, [Eastern Province], British East Africa (Kenya). 

Pipistrellus minusculus G. S. Miller

Proc. Wash. Acad. Sci. 2: 647, 28 Dec 1900c.

= Neoromicia nanus minusculus (G. S. Miller, 1900). See Simmons (2005:494).

Holotype: USNM 84500. In alcohol, [skull removed]. Adult 

female. Collected May 1894 by O. F. Cook. No original 

number.

Type Locality: Mount Coffee, [Montserrado], Liberia. 

Genus Nycticeius Rafinesque, 1819

Nyct[icea]. crepuscularis LeConte

In McMurtrie, The Animal Kingdom 1: 432, 1831.

= Nycticeius humeralis (Rafinesque, 1818). See G. S. Miller 

(1897a:118).

Syntypes: USNM 4735 and USNM 4736. [Skins with skulls 

inside]. Age and sex not given. Collected by [MC: Maj. 

LeConte]. No original number. [MC: Cataloged 13 Apr 

1861].

Type Locality: None given. [MC: United States]. [LeConte 

(1855:433): inhabits Georgia].

Remarks: Following Lyon and Osgood (1909:159), these specimens have been regarded as syntypes more from tradition 

than anything else. There is nothing in the original description to show that they are syntypes. They were presented 

to the museum years ago by Major LeConte as typical, or 

perhaps as original, specimens. Each bears two old labels: 

a small paper tag and a standard Smithsonian Institution 

label. The word “Crepuscularis” is written on each of the 

paper tags in a handwriting that cannot be identified. The 

large label on USNM 4735 is so stained as to be illegible, 

but the one on USNM 4736 is in good condition. Its inscription reads “4736 Vespertilio crepuscularis U. States Maj. 

LeConte” and was written by Baird, according to Lyon and 

Osgood (1909:159). 

Genus Nyctophilus Leach, 1821

Nyctophilus arnhemensis Johnson

Proc. Biol. Soc. Wash. 72: 184, 4 Nov 1959.

Holotype: USNM 284240, not 284249 as in original description. Skin and skull. Adult male. Collected 12 Aug 1948 by 

D. H. Johnson. Original number 5692.

Type Locality: Rocky Bay, south of Yirrkala, Cape Arnhem Peninsula, Northern Territory, Australia, lat. 12o16′S, long. 

136o47′E. 

Genus Parastrellus Hoofer, Van Den Bussche, 

and Horacek, 2006

Pipistrellus hesperus australis G. S. Miller

N. Am. Fauna 13: 90, 16 Oct 1897a.

= Parastrellus hesperus hesperus (H. Allen, 1864). See Findley 

and Traut (1970:756) who placed australis in hesperus

and Hoofer et al. (2006:987) who placed hesperus in Parastrellus.

Holotype: USNM 52112. In alcohol. Adult female. Collected 14 

May 1892 by E. W. Nelson. Original number 2614.

Type Locality: Barranca Ibarra, altitude about 3,000 ft (914 m), 

Jalisco, Mexico. 

Pipistrellus hesperus maximus Hatfield

J. Mammal. 17: 261, 14 Aug 1936.

= Parastrellus hesperus maximus (Hatfield, 1936). See Hoofer et 

al. (2006:988).

Holotype: USNM 21344/36997. Skin and skull. Adult female. 

Collected 6 Jun 1892 by E. A. Mearns and F. X. Holzner. 

Original number 1854.

Type Locality: Dog Spring, Grant (Hidalgo) County, New 

Mexico. 

Pipistrellus hesperus oklahomae Glass and Morse

J. Mammal. 40: 531, 20 Nov 1959.

= Parastrellus hesperus maximus (Hatfield, 1936). See Findley 

and Traut (1970:761) who placed oklahomae in maximus 

and Hoofer et al. (2006:988) who placed hesperus in Parastrellus.

Holotype: USNM 287970. Skin and skull. Adult female. Collected 2 Jun 1955 by B. P. Glass. Original number 1634.

Type Locality: Granite, north side of Granite Mountain, Greer 

County, Oklahoma.

Remarks: Published as Oklahoma State University Museum of 

Zoology number 2863. Cataloged as USNM 287970 on 26 

Mar 1958. 

Pipistrellus hesperus santarosae Hatfield

J. Mammal. 17: 261, 14 Aug 1936.

= Parastrellus hesperus maximus (Hatfield, 1936). See Findley 

and Traut (1970:761) who placed santarosae in maximus

and Hoofer et al. (2006:988) who placed hesperus in Parastrellus.

Holotype: USNM 118756. Skin and skull. Adult female. Collected 6 Oct 1902 by J. H. Gaut. Original number 568.

Type Locality: Santa Rosa, Guadalupe County, New Mexico. 

Scotophilus hesperus H. Allen

Smithson. Misc. Collect. 7(1): 43, Jun 1864.
NUMBER 644 • 8 1

= Parastrellus hesperus hesperus (H. Allen, 1864). See Hoofer et 

al. (2006:987).

Lectotype: USNM 5406/37406 as designated by G. S. Miller 

(1897a:88). In alcohol, [skull removed]. [Lyon and Osgood 

(1909:274): Adult] [MC: male]. Collected in [MC: 1855] by 

G. H. Thomas. No original number.

Type Locality: [Lyon and Osgood (1909:274): Old] Fort Yuma, 

[Imperial County], California.

Remarks: No type was designated by number in the original 

description, which mentioned three specimens by number. 

G. S. Miller (1897a:88) called USNM 5406/37406 the type 

= lectotype (ICZN, 1999: Article 74.5). It was published as 

USNM 5406; USNM 37406 is the skull that belongs with it. 

Vesperugo merriami Dobson

Ann. Mag. Nat. Hist. ser 5, 18: 124, Aug 1886.

= Parastrellus hesperus hesperus (H. Allen, 1864). See True 

(1888:515) who placed merriami in hesperus and Hoofer et 

al. (2006:987) who placed hesperus in Parastrellus.

Holotype: USNM 186445. [MCC: In alcohol. Adult male. Collected 3 Feb 1885 by L. Belding. Original number 11. Merriam collection number 1135].

Type Locality: [MCC: Red Bluff, Tehama County, California].

Remarks: Type not designated by number in the original description. Dobson (1886:125) stated that he had a single 

adult male, that True (1888:515) said was sent to him by 

C. H. Merriam. A perusal of the Merriam collection catalog reveals an entry, number 1135, of Vespertilio without 

a species name. At a later unknown date, Vespertilio was 

changed to Vesperugo and merriami Dobson was added. It 

is marked as the type and the data for the specimen matches 

that published for the type. It is without question the holotype by monotypy (ICZN, 1999: Article 73.1.2). Dobson 

(1886:125) erroneously gave the locality as Locust Grove, 

New York, the location of the Merriam collection at the 

time. This error was corrected by G. S. Miller (1897a:31). 

Cataloged as USNM 186445 on 13 Oct 1911. 

Genus Perimyotis Menu, 1984

Pipistrellus subflavus obscurus G. S. Miller

N. Am. Fauna 13: 93, 16 Oct 1897a.

= Perimyotis subflavus subflavus (F. Cuvier, 1832). See Davis 

(1959:522) who placed obscurus in subflavus and Menu 

(1984:415) who placed subflavus in Perimyotis. See Hoofer 

et al. (2006:991) for current use of Perimyotis.

Holotype: USNM 67723. Skin [and skull]. Adult female. Collected 6 Sep 1894 by W. K. Fisher. Original number 198.

Type Locality: Lake George, Warren County, New York. 

Genus Phoniscus Miller, 1905

Phoniscus atrox G. S. Miller

Proc. Biol. Soc. Wash. 18: 230, 9 Dec 1905b.

Holotype: USNM 123141. In alcohol, [skull removed]. Adult female. Collected 9 Sep 1903 by W. L. Abbott. Original number 2781.

Type Locality: Vicinity of Kateman River (Sungai Kateman), 

[Riau], eastern Sumatra, [Indonesia]. 

Phoniscus rapax G. S. Miller

J. Mammal. 12: 412, 11 Nov 1931a.

= Phoniscus jagorii (Peters, 1866). See Hill (1966:550) who recognized rapax as a subspecies of jagorii.

Holotype: USNM 199834. In alcohol, [skull removed]. Adult 

male. Collected 25 Aug 1914 by H. C. Raven. Original 

number 1670.

Type Locality: [Sungai] Paleleh, about latitude (longitude) 122o, 

[Sulawesi Tengah], north coast of northeastern Celebes (Sulawesi), [Indonesia]. 

Genus Pipistrellus Kaup, 1829

Pipistrellus aero Heller

Smithson. Misc. Collect. 60(12): 3, 4 Nov 1912a.

Holotype: USNM 181812. [Skin and skull]. Adult male. Collected 26 Aug 1911 by E. Heller. Original number 4110.

Type Locality: Summit of Mount Garguez (Warges), altitude 

7,000 ft (2,134 m), [SL: 7,100 ft (2,164 m)], Mathews 

Range, [Rift Valley], British East Africa (Kenya). 

Pipistrellus camortae G. S. Miller

Proc. U.S. Natl. Mus. 24: 779, 29 May 1902a.

= Pipistrellus javanicus camortae G. S. Miller, 1902. See Corbet 

and Hill (1992:136) who considered camortae a synonym 

of javanicus.

Holotype: USNM 111897. In alcohol, [skull removed]. Adult male. 

Collected 12 Feb 1902 by W. L. Abbott. No original number.

Type Locality: Kamorta (Camorta) [Island], Nicobar Islands, 

[Andaman and Nicobar Islands, India]. 

Pipistrellus subulidens G. S. Miller

Proc. Wash. Acad. Sci. 3: 134, 26 Mar 1901a.

= Pipistrellus tenuis subulidens G. S. Miller, 1901. See Koopman 

(1973:115) who considered subulidens a synonym of tenuis.

Holotype: USNM 104758. In alcohol, [skull removed]. Adult 

female. Collected 3 Jun 1900 by [MC: W. L. Abbott]. No 

original number.

Type Locality: Sirhassen Island (Pulau Serasan), South Natunas 

(Kepulauan Natuna Selatan), [Kepulauan Riau, Indonesia]. 

Genus Rhogeessa H. Allen, 1866

Rhogeessa gracilis G. S. Miller

N. Am. Fauna 13: 126, 16 Oct 1897a.

Holotype: USNM 70694. In alcohol. Adult male. Collected 24 

Nov 1894 by E. W. Nelson and E. A. Goldman. Original 

number 7099.
8 2 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Type Locality: Piaxtla, [FC: altitude 3,900 ft (1,189 m)], Puebla, 

[Mexico]. 

Rhogeessa minutilla G. S. Miller

Proc. Biol. Soc. Wash. 11: 139, 13 May 1897b.

Holotype: USNM 63216. [Skin and skull]. Adult male. Collected 

8 Jul 1895 by W. Robinson. Original number 463.

Type Locality: Margarita Island (Isla de Margarita), [Nueva Esparta], Venezuela. 

Rhogeessa parvula H. Allen

Proc. Acad. Nat. Sci. Phila. 18: 285, ordered published 28 Aug 

1866.

Holotype: USNM 7841, ANSP 1832. In alcohol, [skull removed]. 

[Lyon and Osgood (1909:292): Adult] male. Collector and 

date collected unknown. [MC: Received from A. J. Grayson. Cataloged between Feb and May 1865].

Type Locality: Tres Marias [Islands, Nayarit], Mexico.

Remarks: This specimen is currently in the collection of the Academy of Natural Sciences of Philadelphia (Academy of Natural Sciences of Drexel University) as ANSP 1832 (Koopman, 

1976:6), as verified by R. D. Fisher on 19 Jun 1996. 

Rhogeessa tumida H. Allen

Proc. Acad. Nat. Sci. Phila. 18: 286, ordered published 28 Aug 

1866.

