The 339th Rifle Division was first formed in late August, 1941, as a standard Red Army rifle division, at Rostov-on-Don. As it was formed in part from reservists and cadre that included members of the Communist Party from that city, it carried the honorific title "Rostov" for the duration. In late November it was part of the force that counterattacked the German 1st Panzer Army in the Battle of Rostov and forced its retreat from the city, one of the first major setbacks for the invaders. During 1942 the division was forced to retreat into the Caucasus, where it fought to defend the passes leading to the Black Sea ports. In 1943 it fought to liberate the Taman Peninsula, and then in early 1944 to also liberate Crimea. In the following months the division was reassigned to the 1st Belorussian Front, with which it took part in the Battle of Berlin in 1945. Following a distinguished career, the division was disbanded in the summer of that year.