The first anatomically modern humans ("Homo sapiens sapiens") in North Africa are the makers of the Aterian, a Middle Stone Age (or Middle Palaeolithic) stone tool culture. The earliest Aterian lithic assemblages date to around 145,000 years ago, and were discovered at the site of Ifri n'Ammar in Morocco. This industry was followed by the Iberomaurusian culture, a backed bladelet industry found throughout the Maghreb. It was originally described in 1909 at the site of Abri Mouillah. Other names for this Cro-Magnon-associated culture include "Mouillian" and "Oranian". The Epipaleolithic Iberomaurusian makers were centred in prehistoric sites, such as Taforalt and Mechta-Afalou. They were succeeded by the Capsians. The Capsian culture is often thought to have arrived in Africa from the Near East, although it is also suggested that the Iberomaurusians may have been the progenitors of the Capsians. Around 5000 BC, the populations of North Africa were primarily descended from the makers of the Iberomaurusian and Capsian cultures, with a more recent intrusion associated with the Neolithic revolution.