Following Darius the Great's victory over the Magian usurper, Gaumata, in September 522 BC, revolts spread throughout the empire. In December 522 BC, a revolt in support of the Median leader Phraortes erupted in Hyrcania, and in March 521 BC, the Hyrcanian rebels unsuccessfully attacked Hystaspes, satrap of Parthia. In May, Phraortes was defeated and Hyrcania returned to Achaemenid rule. Darius later settled Hyrcanians in the settlement of Dareionkome () in the Hyrcanian Plain in Lydia. Other Hyrcanian settlements in the Hermus valley include Ormoita and Tyanolla. According to Herodotus, Hyrcanian soldiers participated in the Second Persian invasion of Greece in 480 BC under the command of Megapanus. After the war, a garrison composed of Hyrcanian soldiers was posted in the valleys of the rivers Hermus and Caicus. Under Xerxes I, Hyrcania was likely detached from the satrapy of Parthia and administered separately. Some sources imply Hyrcania was later administered as a sub-province of Media. According to Ctesias, Artaxerxes I appointed his illegitimate son Ochus as satrap of Hyrcania in c. 425 BC, who later assumed the throne as Darius II and appointed Idernes as satrap. Idernes' son Terituchmes succeeded him as satrap of Hyrcania following his death and was married to Amestris, daughter of Darius II, but Terituchmes was murdered by Udiastes, a henchman of Terituchmes, for conspiring to murder Amestris and revolt against Darius so he could wed his half-sister Roxana.