The oldest surviving church features in the district are from the Saxon era, found in a string of churches on the edges of the South Downs or in the flat plains around the River Cuckmere: at Arlington, East Dean, Friston and Jevington. Norman work survives in the churches at Alciston, Berwick, Hellingly, Herstmonceux, Isfield, Litlington, Little Horsted, Rotherfield (whose original church was founded in 792), Westdean, Westham and Wilmington. A rare Norman-era piscina survives at East Hoathly in an otherwise largely Victorian church, and at Hartfield a mostly late medieval church retains Norman masonry in one wall. Several Norman features survive at Maresfield but were moved around during Victorian restoration. Wadhurst's church was added to throughout the medieval era, but its Norman tower survives; and part of the Norman chancel survives at Pevensey in an otherwise mostly 13th-century church. Churches largely or wholly of that century—some of which replaced earlier buildings—can be found at Buxted, Chalvington, Chiddingly, Fletching (although Pevnser describes its tower as Norman), Folkington, Framfield (rebuilt in the 16th century after a fire), Lullington and Waldron. Features from the 13th century also survive at Hooe, Laughton, Ripe, Warbleton, Wartling and Willingdon, which were rebuilt or extended later in the medieval era, and at Ninfield in a church subjected to extensive 19th-century rebuilding. At Alfriston, the original Anglo-Saxon church was replaced in the 1360s by the vast cruciform "Cathedral of the Downs", whose flintwork is "some of the best in England". Also from that century is the church at Old Heathfield. Mayfield's ancient church (founded by Dunstan, the saint to which it is dedicated) is mostly 15th-century. English Anglican churches wholly of the 17th and 18th centuries are rare, but Withyham (rebuilt in 1680 after the predecessor burnt down) and Crowborough (newly built in 1744 and altered later) are represented in the district.