United States

Department

of Agriculture

Forest Service

Rocky Mountain

Research Station

General Technical Report

RMRS-GTR-88WWW

January 2002

Recovery History of

Greenback Cutthroat

Trout: Population

Characteristics, Hatchery

Involvement, and

Bibliography

Version 1.0

Michael K. Young

Amy L. Harig

Bruce Rosenlund

Chris Kennedy
Young, Michael K.; Harig, Amy L.; Rosenlund, Bruce; Kennedy, Chris. 2002. Recovery history of

greenback cutthroat trout: population characteristics, hatchery involvement, and bibliography,

version 1.0. [Online]. RMRS-GTR-88WWW. Fort Collins, CO: U.S. Department of Agriculture,

Forest Service, Rocky Mountain Research Station. 25 p.

The greenback cutthroat trout (Oncorhynchus clarki stomias) is native to the mountain and foothill

waters of the South Platte and Arkansas river basins in Colorado. This taxon declined rapidly

beginning near the turn of the century and was listed under the Endangered Species Act in 1973. We

assembled all available information from a variety of sources to document recovery efforts from 1957

to 1999. For each greenback cutthroat trout population, we noted (1) the characteristics of recovery

waters and dates of population discovery or introduction; (2) fish abundance estimates; (3) analyses

of genetic purity; (4) salmonid fauna, removal efforts, and reinvasions; and (5) dates, sources, and

numbers of fish used to establish populations. For each hatchery used to assist recovery, we noted

(1) sources and numbers of fish delivered to them; (2) sources, sex ratios, and production of fish used

to create particular year classes; and (3) dates, numbers, and sizes of hatchery fish shipped to

establish wild populations or supplement other hatchery stocks.

Keywords: threatened species, greenback cutthroat trout, Oncorhynchus clarki stomias, recovery

Acknowledgments ________________________________

We thank the following individuals for providing data or for other contributions to this report: Robert

Behnke, Kevin Bestgen, Mindy Clark, Stephanie Comfort, Karl Cordova, Steven Culver, Pat Dwyer,

Kurt Fausch, Pete Gallagher, Paula Guenther-Gloss, Larry Harris, Ken Kehmeier, Doug Krieger, Bill

Krise, Jim Melby, Tom Nesler, Greg Policky, Nick Schmal, Phil Schler, Ed Stege, Harry Vermillion,

Mary Kay Watry, and David Winters.

Abstract _________________________________________

Authors _________________________________________

Michael K. Young is with the Rocky Mountain Research Station in Laramie, Wyoming. Amy L.

Harig is with Trout Unlimited in Boulder, Colorado. Bruce Rosenlund and Chris Kennedy are

with the Colorado Fish and Wildlife Assistance Office of the U.S. Fish and Wildlife Service in

Lakewood, Colorado.
USDA Forest Service Gen. Tech. Rep. RMRS-GTR-88WWW. 2002 1

Introduction ___________________

The greenback cutthroat trout (Oncorhynchus clarki

stomias) is native to the mountain and foothill waters

of the South Platte and Arkansas river basins in

Colorado (Behnke and Zarn 1976). This taxon was

abundant in the late 19th century when large numbers of European immigrants arrived in and along the

Front Range of Colorado. At that time, fish from 2 to

4.5 kg were relatively common historically and were

notable for their extensive migrations to spawn, rear,

and overwinter (Wiltzius 1985). Subsequent mining in

the Arkansas River basin and southern tributaries of

the South Platte River (Ubbelohde and others 1976)

introduced large amounts of sediment and toxic runoff

that reduced or exterminated many greenback cutthroat trout populations, as did agricultural development in river valleys because of water diversions

(Jordan 1891). Furthermore, harvest of greenback

cutthroat trout, often with explosives, was sufficiently

widespread to have eliminated additional populations

(Wiltzius 1985). Although by 1919 greenback cutthroat trout were still found in many tributaries of the

upper Arkansas River (Carhart 1950), there are no

reports on the status of populations in other locations

at that time.

Concomitant with the extensive commercial and recreational harvest of greenback cutthroat trout in the

late 1880s was the appearance of private and government fish hatcheries in Colorado. Initially, greenback

cutthroat trout were frequently cultured, but their use

diminished by the 1900s (Wiltzius 1985), possibly

because few wild fish were still extant. Brook trout

(Salvelinus fontinalis) were the first nonnative salmoRecovery History of Greenback

Cutthroat Trout: Population

Characteristics, Hatchery

Involvement, and Bibliography

Version 1.0

Michael K. Young

Amy L. Harig

Bruce Rosenlund

Chris Kennedy

nids cultured in Colorado in 1872, but other species

soon followed. Because cutthroat trout are often replaced by brook trout and brown trout (Salmo trutta)

and readily hybridize with rainbow trout (O. mykiss)

and nonindigenous subspecies of cutthroat trout

(Behnke 1992; Behnke and Zarn 1976; Wang and

White 1994), introductions of nonnative trout, or the

invasion of stocked populations into new waters, are

believed to have eliminated greenback cutthroat trout

from nearly all of their remaining historical range.

By the 1930s, the subspecies was considered extinct

(Green 1937), but an apparently pure population in a

portion of the Big Thompson River in Rocky Mountain

National Park was found in 1957 (U.S. Bureau of Sport

Fisheries and Wildlife 1957). Though this population

was later thought to be introgressed with nonnative

species, additional populations detected in 1965 and

1970 were deemed pure. Consequently, the greenback

cutthroat trout was listed as endangered in 1973

under the U.S. Endangered Species Act, and downlisted

to threatened in 1978 (USFWS 1998). The original

recovery plan was completed in 1977 (Greenback Cutthroat Trout Recovery Team 1977), and revisions were

adopted in 1983 (USFWS 1983) and 1998 (USFWS

1998).

Although surveys for remaining populations continue

and are occasionally successful, most recovery efforts

have focused on establishing new populations (USFWS

1998). By 1999, introductions had been attempted in

44 waters (Harig and others 2000). Many of these

attempts have been successful, to the extent that the

greenback cutthroat trout may soon be proposed for
2 USDA Forest Service Gen. Tech. Rep. RMRS-GTR-88WWW. 2002

delisting under the Endangered Species Act within all

or part of its historical range (USFWS 1998).

A requirement for delisting is the completion of a longterm plan to guide management, and central to that

plan is a compilation of the conservation and recovery

history of this subspecies. Although some of this information is readily obtainable (USFWS 1998 and references therein), many data relevant to the discovery,

establishment, and monitoring of wild populations

and the establishment, maintenance, and distribution

of hatchery broodstocks are difficult to find. Consequently, our objective was to compile existing data

from 1957 to 1999 on population characteristics of

greenback cutthroat trout in waters used for recovery

and on hatchery participation in recovery. Additional

information on habitat surveys, habitat modification,

and fishing regulations was excluded. The bibliography contains materials cited in this document as well

as available literature pertinent to the biology and

recovery of greenback cutthroat trout.

Methods ______________________

Information was assembled from peer-reviewed

manuscripts, unpublished reports, student theses,

the Aquatic Data Management System (ADAMAS)

database (CDOW 2000), field data sheets, correspondence, and other paper records in agency offices. Where

data were in conflict, original sources (typically field

data sheets or hatchery logs) were considered the most

accurate, though these were not always obtainable.

Otherwise, conflicts were resolved by using the most

frequently reported data. All entries were checked

against the source data at least twice.

For each greenback cutthroat trout population, we

noted (1) the characteristics of recovery waters and

dates of population discovery or introduction; (2) fish

abundance estimates; (3) analyses of genetic purity;

(4) salmonid fauna, removal efforts, and reinvasions;

and (5) dates, sources, and numbers of fish used to

establish populations. For each hatchery, we noted (1)

sources and numbers of fish delivered to them; (2)

sources, sex ratios, and production of fish used to

create particular year classes; and (3) dates, numbers,

and sizes of hatchery fish shipped to establish wild

populations or supplement other hatchery stocks.

Results were presented separately for the Arkansas

and South Platte river basins because these stocks

may be considered separately for delisting.

Results and Discussion _________

Greenback cutthroat trout were found or introduced

in 68 waters as part of recovery activities (table 1).

Only waters used in an attempt to create a selfsustaining population were included – we did not

consider waters used solely to provide a sport fishery.

We also restricted this list to waters that were believed

to contain genetically pure fish during recovery, though

some of these populations now appear to be hybridized. Nearly all waters were qualitatively surveyed

during recovery efforts, and quantitative estimates

were obtained in 53 waters (table 2). In streams, the

mean abundance of greenback cutthroat trout greater

than 100 mm was 24 fish/100 m (SD, 12). In lakes, the

mean catch rate was 12 fish/gill-net-hour (SD, 9).

Samples from 22 waters were subjected to genetic

analyses (table 3). Most evaluations before 1995 relied

on meristic and morphometric analyses, with more

advanced techniques employed thereafter. It seems

likely that the precision and accuracy of these approaches will continue to improve, and that current

tallies of pure and hybridized populations will change.

For example, the most recent analyses (Evans and

Shiozawa 2001) determined that four populations --

previously used to found broodstocks and other wild

populations -- were introgressed to varying degrees

with Yellowstone cutthroat trout (O. c. bouvieri) or

rainbow trout. The proposed federal policy of protecting hybridized populations (USFWS 1996) may assume a greater role as the ability to estimate the

amount and source of introgression increases.

Of the 68 recovery waters, 41 were treated with antimycin or rotenone to remove nonnative salmonids

prior to introductions of greenback cutthroat trout

(table 4). Because of this intensive use of toxicants,

many protocols for their application were developed

(for example, Tiffan 1992). Although most nonnative

salmonid populations were successfully eliminated,

periodic failures of artificial barriers, illegal introductions by anglers, or incomplete removals have permitted invasions in 15 waters (cf. Harig and others 2000).

Transfers of fish from different waters and different

hatcheries have been extensive (table 5). The median

number of fish introduced into streams was 4,862

(range, 900-28,456) and into lakes 4,696 (range 85-

59,500). We caution that there is substantial uncertainty in these estimates because of the consistent

lack of consensus among literature sources. Most transfers involved fish reared or produced by three hatcheries: the Bellvue Fish Research Hatchery (BFRH) in

Bellvue, Colorado; the Bozeman Fish Technology Center (BFTC) in Bozeman, Montana; and the Saratoga

National Fish Hatchery (SNFH) in Saratoga, Wyoming. The Pueblo State Fish Hatchery in Pueblo,

Colorado, also distributed limited numbers of fish

beginning in 1997, but did not develop broodstocks,
USDA Forest Service Gen. Tech. Rep. RMRS-GTR-88WWW. 2002 3

hence we did not include it in our analysis of hatchery

activities. Roaring Creek was historically stocked,

though the population appears to be genetically pure.

Greenback cutthroat trout arrived in hatcheries as

juveniles, adults, fertilized eggs, and milt (table 6).

Attempts to minimize inbreeding depression involved

the use of milt from wild males to fertilize eggs of

hatchery females at the BFTC (Dwyer and Rosenlund

1988) or avoidance of sister-brother spawning at the

BFRH by using 3-year-old females and 2-year-old

males (Schler, personal communication). Strategies

for developing broodstocks varied among hatcheries

(table 7), as did protocols for selecting spawning groups.

Typical spawning practice at the BFRH was to spawn

three females with two males with milt from males

added sequentially to the eggs of all females, though

1:1 matings were conducted when fish were abundant

(Schler, personal communication). One:one ratios of

females to males were typically attempted at the

SNFH. At the BFTC, milt from males was frequently

limited and occasionally as many as six females were

fertilized with the milt from one male (Dwyer, personal communication). In addition, milt from wild

males was generally stored individually but occasionally mixed in a single vial.

Records of the origin and number of fish spawned were

inconsistent for the BFRH and BFTC. Particularly for

the latter, many females were respawned, thus reports of total spawnings exaggerated the number of

females used. Also, shipments of milt to this hatchery

were occasionally unsuccessful, as were the attempts

to fertilize eggs of hatchery females with milt from

wild males, which accounts for the differences in fish

shipped to hatcheries (table 6) and those used to

produce year classes (table 7).

From 1985 to 1996, over 639,000 fish were distributed

as part of the recovery of greenback cutthroat trout

(table 8). Unfortunately, discrepancies between the

number of fish shipped from hatcheries and the number of fish reported as stocked were typical. In some

cases, more fish were shipped than reported stocked,

which was partly attributable to fish being sent to

nonrecovery waters. In other instances, mortality of

fish in transport was reported and ranged from 5 to 40

percent, in which case our estimates of fish stocked

(table 5) relied on reports from biologists responsible

for stocking. Alternatively, if fewer fish were reported

shipped than reported stocked, stocking totals were

adjusted proportionately downward. Hatchery records

were regarded as more reliable because the weight of

fish shipped and the number of fish per unit weight

were usually measured.

