Task: Question Answering
Input (English): Background Paragraph: Alpine tundra occurs at high enough altitude at any latitude. Portions of montane grasslands and shrublands ecoregions worldwide include alpine tundra. Large regions of alpine tundra occur in the North American Cordillera, the Alps and Pyrenees of Europe, the Himalaya and Karakoram of Asia, the Andes of South America, and the Eastern Rift mountains of Africa.Alpine tundra occupies high-mountain summits, slopes, and ridges above timberline. Aspect plays a role as well; the treeline often occurs at higher elevations on warmer equator-facing slopes. Because the alpine zone is present only on mountains, much of the landscape is rugged and broken, with rocky, snowcapped peaks, cliffs, and talus slopes, but also contains areas of gently rolling to almost flat topography.Averaging over many locations and local microclimates, the treeline rises 75 metres (245 ft) when moving 1 degree south from 70 to 50°N, and 130 metres (430 ft) per degree from 50 to 30°N. Between 30°N and 20°S, the treeline is roughly constant, between 3,500 and 4,000 metres (11,500 and 13,100 ft). 
Story: Gill and Ann were two fellow botanists. Gill studied alpine tundras extensively, while Ann studied tropical forests. They both discovered very interesting facts about nature and plants. 
Question: Which person learned more about shrublands ecoregions?
Output (English): Gill.