When "The Origin of Species" was published FitzRoy was dismayed and apparently felt guilty for his part in the theory's development. He was in Oxford on 30 June 1860 to present a paper on storms and attended the meeting of the British Association for the Advancement of Science at which Samuel Wilberforce attacked Darwin's theory. During the debate FitzRoy, seen as "a grey haired Roman nosed elderly gentleman", stood in the centre of the audience and "lifting an immense Bible first with both and afterwards with one hand over his head, solemnly implored the audience to believe God rather than man". As he admitted that "The Origin of Species" had given him "acutest pain", the crowd shouted him down.