Species of Aquilapolknites 

and Fibulapollis From Two 

Upper Cretaceous Localities 

in Alaska

GEOLOGICAL SURVEY PROFESSIONAL PAPER 643-A
Species of Aquilapollenttes 

and Fibulapollis From Two 

Upper Cretaceous Localities 

in Alaska

By BERNADINE D. TSCHUDY

CONTRIBUTIONS TO PALEONTOLOGY

GEOLOGICAL SURVEY PROFESSIONAL PAPER 643-A

Occurrence, description, and 

illustration of fossil pollen

UNITED STATES GOVERNMENT PRINTING OFFICE, WASHINGTON : 1969
UNITED STATES DEPARTMENT OF THE INTERIOR 

WALTER J. HIGKEL, Secretary

GEOLOGICAL SURVEY 

William T. Pecora, Director

For sale by the Superintendent of Documents, U.S. Government Printing Office 

Washington, D.C. 20402 - Price $1 (paper cover)
CONTENTS

Abstract. __________________________________________

Introduction ______________________________________

Photography.__________________________________

Type slides___________________________________

Sample localities and their respective species of 

Aquilapollenites and Fibulapollis... ______________

Relative abundance of Aquilapollenites, Fibulapollis, 

and other paly nomorphs___ ___________________

Acknowledgments__ _ _ _________________________

Summary. _________________________________________

Systematic descriptions._____________________________

Form genus Aquilapollenites (Rouse) Funkhouser, 

1961________________________________________

Aquilapollenites senonicus (Mtchedlishvili) 

Tschudy and Leopold, 1969_____________

Aquilapollenites clarireticulatus (Samoilovitch) 

n. comb________________________________

Page

Al

1

1

1

Systematic descriptions Continued

Form genus Aqwlapollenites, etc. Continued 

Aquilapollenites trialatus Rouse, 1957____

Aquilapollenites parallelus n. sp_________

Aquilapollenites fusiformis n. sp__ _______

Aquilapollenites dentatus n. sp__ ______

Aquilapollenites colvillensis n. sp_________

Aquilapollenites rectus n. sp _________

Aquilapollenites scabridus n. sp ________

Aquilapollenites contiguus n. sp_ ________

Aquilapollenites laticorpus n. sp_________

Form genus Fibulapollis Chlonova, 1961___._ 

Fibulapollis scabratus n. sp _____________

References cited-_____________-___-__-__---___

Index _____________--____-____--_--------_---

Page 

A7

7

8

9

9

10

11

12

12

13

13

14

17

ILLUSTRATIONS

[Plates 1-16 follow index]

PLATE 1. Aquilapollenites senonicus (Mtchedlishvili) Tschudy and Leopold, 1969.

2. Aquilapollenites darireticulatus (Samoilovitch) n. comb.

3. Aquilapollenites clarireticulatus (Samoilovitch) n. oomb.

4. Aquilapollenites trialatus Rouse, 1957.

5. Aquilapollenites parallelus n. sp.

6. Aquilapollenites parallelus n. sp.

7. Aquilapollenites parallelus n. sp.

8. Aquilapollenites fusiformis n. sp.

9. Aquilapollenites dentatus n. sp.

10. Aquilapollenites dentatus n. sp.

11. Aquilapollenites colvillensis n. sp.

12. Aquilapollenites rectus n. sp.

13. Aquilapollenites scabridus n. sp. and Aquilapollenites rectus n. sp.

14. Aquilapollenites scabridus n. sp.

15. Aquilapollenites contiguus n. sp.

16. Aquilapollenites laticorpus n. sp. and Fibulapollis scabratus n. sp.

FIGURE 1. Index map of Alaska showing sample localities and their respective species of Aquilapollenites and Fibulapollis-...

Page 

A2

TABLES

TABLE 1. Rocky Mountain pollen localities__________________________---__---__-_-

2. Relative abundance of Aquilapollenites, Fibulapollis, and other palynomorphs.

Page

A3

3

in

CONTRIBUTIONS TO PALEONTOLOGY

SPECIES OF AQUILAPOLLENITES AND FIBULAPOLLIS FROM TWO UPPER CRETACEOUS

LOCALITIES IN ALASKA

By BERNADINE D. TSGHUDY

ABSTRACT

Eight new species of Aquilapollenites from northern and 

eastern Upper Cretaceous localities in Alaska are named and 

described. Three previously described species of Aquilapollenites 

also were present in the samples at these localities; two of 

these species were originally reported from Cretaceous sedimentary rocks of Russia and one was described from Upper 

Cretaceous sedimentary rocks from Canada. The latter three 

species are also present in Upper Cretaceous rocks of the northern Rocky Mountains of the United States. None of the eight 

new species of Aquilapollenites has been observed in samples 

from the Rocky Mountain region.

One species of the genus Fibulapollis, a genus probably 

closely allied to Aquilapollenites, was found in the Alaskan 

samples, but it occurs only as rare specimens. The species is 

also present in some Upper Cretaceous samples from the 

Rocky Mountain region and is similar to two previously described species from Siberia.

INTRODUCTION

Since 1957, many species of AquilapoUenites have 

been reported from Upper Cretaceous and (or) Paleocene sedimentary rocks from the Rocky Mountain regions of Canada and the United States and from Russia, 

Japan, Malaysia, Scotland, and Africa. (See summary 

in Tschudy and Leopold (1969).) Recently, forms pertaining to this genus have been recognized in Alaskan 

sedimentary rocks by oil company palynologists and by 

workers in the U.S. Geological Survey Denver Pollen 

Laboratory; however, Alaskan species of Aqufla-pollenites have not been reported in the literature. Judging 

from the diversity of forms found in the few samples 

reported here, the taxa described probably represent a 

very small percentage of the total number of species of 

the genus which exist in the Upper Cretaceous of 

Alaska.

Two species of the genus Fibwlapollis were previously 

described from sediments from the Chulym-Enisei depression (western Siberia) of the U.S.S.R. A species 

of Fibula/pollis similar to the two species reported from

Russia is present in the Upper Cretaceous samples from 

the two Alaskan localities of this report and is also present in Upper Cretaceous sedimentary rocks from localities in the Rocky Mountain region.

PHOTOGRAPHY

The transmitted light photographs which illustrate 

the taxa in this report were taken on a Zeiss photo^cope. 

Apochromatic X 40 and Neofluar X 100 oil immersion 

objectives and 35 millimeter KB-14 film were used. Four 

of the new species are illustrated by scanning electron 

microscope (SEM) photographs as well as by standard 

light microscope photographs. The SEM photographs 

were taken by Charles M. Drew at the Naval Weapons 

Center at China Lake, Calif.; methods used in the preparation and photographing of these pollen grains are 

the same as those described by Drew and Tschudy (1968, 

p.1830).

TYPE SLIDES

Slides containing type specimens of new taxa, as well 

as slides containing nontype grains represented 011 the 

plates, are on file at the U.S. Geological Survey Denver 

Pollen Laboratory. The type specimens are within black 

ink circles marked on the slides; they may also be located by the mechanical stage coordinates which are 

included in the plate explanations and with the individual holotype and paratype assignments. The author's coordinate readings for the center point of a 1- by 

3-inch standard microscope slide are 108.2 X 12.4 mm 

(horizontal X vertical axes); with the slide label placed 

to the left on the microscope stage, the vertical coordinates decrease toward the bottom edge of the slic'e and 

the horizontal coordinates decrease toward the right 

edge of the slide. Conversion of coordinates to those of 

another mechanical stage may be made by the methods 

of Traverse (1958) and Tschudy (1966).

Al
A2 CONTRIBUTIONS TO PALEONTOLOGY

SAMPLE LOCALITIES AND THEIR RESPECTIVE 

SPECIES OF AQUILAPOLLENITES AND FIBULAPOLLIS

The samples used in this study are from the pollen 

slide files of the U.S. Geological Survey Denver Pollen 

Laboratory. The localities from which the Alaskan 

samples were collected and the species of Aquilapottenites and F ibulapoUis which were found at each locality 

are designated in figure 1.

Five samples (A, B, C, G, H) from USGS paleobotany locality D3124 were collected July 18, 1951, along 

the Colville River in northern Alaska, lat 69°40' N., long 

151°25' W., in T. 4 N., R. 2 E. (Umiat quadrangle), by 

C. A. Arnold, R. A. Scott, and J. S. Lowther. These 

samples are from the Colville Group of Late Cretaceous 

age.

One sample, from USGS paleobotany locality D1799,

was collected in 1961 on the east bank of the Nation 

River in eastern Alaska, lat 65°19.6' N., long 141°29.6' 

W., in SEi/4 sec. 23, T. 6 N., R. 30 E. (Charley River 

B-l quadrangle), by E. E. Brabb. This sample was 

taken from rocks of Late Cretaceous age.

Rocky Mountain pollen localities mentioned in this 

report are listed in table 1.

RELATIVE ABUNDANCE OF AQUILAPOLLENITES. 

FIBULAPOLLIS , AND OTHER PALYNOMORPHS

Palynomorph counts were made for four of the more 

productive samples from the two localities in Alaska. 

The species of AquilapoUenites and FibulapoUis present and their abundance in respect to each other and to 

other palynomorphs are shown in table 2. In this table 

the other palynomorphs found in the sampler are classified only under genera or groups.

A. colvillensis 

A. contigit'is 

A. fusiformis 

Micorpus 

A. senonicus 

A. trialatus 

F. scabratus

A. clarireticulatus 

A. dentatus 

A. parallelus 

A. scabridux

A. senonicus

A. rectus

F. scabratus

FIGURE 1. Index map of Alaska showing sample localities and their respective species of Aquilapollenites and FibulapolUs.
SPECIES OF AQUILAPOLLENITES AND FIBULAPOLLIS, ALASKA A3

TABLE 1. Rocky Mountain pollen localities

USGS paleobotany localit; 

No.

D133(M ...

D1330-5.. _.._-. ___ --

State

.__ _do __ -._-

D1330-19.. _.--.. ..do.... __... 

D1330-26.. ......... _.do..___. ...

D 1331-2, -5-__.

D1602--. -_--_. 

D1603. .---.

D1750-B _

D3147-B _ _.

D3472-B, -C, 

-D,-E,-F. 

D3690-B 

D3690-C, -D, 

-H 

D3718-E,-!.-.

D3724-B, -E-.

D3725-A. _..

D3730-A, -D__

D3740-B..._... 

D3754-A ._.

D3980-E-.-_-_.

. -.do. .

......do..... , 

 do.. .

 do... ......

.... .do ...

 do. .

- do ....

.._. do ..

.... ...do . 

. ... do.... 

. .do 

Wyoming ___ 

. Montana. _----

.. Colorado.. ....

County

Niobrara. . . 

_ .do. .

.-_.-do . 

 do......

_-.--do _.

 do 

 do..... .

.....do-. .

 do 

Sheridan. _.

Locality

Sec.

..-- NEMNEX-- 

.... NW^NE^...

NE^NWW.-. 

.... SE&SEK -

.... SW^SWM---

.... C_. _...- - 

.... NEJ^SEJi -

._ . NWJ4- ...

. - NW^NW^.__

.... NEJiSWJi....

.... NWJi- ..

___ NEJ^NWM

NWM-

.... C--. ... 

23 

23 

23 

14 

14 

3 

7 

12 

1 

29

14

14

13 

12 

26 

2 

19

29 

36

T.

38 N. 

38 N. 

38 N. 

38 N. 

38 N. 

37 N. 

37 N. 

15 N. 

21 N. 

21 N.

141 N. 

141 N.

22 N. 

22 N. 

24 N. 

141 N. 

57 N.

21 N.

47 N.

R.

62 W. 

62W. 

62W. 

62W. 

62W. 

64W. 

64W. 

92W. 

37 E. 

34 E.

55 E. 

55 E.

17 E. 

17 E. 

17 E. 

55 E. 

86W.

34 E.

7W.

TABLE 2. Relative abundance, in percent, of the species of 

Aquilapollenites, Fibulapollis, and other palynomorphs

[ Palynomorphs not identified to species are listed merely under genera or larger 

groups. Numbers represent percentages and are based on counts of 200 palynomorphs 

per sample; plus sign indicates presence of palynomorphs in samples where they 

did not occur within counts]

Sample localities

D1799 D3124^B D3124-G D3124-H

Aquilapollenites darmtic.ula.tus. -- ... ... _ ...

colvittensis---. -------- _ . _ _ ..-. ....

contiguus. ............ ... ---------- _ ..

dentatus...... ...................... ... .

fusifor mis... --------------------------- -

laticorpus.. .............................

parallelus... ... ... .....-..-. ... -..-. _ .

rectus-. ............ .......... .........

scabridus .... ... .... . ... ........ ... .....

senonicus. ..............................

trialatus... ............................

Fibulapollis scabratus....... ... . ... ... _ .-.

Fern spores:

Monolete.--------. _ -------- . _ ...-

T|rilete-.-_ .............................