The bibliography contains materials that could be

attributed to specific individuals, groups, or agencies;

with few exceptions this material was not peer-reviewed. Much additional information that could not be

cited was made available through personal communications or from agency files. Although we attempted to

obtain all data relevant to the recovery of greenback

cutthroat trout, we undoubtedly have overlooked some

sources. We trust that our omissions will be corrected

by others with access to additional information. Also,

despite repeated checking of data, there most assuredly are errors in the tables that we hope will be

remedied in later revisions.

After undergoing this effort, our primary recommendation is the creation of a formal process for reporting

data and establishment of an easily accessible database to assist with future management of greenback

cutthroat trout, as has been recommended for other

federally listed species (James 1999; Safford 1995;

Tear and others 1995). The often obscure and disparate sources of data (frequently limited to hand-written

notes and in at least one instance, on a napkin)

consulted to prepare this document are a reminder

that current data reporting strategies risk the loss of

much precious information, and that the availability

of such data to all participants in endangered species

management will lead to better-informed decisions.

References ____________________

Anderson, Rick. 1987. [Letter to Doug Krieger]. Summary of greenback cutthroat restoration efforts for 1987 field season.

November 16. 2 leaves. On file at: U.S. Department of

Agriculture, Forest Service, Rocky Mountain Research Station,

Forestry Sciences Laboratory, RWU 4352, Laramie, WY.

Anderson, Rick. 1989. [Letter to Doug Krieger]. Summary of greenback cutthroat field related work for 1988. January 6. 1 leaf. On

file at: U.S. Department of Agriculture, Forest Service, Rocky

Mountain Research Station, Forestry Sciences Laboratory, RWU

4352, Laramie, WY.

Anderson, Rick; Melby, James L. 1989. Greenback cutthroat trout

recovery project. 1989 progress report. Denver: Colorado Division

of Wildlife, Southeast Region. 20 p.

Anderson, Rick; Melby, Jim; Wurm, Don. 1986. Greenback cutthroat trout restoration and monitoring; SE-8-3. June 1, 1995, to

December 31, 1985; job progress report. Denver, CO: Colorado

Division of Wildlife. 2 p.

Anon. [n.d.]. Current status - greenback cutthroat trout populations

on Arapaho-Roosevelt National Forests. Fort Collins, CO:

Arapaho-Roosevelt National Forest. 5 p.

Azevedo, Robert A; Wallis, O. L. 1962. Inter-agency lake surveys

and trout investigations, Rocky Mountain National Park, 1961

including ten year lake stocking schedule. Washington, DC: U.S.

Department of the Interior, National Park Service. 59 p.

Behnke, Robert. 1985. Greenback cutthroat trout from Hunters

Creek, Rocky Mountain National Park. Fort Collins: Colorado

State University, Department of Fishery and Wildlife Biology. 4 p.

Behnke, Robert. 1986. Greenback cutthroat trout from Upper

Hutcheson Lake, Rocky Mountain National Park. Fort Collins:

Colorado State University, Department of Fishery and Wildlife

Biology. 3 p.

Behnke, Robert. 1988. Three samples of cutthroat trout from Rocky

Mountain National Park. Fort Collins: Colorado State

University, Department of Fishery and Wildlife Biology. 3 p.
4 USDA Forest Service Gen. Tech. Rep. RMRS-GTR-88WWW. 2002

Behnke, Robert. 1989. Identification of seven samples of cutthroat

trout from Rocky Mountain National Park. Fort Collins: Colorado

State University, Department of Fishery and Wildlife Biology. 5 p.

Behnke, Robert J. 1969. Rare and endangered species: the greenback trout, Salmo clarki stomias Cope: sources for introductions

and a recent discovery of a probable pure population. Fort Collins:

Colorado State University, Report for the Colorado Cooperative

Fishery Unit. 5 p.

Behnke, Robert J. 1973. The greenback cutthroat trout, Salmo

clarki stomias. Status report prepared for: U.S. Fish and Wildlife

Service, Region 2, Albuquerque, NM. 11 p.

Behnke, Robert J. 1976. Summary of information on the status of

greenback cutthroat trout, Salmo clarki stomias. Prepared for:

U.S. Fish and Wildlife Service, Salt Lake City, UT. 30 p.

Behnke, Robert J. 1977. Evaluation of recent collections of cutthroat

trout in Colorado. Prepared for: Colorado Division of Wildlife,

Denver. 18 p.

Behnke, Robert J. 1979. Monograph of the native trouts of the

Genus Salmo of western North America. Lakewood, CO: U.S.

Department of Agriculture, Forest Service, Region 2. 163 p.

Behnke, Robert J. 1987. Identification of cutthroat trout samples

from Rocky Mountain National Park. Fort Collins: Colorado

State University, Department of Fishery and Wildlife Biology. 7 p.

Behnke, Robert J. 1992. Native trout of Western North America.

Monograph 6. Bethesda, MD: American Fisheries Society. 275 p.

Behnke, Robert J.; Zarn, M. 1976. Biology and management of

threatened and endangered western trout. Gen. Tech. Rep. RM28. Fort Collins, CO: U.S. Forest Service, Rocky Mountain Forest

and Range Experiment Station.

Bennett, Jim. 1983. [Letter to Greenback Cutthroat Trout Recovery

Team]. Some recent, future business. September 29. 1 leaf. On file

at: U.S. Department of Agriculture, Forest Service, Rocky

Mountain Research Station, Forestry Sciences Laboratory, RWU

4352, Laramie, WY.

Bergersen, Eric P. 1988a. [Letter to Bruce D. Rosenlund]. March 15.

1 leaf. On file at: U.S. Department of the Interior, Fish and

Wildlife Service, Colorado Fish and Wildlife Assistance Office,

Lakewood, CO.

Bergersen, Eric P. 1988b. [Letter to Bruce D. Rosenlund]. May 17.

1 leaf. On file at: U.S. Department of the Interior, Fish and

Wildlife Service, Colorado Fish and Wildlife Assistance Office,

Lakewood, CO.

Bulkley, Ross V. 1957. Survey of waters in Rocky Mountain

National Park for re-establishing greenback trout, Salmo clarki

stomias. Estes Park, CO: U.S. Department of the Interior,

National Park Service. 25 p.

Bulkley, Ross V. 1959. Report of 1958 fishery studies by the Bureau

of Sport Fisheries and Wildlife in Rocky Mountain National Park.

Rocky Mountain Sport Fishery Investigations Administrative

Report. 38 p.

Carhart, Arthur Hawthorne. 1950. Fishing in the West. New York:

MacMillan. 144 p.

Chart, Thomas E.; Gillihan, Scott; Krieger, Douglas A. 1986. Greenback cutthroat trout recovery project: 1986 progress report. Fort

Collins: Colorado Division of Wildlife. 17 p.

Chart, Thomas E.; Haneke, Rock; Kehmeier, Kenneth. 1987. Greenback cutthroat trout recovery project: 1987 progress report. Fort

Collins: Colorado Division of Wildlife. 15 p.

Colorado Division of Wildlife (CDOW). 1992. 1992 greenback cutthroat trout report. Fort Collins, CO: Fish Research Hatchery.

Colorado Division of Wildlife (CDOW). 2000. Aquatic data

management system (ADAMAS). Denver: Colorado Division of

Wildlife.

Cope, E. D. 1872. Report on the recent reptiles and fishes of the

survey, collected by Campbell Carrington and C. M. Dawes. In:

Fifth annual report of the United States Geographical Survey of

the Territories (Hayden's Survey), part 4. Washington, DC: U.S.

Government Printing Office: 467-476.

Cope, Oliver B. 1959. Report on 1959 fishery studies by the Bureau

of Sport Fisheries and Wildlife in Rocky Mountain National Park.

Denver, CO: U.S. Department of the Interior, Bureau of Sport

Fisheries and Wildlife. 5 p.

Culver, Steven R. 1983. Greenback cutthroat trout recovery project:

1983 progress report. Fort Collins: Colorado Division of Wildlife.

29 p.

Culver, Steven R.; Bestgen, Kevin R. 1983. Greenback cutthroat

trout recovery project: 1982 progress report. Fort Collins:

Colorado Division of Wildlife. 93 p.

Culver, Steven R.; Bestgen, Kevin R. 1985. Greenback cutthroat

trout recovery project: 1985 progress report. Fort Collins:

Colorado Division of Wildlife. 37 p.

Culver, Steven R.; Lentsch, Leo D. 1982. Greenback cutthroat trout

recovery project: 1981 progress report. Fort Collins: Colorado

Division of Wildlife. 34 p.

Cummings, Timothy R. 1987. Brook trout competition with greenback cutthroat trout in Hidden Valley Creek, Colorado. Fort

Collins: Colorado State University. 52 p. Thesis.

Davis, James A.; Culver, Steven R. 1984. Greenback cutthroat trout

recovery project: 1984 progress report. Fort Collins: Colorado

Division of Wildlife. 40 p.

Desilet, Ron. 1985. Greenback trout recovery program. [Letter to

Eddie Kochman]. August 23. 4 leaves. On file at: U.S. Department

of Agriculture, Forest Service, Rocky Mountain Research

Station, Forestry Sciences Laboratory, RWU 4352, Laramie, WY.

Dieffenbach, William H. 1964. Taxonomy and selected life history of

the cutthroat trout (Salmo clarki Richardson) of the South Platte

drainage, Colorado. Fort Collins: Colorado State University. 49 p.

Thesis.

Dieffenbach, William H. 1966. Taxonomy of the cutthroat trout

(Salmo clarki Richardson) of the South Platte drainage,

Colorado. Copeia. 3: 414-424.

Dwyer, William P. 1977. [Letter to Bruce Rosenlund]. November 10.

2 leaves. On file at: U.S. Department of Agriculture, Forest

Service, Rocky Mountain Research Station, Forestry Sciences

Laboratory, RWU 4352, Laramie, WY.

Dwyer, William P. 1978. Annual greenback cutthroat trout progress

report for 1978. Bozeman, MT: U.S. Fish and Wildlife Service,

Fish Cultural Development Center. 3 p.

Dwyer, William P. 1979. Annual greenback cutthroat trout progress

report for '79. Bozeman, MT: U.S. Fish and Wildlife Service, Fish

Cultural Development Center. 2 p.

Dwyer, William P. 1980. Greenback cutthroat trout annual report

1980. Bozeman, MT: U.S. Fish and Wildlife Service, Fish

Cultural Development Center. 3 p.

Dwyer, William P. 1981. Greenback cutthroat trout annual report

for 1981. Bozeman, MT: U.S. Fish and Wildlife Service, Fish

Cultural Development Center. 8 p.

Dwyer, William P. 1982. Greenback cutthroat trout annual report

- 1982. Bozeman, MT: U.S. Fish and Wildlife Service, Fish

Cultural Development Center. 4 p.

Dwyer, William P. 1983. Greenback cutthroat trout annual report

- 1983. Bozeman, MT: U.S. Fish and Wildlife Service, Fish

Technology Center. 4 p.

Dwyer, William P. 1984. Greenback cutthroat trout annual report

- 1984. Bozeman, MT: U.S. Fish and Wildlife Service, Fish

Technology Center. 3 p.

Dwyer, William P. 1985. Greenback cutthroat trout annual report

for 1985. Bozeman, MT: U.S. Fish and Wildlife Service, Fish

Technology Center. 3 p.

Dwyer, William P. 1986. Greenback cutthroat trout annual report

- 1986. Bozeman, MT: U.S. Fish and Wildlife Service, Fish

Technology Center. 3 p.

Dwyer, William P. 1987. Greenback cutthroat trout annual report

- 1987. Bozeman, MT: U.S. Fish and Wildlife Service, Fish

Technology Center. 3 p.

Dwyer, William P. 1988. Colorado greenback cutthroat trout 1988

annual report. Bozeman, MT: U.S. Fish and Wildlife Service, Fish

Technology Center. 2 p.

Dwyer, William P. 1989. Colorado greenback cutthroat trout 1989

annual report. Bozeman, MT: U.S. Fish and Wildlife Service, Fish

Technology Center. 4 p.

Dwyer, William P. 1990. Colorado greenback cutthroat trout 1990

annual report. Bozeman, MT: U.S. Fish and Wildlife Service, Fish

Technology Center. 3 p.

Dwyer, William P. 1991. Colorado greenback cutthroat trout 1991

annual report. Bozeman, MT: U.S. Fish and Wildlife Service, Fish

Technology Center. 3 p.

Dwyer, William P. 1992. Colorado greenback cutthroat trout 1992

annual report. Bozeman, MT: U.S. Fish and Wildlife Service, Fish

Technology Center. 2 p.
USDA Forest Service Gen. Tech. Rep. RMRS-GTR-88WWW. 2002 5

Dwyer, William P. 1993. Colorado greenback cutthroat trout 1993

annual report. Bozeman, MT: U.S. Fish and Wildlife Service, Fish

Technology Center. 2 p.