Holotype: USNM 84021, ANSP 1831. In alcohol, [skull removed]. [Lyon and Osgood (1909:276). Adult] male. 

Collected by Dr. Sartorius, date unknown. [MC: Cataloged 

1 Mar 1898].

Type Locality: Mirador, [Veracruz], Mexico.

Remarks: Notes with the skull made by G. S. Miller, Jr. indicate that the skull was returned to the museum by H. Allen 

bearing the original published number, USNM 8195, which 

was determined to be in error and was recataloged USNM 

84021 on 1 Mar 1898. LaVal (1973b:29) determined that 

the cranium currently numbered USNM 37329 and the 

mandible that matches, USNM 84021, comprise the holotype. They had apparently been mismatched. See also Hall 

(1952:231) for additional discussion. The skin to this specimen is currently in the collection of the Academy of Natural 

Sciences of Philadelphia (Academy of Natural Sciences of 

Drexel University) as ANSP 1831 (Koopman, 1976:6), as 

verified by R. D. Fisher on 19 Jun 1996. 

Genus Scotophilus Leach, 1821

Scotophilus nucella Robbins

Ann. Mus. R. Afr. Cent., Sci. Zool. 237: 19, 1984.

Holotype: USNM 412141. Skin and skull. Adult male. Collected 

5 Aug 1967 by B. J. Hayward. Original number 4256.

Type Locality: 1 mi (1.6 km) N Nkawkaw, [SL: 6o33′N, 0o44′W], 

Eastern Region, Ghana. 
Akmali, V., A. Farazmand, J. Darvish, and M. Sharifi. 2011. Phylogeography and Taxonomic Status of the Greater 

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Albuja, V., and A. L. Gardner. 2005. A New Species of Lonchophylla Thomas (Chiroptera: Phyllostomidae) from 

Ecuador. Proceedings of the Biological Society of Washington, 118:442–449.

Alexander, L. F. 1996. A Morphometric Analysis of Geographic Variation within Sorex monticolus (Insectivora: 

Soricidae). Miscellaneous Publications, Natural History Museum, University of Kansas, 88:1–54.

Allen, G. M. 1916. Bats of the Genus Corynorhinus. Bulletin of the Museum of Comparative Zoology, 60:333–

356.

——–. 1917. Two Undescribed West Indian Bats. Proceedings of the Biological Society of Washington, 30:165–

170.

——–. 1927. Lagomorphs Collected by the Asiatic Expedition. American Museum Novitates, 284:1–11.

——–. 1938. “The Mammals of China and Mongolia.” In Natural History of Central Asia, ed. W. Granger, Central Asiatic Expeditions of the American Museum of Natural History, New York, 11: part 1:1–620.

Allen, H. 1861. Description of a New Mexican Bat. Proceedings of the Academy of Natural Sciences of Philadelphia, 13:359–361.

——–. 1862. Description of Two New Species of Vespertilionidae and Some Remarks on the Genus Antrozous.

Proceedings of the Academy of Natural Sciences of Philadelphia, 14:426–428.

——–. 1864. Monograph of the Bats of North America. Smithsonian Miscellaneous Collections, 7(1):1–85. http://

dx.doi.org/10.5479/si.03629236.43.i. 

——–. 1866. Notes on the Vespertilionidae of Tropical America. Proceedings of the Academy of Natural Sciences 

of Philadelphia, 18:279–288.

——–1889a. On the Genus Nyctinomus and Description of Two New Species. Proceedings of the American Philosophical Society, 26:1–6.

——–1889b. On the Genus Nyctinomus and Description of Two New Species. Proceedings of the American Philosophical Society, 26:558–563.

——–. 1890a. Description of a New Species of Pteropus. Proceedings of the American Philosophical Society, 

28:70–72.

——–. 1890b. Description of a New Species of Carollia and Remarks on Carollia brevicauda. Proceedings of the 

American Philosophical Society, 28:19–26.

——–. 1890c. Description of a New Species of Bat, Atalapha semota. Proceedings of the United States National 

Museum, 13:173–175. http://dx.doi.org/10.5479/si.00963801.13-807.173.

——–. 1891. On a New Species of Atalapha. Proceedings of the American Philosophical Society, 29:5–7.

——–. 1892. Description of a New Genus of Phyllostome Bats. Proceedings of the United States National Museum, 15:441–442. http://dx.doi.org/10.5479/si.00963801.15-913.441. 

——–. 1894. A Monograph of the Bats of North America. Bulletin of the United States National Museum, 43:1–

198. http://dx.doi.org/10.5479/si.03629236.43.i. 

——–. 1896. Description of a New Species of Bat of the Genus Glossophaga. Proceedings of the United States 

National Museum, 18:779–781. http://dx.doi.org/10.5479/si.00963801.18-1100.779. 

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Allen, J. A. 1875. Synopsis of the American Leporidae. Proceedings of the Boston Academy of Natural Sciences, 

17:430–436.

——–. 1877. “Family Leporidae.” In Monograph of North American Rodentia, United States Geological Survey 

of the Territories, ed. E. Coues and J. A. Allen, pp. 267–378. Washington, D.C.: Government Printing Office.

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——–. 1897b. Lepus baileyi, a New Cottontail Rabbit from Wyoming. Proceedings 

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——–. 1897c. Descriptions of Five New Shrews from Mexico, Guatemala, and 

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——–. 1897d. Two New Moles from California and Oregon. Proceedings of the 

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——–. 1897e. A New Bat of the Genus Antrozous from California. Proceedings of 

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——–. 1902a. Five New Mammals from Mexico. Proceedings of the Biological 

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——–. 1902b. Two New Shrews of the Sorex tenellus Group from California. Proceedings of the Biological Society of Washington, 15:75–76.