Conifer pollen:

cf. Dacrydium.... .......................

cf. Sequoia............... _---... _ ---..

Inaperturopollenites....... . ........... 

Other conifer pollen...-...-..-.-.---...

Nonconifer pollen:

Monosulcate... ... _ ...................

Erdtmanipollis . ... ............. .... ...

Cranwellia.. ... ....... ... ...............

Loranthacites (sensu Mtchedlishvili,

1961) . .-

Miscellaneous triporate and tricolpate

pollen. .............. _ ....... _ ...

Algae:

Botryococcus.... ... ... ....... .... . ... ...

Incertae sedis _ -.-. --.-.------.... _ .....

Total:

Aquilapollenites ... ..................

Other palynomorphs- --------------

. 4

0

0

7.5

0

0

2.5

10

+

+

0

+

42

3.5

0

0

0 

0

5

.5

11

4.0

10

0

0

24

76

100

0

39.5

2.5

0

.5

.5

0

0

0

1

0

+

2

10.5

0

0

.5 

.5

22.5

0

0

0

.5

10.5

9

44

56

100

0

20.5

6

0

1

0

0

0

0

+

0

+

6

4

5

0

.5 

32.5

8.5

0

+

0

.5

7.5

8

27.5

72.5

100

0

4

0

0

0

0

0

0

0

0

1.5

0

3

8.5

0

2.5

2.5 

1

3

0

+

0

.5

0

73.5

5.5

94.5

100

ACKNOWLEDGMENTS

Glenn E. Rouse furnished a slide of his type material 

from Canada which contained pollen useful for comparison with Alaskan and Rocky Mountain forms. Ivan 

J. Mittin translated the parts of Russian publications 

that are included in this report. A. F. Chlonova's 

prompt replies to correspondence, as well as her examination of and comments about pollen which was 

sent to her, are greatly appreciated. The author is particularly grateful to Estella B. Leopold for helpful

advice and criticisms during the entire preparation of 

this report.

The scanning electron microscope photographs were 

generously contributed by Charles M. Drew of the 

Naval Weapons Center, China Lake, Calif.

SUMMARY

Eight new species of Aquilapollenites and one new 

species of Fibidapollis are named and described from 

two Upper Cretaceous localities in Alaska. Known distributions of three previously named species of Aquilapollenites which were present in these samples are 

noted, and two of these species are redescribed.

The eight new species of Aquilapollenites appear to 

be distinctly different from any species which have been 

encountered in sedimentary rocks from the Rocky 

Mountain region and from species which have been 

reported from the U.S.S.R. Four of the eight new 

species were found in the sample from eastern Alaska; 

the remaining four, in samples from the northern 

Alaska locality.

Of the three previously known species of Aquilapollenites which were found in the Alaskan samples: One 

was described from Senonian sedimentary rocks from 

the Western Siberian lowlands (Sym River) and has 

been found to range from lower Campanian into upper Maestrichtian in the Rocky Mountains; another 

species, previously named from the MaestrichtianDanian of Yakutia (Linde River, central Siberia), has 

been found in the lower Campanian Eagle Sandstone 

in Montana; and the third, described from Upper 

Cretaceous seidmentary rocks of Alberta, Canada, has 

been found in the middle Campanian Claggett Shale in 

Montana. Grains very similar (or equal) to this last 

species have been reported from Siberia.

The one species of Fibulapollis which was present 

in samples from both Alaskan localities has been found 

to range from lower Campanian into lower Maestrichtian, in the Rocky Mountains. This taxon is similar to 

two species of Fibulapollis previously described from 

the Chulym-Enisei depression of Western Siberia.

SYSTEMATIC DESCRIPTIONS

Form genus Aquilapollenites (Rouse) Funkhouser, 19"1

Aquilapollenites Rouse, 1957, p. 370-371. Type species: Aquilapollenites quadrilobus Rouse, 1957. Type specimen: 

Designated by Rouse (1957, p. 371) from the Brazeau 

Formation, Upper Cretaceous, Alberta, Canada [McMaster 

University Palaeobotanical Collection, Hamilton, Ontario, 

Canada, slide Cs-31] ; illustrated in Riadforth and Rouse 

(1954, pi. 1, fig. 14), and in Tschudy and Leopold (1969, 

pi. 2, figs. 1A-B).

Aquilapollenites (Rouse) Funkhouser, 1961 (April), p. 193-194.
A4 (CONTRIBUTIONS TO PALEONTOLOGY

Triprojectacites Mtchedlishvili, 1961 (in Samoilovitch and

others, 1961 (July), p. 203-204) (supergroup of

Mtchedlishvili). 

Triprojectus Mtchedlishvili, 1961 (in Samoilovitch and others,

1961 (July), p. 204-205). 

Aquilapollenites Rouse et Mtchedlishvili, 1961 (in Samioilovitch

and others, 1961 (July), p. 209). 

Integricorpus Mtchedlishvili, 1961 (in Samoilovitch and others,

1961 (July), p. 217). 

Mancicorpus Mtchedlishvili, 1961 (in Samoilovitch and others,

1961 (July), p. 218-219). 

Parviprojectux Mtchedlishvili, 1961 (in Samoilovitch and others,

1961 (July), p. 225). 

Projectoporites Mtchedlishvili, 1961 (in Samoilovitch and

others, 1961'(July), p. 227). 

Tricerapollis Chlonova, 1961 (August), p. 85. 

Taurocephalus Simpson, 1961 (December), p. 440. 

Mancicorpus (Mtchedlishvili) Srivastava, 1968b, 695-697. 

Aquilapollenites (Rouse) Funkhouser, 1961 (in Tschudy and

Leopold, 1969).

Aquilapollenites senonicus (Mtchedlishvili) Tschudy and Leopold,

1969

Plate 1, figures 1-14

Mancicorpus senonicum Mtchedlishvili, 1961 (in Samoilovitch 

and others, 1961, p. 224-225). Holotype: pi. 72, figs. 2a-c; 

Paleophytological Laboratory of the Scientific Research 

Institute of Petroleum and Geological Exploration 

(VNIGRI), preparation 6196; Western Siberian lowlands, 

Sym River, Kolokol'nikov Yar, specimen 8528; lower 

Symian subsuite, Senonian.

Mancicorpus senonicus Mtchedlishvili, 1961 (in Srivastava, 

1968a, p. 1488).

Aquilapollenites senonicus (Mtchedlishvili) Tschudy and 

Leopold, 1969.

The following is a translation by Ivan Mittin of the 

description of Mancicorpus senoniciim Mtchedlishvili, 

1961 (in Samoilovitch and others, 1961, p. 224-225) :

"Diagnosis. Pollen grains are large, three furrowed. 

Proximal (?) end of body cylindrical, with rounded end. 

There is a distal ( '1} node. Equatorial projections in contact with the body form an angle nearly at right angles. 

Exine of medium thickness, clublike (pilata), reticulate.

"Description. Pollen grains 32.5/A to 36.5/* [microns] 

in height, average height 34.6^; its diameter from 

44.2/x to 48.6/t, average 46.5/A; the length of the body from 

30.5/A to 35.4/*, average 32.5/*; width of body 

9.7p. to 14.9/A, average 12.5/*; length of equatorial projections from 15.8/i to 19.1/*, average IT.5/^; width of 

equatorial projections 10.1/u, to 12.1/*, average ll.lju.

"The pollen grains are three furrowed. The proximal 

end of the body is elongated-rectangular in form with 

the rounded end that becomes wider in the direction of 

the equatorial projections. Distal (?) end is represented 

by a small node. Equatorial projections joining the body 

nearly at right angles. Projections are long and rather

wide, furrows long and narrow. Exine about 1.3/* thick, 

the layer of nexine of the grain body is almost one half 

as thick as the sexine and it becomes wider near the 

projections; thick parts of the nexine are long for more 

than one half of the length of the projections, the structure of sexine is clublike with the rods thin, the heads 

small, rounded, and they converge at the edge of the 

body forming a sheath. The greatest thickness of the 

sexine layer is observed at the place of transition of the 

body into the equatorial projections. The sculpture is 

reticulate, the walls are composed of clublike protrusions, in plan having lenticular structure. The size of 

cells decreases toward the end of the body and equatorial projections. The ends of projections are covered with 

the exine that retains clublike structure which can be 

distinguished at magnification, X 1,350.

"Material. Ten well-preserved specimens.

"Variations. The pollen grains of this sp3cies vary a 

little, frequently the end of the body is slightly bent and 

in some specimens the thin ends of equatorial projections may be destroyed or broken.

"Comparison and remarks. By the dimension of 

pollen and the structure of the exine the species in the 

description resembles M. trapeziforme sp. n. It differs 

from it in the presence of a distal node, the form of the 

body and the size of the angle formed at the contact 

of the body with the equatorial projections It differs 

from M. tenue sp. n. in the size and the form of the body 

and in the length of equatorial projections,

"Locality. Western Siberian lowlands, River Vakh 

and drill hole 2T-K, interval 329.90-343.15 m [meters] 

collected by E. IS". Petrov, lower (?) Maestrichtian; 

River Sym, Kolokol'nikov Yar, specimen 8528, collected 

by S. V. Lebedev, Senonian (lower Symian subsuite) ; 

River Sym, upper Symian drill hole 14, specimen 18, 

interval 23.4-26.9 m, Maestrichtian.

"Distribution. Senonian of the eastern part of the 

Western Siberian lowlands."

Records of occurrence. 

Rock unit

TJSGS paleobot- Locality

any locality No. or 

literature reference State and (or) 

country

U.S.S.R. 

Lower Symian Senonian_......... Mtchedlishvili Western Siberia

subsuite. (in Samoilovitch

and others, 

1961). 

(?)..........-..... Lower(?) .....do... .... Do.

Maestrichtian. 

(?) __ Maestrichtian....... -...do- ........ Do.

Canada.

Edmonton Forma- Maestrichtian-_... Srivastava (1968a). Alberta, 

tion. U.S.A. 

(?)- .--. - Upper Cretaceous D1799- ......... Alaska.

Colville Group..-,........do............... D3124-B, -G...... Do.

(?) -..----.-........do ---- D1750-B.......... T'yoming.

Lance Formation, Upper Maestrich- D1602.__._... Do.

upper part. tian.

Hell Creek Forma- Upper and middle D3730-A,-D Montana, 

tion, middle part. Maestrichtian.

See footnote at end of table.
SPECIES OF AQUILAPOLLENITES AND FIBULAPOLLIS, ALASKA A5

USGS paleobot- Locality 1

Rock unit Stage or series any locality No. or 

literature reference State and (or) 

country

U.S.A. 

Hell Creek Forma- Middle Maestrich- D3147-B..... __ Montana.

tion, lower part. tian.

Do.. ............... ...do. ........... D3472-B, -C, -D,

" Do.

Do.... -_.-..._..--.. _.do _ ............ D3690-C, -D, -H. Do.

Do... ................. do.... ........... D3754-A-. ........ Do.

Lance Formation, -...-do.- -.----..-.. D1603. --...----... Wyoming.

lower part. 

Fox H'ills Sandstone, Middle or lower D3690-B... ___ Montana.

Colgate Member. Maestrichtian. 

Fox Hills Sandstone.. Lower Maestrich- D1331-2, -5 _ .... Wyoming.

tian. 

Judith River Forma- Upper Campanian.. D3725-A... __ ... Montana.

tion, upper part. 

Fruitland Formation.. ....do __ . _ ------ D3980-E.. ____ Colorado.

Judith River Forma- Middle Campanian . D3740-B-... __ .Wyoming.

tion, lower part. 

Pierre Shale, Mitten Middle or lower D1330-5.. ......... Do.

Black Shale Campanian.

Member. 

Eagle Sandstone, Lower Campanian.. D3718-E-. .--..... Montana.

middle part.

1 For specific Rocky Mountain localities see table 1.

Description. Based on 55 specimens from the U.S. 

Geological Survey collections listed above. Tricolpate, 

heteropolar pollen grains with three equatorial protrusions. In equatorial view : grains with a domelike major 

pole, a nodelike minor pole, and three equatorial protrusions extending at nearly right angles from polar 

axis of grain or drooping slightly toward minor pole. 