Dwyer, William P. 1994. Colorado greenback cutthroat trout 1994

annual report. Bozeman, MT: U.S. Fish and Wildlife Service, Fish

Technology Center. 2 p.

Dwyer, William P. 2001. [Personal communication]. August 15.

Bozeman, MT; U.S. Fish and Wildlife Service, Fish Technology

Center.

Dwyer, William P.; Rosenlund, Bruce D. 1988. Role of fish culture in

the reestablishment of greenback cutthroat trout. American

Fisheries Society Symposium. 4: 75-80.

Dwyer, William P.; Smith, C. E.; Rosenlund, Bruce D. 1994.

Recovery of the greenback cutthroat trout, a success story. In:

Barnhart, R.; Shake, B.; Hamre, R. H., tech. eds. Wild Trout V:

wild trout in the 21st century. Arlington, VA: Trout Unlimited:

174-176.

Edsall, D. A.; Erdahl, D. A. 1990. Comparison of two fry diets fed to

greenback and Bear Lake cutthroat, and golden trout. Bozeman,

MT: U.S. Fish and Wildlife Service, Fish Technology Center.

Evans, R. Paul; Shiozawa, Dennis K. 2001. The genetic status of

greenback cutthroat trout (Oncorhynchus clarki stomias)

populations in Colorado. Final Report. Denver, CO: Prepared for

the Colorado Division of Wildlife. 12 p.

Fausch, Kurt D.; Cummings, Timothy R. 1985. Effects of brook trout

competition on threatened greenback cutthroat trout. Final Rep.

Contract CX-1200-4-A043. Prepared for: National Park Service,

Rocky Mountain Regional Office, Denver, CO. 20 p.

Fischer, George Richard. 1988. A summary of fisheries activities in

Rocky Mountain National Park. Golden, CO: U.S. Fish and

Wildlife Service. 17 p.

Fittante, Brenda L. 1987. [Letter to Forest Supervisor]. 1986 greenback cutthroat trout recovery progress on Pike & San Isabel

National Forests. January 5. 4 leaves. On file at: U.S.

Department of Agriculture, Forest Service, Rocky Mountain

Research Station, Forestry Sciences Laboratory, RWU 4352,

Laramie, WY.

Gallagher, J. Peter; Winters David S.; Kittrell, Steven. 1994.

Basinwide stream habitat inventory. Volume 1. Analysis,

discussion and conclusions. Pueblo, CO: U.S. Forest Service,

Pike-San Isabel National Forests.

Gold, J. 1978. Untitled report. College Station, TX: Texas A&M

University.

Green, W. S. 1937. Colorado trout. Denver: Colorado Museum of

Natural History. Popular Series No. 2.

Greenback Cutthroat Trout Recovery Team. 1977. Greenback cutthroat trout recovery plan. Denver, CO: U.S. Fish and Wildlife

Service. 26 p.

Gritman, James C. 1977. [Letter to Chief, Permit Branch, Wildlife

Permit Office]. November 16. 1 leaf. On file at: U.S. Department

of Agriculture, Forest Service, Rocky Mountain Research

Station, Forestry Sciences Laboratory, RWU 4352, Laramie, WY.

Guse, N.; Wallis, O. L. 1965. Long range fishery management plan,

Rocky Mountain National Park, 1965-1974. Washington, DC:

U.S. Department of the Interior, National Park Service.

Hammer, J. 1992. [Letter to Wyoming State Supervisor, U.S. Fish

and Wildlife Service]. Report of activities conducted under

authority of subpermit 85-3, permit PRT-704930. December 8. 1

leaf. On file at: U.S. Department of Agriculture, Forest Service,

Rocky Mountain Research Station, Forestry Sciences

Laboratory, RWU 4352, Laramie, WY.

Hammer, J. E. 1988. Annual report: fiscal year 1988. Saratoga, WY:

Saratoga National Fish Hatchery. 8 p.

Hammer, J. E. 1989. Annual report: fiscal year 1989. Saratoga, WY:

Saratoga National Fish Hatchery. 21 p.

Hammer, J. E. 1991. Annual report: fiscal year 1990. Saratoga, WY:

Saratoga National Fish Hatchery. 16 p.

Hammer, J. E. 1991. Annual report: fiscal year 1991. Saratoga, WY:

Saratoga National Fish Hatchery. 21 p.

Hammer, J. E. 1993. Annual report: fiscal year 1993. Saratoga, WY:

Saratoga National Fish Hatchery. 17 p.

Harig, Amy L. 2000. Factors influencing success of cutthroat trout

translocations. Fort Collins: Colorado State University. 140 p.

Dissertation.

Harig, Amy L.; Fausch, Kurt D. 1996. Compilation of data on

Colorado waters containing greenback cutthroat trout. Fort

Collins: Colorado State University, Department of Fishery and

Wildlife Biology. 35 p.

Harig, Amy L.; Fausch, Kurt D. 1996. Habitat for greenback cutthroat trout in Cottonwood Creek, San Isabel National Forest.

Fort Collins: Colorado State University, Department of Fishery

and Wildlife Biology, Colorado State University. 26 p.

Harig, Amy L.; Fausch, Kurt D. 1997. Evaluating risks of extinction

from habitat fragmentation for native cutthroat trout in Colorado

and Wyoming. Fort Collins: Colorado State University,

Department of Fishery and Wildlife Biology. 61 p.

Harig, Amy L.; Fausch, Kurt D. 1998. Evaluating risks of extinction

from habitat fragmentation for native cutthroat trout in

Colorado. Fort Collins: Colorado State University, Department of

Fishery and Wildlife Biology. 59 p.

Harig, Amy L.; Fausch, Kurt D. 1999. Minimum habitat

requirements for establishing translocated cutthroat trout

populations. Fort Collins: Colorado State University,

Department of Fishery and Wildlife Biology, Colorado State

University. 90 p.

Harig, Amy L.; Fausch, Kurt D. 2000. Factors influencing success of

cutthroat trout translocations. Fort Collins: Colorado State

University, Department of Fishery and Wildlife Biology. 140 p.

Harig, Amy L.; Fausch, Kurt D.; Young, Michael K. 2000. Factors

influencing success of greenback cutthroat trout translocations.

North American Journal of Fisheries Management. 20: 994-1004.

Hickman, Terry J.; Miller, David L. 1977. A report on the greenback

cutthroat trout inventory program, July to October, 1977. Fort

Collins: Colorado State University, Department of Fisheries and

Wildlife. 23 p.

James, F. C. 1999. Lessons learned from a study of habitat

conservation planning. BioScience. 49: 871-874.

Johnson, J. E. 1976. Status of endangered and threatened fish

species in Colorado. Tech. Note 280. U.S. Department of the

Interior, Bureau of Land Management.

Jordan, D. S. 1891. Report on explorations in Colorado and Utah

during the summer of 1889, with an account of the fishes found in

each river basin examined. U.S. Fish Commission Bulletin. 9:

1-40.

Juday, C. 1906. A study of Twin Lakes, Colorado, with especial

consideration of the food of the trouts. U.S. Bureau of Fisheries

Bulletin. 26: 147-178.

Kanda, Naohisa; Leary, Robb. 1998. [Letter to Paula GuentherGloss]. October 23. 8 leaves. On file at: U.S. Department of

Agriculture, Forest Service, Rocky Mountain Research Station,

Forestry Sciences Laboratory, RWU 4352, Laramie, WY.

Kehmeier, Ken; Haneke, Rock; Garrison, Tyler. 1988. Greenback

cutthroat trout recovery project: 1988 progress report. Fort Collins:

Colorado Division of Wildlife. 20 p.

Kehmeier, Ken; Van Buren, Randy. 1990. Greenback cutthroat

trout recovery project: 1989 and 1990 progress report. Fort

Collins: Colorado Division of Wildlife. 20 p.

Langlois, David; Hickman, Terry; Miller, David. 1978. Greenback

cutthroat trout restoration program. Endangered Wildlife

Investigations Job Progress Report SE-3-1. Fort Collins:

Colorado Division of Wildlife. 42 p.

Langlois, David; Hickman, Terry; Miller, David. 1979. Greenback

cutthroat trout restoration program. Endangered Wildlife

Investigations. Job Progress Report SE-3-2. Fort Collins:

Colorado Division of Wildlife. 90 p.

Leary, Robb. 1986. [Letter to Bruce Rosenlund]. March 26. 10

leaves. On file at: U.S. Department of Agriculture, Forest Service,

Rocky Mountain Research Station, Forestry Sciences

Laboratory, RWU 4352, Laramie, WY.

Leary, Robb. 1987. [Letter to Jim Melby]. September 14. 8 leaves.

On file at: U.S. Department of Agriculture, Forest Service, Rocky

Mountain Research Station, Forestry Sciences Laboratory, RWU

4352, Laramie, WY.

Loeffler, Chuck; Miller, David; Winters, David; Nelson, Patrick.

1982. Greenback cutthroat trout restoration and monitoring;

January 1, 1981, to June 30, 1982. Job Progress Report. SE-8-1.

Denver: Colorado Division of Wildlife. 3 p.
6 USDA Forest Service Gen. Tech. Rep. RMRS-GTR-88WWW. 2002

Loeffler, Chuck; Young, Douglas; Schoenfeld, Mark; Mellott, Ron;

Mellott, Lisa. 1984. Greenback cutthroat trout restoration and

monitoring; July 1, 1983, to June 30, 1984. Job Progress Report

SE-8-2. Denver, CO: Colorado Division of Wildlife. 6 p.

Mandis, Tom. 1989. Greenback cutthroat trout broodfish spawning

report. [Letter to Larry Harris]. August 1. 1 leaf. On file at: U.S.

Department of Agriculture, Forest Service, Rocky Mountain

Research Station, Forestry Sciences Laboratory, RWU 4352,

Laramie, WY.

Mandis, Tom. 1989. Greenback update. [Letter to Larry Harris].

October 16. 1 leaf. On file at: U.S. Department of Agriculture,

Forest Service, Rocky Mountain Research Station, Forestry

Sciences Laboratory, RWU 4352, Laramie, WY.

Mandis, Tom. 1990. Greenback cutthroat status report. [Letter to

Larry Harris]. January 17. 1 leaf. On file at: U.S. Department of

Agriculture, Forest Service, Rocky Mountain Research Station,

Forestry Sciences Laboratory, RWU 4352, Laramie, WY.

Mandis, Tom. 1991. 1991 spawning of greenback natives at the

research hatchery. [Letter to Larry Harris]. June 25. 1 leaf. On

file at: U.S. Department of Agriculture, Forest Service, Rocky

Mountain Research Station, Forestry Sciences Laboratory, RWU

4352, Laramie, WY.

Mandis, Tom; Harris, Larry E. 1991. Greenback cutthroat trout

(Salmo clarki stomias) propagation. Federal Aid Project SE-13.

Fort Collins: Colorado Division of Wildlife, Fish Research

Section. 2 p.

Mandis, Tom; Martinez, Anita; Schler, Phil. 1996. Greenback cutthroat trout (Oncorhynchus clarki stomias) propagation. Federal

Aid Project SE-13. Fort Collins: Colorado Division of Wildlife,

Fish Research Section. 40 p.

Mandis, Tom; Schler, Phil. 1997. Greenback cutthroat trout

(Oncorhynchus clarki stomias) propagation. Federal Aid Project

SE-13. Fort Collins: Colorado Division of Wildlife, Fish Research

Section. 1 p.

McGrath, C. C. 2000. Mechanisms for displacement of greenback

cutthroat trout by brook trout in montane streams in Rocky

Mountain National Park. Boulder, CO: Center for Limnology,

Cooperative Institute for Research in Environmental Sciences.

Melby, James L. 1990. 1989 stream inventory summary. Vol. 1.

Colorado Springs: Colorado Division of Wildlife, Southeast

Region. 102 p.

Melby, James L.; Policky, Greg A. 1996. Greenback cutthroat trout

recovery efforts. Progress Report. Colorado Springs: Colorado

Division of Wildlife, Southeast Region.

Melby, Jim 1987. [Letter to Doug Krieger]. Greenback cutthroat

trout progress report 1987. December 1. 1 leaf. On file at: U.S.

Department of Agriculture, Forest Service, Rocky Mountain

Research Station, Forestry Sciences Laboratory, RWU 4352,

Laramie, WY.

Melby, Jim 1989. [Letter to Doug Krieger]. Greenback cutthroat

work accomplishments 1988. January 10. 1 leaf. On file at: U.S.

Department of Agriculture, Forest Service, Rocky Mountain

Research Station, Forestry Sciences Laboratory, RWU 4352,

Laramie, WY.

Melton, Brenda L. 1986. [Letter to Forest Supervisor]. 1985 greenback cutthroat trout recovery progress on Pike & San Isabel

National Forests. February 13. 6 leaves. On file at: U.S.