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NUMBER 644 • 9 1

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57:65–66.
abbotti

Galeopterus, 12

Hylobates, 21

Lepus, 25

Myotis, 73

abdita

Thyroptera, 66

abyssinicus

Colobus, 16

Acerodon

celebensis, 51

Acratocnus

comes, 8

acuminatus

Rhinolophus, 54

adusta

Macaca, 16

Monodelphis, 6

adversus

Myotis, 73, 74

aegyptius

Lepus, 25

aeneus

Scalops, 50

aequatoria

Pachyura, 49

aequatorius

Sylvisorex, 49

aereus

Scalopus, 50

aero

Pipistrellus, 81

affinis

Rhinolophus, 54

Vespertilio, 76

africana

Loxodonta, 7

africanus

Elephas, 7

agadiri

Crocidura, 33

agilis

Vespertilio, 76

agnatus

Pithecus, 17

Ailurops

furvus, 7

ursinus, 7

alascensis

Myotis, 75

Sorex, 41, 43, 45

alaskanus

Sorex, 45

alba

Ectophylla, 60

albibarbis

Hylobates, 21

Neosorex, 41

Sorex, 41, 46

albifrons

Cebus, 14

albipes

Heterohyrax, 7

albiventer

Sorex, 42

albiventris

Atelerix, 32

Marmosa, 5

albogularis

Cercopithecus, 15

Lasiopyga, 15

alchemillae

Crocidura, 33

aldabrensis

Pteropus, 52

alecto

Emballonura, 56

Pteropus, 52

aleksandrisi

Crocidura, 33

alethina

Rhinophylla, 64

Index of Taxa
9 6 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

alleni

Lepus, 23

allex

Crocidura, 33

alnorum

Sorex, 41

Alouatta

inconsonans, 14

mexicana, 14

palliata, 14

alpina

Crocidura, 33

Ochotona, 22

alpinus

Scapanus, 51

Sorex, 41

altae

Crocidura, 34

altamirae

Lepus, 26

alticola

Blarina, 37

Cryptotis, 37

Sorex, 41

altifrons

Myotis, 73

amazonicus

Eumops, 67

americanus

Lepus, 24, 25, 26, 27

ammophilus

Sylvilagus, 31

amoenus

Sorex, 41

amotus

Myotis, 75

amplus

Artibeus, 57

anambae

Tupaia, 9

anambensis

Emballonura, 56

andamanensis

Crocidura, 33

angeli

Sturnira, 64

angulatus

Cynopterus, 51

angusticeps

Blarina, 32

angustidens

Lepus, 24

annectens

Ardops, 57

Ochotona, 22

Anoura

cultrata, 57

latidens, 57

Antechinus

bilarni, 6

antillularum

Nyctinomus, 69

Tadarida, 69

Antrozous

cantwelli, 70

minor, 70

obscurus, 70

pacificus, 70

pallidus, 70

anubis

Papio, 19

aoris

Galeopithecus, 11

AOTIDAE, 14

Aotus

zonalis, 14

apaella

Sapajus, 14

aphylla

Phyllonycteris, 63

Reithronycteris, 63

apoensis

Cynomolgus, 16

aquaticus

Scalopus, 50

Sylvilagus, 30

aquilus

Platyrrhinus, 63

Vampyrops, 63

araneus

Sorex, 41, 42

aratathomasi

Sturnira, 64

arboreus

Dendrohyrax, 7

arcticus

Lepus, 24, 25

Sorex, 42, 45, 46, 47

Ardops

annectens, 57

luciae, 57

nichollsi, 57

arenaria

Lasiopyga, 15

arenarius

Miniopterus, 73

argentata

Ochotona, 22

argenteus

Presbytis, 20

arizonae

Lepus, 27, 29

Sylvilagus, 27, 28, 29

arnhemensis

Nyctophilus, 80

arquatus

Pteropus, 51

Artibeus

amplus, 57

aztecus, 59

gnomus, 59

hirsutus, 58

incomitatus, 59

intermedius, 58

jamaicensis, 58

koopmani, 58

lituratus, 58

praeceps, 58

triomylus, 58

arwasca

Presbytis, 19

ascanius

Cercopithecus, 15

Lasiopyga, 15

asellus

Lepus, 24

Atalapha

semota, 72

teliotis, 72

Atelerix

albiventris, 32

Ateles

dariensis, 14

fuscipes, 14

geoffroyi, 15

ornatus, 15

panamensis, 15

rufiventris, 14

tricolor, 15

vellerosus, 15

yucatanensis, 15

ATELIDAE, 14

ater

Hipposideros, 55

aterrimus

Pteropus, 52

Atophyrax

bendirii, 40

atriceps

Macaca, 17

atrox

Phoniscus, 81

attenuata

Crocidura, 34

audubonii

Lepus, 28

Sylvilagus, 27, 28, 29, 30

aurarius

Platyrrhinus, 63

Vampyrops, 63

auratus

Semnopithecus, 21

aurea

Macaca, 19

aurigineus

Lepus, 24

aurispinosus

Dysopes, 68

Nyctinomops, 68

australis

Corynorhinus, 70

Pipistrellus, 80

Presbytis, 19

austroriparius

Myotis, 73, 74

azteca

Dermanura, 59

aztecus

Artibeus, 59

Sylvilagus, 28

babi

Cynopterus, 51

bachmani

Lepus, 28

Sylvilagus, 28, 30

badistes

Sylvilagus, 31

bahamensis

Eptesicus, 71

Vespertilio, 71

baileyi

Lepus, 28

Myotis, 74

Sylvilagus, 28, 30

bairdi

Sorex, 42, 45

bairdii

Lepus, 24

balutus

Pteropus, 52

bancanus

Cephalopachus, 13

Nycticebus, 12

bangsii

Lepus, 25

barbensis

Hipposideros, 55

batamana

Tupaia, 10

batuanus

Presbytes, 19

baveanus

Pteropus, 52

baweanus

Pithecus, 17

beatus

Crocidura, 33

belangeri

Tupaia, 10

bendirii

Atophyrax, 40

Sorex, 40, 42

bergensis

Sorex, 42

berlandieri

Blarina, 37

Cryptotis, 37, 39

besara

Tana, 9

Tupaia, 9

bicolor

Hipposideros, 55

bilarni

Antechinus, 6

Pseudantechinus, 6

bishopi

Marmosa, 5

Marmosops, 5

Blarina

alticola, 37

angusticeps, 32

berlandieri, 37

brevicauda, 32

carolinensis, 32, 33

exilipes, 37

eximius, 37

floridana, 37
NUMBER 644

• 9 7

fossor, 37

goldmani, 38

knoxjonesi, 32

machetes, 38

magna, 38

mayensis, 38

mexicana, 38

nelsoni, 38

obscura, 38

peninsulae, 32

peregrina, 38

shermani, 32

soricina, 38

talpoides, 32

telmalestes, 32

blitchi

Sorex, 42

blossevillii

Lasiurus, 72

bogotensis

Sturnira, 64

bolivari

Crocidura, 33

bombifrons

Erophylla, 60

Kerivoula, 72

Phyllonycteris, 60

borealis

Lasiurus, 72

borneanus

Cephalopachus, 13

Galeopterus, 12

Nycticebus, 12

Tarsius, 13

borneensis

Rhinolophus, 55

botswanae

Laephotis, 72

brachycephalus

Nycticebus, 12

Platyrrhinus, 63, 64

Brachylagus

idahoensis, 23

Brachyphylla

cavernarum, 58

minor, 58

nana, 58

pumila, 58

brachyurus

Lepus, 24

BRADYPODIDAE, 8

Bradypus

ephippiger, 8

ignavus, 8

pygmaeus, 8

variegatus, 8

brasiliense

Lophostoma, 61

brasiliensis

Lepus, 28

Sylvilagus, 30

Tadarida, 69, 70

brevicauda

Blarina, 32

brevimanus

Natalus, 66

broca

Macaca, 16

brooksi

Ochotona, 22

brucei

Heterohyrax, 7

brunnescens

Ochotona, 22

bunoae

Tupaia, 9

Cabassous

centralis, 8

cadenai

Lonchophylla, 61

caecutiens

Sorex, 44

CAENOLESTIDAE, 6

cagayanus

Cynomolgus, 16

Pteropus, 52

californica

Didelphis, 4

californicus

Eumops, 67

Lepus, 24, 25, 26, 27

Macrotus, 61

Molossus, 67

Myotis, 74, 76, 77, 78, 79

Sorex, 42

callida

Lasiopyga, 15

callotis

Lepus, 25

Caluromys

centralis, 3

derbianus, 3

camortae

Pipistrellus, 81

campestris

Glossophaga, 60

Lepus, 25

cana

Presbytis, 19, 20

canescens

Tlacuatzin, 6

caniclunis

Lepus, 28

canicrus

Presbytis, 20

cansus

Ochotona, 22

cantwelli

Antrozous, 70

canus

Lepus, 24

capensis

Lepus, 25

capitalis

Pithecus, 18

capitaneus

Myotis, 74

capito

Taphonycteris, 57

capucinus

Cebus, 14

carbo

Trachypithecus, 20

carimatae

Megaderma, 56

Myotis, 74

Pithecus, 18

Presbytis, 20

Tupaia, 9

Carlito

fraterculus, 13

syrichta, 13

carolinensis

Blarina, 32, 33

Sorex, 33

Carollia

castanea, 58

perspicillata, 58

subrufa, 58

caryi

Scalopus, 50

cascadensis

Sorex, 46

castanea

Carollia, 58

Tupaia, 9

castaneus

Lasiurus, 73

catemana

Presbytis, 20

caucae

Didelphis, 4

Marmosa, 5

caudata

Crocidura, 33

caudatus

Sorex, 46

caurinus

Myotis, 74, 78

cavaticus

Taphozous, 57

cavernarum

Brachyphylla, 58

CEBIDAE, 14

Cebus

albifrons, 14

capucinus, 14

cesarae, 14

limitaneus, 14

margaritae, 14

pleei, 14

cedrophilus

Sylvilagus, 30

celebensis

Acerodon, 51

Harpyionycteris, 52

Megaderma, 56

centralis

Cabassous, 8

Caluromys, 3

Cercopithecus, 15

Myrmecophaga, 8

Philander, 3

Tatoua, 8

Centurio

greenhalli, 59

mcmurtrii, 58

senex, 59

Cephalopachus

bancanus, 13

borneanus, 13

saltator, 13

CERCOPITHECIDAE, 15

Cercopithecus

albogularis, 15

ascanius, 15

centralis, 15

kibonotensis, 15

luteus, 15

mitis, 15

schmidti, 15

stuhlmanni, 15

cervicalis

Tana, 9

Tupaia, 9

cesarae

Cebus, 14

Chacodelphys

formosa, 3

Chaerephon

jobensis, 67

luzonus, 67

naivashae, 67

plicatus, 67

pumilus, 67

pusillus, 67

chaouianensis

Crocidura, 33

chapmani

Sylvilagus, 28, 30

Cheiromeles

parvidens, 67

chiapensis

Lepus, 28

Sorex, 49

Chilonatalus

macer, 66

micropus, 66

tumidifrons, 66

Chilonycteris

gonavensis, 65

mexicana, 65

parnellii, 65

portoricensis, 65

Chilophylla

hirsuta, 55

Chiroderma

gorgasi, 59

isthmicum, 59

jesupi, 59

salvini, 59

scopaeum, 59

trinitatum, 59

villosum, 59

Chironectes

minimus, 3

panamensis, 3
9 8 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

CHIROPTERA, 51

Chlorocebus

hilgerti, 15

pygerythrus, 15

Choeroniscus

periosus, 59

chombolis

Galeopterus, 12

chrysogaster

Tupaia, 9

chrysomalla

Tupaia, 9

chrysomelas

Presbytis, 19

ciliolabrum

Myotis, 77

Vespertilio, 77

cinereus

Hylobates, 21

Lasiurus, 72

Sorex, 40, 43, 46

CINGULATA, 8

cinnamomeus

Lepus, 25

Pipistrellus, 76

circe

Rhinolophus, 54

cirrhosus

Trachops, 64

clinedaphus

Monophyllus, 62, 63

clivosus

Rhinolophus, 54

cobourgiana

Tadarida, 68

cobourgianus

Mormopterus, 68

cocos

Galago, 13

coelophyllus

Rhinolophus, 54

Coelops

frithii, 55

hirsutus, 55

inflata, 55

inflatus, 55

robinsoni, 55

coffini

Trachops, 64

cognatus

Sylvilagus, 30

collaris

Lagomys, 21

Ochotona, 21

Colobus

abyssinicus, 16

guereza, 16

matschiei, 16

occidentalis, 16

percivali, 16

roosevelti, 16

terrestris, 16

colombianus

Metachirus, 5

comes

Acratocnus, 8

compressus

Macrotus, 62

concava

Lonchophylla, 61

concolor

Simias, 20

Condylura

