During the Byzantine civil war of 1352–1357, Bera was captured by the forces of John V Palaiologos in 1355. Bera was conquered by the Ottomans under Lala Shahin Pasha in the early 1370s, and by 1433, it is recorded as a town with a mixed Greek and Turkish population. The fortifications had been party destroyed, and the "katholikon" of the monastery had been converted into a mosque.