Department of Agriculture, Forest Service, Rocky Mountain

Research Station, Forestry Sciences Laboratory, RWU 4352,

Laramie, WY.

Mitchell, Joseph Daniel. 1978. Evaluation and comparison of stream

habitats and populations between greenback cutthroat and

rainbow trout in the Rocky Mountains of Colorado. Boulder:

University of Colorado. 53 p. Thesis.

Mullan, James W. 1969. Fishery management program, Rocky

Mountain National Park, Colorado: 1968 progress report. Vernal,

UT: U.S. Fish and Wildlife Service, Bureau of Sport Fisheries and

Wildlife, Division of Fishery Services. 15 p.

Mullan, James W. 1970. Fishery management program, Rocky

Mountain National Park, Colorado: 1969 annual project report.

Vernal, UT: U.S. Fish and Wildlife Service, Bureau of Sport

Fisheries and Wildlife, Division of Fishery Services. 12 p.

Mullan, James W. 1971. Fishery management program, Rocky

Mountain National Park, Colorado: 1970 annual project report.

Vernal, UT: U.S. Fish and Wildlife Service, Bureau of Sport

Fisheries and Wildlife, Division of Fishery Services. 5 p.

Mullan, James W. 1972. Fishery management program, Rocky

Mountain National Park, Colorado: 1971 annual project report.

Vernal, UT: U.S. Fish and Wildlife Service, Bureau of Sport

Fisheries and Wildlife, Division of Fishery Services. 27 p.

Mullan, James W. 1973. Fishery management program, Rocky

Mountain National Park, Colorado: 1972 annual project report.

Vernal, UT: U.S. Fish and Wildlife Service, Bureau of Sport

Fisheries and Wildlife, Division of Fishery Services. 36 p.

Mullan, James W. 1974. Considerations in perpetuation of greenback cutthroat trout. Fishery management program, Rocky

Mountain National Park, Colorado. 1973 Special Report. Vernal,

UT: U.S. Fish and Wildlife Service, Bureau of Sport Fisheries and

Wildlife, Division of Fishery Services. 26 p.

Mullan, James W. 1976. Fishery management program, Rocky

Mountain National Park, Colorado: 1974-75 annual project

reports. Vernal, UT: U.S. Fish and Wildlife Service. 53 p.

Nelson, W. C. 1972. An unexploited population of greenback trout.

Fort Collins: Colorado Division of Wildlife. 13 p.

Policky, Greg A. 1987. Greenback cutthroat trout status report,

1987. Denver: Colorado Division of Wildlife, Central Region. 3 p.

Policky, Greg A. 1990. Greenback cutthroat trout status report,

1989. Denver: Colorado Division of Wildlife, Central Region. 2 p.

Policky, Greg A. 1993. Greenback cutthroat trout status report,

1990-1992. Denver, CO: Colorado Division of Wildlife, Central

Region. 2 p.

Policky, Greg A.; Melby, James L. 1995. Greenback cutthroat trout

recovery efforts: 1995 progress report. Denver: Colorado Division

of Wildlife, Southeast Region. 13 p.

Policky, Greg A.; Melby, James L.. 1996. Greenback cutthroat trout

recovery efforts: 1996 progress report. Denver: Colorado Division

of Wildlife, Southeast Region. 13 p.

Policky, Greg A.; Melby, James L. 1997. Greenback cutthroat trout

recovery efforts: 1997 progress report. Denver: Colorado Division

of Wildlife, Southeast Region. 13 p.

Policky, Greg A.; Melby, James L.; Dowler, Gary S. 1998. Greenback

cutthroat trout recovery efforts: 1998 progress report. Denver:

Colorado Division of Wildlife, Southeast Region. 14 p.

Policky, Greg A.; Melby, James L.; Dowler, Gary S.; Krieger, Doug.

2000. Greenback cutthroat trout recovery efforts: 2000 progress

report. Denver: Colorado Division of Wildlife, Southeast Region.

27 p.

Proebstel, Don. [n.d.]. Genetic variability of greenback cutthroat

trout (Oncorhynchus clarki stomias): as determined by analysis

of meristic characters and restriction fragment length

polymorphisms of mitochondrial DNA. Unpublished paper on file

at: U.S. Department of Agriculture, Forest Service, Rocky

Mountain Research Station, Forestry Sciences Laboratory, RWU

4352, Laramie, WY. 10 p.

Proebstel, Don S.; Martinez, Anita M.; Ellis, Robert P. 1996.

Taxonomic status of cutthroat trout, Rio Grande suckers, and

Arkansas darters determined through morphometric, meristic,

and mitochondrial DNA analysis. Unpublished paper on file at:

U.S. Department of Agriculture, Forest Service, Rocky Mountain

Research Station, Forestry Sciences Laboratory, RWU 4352,

Laramie, WY. 150 p.

Rosenlund, Bruce D. 1978. Rocky Mountain National Park fisheries

management report. Golden, CO: U.S. Fish and Wildlife Service.

56 p.

Rosenlund, Bruce D. 1979. Rocky Mountain National Park fisheries

management report. Golden, CO: U.S. Fish and Wildlife Service.

79 p.

Rosenlund, Bruce D. 1981. [Letter to Greenback Cutthroat Trout

Recovery Team]. Status and future plans for the greenback

cutthroat trout broodstock. January 16. 6 leaves. On file at: U.S.

Department of Agriculture, Forest Service, Rocky Mountain

Research Station, Forestry Sciences Laboratory, RWU 4352,

Laramie, WY.

Rosenlund, Bruce D. 1984. [Letter to Chuck Loeffler]. August 14. 3

leaves. On file at: U.S. Department of Agriculture, Forest Service,

Rocky Mountain Research Station, Forestry Sciences

Laboratory, RWU 4352, Laramie, WY.

Rosenlund, Bruce D. 1987. [Letter to Greenback Cutthroat Trout

Recovery Team]. August 31. 1 leaf. On file at: U.S. Department of

Agriculture, Forest Service, Rocky Mountain Research Station,

Forestry Sciences Laboratory, RWU 4352, Laramie, WY.
USDA Forest Service Gen. Tech. Rep. RMRS-GTR-88WWW. 2002 7

Rosenlund, Bruce D. 1988. [Letter to Greenback Cutthroat Trout

Recovery Team]. Stocking plan for Como Creek, Cascade Creek,

and Poudre River greenbacks. January 26. 7 leaves. On file at:

U.S. Department of Agriculture, Forest Service, Rocky Mountain

Research Station, Forestry Sciences Laboratory, RWU 4352,

Laramie, WY.

Rosenlund, Bruce D. 1988. [Letter to Greenback Cutthroat Trout

Recovery Team]. August 29. 1 leaf. On file at: U.S. Department of

the Interior, Fish and Wildlife Service, Colorado Fish and

Wildlife Assistance Office, Lakewood, CO.

Rosenlund, Bruce D. 1989. [Letter to the Greenback Cutthroat

Trout Recovery Team]. Collection of eggs for a new Colorado

based greenback broodstock. July 17. 2 leaves. On file at: U.S.

Department of Agriculture, Forest Service, Rocky Mountain

Research Station, Forestry Sciences Laboratory, RWU 4352

Laramie, WY.

Rosenlund, Bruce D. 1993. [Letter to Greenback Cutthroat Trout

Recovery Team]. Greenback Cutthroat Trout Recovery Team

meeting. January 19. 20 leaves. On file at: U.S. Department of

Agriculture, Forest Service, Rocky Mountain Research Station,

Forestry Sciences Laboratory, RWU 4352, Laramie, WY.

Rosenlund, Bruce D. [n.d.]. Genetic study of greenback cutthroat

trout. Golden, CO: U.S. Fish and Wildlife Service. 5 p.

Rosenlund, Bruce D.; Axtell, Craig; Johnson, Terese. 1994.

Fisheries and aquatic management report 1992-1993: Rocky

Mountain National Park. Golden, CO: U.S. Fish and Wildlife

Service. 27 p.

Rosenlund, Bruce D.; Canestorp, K. M.; Sharps, D. E.; Barrett, J.

1994. Fisheries and aquatic management, Rocky Mountain

National Park 1993. Denver, CO: U.S. Fish and Wildlife Service.

Rosenlund, Bruce D.; Dwyer, William P. [n.d.]. Uses of cultured

greenback cutthroat trout in reestablishing stable reproducing

populations of greenback cutthroat trout 1889[sic]-1993. 15 p.

Rosenlund, Bruce D.; Emmons, Steve. 1986. Fisheries management

plan. Colorado Springs, CO: U.S. Army, Fort Carson. 5 p.

Rosenlund, Bruce D.; Kennedy, Chris; Axtell, Craig. 2000. Fisheries

and aquatic management: Rocky Mountain National Park.

Denver, CO: U.S. Fish and Wildlife Service. 182 p.

Rosenlund, Bruce D.; Stevens, David R. 1988. Fisheries and aquatic

management report: Rocky Mountain National Park. Golden,

CO: U.S. Fish and Wildlife Service.

Rosenlund, Bruce D.; Stevens, David R. 1990. Fisheries and aquatic

management report: Rocky Mountain National Park, 1988-1989.

Golden, CO: U.S. Fish and Wildlife Service. 192 p.

Rosenlund, Bruce D.; Stevens, David R. 1991. Fisheries and aquatic

management report: Rocky Mountain National Park. Golden,

CO: U.S. Fish and Wildlife Service. 23 p.

Safford, H. D. 1995. The recovery planning blues. Northwest

Science. 69: 330-335.

Saulters, Todd D. W.; Gallagher, J. Peter; Winters, David S. [n.d.].

Basinwide stream habitat inventory, North Apache Creek: an

analysis of a potential greenback cutthroat trout stream. Pueblo,

CO: U.S. Forest Service, Pike-San Isabel National Forest. 46 p.

Scarnecchia, Dennis L. 1983. Trout and char production in Colorado's

small streams. Fort Collins: Colorado State University. 190 p.

Dissertation.

Scarnecchia, Dennis L.; Bergersen, Eric P. 1986. Production and

habitat of threatened greenback and Colorado River cutthroat

trouts in Rocky Mountain headwater streams. Transactions of

the American Fisheries Society. 115: 382-391.

Scarnecchia, Dennis L.; Bergersen, Eric P. 1987. Trout production

and standing crop in Colorado's small streams, as related to

environmental features. North American Journal Anderson, Rick.

1987. [Letter to Doug Krieger]. Summary of greenback cutthroat

restoration efforts for 1987 field season. November 16. 2 leaves.

On file at: U.S. Department of Agriculture, Forest Service, Rocky

Mountain Research Station, Forestry Sciences Laboratory, RWU

4352, Laramie, WY.

Schler, Phil. 1999. Greenback cutthroat trout (Oncorhynchus clarki

stomias) propagation. Federal Aid Project SE-13. Fort Collins:

Colorado Division of Wildlife, Fish Research Section. 1 p.

Schler, Phil. 2001. [Personal communication]. April 10. Bellvue,

CO; Colorado Division of Wildlife.

Schler, Phil; Mandis, Tom. 1993. Greenback cutthroat trout (Salmo

clarki stomias) propagation. Federal Aid Project SE-13. Fort

Collins: Colorado Division of Wildlife, Fish Research Section. 7 p.

Schler, Phil; Mandis, Tom. 1994. Greenback cutthroat trout (Salmo

clarki stomias) propagation. Federal Aid Project SE-13. Fort

Collins: Colorado Division of Wildlife, Fish Research Section. 4 p.

Shively, R. 1989a. [Letter to Bruce D. Rosenlund]. Summary of

sampling activities conducted in Rocky Mountain National Park

during 1988. January 27. 1 leaf. On file at: U.S. Department of the

Interior, Fish and Wildlife Service, Colorado Fish and Wildlife

Assistance Office, Lakewood, CO.

Shively, R. 1989b. [Letter to Bruce D. Rosenlund]. Population and

biomass estimate for Pear and Ouzel Lakes. January 27. 1 leaf.

On file at: U.S. Department of the Interior, Fish and Wildlife

Service, Colorado Fish and Wildlife Assistance Office, Lakewood, CO.

Staley, J. 1994. Greenback cutthroat trout status report, 1993.

Denver, CO: Division of Wildlife, Central Region. 2 p.

Stevens, D. R. 1970. Greenback cutthroat trout project. Estes Park,

CO: Rocky Mountain National Park.

Stevens, D. R.; Butts, D.; Mullan, J.; Klag, J. 1973. Aquatic resource

survey 1973. Estes Park, CO: Rocky Mountain National Park.

Stevens, D. R.; Mullan, James. 1976. Aquatic resource survey 1975.

ROMO-N-20. Washington, DC: National Park Service. 6 p.

Stevens, David R.; Mullan, James. 1977. Aquatic resource survey

1976. ROMO-N-20. Washington, DC: National Park Service. 3 p.

Stevens, David R.; Rosenlund, Bruce. 1979. Greenback restoration

study 1978. ROMO-N-20. Washington, DC: National Park

Service. 3 p.

