By 1 June, Kesselring tried to concentrate the 90th, the 1st Parachute, 305th Infantry and 94th Infantry divisions into a defensive line between Piglio and Paliano. But by then, these units were very weak and were constantly in contact with Allied forces, making relocation extremely difficult. By 4 June, the 90th were again fighting rear-guard actions on the line Ponte Orsino - Trevi and had retreated further north. At the end of July 1944, the division was again relocated into the Po Valley between Modena and Parma and then to Liguria north of Genoa to re-group. After the Allied landings in southern France, the division was relocated to Piedmont to secure the border of the Alpine crossings from France into Italy.