Polar axis 27/x. to 37/x,; equatorial diameter (including 

equatorial protrusions) 36/x, to 48/x; length of major 

polar protrusion 14/x, to 17.5/x, width (measured halfway 

between base of protrusion and the pole) 14/x. to 17.5/x.; 

length of equatorial protrusions 14/x, to 20/x, width 

(measured at midlength) 10.5/x. to 15/x; major polar protrusions usually slightly larger, in some cases equal to, 

or rarely smaller than equatorial protrusions; minor 

pole protruding slightly or in rare specimens not protruding. In polar view : grains triangular, with slightly 

convex, slightly concave, or irregularly concave-convex 

sides, with rounded or pointed corners. Apertures: 

colpi extending the length of the equatorial protrusions, 

nearly anastomosing at minor pole (in some grains colpi 

appear to anastomose ; in other grains colpi do not quite 

anastomose), colpi extend only a short distance onto 

base of major polar protrusion. Pollen wall: exine two 

layered; endexine (except for endexinal thickenings) 

thin or as much as 0.5/x, thick, very thin on ends of equatorial protrusions at colpal margins ; axillary endexinal 

thickenings, tear shaped to lens shaped, as much as 4/x, 

thick in thickest part, extending onto equatorial protrusions for approximately half the length of the protrusions, extending onto base of major polar protrusion, 

and reaching nearly to minor pole; endexinal thickenings often lacking, some specimens show distinct thin 

areas or cavities from which thickenings have probably 

been dissolved (pi. 1, figs. 1, 2) ; ektexine approximately 

1/x. thick on minor polar protrusion and on basal half of 

major polar protrusion; ektexine usually slightly 

thinner on major polar dome than on remainder of

351-255 O 69 2

body, ektexine on equatorial protrusions as much as 

0.5/x. thick; ektexinal elements columnar, joined at tips 

resulting in a reticulate body sculpture; lumin?, of 

reticulum on minor polar protrusion and on approximate basal half of major polar protrusion as much as 

0.5/x, in diameter; lumina on major polar dome usually 

smaller than on remainder of body; sculpture of equatorial protrusions very finely reticulate (magnification 

X 1,000) or granulate; granula less than 0.5/x. in diameter, with very low relief, scattered over surface of 

equatorial protrusions.

Remarks. The grains described above closely resemble Aquilapollenites (alias Mancicorpus) senov,icus 

as it was described and illustrated by Mtchedlishvili. 

Slightly wider ranges in size and form in the Alaskan 

specimens may be due to the much greater number of 

grains examined (Mtchedlishvili included a totr.l of 

only 10 specimens for his description). Variations in A. 

senonicus as described by Mtchedlishvili and as found 

in this study are discussed below:

1. Some of the Alaskan specimens have wider equatorial and major polar protrusions but a great 

many are as slender as those described by 

Mtchedlishvili.

2. A. senonicus was described by Mtchedlishvili as having equatorial projections joining the body nearly 

at right angles. The Alaskan specimens were 

found to have equatorial protrusions which ranged 

from a nearly right angle position to a slight tilt 

toward the direction of the minor pole.

3. Mtchedlishvili did not mention the shape or maximum thickness of the endexinal thickening?; he 

merely stated that thick parts of the nexine ertend 

for more than half the length of the (equatorial) 

projections. His photograph of the holotype (in 

Samoilovitch and others, 1961, pi. 72, fig. 2a) shows 

a thin area in an axilla which closely resembles the 

endexinal cavities in some of the Alaskan grains 

(pi. 1, figs. 1,2).

Similar species. A. senonicus strikingly resembles 

Aquilapollenites delicatus Stanley (1961, p. 346-347) 

but can easily be distinguished by its lack of spines on 

the major and minor polar protrusions. The shape of 

A. senonicus resembles that of Aquilapollenites quadrilobm Rouse (1957, p. 371) but its coarser body reticulum and lack of spines separate it from the latter 

species. A. senonicus appears to closely resemble "Mancicorpus" trapesi forme Mtchedlishvili (in Samoilovitch 

and others, 1961, p. 221-222) ; however, Mtchedlishvili 

stated that the former differs "in the presence of a distal 

node, the form of the body, and the size of the angle 

formed at the contact of the 'body with the ecumtorial 

projections" (translation by Ivan Mittin).
A6 (CONTRIBUTIONS TO PALEONTOLOGY

Aquilapollenites clarireticulatus (Samoilovitch) n. comb.

Plate 2, figures 1-8; plate 3, figures 1-7

Inteffricorpus clarireticulatus Samoilovitch, 1965, p. 123-124. 

Holotype: figs. 2a-b (p. 128) and pi. I, figs. 2a-d (p. 403) ; 

(Paleophytological Laboratory of the Scientific Research 

Institute of Petroleum :a,nd Geological Exploration 

(VNIGRI), preparation 10535a; Linde River, Yakutia, 

U.S.S.R.; top of Chirimyian suite, Maestrichtian-Danian.

The following is >a translation by Ivan Mittin of the 

description of Integricorpus clarireticulatus Samoilovitch, 1965, p. 123-124:

"Diagnosis. Pollen grains from rather large to large, 

isopolar or subisopolar, with three equatorial furrows 

and three meridianal furrows. Body ellipsoidal, large, 

with three equatorial protrusions. Protrusions inclined 

with respect to body, narrow, not long. Exine rather 

thin, clublike, reticulate, not sheathed.

"Description. Diameter of pollen grains in polar position 49.7/i. In equatorial position: polar axis 56.8/*, equatorial axis (with protrusions) 54.7/A, width of ends of 

body 25.3/A, length of equatorial protrusions 13.5/*, to 

18.7/t, width 5.5/t to 10.0/*.

"Pollen grains isopolar or subisopolar, with three 

meridianal and three equatorial furrows. Body large, 

ellipsoidal, one of its ends is slightly pointed, the other 

rounded. In polar position pollen triangular in outline 

with straight or slightly convex sides. Equatorial protrusions are situated 'at an angle to the 'body and form 

with the polar axis an angle about 30°, they are narrow, 

not. long, rounded in cross section, gradually narrowing toward ends, which may be found destroyed. Six 

furrows: three of them pass through the protrusions 

meridianally, the other three follow the equator between the protrusions; all furrows narrow, slitlike, with 

slightly rough margins. Exine is rather thin on the ends 

of the body (slightly larger than 1.0/x), it thickens to 

2.7/* to 3.5ft (nexine thickenings) in the areas between 

the ends of the body and protrusions. Layers poorly 

distinguishable. Traced iare: endonexine, very thin; ektonexine, becoming thicker in transition to protrusions 

and wedging out toward their ends and on the poles, 'and 

sexine of clublike structure, with heads of irregular 

form; width of the layer of heads does not exceed the 

width of the layer of columns. Sculpture highly clear, 

meshlike; walls of cells thin; cells very large on mesocolpium, sharply decrease in size and become elongate 

near equatorial furrows. Pollen dark yellow in color.

"Material. Four specimens well and medium preserved from three localities.

"Variability. Has not been traced because of insufficient amount of material.

"Comparison and Remarks. Pollen of the species 

under description resembles very little the pollen grains

of other species of subterm Triprojectacites It differs 

from pollen grains of Integricorpus helium N. Mtch., 

with which it is most comparable, in more angular form 

of body that narrows toward the ends, in thinner exine 

and in less coarse mesh appearance, as well as in the 

presence of a noticeable network in the equatorial zone 

which in /. ~beUwm is almost lacking the sculpture. It 

differs from Parviprojectus reticulatus N". Mtch. in 

larger size, in the reclining position of the protrusions, 

in the presence of equatorial furrows and considerable 

coarser and larger meshlike appearance of the exine.

"N". D. Mtchedlishvili (1961), who recojrnized the 

form genus Integricorpus. and to which we refer the 

species in the description, did not indicate tr<} presence 

of equatorial furrows in pollen grains.

"We came to the conclusion that they exist in the 

pollen of Integricorpus on the basis of our late findings 

of individual grain halves which were torn along the 

equator, which can happen only in the presence of 

highly thin exine (membrane of the furrowr) along it.

"The presence of equatorial furrows in pollen of some 

other representatives of the subterm Triprojectacites, 

such as in Aquilapollenites novacolpites Funk., is also 

noted by John Funkhouser (Funkhouser, 19*1).

"Localities. Yakutia, River Linde, exposure 1025, 

specimen 6, lignite; exposure 1028 specimen 2, clay; exposure 1039, specimen 4, clay. Collector V. V. Zabaluev. 

Maestrichtian-Danian (upper part Chirimyian suite).

"Distribution. Maestrichtian-Danian of Yukutia." 

Records of occurrence. 

Rock unit

USGS paleobota-7y 

Stage or series locality No. or 

literature reference

Locality

State 

and(or) 

country

Chirimyian SuiteEagle Sandstone, 

upper part.

Maestrichtian-Danian. Samoilovitch (1965'*... Yakutia,

U.S.S.B.

Lower Campanian.. - - D3718-I. -------------- Montana,

U.S.A.i

Upper Cretaceous_.. D1799.---------------- Alaska,

U.S.A.

1 For specific locality see table 1.

Description. Based on 40 specimens from USGS 

paleobotany localities D1799 and D3718-I. Tricolpate, 

isopolar to subisopolar pollen grains with three equatorial protrusions; polar axis 28/* to 44^, equatorial 

diameter (including equatorial protrusions) 29/* to 42/M. 

In equatorial view: body of grain oval, with broadly 

rounded poles, distended at equator where merging 

with V-shaped to U-shaped equatorial protrusions; 

polar protrusions much larger than equatorial protrusions; many specimens having equatorial protrusions 

bent or broken. In polar view: grains triangular, with 

irregularly concave-convex sides. Apertures: colpi 

reaching full length of equatorial protrusions, extend-
SPECIES OF AQUILAPOLLENITES AND FIBULAPOLLIS, ALASKA A7

ing onto body for approximately three-fourths the 

distance to poles; three furrows present on equator 

between equatorial protrusions. Pollen wall: exine two 

layered; endexine (exclusive of costae) thin; axillary 

endexinal costae present or lacking (probably dependent on chemical processing of sediment) ; costae when 

present bow shaped, as much as 2/i thick in thickest 

part, extending approximately half the length of the 

equatorial protrusions, reaching onto body for approximately three-fourths the distance to poles; ektexine 

on body 0.5ju, to Ip thick, with maximum thickness 

between polar areas and equator; ektexinal elements 

columnar, fused at tips forming round-topped muri; 

surface sculpture of body reticulate to striate; lumina 

of reticulum as much as 1.5/x in diameter in areas 

between poles and equator, usually slightly smaller on 

polar areas than between poles and equator; muri close 

together in equatorial zone, parallel to each other and 

arranged perpendicular to equatorial furrows resulting in an equatorial belt (about 15/u, wide) of differentiated sculpture; striations persisting beyond ends of 

equatorial furrows, fanning out onto equatorial protrusions (SEM photograph, pi. 2, fig. 8; pi. 3, fig. 7), 

striations discontinuous on protrusions at their approximate midlength; on outer ends of equatorial protrusions exine thin, wall layers usually not discernible, 

sculpture smooth or granular. As seen in SEM photographs, the reticulate-striate appearance on the polar 

protrusions results from the joining of the elongate, 

more or less parallel muri of high relief with the horizontal muri of lower relief (SEM photograph, pi. 2, 

fig. 8; pi. 3, fig. 1) ; at high magnification (approximately X 10,800) the elongate muri appear cordlike, 

many branch, and some interweave (SEM photograph, 

pi. 3, fig. 1).

Remarks. The specimens just described appear, 

except for their smaller size, very similar to "Integricorpus" clarireticulatux as it was described and illustrated by Samoilovitch. Difference in size could be the 

result of differences in chemical processing in the two 

laboratories. The Samoilovitch description was based 

on only four specimens; a larger population of the Russian material conceivably might have yielded a wider 

size range.

Because "I'ntegricorp-us" clarireticulatus conforms to 

the genus Aqutlapollemtes (Rouse) Fimkhouser, it has 

been transferred to Aquilapollenites.

Similar species. A. clarireticulatus n. comb, resembles A. reticulatus (Mtchedlishvili) Tschudy and 

Leopold (1969) but can easily be distinguished by its 

equatorial furrows and by the distinct bands of striate 

sculpture adjacent to the furrows at the equator.

A. clarireticulatus resembles A. novcwolpites Funkhouser (1961, p. 196) except that the equatorial protrusions of A. clarireticulatus are not constricted 

between their tips and the body of the grain as they 

are on A. novacolpites (as seen on its type specimens 

and on several additional specimens found on the type 

slide). The equatorial zone of striate sculpture which 

characterizes A. clarireticulatus is not present in the 

type specimens or in other specimens found on the type 

slide of A. novacolpites.

Aquilapollenites trialatus Rouse, 1957 

Plate 4, figures 1-5

Aqutlapollemtes trialatus Rouse, 1957, p. 371, pi. 2, figs. 14, 15. 

Type specimen: Bouse, 1957, pi. 2, fig. 14; McMaster 

University Palaeobotanical Collection; Slide Leth No. 8, 

1: 5 (ii) ; Lethbridge City Collieries, south-^ntral 

Alberta, Canada; Oldman Formation, Upper Cretaceous.

Aquilapollenites triala-tvs Rouse, 1957 (in Tschudy and Leopold.

1969). 

Records of occurrence. 

Rock unit

Locality USGS paleobotany 

Stage or series locality No. or State 

literature reference rnd(or) 

country

Oldman Formation_._ Upper Cretaceous...-- Rouse (1957)..-.-.-.__ Alberta,

Canada.

Colville Group...-__-._ Upper Cretaceous_.- D3124-H.............. Alaska,

U.S.A.

