"標的を検出する方法であって、以下:サンプルを核酸バーコードプローブと接触させるステップ、及び前記核酸バーコードプローブが、前記サンプル中の1つ以上の成分と結合するか否かを決定するステップを含み、前記核酸バーコードプローブと前記サンプルの1つ以上の成分との結合が、前記サンプル中の標的の存在を示し、また、前記核酸バーコードプローブが、標的結合部分と、少なくとも2つの蛍光標識領域とを含む核酸ナノ構造を含む、方法。前記サンプルの1つ以上の成分と結合した前記核酸バーコードプローブの蛍光標識領域の色及び/又は配向に基づいて、標的を識別するステップをさらに含む、請求項1に記載の方法。前記核酸バーコードプローブが、前記サンプル中の1つ以上の成分と結合するか否かが、前記核酸バーコードプローブを、可溶性の、一時的に結合する発蛍光団担持オリゴヌクレオチドと接触させるステップを含む、請求項1又は2に記載の方法。前記核酸バーコードプローブが、安定に結合した発蛍光団担持オリゴヌクレオチドを含む、請求項1又は2に記載の方法。標的を検出する方法であって、前記標的が核酸バーコードプローブと結合するのに十分な条件下で、前記標的を核酸バーコードプローブと接触させるステップ;前記標的と結合していない材料から前記標的を分離するステップ;及び前記標的と結合した前記核酸バーコードプローブの存在を検出するステップを含み、前記核酸バーコードプローブが、標的結合部分と、少なくとも2つの蛍光標識領域とを含む核酸ナノ構造を含む、方法。前記標的と結合した前記核酸バーコードプローブの蛍光標識領域の色及び/又は配向に基づいて、前記標的を識別するステップをさらに含む、請求項5に記載の方法。前記標的と結合した前記核酸バーコードプローブの存在を検出するステップが、前記核酸バーコードプローブを、可溶性の、一時的に結合する発蛍光団担持オリゴヌクレオチドと接触させるステップを含む、請求項5又は6に記載の方法。前記核酸バーコードプローブが、安定に結合した発蛍光団担持オリゴヌクレオチドを含む、請求項5又は6に記載の方法。サンプルと、少なくとも2つの蛍光標識領域を有する核酸ナノ構造を含む核酸バーコードプローブを含む組成物であって、前記核酸ナノ構造上の前記蛍光標識領域の各々の位置は、前記蛍光標識のパターンが、前記蛍光標識領域による可視光線の発光に基づいて決定可能であるようにする、組成物。前記核酸ナノ構造が、スカフォールド鎖と複数のステープル鎖を含む、請求項9に記載の組成物。前記蛍光標識領域が、蛍光標識ステープル鎖を含む、請求項9又は10に記載の組成物。前記蛍光標識ステープル鎖が、aスカフォールド鎖にハイブリダイズしたステープルドメインとbハンドルドメインとを含む、請求項11に記載の組成物。前記ハンドルドメインが、発蛍光団標識オリゴヌクレオチドにハイブリダイズしている、請求項12に記載の組成物。前記核酸ナノ構造が、DNAナノ構造である、請求項9~13のいずれか一項に記載の組成物。前記DNAナノ構造がナノチューブである、請求項14に記載の組成物。前記核酸ナノ構造が、少なくとも3つの蛍光標識領域又は少なくとも4つの蛍光標識領域を含む、請求項9~15のいずれか一項に記載の組成物。前記少なくとも3つ又は少なくとも4つの蛍光標識領域の配向が、前記蛍光標識領域同士の非対称間隔に基づいて決定可能である、請求項16に記載の組成物。前記蛍光標識領域の各々が、前記他の蛍光標識領域の中心から少なくとも約50nm又は少なくとも約100nmに位置する中心を有する、請求項9~17のいずれか一項に記載の組成物。前記蛍光標識領域の各々が、前記他の蛍光標識領域の中心から少なくとも約200nmの地点に位置する中心を有する、請求項9~18のいずれか一項に記載の組成物。前記蛍光標識領域の各々の前記中心が、あらゆる他の蛍光標識領域の中心から少なくとも約250nmに位置する、請求項9~19のいずれか一項に記載の組成物。前記核酸ナノ構造が、標的結合部分をさらに含む、請求項9~20のいずれか一項に記載の組成物。前記標的結合部分が、一本鎖核酸である、請求項21に記載の組成物。前記標的結合部分が、抗体である、請求項21に記載の組成物。前記標的結合部分が、タンパク質又はペプチドである、請求項21に記載の組成物。標的核酸を識別する方法であって、以下:前記標的核酸が核酸バーコードプローブと結合するような条件下で、前記標的核酸を請求項9~24のいずれか一項に記載の核酸バーコードプローブと接触させるステップ;前記標的核酸と結合していない材料から前記標的核酸を分離するステップ;及び前記標的核酸と結合した前記核酸バーコードプローブの存在を検出するステップを含む、方法。標的タンパク質又は細胞を識別する方法であって、以下:前記標的タンパク質又は細胞が核酸バーコードプローブと結合するような条件下で、前記標的タンパク質又は細胞を請求項9~24のいずれか一項に記載の核酸バーコードプローブと接触させるステップ;前記標的タンパク質又は細胞と結合していない材料から前記標的タンパク質又は細胞を分離するステップ;及び前記標的タンパク質又は細胞と結合した前記核酸バーコードプローブの存在を検出するステップを含む、方法。標的タンパク質又は細胞を識別する方法であって、以下:前記標的タンパク質又は細胞が核酸バーコードプローブと結合するような条件下で、前記標的タンパク質又は細胞を請求項9~24のいずれか一項に記載の核酸バーコードプローブと接触させるステップ;前記標的タンパク質又は細胞と結合していない材料から前記標的タンパク質又は細胞を分離するステップ;及び前記標的タンパク質又は細胞と結合した前記核酸バーコードプローブの存在を検出するステップを含む、方法。標的を検出する方法であって、以下:サンプルを請求項9~24のいずれか一項に記載の核酸バーコードプローブと接触させるステップ、及び前記核酸バーコードプローブが、前記サンプル中の1つ以上の成分と結合するか否かを決定するステップを含み、前記核酸バーコードプローブと前記サンプル中の1つ以上の成分との結合が、前記サンプル中の標的の存在を示す、方法。前記接触ステップの後、前記核酸バーコードプローブに結合していないサンプル中の成分を除去するステップをさらに含む、請求項28に記載の方法。前記標的が、核酸、タンパク質、ペプチド、若しくは細胞、又はこれらの組合せである、請求項28又は29に記載の方法。前記核酸バーコードプローブの存在が、蛍光顕微鏡によって検出される、請求項25~30のいずれか一項に記載の組成物。前記核酸バーコードプローブの存在が、前記核酸バーコードプローブを電界に暴露することなく、検出される、請求項25~30のいずれか一項に記載の組成物。核酸バーコードプローブの集団であって、各プローブが、少なくとも2つの蛍光標識領域と標的結合部分を有する核酸ナノ構造を含み、前記核酸ナノ構造上の前記蛍光標識領域の各々の前記位置は、前記核酸バーコードプローブの蛍光標識のパターン及び前記結合部分の同一性が、前記蛍光標識領域による可視光線の発光に基づいて決定可能であるようにする、核酸バーコードプローブの集団。前記蛍光標識領域の各々が、他の蛍光標識領域の中心から少なくとも約50nm、少なくとも約100nm、少なくとも約200nm、又は約250nmに位置する中心を有する、請求項33に記載の集団。本発明は、中でも、核酸ナノ構造を含む核酸バーコードプローブを提供し、この核酸ナノ構造は、発蛍光団担持オリゴヌクレオチドと、安定に、又は一時的に結合従って、これで標識することができる1つ以上の蛍光標識領域を含む。バーコードプローブは、さらに、標的結合部分を含んでもよい。本発明はまた、このようなバーコードプローブの使用方法を提供し、こうした使用には、限定されないが、被検体アッセイ例:1種以上の被検体を検出し、任意に定量するためのアッセイにおけるその使用が含まれる。また、蛍光シグナルの性質例:波長又は色、強度など及びバーコード上での上記シグナルのパターン又は配向、若しくは配置若しくは幾何学的形状を用いて、サンプル中の特定の被検体を識別することもできる。これにより、複数の被検体を同時に又は、最低でも、単一のサンプル又はサンプルからの単一アリコートと一緒に検出することができる多重化アッセイが促進される。一態様では、本発明は、標的を検出する方法を提供し、この方法は、サンプルを核酸バーコードプローブと接触させるステップと、核酸バーコードプローブが、サンプル中の1つ以上の成分と結合するか否かを決定するステップを含み、ここで、核酸バーコードプローブとサンプルの1つ以上の成分との結合は、サンプル中の標的の存在を示し、また、核酸バーコードプローブは、標的結合部分と、少なくとも2つの蛍光標識領域とを含む核酸ナノ構造を含む。ある実施形態では、本方法はさらに、サンプルの1つ以上の成分と結合した核酸バーコードプローブの蛍光標識領域の色及び/又は配向に基づいて、標的を識別するステップを含む。ある実施形態では、核酸バーコードプローブが、サンプル中の1つ以上の成分と結合するか否かは、核酸バーコードプローブを、可溶性の、一時的に結合する発蛍光団担持オリゴヌクレオチドと接触させるステップを含む。ある実施形態では、核酸バーコードプローブは、安定に結合した発蛍光団担持オリゴヌクレオチドを含む。別の態様では、本発明は、標的を検出する方法を提供し、この方法は、標的が核酸バーコードプローブと結合するのに十分な条件下で、標的を核酸バーコードプローブと接触させるステップ;標的と結合していない材料から標的を分離するステップ;及び標的と結合した核酸バーコードプローブの存在を検出するステップを含み、この核酸バーコードプローブは、標的結合部分と、少なくとも2つの蛍光標識領域とを含む核酸ナノ構造を含む。ある実施形態では、本方法はさらに、標的と結合した核酸バーコードプローブの蛍光標識領域の色及び/又は配向に基づいて、標的を識別するステップを含む。ある実施形態では、標的と結合した核酸バーコードプローブの存在を検出するステップは、核酸バーコードプローブを、可溶性の、一時的に結合する発蛍光団担持オリゴヌクレオチドと接触させるステップを含む。ある実施形態では、核酸バーコードプローブは、安定に結合した発蛍光団担持オリゴヌクレオチドを含む。