The present distribution of most perissodactyl species is only a small fraction of their original range. Members of this group are now found only in Central and South America, eastern and southern Africa, and central, southern, and southeastern Asia. During the peak of odd-toed ungulate existence, from the Eocene to the Oligocene, perissodactyls were distributed over much of the globe, the only exceptions being Australia and Antarctica. Horses and tapirs arrived in South America after the formation of the Isthmus of Panama in the Pliocene, around 3 million years ago. In North America, they died out around 10,000 years ago, while in Europe, the tarpans disappeared in the 19th century. Hunting and habitat restriction have reduced the present-day species to fragmented relict populations. In contrast, domesticated horses and donkeys have gained a worldwide distribution, and feral animals of both species are now also found in regions outside of their original range, such as in Australia.