In 480, the last Western Roman emperor, Julius Nepos, was murdered and a Roman general of barbarian origin, Odoacer, declared allegiance to Eastern Roman emperor Zeno. Despite owing nominal allegiance to Constantinople, Odoacer and later the Ostrogoths continued, like the last emperors, to rule Italy as a virtually independent realm from Ravenna. Meanwhile, the Senate, even though long since stripped of wider powers, continued to administer Rome itself, with the Pope usually coming from a senatorial family. This situation continued until Theodahad murdered Amalasuntha, a pro-imperial Gothic queen, and usurped the power in 535. The Eastern Roman emperor, Justinian I (reigned 527–565), used this as a pretext to send forces to Italy under his famed general Belisarius, recapturing the city next year, on December 9, 536 AD. In 537–538, the Eastern Romans successfully defended the city in a year-long siege against the Ostrogoth army, and eventually took Ravenna, too.