Soviet civilian populations in the occupied areas were severely persecuted and endured the treacherous conditions of the Eastern Front, which spawned atrocities such as the siege of Leningrad, when 1.2 million civilians died. Thousands of peasant villages across Russia, Belarus and Ukraine were annihilated by German troops. During the occupation, the Leningrad, Pskov and Novgorod regions lost about a quarter of their populations. An estimated one-quarter of Soviet civilian deaths at the hands of the Nazis and their allies (five million Russians, three million Ukrainians and 1.5 million Belarusians) were racially motivated. In 1995, the Russian Academy of Sciences reported that civilian deaths in the occupied USSR, including Jews, at the hands of the Germans totaled 13.7 million dead (20 percent of the population of 68 million). The figure includes 7.4 million victims of Nazi genocide and reprisals, 2.2 million deaths of persons deported to Germany as forced labour, and 4.1 million famine and disease deaths. An estimated three million people also died of starvation in unoccupied territory. The losses occurred within the 1946–1991 borders of the USSR, and include territories annexed in 1939–40. The deaths of 8.2 million Soviet civilians, including Jews, were documented by the Soviet Extraordinary State Commission.