Most of the Arab tribes in central and southern Syria remained loyal to the Umayyads and selected the non-Sufyanid Marwan ibn al-Hakam from Medina to succeed Mu'awiya II. The proclamation of Marwan as caliph in Damascus marked a turning point against Ibn al-Zubayr. Marwan's partisans, led by Ubayd Allah ibn Ziyad, decisively defeated the pro-Zubayrid Qaysi tribes, led by al-Dahhak ibn Qays al-Fihri, at the Battle of Marj Rahit in July 684. The surviving Qaysi tribesmen fled to the Jazira (Upper Mesopotamia) under the leadership of Zufar ibn al-Harith al-Kilabi, who maintained his recognition of Ibn al-Zubayr's suzerainty. However, in March 685, Ibn al-Zubayr lost the economically important province of Egypt to Marwan.