Claggett Shale, upper Middle Campanian_ D3724-E............._ M-mtma,

part. U.S.A. 1

1 For specific locality see table 1.

Remarks. Specimens of A. trialatus found in the 

Alaskan sample USGS paleobotany locality D3124-H 

closely resemble specimens found on a slide (Leth No. 

8, 1:5 (i) Ger) of type material loaned by Glenn E. 

Rouse to this laboratory.

In samples from the Rocky Mountain area the tf.uthor 

has found only a few specimens of A. trialatus; however, two new varieties of A. trialatus from Campanian 

sedimentary rocks of the Rocky Mountains are described by Tschudy and Leopold (1969).

Grains from the Upper Cretaceous (Lindian suite) 

from Yakutia, central Siberia, U.S.S.R., were identified as A. trialatus Rouse by Bolkhovitina (1959, p 127) 

and grains from the Senonian (lower Symiar subsuite) from a locality near the Sym River in the Western Siberian lowlands, which appear very similar or 

equal to A. trialatus, were described by Mtchedlishvilli 

(in Samoilovitch and others, 1961, p. 217-218) as Integricorpus bellum.

Aquilapollenites parallelus n. sp. 

Plate 5, figures 1-11; plate 6, figures 1-2; and plate 7, figures 1-2

Holotype. USGS paleobotany locality D1799, slide 

(5) at 100.7 X 12.8; plate 5, figure 1; lat 65°19.6' N., long
A8 CONTRIBUTIONS TO PALEONTOLOGY

141°29.6' W.; SE}£ sec. 23, T. 6 N., R. 30 E. (Charley 

River B-l quadrangle) ; east bank of Nation River, 

Alaska; Upper Cretaceous.

Paratype. USGS paleobotany locality D1799, slide 

(7) at 77.7X14.9; plate 5, figures 8, 11; same locality 

data as for holotype.

Paratype. USGS paleobotany locality D1799, slide 

(5) at 79.9X19.8; plate 5, figures 4, 6; same locality 

data as for holotype.

Occurrence. USGS paleobotany locality D1799.

Description. Based on 23 measured specimens and on 

many more examined in less detail. Tricolpate, isopolar 

to subisopolar pollen grains with three equatorial protrusions. In equatorial view: polar axis 33u to 59ju, (commonly 45/u to 55/w), equatorial diameter (including 

equatorial protrusions) 28/x, to 55/j (commonly 42/u to 

50/x), diameter of body 14/u, to 24^, length of equatorial 

protrusions 13/i to 21//, width of equatorial protrusions 

6.5ju. to 10.5,11; body having nearly parallel sides and 

flattened or broadly rounded ends; equatorial protrusions having parallel sides and rounded ends; width of 

equatorial protrusions usually about half the diameter 

of body. In polar view: grains triangular, with concave 

or irregular sides; poles of body triangular, with slightly convex sides (pi. 6, fig. 6). Apertures: colpi extending 

full length of equatorial protrusions, in some specimens 

gaping at ends of protrusions, apparently not extending onto body or, if so, for a very short distance; colpi 

usually visible only in oblique and polar views of grains. 

Pollen wall: exine two layered; endexine (except for 

endexinal costae) less than 0.5/u, thick; axillary endexinal 

costae as much as 2ju thick in thickest part, extending 

approximately two-thirds the distance to ends of equatorial protrusions, extending only slightly onto body of 

grain; ektexine 0.5/u to 2.0/u thick, forming ektexinal 

ridges (lirae) ridges give the grains an overall striate 

appearance; in SEM photographs (pis. 6, 7) surface of 

grains appear fluted (or plaited) ; lirae nearly smooth at 

X 1,000 magnification, at SEM high magnification 

(X 10,800) ridges have scattered wartlike projections 

(pi. 6, fig. 2); lirae oriented nearly perpendicular to 

colpal margins on equatorial protrusions; at ends of 

protrusions lirae endings form toothed colpal margins 

('SEM photographs, pi. 7, figs. 1-2); lirae on the body 

(in equatorial view) parallel to polar axis in the equatorial zone, irregularly arranged or in fingerprint pattern near poles, and parallel to circumference of body 

between poles and equator; in polar view of grains, triangular pole having lirae approximately perpendicular 

to sides of triangle and encircling corners (pi. 5, fig. 6).

Remarks. The grains described above have been 

given the species name parallelus because of the nearly

parallel sides of both the polar and the equatorial protrusions as seen in equatorial view of the grains.

Similar species. A. parallelus most closely resembles 

Aquilapollenites conatus Norton (1965, p. 142) but can 

be distinguished by its proportionally narrower equatorial protrusions and coarser lirae on the equatorial 

protrusions. The ektexinal ridges of A. parallelus do 

not appear to be connected by horizontal muri, whereas 

the ridges of A. conatus are at least in part connected 

by narrow horizontal muri (ladderlike); sometimes 

these muri give the surface of the grains a reticulate to 

striate appearance.

The parallel-sided body of A. parallelus distinguishes 

it from Aquilapollenites bertillonites Funkhouser 

(1961, p. 196) and Aquilapollenites wilfordi Muller 

(1968, p. 16-17).

Aquilapollenites fusiformis n. sp.

Plate 8, figures 1-12

Holotype. USGS paleobotany locality D3124-G, 

slide (6) at 111.9X18.1; plate 8, figures 1, 2; lat 69°40' 

N., long 151°25' W.; T. 4 N., E. 2 E. (Umiat quadrangle) ; along Colville River, Alaska; Colville Group, 

Upper Cretaceous.

Paratype. USGS paleobotany locality D3124-G, 

slide (6) at 109.1X10.2; plate 8, figure 3; same locality 

data as for holotype.

Occurrence. USGS paleobotany locality D3124-B, 

-G.

Description. Based on 22 specimens in equatorial 

view, 13 specimens in polar view, and three in oblique 

views. Tricolpate, isopolar to subisopolar pollen grains 

with three equatorial protrusions. In equatorial view: 

body rhomboidal to spindle shaped, poles narrowly 

rounded to rather pointed; polar axis 35^ to 55/*; equatorial diameter (including equatorial protrusions) 29/u 

to 52/i; equatorial protrusions V-shaped, 12/u to 16^ long 

(measured from free end of protrusion inward to onset 

of body sculpture). In polar view: shape of grains triangular, with plane to irregularly concave-convex sides 

and pointed or rounded corners. Apertures: coT pi slitlike 

or often gaping, extending full length of equatorial protrusions and onto body nearly to poles. Pollen wall: 

exine two layered; endexine on body (exclusive of costae) less than 0.5/* thick; axillary endexinal costae long, 

bow shaped, as much as 3/j thick in thickest part, tapering to very thin ends, extending nearly to poles and 

reaching nearly to ends of equatorial protrueions (costae often lacking, probably having been dissolved during chemical processing of grains); ektexine of body 

less than 1/u, or sometimes as much as 3/j thick; ektexinal 

elements elongate, very thin or as much as about 0.25/j 

wide, irregularly distributed or more or les^ parallel,
-SPECIES OF AQUILAPOLLENITES AND FIBULAPOLLIS, ALASKA A9

sometimes connected by horizontal muri; sculpture 

varying from rugulate to striate-reticulate, or in fingerprint pattern; ektexine sometimes thickened to as much 

as 3/j, at poles and elements more compact than on remainder of body; ektexine of equatorial protrusions less 

than 0.5/j. thick, elements compact, sculpture finely granulate to faintly striate.

Remarks. Except for variation in coarseness of 

ektexinal sculpture the described specimens are quite 

uniform in appearance. The spindlelike shape in equatorial view suggested the name fusiformis for this 

species.

Similar species. A. fusiformis, in common with 

Aquilapollenites trialatus Rouse (195Y, p. 371), Aquilapollenites reticulatus (Mtchedlishvili) Tschudy and 

Leopold (1969), and Aquilapollenites ~bertillonites 

Funkhouser (1961, p. 196) has finely ornamented 

V-shaped equatorial protrusions. The spindlelike shape 

and rugulate to striate-reticulate body sculpture of A. 

fusiformis distinguish it from the reticulate species 

A. trialatm and A. reticulatm; its surface sculpture 

easily distinguishes it from the distinctly striate species 

A. l>ertillonites.

Aquilapollenites dentatus n. sp. 

Plate 9, figures 1-12; plate 10

Holotype. USGrS paleobotany locality D1799, slide 

(Y) at 90.9X8.9; plate 9, figures 1, 4, Y; lat 65°19.6' N., 

long 141°29.6' W.; SEi/i sec, 23, T. 6 N., R. 30 R 

(Charley Eiver B-l quadrangle) ; east bank of Nation 

River, Alaska; Upper Cretaceous.

Paratype. USGS paleobotany locality D1Y99, slide 

(Y) at Y9.9 X Y.2; plate 9, figures 2, 5; same locality data 

as for holotype.

Occurrence. USGS paleobotany locality D1Y99.

Description. Based on over 30 measured specimens 

and on many more grains examined in less detail. Tricolpate, isopolar to subisopolar pollen grains with three 

equatorial protrusions. In equatorial view: polar axis 

29.5/z, to 55/x, (commonly 45/z, to 52/x,), equatorial diameter 

(including equatorial protrusions) 35ju, to 58/x, (commonly 45/x to 58/x), diameter of body 10.5ju to 21/x, diameter or equatorial protrusions Y.5/x to 14/*, length of 

equatorial protrusions 11/x, to 21/x,; shape of body oval; 

diameter of polar protrusions usually slightly greater 

than diameter of equatorial protrusions; lengths of polar 

and equatorial protrusions usually approximately 

equal; polar and equatorial protrusions broadly rounded 

at ends. In polar view: shape of grains triangular, with 

slightly concave sides and rounded corners; equatorial 

protrusions slightly flattened in plane parallel to polar 

axis, of grain. Apertures: colpi reaching total length of 

equatorial protrusions, extending a short distance onto

body, sometimes gaping at ends of equatorial protrusions. Pollen wall: exine two layered, endexine (except 

for endexinal costae) usually less than 0.5/i thick; axillary enclexinal costae bow shaped to lens shaped or irregular (some costae having a bulge or apparent thickening directly at the angle of the axillae), costae as 

much as 4.5/x thick in thickest part, extending one-half 

to two-thirds the length of equatorial protrusion'7 and 

onto the body for one-third to one-half the distance to 

poles; ektexine approximately I/*, thick; ektexinsl elements columnar, their diameter approximately equal to 

the distance between them, covered by a perforate tectum resulting in a foveolate surface sculpture; a row 

of conical spines, 2.5/z to 4/z, long, present along colpal 

margins at tips of equatorial protrusions and pointing 

toward polar axis (retroflexed); in SEM photograph 

(at X 4,200 magnification) the spines at colpal margins 

appear thornlike and somewhat flattened (pi. 10); 

smaller conical spines I/* to 2/x, long ornamenting surface of grain these mainly restricted to polar domes 

and an equatorial band where they extend onto equatorial protrusions for one-half to two-thirds the length 

of the protrusions; tips of equatorial protrusions devoid 

of spines except for the row of large spines at colpal 

margins; spines on polar protrusions standing perpendicular to surface or having a slight tendency to point 

poleward; along equatorial band the spines tend to 

point away from tips of equatorial protrusions.

Remarks. The new species was given the name 

dentatus (toothed, pointed) because of the conspicuous 

row of large, sharp-pointed spines present alorg the 

colpal margins at the tips of the equatorial protrusions.

Similar species. A. dentatus n. sp. resembles Aquilapollenites attenuatus Funkhouser (1961, p. 194,196) but 

can easily be distinguished because it has a single row 

of enlarged spines confined to the colpal margins at 

the ends of the equatorial protrusions, whereas A. attenuatus has spines scattered over the ends of the equatorial protrusions and also continuing along the colpal 

margins as far as, or nearly to the body; the spines at 

the tips of the equatorial protrusions in A. dentatus are 

conspicuously larger than the body spines whereas the 

spines on the equatorial protrusions of A. attenuatus 

are not noticeably larger than the body spines. A. dentatus tends to be smaller than A. attenuatus but size ranges 

do overlap. A. dentatus usually has thicker axillary endexinal thickenings than A. attenuatus.

Aquilapollenites colvillensis n. sp. 

Plate 11, figures 1-18

Holotype. USGS paleobotany locality D3324-G, 

slide (5) at 109.5X21.8; plate 11, figures 1, 2, 3; lat 

69°40' N., long 151°25' W.; T. 4 N., R. 2 E. (Umiat
A10 .CONTRIBUTIONS TO PALEONTOLOGY

quadrangle); along Colville River, Alaska; Colville 

Group, Upper Cretaceous.

Paratype. USGS paleobotany locality D3124-G, 

slide (16) at 83.9X10.4; plate 11, figures 4, 8; same 

locality data as for holotype.

Paratype. USGS paleobotany locality D3124-G, 

slide (6) at 110.1X7.6; plate 11, figures 5, 6; same 

locality data as for holotype.

