Octavian's land redistribution caused widespread tensions, as farmers were dispossessed in favor of soldiers. Antony's brother Lucius Antonius, who was serving as Consul, stood up for the dispossessed farmers. The conflict led to the Perusine War, in which Lucius gathered an army of supporters to challenge Octavian. He was encouraged by Mark Antony's wife Fulvia. Lepidus held Rome with two legions while Octavian left to gather his army, but Lucius defeated Lepidus, who was forced to flee to Octavian. As Octavian advanced on Rome, Lucius withdrew to Perusia (Perugia), where he was besieged by Octavian in the winter of 41–40 BC. He finally surrendered in exchange for clemency. The outcome was that Lepidus was confirmed as governor of Africa, acquiring six of Antony's legions, leaving Octavian as the sole power in Italy, with his own loyal legions in control. When Antony's supporter Calenus, governor of Gaul, died, Octavian took over his legions, further strengthening his control over the west. This new distribution of power among the triumvirs was confirmed by the Treaty of Brundisium in September 40 BC. At around the same time, Antony's wife Fulvia died. Octavian arranged for Antony to marry his sister, Octavia, as a symbol of the renewed alliance.