After Andronikos II took to the throne, the army fell apart and the Byzantines suffered regular defeats at the hands of their eastern opponents, although they would continue to enjoy success against the crusader territories in Greece. By c. 1350, following a destructive civil war and the outbreak of the Black Death, the Empire was no longer capable of raising troops and the supplies to maintain them. The Empire came to rely upon troops provided by Serbs, Bulgarians, Venetians, Latins, Genoans and Ottoman Turks to fight the civil wars that lasted for the greater part of the 14th century, with the latter foe being the most successful in establishing a foothold in Thrace. The Ottomans swiftly expanded through the Balkans and cut off Constantinople, the capital of the Byzantine Empire, from the surrounding land. The last decisive battle was fought by the Palaiologan army in 1453, when Constantinople was besieged and fell on 29 May. The last isolated remnants of the Byzantine state were conquered by 1461.