A total of 16 tropical depressions were recorded, of which 15 further intensified into tropical storms. Of those 15, a total of seven strengthened into hurricanes, while four attained their peaks as major hurricanes. Activity began with Alex which, upon making landfall in the Azores, became the first January landfalling tropical cyclone since Hurricane Alice in the 1954 season. In June, tropical storms Colin and Danielle became the earliest third and fourth named storms, respectively, on record. Hermine moved ashore the coastline of Florida as a Category 1 hurricane on September 2, ending the record hurricane drought that began in the state after the 2005 season's Hurricane Wilma. In late September and early October, Hurricane Matthew wrought destruction throughout the Caribbean Sea and Southeastern United States, resulting in $15.09 billion (2016 USD) in damage and 603 deaths; it remains the ninth-costliest hurricane on record in the Atlantic, as well as the tenth-costliest in the United States. In mid-October, Hurricane Nicole developed to Category 4 intensity and remained a major hurricane while directly impacting Bermuda, the first storm of such strength to do so since Hurricane Fabian in the 2003 season. Following the 2016 season, the names "Matthew" and "Otto" were retired from reuse in the North Atlantic by the World Meteorological Organization.