Starting from 750 AD large urban centers such as Monte Albán fell across the Oaxaca area and smaller dominions grew and evolved until the Spanish Conquest in 1521. Between 700 and 1300, the Mixtec were scattered among various dominions, including those of Achiutla, Tequixtepec-Chazumba, Apoala and Coixtlahuaca. The Zapotecs occupied a large region from Central Valleys region to the Isthmus of Tehuantepec. However, no major city state like Monte Albán arose again, with villages and city-states remaining small, between 1,000 and 3,000 people with a palace, temple, market and residences. In a number of cases, there were Mesoamerican ball courts as well. These and larger centers also functioned as military fortresses in time of invasion. Important Zapotec and Mixtec sites include Yagul, Zaachila, Inguiteria, Yanhuitlan, Tamazulapan, Tejupan, and Teposcolula. During nearly all of this time, these various entities were at war with one another, and faced the threat of Aztec expansion.