On 24 October 1940, Vichy French LeO 451s carried out an air raid against Gibraltar in retaliation for the Free-French attack on Dakar, losing one of their number to British anti-aircraft fire. Two bomber units equipped with LeO 451s, GB I/12 and GB I/31 were based in Syria when Allied forces invaded on 8 June 1941, at the start of the Syria-Lebanon Campaign. These were supplemented by GB I/25, which was dispatched from Tunisia. During this campaign, the LeO 451s flew a total of 855 sorties, losing 29 LeO 451s in the process. After Operation Torch which began on 8 November 1942, surviving French LeO 451 in North Africa were used primarily for freight duties, although they flew a few bombing missions against Axis forces during the Tunisia Campaign. They were replaced by Handley-Page Halifax and B-26 Marauder bombers.