cristata, 49

nigra, 49

parva, 49

connectens

Lepus, 28

Sylvilagus, 28

constanzae

Tadarida, 69

contigua

Lasiopyga, 15

continentis

Myotis, 73

convexum

Uroderma, 65

coronatus

Sorex, 42

corvus

Presbytis, 20

Corynorhinus

australis, 70

ingens, 70

macrotis, 70

megalotis, 70

mexicanus, 70

pallescens, 70

townsendii, 70, 71

virginianus, 71

coucang

Nycticebus, 12, 13

coxi

Pteropus, 52

cozumelae

Didelphis, 4

Mimon, 62

cracens

Marmosa, 5

Marmosops, 5

crawfordi

Notiosorex, 40

Sorex, 40

crenulatum

Mimon, 62

crepuscularis

Nycticea, 80

crepusculus

Trachypithecus, 20

cristata

Condylura, 49

Presbytis, 20

cristatus

Trachypithecus, 21

cristobalensis

Sorex, 47

Crocidura

agadiri, 33

alchemillae, 33

aleksandrisi, 33

allex, 33

alpina, 33

altae, 34

andamanensis, 33

attenuata, 34

beatus, 33

bolivari, 33

caudata, 33

chaouianensis, 33

daphnia, 33

elongata, 33

essaouiranensis, 34

fumosa, 33

gmelini, 35

gouliminensis, 34

grandis, 34

grayi, 34

grisea, 34

grisescens, 34

halconus, 34

heljanensis, 34

hildegardeae, 34, 35

horsfieldi, 34

hosletti, 36

kurodai, 34

lakiundae, 37

lea, 34

lepidura, 34

levicula, 34

lignicolor, 35

lipara, 35

littoralis, 35

longipes, 35

lutrella, 35

lutreola, 35

maanjae, 35

mesatanensis, 35

mimula, 35

mindorus, 35

muricauda, 37

mutesae, 35

nicobarica, 35

nigripes, 35

nigrofusca, 36, 37

nilotica, 36

nisa, 36

nyansae, 33

olivieri, 33, 36

parvipes, 35, 36

pergrisea, 36

planiceps, 36

procera, 34

pullata, 36

quelpartis, 36

raineyi, 36

rapax, 34

rhoditis, 36

roosevelti, 37

russula, 33, 34, 36

safii, 34

shantungensis, 36

sicula, 33, 36

simiolus, 36

suahelae, 36

sururae, 36

tamrinensis, 33

tarfayaensis, 33, 34, 36

tiznitensis, 36

turba, 37

viaria, 33, 36

whitakeri, 34, 35, 37

yebalensis, 33, 34

zaianensis, 37

zaphiri, 36

cruciger

Presbytis, 19

Cryptotis

alticola, 37

berlandieri, 37, 39

floridanus, 37

frontalis, 38

goldmani, 38, 39

goodwini, 38

gracilis, 39

griseoventris, 39

guerrerensis, 39

jacksoni, 39

lacertosus, 39

macer, 39

machetes, 38

magnus, 38

mam, 39

mayensis, 38

merriami, 39

merus, 39

mexicanus, 38

nayaritensis, 39

nelsoni, 38

nigrescens, 39, 40

obscurus, 38

oreoryctes, 39

parvus, 37, 38, 39, 40

peregrinus, 38

pergracilis, 39

pueblensis, 39

soricinus, 38

tamensis, 39

zeteki, 40

cubanus

Monophyllus, 62

cultrata

Anoura, 57

cunicularius

Sylvilagus, 30

cupidus

Pithecus, 18

curasoae

Leptonycteris, 61

curtatus

Pipistrellus, 71

cuyonis

Tupaia, 9

CYCLOPEDIDAE, 8

Cyclopes

didactylus, 8

mexicanus, 8
NUMBER 644

• 9 9

cylindrura

Urogale, 11

cynocephala

Nycticea, 69

Tadarida, 69

CYNOCEPHALIDAE, 11

Cynomolgus

apoensis, 16

cagayanus, 16

mindanensis, 16

suluensis, 16

Cynopithecus

lembicus, 16

Cynopterus

angulatus, 51

babi, 51

harpax, 52

horsfieldii, 52

lyoni, 52

major, 51

minutus, 51

pagensis, 51, 52

princeps, 52

serasani, 52

sphinx, 51, 52

titthaecheilus, 51

dalli

Lepus, 24, 26

daphnia

Crocidura, 33

dariensis

Ateles, 14

DASYPODIDAE, 8

Dasypterus

floridanus, 72

Dasypus

mexicanus, 8

novemcinctus, 8

DASYURIDAE, 6

DASYUROMORPHIA, 6

daubentonii

Myotis, 76

debilis

Molossus, 68

degelidus

Lasiurus, 73

degener

Lichonycteris, 61

deignani

Myotis, 74

demararae

Marmosa, 4

Micoureus, 4

Dendrohyrax

arboreus, 7

validus, 7

dentaneus

Metachirus, 5

dentatus

Tarsius, 13

depressa

Kerivoula, 72

derbianus

Caluromys, 3

Dermanura

azteca, 59

gnoma, 59

phaeotis, 59

rava, 59

tolteca, 60

watsoni, 59

DERMOPTERA, 11

deserticola

Lepus, 27

destructioni

Sorex, 48

diazi

Romerolagus, 27

didactylus

Cyclopes, 8

DIDELPHIDAE, 3

DIDELPHIMORPHIA, 3

Didelphis

californica, 4

caucae, 4

cozumelae, 4

marsupialis, 4

richmondi, 4

tabascensis, 4

texensis, 4

virginiana, 4

yucatanensis, 4

dilatus

Scapanus, 51

dimidiata

Peramys, 6

diminutus

Myotis, 74

DIPROTODONTIA, 7

discifera

Thyroptera, 66

discolor

Tupaia, 10

dispar

Sorex, 42

dobsoni

Sorex, 42

dominicensis

Myotis, 74

Natalus, 66

dundasi

Elephantulus, 7

durangae

Sorex, 42

Dymecodon

pilirostris, 49

Dysopes

aurispinosus, 68

ebenus

Semnopithecus, 21

Trachypithecus, 21

Echinosorex

gymnura, 32

Ectophylla

alba, 60

edarata

Tupaia, 10

elassodon

Sorex, 44

elegantulus

Midas, 14

ELEPHANTIDAE, 7

Elephantulus

dundasi, 7

mariakanae, 7

phaeus, 7

rufescens, 7

Elephas

africanus, 7

rothschildi, 7

elisabethae

Vampyressa, 65

elongata

Crocidura, 33

Glossophaga, 60

emarginatus

Sorex, 42

Emballonura

alecto, 56

anambensis, 56

monticola, 56

peninsularis, 56

pusilla, 56

semicaudata, 57

sulcata, 57

EMBALLONURIDAE, 56

engana

Kerivoula, 72

enganus

Pteropus, 52

Eonycteris

robusta, 52

ephippiger

Bradypus, 8

Eptesicus

bahamensis, 71

fuscus, 71

hispaniolae, 71

lynni, 71

miradorensis, 71

pallens, 71

pallidus, 71

serotinus, 71

ugandae, 79

wetmorei, 71

ERINACEIDAE, 32

ERINACEOMORPHA, 32

Erinaceus

sotikae, 32

Erophylla

bombifrons, 60

mariguanensis, 60

planifrons, 60

sezekorni, 60

Erythrocebus

patas, 16

whitei, 16

erythrotis

Ochotona, 22

esau

Macaca, 19

espiritosantensis

Natalus, 67

essaouiranensis

Crocidura, 34

Eumops

amazonicus, 67

californicus, 67

glaucinus, 68

hansae, 67

perotis, 67

euronotus

Sorex, 42

europaeus

Lepus, 25, 26

europs

Nyctinomops, 69

Nyctinomus, 69

Euroscaptor

grandis, 50

parvidens, 50

everetti

Tupaia, 11

evotis

Myotis, 75, 77

Notiosorex, 40

Sorex, 40

Vespertilio, 77

exiguous

Vespertilio, 77

exiguus

Sylvilagus, 30

exilipes

Blarina, 37

exilis

Vespertilio, 77, 79

eximius

Blarina, 37

Sorex, 43

extremus

Myotis, 75

fascicularis

Macaca, 16, 17, 18, 19

faunulus

Pteropus, 53

femoralis

Presbytis, 19, 20

femorosaccus

Nyctinomops, 69

Nyctinomus, 69

fenisex

Ochotona, 22

ferreus

Monophyllus, 62

ferruginea

Tupaia, 10, 11

festinus

Lepus, 25

filchneri

Lepus, 26

fimbriatus

Myotis, 75

fimbripes

Sorex, 43
100 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

findleyi

Myotis, 74

fisheri

Sorex, 43

flavicauda

Pygathrix, 21

flavicaudus

Trachypithecus, 21

flavus

Myotis, 74

floridana

Blarina, 37

floridanus

Cryptotis, 37

Dasypterus, 72

Lasiurus, 72

Lepus, 28, 29

Sylvilagus, 28, 29, 31

fodiens

Neomys, 40

fontinalis

Sorex, 43

formosa

Chacodelphys, 3

Marmosa, 3

formosus

Myotis, 74

fornicata

Lonchophylla, 61

fortidens

Myotis, 76

fortis

Molossus, 68

fossor

Blarina, 37

fraterculus

Carlito, 13

Tarsius, 13

frithii

Coelops, 55

frons

Lavia, 56

frontalis

Cryptotis, 38

fumeus

Sorex, 43

fumosa

Crocidura, 33

Ochotona, 22

fumosus

Vampyrops, 63

furvus

Ailurops, 7

Phalanger, 7

fusca

Macaca, 17

Murina, 73

fuscior

Tupaia, 11

fuscipes

Ateles, 14

Ochotona, 23

fusciventris

Platyrrhinus, 63

fuscogriseus

Metachirus, 6

Philander, 6

fuscus

Eptesicus, 71

Macacus, 17

Murina, 73

Pithecus, 18

Tarsius, 13

Vespertilio, 71

gabbi

Lepus, 28

Sylvilagus, 28, 30, 31

gaillardi

Lepus, 25

GALAGIDAE, 13

Galago

cocos, 13

moholi, 13

senegalensis, 13

sotikae, 13

Galeopithecus

aoris, 11

gracilis, 12

natunae, 12

pumilus, 12

saturatus, 12

tuancus, 12

Galeopterus

abbotti, 12

borneanus, 12

chombolis, 12

lautensis, 12

variegatus, 11, 12

galeritus

Hipposideros, 55

geminorum

Pteropus, 53

gemmeus

Sylvisorex, 49

gentilis

Hipposideros, 55

geoffroyi

Ateles, 15

germaini

Trachypithecus, 20

gibbsi

Neurotrichus, 50

gibbsii

Neurotrichus, 50

Urotrichus, 50

gigas

Megasorex, 40

Notiosorex, 40

gilberti

Hipposideros, 55

glacialis

Sorex, 43

glaucinus

Eumops, 68

glis

Tupaia, 9, 10, 11

Glossophaga

campestris, 60

elongata, 60

longirostris, 60

microtis, 60

mutica, 60

rostrata, 60

soricina, 60, 61

truei, 61

villosa, 61

gmelini

Crocidura, 35

gnoma

Dermanura, 59

gnomus

Artibeus, 59

godmani

Sorex, 43

goldmani

Blarina, 38

Cryptotis, 38, 39

Lepus, 27

Ochotona, 23

Sylvilagus, 27

gonavensis

Chilonycteris, 65

Pteronotus, 65

goodwini

Cryptotis, 38

gorgasi

Chiroderma, 59

gouliminensis

Crocidura, 34

Gracilinanus

marica, 4

perijae, 4

gracilis

Cryptotis, 39

Galeopithecus, 12

Rhogeessa, 81

Tupaia, 10

grahami

Lepus, 25

grandis

Crocidura, 34

Euroscaptor, 50

grayi

Crocidura, 34

graysoni

Lepus, 29

Sylvilagus, 29, 31

greenhalli

Centurio, 59

grisea

Crocidura, 34

griseoventris

Cryptotis, 39

grisescens

Crocidura, 34

Myotis, 74

griseus

Lepus, 26

groenlandicus

Lepus, 24

gryphus

Vespertilio, 77

guereza

Colobus, 16

guerrerensis

Cryptotis, 39

gymnura

Echinosorex, 32

Gymnura, 32

Gymnura

gymnura, 32

minor, 32

halconus

Crocidura, 34

handleyi

Myotis, 74

hansae

Eumops, 67

hardwickii

Kerivoula, 72

harpax

Cynopterus, 52

Harpyionycteris

celebensis, 52

harrisoni

Rhinopoma, 56

hasseltii

Myotis, 73

haydeni

Sorex, 43

heberfolium

Macrotus, 62

hefneri

Sylvilagus, 31

helios

Neoromicia, 80

Pipistrellus, 80

Heliosorex

roosevelti, 37

heljanensis

Crocidura, 34

Hemiderma

subrufum, 58

tricolor, 58

henshawii

Vespertilio, 78

hercynicus

Sorex, 41