別の態様では、本発明は、少なくとも2つの蛍光標識領域を有する核酸ナノ構造を含む核酸バーコードプローブ、例えば、DNAナノ構造を含むDNAバーコードプローブを提供する。ある実施形態では、ナノ構造は、少なくとも3つの蛍光標識領域を含み、その配向は、蛍光標識領域の非対称間隔によって決定可能である。ナノ構造上の蛍光標識領域の各々の位置は、蛍光標識のパターンが、蛍光標識領域による可視光線の発光に基づいて決定可能であるようにする。ある実施形態では、蛍光標識領域による可視光線の発光に基づいて、バーコードプローブの配向が決定可能である。ある実施形態では、蛍光標識領域の各々は、別の蛍光標識領域の中心から少なくとも200nmの地点に位置する中心を有する。ある実施形態では、蛍光標識領域の各々は、別の蛍光標識領域の中心から少なくとも250nmの地点に位置する中心を有する。ある実施形態では、蛍光標識領域の各々は、別の蛍光標識領域の中心から少なくとも約25nm、少なくとも約50nm、少なくとも約75nm、又は少なくとも約100nm、若しくは少なくとも約150nmの地点に位置する中心を有する。ある実施形態では、上記ナノ構造は、スカフォールド鎖と複数のステープル鎖を有する。ある実施形態では、スカフォールド鎖は、M13バクテリオファージに由来する配列を有する。ある実施形態では、DNAナノ構造の蛍光標識領域は、発蛍光団で標識されたステープル鎖を含む。ある実施形態では、発蛍光団標識ステープル鎖は、一時的に結合した発蛍光団標識ステープル鎖である。ある実施形態では、発蛍光団標識ステープル鎖は、室温で一時的に結合する。ある実施形態では、発蛍光団標識ステープル鎖は、長さが7~12ヌクレオチドである。ある実施形態では、発蛍光団標識ステープル鎖は、長さが約9ヌクレオチドである。ある実施形態では、発蛍光団標識ステープル鎖は、安定に結合した発蛍光団標識ステープル鎖である。ある実施形態では、発蛍光団標識ステープル鎖は、室温で安定に結合する発蛍光団標識ステープル鎖である。ある実施形態では、発蛍光団標識ステープル鎖は、長さが少なくとも18ヌクレオチドであり、例えば、ある実施形態では、長さが18~25ヌクレオチドである。ある実施形態では、上記発蛍光団標識ステープル鎖は、直接標識される。ある実施形態では、発蛍光団標識ステープル鎖は、間接的に標識される。ある実施形態では、発蛍光団標識ステープル鎖は、スカフォールド鎖にハイブリダイズしたステープルドメインと、発蛍光団標識オリゴヌクレオチドにハイブリダイズしたハンドルドメインとを有する。ある実施形態では、ハンドルドメインは、長さが7~12ヌクレオチドである。ある実施形態では、ハンドルドメインは、長さが約9ヌクレオチドである。ある実施形態では、ハンドルドメインは、長さが少なくとも約18ヌクレオチドである。ある実施形態では、ハンドルドメインは、長さが18~25ヌクレオチドである。ある実施形態では、上記ナノ構造は、ナノチューブである。ある実施形態では、ナノ構造は、DNA折り紙法を用いて作製される。ある実施形態では、ナノ構造は、一本鎖DNAタイルを含む。ある実施形態では、ナノ構造は、DNAヘアピンを含む。ある実施形態では、上記核酸バーコードプローブは、さらに、標的結合部分を含む。ある実施形態では、標的結合部分は、核酸標的に相補的な一本鎖核酸である。ある実施形態では、標的結合部分は、抗体である。ある実施形態では、標的結合部分は、タンパク質又はペプチドである。ある実施形態では、ナノ構造は、いったん組み立てられると、静止状態で十分に不変イミュータブルであるため、その静止状態で光学顕微鏡法を用いて、蛍光領域のパターンを検出することができる。ある実施形態では、静止状態は、適用される電界の非存在を含む。別の態様では、本発明は、前述した核酸ナノ構造バーコードプローブのいずれか複数を含む組成物を提供する。ある実施形態では、上記複数のいくつか又は全メンバーは、上記複数の他のメンバーとは異なる。これらのメンバーは、標的結合部分並びにバーコードのシグナル及び配向配置に基づいて、互いに異なるものであってもよい。ある実施形態では、上記複数は、2、3、4、5、6、7、8、9、10、11、12、13、14、15、16、17、18、19、20、25、30、40、50、100、200以上の数と等しいか、それより小さい。ある実施形態では、上記複数は、216以下である。別の態様では、本発明は、標的核酸を識別する方法を提供し、この方法は、標的核酸がバーコードプローブと結合する条件下で、標的核酸を前記バーコードプローブのいずれかと接触させるステップ;標的核酸と結合していない材料から、標的核酸を分離するステップ;及び、標的核酸と結合したバーコードプローブの存在を検出するステップを含む。別の態様では、本発明は、標的タンパク質又は細胞を識別する方法を提供し、この方法は、標的タンパク質又は細胞がバーコードプローブと結合する条件下で、標的タンパク質又は細胞を前記バーコードプローブのいずれかと接触させるステップ;標的タンパク質又は細胞と結合していない材料から、標的タンパク質又は細胞を分離するステップ;及び、標的タンパク質又は細胞と結合したバーコードプローブの存在を検出するステップを含む。別の態様では、本発明は、標的タンパク質又は細胞を識別する方法を提供し、この方法は、標的タンパク質又は細胞がバーコードプローブと結合する条件下で、標的タンパク質又は細胞を前記バーコードプローブのいずれかと接触させるステップ;標的タンパク質又は細胞と結合していない材料から、標的タンパク質又は細胞を分離するステップ;及び、標的タンパク質又は細胞と結合したバーコードプローブの存在を検出するステップを含む。別の態様では、本発明は、標的を検出する方法を提供し、この方法は、サンプルを前記核酸バーコードプローブのいずれかと接触させるステップと、核酸バーコードプローブが、サンプル中の1つ以上の成分と結合するか否かを決定するステップを含み、ここで、核酸バーコードプローブとサンプル中の1つ以上の成分との結合は、サンプル中の標的の存在を示している。ある実施形態では、本方法は、接触ステップの後、核酸バーコードプローブと接触していないサンプル中の成分を除去するステップをさらに含む。ある実施形態では、標的は、核酸、タンパク質、ペプチド、若しくは細胞、又はこれらの組合せである。ある実施形態では、バーコードプローブの存在は、蛍光顕微鏡法によって検出する。ある実施形態では、DNAバーコードプローブを電界に暴露することなく、バーコードプローブの存在を検出する。ある実施形態では、各プローブは、少なくとも2つの蛍光標識領域と、標的結合部分とを含む核酸ナノ構造を含み、ナノ構造上の蛍光標識領域の各々の位置決めは、バーコードプローブの蛍光標識のパターン及び標的結合部分の同一性が、蛍光標識領域による可視光線の発光に基づいて決定可能であるようにし;また、蛍光標識領域の各々は、他の蛍光標識領域の中心から少なくとも200nmの地点に位置する中心を有する。別の態様では、本発明は、DNAバーコードプローブなどの核酸バーコードプローブの集団を提供し、各プローブは、少なくとも2つの蛍光標識領域と、標的結合部分とを有するDNAナノ構造のような核酸ナノ構造を含み、ここで、核酸ナノ構造上の蛍光標識領域の各々の位置決めは、核酸バーコードプローブの蛍光標識のパターン及び標的結合部分の同一性が、蛍光標識領域による可視光線の発光に基づいて、決定可能であるようにする。ある実施形態では、蛍光標識領域の各々は、他の蛍光標識領域の中心から少なくとも約25nm、少なくとも約50nm、少なくとも約100nm、少なくとも約200nm、若しくは少なくとも約250nmの地点に位置する中心を有する。本発明の前記及びその他の態様及び実施形態をここにさらに詳しく説明する。特許又は出願ファイルは、カラーで作成された少なくとも1枚の図面を含む。カラー図面を含む本特許又は特許出願公開公報のコピーは、要請及び必要費用の支払いに応じて、弊所により提供される。本願の公報は、図面のグレースケールレンダリングを含む。これら図面のカラー版は、要請に応じて取得可能である。本明細書において、完全にアドレス指定可能で、しかも標準的蛍光顕微鏡法を用いて、読み取りが可能な核酸例えば、DNAナノ構造バーコードプローブが提供される。これまで記載されている分子バーコードは、以下の2つの戦略のうち1つを用いて構築されている:1複数のスペクトル非重複発蛍光団を、制御されるモル比で組み合わせることにより、混合色シグネチャーを生成するもの;又は2回折限界約200nmを超える発蛍光団を分離して、これらを特定の幾何学模様に配置するもの。第1のカテゴリーのバーコードは、例えば、デンドリマーDNA例えばLietal.,Nat.Biotechnol.23:885-8892005を参照、自己集合2DDNA配列例えば、Linetal.,NanoLett.7:507-5122007を参照及び量子ドット例えば、Hanetal.,Nat.Biotechnol.19:631-6352001を参照から作製されている。このようなバーコードの構築及び解読の成功は、蛍光標識効率と、様々な蛍光濃度レベルを正確に検出するイメージング機器の能力の両方に大きく左右される。対照的に、第2の戦略で構築されたバーコードは、より頑健であり、標識効率及びイメージング条件が準最適である場合でも、構築し、検出することができる。さらに、このような方法を用いると、多重化能力は、含まれるそれぞれの追加蛍光スポットと共に急激に増大する。このカテゴリーに属するバーコードの例として、希土類元素をドーピングしたグラスマイクロファイバー長さ約100μm例えば、Dejnekaetal.,Proc.Nat.Acad.Sci.USA100:389-3932003を参照及び一方の鎖が特定位置で蛍光タグ付けされた二本鎖直鎖状DNA長さ2~5μm例えば、Xiaoetal.,Nat.Methods6:199-2012009及びGeissetal.,Nat.Biotechnol.26:317-3252008を参照がある。