Occurrence. USGS paleobotany locality D1324-B, 

-G, -H.

Description. Based on the examination of more 

than 60 specimens in equatorial, polar, and oblique 

views. Tricolpate, isopolar to subisopolar pollen grains 

with three equatorial protusions. In equatorial view: 

body of grain having nearly parallel sides and broadly 

rounded poles; polar axis approximately two times the 

body diameter; equatorial protrusions having nearly 

parallel sides and broadly rounded ends, extending at 

right angles from body, usually shorter and narrower 

than polar protrusions; polar axis 27/t to 45/t, diameter 

of body 12.5/t to 20/t, equatorial diameter (including 

equatorial protrusions) 28/x to 40/t, length of equatorial 

protusions 8/t to 12/t, width of equatorial protusions 

10/4 to 14/x. In polar view: grains triangular; sides 

slightly concave, slightly convex, or irregular; colpi at 

corners, in some specimens gaping. Apertures: colpi 

reaching full length of equatorial protrusions and onto 

body of grain for approximately one-third the distance 

to poles. Pollen wall: exine two layered; endexine (exclusive of costae) thin; axillary endexinal costae lens 

shaped to sausage shaped (pi. 11, fig. 11), as much as 4/t 

thick in thickest part, occupying approximately half 

the length of equatorial protrusions and extending onto 

body for about one-fourth the distance to poles; costae, 

as viewed from their ends, rounded poleward and flattened equatorially (pi. 11, figs. 16, 17) ; costae usually 

lacking in grains which have had long Schulze treatment during removal from rock matrix; * ektexine 

(except at colpal margins) about 0.5/A thick; ektexinal 

elements columnar, compact, joined at tips resulting in 

a finely foveolate-reticulate to nearly smooth appearance to surface of grain; ektexine at colpal margins very 

thin; broad-based projections, about 0.5/t long, spaced 

at 1.5/A to 2/t intervals, ornamenting body of grain; sharp 

pointed conical spinae (1/t to 1.5/t long), spaced at 1.5^ 

to 2/4 intervals, ornamenting equatorial protrusions, 

tending to point backward (retroflexed) from ends of 

protrusions and toward body of grain.

Similar species. A. colvillensis resembles A. laticorpus n. sp. (p. A12) and A. rectus n. sp. (p. A10)

1 Preliminary experiments carried ont in this laboratory on this 

species indicate that endexinal costae are removed by long Schulze 

treatment.

but is easily distinguishable from the latter two 

species. A. colvillensis has coarser surface omamentation than A. laticorpus and it has a nearly parallelsided body whereas A. laticorpus has a broadly oval 

body. The equatorial protrusions of A. colvillensis are 

nearly always considerably smaller than the polar protrusions whereas the equatorial protrusions of A. laticorpus and A. rectus are usually approximately the 

same size as the polar protrusions. A. colvillensis 

tends to have a polar axis greater than its equatorial 

diameter whereas A. rectus tends to have a polar axis 

shorter than its equatorial diameter.

The shape of A. colvillensis is similar to that of 

Aquilapollenites spinulosus Funkhouser (1961, p. 194), 

but the former has coarser surface ornamentation. 

A. spinulosus has fine, slender spines over its entire 

surface, whereas A. colvillensis has short, broad-based 

projections on its body and longer, conical spines on its 

equatorial protrusions.

The nearly parallel-sided body of A. colvillensis 

resembles that of "Projectoporites" ovalis Mtchedlishvili (in Samoilovitch and others, 1961, p. 228-229); 

however, the two species differ in that A. colvillensis 

is much smaller than "P." ovalis (polar axis 47.5/* to 

50.8/t, diameter of body 23.7/1. to 31.2/*) and the body 

ornamentation of A. colvillensis consists of short, 

broad-based projections whereas that of "P." ovalis 

consists of slender, longer, sharp-pointed spines. "P." 

ovalis was described as having pores rather than colpi; 

however, inasmuch as only four specimen? were examined this feature may not have been certain.

Aquilapollenites rectus n. sp. 

Plate 12, figures 1-13; plate 13, figures 4. 9

Holotype. USGS paleobotany locality D1799, slide 

(4) at 113.3 X 18.1; plate 12, figure 1; lat 65° 19.6' N., 

long 141° 29.6' W.; SE% sec. 23, T. 6 N., R. 30 E. 

(Charley River B-l quadrangle); east bank of Nation 

River, Alaska; Upper Cretaceous.

Paratype. USGS paleobotany locality D1799, slide 

(4) at 98.4X11.7; plate 12, figure 2; same lo^lity data 

as for holotype.

Occurrence. USGS paleobotany locality D1799.

Description. Based on 23 specimens in equatorial 

and oblique views and six in polar view. Tricolpate, 

isopolar to subisopolar pollen grains with three equatorial protrusions; polar axis 30/t to 37/t; equatorial 

diameter (including equatorial protrusions) 33/t to 45/t. 

In equatorial view: body of grain broadly oval; polar 

axis slightly shorter than or about equal to equatorial 

diameter; polar protrusions equal to or slightly larger 

or smaller than equatorial protrusions; polar and equatorial protrusions broadly rounded at their ends; areas
SPECIES OF AQUILAPOLLENITES AND FIBULAPOLLIS, ALASKA All

of attachment of equatorial protrusions to body (the 

axillae) concave. In polar view: grains triangular, with 

rounded corners and plane or irregularly concave-convex sides. Apertures: colpi extending full length of 

equatorial protrusions, apparently not extending onto 

body, or, if they do, for only a very short distance. 

Pollen wall: exine two layered; endexine thin (except 

for endexinal costae), very thin at colpal margins at 

ends of equatorial protrusions; axillary endexinal costae 

lens shaped (in equatorial view of grain), 1.5/* to 2.5ju, 

thick in thickest part, extending one-third to one-half 

the length of the equatorial protrusions, extending a 

short distance onto base of polar protrusions; ektexine 

very thin at colpal margins at ends of equatorial protrusions, 0.5/A to 1.0/A thick on remainder of grain, having maximum thickness on approximate basal half of 

polar protrusions; ektexinal elements columnar, joined 

at tips giving a finely reticulate appearance to surface of grain; reticuhmi slightly coarser on basal part 

of polar protrusions than on remainder of grain; conical 

spines and baculae 1.0/tt to 1.5/x long, distributed over 

entire surface of grain, on equatorial protrusions tending to point toward polar axis of grain.

Remarks. The species was given the name rectus 

(straight) because of its rather stiff or rigid 

appearance.

Similar species. A. rectus n. sp. resembles A. laticorpus n. sp. (this report, p. A12) but differs from it 

in having larger surface projections (spines and 

baculae); A. rectus also has a more concave curvature 

in the axillae than A. latworpus. The equatorial protrusions of A. rectus are more distinctly prominent 

than they are in A. latworpus. When the grains here 

described as A. rectus were found in sample D1T99 

they were believed to be the species A. laticorpus n. sp. 

which was earlier found in sample D3124; however, 

further study of many specimens from the two samples 

showed the two species to be distinct.

^4. rectus, n. sp. resembles A. colvillensis n. sp. (this 

report, p. A9), but it has equatorial protrusions 

which are larger in proportion to the polar protrusions. 

A. rectus has surface projections of nearly uniform size 

distributed over the entire grain, whereas A. colviilensls 

has distinctly larger surface projections on the equatorial protrusions than on the polar protrusions.

In size of grain and type of surface ornamentation, 

A. rectus n. sp. resembles Aquilapollenites spinulos-m 

Funkhouser (1961, p. 194) but differs because it has 

proportionally shorter polar protrusions and proportionally wider equatorial protrusions (in equatorial 

view of grain). A. spinulosm usually has a longer 

polar axis than equatorial diameter and the reverse 

relationship is usually true for specimens of A. rectus.

Aquilapollenites scabridus n. sp. 

Plate 13, figures 1-11; plate 14, figures 1-2

Holotype. USGS paleobotany locality D1799, slide 

(5) at 86.2X11.8; plat© 13, figures 1, 2; lat 65°19.6' N"., 

long 141°29.6' W.; SE^ sec. 23, T. 6 N., R. 30 E. 

(Charley Eiver B-l quadrangle); east bank of Nation 

Eiver, Alaska; Upper Cretaceous.

Paratype. USGS paleobotany locality D1799, slide 

(T) at 90.3X12.0; plate 13, figure 3; same locality data 

as for holotype.

Occurrence. USGS paleobotany locality D1799.

Description. Based on 18 specimens in equatorial 

and oblique views and two specimens in polar view. Tricolpate, isopolar to subisopolar pollen grains with three 

equatorial protrusions. In equatorial view: polar axis 

32ft, to 43/A, equatorial diameter (including equatorial 

protrusions) 32/* to 45ju,; polar axis usually approximately equal to equatorial diameter, polar protrusions 

smaller than, or in some specimens about equal in size 

to, equatorial protrusions; poles broadly rounded, or in 

many specimens flattened; equatorial protrusions with 

broadly rounded ends; attachment areas (axilla0-) of 

equatorial protrusions to body concave. In polar view: 

shape triangular, with plane or irregular cor cavecon vex sides. Apertures: colpi extending the length of 

the equatorial protrusions, at least in some specimens 

extending onto body for approximately one-fourth the 

distance to poles. Pollen wall: exine two layered; endexine less than 0.5/t thick (except for endexinal cortae); 

axillary endexinal costae (in specimens where present) 

bow shaped to sausage shaped (as seen in equatorial 

view of grain), as much as 3ju, thick in thickest part, 

occupying one-third to one-half the length of the equatorial protrusions, extending a short distance onto body 

of grain; costae lacking in some specimens, represented 

only by thin areas from which they were proba^y removed during chemical processing of sample; ektexine 

lp to 1.5/A thick, with maximum thickness on basal part 

of polar protrusions, ektexine very thin at colpal margins at ends of equatorial protrusions; ektexinal elements columnar, tectate; columellae usually visiMe ait 

X 1,000 magnification on polar protrusions of well-preserved grains; polar domes almost smooth, remainder of 

surface of grain densely scabrate; on SEM photograph 

(pi. 14, fig. 2) at X 10,400 magnification, the sculpt-iring 

elements on the equatorial zone and on an equatorial 

protrusion appear approximately twice as high as wide 

and their peaks are rounded to bluntly conical; at 

X 4,200 magnification the sculpturing elements c^.n be 

seen to be more compactly arranged on the polar Monies 

than on the remainder of the grain. (See pi. 14, fig1. 1.)

R&ma/rks. The species was given the name scabridus 

because of its rough surface sculpture.
A12 CONTRIBUTIONS TO PALEONTOLOGY

Similar species. In shape and size A. scabridus is 

very similar to A. rectus n. sp. (this report, p. A10), 

but. it lacks the spinose ornamentation which characterizes the latter species.

The polar domes of A. scabridus^ in common with 

those of A. procerus Samoilovitch (1965, p. 126-128), 

are smooth or nearly so; however, the sculpture of the 

polar domes of A. scabridus is not as sharply distinct 

from that of the remainder of the grain as it is in A. 

procerus. A. scabndus has proportionally shorter 

polar protrusions and broader equatorial protrusions 

(in equatorial view of grains) than has A. proceru.s.

In shape of grain and nearly smooth polar domes, 

A. scabrid'its resembles A. twrbidus Tschudy and Leopold (1969) ; but it differs because it does not have 

the spinose ornamentation in the equatorial zone of the 

body and on the equatorial protrusions.

Aquilapollenites contiguus n. sp.

Plate 15, figures 1-15

Holotype. USGS plaeobotany locality D3124-G, 

slide (5) at 88.7X5.7; plate 15, figures 1, 2; lat 69°40' 

N., long 151°25' W.; T. 4 N., R. 2 E. (Umiat quadrangle) ; along Colville River, Alaska; Colville Group, 

Upper Cretaceous.

Paratype. USGS paleobotany locality D3124-G, 

slide (6) at 100.9X3.5; plate 15, figure 3; same locality 

data as for holotype.

Paratype. USGrS paleobotany locality D3124-G, 

slide (5) at 102.7 X 10.4; plate 15, figure 4; same locality 

data as for holotype.

Occurrence. USGS paleobotany locality D3124-B, 

-G.

Description. Based on 20 specimens in equatorial 

and oblique views and seven in polar view. Tricolpate, 

isopolar to subisopolar pollen grains. In equatorial 

view: polar protrusions shorter than equatorial protrusions and more broadly rounded at ends, area of 

attachment of equatorial protrusions to body slightly 

concave, polar axis usually less than equatorial diameter; polar axis 22.5/i to 28/*, equatorial diameter 

30.5/x to 44/x,. In polar view: shape triangular, with 

plane, slightly convex, slightly concave, or irregular 

sides; most specimens with earlike corners these bearing slitlike or gaping colpi. Apertures: colpi short, dissecting ends of equatorial protrusions, not extending 

onto body of grain, usually not visible in equatorial 

view of grains. Pollen wall: exine two layered, I/* to 1.5/x 

thick, thinner at colpal margins; endexine (except for 

endexinal costae) thin; axillary endexinal costae in 

specimens where present as much as 2.5/^ wide at 

thickest part, usually approximately three times longer 

than wide (in equatorial view of grain), teardrop to

lens shaped; costae sometimes lacking, represented only 

by costal cavities or thin areas from which costae were 

probably removed during chemical processing of 

sample; ektexine two to three times thicke1* than endexine, columnar, apparently tectate; columellae extremely fine, often difficult to distinguish even at 

X 1,000 magnification; surface sculpture granulate to 

scabrate.