hesperus

Parastrellus, 80, 81

Pipistrellus, 80

Scotophilus, 80

Heterohyrax

albipes, 7

brucei, 7

hildegardeae

Crocidura, 34, 35

hilgerti

Chlorocebus, 15

HIPPOSIDERIDAE, 55

Hipposideros

ater, 55

barbensis, 55

bicolor, 55

galeritus, 55

gentilis, 55
NUMBER 644

• 101

gilberti, 55

insolens, 55

larvatus, 55

nicobarulae, 55

pelingensis, 55

saevus, 55

toala, 55

hirsuta

Chilophylla, 55

hirsutus

Artibeus, 58

Coelops, 55

Myotis, 75

Rhinolophus, 54

hispaniolae

Eptesicus, 71

Histiotus

humboldti, 71

hitchensi

Sylvilagus, 31

hollisteri

Sorex, 46

holzneri

Lepus, 29

Sylvilagus, 29

hondurensis

Sylvilagus, 31

horsfieldi

Crocidura, 34

horsfieldii

Cynopterus, 52

Myotis, 74

hosei

Presbytis, 20

hosletti

Crocidura, 36

howelli

Scalopus, 50

hoyi

Microsorex, 40, 41

Sorex, 40, 41, 43, 48

hubbardi

Lepus, 24

humboldtensis

Sorex, 48

humboldti

Histiotus, 71

humeralis

Nycticeius, 80

huttoni

Murina, 73

hydrobadistes

Sorex, 46

Hylobates

abbotti, 21

albibarbis, 21

cinereus, 21

klossii, 21

muelleri, 21

mulleri, 21

HYLOBATIDAE, 21

hypomelanus

Pteropus, 52, 53

hypomicrus

Nesophontes, 32

Hypsugo

macrotis, 71

HYRACOIDEA, 7

idahoensis

Brachylagus, 23

Lepus, 23

Sorex, 43

ignavus

Bradypus, 8

impavidus

Marmosops, 5

impudens

Pithecus, 18

inca

Lestoros, 6

Orolestes, 6

incana

Ochotona, 23

incomitatus

Artibeus, 59

inconsonans

Alouatta, 14

indochinensis

Macaca, 17

inflata

Coelops, 55

Tupaia, 10

inflatus

Coelops, 55

Scalopus, 50

infuscus

Platyrrhinus, 63

ingens

Corynorhinus, 70

inops

Rhinolophus, 54

insolens

Hipposideros, 55

insonus

Lepus, 29

Sylvilagus, 29

insulana

Macaca, 17

insularis

Marmosa, 6

Pteropus, 53

interior

Myotis, 75

intermedia

Mormoops, 65

Tadarida, 69, 70

intermedius

Artibeus, 58

Lasiurus, 72, 73

intervectus

Microsorex, 40

invicta

Marmosa, 5

invictus

Marmosops, 5

irus

Macaca, 17

isolatus

Sorex, 45

isthmica

Marmosa, 4

isthmicum

Chiroderma, 59

itatiayae

Peramys, 6

jacksoni

Cryptotis, 39

jagorii

Phoniscus, 81

jamaicensis

Artibeus, 58

Macrotus, 62

javanicus

Pipistrellus, 81

jesupi

Chiroderma, 59

jewetti

Ochotona, 23

jobensis

Chaerephon, 67

Nyctinomus, 67

juncensis

Sorex, 42

kaimosae

Lasiopyga, 15

kakumegae

Lepus, 25

kalkoae

Lophostoma, 61

keenani

Mimon, 62

keenii

Myotis, 79

Vespertilio, 79

keniensis

Rhinolophus, 54

Kerivoula

bombifrons, 72

depressa, 72

engana, 72

hardwickii, 72

minuta, 72

pellucida, 72

kibonotensis

Cercopithecus, 15

kima

Lasiopyga, 15

klamathensis

Lepus, 25

klossii

Hylobates, 21

Symphalangus, 21

knoxjonesi

Blarina, 32

kohtauensis

Tupaia, 10

konganensis

Sylvisorex, 49

koopmani

Artibeus, 58

koratensis

Presbytis, 20

kunsi

Monodelphis, 6

kurodai

Crocidura, 34

labiatus

Saguinas, 14

labradorius

Lepus, 25

lacernata

Tupaia, 10

lacertosus

Cryptotis, 39

Laephotis

botswanae, 72

namibensis, 72

LAGOMORPHA, 21

Lagomys

collaris, 21

schisticeps, 21

lagunae

Sorex, 44

lakiundae

Crocidura, 37

lanensis

Pteropus, 53

lanigera

Pteropus, 53

lapsus

Pithecus, 18

laricorum

Sorex, 42

larvatus

Hipposideros, 55

lasiae

Macaca, 18

Megaderma, 56

Pithecus, 18

Lasiopyga

albogularis, 15

arenaria, 15

ascanius, 15

callida, 15

contigua, 15

kaimosae, 15

kima, 15

leucampax, 15

maritima, 15

mauae, 15

pygerythra, 15

tumbili, 15

Lasiurus

blossevillii, 72

borealis, 72

castaneus, 73

cinereus, 72

degelidus, 73

floridanus, 72

intermedius, 72, 73

minor, 73

ornatus, 72

semotus, 72

teliotis, 72

laticaudatus

Nyctinomops, 68, 69
102 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

latidens

Anoura, 57

latimanus

Scapanus, 51

latus

Vampyrops, 63, 64

lautensis

Galeopterus, 12

Pithecus, 18

Lavia

frons, 56

rex, 56

lea

Crocidura, 34

leibii

Myotis, 76

lembicus

Cynopithecus, 16

lemhi

Ochotona, 23

leonina

Macaca, 16, 17

lepidura

Crocidura, 34

lepidus

Pteropus, 53

LEPORIDAE, 23

leporinus

Noctilio, 66

Leptonycteris

curasoae, 61

Lepus

abbotti, 25

aegyptius, 25

alleni, 23

altamirae, 26

americanus, 24, 25, 26, 27

angustidens, 24

arcticus, 24, 25

arizonae, 27, 29

asellus, 24

audubonii, 28

aurigineus, 24

bachmani, 28

baileyi, 28

bairdii, 24

bangsii, 25

brachyurus, 24

brasiliensis, 28

californicus, 24, 25, 26, 27

callotis, 25

campestris, 25

caniclunis, 28

canus, 24

capensis, 25

chiapensis, 28

cinnamomeus, 25

connectens, 28

dalli, 24, 26

deserticola, 27

europaeus, 25, 26

festinus, 25

filchneri, 26

floridanus, 28, 29

gabbi, 28

gaillardi, 25

goldmani, 27

grahami, 25

graysoni, 29

griseus, 26

groenlandicus, 24

holzneri, 29

hubbardi, 24

idahoensis, 23

insonus, 29

kakumegae, 25

klamathensis, 25

labradorius, 25

macfarlani, 24

magdalenae, 25

major, 27

margaritae, 29

melanotis, 26

merriami, 26

microtis, 25, 26

minor, 27

monstrabilis, 24

oistolus, 25

orizabae, 29

othus, 26

pacificus, 30

parnassius, 26

pinetis, 29

poadromus, 26

porsildi, 24

quercerus, 26

raineyi, 26

rigidus, 29

robustus, 29

saliens, 26

sanctidiegi, 28

sierrae, 25

simplicicanus, 29

sowerbyae, 26

stoliczkanus, 26

subcinctus, 28

swinhoei, 26

sylvaticus, 29

texianus, 24, 25, 26, 27

tibetanus, 26

tiburonensis, 23

timidus, 27

tolai, 24, 25, 26

townsendii, 25

trowbridgii, 29

truei, 30

tularensis, 27

ubericolor, 28

varronis, 27

veraecrucis, 30

wallawalla, 27

washingtonii, 25, 27

xanti, 25

yucatanicus, 29

lestes

Papio, 19

Lestoros

inca, 6

leucampax

Lasiopyga, 15

leucogenys

Sorex, 44

levicula

Crocidura, 34

levis

Ochotona, 22

Lichonycteris

degener, 61

lignicolor

Crocidura, 35

lilium

Sturnira, 64

limitaneus

Cebus, 14

lingae

Pithecus, 18

Tana, 9

Tupaia, 9

lingungensis

Pithecus, 18

lipara

Crocidura, 35

littoralis

Crocidura, 35

Sylvilagus, 30

lituratus

Artibeus, 58

lixa

Pachyura, 49

Lonchophylla

cadenai, 61

concava, 61

fornicata, 61

robusta, 61

longicauda

Sorex, 44, 45

longicaudus

Sorex, 45

longicrus

Myotis, 73, 75, 77

Vespertilio, 77

longipes

Crocidura, 35

Tupaia, 10

longirostris

Glossophaga, 60

Sorex, 43

Lophostoma

brasiliense, 61

kalkoae, 61

venezuelae, 61

loriae

Mormopterus, 68

Tadarida, 68

LORISIDAE, 12

loukashkini

Myotis, 76

Loxodonta

africana, 7

luciae

Ardops, 57

Monophyllus, 63

Stenoderma, 57

lucifugus

Myotis, 75, 76, 77

Vespertilio, 77

lutescens

Ochotona, 22

luteus

Cercopithecus, 15

lutosus

Myotis, 76

lutrella

Crocidura, 35

lutreola

Crocidura, 35

luzonus

Chaerephon, 67

lyelli

Sorex, 47

lynni

Eptesicus, 71

lyoni

Cynopterus, 52

maanjae

Crocidura, 35

Macaca

adusta, 16

atriceps, 17

aurea, 19

broca, 16

esau, 19

fascicularis, 16, 17, 18, 19

fusca, 17

indochinensis, 17

insulana, 17

irus, 17

lasiae, 18

leonina, 16, 17

mulatta, 17

nemestrina, 16, 17

nigra, 16

pagensis, 17

philippensis, 16, 19

thibetana, 19

tua, 17

umbrosa, 17

Macacus

fuscus, 17

pagensis, 17

phaeura, 17

pumilus, 17

rhesus, 17

umbrosus, 17

villosus, 17

macer

Chilonatalus, 66

Cryptotis, 39

macfarlani

Lepus, 24

machetes

Blarina, 38

Cryptotis, 38

machrinoides

Scalopus, 50

macrocorpus

Sylvilagus, 31

macrodon

Sorex, 44
NUMBER 644

• 103

macropus

Vespertilio, 77

macropygmaeus

Sorex, 44

MACROSCELIDEA, 7

MACROSCELIDIDAE, 7

macrotis

Corynorhinus, 70

Hypsugo, 71

Nyctinomops, 69

Nyctinomus, 69

Rhinolophus, 54

Macrotus

californicus, 61

compressus, 62

heberfolium, 62

jamaicensis, 62

waterhousii, 62

macrourus

Natalus, 67

magdalenae

Lepus, 25

magna

Blarina, 38

magnus

Cryptotis, 38

major

Cynopterus, 51

Lepus, 27

Natalus, 66

Neurotrichus, 50

Thyroptera, 66

mallurus

Sylvilagus, 31

mam

Cryptotis, 39

Manatus

oweni, 8

mandibularis

Pithecus, 18

mansalaris

Pithecus, 19

mansuetus

Sylvilagus, 31

margaritae

Cebus, 14

Lepus, 29

Sapajus, 14

Sylvilagus, 29

mariakanae

Elephantulus, 7

marica

Gracilinanus, 4

mariguanensis

Erophylla, 60

maritima

Lasiopyga, 15

Marmosa

albiventris, 5

bishopi, 5

caucae, 5

cracens, 5

demararae, 4

formosa, 3

insularis, 6

invicta, 5

isthmica, 4

meridae, 4

mexicana, 4

murina, 4

muscula, 3

oaxacae, 6

parvidens, 5

pinheiroi, 5

purui, 5

ruatanica, 4

tatei, 6

woodalli, 5

xerophila, 5

zeledoni, 5

Marmosops

bishopi, 5

cracens, 5

impavidus, 5

invictus, 5

neblina, 5

noctivagus, 5

pinheiroi, 5

marsupialis

Didelphis, 4

masae

Tana, 9

Tupaia, 9

mastivus

Noctilio, 66

matschiei

Colobus, 16

mauae

Lasiopyga, 15

maximus

Parastrellus, 80

Pipistrellus, 80

mayensis

Blarina, 38

Cryptotis, 38

mcmurtrii

Centurio, 58

mearnsi

Pteropus, 53

mearnsii

Sylvilagus, 31

Megaderma

carimatae, 56

celebensis, 56

lasiae, 56

niasense, 56

siumatis, 56

spasma, 56

MEGADERMATIDAE, 56

MEGALONYCHIDAE, 8

megalophylla

Mormoops, 65

megalotis

Corynorhinus, 70

Micronycteris, 62

megalura

Sylvisorex, 49

Megasorex

gigas, 40

mehelyi

Rhinolophus, 54

melalophos

Presbytis, 19

melanopogon

Taphozous, 57

melanops

Peramys, 6

melanorhinus

Myotis, 78

Vespertilio, 78

melanotis

Lepus, 26

melanotus

Pteropus, 53

meridae