しかし、この戦略を用いる既存の方法は、蛍光スポットのアドレス指定可能性の点と、専用装置の非存在下では、蛍光スポットを可視光線の回折限界内にするような立体配座に、標識バーコードが折りたたまれてしまうため、バーコードが蛍光顕微鏡法を用いて読み取り不能になる点の両方で制限される。本発明者らは、既存の分子バーコード戦略に関連する問題が、DNAナノ構造のような核酸ナノ構造を用いて、新規の分子バーコードプローブを形成することによって、解決できることを見出した。DNAなどの核酸は、様々な技術、例えば、DNA折り紙法Rothemund、米国特許出願公開第2007/0117109A1号明細書、一本鎖タイルYinetal.,ProgrammingDNATubeCircumferences,Science2008:321:824-826、DNAヘアピンYinetal.米国特許出願公開第2009/0011956A1号明細書;Yinetal.Programmingbiomolecularself-assemblypathways,Nature2008451:318-323などを用いて、予め定めた1次元、2次元又は3次元ナノ構造に折りたたむことができる。一般に、DNA折り紙法は、1つ以上の長いスカフォールドDNA鎖を、合理的に設計した複数のステープルDNA鎖を用いて、特定の形状に折りたたむことを含む。ステープル鎖の配列は、これらが、スカフォールド鎖の特定部分にハイブリダイズし、これによって、スカフォールド鎖を強制的に特定の形状にするように設計される。DNA折り紙構造を作製する上で有用な方法は、例えば、米国特許出願公開第2007/0117109号明細書、同第2008/0287668号明細書、同第2010/0069621号明細書、及び同第2010/0216978号明細書に見出すことができ、これら文献の各々は、その全文を参照として本明細書に含むものとする。例えば、cadnanoウェブサイトにてインターネット上で入手可能なcaDNAnoソフトウエアを用いれば、ステープル設計を容易にすることができる。特定の実施形態では、DNAナノ構造バーコードプローブは、合理的に設計されたステープル鎖によって特定の形状に保持される1つ以上のスカフォールド鎖から成るDNAナノ構造を含むものでよい。配列表に、配列番号1として、スカフォールド鎖のヌクレオチド配列を記載するが、これを用いて、一例としてのDNA折り紙バーコードプローブを構築することができる。配列表にはまた、配列番号13~361として、ステープル鎖のヌクレオチド配列を記載するが、これらを用いて、配列番号1で示されるスカフォールド鎖と一緒に、一例としてのDNA折り紙バーコードプローブを構築することもできる。本明細書でさらに詳しく説明するが、配列番号13~186は、フロントモノマーを形成するのに用いられるステープル鎖のヌクレオチド配列を示し、また、配列番号187~361は、一例としてのDNA折り紙バーコードプローブ内にリアモノマーを形成するのに用いられるステープル鎖のヌクレオチド配列を示す。本明細書で用いるように、DNAナノ構造という用語は、便宜上用いるものであり、本発明が、核酸ナノ構造全般を考慮していることは理解すべきである。本発明のナノ構造は、直線状例:ナノロッド又は非直線状例:星形、三角形などのいずれであってもよい。DNAナノ構造は、少なくとも2つの非重複蛍光標識領域を有する。特定の実施形態では、DNAナノ構造バーコードプローブは、少なくとも3、4、5、6、7、8、9、10、11若しくは12の非重複蛍光標識領域を有する。プローブの発蛍光団パターンは、光学顕微鏡法を用いて決定する。ある実施形態では、蛍光標識領域の各々は、蛍光光学顕微鏡検査を用いて、視覚により識別することができる。これらの実施形態には、各蛍光標識領域の中心と、他の蛍光標識領域の中心との間隔が、可視光の回折限界より大きい距離すなわち、少なくとも200nmであるものもある。従って、特定の実施形態では、各蛍光標識領域は、他の蛍光標識領域いずれからも少なくとも200nmの地点にある。ある実施形態では、各蛍光標識領域は、他の蛍光標識領域のいずれからも少なくとも250nmの地点にある。ある実施形態では、さらに高い解像度が達成可能であり、蛍光標識領域同士の間隔は、より短い距離すなわち、少なくとも200nm未満の距離であってもよい。従って、ある実施形態では、蛍光標識領域間の距離は、175nm、150nm、125nm、100nm、75nm、50nm、25nm若しくはこれより小さい距離と等しいか、その近似値であってよい。蛍光標識領域同士の間隔が、可視光線の回折限界より小さい距離である場合には、超解像技術を用いて、バーコード化プローブをイメージングするが、こうした技術として、例えば、以下のものがある:PointAccumulationforImaginginNanoscaleTopographyPAINT,DNA-PAINT、StimulatedEmissionDepletionMicroscopySTED、ReversibleSaturableOpticalFluorescenceTransitionsRESOLFT、StochasticOpticalReconstructionMicroscopySTORM、dSTORM、PhotoactivatedLocalizationMicroscopyPALM、BlinkMicroscopyBM、並びにその他あらゆる形態の超解像顕微鏡法。蛍光標識領域という用語は、本明細書にさらに詳しく説明するように、ハンドルドメイン又はドッキング鎖若しくは配列に相補的な、発蛍光団担持部分、例えば、発蛍光団担持オリゴヌクレオチドで、安定に標識される領域及び一時的に標識される領域を包含する。ある形態では、本発明のバーコードプローブは、そこに結合されたこのような発蛍光団担持オリゴヌクレオチドと一緒に提供されるため、これらのプローブは蛍光性である。別の形態では、本発明のバーコードプローブは、そこに結合されたこのような発蛍光団担持オリゴヌクレオチドなしで提供されるため、これらのプローブは、発蛍光団担持オリゴヌクレオチドと結合するまで、蛍光性ではない場合もある。後者の場合には、プローブは、プローブのハンドルドメイン又はドッキング鎖に特異的な例えば、典型的には、相補的な複数の発蛍光団担持オリゴヌクレオチドと一緒に提供することができる。プローブ及びオリゴヌクレオチドは、キットに納めて、任意で各々をキット内の個別の容器に納めて提供してもよい。蛍光標識領域の位置は、DNAナノ構造バーコードの配向が、蛍光顕微鏡で、バーコードの視覚化によって典型的には、標識領域の配置のある非対称性の形態、又はある慣用的配置法、例えば、バーコードの末端を示すために維持される特定の色又はパターンにより決定可能であるように、選択することができる。ある形態では、配向情報は必要でないため、バーコードパターンが必ずしもユニークである必要はない。従って、2つの標識だけを含むバーコードを用いてもよく、様々な標識の存在及び/又は強度が、必要な情報を提供する。ある形態では、蛍光標識領域は、直線状に配置される。一例を例えば、図1に示す。しかし、蛍光標識領域はまた、非直線状に配置してもよい。一例を図8に示す。その他の非直線状配置は、当業者には明らかであり、本発明に含まれる。ある形態では、蛍光標識領域の非直線状配向又は配置若しくは構造を用いて、被検体を検出及び/又は識別する。ある形態では、非直線状に配置した蛍光標識領域のシグナル例えば、色又は色の組合せを含むを用いて、被検体を検出及び/又は識別する。さらに別の形態では、配向及びシグナルを一緒に用いて、被検体を検出及び/又は識別する。従って、ナノ構造が、直線状若しくは非直線状であってよい、及び/又は蛍光標識領域の配置が、直線状若しくは非直線状であってよいことは理解されたい。DNAナノ構造は、いったん組み立てられると、静止状態例えば、電界からの外的影響の非存在下で十分に不変イミュータブルであるため、その静止状態で光学顕微鏡検査を用いて、蛍光領域のパターンを検出することができる。ナノ構造の部分には、柔軟性でありうるが、そのような部分として、例えば非標識部分がある。ある実施形態では、プローブは、標識を担持する比較的剛性の部分と、標識されていない柔軟性の部分とを含み得る。ある実施形態では、プローブは、変形性リンカーによって接続される2つの剛性標識領域を有するものでよい。特定の実施形態では、DNAバーコードプローブのDNAナノ構造は、互いの間隔が少なくとも200nmである少なくとも3つの蛍光標識領域を支持することができる、どんな1次元、2次元又は3次元形状であってもよい。しかし、本明細書に記載するように、本発明は、互いの間隔が200nmに満たない蛍光標識領域を有するナノ構造も考慮されため、本発明はそのように限定されるわけではない。特定の実施形態では、DNAナノ構造は、ナノチューブである。ある実施形態では、ナノチューブは、長さが少なくとも200、300、400、500、600、700又は800nmである。特定の実施形態では、DNAナノ構造は、6へリックスバンドル二量体ナノチューブである。DNAナノチューブ設計DNA折り紙法によるについての記載は、例えば、Douglasetal.,Proc.Natl.Acad.Sci.USA104:6644-66482006及び米国特許出願公開第2010/0216978号明細書に見出すことができ、これらの文献は、各々、その全文を参照として本明細書に組み込む。ある実施形態では、DNAバーコードプローブは、DNAバーコードプローブの配向が、蛍光標識領域による可視光線の発光に基づいて決定可能であるように、設計される。DNAバーコードの配向を決定可能にするために、バーコードのどの特性を用いてもよい。例えば、DNAバーコードプローブが、ナノチューブのような実質的に1次元直線構造であれば、蛍光標識領域は、DNAナノ構造に沿って非対称的に配置することができる。実質的に一次元直線のDNAバーコードプローブの配向は、特定の発蛍光団又は発蛍光団の組合せを非対称に用いて、DNAナノ構造の片側を一貫して標識することにより、決定可能にすることもできる。DNAナノ構造が、2次元又は3次元である場合には、DNAバーコードプローブの配向は、例えば、構造自体の形状の非対称性に基づいて決定可能にすることもできる。