Remarks. The species was given the name contiguus 

(touching, bordering) because it just barely conforms 

to the circumscription of the genus Aquilapollenites 

A. contiguus having only a slight concavity (in equatorial view of grain) in the axillae where the equatorial 

protrusions join the body of the grain, in contrast to 

most species of Aquilapollenites which have riore deeply 

concave axillary areas.

Similar species. A. contiguus resembles Fibula-pollis 

saxbratus n. sp. (this report p. A13) but is distinguishable because of the presence of the slight concavity in 

the axillae where the equatorial protrusions join the 

body of the grain. The endexinal thickenings of A. 

contiguus are usually more elongate than those of F. 

scdbratus. The latter species usually has an equatorial 

diameter approximately equal to the polar axis whereas 

A. contiguus nearly always has an equatorial diameter 

considerably greater than its poplar axis.

Aquilapollenites laticorpus n. sp. 

Plate 16, figures 1-8

Holotype. USGS paleobotany locality D3124-G, 

slide (15) at 90.1X8.0; plate 16, figure 1; lat 69°40/ N., 

long 151°25' W.; T. 4 N., R. 2 E. (Umiat quadrangle) ; 

along Colville River, Alaska; Colville Group, Upper 

Cretaceous.

Pamtype. USGS paleobotany locality D3124-G, 

slide (18) at 116.8X6.9; plate 16, figures 2, 3; same 

locality data as for holotype.

Occurrence. USGS paleobotany locality D3124-B, 

-G.

Description. Based on 20 specimens in equatorial and 

oblique views and two in polar view. Tricolpate, isopolar to subisopolar pollen grains with three equatorial 

protrusions. In equatorial view: polar axis 28/i to 43/x, 

equatorial diameter (including equatorial protrusions) 

28/x to 43/i; polar axis usually approximately equal to 

equatorial diameter; polar protrusions equal to, or 

slightly larger or smaller than equatorial protrusions; 

polar and equatorial protrusions broadly rounded at 

ends; areas of attachment of equatorial protrusions to 

body (the axillae) broadly concave. In polar view: 

grains triangular, with slightly convex, slightly concave, or irregular sides. Apertures: colpi extending full 

length of equatorial protrusions, reaching a short dis-
SPECIES OF AQUILAPOLLENITES AND FIBULAPOLLIS, ALASKA A13

tance onto body. Pollen wall: exine two layered; endexine thin (except for endexinal costae); axillary 

endexinal costae in specimens where present lens shaped 

to sausage shaped (as seen in equatorial view of grains), 

occupying approximately half the length of the equatorial protrusions and extending a short distance onto 

body of grain; costae lacking in some specimens, represented only by thin areas from which costae were probably dissolved during chemical processing of sample; 

ektexine 0.5/A to I/A thick, with maximum thickness on 

basal part of polar protrusions, thin at colpal margins 

on ends of equatorial protrusions; ektexinal elements 

columnar, joined at tips resulting in a finely reticulate 

appearance to surface of grain; reticulum usually 

slightly coarser in a band around approximate basal 

half of polar protrusions; small baculae (or spinae) 

less than I/A long, sparsely distributed over surface of 

grain.

Remarks. The species was given the name laticorpus 

(latus, broad; corpus, body) because of its equatorially 

broadened body.

Five specimens resembling A. laticorpus but differing 

in that they have distinctly larger surface projections 

on the equatorial protrusions were found in USGS 

paleobotany locality D3124-G (pi. 16, figs. 9,10); these, 

specimens have not been named and described here 

owing to insufficient diagnostic material.

Similar species. A. laticorpus n. sp. has slight resemblance to A. colvillensis n. sp. (p. A9, this report,) 

and to Aquilapollentes spinulosus Funkhouser (1961, 

p. 194), but it differs from both in having smaller surface projections and a more broadly oval body. In 

equatorial views the bodies of A. colvillensis and 

A. spimilosus usually have more or less parallel sides.

A. laticorpu-s has smaller surface projections than 

A. rectus n. sp. (p. AlO, this report) and in equatorial 

views of grains the equatorial protrusions of A. laticorpus merge more gradually with the body than do 

those of A. rectus.

Form genus Fibulapollis Chlonova, 1961

Fibulapollis Chlonova, 1961, p. 87.

The following description from Chlonova (1961, p. 

87) is a translation by Ivan Mittin:

'"Description. Outline of the pollen body is more 

or less rounded or rounded-triangular, more rarely four 

angular. There are three pores or more rarely four. Pore 

apertures have a square form (rarely rounded). Exine 

is two-layered, I/A to 1.5/i thick, it is thicker at various 

degrees at the edges or margins of the aperturelike zone. 

Grains are smooth or carry fine dotted banded sculpture, 

relatively small (not large), 15ju. to 48/*. The color is 

light yellow.

351-255 O 69 ,3

"Type species of genus Fibulapollis mirificus 

(Chlonova) comb, nov." See Triporina mirifica Chlonova (1957, p. 44, pi. 1, figs. 4-6).

Description (morphological interpretation of present 

author). Tricolpate, isopolar to subisopolar (or heteropolar?) pollen grains with three (or rarely four 2 ) 

equatorial expansions in apertural areas; equatorial expansions not appearing as distinct equatorial protrusions but merging gradually with body of grain. Shape 

of grains in equatorial view oval to rounded rhomboidal, 

shape in polar view triangular. Colpi slitlike, cutting 

equatorial expansions in plane parallel to polar aris of 

grain. Exine at least two-layered; prominent variously 

shaped endexinal thickenings present at or near ends 

of colpi. Endexinal thickenings lacking in some specimens; probably dissolved during chemical processing 

of sample. Sculpture smooth or finely ornamented.

Remarks. In shape and size Fibulapollis resembles 

Cranwellia Srivastava (1966, p. 537) but is easily distinguishable becaues of its nonstriate surface sculpture.

The oval to rounded rhomboidal shape of Fibulapollis 

in equatorial view separates it from isopolar (or subisopolar) species of Aquilapollenites which in equatorial view have concave contact areas where the equatorial protrusions join the body.

Fibulapollis scabratus n. sp. 

Plate 16, figure 11-20

Holotype. USGS paleobotany locality D3124-A, 

slide (1) at 97.8X7.3; plate 16, figure 11; lat 69°40/ N., 

long 151 °25' W.; T. 4 N., K. 2 E. (Umiat quadrangle); 

along Colville Kiver, Alaska; Colville Group, Upper 

Cretaceous.

Paratype. USGS paleobotany locality D3124-A, 

slide (1) at 103.7X20.2; plate 16, figure 12; same locality data as for holotype.

Pamtype. USGS paleobotany locality D3124-A, 

slide (1) at 96.8 X 20.2; plate 16, figure 13; same locality 

data as for holotype.

Records of occurrence. 

Rock unit Stage or series USGS paleobotany 

locality No.

Lo-jality >

State and 

country

U.8.A. 

Colville Group......... Upper Cretaceous..... D3124-A,-B,-C,-G._ Aliska.

(?)-..-.._--._...-._.._...do. -...-----... D1799................. Do.

Fox Hills Sandstone...- Lower Maestrichtian.. D1331-5..-....--__.--_ Wyoming.

Pierre Shale, upper

unnamed shale

member. 

Pierre Shale, lower

unnamed shale

member. 

Pierre Shale, Mitten

Black Shale member. 

Claggett Shale, middle

part.

Upper Campanian_.._ D1330-26.

-do....---..------.- D1330-19.

Do.

Do.

Middle or lower D1330-4...,__..._-.-_- Do.

Campanian. 

Lower Campanian.... D3724-B.............. Montana.

For specific Rocky Mountain localities see table 1.

2 Chlonova (1961, p. 87). Grains having four equatorial expansions 

wpre not found by the present author.
A14 CONTRIBUTIONS TO PALEONTOLOGY

Description. Based on more than 40 specimens. Tricolpate, isopolartosubisopolar (orheteropolar?) pollen 

grains. In equatorial view: grains oval to rounded 

rhomboidal; polar areas broadly rounded; apertural 

areas distended equatorially but not forming equatorial 

protrusions distinct from body, protruded areas merging gradually onto body of grain not forming angles; 

polar axis 23/j, to 31.5/t, equatorial diameter 25.5^ to 

38ju,; polar axis commonly about equal to equatorial 

diameter, or sometimes slightly less or more than equatorial diameter. In polar view: grains triangular, with 

straight, slightly convex, slightly concave, or irregular 

sides. Apertures: colpi slitlike, present on broadly 

rounded ends of equatorial expansions, in plane parallel 

to polar axis of grain, reaching approximately one-third 

the distance to poles. Pollen wall: exine two layered, 

O.Sju, to lju, thick, thinner at colpal margins; endexine 

(exclusive of costae) thin; endexinal costae lens shaped, 

teardrop shaped or nearly round, as much as 2ju, thick 

in thickest part, usually not much longer than wide, 

located at colpal ends; endexinal costae lacking in some 

specimens but thin areas are present (pi. 16, figs. 15,17) 

and probably indicate positions from which costae were 

dissolved during chemical processing of rock sample; 

ektexine two to three times thicker than endexine, 

columnar, tectate; columellae extremely small, usually 

perceptible at X 1,000 magnification; surface sculpture 

scabrate.

Remarks. Because the specimens described above appeared to conform to the genus Fibulapollis and possibly were equivalent to FibulapoHis mirificm (Chlonova) Chlonova (1961, p. 87) or Fibulapollis pwnctatus 

Chlonova (1961, p. 87), some of the Alaskan material 

was sent to A. F. Chlonova for examination. After a 

study of the Alaskan specimens Dr. Chlonova (written 

commun., Nov. 3, 1964) replied that in size and shape 

they are close to F. jnirificus, but that the exine of F. 

mirificus is thicker and is smooth or slightly granular. 

She also mentioned that the average size of pollen of 

F. mirificus is smaller than that of the Alaskan pollen 

though she believed that this size difference might be 

accounted for by differences in laboratory methods. Dr. 

Chlonova also compared the Alaskan specimens with F. 

punctatus,' she stated that F. punctatus is larger than 

the Alaskan grains but that the ornamentation of the 

two is quite corparable. She said that there were other 

distinguishing features such as form and size of the 

apertures and thickness of exine in the aperturelike 

zone. She concluded that our specimens probably came 

from closely related plants.

REFERENCES CITED

Bolkhovitina, N. A., 1959, Sporovo-pyl'tsevye kompleksy mezozoishikh otlozlienii Viliuiskoi vpadiny i ikh znachenie dlia 

stratigrafii. [Mesozoic sporopollen complexes ir the Vilioui 

Basin and their stratigraphic significance] : Akad. Nauk 

SSSR Geol. Inst. Trudy, v. 24,185 p., 8 pis.

Chlonova, A. F., 1957, O vydelenii rukovodiashchiljh vidov pri 

opredelenii vozrasta otlozhenii po sporovo-pyt'tsevomv 

analizu [On identification of the index species in the determination of the age of deposits by spore-pollen analysis] : 

Akad. Nauk SSSR Vostoeh. Fil. Izv., no. 2, p. 43-46, 1 pi.

 1961, Spory i phl'tsa verkhnei poloviny verkhnego mela 

vostochnoi chasti Zapadno-Sibirskoi nizmennosti [Spores 

and pollen from the upper half of the Upper G etaceous of 

the eastern part of the western Siberian depression] : Akad. 

Nauk 'SSS'R Sibirskoye Otdeleniye, Inst. Geologii i Geofiziki 

Trudy, v. 7,140 p., 17 pis.

Drew, O. M., and Tschudy, B. D., 1968, Aquilapoltenites: Fossil 

pollen as seen under the scanning electron microscope: Geol. 

Soc. America Bull., v. 79, no. 12, p. 1829-1832, 2 pis.

Funkhouser, J.W., 1961, Pollen of the genus Aquilapollenites: 

Micropaleontology, v. 7, no. 2, p. 193-198,2 pis.

Muller, Jan, 1968, Palynology of the Pedawan and Plateau Sandstone Formations (Cretaceous-Eocene) in Sarawak, Malaysia : Micropaleontology, v. 14, no. 1, p. 1-37, 5 pi?.

Norton, N. J., 1965, Three, new species of Aquilapollenites from 

the Hell 'Greek Formation, Garfield County, Montana (!) : 

Pollen et Spores, v. 7, no. 1, p. 135-143, 4 pis.