Marmosa, 4

merriami

Cryptotis, 39

Lepus, 26

Sorex, 44

Vesperugo, 81

merus

Cryptotis, 39

mesatanensis

Crocidura, 35

messorius

Sylvilagus, 31

Metachirus

colombianus, 5

dentaneus, 5

fuscogriseus, 6

nudicaudatus, 5

pallidus, 6

mexicana

Alouatta, 14

Blarina, 38

Chilonycteris, 65

Marmosa, 4

Tadarida, 69

mexicanus

Corynorhinus, 70

Cryptotis, 38

Cyclopes, 8

Dasypus, 8

Micronycteris, 62

Myotis, 76

Natalus, 66

Noctilio, 66

Pteronotus, 65

Micoureus

demararae, 4

Micronycteris

megalotis, 62

mexicanus, 62

microtis, 62

sanborni, 62

micronyx

Myotis, 75

microphyllum

Rhinopoma, 56

micropus

Chilonatalus, 66

Microsorex

hoyi, 40, 41

intervectus, 40

montanus, 41

washingtoni, 41

winnemana, 41

microtis

Glossophaga, 60

Lepus, 25, 26

Micronycteris, 62

Midas

elegantulus, 14

milleri

Sorex, 44

Mimon

cozumelae, 62

crenulatum, 62

keenani, 62

mimula

Crocidura, 35

mindanensis

Cynomolgus, 16

mindorus

Crocidura, 35

Pithecus, 19

minimus

Chironectes, 3

Miniopterus

arenarius, 73

natalensis, 73

paululus, 73

minor

Antrozous, 70

Brachyphylla, 58

Gymnura, 32

Lasiurus, 73

Lepus, 27

Neomys, 40

Niadias, 52

Sylvilagus, 27

Tupaia, 11

minusculus

Neoromicia, 80

Pipistrellus, 80

minuta

Kerivoula, 72

Vampyressa, 65

minutilla

Rhogeessa, 82

minutillus

Rhinolophus, 54

minutus

Cynopterus, 51

Rhinolophus, 54

miradorensis

Eptesicus, 71

Scotophilus, 71

mitis

Cercopithecus, 15

moellendorffi

Tupaia, 9

mohavensis

Nyctinomus, 69

moholi

Galago, 13

MOLOSSIDAE, 67
104 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

molossus

Molossus, 68

Molossus

californicus, 67

debilis, 68

fortis, 68

molossus, 68

nigricans, 68

pretiosus, 68

pygmaeus, 68

rufus, 68

Monodelphis

adusta, 6

kunsi, 6

sorex, 6

Monophyllus

clinedaphus, 62, 63

cubanus, 62

ferreus, 62

luciae, 63

plethodon, 63

portoricensis, 63

redmani, 62, 63

monstrabilis

Lepus, 24

montanus

Microsorex, 41

Sorex, 41

montereyensis

Sorex, 44

monticola

Emballonura, 56

Sorex, 41, 42, 43, 44, 45, 48, 

49

monticolus

Sorex, 44

moorei

Ochotona, 22

mordax

Sturnira, 64

Sturnirops, 64

MORMOOPIDAE, 65

Mormoops

intermedia, 65

megalophylla, 65

senicula, 65

tumidiceps, 65

Mormopterus

cobourgianus, 68

loriae, 68

phrudus, 68

muelleri

Hylobates, 21

mulatta

Macaca, 17

mulleri

Hylobates, 21

mumfordi

Myotis, 73

mundus

Vespertilio, 78

muricauda

Crocidura, 37

Myosorex, 37

muricola

Myotis, 75

murina

Marmosa, 4

Murina

fusca, 73

fuscus, 73

huttoni, 73

murinus

Suncus, 49

muscula

Marmosa, 3

mutabilis

Sorex, 47

mutesae

Crocidura, 35

mutica

Glossophaga, 60

myops

Sorex, 48

Myosorex

muricauda, 37

Myotis

abbotti, 73

adversus, 73, 74

alascensis, 75

altifrons, 73

amotus, 75

austroriparius, 73, 74

baileyi, 74

californicus, 74, 76, 77, 78, 79

capitaneus, 74

carimatae, 74

caurinus, 74, 78

ciliolabrum, 77

continentis, 73

daubentonii, 76

deignani, 74

diminutus, 74

dominicensis, 74

evotis, 75, 77

extremus, 75

fimbriatus, 75

findleyi, 74

flavus, 74

formosus, 74

fortidens, 76

grisescens, 74

handleyi, 74

hasseltii, 73

hirsutus, 75

horsfieldii, 74

interior, 75

keenii, 79

leibii, 76

longicrus, 73, 75, 77

loukashkini, 76

lucifugus, 75, 76, 77

lutosus, 76

melanorhinus, 78

mexicanus, 76

micronyx, 75

mumfordi, 73

muricola, 75

nesopolus, 75

niasensis, 75

nigricans, 75, 77, 78

occultus, 74, 75

pallidus, 74

pernox, 75

petax, 75, 76

phasma, 75

quebecensis, 73

riparius, 76

ruber, 79

saturatus, 76

septentrionalis, 77

siligorensis, 76

simus, 76

sowerbyi, 76

stephensi, 74

thaianus, 76

thysanodes, 76

volans, 73, 74, 75, 77, 79

watasei, 74

winnemana, 76

yumanensis, 75, 76, 77, 78, 79

Myrmecophaga

centralis, 8

tridactyla, 8

MYRMECOPHAGIDAE, 8

naivashae

Chaerephon, 67

namibensis

Laephotis, 72

nana

Brachyphylla, 58

nanus

Neoromicia, 80

Sorex, 48

nasutus

Promops, 69

natalensis

Miniopterus, 73

Natalus, 66

NATALIDAE, 66

Natalus

brevimanus, 66

dominicensis, 66

espiritosantensis, 67

macrourus, 67

major, 66

mexicanus, 66

natalensis, 66

stramineus, 66

tumidirostris, 67

natunae

Galeopithecus, 12

Nycticebus, 13

Tupaia, 10

navigator

Neosorex, 41

Sorex, 41, 45

nayaritensis

Cryptotis, 39

neblina

Marmosops, 5

nelsoni

Blarina, 38

Cryptotis, 38

Romerolagus, 27

Vampyrus, 65

nemestrina

Macaca, 16, 17

neomexicanus

Sorex, 45

Sylvilagus, 30

Neomys

fodiens, 40

minor, 40

Neoromicia

helios, 80

minusculus, 80

nanus, 80

somalicus, 79

ugandae, 79

Neosorex

albibarbis, 41

navigator, 41

nereis

Rhinolophus, 54

nesites

Rhinolophus, 54

Nesophontes

hypomicrus, 32

paramicrus, 32

zamicrus, 32

NESOPHONTIDAE, 32

nesopolus

Myotis, 75

Neurotrichus

gibbsi, 50

gibbsii, 50

major, 50

nevadensis

Nyctinomus, 69

Ochotona, 23

Sorex, 45

Niadias

minor, 52

niadicus

Pteropus, 53

niasense

Megaderma, 56

niasensis

Myotis, 75

Rhinolophus, 55

nichollsi

Ardops, 57

nicobarica

Crocidura, 35

Tupaia, 10

nicobarulae

Hipposideros, 55

nigra

Condylura, 49

Macaca, 16

nigrescens

Cryptotis, 39, 40

Ochotona, 22

nigricans

Molossus, 68

Myotis, 75, 77, 78
NUMBER 644

• 105

nigripes

Crocidura, 35

nigrofusca

Crocidura, 36, 37

nilotica

Crocidura, 36

nisa

Crocidura, 36

nitelinea

Platyrrhinus, 63

nitida

Ochotona, 22

nitidus

Vespertilio, 78, 79

Noctilio

leporinus, 66

mastivus, 66

mexicanus, 66

NOCTILIONIDAE, 66

noctivagus

Marmosops, 5

Notiosorex

crawfordi, 40

evotis, 40

gigas, 40

novemcinctum

Tatu, 8

novemcinctus

Dasypus, 8

nucella

Scotophilus, 82

nudicaudatus

Metachirus, 5

nyansae

Crocidura, 33

Nycticea

crepuscularis, 80

cynocephala, 69

Nycticebus

bancanus, 12

borneanus, 12

brachycephalus, 12

coucang, 12, 13

natunae, 13

Nycticeius

humeralis, 80

Nyctinomops

aurispinosus, 68

europs, 69

femorosaccus, 69

laticaudatus, 68, 69

macrotis, 69

yucatanicus, 68

Nyctinomus

antillularum, 69

europs, 69

femorosaccus, 69

jobensis, 67

macrotis, 69

mohavensis, 69

nevadensis, 69

orthotis, 68

pusillus, 67

Nyctophilus

arnhemensis, 80

oaxacae

Marmosa, 6

Sorex, 47

obscura

Blarina, 38

obscuroides

Sorex, 48

obscurus

Antrozous, 70

Cryptotis, 38

Pipistrellus, 81

Sorex, 45, 48, 49

Sylvilagus, 31

Trachypithecus, 20, 21

Vespertilio, 78

obtusa

Phyllonycteris, 63

occidentalis

Colobus, 16

occultidens

Pachyura, 49

occultus

Myotis, 74, 75

Ochotona

alpina, 22

annectens, 22

argentata, 22

brooksi, 22

brunnescens, 22

cansus, 22

collaris, 21

erythrotis, 22

fenisex, 22

fumosa, 22

fuscipes, 23

goldmani, 23

incana, 23

jewetti, 23

lemhi, 23

levis, 22

lutescens, 22

moorei, 22

nevadensis, 23

nigrescens, 22

nitida, 22

princeps, 21, 22, 23

saxatilis, 22, 23

schisticeps, 21, 23

uinta, 23

ventorum, 23

vulpina, 22

OCHOTONIDAE, 21

oistolus

Lepus, 25

oklahomae

Pipistrellus, 80

olivacea

Tupaia, 10

olivieri

Crocidura, 33, 36

opossum

Philander, 6

orarius

Scapanus, 51

oregonensis

Vespertilio, 78, 79

oreopolus

Sorex, 45

oreoryctes

Cryptotis, 39

orizabae

Lepus, 29

Sorex, 45

Sylvilagus, 28, 29, 31

ornatus

Ateles, 15

Lasiurus, 72

Sorex, 42, 44, 45

Orolestes

inca, 6

orthotis

Nyctinomus, 68

othus

Lepus, 26

oweni

Manatus, 8

Pachyura

aequatoria, 49

lixa, 49

occultidens, 49

pachyurus

Sorex, 45

pacificus

Antrozous, 70

Lepus, 30

Sorex, 46, 49

pagensis

Cynopterus, 51, 52

Macaca, 17

Macacus, 17

pallens

Eptesicus, 71

pallescens

Corynorhinus, 70

palliata

Alouatta, 14

pallidus

Antrozous, 70

Eptesicus, 71

Metachirus, 6

Myotis, 74

Vespertilio, 70

palmeri

Sorex, 42

palustris

Sorex, 46

Sylvilagus, 31

pamana

Promops, 69

panamensis

Ateles, 15

Chironectes, 3

Papio

anubis, 19

lestes, 19

vigilis, 19

paramicrus

Nesophontes, 32

Parastrellus

hesperus, 80, 81

maximus, 80

parnassius

Lepus, 26

parnellii

Chilonycteris, 65

Pteronotus, 65

Parocnus

serus, 8

parva

Condylura, 49

parvidens

Cheiromeles, 67

Euroscaptor, 50

Marmosa, 5

Sorex, 45

Sturnira, 64

Talpa, 50

parvipes

Crocidura, 35, 36

parvula

Rhogeessa, 82

parvus

Cryptotis, 37, 38, 39, 40

patas

Erythrocebus, 16

PAUCITUBERCULATA, 6

paulsoni

Sturnira, 64

paululus

Miniopterus, 73

pelagicus

Pteropus, 53

pelingensis

Hipposideros, 55

pellucida

Kerivoula, 72

pemangilis

Tupaia, 10

peninsulae

Blarina, 32

peninsularis

Emballonura, 56

Peramys

dimidiata, 6

itatiayae, 6

melanops, 6

percivali

Colobus, 16

percura

Presbytis, 20

peregrina

Blarina, 38

peregrinus

Cryptotis, 38

pergracilis

Cryptotis, 39

pergrisea

Crocidura, 36

perijae

Gracilinanus, 4

Perimyotis

subflavus, 81
106 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

periosus

Choeroniscus, 59

permiliensis

Sorex, 45

pernox

Myotis, 75

perotis

Eumops, 67

personatus

Sorex, 46

perspicillata

Carollia, 58

petax

Myotis, 75, 76

Petrodromus

sangi, 7

sultani, 7

tetradactylus, 7

phaeotis

Dermanura, 59

phaeura

Macacus, 17

Tupaia, 10

phaeus

Elephantulus, 7

Phalanger

furvus, 7

PHALANGERIDAE, 7

phasma

Myotis, 75

phayrei

Trachypithecus, 20

