Task: Question Generation
Input (English): During the Cenozoic era, specifically about 25 million years ago during the Miocene and Pliocene epochs, the continental climate became favorable to the evolution of grasslands. Existing forest biomes declined and grasslands became much more widespread. The grasslands provided a new niche for mammals, including many ungulates and glires, that switched from browsing diets to grazing diets. Traditionally, the spread of grasslands and the development of grazers have been strongly linked. However, an examination of mammalian teeth suggests that it is the open, gritty habitat and not the grass itself which is linked to diet changes in mammals, giving rise to the "grit, not grass" hypothesis.
Output (English): the spread of grasslands and what have been strongly linked?