Task: Question Generation
Input (English): Background Paragraph: The brains of all mammals have a unique layer of nerve cells covering the cerebrum. This layer is called the neocortex (the pink region of the brains in Figure above ). The neocortex plays an important role in many complex brain functions. In some mammals, such as rats, the neocortex is relatively smooth. In other mammals, especially humans, the neocortex has many folds. The folds increase the surface area of the neocortex. The larger this area is, the greater the mental abilities of an animal. 
Story: Two colleagues studied at the same university. Nick studied the brains of birds, while Joe studied the brains of mammals.
Output (English): Which colleague studied more about the smooth neocortex of rats?