In 267 BC, Ptolemy II persuaded the Greek cities to revolt against Macedon, in what became the Chremonidean War, after the Athenian leader Chremonides. The cities were defeated and Athens lost her independence and her democratic institutions. This marked the end of Athens as a political actor, although it remained the largest, wealthiest and most cultivated city in Greece. In 225 BC, Macedon defeated the Egyptian fleet at Cos and brought the Aegean islands, except Rhodes, under its rule as well.