特定の実施形態では、DNAバーコードプローブの蛍光標識領域は、蛍光部分でDNAナノ構造の特定のステープルを標識することによって生成する。ステープルは、直接標識してもよいし、間接的に標識してもよい。本明細書で用いる直接標識されるという用語は、検出可能な部分と共有結合される核酸を指す。これに対し、間接的に標識されるという用語は、1つ以上の非共有相互作用により、検出可能な部分に結合される核酸を指す。特定の実施形態では、DNAナノ構造の特定のステープルは、蛍光部分で直接標識する。このような実施形態では、例えば、DNAナノ構造への組込み前に、蛍光部分に結合、又は共有結合した特定の蛍光部分を用いて、ステープルを合成することができる。あらゆる組合せの蛍光部分を単一バーコードに用いてよいが、その際、各標識領域からの蛍光シグナルは、検出可能であるものとする。好ましくは、蛍光部分の組合せは、蛍光部分同士でエネルギー交換が一切ないすなわち、単一バーコードプローブに用いる発蛍光団組合せが、ドナー-アクセプター対として互いに作用する発蛍光団の対を含まないように選択する。ある実施形態では、DNAナノ構造の特定のステープルは、蛍光部分で間接的に標識される。このような実施形態では、こうしたステープルは、少なくとも2つのドメイン、すなわち、ステープルドメインとハンドルドメインを有するように合成することができる。ステープルドメインは、スカフォールド鎖にハイブリダイズして、DNAナノ構造の形成及び安定性に寄与するステープルの領域又はヌクレオチド配列である。ハンドルドメインは、DNAナノ構造の形成に必要ではない追加的核酸配列を含む。DNAナノ構造の形成前、形成中又は形成後、ハンドル配列を利用して、相補性DNA配列を有するオリゴヌクレオチドによりハイブリダイズさせることができる。従って、こうしたステープルは、ハンドルドメインを別の核酸にハイブリダイズすることによって、間接的に標識することができ、この核酸は、ハンドルに相補的な核酸配列を有し、それ自体が、蛍光部分で直接又は間接的のいずれかで標識される。ある実施形態では、DNAバーコードプローブの各蛍光領域は、少なくとも1つの蛍光標識ステープルを含む。特定の実施形態では、蛍光標識領域は、複数の蛍光標識ステープルを含んでいてよい。例えば、特定の実施形態では、各蛍光標識領域は、少なくとも2、3、4、5、6、7、8、9、10、11、12、又は13以上の蛍光標識ステープルを含んでいてよい。複数の標識ステープルを有するように設計された領域では、全てのステープルを単一タイプの発蛍光団によって標識してもよいし、あるいは、複数の発蛍光団タイプの組合せでステープルを標識してもよい。例えば、特定の実施形態では、蛍光標識領域は、予め定めた比で存在する複数のタイプの発蛍光団によって標識されるように設計する。後者のタイプの標識二重標識と呼ぶの例を実施例に示す。二重標識では、予め定めた比の2つの発蛍光団で単一領域を標識することによって、寄与する発蛍光団のいずれかから識別可能な蛍光シグナルを生成する。このように、バーコードプローブの多様性が増加している。さらに別の実施形態では、バーコードプローブは、発蛍光団担持オリゴヌクレオチド鎖で一時的に標識される1つ以上の領域を含んでもよい。これらの実施形態は、蛍光標識領域同士の間隔が、200nm以内である場合に用いてよい。ある実施形態では、バーコードに寄与する全ての領域を一時的に標識されるように設計する。一領域の一時的標識は、蛍光標識オリゴヌクレオチドの持続的結合に用いられるものより長さが短いハンドルドメインを含むステープル鎖から成るナノ構造を用いることによって達成される。一例として、発蛍光団担持オリゴヌクレオチドとステープル鎖のハンドルドメインの持続的又は安定結合は、長さが約21ヌクレオチドのオリゴヌクレオチド及びハンドルドメインを用いて、達成することができる。より短いオリゴヌクレオチド及びハンドルドメインを用いると、両者間の結合強度が低下するため、長い鎖より解離しやすくなる。室温で、長さが約9ヌクレオチドのオリゴヌクレオチド及びハンドルドメインは、一時的にしか互いに結合しない。当該技術分野では周知のように、より高い温度で、同じ会合/解離の動力学を達成するためには、概して、オリゴヌクレオチド及びハンドルドメインの長さを長くする。従って、本発明は、発蛍光団で直接又は間接的に標識されるハンドルドメイン、及び発蛍光団で一時的に標識されるものを考慮する。ハンドルドメインの長さ及び同様にその相補的オリゴヌクレオチドは、結合の性質すなわち、持続的又は一時的結合のいずれを意図するか、及び限定されないが、温度、塩濃度などの反応条件に応じて変動する。こうした長さは、限定しないが、約5ヌクレオチド~30ヌクレオチド、又は約7ヌクレオチド~約25ヌクレオチド、又は約9ヌクレオチド~約21ヌクレオチドの範囲であってよい。ある実施形態では、DNAバーコードプローブは、目的とする標的又は被検体の結合パートナーとして作用する標的結合部分をさらに含んでもよい。本明細書で用いる用語結合という用語は、例えば、生理的条件下での静電、疎水、イオン及び/又は水素結合相互作用による少なくとも2つの分子同士の会合を指す。このような標的結合部分は、例えば、限定しないが、オリゴヌクレオチドのような核酸、抗体若しくは抗体フラグメントのようなタンパク質若しくはペプチド、炭水化物若しくは多糖、又は小分子であってよい。また、標的結合部分と、これが結合する標的は、受容体とリガンドの関係で考察することもできる。特定の実施形態では、DNAバーコードプローブは、複数の標的捕捉部分を含んでもよく、これらは、同じでも異なっていてもよい。標的捕捉部分は、当該技術分野で公知のあらゆる方法を用いて、DNAバーコードプローブのDNAナノ構造に結合させることができる。例えば、標的捕捉部分は、ステープル鎖又はスカフォールド鎖に共有結合させることができる。捕捉部分はまた、例えば、前述のように、ステープル鎖のハンドルドメインにハイブリダイズさせることにより、ステープル鎖又はスカフォールド鎖のいずれかに間接的に結合させることもできる。特定の実施形態では、DNAバーコードプローブは、標的核酸にハイブリダイズすることができるオリゴヌクレオチド又は他の核酸である標的結合部分を含んでもよい。一般に、オリゴヌクレオチドは、それが、標的核酸の配列に実質的に相補的な核酸配列を含んでいれば、標的核酸にハイブリダイズすることができる。特定の実施形態では、オリゴヌクレオチドは、標的に完全に相補的なすなわち、標的配列と全てのヌクレオチドで塩基対合することができる配列を含んでもよい。ある実施形態では、オリゴヌクレオチドは、標的に対し、完全ではない相補性を有するが、特定の条件下ではやはり標的核酸とハイブリダイズすることができる。従って、特定の実施形態では、オリゴヌクレオチドの配列は、標的核酸の配列に、少なくとも10、20、30、40、50、60、65、70、75、80、85、90、又は95相補的である。特定の実施形態では、DNAバーコードプローブは、アプタマーである標的結合部分を含んでもよい。特定の実施形態では、DNAバーコードプローブは、抗体である標的結合部分を含んでもよい。本明細書で用いる抗体という用語は、全長抗体及び抗原結合フラグメントすなわち、抗原結合部分又はその一本鎖を包含する。抗体という用語は、限定されないが、ジスルフィド結合で相互結合される少なくとも2つの重H鎖と2つの軽L鎖を含む糖タンパク質、又はその抗原結合部分を含む。抗体は、ポリクローナル又はモノクローナル;異種間、同種異系、若しくは同系間;又はこれらの修飾形態例えば、ヒト化、キメラのいずれであってもよい。本明細書で用いる抗体の抗原結合部分というフレーズは、抗原に特異的に結合する能力を保持する抗体の1つ以上のフラグメントを指す。抗体の抗原結合機能は、完全長抗体のフラグメントによって達成される。抗体の抗原結合部分という用語に包含される結合フラグメントの例には、以下のものが含まれる:iFabフラグメント、すなわち、VH、VL、CL及びCH1ドメインからなる一価フラグメント;iiFab’2フラグメント、すなわち、ヒンジ領域でジスルフィド架橋によって結合された2つのFabフラグメントを含む二価フラグメント;iiiVH及びCH1ドメインからなるFdフラグメント;iv抗体の単一アームのVH及びVLドメインからなるFvフラグメント;vVHドメインからなるdAbフラグメントWardetal.,1989Nature341;544546;vi単離された相補性決定領域CDR又はvi任意で、合成リンカーによって結合されうる2つ以上の単離されたCDRの組合せ。さらに、Fvフラグメントの2つのドメイン、すなわちVH及びVLが、別の遺伝子によってコードされていても、これらは、組換え方法を用いて、合成リンカーにより結合することができ、このリンカーは、両ドメインを、VH及びVL領域が対合して一価分子を形成する一本のタンパク質鎖にすることができる一本鎖FvscFvとして知られる;例えば、Birdetal.1988Science242:423426;及びHustonetal.1988Proc.Natl.Acad.Sci.USA85:58795883を参照。このような一本鎖抗体もまた、抗体の抗原結合部分という用語に包含されるものとする。これらの抗体フラグメントは、当業者には公知の従来の技術を用いて得た後、インタクトな抗体と同じように、効用について上記フラグメントをスクリーニングする。特定の実施形態では、標的結合部分はタンパク質又はペプチドである。例えば、タンパク質又はペプチドを用いて、DNAバーコードプローブをタンパク質のリガンドと結合させることができる。また、タンパク質又はペプチド標的結合部分を有するDNAバーコードを用いて、限定するものではないが、以下のような抗体又は受容体を識別することもできる:細胞表面受容体例えば、限定されないが、T細胞受容体又はB細胞受容体、及び細胞内受容体例えば、限定されないが、タンパク質又はペプチドに存在するエピトープに結合することができるホルモン受容体。