Stevens, David R.; Rosenlund, Bruce D. 1986. Greenback cutthroat

trout restoration in Rocky Mountain National Park. In: Larson,

G.; Soukup, M., eds. Proceedings of the conference on science in

the National Parks. Fort Collins: Colorado State University. 104-118.

Stuber, R. J.; Rosenlund, Bruce D.; Bennett, J. R. 1988. Greenback

cutthroat trout recovery program: management overview.

American Fisheries Society Symposium. 4: 71-74.

Tear, T. H.; Scott, J. M.; Hayward, P. H.; Griffith, B. 1995. Recovery

plans and the Endangered Species Act: are criticisms supported

by data? Conservation Biology. 9: 182-195.

Tiffan, K. R. 1992. Oxidation and performance of antimycin in high

gradient streams. Fort Collins: Colorado State University. 78 p.

Thesis.

Ubbelohde, Carl; Benson, Maxine; Smith, Duane A. 1976. A

Colorado history. Boulder, CO: Pruett Publishing. 390 p.

U.S. Bureau of Sport Fisheries and Wildlife. 1957. Report on 1957

studies on the green-back trout in Rocky Mountain National

Park. 12 p.

U.S. Fish and Wildlife Service (USFWS). 1982. Butler-Borgeson

creek census, Hidden Valley Creek, Rocky Mountain National

Park. Lakewood: Colorado Fish and Wildlife Assistance Office. 7 p.

U.S. Fish and Wildlife Service (USFWS). 1983. Greenback cutthroat trout recovery plan. Denver, CO. 45 p.

U.S. Fish and Wildlife Service (USFWS). 1984. Annual report - 84.

Saratoga, WY: National Fish Hatchery. 2 p.

U.S. Fish and Wildlife Service (USFWS). 1985. Annual report - 85.

Saratoga, WY: National Fish Hatchery. 3 p.

U.S. Fish and Wildlife Service (USFWS). 1985. Endangered Species

Technical Bulletin. 10(8): 7.

U.S. Fish and Wildlife Service (USFWS). 1986. Annual report - 86.

Saratoga, WY: National Fish Hatchery. 2 p.

U.S. Fish and Wildlife Service (USFWS). 1996. Proposed policy and

proposed rule on the treatment of intercrosses and intercross

progeny (the issue of “hybridization”). Federal Register. 61:

4710-4713.

U.S. Fish and Wildlife Service (USFWS). 1998. Greenback cutthroat trout recovery plan. Denver, CO. 62 p.

U.S. Fish and Wildlife Service (USFWS). [n.d.]. Broodstock

notebook. Saratoga, WY: Saratoga National Fish Hatchery.

U.S. Fish and Wildlife Service (USFWS). [n.d.]. Lot history -

broodstock. Saratoga, WY: Saratoga National Fish Hatchery.

Wagner, Eric J. 1982. Taxonomic analysis of cutthroat trout from

Rocky Mountain National Park. Fort Collins: Colorado State

University, Department of Fishery and Wildlife Biology. 9 p.
8 USDA Forest Service Gen. Tech. Rep. RMRS-GTR-88WWW. 2002

Wallis, O. L. 1961. An evaluation of the fishery resources of Rocky

Mountain National Park and needs for interpretation, research

and management. Washington, DC: U.S. Department of the

Interior, National Park Service. 50 p.

Wallis, O. L. 1967. [Letter to Assistant Director, Operations].

Operations concerning circumstances surrounding removal of

greenback cutthroat trout from Forest Canyon in 1965. May 24.

1 leaf. On file at: U.S. Department of the Interior, Fish and

Wildlife Service, Colorado Fish and Wildlife Assistance Office,

Lakewood, CO.

Wallis, O. L.; Azevedo, Robert A. 1961. Inter-agency lake surveys

and trout investigations, Rocky Mountain National Park.

Washington, DC: U.S. Department of the Interior, National Park

Service. 17 p.

Wang, L. 1989. Behavior and microhabitat competition of brown

trout and greenback cutthroat trout in an artificial stream.

Bozeman: Montana State University. Thesis.

Wang, L.; White, R. J. 1994. Competition between wild brown trout

and hatchery greenback cutthroat trout of largely wild

parentage. North American Journal of Fisheries Management.

14: 475-487.

Washington, Ted. 1981. Greenback cutthroat trout restoration

program. Endangered wildlife investigations. Job Progress

Report SE-3-1 to SE-3-3. Fort Collins: Colorado Division of

Wildlife. 26 p.

Wernsman, Gary R. 1973. Systematics of native Colorado cutthroat

trout. Fort Collins: Colorado State University. Thesis. 57 p.

Wiltzius, W. J. 1985. Fish culture and stocking in Colorado, 1872 -

1978. Division Report 12. Denver: Colorado Division of Wildlife.

Winters, David S.; Bennett, Eddie N.; Gallagher, J. Peter. 1995.

Basinwide stream habitat inventory: greenback cutthroat trout

streams of the Pike and San Isabel National Forests in Colorado.

Pueblo: U.S. Forest Service, Pike-San Isabel National Forest. 59

p.

Woodward, D. F.; Farig, A. M.; Little, E. E.; Steadman, B.; Yancik,

R. 1991. Sensitivity of greenback cutthroat trout to acidic pH and

elevated aluminum. Transactions of the American Fisheries

Society. 120: 34-42.

Young, M. K. 1998. Population characteristics of greenback cutthroat trout, Pennock Creek drainage, Arapaho-Roosevelt

National Forest. Laramie, WY: U.S. Department of Agriculture,

Rocky Mountain Research Station. 12 p.

Young, M. K.; Harig, A. L. 2001. A critique of the recovery of

greenback cutthroat trout. Conservation Biology. 15: 1575-1584.
USDA Forest Service Gen. Tech. Rep. RMRS-GTR-88WWW. 2002 9

Tables
10 USDA Forest Service Gen. Tech. Rep. RMRS-GTR-88WWW. 2002

Table 1 -- Waters used for recovery of greenback cutthroat trout from 1957 to 1999. We identified the major river basin, name, drainage, and Colorado Division of Wildlife (CDOW) water code of

each recovery water. Additional information on each water can be obtained by searching for this water code in the ADAMAS database (CDOW 2000). Elevations (m) were obtained from U.S.

Geological Survey 1:24,000 topographic maps or the ADAMAS database. For streams, elevations are the downstream limits of allopatric populations of greenback cutthroat trout. If populations

were largely sympatric with nonnative species or consisted of hybridized fish, the downstream-most location where regularly reported is given. Lake sizes (ha) were obtained from the ADAMAS

database. Short reaches of lake inlets or outlets that also contained greenback cutthroat trout were not included. The lengths (km) of occupied portions of streams were obtained from reports.

Single values for streams indicate a field measurement of fish distribution or consensus on a single value, whereas ranges reflect different estimates given in the absence of field measurements.

The year that fish were discovered or introduced into recovery waters is also noted. If establishment of greenback cutthroat trout was attempted more than once in a water (e.g., following a

chemical treatment to remove all fish), additional dates are included. An asterisk (*) indicates that recovery efforts have been abandoned or greenback cutthroat trout have been extirpated in a

water because of invasion by nonnative salmonids, unsuitable habitat, or a lack of reproduction by greenback cutthroat trout. Missing data are denoted by a dashed line.

Water Drainage

CDOW

Code

Elevation

(m)

Area

(ha)

Length

(km) Found Founded

Arkansas Basin

North Apache Creek Huerfano River 31616 2,424 3.9 1996

South Apache Creek Huerfano River 31618 2,195 4.2 1977

Lake Fork Arkansas River Arkansas River 33821 3,069 6.2 1987,99

Boehmer Reservoir Beaver Creek 82076 3,438 9.0 1985

Cascade Creek (1) Huerfano River 29315 3,109 1.6-1.8 1976

Cascade Creek (2) Rock Creek 29320 3,341 0.9 1991

Cottonwood Creek* Grape Creek 32742 2,847 5.3 1983

Duck Pond Turkey Creek 83078 1,896 1.0 1995

Elk Creek Rock Creek 29712 3,359 1.2-2.5 1991

Greenhorn Creek St. Charles River 29884 3,109 3.4 1989

South Prong Hayden Creek Arkansas River 32487 --- --- 1996

South Fork Huerfano River* Huerfano River 30154 2,926 --- 1963

Lytle Pond Turkey Creek 82949 1,905 0.8 1981

Mason Reservoir Beaver Creek 79966 3,325 45.4 1996

McAlpine Lake* Deer Creek 82672 2,868 0.3 1980

Native Lake Rock Creek 80919 3,417 2.0 1991

Newlin Creek Hardscrabble Creek 30514 2,020 6.9 1997

Rainbow Lake Rock Creek 81315 3,654 3.3 1991

Rock Creek Arkansas River 30659 2,975 5.8-9.0 1991

East Branch Sayers Gulch South Fork Lake Creek 31890 3,218 2.4 1996

Severy Creek Cascade Creek (3) 31312 2,682 1.2-2.0 1997

Swamp Lakes Rock Creek 81276 3,310 0.7 1991

Timberline Lake Lake Fork Arkansas River 81341 3,309 11.0 1987, 99

Virginia Lake Lake Fork Arkansas River 81416 3,481 1.0 1987

South Platte Basin

Albion Creek* Boulder Creek 10001 3,121 2.4 1955

Bard Creek Clear Creek 10051 2,780 6.1-8.0 1982

Bear Lake Big Thompson River 53811 2,888 3.9 1975

Black Hollow Creek Cache la Poudre River 12916 2,323 1.6 1963 1969,80

Bruno Gulch* South Platte River 13920 2,975 3.0-9.0 1986

Caddis Lake Big Thompson River 57340 3,290 0.2 1959

Como Creek Boulder Creek 13184 2,658 3.5 1969

Cony Creek St. Vrain Creek 10621 2,865 4.9 1989

Cornelius Creek* Cache la Poudre River 12827 2,390 6.6 1983

Crystal Lake Big Thompson River 57338 3,508 10.3 1984

Dream Lake Big Thompson River 54685 3,081 1.4 1997

Fern Creek Big Thompson River 11019 2,713 2.4 1982

Fern Lake Big Thompson River 57326 2,905 2.9 1982

George Creek Cache la Poudre River 11003 2,500 10 1983

Hidden Valley Creek Big Thompson River 12954 2,727 2.4 1.6 1973

Hourglass Creek* Cache la Poudre River 13627 2,865 2.8 1966

Hunters Creek St. Vrain Creek 11205 2,865 1.8 1985

Husted Lake Big Thompson River 57415 3,380 4.4 1986,91

Lower Hutcheson Lake St. Vrain Creek 60385 3,308 2.7 1989

Middle Hutcheson Lake St. Vrain Creek 60397 3,411 0.6 1987

Upper Hutcheson Lake St. Vrain Creek 60400 3,509 1.8 1986

Lake Louise Big Thompson River 59801 3,362 2.5 1989

Lawn Lake Big Thompson River 57403 3,349 17.3 1984

Loomis Lake Big Thompson River 57352 3,118 1.2 1991

Lost Lake Big Thompson River 57439 3,266 2.9 1987

May Creek Cache la Poudre River 12978 2,926 4.1 1980

Odessa Lake Big Thompson River 55827 3,054 3.3 1984

(con.)
USDA Forest Service Gen. Tech. Rep. RMRS-GTR-88WWW. 2002 11

Table 1 (Con.)

Ouzel Creek St. Vrain Creek 14085 2,865 4.4 1981

Ouzel Lake St. Vrain Creek 59572 3,048 2 1981

Pear Lake St. Vrain Creek 59584 3,225 6.4 1989

Pennock Creek Cache la Poudre River 11736 2,609 3.4 1986

Little South Fork Poudre River Cache la Poudre River 11940 2,780 1.6 1965

Roaring Creek Cache la Poudre River 12081 2,609 13.2 1963

Roaring River Big Thompson River 13069 2,774 6.9 1984

Sandbeach Lake St. Vrain Creek 59609 3,134 4 1989

East Fork Sheep Creek Cache la Poudre River 13730 2,880 5.4 1982

West Fork Sheep Creek Cache la Poudre River 13742 2,880 7.5 1982

Spruce Lake Big Thompson River 57364 2,947 1.6 1991

Big Thompson River Big Thompson River 12372 2,573 12.5 1957

North Fork Big Thompson River Big Thompson River 13300 3,011 4.7 1970

West Creek Big Thompson River 12524 2,499 5 1979

Williams Gulch Cache la Poudre River 14263 2,804 4.7 1996

Zimmerman Lake Cache la Poudre River 57059 3,199 4.3 1996

Zinn Ranch Ponds* Plum Creek 60943 2,505 0.2-0.7 _ _ 1985

Water Drainage

CDOW

Code

Elevation

(m)

Area

(ha)

Length

(km) Found Founded
12 USDA Forest Service Gen. Tech. Rep. RMRS-GTR-88WWW. 2002

Table 2 -- Abundance estimates for greenback cutthroat trout in recovery waters from 1955 to 1999. Only surveys detecting greenback cutthroat trout are included. We defined qualitative surveys

as visual, angling, or one-pass electrofishing counts used to estimate relative abundance, or as samples taken for other purposes (e.g., genetic analyses, but not including creel surveys).