Raclforth, N. W., and Rouse, G. B., 1954, The classification of 

recently discovered Cretaceous plant microfossils of potential importance to the stratigraphy of western Canadian 

coals [Alberta] : Canadian Jour. Botany, v. "2, no. 1, p. 

187-201,1 pi.

Rouse, G. E., 1957, The application of a new nomerNatural approach to Upper Cretaceous plant microfossils from western 

Canada: Canadian Jour. Botany, v. 35, no. 3, p. 349-375, 

3 pis.

Samoilovitch, S. R., 1965, Pyl'tsa novykh vidov verkl nemelovykh 

pokrytosemennykh rastenii lAkutii [Pollen of new species 

of Upper Cretaceous angiosperm plants of Yakrtia], p. 121- 

141, 402-411, in. Samoilovitch, S. R., ed., Paleofi^ologicheskii 

sbornik: Vses. Neft. Nauchno-Issled. Geol.-Razved. Inst. 

Trudy, v. 239, 507 p., 113 pis.

Samoilovitch, S. R., and others, 1961, Pyl'tsa i spory zapadnoi 

Sibiri, Uira-Paleotsen [Pollen and spores of western Siberia, 

Jurassic-Paleocene]: Yses. Neft. Nauchno-Issled. Geol.- 

Razved. Inst. Trudy, v. 177, 352 p., 149 pis.

Simpson, J. B., 1961, The Tertiary pollen-flora of Mull and 

Ardnamurchan: Royal Soc. Edinburgh Trans., v. 64, no. 16, 

p. 421-464, 20 pis.

Srivastava, S. K., 1966, Upper Cretaceous microflora (Maestrichtian) from Scollard, Alberta, Canada : Pollen et Spores, v. 8, 

no. 3, p. 497-552,11 pis.

Srivastava. S. K., 1968a, Eight species of Mancieorpns from the 

Edmonton Formation (Maestrichtian). Alberta, Canada: 

Canadian Jour. Botany, v. 46, no. 12, p. 1485-1490, 2 pis.

Srivastava, S. K., 1968b, Reticulate species of Aqi'ilapollenites 

and emendation of genus Mancicorpus Mtchedlishvili: Pollen et Spores, v. 10, no. 3, p. 665-699.

Stanley, E. A., 1961, The fossil pollen genus Aqitilapoltenitcs: 

Pollen et Spores, v. 3, no. 2, p. 329-352, 8 pis.

Traverse, A. F., Jr., 1958, Locating plant microfossils on mixed 

slides: Micropaleontology, v. 4, no. 2. p. 207-208.
S'PEGIES OF AQUILAPOLLENITES AND FIBULAPOLLIS, ALASKA A15

Tsehudy, B. D., and Leopold, E. B., 1969, Aquilapollenites Tschudy, R. H., 1966, Associated megaspores and microspores of 

(Rouse) Funkhouser Selected Rocky Mountain taxa and the Cretaceous genus Ariadnaesporites Potonie, 1956, 

their stratigraphic ranges in Symposium on Palynology of emend., in Geological Survey research 1966: U.S. Geo1 . Surthe Late Cretaceous and Early Tertiary : Geol. Soc. America vey Prof. Paper 550-D, p. D76-D82. 

Spec. Paper 127. (In press.)

INDEX

Page 

Aquila polkn ites............................... A3

attenuatus................................ it

bertillon ites............................... 8,9

elarireticulatus.-.-................. 3, 6; pis. 2, 3

colvillensis .. __________________ 3,9, 11, 13; pi. 11

conatus ___________________________________ 8

contiguus........................... 3, ;#; pi. 15

dedicate._________________________________ 5

dentatus.......................... 3, 9; pis. 9,10

fusiformis............................. 3, 8; pi. 8

laticorpus.. __________________ 3, 10, 11, W; pi. 16

« ovacolpites............................... 6,7

parattelus... ____________________ 3; pis. 5, 6, 7

procerus.................................. 12

quadrilobus............................... 3,5

rectus.. _____________________ 3,10,13; pis. 12, 13

reHcufa.Ms_______________________________ 7,9

scabridus.. _____________________ 3, 11; pis. 13,14

se7-£m.cMS-__-----_---_-_--____________ 3, 4; pi. 1

spinulosus.. ______________________________ 10,13

trialatus--.......................... 3,7, 9; pi. 4

turbidm.................................. 12

wilfordi. -.-____-__--____-___________._.._. 8

attenuatus, Aquilapollmites................... 9

helium, Integricorpus.......................... 6,7

bertillon ites, Aquilapollen ites.................. 8,9

Bibliography. __________ ______________________ 14

Botryococcus.................................. 3

darireticulatus, Aquilapollenites.-...... 3, 6; pis. 2,3

Integricorpus......................... 6, 7; pi. 2

colvillensis, Aquilapollenites------- 3, 9, 11, 13; pi. 11

[Italic page numbers indicate descriptions]

Page 

conatus, Aquilapollenites...................... A8

contiguus, Aquilapollenites..,......-.-.. 3, IS; pi. 15

Granwellia.................................... 3,13

Dacrydium................................... 3

defiea.us, Aquilapollenites..................... 5

dentatus, Aquilapollenites............. -3, 9; pis. 9, 10

Erdtmanipottis............................... 3

Fibulapollis mirificus ......................... 13, 14

punctatus ... ... ........................... 14

scabratus... ..................... 3, 12, /S; pi. 16

fusiformis, Aquilapollenites. ............... 3, 8; pi. 8

Inaperturopollenites ........................... 3

Integricorpus _-_____________--_-___.___-.--___ 4

bettum. -.--.-- ... ......................... 6,7

clarireticulatus. ....................... 6, 7; pi. 2

laticorpus, AquilapoUeniUs . 

Loranthacites _.____________

3, 10, 11, 7^; pi. 16 

__-__.___.... 3

Mancicorpus senonicum,- ...................... 4

senonicus ._..._..........._....-._..--,... 4

trapeziforme ... ___________._..______-_-.--_ 4, 5

mirifica, Triporina.. .......................... 13

mirificus, Fibulapollis . ---.-........-----.-.... 13, 14

novacolpites, Aquilapollenites. ----------------- 6, 7

ovalis, Projectoporites ...... .................... 10

Page 

s, Aquilapollenites.. ....... .-.A3; pis. 5, 6, 7

Parviprojectus. ............................... 4

reticulatus. ............................... 6

procerws, 4gw.ZapoZfen8es -...___._-_-....____. 12

Projectopor ites. ............................... 4

ovalis. .-.......--.----.-----.....-------.- 10

punctatus, Fibulapollis ... ..................... 14

quadrilobus, Aquilapollenites. ................. 3,5

reticulatus, Aquilapollenites. .................. 7,9

Parviprojectus. ... .......... ... . ... ....... 6

rectus, Aquilapollenites.. ........ 3, 10, 13; pis. 12, 13

scabratus, Fibulapollis. .............. 3, 12, IS; pi. 16

scabridus, Aquilapolknites.... .... .. 3, //; pis. 13, 14

senonicum, Manicorpus . . . .................... 4

senonicus, Aquilapolknites. ............... 3, 4; pi. 1

Mancicorpus....- ....... ... ............... 4

Sequoia.. .............. ....................... 3

spinulosus, Aquilapolknites. -...__..-....__.__ 10, 13

Taurocephalus . . .-............-.----.......... 4

trapeziforme, Mancicorpus. .................... 4, 5

trialatus, Aquilapollenites- ... ........... 3, 7, 9; pi. 4

Tricerapollis .................................. 4

Triporina mirifica ... .......................... 13

Triprojectacites- ----------------------------- 4

Triprojectus...... ................ .......... 4

turbidus, Aquilapollenites -____-___--,_.__-____ 12

wilfordi, Aquilapollenites. 8

A17

PLATES 1-16
PLATE 1

FIGURES 1-14. Aquilapollenites senonicus (Mtchedlishvili) Tschudy and Leopold, 1969 

(p. A4).

1. Equatorial view. USGS paleobotany loc. D3124-G, slide (5) 

at 112.6X3.1.

2. Equatorial view. USGS paleobotany loc. D3124-G, slide (5) at 

110.7X3.0.

3. Equatorial view. USGS paleobotany loc. D3124-G, slide (25) at 

107.2X22.3.

4. Equatorial view. USGS paleobotany loc. D3124-G, slide (6) 

at 81.2X19.7.

5. Equatorial view. USGS paleobotany loc. D1602, slide (2) at 90.9 

X16.2.

6. Polar view. USGS paleobotany loc. D3124-G, slide (15) at 98.9

X20.8. 

7, 8. Oblique equatorial view, high and low focus. USGS paleobotany

loc. D3124-B, slide (2) at 91.5X10.7. 

9. Oblique view. USGS paleobotany loc. D3124-G, slide (5) at

100.5X2.2.

10, 11. Oblique equatorial view, mid and high focus. USGS paleobotany 

loc. D3472-D, slide (1) at 76.9X15.4.

12. Oblique view. USGS paleobotany loc. D3124-G, slide (5) at 

113.0X6.8.

13. Oblique view. USGS paleobotany loc. D3124-G, slide (5) at 105.5 

X7.5.

14. Oblique view. USGS paleobotany loc. D3124-G, slide (5) at 95.9 

X8.8.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 643-A PLATE 1

12

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AQUILAPOLLENITES SENONICUS (MTCHEDLISHVILI) TSCHUDY AND LEOPOLD
PLATE 2

FIGURES 1-8. Aquilapollenites clarireticulatus (Samoilovitch) n. comb. (p. A6)

1. Equatorial view. USGS paleobotany loc. D1799, slide (4) at 95.3X4.9.

2. Equatorial view. USGS paleobotany loc. D1799, slide (4) at 111.6 

X21.3.

3. Equatorial view. USGS paleobotany loc. D1799, slide (4) at 97.3 

X14.1.

4. Equatorial view. USGS paleobotany loc. D1799, slide (4) at 92.7

X19.4.

5, 6. Oblique view, low and high focus. USGS paleobotany loc. D1799, 

slide (4) at 95.3X12.0.

7. Copy of drawings of holotype of Integricorpus clarireticulatus (from 

Samoilovitch, 1965, p. 123). A, X 600 (approximate), B, X 1,350 

(approximate).

8. Scanning electron microscope photograph. Equatorial view; grain 

partially covered by debris. X indicates position X on specimen 

at higher magnification illustrated on pi. 3, fig. 1. USGS paleobotany 

loc. D1799. Photograph by Charles M. Drew.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 643-A PLATE 2

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AQUILAPOLLENITES CLARIRETICULATUS (SAMOILOVITCH) N. COMB.
PLATE 3

FIGURES 1-7. Aquilapollenites clarireticulatus (Samoilovitch) n. comb. (p. A6).

1. Scanning electron microscope photograph (X 10,800) showing 

surface detail. X indicates position X on specimen at lower magnification illustrated on pi. 2, fig. 8. USGS paleobotany loc. D1799. 

Photograph by Charles M. Drew.

2. Equatorial view. USGS paleobotany loc. D1799, slide (4) at 108.2X6.9.

3. Polar view. USGS paleobotany loc. D1799, slide (4) at 102.6X2.9.

4. Polar view. USGS paleobotany loc. D1799, slide (4) at 92.8X15.8.

5. Equatorial view. USGS paleobotany loc. D1799, slide (4) at 

81.4X20.8.

6. Equatorial view. USGS paleobotany loc. D1799, slide (4) at 97.2X6.9.

7. Scanning electron microscope photograph. High magnification 

(X10,800) of the equatorial protrusion shown at the lower left on 

pi. 2, fig. 8. USGS paleobotany loc. D1799. Photograph by Charles 

M. Drew.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 643-A PLATE 3

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AQUILAPOLLENITES CLARIRETICULATUS (SAMOILOVITCH) N. COMB.
PLATE 4

FIGURES 1-5. Aquilapollenites trialatus Rouse (p. A7).

1. Equatorial view. A specimen from Rouse's type material. Rouse 

slide Leth No. 8, l:5(i) Ger; at 78X15.1.

2. Equatorial view. USGS paleobotany loc. D3124-H, slide (1) at 

94.8X19.3.

3. Equatorial view. USGS paleobotany loc. D3124-H, slide (3) at 

115.4X19.8.

4. Equatorial view. USGS paleobotany loc. D3124-H, slide (1) at 

101.3X19.4.

5. Polar view. USGS paleobotany loc. D3124-H, slide (3) at 104.4X 17.2.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 643-A PLATE 4

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AQUILAPOLLENITES TRIALATUS ROUSE
PLATE 5

FIGURES 1-11. Aguilapollenites parallelus n. sp. (p. A7).

1. Equatorial view of holotype. USGS paleobotany loc. D1799, 

slide (5) at 100.7X12.8.

2. Equatorial view. USGS paleobotany loc. D1799, slide (4) at 

107.5X16.1.

3. Equatorial view. USGS paleobotany loc. D1799, slide (4) at 

110.9X19.9.