特定の実施形態では、標的結合部分は小分子である。小分子標的結合部分を有するDNAバーコードプローブを用いて、例えば、目的とする小分子の分子又は細胞標的を識別することができる。さらに、各小分子がDNAバーコードで個別にタグ付けされている、小分子のライブラリーを小分子ライブラリースクリーンに用いて、目的とする標的に特異的に結合する小分子を識別することができる。DNAバーコードプローブは、異なる種の集団として提供してもよい。異なるバーコードプローブの数は、上記集団内の特定のバーコードの多重化能力によってのみ限定される。特定の実施形態では、上記集団は、少なくとも10、50、100、500、1000、2000、3000、4000、5000、104、50000、105、105、106、107、108、109、1010、1011の異なるバーコードプローブを含む。上記集団は、1、2、3、4、5、6、7、8、9、10、11、12、13、14、15、16、17、18、19、20、21、22、23、24、25、26、27、28、29、30、又は31以上の異なる種を含んでもよい。上記集団は、約5、10、15、20、25、30、35、40、45、50、55、60、65、70、75、80、85、90、95、100以上の数以下の異なる種を含んでもよい。ある実施形態では、上記集団は、216の異なるバーコードプローブを含む。本明細書に記載するバーコードプローブは、識別可能な種のこのようなラージスケール集団の形成を可能にするのに十分適応可能かつ拡張可能である。使用方法DNAナノ構造バーコードプローブは、とりわけ、既存の分子バーコード技術が用いられるあらゆるアッセイに用いることができる。例えば、DNAバーコードプローブは、米国特許出願公開第2010/0015607号明細書に記載の方法に従って用いることができる。尚、上記文献の内容は、その全文を参照として本明細書に組み込む。典型的には、アッセイとしては、検出アッセイがあり、例えば、診断アッセイ、予後アッセイ、患者モニタリングアッセイ、スクリーニングアッセイ、細菌戦アッセイ、法医学分析アッセイ、出生前ゲノム診断アッセイなどが含まれる。アッセイは、invitroアッセイ又はinvivoアッセイのいずれであってもよい。分析するサンプルは、生体サンプル、例えば、血液、唾液、リンパ液、粘液、便、尿などであってよい。サンプルは、環境サンプル、例えば、水サンプル、空気サンプル、食品サンプルなどであってもよい。アッセイは、固定化された結合反応物の1つ以上の成分と一緒に実施してよい。従って、標的又はバーコードを固定化してよい。アッセイは、非固定化結合反応物の1つ以上の成分と一緒に実施することも可能である。本発明のバーコードプローブによって提供される多重化能力を考慮して、アッセイは、ほぼ同時に行う、サンプル中のいくつかの標的の検出を含むものであってもよい。一例として、特定の細胞型例えば、特異的細胞表面受容体に基づいて、さらには、この特定の細胞型における特定の遺伝的突然変異を検出するために、アッセイを用いてもよい。このようにして、エンドユーザーは、例えば、何個の特定の型の細胞が、対象の突然変異を保有しているかを決定することができる。特定の実施形態では、標的核酸を同定する方法は、オリゴヌクレオチド標的捕捉部分を有するDNAバーコードと、標的核酸を接触させることを含み、上記の部分は、標的核酸の配列にハイブリダイズすることができる核酸配列を含む。特定の実施形態では、標的核酸がDNAバーコードプローブと結合するような条件下で、標的核酸をDNAバーコードプローブと接触させる。ある実施形態では、次に、標的核酸と結合していない少なくとも一部の材料から、標的核酸を分離する。特定の実施形態では、続いて、標的核酸と結合したDNAバーコードプローブの存在を検出することによって、標的核酸の存在を決定する。DNAバーコードプローブの検出は、当該技術分野では公知のあらゆる方法により、例えば、蛍光顕微鏡法を用いて達成することができる。ある実施形態では、前述の方法は、第2標的核酸配列に相補的な核酸配列を含む捕捉プローブを固体支持体に固定化するステップをさらに含んでもよい。ある実施形態では、捕捉プローブ及びDNAバーコードプローブの両方と標的核酸を接触させる。標的核酸は、最初に捕捉プローブと、最初にDNAバーコードプローブと、又は同時に両プローブと接触させてもよい。標的核酸にハイブリダイズする前、標的核酸にハイブリダイズした後、又は標的核酸にハイブリダイズするのと同時に、捕捉プローブを固体支持体に固定化することができる。このような実施形態では、標的核酸の存在によって、固体支持体に固定化された、捕捉プローブ、標的核酸及びDNAバーコードプローブを含む第3複合体が形成されることになる。前述したように、次に、当該技術分野では公知のあらゆる適切な方法により、例えば、蛍光顕微鏡法を用いて、DNAバーコードプローブの存在を検出することによって、標的核酸の存在を決定する。特定の実施形態では、標的タンパク質又は細胞を同定する方法は、標的タンパク質又は細胞を、本明細書に記載のDNAバーコードプローブと接触させるステップを含んでもよい。特定の実施形態では、DNAプローブは、タンパク質、ペプチド又は抗体標的捕捉部分を含む。ある実施形態では、標的細胞又はタンパク質がDNAバーコードプローブと結合するような条件下で、DNAバーコードプローブを標的細胞又はタンパク質と接触させる。ある実施形態では、上記の方法はさらに、標的タンパク質又は細胞と結合していない少なくとも一部の材料から、標的タンパク質又は細胞を分離するステップを含む。特定の実施形態では、本方法はさらに、標的タンパク質又は細胞と結合したDNAバーコードプローブの存在を検出することによって、標的タンパク質又は細胞を同定するステップを含む。ある実施形態では、標的タンパク質は、抗体又は受容体、例えば、限定されないが、T細胞受容体である。本発明の態様及び実施形態のいくつかについては、画像取得の際、特に超解像アプローチを用いる場合及び/又はイメージング技術が、個別に取得した画像の重ね合わせを必要とする場合には、バーコードプローブ及び/又は標的を静止状態に維持することが好ましいことは理解されよう。超解像イメージング本明細書で述べるように、本発明のある態様では、バーコードプローブは、200nmすなわち、可視光線の回折限界未満の間隔で位置する蛍光標識領域を含む。これらの形態のいくつかでは、プローブをサンプルと接触させ、このサンプルの1つ以上の成分といったん結合したら、これを可溶性オリゴヌクレオチドに暴露するが、その各々は、上記領域の1つに特異的であり、発蛍光団で標識されている。このような可溶性の、一時的に結合するオリゴヌクレオチドは、本明細書ではイメージャー鎖と呼ぶこともある。ある形態では、サンプル中の非結合成分は、発蛍光団担持オリゴヌクレオチドの添加前に除去する。次に、発蛍光団担持オリゴヌクレオチドとプローブ上のその補体との結合を検出するために、一連の単一微速度撮影画像又は微速度撮影動画を用いて、残るサンプル成分をイメージングすることによって、バーコードプローブの存在及びそのバーコードに基づくその同一性を決定する。プローブ上の補体は、本明細書ではドッキング鎖と呼ぶこともある。任意の所定時点で、単一発蛍光団担持オリゴヌクレオチドだけがバーコードプローブと結合するように、条件を設定する。任意の所定時点で、単一発蛍光団担持オリゴヌクレオチドだけが結合すれば、たとえ隣接領域同士の間隔が200nm未満であっても、その色同一性を決定することができる同時に隣接領域に結合する発蛍光団がないため。このようにして、200nm未満の間隔で隣接する蛍光標識領域を配置することができ、これによって、バーコードのさらに高い多様性を生み出すことができる。これらの実施形態において、蛍光標識領域は、一時的に蛍光性であるにすぎず、バーコードは、イメージャー鎖が結合しているときだけ蛍光性であることは理解すべきである。ある実施形態では、イメージャー及び/又はドッキング鎖は、長さ約5~約18ヌクレオチド、又は長さ約6~約15ヌクレオチド、又は長さ約7~約12ヌクレオチドである。ある実施形態では、長さは、約8、9、10、11又は12ヌクレオチドである。当該技術分野では、他の局在化に基づく超解像蛍光技術、例えば、米国特許第7838302号明細書STORMに関するその開示内容は参照として本明細書に組み込むに記載のSTORM技術などが公知である。本発明は、あらゆる超解像技術に十分な方式でのナノ構造バーコードプローブの標識を考慮する。本発明の別の態様では、本発明のナノ構造バーコードプローブを用いて、蛍光顕微鏡法装置を較正及び/又はその較正を確認することもでき、こうした装置として、標準的又は従来の蛍光顕微鏡、同焦点顕微鏡、超解像蛍光顕微鏡などがある。従って、ナノ構造バーコードプローブは、較正基準又はツールとして用いてもよい。上記の態様で特に興味深いのは、可視光線の回折限界を下回る間隔すなわち、約200nm未満で位置する蛍光標識領域を有するバーコードプローブである。このように用いる場合には、プローブは、標的結合部分を有してもよいし、有していなくてもよい。一例として、プローブは、イメージングを目的とする固定化を促進する標的結合部分、例えば、ビオチン又はストレプトアビジンを含んでいてもよい。実施例1:DNAバーコードの形成800nmの6へリックバンドル二量体ナノチューブの3つの個別の領域を蛍光標識することによって、DNA折り紙バーコードプローブを構築した図1。長さ約28nmの蛍光標識領域の各々を、3つの発蛍光団の1つローダミン-グリーン、Cy3又はCy5に結合させた12のオリゴヌクレオチドで修飾した。DNA折り紙バーコードプローブは、最初の2つの標識領域同士の間隔約460nmを最後の2つの間隔約270nmより大きくして配置することによって、非対称になるように設計した。尚、対称配置を含む他の配置も本発明によって考慮されることは、理解すべきである。DNA折り紙バーコードプローブの主要構造は、cadnanoウェブサイトで入手可能なcaDNAnoソフトウエアを用いて、設計されるDNA6へリックスバンドルナノチューブ二量体である。