Quantitative surveys comprised two- or three-pass depletion or mark-recapture counts in streams and catches per gill-net-hour in lakes. Numbers in parentheses denote the number of quantitative

estimates that were reported to exist in additional reaches or years but were unavailable. Sampling intensity is the length (m) of stream sampled or number of nets used in a lake per year. Mean

abundance is the estimated number of greenback cutthroat trout per 100 m in streams or per gill-net-hour in lakes for fish greater than 100 mm total length. The few mark-recapture estimates for

lakes (Bergersen 1988a,b; Shively 1989b) are not included. Means were obtained by taking an average for all reaches or nets for each year, then calculating a mean for all years. (Standard

deviations, SD, are given.) Missing data are denoted by a dashed line.

Water Qualitative Quantitative

Sampling

intensity

(m) Mean SD

Arkansas Basin

South Apache Creek 1977,80-84,92 1989,94,95 191 60 34

Lake Fork Arkansas River 1987,89,97 1995(4) 845 16

Boehmer Reservoir 1987,94-97,99 (3)

Cascade Creek (1) 1976,78,80,81,96,99 1994(8) 78 19

Cascade Creek (2) 1995(1) 165 10

Cottonwood Creek 1984,89,95,96,99

Duck Pond 1987

Dutch Creek 1978

Elk Creek 1995(1) 130 20

Greenhorn Creek 1997 1992,94,95(3) 274 33 16

South Prong Hayden Creek 1996,99

South Fork Huerfano River 1963,76,78

Lytle Pond 1982-88

McAlpine Lake 1981-85

Native Lake 1991,92 1991-93,95 1 5 5

Newlin Creek 1998

Rainbow Lake 1991-93,95 1 5 1

Rock Creek 1992,99 1993,95(3) 330-350 15 3

East Branch Sayres Gulch 1998 --- 6

Severy Creek 1997 (1)

Swamp Lakes 1993,94 2-3 7 9

Timberline Lake 1989,95 1-2 2 2

Virginia Lake 1989,95 1 1 1

South Platte Basin

Albion Creek 1955,57,63,64,69

Bard Creek 1983 1985,94,99(3) 100-275 33 17

Bear Lake 1977-82,84-89

Black Hollow Creek 1963,64,66,67,71,72,78,79,82,83,96 1985-89,99(1) 91-261 21 9

Bruno Gulch 1991 (1)

Caddis Lake 1972 1979,88,99 1 12 5

Como Creek 1969,70,73,75,77-80,82-89 1982,85,91,95,99 201-425 30 4

Cony Creek 1996,98

Cornelius Creek 1984 1985-89,99 91-675 11 5

Crystal Lake 1991-93 1986,88,91-93,98 1-2 1 2

Dream Lake 1998,99 2 9 5

Fern Creek 1985,96 1988,91 45-91 33 11

Fern Lake 1993,98 1984,85,87,90-93 1 7 6

George Creek 1984 1985-89,99 91-750 30 11

Hidden Valley Creek 1974-86,88-90,95,97-99

Hourglass Creek 1969,81,82 1984 100 7

Hunters Creek 1985-89,92,93,97 1988,99 183-450 32 6

Husted Lake 1998 1993,98 1-2 21 2

Lower Hutcheson Lake 1989,91,97 1989,91,98 1 7 5

Middle Hutcheson Lake 1987 1981,86,87,98 1-2 3 6

(con.)
USDA Forest Service Gen. Tech. Rep. RMRS-GTR-88WWW. 2002 13

Table 2 (Con.)

Water Qualitative Quantitative

Sampling

intensity

(m) Mean SD

Upper Hutcheson Lake 1986,96,97 1963,81,86,90,98 1 18 18

Lake Louise 1993,98 1 16 10

Lawn Lake 1986,89,91-93 1986-89,91,92,96 1-2 10 6

Loomis Lake 1998 1993,95 1 16 4

Lost Lake 1993 1989,90,93,98 1 14 16

May Creek 1982,83,96 1986-89,99(1) 91-450 18 14

Odessa Lake 1986,91-93,98 1987,91-93,98 1-2 5 3

Ouzel Creek 1982-89,91,93,97-99 1998 110 39

Ouzel Lake 1985-89,93,98 1983-89,91,93,98 1-2 20 21

Pear Lake 1989,90 1989,98(1) 1 27 27

Pennock Creek 1987,88 1989,98 88-450 11 11

Little South Fork Poudre River 1965,70,75,78-81,84-86,89,90,92,93 1983,86-89,98,99 91-560 13 9

Roaring Creek 1963,64,77,81,97 1999(1) 825 47

Roaring River 1986,91 1988,89 99-183 23 10

Sandbeach Lake 1990,91,93,98 1989-91,93,98 1 34 45

East Fork Sheep Creek 1983,98 1986-89,99(2) 91-475 30 9

West Fork Sheep Creek 1983,98 1986-89,99(2) 91-575 34 11

Spruce Lake 1993,95,98 1992,93,95 1 19 21

Big Thompson River 1957-59,65,67-69,75,78,80,81,85,99 1980(2) 671 29

North Fork Big Thompson River 1973,78,83,84,89,93,97 1986 --- 16

West Creek 1980,83,85-89,98 1999 91 12

Williams Gulch 1982,83,96,97 1985-89,99(1) 100-475 26 12

Zimmerman Lake 1996,97

Zinn Ranch Ponds 1986,87,89 _ _ _ _
14 USDA Forest Service Gen. Tech. Rep. RMRS-GTR-88WWW. 2002

Table 3 -- Genetic assessments of greenback cutthroat trout. Under results, h denotes that the sample was believed to

contain hybridized fish and p denotes the sample was deemed genetically pure. Method refers to the technique used to

ascertain genetic status: m, meristic-morphometric assessment; e, electrophoretic analysis of allozymes; and d, examination

of mitochondrial or nuclear DNA. For each analysis, the year or years from which samples were drawn and the sample size

(N) are noted. Within a particular water, identical samples evaluated on more than one occasion are marked with an asterisk

(*). Missing data are denoted by a dashed line.

Water Results Method Year N

Arkansas Basin

South Apache Creek h m 1977 26

p e 1987 30

h m 1996 10*

p d 1996 10*

p e 1999 16

h d 2001 32

Boehmer Reservoir p m,d 1996 30

h e 1999 30

h d 2000 30

h d 2000 28

h d 2001 27

Cascade Creek (1) p m 1976 15*

p m 1976 15*

p m,d 1996 12

h d 2001 20

Cottonwood Creek h e 1999 7

South Prong Hayden Creek h m 1998 10*

p d 1998 10*

p e 1999 12

South Fork Huerfano River h m 1963 15

p m 1976 15

h m 1978 2

Severy Creek p m,d 1999 12

p d 2001 2

South Platte Basin

Albion Creek p m 1957-64 39

Black Hollow Creek p m 1963 40*

h m 1963 40*

h m 1964-72 ---

p m 1978 8

h d 1996 5

Caddis Lake h m 1972 6

p d 1999 15

(con.)
USDA Forest Service Gen. Tech. Rep. RMRS-GTR-88WWW. 2002 15

Table 3 (Con.)

Como Creek p m 1969 18

p m 1978 10

p d 2001 30

Hidden Valley Creek p e 1985 28

Hunters Creek p m 1985 7

p d 2001 25

Middle Hutcheson Lake p m 1986 3

Upper Hutcheson Lake p m 1987 27

h d 2001 25

May Creek h d 1996 10

Ouzel Creek p e 1985 25

Little South Fork Poudre River p m 1965-70 18

p d 2001 32

Roaring Creek h m 1977 8

p e 1998 25

p d 2001 32

Sheep Creek h d 1996 15

Big Thompson River h m 1959-67 40

p m 1968 ---

p m 1978 12

h m 1982 43

h d 1999 15

Williams Gulch h d 1996 5

Water Results Method Year N
16 USDA Forest Service Gen. Tech. Rep. RMRS-GTR-88WWW. 2002

(con.)

Table 4 -- Salmonid fauna of recovery waters for greenback cutthroat trout. For species present at the time recovery efforts

began in a particular water, abbreviations are: b, historically barren; gbct, greenback cutthroat trout; bk, brook trout; bn,

brown trout; ct, hybridized or nonnative cutthroat trout; lk, lake trout; and rb, rainbow trout. We also note when a water was

chemically treated to remove nonnative species, and if nonnative species reappeared, the year they were detected.

Systems that were chemically treated more than once have multiple entries. For the form of barrier inhibiting upstream

migration by nonnative fish, abbreviations are: a, artificial; g, geomorphic; and n, none. Artificial barriers consist of dams,

water diversions, weirs, constructed ponds, or toxic mine outflows, whereas geomorphic barriers include waterfalls,

cascades, dry stream channels, or naturally landlocked waters.

Water

Species

present

Treatment

year

Nonnative

species

detected Barrier

Arkansas Basin

North Apache Creek b g

South Apache Creek gbct g

Lake Fork Arkansas River bk,bn,ct 1986 1987 a

bk,bn 1998

Boehmer Reservoir ct 1984 a

Cascade Creek (1) gbct g

Cascade Creek (2) b g

Cottonwood Creek b g

Duck Pond bk 1984 a

Elk Creek bk 1990 1995 a

Greenhorn Creek b g

South Prong Hayden Creek gbct g

South Fork Huerfano River bk n

Lytle Pond b 1982(bk) a

McAlpine Lake b a

Native Lake bk 1990 1995 a

Newlin Creek b a

Rainbow Lake bk 1990 g

Rock Creek bk 1990 1995 a

East Branch Sayres Gulch b a

Severy Creek gbct g

Swamp Lakes bk 1990 g

Timberline Lake bk,ct,lk 1986 1987 a

bk 1998

Virginia Lake bk 1986 g

South Platte Basin

Albion Creek bk n

Bard Creek b g

Bear Lake bk 1975 g
USDA Forest Service Gen. Tech. Rep. RMRS-GTR-88WWW. 2002 17

Table 4 (Con.)

Black Hollow Creek bk 1967 1973 a

bk 1979

Bruno Gulch bk 1985 1985 a

Caddis Lake ct 1958 g

Como Creek gbct g

Cony Creek ct,rb 1987,88 g

Cornelius Creek bk,bn 1981,82 1985 a

Crystal Lake ct 1983 g

Dream Lake ct 1996 g

Fern Creek bk,ct 1982 g

Fern Lake bk,ct 1982 g

George Creek bk,bn 1981,82 1985(bk) a,g

Hidden Valley Creek bk 1973 1976 g

Hourglass Creek b 1977(bk) g

Hunters Creek gbct g

Husted Lake bk,ct 1986 1989 g

bk 1990

Lower Hutcheson Lake ct,rb 1986 g

Middle Hutcheson Lake gbct g

Upper Hutcheson Lake gbct g

Lake Louise b g

Lawn Lake bk 1983 g

Loomis Lake bk 1990 g

Lost Lake bk,ct 1986 g

May Creek bk 1979 g

Odessa Lake b g

Ouzel Creek bk 1980 1985 g

Ouzel Lake bk 1980 1985 g

Pear Lake ct,rb 1988 g

Pennock Creek bk,bn 1979,85 1989 a

Little South Fork Poudre River gbct g

Roaring Creek gbct g

Roaring River bk 1983 g

Sandbeach Lake rb 1988 g

East Fork Sheep Creek rb 1980 g

West Fork Sheep Creek rb 1980 g

Spruce Lake rb 1990 g

Big Thompson River b g

North Fork Big Thompson River bk 1986 1993 g

West Creek bk 1978 g

Williams Gulch b g

Zimmerman Lake b g

Zinn Ranch Ponds bk 1984 _ n

Water

Species

present

Treatment

year

Nonnative

species

detected Barrier
18 USDA Forest Service Gen. Tech. Rep. RMRS-GTR-88WWW. 2002

Table 5 -- Destination, date, source, number, and mean length (mm) of greenback cutthroat trout stocked for purposes of recovery. Only waters receiving fish from elsewhere are included.

Dates are given to the nearest day, when known. Under source, year taken from a contributing water (and whether it was held or reared at another location) or year classes from each

hatchery are noted (see tables 7 and 8 for more information on hatchery year classes). Stocking in lake inlets or outlets is included with lake totals. Abbreviations are: BFRH, Bellvue Fish

Research Hatchery; BFTC, Bozeman Fish Technology Center; CNRL, Columbia National Research Lab; PSFH, Pueblo State Fish Hatchery; and SNFH, Saratoga National Fish Hatchery.

