表(4)-1 フタバガキ科で最も大きな属である Shorea 属の種について、マレーシア森林研究所樹木園、サバ森林研究センター樹木園、サラワク州ランビル国立公園及び天然林から収集を行った。またドイツのゲッチンゲン大学が収集したサンプルについても分譲を受け解析を行った。

表-1 種識別で收集した材料
属	種	材分類のグループ名	Section no.²	採取地または起源	国と地域
Shorea	atrinervosa	Balau	1a	Forest Research Institute Malaysia	Peninsular, Malaysia
Shorea	atrinervosa	Balau	1a	Forest Research Institute Malaysia	Peninsular, Malaysia
Shorea	biawak	Balau	1b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	biawak	Balau	1b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	biawak	Balau	1b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	coltina	Balau	1a	Forest Research Institute Malaysia arboretum	Peninsular, Malaysia
Shorea	crassa	Balau	1a	Lambir Hills National Park	Sarawak, Malaysia
Shorea	domatiosa	Balau	1a	Forest Research Institute Malaysia arboretum	Peninsular, Malaysia
Shorea	falcifera	Balau	1a	Kob Moi	Peninsular, Malaysia
Shorea	falcifera	Balau	1a	Forest Research Centre arboretum	Sabah, Malaysia
Shorea	foxworthyi	Balau	1a	Forest Research Institute Malaysia	Peninsular, Malaysia
Shorea	guiso	Balau	1a	Forest Research Institute Malaysia arboretum	Peninsular, Malaysia
Shorea	havilandii	Balau	1a	Lambir Hills National Park	Sarawak, Malaysia
Shorea	havilandii	Balau	1a	Lambir Hills National Park	Sarawak, Malaysia
Shorea	havilandii	Balau	1a	Lambir Hills National Park	Sarawak, Malaysia
Shorea	inappendiculata	Balau	1a	Lambir Hills National Park	Sarawak, Malaysia
Shorea	isoptera	Balau	3	Forest Research Institute Malaysia arboretum	Peninsular, Malaysia
Shorea	laevis	Balau	1b	Georg-August University of Goettingen	Indonesia
Shorea	laevis	Balau	1b	PT Sari Bumi Kusuma	Indonesia, Kalimantan
Shorea	laevis	Balau	1b	PT Sari Bumi Kusuma	Indonesia, Kalimantan
Shorea	materialis	Balau	1a	Forest Research Institute Malaysia	Peninsular, Malaysia
Shorea	materialis	Balau	1a	Forest Research Institute Malaysia	Peninsular, Malaysia
Shorea	materialis	Balau	1a	Forest Research Institute Malaysia	Peninsular, Malaysia
Shorea	materialis	Balau	1a	Forest Research Institute Malaysia	Peninsular, Malaysia
Shorea	maxwelliana	Balau	1b	Georg-August University of Goettingen	Indonesia
Shorea	obscura	Balau	1a	Forest Research Institute Malaysia arboretum	Peninsular, Malaysia
Shorea	ochrophloia	Balau	1a	Forest Research Institute Malaysia arboretum	Peninsular, Malaysia
Shorea	seminis	Balau	1a	Georg-August University of Goettingen	Indonesia
Shorea	sumatrana	Balau	1a	Forest Research Institute Malaysia	Peninsular, Malaysia
Shorea	sumatrana	Balau	1a	Forest Research Institute Malaysia	Peninsular, Malaysia
Shorea	sumatrana	Balau	1a	Forest Research Institute Malaysia	Peninsular, Malaysia
Shorea	superba	Balau	1a	Lambir Hills National Park	Sarawak, Malaysia
Shorea	superba	Balau	1a	Lambir Hills National Park	Sarawak, Malaysia
Shorea	superba	Balau	1a	Lambir Hills National Park	Sarawak, Malaysia
Shorea	acuminata	Red Meranti	9b	Forest Research Institute Malaysia	Peninsular, Malaysia
Shorea	acuminata	Red Meranti	9b	Forest Research Institute Malaysia	Peninsular, Malaysia
Shorea	acuminata	Red Meranti	9b	Georg-August University of Goettingen	Indonesia
Shorea	acuma	Red Meranti	9a	Lambir Hills National Park	Sarawak, Malaysia
Shorea	acuta	Red Meranti	9a	Lambir Hills National Park	Sarawak, Malaysia
Shorea	acuta	Red Meranti	9a	Lambir Hills National Park	Sarawak, Malaysia
Shorea	acuta	Red Meranti	9a	Lambir Hills National Park	Sarawak, Malaysia
Shorea	albida	Red Meranti	6	Forest Research Centre Sarawak	Sarawak, Malaysia
Shorea	albida	Red Meranti	6	Forest Research Centre Sarawak	Sarawak, Malaysia
Shorea	almon	Red Meranti	7b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	almon	Red Meranti	7b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	almon	Red Meranti	7b	Forest Research Centre arboretum	Sabah, Malaysia