各モノマーに用いるスカフォールド鎖の配列は、配列番号1として記載されている。DNA折り紙バーコードの構築に用いられるステープルの配列は、配列番号13~361として記載される。特定の選択されたステープルを伸長して合理的設計後、蛍光標識のための21nt一本鎖突出部ハンドルを含有させた。表1に、用いたハンドル及び抗ハンドル配列を記載する。従って、配列表に記載するステープル鎖のうち、配列番号:16、17、70~73、126-129、182及び183は青色に標識され;配列番号:195、196、239、240、251、252、295、296、307、308、351及び352は赤色に標識され;また、配列番号:216~219、272~275及び328~331は緑色に標識される。プローブのフロント及びリアモノマーの配置を示すcaDNAnoJSONファイルが、それぞれ6ht-v5-1color-1.json及び6ht-v6-2color-1.jsonとしてコンピュータープログラム一覧表の付録に提供されている。DNA折り紙バーコードプローブを組み立てるために、フロント及びリアモノマーを各々組み立て、個別の試験管中で蛍光標識した後、一緒に混合して、二量体を形成した。各モノマーの組み立ては、M13バクテリオファージ由来の100nMスカフォールド鎖p7308と称する配列番号1を、オリゴヌクレオチドステープル鎖のプールフロントモノマーについては配列番号13~186及びリアモノマーについては187~361;各々600nM;逆相カートリッジで精製、BioneerInc.と、5mMTris、1mMEDTA、20mMMgCl2、50mMNaClpH8を含むフォールディングバッファー中で混合した後、混合物を熱アニーリングランプ80分の時間をかけて80°Cから60°Cに冷却した後、15時間かけて60°Cから24°Cに冷却したに付すことによって、ワンポット反応で達成する。過剰ステープルをポリエチレングリコールPEG分画によって除去した後、各モノマーを、適切な発蛍光団を担持する抗ハンドルと一緒に、1:1.2モル比で、標識のためにインキュベートした。二量化のために、化学量の蛍光標識フロント及びリアモノマーを混合して、37°Cで2時間インキュベートした。最終生成物をアガロースゲル非変性、1.0電気泳動によって精製した。精製したナノバーコードは、-20°Cで少なくとも3か月保存することができる。この実施例では、2つのモノマーから成るバーコードを用いたが、DNAバーコードプローブは、3、4、5、又は6以上のモノマーを含むこともできる。従って、プラットフォームは、あらゆる所望のレベルに大きさを変更可能であり、あらゆる所望の数の異なるプローブを提供することができる。実施例2:蛍光顕微鏡を用いた単一標識DNAバーコードの検出前述のように、モノマーナノチューブの2つを折りたたんで、個別に蛍光標識した後、一緒にして、ナノバーコードを取得した。最終生成物をアガロースゲル電気泳動によって精製した後、全反射照明蛍光顕微鏡TIRFMを用いたイメージングのために、ガラススライド上に直接載せた。手短に言うと、5μLの精製ナノバーコード約20pMをガラススライド上に載せ、カバーガラスNo.1.5、18×18mm2、厚さ約0.17mmで挟み、5分静置した後、LeicaDM16000BTIRFMでイメージングした。各々擬色青、緑及び赤を割り当てた3つのチャンネルを通じて、サンプルを順次イメージングした。青色チャンネルについては、488nmレーザービームをダイクロイックミラー430/505/575/670で反射させ、100×対物レンズHCXPLAPO100×/1.47oilCORRTIRF,Leicaを介してサンプルに照射した。放射光は、同じ対物レンズを介して収集し、同じダイクロイックミラー及び外部吸収フィルターを介してフィルタリングした後、EM-CCDカメラHamamatsuC9100-02で、800ミリ秒間積分した。緑色及び赤色チャンネルについては、励起レーザーそれぞれ561及び635nm、吸収フィルターそれぞれ605/52及び705/72及び露光時間それぞれ500及び700ミリ秒を除いて同様の設計を用いた。イメージング工程は、LeicaLASソフトウエアによって自動的に制御した。画像は、ImageJ及び解読のためのカスタムソフトウエアにより処理した。以下の擬色を蛍光標識領域に割り当てた:AlexaFluor488/ローダミングリーン青色、Cy3緑色及びCy5赤色。TIRFM画像は、DNA折り紙バーコードを3つの異なるブライトスポットのストリップとして分解した。特に、青色及び緑色スポット間の距離は、緑色及び赤色間の距離より長く、これはバーコードの非対称性を示しているデータは示していない。各々が3つの異なる発蛍光団の1つによって標識されている3つの蛍光標識領域を有するDNA折り紙バーコードシステムは、3327の異なるDNA折り紙バーコードプローブを生成することができる。生成しうる27全てのDNA折り紙バーコードプローブを個別に組み立てて、アガロースゲル電気泳動により一緒に精製した。バーコードファミリー中の27のメンバーからの5つの異なるバーコードを品質管理実験のためにランダムに選択した。同じ実験条件下で、バーコードを組み立てて、個別に精製した後、イメージングした。バーコードの2つの異なる特徴をTIRF画像図2A,上のパネルからはっきりと見ることができた:第1に、バーコード上の各蛍光標識ゾーンが、単色スポットとして分解されたが、完全バーコードの各々は、このようなスポットの3つから構成された;第2に、隣接するスポットの2つが、小さな隙間によって分離されたが、他の7組の隣接する2つのスポットは、互いに接近したままであった。従って、専用の解読ソフトウエアを一切用いなくても、スポットの色同一性及びそれらの相対空間位置に基づき、幾何学的にコードされたバーコードを視覚によって認識し、解読することができる。BRGバーコード上の各スポットの中心を位置測定するカスタムソフトウエアを用いて、隣接スポット同士の平均中心間距離が、433±53nm平均値±s.d.、N=70;大きい距離及び264±52nm平均値±s.d.、N=70;小さい距離であることを測定し、バーコードの正確な形成を確認した。実験によって測定したこれらの距離は、設計した値478nm及び298nmよりやや小さかった。この相違は、ナノチューブ持続長:約1~2μmのランダムな熱曲げに起因するものと考えられ、これは他者23、28によって既に観測されており、本発明者らにより透過型電子顕微鏡法TEMを用いて確認されたデータは示さず。他のいくつかの幾何学的コード化バーコードシステム例:NanoStringnCounter12とは異なり、サンプルの調製に分子組合せのステップが一切なかったことに留意する必要がある。蛍光スポット同士の分離は、6へリックスバンドルナノチューブの固有に剛性の構造によってのみ引き起こされた。また、スポット強度は、画像全体で完全に均一ではないことにも注目すべきであり、これは、サンプル段階の不均質な照度及び蛍光効率の差などの要因によって説明することができる。それでも尚、TIRF画像から、バーコードが好適に組み立てられ、明瞭に分解できることが証明された。次に、選択した5つのバーコードについて、面積50×50μm2のTIRF画像を手動式で調べた図2A、下のパネル。画像内に見出される対象をまず、認定すなわち、設計通りにほぼ直線かつ非対称様式で配置された3つの単色スポットバーコードと、非認定すなわち、その他全ての対称バーコードに分類した。認定された8つのバーコードは、構成スポットの蛍光シグネチャーに基づいて適正及び非適正偽陽性バーコードにさらに分類して、バーコードが予想されるタイプであるか否かを表した。非認定バーコードは、1モノマーナノチューブ単一スポット又は2つのキッシングスポット、2[間違った形状すなわち、極端な曲げを有するバーコード、及び3複数の色を有する少なくとも1つのスポットを含むバーコードにさらに分類した。本発明者らの統計により、可視対象の70超が、完全に予想タイプすなわち、観察した188の認定バーコードのうち偽陽性がゼロであるとさらに決定された認定バーコードであることが明らかになった。非認定バーコードは、折りたたみの欠陥、取扱い中のサンプル損傷、及び表面へのナノチューブの重なりによって起こったと考えられるが、これらは、サンプル調製及びイメージングプロトコルを最適化することによって大幅に軽減することができる。典型的には、いくつかの異なるバーコード種を1つのプールに共存させる。従って、様々なタイプのバーコードを混合することにより、本システムの頑健性を試験することが重要である。最初の試験では、BRG及びRGBバーコードを個別に合成し、等モル比で互いに混合して、ゲル電気泳動によりこれらを共精製した。精製した混合物のTIRF分析から、2つのバーコードの1:1化学量論と、全体組み立て成功率認定バーコード/全対象約80が確認され、両バーコードが、混合及び共精製工程中その完全性を維持したことを示していた。さらに、認定バーコードの98超が、2つの予想タイプBRG及びRGBの一方に属した。2の偽陽性は、非予想バーコード、すなわちBGBによるものであったが、これは、BRGバーコードのフロントモノマーが、RGBバーコードのリアモノマーの近傍に位置した稀なケースに起因する可能性がある。これは、よりストリンジェントな精製条件を用いて、残留モノマーを最小限にすることにより、排除することができる。次に、全種を等モル量で混合したバーコードファミリーの全27メンバーのプールをイメージングすることにより、本システムを試験した。TIRF画像図2Bは、全タイプのバーコードが分解されたことを示している。27の50×50μm2画像合計約1,500バーコードをサンプリングすることによる統計から、標準偏差9で、バーコードタイプ毎に55の平均総数が明らかになり図2C、ピペット操作及びサンプリング誤差を考慮して予測した化学量論と十分に一致した。4つのカテゴリーこの場合、全27タイプが含まれていることから、適正及び非適正バーコードは識別不可能であることに注意されたいについて観察された対象の分布は、単一タイプのバーコードサンプルから測定した値と一致していた。