Altogether, 3 million Polish Jews – approximately 90% of Poland's pre-war Jewry – and between 1.8 and 2.8 million ethnic Poles were killed during the German occupation of Poland, including between 50,000 and 100,000 members of the Polish intelligentsia – academics, doctors, lawyers, nobility and priesthood. During the Warsaw Uprising alone, over 150,000 Polish civilians were killed, most were murdered by the Germans during the Wola and Ochota massacres. Around 150,000 Polish civilians were killed by Soviets between 1939 and 1941 during the Soviet Union's occupation of eastern Poland (Kresy), and another estimated 100,000 Poles were murdered by the Ukrainian Insurgent Army (UPA) between 1943 and 1944 in what became known as the Wołyń Massacres. Of all the countries in the war, Poland lost the highest percentage of its citizens: around 6 million perished – more than one-sixth of Poland's pre-war population – half of them Polish Jews. About 90% of deaths were non-military in nature.