Also noted are cases where nonnative or unknown sources of cutthroat trout were stocked. In some instances, fish lengths were not specified in the literature, but were given as fry,

juveniles, adults, or mixtures. Missing data are denoted by a dashed line.

Water Date Source Number

Length

(mm)

Arkansas Basin

North Apache Creek 6/27/1996 1995 Boehmer Reservoir (via BFRH) 360 111

9/24/1997 1997 Boehmer Reservoir (via PSFH) 600 75

Lake Fork Arkansas River 9/9/1987 1987 SNFH 1,500 50

9/12/1988 1988 SNFH 2,000 25

8/30/1989 1988-1989 SNFH 2,000 121

8/28/1990 1990 SNFH 18,000 33

7/23/1996 1995 Boehmer Reservoir (via BFRH) 3,548 116

9/18/1997 1997 Boehmer Reservoir (via PSFH) 1,000 71

Boehmer Reservoir 9/19/1985 1984 McAlpine Lake (via SNFH) 1,670 93

9/19/1985 1984 Lytle Pond (via SNFH) 125 124

10/3/1985 Lytle Pond 100 267

9/9/1987 1987 SNFH 3,500 50

9/12/1988 1988 SNFH 2,500 25

8/30/1989 1989 SNFH 15,320 38

Cottonwood Creek 8/1983 Cascade Creek 32 126

7/25/1985 1984 Lytle Pond (via SNFH) 225 120

7/25/1985 1984 McAlpine Lake (via SNFH) 456 97

9/12/1988 1988 SNFH 3,000 25

8/30/1989 1989 SNFH 1,149 38

Duck Pond 7/25/1985 Lytle Pond 77 240

6/3/1987 Lytle Pond 7 363

1988 Lytle Pond 143 ---

5/19/1993 1992 South Apache Creek & 1992 SNFH (via BFRH) --- 150

7/18/1994 1993 Boehmer Reservoir (via BFRH) 1,523 147

Elk Creek 6/26/1991 1989 SNFH 300 235

9/17/1991 1991 SNFH 750 49

9/22/1992 1992 SNFH 750 38

7/23/1996 1995 Boehmer Reservoir (via BFRH) 728 116

Greenhorn Creek 8/30/1989 1989 SNFH 153 38

9/24/1997 1997 Boehmer Reservoir (via PSFH) 1,000 75

South Fork Huerfano Creek 9/1/1976 Pikes Peak cutthroat trout 4,500 30

Lytle Pond 9/22/1981 Cascade Creek 40 mixture

7/1983 1983 Lytle Pond (via unknown hatchery) 92 fry

6/9/1984 1984 Lytle Pond (via CNRL) 1,100 fry

7/25/1985 1984 McAlpine Lake (via SNFH) 50 97

5/19/1993 1992 South Apache Creek & 1992 SNFH (via BFRH) 2,350 150

McAlpine Lake 1980 Cascade Creek 25 mixture

9/19/1985 1884 Lytle Pond (via SNFH) 30 124

9/19/1985 1984 McAlpine Lake (via SNFH) 30 97

Native Lake 6/26/1991 1989 SNFH 400 235

9/17/1991 1991 SNFH 1,140 49

9/22/1992 1992 SNFH 1,400 38

7/23/1996 1995 Boehmer Reservoir (via BFRH) 728 116

(con.)
USDA Forest Service Gen. Tech. Rep. RMRS-GTR-88WWW. 2002 19

Table 5 (Con.)

7/23/1996 1996a BFRH 700 41

Newlin Creek 9/24/1997 1997 Boehmer Reservoir (via PSFH) 900 75

Rainbow Lake 6/26/1991 1989 SNFH 500 235

9/17/1991 1991 SNFH 1,660 49

9/22/1992 1992 SNFH 1,600 38

7/23/1996 1995 Boehmer Reservoir (via BFRH) 936 116

Rock Creek 6/26/1991 1989 SNFH 1,044 235

9/17/1991 1991 SNFH 8,600 49

9/22/1992 1992 SNFH 4,900 38

7/23/1996 1995 Boehmer Reservoir (via BFRH) 1,612 116

East Branch Sayres Gulch 7/23/1996 1995 Boehmer Reservoir (via BFRH) 728 116

7/23/1996 1996a BFRH 700 41

9/18/1997 1997 Boehmer Reservoir (via PSFH) 1,500 71

Swamp Lakes, Upper and Lower 6/26/1991 1989 SNFH 200 235

9/17/1991 1991 SNFH 1,600 49

9/22/1992 1992 SNFH 900 38

Timberline Lake 9/9/1987 1987 SNFH 7,000 50

9/12/1988 1988 SNFH 12,500 25

8/30/1989 1989 SNFH 20,000 38

8/28/1990 1990 SNFH 20,000 33

Virginia Lake 9/9/1987 1987 SNFH 2,500 50

9/12/1988 1988 SNFH 1,000 25

8/30/1989 1989 SNFH 1,000 38

8/28/1990 1990 SNFH 250 33

South Platte Basin

Bard Creek 10/5/1982 1982 BFTC 6,900 43

11/16/1982 1982 BFTC 980 ---

9/28/1983 1983 BFTC 2,056 46

7/11/1984 1983 BFTC 400 140

9/11/1984 1984 BFTC 1,600 36

9/25/1985 1985 BFTC 5,000 36

9/16/1987 1987 BFTC 5,000 34

9/23/1992 1992 BFTC 5,000 67

6/21/1993 1992 BFRH 1,350 127

Bear Lake 11/4/1975 Como Creek 51 adults

11/4/1975 Como Creek 14 fry

6/26/1981 1980 BFTC 432 155

7/11/1984 1983 BFTC 200 140

9/16/1987 1987 BFTC 500 34

6/30/1989 1988 BFTC 271 161

5/19/1993 1992 Little South Fork Poudre River (via BFRH) 54 99

Big Thompson River 1922-23 Yellowstone cutthroat trout --- ---

9/23/1992 1992 BFTC 2,700 67

9/23/1992 1992 BFRH 100 fry

N.F. Big Thompson River 10/19/1970 Como Creek 50 125

9/16/1987 1987 BFTC 3,000 34

6/30/1989 1988 BFTC 90 161

9/14/1989 1989 BFTC 7,000 40

10/11/1990 1990 BFTC 1,500 64

Black Hollow Creek 9/1969 Albion Creek 10 mixture

1970 Como Creek 40 mixture

8/15/1980 Como Creek 37 115

10/6/1982 1982 BFTC 850 43

9/26/1983 1983 BFTC 935 46

7/11/1984 1983 BFTC 75 140

9/11/1984 1984 BFTC 600 36

9/16/1987 1987 BFTC 3,000 34

Bruno Gulch 9/1986 Zinn Ranch Ponds 30 100

9/24/1986 1986 BFTC 1,180 36

9/16/1987 1987 BFTC 6,750 34

9/14/1989 1989 BFTC 5,120 40

10/11/1990 1990 BFTC 5,000 64

Caddis Lake 8/6/1959 Big Thompson River 97 175

9/4/1959 Big Thompson River 112 175

1962 Pikes Peak cutthroat trout --- ---

Cony Creek 9/14/1989 1989 BFTC 8,500 40

10/11/1990 1990 BFTC 1,500 64

Water Date Source Number

Length

(mm)

(con.)
20 USDA Forest Service Gen. Tech. Rep. RMRS-GTR-88WWW. 2002

7/2/1991 Pear Lake 130 224

Cornelius Creek 9/26/1983 1983 BFTC 2,308 46

7/11/1984 1983 BFTC 310 140

9/11/1984 1984 BFTC 2,000 36

Crystal Lake 9/11/1984 1984 BFTC 685 36

9/25/1985 1985 BFTC 1,300 36

9/24/1986 1986 BFTC 1,090 36

9/14/1989 1989 BFTC 11,764 40

Dream Lake 7/11/1997 Upper Hutcheson Lake 190 adults

7/11/1997 Upper Hutcheson Lake 5 juveniles

Fern Creek 9/26/1983 1983 BFTC 560 46

9/11/1984 1984 BFTC 414 36

9/25/1985 1985 BFTC 1,100 36

Fern Lake 10/5/1982 1982 BFTC 900 43

9/26/1983 1983 BFTC 4,699 46

9/11/1984 1984 BFTC 3,679 36

George Creek 9/26/1983 1983 BFTC 2,618 46

7/11/1984 1983 BFTC 200 140

9/11/1984 1984 BFTC 2,000 36

Hidden Valley Creek 10/16/1973 Como Creek 82 155

6/30/1989 1988 BFTC 814 161

10/11/1990 1990 BFTC 736 64

Hourglass Creek 10/1966 Black Hollow Creek 54 mixture

8/15/1980 Como Creek 34 115

7/15/1981 1980 BFTC 158 155

10/5/1982 1982 BFTC 1,110 43

Husted Lake 9/24/1986 1986 BFTC 1,820 36

9/16/1987 1987 BFTC 2,000 34

9/25/1991 1991 BFTC 4,000 58

9/23/1992 1992 BFTC 2,740 67

9/23/1992 1992 BFRH 260 fry

Hutcheson Lake, Lower 6/30/1989 1988 BFTC 1,340 161

9/14/1989 1989 BFTC 1,700 40

9/25/1991 1991 BFTC 1,600 58

Hutcheson Lake, Upper 1952 Cutthroat trout 1,200 ---

1964 Cutthroat trout 3,000 ---

Lake Louise 9/14/1989 1989 BFTC 3,500 40

Lawn Lake 9/11/1984 1984 BFTC 6,415 36

9/25/1985 1985 BFTC 8,000 36

9/24/1986 1986 BFTC 6,830 36

9/14/1989 1989 BFTC 1,000 40

Loomis Lake 9/25/1991 1991 BFTC 1,000 58

9/23/1992 1992 BFTC 639 67

9/23/1992 1992 BFRH 361 fry

Lost Lake 9/16/1987 1987 BFTC 4,000 34

6/30/1989 1988 BFTC 1,356 161

9/14/1989 1989 BFTC 3,000 40

May Creek 8/15/1980 Como Creek 54 115

10/5/1982 1982 BFTC 1,700 43

9/27/1983 1983 BFTC 701 46

7/11/1984 1983 BFTC 90 140

9/11/1984 1984 BFTC 800 36

9/16/1987 1987 BFTC 3,000 34

Odessa Lake 9/11/1984 1984 BFTC 619 36

9/25/1985 1985 BFTC 1,100 36

9/14/1989 1989 BFTC 4,000 40

Ouzel Creek 9/15/1981 1981 BFTC 8,570 29

10/6/1982 1982 BFTC 1,733 43

9/26/1983 1983 BFTC 970 46

Ouzel Lake 9/15/1981 1981 BFTC 6,420 29

10/6/1982 1982 BFTC 3,467 43

9/26/1983 1983 BFTC 3,269 46

Pear Lake 6/30/1989 1988 BFTC 3,164 161

9/14/1989 1989 BFTC 5,000 40

10/11/1990 1990 BFTC 1,500 64

Pennock Creek 9/24/1986 1986 BFTC 4,560 36

9/16/1987 1987 BFTC 8,400 34 (con.)

Table 5 (Con.)

Water Date Source Number

Length

(mm)
USDA Forest Service Gen. Tech. Rep. RMRS-GTR-88WWW. 2002 21

Table 5 (Con.)