以上の観察内容は、マイクロメートルに満たない長さのDNAナノチューブが、内在性の構造的剛性を備える幾何学的コード化バーコードを構築する上で信頼できるプラットフォームを呈示することを示している。ソフトウエアを用いて、様々なタイプのDNA折り紙バーコードプローブの画像を自動的に認識し、解読することができる。典型的には、このようなソフトウエアは、3つの基本的ステップ:認識、フィルタリング及び解読を有する。第1ステップでは、蛍光スポットを同定する。各蛍光チャンネル青色、赤色又は緑色について、ソフトウエアは、強度及びサイズに基づき蛍光スポットを同定し、続いて、全画像強度及び予め定めたスポットサイズを用いて自動的に決定された閾値と、上記スポットを比較する。こうした閾値を下回る蛍光スポットは、一般的バックグラウンドノイズと一緒に除去される。3つのチャンネル全てを処理した後、3つのチャンネルで識別したスポットを合わせることにより、バーコード画像を構成する。このようなバーコード画像の一例を図3に示す。第2ステップでは、DNA折り紙バーコードプローブの各領域間の距離蛍光スポットのx,y座標から計算すると、バーコード曲げ角中心スポットに基づくを計算する。次に、ソフトウエアは、DNA折り紙バーコードプローブをフィルタリングして、理論上の領域距離と一致しない領域距離を有するもの、又は現在のところ任意に設定された閾値角度140度より小さいバーコード曲げ角を有するものを除去する。一例として、以下の3つの距離規則を用いて、BRGバーコードをフィルタリングした:1青色及び緑色スポット間の距離は、15画素を超えてはならない理論上の距離は、11画素であった;2赤色及び青色スポット間の距離は、9画素を超えてはならない理論上の距離は、6.8画素であった;3赤色及び緑色スポット間の距離は、6画素を超えてはならない理論上の距離は、4.2画素であった。140度を下回る曲げ角を有する赤色スポットでBRGバーコードを除去した。最後のステップでは、全バーコードを識別し、フィルタリングした後、ソフトウエアは、様々な基準理論上の距離及び色に基づきを用いて、バーコードを特定のタイプに分類した。従って、複数のタイプのDNAバーコードを、それらが本来どのように設計されたかに基づいて解読することができる。最終的に構成されたDNAバーコードプローブ画像に加えて、ソフトウエアは、各DNAバーコードプローブ蛍光スポットのx,y座標、隣接スポット間の距離、バーコードの曲げ角、及びバーコードタイプに関する統計的情報を出力することができる。前述したソフトウエアを用いて、BRG及びGRGDNA折り紙バーコードプローブを検出した。図3に示すように、BRG及びGRGDNA折り紙バーコードプローブは、自動化解読ソフトウエアによって明瞭に同定された。ソフトウエアは、いくつかのフィルターを用いて、有効なDNA折り紙バーコードプローブを選択した。その結果、BRG及びGRGナノバーコードの88及び85がカウントされた。ソフトウエアは、BRG及びGRGバーコードに関する統計的情報を生成した表2が、これは、これらのDNA折り紙バーコードプローブ上の各領域間の測定距離が、設計通りの理論上の領域距離と十分に一致していることを示していた図3E。実施例3:DNAバーコードプローブを用いたオリゴヌクレオチド標的の検出DNAバーコードプローブを用いて、42-ntDNAオリゴヌクレオチド標的を検出した。2つの異なる精製バーコード、すなわち、42-ntオリゴヌクレオチド標的に特異的な21-ntオリゴヌクレオチド標的捕捉部分を含む緑色、赤色、緑色GRGと、標的捕捉部分を含まない青色、赤色、緑色BRGを1:1の比で混合して、フォールディングバッファーで20pMまで希釈した。標的1に特異的なビオチンプローブ100nM、0.2μLと標的110nM、0.4μLを100μLのバーコード混合物に添加した。100μLのバーコード混合物を含む個別の試験管に、0.2μLの標的用の100nMビオチンプローブ及び0.4μLの1×フォールディングバッファーを負の対照として添加した。上記サンプルをいずれも37°Cで一晩インキュベートした後、ストレプトアビジンを塗布したカバーガラスXenopore支持体を含むミクロ流体セルの個別チャンネルを介して流した。次に、両チャンネルを40μLの洗浄バッファーで5mMTris-HClpH8.0、500mMNaCl洗浄した後、前述したようにLeicaTIRFMでイメージングした。各チャンネルにおいてランダムな位置で10の画像を撮影し、DNA折り紙バーコードプローブの総数を手動式でカウントした。標的オリゴヌクレオチドの存在下では、ビオチンプローブは、1つの標的オリゴヌクレオチドの21nt配列にハイブリダイズし、また、GRGバーコードは、標的オリゴヌクレオチドの第2の21nt配列にハイブリダイズした。しかし、標的オリゴヌクレオチドに特異的な標的捕捉部分が欠如したBRGプローブは、標的にハイブリダイズせず、従って、ビオチンプローブによって結合されていない図4A。その結果、検出反応混合物をストレプトアビジン塗布ガラス表面全体に流すと、GRGバーコードは表面と結合するが、BRGバーコードは、洗い流される。TIRF画像及び統計分析は、検出の特異性を支持した図4B及びC。40pMオリゴヌクレオチド標的の存在下では、BRGバーコードの約20倍のGRGバーコードが表面と結合した。標的の非存在下では、いずれのバーコードも有意な表面結合を示さなかった。実施例4:二重標識バーコードプローブ多重化能力をさらに増大するために、ゾーン毎の6ステープル伸長部を変更して、標識のために12の同じ蛍光オリゴヌクレオチドを用いるのではなく、2つ以下の発蛍光団の組合せを用いて、各ゾーンについてユニークな蛍光シグネチャー擬色を生み出すようにした。スペクトルが識別可能な3つの発蛍光団を用いたこの二重標識戦略によって、6つの擬色B、R、G、BG、及びGRが生み出された。その結果、異なるバーコードの総数は、27から63=216へと増加したが、これは、多重化能力の大きさの増大を表すものである。単一標識ゾーンバーコードファミリーと同様に、二重標識ゾーンバーコードファミリーから5メンバーを品質管理の目的で選択した。バーコードは、重ね合わせた画像からのみ、又は異なるチャンネル全てを同時に検査するかのいずれかで、視覚により解読した。例えば、図5Aの第1列に示すように、バーコードBG--GR-BR--及び-は、それぞれ、バーコード内の大小のゾーン間距離を示すは、青色、緑色及び赤色チャンネルにおいて各々2つのスポットを呈示しているが、両者の隙間は小さくなっており、その設計と一致している。重ね合わせた画像では、バーコードは、シアン--黄色-ピンクとして見えたが、これは、予想通り、二重標識戦略によって起こる色の混合の結果であった。同様に、本発明者らは、さらに、他の4つの選択したバーコードの適正な形成も確認した図5A。ばらばらな標識効率と、イメージング中のわずかなサンプルのずれのために、二重標識ゾーンからの最終擬色は必ずしも均一ではなかった例えば、黄色スポットには緑がかっているものもあれば、赤みがかっているものもあったが、任意の特定スポットの蛍光シグネチャーは、3つのイメージングチャンネルから得た原画像を確認することにより、識別することができた。本発明者らは、各二重標識ゾーンバーコードの2つの50×50μm2画像を手動式で分析し、図5Bに統計データをプロットした。ここでは、対象を、その形状に基づいて認定バーコードと非認定バーコードに分類し、認定バーコードは、さらに適正及び非適正として類別した。対象の75~95は認定バーコードであり、その80~90は、適正タイプであったパーセンテージは、バーコードの具体的種類に応じて変動する。単一標識ゾーンバーコードファミリーと比較して、認定バーコードのパーセンテージは、同じままであったが、偽陽性率は、ゼロから10~20へと有意に増加した。観測されたこの偽陽性率の増加は、二重標識戦略の予想した頑健性低下単一標識と比較してと一致している。1つの設計では、二重標識ゾーンは、各発蛍光団種について、6つのステープル伸長部を有していたが、これは、単一標識ゾーンの半分の数に過ぎなかった。その結果、二重標識ゾーンバーコードは、光退色のような損傷をより受けやすい二量体スポットから構成された。この意味で、単一標識ゾーンバーコードは、冗長的にコード化された二重標識ゾーンバーコードとして考えることができる。例えば、単一標識ゾーンバーコードは、6つの発蛍光団が各ゾーンから欠失した場合でも、依然として適正タイプとして認識することができるが、そのようなシナリオにおいて、二重標識ゾーンバーコードは、認定を取り消されるか、又は非適正タイプとして類別されうる。後者の場合、バーコードは、偽陽性率を高めるであろう。原則として、偽陽性率は、二重標識ゾーンにおける各発蛍光団種のコピー数を増加することによって、下げることができる。試験した5つのバーコードは、2つの異なる発蛍光団種で標識されたそれらのゾーンの各々を有しており、そのため、二重標識ゾーンバーコードファミリーの最も誤りがちなメンバーに含まれる傾向が高くなる。従って、本発明者らは、ファミリー全体からの平均で、より小さい偽陽性率を予測する。本発明者らは、個別に組み立てて、共精製した72のバーコード種を含む混合物をイメージングすることにより、二重標識ゾーンバーコードシステムをさらに試験した。2ステップでの解読プロセスを支援するために、カスタムMATLSBスクリプトを用いた。ステップ1では、バーコードを含む3チャンネル赤色、緑色、青色TIRF画像を前処理することにより、バックグラウンドを除去した後、認定バーコードを含む画素だけが残るように、二値化した。得られる3チャンネル二値画像を合わせることにより、単一チャンネル二値画像を生成した。次に、ソフトウエアは、二値画像におけるその形状に基づいて、幾何学的に適合するバーコードの位置及び配向を識別した。ステップ2では、ステップ1で位置決定した各バーコードについて、3チャンネル画像の対応領域を、考えられるあらゆる基準バーコードのライブラリーと比較した。