As a result, Henry of Meissen gained the bulk of Thuringia in 1264, while the Hessian possessions of the landgraves was separated as the Landgraviate of Hesse under the rule of Sophie's son Henry I. The Meissen margraves of the Wettin dynasty retained the landgravial title. Upon the death of Margrave Frederick III of Meissen his younger brothers divided their heritage in the 1382 Division of Chemnitz, whereby Thuringia passed to Balthasar. Upon the death of Landgrave Frederick IV in 1440, Thuringia fell to his nephew Elector Frederick II of Saxony, the inheritance conflict with his brother William III led to the 1445 Division of Altenburg and the Saxon Fratricidal War over the Wettin lands. The Thuringian lands fell to William III, when he died childless in 1482, Elector Ernest, inherited the landgraviate, uniting the Wettin lands under his rule. After the 1485 Treaty of Leipzig, Thuringia split into the Saxon Ernestine and Albertine duchies.