In pre-Columbian times, a number of Mixtecan city states competed with each other and with the Zapotec kingdoms. The major Mixtec polity was Tututepec which rose to prominence in the 11th century under the leadership of Eight Deer Jaguar Claw, the only Mixtec king who ever united the Highland and Lowland polities into a single state. Like the rest of the indigenous peoples of Mexico, the Mixtec were conquered by the Spanish invaders and their indigenous allies in the 16th century. Pre-Columbian Mixtecs numbered around 1.5 million. Today there are approximately 800,000 Mixtec people in Mexico, and there are also large populations in the United States.