Roaring Creek 1912 Cutthroat trout 9,000 ---

Roaring River 9/11/1984 1984 BFTC 1,000 36

9/25/1985 1985 BFTC 3,700 36

9/24/1986 1986 BFTC 3,640 36

9/16/1987 1987 BFTC 6,000 34

Sandbeach Lake 6/30/1989 1988 BFTC 1,862 161

9/14/1989 1989 BFTC 4,000 40

East Fork Sheep Creek 10/5/1982 1982 BFTC 3,600 43

9/27/1983 1983 BFTC 1,100 46

7/11/1984 1983 BFTC 150 140

9/11/1984 1984 BFTC 500 36

9/25/1985 1985 BFTC 2,500 36

9/16/1987 1987 BFTC 3,000 34

West Fork Sheep Creek 10/5/1982 1982 BFTC 3,600 43

9/27/1983 1983 BFTC 1,000 46

7/11/1984 1983 BFTC 150 140

9/11/1984 1984 BFTC 500 36

9/25/1985 1985 BFTC 2,500 36

9/16/1987 1987 BFTC 3,000 34

Spruce Lake 9/25/1991 1991 BFTC 1,000 58

9/23/1992 1992 BFTC 1,005 67

9/23/1992 1992 BFRH 495 fry

West Creek 6/20/1979 Hidden Valley Creek 58 180

7/11/1984 1983 BFTC 100 140

9/11/1984 1984 BFTC 276 36

9/16/1987 1987 BFTC 2,000 34

9/14/1989 1989 BFTC 1,500 40

Williams Gulch 6/26/1981 1980 BFTC 48 155

10/6/1982 1982 BFTC 1,700 43

9/29/1983 1983 BFTC 421 46

7/11/1984 1983 BFTC 230 140

9/11/1984 1984 BFTC 800 36

9/16/1987 1987 BFTC 500 34

Zimmerman Lake 7/2/1996 1995 Hunters Creek-Little South Fork Poudre River (via

BFRH)

1,622 137

Zinn Ranch Ponds 9/25/1985 1985 BFTC 810 36

9/24/1986 1986 BFTC 730 36

_ 9/16/1987 1987 BFTC 700 34

Water Date Source Number

Length

(mm)
22 USDA Forest Service Gen. Tech. Rep. RMRS-GTR-88WWW. 2002

Table 6 -- Characteristics of greenback cutthroat trout delivered to hatcheries, including hatchery used, year spawned, and source and number of eggs or milt sent to hatcheries for

recovery efforts. If eggs were used, the number of females and males is given in parentheses. A number followed by a plus sign (+) indicates the minimum number of a sex that was

used. If milt was used, the number of males providing milt is noted, and the number of containers used to ship the milt is given in parentheses. Hatchery abbreviations are: BFRH, Bellvue

Fish Research Hatchery; BFTC, Bozeman Fish Technology Center; and SNFH, Saratoga National Fish Hatchery. Missing data are denoted by a dashed line.

Hatchery Year Source Eggs Milt Comments

Arkansas Basin

BFRH 1992 South Apache Creek 2,622 (---,---)

1992 SNFH 12,006 (264,264)

1993 Boehmer Reservoir 29,700 (---,---)

1994 Boehmer Reservoir 58,996 (---,---) In 1996, 660 offspring were mixed

with 56, 1993 Boehmer Reservoir

offspring.

1995 Boehmer Reservoir 89,472 (---,---)

1996 Boehmer Reservoir 9,824 (---,---)

SNFH 1984 Lytle Pond 1,743 (---,---) Fish from both sources originated

from Cascade Creek.

1984 McAlpine Lake 5,556 (5,6) In 1985, 504 offspring from both

sources were mixed.

South Platte Basin

BFRH 1989 Como Creek 847 (4,18) In 1989, 1020 offspring (77 from

Como Creek, 688 from Hunters

Creek, and 255 from Bear Lake) were

mixed.

1989 Bear Lake 602 (1,---)

1989 Hunters Creek 3,688 (11,10)

1989 Little South Fork Poudre River 282 (1,---) Eggs infertile.

1990 Upper Hutcheson Lake 305 (---,---)

1992 Hunters Creek 1,185 (9,---)

1992 Little South Fork Poudre River 760 (2,2)

1993 Hunters Creek 1,768 (11,23)

1994 Como Creek 580 (6,3+) In 1996, 10 offspring were mixed

with 100, 1993 Hunters Creek

offspring.

1995 Hunters Creek 4,734 (27,---) In 1996, 800 offspring were mixed

with 45, 1995 Little South Fork

Poudre River offspring.

1995 Little South Fork Poudre River 1,632 (---,---)

1996 Upper Hutcheson Lake 6,948 (20,---) In 1996, mixed with offspring of fish

spawned in hatchery.

1997 Hunters Creek 2,016 (13,18)

1997 Upper Hutcheson Lake 4,600 (---,---)

BFTC 1977 Como Creeka

1982 Hidden Valley Creek 12

1983 Como Creek 8

1983 Hidden Valley Creek 21

1984 Como Creek 12(8)

1984 Hidden Valley Creek 46(40)

1985 Como Creek 40(---) Up to 3 males per vial.

1985 Hidden Valley Creek 24

1986 Como Creek 18

1986 Hunters Creek 9

1986 Little South Fork Poudre River 2

1987 Como Creek 16

1987 Hunters Creek 17

1988 Como Creek 26(23)

1988 Hunters Creek 14

1989 Como Creek 22

SNFH 1984 Little South Fork Poudre River 500 (3,1) All eggs died.

_ 1985 Little South Fork Poudre River 353 (9,24) _ All fry died by February 1986.

a In 1977, 66 juvenile and adult fish -- of which 64 survived -- were sent from Como Creek to the BFTC to found its South Platte Basin broodstock.
USDA Forest Service Gen. Tech. Rep. RMRS-GTR-88WWW. 2002 23

Table 7 -- Characteristics of greenback cutthroat trout spawned in hatcheries as part of recovery efforts, including hatchery used, year spawned, source, year class name, number, and sex of fish

used, and eggs and fry produced. Source denotes the origin of fish being spawned. If the hatchery received fish or eggs from a wild population to support a broodstock, the year of arrival in the

hatchery and the water providing the fish are noted. If fish were derived from spawning in a hatchery, the year of spawning and hatchery are noted. To the extent possible, only numbers of fish

successfully producing viable offspring are included. If the sexes of fish of different sources were known, it is indicated in parentheses (F, female; M, male) or overall in the following column.

Abbreviations are: BFRH, Bellvue Fish Research Hatchery; BFTC, Bozeman Fish Technology Center; and SNFH, Saratoga National Fish Hatchery. Missing data are denoted by a dashed line.

Hatchery Year Source Year class F,M Eggs Fry Comments

Arkansas Basin

BFRH 1996 1993 Boehmer Reservoir 1996a BFRH 118,201 75,688 ---

SNFH 1987 1984 Lytle Pond-McAlpine

Lake

1987 SNFH 169,219 182,336 ---

1988 1984 Lytle Pond-McAlpine

Lake

1988 SNFH 151,200 281,626 ---

1989 1984 Lytle Pond-McAlpine

Lake

1989 SNFH 108,190 244,422 ---

1990 1984 Lytle Pond-McAlpine

Lake

1990 SNFH 32,40 76,930 ---

1991 1988 SNFH 1991 SNFH 236,225 273,383 ---

1992 1988 SNFH 1992 SNFH 150,150 292,113 --- All fry produced this

year were combined.

1992 1989 SNFH 1992 SNFH 114,114 94,280 ---

South Platte Basin

BFRH 1991 1989 Bear Lake-Como CreekHunters Creek

1991 BFRH 7,--- 4,006 446

1992 1989 Bear Lake-Como CreekHunters Creek

1992 BFRH 17,--- 9,590 3,500

1993 1990 Upper Hutcheson

Lake-1991 BFRH

1993 BFRH 30,--- 33,406 2,250

1996 1993 Hunters Creek 1996b BFRH 52,62 41,993 ---

BFTC 1978 1977 Como Creek 1978 BFTC 2,--- 1,005

1979 1977 Como Creek 1979 BFTC 2,--- 1,377 268

1980 1977 Como Creek 1980 BFTC 7,--- 3,704 900

1981 1978 BFTC 1981 BFTC 11,--- 6,065 ---

1981 1979 BFTC 1981 BFTC 90,--- 35,021 ---

1982 1979 BFTC(F),1982 Hidden

Valley Creek(M)

1982 BFTC 30,12 --- ---

1982 1979 BFTC 1982 BFTC ---,--- --- --- In 1982, 32,000 fry

produced overall.

1983 1979 BFTC 1983 BFTC 25,13 --- ---

1983 1979 BFTC(F),1983 Como

Creek(M)

1983 BFTC 7,8 --- ---

1983 1979 BFTC(F),1983 Hidden

Valley Creek(M)

1983 BFTC 35,18 --- ---

1984 Mixed BFTC(F),1984 Hidden

Valley Creek(M)

1984 BFTC 25,33 --- ---

1984 Mixed BFTC 1984 BFTC 126,58

1985 Mixed BFTC(F),1985 Como

Creek(M)

1985 BFTC 95,25 --- ---

1985 Mixed BFTC(F),1985 Hidden

Valley Creek(M)

1985 BFTC 120,24 --- ---

1986 Mixed BFTC(F),1986 Como

Creek(M)

1986 BFTC 52,18 --- ---

1986 Mixed BFTC(F),1986

Hunters Creek(M)

1986 BFTC 20,9 --- ---

1986 Mixed BFTC(F),1986 LSF

Poudre River(M)

1986 BFTC 5,2 --- ---

1986 Mixed BFTC 1986 BFTC 164,--- --- --- In 1986, 210 females

used overall.

1987 Mixed BFTC,1987 Como

Creek(M)

1987 BFTC 64,16 75,087 ---

1987 Mixed BFTC,1987 Hunters

Creek(M)

1987 BFTC 65,17 36,825 ---

1987 Mixed BFTC 1987 BFTC 95,60 37,621 --- In 1987, 63,633 fry

produced overall.

1988 Mixed BFTC,1988 Como

Creek(M)

1988 BFTC 53,26 --- ---

1988 Mixed BFTC,1988 Hunters

Creek(M)

1988 BFTC ---,14 --- ---

1988 Mixed BFTC 1988 BFTC 146+,70+ --- ---

1989 Mixed BFTC,1989 Como

Creek(M)

1989 BFTC ---,22+ --- ---

1989 Mixed BFTC 1989 BFTC ---,--- --- --- 1n 1989, 315 females

used overall.

1990 Mixed BFTC 1990 BFTC 51,86 85,000 ---

1991 Mixed BFTC 1991 BFTC 65,--- 98,000 ---

1992 Mixed BFTC 1992 BFTC 50,--- 60,000 ---
24 USDA Forest Service Gen. Tech. Rep. RMRS-GTR-88WWW. 2002

Table 8 -- Characteristics of greenback cutthroat trout distributed by hatcheries as part of recovery efforts, including hatchery used, year

shipped, year class name, number, and mean length (mm) of fish shipped. Shipments for disease testing, for genetic analyses, for fish

collections, or to support sport fisheries were excluded when possible. See tables 6 and 7 for origins of year classes. Multiple shipments of

single sources in one year were distributed on different dates. Hatchery abbreviations are: BFRH, Bellvue Fish Research Hatchery; BFTC,

Bozeman Fish Technology Center; and SNFH, Saratoga National Fish Hatchery. Missing data are denoted by a dashed line.

Hatchery Year Year class Number Length

Arkansas Basin

BFRH 1992 1992 SNFH 5,100 38

1993 1992 South Apache Creek 1,750 150

1993 1992 SNFH 600 150

1994 1993 Boehmer Reservoir 6,750 147

1995 1994 Boehmer Reservoir 2,080 ---

1995 1994 Boehmer Reservoir 4,020 ---

1996 1994 Boehmer Reservoir 4,445 ---

1996 1995 Boehmer Reservoir 360 111

1996 1995 Boehmer Reservoir 26,140 116

1996 1996a BFRH 1,400 41

SNFH 1985 1984 Lytle Pond 225 120

1985 1984 McAlpine Lake 506 97

1985 1984 Lytle Pond 155 124

1985 1984 McAlpine Lake 1,700 93

1987 1987 SNFH 14,581 50

1988 1988 SNFH 23,162 eggs

1988 1988 SNFH 59,334 25

1989 1989 SNFH 66,684 38

1989 1988 SNFH 2,394 203

1989 1988 SNFH 1,680 213

1990 1989 SNFH 2,157 120

1990 1989 SNFH 3,505 120

1990 1990 SNFH 15,843 33

1990 1989 SNFH 1,250 141

1991 1989 SNFH 2,273 235

1991 1991 SNFH 22,983 49

1991 1991 SNFH 34,514 50

1992 1992 SNFH 12,006 eggs

1992 1992 SNFH 9,800 38

1992 1992 SNFH 2,962 43

1992 1989 SNFH 15 ---

South Platte Basin (con.)
USDA Forest Service Gen. Tech. Rep. RMRS-GTR-88WWW. 2002 25

South Platte Basin

Table 8 (Con.)

BFRH 1992 1992 BFRH 2,000 ---

1993 1992 Little South Fork Poudre River 54 99

1993 1992 Hunters Creek 900 ---

1993 1992 BFRH 1,350 127

1993 1993 BFRH 2,250 ---

1996 1995 Hunters Creek 1,600 137

1996 1995 Little South Fork Poudre River 200 137

BFTC 1981 1980 BFTC 865 155

1981 1981 BFTC 16,570 29

1982 1982 BFTC 26,200 43

1982 1982 BFTC 1,150 ---

1983 1983 BFTC 23,000 46

1984 1983 BFTC 1,859 140

1984 1984 BFTC 39,623 36

1985 1985 BFTC 27,000 36

1986 1986 BFTC 29,724 36

1987 1987 BFTC 60,220 34

1989 1988 BFTC 9,221 161

1989 1989 BFTC 55,764 40

1990 1990 BFTC 12,920 64

1991 1991 BFTC 12,498 58

_ 1992 1992 BFTC 19,448 67

Hatchery Year Year class Number Length
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