Philander

centralis, 3

fuscogriseus, 6

opossum, 6

philippensis

Macaca, 16, 19

philippinensis

Taphozous, 57

Phoniscus

atrox, 81

jagorii, 81

rapax, 81

phrudus

Mormopterus, 68

Phyllonycteris

aphylla, 63

bombifrons, 60

obtusa, 63

planifrons, 60

poeyi, 63

PHYLLOSTOMIDAE, 57

picta

Tupaia, 11

pilirostris

Dymecodon, 49

PILOSA, 8

pinetis

Lepus, 29

pinheiroi

Marmosa, 5

Marmosops, 5

Pipistrellus

aero, 81

australis, 80

camortae, 81

cinnamomeus, 76

curtatus, 71

helios, 80

hesperus, 80

javanicus, 81

maximus, 80

minusculus, 80

obscurus, 81

oklahomae, 80

santarosae, 80

subflavus, 81

subulidens, 81

tenuis, 81

Pithecus

agnatus, 17

baweanus, 17

capitalis, 18

carimatae, 18

cupidus, 18

fuscus, 18

impudens, 18

lapsus, 18

lasiae, 18

lautensis, 18

lingae, 18

lingungensis, 18

mandibularis, 18

mansalaris, 19

mindorus, 19

pullus, 19

sirhassenensis, 19

vitis, 19

planiceps

Crocidura, 36

Sorex, 46

planifrons

Erophylla, 60

Phyllonycteris, 60

Platyrrhinus

aquilus, 63

aurarius, 63

brachycephalus, 63,64

fusciventris, 63

infuscus, 63

nitelinea, 63

saccharus, 64

pleei

Cebus, 14

plethodon

Monophyllus, 63

plicatus

Chaerephon, 67

pluto

Saccolaimus, 57

Taphozous, 57

poadromus

Lepus, 26

Podogymnura

truei, 32

poeyi

Phyllonycteris, 63

polulus

Surdisorex, 49

porsildi

Lepus, 24

portoricensis

Chilonycteris, 65

Monophyllus, 63

Pteronotus, 65

praeceps

Artibeus, 58

preblei

Sorex, 46

Presbytes

batuanus, 19

rhionis, 19

Presbytis

argenteus, 20

arwasca, 19

australis, 19

cana, 19, 20

canicrus, 20

carimatae, 20

catemana, 20

chrysomelas, 19

corvus, 20

cristata, 20

cruciger, 19

femoralis, 19, 20

hosei, 20

koratensis, 20

melalophos, 19

percura, 20

rhionis, 19

rubicunda, 20

rubida, 20

siamensis, 19, 20

sumatrana, 19

vigilans, 21

pretiosus

Molossus, 68

prevostensis

Sorex, 44

pribilofensis

Sorex, 46

PRIMATES, 12

princeps

Cynopterus, 52

Ochotona, 21, 22, 23

PROBOSCIDEA, 7

PROCAVIIDAE, 7

procera

Crocidura, 34

Promops

nasutus, 69

pamana, 69

Pseudantechinus

bilarni, 6

Pteronotus

gonavensis, 65

mexicanus, 65

parnellii, 65

portoricensis, 65

PTEROPODIDAE, 51

Pteropus

aldabrensis, 52

alecto, 52

arquatus, 51

aterrimus, 52

balutus, 52

baveanus, 52

cagayanus, 52

coxi, 52

enganus, 52

faunulus, 53

geminorum, 53

hypomelanus, 52, 53

insularis, 53

lanensis, 53

lanigera, 53

lepidus, 53

mearnsi, 53

melanotus, 53

niadicus, 53

pelagicus, 53

pumilus, 52, 53

samoensis, 53

speciosus, 53

vampyrus, 53

vociferus, 53

pueblensis

Cryptotis, 39

pulcher

Scalopus, 50

pullata

Crocidura, 36

pullus

Pithecus, 19

pulonis

Tupaia, 11

pumila

Brachyphylla, 58

pumilus

Chaerephon, 67

Galeopithecus, 12

Macacus, 17

Pteropus, 52, 53

Tarsius, 13

purui

Marmosa, 5

pusilla

Emballonura, 56

pusillus

Chaerephon, 67

Nyctinomus, 67

Rhinolophus, 54

Pygathrix

flavicauda, 21

rubicunda, 20

rubida, 20

sanctorum, 21

ultima, 21

pygerythra

Lasiopyga, 15

pygerythrus

Chlorocebus, 15

pygmaeus

Bradypus, 8

Molossus, 68

quebecensis

Myotis, 73
NUMBER 644

• 107

quelpartis

Crocidura, 36

quercerus

Lepus, 26

raineyi

Crocidura, 36

Lepus, 26

rapax

Crocidura, 34

Phoniscus, 81

rava

Dermanura, 59

raviana

Tupaia, 11

redmani

Monophyllus, 62, 63

Reithronycteris

aphylla, 63

restrictus

Sylvilagus, 31

rex

Lavia, 56

rhesus

Macacus, 17

RHINOLOPHIDAE, 54

Rhinolophus

acuminatus, 54

affinis, 54

borneensis, 55

circe, 54

clivosus, 54

coelophyllus, 54

hirsutus, 54

inops, 54

keniensis, 54

macrotis, 54

mehelyi, 54

minutillus, 54

minutus, 54

nereis, 54

nesites, 54

niasensis, 55

pusillus, 54

shameli, 54

solitarius, 54

spadix, 55

trifoliatus, 54, 55

tuneti, 54

virgo, 55

Rhinophylla

alethina, 64

Rhinopoma

harrisoni, 56

microphyllum, 56

RHINOPOMATIDAE, 56

rhionis

Presbytes, 19

Presbytis, 19

rhoditis

Crocidura, 36

Rhogeessa

gracilis, 81

minutilla, 82

parvula, 82

tumida, 82

riabus

Tupaia, 11

richmondi

Didelphis, 4

rigidus

Lepus, 29

riparius

Myotis, 76

robinsoni

Coelops, 55

roboratus

Sorex, 46

robusta

Eonycteris, 52

Lonchophylla, 61

robustus

Lepus, 29

Sylvilagus, 29

Romerolagus

diazi, 27

nelsoni, 27

roosevelti

Colobus, 16

Crocidura, 37

Heliosorex, 37

rostrata

Glossophaga, 60

rothschildi

Elephas, 7

ruatanica

Marmosa, 4

ruber

Myotis, 79

Vespertilio, 79

rubicunda

Presbytis, 20

Pygathrix, 20

rubida

Presbytis, 20

Pygathrix, 20

rufescens

Elephantulus, 7

rufiventris

Ateles, 14

rufus

Molossus, 68

russula

Crocidura, 33, 34, 36

saccharus

Platyrrhinus, 64

Vampyrops, 64

saccolaimus

Saccolaimus, 57

Saccolaimus

pluto, 57

saccolaimus, 57

saevus

Hipposideros, 55

safii

Crocidura, 34

Saguinas

labiatus, 14

salatana

Tupaia, 10

saliens

Lepus, 26

saltator

Cephalopachus, 13

Tarsius, 13

salvini

Chiroderma, 59

Sorex, 43, 46, 47

samoensis

Pteropus, 53

sanborni

Micronycteris, 62

sanctaemartae

Sylvilagus, 30

sanctidiegi

Lepus, 28

sanctorum

Pygathrix, 21

Trachypithecus, 21

sangi

Petrodromus, 7

santarosae

Pipistrellus, 80

Sapajus

apaella, 14

margaritae, 14

saturatus

Galeopithecus, 12

Myotis, 76

saussurei

Sorex, 46, 47

saxatilis

Ochotona, 22, 23

Scalops

aeneus, 50

Scalopus

aereus, 50

aquaticus, 50

caryi, 50

howelli, 50

inflatus, 50

machrinoides, 50

pulcher, 50

SCANDENTIA, 9

Scapanus

alpinus, 51

dilatus, 51

latimanus, 51

orarius, 51

schefferi, 51

sericatus, 51

townsendii, 50

truei, 51

schefferi

Scapanus, 51

schisticeps

Lagomys, 21

Ochotona, 21, 23

schmidti

Cercopithecus, 15

sclateri

Sorex, 47

scopaeum

Chiroderma, 59

Scotophilus

hesperus, 80

miradorensis, 71

nucella, 82

semicaudata

Emballonura, 57

semicaudatus

Vespertilio, 57

Semnopithecus

auratus, 21

ebenus, 21

semota

Atalapha, 72

semotus

Lasiurus, 72

senegalensis

Galago, 13

Trichechus, 8

senex

Centurio, 59

senicula

Mormoops, 65

septentrionalis

Myotis, 77

Vespertilio, 77

serasani

Cynopterus, 52

sericatus

Scapanus, 51

serotinus

Eptesicus, 71

serus

Parocnus, 8

sezekorni

Erophylla, 60

shameli

Rhinolophus, 54

shantungensis

Crocidura, 36

shastensis

Sorex, 47

shermani

Blarina, 32

shumaginensis

Sorex, 41

siaca

Tupaia, 11

siamensis

Presbytis, 19, 20

sicula

Crocidura, 33, 36

sierrae

Lepus, 25

siligorensis

Myotis, 76

Simias

concolor, 20

similis

Sorex, 48

Sylvilagus, 31

simiolus

Crocidura, 36
108 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

simplicicanus

Lepus, 29

simus

Myotis, 76

sincepis

Tupaia, 11

SIRENIA, 8

sirhassenensis

Pithecus, 19

Tupaia, 11

siumatis

Megaderma, 56

solifer

Taphozous, 57

solitarius

Rhinolophus, 54

somalicus

Neoromicia, 79

sonomae

Sorex, 47

sordida

Tupaia, 11

sorex

Monodelphis, 6

Sorex

alascensis, 41, 43, 45

alaskanus, 45

albibarbis, 41, 46

albiventer, 42

alnorum, 41

alpinus, 41

alticola, 41

amoenus, 41

araneus, 41, 42

arcticus, 42, 45, 46, 47

bairdi, 42, 45

bendirii, 40, 42

bergensis, 42

blitchi, 42

caecutiens, 44

californicus, 42

carolinensis, 33

cascadensis, 46

caudatus, 46

chiapensis, 49

cinereus, 40, 43, 46

coronatus, 42

crawfordi, 40

cristobalensis, 47

destructioni, 48

dispar, 42

dobsoni, 42

durangae, 42

elassodon, 44

emarginatus, 42

euronotus, 42

evotis, 40

eximius, 43

fimbripes, 43

fisheri, 43

fontinalis, 43

fumeus, 43

glacialis, 43

godmani, 43

haydeni, 43

hercynicus, 41

hollisteri, 46

hoyi, 40, 41, 43, 48

humboldtensis, 48

hydrobadistes, 46

idahoensis, 43

isolatus, 45

juncensis, 42

lagunae, 44

laricorum, 42

leucogenys, 44

longicauda, 44, 45

longicaudus, 45

longirostris, 43

lyelli, 47

macrodon, 44

macropygmaeus, 44

merriami, 44

milleri, 44

montanus, 41

montereyensis, 44

monticola, 41, 42, 43, 44, 45, 

48, 49

monticolus, 44

mutabilis, 47

myops, 48

nanus, 48

navigator, 41, 45

neomexicanus, 45

nevadensis, 45

oaxacae, 47

obscuroides, 48

obscurus, 45, 48, 49

oreopolus, 45

orizabae, 45

ornatus, 42, 44, 45

pachyurus, 45

pacificus, 46, 49

palmeri, 42

palustris, 46

parvidens, 45

permiliensis, 45

personatus, 46

planiceps, 46

preblei, 46

prevostensis, 44

pribilofensis, 46

roboratus, 46

salvini, 43, 46, 47

saussurei, 46, 47

sclateri, 47

shastensis, 47

shumaginensis, 41

similis, 48

sonomae, 47

sphagnicola, 47

stizodon, 47

streatori, 46

suckleyi, 47

tenelliodus, 47

tenellus, 47, 48

thompsoni, 48

trigonirostris, 48

trowbridgii, 44, 48

tundrensis, 48

turneri, 46

umbrosus, 43

vagrans, 41, 42, 45, 47, 48, 

49

vancouverensis, 49

ventralis, 45

veraecrucis, 47

veraepacis, 47, 49

winnemana, 41

yaquinae, 49

SORICIDAE, 32

soricina

Blarina, 38

Glossophaga, 60, 61

soricinus

Cryptotis, 38

SORICOMORPHA, 32

sotikae

Erinaceus, 32

Galago, 13

sowerbyae

Lepus, 26

sowerbyi

Myotis, 76

spadix

Rhinolophus, 55

spasma

Megaderma, 56

speciosa

Tupaia, 9

speciosus

Pteropus,53 

spectrum

Vampyrum, 65

Vampyrus, 65

sphagnicola

Sorex, 47

sphinx

Cynopterus, 51, 52

splendidula

Tupaia, 9, 10

splendidulus

Tupaia, 11

Stenoderma

luciae, 57

tolteca, 60