観察したバーコードには、最も高い相関を有する基準バーコードのアイデンティティーを付与した。完全自動化解読プロセス非監視モードは、前述した2つのステップの後に終了した。任意の監視モードにおいて、ソフトウエアは、ユーザーに、観察したバーコードと、承認のためのその最も可能性の高いアイデンティティーを提供した。監視及び非監視解読を比較して、コンピューターとユーザーとの間で、>80の同意が確認された。36の64×64μm23チャンネル画像のコンピューター支援監視モード分析によって、116の異なる種に属する約2,600の認定バーコードを登録した図5C、上のパネル。予想した72種が、全バーコード集団の約98を占め、種当たりの平均バーコードカウント数が36で、標準偏差は8であった。これに対し、非予想種は、平均して種当たり約1.4最大カウント数4バーコードに過ぎなかった。最後に、本発明者らは、二重標識ゾーンバーコードの全216メンバーを含む混合物を分析した。この混合物の60の64×64μm2画像を、非監視モードの解読ソフトウエアで処理した。完全自動化解読プロセスによって、標準偏差17で、種当たり約34バーコード合計約7,200バーコードが登録された図5C、下のパネル。比較的大きな標準偏差は、完全自動化データ分析における解読エラーによるものと考えられる。本発明者らの試験によって、216のバーコード種が好適に構築され、分解されたことが証明された図5D。実施例5:超解像度バーコードプローブ約100nm間隔で配置される蛍光要素を有する幾何学的コード化超解像度バーコードを用いて、より高い空間情報密度を有するバーコードも作製した。後者のアプローチの実現可能性実証として、本発明者らは、近年開発された超解像蛍光技術であるDNA-PAINT31を適用して、バーコードをイメージングした。この数年で、遠視野蛍光顕微鏡法40~44を用いて、回折限界を超えるイメージングを可能にする複数の技術が開発されている。ほとんどの超解像の実施において、発蛍光団を、蛍光ON及びOFF状態にスイッチすることにより、個々の分子を連続的に位置測定することができる。目標読み出し計画、例えば、StimulatedEmissionDepletionMicroscopy45STED又は他のReversibleSaturableOpticalFluorescenceTransitions40RESOLFT技術に依存する方法では、蛍光発生は、回折限界を下回る面積に能動的に限定される。蛍光分子のスイッチングはまた、推計学的に実施することができ、これは、例えば、直接StochasticOpticalReconstructionMicroscopy46、47STORM,dSTORM、PhotoactivatedLocalizationMicroscopy48PALM及びBlinkMicroscopy49BMにおいて行うが、これらにおいて、ほとんどの蛍光分子は、暗状態で作製され、蛍光を発するために推計学的にスイッチオンされるに過ぎない。PointAccumulationforImaginginNanoscaleTopography50PAINTでは、蛍光スイッチングは、蛍光分子で表面をターゲティングすることにより達成される。全ての推計学的アプローチでは、単一分子からの蛍光を回折限界面積内で位置測定することにより51、52、超解像画像を得る。DNA-PAINTは、蛍光標識オリゴヌクレオチドイメージャー鎖と、DNAナノ構造上の相補的ドッキング鎖の一時的結合を用いて、位置測定に基づく超解像顕微鏡法に必要な蛍光ON及びOFF状態間のスイッチングを達成する図6A参照。イメージャー/ドッキング鎖二重らせんの長さ及び溶液中のイメージャー鎖の濃度を調節することによって、蛍光ON及びOFF時間を独立に調節することができる。この試験のために、本発明者らは、スペクトルが異なる3つの色素に結合させた直交イメージャー鎖配列を用いた3色イメージング青色励起にはAtto488、緑色励起にはCy3B、また赤色励起にはAtto655に、DNA-PAINT技術を拡大した。3色超解像度バーコードシステムの実現可能性を証明するために、本発明者らは、対称配置で4つの結合ゾーンを含むDNAナノチューブモノマーを設計した。隣接ゾーン同士の間隔は、約114nmすなわち、回折限界を十分下回るであった。各結合ゾーンは、18のステープル鎖から構成され、これらは、伸長されて、赤色、緑色又は青色イメージャー鎖が結合するための3群の直交配列1群当たり6つを展示することができる。原理証明実験として、本発明者らは、5つの異なるバーコードを設計した図6A及び6Bの上のパネル。図6Bの下のパネルは、各チャンネルについて個別に、及び全チャンネルの重なりについて5つのバーコードの超解像再構成を示す。図6Cは、5つのバーコード全てを含む、より大きな面積を示す。3つのチャンネル全てにおいてバーコードのユニークなパターンを分解することができる。バーコードの中には、3色チャンネル全ての連続的イメージング中に動いたものもあったが、それでも分解可能であった。励起を変更すると共に、画像取得を高速化して、上記の作用を防止することによって、イメージングを改善することができた。DNA-PAINTは、光退色又は間違った標識鎖を被りにいため、イメージャー鎖とナノチューブ上のドッキング配列のイメージャー鎖の一時的で反復性の結合は、位置測定に必要なブリンキング挙動を生み出すだけではなく、イメージングプロトコルをより頑健にする。用いた顕微鏡セットアップでは、DNA-PAINTは、赤色チャンネルでは約46nm再構成PSFに対するガウス関数当てはめのFWHM、緑色チャンネルでは約25nm及び青色チャンネルでは約29nmの解像度を提供する。赤色イメージングチャンネルの解像度が低いのは、レーザー励起強度が弱いためである。より強力なTIRFシステムを用いると、緑色及び青色チャンネルと同様の、24nmの解像度を得ることができる。取得可能な解像度及びイメージング特異性から、バーコードの幾何学的非対称性を維持しながら、一ナノチューブモノマー上で6位置を頑健に分解できることが示され、これは、67=279,936の異なるバーコードをもたらしうる。さらに、ナノチューブ設計のモジュール性のために、バーコードのカスタマイズされたリエンジニアリングが可能になり、ここで、バーコードは、用いる顕微鏡の分解能に合わせて調節されるゾーン間距離を有しており、これによって、広範な顕微鏡セットアップに適用することが可能になる。実施例6:InSitu標識原理証明実験において、GRGバーコードを用いて、野生型カンジダアルビカンスCandidaalbicans酵母をタグ付けした。この酵母細胞を、まず、カンジダアルビカンスC.albicansに特異的なビオチン化ポリクローナル抗体と混合し、次いで、ストレプトアビジンの層を塗布した後、最後に、ビオチン化GRGバーコード図7Aと一緒にインキュベートした。TIRF顕微鏡法によって、酵母細胞の底部表面に結合したバーコードが明らかになった図7B、上のパネル。不都合にも酵母細胞の平らではない細胞壁上に落下したナノチューブもあったが、いくつかのGRGバーコードは、はっきりと視覚化することができる。対照的に、非ビオチン化抗体又はバーコードを用いて酵母を処理した場合、バーコードタグ付けは全く観察されなかった図7B、下のパネルが、これは、バーコードと細胞表面の間に非特異的相互作用がほとんど又は全く存在しなかったことを示している。実施例7:非直線状標識ナノ構造非直線構造のDNAナノ構造を組み立てて、より洗練されたバーコードを作製することができる。図8は、ステープル伸長部同士のハイブリダイゼーションにより図8A、挿入図、3つの約400nmDNAチューブを約60nmDNA環状DNAの外縁に結合させた例を示す。上記の環とナノチューブの遠端を蛍光標識することにより、蛍光顕微鏡法で明瞭に分解可能な3点星様構造が形成された。TIRF顕微鏡法及びTEM試験図8Bから、好適に折りたたまれたバーコードの約50が、設計通りに互いの角度120°前後で、環を取り囲む3つのナノチューブを特徴としたが、他の多くのバーコードは、環とナノチューブの間の半柔軟性二本鎖DNAリンカーのために、隣接するナノチューブの間の角度が有意に偏っていた。3つの同じサテライト直線状バーコードを中心ハブと接続させる類似の設計ここで、3つのサテライトバーコードは、共通の蛍光標識ゾーンとしてハブを共有しうるを用いて、3倍のコード化冗長性を有するバーコードを構築することができ、これらは優れた信頼性を特徴とすると考えられる。さらに、環と突起の間例えば、鎖交差を有する多重らせんDNAに、より剛性のリンカーを用いて、より確実に画定されたバーコード形状を強化することも可能である。参照文献1Han,M.,Gao,X.,Su,J.Z.Nie,S.Quantum-dot-taggedmicrobeadsformultiplexedopticalcodingofbiomolecules.Naturebiotechnology19,631-635,doi:10.1038/902282001.2Xu,H.etal.MultiplexedSNPgenotypingusingtheQbeadsystem:aquantumdot-encodedmicrosphere-basedassay.NucleicAcidsResearch31,e432003.3Li,Y.,Cu,Y.T.H.Luo,D.MultiplexeddetectionofpathogenDNAwithDNA-basedfluorescencenanobarcodes.Naturebiotechnology23,885-889,doi:10.1038/nbt11062005.4Livet,J.etal.Transgenicstrategiesforcombinatorialexpressionoffluorescentproteinsinthenervoussyst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