Shorea	amplexicaulis	Red Meranti	8	Lambir Hills National Park	Sarawak, Malaysia
Shorea	amplexicaulis	Red Meranti	8	Lambir Hills National Park	Sarawak, Malaysia
Shorea	andulensis	Red Meranti	7b	Georg-August University of Goettingen	Indonesia
Shorea	argentifolia	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	beccariana	Red Meranti	8	Forest Research Institute Malaysia arboretum	Peninsular, Malaysia
Shorea	hullata	Red Meranti	7b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	curisii	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	curisii	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	dasyphylla	Red Meranti	9b	Georg-August University of Goettingen	Indonesia
Shorea	dasyphylla	Red Meranti	9b	Forest Research Institute Malaysia arboretum	Peninsular, Malaysia
Shorea	fallax	Red Meranti	7b	Forest Research Institute Malaysia arboretum	Peninsular, Malaysia
Shorea	fallax	Red Meranti	7b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	fallax	Red Meranti	7b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	fallax	Red Meranti	7b	Georg-August University of Goettingen	Indonesia
Shorea	fallax	Red Meranti	7b	Plot, Deramakot forest reserve	Sabah, Malaysia
Shorea	fallax	Red Meranti	7b	PT Sari Bumi Kusuma	Indonesia, Kalimantan
Shorea	ferruginea	Red Meranti	9a	Lambir Hills National Park	Sarawak, Malaysia
Shorea	ferruginea	Red Meranti	9a	Lambir Hills National Park	Sarawak, Malaysia
Shorea	flaviflora	Red Meranti	7b	Forest Research Institute Malaysia arboretum	Peninsular, Malaysia
Shorea	johrensis	Red Meranti	7b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	johrensis	Red Meranti	7b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	johrensis	Red Meranti	7b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	johrensis	Red Meranti	7b	Georg-August University of Goettingen	Indonesia
Shorea	johrensis	Red Meranti	7b	Forest Research Centre arboretum	Sabah, Malaysia
Shorea	johrensis	Red Meranti	7b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	johrensis	Red Meranti	7b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	johrensis	Red Meranti	7b	PT Sari Bumi Kusuma	Indonesia, Kalimantan
Shorea	kunstleri	Red Meranti	7b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	kunstleri	Red Meranti	7b	Forest Research Institute Malaysia	Peninsular, Malaysia
Shorea	kunstleri	Red Meranti	7b	Forest Research Institute Malaysia	Peninsular, Malaysia
Shorea	lepidota	Red Meranti	9b	Georg-August University of Goettingen	Indonesia
Shorea	leprosula	Red Meranti	9b	Georg-August University of Goettingen	Indonesia
Shorea	leprosula	Red Meranti	9b	Forest Research Centre arboretum	Sabah, Malaysia
Shorea	leprosula	Red Meranti	9b	Forest Research Institute Malaysia	Peninsular, Malaysia
Shorea	leprosula	Red Meranti	9b	Forest Research Institute Malaysia	Peninsular, Malaysia
Shorea	leprosula	Red Meranti	9b	Forest Research Institute Malaysia	Peninsular, Malaysia
Shorea	leprosula	Red Meranti	9b	Forest Research Institute Malaysia	Peninsular, Malaysia
Shorea	leprosula	Red Meranti	9b	Forest Research Institute Malaysia	Peninsular, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Georg-August University of Goettingen	Indonesia
Shorea	leprosula	Red Meranti	9b	Georg-August University of Goettingen	Indonesia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lambir Hills National Park	Sarawak, Malaysia
Shorea	leprosula	Red Meranti	9b	Lamb

Ashton (1982), 1; Shorca (1a; Shorca, 1b; Barbata), 2; Pentacme, 3; Neohopea, 4; Richetioides (4a; Polyandrae, 4b; Richetioides), 5; Anthoshorea, 6; Rubella, 7; Brachypterae (7a; Smithiana, Brachypterae), 8; Pachycarpae, 9; Mutica (9a; Auriculatae, 9b; Mutica) and 10; Ovalis