Except for the royal city of Amathus, the Kingdoms of Cyprus took part in the Ionian Revolt in 499 BC. The revolt on Cyprus was led by Onesilos of Salamis, brother of the King of Salamis, whom he dethroned for not wanting to fight for independence. The Persians crushed the Cypriot armies and laid siege to the fortified towns in 498 BCE. Soloi surrendered after a five-month siege.