The Huns crossed the Volga River from the east and forced large groups of Alans and Goths to abandon their homelands in the Pontic steppes. Hun, Alan and Goth troops pillaged Pannonia for the first time in the winter of 379–380. The Romans allowed the Marcomanni to settle in Pannonia Prima when their queen, Fritigil, converted to Christianity around 396. The Huns' center of power was transferred to the lowlands along the Middle Danube in the 420s, but their empire collapsed after the Germanic and Iranian subject peoples defeated them in the Battle of Nedao in 454. The Gepids took control of the lands to the east of the river Tisza and the Romans allowed the Ostrogoths to settle in Pannonia. in the early 470s, the Ostrogoths left for Italy and less powerful Germanic peoples, like the Heruli and Suebi, seized parts of Pannonia. The Lombards occupied the province in the 500s.