4. 6. Polar view of a paratype, mid and high focus. USGS paleobotany 

loc. D1799, slide (5) at 79.9X19.8.

5. 7. Polar view, mid and high focus. USGS paleobotany loc. D1799,

slide (7) at 103.1X21.8.

8, 11. Equatorial view of a paratype, high and mid focus. USGS paleobotany loc. D1799, slide (7) at 77.7X14.9.

9. Equatorial view. USGS paleobotany loc. D1799, slide (7) at 

96.2X11.9.

10. Equatorial view. USGS paleobotany loc. D1799, slide (5) at 

104.1X14.1.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 643-A PLATE 5

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AQUILAPOLLENITES PARALLELUS N. SP.
PLATE 6

FIGURES 1-2. Aquilapollenites parallelus n. sp. (p. A7).

1. Scanning electron microscope photograph. Equatorial view (X 4,300 

magnification). USGS paleobotany loc. D1799.

2. Scanning electron microscope photograph. Higher magnification 

(X 10,800) of a portion of grain in fig. 1 (arrows denote corresponding 

positions on figs. 1 and 2). Photographs by Charles M. Drew.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 643-A PLATE 6

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PLATE 7

FIGURES 1-2. Aguilapollentites parallelus n. sp. (p. A7).

1. Scanning electron microscope photograph. Equatorial view of a 

specimen partially covered by debris showing an equatorial protrusion with open colpus at end (X 4,200 magnification). USGS 

paleobotany loc. D1799.

2. Scanning electron microscope photograph. Portion of grain shown in 

fig. 1 at greater magnification (X 10,400). Photographs by Charles 

M. Drew.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 643-A PLATE 7

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PLATE 8

FIGURES 1-12. Aquilapollenites fusiformis n. sp. (p. A8).

1, 2. Equatorial view of holotype, low and high focus. USGS

paleobotany loc. D3124-G, slide (6) at 111.9X18.1. 

3. Polar view of paratype. USGS paleobotany loc. D3124-G, slide

(6) at 109.1X10.2.

4, 5. Polar view, high and low focus. USGS paleobotany loc. D3124- 

G, slide (6) at 117.0X16.5.

6. Equatorial view. USGS paleobotany loc. D3124-G, slide (6) 

at 78.4X5.8.

7. Equatorial view. USGS paleobotany loc. D3124-G, slide (20) 

at 107.1X23.0.

8. Equatorial view. USGS paleobotany loc. D3124-G, slide (36) 

at 105.8X17.1.

9. Equatorial view. USGS paleobotany loc. D3124-G, slide (6) 

at 89.2X21.2.

10. Equatorial view. USGS paleobotany loc. D3124-G, slide (15)

at 93.1X17.6.

11, 12. Oblique view, low and mid focus. USGS paleobotany loc. 

D3124-G, slide (6) at 92.2X21.0.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 643-A PLATE 8

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AQUILAPOLLENITES FUSIFORMIS N. SP.
PLATE 9

FIGURES 1-12. Aquilapollenites dentatus n. sp. (p. A9).

1, 4, 7. Equatorial view of holotype, low, mid and high focus. USGS 

paleobotany loc. D1799, slide (7) at 90.9X8.9.

2. 5. Polar view of paratype, mid and high focus. USGS paleobotany 

loc. D1799, slide (7) at 79.9X7.2.

3. 6. Equatorial view, mid and low focus. USGS paleobotany loc. 

D1799, slide (7) at 90.4X22.0.

8. Polar view. USGS paleobotany loc. D1799, slide (4) at 

113.3X12.8.

9. Equatorial view. USGS paleobotany loc. D1799, slide (5) at

101.7X6.6. 

10, 11. Oblique equatorial view, mid and high focus. USGS paleobotany

loc. D1799, slide (4) at 108.1X14.1.

12. Equatorial view. USGS paleobotany loc. D1799, slide (4) at 

109.2X4.1.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 643-A PLATE. 9

AQUILAPOLLENITES DENTATUS N. SP.
PLATE 10

FIGURE 1. Aquilapollenites dentatus n. sp. (p. A9).

Scanning electron microscope photograph. Oblique view (X 4,200). USGS 

paleobotany loc. D1799. Photograph by Charles M. Drew.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 643-A PLATE 10

AQUILAPOLLENITES DENTATUS N. SP.
PLATE 11

FIGURES 1-18. Aquilapollenites colvillensis n. sp. (p. A9).

1, 2, 3. Equatorial view of holotype, high, mid, and low focus. USGS 

paleobotany loc. D3124-G, slide (5) at 109.5X21.8.

4. 8. Equatorial view of a paratype, mid and low focus. USGS 

paleobotany loc. D3124-G, slide (16) at 83.9X10.4.

5. 6. Polar view of a paratype, high and mid focus. USGS paleobotany loc. D3124-G, slide (6) at 110.1X7.6.

7. Polar view. USGS paleobotany loc. D3124-G, slide (6) at 

90.4X 15.6.

9. Equatorial view. USGS paleobotany loc. D3124-G, slide (36) 

at 93.6X2.2.

10. Oblique view. USGS paleobotany loc. D3124-G, slide (6) at 

84.9X5.8.

11. Equatorial view. USGS paleobotany loc. D3124-G, slide (25)

at 100.1X8.6. 

12, 14. Equatorial view, low and mid focus. USGS paleobotany loc.

D3124-G, slide (6) at 82.9X5.7. 

13. Oblique view. USGS paleobotany loc. D3124-B, slide (2) at

119.0X7.5. 

15. Equatorial view. USGS paleobotany loc. D3124-H, slide (1) at

81.8X7.8. 

16, 17. Equatorial view, high and mid focus. USGS paleobotany loc.

D3124-B, slide (2) at 93.1X15.2. 

18. Oblique equatorial view. USGS paleobotany loc. D3124-B,

slide (17) at 109.0X7.1.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 643-A PLATE 11

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AQUILAPOLLENITES COLVILLENSIS N. SP.
PLATE 12

FIGURES 1-13. Aguilapollenites rectus n. sp. (p. AID)

1. Equatorial view of holotype. USGS paleobotany loc. D1799, slide 

(4) at 113.3X18.1.

2. Polar view of paratype. USGS paleobotany loc. D1799, slide (4)

at 98.4X11.7.

3, 6. Polar view, low and high focus. USGS paleobotany loc. D1799, 

slide (7) at 89.4X22.1.

4. Equatorial view. USGS paleobotany loc. D1799, slide (2) at 111.4 

X8.8.

5. Oblique equatorial view. USGS paleobotany loc. D1799, slide (4) 

at 98.5X4.3.

7. Equatorial view. USGS paleobotany loc. D1799, slide (6) at 86.3 

X17.1.

8. Equatorial view. USGS paleobotany loc. D1799, slide (4) at 101.2 

X2.0.

9. 12. Polar view, mid and low focus. USGS paleobotany loc. D1799, 

slide (6) at 84.3X17.0.

10. 11. Equatorial view, high and mid focus. USGS paleobotany loc. D1799,

slide (7) at 86.5X19.5. 

13. Polar view. USGS paleobotany loc. D1799, slide (4) at 100.0X2.2.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 643-A PLATE 12

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AQUILAPOLLENITES RECTUS N. SP.
PLATE 13

FIGURES 1-3, 5-8, 10-11. Aquilapollenites scabridus n. sp. (p. All).

1, 2. Equatorial view of holotype, two levels. USGS 

paleobotany loc. D1799, slide (5) at 86.2X11.8. 

3. Polar view of paratype. USGS paleobotany loc. 

D1799, slide (7) at 90.3X12.0.

5. Polar view. USGS paleobotany loc. D1799, slide (6) 

at 87.3X9.7.

6. Equatorial view. USGS paleobotany loc. D1799, 

slide (7) at 89.9X3.9.

7. Polar view. USGS paleobotany loc. D1799, slide (5)

at 81.1X10.6.

8, 11. Equatorial view, mid and high focus. USGS paleobotany loc. D1799, slide (7) at 86.8X19.3. 

10. Oblique equatorial view. USGS paleobotany loc.

D1799, slide (4) at 101.3X2.3.

4, 9. Aquilapollenites scabridus n. sp. (p. All) with Aquilapollenites 

rectus n. sp. (p. AlO).

4, 9. Equatorial views of A. scabridus (above) and A. 

rectus (below), high and mid focus. USGS paleobotany loc. D1799, slide (4) at 87.9X13.7.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 643-A PLATE 13

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AQUILAPOLLENITES SCABRIDUS N. SP. AND AQUILAPOLLENITES RECTUS N. SP.
PLATE 14

FIGURES 1-2. Aquilapollenites scabridus n. sp. (p. All).

1. Scanning electron microscope photograph, oblique view (X 4,200). 

USGS paleobotany loc. D1799.

2. Scanning electron microscope photograph. Higher magnification 

(X 10,400) of lower portion of specimen shown in fig. 1. Photographs by Charles M. Drew.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 643-A PLATE 14

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PLATE 15

FIGURES 1-15. Aguilapollenites contiguus n. sp. (p. A12).

1, 2. Equatorial view of holotype, high and mid focus. USGS paleobotany loc. D3124-G, slide (5) at 88.7X5.7.

3. Oblique view of a paratype. USGS paleobotany loc. D3124-G, 

slide (6) at 100.9X3.5.

4. Polar view of a paratype. USGS paleobotany loc. D3124-G, 

slide (5) at 102.7X10.4.

5. Polar view. USGS paleobotany loc. D3124-G, slide (6) at 

92.3X4.6.

6. Oblique view. USGS paleobotany loc. D3124-G, slide (6) at 

111.7X17.3.

7. Equatorial view. USGS paleobotany loc. D3124-G, slide (6) at

82.8X8.8.

8, 11. Oblique view, high and mid focus. USGS paleobotany 

loc. D3124-G, slide (15) at 101.5X14.6.

9. Equatorial view. USGS paleobotany loc. D3124-G, slide (5) at 

87.2X19.8.

10. Equatorial view. USGS paleobotany loc. D3124-G, slide (15) 

at 105.1X11.7.

12. Polar view. USGS paleobotany loc. D3124-G, slide (5) at 

97.2X4.8.

13. Equatorial view. USGS paleobotany loc. D3124-G, slide (5) at 

85.4X15.0.

14. Oblique equatorial view. USGS paleobotany loc. D3124-G, slide 

(15) at 106.1X14.6.

15. Polar view. USGS paleobotany loc. D3124-G, slide (6) at 

117.3X16.4.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 643-A PLATE 15

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AQUILAPOLLENITES CONTIGUUS N. SP.
PLATE 16

FIGURES 1-8. Aquilapollenites laticorpus n. sp. (p. A12).

1. Equatorial view of holotype. USGS paleobotany loc. D3124-G,

slide (15) at 90.1X8.0. 

2, 3. Polar view of paratype, high and mid focus. USGS paleobotany

loc. D3124-G, slide (18) at 116. 8X6. 9. 

4. Oblique equatorial view. USGS paleobotany loc. D3124-G, slide

(15) at 114.5X10.2.

5, 6. Equatorial view, high and mid focus. USGS paleobotany loc. 

D3124-G, slide (6) at 86.9X14.8.

7. Equatorial view. USGS paleobotany loc. D3124-G, slide (5) at 

99.1X21.2.

8. Equatorial view. USGS paleobotany loc. D3124-G, slide (5) at

80.5X16.9. 

FIGURES 9-10. Cf. Aquilapollenites laticorpus (p. A13).

9, 10. Equatorial view, mid and high focus. USGS paleobotany loc.

D3124-G, slide (5) at 101.1X4.8. 

FIGURES 11-20. Fibulapottis scabratus n. sp. (p. A13).

11. Equatorial view of holotype. USGS paleobotany loc. D3124-A, 

slide (1) at 97.8X7.3.

12. Polar view of a paratype. USGS paleobotany loc. D3124-A, 

slide (1) at 103.7X20.2.

13. Oblique equatorial view of a paratype. USGS paleobotany loc. 

D3124-A, slide (1) at 96.8X20.2.

14. Oblique equatorial view. USGS paleobotany loc. D3124-G, slide 

(5) at 81.9X10.2.

15. Equatorial view. USGS paleobotany loc. D3124-G, slide (5) 

at 94.8X2.1.

16. Equatorial view. USGS paleobotany loc. D1330-4, slide (3) at 

95.4X3.9.

17. Oblique equatorial view. USGS paleobotany loc. D3124-G, 

slide (5) at 107.3X21.0.

18. Polar view. USGS paleobotany loc. D1330-19, slide (3) at 84.8X

11.5.

19, 20. Oblique view, low and high focus. USGS paleobotany loc. D3124- 

G, slide (15) at 97.7X17.7.
GEOLOGICAL SURVEY PROFESSIONAL PAPER 643-A PLATE 16

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AQUILAPOLLENITES LATICORPUS N. SP. AND FIBULAPOLLIS SCABRATUS N. SP.