stephensi

Myotis, 74

stizodon

Sorex, 47

stoliczkanus

Lepus, 26

stramineus

Natalus, 66

streatori

Sorex, 46

stuhlmanni

Cercopithecus, 15

Sturnira

angeli, 64

aratathomasi, 64

bogotensis, 64

lilium, 64

mordax, 64

parvidens, 64

paulsoni, 64

thomasi, 64

Sturnirops

mordax, 64

suahelae

Crocidura, 36

subcinctus

Lepus, 28

subflavus

Perimyotis, 81

Pipistrellus, 81

subrufa

Carollia, 58

subrufum

Hemiderma, 58

subulatus

Vespertilio, 79

subulidens

Pipistrellus, 81

suckleyi

Sorex, 47

sulcata

Emballonura, 57

sultani

Petrodromus, 7

suluensis

Cynomolgus, 16

sumatrana

Presbytis, 19

Suncus

murinus, 49

surda

Tupaia, 10

Surdisorex

polulus, 49

sururae

Crocidura, 36

swinhoei

Lepus, 26

sylvaticus

Lepus, 29

Sylvilagus

ammophilus, 31

aquaticus, 30

arizonae, 27, 28, 29

audubonii, 27, 28, 29, 30

aztecus, 28

bachmani, 28, 30

badistes, 31

baileyi, 28, 30

brasiliensis, 30

cedrophilus, 30

chapmani, 28, 29

cognatus, 30

connectens, 28

cunicularius, 30

exiguus, 30

floridanus, 28, 29, 31

gabbi, 28, 30, 31

goldmani, 27

graysoni, 29, 31

hefneri, 31

hitchensi, 31

holzneri, 29

hondurensis, 31
NUMBER 644

• 109

insonus, 29

littoralis, 30

macrocorpus, 31

mallurus, 31

mansuetus, 31

margaritae, 29

mearnsii, 31

messorius, 31

minor, 27

neomexicanus, 30

obscurus, 31

orizabae, 28, 29, 31

palustris, 31

restrictus, 31

robustus, 29

sanctaemartae, 30

similis, 31

truei, 30

ubericolor, 28

vallicola, 30

warreni, 30

yucatanicus, 29

Sylvisorex

aequatorius, 49

gemmeus, 49

konganensis, 49

megalura, 49

Symphalangus

klossii, 21

syrichta

Carlito, 13

tabascensis

Didelphis, 4

Tadarida

antillularum, 69

brasiliensis, 69, 70

cobourgiana, 68

constanzae, 69

cynocephala, 69

intermedia, 69, 70

loriae, 68

mexicana, 69

Talpa

parvidens, 50

TALPIDAE, 49

talpoides

Blarina, 32

tamensis

Cryptotis, 39

tamrinensis

Crocidura, 33

tana

Tana, 9

Tupaia, 9, 11

Tana

besara, 9

cervicalis, 9

lingae, 9

masae, 9

tana, 9

tuancus, 9

Taphonycteris

capito, 57

Taphozous

cavaticus, 57

melanopogon, 57

philippinensis, 57

pluto, 57

solifer, 57

tarfayaensis

Crocidura, 33, 34, 36

TARSIIDAE, 13

Tarsius

borneanus, 13

dentatus, 13

fraterculus, 13

fuscus, 13

pumilus, 13

saltator, 13

tatei

Marmosa, 6

Thylamys, 6

Tatoua

centralis, 8

Tatu

novemcinctum, 8

texanum, 8

teliotis

Atalapha, 72

Lasiurus, 72

telmalestes

Blarina, 32

tenelliodus

Sorex, 47

tenellus

Sorex, 47, 48

tenuidorsalis

Vespertilio, 79

tenuis

Pipistrellus, 81

tephrura

Tupaia, 11

terrestris

Colobus, 16

tetradactylus

Petrodromus, 7

texanum

Tatu, 8

texensis

Didelphis, 4

texianus

Lepus, 24, 25, 26, 27

thaianus

Myotis, 76

thibetana

Macaca, 19

thomasi

Sturnira, 64

thompsoni

Sorex, 48

Thylamys

tatei, 6

thyone

Vampyressa, 65

Thyroptera

abdita, 66

discifera, 66

major, 66

THYROPTERIDAE, 66

thysanodes

Myotis, 76

tibetanus

Lepus, 26

tiburonensis

Lepus, 23

timidus

Lepus, 27

titthaecheilus

Cynopterus, 51

tiznitensis

Crocidura, 36

Tlacuatzin

canescens, 6

toala

Hipposideros, 55

tolai

Lepus, 24, 25, 26

tolteca

Dermanura, 60

Stenoderma, 60

townsendii

Corynorhinus, 70, 71

Lepus, 25

Scapanus, 50

Trachops

cirrhosus, 64

coffini, 64

Trachypithecus

carbo, 20

crepusculus, 20

cristatus, 21

ebenus, 21

flavicaudus, 21

germaini, 20

obscurus, 20, 21

phayrei, 20

sanctorum, 21

vigilans, 21

TRICHECHIDAE, 8

Trichechus

senegalensis, 8

tricolor

Ateles, 15

Hemiderma, 58

tridactyla

Myrmecophaga, 8

trifoliatus

Rhinolophus, 54, 55

trigonirostris

Sorex, 48

trinitatum

Chiroderma, 59

triomylus

Artibeus, 58

trowbridgii

Lepus, 29

Sorex, 44, 48

truei

Glossophaga, 61

Lepus, 30

Podogymnura, 32

Scapanus, 51

Sylvilagus, 30

tua

Macaca, 17

tuancus

Galeopithecus, 12

Tana, 9

tularensis

Lepus, 27

tumbili

Lasiopyga, 15

tumida

Rhogeessa, 82

tumidiceps

Mormoops, 65

tumidifrons

Chilonatalus, 66

tumidirostris

Natalus, 67

tundrensis

Sorex, 48

tuneti

Rhinolophus, 54

Tupaia

anambae, 9

batamana, 10

belangeri, 10

besara, 9

bunoae, 9

carimatae, 9

castanea, 9

cervicalis, 9

chrysogaster, 9

chrysomalla, 9

cuyonis, 9

discolor, 10

edarata, 10

everetti, 11

ferruginea, 10, 11

fuscior, 11

glis, 9, 10, 11

gracilis, 10

inflata, 10

kohtauensis, 10

lacernata, 10

lingae, 9

longipes, 10

masae, 9

minor, 11

moellendorffi, 9

natunae, 10

nicobarica, 10

olivacea, 10

pemangilis, 10

phaeura, 10

picta, 11

pulonis, 11

raviana, 11

riabus, 11

salatana, 10

siaca, 11

sincepis, 11

sirhassenensis, 11

sordida, 11

speciosa, 9

splendidula, 9, 10

splendidulus, 11
110 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY

Tupaia (continued)

surda, 10

tana, 9, 11

tephrura, 11

TUPAIIDAE, 9

turba

Crocidura, 37

turneri

Sorex, 46

ubericolor

Lepus, 28

Sylvilagus, 28

ugandae

Eptesicus, 79

Neoromicia, 79

uinta

Ochotona, 23

ultima

Pygathrix, 21

umbrosa

Macaca, 17

umbrosus

Macacus, 17

Sorex, 43

Uroderma

convexum, 65

Urogale

cylindrura, 11

Urotrichus

gibbsii, 50

ursinus

Ailurops, 7

vagrans

Sorex, 41, 42, 45, 47, 48, 49

validus

Dendrohyrax, 7

vallicola

Sylvilagus, 30

Vampyressa

elisabethae, 65

minuta, 65

thyone, 65

Vampyrops

aquilus, 63

aurarius, 63

fumosus, 63

latus, 63, 64

saccharus, 64

Vampyrum

spectrum, 65

vampyrus

Pteropus, 53

Vampyrus

nelsoni, 65

spectrum, 65

vancouverensis

Sorex, 49

variegatus

Bradypus, 8

Galeopterus, 11, 12

varronis

Lepus, 27

vellerosus

Ateles, 15

venezuelae

Lophostoma, 61

ventorum

Ochotona, 23

ventralis

Sorex, 45

veraecrucis

Lepus, 30

Sorex, 47

veraepacis

Sorex, 47, 49

Vespertilio

affinis, 76

agilis, 76

bahamensis, 71

ciliolabrum, 77

evotis, 77

exiguous, 77

exilis, 77, 79

fuscus, 71

gryphus, 77

henshawii, 78

keenii, 79

longicrus, 77

lucifugus, 77

macropus, 77

melanorhinus, 78

mundus, 78

nitidus, 78, 79

obscurus, 78

oregonensis, 78, 79

pallidus, 70

ruber, 79

semicaudatus, 57

septentrionalis, 77

subulatus, 79

tenuidorsalis, 79

volans, 79

yumanensis, 79

VESPERTILIONIDAE, 70

Vesperugo

merriami, 81

viaria

Crocidura, 33, 36

vigilans

Presbytis, 21

Trachypithecus, 21

vigilis

Papio, 19

villosa

Glossophaga, 61

villosum

Chiroderma, 59

villosus

Macacus, 17

virginiana

Didelphis, 4

virginianus

Corynorhinus, 71

virgo

Rhinolophus, 55

vitis

Pithecus, 19

vociferus

Pteropus, 53

volans

Myotis, 73, 74, 75, 77, 79

Vespertilio, 79

vulpina

Ochotona, 22

wallawalla

Lepus, 27

warreni

Sylvilagus, 30

washingtoni

Microsorex, 41

washingtonii

Lepus, 25, 27

watasei

Myotis, 74

waterhousii

Macrotus, 62

watsoni

Dermanura, 59

wetmorei

Eptesicus, 71

whitakeri

Crocidura, 34, 35, 37

whitei

Erythrocebus, 16

winnemana

Microsorex, 41

Myotis, 76

Sorex, 41

woodalli

Marmosa, 5

xanti

Lepus, 25

xerophila

Marmosa, 5

yaquinae

Sorex, 49

yebalensis

Crocidura, 33, 34

yucatanensis

Ateles, 15

Didelphis, 4

yucatanicus

Lepus, 29

Nyctinomops, 68

Sylvilagus, 29

yumanensis

Myotis, 75, 76, 77, 78, 79

Vespertilio, 79

zaianensis

Crocidura, 37

zamicrus

Nesophontes, 32

zaphiri

Crocidura, 36

zeledoni

Marmosa, 5

zeteki

Cryptotis, 40

zonalis

Aotus, 14
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visit www.scholarlypress.si.edu or Open SI (http://

opensi.si.edu). 

ABSTRACTS must not exceed 300 words. 

TEXT must be prepared in a recent version of 

Microsoft Word; use a Times font in 12 point for 

regular text; be double spaced; and have 1" margins. 

REQUIRED ELEMENTS are title page, abstract, 

table of contents, main text, and references.

FIGURES should be numbered sequentially (1, 2, 

3, etc.) in the order called out; have components 

lettered consistently (in size, font, and style) and 

described in captions; include a scale bar or scale 

description, if appropriate; include any legends in 

or on the figure rather than in a caption. Figures 

must be original and submitted as individual TIF or 

EPS files.

FIGURE FILES must not be embedded in the main 

text. Resolution for art files must be at least 300 dpi 

for grayscale and color images, and at least 1200 

dpi for line art. Color images should be requested 

only if required. 

TAXONOMIC KEYS in natural history papers 

should use the aligned-couplet form for zoology. If 

cross referencing is required between key and text, 

do not include page references within the key but 

number the keyed-out taxa, using the same numbers with their corresponding heads in the text.

SYNONYMY IN ZOOLOGY must use the short 

form (taxon, author, year:page), with full reference 

at the end of the paper under “References.”

REFERENCES should be in alphabetical order, 

and in chronological order for same-author entries. 

Each reference should be cited at least once in main 

text. Complete bibliographic information must 

be included in all citations. Examples of the most 

common types of citations are provided on SISP’s 

“Author Resources” page at www.scholarlypress

.si.edu.

SUMMARY OF REQUIREMENTS FOR SMITHSONIAN CONTRIBUTIONS SERIES