In the Pacific, the Allies returned to the Philippines, recaptured Burma, and invaded Borneo. Offensives were undertaken to reduce the Japanese forces remaining in Bougainville, New Guinea and the Philippines. In April 1945, American forces landed on Okinawa, where heavy fighting continued until June. Along the way, the ratio of Japanese to American casualties dropped from five to one in the Philippines to two to one on Okinawa. Although some Japanese soldiers were taken prisoner, most fought until they were killed or committed suicide. Nearly 99 percent of the 21,000 defenders of Iwo Jima were killed. Of the 117,000 Okinawan and Japanese troops defending Okinawa in April to June 1945, 94 percent were killed; 7,401 Japanese soldiers surrendered, an unprecedentedly large number.