Philadelphia is one of the oldest municipalities in the United States. William Penn, an English Quaker, founded the city in 1682 to serve as capital of the Pennsylvania Colony. Philadelphia played an instrumental role in the American Revolution as a meeting place for the Founding Fathers of the United States, who signed the Declaration of Independence in 1776 at the Second Continental Congress, and the Constitution at the Philadelphia Convention of 1787. Several other key events occurred in Philadelphia during the Revolutionary War including the First Continental Congress, the preservation of the Liberty Bell, the Battle of Germantown, and the Siege of Fort Mifflin. Philadelphia remained the nation's largest city until being overtaken by New York City in 1790; the city was also one of the nation's capitals during the revolution, serving as temporary U.S. capital while Washington, D.C. was under construction. In the 19th and 20th centuries, Philadelphia became a major industrial center and a railroad hub. The city grew due to an influx of European immigrants, most of whom initially came from Ireland and Germany—the two largest reported ancestry groups in the city . Later immigrant groups in the 20th century came from Italy (Italian being the third largest European ethnic ancestry currently reported in Philadelphia) and other Southern European and Eastern European countries. In the early 20th century, Philadelphia became a prime destination for African Americans during the Great Migration after the Civil War. Puerto Ricans began moving to the city in large numbers in the period between World War I and II, and in even greater numbers in the post-war period. The city's population doubled from one million to two million people between 1890 and 1950.